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<mods:namePart>Tongkerd, Piyoros</mods:namePart>
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<mods:namePart>Tumpeesuwan, Sakboworn</mods:namePart>
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<mods:namePart>Inkhavilay, Khamla</mods:namePart>
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<mods:namePart>Prasankok, Pongpun</mods:namePart>
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11.
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<taxonomicName LSID="https://zoobank.org/24A470B8-4449-4FF9-802F-AC4DC0BCBB11" authority="Tongkerd & Panha" authorityName="Tongkerd & Panha" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma ebenozostera" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ebenozostera" status="sp. nov.">Rhiostoma ebenozostera Tongkerd & Panha</taxonomicName>
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution map of Rhiostoma housei - species group." figureDoi="10.3897/zookeys.1142.90097.figure18" httpUri="https://binary.pensoft.net/fig/799876" pageId="0" pageNumber="1">Figs 18</figureCitation>
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<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shell of Rhiostoma ebenozostera sp. nov. A holotype CUMZ 4703 from Tham Pha Poo, Loei B, C paratypes from type locality B paratype CUMZ 4704 and C paratype CUMZ 3889 D specimen CUMZ 10211 from Lom Kao, Phetchabun E specimen CUMZ 4765 from Suwannakhuha, Nongboa Lumphu F specimen CUMZ 10166 / 2 from Phu Wiang, Khon Kaen." figureDoi="10.3897/zookeys.1142.90097.figure29" httpUri="https://binary.pensoft.net/fig/799887" pageId="0" pageNumber="1">, 29</figureCitation>
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<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Radula morphology A Rhiostoma housei specimen CUMZ 3982 from Wat Tham Srivilai, Saraburi B Rhiostoma hainesi specimen from Chanthaburi C Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sua, Krabi D Rhiostoma ebenozostera sp. nov. paratype CUMZ 4704 from Tham Pha Poo, Loei E Rhiostoma lannaense sp. nov. paratype CUMZ 4701 from Chiang Dao, Chiang Mai F Rhiostoma tigrina sp. nov. paratype CUMZ 4497 / 2 from Mae Sai, Chiang Rai." figureDoi="10.3897/zookeys.1142.90097.figure32" httpUri="https://binary.pensoft.net/fig/799890" pageId="0" pageNumber="1">, 32D</figureCitation>
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|
||||
<figureCitation id="417116D3C7C8D340CEE21082ACEF171D" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shell of Rhiostoma ebenozostera sp. nov. A holotype CUMZ 4703 from Tham Pha Poo, Loei B, C paratypes from type locality B paratype CUMZ 4704 and C paratype CUMZ 3889 D specimen CUMZ 10211 from Lom Kao, Phetchabun E specimen CUMZ 4765 from Suwannakhuha, Nongboa Lumphu F specimen CUMZ 10166 / 2 from Phu Wiang, Khon Kaen." figureDoi="10.3897/zookeys.1142.90097.figure29" httpUri="https://binary.pensoft.net/fig/799887" pageId="0" pageNumber="1">, 29</figureCitation>
|
||||
<figureCitation id="4018C462045A362B1446C016798C6A75" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Radula morphology A Rhiostoma housei specimen CUMZ 3982 from Wat Tham Srivilai, Saraburi B Rhiostoma hainesi specimen from Chanthaburi C Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sua, Krabi D Rhiostoma ebenozostera sp. nov. paratype CUMZ 4704 from Tham Pha Poo, Loei E Rhiostoma lannaense sp. nov. paratype CUMZ 4701 from Chiang Dao, Chiang Mai F Rhiostoma tigrina sp. nov. paratype CUMZ 4497 / 2 from Mae Sai, Chiang Rai." figureDoi="10.3897/zookeys.1142.90097.figure32" httpUri="https://binary.pensoft.net/fig/799890" pageId="0" pageNumber="1">, 32D</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="type material">
|
||||
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
|
||||
<subSubSection id="DC817132AFF90FB6D57BFB26215C8AF7" pageId="0" pageNumber="1" type="type material">
|
||||
<paragraph id="375B4785CFB624000DC0721C7D16C893" pageId="0" pageNumber="1">Type material.</paragraph>
|
||||
<paragraph id="6F09401B1615626198A3721BF2FF9A15" pageId="0" pageNumber="1">
|
||||
<emphasis id="449A4D7D8721300E154FDF93CA288C7F" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
|
||||
CUMZ 4703 (cW 19.4 mm, cH 11.1 mm, dL 13.7 mm; Fig.
|
||||
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shell of Rhiostoma ebenozostera sp. nov. A holotype CUMZ 4703 from Tham Pha Poo, Loei B, C paratypes from type locality B paratype CUMZ 4704 and C paratype CUMZ 3889 D specimen CUMZ 10211 from Lom Kao, Phetchabun E specimen CUMZ 4765 from Suwannakhuha, Nongboa Lumphu F specimen CUMZ 10166 / 2 from Phu Wiang, Khon Kaen." figureDoi="10.3897/zookeys.1142.90097.figure29" httpUri="https://binary.pensoft.net/fig/799887" pageId="0" pageNumber="1">29A</figureCitation>
|
||||
<figureCitation id="2DA2E94CF0538604F2067A0A5494B0F7" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shell of Rhiostoma ebenozostera sp. nov. A holotype CUMZ 4703 from Tham Pha Poo, Loei B, C paratypes from type locality B paratype CUMZ 4704 and C paratype CUMZ 3889 D specimen CUMZ 10211 from Lom Kao, Phetchabun E specimen CUMZ 4765 from Suwannakhuha, Nongboa Lumphu F specimen CUMZ 10166 / 2 from Phu Wiang, Khon Kaen." figureDoi="10.3897/zookeys.1142.90097.figure29" httpUri="https://binary.pensoft.net/fig/799887" pageId="0" pageNumber="1">29A</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
|
||||
<emphasis id="4717199027FE4E29D40AC7988E820B1B" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
|
||||
CUMZ 3887 (228 shells), CUMZ 3889 (53 adults + 25 juveniles; Fig.
|
||||
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shell of Rhiostoma ebenozostera sp. nov. A holotype CUMZ 4703 from Tham Pha Poo, Loei B, C paratypes from type locality B paratype CUMZ 4704 and C paratype CUMZ 3889 D specimen CUMZ 10211 from Lom Kao, Phetchabun E specimen CUMZ 4765 from Suwannakhuha, Nongboa Lumphu F specimen CUMZ 10166 / 2 from Phu Wiang, Khon Kaen." figureDoi="10.3897/zookeys.1142.90097.figure29" httpUri="https://binary.pensoft.net/fig/799887" pageId="0" pageNumber="1">29C</figureCitation>
|
||||
<figureCitation id="987F6BF9E41203BA4B9A6EB4B3F498DF" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shell of Rhiostoma ebenozostera sp. nov. A holotype CUMZ 4703 from Tham Pha Poo, Loei B, C paratypes from type locality B paratype CUMZ 4704 and C paratype CUMZ 3889 D specimen CUMZ 10211 from Lom Kao, Phetchabun E specimen CUMZ 4765 from Suwannakhuha, Nongboa Lumphu F specimen CUMZ 10166 / 2 from Phu Wiang, Khon Kaen." figureDoi="10.3897/zookeys.1142.90097.figure29" httpUri="https://binary.pensoft.net/fig/799887" pageId="0" pageNumber="1">29C</figureCitation>
|
||||
), CUMZ 4704 (2 shells; Figs
|
||||
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shell of Rhiostoma ebenozostera sp. nov. A holotype CUMZ 4703 from Tham Pha Poo, Loei B, C paratypes from type locality B paratype CUMZ 4704 and C paratype CUMZ 3889 D specimen CUMZ 10211 from Lom Kao, Phetchabun E specimen CUMZ 4765 from Suwannakhuha, Nongboa Lumphu F specimen CUMZ 10166 / 2 from Phu Wiang, Khon Kaen." figureDoi="10.3897/zookeys.1142.90097.figure29" httpUri="https://binary.pensoft.net/fig/799887" pageId="0" pageNumber="1">29B</figureCitation>
|
||||
<figureCitation id="101ED285C8AED0FC3C0A6EE60869ED9D" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shell of Rhiostoma ebenozostera sp. nov. A holotype CUMZ 4703 from Tham Pha Poo, Loei B, C paratypes from type locality B paratype CUMZ 4704 and C paratype CUMZ 3889 D specimen CUMZ 10211 from Lom Kao, Phetchabun E specimen CUMZ 4765 from Suwannakhuha, Nongboa Lumphu F specimen CUMZ 10166 / 2 from Phu Wiang, Khon Kaen." figureDoi="10.3897/zookeys.1142.90097.figure29" httpUri="https://binary.pensoft.net/fig/799887" pageId="0" pageNumber="1">29B</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Radula morphology A Rhiostoma housei specimen CUMZ 3982 from Wat Tham Srivilai, Saraburi B Rhiostoma hainesi specimen from Chanthaburi C Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sua, Krabi D Rhiostoma ebenozostera sp. nov. paratype CUMZ 4704 from Tham Pha Poo, Loei E Rhiostoma lannaense sp. nov. paratype CUMZ 4701 from Chiang Dao, Chiang Mai F Rhiostoma tigrina sp. nov. paratype CUMZ 4497 / 2 from Mae Sai, Chiang Rai." figureDoi="10.3897/zookeys.1142.90097.figure32" httpUri="https://binary.pensoft.net/fig/799890" pageId="0" pageNumber="1">32D</figureCitation>
|
||||
<figureCitation id="E004ECD4B1CD1D5F7F226EB2D29B835D" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Radula morphology A Rhiostoma housei specimen CUMZ 3982 from Wat Tham Srivilai, Saraburi B Rhiostoma hainesi specimen from Chanthaburi C Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sua, Krabi D Rhiostoma ebenozostera sp. nov. paratype CUMZ 4704 from Tham Pha Poo, Loei E Rhiostoma lannaense sp. nov. paratype CUMZ 4701 from Chiang Dao, Chiang Mai F Rhiostoma tigrina sp. nov. paratype CUMZ 4497 / 2 from Mae Sai, Chiang Rai." figureDoi="10.3897/zookeys.1142.90097.figure32" httpUri="https://binary.pensoft.net/fig/799890" pageId="0" pageNumber="1">32D</figureCitation>
|
||||
), CUMZ 4766 (99 adults + 24 juveniles), NHMUK 20220439 (5 shells), and SMF 368674 (5 shells). All paratypes are from the type locality.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.1142.90097.figure29" httpUri="https://binary.pensoft.net/fig/799887" pageId="0" pageNumber="1" start="Figure 29" startId="F29">
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 29.</emphasis>
|
||||
<caption id="90048D423AC07D34A5EC726AE2DEDBAB" doi="10.3897/zookeys.1142.90097.figure29" httpUri="https://binary.pensoft.net/fig/799887" pageId="0" pageNumber="1" start="Figure 29" startId="F29">
|
||||
<paragraph id="4318FDB8F72A7B09B37A1FEB281051CB" pageId="0" pageNumber="1">
|
||||
<emphasis id="14F5F43EA44DFA206505F61556374A24" bold="true" pageId="0" pageNumber="1">Figure 29.</emphasis>
|
||||
Shell of
|
||||
<taxonomicName authorityName="Tongkerd & Panha" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma ebenozostera" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ebenozostera">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma ebenozostera</emphasis>
|
||||
<taxonomicName id="CB922899665B04D787C7911F154C64BE" authorityName="Tongkerd & Panha" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma ebenozostera" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ebenozostera">
|
||||
<emphasis id="565E4FBCED72FC7E6FA04B499CFC1BD3" italics="true" pageId="0" pageNumber="1">Rhiostoma ebenozostera</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov.
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||||
<emphasis id="73DD0C242E575A22280512DC14DFA64B" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||||
holotype CUMZ 4703 from Tham Pha Poo, Loei
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">B, C</emphasis>
|
||||
<emphasis id="A4E224905DB6507D840FA5369A3F2308" bold="true" pageId="0" pageNumber="1">B, C</emphasis>
|
||||
paratypes from type locality
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||||
<emphasis id="AF15AEBD90609FEBFF35F962A90D72BB" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||||
paratype CUMZ 4704 and
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||||
<emphasis id="62E23587A199DB69707C52A61E6E55E3" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||||
paratype CUMZ 3889
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||||
<emphasis id="FF61FCD0C630006DE5BDDD05CAF4B374" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||||
specimen CUMZ 10211 from Lom Kao, Phetchabun
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||||
<emphasis id="68B2D940DC721C8D7F8AFD69CE02AAF8" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||||
specimen CUMZ 4765 from Suwannakhuha, Nongboa Lumphu
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||||
<emphasis id="60A5BEE92E2F990828CDA7334FEEB31E" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||||
specimen CUMZ 10166/2 from Phu Wiang, Khon Kaen.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="type locality">
|
||||
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<subSubSection id="F31DE7260737B521B1C703289058B249" pageId="0" pageNumber="1" type="type locality">
|
||||
<paragraph id="35F85F41BAD02B76684478F0BA5765E6" pageId="0" pageNumber="1">Type locality.</paragraph>
|
||||
<paragraph id="F280AAC9B2FF99B1E13459ABB47D7C90" pageId="0" pageNumber="1">
|
||||
Tham Pha Poo, Muang District, Loei Province, Thailand (
|
||||
<geoCoordinate degrees="17" direction="north" minutes="34" orientation="latitude" precision="1" seconds="45.1" value="17.579195">17°34'45.1"N</geoCoordinate>
|
||||
<geoCoordinate id="927C694BE353146C0F09756DEEC0A76D" degrees="17" direction="north" minutes="34" orientation="latitude" precision="1" seconds="45.1" value="17.579195">17°34'45.1"N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="101" direction="east" minutes="42" orientation="longitude" precision="1" seconds="36.2" value="101.71005">101°42'36.2"E</geoCoordinate>
|
||||
<geoCoordinate id="8C59C229291F13ABCA769FF119B74358" degrees="101" direction="east" minutes="42" orientation="longitude" precision="1" seconds="36.2" value="101.71005">101°42'36.2"E</geoCoordinate>
|
||||
). Dry dipterocarp forest on limestone hill.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
|
||||
<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Thailand" location="Loei" specimenCount="1" stateProvince="Loei">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<collectingCountry name="Thailand">Thailand</collectingCountry>
|
||||
<subSubSection id="02807B95D420C37D426931B185A2FD82" pageId="0" pageNumber="1" type="materials_examined">
|
||||
<paragraph id="ECDE1A90CCF71CAEF14307D9AA7B922F" pageId="0" pageNumber="1">Other material examined.</paragraph>
|
||||
<paragraph id="FFE401E5D2174152C22748632C98D0D1" pageId="0" pageNumber="1">
|
||||
<materialsCitation id="7AD63DE9A61662BDC7393396F2BA0898" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Thailand" location="Loei" specimenCount="1" stateProvince="Loei">
|
||||
<emphasis id="28FC2D2680FB8652FA848263A197C4E9" bold="true" pageId="0" pageNumber="1">
|
||||
<collectingCountry id="F85C039B986280AE676777D7B4D02DAA" name="Thailand">Thailand</collectingCountry>
|
||||
</emphasis>
|
||||
: Wat Tham Pha Ban, Chiangkan,
|
||||
<collectingRegion country="Thailand" name="Loei">Loei</collectingRegion>
|
||||
<collectingRegion id="BAE0212DDFC4C89910254D0159574456" country="Thailand" name="Loei">Loei</collectingRegion>
|
||||
: CUMZ 4444. Phu Pha Lom, Muang,
|
||||
<collectingRegion country="Thailand" name="Loei">Loei</collectingRegion>
|
||||
<collectingRegion id="C8477981BD220D9CAAA0D28570B36D08" country="Thailand" name="Loei">Loei</collectingRegion>
|
||||
: CUMZ 10013. Wat Tham Phu Pha Lom, Muang,
|
||||
<collectingRegion country="Thailand" name="Loei">Loei</collectingRegion>
|
||||
<collectingRegion id="2E5C430DA67D14E7DCAB73A8FA9D47FD" country="Thailand" name="Loei">Loei</collectingRegion>
|
||||
: CUMZ 4443. Tham Khuha Waree, Wang Sapung,
|
||||
<collectingRegion country="Thailand" name="Loei">Loei</collectingRegion>
|
||||
<collectingRegion id="31E19824DDEABBDE64554377B6009730" country="Thailand" name="Loei">Loei</collectingRegion>
|
||||
: CUMZ 3925, 3946. Tham Pha Fang, Wang Sapung,
|
||||
<collectingRegion country="Thailand" name="Loei">Loei</collectingRegion>
|
||||
<collectingRegion id="3FE5411FE232A1DBDF120C9FB70E95E8" country="Thailand" name="Loei">Loei</collectingRegion>
|
||||
: CUMZ 10155. Khao Wangpha, Na Wang, Nongboa Lumphu: CUMZ 3935. Tham Suwannakhuha, Na Wang, Nongboa Lumphu: CUMZ 4765 (Fig.
|
||||
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shell of Rhiostoma ebenozostera sp. nov. A holotype CUMZ 4703 from Tham Pha Poo, Loei B, C paratypes from type locality B paratype CUMZ 4704 and C paratype CUMZ 3889 D specimen CUMZ 10211 from Lom Kao, Phetchabun E specimen CUMZ 4765 from Suwannakhuha, Nongboa Lumphu F specimen CUMZ 10166 / 2 from Phu Wiang, Khon Kaen." figureDoi="10.3897/zookeys.1142.90097.figure29" httpUri="https://binary.pensoft.net/fig/799887" pageId="0" pageNumber="1">29E</figureCitation>
|
||||
<figureCitation id="0D97267095296CD52044B457275476D9" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shell of Rhiostoma ebenozostera sp. nov. A holotype CUMZ 4703 from Tham Pha Poo, Loei B, C paratypes from type locality B paratype CUMZ 4704 and C paratype CUMZ 3889 D specimen CUMZ 10211 from Lom Kao, Phetchabun E specimen CUMZ 4765 from Suwannakhuha, Nongboa Lumphu F specimen CUMZ 10166 / 2 from Phu Wiang, Khon Kaen." figureDoi="10.3897/zookeys.1142.90097.figure29" httpUri="https://binary.pensoft.net/fig/799887" pageId="0" pageNumber="1">29E</figureCitation>
|
||||
). Wat Paphaya, Suwannakhuha, Nongboa Lumphu: CUMZ 10031. Wat Suwannakhuha, Ban Nasi, Suwannakhuha, Nongboa Lumphu: CUMZ 3889, 3933, 10021. Wat Phu Poon, Lomkao,
|
||||
<collectingRegion country="Thailand" name="Phetchabun">Phetchabun</collectingRegion>
|
||||
<collectingRegion id="F54F2BE4450BBF63BB478524EEEBF729" country="Thailand" name="Phetchabun">Phetchabun</collectingRegion>
|
||||
: CUMZ 10211 (Fig.
|
||||
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shell of Rhiostoma ebenozostera sp. nov. A holotype CUMZ 4703 from Tham Pha Poo, Loei B, C paratypes from type locality B paratype CUMZ 4704 and C paratype CUMZ 3889 D specimen CUMZ 10211 from Lom Kao, Phetchabun E specimen CUMZ 4765 from Suwannakhuha, Nongboa Lumphu F specimen CUMZ 10166 / 2 from Phu Wiang, Khon Kaen." figureDoi="10.3897/zookeys.1142.90097.figure29" httpUri="https://binary.pensoft.net/fig/799887" pageId="0" pageNumber="1">29D</figureCitation>
|
||||
<figureCitation id="DC7F7B495027AB50FDBB5C827223ABD7" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shell of Rhiostoma ebenozostera sp. nov. A holotype CUMZ 4703 from Tham Pha Poo, Loei B, C paratypes from type locality B paratype CUMZ 4704 and C paratype CUMZ 3889 D specimen CUMZ 10211 from Lom Kao, Phetchabun E specimen CUMZ 4765 from Suwannakhuha, Nongboa Lumphu F specimen CUMZ 10166 / 2 from Phu Wiang, Khon Kaen." figureDoi="10.3897/zookeys.1142.90097.figure29" httpUri="https://binary.pensoft.net/fig/799887" pageId="0" pageNumber="1">29D</figureCitation>
|
||||
). Tham Poo Loup, Chum Phae,
|
||||
<collectingRegion country="Thailand" name="Khon Kaen">Khon Kaen</collectingRegion>
|
||||
<collectingRegion id="63EAEDF3823EEB497A642452838776AF" country="Thailand" name="Khon Kaen">Khon Kaen</collectingRegion>
|
||||
: CUMZ 3931. Tham Nennoi, Chum Phae,
|
||||
<collectingRegion country="Thailand" name="Khon Kaen">Khon Kaen</collectingRegion>
|
||||
<collectingRegion id="33A635A9893F14E2CC73EF1AF04ECC50" country="Thailand" name="Khon Kaen">Khon Kaen</collectingRegion>
|
||||
: CUMZ 10015. Tham Phayanakarat, Chum Phae,
|
||||
<collectingRegion country="Thailand" name="Khon Kaen">Khon Kaen</collectingRegion>
|
||||
<collectingRegion id="951E487A8B0EF5B588421C5E8C9F40C5" country="Thailand" name="Khon Kaen">Khon Kaen</collectingRegion>
|
||||
: CUMZ 10025. Phu Wiang, Wiang Kao,
|
||||
<collectingRegion country="Thailand" name="Khon Kaen">Khon Kaen</collectingRegion>
|
||||
<collectingRegion id="D89FCD514676204A057F67D401443A27" country="Thailand" name="Khon Kaen">Khon Kaen</collectingRegion>
|
||||
: CUMZ 10166/1, 10166/2 (Fig.
|
||||
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shell of Rhiostoma ebenozostera sp. nov. A holotype CUMZ 4703 from Tham Pha Poo, Loei B, C paratypes from type locality B paratype CUMZ 4704 and C paratype CUMZ 3889 D specimen CUMZ 10211 from Lom Kao, Phetchabun E specimen CUMZ 4765 from Suwannakhuha, Nongboa Lumphu F specimen CUMZ 10166 / 2 from Phu Wiang, Khon Kaen." figureDoi="10.3897/zookeys.1142.90097.figure29" httpUri="https://binary.pensoft.net/fig/799887" pageId="0" pageNumber="1">29F</figureCitation>
|
||||
<figureCitation id="19CBE8FC8AD34941E9033B28DF6D5E05" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shell of Rhiostoma ebenozostera sp. nov. A holotype CUMZ 4703 from Tham Pha Poo, Loei B, C paratypes from type locality B paratype CUMZ 4704 and C paratype CUMZ 3889 D specimen CUMZ 10211 from Lom Kao, Phetchabun E specimen CUMZ 4765 from Suwannakhuha, Nongboa Lumphu F specimen CUMZ 10166 / 2 from Phu Wiang, Khon Kaen." figureDoi="10.3897/zookeys.1142.90097.figure29" httpUri="https://binary.pensoft.net/fig/799887" pageId="0" pageNumber="1">29F</figureCitation>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">Shell medium, thick and depressed. Detached whorl medium in length, curved, and slightly descending. Breathing device tubular. Broad dark brown spiral band on periphery present.</paragraph>
|
||||
<subSubSection id="0B9D5686352FE24CD727E46650E061E9" pageId="0" pageNumber="1" type="diagnosis">
|
||||
<paragraph id="34449B73D9F2509EADCC7C9E28A54DE7" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||||
<paragraph id="947BE446537551C5381E9FAF9B51FF72" pageId="0" pageNumber="1">Shell medium, thick and depressed. Detached whorl medium in length, curved, and slightly descending. Breathing device tubular. Broad dark brown spiral band on periphery present.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
|
||||
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<taxonomicName authorityName="Tongkerd & Panha" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma ebenozostera" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ebenozostera">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma ebenozostera</emphasis>
|
||||
<subSubSection id="46C03E423FBC34631E498076B0780907" pageId="0" pageNumber="1" type="differential diagnosis">
|
||||
<paragraph id="631175B4E9C9406040BCC354B728EB38" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
|
||||
<paragraph id="0089FB114E725696E2EB73AEA7A73A5E" pageId="0" pageNumber="1">
|
||||
<taxonomicName id="0629FAD006DD005959C6A26F2ECE038F" authorityName="Tongkerd & Panha" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma ebenozostera" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ebenozostera">
|
||||
<emphasis id="98A4F4859B5A38132376B9A2E838B9A4" italics="true" pageId="0" pageNumber="1">Rhiostoma ebenozostera</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. can be distinguished from all other recognised
|
||||
<taxonomicName authorityName="Tongkerd & Tumpeesuwan & Inkhavilay & Prasankok & Jeratthitikul & Panha & Sutcharit" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma</emphasis>
|
||||
<taxonomicName id="148B0EBEBCD1702E48ECAA908748D77E" authorityName="Tongkerd & Tumpeesuwan & Inkhavilay & Prasankok & Jeratthitikul & Panha & Sutcharit" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="2705A208EC88BB42FF84008F392E3B2D" italics="true" pageId="0" pageNumber="1">Rhiostoma</emphasis>
|
||||
</taxonomicName>
|
||||
species by having a prominent wide dark brown to black peripheral band. This colour pattern persists in all populations recognised for this species. Regardless of the peripheral band, this new species can be distinguished from
|
||||
<taxonomicName lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
|
||||
<taxonomicName id="26DEC9A727B8F1CA65E6AC1D539E39F3" lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
|
||||
<emphasis id="ABEE7A227497B7736336955C3880A479" italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
|
||||
</taxonomicName>
|
||||
in having straight and slender tubular breathing device, and relatively longer detached whorl. In contrast,
|
||||
<taxonomicName lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
|
||||
<taxonomicName id="C0ADB3F4B67FACFD20368DDC09FC2F0D" lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
|
||||
<emphasis id="2C6CF17DA007E9B69F782C90110D060F" italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
|
||||
</taxonomicName>
|
||||
has a stout and curved tubular breathing device and relatively shorter detached whorl.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="description">
|
||||
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Shell</emphasis>
|
||||
<subSubSection id="61BFAF7AF8E6963BF6428729FD1AAF82" pageId="0" pageNumber="1" type="description">
|
||||
<paragraph id="1F290F0AC91F776EB72B8EC044B291CA" pageId="0" pageNumber="1">Description.</paragraph>
|
||||
<paragraph id="20FB8AD1A4EB123779D29765A6EFD1B0" pageId="0" pageNumber="1">
|
||||
<emphasis id="37AFB6C1DD748C1374E537B3BE185C29" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="C8D2F0296B73A8A0130C5AEFBE770CD1" italics="true" pageId="0" pageNumber="1">Shell</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Shell medium, cW 17.7-22.6 mm, cH 9.4-12.1 mm, thick, and sub-discoidal to discoidal shape; detached-whorl length 8.5-14.5 mm. Apex acute; spire nearly flat to elevated. Whorls 4 to 5, convex, increasing regularly; suture wide and shallow; last whorl rounded and stout. Detached whorl medium in length, approximately the same length as apertural width, curved and descending. Shell surface with fine growth lines. Periostracum thin, corneous, and transparent. Shell colour uniformly brownish or with pattern of dark brown blotches, and faded on ventral shell surface; dark spiral band on periphery always present and broad. Peristome circular and double; lip thickened, expanded and multi-layered. Aperture opened sub-ventrally. Breathing device tubular, perpendicular to detached whorl and its tip attached to preceding whorl; outer lip forming long and closed tube and located just behind apertural lip; inner lip with deep incision or small hole inside aperture. Umbilicus widely opened and deep. Operculum calcareous, cup-shaped, and multispiral (Fig.
|
||||
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shell of Rhiostoma ebenozostera sp. nov. A holotype CUMZ 4703 from Tham Pha Poo, Loei B, C paratypes from type locality B paratype CUMZ 4704 and C paratype CUMZ 3889 D specimen CUMZ 10211 from Lom Kao, Phetchabun E specimen CUMZ 4765 from Suwannakhuha, Nongboa Lumphu F specimen CUMZ 10166 / 2 from Phu Wiang, Khon Kaen." figureDoi="10.3897/zookeys.1142.90097.figure29" httpUri="https://binary.pensoft.net/fig/799887" pageId="0" pageNumber="1">29</figureCitation>
|
||||
<figureCitation id="D94BDD72E558105D3ABF71996C1A5FBE" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shell of Rhiostoma ebenozostera sp. nov. A holotype CUMZ 4703 from Tham Pha Poo, Loei B, C paratypes from type locality B paratype CUMZ 4704 and C paratype CUMZ 3889 D specimen CUMZ 10211 from Lom Kao, Phetchabun E specimen CUMZ 4765 from Suwannakhuha, Nongboa Lumphu F specimen CUMZ 10166 / 2 from Phu Wiang, Khon Kaen." figureDoi="10.3897/zookeys.1142.90097.figure29" httpUri="https://binary.pensoft.net/fig/799887" pageId="0" pageNumber="1">29</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Radula</emphasis>
|
||||
<paragraph id="5BEB58AF8B52A92D06C2E4A5D0FAB408" pageId="0" pageNumber="1">
|
||||
<emphasis id="E987EE1F4537E051E6A92C33FEA90F4B" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="AE3460F21EB350C5E90F87AE9AFC51C9" italics="true" pageId="0" pageNumber="1">Radula</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Teeth arrangement and shape are similar to those of
|
||||
<taxonomicName lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
|
||||
<taxonomicName id="F4447CE5D3BB677904EC9E5356FBA47D" lsidName="R. housei" pageId="0" pageNumber="1" rank="species" species="housei">
|
||||
<emphasis id="854AD9F3463EE422C34C5A92B36671CC" italics="true" pageId="0" pageNumber="1">R. housei</emphasis>
|
||||
</taxonomicName>
|
||||
. Central tooth with large central cup and two lateral cusps on each side. Lateral teeth composed of three cusps; central cusp large, with convex tip, and flanked by small pointed tips of one outer cusp and one inner cusp. Marginal teeth each consisting of three pointed cusps (Fig.
|
||||
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Radula morphology A Rhiostoma housei specimen CUMZ 3982 from Wat Tham Srivilai, Saraburi B Rhiostoma hainesi specimen from Chanthaburi C Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sua, Krabi D Rhiostoma ebenozostera sp. nov. paratype CUMZ 4704 from Tham Pha Poo, Loei E Rhiostoma lannaense sp. nov. paratype CUMZ 4701 from Chiang Dao, Chiang Mai F Rhiostoma tigrina sp. nov. paratype CUMZ 4497 / 2 from Mae Sai, Chiang Rai." figureDoi="10.3897/zookeys.1142.90097.figure32" httpUri="https://binary.pensoft.net/fig/799890" pageId="0" pageNumber="1">32D</figureCitation>
|
||||
<figureCitation id="F4A0D0728D0401932BB49BA5C49B1E2E" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Radula morphology A Rhiostoma housei specimen CUMZ 3982 from Wat Tham Srivilai, Saraburi B Rhiostoma hainesi specimen from Chanthaburi C Rhiostoma jalorensis specimen CUMZ 3994 from Wat Tham Sua, Krabi D Rhiostoma ebenozostera sp. nov. paratype CUMZ 4704 from Tham Pha Poo, Loei E Rhiostoma lannaense sp. nov. paratype CUMZ 4701 from Chiang Dao, Chiang Mai F Rhiostoma tigrina sp. nov. paratype CUMZ 4497 / 2 from Mae Sai, Chiang Rai." figureDoi="10.3897/zookeys.1142.90097.figure32" httpUri="https://binary.pensoft.net/fig/799890" pageId="0" pageNumber="1">32D</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
||||
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<subSubSection id="26B157AA4AB22880E1C770572E9502E8" pageId="0" pageNumber="1" type="etymology">
|
||||
<paragraph id="30A9AE0AB403CFA71D29C2912B0DFB05" pageId="0" pageNumber="1">Etymology.</paragraph>
|
||||
<paragraph id="600211A65D9C7126CCBE9D9D003966D3" pageId="0" pageNumber="1">
|
||||
The specific name
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">ebenozostera</emphasis>
|
||||
<emphasis id="990A6BCD42D40C73847C46194D7FA77E" italics="true" pageId="0" pageNumber="1">ebenozostera</emphasis>
|
||||
is derived from the two Greek words
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">ebenos</emphasis>
|
||||
<emphasis id="C6C690E864D6C3B8291E6C0BA691C24D" italics="true" pageId="0" pageNumber="1">ebenos</emphasis>
|
||||
meaning black colour, and
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">zoste</emphasis>
|
||||
<emphasis id="0AF08306D013041D2BFBA29B9BE73DD6" italics="true" pageId="0" pageNumber="1">zoste</emphasis>
|
||||
meaning girdle or belt. This name refers to the prominence of a dark, wide, peripheral band, which is the distinguishing character of this species.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<subSubSection id="B4A9486CC7475AF5039A2A7253104FBF" pageId="0" pageNumber="1" type="distribution">
|
||||
<paragraph id="928EF8B0ABA0ACDABFF400FCED88F8C6" pageId="0" pageNumber="1">Distribution.</paragraph>
|
||||
<paragraph id="4B499C55D9E73F4AD37573F86110DC25" pageId="0" pageNumber="1">
|
||||
This new species has narrow distribution; it is known from several limestone outcrops in dry deciduous forests of Loei, Nong Bua Lam Phu, and Khonkaen provinces (Fig.
|
||||
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution map of Rhiostoma housei - species group." figureDoi="10.3897/zookeys.1142.90097.figure18" httpUri="https://binary.pensoft.net/fig/799876" pageId="0" pageNumber="1">18</figureCitation>
|
||||
<figureCitation id="05C55BF11CAAE8446C302E2710A8769C" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution map of Rhiostoma housei - species group." figureDoi="10.3897/zookeys.1142.90097.figure18" httpUri="https://binary.pensoft.net/fig/799876" pageId="0" pageNumber="1">18</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
||||
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<subSubSection id="D09C453283C9C2FB02D3CEFC69B1D89E" pageId="0" pageNumber="1" type="remarks">
|
||||
<paragraph id="E6799D7169F2FB9D1756A49904D52E83" pageId="0" pageNumber="1">Remarks.</paragraph>
|
||||
<paragraph id="236F529630DE9FC26D6D0C8B4EBD30AD" pageId="0" pageNumber="1">
|
||||
The specimens from Nong Bua Lam Phu possess a stout shell, slightly shorter detached whorl, and paler shell colour (Fig.
|
||||
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shell of Rhiostoma ebenozostera sp. nov. A holotype CUMZ 4703 from Tham Pha Poo, Loei B, C paratypes from type locality B paratype CUMZ 4704 and C paratype CUMZ 3889 D specimen CUMZ 10211 from Lom Kao, Phetchabun E specimen CUMZ 4765 from Suwannakhuha, Nongboa Lumphu F specimen CUMZ 10166 / 2 from Phu Wiang, Khon Kaen." figureDoi="10.3897/zookeys.1142.90097.figure29" httpUri="https://binary.pensoft.net/fig/799887" pageId="0" pageNumber="1">29E</figureCitation>
|
||||
<figureCitation id="BD889C79C55CE4097D35550188AB0D03" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shell of Rhiostoma ebenozostera sp. nov. A holotype CUMZ 4703 from Tham Pha Poo, Loei B, C paratypes from type locality B paratype CUMZ 4704 and C paratype CUMZ 3889 D specimen CUMZ 10211 from Lom Kao, Phetchabun E specimen CUMZ 4765 from Suwannakhuha, Nongboa Lumphu F specimen CUMZ 10166 / 2 from Phu Wiang, Khon Kaen." figureDoi="10.3897/zookeys.1142.90097.figure29" httpUri="https://binary.pensoft.net/fig/799887" pageId="0" pageNumber="1">29E</figureCitation>
|
||||
) than the typical population. However, a prominent broad, dark, peripheral band suggests that they are conspecific.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
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