diff --git a/data/A2/2B/48/A22B481BFFDEFF9AE4E62F47FE72FDAC.xml b/data/A2/2B/48/A22B481BFFDEFF9AE4E62F47FE72FDAC.xml new file mode 100644 index 00000000000..aae4b75256d --- /dev/null +++ b/data/A2/2B/48/A22B481BFFDEFF9AE4E62F47FE72FDAC.xml @@ -0,0 +1,1342 @@ +<document id="9364B4D3F9365A1DAEDF9DFCA2E22F5E" ID-DOI="10.1649/0010-065X-76.1.85" ID-ISSN="1938-4394" ID-Zenodo-Dep="13252164" ID-ZooBank="urn:lsid:zoobank.org:pub:15321DC6-D04B-4D26-9751-B0D3E9C2BCC7" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1723002119180" checkinUser="felipe" docAuthor="Suzumura, Alyssa Lee, Henegouwen, Arno van Berge & Budi, Anang Setyo" docDate="2022" docId="A22B481BFFDEFF9AE4E62F47FE72FDAC" docLanguage="en" docName="TheColeopBull.76.1.85-94.pdf" docOrigin="The 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E) Male protarsus, F) Mesoventral process, ventral.Abbreviations: bs = basal segment of tegmen; cor = corona; ls = lateral struts of tergite IX; mkp = mesoventral protrusion; ml = median lobe; mr = median ridge of ventrite IX; oc = outer claw; par = paramera; pml = posterior mesoventral lip; S5 = tarsomere 5; T10 = tergite X; V9 = sternite IX." figureDoi-0="http://doi.org/10.5281/zenodo.13252166" figureDoi-1="http://doi.org/10.5281/zenodo.13252170" figureDoi-2="http://doi.org/10.5281/zenodo.13252172" httpUri-0="https://zenodo.org/record/13252166/files/figure.png" httpUri-1="https://zenodo.org/record/13252170/files/figure.png" httpUri-2="https://zenodo.org/record/13252172/files/figure.png" pageId="3" pageNumber="87">Figs. 1–3</figureCitation> +, +<figureCitation id="B2B9E588FFDEFF93E59D2FDCFC35FA3E" box="[937,969,1481,1502]" captionStart="Fig" captionStartId="7.[197,228,990,1009]" captionTargetBox="[177,655,196,960]" captionTargetId="figure-561@7.[173,659,193,963]" captionTargetPageId="7" captionText="Fig. 4. Pronotal coloration, left lateral view. A) Sphaeridium sundense, new species, B) S. quinquemaculatum." figureDoi="http://doi.org/10.5281/zenodo.13252174" httpUri="https://zenodo.org/record/13252174/files/figure.png" pageId="3" pageNumber="87">4A</figureCitation> +, +<figureCitation id="B2B9E588FFDEFF93E5E72FDCFBD5FA3F" box="[979,1065,1481,1503]" captionStart="Fig" captionStartId="8.[176,207,1360,1379]" captionTargetBox="[152,1172,193,1333]" captionTargetId="figure-172@8.[152,1172,193,1333]" captionTargetPageId="8" captionText="Fig. 5. Morphology of Sphaeridium species.A, C, E) Sphaeridium sundense, new species, B, D, F) S. quinquemaculatum. A–B) Procoxae, C–D) Mesoventral keel, ventral view, E–F) Mesoventral keel, oblique view." figureDoi="http://doi.org/10.5281/zenodo.13252176" httpUri="https://zenodo.org/record/13252176/files/figure.png" pageId="3" pageNumber="87">5A, C, E</figureCitation> +, +<figureCitation id="B2B9E588FFDEFF93E2072FDCFBBCFA3F" box="[1075,1088,1481,1503]" captionStart="Fig" captionStartId="9.[197,228,642,661]" captionTargetBox="[173,1193,193,615]" captionTargetId="figure-592@9.[173,1193,193,615]" captionTargetPageId="9" captionText="Fig. 6. 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( +<specimenCount id="3C843284FFDAFF97E56F2B69FC85FE71" box="[859,889,380,401]" count="1" pageId="7" pageNumber="91" type="male">1♂</specimenCount> +, +<collectionCode id="4C9361C8FFDAFF97E5BE2B69FC29FE71" box="[906,981,380,401]" collectionName="Netherlands, Amsterdam, Universiteit van Amsterdam, Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="7" pageNumber="91">ZMAN</collectionCode> +); +</materialsCitation> +<materialsCitation id="9AEAF350FFDAFF97E5DF2B69FBEBFE4C" pageId="7" pageNumber="91"> +<collectorName id="87779CDBFFDAFF97E5DF2B69FB5CFE71" box="[1003,1184,380,402]" pageId="7" pageNumber="91">Tandjong Morawa</collectorName> +/ +<collectorName id="87779CDBFFDAFF97E4F72B82FCE8FE4D" box="[707,788,407,429]" pageId="7" pageNumber="91">Serdang</collectorName> +, coll. +<collectorName id="87779CDBFFDAFF97E5792B82FC70FE4C" box="[845,908,407,428]" pageId="7" pageNumber="91">Hagen</collectorName> +( +<specimenCount id="3C843284FFDAFF97E5A82B82FC44FE4C" box="[924,952,407,428]" count="1" pageId="7" pageNumber="91" type="female">1♀</specimenCount> +, +<collectionCode id="4C9361C8FFDAFF97E5F02B82FBEDFE4C" box="[964,1041,407,428]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]" pageId="7" pageNumber="91">RMNH</collectionCode> +) +</materialsCitation> +. +</paragraph> +</materialsCitation> +</paragraph> +</subSubSection> +<caption id="7EFDA985FFD8FF95E6F12F7CFD47FA24" ID-DOI="http://doi.org/10.5281/zenodo.13252170" ID-Zenodo-Dep="13252170" httpUri="https://zenodo.org/record/13252170/files/figure.png" pageId="5" pageNumber="89" startId="5.[197,228,1385,1404]" targetBox="[173,1193,193,1358]" targetPageId="5" targetType="figure"> +<paragraph id="2A3DF90DFFD8FF95E6F12F7CFD47FA24" blockId="5.[173,1193,1385,1476]" pageId="5" pageNumber="89"> +<emphasis id="18F6251FFFD8FF95E6F12F7CFF02FA9C" bold="true" box="[197,254,1385,1404]" pageId="5" pageNumber="89">Fig. 2.</emphasis> +SEM of +<taxonomicName id="ED82828EFFD8FF95E7682F7CFDE1FA9C" authorityName="Suzumura & Henegouwen & Budi" authorityYear="2022" box="[348,541,1385,1404]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="89" phylum="Arthropoda" rank="species" species="sundense" status="sp. nov."> +<emphasis id="18F6251FFFD8FF95E7682F7CFDE1FA9C" box="[348,541,1385,1404]" italics="true" pageId="5" pageNumber="89">Sphaeridium sundense</emphasis> +</taxonomicName> +, +<emphasis id="18F6251FFFD8FF95E41D2F7CFD6EFA9C" bold="true" box="[553,658,1385,1404]" pageId="5" pageNumber="89"> +<taxonomicNameLabel id="03C59864FFD8FF95E41D2F7CFD6EFA9C" box="[553,658,1385,1404]" pageId="5" pageNumber="89" rank="species">new species</taxonomicNameLabel> +</emphasis> +. A) Head, ventral, inset shows microrugae on surface of mentum, B) Pronotum, dorso-lateral, inset shows microsculpture on surface, C) Proventrite, D) Mesoventral keel, ventral oblique, E) Metaventral field. F–G) Detail of types of microsculpture: F) Granulate microsculpture on pronotum, G) Rugose microsculpture (microrugae) on metaventral field. +</paragraph> +</caption> +<caption id="7EFDA985FFDBFF96E69B2E10FC91FB99" ID-DOI="http://doi.org/10.5281/zenodo.13252172" ID-Zenodo-Dep="13252172" httpUri="https://zenodo.org/record/13252172/files/figure.png" pageId="6" pageNumber="90" startId="6.[175,206,1029,1048]" targetBox="[156,1168,197,999]" targetPageId="6" targetType="figure"> +<paragraph id="2A3DF90DFFDBFF96E69B2E10FC91FB99" blockId="6.[152,1172,1029,1145]" pageId="6" pageNumber="90"> +<emphasis id="18F6251FFFDBFF96E69B2E10FF1BFBF8" bold="true" box="[175,231,1029,1048]" pageId="6" pageNumber="90">Fig. 3.</emphasis> +<taxonomicName id="ED82828EFFDBFF96E6C92E13FE41FBF9" authorityName="Suzumura & Henegouwen & Budi" authorityYear="2022" box="[253,445,1030,1049]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="90" phylum="Arthropoda" rank="species" species="sundense" status="sp. nov."> +<emphasis id="18F6251FFFDBFF96E6C92E13FE41FBF9" box="[253,445,1030,1049]" italics="true" pageId="6" pageNumber="90">Sphaeridium sundense</emphasis> +</taxonomicName> +, +<emphasis id="18F6251FFFDBFF96E7FC2E13FDCDFBF8" bold="true" box="[456,561,1029,1049]" pageId="6" pageNumber="90"> +<taxonomicNameLabel id="03C59864FFDBFF96E7FC2E13FDCDFBF8" box="[456,561,1029,1049]" pageId="6" pageNumber="90" rank="species">new species</taxonomicNameLabel> +</emphasis> +. A–D) Male genitalia:A) Abdominal tergite IX, dorsal, B) Median lobe and tegmen, C) Ventral detail of apex of median lobe, D) Lateral profile of median lobe and tegmen. E) Male protarsus, F) Mesoventral process, ventral.Abbreviations: bs = basal segment of tegmen; cor = corona; ls = lateral struts of tergite IX; mkp = mesoventral protrusion; ml = median lobe; mr = median ridge of ventrite IX; oc = outer claw; par = paramera; pml = posterior mesoventral lip; S5 = tarsomere 5; T10 = tergite X; V9 = sternite IX. +</paragraph> +</caption> +<subSubSection id="6298AA86FFDAFF99E4EF2BA4FC82FCAF" lastPageId="9" lastPageNumber="93" pageId="7" pageNumber="91" type="description"> +<paragraph id="2A3DF90DFFDAFF97E4EF2BA4FB79FE1D" blockId="7.[706,1193,194,1736]" pageId="7" pageNumber="91"> +<emphasis id="18F6251FFFDAFF97E4EF2BA4FCADFE27" bold="true" box="[731,849,433,455]" pageId="7" pageNumber="91">Description.</emphasis> +Small, body length 3.0–5.0 mm, body width 2.0–3.0 mm. Body shape oval to slightly sub- quadrate, moderately compressed dorsoventrally. +</paragraph> +<paragraph id="2A3DF90DFFDAFF97E4EF2817FCCAFBCD" blockId="7.[706,1193,194,1736]" pageId="7" pageNumber="91"> +Coloration not widely variable, usually as in +<figureCitation id="B2B9E588FFDAFF97E4F72809FD05FDD1" box="[707,761,540,562]" captionStart="Fig" captionStartId="4.[176,207,1304,1323]" captionTargetBox="[152,1172,193,1277]" captionTargetId="figure-157@4.[152,1172,193,1277]" captionTargetPageId="4" captionText="Fig. 1. Sphaeridium sundense, new species, habitus. A) Dorsal, B) Ventral, C) Lateral, D) Posterior." figureDoi="http://doi.org/10.5281/zenodo.13252166" httpUri="https://zenodo.org/record/13252166/files/figure.png" pageId="7" pageNumber="91">Fig. 1</figureCitation> +, head black, sometimes with a small indistinct reddish spot at middle behind level of eyes, labrum reddish, pronotum black with wide yellow stripe along lateral margins ( +<figureCitation id="B2B9E588FFDAFF97E5AC2879FBF4FD61" box="[920,1032,620,642]" captionStart="Fig" captionStartId="4.[176,207,1304,1323]" captionTargetBox="[152,1172,193,1277]" captionTargetId="figure-157@4.[152,1172,193,1277]" captionTargetPageId="4" captionText="Fig. 1. Sphaeridium sundense, new species, habitus. A) Dorsal, B) Ventral, C) Lateral, D) Posterior." figureDoi="http://doi.org/10.5281/zenodo.13252166" httpUri="https://zenodo.org/record/13252166/files/figure.png" pageId="7" pageNumber="91">Figs. 1A, C</figureCitation> +, +<figureCitation id="B2B9E588FFDAFF97E2252879FBCDFD61" box="[1041,1073,620,641]" captionStart="Fig" captionStartId="7.[197,228,990,1009]" captionTargetBox="[177,655,196,960]" captionTargetId="figure-561@7.[173,659,193,963]" captionTargetPageId="7" captionText="Fig. 4. Pronotal coloration, left lateral view. A) Sphaeridium sundense, new species, B) S. quinquemaculatum." figureDoi="http://doi.org/10.5281/zenodo.13252174" httpUri="https://zenodo.org/record/13252174/files/figure.png" pageId="7" pageNumber="91">4A</figureCitation> +), continuing around anterior and posterior angles but not con- tacting posterior margin; elytra black to reddish in more freshly emerged adults, lateral margins with yellow border beginning slightly posterior to base, widening before midlength, continuing around posterior tip and ending abruptly at about 1/4 to 1/3 the length of elytral suture, border of this stripe indistinct posteriorly; lacking reddish humeral spots; epipleuron fulvous. Pygidium dark brown on anterior 1/2–1/4, yellow posteriorly. Ventral base color dark brown except narrow yellow stripe on gula, lateral portions of pronotum, and posterior margins of ventrites ( +<figureCitation id="B2B9E588FFDAFF97E50E29D2FC7FFC3C" box="[826,899,967,989]" captionStart="Fig" captionStartId="4.[176,207,1304,1323]" captionTargetBox="[152,1172,193,1277]" captionTargetId="figure-157@4.[152,1172,193,1277]" captionTargetPageId="4" captionText="Fig. 1. Sphaeridium sundense, new species, habitus. A) Dorsal, B) Ventral, C) Lateral, D) Posterior." figureDoi="http://doi.org/10.5281/zenodo.13252166" httpUri="https://zenodo.org/record/13252166/files/figure.png" pageId="7" pageNumber="91">Fig. 1B</figureCitation> +); legs yellow with brown spots at centers of femora. Mentum piceous-testaceous, maxillary palps yellow, antennae yellow with dark brown club. +</paragraph> +<caption id="7EFDA985FFDAFF97E6F129CBFEEDFBC1" ID-DOI="http://doi.org/10.5281/zenodo.13252174" ID-Zenodo-Dep="13252174" httpUri="https://zenodo.org/record/13252174/files/figure.png" pageId="7" pageNumber="91" startId="7.[197,228,990,1009]" targetBox="[177,655,196,960]" targetPageId="7" targetType="figure"> +<paragraph id="2A3DF90DFFDAFF97E6F129CBFEEDFBC1" blockId="7.[173,660,990,1057]" pageId="7" pageNumber="91"> +<emphasis id="18F6251FFFDAFF97E6F129CBFEF0FC11" bold="true" box="[197,268,990,1009]" pageId="7" pageNumber="91">Fig. 4.</emphasis> +Pronotal coloration, left lateral view. A) +<taxonomicName id="ED82828EFFDAFF97E6FF29E3FE72FBE9" authorityName="Suzumura & Henegouwen & Budi" authorityYear="2022" box="[203,398,1014,1033]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="91" phylum="Arthropoda" rank="species" species="sundense" status="sp. nov."> +<emphasis id="18F6251FFFDAFF97E6FF29E3FE72FBE9" box="[203,398,1014,1033]" italics="true" pageId="7" pageNumber="91">Sphaeridium sundense</emphasis> +</taxonomicName> +, +<emphasis id="18F6251FFFDAFF97E7AF29E3FDFAFBE9" bold="true" box="[411,518,1014,1033]" pageId="7" pageNumber="91"> +<taxonomicNameLabel id="03C59864FFDAFF97E7AF29E3FDFAFBE9" box="[411,518,1014,1033]" pageId="7" pageNumber="91" rank="species">new species</taxonomicNameLabel> +</emphasis> +, B) +<taxonomicName id="ED82828EFFDAFF97E41B29E3FEF0FBC1" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="91" phylum="Arthropoda" rank="species" species="quinquemaculatum"> +<emphasis id="18F6251FFFDAFF97E41B29E3FEF0FBC1" italics="true" pageId="7" pageNumber="91">S. quinquemaculatum</emphasis> +</taxonomicName> +. +</paragraph> +</caption> +<paragraph id="2A3DF90DFFDAFF97E4EF2E27FB55FA42" blockId="7.[706,1193,194,1736]" pageId="7" pageNumber="91"> +Dorsal surface of head with dense, uniform punctures separated by less than their diameter except around smooth epicranial sutures, interstices with shallow microsculpture. Eyes of normal size, excised on anterior margin. Labrum exposed, anterior margin straight medially, with dense, flat, golden pubesence anteriorly, surface covered in dense transverse microrugae. Mentum +<emphasis id="18F6251FFFDAFF97E54A2EF8FC67FAE2" box="[894,923,1261,1282]" italics="true" pageId="7" pageNumber="91">ca.</emphasis> +0.6 times as long as wide, slightly bisinuate on anterior margin, lateral margins gently curved, surface with deep, coarse setiferous punctures, interspaces with dense, conspicuous microrugae ( +<figureCitation id="B2B9E588FFDAFF97E5112F42FC89FA8C" box="[805,885,1367,1389]" captionStart="Fig" captionStartId="5.[197,228,1385,1404]" captionTargetBox="[173,1193,193,1358]" captionTargetId="figure-110@5.[173,1193,193,1358]" captionTargetPageId="5" captionText="Fig. 2. SEM of Sphaeridium sundense, new species. A) Head, ventral, inset shows microrugae on surface of mentum, B) Pronotum, dorso-lateral, inset shows microsculpture on surface, C) Proventrite, D) Mesoventral keel, ventral oblique, E) Metaventral field. F–G) Detail of types of microsculpture: F) Granulate microsculpture on pronotum, G) Rugose microsculpture (microrugae) on metaventral field." figureDoi="http://doi.org/10.5281/zenodo.13252170" httpUri="https://zenodo.org/record/13252170/files/figure.png" pageId="7" pageNumber="91">Fig. 2A</figureCitation> +); gula parallel-sided or slightly wider posteriorly, of normal width, not distinctly wide or narrow.Antennal club symmetrical, compact. +</paragraph> +<paragraph id="2A3DF90DFFDAFF98E4EF2FB2FE5FF9E7" blockId="7.[706,1193,194,1736]" lastBlockId="8.[152,638,1442,1757]" lastPageId="8" lastPageNumber="92" pageId="7" pageNumber="91"> +Pronotum with bisinuate posterior margin; lateral margins evenly curved, with irregular row of small blunt spines along posterior 2/3 ( +<figureCitation id="B2B9E588FFDAFF97E5CC2FC9FBBEFA11" box="[1016,1090,1500,1522]" captionStart="Fig" captionStartId="5.[197,228,1385,1404]" captionTargetBox="[173,1193,193,1358]" captionTargetId="figure-110@5.[173,1193,193,1358]" captionTargetPageId="5" captionText="Fig. 2. SEM of Sphaeridium sundense, new species. A) Head, ventral, inset shows microrugae on surface of mentum, B) Pronotum, dorso-lateral, inset shows microsculpture on surface, C) Proventrite, D) Mesoventral keel, ventral oblique, E) Metaventral field. F–G) Detail of types of microsculpture: F) Granulate microsculpture on pronotum, G) Rugose microsculpture (microrugae) on metaventral field." figureDoi="http://doi.org/10.5281/zenodo.13252170" httpUri="https://zenodo.org/record/13252170/files/figure.png" pageId="7" pageNumber="91">Fig. 2B</figureCitation> +); marginal beading wide, flat, continued along anterior margin, ending at posterior margin; posterolateral corners forming an oblique angle. Surface covered in dense punctures separated by less than their diameter, slightly larger than those on head, interspaces with dense, granulate microsculpture ( +<figureCitation id="B2B9E588FFDAFF97E2282C69FB61F971" box="[1052,1181,1660,1682]" captionStart="Fig" captionStartId="5.[197,228,1385,1404]" captionTargetBox="[173,1193,193,1358]" captionTargetId="figure-110@5.[173,1193,193,1358]" captionTargetPageId="5" captionText="Fig. 2. SEM of Sphaeridium sundense, new species. A) Head, ventral, inset shows microrugae on surface of mentum, B) Pronotum, dorso-lateral, inset shows microsculpture on surface, C) Proventrite, D) Mesoventral keel, ventral oblique, E) Metaventral field. F–G) Detail of types of microsculpture: F) Granulate microsculpture on pronotum, G) Rugose microsculpture (microrugae) on metaventral field." figureDoi="http://doi.org/10.5281/zenodo.13252170" httpUri="https://zenodo.org/record/13252170/files/figure.png" pageId="7" pageNumber="91">Figs. 2B, F</figureCitation> +). Proventral process convex, weakly tectiform posteriorly between coxae, forming a large spine at posterior apex, and bearing 2−4 strong spines along longitudinal median line, the anterior of which is flanked by several weaker spines, golden setae, and sparse pubesence ( +<figureCitation id="B2B9E588FFD5FF98E77F2FE7FE6BF9E7" box="[331,407,1522,1544]" captionStart="Fig" captionStartId="5.[197,228,1385,1404]" captionTargetBox="[173,1193,193,1358]" captionTargetId="figure-110@5.[173,1193,193,1358]" captionTargetPageId="5" captionText="Fig. 2. SEM of Sphaeridium sundense, new species. A) Head, ventral, inset shows microrugae on surface of mentum, B) Pronotum, dorso-lateral, inset shows microsculpture on surface, C) Proventrite, D) Mesoventral keel, ventral oblique, E) Metaventral field. F–G) Detail of types of microsculpture: F) Granulate microsculpture on pronotum, G) Rugose microsculpture (microrugae) on metaventral field." figureDoi="http://doi.org/10.5281/zenodo.13252170" httpUri="https://zenodo.org/record/13252170/files/figure.png" pageId="8" pageNumber="92">Fig. 2C</figureCitation> +). +</paragraph> +<caption id="7EFDA985FFD5FF98E6842F45FC09FA9B" ID-DOI="http://doi.org/10.5281/zenodo.13252176" ID-Zenodo-Dep="13252176" httpUri="https://zenodo.org/record/13252176/files/figure.png" pageId="8" pageNumber="92" startId="8.[176,207,1360,1379]" targetBox="[152,1172,193,1333]" targetPageId="8" targetType="figure"> +<paragraph id="2A3DF90DFFD5FF98E6842F45FC09FA9B" blockId="8.[152,1170,1360,1404]" pageId="8" pageNumber="92"> +<emphasis id="18F6251FFFD5FF98E6842F45FF1BFA83" bold="true" box="[176,231,1360,1379]" pageId="8" pageNumber="92">Fig. 5.</emphasis> +Morphology of +<taxonomicName id="ED82828EFFD5FF98E7B72F45FDB1FA83" box="[387,589,1360,1379]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="92" phylum="Arthropoda" rank="species" species="undefined-A"> +<emphasis id="18F6251FFFD5FF98E7B72F45FE0CFA83" box="[387,496,1360,1379]" italics="true" pageId="8" pageNumber="92">Sphaeridium</emphasis> +species.A +</taxonomicName> +, C, E) +<taxonomicName id="ED82828EFFD5FF98E4B32F45FCBBFA83" authorityName="Suzumura & Henegouwen & Budi" authorityYear="2022" box="[647,839,1360,1379]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="92" phylum="Arthropoda" rank="species" species="sundense" status="sp. nov."> +<emphasis id="18F6251FFFD5FF98E4B32F45FCBBFA83" box="[647,839,1360,1379]" italics="true" pageId="8" pageNumber="92">Sphaeridium sundense</emphasis> +</taxonomicName> +, +<emphasis id="18F6251FFFD5FF98E5662F45FC46FA83" bold="true" box="[850,954,1360,1379]" pageId="8" pageNumber="92"> +<taxonomicNameLabel id="03C59864FFD5FF98E5662F45FC46FA83" box="[850,954,1360,1379]" pageId="8" pageNumber="92" rank="species">new species</taxonomicNameLabel> +</emphasis> +, B, D, F) +<taxonomicName id="ED82828EFFD5FF98E23A2F44FF2FFA9B" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="92" phylum="Arthropoda" rank="species" species="quinquemaculatum"> +<emphasis id="18F6251FFFD5FF98E23A2F44FF2FFA9B" italics="true" pageId="8" pageNumber="92">S. quinquemaculatum</emphasis> +</taxonomicName> +. A–B) Procoxae, C–D) Mesoventral keel, ventral view, E–F) Mesoventral keel, oblique view. +</paragraph> +</caption> +<paragraph id="2A3DF90DFFD5FF98E6842C19FD9EF96D" blockId="8.[152,638,1442,1757]" pageId="8" pageNumber="92"> +Scutellar shield about twice as long as wide, sides almost parallel-sided anteriorly, gently curved posteriorly ( +<figureCitation id="B2B9E588FFD5FF98E6D82C57FEC4F9B7" box="[236,312,1602,1624]" captionStart="Fig" captionStartId="4.[176,207,1304,1323]" captionTargetBox="[152,1172,193,1277]" captionTargetId="figure-157@4.[152,1172,193,1277]" captionTargetPageId="4" captionText="Fig. 1. Sphaeridium sundense, new species, habitus. A) Dorsal, B) Ventral, C) Lateral, D) Posterior." figureDoi="http://doi.org/10.5281/zenodo.13252166" httpUri="https://zenodo.org/record/13252166/files/figure.png" pageId="8" pageNumber="92">Fig. 1A</figureCitation> +); surface densely punctured, con- sistent with surrounding punctation on elytra, interspaces without discernable microsculpture. +</paragraph> +<paragraph id="2A3DF90DFFD5FF99E6842C87FD72FCAD" blockId="8.[152,638,1442,1757]" lastBlockId="9.[173,660,716,1751]" lastPageId="9" lastPageNumber="93" pageId="8" pageNumber="92"> +Elytra ovoid, sides strongly rounded, apices rounded separately; sutural interval slightly raised, surface flat, continuous with marginal beading which is distinct along lateral margins and around posterior apices ( +<figureCitation id="B2B9E588FFD5FF98E5602FA9FC35FA32" box="[852,969,1468,1490]" captionStart="Fig" captionStartId="4.[176,207,1304,1323]" captionTargetBox="[152,1172,193,1277]" captionTargetId="figure-157@4.[152,1172,193,1277]" captionTargetPageId="4" captionText="Fig. 1. Sphaeridium sundense, new species, habitus. A) Dorsal, B) Ventral, C) Lateral, D) Posterior." figureDoi="http://doi.org/10.5281/zenodo.13252166" httpUri="https://zenodo.org/record/13252166/files/figure.png" pageId="8" pageNumber="92">Figs. 1C, D</figureCitation> +). Surface covered in dense, uniform punctures separated by less than their diameter, interspaces with distinct granulate microsculpture and sparse microrugae throughout; striae distinct, though larger strial punctures not evenly aligned, without grooves, interspaces flat ( +<figureCitation id="B2B9E588FFD5FF98E4802C49FCFDF991" box="[692,769,1628,1650]" captionStart="Fig" captionStartId="4.[176,207,1304,1323]" captionTargetBox="[152,1172,193,1277]" captionTargetId="figure-157@4.[152,1172,193,1277]" captionTargetPageId="4" captionText="Fig. 1. Sphaeridium sundense, new species, habitus. A) Dorsal, B) Ventral, C) Lateral, D) Posterior." figureDoi="http://doi.org/10.5281/zenodo.13252166" httpUri="https://zenodo.org/record/13252166/files/figure.png" pageId="8" pageNumber="92">Fig. 1A</figureCitation> +). Mesoventral process as in +<figureCitation id="B2B9E588FFD5FF98E2222C49FB96F991" box="[1046,1130,1628,1650]" captionStart="Fig" captionStartId="5.[197,228,1385,1404]" captionTargetBox="[173,1193,193,1358]" captionTargetId="figure-110@5.[173,1193,193,1358]" captionTargetPageId="5" captionText="Fig. 2. SEM of Sphaeridium sundense, new species. A) Head, ventral, inset shows microrugae on surface of mentum, B) Pronotum, dorso-lateral, inset shows microsculpture on surface, C) Proventrite, D) Mesoventral keel, ventral oblique, E) Metaventral field. F–G) Detail of types of microsculpture: F) Granulate microsculpture on pronotum, G) Rugose microsculpture (microrugae) on metaventral field." figureDoi="http://doi.org/10.5281/zenodo.13252170" httpUri="https://zenodo.org/record/13252170/files/figure.png" pageId="8" pageNumber="92">Figs. 2D</figureCitation> +and +<figureCitation id="B2B9E588FFD5FF98E4992C62FD3BF96C" box="[685,711,1655,1676]" captionStart="Fig" captionStartId="6.[175,206,1029,1048]" captionTargetBox="[156,1168,197,999]" captionTargetId="figure-241@6.[152,1172,193,1002]" captionTargetPageId="6" captionText="Fig. 3. Sphaeridium sundense, new species. A–D) Male genitalia:A) Abdominal tergite IX, dorsal, B) Median lobe and tegmen, C) Ventral detail of apex of median lobe, D) Lateral profile of median lobe and tegmen. E) Male protarsus, F) Mesoventral process, ventral.Abbreviations: bs = basal segment of tegmen; cor = corona; ls = lateral struts of tergite IX; mkp = mesoventral protrusion; ml = median lobe; mr = median ridge of ventrite IX; oc = outer claw; par = paramera; pml = posterior mesoventral lip; S5 = tarsomere 5; T10 = tergite X; V9 = sternite IX." figureDoi="http://doi.org/10.5281/zenodo.13252172" httpUri="https://zenodo.org/record/13252172/files/figure.png" pageId="8" pageNumber="92">3F</figureCitation> +, anterior protrusion triangular, distinctly longer than wide, with a distinctive symmetrical pattern of long, strong spines: four rows of strong spines of increasing size numbering 3-6-4-2 from anterior to posterior. Posterior lip slightly convex, slightly shorter than protrusion, separated from lateral areas of mesoventrite by posterolateral angles of protrusion, distinctly raised above level of metaventrite, lacking pubesence on surface ( +<figureCitation id="B2B9E588FFD4FF99E7E02922FDD6FCAC" box="[468,554,823,845]" captionStart="Fig" captionStartId="5.[197,228,1385,1404]" captionTargetBox="[173,1193,193,1358]" captionTargetId="figure-110@5.[173,1193,193,1358]" captionTargetPageId="5" captionText="Fig. 2. SEM of Sphaeridium sundense, new species. A) Head, ventral, inset shows microrugae on surface of mentum, B) Pronotum, dorso-lateral, inset shows microsculpture on surface, C) Proventrite, D) Mesoventral keel, ventral oblique, E) Metaventral field. F–G) Detail of types of microsculpture: F) Granulate microsculpture on pronotum, G) Rugose microsculpture (microrugae) on metaventral field." figureDoi="http://doi.org/10.5281/zenodo.13252170" httpUri="https://zenodo.org/record/13252170/files/figure.png" pageId="9" pageNumber="93">Figs. 2D</figureCitation> +, +<figureCitation id="B2B9E588FFD4FF99E4002922FDADFCAC" box="[564,593,823,844]" captionStart="Fig" captionStartId="6.[175,206,1029,1048]" captionTargetBox="[156,1168,197,999]" captionTargetId="figure-241@6.[152,1172,193,1002]" captionTargetPageId="6" captionText="Fig. 3. Sphaeridium sundense, new species. A–D) Male genitalia:A) Abdominal tergite IX, dorsal, B) Median lobe and tegmen, C) Ventral detail of apex of median lobe, D) Lateral profile of median lobe and tegmen. E) Male protarsus, F) Mesoventral process, ventral.Abbreviations: bs = basal segment of tegmen; cor = corona; ls = lateral struts of tergite IX; mkp = mesoventral protrusion; ml = median lobe; mr = median ridge of ventrite IX; oc = outer claw; par = paramera; pml = posterior mesoventral lip; S5 = tarsomere 5; T10 = tergite X; V9 = sternite IX." figureDoi="http://doi.org/10.5281/zenodo.13252172" httpUri="https://zenodo.org/record/13252172/files/figure.png" pageId="9" pageNumber="93">3F</figureCitation> +: pml). +</paragraph> +<caption id="7EFDA985FFD4FF99E6F12897FBCFFD75" ID-DOI="http://doi.org/10.5281/zenodo.13252180" ID-Zenodo-Dep="13252180" box="[197,1075,642,662]" httpUri="https://zenodo.org/record/13252180/files/figure.png" pageId="9" pageNumber="93" startId="9.[197,228,642,661]" targetBox="[173,1193,193,615]" targetPageId="9" targetType="figure"> +<paragraph id="2A3DF90DFFD4FF99E6F12897FBCFFD75" blockId="9.[197,1075,642,662]" box="[197,1075,642,662]" pageId="9" pageNumber="93"> +<emphasis id="18F6251FFFD4FF99E6F12897FF01FD75" bold="true" box="[197,253,642,661]" pageId="9" pageNumber="93">Fig. 6.</emphasis> +Map of locations from which +<taxonomicName id="ED82828EFFD4FF99E4252897FD2DFD75" authorityName="Suzumura & Henegouwen & Budi" authorityYear="2022" box="[529,721,642,661]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="93" phylum="Arthropoda" rank="species" species="sundense" status="sp. nov."> +<emphasis id="18F6251FFFD4FF99E4252897FD2DFD75" box="[529,721,642,661]" italics="true" pageId="9" pageNumber="93">Sphaeridium sundense</emphasis> +</taxonomicName> +, +<emphasis id="18F6251FFFD4FF99E4E82896FCB9FD75" bold="true" box="[732,837,642,662]" pageId="9" pageNumber="93"> +<taxonomicNameLabel id="03C59864FFD4FF99E4E82896FCB9FD75" box="[732,837,642,662]" pageId="9" pageNumber="93" rank="species">new species</taxonomicNameLabel> +</emphasis> +was collected in Indonesia. +</paragraph> +</caption> +<paragraph id="2A3DF90DFFD4FF99E6F12944FDDFFB6D" blockId="9.[173,660,716,1751]" pageId="9" pageNumber="93"> +Metaventrite with metaventral field slightly raised as in +<figureCitation id="B2B9E588FFD4FF99E6EA2979FED9FC61" box="[222,293,876,898]" captionStart="Fig" captionStartId="5.[197,228,1385,1404]" captionTargetBox="[173,1193,193,1358]" captionTargetId="figure-110@5.[173,1193,193,1358]" captionTargetPageId="5" captionText="Fig. 2. SEM of Sphaeridium sundense, new species. A) Head, ventral, inset shows microrugae on surface of mentum, B) Pronotum, dorso-lateral, inset shows microsculpture on surface, C) Proventrite, D) Mesoventral keel, ventral oblique, E) Metaventral field. F–G) Detail of types of microsculpture: F) Granulate microsculpture on pronotum, G) Rugose microsculpture (microrugae) on metaventral field." figureDoi="http://doi.org/10.5281/zenodo.13252170" httpUri="https://zenodo.org/record/13252170/files/figure.png" pageId="9" pageNumber="93">Fig. 2E</figureCitation> +, convex, longer than wide, lateral margins not distinctly separated from surrounding lateral areas, median furrow weakly continuing posterior to transverse furrows which are only distinct centrally, curved laterally, meeting at an oblique angle; posterior margin between metacoxae gently curved, emar- ginate medially; surface covered in deep uniform punctures, separated by about 2× their diameter, interspaces with dense microrugae ( +<figureCitation id="B2B9E588FFD4FF99E7C72E54FDBCFBB6" box="[499,576,1089,1111]" captionStart="Fig" captionStartId="5.[197,228,1385,1404]" captionTargetBox="[173,1193,193,1358]" captionTargetId="figure-110@5.[173,1193,193,1358]" captionTargetPageId="5" captionText="Fig. 2. SEM of Sphaeridium sundense, new species. A) Head, ventral, inset shows microrugae on surface of mentum, B) Pronotum, dorso-lateral, inset shows microsculpture on surface, C) Proventrite, D) Mesoventral keel, ventral oblique, E) Metaventral field. F–G) Detail of types of microsculpture: F) Granulate microsculpture on pronotum, G) Rugose microsculpture (microrugae) on metaventral field." figureDoi="http://doi.org/10.5281/zenodo.13252170" httpUri="https://zenodo.org/record/13252170/files/figure.png" pageId="9" pageNumber="93">Fig. 2G</figureCitation> +), central area posterior to transverse furrow with fewer punctures but microrugae strong throughout. +</paragraph> +<paragraph id="2A3DF90DFFD4FF99E6F12E87FD6FFA06" blockId="9.[173,660,716,1751]" pageId="9" pageNumber="93"> +Procoxae with strong spines and sparse short indistinct setae, lacking long golden pubesence interspersed among the spines, with dentiform process at internal posterior apex. Posterior margin of trochan- ters with a row of spines continuing along posterior margin of femora. Male protarsus modified, with tarsomeres 1–4 shortened and slightly widened and tarsomere 5 almost as long as segments 1–4 com- bined, tarsomere 5 a little wider than tarsomere 4, with apical outer margin straight to slightly curved, slightly longer along ventral margin, without pointed angular process ( +<figureCitation id="B2B9E588FFD4FF99E77E2FA2FE6EFA2C" box="[330,402,1463,1485]" captionStart="Fig" captionStartId="6.[175,206,1029,1048]" captionTargetBox="[156,1168,197,999]" captionTargetId="figure-241@6.[152,1172,193,1002]" captionTargetPageId="6" captionText="Fig. 3. Sphaeridium sundense, new species. A–D) Male genitalia:A) Abdominal tergite IX, dorsal, B) Median lobe and tegmen, C) Ventral detail of apex of median lobe, D) Lateral profile of median lobe and tegmen. E) Male protarsus, F) Mesoventral process, ventral.Abbreviations: bs = basal segment of tegmen; cor = corona; ls = lateral struts of tergite IX; mkp = mesoventral protrusion; ml = median lobe; mr = median ridge of ventrite IX; oc = outer claw; par = paramera; pml = posterior mesoventral lip; S5 = tarsomere 5; T10 = tergite X; V9 = sternite IX." figureDoi="http://doi.org/10.5281/zenodo.13252172" httpUri="https://zenodo.org/record/13252172/files/figure.png" pageId="9" pageNumber="93">Fig. 3E</figureCitation> +). Hind tibia with one small, moderately strong spine on ventral surface ( +<figureCitation id="B2B9E588FFD4FF99E40B2FC4FD7AFA06" box="[575,646,1489,1511]" captionStart="Fig" captionStartId="4.[176,207,1304,1323]" captionTargetBox="[152,1172,193,1277]" captionTargetId="figure-157@4.[152,1172,193,1277]" captionTargetPageId="4" captionText="Fig. 1. Sphaeridium sundense, new species, habitus. A) Dorsal, B) Ventral, C) Lateral, D) Posterior." figureDoi="http://doi.org/10.5281/zenodo.13252166" httpUri="https://zenodo.org/record/13252166/files/figure.png" pageId="9" pageNumber="93">Fig. 1B</figureCitation> +). +</paragraph> +<paragraph id="2A3DF90DFFD4FF99E6F12FF9FDB8F9D7" blockId="9.[173,660,716,1751]" pageId="9" pageNumber="93">Pygidium weakly tectiform, apex obtusely point- ed; abdominal ventrite 5 with apex evenly rounded, surface bare, without fringe of short setae.</paragraph> +<paragraph id="2A3DF90DFFD4FF99E6F12C29FECFF98D" blockId="9.[173,660,716,1751]" pageId="9" pageNumber="93">Abdominal apex simply rounded, without emargination.</paragraph> +<paragraph id="2A3DF90DFFD4FF99E6F12C64FC82FCAF" blockId="9.[173,660,716,1751]" lastBlockId="9.[706,1193,716,1751]" pageId="9" pageNumber="93"> +Male genitalia as in +<figureCitation id="B2B9E588FFD4FF99E7B62C64FE0FF966" box="[386,499,1649,1671]" captionStart="Fig" captionStartId="6.[175,206,1029,1048]" captionTargetBox="[156,1168,197,999]" captionTargetId="figure-241@6.[152,1172,193,1002]" captionTargetPageId="6" captionText="Fig. 3. Sphaeridium sundense, new species. A–D) Male genitalia:A) Abdominal tergite IX, dorsal, B) Median lobe and tegmen, C) Ventral detail of apex of median lobe, D) Lateral profile of median lobe and tegmen. E) Male protarsus, F) Mesoventral process, ventral.Abbreviations: bs = basal segment of tegmen; cor = corona; ls = lateral struts of tergite IX; mkp = mesoventral protrusion; ml = median lobe; mr = median ridge of ventrite IX; oc = outer claw; par = paramera; pml = posterior mesoventral lip; S5 = tarsomere 5; T10 = tergite X; V9 = sternite IX." figureDoi="http://doi.org/10.5281/zenodo.13252172" httpUri="https://zenodo.org/record/13252172/files/figure.png" pageId="9" pageNumber="93">Figs. 3A–D</figureCitation> +. Ratio of median lobe:tegmen = 1.73. Median lobe with sides parallel on basal half, slightly constricted at about apical 1/3–1/4, slightly widened before tip, with truncate apical process; parameres about half the length of median lobe, rather wide on ventral side, with rounded tips.Abdominal segment IX about half the length of the lateral struts, apex roundly pointed, median ridge short. +</paragraph> +</subSubSection> +<subSubSection id="6298AA86FFD4FF99E4EF2942FCA8FC39" pageId="9" pageNumber="93" type="etymology"> +<paragraph id="2A3DF90DFFD4FF99E4EF2942FCA8FC39" blockId="9.[706,1193,716,1751]" pageId="9" pageNumber="93"> +<emphasis id="18F6251FFFD4FF99E4EF2942FCB2FC8C" bold="true" box="[731,846,855,876]" pageId="9" pageNumber="93">Etymology.</emphasis> +The specific epithet +<taxonomicName id="ED82828EFFD4FF99E2162942FB79FC8C" authorityName="Suzumura & Henegouwen & Budi" authorityYear="2022" box="[1058,1157,855,876]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="93" phylum="Arthropoda" rank="species" species="sundense"> +“ +<emphasis id="18F6251FFFD4FF99E2192942FB79FC8C" box="[1069,1157,855,876]" italics="true" pageId="9" pageNumber="93">sundense</emphasis> +</taxonomicName> +” is an adjective taken from the geographical name Sundaland, the former landmass that fell apart into the many islands including what is now called the Lesser Sundas. +</paragraph> +</subSubSection> +<subSubSection id="6298AA86FFD4FF99E4EF29F4FC7DFB7A" pageId="9" pageNumber="93" type="distribution"> +<paragraph id="2A3DF90DFFD4FF99E4EF29F4FC7DFB7A" blockId="9.[706,1193,716,1751]" pageId="9" pageNumber="93"> +<emphasis id="18F6251FFFD4FF99E4EF29F4FCA2FC17" bold="true" box="[731,862,993,1015]" pageId="9" pageNumber="93">Distribution.</emphasis> +<collectingCountry id="5295B99DFFD4FF99E55E29F4FC37FC17" box="[874,971,993,1015]" name="Indonesia" pageId="9" pageNumber="93">Indonesia</collectingCountry> +: +<collectingRegion id="E84637EFFFD4FF99E5EF29F4FBD1FC16" box="[987,1069,993,1014]" country="Indonesia" name="Sumatera Utara" pageId="9" pageNumber="93">Sumatra</collectingRegion> +, +<collectingRegion id="E84637EFFFD4FF99E20929F7FB96FC17" box="[1085,1130,994,1015]" country="Indonesia" name="Jawa Timur" pageId="9" pageNumber="93">Java</collectingRegion> +, +<collectingRegion id="E84637EFFFD4FF99E24E29F4FB58FC17" box="[1146,1188,993,1015]" country="Indonesia" name="Bali" pageId="9" pageNumber="93">Bali</collectingRegion> +, Lombok, Sumbawa, Sumba, +<collectingCountry id="5295B99DFFD4FF99E5EF29E9FBE5FBF2" box="[987,1049,1020,1042]" name="Indonesia" pageId="9" pageNumber="93">Flores</collectingCountry> +, Timor, Wetar, Kei ( +<figureCitation id="B2B9E588FFD4FF99E4C52E02FCD7FBCD" box="[753,811,1047,1069]" captionStart="Fig" captionStartId="9.[197,228,642,661]" captionTargetBox="[173,1193,193,615]" captionTargetId="figure-592@9.[173,1193,193,615]" captionTargetPageId="9" captionText="Fig. 6. Map of locations from which Sphaeridium sundense, new species was collected in Indonesia." figureDoi="http://doi.org/10.5281/zenodo.13252180" httpUri="https://zenodo.org/record/13252180/files/figure.png" pageId="9" pageNumber="93">Fig. 6</figureCitation> +). AVBH recorded it from +<collectingRegion id="E84637EFFFD4FF99E22B2E0DFB8DFBCD" box="[1055,1137,1048,1069]" country="Indonesia" name="Sumatera Utara" pageId="9" pageNumber="93">Sumatra</collectingRegion> +, +<collectingRegion id="E84637EFFFD4FF99E24E2E02FB58FBCD" box="[1146,1188,1047,1069]" country="Indonesia" name="Bali" pageId="9" pageNumber="93">Bali</collectingRegion> +, Wetar, and Pulau Kei based on material in museum collections. This indicates that as far as we know, the new species reaches eastwards to what is known as Lydekker’s Line. +</paragraph> +</subSubSection> +<subSubSection id="6298AA86FFD4FF9AE4EF2EB7FE72FDAC" lastPageId="10" lastPageNumber="94" pageId="9" pageNumber="93" type="discussion"> +<paragraph id="2A3DF90DFFD4FF99E4EF2EB7FC84FA27" blockId="9.[706,1193,716,1751]" pageId="9" pageNumber="93"> +<emphasis id="18F6251FFFD4FF99E4EF2EB7FCC1FB57" bold="true" box="[731,829,1186,1207]" pageId="9" pageNumber="93">Remarks.</emphasis> +While there are several small (> +<quantity id="ED7A54E8FFD4FF99E2462EB7FB54FB57" box="[1138,1192,1186,1207]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="93" unit="mm" value="8.0">8 mm</quantity> +in length) species of +<taxonomicName id="ED82828EFFD4FF99E5B72EA8FC00FB32" authorityName="FABRICIUS" authorityYear="1775" box="[899,1020,1213,1234]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="93" phylum="Arthropoda" rank="genus"> +<emphasis id="18F6251FFFD4FF99E5B72EA8FC00FB32" box="[899,1020,1213,1234]" italics="true" pageId="9" pageNumber="93">Sphaeridium</emphasis> +</taxonomicName> +recorded from the Indonesian archipelago, +<taxonomicName id="ED82828EFFD4FF99E5852ECCFBDEFB0D" authorityName="Suzumura & Henegouwen & Budi" authorityYear="2022" box="[945,1058,1240,1262]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="93" phylum="Arthropoda" rank="species" species="sundense"> +<emphasis id="18F6251FFFD4FF99E5852ECCFBDEFB0D" box="[945,1058,1240,1262]" italics="true" pageId="9" pageNumber="93">S. sundense</emphasis> +</taxonomicName> +can be easily differentiated from most of the others by the follow- ing characters: ventral coloration dark, base color black to dark brown, never yellow (as in +<taxonomicName id="ED82828EFFD4FF99E25F2F3FFCB8FABA" authorityName="OrchYmont" authorityYear="1924" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="93" phylum="Arthropoda" rank="species" species="flavomaculatum"> +<emphasis id="18F6251FFFD4FF99E25F2F3FFCB8FABA" italics="true" pageId="9" pageNumber="93">S. flavomaculatum</emphasis> +</taxonomicName> +and +<taxonomicName id="ED82828EFFD4FF99E5422F50FC0BFABA" baseAuthorityName="van Berge Henegouwen" baseAuthorityYear="1986" box="[886,1015,1349,1370]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="93" phylum="Arthropoda" rank="species" species="huijbregtsi"> +<emphasis id="18F6251FFFD4FF99E5422F50FC0BFABA" box="[886,1015,1349,1370]" italics="true" pageId="9" pageNumber="93">S. huijbregtsi</emphasis> +</taxonomicName> +); posterior angles of pronotum without emargination (as in +<taxonomicName id="ED82828EFFD4FF99E2652F74FD06FA71" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="93" phylum="Arthropoda" rank="species" species="dimidiatum"> +<emphasis id="18F6251FFFD4FF99E2652F74FD06FA71" italics="true" pageId="9" pageNumber="93">S. dimidiatum</emphasis> +</taxonomicName> +and the larger species +<taxonomicName id="ED82828EFFD4FF99E5E32F69FBC3FA70" authorityName="OrchYmont" authorityYear="1933" box="[983,1087,1403,1425]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="93" phylum="Arthropoda" rank="species" species="discolor"> +<emphasis id="18F6251FFFD4FF99E5E32F69FBC3FA70" box="[983,1087,1403,1425]" italics="true" pageId="9" pageNumber="93">S. discolor</emphasis> +</taxonomicName> +); and with distinct elytral striae (lacking in +<taxonomicName id="ED82828EFFD4FF99E5C22F82FB54FA4B" authorityName="OrchYmont" authorityYear="1924" box="[1014,1192,1430,1452]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="93" phylum="Arthropoda" rank="species" species="flavomaculatum"> +<emphasis id="18F6251FFFD4FF99E5C22F82FB54FA4B" box="[1014,1192,1430,1452]" italics="true" pageId="9" pageNumber="93">S. flavomaculatum</emphasis> +</taxonomicName> +and +<taxonomicName id="ED82828EFFD4FF99E4DF2FA7FC96FA27" baseAuthorityName="van Berge Henegouwen" baseAuthorityYear="1986" box="[747,874,1458,1479]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="93" phylum="Arthropoda" rank="species" species="huijbregtsi"> +<emphasis id="18F6251FFFD4FF99E4DF2FA7FC96FA27" box="[747,874,1458,1479]" italics="true" pageId="9" pageNumber="93">S. huijbregtsi</emphasis> +</taxonomicName> +). +</paragraph> +<paragraph id="2A3DF90DFFD4FF9AE4EF2FD8FE72FDAC" blockId="9.[706,1193,716,1751]" lastBlockId="10.[152,638,193,589]" lastPageId="10" lastPageNumber="94" pageId="9" pageNumber="93"> +Perhaps the most morphologically similar species to +<taxonomicName id="ED82828EFFD4FF99E4EF2FFDFCB5FA1D" authorityName="Suzumura & Henegouwen & Budi" authorityYear="2022" box="[731,841,1512,1533]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="93" phylum="Arthropoda" rank="species" species="sundense"> +<emphasis id="18F6251FFFD4FF99E4EF2FFDFCB5FA1D" box="[731,841,1512,1533]" italics="true" pageId="9" pageNumber="93">S. sundense</emphasis> +</taxonomicName> +is +<taxonomicName id="ED82828EFFD4FF99E5502FFDFBCCFA1D" box="[868,1072,1512,1533]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="93" phylum="Arthropoda" rank="species" species="quinquemaculatum"> +<emphasis id="18F6251FFFD4FF99E5502FFDFBCCFA1D" box="[868,1072,1512,1533]" italics="true" pageId="9" pageNumber="93">S. quinquemaculatum</emphasis> +</taxonomicName> +, particularly in size and often coloration. +<taxonomicName id="ED82828EFFD4FF99E5E72C16FB54F9F8" authorityName="Suzumura & Henegouwen & Budi" authorityYear="2022" box="[979,1192,1539,1560]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="93" phylum="Arthropoda" rank="species" species="sundense"> +<emphasis id="18F6251FFFD4FF99E5E72C16FB54F9F8" box="[979,1192,1539,1560]" italics="true" pageId="9" pageNumber="93">Sphaeridium sundense</emphasis> +</taxonomicName> +can, however, be easily separated from the former by the structure of the mesoventral process, of which the anterior protrusion is distinctly longer than wide, bears a unique pattern of strong spines on the ventral surface, and the posterior lip lacks pubescence ( +<figureCitation id="B2B9E588FFD4FF99E24D2C9EFD1EF95B" captionStart="Fig" captionStartId="5.[197,228,1385,1404]" captionTargetBox="[173,1193,193,1358]" captionTargetId="figure-110@5.[173,1193,193,1358]" captionTargetPageId="5" captionText="Fig. 2. SEM of Sphaeridium sundense, new species. A) Head, ventral, inset shows microrugae on surface of mentum, B) Pronotum, dorso-lateral, inset shows microsculpture on surface, C) Proventrite, D) Mesoventral keel, ventral oblique, E) Metaventral field. F–G) Detail of types of microsculpture: F) Granulate microsculpture on pronotum, G) Rugose microsculpture (microrugae) on metaventral field." figureDoi="http://doi.org/10.5281/zenodo.13252170" httpUri="https://zenodo.org/record/13252170/files/figure.png" pageId="9" pageNumber="93">Figs. 2D</figureCitation> +, +<figureCitation id="B2B9E588FFD4FF99E4D92CB3FCFBF95B" box="[749,775,1702,1723]" captionStart="Fig" captionStartId="6.[175,206,1029,1048]" captionTargetBox="[156,1168,197,999]" captionTargetId="figure-241@6.[152,1172,193,1002]" captionTargetPageId="6" captionText="Fig. 3. Sphaeridium sundense, new species. A–D) Male genitalia:A) Abdominal tergite IX, dorsal, B) Median lobe and tegmen, C) Ventral detail of apex of median lobe, D) Lateral profile of median lobe and tegmen. E) Male protarsus, F) Mesoventral process, ventral.Abbreviations: bs = basal segment of tegmen; cor = corona; ls = lateral struts of tergite IX; mkp = mesoventral protrusion; ml = median lobe; mr = median ridge of ventrite IX; oc = outer claw; par = paramera; pml = posterior mesoventral lip; S5 = tarsomere 5; T10 = tergite X; V9 = sternite IX." figureDoi="http://doi.org/10.5281/zenodo.13252172" httpUri="https://zenodo.org/record/13252172/files/figure.png" pageId="9" pageNumber="93">3F</figureCitation> +, +<figureCitation id="B2B9E588FFD4FF99E5262CB3FCCCF95C" box="[786,816,1702,1724]" captionStart="Fig" captionStartId="8.[176,207,1360,1379]" captionTargetBox="[152,1172,193,1333]" captionTargetId="figure-172@8.[152,1172,193,1333]" captionTargetPageId="8" captionText="Fig. 5. Morphology of Sphaeridium species.A, C, E) Sphaeridium sundense, new species, B, D, F) S. quinquemaculatum. A–B) Procoxae, C–D) Mesoventral keel, ventral view, E–F) Mesoventral keel, oblique view." figureDoi="http://doi.org/10.5281/zenodo.13252176" httpUri="https://zenodo.org/record/13252176/files/figure.png" pageId="9" pageNumber="93">5C</figureCitation> +). In +<taxonomicName id="ED82828EFFD4FF99E56B2CB2FBD0F95B" box="[863,1068,1702,1724]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="93" phylum="Arthropoda" rank="species" species="quinquemaculatum"> +<emphasis id="18F6251FFFD4FF99E56B2CB2FBD0F95B" box="[863,1068,1702,1724]" italics="true" pageId="9" pageNumber="93">S. quinquemaculatum</emphasis> +</taxonomicName> +, the anterior protrusion is about as wide as long and the posterior lip is sparsely but distinctly pubescent ( +<figureCitation id="B2B9E588FFD7FF9AE4172AD7FD8FFF37" box="[547,627,193,216]" captionStart="Fig" captionStartId="8.[176,207,1360,1379]" captionTargetBox="[152,1172,193,1333]" captionTargetId="figure-172@8.[152,1172,193,1333]" captionTargetPageId="8" captionText="Fig. 5. Morphology of Sphaeridium species.A, C, E) Sphaeridium sundense, new species, B, D, F) S. quinquemaculatum. A–B) Procoxae, C–D) Mesoventral keel, ventral view, E–F) Mesoventral keel, oblique view." figureDoi="http://doi.org/10.5281/zenodo.13252176" httpUri="https://zenodo.org/record/13252176/files/figure.png" pageId="10" pageNumber="94">Fig. 5D</figureCitation> +). Additionally, the procoxae of +<taxonomicName id="ED82828EFFD7FF9AE78D2AC8FDD5FF11" authorityName="Suzumura & Henegouwen & Budi" authorityYear="2022" box="[441,553,220,242]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="94" phylum="Arthropoda" rank="species" species="sundense"> +<emphasis id="18F6251FFFD7FF9AE78D2AC8FDD5FF11" box="[441,553,220,242]" italics="true" pageId="10" pageNumber="94">S. sundense</emphasis> +</taxonomicName> +lack the abundant long yellowish pubescence interspersed among the strong spines as is seen in +<taxonomicName id="ED82828EFFD7FF9AE7D92B07FF25FEA1" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="94" phylum="Arthropoda" rank="species" species="quinquemaculatum"> +<emphasis id="18F6251FFFD7FF9AE7D92B07FF25FEA1" italics="true" pageId="10" pageNumber="94">S. quinquemaculatum</emphasis> +</taxonomicName> +( +<figureCitation id="B2B9E588FFD7FF9AE6DD2B39FEC9FEA2" box="[233,309,300,322]" captionStart="Fig" captionStartId="8.[176,207,1360,1379]" captionTargetBox="[152,1172,193,1333]" captionTargetId="figure-172@8.[152,1172,193,1333]" captionTargetPageId="8" captionText="Fig. 5. Morphology of Sphaeridium species.A, C, E) Sphaeridium sundense, new species, B, D, F) S. quinquemaculatum. A–B) Procoxae, C–D) Mesoventral keel, ventral view, E–F) Mesoventral keel, oblique view." figureDoi="http://doi.org/10.5281/zenodo.13252176" httpUri="https://zenodo.org/record/13252176/files/figure.png" pageId="10" pageNumber="94">Fig. 5B</figureCitation> +). +<taxonomicName id="ED82828EFFD7FF9AE77C2B39FD81FEA1" box="[328,637,300,321]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="94" phylum="Arthropoda" rank="species" species="quinquemaculatum"> +<emphasis id="18F6251FFFD7FF9AE77C2B39FD81FEA1" box="[328,637,300,321]" italics="true" pageId="10" pageNumber="94">Sphaeridium quinquemaculatum</emphasis> +</taxonomicName> +shows remarkable variation in color pattern, and some specimens are rather dark and quite similar to +<taxonomicName id="ED82828EFFD7FF9AE6AC2B68FEFBFE71" authorityName="Suzumura & Henegouwen & Budi" authorityYear="2022" box="[152,263,380,402]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="94" phylum="Arthropoda" rank="species" species="sundense"> +<emphasis id="18F6251FFFD7FF9AE6AC2B68FEFBFE71" box="[152,263,380,402]" italics="true" pageId="10" pageNumber="94">S. sundense</emphasis> +</taxonomicName> +. Differences in the extent of yellow on the lateral margins of the pronotum, however, are highly constant, with +<taxonomicName id="ED82828EFFD7FF9AE76B2BA7FDD5FE27" box="[351,553,434,455]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="94" phylum="Arthropoda" rank="species" species="quinquemaculatum"> +<emphasis id="18F6251FFFD7FF9AE76B2BA7FDD5FE27" box="[351,553,434,455]" italics="true" pageId="10" pageNumber="94">S. quinquemaculatum</emphasis> +</taxonomicName> +having a yellow border which does not continue around anterior and posterior angles, whereas the yellow border of +<taxonomicName id="ED82828EFFD7FF9AE6862817FEDCFDF7" authorityName="Suzumura & Henegouwen & Budi" authorityYear="2022" box="[178,288,514,535]" class="Insecta" family="Hydrophilidae" genus="Sphaeridium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="94" phylum="Arthropoda" rank="species" species="sundense"> +<emphasis id="18F6251FFFD7FF9AE6862817FEDCFDF7" box="[178,288,514,535]" italics="true" pageId="10" pageNumber="94">S. sundense</emphasis> +</taxonomicName> +does continue around the angles and reaches some distance inward from both anterior and posterior margins ( +<figureCitation id="B2B9E588FFD7FF9AE77D2822FE7EFDAC" box="[329,386,567,589]" captionStart="Fig" captionStartId="7.[197,228,990,1009]" captionTargetBox="[177,655,196,960]" captionTargetId="figure-561@7.[173,659,193,963]" captionTargetPageId="7" captionText="Fig. 4. Pronotal coloration, left lateral view. A) Sphaeridium sundense, new species, B) S. quinquemaculatum." figureDoi="http://doi.org/10.5281/zenodo.13252174" httpUri="https://zenodo.org/record/13252174/files/figure.png" pageId="10" pageNumber="94">Fig. 4</figureCitation> +). +</paragraph> +</subSubSection> +</treatment> +</document> \ No newline at end of file