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<document ENCODING="UTF-8" ID-GBIF-Dataset="dd1fc5a5-a514-4720-a35e-2866e04e6a48" ModsDocID="z01639p001" checkinTime="1247323225858" checkinUser="thomas" docAuthor="William T. White, Peter R. Last & John D. Stevens" docDate="2007" docId="17C821753D8B2C09EDB94FD7607324A0" docLanguage="en" docName="2007_White_Last_Stevens_gg1.xml" docOrigin="Zootaxa 1639" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:84152FF4-E49B-4049-A9FC-E558F585D210" docTitle="Halaelurus maculosus White, Last & Stevens, 2007, new species" docType="treatment" docVersion="13" lastPageNumber="10" masterDocId="1B382B290DCA6874CFBE13D02DA0DAB8" masterDocTitle="Halaelurus maculosus n. sp. and H. sellus n. sp., two new species of catshark (Carcharhiniformes: Scyliorhinidae) from the Indo-West Pacific." masterLastPageNumber="21" masterPageNumber="1" pageNumber="2" updateTime="1677102616916" updateUser="ExternalLinkService">
|
||||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo>
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||||
<mods:title>Halaelurus maculosus n. sp. and H. sellus n. sp., two new species of catshark (Carcharhiniformes: Scyliorhinidae) from the Indo-West Pacific.</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>William T. White</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
|
||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>Peter R. Last</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:name type="personal">
|
||||
<mods:role>
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||||
<mods:roleTerm>Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>John D. Stevens</mods:namePart>
|
||||
</mods:name>
|
||||
<mods:typeOfResource>text</mods:typeOfResource>
|
||||
<mods:relatedItem type="host">
|
||||
<mods:titleInfo>
|
||||
<mods:title>Zootaxa</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:part>
|
||||
<mods:date>2007</mods:date>
|
||||
<mods:detail type="volume">
|
||||
<mods:number>1639</mods:number>
|
||||
</mods:detail>
|
||||
<mods:extent unit="page">
|
||||
<mods:start>1</mods:start>
|
||||
<mods:end>21</mods:end>
|
||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:location>
|
||||
<mods:url>http://www.zoobank.org/urn:lsid:zoobank.org:pub:84152FF4-E49B-4049-A9FC-E558F585D210</mods:url>
|
||||
</mods:location>
|
||||
<mods:classification>journal article</mods:classification>
|
||||
<mods:identifier type="ZooBank-PUB">z01639p001</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6243381" ID-GBIF-Taxon="100117182" ID-Zenodo-Dep="6243381" LSID="urn:lsid:plazi:treatment:17C821753D8B2C09EDB94FD7607324A0" httpUri="http://treatment.plazi.org/id/17C821753D8B2C09EDB94FD7607324A0" lastPageNumber="10" pageNumber="2">
|
||||
<subSubSection type="nomenclature">
|
||||
<paragraph pageNumber="2">
|
||||
<taxonomicName ID-CoL="6KXN4" class="Elasmobranchii" family="Scyliorhinidae" genus="Halaelurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="2" phylum="Chordata" rank="species" species="maculosus">Halaelurus maculosus</taxonomicName>
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
<paragraph pageNumber="2">Indonesian Speckled Catshark</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph pageNumber="2">Figs 1-4; Tables 1 and 2.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="reference_group">
|
||||
<paragraph pageNumber="2">
|
||||
<taxonomicName class="Elasmobranchii" family="Scyliorhinidae" genus="Halaelurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="2" phylum="Chordata" rank="species" species="buergeri">Halaelurus cf buergeri</taxonomicName>
|
||||
: White et al., 2006: 178, 179.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph pageNumber="2">
|
||||
<materialsCitation ID-GBIF-Occurrence="923907785" collectingDate="2002-07" collectionCode="MZB" country="Indonesia" latitude="-8.75" location="Kedonganan fish market" longitude="115.166664" specimenCode="MZB 15499" stateProvince="Bali" typeStatus="Holotype">
|
||||
<typeStatus>Holotype</typeStatus>
|
||||
.
|
||||
<specimenCode>
|
||||
<collectionCode>MZB</collectionCode>
|
||||
15499
|
||||
</specimenCode>
|
||||
, adult male 436 mm TL,
|
||||
<location LSID="urn:lsid:plazi:treatment:17C821753D8B2C09EDB94FD7607324A0:6152CCD7D604CA4567EC40B37DBF0C36" country="Indonesia" latitude="-8.75" longitude="115.166664" name="Kedonganan fish market" stateProvince="Bali">Kedonganan fish market</location>
|
||||
,
|
||||
<collectingRegion>Bali</collectingRegion>
|
||||
,
|
||||
<collectingCountry>Indonesia</collectingCountry>
|
||||
,
|
||||
<geoCoordinate direction="south" orientation="latitude" value="-8.75">08°45' S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate direction="east" orientation="longitude" value="115.166664">115° 10' E</geoCoordinate>
|
||||
,
|
||||
<date month="07" value="2002-07" year="2002">
|
||||
<collectingDate month="07" value="2002-07" year="2002">July 2002</collectingDate>
|
||||
</date>
|
||||
.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph pageNumber="2">
|
||||
<materialsCitation ID-GBIF-Occurrence="923907784" collectingDate="2002-07" collectionCode="CSIRO, MZB" country="Indonesia" latitude="-8.75" location="Kedonganan fish market" longitude="115.166664" specimenCode="CSIRO H 5889 - 09, CSIRO H 5889 - 16, CSIRO H 5889 - 17, CSIRO H 5889 - 18, CSIRO H 5890 - 01, MZB 15550" stateProvince="Bali" typeStatus="Paratypes">
|
||||
<typeStatus>Paratypes</typeStatus>
|
||||
. 7 specimens:
|
||||
<specimenCode>
|
||||
<collectionCode>CSIRO</collectionCode>
|
||||
H 5889-09
|
||||
</specimenCode>
|
||||
, adult male 457 mm TL,
|
||||
<specimenCode>
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||||
<collectionCode>CSIRO</collectionCode>
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||||
H 5889-16
|
||||
</specimenCode>
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||||
, adult male 439 mm TL,
|
||||
<specimenCode>
|
||||
<collectionCode>CSIRO</collectionCode>
|
||||
H 5889-17
|
||||
</specimenCode>
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||||
, adult male 444 mm TL,
|
||||
<specimenCode>
|
||||
<collectionCode>CSIRO</collectionCode>
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||||
H 5889-18
|
||||
</specimenCode>
|
||||
, adult male 430 mm TL,
|
||||
<specimenCode>
|
||||
<collectionCode>CSIRO</collectionCode>
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||||
H 5890-01
|
||||
</specimenCode>
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||||
, immature male 315 mm TL,
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||||
<specimenCode>
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||||
<collectionCode>MZB</collectionCode>
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||||
15550
|
||||
</specimenCode>
|
||||
, adult male 421 mm TL,
|
||||
<location LSID="urn:lsid:plazi:treatment:17C821753D8B2C09EDB94FD7607324A0:76E46E03C6432FE2DF79747617CBCD0A" country="Indonesia" latitude="-8.75" longitude="115.166664" name="Kedonganan fish market" stateProvince="Bali">collected at same locality as holotype</location>
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation ID-GBIF-Occurrence="923907794" collectingDate="2001" collectionCode="CSIRO" country="Indonesia" latitude="-7.6666665" location="Cilacap landing site" longitude="109.0" specimenCode="CSIRO H 5858 - 01" stateProvince="Central Java" typeStatus="Paratype">
|
||||
<specimenCode>
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||||
<collectionCode>CSIRO</collectionCode>
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||||
H 5858-01
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||||
</specimenCode>
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||||
, female 426 mm TL,
|
||||
<location LSID="urn:lsid:plazi:treatment:17C821753D8B2C09EDB94FD7607324A0:8D00F69EF0C85E08605559602BFF358E" country="Indonesia" latitude="-7.6666665" longitude="109.0" name="Cilacap landing site" stateProvince="Central Java">Cilacap landing site</location>
|
||||
,
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||||
<collectingRegion>Central Java</collectingRegion>
|
||||
,
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||||
<collectingCountry>Indonesia</collectingCountry>
|
||||
,
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||||
<geoCoordinate direction="south" orientation="latitude" value="-7.6666665">07°40' S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate direction="east" orientation="longitude" value="109.0">109°00' E</geoCoordinate>
|
||||
,
|
||||
<collectingDate>2001</collectingDate>
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation ID-GBIF-Occurrence="923907788" collectingDate="2006" collectionCode="CSIRO" country="Indonesia" latitude="-8.75" location="Tanjung Luar landing site" longitude="116.583336" specimenCode="CSIRO H 6497 - 02" stateProvince="Lombok" typeStatus="Paratypes">
|
||||
<specimenCode>
|
||||
<collectionCode>CSIRO</collectionCode>
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||||
H 6497-02
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||||
</specimenCode>
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||||
, 17 egg cases,
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||||
<location LSID="urn:lsid:plazi:treatment:17C821753D8B2C09EDB94FD7607324A0:C0EBDD219825F45861FD2A66B632E3D4" country="Indonesia" latitude="-8.75" longitude="116.583336" name="Tanjung Luar landing site" stateProvince="Lombok">Tanjung Luar landing site</location>
|
||||
,
|
||||
<collectingRegion>Lombok</collectingRegion>
|
||||
,
|
||||
<collectingCountry>Indonesia</collectingCountry>
|
||||
,
|
||||
<geoCoordinate direction="south" orientation="latitude" value="-8.75">08°45'S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate direction="east" orientation="longitude" value="116.583336">116°35'E</geoCoordinate>
|
||||
,
|
||||
<collectingDate>2006</collectingDate>
|
||||
.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph pageNumber="2">
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Other material. 14 specimens:
|
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<materialsCitation ID-GBIF-Occurrence="923907787" collectingDate="2002-07" collectionCode="CSIRO, MZB" country="Indonesia" latitude="-8.75" location="Kedonganan fish market" longitude="115.166664" specimenCode="CSIRO H 5889 - 14, CSIRO H 5889 - 25, CSIRO H 5890 - 02, MZB 15446, MZB 15448, MZB 15449, MZB 15459, MZB 15462, MZB 15464, MZB 15465, MZB 15466, MZB 15467, MZB 15490" stateProvince="Bali">
|
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<specimenCode>
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<collectionCode>CSIRO</collectionCode>
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H 5889-14
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</specimenCode>
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, adult male 434 mm TL,
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<specimenCode>
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<collectionCode>CSIRO</collectionCode>
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H 5889-25
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</specimenCode>
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, immature male 344 mm TL,
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<specimenCode>
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<collectionCode>CSIRO</collectionCode>
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H 5890-02
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</specimenCode>
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, adolescent male, 360 mm TL,
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<specimenCode>
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<collectionCode>MZB</collectionCode>
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15446
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</specimenCode>
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, adult male 426 mm TL,
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<specimenCode>
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<collectionCode>MZB</collectionCode>
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15448
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</specimenCode>
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, adult male 411 mm TL,
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<specimenCode>
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<collectionCode>MZB</collectionCode>
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15449
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</specimenCode>
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, adolescent male 413 mm TL,
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<specimenCode>
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<collectionCode>MZB</collectionCode>
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15459
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</specimenCode>
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, adult male 430 mm TL,
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<specimenCode>
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<collectionCode>MZB</collectionCode>
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15462
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</specimenCode>
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, adult male 433 mm TL,
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<specimenCode>
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<collectionCode>MZB</collectionCode>
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15464
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</specimenCode>
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, adult male 444 mm TL,
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<specimenCode>
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<collectionCode>MZB</collectionCode>
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15465
|
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</specimenCode>
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, adult male 400 mm TL,
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<specimenCode>
|
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<collectionCode>MZB</collectionCode>
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15466
|
||||
</specimenCode>
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, female 453 mm TL,
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<specimenCode>
|
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<collectionCode>MZB</collectionCode>
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15467
|
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</specimenCode>
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, adult male 412 mm TL,
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<specimenCode>
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<collectionCode>MZB</collectionCode>
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15490
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</specimenCode>
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, adult male 441 mm TL,
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<location LSID="urn:lsid:plazi:treatment:17C821753D8B2C09EDB94FD7607324A0:201CC53E7CDA59F748CE991C54482CB7" country="Indonesia" latitude="-8.75" longitude="115.166664" name="Kedonganan fish market" stateProvince="Bali">collected at same locality as holotype</location>
|
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</materialsCitation>
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||||
;
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<materialsCitation ID-GBIF-Occurrence="923907795" collectingDate="1999" collectionCode="JPAG" country="Philippines" location="Philippines" specimenCode="JPAG 114">
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<specimenCode>
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<collectionCode>JPAG</collectionCode>
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114
|
||||
</specimenCode>
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||||
, female 507 mm TL,
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<location LSID="urn:lsid:plazi:treatment:17C821753D8B2C09EDB94FD7607324A0:7BCAF8D390B5C966BDB21A2011920D1F" country="Philippines" name="Philippines">
|
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<collectingCountry>Philippines</collectingCountry>
|
||||
</location>
|
||||
,
|
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<collectingDate>1999</collectingDate>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
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||||
<subSubSection type="diagnosis">
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||||
<paragraph pageNumber="5">
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||||
Diagnosis. Member of the genus
|
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:EF8B4C66-B5D8-494A-A976-671A4547EB5E" class="Elasmobranchii" family="Scyliorhinidae" genus="Halaelurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Halaelurus Gill 1862:407" order="Carcharhiniformes" pageNumber="5" phylum="Chordata" rank="genus">Halaelurus</taxonomicName>
|
||||
with the following combination of characters: a relatively short, broadly rounded snout, preoral length 3.9-4.9% TL; predorsal length 41.6-42.8% TL, 3.8-4.7 times anal -caudal space; anal -caudal space 9.0-10.9% TL; mouth moderately long, length 2.8-3.6% TL; upper labial furrows very short (8.1-14.9 in eye length), sometimes inconspicuous; eyes moderately large, eye length 3.1-4.0% TL; pectoral fin with broadly rounded apex, anterior margin 10.8-11.7% TL, 1.9-2.3 times inner margin; anal fin relatively large, length 11.0-12.2% TL, anterior margin 8.0-9.1% TL; adult clasper elongate and relatively narrow, outer length 6.4-7.5% TL, 3.6-5.0 times clasper base length, extending past pelvic-fin free rear tip by 0.25-0.36 times pelvic-fin inner margin length, clasper tip narrowly rounded; total vertebral centra 127-132, precaudal centra 81-90; dorsal surface with broad, dark brownish saddles, densely covered with numerous, dark brown spots; spots present on interspaces between saddles. Small adult size, attaining about 530 mm TL.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph pageNumber="5">Description. Proportions as percentages of total length for the holotype and paratypes (7) are presented in Table 1.</paragraph>
|
||||
<paragraph pageNumber="5">Body tapering, anterior trunk wide and somewhat depressed; length of trunk from fifth gill openings to vent 1.06 (1.02-1.30) times head length; tail long and slender, almost circular in cross-section at second dorsal-fin insertion; no lateral keels; height 1.21 (0.91-1.10) in width at second dorsal-fin insertion, 3.28 (2.62- 3.08) in dorsal -caudal space; no postnatal ridge between anal fin base and lower caudal-fin origin; no longitudinal ridges on dorsal surface, no obvious lateral ridges. Head short, length 1.31 (1.04-1.33) to pectoral -pelvic space; wide and somewhat depressed, more so between eyes and ventrally; narrowly pointed, not upturned or knob-like in lateral view (slightly upturned in some paratypes); in dorsoventral view anterior to gill openings broadly parabolic; subocular ridge evident, variably angular. Preoral snout short, 0.68 (0.49-0.76) times mouth width; broadly rounded in dorsoventral view, no indentations adjacent to anterior of orbits. Eyes small, spindle-shaped; dorsolateral on head, with lower edges well medial to lateral head margin in dorsal view; subocular ridges weak, not well defined, subequal in length to eye, almost a spiracle length from eye margin; no supraorbital crest; length 4.95 (4.80-6.00) in head length, 6.90 (4.42-6.14) times eye height.</paragraph>
|
||||
<paragraph pageNumber="6">External eye openings with or without weak anterior and posterior notches. Nictitating lower eyelids of rudimentary type. Spiracles small, variable, subcircular, cresentic to suboval, length 3.65 (3.17-5.05) in eye length, 0.17 (0.16-0.26) eye length behind and below posterior eye notch. Gill openings straight to slightly concave; slightly elevated on dorsolateral surface of head, upper margins slightly below lower edge of eye; anteriormost openings slightly larger than those posterior (heights variable, first four subequal or with anteriormost taller in paratypes); fifth opening much shorter than anterior four; fourth opening over origin of pectoral fin; height of fifth 0.57 (0.54-0.61) of third; height of third 14.32 (11.09-14.91) in head, 0.35 (0.35-0.51) of eye length. Gill filaments not visible from outside. Nostrils separated from each other and not reaching mouth; very small incurrent apertures; width 0.94 (1.02-1.29) in internarial space, 1.42 (1.40-1.66) in eye length, 0.49 (0.50-0.77) in third gill-slit height. Anterior nasal flaps large, forming a right angle at apex (sometimes with a weak lobe in paratypes); posterior margin nearly straight or weakly corrugated, innermost edge short of upper lip by more than a tooth length; covering excurrent apertures; posterior nasal flaps present but inconspicuous, directed internally, visible when upper flap is lifted (but exposed in paratype CSIRO H 5889-17); skirt-shaped fold around excurrent aperture (nasal curtain and fold overlapping).</paragraph>
|
||||
<paragraph pageNumber="6">Mouth moderately large, very broadly arched; upper jaw horizontal at symphysis, forming a sharp angle just lateral to insertion of nasal curtain, posterior half of jaw almost straight, directed posterolaterally; central part of upper jaw overhanging lower jaw so that upper teeth are exposed in ventral view, edge of lower jaw similar in shape to edge of upper jaw; width 2.64 (2.27-2.99) in head length; length 2.19 (1.98-2.84) in width. Tongue large, flat, rounded anteriorly, filling most of floor of mouth (from paratype MZB 15550). Labial furrows very short but usually conspicuous (labial furrows on left side inconspicuous in some paratypes), confined to jaw angle; upper furrows shorter than lower furrows, 0.58 (0.32-0.69) of lower furrows. Teeth in 56/ 53 rows; 5/4 series functional; usually with long central cusp flanked by either one or two short cusplets on each side; lateral teeth with progressively smaller cusps; no toothless spaces at symphysis; not strongly differentiated in upper and lower jaws; medials, anterolaterals and posteriors in both jaws weakly defined; central upper anterolateral teeth largest. Tooth formula (from paratype MZB 15550) is: 27-2-27 / 25-4-24</paragraph>
|
||||
<paragraph pageNumber="6">Sexual heterodonty present; upper anterolateral teeth in an adult male paratype (MZB 15550) with enlarged central cusp compared to those in the female paratype (CSIRO H 5858-01), teeth height ~1.3 vs. 0.6-0.7 mm.</paragraph>
|
||||
<paragraph pageNumber="7">Lateral trunk denticles below first dorsal fin small (0.5-0.6 mm long), dense, strongly imbricate. Crowns mostly tricuspid, with long pungent central cusp flanked by short, sharp lateral cusps; pair of strong, converging medial ridges extending entire length of crown; pair of low but prominent ridges on lateral cusps; crown length 1.5 to 2 times width, medial cusp about half or less total length of denticle.</paragraph>
|
||||
<paragraph pageNumber="7">First dorsal fin low, angled rearwards, not falcate; slightly convex anterior margin, apex broadly rounded; posterior margin almost straight (sometimes weakly convex), sloping slightly posteroventrally from apex (upright in some paratypes); free rear tip angular, inner margin nearly straight; insertion slightly anterior to fin apex; origin barely anterior to pelvic-fin insertions (further forward in female paratype CSIRO H 5858-01); insertion slightly posterior to pelvic-fin free rear tips (over free rear tips in female paratype), well anterior to anal-fin origin. First dorsal-fin base 1.87 (1.68-1.92) in interdorsal space, 2.68 (2.08-2.81) in dorsal caudalfin margin; height 1.59 (1.55-1.96) in base length; inner margin 2.14 (1.87-2.55) in height, 3.41 (3.09-4.51) in base length. Second dorsal fin low, subtriangular, generally lower and more elongate than first dorsal fin; height 0.88 (0.77-0.87) of first dorsal-fin height, base length 1.08 (0.87-1.13) of first dorsal-fin base length; anterior margin nearly straight or slightly convex, with bluntly angular to narrowly rounded apex; posterior margin sloping posteroventrally from apex, weakly concave; free rear tip angular, extended slightly; inner margin straight; origin slightly forward to insertion of anal-fin (over anal-fin insertion in smallest paratype); insertion well posterior to anal-fin free rear tip, about opposite second dorsal-fin apex. Second dorsal-fin base length 1.10 (0.91-1.12) in dorsal -caudal space; height 1.94 (1.82-2.34) base length; inner margin 2.00 (1.50- 2.13) in height, 3.89 (3.26-4.26) in base length.</paragraph>
|
||||
<paragraph pageNumber="7">Anal fin low, subtriangular, smaller in area to second dorsal fin; height 0.76 (0.67-0.80) in second dorsalfin height, base 1.27 (1.06-1.21) times second dorsal-fin base; base sometimes with a preanal ridge; anterior margin nearly straight or weakly convex, apex broadly rounded; posterior margin weakly concave, slanting strongly posterodorsally; free rear tip angular, extended slightly, inner margin nearly straight; origin about 1.26 (1.26-1.48) times base length behind pelvic-fin insertions; insertion well posterior to apex. Anal-fin base 0.98 (0.83-1.07) in anal-caudal space; height 3.25 (3.07-3.73) in base length; inner margin 1.43 (1.14-1.45) in height, 4.66 (3.77-4.46) in base length.</paragraph>
|
||||
<paragraph pageNumber="8">Pectoral fin small, somewhat lobate, apex almost forming a right angle; anterior margin weakly convex, 1.02 (0.97-1.07) times its length; base narrow; posterior margin almost straight; free rear tip broadly rounded, inner margin moderately to strongly convex. Pelvic fin weakly triangular; anterior margin slightly convex, 0.60 (0.58-0.64) of pectoral-fin anterior margin; apex broadly rounded; posterior margin slightly convex; free rear tip angular, slightly attenuated in adult males; inner margins straight, not fused together over claspers of adult males.</paragraph>
|
||||
<paragraph lastPageNumber="9" pageNumber="8">Claspers of the adult male holotype elongate, moderately slender, nearly straight and tapering on lateral edge, not undulated; extending well behind pelvic-fin free rear tips, but terminating well anterior to anal-fin origin; tip sharply pointed, not fleshy. Most of ventral and distal portion of dorsal clasper with small clasper denticles with anteriorly directed cusps; anterior dorsal surface of glans largely naked. Apopyle and hypopyle connected by long clasper groove, with its dorsal margins fused over clasper canal. Lateral fold well developed, originating along length of dorsal midline and enveloping part of inner dorsal half of clasper; terminating distally just posterior to origin of cover rhipidion. Cover rhipidion very short, narrow, formed as distally tapering wedge, mesial margin with fleshy, longitudinal fold, without mid-lateral tab; separated posteriorly from posterior end of external edge of clasper by a deep groove; partly posterior to exorhipidion, well ahead of the rear end of the rhipidion. Rhipidion present, large, extending along most of clasper glans; formed as a flat blade with posterior tip beside apex of exorhipidion. Pseudosiphon absent. Pseudopera indistinct. Exorhipidion elongate, tapering distally, not flap-like, originating near posterolateral edge of cover rhipidion; a series of about 20 small clasper hooks along outer rim of hypopyle, mostly in multiple rows; apex sharply pointed.</paragraph>
|
||||
<paragraph pageNumber="9">Caudal fin narrow and asymmetrical; terminal lobe large; ventral lobe very low; dorsal margin moderately long, nearly straight with weak lateral undulations, 3.96 (3.67-4.60) in precaudal length; preventral margin nearly straight, 1.68 (1.53-1.82) in dorsal caudal-fin margin; tip of ventral caudal-fin lobe broadly rounded; postventral margin weakly concave, not well differentiated into upper and lower parts. Subterminal notch a narrow, shallow slot; margin straight, its length 0.71 (0.82-1.16) in terminal region. Terminal margin slightly convex, sometimes notched, dorsal and ventral tips broadly rounded, its length 3.66 (3.39-4.62) times dorsal caudal-fin margin.</paragraph>
|
||||
<paragraph pageNumber="9">Egg cases medium-sized, 45-50 mm in length from anterior to posterior margins (excluding horns); smooth-walls without longitudinal striations or ridges; relatively flat, its height 18.1 (16.0-19.4)% of case length; posterior width 39.2 (38.3-40.2)% of case length; lateral flanges absent. Anterior border nearly straight, broad; anterior horns small and nearly vestigial; tendrils long, wiry and tightly coiled. Posterior border narrow, concave; posterior horns short, curved strongly inwards but not touching; tendrils very long, wiry, highly convoluted, tightly coiled. Egg cases, in alcohol, are uniformly golden yellow to brown in colour; similar, but slightly darker, when fresh.</paragraph>
|
||||
<paragraph pageNumber="9">Vertebral centra 132 (127-132 in paratypes), monospondylous 33 (33-34), precaudal 90 (81-88, mostly less than 83-85) and caudal 37 (38-41). Last few MP centra before MP-DP transition hardly enlarged, not forming ‘stutter zone’ of alternating long and short centra.</paragraph>
|
||||
<paragraph lastPageNumber="10" pageNumber="9">Colour. In preservative: Dorsal surface pale brown, with about 10 vague, widely separated, darker brown saddles and densely peppered with brownish black spots; margins of saddles not demarcated by darker transverse lines; saddles mostly subequal in breadth to their interspaces, extending to about lateral midline on tail and slightly more ventrally on trunk; interspaces densely spotted; spots variable in size, largest (slightly larger than spiracle) mostly widely spaced on side on abdomen, smallest (about quarter of spiracle diameter) densest along dorsal midline and on head (spots consistently much smaller than spiracles in some paratypes). Head with two weakly defined saddles, across orbito -spiracular region and between gills; anterior margin (origin) of first saddle through middle of interorbit, posterior margin (insertion) between spiracles, well-developed extensions of saddle directed antero-ventrally below eyes; second saddle not obviously recurved, origin through second gill slit, insertion well posterior to last gill slit; small dark spots densely scattered over snout (~10-13 spots per cm2, denser in some paratypes). Supra-abdominal region with two similar saddles; anterior saddle originating just posterior to pectoral-fin insertion, terminating well-behind level of its free rear tip; second saddle centered over pelvic-fin origin. Tail and caudal fin with 6 saddles; one saddle centered under each dorsal-fin base; slightly broader, central interdorsal saddle; narrower saddle centered on dorsal -caudal space; two broad saddles on caudal fin, anterior saddle above lower caudal lobe, narrower than posterior saddle at tail apex. Dorsal fins densely spotted, with evidence of rusty saddles anteriorly, paler posteriorly. Paired fins evenly spotted on dorsal surface, not paler distally; ventral surface mostly uniformly yellowish. Lower lobe of caudal fin spotted. Anal fin uniform yellowish. Ventral surface of body pale yellow or creamish, dusky on tail (holotype and most paratypes with dark, embedded, sand granules); mouth white (more yellowish is some paratypes).</paragraph>
|
||||
<paragraph pageNumber="10">Size. Attains a maximum size of about 530 mm TL; males mature at ~400 mm TL; five females between 476 and 528 mm TL were pregnant, with between three and six egg cases in each of the two uteri.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="distribution">
|
||||
<paragraph pageNumber="10">Distribution. Known from the southern coast of Java, and off the islands of Bali and Lombok in eastern Indonesia, and has also been recorded from several landing sites in the Philippines (see Fig. 9).</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="biology_ecology">
|
||||
<paragraph pageNumber="10">Habitat. Caught occasionally by demersal fisheries, but no depth distribution information available.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="etymology">
|
||||
<paragraph pageNumber="10">Etymology. Derived from the Latin maculosus (speckled) in allusion to the speckling of small dark spots over the dorsal and lateral surfaces of the body.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph pageNumber="10">
|
||||
Comparisons.
|
||||
<taxonomicName class="Elasmobranchii" family="Scyliorhinidae" genus="Halaelurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="10" phylum="Chordata" rank="species" species="maculosus">Halaelurus maculosus</taxonomicName>
|
||||
was first recognised from the Philippines as being similar but possible distinct from
|
||||
<taxonomicName class="Elasmobranchii" family="Scyliorhinidae" genus="Halaelurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="10" phylum="Chordata" rank="species" species="buergeri">H. buergeri</taxonomicName>
|
||||
(Compagno et al., 2005). However, it differs from
|
||||
<taxonomicName class="Elasmobranchii" family="Scyliorhinidae" genus="Halaelurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="10" phylum="Chordata" rank="species" species="buergeri">H. buergeri</taxonomicName>
|
||||
in having a colour pattern of numerous, small to moderately large dark spots (usually much smaller than spiracle and covering the entire dorsal and lateral surfaces of the body), rather than fewer, larger dark spots (usually larger than spiracle and arranged along the saddle-marking borders but which are absent from the paler areas between saddles) (see Figs 1 and 11).
|
||||
<taxonomicName class="Elasmobranchii" family="Scyliorhinidae" genus="Halaelurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="10" phylum="Chordata" rank="species" species="maculosus">Halaelurus maculosus</taxonomicName>
|
||||
also differs from
|
||||
<taxonomicName class="Elasmobranchii" family="Scyliorhinidae" genus="Halaelurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="10" phylum="Chordata" rank="species" species="buergeri">H. buergeri</taxonomicName>
|
||||
in a number of morphological characters: more posteriorly positioned first dorsal fin (predorsal length 41.6-42.8 vs. 39.3-41.0% TL), dorsal fins closer together (interdorsal space 13.1-15.0 vs. 16.0-17.1% TL), slightly longer head (head length 17.8- 19.4 vs. 16.5-17.8% TL), shorter caudal peduncle (dorsal-caudal space 7.7-8.9 vs. 9.2-9.9% TL, anal-caudal space 9.0-10.9 vs. 11.3-12.8% TL), longer pectoral fin (anterior margin 10.8-11.7 vs. 9.6-10.6% TL) and longer second dorsal (length 9.7-11.3 vs. 8.8-9.5% TL) and anal (length 11.0-12.2 vs. 10.3-10.5% TL) fins.
|
||||
</paragraph>
|
||||
<paragraph pageNumber="10">
|
||||
The length of the labial furrows has been previously used to separate Indo-West Pacific species
|
||||
<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:7B93E5DF-87DD-483D-8281-E80D93969483" class="Elasmobranchii" family="Scyliorhinidae" genus="Halaelurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Halaelurus boesemani Springer & D'Aubrey 1972:11" order="Carcharhiniformes" pageNumber="10" phylum="Chordata" rank="species" species="boesemani">H. boesemani</taxonomicName>
|
||||
(long furrows, up to 5 mm in length) and
|
||||
<taxonomicName class="Elasmobranchii" family="Scyliorhinidae" genus="Halaelurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="10" phylum="Chordata" rank="species" species="buergeri">H. buergeri</taxonomicName>
|
||||
(furrows absent or very short, <2 mm) (see e.g. Springer, 1979; Compagno 1984; Figs 10b and 11b).
|
||||
<taxonomicName class="Elasmobranchii" family="Scyliorhinidae" genus="Halaelurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="10" phylum="Chordata" rank="species" species="maculosus">Halaelurus maculosus</taxonomicName>
|
||||
possess very short (<1.8 mm) labial furrows, similar to
|
||||
<taxonomicName class="Elasmobranchii" family="Scyliorhinidae" genus="Halaelurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="10" phylum="Chordata" rank="species" species="buergeri">H. buergeri</taxonomicName>
|
||||
. Springer and D’Aubrey (1972) noted that the degree of development of the labial furrows in
|
||||
<taxonomicName class="Elasmobranchii" family="Scyliorhinidae" genus="Halaelurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="10" phylum="Chordata" rank="species" species="buergeri">H. buergeri</taxonomicName>
|
||||
is variable, with some specimens lacking noticeable furrows and one specimen with furrows present on only one of the mouth corners. The labial furrows of
|
||||
<taxonomicName class="Elasmobranchii" family="Scyliorhinidae" genus="Halaelurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="10" phylum="Chordata" rank="species" species="maculosus">H. maculosus</taxonomicName>
|
||||
are also variably developed in some paratypes and are barely visible in the holotype (Fig. 2).
|
||||
</paragraph>
|
||||
<paragraph pageNumber="10">
|
||||
The colour pattern of
|
||||
<taxonomicName class="Elasmobranchii" family="Scyliorhinidae" genus="Halaelurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="10" phylum="Chordata" rank="species" species="maculosus">H. maculosus</taxonomicName>
|
||||
is very similar to that of
|
||||
<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:7B93E5DF-87DD-483D-8281-E80D93969483" class="Elasmobranchii" family="Scyliorhinidae" genus="Halaelurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Halaelurus boesemani Springer & D'Aubrey 1972:11" order="Carcharhiniformes" pageNumber="10" phylum="Chordata" rank="species" species="boesemani">H. boesemani</taxonomicName>
|
||||
, but differs in having a denser coverage of variable-sized dark spots, with those over the darker saddle markings often much larger (Figs 1 and 10).
|
||||
<taxonomicName class="Elasmobranchii" family="Scyliorhinidae" genus="Halaelurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="10" phylum="Chordata" rank="species" species="maculosus">Halaelurus maculosus</taxonomicName>
|
||||
also differs from
|
||||
<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:7B93E5DF-87DD-483D-8281-E80D93969483" class="Elasmobranchii" family="Scyliorhinidae" genus="Halaelurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Halaelurus boesemani Springer & D'Aubrey 1972:11" order="Carcharhiniformes" pageNumber="10" phylum="Chordata" rank="species" species="boesemani">H. boesemani</taxonomicName>
|
||||
in a number of morphometric values and ratios: mouth length 2.8-3.6 vs. 2.2% TL, 2.0-2.8 vs. 3.7 times mouth width; lower labial furrow length 0.4- 0.9 vs. 1.6% TL, 4.0-9.6 vs. 2.0 times in eye length; interdorsal space 13.1-15.0 vs. 17.1% TL; anterior nasal flap length 1.3-1.7 vs. 2.0% TL; head width 11.1-12.2 vs. 13.6% TL,>28 vs. 7.4 times lower labial furrow length; caudal peduncle height 0.9-1.2 vs. 1.6 times its width; clasper outer length 3.6-5.0 vs. 5.7 times clasper base; claspers extending past pelvic-fin free tips by 5.8-7.1 vs. 0.9 times eye length. They also differ in the total number of vertebrae, 127-136 (usually 130-132) vs. 135-139 (usually 136-138).
|
||||
</paragraph>
|
||||
<paragraph pageNumber="10">
|
||||
<taxonomicName class="Elasmobranchii" family="Scyliorhinidae" genus="Halaelurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="10" phylum="Chordata" rank="species" species="maculosus">Halaelurus maculosus</taxonomicName>
|
||||
differs from two southern African endemics,
|
||||
<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:37E93C92-A664-4763-B9FD-2C06F52224D8" class="Elasmobranchii" family="Scyliorhinidae" genus="Halaelurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Halaelurus lineatus Bass, D'Aubrey & Kistnasamy 1975:12" order="Carcharhiniformes" pageNumber="10" phylum="Chordata" rank="species" species="lineatus">H. lineatus</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Elasmobranchii" family="Scyliorhinidae" genus="Halaelurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="10" phylum="Chordata" rank="species" species="natalensis">H. natalensis</taxonomicName>
|
||||
, in having a rounded snout tip which is not upturned, compared to a knob-like, upturned snout tip in the latter two species. It differs from
|
||||
<taxonomicName class="Elasmobranchii" family="Scyliorhinidae" genus="Halaelurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Carcharhiniformes" pageNumber="10" phylum="Chordata" rank="species" species="quagga">Halaelurus quagga</taxonomicName>
|
||||
from the northwestern Indian Ocean, in having a more heavily spotted colour pattern (rather than with>20 dark vertical bars without dark spots).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
58
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<taxonomicName id="75F515395F023D8040CE404CE056281B" ID-CoL="4JRGH" authority="Walker, 1835" authorityName="Walker" authorityYear="1835" class="Insecta" family="Platygastridae" genus="Platygaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platygaster (Platygaster) pleuron" order="Hymenoptera" pageId="0" pageNumber="7991" phylum="Arthropoda" rank="species" species="pleuron" subGenus="Platygaster">Platygaster (Platygaster) pleuron Walker, 1835</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="4AC7B76154633DEC0117927FEFC58276" pageId="0" pageNumber="7991" type="distribution">
|
||||
<paragraph id="36989F8185C87A8EB6ED24974A53A2A6" pageId="0" pageNumber="7991">Distribution</paragraph>
|
||||
<paragraph id="2BE86D08F675B3579A2F3D271B80C4AB" pageId="0" pageNumber="7991">England</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
82
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Normal file
82
data/17/C8/6D/17C86D3C356B499CED9222174C0A1AD4.xml
Normal file
|
|
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<taxonomicName id="E875113AEDF147407DB7C984839EDE5B" ID-CoL="6RLPM" ID-ENA="2495090" authority="Jurine, 1807" authorityName="Jurine" authorityYear="1807" class="Insecta" family="Crabronidae" genus="Miscophus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Miscophus bicolor" order="Hymenoptera" pageId="0" pageNumber="8050" phylum="Arthropoda" rank="species" species="bicolor">Miscophus bicolor Jurine, 1807</taxonomicName>
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|
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|
||||
<taxonomicName id="EE4433AE741E68B104AE80F997E9EDB5" genus="Apoidea" lsidName="Apoidea dubius" pageId="0" pageNumber="8050" rank="species" species="dubius">dubius</taxonomicName>
|
||||
(Panzer, 1809,
|
||||
<taxonomicName id="288C57D442637BB85F3061F1F7D8B915" class="Insecta" family="Crabronidae" genus="Larra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Larra" order="Hymenoptera" pageId="0" pageNumber="8050" phylum="Arthropoda" rank="genus">Larra</taxonomicName>
|
||||
)
|
||||
</paragraph>
|
||||
<paragraph id="8A9E4C67674075C3B905112EC1875822" pageId="0" pageNumber="8050">
|
||||
<taxonomicName id="17F4C7FC6C6EBD8787683BB6706799E2" genus="Apoidea" lsidName="Apoidea metallicus" pageId="0" pageNumber="8050" rank="species" species="metallicus">metallicus</taxonomicName>
|
||||
<normalizedToken id="FE03740F61D6365BA9C66855BB55E5B9" originalValue="Verhöff">Verhoeff</normalizedToken>
|
||||
, 1890
|
||||
</paragraph>
|
||||
<paragraph id="40BF2B9F1AF250F3F343514F5D4F4919" pageId="0" pageNumber="8050">
|
||||
<taxonomicName id="72E71E823E08DE8E471ADFE74C59A303" genus="Apoidea" lsidName="Apoidea tsunekii" pageId="0" pageNumber="8050" rank="species" species="tsunekii">tsunekii</taxonomicName>
|
||||
de Andrade, 1960
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="86595E6CAF6BF5548A55DF76F8EB6C68" pageId="0" pageNumber="8050" type="distribution">
|
||||
<paragraph id="1C009E57CCFB08E2D32B9B3623A5D274" pageId="0" pageNumber="8050">Distribution</paragraph>
|
||||
<paragraph id="8F8FFCA9DE14FA8A0AE0B711825C047F" pageId="0" pageNumber="8050">England</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="E019E5D634237779D7D4A134B400BBA3" pageId="0" pageNumber="8050" type="notes">
|
||||
<paragraph id="BFA71B48C9C988C110C5978C200E5843" pageId="0" pageNumber="8050">Notes</paragraph>
|
||||
<paragraph id="307060014189A14F22F533A61B4A4171" pageId="0" pageNumber="8050">
|
||||
added by
|
||||
<bibRefCitation id="2035C9FF20D058E996FFEFD3F1630DC2" author="Knowles, A. P." journalOrPublisher="British Journal of Entomology and Natural History" pageId="0" pageNumber="8050" pagination="229 - 232" title="Miscophusbicolor Jurine (Hymenoptera: Crabronidae), a wasp new to Britain" volume="18" year="2006">Knowles and Else (2006)</bibRefCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
206
data/17/C8/78/17C878AA46299D90FEFD242DF1CC6B59.xml
Normal file
206
data/17/C8/78/17C878AA46299D90FEFD242DF1CC6B59.xml
Normal file
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@ -0,0 +1,206 @@
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<mods:namePart>Qiao, Penghai</mods:namePart>
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|
||||
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||||
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||||
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||||
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||||
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||||
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|
||||
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||||
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||||
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||||
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||||
<mods:identifier type="ZooBank">571F329FDB87411EAC761822D44B1E94</mods:identifier>
|
||||
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|
||||
<treatment ID-GBIF-Taxon="141125908" LSID="urn:lsid:zoobank.org:act:9D93BD0E-90DE-4516-8C8C-BFF5BA0530CB" httpUri="http://treatment.plazi.org/id/17C878AA46299D90FEFD242DF1CC6B59" lastPageId="6" lastPageNumber="71" pageId="3" pageNumber="68">
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|
||||
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|
||||
<taxonomicName LSID="http://zoobank.org/9D93BD0E-90DE-4516-8C8C-BFF5BA0530CB" class="Chilopoda" family="Lithobiidae" genus="Hessebius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hessebius luquensis" order="Lithobiomorpha" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="luquensis">Hessebius luquensis</taxonomicName>
|
||||
<taxonomicNameLabel pageId="3" pageNumber="68">sp. n.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="3" pageNumber="68" type="type data">
|
||||
<paragraph pageId="3" pageNumber="68">Type data.</paragraph>
|
||||
<paragraph pageId="3" pageNumber="68">
|
||||
Holotype: female numbered LQ 8 (Fig. 2
|
||||
<normalizedToken originalValue="A–F">A-F</normalizedToken>
|
||||
), body length 10.4 mm, from Luqu County, the Gannan Prefecture, Gansu province, China,
|
||||
<geoCoordinate direction="north" orientation="latitude" precision="1" value="34.75647">34.75647°N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate direction="east" orientation="longitude" precision="1" value="102.57245">102.57245°E</geoCoordinate>
|
||||
, 13 May 2012, 3192 meters above sea level, leg. Gonghua Lin, Weiping Li. Paratypes: 8 females, 2 males, same data as holotype.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="3" pageNumber="68" type="habitat">
|
||||
<paragraph pageId="3" pageNumber="68">Habitat.</paragraph>
|
||||
<paragraph pageId="3" pageNumber="68">Speciemens were collected under stones along roadside on steppes from Luqu.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="3" pageNumber="68" type="etymology">
|
||||
<paragraph pageId="3" pageNumber="68">Etymology.</paragraph>
|
||||
<paragraph pageId="3" pageNumber="68">The name is derived from the locality Luqu where the species is discovered. Luqu country is situated in the eastern edge of the Tibetan Plateau standing on the junction of Gansu, Qinghai and Sichuan Provinces.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="4" pageNumber="69" type="diagnosis">
|
||||
<paragraph pageId="4" pageNumber="69">
|
||||
<pageBreakToken pageId="4" pageNumber="69" start="start">Diagnosis</pageBreakToken>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph pageId="4" pageNumber="69">
|
||||
Body length 8.5-12.3 mm; head slightly widened; antennae of 20 antennomeres; 7-10 ocelli arranged in three rows;
|
||||
<normalizedToken originalValue="Tömösváry’s">Toemoesvary's</normalizedToken>
|
||||
organ oval, almost equal in size to neighboring ocelli (Fig. 2B); lateral margins of forcipular coxosternite slanting; anterior margin with 2 + 2 sharp teeth and with setiform porodonts; tergites without triangular posterolateral process, a line of setae along posterior border of TT 8 and 10; legs 14 and 15 thicker than anterior ones in both sexes; a dorsal furrow on the tibia of legs 14-15 on male; coxal pores 3-6, round, arranged in one row; female gonopods with two moderately long, bullet-shaped spurs, the second article of the female gonopods having a massive process; terminal claw of the third article simple, with a small triangular protuberance on basal ventral side; male gonopods short and small.
|
||||
</paragraph>
|
||||
<caption pageId="4" pageNumber="69">
|
||||
<paragraph pageId="4" pageNumber="69">
|
||||
Figure 2.
|
||||
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Hessebius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hessebius luquensis" order="Lithobiomorpha" pageId="4" pageNumber="69" phylum="Arthropoda" rank="species" species="luquensis">Hessebius luquensis</taxonomicName>
|
||||
sp. n., holotype, female: A dorsal view, scale 1 mm B left ocelli and
|
||||
<normalizedToken originalValue="Tömösváry’s">Toemoesvary's</normalizedToken>
|
||||
organ (To), scale 250
|
||||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||||
; C, forcipular coxosternite, ventral view, scale bar 1 mm D posterior segments and gonopods, internal view, scale bar 500
|
||||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||||
E right gonopod, dorsal view, scale bar 250
|
||||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||||
F, right gonopod, ventral lateral view, scale bar 250
|
||||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||||
. Paratype, LQ 9, male G posterior segments and gonopods, internal view, scale bar 500
|
||||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||||
.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection lastPageId="6" lastPageNumber="71" pageId="4" pageNumber="69" type="description">
|
||||
<paragraph pageId="4" pageNumber="69">Description.</paragraph>
|
||||
<paragraph pageId="4" pageNumber="69">Holotype (female), body 10.4 mm long, cephalic plate 1.3 mm width, 1.2 mm length.</paragraph>
|
||||
<paragraph pageId="4" pageNumber="69">Colour (based on specimens in 95 % ethanol): antennomeres yellow; tergites pale yellow, with brown margin; cephalic plate and T 1 brown; pleural region and sternites pale yellow; distal part of forcipules dark brown, maxillipede coxosternum and S 15 yellow; legs pale yellow with gray hue, pretarsal claw dark brown.</paragraph>
|
||||
<paragraph pageId="4" pageNumber="69">Cephalic plate smooth, convex, slightly longer than wide; short to long setae scattered along the marginal ridge of the cephalic plate; setae on head shield symmetrically arranged, three pairs between antennocellar and transverse suture, two pairs behind these sutures; frontal marginal ridge with clear transverse suture; projection of lateral marginal conspicuously discontinuous; posterior marginal ridge slightly concave without median thickening.</paragraph>
|
||||
<paragraph pageId="4" pageNumber="69">
|
||||
Ocelli (Fig. 2B): translucent with dark pigment, 1 + 4, 3, 2 ocelli on each cephalic plateau, arranged in three rows. The posterior ocellus is the biggest, seriate ocelli smaller.
|
||||
<normalizedToken originalValue="Tömösváry’s">Toemoesvary's</normalizedToken>
|
||||
organ oval, nearly the same size as the adjacent ocelli, not remote, situated ventraly on anterolateral margin of cephalic pleurite.
|
||||
</paragraph>
|
||||
<paragraph pageId="4" pageNumber="69">Antennae length 3.46 mm, extending back to anterior margin of T 5, consisting of 20 elongate antennomeres covered with dense pale setae. The basal two articles enlarged, then elongate and tapering. The first article wider than long, the second article has the equal width to length, terminal article approximately 2.5 times length to width. Setation: the first two articles has fewer setae than succeeding articles especially on anterior side, then increasing, till 6 or 7, the density become constant.</paragraph>
|
||||
<paragraph pageId="4" pageNumber="69">Forcipular coxosternites subtrapezoidal, coxosternite with narrow, straight or slightly recurved dental margin; 2+2 teeth on dental margin, small, blunt knobs with independent sclerotization from coxosternite; porodont setiform towards its apex, much stouter than a seta at its base; no shoulders lateral to the porodont; 3 to 4 long setae along the slope, some setae scattered on ventral side of coxosternite.</paragraph>
|
||||
<paragraph lastPageId="5" lastPageNumber="70" pageId="4" pageNumber="69">
|
||||
Tergites smooth, T1 narrower than head and T3, subrectangular; on TT 8 and 10 there is a line of setae along posterior borders; posterior margin of TT. 1, 3, 5, 8, 10, 12, 14 a little concave, of TT. 6, 7 straight; posterior angles of all tergites rounded without triangular projections; marginal ridge narrow, entire on TT 1, 3 and 5, interrupted posteriorly on TT 7, 8, 10, 12, and 14 (Fig. 2A); tiny setae inserted in pores
|
||||
<pageBreakToken pageId="5" pageNumber="70" start="start">scattered</pageBreakToken>
|
||||
very sparsely over the surface, more setae on anterior and posterior angles of each tergite.
|
||||
</paragraph>
|
||||
<paragraph pageId="5" pageNumber="70">
|
||||
Sternites trapeziform, setae scattered very sparsely on the surface; four pairs of long bristles approximately symmetrical on the anterior corner and margin, one pair on the posterior corner; among long bristles
|
||||
<normalizedToken originalValue="there’s">there's</normalizedToken>
|
||||
small pairs of short bristles; short to long setae along posterior border, in some individuals SS 13, 14, 15 more dense.
|
||||
</paragraph>
|
||||
<paragraph pageId="5" pageNumber="70">Legs: tarsal articulations only visible with shallow ventral suture on 1st to 11th, distinct on 12th and 13th, well-defined on legs 14 and 15; leg pairs 14 and 15 thicker and longer with sparse setae in contrast to legs 1-13; pretarsus of legs 1-13 with a slightly curved, long, principal claw and smaller and thinner anterior and posterior accessory spines, anterior accessory spines long and slender, 0.33-0.5 the length of principal claw, posterior one stouter, 0.25 the length of principal claw, forming slightly larger angles with tarsal claws; accessory apical spines on the 14th vestigial, absent on the 15th; abundant glandular pores on surfaces of femur, tibia and tarsus of legs 14 and 15; short to moderately long setae scattered over the surface of legs 1-13, latter half of tarsi generally more setose with two rows of setae along ventral side, fewer setae on legs 14-15.</paragraph>
|
||||
<paragraph pageId="5" pageNumber="70">14th and 15th legs: swollen, 15th leg 30% of body-length, tarsus 1 4.3 times longer than wide, tarsus 2 48% length of tarsus on leg 15. Data on the leg plectrotaxy are compiled in Table 1. In the male the 14th and 15th tibia has a dorsal sulcus extending along its whole length.</paragraph>
|
||||
<caption ID-Table-UUID="6C9823FE98BE7B3937ED70178290A581" httpUri="http://table.plazi.org/id/6C9823FE98BE7B3937ED70178290A581" pageId="5" pageNumber="70">
|
||||
<paragraph pageId="5" pageNumber="70">
|
||||
Table 1.
|
||||
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Hessebius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hessebius luquensis" order="Lithobiomorpha" pageId="5" pageNumber="70" phylum="Arthropoda" rank="species" species="luquensis">Hessebius luquensis</taxonomicName>
|
||||
sp. n. leg plectrotaxy; letters in brackets indicate variable spines.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph pageId="5" pageNumber="70">
|
||||
<table pageId="5" pageNumber="70">
|
||||
<tr pageId="5" pageNumber="70">
|
||||
<th colspan="1" pageId="5" pageNumber="70" rowspan="2">Legs</th>
|
||||
<th colspan="5" pageId="5" pageNumber="70" rowspan="1">Ventral</th>
|
||||
<th colspan="5" pageId="5" pageNumber="70" rowspan="1">Dorsal</th>
|
||||
</tr>
|
||||
<tr pageId="5" pageNumber="70">
|
||||
<th colspan="1" pageId="5" pageNumber="70" rowspan="1">C</th>
|
||||
<th colspan="1" pageId="5" pageNumber="70" rowspan="1">
|
||||
<abbrev id="ABBRID0EAGAE" pageId="5" pageNumber="70">Tr</abbrev>
|
||||
</th>
|
||||
<th colspan="1" pageId="5" pageNumber="70" rowspan="1">P</th>
|
||||
<th colspan="1" pageId="5" pageNumber="70" rowspan="1">F</th>
|
||||
<th colspan="1" pageId="5" pageNumber="70" rowspan="1">
|
||||
<abbrev id="ABBRID0EOGAE" pageId="5" pageNumber="70">Ti</abbrev>
|
||||
</th>
|
||||
<th colspan="1" pageId="5" pageNumber="70" rowspan="1">C</th>
|
||||
<th colspan="1" pageId="5" pageNumber="70" rowspan="1">
|
||||
<abbrev id="ABBRID0EZGAE" pageId="5" pageNumber="70">Tr</abbrev>
|
||||
</th>
|
||||
<th colspan="1" pageId="5" pageNumber="70" rowspan="1">P</th>
|
||||
<th colspan="1" pageId="5" pageNumber="70" rowspan="1">F</th>
|
||||
<th colspan="1" pageId="5" pageNumber="70" rowspan="1">
|
||||
<abbrev id="ABBRID0EHHAE" pageId="5" pageNumber="70">Ti</abbrev>
|
||||
</th>
|
||||
</tr>
|
||||
</table>
|
||||
</paragraph>
|
||||
<paragraph pageId="5" pageNumber="70">Coxal pores on legs 12-15, circular; inner pores smaller. Distance between pores 2-3 times bigger than diameter of pore; formula 4, 4, 4, 4. Coxal pores set in a shallow groove arranged in a row with short to long setae scattered over the surface of apophysis.</paragraph>
|
||||
<paragraph pageId="5" pageNumber="70">Female S15 generally trapeziform, straight posteromedially; sternite of genital segment well sclerotised, wider than long; sternite of genital segment with posterior margin moderately concave between condyles of gonopods, except for a small, median approximately circular bulge, distal lightly sclerotised; short to long setae scattered over the surface of genital segment and lateral margins.</paragraph>
|
||||
<paragraph lastPageId="6" lastPageNumber="71" pageId="5" pageNumber="70">
|
||||
Female gonopods divided into three articles, the first article moderately broad, bearing 11-17 short to moderately long setae, arranged in three rows; the first article
|
||||
<pageBreakToken pageId="6" pageNumber="71" start="start">also</pageBreakToken>
|
||||
bearing 2+2 moderately long, bullet-shaped spurs, inner spur slightly smaller and more anterior than the outer (Fig. 2D); the second article with 6 setae arranged in one rows(Fig. 2F); dorsolateral ridge of second article with a massive expansion projecting distally over the base of the third article (Fig. 2F), six short blunt spines along the dorsolateral ridge, one on the ventral side of dorsodistal projection (Fig. 2F); three moderately long setae on third article; dorsolateral setae one on the first article, eight on the second article four of which short and blunt and four on the third (Fig. 2E); one long dorsomedial setae on each article (Fig. 2E); terminal claw simple, slender and sharp, having small triangular protuberance on ventral side (Fig. 2D).
|
||||
</paragraph>
|
||||
<paragraph pageId="6" pageNumber="71">Male S15: subsemicircular, well chitinized, long setae scattered sparsely over its surface and posterior margins. Male genitalia: first genital sternite wider than long, well chitinized; posterior margin quite deeply concave between the gonopods, no bulge medially; 24 short to medium setae scattered sparsely over its surface and at lateral margins, second genital sternite with abundance seta; gonopod of a single small article with 2 seta on its surface, apically slightly chitinized, flat (Fig. 2H).</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="6" pageNumber="71" type="variation">
|
||||
<paragraph pageId="6" pageNumber="71">Variations.</paragraph>
|
||||
<paragraph pageId="6" pageNumber="71">The length of the body (from anterior to posterior) range from 8.5 mm (LQ 9) to (LQ 12) 12.3 mm. Colour of body from pale yellow to yellow brown to ferruginous. Ocelli 1 + 4, 3, 2 or 1 + 4, 2, 1 or 1 + 3, 2, 1 on the cephalic plateau. Coxal pores 4444, 4443, 5466, 6466, 5555, 4555 in female; 4444 in male. 15th legs of LQ 9 (♂): length of each of the three distal articles of the 15th legs in comparison with their own diameter, 15th tibiae: 0.76 mm/0.31 mm = 2.45x; 15th tarsus 1: 0.69 mm/0.22 mm =3.14x; 15th tarsus 2: 0.62-0.13 mm/0.06 mm = 4.77x.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="6" pageNumber="71" type="remarks">
|
||||
<paragraph pageId="6" pageNumber="71">Remarks.</paragraph>
|
||||
<paragraph pageId="6" pageNumber="71">
|
||||
The female of
|
||||
<taxonomicName lsidName="H. luquensis" pageId="6" pageNumber="71" rank="species" species="luquensis">H. luquensis</taxonomicName>
|
||||
sp. n. is mostly similar to
|
||||
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Hessebius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hessebius longispinipes" order="Lithobiomorpha" pageId="6" pageNumber="71" phylum="Arthropoda" rank="species" species="longispinipes">Hessebius longispinipes</taxonomicName>
|
||||
Ma, Pei and Zhu, 2009, but can be readily distinguished by the following characters: more antennomeres (20 + 20, vs. 18 + 18 in
|
||||
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Hessebius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hessebius longispinipes" order="Lithobiomorpha" pageId="6" pageNumber="71" phylum="Arthropoda" rank="species" species="longispinipes">Hessebius longispinipes</taxonomicName>
|
||||
), more ocelli in three rows, a bulge exists near the base of the porodont; 14th accessory spines present, apical claw of female gonopods with triangular protuberance only on the ventral side and the apex of the male gonopod flat versus hemisphere in
|
||||
<taxonomicName lsidName="H. longispinipes" pageId="6" pageNumber="71" rank="species" species="longispinipes">H. longispinipes</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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