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<document id="E7BAB67F84C6254B1F2CE41E37C0E5D4" ID-CLB-Dataset="30221" ID-DOI="http://dx.doi.org/10.3897/zookeys.751.20611" ID-GBIF-Dataset="9c2aaaed-3a33-47c3-9105-982e2d787d67" ID-PMC="PMC5919911" ID-Pensoft-Pub="1313-2970-751-41" ID-PubMed="29706784" ID-ZooBank="FF9D82A9CB464D78BD6F8C9F69C3CAB8" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-751-41" ModsDocOrigin="ZooKeys 751" ModsDocTitle="Madicolous Chironomidae from the Brazilian Atlantic Forest: a checklist with notes on altitudinal distributions (Diptera, Insecta)" checkinTime="1524205949714" checkinUser="pensoft" docAuthor="Shimabukuro, Erika Mayumi &amp; Trivinho-Strixino, Susana" docDate="2018" docId="B0720D42C49512691290777A8CAE81E6" docLanguage="en" docName="ZooKeys 751: 41-73" docOrigin="ZooKeys 751" docSource="http://dx.doi.org/10.3897/zookeys.751.20611" docTitle="O. japi Wiedenbrug, Mendes, Pepinelli &amp; Trivinho-Strixino 2009" docType="treatment" docVersion="6" lastPageNumber="52" masterDocId="FFE2C733FFE9FFA1A34B5135332ED36E" masterDocTitle="Madicolous Chironomidae from the Brazilian Atlantic Forest: a checklist with notes on altitudinal distributions (Diptera, Insecta)" masterLastPageNumber="73" masterPageNumber="41" pageNumber="52" updateTime="1701385394021" updateUser="ExternalLinkService">
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<mods:title id="3674C51577CADC92196552EEE451E90C">Madicolous Chironomidae from the Brazilian Atlantic Forest: a checklist with notes on altitudinal distributions (Diptera, Insecta)</mods:title>
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<mods:namePart id="40B66E4D6B452E5D9BC0CC418ED58DC7">Shimabukuro, Erika Mayumi</mods:namePart>
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<mods:namePart id="35CC243F6DC0897D080BD1E89FE30BAA">Trivinho-Strixino, Susana</mods:namePart>
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<mods:typeOfResource id="863FB6B55398D9518AC3BFDE1F08EF2A">text</mods:typeOfResource>
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<treatment id="B0720D42C49512691290777A8CAE81E6" ID-GBIF-Taxon="143619813" LSID="urn:lsid:plazi:treatment:B0720D42C49512691290777A8CAE81E6" httpUri="http://treatment.plazi.org/id/B0720D42C49512691290777A8CAE81E6" lastPageNumber="52" pageId="11" pageNumber="52">
<subSubSection id="0C01C3E0AD4A03826829123FF44853D1" pageId="11" pageNumber="52" type="nomenclature">
<paragraph id="1ECCE55BAD517097E2137C5369EC1111" pageId="11" pageNumber="52">
<taxonomicName id="F343642DCF1926C9ADC4DF684320F380" ID-CoL="K3JZ" authority="Wiedenbrug, Mendes, Pepinelli &amp; Trivinho-Strixino, 2009" authorityName="Wiedenbrug, Mendes, Pepinelli &amp; Trivinho-Strixino" authorityYear="2009" family="Chironomidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="O. japi" order="Diptera" pageId="11" pageNumber="52" rank="species" species="japi">O. japi Wiedenbrug, Mendes, Pepinelli &amp; Trivinho-Strixino, 2009</taxonomicName>
</paragraph>
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<subSubSection id="602E5924604F2CCAFEE6D9B2C8C73BB7" pageId="11" pageNumber="52" type="distribution">
<paragraph id="483762C59161DD56663F6760096B8C48" pageId="11" pageNumber="52">Distribution.</paragraph>
<paragraph id="702989F77C2D96AE7C12049C59CFE22D" pageId="11" pageNumber="52">
BRAZIL, Serra do Japi,
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,
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Paulo State (
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,
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); PESM, Ubatuba,
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Paulo State (
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,
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and
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,
<geoCoordinate id="2FB8AF0BD2D1369C7D53D165BBF5DA5A" direction="west" orientation="longitude" precision="1" value="-45.2407">45°14.442'W</geoCoordinate>
).
</paragraph>
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<paragraph id="E5950391FB00C57E9DDB560D7B75BE61" pageId="11" pageNumber="52">Elevation.</paragraph>
<paragraph id="400A40A4C71EFC8D6A632972B5A58160" pageId="11" pageNumber="52">1058 m a.s.l.</paragraph>
</subSubSection>
<subSubSection id="5D6B59B7FF1C46235C6423C792E70126" pageId="11" pageNumber="52" type="habitat">
<paragraph id="5CC48D3CF908EA55B43BE1129BF3426D" pageId="11" pageNumber="52">Habitats.</paragraph>
<paragraph id="1C15A679B456CD51AFF17A7D9381453D" pageId="11" pageNumber="52">Litter below a waterfall of a first-order stream.</paragraph>
</subSubSection>
<subSubSection id="853033DD345209CA179089B04DD6F301" pageId="11" pageNumber="52" type="known stages">
<paragraph id="0ECB2EA19E053DF8D037F3FCC643FDAF" pageId="11" pageNumber="52">Known stages.</paragraph>
<paragraph id="E6C610BCDE93582C8AD41A5ED79F1D0A" pageId="11" pageNumber="52">L, P, F, M.</paragraph>
</subSubSection>
<subSubSection id="7AA89631DF0AEFF9E39B4B533CAB56EE" pageId="11" pageNumber="52" type="references">
<paragraph id="5347A6AD8AA478ED872D9381BD743FBB" pageId="11" pageNumber="52">References.</paragraph>
<paragraph id="78B324782C7FDBE332764A32AEA15FCD" pageId="11" pageNumber="52">
<bibRefCitation id="D3439DC0E9C943E6A23BF6A727BA2B09" author="Wiedenbrug, S" journalOrPublisher="Zootaxa" pageId="30" pageNumber="71" pagination="1 - 26" title="Review of the genus Onconeura Andersen et Saether (Diptera: Chironomidae), with the description of four new species from Brazil." url="https://doi.org/10.5281/zenodo.190820" volume="2265" year="2009">Wiedenbrug et al. (2009)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3EE81D305D5F2BAAC7616B3C9561377F" pageId="11" pageNumber="52" type="remarks">
<paragraph id="45A696DE99DFF896D4E333618C7FA021" pageId="11" pageNumber="52">Remarks.</paragraph>
<paragraph id="B646AE931731B064FC80B1690365C08B" pageId="11" pageNumber="52">
<taxonomicName id="8C3FD45D50819656EF1B892355E2B6E3" lsidName="O. japi" pageId="11" pageNumber="52" rank="species" species="japi">O. japi</taxonomicName>
was found on marginal rocks of small streams, at 1570 m a.s.l., extending the altitudinal record of the species. Environmental characterization: Water temperature 16.3 °C; dissolved oxygen 8.4 mg.l-1; pH 6.4; slow flowing; vegetal canopy completely absent. The species was found in APASM (Minas Gerais State), extending the geographical distribution of the species. Low altitudinal specificity (IndVal: 0.14; p = 1.0).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:namePart>Chen, Zhe-Yu</mods:namePart>
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<mods:namePart>Wu, Min</mods:namePart>
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<mods:affiliation>School of Life Sciences, Nanjing University, Nanjing 210023, China</mods:affiliation>
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<taxonomicName LSID="B0722042-8149-51EB-A94D-D218446E36F0" authority="(Martens, 1875)" baseAuthorityName="Martens" baseAuthorityYear="1875" class="Gastropoda" family="Camaenidae" genus="Stegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegodera angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis">Stegodera angusticollis (Martens, 1875)</taxonomicName>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Bayesian Inferences of camaenids (representatives of genera in Table 1) based on partial mitochondrial 16 S and CO 1 sequences. Numbers near nodes indicate Bayesian posterior probabilities. The squares on the right of each taxon name indicate its character states. Abbreviations: PS: penial sheath; EpP: epiphallic papilla; PC: penial caecum; Fl: flagellum; DS: dart sac apparatus A phylogram without Australian camaenids B the results of ancestral states (tagged on branch in red, blue, or black) reconstruction mapping on the phylogram with some Australian camaenid genera added. Orange part: the topologies supported by the results respectively by using both Bayesian Inference and Maximum Likelihood methods. Character states are from Schileyko (2003) and Wu (2019). Scale bars for substitutions per site." figureDoi="10.3897/zookeys.1059.68385.figure1" httpUri="https://binary.pensoft.net/fig/584114" pageId="0" pageNumber="1">Figures 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution of Stegodera and Pseudostegodera qiului gen. et sp. nov. Triangle: new locality of Stegodera angusticollis (Martens, 1875); dashed range: area of Stegodera angusticollis; circle: type locality of Pseudostegodera qiului gen. et sp. nov." figureDoi="10.3897/zookeys.1059.68385.figure2" httpUri="https://binary.pensoft.net/fig/584115" pageId="0" pageNumber="1">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells A Stegodera angusticollis Martens, 1876, HBUMM 08435 B-D Pseudostegodera qiului gen. et sp. nov., holotype IZCAS TM 206978 C p rotoconch, magnified D shell surface, magnified. Scale bar: 5 mm (A, B)." figureDoi="10.3897/zookeys.1059.68385.figure3" httpUri="https://binary.pensoft.net/fig/584116" pageId="0" pageNumber="1">, 3A</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. General anatomy A-D Stegodera angusticollis (Martens, 1875), HBUMM 08435 A lobe (arrowed) on the left side of mantle edge B lobe (arrowed) on the right side of mantle edge C internal surface of head region D jaw E-H Pseudostegodera qiului gen. et sp. nov., holotype IZCAS TM 206978 E jaw F internal surface of head region G lobe (arrowed) on the left side of mantle edge H lobe (arrowed) on the right side of mantle edge." figureDoi="10.3897/zookeys.1059.68385.figure4" httpUri="https://binary.pensoft.net/fig/584117" pageId="0" pageNumber="1">, 4A-D</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A general view B male part, showing the vas deferens connecting epiphallus (arrowed) C female part, showing the dart sac, accessory sac, and mucous glands." figureDoi="10.3897/zookeys.1059.68385.figure5" httpUri="https://binary.pensoft.net/fig/584118" pageId="0" pageNumber="1">, 5A-C</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A exposed dart sac apparatus B magnified partial penis and partial epiphallus. Arrow showing a very thin membranous sac wrapping distal half of penis with distal end connecting basal penial retractor muscle." figureDoi="10.3897/zookeys.1059.68385.figure6" httpUri="https://binary.pensoft.net/fig/584119" pageId="0" pageNumber="1">, 6A, B</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitat and living specimens of Stegodera angusticollis (Martens, 1875) A habitat (photograph Kai-Chen Ouyang) B the living specimen of HBUMM 08435 C a living specimen from Chibi City (photograph Lu Qiu)." figureDoi="10.3897/zookeys.1059.68385.figure8" httpUri="https://binary.pensoft.net/fig/584121" pageId="0" pageNumber="1">, 8A-C</figureCitation>
</paragraph>
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<taxonomicName authorityName="Martens" authorityYear="1875" class="Gastropoda" family="Camaenidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis">Helix angusticollis</taxonomicName>
Martens, 1875a: 2.
<bibRefCitation author="Martens, E von" journalOrPublisher="Malakozoologische Blaetter" pageId="0" pageNumber="1" pagination="185 - 188" refId="B22" refString="Martens, E von, 1875b. Binnen-Mollusken aus dem mittlern China. Malakozoologische Blaetter 22: 185 - 188" title="Binnen-Mollusken aus dem mittlern China." volume="22" year="1875 b">Martens 1875b</bibRefCitation>
: 185; Pfeiffer 1875: 449, 1876: 149, pl. 134, figs 7-10;
<bibRefCitation author="Gredler, V" journalOrPublisher="Malakozoologische Blaetter, Neue Folge" pageId="0" pageNumber="1" pagination="165 - 187" refId="B10" refString="Gredler, V, 1882. Uebersicht der Binnenschnecken von China. Malakozoologische Blaetter, Neue Folge 5: 165 - 187" title="Uebersicht der Binnenschnecken von China." volume="5" year="1882">Gredler 1882</bibRefCitation>
: 175;
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.50365" author="Heude, PM" journalOrPublisher="Memoires concernant l'histoire naturelle de l'Empire chinois" pageId="0" pageNumber="1" pagination="1 - 84" refId="B13" refString="Heude, PM, 1882. Notes sur les mollusques terrestres de la vallee du Fleuve Bleu. Memoires concernant l'histoire naturelle de l'Empire chinois 1: 1 - 84, DOI: https://doi.org/10.5962/bhl.title.50365" title="Notes sur les mollusques terrestres de la vallee du Fleuve Bleu." url="https://doi.org/10.5962/bhl.title.50365" volume="1" year="1882">Heude 1882</bibRefCitation>
: 36, pl. 15, fig. 8;
<bibRefCitation author="Moellendorff, OF" journalOrPublisher="Jahrbuecher der Deutschen Malakozoologischen Gesellschaft" pageId="0" pageNumber="1" pagination="307 - 391" refId="B23" refString="Moellendorff, OF, 1884. Materialien zur Fauna von China. Jahrbuecher der Deutschen Malakozoologischen Gesellschaft 11: 307 - 391" title="Materialien zur Fauna von China." volume="11" year="1884">
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1884
</bibRefCitation>
: 387.
</paragraph>
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<taxonomicName authorityName="Martens" authorityYear="1875" class="Gastropoda" family="Camaenidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix (Stegodera) angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis" subGenus="Stegodera">Helix (Stegodera) angusticollis</taxonomicName>
. - Martens, 1876 (in
<bibRefCitation author="Pfeiffer, L" journalOrPublisher="Jahrbuecher der Deutschen Malakozoologischen Gesellschaft" pageId="0" pageNumber="1" refId="B26" refString="Pfeiffer, L, 1870-1876. Novitates Conchologicae. Series prima. Mollusca extramarina. Descriptions et figures de coquilles extramarines nouvelles ou peu connues. Tome IV, 1-8, pls 109-111 [1870], 9-40, pls 112-117 [1871], 41-88, pls 118-125 [1872], 89-112, pls 126-130 [1874], 113-144, pls 131-134 [1875], 145-171, pls 135-137." title="Novitates Conchologicae. Series prima. Mollusca extramarina. Descriptions et figures de coquilles extramarines nouvelles ou peu connues. Tome IV, 1 - 8, pls 109 - 111 [1870], 9 - 40, pls 112 - 117 [1871], 41 - 88, pls 118 - 125 [1872], 89 - 112, pls 126 - 130 [1874], 113 - 144, pls 131 - 134 [1875], 145 - 171, pls 135 - 137." year="1870 - 1876">Pfeiffer 1870-1876</bibRefCitation>
): 149-150;
<bibRefCitation author="Pilsbry, HA" journalOrPublisher="Academy of Natural Science Philadelphia, PA" pageId="0" pageNumber="1" refId="B27" refString="Pilsbry, HA, 1890. [1890-1891] Manual of conchology, Structural and systematic, with illustrations of the species, Second series, Volume 6 (Helicidae, Vol. IV). Academy of Natural Science Philadelphia, PA" title="[1890 - 1891] Manual of conchology, Structural and systematic, with illustrations of the species, Second series, Volume 6 (Helicidae, Vol. IV)." year="1890">Pilsbry 1890</bibRefCitation>
: 7, pl. 1, figs 15-17.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Camaenidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis (Stegodera) angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis" subGenus="Stegodera">Plectopylis (Stegodera) angusticollis</taxonomicName>
. - Pilsbry 1894: 147.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Camaenidae" genus="Planispira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Planispira (Stegodera) angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis" subGenus="Stegodera">Planispira (Stegodera) angusticollis</taxonomicName>
. -
<bibRefCitation author="Thiele, J" journalOrPublisher="Journal of Molluscan Studies" pageId="0" pageNumber="1" refId="B40" refString="Thiele, J, 1931. Handbuch der systematischen Weichtierkunde. Vol. 1, Teil 2: 377-778. Gustav Fischer, Jena." title="Handbuch der systematischen Weichtierkunde. Vol. 1, Teil 2: 377 - 778. Gustav Fischer, Jena." year="1931">Thiele 1931</bibRefCitation>
: 681.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Martens" baseAuthorityYear="1875" class="Gastropoda" family="Camaenidae" genus="Stegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegodera angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis">Stegodera angusticollis</taxonomicName>
. -
<bibRefCitation author="Gude, GK" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="1" pagination="1 - 11" refId="B11" refString="Gude, GK, 1902. A classified list of the helicoid land shells of Asia. Proceedings of the Malacological Society of London 9 (1): 1 - 11" title="A classified list of the helicoid land shells of Asia." volume="9" year="1902">Gude 1902</bibRefCitation>
: 4; Kobelt 1905: 78;
<bibRefCitation author="Pilsbry, HA" journalOrPublisher="The Nautilus" pageId="0" pageNumber="1" pagination="63 - 67" refId="B29" refString="Pilsbry, HA, 1905. Notes on Moellendorffia and Stegodera. The Nautilus 19: 63 - 67" title="Notes on Moellendorffia and Stegodera." volume="19" year="1905">Pilsbry 1905</bibRefCitation>
: 64, 66, pl. 2, figs 1-3;
<bibRefCitation author="Gude, GK" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="1" pagination="52 - 73" refId="B12" refString="Gude, GK, 1920. The armature of land Mollusca. Proceedings of the Malacological Society of London 14 (1): 52 - 73" title="The armature of land Mollusca." volume="14" year="1920">Gude 1920</bibRefCitation>
: 60;
<bibRefCitation author="Yen, TC" journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft" pageId="0" pageNumber="1" pagination="1 - 234" refId="B50" refString="Yen, TC, 1939. Die chinesischen Land- und Suesswasser-Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 444: 1 - 234" title="Die chinesischen Land- und Suesswasser-Gastropoden des Natur-Museums Senckenberg." volume="444" year="1939">Yen 1939</bibRefCitation>
: 126, pl. 13, fig. 9;
<bibRefCitation author="Zilch, A" journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft" pageId="0" pageNumber="1" refId="B51" refString="Zilch, A, 1960. Teil 2, Euthyneura. In: Wenz W (Ed.) Gastropoda, Handbuch der Palaeozoologie (ed. Schindewolf OH), Band 6, Gastropoda. Berlin: Gebrueder Borntraeger, 835 pp. [(4): 601-835 (1960)] [Abb. 2112-2515, I-XII]" title="Teil 2, Euthyneura. In: Wenz W (Ed.) Gastropoda, Handbuch der Palaeozoologie (ed. Schindewolf OH), Band 6, Gastropoda. Berlin: Gebrueder Borntraeger, 835 pp. [(4): 601 - 835 (1960)] [Abb. 2112 - 2515, I-XII]" year="1960">Zilch 1960</bibRefCitation>
: fig. 2138;
<bibRefCitation author="Richardson, CL" journalOrPublisher="Tryonia" pageId="0" pageNumber="1" pagination="1 - 253" refId="B34" refString="Richardson, CL, 1983. Bradybaenidae: Catalog of species. Tryonia 12: 1 - 253" title="Bradybaenidae: Catalog of species." volume="12" year="1983">Richardson 1983</bibRefCitation>
: 291;
<bibRefCitation author="Schileyko, AA" journalOrPublisher="Ruthenica, Supplement" pageId="0" pageNumber="1" pagination="1467 - 1626" refId="B36" refString="Schileyko, AA, 2003. Treatise on recent terrestrial pulmonate molluscs, Part 11. Trigonochlamydidae, Papillodermidae, Vitrinidae, Limacidae, Bielziidae, Agriolimacidae, Boettgerillidae, Camaenidae. Ruthenica, Supplement 2 (11): 1467 - 1626" title="Treatise on recent terrestrial pulmonate molluscs, Part 11. Trigonochlamydidae, Papillodermidae, Vitrinidae, Limacidae, Bielziidae, Agriolimacidae, Boettgerillidae, Camaenidae." volume="2" year="2003">Schileyko 2003</bibRefCitation>
: 1512, fig. 1948;
<bibRefCitation author="Chen, DN" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="1" refId="B4" refString="Chen, DN, Gao, JX, 1987. Economic Fauna of China (Terrestrial Molluscs). Science Press, Beijing" title="Economic Fauna of China (Terrestrial Molluscs)." year="1987">Chen and Gao 1987</bibRefCitation>
: 107, fig. 137; Qian et al. 2008: 289, fig. 169;
<bibRefCitation author="Qian, ZX" journalOrPublisher="Zhejiang People's Fine Arts Publishing House, Hangzhou" pageId="0" pageNumber="1" refId="B30" refString="Qian, ZX, Zhou, WC, 2014. Illustrated Handbook of Common Terrestrial Mollusks in China. Zhejiang People's Fine Arts Publishing House, Hangzhou" title="Illustrated Handbook of Common Terrestrial Mollusks in China." year="2014">Qian and Zhou 2014</bibRefCitation>
: 120, figure in text.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Museum material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation accessionNumber="SMF27113, SMF27114, SMF42579, TM108733, TM158903, TM159272" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="v. Richthofen, O. v. Moellendorff, K. Hashagen" country="China" location="Hubei" specimenCount="1" stateProvince="Hubei" typeStatus="syntype">
• ZMB. Moll. 31044,
<typeStatus>syntype</typeStatus>
, Poyangsee,
<collectingCountry name="China">China</collectingCountry>
, slg.
<collectorName>v. Richthofen</collectorName>
; ZMB. • ZMB. Moll. 3710/1
<collectingCountry name="China">China</collectingCountry>
, slg. Paetel; •
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF27113">SMF27113</accessionNumber>
/3,
<collectingCountry name="China">China</collectingCountry>
: Prov. Hupei, slg.
<collectorName>O. v. Moellendorff</collectorName>
; •
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF27114">SMF27114</accessionNumber>
/2,
<collectingCountry name="China">China</collectingCountry>
:
<normalizedToken originalValue="Wǔ-chang-fú">Wǔ-chang-fu</normalizedToken>
, slg.
<collectorName>K. Hashagen</collectorName>
(ex. Schmacker); •
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF42579">SMF42579</accessionNumber>
/3,
<collectingCountry name="China">China</collectingCountry>
:
<normalizedToken originalValue="Húbei">
<collectingRegion country="China" name="Hubei">Hubei</collectingRegion>
</normalizedToken>
(Hupe), slg. Ehrmann; • IZCAS
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/TM108733">TM108733</accessionNumber>
/1, Kieou-Kiang; • IZCAS
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/TM158903">TM158903</accessionNumber>
-158920/18, Ou-tchang h[ien], ex.
<normalizedToken originalValue="Musée">Musee</normalizedToken>
Heude; • IZCAS
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/TM159272">TM159272</accessionNumber>
-159397/126, Kieou-Kiang, ex.
<normalizedToken originalValue="Musée">Musee</normalizedToken>
Heude
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">New material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2020-10-15" collectorName="Xiao-Long Wang, Zi-Hao Shen" country="China" county="Yangxin County" elevation="140" latitude="30.00106" location="One" longLatPrecision="1" longitude="115.19012" municipality="Huangshi City" specimenCount="1" stateProvince="Hubei Province">
•
<location LSID="urn:lsid:plazi:treatment:B0722042814951EBA94DD218446E36F0:DB5BE5FCBEEEBDAAC388CC85D09E8027" country="China" county="Yangxin County" latitude="30.00106" longLatPrecision="1" longitude="115.19012" municipality="Huangshi City" name="One" stateProvince="Hubei Province">One</location>
shell of HBUMM08435 (dissected),
<location LSID="urn:lsid:plazi:treatment:B0722042814951EBA94DD218446E36F0:0662AF0339E1A57D283FB9415CC034FC" country="China" county="Yangxin County" latitude="30.00106" longLatPrecision="1" longitude="115.19012" municipality="Huangshi City" name="Guanyin Cave" stateProvince="Hubei Province">Guanyin Cave</location>
[观音洞],
<location LSID="urn:lsid:plazi:treatment:B0722042814951EBA94DD218446E36F0:2FC890E59434868C23A203998F2BB7C1" country="China" county="Yangxin County" latitude="30.00106" longLatPrecision="1" longitude="115.19012" municipality="Huangshi City" name="Taizi Town" stateProvince="Hubei Province">Taizi Town</location>
[太子镇],
<collectingCounty>Yangxin County</collectingCounty>
[阳新县],
<collectingMunicipality>Huangshi City</collectingMunicipality>
[黄石市],
<collectingRegion country="China" name="Hubei">Hubei Province</collectingRegion>
,
<collectingCountry name="China">China</collectingCountry>
,
<geoCoordinate degrees="30" direction="north" minutes="0" orientation="latitude" precision="1" seconds="3.816" value="30.00106">30°0'3.816&quot;N</geoCoordinate>
,
<geoCoordinate degrees="115" direction="east" minutes="11" orientation="longitude" precision="1" seconds="24.428" value="115.19012">115°11'24.428&quot;E</geoCoordinate>
,
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.4" unit="m" value="140.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.4" unit="m" value="140.0">140 m</quantity>
a.s.l.
</elevation>
,
<collectingDate value="2020-10-15">2020-X-15</collectingDate>
, leg.
<collectorName>Xiao-Long Wang</collectorName>
&amp;
<collectorName>Zi-Hao Shen</collectorName>
(SDmaj =
<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.67" unit="mm" value="26.7">26.7 mm</quantity>
, SDmin =
<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.13" unit="mm" value="21.3">21.3 mm</quantity>
, SH =
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.13" unit="mm" value="11.3">11.3 mm</quantity>
)
</materialsCitation>
. •
<materialsCitation collectingDate="2020-10-03" collectorName="Di Yu, Kai-Chen Ouyang." country="China" county="Xianning City" latitude="29.67" location="Three" longLatPrecision="20" longitude="113.97639" municipality="Chibi City" specimenCount="1" stateProvince="Hubei Province">
<location LSID="urn:lsid:plazi:treatment:B0722042814951EBA94DD218446E36F0:B1C6973FC7A51022A15687C0556A357C" country="China" county="Xianning City" latitude="29.67" longLatPrecision="20" longitude="113.97639" municipality="Chibi City" name="Three" stateProvince="Hubei Province">Three</location>
shells of OYKC, one shell of QL,
<location LSID="urn:lsid:plazi:treatment:B0722042814951EBA94DD218446E36F0:DEC5B63CA9E0187F0D34A8261F45FB1E" country="China" county="Xianning City" latitude="29.67" longLatPrecision="20" longitude="113.97639" municipality="Chibi City" name="Lushui Lake" stateProvince="Hubei Province">Lushui Lake</location>
[陆水湖],
<collectingMunicipality>Chibi City</collectingMunicipality>
[赤壁市],
<collectingCounty>Xianning City</collectingCounty>
[咸宁市],
<collectingRegion country="China" name="Hubei">Hubei Province</collectingRegion>
,
<collectingCountry name="China">China</collectingCountry>
,
<geoCoordinate degrees="29" direction="north" minutes="40" orientation="latitude" precision="15" seconds="12" value="29.67">29°40'12&quot;N</geoCoordinate>
,
<geoCoordinate degrees="113" direction="east" minutes="58" orientation="longitude" precision="15" seconds="35" value="113.97639">113°58'35&quot;E</geoCoordinate>
,
<collectingDate value="2020-10-03">2020-X-03</collectingDate>
, leg.
<collectorName>Di Yu</collectorName>
&amp;
<collectorName>Kai-Chen Ouyang.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">Poyang-See (= Poyang Lake [鄱阳湖], Jiangxi).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Measurements of new material.</paragraph>
<paragraph pageId="0" pageNumber="1">
SDmaj =
<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.8649999999999998" metricValueMax="3.06" metricValueMin="2.67" unit="mm" value="28.65" valueMax="30.6" valueMin="26.7">26.7-30.6 mm</quantity>
, SDmin =
<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.37" metricValueMin="2.13" unit="mm" value="22.5" valueMax="23.7" valueMin="21.3">21.3-23.7 mm</quantity>
, SH =
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.175" metricValueMax="1.22" metricValueMin="1.13" unit="mm" value="11.75" valueMax="12.2" valueMin="11.3">11.3-12.2 mm</quantity>
(n = 6).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">Body whorl completely covering partial penultimate whorl. Penial sheath weakly present. Epiphallic papilla absent. Dart sac apparatus and flagellum present. Mucous glands with numerous tubes.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="redescription">
<paragraph pageId="0" pageNumber="1">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="1">
Shell (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells A Stegodera angusticollis Martens, 1876, HBUMM 08435 B-D Pseudostegodera qiului gen. et sp. nov., holotype IZCAS TM 206978 C p rotoconch, magnified D shell surface, magnified. Scale bar: 5 mm (A, B)." figureDoi="10.3897/zookeys.1059.68385.figure3" httpUri="https://binary.pensoft.net/fig/584116" pageId="0" pageNumber="1">3A</figureCitation>
). Sinistral, large, solid, rather flat, five whorls, in chestnut and darker near aperture. Suture impressed. Protoconch 11/4 whorls. After ~ 31/2 whorls, growth lines broken into regularly arranged tubercles. Body whorl compressed from the last 1/4 to the last 1/8 whorls, completely covering corresponding part of penultimate whorl including the suture. After the last 1/8 whorls, body whorl becoming as broad as normal again. On the above compressed region, one shorter ventral depression and one longer apical depression present. Aperture semilunar, slightly descending. Peristome chestnut, thickened, expanded, and slightly reflexed. Umbilicus moderately broad; approximately 1/5 of shell major diameter. Protoconch visible through umbilicus.
</paragraph>
<paragraph pageId="0" pageNumber="1">
General anatomy (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. General anatomy A-D Stegodera angusticollis (Martens, 1875), HBUMM 08435 A lobe (arrowed) on the left side of mantle edge B lobe (arrowed) on the right side of mantle edge C internal surface of head region D jaw E-H Pseudostegodera qiului gen. et sp. nov., holotype IZCAS TM 206978 E jaw F internal surface of head region G lobe (arrowed) on the left side of mantle edge H lobe (arrowed) on the right side of mantle edge." figureDoi="10.3897/zookeys.1059.68385.figure4" httpUri="https://binary.pensoft.net/fig/584117" pageId="0" pageNumber="1">4A-D</figureCitation>
). Head wart absent. On internal body wall of head region between ommatophore insertions with tiny pits rather than glands (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. General anatomy A-D Stegodera angusticollis (Martens, 1875), HBUMM 08435 A lobe (arrowed) on the left side of mantle edge B lobe (arrowed) on the right side of mantle edge C internal surface of head region D jaw E-H Pseudostegodera qiului gen. et sp. nov., holotype IZCAS TM 206978 E jaw F internal surface of head region G lobe (arrowed) on the left side of mantle edge H lobe (arrowed) on the right side of mantle edge." figureDoi="10.3897/zookeys.1059.68385.figure4" httpUri="https://binary.pensoft.net/fig/584117" pageId="0" pageNumber="1">4C</figureCitation>
). Each side of mantle edge with a leaf-shaped appendage (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. General anatomy A-D Stegodera angusticollis (Martens, 1875), HBUMM 08435 A lobe (arrowed) on the left side of mantle edge B lobe (arrowed) on the right side of mantle edge C internal surface of head region D jaw E-H Pseudostegodera qiului gen. et sp. nov., holotype IZCAS TM 206978 E jaw F internal surface of head region G lobe (arrowed) on the left side of mantle edge H lobe (arrowed) on the right side of mantle edge." figureDoi="10.3897/zookeys.1059.68385.figure4" httpUri="https://binary.pensoft.net/fig/584117" pageId="0" pageNumber="1">4A, B</figureCitation>
). Body reddish brown, central dorsa with light longitudinal stripes. Sole dirty white. Jaw arcuate; with 12 more or less projecting ribs (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. General anatomy A-D Stegodera angusticollis (Martens, 1875), HBUMM 08435 A lobe (arrowed) on the left side of mantle edge B lobe (arrowed) on the right side of mantle edge C internal surface of head region D jaw E-H Pseudostegodera qiului gen. et sp. nov., holotype IZCAS TM 206978 E jaw F internal surface of head region G lobe (arrowed) on the left side of mantle edge H lobe (arrowed) on the right side of mantle edge." figureDoi="10.3897/zookeys.1059.68385.figure4" httpUri="https://binary.pensoft.net/fig/584117" pageId="0" pageNumber="1">4D</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Genitalia (Figs
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A general view B male part, showing the vas deferens connecting epiphallus (arrowed) C female part, showing the dart sac, accessory sac, and mucous glands." figureDoi="10.3897/zookeys.1059.68385.figure5" httpUri="https://binary.pensoft.net/fig/584118" pageId="0" pageNumber="1">5A-C</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A exposed dart sac apparatus B magnified partial penis and partial epiphallus. Arrow showing a very thin membranous sac wrapping distal half of penis with distal end connecting basal penial retractor muscle." figureDoi="10.3897/zookeys.1059.68385.figure6" httpUri="https://binary.pensoft.net/fig/584119" pageId="0" pageNumber="1">6A, B</figureCitation>
). Penis sheath present but thin and very short. Penis medially slightly thickened, moderately long, ~ 8.8 mm, externally simple. Inside penis with several thin longitudinal pilasters. Epiphallus slightly longer than penis. A very thin membranous sac wrapping distal half of penis with distal end connecting basal penial retractor muscle (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A exposed dart sac apparatus B magnified partial penis and partial epiphallus. Arrow showing a very thin membranous sac wrapping distal half of penis with distal end connecting basal penial retractor muscle." figureDoi="10.3897/zookeys.1059.68385.figure6" httpUri="https://binary.pensoft.net/fig/584119" pageId="0" pageNumber="1">6B</figureCitation>
). Flagellum cylindrical, tapering. Vas deferens thin throughout. Dart sac apparatus distally inserting on vagina. Dart sac small. Dart not observed. Membranous sac surrounding terminal genitalia absent. Accessory sac relatively large. Tubes of mucous glands more than twenty, neatly inserting in a single row on distal and same side of dart sac on accessory sac (Figs
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A general view B male part, showing the vas deferens connecting epiphallus (arrowed) C female part, showing the dart sac, accessory sac, and mucous glands." figureDoi="10.3897/zookeys.1059.68385.figure5" httpUri="https://binary.pensoft.net/fig/584118" pageId="0" pageNumber="1">5C</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A exposed dart sac apparatus B magnified partial penis and partial epiphallus. Arrow showing a very thin membranous sac wrapping distal half of penis with distal end connecting basal penial retractor muscle." figureDoi="10.3897/zookeys.1059.68385.figure6" httpUri="https://binary.pensoft.net/fig/584119" pageId="0" pageNumber="1">6B</figureCitation>
). From proximal to distal accessory sac, mucous gland tubes increased gradually in length (Figs
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A general view B male part, showing the vas deferens connecting epiphallus (arrowed) C female part, showing the dart sac, accessory sac, and mucous glands." figureDoi="10.3897/zookeys.1059.68385.figure5" httpUri="https://binary.pensoft.net/fig/584118" pageId="0" pageNumber="1">5C</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A exposed dart sac apparatus B magnified partial penis and partial epiphallus. Arrow showing a very thin membranous sac wrapping distal half of penis with distal end connecting basal penial retractor muscle." figureDoi="10.3897/zookeys.1059.68385.figure6" httpUri="https://binary.pensoft.net/fig/584119" pageId="0" pageNumber="1">6B</figureCitation>
). Papilla distally leading to mucous glands on inner wall of accessory sac integrated into one long thick spongious pilaster on the side of mucous gland tube insertion (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A exposed dart sac apparatus B magnified partial penis and partial epiphallus. Arrow showing a very thin membranous sac wrapping distal half of penis with distal end connecting basal penial retractor muscle." figureDoi="10.3897/zookeys.1059.68385.figure6" httpUri="https://binary.pensoft.net/fig/584119" pageId="0" pageNumber="1">6A</figureCitation>
). Bursa copulatrix ball-shaped.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="ecology">
<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="1">This species was observed under litter layer, on nearby rocks and crevices.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Central China: Hubei, Jiangxi (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution of Stegodera and Pseudostegodera qiului gen. et sp. nov. Triangle: new locality of Stegodera angusticollis (Martens, 1875); dashed range: area of Stegodera angusticollis; circle: type locality of Pseudostegodera qiului gen. et sp. nov." figureDoi="10.3897/zookeys.1059.68385.figure2" httpUri="https://binary.pensoft.net/fig/584115" pageId="0" pageNumber="1">2</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1059.68385.figure2" httpUri="https://binary.pensoft.net/fig/584115" pageId="0" pageNumber="1" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 2.</emphasis>
Distribution of
<taxonomicName authorityName="Martens" authorityYear="1876" class="Gastropoda" family="Camaenidae" genus="Stegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegodera" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Stegodera</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Chen" authorityYear="2021" class="Gastropoda" family="Camaenidae" genus="Pseudostegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudostegodera qiului" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="qiului">
<emphasis italics="true" pageId="0" pageNumber="1">Pseudostegodera qiului</emphasis>
</taxonomicName>
gen. et sp. nov. Triangle: new locality of
<taxonomicName baseAuthorityName="Martens" baseAuthorityYear="1875" class="Gastropoda" family="Camaenidae" genus="Stegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegodera angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis">
<emphasis italics="true" pageId="0" pageNumber="1">Stegodera angusticollis</emphasis>
</taxonomicName>
(Martens, 1875); dashed range: area of
<taxonomicName baseAuthorityName="Martens" baseAuthorityYear="1875" class="Gastropoda" family="Camaenidae" genus="Stegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegodera angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis">
<emphasis italics="true" pageId="0" pageNumber="1">Stegodera angusticollis</emphasis>
</taxonomicName>
; circle: type locality of
<taxonomicName authorityName="Chen" authorityYear="2021" class="Gastropoda" family="Camaenidae" genus="Pseudostegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudostegodera qiului" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="qiului">
<emphasis italics="true" pageId="0" pageNumber="1">Pseudostegodera qiului</emphasis>
</taxonomicName>
gen. et sp. nov.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
The peristome of
<taxonomicName baseAuthorityName="Martens" baseAuthorityYear="1875" class="Gastropoda" family="Camaenidae" genus="Stegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegodera angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis">
<emphasis italics="true" pageId="0" pageNumber="1">Stegodera angusticollis</emphasis>
</taxonomicName>
is in chestnut in fresh specimens, but in most museum specimens this color gets faint to white. In
<taxonomicName baseAuthorityName="Martens" baseAuthorityYear="1875" class="Gastropoda" family="Camaenidae" genus="Stegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegodera angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis">
<emphasis italics="true" pageId="0" pageNumber="1">Stegodera angusticollis</emphasis>
</taxonomicName>
, the compressed part of body whorl usually completely cover the penultimate whorl, while in
<taxonomicName authorityName="Chen" authorityYear="2021" class="Gastropoda" family="Camaenidae" genus="Pseudostegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudostegodera qiului" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="qiului">
<emphasis italics="true" pageId="0" pageNumber="1">Pseudostegodera qiului</emphasis>
</taxonomicName>
gen. et sp. nov. the corresponding part is not so compressed and the compressed part of body whorl can hardly cover the penultimate whorl.
</paragraph>
<caption doi="10.3897/zookeys.1059.68385.figure3" httpUri="https://binary.pensoft.net/fig/584116" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
Shells
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName baseAuthorityName="Martens" baseAuthorityYear="1875" class="Gastropoda" family="Camaenidae" genus="Stegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegodera angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis">
<emphasis italics="true" pageId="0" pageNumber="1">Stegodera angusticollis</emphasis>
</taxonomicName>
Martens, 1876, HBUMM08435
<emphasis bold="true" pageId="0" pageNumber="1">B-D</emphasis>
<taxonomicName authorityName="Chen" authorityYear="2021" class="Gastropoda" family="Camaenidae" genus="Pseudostegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudostegodera qiului" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="qiului">
<emphasis italics="true" pageId="0" pageNumber="1">Pseudostegodera qiului</emphasis>
</taxonomicName>
gen. et sp. nov., holotype IZCAS TM206978
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
p rotoconch, magnified
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
shell surface, magnified. Scale bar: 5 mm (
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1059.68385.figure4" httpUri="https://binary.pensoft.net/fig/584117" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
General anatomy
<emphasis bold="true" pageId="0" pageNumber="1">A-D</emphasis>
<taxonomicName baseAuthorityName="Martens" baseAuthorityYear="1875" class="Gastropoda" family="Camaenidae" genus="Stegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegodera angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis">
<emphasis italics="true" pageId="0" pageNumber="1">Stegodera angusticollis</emphasis>
</taxonomicName>
(Martens, 1875), HBUMM08435
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
lobe (arrowed) on the left side of mantle edge
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
lobe (arrowed) on the right side of mantle edge
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
internal surface of head region
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
jaw
<emphasis bold="true" pageId="0" pageNumber="1">E-H</emphasis>
<taxonomicName authorityName="Chen" authorityYear="2021" class="Gastropoda" family="Camaenidae" genus="Pseudostegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudostegodera qiului" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="qiului">
<emphasis italics="true" pageId="0" pageNumber="1">Pseudostegodera qiului</emphasis>
</taxonomicName>
gen. et sp. nov., holotype IZCAS TM206978
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
jaw
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
internal surface of head region
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
lobe (arrowed) on the left side of mantle edge
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
lobe (arrowed) on the right side of mantle edge.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
Some authors (
<bibRefCitation author="Chen, DN" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="1" refId="B4" refString="Chen, DN, Gao, JX, 1987. Economic Fauna of China (Terrestrial Molluscs). Science Press, Beijing" title="Economic Fauna of China (Terrestrial Molluscs)." year="1987">Chen and Gao 1987</bibRefCitation>
; Qian et al. 2008;
<bibRefCitation author="Qian, ZX" journalOrPublisher="Zhejiang People's Fine Arts Publishing House, Hangzhou" pageId="0" pageNumber="1" refId="B30" refString="Qian, ZX, Zhou, WC, 2014. Illustrated Handbook of Common Terrestrial Mollusks in China. Zhejiang People's Fine Arts Publishing House, Hangzhou" title="Illustrated Handbook of Common Terrestrial Mollusks in China." year="2014">Qian and Zhou 2014</bibRefCitation>
) recorded this species in Jiangsu, Zhejiang and Anhui, but voucher specimens from these provinces were neither recorded therein nor being found in the mollusk collection deposited in IZCAS where some of them once worked. Therefore, the accuracy of these records needs to be further confirmed.
</paragraph>
<caption doi="10.3897/zookeys.1059.68385.figure5" httpUri="https://binary.pensoft.net/fig/584118" pageId="0" pageNumber="1" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 5.</emphasis>
Genitalia of
<taxonomicName baseAuthorityName="Martens" baseAuthorityYear="1875" class="Gastropoda" family="Camaenidae" genus="Stegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegodera angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis">
<emphasis italics="true" pageId="0" pageNumber="1">Stegodera angusticollis</emphasis>
</taxonomicName>
(Martens, 1875), HBUMM08435
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
general view
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
male part, showing the vas deferens connecting epiphallus (arrowed)
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
female part, showing the dart sac, accessory sac, and mucous glands.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1059.68385.figure6" httpUri="https://binary.pensoft.net/fig/584119" pageId="0" pageNumber="1" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 6.</emphasis>
Genitalia of
<taxonomicName baseAuthorityName="Martens" baseAuthorityYear="1875" class="Gastropoda" family="Camaenidae" genus="Stegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegodera angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis">
<emphasis italics="true" pageId="0" pageNumber="1">Stegodera angusticollis</emphasis>
</taxonomicName>
(Martens, 1875), HBUMM08435
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
exposed dart sac apparatus
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
magnified partial penis and partial epiphallus. Arrow showing a very thin membranous sac wrapping distal half of penis with distal end connecting basal penial retractor muscle.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

60
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<taxonomicName id="6C5D7278A8C86B294608A4147B645383" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181733" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole hamtoni Wilson" lsidName-HNS="Pheidole hamtoni Wilson" order="Hymenoptera" pageNumber="296" phylum="Arthropoda" rank="species" species="hamtoni">Pheidole hamtoni</taxonomicName>
<taxonomicNameLabel id="4B962094E967D554908AB85C3A26C49E" rank="species">new species</taxonomicNameLabel>
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<paragraph id="5DD2449DDA5CBD7AF4DB37D7253E5732" pageNumber="296">Types Mus. Comp. Zool. Harvard.</paragraph>
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<subSubSection id="B5ACFE1C61DE84BA7D9A79DC51861B80" type="etymology">
<paragraph id="7461E472CFF6D7E7A5BE5BC172C61CDA" pageNumber="296">Etymology Named after the collector, Robert J. Hamton.</paragraph>
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<paragraph id="A7159945C514450B6650CE3BE8927227" pageNumber="296">
Diagnosis A member of the
<taxonomicName id="013D0275E6B3AC340BAAF041CE99415B" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33735" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole fallax Mayr" lsidName-HNS="Pheidole fallax Mayr" order="Hymenoptera" pageNumber="296" phylum="Arthropoda" rank="species" species="fallax">fallax</taxonomicName>
group, close to
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but differing from it and other species of the group by the following combination of traits. Both major and minor brownish yellow.
</paragraph>
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<subSubSection id="309AF5AAAE07F42B329EF350B1EA01FE" type="description">
<paragraph id="2A1439531CFC53E577664F7A0538EAD9" pageNumber="296">Major: head capsule 1.14X longer than broad; hypostomum with 5 teeth; propodeal spine greatly reduced, to a rudimentary denticle; postpetiolar node from above short and bell-shaped; pilosity erect and very dense; rugoreticulum present between each eye and the nearby antennal fossa; carinulae mesad to the eye extend almost all the way to the occipital border. Minor: propodeal spines reduced to denticles; occiput narrowed slightly, with a nuchal collar. Measurements (mm) Holotype major: HW 1.32, HL 1.50, SL 1.22, EL 0.22, PW 0.74. Paratype minor: HW 0.64, HL 0.78, SL 1.02, EL 0.16, PW 0.44.</paragraph>
<paragraph id="945527998C154F6FDDB887CDE6F0D3A1" pageNumber="296">Color Major: body concolorous brownish yellow, except for gaster, which is a shade darker. Minor: concolorous brownish yellow.</paragraph>
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<subSubSection id="CAB0BC8D4A7F585BB3803FF6AA807850" type="distribution">
<paragraph id="BA558DF37318E7CEA088EF1DDD8E4F98" pageNumber="296">Range Known from the type locality and from 8 km northwest of Tequila, Jalisco, 1220 m (R. J. Hamton).</paragraph>
</subSubSection>
<subSubSection id="53309E84465FE70CC21D50B4F6CF9815" type="biology_ecology">
<paragraph id="2482CC99287285A126EAF937A56263B0" pageNumber="296">Biology Unknown.</paragraph>
</subSubSection>
<subSubSection id="9E467D374C042E2C4421CFCDE9AA7219" type="materials_examined">
<paragraph id="B058648053608AD53CDE6DC560B5EB6F" pageNumber="296">Figure Upper: holotype, major. Lower: paratype, minor. MEXICO: 10 km northwest of Ixtlan, Nayarit (Robert J. Hamton). Scale bars = 1 mm.</paragraph>
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