| Dominican Republic | +Oise, France | +Mexico | +
+ |
+|
|---|---|---|---|---|
| (VP coll.) | +
+(
+ |
+(VP coll.) | +
+ |
+|
| Bicyclic acidsa | +15 | +59 | +4 | +
+ |
+
| Bicyclic alcoholsb | +17 | +14 | +2 | +
+ |
+
| Bicyclic hydrocarbons | +47 | +7 | +53 | +
+ |
+
| 1-Methylbicyclic hydrocarbons | +22 | +19 | +41 | +28 | +
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|---|
|
+ |
+Adult male | +Adult male | +Adult male | +Adult female | +Adult female | +
|
+ |
+43.8 | +47.1 | +39.8 | +35.5 | +39.9 | +
|
+ |
+62.7 | +36.3* | +35.0* | +41.6 | +45.2 | +
|
+ |
+20.5 | +19.5 | +17.4 | +15.5 | +19.7 | +
|
+ |
+4.6 | +5.2 | +4.2 | +3.5 | +3.3 | +
|
+ |
+9.4 | +11.1 | +8.9 | +9.4 | +8.9 | +
|
+ |
+12.2 | +12.9 | +11.6 | +9.7 | +11.4 | +
|
+ |
+8.5 | +9.7 | +7.9 | +6.9 | +7.6 | +
|
+ |
+5.6 | +5.9 | +5.2 | +4.2 | +4.4 | +
|
+ |
+1.6 | +1.9 | +1.2 | +1.2 | +1.5 | +
|
+ |
+7.3 | +8.0 | +6.6 | +5.5 | +6.0 | +
|
+ |
+6.2 | +5.6 | +4.8 | +4.3 | +4.9 | +
|
+ |
+3.4 | +3.3 | +2.9 | +2.2 | +3.1 | +
|
+ |
+3.0 | +3.4 | +2.9 | +2.8 | +2.6 | +
|
+ |
+5.1 | +5.0 | +4.2 | +3.7 | +4.0 | +
|
+ |
+3.4 | +3.5 | +3.1 | +2.5 | +3.1 | +
|
+ |
+1.3 | +1.6 | +1.7 | +1.2 | +1.5 | +
|
+ |
+1.7 | +1.8 | +1.3 | +1.4 | +1.3 | +
|
+ |
+24 | +23 | +23 | +23 | +26 | +
|
+ |
+20 | +21 | +21 | +20 | +19 | +
|
+ |
+35 | +40 | +36 | +39 | +36 | +
|
+ |
+21 | +19 | +21 | +24 | +27 | +
|
+ |
+14 | +15 | +15 | +15 | +13 | +
|
+ |
+8 | +8 | +8 | +8 | +7 | +
|
+ |
+27 | +25 | +26 | +27 | +28 | +
|
+ |
+7/7 | +8/8 | +8/8 | +8/8 | +8/8 | +
|
+ |
+5/5 | +6/6 | +5/6 | +6/5 | +6/6 | +
|
+ |
+6/6 | +6/6 | +6/6 | +6/6 | +6/6 | +
|
+ |
+6 | +6 | +5 | +- | +- | +
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|---|---|---|---|---|---|---|
|
+ |
+
+ |
+40.6–47.5 | +36.6–42.1 | +
+ |
+
+ |
+31.8–38.6 | +35.0–42.5 | +31.3–41.6 | +36.6–43.9 | +
+ |
+37.6–47.8 | +40.7–42.6 | +
|
+ |
+41.6–62.7 | +51.0–53.2 | +33.5–51.5 | +2.5–37.8 | +58 | +27.7–37.1 | +39.0– 56.0 | +>8.0–61.5 | +46.4–54.1 | +52.1–69.0 | +>20.3–57.5 | +>46.1–57.3 | +
|
+ |
+
+ |
+3.7–4.4 | +3.3–4.4 | +2.3–5.2 | +- | +
+ |
+- | +3.1–4.6 | +2.8–4.4 | +
+ |
+3.2–5.0 | +4.1–4.5 | +
|
+ |
+8.9–11.1 | +8.5–9.7 | +- | +4.0–9.1 | +- | +- | +- | +- | +8.0–9.2 | +9.7–11.5 | +8.4–10.6 | +8.8–9.5 | +
|
+ |
+
+ |
+
+ |
+
+ |
+11.8–21.0 | +- | +
+ |
+- | +- | +- | +
+ |
+15.5–22.2 | +17.3–18.4 | +
|
+ |
+
+ |
+
+ |
+10.2–11.3 | +
+ |
+- | +8.1–9.8 | +10.5–12.8 | +9.0–13.0 | +
+ |
+
+ |
+12.0–14.8 | +12.7–13.6 | +
|
+ |
+6.9–9.7 | +6.0–7.7 | +6.2–7.6 | +5.1–7.9 | +- | +5.1–6.2 | +7.0–8.2 | +6.5–8.0 | +5.4–7.3 | +9.2–9.7 | +7.8–9.3 | +7.8–8.0 | +
|
+ |
+
+ |
+4.3–5.2 | +4.2–4.7 | +3.3–6.0 | +- | +
+ |
+- | +4.2–5.7 | +3.7–4.9 | +5.3–6.2 | +4.3–6.0 | +4.7–4.8 | +
|
+ |
+
+ |
+1.2–1.5 | +
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+0.8–1.2 | +1.4–1.7 | +0.9–1.3 | +0.6–0.7 | +
|
+ |
+5.5–8.0 | +- | +5.7–6.3 | +- | +- | +4.6–6.0 | +- | +- | +- | +7.6–8.5 | +6.2–8.0 | +6.5–6.6 | +
|
+ |
+
+ |
+5.9–6.7 | +4.9–5.6 | +4.0–6.2 | +- | +
+ |
+- | +4.8–5.6 | +4.9–6.2 | +
+ |
+4.4–6.8 | +5.1–6.1 | +
|
+ |
+
+ |
+2.2–3.3 | +2.6–3.0 | +1.8–2.9 | +- | +1.8–2.2 | +- | +2.6–3.8 | +2.4–2.9 | +
+ |
+2.5–3.3 | +2.9–3.0 | +
|
+ |
+
+ |
+3.2–3.8 | +2.6–3.0 | +2.2–3.7 | +- | +
+ |
+- | +3.1–3.9 | +3.2–3.8 | +
+ |
+3.1–4.1 | +3.5–3.8 | +
|
+ |
+
+ |
+4.4–5.2 | +4.0–4.4 | +3.0–4.6 | +- | +
+ |
+- | +3.8–4.6 | +4.1–5.1 | +
+ |
+4.1–5.6 | +4.5–5.1 | +
|
+ |
+
+ |
+3.4–3.9 | +2.6–3.2 | +2.2–4.0 | +- | +2.4–2.7 | +- | +2.8–3.7 | +3.1–4.0 | +
+ |
+3.1–4.3 | +3.5–3.9 | +
|
+ |
+
+ |
+1.3–1.6 | +1.0–1.3 | +0.9–2.1 | +- | +
+ |
+- | +0.9–1.5 | +1.2–1.7 | +1.7–1.8 | +1.6–2.1 | +1.5 | +
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+1.5–1.7 | +1.7–2.4 | +1.7–1.9 | +
|
+ |
+
+ |
+
+ |
+22–26 | +
+ |
+18–25 | +
+ |
+22 | +24 | +
+ |
+23–24 | +
+ |
+
+ |
+
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+20–23 | +
+ |
+
+ |
+
|
+ |
+
+ |
+38–44 | +
+ |
+- | +- | +
+ |
+- | +- | +31–42 | +
+ |
+33–35 | +
+ |
+
|
+ |
+
+ |
+23–26 | +- | +- | +- | +
+ |
+
+ |
+- | +18–25 | +24–25 | +20–25 | +18–21 | +
|
+ |
+13–15 | +13–15 | +- | +
+ |
+12–16 | +13–15 | +15 | +15 | +12–14 | +15 | +12–15 | +14–15 | +
|
+ |
+
+ |
+7–10 | +
+ |
+8–10 | +- | +8–10 | +- | +6–7 | +7–10 | +8–9 | +
+ |
+7 | +
|
+ |
+
+ |
+25–29 | +- | +- | +- | +- | +
+ |
+- | +26–31 | +26–27 | +
+ |
+24–26 | +
|
+ |
+
+ |
+7–9 | +8 | +
+ |
+7–8 | +8–9 | +
+ |
+
+ |
+7–8 | +8–10 | +8 | +7–8 | +
|
+ |
+
+ |
+6–7 | +
+ |
+6–7 | +6–7 | +6–8 | +
+ |
+6–8 | +5–7 | +
+ |
+
+ |
+6–7 | +
|
+ |
+6 | +5–6 | +6 | +- | +- | +6–7 | +7 | +- | +5– 6 | +7–8 | +6 | +5–6 | +
|
+ |
+
+ |
+
+ |
+
+ |
+5–7 | +6–7 | +
+ |
+
+ |
+
+ |
+5–6 | +
+ |
+6 - 7 | +6 | +
| Male holotype | +5 male | +5 female | +
|---|---|---|
| L 2922 | +3022±73.1 (2922–3082) | +3183±244.3 (2931–3447) | +
| a 22.2 | +22.9±3.1 (20.1–26.2) | +19.3±4.2 (16.7–25.5) | +
| b 4.8 | +5.3±0.1 (4.8–5.4) | +5.9±0.3 (5.4–6.3) | +
| c 17.2 | +19±1.3 (17.2–20.6) | +21.1±1.3 (19.3–22.3) | +
| c' 1.9 | +1.7±0.3 (1.5–2) | +2±0.3 (1.7–2.4) | +
| TL 170 | +159±10.3 (148–170) | +152±15.7 (139–172) | +
| M 132 | +134±20.8 (112–153) | +169±26.5 (131–191) | +
| HD 17 | +17±1 (16– 18) | +15±2.8 (11.6–18.1) | +
| a.b.d. 88 | +93±15 (79–109) | +78±6.2 (72–86) | +
| OLS 17 | +16±1 (15–17) | +16±2.8 (15–19) | +
| CSL 14 | +14 | +13±3 (11–15) | +
| E-abe 58 | +66± 6.8 (58–71) | +67±1.7 (65–69) | +
| gub 24 | +29±1.5 (24–31) | +|
| NR 277 | +256±14.1 (243–277) | +254±35.9 (210–290) | +
| ph L 606 | +574±8.1 (566–606) | +539±46.9 (469–568) | +
| ph ebd 116 | +115±15.4 (100–130) | +124±19.2 (99–145) | +
| s’ 1.4 | +1.5±0.3 (1.2–1.7) | +|
| spic 127 | +142±15.9 (127–160) | +|
| Egg (length x width) | +120 x 140 | +|
| V | +1707±114.3 (1550–1825) | +|
| V% | +53.7±3.5 (51.2–58.9) | +|
| GL | +942±135.6 (809–1066) | +|
| amph D 5 | +5±0.9 (5–5.2) | +6±0.8 (5.4–6.6) | +
| A-a.b.e. 11 | +10±1.2 (9–11) | +10±1 (10–11) | +
|
+ |
+
+ |
+|
|---|---|---|
| Carina begins Blunt teeth present | +Pereon 6 Pereon 7–Epimeron 2 | +Pereon 5 Pereon 6–Epimeron 3 | +
| Coxa 1–3 | +Similar in shape | +Coxa 3 longer and wider than coxa 1–2 | +
| Coxa 6 | +Smooth | +With lateral protrusion | +
|---|---|---|
| Pereopod 7 merus | +Rectangular | +Posterior corner pointed | +
| Discussion | +
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +warm-kollin | +||
|
+ |
+
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||
|
+ |
+0 - unbedeutend, keine Bindung. | +
| Ruhiges Wasser | +2 - Schwerpunktlebensraum | +
| Grundwasser | +0 - unbedeutend, keine Bindung. | +
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+209300 | +
| = | +
+ |
+
+ |
+2389 | +
| = | +
+ |
+
+ |
+2375 | +
| = | +
+ |
+
+ |
+2375 | +
| = | +
+ |
+
+ |
+209300 | +
| = | +
+ |
+
+ |
+153 | +
| = | +
+ |
+
+ |
+144 | +
| = | +
+ |
+
+ |
+209300 | +
| = | +
+ |
+
+ |
+2025 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +
+stark
+ |
+B2ab(iii,iv) | +
| Mittelland (MP) | +
+stark
+ |
+B2ab(iii,iv) | +
| Alpennordflanke (NA) | +
+stark
+ |
+B2ab(iii,iv) | +
|
+ |
+regional beziehungsweise in der Schweiz ausgestorben (Regionally Extinct) | +|
|
+ |
+-- | +|
| Westliche Zentralalpen (WA) | +-- | +
|
+ |
+
+3 - Mittlere nationale
+ |
+
|
+ |
+2 - Klarer Massnahmebedarf | +
|
+ |
+1 - Gering | +
|
+ |
+
+2 -
+ |
+
|
+ |
+Nein | +|
|
+ |
+
+ |
+(01.01.2018) | +
|
+ |
+
+ |
+(02.03.2005) | +
|
+ |
+-- | +|
|
+ |
+
+ |
+(23.01.2013) | +
| - | +
+ |
+
+ |
+
|---|---|---|
| Flowering pedicel length | +18-55 mm | +2-8 mm | +
| Inflorescence position | +supra-axillary | +extra-axillary/leaf opposed | +
| Number of petals visible in bud | +3 | +6 | +
| Outer petals form | +broadly ovate | +ovate to elliptic | +
| Inner petals form | +ovate | +elliptic | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +
+montan (
+ |
+||
|
+ |
+
+ |
+
+ |
+
+subozeanisch bis subkontinental (mittlere Luftfeuchtigkeit,
+ |
+||
| Head | +Tergites | +Appendages | +|||
|---|---|---|---|---|---|
| Ant. I | +192 | +Th. II | +200 | +Man | +540 | +
| Ant. II | +390 | +Th. III | +212 | +Dens | +580 | +
| Ant. III | +390 | +Abd. I | +110 | +Mucro | +48 | +
| Ant. IVa | +322 | +Abd. II | +120 | +
+ |
+1,168 | +
| Ant. IVb | +230 | +Abd. III | +140 | +Claw I | +62 | +
| Ant. | +1,524 | +Abd. IV | +655 | +Claw II | +62 | +
| Head | +465 | +Abd. V | +80 | +Claw III | +65 | +
| - | +- | +Abd. VI | +60 | +- | +- | +
| - | +- | +Body | +1,577 | +- | +- | +
| Characters | +
+ |
+
+ |
+
+ |
+
|---|---|---|---|
| Head dorsal surface | +clearly concave (fig. 5) | +almost flat (fig. 10) | +clearly concave (fig. 15) | +
| Preocular part of head | +swollen with very small sharp process (fig. 5) | +flat with sharp process (fig. 10) | +very swollen without process | +
| Interocelar distance | +1.8–2.1 | +1.8–1.9 | +1.45–1.55 | +
| Third antennal segment | +flat and broad | +flat and only slightly broad- ened | +flat and only slightly broadened | +
| Apex of the 4th antennal segment | +of the same color as its remaining part | +clearly lighter than its remaining part | +clearly lighter than its remaining part | +
| Pronotum | +anterior median tuberosity swollen, lateral pronotal mar- gins irregularly rugged with pointed projection (fig. 6) | +anterior median tuberosity flat and almost invisible, lateral pronotal margins smooth with rounded projection (fig. 11) | +anterior median tuberosity strongly swollen, lateral pronotal margins irregularly rugged with obtuse projection (fig. 16) | +
| Lateral margins of sterna | +with small posterior apical projection and very small median lobe, yellow spot absent (fig. 7) | +with a single pyramid-shaped apical projection bearing yellow spot, and a small median lobe (fig. 12) | +with a single pyramid-shaped apical projection bearing yellow spot, and a small median lobe (fig. 17) | +
| Paramere | +rounded, bearing small hypo- physis with shallow incision (fig. 9) | +oval with elongate hypophy- sis (fig. 13) | +elongate with sharp and prolonged hypophysis (fig. 18) | +
| Anterior part of ejaculatory reservoir | +strongly coiled (fig. 8) | +strongly coiled (fig. 14) | +straight (fig.19) | +
| Numbers | +Definition | +
|---|---|
| 1 | +Most anterior points of the premaxilla | +
| 2–3 | +Most anterior points in the joint between the | +
| premaxilla and maxilla | +|
| 4–5 | +Most lateral points in the joint between the | +
| prefrontal and maxilla | +|
| 6–7 | +Most posterior points of the maxilla | +
| 8–9 | +Most anterior points on the suture between | +
| the parietal and pterygoid | +|
| 10–11 | +Most anterior points of the pterygoid | +
| 12 | +Centre of the joint between the frontal and | +
| parietal | +|
| 13–14 | +Most posterolateral points of the pterygoid | +
| 15–16 | +Most anterolateral points of the jaw joint | +
| 17–18 | +Most posteromedial points of squamosal | +
| 19–20 | +Most posterolateral points of the jaw joint | +
| 21 | +Most posteromedial point of the parietal | +
| 22–23 | +Most posterior points of the exoccipital | +
| Phase | +
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|
|
+ |
+||||
| I | +– | +– | +– | +– | +
| II | +Columella, dentary, | +– | +– | +– | +
| parasphenoid, | +||||
| prearticular, | +||||
| premaxilla, | +||||
| squamosal, vomer | +||||
| III | +Angular, coronoid | +– | +– | +– | +
| IV | +Parietal | +– | +– | +– | +
| V | +Maxilla | +– | +– | +– | +
| VI | +Exoccipital, frontal, | +– | +Dentary, exoccipital, | ++Coronoid, + |
| dennasal, pterygoid | +opisthotic, palatine, | +tary, exoccipital, | +||
| parasphenoid, parietal, | +paraphenoid, | +|||
| prootic, prearticular, | +premaxilla, prootic, | +|||
| premaxilla, pterygoid, | +vomer | +|||
| squamosal, vomer | +||||
| VII | +Exoccipital, | +– | +Angular, columella, cor- | +Maxilla, palatine, | +
| hypobranchial II, | +onoid, frontal, maxilla, | ++parasphenoid, + | ||
| parprefrontal, quadrate | +prefrontal, quadrate | +ietal, squamosal | +||
| VIII | +Orbitosphenoid | +Exoccipital | +Basibranchial II, nasal, | +Angular, columella, | +
| orbitosphenoid | +frontal, opisthotic, | +|||
| IX | +– | +Parietal | +– | ++prearticular, + |
| pteryX | +– | +Angular, articular, colu- | +– | +goid | +
| mella, coronoid, dentary, | +||||
| opisthotic, hypobranchial II, | +||||
| prearticular, premaxilla, | +||||
| pterygoid, prootic, quadrate | +||||
| XI | +– | +Ceratobranchial II, frontal, | +– | +|
| +maxilla, nasal, + | ||||
| orbitosphenoid, parasphenoid | +||||
| XII | +Articular, | +– | +Articular, ceratobran- | +Articular, | +
| ceratobranchial II | +chial II, ceratohyal, | +basibranchial II, | +||
| hypobranchial II | ++nasal, + | |||
| orbitosphenoid, prefrontal | +||||
| XIII | +– | +– | +– | +– | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Range | +|||||
|---|---|---|---|---|---|
| Measurements | +H | +Min | +Max | +Mean | +SD | +
| Standard length | +30.1 | +17.6 | +30.1 | +23.7 | +- | +
| Percent of Standard length | +|||||
| Body depth | +15.5 | +14.2 | +16.8 | +15.7 | +0.96 | +
| Predorsal distance | +46.7 | +45.6 | +51.1 | +48.1 | +1.67 | +
| Prepelvic distance | +42.7 | +42.2 | +48.9 | +44.9 | +2.15 | +
| Preanal distance | +64.6 | +64.1 | +68.3 | +65.9 | +1.09 | +
| Preadipose distance | +77.5 | +74.7 | +78.6 | +76.8 | +1.22 | +
| Dorsal-fin spine length | +23.7 | +20.1 | +26.0 | +24.1 | +1.65 | +
| Anal-fin unbranched ray length | +13.2 | +12.9 | +16.2 | +14.8 | +1.02 | +
| Adipose-fin spine length | +6.3 | +6.0 | +9.2 | +7.5 | +0.91 | +
| Pectoral-fin spine length – males (9) | +27.2 | +32.1 | +28.9 | +1.60 | +|
| Pectoral-fin spine length – females (7) | +28.4 | +22.8 | +29.1 | +25.8 | +2.18 | +
| Pectoral-fin spine depth | +1.7 | +1.1 | +2.0 | +1.6 | +0.22 | +
| Caudal peduncle depth | +6.5 | +6.5 | +8.1 | +7.1 | +0.42 | +
| Dorsal-adipose fin distance | +16.4 | +13.3 | +17.5 | +15.9 | +1.27 | +
| Dorsal-fin base length | +12.9 | +10.8 | +14.2 | +12.6 | +0.88 | +
| Lower caudal-fin principal ray | +22.7 | +22.7 | +28.6 | +25.8 | +1.50 | +
| Cleithral width | +26.1 | +25.0 | +26.7 | +25.9 | +0.53 | +
| Head length | +35.9 | +34.8 | +42.0 | +38.8 | +2.16 | +
| Percent of Head length | +|||||
| Head depth | +43.8 | +37.9 | +47.0 | +42.2 | +2.95 | +
| Interorbital distance | +37.3 | +32.1 | +38.4 | +35.0 | +1.95 | +
| Orbital horizontal diameter | +16.0 | +15.1 | +17.1 | +16.0 | +0.59 | +
| Snout length | +54.0 | +52.0 | +55.9 | +54.1 | +1.28 | +
| Internareal distance | +9.6 | +6.3 | +9.9 | +8.0 | +1.19 | +
|
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|
| Length of ears | +<20 mm | +<21 mm | +> 21 mm | +
| Connecting band between ears | +Absent | +Absent | +Present, with variable development | +
| Skull | +Delicate to robust, wide and short | +Very robust, wide and short | +Delicate, long and narrow | +
| Development of crests | +Sagittal and lambdoidal crests weakly to well developed, united or not | +Sagittal and lambdoidal crests well developed, united | +Sagittal and lambdoidal crests weakly developed, separated | +
| Postorbital process of the jugal | +Weakly developed | +Weakly developed | +Well developed | +
| Eyes | +Small | +Small | +Large | +
| Auditory bullae | +Small | +Small | +Large | +
| Length of dorsal fur (mean) | +4.5–13.1 mm (8.1 mm) | +8.1–13.7 mm (10.1 mm) | +8.0–14.5 mm (11.4 mm) | +
| Length of forearm (mean) | +31.4–48.8 mm (40.1 mm) | +40.1–54.0 mm (46.6 mm) | +44.0–52.6 mm (47.8 mm) | +
| Tragus | +Wider at the base, short, rounded, smooth and slender | +Wider at the base, short, rounded, smooth and broad | +Wider at the base, long, pointed and notched | +
| Occipital helmet | +Absent to well developed | +Well developed | +Absent | +
| GLS (mean) | +12.9–18.0 mm (15.8 mm) | +16.9–20.8 mm (19.1 mm) | +17.4–19.8 mm (18.4 mm) | +
| MTL (mean) | +4.5–7.0 mm (5.8 mm) | +6.2–7.7 mm (7.1 mm) | +5.6–7.1 mm (6.3 mm) | +
|
+ |
+Type | +locality: | +«Tlacotalpam | +[= | +Tlacotalpan, | +Veracruz], | +Mexique» | +(original | +
|---|---|---|---|---|---|---|---|---|
| citation). | +||||||||
|
+ |
+
| Species | +N | +WH/LV (mean) | +LE/LP (mean) | +LE/WE (mean) | +
|---|---|---|---|---|
|
+ |
+24 | +2.18–2.50 (2.36) | +2.53–2.73 (2.62) | +1.45–1.58 (1.51) | +
|
+ |
+101 | +1.10–1.44 (1.29) | +2.10–2.69 (2.37) | +1.45–1.62 (1.53) | +
| Eastern U.S.A. | +24 | +1.10–1.21 (1.15) | +2.10–2.30 (2.23) | +1.49–1.62 (1.56) | +
| Texas | +24 | +1.15–1.35 (1.26) | +2.28–2.49 (2.38) | +1.47–1.61 (1.53) | +
| Arizona + New Mexico | +24 | +1.29–1.44 (1.38) | +2.31–2.62 (2.46) | +1.45–1.61 (1.52) | +
| Mexico | +24 | +1.24–1.42 (1.36) | +2.24–2.69 (2.41) | +1.47–1.59 (1.52) | +
| Cuba | +5 | +1.23–1.32 (1.26) | +2.15–2.31 (2.24) | +1.41–1.50 (1.46) | +
|
+ |
+2 | +1.47–1.49 | +2.26 | +1.49 | +
|
+ |
+24 | +1.67–1.80 (1.73) | +2.43–2.53 (2.48) | +1.44–1.55 (1.48) | +
|
+ |
+24 | +1.17–1.40 (1.27) | +1.85–2.07 (1.95) | +1.81–2.10 (1.93) | +
|
+ |
+30 | +1.41–1.74 (1.55) | +1.93–2.18 (2.02) | +1.60–1.80 (1.70) | +
| Northern states | +20 | +1.41–1.58 (1.50) | +1.93–2.08 (2.02) | +1.60–1.80 (1.69) | +
| Southern states | +10 | +1.53–1.74 (1.65) | +1.99–2.18 (2.07) | +1.68–1.78 (1.74) | +
|
+ |
+34 | +1.46–1.61 (1.52) | +1.78–1.90 (1.83) | +1.69–1.89 (1.77) | +
| Mexico | +24 | +1.46–1.61 (1.51) | +1.78–1.86 (1.82) | +1.69–1.86 (1.76) | +
| Jamaica | +10 | +1.47–1.58 (1.53) | +1.80–1.90 (1.86) | +1.71–1.89 (1.78) | +
|
+ |
+15 | +1.53–1.73 (1.66) | +1.95–2.19 (2.09) | +1.46–1.68 (1.60) | +
| Cuba + Cayman Islands | +5 | +1.53–1.71 (1.61) | +1.95–2.04 (2.00) | +1.46–1.65 (1.55) | +
| Haiti | +10 | +1.62–1.73 (1.69) | +2.08–2.19 (2.13) | +1.55–1.68 (1.62) | +
|
+ |
+15 | +1.38–1.53 (1.47) | +2.05–2.20 (2.11) | +1.68–1.85 (1.76) | +
|
+ |
+24 | +1.40–1.59 (1.50) | +1.65–1.81 (1.73) | +1.74–1.90 (1.80) | +
|
+ |
+24 | +1.48–1.71 (1.59) | +1.98–2.13 (2.07) | +1.65–1.85 (1.77) | +
|
+ |
+3 | +2.35–2.41 (2.38) | +1.98–2.04 (2.01) | +1.61–1.68 (1.65) | +
|
+ |
+24 | +2.27–2.60 (2.43) | +1.64–1.84 (1.75) | +1.48–1.64 (1.58) | +
|
+ |
+24 | +1.28–1.41 (1.34) | +1.84–2.00 (1.92) | +1.73–1.93 (1.82) | +
|
+ |
+1 | +2.27 | +2.33 | +1.86 | +
|
+ |
+11 | +2.25–2.71 (2.45) | +2.17–2.36 (2.26) | +1.54–1.75 (1.64) | +
|
+ |
+24 | +2.19–2.47 (2.31) | +1.67–1.91 (1.78) | +1.43–1.55 (1.49) | +
|
+ |
+3 | +2.14–2.32 (2.24) | +1.43–1.66 (1.54) | +1.39–1.56 (1.50) | +
|
+ |
+locality: «Yucatan | +[Mexico]» | +(original | +citation). | +
|---|---|---|---|---|
|
+ |
+||||
|
+ |
+
| Character | +
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|---|---|---|---|---|---|---|
|
+ |
+|||||||||||
| Stem height (cm) | +150–300 | +100–200 | +50–120 | +100–250 | +80–150 | +40–100(130) | +up to 300 | +100–200 | +150 | +(40) 80–100 | +80–150 | +
| Leaf shape and | +sub-opaque | +shining green | +shining green | +shining green | +shining | +green | +green | +shining | +shining | +glaucescens | +green | +
| color | +green | +adaxial, | +green | +glaucos | +green | +green | +adaxial, pale | +||||
| glaucos | +glaucos | +glacous | +green abaxial | +||||||||
| abaxial | +|||||||||||
| Leaf blades | +rhombic | +triangular | +triangular | +rhombic | +triangular | +triangular | +rhombic | +rhombic | +triangular | +triangular- | +rhombic | +
| outline | +ovate | +||||||||||
| Leaf blade lenght | +100 | +40–60 | +18–70 | +20–50 | +20–40 | +40–50 | +20–50 | +40–60 | +15–40 | +8–20 | +up to 20 | +
| (cm) | +|||||||||||
| Leaf | +glabrous | +glabrous | +scabrous- | +glabrous | +glabrous | +glabrous | +glabrous | +glabrous | +glabrous | +glabrous | +glabrous | +
| indumentum | +hispidulous | +or rar. | +to slightly | +||||||||
| scabridulous | +scabrous | +||||||||||
| Leaf pinnation | +4–5 | +3 | +5–6 | +3–4 | +2–3 | +5–6(7) | +5–6 | +3–4(5) | +3–5 | +3–4 | +4 | +
| Leaf segments | +20–40(50) x | +5–10(30) x | +0.5–5 x 0.5–1 | +(1)3–15 x | +2–10 x | +0.5–2(4) x | +1.2–5 x | +2–6 x 1.5–2 | +1–2 (5) x | +3–10 x | +0.5–2 x | +
| size (mm) | +0.4–1 | +1–2(3) | +1.3–1.5 | +0.8–1.1 | +0.4–0.7(1) | +0.5–0.6 | +1–1.3 | +0.5–2 | +0.5–1 | +||
| Hermaphrodite | +12–26(50) | +25–40 | +10–30 | +16–28 | +16–18 | +7–14(20) | +16–28 | +8–16(22) | +10–15 | +7–12 | +6–11 | +
| umbel rays | +|||||||||||
| number | +|||||||||||
| Hermaphrodite | +(13)22–45 | +10–30 | +15–40 | +20–30 | +15–30 | +37–66(90) | +20–35 | +22–35 | +25–45 | +18–45 | +26–32 | +
| umbels rays | +|||||||||||
| lenght (mm) | +|||||||||||
| Staminate | +14–20 | +13–18 | +10–25 | +18 | +<20 | +6–10 | +14–18 | +16–18 | +9–12 | +13–30 | +7–10 | +
| umbels rays | +|||||||||||
| number | +|||||||||||
| Staminate | +5–9(13) | +10–15 | +5–24 | +10–20 | +15–30 | +16–27 | +8–14 | +4–6 | +10–15 | +6–11 | +12–16 | +
| umbels rays | +|||||||||||
| lenght (mm) | +|||||||||||
| Mericarp shape | +obovate to | +elliptical | +rounded to | +rounded to | +obovate- | +elliptical | +elliptical | +elliptical | +elliptical to | +elliptical to | +rounded to | +
| obovate- | +rounded-oblong | +rounded-oblong | +oblong to | +rounded | +orbicular | +obovate | +|||||
| elliptical | +elliptical | +||||||||||
| Mericarp size | +(11)13–17 x | +13–15 x 7–8 | +9–16 x 8–14 | +11–14 x 9–11 | +7–10 x 3–6 | +6.5–11 x | +10–13 x | +8–15 x 6–8 | +12–18 x | +8–12 x 6–8 | +6–8 x 4–6 | +
| (mm) | +7–11 | +5–6.5 | +6.5–8.5 | +12–14 | +|||||||
| Mericarp wings | +1.7–2.5 | +1–1.2 | +1–1.6 | +1.5–2 | +0.5–1 | +0.5–1 | +1.2–1.7 | +0.25–1 | +1.6–2 | +0.7–1 | +0.5–0.8 | +
| width (mm) | +|||||||||||
| Vittae per | +1–3 | +3–4 | +2–3 | +1–4 | +1–3 | +2–4 | +2–3 | +3 | +3 | +2–3 | +1–3 | +
| valecula | +|||||||||||
| Commissural | +2–4 | +4–6 | +4–6 | +6–8 | +4–5 | +8–10(12) | +4–6 | +4–6 | +4 | +4 | +5–6 | +
| vittae | +
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +warm-kollin | +||
|
+ |
+
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+77700 | +
| = | +
+ |
+
+ |
+264 | +
| = | +
+ |
+
+ |
+350 | +
| = | +
+ |
+
+ |
+350 | +
| = | +
+ |
+
+ |
+77700 | +
| = | +
+ |
+
+ |
+848 | +
| = | +
+ |
+
+ |
+77700 | +
| = | +
+ |
+
+ |
+151 | +
| CIB WB2020190 | +CIB WB2020081511 | +CIB WB2020191 | +CIB TZ20160714005 | +Ranging (Mean±SD) | +CIB TZ20160714002 | +CIB TZ20160714010 | +CIB WB2020201 | +CIB WB2020202 | +|
|---|---|---|---|---|---|---|---|---|---|
|
+ |
+Male | +Male | +Male | +Male | +Male (n=4) | +Female | +Female | +Larva | +Larva | +
|
+ |
+131.4 | +135.1 | +120.5 | +121.6 | +120.5–135.1 (127.1 ± 7.2) | +127.6 | +123.5 | +36.9 | +41.3 | +
|
+ |
+62.9 | +65.7 | +61.1 | +65.9 | +61.1–65.9 (63.9 ± 2.3) | +69.2 | +66.7 | +20.6 | +21.4 | +
|
+ |
+16.1 | +16.1 | +14.2 | +15.9 | +14.2–16.1 (15.6 ± 0.9) | +16.1 | +15.7 | +7.1 | +7.5 | +
|
+ |
+15.8 | +15.8 | +15 | +15.1 | +15.0–15.8 (15.4± 0.4) | +14.7 | +14.9 | +5.6 | +6.3 | +
|
+ |
+6.4 | +6.9 | +6.8 | +6.5 | +6.4–6.9 (6.6± 0.2) | +5.8 | +6.1 | +2.1 | +2 | +
|
+ |
+4.6 | +5.1 | +4.4 | +3.5 | +3.5–5.1 (4.4± 0.6) | +4.9 | +4.7 | +1.4 | +1.8 | +
|
+ |
+7.2 | +7.6 | +7.3 | +8.1 | +7.2–8.1 (7.5± 0.4) | +8.5 | +8.5 | +1.7 | +2.1 | +
|
+ |
+2 | +2 | +1.8 | +1.8 | +1.8–2.0 (1.9± 0.1) | +1.9 | +2 | +0.7 | +0.9 | +
|
+ |
+5.8 | +5.6 | +4.7 | +5.7 | +4.7–5.8 (5.4± 0.5) | +5.1 | +5.1 | +1.3 | +1.3 | +
|
+ |
+8.8 | +8.3 | +8.3 | +8.9 | +8.3–8.9 (8.8± 0.3) | +9 | +8.3 | +2.6 | +2.8 | +
|
+ |
+11.9 | +12.2 | +11.9 | +13.3 | +11.9–13.3 (12.3± 0.7) | +13.1 | +12.2 | +3.5 | +4 | +
|
+ |
+46.6 | +50.6 | +45.1 | +49.9 | +45.1–50.6 (48.0± 2.6) | +53.6 | +51.9 | +13.4 | +14.5 | +
|
+ |
+64 | +62.9 | +56.7 | +54.5 | +54.5–64.0 (59.5± 4.6) | +58.8 | +55.8 | +15.8 | +18.2 | +
|
+ |
+10.7 | +8.1 | +7.7 | +8.4 | +7.7–10.7 (8.7± 1.4) | +8.2 | +7.5 | +5.2 | +5.6 | +
|
+ |
+7.7 | +7.8 | +7.7 | +8.1 | +7.7–8.1 (7.8± 0.2) | +7.9 | +7.2 | +2.3 | +2.1 | +
|
+ |
+7.1 | +7.7 | +6.4 | +7.6 | +6.4–7.7 (7.2± 0.6) | +5.6 | +6.3 | +1.7 | +1.9 | +
|
+ |
+4.7 | +4.9 | +4.8 | +4.7 | +4.7–4.9 (4.8± 0.1) | +4.4 | +3.5 | +1.5 | +1.8 | +
|
+ |
+5.6 | +5.4 | +5.5 | +4.3 | +4.3–5.6 (5.2± 0.6) | +5.1 | +4.7 | +1.2 | +1 | +
|
+ |
+5.5 | +6.2 | +6.7 | +5.9 | +5.5–6.7 (6.1± 0.5) | +6.2 | +5.9 | +1.9 | +1.8 | +
| ID | +Species | +1 | +2 | +3 | +4 | +5 | +6 | +7 | +8 | +9 | +10 | +11 | +12 | +13 | +14 | +15 | +16 | +
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| 1 | +
+ |
+||||||||||||||||
| 2 | +
+ |
+2.7 | +|||||||||||||||
| 3 | +
+ |
+2.9 | +4.0 | +||||||||||||||
| 4 | +
+ |
+2.7 | +3.5 | +3.8 | +|||||||||||||
| 5 | +
+ |
+2.8 | +3.8 | +3.6 | +3.1 | +||||||||||||
| 6 | +
+ |
+2.5 | +3.5 | +3.7 | +2.9 | +2.2 | +|||||||||||
| 7 | +
+ |
+3.9 | +4.7 | +4.9 | +4.4 | +4.6 | +4.7 | +||||||||||
| 8 | +
+ |
+3.8 | +5.0 | +4.3 | +4.4 | +4.3 | +4.4 | +3.9 | +|||||||||
| 9 | +
+ |
+6.9 | +7.8 | +7.7 | +7.5 | +7.6 | +7.7 | +8.0 | +7.4 | +||||||||
| 10 | +
+ |
+8.8 | +9.5 | +9.0 | +9.1 | +8.8 | +8.9 | +9.4 | +8.5 | +7.6 | +|||||||
| 11 | +
+ |
+11.1 | +12.0 | +11.9 | +12.1 | +12.3 | +11.6 | +12.4 | +11.2 | +11.8 | +12.9 | +||||||
| 12 | +
+ |
+10.0 | +10.4 | +10.4 | +10.2 | +11.1 | +10.8 | +11.3 | +11.1 | +10.2 | +10.7 | +12.4 | +|||||
| 13 | +
+ |
+8.8 | +10.2 | +9.1 | +9.0 | +9.8 | +9.7 | +10.6 | +10.4 | +9.3 | +11.5 | +13.0 | +11.6 | +||||
| 14 | +
+ |
+8.7 | +10.1 | +9.0 | +8.9 | +9.7 | +9.6 | +10.5 | +10.3 | +8.9 | +10.9 | +12.8 | +11.5 | +0.5 | +|||
| 15 | +
+ |
+8.4 | +9.8 | +8.9 | +9.2 | +9.3 | +9.3 | +10.1 | +9.4 | +9.4 | +11.5 | +11.9 | +12.3 | +3.6 | +3.5 | +||
| 16 | +
+ |
+7.9 | +9.4 | +8.8 | +8.5 | +8.7 | +8.8 | +9.8 | +9.3 | +9.1 | +10.4 | +13.1 | +11.2 | +5.1 | +4.8 | +5.6 | +|
| 17 | +
+ |
+8.5 | +10.0 | +9.1 | +9.0 | +9.3 | +9.4 | +10.3 | +9.6 | +9.4 | +10.9 | +12.7 | +11.3 | +5.1 | +4.8 | +5.2 | +2.9 | +
| 18 | +
+ |
+8.0 | +9.8 | +8.8 | +9.3 | +9.0 | +9.3 | +9.5 | +9.9 | +8.4 | +9.7 | +12.1 | +11.3 | +5.9 | +5.4 | +5.5 | +6.2 | +
| 19 | +
+ |
+7.9 | +9.5 | +8.4 | +8.1 | +8.9 | +8.8 | +10.0 | +9.4 | +8.2 | +9.4 | +11.8 | +10.5 | +5.6 | +5.0 | +5.3 | +4.8 | +
| 20 | +
+ |
+8.0 | +9.5 | +8.7 | +8.5 | +9.1 | +9.0 | +10.1 | +9.5 | +9.0 | +11.0 | +12.3 | +10.9 | +5.3 | +5.0 | +4.6 | +5.0 | +
| 21 | +
+ |
+10.8 | +12.0 | +11.6 | +11.2 | +11.0 | +11.8 | +11.9 | +10.5 | +11.3 | +12.7 | +13.4 | +14.0 | +13.4 | +13.0 | +12.7 | +12.1 | +
| 22 | +
+ |
+10.8 | +12.0 | +11.4 | +11.2 | +11.0 | +11.8 | +11.7 | +10.7 | +11.5 | +12.8 | +13.6 | +14.2 | +13.4 | +13.0 | +12.6 | +12.1 | +
| 23 | +
+ |
+9.9 | +11.4 | +10.8 | +10.7 | +10.6 | +11.1 | +11.2 | +9.8 | +10.7 | +11.9 | +13.0 | +13.4 | +12.7 | +12.3 | +12.0 | +11.2 | +
| 24 | +
+ |
+10.2 | +11.7 | +10.7 | +10.8 | +10.9 | +11.4 | +11.1 | +10.4 | +11.0 | +12.3 | +14.4 | +13.6 | +12.6 | +12.3 | +11.8 | +11.7 | +
| 25 | +
+ |
+11.1 | +13.3 | +11.9 | +12.0 | +11.9 | +12.6 | +13.0 | +11.8 | +11.0 | +13.6 | +14.6 | +14.5 | +12.9 | +12.5 | +12.4 | +11.7 | +
| 26 | +
+ |
+11.5 | +13.3 | +12.6 | +12.2 | +12.2 | +13.2 | +13.7 | +12.2 | +11.3 | +13.9 | +15.5 | +14.4 | +13.7 | +13.4 | +13.3 | +11.8 | +
| 27 | +
+ |
+12.9 | +14.7 | +14.2 | +14.3 | +14.0 | +14.2 | +14.9 | +13.9 | +14.0 | +14.3 | +16.8 | +16.3 | +15.0 | +14.6 | +14.3 | +14.1 | +
| 28 | +
+ |
+10.0 | +12.0 | +11.1 | +11.0 | +10.5 | +11.3 | +11.4 | +10.5 | +10.2 | +12.1 | +13.4 | +13.9 | +11.8 | +11.5 | +11.0 | +11.3 | +
| 29 | +
+ |
+10.5 | +12.1 | +11.9 | +11.0 | +11.3 | +11.6 | +12.6 | +12.2 | +11.3 | +13.0 | +13.5 | +13.8 | +12.9 | +12.3 | +12.4 | +12.0 | +
| 30 | +
+ |
+12.0 | +13.8 | +13.3 | +12.7 | +12.7 | +13.1 | +13.9 | +13.4 | +12.6 | +14.4 | +14.5 | +14.8 | +14.4 | +13.8 | +13.7 | +13.3 | +
| 31 | +
+ |
+10.9 | +12.3 | +11.9 | +11.4 | +11.7 | +12.2 | +12.1 | +11.5 | +10.9 | +12.6 | +14.2 | +13.7 | +12.9 | +12.3 | +12.7 | +11.5 | +
| 32 | +
+ |
+9.8 | +11.3 | +10.7 | +9.7 | +10.5 | +10.8 | +11.7 | +10.6 | +10.2 | +11.8 | +13.3 | +11.8 | +11.1 | +10.8 | +11.0 | +10.8 | +
| 33 | +
+ |
+8.7 | +10.0 | +9.7 | +10.1 | +9.7 | +10.4 | +10.3 | +9.3 | +9.7 | +10.3 | +12.4 | +11.6 | +11.0 | +10.6 | +10.0 | +9.8 | +
| 34 | +
+ |
+8.8 | +10.4 | +10.3 | +10.1 | +10.1 | +10.8 | +10.6 | +10.1 | +9.8 | +10.9 | +13.0 | +11.5 | +10.7 | +10.2 | +10.5 | +9.6 | +
| ID | +Species | +17 | +18 | +19 | +20 | +21 | +22 | +23 | +24 | +25 | +26 | +27 | +28 | +29 | +30 | +31 | +32 | +33 | +
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| 1 | +
+ |
+|||||||||||||||||
| 2 | +
+ |
+|||||||||||||||||
| 3 | +
+ |
+|||||||||||||||||
| 4 | +
+ |
+|||||||||||||||||
| 5 | +
+ |
+|||||||||||||||||
| 6 | +
+ |
+|||||||||||||||||
| 7 | +
+ |
+|||||||||||||||||
| 8 | +
+ |
+|||||||||||||||||
| 9 | +
+ |
+|||||||||||||||||
| 10 | +
+ |
+|||||||||||||||||
| 11 | +
+ |
+|||||||||||||||||
| 12 | +
+ |
+|||||||||||||||||
| 13 | +
+ |
+|||||||||||||||||
| 14 | +
+ |
+|||||||||||||||||
| 15 | +
+ |
+|||||||||||||||||
| 16 | +
+ |
+|||||||||||||||||
| 17 | +
+ |
+|||||||||||||||||
| 18 | +
+ |
+5.8 | +||||||||||||||||
| 19 | +
+ |
+5.0 | +3.3 | +|||||||||||||||
| 20 | +
+ |
+5.1 | +5.5 | +4.5 | +||||||||||||||
| 21 | +
+ |
+12.1 | +10.9 | +11.1 | +12.3 | +|||||||||||||
| 22 | +
+ |
+12.1 | +11.1 | +11.1 | +12.3 | +1.2 | +||||||||||||
| 23 | +
+ |
+11.1 | +10.0 | +10.2 | +11.3 | +2.2 | +2.6 | +|||||||||||
| 24 | +
+ |
+11.7 | +10.4 | +11.0 | +11.7 | +4.2 | +4.4 | +3.8 | +||||||||||
| 25 | +
+ |
+11.6 | +10.3 | +10.8 | +11.8 | +4.5 | +4.5 | +4.1 | +5.6 | +|||||||||
| 26 | +
+ |
+12.1 | +11.6 | +11.6 | +12.3 | +5.2 | +5.6 | +4.6 | +6.6 | +4.4 | +||||||||
| 27 | +
+ |
+14.6 | +13.6 | +13.8 | +14.9 | +7.7 | +7.7 | +7.2 | +8.8 | +7.7 | +7.7 | +|||||||
| 28 | +
+ |
+11.1 | +9.4 | +10.3 | +10.9 | +4.1 | +4.5 | +3.9 | +5.4 | +4.3 | +5.8 | +8.1 | +||||||
| 29 | +
+ |
+11.6 | +10.3 | +10.5 | +11.9 | +6.5 | +7.0 | +6.5 | +7.9 | +7.1 | +8.1 | +9.8 | +6.7 | +|||||
| 30 | +
+ |
+12.9 | +11.7 | +12.0 | +13.3 | +7.4 | +7.8 | +7.4 | +8.9 | +8.2 | +9.7 | +11.0 | +7.8 | +1.8 | +||||
| 31 | +
+ |
+11.6 | +10.2 | +10.6 | +11.9 | +6.9 | +6.6 | +6.5 | +7.2 | +7.3 | +7.5 | +9.2 | +7.0 | +5.2 | +6.8 | +|||
| 32 | +
+ |
+10.7 | +9.5 | +9.2 | +9.9 | +6.0 | +6.3 | +5.7 | +6.4 | +6.5 | +6.7 | +8.6 | +6.5 | +6.2 | +7.4 | +5.6 | +||
| 33 | +
+ |
+10.3 | +9.5 | +9.2 | +10.2 | +7.9 | +7.8 | +7.7 | +8.3 | +9.0 | +9.2 | +10.8 | +8.4 | +8.7 | +9.8 | +8.0 | +6.8 | +|
| 34 | +
+ |
+10.0 | +9.1 | +9.1 | +10.0 | +8.0 | +8.3 | +7.1 | +7.7 | +9.2 | +9.0 | +10.8 | +7.9 | +8.5 | +9.4 | +7.5 | +6.3 | +2.8 | +
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|---|---|
|
+ |
+120.5–135.1 | +123.5–127.6 | +61.1–65.9 | +66.7–69.2 | +present | +beyond the snout | +
|
+ |
+118.9–145.7 | +103.8–165.4 | +59.4–74.8 | +51.0–96.1 | +present | +/ | +
|
+ |
+118.0-138.0 | +/ | +55.8–63.3 | +/ | +present | +/ | +
|
+ |
+110.4–140.3 | +138.9–162.5 | +61.1–72.6 | +79.5–90.4 | +absent | +/ | +
|
+ |
+/ | +134.9–155.5 | +/ | +72.6–82.4 | +present | +anterior corner of eyes | +
|
+ |
+/ | +/ | +64.7–83.6 | +70.5–100.3 | +present | +eyes to nostrils | +
|
+ |
+137.0–148.0 | +125.0–140.0 | +55.7–57.7 | +/ | +present | +eye | +
|
+ |
+110.1–146.5 | +138.6–154.2 | +64.2–82.0 | +80.8–88.0 | +present | +eye | +
|
+ |
+145.6–173.0 | +150.0–156.5 | +/ | +/ | +present | +/ | +
|
+ |
+151.0–172.0 | +142.7–170.5 | +76.8–85.2 | +76.3–87.4 | +present | +beyond the snout | +
|
+ |
+109.1–130.4 | +141.8 | +56.5–60.2 | +73.4 | +present | +/ | +
|
+ |
+/ | +/ | +/ | +71.6 | +present | +/ | +
|
+ |
+/ | +/ | +61.57 –70.69 | +76.8 | +present | +eye | +
|
+ |
+118.4–124.5 | +144.5 | +62.0 –66.5 | +79.3 | +present | +/ | +
|
+ |
+/ | +/ | +63.20–68.71 | +/ | +present | +nostril | +
|
+ |
+/ | +/ | +59.3–68.6 | +/ | +present | +/ | +
|
+ |
+113.0–121.8 | +/ | +48.3–53.6 | +/ | +absent | +/ | +
|
+ |
+126.0–133.0 | +/ | +61.9–77.3 | +/ | +absent | +/ | +
|
+ |
+/ | +/ | +54.4–68.3 | +70.8 | +present | +/ | +
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|---|
|
+ |
+overlapping | +slightly flattened | +not segmented | +absent | +absent | +
|
+ |
+meeting | +slightly flattened | +not segmented | +absent | +absent | +
|
+ |
+meeting or overlapping | +knob-like | +not segmented | +absent | +absent | +
|
+ |
+overlapping | +slightly flattened | +not segmented | +absent | +absent | +
|
+ |
+not touched | +slightly flattened | +not segmented | +absent | +absent | +
|
+ |
+overlapping | +slightly flattened | +not segmented | +absent | +absent | +
|
+ |
+meeting or overlapping | +slightly flattened | +not segmented | +absent | +absent | +
|
+ |
+not touched | +slightly flattened | +not segmented | +absent | +absent | +
|
+ |
+overlapping | +slightly flattened | +not segmented | +present | +absent | +
|
+ |
+overlapping | +knob-like | +not segmented | +absent | +absent | +
|
+ |
+/ | +knob-like | +not segmented | +present | +present | +
|
+ |
+/ | +knob-like | +not segmented | +present | +present | +
|
+ |
+/ | +knob-like | +not segmented | +present | +absent | +
|
+ |
+overlapping | +knob-like | +not segmented | +absent | +absent | +
|
+ |
+/ | +knob-like | +not segmented | +absent | +absent | +
|
+ |
+overlapping | +knob-like, in irregular series | +not segmented | +absent | +absent | +
|
+ |
+/ | +slightly flattened | +not segmented | +absent | +absent | +
|
+ |
+meeting or overlapping | +slightly flattened | +not segmented | +absent | +absent | +
|
+ |
+overlapping | +knob-like | +segmented tuberculate | +absent | +absent | +
| 2a | +
+ |
+
|
+2
+ |
+2 | +
| th | +3 | +
| th | +4 | +
|
+ |
+|
|
+ |
+|
|
+ |
+|
| 5 | +|
|
+th3
+ |
+6 | +
| th3e7a | +14 | +
|
+ |
+|
| 7 | +|
|
+ |
+|
|
+ |
+|
| th3b5a9a | +9 | +
|
+ |
+13 | +
| th | +
+ |
+
| th | +10 | +
| 11 | +|
| 12 | +|
| 5b | +
+ |
+
| 3b | +
+ |
+
|
+ |
+|
| 5d | +
+ |
+
| th3c | +
+ |
+
| th3d | +
+ |
+
| 3e | +
+ |
+
| 15 | +|
| 16 | +|
| 20 | +|
| th | +
+ |
+
| th | +17 | +
|
+ |
+|
| 18 | +|
|
+ |
+|
| 19 | +|
| th7e | +
+ |
+
| th | +
+ |
+
|
+ |
+|
|
+ |
+
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +sehr trocken | +Lichtzahl L | +sehr hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +
+montan (
+ |
+||
|
+ |
+
+ |
+
+ |
+kontinental (sehr niedrige Luftfeuchtigkeit, sehr grosse Temperaturschwankungen, kalte Winter) | +||
|
+ |
+
+ |
+
+ |
+
|---|---|---|
| Habit | +Prostrate | +Erect or decumbent | +
| Stem | +Leaf scars indistinct, internodes long, up to 3 cm | +Leaf scars prominent, internodes short, up to 1 cm | +
| Leaf | +Elliptic-lanceolate, nerves prominent at both surfaces, | +Linear-oblong to lanceolate, nerves prominent at abaxial | +
| grooved along the lateral nerves on the adaxial surface | +surface and indistinct on adaxial surface | +|
| Inflorescence | +Spikes sometimes aggregated at base, globose-ovoid, | +Spikes not aggregated at base, oblong, 3-6 cm long | +
| up to 2 cm long | +||
| Calyx | +7 × 3 mm, anterior sepals united � of the way | +9 × 4 mm, anterior sepals united � of the way | +
| Pollen | +
+Size 34.66 × 23.60 µm, exine thick
+ |
+
+Size 41.81 × 28.22 µm, exine thin
+ |
+
| Capsule | +6 × 2 mm | +6 × 4 mm | +
| Seeds | +Oblong, 3 × 1.5 mm | +Ovoid-suborbicular, 3.5 × 3 mm | +
|
+ |
+
+ |
+
+ |
+
|---|---|---|
|
+ |
+||
| Cuticle | +Thick | +Thin | +
| Epidermis | +Thick walled cells, variable in size and shape | +Thin walled cells, (almost) uniform in size and shape | +
| Hairs | +Simple non-glandular | +Four lobed glandular and simple non-glandular | +
| Cystoliths | +Present on both surfaces | +Present on abaxial surface | +
| Stomata type | +Anisocytic and diacytic | +Diacytic | +
| Midrib | +Parenchyma cells 3-layered | +Parenchyma cells 5-layered | +
| Vascular bundles | +Surrounded by 2-layered sclerenchyma cells | +Surrounded by 3-layered sclerenchyma cells | +
|
+ |
+||
| Medullary rays | +Uniseriate | +Uni and biseriate | +
| Intra-xylary Phloem | +Absent | +Present | +
| Pith parenchyma | +Thick walled with tannin | +Thin walled | +
| Acicular crystals | +Present | +Absent | +
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+13750 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +nicht beurteilt (Not Evaluated) | +|
| Mittelland (MP) | +nicht beurteilt (Not Evaluated) | +|
| Alpennordflanke (NA) | +nicht beurteilt (Not Evaluated) | +|
|
+ |
+nicht beurteilt (Not Evaluated) | +|
|
+ |
+nicht beurteilt (Not Evaluated) | +|
| Westliche Zentralalpen (WA) | +nicht beurteilt (Not Evaluated) | +
|
+ |
+-- | +
|
+ |
+-- | +
|
+ |
+-- | +
|
+ |
+-- | +
| Kein internationaler, nationaler oder kantonaler Schutz | +
| Character | +
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+|
|---|---|---|---|---|---|---|---|---|
| Holotype | +Paratypes | +(1-4)* | +(4, 5)* | +(6, 7)* | +(4, 5)* | +(4)* | +(8)* | +|
|
+6.3.5 - Hopfenbuchenwald (
+ |
+
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +schattig | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +warm-kollin | +||
|
+ |
+
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||
|
+ |
+0 - unbedeutend, keine Bindung. | +
| Ruhiges Wasser | +0 - unbedeutend, keine Bindung. | +
| Grundwasser | +0 - unbedeutend, keine Bindung. | +
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+231100 | +
| = | +
+ |
+
+ |
+1238 | +
| = | +
+ |
+
+ |
+582 | +
| = | +
+ |
+
+ |
+582 | +
| = | +
+ |
+
+ |
+231100 | +
| = | +
+ |
+
+ |
+1871 | +
| = | +
+ |
+
+ |
+1539 | +
| = | +
+ |
+
+ |
+231100 | +
| = | +
+ |
+
+ |
+843 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +-- | +|
| Mittelland (MP) | +-- | +|
| Alpennordflanke (NA) | +-- | +|
|
+ |
+
+stark
+ |
+C2a(i) | +
|
+ |
+-- | +|
| Westliche Zentralalpen (WA) | +-- | +
|
+ |
+
+3 - Mittlere nationale
+ |
+
|
+ |
+2 - Klarer Massnahmebedarf | +
|
+ |
+1 - Gering | +
|
+ |
+
+2 -
+ |
+
| Kein internationaler, nationaler oder kantonaler Schutz | +
|
+ |
+- | +weitere Informationen | +
| ♀ | +Fe | +
+ |
+Ti | +Mt | +Ta | +Total | +
|---|
| Interval (ms) | +326 ± 116 | +186%906 | +94 | +- | +- | +- | +51.20 ± 7.07 | +26%63 1340 | +
|---|---|---|---|---|---|---|---|---|
|
+ |
+Peak frequency | +Min frequency | +Max frequency | +Banđwiđth | +Quartile 25 | +Quartile 50 | +Quartile 75 | +Quartile (75%-25%) | +
| Mean ± SD Min%Max | +11.72 ± 0.79 7.21%14.25 | +4.14 ± 0.44 3.93%8.81 | +18.45 ± 1.74 15.46%23.81 | +14.30 ± 1.87 7.68%19.87 | +9.93 ± 0.56 7.59%10.96 | +11.50 ± 0.48 10.03%12.75 | +12.82 ± 0.45 11.25%14.34 | +2.89 ± 0.55 1.41%4.69 | +
| Type status | +Holotype | +Paratypes | +|||
|---|---|---|---|---|---|
| Specimen No. | +NCBS-BH723 | +NCBS-BH724 | +NCBS-BH725 | +NCBS-BH726 | +NCBS-BH727 | +
| Sex | +M | +F | +F | +F | +F | +
| SVL | +30.8 | +30.7 | +29.0 | +25.5 | +27.1 | +
| TL | +37.7 | +38.5 | +2.0* | +24.6 | +31.0 | +
| TW | +3.4 | +3.3 | +3.8 | +2.5 | +2.7 | +
| LAL | +4.2 | +4.3 | +3.8 | +3.6 | +3.8 | +
| CL | +5.0 | +4.9 | +4.7 | +4.6 | +4.7 | +
| AGL | +12.7 | +13.9 | +12.3 | +10.3 | +10.5 | +
| BH | +3.2 | +2.6 | +2.3 | +3.1 | +3.0 | +
| BW | +7.1 | +7.0 | +5.7 | +5.3 | +5.5 | +
| HL | +8.1 | +8.4 | +7.3 | +6.6 | +6.8 | +
| HW | +5.4 | +5.5 | +4.5 | +4.7 | +4.6 | +
| HD | +3.5 | +3.3 | +2.6 | +2.7 | +3.0 | +
| ED | +1.5 | +1.4 | +1.4 | +1.4 | +1.3 | +
| EE | +2.6 | +2.7 | +2.3 | +2.3 | +2.3 | +
| ES | +3.8 | +3.9 | +3.5 | +3.1 | +2.3 | +
| EN | +3.0 | +3.0 | +2.5 | +2.5 | +2.7 | +
| IN | +1.0 | +1.0 | +0.8 | +0.8 | +0.8 | +
| IO | +1.3 | +1.3 | +1.3 | +1.1 | +1.2 | +
| EL | +0.4 | +0.5 | +0.5 | +0.5 | +0.5 | +
| Type status | +Holotype | +Paratypes | +|||
|---|---|---|---|---|---|
| Specimen No. | +NCBS- BH723 | +NCBS- BH724 | +NCBS- BH725 | +NCBS- BH726 | +NCBS- BH727 | +
| SL L&R | +9&8 | +9&9 | +8&8 | +8&8 | +8&8 | +
| IL L&R | +7&6 | +8&7 | +6&6 | +7&7 | +7&7 | +
| SL M L&R | +6&6 | +6&6 | +6&6 | +6&6 | +6&6 | +
| IL M L&R | +5&5 | +5&5 | +5&5 | +5&5 | +5&5 | +
| PVT | +A | +A | +A | +A | +A | +
| DTR | +6 | +6 | +4 | +4 | +6 | +
| MVSR | +25 | +23 | +24 | +23 | +23 | +
| VS | +103 | +112 | +105 | +104 | +102 | +
| LamF1 L&R | +10&10 | +10&10 | +9&9 | +10&10 | +10&10 | +
| LamF4 L&R | +17&17 | +17&17 | +16&16 | +17&17 | +16&16 | +
| LamT1L&R | +10&10 | +9&9 | +9&9 | +10&10 | +10&10 | +
| LamT4 L&R | +20&20 | +19&19 | +19&19 | +18&19 | +17&19 | +
| LamT5 L&R | +18&18 | +16&17 | +16&15 | +16&15 | +17&17 | +
| PP L&R | +3 | +A | +A | +A | +A | +
| SB PP&FP L&R | +10&10 | +A | +A | +A | +A | +
| FP L&R | +1 | +A | +A | +A | +A | +
| Number of postcloacal tubercles | +1 | +1 | +1 | +1 | +1 | +
| Anterior extra-brillar fringe scales enlarged (1) or not enlarged (0) | +1 | +1 | +1 | +1 | +1 | +
| Ventral scales keeled (1) or smooth (0) | +0 | +0 | +0 | +0 | +0 | +
| Gular scales keeled (1) or smooth (0) | +0 | +0 | +0 | +0 | +0 | +
| Pectoral scales keeled (1) or smooth (0) | +0 | +0 | +0 | +0 | +0 | +
| Precloacal pores continuous (1) or separated (0) | +1 | +A | +A | +A | +A | +
| Precloacal pores elongate (1) or round (0) | +0 | +A | +A | +A | +A | +
| femoral pores elongate (1) or round (0) | +1 | +A | +A | +A | +A | +
| Dorsal pholidosis homogeneous (1) or heterogeneous (0) | +0 | +0 | +0 | +0 | +0 | +
| Dorsal tubercles keeled (1) or not keeled (0) | +1 | +1 | +1 | +1 | +1 | +
| Tubercles linearly arranged (1) or more random (0) | +1 | +1 | +1 | +1 | +1 | +
| Spine-like tubercles on flank present (1) or absent (0) | +1 | +1 | +1 | +1 | +1 | +
| Lateral caudal furrows present (1) or absent (0) | +1 | +1 | +1 | +1 | +1 | +
| Caudal tubercles in lateral furrow (1) or not (0) | +1 | +1 | +1 | +1 | +1 | +
| Subcaudals keeled (1) or smooth (0) | +0 | +0 | +0 | +0 | +0 | +
| Single median row of keeled subcaudals (1) or smooth (0) | +0 | +0 | +0 | +0 | +0 | +
| Caudal tubercles encircle tail (1) or not (0) | +1 | +1 | +1 | +1 | +1 | +
| Slightly enlarged median subcaudal scale row (1) or not (0) | +1 | +1 | +1 | +1 | +1 | +
| Enlarged femoral scales present (1) or absent (0) | +1 | +1 | +1 | +1 | +1 | +
| 1 | +2 | +3 | +4 | +5 | +6 | +7 | +8 | +9 | +10 | +11 | +12 | +13 | +14 | +|
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
|
+ |
+4–8 | +1 (2) | +– | ++ | ++ | ++ | +e | +– | ++ | ++ | +24 | +~40 | +a | +C | +
|
+ |
+3–5 | +3(0–5) | ++ | +– | +– | +– | +r | ++ | +– | +– | +50 | +6–50 | +b | +NE | +
|
+ |
+2–3 | +3 (5,6) | ++ | +– | ++ | +– | +e | ++ | ++ | +– | +24 | +11–18 | +c | +B | +
|
+ |
+1.1–1.4 | +3 | +– | ++ | +– | +– | +r | ++ | +– | +– | +33 | +2 | +d | +NC | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +
+kollin (
+ |
+||
|
+ |
+
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||
| Leg | +Cox. | +Troc. | +Fem. | +Pat. | +Tib. | +Met | +Tars | +Total | +
|---|---|---|---|---|---|---|---|---|
| I | +1.12 | +0.54 | +3.52 | +1.66 | +3.22 | +2.44 | +1.76 | +14.26 | +
| II | +1.12 | +0.54 | +2.17 | +1.46 | +2.83 | +2.44 | +1.76 | +13.32 | +
| III | +1.07 | +0.54 | +2.68 | +1.46 | +2.35 | +2.64 | +1.76 | +12.50 | +
| IV | +1.12 | +0.54 | +3.76 | +1.46 | +3.42 | +4.15 | +2.30 | +16.75 | +
| Palp | +----- | +0.34 | +1.37 | +0.73 | +0.93 | +----- | +1.46 | +4.83 | +
| Fem | +d | +v | +pl | +rl | +Tib | +d | +v | +pl | +rl | +Met | +d | +v | +pl | +rl | +
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| I | +1 1 0 | +0 | +1 1 1 | +0 | +I | +0 | +2222 | +0 | +0 | +I | +0 | +200 | +0 | +0 | +
| II | +1 1 0 | +0 | +1 1 1 | +0 | +II | +0 | +2222 | +0 | +0 | +II | +0 | +200 | +0 | +0 | +
| III | +1 1 0 | +0 | +0 1 1 | +0 1 1 | +III | +11 | +222 | +110 | +110 | +III | +11 | +22 | +111 | +111 | +
| IV | +1 1 1 | +0 | +1 1 1 | +0 1 1 | +IV | +111 | +222 | +1 1 | +1 1 | +IV | +111 | +222 | +1 1 | +1 1 | +
| Leg Cox. | +Troc. | +Fem. | +Pat. | +Tib. | +Met. | +Tars. | +Total | +
|---|---|---|---|---|---|---|---|
| I 1.08 II 1.03 III 1.03 IV 1.03 Palp ----- | +0.44 0.44 0.44 0.44 0.34 | +2.93 2.59 2.29 3.27 1.12 | +1.40 1.22 1.22 1.22 0.63 | +2.83 2.59 2.29 3.27 0.83 | +2.15 2.15 2.35 3.76 ----- | +1.56 1.56 1.56 2.15 1.12 | +12.39 11.63 11.04 14.95 4.04 | +
|
+ |
+
| Fem | +d | +V | +pl | +rl | +Tib | +d | +v | +pl | +rl | +Met | +d | +v | +pl | +rl | +
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| I | +1 1 0 | +0 | +0 1 1 | +0 | +I | +0 | +2222 | +0 | +0 | +I | +0 | +200 | +0 | +0 | +
| II | +1 1 0 | +0 | +1 1 1 | +0 | +II | +0 | +2222 | +0 | +0 | +II | +0 | +200 | +0 | +0 | +
| III | +1 1 0 | +0 | +0 1 1 | +0 1 1 | +III | +11 | +222 | +11 | +11 | +III | +11 | +22 | +11 | +11 | +
| IV | +1 1 0 | +0 | +1 1 1 | +1 1 1 | +IV | +111 | +222 | +11 | +11 | +IV | +111 | +222 | +11 | +11 | +
| Leg Cox. | +Troc. | +Fem. | +Pat. | +Tib. | +Met | +Tars | +Total | +
|---|---|---|---|---|---|---|---|
| I 0.88 II 0.83 III 0.83 IV 0.88 Palp ----- | +0.29 0.29 0.29 0.29 0.24 | +2.50 2.29 2.05 2.83 0.88 | +1.12 1.12 0.97 1.12 0.39 | +2.50 2.29 1.80 2.73 0.34 | +2.20 2.25 1.15 3.42 ----- | +1.50 1.50 1.50 2.00 0.83 | +10.99 10.57 9.59 13.27 2.68 | +
| Leg spination | +
| Fem | +d | +V | +pl | +rl | +Tib | +d | +v | +pl | +rl | +Met | +d | +v | +pl | +rl | +
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| I | +1 1 1 | +0 | +1 1 | +1 1 | +I | +1 | +2222 | +11 | +11 | +I | +0 | +200 | +0 | +0 | +
| II | +1 1 1 | +0 | +1 1 | +1 1 | +II | +111 | +2222 | +111 | +11 | +II | +0 | +200 | +0 | +0 | +
| III | +1 1 1 | +0 | +1 1 | +1 1 | +III | +11 | +222 | +11 | +11 | +III | +11 | +222 | +11 | +11 | +
| IV | +1 1 1 | +0 | +1 1 | +1 1 1 | +IV | +11 | +222 | +11 | +11 | +IV | +111 | +222 | +11 | +11 | +
| 52' | +
| 53 | +
|
+ |
+
|
+ |
+
| 1 | +Leaf blades distinctly scabrous on their margins; spikelets tightly appressed and strongly overlapping | +2 | +
| - | +Leaf blades glabrous on their margins (occasionally with a few scattered teeth, but teeth never abundant); spikelets weakly appressed and weakly overlapping | +5 | +
| 2 | +Culms growing in tufts (i.e., cespitose) from hard knotty bases, rhizomes absent, rarely short; branches appressed, usually not readily discernible within an inflorescence, not distinctly one-sided | +
+ |
+
| - | +Culms growing from rhizomes; branches appressed, ascending, or spreading, readily discernible within an inflorescence, distinctly one-sided | +3 | +
| 3 | +Upper glumes distinctly awned, awns 3-8 mm long; blades 5-15 mm wide; ligules 1-3 mm long; branches pedunculate, rarely sessile | +
+ |
+
| - | +Upper glumes unawned or short-awned, when present awns to 2 mm long; blades 0.5-8 mm wide; ligules 0.5-1 mm long; branches sessile, rarely pedunculate | +4 | +
|
+ |
+Glume keels ciliate, hairs stiff, 0.5-0.8(-1) mm long; glumes with two inconspicuous lateral veins on one side of the keel; branches appressed to the main axis; most branches 3-6 mm wide; inflorescences 8-25 cm long; spikelets ovate to lanceolate; florets more or less equaling the upper glumes in length | +
+ |
+
| - | +Glume keels scabrous, teeth 0.1-0.2 mm long; glumes with two conspicuous lateral veins on one side of the keel; branches appressed, ascending, or spreading from main axis; most branches 2-2.5 mm wide; inflorescences 3-15 cm long; spikelets linear lanceolate to ovate lanceolate; florets shorter than the upper glumes | +
+ |
+
| 5 | +Spikelets 8-14(-16.5) mm long; branch rachises 0.4-1 mm wide between spikelets; glumes glabrous or weakly pubescent; leaf blades more or less erect, forming an angle 15-18° with the culm | +
+ |
+
| - | +Spikelets 14-25 mm long; branch rachises 1-2.2 mm wide between spikelets; glumes moderately to densely pubescent; leaf blades ascending to spreading, forming an angle 30-60° with the culm | +6 | +
| 6 | +Spikelets (15-)16.5-25 mm long; anthers 7-10 mm long, usually fully exserted at maturity; pollen fertile; ligules 1-3 mm long; upper glumes 3-6-veined, 13-22 mm long; glumes (weakly) moderately to densely pubescent with hairs 0.1-0.3 mm long, hairs to 0.6 mm long and usually denser proximally; calluses (1.5-)2-4.5 mm long; branches (3-)4-5(-6) mm wide | +
+ |
+
| - | +Spikelets 14-17.5 mm long; anthers 5-7(-8.5) mm long, not or incompletely exserted at maturity, indehiscent; pollen sterile; ligules 1-1.5 mm long; upper glumes 3-veined, 12.5-16.5 mm long; glumes weakly to moderately pubescent with hairs 0.1-0.2 mm long, occasionally to 0.6 mm long proximally; calluses 0.6-1.5(-2) mm long; branches (2.5-)3-4 mm wide | +
+ |
+
|
+ |
+
+ |
+
+ |
+
|---|---|---|
| Shell | +Semiglobose-globose peripheral keel absent fine radial growth lines | +Semidepressed-depressed peripheral keel present distinct radial ribs | +
| Vas deferens | +Short, entering epiphallus subapically | +Long, entering epiphallus apically | +
| Epiphallus | +Slender and elongate | +Swollen | +
| Penis | +Clavate | +Short cylindrical | +
| Vagina | +Outer surface with longitudinal rugose sculpture | +Outer surface smooth | +
| Gland of amatorial organ | +Ovate The lobes fuse as very thin sac Consists of 5–6 clusters of internal ducts | +Crescent-shaped The lobes fuse as very thick sac Consists of 2–4 clusters of internal ducts | +
| Radula: marginal teeth | +Short, finger shaped with blunt tip | +Elongate, slender finger shaped with sharp pointed tip | +
| Standard length (mm) | +130.7 | +29 | +75.0–337.5 | +261.7 | +* | +
|---|---|---|---|---|---|
| Percentage of standard length | +|||||
| Body depth | +60.6 | +28 | +53.0–64.5 | +58.9 | +2.5 | +
| Head length | +24.4 | +28 | +22.5–25.4 | +24.0 | +0.8 | +
| Supraoccipital process length | +29.9 | +28 | +24.0–30.7 | +27.2 | +2.1 | +
| Predorsal length | +54.5 | +28 | +47.4–57.6 | +53.3 | +1.9 | +
| Dorsal-fin base length | +30.4 | +28 | +23.2–32.4 | +30.1 | +1.7 | +
| Interdorsal length | +13.6 | +28 | +6.6–14.1 | +12.8 | +1.4 | +
| Adipose-fin base length | +6.4 | +28 | +5.5–9.2 | +6.2 | +0.7 | +
| Caudal-peduncle depth | +10.6 | +28 | +9.1–10.7 | +9.9 | +0.5 | +
| Anal-fin base length | +33.5 | +28 | +30.2–34.7 | +32.8 | +1.0 | +
| Preanal length | +74.2 | +28 | +73.9–81.3 | +76.6 | +1.8 | +
| Prepelvic length | +56.5 | +28 | +55.1–61.8 | +57.3 | +1.5 | +
| Prepectoral length | +25.3 | +28 | +24.0–29.5 | +25.9 | +1.3 | +
| Anal-pelvic distance | +20.5 | +28 | +19.7–27.3 | +21.7 | +1.5 | +
| Pelvic-pectoral distance | +31.1 | +28 | +29.6–35.1 | +32.2 | +1.3 | +
| Width of peduncle | +3.3 | +28 | +2.8–5.0 | +4.0 | +0.5 | +
| Pectoral-fin length | +19.6 | +28 | +14.8–21.7 | +19.3 | +1.3 | +
| Pelvic-fin length | +15.4 | +27 | +11.6–15.6 | +14.1 | +0.9 | +
| First anal-fin lobe length | +22.9 | +21 | +20.1–36.3 | +28.1 | +4.5 | +
| Second anal-fin lobe length | +* | +6 | +14.9–19.4 | +16.6 | +1.7 | +
| Dorsal-fin length | +29.2 | +27 | +20.1–32.1 | +25.4 | +3.0 | +
| First oblique depth | +66.2 | +28 | +61.2–68.7 | +64.4 | +1.9 | +
| Second oblique depth | +49.2 | +28 | +44.6–50.5 | +47.5 | +1.6 | +
| Third oblique depth | +27.3 | +28 | +24.9–27.4 | +26.1 | +0.7 | +
| Percentage of head length | +|||||
| Snout length | +27.0 | +28 | +25.5–37.2 | +31.2 | +3.4 | +
| Interorbital width | +52.0 | +28 | +44.8–58.5 | +53.7 | +3.6 | +
| Width head | +63.1 | +27 | +58.5–73.0 | +65.9 | +3.5 | +
| Postorbital distance | +30.2 | +28 | +28.1–36.9 | +32.9 | +2.4 | +
| Fourth infraorbital width | +14.7 | +28 | +11.5–17.8 | +15.2 | +1.3 | +
| Eye vertical diameter | +40.2 | +28 | +29.8–42.9 | +33.6 | +3.7 | +
| Mouth length | +18.5 | +28 | +14.2–20.7 | +18.0 | +1.4 | +
| Third infraorbital width | +9.6 | +28 | +8.1–13.9 | +11.0 | +1.3 | +
| Cheek gap width | +11.0 | +28 | +8.5–13.9 | +10.5 | +1.3 | +
| Mouth width | +31.0 | +28 | +29.9–38.5 | +35.5 | +2.5 | +
| Legs | +Coxa | +Basis | +Exopod | +Endopod | +
|---|
| 2.5 - Wechselfeuchte Pionierfluren | +
|
+ |
+
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+sehr feucht; Feuchtigkeit
+ |
+Lichtzahl L | +halbschattig | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +
+kollin (
+ |
+||
|
+ |
+
+ |
+
+ |
+subozeanisch (hohe Luftfeuchtigkeit, geringe Temperaturschwankungen, eher milde Winter) | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+315300 | +
| = | +
+ |
+
+ |
+459 | +
| = | +
+ |
+
+ |
+1273 | +
| = | +
+ |
+
+ |
+1273 | +
| = | +
+ |
+
+ |
+315300 | +
| = | +
+ |
+
+ |
+905 | +
| = | +
+ |
+
+ |
+787 | +
| = | +
+ |
+
+ |
+315300 | +
| = | +
+ |
+
+ |
+166 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +
+potenziell
+ |
+B2b(iii) | +
| Mittelland (MP) | +
+potenziell
+ |
+B2b(iii) | +
| Alpennordflanke (NA) | +
+potenziell
+ |
+B2b(iii) | +
|
+ |
+
+potenziell
+ |
+B2b(iii) | +
|
+ |
+
+ |
+|
| Westliche Zentralalpen (WA) | +
+potenziell
+ |
+B2b(iii) | +
|
+Keine nationale
+ |
+
|
+ |
+Nein | +|
|
+ |
+
+ |
+(25.07.2007) | +
|
+ |
+-- | + + +
| KEINE ANGABE | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +
+montan (
+ |
+||
|
+ |
+
+ |
+
+ |
+
+subozeanisch bis subkontinental (mittlere Luftfeuchtigkeit,
+ |
+||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+236400 | +
| = | +
+ |
+
+ |
+2129 | +
| = | +
+ |
+
+ |
+236400 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +
+ |
+|
| Mittelland (MP) | +
+ |
+|
| Alpennordflanke (NA) | +
+ |
+|
|
+ |
+
+ |
+|
|
+ |
+
+ |
+|
| Westliche Zentralalpen (WA) | +
+ |
+
|
+Keine nationale
+ |
+
| Kein internationaler, nationaler oder kantonaler Schutz | +
| mm | +% BDL | +mm | +% BDL | +|
|---|---|---|---|---|
| CDH | +32.2 | +4 | +25.7–38.1 | +3–5 | +
| CVM | +305 | +39 | +242.3–299.3 | +30–36 | +
| CVH | +29.26 | +4 | +21.7–31.2 | +3–4 | +
| CTL | +260 | +33 | +209.7–267.3 | +25–32 | +
| CLT | +151.5 | +19 | +151.5–164.2 | +19–20 | +
| CLM | +25.3 | +3 | +30.43–36 | +3–4 | +
| CLL | +37.7 | +5 | +28.6–37.7 | +3–4 | +
| CLO | +46.8 | +6 | +107.7–121.1 | +14 | +
| CLI | +113.1 | +14 | +113.1–152.1 | +17–20 | +
| CLB | +111 | +14 | +20.5–25 | +2–3 | +
| FTL | +41.8 | +5 | +36.6–42.4 | +4–5 | +
| TBH | +11.6 | +1 | +6.9–12.7 | +1 | +
| TBL | +17.5 | +2 | +11.2–20.7 | +1–2 | +
| TBW | +11.1 | +1 | +8.3–12.1 | +1 | +
| TsW | +7.5 | +1 | +4.6–7.3 | +1 | +
| ONC | +25.3 | +3 | +19.5–32 | +3–4 | +
| LRC | +13 | +2 | +11.1–25 | +1–3 | +
| LNC | +54.4 | +7 | +26.2–64.3 | +3–7 | +
| IOA | +26.1 | +3 | +5.3–38.6 | +1–4 | +
| OTM | +77.2 | +10 | +73.1–88.8 | +8–11 | +
| OCL | +26 | +3 | +31.4–54.5 | +4–6 | +
| STL | +50.6 | +6 | +39.1–74 | +5–8 | +
| SPS | +21.7 | +3 | +36.7–57 | +4–6 | +
|
+ |
+
+ |
+
+ |
+
+ |
+||
|---|---|---|---|---|---|
|
+ |
+
+ |
+||||
| Holotype | +Paratype | +Lectotype | +Lectotype | +Holotype | +|
| Total length | +27.3 | +28.6 | +44.0 | +33.9 | +31.0 | +
| Carapace: | +|||||
| length | +3.9 | +4.1 | +5.2 | +4.3 | +4.1 | +
| anterior width | +2.8 | +3.2 | +3.8 | +3.2 | +2.9 | +
| posterior width | +3.6 | +4.5 | +5.5 | +4.7 | +4.3 | +
| Metasomal segment I: | +|||||
| length | +2.1 | +2.6 | +2.8 | +2.3 | +2.0 | +
| width | +2.2 | +2.5 | +3.0 | +2.7 | +2.5 | +
| Metasomal segment V: | +|||||
| length | +4.7 | +5.2 | +6.2 | +5.2 | +4.9 | +
| width | +1.7 | +2.1 | +2.5 | +2.2 | +1.9 | +
| depth | +1.7 | +2.1 | +2.4 | +2.1 | +1.9 | +
| Vesicle: | +|||||
| width | +1.2 | +1.4 | +2.0 | +1.8 | +1.4 | +
| depth | +1.4 | +1.5 | +1.9 | +1.8 | +1.5 | +
| Pedipalp: | +|||||
| Femur length | +3.8 | +4.3 | +4.8 | +4.0 | +3.8 | +
| Femur width | +1.1 | +1.2 | +1.4 | +1.2 | +1.1 | +
| Patella length | +4.3 | +5.1 | +5.6 | +4.5 | +4.3 | +
| Patella width | +1.4 | +1.6 | +2.1 | +1.6 | +1.5 | +
| Chela length | +6.1 | +6.9 | +8.4 | +6.7 | +6.8 | +
| Chela width | +1.0 | +1.1 | +1.7 | +1.3 | +1.3 | +
| Chela depth | +1.0 | +1.1 | +1.6 | +1.3 | +1.3 | +
| Movable finger: | +|||||
| length | +4.2 | +4.9 | +5.9 | +4.8 | +4.5 | +
|
+4.2 -
+ |
+
|
+4.2.4 -
+ |
+
|
+4.3.6 - Buntschwingelhalde (
+ |
+
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +trocken | +Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +warm-kollin | +||
|
+ |
+
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+135600 | +
| = | +
+ |
+
+ |
+405 | +
| = | +
+ |
+
+ |
+1257 | +
| = | +
+ |
+
+ |
+1257 | +
| = | +
+ |
+
+ |
+135600 | +
| = | +
+ |
+
+ |
+135600 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +
+potenziell
+ |
+B2b(iii) | +
| Mittelland (MP) | +verletzlich (Vulnerable) | +A3c | +
| Alpennordflanke (NA) | +
+potenziell
+ |
+A3c | +
|
+ |
+
+nicht
+ |
+|
|
+ |
+
+nicht
+ |
+|
| Westliche Zentralalpen (WA) | +
+nicht
+ |
+
|
+Keine nationale
+ |
+
|
+ |
+Nein | +|
|
+ |
+
+ |
+(01.01.2010) | +
|
+ |
+
+ |
+(01.01.2016) | +
|
+ |
+
+Teilweise
+ |
+(06.12.1978) | +
|
+ |
+
+Teilweise
+ |
+(01.08.2013) | +
|
+ |
+-- | +|
|
+ |
+
+ |
+(06.03.1979) | +
|
+ |
+
+ |
+(01.01.2018) | +
|
+ |
+
+ |
+(03.12.1964) | +
|
+ |
+
+ |
+(01.01.2012) | +
|
+ |
+L - Leitart | +weitere Informationen | +
| KEINE ANGABE | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+388630 | +
|
+ |
+|
|
+2.
+ |
+
+ |
+
|
+2*.
+ |
+
+ |
+
|
+1*.
+ |
+|
|
+3.
+ |
+
+ |
+
|
+3*.
+ |
+
+ |
+
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +schattig | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +warm-kollin | +||
|
+ |
+
+ |
+
+ |
+
+subozeanisch bis subkontinental (mittlere Luftfeuchtigkeit,
+ |
+||
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+feucht; Feuchtigkeit stark wechselnd (mehr als
+ |
+Lichtzahl L | +hell | +Salzzeichen | +1 | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +warm-kollin | +||
|
+ |
+
+ |
+
+ |
+subozeanisch (hohe Luftfeuchtigkeit, geringe Temperaturschwankungen, eher milde Winter) | +||
| Morphological feature | +
+ |
+||||||||||||
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| Adult ♂ (HT) | +Adult ♂ | +Adult ♂ | +Adult ♂ | +Adult ♂ | +Adult ♂ | +Adult ♂ | +Adult ♀ | +Adult ♀ | +Adult ♀ | +Adult ♀ | +|||
| DECBA2109 | +DECBA1967 | +DECBA0801 | +DECBA1958 | +DECBA2182 | +DECBA2092 | +DECBA2039 | +DECBA1787 | +DECBA1974 | +DECBA1793 | +DECBA2074 | +|||
| Holotype | +Paratypes | +||
|---|---|---|---|
| male | +males (6) | +females (4) | +|
| Standard length (mm) | +36.9 | +29.7-36.1 | +26.6-30.0 | +
| Percent of standard length | +|||
| Percent of head length | +|||
| NLNBNSQCONMBSKABYTNTBC | +
|
+ |
+
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Character | +
+ |
+
+ |
+Z | +p | +
|---|---|---|---|---|
| Scales between second canthals | +||||
| Postrostrals | +||||
| Loreal rows | +1.618E-05 | +|||
| Scales between supraorbital semicircles | +||||
| Scales between interparietal and semicircles | +0.01639 | +|||
| Supralabials to below center of eye | +2.19E-08 | +|||
| Postmentals | +||||
|
+Lamellae under phalanges
+ |
+0.01107 | +
| Character | +
+ |
+
+ |
+F | +p | +t | +p | +
|---|---|---|---|---|---|---|
| 0.03328 | +||||||
| 0.003981 | +||||||
| 0.05 | +||||||
| 0.001056 | +||||||
| 0.003917 | +||||||
| 0.00431 | +
| Character | +Males | +Females | +
|---|---|---|
| N = 21 | +N = 13 | +
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +sehr trocken | +Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +warm-kollin | +||
|
+ |
+
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+174700 | +
| = | +
+ |
+
+ |
+523 | +
| = | +
+ |
+
+ |
+835 | +
| = | +
+ |
+
+ |
+835 | +
| = | +
+ |
+
+ |
+174700 | +
| = | +
+ |
+
+ |
+2049 | +
| = | +
+ |
+
+ |
+1672 | +
| = | +
+ |
+
+ |
+174700 | +
| = | +
+ |
+
+ |
+1075 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +verletzlich (Vulnerable) | +A3c | +
| Mittelland (MP) | +
+stark
+ |
+A3c | +
| Alpennordflanke (NA) | +verletzlich (Vulnerable) | +B2ab(iii) | +
|
+ |
+
+potenziell
+ |
+B2b(iii) | +
|
+ |
+verletzlich (Vulnerable) | +B2ab(iii) | +
| Westliche Zentralalpen (WA) | +
+potenziell
+ |
+B2b(iii) | +
|
+Keine nationale
+ |
+
|
+ |
+Nein | +|
|
+ |
+
+ |
+(02.03.2005) | +
|
+ |
+-- | +|
|
+ |
+
+ |
+(25.07.2007) | +
|
+ |
+L - Leitart | +weitere Informationen | +
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +sauer (pH 3.5-6.5) | +Temperaturzahl T | +
+montan (
+ |
+||
|
+ |
+
+sehr
+ |
+
+ |
+
+subozeanisch bis subkontinental (mittlere Luftfeuchtigkeit,
+ |
+||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+89600 | +
| = | +
+ |
+
+ |
+2565 | +
| = | +
+ |
+
+ |
+2741 | +
| = | +
+ |
+
+ |
+2741 | +
| = | +
+ |
+
+ |
+89600 | +
| = | +
+ |
+
+ |
+534 | +
| = | +
+ |
+
+ |
+470 | +
| = | +
+ |
+
+ |
+89600 | +
| = | +
+ |
+
+ |
+2499 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +verletzlich (Vulnerable) | +A3c; B2ab(iii) | +
| Mittelland (MP) | +
+stark
+ |
+A3c; B2ab(iii) | +
| Alpennordflanke (NA) | +
+potenziell
+ |
+B2b(iii) | +
|
+ |
+verletzlich (Vulnerable) | +D2 | +
|
+ |
+
+potenziell
+ |
+B2b(iii) | +
| Westliche Zentralalpen (WA) | +
+stark
+ |
+B2ab(iii) | +
|
+Keine nationale
+ |
+
|
+ |
+Nein | +|
|
+ |
+
+ |
+(02.03.2005) | +
|
+ |
+-- | + + +
| 4.5 - Fettwiesen und -weiden | +
|
+ |
+
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +feucht | +Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +
+montan (
+ |
+||
|
+ |
+
+ |
+
+ |
+
+subozeanisch bis subkontinental (mittlere Luftfeuchtigkeit,
+ |
+||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+328100 | +
| = | +
+ |
+
+ |
+1645 | +
| = | +
+ |
+
+ |
+1595 | +
| = | +
+ |
+
+ |
+1595 | +
| = | +
+ |
+
+ |
+328100 | +
| = | +
+ |
+
+ |
+2494 | +
| = | +
+ |
+
+ |
+2026 | +
| = | +
+ |
+
+ |
+328100 | +
| = | +
+ |
+
+ |
+1416 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +
+nicht
+ |
+|
| Mittelland (MP) | +
+nicht
+ |
+|
| Alpennordflanke (NA) | +
+nicht
+ |
+|
|
+ |
+
+nicht
+ |
+|
|
+ |
+
+nicht
+ |
+|
| Westliche Zentralalpen (WA) | +
+nicht
+ |
+
|
+Keine nationale
+ |
+
| Kein internationaler, nationaler oder kantonaler Schutz | +
| - | +
+ |
+
+ |
+
+ |
+
|
+ |
++ | +- | ++ | +
|
+ |
++ | +- | ++ | +
|
+ |
++ | ++ | +- | +
|
+ |
++ | ++ | +- | +
|
+ |
++ | ++ | ++ | +
|
+ |
++ | +- | +- | +
|
+ |
+- | ++ | ++ | +
|
+ |
++ | ++ | +- | +
|
+ |
++ | ++ | +- | +
|
+ |
++ | +- | ++ | +
|
+ |
++ | ++ | +- | +
|
+ |
++ | +- | +- | +
|
+ |
++ | ++ | ++ | +
|
+ |
+- | ++ | ++ | +
|
+ |
++ | +- | +- | +
|
+ |
++ | ++ | ++ | +
|
+ |
++ | ++ | +- | +
|
+ |
++ | ++ | +- | +
| - | +
+ |
+
+ |
+
+ |
+||
| - | +
+ |
+
+ |
+
+ |
+||
| Standard length (mm) | +80.5 | +62.4 | +74.5 | +77.3 | +62.4-80.5 | +
| % SL | +- | +- | +- | +- | +- | +
| Predorsal length | +40.4 | +41.2 | +39.1 | +39.6 | +39.1-41.2 | +
| Preanal length | +71.1 | +71.3 | +70.9 | +71.7 | +70.9-71.7 | +
| Prepelvic length | +55.2 | +55.4 | +58.4 | +57.2 | +55.2-58.4 | +
| Prepectoral length | +25.2 | +26.1 | +26.7 | +26.1 | +25.2-26.7 | +
| Length of dorsal-fin base | +17.3 | +18.9 | +15.4 | +15.7 | +15.4-18.9 | +
| Length of dorsal spine | +11.9 | +16.7 | +12.2 | +12.8 | +11.9-16.7 | +
| Length of anal-fin base | +11.1 | +13.5 | +12.8 | +13.5 | +11.1-13.5 | +
| Pelvic-fin length | +16.4 | +16.8 | +16.2 | +16.8 | +16.2-16.8 | +
| Pectoral-fin length | +19.3 | +23.1 | +20.0 | +19.4 | +19.3-23.1 | +
| Pectoral-spine length | +17.6 | +20.4 | +18.7 | +16.2 | +16.2-20.4 | +
| Caudal-fin length | +30.4 | +28.7 | +26.0 | +26.9 | +26.0-30.4 | +
| Length of adipose-fin base | +20.9 | +20.3 | +22.8 | +23.3 | +20.3-23.3 | +
| Dorsal to adipose distance | +12.5 | +8.5 | +10.6 | +8.9 | +8.5-12.5 | +
| Post-adipose distance | +14.2 | +14.7 | +14.1 | +15.3 | +14.1-15.3 | +
| Length of caudal peduncle | +18.4 | +16.1 | +16.9 | +17.5 | +16.1-18.4 | +
| Depth of caudal peduncle | +11.1 | +9.9 | +10.2 | +9.8 | +9.8-11.1 | +
| Body depth at anus | +20.7 | +21.5 | +23.1 | +21.0 | +20.7-23.1 | +
| Head length | +27.8 | +31.6 | +29.3 | +28.5 | +27.8-31.6 | +
| Head width | +20.1 | +19.4 | +20.7 | +19.7 | +19.4-20.7 | +
| Head depth | +18.9 | +19.7 | +19.3 | +19.8 | +18.9-19.8 | +
| % HL | +- | +- | +- | +- | +- | +
| Snout length | +35 | +33 | +34 | +37 | +33-37 | +
| Interorbital distance | +42 | +37 | +35 | +36 | +35-42 | +
| Eye diameter | +17 | +24 | +20 | +20 | +17-24 | +
| Nasal barbel length | +68 | +55 | +66 | +77 | +55-77 | +
| Maxillary barbel length | +261 | +290 | +273 | +295 | +261-295 | +
| Inner mandibular barbel length | +104 | +83 | +96 | +98 | +83-104 | +
| Outer mandibular barbel length | +150 | +189 | +136 | +144 | +136-189 | +
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+sehr feucht; Feuchtigkeit stark wechselnd (mehr als
+ |
+Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +unter-montan und ober-kollin | +||
|
+ |
+
+ |
+
+ |
+subozeanisch (hohe Luftfeuchtigkeit, geringe Temperaturschwankungen, eher milde Winter) | +||
|
+ |
+0 - unbedeutend, keine Bindung. | +
| Ruhiges Wasser | +2 - Schwerpunktlebensraum | +
| Grundwasser | +0 - unbedeutend, keine Bindung. | +
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+398900 | +
| = | +
+ |
+
+ |
+605 | +
| = | +
+ |
+
+ |
+843 | +
| = | +
+ |
+
+ |
+843 | +
| = | +
+ |
+
+ |
+398900 | +
| = | +
+ |
+
+ |
+398900 | +
| = | +
+ |
+
+ |
+457 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +vom Aussterben bedroht (Critically Endangered) | +B2ab(iii,v); C2a(i)b | +
| Mittelland (MP) | +-- | +|
| Alpennordflanke (NA) | +-- | +|
|
+ |
+-- | +|
|
+ |
+-- | +|
| Westliche Zentralalpen (WA) | +-- | +
|
+ |
+
+1 - Sehr hohe nationale
+ |
+
|
+ |
+2 - Klarer Massnahmebedarf | +
|
+ |
+2 - Mittel | +
|
+ |
+
+2 -
+ |
+
|
+ |
+Ja | +|
|
+ |
+
+ |
+(02.03.2005) | +
|
+ |
+
+ |
+
+
+
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+356000 | +
| = | +
+ |
+
+ |
+356000 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +nicht beurteilt (Not Evaluated) | +|
| Mittelland (MP) | +nicht beurteilt (Not Evaluated) | +|
| Alpennordflanke (NA) | +nicht beurteilt (Not Evaluated) | +|
|
+ |
+nicht beurteilt (Not Evaluated) | +|
|
+ |
+nicht beurteilt (Not Evaluated) | +|
| Westliche Zentralalpen (WA) | +nicht beurteilt (Not Evaluated) | +
|
+Keine nationale
+ |
+
| Kein internationaler, nationaler oder kantonaler Schutz | +
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+20200 | +
| = | +
+ |
+
+ |
+20200 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +nicht beurteilt (Not Evaluated) | +|
| Mittelland (MP) | +nicht beurteilt (Not Evaluated) | +|
| Alpennordflanke (NA) | +nicht beurteilt (Not Evaluated) | +|
|
+ |
+nicht beurteilt (Not Evaluated) | +|
|
+ |
+nicht beurteilt (Not Evaluated) | +|
| Westliche Zentralalpen (WA) | +nicht beurteilt (Not Evaluated) | +
|
+Keine nationale
+ |
+
| Kein internationaler, nationaler oder kantonaler Schutz | +
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +frisch | +Lichtzahl L | +sehr hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +
+unter-alpin, supra-subalpin und ober-subalpin (
+ |
+||
|
+ |
+
+ |
+
+ |
+
+subozeanisch bis subkontinental (mittlere Luftfeuchtigkeit,
+ |
+||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+339900 | +
| = | +
+ |
+
+ |
+171 | +
| = | +
+ |
+
+ |
+188 | +
| = | +
+ |
+
+ |
+188 | +
| = | +
+ |
+
+ |
+339900 | +
| = | +
+ |
+
+ |
+1168 | +
| = | +
+ |
+
+ |
+1008 | +
| = | +
+ |
+
+ |
+339900 | +
| = | +
+ |
+
+ |
+416 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +-- | +|
| Mittelland (MP) | +-- | +|
| Alpennordflanke (NA) | +
+nicht
+ |
+|
|
+ |
+
+potenziell
+ |
+D2 | +
|
+ |
+
+nicht
+ |
+|
| Westliche Zentralalpen (WA) | +
+potenziell
+ |
+D2 | +
|
+Keine nationale
+ |
+
|
+ |
+Nein | +|
|
+ |
+
+ |
+(12.03.1973) | +
|
+ |
+-- | + + +
|
+1.
+ |
+
+ |
+
|
+1*.
+ |
+
+ |
+
| - | +Holotype | +paratypes | +|||
|---|---|---|---|---|---|
| - | +mean | +min | +max | +SD | +|
| Standard length (mm) | +58 | +- | +51 | +62 | +- | +
| In percent of standard length | +|||||
| Head length | +23.8 | +23.1 | +20.4 | +24.8 | +1.1 | +
| Body depth at dorsal-fin origin | +14.2 | +16.5 | +14.2 | +17.7 | +0.8 | +
| Predorsal length | +52.7 | +50.3 | +47.7 | +53.0 | +1.7 | +
| Postdorsal length | +35.9 | +35.6 | +32.0 | +39.7 | +2.0 | +
| Preanal length | +74.2 | +71.0 | +68.4 | +74.2 | +1.5 | +
| Prepelvic length | +52.4 | +49.5 | +46.6 | +52.4 | +1.4 | +
| Distance between pectoral and pelvic-fin origins | +32.0 | +29.6 | +27.0 | +32.0 | +1.1 | +
| Distance between pelvic and anal-fin origins | +22.0 | +21.1 | +18.0 | +27.5 | +1.9 | +
| Distance between vent and anal-fin origin | +3.8 | +4.0 | +3.2 | +5.1 | +0.5 | +
| Depth of caudal peduncle | +6.3 | +7.2 | +6.4 | +7.9 | +0.4 | +
| Length of caudal peduncle | +23.1 | +21.4 | +19.7 | +23.7 | +1.3 | +
| Dorsal-fin depth | +17.6 | +18.8 | +16.6 | +20.7 | +1.2 | +
| Anal-fin depth | +15.0 | +16.3 | +14.5 | +18.2 | +0.9 | +
| Pectoral-fin length | +20.0 | +22.2 | +18.3 | +26.3 | +1.7 | +
| Pelvic-fin length | +17.8 | +16.8 | +14.6 | +17.9 | +0.9 | +
| In percent of head length | +|||||
| Head depth at eye | +46.5 | +49.3 | +45.4 | +58.1 | +3.2 | +
| Snout length | +33.6 | +42.2 | +35.6 | +37.9 | +3.1 | +
| Eye diameter | +19.4 | +20.6 | +17.3 | +24.5 | +1.9 | +
| Postorbital distance | +45.7 | +46.9 | +42.5 | +55.3 | +3.2 | +
| Maximum head width | +61.6 | +62.0 | +54.6 | +69.5 | +3.3 | +
| Interorbital width | +25.3 | +28.0 | +23.1 | +35.8 | +3.2 | +
| Length of inner rostral barbel | +23.9 | +25.9 | +20.5 | +32.7 | +2.8 | +
| Length of outer rostral barbel | +28.3 | +30.6 | +24.8 | +37.6 | +3.4 | +
| Length of maxillary barbel | +26.7 | +26.6 | +22.4 | +34.1 | +2.7 | +
| Caudal peduncle length/depth | +3.0 | +2.9 | +2.8 | +3.2 | +0.1 | +
| - | +
+"
+ |
+
+"
+ |
+||
|---|---|---|---|---|
|
+males (
+ |
+
+females (
+ |
+
+males (
+ |
+
+females (
+ |
+|
| Standard length (mm) | +29.0-34.6 | +27.7-35.6 | +31.3-38.1 | +30.2-39.0 | +
|
+ |
+||||
| Body depth | +20.6-24.4 | +19.7-22.5 | +26.0-30.9 | +22.8-25.1 | +
| Caudal peduncle depth | +12.6-14.5 | +11.2-12.7 | +14.0-17.1 | +12.9-13.7 | +
| Pre-dorsal length | +67.6-73.9 | +71.6-73.9 | +67.5-72.3 | +68.5-70.7 | +
| Pre-pelvic length | +43.0-48.6 | +44.9-49.7 | +45.1-49.5 | +46.4-47.8 | +
| Length of dorsal-fin base | +8.7-11.6 | +7.3-10.1 | +11.9-13.1 | +10.8-11.6 | +
| Length of anal-fin base | +16.8-21.1 | +13.2-16.6 | +16.0-20.0 | +15.0-16.2 | +
| Caudal-fin length | +30.1-33.8 | +28.9-31.2 | +27.5-30.2 | +25.3-28.5 | +
| Pectoral-fin length | +18.6-22.1 | +18.8-20.8 | +18.7-22.4 | +17.5-19.5 | +
| Pelvic-fin length | +15.5-22.8 | +13.2-14.6 | +14.0-18.0 | +11.9-12.9 | +
| Head length (mm) | +6.9-8.4 | +6.2-7.9 | +8.2-9.5 | +7.2-9.8 | +
|
+ |
+||||
| Head depth | +61.9-67.9 | +59.4-63.5 | +70.7-79.8 | +66.6-69.6 | +
| Head width | +57.1-63.2 | +59.0-64.7 | +63.1-67.9 | +65.3-68.1 | +
| Snout length | +21.0-23.9 | +19.2-23.4 | +20.7-22.8 | +22.2-24.4 | +
| Lower jaw length | +9.5-12.3 | +8.7-11.0 | +9.8-14.3 | +8.9-11.2 | +
| Eye diameter | +35.7-39.5 | +36.2-41.1 | +35.8-38.5 | +37.2-39.2 | +
|
+7.1.4 -
+ |
+
|
+8.2.1.2 - Kalkreiche
+ |
+
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +sehr trocken | +Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +warm-kollin | +||
|
+ |
+
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||
|
+ |
+0 - unbedeutend, keine Bindung. | +
| Ruhiges Wasser | +0 - unbedeutend, keine Bindung. | +
| Grundwasser | +0 - unbedeutend, keine Bindung. | +
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+312200 | +
| = | +
+ |
+
+ |
+287 | +
| = | +
+ |
+
+ |
+1101 | +
| = | +
+ |
+
+ |
+1101 | +
| = | +
+ |
+
+ |
+312200 | +
| = | +
+ |
+
+ |
+928 | +
| = | +
+ |
+
+ |
+809 | +
| = | +
+ |
+
+ |
+312200 | +
| = | +
+ |
+
+ |
+199 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +vom Aussterben bedroht (Critically Endangered) | +B2b(iii)c(iii) | +
| Mittelland (MP) | +vom Aussterben bedroht (Critically Endangered) | +B2b(iii)c(iii) | +
| Alpennordflanke (NA) | +verschollen, vermutlich in der Schweiz ausgestorben (Critically Endangered, Probably Extinct) | +|
|
+ |
+verschollen, vermutlich in der Schweiz ausgestorben (Critically Endangered, Probably Extinct) | +|
|
+ |
+-- | +|
| Westliche Zentralalpen (WA) | +
+stark
+ |
+B2b(iii)c(iii) | +
|
+ |
+
+3 - Mittlere nationale
+ |
+
|
+ |
+2 - Klarer Massnahmebedarf | +
|
+ |
+1 - Gering | +
|
+ |
+
+1 -
+ |
+
|
+ |
+Nein | +|
|
+ |
+
+ |
+(25.07.2007) | +
|
+ |
+-- | +|
|
+ |
+
+ |
+(23.01.2013) | +
|
+ |
+Z - Zielart | +weitere Informationen | +
| Genera | +Diagnostic nucleotide positions | +|||||||||||||||||||||||||||||||||
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| - | +- | +- | +- | +- | +- | +- | +- | +- | +- | +- | +- | +- | +- | +- | +- | +- | +- | +- | +- | +- | +- | +- | +- | +- | +- | +1 | +1 | +1 | +1 | +1 | +1 | +1 | +1 | +1 | +
| - | +- | +- | +1 | +1 | +2 | +2 | +3 | +3 | +3 | +4 | +4 | +4 | +6 | +6 | +7 | +7 | +8 | +8 | +8 | +8 | +8 | +9 | +9 | +9 | +9 | +0 | +0 | +0 | +1 | +2 | +2 | +2 | +2 | +2 | +
| - | +6 | +8 | +0 | +8 | +0 | +9 | +4 | +7 | +8 | +1 | +7 | +9 | +1 | +4 | +1 | +8 | +4 | +5 | +5 | +8 | +9 | +4 | +9 | +7 | +9 | +3 | +4 | +4 | +5 | +0 | +1 | +4 | +8 | +9 | +
| - | +0 | +0 | +6 | +4 | +5 | +5 | +6 | +0 | +5 | +8 | +8 | +3 | +4 | +9 | +5 | +4 | +1 | +3 | +9 | +3 | +2 | +1 | +4 | +0 | +7 | +6 | +3 | +6 | +3 | +4 | +4 | +0 | +5 | +7 | +
|
+ |
+
+ |
+
+ |
+T | +
+ |
+
+ |
+
+ |
+G | +
+ |
+T | +T | +T | +T | +T | +
+ |
+
+ |
+C | +C | +G | +
+ |
+C | +C | +
+ |
+
+ |
+G | +
+ |
+T | +T | +T | +C | +G | +
+ |
+
+ |
+
+ |
+T | +
|
+ |
+T | +G | +C | +T | +A/G | +C | +A | +G | +C | +C | +C | +C | +
+ |
+C | +C | +
+ |
+
+ |
+
+ |
+T | +
+ |
+
+ |
+C | +G/A | +A | +T | +C | +
+ |
+C | +
+ |
+A | +C | +A/G | +C | +C | +
|
+ |
+T | +G | +A/C/T | +T | +A/G | +C | +A | +G | +T/C | +T/C | +C | +C | +T | +C | +C | +C | +C | +A | +T | +C | +C | +C | +G/A | +G/A | +T | +C/T | +C | +C/T | +C | +A | +C/A | +A/G | +C/T | +C/T | +
|
+ |
+T | +G | +C/T | +T | +A/G | +C | +G | +G | +T/C | +T/C | +T | +T | +T | +C | +C | +C | +C | +A/G | +T | +C | +C | +C | +G/A | +A | +T | +T | +C | +C | +C | +A | +C | +A/G | +C | +C/T | +
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+326300 | +
| = | +
+ |
+
+ |
+326300 | +
| = | +
+ |
+
+ |
+2305 | +
| = | +
+ |
+
+ |
+326300 | +
| +P +1 + | ++P +2 + | ++P +3 + | +
|---|---|---|
| 1 | +||
| 2 | +||
| 3 | +||
| 4 | +||
| 5 | +