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: Willmann 1923
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Diagnose.
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des
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470-550 µm. Sensillus leicht
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und mit einem Fortsatz endend, mit apikalen Dornen. Dorsalborsten glatt, gerade und spitz endend. 6-8 Paar Genitalborsten. 1 Paar Analborsten, 3 Paar Adanalborsten.
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Verbreitung und
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. Holarktische Art. Lebt vor allem in feuchtem Moos und Flechten, kommt aber auch in Waldstreu und in
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vor.
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aus der Kneissl-Sammlung
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1. [K 1033,
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sp.]; 1 ex (ad), (C), det. A. Szywilewska.
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Bemerkung: Im Hinblick auf
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Prodorsalborsten Bestimmung nicht sicher.
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2. [K 1034,
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sp.]; 1 ex (ad), (C), det. A. Szywilewska.
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Bemerkung: Im Hinblick auf
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Prodorsalborsten Bestimmung nicht sicher.
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<taxonomicName authorityName="Osten Sacken" authorityYear="1860" class="Insecta" family="Limoniidae" genus="Elephantomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elephantomyia" order="Diptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">Elephantomyia</taxonomicName>
Osten Sacken, 1860: 220;
<bibRefCitation author="Alexander, CP" journalOrPublisher="Revista de Entomologia" pageId="0" pageNumber="41" pagination="509 - 556" refId="B11" refString="Alexander, CP, 1948. Notes on the tropical American species of Tipulidae (Diptera). V. The specialized Hexatomini: Limnophila, Shannonomyia, Gynoplistia, Hexatoma, Atarba, Elephantomyia, and allies. Revista de Entomologia 19: 509 - 556" title="Notes on the tropical American species of Tipulidae (Diptera). V. The specialized Hexatomini: Limnophila, Shannonomyia, Gynoplistia, Hexatoma, Atarba, Elephantomyia, and allies." volume="19" year="1948">Alexander 1948</bibRefCitation>
: 522;
<bibRefCitation author="Ito, S" journalOrPublisher="Mushi" pageId="0" pageNumber="41" pagination="89 - 92" refId="B15" refString="Ito, S, 1948. Two undescribed species of the tipulid genus Elephantomyia from Japan. Mushi 18: 89 - 92" title="Two undescribed species of the tipulid genus Elephantomyia from Japan." volume="18" year="1948">Ito 1948</bibRefCitation>
: 89;
<bibRefCitation author="Ishida, H" journalOrPublisher="Science Report of the Hyogo University of Agriculture, Serie Natural Sciences" pageId="0" pageNumber="41" pagination="3 - 11" refId="B14" refString="Ishida, H, 1959. The catalogue of the Japanese Tipulidae, with the keys to the genera and subgenera (Diptera). V. Limoniinae, tribe Hexatomini. Science Report of the Hyogo University of Agriculture, Serie Natural Sciences 4 (1): 3 - 11" title="The catalogue of the Japanese Tipulidae, with the keys to the genera and subgenera (Diptera). V. Limoniinae, tribe Hexatomini." volume="4" year="1959">Ishida 1959</bibRefCitation>
: 2;
<bibRefCitation author="Savchenko, EN" journalOrPublisher="Akademiya Nauk Ukrainskoy SSR, Kiev" pageId="0" pageNumber="41" refId="B26" refString="Savchenko, EN, Krivolutskaya, GO, 1976. Limoniidae of the South Kuril Islands and South Sakhalin. Akademiya Nauk Ukrainskoy SSR, Kiev" title="Limoniidae of the South Kuril Islands and South Sakhalin." year="1976">Savchenko and Krivolutskaya 1976</bibRefCitation>
: 74;
<bibRefCitation author="Savchenko, EN" journalOrPublisher="Schmalhausen Institute of Zoology of Academy of Sciences of Ukraine, Naukova Dumka, Kiev" pageId="0" pageNumber="41" refId="B27" refString="Savchenko, EN, 1983. Limoniidae of South Primorye. Akademiy Nauk Ukrainskoy SSR, I.I. Schmalhausen Institute of Zoology of Academy of Sciences of Ukraine, Naukova Dumka, Kiev" title="Limoniidae of South Primorye. Akademiy Nauk Ukrainskoy SSR, I. I." year="1983">Savchenko 1983</bibRefCitation>
: 62,
<bibRefCitation author="Savchenko, EN" journalOrPublisher="Fauna Ukrainy" pageId="0" pageNumber="41" pagination="1 - 380" refId="B28" refString="Savchenko, EN, 1986. Komary-limoniidy [limoniid-flies]. (General description, subfamilies Pediciinae and Hexatominae). Fauna Ukrainy 14 (2): 1 - 380" title="Komary-limoniidy [limoniid-flies]. (General description, subfamilies Pediciinae and Hexatominae)." volume="14" year="1986">1986</bibRefCitation>
: 202,
<bibRefCitation author="Savchenko, EN" journalOrPublisher="Schmalhausen Institute of Zoology of Academy of Sciences of Ukraine, Naukova Dumka, Kiev" pageId="0" pageNumber="41" refId="B29" refString="Savchenko, EN, 1989. Komary-limoniidy fauny SSSR [Limoniidae fauna of the USSR]. Determination tables of superspecies taxa with catalogue survey of species. Akadimiya Nauk Ukrainian SSR, I.I. Schmalhausen Institute of Zoology of Academy of Sciences of Ukraine, Naukova Dumka, Kiev" title="Komary-limoniidy fauny SSSR [Limoniidae fauna of the USSR]. Determination tables of superspecies taxa with catalogue survey of species. Akadimiya Nauk Ukrainian SSR, I. I." year="1989">1989</bibRefCitation>
: 49;
<bibRefCitation DOI="https://doi.org/10.12651/JSR.2015.4.2.061" author="Podenas, S" journalOrPublisher="Journal of Species Research" pageId="0" pageNumber="41" pagination="61 - 96" refId="B24" refString="Podenas, S, Byun, HW, Kim, SK, 2015. Limoniinae crane flies (Diptera: Limoniidae) new to Korea. Journal of Species Research 4: 61 - 96, DOI: https://doi.org/10.12651/JSR.2015.4.2.061" title="Limoniinae crane flies (Diptera: Limoniidae) new to Korea." url="https://doi.org/10.12651/JSR.2015.4.2.061" volume="4" year="2015">Podenas et al. 2015</bibRefCitation>
: 70.
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<paragraph pageId="0" pageNumber="41">Type species.</paragraph>
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<emphasis italics="true" pageId="0" pageNumber="41">Limnobiorhynchus canadensis</emphasis>
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Westwood, 1836 (=
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Osten Sacken, 1869).
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<paragraph pageId="0" pageNumber="41">
A total of 16 species of
<taxonomicName authorityName="Osten Sacken" authorityYear="1860" class="Insecta" family="Limoniidae" genus="Elephantomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elephantomyia" order="Diptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="41">Elephantomyia</emphasis>
</taxonomicName>
are known from the East Palaearctic (
<bibRefCitation author="Oosterbroek, P" journalOrPublisher="Monograph 27 (1). Research Branch, Agriculture Canada, Ottawa" pageId="0" pageNumber="41" publicationUrl="https://ccw.naturalis.nl/index.php" refId="B20" refString="Oosterbroek, P, 2020. Catalogue of the Craneflies of the World (CCW). https://ccw.naturalis.nl/index.php" title="Catalogue of the Craneflies of the World (CCW)." url="https://ccw.naturalis.nl/index.php" year="2020">Oosterbroek 2020</bibRefCitation>
). They belong to two subgenera:
<taxonomicName authority="" lsidName="E. (Elephantomyia)" pageId="0" pageNumber="41" rank="subGenus" subGenus="Elephantomyia">E. (Elephantomyia)</taxonomicName>
Osten Sacken, 1860 and
<taxonomicName authority="" lsidName="E. (Elephantomyodes)" pageId="0" pageNumber="41" rank="subGenus" subGenus="Elephantomyodes">E. (Elephantomyodes)</taxonomicName>
Alexander, 1923. The nominate subgenus includes 14 species, and the subgenus
<taxonomicName lsidName="(Elephantomyodes)" pageId="0" pageNumber="41" rank="subGenus" subGenus="Elephantomyodes">Elephantomyodes</taxonomicName>
includes two species (
<taxonomicName lsidName="E. sophiarum" pageId="0" pageNumber="41" rank="species" species="sophiarum">
<emphasis italics="true" pageId="0" pageNumber="41">E. sophiarum</emphasis>
</taxonomicName>
Ito, 1948 from Honshu and Kyushu, Japan, and
<taxonomicName lsidName="E. tianmushana" pageId="0" pageNumber="41" rank="species" species="tianmushana">
<emphasis italics="true" pageId="0" pageNumber="41">E. tianmushana</emphasis>
</taxonomicName>
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3919.3.6" author="Zhang, X" journalOrPublisher="Zootaxa" pageId="0" pageNumber="41" pagination="553 - 572" refId="B33" refString="Zhang, X, Li, Y, Yang, D, 2015. A review of the genus Elephantomyia Osten Sacken from China, with descriptions of two new species (Diptera, Limoniidae). Zootaxa 3919: 553 - 572, DOI: https://doi.org/10.11646/zootaxa.3919.3.6" title="A review of the genus Elephantomyia Osten Sacken from China, with descriptions of two new species (Diptera, Limoniidae)." url="https://doi.org/10.11646/zootaxa.3919.3.6" volume="3919" year="2015">Zhang et al., 2015</bibRefCitation>
from Zhejiang, China). Eastern Palaearctic species of the subgenus
<taxonomicName authority="" lsidName="E. (Elephantomyodes)" pageId="0" pageNumber="41" rank="subGenus" subGenus="Elephantomyodes">E. (Elephantomyodes)</taxonomicName>
can be easily distinguished from species belonging to the subgenus
<taxonomicName authority="" lsidName="E. (Elephantomyia)" pageId="0" pageNumber="41" rank="subGenus" subGenus="Elephantomyia">E. (Elephantomyia)</taxonomicName>
, as they have snowy white tarsal segments and a very narrow anal angle of the wing. Only one species of the genus
<taxonomicName authorityName="Osten Sacken" authorityYear="1860" class="Insecta" family="Limoniidae" genus="Elephantomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elephantomyia" order="Diptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="41">Elephantomyia</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="E. edwardsi" pageId="0" pageNumber="41" rank="species" species="edwardsi">
<emphasis italics="true" pageId="0" pageNumber="41">E. edwardsi</emphasis>
</taxonomicName>
Lackschewitz, 1932 was previously recorded from the Korean Peninsula (
<bibRefCitation DOI="https://doi.org/10.12651/JSR.2015.4.2.061" author="Podenas, S" journalOrPublisher="Journal of Species Research" pageId="0" pageNumber="41" pagination="61 - 96" refId="B24" refString="Podenas, S, Byun, HW, Kim, SK, 2015. Limoniinae crane flies (Diptera: Limoniidae) new to Korea. Journal of Species Research 4: 61 - 96, DOI: https://doi.org/10.12651/JSR.2015.4.2.061" title="Limoniinae crane flies (Diptera: Limoniidae) new to Korea." url="https://doi.org/10.12651/JSR.2015.4.2.061" volume="4" year="2015">Podenas et al. 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,241 @@
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.913.48650" ID-GBIF-Dataset="81713c0c-9aa1-4a4e-8c5e-ea568c065442" ID-PMC="PMC7044255" ID-Pensoft-Pub="1313-2970-913-1" ID-Pensoft-UUID="2AC899BE84CE53E0B7461D63DA3F0A03" ID-PubMed="32132849" ID-ZooBank="575D3DC1CBF44E99816D0F68F931F927" ModsDocID="1313-2970-913-1" checkinTime="1582137866924" checkinUser="pensoft" docAuthor="Chang, Wan-Jin &amp; Li, Shuqiang" docDate="2020" docId="C25A5D320D8256BAA313FBA2441EEEAA" docLanguage="en" docName="ZooKeys 913: 1-87" docOrigin="ZooKeys 913" docSource="http://dx.doi.org/10.3897/zookeys.913.48650" docTitle="Leclercera shergylaensis Chang &amp; Li 2020, sp. nov." docType="treatment" docUuid="5D098902-6A00-480F-91C4-7706C93EECEE" docUuidSource="ZooBank" docVersion="4" id="2AC899BE84CE53E0B7461D63DA3F0A03" lastPageNumber="1" masterDocId="2AC899BE84CE53E0B7461D63DA3F0A03" masterDocTitle="Thirty-one new species of the spider genus Leclercera from Southeast Asia (Araneae, Psilodercidae)" masterLastPageNumber="87" masterPageNumber="1" pageNumber="1" updateTime="1668168492669" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Thirty-one new species of the spider genus Leclercera from Southeast Asia (Araneae, Psilodercidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chang, Wan-Jin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Shuqiang</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>913</mods:number>
</mods:detail>
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<mods:start>1</mods:start>
<mods:end>87</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.913.48650</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.913.48650</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-913-1</mods:identifier>
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<treatment ID-GBIF-Taxon="162174138" LSID="urn:lsid:zoobank.org:act:5D098902-6A00-480F-91C4-7706C93EECEE" httpUri="http://treatment.plazi.org/id/C25A5D320D8256BAA313FBA2441EEEAA" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="C25A5D32-0D82-56BA-A313-FBA2441EEEAA" authority="Chang &amp; Li, 2020" authorityName="Chang &amp; Li" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Leclercera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leclercera shergylaensis" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="shergylaensis" status="sp. nov.">Leclercera shergylaensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Leclercera shergylaensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure45" httpUri="https://binary.pensoft.net/fig/380987" pageId="0" pageNumber="1">Figs 45</figureCitation>
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Leclercera shergylaensis sp. nov. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: CO = conductor, EM = embolus, LA = laminar apophysis, RA = retrolateral apophyses, SN = spines." figureDoi="10.3897/zookeys.913.48650.figure46" httpUri="https://binary.pensoft.net/fig/380988" pageId="0" pageNumber="1">, 46</figureCitation>
<figureCitation captionStart="Figure 57" captionStartId="F57" captionText="Figure 57. Cheliceral retromargin, posterior view. A Leclercera aniensis sp. nov. B L. renqinensis sp. nov. C L. shergylaensis sp. nov. D L. pulongensis sp. nov. E L. tudao sp. nov. F L. duibaensis sp. nov. G L. jiazhongensis sp. nov. Abbreviations: PT = promargin teeth, RT = retromargin teeth." figureDoi="10.3897/zookeys.913.48650.figure57" httpUri="https://binary.pensoft.net/fig/380999" pageId="0" pageNumber="1">, 57C</figureCitation>
<figureCitation captionStart="Figure 58" captionStartId="F58" captionText="Figure 58. Distribution of new Leclercera species in Southeast Asia. 1 L. selasihensis sp. nov. 2 L. mianqiu sp. nov. 3 L. thamsangensis sp. nov. 4 L. yandou sp. nov. 5 L. thamkaewensis sp. nov. 6 L. xiangbabang sp. nov. 7 L. jianzuiyu sp. nov. 8 L. yamaensis sp. nov. 9 L. banensis sp. nov. 10 L. dumuzhou sp. nov. 11 L. suwanensis sp. nov. 12 L. maochong sp. nov. 13 L. yuanzhui sp. nov. 14 L. shanzi sp. nov. 15 L. duandai sp. nov. 16 L. hponensis sp. nov. 17 L. lizi sp. nov. 18 L. xiaodai sp. nov. 19 L. paiensis sp. nov. 20 L. yanjing sp. nov. 21 L. ekteenensis sp. nov. 22 L. zanggaensis sp. nov. 23 L. zhamensis sp. nov. 24 L. sanjiao sp. nov. 25 L. aniensis sp. nov. 26 L. renqinensis sp. nov. 27 L. shergylaensis sp. nov. 28 L. pulongensis sp. nov. 29 L. tudao sp. nov. 30 L. duibaensis sp. nov. 31 L. jiazhongensis sp. nov." figureDoi="10.3897/zookeys.913.48650.figure58" httpUri="https://binary.pensoft.net/fig/381000" pageId="0" pageNumber="1">, 58</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="types">
<paragraph pageId="0" pageNumber="1">Types.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: ♂ (IZCAS), China, Tibet Autonomous Region, Nyingchi, Shergyla Mountain,
<geoCoordinate degrees="29" direction="north" minutes="33.7980" orientation="latitude" precision="1" value="29.5633">29°33.7980'N</geoCoordinate>
,
<geoCoordinate degrees="94" direction="east" minutes="34.2060" orientation="longitude" precision="1" value="94.5701">94°34.2060'E</geoCoordinate>
, elevation ca 3764 m, 15.VII.2019, X. Zhang, Z. Bai and J. Liu leg.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
: 1♀ (IZCAS), same data as holotype.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">The species name is an adjective referring to the type locality.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Males of
<taxonomicName lsidName="L. shergylaensis" pageId="0" pageNumber="1" rank="species" species="shergylaensis">
<emphasis italics="true" pageId="0" pageNumber="1">L. shergylaensis</emphasis>
</taxonomicName>
sp. nov. resemble
<taxonomicName lsidName="L. pulongensis" pageId="0" pageNumber="1" rank="species" species="pulongensis">
<emphasis italics="true" pageId="0" pageNumber="1">L. pulongensis</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName lsidName="L. duibaensis" pageId="0" pageNumber="1" rank="species" species="duibaensis">
<emphasis italics="true" pageId="0" pageNumber="1">L. duibaensis</emphasis>
</taxonomicName>
sp. nov. but can be distinguished by the presence of a conductor (Fig.
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Leclercera shergylaensis sp. nov. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: CO = conductor, EM = embolus, LA = laminar apophysis, RA = retrolateral apophyses, SN = spines." figureDoi="10.3897/zookeys.913.48650.figure46" httpUri="https://binary.pensoft.net/fig/380988" pageId="0" pageNumber="1">46B</figureCitation>
) (vs. absence of a conductor in
<taxonomicName lsidName="L. pulongensis" pageId="0" pageNumber="1" rank="species" species="pulongensis">
<emphasis italics="true" pageId="0" pageNumber="1">L. pulongensis</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Leclercera pulongensis sp. nov. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: EM = embolus, LA = laminar apophysis, RA = retrolateral apophysis, SN = spine, ST = strong setae." figureDoi="10.3897/zookeys.913.48650.figure48" httpUri="https://binary.pensoft.net/fig/380990" pageId="0" pageNumber="1">48B</figureCitation>
) and
<taxonomicName lsidName="L. duibaensis" pageId="0" pageNumber="1" rank="species" species="duibaensis">
<emphasis italics="true" pageId="0" pageNumber="1">L. duibaensis</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Leclercera duibaensis sp. nov. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: BG = bulge, EM = embolus, LA = laminar apophysis, RA = retrolateral apophyses, SN = spines." figureDoi="10.3897/zookeys.913.48650.figure52" httpUri="https://binary.pensoft.net/fig/380994" pageId="0" pageNumber="1">52B</figureCitation>
)), two retrolateral apophyses bearing a spine on the tibia (Fig.
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Leclercera shergylaensis sp. nov. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: CO = conductor, EM = embolus, LA = laminar apophysis, RA = retrolateral apophyses, SN = spines." figureDoi="10.3897/zookeys.913.48650.figure46" httpUri="https://binary.pensoft.net/fig/380988" pageId="0" pageNumber="1">46D</figureCitation>
) (vs. a prolateral apophysis bearing a hooked spine on the tibia in
<taxonomicName lsidName="L. pulongensis" pageId="0" pageNumber="1" rank="species" species="pulongensis">
<emphasis italics="true" pageId="0" pageNumber="1">L. pulongensis</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Leclercera pulongensis sp. nov. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: EM = embolus, LA = laminar apophysis, RA = retrolateral apophysis, SN = spine, ST = strong setae." figureDoi="10.3897/zookeys.913.48650.figure48" httpUri="https://binary.pensoft.net/fig/380990" pageId="0" pageNumber="1">48C</figureCitation>
)) and a retrolateral apophysis bearing spine on tibia in
<taxonomicName lsidName="L. duibaensis" pageId="0" pageNumber="1" rank="species" species="duibaensis">
<emphasis italics="true" pageId="0" pageNumber="1">L. duibaensis</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Leclercera duibaensis sp. nov. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: BG = bulge, EM = embolus, LA = laminar apophysis, RA = retrolateral apophyses, SN = spines." figureDoi="10.3897/zookeys.913.48650.figure52" httpUri="https://binary.pensoft.net/fig/380994" pageId="0" pageNumber="1">52D</figureCitation>
)), absence of strong setae in both
<taxonomicName lsidName="L. shergylaensis" pageId="0" pageNumber="1" rank="species" species="shergylaensis">
<emphasis italics="true" pageId="0" pageNumber="1">L. shergylaensis</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName lsidName="L. duibaensis" pageId="0" pageNumber="1" rank="species" species="duibaensis">
<emphasis italics="true" pageId="0" pageNumber="1">L. duibaensis</emphasis>
</taxonomicName>
sp. nov. (vs. the presence of strong setae on the tibia anteriorly in
<taxonomicName lsidName="L. pulongensis" pageId="0" pageNumber="1" rank="species" species="pulongensis">
<emphasis italics="true" pageId="0" pageNumber="1">L. pulongensis</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Leclercera pulongensis sp. nov. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: EM = embolus, LA = laminar apophysis, RA = retrolateral apophysis, SN = spine, ST = strong setae." figureDoi="10.3897/zookeys.913.48650.figure48" httpUri="https://binary.pensoft.net/fig/380990" pageId="0" pageNumber="1">48D</figureCitation>
)), and the absence of a bulge on the bulb of both
<taxonomicName lsidName="L. shergylaensis" pageId="0" pageNumber="1" rank="species" species="shergylaensis">
<emphasis italics="true" pageId="0" pageNumber="1">L. shergylaensis</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName lsidName="L. pulongensis" pageId="0" pageNumber="1" rank="species" species="pulongensis">
<emphasis italics="true" pageId="0" pageNumber="1">L. pulongensis</emphasis>
</taxonomicName>
sp. nov. (vs. the presence of a bulge on the bulb of
<taxonomicName lsidName="L. duibaensis" pageId="0" pageNumber="1" rank="species" species="duibaensis">
<emphasis italics="true" pageId="0" pageNumber="1">L. duibaensis</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Leclercera duibaensis sp. nov. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: BG = bulge, EM = embolus, LA = laminar apophysis, RA = retrolateral apophyses, SN = spines." figureDoi="10.3897/zookeys.913.48650.figure52" httpUri="https://binary.pensoft.net/fig/380994" pageId="0" pageNumber="1">52B</figureCitation>
)), absence of an apophysis on the cymbium in both
<taxonomicName lsidName="L. shergylaensis" pageId="0" pageNumber="1" rank="species" species="shergylaensis">
<emphasis italics="true" pageId="0" pageNumber="1">L. shergylaensis</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName lsidName="L. pulongensis" pageId="0" pageNumber="1" rank="species" species="pulongensis">
<emphasis italics="true" pageId="0" pageNumber="1">L. pulongensis</emphasis>
</taxonomicName>
sp. nov. (vs. the presence of a retrolateral apophysis bearing four spines on the cymbium in
<taxonomicName lsidName="L. duibaensis" pageId="0" pageNumber="1" rank="species" species="duibaensis">
<emphasis italics="true" pageId="0" pageNumber="1">L. duibaensis</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Leclercera duibaensis sp. nov. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: BG = bulge, EM = embolus, LA = laminar apophysis, RA = retrolateral apophyses, SN = spines." figureDoi="10.3897/zookeys.913.48650.figure52" httpUri="https://binary.pensoft.net/fig/380994" pageId="0" pageNumber="1">52D</figureCitation>
)), and the tibia swollen angularly (Fig.
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Leclercera shergylaensis sp. nov. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: CO = conductor, EM = embolus, LA = laminar apophysis, RA = retrolateral apophyses, SN = spines." figureDoi="10.3897/zookeys.913.48650.figure46" httpUri="https://binary.pensoft.net/fig/380988" pageId="0" pageNumber="1">46C</figureCitation>
) (vs. tibia roundly swollen in both
<taxonomicName lsidName="L. pulongensis" pageId="0" pageNumber="1" rank="species" species="pulongensis">
<emphasis italics="true" pageId="0" pageNumber="1">L. pulongensis</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName lsidName="L. duibaensis" pageId="0" pageNumber="1" rank="species" species="duibaensis">
<emphasis italics="true" pageId="0" pageNumber="1">L. duibaensis</emphasis>
</taxonomicName>
sp. nov.).
</paragraph>
<caption doi="10.3897/zookeys.913.48650.figure45" httpUri="https://binary.pensoft.net/fig/380987" pageId="0" pageNumber="1" start="Figure 45" startId="F45">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 45.</emphasis>
<taxonomicName authorityName="Chang &amp; Li" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Leclercera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leclercera shergylaensis" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="shergylaensis">
<emphasis italics="true" pageId="0" pageNumber="1">Leclercera shergylaensis</emphasis>
</taxonomicName>
sp. nov., male holotype and female paratype.
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
Endogyne, dorsal view
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
female epigastric area, ventral view
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
male habitus, dorsal view
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
female habitus, dorsal view
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
female habitus, ventral view. Abbreviation: SP = spermatheca.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Male</emphasis>
(Holotype). Total length 3.13; carapace 1.00 long, 1.13 wide; abdomen 2.13 long, 1.09 wide. Carapace round and brown, with median dark brown band (Fig.
<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Leclercera shergylaensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure45" httpUri="https://binary.pensoft.net/fig/380987" pageId="0" pageNumber="1">45C</figureCitation>
). Chelicerae brown (Fig.
<figureCitation captionStart="Figure 57" captionStartId="F57" captionText="Figure 57. Cheliceral retromargin, posterior view. A Leclercera aniensis sp. nov. B L. renqinensis sp. nov. C L. shergylaensis sp. nov. D L. pulongensis sp. nov. E L. tudao sp. nov. F L. duibaensis sp. nov. G L. jiazhongensis sp. nov. Abbreviations: PT = promargin teeth, RT = retromargin teeth." figureDoi="10.3897/zookeys.913.48650.figure57" httpUri="https://binary.pensoft.net/fig/380999" pageId="0" pageNumber="1">57C</figureCitation>
). Clypeus pale brown. Endites dark brown, light brown basally. Labium dark brown. Sternum dark brown, with short light brown band medially. Abdomen elongated, dorsum with dark brown spots, posterior with dark brown horizontal stripes, antero-ventrally with dark brown inverted fan-shaped pattern medially and kidney-shaped pattern laterally, posterior dark brown with two light vertical traces medially. Legs uniformly brown; measurements: I 9.60 (2.60. 0.40. 3.00, 2.40, 1.20), II 8.94 (2.60, 0.40, 2.50, 2.24, 1.20), III missing, IV 9.20 (2.60, 0.40, 2.80, 2.20, 1.20). Palp (Fig.
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Leclercera shergylaensis sp. nov. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: CO = conductor, EM = embolus, LA = laminar apophysis, RA = retrolateral apophyses, SN = spines." figureDoi="10.3897/zookeys.913.48650.figure46" httpUri="https://binary.pensoft.net/fig/380988" pageId="0" pageNumber="1">46A-D</figureCitation>
): femur slender, four times longer than patella; patella not swollen; tibia swollen pentagonally, similar length to, and 2 times wider than femur, with two retrolateral apophyses bearing spine (Fig.
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Leclercera shergylaensis sp. nov. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: CO = conductor, EM = embolus, LA = laminar apophysis, RA = retrolateral apophyses, SN = spines." figureDoi="10.3897/zookeys.913.48650.figure46" httpUri="https://binary.pensoft.net/fig/380988" pageId="0" pageNumber="1">46D</figureCitation>
) (one apophysis on anterior, the other medially on tibia), spine and apophysis similar in length; cymbium two times shorter than femur, dark brown distally; bulb brown, ovate, with embolus, laminar apophysis, and conductor arising distally; embolus thin, two times longer than conductor and laminar apophysis; branched laminar apophysis, two times wider than embolus, adjacent to conductor; conductor branched, with the longer branch longer than laminar apophysis, attached to embolus (Fig.
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Leclercera shergylaensis sp. nov. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: CO = conductor, EM = embolus, LA = laminar apophysis, RA = retrolateral apophyses, SN = spines." figureDoi="10.3897/zookeys.913.48650.figure46" httpUri="https://binary.pensoft.net/fig/380988" pageId="0" pageNumber="1">46B</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Female</emphasis>
(Paratype). General features and coloration similar to those of male (Fig.
<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Leclercera shergylaensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure45" httpUri="https://binary.pensoft.net/fig/380987" pageId="0" pageNumber="1">45D, E</figureCitation>
). Measurements: total length 2.44; carapace 0.94 long, 1.00 wide; abdomen 1.50 long, 0.88 wide. Leg measurements: I 7.69 (2.24, 0.31, 2.34, 1.80, 1.00), II 6.70 (1.80, 0.31, 2.03, 1.56, 1.00), III 5.29 (1.41, 0.31, 1.38, 1.25, 0.94), IV 6.10 (1.80, 0.31, 2.19, 1.80, 1.00). Epigastric area (Fig.
<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Leclercera shergylaensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure45" httpUri="https://binary.pensoft.net/fig/380987" pageId="0" pageNumber="1">45B</figureCitation>
): elliptical dark brown patch with distinct whitish slit. Endogyne (Fig.
<figureCitation captionStart="Figure 45" captionStartId="F45" captionText="Figure 45. Leclercera shergylaensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure45" httpUri="https://binary.pensoft.net/fig/380987" pageId="0" pageNumber="1">45A</figureCitation>
): a pair of ovoid spermathecae, thickening at margins.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Known only from the type locality (Fig.
<figureCitation captionStart="Figure 58" captionStartId="F58" captionText="Figure 58. Distribution of new Leclercera species in Southeast Asia. 1 L. selasihensis sp. nov. 2 L. mianqiu sp. nov. 3 L. thamsangensis sp. nov. 4 L. yandou sp. nov. 5 L. thamkaewensis sp. nov. 6 L. xiangbabang sp. nov. 7 L. jianzuiyu sp. nov. 8 L. yamaensis sp. nov. 9 L. banensis sp. nov. 10 L. dumuzhou sp. nov. 11 L. suwanensis sp. nov. 12 L. maochong sp. nov. 13 L. yuanzhui sp. nov. 14 L. shanzi sp. nov. 15 L. duandai sp. nov. 16 L. hponensis sp. nov. 17 L. lizi sp. nov. 18 L. xiaodai sp. nov. 19 L. paiensis sp. nov. 20 L. yanjing sp. nov. 21 L. ekteenensis sp. nov. 22 L. zanggaensis sp. nov. 23 L. zhamensis sp. nov. 24 L. sanjiao sp. nov. 25 L. aniensis sp. nov. 26 L. renqinensis sp. nov. 27 L. shergylaensis sp. nov. 28 L. pulongensis sp. nov. 29 L. tudao sp. nov. 30 L. duibaensis sp. nov. 31 L. jiazhongensis sp. nov." figureDoi="10.3897/zookeys.913.48650.figure58" httpUri="https://binary.pensoft.net/fig/381000" pageId="0" pageNumber="1">58</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.913.48650.figure46" httpUri="https://binary.pensoft.net/fig/380988" pageId="0" pageNumber="1" start="Figure 46" startId="F46">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 46.</emphasis>
<taxonomicName authorityName="Chang &amp; Li" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Leclercera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leclercera shergylaensis" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="shergylaensis">
<emphasis italics="true" pageId="0" pageNumber="1">Leclercera shergylaensis</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
Palp, ventral view
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
bulb, ventral view
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
palp, prolateral view
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
palp, retrolateral view. Abbreviations: CO = conductor, EM = embolus, LA = laminar apophysis, RA = retrolateral apophyses, SN = spines.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

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<mods:affiliation>Cornell University Insect Collection, John H. and Anna B. Comstock Hall, 129 Garden Ave., Cornell University, Ithaca NY 14853 - 2601 U. S. A.</mods:affiliation>
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<taxonomicName authority="Valentine, 1987" authorityName="Valentine" authorityYear="1987" class="Insecta" family="Carabidae" genus="Bryanites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryanites" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Bryanites Valentine, 1987</taxonomicName>
</paragraph>
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<subSubSection pageId="2" pageNumber="3" type="type species">
<paragraph pageId="2" pageNumber="3">Type species.</paragraph>
<paragraph pageId="2" pageNumber="3">
<taxonomicName authorityName="Valentine" authorityYear="1987" class="Insecta" family="Carabidae" genus="Bryanites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryanites samoaensis" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="samoaensis">
<emphasis italics="true" pageId="2" pageNumber="3">Bryanites samoaensis</emphasis>
</taxonomicName>
Valentine (by original designation).
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="diagnosis">
<paragraph pageId="2" pageNumber="3">Diagnosis.</paragraph>
<paragraph pageId="2" pageNumber="3">
These beetles can be diagnosed from other Pacific platynine taxa by the much reduced subapical sinuation of the elytra, with the elytral margins straight to convex near the apex (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Male holotype, Bryanites graeffii, dorsal view." figureDoi="10.3897/zse.93.10802.figure1" httpUri="https://binary.pensoft.net/fig/116946" pageId="2" pageNumber="3">1</figureCitation>
). The pronotum is broadly orbiculate, with the lateral margins convex to only slightly concave anterad the rounded hind angles. The pronotum has the basal seta present and situated anterad the rounded hind angle. The elytra are moderately narrowed basally, with the basal elytral groove meeting the lateral elytral depression at an angle; i.e. the humeri are angulate. The male aedeagus exhibits characters typical of tribe
<taxonomicName authorityName="Bonelli" authorityYear="1810" lsidName="" pageId="2" pageNumber="3" rank="tribe" tribe="Platynini">Platynini</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male aedeagus, internal sac everted, of Bryanites graeffii male holotype, right lateral view. Abbreviations include: ae, aedeagal median lobe; is, internal sac; lp, left paramere; rp, right paramere." figureDoi="10.3897/zse.93.10802.figure4" httpUri="https://binary.pensoft.net/fig/116951" pageId="2" pageNumber="3">4</figureCitation>
): 1, both parameres are rounded apically, or conchoid in shape; 2, the right paramere is smaller than the left, and the aedeagus lies with the right side ventral in repose; 3, the basal bulb is closed and bears a sagittal crest; and 4, the internal sac is tubular and unarmored, with slightly more melanized spicules present apically near the gonopore.
</paragraph>
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<paragraph pageId="2" pageNumber="3">
<emphasis bold="true" pageId="2" pageNumber="3">Figure 4.</emphasis>
Male aedeagus, internal sac everted, of
<taxonomicName authorityName="Liebherr" authorityYear="2017" class="Insecta" family="Carabidae" genus="Bryanites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryanites graeffii" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="graeffii">
<emphasis italics="true" pageId="2" pageNumber="3">Bryanites graeffii</emphasis>
</taxonomicName>
male holotype, right lateral view. Abbreviations include: ae, aedeagal median lobe; is, internal sac; lp, left paramere; rp, right paramere.
</paragraph>
</caption>
<paragraph pageId="2" pageNumber="3">
Beetles of all three
<taxonomicName authorityName="Valentine" authorityYear="1987" class="Insecta" family="Carabidae" genus="Bryanites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryanites" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="2" pageNumber="3">Bryanites</emphasis>
</taxonomicName>
species are brachypterous, with the wing rudiments narrow and elongate stenopterous straps, the length of the rudiments more than 4
<normalizedToken originalValue="×">x</normalizedToken>
the rudiment breadth. Beetle size is moderate to large, ranging from 11.7 mm in
<taxonomicName authorityName="Valentine" authorityYear="1987" class="Insecta" family="Carabidae" genus="Bryanites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryanites barri" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="barri">
<emphasis italics="true" pageId="2" pageNumber="3">Bryanites barri</emphasis>
</taxonomicName>
(measurement of Barr 1987) to 16.2 mm in
<taxonomicName authorityName="Liebherr" authorityYear="2017" class="Insecta" family="Carabidae" genus="Bryanites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryanites graeffii" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="graeffii">
<emphasis italics="true" pageId="2" pageNumber="3">Bryanites graeffii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="key">
<paragraph pageId="2" pageNumber="3">
Key to the Adults of
<taxonomicName authorityName="Valentine" authorityYear="1987" class="Insecta" family="Carabidae" genus="Bryanites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryanites" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="2" pageNumber="3">Bryanites</emphasis>
</taxonomicName>
Valentine
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<paragraph pageId="2" pageNumber="3">
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<td colspan="1" pageId="2" pageNumber="3" rowspan="1">1.</td>
<td colspan="1" pageId="2" pageNumber="3" rowspan="1">Beetles of moderate size, standardized body length 11.7-12.5 mm; pronotum quadrisetose both lateral and basal setae present; parascutellar seta present</td>
<td colspan="1" pageId="2" pageNumber="3" rowspan="1">2</td>
</tr>
<tr pageId="2" pageNumber="3">
<td colspan="1" pageId="2" pageNumber="3" rowspan="1">-</td>
<td colspan="1" pageId="2" pageNumber="3" rowspan="1">Larger beetles, standardized body length 16.2 mm; pronotum bisetose, only basal seta present both sides; parascutellar seta absent</td>
<td colspan="1" pageId="2" pageNumber="3" rowspan="1">
<taxonomicName authorityName="Liebherr" authorityYear="2017" class="Insecta" family="Carabidae" genus="Bryanites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryanites graeffii" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="graeffii">
<emphasis italics="true" pageId="2" pageNumber="3">Bryanites graeffii</emphasis>
</taxonomicName>
sp. n.
</td>
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<tr pageId="2" pageNumber="3">
<td colspan="1" pageId="2" pageNumber="3" rowspan="1">2.</td>
<td colspan="1" pageId="2" pageNumber="3" rowspan="1">Third elytral interval with 6 setae along length, the anterior seta associated with the third stria, the trailing five setae associated with the second stria; elytral basal groove meeting lateral elytral depression at right angle; pronotum without any evident laterobasal depressions, at most a slight depression near hind margin (left side of holotype)</td>
<td colspan="1" pageId="2" pageNumber="3" rowspan="1">
<taxonomicName authorityName="Valentine" authorityYear="1987" class="Insecta" family="Carabidae" genus="Bryanites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryanites samoaensis" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="samoaensis">
<emphasis italics="true" pageId="2" pageNumber="3">Bryanites samoaensis</emphasis>
</taxonomicName>
Valentine
</td>
</tr>
<tr pageId="2" pageNumber="3">
<td colspan="1" pageId="2" pageNumber="3" rowspan="1">-</td>
<td colspan="1" pageId="2" pageNumber="3" rowspan="1">Third elytral interval with 3 setae along length, the anterior seta associated with the third stria, the posterior two setae associated with the second stria; elytral basal groove meeting lateral elytral depression at acute angle; pronotum with broad, shallow linear laterobasal depressions extended from hind angles toward center of pronotal disc</td>
<td colspan="1" pageId="2" pageNumber="3" rowspan="1">
<taxonomicName authorityName="Valentine" authorityYear="1987" class="Insecta" family="Carabidae" genus="Bryanites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryanites barri" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="barri">
<emphasis italics="true" pageId="2" pageNumber="3">Bryanites barri</emphasis>
</taxonomicName>
Valentine
</td>
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</paragraph>
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Genus
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(
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species:
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<mods:title id="031EAB6B07490450645A2C869AA4447A">A new fossil species of the reticulated beetle genus Cupes (Coleoptera: Archostemata: Cupedidae) from Eocene Baltic amber</mods:title>
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<heading id="11048F68FFE7FFA12CFB998F9BD0E2DD" bold="true" box="[151,402,1471,1498]" fontSize="11" level="1" pageId="1" pageNumber="452" reason="1">
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<figureCitation id="D2C82481FFE7FFA12CFB99AB9B41E2F9" box="[151,259,1508,1534]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,250,1930,1954]" captionStartId-1="4.[151,250,1854,1878]" captionStartId-2="5.[151,250,1142,1166]" captionStartId-3="6.[151,250,1909,1933]" captionTargetBox-0="[155,1436,179,1904]" captionTargetBox-1="[164,1423,181,1830]" captionTargetBox-2="[164,1423,181,1118]" captionTargetBox-3="[176,1412,181,1885]" captionTargetId-0="figure-15@3.[151,1436,179,1904]" captionTargetId-1="figure-28@4.[164,1423,181,1830]" captionTargetId-2="figure-15@5.[164,1423,181,1118]" captionTargetId-3="figure-28@6.[176,1412,181,1885]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. General habitus of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Dorsal view. B, Ventral view. Scale bars: 3.0 mm." captionText-1="FIGURE 2. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Head and pronotum, dorsal view. B, Head, dorsal view. C, Right lateral half of pronotum, dorsal view. D, Left elytron and scutellum, dorsal view. E, Right metatarsus, dorsal view. F, Left antenna, dorsal view. Abbreviations: a111, antennomeres 111; ey, eyes; fv, longitudinal median furrow on vertex; mtt15, metatarsomeres 15; p1, supraantennal tubercle; p2, supraocular tubercle; tmp, temple. Scale bars: 1.0 mm (A); 0.5 mm (B, C, E); 2.0 mm (D, F)." captionText-2="FIGURE 3. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Head with morphological interpretations, dorsal view. B, Head without morphological interpretations, dorsal view. C, Pronotum with morphological interpretations, dorsal view. C, Pronotum without morphological interpretations, dorsal view.Abbreviations: a12, antennomeres 12; ey, eyes; fs, paramedian fossae at anterior pronotal edge; fv, longitudinal median furrow on vertex; mrp, median elevated ridge of pronotum; p1, supraantennal tubercle; p2, supraocular tubercle; pn, pronotum; tmp, temple." captionText-3="FIGURE 4. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237.A, Head, right antenna, andprothorax, ventral view. B, Forelegs, ventral view. C, Basal part of left antenna, ventral view. D, Mid- and hindlegs, ventral view. E, Abdomen, ventral view. Abbreviations: a36, antennomeres 36; v15, ventrites 15. Scale bars: 1.0 mm (AD); 2.0 mm (E)." figureDoi-0="http://doi.org/10.5281/zenodo.10910593" figureDoi-1="http://doi.org/10.5281/zenodo.10910595" figureDoi-2="http://doi.org/10.5281/zenodo.10910599" figureDoi-3="http://doi.org/10.5281/zenodo.10910601" httpUri-0="https://zenodo.org/record/10910593/files/figure.png" httpUri-1="https://zenodo.org/record/10910595/files/figure.png" httpUri-2="https://zenodo.org/record/10910599/files/figure.png" httpUri-3="https://zenodo.org/record/10910601/files/figure.png" pageId="1" pageNumber="452">Figs. 14</figureCitation>
.
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<emphasis id="7887E416FFE7FFA12CFB9A639B0AE141" bold="true" box="[151,328,1580,1606]" pageId="1" pageNumber="452">Type material.</emphasis>
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amber, with the accession number SEHU-0000121237, deposited at the
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(
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(
<collectionCode id="2CE2A0C1FFE7FFA12D649AD79B0DE1B5" box="[264,335,1688,1714]" country="Germany" httpUri="http://grbio.org/cool/kch5-hv72" name="Humboldt University Zoologisches Museum" pageId="1" pageNumber="452">HUM</collectionCode>
),
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<location id="4F2C6EDFFFE7FFA12DB49AD7980DE1B5" LSID="urn:lsid:plazi:treatment:C25A8912FFE7FFA72CFB998F98E7E582:4F2C6EDFFFE7FFA12DB49AD7980DE1B5" box="[472,591,1688,1714]" country="Japan" county="Syninclusion" municipality="Sapporo" name="University" pageId="1" pageNumber="452" stateProvince="Hokkaido">University</location>
,
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,
<collectingCountry id="32E47894FFE7FFA12EA99AD6994AE1B4" box="[709,776,1689,1715]" name="Japan" pageId="1" pageNumber="452">Japan</collectingCountry>
.
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: a tiny Collembola.
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<typeStatus id="954886A6FFE7FFA12CAB9AF39B40E1D1" box="[199,258,1724,1750]" pageId="1" pageNumber="452">Type</typeStatus>
locality and horizon.
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<collectingCountry id="32E47894FFE7FFA12E699AF39816E1D1" box="[517,596,1724,1750]" name="Russia" pageId="1" pageNumber="452">Russia</collectingCountry>
:
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; middle to
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.
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<subSubSection id="02E96B8FFFE7FFA12CAB9AAF9B7FE045" pageId="1" pageNumber="452" type="etymology">
<paragraph id="4A4C3804FFE7FFA12CAB9AAF9B7FE045" blockId="1.[151,1437,1580,2039]" pageId="1" pageNumber="452">
<emphasis id="7887E416FFE7FFA12CAB9AAF9B0CE1FD" bold="true" box="[199,334,1760,1786]" pageId="1" pageNumber="452">Etymology.</emphasis>
The specific epithet
<taxonomicName id="8DF34387FFE7FFA12E2F9AAF98DEE1FD" authorityName="Yamamoto" authorityYear="2024" box="[579,668,1760,1786]" class="Insecta" family="Cupedidae" genus="Cupes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="balticus">
<emphasis id="7887E416FFE7FFA12E2F9AAF98DEE1FD" box="[579,668,1760,1786]" italics="true" pageId="1" pageNumber="452">balticus</emphasis>
</taxonomicName>
refers to the occurrence of the
<typeStatus id="954886A6FFE7FFA128679AAE9E79E1FC" box="[1035,1083,1761,1787]" pageId="1" pageNumber="452">type</typeStatus>
material in Baltic amber from near the Baltic Sea Coast, also emphasizing the potentially higher diversity and abundance of
<taxonomicName id="8DF34387FFE7FFA128D99B4B9EB9E019" authorityName="Fabricius" authorityYear="1801" box="[1205,1275,1796,1822]" class="Insecta" family="Cupedidae" genus="Cupes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="452" phylum="Arthropoda" rank="genus">
<emphasis id="7887E416FFE7FFA128D99B4B9EB9E019" box="[1205,1275,1796,1822]" italics="true" pageId="1" pageNumber="452">Cupes</emphasis>
</taxonomicName>
beetles in this amber deposit.
</paragraph>
</subSubSection>
<subSubSection id="02E96B8FFFE7FFA22CAB9B0398C0E7F0" lastPageId="2" lastPageNumber="453" pageId="1" pageNumber="452" type="diagnosis">
<paragraph id="4A4C3804FFE7FFA22CAB9B0398C0E7F0" blockId="1.[151,1437,1580,2039]" lastBlockId="2.[151,1437,151,1981]" lastPageId="2" lastPageNumber="453" pageId="1" pageNumber="452">
<emphasis id="7887E416FFE7FFA12CAB9B039B00E061" bold="true" box="[199,322,1868,1894]" pageId="1" pageNumber="452">Diagnosis.</emphasis>
Body of relatively large size (
<quantity id="8D0B95E1FFE7FFA12EF29B0398B5E060" box="[670,759,1868,1895]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="1" pageNumber="452" unit="mm" value="9.3">9.3 mm</quantity>
long) as
<taxonomicName id="8DF34387FFE7FFA12F089B039E41E061" box="[868,1027,1868,1894]" pageId="1" pageNumber="452">
<emphasis id="7887E416FFE7FFA12F089B0399E8E061" box="[868,938,1868,1894]" italics="true" pageId="1" pageNumber="452">Cupes</emphasis>
species
</taxonomicName>
in Baltic amber, with a peculiar bi-colored pattern on elytra by scales, consisting of brownish and yellowish grey spots (head, median longitudinal stripe on pronotum, and elytral apex brownish); head with 2 pairs of dorsal paramedian tubercles well raised, with temples only slightly shorter than eyes; antenna with antennomere 3 almost twice as long as antennomere 2, as well as moderately narrowly elongate terminal antennomere (about 3.8 times as long as thick); pronotum wider than head, anterolaterally projecting and laterally broadly angulate, with an anteriorly widened medial elevated stripe without a clear longitudinal furrow; hind leg with comaratively short metatarsomere 5 (metatarsomere 1 distinctly less than twice as long as metatarsomere 5).
</paragraph>
</subSubSection>
<subSubSection id="02E96B8FFFE4FFA72CAB9CB198E7E582" lastPageId="7" lastPageNumber="458" pageId="2" pageNumber="453" type="description">
<paragraph id="4A4C3804FFE4FFA22CAB9CB199B2E6C0" blockId="2.[151,1437,151,1981]" pageId="2" pageNumber="453">
<emphasis id="7887E416FFE4FFA22CAB9CB19B1BE61F" bold="true" box="[199,345,254,280]" pageId="2" pageNumber="453">Description.</emphasis>
Body (
<figureCitation id="D2C82481FFE4FFA22DDE9CB09BB7E61E" box="[434,501,255,281]" captionStart="FIGURE 1" captionStartId="3.[151,250,1930,1954]" captionTargetBox="[155,1436,179,1904]" captionTargetId="figure-15@3.[151,1436,179,1904]" captionTargetPageId="3" captionText="FIGURE 1. General habitus of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Dorsal view. B, Ventral view. Scale bars: 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10910593" httpUri="https://zenodo.org/record/10910593/files/figure.png" pageId="2" pageNumber="453">Fig. 1</figureCitation>
) narrowly elongate, relatively large,
<quantity id="8D0B95E1FFE4FFA22FCA9CB09E4FE61E" box="[934,1037,255,281]" metricMagnitude="-3" metricUnit="m" metricValue="9.32" pageId="2" pageNumber="453" unit="mm" value="9.32">9.32 mm</quantity>
long, moderately convex dorsally, flattened ventrally, widest at posterior part of elytra. Surface with extremely dense scales, masking coloration of body surface; scales on head, pronotum (including pronotal median longitudinal stripe) dark brown; elytra bi-colored, with a characteristic pattern of brownish and yellowish grey spots, with brownish apex (both subapical and apical bands without interruption); body ventrally with extremely dense, yellowish grey scales; surface on antennae and legs scales gradually transforming into subrecumbent, rather stout setae.
</paragraph>
<paragraph id="4A4C3804FFE4FFA22CAB9D809F3AE48E" blockId="2.[151,1437,151,1981]" pageId="2" pageNumber="453">
Head (
<figureCitation id="D2C82481FFE4FFA22D7C9D809BD3E6EE" box="[272,401,463,489]" captionStart="FIGURE 2" captionStartId="4.[151,250,1854,1878]" captionTargetBox="[164,1423,181,1830]" captionTargetId="figure-28@4.[164,1423,181,1830]" captionTargetPageId="4" captionText="FIGURE 2. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Head and pronotum, dorsal view. B, Head, dorsal view. C, Right lateral half of pronotum, dorsal view. D, Left elytron and scutellum, dorsal view. E, Right metatarsus, dorsal view. F, Left antenna, dorsal view. Abbreviations: a111, antennomeres 111; ey, eyes; fv, longitudinal median furrow on vertex; mtt15, metatarsomeres 15; p1, supraantennal tubercle; p2, supraocular tubercle; tmp, temple. Scale bars: 1.0 mm (A); 0.5 mm (B, C, E); 2.0 mm (D, F)." figureDoi="http://doi.org/10.5281/zenodo.10910595" httpUri="https://zenodo.org/record/10910595/files/figure.png" pageId="2" pageNumber="453">Figs. 2A, B</figureCitation>
,
<figureCitation id="D2C82481FFE4FFA22DF09D809B9FE6EE" box="[412,477,463,489]" captionStart="FIGURE 3" captionStartId="5.[151,250,1142,1166]" captionTargetBox="[164,1423,181,1118]" captionTargetId="figure-15@5.[164,1423,181,1118]" captionTargetPageId="5" captionText="FIGURE 3. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Head with morphological interpretations, dorsal view. B, Head without morphological interpretations, dorsal view. C, Pronotum with morphological interpretations, dorsal view. C, Pronotum without morphological interpretations, dorsal view.Abbreviations: a12, antennomeres 12; ey, eyes; fs, paramedian fossae at anterior pronotal edge; fv, longitudinal median furrow on vertex; mrp, median elevated ridge of pronotum; p1, supraantennal tubercle; p2, supraocular tubercle; pn, pronotum; tmp, temple." figureDoi="http://doi.org/10.5281/zenodo.10910599" httpUri="https://zenodo.org/record/10910599/files/figure.png" pageId="2" pageNumber="453">3A, B</figureCitation>
) weakly transverse, widest across eyes,
<quantity id="8D0B95E1FFE4FFA22FF49D8099B9E6EE" box="[920,1019,463,489]" metricMagnitude="-3" metricUnit="m" metricValue="1.06" pageId="2" pageNumber="453" unit="mm" value="1.06">1.06 mm</quantity>
long,
<quantity id="8D0B95E1FFE4FFA228529D809EE3E6EE" box="[1086,1185,463,489]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="453" unit="mm" value="1.6">1.60 mm</quantity>
wide (including eyes), moderately narrower than pronotum. Surface of head with 2 paramedian pairs of dorsal tubercles, both well raised dorsally and swollen; anterior tubercles or protuberances (
<emphasis id="7887E416FFE4FFA22F4F9E5B997DE529" box="[803,831,532,558]" italics="true" pageId="2" pageNumber="453">p1</emphasis>
) rather rectangular, transverse, with their base closed; posterior protuberances (
<emphasis id="7887E416FFE4FFA22DC19E789B88E556" box="[429,458,567,593]" italics="true" pageId="2" pageNumber="453">p2</emphasis>
) elongate-oval, with more raised anterior half, their bases separated by a relatively wide and elevated median stripe divided by a longitudinal furrow (
<emphasis id="7887E416FFE4FFA22F2C9E169916E574" box="[832,852,601,627]" italics="true" pageId="2" pageNumber="453">fv</emphasis>
). Eyes (
<figureCitation id="D2C82481FFE4FFA22FDC9E169E71E573" box="[944,1075,601,628]" captionStart="FIGURE 2" captionStartId="4.[151,250,1854,1878]" captionTargetBox="[164,1423,181,1830]" captionTargetId="figure-28@4.[164,1423,181,1830]" captionTargetPageId="4" captionText="FIGURE 2. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Head and pronotum, dorsal view. B, Head, dorsal view. C, Right lateral half of pronotum, dorsal view. D, Left elytron and scutellum, dorsal view. E, Right metatarsus, dorsal view. F, Left antenna, dorsal view. Abbreviations: a111, antennomeres 111; ey, eyes; fv, longitudinal median furrow on vertex; mtt15, metatarsomeres 15; p1, supraantennal tubercle; p2, supraocular tubercle; tmp, temple. Scale bars: 1.0 mm (A); 0.5 mm (B, C, E); 2.0 mm (D, F)." figureDoi="http://doi.org/10.5281/zenodo.10910595" httpUri="https://zenodo.org/record/10910595/files/figure.png" pageId="2" pageNumber="453">Figs. 2A, B</figureCitation>
,
<figureCitation id="D2C82481FFE4FFA228529E169EC1E573" box="[1086,1155,601,628]" captionStart="FIGURE 3" captionStartId="5.[151,250,1142,1166]" captionTargetBox="[164,1423,181,1118]" captionTargetId="figure-15@5.[164,1423,181,1118]" captionTargetPageId="5" captionText="FIGURE 3. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Head with morphological interpretations, dorsal view. B, Head without morphological interpretations, dorsal view. C, Pronotum with morphological interpretations, dorsal view. C, Pronotum without morphological interpretations, dorsal view.Abbreviations: a12, antennomeres 12; ey, eyes; fs, paramedian fossae at anterior pronotal edge; fv, longitudinal median furrow on vertex; mrp, median elevated ridge of pronotum; p1, supraantennal tubercle; p2, supraocular tubercle; pn, pronotum; tmp, temple." figureDoi="http://doi.org/10.5281/zenodo.10910599" httpUri="https://zenodo.org/record/10910599/files/figure.png" pageId="2" pageNumber="453">3A, B</figureCitation>
,
<emphasis id="7887E416FFE4FFA228E29E159EE5E574" box="[1166,1191,602,627]" italics="true" pageId="2" pageNumber="453">ey</emphasis>
) rather small, as wide as scape, only slightly longer than temples, moderately protuberant laterally. Temples (
<figureCitation id="D2C82481FFE4FFA228109E339F47E590" box="[1148,1285,636,663]" captionStart="FIGURE 2" captionStartId="4.[151,250,1854,1878]" captionTargetBox="[164,1423,181,1830]" captionTargetId="figure-28@4.[164,1423,181,1830]" captionTargetPageId="4" captionText="FIGURE 2. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Head and pronotum, dorsal view. B, Head, dorsal view. C, Right lateral half of pronotum, dorsal view. D, Left elytron and scutellum, dorsal view. E, Right metatarsus, dorsal view. F, Left antenna, dorsal view. Abbreviations: a111, antennomeres 111; ey, eyes; fv, longitudinal median furrow on vertex; mtt15, metatarsomeres 15; p1, supraantennal tubercle; p2, supraocular tubercle; tmp, temple. Scale bars: 1.0 mm (A); 0.5 mm (B, C, E); 2.0 mm (D, F)." figureDoi="http://doi.org/10.5281/zenodo.10910595" httpUri="https://zenodo.org/record/10910595/files/figure.png" pageId="2" pageNumber="453">Figs. 2A, B</figureCitation>
,
<figureCitation id="D2C82481FFE4FFA2297F9E339F19E590" box="[1299,1371,636,663]" captionStart="FIGURE 3" captionStartId="5.[151,250,1142,1166]" captionTargetBox="[164,1423,181,1118]" captionTargetId="figure-15@5.[164,1423,181,1118]" captionTargetPageId="5" captionText="FIGURE 3. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Head with morphological interpretations, dorsal view. B, Head without morphological interpretations, dorsal view. C, Pronotum with morphological interpretations, dorsal view. C, Pronotum without morphological interpretations, dorsal view.Abbreviations: a12, antennomeres 12; ey, eyes; fs, paramedian fossae at anterior pronotal edge; fv, longitudinal median furrow on vertex; mrp, median elevated ridge of pronotum; p1, supraantennal tubercle; p2, supraocular tubercle; pn, pronotum; tmp, temple." figureDoi="http://doi.org/10.5281/zenodo.10910599" httpUri="https://zenodo.org/record/10910599/files/figure.png" pageId="2" pageNumber="453">3A, B</figureCitation>
,
<emphasis id="7887E416FFE4FFA229059E329FD1E591" box="[1385,1427,637,662]" italics="true" pageId="2" pageNumber="453">tmp</emphasis>
) long, prominent. Gular sutures apparently subparallel-sided, narrowly separated. Antennal insertions divided by distance about as wide as length of scape (
<figureCitation id="D2C82481FFE4FFA22E059E8E98A3E5DC" box="[617,737,705,731]" captionStart="FIGURE 3" captionStartId="5.[151,250,1142,1166]" captionTargetBox="[164,1423,181,1118]" captionTargetId="figure-15@5.[164,1423,181,1118]" captionTargetPageId="5" captionText="FIGURE 3. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Head with morphological interpretations, dorsal view. B, Head without morphological interpretations, dorsal view. C, Pronotum with morphological interpretations, dorsal view. C, Pronotum without morphological interpretations, dorsal view.Abbreviations: a12, antennomeres 12; ey, eyes; fs, paramedian fossae at anterior pronotal edge; fv, longitudinal median furrow on vertex; mrp, median elevated ridge of pronotum; p1, supraantennal tubercle; p2, supraocular tubercle; pn, pronotum; tmp, temple." figureDoi="http://doi.org/10.5281/zenodo.10910599" httpUri="https://zenodo.org/record/10910599/files/figure.png" pageId="2" pageNumber="453">Fig. 3A, B</figureCitation>
). Antennae (
<figureCitation id="D2C82481FFE4FFA22F009E8E99FBE5DC" box="[876,953,705,731]" captionStart="FIGURE 1" captionStartId="3.[151,250,1930,1954]" captionTargetBox="[155,1436,179,1904]" captionTargetId="figure-15@3.[151,1436,179,1904]" captionTargetPageId="3" captionText="FIGURE 1. General habitus of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Dorsal view. B, Ventral view. Scale bars: 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10910593" httpUri="https://zenodo.org/record/10910593/files/figure.png" pageId="2" pageNumber="453">Figs. 1</figureCitation>
,
<figureCitation id="D2C82481FFE4FFA22FAA9E8E99A1E5DC" box="[966,995,705,731]" captionStart="FIGURE 2" captionStartId="4.[151,250,1854,1878]" captionTargetBox="[164,1423,181,1830]" captionTargetId="figure-28@4.[164,1423,181,1830]" captionTargetPageId="4" captionText="FIGURE 2. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Head and pronotum, dorsal view. B, Head, dorsal view. C, Right lateral half of pronotum, dorsal view. D, Left elytron and scutellum, dorsal view. E, Right metatarsus, dorsal view. F, Left antenna, dorsal view. Abbreviations: a111, antennomeres 111; ey, eyes; fv, longitudinal median furrow on vertex; mtt15, metatarsomeres 15; p1, supraantennal tubercle; p2, supraocular tubercle; tmp, temple. Scale bars: 1.0 mm (A); 0.5 mm (B, C, E); 2.0 mm (D, F)." figureDoi="http://doi.org/10.5281/zenodo.10910595" httpUri="https://zenodo.org/record/10910595/files/figure.png" pageId="2" pageNumber="453">2F</figureCitation>
,
<figureCitation id="D2C82481FFE4FFA22F829E8E9E4DE5DC" box="[1006,1039,705,731]" captionStart="FIGURE 4" captionStartId="6.[151,250,1909,1933]" captionTargetBox="[176,1412,181,1885]" captionTargetId="figure-28@6.[176,1412,181,1885]" captionTargetPageId="6" captionText="FIGURE 4. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237.A, Head, right antenna, andprothorax, ventral view. B, Forelegs, ventral view. C, Basal part of left antenna, ventral view. D, Mid- and hindlegs, ventral view. E, Abdomen, ventral view. Abbreviations: a36, antennomeres 36; v15, ventrites 15. Scale bars: 1.0 mm (AD); 2.0 mm (E)." figureDoi="http://doi.org/10.5281/zenodo.10910601" httpUri="https://zenodo.org/record/10910601/files/figure.png" pageId="2" pageNumber="453">4C</figureCitation>
) 11-segmented, filiform, somewhat slender, about 0.6 time as long as body (
<quantity id="8D0B95E1FFE4FFA22E379EAB9881E5F8" box="[603,707,740,767]" metricMagnitude="-3" metricUnit="m" metricValue="5.53" pageId="2" pageNumber="453" unit="mm" value="5.53">5.53 mm</quantity>
, left), barely reaching to middle of elytra; scape (antennomere 1,
<emphasis id="7887E416FFE4FFA22CFB9F489AF1E426" box="[151,179,775,801]" italics="true" pageId="2" pageNumber="453">a1</emphasis>
) approximately oval, 1.5 times as long as thick; pedicel (antennomere 2) short, as long as thick, wider than scape; antennomere 3 narrowly elongate, moderately dilated towards apex, nearly about twice as long as 2nd, slightly longer than 4th; antennomeres 410 each narrowly elongate, subcylindrical, gradually narrowing distally; ultimate antennomere less than 4 times as long as thick (3.8 times as long as thick on left side). Labrum scarcely visible.
</paragraph>
<paragraph id="4A4C3804FFE4FFA22CAB9FDD9B1AE399" blockId="2.[151,1437,151,1981]" pageId="2" pageNumber="453">
Pronotum (
<figureCitation id="D2C82481FFE4FFA22D289FDE9B8AE4AC" box="[324,456,913,939]" captionStart="FIGURE 2" captionStartId="4.[151,250,1854,1878]" captionTargetBox="[164,1423,181,1830]" captionTargetId="figure-28@4.[164,1423,181,1830]" captionTargetPageId="4" captionText="FIGURE 2. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Head and pronotum, dorsal view. B, Head, dorsal view. C, Right lateral half of pronotum, dorsal view. D, Left elytron and scutellum, dorsal view. E, Right metatarsus, dorsal view. F, Left antenna, dorsal view. Abbreviations: a111, antennomeres 111; ey, eyes; fv, longitudinal median furrow on vertex; mtt15, metatarsomeres 15; p1, supraantennal tubercle; p2, supraocular tubercle; tmp, temple. Scale bars: 1.0 mm (A); 0.5 mm (B, C, E); 2.0 mm (D, F)." figureDoi="http://doi.org/10.5281/zenodo.10910595" httpUri="https://zenodo.org/record/10910595/files/figure.png" pageId="2" pageNumber="453">Figs. 2A, C</figureCitation>
,
<figureCitation id="D2C82481FFE4FFA22DBF9FDE985AE4AB" box="[467,536,913,940]" captionStart="FIGURE 3" captionStartId="5.[151,250,1142,1166]" captionTargetBox="[164,1423,181,1118]" captionTargetId="figure-15@5.[164,1423,181,1118]" captionTargetPageId="5" captionText="FIGURE 3. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Head with morphological interpretations, dorsal view. B, Head without morphological interpretations, dorsal view. C, Pronotum with morphological interpretations, dorsal view. C, Pronotum without morphological interpretations, dorsal view.Abbreviations: a12, antennomeres 12; ey, eyes; fs, paramedian fossae at anterior pronotal edge; fv, longitudinal median furrow on vertex; mrp, median elevated ridge of pronotum; p1, supraantennal tubercle; p2, supraocular tubercle; pn, pronotum; tmp, temple." figureDoi="http://doi.org/10.5281/zenodo.10910599" httpUri="https://zenodo.org/record/10910599/files/figure.png" pageId="2" pageNumber="453">3C, D</figureCitation>
,
<emphasis id="7887E416FFE4FFA22E4F9FDD987DE4AC" box="[547,575,914,939]" italics="true" pageId="2" pageNumber="453">pn</emphasis>
) subtrapezoidal, moderately transverse, widest at about half length, narrower than elytra,
<quantity id="8D0B95E1FFE4FFA22D4E9FFB9BCAE4C8" box="[290,392,948,975]" metricMagnitude="-3" metricUnit="m" metricValue="1.29" pageId="2" pageNumber="453" unit="mm" value="1.29">1.29 mm</quantity>
long (along midline),
<quantity id="8D0B95E1FFE4FFA22EE29FFB98B6E4C8" box="[654,756,948,975]" metricMagnitude="-3" metricUnit="m" metricValue="1.94" pageId="2" pageNumber="453" unit="mm" value="1.94">1.94 mm</quantity>
wide; lateral edges broadly angulate, strongly narrowed in anterior 2/5; anterolateral angles right to obtuse, slightly projecting; posterolateral corners broadly arcuate; posterior margin weakly bisinuate, slightly projecting posteriorly, with a small and weak emargination medially; disc moderately convex dorsally (medial area only, outer areas somewhat explanate), with a median elevated ridge, or stripe, along entire length (
<figureCitation id="D2C82481FFE4FFA22DA598709807E35E" box="[457,581,1087,1113]" captionStart="FIGURE 3" captionStartId="5.[151,250,1142,1166]" captionTargetBox="[164,1423,181,1118]" captionTargetId="figure-15@5.[164,1423,181,1118]" captionTargetPageId="5" captionText="FIGURE 3. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Head with morphological interpretations, dorsal view. B, Head without morphological interpretations, dorsal view. C, Pronotum with morphological interpretations, dorsal view. C, Pronotum without morphological interpretations, dorsal view.Abbreviations: a12, antennomeres 12; ey, eyes; fs, paramedian fossae at anterior pronotal edge; fv, longitudinal median furrow on vertex; mrp, median elevated ridge of pronotum; p1, supraantennal tubercle; p2, supraocular tubercle; pn, pronotum; tmp, temple." figureDoi="http://doi.org/10.5281/zenodo.10910599" httpUri="https://zenodo.org/record/10910599/files/figure.png" pageId="2" pageNumber="453">Fig. 3C, D</figureCitation>
,
<emphasis id="7887E416FFE4FFA22E3E9870983DE35F" box="[594,639,1087,1112]" italics="true" pageId="2" pageNumber="453">mrp</emphasis>
), as well as a pair of shallow paramedian fossae at anterior edge (
<figureCitation id="D2C82481FFE4FFA2290398709A9EE37B" captionStart="FIGURE 3" captionStartId="5.[151,250,1142,1166]" captionTargetBox="[164,1423,181,1118]" captionTargetId="figure-15@5.[164,1423,181,1118]" captionTargetPageId="5" captionText="FIGURE 3. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Head with morphological interpretations, dorsal view. B, Head without morphological interpretations, dorsal view. C, Pronotum with morphological interpretations, dorsal view. C, Pronotum without morphological interpretations, dorsal view.Abbreviations: a12, antennomeres 12; ey, eyes; fs, paramedian fossae at anterior pronotal edge; fv, longitudinal median furrow on vertex; mrp, median elevated ridge of pronotum; p1, supraantennal tubercle; p2, supraocular tubercle; pn, pronotum; tmp, temple." figureDoi="http://doi.org/10.5281/zenodo.10910599" httpUri="https://zenodo.org/record/10910599/files/figure.png" pageId="2" pageNumber="453">Fig. 3C, D</figureCitation>
,
<emphasis id="7887E416FFE4FFA22C8A982E9ABBE37C" box="[230,249,1121,1147]" italics="true" pageId="2" pageNumber="453">fs</emphasis>
), but lacking a longitudinal furrow along midline. Prosternal process seemingly narrow. Scutellum minute, subquadrangular.
</paragraph>
<paragraph id="4A4C3804FFE4FFA22CAB98E89A96E268" blockId="2.[151,1437,151,1981]" pageId="2" pageNumber="453">
Elytra (
<figureCitation id="D2C82481FFE4FFA22D7798E89B3DE3C6" box="[283,383,1191,1217]" captionStart="FIGURE 1" captionStartId="3.[151,250,1930,1954]" captionTargetBox="[155,1436,179,1904]" captionTargetId="figure-15@3.[151,1436,179,1904]" captionTargetPageId="3" captionText="FIGURE 1. General habitus of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Dorsal view. B, Ventral view. Scale bars: 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10910593" httpUri="https://zenodo.org/record/10910593/files/figure.png" pageId="2" pageNumber="453">Figs. 1A</figureCitation>
,
<figureCitation id="D2C82481FFE4FFA22DE798E89BECE3C6" box="[395,430,1191,1217]" captionStart="FIGURE 2" captionStartId="4.[151,250,1854,1878]" captionTargetBox="[164,1423,181,1830]" captionTargetId="figure-28@4.[164,1423,181,1830]" captionTargetPageId="4" captionText="FIGURE 2. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Head and pronotum, dorsal view. B, Head, dorsal view. C, Right lateral half of pronotum, dorsal view. D, Left elytron and scutellum, dorsal view. E, Right metatarsus, dorsal view. F, Left antenna, dorsal view. Abbreviations: a111, antennomeres 111; ey, eyes; fv, longitudinal median furrow on vertex; mtt15, metatarsomeres 15; p1, supraantennal tubercle; p2, supraocular tubercle; tmp, temple. Scale bars: 1.0 mm (A); 0.5 mm (B, C, E); 2.0 mm (D, F)." figureDoi="http://doi.org/10.5281/zenodo.10910595" httpUri="https://zenodo.org/record/10910595/files/figure.png" pageId="2" pageNumber="453">2D</figureCitation>
) narrowly elongate, gradually divergent posteriorly, widest in posterior 1/3 (right and left elytron
<quantity id="8D0B95E1FFE4FFA22C8298869B11E3E3" box="[238,339,1225,1252]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="2" pageNumber="453" unit="mm" value="6.7">6.70 mm</quantity>
long from base to apex, right elytron
<quantity id="8D0B95E1FFE4FFA22E9398869926E3E3" box="[767,868,1225,1252]" metricMagnitude="-3" metricUnit="m" metricValue="1.48" pageId="2" pageNumber="453" unit="mm" value="1.48">1.48 mm</quantity>
wide, left elytron
<quantity id="8D0B95E1FFE4FFA2285898869ED9E3E3" box="[1076,1179,1225,1252]" metricMagnitude="-3" metricUnit="m" metricValue="1.79" pageId="2" pageNumber="453" unit="mm" value="1.79">1.79 mm</quantity>
; the distinct difference of the width of the elytra must be an artefact), with acute elytral apices (not emarginate); surface each with 9 longitudinal rows of sub-oval to rounded window-punctures (window-punctures regularly arranged; some punctures rather transverse near base of elytra); 5 interval elevations present, all distinctly raised from base or subbase to apex.
</paragraph>
<paragraph id="4A4C3804FFE4FFA22CAB993899F1E054" blockId="2.[151,1437,151,1981]" pageId="2" pageNumber="453">
Legs (
<figureCitation id="D2C82481FFE4FFA22D6199389B18E296" box="[269,346,1399,1425]" captionStart="FIGURE 1" captionStartId="3.[151,250,1930,1954]" captionTargetBox="[155,1436,179,1904]" captionTargetId="figure-15@3.[151,1436,179,1904]" captionTargetPageId="3" captionText="FIGURE 1. General habitus of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Dorsal view. B, Ventral view. Scale bars: 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10910593" httpUri="https://zenodo.org/record/10910593/files/figure.png" pageId="2" pageNumber="453">
Figs.
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</figureCitation>
,
<figureCitation id="D2C82481FFE4FFA22D0B99389BCBE296" box="[359,393,1399,1425]" captionStart="FIGURE 2" captionStartId="4.[151,250,1854,1878]" captionTargetBox="[164,1423,181,1830]" captionTargetId="figure-28@4.[164,1423,181,1830]" captionTargetPageId="4" captionText="FIGURE 2. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Head and pronotum, dorsal view. B, Head, dorsal view. C, Right lateral half of pronotum, dorsal view. D, Left elytron and scutellum, dorsal view. E, Right metatarsus, dorsal view. F, Left antenna, dorsal view. Abbreviations: a111, antennomeres 111; ey, eyes; fv, longitudinal median furrow on vertex; mtt15, metatarsomeres 15; p1, supraantennal tubercle; p2, supraocular tubercle; tmp, temple. Scale bars: 1.0 mm (A); 0.5 mm (B, C, E); 2.0 mm (D, F)." figureDoi="http://doi.org/10.5281/zenodo.10910595" httpUri="https://zenodo.org/record/10910595/files/figure.png" pageId="2" pageNumber="453">2E</figureCitation>
,
<figureCitation id="D2C82481FFE4FFA22DF899389BB4E296" box="[404,502,1399,1425]" captionStart="FIGURE 4" captionStartId="6.[151,250,1909,1933]" captionTargetBox="[176,1412,181,1885]" captionTargetId="figure-28@6.[176,1412,181,1885]" captionTargetPageId="6" captionText="FIGURE 4. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237.A, Head, right antenna, andprothorax, ventral view. B, Forelegs, ventral view. C, Basal part of left antenna, ventral view. D, Mid- and hindlegs, ventral view. E, Abdomen, ventral view. Abbreviations: a36, antennomeres 36; v15, ventrites 15. Scale bars: 1.0 mm (AD); 2.0 mm (E)." figureDoi="http://doi.org/10.5281/zenodo.10910601" httpUri="https://zenodo.org/record/10910601/files/figure.png" pageId="2" pageNumber="453">4B, D, E</figureCitation>
) moderately long, relatively thick; femora stout, clavate, metafemora not extending beyond lateral edge of posterior body; tibiae slender, bar-like, gradually slightly widened toward apex, much thinner than corresponding femora (but not less than half as wide), with metatibial length of
<quantity id="8D0B95E1FFE4FFA2283C99F39EF6E2D0" box="[1104,1204,1468,1495]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="2" pageNumber="453" unit="mm" value="0.64">0.64 mm</quantity>
(left). Tarsi (
<figureCitation id="D2C82481FFE4FFA2292499F39FD7E2D1" box="[1352,1429,1468,1494]" captionStart="FIGURE 1" captionStartId="3.[151,250,1930,1954]" captionTargetBox="[155,1436,179,1904]" captionTargetId="figure-15@3.[151,1436,179,1904]" captionTargetPageId="3" captionText="FIGURE 1. General habitus of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Dorsal view. B, Ventral view. Scale bars: 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10910593" httpUri="https://zenodo.org/record/10910593/files/figure.png" pageId="2" pageNumber="453">
Figs.
<geoCoordinate id="2FC75EC3FFE4FFA229EB99F39FD7E2D1" box="[1415,1429,1468,1494]" degrees="1, 2" direction="east" orientation="longitude" pageId="2" pageNumber="453" precision="5555" value="1.2">1</geoCoordinate>
</figureCitation>
,
<figureCitation id="D2C82481FFE4FFA22CFB99909AFBE2FE" box="[151,185,1503,1529]" captionStart="FIGURE 2" captionStartId="4.[151,250,1854,1878]" captionTargetBox="[164,1423,181,1830]" captionTargetId="figure-28@4.[164,1423,181,1830]" captionTargetPageId="4" captionText="FIGURE 2. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Head and pronotum, dorsal view. B, Head, dorsal view. C, Right lateral half of pronotum, dorsal view. D, Left elytron and scutellum, dorsal view. E, Right metatarsus, dorsal view. F, Left antenna, dorsal view. Abbreviations: a111, antennomeres 111; ey, eyes; fv, longitudinal median furrow on vertex; mtt15, metatarsomeres 15; p1, supraantennal tubercle; p2, supraocular tubercle; tmp, temple. Scale bars: 1.0 mm (A); 0.5 mm (B, C, E); 2.0 mm (D, F)." figureDoi="http://doi.org/10.5281/zenodo.10910595" httpUri="https://zenodo.org/record/10910595/files/figure.png" pageId="2" pageNumber="453">2E</figureCitation>
,
<figureCitation id="D2C82481FFE4FFA22CA899909B64E2FE" box="[196,294,1503,1529]" captionStart="FIGURE 4" captionStartId="6.[151,250,1909,1933]" captionTargetBox="[176,1412,181,1885]" captionTargetId="figure-28@6.[176,1412,181,1885]" captionTargetPageId="6" captionText="FIGURE 4. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237.A, Head, right antenna, andprothorax, ventral view. B, Forelegs, ventral view. C, Basal part of left antenna, ventral view. D, Mid- and hindlegs, ventral view. E, Abdomen, ventral view. Abbreviations: a36, antennomeres 36; v15, ventrites 15. Scale bars: 1.0 mm (AD); 2.0 mm (E)." figureDoi="http://doi.org/10.5281/zenodo.10910601" httpUri="https://zenodo.org/record/10910601/files/figure.png" pageId="2" pageNumber="453">4B, D, E</figureCitation>
) 5-segmented, slightly narrower than corresponding tibiae; protarsi (
<figureCitation id="D2C82481FFE4FFA2284A99909EDFE2FE" box="[1062,1181,1503,1529]" captionStart="FIGURE 4" captionStartId="6.[151,250,1909,1933]" captionTargetBox="[176,1412,181,1885]" captionTargetId="figure-28@6.[176,1412,181,1885]" captionTargetPageId="6" captionText="FIGURE 4. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237.A, Head, right antenna, andprothorax, ventral view. B, Forelegs, ventral view. C, Basal part of left antenna, ventral view. D, Mid- and hindlegs, ventral view. E, Abdomen, ventral view. Abbreviations: a36, antennomeres 36; v15, ventrites 15. Scale bars: 1.0 mm (AD); 2.0 mm (E)." figureDoi="http://doi.org/10.5281/zenodo.10910601" httpUri="https://zenodo.org/record/10910601/files/figure.png" pageId="2" pageNumber="453">Fig. 4B, D</figureCitation>
) shortened and robust; protarsomere 1 slightly longer than wide, protarsomeres 24 subequal in length, each shorter than 1st, ventrally expanded; protarsomere 5 apparently as long as 1st, but apical part not well visible due to milky coarting of micro airbubbles; mesotarsi (
<figureCitation id="D2C82481FFE4FFA22DF89A089848E166" box="[404,522,1607,1633]" captionStart="FIGURE 4" captionStartId="6.[151,250,1909,1933]" captionTargetBox="[176,1412,181,1885]" captionTargetId="figure-28@6.[176,1412,181,1885]" captionTargetPageId="6" captionText="FIGURE 4. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237.A, Head, right antenna, andprothorax, ventral view. B, Forelegs, ventral view. C, Basal part of left antenna, ventral view. D, Mid- and hindlegs, ventral view. E, Abdomen, ventral view. Abbreviations: a36, antennomeres 36; v15, ventrites 15. Scale bars: 1.0 mm (AD); 2.0 mm (E)." figureDoi="http://doi.org/10.5281/zenodo.10910601" httpUri="https://zenodo.org/record/10910601/files/figure.png" pageId="2" pageNumber="453">Fig. 4D, E</figureCitation>
) slender; mesotarsomere 1 elongate, about twice as long as wide, mesotarsomeres 23 subequal in length, each wider than long, shorter than 1st, mesotarsomere 4 slightly longer than 3rd but shorter than 1st, as wide as two preceding segments, mesotarsomere 5 seemingly as long as 1st; metatarsi (
<figureCitation id="D2C82481FFE4FFA2288B9AC39F05E1A1" box="[1255,1351,1676,1702]" captionStart="FIGURE 2" captionStartId="4.[151,250,1854,1878]" captionTargetBox="[164,1423,181,1830]" captionTargetId="figure-28@4.[164,1423,181,1830]" captionTargetPageId="4" captionText="FIGURE 2. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Head and pronotum, dorsal view. B, Head, dorsal view. C, Right lateral half of pronotum, dorsal view. D, Left elytron and scutellum, dorsal view. E, Right metatarsus, dorsal view. F, Left antenna, dorsal view. Abbreviations: a111, antennomeres 111; ey, eyes; fv, longitudinal median furrow on vertex; mtt15, metatarsomeres 15; p1, supraantennal tubercle; p2, supraocular tubercle; tmp, temple. Scale bars: 1.0 mm (A); 0.5 mm (B, C, E); 2.0 mm (D, F)." figureDoi="http://doi.org/10.5281/zenodo.10910595" httpUri="https://zenodo.org/record/10910595/files/figure.png" pageId="2" pageNumber="453">Figs. 2E</figureCitation>
,
<figureCitation id="D2C82481FFE4FFA2293E9AC39FD6E1A1" box="[1362,1428,1676,1702]" captionStart="FIGURE 4" captionStartId="6.[151,250,1909,1933]" captionTargetBox="[176,1412,181,1885]" captionTargetId="figure-28@6.[176,1412,181,1885]" captionTargetPageId="6" captionText="FIGURE 4. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237.A, Head, right antenna, andprothorax, ventral view. B, Forelegs, ventral view. C, Basal part of left antenna, ventral view. D, Mid- and hindlegs, ventral view. E, Abdomen, ventral view. Abbreviations: a36, antennomeres 36; v15, ventrites 15. Scale bars: 1.0 mm (AD); 2.0 mm (E)." figureDoi="http://doi.org/10.5281/zenodo.10910601" httpUri="https://zenodo.org/record/10910601/files/figure.png" pageId="2" pageNumber="453">4D, E</figureCitation>
) slender; metatarsomere 1 (
<figureCitation id="D2C82481FFE4FFA22DAE9AE0985AE1CE" box="[450,536,1711,1737]" captionStart="FIGURE 2" captionStartId="4.[151,250,1854,1878]" captionTargetBox="[164,1423,181,1830]" captionTargetId="figure-28@4.[164,1423,181,1830]" captionTargetPageId="4" captionText="FIGURE 2. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Head and pronotum, dorsal view. B, Head, dorsal view. C, Right lateral half of pronotum, dorsal view. D, Left elytron and scutellum, dorsal view. E, Right metatarsus, dorsal view. F, Left antenna, dorsal view. Abbreviations: a111, antennomeres 111; ey, eyes; fv, longitudinal median furrow on vertex; mtt15, metatarsomeres 15; p1, supraantennal tubercle; p2, supraocular tubercle; tmp, temple. Scale bars: 1.0 mm (A); 0.5 mm (B, C, E); 2.0 mm (D, F)." figureDoi="http://doi.org/10.5281/zenodo.10910595" httpUri="https://zenodo.org/record/10910595/files/figure.png" pageId="2" pageNumber="453">Fig. 2E</figureCitation>
,
<emphasis id="7887E416FFE4FFA22E489AE09814E1CE" box="[548,598,1711,1737]" italics="true" pageId="2" pageNumber="453">mtt1</emphasis>
) elongate, about 2.9 times as long as wide, about 1.6 times as long as 5th, metatarsomere 2 elongate, sub-cylindrical, about 1.6 times as long as wide, about half as long as 1st, metatarsomere 3 short, very slightly wider than long, distinctly shorter than 2nd, metatarsomere 4 short, widest, bilobed ventrally, moderately longer than 3rd, as long as 2nd, metatarsomere 5 narrowly elongate, slender, strongly dilated apically, widest near apex, about 2.7 times as long as wide. Claws short, simple.
</paragraph>
<paragraph id="4A4C3804FFE4FFA22CAB9B139E0FE0BA" blockId="2.[151,1437,151,1981]" pageId="2" pageNumber="453">
Abdomen (
<figureCitation id="D2C82481FFE4FFA22D269B139BDFE071" box="[330,413,1884,1910]" captionStart="FIGURE 1" captionStartId="3.[151,250,1930,1954]" captionTargetBox="[155,1436,179,1904]" captionTargetId="figure-15@3.[151,1436,179,1904]" captionTargetPageId="3" captionText="FIGURE 1. General habitus of Cupes balticus sp. n., holotype, SEHU-0000121237. A, Dorsal view. B, Ventral view. Scale bars: 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10910593" httpUri="https://zenodo.org/record/10910593/files/figure.png" pageId="2" pageNumber="453">Figs. 1</figureCitation>
,
<figureCitation id="D2C82481FFE4FFA22DDC9B139BBAE071" box="[432,504,1884,1910]" captionStart="FIGURE 4" captionStartId="6.[151,250,1909,1933]" captionTargetBox="[176,1412,181,1885]" captionTargetId="figure-28@6.[176,1412,181,1885]" captionTargetPageId="6" captionText="FIGURE 4. Morphological details of Cupes balticus sp. n., holotype, SEHU-0000121237.A, Head, right antenna, andprothorax, ventral view. B, Forelegs, ventral view. C, Basal part of left antenna, ventral view. D, Mid- and hindlegs, ventral view. E, Abdomen, ventral view. Abbreviations: a36, antennomeres 36; v15, ventrites 15. Scale bars: 1.0 mm (AD); 2.0 mm (E)." figureDoi="http://doi.org/10.5281/zenodo.10910601" httpUri="https://zenodo.org/record/10910601/files/figure.png" pageId="2" pageNumber="453">4D, E</figureCitation>
) elongate, apex acute, with five visible ventrites; tergite VIII well sclerotized, semicircular, closely covered with minute setae; ventrite 5 twice as long as 4th; all ventrites except the 1st markedly depressed anteriorly and overlapping the anterior margin of each successive ventrite.
</paragraph>
<caption id="1E8C688CFFE5FFA32CFB9BC59B61E0C1" ID-DOI="http://doi.org/10.5281/zenodo.10910593" ID-Zenodo-Dep="10910593" httpUri="https://zenodo.org/record/10910593/files/figure.png" pageId="3" pageNumber="454" startId="3.[151,250,1930,1954]" targetBox="[155,1436,179,1904]" targetPageId="3" targetType="figure">
<paragraph id="4A4C3804FFE5FFA32CFB9BC59B61E0C1" blockId="3.[151,1436,1930,1990]" pageId="3" pageNumber="454">
<emphasis id="7887E416FFE5FFA32CFB9BC59B57E0A5" bold="true" box="[151,277,1930,1954]" pageId="3" pageNumber="454">FIGURE 1.</emphasis>
General habitus of
<taxonomicName id="8DF34387FFE5FFA32D889BC5983EE0A5" authority="Yamamoto, 2024" authorityName="Yamamoto" authorityYear="2024" box="[484,636,1930,1954]" class="Insecta" family="Cupedidae" genus="Cupes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="balticus" status="sp. nov.">
<emphasis id="7887E416FFE5FFA32D889BC5983EE0A5" box="[484,636,1930,1954]" italics="true" pageId="3" pageNumber="454">Cupes balticus</emphasis>
</taxonomicName>
<emphasis id="7887E416FFE5FFA32EE89BC598FDE0A5" bold="true" box="[644,703,1930,1954]" pageId="3" pageNumber="454">
<taxonomicNameLabel id="63B4596DFFE5FFA32EE89BC598FDE0A5" box="[644,703,1930,1954]" pageId="3" pageNumber="454" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype, SEHU-0000121237.
<emphasis id="7887E416FFE5FFA328609BC59E5CE0A5" bold="true" box="[1036,1054,1930,1954]" pageId="3" pageNumber="454">A</emphasis>
, Dorsal view.
<emphasis id="7887E416FFE5FFA328DB9BC59E8AE0A5" bold="true" box="[1207,1224,1930,1954]" pageId="3" pageNumber="454">B</emphasis>
, Ventral view. Scale bars: 3.0 mm.
</paragraph>
</caption>
<caption id="1E8C688CFFE2FFA42CFB9B7198EAE0E1" ID-DOI="http://doi.org/10.5281/zenodo.10910595" ID-Zenodo-Dep="10910595" httpUri="https://zenodo.org/record/10910595/files/figure.png" pageId="4" pageNumber="455" startId="4.[151,250,1854,1878]" targetBox="[164,1423,181,1830]" targetPageId="4" targetType="figure">
<paragraph id="4A4C3804FFE2FFA42CFB9B7198EAE0E1" blockId="4.[151,1436,1854,2022]" pageId="4" pageNumber="455">
<emphasis id="7887E416FFE2FFA42CFB9B719B54E051" bold="true" box="[151,278,1854,1878]" pageId="4" pageNumber="455">FIGURE 2.</emphasis>
Morphological details of
<taxonomicName id="8DF34387FFE2FFA42E479B709886E051" authority="Yamamoto, 2024" authorityName="Yamamoto" authorityYear="2024" box="[555,708,1854,1878]" class="Insecta" family="Cupedidae" genus="Cupes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="balticus" status="sp. nov.">
<emphasis id="7887E416FFE2FFA42E479B709886E051" box="[555,708,1854,1878]" italics="true" pageId="4" pageNumber="455">Cupes balticus</emphasis>
</taxonomicName>
<emphasis id="7887E416FFE2FFA42EA29B719948E051" bold="true" box="[718,778,1854,1878]" pageId="4" pageNumber="455">
<taxonomicNameLabel id="63B4596DFFE2FFA42EA29B719948E051" box="[718,778,1854,1878]" pageId="4" pageNumber="455" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype, SEHU-0000121237.
<emphasis id="7887E416FFE2FFA428309B719E2CE051" bold="true" box="[1116,1134,1854,1878]" pageId="4" pageNumber="455">A</emphasis>
, Head and pronotum, dorsal view.
<emphasis id="7887E416FFE2FFA42CBF9B2D9AA6E07D" bold="true" box="[211,228,1890,1914]" pageId="4" pageNumber="455">B</emphasis>
, Head, dorsal view.
<emphasis id="7887E416FFE2FFA42DDF9B2D9B87E07D" bold="true" box="[435,453,1890,1914]" pageId="4" pageNumber="455">C</emphasis>
, Right lateral half of pronotum, dorsal view.
<emphasis id="7887E416FFE2FFA42FE39B2D99E3E07D" bold="true" box="[911,929,1890,1914]" pageId="4" pageNumber="455">D</emphasis>
, Left elytron and scutellum, dorsal view.
<emphasis id="7887E416FFE2FFA4292A9B2D9F15E07D" bold="true" box="[1350,1367,1890,1914]" pageId="4" pageNumber="455">E</emphasis>
, Right metatarsus, dorsal view.
<emphasis id="7887E416FFE2FFA42DF79BC99BE8E099" bold="true" box="[411,426,1926,1950]" pageId="4" pageNumber="455">F</emphasis>
, Left antenna, dorsal view. Abbreviations: a111, antennomeres 111; ey, eyes; fv, longitudinal median furrow on vertex; mtt15, metatarsomeres 15; p1, supraantennal tubercle; p2, supraocular tubercle; tmp, temple. Scale bars: 1.0 mm (A); 0.5 mm (B, C, E); 2.0 mm (D, F).
</paragraph>
</caption>
<caption id="1E8C688CFFE3FFA52CFB98399ECFE219" ID-DOI="http://doi.org/10.5281/zenodo.10910599" ID-Zenodo-Dep="10910599" httpUri="https://zenodo.org/record/10910599/files/figure.png" pageId="5" pageNumber="456" startId="5.[151,250,1142,1166]" targetBox="[164,1423,181,1118]" targetPageId="5" targetType="figure">
<paragraph id="4A4C3804FFE3FFA52CFB98399ECFE219" blockId="5.[151,1437,1142,1310]" pageId="5" pageNumber="456">
<emphasis id="7887E416FFE3FFA52CFB98399B5AE389" bold="true" box="[151,280,1142,1166]" pageId="5" pageNumber="456">FIGURE 3.</emphasis>
Morphological details of
<taxonomicName id="8DF34387FFE3FFA52E5E9838988FE389" authority="Yamamoto, 2024" authorityName="Yamamoto" authorityYear="2024" box="[562,717,1142,1166]" class="Insecta" family="Cupedidae" genus="Cupes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="balticus" status="sp. nov.">
<emphasis id="7887E416FFE3FFA52E5E9838988FE389" box="[562,717,1142,1166]" italics="true" pageId="5" pageNumber="456">Cupes balticus</emphasis>
</taxonomicName>
<emphasis id="7887E416FFE3FFA52EB498399954E389" bold="true" box="[728,790,1142,1166]" pageId="5" pageNumber="456">
<taxonomicNameLabel id="63B4596DFFE3FFA52EB498399954E389" box="[728,790,1142,1166]" pageId="5" pageNumber="456" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype, SEHU-0000121237.
<emphasis id="7887E416FFE3FFA5280098399E3CE389" bold="true" box="[1132,1150,1142,1166]" pageId="5" pageNumber="456">A</emphasis>
, Head with morphological interpretations, dorsal view.
<emphasis id="7887E416FFE3FFA52DAD98D59B90E3B5" bold="true" box="[449,466,1178,1202]" pageId="5" pageNumber="456">B</emphasis>
, Head without morphological interpretations, dorsal view.
<emphasis id="7887E416FFE3FFA5282D98D59E11E3B5" bold="true" box="[1089,1107,1178,1202]" pageId="5" pageNumber="456">C</emphasis>
, Pronotum with morphological interpretations, dorsal view.
<emphasis id="7887E416FFE3FFA52DDD98F19B81E3D1" bold="true" box="[433,451,1214,1238]" pageId="5" pageNumber="456">C</emphasis>
, Pronotum without morphological interpretations, dorsal view.Abbreviations: a12, antennomeres 12; ey, eyes; fs, paramedian fossae at anterior pronotal edge; fv, longitudinal median furrow on vertex; mrp, median elevated ridge of pronotum; p1, supraantennal tubercle; p2, supraocular tubercle; pn, pronotum; tmp, temple.
</paragraph>
</caption>
<paragraph id="4A4C3804FFE3FFA52CAB99039E29E1AD" blockId="5.[151,1437,1356,2031]" pageId="5" pageNumber="456">
<emphasis id="7887E416FFE3FFA52CAB99039BECE260" bold="true" box="[199,430,1356,1383]" pageId="5" pageNumber="456">Comparative notes.</emphasis>
The new species is most closely related to
<taxonomicName id="8DF34387FFE3FFA52FEC99029EBCE260" authority="Kirejtshuk, 2005" authorityName="Kirejtshuk" authorityYear="2005" box="[896,1278,1356,1383]" class="Insecta" family="Cupedidae" genus="Cupes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="weitschati">
<emphasis id="7887E416FFE3FFA52FEC99029E7FE261" box="[896,1085,1356,1383]" italics="true" pageId="5" pageNumber="456">Cupes weitschati</emphasis>
<bibRefCitation id="2E6245F5FFE3FFA5282D99039EBCE260" author="Kirejtshuk, A. G." box="[1089,1278,1356,1383]" pageId="5" pageNumber="456" pagination="55 - 84" refId="ref4823" refString="Kirejtshuk, A. G. (2005) A revision of the genus Cupes Fabricius, 1801 from Baltic amber and some notes on taxonomy and composition of the family Cupedidae (Coleoptera, Archostemata). Mitteilungen aus dem Geologisch-Palaontologischen Institut an der Universitat Hamburg, 89, 55 - 84." type="journal article" year="2005">Kirejtshuk, 2005</bibRefCitation>
</taxonomicName>
, characterized by two paramedian pairs of similar dorsal tubercles on the head, comparable eye size, and bi-colored elytra with a distinct pattern. It is differentiated by its anteriorly pointed pronotal corners with angulate lateral margins, uniformly brownish head and pronotum, the presence of two interrupted bands in the apical and subapical parts of the elytra, and a more narrowly elongate ultimate antennomere. Among the Baltic amber cupedid taxa,
<taxonomicName id="8DF34387FFE3FFA528C299929F17E2F1" authority="Yamamoto, 2024" authorityName="Yamamoto" authorityYear="2024" box="[1198,1365,1500,1527]" class="Insecta" family="Cupedidae" genus="Cupes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="balticus" status="sp. nov.">
<emphasis id="7887E416FFE3FFA528C299929F17E2F1" box="[1198,1365,1500,1527]" italics="true" pageId="5" pageNumber="456">Cupes balticus</emphasis>
</taxonomicName>
<emphasis id="7887E416FFE3FFA5293099929FDFE2F0" bold="true" box="[1372,1437,1501,1527]" pageId="5" pageNumber="456">
<taxonomicNameLabel id="63B4596DFFE3FFA5293099929FDFE2F0" box="[1372,1437,1501,1527]" pageId="5" pageNumber="456" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
does not align with the monospecific extinct genus
<taxonomicName id="8DF34387FFE3FFA52EAA9A4E9965E11D" authorityName="Kirejtshuk, Nel &amp; Kirejtshuk" authorityYear="2016" box="[710,807,1537,1562]" class="Insecta" family="Cupedidae" genus="Taxopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="genus">
<emphasis id="7887E416FFE3FFA52EAA9A4E9965E11D" box="[710,807,1537,1562]" italics="true" pageId="5" pageNumber="456">Taxopsis</emphasis>
</taxonomicName>
due to its much smaller body size (body length
<quantity id="8D0B95E1FFE3FFA529409A4F9A81E138" metricMagnitude="-2" metricUnit="m" metricValue="1.71" metricValueMax="1.81" metricValueMin="1.61" pageId="5" pageNumber="456" unit="mm" value="17.1" valueMax="18.1" valueMin="16.1">16.118.1 mm</quantity>
in
<emphasis id="7887E416FFE3FFA52C849A6A9B32E139" box="[232,368,1573,1598]" italics="true" pageId="5" pageNumber="456">
<taxonomicName id="8DF34387FFE3FFA52C849A6A9B0BE139" authorityName="Kirejtshuk, Nel &amp; Kirejtshuk" authorityYear="2016" box="[232,329,1573,1598]" class="Insecta" family="Cupedidae" genus="Taxopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="genus">Taxopsis</taxonomicName>
vs.
</emphasis>
<quantity id="8D0B95E1FFE3FFA52D1B9A6A9B8CE138" box="[375,462,1573,1599]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="5" pageNumber="456" unit="mm" value="9.3">9.3 mm</quantity>
in the new species), narrowly separated procoxae, and more or less oval-to-polygonal elytral cells (compared to strongly transverse subquadrangular cells in
<taxonomicName id="8DF34387FFE3FFA52FAF9A069E66E165" authorityName="Kirejtshuk, Nel &amp; Kirejtshuk" authorityYear="2016" box="[963,1060,1609,1634]" class="Insecta" family="Cupedidae" genus="Taxopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="genus">
<emphasis id="7887E416FFE3FFA52FAF9A069E66E165" box="[963,1060,1609,1634]" italics="true" pageId="5" pageNumber="456">Taxopsis</emphasis>
</taxonomicName>
) (
<bibRefCitation id="2E6245F5FFE3FFA528529A079EB9E164" author="Kirejtshuk, A. G." box="[1086,1275,1608,1635]" pageId="5" pageNumber="456" pagination="55 - 84" refId="ref4823" refString="Kirejtshuk, A. G. (2005) A revision of the genus Cupes Fabricius, 1801 from Baltic amber and some notes on taxonomy and composition of the family Cupedidae (Coleoptera, Archostemata). Mitteilungen aus dem Geologisch-Palaontologischen Institut an der Universitat Hamburg, 89, 55 - 84." type="journal article" year="2005">Kirejtshuk 2005</bibRefCitation>
;
<bibRefCitation id="2E6245F5FFE3FFA529659A079ABAE181" author="Kirejtshuk, A. G. &amp; Nel, A. &amp; Kirejtshuk, P. A." pageId="5" pageNumber="456" pagination="61 - 190" refId="ref4926" refString="Kirejtshuk, A. G., Nel, A. &amp; Kirejtshuk, P. A. (2016) Taxonomy of the reticulate beetles of the subfamily Cupedinae (Coleoptera: Archostemata), with a review of the historical development. Invertebrate Zoology, 13 (2), 61 - 190. https: // doi. org / 10.15298 / invertzool. 13.2.01" type="journal article" year="2016">
Kirejtshuk
<emphasis id="7887E416FFE3FFA529E49A069AEFE181" italics="true" pageId="5" pageNumber="456">et al</emphasis>
. 2016
</bibRefCitation>
). Distinguished from the other six species of
<taxonomicName id="8DF34387FFE3FFA52E999A229979E180" authorityName="Fabricius" authorityYear="1801" box="[757,827,1645,1671]" class="Insecta" family="Cupedidae" genus="Cupes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="genus">
<emphasis id="7887E416FFE3FFA52E999A229979E180" box="[757,827,1645,1671]" italics="true" pageId="5" pageNumber="456">Cupes</emphasis>
</taxonomicName>
in Baltic amber, the new species is differentiated by a combination of the following character states (
<bibRefCitation id="2E6245F5FFE3FFA52ECC9ADF9918E1AC" author="Kirejtshuk, A. G." box="[672,858,1680,1707]" pageId="5" pageNumber="456" pagination="55 - 84" refId="ref4823" refString="Kirejtshuk, A. G. (2005) A revision of the genus Cupes Fabricius, 1801 from Baltic amber and some notes on taxonomy and composition of the family Cupedidae (Coleoptera, Archostemata). Mitteilungen aus dem Geologisch-Palaontologischen Institut an der Universitat Hamburg, 89, 55 - 84." type="journal article" year="2005">Kirejtshuk 2005</bibRefCitation>
;
<bibRefCitation id="2E6245F5FFE3FFA52F0A9ADF9E1DE1AD" author="Kirejtshuk, A. G. &amp; Nel, A. &amp; Kirejtshuk, P. A." box="[870,1119,1680,1707]" pageId="5" pageNumber="456" pagination="61 - 190" refId="ref4926" refString="Kirejtshuk, A. G., Nel, A. &amp; Kirejtshuk, P. A. (2016) Taxonomy of the reticulate beetles of the subfamily Cupedinae (Coleoptera: Archostemata), with a review of the historical development. Invertebrate Zoology, 13 (2), 61 - 190. https: // doi. org / 10.15298 / invertzool. 13.2.01" type="journal article" year="2016">
Kirejtshuk
<emphasis id="7887E416FFE3FFA52F8F9ADE9E56E1AD" box="[995,1044,1680,1706]" italics="true" pageId="5" pageNumber="456">et al</emphasis>
. 2016
</bibRefCitation>
):
</paragraph>
<paragraph id="4A4C3804FFE3FFA52CAB9AFA9987E011" blockId="5.[151,1437,1356,2031]" pageId="5" pageNumber="456">
—From
<taxonomicName id="8DF34387FFE3FFA52D419AFA9905E1C9" authority="Iablokoff-Khnzorian, 1960" authorityName="Iablokoff-Khnzorian" authorityYear="1960" box="[301,839,1716,1743]" class="Insecta" family="Cupedidae" genus="Cupes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="rohdendorfi">
<emphasis id="7887E416FFE3FFA52D419AFA9846E1C9" box="[301,516,1716,1743]" italics="true" pageId="5" pageNumber="456">Cupes rohdendorfi</emphasis>
<bibRefCitation id="2E6245F5FFE3FFA52E7D9AFB9905E1C9" author="Iablokoff-Khnzorian, S. M." box="[529,839,1716,1742]" pageId="5" pageNumber="456" pagination="90 - 101" refId="ref4785" refString="Iablokoff-Khnzorian, S. M. (1960) Novye zhestkokrylye iz Baltiyskogo yantarya [New beetles from Baltic amber]. Paleontologicheskij Zhurnal, 1960 (3), 90 - 101. [in Russian]" type="journal article" year="1960">Iablokoff-Khnzorian, 1960</bibRefCitation>
</taxonomicName>
, by having only two paramedian pairs of suboval tubercles on the head dorsum, distinctly smaller eyes, much longer temples, a less elongate terminal antennomere, and the absence of a longitudinal division of the pronotoum by a furrow;
</paragraph>
<paragraph id="4A4C3804FFE3FFA52CAB9B6E9878E084" blockId="5.[151,1437,1356,2031]" pageId="5" pageNumber="456">
—From
<taxonomicName id="8DF34387FFE3FFA52D419B6E98FDE03C" authority="Kirejtshuk, 2005" authorityName="Kirejtshuk" authorityYear="2005" box="[301,703,1824,1851]" class="Insecta" family="Cupedidae" genus="Cupes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="komissari">
<emphasis id="7887E416FFE3FFA52D419B6E9BAFE03D" box="[301,493,1824,1851]" italics="true" pageId="5" pageNumber="456">Cupes komissari</emphasis>
<bibRefCitation id="2E6245F5FFE3FFA52D969B6F98FDE03C" author="Kirejtshuk, A. G." box="[506,703,1824,1851]" pageId="5" pageNumber="456" pagination="55 - 84" refId="ref4823" refString="Kirejtshuk, A. G. (2005) A revision of the genus Cupes Fabricius, 1801 from Baltic amber and some notes on taxonomy and composition of the family Cupedidae (Coleoptera, Archostemata). Mitteilungen aus dem Geologisch-Palaontologischen Institut an der Universitat Hamburg, 89, 55 - 84." type="journal article" year="2005">Kirejtshuk, 2005</bibRefCitation>
</taxonomicName>
, by the bi-colored scales on the elytra, only two paramedian prominent pairs of suboval tubercles on the head dorsum, smaller eyes, longer temples, and the absence of a median longitudinal furrow on the pronotum;
</paragraph>
<paragraph id="4A4C3804FFE3FFA52CAB9BC29887E0E8" blockId="5.[151,1437,1356,2031]" pageId="5" pageNumber="456">
—From
<taxonomicName id="8DF34387FFE3FFA52D459BC298FFE0A0" authority="Kirejtshuk, 2005" authorityName="Kirejtshuk" authorityYear="2005" box="[297,701,1932,1959]" class="Insecta" family="Cupedidae" genus="Cupes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="kerneggeri">
<emphasis id="7887E416FFE3FFA52D459BC29BB0E0A1" box="[297,498,1932,1959]" italics="true" pageId="5" pageNumber="456">Cupes kerneggeri</emphasis>
<bibRefCitation id="2E6245F5FFE3FFA52D979BC398FFE0A0" author="Kirejtshuk, A. G." box="[507,701,1932,1959]" pageId="5" pageNumber="456" pagination="55 - 84" refId="ref4823" refString="Kirejtshuk, A. G. (2005) A revision of the genus Cupes Fabricius, 1801 from Baltic amber and some notes on taxonomy and composition of the family Cupedidae (Coleoptera, Archostemata). Mitteilungen aus dem Geologisch-Palaontologischen Institut an der Universitat Hamburg, 89, 55 - 84." type="journal article" year="2005">Kirejtshuk, 2005</bibRefCitation>
</taxonomicName>
, by much more prominent protuberances on the head dorsum, a large scape as wide as the eye, a more narrowly elongate terminal antennomere, and a prontum with a rounded posterior edge with a weakly sinuate median part;
</paragraph>
<caption id="1E8C688CFFE0FFA62CFB9B3A9FD1E0D2" ID-DOI="http://doi.org/10.5281/zenodo.10910601" ID-Zenodo-Dep="10910601" httpUri="https://zenodo.org/record/10910601/files/figure.png" pageId="6" pageNumber="457" startId="6.[151,250,1909,1933]" targetBox="[176,1412,181,1885]" targetPageId="6" targetType="figure">
<paragraph id="4A4C3804FFE0FFA62CFB9B3A9FD1E0D2" blockId="6.[151,1437,1909,2005]" pageId="6" pageNumber="457">
<emphasis id="7887E416FFE0FFA62CFB9B3A9B53E08A" bold="true" box="[151,273,1909,1933]" pageId="6" pageNumber="457">FIGURE 4.</emphasis>
Morphological details of
<taxonomicName id="8DF34387FFE0FFA62E7C9B3A98E1E08A" authority="Yamamoto, 2024" authorityName="Yamamoto" authorityYear="2024" box="[528,675,1909,1933]" class="Insecta" family="Cupedidae" genus="Cupes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="balticus" status="sp. nov.">
<emphasis id="7887E416FFE0FFA62E7C9B3A98E1E08A" box="[528,675,1909,1933]" italics="true" pageId="6" pageNumber="457">Cupes balticus</emphasis>
</taxonomicName>
<emphasis id="7887E416FFE0FFA62EC49B3A989DE08A" bold="true" box="[680,735,1909,1933]" pageId="6" pageNumber="457">
<taxonomicNameLabel id="63B4596DFFE0FFA62EC49B3A989DE08A" box="[680,735,1909,1933]" pageId="6" pageNumber="457" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype, SEHU-0000121237.
<emphasis id="7887E416FFE0FFA6284C9B3A9E70E08A" bold="true" box="[1056,1074,1909,1933]" pageId="6" pageNumber="457">A</emphasis>
, Head, right antenna, and prothorax, ventral view.
<emphasis id="7887E416FFE0FFA62D4A9BD69B75E0B6" bold="true" box="[294,311,1945,1969]" pageId="6" pageNumber="457">B</emphasis>
, Forelegs, ventral view.
<emphasis id="7887E416FFE0FFA62E509BD6980CE0B6" bold="true" box="[572,590,1945,1969]" pageId="6" pageNumber="457">C</emphasis>
, Basal part of left antenna, ventral view.
<emphasis id="7887E416FFE0FFA6286A9BD69E5AE0B6" bold="true" box="[1030,1048,1945,1969]" pageId="6" pageNumber="457">D</emphasis>
, Mid- and hindlegs, ventral view.
<emphasis id="7887E416FFE0FFA629E99BD69FD4E0B6" bold="true" box="[1413,1430,1945,1969]" pageId="6" pageNumber="457">E</emphasis>
, Abdomen, ventral view. Abbreviations: a36, antennomeres 36; v15, ventrites 15. Scale bars: 1.0 mm (AD); 2.0 mm (E).
</paragraph>
</caption>
<paragraph id="4A4C3804FFE1FFA72CAB9CD89F49E7FE" blockId="7.[151,1436,150,645]" pageId="7" pageNumber="458">
—From
<taxonomicName id="8DF34387FFE1FFA72D4B9CD898D0E7B6" authority="Kirejtshuk, 2005" authorityName="Kirejtshuk" authorityYear="2005" box="[295,658,150,177]" class="Insecta" family="Cupedidae" genus="Cupes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="groehni">
<emphasis id="7887E416FFE1FFA72D4B9CD89B89E7B6" box="[295,459,151,177]" italics="true" pageId="7" pageNumber="458">Cupes groehni</emphasis>
<bibRefCitation id="2E6245F5FFE1FFA72DBE9CD898D0E7B6" author="Kirejtshuk, A. G." box="[466,658,150,177]" pageId="7" pageNumber="458" pagination="55 - 84" refId="ref4823" refString="Kirejtshuk, A. G. (2005) A revision of the genus Cupes Fabricius, 1801 from Baltic amber and some notes on taxonomy and composition of the family Cupedidae (Coleoptera, Archostemata). Mitteilungen aus dem Geologisch-Palaontologischen Institut an der Universitat Hamburg, 89, 55 - 84." type="journal article" year="2005">Kirejtshuk, 2005</bibRefCitation>
</taxonomicName>
, by smaller eyes and longer temples, a shorter and less elongate third antennomere compared to the second, a much less narrowly elongate ultimate antennomere (
<typeStatus id="954886A6FFE1FFA728FB9CF49EBFE7D2" box="[1175,1277,187,213]" pageId="7" pageNumber="458" type="paratype">paratype</typeStatus>
), an anteriorly projecting pronotal margin, a less elongate first protarsomere, and a less elongate first metatarsomere;
</paragraph>
<paragraph id="4A4C3804FFE1FFA72CAB9D4C9871E68E" blockId="7.[151,1436,150,645]" pageId="7" pageNumber="458">
—From
<taxonomicName id="8DF34387FFE1FFA72D409D4C98A0E61A" authority="Kirejtshuk, 2005" authorityName="Kirejtshuk" authorityYear="2005" box="[300,738,258,285]" class="Insecta" family="Cupedidae" genus="Cupes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="hoffeinsorum">
<emphasis id="7887E416FFE1FFA72D409D4C9850E61A" box="[300,530,259,285]" italics="true" pageId="7" pageNumber="458">Cupes hoffeinsorum</emphasis>
<bibRefCitation id="2E6245F5FFE1FFA72E729D4C98A0E61A" author="Kirejtshuk, A. G." box="[542,738,258,285]" pageId="7" pageNumber="458" pagination="55 - 84" refId="ref4823" refString="Kirejtshuk, A. G. (2005) A revision of the genus Cupes Fabricius, 1801 from Baltic amber and some notes on taxonomy and composition of the family Cupedidae (Coleoptera, Archostemata). Mitteilungen aus dem Geologisch-Palaontologischen Institut an der Universitat Hamburg, 89, 55 - 84." type="journal article" year="2005">Kirejtshuk, 2005</bibRefCitation>
</taxonomicName>
, by partially lighter scales, smaller eyes, longer temples, a vertex with a median longitudinal furrow, a shorter and less elongate third antennomere compared to the second, a much less elongate terminal antennomere, a pronotum wider than the head at temples, and a much shorter fifth metatarsomere compared to the first;
</paragraph>
<paragraph id="4A4C3804FFE1FFA72CAB9DDC9A8EE53A" blockId="7.[151,1436,150,645]" pageId="7" pageNumber="458">
—From
<taxonomicName id="8DF34387FFE1FFA72D449DDC98A6E6AA" authority="(Motschoulsky, 1856)" baseAuthorityName="Motschoulsky" baseAuthorityYear="1856" box="[296,740,402,429]" class="Insecta" family="Cupedidae" genus="Cupes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="tesselatus">
<emphasis id="7887E416FFE1FFA72D449DDC9BA1E6AA" box="[296,483,403,429]" italics="true" pageId="7" pageNumber="458">Cupes tesselatus</emphasis>
(Motschoulsky, 1856)
</taxonomicName>
, by the uniformly brownish head and pronotum, smaller eyes with longer temples, a shorter and less elongate third antennomere compared to the second, a more elongate terminal antennomere, a wider pronotum compared to the head at temples, a curved posterior pronotal edge weakly protruding medially, a more narrowly elongate first metatarsomere, and a much shorter fifth metatarsomere compared to the first.
</paragraph>
<paragraph id="4A4C3804FFE1FFA72CAB9E0898E7E582" blockId="7.[151,1436,150,645]" pageId="7" pageNumber="458">Therefore, the specimen described here differs distinctly from any cupedids previously described from Baltic amber and is thus recognized as a new species.</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,72 @@
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<mods:title id="2FA7B8BC70F8336F5764B584EF6DC000">Biodiversity Data Journal</mods:title>
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<mods:part id="4682D85A911488759B4C3EF2DD57BB4B">
<mods:date id="4D8FE3D748F345B092EE28FDB5787F78">2016</mods:date>
<mods:detail id="D87935EBBE52E16472589CEF5AE295B0" type="volume">
<mods:number id="AEF195CDA1C2651FFF83592311B79079">4</mods:number>
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<mods:start id="D151CBEB7993810EFE076613E0419A54">8013</mods:start>
<mods:end id="AA4653EB25AAF23DB22A84E6FB945CE3">8013</mods:end>
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<mods:location id="5FA8A337F2B9480CE313C7BC87338FA5">
<mods:url id="20549565DCA4AF8F8E553A6A452BD784">http://dx.doi.org/10.3897/BDJ.4.e8013</mods:url>
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<mods:classification id="984CC1012DE0D013818CC21C9DDBB8C8">journal article</mods:classification>
<mods:identifier id="10E83E491113353C2ABBEC7E49E48BFB" type="DOI">http://dx.doi.org/10.3897/BDJ.4.e8013</mods:identifier>
<mods:identifier id="3C6A477B59F5D44DC409A1BFF0D28A34" type="Pensoft-Pub">1314-2828-4-8013</mods:identifier>
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<taxonomicName id="DC5ABF155771F15F0524DC53179D955D" ID-CoL="62L74" authority="Walker, 1846" authorityName="Walker" authorityYear="1846" class="Insecta" family="Mymaridae" genus="Caraphractus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caraphractus" order="Hymenoptera" pageId="0" pageNumber="8013" phylum="Arthropoda" rank="genus">Caraphractus Walker, 1846</taxonomicName>
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<taxonomicName id="89D5CA8C451B47C142303FA7DEB4DE68" class="Insecta" family="Mymaridae" genus="Valkerella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="VALKERELLA" order="Hymenoptera" pageId="0" pageNumber="8013" phylum="Arthropoda" rank="genus">VALKERELLA</taxonomicName>
Westwood, 1879
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