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<document id="390FE30792431CAA763AD6C8313FE4FD" ID-DOI="10.1093/zoolinnean/zlae167" ID-ISSN="0024-4082" ID-ZooBank="3C07EBC-1845-4356-B182-E801A3BDC7EC" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="operationResults" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1738158306083" checkinUser="plazi" docAuthor="Gu, Xinyu, Chen, Chong, Gao, Kexin, Zhou, Yadong & Sun, Jin" docDate="2025" docId="03946B0DFFCB472724A1FCC57182FC5A" docLanguage="en" docName="zlae167.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 203 (1)" docSource="https://doi.org/10.1093/zoolinnean/zlae167" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.10:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="10" docTitle="Shinkailepas cornuthauma Gu, Chen, Gao, Zhou & Sun, 2025, sp. nov." docType="treatment" docVersion="2" lastPageNumber="12" masterDocId="FFAD1375FFCC472C2708FFBD717BFFAF" masterDocTitle="Integrative taxonomy of new neritimorph limpets from Indian Ocean deep-sea hot vents shed light on their biogeographic history" masterLastPageNumber="15" masterPageNumber="1" pageNumber="8" updateTime="1738921431350" updateUser="carolina">
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<mods:titleInfo id="FE4DF0D9B7843E13C20454E0738717A8">
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<mods:title id="7CA215944DD18E9870D6927D48864B7A">Integrative taxonomy of new neritimorph limpets from Indian Ocean deep-sea hot vents shed light on their biogeographic history</mods:title>
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<mods:roleTerm id="25EF338DF7CD19125D590DCCC8655EEA">Author</mods:roleTerm>
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<mods:namePart id="1F3BCCAB97F7CD748F6335B22E576DD2">Gu, Xinyu</mods:namePart>
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<mods:affiliation id="6B3346053C0C956E67CB8EE56AD973A8">Key Laboratory of Evolution & Marine Biodiversity (Ministry of Education) and Institute of Evolution & Marine Biodiversity, Ocean University of China, Qingdao 266003, China & Laboratory for Marine Biology and Biotechnology, Qingdao Marine Science and Technology Center, Laoshan Laboratory, Qingdao 266237, China & Academy of the Future Ocean, Ocean University of China, Qingdao 266100, China</mods:affiliation>
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<mods:roleTerm id="6E729B17B0188760D5BFF9A34BF34531">Author</mods:roleTerm>
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<mods:namePart id="E8D61C67A56988C7067DD57C74E12A78">Chen, Chong</mods:namePart>
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<mods:affiliation id="ACFABA457D82A8E272DCB7A0614D2839">X-STAR, Japan Agency for Marine-Earth Science and Technology (JAMSTEC), 2 - 15 Natsushima-cho, Yokosuka, Kanagawa 237 - 0061, Japan</mods:affiliation>
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<mods:namePart id="303CE47DD4E0A4B589A05E4ECF3E6E65">Gao, Kexin</mods:namePart>
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<mods:affiliation id="21D901A646A99743D794BF1877F6336E">Key Laboratory of Evolution & Marine Biodiversity (Ministry of Education) and Institute of Evolution & Marine Biodiversity, Ocean University of China, Qingdao 266003, China & Laboratory for Marine Biology and Biotechnology, Qingdao Marine Science and Technology Center, Laoshan Laboratory, Qingdao 266237, China</mods:affiliation>
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<mods:namePart id="39E1792B8473D01B58458C98F7AA119E">Zhou, Yadong</mods:namePart>
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<mods:affiliation id="FAAB637E82244D705B12150A46639CA5">Key Laboratory of Marine Ecosystem Dynamics, Second Institute of Oceanography, Ministry of Natural Resources, Hangzhou 310012, China & Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai), Zhuhai 519082, China</mods:affiliation>
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<mods:nameIdentifier id="F73D2F237221F49A6BDFFCB59D43D7E1" type="email">jin_sun@ouc.edu.cn</mods:nameIdentifier>
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<mods:namePart id="30D892D5CFEC81C74B719D8CF5E9898F">Sun, Jin</mods:namePart>
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<mods:affiliation id="702DBA6CADD565F92AFB6599987807A5">Key Laboratory of Evolution & Marine Biodiversity (Ministry of Education) and Institute of Evolution & Marine Biodiversity, Ocean University of China, Qingdao 266003, China & Laboratory for Marine Biology and Biotechnology, Qingdao Marine Science and Technology Center, Laoshan Laboratory, Qingdao 266237, China & Academy of the Future Ocean, Ocean University of China, Qingdao 266100, China</mods:affiliation>
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<mods:nameIdentifier id="E943730C5096A336E6BE5BC341CEE1E2" type="email">jin_sun@ouc.edu.cn</mods:nameIdentifier>
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<mods:title id="533A2173958C4861E240541CE05FC095">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="1D1D3192307CDD637CF4A6FA38A5D13C">2025</mods:date>
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<mods:title id="E0F396EEC9A0794A1A3DEA0C7E6D3DE9">Zool. J. Linn. Soc.</mods:title>
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<mods:number id="DD8831C810DF3AEB3CC3C6C36844EFE2">2025-01-02</mods:number>
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<mods:url id="3E80EE408899730217BF9C4490A6B23C">https://doi.org/10.1093/zoolinnean/zlae167</mods:url>
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<mods:identifier id="18FD0334C5FC16E94887F164D0984E11" type="DOI">10.1093/zoolinnean/zlae167</mods:identifier>
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<mods:identifier id="4D8A7647707104E4C7E148B6E35C6F33" type="ISSN">0024-4082</mods:identifier>
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<mods:identifier id="38D00F803B7EA24C25200038B851BF02" type="ZooBank">3C07EBC-1845-4356-B182-E801A3BDC7EC</mods:identifier>
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<treatment id="03946B0DFFCB472724A1FCC57182FC5A" LSID="urn:lsid:plazi:treatment:03946B0DFFCB472724A1FCC57182FC5A" httpUri="http://treatment.plazi.org/id/03946B0DFFCB472724A1FCC57182FC5A" lastPageId="11" lastPageNumber="12" pageId="7" pageNumber="8">
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<subSubSection id="C3278990FFCB472B24A1FCC5744FFC3C" box="[937,1332,888,915]" pageId="7" pageNumber="8" type="nomenclature">
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<paragraph id="8B82DA1BFFCB472B24A1FCC5744FFC3C" blockId="7.[937,1332,888,915]" box="[937,1332,888,915]" pageId="7" pageNumber="8">
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<heading id="D0CA6D77FFCB472B24A1FCC5744FFC3C" bold="true" box="[937,1332,888,915]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="8" reason="2">
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<emphasis id="B9490609FFCB472B24A1FCC5744FFC3C" bold="true" box="[937,1332,888,915]" pageId="7" pageNumber="8">
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<taxonomicName id="4C3DA198FFCB472B24A1FCC575ACFC3C" authorityName="Gu & Chen & Gao & Zhou & Sun" authorityYear="2025" box="[937,1239,888,915]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="cornuthauma" status="sp. nov.">
|
||||
<emphasis id="B9490609FFCB472B24A1FCC575ACFC3C" bold="true" box="[937,1239,888,915]" italics="true" pageId="7" pageNumber="8">Shinkailepas cornuthauma</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27ABB72FFCB472B23D5FCC4744FFC3C" box="[1245,1332,889,915]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3278990FFCB472B233FFC0A75DDFC7E" box="[1079,1190,951,978]" pageId="7" pageNumber="8" type="description">
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<paragraph id="8B82DA1BFFCB472B233FFC0A75DDFC7E" blockId="7.[809,1461,951,1077]" box="[1079,1190,951,978]" pageId="7" pageNumber="8">
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||||
<emphasis id="B9490609FFCB472B233FFC0A75DDFC7E" bold="true" box="[1079,1190,951,978]" pageId="7" pageNumber="8">
|
||||
(
|
||||
<figureCitation id="1306C69EFFCB472B2349FC0A75E7FC7D" box="[1089,1180,951,978]" captionStart-0="Figure 5" captionStart-1="Figure 6" captionStart-2="Figure 7" captionStartId-0="8.[129,194,1562,1586]" captionStartId-1="9.[113,178,1683,1707]" captionStartId-2="10.[129,194,1040,1064]" captionTargetBox-0="[129,1473,144,1535]" captionTargetBox-1="[114,1458,144,1655]" captionTargetBox-2="[129,1473,144,1012]" captionTargetId-0="figure-187@8.[129,1473,144,1535]" captionTargetId-1="figure-69@9.[114,1458,144,1655]" captionTargetId-2="figure-475@10.[129,1473,144,1012]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="Figure 5. Shinkailepas cornuthauma, specimens examined in this study. A, specimen collected from Wocan field (RSIO 38229), female. B, holotype (RSIO 38227), male. White arrow indicates the penis. C, paratype 1 (RSIO 38228), female, shell damaged during collection." captionText-1="Figure 6. Shinkailepas cornuthauma, external anatomy of the holotype male (RSIO 38227). A, dorsal aspect with shell removed.B, enlarged dorsal view of the head with mantle edge removed, female. C, enlarged ventral of the head.D, operculum shown attached to dorsal aspect of foot.Abbreviations:ang, annex gland; cl, cephalic lappet; ct, ctenidium; dg, digestive gland; e, eye; f, foot; lsm, left shell muscle; m, mouth; me, mantle edge; nl, neck lobe; ol, oral lappet; pe, penis; pc, pericardium; pr, prostate; rsm, right shell muscle; sn, snout; st, stomach; t, cephalic tentacle; te, testis." captionText-2="Figure 7. Shinkailepas cornuthauma, scanning electron micrographs of the holotype (RSIO 38227). A, shell microstructure. B, overview of the radula. C, close-up of the left half of the radula. D, close-up of the outer laterals and marginal teeth." pageId="7" pageNumber="8">Figs 5–7</figureCitation>
|
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)
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</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3278990FFCB472B2422FC6372F8FB9A" pageId="7" pageNumber="8" type="reference_group">
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||||
<paragraph id="8B82DA1BFFCB472B2422FC6372F8FB9A" blockId="7.[809,1461,951,1077]" pageId="7" pageNumber="8">
|
||||
<taxonomicName id="4C3DA198FFCB472B2422FC6372B3FC5A" box="[810,968,990,1014]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="undetermined">
|
||||
<emphasis id="B9490609FFCB472B2422FC6372DEFC59" box="[810,933,990,1014]" italics="true" pageId="7" pageNumber="8">Shinkailepas</emphasis>
|
||||
sp.
|
||||
</taxonomicName>
|
||||
<bibRefCitation id="EFACA7EAFFCB472B24D8FC6375F4FC59" author="Zhou Y & Chen C & Zhang D" box="[976,1167,990,1014]" pageId="7" pageNumber="8" pagination="2858 - 70" refId="ref12257" refString="Zhou Y, Chen C, Zhang D et al. Delineating biogeographic regions in Indian Ocean deep-sea vents and implications for conservation. Diversity and Distributions 2022; 28: 2858 - 70. https: // doi. org / 10.1111 / ddi. 13535" type="journal article" year="2022">
|
||||
Zhou
|
||||
<emphasis id="B9490609FFCB472B2306FC627545FC59" box="[1038,1086,990,1014]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
|
||||
(2022)
|
||||
</bibRefCitation>
|
||||
: supplementary table S5 (at least the sequenced individual from the Daxi field belongs to this species).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3278990FFCB472B2422FBEB75DFFB22" pageId="7" pageNumber="8" type="description">
|
||||
<paragraph id="8B82DA1BFFCB472B2422FBEB75DFFB22" blockId="7.[810,1459,1110,1165]" pageId="7" pageNumber="8">
|
||||
<emphasis id="B9490609FFCB472B2422FBEB7568FBC1" box="[810,1043,1110,1134]" italics="true" pageId="7" pageNumber="8">ZooBank registration:</emphasis>
|
||||
<uri id="FFACD619FFCB472B2327FBEB75DFFB22" pageId="7" pageNumber="8">
|
||||
urn:lsid:zoobank.org:act:
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<uuid id="FF9BE0CEFFCB472B2225FBEB75DFFB22" pageId="7" pageNumber="8">BD73673D- FE2F-449C-82CE-DA2F6A4B989D</uuid>
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</uri>
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</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3278990FFCB472B2422FB1275AAFA0D" pageId="7" pageNumber="8" type="diagnosis">
|
||||
<paragraph id="8B82DA1BFFCB472B2422FB1275AAFA0D" blockId="7.[810,1460,1199,1442]" pageId="7" pageNumber="8">
|
||||
<emphasis id="B9490609FFCB472B2422FB1272EBFB69" box="[810,912,1199,1222]" italics="true" pageId="7" pageNumber="8">Diagnosis:</emphasis>
|
||||
Medium-sized
|
||||
<taxonomicName id="4C3DA198FFCB472B233FFB1275C9FB68" box="[1079,1202,1199,1223]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B9490609FFCB472B233FFB1275C9FB68" box="[1079,1202,1199,1223]" italics="true" pageId="7" pageNumber="8">Shinkailepas</emphasis>
|
||||
</taxonomicName>
|
||||
(SL ≤
|
||||
<quantity id="4CC577FEFFCB472B23F7FB127432FB68" box="[1279,1353,1199,1223]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="8" unit="mm" value="12.0">12 mm</quantity>
|
||||
) with the apex weakly inclined to the right and positioned at approximately 70% from the anterior shell edge. Shell sculpture weakly cancellate with radial ribs much stronger than concentric ribs, raised nodes are formed where the two directions meet. Septum (‘deck’) reduced and minute. Epipodial fold carries 15–18 small triangular tentacles. Operculum entirely corneous. Diameter of opercular nucleus approximately 500 μm.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3278990FFCB472B2422FA7A759EF819" pageId="7" pageNumber="8" type="materials_examined">
|
||||
<paragraph id="8B82DA1BFFCB472B2422FA7A74F7FA70" blockId="7.[810,1420,1479,1503]" box="[810,1420,1479,1503]" pageId="7" pageNumber="8">
|
||||
<emphasis id="B9490609FFCB472B2422FA7A72D7FA70" box="[810,940,1479,1503]" italics="true" pageId="7" pageNumber="8">
|
||||
<typeStatus id="548664B9FFCB472B2422FA7A7221FA71" box="[810,858,1479,1502]" pageId="7" pageNumber="8">Type</typeStatus>
|
||||
locality:
|
||||
</emphasis>
|
||||
Daxi hydrothermal vent field, Carlsberg Ridge.
|
||||
</paragraph>
|
||||
<paragraph id="8B82DA1BFFCB472B2422F9B47457F953" blockId="7.[810,1459,1544,1788]" pageId="7" pageNumber="8">
|
||||
<emphasis id="B9490609FFCB472B2422F9B472C6F98E" box="[810,957,1545,1569]" italics="true" pageId="7" pageNumber="8">Type material:</emphasis>
|
||||
<materialsCitation id="3B55D046FFCB472B24C0F9B47435F931" collectingDate="2017-03" collectionCode="R, V" elevation="3480" latitude="6.8" longLatPrecision="129" longitude="60.18" pageId="7" pageNumber="8" specimenCode="RSIO 38227" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||||
<typeStatus id="548664B9FFCB472B24C0F9B47550F98E" box="[968,1067,1545,1569]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
|
||||
(
|
||||
<specimenCode id="DB9B7260FFCB472B2335F9B475B1F98E" box="[1085,1226,1545,1569]" collectionCode="RSIO" pageId="7" pageNumber="8">RSIO 38227</specimenCode>
|
||||
;
|
||||
<figureCitation id="1306C69EFFCB472B23D1F9B47452F98F" box="[1241,1321,1544,1569]" captionStart="Figure 5" captionStartId="8.[129,194,1562,1586]" captionTargetBox="[129,1473,144,1535]" captionTargetId="figure-187@8.[129,1473,144,1535]" captionTargetPageId="8" captionText="Figure 5. Shinkailepas cornuthauma, specimens examined in this study. A, specimen collected from Wocan field (RSIO 38229), female. B, holotype (RSIO 38227), male. White arrow indicates the penis. C, paratype 1 (RSIO 38228), female, shell damaged during collection." pageId="7" pageNumber="8">Figs 5B</figureCitation>
|
||||
,
|
||||
<figureCitation id="1306C69EFFCB472B2232F9B4741EF98E" box="[1338,1381,1545,1569]" captionStart-0="Figure 6" captionStart-1="Figure 7" captionStartId-0="9.[113,178,1683,1707]" captionStartId-1="10.[129,194,1040,1064]" captionTargetBox-0="[114,1458,144,1655]" captionTargetBox-1="[129,1473,144,1012]" captionTargetId-0="figure-69@9.[114,1458,144,1655]" captionTargetId-1="figure-475@10.[129,1473,144,1012]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="Figure 6. Shinkailepas cornuthauma, external anatomy of the holotype male (RSIO 38227). A, dorsal aspect with shell removed.B, enlarged dorsal view of the head with mantle edge removed, female. C, enlarged ventral of the head.D, operculum shown attached to dorsal aspect of foot.Abbreviations:ang, annex gland; cl, cephalic lappet; ct, ctenidium; dg, digestive gland; e, eye; f, foot; lsm, left shell muscle; m, mouth; me, mantle edge; nl, neck lobe; ol, oral lappet; pe, penis; pc, pericardium; pr, prostate; rsm, right shell muscle; sn, snout; st, stomach; t, cephalic tentacle; te, testis." captionText-1="Figure 7. Shinkailepas cornuthauma, scanning electron micrographs of the holotype (RSIO 38227). A, shell microstructure. B, overview of the radula. C, close-up of the left half of the radula. D, close-up of the outer laterals and marginal teeth." pageId="7" pageNumber="8">6–7</figureCitation>
|
||||
),
|
||||
<specimenCount id="9D3B1192FFCB472B2274F9B474D4F98E" box="[1404,1455,1545,1569]" pageId="7" pageNumber="8" type="male">male</specimenCount>
|
||||
, 95% ethanol, SL
|
||||
<quantity id="4CC577FEFFCB472B24D7F9957547F990" box="[991,1084,1576,1600]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="7" pageNumber="8" unit="mm" value="11.7">11.7 mm</quantity>
|
||||
, SW
|
||||
<quantity id="4CC577FEFFCB472B237EF99575BDF990" box="[1142,1222,1576,1600]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="7" pageNumber="8" unit="mm" value="8.4">8.4 mm</quantity>
|
||||
, male; Daxi vent field, Carlsberg Ridge,
|
||||
<geoCoordinate id="EE09BCDCFFCB472B24E6F9FA7528F9F0" box="[1006,1107,1606,1631]" degrees="60" direction="east" minutes="10.8" orientation="longitude" pageId="7" pageNumber="8" precision="92" value="60.18">
|
||||
60°10.8
|
||||
<emphasis id="B9490609FFCB472B2335F9FB7538F9F0" box="[1085,1091,1606,1631]" italics="true" pageId="7" pageNumber="8">ʹ</emphasis>
|
||||
E
|
||||
</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE09BCDCFFCB472B236FF9FA75B8F9F0" box="[1127,1219,1606,1631]" degrees="6" direction="north" minutes="48.0" orientation="latitude" pageId="7" pageNumber="8" precision="92" value="6.8">
|
||||
6°48.0
|
||||
<emphasis id="B9490609FFCB472B23A1F9FB75D4F9F0" box="[1193,1199,1606,1631]" italics="true" pageId="7" pageNumber="8">ʹ</emphasis>
|
||||
N
|
||||
</geoCoordinate>
|
||||
,
|
||||
<quantity id="4CC577FEFFCB472B23DFF9FA7454F9F0" box="[1239,1327,1607,1631]" metricMagnitude="3" metricUnit="m" metricValue="3.4799999999999995" pageId="7" pageNumber="8" unit="m" value="3480.0">
|
||||
<elevation id="00103D28FFCB472B23DFF9FA7454F9F0" box="[1239,1327,1607,1631]" metricMagnitude="3" metricUnit="m" metricValue="3.4799999999999995" pageId="7" pageNumber="8" unit="m" value="3480.0">3480 m</elevation>
|
||||
</quantity>
|
||||
deep, R/V
|
||||
<emphasis id="B9490609FFCB472B2422F9DA72A8F9D0" box="[810,979,1639,1663]" italics="true" pageId="7" pageNumber="8">Xiangyanghong 9</emphasis>
|
||||
cruise DY38, collected by a seven-function manipulator of HOV
|
||||
<emphasis id="B9490609FFCB472B24E5F93B7540F931" box="[1005,1083,1670,1694]" italics="true" pageId="7" pageNumber="8">Jiaolong</emphasis>
|
||||
on dive 126,
|
||||
<date id="FF83FCDBFFCB472B23C5F93B7435F931" box="[1229,1358,1670,1694]" pageId="7" pageNumber="8" value="2017-03">
|
||||
<collectingDate id="EFC70533FFCB472B23C5F93B7435F931" box="[1229,1358,1670,1694]" pageId="7" pageNumber="8" value="2017-03">March 2017</collectingDate>
|
||||
</date>
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="3B55D046FFCB472B2251F93B7453F953" collectingDate="2017-03" collectionCode="RSIO" elevation="3480" latitude="6.8" longLatPrecision="129" longitude="60.18" pageId="7" pageNumber="8" specimenCode="RSIO 38228" specimenCount="1" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
|
||||
<typeStatus id="548664B9FFCB472B2251F93B74C8F931" box="[1369,1459,1670,1694]" pageId="7" pageNumber="8" type="paratype">Paratype</typeStatus>
|
||||
1 (
|
||||
<specimenCode id="DB9B7260FFCB472B2440F91872AEF912" box="[840,981,1701,1725]" collectionCode="RSIO" pageId="7" pageNumber="8">RSIO 38228</specimenCode>
|
||||
;
|
||||
<figureCitation id="1306C69EFFCB472B24EBF918754AF912" box="[995,1073,1701,1725]" captionStart="Figure 5" captionStartId="8.[129,194,1562,1586]" captionTargetBox="[129,1473,144,1535]" captionTargetId="figure-187@8.[129,1473,144,1535]" captionTargetPageId="8" captionText="Figure 5. Shinkailepas cornuthauma, specimens examined in this study. A, specimen collected from Wocan field (RSIO 38229), female. B, holotype (RSIO 38227), male. White arrow indicates the penis. C, paratype 1 (RSIO 38228), female, shell damaged during collection." pageId="7" pageNumber="8">Fig. 5C</figureCitation>
|
||||
),
|
||||
<specimenCount id="9D3B1192FFCB472B2344F91875EAF912" box="[1100,1169,1701,1725]" pageId="7" pageNumber="8" type="female">female</specimenCount>
|
||||
, 95% ethanol, shell broken during collection, animal approximately
|
||||
<quantity id="4CC577FEFFCB472B23D9F9797464F973" box="[1233,1311,1732,1756]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="7" pageNumber="8" unit="mm" value="8.5">8.5 mm</quantity>
|
||||
in length and
|
||||
<quantity id="4CC577FEFFCB472B2422F9597203F954" box="[810,888,1764,1788]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="7" pageNumber="8" unit="mm" value="5.5">5.5 mm</quantity>
|
||||
in width, same collecting data as holotype
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8B82DA1BFFCB472B2422F89C759EF819" blockId="7.[810,1459,1824,1974]" pageId="7" pageNumber="8">
|
||||
<emphasis id="B9490609FFCB472B2422F89C7297F896" box="[810,1004,1825,1849]" italics="true" pageId="7" pageNumber="8">Material examined:</emphasis>
|
||||
<materialsCitation id="3B55D046FFCB472B24FDF89C759AF819" collectingDate="2017-03" collectionCode="A" elevation="2920" latitude="6.36" longLatPrecision="129" longitude="60.53" pageId="7" pageNumber="8" specimenCode="RSIO 38229" specimenCount="2" specimenCount-female="1">
|
||||
<specimenCount id="9D3B1192FFCB472B24FDF89C75F7F896" box="[1013,1164,1825,1849]" count="1" pageId="7" pageNumber="8" type="generic">One specimen</specimenCount>
|
||||
(
|
||||
<specimenCode id="DB9B7260FFCB472B2394F89C745CF896" box="[1180,1319,1825,1849]" collectionCode="RSIO" pageId="7" pageNumber="8">RSIO 38229</specimenCode>
|
||||
;
|
||||
<figureCitation id="1306C69EFFCB472B223CF89C74FBF897" box="[1332,1408,1824,1849]" captionStart="Figure 5" captionStartId="8.[129,194,1562,1586]" captionTargetBox="[129,1473,144,1535]" captionTargetId="figure-187@8.[129,1473,144,1535]" captionTargetPageId="8" captionText="Figure 5. Shinkailepas cornuthauma, specimens examined in this study. A, specimen collected from Wocan field (RSIO 38229), female. B, holotype (RSIO 38227), male. White arrow indicates the penis. C, paratype 1 (RSIO 38228), female, shell damaged during collection." pageId="7" pageNumber="8">
|
||||
Fig. 5
|
||||
<collectionCode id="ED2C42DEFFCB472B2265F89D74FBF897" box="[1389,1408,1824,1848]" pageId="7" pageNumber="8">A</collectionCode>
|
||||
</figureCitation>
|
||||
),
|
||||
<specimenCount id="9D3B1192FFCB472B2290F89C7226F8F7" pageId="7" pageNumber="8" type="female">female</specimenCount>
|
||||
, 95% ethanol, SL
|
||||
<quantity id="4CC577FEFFCB472B2315F8FD7516F8F8" box="[1053,1133,1856,1880]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="7" pageNumber="8" unit="mm" value="6.7">6.7 mm</quantity>
|
||||
, SW 5.0 mm, Wocan vent field (
|
||||
<geoCoordinate id="EE09BCDCFFCB472B2439F8E272E0F8D8" box="[817,923,1886,1911]" degrees="60" direction="east" minutes="31.8" orientation="longitude" pageId="7" pageNumber="8" precision="92" value="60.53">
|
||||
60°31.8
|
||||
<emphasis id="B9490609FFCB472B248CF8E372F1F8D8" box="[900,906,1886,1911]" italics="true" pageId="7" pageNumber="8">ʹ</emphasis>
|
||||
E
|
||||
</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE09BCDCFFCB472B24A4F8E27571F8D8" box="[940,1034,1886,1911]" degrees="6" direction="north" minutes="21.6" orientation="latitude" pageId="7" pageNumber="8" precision="92" value="6.36">
|
||||
6°21.6
|
||||
<emphasis id="B9490609FFCB472B24E5F8E37288F8D8" box="[1005,1011,1886,1911]" italics="true" pageId="7" pageNumber="8">ʹ</emphasis>
|
||||
N
|
||||
</geoCoordinate>
|
||||
),
|
||||
<quantity id="4CC577FEFFCB472B232CF8E27500F8D8" box="[1060,1147,1887,1911]" metricMagnitude="3" metricUnit="m" metricValue="2.92" pageId="7" pageNumber="8" unit="m" value="2920.0">
|
||||
<elevation id="00103D28FFCB472B232CF8E27500F8D8" box="[1060,1147,1887,1911]" metricMagnitude="3" metricUnit="m" metricValue="2.92" pageId="7" pageNumber="8" unit="m" value="2920.0">2920 m</elevation>
|
||||
</quantity>
|
||||
deep, collected by a seven-function manipulator of HOV
|
||||
<emphasis id="B9490609FFCB472B238AF8C275ABF838" box="[1154,1232,1919,1943]" italics="true" pageId="7" pageNumber="8">Jiaolong</emphasis>
|
||||
on dive #125, R/V
|
||||
<emphasis id="B9490609FFCB472B2422F82372AFF819" box="[810,980,1950,1974]" italics="true" pageId="7" pageNumber="8">Xiangyanghong 9</emphasis>
|
||||
cruise DY38,
|
||||
<date id="FF83FCDBFFCB472B236BF823759AF819" box="[1123,1249,1950,1974]" pageId="7" pageNumber="8" value="2017-03">
|
||||
<collectingDate id="EFC70533FFCB472B236BF823759AF819" box="[1123,1249,1950,1974]" pageId="7" pageNumber="8" value="2017-03">March 2017</collectingDate>
|
||||
</date>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF428A93FFC447242789F9A77408F9E1" pageId="8" pageNumber="9" startId="8.[129,194,1562,1586]" targetBox="[129,1473,144,1535]" targetPageId="8" targetType="figure">
|
||||
<paragraph id="8B82DA1BFFC447242789F9A77408F9E1" blockId="8.[129,1417,1562,1615]" pageId="8" pageNumber="9">
|
||||
<emphasis id="B9490609FFC447242789F9A771A3F99C" bold="true" box="[129,216,1562,1587]" pageId="8" pageNumber="9">Figure 5.</emphasis>
|
||||
<taxonomicName id="4C3DA198FFC4472427D5F9A770B6F99D" authorityName="Gu & Chen & Gao & Zhou & Sun" authorityYear="2025" box="[221,461,1562,1586]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="cornuthauma">
|
||||
<emphasis id="B9490609FFC4472427D5F9A770B6F99D" box="[221,461,1562,1586]" italics="true" pageId="8" pageNumber="9">Shinkailepas cornuthauma</emphasis>
|
||||
</taxonomicName>
|
||||
, specimens examined in this study. A, specimen collected from Wocan field (RSIO 38229), female. B, holotype (RSIO 38227), male. White arrow indicates the penis. C, paratype 1 (RSIO 38228), female, shell damaged during collection.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="C3278990FFC447262789F9C273B5F950" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="description">
|
||||
<paragraph id="8B82DA1BFFC447242789F9C27538F81F" blockId="8.[129,779,1662,1969]" lastBlockId="8.[824,1475,1662,1969]" pageId="8" pageNumber="9">
|
||||
<emphasis id="B9490609FFC447242789F9C27180F939" box="[129,251,1663,1686]" italics="true" pageId="8" pageNumber="9">Description:</emphasis>
|
||||
Shell (
|
||||
<figureCitation id="1306C69EFFC44724265FF9C370E2F939" box="[343,409,1662,1686]" captionStart="Figure 5" captionStartId="8.[129,194,1562,1586]" captionTargetBox="[129,1473,144,1535]" captionTargetId="figure-187@8.[129,1473,144,1535]" captionTargetPageId="8" captionText="Figure 5. Shinkailepas cornuthauma, specimens examined in this study. A, specimen collected from Wocan field (RSIO 38229), female. B, holotype (RSIO 38227), male. White arrow indicates the penis. C, paratype 1 (RSIO 38228), female, shell damaged during collection." pageId="8" pageNumber="9">Fig. 5</figureCitation>
|
||||
) medium-sized for genus, adult limpet-formed, with evidence of coiling when juvenile. Shell length 1.2–1.4 times of width; shell height approximately onequarter of length. Aperture oval, reduced and minute septum (‘deck’) present posteriorly. Apex positioned at approximately 30% from posterior shell edge, weakly recurved to right side. Shell sculpture dominated by very strong radial ribs crossing with concentric growth lines to form weakly cancellate sculpture; raised protuberances form where two directions converge. Radial sculpture with three to five minor ribs between each pair of major ribs. Shell margin rather flat. Shell interior translucent-white, with numerous shell pores. Ostracum moderately thin, translucent. Numerous shell pores present on inner ostracum surface but absent on septum. Shell microstructure (
|
||||
<figureCitation id="1306C69EFFC4472424A0F9417288F8BB" box="[936,1011,1788,1812]" captionStart="Figure 7" captionStartId="10.[129,194,1040,1064]" captionTargetBox="[129,1473,144,1012]" captionTargetId="figure-475@10.[129,1473,144,1012]" captionTargetPageId="10" captionText="Figure 7. Shinkailepas cornuthauma, scanning electron micrographs of the holotype (RSIO 38227). A, shell microstructure. B, overview of the radula. C, close-up of the left half of the radula. D, close-up of the outer laterals and marginal teeth." pageId="8" pageNumber="9">Fig. 7A</figureCitation>
|
||||
) with three layers dorsal to the myostracum, a thin outermost granular layer, followed by a thick simple prismatic layer, then by an equally thick crossed lamellar layer. Protoconch unknown as corroded in all specimens available for study. Periostracum moderately thick, greenish, often covered in rusty mineral deposits.
|
||||
</paragraph>
|
||||
<caption id="DF428A93FFC547252779F92E7061F8B4" pageId="9" pageNumber="10" startId="9.[113,178,1683,1707]" targetBox="[114,1458,144,1655]" targetPageId="9" targetType="figure">
|
||||
<paragraph id="8B82DA1BFFC547252779F92E7061F8B4" blockId="9.[113,1457,1683,1819]" pageId="9" pageNumber="10">
|
||||
<emphasis id="B9490609FFC547252779F92E71B2F904" bold="true" box="[113,201,1683,1707]" pageId="9" pageNumber="10">Figure 6.</emphasis>
|
||||
<taxonomicName id="4C3DA198FFC5472527C6F92E70C5F904" authorityName="Gu & Chen & Gao & Zhou & Sun" authorityYear="2025" box="[206,446,1683,1707]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="cornuthauma">
|
||||
<emphasis id="B9490609FFC5472527C6F92E70C5F904" box="[206,446,1683,1707]" italics="true" pageId="9" pageNumber="10">Shinkailepas cornuthauma</emphasis>
|
||||
</taxonomicName>
|
||||
, external anatomy of the holotype male (RSIO 38227). A, dorsal aspect with shell removed. B, enlarged dorsal view of the head with mantle edge removed, female. C, enlarged ventral of the head. D, operculum shown attached to dorsal aspect of foot. Abbreviations: ang, annex gland; cl, cephalic lappet; ct, ctenidium; dg, digestive gland; e, eye; f, foot; lsm, left shell muscle; m, mouth; me, mantle edge; nl, neck lobe; ol, oral lappet; pe, penis; pc, pericardium; pr, prostate; rsm, right shell muscle; sn, snout; st, stomach; t, cephalic tentacle; te, testis.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B82DA1BFFC547262785F8F670C6FB64" blockId="9.[113,763,1867,1954]" lastBlockId="10.[127,779,1140,1791]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
|
||||
External anatomy (
|
||||
<figureCitation id="1306C69EFFC54725265CF8F670C4F8CC" box="[340,447,1867,1891]" captionStart="Figure 6" captionStartId="9.[113,178,1683,1707]" captionTargetBox="[114,1458,144,1655]" captionTargetId="figure-69@9.[114,1458,144,1655]" captionTargetPageId="9" captionText="Figure 6. Shinkailepas cornuthauma, external anatomy of the holotype male (RSIO 38227). A, dorsal aspect with shell removed.B, enlarged dorsal view of the head with mantle edge removed, female. C, enlarged ventral of the head.D, operculum shown attached to dorsal aspect of foot.Abbreviations:ang, annex gland; cl, cephalic lappet; ct, ctenidium; dg, digestive gland; e, eye; f, foot; lsm, left shell muscle; m, mouth; me, mantle edge; nl, neck lobe; ol, oral lappet; pe, penis; pc, pericardium; pr, prostate; rsm, right shell muscle; sn, snout; st, stomach; t, cephalic tentacle; te, testis." pageId="9" pageNumber="10">Fig. 6A–C</figureCitation>
|
||||
) typical for genus, most characters agree well with those described under
|
||||
<taxonomicName id="4C3DA198FFC547252531F8D673CFF82D" box="[569,692,1899,1922]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="tiarasimia">
|
||||
<emphasis id="B9490609FFC547252531F8D673CFF82D" box="[569,692,1899,1922]" italics="true" pageId="9" pageNumber="10">S. tiarasimia</emphasis>
|
||||
</taxonomicName>
|
||||
above. Epipodial fold (
|
||||
<figureCitation id="1306C69EFFC54725261FF837702AF80D" box="[279,337,1930,1954]" captionStart="Figure 5" captionStartId="8.[129,194,1562,1586]" captionTargetBox="[129,1473,144,1535]" captionTargetId="figure-187@8.[129,1473,144,1535]" captionTargetPageId="8" captionText="Figure 5. Shinkailepas cornuthauma, specimens examined in this study. A, specimen collected from Wocan field (RSIO 38229), female. B, holotype (RSIO 38227), male. White arrow indicates the penis. C, paratype 1 (RSIO 38228), female, shell damaged during collection." pageId="9" pageNumber="10">Fig. 5</figureCitation>
|
||||
) carries 15–18 small triangular tentacles posteriorly. Operculum (
|
||||
<figureCitation id="1306C69EFFC547252335F8F675E9F8CC" box="[1085,1170,1867,1891]" captionStart="Figure 6" captionStartId="9.[113,178,1683,1707]" captionTargetBox="[114,1458,144,1655]" captionTargetId="figure-69@9.[114,1458,144,1655]" captionTargetPageId="9" captionText="Figure 6. Shinkailepas cornuthauma, external anatomy of the holotype male (RSIO 38227). A, dorsal aspect with shell removed.B, enlarged dorsal view of the head with mantle edge removed, female. C, enlarged ventral of the head.D, operculum shown attached to dorsal aspect of foot.Abbreviations:ang, annex gland; cl, cephalic lappet; ct, ctenidium; dg, digestive gland; e, eye; f, foot; lsm, left shell muscle; m, mouth; me, mantle edge; nl, neck lobe; ol, oral lappet; pe, penis; pc, pericardium; pr, prostate; rsm, right shell muscle; sn, snout; st, stomach; t, cephalic tentacle; te, testis." pageId="9" pageNumber="10">Fig. 6D</figureCitation>
|
||||
) large, entirely corneous, occupying over half of dorsal surface of foot, exact shape could not be determined due to its folded condition in examined specimens from preservation issues. Diameter of opercular nucleus approximately 500 μm, but due to its folded state, exact measurement was not possible.
|
||||
</paragraph>
|
||||
<caption id="DF428A93FFC647262789FBAD7567FBEB" pageId="10" pageNumber="11" startId="10.[129,194,1040,1064]" targetBox="[129,1473,144,1012]" targetPageId="10" targetType="figure">
|
||||
<paragraph id="8B82DA1BFFC647262789FBAD7567FBEB" blockId="10.[129,1462,1040,1092]" pageId="10" pageNumber="11">
|
||||
<emphasis id="B9490609FFC647262789FBAD71A3FB87" bold="true" box="[129,216,1040,1064]" pageId="10" pageNumber="11">Figure 7.</emphasis>
|
||||
<taxonomicName id="4C3DA198FFC6472627D5FBAD70B6FB87" authorityName="Gu & Chen & Gao & Zhou & Sun" authorityYear="2025" box="[221,461,1040,1064]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="cornuthauma">
|
||||
<emphasis id="B9490609FFC6472627D5FBAD70B6FB87" box="[221,461,1040,1064]" italics="true" pageId="10" pageNumber="11">Shinkailepas cornuthauma</emphasis>
|
||||
</taxonomicName>
|
||||
, scanning electron micrographs of the holotype (RSIO 38227). A, shell microstructure. B, overview of the radula. C, close-up of the left half of the radula. D, close-up of the outer laterals and marginal teeth.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B82DA1BFFC647262794FB6F73B5F950" blockId="10.[127,779,1140,1791]" pageId="10" pageNumber="11">
|
||||
Radula (
|
||||
<figureCitation id="1306C69EFFC6472627FDFB6F7027FB45" box="[245,348,1234,1258]" captionStart="Figure 7" captionStartId="10.[129,194,1040,1064]" captionTargetBox="[129,1473,144,1012]" captionTargetId="figure-475@10.[129,1473,144,1012]" captionTargetPageId="10" captionText="Figure 7. Shinkailepas cornuthauma, scanning electron micrographs of the holotype (RSIO 38227). A, shell microstructure. B, overview of the radula. C, close-up of the left half of the radula. D, close-up of the outer laterals and marginal teeth." pageId="10" pageNumber="11">Fig. 7B–D</figureCitation>
|
||||
) rhipidoglossate, overall dentition pattern typical for genus (
|
||||
<bibRefCitation id="EFACA7EAFFC64726263AFB4C7085FAA5" author="Fukumori H & Yahagi T & Waren A" box="[306,510,1265,1290]" pageId="10" pageNumber="11" pagination="636 - 55" refId="ref10278" refString="Fukumori H, Yahagi T, Waren A et al. Amended generic classification of themarinegastropodfamilyPhenacolepadidae: transitionsfromsnails to limpets and shallow-water to deep-sea hydrothermal vents and cold seeps. Zoological Journal of the Linnean Society 2019; 185: 636 - 55. https: // doi. org / 10.1093 / zoolinnean / zly 078" type="journal article" year="2019">
|
||||
Fukumori
|
||||
<emphasis id="B9490609FFC647262693FB4F70B3FAA6" box="[411,456,1265,1289]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
|
||||
2019
|
||||
</bibRefCitation>
|
||||
), formula
|
||||
<emphasis id="B9490609FFC64726256DFB4F7315FAA6" box="[613,622,1266,1289]" italics="true" pageId="10" pageNumber="11">c</emphasis>
|
||||
. 100–4–1–4–
|
||||
<emphasis id="B9490609FFC6472625F3FB4F727FFAA6" box="[763,772,1266,1289]" italics="true" pageId="10" pageNumber="11">c</emphasis>
|
||||
. 100. Central tooth subquadrate, about 1.5 times as long as wide, with thickened central ridge running horizontally. Innermost first lateral film-like, strongly oblique, twice as wide as central tooth, cusp indistinct, lacking serration, curled over distally to form overhanging flap, outermost side with raised fold. Second lateral narrower than central tooth, hook-shaped with shaft recurved inwards, inner side of shaft with one raised hump; double-cusped with upper one smooth, lower one carrying four sharp minor cusps. Third lateral (
|
||||
<figureCitation id="1306C69EFFC64726264FF9B670F5F98C" box="[327,398,1547,1571]" captionStart="Figure 7" captionStartId="10.[129,194,1040,1064]" captionTargetBox="[129,1473,144,1012]" captionTargetId="figure-475@10.[129,1473,144,1012]" captionTargetPageId="10" captionText="Figure 7. Shinkailepas cornuthauma, scanning electron micrographs of the holotype (RSIO 38227). A, shell microstructure. B, overview of the radula. C, close-up of the left half of the radula. D, close-up of the outer laterals and marginal teeth." pageId="10" pageNumber="11">Fig.7D</figureCitation>
|
||||
) narrow with inward-rolled shaft, with one minute denticle, otherwise smooth. Outermost fourth lateral large, with solid, smooth, very slightly inwardly recurved shaft, distally enlarging to form wide cusp divided into five denticles, of which second from inside strongest by far. Inner marginals with shafts similar in length to fourth lateral, serrated into 4–5 denticles distally. Outer marginals much smaller, cusp rake-like, serrated into many fine denticles whose numbers increase outwards.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3278990FFC647262789F89D7588FB04" pageId="10" pageNumber="11" type="etymology">
|
||||
<paragraph id="8B82DA1BFFC647262789F89D7588FB04" blockId="10.[128,779,1824,1973]" lastBlockId="10.[825,1474,1140,1196]" pageId="10" pageNumber="11">
|
||||
<emphasis id="B9490609FFC647262789F89D718BF897" box="[129,240,1824,1848]" italics="true" pageId="10" pageNumber="11">Etymology:</emphasis>
|
||||
The specific epithet is a combination of Latin
|
||||
<emphasis id="B9490609FFC6472625DCF89C7270F897" box="[724,779,1825,1848]" italics="true" pageId="10" pageNumber="11">cornū</emphasis>
|
||||
(‘horn’) and Ancient Greek
|
||||
<emphasis id="B9490609FFC6472626A9F8827096F8F8" box="[417,493,1855,1879]" italics="true" pageId="10" pageNumber="11">thaûma</emphasis>
|
||||
(‘wonder, marvel, astonishment’), referring to its horn-like, curved apex and the wonderful happening of collecting the few specimens known, allowing its formal description. This name was also partly inspired by the magical helm ‘
|
||||
<emphasis id="B9490609FFC6472624D9FBC9753CFB23" box="[977,1095,1140,1164]" italics="true" pageId="10" pageNumber="11">cornuthaum</emphasis>
|
||||
’ that appears in the roguelike video game
|
||||
<emphasis id="B9490609FFC64726247DFB2E72B7FB04" box="[885,972,1171,1195]" italics="true" pageId="10" pageNumber="11">NetHack</emphasis>
|
||||
. Used as noun in apposition.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3278990FFC647262431FB7072D7FAAB" pageId="10" pageNumber="11" type="distribution">
|
||||
<paragraph id="8B82DA1BFFC647262431FB7072D7FAAB" blockId="10.[825,1474,1229,1284]" pageId="10" pageNumber="11">
|
||||
<emphasis id="B9490609FFC647262431FB7072CCFB4A" box="[825,951,1229,1253]" italics="true" pageId="10" pageNumber="11">Distribution:</emphasis>
|
||||
Only known from the Daxi and Wocan vent fields on the CR.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3278990FFC647272431FA987182FC5A" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="discussion">
|
||||
<paragraph id="8B82DA1BFFC647262431FA987572F95B" blockId="10.[825,1475,1317,1780]" pageId="10" pageNumber="11">
|
||||
<emphasis id="B9490609FFC647262431FA9875E4FA92" box="[825,1183,1317,1341]" italics="true" pageId="10" pageNumber="11">
|
||||
Remarks:
|
||||
<taxonomicName id="4C3DA198FFC647262494FA9875E4FA92" authorityName="Gu & Chen & Gao & Zhou & Sun" authorityYear="2025" box="[924,1183,1317,1341]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="cornuthauma">Shinkailepas cornuthauma</taxonomicName>
|
||||
</emphasis>
|
||||
is clearly distinct from other described
|
||||
<taxonomicName id="4C3DA198FFC6472624AAFAF87510FAF2" box="[930,1131,1349,1373]" pageId="10" pageNumber="11">
|
||||
<emphasis id="B9490609FFC6472624AAFAF87566FAF2" box="[930,1053,1349,1373]" italics="true" pageId="10" pageNumber="11">Shinkailepas</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
by its relatively anterior apex position at approximately 30% of SL from the posterior shell edge, a very reduced septum (‘deck’), combined with its strong radial shell sculpture with three to five minor ribs between two major ribs.
|
||||
<taxonomicName id="4C3DA198FFC647262462FA7F753CFA75" baseAuthorityName="Beck" baseAuthorityYear="1992" box="[874,1095,1474,1498]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="tollmanni">
|
||||
<emphasis id="B9490609FFC647262462FA7F753CFA75" box="[874,1095,1474,1498]" italics="true" pageId="10" pageNumber="11">Shinkailepas tollmanni</emphasis>
|
||||
</taxonomicName>
|
||||
is the only similar species with a similarly reduced condition of the septum, but its sculpture consists solely of fine radial ribs which are smooth instead of strongly granulated as in
|
||||
<taxonomicName id="4C3DA198FFC6472624E4F99D75F1F997" authorityName="Gu & Chen & Gao & Zhou & Sun" authorityYear="2025" box="[1004,1162,1568,1592]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="cornuthauma">
|
||||
<emphasis id="B9490609FFC6472624E4F99D75F1F997" box="[1004,1162,1568,1592]" italics="true" pageId="10" pageNumber="11">S. cornuthauma</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<bibRefCitation id="EFACA7EAFFC647262397F99D7474F997" author="Beck LA" box="[1183,1295,1568,1592]" pageId="10" pageNumber="11" pagination="259 - 75" refId="ref9697" refString="Beck LA. Two new neritacean limpets (Gastropoda: Prosobranchia: Neritacea: Phenacolepadidae) from active hydrothermal vents at Hydrothermal Field 1 ' Wienerwald' in the Manus Back-Arc Basin (Bismarck Sea, Papua-New Guinea). Annalen des Naturhistorischen Museums in Wien. Serie B fur Botanik und Zoologie 1992; 93: 259 - 75." type="journal article" year="1992">Beck 1992</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EFACA7EAFFC647262217F99D7216F9F7" author="Fukumori H & Yahagi T & Waren A" pageId="10" pageNumber="11" pagination="636 - 55" refId="ref10278" refString="Fukumori H, Yahagi T, Waren A et al. Amended generic classification of themarinegastropodfamilyPhenacolepadidae: transitionsfromsnails to limpets and shallow-water to deep-sea hydrothermal vents and cold seeps. Zoological Journal of the Linnean Society 2019; 185: 636 - 55. https: // doi. org / 10.1093 / zoolinnean / zly 078" type="journal article" year="2019">
|
||||
Fukumori
|
||||
<emphasis id="B9490609FFC647262287F99C74B8F997" box="[1423,1475,1568,1592]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
|
||||
2019
|
||||
</bibRefCitation>
|
||||
). Furthermore, the epipodial tentacles of
|
||||
<taxonomicName id="4C3DA198FFC647262248F98274B9F9F8" authorityName="Gu & Chen & Gao & Zhou & Sun" authorityYear="2025" box="[1344,1474,1599,1623]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="cornuthauma">
|
||||
<emphasis id="B9490609FFC647262248F98274B9F9F8" box="[1344,1474,1599,1623]" italics="true" pageId="10" pageNumber="11">cornuthauma</emphasis>
|
||||
</taxonomicName>
|
||||
are small, triangular, and number between 15–18, which aids to distinguish it from species like
|
||||
<taxonomicName id="4C3DA198FFC64726239EF9C3747BF939" box="[1174,1280,1662,1686]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="conspira">
|
||||
<emphasis id="B9490609FFC64726239EF9C3747BF939" box="[1174,1280,1662,1686]" italics="true" pageId="10" pageNumber="11">S. conspira</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C3DA198FFC64726223BF9C37400F939" box="[1331,1403,1662,1686]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="gigas">
|
||||
<emphasis id="B9490609FFC64726223BF9C37400F939" box="[1331,1403,1662,1686]" italics="true" pageId="10" pageNumber="11">S. gigas</emphasis>
|
||||
</taxonomicName>
|
||||
whose epipodial folds are divided into over 50 large, paddle-like projections (
|
||||
<bibRefCitation id="EFACA7EAFFC647262493F90075E8F97A" author="Chen C & Sigwart JD" box="[923,1171,1725,1749]" pageId="10" pageNumber="11" pagination="401 - 36" refId="ref9891" refString="Chen C, Sigwart JD. The lost vent gastropod species of Lothar A. Beck. Zootaxa 2023; 5270: 401 - 36. https: // doi. org / 10.11646 / zootaxa. 5270.3.2" type="journal article" year="2023">Chen and Sigwart 2023</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EFACA7EAFFC6472623A8F9007282F95B" author="Senckenberg Ocean Species Alliance & SOSA & Brandt A & Chen C" pageId="10" pageNumber="11" refId="ref11378" refString="Senckenberg Ocean Species Alliance (SOSA), Brandt A, Chen C, et al. Ocean species discoveries 1 - 12 - a primer for accelerating marine invertebrate taxonomy. Biodiversity Data Journal 2024; 12: e 128431. https: // doi. org / 10.3897 / BDJ. 12. e 128431" type="journal volume" year="2024">
|
||||
Senckenberg Ocean Species Alliance
|
||||
<emphasis id="B9490609FFC64726249AF96072BAF95B" box="[914,961,1756,1780]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
|
||||
2024
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8B82DA1BFFC647262311F8A97599F881" blockId="10.[1049,1250,1812,1838]" box="[1049,1250,1812,1838]" pageId="10" pageNumber="11">
|
||||
<emphasis id="B9490609FFC647262311F8A97599F881" bold="true" box="[1049,1250,1812,1838]" pageId="10" pageNumber="11">Molecular analyses</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="8B82DA1BFFC647272431F88673C1FD12" blockId="10.[825,1474,1851,1969]" lastBlockId="11.[112,763,144,1014]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
|
||||
Our phylogenetic tree reconstructed using the
|
||||
<emphasis id="B9490609FFC64726225EF88674F8F8FD" box="[1366,1411,1851,1874]" italics="true" pageId="10" pageNumber="11">COI</emphasis>
|
||||
gene (
|
||||
<figureCitation id="1306C69EFFC64726244CF8E772FAF8DD" box="[836,897,1882,1906]" captionStart="Figure 8" captionStartId="11.[113,178,1869,1893]" captionTargetBox="[114,1458,1114,1839]" captionTargetId="figure-578@11.[114,1458,1113,1841]" captionTargetPageId="11" captionText="Figure 8. Phylogenetic tree of Shinkailepas reconstructed using Bayesian inference from 658 bp of the mitochondrial cytochrome c oxidase subunit I (COI) gene.GenBank accession numbers of the sequences included are shown in parentheses. Node values indicate Bayesian posterior probabilities, only those higher than 0.7 are shown. Representation of undescribed species follows Poitrimol et al.(2022); Woodlark, Lau, and Futuna refer to Woodlark Basin, Lau Basin, and Futuna Arc in the southwestern Pacific, respectively." pageId="10" pageNumber="11">Fig. 8</figureCitation>
|
||||
) recovered a strongly supported monophyletic genus
|
||||
<taxonomicName id="4C3DA198FFC647262431F8C472CCF83E" box="[825,951,1913,1937]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B9490609FFC647262431F8C472CCF83E" box="[825,951,1913,1937]" italics="true" pageId="10" pageNumber="11">Shinkailepas</emphasis>
|
||||
</taxonomicName>
|
||||
(Bayesian Posterior Probability, BPP = 0.9) including
|
||||
<taxonomicName id="4C3DA198FFC6472624AEF824755CF81F" box="[934,1063,1945,1968]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="tiarasimia">
|
||||
<emphasis id="B9490609FFC6472624AEF824755CF81F" box="[934,1063,1945,1968]" italics="true" pageId="10" pageNumber="11">S. tiarasimia</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C3DA198FFC64726236AF824747FF81E" authorityName="Gu & Chen & Gao & Zhou & Sun" authorityYear="2025" box="[1122,1284,1945,1969]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="cornuthauma">
|
||||
<emphasis id="B9490609FFC64726236AF824747FF81E" box="[1122,1284,1945,1969]" italics="true" pageId="10" pageNumber="11">S. cornuthauma</emphasis>
|
||||
</taxonomicName>
|
||||
, supporting their inclusion in this genus. Two major clades were recovered within
|
||||
<taxonomicName id="4C3DA198FFC7472727B4FF127041FF68" box="[188,314,175,199]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B9490609FFC7472727B4FF127041FF68" box="[188,314,175,199]" italics="true" pageId="11" pageNumber="12">Shinkailepas</emphasis>
|
||||
</taxonomicName>
|
||||
, one containing
|
||||
<taxonomicName id="4C3DA198FFC7472726E3FF0D730AFF68" authorityName="Okutani, Saito & Hashimoto" authorityYear="1989" box="[491,625,175,199]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="kaikatensis">
|
||||
<emphasis id="B9490609FFC7472726E3FF0D730AFF68" box="[491,625,175,199]" italics="true" pageId="11" pageNumber="12">S. kaikatensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C3DA198FFC747272575FF0D73BCFF68" box="[637,711,176,199]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="gigas">
|
||||
<emphasis id="B9490609FFC747272575FF0D73BCFF68" box="[637,711,176,199]" italics="true" pageId="11" pageNumber="12">S. gigas</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="4C3DA198FFC74727277AFF72706FFF48" authorityName="Gu & Chen & Gao & Zhou & Sun" authorityYear="2025" box="[114,276,207,231]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="cornuthauma">
|
||||
<emphasis id="B9490609FFC74727277AFF72706FFF48" box="[114,276,207,231]" italics="true" pageId="11" pageNumber="12">S. cornuthauma</emphasis>
|
||||
</taxonomicName>
|
||||
(‘Clade A’ hereafter) and the other with all remaining species (‘Clade B’ hereafter); both were strongly supported (BPP = 0.94 and 0.96, respectively). Within Clade A,
|
||||
<taxonomicName id="4C3DA198FFC74727279AFE907049FEEA" authorityName="Gu & Chen & Gao & Zhou & Sun" authorityYear="2025" box="[146,306,301,325]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="cornuthauma">
|
||||
<emphasis id="B9490609FFC74727279AFE907049FEEA" box="[146,306,301,325]" italics="true" pageId="11" pageNumber="12">S. cornuthauma</emphasis>
|
||||
</taxonomicName>
|
||||
was the earliest diverging species sister to the
|
||||
<emphasis id="B9490609FFC747272796FEF170E8FECB" box="[158,403,332,356]" italics="true" pageId="11" pageNumber="12">
|
||||
<taxonomicName id="4C3DA198FFC747272796FEF17057FECB" authorityName="Okutani, Saito & Hashimoto" authorityYear="1989" box="[158,300,332,356]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="kaikatensis">S. kaikatensis</taxonomicName>
|
||||
—
|
||||
<taxonomicName id="4C3DA198FFC74727264FFEF170E8FECB" box="[327,403,332,356]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="gigas">S. gigas</taxonomicName>
|
||||
</emphasis>
|
||||
pair, but only with low support (BPP <70). Within Clade B, there was a basal split between the fully supported (BPP = 1)
|
||||
<taxonomicName id="4C3DA198FFC7472726A7FE3671D1FE6D" authority="Woodlark Basin" authorityName="Woodlark Basin" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" isUncertain="true" kingdom="Animalia" order="Cycloneritida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="tufari">
|
||||
<emphasis id="B9490609FFC7472726A7FE367356FE0C" box="[431,557,395,419]" italics="true" pageId="11" pageNumber="12">Shinkailepas</emphasis>
|
||||
aff.
|
||||
<emphasis id="B9490609FFC74727255EFE3673F5FE0C" box="[598,654,395,419]" italics="true" pageId="11" pageNumber="12">tufari</emphasis>
|
||||
Woodlark Basin
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C3DA198FFC7472727EEFE1770F0FE6E" box="[230,395,426,450]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="undetermined">
|
||||
<emphasis id="B9490609FFC7472727EEFE17701FFE6D" box="[230,356,426,450]" italics="true" pageId="11" pageNumber="12">Shinkailepas</emphasis>
|
||||
sp.
|
||||
</taxonomicName>
|
||||
Mariana Trough (
|
||||
<emphasis id="B9490609FFC747272550FE1673F7FE6D" box="[600,652,427,450]" italics="true" pageId="11" pageNumber="12">sensu</emphasis>
|
||||
<bibRefCitation id="EFACA7EAFFC74727259EFE17719CFE4E" author="Poitrimol C & Thiebaut E & Daguin-Thiebaut C" pageId="11" pageNumber="12" refId="ref11104" refString="Poitrimol C, Thiebaut E, Daguin-Thiebaut C et al. Contrasted phylogeographic patterns of hydrothermal vent gastropods along South West Pacific: Woodlark Basin, a possible contact zone and / or stepping-stone. PLoS One 2022; 17: e 0275638. https: // doi. org / 10.1371 / journal. pone. 0275638" type="journal volume" year="2022">
|
||||
Poitrimol
|
||||
<emphasis id="B9490609FFC747272779FE7771D3FE4E" box="[113,168,457,481]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
|
||||
2022
|
||||
</bibRefCitation>
|
||||
) pair and all other taxa.
|
||||
<taxonomicName id="4C3DA198FFC747272503FE747381FE4E" box="[523,762,457,481]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="tiarasimia">
|
||||
<emphasis id="B9490609FFC747272503FE747381FE4E" box="[523,762,457,481]" italics="true" pageId="11" pageNumber="12">Shinkailepas tiarasimia</emphasis>
|
||||
</taxonomicName>
|
||||
was recovered as sister to
|
||||
<taxonomicName id="4C3DA198FFC747272697FE54736AFDAF" box="[415,529,489,512]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="conspira">
|
||||
<emphasis id="B9490609FFC747272697FE54736AFDAF" box="[415,529,489,512]" italics="true" pageId="11" pageNumber="12">S. conspira</emphasis>
|
||||
</taxonomicName>
|
||||
with strong support (BPP = 0.98). This pair was in turn clustered with
|
||||
<taxonomicName id="4C3DA198FFC74727258DFDB571A3FD90" authority="Lau Basin" authorityName="Lau Basin" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" isUncertain="true" kingdom="Animalia" order="Cycloneritida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="tufari">
|
||||
<emphasis id="B9490609FFC74727258DFDB573E3FDB0" box="[645,664,520,543]" italics="true" pageId="11" pageNumber="12">S.</emphasis>
|
||||
aff.
|
||||
<emphasis id="B9490609FFC7472725CBFDB57380FD8F" box="[707,763,520,544]" italics="true" pageId="11" pageNumber="12">tufari</emphasis>
|
||||
Lau Basin
|
||||
</taxonomicName>
|
||||
&
|
||||
<collectingCountry id="F32A9A8BFFC7472727FFFD957039FD90" box="[247,322,552,575]" name="Wallis and Futuna" pageId="11" pageNumber="12">Futuna</collectingCountry>
|
||||
Arc (
|
||||
<bibRefCitation id="EFACA7EAFFC747272675FD9A732AFD90" author="Poitrimol C & Thiebaut E & Daguin-Thiebaut C" box="[381,593,551,575]" pageId="11" pageNumber="12" refId="ref11104" refString="Poitrimol C, Thiebaut E, Daguin-Thiebaut C et al. Contrasted phylogeographic patterns of hydrothermal vent gastropods along South West Pacific: Woodlark Basin, a possible contact zone and / or stepping-stone. PLoS One 2022; 17: e 0275638. https: // doi. org / 10.1371 / journal. pone. 0275638" type="journal volume" year="2022">
|
||||
Poitrimol
|
||||
<emphasis id="B9490609FFC7472726EFFD95736CFD90" box="[487,535,551,575]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
|
||||
2022
|
||||
</bibRefCitation>
|
||||
) with moderate support (BPP = 0.84). These three taxa were weakly supported (BPP <0.70) as sister to the
|
||||
<taxonomicName id="4C3DA198FFC7472726BDFDDB7338FDD1" box="[437,579,614,638]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="myojinensis">
|
||||
<emphasis id="B9490609FFC7472726BDFDDB7338FDD1" box="[437,579,614,638]" italics="true" pageId="11" pageNumber="12">S. myojinensis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C3DA198FFC747272576FDDB7381FDD1" baseAuthorityName="Beck" baseAuthorityYear="1992" box="[638,762,614,638]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="tollmanni">
|
||||
<emphasis id="B9490609FFC747272576FDDB7381FDD1" box="[638,762,614,638]" italics="true" pageId="11" pageNumber="12">S. tollmanni</emphasis>
|
||||
</taxonomicName>
|
||||
pair.
|
||||
<taxonomicName id="4C3DA198FFC7472727AEFD387018FD32" box="[166,355,645,669]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="tufari">
|
||||
<emphasis id="B9490609FFC7472727AEFD387018FD32" box="[166,355,645,669]" italics="true" pageId="11" pageNumber="12">Shinkailepas tufari</emphasis>
|
||||
</taxonomicName>
|
||||
was found to be the closest relative of these five species, with moderate support (BPP = 0.83).
|
||||
</paragraph>
|
||||
<paragraph id="8B82DA1BFFC747272785FD797182FC5A" blockId="11.[112,763,144,1014]" pageId="11" pageNumber="12">
|
||||
From estimates of genetic divergence between
|
||||
<taxonomicName id="4C3DA198FFC747272570FD7971C6FD54" pageId="11" pageNumber="12">
|
||||
<emphasis id="B9490609FFC747272570FD797382FD73" box="[632,761,708,732]" italics="true" pageId="11" pageNumber="12">Shinkailepas</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
using the
|
||||
<emphasis id="B9490609FFC747272639FD597025FD54" box="[305,350,740,763]" italics="true" pageId="11" pageNumber="12">COI</emphasis>
|
||||
gene (Table 1), the K2P distance between
|
||||
<taxonomicName id="4C3DA198FFC7472727B2FCBE7040FCB5" box="[186,315,771,794]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="tiarasimia">
|
||||
<emphasis id="B9490609FFC7472727B2FCBE7040FCB5" box="[186,315,771,794]" italics="true" pageId="11" pageNumber="12">S. tiarasimia</emphasis>
|
||||
</taxonomicName>
|
||||
and other known congeners ranges from 6.65 to 8.86% and the same for
|
||||
<taxonomicName id="4C3DA198FFC7472726C7FC9F7309FC95" authorityName="Gu & Chen & Gao & Zhou & Sun" authorityYear="2025" box="[463,626,802,826]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="cornuthauma">
|
||||
<emphasis id="B9490609FFC7472726C7FC9F7309FC95" box="[463,626,802,826]" italics="true" pageId="11" pageNumber="12">S. cornuthauma</emphasis>
|
||||
</taxonomicName>
|
||||
ranges from 7.05 to 11.75%. This is roughly equivalent to the range observed among other
|
||||
<taxonomicName id="4C3DA198FFC74727266DFCDC7339FCD6" box="[357,578,865,889]" pageId="11" pageNumber="12">
|
||||
<emphasis id="B9490609FFC74727266DFCDC709DFCD6" box="[357,486,865,889]" italics="true" pageId="11" pageNumber="12">Shinkailepas</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
(5.50–11.42%), and above the lowest value of 5.50% between two described species (
|
||||
<emphasis id="B9490609FFC7472727DBFC1D70BEFC18" box="[211,453,927,951]" italics="true" pageId="11" pageNumber="12">
|
||||
<taxonomicName id="4C3DA198FFC7472727DBFC1D702DFC18" baseAuthorityName="Beck" baseAuthorityYear="1992" box="[211,342,927,951]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="tollmanni">S. tollmanni</taxonomicName>
|
||||
—
|
||||
<taxonomicName id="4C3DA198FFC747272679FC2270BEFC18" box="[369,453,927,951]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="tufari">S. tufari</taxonomicName>
|
||||
</emphasis>
|
||||
). These support the recognition of
|
||||
<taxonomicName id="4C3DA198FFC7472727D0FC027021FC79" box="[216,346,959,982]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="tiarasimia">
|
||||
<emphasis id="B9490609FFC7472727D0FC027021FC79" box="[216,346,959,982]" italics="true" pageId="11" pageNumber="12">S. tiarasimia</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C3DA198FFC74727269BFC02734DFC78" authorityName="Gu & Chen & Gao & Zhou & Sun" authorityYear="2025" box="[403,566,959,983]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="cornuthauma">
|
||||
<emphasis id="B9490609FFC74727269BFC02734DFC78" box="[403,566,959,983]" italics="true" pageId="11" pageNumber="12">S. cornuthauma</emphasis>
|
||||
</taxonomicName>
|
||||
as new species of
|
||||
<taxonomicName id="4C3DA198FFC747272779FC637189FC59" box="[113,242,990,1014]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B9490609FFC747272779FC637189FC59" box="[113,242,990,1014]" italics="true" pageId="11" pageNumber="12">Shinkailepas</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
691
data/03/94/6B/03946B0DFFCF472B24B3FE6075ADFCF6.xml
Normal file
691
data/03/94/6B/03946B0DFFCF472B24B3FE6075ADFCF6.xml
Normal file
|
|
@ -0,0 +1,691 @@
|
|||
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<mods:affiliation id="0DE3E58A06D3CF72B04C98048135D012">Key Laboratory of Marine Ecosystem Dynamics, Second Institute of Oceanography, Ministry of Natural Resources, Hangzhou 310012, China & Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai), Zhuhai 519082, China</mods:affiliation>
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|
||||
<heading id="D0CA6D77FFCF472F24B3FE607459FE57" bold="true" box="[955,1314,477,504]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="4" reason="2">
|
||||
<emphasis id="B9490609FFCF472F24B3FE607459FE57" bold="true" box="[955,1314,477,504]" pageId="3" pageNumber="4">
|
||||
<taxonomicName id="4C3DA198FFCF472F24B3FE6075BDFE57" authorityName="Gu & Chen & Gao & Zhou & Sun" authorityYear="2025" box="[955,1222,477,504]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="tiarasimia" status="sp. nov.">
|
||||
<emphasis id="B9490609FFCF472F24B3FE6075BDFE57" bold="true" box="[955,1222,477,504]" italics="true" pageId="3" pageNumber="4">Shinkailepas tiarasimia</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27ABB72FFCF472F23C3FE637459FE57" box="[1227,1314,478,504]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3278990FFCF472F233FFDB275DDFD86" box="[1079,1190,527,554]" pageId="3" pageNumber="4" type="description">
|
||||
<paragraph id="8B82DA1BFFCF472F233FFDB275DDFD86" blockId="3.[810,1460,527,716]" box="[1079,1190,527,554]" pageId="3" pageNumber="4">
|
||||
<emphasis id="B9490609FFCF472F233FFDB275DDFD86" bold="true" box="[1079,1190,527,554]" pageId="3" pageNumber="4">
|
||||
(
|
||||
<figureCitation id="1306C69EFFCF472F2349FDAD75E7FD85" box="[1089,1180,528,554]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStartId-0="4.[129,194,1543,1567]" captionStartId-1="5.[113,178,1532,1556]" captionStartId-2="6.[130,195,1604,1628]" captionTargetBox-0="[129,1473,144,1515]" captionTargetBox-1="[114,1458,144,1505]" captionTargetBox-2="[129,1473,144,1577]" captionTargetId-0="figure-171@4.[129,1473,144,1515]" captionTargetId-1="figure-148@5.[114,1458,144,1505]" captionTargetId-2="figure-150@6.[129,1473,144,1577]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="Figure 2. Shinkailepas tiarasimia, representative specimens. A, holotype (RSIO 38219), female, Wocan field.B, paratype 1 (RSIO 38220), female, Wocan field.C, paratype 2 (RSIO 38221), female, Wocan field.D, paratype 4 (RSIO 38223), male, Wocan field. E, paratype 6 (RSIO 38225), female, Wocan field, partly covered in rusty mineral deposits. F, damaged shell collected from Kairei field (NSMT-Mo 79568), completely covered in rusty mineral deposits. Black arrowhead indicates an egg case (enlarged in Fig. 3D); white arrowhead indicates the penis." captionText-1="Figure 3. Shinkailepas tiarasimia, external anatomy. A, dorsal aspect with shell removed, paratype 6 (RSIO 38225), female. B, enlarged dorsal view of the head with mantle edge removed, paratype 3 (RSIO 38222), female. C, operculum shown attached to dorsal aspect of foot with dashed line indicating the approximate outline, paratype 2 (RSIO 38221), female.D, egg capsule found on the shell of paratype 1 (RSIO 38220), female.Abbreviations:ag, albumen gland; cg, capsule gland; cl, cephalic lappet; ct, ctenidium; dg, digestive gland; e, eye; lsm, left shell muscle; me, mantle edge; nl, neck lobe; ol, oral lappet; ov, ovary; pc, pericardium; rs; radula sac; rsm, right shell muscle; sn, snout; st, stomach; t, cephalic tentacle." captionText-2="Figure 4.Shinkailepas tiarasimia, scanning electron micrographs. A, protoconch of a specimen from Kairei field (NSMT-Mo 79568). B, enlargement of part A showing fine pitting on the protoconch. C, shell microstructure, paratype 5 (RSIO 38224). D, shell surface sculpture, paratype 5 (RSIO 38224).E, overview of the radula, paratype 4 (RSIO 38223).F, close-up of the outer lateral teeth and marginal teeth, paratype 4 (RSIO 38223)." pageId="3" pageNumber="4">Figs 2–4</figureCitation>
|
||||
)
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3278990FFCF472F2422FD8B75E7FC8B" pageId="3" pageNumber="4" type="reference_group">
|
||||
<paragraph id="8B82DA1BFFCF472F2422FD8B7459FDE1" blockId="3.[810,1460,527,716]" box="[810,1314,566,590]" pageId="3" pageNumber="4">
|
||||
<treatmentCitationGroup id="AB2DFD35FFCF472F2422FD8B7459FDE1" box="[810,1314,566,590]" pageId="3" pageNumber="4">
|
||||
<treatmentCitation id="0A9CFC0AFFCF472F2422FD8B75B5FDE1" author="Van Dover CL & Humphris SE & Fornari D" box="[810,1230,566,590]" page="821" pageId="3" pageNumber="4" year="2001">
|
||||
<taxonomicName id="4C3DA198FFCF472F2422FD8B75B5FDE1" ID-CoL="7PGQ2" authority="Van Dover et al. (2001)" authorityName="Van Dover" authorityYear="2001" box="[810,1230,566,590]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B9490609FFCF472F2422FD8B72DEFDE1" box="[810,933,566,590]" italics="true" pageId="3" pageNumber="4">Shinkailepas</emphasis>
|
||||
<bibRefCitation id="EFACA7EAFFCF472F24A3FD8A75E2FDE1" author="Van Dover CL & Humphris SE & Fornari D" box="[939,1177,566,590]" pageId="3" pageNumber="4" pagination="818 - 23" refId="ref11829" refString="Van Dover CL, Humphris SE, Fornari D et al. Biogeography and ecological setting of Indian Ocean hydrothermal vents. Science 2001; 294: 818 - 23. https: // doi. org / 10.1126 / science. 1064574" type="journal article" year="2001">
|
||||
Van Dover
|
||||
<emphasis id="B9490609FFCF472F2314FD8A7530FDE1" box="[1052,1099,566,590]" italics="true" pageId="3" pageNumber="4">et al.</emphasis>
|
||||
(2001)
|
||||
</bibRefCitation>
|
||||
: 821
|
||||
</taxonomicName>
|
||||
</treatmentCitation>
|
||||
, table 2.
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
<paragraph id="8B82DA1BFFCF472F2422FDEB7564FD03" blockId="3.[810,1460,527,716]" pageId="3" pageNumber="4">
|
||||
<treatmentCitationGroup id="AB2DFD35FFCF472F2422FDEB7564FD03" pageId="3" pageNumber="4">
|
||||
<taxonomicName id="4C3DA198FFCF472F2422FDEB72BCFDC2" box="[810,967,598,622]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="undetermined">
|
||||
<emphasis id="B9490609FFCF472F2422FDEB72DEFDC1" box="[810,933,598,622]" italics="true" pageId="3" pageNumber="4">Shinkailepas</emphasis>
|
||||
sp.
|
||||
</taxonomicName>
|
||||
<treatmentCitation id="0A9CFC0AFFCF472F24C3FDEB743BFDC1" author="Watanabe H & Beedessee G" box="[971,1344,597,622]" page="16" pageId="3" pageNumber="4" year="2015">
|
||||
<bibRefCitation id="EFACA7EAFFCF472F24C3FDEB7462FDC2" author="Watanabe H & Beedessee G" box="[971,1305,597,622]" pageId="3" pageNumber="4" pagination="205 - 12" refId="ref11962" refString="Watanabe H, Beedessee G. Vent fauna on the Central Indian Ridge. In: Ishibashi J-I, Okino K, Sunamura M (eds), Subseafloor Biosphere Linked to Hydrothermal Systems: TAIGA Concept. Tokyo: Springer Japan, 2015, 205 - 12." type="book chapter" year="2015">Watanabe and Beedessee (2015)</bibRefCitation>
|
||||
: 16
|
||||
</treatmentCitation>
|
||||
, table 16.1.
|
||||
<taxonomicName id="4C3DA198FFCF472F2422FDC872B2FD23" box="[810,969,629,653]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="undetermined">
|
||||
<emphasis id="B9490609FFCF472F2422FDC872DEFD22" box="[810,933,629,653]" italics="true" pageId="3" pageNumber="4">Shinkailepas</emphasis>
|
||||
sp.
|
||||
</taxonomicName>
|
||||
CIR (Central Indian Ridge)
|
||||
<bibRefCitation id="EFACA7EAFFCF472F23F1FDC874D7FD22" author="Kim J & Son S-K & Kim D" box="[1273,1452,629,653]" pageId="3" pageNumber="4" refId="ref10727" refString="Kim J, Son S-K, Kim D et al. Discovery of active hydrothermal vent fields along the Central Indian Ridge, 8 - 12 ° S. Geochemistry, Geophysics, Geosystems 2020; 21: e 2020 GC 009058. https: // doi. org / 10.1029 / 2020 GC 009058" type="journal volume" year="2020">
|
||||
Kim
|
||||
<emphasis id="B9490609FFCF472F2223FDCB7420FD22" box="[1323,1371,629,653]" italics="true" pageId="3" pageNumber="4">et al.</emphasis>
|
||||
(2020)
|
||||
</bibRefCitation>
|
||||
: supplementary table S1.
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
<paragraph id="8B82DA1BFFCF472F2422FD0974C8FD63" blockId="3.[810,1460,527,716]" box="[810,1459,692,716]" pageId="3" pageNumber="4">
|
||||
<treatmentCitationGroup id="AB2DFD35FFCF472F2422FD0974C8FD63" box="[810,1459,692,716]" pageId="3" pageNumber="4">
|
||||
‘
|
||||
<taxonomicName id="4C3DA198FFCF472F2426FD0972BAFD63" ID-CoL="7NJ9L" authorityName="J. H. McLean" authorityYear="1988" box="[814,961,692,716]" class="Gastropoda" family="Lepetodrilidae" kingdom="Animalia" order="Lepetellida" pageId="3" pageNumber="4" phylum="Mollusca" rank="family">Lepetodrilidae</taxonomicName>
|
||||
fam. inc.’
|
||||
<treatmentCitation id="0A9CFC0AFFCF472F2315FD09746FFD63" author="Gerdes K & Kihara TC & Martinez Arbizu P" box="[1053,1300,692,716]" page="219" pageId="3" pageNumber="4" year="2021">
|
||||
<bibRefCitation id="EFACA7EAFFCF472F2315FD09759AFD63" author="Gerdes K & Kihara TC & Martinez Arbizu P" box="[1053,1249,692,716]" pageId="3" pageNumber="4" refId="ref10386" refString="Gerdes K, Kihara TC, Martinez Arbizu P et al. Megafauna of the German exploration licence area for seafloor massive sulphides along the Central and South East Indian Ridge (Indian Ocean). Biodiversity Data Journal 2021; 9: e 69955. https: // doi. org / 10.3897 / BDJ. 9. e 69955" type="journal volume" year="2021">
|
||||
Gerdes
|
||||
<emphasis id="B9490609FFCF472F2361FD0975EDFD63" box="[1129,1174,692,716]" italics="true" pageId="3" pageNumber="4">et al.</emphasis>
|
||||
(2021)
|
||||
</bibRefCitation>
|
||||
: 219
|
||||
</treatmentCitation>
|
||||
, fig. 201, table 3.
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
<paragraph id="8B82DA1BFFCF472F2422FD5075E7FC8B" blockId="3.[810,1459,749,804]" pageId="3" pageNumber="4">
|
||||
<emphasis id="B9490609FFCF472F2422FD507563FCAA" box="[810,1048,749,773]" italics="true" pageId="3" pageNumber="4">ZooBank registration:</emphasis>
|
||||
<uri id="FFACD619FFCF472F2330FD5075E7FC8B" pageId="3" pageNumber="4">
|
||||
urn:lsid:zoobank.org:act:
|
||||
<uuid id="FF9BE0CEFFCF472F2232FD5075E7FC8B" pageId="3" pageNumber="4">8B81A710- BFCB-4F5E-A588-1A1CA44949F2</uuid>
|
||||
</uri>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3278990FFCF472F2422FCFB751EFB38" pageId="3" pageNumber="4" type="diagnosis">
|
||||
<paragraph id="8B82DA1BFFCF472F2422FCFB751EFB38" blockId="3.[810,1460,837,1175]" pageId="3" pageNumber="4">
|
||||
<emphasis id="B9490609FFCF472F2422FCFB72EBFCF2" box="[810,912,838,861]" italics="true" pageId="3" pageNumber="4">Diagnosis:</emphasis>
|
||||
Medium-sized
|
||||
<taxonomicName id="4C3DA198FFCF472F2341FCFB75BFFCF1" box="[1097,1220,838,862]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B9490609FFCF472F2341FCFB75BFFCF1" box="[1097,1220,838,862]" italics="true" pageId="3" pageNumber="4">Shinkailepas</emphasis>
|
||||
</taxonomicName>
|
||||
(SL ≤
|
||||
<quantity id="4CC577FEFFCF472F2213FCF87411FCF1" box="[1307,1386,837,862]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="3" pageNumber="4" unit="mm" value="8.5">8.5 mm</quantity>
|
||||
) with apex slightly bent to left, positioned near posterior shell margin at about 90% from anterior edge. Shell sculpture cancellate with concentric sculpture equal in strength or stronger than radial ones, drawn out to form weak nodes where they cross. Protoconch bulbous, smooth except for micro-pustules, about 660 μm in size when measured from adult shell. Diameter of opercular nucleus (see
|
||||
<bibRefCitation id="EFACA7EAFFCF472F232FFB9C7438FB96" author="Fukumori H & Kano Y" box="[1063,1347,1057,1081]" pageId="3" pageNumber="4" pagination="213 - 27" refId="ref10235" refString="Fukumori H, Kano Y. Evolutionary ecology of settlement size in planktotrophic neritimorph gastropods. Marine Biology 2014; 161: 213 - 27. https: // doi. org / 10.1007 / s 00227 - 013 - 2330 - 5" type="journal article" year="2014">Fukumori and Kano 2014</bibRefCitation>
|
||||
) 475 μm. Ostracum translucent, with silvery sheen. Operculum entirely corneous, not calcified. Epipodial fold with 12–15 short, triangular, tentacular projections.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3278990FFCF47282422FB047347F955" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="materials_examined">
|
||||
<paragraph id="8B82DA1BFFCF472F2422FB0474D8FB7F" blockId="3.[810,1443,1208,1232]" box="[810,1443,1208,1232]" pageId="3" pageNumber="4">
|
||||
<emphasis id="B9490609FFCF472F2422FB0472D7FB7F" box="[810,940,1208,1232]" italics="true" pageId="3" pageNumber="4">
|
||||
<typeStatus id="548664B9FFCF472F2422FB047221FB7F" box="[810,858,1209,1232]" pageId="3" pageNumber="4">Type</typeStatus>
|
||||
locality:
|
||||
</emphasis>
|
||||
Wocan hydrothermal vent field, Carlsberg Ridge.
|
||||
</paragraph>
|
||||
<paragraph id="8B82DA1BFFCF472F2422FB4F7502F96F" blockId="3.[810,1460,1265,1728]" pageId="3" pageNumber="4">
|
||||
<emphasis id="B9490609FFCF472F2422FB4F72C0FAA5" box="[810,955,1266,1290]" italics="true" pageId="3" pageNumber="4">Type material:</emphasis>
|
||||
<materialsCitation id="3B55D046FFCF472F24CCFB4F7547FA28" collectedFrom="Wocan vent field" collectingDate="2017-03" collectionCode="R, V" elevation="2920" latitude="6.36" longLatPrecision="129" longitude="60.53" pageId="3" pageNumber="4" specimenCode="RSIO 38219" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
|
||||
<typeStatus id="548664B9FFCF472F24CCFB4F755CFAA5" box="[964,1063,1266,1290]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
|
||||
(
|
||||
<specimenCode id="DB9B7260FFCF472F233FFB4F75B9FAA5" box="[1079,1218,1266,1290]" collectionCode="RSIO" pageId="3" pageNumber="4">RSIO 38219</specimenCode>
|
||||
;
|
||||
<figureCitation id="1306C69EFFCF472F23C7FB4F7460FAA5" box="[1231,1307,1266,1290]" captionStart="Figure 2" captionStartId="4.[129,194,1543,1567]" captionTargetBox="[129,1473,144,1515]" captionTargetId="figure-171@4.[129,1473,144,1515]" captionTargetPageId="4" captionText="Figure 2. Shinkailepas tiarasimia, representative specimens. A, holotype (RSIO 38219), female, Wocan field.B, paratype 1 (RSIO 38220), female, Wocan field.C, paratype 2 (RSIO 38221), female, Wocan field.D, paratype 4 (RSIO 38223), male, Wocan field. E, paratype 6 (RSIO 38225), female, Wocan field, partly covered in rusty mineral deposits. F, damaged shell collected from Kairei field (NSMT-Mo 79568), completely covered in rusty mineral deposits. Black arrowhead indicates an egg case (enlarged in Fig. 3D); white arrowhead indicates the penis." pageId="3" pageNumber="4">Fig. 2A</figureCitation>
|
||||
),
|
||||
<specimenCount id="9D3B1192FFCF472F223CFB4F7402FAA5" box="[1332,1401,1266,1290]" pageId="3" pageNumber="4" type="female">female</specimenCount>
|
||||
, 95% ethanol; SL
|
||||
<quantity id="4CC577FEFFCF472F24A0FAAC7283FA87" box="[936,1016,1297,1321]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="3" pageNumber="4" unit="mm" value="5.3">5.3 mm</quantity>
|
||||
, SW
|
||||
<quantity id="4CC577FEFFCF472F2327FAAC75FBFA87" box="[1071,1152,1297,1321]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="3" pageNumber="4" unit="mm" value="4.2">4.2 mm</quantity>
|
||||
;
|
||||
<collectedFrom id="382199E6FFCF472F2383FAAC7443FA86" box="[1163,1336,1297,1321]" pageId="3" pageNumber="4">Wocan vent field</collectedFrom>
|
||||
(
|
||||
<geoCoordinate id="EE09BCDCFFCF472F224EFAAC74D4FA87" box="[1350,1455,1295,1321]" degrees="60" direction="east" minutes="31.8" orientation="longitude" pageId="3" pageNumber="4" precision="92" value="60.53">
|
||||
60°31.8
|
||||
<emphasis id="B9490609FFCF472F2290FAB274E5FA87" box="[1432,1438,1295,1320]" italics="true" pageId="3" pageNumber="4">ʹ</emphasis>
|
||||
E
|
||||
</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE09BCDCFFCF472F2422FA8D72F3FAE7" box="[810,904,1327,1352]" degrees="6" direction="north" minutes="21.6" orientation="latitude" pageId="3" pageNumber="4" precision="92" value="6.36">
|
||||
6°21.6
|
||||
<emphasis id="B9490609FFCF472F2464FA927209FAE7" box="[876,882,1327,1352]" italics="true" pageId="3" pageNumber="4">ʹ</emphasis>
|
||||
N
|
||||
</geoCoordinate>
|
||||
),
|
||||
<quantity id="4CC577FEFFCF472F24AAFA8D728DFAE7" box="[930,1014,1328,1352]" metricMagnitude="3" metricUnit="m" metricValue="2.92" pageId="3" pageNumber="4" unit="m" value="2920.0">
|
||||
<elevation id="00103D28FFCF472F24AAFA8D728DFAE7" box="[930,1014,1328,1352]" metricMagnitude="3" metricUnit="m" metricValue="2.92" pageId="3" pageNumber="4" unit="m" value="2920.0">2920 m</elevation>
|
||||
</quantity>
|
||||
deep, collected by a seven-function manipulator of HOV
|
||||
<emphasis id="B9490609FFCF472F24E4FAED7541FAC7" box="[1004,1082,1360,1384]" italics="true" pageId="3" pageNumber="4">Jiaolong</emphasis>
|
||||
on dive #129, R/V
|
||||
<emphasis id="B9490609FFCF472F2201FAED74CFFAC7" box="[1289,1460,1360,1384]" italics="true" pageId="3" pageNumber="4">Xiangyanghong 9</emphasis>
|
||||
cruise DY38,
|
||||
<date id="FF83FCDBFFCF472F24B2FAD27547FA28" box="[954,1084,1391,1415]" pageId="3" pageNumber="4" value="2017-03">
|
||||
<collectingDate id="EFC70533FFCF472F24B2FAD27547FA28" box="[954,1084,1391,1415]" pageId="3" pageNumber="4" value="2017-03">March 2017</collectingDate>
|
||||
</date>
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="3B55D046FFCF472F2340FAD27440FA09" collectedFrom="Wocan vent field" collectingDate="2017-03" collectionCode="RSIO" elevation="2920" latitude="6.36" longLatPrecision="129" longitude="60.53" pageId="3" pageNumber="4" specimenCode="RSIO 38220" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||||
<typeStatus id="548664B9FFCF472F2340FAD275D9FA28" box="[1096,1186,1391,1415]" pageId="3" pageNumber="4" type="paratype">Paratype</typeStatus>
|
||||
1 (
|
||||
<specimenCode id="DB9B7260FFCF472F23CFFAD27429FA28" box="[1223,1362,1391,1415]" collectionCode="RSIO" pageId="3" pageNumber="4">RSIO 38220</specimenCode>
|
||||
;
|
||||
<figureCitation id="1306C69EFFCF472F2257FAD274D6FA28" box="[1375,1453,1391,1415]" captionStart="Figure 2" captionStartId="4.[129,194,1543,1567]" captionTargetBox="[129,1473,144,1515]" captionTargetId="figure-171@4.[129,1473,144,1515]" captionTargetPageId="4" captionText="Figure 2. Shinkailepas tiarasimia, representative specimens. A, holotype (RSIO 38219), female, Wocan field.B, paratype 1 (RSIO 38220), female, Wocan field.C, paratype 2 (RSIO 38221), female, Wocan field.D, paratype 4 (RSIO 38223), male, Wocan field. E, paratype 6 (RSIO 38225), female, Wocan field, partly covered in rusty mineral deposits. F, damaged shell collected from Kairei field (NSMT-Mo 79568), completely covered in rusty mineral deposits. Black arrowhead indicates an egg case (enlarged in Fig. 3D); white arrowhead indicates the penis." pageId="3" pageNumber="4">Figs 2B</figureCitation>
|
||||
,
|
||||
<figureCitation id="1306C69EFFCF472F2422FA337231FA09" box="[810,842,1422,1446]" captionStart="Figure 3" captionStartId="5.[113,178,1532,1556]" captionTargetBox="[114,1458,144,1505]" captionTargetId="figure-148@5.[114,1458,144,1505]" captionTargetPageId="5" captionText="Figure 3. Shinkailepas tiarasimia, external anatomy. A, dorsal aspect with shell removed, paratype 6 (RSIO 38225), female. B, enlarged dorsal view of the head with mantle edge removed, paratype 3 (RSIO 38222), female. C, operculum shown attached to dorsal aspect of foot with dashed line indicating the approximate outline, paratype 2 (RSIO 38221), female.D, egg capsule found on the shell of paratype 1 (RSIO 38220), female.Abbreviations:ag, albumen gland; cg, capsule gland; cl, cephalic lappet; ct, ctenidium; dg, digestive gland; e, eye; lsm, left shell muscle; me, mantle edge; nl, neck lobe; ol, oral lappet; ov, ovary; pc, pericardium; rs; radula sac; rsm, right shell muscle; sn, snout; st, stomach; t, cephalic tentacle." pageId="3" pageNumber="4">3D</figureCitation>
|
||||
),
|
||||
<specimenCount id="9D3B1192FFCF472F2469FA3372DDFA09" box="[865,934,1422,1446]" pageId="3" pageNumber="4" type="female">female</specimenCount>
|
||||
, 95% ethanol; SL
|
||||
<quantity id="4CC577FEFFCF472F236CFA3375CFFA09" box="[1124,1204,1422,1446]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="3" pageNumber="4" unit="mm" value="6.5">6.5 mm</quantity>
|
||||
, SW
|
||||
<quantity id="4CC577FEFFCF472F23E3FA337440FA09" box="[1259,1339,1422,1446]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="3" pageNumber="4" unit="mm" value="5.4">5.4 mm</quantity>
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="3B55D046FFCF472F224DFA3272D1FA4B" collectedFrom="Wocan vent field" collectingDate="2017-03" collectionCode="RSIO" elevation="2920" latitude="6.36" longLatPrecision="129" longitude="60.53" pageId="3" pageNumber="4" specimenCode="RSIO 38221" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||||
<typeStatus id="548664B9FFCF472F224DFA3274E4FA08" box="[1349,1439,1423,1447]" pageId="3" pageNumber="4" type="paratype">Paratype</typeStatus>
|
||||
2 (RSIO 38221;
|
||||
<figureCitation id="1306C69EFFCF472F24C2FA137563FA69" box="[970,1048,1454,1478]" captionStart="Figure 2" captionStartId="4.[129,194,1543,1567]" captionTargetBox="[129,1473,144,1515]" captionTargetId="figure-171@4.[129,1473,144,1515]" captionTargetPageId="4" captionText="Figure 2. Shinkailepas tiarasimia, representative specimens. A, holotype (RSIO 38219), female, Wocan field.B, paratype 1 (RSIO 38220), female, Wocan field.C, paratype 2 (RSIO 38221), female, Wocan field.D, paratype 4 (RSIO 38223), male, Wocan field. E, paratype 6 (RSIO 38225), female, Wocan field, partly covered in rusty mineral deposits. F, damaged shell collected from Kairei field (NSMT-Mo 79568), completely covered in rusty mineral deposits. Black arrowhead indicates an egg case (enlarged in Fig. 3D); white arrowhead indicates the penis." pageId="3" pageNumber="4">Figs 2C</figureCitation>
|
||||
,
|
||||
<figureCitation id="1306C69EFFCF472F232CFA137538FA69" box="[1060,1091,1454,1478]" captionStart="Figure 3" captionStartId="5.[113,178,1532,1556]" captionTargetBox="[114,1458,144,1505]" captionTargetId="figure-148@5.[114,1458,144,1505]" captionTargetPageId="5" captionText="Figure 3. Shinkailepas tiarasimia, external anatomy. A, dorsal aspect with shell removed, paratype 6 (RSIO 38225), female. B, enlarged dorsal view of the head with mantle edge removed, paratype 3 (RSIO 38222), female. C, operculum shown attached to dorsal aspect of foot with dashed line indicating the approximate outline, paratype 2 (RSIO 38221), female.D, egg capsule found on the shell of paratype 1 (RSIO 38220), female.Abbreviations:ag, albumen gland; cg, capsule gland; cl, cephalic lappet; ct, ctenidium; dg, digestive gland; e, eye; lsm, left shell muscle; me, mantle edge; nl, neck lobe; ol, oral lappet; ov, ovary; pc, pericardium; rs; radula sac; rsm, right shell muscle; sn, snout; st, stomach; t, cephalic tentacle." pageId="3" pageNumber="4">3C</figureCitation>
|
||||
),
|
||||
<specimenCount id="9D3B1192FFCF472F2353FA1375DBFA69" box="[1115,1184,1454,1478]" pageId="3" pageNumber="4" type="female">female</specimenCount>
|
||||
, 95% ethanol; SL
|
||||
<quantity id="4CC577FEFFCF472F2257FA1074D4FA6A" box="[1375,1455,1453,1477]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="4" unit="mm" value="4.5">4.5 mm</quantity>
|
||||
, SW
|
||||
<quantity id="4CC577FEFFCF472F2453FA7072D1FA4B" box="[859,938,1485,1509]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="3" pageNumber="4" unit="mm" value="3.6">3.6 mm</quantity>
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="3B55D046FFCF472F24BDFA707200F98C" collectedFrom="Wocan vent field" collectingDate="2017-03" collectionCode="RSIO" elevation="2920" latitude="6.36" longLatPrecision="129" longitude="60.53" pageId="3" pageNumber="4" specimenCode="RSIO 38222" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||||
<typeStatus id="548664B9FFCF472F24BDFA707574FA4A" box="[949,1039,1485,1509]" pageId="3" pageNumber="4" type="paratype">Paratype</typeStatus>
|
||||
3 (
|
||||
<specimenCode id="DB9B7260FFCF472F233EFA7075B9FA4A" box="[1078,1218,1485,1509]" collectionCode="RSIO" pageId="3" pageNumber="4">RSIO 38222</specimenCode>
|
||||
;
|
||||
<figureCitation id="1306C69EFFCF472F23D8FA707461FA4A" box="[1232,1306,1485,1509]" captionStart="Figure 3" captionStartId="5.[113,178,1532,1556]" captionTargetBox="[114,1458,144,1505]" captionTargetId="figure-148@5.[114,1458,144,1505]" captionTargetPageId="5" captionText="Figure 3. Shinkailepas tiarasimia, external anatomy. A, dorsal aspect with shell removed, paratype 6 (RSIO 38225), female. B, enlarged dorsal view of the head with mantle edge removed, paratype 3 (RSIO 38222), female. C, operculum shown attached to dorsal aspect of foot with dashed line indicating the approximate outline, paratype 2 (RSIO 38221), female.D, egg capsule found on the shell of paratype 1 (RSIO 38220), female.Abbreviations:ag, albumen gland; cg, capsule gland; cl, cephalic lappet; ct, ctenidium; dg, digestive gland; e, eye; lsm, left shell muscle; me, mantle edge; nl, neck lobe; ol, oral lappet; ov, ovary; pc, pericardium; rs; radula sac; rsm, right shell muscle; sn, snout; st, stomach; t, cephalic tentacle." pageId="3" pageNumber="4">Fig. 3B</figureCitation>
|
||||
),
|
||||
<specimenCount id="9D3B1192FFCF472F223CFA707402FA4A" box="[1332,1401,1485,1509]" pageId="3" pageNumber="4" type="female">female</specimenCount>
|
||||
, 95% ethanol, radula dissected for SEM (
|
||||
<figureCitation id="1306C69EFFCF472F239DFA517582F9AB" box="[1173,1273,1516,1540]" captionStart="Figure 4" captionStartId="6.[130,195,1604,1628]" captionTargetBox="[129,1473,144,1577]" captionTargetId="figure-150@6.[129,1473,144,1577]" captionTargetPageId="6" captionText="Figure 4.Shinkailepas tiarasimia, scanning electron micrographs. A, protoconch of a specimen from Kairei field (NSMT-Mo 79568). B, enlargement of part A showing fine pitting on the protoconch. C, shell microstructure, paratype 5 (RSIO 38224). D, shell surface sculpture, paratype 5 (RSIO 38224).E, overview of the radula, paratype 4 (RSIO 38223).F, close-up of the outer lateral teeth and marginal teeth, paratype 4 (RSIO 38223)." pageId="3" pageNumber="4">Fig. 4E–F</figureCitation>
|
||||
); SL
|
||||
<quantity id="4CC577FEFFCF472F223CFA5174F8F9AB" box="[1332,1411,1516,1540]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="3" pageNumber="4" unit="mm" value="4.1">4.1 mm</quantity>
|
||||
, SW
|
||||
<quantity id="4CC577FEFFCF472F2423F9B17200F98C" box="[811,891,1548,1572]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="3" pageNumber="4" unit="mm" value="3.1">3.1 mm</quantity>
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="3B55D046FFCF472F248BF9B17559F9ED" collectedFrom="Wocan vent field" collectingDate="2017-03" collectionCode="RSIO" elevation="2920" latitude="6.36" longLatPrecision="129" longitude="60.53" pageId="3" pageNumber="4" specimenCode="RSIO 38223" specimenCount="1" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
|
||||
<typeStatus id="548664B9FFCF472F248BF9B172A6F98B" box="[899,989,1548,1572]" pageId="3" pageNumber="4" type="paratype">Paratype</typeStatus>
|
||||
4 (
|
||||
<specimenCode id="DB9B7260FFCF472F24F4F9B175FEF98B" box="[1020,1157,1548,1572]" collectionCode="RSIO" pageId="3" pageNumber="4">RSIO 38223</specimenCode>
|
||||
;
|
||||
<figureCitation id="1306C69EFFCF472F2387F9B175A1F98B" box="[1167,1242,1548,1572]" captionStart="Figure 2" captionStartId="4.[129,194,1543,1567]" captionTargetBox="[129,1473,144,1515]" captionTargetId="figure-171@4.[129,1473,144,1515]" captionTargetPageId="4" captionText="Figure 2. Shinkailepas tiarasimia, representative specimens. A, holotype (RSIO 38219), female, Wocan field.B, paratype 1 (RSIO 38220), female, Wocan field.C, paratype 2 (RSIO 38221), female, Wocan field.D, paratype 4 (RSIO 38223), male, Wocan field. E, paratype 6 (RSIO 38225), female, Wocan field, partly covered in rusty mineral deposits. F, damaged shell collected from Kairei field (NSMT-Mo 79568), completely covered in rusty mineral deposits. Black arrowhead indicates an egg case (enlarged in Fig. 3D); white arrowhead indicates the penis." pageId="3" pageNumber="4">Fig. 2D</figureCitation>
|
||||
),
|
||||
<specimenCount id="9D3B1192FFCF472F23F8F9B17458F98B" box="[1264,1315,1548,1572]" pageId="3" pageNumber="4" type="male">male</specimenCount>
|
||||
, 95% ethanol; SL
|
||||
<quantity id="4CC577FEFFCF472F2444F99672E7F9ED" box="[844,924,1579,1603]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="3" pageNumber="4" unit="mm" value="3.7">3.7 mm</quantity>
|
||||
, SW
|
||||
<quantity id="4CC577FEFFCF472F24D9F9967559F9ED" box="[977,1058,1579,1603]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="4" unit="mm" value="2.9">2.9 mm</quantity>
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="3B55D046FFCF472F2322F99672DDF92E" collectedFrom="Wocan vent field" collectingDate="2017-03" collectionCode="RSIO" elevation="2920" latitude="6.36" longLatPrecision="129" longitude="60.53" pageId="3" pageNumber="4" specimenCode="RSIO 38224" specimenCount="1" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
|
||||
<typeStatus id="548664B9FFCF472F2322F99675FFF9EC" box="[1066,1156,1579,1603]" pageId="3" pageNumber="4" type="paratype">Paratype</typeStatus>
|
||||
5 (
|
||||
<specimenCode id="DB9B7260FFCF472F23ACF9967456F9EC" box="[1188,1325,1579,1603]" collectionCode="RSIO" pageId="3" pageNumber="4">RSIO 38224</specimenCode>
|
||||
;
|
||||
<figureCitation id="1306C69EFFCF472F2230F99674D9F9EC" box="[1336,1442,1579,1603]" captionStart="Figure 4" captionStartId="6.[130,195,1604,1628]" captionTargetBox="[129,1473,144,1577]" captionTargetId="figure-150@6.[129,1473,144,1577]" captionTargetPageId="6" captionText="Figure 4.Shinkailepas tiarasimia, scanning electron micrographs. A, protoconch of a specimen from Kairei field (NSMT-Mo 79568). B, enlargement of part A showing fine pitting on the protoconch. C, shell microstructure, paratype 5 (RSIO 38224). D, shell surface sculpture, paratype 5 (RSIO 38224).E, overview of the radula, paratype 4 (RSIO 38223).F, close-up of the outer lateral teeth and marginal teeth, paratype 4 (RSIO 38223)." pageId="3" pageNumber="4">Fig. 4C–D</figureCitation>
|
||||
),
|
||||
<specimenCount id="9D3B1192FFCF472F2422F9F77226F9CD" box="[810,861,1610,1634]" pageId="3" pageNumber="4" type="male">male</specimenCount>
|
||||
, 95% ethanol, periostracum removed for SEM; SL
|
||||
<quantity id="4CC577FEFFCF472F2257F9F774D4F9CD" box="[1375,1455,1610,1634]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="3" pageNumber="4" unit="mm" value="5.9">5.9 mm</quantity>
|
||||
, SW
|
||||
<quantity id="4CC577FEFFCF472F245DF9D772DDF92E" box="[853,934,1642,1666]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="3" pageNumber="4" unit="mm" value="3.9">3.9 mm</quantity>
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="3B55D046FFCF472F24A5F9D7750EF96F" collectedFrom="Wocan vent field" collectingDate="2017-03" collectionCode="RSIO" elevation="2920" latitude="6.36" longLatPrecision="129" longitude="60.53" pageId="3" pageNumber="4" specimenCode="RSIO 38225" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||||
<typeStatus id="548664B9FFCF472F24A5F9D7757CF92D" box="[941,1031,1642,1666]" pageId="3" pageNumber="4" type="paratype">Paratype</typeStatus>
|
||||
6 (
|
||||
<specimenCode id="DB9B7260FFCF472F232EF9D775D5F92E" box="[1062,1198,1641,1666]" collectionCode="RSIO" pageId="3" pageNumber="4">RSIO 38225</specimenCode>
|
||||
;
|
||||
<figureCitation id="1306C69EFFCF472F23BFF9D77459F92D" box="[1207,1314,1642,1666]" captionStart="Figure 4" captionStartId="6.[130,195,1604,1628]" captionTargetBox="[129,1473,144,1577]" captionTargetId="figure-150@6.[129,1473,144,1577]" captionTargetPageId="6" captionText="Figure 4.Shinkailepas tiarasimia, scanning electron micrographs. A, protoconch of a specimen from Kairei field (NSMT-Mo 79568). B, enlargement of part A showing fine pitting on the protoconch. C, shell microstructure, paratype 5 (RSIO 38224). D, shell surface sculpture, paratype 5 (RSIO 38224).E, overview of the radula, paratype 4 (RSIO 38223).F, close-up of the outer lateral teeth and marginal teeth, paratype 4 (RSIO 38223)." pageId="3" pageNumber="4">Fig. 4C–D</figureCitation>
|
||||
),
|
||||
<specimenCount id="9D3B1192FFCF472F2230F9D77406F92D" box="[1336,1405,1642,1666]" pageId="3" pageNumber="4" type="female">female</specimenCount>
|
||||
, 95% ethanol (
|
||||
<figureCitation id="1306C69EFFCF472F2481F93472AFF90E" box="[905,980,1673,1697]" captionStart="Figure 3" captionStartId="5.[113,178,1532,1556]" captionTargetBox="[114,1458,144,1505]" captionTargetId="figure-148@5.[114,1458,144,1505]" captionTargetPageId="5" captionText="Figure 3. Shinkailepas tiarasimia, external anatomy. A, dorsal aspect with shell removed, paratype 6 (RSIO 38225), female. B, enlarged dorsal view of the head with mantle edge removed, paratype 3 (RSIO 38222), female. C, operculum shown attached to dorsal aspect of foot with dashed line indicating the approximate outline, paratype 2 (RSIO 38221), female.D, egg capsule found on the shell of paratype 1 (RSIO 38220), female.Abbreviations:ag, albumen gland; cg, capsule gland; cl, cephalic lappet; ct, ctenidium; dg, digestive gland; e, eye; lsm, left shell muscle; me, mantle edge; nl, neck lobe; ol, oral lappet; ov, ovary; pc, pericardium; rs; radula sac; rsm, right shell muscle; sn, snout; st, stomach; t, cephalic tentacle." pageId="3" pageNumber="4">Fig. 3A</figureCitation>
|
||||
); SL
|
||||
<quantity id="4CC577FEFFCF472F231AF9347519F90F" box="[1042,1122,1673,1697]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="3" pageNumber="4" unit="mm" value="4.8">4.8 mm</quantity>
|
||||
, SW
|
||||
<quantity id="4CC577FEFFCF472F2392F9347590F90F" box="[1178,1259,1673,1697]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="3" pageNumber="4" unit="mm" value="3.7">3.7 mm</quantity>
|
||||
. All paratypes with same collecting data as holotype
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8B82DA1BFFCF47282422F95F7347F955" blockId="3.[810,1461,1761,1974]" lastBlockId="4.[129,777,1698,1786]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
|
||||
<emphasis id="B9490609FFCF472F2422F95F7296F955" box="[810,1005,1762,1786]" italics="true" pageId="3" pageNumber="4">Material examined:</emphasis>
|
||||
One shell (NSMT-Mo 79568;
|
||||
<figureCitation id="1306C69EFFCF472F2233F95F74D9F955" box="[1339,1442,1762,1786]" captionStart="Figure 4" captionStartId="6.[130,195,1604,1628]" captionTargetBox="[129,1473,144,1577]" captionTargetId="figure-150@6.[129,1473,144,1577]" captionTargetPageId="6" captionText="Figure 4.Shinkailepas tiarasimia, scanning electron micrographs. A, protoconch of a specimen from Kairei field (NSMT-Mo 79568). B, enlargement of part A showing fine pitting on the protoconch. C, shell microstructure, paratype 5 (RSIO 38224). D, shell surface sculpture, paratype 5 (RSIO 38224).E, overview of the radula, paratype 4 (RSIO 38223).F, close-up of the outer lateral teeth and marginal teeth, paratype 4 (RSIO 38223)." pageId="3" pageNumber="4">Fig. 4A–B</figureCitation>
|
||||
), periostracum of the protoconch removed for SEM, 99% ethanol; SL
|
||||
<quantity id="4CC577FEFFCF472F24ACF89D728FF897" box="[932,1012,1824,1848]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="3" pageNumber="4" unit="mm" value="8.4">8.4 mm</quantity>
|
||||
, SW
|
||||
<quantity id="4CC577FEFFCF472F232EF89D750CF897" box="[1062,1143,1824,1848]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="3" pageNumber="4" unit="mm" value="6.1">6.1 mm</quantity>
|
||||
; Kairei vent field (
|
||||
<geoCoordinate id="EE09BCDCFFCF472F2224F89D74D4F897" box="[1324,1455,1823,1848]" degrees="70" direction="east" minutes="2.4187" orientation="longitude" pageId="3" pageNumber="4" precision="1" value="70.04031">
|
||||
70°2.4187
|
||||
<emphasis id="B9490609FFCF472F2290F8A274E5F897" box="[1432,1438,1823,1848]" italics="true" pageId="3" pageNumber="4">ʹ</emphasis>
|
||||
E
|
||||
</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE09BCDCFFCF472F2422F88272CDF8F7" box="[810,950,1854,1880]" degrees="25" direction="south" minutes="19.2315" orientation="latitude" pageId="3" pageNumber="4" precision="1" value="-25.320524">
|
||||
25°19.2315
|
||||
<emphasis id="B9490609FFCF472F24A8F88372DDF8F8" box="[928,934,1854,1879]" italics="true" pageId="3" pageNumber="4">ʹ</emphasis>
|
||||
S
|
||||
</geoCoordinate>
|
||||
),
|
||||
<quantity id="4CC577FEFFCF472F24D8F8FD755CF8F8" box="[976,1063,1856,1880]" metricMagnitude="3" metricUnit="m" metricValue="2.424" pageId="3" pageNumber="4" unit="m" value="2424.0">
|
||||
<elevation id="00103D28FFCF472F24D8F8FD755CF8F8" box="[976,1063,1856,1880]" metricMagnitude="3" metricUnit="m" metricValue="2.424" pageId="3" pageNumber="4" unit="m" value="2424.0">2424 m</elevation>
|
||||
</quantity>
|
||||
deep, collected by suction sampler mounted on HOV
|
||||
<emphasis id="B9490609FFCF472F24FCF8E27501F8D8" box="[1012,1146,1887,1911]" italics="true" pageId="3" pageNumber="4">
|
||||
<taxonomicName id="4C3DA198FFCF472F24FCF8E27544F8D8" authorityName="Miura & Laubier" authorityYear="1990" box="[1012,1087,1887,1911]" class="Polychaeta" family="Chrysopetalidae" genus="Shinkai" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">Shinkai</taxonomicName>
|
||||
6500
|
||||
</emphasis>
|
||||
on dive #1449,
|
||||
<collectionCode id="ED2C42DEFFCF472F222AF8E2744FF8D8" box="[1314,1332,1887,1911]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="4">R</collectionCode>
|
||||
/
|
||||
<collectionCode id="ED2C42DEFFCF472F2234F8E27434F8D8" box="[1340,1359,1887,1911]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="4" type="Museum">V</collectionCode>
|
||||
<emphasis id="B9490609FFCF472F2251F8E274CEF8D8" box="[1369,1461,1887,1911]" italics="true" pageId="3" pageNumber="4">Yokosuka</emphasis>
|
||||
cruise YK16-E02,
|
||||
<date id="FF83FCDBFFCF472F24E3F8C3750FF839" box="[1003,1140,1918,1942]" pageId="3" pageNumber="4" value="2016-02-13">
|
||||
<collectingDate id="EFC70533FFCF472F24E3F8C3750FF839" box="[1003,1140,1918,1942]" pageId="3" pageNumber="4" value="2016-02-13">Feb 13 2016</collectingDate>
|
||||
</date>
|
||||
. Lot of
|
||||
<specimenCount id="9D3B1192FFCF472F23C4F8C3741FF839" box="[1228,1380,1918,1942]" count="5" pageId="3" pageNumber="4" type="generic">five specimens</specimenCount>
|
||||
(
|
||||
<specimenCode id="DB9B7260FFCF472F227CF8C3720BF819" collectionCode="RSIO" pageId="3" pageNumber="4">RSIO 38226</specimenCode>
|
||||
), 95% ethanol, up to SL
|
||||
<quantity id="4CC577FEFFCF472F2389F82075AAF81A" box="[1153,1233,1949,1973]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="3" pageNumber="4" unit="mm" value="5.1">5.1 mm</quantity>
|
||||
, SW
|
||||
<quantity id="4CC577FEFFCF472F2205F8237425F81A" box="[1293,1374,1950,1974]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="3" pageNumber="4" unit="mm" value="3.9">3.9 mm</quantity>
|
||||
; Wocan vent field (
|
||||
<geoCoordinate id="EE09BCDCFFC8472827FBF9197027F914" box="[243,348,1698,1724]" degrees="60" direction="east" minutes="31.8" orientation="longitude" pageId="4" pageNumber="5" precision="92" value="60.53">
|
||||
60°31.8
|
||||
<emphasis id="B9490609FFC84728264DF91F7030F914" box="[325,331,1698,1723]" italics="true" pageId="4" pageNumber="5">ʹ</emphasis>
|
||||
E
|
||||
</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE09BCDCFFC847282660F91970BCF913" box="[360,455,1698,1724]" degrees="6" direction="north" minutes="21.6" orientation="latitude" pageId="4" pageNumber="5" precision="92" value="6.36">
|
||||
6°21.6
|
||||
<emphasis id="B9490609FFC8472826A2F91F70CBF914" box="[426,432,1698,1723]" italics="true" pageId="4" pageNumber="5">ʹ</emphasis>
|
||||
N
|
||||
</geoCoordinate>
|
||||
),
|
||||
<quantity id="4CC577FEFFC8472826D5F9197354F914" box="[477,559,1700,1724]" metricMagnitude="3" metricUnit="m" metricValue="2.9699999999999998" pageId="4" pageNumber="5" unit="m" value="2970.0">
|
||||
<elevation id="00103D28FFC8472826D5F9197354F914" box="[477,559,1700,1724]" metricMagnitude="3" metricUnit="m" metricValue="2.9699999999999998" pageId="4" pageNumber="5" unit="m" value="2970.0">2970 m</elevation>
|
||||
</quantity>
|
||||
deep, collected by a seven-function manipulator of HOV
|
||||
<emphasis id="B9490609FFC8472826F1F97E733CF974" box="[505,583,1731,1755]" italics="true" pageId="4" pageNumber="5">Jiaolong</emphasis>
|
||||
on dive #125,
|
||||
<collectionCode id="ED2C42DEFFC8472825D3F97E7396F974" box="[731,749,1731,1755]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="5">R</collectionCode>
|
||||
/
|
||||
<collectionCode id="ED2C42DEFFC8472825FEF97E7272F974" box="[758,777,1731,1755]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="5" type="Museum">V</collectionCode>
|
||||
<emphasis id="B9490609FFC847282789F95F7051F955" box="[129,298,1762,1786]" italics="true" pageId="4" pageNumber="5">Xiangyanghong 9</emphasis>
|
||||
cruise DY38,
|
||||
<date id="FF83FCDBFFC8472826B1F95F734CF955" box="[441,567,1762,1786]" pageId="4" pageNumber="5" value="2017-03">
|
||||
<collectingDate id="EFC70533FFC8472826B1F95F734CF955" box="[441,567,1762,1786]" pageId="4" pageNumber="5" value="2017-03">March 2017</collectingDate>
|
||||
</date>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF428A93FFC847282789F9BA74B9F9DC" pageId="4" pageNumber="5" startId="4.[129,194,1543,1567]" targetBox="[129,1473,144,1515]" targetPageId="4" targetType="figure">
|
||||
<paragraph id="8B82DA1BFFC847282789F9BA74B9F9DC" blockId="4.[129,1474,1543,1652]" pageId="4" pageNumber="5">
|
||||
<emphasis id="B9490609FFC847282789F9BA71A3F9B0" bold="true" box="[129,216,1543,1567]" pageId="4" pageNumber="5">Figure 2.</emphasis>
|
||||
<taxonomicName id="4C3DA198FFC8472827D5F9BA70D4F9B0" authorityName="Gu & Chen & Gao & Zhou & Sun" authorityYear="2025" box="[221,431,1543,1567]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="tiarasimia">
|
||||
<emphasis id="B9490609FFC8472827D5F9BA70D4F9B0" box="[221,431,1543,1567]" italics="true" pageId="4" pageNumber="5">Shinkailepas tiarasimia</emphasis>
|
||||
</taxonomicName>
|
||||
, representative specimens. A, holotype (RSIO 38219), female, Wocan field. B, paratype 1 (RSIO 38220), female, Wocan field. C, paratype 2 (RSIO 38221), female, Wocan field. D, paratype 4 (RSIO 38223), male, Wocan field. E, paratype 6 (RSIO 38225), female, Wocan field, partly covered in rusty mineral deposits. F, damaged shell collected from Kairei field (NSMT-Mo 79568), completely covered in rusty mineral deposits. Black arrowhead indicates an egg case (enlarged in Fig. 3D); white arrowhead indicates the penis.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="C3278990FFC8472B2789F8A1737FFA80" lastPageId="7" lastPageNumber="8" pageId="4" pageNumber="5" type="description">
|
||||
<paragraph id="8B82DA1BFFC847292789F8A175B7F8E8" blockId="4.[128,778,1820,1969]" lastBlockId="5.[810,1461,1744,1956]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
|
||||
<emphasis id="B9490609FFC847282789F8A17182F89C" box="[129,249,1820,1843]" italics="true" pageId="4" pageNumber="5">Description:</emphasis>
|
||||
Shell (
|
||||
<figureCitation id="1306C69EFFC84728265BF8A170EEF89B" box="[339,405,1820,1844]" captionStart="Figure 2" captionStartId="4.[129,194,1543,1567]" captionTargetBox="[129,1473,144,1515]" captionTargetId="figure-171@4.[129,1473,144,1515]" captionTargetPageId="4" captionText="Figure 2. Shinkailepas tiarasimia, representative specimens. A, holotype (RSIO 38219), female, Wocan field.B, paratype 1 (RSIO 38220), female, Wocan field.C, paratype 2 (RSIO 38221), female, Wocan field.D, paratype 4 (RSIO 38223), male, Wocan field. E, paratype 6 (RSIO 38225), female, Wocan field, partly covered in rusty mineral deposits. F, damaged shell collected from Kairei field (NSMT-Mo 79568), completely covered in rusty mineral deposits. Black arrowhead indicates an egg case (enlarged in Fig. 3D); white arrowhead indicates the penis." pageId="4" pageNumber="5">Fig. 2</figureCitation>
|
||||
) medium-sized for genus, nearly limpet-form as adults but conspicuously coiled at juvenile stage. Aperture oval, 1.1–1.3 times as long as wide. Height about one-third of length. Apex overhanging, close to shell edge at approximately 10% of length from posterior margin, recurved slightly to left side. Shell sculpture cancellate from crossing of concentric sculpture with radials, forming low protrusions where two directions meet. Concentric sculptures stronger than or equal in strength to radials. Shell margin nearly flat to slightly raised (
|
||||
<figureCitation id="1306C69EFFC8472824DFF89D7547F896" box="[983,1084,1824,1849]" captionStart="Figure 2" captionStartId="4.[129,194,1543,1567]" captionTargetBox="[129,1473,144,1515]" captionTargetId="figure-171@4.[129,1473,144,1515]" captionTargetPageId="4" captionText="Figure 2. Shinkailepas tiarasimia, representative specimens. A, holotype (RSIO 38219), female, Wocan field.B, paratype 1 (RSIO 38220), female, Wocan field.C, paratype 2 (RSIO 38221), female, Wocan field.D, paratype 4 (RSIO 38223), male, Wocan field. E, paratype 6 (RSIO 38225), female, Wocan field, partly covered in rusty mineral deposits. F, damaged shell collected from Kairei field (NSMT-Mo 79568), completely covered in rusty mineral deposits. Black arrowhead indicates an egg case (enlarged in Fig. 3D); white arrowhead indicates the penis." pageId="4" pageNumber="5">Fig. 2A–B</figureCitation>
|
||||
). Septum (‘deck’) large, conspicuous, occupying anterior one-fourth of aperture at posterior (
|
||||
<figureCitation id="1306C69EFFC847282294F8FD720EF8D8" captionStart="Figure 2" captionStartId="4.[129,194,1543,1567]" captionTargetBox="[129,1473,144,1515]" captionTargetId="figure-171@4.[129,1473,144,1515]" captionTargetPageId="4" captionText="Figure 2. Shinkailepas tiarasimia, representative specimens. A, holotype (RSIO 38219), female, Wocan field.B, paratype 1 (RSIO 38220), female, Wocan field.C, paratype 2 (RSIO 38221), female, Wocan field.D, paratype 4 (RSIO 38223), male, Wocan field. E, paratype 6 (RSIO 38225), female, Wocan field, partly covered in rusty mineral deposits. F, damaged shell collected from Kairei field (NSMT-Mo 79568), completely covered in rusty mineral deposits. Black arrowhead indicates an egg case (enlarged in Fig. 3D); white arrowhead indicates the penis." pageId="4" pageNumber="5">Fig. 2B, E</figureCitation>
|
||||
). Inner shell surface carry numerous shell pores, absent on septum. Ostracum thin, translucent-white with silvery sheen. Shell microstructure (
|
||||
<figureCitation id="1306C69EFFC847282362F82375CCF819" box="[1130,1207,1950,1974]" captionStart="Figure 4" captionStartId="6.[130,195,1604,1628]" captionTargetBox="[129,1473,144,1577]" captionTargetId="figure-150@6.[129,1473,144,1577]" captionTargetPageId="6" captionText="Figure 4.Shinkailepas tiarasimia, scanning electron micrographs. A, protoconch of a specimen from Kairei field (NSMT-Mo 79568). B, enlargement of part A showing fine pitting on the protoconch. C, shell microstructure, paratype 5 (RSIO 38224). D, shell surface sculpture, paratype 5 (RSIO 38224).E, overview of the radula, paratype 4 (RSIO 38223).F, close-up of the outer lateral teeth and marginal teeth, paratype 4 (RSIO 38223)." pageId="4" pageNumber="5">Fig. 4C</figureCitation>
|
||||
) of three layers dorsal to myostracum, outermost homogeneous layer thin, followed by another thin granular layer, innermost crossed-lamellar layer thick. Periostracum thin, translucent, light to bright green (
|
||||
<figureCitation id="1306C69EFFC9472925DCF8B27105F8E8" captionStart="Figure 2" captionStartId="4.[129,194,1543,1567]" captionTargetBox="[129,1473,144,1515]" captionTargetId="figure-171@4.[129,1473,144,1515]" captionTargetPageId="4" captionText="Figure 2. Shinkailepas tiarasimia, representative specimens. A, holotype (RSIO 38219), female, Wocan field.B, paratype 1 (RSIO 38220), female, Wocan field.C, paratype 2 (RSIO 38221), female, Wocan field.D, paratype 4 (RSIO 38223), male, Wocan field. E, paratype 6 (RSIO 38225), female, Wocan field, partly covered in rusty mineral deposits. F, damaged shell collected from Kairei field (NSMT-Mo 79568), completely covered in rusty mineral deposits. Black arrowhead indicates an egg case (enlarged in Fig. 3D); white arrowhead indicates the penis." pageId="5" pageNumber="6">Fig. 2</figureCitation>
|
||||
A-D), typically with some sulphide mineral deposits on top (
|
||||
<figureCitation id="1306C69EFFC947292774F8F371B0F8C9" box="[124,203,1870,1894]" captionStart="Figure 3" captionStartId="5.[113,178,1532,1556]" captionTargetBox="[114,1458,144,1505]" captionTargetId="figure-148@5.[114,1458,144,1505]" captionTargetPageId="5" captionText="Figure 3. Shinkailepas tiarasimia, external anatomy. A, dorsal aspect with shell removed, paratype 6 (RSIO 38225), female. B, enlarged dorsal view of the head with mantle edge removed, paratype 3 (RSIO 38222), female. C, operculum shown attached to dorsal aspect of foot with dashed line indicating the approximate outline, paratype 2 (RSIO 38221), female.D, egg capsule found on the shell of paratype 1 (RSIO 38220), female.Abbreviations:ag, albumen gland; cg, capsule gland; cl, cephalic lappet; ct, ctenidium; dg, digestive gland; e, eye; lsm, left shell muscle; me, mantle edge; nl, neck lobe; ol, oral lappet; ov, ovary; pc, pericardium; rs; radula sac; rsm, right shell muscle; sn, snout; st, stomach; t, cephalic tentacle." pageId="5" pageNumber="6">Fig. 3D</figureCitation>
|
||||
), in extreme cases shell may appear to become reddish in appearance due to rusty deposits, as known for some other
|
||||
<taxonomicName id="4C3DA198FFC9472927B8F8317006F80B" box="[176,381,1932,1956]" pageId="5" pageNumber="6">
|
||||
<emphasis id="B9490609FFC9472927B8F8317056F80B" box="[176,301,1932,1956]" italics="true" pageId="5" pageNumber="6">Shinkailepas</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
(
|
||||
<bibRefCitation id="EFACA7EAFFC947292687F8317313F80A" author="Fukumori H & Yahagi T & Waren A" box="[399,616,1932,1957]" pageId="5" pageNumber="6" pagination="636 - 55" refId="ref10278" refString="Fukumori H, Yahagi T, Waren A et al. Amended generic classification of themarinegastropodfamilyPhenacolepadidae: transitionsfromsnails to limpets and shallow-water to deep-sea hydrothermal vents and cold seeps. Zoological Journal of the Linnean Society 2019; 185: 636 - 55. https: // doi. org / 10.1093 / zoolinnean / zly 078" type="journal article" year="2019">
|
||||
Fukumori
|
||||
<emphasis id="B9490609FFC9472926F6F8307355F80B" box="[510,558,1932,1956]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
|
||||
2019
|
||||
</bibRefCitation>
|
||||
). Protoconch (
|
||||
<figureCitation id="1306C69EFFC94729243DF96D72FBF946" box="[821,896,1744,1769]" captionStart="Figure 2" captionStartId="4.[129,194,1543,1567]" captionTargetBox="[129,1473,144,1515]" captionTargetId="figure-171@4.[129,1473,144,1515]" captionTargetPageId="4" captionText="Figure 2. Shinkailepas tiarasimia, representative specimens. A, holotype (RSIO 38219), female, Wocan field.B, paratype 1 (RSIO 38220), female, Wocan field.C, paratype 2 (RSIO 38221), female, Wocan field.D, paratype 4 (RSIO 38223), male, Wocan field. E, paratype 6 (RSIO 38225), female, Wocan field, partly covered in rusty mineral deposits. F, damaged shell collected from Kairei field (NSMT-Mo 79568), completely covered in rusty mineral deposits. Black arrowhead indicates an egg case (enlarged in Fig. 3D); white arrowhead indicates the penis." pageId="5" pageNumber="6">Figs 2F</figureCitation>
|
||||
,
|
||||
<figureCitation id="1306C69EFFC947292484F96C72D1F946" box="[908,938,1745,1769]" captionStart="Figure 4" captionStartId="6.[130,195,1604,1628]" captionTargetBox="[129,1473,144,1577]" captionTargetId="figure-150@6.[129,1473,144,1577]" captionTargetPageId="6" captionText="Figure 4.Shinkailepas tiarasimia, scanning electron micrographs. A, protoconch of a specimen from Kairei field (NSMT-Mo 79568). B, enlargement of part A showing fine pitting on the protoconch. C, shell microstructure, paratype 5 (RSIO 38224). D, shell surface sculpture, paratype 5 (RSIO 38224).E, overview of the radula, paratype 4 (RSIO 38223).F, close-up of the outer lateral teeth and marginal teeth, paratype 4 (RSIO 38223)." pageId="5" pageNumber="6">4A</figureCitation>
|
||||
) white, covered by dark purple-red periostracum, bulbous, approximately 660 μm in length when measuring exposed part from adult shells, completely smooth except for minute dimples of 1–2 μm in size (
|
||||
<figureCitation id="1306C69EFFC94729237CF89375C0F8E8" box="[1140,1211,1838,1863]" captionStart="Figure 4" captionStartId="6.[130,195,1604,1628]" captionTargetBox="[129,1473,144,1577]" captionTargetId="figure-150@6.[129,1473,144,1577]" captionTargetPageId="6" captionText="Figure 4.Shinkailepas tiarasimia, scanning electron micrographs. A, protoconch of a specimen from Kairei field (NSMT-Mo 79568). B, enlargement of part A showing fine pitting on the protoconch. C, shell microstructure, paratype 5 (RSIO 38224). D, shell surface sculpture, paratype 5 (RSIO 38224).E, overview of the radula, paratype 4 (RSIO 38223).F, close-up of the outer lateral teeth and marginal teeth, paratype 4 (RSIO 38223)." pageId="5" pageNumber="6">Fig. 4B</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="DF428A93FFC947292779FA41705DF90E" pageId="5" pageNumber="6" startId="5.[113,178,1532,1556]" targetBox="[114,1458,144,1505]" targetPageId="5" targetType="figure">
|
||||
<paragraph id="8B82DA1BFFC947292779FA41705DF90E" blockId="5.[113,1456,1532,1697]" pageId="5" pageNumber="6">
|
||||
<emphasis id="B9490609FFC947292779FA4171B2F9BB" bold="true" box="[113,201,1532,1556]" pageId="5" pageNumber="6">Figure 3.</emphasis>
|
||||
<taxonomicName id="4C3DA198FFC9472927C6FA4070DBF9BB" authorityName="Gu & Chen & Gao & Zhou & Sun" authorityYear="2025" box="[206,416,1533,1557]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="tiarasimia">
|
||||
<emphasis id="B9490609FFC9472927C6FA4070DBF9BB" box="[206,416,1533,1557]" italics="true" pageId="5" pageNumber="6">Shinkailepas tiarasimia</emphasis>
|
||||
</taxonomicName>
|
||||
, external anatomy. A, dorsal aspect with shell removed, paratype 6 (RSIO 38225), female. B, enlarged dorsal view of the head with mantle edge removed, paratype 3 (RSIO 38222), female. C, operculum shown attached to dorsal aspect of foot with dashed line indicating the approximate outline, paratype 2 (RSIO 38221), female. D, egg capsule found on the shell of paratype 1 (RSIO 38220), female. Abbreviations: ag, albumen gland; cg, capsule gland; cl, cephalic lappet; ct, ctenidium; dg, digestive gland; e, eye; lsm, left shell muscle; me, mantle edge; nl, neck lobe; ol, oral lappet; ov, ovary; pc, pericardium; rs; radula sac; rsm, right shell muscle; sn, snout; st, stomach; t, cephalic tentacle.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B82DA1BFFC9472B244DF8F3731AFDAE" blockId="5.[810,1461,1744,1956]" lastBlockId="7.[113,764,144,1327]" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6">
|
||||
External anatomy (
|
||||
<figureCitation id="1306C69EFFC947292312F8F375EBF8C9" box="[1050,1168,1870,1894]" captionStart="Figure 2" captionStartId="4.[129,194,1543,1567]" captionTargetBox="[129,1473,144,1515]" captionTargetId="figure-171@4.[129,1473,144,1515]" captionTargetPageId="4" captionText="Figure 2. Shinkailepas tiarasimia, representative specimens. A, holotype (RSIO 38219), female, Wocan field.B, paratype 1 (RSIO 38220), female, Wocan field.C, paratype 2 (RSIO 38221), female, Wocan field.D, paratype 4 (RSIO 38223), male, Wocan field. E, paratype 6 (RSIO 38225), female, Wocan field, partly covered in rusty mineral deposits. F, damaged shell collected from Kairei field (NSMT-Mo 79568), completely covered in rusty mineral deposits. Black arrowhead indicates an egg case (enlarged in Fig. 3D); white arrowhead indicates the penis." pageId="5" pageNumber="6">Figs 2C–D</figureCitation>
|
||||
,
|
||||
<figureCitation id="1306C69EFFC9472923ABF8F3759AF8C9" box="[1187,1249,1870,1894]" captionStart="Figure 3" captionStartId="5.[113,178,1532,1556]" captionTargetBox="[114,1458,144,1505]" captionTargetId="figure-148@5.[114,1458,144,1505]" captionTargetPageId="5" captionText="Figure 3. Shinkailepas tiarasimia, external anatomy. A, dorsal aspect with shell removed, paratype 6 (RSIO 38225), female. B, enlarged dorsal view of the head with mantle edge removed, paratype 3 (RSIO 38222), female. C, operculum shown attached to dorsal aspect of foot with dashed line indicating the approximate outline, paratype 2 (RSIO 38221), female.D, egg capsule found on the shell of paratype 1 (RSIO 38220), female.Abbreviations:ag, albumen gland; cg, capsule gland; cl, cephalic lappet; ct, ctenidium; dg, digestive gland; e, eye; lsm, left shell muscle; me, mantle edge; nl, neck lobe; ol, oral lappet; ov, ovary; pc, pericardium; rs; radula sac; rsm, right shell muscle; sn, snout; st, stomach; t, cephalic tentacle." pageId="5" pageNumber="6">3A–C</figureCitation>
|
||||
) typical for genus (
|
||||
<bibRefCitation id="EFACA7EAFFC94729243DF8D0729DF82A" author="Sasaki T & Okutani T & Fujikura K" box="[821,998,1901,1925]" pageId="5" pageNumber="6" pagination="1 - 26" refId="ref11343" refString="Sasaki T, Okutani T, Fujikura K. Anatomy of Shinkailepas myojinensis Sasaki, Okutani & Fujikura, 2003 (Gastropoda: Neritopsina). Malacologia 2006; 48: 1 - 26." type="journal article" year="2006">
|
||||
Sasaki
|
||||
<emphasis id="B9490609FFC947292473F8D372D7F82A" box="[891,940,1901,1925]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
|
||||
2006
|
||||
</bibRefCitation>
|
||||
). Cephalic tentacles simple conical, tapered, elongate. Cephalic lappets small, positioned immediately anterior of cephalic tentacle. Left cephalic lappet develops into conical penis (
|
||||
<figureCitation id="1306C69EFFCA472A2613F8BD7013F8B7" box="[283,360,1792,1816]" captionStart="Figure 2" captionStartId="4.[129,194,1543,1567]" captionTargetBox="[129,1473,144,1515]" captionTargetId="figure-171@4.[129,1473,144,1515]" captionTargetPageId="4" captionText="Figure 2. Shinkailepas tiarasimia, representative specimens. A, holotype (RSIO 38219), female, Wocan field.B, paratype 1 (RSIO 38220), female, Wocan field.C, paratype 2 (RSIO 38221), female, Wocan field.D, paratype 4 (RSIO 38223), male, Wocan field. E, paratype 6 (RSIO 38225), female, Wocan field, partly covered in rusty mineral deposits. F, damaged shell collected from Kairei field (NSMT-Mo 79568), completely covered in rusty mineral deposits. Black arrowhead indicates an egg case (enlarged in Fig. 3D); white arrowhead indicates the penis." pageId="6" pageNumber="7">Fig. 2D</figureCitation>
|
||||
) in males. Triangular neck lobe present posterior of cephalic tentacles. Eyes present, with dark pigments. Oral disc (
|
||||
<figureCitation id="1306C69EFFCA472A27E6F8827021F8F8" box="[238,346,1855,1879]" captionStart="Figure 2" captionStartId="4.[129,194,1543,1567]" captionTargetBox="[129,1473,144,1515]" captionTargetId="figure-171@4.[129,1473,144,1515]" captionTargetPageId="4" captionText="Figure 2. Shinkailepas tiarasimia, representative specimens. A, holotype (RSIO 38219), female, Wocan field.B, paratype 1 (RSIO 38220), female, Wocan field.C, paratype 2 (RSIO 38221), female, Wocan field.D, paratype 4 (RSIO 38223), male, Wocan field. E, paratype 6 (RSIO 38225), female, Wocan field, partly covered in rusty mineral deposits. F, damaged shell collected from Kairei field (NSMT-Mo 79568), completely covered in rusty mineral deposits. Black arrowhead indicates an egg case (enlarged in Fig. 3D); white arrowhead indicates the penis." pageId="6" pageNumber="7">Fig. 2C–D</figureCitation>
|
||||
) circular with strongly muscular outer lip, surrounded by well-developed oral lappet as wide as oral disc to either side. Dorsal surface (
|
||||
<figureCitation id="1306C69EFFCA472A269DF8C070A5F83A" box="[405,478,1917,1941]" captionStart="Figure 3" captionStartId="5.[113,178,1532,1556]" captionTargetBox="[114,1458,144,1505]" captionTargetId="figure-148@5.[114,1458,144,1505]" captionTargetPageId="5" captionText="Figure 3. Shinkailepas tiarasimia, external anatomy. A, dorsal aspect with shell removed, paratype 6 (RSIO 38225), female. B, enlarged dorsal view of the head with mantle edge removed, paratype 3 (RSIO 38222), female. C, operculum shown attached to dorsal aspect of foot with dashed line indicating the approximate outline, paratype 2 (RSIO 38221), female.D, egg capsule found on the shell of paratype 1 (RSIO 38220), female.Abbreviations:ag, albumen gland; cg, capsule gland; cl, cephalic lappet; ct, ctenidium; dg, digestive gland; e, eye; lsm, left shell muscle; me, mantle edge; nl, neck lobe; ol, oral lappet; ov, ovary; pc, pericardium; rs; radula sac; rsm, right shell muscle; sn, snout; st, stomach; t, cephalic tentacle." pageId="6" pageNumber="7">Fig. 3A</figureCitation>
|
||||
) covered by thin mantle with thickened edge lacking major tentacles. Hypobranchial gland lacking. Shell muscle divided into two, left shell muscle reaching slightly anterior of right shell muscle. Pallial edge slightly anterior of left shell muscle edge. Single ctenidium bipectinate, oblique in position with base at posterior left extending to anterior right. Intestine visible on dorsal mantle wall immediately posterior of ctenidium. Pallial gonoduct attached on dorsal mantle ceiling, surrounded anteriorly by capsule gland and posteriorly by albumen gland in females; in males replaced by prostate anteriorly and annex gland posteriorly. Pericardium triangular, on posterior left just posterior or ctenidium. Part of stomach visible on most posterior part of dorsal visceral mass, surrounded by extensive digestive gland dorsal to extensively developed gonad. Foot (
|
||||
<figureCitation id="1306C69EFFCB472B27BBFE907185FEEA" box="[179,254,301,325]" captionStart="Figure 2" captionStartId="4.[129,194,1543,1567]" captionTargetBox="[129,1473,144,1515]" captionTargetId="figure-171@4.[129,1473,144,1515]" captionTargetPageId="4" captionText="Figure 2. Shinkailepas tiarasimia, representative specimens. A, holotype (RSIO 38219), female, Wocan field.B, paratype 1 (RSIO 38220), female, Wocan field.C, paratype 2 (RSIO 38221), female, Wocan field.D, paratype 4 (RSIO 38223), male, Wocan field. E, paratype 6 (RSIO 38225), female, Wocan field, partly covered in rusty mineral deposits. F, damaged shell collected from Kairei field (NSMT-Mo 79568), completely covered in rusty mineral deposits. Black arrowhead indicates an egg case (enlarged in Fig. 3D); white arrowhead indicates the penis." pageId="7" pageNumber="8">Fig. 2C</figureCitation>
|
||||
) with circular, flattened sole, anterior pedal gland present. Epipodial fold carries 12–15 short, triangular tentacles. Operculum (
|
||||
<figureCitation id="1306C69EFFCB472B27F0FED67038FE2C" box="[248,323,363,387]" captionStart="Figure 3" captionStartId="5.[113,178,1532,1556]" captionTargetBox="[114,1458,144,1505]" captionTargetId="figure-148@5.[114,1458,144,1505]" captionTargetPageId="5" captionText="Figure 3. Shinkailepas tiarasimia, external anatomy. A, dorsal aspect with shell removed, paratype 6 (RSIO 38225), female. B, enlarged dorsal view of the head with mantle edge removed, paratype 3 (RSIO 38222), female. C, operculum shown attached to dorsal aspect of foot with dashed line indicating the approximate outline, paratype 2 (RSIO 38221), female.D, egg capsule found on the shell of paratype 1 (RSIO 38220), female.Abbreviations:ag, albumen gland; cg, capsule gland; cl, cephalic lappet; ct, ctenidium; dg, digestive gland; e, eye; lsm, left shell muscle; me, mantle edge; nl, neck lobe; ol, oral lappet; ov, ovary; pc, pericardium; rs; radula sac; rsm, right shell muscle; sn, snout; st, stomach; t, cephalic tentacle." pageId="7" pageNumber="8">Fig. 3C</figureCitation>
|
||||
) present, attached to dorsal surface of foot below visceral mass, thin, translucent, rather symmetrical in outline but with thickened nucleus area at left side, entirely corneous with no evidence of calcification. Diameter of opercular nucleus (see
|
||||
<bibRefCitation id="EFACA7EAFFCB472B27FCFE547086FDAE" author="Fukumori H & Kano Y" box="[244,509,489,513]" pageId="7" pageNumber="8" pagination="213 - 27" refId="ref10235" refString="Fukumori H, Kano Y. Evolutionary ecology of settlement size in planktotrophic neritimorph gastropods. Marine Biology 2014; 161: 213 - 27. https: // doi. org / 10.1007 / s 00227 - 013 - 2330 - 5" type="journal article" year="2014">Fukumori and Kano 2014</bibRefCitation>
|
||||
) 475 μm.
|
||||
</paragraph>
|
||||
<caption id="DF428A93FFCA472A278AF9F97007F91E" pageId="6" pageNumber="7" startId="6.[130,195,1604,1628]" targetBox="[129,1473,144,1577]" targetPageId="6" targetType="figure">
|
||||
<paragraph id="8B82DA1BFFCA472A278AF9F97007F91E" blockId="6.[129,1438,1604,1713]" pageId="6" pageNumber="7">
|
||||
<emphasis id="B9490609FFCA472A278AF9F971A1F9F2" bold="true" box="[130,218,1604,1629]" pageId="6" pageNumber="7">Figure 4.</emphasis>
|
||||
<taxonomicName id="4C3DA198FFCA472A27D5F9F870D4F9F3" authorityName="Gu & Chen & Gao & Zhou & Sun" authorityYear="2025" box="[221,431,1605,1629]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="tiarasimia">
|
||||
<emphasis id="B9490609FFCA472A27D5F9F870D4F9F3" box="[221,431,1605,1629]" italics="true" pageId="6" pageNumber="7">Shinkailepas tiarasimia</emphasis>
|
||||
</taxonomicName>
|
||||
, scanning electron micrographs. A, protoconch of a specimen from Kairei field (NSMT-Mo 79568). B, enlargement of part A showing fine pitting on the protoconch. C, shell microstructure, paratype 5 (RSIO 38224). D, shell surface sculpture, paratype 5 (RSIO 38224). E, overview of the radula, paratype 4 (RSIO 38223). F, close-up of the outer lateral teeth and marginal teeth, paratype 4 (RSIO 38223).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B82DA1BFFCB472B2785FDB57330FB3C" blockId="7.[113,764,144,1327]" pageId="7" pageNumber="8">
|
||||
Radula (
|
||||
<figureCitation id="1306C69EFFCB472B27E6FDB57022FD8F" box="[238,345,520,544]" captionStart="Figure 4" captionStartId="6.[130,195,1604,1628]" captionTargetBox="[129,1473,144,1577]" captionTargetId="figure-150@6.[129,1473,144,1577]" captionTargetPageId="6" captionText="Figure 4.Shinkailepas tiarasimia, scanning electron micrographs. A, protoconch of a specimen from Kairei field (NSMT-Mo 79568). B, enlargement of part A showing fine pitting on the protoconch. C, shell microstructure, paratype 5 (RSIO 38224). D, shell surface sculpture, paratype 5 (RSIO 38224).E, overview of the radula, paratype 4 (RSIO 38223).F, close-up of the outer lateral teeth and marginal teeth, paratype 4 (RSIO 38223)." pageId="7" pageNumber="8">Fig. 4E–F</figureCitation>
|
||||
) rhipidoglossate, formula
|
||||
<emphasis id="B9490609FFCB472B2589FDB473EBFD8F" box="[641,656,521,544]" italics="true" pageId="7" pageNumber="8">c.</emphasis>
|
||||
80–4–1– 4–
|
||||
<emphasis id="B9490609FFCB472B2784FD9571E0FD90" box="[140,155,552,575]" italics="true" pageId="7" pageNumber="8">c.</emphasis>
|
||||
80. Central tooth oblong-quadrate, twice as long as wide, dorsal surface smooth, distally with very weak anterior ridge, overhanging cusp lacking. Innermost first lateral large, film-like, obliquely positioned with outer edge much higher than central tooth. Anterior edge of first lateral rolled to form weakly overhanging, smooth cusp; outermost edge raised to form vertical ridge. Second lateral small with inwardly recurved shaft, positionally occupying anterior half of first lateral, distally equipped with double overhanging cusps. Anterior cusp of second lateral smooth, posterior cusp with five to six denticles. Third lateral with strongly rolled shaft enveloping second lateral, cusp oblique with single major denticle whose inner side carries irregular serration. Outermost fourth lateral large, 1.5 times as long as third lateral, with slightly inwardly recurved shaft that gradually expands distally to form large, overhanging cusp with one inner, major denticle plus three outer, minor denticles. Inner marginals (
|
||||
<figureCitation id="1306C69EFFCB472B27ECFBA07051FB9A" box="[228,298,1053,1077]" captionStart="Figure 4" captionStartId="6.[130,195,1604,1628]" captionTargetBox="[129,1473,144,1577]" captionTargetId="figure-150@6.[129,1473,144,1577]" captionTargetPageId="6" captionText="Figure 4.Shinkailepas tiarasimia, scanning electron micrographs. A, protoconch of a specimen from Kairei field (NSMT-Mo 79568). B, enlargement of part A showing fine pitting on the protoconch. C, shell microstructure, paratype 5 (RSIO 38224). D, shell surface sculpture, paratype 5 (RSIO 38224).E, overview of the radula, paratype 4 (RSIO 38223).F, close-up of the outer lateral teeth and marginal teeth, paratype 4 (RSIO 38223)." pageId="7" pageNumber="8">Fig. 4F</figureCitation>
|
||||
) as long as fourth lateral, distally serrated into rake-like cusp with five to six denticles. Outer marginals much longer than inner marginals with elongate shafts ending in very finely serrated cusps, increasingly so outwards.
|
||||
</paragraph>
|
||||
<paragraph id="8B82DA1BFFCB472B2785FB27737FFA80" blockId="7.[113,764,144,1327]" pageId="7" pageNumber="8">
|
||||
Egg capsule (
|
||||
<figureCitation id="1306C69EFFCB472B2617FB277014FB1D" box="[287,367,1178,1202]" captionStart="Figure 3" captionStartId="5.[113,178,1532,1556]" captionTargetBox="[114,1458,144,1505]" captionTargetId="figure-148@5.[114,1458,144,1505]" captionTargetPageId="5" captionText="Figure 3. Shinkailepas tiarasimia, external anatomy. A, dorsal aspect with shell removed, paratype 6 (RSIO 38225), female. B, enlarged dorsal view of the head with mantle edge removed, paratype 3 (RSIO 38222), female. C, operculum shown attached to dorsal aspect of foot with dashed line indicating the approximate outline, paratype 2 (RSIO 38221), female.D, egg capsule found on the shell of paratype 1 (RSIO 38220), female.Abbreviations:ag, albumen gland; cg, capsule gland; cl, cephalic lappet; ct, ctenidium; dg, digestive gland; e, eye; lsm, left shell muscle; me, mantle edge; nl, neck lobe; ol, oral lappet; ov, ovary; pc, pericardium; rs; radula sac; rsm, right shell muscle; sn, snout; st, stomach; t, cephalic tentacle." pageId="7" pageNumber="8">Fig. 3D</figureCitation>
|
||||
) about
|
||||
<quantity id="4CC577FEFFCB472B26C3FB277362FB1E" box="[459,537,1178,1202]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="8" unit="mm" value="0.6">0.6 mm</quantity>
|
||||
in size, oblong oval, semi-transparent, similar in shape to other
|
||||
<taxonomicName id="4C3DA198FFCB472B252DFB0473DBFB7E" box="[549,672,1209,1233]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B9490609FFCB472B252DFB0473DBFB7E" box="[549,672,1209,1233]" italics="true" pageId="7" pageNumber="8">Shinkailepas</emphasis>
|
||||
</taxonomicName>
|
||||
egg capsules previously illustrated in literature (
|
||||
<bibRefCitation id="EFACA7EAFFCB472B2512FB6473D6FB5E" author="Metaxas A" box="[538,685,1241,1265]" pageId="7" pageNumber="8" pagination="103 - 17" refId="ref10937" refString="Metaxas A. Spatial patterns of larval abundance at hydrothermal vents on seamounts: evidence for recruitment limitation. Marine Ecology Progress Series 2011; 437: 103 - 17. https: // doi. org / 10.3354 / meps 09283" type="journal article" year="2011">Metaxas 2011</bibRefCitation>
|
||||
). Only one found on shell surface (of
|
||||
<typeStatus id="548664B9FFCB472B26ACFB457086FABF" box="[420,509,1272,1296]" pageId="7" pageNumber="8" type="paratype">paratype</typeStatus>
|
||||
1;
|
||||
<figureCitation id="1306C69EFFCB472B2513FB457319FABF" box="[539,610,1272,1296]" captionStart="Figure 2" captionStartId="4.[129,194,1543,1567]" captionTargetBox="[129,1473,144,1515]" captionTargetId="figure-171@4.[129,1473,144,1515]" captionTargetPageId="4" captionText="Figure 2. Shinkailepas tiarasimia, representative specimens. A, holotype (RSIO 38219), female, Wocan field.B, paratype 1 (RSIO 38220), female, Wocan field.C, paratype 2 (RSIO 38221), female, Wocan field.D, paratype 4 (RSIO 38223), male, Wocan field. E, paratype 6 (RSIO 38225), female, Wocan field, partly covered in rusty mineral deposits. F, damaged shell collected from Kairei field (NSMT-Mo 79568), completely covered in rusty mineral deposits. Black arrowhead indicates an egg case (enlarged in Fig. 3D); white arrowhead indicates the penis." pageId="7" pageNumber="8">Fig. 2B</figureCitation>
|
||||
), at least seven embryos can be seen inside the capsule.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3278990FFCB472B2779FAEC7189FA68" pageId="7" pageNumber="8" type="etymology">
|
||||
<paragraph id="8B82DA1BFFCB472B2779FAEC7189FA68" blockId="7.[113,764,1361,1479]" pageId="7" pageNumber="8">
|
||||
<emphasis id="B9490609FFCB472B2779FAEC719AFAC6" box="[113,225,1361,1385]" italics="true" pageId="7" pageNumber="8">Etymology:</emphasis>
|
||||
Greek,
|
||||
<emphasis id="B9490609FFCB472B2668FAEC70F5FAC7" box="[352,398,1361,1384]" italics="true" pageId="7" pageNumber="8">tiára</emphasis>
|
||||
(‘headdress’) +
|
||||
<emphasis id="B9490609FFCB472B255AFAEC73E9FAC7" box="[594,658,1361,1384]" italics="true" pageId="7" pageNumber="8">asimía</emphasis>
|
||||
(‘silvercoloured’), feminine adjective. Refers to the silver sheen of the ostracum and its cap-like shape, reminiscent of oriental silver headdresses.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3278990FFCB472B2779FA5573ECF9B0" pageId="7" pageNumber="8" type="distribution">
|
||||
<paragraph id="8B82DA1BFFCB472B2779FA5573ECF9B0" blockId="7.[113,762,1512,1567]" pageId="7" pageNumber="8">
|
||||
<emphasis id="B9490609FFCB472B2779FA557194F9AF" box="[113,239,1512,1536]" italics="true" pageId="7" pageNumber="8">Distribution:</emphasis>
|
||||
Known with certainty from hydrothermal vents on both the CR (Wocan field) and the CIR (Kairei field).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3278990FFCB472B268AF98275ADFCF6" pageId="7" pageNumber="8" type="discussion">
|
||||
<paragraph id="8B82DA1BFFCB472B268AF9827092F9F6" blockId="7.[113,764,1599,1975]" box="[386,489,1599,1625]" pageId="7" pageNumber="8">
|
||||
<emphasis id="B9490609FFCB472B268AF9827092F9F6" bold="true" box="[386,489,1599,1625]" pageId="7" pageNumber="8">Remarks:</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="8B82DA1BFFCB472B2779F9D875F2FE8A" blockId="7.[113,764,1599,1975]" lastBlockId="7.[809,1460,144,857]" pageId="7" pageNumber="8">
|
||||
Morphologically
|
||||
<taxonomicName id="4C3DA198FFCB472B262AF9DB70E6F9D2" authorityName="Gu & Chen & Gao & Zhou & Sun" authorityYear="2025" box="[290,413,1638,1661]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="tiarasimia">
|
||||
<emphasis id="B9490609FFCB472B262AF9DB70E6F9D2" box="[290,413,1638,1661]" italics="true" pageId="7" pageNumber="8">S. tiarasimia</emphasis>
|
||||
</taxonomicName>
|
||||
most closely resembles
|
||||
<taxonomicName id="4C3DA198FFCB472B259AF9DB7380F9D2" box="[658,763,1638,1661]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="conspira">
|
||||
<emphasis id="B9490609FFCB472B259AF9DB7380F9D2" box="[658,763,1638,1661]" italics="true" pageId="7" pageNumber="8">S. conspira</emphasis>
|
||||
</taxonomicName>
|
||||
from the Lau Basin in the southwestern Pacific (
|
||||
<bibRefCitation id="EFACA7EAFFCB472B2576F93871BFF913" author="Poitrimol C & Thiebaut E & Daguin-Thiebaut C" pageId="7" pageNumber="8" refId="ref11104" refString="Poitrimol C, Thiebaut E, Daguin-Thiebaut C et al. Contrasted phylogeographic patterns of hydrothermal vent gastropods along South West Pacific: Woodlark Basin, a possible contact zone and / or stepping-stone. PLoS One 2022; 17: e 0275638. https: // doi. org / 10.1371 / journal. pone. 0275638" type="journal volume" year="2022">
|
||||
Poitrimol
|
||||
<emphasis id="B9490609FFCB472B25E1F93871F1F913" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
|
||||
2022
|
||||
</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EFACA7EAFFCB472B27D9F91970B3F913" author="Chen C & Sigwart JD" box="[209,456,1700,1724]" pageId="7" pageNumber="8" pagination="401 - 36" refId="ref9891" refString="Chen C, Sigwart JD. The lost vent gastropod species of Lothar A. Beck. Zootaxa 2023; 5270: 401 - 36. https: // doi. org / 10.11646 / zootaxa. 5270.3.2" type="journal article" year="2023">Chen and Sigwart 2023</bibRefCitation>
|
||||
), which exhibits similar shell sculpture with concentric ribs stronger than radial ones equal in strength, a similar apex position near the shell posterior, as well as a similar radula morphology. These two species, however, can be clearly differentiated in that the epipodial lobe of
|
||||
<taxonomicName id="4C3DA198FFCB472B259AF89F7387F896" box="[658,764,1826,1849]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="conspira">
|
||||
<emphasis id="B9490609FFCB472B259AF89F7387F896" box="[658,764,1826,1849]" italics="true" pageId="7" pageNumber="8">S. conspira</emphasis>
|
||||
</taxonomicName>
|
||||
is well developed and bears up to 60 paddle-like flaps (
|
||||
<bibRefCitation id="EFACA7EAFFCB472B259EF8FC718CF8D7" author="Chen C & Sigwart JD" pageId="7" pageNumber="8" pagination="401 - 36" refId="ref9891" refString="Chen C, Sigwart JD. The lost vent gastropod species of Lothar A. Beck. Zootaxa 2023; 5270: 401 - 36. https: // doi. org / 10.11646 / zootaxa. 5270.3.2" type="journal article" year="2023">Chen and Sigwart 2023</bibRefCitation>
|
||||
) compared to
|
||||
<taxonomicName id="4C3DA198FFCB472B2683F8DD737FF8D7" authorityName="Gu & Chen & Gao & Zhou & Sun" authorityYear="2025" box="[395,516,1888,1912]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="tiarasimia">
|
||||
<emphasis id="B9490609FFCB472B2683F8DD737FF8D7" box="[395,516,1888,1912]" italics="true" pageId="7" pageNumber="8">S. tiarasimia</emphasis>
|
||||
</taxonomicName>
|
||||
where only 12–15 small triangular tentacles are present.
|
||||
<taxonomicName id="4C3DA198FFCB472B26BDF8C273FCF838" box="[437,647,1919,1943]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="conspira">
|
||||
<emphasis id="B9490609FFCB472B26BDF8C273FCF838" box="[437,647,1919,1943]" italics="true" pageId="7" pageNumber="8">Shinkailepas conspira</emphasis>
|
||||
</taxonomicName>
|
||||
is also notably a much larger species reaching up to
|
||||
<quantity id="4CC577FEFFCB472B2512F822730EF819" box="[538,629,1951,1975]" metricMagnitude="-2" metricUnit="m" metricValue="2.13" pageId="7" pageNumber="8" unit="mm" value="21.3">21.3 mm</quantity>
|
||||
in SL (
|
||||
<bibRefCitation id="EFACA7EAFFCB472B25C9F82272A6FF07" author="Chen C & Sigwart JD" box="[705,989,144,1975]" pageId="7" pageNumber="8" pagination="401 - 36" refId="ref9891" refString="Chen C, Sigwart JD. The lost vent gastropod species of Lothar A. Beck. Zootaxa 2023; 5270: 401 - 36. https: // doi. org / 10.11646 / zootaxa. 5270.3.2" type="journal article" year="2023">Chen and Sigwart 2023</bibRefCitation>
|
||||
). Another species sharing a similar adult shell sculpture is
|
||||
<taxonomicName id="4C3DA198FFCB472B24A5FF0D7284FF68" authorityName="Beck" authorityYear="1992" box="[941,1023,175,199]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="tufari">
|
||||
<emphasis id="B9490609FFCB472B24A5FF0D7284FF68" box="[941,1023,175,199]" italics="true" pageId="7" pageNumber="8">S. tufari</emphasis>
|
||||
</taxonomicName>
|
||||
from the Manus Basin vents also in the southwestern Pacific (
|
||||
<bibRefCitation id="EFACA7EAFFCB472B2318FF72759EFF48" author="Poitrimol C & Thiebaut E & Daguin-Thiebaut C" box="[1040,1253,207,231]" pageId="7" pageNumber="8" refId="ref11104" refString="Poitrimol C, Thiebaut E, Daguin-Thiebaut C et al. Contrasted phylogeographic patterns of hydrothermal vent gastropods along South West Pacific: Woodlark Basin, a possible contact zone and / or stepping-stone. PLoS One 2022; 17: e 0275638. https: // doi. org / 10.1371 / journal. pone. 0275638" type="journal volume" year="2022">
|
||||
Poitrimol
|
||||
<emphasis id="B9490609FFCB472B2372FF7275D0FF48" box="[1146,1195,207,231]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
|
||||
2022
|
||||
</bibRefCitation>
|
||||
); however,
|
||||
<taxonomicName id="4C3DA198FFCB472B226BFF7274C9FF48" authorityName="Beck" authorityYear="1992" box="[1379,1458,207,231]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="tufari">
|
||||
<emphasis id="B9490609FFCB472B226BFF7274C9FF48" box="[1379,1458,207,231]" italics="true" pageId="7" pageNumber="8">S. tufari</emphasis>
|
||||
</taxonomicName>
|
||||
has a much larger septum positioned at approximately the midpoint of the aperture (
|
||||
<bibRefCitation id="EFACA7EAFFCB472B2305FEB07503FE8A" author="Beck LA" box="[1037,1144,269,293]" pageId="7" pageNumber="8" pagination="259 - 75" refId="ref9697" refString="Beck LA. Two new neritacean limpets (Gastropoda: Prosobranchia: Neritacea: Phenacolepadidae) from active hydrothermal vents at Hydrothermal Field 1 ' Wienerwald' in the Manus Back-Arc Basin (Bismarck Sea, Papua-New Guinea). Annalen des Naturhistorischen Museums in Wien. Serie B fur Botanik und Zoologie 1992; 93: 259 - 75." type="journal article" year="1992">Beck 1992</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8B82DA1BFFCB472B244DFE9075ADFCF6" blockId="7.[809,1460,144,857]" pageId="7" pageNumber="8">
|
||||
While
|
||||
<taxonomicName id="4C3DA198FFCB472B2487FE907562FEEB" authorityName="Sasaki, Okutani & Fujikura" authorityYear="2003" box="[911,1049,301,324]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="myojinensis">
|
||||
<emphasis id="B9490609FFCB472B2487FE907562FEEB" box="[911,1049,301,324]" italics="true" pageId="7" pageNumber="8">S. myojinensis</emphasis>
|
||||
</taxonomicName>
|
||||
typically exhibits stronger shell sculpture than
|
||||
<taxonomicName id="4C3DA198FFCB472B249BFEF1759EFECB" authority="(Sasaki et al. 2003)" baseAuthorityName="Sasaki" baseAuthorityYear="2003" box="[915,1253,332,356]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="tiarasimia">
|
||||
<emphasis id="B9490609FFCB472B249BFEF1756AFECB" box="[915,1041,332,356]" italics="true" pageId="7" pageNumber="8">S. tiarasimia</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFACA7EAFFCB472B232CFEF175A1FECB" author="Sasaki T & Okutani T & Fujikura K" box="[1060,1242,332,356]" pageId="7" pageNumber="8" pagination="189 - 210" refId="ref11309" refString="Sasaki T, Okutani T, Fujikura K. New taxa and new records of patelliform gastropods associated with hemoautosynthesis-based communities in Japanese waters. Veliger 2003; 46: 189 - 210." type="journal article" year="2003">
|
||||
Sasaki
|
||||
<emphasis id="B9490609FFCB472B2364FEF075E5FECB" box="[1132,1182,332,356]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
|
||||
2003
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
this species can be morphologically very variable and some specimens may resemble
|
||||
<taxonomicName id="4C3DA198FFCB472B2475FE3675A0FE0C" authority="(Yahagi et al. 2017)" baseAuthorityName="Yahagi" baseAuthorityYear="2017" box="[893,1243,395,419]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="tiarasimia">
|
||||
<emphasis id="B9490609FFCB472B2475FE367287FE0D" box="[893,1020,395,418]" italics="true" pageId="7" pageNumber="8">S. tiarasimia</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFACA7EAFFCB472B2319FE3675ABFE0C" author="Yahagi T & Kayama Watanabe H & Kojima S" box="[1041,1232,395,419]" pageId="7" pageNumber="8" pagination="1524 - 34" refId="ref12036" refString="Yahagi T, Kayama Watanabe H, Kojima S et al. Do larvae from deep-sea hydrothermal vents disperse in surface waters? Ecology 2017; 98: 1524 - 34. https: // doi. org / 10.1002 / ecy. 1800" type="journal article" year="2017">
|
||||
Yahagi
|
||||
<emphasis id="B9490609FFCB472B2357FE3675E8FE0C" box="[1119,1171,395,419]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
|
||||
2017
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
, but the operculum of
|
||||
<taxonomicName id="4C3DA198FFCB472B2440FE1772AAFE6D" authorityName="Sasaki, Okutani & Fujikura" authorityYear="2003" box="[840,977,426,450]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="myojinensis">
|
||||
<emphasis id="B9490609FFCB472B2440FE1772AAFE6D" box="[840,977,426,450]" italics="true" pageId="7" pageNumber="8">S. myojinensis</emphasis>
|
||||
</taxonomicName>
|
||||
is partly calcified (likewise in
|
||||
<taxonomicName id="4C3DA198FFCB472B2210FE1774FFFE6D" box="[1304,1412,426,450]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="conspira">
|
||||
<emphasis id="B9490609FFCB472B2210FE1774FFFE6D" box="[1304,1412,426,450]" italics="true" pageId="7" pageNumber="8">S. conspira</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C3DA198FFCB472B2422FE777200FE4E" authorityName="Beck" authorityYear="1992" box="[810,891,457,481]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="tufari">
|
||||
<emphasis id="B9490609FFCB472B2422FE777200FE4E" box="[810,891,457,481]" italics="true" pageId="7" pageNumber="8">S. tufari</emphasis>
|
||||
</taxonomicName>
|
||||
), whereas in
|
||||
<taxonomicName id="4C3DA198FFCB472B2307FE7775F5FE4E" box="[1039,1166,458,481]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="tiarasimia">
|
||||
<emphasis id="B9490609FFCB472B2307FE7775F5FE4E" box="[1039,1166,458,481]" italics="true" pageId="7" pageNumber="8">S. tiarasimia</emphasis>
|
||||
</taxonomicName>
|
||||
it is entirely corneous. The protoconch of both
|
||||
<taxonomicName id="4C3DA198FFCB472B24F4FE54751DFDAF" box="[1020,1126,489,512]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="conspira">
|
||||
<emphasis id="B9490609FFCB472B24F4FE54751DFDAF" box="[1020,1126,489,512]" italics="true" pageId="7" pageNumber="8">S. conspira</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C3DA198FFCB472B2391FE547593FDAE" authorityName="Beck" authorityYear="1992" box="[1177,1256,489,513]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="tufari">
|
||||
<emphasis id="B9490609FFCB472B2391FE547593FDAE" box="[1177,1256,489,513]" italics="true" pageId="7" pageNumber="8">S. tufari</emphasis>
|
||||
</taxonomicName>
|
||||
are both about 800 μm in size (
|
||||
<bibRefCitation id="EFACA7EAFFCB472B24A0FDB5756CFD8F" author="Beck LA" box="[936,1047,520,544]" pageId="7" pageNumber="8" pagination="259 - 75" refId="ref9697" refString="Beck LA. Two new neritacean limpets (Gastropoda: Prosobranchia: Neritacea: Phenacolepadidae) from active hydrothermal vents at Hydrothermal Field 1 ' Wienerwald' in the Manus Back-Arc Basin (Bismarck Sea, Papua-New Guinea). Annalen des Naturhistorischen Museums in Wien. Serie B fur Botanik und Zoologie 1992; 93: 259 - 75." type="journal article" year="1992">Beck 1992</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EFACA7EAFFCB472B232DFDB57465FD8F" author="Chen C & Sigwart JD" box="[1061,1310,520,544]" pageId="7" pageNumber="8" pagination="401 - 36" refId="ref9891" refString="Chen C, Sigwart JD. The lost vent gastropod species of Lothar A. Beck. Zootaxa 2023; 5270: 401 - 36. https: // doi. org / 10.11646 / zootaxa. 5270.3.2" type="journal article" year="2023">Chen and Sigwart 2023</bibRefCitation>
|
||||
), much larger than the 660 μm in
|
||||
<taxonomicName id="4C3DA198FFCB472B24F3FD957503FD90" box="[1019,1144,552,575]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="tiarasimia">
|
||||
<emphasis id="B9490609FFCB472B24F3FD957503FD90" box="[1019,1144,552,575]" italics="true" pageId="7" pageNumber="8">S. tiarasimia</emphasis>
|
||||
</taxonomicName>
|
||||
; while that of
|
||||
<taxonomicName id="4C3DA198FFCB472B221AFD9574E0FD90" authorityName="Sasaki, Okutani & Fujikura" authorityYear="2003" box="[1298,1435,552,575]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="myojinensis">
|
||||
<emphasis id="B9490609FFCB472B221AFD9574E0FD90" box="[1298,1435,552,575]" italics="true" pageId="7" pageNumber="8">S. myojinensis</emphasis>
|
||||
</taxonomicName>
|
||||
is similar and also 660 μm (
|
||||
<bibRefCitation id="EFACA7EAFFCB472B233AFDFA7599FDF0" author="Sasaki T & Okutani T & Fujikura K" box="[1074,1250,583,607]" pageId="7" pageNumber="8" pagination="189 - 210" refId="ref11309" refString="Sasaki T, Okutani T, Fujikura K. New taxa and new records of patelliform gastropods associated with hemoautosynthesis-based communities in Japanese waters. Veliger 2003; 46: 189 - 210." type="journal article" year="2003">
|
||||
Sasaki
|
||||
<emphasis id="B9490609FFCB472B2370FDFA75D3FDF0" box="[1144,1192,583,607]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
|
||||
2003
|
||||
</bibRefCitation>
|
||||
). However, we note that our protoconch size is measured from the exposed part in adult shells unlike those of
|
||||
<taxonomicName id="4C3DA198FFCB472B234AFD3B75B2FD32" authorityName="Sasaki, Okutani & Fujikura" authorityYear="2003" box="[1090,1225,646,669]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="myojinensis">
|
||||
<emphasis id="B9490609FFCB472B234AFD3B75B2FD32" box="[1090,1225,646,669]" italics="true" pageId="7" pageNumber="8">S. myojinensis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C3DA198FFCB472B23F4FD3B7431FD32" authorityName="Beck" authorityYear="1992" box="[1276,1354,645,669]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="tufari">
|
||||
<emphasis id="B9490609FFCB472B23F4FD3B7431FD32" box="[1276,1354,645,669]" italics="true" pageId="7" pageNumber="8">S. tufari</emphasis>
|
||||
</taxonomicName>
|
||||
which are measured using juveniles (
|
||||
<bibRefCitation id="EFACA7EAFFCB472B2331FD1875DFFD12" author="Beck LA" box="[1081,1188,677,701]" pageId="7" pageNumber="8" pagination="259 - 75" refId="ref9697" refString="Beck LA. Two new neritacean limpets (Gastropoda: Prosobranchia: Neritacea: Phenacolepadidae) from active hydrothermal vents at Hydrothermal Field 1 ' Wienerwald' in the Manus Back-Arc Basin (Bismarck Sea, Papua-New Guinea). Annalen des Naturhistorischen Museums in Wien. Serie B fur Botanik und Zoologie 1992; 93: 259 - 75." type="journal article" year="1992">Beck 1992</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EFACA7EAFFCB472B23A7FD187427FD12" author="Sasaki T & Okutani T & Fujikura K" box="[1199,1372,677,701]" pageId="7" pageNumber="8" pagination="189 - 210" refId="ref11309" refString="Sasaki T, Okutani T, Fujikura K. New taxa and new records of patelliform gastropods associated with hemoautosynthesis-based communities in Japanese waters. Veliger 2003; 46: 189 - 210." type="journal article" year="2003">
|
||||
Sasaki
|
||||
<emphasis id="B9490609FFCB472B23FCFD187458FD12" box="[1268,1315,677,701]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
|
||||
2003
|
||||
</bibRefCitation>
|
||||
). As the teleoconch of
|
||||
<taxonomicName id="4C3DA198FFCB472B24BFFD797549FD73" box="[951,1074,708,732]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B9490609FFCB472B24BFFD797549FD73" box="[951,1074,708,732]" italics="true" pageId="7" pageNumber="8">Shinkailepas</emphasis>
|
||||
</taxonomicName>
|
||||
often envelops the protoconch during growth, our measurement of the protoconch size is likely less accurate than those measured from juveniles. The central teeth of
|
||||
<taxonomicName id="4C3DA198FFCB472B2423FC9F72DCFC95" box="[811,935,802,826]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="tiarasimia">
|
||||
<emphasis id="B9490609FFCB472B2423FC9F72DCFC95" box="[811,935,802,826]" italics="true" pageId="7" pageNumber="8">S. tiarasimia</emphasis>
|
||||
</taxonomicName>
|
||||
also appear to lack a horizontal ridge that is prominent in
|
||||
<taxonomicName id="4C3DA198FFCB472B2476FCFF75A9FCF6" authority="(Sasaki et al., 2003)" baseAuthorityName="Sasaki" baseAuthorityYear="2003" box="[894,1234,833,857]" class="Gastropoda" family="Phenacolepadidae" genus="Shinkailepas" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="myojinensis">
|
||||
<emphasis id="B9490609FFCB472B2476FCFF757FFCF6" box="[894,1028,834,857]" italics="true" pageId="7" pageNumber="8">S. myojinensis</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFACA7EAFFCB472B231CFCFC75BDFCF6" author="Sasaki T & Okutani T & Fujikura K" box="[1044,1222,833,857]" pageId="7" pageNumber="8" pagination="189 - 210" refId="ref11309" refString="Sasaki T, Okutani T, Fujikura K. New taxa and new records of patelliform gastropods associated with hemoautosynthesis-based communities in Japanese waters. Veliger 2003; 46: 189 - 210." type="journal article" year="2003">
|
||||
Sasaki
|
||||
<emphasis id="B9490609FFCB472B2351FCFF75FAFCF6" box="[1113,1153,833,857]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
|
||||
., 2003
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
Loading…
Reference in a new issue