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24.
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<figureCitation id="D683DCB8FBF5D816A5AEBD131CBB4DA6" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Shell of Rhiostoma prestoni A-D originally labelled as &quot; Pterocyclos fruhstorferi Kobelt, 1909 &quot; A lectotype SMF 130353 from Chiem-hoa, Tonkin B paralectotype SMF 130354 from type locality and C, D paralectotype SMF 130355 from type locality E specimen SMF 130358 from Muong-Hum." figureDoi="10.3897/zookeys.1142.90097.figure49" httpUri="https://binary.pensoft.net/fig/799907" pageId="0" pageNumber="1">, 49</figureCitation>
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Bavay &amp; Dautzenberg, 1909b: 248, 249. Type locality: Binh-Lu, Tonkin [Binh Lieu, Quang Ninh Province, Vietnam].
<bibRefCitation author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="279 - 288" refId="B8" refString="Bavay, A, Dautzenberg, P, 1909a. Description de coquilles nouvelles de ľIndo-Chine (6e suite). Journal de Conchyliologie 57: 279 - 288" title="Description de coquilles nouvelles de ľIndo-Chine (6 e suite)." volume="57" year="1909 a">Bavay and Dautzenberg 1909a</bibRefCitation>
<bibRefCitation id="237024AECBD08F02568BCA787753F5AF" author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="279 - 288" refId="B8" refString="Bavay, A, Dautzenberg, P, 1909a. Description de coquilles nouvelles de ľIndo-Chine (6e suite). Journal de Conchyliologie 57: 279 - 288" title="Description de coquilles nouvelles de ľIndo-Chine (6 e suite)." volume="57" year="1909 a">Bavay and Dautzenberg 1909a</bibRefCitation>
: 283, 284, pl. 11, figs 1-3. Kobelt 1913: 969, 970, pl. 112, figs 5-7.
<bibRefCitation author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="171 - 196" refId="B188" refString="Zilch, A, 1956. Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4). Archiv fuer Molluskenkunde 85: 171 - 196" title="Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4)." volume="85" year="1956">Zilch 1956</bibRefCitation>
<bibRefCitation id="3C88D0E0390CCB2DE02DFC9BE209D90C" author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="171 - 196" refId="B188" refString="Zilch, A, 1956. Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4). Archiv fuer Molluskenkunde 85: 171 - 196" title="Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4)." volume="85" year="1956">Zilch 1956</bibRefCitation>
: 172.
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<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pterocyclos prestoni subsp. var. var. depicta" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="prestoni" subSpecies="var." variety="depicta">Pterocyclos prestoni var. depicta</taxonomicName>
<paragraph id="2B4A5295AF02962F91F82700B8FA3C84" pageId="0" pageNumber="1">
<taxonomicName id="635FC8BCF916EFFFBE4201414DD3EE2F" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pterocyclos prestoni subsp. var. var. depicta" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="prestoni" subSpecies="var." variety="depicta">Pterocyclos prestoni var. depicta</taxonomicName>
Bavay &amp; Dautzenberg, 1909b: 249. Type locality: Phong-Tho, Tonkin [Phong Tho, Lai Chau Province, Vietnam].
<bibRefCitation author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="279 - 288" refId="B8" refString="Bavay, A, Dautzenberg, P, 1909a. Description de coquilles nouvelles de ľIndo-Chine (6e suite). Journal de Conchyliologie 57: 279 - 288" title="Description de coquilles nouvelles de ľIndo-Chine (6 e suite)." volume="57" year="1909 a">Bavay and Dautzenberg 1909a</bibRefCitation>
<bibRefCitation id="FE78FD32D5A58C98E66B6930734B3BFA" author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="279 - 288" refId="B8" refString="Bavay, A, Dautzenberg, P, 1909a. Description de coquilles nouvelles de ľIndo-Chine (6e suite). Journal de Conchyliologie 57: 279 - 288" title="Description de coquilles nouvelles de ľIndo-Chine (6 e suite)." volume="57" year="1909 a">Bavay and Dautzenberg 1909a</bibRefCitation>
: 284, pl. 11, fig. 4.
<bibRefCitation author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="171 - 196" refId="B188" refString="Zilch, A, 1956. Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4). Archiv fuer Molluskenkunde 85: 171 - 196" title="Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4)." volume="85" year="1956">Zilch 1956</bibRefCitation>
<bibRefCitation id="AE2A9EDA5EBEF9AD96DB023C0D8DCF65" author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="171 - 196" refId="B188" refString="Zilch, A, 1956. Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4). Archiv fuer Molluskenkunde 85: 171 - 196" title="Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4)." volume="85" year="1956">Zilch 1956</bibRefCitation>
: 173. Kobelt 1913: 970, pl. 112, figs 8-9. New synonym.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pterocyclos fruhstorferi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fruhstorferi">Pterocyclos fruhstorferi</taxonomicName>
<paragraph id="7EE0920652312EF9FDED3FE2220CF8D5" pageId="0" pageNumber="1">
<taxonomicName id="131036F5F7E50A66D930BBC06AECF9F0" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pterocyclos fruhstorferi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fruhstorferi">Pterocyclos fruhstorferi</taxonomicName>
Kobelt, 1909: 82. Type locality: Chiem-hoa, Tonkin [Chiem Hoa, Tuyen Quang Province, Vietnam]. Kobelt 1911: 739, 740, pl. 107, figs 7-9.
<bibRefCitation author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="171 - 196" refId="B188" refString="Zilch, A, 1956. Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4). Archiv fuer Molluskenkunde 85: 171 - 196" title="Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4)." volume="85" year="1956">Zilch 1956</bibRefCitation>
<bibRefCitation id="76F62CA85ABEA6E002A740AE73BAFD4B" author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="171 - 196" refId="B188" refString="Zilch, A, 1956. Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4). Archiv fuer Molluskenkunde 85: 171 - 196" title="Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4)." volume="85" year="1956">Zilch 1956</bibRefCitation>
: 172.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Syntype</emphasis>
<subSubSection id="F1A1F89321EB763CB1BDEF0CE6140D61" pageId="0" pageNumber="1" type="type material">
<paragraph id="02EE9F7AA630A0573C59250B4C7D2B8A" pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph id="9CA2A6FB938573D3B84A1854C5293F00" pageId="0" pageNumber="1">
<emphasis id="998948582864654C7FA0D09EAFB4F947" bold="true" italics="true" pageId="0" pageNumber="1">Syntype</emphasis>
MNHN-IM-2000-30754 (1 shell; Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Shell of Rhiostoma prestoni A syntype MNHN-IM- 2000 - 30754 from Binh-Lu B syntype RBINS / MT / 525354 from Binh-Lu C-F originally labelled as &quot; Pterocyclus prestoni depictus Bavay &amp; Dautzenberg, 1908 &quot; C syntype MNHN-IM- 2000 - 30755 from Phong-Tho and D-F syntype RBINS / MT / 525355 from Phong Tho. Photograph: M Caballer and V Heros, MNHN (A, C)." figureDoi="10.3897/zookeys.1142.90097.figure48" httpUri="https://binary.pensoft.net/fig/799906" pageId="0" pageNumber="1">48A</figureCitation>
<figureCitation id="CAE0989F935F4CF16A09D6F562604D14" captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Shell of Rhiostoma prestoni A syntype MNHN-IM- 2000 - 30754 from Binh-Lu B syntype RBINS / MT / 525354 from Binh-Lu C-F originally labelled as &quot; Pterocyclus prestoni depictus Bavay &amp; Dautzenberg, 1908 &quot; C syntype MNHN-IM- 2000 - 30755 from Phong-Tho and D-F syntype RBINS / MT / 525355 from Phong Tho. Photograph: M Caballer and V Heros, MNHN (A, C)." figureDoi="10.3897/zookeys.1142.90097.figure48" httpUri="https://binary.pensoft.net/fig/799906" pageId="0" pageNumber="1">48A</figureCitation>
) from Binh-Lu (figured in
<bibRefCitation author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="279 - 288" refId="B8" refString="Bavay, A, Dautzenberg, P, 1909a. Description de coquilles nouvelles de ľIndo-Chine (6e suite). Journal de Conchyliologie 57: 279 - 288" title="Description de coquilles nouvelles de ľIndo-Chine (6 e suite)." volume="57" year="1909 a">Bavay and Dautzenberg 1909a</bibRefCitation>
<bibRefCitation id="F4DB469F2FD0F4FA46D5B5C1DE9EB609" author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="279 - 288" refId="B8" refString="Bavay, A, Dautzenberg, P, 1909a. Description de coquilles nouvelles de ľIndo-Chine (6e suite). Journal de Conchyliologie 57: 279 - 288" title="Description de coquilles nouvelles de ľIndo-Chine (6 e suite)." volume="57" year="1909 a">Bavay and Dautzenberg 1909a</bibRefCitation>
: pl. 11, figs 1-3) and
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">syntype</emphasis>
<emphasis id="1519816B147519F15B84BB9A9EC257CD" bold="true" italics="true" pageId="0" pageNumber="1">syntype</emphasis>
RBINS 525354 (1 shell; Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Shell of Rhiostoma prestoni A syntype MNHN-IM- 2000 - 30754 from Binh-Lu B syntype RBINS / MT / 525354 from Binh-Lu C-F originally labelled as &quot; Pterocyclus prestoni depictus Bavay &amp; Dautzenberg, 1908 &quot; C syntype MNHN-IM- 2000 - 30755 from Phong-Tho and D-F syntype RBINS / MT / 525355 from Phong Tho. Photograph: M Caballer and V Heros, MNHN (A, C)." figureDoi="10.3897/zookeys.1142.90097.figure48" httpUri="https://binary.pensoft.net/fig/799906" pageId="0" pageNumber="1">48B</figureCitation>
<figureCitation id="F638B791C33E7EC79B29CCABB22C0155" captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Shell of Rhiostoma prestoni A syntype MNHN-IM- 2000 - 30754 from Binh-Lu B syntype RBINS / MT / 525354 from Binh-Lu C-F originally labelled as &quot; Pterocyclus prestoni depictus Bavay &amp; Dautzenberg, 1908 &quot; C syntype MNHN-IM- 2000 - 30755 from Phong-Tho and D-F syntype RBINS / MT / 525355 from Phong Tho. Photograph: M Caballer and V Heros, MNHN (A, C)." figureDoi="10.3897/zookeys.1142.90097.figure48" httpUri="https://binary.pensoft.net/fig/799906" pageId="0" pageNumber="1">48B</figureCitation>
) from Binh Lu.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Syntype</emphasis>
<emphasis id="646A81A01F50169F765C4B5A6DD114D0" bold="true" italics="true" pageId="0" pageNumber="1">Syntype</emphasis>
MNHN-IM-2000-30755 (1 shell; Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Shell of Rhiostoma prestoni A syntype MNHN-IM- 2000 - 30754 from Binh-Lu B syntype RBINS / MT / 525354 from Binh-Lu C-F originally labelled as &quot; Pterocyclus prestoni depictus Bavay &amp; Dautzenberg, 1908 &quot; C syntype MNHN-IM- 2000 - 30755 from Phong-Tho and D-F syntype RBINS / MT / 525355 from Phong Tho. Photograph: M Caballer and V Heros, MNHN (A, C)." figureDoi="10.3897/zookeys.1142.90097.figure48" httpUri="https://binary.pensoft.net/fig/799906" pageId="0" pageNumber="1">48C</figureCitation>
<figureCitation id="42158134F5296C27CD7BBD2878394C4E" captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Shell of Rhiostoma prestoni A syntype MNHN-IM- 2000 - 30754 from Binh-Lu B syntype RBINS / MT / 525354 from Binh-Lu C-F originally labelled as &quot; Pterocyclus prestoni depictus Bavay &amp; Dautzenberg, 1908 &quot; C syntype MNHN-IM- 2000 - 30755 from Phong-Tho and D-F syntype RBINS / MT / 525355 from Phong Tho. Photograph: M Caballer and V Heros, MNHN (A, C)." figureDoi="10.3897/zookeys.1142.90097.figure48" httpUri="https://binary.pensoft.net/fig/799906" pageId="0" pageNumber="1">48C</figureCitation>
) of
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos prestoni subsp. var. var. depicta" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="prestoni" subSpecies="var." variety="depicta">Pterocyclos prestoni var. depicta</taxonomicName>
<taxonomicName id="7D63F5EAD9D04651833B65D5B3A81A02" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos prestoni subsp. var. var. depicta" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="prestoni" subSpecies="var." variety="depicta">Pterocyclos prestoni var. depicta</taxonomicName>
Bavay &amp; Dautzenberg, 1909, from Phong-Tho (figured in
<bibRefCitation author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="279 - 288" refId="B8" refString="Bavay, A, Dautzenberg, P, 1909a. Description de coquilles nouvelles de ľIndo-Chine (6e suite). Journal de Conchyliologie 57: 279 - 288" title="Description de coquilles nouvelles de ľIndo-Chine (6 e suite)." volume="57" year="1909 a">Bavay and Dautzenberg 1909a</bibRefCitation>
<bibRefCitation id="DE43CE8F015FE5BA555C3481608F5BE9" author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="279 - 288" refId="B8" refString="Bavay, A, Dautzenberg, P, 1909a. Description de coquilles nouvelles de ľIndo-Chine (6e suite). Journal de Conchyliologie 57: 279 - 288" title="Description de coquilles nouvelles de ľIndo-Chine (6 e suite)." volume="57" year="1909 a">Bavay and Dautzenberg 1909a</bibRefCitation>
: pl. 11, fig. 4) and
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">syntype</emphasis>
<emphasis id="661962634DD41D07FEC10BE6B2A7722D" bold="true" italics="true" pageId="0" pageNumber="1">syntype</emphasis>
RBINS 525355 (3 shells; Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Shell of Rhiostoma prestoni A syntype MNHN-IM- 2000 - 30754 from Binh-Lu B syntype RBINS / MT / 525354 from Binh-Lu C-F originally labelled as &quot; Pterocyclus prestoni depictus Bavay &amp; Dautzenberg, 1908 &quot; C syntype MNHN-IM- 2000 - 30755 from Phong-Tho and D-F syntype RBINS / MT / 525355 from Phong Tho. Photograph: M Caballer and V Heros, MNHN (A, C)." figureDoi="10.3897/zookeys.1142.90097.figure48" httpUri="https://binary.pensoft.net/fig/799906" pageId="0" pageNumber="1">48D-F</figureCitation>
<figureCitation id="290CB28AB5EC2828BB94FFEE99859621" captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Shell of Rhiostoma prestoni A syntype MNHN-IM- 2000 - 30754 from Binh-Lu B syntype RBINS / MT / 525354 from Binh-Lu C-F originally labelled as &quot; Pterocyclus prestoni depictus Bavay &amp; Dautzenberg, 1908 &quot; C syntype MNHN-IM- 2000 - 30755 from Phong-Tho and D-F syntype RBINS / MT / 525355 from Phong Tho. Photograph: M Caballer and V Heros, MNHN (A, C)." figureDoi="10.3897/zookeys.1142.90097.figure48" httpUri="https://binary.pensoft.net/fig/799906" pageId="0" pageNumber="1">48D-F</figureCitation>
) from Phong-Tho.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Lectotype</emphasis>
<emphasis id="579D816D8D79591F424337DDFAB8C492" bold="true" italics="true" pageId="0" pageNumber="1">Lectotype</emphasis>
(designation in
<bibRefCitation author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="171 - 196" refId="B188" refString="Zilch, A, 1956. Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4). Archiv fuer Molluskenkunde 85: 171 - 196" title="Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4)." volume="85" year="1956">Zilch (1956)</bibRefCitation>
<bibRefCitation id="67959E2F125011EA32ACEF1DF385FC2F" author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="171 - 196" refId="B188" refString="Zilch, A, 1956. Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4). Archiv fuer Molluskenkunde 85: 171 - 196" title="Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4)." volume="85" year="1956">Zilch (1956)</bibRefCitation>
) SMF 130353 (1 shell; Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Shell of Rhiostoma prestoni A-D originally labelled as &quot; Pterocyclos fruhstorferi Kobelt, 1909 &quot; A lectotype SMF 130353 from Chiem-hoa, Tonkin B paralectotype SMF 130354 from type locality and C, D paralectotype SMF 130355 from type locality E specimen SMF 130358 from Muong-Hum." figureDoi="10.3897/zookeys.1142.90097.figure49" httpUri="https://binary.pensoft.net/fig/799907" pageId="0" pageNumber="1">49A</figureCitation>
<figureCitation id="ECD7AF1E7D04BAA94CD9BDA3204CB50C" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Shell of Rhiostoma prestoni A-D originally labelled as &quot; Pterocyclos fruhstorferi Kobelt, 1909 &quot; A lectotype SMF 130353 from Chiem-hoa, Tonkin B paralectotype SMF 130354 from type locality and C, D paralectotype SMF 130355 from type locality E specimen SMF 130358 from Muong-Hum." figureDoi="10.3897/zookeys.1142.90097.figure49" httpUri="https://binary.pensoft.net/fig/799907" pageId="0" pageNumber="1">49A</figureCitation>
) of
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos fruhstorferi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fruhstorferi">
<emphasis italics="true" pageId="0" pageNumber="1">Pterocyclos fruhstorferi</emphasis>
<taxonomicName id="924AAE0ACE0E7AFF9F4EACB27507D4A3" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos fruhstorferi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fruhstorferi">
<emphasis id="AF73E0395A81C8B8404318F8B8C18EBD" italics="true" pageId="0" pageNumber="1">Pterocyclos fruhstorferi</emphasis>
</taxonomicName>
Kobelt, 1909, from Chiem-hoa, Tonkin,
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">paralectotype</emphasis>
<emphasis id="B60DB7C69BBED6A0C8983AF9CEC8C8F3" bold="true" italics="true" pageId="0" pageNumber="1">paralectotype</emphasis>
SMF 130354 (1 shell; Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Shell of Rhiostoma prestoni A-D originally labelled as &quot; Pterocyclos fruhstorferi Kobelt, 1909 &quot; A lectotype SMF 130353 from Chiem-hoa, Tonkin B paralectotype SMF 130354 from type locality and C, D paralectotype SMF 130355 from type locality E specimen SMF 130358 from Muong-Hum." figureDoi="10.3897/zookeys.1142.90097.figure49" httpUri="https://binary.pensoft.net/fig/799907" pageId="0" pageNumber="1">49B</figureCitation>
<figureCitation id="0CA18E3ED5B0A6E5AFFA6760B666C8FD" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Shell of Rhiostoma prestoni A-D originally labelled as &quot; Pterocyclos fruhstorferi Kobelt, 1909 &quot; A lectotype SMF 130353 from Chiem-hoa, Tonkin B paralectotype SMF 130354 from type locality and C, D paralectotype SMF 130355 from type locality E specimen SMF 130358 from Muong-Hum." figureDoi="10.3897/zookeys.1142.90097.figure49" httpUri="https://binary.pensoft.net/fig/799907" pageId="0" pageNumber="1">49B</figureCitation>
) and
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">paralectotype</emphasis>
<emphasis id="81AD662F0F010DBA9849EDE7DA41EA38" bold="true" italics="true" pageId="0" pageNumber="1">paralectotype</emphasis>
SMF 130355/2 (2 shells; Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Shell of Rhiostoma prestoni A-D originally labelled as &quot; Pterocyclos fruhstorferi Kobelt, 1909 &quot; A lectotype SMF 130353 from Chiem-hoa, Tonkin B paralectotype SMF 130354 from type locality and C, D paralectotype SMF 130355 from type locality E specimen SMF 130358 from Muong-Hum." figureDoi="10.3897/zookeys.1142.90097.figure49" httpUri="https://binary.pensoft.net/fig/799907" pageId="0" pageNumber="1">49C, D</figureCitation>
<figureCitation id="EA9E26ED07C4BA1B94578056E2AA01BF" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Shell of Rhiostoma prestoni A-D originally labelled as &quot; Pterocyclos fruhstorferi Kobelt, 1909 &quot; A lectotype SMF 130353 from Chiem-hoa, Tonkin B paralectotype SMF 130354 from type locality and C, D paralectotype SMF 130355 from type locality E specimen SMF 130358 from Muong-Hum." figureDoi="10.3897/zookeys.1142.90097.figure49" httpUri="https://binary.pensoft.net/fig/799907" pageId="0" pageNumber="1">49C, D</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1142.90097.figure48" httpUri="https://binary.pensoft.net/fig/799906" pageId="0" pageNumber="1" start="Figure 48" startId="F48">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 48.</emphasis>
<caption id="E20F5804ACB9947A1BB9E9765952C369" doi="10.3897/zookeys.1142.90097.figure48" httpUri="https://binary.pensoft.net/fig/799906" pageId="0" pageNumber="1" start="Figure 48" startId="F48">
<paragraph id="5913FCE51D5F32A97D00614EFA311884" pageId="0" pageNumber="1">
<emphasis id="2C469F9CDF28A4390991AB86BE6292C9" bold="true" pageId="0" pageNumber="1">Figure 48.</emphasis>
Shell of
<taxonomicName baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1909" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma prestoni" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="prestoni">
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma prestoni</emphasis>
<taxonomicName id="6219D1C205FFDCE93A27C4D9342F3BE6" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1909" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma prestoni" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="prestoni">
<emphasis id="C91B34E657C09F8EE0CA104AD16664EA" italics="true" pageId="0" pageNumber="1">Rhiostoma prestoni</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<emphasis id="729EBB2E56929395D4681FAFB6608452" bold="true" pageId="0" pageNumber="1">A</emphasis>
syntype MNHN-IM-2000-30754 from Binh-Lu
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
<emphasis id="204D7392C1AFD4C9AF415536EFEF9430" bold="true" pageId="0" pageNumber="1">B</emphasis>
syntype RBINS/MT/525354 from Binh-Lu
<emphasis bold="true" pageId="0" pageNumber="1">C-F</emphasis>
<emphasis id="4536D4CE2AD6986E024BB5A054BC5B89" bold="true" pageId="0" pageNumber="1">C-F</emphasis>
originally labelled as &quot;
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Pterocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterocyclus prestoni subsp. depictus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="prestoni" subSpecies="depictus">
<emphasis italics="true" pageId="0" pageNumber="1">Pterocyclus prestoni depictus</emphasis>
<taxonomicName id="3881B9BC36649449C27D9E9336A33CBB" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterocyclus prestoni subsp. depictus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="prestoni" subSpecies="depictus">
<emphasis id="81F4CB3DA14993432D665963950F6A6B" italics="true" pageId="0" pageNumber="1">Pterocyclus prestoni depictus</emphasis>
</taxonomicName>
Bavay &amp; Dautzenberg, 1908&quot;
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
<emphasis id="F59FB94218296E752481AFBECE57D6F8" bold="true" pageId="0" pageNumber="1">C</emphasis>
syntype MNHN-IM-2000-30755 from Phong-Tho and
<emphasis bold="true" pageId="0" pageNumber="1">D-F</emphasis>
<emphasis id="38BFE3F613BC76FC90A84BBF8879DC68" bold="true" pageId="0" pageNumber="1">D-F</emphasis>
syntype RBINS/MT/525355 from Phong Tho. Photograph: M Caballer and V Heros, MNHN (
<emphasis bold="true" pageId="0" pageNumber="1">A, C</emphasis>
<emphasis id="DD9D5F4C49FAA24B9E0EF4F8342BC347" bold="true" pageId="0" pageNumber="1">A, C</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1142.90097.figure49" httpUri="https://binary.pensoft.net/fig/799907" pageId="0" pageNumber="1" start="Figure 49" startId="F49">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 49.</emphasis>
<caption id="F496214110A9124262770666D4EEBD3A" doi="10.3897/zookeys.1142.90097.figure49" httpUri="https://binary.pensoft.net/fig/799907" pageId="0" pageNumber="1" start="Figure 49" startId="F49">
<paragraph id="1C2CAF01B9208763C1D985D02A8DBBF3" pageId="0" pageNumber="1">
<emphasis id="51CFFF0E0543925F629141C07139E6BE" bold="true" pageId="0" pageNumber="1">Figure 49.</emphasis>
Shell of
<taxonomicName baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1909" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma prestoni" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="prestoni">
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma prestoni</emphasis>
<taxonomicName id="CB6C2A0FCAC3C262356A3D60E2ED2049" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1909" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma prestoni" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="prestoni">
<emphasis id="68B58D928FE827ACE41DC212ED4C490E" italics="true" pageId="0" pageNumber="1">Rhiostoma prestoni</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">A-D</emphasis>
<emphasis id="25F52CEBEAA080452B4AC1639ACDAAF8" bold="true" pageId="0" pageNumber="1">A-D</emphasis>
originally labelled as &quot;
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos fruhstorferi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fruhstorferi">
<emphasis italics="true" pageId="0" pageNumber="1">Pterocyclos fruhstorferi</emphasis>
<taxonomicName id="F5DC3A79919EEF1BDC5A989575387805" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos fruhstorferi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fruhstorferi">
<emphasis id="6DA9EC8A155FD3C8EE78A2A8510C0876" italics="true" pageId="0" pageNumber="1">Pterocyclos fruhstorferi</emphasis>
</taxonomicName>
Kobelt, 1909&quot;
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<emphasis id="6846A89187AC0336330DA75D47E90BF9" bold="true" pageId="0" pageNumber="1">A</emphasis>
lectotype SMF 130353 from Chiem-hoa, Tonkin
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
<emphasis id="D28841C1627D49377A83DE453A0D1D4B" bold="true" pageId="0" pageNumber="1">B</emphasis>
paralectotype SMF 130354 from type locality and
<emphasis bold="true" pageId="0" pageNumber="1">C, D</emphasis>
<emphasis id="6E4CA0A12BA6F7558EB2BBDFC00FBD4C" bold="true" pageId="0" pageNumber="1">C, D</emphasis>
paralectotype SMF 130355 from type locality
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
<emphasis id="814E10B5EA6AF4344E498336EB378589" bold="true" pageId="0" pageNumber="1">E</emphasis>
specimen SMF 130358 from Muong-Hum.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingCountry name="Vietnam">Vietnam</collectingCountry>
<subSubSection id="A70ADE0386A6D86CE8D951A50372174A" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="F1471CDAFC0C2366B8DB7C37536D95A0" pageId="0" pageNumber="1">Other material examined.</paragraph>
<paragraph id="81CC05DC8152A280FE4D45C613551E40" pageId="0" pageNumber="1">
<emphasis id="CAB2129661B8BF64BD8E45E083981ABC" bold="true" pageId="0" pageNumber="1">
<collectingCountry id="CC83E36849C71EA4B0755ED66121027E" name="Vietnam">Vietnam</collectingCountry>
</emphasis>
: Trinh-Loung, Tonkin: SMF 130356. Muong-Hum, Tonkin: SMF 130357, 130358 (Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Shell of Rhiostoma prestoni A-D originally labelled as &quot; Pterocyclos fruhstorferi Kobelt, 1909 &quot; A lectotype SMF 130353 from Chiem-hoa, Tonkin B paralectotype SMF 130354 from type locality and C, D paralectotype SMF 130355 from type locality E specimen SMF 130358 from Muong-Hum." figureDoi="10.3897/zookeys.1142.90097.figure49" httpUri="https://binary.pensoft.net/fig/799907" pageId="0" pageNumber="1">49E</figureCitation>
<figureCitation id="11207851FF5D5DB718551D9345762AE2" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Shell of Rhiostoma prestoni A-D originally labelled as &quot; Pterocyclos fruhstorferi Kobelt, 1909 &quot; A lectotype SMF 130353 from Chiem-hoa, Tonkin B paralectotype SMF 130354 from type locality and C, D paralectotype SMF 130355 from type locality E specimen SMF 130358 from Muong-Hum." figureDoi="10.3897/zookeys.1142.90097.figure49" httpUri="https://binary.pensoft.net/fig/799907" pageId="0" pageNumber="1">49E</figureCitation>
). Ba-Xot, Tonkin: NHMW 71770/R/9406 (1 shell). Not-Son, Tonkin: NHMW 71770/R/9405 (1 shell). Tuah-Tuong, Tonkin: NHMW 71770/R/9404 (1 shell). Khe-So, Tonkin: NHMW 47842 (2 shells). Muong Bo, Tonkin: NHMW 71770/R/9407 (1 shell). Bo Xat, Tonkin: NHMW 71770/R/9403 (1 shell).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">Shell large, flattened to depressed shell, and without detached whorl. Breathing device canal-shaped, pointed anteriorly, and attached to preceding whorl. Peristome double, outer lip with wide canal, inner lip with wide and shallow incision. Operculum calcareous, low cup-shaped.</paragraph>
<subSubSection id="C463F7EBF5580B8E90E6C4BA4335F84F" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="B9BF3AD798838B49C9EDCF6906447A79" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="63581E99266628D37F9988802DAE11BE" pageId="0" pageNumber="1">Shell large, flattened to depressed shell, and without detached whorl. Breathing device canal-shaped, pointed anteriorly, and attached to preceding whorl. Peristome double, outer lip with wide canal, inner lip with wide and shallow incision. Operculum calcareous, low cup-shaped.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1909" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma prestoni" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="prestoni">
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma prestoni</emphasis>
<subSubSection id="A718615FC9C8DF57A8DFA39DD3986746" pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph id="10761B8D3CB10FE9EA0195E914B62EB8" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph id="B67A535138016CC1C3C35A7098F3BE16" pageId="0" pageNumber="1">
<taxonomicName id="C9EB895DDC97A7EAD5E2041DD63FE610" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1909" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma prestoni" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="prestoni">
<emphasis id="5569C0A910DFC7F834E9840EF0AAEF47" italics="true" pageId="0" pageNumber="1">Rhiostoma prestoni</emphasis>
</taxonomicName>
has a shell morphology that resembles that of
<taxonomicName lsidName="R. morleti" pageId="0" pageNumber="1" rank="species" species="morleti">
<emphasis italics="true" pageId="0" pageNumber="1">R. morleti</emphasis>
<taxonomicName id="2AFEE39DD20539D9DCF597C3988BDFFD" lsidName="R. morleti" pageId="0" pageNumber="1" rank="species" species="morleti">
<emphasis id="B89F8EA5BD476700B848DC4336602372" italics="true" pageId="0" pageNumber="1">R. morleti</emphasis>
</taxonomicName>
, but it differs in having the canal-shaped breathing device completely attached to the preceding whorl, while
<taxonomicName lsidName="R. morleti" pageId="0" pageNumber="1" rank="species" species="morleti">
<emphasis italics="true" pageId="0" pageNumber="1">R. morleti</emphasis>
<taxonomicName id="E521D8713DAC85BD64FD651629449FF1" lsidName="R. morleti" pageId="0" pageNumber="1" rank="species" species="morleti">
<emphasis id="F50AE0FC8118BDFA8BB007D647FC0D8C" italics="true" pageId="0" pageNumber="1">R. morleti</emphasis>
</taxonomicName>
has an incomplete tube or a canal-shaped breathing device not attached to the preceding whorl.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Shell</emphasis>
<subSubSection id="8ECFE04991AA0AFE5E285DB60BABACD8" pageId="0" pageNumber="1" type="description">
<paragraph id="3E91AC0353519DCE1E50DAE85CCFC304" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="B988587C72FCFD2ECA76DD0668D2BE4E" pageId="0" pageNumber="1">
<emphasis id="88890DA23CC110F3F6BA352ED2F6D9FD" bold="true" pageId="0" pageNumber="1">
<emphasis id="FAE967E8DBCC07572AAC474383587E60" italics="true" pageId="0" pageNumber="1">Shell</emphasis>
.
</emphasis>
Shell medium to large, width 22.5-29.1 mm, height 7.8-12.0 mm. Shell thickened, and flattened shape. Apex acute; spire flat to slightly elevated. Whorls 4 to 5, convex, increasing regularly; suture wide and deep; last whorl rounded, stout and wider than penultimate whorl. Shell surface with fine growth lines. Periostracum thick, corneous, and brownish. Shell colour varies from uniformly brownish to brownish zigzag patterns; brownish peripheral band present or absent. Detached whorl absent. Peristome circular and double; lip thickened and slightly expanded. Aperture opened sub-laterally. Breathing device canal-shaped and attached to preceding whorl, causing it to appear without a detached whorl; outer lip protruding, expanded at base and forming wide canal; inner lip with a wide and shallow incision. Umbilicus widely opened. Operculum calcareous, cup-shaped, and multispiral (Figs
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Shell of Rhiostoma prestoni A syntype MNHN-IM- 2000 - 30754 from Binh-Lu B syntype RBINS / MT / 525354 from Binh-Lu C-F originally labelled as &quot; Pterocyclus prestoni depictus Bavay &amp; Dautzenberg, 1908 &quot; C syntype MNHN-IM- 2000 - 30755 from Phong-Tho and D-F syntype RBINS / MT / 525355 from Phong Tho. Photograph: M Caballer and V Heros, MNHN (A, C)." figureDoi="10.3897/zookeys.1142.90097.figure48" httpUri="https://binary.pensoft.net/fig/799906" pageId="0" pageNumber="1">48</figureCitation>
<figureCitation id="93FF5A305EBD664A1BD84FD6B01B458E" captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Shell of Rhiostoma prestoni A syntype MNHN-IM- 2000 - 30754 from Binh-Lu B syntype RBINS / MT / 525354 from Binh-Lu C-F originally labelled as &quot; Pterocyclus prestoni depictus Bavay &amp; Dautzenberg, 1908 &quot; C syntype MNHN-IM- 2000 - 30755 from Phong-Tho and D-F syntype RBINS / MT / 525355 from Phong Tho. Photograph: M Caballer and V Heros, MNHN (A, C)." figureDoi="10.3897/zookeys.1142.90097.figure48" httpUri="https://binary.pensoft.net/fig/799906" pageId="0" pageNumber="1">48</figureCitation>
,
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Shell of Rhiostoma prestoni A-D originally labelled as &quot; Pterocyclos fruhstorferi Kobelt, 1909 &quot; A lectotype SMF 130353 from Chiem-hoa, Tonkin B paralectotype SMF 130354 from type locality and C, D paralectotype SMF 130355 from type locality E specimen SMF 130358 from Muong-Hum." figureDoi="10.3897/zookeys.1142.90097.figure49" httpUri="https://binary.pensoft.net/fig/799907" pageId="0" pageNumber="1">49</figureCitation>
<figureCitation id="D03460268A69F07C58D284E75FE3893C" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Shell of Rhiostoma prestoni A-D originally labelled as &quot; Pterocyclos fruhstorferi Kobelt, 1909 &quot; A lectotype SMF 130353 from Chiem-hoa, Tonkin B paralectotype SMF 130354 from type locality and C, D paralectotype SMF 130355 from type locality E specimen SMF 130358 from Muong-Hum." figureDoi="10.3897/zookeys.1142.90097.figure49" httpUri="https://binary.pensoft.net/fig/799907" pageId="0" pageNumber="1">49</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
<subSubSection id="87F3373FE00F92BE8D084F47C9F60F09" pageId="0" pageNumber="1" type="distribution">
<paragraph id="FCF03D960C53B9B0700B17878C5707B9" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="EC8B67608C8CC2DF6532C6BB8C98DD2D" pageId="0" pageNumber="1">
This species is mainly distributed in Lai Chau, Quang Ninh, and Tuyen Quang provinces of northern Vietnam (Fig.
<figureCitation captionStart="Figure 44" captionStartId="F44" captionText="Figure 44. Distribution map of Rhiostoma morleti - species group." figureDoi="10.3897/zookeys.1142.90097.figure44" httpUri="https://binary.pensoft.net/fig/799902" pageId="0" pageNumber="1">44</figureCitation>
<figureCitation id="D85474BFB83BB7BDCAAE28867C643A28" captionStart="Figure 44" captionStartId="F44" captionText="Figure 44. Distribution map of Rhiostoma morleti - species group." figureDoi="10.3897/zookeys.1142.90097.figure44" httpUri="https://binary.pensoft.net/fig/799902" pageId="0" pageNumber="1">44</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1909" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma prestoni" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="prestoni">
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma prestoni</emphasis>
<subSubSection id="13EEB30C84C42422F97AD12466D24E27" pageId="0" pageNumber="1" type="remarks">
<paragraph id="0DD61E0649ACA53C01CA077360168ED1" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="B6CA266583FDC11641D278306D291FD3" pageId="0" pageNumber="1">
<taxonomicName id="F57A80CED571158ECC6E1658DC4E1B24" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1909" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma prestoni" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="prestoni">
<emphasis id="E1E0BCBE1792B2280F0D24EBFD679928" italics="true" pageId="0" pageNumber="1">Rhiostoma prestoni</emphasis>
</taxonomicName>
has long been classified as a member of the
<taxonomicName authorityName="Benson" authorityYear="1832" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pterocyclos</emphasis>
<taxonomicName id="5D6E8AD6FDE8F34B99A807CC63EBAD0D" authorityName="Benson" authorityYear="1832" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="75320FDE5764E89B79DE74BB2106AEB7" italics="true" pageId="0" pageNumber="1">Pterocyclos</emphasis>
</taxonomicName>
(see Kobelt 1911;
<bibRefCitation author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="171 - 196" refId="B188" refString="Zilch, A, 1956. Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4). Archiv fuer Molluskenkunde 85: 171 - 196" title="Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4)." volume="85" year="1956">Zilch 1956</bibRefCitation>
<bibRefCitation id="ED0EE34723EF9353E3B115B881EEE4CA" author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="171 - 196" refId="B188" refString="Zilch, A, 1956. Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4). Archiv fuer Molluskenkunde 85: 171 - 196" title="Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4)." volume="85" year="1956">Zilch 1956</bibRefCitation>
). The operculum stuck inside the last whorl of the syntype is a calcareous cup shape and multispiral with elevated lamellae (Figs
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Shell of Rhiostoma prestoni A syntype MNHN-IM- 2000 - 30754 from Binh-Lu B syntype RBINS / MT / 525354 from Binh-Lu C-F originally labelled as &quot; Pterocyclus prestoni depictus Bavay &amp; Dautzenberg, 1908 &quot; C syntype MNHN-IM- 2000 - 30755 from Phong-Tho and D-F syntype RBINS / MT / 525355 from Phong Tho. Photograph: M Caballer and V Heros, MNHN (A, C)." figureDoi="10.3897/zookeys.1142.90097.figure48" httpUri="https://binary.pensoft.net/fig/799906" pageId="0" pageNumber="1">48</figureCitation>
<figureCitation id="404208305E881545C35B78F3B7935BFF" captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Shell of Rhiostoma prestoni A syntype MNHN-IM- 2000 - 30754 from Binh-Lu B syntype RBINS / MT / 525354 from Binh-Lu C-F originally labelled as &quot; Pterocyclus prestoni depictus Bavay &amp; Dautzenberg, 1908 &quot; C syntype MNHN-IM- 2000 - 30755 from Phong-Tho and D-F syntype RBINS / MT / 525355 from Phong Tho. Photograph: M Caballer and V Heros, MNHN (A, C)." figureDoi="10.3897/zookeys.1142.90097.figure48" httpUri="https://binary.pensoft.net/fig/799906" pageId="0" pageNumber="1">48</figureCitation>
,
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Shell of Rhiostoma prestoni A-D originally labelled as &quot; Pterocyclos fruhstorferi Kobelt, 1909 &quot; A lectotype SMF 130353 from Chiem-hoa, Tonkin B paralectotype SMF 130354 from type locality and C, D paralectotype SMF 130355 from type locality E specimen SMF 130358 from Muong-Hum." figureDoi="10.3897/zookeys.1142.90097.figure49" httpUri="https://binary.pensoft.net/fig/799907" pageId="0" pageNumber="1">49</figureCitation>
<figureCitation id="37FE0060FF8D1864C1FB81B2455985BD" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Shell of Rhiostoma prestoni A-D originally labelled as &quot; Pterocyclos fruhstorferi Kobelt, 1909 &quot; A lectotype SMF 130353 from Chiem-hoa, Tonkin B paralectotype SMF 130354 from type locality and C, D paralectotype SMF 130355 from type locality E specimen SMF 130358 from Muong-Hum." figureDoi="10.3897/zookeys.1142.90097.figure49" httpUri="https://binary.pensoft.net/fig/799907" pageId="0" pageNumber="1">49</figureCitation>
); these are the distinguishing characters of the genus
<taxonomicName authorityName="Tongkerd &amp; Tumpeesuwan &amp; Inkhavilay &amp; Prasankok &amp; Jeratthitikul &amp; Panha &amp; Sutcharit" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma</emphasis>
<taxonomicName id="1E966970568BA76A393B1AF19D9D89F0" authorityName="Tongkerd &amp; Tumpeesuwan &amp; Inkhavilay &amp; Prasankok &amp; Jeratthitikul &amp; Panha &amp; Sutcharit" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="5D842C1783D4357799B503BEEA8C1689" italics="true" pageId="0" pageNumber="1">Rhiostoma</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of breathing device and operculum characteristics among the five genera that closely resemble Rhiostoma. The specimens and opercula represented herein are the type specimen or topotype of the respective type species of each genus; Pterocyclos: P. rupestris Benson, 1832 (lectotype UMZC I. 102070, no locality data), Spiraculum: S. hispidum Pearson, 1833 (specimen NHMUK 1906.4.4.72, from Khasi, Teria Ghat), Rhiostoma: R. haughtoni Benson, 1860 (syntype UMZC I. 103445, from Moulmein), Cyclotus: C. variegatus Pearson, 1833 (specimen NHMUK 1896.6.7.34 - 35, from Palawan), Opisthoporus: Cyclostoma taylorianum Pfeiffer, 1851 = junior synonym of Pterocyclos biciliatum Mousson, 1849 (specimen NHMUK 20210200 for shell, from Basan, Sarawak) and Pterocyclos tener Menke, 1856 (NHMUK ex. Cuming collection for operculum, from Cochin China), and Ptychopoma: Cyclophorus chinensis Moellendorff, 1875 (lectotype SMF 39415, from Lushan bei Kiukiang, China). Red arrows indicate the expansion of outer lip at upper junction of the peristome. Blue arrows indicate breathing tubes (apertural tube in Rhiostoma and sutural tubes in Spiraculum and Opisthoporus)." figureDoi="10.3897/zookeys.1142.90097.figure5" httpUri="https://binary.pensoft.net/fig/799863" pageId="0" pageNumber="1">5</figureCitation>
<figureCitation id="6934E4E796A7FE92B5E63E877EE79661" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of breathing device and operculum characteristics among the five genera that closely resemble Rhiostoma. The specimens and opercula represented herein are the type specimen or topotype of the respective type species of each genus; Pterocyclos: P. rupestris Benson, 1832 (lectotype UMZC I. 102070, no locality data), Spiraculum: S. hispidum Pearson, 1833 (specimen NHMUK 1906.4.4.72, from Khasi, Teria Ghat), Rhiostoma: R. haughtoni Benson, 1860 (syntype UMZC I. 103445, from Moulmein), Cyclotus: C. variegatus Pearson, 1833 (specimen NHMUK 1896.6.7.34 - 35, from Palawan), Opisthoporus: Cyclostoma taylorianum Pfeiffer, 1851 = junior synonym of Pterocyclos biciliatum Mousson, 1849 (specimen NHMUK 20210200 for shell, from Basan, Sarawak) and Pterocyclos tener Menke, 1856 (NHMUK ex. Cuming collection for operculum, from Cochin China), and Ptychopoma: Cyclophorus chinensis Moellendorff, 1875 (lectotype SMF 39415, from Lushan bei Kiukiang, China). Red arrows indicate the expansion of outer lip at upper junction of the peristome. Blue arrows indicate breathing tubes (apertural tube in Rhiostoma and sutural tubes in Spiraculum and Opisthoporus)." figureDoi="10.3897/zookeys.1142.90097.figure5" httpUri="https://binary.pensoft.net/fig/799863" pageId="0" pageNumber="1">5</figureCitation>
). Therefore, we have reclassified this species as a member of
<taxonomicName authorityName="Tongkerd &amp; Tumpeesuwan &amp; Inkhavilay &amp; Prasankok &amp; Jeratthitikul &amp; Panha &amp; Sutcharit" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma</emphasis>
<taxonomicName id="E5DC70B651A51D1CE7E702FDFD3908DC" authorityName="Tongkerd &amp; Tumpeesuwan &amp; Inkhavilay &amp; Prasankok &amp; Jeratthitikul &amp; Panha &amp; Sutcharit" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="931133070B9CAB2A454C35872A4C4F54" italics="true" pageId="0" pageNumber="1">Rhiostoma</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<paragraph id="727C548230B4CB273137C7058EF87108" pageId="0" pageNumber="1">
We examined specimens labelled as
<taxonomicName lsidName="R. prestoni" pageId="0" pageNumber="1" rank="species" species="prestoni">
<emphasis italics="true" pageId="0" pageNumber="1">R. prestoni</emphasis>
<taxonomicName id="64DB6AB0B805E66D285610446009A9AD" lsidName="R. prestoni" pageId="0" pageNumber="1" rank="species" species="prestoni">
<emphasis id="572163D4453E16C6BA6FB6081805973E" italics="true" pageId="0" pageNumber="1">R. prestoni</emphasis>
</taxonomicName>
in the RBINS ex. Dautzenberg collection from northern Vietnam that show a high variation in shell colour, from monochrome yellowish to brownish zigzag pattern. However, the shape of the breathing device has less variation than the shell characters and colour patterns. Therefore, we retain
<taxonomicName lsidName="R. prestoni" pageId="0" pageNumber="1" rank="species" species="prestoni">
<emphasis italics="true" pageId="0" pageNumber="1">R. prestoni</emphasis>
<taxonomicName id="F00BD3786703DEB862D01CC9E9705011" lsidName="R. prestoni" pageId="0" pageNumber="1" rank="species" species="prestoni">
<emphasis id="C8B9E5A321B61D216373EA54DC6EA1D4" italics="true" pageId="0" pageNumber="1">R. prestoni</emphasis>
</taxonomicName>
as a distinct species until the DNA phylogeny becomes available and confirms their status and relationship.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<paragraph id="87847ACB7A9355BE71900EF89D39D57F" pageId="0" pageNumber="1">
The nominal species,
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos fruhstorferi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fruhstorferi">
<emphasis italics="true" pageId="0" pageNumber="1">Pterocyclos fruhstorferi</emphasis>
<taxonomicName id="B735694C7B71E2168F78039C201609FC" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos fruhstorferi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fruhstorferi">
<emphasis id="EA6AAA83C75AE01A895F86A06CE03D50" italics="true" pageId="0" pageNumber="1">Pterocyclos fruhstorferi</emphasis>
</taxonomicName>
Kobelt, 1909, was described based on specimens from northern Vietnam. The unique name-bearing type was subsequently designated in
<bibRefCitation author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="171 - 196" refId="B188" refString="Zilch, A, 1956. Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4). Archiv fuer Molluskenkunde 85: 171 - 196" title="Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4)." volume="85" year="1956">Zilch (1956</bibRefCitation>
<bibRefCitation id="CA79607FEEFBD5DA2AF4E1689ACC9F52" author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="171 - 196" refId="B188" refString="Zilch, A, 1956. Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4). Archiv fuer Molluskenkunde 85: 171 - 196" title="Die typen und typoide des Natur-Museums Senckenberg, 18, Mollusca Cyclophoridae, Cyclophorinae (4)." volume="85" year="1956">Zilch (1956</bibRefCitation>
: 172). The lectotype has a shell form similar to that of
<taxonomicName lsidName="R. prestoni" pageId="0" pageNumber="1" rank="species" species="prestoni">
<emphasis italics="true" pageId="0" pageNumber="1">R. prestoni</emphasis>
<taxonomicName id="50CC687FF48734C01FF4E1DB7765416B" lsidName="R. prestoni" pageId="0" pageNumber="1" rank="species" species="prestoni">
<emphasis id="73D27E87556D35506C5BD14DEA4C653E" italics="true" pageId="0" pageNumber="1">R. prestoni</emphasis>
</taxonomicName>
, and the only noticeable difference is the brownish zigzag colour pattern. In contrast, the syntype of
<taxonomicName lsidName="R. prestoni" pageId="0" pageNumber="1" rank="species" species="prestoni">
<emphasis italics="true" pageId="0" pageNumber="1">R. prestoni</emphasis>
<taxonomicName id="9AF558BEFA7EB4D06991AEE5F5688D1B" lsidName="R. prestoni" pageId="0" pageNumber="1" rank="species" species="prestoni">
<emphasis id="033B87B7EBB148B3F7DBE2C0F66F9602" italics="true" pageId="0" pageNumber="1">R. prestoni</emphasis>
</taxonomicName>
has a monochrome yellowish brown shell colour. Therefore, we agree with Kobelt (1911) in recognising
<taxonomicName lsidName="P. fruhstorferi" pageId="0" pageNumber="1" rank="species" species="fruhstorferi">
<emphasis italics="true" pageId="0" pageNumber="1">P. fruhstorferi</emphasis>
<taxonomicName id="90C68B0F5DB7E7CF68665C0010A87D96" lsidName="P. fruhstorferi" pageId="0" pageNumber="1" rank="species" species="fruhstorferi">
<emphasis id="D5BA715FEEC90CDFAE525DC9D1A9C55D" italics="true" pageId="0" pageNumber="1">P. fruhstorferi</emphasis>
</taxonomicName>
as a junior subjective synonym of
<taxonomicName lsidName="R. prestoni" pageId="0" pageNumber="1" rank="species" species="prestoni">
<emphasis italics="true" pageId="0" pageNumber="1">R. prestoni</emphasis>
<taxonomicName id="8F6FA5A4E92311CD2126F908A47FBA1C" lsidName="R. prestoni" pageId="0" pageNumber="1" rank="species" species="prestoni">
<emphasis id="ED88B542E8CB81C00FD91E20751FF8B3" italics="true" pageId="0" pageNumber="1">R. prestoni</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<paragraph id="E616BD85CB66603799A6F01563246BFA" pageId="0" pageNumber="1">
The specimen figured in
<bibRefCitation author="Do, DS" journalOrPublisher="Ruthenica" pageId="0" pageNumber="1" pagination="117 - 132" refId="B34" refString="Do, DS, Nguyen, THT, Do, VN, 2015. A checklist and classification of terrestrial prosobranch snails from Son La, north-western Vietnam. Ruthenica 25: 117 - 132" title="A checklist and classification of terrestrial prosobranch snails from Son La, north-western Vietnam." volume="25" year="2015">Do et al. (2015</bibRefCitation>
<bibRefCitation id="818DFE192F781DC0C4E597E6FBE9EB18" author="Do, DS" journalOrPublisher="Ruthenica" pageId="0" pageNumber="1" pagination="117 - 132" refId="B34" refString="Do, DS, Nguyen, THT, Do, VN, 2015. A checklist and classification of terrestrial prosobranch snails from Son La, north-western Vietnam. Ruthenica 25: 117 - 132" title="A checklist and classification of terrestrial prosobranch snails from Son La, north-western Vietnam." volume="25" year="2015">Do et al. (2015</bibRefCitation>
: 122, fig. 4b) as
<taxonomicName authorityName="Bavay &amp; Dautzenberg" authorityYear="1909" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos prestoni" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="prestoni">
<emphasis italics="true" pageId="0" pageNumber="1">Pterocyclos prestoni</emphasis>
<taxonomicName id="42D7664ED62E6A9E26FE8C86BCF7ECBF" authorityName="Bavay &amp; Dautzenberg" authorityYear="1909" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos prestoni" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="prestoni">
<emphasis id="002EF49F93AB76DFBA61F2DB79920F20" italics="true" pageId="0" pageNumber="1">Pterocyclos prestoni</emphasis>
</taxonomicName>
Bavay &amp; Dautzenberg, 1909 exhibits a corrugated shell surface with nodules arranged on growth lines over the entire shell, and with canal-shaped breathing device protruding apically and attached to preceding whorl. These characters clearly differ from the typical
<taxonomicName lsidName="R. prestoni" pageId="0" pageNumber="1" rank="species" species="prestoni">
<emphasis italics="true" pageId="0" pageNumber="1">R. prestoni</emphasis>
<taxonomicName id="5586954FBB3844E33DB1A5AB56ACC5BB" lsidName="R. prestoni" pageId="0" pageNumber="1" rank="species" species="prestoni">
<emphasis id="C14B0FD87656CF5E3695217460CBA645" italics="true" pageId="0" pageNumber="1">R. prestoni</emphasis>
</taxonomicName>
and represent an undescribed species. However, this specimen has no operculum for comparison; therefore, the generic placement is still provisional.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<paragraph id="31DD85B7458A5C6FC5ABC75C6AA8919F" pageId="0" pageNumber="1">
Finally, the 'var.
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma prestoni var. depicta" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="prestoni" variety="depicta">Rhiostoma prestoni depicta</taxonomicName>
<taxonomicName id="F48D1F797DD3F7E9EA646A71EB934FD4" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma prestoni var. depicta" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="prestoni" variety="depicta">Rhiostoma prestoni depicta</taxonomicName>
' was identified based on the distinct shell colour having brownish zigzags or blotches. However, the shell pattern alone cannot be considered the distinguishing character for recognising species or subspecies, since several species of the genus
<taxonomicName authorityName="Tongkerd &amp; Tumpeesuwan &amp; Inkhavilay &amp; Prasankok &amp; Jeratthitikul &amp; Panha &amp; Sutcharit" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma</emphasis>
<taxonomicName id="313443669F545633788EC02FAB766232" authorityName="Tongkerd &amp; Tumpeesuwan &amp; Inkhavilay &amp; Prasankok &amp; Jeratthitikul &amp; Panha &amp; Sutcharit" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis id="1ED749D4572027350C0767FDDE305368" italics="true" pageId="0" pageNumber="1">Rhiostoma</emphasis>
</taxonomicName>
have high variation in shell colour and pattern.
</paragraph>