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<document id="C1AFF135B64FE4646BE5EBE551C36EF4" ID-CLB-Dataset="35031" ID-DOI="http://dx.doi.org/10.3897/zookeys.400.6878" ID-GBIF-Dataset="a2263d9b-3085-4c8e-952c-7d0627e9aeda" ID-PMC="PMC4023241" ID-Pensoft-Pub="1313-2970-400-1" ID-PubMed="24843254" ID-ZooBank="E803925E0418463D863E183EDDAAA487" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-400-1" ModsDocOrigin="ZooKeys 400" ModsDocTitle="Review of the fish parasitic genus Ceratothoa Dana, 1852 (Crustacea, Isopoda, Cymothoidae) from South Africa, including the description of two new species" checkinTime="1451245997952" checkinUser="pensoft" docAuthor="Hadfield, Kerry A., Bruce, Niel L. &amp; Smit, Nico J." docDate="2014" docId="C2F18C373ECF6279BDF2E0CC04B92A9B" docLanguage="en" docName="ZooKeys 400: 1-42" docOrigin="ZooKeys 400" docSource="http://dx.doi.org/10.3897/zookeys.400.6878" docTitle="Ceratothoa imbricata Fabricius 1775" docType="treatment" docVersion="7" lastPageNumber="13" masterDocId="1421D967FF94FFF2FF88E144FFBBFF8C" masterDocTitle="Review of the fish parasitic genus Ceratothoa Dana, 1852 (Crustacea, Isopoda, Cymothoidae) from South Africa, including the description of two new species" masterLastPageNumber="42" masterPageNumber="1" pageNumber="11" updateTime="1701384070007" updateUser="ExternalLinkService">
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<mods:title id="5328760DEEE81F65989C78D9DD631864">Review of the fish parasitic genus Ceratothoa Dana, 1852 (Crustacea, Isopoda, Cymothoidae) from South Africa, including the description of two new species</mods:title>
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<mods:namePart id="3D1B0BA73033688647F8243D1FA644FB">Hadfield, Kerry A.</mods:namePart>
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<mods:namePart id="97D8313D1D9A896F8D632F5AA2828689">Bruce, Niel L.</mods:namePart>
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<mods:namePart id="BDBAA7D4DAD5E7ECD21CE8635289EA13">Smit, Nico J.</mods:namePart>
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<treatment id="C2F18C373ECF6279BDF2E0CC04B92A9B" ID-DOI="http://doi.org/10.5281/zenodo.3501623" ID-GBIF-Taxon="152052492" ID-Zenodo-Dep="3501623" LSID="urn:lsid:plazi:treatment:C2F18C373ECF6279BDF2E0CC04B92A9B" httpUri="http://treatment.plazi.org/id/C2F18C373ECF6279BDF2E0CC04B92A9B" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
<subSubSection id="3DC673E4221BD800376CD1833EC9BBD7" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="FDB4B5338FF28B4C80F3200553464280" pageId="10" pageNumber="11">
<taxonomicName id="1B2FD155C076AA6EE66C2EE927927E0A" ID-CoL="SQG4" LSID="http://species-id.net/wiki/Ceratothoa_imbricata" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa imbricata" order="Isopoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="imbricata">Ceratothoa imbricata (Fabricius, 1775)</taxonomicName>
Figs 15-18, 21
</paragraph>
</subSubSection>
<subSubSection id="AF92F8F3C3C08C0BFB3EFFAF48DE03BF" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="2D5A66AF1FED38B346B55FCD26CF21BA" pageId="10" pageNumber="11">
<taxonomicName id="4C049284877CB0E8939DFE3C23DE9B7A" class="Malacostraca" family="Oniscidae" genus="Oniscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oniscus umbricatus" order="Isopoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="umbricatus">Oniscus umbricatus</taxonomicName>
Fabricius, 1775: 296.
</paragraph>
<paragraph id="99C3096767010C2327D46BCE82883990" pageId="10" pageNumber="11">
<taxonomicName id="19EC538E5286608435B3BDC37D2F8DEE" class="Malacostraca" family="Oniscidae" genus="Oniscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oniscus imbricatus" order="Isopoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="imbricatus">Oniscus imbricatus</taxonomicName>
. -
<bibRefCitation id="54777CD8E703FAB499CBCFF33F68783C" author="Fabricius, JC" journalOrPublisher="Christ. Gottl. Proft., Hafniae" pageId="16" pageNumber="17" title="Mantissa Insectorum sistens eorum species nuper detectas adiectis characteribus genericus, differentiis specificis, emendationibus, observationibus." url="10.5962/bhl.title.36471" year="1787">Fabricius 1787</bibRefCitation>
: 241.
</paragraph>
<paragraph id="8F39C0033CCD91D9C4D9B0541ACC94AE" pageId="10" pageNumber="11">
<taxonomicName id="05CCB1BF0FF5A4A246EC30D0BC286280" class="Malacostraca" family="Cymothoidae" genus="Cymothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymothoa imbricata" order="Isopoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="imbricata">Cymothoa imbricata</taxonomicName>
. -
<bibRefCitation id="3536888603A26C63B0C0DC6F66362EF8" author="Fabricius, JC" journalOrPublisher="Christ. Gottl. Proft., Hafniae" pageId="17" pageNumber="18" title="Entomologia Systematica emendata et acuta, secundum classes, ordines, genera, species adjectis synonimis, locis, observationibus, descriptionibus." year="1793">Fabricius 1793</bibRefCitation>
: 503; 1798: 304.
</paragraph>
<paragraph id="90395E58A5531347C04BCCD476BB1205" pageId="10" pageNumber="11">
<taxonomicName id="315C894015241A9A0D801D17BB1F896E" class="Malacostraca" family="Cymothoidae" genus="Cymothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymothoa (Banksii)" order="Isopoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="subGenus" subGenus="Banksii">Cymothoa Banksii</taxonomicName>
Leach, 1818: 353.
</paragraph>
<paragraph id="5FA667B477F91E1BAC44F6E159797F95" pageId="10" pageNumber="11">
<taxonomicName id="FB123DC098A86171789C76F1530FCAD1" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa (Banksii)" order="Isopoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="subGenus" subGenus="Banksii">Ceratothoa Banksii</taxonomicName>
. -
<bibRefCitation id="25B100125D0FD4A80206E5E6E5477256" author="Schioedte, JC" journalOrPublisher="Kjobenhavn" pageId="19" pageNumber="20" pagination="281 - 378" title="Symbolae ad monographium Cymothoarum crustaceorum familiae. III. Saophridae. IV. Ceratothoinae" volumeTitle="Naturhistorisk Tidsskrift" year="1883">Schioedte and Meinert 1883</bibRefCitation>
: 340-347, tab. XIV (Cym. XXI), Figs 6-21.
</paragraph>
<paragraph id="BE8229CEF4247CCB4B1002614E1345BA" pageId="10" pageNumber="11">
<taxonomicName id="82FDFA3DE1F46D89D21D597CB8C49403" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa imbricata" order="Isopoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="imbricata">Ceratothoa imbricata</taxonomicName>
. -
<bibRefCitation id="EC58B74C4E99C0F7985CEA9A58640F53" author="Ellis, JP" journalOrPublisher="Bulletin of the British Museum (Natural History)" pageId="16" pageNumber="17" pagination="121 - 128" title="Some type specimens of Isopoda (Flabellifera) in the British Museum (Natural History), and the isopods in the Linnaean Collection." volume="40" year="1981">Ellis 1981</bibRefCitation>
: 123.
</paragraph>
<paragraph id="1D78D1E23D253A601BFB937DF4702A25" pageId="10" pageNumber="11">
<taxonomicName id="769709098F7C812A7A8801C26D147B15" class="Malacostraca" family="Cymothoidae" genus="Codonophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Codonophilus imbricatus" order="Isopoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="imbricatus">Codonophilus imbricatus</taxonomicName>
. -
<bibRefCitation id="A3A5E8ECFC2920D3308526A0FBD7F450" author="Hale, HM" journalOrPublisher="Transactions of the Royal Society of South Australia" pageId="17" pageNumber="18" pagination="201 - 234" title="Review of Australian isopods of the cymothoid group. Part II." volume="50" year="1926">Hale 1926</bibRefCitation>
: 223-226, Figs 15-16;
<bibRefCitation id="3A5842A25B21BAFA2B356A0835B6FB7E" author="Hale, HM" journalOrPublisher="Transactions and proceedings of the Royal Society of South Australia" pageId="17" pageNumber="18" pagination="307 - 321" title="The fauna of Kangaroo Island, South Australia. 1. The Crustacea." volume="51" year="1927">1927</bibRefCitation>
: 315;
<bibRefCitation id="3CEB79DCEE7C188421991A1113BCAF5A" author="Hale, HM" journalOrPublisher="Weir, Government Printer, Adelaide" pageId="17" pageNumber="18" pagination="202 - 380" title="The Crustaceans of South Australia. Part II" year="1929">1929</bibRefCitation>
: 263-264, fig. 262;
<bibRefCitation id="169DC5578C2406D469A3D3BC8706E7CB" author="Hale, HM" journalOrPublisher="Australasian Antarctic Expedition 1911 - 14 Under the leadership of Sir Douglas Mawson, OBE, BE, DSc, FRS Scientific Reports (Series C - Zoology and Botany)" pageId="17" pageNumber="18" pagination="1 - 45" title="Isopoda and Tanaidacea" volume="2" year="1937">1937</bibRefCitation>
: 19;
<bibRefCitation id="E6D9BF29CD762E0EBE9BE20C5D41BD99" author="Hale, HM" journalOrPublisher="Transactions of the Royal Society of South Australia" pageId="17" pageNumber="18" pagination="288 - 304" title="Report on the cymothoid Isopoda obtained by the F. I. S. &quot; Endeavour &quot; on the coasts of Queensland, New South Wales, Victoria, Tasmania, and South Australia." volume="64" year="1940">1940</bibRefCitation>
: 303.
</paragraph>
<paragraph id="3018E1E5A9D64D7BADBCD06E7A97A6B6" pageId="10" pageNumber="11">
<taxonomicName id="3F73CF9021388AF99DFEAD79AE137417" class="Malacostraca" family="Cymothoidae" genus="Cymothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymothoa banksii" order="Isopoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="banksii">Cymothoa banksii</taxonomicName>
. -
<bibRefCitation id="E1F17F65DBA1C280A81EBB02A57BBE2A" author="Ellis, JP" journalOrPublisher="Bulletin of the British Museum (Natural History)" pageId="16" pageNumber="17" pagination="121 - 128" title="Some type specimens of Isopoda (Flabellifera) in the British Museum (Natural History), and the isopods in the Linnaean Collection." volume="40" year="1981">Ellis 1981</bibRefCitation>
: 124.
</paragraph>
</subSubSection>
<subSubSection id="3C7CAF77E29402BBB3CE8881C5958CD2" pageId="10" pageNumber="11" type="material examined">
<paragraph id="DB963720D97AA3BA8F24B4CEEBED4804" pageId="10" pageNumber="11">Material examined.</paragraph>
<paragraph id="4FBD174459F9A4AFF538D42E5F053DF1" pageId="10" pageNumber="11">
Holotype of
<taxonomicName id="EB83656C59433DC5B8F337B571FC4BC6" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa imbricata" order="Isopoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="imbricata">Ceratothoa imbricata</taxonomicName>
. The Natural History Museum, London (BMNH 1979.403.1) - female (34 mm TL; 16 mm W) collection of Sir Joseph Banks, Linnean Society, from New Zealand, coll. S.W.J. Banks, host unknown (
<bibRefCitation id="78566EB4BB59D30C89189300A0AA7601" author="Fabricius, JC" journalOrPublisher="Officina Libraria Kortii, Flensburgi &amp; Lipsiae" pageId="16" pageNumber="17" title="Systema Entomologiae, sistens Insectorum Classes, Ordines, Genera, Species, adiectis Synonymis, Locis, Descriptionibus, Observationibus" url="10.5962/bhl.title.36510" year="1775">Fabricius 1775</bibRefCitation>
). Noted: there is a hole in pereonite 4 and 5.
</paragraph>
<paragraph id="F708AF87793BB5740280BD3D29327D57" pageId="10" pageNumber="11">
Holotype of
<taxonomicName id="C0FF7D39FF185EE305820A8B6800BD30" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa banksii" order="Isopoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="banksii">Ceratothoa banksii</taxonomicName>
. The Natural History Museum, London (BMNH 1979.402.1) - female (37 mm TL; 18 mm W), presented by Leach to the Museum of the Linnean Society, from New Zealand,
<normalizedToken id="1C4C2F529D4700127208179671B025D6" originalValue="White’s">White's</normalizedToken>
MS Cat No. 222, Coll. W.E. Leach, host unknown.
</paragraph>
</subSubSection>
<subSubSection id="9F1B7E6DA7651656839DFAAC7B43E0B8" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="description of holotype">
<paragraph id="AB594F5377F3291B3A483A0DF2DA3B83" pageId="10" pageNumber="11">Description of holotype.</paragraph>
<paragraph id="121BEEB885F326231001322091B2D2C3" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
Body ovoid, 2.1 times as long as greatest width, dorsal surfaces slightly bumpy, widest at pereonite 5, most narrow at pereonite 1, lateral margins posteriorly ovate. Cephalon 0.7 times longer than wide, visible from dorsal view, triangular. Frontal margin rounded to form blunt rostrum. Eyes oval with distinct margins. Pereonite 1 with slight indentations, anterior border straight, anterolateral angle with distinct produced point extending to or beyond the eye margin, posterior
<pageBreakToken id="0834D0EAA5ECC8630889A60B65976415" pageId="11" pageNumber="12" start="start">margins</pageBreakToken>
of pereonites smooth and slightly curved laterally. Pereonites 1-5 increasing in length and width; 6-7 decreasing in length and width; 6 and 7 narrower. Pleon with pleonite 1 most narrow, visible in dorsal view; pleonites posterior margin smooth, mostly concave; posterolateral angles of pleonite 2 narrowly rounded, not posteriorly produced. Pleonites 3-5 similar in form to pleonite 2. Pleonite 5 with posterolateral angles free, not overlapped by lateral margins of pleonite 4, posterior margin produced medially. Pleotelson 2 times as long as anterior width, dorsal surface with lateral indent, lateral margins weakly convex, posterior margin rounded, without median point. Antennule more stout than antenna, comprised of 8 articles. Antenna comprised of 4 articles. Pereopod 1 basis 1.5 times as long as greatest width; ischium 0.8 times as long as basis; merus proximal margin with bulbous protrusion; carpus with straight proximal margin; propodus 1.5 times as long as wide; dactylus slender, 0.9 as long as propodus, 2.3 times as long as basal width. Pereopod 2 propodus 1.3 as long as wide; dactylus 0.6 as long as propodus. Pereopods 3 similar to pereopod 2. Pereopod 6 basis 1.4 times as long as greatest width, ischium 0.7 times as long as basis, propodus 1.5 as long as wide, dactylus 1 as long as propodus. Pereopod 7 basis 1.2 times as long as greatest width; ischium 0.7 as long as basis, without protrusions; merus proximal margin with slight bulbous protrusion, merus 0.4 as long as ischium, 0.6 times as long as wide; carpus 0.6 as long as ischium, without bulbous protrusion, 0.5 times as long as wide; propodus 0.6 as long as ischium, 1.4 times as long as wide; dactylus slender, 1.1 as long as propodus, 2.5 times as long as basal width. Uropod longer than the pleotelson, peduncle 0.7 times longer than rami, peduncle lateral margin without setae; rami extending beyond pleotelson, marginal setae absent, apices narrowly rounded.
</paragraph>
<caption id="8A9CB5734CAE7D43055518ACBA7FF6E0" pageId="11" pageNumber="12">
<paragraph id="BCDC9CA5C62FB433C738F8EBAE42C54B" pageId="11" pageNumber="12">
Figure 15.
<taxonomicName id="FB3279FA2763643EE63AD16E69CD89C2" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa imbricata" order="Isopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="imbricata">Ceratothoa imbricata</taxonomicName>
(Fabricius, 1775), female holotype (34 mm) (BMNH 1979.403.1): A dorsal view B antero-dorsal view of pereonite 1 and cephalon C dorsal view of pleotelson D lateral view.
</paragraph>
</caption>
<caption id="F5608049A00474C4B4A7F88C44C27E14" pageId="11" pageNumber="12">
<paragraph id="673B24D0114DF63B42557DAC58919920" pageId="11" pageNumber="12">
Figure 16.
<taxonomicName id="43DF36100EB2580BF51350E14802D81F" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa imbricata" order="Isopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="imbricata">Ceratothoa imbricata</taxonomicName>
(Fabricius, 1775), female holotype (34 mm) (BMNH 1979.403.1): A pereopod 1 B pereopod 2 C pereopod 6 D pereopod 7.
</paragraph>
</caption>
<caption id="68A8A37CAD7D08796B5AE1EE4037A072" pageId="11" pageNumber="12">
<paragraph id="2CCFD303C86D2BAF9E28B38C781B8717" pageId="11" pageNumber="12">
Figure 17.
<taxonomicName id="62C7A1629C4256C97714A015B34E0AA4" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa imbricata" order="Isopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="imbricata">Ceratothoa imbricata</taxonomicName>
(Fabricius, 1775), female (37 mm), (BMNH 1979.402.1 originally designated as holotype of
<taxonomicName id="A0496A56E23E7C074F92CC758D7D0CDC" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa banksii" order="Isopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="banksii">Ceratothoa banksii</taxonomicName>
Leach, 1818): A dorsal view B antero-dorsal view of pereonite 1 and cephalon C ventral view of cephalon D dorsal view of pleotelson E lateral view.
</paragraph>
</caption>
<caption id="E20FB59ACD9975070E718679933418D6" pageId="11" pageNumber="12">
<paragraph id="E05265BE90EED815D8848B696D1EC2BD" pageId="11" pageNumber="12">
Figure 18.
<taxonomicName id="A8E785CE50930594898A7401A2BEAE17" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa imbricata" order="Isopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="imbricata">Ceratothoa imbricata</taxonomicName>
(Fabricius, 1775), female (37 mm), (BMNH 1979.402.1 originally designated as holotype of
<taxonomicName id="542612D1A6EF9A7A6A4042B0B99B07B3" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa banksii" order="Isopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="banksii">Ceratothoa banksii</taxonomicName>
Leach, 1818): A pereopod 1 B pereopod 2 C pereopod 6 D pereopod 7.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="E9725E141FBFB4D5D8AE513BB5E08019" pageId="11" pageNumber="12" type="distribution">
<paragraph id="C882E5594F49B776D7A51AA6416C139B" pageId="11" pageNumber="12">Distribution.</paragraph>
<paragraph id="2C58BDC56A9479A3BA6D9E7AFE5B2CF0" pageId="11" pageNumber="12">
Australia (
<bibRefCitation id="9CEDEEB5857F0FE161B45A5ACA05EF6D" author="Schioedte, JC" journalOrPublisher="Kjobenhavn" pageId="19" pageNumber="20" pagination="281 - 378" title="Symbolae ad monographium Cymothoarum crustaceorum familiae. III. Saophridae. IV. Ceratothoinae" volumeTitle="Naturhistorisk Tidsskrift" year="1883">Schioedte and Meinert 1883</bibRefCitation>
,
<bibRefCitation id="64915958DBDE9AD0BEA016EEF566B5C9" author="Miers, EJ" journalOrPublisher="BMNH, London" pageId="18" pageNumber="19" pagination="299 - 311" title="Crustacea. Report on the zoological collections made in the Indo-Pacific Ocean during the voyage of HMS &quot; Alert &quot;, 1881 - 1882" year="1884">Miers 1884</bibRefCitation>
,
<bibRefCitation id="5E010A2CCA7DC92191220523B808127A" author="Hale, HM" journalOrPublisher="Transactions and proceedings of the Royal Society of South Australia" pageId="17" pageNumber="18" pagination="307 - 321" title="The fauna of Kangaroo Island, South Australia. 1. The Crustacea." volume="51" year="1927">Hale 1927</bibRefCitation>
,
<bibRefCitation id="B02E113F5D23F9989674E8BD6256DEA6" author="Hale, HM" journalOrPublisher="Transactions of the Royal Society of South Australia" pageId="17" pageNumber="18" pagination="288 - 304" title="Report on the cymothoid Isopoda obtained by the F. I. S. &quot; Endeavour &quot; on the coasts of Queensland, New South Wales, Victoria, Tasmania, and South Australia." volume="64" year="1940">1940</bibRefCitation>
), New Zealand (
<bibRefCitation id="4CB93E4E96802A299CD74D3E146924A4" author="Fabricius, JC" journalOrPublisher="Officina Libraria Kortii, Flensburgi &amp; Lipsiae" pageId="16" pageNumber="17" title="Systema Entomologiae, sistens Insectorum Classes, Ordines, Genera, Species, adiectis Synonymis, Locis, Descriptionibus, Observationibus" url="10.5962/bhl.title.36510" year="1775">Fabricius 1775</bibRefCitation>
,
<bibRefCitation id="46245DBBD71221213174021FFE48F318" author="Fabricius, JC" journalOrPublisher="Christ. Gottl. Proft., Hafniae" pageId="17" pageNumber="18" title="Entomologia Systematica emendata et acuta, secundum classes, ordines, genera, species adjectis synonimis, locis, observationibus, descriptionibus." year="1793">1793</bibRefCitation>
) and Indonesia (
<bibRefCitation id="52B7D34908027D6FDD825B646E9A6599" author="Schioedte, JC" journalOrPublisher="Kjobenhavn" pageId="19" pageNumber="20" pagination="281 - 378" title="Symbolae ad monographium Cymothoarum crustaceorum familiae. III. Saophridae. IV. Ceratothoinae" volumeTitle="Naturhistorisk Tidsskrift" year="1883">Schioedte and Meinert 1883</bibRefCitation>
).
</paragraph>
<paragraph id="1115AFBD6EEA9AF32793F37734CE6FF7" pageId="11" pageNumber="12">
<bibRefCitation id="E38E9E444FA177EBCE1EEB681BB6390B" author="Yu, H" journalOrPublisher="Studia Marina Sinica" pageId="19" pageNumber="20" pagination="223 - 238" title="Study on the Cymothoidae from Chinese waters" volume="45" year="2003">Yu and Li (2003)</bibRefCitation>
included a figure of a specimen described as
<taxonomicName id="771B46BAC0E50870F037E66D81971220" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa imbricata" order="Isopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="imbricata">Ceratothoa imbricata</taxonomicName>
from Chinese waters; the figures show that the antenna bases do not touch, which excludes the species from
<taxonomicName id="4B1101BA590CA50A197D3DCD52AB57ED" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa" order="Isopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">Ceratothoa</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="CE90FB8A979F8152663F2A08EB19CA28" pageId="11" pageNumber="12" type="hosts">
<paragraph id="E89722D530EB57AE34D1DE64A56226A9" pageId="11" pageNumber="12">Hosts.</paragraph>
<paragraph id="FFDB803EA087B7201914D3C71A3B151E" pageId="11" pageNumber="12">
From the mouth of a
<normalizedToken id="7088742F2B638D4DD123D0AC3B7006AF" originalValue="salmon–trout">salmon-trout</normalizedToken>
; from a
<taxonomicName id="D748D3C4A7CBA0260738BFE456E9B05E" family="Monacanthidae" genus="Monacanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monacanthus" order="Tetraodontiformes" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">Monacanthus</taxonomicName>
sp. (
<bibRefCitation id="D43994FC9D415DAEEAACF5BF8F003A40" author="Miers, EJ" journalOrPublisher="BMNH, London" pageId="18" pageNumber="19" pagination="299 - 311" title="Crustacea. Report on the zoological collections made in the Indo-Pacific Ocean during the voyage of HMS &quot; Alert &quot;, 1881 - 1882" year="1884">Miers 1884</bibRefCitation>
); in the Australian jack mackerel,
<taxonomicName id="8366B31AB205C7261E452C830EA2C718" family="Carangidae" genus="Trachurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trachurus declivis" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="declivis">Trachurus declivis</taxonomicName>
(
<bibRefCitation id="7184B1D8921758927A97E9AE722EEDB7" author="Hale, HM" journalOrPublisher="Transactions of the Royal Society of South Australia" pageId="17" pageNumber="18" pagination="201 - 234" title="Review of Australian isopods of the cymothoid group. Part II." volume="50" year="1926">Hale 1926</bibRefCitation>
,
<bibRefCitation id="AC98002404AACDC5EF880B906865C90B" author="Hale, HM" journalOrPublisher="Weir, Government Printer, Adelaide" pageId="17" pageNumber="18" pagination="202 - 380" title="The Crustaceans of South Australia. Part II" year="1929">1929</bibRefCitation>
); in snapper
<taxonomicName id="8301A5AF715C2DA1DE0B311D83FDD84F" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chrysophrys auratus" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="auratus">Chrysophrys auratus</taxonomicName>
(previously
<taxonomicName id="5533A4456F6D7273A71DFC12E1B18ACB" family="Sparidae" genus="Pagrosomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pagrosomus auratus" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="auratus">Pagrosomus auratus</taxonomicName>
), red gurnard (
<taxonomicName id="8F29AA01C3DE3C6524EB6A73A7BD8433" family="Triglidae" genus="Chelidonichthys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chelidonichthys kumu" order="Scorpaeniformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="kumu">Chelidonichthys kumu</taxonomicName>
), and mullet (
<taxonomicName id="1E86F150738E3AF70332C45AA3D2E689" family="Mugilidae" genus="Mugil" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mugil" order="Mugiliformes" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">Mugil</taxonomicName>
sp.) (
<bibRefCitation id="FA726BCACE473A9E2BD1DB158C802EA6" author="Hale, HM" journalOrPublisher="Transactions of the Royal Society of South Australia" pageId="17" pageNumber="18" pagination="201 - 234" title="Review of Australian isopods of the cymothoid group. Part II." volume="50" year="1926">Hale 1926</bibRefCitation>
,
<bibRefCitation id="922194E197C00365BC8F96A2AB0D7049" author="Hale, HM" journalOrPublisher="Weir, Government Printer, Adelaide" pageId="17" pageNumber="18" pagination="202 - 380" title="The Crustaceans of South Australia. Part II" year="1929">1929</bibRefCitation>
); in mouth of
<taxonomicName id="3718895EB39075B43B190E7AF2C7FEBF" family="Kyphosidae" genus="Girella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Girella tricuspidata" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="tricuspidata">Girella tricuspidata</taxonomicName>
(
<bibRefCitation id="D81F40B8F919921AFC28AAA9CAC8CF20" author="Hale, HM" journalOrPublisher="Transactions of the Royal Society of South Australia" pageId="17" pageNumber="18" pagination="201 - 234" title="Review of Australian isopods of the cymothoid group. Part II." volume="50" year="1926">Hale 1926</bibRefCitation>
,
<bibRefCitation id="A1EA053E14645938F2EC0A4A0F2A211F" author="Hale, HM" journalOrPublisher="Weir, Government Printer, Adelaide" pageId="17" pageNumber="18" pagination="202 - 380" title="The Crustaceans of South Australia. Part II" year="1929">1929</bibRefCitation>
); trevally
<taxonomicName id="5A75881298F5B0B75D9EAC470EB63B6C" genus="Psuedocaranx" lsidName="Psuedocaranx dentex" pageId="11" pageNumber="12" rank="species" species="dentex">Psuedocaranx dentex</taxonomicName>
(previously
<taxonomicName id="8C132DFAAB8EBA59044A97768A0148AE" family="Carangidae" genus="Caranx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caranx georgianus" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="georgianus">Caranx georgianus</taxonomicName>
) (
<bibRefCitation id="D5367205E7E74A18FECE7F0AF900A20F" author="Hale, HM" journalOrPublisher="Transactions of the Royal Society of South Australia" pageId="17" pageNumber="18" pagination="201 - 234" title="Review of Australian isopods of the cymothoid group. Part II." volume="50" year="1926">Hale 1926</bibRefCitation>
,
<bibRefCitation id="F1E27B046760685F370F20529C101456" author="Hale, HM" journalOrPublisher="Weir, Government Printer, Adelaide" pageId="17" pageNumber="18" pagination="202 - 380" title="The Crustaceans of South Australia. Part II" year="1929">1929</bibRefCitation>
).
</paragraph>
<paragraph id="53E040D70D9E0D511C5BD6D21D70ECA6" pageId="11" pageNumber="12">
<bibRefCitation id="5E97DE8D802182283D06A3AEC831D9BA" author="Schioedte, JC" journalOrPublisher="Kjobenhavn" pageId="19" pageNumber="20" pagination="281 - 378" title="Symbolae ad monographium Cymothoarum crustaceorum familiae. III. Saophridae. IV. Ceratothoinae" volumeTitle="Naturhistorisk Tidsskrift" year="1883">Schioedte and Meinert (1883)</bibRefCitation>
mention a fish they thought may be a &quot;red hottentot (
<taxonomicName id="9BA5EE5A6DD015A7BB3625689999EEC9" genus="Sargi" lsidName="Sargi hottentotti" pageId="11" pageNumber="12" rank="species" species="hottentotti">Sargi hottentotti</taxonomicName>
Sm.??)&quot; collected from the Cape of Good Hope in South Africa. This could refer to the red roman fish,
<taxonomicName id="7D280C5E0B7310A0D1EA3CF3B1FC54F0" family="Sparidae" genus="Chrysoblephus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chrysoblephus laticeps" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="laticeps">Chrysoblephus laticeps</taxonomicName>
(see
<bibRefCitation id="82B7DD46EA27178ADF295D19EA03783D" author="Kensley, B" journalOrPublisher="South African Museum &amp; The Rustica Press, Wynberg, Cape Town" pageId="18" pageNumber="19" title="Guide to the Marine Isopods of Southern Africa." year="1978">Kensley 1978</bibRefCitation>
), the Zebra (
<taxonomicName id="D61435B58CC16637A875C1399172BD3F" family="Sparidae" genus="Diplodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplodus cervinus subsp. hottentotus" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="subSpecies" species="cervinus" subSpecies="hottentotus">Diplodus cervinus hottentotus</taxonomicName>
) or the Hottentot (
<taxonomicName id="B48F3F916127928ECE6208519C540C37" family="Sparidae" genus="Pachymetopon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachymetopon blochii" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="blochii">Pachymetopon blochii</taxonomicName>
) as &quot;
<taxonomicName id="3A22BA1902A2C364A46BF164C3A8BED5" genus="Sargi" lsidName="Sargi hottentotti" pageId="11" pageNumber="12" rank="species" species="hottentotti">Sargi hottentotti</taxonomicName>
&quot; is not a valid taxonomic name and cannot be found in current fish database searches. No fresh material or museum material of these specimens from the red roman or those collected by
<bibRefCitation id="6339B1CEEA1A8865E1F936325E79BD8D" author="Kensley, B" journalOrPublisher="South African Museum &amp; The Rustica Press, Wynberg, Cape Town" pageId="18" pageNumber="19" title="Guide to the Marine Isopods of Southern Africa." year="1978">Kensley (1978)</bibRefCitation>
could be found so these records are not accepted.
</paragraph>
</subSubSection>
<subSubSection id="5A344D3AF14EC015E19EC13EF870D981" pageId="12" pageNumber="13" type="remarks">
<paragraph id="659C8B814DFB38DD6381BC32EAD4AFF7" pageId="12" pageNumber="13">
<pageBreakToken id="0C0C11C9E9FCFA320800338F7D04A470" pageId="12" pageNumber="13" start="start">Remarks</pageBreakToken>
.
</paragraph>
<paragraph id="42C9647B3D5B62D3F78A4CB35160BE46" pageId="12" pageNumber="13">
<taxonomicName id="BC18294F0F5589116CA64B3723AD76E7" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa imbricata" order="Isopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="imbricata">Ceratothoa imbricata</taxonomicName>
can be identified by a large pereonite 1 with anterolateral margins extending past the eyes; uropods as long or longer than the pleotelson margin; merus with bulbous protrusion; a blunt rostrum; and body widest at pereonite 5.
</paragraph>
<paragraph id="AA3E4F36C3548A341533E29C1DE1C59E" pageId="12" pageNumber="13">
It is apparent that over the years there have been many misidentifications of
<taxonomicName id="8C3B390C16DAB5F17FD8914C4871F5F7" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa imbricata" order="Isopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="imbricata">Ceratothoa imbricata</taxonomicName>
,
<taxonomicName id="D934482D7569C2F20AAD01F5DB0915AF" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa banksii" order="Isopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="banksii">Ceratothoa banksii</taxonomicName>
and
<taxonomicName id="9F9763CE474F79750814B02F72BC073E" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa trigonocephala" order="Isopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="trigonocephala">Ceratothoa trigonocephala</taxonomicName>
, with these names being widely misapplied. The description of
<taxonomicName id="6CEB5A38E07780FF0EC84D7F72A685F2" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa banksii" order="Isopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="banksii">Ceratothoa banksii</taxonomicName>
from New Zealand, given by
<bibRefCitation id="135938CBF4233E20AA1FA3A81406A89F" author="Miers, EJ" journalOrPublisher="Colonial Museum and Geological Department of New Zealand, National History Publication" pageId="18" pageNumber="19" pagination="1 - 133" title="Catalogue of the stalk and sessile-eyed Crustacea of New Zealand." volume="10" year="1876">Miers (1876)</bibRefCitation>
can also be applied to the small Australian
<taxonomicName id="21EEF7E2B29DED4CDF7FF151EAC8AFE2" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa imbricata" order="Isopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="imbricata">Ceratothoa imbricata</taxonomicName>
specimens of
<bibRefCitation id="2481177CF5DE67CA3E0AFE01E3832CFA" author="Miers, EJ" journalOrPublisher="BMNH, London" pageId="18" pageNumber="19" pagination="299 - 311" title="Crustacea. Report on the zoological collections made in the Indo-Pacific Ocean during the voyage of HMS &quot; Alert &quot;, 1881 - 1882" year="1884">Miers (1884)</bibRefCitation>
, with only some slight variations in eyes, smaller anterolateral extensions on pereonite, 1 and a slightly arched pleotelson posterior margin. The original description of
<taxonomicName id="006209C722253FB96D7E626EFFF0CE34" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa banksii" order="Isopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="banksii">Ceratothoa banksii</taxonomicName>
by
<bibRefCitation id="02AB88769F87201F7141C21E56CB5DE8" author="Leach, WE" editor="Cuvier, F" journalOrPublisher="Strasbourg et Levrault, Paris" pageId="18" pageNumber="19" pagination="338 - 354" title="Cymothoadees." volumeTitle="Dictionnaire des Sciences Naturelles" year="1818">Leach (1818)</bibRefCitation>
also described the pleotelson as &quot;nearly straight&quot; but according to
<bibRefCitation id="755031EE116E9E1A14AEF3621C34C111" author="Miers, EJ" journalOrPublisher="BMNH, London" pageId="18" pageNumber="19" pagination="299 - 311" title="Crustacea. Report on the zoological collections made in the Indo-Pacific Ocean during the voyage of HMS &quot; Alert &quot;, 1881 - 1882" year="1884">Miers (1884)</bibRefCitation>
, desiccation had caused the specimen to roll slightly. Many authors agreed with the synonymy of
<taxonomicName id="CFDC0B46B110E816358E529AB77F0272" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa banksii" order="Isopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="banksii">Ceratothoa banksii</taxonomicName>
with
<taxonomicName id="D8EAC41E6FD80028E1C2BDDB2B70B754" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa imbricata" order="Isopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="imbricata">Ceratothoa imbricata</taxonomicName>
including
<bibRefCitation id="4D4AD948C6C16E6B7D2AF5DD55FB4AD2" author="Stebbing, TRR" journalOrPublisher="Kegan Paul, Trench, Trubner &amp; Co. Ltd., London" pageId="19" pageNumber="20" title="A History of Crustacea. Recent Malacostraca." year="1893">Stebbing (1893)</bibRefCitation>
,
<bibRefCitation id="E1DCF34057AC988164B96144CC69FCC0" author="Nierstrasz, HF" journalOrPublisher="Zoologische Mededelingen (Leiden)" pageId="19" pageNumber="20" pagination="71 - 108" title="Die Isopoden-Sammlung im Naturhistorischen Reichsmuseum zu Leiden - 1. Cymothoidae" volume="1" year="1915">Nierstrasz (1915)</bibRefCitation>
and
<bibRefCitation id="B5261D82C5A02B1CC3C26F3A58FBA124" author="Trilles, J-P" journalOrPublisher="Paris" pageId="19" pageNumber="20" pagination="1231 - 1268" title="Les Cymothoidae (Isopoda, Flabellifera) du Museum national d'Histoire naturelle de Paris. Etude critique accompagnee de precisions en particulier sur la repartition geographique et l'ecologie des differentes especes representees. I. Les Ceratothoinae Schioedte &amp; Meinert, 1883." volumeTitle="Bulletin du Museum national d'Histoire naturelle" year="1973">Trilles (1973)</bibRefCitation>
and we maintain this synonymy, however this needs further investigation, especially when fresh material becomes available.
</paragraph>
<paragraph id="8591B3B0690742A09071165D5EFC6D3C" pageId="12" pageNumber="13">
<bibRefCitation id="41672182C4DB3FDEE4842D04AA475F26" author="Trilles, J-P" journalOrPublisher="Studia Marina" pageId="19" pageNumber="20" pagination="1 - 288" title="Les Cymothoidae (Crustacea, Isopoda) du Monde. Podrome pour une faune." volume="21 / 22" year="1994">Trilles (1994)</bibRefCitation>
placed
<bibRefCitation id="C57CD29FE1C475D6272B97D56F450D0C" author="Hale, HM" journalOrPublisher="Transactions of the Royal Society of South Australia" pageId="17" pageNumber="18" pagination="201 - 234" title="Review of Australian isopods of the cymothoid group. Part II." volume="50" year="1926">
<normalizedToken id="AB7E0FBB97DD44DA86E3C17D9133C42C" originalValue="Hale’s">Hale's</normalizedToken>
(1926
</bibRefCitation>
,
<bibRefCitation id="DF93EF32F4321D9BBD288BAF805AE8F1" author="Hale, HM" journalOrPublisher="Transactions and proceedings of the Royal Society of South Australia" pageId="17" pageNumber="18" pagination="307 - 321" title="The fauna of Kangaroo Island, South Australia. 1. The Crustacea." volume="51" year="1927">1927</bibRefCitation>
,
<bibRefCitation id="E96AC268F2643D4334185FE8FDCA460A" author="Hale, HM" journalOrPublisher="Weir, Government Printer, Adelaide" pageId="17" pageNumber="18" pagination="202 - 380" title="The Crustaceans of South Australia. Part II" year="1929">1929</bibRefCitation>
,
<bibRefCitation id="918AE7B3FF656EEC1DF9ADE30510ACB1" author="Hale, HM" journalOrPublisher="Transactions of the Royal Society of South Australia" pageId="17" pageNumber="18" pagination="288 - 304" title="Report on the cymothoid Isopoda obtained by the F. I. S. &quot; Endeavour &quot; on the coasts of Queensland, New South Wales, Victoria, Tasmania, and South Australia." volume="64" year="1940">1940</bibRefCitation>
) records of
<taxonomicName id="A6C09BECCFFDB4D611E0BAF324A749E3" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa imbricata" order="Isopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="imbricata">Ceratothoa imbricata</taxonomicName>
into synonymy with
<taxonomicName id="FB4FCF3D6E97C79286E1699A994E8761" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa trigonocephala" order="Isopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="trigonocephala">Ceratothoa trigonocephala</taxonomicName>
. After reviewing
<bibRefCitation id="B4D1D4EC031C8B407E312F1499B61950" author="Hale, HM" journalOrPublisher="Transactions of the Royal Society of South Australia" pageId="17" pageNumber="18" pagination="201 - 234" title="Review of Australian isopods of the cymothoid group. Part II." volume="50" year="1926">
<normalizedToken id="ABD347CF5794EA07669F3AA0736F2DE8" originalValue="Hale’s">Hale's</normalizedToken>
(1926)
</bibRefCitation>
figures, we conclude that his original identification of
<taxonomicName id="9DD3CDF083BE98EC3173CA3A35AC0E84" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa imbricata" order="Isopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="imbricata">Ceratothoa imbricata</taxonomicName>
is correct.
</paragraph>
<paragraph id="4E1F2B6173B908F13A43B8BE8DB6B04D" pageId="12" pageNumber="13">
No South African specimens were found, fresh or from museum collections, that could be identified as
<taxonomicName id="3EBEE5B61509EB7AE7F0A7AE68203EE1" class="Malacostraca" family="Cymothoidae" genus="Ceratothoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratothoa imbricata" order="Isopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="imbricata">Ceratothoa imbricata</taxonomicName>
, and the species is here excluded from the South Africa fauna.
</paragraph>
</subSubSection>
</treatment>
</document>