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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1198.119765" ID-Pensoft-Pub="1313-2970-1198-135" ID-Pensoft-UUID="74CF9BE973865BDDAC730C6A8DFEC07F" ID-ZooBank="4A6656D0E2FD4955A7BECC1F4128AE14" ModsDocID="1313-2970-1198-135" checkinTime="1713961751984" checkinUser="pensoft" docAuthor="Webb, Michael D. &amp; Xu, Ye" docDate="2024" docId="17410AFDB79D50AE897405195CD017A8" docLanguage="en" docName="ZooKeys 1198: 135-141" docOrigin="ZooKeys 1198" docPubDate="2024-04-23" docSource="http://dx.doi.org/10.3897/zookeys.1198.119765" docTitle="Smyga Dworakowska 1995" docType="treatment" docVersion="1" id="74CF9BE973865BDDAC730C6A8DFEC07F" lastPageNumber="135" masterDocId="74CF9BE973865BDDAC730C6A8DFEC07F" masterDocTitle="Revision of the leafhopper genus Smyga (Hemiptera, Cicadellidae, Typhlocybinae)" masterLastPageNumber="141" masterPageNumber="135" pageNumber="135" updateTime="1713961751984" updateUser="pensoft">
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<mods:title id="3760FAA38D58A2FDA5F793EC73BEF067">Revision of the leafhopper genus Smyga (Hemiptera, Cicadellidae, Typhlocybinae)</mods:title>
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<mods:namePart id="B81C263BBA3ABFE1C065EA469F7CA899">Webb, Michael D.</mods:namePart>
<mods:nameIdentifier id="D75EF72B3A77AEC93A7FA1FC36FEBB79" type="ORCID">https://orcid.org/0000-0002-1312-6142</mods:nameIdentifier>
<mods:affiliation id="C36A2C991D573E3747E0DEC9A522E68B">Department of Life Sciences (Insects), The Natural History Museum, London, SW 7 5 BD, UK</mods:affiliation>
<mods:nameIdentifier id="BE54D50163F9F08396B5A306AD7E063B" type="email">m.webb@nhm.ac.uk</mods:nameIdentifier>
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<mods:namePart id="9C2A692DAFB2E176BB71B959F5F5E161">Xu, Ye</mods:namePart>
<mods:nameIdentifier id="0A40534CFCA9915FF36D5392CBD55270" type="ORCID">https://orcid.org/0000-0003-3230-7923</mods:nameIdentifier>
<mods:affiliation id="34B83A95532415180D779652DC6D57A7">School of Agricultural Science, Jiangxi Agricultural University, Nanchang 330045, China</mods:affiliation>
<mods:nameIdentifier id="A16BAE2267F5463899763AAD69D1B688" type="email">yexu@jxau.edu.cn</mods:nameIdentifier>
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<mods:date id="9F5670D965C4FE4D44B1A3DD7033521F">2024</mods:date>
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<treatment id="17410AFDB79D50AE897405195CD017A8" LSID="urn:lsid:plazi:treatment:17410AFDB79D50AE897405195CD017A8" httpUri="http://treatment.plazi.org/id/17410AFDB79D50AE897405195CD017A8" lastPageNumber="135" pageId="0" pageNumber="135">
<subSubSection id="52B1A20EBC77931B68BA3ECF6CDC5875" pageId="0" pageNumber="135" type="nomenclature">
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<taxonomicName id="C2E253CACBA6EF529FA2C814D8C725AD" ID-CoL="8HR68" LSID="17410AFD-B79D-50AE-8974-05195CD017A8" authority="Dworakowska, 1995" authorityName="Dworakowska" authorityYear="1995" class="Insecta" family="Cicadellidae" genus="Smyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smyga" order="Hemiptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">Smyga Dworakowska, 1995</taxonomicName>
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<subSubSection id="F8DFB6FDC9182F157AD9C07C7BDF201F" pageId="0" pageNumber="135" type="reference_group">
<paragraph id="2BFE4C5C7EB2DEA27BA2305EBA320F3D" pageId="0" pageNumber="135">
<taxonomicName id="2D969332EAF2ACFC3A9BCAD1634C2903" authorityName="Dworakowska" authorityYear="1995" class="Insecta" family="Cicadellidae" genus="Smyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smyga" order="Hemiptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">Smyga</taxonomicName>
Dworakowska, 1995: 151.
</paragraph>
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<subSubSection id="AC58C2AB6F42CB74F1AA6D7FC0049F0A" pageId="0" pageNumber="135" type="type species">
<paragraph id="8DD8E73DB7CB7E3057870B01006D29A5" pageId="0" pageNumber="135">Type species.</paragraph>
<paragraph id="304767CCF4EE0D9CCC68F7485E7D5703" pageId="0" pageNumber="135">
<taxonomicName id="6589A8C54A9260659FAEF45280FA935E" authorityName="Webb &amp; Xu" authorityYear="2024" class="Insecta" family="Cicadellidae" genus="Smyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smyga distincta" order="Hemiptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="distincta">
<emphasis id="5C9FD702C83964A40D3F65BC1E6B3E4C" italics="true" pageId="0" pageNumber="135">Smyga distincta</emphasis>
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Dworakowska, 1995 by original designation.
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<subSubSection id="42B7BF69D67D7D7C753F3A5736F3E7AA" pageId="0" pageNumber="135" type="description">
<paragraph id="B6F244D8300008143B6F7DD7B6183A10" pageId="0" pageNumber="135">Description.</paragraph>
<paragraph id="6BEA2D63D15105F12A185F21308D22EB" pageId="0" pageNumber="135">
Body relatively robust. Pale yellow; head with a brown patch anteriorly (Figs
<figureCitation id="4F76A3F9E56F4783C33F750011B68356" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">1</figureCitation>
,
<figureCitation id="F85A2E2DA529D1FDFC1893F86DF26886" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">9</figureCitation>
); face with anteclypeus brown distally (Fig.
<figureCitation id="12906DC4D93EB12D3CD3EDD09887C46F" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">2</figureCitation>
), sometimes with more extensive brown marking (
<taxonomicName id="9C15566525CA6232EC6413E68AE42C9A" lsidName="S. divergens" pageId="0" pageNumber="135" rank="species" species="divergens">
<emphasis id="B339094D86557F2612090DBC44C2AB72" italics="true" pageId="0" pageNumber="135">S. divergens</emphasis>
</taxonomicName>
); pronotum with a brown transverse narrow band at midlength, pale yellow anterior to band and silvery posterior to band (Figs
<figureCitation id="C139CD485E119B74BDE2347B3C58FD63" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">1</figureCitation>
,
<figureCitation id="1E4441246D4DE8757F29759CEFD5CF34" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">9</figureCitation>
); mesonotum with yellow to brown basal triangles (Figs
<figureCitation id="5169287CEFFD74C0AA588AC4E7CE5F4C" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">1</figureCitation>
,
<figureCitation id="BEAA6D9C450FFB085B8AE2961C1C981D" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">9</figureCitation>
).
</paragraph>
<paragraph id="C502D73F9D84555B8E04179E5EF81983" pageId="0" pageNumber="135">
Head including eyes broader than pronotum in dorsal view, crown short and broad, round anteriorly, length along midline shorter than one-half width between eyes; coronal suture well developed, extended onto face, and terminating at level of antennal bases (Fig.
<figureCitation id="71059D454D068B4A86E207FF1BD21C49" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">2</figureCitation>
). Ocelli distinct, well separated from eyes (Fig.
<figureCitation id="C87D8400EE60919AD5220E9B67A45DA1" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">2</figureCitation>
). Face moderately broad; lateral frontal suture well developed, curved mesad above antennal pit and meeting coronal suture at midline ventromesad of ocelli; anteclypeus slightly convex, not expanded (Fig.
<figureCitation id="FF74F75A7B4BBE6F93FB06E1BFC912AC" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">2</figureCitation>
). Pronotum large with sinuate transverse depression (Figs
<figureCitation id="CA54E1142C4EA12373DEEAD19DC71461" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">1</figureCitation>
,
<figureCitation id="0C073FDF12ABE13D8DA088F6B78D6C59" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">9</figureCitation>
). Forewing narrow, rounded apically; apical cells occupying almost one-third of total length; vein R2 and RM dissociated at bases, joined by cross-vein, both arising from r cell; vein ScP+RA is not detectable; vein MCu almost parallel with vein RM apically. Hindwing with MP+CuA confluent. Front femur seta AM1 stout, situated near ventral margin; intercalary row with one large basal seta and eight or nine smaller setae near tip of femur. Hind femur with macrosetal formula 2 + 1 + 1; tibia row AV with six or seven preapical macrosetae.
</paragraph>
<paragraph id="5242D719E39504312A9E06585D192B6C" pageId="0" pageNumber="135">
Male basal abdominal sternal apodemes (2S apodemes) well developed (Fig.
<figureCitation id="ED61A95D591F7DEF7BDE9E9DE588FB10" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">3</figureCitation>
). Male pygofer elongated, strongly narrowing caudad, posterior margin acute with few rigid microsetae distally, dorsal margin with macrosetae, long fine ventrolateral setae present, ventral appendage absent (Figs
<figureCitation id="F4835242E2AD3C57B17E5AF687022AEA" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">4</figureCitation>
,
<figureCitation id="31CCBDF51BD6F21E02FA467345FE140F" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">10</figureCitation>
). Anal tube process elongate, extended to ventral margin of genital capsule (Figs
<figureCitation id="C5FC2F7EA5F4C1C90008FCEBE1DD8BD1" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">4</figureCitation>
,
<figureCitation id="4DBF00E48DDA8DACF1264F92C5D5C826" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">10</figureCitation>
). Subgenital plate broadest near base (Fig.
<figureCitation id="76610A875C761E2D261520E00E9387FD" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">10</figureCitation>
) or subapically (Fig.
<figureCitation id="3AD9295B4E3F83B2CFD68772A63A8A11" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">4</figureCitation>
), all categories of setae well differentiated; basal setae encompassing mid-length of plate; marginal setae well defined; macrosetae uniserate; feeble microsetae arranged in 2-4 irregular rows apically (Fig.
<figureCitation id="7545EDD7F2CC95EC6AE9BDFB517F5D0C" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">4</figureCitation>
). Connective with media sclerotization, anterior margin and posterior margin deeply emarginated (Fig.
<figureCitation id="0C5217CCA2EB3677CDDD64734A155C13" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">7</figureCitation>
). Style short, sinuate, with tiny teeth and microsetae preapically (Fig.
<figureCitation id="C728257DD10E0903A7811D19ABB13AFC" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">11</figureCitation>
). Aedeagus shaft short, tubular, with basal apodeme long (Figs
<figureCitation id="BDDCE69134F4A76597329E008A50DDD7" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">5</figureCitation>
,
<figureCitation id="6F72B5DA38D8F461111AE62701825789" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">13</figureCitation>
) or laterally compressed with basal apodeme short (Figs
<figureCitation id="C1FDB6F6DB901403EF6B6364EDF5D394" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">6</figureCitation>
,
<figureCitation id="946437407065BCC6E8DD7CA44B77DF5B" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">12</figureCitation>
); gonopore apical on ventral surface (Fig.
<figureCitation id="14D71BAC429508FA23F122386320FF77" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">12</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C1FF177227BCAC1EC5D5BC440E7DBDEC" pageId="0" pageNumber="135" type="notes">
<paragraph id="D86C41A179977413AAE10D7B55C2664F" pageId="0" pageNumber="135">Notes.</paragraph>
<paragraph id="F0DC8210C4F14A6C6E0466BB987B7537" pageId="0" pageNumber="135">
<taxonomicName id="35F970DE847CFB3473BBD3A5BE307E2C" authorityName="Dworakowska" authorityYear="1995" class="Insecta" family="Cicadellidae" genus="Smyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smyga" order="Hemiptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="BF4C3835B88E7E4083A4D64A4139587D" italics="true" pageId="0" pageNumber="135">Smyga</emphasis>
</taxonomicName>
superficially resembles
<taxonomicName id="2163160AB35057C44E52B3772AAF35B8" authorityName="Mahmood" authorityYear="1967" class="Insecta" family="Cicadellidae" genus="Dapitana" kingdom="Animalia" lsidName="Dapitana" order="Hemiptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="407DA4F49AA508640C1FB398176EEAE5" italics="true" pageId="0" pageNumber="135">Dapitana</emphasis>
</taxonomicName>
Mahmood, 1967 in the features of the head and wings (coronal suture extended onto face and terminating at level of antennal bases (Fig.
<figureCitation id="03341DC5452DAF4D4EC86A30468BFB90" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">2</figureCitation>
), forewing with vein R2 and RM dissociated at bases, joined by cross-vein, both arising from r cell and vein ScP+RA is not detectable). It differs from
<taxonomicName id="20173341DB89F27F9B44A01A65F6820F" authorityName="Mahmood" authorityYear="1967" class="Insecta" family="Cicadellidae" genus="Dapitana" kingdom="Animalia" lsidName="Dapitana" order="Hemiptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="61424358EAEEA804772D2C2E98B2D784" italics="true" pageId="0" pageNumber="135">Dapitana</emphasis>
</taxonomicName>
in color pattern (see generic description) and in having the male pygofer with one or two dorsal macrosetae and long fine ventrolateral setae (Figs
<figureCitation id="D5CD5E23DFAA071B443E7F421268C4D7" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">4</figureCitation>
,
<figureCitation id="912325B44DACEC025C90D2F47EF5E68F" captionStart="Figures 1–13" captionStartId="F1" captionText="Figures 1 - 13. Smyga species 1 - 8 S. brevipenis 1 head and thorax, dorsal view (holotype) 2 face (holotype) 3 - 8 male genitalia after Dworakowska's (1995) figures of aberrant form of S. distincta 3 basal sternal apodemes 4 male genital capsule, left lateral view 5 aedeagus left lateral view 6 aedeagus, ventral view 7 connective 8 connective, style and subgenital plate 9 - 13 S. distincta 9 habitus (holotype) 10 - 13 male genitalia, after Dworakowska (1995) 10 genital capsule 11 style 12 aedeagus ventral view 13 aedeagus, left lateral view." figureDoi="10.3897/zookeys.1198.119765.figures1-13" httpUri="https://binary.pensoft.net/fig/1031824" pageId="0" pageNumber="135">10</figureCitation>
) (pygofer without macrosetae and long fine setae in
<taxonomicName id="6821DA491ADEA3047CA6292618CA41DC" authorityName="Mahmood" authorityYear="1967" class="Insecta" family="Cicadellidae" genus="Dapitana" kingdom="Animalia" lsidName="Dapitana" order="Hemiptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="1A5F850C3C650A8D22CF4EA439BE5251" italics="true" pageId="0" pageNumber="135">Dapitana</emphasis>
</taxonomicName>
). Both genera occur on both sides of
<normalizedToken id="AA1B5678012A6E0BE068F6A01C5FE58F" originalValue="Wallace’s">Wallace's</normalizedToken>
Line, separating the Oriental from Australian regions.
</paragraph>
<paragraph id="90FA532B6F2B7FE18A92B5A15ACC7470" pageId="0" pageNumber="135">
<taxonomicName id="211A42B9B19A5CCCE501276245D254DA" authorityName="Dworakowska" authorityYear="1995" class="Insecta" family="Cicadellidae" genus="Smyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smyga" order="Hemiptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="BF596397BEBF4EDC0D1DC5A1A1CBC209" italics="true" pageId="0" pageNumber="135">Smyga</emphasis>
</taxonomicName>
includes five previously known species, all from Borneo, described by
<bibRefCitation id="846EA42D0B1E7062A4F5C7F15B4BDC13" author="Dworakowska, I" journalOrPublisher="Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden" pageId="0" pageNumber="135" pagination="129 - 160" refId="B1" refString="Dworakowska, I, 1995. Szara gen. nov. and some other Oriental Empoascini (Insecta: Auchenorrhyncha: Cicadellidae: Typhlocybinae). Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 56: 129 - 160" title="Szara gen. nov. and some other Oriental Empoascini (Insecta: Auchenorrhyncha: Cicadellidae: Typhlocybinae)." volume="56" year="1995">Dworakowska (1995)</bibRefCitation>
. In her treatment of the type species,
<taxonomicName id="F9B911D2BAB6A73A4821E7B2036ADCA2" lsidName="S. distincta" pageId="0" pageNumber="135" rank="species" species="distincta">
<emphasis id="41730EEEC961B5DEEDF610B978CEA14E" italics="true" pageId="0" pageNumber="135">S. distincta</emphasis>
</taxonomicName>
, Dworakowska also described and illustrated some &quot;aberrant specimens&quot; that she excluded from the type series because she considered the small, ventrally positioned aedeagal shaft to be &quot;not functional.&quot; Here we recognize these specimens as a valid species:
<taxonomicName id="02208098018535F827EB0FB332773982" lsidName="S. brevipenis" pageId="0" pageNumber="135" rank="species" species="brevipenis">
<emphasis id="66944CDC863C133D6D8E9AB597447398" italics="true" pageId="0" pageNumber="135">S. brevipenis</emphasis>
</taxonomicName>
sp. nov., as similar specimens of two new species have been seen in
<bibRefCitation id="F5499FCD4CA56E8C1E4BB07A67493F8C" author="Xu, Y" journalOrPublisher="Pacific Insects Monographs" pageId="0" pageNumber="135" refId="B4" refString="Xu, Y, 2019. Taxonomy and Phylogeny of the Tribe Empoascini Worldwide (Hemiptera: Cicadellidae: Typhlocybinae). PhD thesis, Northwest A&amp;F University, Shaanxi, China." title="Taxonomy and Phylogeny of the Tribe Empoascini Worldwide (Hemiptera: Cicadellidae: Typhlocybinae). PhD thesis, Northwest A &amp; F University, Shaanxi, China." year="2019">
<normalizedToken id="6A66C830249D881545AF854ECB610B2D" originalValue="Xu’s">Xu's</normalizedToken>
(2019)
</bibRefCitation>
PhD thesis.
</paragraph>
<paragraph id="DFE90153C211536BEFEDFD49D7C1BA94" pageId="0" pageNumber="135">
As most
<taxonomicName id="4D4667AE1C3B1D74A1130A20E1A85D2F" authorityName="Dworakowska" authorityYear="1995" class="Insecta" family="Cicadellidae" genus="Smyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smyga" order="Hemiptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="8D9C0B544B6D042CB2B63AB2ED3589F6" italics="true" pageId="0" pageNumber="135">Smyga</emphasis>
</taxonomicName>
species are from similar localities in Borneo (see Checklist) and as males are needed for identification, the female paratypes of two species (
<taxonomicName id="F0A26398439B045040CB42C27391E876" lsidName="S. distincta" pageId="0" pageNumber="135" rank="species" species="distincta">
<emphasis id="738EE9AFA595FCA94772B0E7260846D3" italics="true" pageId="0" pageNumber="135">S. distincta</emphasis>
</taxonomicName>
and
<taxonomicName id="D57A0E790AA86C62CF3301609F739873" lsidName="S. zonata" pageId="0" pageNumber="135" rank="species" species="zonata">
<emphasis id="7AD5D5DD83FBDE69308300C99D23F72A" italics="true" pageId="0" pageNumber="135">S. zonata</emphasis>
</taxonomicName>
) must be regarded as of uncertain identity (see also comments under
<taxonomicName id="B649AEBFE07559CC184D6D824D364CC7" lsidName="S. distincta" pageId="0" pageNumber="135" rank="species" species="distincta">
<emphasis id="B03844BA6A4EB2F573F8953CBE182188" italics="true" pageId="0" pageNumber="135">S. distincta</emphasis>
</taxonomicName>
and
<taxonomicName id="5FF7A82C02286063F79BB12BB40D850C" lsidName="S. zonata" pageId="0" pageNumber="135" rank="species" species="zonata">
<emphasis id="E74EECEB663C811DDA7D2DDCD43554F3" italics="true" pageId="0" pageNumber="135">S. zonata</emphasis>
</taxonomicName>
). It is also of some interest that, compared to males, the number of known females is very low.
</paragraph>
</subSubSection>
<subSubSection id="BDBDEF57E111C0BA5E4ED8779606CBE5" pageId="0" pageNumber="135" type="distribution">
<paragraph id="8ED48FE48FE87697E70BFDE9AD5DCAF1" pageId="0" pageNumber="135">Distribution.</paragraph>
<paragraph id="15DE27B6F314283D4F9BEA2D0A01D7AF" pageId="0" pageNumber="135">Oriental Region (Brunei, Malaysia).</paragraph>
</subSubSection>
<subSubSection id="016ECF0082AC40CF3F55BD276D5049E5" pageId="0" pageNumber="135" type="checklist to species of smyga">
<paragraph id="BC0DE8A0009B44C123D17D3D4A2BAFD8" pageId="0" pageNumber="135">
Checklist to species of
<taxonomicName id="2D5E01A4354C129588757DB9D0523F80" authorityName="Dworakowska" authorityYear="1995" class="Insecta" family="Cicadellidae" genus="Smyga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Smyga" order="Hemiptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="284975C4D795E2A6E93BCA56B117008C" italics="true" pageId="0" pageNumber="135">Smyga</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="543BDC98AAD04AFACE82951F6BFA1D6A" pageId="0" pageNumber="135">
<taxonomicName id="23526166BBE8EAF4E4EE52C44AF3C051" lsidName="S. brevipenis" pageId="0" pageNumber="135" rank="species" species="brevipenis">
<emphasis id="917B86CF8484A8CD60CBF89B070E3FA9" italics="true" pageId="0" pageNumber="135">S. brevipenis</emphasis>
</taxonomicName>
Webb &amp; Xu, sp. nov. (Brunei, Ulu Temburong; Sarawak, Gunong Mulu National Park)
</paragraph>
<paragraph id="A2FA8889343BFA7D54A365F4AD5ACA1F" pageId="0" pageNumber="135">
<taxonomicName id="FB8437358C05DB9E5772E6A2303037EC" lsidName="S. distincta" pageId="0" pageNumber="135" rank="species" species="distincta">
<emphasis id="BA8C9987B67A5743C662333AF62EE237" italics="true" pageId="0" pageNumber="135">S. distincta</emphasis>
</taxonomicName>
Dworakowska, 1995: 153-155, figs 168-177 (Brunei, Ulu Temburong)
</paragraph>
<paragraph id="2DA48B78ADB41204D017CE1F1A04BC9F" pageId="0" pageNumber="135">
<taxonomicName id="8A64A76A323EFAF40EC9689559F4B8D0" lsidName="S. exhibita" pageId="0" pageNumber="135" rank="species" species="exhibita">
<emphasis id="8D80C85DFE4C53E3310454664FC85240" italics="true" pageId="0" pageNumber="135">S. exhibita</emphasis>
</taxonomicName>
Dworakowska, 1995: 155, figs 198-205 (Brunei, Ulu Temburong)
</paragraph>
<paragraph id="5555BC84C790833E3411157A66AB67B3" pageId="0" pageNumber="135">
<taxonomicName id="2AFEB3B4EB2304208531C479CE26AB51" lsidName="S. niema" pageId="0" pageNumber="135" rank="species" species="niema">
<emphasis id="151BDD6AE8E727471004485A9E143FEA" italics="true" pageId="0" pageNumber="135">S. niema</emphasis>
</taxonomicName>
Dworakowska, 1995: 155-156, figs 206-212 (Sabah)
</paragraph>
<paragraph id="3A4B3C5B8C40D1DE72790187FA4821AD" pageId="0" pageNumber="135">
<taxonomicName id="12DC49451D8B905123CA0A394DA1E44A" lsidName="S. ziewa" pageId="0" pageNumber="135" rank="species" species="ziewa">
<emphasis id="FFC46FF926256606B2F70696F5DDA992" italics="true" pageId="0" pageNumber="135">S. ziewa</emphasis>
</taxonomicName>
Dworakowska, 1995: 155, figs 188-192 (Brunei, Bukit Sulang)
</paragraph>
<paragraph id="B90C1C979BA66A6343E7E4BB0E59162B" pageId="0" pageNumber="135">
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<emphasis id="A3CF22EBF0E63D6B30BAEFCF7F6074D8" italics="true" pageId="0" pageNumber="135">S. zonata</emphasis>
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Dworakowska, 1995: 155, figs 193-197 (Sarawak, Gunung Mulu National Park)
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<paragraph id="2095B1395F0E2858369DEF32AC91223F" pageNumber="372">3. Rheum foliis granulatis, petiolis aequalibus.</paragraph>
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Lapathum orientale, aspero &amp; verrucoso folio, Ribes arabibus dictum.
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Lapathum orientale tomentosum rotundifolium, Ribes arabum dictum.
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Lapathum orientale asperum, folio subrotundo, fructu magno purpureo.
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Ribes arabum.
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Ribes arabum, foliis petasitidis.
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Persia
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Libano
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Carmelo
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<mods:number>2022-09-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>10</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>90015</mods:start>
<mods:end>90015</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/BDJ.10.e90015</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/BDJ.10.e90015</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2828-10-e90015</mods:identifier>
<mods:identifier type="Pensoft-UUID">6DA101C8AAF151B081811854C477EAA8</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:1741F56A5BC958EDAF7C573C2BEF18A1" httpUri="http://treatment.plazi.org/id/1741F56A5BC958EDAF7C573C2BEF18A1" lastPageNumber="90015" pageId="0" pageNumber="90015">
<subSubSection pageId="0" pageNumber="90015" type="nomenclature">
<paragraph pageId="0" pageNumber="90015">
<taxonomicName LSID="1741F56A-5BC9-58ED-AF7C-573C2BEF18A1" authority="(Abonnenc, Léger &amp; Fauran 1979)" baseAuthorityName="Abonnenc, Leger &amp; Fauran" baseAuthorityYear="1979" class="Insecta" family="Asilidae" genus="Nyssomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nyssomyia yuilli pajoti" order="Diptera" pageId="0" pageNumber="90015" phylum="Arthropoda" rank="species" species="yuilli pajoti">
Nyssomyia yuilli pajoti (Abonnenc,
<normalizedToken originalValue="Léger">Leger</normalizedToken>
&amp; Fauran 1979)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="90015" type="distribution">
<paragraph pageId="0" pageNumber="90015">Distribution</paragraph>
<paragraph pageId="0" pageNumber="90015">Porto Velho</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="90015" type="notes">
<paragraph pageId="0" pageNumber="90015">Notes</paragraph>
<paragraph pageId="0" pageNumber="90015">
<bibRefCitation author="Galardo, Allan Kardec Ribeiro" journalOrPublisher="Revista da Sociedade Brasileira de Medicina Tropical" pageId="0" pageNumber="90015" pagination="265 - 271" refId="B7910410" refString="Galardo, Allan Kardec Ribeiro, Galardo, Clicia Denis, Silveira, Guilherme Abbad, Ribeiro, Kaio Augusto Nabas, Hijjar, Andrea Valadao, Oliveira, Liliane Leite, Dos Santos, Thiago Vasconcelos, 2015. Phlebotominae sand flies (Diptera: Psychodidae): potential vectors of American cutaneous leishmaniasis agents in the area associated with the Santo Antonio Hydroelectric System in Western Amazonian Brazil. Revista da Sociedade Brasileira de Medicina Tropical 48 (3): 265 - 271" title="Phlebotominae sand flies (Diptera: Psychodidae): potential vectors of American cutaneous leishmaniasis agents in the area associated with the Santo Antonio Hydroelectric System in Western Amazonian Brazil." volume="48" year="2015">Galardo et al. 2015</bibRefCitation>
</paragraph>
</subSubSection>
</treatment>
</document>