plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity

Add updates up until 2025-01-27 13:13:26

Browse source
This commit is contained in:
ggserver 2025-01-27 13:18:33 +00:00
parent 34a82f1014
commit 172587dc3a
1 changed files with 846 additions and 0 deletions
Show stats Download patch file Download diff file Expand all files Collapse all files

846
data/2A/5A/87/2A5A87D7FF9F23396DFA3F61FBB3FA45.xml Normal file
View file

@ -0,0 +1,846 @@
<document id="81835D388E5C8C3DD2FECDD0A1FF35D4" ID-DOI="10.11646/zootaxa.5569.2.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="14734064" ID-ZooBank="BD58BB2E-A227-497D-AEDF-4EFC37CA89F6" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1737734540660" checkinUser="plazi" docAuthor="Pauwels, Olivier S. G., Donbundit, Nattasuda, Sumontha, Montri &amp; Meesook, Worawitoo" docDate="2025" docId="2A5A87D7FF9F23396DFA3F61FBB3FA45" docLanguage="en" docName="zootaxa.5569.2.7.pdf" docOrigin="Zootaxa 5569 (2)" docSource="https://doi.org/10.11646/zootaxa.5569.2.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oligodon cicadophagus Pauwels, Donbundit, Sumontha &amp; Meesook, 2025, sp. nov." docType="treatment" docVersion="1" lastPageNumber="357" masterDocId="D663FFAFFF9D23356D6D3A3FFFA3FFAC" masterDocTitle="Oligodon cicadophagus, a new kukri snake from southern peninsular Thailand (Squamata: Colubridae)" masterLastPageNumber="364" masterPageNumber="345" pageNumber="347" updateTime="1737983605254" updateUser="carolina" zenodo-license-document="CLOSED">
<mods:mods id="3B0D95D324065C0403C46B5E1CD0B67F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F26F3F9BB3D7F05C51FAF10553A4F917">
<mods:title id="150044ECE4C89AD6B74628D37CC52C89">Oligodon cicadophagus, a new kukri snake from southern peninsular Thailand (Squamata: Colubridae)</mods:title>
</mods:titleInfo>
<mods:name id="0F0C2EBCD6FEDB0DD07833B05B814641" type="personal">
<mods:role id="F0B85148446C4B00EFA440FAED339AA2">
<mods:roleTerm id="EDD0857C70E2D903366E4B394C7C1CB5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E83F7792BA9EE6CAD3A2BEEA4BE59A1F">Pauwels, Olivier S. G.</mods:namePart>
<mods:affiliation id="9D667678DECB9A4B2B2EC7A86F8111C5">Royal Belgian Institute of Natural Sciences, Rue Vautier 29, B- 1000 Brussels, Belgium.</mods:affiliation>
</mods:name>
<mods:name id="3E7EBB074F89CA64A3FF7F489B36F4B1" type="personal">
<mods:role id="34B451124B637693136B61AA01C5BA0B">
<mods:roleTerm id="91C627D7C050A622421D37499EC8B052">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3629526766B4E094D2BA1FD7CC2A5F1B">Donbundit, Nattasuda</mods:namePart>
<mods:affiliation id="06232BF5EAE12E46E6A4856E51DF1DFC">Department of Biology, Faculty of Science, Khon Kaen University, Muang, Khon Kaen 40002, Thailand.</mods:affiliation>
</mods:name>
<mods:name id="9F4CE73F6E1DB57E399E881FA4D2BE80" type="personal">
<mods:role id="E0690B63223695621132E0ECE8604831">
<mods:roleTerm id="3D9183EE4518958C49175272ABC9CF1B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4B0E34E84BF59E49FA067CD04B715E63">Sumontha, Montri</mods:namePart>
<mods:affiliation id="217377289FF3B29AB4AE519A53DCC9C7">Ranong Marine Fisheries Research and Development Center, 157 Chaloem Phrakiat Rd., Paknam, Muang, Ranong 85000, Thailand.</mods:affiliation>
</mods:name>
<mods:name id="3DC57C43E87552990EB873D342C0FA23" type="personal">
<mods:role id="48CE5CCC8165C1DA4C12874B8892F25A">
<mods:roleTerm id="C0DA7ECD6626AAD742EFD1DEBB6D5B2F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D98415A08C08FD9A352C98618243DDFA">Meesook, Worawitoo</mods:namePart>
<mods:affiliation id="0556B7926162B51C3E19504CDD99F748">Faculty of Science and Technology, Rajamangala University of Technology Srivijaya, 109 M. 2, Tham Yai, Thung Song, Nakhon Si Thammarat 80110, Thailand.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="855A6902E8BB641D1FBE165D195E33BF">text</mods:typeOfResource>
<mods:relatedItem id="0AE407D9B199075DAF30D9D7E195FAEA" type="host">
<mods:titleInfo id="F02AB590C2E1A089DB865309A9CD24B5">
<mods:title id="CC80B92664E88A3E4F7C2B3ED5BD97CC">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="314855BC9B8D4AE9F9E3B359D7751821">
<mods:date id="2CE13DAD0B14E9815D58259CC5ED3E3C">2025</mods:date>
<mods:detail id="C9359D61F4886C4463FBB67DC4ED492E" type="pubDate">
<mods:number id="1BAC3438F0E437089801A774F5A21E48">2025-01-14</mods:number>
</mods:detail>
<mods:detail id="A496953C144573C40E6F26288D4652DD" type="volume">
<mods:number id="AC6D74832CE58D61056101721DBC8DC2">5569</mods:number>
</mods:detail>
<mods:detail id="0569D968E3AE09F53389250A1204E752" type="issue">
<mods:number id="1850290C9E8DBEF70AE95BE2DEE33016">2</mods:number>
</mods:detail>
<mods:extent id="6B833FC326E36F19BCEF107E6E312F73" unit="page">
<mods:start id="E3A03B5640C35EC8A263A1D3D992FCED">345</mods:start>
<mods:end id="D5512B4D91CD518BC5E26A6C9FA08FDB">364</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="A00997A070D1AFB6B3D0C6F61CC482AF">
<mods:url id="F1DBB17C79A249385EF8405F3745D020">https://doi.org/10.11646/zootaxa.5569.2.7</mods:url>
</mods:location>
<mods:classification id="049F0D0F0AAE476CDD487D30DC4601D4">journal article</mods:classification>
<mods:identifier id="1D0FF67A24476533FC5FC4D121C0E16D" type="DOI">10.11646/zootaxa.5569.2.7</mods:identifier>
<mods:identifier id="54A7CF517BBA71D331DCAEDC9FB03E04" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3F4D6566F698784D37A00FA9D901C175" type="Zenodo-Dep">14734064</mods:identifier>
<mods:identifier id="4E2BDEAD2CD2A224B87D52C35AF92216" type="ZooBank">BD58BB2E-A227-497D-AEDF-4EFC37CA89F6</mods:identifier>
</mods:mods>
<treatment id="2A5A87D7FF9F23396DFA3F61FBB3FA45" LSID="urn:lsid:plazi:treatment:2A5A87D7FF9F23396DFA3F61FBB3FA45" httpUri="http://treatment.plazi.org/id/2A5A87D7FF9F23396DFA3F61FBB3FA45" lastPageId="12" lastPageNumber="357" pageId="2" pageNumber="347">
<subSubSection id="EAE9654AFF9F23376DFA3F61FDB9FAD5" box="[151,538,1374,1401]" pageId="2" pageNumber="347" type="nomenclature">
<paragraph id="A24C36C1FF9F23376DFA3F61FDB9FAD5" blockId="2.[151,538,1374,1437]" box="[151,538,1374,1401]" pageId="2" pageNumber="347">
<heading id="F90481ADFF9F23376DFA3F61FDB9FAD5" bold="true" box="[151,538,1374,1401]" fontSize="11" level="1" pageId="2" pageNumber="347" reason="1">
<emphasis id="9087EAD3FF9F23376DFA3F61FDB9FAD5" bold="true" box="[151,538,1374,1401]" pageId="2" pageNumber="347">
<taxonomicName id="65F34D42FF9F23376DFA3F61FE16FAD5" authorityName="Pauwels &amp; Donbundit &amp; Sumontha &amp; Meesook" authorityYear="2025" box="[151,437,1374,1401]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="2" pageNumber="347" phylum="Chordata" rank="species" species="cicadophagus" status="sp. nov.">
<emphasis id="9087EAD3FF9F23376DFA3F61FE16FAD5" bold="true" box="[151,437,1374,1401]" italics="true" pageId="2" pageNumber="347">Oligodon cicadophagus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8BB457A8FF9F23376CD03F60FDB9FAD5" box="[445,538,1375,1401]" pageId="2" pageNumber="347" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="EAE9654AFF9F23376DFA3FBCFE15FA31" box="[151,438,1410,1437]" pageId="2" pageNumber="347" type="description">
<paragraph id="A24C36C1FF9F23376DFA3FBCFE15FA31" blockId="2.[151,538,1374,1437]" box="[151,438,1410,1437]" pageId="2" pageNumber="347">
(
<figureCitation id="3AC82A44FF9F23376DF23FBCFEA8FA31" box="[159,267,1410,1437]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="3.[151,250,1940,1964]" captionStartId-1="4.[151,250,1045,1069]" captionStartId-2="5.[151,250,1964,1988]" captionStartId-3="6.[151,250,1927,1951]" captionStartId-4="7.[151,250,1943,1967]" captionTargetBox-0="[204,1383,201,1914]" captionTargetBox-1="[164,1423,181,1021]" captionTargetBox-2="[211,1322,265,1906]" captionTargetBox-3="[151,1436,181,1903]" captionTargetBox-4="[174,1331,214,1916]" captionTargetId-0="figure-23@3.[204,1383,179,1914]" captionTargetId-1="figure-35@4.[164,1423,181,1021]" captionTargetId-2="figure-49@5.[175,1412,237,1941]" captionTargetId-3="figure-35@6.[151,1436,181,1903]" captionTargetId-4="figure-23@7.[174,1414,181,1919]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURE 1. Live holotype of Oligodon cicadophagus sp. nov. (CUMZ-R-2625) in dorsal (A) and ventral (B) views. Photographs by M. Sumontha." captionText-1="FIGURE 2. Live holotype of Oligodon cicadophagus sp. nov. (CUMZ-R-2625) in left profile (A), right profile (B), frontodorsal view of the head (C) and right flank (D). Photographs by M. Sumontha. Note the purple brown tongue in A." captionText-2="FIGURE 3. Preserved holotype of Oligodon cicadophagus sp. nov. (CUMZ-R-2625) with everted hemipenes in dorsal (A) and ventral (B) views. Photographs by M. Sumontha." captionText-3="FIGURE 4. Live paratype of Oligodon cicadophagus sp. nov. (CUMZ-R-2626) in dorsal (A) and ventral (B) views. Photographs by M. Sumontha." captionText-4="FIGURE 5. Live paratype of Oligodon cicadophagus sp. nov. (CUMZ-R-2626) in dorsal view (A) and right profile (B) of the head. Photographs by M. Sumontha." figureDoi-0="http://doi.org/10.5281/zenodo.14734066" figureDoi-1="http://doi.org/10.5281/zenodo.14734070" figureDoi-2="http://doi.org/10.5281/zenodo.14734072" figureDoi-3="http://doi.org/10.5281/zenodo.14734076" figureDoi-4="http://doi.org/10.5281/zenodo.14734078" httpUri-0="https://zenodo.org/record/14734066/files/figure.png" httpUri-1="https://zenodo.org/record/14734070/files/figure.png" httpUri-2="https://zenodo.org/record/14734072/files/figure.png" httpUri-3="https://zenodo.org/record/14734076/files/figure.png" httpUri-4="https://zenodo.org/record/14734078/files/figure.png" pageId="2" pageNumber="347">Figs. 1–5</figureCitation>
,
<figureCitation id="3AC82A44FF9F23376C7A3FBCFE84FA31" box="[279,295,1411,1437]" captionStart="FIGURE 7" captionStartId="9.[151,250,1020,1044]" captionTargetBox="[432,1156,181,996]" captionTargetId="figure-23@9.[432,1156,181,996]" captionTargetPageId="9" captionText="FIGURE 7. Live adult Oligodon cicadophagus sp. nov. (unvouchered) from Khao Phung Chang, Phang-Nga Province, southern Thailand. When caught, it regurgitated a cicada nymph. Photograph by T. Butmee." figureDoi="http://doi.org/10.5281/zenodo.14734084" httpUri="https://zenodo.org/record/14734084/files/figure.png" pageId="2" pageNumber="347">7</figureCitation>
;
<tableCitation id="EF71037AFF9F23376C5E3FBCFE0EFA31" box="[307,429,1411,1437]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStartId-0="13.[151,239,151,175]" captionStartId-1="14.[150,238,485,511]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="TABLE 1. Meristic and morphometric (in mm) data for the type-series of Oligodon cicadophagus sp. nov. Paired meristic characters are given left/right. Paired measurements are given for the right side. Supralabial numbers are followed in brackets by the ones contacting the orbit. Infralabial numbers are followed in brackets by how many contact the anterior chin shields. A = anal plate; NA = not assessed; S = single. For the other abbreviations, see Material and methods." captionText-1="TABLE 2. Comparison of Oligodon species of the southern Indochinese and Thai-Malay region showing no distinct nuchal chevron, no distinct transversal pattern and no dorsal or supracaudal blotches. Bolded values are diagnostic differences from Oligodon cicadophagus sp. nov. NA = not available." pageId="2" pageNumber="347">Tables 1–2</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="EAE9654AFF9F23376DFA3FF4FAFBF9FD" pageId="2" pageNumber="347" type="materials_examined">
<paragraph id="A24C36C1FF9F23376DFA3FF4FA9BF9A5" blockId="2.[151,1437,1482,2013]" pageId="2" pageNumber="347">
<materialsCitation id="129B3C9CFF9F23376DFA3FF4FA9BF9A5" collectedFrom="limestone cave" collectingDate="2021-10-28" collectionCode="CUMZ-R" collectorName="Punya Tulyasuk" country="Thailand" latitude="8.086611" location="Sai Thai" longLatPrecision="1" longitude="98.87769" municipality="Muang Krabi District" pageId="2" pageNumber="347" specimenCode="2625" specimenCount="1" stateProvince="Krabi Province" typeStatus="holotype">
<emphasis id="9087EAD3FF9F23376DFA3FF4FEA8FA49" bold="true" box="[151,267,1483,1509]" pageId="2" pageNumber="347">
<typeStatus id="7D488863FF9F23376DFA3FF4FEA5FA49" box="[151,262,1483,1509]" pageId="2" pageNumber="347" type="holotype">Holotype</typeStatus>
.
</emphasis>
CUMZ-R-2625(field no.MS 922),
<specimenCount id="B4F5FD48FF9F23376FFC3FF4FCA1FA49" box="[657,770,1483,1509]" pageId="2" pageNumber="347" type="adult">adult male</specimenCount>
caught on
<date id="D64D1001FF9F23376E193FF4FB8AFA49" box="[884,1065,1483,1509]" pageId="2" pageNumber="347" value="2021-10-28">
<collectingDate id="C609E9E9FF9F23376E193FF4FB8AFA49" box="[884,1065,1483,1509]" pageId="2" pageNumber="347" value="2021-10-28">
28October
<quantity id="650B9B24FF9F23376E9C3FF4FB8AFA49" box="[1009,1065,1483,1509]" metricMagnitude="1" metricUnit="m" metricValue="5.13334" pageId="2" pageNumber="347" unit="in" value="2021.0">2021</quantity>
</collectingDate>
</date>
in a
<collectedFrom id="11EF753CFF9F2337693B3FF4FB5AFA49" box="[1110,1273,1483,1509]" pageId="2" pageNumber="347">limestone cave</collectedFrom>
(
<geoCoordinate id="C7C75006FF9F233768683FF5FA34FA49" box="[1285,1431,1482,1509]" degrees="8" direction="north" minutes="05" orientation="latitude" pageId="2" pageNumber="347" precision="1" seconds="11.8" value="8.086611">8°05’11.8”N</geoCoordinate>
,
<geoCoordinate id="C7C75006FF9F23376DFA3FD1FE97F9A5" box="[151,308,1518,1545]" degrees="98" direction="east" minutes="52" orientation="longitude" pageId="2" pageNumber="347" precision="1" seconds="39.7" value="98.87769">98°52’39.7”E</geoCoordinate>
) at
<location id="A72C601AFF9F23376C333FD0FE1DF9A5" LSID="urn:lsid:plazi:treatment:2A5A87D7FF9F23396DFA3F61FBB3FA45:A72C601AFF9F23376C333FD0FE1DF9A5" box="[350,446,1519,1545]" country="Thailand" latitude="8.086611" longLatPrecision="1" longitude="98.87769" municipality="Muang Krabi District" name="Sai Thai" pageId="2" pageNumber="347" stateProvince="Krabi Province">Sai Thai</location>
,
<collectingMunicipality id="4228ACBBFF9F23376CA43FD0FD1EF9A5" box="[457,701,1519,1545]" pageId="2" pageNumber="347">Muang Krabi District</collectingMunicipality>
,
<collectingRegion id="6037F823FF9F23376FA53FD0FCD6F9A5" box="[712,885,1519,1545]" country="Thailand" name="Krabi" pageId="2" pageNumber="347">Krabi Province</collectingRegion>
, southern
<collectingCountry id="DAE47651FF9F23376E853FD0FBEDF9A5" box="[1000,1102,1519,1545]" name="Thailand" pageId="2" pageNumber="347">Thailand</collectingCountry>
, by
<collectorName id="0F065317FF9F233769113FD0FA97F9A5" box="[1148,1332,1519,1545]" pageId="2" pageNumber="347">Punya Tulyasuk</collectorName>
.
</materialsCitation>
</paragraph>
<paragraph id="A24C36C1FF9F23376DAA3C2CFAFBF9FD" blockId="2.[151,1437,1482,2013]" pageId="2" pageNumber="347">
<materialsCitation id="129B3C9CFF9F23376DAA3C2CFAF7F9FD" collectingDate="2021-12-02" collectionCode="CUMZ-R" collectorName="Punya Tulyasuk" country="Thailand" county="Punya Tulyasuk" latitude="8.084027" location="Nong Thalay" longLatPrecision="1" longitude="98.85983" municipality="Muang Krabi District" pageId="2" pageNumber="347" specimenCode="2626" specimenCount="1" stateProvince="Krabi Province" typeStatus="paratype">
<emphasis id="9087EAD3FF9F23376DAA3C2CFE98F981" bold="true" box="[199,315,1555,1581]" pageId="2" pageNumber="347">
<typeStatus id="7D488863FF9F23376DAA3C2CFE95F981" box="[199,310,1555,1581]" pageId="2" pageNumber="347" type="paratype">Paratype</typeStatus>
.
</emphasis>
CUMZ-R-2626 (field no. MS 921),
<specimenCount id="B4F5FD48FF9F23376F973C2CFCD6F981" box="[762,885,1555,1581]" pageId="2" pageNumber="347" type="adult">adult male</specimenCount>
caught on
<date id="D64D1001FF9F2337696E3C2CFB75F981" box="[1027,1238,1555,1581]" pageId="2" pageNumber="347" value="2021-12-02">
<collectingDate id="C609E9E9FF9F2337696E3C2CFB75F981" box="[1027,1238,1555,1581]" pageId="2" pageNumber="347" value="2021-12-02">2 December 2021</collectingDate>
</date>
at
<location id="A72C601AFF9F233768693C2CFA3FF981" LSID="urn:lsid:plazi:treatment:2A5A87D7FF9F23396DFA3F61FBB3FA45:A72C601AFF9F233768693C2CFA3FF981" box="[1284,1436,1555,1581]" country="Thailand" county="Punya Tulyasuk" latitude="8.084027" longLatPrecision="1" longitude="98.85983" municipality="Muang Krabi District" name="Nong Thalay" pageId="2" pageNumber="347" stateProvince="Krabi Province">Nong Thalay</location>
(
<geoCoordinate id="C7C75006FF9F23376DCD3C09FE92F9FD" box="[160,305,1590,1617]" degrees="8" direction="north" minutes="05" orientation="latitude" pageId="2" pageNumber="347" precision="1" seconds="02.5" value="8.084027">8°05’02.5”N</geoCoordinate>
<geoCoordinate id="C7C75006FF9F23376C553C09FE75F9FD" box="[312,470,1590,1617]" degrees="98" direction="east" minutes="51" orientation="longitude" pageId="2" pageNumber="347" precision="1" seconds="35.4" value="98.85983">98°51’35.4”E</geoCoordinate>
),
<collectingMunicipality id="4228ACBBFF9F23376C873C08FD7DF9FD" box="[490,734,1591,1617]" pageId="2" pageNumber="347">Muang Krabi District</collectingMunicipality>
,
<collectingRegion id="6037F823FF9F23376F843C08FC35F9FD" box="[745,918,1591,1617]" country="Thailand" name="Krabi" pageId="2" pageNumber="347">Krabi Province</collectingRegion>
, southern
<collectingCountry id="DAE47651FF9F233769653C08FBCDF9FD" box="[1032,1134,1591,1617]" name="Thailand" pageId="2" pageNumber="347">Thailand</collectingCountry>
, by
<collectorName id="0F065317FF9F233769F03C08FAF7F9FD" box="[1181,1364,1591,1617]" pageId="2" pageNumber="347">Punya Tulyasuk</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EAE9654AFF9F23376DAA3C65FCBFF885" pageId="2" pageNumber="347" type="diagnosis">
<paragraph id="A24C36C1FF9F23376DAA3C65FCBFF885" blockId="2.[151,1437,1482,2013]" pageId="2" pageNumber="347">
<emphasis id="9087EAD3FF9F23376DAA3C65FEE1F9D8" bold="true" box="[199,322,1626,1652]" pageId="2" pageNumber="347">Diagnosis.</emphasis>
<taxonomicName id="65F34D42FF9F23376C213C64FDFEF9D9" authorityName="Pauwels &amp; Donbundit &amp; Sumontha &amp; Meesook" authorityYear="2025" box="[332,605,1627,1653]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="2" pageNumber="347" phylum="Chordata" rank="species" species="cicadophagus" status="sp. nov.">
<emphasis id="9087EAD3FF9F23376C213C64FDFEF9D9" box="[332,605,1627,1653]" italics="true" pageId="2" pageNumber="347">Oligodon cicadophagus</emphasis>
</taxonomicName>
<emphasis id="9087EAD3FF9F23376F0A3C64FD67F9D9" bold="true" box="[615,708,1627,1653]" pageId="2" pageNumber="347">
<taxonomicNameLabel id="8BB457A8FF9F23376F0A3C64FD67F9D9" box="[615,708,1627,1653]" pageId="2" pageNumber="347" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from all other congeneric species by the combination of its maximal known SVL of
<quantity id="650B9B24FF9F23376FE83C41FD46F935" box="[645,741,1662,1689]" metricMagnitude="-1" metricUnit="m" metricValue="5.37" pageId="2" pageNumber="347" unit="mm" value="537.0">537 mm</quantity>
; 11 maxillary teeth; eight supralabials; one preocular and one presubocular; 17-17-15 dorsal scale rows; 159–168 ventrals and 50 subcaudals in males; a single anal; hemipenes bilobed with large broad lobes, no distinct flounced calyces and unforked sulcus spermaticus; dorsal surface of head, body and tail uniformly olive-gray, without nuchal chevron, dorsal stripes, crossbands, blotches or reticulations; and a venter cream anteriorly turning to dark gray posteriorly.
</paragraph>
</subSubSection>
<subSubSection id="EAE9654AFF9F233D6DAA3D0DFE3FFEED" lastPageId="8" lastPageNumber="353" pageId="2" pageNumber="347" type="description">
<paragraph id="A24C36C1FF9F23376DAA3D0DFCDDF871" blockId="2.[151,1437,1482,2013]" pageId="2" pageNumber="347">
<emphasis id="9087EAD3FF9F23376DAA3D0DFDB9F8E1" bold="true" box="[199,538,1842,1869]" pageId="2" pageNumber="347">
Description of the
<typeStatus id="7D488863FF9F23376CC03D0CFDB5F8E1" box="[429,534,1843,1869]" pageId="2" pageNumber="347" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male (
<figureCitation id="3AC82A44FF9F23376FDB3D0CFC86F8E1" box="[694,805,1843,1869]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1940,1964]" captionStartId-1="4.[151,250,1045,1069]" captionStartId-2="5.[151,250,1964,1988]" captionTargetBox-0="[204,1383,201,1914]" captionTargetBox-1="[164,1423,181,1021]" captionTargetBox-2="[211,1322,265,1906]" captionTargetId-0="figure-23@3.[204,1383,179,1914]" captionTargetId-1="figure-35@4.[164,1423,181,1021]" captionTargetId-2="figure-49@5.[175,1412,237,1941]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Live holotype of Oligodon cicadophagus sp. nov. (CUMZ-R-2625) in dorsal (A) and ventral (B) views. Photographs by M. Sumontha." captionText-1="FIGURE 2. Live holotype of Oligodon cicadophagus sp. nov. (CUMZ-R-2625) in left profile (A), right profile (B), frontodorsal view of the head (C) and right flank (D). Photographs by M. Sumontha. Note the purple brown tongue in A." captionText-2="FIGURE 3. Preserved holotype of Oligodon cicadophagus sp. nov. (CUMZ-R-2625) with everted hemipenes in dorsal (A) and ventral (B) views. Photographs by M. Sumontha." figureDoi-0="http://doi.org/10.5281/zenodo.14734066" figureDoi-1="http://doi.org/10.5281/zenodo.14734070" figureDoi-2="http://doi.org/10.5281/zenodo.14734072" httpUri-0="https://zenodo.org/record/14734066/files/figure.png" httpUri-1="https://zenodo.org/record/14734070/files/figure.png" httpUri-2="https://zenodo.org/record/14734072/files/figure.png" pageId="2" pageNumber="347">Figs. 1–3</figureCitation>
). Body robust but elongate. SVL
<quantity id="650B9B24FF9F233769C23D0DFA84F8E1" box="[1199,1319,1842,1869]" metricMagnitude="-1" metricUnit="m" metricValue="5.237" pageId="2" pageNumber="347" unit="mm" value="523.7">523.7 mm</quantity>
; TaL
<quantity id="650B9B24FF9F233768063D0DFF65F8DD" metricMagnitude="-2" metricUnit="m" metricValue="9.54" pageId="2" pageNumber="347" unit="mm" value="95.4">95.4 mm</quantity>
; TL
<quantity id="650B9B24FF9F23376D933D68FED6F8DD" box="[254,373,1879,1905]" metricMagnitude="-1" metricUnit="m" metricValue="6.191" pageId="2" pageNumber="347" unit="mm" value="619.1">619.1 mm</quantity>
. Head short, barely distinct from the poorly marked neck. Pupil round. Tail robust, tapering, accounting for 15.4 % of the TL (
<quantity id="650B9B24FF9F23376F603D44FD26F839" box="[525,645,1915,1941]" metricMagnitude="-1" metricUnit="m" metricValue="6.191" pageId="2" pageNumber="347" unit="mm" value="619.1">619.1 mm</quantity>
). HL
<quantity id="650B9B24FF9F23376FAE3D44FC89F839" box="[707,810,1915,1941]" metricMagnitude="-2" metricUnit="m" metricValue="1.69" pageId="2" pageNumber="347" unit="mm" value="16.9">16.9 mm</quantity>
; HW
<quantity id="650B9B24FF9F23376E073D44FC72F839" box="[874,977,1915,1941]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="2" pageNumber="347" unit="mm" value="10.6">10.6 mm</quantity>
; HD
<quantity id="650B9B24FF9F233769663D44FBC7F839" box="[1035,1124,1915,1941]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="2" pageNumber="347" unit="mm" value="7.4">7.4 mm</quantity>
. SnOrb
<quantity id="650B9B24FF9F233769D23D45FABBF839" box="[1215,1304,1914,1941]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="2" pageNumber="347" unit="mm" value="5.9">5.9 mm</quantity>
. Snout long (35 % of HL, 2.3 times as long as OrbD). OrbD
<quantity id="650B9B24FF9F23376FA73DA0FC86F815" box="[714,805,1951,1977]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="347" unit="mm" value="2.6">2.6 mm</quantity>
; OrbLip
<quantity id="650B9B24FF9F23376EE33DA1FC4AF815" box="[910,1001,1950,1977]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="347" unit="mm" value="2.5">2.5 mm</quantity>
; distance eye-nostril
<quantity id="650B9B24FF9F233769B13DA1FA94F815" box="[1244,1335,1950,1977]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="347" unit="mm" value="3.5">3.5 mm</quantity>
. Straight distance between nostrils
<quantity id="650B9B24FF9F23376CD53DFCFDB2F871" box="[440,529,1987,2013]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="2" pageNumber="347" unit="mm" value="4.4">4.4 mm</quantity>
. Distance between eyes
<quantity id="650B9B24FF9F23376E4C3DFCFCD9F871" box="[801,890,1987,2013]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="2" pageNumber="347" unit="mm" value="6.7">6.7 mm</quantity>
.
</paragraph>
<caption id="F68C6649FF9E23366DFA3DABFEEBF87C" ID-DOI="http://doi.org/10.5281/zenodo.14734066" ID-Zenodo-Dep="14734066" httpUri="https://zenodo.org/record/14734066/files/figure.png" pageId="3" pageNumber="348" startId="3.[151,250,1940,1964]" targetBox="[204,1383,201,1914]" targetPageId="3" targetType="figure">
<paragraph id="A24C36C1FF9E23366DFA3DABFEEBF87C" blockId="3.[151,1437,1940,2000]" pageId="3" pageNumber="348">
<emphasis id="9087EAD3FF9E23366DFA3DABFEB2F800" bold="true" box="[151,273,1940,1964]" pageId="3" pageNumber="348">FIGURE 1.</emphasis>
Live holotype of
<taxonomicName id="65F34D42FF9E23366CD23DABFD13F800" authorityName="Pauwels &amp; Donbundit &amp; Sumontha &amp; Meesook" authorityYear="2025" box="[447,688,1940,1964]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="3" pageNumber="348" phylum="Chordata" rank="species" species="cicadophagus" status="sp. nov.">
<emphasis id="9087EAD3FF9E23366CD23DABFD13F800" box="[447,688,1940,1964]" italics="true" pageId="3" pageNumber="348">Oligodon cicadophagus</emphasis>
</taxonomicName>
<emphasis id="9087EAD3FF9E23366FD83DABFCA0F800" bold="true" box="[693,771,1940,1964]" pageId="3" pageNumber="348">
<taxonomicNameLabel id="8BB457A8FF9E23366FD83DABFCA0F800" box="[693,771,1940,1964]" pageId="3" pageNumber="348" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CUMZ-R-2625) in dorsal (A) and ventral (B) views. Photographs by M. Sumontha.
</paragraph>
</caption>
<caption id="F68C6649FF9923316DFA3E2AFB7FFBFD" ID-DOI="http://doi.org/10.5281/zenodo.14734070" ID-Zenodo-Dep="14734070" httpUri="https://zenodo.org/record/14734070/files/figure.png" pageId="4" pageNumber="349" startId="4.[151,250,1045,1069]" targetBox="[164,1423,181,1021]" targetPageId="4" targetType="figure">
<paragraph id="A24C36C1FF9923316DFA3E2AFB7FFBFD" blockId="4.[151,1436,1045,1105]" pageId="4" pageNumber="349">
<emphasis id="9087EAD3FF9923316DFA3E2AFEB1FB81" bold="true" box="[151,274,1045,1069]" pageId="4" pageNumber="349">FIGURE 2.</emphasis>
Live holotype of
<taxonomicName id="65F34D42FF9923316CAF3E2AFD10FB81" authorityName="Pauwels &amp; Donbundit &amp; Sumontha &amp; Meesook" authorityYear="2025" box="[450,691,1045,1069]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="4" pageNumber="349" phylum="Chordata" rank="species" species="cicadophagus" status="sp. nov.">
<emphasis id="9087EAD3FF9923316CAF3E2AFD10FB81" box="[450,691,1045,1069]" italics="true" pageId="4" pageNumber="349">Oligodon cicadophagus</emphasis>
</taxonomicName>
<emphasis id="9087EAD3FF9923316FD53E2AFCA4FB81" bold="true" box="[696,775,1045,1069]" pageId="4" pageNumber="349">
<taxonomicNameLabel id="8BB457A8FF9923316FD53E2AFCA4FB81" box="[696,775,1045,1069]" pageId="4" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CUMZ-R-2625) in left profile (A), right profile (B), frontodorsal view of the head (C) and right flank (D). Photographs by M. Sumontha. Note the purple brown tongue in A.
</paragraph>
</caption>
<paragraph id="A24C36C1FF9923316DAA3E9CFE19FAA9" blockId="4.[151,1437,1187,2042]" pageId="4" pageNumber="349">
<emphasis id="9087EAD3FF9923316DAA3E9CFECDFB11" box="[199,366,1187,1213]" italics="true" pageId="4" pageNumber="349">Body scalation</emphasis>
. DSR 17-17-15, all smooth. DSR reduction from 17 to 15 occurs above the 104
<superScript id="55869B89FF99233169883E9CFB51FB1D" attach="left" box="[1253,1266,1187,1201]" fontSize="6" pageId="4" pageNumber="349">th</superScript>
(left) and 106
<superScript id="55869B89FF99233168FD3E9CFA3EFB1D" attach="none" box="[1424,1437,1187,1201]" fontSize="6" pageId="4" pageNumber="349">th</superScript>
(right) VEN by fusion of DSR 4 and 5 (left) or 3 and 4 (right). 2 PV + 159 VEN, laterally angulated. Anal plate single. SC 50, all divided.
</paragraph>
<paragraph id="A24C36C1FF9923316DAA3F30FC4AF975" blockId="4.[151,1437,1187,2042]" pageId="4" pageNumber="349">
<emphasis id="9087EAD3FF9923316DAA3F30FED1FA85" box="[199,370,1295,1321]" italics="true" pageId="4" pageNumber="349">Head scalation</emphasis>
. Rostral thick, curved onto upper snout surface, well visible from above, separating internasals by about one half of their length (
<figureCitation id="3AC82A44FF9923316F763F0CFDC0FAE1" box="[539,611,1331,1357]" captionStart="FIGURE 2" captionStartId="4.[151,250,1045,1069]" captionTargetBox="[164,1423,181,1021]" captionTargetId="figure-35@4.[164,1423,181,1021]" captionTargetPageId="4" captionText="FIGURE 2. Live holotype of Oligodon cicadophagus sp. nov. (CUMZ-R-2625) in left profile (A), right profile (B), frontodorsal view of the head (C) and right flank (D). Photographs by M. Sumontha. Note the purple brown tongue in A." figureDoi="http://doi.org/10.5281/zenodo.14734070" httpUri="https://zenodo.org/record/14734070/files/figure.png" pageId="4" pageNumber="349">Fig. 2</figureCitation>
). RosW
<quantity id="650B9B24FF9923316FAA3F0CFC82FAE2" box="[711,801,1331,1358]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="4" pageNumber="349" unit="mm" value="4.4">4.4 mm</quantity>
, RosH
<quantity id="650B9B24FF9923316E1A3F0CFC72FAE2" box="[887,977,1331,1358]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="4" pageNumber="349" unit="mm" value="3.7">3.7 mm</quantity>
. Nasals vertically divided. Nostril large, piercing middle of nasal. Two internasals, in broad contact, shorter than prefrontals. Two prefrontals, subrectangular, distinctly wider than long. Length of suture between prefrontals 2.4 times the length of suture between internasals. Frontal pentagonal, 1.2 times as long as wide; 1/1 supraoculars, distinctly longer than wide; SL 8/8, 2
<superScript id="55869B89FF99233168773FA0FA89FA01" attach="left" box="[1306,1322,1439,1453]" fontSize="6" pageId="4" pageNumber="349">nd</superScript>
and 3
<superScript id="55869B89FF992331681C3FA0FADCFA01" attach="left" box="[1393,1407,1439,1453]" fontSize="6" pageId="4" pageNumber="349">rd</superScript>
in contact with LOR, 4
<superScript id="55869B89FF9923316C103FFCFE29FA7D" attach="left" box="[381,394,1475,1489]" fontSize="6" pageId="4" pageNumber="349">th</superScript>
and 5
<superScript id="55869B89FF9923316CA23FFCFE7FFA7D" attach="left" box="[463,476,1475,1489]" fontSize="6" pageId="4" pageNumber="349">th</superScript>
in contact with orbit, posterior four distinctly larger than the four anterior ones. LOR 1/1, distinctly longer than high. PreOc 1/1; PreSubOc 1/1; no SubOc; PosOc 2/2, the upper one slightly larger; no PosSubOc. Temporals 2+2 on each side; the anterior upper temporal small and not in contact with the posterior upper temporal. Parietals larger than the frontal, in contact on more than 2/3 of their length behind the frontal. Parietals bordered posteriorly by undifferentiated dorsal scales. Mental width
<quantity id="650B9B24FF992331697D3C6CFBC9F9C2" box="[1040,1130,1619,1646]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="349" unit="mm" value="2.7">2.7 mm</quantity>
, mental length
<quantity id="650B9B24FF99233168763C6CFAD5F9C2" box="[1307,1398,1619,1646]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="349" unit="mm" value="1.8">1.8 mm</quantity>
. IL 8/9, 1
<superScript id="55869B89FF9923316DBA3C48FF41F929" attach="left" box="[215,226,1655,1669]" fontSize="6" pageId="4" pageNumber="349">st</superScript>
pair in broad contact behind mental, IL
<quantity id="650B9B24FF9923316FCA3C48FCA7F93D" box="[679,772,1655,1682]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="7.62" metricValueMin="2.54" pageId="4" pageNumber="349" unit="in" value="2.0" valueMax="3.0" valueMin="1.0">1 to 3 in</quantity>
contact with anterior chin shields on the left side, IL 1 to 4 on the right side. First pair of chin shields distinctly longer than 2
<superScript id="55869B89FF9923316E1F3CA4FC21F905" attach="left" box="[882,898,1691,1705]" fontSize="6" pageId="4" pageNumber="349">nd</superScript>
pair. Second pair of chin shields in contact with the 3
<superScript id="55869B89FF9923316DA33C80FF7FF961" attach="left" box="[206,220,1727,1741]" fontSize="6" pageId="4" pageNumber="349">rd</superScript>
and 4
<superScript id="55869B89FF9923316C4D3C80FE8EF961" attach="left" box="[288,301,1727,1741]" fontSize="6" pageId="4" pageNumber="349">th</superScript>
IL on the left side, and with the 4
<superScript id="55869B89FF9923316FC53C80FD16F961" attach="left" box="[680,693,1727,1741]" fontSize="6" pageId="4" pageNumber="349">th</superScript>
and 5
<superScript id="55869B89FF9923316F943C80FCA5F961" attach="left" box="[761,774,1727,1741]" fontSize="6" pageId="4" pageNumber="349">th</superScript>
IL on the right side.
</paragraph>
<paragraph id="A24C36C1FF9923316DAA3CDCFCD6F88D" blockId="4.[151,1437,1187,2042]" pageId="4" pageNumber="349">
<emphasis id="9087EAD3FF9923316DAA3CDCFE28F951" box="[199,395,1763,1789]" italics="true" pageId="4" pageNumber="349">Maxilla dentition</emphasis>
. Eleven MT (six teeth present + five empty sockets), increasing in size posteriorly, without diastema. Three posteriormost MT distinctly enlarged, kukri-like.
</paragraph>
<paragraph id="A24C36C1FF9923316DAA3D13FED1F8C5" blockId="4.[151,1437,1187,2042]" pageId="4" pageNumber="349">
<emphasis id="9087EAD3FF9923316DAA3D13FEE0F8E9" box="[199,323,1836,1861]" italics="true" pageId="4" pageNumber="349">Hemipenes</emphasis>
. Long (
<figureCitation id="3AC82A44FF9923316CF33D14FE41F8E9" box="[414,482,1835,1861]" captionStart="FIGURE 3" captionStartId="5.[151,250,1964,1988]" captionTargetBox="[211,1322,265,1906]" captionTargetId="figure-49@5.[175,1412,237,1941]" captionTargetPageId="5" captionText="FIGURE 3. Preserved holotype of Oligodon cicadophagus sp. nov. (CUMZ-R-2625) with everted hemipenes in dorsal (A) and ventral (B) views. Photographs by M. Sumontha." figureDoi="http://doi.org/10.5281/zenodo.14734072" httpUri="https://zenodo.org/record/14734072/files/figure.png" pageId="4" pageNumber="349">Fig. 3</figureCitation>
); bilobed with large broad awn-shaped lobes; no distinct flounced calyces; unforked sulcus spermaticus.
</paragraph>
<paragraph id="A24C36C1FF9923306DAA3D4CFCECFF79" blockId="4.[151,1437,1187,2042]" lastBlockId="5.[151,1436,151,213]" lastPageId="5" lastPageNumber="350" pageId="4" pageNumber="349">
<emphasis id="9087EAD3FF9923316DAA3D4CFE2DF821" box="[199,398,1907,1933]" italics="true" pageId="4" pageNumber="349">Coloration in life</emphasis>
. Dorsal surface of the head uniformly dark olive-gray, without marks. No nuchal chevron. Lateral sides of the head olive-gray, lower half of supralabials cream.
<collectingRegion id="6037F823FF9923316ECD3DA8FC75F81D" box="[928,982,1943,1969]" country="Myanmar" name="Chin" pageId="4" pageNumber="349">Chin</collectingRegion>
and throat cream. Tongue purple brown. Dorsum and dorsal surface of the tail uniformly olive-gray, without marks or reticulations. Flanks olive-gray, turning to dark gray on the lower dorsal row. Venter cream anteriorly, progressively turning to dark gray posteriorly. Underside of tail dark gray with white blotches anteriorly, completely white posteriorly. In preservative, the dorsal color darkens, and the ventral colors become more contrasted.
</paragraph>
<caption id="F68C6649FF9823306DFA3D93FD29F844" ID-DOI="http://doi.org/10.5281/zenodo.14734072" ID-Zenodo-Dep="14734072" httpUri="https://zenodo.org/record/14734072/files/figure.png" pageId="5" pageNumber="350" startId="5.[151,250,1964,1988]" targetBox="[211,1322,265,1906]" targetPageId="5" targetType="figure">
<paragraph id="A24C36C1FF9823306DFA3D93FD29F844" blockId="5.[151,1437,1964,2024]" pageId="5" pageNumber="350">
<emphasis id="9087EAD3FF9823306DFA3D93FEB1F868" bold="true" box="[151,274,1964,1988]" pageId="5" pageNumber="350">FIGURE 3.</emphasis>
Preserved holotype of
<taxonomicName id="65F34D42FF9823306C963D93FD4EF868" authorityName="Pauwels &amp; Donbundit &amp; Sumontha &amp; Meesook" authorityYear="2025" box="[507,749,1964,1988]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="5" pageNumber="350" phylum="Chordata" rank="species" species="cicadophagus" status="sp. nov.">
<emphasis id="9087EAD3FF9823306C963D93FD4EF868" box="[507,749,1964,1988]" italics="true" pageId="5" pageNumber="350">Oligodon cicadophagus</emphasis>
</taxonomicName>
<emphasis id="9087EAD3FF9823306F9E3D93FCE1F868" bold="true" box="[755,834,1964,1988]" pageId="5" pageNumber="350">
<taxonomicNameLabel id="8BB457A8FF9823306F9E3D93FCE1F868" box="[755,834,1964,1988]" pageId="5" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CUMZ-R-2625) with everted hemipenes in dorsal (A) and ventral (B) views. Photographs by M. Sumontha.
</paragraph>
</caption>
<caption id="F68C6649FF9B23336DFA3DB8FEEBF86F" ID-DOI="http://doi.org/10.5281/zenodo.14734076" ID-Zenodo-Dep="14734076" httpUri="https://zenodo.org/record/14734076/files/figure.png" pageId="6" pageNumber="351" startId="6.[151,250,1927,1951]" targetBox="[151,1436,181,1903]" targetPageId="6" targetType="figure">
<paragraph id="A24C36C1FF9B23336DFA3DB8FEEBF86F" blockId="6.[151,1437,1927,1987]" pageId="6" pageNumber="351">
<emphasis id="9087EAD3FF9B23336DFA3DB8FEB2F833" bold="true" box="[151,273,1927,1951]" pageId="6" pageNumber="351">FIGURE 4.</emphasis>
Live paratype of
<taxonomicName id="65F34D42FF9B23336CD33DB8FD13F833" authorityName="Pauwels &amp; Donbundit &amp; Sumontha &amp; Meesook" authorityYear="2025" box="[446,688,1927,1951]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="6" pageNumber="351" phylum="Chordata" rank="species" species="cicadophagus" status="sp. nov.">
<emphasis id="9087EAD3FF9B23336CD33DB8FD13F833" box="[446,688,1927,1951]" italics="true" pageId="6" pageNumber="351">Oligodon cicadophagus</emphasis>
</taxonomicName>
<emphasis id="9087EAD3FF9B23336FD93DB8FCA0F833" bold="true" box="[692,771,1927,1951]" pageId="6" pageNumber="351">
<taxonomicNameLabel id="8BB457A8FF9B23336FD93DB8FCA0F833" box="[692,771,1927,1951]" pageId="6" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CUMZ-R-2626) in dorsal (A) and ventral (B) views. Photographs by M. Sumontha.
</paragraph>
</caption>
<caption id="F68C6649FF9A23326DFA3DA8FDA9F87F" ID-DOI="http://doi.org/10.5281/zenodo.14734078" ID-Zenodo-Dep="14734078" httpUri="https://zenodo.org/record/14734078/files/figure.png" pageId="7" pageNumber="352" startId="7.[151,250,1943,1967]" targetBox="[174,1331,214,1916]" targetPageId="7" targetType="figure">
<paragraph id="A24C36C1FF9A23326DFA3DA8FDA9F87F" blockId="7.[151,1436,1943,2003]" pageId="7" pageNumber="352">
<emphasis id="9087EAD3FF9A23326DFA3DA8FEB0F803" bold="true" box="[151,275,1943,1967]" pageId="7" pageNumber="352">FIGURE 5.</emphasis>
Live paratype of
<taxonomicName id="65F34D42FF9A23326CA53DA8FD18F803" authorityName="Pauwels &amp; Donbundit &amp; Sumontha &amp; Meesook" authorityYear="2025" box="[456,699,1943,1967]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="7" pageNumber="352" phylum="Chordata" rank="species" species="cicadophagus" status="sp. nov.">
<emphasis id="9087EAD3FF9A23326CA53DA8FD18F803" box="[456,699,1943,1967]" italics="true" pageId="7" pageNumber="352">Oligodon cicadophagus</emphasis>
</taxonomicName>
<emphasis id="9087EAD3FF9A23326FAC3DA8FCB1F803" bold="true" box="[705,786,1943,1967]" pageId="7" pageNumber="352">
<taxonomicNameLabel id="8BB457A8FF9A23326FAC3DA8FCB1F803" box="[705,786,1943,1967]" pageId="7" pageNumber="352" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CUMZ-R-2626) in dorsal view (A) and right profile (B) of the head. Photographs by M. Sumontha.
</paragraph>
</caption>
<paragraph id="A24C36C1FF95233D6DAA3AA9FE3FFEED" blockId="8.[151,1437,150,609]" pageId="8" pageNumber="353">
<emphasis id="9087EAD3FF95233D6DAA3AA9FE9CFF1C" bold="true" box="[199,319,150,176]" pageId="8" pageNumber="353">Variation.</emphasis>
Main morphometric and meristic characters of the
<specimenCount id="B4F5FD48FF95233D6E153AA8FC40FF1D" box="[888,995,151,177]" count="2" pageId="8" pageNumber="353" type="generic">two types</specimenCount>
are provided in
<tableCitation id="EF71037AFF95233D69FA3AA8FB49FF1D" box="[1175,1258,151,177]" captionStart="TABLE 1" captionStartId="13.[151,239,151,175]" captionTargetPageId="13" captionText="TABLE 1. Meristic and morphometric (in mm) data for the type-series of Oligodon cicadophagus sp. nov. Paired meristic characters are given left/right. Paired measurements are given for the right side. Supralabial numbers are followed in brackets by the ones contacting the orbit. Infralabial numbers are followed in brackets by how many contact the anterior chin shields. A = anal plate; NA = not assessed; S = single. For the other abbreviations, see Material and methods." pageId="8" pageNumber="353">Table 1</tableCitation>
. In the
<typeStatus id="7D488863FF95233D68513AA8FA3FFF1D" box="[1340,1436,151,177]" pageId="8" pageNumber="353" type="paratype">paratype</typeStatus>
(
<figureCitation id="3AC82A44FF95233D6DF23A84FEAAFF79" box="[159,265,186,213]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="6.[151,250,1927,1951]" captionStartId-1="7.[151,250,1943,1967]" captionTargetBox-0="[151,1436,181,1903]" captionTargetBox-1="[174,1331,214,1916]" captionTargetId-0="figure-35@6.[151,1436,181,1903]" captionTargetId-1="figure-23@7.[174,1414,181,1919]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 4. Live paratype of Oligodon cicadophagus sp. nov. (CUMZ-R-2626) in dorsal (A) and ventral (B) views. Photographs by M. Sumontha." captionText-1="FIGURE 5. Live paratype of Oligodon cicadophagus sp. nov. (CUMZ-R-2626) in dorsal view (A) and right profile (B) of the head. Photographs by M. Sumontha." figureDoi-0="http://doi.org/10.5281/zenodo.14734076" figureDoi-1="http://doi.org/10.5281/zenodo.14734078" httpUri-0="https://zenodo.org/record/14734076/files/figure.png" httpUri-1="https://zenodo.org/record/14734078/files/figure.png" pageId="8" pageNumber="353">Figs. 4–5</figureCitation>
) the
<collectionCode id="C4E2AE04FF95233D6C523A84FED6FF79" box="[319,373,187,213]" pageId="8" pageNumber="353">DSR</collectionCode>
reduction from 17 to 15 rows occurs above the 99
<superScript id="55869B89FF95233D6ECD3A85FC0EFF64" attach="left" box="[928,941,186,200]" fontSize="6" pageId="8" pageNumber="353">th</superScript>
(left) and 97
<superScript id="55869B89FF95233D69563A85FBEBFF64" attach="left" box="[1083,1096,186,200]" fontSize="6" pageId="8" pageNumber="353">th</superScript>
(right)
<collectionCode id="C4E2AE04FF95233D69F63A84FB76FF79" box="[1179,1237,187,213]" pageId="8" pageNumber="353">VEN</collectionCode>
by fusion of
<specimenCode id="F2559EBAFF95233D680B3A84FF06FF55" collectionCode="DSR" pageId="8" pageNumber="353">DSR 3</specimenCode>
and 4. The lower anterior temporal of the
<typeStatus id="7D488863FF95233D6FE93AE0FD47FF55" box="[644,740,223,249]" pageId="8" pageNumber="353" type="paratype">paratype</typeStatus>
is very small and does not contact the posterior temporal row (
<figureCitation id="3AC82A44FF95233D6DCD3B3CFF46FEB1" box="[160,229,258,285]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[174,1331,214,1916]" captionTargetId="figure-23@7.[174,1414,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Live paratype of Oligodon cicadophagus sp. nov. (CUMZ-R-2626) in dorsal view (A) and right profile (B) of the head. Photographs by M. Sumontha." figureDoi="http://doi.org/10.5281/zenodo.14734078" httpUri="https://zenodo.org/record/14734078/files/figure.png" pageId="8" pageNumber="353">Fig. 5</figureCitation>
). Similarly to the
<typeStatus id="7D488863FF95233D6CDB3B3CFDB8FEB1" box="[438,539,259,285]" pageId="8" pageNumber="353" type="holotype">holotype</typeStatus>
, the
<typeStatus id="7D488863FF95233D6F3F3B3CFD11FEB1" box="[594,690,259,285]" pageId="8" pageNumber="353" type="paratype">paratype</typeStatus>
shows six
<collectionCode id="C4E2AE04FF95233D6E5E3B3CFCFEFEB1" box="[819,861,259,285]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="8" pageNumber="353">MT</collectionCode>
present and five empty sockets, without diastema. Females still unknown.
</paragraph>
</subSubSection>
<subSubSection id="EAE9654AFF95233D6DAA3B75FA9CFDCD" pageId="8" pageNumber="353" type="multiple">
<paragraph id="A24C36C1FF95233D6DAA3B75FA9CFDCD" blockId="8.[151,1437,150,609]" pageId="8" pageNumber="353">
<emphasis id="9087EAD3FF95233D6DAA3B75FDF5FEC8" bold="true" box="[199,598,330,357]" pageId="8" pageNumber="353">Distribution and natural history.</emphasis>
Within the cave at the type locality (
<figureCitation id="3AC82A44FF95233D696A3B74FBEDFEC9" box="[1031,1102,331,357]" captionStart="FIGURE 6" captionStartId="8.[151,250,1446,1470]" captionTargetBox="[201,1386,633,1423]" captionTargetId="figure-274@8.[201,1386,633,1423]" captionTargetPageId="8" captionText="FIGURE 6. Biotope of Oligodon cicadophagus sp. nov. at the type-locality in a limestone cave at Sai Thai, Muang Krabi District, Krabi Province, southern Thailand. Photograph by N. Donbundit." figureDoi="http://doi.org/10.5281/zenodo.14734082" httpUri="https://zenodo.org/record/14734082/files/figure.png" pageId="8" pageNumber="353">Fig. 6</figureCitation>
), the
<typeStatus id="7D488863FF95233D69FC3B74FB50FEC9" box="[1169,1267,331,357]" pageId="8" pageNumber="353" type="holotype">holotype</typeStatus>
was caught in strict syntopy with the cave-dwelling colubrid
<taxonomicName id="65F34D42FF95233D6FCC3B50FBABFE25" authority="(Butler)" baseAuthorityName="Butler" baseAuthorityYear="1899" box="[673,1032,367,393]" class="Squamata" family="Colubridae" genus="Elaphe" kingdom="Animalia" pageId="8" pageNumber="353" phylum="Chordata" rank="subSpecies" species="taeniura" subSpecies="ridleyi">
<emphasis id="9087EAD3FF95233D6FCC3B50FC05FE25" box="[673,934,367,393]" italics="true" pageId="8" pageNumber="353">Elaphe taeniura ridleyi</emphasis>
(Butler)
</taxonomicName>
, and a few meters from the colubrid
<taxonomicName id="65F34D42FF95233D6DFA3BACFE0EFE01" authority="(Boie)" baseAuthorityName="Boie" baseAuthorityYear="1827" box="[151,429,403,429]" class="Squamata" family="Colubridae" genus="Ahaetulla" kingdom="Animalia" pageId="8" pageNumber="353" phylum="Chordata" rank="species" species="prasina">
<emphasis id="9087EAD3FF95233D6DFA3BACFEC3FE00" box="[151,352,403,429]" italics="true" pageId="8" pageNumber="353">Ahaetulla prasina</emphasis>
(Boie)
</taxonomicName>
and the pitviper
<taxonomicName id="65F34D42FF95233D6F0B3BACFC76FE01" authority="Vogel. An" authorityName="Vogel. An" box="[614,981,403,429]" class="Squamata" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" pageId="8" pageNumber="353" phylum="Chordata" rank="species" species="venustus">
<emphasis id="9087EAD3FF95233D6F0B3BACFCC2FE00" box="[614,865,403,428]" italics="true" pageId="8" pageNumber="353">Trimeresurus venustus</emphasis>
Vogel. An
</taxonomicName>
adult individual caught at night by snake rescuers at Khao Phung Chang (
<geoCoordinate id="C7C75006FF95233D6F6E3B88FD35FE7D" box="[515,662,439,465]" degrees="8" direction="north" minutes="26" orientation="latitude" pageId="8" pageNumber="353" precision="1" seconds="32.7" value="8.442417">8°26’32.7”N</geoCoordinate>
,
<geoCoordinate id="C7C75006FF95233D6FCE3B89FC9CFE7D" box="[675,831,438,465]" degrees="98" direction="east" minutes="30" orientation="longitude" pageId="8" pageNumber="353" precision="1" seconds="57.5" value="98.515976">98°30’57.5”E</geoCoordinate>
) in Phang-Nga Province regurgitated a cicada nymph (Cicadoidea) (T. Butmee, pers. comm.). Unfortunately, this latter individual escaped and could not be preserved, but was photographed (
<figureCitation id="3AC82A44FF95233D6CF63BC0FE7CFDB5" box="[411,479,511,537]" captionStart="FIGURE 7" captionStartId="9.[151,250,1020,1044]" captionTargetBox="[432,1156,181,996]" captionTargetId="figure-23@9.[432,1156,181,996]" captionTargetPageId="9" captionText="FIGURE 7. Live adult Oligodon cicadophagus sp. nov. (unvouchered) from Khao Phung Chang, Phang-Nga Province, southern Thailand. When caught, it regurgitated a cicada nymph. Photograph by T. Butmee." figureDoi="http://doi.org/10.5281/zenodo.14734084" httpUri="https://zenodo.org/record/14734084/files/figure.png" pageId="8" pageNumber="353">Fig. 7</figureCitation>
). This snake does not seem aggressive; none of the three known individuals attempted to bite when handled. The species is so far known only from the above-mentioned three localities within
<collectingRegion id="6037F823FF95233D6840381CFACEFD91" box="[1325,1389,547,573]" country="Thailand" name="Krabi" pageId="8" pageNumber="353">Krabi</collectingRegion>
and Phang-Nga provinces (
<figureCitation id="3AC82A44FF95233D6CF53878FE7EFDCD" box="[408,477,583,609]" captionStart="FIGURE 8" captionStartId="10.[151,250,1968,1992]" captionTargetBox="[206,1381,315,1939]" captionTargetId="figure-111@10.[200,1386,309,1944]" captionTargetPageId="10" captionText="FIGURE 8. Map of Thailand showing the geographical locations of the known localities of Oligodon cicadophagus sp. nov. in Krabi Province (full yellow circle, type locality) and Phang-Nga province (empty yellow circle). Map by W. Sodob." figureDoi="http://doi.org/10.5281/zenodo.14734086" httpUri="https://zenodo.org/record/14734086/files/figure.png" pageId="8" pageNumber="353">Fig. 8</figureCitation>
). The Phang-Nga locality is situated about
<quantity id="650B9B24FF95233D6EAC3879FBABFDCD" box="[961,1032,582,609]" metricMagnitude="4" metricUnit="m" metricValue="5.5" pageId="8" pageNumber="353" unit="km" value="55.0">55 km</quantity>
NW of the
<collectingRegion id="6037F823FF95233D69E63878FB68FDCD" box="[1163,1227,583,609]" country="Thailand" name="Krabi" pageId="8" pageNumber="353">Krabi</collectingRegion>
localities.
</paragraph>
</subSubSection>
<caption id="F68C6649FF95233D6DFA3F99FC2BFA4E" ID-DOI="http://doi.org/10.5281/zenodo.14734082" ID-Zenodo-Dep="14734082" httpUri="https://zenodo.org/record/14734082/files/figure.png" pageId="8" pageNumber="353" startId="8.[151,250,1446,1470]" targetBox="[201,1386,633,1423]" targetPageId="8" targetType="figure">
<paragraph id="A24C36C1FF95233D6DFA3F99FC2BFA4E" blockId="8.[151,1437,1446,1506]" pageId="8" pageNumber="353">
<emphasis id="9087EAD3FF95233D6DFA3F99FEB6FA12" bold="true" box="[151,277,1446,1470]" pageId="8" pageNumber="353">FIGURE 6.</emphasis>
Biotope of
<taxonomicName id="65F34D42FF95233D6CFB3F99FD28FA12" authorityName="Pauwels &amp; Donbundit &amp; Sumontha &amp; Meesook" authorityYear="2025" box="[406,651,1446,1470]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="8" pageNumber="353" phylum="Chordata" rank="species" species="cicadophagus" status="sp. nov.">
<emphasis id="9087EAD3FF95233D6CFB3F99FD28FA12" box="[406,651,1446,1470]" italics="true" pageId="8" pageNumber="353">Oligodon cicadophagus</emphasis>
</taxonomicName>
<emphasis id="9087EAD3FF95233D6FF93F99FD44FA12" bold="true" box="[660,743,1446,1470]" pageId="8" pageNumber="353">
<taxonomicNameLabel id="8BB457A8FF95233D6FF93F99FD44FA12" box="[660,743,1446,1470]" pageId="8" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
at the type-locality in a limestone cave at Sai Thai, Muang Krabi District, Krabi Province, southern Thailand. Photograph by N. Donbundit.
</paragraph>
</caption>
<subSubSection id="EAE9654AFF95233D6DAA3C0AFF5CF916" pageId="8" pageNumber="353" type="etymology">
<paragraph id="A24C36C1FF95233D6DAA3C0AFF5CF916" blockId="8.[151,1436,1588,1722]" pageId="8" pageNumber="353">
<emphasis id="9087EAD3FF95233D6DAA3C0AFEEDF9E3" bold="true" box="[199,334,1589,1615]" pageId="8" pageNumber="353">Etymology.</emphasis>
The specific epithet is a Latinized adjective meaning “cicada eater” and refers to the unusual prey item recovered from an individual in Phang-Nga Province,
<collectingCountry id="DAE47651FF95233D6EFD3C67FC54F9DE" box="[912,1015,1624,1650]" name="Thailand" pageId="8" pageNumber="353">Thailand</collectingCountry>
. We suggest the following common names: งูปี่แก้วกระบี่ (
<emphasis id="9087EAD3FF95233D6C163C42FDFEF93A" box="[379,605,1660,1686]" italics="true" pageId="8" pageNumber="353">
Ngu Pi Kaew
<collectingRegion id="6037F823FF95233D6F713C43FDFEF93A" box="[540,605,1660,1686]" country="Thailand" name="Krabi" pageId="8" pageNumber="353">Krabi</collectingRegion>
</emphasis>
, Thai), Cicada-eating Kukri Snake (English), and
<taxonomicName id="65F34D42FF95233D69F63C43FF58F916" authority="(French)" baseAuthorityName="French" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="8" pageNumber="353" phylum="Chordata" rank="species" species="cicadophage">
<emphasis id="9087EAD3FF95233D69F63C43FA3FF93A" box="[1179,1436,1660,1686]" italics="true" pageId="8" pageNumber="353">Oligodon cicadophage</emphasis>
(French)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="EAE9654AFF9523396DFA3D3DFBB3FA45" lastPageId="12" lastPageNumber="357" pageId="8" pageNumber="353" type="discussion">
<paragraph id="A24C36C1FF95233D6DFA3D3DFD80F8B0" blockId="8.[151,547,1794,1820]" box="[151,547,1794,1820]" pageId="8" pageNumber="353">
<heading id="F90481ADFF95233D6DFA3D3DFD80F8B0" bold="true" box="[151,547,1794,1820]" fontSize="11" level="1" pageId="8" pageNumber="353" reason="1">
<emphasis id="9087EAD3FF95233D6DFA3D3DFD80F8B0" bold="true" box="[151,547,1794,1820]" pageId="8" pageNumber="353">Comparisons with other species</emphasis>
</heading>
</paragraph>
<paragraph id="A24C36C1FF95233D6DFA3D75FC90F858" blockId="8.[151,1436,1866,2036]" pageId="8" pageNumber="353">
Among the
<taxonomicName id="65F34D42FF95233D6C4B3D75FE4EF8C8" box="[294,493,1866,1892]" pageId="8" pageNumber="353">
<emphasis id="9087EAD3FF95233D6C4B3D75FE33F8C8" box="[294,400,1866,1892]" italics="true" pageId="8" pageNumber="353">Oligodon</emphasis>
species
</taxonomicName>
found in the southern Indochinese and Thai-Malay regions, only the following seven ones consistently show no distinct nuchal chevron, no distinct dorsal transversal pattern (bands, marked reticulations), and no dorsal or supracaudal blotches:
<taxonomicName id="65F34D42FF95233D6E653DADFC26F800" authorityName="Leong &amp; Grismer" authorityYear="2004" box="[776,901,1938,1964]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="8" pageNumber="353" phylum="Chordata" rank="species" species="booliati">
<emphasis id="9087EAD3FF95233D6E653DADFC26F800" box="[776,901,1938,1964]" italics="true" pageId="8" pageNumber="353">O. booliati</emphasis>
</taxonomicName>
,
<taxonomicName id="65F34D42FF95233D6EFA3DADFBE5F800" authorityName="Nguyen, Nguyen, Le &amp; Murphy" authorityYear="2016" box="[919,1094,1938,1964]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="8" pageNumber="353" phylum="Chordata" rank="species" species="condaoensis">
<emphasis id="9087EAD3FF95233D6EFA3DADFBE5F800" box="[919,1094,1938,1964]" italics="true" pageId="8" pageNumber="353">O. condaoensis</emphasis>
</taxonomicName>
,
<taxonomicName id="65F34D42FF95233D69343DADFB6EF800" authorityName="Pauwels, Larsen, Suthanthangjai, David &amp; Sumontha" authorityYear="2017" box="[1113,1229,1938,1964]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="8" pageNumber="353" phylum="Chordata" rank="species" species="huahin">
<emphasis id="9087EAD3FF95233D69343DADFB6EF800" box="[1113,1229,1938,1964]" italics="true" pageId="8" pageNumber="353">O. huahin</emphasis>
</taxonomicName>
,
<taxonomicName id="65F34D42FF95233D69B33DADFACCF800" baseAuthorityName="Boulenger" baseAuthorityYear="1914" box="[1246,1391,1938,1964]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="8" pageNumber="353" phylum="Chordata" rank="species" species="inornatus">
<emphasis id="9087EAD3FF95233D69B33DADFACCF800" box="[1246,1391,1938,1964]" italics="true" pageId="8" pageNumber="353">O. inornatus</emphasis>
</taxonomicName>
,
<taxonomicName id="65F34D42FF95233D68EC3DADFF5AF87C" authorityName="Pauwels, Thongyai, Chantong &amp; Sumontha" authorityYear="2021" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="8" pageNumber="353" phylum="Chordata" rank="species" species="phangan">
<emphasis id="9087EAD3FF95233D68EC3DADFF5AF87C" italics="true" pageId="8" pageNumber="353">O. phangan</emphasis>
</taxonomicName>
,
<taxonomicName id="65F34D42FF95233D6C6B3D89FE19F87C" authorityName="Pauwels, Thongyai, Chantong &amp; Sumontha" authorityYear="2021" box="[262,442,1974,2000]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="8" pageNumber="353" phylum="Chordata" rank="species" species="promsombuti">
<emphasis id="9087EAD3FF95233D6C6B3D89FE19F87C" box="[262,442,1974,2000]" italics="true" pageId="8" pageNumber="353">O. promsombuti</emphasis>
</taxonomicName>
and
<taxonomicName id="65F34D42FF95233D6C803D89FD08F87C" authorityName="Pawangkhanant, Poyarkov, Ward-Smith, Grassby-Lewis, Sumontha, Kliukin, Idiiatullina, Trofimets, Suwannapoom &amp; Lee" authorityYear="2024" box="[493,683,1974,2000]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="8" pageNumber="353" phylum="Chordata" rank="species" species="speleoserpens">
<emphasis id="9087EAD3FF95233D6C803D89FD08F87C" box="[493,683,1974,2000]" italics="true" pageId="8" pageNumber="353">O. speleoserpens</emphasis>
</taxonomicName>
. It should be noted that a hatchling identified as
<taxonomicName id="65F34D42FF95233D69AC3D89FA3FF87C" baseAuthorityName="Boulenger" baseAuthorityYear="1914" box="[1217,1436,1974,2000]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="8" pageNumber="353" phylum="Chordata" rank="species" species="inornatus">
<emphasis id="9087EAD3FF95233D69AC3D89FA3FF87C" box="[1217,1436,1974,2000]" italics="true" pageId="8" pageNumber="353">Oligodon inornatus</emphasis>
</taxonomicName>
showed some black dorsal reticulations (
<bibRefCitation id="C6624B30FF95233D6F333DE5FC84F858" author="David, P. &amp; Teynie, A. &amp; Vogel, G." box="[606,807,2010,2036]" pageId="8" pageNumber="353" refId="ref7186" refString="David, P., Teynie, A. &amp; Vogel, G. (2023) The snakes of Laos. Edition Chimaira, Frankfurt am Main, 960 pp." type="book" year="2023">
David
<emphasis id="9087EAD3FF95233D6FC63DE4FD40F858" box="[683,739,2010,2036]" italics="true" pageId="8" pageNumber="353">et al.</emphasis>
2023
</bibRefCitation>
).
</paragraph>
<caption id="F68C6649FF94233C6DFA39C3FC7FFB9B" ID-DOI="http://doi.org/10.5281/zenodo.14734084" ID-Zenodo-Dep="14734084" httpUri="https://zenodo.org/record/14734084/files/figure.png" pageId="9" pageNumber="354" startId="9.[151,250,1020,1044]" targetBox="[432,1156,181,996]" targetPageId="9" targetType="figure">
<paragraph id="A24C36C1FF94233C6DFA39C3FC7FFB9B" blockId="9.[151,1436,1020,1080]" pageId="9" pageNumber="354">
<emphasis id="9087EAD3FF94233C6DFA39C3FEB2FBB8" bold="true" box="[151,273,1020,1044]" pageId="9" pageNumber="354">FIGURE 7.</emphasis>
Live adult
<taxonomicName id="65F34D42FF94233C6CEC39C3FDD0FBB8" authorityName="Pauwels &amp; Donbundit &amp; Sumontha &amp; Meesook" authorityYear="2025" box="[385,627,1020,1044]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="9" pageNumber="354" phylum="Chordata" rank="species" species="cicadophagus" status="sp. nov.">
<emphasis id="9087EAD3FF94233C6CEC39C3FDD0FBB8" box="[385,627,1020,1044]" italics="true" pageId="9" pageNumber="354">Oligodon cicadophagus</emphasis>
</taxonomicName>
<emphasis id="9087EAD3FF94233C6F1A39C3FD65FBB8" bold="true" box="[631,710,1020,1044]" pageId="9" pageNumber="354">
<taxonomicNameLabel id="8BB457A8FF94233C6F1A39C3FD65FBB8" box="[631,710,1020,1044]" pageId="9" pageNumber="354" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(unvouchered) from Khao Phung Chang, Phang-Nga Province, southern Thailand. When caught, it regurgitated a cicada nymph. Photograph by T. Butmee.
</paragraph>
</caption>
<paragraph id="A24C36C1FF94233C6DAA3EB5FDA2FA45" blockId="9.[151,1437,1162,2017]" pageId="9" pageNumber="354">
<taxonomicName id="65F34D42FF94233C6DAA3EB5FE74FB08" authorityName="Pauwels &amp; Donbundit &amp; Sumontha &amp; Meesook" authorityYear="2025" box="[199,471,1162,1188]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="9" pageNumber="354" phylum="Chordata" rank="species" species="cicadophagus" status="sp. nov.">
<emphasis id="9087EAD3FF94233C6DAA3EB5FE74FB08" box="[199,471,1162,1188]" italics="true" pageId="9" pageNumber="354">Oligodon cicadophagus</emphasis>
</taxonomicName>
<emphasis id="9087EAD3FF94233C6C8D3EB5FD9EFB08" bold="true" box="[480,573,1162,1188]" pageId="9" pageNumber="354">
<taxonomicNameLabel id="8BB457A8FF94233C6C8D3EB5FD9EFB08" box="[480,573,1162,1188]" pageId="9" pageNumber="354" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="65F34D42FF94233C6E163EB5FC54FB08" authorityName="Leong &amp; Grismer" authorityYear="2004" box="[891,1015,1162,1188]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="9" pageNumber="354" phylum="Chordata" rank="species" species="booliati">
<emphasis id="9087EAD3FF94233C6E163EB5FC54FB08" box="[891,1015,1162,1188]" italics="true" pageId="9" pageNumber="354">O. booliati</emphasis>
</taxonomicName>
of Pulau Tioman (= Tioman Island) by its distinctly higher maximum known SVL (
<quantity id="650B9B24FF94233C6FC33E91FCAEFB65" box="[686,781,1198,1225]" metricMagnitude="-1" metricUnit="m" metricValue="5.37" pageId="9" pageNumber="354" unit="mm" value="537.0">537 mm</quantity>
<emphasis id="9087EAD3FF94233C6E793E90FCFFFB64" box="[788,860,1199,1224]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
<quantity id="650B9B24FF94233C6E093E91FC65FB64" box="[868,966,1198,1224]" metricMagnitude="-1" metricUnit="m" metricValue="3.8899999999999997" pageId="9" pageNumber="354" unit="mm" value="389.0">389 mm</quantity>
), a distinctly lower TaL/TL ratio in males (0.15–0.17
<emphasis id="9087EAD3FF94233C6C763EECFEC0FB40" box="[283,355,1235,1260]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
0.25), higher
<collectionCode id="C4E2AE04FF94233C6F643EECFDE0FB41" box="[521,579,1235,1261]" pageId="9" pageNumber="354">VEN</collectionCode>
number (159–168
<emphasis id="9087EAD3FF94233C6E4C3EECFCCAFB40" box="[801,873,1235,1260]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
143–153), lower
<collectionCode id="C4E2AE04FF94233C69543EEDFBF8FB40" box="[1081,1115,1234,1260]" pageId="9" pageNumber="354">SC</collectionCode>
number (50
<emphasis id="9087EAD3FF94233C69993EECFA9FFB40" box="[1268,1340,1235,1260]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
54–60), higher
<collectionCode id="C4E2AE04FF94233C6D843EC9FEA9FABC" box="[233,266,1270,1296]" pageId="9" pageNumber="354">SL</collectionCode>
number (eight
<emphasis id="9087EAD3FF94233C6CD33EC8FDA5FABC" box="[446,518,1271,1296]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
six or seven), higher
<collectionCode id="C4E2AE04FF94233C6E653EC8FC81FABD" box="[776,802,1271,1297]" pageId="9" pageNumber="354">IL</collectionCode>
number (eight or nine
<emphasis id="9087EAD3FF94233C695E3EC8FBD8FABC" box="[1075,1147,1271,1296]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
seven), presence (
<emphasis id="9087EAD3FF94233C68383EC8FA3EFABC" box="[1365,1437,1271,1296]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
absence) of a presubocular, an olive-gray (
<emphasis id="9087EAD3FF94233C6F013F24FD17FA98" box="[620,692,1307,1332]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
red) dorsum, and a cream to gray (
<emphasis id="9087EAD3FF94233C695E3F24FBD8FA98" box="[1075,1147,1307,1332]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
dark orange) venter; from
<taxonomicName id="65F34D42FF94233C6DFA3F01FEE6FAF4" authorityName="Nguyen, Nguyen, Le &amp; Murphy" authorityYear="2016" box="[151,325,1342,1368]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="9" pageNumber="354" phylum="Chordata" rank="species" species="condaoensis">
<emphasis id="9087EAD3FF94233C6DFA3F01FEE6FAF4" box="[151,325,1342,1368]" italics="true" pageId="9" pageNumber="354">O. condaoensis</emphasis>
</taxonomicName>
from the Con Dao Islands by its higher TaL/TL ratio in males (0.15–0.17
<emphasis id="9087EAD3FF94233C69F73F00FB41FAF4" box="[1178,1250,1343,1368]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
0.13), distinctly higher
<collectionCode id="C4E2AE04FF94233C6D893F5DFEA5FAD0" box="[228,262,1378,1404]" pageId="9" pageNumber="354">SC</collectionCode>
number (50
<emphasis id="9087EAD3FF94233C6CFF3F5CFE79FAD0" box="[402,474,1379,1404]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
33–37), presence (
<emphasis id="9087EAD3FF94233C6FC13F5CFD57FAD0" box="[684,756,1379,1404]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
absence) of a presubocular, and olive-gray (
<emphasis id="9087EAD3FF94233C69B03F5CFA86FAD0" box="[1245,1317,1379,1404]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
dark gray) dorsum; from the western Thai
<taxonomicName id="65F34D42FF94233C6C983FB9FDC7FA0C" authorityName="Pauwels, Larsen, Suthanthangjai, David &amp; Sumontha" authorityYear="2017" box="[501,612,1414,1440]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="9" pageNumber="354" phylum="Chordata" rank="species" species="huahin">
<emphasis id="9087EAD3FF94233C6C983FB9FDC7FA0C" box="[501,612,1414,1440]" italics="true" pageId="9" pageNumber="354">O. huahin</emphasis>
</taxonomicName>
by its slightly higher TaL/TL ratio in males (0.15–0.17
<emphasis id="9087EAD3FF94233C69A33FB8FAB5FA0C" box="[1230,1302,1415,1440]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
0.12–0.14), distinctly higher
<collectionCode id="C4E2AE04FF94233C6C3B3F95FEDBFA68" box="[342,376,1450,1476]" pageId="9" pageNumber="354">SC</collectionCode>
number (50
<emphasis id="9087EAD3FF94233C6F673F94FDF1FA68" box="[522,594,1451,1476]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
35–41), olive-gray (
<emphasis id="9087EAD3FF94233C6E573F94FC21FA68" box="[826,898,1451,1476]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
light gray, tan or orange) dorsum, and cream to gray (
<emphasis id="9087EAD3FF94233C6DB43FF0FE82FA44" box="[217,289,1487,1512]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
plain white) venter.
</paragraph>
<paragraph id="A24C36C1FF94233C6DAA3FCCFE85F948" blockId="9.[151,1437,1162,2017]" pageId="9" pageNumber="354">
It is distinct from the Indochinese
<taxonomicName id="65F34D42FF94233C6F2C3FCDFD6DF9A0" baseAuthorityName="Boulenger" baseAuthorityYear="1914" box="[577,718,1522,1548]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="9" pageNumber="354" phylum="Chordata" rank="species" species="inornatus">
<emphasis id="9087EAD3FF94233C6F2C3FCDFD6DF9A0" box="[577,718,1522,1548]" italics="true" pageId="9" pageNumber="354">O. inornatus</emphasis>
</taxonomicName>
by its 17 (
<emphasis id="9087EAD3FF94233C6E2F3FCCFC29F9A0" box="[834,906,1523,1548]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
15)
<collectionCode id="C4E2AE04FF94233C6ED63FCDFC55F9A0" box="[955,1014,1522,1548]" pageId="9" pageNumber="354">MSR</collectionCode>
, lower
<collectionCode id="C4E2AE04FF94233C69253FCCFB21F9A1" box="[1096,1154,1523,1549]" pageId="9" pageNumber="354">VEN</collectionCode>
number (159–168
<emphasis id="9087EAD3FF94233C68383FCCFA3EF9A0" box="[1365,1437,1523,1548]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
169–173), higher
<collectionCode id="C4E2AE04FF94233C6C333C29FE23F99C" box="[350,384,1558,1584]" pageId="9" pageNumber="354">SC</collectionCode>
number (50
<emphasis id="9087EAD3FF94233C6F623C28FDF4F99C" box="[527,599,1559,1584]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
31–43), presence (
<emphasis id="9087EAD3FF94233C6E413C28FCD7F99C" box="[812,884,1559,1584]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
absence) of a presubocular, an olive-gray (
<emphasis id="9087EAD3FF94233C68383C28FA3EF99C" box="[1365,1437,1559,1584]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
dark greyish brown or reddish-brown) dorsum, and a cream to gray venter (
<emphasis id="9087EAD3FF94233C69653C04FBC6F9F8" box="[1032,1125,1595,1620]" italics="true" pageId="9" pageNumber="354">versus c</emphasis>
ream with rectangular dark blotches on sides; see
<figureCitation id="3AC82A44FF94233C6CE63C61FE75F9D4" box="[395,470,1630,1656]" captionStart="FIGURE 9" captionStartId="11.[151,250,1841,1865]" captionTargetBox="[176,1411,181,1817]" captionTargetId="figure-23@11.[176,1411,181,1817]" captionTargetPageId="11" captionText="FIGURE 9. General dorsal and ventral views of BMNH 1946.1.4.25, holotype of Oligodon inornatus, from Nong Kai Ploi, Chonburi Province, southeastern Thailand. Photographs by P. Campbell." figureDoi="http://doi.org/10.5281/zenodo.14734088" httpUri="https://zenodo.org/record/14734088/files/figure.png" pageId="9" pageNumber="354">Figs. 9</figureCitation>
and
<figureCitation id="3AC82A44FF94233C6F643C61FD8BF9D4" box="[521,552,1630,1656]" captionStart="FIGURE 10" captionStartId="12.[151,250,1021,1045]" captionTargetBox="[151,1436,181,997]" captionTargetId="figure-35@12.[151,1436,181,997]" captionTargetPageId="12" captionText="FIGURE 10. Right profile of BMNH 1969.1862, paratype of Oligodon inornatus, from “SE Siam”. Photograph by P. Campbell. Note the absence of presubocular scale." figureDoi="http://doi.org/10.5281/zenodo.14734090" httpUri="https://zenodo.org/record/14734090/files/figure.png" pageId="9" pageNumber="354">10</figureCitation>
). It is to be noted that the photograph of an individual from “
<collectingCountry id="DAE47651FF94233C69AE3C61FA8AF9D4" box="[1219,1321,1630,1656]" name="Thailand" pageId="9" pageNumber="354">Thailand</collectingCountry>
” provided by
<bibRefCitation id="C6624B30FF94233C6DD13CBDFE77F930" author="David, P. &amp; Teynie, A. &amp; Vogel, G." box="[188,468,1666,1693]" pageId="9" pageNumber="354" refId="ref7186" refString="David, P., Teynie, A. &amp; Vogel, G. (2023) The snakes of Laos. Edition Chimaira, Frankfurt am Main, 960 pp." type="book" year="2023">
David
<emphasis id="9087EAD3FF94233C6C673CBCFE9DF930" box="[266,318,1666,1692]" italics="true" pageId="9" pageNumber="354">et al</emphasis>
. (2023: 369)
</bibRefCitation>
to illustrate
<taxonomicName id="65F34D42FF94233C6F0A3CBDFCE6F930" baseAuthorityName="Boulenger" baseAuthorityYear="1914" box="[615,837,1666,1692]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="9" pageNumber="354" phylum="Chordata" rank="species" species="inornatus">
<emphasis id="9087EAD3FF94233C6F0A3CBDFCE6F930" box="[615,837,1666,1692]" italics="true" pageId="9" pageNumber="354">Oligodon inornatus</emphasis>
</taxonomicName>
was taken at the
<collectingCountry id="DAE47651FF94233C697F3CBDFBEFF930" box="[1042,1100,1666,1692]" name="Thailand" pageId="9" pageNumber="354">Siam</collectingCountry>
Serpentarium in
<collectingRegion id="6037F823FF94233C68793CBDFADAF930" box="[1300,1401,1666,1692]" country="Thailand" name="Krung Thep Mahanakhon" pageId="9" pageNumber="354">Bangkok</collectingRegion>
(I. Das, pers. comm.), and that its precise geographic origin is hence indeed unknown, and its taxonomic identity unverifiable.
</paragraph>
<paragraph id="A24C36C1FF94233F6DAA3CD1FC87FEB1" blockId="9.[151,1437,1162,2017]" lastBlockId="10.[151,1437,151,285]" lastPageId="10" lastPageNumber="355" pageId="9" pageNumber="354">
<taxonomicName id="65F34D42FF94233C6DAA3CD1FE77F8A4" authorityName="Pauwels &amp; Donbundit &amp; Sumontha &amp; Meesook" authorityYear="2025" box="[199,468,1774,1800]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="9" pageNumber="354" phylum="Chordata" rank="species" species="cicadophagus" status="sp. nov.">
<emphasis id="9087EAD3FF94233C6DAA3CD1FE77F8A4" box="[199,468,1774,1800]" italics="true" pageId="9" pageNumber="354">Oligodon cicadophagus</emphasis>
</taxonomicName>
<emphasis id="9087EAD3FF94233C6CB63CD1FD96F8A4" bold="true" box="[475,565,1774,1800]" pageId="9" pageNumber="354">
<taxonomicNameLabel id="8BB457A8FF94233C6CB63CD1FD96F8A4" box="[475,565,1774,1800]" pageId="9" pageNumber="354" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is readily separated from
<taxonomicName id="65F34D42FF94233C6E363CD1FC7CF8A4" authorityName="Pauwels, Thongyai, Chantong &amp; Sumontha" authorityYear="2021" box="[859,991,1774,1800]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="9" pageNumber="354" phylum="Chordata" rank="species" species="phangan">
<emphasis id="9087EAD3FF94233C6E363CD1FC7CF8A4" box="[859,991,1774,1800]" italics="true" pageId="9" pageNumber="354">O. phangan</emphasis>
</taxonomicName>
of Phangan Island (and possibly with a wider distribution, see
<bibRefCitation id="C6624B30FF94233C6CF43D2DFD6FF880" author="Pawangkhanant, P. &amp; Poyarkov, N. A. &amp; Ward-Smith, H. &amp; Grassby-Lewis, R. &amp; Sumontha, M. &amp; Kliukin, N. S. &amp; Idiiatullina, S. S. &amp; Trofimets, A. V. &amp; Suwannapoom, C. &amp; Lee, J. L." box="[409,716,1810,1837]" pageId="9" pageNumber="354" pagination="359 - 379" refId="ref9206" refString="Pawangkhanant, P., Poyarkov, N. A., Ward-Smith, H., Grassby-Lewis, R., Sumontha, M., Kliukin, N. S., Idiiatullina, S. S., Trofimets, A. V., Suwannapoom, C. &amp; Lee, J. L. (2024) A new species of karst-associated kukri snake (Reptilia: Squamata: Colubridae: Oligodon Fitzinger, 1826) from southern Thailand. Vertebrate Zoology, 74, 359 - 379. https: // doi. org / 10.3897 / vz. 74. e 112132" type="journal article" year="2024">
Pawangkhanant
<emphasis id="9087EAD3FF94233C6F3F3D2CFD27F880" box="[594,644,1810,1836]" italics="true" pageId="9" pageNumber="354">et al</emphasis>
. 2024
</bibRefCitation>
) by its distinctly higher maximum known SVL (
<quantity id="650B9B24FF94233C69823D2DFAEDF881" box="[1263,1358,1810,1837]" metricMagnitude="-1" metricUnit="m" metricValue="5.37" pageId="9" pageNumber="354" unit="mm" value="537.0">537 mm</quantity>
<emphasis id="9087EAD3FF94233C68383D2CFA3EF880" box="[1365,1437,1811,1836]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
<quantity id="650B9B24FF94233C6DFA3D09FF5AF8FC" box="[151,249,1846,1872]" metricMagnitude="-1" metricUnit="m" metricValue="3.69" pageId="9" pageNumber="354" unit="mm" value="369.0">369 mm</quantity>
), higher
<collectionCode id="C4E2AE04FF94233C6C333D09FE23F8FC" box="[350,384,1846,1872]" pageId="9" pageNumber="354">SC</collectionCode>
number (50
<emphasis id="9087EAD3FF94233C6F793D08FDFFF8FC" box="[532,604,1847,1872]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
33–42), presence (
<emphasis id="9087EAD3FF94233C6E5B3D08FCDDF8FC" box="[822,894,1847,1872]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
absence) of a presubocular, 11 (
<emphasis id="9087EAD3FF94233C699D3D08FA9BF8FC" box="[1264,1336,1847,1872]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
12)
<collectionCode id="C4E2AE04FF94233C68003D08FA36F8FD" box="[1389,1429,1847,1873]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="9" pageNumber="354">MT</collectionCode>
, olive-gray (
<emphasis id="9087EAD3FF94233C6C743D64FEC2F8D8" box="[281,353,1883,1908]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
light orange-brown) dorsum, and cream to gray (
<emphasis id="9087EAD3FF94233C6EEE3D64FC68F8D8" box="[899,971,1883,1908]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
pinkish orange) venter; from the southern Thai
<taxonomicName id="65F34D42FF94233C6DB83D41FE2EF834" authorityName="Pauwels, Thongyai, Chantong &amp; Sumontha" authorityYear="2021" box="[213,397,1918,1944]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="9" pageNumber="354" phylum="Chordata" rank="species" species="promsombuti">
<emphasis id="9087EAD3FF94233C6DB83D41FE2EF834" box="[213,397,1918,1944]" italics="true" pageId="9" pageNumber="354">O. promsombuti</emphasis>
</taxonomicName>
by its slightly higher TaL/TL ratio in males (0.15–0.17
<emphasis id="9087EAD3FF94233C69713D40FBC7F834" box="[1052,1124,1919,1944]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
0.14), lower
<collectionCode id="C4E2AE04FF94233C686E3D40FA9EF835" box="[1283,1341,1919,1945]" pageId="9" pageNumber="354">VEN</collectionCode>
number (159–168
<emphasis id="9087EAD3FF94233C6C673D9CFEF1F810" box="[266,338,1955,1980]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
177), higher
<collectionCode id="C4E2AE04FF94233C6C863D9DFDAEF810" box="[491,525,1954,1980]" pageId="9" pageNumber="354">SC</collectionCode>
number (50
<emphasis id="9087EAD3FF94233C6FF23D9CFD44F810" box="[671,743,1955,1980]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
40), single preocular (
<emphasis id="9087EAD3FF94233C6E8A3D9CFB8CF810" box="[999,1071,1955,1980]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
two), presence (
<emphasis id="9087EAD3FF94233C69863D9CFA90F810" box="[1259,1331,1955,1980]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
absence) of a presubocular, 11 (
<emphasis id="9087EAD3FF94233C6CF33DF8FE45F84C" box="[414,486,1991,2016]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
12)
<collectionCode id="C4E2AE04FF94233C6F4D3DF8FDEBF84D" box="[544,584,1991,2017]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="9" pageNumber="354">MT</collectionCode>
, olive-gray (
<emphasis id="9087EAD3FF94233C6FB23DF8FC84F84C" box="[735,807,1991,2016]" italics="true" pageId="9" pageNumber="354">versus</emphasis>
blackish brown) dorsum, and venter cream anteriorly, dark gray posteriorly (
<emphasis id="9087EAD3FF97233F6CFE3AA8FE78FF1C" box="[403,475,151,176]" italics="true" pageId="10" pageNumber="355">versus</emphasis>
ivory, heavily speckled with subrectangular blotches); and from the southern Thai
<taxonomicName id="65F34D42FF97233F68EC3AA8FE97FF79" authorityName="Pawangkhanant, Poyarkov, Ward-Smith, Grassby-Lewis, Sumontha, Kliukin, Idiiatullina, Trofimets, Suwannapoom &amp; Lee" authorityYear="2024" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="10" pageNumber="355" phylum="Chordata" rank="species" species="speleoserpens">
<emphasis id="9087EAD3FF97233F68EC3AA8FE97FF79" italics="true" pageId="10" pageNumber="355">O. speleoserpens</emphasis>
</taxonomicName>
by its higher TaL/TL ratio in males (0.15–0.17
<emphasis id="9087EAD3FF97233F6E2F3A84FC29FF78" box="[834,906,187,212]" italics="true" pageId="10" pageNumber="355">versus</emphasis>
0.14), distinctly smaller maximum known SVL (
<quantity id="650B9B24FF97233F6DF23AE1FF5CFF55" box="[159,255,222,249]" metricMagnitude="-1" metricUnit="m" metricValue="5.37" pageId="10" pageNumber="355" unit="mm" value="537.0">537 mm</quantity>
<emphasis id="9087EAD3FF97233F6C653AE0FEF3FF54" box="[264,336,223,248]" italics="true" pageId="10" pageNumber="355">versus</emphasis>
<quantity id="650B9B24FF97233F6C353AE0FE18FF55" box="[344,443,223,249]" metricMagnitude="-1" metricUnit="m" metricValue="6.81" pageId="10" pageNumber="355" unit="mm" value="681.0">681 mm</quantity>
), 17 (
<emphasis id="9087EAD3FF97233F6C933AE0FDE5FF54" box="[510,582,223,248]" italics="true" pageId="10" pageNumber="355">versus</emphasis>
generally 19)
<collectionCode id="C4E2AE04FF97233F6F863AE0FC82FF55" box="[747,801,223,249]" pageId="10" pageNumber="355">ASR</collectionCode>
, generally eight (
<emphasis id="9087EAD3FF97233F6E803AE0FB96FF54" box="[1005,1077,223,248]" italics="true" pageId="10" pageNumber="355">versus</emphasis>
nine)
<collectionCode id="C4E2AE04FF97233F69123AE0FB3AFF55" box="[1151,1177,223,249]" pageId="10" pageNumber="355">IL</collectionCode>
, distinctly lower
<collectionCode id="C4E2AE04FF97233F680E3AE0FA3EFF55" box="[1379,1437,223,249]" pageId="10" pageNumber="355">VEN</collectionCode>
number (159–168
<emphasis id="9087EAD3FF97233F6C0B3B3CFE0DFEB0" box="[358,430,259,284]" italics="true" pageId="10" pageNumber="355">versus</emphasis>
189–193), and 11 (
<emphasis id="9087EAD3FF97233F6FE43B3CFD72FEB0" box="[649,721,259,284]" italics="true" pageId="10" pageNumber="355">versus</emphasis>
8)
<collectionCode id="C4E2AE04FF97233F6F983B3CFCBEFEB1" box="[757,797,259,285]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="10" pageNumber="355">MT</collectionCode>
.
</paragraph>
<caption id="F68C6649FF97233F6DFA3D8FFA8DF840" ID-DOI="http://doi.org/10.5281/zenodo.14734086" ID-Zenodo-Dep="14734086" httpUri="https://zenodo.org/record/14734086/files/figure.png" pageId="10" pageNumber="355" startId="10.[151,250,1968,1992]" targetBox="[206,1381,315,1939]" targetPageId="10" targetType="figure">
<paragraph id="A24C36C1FF97233F6DFA3D8FFA8DF840" blockId="10.[151,1437,1968,2029]" pageId="10" pageNumber="355">
<emphasis id="9087EAD3FF97233F6DFA3D8FFEB1F864" bold="true" box="[151,274,1968,1992]" pageId="10" pageNumber="355">FIGURE 8.</emphasis>
Map of Thailand showing the geographical locations of the known localities of
<taxonomicName id="65F34D42FF97233F69573D8FFA8FF864" authorityName="Pauwels &amp; Donbundit &amp; Sumontha &amp; Meesook" authorityYear="2025" box="[1082,1324,1968,1992]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="10" pageNumber="355" phylum="Chordata" rank="species" species="cicadophagus" status="sp. nov.">
<emphasis id="9087EAD3FF97233F69573D8FFA8FF864" box="[1082,1324,1968,1992]" italics="true" pageId="10" pageNumber="355">Oligodon cicadophagus</emphasis>
</taxonomicName>
<emphasis id="9087EAD3FF97233F685F3D8FFA21F864" bold="true" box="[1330,1410,1968,1992]" pageId="10" pageNumber="355">
<taxonomicNameLabel id="8BB457A8FF97233F685F3D8FFA21F864" box="[1330,1410,1968,1992]" pageId="10" pageNumber="355" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in Krabi Province (full yellow circle, type locality) and Phang-Nga province (empty yellow circle). Map by W. Sodob.
</paragraph>
</caption>
<caption id="F68C6649FF96233E6DFA3D0EFCD5F8C1" ID-DOI="http://doi.org/10.5281/zenodo.14734088" ID-Zenodo-Dep="14734088" httpUri="https://zenodo.org/record/14734088/files/figure.png" pageId="11" pageNumber="356" startId="11.[151,250,1841,1865]" targetBox="[176,1411,181,1817]" targetPageId="11" targetType="figure">
<paragraph id="A24C36C1FF96233E6DFA3D0EFCD5F8C1" blockId="11.[151,1436,1841,1901]" pageId="11" pageNumber="356">
<emphasis id="9087EAD3FF96233E6DFA3D0EFEB6F8E5" bold="true" box="[151,277,1841,1865]" pageId="11" pageNumber="356">FIGURE 9.</emphasis>
General dorsal and ventral views of BMNH 1946.1.4.25, holotype of
<taxonomicName id="65F34D42FF96233E6E823D0EFB1BF8E5" baseAuthorityName="Boulenger" baseAuthorityYear="1914" box="[1007,1208,1841,1865]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="11" pageNumber="356" phylum="Chordata" rank="species" species="inornatus">
<emphasis id="9087EAD3FF96233E6E823D0EFB1BF8E5" box="[1007,1208,1841,1865]" italics="true" pageId="11" pageNumber="356">Oligodon inornatus</emphasis>
</taxonomicName>
, from Nong Kai Ploi, Chonburi Province, southeastern Thailand. Photographs by P. Campbell.
</paragraph>
</caption>
<caption id="F68C6649FF9123396DFA39C2FD8AFB95" ID-DOI="http://doi.org/10.5281/zenodo.14734090" ID-Zenodo-Dep="14734090" httpUri="https://zenodo.org/record/14734090/files/figure.png" pageId="12" pageNumber="357" startId="12.[151,250,1021,1045]" targetBox="[151,1436,181,997]" targetPageId="12" targetType="figure">
<paragraph id="A24C36C1FF9123396DFA39C2FD8AFB95" blockId="12.[151,1437,1021,1081]" pageId="12" pageNumber="357">
<emphasis id="9087EAD3FF9123396DFA39C2FEBCFBB9" bold="true" box="[151,287,1021,1045]" pageId="12" pageNumber="357">FIGURE 10.</emphasis>
Right profile of BMNH 1969.1862, paratype of
<taxonomicName id="65F34D42FF9123396E6E39C2FC6AFBB9" baseAuthorityName="Boulenger" baseAuthorityYear="1914" box="[771,969,1021,1045]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="12" pageNumber="357" phylum="Chordata" rank="species" species="inornatus">
<emphasis id="9087EAD3FF9123396E6E39C2FC6AFBB9" box="[771,969,1021,1045]" italics="true" pageId="12" pageNumber="357">Oligodon inornatus</emphasis>
</taxonomicName>
, from “SE Siam”. Photograph by P. Campbell. Note the absence of presubocular scale.
</paragraph>
</caption>
<paragraph id="A24C36C1FF9123396DAA3EB4FDA4FA99" blockId="12.[151,1437,1163,1514]" pageId="12" pageNumber="357">
Besides these species which consistently show a uniform pattern,
<taxonomicName id="65F34D42FF9123396ED83EB4FB26FB09" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[949,1157,1163,1189]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="12" pageNumber="357" phylum="Chordata" rank="species" species="cinereus">
<emphasis id="9087EAD3FF9123396ED83EB4FB26FB09" box="[949,1157,1163,1189]" italics="true" pageId="12" pageNumber="357">Oligodon cinereus</emphasis>
</taxonomicName>
, pending a confirmation of its current list of subjective synonyms (see
<bibRefCitation id="C6624B30FF9123396FF33E90FCC7FB65" author="David, P. &amp; Teynie, A. &amp; Vogel, G." box="[670,868,1199,1226]" pageId="12" pageNumber="357" refId="ref7186" refString="David, P., Teynie, A. &amp; Vogel, G. (2023) The snakes of Laos. Edition Chimaira, Frankfurt am Main, 960 pp." type="book" year="2023">
David
<emphasis id="9087EAD3FF9123396F873E8FFCB8FB65" box="[746,795,1199,1225]" italics="true" pageId="12" pageNumber="357">et al</emphasis>
. 2023
</bibRefCitation>
) displays a variety of dorsal patterns, among them an immaculate morph, the typical
<taxonomicName id="65F34D42FF9123396F7B3EECFD22FB41" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[534,641,1235,1261]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="12" pageNumber="357" phylum="Chordata" rank="species" species="cinereus">
“
<emphasis id="9087EAD3FF9123396F4E3EEBFD22FB41" box="[547,641,1236,1261]" italics="true" pageId="12" pageNumber="357">cinereus</emphasis>
</taxonomicName>
morph”, originally described from
<collectingCountry id="DAE47651FF912339697D3EECFB24FB41" box="[1040,1159,1235,1261]" name="Cambodia" pageId="12" pageNumber="357">Cambodia</collectingCountry>
. This immaculate morph differs from
<taxonomicName id="65F34D42FF9123396C4F3EC8FD8DFABD" authorityName="Pauwels &amp; Donbundit &amp; Sumontha &amp; Meesook" authorityYear="2025" box="[290,558,1271,1297]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="12" pageNumber="357" phylum="Chordata" rank="species" species="cicadophagus" status="sp. nov.">
<emphasis id="9087EAD3FF9123396C4F3EC8FD8DFABD" box="[290,558,1271,1297]" italics="true" pageId="12" pageNumber="357">Oligodon cicadophagus</emphasis>
</taxonomicName>
<emphasis id="9087EAD3FF9123396F593EC8FD2EFABD" bold="true" box="[564,653,1271,1297]" pageId="12" pageNumber="357">
<taxonomicNameLabel id="8BB457A8FF9123396F593EC8FD2EFABD" box="[564,653,1271,1297]" pageId="12" pageNumber="357" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by its lower
<collectionCode id="C4E2AE04FF9123396E763EC8FC9EFABD" box="[795,829,1271,1297]" pageId="12" pageNumber="357">SC</collectionCode>
number (27–43
<emphasis id="9087EAD3FF9123396E993EC7FB9FFABD" box="[1012,1084,1272,1297]" italics="true" pageId="12" pageNumber="357">versus</emphasis>
50) and its pinkish gray or dark gray (
<emphasis id="9087EAD3FF9123396DB43F23FE82FA99" box="[217,289,1308,1333]" italics="true" pageId="12" pageNumber="357">versus</emphasis>
olive-gray) dorsum.
</paragraph>
<paragraph id="A24C36C1FF9123396DAA3F00FBB3FA45" blockId="12.[151,1437,1163,1514]" pageId="12" pageNumber="357">
As shown by recent molecular analyses,
<emphasis id="9087EAD3FF9123396FE83F00FBAAFAF5" box="[645,1033,1343,1370]" italics="true" pageId="12" pageNumber="357">
<taxonomicName id="65F34D42FF9123396FE83F00FCF6FAF5" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[645,853,1343,1369]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="12" pageNumber="357" phylum="Chordata" rank="species" species="cinereus">Oligodon cinereus</taxonomicName>
,
<taxonomicName id="65F34D42FF9123396E323F7FFBAAFAF5" authorityName="Nguyen, Nguyen, Le &amp; Murphy" authorityYear="2016" box="[863,1033,1343,1370]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="12" pageNumber="357" phylum="Chordata" rank="species" species="condaoensis">O. condaoensis</taxonomicName>
</emphasis>
(if distinct from the former species),
<taxonomicName id="65F34D42FF9123396DFA3F5BFEAAFAD1" authorityName="Pauwels, Larsen, Suthanthangjai, David &amp; Sumontha" authorityYear="2017" box="[151,265,1379,1406]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="12" pageNumber="357" phylum="Chordata" rank="species" species="huahin">
<emphasis id="9087EAD3FF9123396DFA3F5BFEAAFAD1" box="[151,265,1379,1406]" italics="true" pageId="12" pageNumber="357">O. huahin</emphasis>
</taxonomicName>
,
<taxonomicName id="65F34D42FF9123396C743F5BFE0BFAD1" baseAuthorityName="Boulenger" baseAuthorityYear="1914" box="[281,424,1380,1406]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="12" pageNumber="357" phylum="Chordata" rank="species" species="inornatus">
<emphasis id="9087EAD3FF9123396C743F5BFE0BFAD1" box="[281,424,1380,1406]" italics="true" pageId="12" pageNumber="357">O. inornatus</emphasis>
</taxonomicName>
,
<taxonomicName id="65F34D42FF9123396CD53F5BFD9DFAD1" authorityName="Pauwels, Thongyai, Chantong &amp; Sumontha" authorityYear="2021" box="[440,574,1379,1406]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="12" pageNumber="357" phylum="Chordata" rank="species" species="phangan">
<emphasis id="9087EAD3FF9123396CD53F5BFD9DFAD1" box="[440,574,1379,1406]" italics="true" pageId="12" pageNumber="357">O. phangan</emphasis>
</taxonomicName>
,
<taxonomicName id="65F34D42FF9123396F233F5BFCA6FAD1" authorityName="Pauwels, Thongyai, Chantong &amp; Sumontha" authorityYear="2021" box="[590,773,1379,1406]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="12" pageNumber="357" phylum="Chordata" rank="species" species="promsombuti">
<emphasis id="9087EAD3FF9123396F233F5BFCA6FAD1" box="[590,773,1379,1406]" italics="true" pageId="12" pageNumber="357">O. promsombuti</emphasis>
</taxonomicName>
and
<taxonomicName id="65F34D42FF9123396E523F5BFBA3FAD1" authorityName="Pawangkhanant, Poyarkov, Ward-Smith, Grassby-Lewis, Sumontha, Kliukin, Idiiatullina, Trofimets, Suwannapoom &amp; Lee" authorityYear="2024" box="[831,1024,1379,1406]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="12" pageNumber="357" phylum="Chordata" rank="species" species="speleoserpens">
<emphasis id="9087EAD3FF9123396E523F5BFBA3FAD1" box="[831,1024,1379,1406]" italics="true" pageId="12" pageNumber="357">O. speleoserpens</emphasis>
</taxonomicName>
all belong to the
<taxonomicName id="65F34D42FF91233969A13F5CFA3FFAD1" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[1228,1436,1379,1405]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="12" pageNumber="357" phylum="Chordata" rank="species" species="cinereus">
<emphasis id="9087EAD3FF91233969A13F5CFA3FFAD1" box="[1228,1436,1379,1405]" italics="true" pageId="12" pageNumber="357">Oligodon cinereus</emphasis>
</taxonomicName>
group which is widely distributed from eastern
<collectingCountry id="DAE47651FF9123396FC03FB8FD44FA0D" box="[685,743,1415,1441]" name="India" pageId="12" pageNumber="357">India</collectingCountry>
to Sundaland (
<bibRefCitation id="C6624B30FF9123396EFD3FB8FBFBFA0D" author="David, P. &amp; Teynie, A. &amp; Vogel, G." box="[912,1112,1415,1442]" pageId="12" pageNumber="357" refId="ref7186" refString="David, P., Teynie, A. &amp; Vogel, G. (2023) The snakes of Laos. Edition Chimaira, Frankfurt am Main, 960 pp." type="book" year="2023">
David
<emphasis id="9087EAD3FF9123396EB03FB7FBACFA0D" box="[989,1039,1415,1441]" italics="true" pageId="12" pageNumber="357">et al</emphasis>
. 2023
</bibRefCitation>
;
<bibRefCitation id="C6624B30FF91233969093FB8FA3BFA0D" author="Pawangkhanant, P. &amp; Poyarkov, N. A. &amp; Ward-Smith, H. &amp; Grassby-Lewis, R. &amp; Sumontha, M. &amp; Kliukin, N. S. &amp; Idiiatullina, S. S. &amp; Trofimets, A. V. &amp; Suwannapoom, C. &amp; Lee, J. L." box="[1124,1432,1415,1441]" pageId="12" pageNumber="357" pagination="359 - 379" refId="ref9206" refString="Pawangkhanant, P., Poyarkov, N. A., Ward-Smith, H., Grassby-Lewis, R., Sumontha, M., Kliukin, N. S., Idiiatullina, S. S., Trofimets, A. V., Suwannapoom, C. &amp; Lee, J. L. (2024) A new species of karst-associated kukri snake (Reptilia: Squamata: Colubridae: Oligodon Fitzinger, 1826) from southern Thailand. Vertebrate Zoology, 74, 359 - 379. https: // doi. org / 10.3897 / vz. 74. e 112132" type="journal article" year="2024">
Pawangkhanant
<emphasis id="9087EAD3FF91233968703FB7FAF6FA0D" box="[1309,1365,1415,1441]" italics="true" pageId="12" pageNumber="357">et al.</emphasis>
2024
</bibRefCitation>
;
<bibRefCitation id="C6624B30FF9123396DFA3F93FEE9FA69" author="Lee, J. L. &amp; Yushchenko, P. V. &amp; Suwannapoom, C. &amp; Pawangkhanant, P. &amp; Grismer, L. L. &amp; Nguyen, T. V. &amp; Deepak, V. &amp; Narayanan, S. &amp; Das, S. &amp; Neang, T. &amp; Lalremsanga, H. T. &amp; Yang, J. - H. &amp; Jablonski, D. &amp; Erkaya, M. &amp; Vogel, G. &amp; Bauer, A. M. &amp; Poyarkov, N. A." box="[151,330,1451,1478]" pageId="12" pageNumber="357" refId="ref8160" refString="Lee, J. L., Yushchenko, P. V., Suwannapoom, C., Pawangkhanant, P., Grismer, L. L., Nguyen, T. V., Deepak, V., Narayanan, S., Das, S., Neang, T., Lalremsanga, H. T., Yang, J. - H., Jablonski, D., Erkaya, M., Vogel, G., Bauer, A. M. &amp; Poyarkov, N. A. (2024) A multi-gene phylogeny of the Asian kukri snakes (Oligodon Fitzinger, 1826): sharpening the blade of the second largest serpent radiation (Reptilia: Squamata: Colubridae). Molecular Phylogenetics and Evolution, 201, 108215. https: // doi. org / 10.1016 / j. ympev. 2024.108215" type="journal volume" year="2024">
Lee
<emphasis id="9087EAD3FF9123396DA73F93FF5DFA69" box="[202,254,1451,1477]" italics="true" pageId="12" pageNumber="357">et al</emphasis>
. 2024
</bibRefCitation>
). The hemipenes of
<taxonomicName id="65F34D42FF9123396F593F94FCE0FA69" authorityName="Pauwels &amp; Donbundit &amp; Sumontha &amp; Meesook" authorityYear="2025" box="[564,835,1451,1477]" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="12" pageNumber="357" phylum="Chordata" rank="species" species="cicadophagus" status="sp. nov.">
<emphasis id="9087EAD3FF9123396F593F94FCE0FA69" box="[564,835,1451,1477]" italics="true" pageId="12" pageNumber="357">Oligodon cicadophagus</emphasis>
</taxonomicName>
<emphasis id="9087EAD3FF9123396E213F94FC04FA69" bold="true" box="[844,935,1451,1477]" pageId="12" pageNumber="357">
<taxonomicNameLabel id="8BB457A8FF9123396E213F94FC04FA69" box="[844,935,1451,1477]" pageId="12" pageNumber="357" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resemble those of other members of the
<taxonomicName id="65F34D42FF91233968EC3F93FF56FA45" baseAuthorityName="Gunther" baseAuthorityYear="1864" class="Squamata" family="Colubridae" genus="Oligodon" kingdom="Animalia" pageId="12" pageNumber="357" phylum="Chordata" rank="species" species="cinereus">
<emphasis id="9087EAD3FF91233968EC3F93FF56FA45" italics="true" pageId="12" pageNumber="357">O. cinereus</emphasis>
</taxonomicName>
group (
<tableCitation id="EF71037AFF9123396C233FF0FE00FA45" box="[334,419,1487,1513]" captionStart="TABLE 2" captionStartId="14.[150,238,485,511]" captionTargetPageId="14" captionText="TABLE 2. Comparison of Oligodon species of the southern Indochinese and Thai-Malay region showing no distinct nuchal chevron, no distinct transversal pattern and no dorsal or supracaudal blotches. Bolded values are diagnostic differences from Oligodon cicadophagus sp. nov. NA = not available." pageId="12" pageNumber="357">Table 2</tableCitation>
;
<bibRefCitation id="C6624B30FF9123396CC33FF0FD66FA45" author="Yushchenko, P. V. &amp; Lee, J. L. &amp; Neang, T. &amp; Hun, S. &amp; Nguyen, T. V. &amp; Vogel, G. &amp; Poyarkov, N. A. Jr." box="[430,709,1487,1514]" pageId="12" pageNumber="357" pagination="75 - 96" refId="ref10637" refString="Yushchenko, P. V., Lee, J. L., Neang, T., Hun, S., Nguyen, T. V., Vogel, G. &amp; Poyarkov, N. A. Jr. (2023 a) A taxonomic re-assessment of Oligodon cinereus (Gunther, 1864) (Squamata, Serpentes, Colubridae) populations from southern Indochina. Vertebrate Zoology, 73, 75 - 96. https: // doi. org / 10.3897 / vz. 73. e 91230" type="journal article" year="2023">
Yushchenko
<emphasis id="9087EAD3FF9123396F533FEFFDCCFA45" box="[574,623,1487,1513]" italics="true" pageId="12" pageNumber="357">et al</emphasis>
. 2023a
</bibRefCitation>
;
<bibRefCitation id="C6624B30FF9123396FBD3FF0FBA6FA45" author="Pawangkhanant, P. &amp; Poyarkov, N. A. &amp; Ward-Smith, H. &amp; Grassby-Lewis, R. &amp; Sumontha, M. &amp; Kliukin, N. S. &amp; Idiiatullina, S. S. &amp; Trofimets, A. V. &amp; Suwannapoom, C. &amp; Lee, J. L." box="[720,1029,1487,1514]" pageId="12" pageNumber="357" pagination="359 - 379" refId="ref9206" refString="Pawangkhanant, P., Poyarkov, N. A., Ward-Smith, H., Grassby-Lewis, R., Sumontha, M., Kliukin, N. S., Idiiatullina, S. S., Trofimets, A. V., Suwannapoom, C. &amp; Lee, J. L. (2024) A new species of karst-associated kukri snake (Reptilia: Squamata: Colubridae: Oligodon Fitzinger, 1826) from southern Thailand. Vertebrate Zoology, 74, 359 - 379. https: // doi. org / 10.3897 / vz. 74. e 112132" type="journal article" year="2024">
Pawangkhanant
<emphasis id="9087EAD3FF9123396EE43FEFFC19FA45" box="[905,954,1487,1513]" italics="true" pageId="12" pageNumber="357">et al</emphasis>
. 2024
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>