diff --git a/data/03/D6/87/03D687B8FF92FFE6FDD8F9FDFA8FF976.xml b/data/03/D6/87/03D687B8FF92FFE6FDD8F9FDFA8FF976.xml index 10b216a2854..e741006e144 100644 --- a/data/03/D6/87/03D687B8FF92FFE6FDD8F9FDFA8FF976.xml +++ b/data/03/D6/87/03D687B8FF92FFE6FDD8F9FDFA8FF976.xml @@ -1,57 +1,65 @@ - - - -Two new species of the planthopper genus Eponisiella Emeljanov from China (Hemiptera, Fulgoromorpha, Meenoplidae) + + + +Two new species of the planthopper genus Eponisiella Emeljanov from China (Hemiptera, Fulgoromorpha, Meenoplidae) - - -Author + + +Author -Yang, Lin -17FAF564-8FDA-4303-8848-346AB8EB7DE4 -Institute of Entomology, Guizhou University, Guiyang, Guizhou 550025, P. R. China. & The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University, Guiyang, Guizhou 550025, P. R. China. -yanglin6626@163.com +Lv, Sha-sha +4E2050EC-EB1E-4A61-8761-BFB96DB5152D +Institute of Entomology, Guizhou University, Guiyang, Guizhou 550025, P. R. China. & The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University, Guiyang, Guizhou 550025, P. R. China. +ls13116322648@163.com - - -Author + + +Author -Chen, Xiang-Sheng -D9953BEB-30E6-464A-86F2-F325EA2E4B7C -Institute of Entomology, Guizhou University, Guiyang, Guizhou 550025, P. R. China. & The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University, Guiyang, Guizhou 550025, P. R. China. -chenxs3218@163.com +Yang, Lin +17FAF564-8FDA-4303-8848-346AB8EB7DE4 +Institute of Entomology, Guizhou University, Guiyang, Guizhou 550025, P. R. China. & The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University, Guiyang, Guizhou 550025, P. R. China. +yanglin6626@163.com -text - - -European Journal of Taxonomy + + +Author + +Chen, Xiang-Sheng +D9953BEB-30E6-464A-86F2-F325EA2E4B7C +Institute of Entomology, Guizhou University, Guiyang, Guizhou 550025, P. R. China. & The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University, Guiyang, Guizhou 550025, P. R. China. +chenxs3218@163.com + +text + + +European Journal of Taxonomy - -2021 - -2021-09-14 + +2021 + +2021-09-14 - -767 + +767 - -83 -93 + +83 +93 - -http://dx.doi.org/10.5852/ejt.2021.767.1489 + +http://dx.doi.org/10.5852/ejt.2021.767.1489 -journal article -4207 -10.5852/ejt.2021.767.1489 -8872c843-c80e-4644-a9d4-71ebf26a550b -2118-9773 -5514865 -02EA124E-D4F6-4237-95F1-A1816A54BFC1 +journal article +10.5852/ejt.2021.767.1489 +8872c843-c80e-4644-a9d4-71ebf26a550b +2118-9773 +5514865 +02EA124E-D4F6-4237-95F1-A1816A54BFC1 - + @@ -77,7 +85,7 @@ Genus - + Eponisiella – diff --git a/data/08/2A/87/082A87C1FFD16C3A80176A809739FB9E.xml b/data/08/2A/87/082A87C1FFD16C3A80176A809739FB9E.xml new file mode 100644 index 00000000000..175db8d973c --- /dev/null +++ b/data/08/2A/87/082A87C1FFD16C3A80176A809739FB9E.xml @@ -0,0 +1,714 @@ + + + +Review of the genus Chinoperla Zwick, 1980 (Plecoptera: Perlidae: Perlinae) from China + + + +Author + +Mo, Rao-Rao +9F30237C-B493-4A90-A545-EDB2EDF11BF2 +Guangxi key laboratory of Agric-Environment and Agric-Products Safety and National Demonstration Center for Experimental Plant Science Education, Agricultural College, Guangxi University, Nanning, Guangxi 530004, China. & Department of Plant Protection, Henan Institute of Science and Technology, Xinxiang, Henan 453003, China. +1689832746@qq.com + + + +Author + +Liu, Rui-Jun +1C00450A-AAE4-4F41-8E9D-11FB7A1E05FE +Henan Institute of Science and Technology Herbarium, Xinxiang, Henan 453003, China. +histlruijun@163.com + + + +Author + +Wang, Guo-Quan +2FAFA732-C8F7-4823-949A-EF81D6B6A2F4 +Guangxi key laboratory of Agric-Environment and Agric-Products Safety and National Demonstration Center for Experimental Plant Science Education, Agricultural College, Guangxi University, Nanning, Guangxi 530004, China. +wangguoquan0@163.com(GQW);lwh7969@163.com(WHL) + + + +Author + +Li, Wei-Hai +81D57DE5-E00B-4F6B-A0B3-E81D3C865208 +Department of Plant Protection, Henan Institute of Science and Technology, Xinxiang, Henan 453003, China. +wangguoquan0@163.com(GQW);lwh7969@163.com(WHL) + + + +Author + +Murányi, Dávid +86C43175-8A78-44EB-A505-19553C6246CA +Department of Zoology, Eszterházy Károly University, Leányka u. 6, Eger H- 3300, Hungary. +d.muranyi@gmail.com + +text + + +European Journal of Taxonomy + + +2021 + +2021-10-19 + + +775 + + +62 +85 + + + +journal article +10.5852/ejt.2021.775.1547 +41dd2703-d6ec-4d78-9b3d-c06ff2ddc23f +2118-9773 +5578472 +1EB4BD2A-CDC8-4383-B693-CF2A089F85D5 + + + + + + +Chinoperla nigrifrons +( +Banks, 1939 +) + + + + + + +Figs 6–10 + + + + + + + +Ochthopetina nigrifrons +Banks, 1939: 449 + + +, fig. 73 (original description). + + + + + + +Sinoperla furcomacula +Wu, 1973: 110 + + +, figs 91–93 (original description). + + + + + +Ochthopetina nigrifrons + +– + +Illies 1966: 274 + +(catalog). + + + + + +Chinoperla furcomacula + +– + +Zwick & Sivec 1980: 130 + +(transferred from +Sinoperla +). — + +Zwick 1982: 168 + +(synopsis). — + + +Sivec +et al. +1988: 54 + + +(provisionally possible conspecifity with + +C. nigrifrons + +). — + +Sivec & Zwick 1989: 14 + +(three theories about possible synonymy or distinctiveness with + +C. nigrifrons + +). + + + + + +Chinoperla nigrifrons + +– + + +Sivec +et al. +1988: 54 + + +(transferred from +Ochthopetina +, provisionally possible conspecifity with + +C. furcomacula + +). — + +Sivec & Zwick 1989: 14 + +, fig. 3a, c–e (redescription of male with uneverted aedeagus and female based on types). — + + +Du +et al. +1999: 62 + + +(catalog). — + +Yang & Li 2018: 36 + +(catalog). + + + + + + +Diagnosis + + +Head with a heart-shaped dark brown patch and a subtriangular dark brown spot. Male tergum 9 with a trumpet-shaped median sclerite. Hemitergal processes medially indented in lateral view. Aedeagal tube with a T-shaped sclerotized dorsal marking. Aedeagal sac dorsoapically with a low membranous median lobe; ventrobasally with a large patch of spinules; ventroapically with one long and one shorter spine, or the two of equal length; apically with a long straight spine and a large membranous lobe; laterally with dense large, curved spines arranged on each side. Posterior margin of female sternum 8 slightly produced, forming a small subtriangular subgenital plate with an apical notch. + + + + +Material examined + + + + + +Holotype + +CHINA +• +♂ +; “Hainan, +Ta Han +” [ +Hainan Province +, +Qiongzhong Li and Miao Autonomous County +, located in +Limushan Town +]; + +23 Jun. 1935 + +; +L. Gressitt +leg.; +MCZ +Type 22693. + + + + + +Paratypes + +CHINA +• +1 ♀ +; same locality data as for holotype; + +2 Aug. 1935 + +; +L. Gressitt +leg.; +MCZ + +• + +1 ♀ +; same locality data as for holotype; + +13 Jul. 1935 + +; +L. Gressitt +leg.; +MCZ + +. + + +Other material + + + +CHINA +• +1 ♂ +, +1 ♀ +(holotype and paratype of + +S. furcomacula + +); “ +Kwangtung +, +Hainan +, +Tung-shih +” [ +Hainan Province +belonging to Guangdong at that time, +Wuzhishan City +, +Tongshen Town +]; alt. + +350 m + +; + +27 May 1960 + +; +IZCAS + +• + +2 ♂♂ +; +Hainan Province +, +Baisha Li Autonomous County +, +Mount Yinggeling +; +19°14′0.24″ N +, +109°26′27.96″ E +; alt. + +430 m + +; + +1 Apr. 2013 + +; +Yu-Yu Wang +and +Yun-Lan Jiang +leg.; +HIST + +. + + + + +Fig +. +6 +. + +Chinoperla nigrifrons +( +Banks, 1939 +) + +, ♂, holotype (MCZ Type 22693). +A +. Adult habitus, dorsal view. +B +. Head and pronotum, dorsal view. +C +. Terminalia, dorsal view. +D +. Cleared terminalia, dorsal view. +E +. Cleared terminalia, lateral view. Abbreviations: see Material and methods. Scale bars: A = 1.0 mm; B–E = 0.5 mm. + + + + + +Redescription + + + +Male + + +ADULT HABITUS ( +Figs 6A–B +, +8C +, +9A +). General color brown. Forewing length 7.7–8.0 mm, hindwing length +6.7–7.1 mm +. Head brownish, with a dark brown heart-shaped spot, but specimens from Mount Yinggeling with a long brown band located on the occiput, and with a subtriangular dark brown spot in front of M-line; biocellate, ocelli about 3 diameters apart of each other, slightly closer to each other than to the compound eye; antennae brown. Pronotum trapezoidal, brown with paler lateral margin and with darker, distinct rugosities; anterior corners pointed but posterior corners obtuse. Legs generally brown, proximal part of femora mostly lighter; wings subhyaline and brownish, veins brown, Sc of forewing short, typical of the genus ( +Fig. 8C +); cerci brownish. + + + +Fig. 7. + +Chinoperla nigrifrons +( +Banks, 1939 +) + +, ♂, holotype (MCZ Type 22693). +A +. Aedeagus, dorsal view. +B +. Aedeagus, lateral view. +C +. Aedeagal sac, ventral view. +D +. Apex of aedeagal sac, oblique ventral view. Abbreviations: see Material and methods. Scale bars = 0.25 mm. + + + +ABDOMEN ( +Figs 6C–E +, +8A–B +, +9B–E +). Sterna 3–5 with hair brushes. Dark median sclerite of tergum 9 trumpet-shaped, about half as long as the segment, heavily sclerotized, distinctly forked distally with two sharp points. Hemitergal processes slender, about 2.5× as long as basal width, distinctly narrowing subapically; medially indented in lateral view; basal callus fully covered by sensilla basiconica. + + +AEDEAGUS ( +Figs 7 +, +10 +). Aedeagal tube partially sclerotized dorsally, forming a distinctly T-shaped marking on the +holotype +( +Fig. 7A–B +), but marking in specimens from Mount Yinggeling indistinct, well seen only in lateral view ( +Fig.10A–B +). Fully everted sac as long as the tube, curved dorsad; ventroapical and dorsoapical surface heart-shaped in specimens from Mount Yinggeling ( +Fig. 10 +), less definite in the +holotype +( +Fig. 7 +); dorsoapical surface with a small bulgy membranous median lobe; a single large spine about as long as ¼ aedeagus length, and a large membranous lobe medially at apex, slightly directed dorsally; ventral surface with a large basal patch of spinule, and one long and one shorter subapical spines ( +Fig. 7 +) but in fresh specimens from Mount Yinggeling the two of approximately equal length ( +Fig. 10 +); lateroapical part with dense, large and curved spines arranged on each side. + + + + +Fig. 8. + +Chinoperla nigrifrons +( +Banks, 1939 +) + +. +A–B +. ♂, holotype of + +C. furcomacula +( +Wu, 1973 +) (IZCAS) + +. +A +. Cleared terminalia, dorsal view. +B +. Cleared terminalia, lateral view. — +C +. ♂ from Mount Yinggeling (HIST), forewing. Abbreviations: see Material and methods. Scale bars = 0.5 mm. + + + + + +Distribution + + + +China +: +Hainan Province +, found in three counties. + + + + +Fig. 9. + +Chinoperla nigrifrons +( +Banks, 1939 +) + +, ♂ from Mount Yinggeling (HIST). +A +. Head and pronotum, dorsal view. +B +. Terminalia, dorsal view. +C +. Cleared terminalia, dorsal view. +D +. Cleared terminalia, lateral view. +E +. Cleared terminalia, oblique lateral view. Abbreviations: see Material and methods. Scale bars = 0.5 mm. + + + + + +Remarks + + + +We carefully compared the type specimens of + +O. nigrifrons + +deposited in MCZ with the illustrations in +Sivec & Zwick (1989) +. The head pattern, direction of antennae and shape of the pronotum of the female +paratype +collected on +13 July 1935 +are consistent with those illustrated in +Sivec & Zwick (1989 +: fig. 3a, d–e). The figures of the terminalia and vagina also refer to the same female +paratype +, because the second female +paratype +collected on +2 August 1935 +had not been dissected, and its sternum 9 is more narrow. The figure of the male terminalia in +Sivec & Zwick (1989 +: fig. 3b) shows a shallow apical notch of the median sclerite of tergum 9, and tapering and curving hemitergal processes without a straight apical half. These characters are consistent with those in the male +paratype +, that is not conspecific with the +holotype +and has been reassigned to + +C. changjiangensis + +sp. nov. +above ( +Fig. 3B–E +). However, the small basal sclerite in the aedeagus of the +holotype +( +Fig. 7B +) corresponds to that illustrated in +Sivec & Zwick (1989 +: fig. 3c). Therefore, we believe that the half-everted aedeagus of the +holotype +, the terminalia of the male +paratype +( + +C. changjiangensis + +sp. nov. +), and the head, pronotum, sterna 7–9 and vagina of the female +paratype +of + +Chinoperla nigrifrons + +were redescribed by +Sivec & Zwick (1989) +. Color photographs and illustrations of the male +holotype +are provided in the present study ( +Figs 6–7 +). The characteristics of both female +paratypes +agree well with the description and illustrations provided by +Sivec & Zwick (1989) +. + + + +Fig. 10. + +Chinoperla nigrifrons +( +Banks, 1939 +) + +, ♂ from Mount Yinggeling (HIST). +A +. Aedeagus, lateral view. +B +. Aedeagus, dorsal view. +C +. Aedeagus, ventral view. Abbreviations: see Material and methods. Scale bars = 0.25 mm. + + + +We checked the types of + +S. furcomacula + +in IZCAS, but the aedeagus of + +S. furcomacula + +with the +holotype +label was difficult to fully evert due to the poor condition of the specimen. Based on the original description and redescription of + +C. nigrifrons + +in the present study, we believe that the +holotype +of + +S. furcomacula + +is conspecific with the +holotype +of + +O. nigrifrons + +, and is therefore a junior synonym of + +C. nigrifrons + +( +Banks 1939 +; +Wu 1973 +; + +Sivec +et al. +1988 + +; +Sivec & Zwick 1989 +). The two recently collected males from +Hainan Province +examined in this study agree well with the type specimens in general characteristics such as head pattern, lateral aspect of the hemitergal lobes, and armatures of the aedeagus. However, these fresh specimens from Mount Yinggeling show slight differences in the head pattern and aedeagus, as they have a long brown posterior band on the head, and the two ventroapical spines of the aedeagal sac are straight and nearly equal in length. We believe this brown occipital band disappears in fully aged adults, such as the +holotype +. Similar differences were documented in the case of + +Kamimuria guangxia +Li & +Wang, 2013 + +and + +Neoperla lihuae +Li & Murányi, 2014 + +( + +Wang +et al. +2013 + +; + +Li +et al. +2014 + +). The differences in the aedeagal armatures can be regarded as intraspecific differences. We provide illustrations of the fully everted sac of the + +C. nigrifrons + +holotype +and the fresh adults to allow a reliable identification for future studies ( +Figs 7 +, +10 +). + + +Unfortunately, the +paratype +female of + +S. furcomacula + +in IZCAS is badly damaged and difficult to study. According to the original description and figures of + +S. furcomacula + +( +Wu 1973 +: fig. 93), the terminalia of the +paratype +female distinctly differ from those of + +C. nigrifrons + +by the unmodified subgenital plate. +Sivec & Zwick (1989) +drew up three possibilities for the specific identity of this female; we support the third, which suggests that the +paratype +female may not be conspecific with the +holotype +of + +S. furcomacula + +. Further specimens from +Hainan +are needed to confirm the identity of the female. + + + + \ No newline at end of file diff --git a/data/08/2A/87/082A87C1FFD76C3A806B6AFB9708F8C5.xml b/data/08/2A/87/082A87C1FFD76C3A806B6AFB9708F8C5.xml new file mode 100644 index 00000000000..184b2782b02 --- /dev/null +++ b/data/08/2A/87/082A87C1FFD76C3A806B6AFB9708F8C5.xml @@ -0,0 +1,204 @@ + + + +Review of the genus Chinoperla Zwick, 1980 (Plecoptera: Perlidae: Perlinae) from China + + + +Author + +Mo, Rao-Rao +9F30237C-B493-4A90-A545-EDB2EDF11BF2 +Guangxi key laboratory of Agric-Environment and Agric-Products Safety and National Demonstration Center for Experimental Plant Science Education, Agricultural College, Guangxi University, Nanning, Guangxi 530004, China. & Department of Plant Protection, Henan Institute of Science and Technology, Xinxiang, Henan 453003, China. +1689832746@qq.com + + + +Author + +Liu, Rui-Jun +1C00450A-AAE4-4F41-8E9D-11FB7A1E05FE +Henan Institute of Science and Technology Herbarium, Xinxiang, Henan 453003, China. +histlruijun@163.com + + + +Author + +Wang, Guo-Quan +2FAFA732-C8F7-4823-949A-EF81D6B6A2F4 +Guangxi key laboratory of Agric-Environment and Agric-Products Safety and National Demonstration Center for Experimental Plant Science Education, Agricultural College, Guangxi University, Nanning, Guangxi 530004, China. +wangguoquan0@163.com(GQW);lwh7969@163.com(WHL) + + + +Author + +Li, Wei-Hai +81D57DE5-E00B-4F6B-A0B3-E81D3C865208 +Department of Plant Protection, Henan Institute of Science and Technology, Xinxiang, Henan 453003, China. +wangguoquan0@163.com(GQW);lwh7969@163.com(WHL) + + + +Author + +Murányi, Dávid +86C43175-8A78-44EB-A505-19553C6246CA +Department of Zoology, Eszterházy Károly University, Leányka u. 6, Eger H- 3300, Hungary. +d.muranyi@gmail.com + +text + + +European Journal of Taxonomy + + +2021 + +2021-10-19 + + +775 + + +62 +85 + + + +journal article +10.5852/ejt.2021.775.1547 +41dd2703-d6ec-4d78-9b3d-c06ff2ddc23f +2118-9773 +5578472 +1EB4BD2A-CDC8-4383-B693-CF2A089F85D5 + + + + + + +Chinoperla nigroflavata +( +Wu, 1948 +) + + + + + + + + + +Sinoperla nigroflavata +Wu, 1948: 78 + + +, figs 8–9 (original description of the male). + + + + + +Sinoperla nigroflavata + +– + +Illies 1966: 274 + +(catalog). + + + + + +Chinoperla nigroflavata + +– Zwick 1980: 130 (transferred from +Sinoperla +). — + +Zwick 1982: 167 + +(synopsis). — + + +Sivec +et al. +1988: 54 + + +(synopsis). — + + +Du +et al. +1999: 62 + + +(catalog). — + +Yang & Li 2018: 36 + +(catalog). + + + + + + +Distribution + + + +China +: +Fujian Province +, known only from the (now lost) +holotype +. + + + + + +Remarks + + + +The details of the aedeagal structures of this species are still unknown and the type material is lost ( +Wu 1948 +; + +Du +et al. +1999 + +; +Yang & Li 2018 +). No new material was available for the present study, but it seems to differ from the remaining Chinese species of + +Chinoperla + +based on the absence of the short Sc of the forewing and the shape of the dark median sclerite of tergum 9 ( +Wu 1948 +: figs 8–9). + +Chinoperla gorohovi +Sivec & Stark, 2010 + +from Guangdong and +Vietnam +is easily distinguishable from + +C. nigroflavata + +by the slender nail-shaped median sclerite with a slightly enlarged apex ( +Fig. 11B–D +). + + + + \ No newline at end of file diff --git a/data/BE/1A/87/BE1A87C3BC53BC1DFF1FFDA6FAE8F8D5.xml b/data/BE/1A/87/BE1A87C3BC53BC1DFF1FFDA6FAE8F8D5.xml new file mode 100644 index 00000000000..12f653fbbfe --- /dev/null +++ b/data/BE/1A/87/BE1A87C3BC53BC1DFF1FFDA6FAE8F8D5.xml @@ -0,0 +1,560 @@ + + + +Three new species of tiny tarantulas of the genus Thalerommata Ausserer, 1875 (Araneae, Theraphosidae) from northern Colombia + + + +Author + +Osorio, Luis M. +Grupo de Investigación Biodiversidad del Caribe Colombiano, Facultad de Ciencias Básicas, Universidad del Atlántico, Barranquilla, Colombia + + + +Author + +Benavides, Léiner +Grupo de Investigación en Insectos Neotropicales. Universidad del Magdalena, Santa Marta Colombia + + + +Author + +García-Atencia, Sandy +Grupo de Investigación Biodiversidad del Caribe Colombiano, Facultad de Ciencias Básicas, Universidad del Atlántico, Barranquilla, Colombia + + + +Author + +Bertani, Rogério +Laboratório de Ecologia e Evolução, Instituto Butantan. 05503 - 900 São Paulo, SP, Brazil + +text + + +Zootaxa + + +2024 + +2024-11-07 + + +5537 + + +1 + + +95 +114 + + + + +http://dx.doi.org/10.11646/zootaxa.5537.1.5 + +journal article +10.11646/zootaxa.5537.1.5 +1175-5326 +14239357 +5AD70C1C-9942-471A-857B-D082C17C4F4D + + + + + + + +Thalerommata margarita + +sp. nov. + + + + + + +( +Figs 9–12 +, +Fig. 13F +). + + + + +Type material. + +Holotype +♂ +from +Colombia +: + +Magdalena +: + +Santa Marta +, +Minca +, +Vda. Bella Vista +, +ground hand collecting +, subtropical humid forest ( +11°06’11.6”N +, +74°04’18.1”W +), + +1880 m + +.a.s.l, + +06 May 2023 + +, +L. Benavides +& +E. Bolaño +leg. ( +ICN-Ar12996 +) + +. + +Paratype +1 ♀ +, +2 immatures +, same data as holotype ( +ICN-Ar12997 +) + +. + + + + +Diagnosis. +Males of + +T. margarita + + +sp. nov. + +resemble those of + +T. kogui + + +sp. nov. + +in having a curved metatarsus I with a protuberance in the distal prolateral region ( +Fig. 9D, E +) and + +T. anae + +in the general aspect of the bulb ( +Fig. 10A, B +). They can be distinguished from most of their congeners by having the tibia of the palpus elongate, not enlarged in the basal half, absence of spiniform setae and embolus more than twice the length of the tegulum ( +Fig. 10A–E +) and of + +T. anae + +by having a curved metatarsus I with a protuberance in the distal prolateral region, tibial apophysis shape and embolus strongly curved at its middle half ( +Fig. 9D +, +Fig. 10 A, B +). Females resemble + +T. maculata + +, + +T. caudicula +, +T. yukpa + + +sp. nov. + +and + +T. kogui + + +sp. nov. + +by having the distal article of PLS digitiform ( +Fig. 11D +). They differ from all of the above species in having elongate spermathecae, thickened at their base and the absence of distal lobes ( +Fig. 12 +). + + + + +Etymology. +The specific epithet comes from the Latin +“margarita +,” which translates to “Perla” in Spanish. This term, in turn, is an appellation given to the city of Santa Marta “La Perla de América”. + + + + +Description. +Male +holotype +(ICN-Ar12996). Carapace 6.81 long, 5.97 wide. Abdomen 6.64 long, 3.53 wide. Total length 15.27. Carapace: fovea straight 0.76 wide ( +Fig. 9A +). Clypeus absent. Eight eyes arranged on tubercle 1.28 wide, 0.78 long. MOQ 0.69 wide, 0.56 long. Anterior eye row slightly procurved, posterior row slightly recurved ( +Fig. 9A +). AME 0.25, ALE 0.42, PME 0.18, PLE 0.33. Eye interspaces: AME–AME 0.14, AME–ALE 0.07, ALE–ALE 0.63, PME–PLE 0.04, PME–PME 0.53, ALE–PLE 0.12, PLE–PLE 0.95. Chelicerae: 1.82 long, with 12 large, spaced teeth and numerous tiny on basal inner edge. Intercheliceral intumescence absent. Rastellum composed of long thick setae. Labium ellipsoid, 0.65 long, 1.00 wide, with 44 cuspules rounded on anterior half ( +Fig. 9B +). Labiosternal groove shallow, with a pair of sigilla ( +Fig. 9B +). Maxillae ( +Fig. 9B +): 1.90 long in front, 2.54 long behind, 1.20 wide, with +ca +. 56 cuspules on upper mound on inner angle. Heel distinct. Frontal lobe distinct, short. Prolateral edge whitish: ( +Fig. 9B +). Sternum ( +Fig. 9B +): 2.76 long, 2.81 wide. Three pairs of sigilla, rounded. First pair close to coxa I, second pair close to coxa II, and third pair close to coxa III. All sigilla distant, about one diameter from margin. Lyra and stridulatory apparatus absent. Abdomen: booklungs small with elliptical aperture +ca +. +0.70 in +length, booklung combs absent. Spinnerets: PMS small, 0.55 long, 0.25 wide, 0.11 apart. PLS, basal segment, 0.73 long, 0.45 wide; middle 0.56 long, 0.41 wide; apical 0.76 long, 0.34 wide. Apical segment digitiform. Legs: IV–I–III–II. Legs (femur, patella, tibia, metatarsus, tarsus, total): I: 5.57, 3.50, 5.48, 3.97, 3.12, 21.64; II: 5.43, 3.19, 4.52, 4.28, 2.84, 20.26; III: 5.07, 2.52, 3.58, 4.76, 2.74, 18.67; IV: 6.57, 2.78, 5.27, 6.89, 2.90, 24.41; Palp: 2.46, 2.15, 2.96, -, 1.10, 8.67. Widths: I: 1.30, 1.38, 1.34, 0.73, 0.69; II: 1.26, 1.29, 0.77, 0.74, 0.52; III: 1.43, 1.25, 1.07, 0.70, 0.56; IV: 1.24, 1.12, 1.04, 0.61, 0.52; Palp: 1.14, 1.11, 1.33, -, 0.41. Spines: tarsi of legs lacking spines. Leg I: fe p0-0-2 r0-0-2, pa 0, ti p 0-1-2 v2-2-2, me p1-r1; leg II: fe p0-1-3, pa 0, ti p1-1-0 v2-1-3ap, me p0-1-0, v1-1- 2ap; leg III: fe d3-4-2, pa0, ti p1-1-1 r1-1-1 v2-2-3ap, me v4-3-3ap p1-1-1 r2-2-1ap; leg IV: fe d1-4-2, pa0, ti p1-2-1 r1-2-2 v3-3-3ap, me v3-4-3ap r1-1-1 p2-2-1; palp: fe 0, pa 0, ti p0-2-1 r0-0-1ap; cymbium 12 closely positioned at the apex. PC absent. Clavate trichobothria in two rows, on distal half: 10 on tarsus I; 7 on tarsus II, 14 on tarsus III; 19 on tarsus IV; 15 on palpal tarsus. Claws: STC with double row of 7–7 teeth on leg I; 8–7 on leg II; 7–7 on leg III; 8–8 on leg IV. Scopula: sparse on tarsi of legs I–IV; divided by broad row of setae on tarsi III–IV. Metatarsi I–III scopulate on apical third. Tarsi III–IV cracked. Tibial apophysis ( +Fig. 9D, E +): Upper and lower process roughly at same level. Small lower process with small contiguous sinuous spine, two times process length. Upper process large, curved, with curved spine roughly same length of process. Very short sinuous upper spine, and more basal, long and sinuous spine, twice as long as short spine, positioned on broad and strong base. Both structures on prolateral face of tibia. Tibia I excavated prolaterally from area of apophysis to its tip. Metatarsus I curved, with small prolateral bulge ( +Fig. 9D, E +), when folded it passes retrolaterally by apophysis. Palp ( +Fig. 10A–E +): Bulb pyriform, with embolus that tapers abruptly in middle portion to its tip, with slight curvature to prolateral side. Tegulum 0.49 long, 0.20 wide, embolus 2.64 long, 0.31 wide at its base, 0.09 at middle, 0.03 at tip. Cymbium triangular in retrolateral view, not elongated, with deep incision. Palpal tibia with narrow excavation, spiniform setae absent. Color pattern (preserved in alcohol) ( +Fig. 9A, C +): Carapace, chelicerae and legs brown, abdomen darker with ten whitish oval spots dorsally. Setae Type II ( +Ríos-Tamayo, 2024 +) only on abdomen, occurring both dorsally and ventrally. + + +Description. +Female +paratype +(ICN-Ar 12997). Carapace 7.20 long, 6.45 wide.Abdomen 12.85 long, 8.16 wide. Total length 23.33. Carapace: Fovea straight 1.18 wide ( +Fig. 11A +). Clypeus absent. Eight eyes arranged on tubercle 1.39 wide, 0.93 long. MOQ 0.93 wide, 0.67 long. Anterior eye row straight, posterior row slightly recurved ( +Fig. 11A +). AME 0.22, ALE 0.39, PME 0.28, PLE 0.36. Eye interspaces: AME–AME 0.18, AME–ALE 0.10, ALE–ALE 0.77, PME–PLE 0.07, PME–PME 0.60, ALE–PLE 0.13, PLE–PLE 1.05. Chelicerae: 3.28 long, with 12 spaced teeth and numerous very small denticles on its base. Rastellum composed of long thick setae. Labium ellipsoid, 0.94 long, 1.37 wide, having 67 cuspules rounded, on the distal half ( +Fig. 11B +). Labiosternal groove shallow, with pair of sigilla ( +Fig. 11B +). Maxillae ( +Fig. 11B +): 2.28 long in front, 3.52 long behind, 1.56 wide, with ca. 60 cuspules on upper mound in inner angle. Heel distinct. Frontal lobe distinct, short. Prolateral edge whitish. Sternum ( +Fig. 11B +): 3.27 long, 3.63 wide. Three pairs of sigilla, rounded. First pair close to coxa I, second pair close to coxa II, and third pair close to coxa III. All sigilla distant, about one diameter from margin. Lyra and stridulatory apparatus absent. Abdomen: booklungs small with elliptical aperture +ca. +0.81 in +length, with booklung combs hardly visible. Spinnerets ( +Fig. 11D +): PMS small, 0.84 long, 0.43 wide, 0.51 apart. Basal, middle, and apical segments of PLS, 1.37 long, 0.70 wide; 0.90 long, 0.68 wide; 1.27 long, 0.43 wide, respectively; apical segment digitiform. Legs: formula IV–I–II–III. Legs (femur, patella, tibia, metatarsus, tarsus, total): I: 4.98, 3.54, 3.66, 2.71, 1.91, 16.8; II: 4.56, 2.98, 3.29, 2.77, 2.24, 15.94; III: 4.17, 2.62, 2.60, 3.13, 2.16, 14.68; IV: 5.48, 3.01, 4.08, 4.86, 2.54, 19.97. Palp: 3.73, 2.71, 2.26, -, 2.42, 11.12. Widths: I: 1.12, 1.44, 1.46, 1.02, 0.92; II: 1.38, 1.28, 1.33, 0.96, 0.94; III: 1.72, 1.38, 1.40, 0.97, 0.77; IV: 1.32, 1.40, 1.35, 0.99, 0.83. Palp: 0.97, 1.24, 1.29, -, 0.94. Spines: leg I: fe:0, pa:0, ti0, me v1-0-1ap; leg II: fe0, pa0, ti0, me v1-0-1ap; leg III: fe0, pa0, ti p1-1-0 r0-0-1 v0-0-2ap, me p1-0-0 r0-2-2 v2-2-3ap; leg IV: fe0, pa0, ti p0 r0-1-1 v2-2-3ap, me p0-1-1 v2-2-3ap r0-1-1; palp: 0. PC absent. Clavate trichobothria in two rows on distal two thirds: 17 on tarsus I; 22 on tarsus II; 17 on tarsus III; 23 on tarsus IV; 16 on palpal tarsus. Claws: STC bare on all legs; palpal claw single, bare. Tarsi I–IV scopulate, divided by broad row of setae, metatarsus I–IV apically scopulate, on IV inconspicuous. Tarsi IV cracked. Spermathecae ( +Fig. 12 +): two, each with thick peduncle at base that tapers abruptly as it curves inward, with prolateral lobe. Color pattern (preserved in alcohol) ( +Fig. 11A–C +): as in male. Setae Type II ( +Ríos-Tamayo, 2024 +) only on abdomen, occurring both dorsally and ventrally. + + + + +Distribution. +Colombia +: +Magdalena Department +, Minca, Vereda Bellavista, Sierra Nevada de Santa Marta ( +Fig. 13B +, +14 +). + + +Natural history. +Most specimens were collected in the San Lorenzo area ( +Fig 13B +), with the exception of +one male +from Bella Vista area. The life zones of these localities belong to the very humid low montane forest and very humid subtropical forest, respectively ( +Tamarís-Turizo & López-Salgado, 2006 +). These sectors are located at altitudes between 1800 and +2900 m +.a.s.l. and the climatic conditions in them give rise to average temperatures between 12–18⁰C, with average annual rainfall ranging between +2000–4000 mm +and a geographical conformation that allows it to trap and condense cold air masses, so dense fogs usually form ( +Espinal, 1977 +; Guzmán, 1996). The vegetation is composed mainly of plants from the +Melastomataceae +, +Orchidaceae +, +Araceae +, +Arecaceae +, +Chrysobalanaceae +and +Bromeliaceae +families, as well as lichens and mosses ( +Cuadrado-Peña, 2005 +; +Espinal, 1977 +). However, the abundance of + +Pinus patula + +, a species of pine introduced by the San Lorenzo Experimental Station, is notable ( +Granda & Del Portillo, 2007 +). + + + +FIGURE 9A–E. + +Thalerommata margarita + + +sp. nov. + +, male holotype (ICN-Ar12996). A. Carapace. B. Chelicerae, labium, maxillae and sternum. C. Abdomen, dorsum. D–E. Left leg I tibial apophysis in ventral (D) and prolateral (E). Scale bars = 1 mm. Arrow points to the metatarsal distal bulge. + + + + +FIGURE 10A–E. + +Thalerommata margarita + + +sp. nov. + +, male holotype (ICN-Ar12996). A–B. Left palpal bulb in prolateral (A), retrolateral (B) views. C–E. Left palp in prolateral (C), ventral (D) and retrolateral (E) views. Scale bars = 0.5 mm (A–B), 1mm (C–E). + + + +The individuals were found under logs and stones, in some cases in aggregate form, registering up to 5 individuals in an area of +0.06 m +2 +. In these conditions, mainly adult females and juveniles were found, with few males recorded. They were observed inside cavities in the form of ditches in the ground covered with silk and on the ground in slopes with abundant bromeliads. When disturbed, they tend to escape with rapid movements. Small beetles, crickets, and cockroaches were observed in the same microhabitat as these, as well as other mygalomorphs such as + +Kankuamo marquezi +Perafán, Galvis & Gutiérrez + +, + +Striamea +sp. + +and families of spiders of the infraorder +Araneomorphae +(LB pers. obs.). + + + + +Discussion. +The species described here exhibit a digitiform apical segment of the PLS and a high number of labial cuspules ( +Figs. 3C–D +; +7B–D +; +11B–D +), showing morphological affinities with + +T. maculata + +and + +T. caudicula + +. This suggests a close relationship between these species and supports the hypothesis of +Nicoletta & Ferretti (2024) +, who consider that the +type +locality of + +T. caudicula + +may be an error and that its true geographic distribution could be associated with northern South America or the Caribbean. Records of the genus + +Thalerommata + +indicate a broad geographic distribution related to Caribbean countries, ranging from its northernmost points in +the Bahamas +to its southernmost points in +Colombia +( +Bertani & Raven 2023 +). This suggests the potential distribution of these tarantulas between Central America and northern South America. + + + +FIGURE 11A–D. + +Thalerommata margarita + + +sp. nov. + +, female paratype (ICN-Ar12997). A. Carapace. B. Chelicerae, labium, maxillae and sternum. C. Abdomen, dorsum. D. Spinnerets. Scale bars = 1 mm. + + + + +FIGURE 12A–B. +Spermathecae + +Thalerommata margarita + + +sp. nov. + +, female paratype (ICN-Ar12997). Scale bars = 0.5 mm. + + + +Shared characters between the genera + +Trichopelma + +and + +Thalerommata + +are evident, e.g., the distal article of PLS being short ( +Bertani & Raven 2023 +). However, the new species described herein confirms the variability of this character in + +Thalerommata + +. Additionally, +Mori & Bertani (2020) +recorded the presence of a thickening in the metatarsus of leg I of + +Trichopelma nitidum + +, referred to as the metatarsal bulge; this character is also present in + +T. kogui + + +sp. nov. + +and + +T. margarita + + +sp. nov. + +Similarly, +Raven (1985) +grouped + +Thalerommata + +and + +Trichopelma + +within the basal barychelids, suggesting that this should be considered in future phylogenetic studies involving these two genera. Fruitful studies will no doubt emerge on a possible sexual function of this characteristic, due to its exclusive position on leg I of males and its proximity to the tibial apophyses, important features in the sexual behavior of several mygalomorph spiders ( + +Ferretti +et al. +, 2013 + +). + + +The species + +T. anae + +and + +T. margarita + + +sp. nov. + +exhibit bulbs with a similar appearance but differ by the embolus being strongly curved towards the ventral face in + +T. margarita + + +sp. nov. + +This characteristic, along with the vast geographical distance separating these species by a barrier like the Caribbean Sea, the altitudinal differences with respect to sea level in the ecosystems they inhabit, and the difference in the number of labial cuspules, support the hypothesis that they are two distinct species. The long and slender appearance of the embolus in males of + +T. margarita + + +sp. nov. + +and + +T. yukpa + + +sp. nov. + +could be an adaptation to the long spermathecal receptacles of their conspecific females, a common phenomenon in various spiders ( +Eberhard & Huber, 2010 +). It is hypothesized that the unknown female of + +T. anae + +will also have elongated spermathecal receptacles. Other morphological characters such as tibial apophysis shape, bulb morphology, modified abdominal setae, presence or absence of spines on the cymbium, leg scopula, presence or absence of teeth on STC, presence of palpal claw tufts on females, number of cuspules on labium and maxillae are in agreement with the other species in the genus ( +Bertani & Raven 2023 +). + + +Two of the three newly described species ( + +T. kogui + +and + +T. margarita + +) come from the Sierra Nevada de Santa Marta ( +Fig. 13A +), a small region of +Colombia +recognized as a biodiversity hotspot. This highlights how the genus + +Thalerommata + +remains poorly known. With the inclusion of these three species, +Colombia +now hosts 7 of the 11 known + +Thalerommata +species. + +Undoubtedly, more new species will be discovered with further fieldwork in South America and the Caribbean, as well as investigations of unidentified specimens within collections. + +Thalerommata + +appears to be a very diverse genus with the potential to harbor additional species awaiting description. + + + + \ No newline at end of file diff --git a/data/BE/1A/87/BE1A87C3BC59BC0BFF1FF87CFBD2FDF5.xml b/data/BE/1A/87/BE1A87C3BC59BC0BFF1FF87CFBD2FDF5.xml index d3655e92986..8d4d10a4937 100644 --- a/data/BE/1A/87/BE1A87C3BC59BC0BFF1FF87CFBD2FDF5.xml +++ b/data/BE/1A/87/BE1A87C3BC59BC0BFF1FF87CFBD2FDF5.xml @@ -1,66 +1,66 @@ - - - -Three new species of tiny tarantulas of the genus Thalerommata Ausserer, 1875 (Araneae, Theraphosidae) from northern Colombia + + + +Three new species of tiny tarantulas of the genus Thalerommata Ausserer, 1875 (Araneae, Theraphosidae) from northern Colombia - - -Author + + +Author -Osorio, Luis M. -Grupo de Investigación Biodiversidad del Caribe Colombiano, Facultad de Ciencias Básicas, Universidad del Atlántico, Barranquilla, Colombia +Osorio, Luis M. +Grupo de Investigación Biodiversidad del Caribe Colombiano, Facultad de Ciencias Básicas, Universidad del Atlántico, Barranquilla, Colombia - - -Author + + +Author -Benavides, Léiner -Grupo de Investigación en Insectos Neotropicales. Universidad del Magdalena, Santa Marta Colombia +Benavides, Léiner +Grupo de Investigación en Insectos Neotropicales. Universidad del Magdalena, Santa Marta Colombia - - -Author + + +Author -García-Atencia, Sandy -Grupo de Investigación Biodiversidad del Caribe Colombiano, Facultad de Ciencias Básicas, Universidad del Atlántico, Barranquilla, Colombia +García-Atencia, Sandy +Grupo de Investigación Biodiversidad del Caribe Colombiano, Facultad de Ciencias Básicas, Universidad del Atlántico, Barranquilla, Colombia - - -Author + + +Author -Bertani, Rogério -Laboratório de Ecologia e Evolução, Instituto Butantan. 05503 - 900 São Paulo, SP, Brazil +Bertani, Rogério +Laboratório de Ecologia e Evolução, Instituto Butantan. 05503 - 900 São Paulo, SP, Brazil -text - - -Zootaxa +text + + +Zootaxa - -2024 - -2024-11-07 + +2024 + +2024-11-07 - -5537 + +5537 - -1 + +1 - -95 -114 + +95 +114 - -http://dx.doi.org/10.11646/zootaxa.5537.1.5 + +http://dx.doi.org/10.11646/zootaxa.5537.1.5 -journal article -10.11646/zootaxa.5537.1.5 -1175-5326 -14239357 -5AD70C1C-9942-471A-857B-D082C17C4F4D +journal article +10.11646/zootaxa.5537.1.5 +1175-5326 +14239357 +5AD70C1C-9942-471A-857B-D082C17C4F4D @@ -88,7 +88,7 @@ Type material . - + Holotype ♂ from @@ -101,9 +101,13 @@ from La Jagua de Ibirico , Vda. La Victoria -, Nueva Granada, +, +Nueva Granada +, Forest -, ground hand collecting ( +, +ground hand collecting +( 9°33’33.9”N , 73°09’39.5”W @@ -119,10 +123,12 @@ from L. Osorio & S. García-Atencia -leg. (ICN-Ar 12988) +leg. ( +ICN-Ar 12988 +) . - + Paratype (s) 3 ♂ @@ -137,8 +143,8 @@ from , La Jagua de Ibirico , -Alto -de las Flores, natural regeneration, +Alto de las Flores, natural regeneration +, pitfall ( 9°34’44.1”N @@ -148,11 +154,20 @@ de las Flores, natural regeneration, 1330 m - .a.s.l, -20 March 2021 -, L. Osorio & S. García leg. (ICN-Ar 12989), - + +20 March 2021 + +, +L. Osorio +& +S. García +leg. ( +ICN-Ar 12989 +) + +, + 1 ♀ dissected, same data as preceding ( 9°34’45.3”N @@ -162,35 +177,62 @@ dissected, same data as preceding ( 1282 m - .a.s.l, -8–11April 2022 -, J. Márquez, Y. Domínguez, S. García-Atencia & L. Osorio leg. (ICN-Ar 12990), - + +8–11April 2022 + +, +J. Márquez +, +Y. Domínguez +, +S. García-Atencia +& +L. Osorio +leg. ( +ICN-Ar 12990 +) + +, + 1 ♀ , Becerril , -Manantial -, natural regeneration, +Manantial, natural regeneration +, pitfall ( 9°38’07.0”N , 73°07’04.0”W ) - - -852 m + +852 m +. - +a.s.l, + +3 September 2021 + +, +J. Márquez +, +Y. Domínguez +, +S. García +& +L. Osorio +leg. ( +ICN-Ar 12991 +) -.a.s.l, -3 September 2021 -, J. Márquez, Y. Domínguez, S. García & L. Osorio leg. (ICN-Ar 12991), - +, + 1 ♀ -same data as preceding, coffee agroforestry system, +same data as preceding, +coffee agroforestry system +, pitfall ( 9°38’03.2”N @@ -198,15 +240,26 @@ same data as preceding, coffee agroforestry system, 73°06’56.4”W ) - 987 m - - .a.s.l, -3 September 2021 -, J. Márquez, Y. Domínguez, S. García & L. Osorio leg. (ICN-Ar 12992), - + +3 September 2021 + +, +J. Márquez +, +Y. Domínguez +, +S. García +& +L. Osorio +leg. ( +ICN-Ar 12992 +) + +, + 1 ♀ same data as preceding, Nueva Granada @@ -232,12 +285,14 @@ same data as preceding, S. García-Atencia & L. Osorio -leg. (ICN-Ar 12993). +leg. ( +ICN-Ar 12993 +). Other material examined. - + COLOMBIA : @@ -247,19 +302,19 @@ leg. (ICN-Ar 12993). La Jagua de Ibirico , Mina La Divisa -, oil palm monoculture, +, +oil palm monoculture +, pitfall trap ( 9°36’12.0”N , 73°15’49.1”W ) - - -146 m - + +146 m . - + a.s.l, 1 ♂ , @@ -268,7 +323,9 @@ a.s.l, , K. Caballero -leg. (CIUA-Ara0102) +leg. ( +CIUA-Ara0102 +) . diff --git a/data/BE/1A/87/BE1A87C3BC5EBC04FF1FFA49FEB1F85D.xml b/data/BE/1A/87/BE1A87C3BC5EBC04FF1FFA49FEB1F85D.xml index a2dbffb3553..042b81d3961 100644 --- a/data/BE/1A/87/BE1A87C3BC5EBC04FF1FFA49FEB1F85D.xml +++ b/data/BE/1A/87/BE1A87C3BC5EBC04FF1FFA49FEB1F85D.xml @@ -1,66 +1,66 @@ - - - -Three new species of tiny tarantulas of the genus Thalerommata Ausserer, 1875 (Araneae, Theraphosidae) from northern Colombia + + + +Three new species of tiny tarantulas of the genus Thalerommata Ausserer, 1875 (Araneae, Theraphosidae) from northern Colombia - - -Author + + +Author -Osorio, Luis M. -Grupo de Investigación Biodiversidad del Caribe Colombiano, Facultad de Ciencias Básicas, Universidad del Atlántico, Barranquilla, Colombia +Osorio, Luis M. +Grupo de Investigación Biodiversidad del Caribe Colombiano, Facultad de Ciencias Básicas, Universidad del Atlántico, Barranquilla, Colombia - - -Author + + +Author -Benavides, Léiner -Grupo de Investigación en Insectos Neotropicales. Universidad del Magdalena, Santa Marta Colombia +Benavides, Léiner +Grupo de Investigación en Insectos Neotropicales. Universidad del Magdalena, Santa Marta Colombia - - -Author + + +Author -García-Atencia, Sandy -Grupo de Investigación Biodiversidad del Caribe Colombiano, Facultad de Ciencias Básicas, Universidad del Atlántico, Barranquilla, Colombia +García-Atencia, Sandy +Grupo de Investigación Biodiversidad del Caribe Colombiano, Facultad de Ciencias Básicas, Universidad del Atlántico, Barranquilla, Colombia - - -Author + + +Author -Bertani, Rogério -Laboratório de Ecologia e Evolução, Instituto Butantan. 05503 - 900 São Paulo, SP, Brazil +Bertani, Rogério +Laboratório de Ecologia e Evolução, Instituto Butantan. 05503 - 900 São Paulo, SP, Brazil -text - - -Zootaxa +text + + +Zootaxa - -2024 - -2024-11-07 + +2024 + +2024-11-07 - -5537 + +5537 - -1 + +1 - -95 -114 + +95 +114 - -http://dx.doi.org/10.11646/zootaxa.5537.1.5 + +http://dx.doi.org/10.11646/zootaxa.5537.1.5 -journal article -10.11646/zootaxa.5537.1.5 -1175-5326 -14239357 -5AD70C1C-9942-471A-857B-D082C17C4F4D +journal article +10.11646/zootaxa.5537.1.5 +1175-5326 +14239357 +5AD70C1C-9942-471A-857B-D082C17C4F4D @@ -90,7 +90,7 @@ Figs 5– Type material. - + Holotype ♂ from @@ -103,9 +103,11 @@ from Santa Marta , Minca -, vda. -San Lorenzo -, ground hand collecting, cloud forest ( +, +vda. San Lorenzo +, +ground hand collecting +, cloud forest ( 11°06’18.2”N , 74°03’31.8”W @@ -122,13 +124,17 @@ from L. Benavides & M. J. García -leg. (ICN-Ar 12994) +leg. ( +ICN-Ar 12994 +) . - + Paratype 1 ♀ -same data as holotype (ICN-Ar 12995) +same data as holotype ( +ICN-Ar 12995 +) .