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<document id="F34F1558F5263F43D6969BAC09F79FC2" ID-DOI="10.3897/phytokeys.248.133202" ID-publisher-id="133202" URI-arpha="0CD2E4EE-7B71-5BC1-B3DA-23E14763EDA2" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" article-type="research-article" checkinTime="1731001061505" checkinUser="pensoft" docAuthor="Lavandero, Nicolás, Pérez, Fernanda &amp; Pinilla, Nicolás" docDate="2024" docId="1EFA9A81221F5D47A937FE8F2B86B936" docLanguage="en" docName="PhytoKeys 248: 315-337" docOrigin="PhytoKeys 248" docSource="https://phytokeys.pensoft.net/article/133202/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.phytokeys" docStyleName="PensoftTaxPub.0000.journal_article.phytokeys" docTitle="Leucheria peteroana Lavandero 2024, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="337" masterDocId="0CD2E4EE7B715BC1B3DA23E14763EDA2" masterDocTitle="Leucheria peteroana (Nassauvieae, Asteraceae), a new species of Leucheria endemic to the Andes of Central Chile, and insights into the systematics of Nassauviae" masterLastPageNumber="337" masterPageNumber="315" pageNumber="315" updateTime="1731005913234" updateUser="admin">
<mods:mods id="2C4D58E4D16FE35E71893DF50627DBED" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2BA95D89BB7DB0D758AE28F6EB3C70E4">
<mods:title id="90D42A6E0F4D65A6383AF149A06247A6">Leucheria peteroana (Nassauvieae, Asteraceae), a new species of Leucheria endemic to the Andes of Central Chile, and insights into the systematics of Nassauviae</mods:title>
</mods:titleInfo>
<mods:name id="592B03206A0DDFC5189F5C6CE9C2A463" type="personal">
<mods:role id="7CD12F00F975CA5463F5C12CB3F05A64">
<mods:roleTerm id="E4C54EE830E89EE8DEE6A4298D96990E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9B5B761FDC50C6B2087DBD2C7686EE43">Lavandero, Nicolás</mods:namePart>
<mods:nameIdentifier id="3EB604F2B92AB2679E8EAF4D1FAF99B9" type="ORCID">0000-0002-2390-5078</mods:nameIdentifier>
<mods:affiliation id="6AE7F48EB62DF30C567E4319F3A17401">Facultad de Ciencias Biológicas, Pontificia Universidad Católica de Chile, Avenida Libertador B. O'Higgins 340, Santiago, Chile</mods:affiliation>
</mods:name>
<mods:name id="07D1DCF7BA84B91A34A4DF3AC3907B4B" type="personal">
<mods:role id="24FD7F335D8EA2A44E965162C10F8F41">
<mods:roleTerm id="53E70704488C6C582F0DB65C80EF046D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3F9288CF13BDECA4843D75941FF77EFD">Pérez, Fernanda</mods:namePart>
<mods:nameIdentifier id="964F797D50CEBB423BC6D1884664F3FB" type="ORCID">0000-0001-5730-9760</mods:nameIdentifier>
<mods:affiliation id="3909FF881C9C0AFDB19983D8B368C729">Facultad de Ciencias Biológicas, Pontificia Universidad Católica de Chile, Avenida Libertador B. O'Higgins 340, Santiago, Chile</mods:affiliation>
</mods:name>
<mods:name id="0999057202A58679218238F32E16BB25" type="personal">
<mods:role id="9FD3E75134BD743D1D2A849A356AD08F">
<mods:roleTerm id="A3636E4C19CF015662B515B88D6D4041">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="86B18E17FEC3555712698539D39273E0">Pinilla, Nicolás</mods:namePart>
<mods:nameIdentifier id="E54D01CD44D6FD85EBF418F498CE3F54" type="ORCID">0000-0002-1494-215X</mods:nameIdentifier>
<mods:affiliation id="D6A776CEC0FF5064EE3C38C81C1DA4F1">Facultad de Ciencias Biológicas, Pontificia Universidad Católica de Chile, Avenida Libertador B. O'Higgins 340, Santiago, Chile</mods:affiliation>
</mods:name>
<mods:typeOfResource id="1C1A6D5F2029A61F6ECD2176A4541D50">text</mods:typeOfResource>
<mods:relatedItem id="2F83584C0C57A550805B95EEA48EB335" type="host">
<mods:titleInfo id="DDECCDBE7F086F3949EDED034A8D4300">
<mods:title id="480FCCFE256A2980C28FDECFD2CFD469">PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part id="D8AA35F1FB2C3EDD1FBEE22D024454B1">
<mods:date id="FEED90664904B36E26DAC01271D7EE6C">2024</mods:date>
<mods:detail id="41882A9A043814847C593FBFEC41DB89" type="pubDate">
<mods:number id="8D0C677A24C6DF34FD8373F4FAD1D557">2024-11-07</mods:number>
</mods:detail>
<mods:detail id="76B3CE97E7E9B3F52BEA3EBEE21C3592" type="volume">
<mods:number id="8AC510CC1D23510EB4031672BBCE72C3">248</mods:number>
</mods:detail>
<mods:extent id="A420BDFD8C0E18AA226CF3FB0F06E17B" unit="page">
<mods:start id="B8A3D98E3991DAD0EDDD978844F38B3E">315</mods:start>
<mods:end id="7C82D45ED341B69715CFFE3D9BA17B0D">337</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="BCC3B467D00A294DB84DF882C52F16FF">journal article</mods:classification>
<mods:identifier id="57605B69E569CD732B824DFB35D5FD51" type="DOI">10.3897/phytokeys.248.133202</mods:identifier>
</mods:mods>
<treatment id="1EFA9A81221F5D47A937FE8F2B86B936" ID-arpha="1EFA9A81-221F-5D47-A937-FE8F2B86B936" ID-ipni="urn:lsid:ipni.org:names:77351414-1" LSID="urn:lsid:plazi:treatment:1EFA9A81221F5D47A937FE8F2B86B936" LSID-IPNI="urn:lsid:ipni.org:names:77351414-1" httpUri="http://treatment.plazi.org/id/1EFA9A81221F5D47A937FE8F2B86B936">
<subSubSection id="AEE997B6587BD833C8D7C72DB8D199EC" type="nomenclature">
<paragraph id="7D092F38D6918869AAB27E19CBB521E9">
<taxonomicName id="0E19AD3CD554DD24CCDE1BF22FC8230A" authority="Lavandero" authorityName="Lavandero" authorityYear="2024" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="peteroana" status="sp. nov.">
<emphasis id="6A34F7847255C356345F5002B9F6610B" italics="true">Leucheria peteroana</emphasis>
Lavandero
</taxonomicName>
<taxonomicNameLabel id="4AC6FE4D8ABFDC654E532DC0E0A694A1" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="851BCE26CD8992AB09F73ECEFF3C8C8E" type="description">
<paragraph id="D1C5B16AC5BD403E4359192592EA9EB3">
<figureCitation id="3714BD8305E52242029356C5DF8F81A3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Leucheria peteroana Lavandero, sp. nov. (Lavandero &amp; Pérez 1504, SGO) A habit B detail of the capitulum, apical anther appendages and styles C capitulum, mid-upper view D capitulum, lateral view E leaf, abaxial side F leaf, detail of adaxial side G leaf, detail of abaxial side. All photographs by Nicolás Lavandero." figureDoi="10.3897/phytokeys.248.133202.figure3" httpUri="https://binary.pensoft.net/fig/1169960">Figs 3</figureCitation>
,
<figureCitation id="C12095F9C755D32DBDA12939D5D76176" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habit and habitat of Leucheria peteroana Lavandero, sp. nov. A overview of Laguna El Planchón and the volcanic complex Planchón-Peteroa B detail of the plant C another plant, growing among rocks D detail of plant clump at full sun exposure, but right next to an Andean spring. All photographs by Nicolás Lavandero." figureDoi="10.3897/phytokeys.248.133202.figure4" httpUri="https://binary.pensoft.net/fig/1169961">4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWNAG" type="diagnosis">
<paragraph id="97964A91DE538CCAC6F487E37CE22A02">
<heading id="A0E10C9135CEBED1B1734630F73ED18E" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="E573ADEE49AB5562ECFF285CF357DA28">
<taxonomicName id="26C642E83076649FCBD8173BEFC11073" authorityName="Lavandero" authorityYear="2024" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="peteroana">
<emphasis id="F9C22F4F6316B719880D08D1A2F08C55" italics="true">Leucheria peteroana</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="2D6C40BD677F0D28D0E18A54ADB08FA2" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="runcinata">
<emphasis id="1204D3079FDFC5B44AE70DFB60FF28C5" italics="true">Leucheria runcinata</emphasis>
</taxonomicName>
but differs by its simple aboveground stems (vs. branching stems), solely glandular indumentum (vs. lanose and glandular), lack of any type of scent (vs. strongly pungent or fetid odour), completely white corollas on the adaxial side (vs. lilac to blue), pink-purplish anther apical appendages (vs. blue), pink style branches (vs. white styles), and pappus pectines of 250–520 µm long (vs. 130–160 µm) (Figs
<figureCitation id="DC52733C813979934E37A215FDA7FD62" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison between Leucheria peteroana (NL- 1873, SGO) (A, C, E) and Leucheria runcinata (NL- 1867, SGO) (B, D, F) A, B frontal view of capitula C, D detail of capitula and flowers E, F cypselae and detail of pappus bristles. All photographs by Nicolás Lavandero, except E and F (Nicolás Pinilla)." figureDoi="10.3897/phytokeys.248.133202.figure5" httpUri="https://binary.pensoft.net/fig/1169962">5</figureCitation>
,
<figureCitation id="DF62E674D815ED8D5FA527E98F9A601C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Leaf morphology comparison between Leucheria peteroana and Leucheria runcinata. For each leaf, both adaxial and abaxial sides are shown A Leucheria peteroana, adaxial side (NL 1873, SGO) B Leucheria peteroana, abaxial side (NL 1873, SGO) C Leucheria runcinata, adaxial side (NL- 1867, SGO) D Leucheria runcinata, abaxial side (NL- 1867, SGO)." figureDoi="10.3897/phytokeys.248.133202.figure6" httpUri="https://binary.pensoft.net/fig/1169963">6</figureCitation>
).
<taxonomicName id="578DBD157C08DD903F1F473E5E2BAC71" authorityName="Lavandero" authorityYear="2024" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="peteroana">
<emphasis id="A4D8D9C7F7B7C79A0971332038AC24F0" italics="true">Leucheria peteroana</emphasis>
</taxonomicName>
also differs from
<taxonomicName id="0454CDF61683E53011E3A856DE9DAE0B" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="apiifolia">
<emphasis id="284620B84500A89DB9FADEA06867BCD3" italics="true">Leucheria apiifolia</emphasis>
</taxonomicName>
by its larger height (vs. plants not taller than
<quantity id="5E99254289731163BE630F86DDDE079B" metricMagnitude="-1" metricUnit="m" metricValue="3.0" unit="cm" value="30.0">30 cm</quantity>
.),
<specimenCount id="CE0BD45B417CA22EB9A1262B691C8EFF" count="2" type="generic" typeStatus="types">two types</specimenCount>
of glandular trichomes in the vegetative part (vs. only
<specimenCount id="3A4089D1A1A577AC371F4D0F0F04CB51" count="1" type="generic" typeStatus="type">one type</specimenCount>
of glandular trichome), lack of any type of scent (vs. soft and lemony odour), completely white corollas on the adaxial side (vs. pale gold corollas), outer lip completely extended at full anthesis (vs. outer lip revolute), and pink-purplish anther apical appendages (vs. beige to dark brown) (Fig.
<figureCitation id="4C6F0DED2CCC83932B73DC66BFC32C4B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Other Leucheria species found along the Río Teno Basin in the Maule Region, Central Chile A Leucheria amoena B Leucheria apiifolia C Leucheria candidissima D Leucheria garciana E Leucheria glacialis F Leucheria integrifolia G Leucheria lithospermifolia H Leucheria millefolium. All photographs by Nicolás Lavandero, except L. amoena (Joaquín E. Sepúlveda) and L. candidissima (Guillermo Debandi)." figureDoi="10.3897/phytokeys.248.133202.figure7" httpUri="https://binary.pensoft.net/fig/1169964">7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUPAG" type="material">
<paragraph id="7ECDC9B38E8B8EC75527FA613E8972CB">
<heading id="EF4703FC79571BEF34F7A88FBFCACE2F" reason="title">Type.</heading>
</paragraph>
<paragraph id="0782C2B30CC80D4D303F83D949ECB589">
<materialsCitation id="02920D7D018C13A5BBDFA0D846F8EEB6" collectingDate="2019-03-05" collectionCode="CONC" collectorName="N. Lavandero &amp; L. Santilli &amp; C. Ossa" country="Chile" county="Provincia de Curico" elevation="2593" latitude="-35.16696" location="Lagunas de Teno, alrededores de la Laguna Planchon, sector Norte" longLatPrecision="1" longitude="-70.54383" municipality="comuna de Romeral" specimenCode="SGO 171859" specimenCount="2" stateProvince="Region del Maule" typeStatus="holotype">
<collectingCountry id="9296F2B1CE93507EDC62DB93881A2AE3" name="Chile">Chile</collectingCountry>
•
<collectingRegion id="62F136674E297BF490D48E14140EA308" country="Chile" name="Maule">Región del Maule</collectingRegion>
:
<collectingCounty id="CF9A95945B7DD648E361BFDB8D90EE4A">Provincia de Curicó</collectingCounty>
,
<collectingMunicipality id="366228488A16537BDB49A97FFDCDE747">comuna de Romeral</collectingMunicipality>
.
<location id="B94E6F90FEE7429A76636E04B945C723" LSID="urn:lsid:plazi:treatment:1EFA9A81221F5D47A937FE8F2B86B936:B94E6F90FEE7429A76636E04B945C723" country="Chile" county="Provincia de Curico" latitude="-35.16696" longLatPrecision="1" longitude="-70.54383" municipality="comuna de Romeral" name="Lagunas de Teno, alrededores de la Laguna Planchon, sector Norte" stateProvince="Region del Maule">Lagunas de Teno, alrededores de la Laguna Planchón, sector Norte</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E4PAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-70.543831,-35.166961]}">
<geoCoordinate id="DB35C988BD3B1CBF0329EB2B8A0EA002" degrees="35" direction="south" minutes="10" orientation="latitude" precision="1" seconds="1.06" value="-35.16696">35 ° 10 ' 1.06 &quot; S</geoCoordinate>
,
<geoCoordinate id="9A3E6840900158E9E6DEC348FFAA9F07" degrees="70" direction="west" minutes="32" orientation="longitude" precision="1" seconds="37.79" value="-70.54383">70 ° 32 ' 37.79 &quot; W</geoCoordinate>
</named-content>
,
<quantity id="81A0F457C6802FF95DA056E99C547602" metricMagnitude="3" metricUnit="m" metricValue="2.593" unit="m" value="2593.0">
<elevation id="B5C13BFAE238A8BE7525332930FF825C" metricMagnitude="3" metricUnit="m" metricValue="2.593" unit="m" value="2593.0">2593 m</elevation>
</quantity>
.,
<date id="70E2B6AF9ADD2A6297827F3FA2CD55BD" value="2019-03-05">
<collectingDate id="224CB307FE59F0D84DEFA7A6DA05A051" value="2019-03-05">5 March 2019</collectingDate>
</date>
, fl. And fr.,
<emphasis id="088E0C579B8918347AD1948A2C67AA59" italics="true">
<collectorName id="DAC26B407378773D2A2E5BC59BAD9613">N. Lavandero</collectorName>
</emphasis>
,
<emphasis id="CE0742CD5B1C6D1F07655C96CC90F0D4" italics="true">
<collectorName id="D65E933B6768A21141A70FDDD306E535">L. Santilli</collectorName>
y
<collectorName id="25D31617462F5702B65CD1C8823AB1C5">C. Ossa</collectorName>
</emphasis>
1873 (
<typeStatus id="7BEF2BCCE10A1A1B71CE2923B71FF743" type="holotype">holotype</typeStatus>
:
<specimenCode id="D1B70B1479EDFECCD552D8E5A123AAD4" collectionCode="SGO" country="Chile" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15312" lsid="urn:lsid:biocol.org:col:15312" name="Museo Nacional de Historia Natural" type="Herbarium">SGO 171859</specimenCode>
!;
<typeStatus id="8981D6C00D6E981704B3A9A5B20254C1" type="isotype">Isotype</typeStatus>
<collectionCode id="FA70F8716E49714AB5C64776FC0D82D4" country="Chile" httpUri="http://grbio.org/cool/mhpx-qnxz" name="Universidad de Concepción" type="Herbarium">CONC</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGQAG" type="description">
<paragraph id="EC761640EF415B80027B108469DF6573">
<heading id="1A5724A4B7361C9AFE653734F0373948" reason="title">Description.</heading>
</paragraph>
<paragraph id="FF4141577A4A647F50B3243735AA5C33">
<emphasis id="810FC77813F31911F6211FA0562AB1A2" bold="true" italics="true">Perennial</emphasis>
herb
<quantity id="280EB629C0B793F98F0ADA8F82979FB5" metricMagnitude="-1" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" unit="cm" value="55.0" valueMax="70.0" valueMin="40.0">40–70 cm</quantity>
tall, forming clumps of seasonally persistent annual stems.
<emphasis id="C9F6E290410F884DCEECE034210E8421" bold="true" italics="true">Rhizome</emphasis>
dark brown, round,
<quantity id="E90FCF35802E96BE3C5A15AAD032517E" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="5.0" metricValueMin="2.5" unit="mm" value="37.5" valueMax="50.0" valueMin="25.0">25–50 mm</quantity>
wide, bifurcating, oblique to vertical.
<emphasis id="9272BB0B4007718D5561CA6F67553A77" bold="true" italics="true">Roots</emphasis>
brown, ca.
<quantity id="F076C772345A41C58CCBFC2BD6CEFE07" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
wide.
<emphasis id="46C1B78032CE5DD014C0F21A032EC686" bold="true" italics="true">Stems</emphasis>
green, erect, fistulose, 2.5–5.0 mm wide, simple, never branching, round, internodes up to
<quantity id="4908BECAA91F73DAF790FCE5588A074B" metricMagnitude="-1" metricUnit="m" metricValue="1.0" unit="cm" value="10.0">10 cm</quantity>
long, densely covered by
<specimenCount id="BA8107339ACA2AB30911D1631802C9D0" count="2" type="generic" typeStatus="types">two types</specimenCount>
of trichomes with clear and sticky resin, not fragrant, and without any pungent or noticeable scent when touched or pressed (same indumentum up to the corolla tube): short glandular, capitate, (60 –) 90–150 µm long, multicellular 8–15 - celled trichomes; long glandular, (300 –) 500–1500 (– 2300) µm long, multicellular 10–30 (– 50) - celled trichomes.
<emphasis id="4C7D6A592BBA68145BC23E773215261E" bold="true" italics="true">Leaves</emphasis>
green, alternate; basal leaves attenuate, more densely arranged at the base, but not forming a conspicuous rosette; upper leaves sessile, amplexicaul, loosely arranged, gradually reduced in size towards the capitulescence.
<emphasis id="A58C2ADB2AFBE978AA8EFC668AF55261" bold="true" italics="true">Lamina</emphasis>
oblanceolate, pinnatipartite to pinnatisect, with 6–9 segments per side, almost tripartite towards the apex, (70 –) 130–180 (– 190) × (20 –) 50–60 (– 65) mm; base attenuate, amplexicaul, apex mucronate; margin serrate, texture coriaceous, densely glandulous on both surfaces; segments at the base 1 (– 2) - dentate, apex mucronate; segments in the middle 4–7 - dentate; apical segments fused, 3–7 - dentate; venation prominent on abaxial side, with primary vein ending in apical mucro, secondary veins ending in apical mucro of each segment, and tertiary veins ending in lateral teeth of each segment.
<emphasis id="6E9BF537E3898D4205C9CE92D7799BE9" bold="true" italics="true">Capitulescence</emphasis>
a single corymbiform cyme per stem.
<emphasis id="635D2C3A9C3AA14B8F8259EA971E60BB" bold="true" italics="true">Capitula</emphasis>
5–9 per stem, pedunculate, homogamous, discoid; pedicels (2 –) 8–10 (– 14) cm long.
<emphasis id="FE87E3024328382281205B92B9CC1AFE" bold="true" italics="true">Involucres</emphasis>
hemispheric 10.1–11.0 ×
<quantity id="B8F540B07B125207463AFF98323C501E" metricMagnitude="-2" metricUnit="m" metricValue="1.47" metricValueMax="1.52" metricValueMin="1.42" unit="mm" value="14.7" valueMax="15.2" valueMin="14.2">14.2–15.2 mm</quantity>
, two-seriate, alternate.
<emphasis id="8E75F590D5AE704055BAC1AB6AD51A7B" bold="true" italics="true">Receptacle</emphasis>
convex, epaleate (no flowers between bracts), glabrous.
<emphasis id="F73E34D3A737C479837C34A542EDC0D0" bold="true" italics="true">Outer involucral bracts</emphasis>
(10 –) 12 (– 14), green, lanceolate, concave on the inner face, 8.1–9.2 ×
<quantity id="D5722B3CF67AB0A38106C0D6DB1C9CEE" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" unit="mm" value="1.85" valueMax="1.9" valueMin="1.8">1.8–1.9 mm</quantity>
, with 3 dark-green longitudinal veins (including the midrib), margin ciliate, apex ciliate, texture coriaceous to hyaline-membranaceous towards the margins, abaxial side densely covered by short and long glandular trichomes, adaxial side glabrous.
<emphasis id="EF5E65CC6F01123BAAA5F83BD5C908DC" bold="true" italics="true">Inner involucral bracts</emphasis>
half the number of outer involucral bracts, (5 –) 6 (– 7), green, lanceolate, concave to flat 9.0–9.2 ×
<quantity id="49FF7D0A1EB3186973FC610CCF908297" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" unit="mm" value="2.05" valueMax="2.2" valueMin="1.9">1.9–2.2 mm</quantity>
, with one dark-green longitudinal stripe (midrib), apex acute, texture leaf-like to hyaline-membranaceous towards both lateral margins, margin ciliate, central portion of the abaxial side sparsely covered by short glandular trichomes, hyaline lamina glabrous, adaxial side glabrous.
<emphasis id="5E98B9D6FC7D160970007B0DE3F32EEF" bold="true" italics="true">Flowers</emphasis>
isomorphic, bisexual, (40 –) 43 (– 45) per capitulum.
<emphasis id="B60F906D6E70C09ACD2E5F7CB75DF8A3" bold="true" italics="true">Corollas</emphasis>
bilabiate, white, sometimes pinkish white on the abaxial side, tube
<quantity id="3F72BFBBBA2AB9A8AFFEA1FE5E07CF80" metricMagnitude="-3" metricUnit="m" metricValue="4.449999999999999" metricValueMax="4.6" metricValueMin="4.3" unit="mm" value="4.449999999999999" valueMax="4.6" valueMin="4.3">4.3–4.6 mm</quantity>
long, 1.0–1.1 wide; corolla tube sparsely covered by glandular trichomes.
<emphasis id="12A2BA0C5A641DA18B246A7304E87DE8" bold="true" italics="true">Outer lip</emphasis>
oblanceolate, 6.8–7.1 ×
<quantity id="9295FB3BF25BD4819F71EF1B73CC4C80" metricMagnitude="-3" metricUnit="m" metricValue="3.35" metricValueMax="3.5" metricValueMin="3.2" unit="mm" value="3.35" valueMax="3.5" valueMin="3.2">3.2–3.5 mm</quantity>
at its widest, apex 3 - toothed, teeth equal, 4 - veined, glabrous.
<emphasis id="F87D92AD40189E2979E5E545627EF1F3" bold="true" italics="true">Inner lip</emphasis>
bifid, lacinae linear, 3.4–3.6 ×
<quantity id="6B62213CD7732F55BCCAB8756F5FE6F3" metricMagnitude="-4" metricUnit="m" metricValue="2.45" metricValueMax="2.9" metricValueMin="2.0" unit="mm" value="0.245" valueMax="0.29" valueMin="0.2">0.20–0.29 mm</quantity>
at its widest, connivent, glabrous.
<emphasis id="0A0F5FAB71783E29CB1831F1B42650A2" bold="true" italics="true">Stamens</emphasis>
5,
<quantity id="7CA8CC26F80482B144F0F54731D0C9DF" metricMagnitude="-3" metricUnit="m" metricValue="6.45" metricValueMax="6.7" metricValueMin="6.2" unit="mm" value="6.45" valueMax="6.7" valueMin="6.2">6.2–6.7 mm</quantity>
long, glabrous.
<emphasis id="2CE309B7D78F4305693B2B996733FB9C" bold="true" italics="true">Anthers</emphasis>
sagittate, 3.0–
<quantity id="2B1EC2725FF793BCDCA89043A2EBF90A" metricMagnitude="-3" metricUnit="m" metricValue="3.3" unit="mm" value="3.3">3.3 mm</quantity>
long; apical appendages pink-purplish, lanceolate,
<quantity id="5D52495825CC93714F424133EE4AFB1D" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" unit="mm" value="1.75" valueMax="1.9" valueMin="1.6">1.6–1.9 mm</quantity>
long, apex acute; tails long, lanceolate,
<quantity id="DA74002199253E5BC9FE59D72C09F50A" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.1–1.2 mm</quantity>
long, apex rounded, smooth.
<emphasis id="260C1AEABC1E2688663D8E563EA8172B" bold="true" italics="true">Style</emphasis>
pink, 6.5–7.0 mm long, cleft into two truncate branches of pink colour, branches
<quantity id="5B6461E61C193205325BB50EA6FEF534" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" unit="mm" value="1.2" valueMax="1.3" valueMin="1.1">1.1–1.3 mm</quantity>
long, with stigmatic papillae on internal surface and apical crown papillose.
<emphasis id="98775AC99C1C04AC378EF093BBD71908" bold="true" italics="true">Cypselae</emphasis>
dark-brown, 3.5–3.6 ×
<quantity id="51533E55327C32268579E147D1CD06A9" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.1–1.2 mm</quantity>
, obovoid, strigose; covered by
<specimenCount id="328F02CE32E2556A427A1E48FFD8FD3B" count="2" type="generic" typeStatus="types">two types</specimenCount>
of trichomes: glandular biseriate trichomes, 100–130 µm long, and twin trichomes, 230–280 µm long.
<emphasis id="4DF542D3D42BBBB60146DA4E028D9126" bold="true" italics="true">Pappus</emphasis>
uniseriate, fused at their bases into a ring, deciduous; bristles 23–30, white, sub-plumose,
<quantity id="EFFB1CD059000C48D0A4FF9B9AD61534" metricMagnitude="-3" metricUnit="m" metricValue="7.55" metricValueMax="7.7" metricValueMin="7.4" unit="mm" value="7.55" valueMax="7.7" valueMin="7.4">7.4–7.7 mm</quantity>
long; pectines long, filiform, 250–400 (– 520) µm long, laterally inserted.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGTAG" type="distribution">
<paragraph id="1BB9BD03E33DE3BEA05CBE1757548245">
<heading id="431E3E265D482F811B9F6DE2247C67F2" reason="title">Distribution and habitat.</heading>
</paragraph>
<paragraph id="5B05801077D3F07F8C5FE79DE42655F8">
<taxonomicName id="CBC69AA62214CB29DAAC46B8BCFB28FD" authorityName="Lavandero" authorityYear="2024" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="peteroana">
<emphasis id="109F736D5E24F71F89772FBBDF70F871" italics="true">Leucheria peteroana</emphasis>
</taxonomicName>
is endemic to the Andes of Central
<collectingCountry id="5536667EAD2073CE1B7DE7F7DB764E12" name="Chile">Chile</collectingCountry>
. It is known only from the type locality on the whereabouts of Laguna El Planchón,
<collectingRegion id="E85DC88DCC1FF3475190336AD138C0B5" country="Chile" name="Maule">Maule Region</collectingRegion>
(Fig.
<figureCitation id="7AA08FDFC73486D44AC27A1C70FD2294" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution map of Leucheria peteroana (white triangles) in Chile, Maule Region, based on the type locality and collections." figureDoi="10.3897/phytokeys.248.133202.figure1" httpUri="https://binary.pensoft.net/fig/1169958">1</figureCitation>
). It grows at full sun in margins of Andean wetlands or shaded by large boulders and rock walls near
<quantity id="3077E4B885F062C3A2E725A2A53F0F6A" metricMagnitude="3" metricUnit="m" metricValue="2.5" unit="m" value="2500.0">2500 m</quantity>
a. s. l. with SE orientation (Fig.
<figureCitation id="A5B59D18BDB808617A78719A593D16EB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habit and habitat of Leucheria peteroana Lavandero, sp. nov. A overview of Laguna El Planchón and the volcanic complex Planchón-Peteroa B detail of the plant C another plant, growing among rocks D detail of plant clump at full sun exposure, but right next to an Andean spring. All photographs by Nicolás Lavandero." figureDoi="10.3897/phytokeys.248.133202.figure4" httpUri="https://binary.pensoft.net/fig/1169961">4</figureCitation>
).
<taxonomicName id="481CBC245297201EA646EE52FB385DC3" authorityName="Lavandero" authorityYear="2024" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="peteroana">
<emphasis id="F6056E65FC8BA06D4BA0125106B91FE2" italics="true">L. peteroana</emphasis>
</taxonomicName>
occurs associated with other high Andean plants such as
<taxonomicName id="9EA0BF4B0ADD90B202F12C871E048EF5" authority="(Gillies ex Arn.) Weigend &amp; R. H. Acuna" authorityName="Weigend &amp; R. H. Acuna" baseAuthorityName="Arn." class="Magnoliopsida" family="Loasaceae" genus="Grausa" kingdom="Plantae" order="Cornales" phylum="Tracheophyta" rank="species" species="lateritia">
<emphasis id="A139DED6C2928669B96F32AC6392644F" italics="true">Grausa lateritia</emphasis>
(Gillies ex Arn.) Weigend &amp; R. H. Acuña
</taxonomicName>
,
<taxonomicName id="1639AB42CA27529D094F1F64BC9972AA" authority="Phil." authorityName="Phil." class="Magnoliopsida" family="Calceolariaceae" genus="Calceolaria" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="williamsii">
<emphasis id="67A59DA2CA9D796750F2D72806AAB60A" italics="true">Calceolaria williamsii</emphasis>
Phil.
</taxonomicName>
,
<taxonomicName id="46F1A40A6C89227DE488882104C44F32" authority="Ruiz &amp; Pav." authorityName="Ruiz &amp; Pav." class="Magnoliopsida" family="Rosaceae" genus="Acaena" kingdom="Plantae" order="Rosales" phylum="Tracheophyta" rank="species" species="ovalifolia">
<emphasis id="2C6CFBD22E420FB05EDB3EA552F1DE95" italics="true">Acaena ovalifolia</emphasis>
Ruiz &amp; Pav.
</taxonomicName>
,
<taxonomicName id="EE12EDA5B8B17074541389D8C43078C8" authority="Clos" authorityName="Clos" class="Magnoliopsida" family="Calceolariaceae" genus="Calceolaria" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="filicaulis">
<emphasis id="E73C3DF4AF12AFCE238B68DD8F52EE10" italics="true">Calceolaria filicaulis</emphasis>
Clos
</taxonomicName>
, and
<taxonomicName id="960902FE523E2204AC5F2AE0B889179B" class="Magnoliopsida" family="Phrymaceae" genus="Erythranthe" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="lutea">
<emphasis id="8496EB2162466128857F4F302985C881" italics="true">Erythranthe lutea</emphasis>
</taxonomicName>
(L.) G. L. Nesom.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBWAG" type="phenology">
<paragraph id="4DB576475D92AA347A0AD3036F866401">
<heading id="4F57BC8D65AF9A2F4F018852BEAAC40F" reason="title">Phenology.</heading>
</paragraph>
<paragraph id="ABC9AD7C9ADFAE3867052A8CA67B9D00">Flowering between December and March. Fruiting in March.</paragraph>
</subSubSection>
<subSubSection id="SECID0EGWAG" type="etymology">
<paragraph id="7333A9220D6486F9A78F055561013B91">
<heading id="EE573E4FC29D53686223B4BF897F83EA" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="D1CA23E0A06540DB45BD58AED470F087">The specific epithet refers to the active volcanic complex Planchón-Peteroa. The Andean Lake where the species occurs lies at the foot of this volcano.</paragraph>
<caption id="540917657D468D61096B57775DA33C27" ID-DOI="10.3897/phytokeys.248.133202.figure3" ID-arpha="CDC0572C-D1E8-5FCD-84E5-DFB17C623E5D" httpUri="https://binary.pensoft.net/fig/1169960" startId="F3">
<paragraph id="C09A8772E65230A661B6E31B9A3587CF">
<label id="651B2C0A905E8E2DB03809E2E7373875">Figure 3.</label>
</paragraph>
<paragraph id="34F89C952A5BEEA43FB5E5DCF9065D58">
<taxonomicName id="C9767B158DA0E4B96D6C678445547D48" authority="Lavandero" authorityName="Lavandero" authorityYear="2024" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="peteroana" status="sp. nov.">
<emphasis id="81859C542C7926C3B6CBB57622D4672B" italics="true">Leucheria peteroana</emphasis>
Lavandero
</taxonomicName>
,
<taxonomicNameLabel id="A7F82A8025B1BB90210EC22A53CCFE86" rank="species">sp. nov.</taxonomicNameLabel>
(Lavandero &amp; Pérez 1504, SGO)
<emphasis id="DF4530BFC12A94ED32A52A9CAC90C0DC" bold="true">A</emphasis>
habit
<emphasis id="13DA67F3A85D82179FFF8E2E251673A2" bold="true">B</emphasis>
detail of the capitulum, apical anther appendages and styles
<emphasis id="AD8ACBE58B4F12885EAE474F77E2E415" bold="true">C</emphasis>
capitulum, mid-upper view
<emphasis id="BCFEFD98D8F7AE4505B7E65E15D71D73" bold="true">D</emphasis>
capitulum, lateral view
<emphasis id="DA552079F8D357B1174C49FBF25865E0" bold="true">E</emphasis>
leaf, abaxial side
<emphasis id="423B4EAECA91D513CE05D9504ED5D39C" bold="true">F</emphasis>
leaf, detail of adaxial side
<emphasis id="50B4F5524A2CC18D36E9162288C8BE59" bold="true">G</emphasis>
leaf, detail of abaxial side. All photographs by Nicolás Lavandero.
</paragraph>
</caption>
<caption id="A57BEC9616771836EE0A327C9A0FA87E" ID-DOI="10.3897/phytokeys.248.133202.figure4" ID-arpha="A245306D-C031-5116-B1CA-B72FE35EEEA2" httpUri="https://binary.pensoft.net/fig/1169961" startId="F4">
<paragraph id="E974AF29D02B15634629CC60CF0BA709">
<label id="D72CDE04869FAF56CBDA0D9D8AC091A7">Figure 4.</label>
</paragraph>
<paragraph id="30D602CFE3ABABC8538A94ED4D1891EB">
Habit and habitat of
<taxonomicName id="887B765791BA8B331ABCF48B5C659181" authority="Lavandero" authorityName="Lavandero" authorityYear="2024" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="peteroana" status="sp. nov.">
<emphasis id="42F392ADC1D95F9490D0C4D0607B9961" italics="true">Leucheria peteroana</emphasis>
Lavandero
</taxonomicName>
,
<taxonomicNameLabel id="FB00AE20E5CDF7B8EA536BC3A9A0C79F" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="57895376D455DD157C55FC2B776C339F" bold="true">A</emphasis>
overview of Laguna El Planchón and the volcanic complex Planchón-Peteroa
<emphasis id="F6113448DC61A25C51126EE16E119B86" bold="true">B</emphasis>
detail of the plant
<emphasis id="0B3F085C1060BFAD7ED94B4C9FF6431F" bold="true">C</emphasis>
another plant, growing among rocks
<emphasis id="EF6B89D0A04E7358501C175C9B24211B" bold="true">D</emphasis>
detail of plant clump at full sun exposure, but right next to an Andean spring. All photographs by Nicolás Lavandero.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ERYAG" type="Informal conservation status">
<paragraph id="FF417EF931899A14D21F8F4DCFF9443F">
<heading id="3F5C836CC3576983555EA18ECD35FF07" reason="title">Informal conservation status.</heading>
</paragraph>
<paragraph id="8241D180401AA35FCD1512E2CA781330">
<taxonomicName id="7879A6C5CBC4D6EEE8E28125141FA363" authorityName="Lavandero" authorityYear="2024" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="peteroana">
<emphasis id="29BFA20B148D099763F8FBB0C9A1D33E" italics="true">Leucheria peteroana</emphasis>
</taxonomicName>
can be tentatively considered as Critically Endangered (CR) under the IUCN categories and criteria B 2 ab (ii, iii). Criterion B 2 was selected because its Area of Occupancy is &lt;
<quantity id="FD6AC7FEC7CC458B5B2A602A3A796DCC" metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10 km</quantity>
<superScript id="0BA8CEFD6A55D56C3E90E9B109EF8409">2</superScript>
(
<quantity id="B5C95E7D1635C41703EDEA1A375F32F6" metricMagnitude="3" metricUnit="m" metricValue="4.0" unit="km" value="4.0">4 km</quantity>
<superScript id="7FD69E8C36BA4B9E79B6EDFEF70EF951">2</superScript>
). Criterion “ a ” was selected because it is known to exist at only a single location, with only two known subpopulations. Criterion b (ii, iii) was selected because we expect a continuing decline of suitable conditions for the species to thrive. There is evidence of a decreasing snow cover extent during the dry season of near 15 % per decade in the Andes at mid-latitudes (
<bibRefCitation id="28301B9FFA75D494C9812952A259E047" DOI="10.1038/s41598-019-53486-7" author="Cordero" etAl="et al." firstAuthor="Cordero" issue="1" journalOrPublisher="Scientific Reports" pagination="16945" refId="B7" refString="Cordero RR, Asencio V, Feron S, Damiani A, Llanillo PJ, Sepulveda E, Jorquera J, Carrasco J, Casassa G (2019) Dry-season snow cover losses in the Andes (18–40 S) driven by changes in large-scale climate modes. Scientific Reports 9 (1): 16945. https://doi.org/10.1038/s41598-019-53486-7" title="Dry-season snow cover losses in the Andes (18-40 S) driven by changes in large-scale climate modes." volume="9" year="2019">Cordero et al. 2019</bibRefCitation>
). It is also likely that explosive volcanic eruptions of the Planchón-Peteroa complex, close to the only known locality of the species, may wipe out the whole population. These events are relatively common, with at least 20 eruptions documented since 1600 CE, the most recent occurring between
<date id="07F49A847A129653439256DB317DC0E1" value="2018-09">September 2018</date>
and
<date id="14D40EA99D44520C7E052AEF11F364E8" value="2019-04">April 2019</date>
(
<bibRefCitation id="174A7A95063972492B213D7C4FDB3011" DOI="10.1016/j.jvolgeores.2020.106984" author="Romero" etAl="et al." firstAuthor="Romero" refId="B45" refString="Romero JE, Aguilera F, Delgado F, Guzmán D, Van Eaton AR, Luengo N, Caro J, Bustillos J, Guevara A, Holbik S, Tormey D, Zegarra I (2020) Combining ash analyses with remote sensing to identify juvenile magma involvement and fragmentation mechanisms during the 2018 / 19 small eruption of Peteroa volcano (Southern Andes). Journal of Volcanology and Geothermal Research 402: 106984. https://doi.org/10.1016/j.jvolgeores.2020.106984" year="2020">Romero et al. 2020</bibRefCitation>
).
<taxonomicName id="0D355A5547F1363CB268AC50C5BC73A3" authorityName="Lavandero" authorityYear="2024" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="peteroana">
<emphasis id="E2438875C5D27FB7CA900A3DCA0E54C2" italics="true">Leucheria peteroana</emphasis>
</taxonomicName>
is not present in any known protected area. Although there have not been appropriate efforts to exhaustively locate more populations of
<taxonomicName id="D11062246AC30047AC866EAE6A72838A" authorityName="Lavandero" authorityYear="2024" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="peteroana">
<emphasis id="A208C96166A993A47517EBE08E0E41A3" italics="true">L. peteroana</emphasis>
</taxonomicName>
in the area, it is likely that these would be subject to the same threats as the already known populations. The extent of occurrence (
<abbrev id="ABBRID0EE1AG" xlink_title="extent of occurrence">EOO</abbrev>
) could not be calculated since only two populations are known.
</paragraph>
<caption id="538286B59DA4F6E0DD16D6B74E2EA5FB" ID-DOI="10.3897/phytokeys.248.133202.figure5" ID-arpha="CAA5BAE3-E108-5D17-8EBB-4B47E659DD4B" httpUri="https://binary.pensoft.net/fig/1169962" startId="F5">
<paragraph id="22020CB37CC528943DBBD325D44FA5D3">
<label id="1C2198551336607D6B012212C908742C">Figure 5.</label>
</paragraph>
<paragraph id="A86BFEA92321674076896EA52AF4B8A9">
Comparison between
<taxonomicName id="9AFCA06428B5CADF516A4369B75FC35C" authorityName="Lavandero" authorityYear="2024" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="peteroana">
<emphasis id="4D211B99B5BE9C83750F5F026559756F" italics="true">Leucheria peteroana</emphasis>
</taxonomicName>
(NL- 1873, SGO) (
<emphasis id="F9CFAA7CEAF917C962E47A70736CC574" bold="true">A, C, E</emphasis>
) and
<taxonomicName id="C5517688FF56B3714FA5B4045CDC0AE9" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="runcinata">
<emphasis id="F0AFD8FF42FDB8B37D278CF06287494D" italics="true">Leucheria runcinata</emphasis>
</taxonomicName>
(NL- 1867, SGO) (
<emphasis id="636BEC3E03FE388204968B65FCFEE8B0" bold="true">B, D, F</emphasis>
)
<emphasis id="F6360DA8F6ED227EC7AB41FF09DE4288" bold="true">A, B</emphasis>
frontal view of capitula
<emphasis id="1A20115543B70458E14739275DA77CBF" bold="true">C, D</emphasis>
detail of capitula and flowers
<emphasis id="5034BFFD2BED88594B0BA553F9718275" bold="true">E, F</emphasis>
cypselae and detail of pappus bristles. All photographs by Nicolás Lavandero, except E and F (Nicolás Pinilla).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EV2AG" type="material">
<paragraph id="994EB294E2988F79F59DBD100D9A08CD">
<heading id="136FB540FDB1AD2CB227ECAE689D637E" reason="title">Additional specimens examined.</heading>
</paragraph>
<paragraph id="26944C0541180F7C9991209311E7BCA3">
<materialsCitation id="536C10F48C35570A52D7B44F7C91BEA2" collectingDate="1967-03-10" collectingDateMax="2022-01-08" collectingDateMin="1967-03-10" collectionCode="CONC" collectorName="Marticorena &amp; Matthei" country="Chile" county="Provincia de Curico" elevation="2500" location="A Orillas de la Laguna Teno" municipality="departamento de Curico" specimenCount="1" stateProvince="Region del Maule">
<collectingCountry id="77BD0910A9D27CB978D1B4955C59879D" name="Chile">Chile</collectingCountry>
•
<collectingRegion id="37188FCD43D19B31F06F57F1506C2B04" country="Chile" name="Maule">Región del Maule</collectingRegion>
:
<collectingCounty id="4523F8B0D8AC63F2C5043E36FEC5D99D">Provincia de Curicó</collectingCounty>
,
<collectingMunicipality id="8CAC0FB760366813489D7214E36023B9">departamento de Curicó</collectingMunicipality>
.
<location id="70429BE1AC3505076C8EACB5016BBA88" LSID="urn:lsid:plazi:treatment:1EFA9A81221F5D47A937FE8F2B86B936:70429BE1AC3505076C8EACB5016BBA88" country="Chile" county="Provincia de Curico" municipality="departamento de Curico" name="A Orillas de la Laguna Teno" stateProvince="Region del Maule">A Orillas de la Laguna Teno</location>
.
<quantity id="D2A2FED088CBA2B7146E2E9859B6AFEE" metricMagnitude="3" metricUnit="m" metricValue="2.5" unit="m" value="2500.0">
<elevation id="2861F4BC01859982D8196216E53E8170" metricMagnitude="3" metricUnit="m" metricValue="2.5" unit="m" value="2500.0">2500 m</elevation>
</quantity>
.
<date id="7403570855CF0C59A5C8D6C3E2842CA7" value="1967-03-10">
<collectingDate id="971F2434FECE52602BE9B41E70829100" value="1967-03-10">10 March 1967</collectingDate>
</date>
.
<emphasis id="7AA3E513D36AA12BA19C49255845AF81" italics="true">
<collectorName id="9EBCF9D572EF743F3AF2787AF9D4E0C3">Marticorena</collectorName>
&amp;
<collectorName id="C2A2B7D6CACE44C068031880C48F097A">Matthei</collectorName>
892
</emphasis>
(
<collectionCode id="3C2351C9A649D22068D304936293D0A0" country="Chile" httpUri="http://grbio.org/cool/mhpx-qnxz" name="Universidad de Concepción" type="Herbarium">CONC</collectionCode>
!)
</materialsCitation>
;
<materialsCitation id="5262BD99583DDCD59A75718CEB9B7A5F" collectingDate="2022-01-08" collectingDateMax="2022-01-08" collectingDateMin="1967-03-10" collectionCode="SGO" collectorName="Lavandero &amp; Perez" country="Chile" county="Provincia de Curico" elevation="2570" location="En los alrededores de la Laguna Teno" specimenCount="1" stateProvince="Region del Maule">
<location id="52D76F4C5667BF061C8143C472EBE9DB" LSID="urn:lsid:plazi:treatment:1EFA9A81221F5D47A937FE8F2B86B936:52D76F4C5667BF061C8143C472EBE9DB" country="Chile" county="Provincia de Curico" municipality="departamento de Curico" name="En los alrededores de la Laguna Teno" stateProvince="Region del Maule">En los alrededores de la Laguna Teno</location>
.
<quantity id="70F5717A5608E31D81BDBBB8EFDB63A8" metricMagnitude="3" metricUnit="m" metricValue="2.57" unit="m" value="2570.0">
<elevation id="09AF23C5919B2075DC3813DF56070F3C" metricMagnitude="3" metricUnit="m" metricValue="2.57" unit="m" value="2570.0">2570 m</elevation>
</quantity>
.
<emphasis id="35165E267688F59B1DD83771F9AC9E8B" italics="true">
<collectorName id="2A61C45097BECACA1671C566CE858F89">Lavandero</collectorName>
&amp;
<collectorName id="75E92EBC6CAECB0BF12E01DA633BA8CA">Pérez</collectorName>
1504
</emphasis>
.
<date id="FA5961690789B953AD2C01CE333469AE" value="2022-01-08">
<collectingDate id="39A4BA779DF859EBD71AA090EC53DB9E" value="2022-01-08">8 January 2022</collectingDate>
</date>
(
<collectionCode id="D05B5EECD22A6BDA9188C5C74ED3BD37" country="Chile" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15312" lsid="urn:lsid:biocol.org:col:15312" name="Museo Nacional de Historia Natural" type="Herbarium">SGO</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E62AG" type="notes">
<paragraph id="6611C102DA7F402D39BD4CDB0F632925">
<heading id="7D923C85EAB7114C32CADA2547BC9835" reason="title">Notes.</heading>
</paragraph>
<paragraph id="8E11CDE7ECDFEF2E8F2D38E45E7067BF">
<bibRefCitation id="6DE7BB80876DDDE11A245810EBDEB667" author="Crisci JV" firstAuthor="Crisci" journalOrPublisher="Darwiniana" pagination="9-126" refId="B9" refString="Crisci JV (1976) Revisión del género Leucheria (Compositae: Mutisieae). Darwiniana 20: 9–126." title="Revisión del género Leucheria (Compositae: Mutisieae)." volume="20" year="1976">Crisci (1976)</bibRefCitation>
and posteriorly
<bibRefCitation id="FA82796150624BFDB020A95475DCC6CE" DOI="10.5479/si.19773379" author="Katinas L &amp; Apodaca MJ &amp; Crisci JV" etAl="et al." firstAuthor="Katinas" refId="B26" refString="Katinas L, Apodaca MJ, Crisci JV (2022) A synopsis of Leucheria (Asteraceae, Nassauvieae), with notes on the morphology. Smithsonian Contributions to Botany, No. 115. Smithsonian Institution Scholarly Press, Washington, D. C. https://doi.org/10.5479/si.19773379" year="2022">Katinas et al. (2022)</bibRefCitation>
identified
<emphasis id="E10C107257CA0E87EE1E8DCA23AC4542" italics="true">Marticorena &amp; Matthei 892</emphasis>
as
<taxonomicName id="B84247FE7D0B4E033F472130E91A9F6D" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="apiifolia">
<emphasis id="434987829D8C6D5EAAB7DB058205C1B2" italics="true">Leucheria apiifolia</emphasis>
</taxonomicName>
. The differences between these species are notorious (Fig.
<figureCitation id="556B6283D0ED5BFE60D2C1F0CB956151" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Other Leucheria species found along the Río Teno Basin in the Maule Region, Central Chile A Leucheria amoena B Leucheria apiifolia C Leucheria candidissima D Leucheria garciana E Leucheria glacialis F Leucheria integrifolia G Leucheria lithospermifolia H Leucheria millefolium. All photographs by Nicolás Lavandero, except L. amoena (Joaquín E. Sepúlveda) and L. candidissima (Guillermo Debandi)." figureDoi="10.3897/phytokeys.248.133202.figure7" httpUri="https://binary.pensoft.net/fig/1169964">7</figureCitation>
), since the leaf shape and flower colour differ, but the fact that both plants lack lanate indumentum and the dark colour both species acquire once pressed, may have led to this misidentification.
</paragraph>
<caption id="69DCB4C5D770987DFD1949E44E1D4AD7" ID-DOI="10.3897/phytokeys.248.133202.figure6" ID-arpha="06F5CA31-0B04-5DD7-B270-D53B6C7C81E9" httpUri="https://binary.pensoft.net/fig/1169963" startId="F6">
<paragraph id="CB27F6B777378AE0FA5240301A3E207C">
<label id="8F54D7867A667BBBF21D989FF4745191">Figure 6.</label>
</paragraph>
<paragraph id="E8EF61ABC004CE64E7E45CF5285586CB">
Leaf morphology comparison between
<taxonomicName id="B39276C902D16400676FB468585566C1" authorityName="Lavandero" authorityYear="2024" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="peteroana">
<emphasis id="24B9C652297367745550584182FB95CE" italics="true">Leucheria peteroana</emphasis>
</taxonomicName>
and
<taxonomicName id="D74FD95540C821B6CF6ADED81CE579D9" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="runcinata">
<emphasis id="369B4BBBE688B354A0EFE353662FE83F" italics="true">Leucheria runcinata</emphasis>
</taxonomicName>
. For each leaf, both adaxial and abaxial sides are shown
<emphasis id="7FCA360ED2C3A7E6BF96ECC1337C56F2" bold="true">A</emphasis>
<taxonomicName id="61D3033E06BCF8F0E8C10911AA235DC8" authorityName="Lavandero" authorityYear="2024" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="peteroana">
<emphasis id="FED074C98F05B8AF8F123EAB6D3C5FE7" italics="true">Leucheria peteroana</emphasis>
</taxonomicName>
, adaxial side (NL 1873, SGO)
<emphasis id="7395E8477FC9D3C8B346871CC275E0BC" bold="true">B</emphasis>
<taxonomicName id="68809F27F8FD9EDF543B6C45F6FD5460" authorityName="Lavandero" authorityYear="2024" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="peteroana">
<emphasis id="EC6F0F4ADF2A9F6AE4DFB49F4C2323E9" italics="true">Leucheria peteroana</emphasis>
</taxonomicName>
, abaxial side (NL 1873, SGO)
<emphasis id="7257B191486AACA5F8558F94BFFB3B22" bold="true">C</emphasis>
<taxonomicName id="AE4E7A99D292D77A4B768AF661879DB8" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="runcinata">
<emphasis id="E3CF94CEC44E6BEA02F5CB38205CA9B2" italics="true">Leucheria runcinata</emphasis>
</taxonomicName>
, adaxial side (NL- 1867, SGO)
<emphasis id="BFA896497B3F0A578CB6BE76DBC63383" bold="true">D</emphasis>
<taxonomicName id="40E21C5035FFF6580D7D8D4CB1AAFCB4" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="runcinata">
<emphasis id="4BD8E1DF75074A1842C3EE804E0BAC01" italics="true">Leucheria runcinata</emphasis>
</taxonomicName>
, abaxial side (NL- 1867, SGO).
</paragraph>
</caption>
<caption id="97EFF5593CECC4ECC5A40FB8821F5C71" ID-DOI="10.3897/phytokeys.248.133202.figure7" ID-arpha="4DA4616D-7E41-527B-99EE-6BE7D8769F10" httpUri="https://binary.pensoft.net/fig/1169964" startId="F7">
<paragraph id="EF332D75FCF719E2251378AA87FFBAD7">
<label id="9D46A83CA3797065897AF2ED306E46E1">Figure 7.</label>
</paragraph>
<paragraph id="F0791780272C354A37A3E9B8F3C4C37C">
Other
<taxonomicName id="FDC2806963BC7BF90A33D9D8D95C91C0">
<emphasis id="58982BA6CA9C21F81A981DB6FC242E07" italics="true">Leucheria</emphasis>
species
</taxonomicName>
found along the Río Teno Basin in the Maule Region, Central Chile
<emphasis id="65DA6042454F62724A00879756A79B68" bold="true">A</emphasis>
<taxonomicName id="651835FEB8ADFFE2D8A8EB68FD26AB87" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="amoena">
<emphasis id="AB84197B3A5763460207AB26FA4E8631" italics="true">Leucheria amoena</emphasis>
</taxonomicName>
<emphasis id="7F95714851F2C97A9D16806629C25FAD" bold="true">B</emphasis>
<taxonomicName id="788687527D454A458924D978168C2215" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="apiifolia">
<emphasis id="4F637AD5D6C202520489563EAAD864D2" italics="true">Leucheria apiifolia</emphasis>
</taxonomicName>
<emphasis id="3AB6B204ED1DC77B96A13121E20B783F" bold="true">C</emphasis>
<taxonomicName id="E57D2A353B92B6C424447F24C0ACD647" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="candidissima">
<emphasis id="99ADF91BA18AB8F53667028E33446B9B" italics="true">Leucheria candidissima</emphasis>
</taxonomicName>
<emphasis id="081B3389E1BB03E4B92FF207CFEC7321" bold="true">D</emphasis>
<taxonomicName id="0D34BED833DC443771C5E5D060D33399" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="garciana">
<emphasis id="6F7A93D85062CB5891E89D14867A89DF" italics="true">Leucheria garciana</emphasis>
</taxonomicName>
<emphasis id="071142FD8487E9F9179FB5B462F717A8" bold="true">E</emphasis>
<taxonomicName id="6462749654C1EE5EC4B118874385CBF5" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="glacialis">
<emphasis id="D7476181FD4C81619E26FED2C2C1B56B" italics="true">Leucheria glacialis</emphasis>
</taxonomicName>
<emphasis id="541F9E9D45A7F12B86AA7061435EB43C" bold="true">F</emphasis>
<taxonomicName id="DAB8568BD40AC79C58DC3FFB67CE1489" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="integrifolia">
<emphasis id="F2BD5D61D53358870746FAAE39D74E86" italics="true">Leucheria integrifolia</emphasis>
</taxonomicName>
<emphasis id="D128FB5BB30D2274F1174DD8FC539289" bold="true">G</emphasis>
<taxonomicName id="789ED8A6173E3F6379F59EC20CA7CCCC" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="lithospermifolia">
<emphasis id="5D6063E4304D2E4F48DDC5161882E7BA" italics="true">Leucheria lithospermifolia</emphasis>
</taxonomicName>
<emphasis id="7EF83F76A3723E4600D2F8C0E956E2AA" bold="true">H</emphasis>
<taxonomicName id="2315E4BDF8C9DB8F453199932C088CD4" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="millefolium">
<emphasis id="352EE4CD5F62FF3C34C12FA527E0CD89" italics="true">Leucheria millefolium</emphasis>
</taxonomicName>
. All photographs by Nicolás Lavandero, except
<taxonomicName id="AA99C53C19425D023ED86DE6B4C10CA1" authority="(Joaquin E. Sepulveda)" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="amoena">
<emphasis id="070BEBE9D88614D7AFBDA7386B52554E" italics="true">L. amoena</emphasis>
(Joaquín E. Sepúlveda)
</taxonomicName>
and
<taxonomicName id="67333EF1134D3443DC4AD9552CDA800D" authority="(Guillermo Debandi)" baseAuthorityName="Guillermo Debandi" class="Magnoliopsida" family="Asteraceae" genus="Leucheria" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="candidissima">
<emphasis id="828BED005AD5F9B400FFB2D83638748B" italics="true">L. candidissima</emphasis>
(Guillermo Debandi)
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>