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<document id="F452BED38D1A06E9F7792C156FF6C21F" ID-DOI="10.1649/0010-065X-74.3.542" ID-ISSN="1938-4394" ID-Zenodo-Dep="5329407" ID-ZooBank="urn:lsid:zoobank.org:pub:526D70F6-D98D-4ACE-8D29-108D7B13B31D" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630163227317" checkinUser="felipe" docAuthor="Löbl, Ivan &amp; Smetana, Aleš" docDate="2020" docId="B0611772FFC2FF84FD78FB3B57D45119" docLanguage="en" docName="0010-065X-74.3.542.pdf" docOrigin="The Coleopterists Bulletin 74 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-74.3.542" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Praehelichus Lobl and Smetana 2020, new genus" docType="treatment" docUuid="444D6929-8323-4092-A82A-14A4BC0469A1" docUuidSource="ZooBank" docVersion="4" lastPageNumber="543" masterDocId="4C586F0AFFC3FF86FFB9FF8F5636551F" masterDocTitle="On the Names of Three Genera of Dryopidae and Elateridae (Coleoptera)" masterLastPageNumber="543" masterPageNumber="542" pageNumber="542" updateTime="1699169883525" updateUser="plazi">
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<mods:title id="5272192FCDEA7465B8AEF17E18851DBB">On the Names of Three Genera of Dryopidae and Elateridae (Coleoptera)</mods:title>
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<mods:namePart id="C920C5EF0CFCA682C453C816F49C3B43">Löbl, Ivan</mods:namePart>
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<mods:namePart id="96476022E8C610644521BC89DF4F0D19">Smetana, Aleš</mods:namePart>
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<mods:title id="80AEF2EE2C13372CBF7619FDD6501BBE">The Coleopterists Bulletin</mods:title>
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<mods:date id="EDAF631176468B271AE6F8C6D51ACE43">2020</mods:date>
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<mods:number id="918A4857BAA01D00E14533210965CC3B">2020-09-09</mods:number>
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<mods:number id="E57DDCB5C8191AEC53DA7FD293A4DEB2">74</mods:number>
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<emphasis id="0ABC7A76FFC2FF87FD78FB3B52B651D5" bold="true" box="[705,1152,1204,1226]" pageId="1" pageNumber="542">
<taxonomicName id="FFC8DDE7FFC2FF87FD78FB3B523C51D5" ID-CoL="6VCJ" authority="Lobl and Smetana" authorityName="Lobl and Smetana" authorityYear="2020" box="[705,1034,1204,1226]" class="Insecta" family="Dryopidae" genus="Praehelichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="542" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="0ABC7A76FFC2FF87FD78FB3B557551D5" bold="true" box="[705,835,1204,1226]" italics="true" pageId="1" pageNumber="542">Praehelichus</emphasis>
Löbl and Smetana
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,
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zoobank.org/
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<paragraph id="3877A664FFC2FF84FD7CFA9B547B5439" blockId="1.[685,1172,1299,1730]" lastBlockId="2.[173,659,193,294]" lastPageId="2" lastPageNumber="543" pageId="1" pageNumber="542">
Similarly to the above case,
<taxonomicName id="FFC8DDE7FFC2FF87FC71FA9C54EB505B" authority="Bollow, 1940" authorityName="Bollow" authorityYear="1940" class="Insecta" family="Dryopidae" genus="Praehelichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="542" phylum="Arthropoda" rank="genus">
<emphasis id="0ABC7A76FFC2FF87FC71FA9C52735036" box="[968,1093,1299,1321]" italics="true" pageId="1" pageNumber="542">Praehelichus</emphasis>
<bibRefCitation id="5C59DB95FFC2FF87FBF0FA9B54EB505B" author="Bollow, H." pageId="1" pageNumber="542" pagination="24 - 71" refId="ref1046" refString="Bollow, H. 1940. Monographie der palaearktischen Dryopidae mit Berucksichtigung der eventuell transgredieren den Arten (Col.). Mitteilungen der Munchner Entomologischen Gesellschaft 30: 24 - 71." type="journal article" year="1940">Bollow, 1940</bibRefCitation>
</taxonomicName>
was described as a new subgenus of
<taxonomicName id="FFC8DDE7FFC2FF87FB86FAA152A5505B" authorityName="Erichson" authorityYear="1847" box="[1087,1171,1326,1348]" class="Insecta" family="Dryopidae" genus="Helichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="542" phylum="Arthropoda" rank="genus">
<emphasis id="0ABC7A76FFC2FF87FB86FAA152A5505B" box="[1087,1171,1326,1348]" italics="true" pageId="1" pageNumber="542">Helichus</emphasis>
</taxonomicName>
to include four species,
<taxonomicName id="FFC8DDE7FFC2FF87FC26FAC654D15065" authority="(Solsky, 1876)" baseAuthorityName="Solsky" baseAuthorityYear="1876" class="Insecta" family="Dryopidae" genus="Helichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="542" phylum="Arthropoda" rank="species" species="longus">
<emphasis id="0ABC7A76FFC2FF87FC26FAC6520A5040" box="[927,1084,1353,1375]" italics="true" pageId="1" pageNumber="542">Helichus longus</emphasis>
(Solsky, 1876)
</taxonomicName>
,
<taxonomicName id="FFC8DDE7FFC2FF87FD48FAEA52005065" authority="Fairmaire, 1888" authorityName="Fairmaire" authorityYear="1888" box="[753,1078,1381,1403]" class="Insecta" family="Dryopidae" genus="Helichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="542" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="0ABC7A76FFC2FF87FD48FAEA55A05065" box="[753,918,1381,1403]" italics="true" pageId="1" pageNumber="542">Helichus sinensis</emphasis>
Fairmaire, 1888
</taxonomicName>
,
<taxonomicName id="FFC8DDE7FFC2FF87FB86FAEA523E5089" authority="(C. O. Waterhouse, 1881)" baseAuthorityName="C. O. Waterhouse" baseAuthorityYear="1881" class="Insecta" family="Dryopidae" genus="Helichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="542" phylum="Arthropoda" rank="species" species="sericatus">
<emphasis id="0ABC7A76FFC2FF87FB86FAEA55345089" italics="true" pageId="1" pageNumber="542">Helichus sericatus</emphasis>
(C. O. Waterhouse, 1881)
</taxonomicName>
, and
<taxonomicName id="FFC8DDE7FFC2FF87FB86FA0F55B350AE" authority="(Zaitzev, 1908)" baseAuthorityName="Zaitzev" baseAuthorityYear="1908" class="Insecta" family="Dryopidae" genus="Helichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="542" phylum="Arthropoda" rank="species" species="solskyi">
<emphasis id="0ABC7A76FFC2FF87FB86FA0F54DB50AE" italics="true" pageId="1" pageNumber="542">Helichus solskyi</emphasis>
(Zaitzev, 1908)
</taxonomicName>
. Subsequently,
<taxonomicName id="FFC8DDE7FFC2FF87FBAEFA1452A250AE" authorityName="Bollow" authorityYear="1940" box="[1047,1172,1435,1457]" class="Insecta" family="Dryopidae" genus="Praehelichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="542" phylum="Arthropoda" rank="genus">
<emphasis id="0ABC7A76FFC2FF87FBAEFA1452A250AE" box="[1047,1172,1435,1457]" italics="true" pageId="1" pageNumber="542">Praehelichus</emphasis>
</taxonomicName>
was raised to genus rank and was used as valid, although Bollow and subsequent workers failed to designate the
<typeStatus id="E77318C6FFC2FF87FC87FA615551531C" box="[830,871,1518,1539]" pageId="1" pageNumber="542">type</typeStatus>
species. According to
<bibRefCitation id="5C59DB95FFC2FF87FBE5FA6154D15301" author="ICZN &amp; The International Commission on Zoological Nomenclature" pageId="1" pageNumber="542" refId="ref1079" refString="ICZN (The International Commission on Zoological Nomenclature). 1999. International Code of Zoological Nomenclature. Fourth Edition. The International Trust of Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN (1999</bibRefCitation>
, Art. 13.3),
<taxonomicName id="FFC8DDE7FFC2FF87FCCCF98752715301" authority="Bollow" authorityName="Bollow" authorityYear="1940" box="[885,1095,1544,1566]" class="Insecta" family="Dryopidae" genus="Praehelichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="542" phylum="Arthropoda" rank="genus">
<emphasis id="0ABC7A76FFC2FF87FCCCF98755C45301" box="[885,1010,1544,1566]" italics="true" pageId="1" pageNumber="542">Praehelichus</emphasis>
Bollow
</taxonomicName>
is unavailable. Löbl and
<taxonomicName id="FFC8DDE7FFC2FF87FCD2F9AB52115326" authority="Löbl &amp; Smetana, 2020" authorityName="Löbl &amp; Smetana" authorityYear="2020" box="[875,1063,1572,1593]" genus="Smetana" pageId="1" pageNumber="543" rank="genus" status="gen. nov.">Smetana (2006: 61)</taxonomicName>
designated
<taxonomicName id="FFC8DDE7FFC2FF87FD14F9CF5279534A" authority="C. O. Waterhouse, 1881" authorityName="C. O. Waterhouse" authorityYear="1881" box="[685,1103,1600,1621]" class="Insecta" family="Dryopidae" genus="Dryops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="542" phylum="Arthropoda" rank="species" species="sericatus">
<emphasis id="0ABC7A76FFC2FF87FD14F9CF5565534A" box="[685,851,1600,1621]" italics="true" pageId="1" pageNumber="542">Dryops sericatus</emphasis>
C. O. Waterhouse, 1881
</taxonomicName>
as the
<typeStatus id="E77318C6FFC2FF87FD14F9D454E0536F" box="[685,726,1627,1648]" pageId="1" pageNumber="542">type</typeStatus>
species of
<taxonomicName id="FFC8DDE7FFC2FF87FCFFF9D555F5536F" authorityName="Bollow" authorityYear="1940" box="[838,963,1626,1648]" class="Insecta" family="Dryopidae" genus="Praehelichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="542" phylum="Arthropoda" rank="genus">
<emphasis id="0ABC7A76FFC2FF87FCFFF9D555F5536F" box="[838,963,1626,1648]" italics="true" pageId="1" pageNumber="542">Praehelichus</emphasis>
</taxonomicName>
. As they had not explicitly indicated that the taxon was new, this name remained unavailable (
<bibRefCitation id="5C59DB95FFC2FF87FC34F91D523753B8" author="ICZN &amp; The International Commission on Zoological Nomenclature" box="[909,1025,1682,1703]" pageId="1" pageNumber="542" refId="ref1079" refString="ICZN (The International Commission on Zoological Nomenclature). 1999. International Code of Zoological Nomenclature. Fourth Edition. The International Trust of Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
, Art. 16.1). In order to make the name available, the new genus
<taxonomicName id="FFC8DDE7FFC1FF84FF14FF4E571C55C8" authorityName="Bollow" authorityYear="1940" box="[173,298,193,215]" class="Insecta" family="Dryopidae" genus="Praehelichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="543" phylum="Arthropoda" rank="genus">
<emphasis id="0ABC7A76FFC1FF84FF14FF4E571C55C8" box="[173,298,193,215]" italics="true" pageId="2" pageNumber="543">Praehelichus</emphasis>
</taxonomicName>
is here established, with
<taxonomicName id="FFC8DDE7FFC1FF84FDF4FF4D57DA55EE" authority="C. O. Waterhouse, 1881" authorityName="C. O. Waterhouse" authorityYear="1881" class="Insecta" family="Dryopidae" genus="Dryops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="543" phylum="Arthropoda" rank="species" species="sericatus">
<emphasis id="0ABC7A76FFC1FF84FDF4FF4D573455EE" italics="true" pageId="2" pageNumber="543">Dryops sericatus</emphasis>
C. O. Waterhouse, 1881
</taxonomicName>
designated as the
<typeStatus id="E77318C6FFC1FF84FF14FF7856E05413" box="[173,214,247,268]" pageId="2" pageNumber="543">type</typeStatus>
species. The original description given by
<bibRefCitation id="5C59DB95FFC1FF84FF14FE9E57565439" author="Bollow, H." box="[173,352,273,294]" pageId="2" pageNumber="543" pagination="24 - 71" refId="ref1046" refString="Bollow, H. 1940. Monographie der palaearktischen Dryopidae mit Berucksichtigung der eventuell transgredieren den Arten (Col.). Mitteilungen der Munchner Entomologischen Gesellschaft 30: 24 - 71." type="journal article" year="1940">Bollow (1940: 36)</bibRefCitation>
characterized the taxon.
</paragraph>
<paragraph id="3877A664FFC1FF84FEDAFEDE57EA5478" blockId="2.[355,476,337,360]" box="[355,476,337,360]" pageId="2" pageNumber="543">
<taxonomicName id="FFC8DDE7FFC1FF84FEDAFEDE57EA5478" authorityName="Leach" authorityYear="1815" box="[355,476,337,360]" class="Insecta" family="Elateridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="543" phylum="Arthropoda" rank="family">ELATERIDAE</taxonomicName>
</paragraph>
<paragraph id="3877A664FFC1FF84FF70FEF2544954D7" blockId="2.[193,639,381,456]" pageId="2" pageNumber="543">
<emphasis id="0ABC7A76FFC1FF84FF70FEF25440548C" bold="true" box="[201,630,381,403]" pageId="2" pageNumber="543">
<taxonomicName id="FFC8DDE7FFC1FF84FF70FEF25436548C" authority="Lobl and Smetana" authorityName="Lobl and Smetana" authorityYear="2020" box="[201,512,381,403]" class="Insecta" family="Elateridae" genus="Paulusiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="543" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="0ABC7A76FFC1FF84FF70FEF2570F548C" bold="true" box="[201,313,381,403]" italics="true" pageId="2" pageNumber="543">Paulusiella</emphasis>
Löbl and Smetana
</taxonomicName>
,
<taxonomicNameLabel id="118FC70DFFC1FF84FDB5FEF15440548C" box="[524,630,382,403]" pageId="2" pageNumber="543" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
zoobank.org/
<uri id="4C59AA66FFC1FF84FED5FE17544954D7" pageId="2" pageNumber="543">
urn:lsid:zoobank.org:act:
<uuid id="4C6E9CB1FFC1FF84FF78FE3C544954D7" box="[193,639,435,456]" pageId="2" pageNumber="543">085F6DFA-9767-4A66-A84E-D7805FF82855</uuid>
</uri>
</paragraph>
<paragraph id="3877A664FFC1FF84FF7CFE52542256A9" blockId="2.[173,660,477,1030]" pageId="2" pageNumber="543">
<bibRefCitation id="5C59DB95FFC1FF84FF7CFE52574E54ED" author="Mandl, K." box="[197,376,477,498]" pageId="2" pageNumber="543" pagination="635 - 645" refId="ref1463" refString="Mandl, K. 1974. Eine neue Gattung, drei neue Arten und eine neue Unterart aus der Familie Karumiidae (Dascilloidea, Col.). Verhandlungen der Naturforschenden Gesellschaft Basel 84: 635 - 645." type="journal article" year="1974">Mandl (1974: 640)</bibRefCitation>
stated he was establishing the new genus
<taxonomicName id="FFC8DDE7FFC1FF84FEA1FE7857B25712" authorityName="Lobl and Smetana" authorityYear="2020" box="[280,388,503,525]" class="Insecta" family="Elateridae" genus="Paulusiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="543" phylum="Arthropoda" rank="genus">
<emphasis id="0ABC7A76FFC1FF84FEA1FE7857B25712" box="[280,388,503,525]" italics="true" pageId="2" pageNumber="543">Paulusiella</emphasis>
</taxonomicName>
for
<taxonomicName id="FFC8DDE7FFC1FF84FE15FE78571A5738" authority="Paulus, 1972" authorityName="Paulus" authorityYear="1972" class="Insecta" family="Dascillidae" genus="Escalerina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="543" phylum="Arthropoda" rank="species" species="serraticornis">
<emphasis id="0ABC7A76FFC1FF84FE15FE7854A55712" box="[428,659,503,525]" italics="true" pageId="2" pageNumber="543">Escalerina serraticornis</emphasis>
Paulus, 1972
</taxonomicName>
and a new species,
<taxonomicName id="FFC8DDE7FFC1FF84FE43FD9D5796575D" authority="Mandl, 1974" authorityName="Mandl" authorityYear="1974" class="Insecta" family="Elateridae" genus="Paulusiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="543" phylum="Arthropoda" rank="species" species="fossulatipennis">
<emphasis id="0ABC7A76FFC1FF84FE43FD9D5729575D" italics="true" pageId="2" pageNumber="543">Paulusiella fossulatipennis</emphasis>
<bibRefCitation id="5C59DB95FFC1FF84FE9DFDA25796575D" author="Mandl, K." box="[292,416,557,578]" pageId="2" pageNumber="543" pagination="635 - 645" refId="ref1463" refString="Mandl, K. 1974. Eine neue Gattung, drei neue Arten und eine neue Unterart aus der Familie Karumiidae (Dascilloidea, Col.). Verhandlungen der Naturforschenden Gesellschaft Basel 84: 635 - 645." type="journal article" year="1974">Mandl, 1974</bibRefCitation>
</taxonomicName>
. In the same paper he also described
<taxonomicName id="FFC8DDE7FFC1FF84FEB6FDC854785743" authority="Mandl, 1974" authorityName="Mandl" authorityYear="1974" box="[271,590,583,605]" class="Insecta" family="Elateridae" genus="Paulusiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="543" phylum="Arthropoda" rank="species" species="richteri">
<emphasis id="0ABC7A76FFC1FF84FEB6FDC857FC5742" box="[271,458,583,605]" italics="true" pageId="2" pageNumber="543">Paulusiella richteri</emphasis>
<bibRefCitation id="5C59DB95FFC1FF84FE6BFDC854785743" author="Mandl, K." box="[466,590,583,604]" pageId="2" pageNumber="543" pagination="635 - 645" refId="ref1463" refString="Mandl, K. 1974. Eine neue Gattung, drei neue Arten und eine neue Unterart aus der Familie Karumiidae (Dascilloidea, Col.). Verhandlungen der Naturforschenden Gesellschaft Basel 84: 635 - 645." type="journal article" year="1974">Mandl, 1974</bibRefCitation>
</taxonomicName>
with a subspecies,
<taxonomicName id="FFC8DDE7FFC1FF84FE99FDEE56E9578D" authority="Mandl, 1974" authorityName="Mandl" authorityYear="1974" class="Insecta" family="Elateridae" genus="Paulusiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="543" phylum="Arthropoda" rank="subSpecies" species="richteri" subSpecies="acuticollis">
<emphasis id="0ABC7A76FFC1FF84FE99FDEE54705768" box="[288,582,609,631]" italics="true" pageId="2" pageNumber="543">Paulusiella richteri acuticollis</emphasis>
<bibRefCitation id="5C59DB95FFC1FF84FDF6FDED56E9578D" author="Mandl, K." pageId="2" pageNumber="543" pagination="635 - 645" refId="ref1463" refString="Mandl, K. 1974. Eine neue Gattung, drei neue Arten und eine neue Unterart aus der Familie Karumiidae (Dascilloidea, Col.). Verhandlungen der Naturforschenden Gesellschaft Basel 84: 635 - 645." type="journal article" year="1974">Mandl, 1974</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="FFC8DDE7FFC1FF84FEA3FDF35457578D" authority="Mandl, 1974" authorityName="Mandl" authorityYear="1974" box="[282,609,636,658]" class="Insecta" family="Elateridae" genus="Paulusiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="543" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="0ABC7A76FFC1FF84FEA3FDF357E2578D" box="[282,468,636,658]" italics="true" pageId="2" pageNumber="543">Paulusiella pallida</emphasis>
<bibRefCitation id="5C59DB95FFC1FF84FE66FDF25457578D" author="Mandl, K." box="[479,609,637,658]" pageId="2" pageNumber="543" pagination="635 - 645" refId="ref1463" refString="Mandl, K. 1974. Eine neue Gattung, drei neue Arten und eine neue Unterart aus der Familie Karumiidae (Dascilloidea, Col.). Verhandlungen der Naturforschenden Gesellschaft Basel 84: 635 - 645." type="journal article" year="1974">Mandl, 1974</bibRefCitation>
</taxonomicName>
. The
<typeStatus id="E77318C6FFC1FF84FF14FD1856E057B3" box="[173,214,663,684]" pageId="2" pageNumber="543">type</typeStatus>
species was not designated and
<taxonomicName id="FFC8DDE7FFC1FF84FD9EFD1956DD57D8" authority="Mandl" authorityName="Mandl" authorityYear="1974" class="Insecta" family="Elateridae" genus="Paulusiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="543" phylum="Arthropoda" rank="genus">
<emphasis id="0ABC7A76FFC1FF84FD9EFD1954A557B3" box="[551,659,662,684]" italics="true" pageId="2" pageNumber="543">Paulusiella</emphasis>
Mandl
</taxonomicName>
is therefore, according to
<bibRefCitation id="5C59DB95FFC1FF84FE53FD3D545557D8" author="ICZN &amp; The International Commission on Zoological Nomenclature" box="[490,611,690,711]" pageId="2" pageNumber="543" refId="ref1079" refString="ICZN (The International Commission on Zoological Nomenclature). 1999. International Code of Zoological Nomenclature. Fourth Edition. The International Trust of Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN (1999</bibRefCitation>
, Art. 13.3), unavailable.
<bibRefCitation id="5C59DB95FFC1FF84FECFFD43542957FE" author="Lobl, I." box="[374,543,716,737]" pageId="2" pageNumber="543" refId="ref1333" refString="Lobl, I. 2007. New nomenclatorial and taxonomic acts, and comments. Elateridae: Cebrioninae [p. 32]. In: Catalogue of Palaearctic Coleoptera. Volume 4. Elateroidea - Derodontoidea - Lymexyloidea - Cleroidea - Cucujoidea (I. Lobl and A. Smetana, editors). Apollo Books, Stenstrup, 935 pp." type="book" year="2007">Löbl (2007: 32)</bibRefCitation>
designated
<taxonomicName id="FFC8DDE7FFC1FF84FF14FD69541C57E3" authority="Mandl, 1974" authorityName="Mandl" authorityYear="1974" box="[173,554,742,764]" class="Insecta" family="Elateridae" genus="Paulusiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="543" phylum="Arthropoda" rank="species" species="fossulatipennis">
<emphasis id="0ABC7A76FFC1FF84FF14FD69579A57E3" box="[173,428,742,764]" italics="true" pageId="2" pageNumber="543">Paulusiella fossulatipennis</emphasis>
<bibRefCitation id="5C59DB95FFC1FF84FE08FD68541C57E3" author="Mandl, K." box="[433,554,743,764]" pageId="2" pageNumber="543" pagination="635 - 645" refId="ref1463" refString="Mandl, K. 1974. Eine neue Gattung, drei neue Arten und eine neue Unterart aus der Familie Karumiidae (Dascilloidea, Col.). Verhandlungen der Naturforschenden Gesellschaft Basel 84: 635 - 645." type="journal article" year="1974">Mandl, 1974</bibRefCitation>
</taxonomicName>
as the
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species. As he had not explicitly indicated that the taxon was new, this name remained unavailable (
<bibRefCitation id="5C59DB95FFC1FF84FF0AFCB857125653" author="ICZN &amp; The International Commission on Zoological Nomenclature" box="[179,292,823,844]" pageId="2" pageNumber="543" refId="ref1079" refString="ICZN (The International Commission on Zoological Nomenclature). 1999. International Code of Zoological Nomenclature. Fourth Edition. The International Trust of Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
, Art. 16.1). In order to make the name available, the new genus
<taxonomicName id="FFC8DDE7FFC1FF84FE72FCDF54015679" authorityName="Mandl" authorityYear="1974" box="[459,567,848,870]" class="Insecta" family="Elateridae" genus="Paulusiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="543" phylum="Arthropoda" rank="genus">
<emphasis id="0ABC7A76FFC1FF84FE72FCDF54015679" box="[459,567,848,870]" italics="true" pageId="2" pageNumber="543">Paulusiella</emphasis>
</taxonomicName>
is here established, with
<taxonomicName id="FFC8DDE7FFC1FF84FEF6FCE456EA5684" authority="Mandl, 1974" authorityName="Mandl" authorityYear="1974" class="Insecta" family="Elateridae" genus="Paulusiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="543" phylum="Arthropoda" rank="species" species="fossulatipennis">
<emphasis id="0ABC7A76FFC1FF84FEF6FCE4547C569E" box="[335,586,875,897]" italics="true" pageId="2" pageNumber="543">Paulusiella fossulatipennis</emphasis>
<bibRefCitation id="5C59DB95FFC1FF84FDE9FCE356EA5684" author="Mandl, K." pageId="2" pageNumber="543" pagination="635 - 645" refId="ref1463" refString="Mandl, K. 1974. Eine neue Gattung, drei neue Arten und eine neue Unterart aus der Familie Karumiidae (Dascilloidea, Col.). Verhandlungen der Naturforschenden Gesellschaft Basel 84: 635 - 645." type="journal article" year="1974">Mandl, 1974</bibRefCitation>
</taxonomicName>
as the
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species. The description given by
<bibRefCitation id="5C59DB95FFC1FF84FF14FC2E571956A9" author="Mandl, K." box="[173,303,929,950]" pageId="2" pageNumber="543" pagination="635 - 645" refId="ref1463" refString="Mandl, K. 1974. Eine neue Gattung, drei neue Arten und eine neue Unterart aus der Familie Karumiidae (Dascilloidea, Col.). Verhandlungen der Naturforschenden Gesellschaft Basel 84: 635 - 645." type="journal article" year="1974">Mandl (1974)</bibRefCitation>
differentiates the taxon.
</paragraph>
<paragraph id="3877A664FFC1FF84FF7CFC3457D45119" blockId="2.[173,660,477,1030]" pageId="2" pageNumber="543">
The relationships of
<taxonomicName id="FFC8DDE7FFC1FF84FE3DFC3457C656CE" authorityName="Mandl" authorityYear="1974" box="[388,496,955,977]" class="Insecta" family="Elateridae" genus="Paulusiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="543" phylum="Arthropoda" rank="genus">
<emphasis id="0ABC7A76FFC1FF84FE3DFC3457C656CE" box="[388,496,955,977]" italics="true" pageId="2" pageNumber="543">Paulusiella</emphasis>
</taxonomicName>
are doubtful, and the genus was placed by Ivie and Barclay (2011) within
<taxonomicName id="FFC8DDE7FFC1FF84FF48FC7E57665119" authorityName="Leach" authorityYear="1815" box="[241,336,1009,1030]" class="Insecta" family="Elateridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="543" phylum="Arthropoda" rank="family">Elateridae</taxonomicName>
<emphasis id="0ABC7A76FFC1FF84FEE1FC7E57EA5119" box="[344,476,1008,1030]" italics="true" pageId="2" pageNumber="543">incertae sedis</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>

114
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<taxonomicName LSID="http://species-id.net/wiki/Torrenticola_dentifera" authority="Wiles, 1991" authorityName="Wiles" authorityYear="1991" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola dentifera" order="Trombidiformes" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="dentifera">
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dentifera Wiles, 1991
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Figs 2, 7B
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<paragraph pageId="3" pageNumber="24">
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola dentifera" order="Trombidiformes" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="dentifera">Torrenticola dentifera</taxonomicName>
Wiles, 1991: 43.
</paragraph>
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<subSubSection pageId="3" pageNumber="24" type="material examined">
<paragraph pageId="3" pageNumber="24">Material examined.</paragraph>
<paragraph pageId="3" pageNumber="24">
SOUTH KOREA: CR9 Ne myeon Mt, Naebyeansan NP, stream near Naebyeansan Info Center,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="35.640453">35°38'25.623&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="126.58143">126°34'53.1438&quot;E</geoCoordinate>
, 10.x.2012,
<normalizedToken originalValue="Pešić">Pesic</normalizedToken>
&amp; Choi 1/0/0 (mounted, NIBRIV0000268845).
</paragraph>
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<subSubSection pageId="3" pageNumber="24" type="description">
<paragraph pageId="3" pageNumber="24">Morphology.</paragraph>
<paragraph pageId="3" pageNumber="24">Male. General features. Idiosoma elongated; frontal platelets anteriorly bulging (Figs 2A, 7B); Cxgl-4 subapical; medial suture line of Cx-II+III relatively long; posterior suture line of Cx-IV weakly curved; excretory pore and Vgl-2 on the line of primary sclerotization near posterior idiosoma margin; ejaculatory complex conventional in shape (Fig. 2D); gnathosoma ventral margin only slightly curved, rostrum well developed (Fig. 2E); P-2 shorter than P-4, ventral margin with a fine denticulation also in proximal half of the segment, distally with a laterally compressed, anteriorly directed hyaline extension; P-3 with a subrectangular, apically serrated ventrodistal projection with a fine denticles; P-4 with long and broadly rounded distal seta (Fig. 2C), ventral tubercles well developed and separated.</paragraph>
<paragraph pageId="3" pageNumber="24">Measurements. Idiosoma (ventral view: Fig. 2B) L 591, W 341; dorsal shield (Figs 2A, 7B) L 472, W 305, L/W ratio 1.55; dorsal plate L 434; shoulder plate L 141-144, W 37-40, L/W ratio 3.6-3.8; frontal plate L 101-108, W 47-48, L/W ratio 2.2-2.3; shoulder/frontal plate L ratio 1.3-1.4. Gnathosomal bay L 81, Cx-I total L 200, Cx-I mL 117, Cx-II+III mL 113; ratio Cx-I L/Cx-II+III mL 1.8; Cx-I mL/Cx-II+III mL 1.04. Genital field L/W 113/95, ratio 1.19; ejaculatory complex L 166; distance genital field-excretory pore 133, genital field-caudal idiosoma margin 161. Gnathosoma vL 247; palp total L 226, dL: P-1, 22; P-2, 62; P-3, 48; P-4, 77; P-5, 17; P-2/P-4 ratio 0.8.</paragraph>
<caption pageId="3" pageNumber="24">
<paragraph pageId="3" pageNumber="24">
Figure 2.
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola dentifera" order="Trombidiformes" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="dentifera">Torrenticola dentifera</taxonomicName>
Wiles, 1991, male, stream in Naebyeansan NP, Korea: A dorsal shield B ventral shield C palp, medial view D ejaculatory complex E gnathosoma. Scale bars = 100
<normalizedToken originalValue="μm">μm</normalizedToken>
.
</paragraph>
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<subSubSection pageId="3" pageNumber="24" type="remarks">
<paragraph pageId="3" pageNumber="24">Remarks.</paragraph>
<paragraph pageId="3" pageNumber="24">
The single male specimen examined fits the original description of
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola dentifera" order="Trombidiformes" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="dentifera">Torrenticola dentifera</taxonomicName>
,which was based on two male specimens from Selangor, Peninsular Malaysia (
<bibRefCitation author="Wiles, PR" journalOrPublisher="Acarologia" pageId="24" pageNumber="45" pagination="41 - 56" title="Rheophilic watermites (Acari: Hydrachnidia) from mainland Malaysia." volume="32" year="1991">Wiles 1991</bibRefCitation>
). The only differences are found in larger dimensions of idiosoma and palps of the South Korea specimen.
</paragraph>
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<subSubSection pageId="3" pageNumber="24" type="habitat">
<paragraph pageId="3" pageNumber="24">Habitat.</paragraph>
<paragraph pageId="3" pageNumber="24">A permanent sandy/bouldary stream, shaded by riparian vegetation (Fig. 14B).</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="24" type="distribution">
<paragraph pageId="3" pageNumber="24">Distribution.</paragraph>
<paragraph pageId="3" pageNumber="24">
Malaysia (
<bibRefCitation author="Wiles, PR" journalOrPublisher="Acarologia" pageId="24" pageNumber="45" pagination="41 - 56" title="Rheophilic watermites (Acari: Hydrachnidia) from mainland Malaysia." volume="32" year="1991">Wiles 1991</bibRefCitation>
,
<bibRefCitation author="Wiles, PR" journalOrPublisher="Journal of Natural History" pageId="24" pageNumber="45" pagination="191 - 236" title="Asian and Oriental Torrenticolidae Piersig, 1902 (Acari: Hydrachnidia: Lebertioidea): a revision of the family and description of new species of Torrenticola Piersig and Pseudotorrenticola Walter, from Southeast Asia." url="10.1080/00222939700770121" volume="31" year="1997">1997</bibRefCitation>
). New for the fauna of Korea.
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<mods:classification id="B19988A6AC7827628E98156E2B6CE037">journal article</mods:classification>
<mods:identifier id="5D8030F83B6FD2BC47B9725FE77A963F" type="CLB-Dataset">173356</mods:identifier>
<mods:identifier id="C91EC604076483922260F58E13DFE587" type="DOI">10.5281/zenodo.5272840</mods:identifier>
<mods:identifier id="574BD62AC056470E1B22377599845F29" type="GBIF-Dataset">e508c8df-ec3b-45b4-8eb1-b000961914fa</mods:identifier>
<mods:identifier id="6613170A7315A155539C704483D0F083" type="ISSN">0253-116X</mods:identifier>
<mods:identifier id="6A273BE16F2D23054B3179E9BAC0FB57" type="Zenodo-Dep">5272840</mods:identifier>
</mods:mods>
<treatment id="B061879F9F18FFD133FFFE30FDEDFEEB" ID-DOI="http://doi.org/10.5281/zenodo.5272312" ID-GBIF-Taxon="203622998" ID-Zenodo-Dep="5272312" LSID="urn:lsid:plazi:treatment:B061879F9F18FFD133FFFE30FDEDFEEB" httpUri="http://treatment.plazi.org/id/B061879F9F18FFD133FFFE30FDEDFEEB" lastPageId="3" lastPageNumber="330" pageId="2" pageNumber="329">
<subSubSection id="70D265029F18FFD033FFFE30FD93FE6D" box="[151,619,459,481]" pageId="2" pageNumber="329" type="nomenclature">
<paragraph id="387736899F18FFD033FFFE30FD93FE6D" blockId="2.[151,619,459,481]" box="[151,619,459,481]" pageId="2" pageNumber="329">
<heading id="633F81E59F18FFD033FFFE30FD93FE6D" box="[151,619,459,481]" fontSize="8" level="2" pageId="2" pageNumber="329" reason="2">
<emphasis id="0ABCEA9B9F18FFD033FFFE30FD93FE6D" bold="true" box="[151,619,459,481]" pageId="2" pageNumber="329">
<taxonomicName id="FFC84D0A9F18FFD033FFFE30FE53FE6D" ID-CoL="8Y9D2" authority="Löbl, 2020" authorityName="Löbl" authorityYear="2020" box="[151,427,459,481]" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="329" phylum="Arthropoda" rank="species" species="insinuata" status="sp. nov.">
<emphasis id="0ABCEA9B9F18FFD033FFFE30FE53FE6D" bold="true" box="[151,427,459,481]" italics="true" pageId="2" pageNumber="329">Scaphobaeocera insinuata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="118F57E09F18FFD032DAFE37FE06FE6D" box="[434,510,460,481]" pageId="2" pageNumber="329" rank="species">nov.sp.</taxonomicNameLabel>
(
<figureCitation id="A0F32A0C9F18FFD03163FE30FD9BFE6D" box="[523,611,459,481]" captionStart="Figs 1-7" captionStartId="7.[151,191,1465,1487]" captionTargetBox="[173,1034,215,1445]" captionTargetId="figure-73@7.[153,1060,200,1452]" captionTargetPageId="7" captionText="Figs 1-7: Scaphobaeocera, aedeagi; (1, 2) S. furva nov.sp., in dorsal and lateral views, scale = 0.1 mm; (3, 4) Ditto, flagellum in dorsal and lateral views, scale = 0.05 mm; (5, 6) S. insinuata nov.sp., in dorsal and lateral views, scale = 0.1 mm; (7) Ditto, flagellum in lateral view, scale = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5272842" httpUri="https://zenodo.org/record/5272842/files/figure.png" pageId="2" pageNumber="329">Figs 5-7</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="70D265029F18FFD033FFFE09FEE2FD6C" pageId="2" pageNumber="329" type="materials_examined">
<paragraph id="387736899F18FFD033FFFE09FEE2FD6C" blockId="2.[151,1059,498,736]" pageId="2" pageNumber="329">
<collectionCode id="5ED9AE4C9F18FFD033FFFE09FF26FD8B" box="[151,222,498,519]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="329">Type</collectionCode>
material:
<materialsCitation id="88A03CD49F18FFD032F0FE09FDDAFDAC" ID-GBIF-Occurrence="3947022302" collectedFrom="Canacea litter" collectingDate="2009-04-13" collectionCode="MHNG" country="Taiwan" county="Taitung Co." location="rd. 20 km 173" municipality="Vit" pageId="2" pageNumber="329" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="E773882B9F18FFD032F0FE09FE11FD84" box="[408,489,498,520]" pageId="2" pageNumber="329" type="holotype">Holotype</typeStatus>
<specimenCount id="2ECEFD009F18FFD0329CFE09FDFFFD8B" box="[500,519,498,519]" pageId="2" pageNumber="329" type="male">♂</specimenCount>
:
<collectingCountry id="40DF76199F18FFD0317EFE09FD88FD8B" box="[534,624,498,519]" name="Taiwan" pageId="2" pageNumber="329">TAIWAN</collectingCountry>
<collectingCounty id="D1164E059F18FFD03113FE09FD12FD8B" box="[635,746,498,520]" pageId="2" pageNumber="329">Taitung Co.</collectingCounty>
<location id="3D1760529F18FFD0319CFE09FC7BFD8B" LSID="urn:lsid:plazi:treatment:B061879F9F18FFD133FFFE30FDEDFEEB:3D1760529F18FFD0319CFE09FC7BFD8B" box="[756,899,498,520]" country="Taiwan" county="Taitung Co." municipality="Vit" name="rd. 20 km 173" pageId="2" pageNumber="329">
rd.
<quantity id="FF309B6C9F18FFD0307FFE09FCADFD84" box="[791,853,498,520]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="329" unit="km" value="20.0">20 km</quantity>
173
</location>
,
<locationDeviation id="29C52FAB9F18FFD030F9FE09FC0AFD8B" bearing="180.0" box="[913,1010,498,519]" distance="1200.0" pageId="2" pageNumber="329">
<quantity id="FF309B6C9F18FFD030F9FE09FC22FD8B" box="[913,986,498,519]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="329" unit="m" value="1200.0">1200 m</quantity>
S
</locationDeviation>
.
<collectingMunicipality id="D813ACF39F18FFD03769FE09FBE7FD84" box="[1025,1055,498,520]" pageId="2" pageNumber="329">Vít</collectingMunicipality>
,
<date id="4C7610499F18FFD033ACFDF1FEDAFDAC" box="[196,290,522,544]" pageId="2" pageNumber="329" value="2009-04-13">
<collectingDate id="5C32E9A19F18FFD033ACFDF1FEDAFDAC" box="[196,290,522,544]" pageId="2" pageNumber="329" value="2009-04-13">13.iv.2009</collectingDate>
</date>
<collectedFrom id="8BD475749F18FFD03241FDF1FE5CFDAC" box="[297,420,522,544]" pageId="2" pageNumber="329">Canacea litter</collectedFrom>
10 (
<collectionCode id="5ED9AE4C9F18FFD032A6FDF1FDEFFDAC" box="[462,535,522,544]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="329" type="Museum">MHNG</collectionCode>
).
</materialsCitation>
<materialsCitation id="88A03CD49F18FFD03140FDF1FC79FDB4" ID-GBIF-Occurrence="3947022308" collectedFrom="mountain forest litter" collectingDate="2007-04-08" collectionCode="MHNG" collectorName="S. Vit" country="Taiwan" county="Taitung Co." elevation="1300" location="Rd no 20, km 174" municipality="Litao" pageId="2" pageNumber="329" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="E773882B9F18FFD03140FDF1FD78FD93" box="[552,640,522,543]" pageId="2" pageNumber="329" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="2ECEFD009F18FFD031E2FDF1FD5FFD93" box="[650,679,522,543]" pageId="2" pageNumber="329" type="male">1♂</specimenCount>
, S.
<collectingCountry id="40DF76199F18FFD031A3FDF1FCDFFD93" box="[715,807,522,543]" name="Taiwan" pageId="2" pageNumber="329">TAIWAN</collectingCountry>
,
<collectingCounty id="D1164E059F18FFD0305AFDF1FC66FD93" box="[818,926,522,544]" pageId="2" pageNumber="329">Taitung Co.</collectingCounty>
<location id="3D1760529F18FFD030CCFDF1FF1FFDBB" LSID="urn:lsid:plazi:treatment:B061879F9F18FFD133FFFE30FDEDFEEB:3D1760529F18FFD030CCFDF1FF1FFDBB" country="Taiwan" county="Taitung Co." municipality="Litao" name="Rd no 20, km 174" pageId="2" pageNumber="329">Rd no 20, km 174</location>
, after
<collectingMunicipality id="D813ACF39F18FFD03248FDD9FEB6FDB4" box="[288,334,546,568]" pageId="2" pageNumber="329">Litao</collectingMunicipality>
<date id="4C7610499F18FFD0323DFDD9FE51FDB4" box="[341,425,546,568]" pageId="2" pageNumber="329" value="2007-04-08">
<collectingDate id="5C32E9A19F18FFD0323DFDD9FE51FDB4" box="[341,425,546,568]" pageId="2" pageNumber="329" value="2007-04-08">8.iv.2007</collectingDate>
</date>
,
<collectorName id="953D535F9F18FFD032DBFDD9FE11FDB4" box="[435,489,546,568]" pageId="2" pageNumber="329">S. Vít</collectorName>
, env.
<quantity id="FF309B6C9F18FFD03177FDD9FD9CFDBB" box="[543,612,546,567]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="329" unit="m" value="1300.0">
<elevation id="B3E5D1BA9F18FFD03177FDD9FD9CFDBB" box="[543,612,546,567]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="329" unit="m" value="1300.0">1300 m</elevation>
</quantity>
<collectedFrom id="8BD475749F18FFD03102FDD9FCDEFDB4" box="[618,806,546,568]" pageId="2" pageNumber="329">mountain forest litter</collectedFrom>
(
<collectionCode id="5ED9AE4C9F18FFD0305AFDD9FC82FDB4" box="[818,890,546,568]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="329" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="88A03CD49F18FFD030E3FDD9FE23FDE4" ID-GBIF-Occurrence="3947022307" collectedFrom="Canacea litter" collectingDate="2008-03-25" collectionCode="MHNG" country="Taiwan" county="Hsinchu Co." location="Jienshih Twnsh" municipality="Vit" pageId="2" pageNumber="329" specimenCount="2" specimenCount-male="2">
<specimenCount id="2ECEFD009F18FFD030E3FDD9FC41FDBB" box="[907,953,546,567]" pageId="2" pageNumber="329" type="male">2♂♂</specimenCount>
,
<collectingCountry id="40DF76199F18FFD030ABFDD9FBE7FDBB" box="[963,1055,546,567]" name="Taiwan" pageId="2" pageNumber="329">TAIWAN</collectingCountry>
,
<collectingCounty id="D1164E059F18FFD033ACFDC1FECBFDC3" box="[196,307,570,592]" pageId="2" pageNumber="329">Hsinchu Co.</collectingCounty>
<location id="3D1760529F18FFD0325FFDC1FE3AFDDC" LSID="urn:lsid:plazi:treatment:B061879F9F18FFD133FFFE30FDEDFEEB:3D1760529F18FFD0325FFDC1FE3AFDDC" box="[311,450,570,592]" country="Taiwan" county="Hsinchu Co." municipality="Vit" name="Jienshih Twnsh" pageId="2" pageNumber="329">Jienshih Twnsh</location>
. near Hsinkuang vill.,
<location id="3D1760529F18FFD031EDFDC1FCF5FDDC" LSID="urn:lsid:plazi:treatment:B061879F9F18FFD133FFFE30FDEDFEEB:3D1760529F18FFD031EDFDC1FCF5FDDC" box="[645,781,570,592]" country="Taiwan" county="Hsinchu Co." municipality="Vit" name="km 44 rd no 60" pageId="2" pageNumber="329">km 44 rd no 60</location>
, env.
<locationDeviation id="29C52FAB9F18FFD03057FDC1FC6DFDC3" bearing="180.0" box="[831,917,570,592]" distance="1600.0" pageId="2" pageNumber="329">
<quantity id="FF309B6C9F18FFD03057FDC1FC7AFDC3" box="[831,898,570,592]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="329" unit="m" value="1600.0">1600 m</quantity>
S
</locationDeviation>
.
<collectingMunicipality id="D813ACF39F18FFD030F6FDC1FC44FDDC" box="[926,956,570,592]" pageId="2" pageNumber="329">Vít</collectingMunicipality>
,
<date id="4C7610499F18FFD030ACFDC1FBDBFDDC" box="[964,1059,570,592]" pageId="2" pageNumber="329" value="2008-03-25">
<collectingDate id="5C32E9A19F18FFD030ACFDC1FBDBFDDC" box="[964,1059,570,592]" pageId="2" pageNumber="329" value="2008-03-25">25.iii.2008</collectingDate>
</date>
<collectedFrom id="8BD475749F18FFD033ACFDA9FEB9FDE4" box="[196,321,594,616]" pageId="2" pageNumber="329">Canacea litter</collectedFrom>
, ♂ iii6 (
<collectionCode id="5ED9AE4C9F18FFD032E4FDA9FE2DFDE4" box="[396,469,594,616]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="329" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="88A03CD49F18FFD0328DFDA9FBE7FDE4" ID-GBIF-Occurrence="3947022304" box="[485,1055,594,616]" collectingDate="1977-06-12" collectionCode="MHNG" collectorName="J. Klapperich" country="Taiwan" elevation="1400" location="Fenchihu" municipality="Fenchihu" pageId="2" pageNumber="329" specimenCount="1" specimenCount-male="1">
<specimenCount id="2ECEFD009F18FFD0328DFDA9FDFAFDEB" box="[485,514,594,615]" pageId="2" pageNumber="329" type="male">1♂</specimenCount>
,
<collectingCountry id="40DF76199F18FFD03165FDA9FDB7FDE4" box="[525,591,594,616]" name="Taiwan" pageId="2" pageNumber="329">Taiwan</collectingCountry>
<collectingMunicipality id="D813ACF39F18FFD0313EFDA9FD50FDE4" box="[598,680,594,616]" country="Taiwan" name="Fenchihu" pageId="2" pageNumber="329">Fenchihu</collectingMunicipality>
<quantity id="FF309B6C9F18FFD031C6FDA9FD0CFDEB" box="[686,756,594,615]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="329" unit="m" value="1400.0">
<elevation id="B3E5D1BA9F18FFD031C6FDA9FD0CFDEB" box="[686,756,594,615]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="329" unit="m" value="1400.0">1400 m</elevation>
</quantity>
, 12.vi.77
<collectorName id="953D535F9F18FFD03025FDA9FC3BFDE4" box="[845,963,594,616]" pageId="2" pageNumber="329">J. Klapperich</collectorName>
(
<collectionCode id="5ED9AE4C9F18FFD030B8FDA9FBE1FDE4" box="[976,1049,594,616]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="329" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="88A03CD49F18FFD033ACFD91FC0EFD0C" ID-GBIF-Occurrence="3947022301" box="[196,1014,617,640]" collectingDate="1990-05-16" collectionCode="MHNG" collectorName="A. Smetana" country="Taiwan" elevation="1650" location="Shanlinchi" municipality="Shanlinchi" pageId="2" pageNumber="329" specimenCount="2" specimenCount-male="2" stateProvince="Nantou Hsien">
<specimenCount id="2ECEFD009F18FFD033ACFD91FF0AFDF3" box="[196,242,618,639]" pageId="2" pageNumber="329" type="male">2♂♂</specimenCount>
,
<collectingCountry id="40DF76199F18FFD03395FD91FEAFFDF3" box="[253,343,618,639]" name="Taiwan" pageId="2" pageNumber="329">TAIWAN</collectingCountry>
<collectingRegion id="FA0CF86B9F18FFD03236FD91FE23FD0C" box="[350,475,618,640]" country="Taiwan" name="Nantou" pageId="2" pageNumber="329">Nantou Hsien</collectingRegion>
,
<collectingMunicipality id="D813ACF39F18FFD0328EFD91FDBBFD0C" box="[486,579,618,640]" country="Taiwan" name="Shanlinchi" pageId="2" pageNumber="329" stateProvince="Nantou">Shanlinchi</collectingMunicipality>
<quantity id="FF309B6C9F18FFD03122FD91FD77FDF3" box="[586,655,618,640]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="2" pageNumber="329" unit="m" value="1650.0">
<elevation id="B3E5D1BA9F18FFD03122FD91FD77FDF3" box="[586,655,618,640]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="2" pageNumber="329" unit="m" value="1650.0">1650 m</elevation>
</quantity>
<date id="4C7610499F18FFD031FEFD91FD0AFD0C" box="[662,754,618,640]" pageId="2" pageNumber="329" value="1990-05-16">
<collectingDate id="5C32E9A19F18FFD031FEFD91FD0AFD0C" box="[662,754,618,640]" pageId="2" pageNumber="329" value="1990-05-16">16.V.1990</collectingDate>
</date>
<collectorName id="953D535F9F18FFD03191FD91FC9AFDF3" box="[761,866,618,639]" pageId="2" pageNumber="329">A. Smetana</collectorName>
[T60] (
<collectionCode id="5ED9AE4C9F18FFD030C0FD91FC08FD0C" box="[936,1008,618,640]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="329" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="88A03CD49F18FFD03769FD91FEE2FD6C" ID-GBIF-Occurrence="3947022306" collectedFrom="primary forest on the slope with sparse understory: sifting of small accumulation of leaves" collectionCode="NMNS" collectorName="Damaska &amp; Fikacek &amp; Hu &amp; Liu" country="Taiwan" elevation="1150" latitude="24.074461" location="Huisun Forest reserve" longLatPrecision="1" longitude="121.03664" pageId="2" pageNumber="329" specimenCount="1" specimenCount-male="1" stateProvince="Nantou County">
<specimenCount id="2ECEFD009F18FFD03769FD91FBE7FDF3" box="[1025,1055,618,639]" pageId="2" pageNumber="329" type="male">1♂</specimenCount>
.
<collectingCountry id="40DF76199F18FFD033ACFD79FED9FD1B" box="[196,289,642,663]" name="Taiwan" pageId="2" pageNumber="329">TAIWAN</collectingCountry>
:
<collectingRegion id="FA0CF86B9F18FFD03247FD79FE44FD1B" box="[303,444,642,663]" country="Taiwan" name="Nantou" pageId="2" pageNumber="329">Nantou County</collectingRegion>
<location id="3D1760529F18FFD032AFFD79FD6DFD1B" LSID="urn:lsid:plazi:treatment:B061879F9F18FFD133FFFE30FDEDFEEB:3D1760529F18FFD032AFFD79FD6DFD1B" box="[455,661,642,664]" country="Taiwan" latitude="24.074461" longLatPrecision="1" longitude="121.03664" name="Huisun Forest reserve" pageId="2" pageNumber="329" stateProvince="Nantou County">Huisun Forest reserve</location>
, track to
<location id="3D1760529F18FFD0319FFD79FC6FFD1B" LSID="urn:lsid:plazi:treatment:B061879F9F18FFD133FFFE30FDEDFEEB:3D1760529F18FFD0319FFD79FC6FFD1B" box="[759,919,642,664]" country="Taiwan" latitude="24.074461" longLatPrecision="1" longitude="121.03664" municipality="Huisun Forest reserve" name="Xiaochushan Mt." pageId="2" pageNumber="329" stateProvince="Nantou County">Xiaochushan Mt.</location>
,
<geoCoordinate id="5DFC504E9F18FFD030CDFD79FBD8FD14" box="[933,1056,642,664]" degrees="24.0744602" direction="north" orientation="latitude" pageId="2" pageNumber="329" precision="1" value="24.074461">24.0744602N</geoCoordinate>
,
<geoCoordinate id="5DFC504E9F18FFD033ACFD61FEB0FD3C" box="[196,328,666,688]" degrees="121.0366337" direction="east" orientation="longitude" pageId="2" pageNumber="329" precision="1" value="121.03664">121.0366337E</geoCoordinate>
;
<quantity id="FF309B6C9F18FFD03239FD61FE6EFD23" box="[337,406,666,688]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="2" pageNumber="329" unit="m" value="1150.0">
<elevation id="B3E5D1BA9F18FFD03239FD61FE6EFD23" box="[337,406,666,688]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="2" pageNumber="329" unit="m" value="1150.0">1150 m</elevation>
</quantity>
4.v.2919;
<collectorName id="953D535F9F18FFD0329DFD61FDB1FD3C" box="[501,585,666,688]" pageId="2" pageNumber="329">Damaška</collectorName>
,
<collectorName id="953D535F9F18FFD0313BFD61FD63FD3C" box="[595,667,666,688]" pageId="2" pageNumber="329">Fikáček</collectorName>
,
<collectorName id="953D535F9F18FFD031CCFD61FD47FD23" box="[676,703,666,687]" pageId="2" pageNumber="329">Hu</collectorName>
&amp;
<collectorName id="953D535F9F18FFD031B5FD61FD04FD3C" box="[733,764,666,688]" pageId="2" pageNumber="329">Liu</collectorName>
lgt. /
<collectedFrom id="8BD475749F18FFD0305AFD61FD1FFD44" pageId="2" pageNumber="329">primary forest on the slope with sparse understory: sifting of small accumulation of leaves</collectedFrom>
/ Huisun Leaf Litter Beetles Project (
<collectionCode id="5ED9AE4C9F18FFD033A3FD31FEE8FD6C" box="[203,272,714,736]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="2" pageNumber="329">NMNS</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="70D265029F18FFD033FFFD15FD2FFA40" pageId="2" pageNumber="329" type="description">
<paragraph id="387736899F18FFD033FFFD15FD60FB55" blockId="2.[151,1059,750,1241]" pageId="2" pageNumber="329">
Description: Length
<quantity id="FF309B6C9F18FFD032ADFD15FDA0FC88" box="[453,600,750,772]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.25" metricValueMin="1.15" pageId="2" pageNumber="329" unit="mm" value="1.2" valueMax="1.25" valueMin="1.15">1.15-1.25 mm</quantity>
, width
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, dorsoventral diameter
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. Head, pronotum, elytra and ventral side of thorax light reddish-brown to blackish-brown, elytra usually slightly lighter near apices. Abdomen about as thorax, with apical segments ochraceous. Appendages ochraceous. Length/width ratios of antennomeres as: III 18/7: IV 23/7: V 28/7: VI 23/8: VII 25/13: VIII 18/11: IX 23/15: X 25/17: XI 38/19. Pronotum very finely punctate, not microsculptured. Scutellum concealed or its tip exposed. Elytra not iridescent, with hardly visible parasutural striae, sutural striae deep, curved at base, discal punctation about as fine as pronotal punctation, microsculpture absent. Hypomera not microsculptured, smooth, lacking longitudinal stria. Middle area of metaventrite flat, entirely densely, distinctly punctate, with punctures to part about as large as puncture intervals; pubescence long; median stria and median impression absent. Lateral parts of metaventrite very finely and sparsely punctate, not microsculptured. Submesocoxal areas about
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, submesocoxal lines appearing impunctate. Metanepisterna about
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wide, flat, not narrowed posteriad, impunctate, with straight suture reaching metepimera. Tibiae straight. Ventrite 1 not microsculptured, very finely punctate, basal punctures distinct, slightly elongate. Following sternites with strigulate microsculpture.
</paragraph>
<paragraph id="387736899F18FFD033FFFB12FD2FFA40" blockId="2.[151,1059,1257,1484]" pageId="2" pageNumber="329">
Male characters: Protarsomeres 1 to 3 strongly widened, wide almost as apices of protibiae. Aedeagus (
<figureCitation id="A0F32A0C9F18FFD032B2FAFDFDCBFA90" box="[474,563,1286,1308]" captionStart="Figs 1-7" captionStartId="7.[151,191,1465,1487]" captionTargetBox="[173,1034,215,1445]" captionTargetId="figure-73@7.[153,1060,200,1452]" captionTargetPageId="7" captionText="Figs 1-7: Scaphobaeocera, aedeagi; (1, 2) S. furva nov.sp., in dorsal and lateral views, scale = 0.1 mm; (3, 4) Ditto, flagellum in dorsal and lateral views, scale = 0.05 mm; (5, 6) S. insinuata nov.sp., in dorsal and lateral views, scale = 0.1 mm; (7) Ditto, flagellum in lateral view, scale = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5272842" httpUri="https://zenodo.org/record/5272842/files/figure.png" pageId="2" pageNumber="329">Figs 5-7</figureCitation>
)
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long. Median lobe with narrow basal bulb, apical process asymmetrical, moderately inflexed, long almost as basal bulb, ventral side irregularly concave, tip acute and bent in lateral view, blunt in dorsal view. Articular process not prominent. Parameres symmetrical, oblique, reaching posterior level of tip of median lobe, almost evenly broad in dorsal view, strongly widened apically in lateral view. Internal sac with narrow, convoluted flagellum forming two complete loops, widened and folded in proximal section.
</paragraph>
</subSubSection>
<subSubSection id="70D265029F18FFD133FFFA20FD9AFEA8" lastPageId="3" lastPageNumber="330" pageId="2" pageNumber="329" type="diagnosis">
<paragraph id="387736899F18FFD133FFFA20FD9AFEA8" blockId="2.[151,1058,1499,1580]" lastBlockId="3.[151,1058,211,292]" lastPageId="3" lastPageNumber="330" pageId="2" pageNumber="329">
Differential diagnosis: This species is very similar and likely related with
<taxonomicName id="FFC84D0A9F18FFD033A4FA02FEE5F982" authorityName="Löbl" authorityYear="2020" box="[204,285,1529,1550]" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="329" phylum="Arthropoda" rank="species" species="furva">
<emphasis id="0ABCEA9B9F18FFD033A4FA02FEE5F982" box="[204,285,1529,1550]" italics="true" pageId="2" pageNumber="329">S. furva</emphasis>
</taxonomicName>
. It differs in external characters from the latter by the wider antennomeres IX to XI and the setae on the median part of metaventrite distinctly longer. The aedeagi provide reliable distinguish characters, the apical process of the median lobe is notably longer, the parameres are more expanded apically, and the flagellum forming only two loops is distinctly folded in proximal section.
</paragraph>
</subSubSection>
<subSubSection id="70D265029F19FFD133FFFEC8FDEDFEEB" pageId="3" pageNumber="330" type="etymology">
<paragraph id="387736899F19FFD133FFFEC8FDEDFEEB" blockId="3.[151,1058,307,359]" pageId="3" pageNumber="330">Etymology: The species epithet is a Latin adjective meaning not sinuate, referring to the proximal section of the flagellum.</paragraph>
</subSubSection>
</treatment>
</document>

140
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<mods:title id="AC4AAF1EDF5068A07F551E79FE706CAF">Four new Taiwanese species of Scaphobaeocera CSIKI (Coleoptera, Staphylinidae, Scaphidiinae)</mods:title>
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<emphasis id="0ABCEA9B9F19FFD133FFFE6AFE5EFE2B" bold="true" box="[151,422,401,423]" italics="true" pageId="3" pageNumber="330">Scaphobaeocera montana</emphasis>
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(
<figureCitation id="A0F32A0C9F19FFD1316EFE6AFD92FE2B" box="[518,618,401,423]" captionStart="Figs 8-14" captionStartId="8.[151,191,1496,1518]" captionTargetBox="[169,1050,229,1467]" captionTargetId="figure-79@8.[154,1061,203,1485]" captionTargetPageId="8" captionText="Figs 8-14: Scaphobaeocera, aedeagi; (8, 9) S. montana nov.sp., in dorsal and lateral views, scale = 0.1 mm; (10) Ditto, flagellum and proximal sclerite of internal sac, dorsal view, scle = 0.05 mm; (11, 12) S. tengchinensis nov.sp., in dorsal and lateral views, scale = 0.1 mm; (13, 14) Ditto, flagellum in dorsal and lateral views, scale = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5272846" httpUri="https://zenodo.org/record/5272846/files/figure.png" pageId="3" pageNumber="330">Figs 8-10</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="70D265029F19FFD133FFFE4CFDD4FE71" pageId="3" pageNumber="330" type="materials_examined">
<paragraph id="387736899F19FFD133FFFE4CFDD4FE71" blockId="3.[151,1059,439,509]" pageId="3" pageNumber="330">
Type material:
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<specimenCount id="2ECEFD009F19FFD1328CFE4CFE00FE40" box="[484,504,439,460]" pageId="3" pageNumber="330" type="male">♂</specimenCount>
:
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<collectingRegion id="FA0CF86B9F19FFD13137FE4CFD27FE41" box="[607,735,439,461]" country="Taiwan" name="Hualien" pageId="3" pageNumber="330">Hualien Hsien</collectingRegion>
,
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<quantity id="FF309B6C9F19FFD130B7FE4CFBDBFE40" box="[991,1059,439,460]" metricMagnitude="3" metricUnit="m" metricValue="2.2199999999999998" pageId="3" pageNumber="330" unit="m" value="2220.0">
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<collectorName id="953D535F9F19FFD1324EFE34FE76FE68" box="[294,398,463,484]" pageId="3" pageNumber="330">A. Smetana</collectorName>
[T55] (
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)
</materialsCitation>
.
<materialsCitation id="88A03CD49F19FFD13141FE34FDD4FE71" ID-GBIF-Occurrence="3947022305" collectingDate="1990-05-04" collectionCode="MHNG" collectorName="A. Smetana" country="Taiwan" elevation="2230" location="Anmachan" municipality="Hsien" pageId="3" pageNumber="330" specimenCount="1" specimenCount-male="1" stateProvince="Taichung" typeStatus="paratype">
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:
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<collectingMunicipality id="D813ACF39F19FFD1303AFE34FC7FFE69" box="[850,903,463,485]" pageId="3" pageNumber="330">Hsien</collectingMunicipality>
,
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<quantity id="FF309B6C9F19FFD1309EFE34FF2EFE70" metricMagnitude="3" metricUnit="m" metricValue="2.23" pageId="3" pageNumber="330" unit="m" value="2230.0">
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[T43] (
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).
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</paragraph>
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<paragraph id="387736899F19FFD133FFFDF0FDD1FBBD" blockId="3.[151,1059,523,1073]" pageId="3" pageNumber="330">
Description: Length
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, width
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, dorsoventral diameter
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. Frons, pronotum and elytra, elytral apices excepted, blackish. Elytral with narrowly lighter apices. Ventral side of thorax, most of exposed tergites and ventrites somewhat lighter than pronotum and elytra. Femora and tibiae dark reddish-brown, apical abdominal segments, tarsi and antennae lighter than tibiae, yellowish. Length/width ratios of antennomeres as: III 16/7: IV 22/7: V 28/7: VI 20/8: VII 35/15: VIII 24/12: IX 35/15: X 34/18: XI 55/18. Pronotum extremely finely punctate, punctured hardly visible at 100 times magnification, not microsculptured. Minute point of scutellum exposed. Elytra weakly iridescent, with distinct parasutural striae, sutural striae deep, curved at base, discal punctation about as fine as pronotal punctation, microsculpture appearing absent. Hypomeron not microsculptured, smooth, with longitudinal stria separating upper oblique part from lower vertical part. Middle area of metaventrite convex, with large centre smooth, densely, distinctly punctate around impunctate centre and with punctures to part about as large as puncture intervals; pubescence short; median stria and median impression absent. Lateral parts of metaventrite very finely and sparsely punctate, not microsculptured. Submesocoxal areas about
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, submesocoxal lines appearing impunctate. Tibiae straight. Ventrite 1 not microsculptured, very finely punctate, basal punctures distinct, not elongate. Following sternites with strigulate microsculpture.
</paragraph>
<paragraph id="387736899F19FFD133FFFBBAFD8DFAA8" blockId="3.[151,1059,1089,1316]" pageId="3" pageNumber="330">
Male characters: Protarsomeres slightly widened, much narrower than apices of tibiae. Aedeagus (
<figureCitation id="A0F32A0C9F19FFD13219FBA5FE0CFBF8" box="[369,500,1118,1140]" captionStart="Figs 8-14" captionStartId="8.[151,191,1496,1518]" captionTargetBox="[169,1050,229,1467]" captionTargetId="figure-79@8.[154,1061,203,1485]" captionTargetPageId="8" captionText="Figs 8-14: Scaphobaeocera, aedeagi; (8, 9) S. montana nov.sp., in dorsal and lateral views, scale = 0.1 mm; (10) Ditto, flagellum and proximal sclerite of internal sac, dorsal view, scle = 0.05 mm; (11, 12) S. tengchinensis nov.sp., in dorsal and lateral views, scale = 0.1 mm; (13, 14) Ditto, flagellum in dorsal and lateral views, scale = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5272846" httpUri="https://zenodo.org/record/5272846/files/figure.png" pageId="3" pageNumber="330">Figs 8-10.46</figureCitation>
-
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long. Median lobe with narrow basal bulb, apical process asymmetrical, weakly inflexed, almost long much shorter than basal bulb, ventral side weakly concave, tip blunt and not bent in lateral view, blunt in dorsal view. Articular process not prominent. Parameres symmetrical, almost in axis with the median lobe, reaching posterior level of tip of median lobe, almost evenly broad in dorsal view, strongly widened apically in lateral view. Internal sac with large basal sclerite joined with narrow, bent flagellum not forming loops.
</paragraph>
</subSubSection>
<subSubSection id="70D265029F19FFD133FFFAC8FC30F99A" pageId="3" pageNumber="330" type="diagnosis">
<paragraph id="387736899F19FFD133FFFAC8FC30F99A" blockId="3.[151,1058,1331,1559]" pageId="3" pageNumber="330">
Differential diagnosis: The aedeagal characters of this species suggest relationships with the Japanese
<taxonomicName id="FFC84D0A9F19FFD132B4FAAAFD2CFAEA" authority="LOBL, 1981" authorityName="LOBL" authorityYear="1981" box="[476,724,1359,1382]" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="330" phylum="Arthropoda" rank="species" species="smetanai">
<emphasis id="0ABCEA9B9F19FFD132B4FAAAFDA9FAEA" box="[476,593,1361,1382]" italics="true" pageId="3" pageNumber="330">S. smetanai</emphasis>
<bibRefCitation id="5C594B789F19FFD13131FAB4FD2CFAEA" author="LOBL I" box="[601,724,1359,1382]" pageId="3" pageNumber="330" pagination="229 - 244" refId="ref4053" refString="LOBL I. (1981): Uber die japanische Arten der Gattungen Scaphobaeocera CSIKI und Scaphoxium LOBL (Col., Scaphidiidae). - Mitteilungen der Schweizerischen entomologischen Gesellschaft 54: 229 - 244." type="journal article" year="1981">LÖBL, 1981</bibRefCitation>
</taxonomicName>
. The latter may be distinguished by the well visible elytral and abdominal microsculpture, the distinctly iridescent elytra, and the length/width ratios of the antennomeres, notably by the antennomeres VIII about as narrow as the antennomeres VI and the antennomeres XI about 2.5 times as long as wide. The aedeagi of these two species differ by apical process of the median lobe narrower (lateral view), the margin of the basal sclerite of the internal sac broadly convex, and the proximal section of the flagellum much narrower in
<taxonomicName id="FFC84D0A9F19FFD13038F9FAFC3AF99A" authorityName="Löbl" authorityYear="2020" box="[848,962,1537,1558]" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="330" phylum="Arthropoda" rank="species" species="montana">
<emphasis id="0ABCEA9B9F19FFD13038F9FAFC3AF99A" box="[848,962,1537,1558]" italics="true" pageId="3" pageNumber="330">S. montana</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="70D265029F19FFD133FFF9DDFC4BF9B0" box="[151,947,1574,1596]" pageId="3" pageNumber="330" type="etymology">
<paragraph id="387736899F19FFD133FFF9DDFC4BF9B0" blockId="3.[151,947,1574,1596]" box="[151,947,1574,1596]" pageId="3" pageNumber="330">Etymology: The species epithet is a Latin adjective meaning mountain.</paragraph>
</subSubSection>
</treatment>
</document>

196
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<emphasis id="0ABCEA9B9F1BFFD333FFFE27FE79FE7E" bold="true" box="[151,385,476,498]" italics="true" pageId="1" pageNumber="328">Scaphobaeocera furva</emphasis>
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(
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)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="70D265029F1BFFD333FFFDF9FEE5FDEC" pageId="1" pageNumber="328" type="materials_examined">
<paragraph id="387736899F1BFFD333FFFDF9FEE5FDEC" blockId="1.[151,1059,514,608]" pageId="1" pageNumber="328">
<collectionCode id="5ED9AE4C9F1BFFD333FFFDF9FF26FD9B" box="[151,222,514,535]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="328">Type</collectionCode>
material:
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<typeStatus id="E773882B9F1BFFD332E5FDF9FE27FD94" box="[397,479,514,536]" pageId="1" pageNumber="328" type="holotype">Holotype</typeStatus>
<specimenCount id="2ECEFD009F1BFFD3328CFDF9FE0FFD9B" box="[484,503,514,535]" pageId="1" pageNumber="328" type="male">♂</specimenCount>
:
<collectingCountry id="40DF76199F1BFFD33169FDF9FDA3FD9B" box="[513,603,514,535]" name="Taiwan" pageId="1" pageNumber="328">TAIWAN</collectingCountry>
<collectingCounty id="D1164E059F1BFFD33108FDF9FD46FD9B" box="[608,702,514,536]" pageId="1" pageNumber="328">Taipei Co.</collectingCounty>
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slopes E of
<collectingMunicipality id="D813ACF39F1BFFD330B4FDF9FF06FDBC" pageId="1" pageNumber="328">Shamao S. Vít</collectingMunicipality>
,
<date id="4C7610499F1BFFD33263FDE1FE90FDA3" box="[267,360,538,559]" pageId="1" pageNumber="328" value="2007-10-24">
<collectingDate id="5C32E9A19F1BFFD33263FDE1FE90FDA3" box="[267,360,538,559]" pageId="1" pageNumber="328" value="2007-10-24">24.X.2007</collectingDate>
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</quantity>
, rotten bole (
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)
</materialsCitation>
.
<materialsCitation id="88A03CD49F1BFFD331EBFDE1FEE1FDEC" ID-GBIF-Occurrence="3947022311" collectingDate="2008-03-25" collectionCode="MHNG" collectorName="S. Vit" country="Taiwan" county="Hsinchu Co." elevation="1800" location="Jienshih Twnsh. env" pageId="1" pageNumber="328" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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<specimenCount id="2ECEFD009F1BFFD331B1FDE1FD15FDA3" box="[729,749,538,559]" pageId="1" pageNumber="328" type="male">♂</specimenCount>
:
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<collectingCounty id="D1164E059F1BFFD33036FDE1FC2AFDA3" box="[862,978,538,560]" pageId="1" pageNumber="328">Hsinchu Co.</collectingCounty>
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.
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;
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<collectingDate id="5C32E9A19F1BFFD3315CFDC9FD6AFDC4" box="[564,658,562,584]" pageId="1" pageNumber="328" value="2008-03-25">25.iii.2008</collectingDate>
</date>
Hsingkuang trib. vill., leg.
<collectorName id="953D535F9F1BFFD330ECFDC9FC40FDC4" box="[900,952,562,584]" pageId="1" pageNumber="328">S. Vít</collectorName>
Forest litter (
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)
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.
</paragraph>
</subSubSection>
<subSubSection id="70D265029F1BFFD333FFFD95FDEAFAC0" pageId="1" pageNumber="328" type="description">
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Description: Length
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, width
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, dorsoventral diameter
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. Head, pronotum, elytra and ventral side of thorax dark brown to black, elytra lighter near apices. Abdomen dark brown, with apical segments ochraceous. Appendages ochraceous. Length/width ratios of antennomeres as: III 14/6: IV 20/6: V 27/6: VI 20/7: VII 27/8: VIII 13/9: IX 28/12: X 26/14: XI 38/12. Pronotum very finely punctate, not microsculptured. Tip of scutellum exposed. Elytra not iridescent, with parasutural striae, sutural striae deep, curved at base, discal punctation about as fine as pronotal punctation, microsculpture hardly visible. Hypomera not microsculptured, smooth, lacking longitudinal stria. Middle area of metaventrite hardly convex, entirely densely, distinctly punctate, with punctures to part about as large as puncture intervals; pubescence very short, median stria and median impression absent. Lateral parts of metaventrite very finely and sparsely punctate, not microsculptured. Submesocoxal areas about
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, submesocoxal lines appearing impunctate. Metanepisterna about
<quantity id="FF309B6C9F1BFFD33085FC35FF09FB8D" metricMagnitude="-5" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="1" pageNumber="328" unit="mm" value="0.025" valueMax="0.03" valueMin="0.02">0.02- 0.03 mm</quantity>
wide, flat, slightly narrowed posteriad, impunctate, with slightly concave suture reaching metepimera. Tibiae straight. Ventrite 1 not microsculptured, very finely punctate, basal punctures distinct, not elongate. Following sternites with strigulate microsculpture.
</paragraph>
<paragraph id="387736899F1BFFD333FFFB92FDEAFAC0" blockId="1.[151,1058,1129,1356]" pageId="1" pageNumber="328">
Male characters: Protarsomeres 1 to 3 strongly widened, wide almost as apices of protibiae. Aedeagus (
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)
<quantity id="FF309B6C9F1BFFD3312BFB7DFD2CFB10" box="[579,724,1158,1180]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" metricValueMax="4.6" metricValueMin="4.0" pageId="1" pageNumber="328" unit="mm" value="0.43" valueMax="0.46" valueMin="0.4">0.40-0.46 mm</quantity>
long. Median lobe with narrow basal bulb, apical process asymmetrical, moderately inflexed, much shorter than basal bulb, concave ventral side and tip acute in lateral view, blunt in dorsal view. Articular process not prominent. Parameres symmetrical, straight, reaching level of tip of median lobe, almost evenly broad in dorsal view, strongly widened dorso-apically in lateral view. Internal sac with narrow, convoluted flagellum forming three complete loops, moderately widened at proximal end.
</paragraph>
</subSubSection>
<subSubSection id="70D265029F1BFFD033FFFAA0FDB4FEF0" lastPageId="2" lastPageNumber="329" pageId="1" pageNumber="328" type="diagnosis">
<paragraph id="387736899F1BFFD033FFFAA0FDB4FEF0" blockId="1.[151,1059,1371,1599]" lastBlockId="2.[151,1059,211,380]" lastPageId="2" lastPageNumber="329" pageId="1" pageNumber="328">
Differential diagnosis: The key given below provides means to distinguish this new species from its Taiwanese congeners. The aedeagal characters suggest relationships with
<taxonomicName id="FFC84D0A9F1BFFD332C4FA6DFCC3FA27" authority="HOSHINA &amp; SUGAYA, 2003" authorityName="HOSHINA &amp; SUGAYA" authorityYear="2003" box="[428,827,1430,1452]" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="328" phylum="Arthropoda" rank="species" species="sunadai">
<emphasis id="0ABCEA9B9F1BFFD332C4FA6DFDE0FA27" box="[428,536,1430,1451]" italics="true" pageId="1" pageNumber="328">S. sunadai</emphasis>
<bibRefCitation id="5C594B789F1BFFD33149FA6CFCC3FA27" author="HOSHINA H &amp; H. SUGAYA" box="[545,827,1430,1452]" firstAuthor="HOSHINA" journalOrPublisher="Entomological Review of Japan" pageId="1" pageNumber="328" pagination="35 - 41" part="58" refId="ref3977" refString="HOSHINA H. &amp; H. SUGAYA (2003): New records of the genera Scaphobaeocera and Scaphoxium (Coleoptera: Staphylinidae) from the Ryukyus, Japan, with descriptions of two new species. - Entomological Review of Japan 58: 35 - 41." title="New records of the genera Scaphobaeocera and Scaphoxium (Coleoptera: Staphylinidae) from the Ryukyus, Japan, with descriptions of two new species" type="journal article" year="2003">HOSHINA &amp; SUGAYA, 2003</bibRefCitation>
</taxonomicName>
from the Ryukyus,
<taxonomicName id="FFC84D0A9F1BFFD33767FA6DFE98FA44" authority="LOBL, 2018" authorityName="LOBL" authorityYear="2018" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="328" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="0ABCEA9B9F1BFFD33767FA6DFF22FA44" italics="true" pageId="1" pageNumber="328">S. glabra</emphasis>
LÖBL, 2018
</taxonomicName>
from Yunnan, and
<taxonomicName id="FFC84D0A9F1BFFD3315CFA4FFCDBFA44" authority="LOBL, 1981" authorityName="LOBL" authorityYear="1981" box="[564,803,1458,1481]" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="328" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="0ABCEA9B9F1BFFD3315CFA4FFD67FA44" box="[564,671,1459,1481]" italics="true" pageId="1" pageNumber="328">S. gracilis</emphasis>
<bibRefCitation id="5C594B789F1BFFD331C1FA49FCDBFA44" author="LOBL I" box="[681,803,1458,1481]" pageId="1" pageNumber="328" pagination="229 - 244" refId="ref4053" refString="LOBL I. (1981): Uber die japanische Arten der Gattungen Scaphobaeocera CSIKI und Scaphoxium LOBL (Col., Scaphidiidae). - Mitteilungen der Schweizerischen entomologischen Gesellschaft 54: 229 - 244." type="journal article" year="1981">LÖBL, 1981</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="FFC84D0A9F1BFFD33034FA4FFF32FA6A" authority="LOBL, 1981" authorityName="LOBL" authorityYear="1981" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="328" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="0ABCEA9B9F1BFFD33034FA4FFC22FA44" box="[860,986,1459,1481]" italics="true" pageId="1" pageNumber="328">S. variabilis</emphasis>
<bibRefCitation id="5C594B789F1BFFD3308CFA49FF32FA6A" author="LOBL I" pageId="1" pageNumber="328" pagination="229 - 244" refId="ref4053" refString="LOBL I. (1981): Uber die japanische Arten der Gattungen Scaphobaeocera CSIKI und Scaphoxium LOBL (Col., Scaphidiidae). - Mitteilungen der Schweizerischen entomologischen Gesellschaft 54: 229 - 244." type="journal article" year="1981">LÖBL, 1981</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="40DF76199F1BFFD33265FA2AFEB2FA6A" box="[269,330,1489,1510]" name="Japan" pageId="1" pageNumber="328">Japan</collectingCountry>
.
<taxonomicName id="FFC84D0A9F1BFFD3323EFA2AFDA0FA6A" authorityName="HOSHINA &amp; SUGAYA" authorityYear="2003" box="[342,600,1489,1510]" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="328" phylum="Arthropoda" rank="species" species="sunadai">
<emphasis id="0ABCEA9B9F1BFFD3323EFA2AFDA0FA6A" box="[342,600,1489,1510]" italics="true" pageId="1" pageNumber="328">Scaphobaeocera sunadai</emphasis>
</taxonomicName>
may be readily distinguished from
<taxonomicName id="FFC84D0A9F1BFFD330B9FA2AFBDAFA6A" authorityName="Löbl" authorityYear="2020" box="[977,1058,1489,1510]" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="328" phylum="Arthropoda" rank="species" species="furva">
<emphasis id="0ABCEA9B9F1BFFD330B9FA2AFC26FA6A" box="[977,990,1489,1510]" italics="true" pageId="1" pageNumber="328">S</emphasis>
.
<emphasis id="0ABCEA9B9F1BFFD33085FA2AFBDAFA6A" box="[1005,1058,1489,1510]" italics="true" pageId="1" pageNumber="328">furva</emphasis>
</taxonomicName>
by the parameres gradually widened apically, the apical process of the median lobe strongly inflexed, near apex almost perpendicular to the axis of aedeagus (lateral view) and the flagellum forming a single complete loop. The apical process of the median lobe is much shorter and has blunt apex in
<taxonomicName id="FFC84D0A9F18FFD0313CFF2FFD40FF64" authorityName="LOBL" authorityYear="2018" box="[596,696,211,233]" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="329" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="0ABCEA9B9F18FFD0313CFF2FFD40FF64" box="[596,696,211,233]" italics="true" pageId="2" pageNumber="329">S. glabra</emphasis>
</taxonomicName>
. This species differs in external characters drastically by its strongly elongate antennomeres XI. The
<taxonomicName id="FFC84D0A9F18FFD03005FF0AFC2FFE8A" authorityName="LOBL" authorityYear="1981" box="[877,983,241,262]" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="329" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="0ABCEA9B9F18FFD03005FF0AFC2FFE8A" box="[877,983,241,262]" italics="true" pageId="2" pageNumber="329">S. gracilis</emphasis>
</taxonomicName>
and
<taxonomicName id="FFC84D0A9F18FFD03767FF0AFF00FEAF" authorityName="LOBL" authorityYear="1981" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="329" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="0ABCEA9B9F18FFD03767FF0AFF00FEAF" italics="true" pageId="2" pageNumber="329">S. variabilis</emphasis>
</taxonomicName>
described from
<collectingCountry id="40DF76199F18FFD032DAFEF4FE14FEA8" box="[434,492,271,292]" name="Japan" pageId="2" pageNumber="329">Japan</collectingCountry>
have the pronotum and elytra microsculptured and iridescent, unlike those in
<taxonomicName id="FFC84D0A9F18FFD032CBFED7FE0BFECC" authorityName="Löbl" authorityYear="2020" box="[419,499,299,321]" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="329" phylum="Arthropoda" rank="species" species="furva">
<emphasis id="0ABCEA9B9F18FFD032CBFED7FE0BFECC" box="[419,499,299,321]" italics="true" pageId="2" pageNumber="329">S. furva</emphasis>
</taxonomicName>
. In addition, the parameres are narrower and only near apices widened in
<taxonomicName id="FFC84D0A9F18FFD0323FFEB2FE46FED2" authorityName="LOBL" authorityYear="1981" box="[343,446,329,350]" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="329" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="0ABCEA9B9F18FFD0323FFEB2FE46FED2" box="[343,446,329,350]" italics="true" pageId="2" pageNumber="329">S. gracilis</emphasis>
</taxonomicName>
, and the oblique proximal section of the flagellum is much shorter and hardly widened in
<taxonomicName id="FFC84D0A9F18FFD032A4FE9DFDBFFEF7" authorityName="LOBL" authorityYear="1981" box="[460,583,358,379]" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="329" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="0ABCEA9B9F18FFD032A4FE9DFDBFFEF7" box="[460,583,358,379]" italics="true" pageId="2" pageNumber="329">S. variabilis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="70D265029F18FFD033FFFE70FC78FE2D" box="[151,896,395,417]" pageId="2" pageNumber="329" type="etymology">
<paragraph id="387736899F18FFD033FFFE70FC78FE2D" blockId="2.[151,896,395,417]" box="[151,896,395,417]" pageId="2" pageNumber="329">Etymology: The species epithet is a Latin adjective meaning dark.</paragraph>
</subSubSection>
</treatment>
</document>

139
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<paragraph id="387736899F1EFFD633FFFF2AFD49FF6B" blockId="4.[151,689,209,231]" box="[151,689,209,231]" pageId="4" pageNumber="331">
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<emphasis id="0ABCEA9B9F1EFFD633FFFF2AFD49FF6B" bold="true" box="[151,689,209,231]" pageId="4" pageNumber="331">
<taxonomicName id="FFC84D0A9F1EFFD633FFFF2AFE20FF6B" ID-CoL="8Y9D7" authority="Löbl, 2020" authorityName="Löbl" authorityYear="2020" box="[151,472,209,231]" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="331" phylum="Arthropoda" rank="species" species="tengchinensis" status="sp. nov.">
<emphasis id="0ABCEA9B9F1EFFD633FFFF2AFE20FF6B" bold="true" box="[151,472,209,231]" italics="true" pageId="4" pageNumber="331">Scaphobaeocera tengchinensis</emphasis>
</taxonomicName>
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(
<figureCitation id="A0F32A0C9F1EFFD63150FF2AFD51FF6B" box="[568,681,209,231]" captionStart="Figs 8-14" captionStartId="8.[151,191,1496,1518]" captionTargetBox="[169,1050,229,1467]" captionTargetId="figure-79@8.[154,1061,203,1485]" captionTargetPageId="8" captionText="Figs 8-14: Scaphobaeocera, aedeagi; (8, 9) S. montana nov.sp., in dorsal and lateral views, scale = 0.1 mm; (10) Ditto, flagellum and proximal sclerite of internal sac, dorsal view, scle = 0.05 mm; (11, 12) S. tengchinensis nov.sp., in dorsal and lateral views, scale = 0.1 mm; (13, 14) Ditto, flagellum in dorsal and lateral views, scale = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5272846" httpUri="https://zenodo.org/record/5272846/files/figure.png" pageId="4" pageNumber="331">Figs 11-14</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="70D265029F1EFFD633FFFF0CFC66FEA9" pageId="4" pageNumber="331" type="materials_examined">
<paragraph id="387736899F1EFFD633FFFF0CFC66FEA9" blockId="4.[151,1059,247,293]" pageId="4" pageNumber="331">
Type material:
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<typeStatus id="E773882B9F1EFFD632E5FF0CFE27FE81" box="[397,479,247,269]" pageId="4" pageNumber="331" type="holotype">Holotype</typeStatus>
<specimenCount id="2ECEFD009F1EFFD6328CFF0CFE0FFE80" box="[484,503,247,268]" pageId="4" pageNumber="331" type="male">♂</specimenCount>
:
<collectingCountry id="40DF76199F1EFFD63169FF0CFDA3FE80" box="[513,603,247,268]" name="Taiwan" pageId="4" pageNumber="331">TAIWAN</collectingCountry>
<collectingRegion id="FA0CF86B9F1EFFD63108FF0CFD01FE81" box="[608,761,247,269]" pageId="4" pageNumber="331">Kaohsiung Hsien</collectingRegion>
,
<collectingMunicipality id="D813ACF39F1EFFD6306BFF0CFCAEFE81" box="[771,854,247,269]" pageId="4" pageNumber="331">Tengchin</collectingMunicipality>
<quantity id="FF309B6C9F1EFFD63033FF0CFC66FE80" box="[859,926,247,269]" metricMagnitude="3" metricUnit="m" metricValue="1.61" pageId="4" pageNumber="331" unit="m" value="1610.0">
<elevation id="B3E5D1BA9F1EFFD63033FF0CFC66FE80" box="[859,926,247,269]" metricMagnitude="3" metricUnit="m" metricValue="1.61" pageId="4" pageNumber="331" unit="m" value="1610.0">1610 m</elevation>
</quantity>
<date id="4C7610499F1EFFD630CCFF0CFBFFFE80" box="[932,1031,247,268]" pageId="4" pageNumber="331" value="1990-04-24">
<collectingDate id="5C32E9A19F1EFFD630CCFF0CFBFFFE80" box="[932,1031,247,268]" pageId="4" pageNumber="331" value="1990-04-24">24.IV.1990</collectingDate>
</date>
<collectorName id="953D535F9F1EFFD63765FF0CFEE8FEA8" pageId="4" pageNumber="331">A. Smetana</collectorName>
[T20] (
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)
</materialsCitation>
.
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<typeStatus id="E773882B9F1EFFD632C0FEF4FE0DFEA8" box="[424,501,271,292]" pageId="4" pageNumber="331" type="paratype">Paratype</typeStatus>
<specimenCount id="2ECEFD009F1EFFD63291FEF4FDF4FEA8" box="[505,524,271,292]" pageId="4" pageNumber="331" type="male">♂</specimenCount>
: with the same data as the holotype (
<collectionCode id="5ED9AE4C9F1EFFD63024FEF4FC6DFEA9" box="[844,917,271,293]" pageId="4" pageNumber="331">MHNG</collectionCode>
).
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</paragraph>
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Description: Length
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, width
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, dorsoventral diameter
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. Head, pronotum and elytra blackish with weak reddish shine, ventral side of thorax and ventrite I dark brown, abdomen gradually light apically, with apical segments yellowish. Femora reddish-brown, tibiae, tarsi and antennae ochraceous to yellowish. Length/width ratios of antennomeres as: III 22/7: IV 23/7: V 28/7: VI 26/8: VII 33/12: VIII 26/9: IX 33/13: X 34/13: XI 42/15. Pronotum very finely punctate, not microsculptured. Scutellum concealed. Elytra not iridescent, with hardly visible parasutural striae, sutural striae deep, curved at base, discal punctation about as fine as pronotal punctation, microsculpture absent. Hypomera not microsculptured, smooth, lacking longitudinal stria. Middle area of metaventrite flat, entirely densely, distinctly punctate, with punctures to part about as large as puncture intervals; pubescence long, median stria and median impression absent. Lateral parts of metaventrite very finely and sparsely punctate, not microsculptured. Submesocoxal areas about
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, submesocoxal lines distinctly punctate. Metanepisterna about
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wide, flat, not narrowed posteriad, impunctate, with straight suture reaching metepimera. Tibiae straight. Exposed ventrites bearing strigulate microsculpture, very finely punctate, ventrite I with not elongate basal punctures.
</paragraph>
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Male characters: Protarsomeres 1 to 3 widened, distinctly narrower than apices of tibiae. Aedeagus (
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)
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long. Median lobe with basal bulb moderately wide, apical process asymmetrical, strongly inflexed, almost long as basal bulb, ventral side oblique, tip acute, hardly bent in lateral view, blunt in dorsal view. Articular process not prominent. Parameres symmetrical, oblique, reaching posterior level of tip of median lobe, slightly narrowed apically in dorsal view, oblique and widened apically in lateral view. Internal sac with narrow, convoluted flagellum forming two complete loops, widened and sinuate in proximal section (lateral view), capsular in dorsal view.
</paragraph>
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<paragraph id="387736899F1EFFD633FFFBC5FE7FFA88" blockId="4.[151,1058,1086,1284]" pageId="4" pageNumber="331">
Differential diagnosis: This species has an aedeagus similar with that of
<taxonomicName id="FFC84D0A9F1EFFD633DDFBA7FE3DFBFC" authority="LOBL, 1990" authorityName="LOBL" authorityYear="1990" box="[181,453,1114,1137]" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="331" phylum="Arthropoda" rank="species" species="sabapensis">
<emphasis id="0ABCEA9B9F1EFFD633DDFBA7FEB8FBFC" box="[181,320,1115,1137]" italics="true" pageId="4" pageNumber="331">S. sabapensis</emphasis>
LÖBL, 1990
</taxonomicName>
, described from
<collectingCountry id="40DF76199F1EFFD6311EFBA0FD28FBFD" box="[630,720,1115,1137]" name="Thailand" pageId="4" pageNumber="331">Thailand</collectingCountry>
and subsequently reported from the
<collectingCountry id="40DF76199F1EFFD633D4FB82FED6FB03" box="[188,302,1145,1167]" name="Philippines" pageId="4" pageNumber="331">Philippines</collectingCountry>
(
<bibRefCitation id="5C594B789F1EFFD63254FB8CFE46FB03" author="LOBL I" box="[316,446,1143,1167]" pageId="4" pageNumber="331" pagination="695 - 721" refId="ref4208" refString="LOBL I. (2011 b): On the Scaphisomatini (Coleoptera: Staphylinidae: Scaphidiinae) of the Philippines, II. - Revue suisse de Zoologie 118: 695 - 721." type="journal article" year="2011">LÖBL 2011b</bibRefCitation>
). The proximal sections of the flagellum are, however, quite distinctive, being in
<taxonomicName id="FFC84D0A9F1EFFD63204FB6DFE0FFB27" authorityName="LOBL" authorityYear="1990" box="[364,503,1174,1195]" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="331" phylum="Arthropoda" rank="species" species="sabapensis">
<emphasis id="0ABCEA9B9F1EFFD63204FB6DFE0FFB27" box="[364,503,1174,1195]" italics="true" pageId="4" pageNumber="331">S. sabapensis</emphasis>
</taxonomicName>
oblique, weakly widened and not joint to a proximal capsule. The new species may be also distinguished by the median part of the metaventrite not impressed, and the pronotum and elytra not iridescent and lacking distinct microsculpture.
</paragraph>
</subSubSection>
<subSubSection id="70D265029F1EFFD633FFFAE8FC3AFAA5" box="[151,962,1299,1321]" pageId="4" pageNumber="331" type="etymology">
<paragraph id="387736899F1EFFD633FFFAE8FC3AFAA5" blockId="4.[151,962,1299,1321]" box="[151,962,1299,1321]" pageId="4" pageNumber="331">
Etymology: The species epithet is derived from the name of the
<typeStatus id="E773882B9F1EFFD63055FAEFFC91FAA5" box="[829,873,1300,1321]" pageId="4" pageNumber="331">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>

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Key to the
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<emphasis id="0ABCEA9B9F1FFFD7316CFF34FD40FF6B" bold="true" box="[516,696,207,231]" italics="true" pageId="5" pageNumber="332">Scaphobaeocera</emphasis>
</taxonomicName>
of
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<keyLead id="833995BC9F1FFFD733FFFF05FBDAFEA6" pageId="5" pageNumber="332">1 Hypomera each with longitudinal stria separating upper oblique surface from lower surface .................................................................................................................................2</keyLead>
</paragraph>
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<keyLead id="833995BC9F1FFFD733FFFECFFBDAFEC5" box="[151,1058,307,329]" pageId="5" pageNumber="332">- Hypomera lacking longitudinal striae..................................................................................5</keyLead>
</paragraph>
</keyStep>
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<paragraph id="387736899F1FFFD733FFFEA9FBDAFEF2" blockId="5.[151,1059,254,940]" pageId="5" pageNumber="332">
<keyLead id="833995BC9F1FFFD733FFFEA9FBDAFEF2" pageId="5" pageNumber="332">
2 Antennomere XI long about as antennomeres IX and X combined. Aedeagus with flagellum wide, convoluted, forming three complete loops........................
<taxonomicName id="FFC84D0A9F1FFFD730EBFE92FBDAFEF2" ID-CoL="79SY6" authority="LOBL" authorityName="LOBL" box="[899,1058,361,382]" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="332" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="0ABCEA9B9F1FFFD730EBFE92FC10FEF2" box="[899,1000,361,382]" italics="true" pageId="5" pageNumber="332">S. dorsalis</emphasis>
LÖBL
</taxonomicName>
</keyLead>
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<paragraph id="387736899F1FFFD733FFFE73FBDAFE3F" blockId="5.[151,1059,254,940]" pageId="5" pageNumber="332">
<keyLead id="833995BC9F1FFFD733FFFE73FBDAFE3F" pageId="5" pageNumber="332">- Antennomere XI shorter than antennomeres IX and X combined. Aedeagus with flagellum narrow, bent ........................................................................................................3</keyLead>
</paragraph>
</keyStep>
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<keyLead id="833995BC9F1FFFD733FFFE46FBDAFD92" pageId="5" pageNumber="332">
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3 Tip of parameres reaching posterior tip of median lobe of aedeagus, parameres distinctly widened apically in lateral view ..........................................................................4 Tip of median lobe of aedeagus reaching posterior tip of parameres, parameres not or slightly widened apically.................................................................................
<taxonomicName id="FFC84D0A9F1FFFD730F7FDF2FBDAFD92" ID-CoL="6XT9P" authority="LOBL" authorityName="LOBL" box="[927,1058,521,542]" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="332" phylum="Arthropoda" rank="species" species="cyrta">
<emphasis id="0ABCEA9B9F1FFFD730F7FDF2FC10FD92" box="[927,1000,521,542]" italics="true" pageId="5" pageNumber="332">S. cyrta</emphasis>
LÖBL
</taxonomicName>
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4 Proximal sclerites of internal sac narrow, with prominent apophysis............
<taxonomicName id="FFC84D0A9F1FFFD730FBFDD3FBDAFDB1" ID-CoL="6Y4Y6" authority="LOBL" authorityName="LOBL" box="[915,1058,551,573]" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="332" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="0ABCEA9B9F1FFFD730FBFDD3FC10FDB0" box="[915,1000,551,573]" italics="true" pageId="5" pageNumber="332">S. dispar</emphasis>
LÖBL
</taxonomicName>
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<keyLead id="833995BC9F1FFFD733FFFDBDFBDBFDFE" pageId="5" pageNumber="332">
- Proximal sclerites of internal sac wide, with broadly rounded anterior margin, lacking apophysis................................................................................................
<taxonomicName id="FFC84D0A9F1FFFD73003FDA6FC2FFDFE" ID-CoL="8Y9D6" authority="Löbl, 2020" authorityName="Löbl" authorityYear="2020" box="[875,983,605,626]" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="332" phylum="Arthropoda" rank="species" species="montana" status="sp. nov.">
<emphasis id="0ABCEA9B9F1FFFD73003FDA6FC2FFDFE" box="[875,983,605,626]" italics="true" pageId="5" pageNumber="332">S. montana</emphasis>
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<keyLead id="833995BC9F1FFFD733FFFD80FBDAFD1D" box="[151,1058,635,657]" pageId="5" pageNumber="332">
5 Metaventrite with elongate median impression ...................
<taxonomicName id="FFC84D0A9F1FFFD7319DFD87FBDAFD1D" ID-CoL="6Y4Y7" authority="MIWA &amp; MITONO" authorityName="MIWA &amp; MITONO" box="[757,1058,635,657]" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="332" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="0ABCEA9B9F1FFFD7319DFD87FC8CFD1C" box="[757,884,635,657]" italics="true" pageId="5" pageNumber="332">S. formosana</emphasis>
MIWA &amp; MITONO
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="387736899F1FFFD733FFFD61FBDAFD3C" blockId="5.[151,1059,254,940]" box="[151,1058,666,688]" pageId="5" pageNumber="332">
<keyLead id="833995BC9F1FFFD733FFFD61FBDAFD3C" box="[151,1058,666,688]" pageId="5" pageNumber="332">- Metaventrite without median impression ............................................................................6</keyLead>
</paragraph>
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<paragraph id="387736899F1FFFD733FFFD42FBDBFD42" blockId="5.[151,1059,254,940]" box="[151,1059,697,719]" pageId="5" pageNumber="332">
<keyLead id="833995BC9F1FFFD733FFFD42FBDBFD42" box="[151,1059,697,719]" pageId="5" pageNumber="332">
6 Aedeagus with flagellum forming three complete loops ..............................
<taxonomicName id="FFC84D0A9F1FFFD730E4FD42FC20FD42" ID-CoL="8Y9CZ" authority="Löbl, 2020" authorityName="Löbl" authorityYear="2020" box="[908,984,697,718]" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="332" phylum="Arthropoda" rank="species" species="furva" status="sp. nov.">
<emphasis id="0ABCEA9B9F1FFFD730E4FD42FC20FD42" box="[908,984,697,718]" italics="true" pageId="5" pageNumber="332">S. furva</emphasis>
</taxonomicName>
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</keyLead>
</paragraph>
<paragraph id="387736899F1FFFD733FFFD23FBDAFD61" blockId="5.[151,1059,254,940]" box="[151,1058,727,749]" pageId="5" pageNumber="332">
<keyLead id="833995BC9F1FFFD733FFFD23FBDAFD61" box="[151,1058,727,749]" pageId="5" pageNumber="332">- Aedeagus with flagellum forming two complete loops .......................................................7</keyLead>
</paragraph>
</keyStep>
<keyStep id="833C2E2C9F1FFFD733FFFD0DFBDAFCCC" pageId="5" pageNumber="332">
<paragraph id="387736899F1FFFD733FFFD0DFBDBFCAE" blockId="5.[151,1059,254,940]" pageId="5" pageNumber="332">
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7 Aedeagus with proximal section of flagellum folded and evenly wide ................................. ...............................................................................................................
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<keyLead id="833995BC9F1FFFD733FFFCD7FBDAFCCC" box="[151,1058,811,833]" pageId="5" pageNumber="332">- Aedeagus with proximal section of flagellum not folded and widened...............................8</keyLead>
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8 Aedegus with apices of parameres slightly widened (in lateral view), flagellum slightly widened and simple in proximal section ........................................
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sp. near
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LÖBL
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- Aedegus with parameres gradually widened apically (in lateral view), flagellum strongly widened proximally and with bulbous vesicle...................
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<emphasis id="0ABCEA9B9F1FFFD73054FC6DFC2FFC27" box="[828,983,918,939]" italics="true" pageId="5" pageNumber="332">S. tengchinensis</emphasis>
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.
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Phalangopsites
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Phalangopsitidae
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.
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genus:
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<emphasis id="0ABCEAA7FFCC8D012A40CC552DE8F9BA" box="[806,914,1544,1565]" italics="true" pageId="1" pageNumber="57">Remarks:</emphasis>
Priority for family-group names based on the
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dates from Phalangopsites
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, and not from Phalangopsites
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, as has been widely purported in the literature.
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TRIBE
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LUZARINI
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Luzarae
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Luzarinae
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genus: Luzara
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<emphasis id="0ABCEAA7FFCF8D0229C2CA9B2F75FF7C" box="[164,271,198,219]" italics="true" pageId="2" pageNumber="58">Remarks:</emphasis>
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defined the
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based on the morphology of the male genitalia, stating that ‘chez les
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[
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], toute la partie médiane de l’épiphalle est régressée, notamment les lophi médians; d’autre part, les “bras épiphalliques” ainsi formés acquièrent une structure complex, avec généralement une ou deux zones de rupture ou d’articulation et des épines latérales’. The retrograde median region of the epiphallus is unique amongst
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, and is likely to be apomorphic. The complex ‘bras épiphalliques’, or ‘epiphallic arms’, and their lateral lobes are highly variable: their morphology and spatial associations with the epiphallic parameres have proved very useful at lower taxonomic levels (e.g.
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,
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;
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Amphiacustae
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<emphasis id="0ABCEAA7FFCF8D022855C9152FB3FCFA" box="[307,457,840,861]" italics="true" pageId="2" pageNumber="58">Amphiacusta</emphasis>
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<emphasis id="0ABCEAA7FFCF8D0229C2C93A2F75FCDB" box="[164,271,871,892]" italics="true" pageId="2" pageNumber="58">Remarks:</emphasis>
The priority of family-group names based on the
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dates from Amphiacustae
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. This name has subsequently been used to refer to a group of genera comprising
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<emphasis id="0ABCEAA7FFCF8D0229C2C9BC2F40FC51" box="[164,314,993,1014]" italics="true" pageId="2" pageNumber="58">Amphiacusta</emphasis>
</taxonomicName>
,
<taxonomicName id="FFC84D36FFCF8D02282EC9BC2F9AFC51" authorityName="Desutter-Grandcolas &amp; Hubbell" authorityYear="1993" box="[328,480,993,1014]" class="Insecta" family="Haglotettigoniidae" genus="Arachnopsita" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="0ABCEAA7FFCF8D02282EC9BC2F9AFC51" box="[328,480,993,1014]" italics="true" pageId="2" pageNumber="58">Arachnopsita</emphasis>
</taxonomicName>
,
<taxonomicName id="FFC84D36FFCF8D022888C9BC2C1FFC51" authorityName="Desutter-Grandcolas" authorityYear="1994" box="[494,613,993,1014]" class="Insecta" family="Haglotettigoniidae" genus="Cantrallia" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="0ABCEAA7FFCF8D022888C9BC2C1FFC51" box="[494,613,993,1014]" italics="true" pageId="2" pageNumber="58">Cantrallia</emphasis>
</taxonomicName>
,
<taxonomicName id="FFC84D36FFCF8D022B12C9BC2D7EFC51" authorityName="Desutter-Grandcolas" authorityYear="1994" box="[628,772,993,1014]" class="Insecta" family="Haglotettigoniidae" genus="Leptopedetes" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="0ABCEAA7FFCF8D022B12C9BC2D7EFC51" box="[628,772,993,1014]" italics="true" pageId="2" pageNumber="58">Leptopedetes</emphasis>
</taxonomicName>
,
<taxonomicName id="FFC84D36FFCF8D0229C2CE5C2F59FBB1" authorityName="Desutter-Grandcolas &amp; Hubbell" authorityYear="1993" box="[164,291,1025,1046]" class="Insecta" family="Haglotettigoniidae" genus="Longuripes" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="0ABCEAA7FFCF8D0229C2CE5C2F59FBB1" box="[164,291,1025,1046]" italics="true" pageId="2" pageNumber="58">Longuripes</emphasis>
</taxonomicName>
,
<taxonomicName id="FFC84D36FFCF8D022859CE5D2FB4FBB2" authorityName="Desutter-Grandcolas &amp; Hubbell" authorityYear="1993" box="[319,462,1024,1045]" class="Insecta" family="Haglotettigoniidae" genus="Mayagryllus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="0ABCEAA7FFCF8D022859CE5D2FB4FBB2" box="[319,462,1024,1045]" italics="true" pageId="2" pageNumber="58">Mayagryllus</emphasis>
</taxonomicName>
,
<taxonomicName id="FFC84D36FFCF8D02288CCE5C2CFCFBB1" authorityName="Desutter-Grandcolas &amp; Hubbell" authorityYear="1993" box="[490,646,1025,1046]" class="Insecta" family="Haglotettigoniidae" genus="Nemoricantor" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="0ABCEAA7FFCF8D02288CCE5C2CFCFBB1" box="[490,646,1025,1046]" italics="true" pageId="2" pageNumber="58">Nemoricantor</emphasis>
</taxonomicName>
,
<taxonomicName id="FFC84D36FFCF8D022BC4CE5C2D7EFBB1" authorityName="Desutter-Grandcolas &amp; Hubbell" authorityYear="1993" box="[674,772,1025,1046]" class="Insecta" family="Haglotettigoniidae" genus="Noctivox" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="0ABCEAA7FFCF8D022BC4CE5C2D7EFBB1" box="[674,772,1025,1046]" italics="true" pageId="2" pageNumber="58">Noctivox</emphasis>
</taxonomicName>
, and
<taxonomicName id="FFC84D36FFCF8D0229B0CE422F0FFB93" authorityName="Desutter-Grandcolas" authorityYear="1993" box="[214,373,1055,1076]" class="Insecta" family="Phalangopsidae" genus="Prolonguripes" higherTaxonomySource="CoL" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="0ABCEAA7FFCF8D0229B0CE422F0FFB93" box="[214,373,1055,1076]" italics="true" pageId="2" pageNumber="58">Prolonguripes</emphasis>
</taxonomicName>
(
<bibRefCitation id="5C594B44FFCF8D0228E2CE422CC3FB93" author="Desutter-Grandcolas L" box="[388,697,1055,1077]" pageId="2" pageNumber="58" pagination="1 - 82" refId="ref5975" refString="Desutter-Grandcolas L. 1993. The cricket fauna of Chipanecan caves (Mexico), systematics, phylogeny and the evolution towards troglobitic life (Orthoptera, Grylloidea, Phalangopsidae, Luzarinae). International Journal of Speleology 22: 1 - 82." type="journal article" year="1993">Desutter-Grandcolas, 1993</bibRefCitation>
,
<bibRefCitation id="5C594B44FFCF8D022BA3CE422C85FB92" author="Desutter-Grandcolas L" box="[709,767,1055,1077]" pageId="2" pageNumber="58" pagination="1 - 19" refId="ref6090" refString="Desutter-Grandcolas L. 1999. Are troglobitic taxa troglobiomorphic? A tes using phylogenetic inference. International Journal of Speleology 26: 1 - 19." type="journal article" year="1999">1999</bibRefCitation>
). The monophyly of this group is well supported by the apical migration of sclerites in the epiphallic arms (termed the ‘C-sclerites’ by
<bibRefCitation id="5C594B44FFCF8D022B71CE272EA1FB08" author="Desutter-Grandcolas L" pageId="2" pageNumber="58" pagination="1 - 82" refId="ref5975" refString="Desutter-Grandcolas L. 1993. The cricket fauna of Chipanecan caves (Mexico), systematics, phylogeny and the evolution towards troglobitic life (Orthoptera, Grylloidea, Phalangopsidae, Luzarinae). International Journal of Speleology 22: 1 - 82." type="journal article" year="1993">Desutter-Grandcolas, 1993</bibRefCitation>
) and the endophallic apodeme without a mediodorsal crest. The group includes some 84 species in the aforementioned genera, distributed throughout Central America and the West Indies.
</paragraph>
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<bibRefCitation id="5C594B44FFCF8D0229DACF4E2F0AFA8D" author="Hubbell TH" box="[188,368,1299,1322]" pageId="2" pageNumber="58" pagination="191 - 233" refId="ref6412" refString="Hubbell TH. 1938. New cave-crickets from Yucatan, with a review of the Pentacentrinae, and studies on the genus Amphiacusta (Orthoptera: Gryllidae). Carnegie Institute of Washington Publication 491: 191 - 233." type="journal article" year="1938">Hubbell (1938)</bibRefCitation>
formed the name Amphiacustae using his standard suffix for family-group names beneath the rank of subfamily and although the group has been intensively studied since then (
<bibRefCitation id="5C594B44FFCF8D0229CDCFD32F90FA04" author="Desutter-Grandcolas L" box="[171,490,1422,1444]" pageId="2" pageNumber="58" pagination="1 - 82" refId="ref5975" refString="Desutter-Grandcolas L. 1993. The cricket fauna of Chipanecan caves (Mexico), systematics, phylogeny and the evolution towards troglobitic life (Orthoptera, Grylloidea, Phalangopsidae, Luzarinae). International Journal of Speleology 22: 1 - 82." type="journal article" year="1993">Desutter-Grandcolas, 1993</bibRefCitation>
, 1997, 1999; Desutter- Grandcolas &amp; Otte, 1997), it has never been assigned a rank. Thus,
<bibRefCitation id="5C594B44FFCF8D022820CF962C7BFA46" author="Hubbell TH" box="[326,513,1483,1505]" pageId="2" pageNumber="58" pagination="191 - 233" refId="ref6412" refString="Hubbell TH. 1938. New cave-crickets from Yucatan, with a review of the Pentacentrinae, and studies on the genus Amphiacusta (Orthoptera: Gryllidae). Carnegie Institute of Washington Publication 491: 191 - 233." type="journal article" year="1938">Hubbell’s (1938)</bibRefCitation>
original spelling of the name (using the suffix -ae) has never been questioned. The suffixes of family-group names are regulated by the International Code of Zoological Nomenclature (
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) at the ranks of superfamily, family, subfamily, tribe, and subtribe (
<collectionCode id="5ED9AE70FFCF8D0228FDCC392FA4F9DE" box="[411,478,1636,1657]" pageId="2" pageNumber="58">ICZN</collectionCode>
, 1999: article 29.2). The suffixes of names at other ranks within the family group are not formally regulated, but do have widely accepted standard suffixes (e.g. -oidae for an epifamily, -iti for a supertribe, -ita for an infratribe, etc.). The suffixes of rankless taxa are likewise unregulated by the
<collectionCode id="5ED9AE70FFCF8D02299ECD412F40F896" box="[248,314,1820,1841]" pageId="2" pageNumber="58">ICZN</collectionCode>
, and their use has become increasingly desirable in recent years because of the widespread use of cladistic methodology and the need to name important nodes within phylogenies; particularly in cases where the assignment of a specific rank would be superfluous. Nevertheless, the use of the name Amphiacustae as a rankless taxon retaining its original suffix is somewhat problematic because of the similarity of the name with that of the
<typeStatus id="E7738817FFCF8D022C7CCB022B31FED3" box="[1306,1355,351,372]" pageId="2" pageNumber="58">type</typeStatus>
genus. This has the potential to cause confusion, and so the group is here assigned the rank of subtribe and the suffix is emended to ‘-ina’, in accordance with
<collectionCode id="5ED9AE70FFCF8D022C04CBE62BDBFE77" box="[1378,1441,443,464]" pageId="2" pageNumber="58">ICZN</collectionCode>
regulations.
</paragraph>
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77
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GENUS
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<emphasis id="0ABCEAA7FFCF8D022A5CC8092DC2FDCE" box="[826,952,596,617]" italics="true" pageId="2" pageNumber="58">Etymology:</emphasis>
The new genus-group name is formed from a combination of the Latin words
<emphasis id="0ABCEAA7FFCF8D022C2DC82E2BE0FD2F" box="[1355,1434,627,648]" italics="true" pageId="2" pageNumber="58">aranea</emphasis>
, meaning ‘spider’, and
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, meaning ‘cricket’: literally ‘spider cricket’, in reference to the vernacular name of phalangopsines. The gender is masculine.
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species:
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<emphasis id="0ABCEAA7FFCF8D022A5CC9512DCBFC86" box="[826,945,780,801]" italics="true" pageId="2" pageNumber="58">Diagnosis:</emphasis>
Female: head with comparatively small fastigium, protruding only slightly beyond the antennal fossae and in line with the vertex; subocular furrow prominent, distally curved, meeting weak frontal costa near frontoclypeal suture/anterior tentorial pit; compound eyes suboval; ocelli absent; pronotum wider than long, saddle-shaped, with prominent dark band near pleural margins and rows of long marginal setae; procoxa markedly more robust than mesocoxa; mesotibia with two apical spurs; metatibia with two parallel rows of between ten and 12 small dorsal spines, four long and robust subapical spurs, and three apical spurs (median one markedly longer than the others); subgenital plate broadly triangular, with a small but distinct median notch in the posterior margin. Male: unknown.
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<subSubSection id="70D2653EFFCF8D022A07CF772B00FAE7" box="[865,1402,1322,1344]" pageId="2" pageNumber="58" type="nomenclature">
<paragraph id="387736B5FFCF8D022A07CF772B00FAE7" blockId="2.[865,1402,1322,1344]" box="[865,1402,1322,1344]" pageId="2" pageNumber="58">
<heading id="633F81D9FFCF8D022A07CF772B00FAE7" box="[865,1402,1322,1344]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="58" reason="4">
<taxonomicName id="FFC84D36FFCF8D022A07CF772AF3FA98" authorityName="HEADS" authorityYear="2010" box="[865,1161,1322,1343]" class="Insecta" family="Haglotettigoniidae" genus="Araneagryllus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="58" phylum="Arthropoda" rank="species" species="dylani">ARANEAGRYLLUS DYLANI</taxonomicName>
<taxonomicNameLabel id="118F57DCFFCF8D022DF2CF702A83FA98" box="[1172,1273,1322,1343]" pageId="2" pageNumber="58" rank="species">
<emphasis id="0ABCEAA7FFCF8D022DF2CF702ACDFA98" bold="true" box="[1172,1207,1322,1343]" pageId="2" pageNumber="58">SP.</emphasis>
<emphasis id="0ABCEAA7FFCF8D022DA7CF702A83FA98" bold="true" box="[1217,1273,1322,1343]" pageId="2" pageNumber="58">NOV.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="A0F32A30FFCF8D022C6DCF772B0BFAE7" box="[1291,1393,1322,1344]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="3.[144,223,1044,1063]" captionStartId-1="4.[164,243,1278,1297]" captionStartId-2="5.[144,223,1132,1151]" captionTargetBox-0="[144,761,195,1015]" captionTargetBox-1="[378,1232,197,1246]" captionTargetBox-2="[144,758,198,1101]" captionTargetId-0="figure-579@3.[144,761,195,1015]" captionTargetId-1="figure-264@4.[375,1232,195,1249]" captionTargetId-2="figure-137@5.[144,760,196,1103]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="Figure 1. Araneagryllus dylani gen. et sp. nov., photomicrograph of holotype AMNH DR-12-32. Scale bar: 3 mm." captionText-1="Figure 2. Araneagryllus dylani gen. et sp. nov., camera lucida drawings of holotype AMNH DR-12-32. A, general habitus of specimen. Scale bar: 3 mm. B, left lateral view of head. Scale bar: 1 mm." captionText-2="Figure 3. Schematic reconstructions of the legs of Araneagryllus dylani gen. et sp. nov. showing the tibial armature and the distribution of the colour pattern. A, prothoracic leg. B, mesothoracic leg. C, metathoracic leg. Scale bar: 3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4720038" figureDoi-1="http://doi.org/10.5281/zenodo.4720042" figureDoi-2="http://doi.org/10.5281/zenodo.4720044" httpUri-0="https://zenodo.org/record/4720038/files/figure.png" httpUri-1="https://zenodo.org/record/4720042/files/figure.png" httpUri-2="https://zenodo.org/record/4720044/files/figure.png" pageId="2" pageNumber="58">FIGS 1–3</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="70D2653EFFCF8D032A5CCF0C2DA0FA46" lastPageId="3" lastPageNumber="59" pageId="2" pageNumber="58" type="etymology">
<paragraph id="387736B5FFCF8D022A5CCF0C2AC2FA03" blockId="2.[826,1442,1361,1903]" pageId="2" pageNumber="58">
<emphasis id="0ABCEAA7FFCF8D022A5CCF0C2DC2FAC1" box="[826,952,1361,1382]" italics="true" pageId="2" pageNumber="58">Etymology:</emphasis>
The specific epithet is a noun in the genitive case, and is named in honour of my son, Dylan L. Heads. The gender is masculine.
</paragraph>
<paragraph id="387736B5FFCF8D022A5CCFF02AE1F910" blockId="2.[826,1442,1361,1903]" pageId="2" pageNumber="58">
<emphasis id="0ABCEAA7FFCF8D022A5CCFF02D0FFA65" box="[826,885,1453,1474]" italics="true" pageId="2" pageNumber="58">Type:</emphasis>
<materialsCitation id="88A03CE8FFCF8D022AE3CFF12AE1F910" collectionCode="AMNH" country="Dominican Republic" location="Early Miocene" municipality="Dominican" pageId="2" pageNumber="58" specimenCount="1" typeStatus="holotype">
<typeStatus id="E7738817FFCF8D022AE3CFF12D91FA65" box="[901,1003,1452,1474]" pageId="2" pageNumber="58" type="holotype">Holotype</typeStatus>
female:
<collectingCountry id="40DF7625FFCF8D022D3BCFF12B34FA65" box="[1117,1358,1452,1474]" name="Dominican Republic" pageId="2" pageNumber="58">Dominican Republic</collectingCountry>
,
<location id="3D17606EFFCF8D022C04CFF12DE2FA46" LSID="urn:lsid:plazi:treatment:B06187A3FFCF8D032A07CF772DA0FA46:3D17606EFFCF8D022C04CFF12DE2FA46" country="Dominican Republic" municipality="Dominican" name="Early Miocene" pageId="2" pageNumber="58">Early Miocene</location>
(Burdigalian, 18–20 Mya)
<collectingMunicipality id="D813ACCFFFCF8D022DA8CF962B30FA46" box="[1230,1354,1483,1505]" pageId="2" pageNumber="58">Dominican</collectingMunicipality>
amber;
<collectionCode id="5ED9AE70FFCF8D022A5CCFB72DF6FA58" box="[826,908,1514,1535]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="2" pageNumber="58">AMNH</collectionCode>
specimen number DR-12-32. Syninclusions: this piece of amber is particularly rich in biotic inclusions, including an isopod, a thysanuran, a collembolan, an auchenorrhynchan, two chalcidoid wasps, and numerous Diptera. The specimen is notable for the development of a white emulsion, particularly around the abdomen and mouthparts.
</materialsCitation>
</paragraph>
<paragraph id="387736B5FFCF8D022A5CCC9D2B62F972" blockId="2.[826,1442,1361,1903]" box="[826,1304,1728,1750]" pageId="2" pageNumber="58">
<emphasis id="0ABCEAA7FFCF8D022A5CCC9D2DCBF972" box="[826,945,1728,1749]" italics="true" pageId="2" pageNumber="58">Diagnosis:</emphasis>
As for the genus (
<emphasis id="0ABCEAA7FFCF8D022DE9CC9D2B73F972" box="[1167,1289,1728,1749]" italics="true" pageId="2" pageNumber="58">vide supra</emphasis>
).
</paragraph>
<paragraph id="387736B5FFCF8D032A5CCC822DA0FA46" blockId="2.[826,1442,1361,1903]" lastBlockId="3.[806,1422,198,1505]" lastPageId="3" lastPageNumber="59" pageId="2" pageNumber="58">
<emphasis id="0ABCEAA7FFCF8D022A5CCC822DB8F953" box="[826,962,1759,1780]" italics="true" pageId="2" pageNumber="58">Description:</emphasis>
Female: body form robust (
<figureCitation id="A0F32A30FFCF8D022C7FCC832B25F953" box="[1305,1375,1758,1780]" captionStart="Figure 1" captionStartId="3.[144,223,1044,1063]" captionTargetBox="[144,761,195,1015]" captionTargetId="figure-579@3.[144,761,195,1015]" captionTargetPageId="3" captionText="Figure 1. Araneagryllus dylani gen. et sp. nov., photomicrograph of holotype AMNH DR-12-32. Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.4720038" httpUri="https://zenodo.org/record/4720038/files/figure.png" pageId="2" pageNumber="58">Figs 1</figureCitation>
,
<figureCitation id="A0F32A30FFCF8D022C14CC822BEFF952" box="[1394,1429,1759,1781]" captionStart="Figure 2" captionStartId="4.[164,243,1278,1297]" captionTargetBox="[378,1232,197,1246]" captionTargetId="figure-264@4.[375,1232,195,1249]" captionTargetPageId="4" captionText="Figure 2. Araneagryllus dylani gen. et sp. nov., camera lucida drawings of holotype AMNH DR-12-32. A, general habitus of specimen. Scale bar: 3 mm. B, left lateral view of head. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4720042" httpUri="https://zenodo.org/record/4720042/files/figure.png" pageId="2" pageNumber="58">2A</figureCitation>
); 12.11-mm long, measured from head to abdominal apex (excluding ovipositor). Head hypognathous (
<figureCitation id="A0F32A30FFCF8D022A24CD672DEDF8F7" box="[834,919,1850,1872]" captionStart="Figure 2" captionStartId="4.[164,243,1278,1297]" captionTargetBox="[378,1232,197,1246]" captionTargetId="figure-264@4.[375,1232,195,1249]" captionTargetPageId="4" captionText="Figure 2. Araneagryllus dylani gen. et sp. nov., camera lucida drawings of holotype AMNH DR-12-32. A, general habitus of specimen. Scale bar: 3 mm. B, left lateral view of head. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4720042" httpUri="https://zenodo.org/record/4720042/files/figure.png" pageId="2" pageNumber="58">Fig. 2B</figureCitation>
); vertex broad, with numerous dark spots extending towards the fastigium, which protrudes between the antennal fossae and is covered with a patch of short setae; frons gently sloping; frontal costa shallow, indistinct; subocular furrow distinct, curved medially towards the frontoclypeal suture; genae almost triangular in shape, covered in minute pits, with two dark spots beneath the compound eyes and a much smaller dark spot towards its ventral margin; compound eyes well developed, globose, situated high on the head, and protuding laterally; ocelli absent; antennal fossae situated directly anterior of the compound eyes; scape large, longer than wide and pedicel markedly smaller, only slightly longer than wide; first, second, and third flagellomeres small, as long as wide; subsequent flagellomeres markedly longer than wide; clypeus, labrum, and other mouthparts largely obscured by white emulsion; mandible apparently strong; maxillary palp long, terminal palpomere markedly inflated apically; labial palp smaller, largely obscured by emulsion. Pronotum shield-like, wider than long, with prominent median and marginal carinae and irregularly spaced short setae; pleural margin with prominent dark band and pale ‘lip’ on anterior pleural lobe; median carina with dark stripe, bordered laterally by several dark spots; anterior and posterior margins of pronotum with prominent rows of long setae. Meso- and metanotae and abdominal tergites broad, decreasing in size posteriorly and of largely uniform dark coloration; posterior margins of abdominal tergites with rows of setae. Abdomen largely obscured by emulsion; cerci incomplete (truncated by surface of amber), broad basally, narrowing apically, with a dense covering of setae; epiproct triangular, covered with emulsion; ovipositor incomplete (truncated by surface of amber), strong basally, narrowing apically; subgenital plate broad basally, narrowing slightly posteriorly, with a small median posterior notch. Pro- and mesothoracic legs long, robust (
<figureCitation id="A0F32A30FFCE8D032A19C8092D8EFDCE" box="[895,1012,596,618]" captionStart="Figure 3" captionStartId="5.[144,223,1132,1151]" captionTargetBox="[144,758,198,1101]" captionTargetId="figure-137@5.[144,760,196,1103]" captionTargetPageId="5" captionText="Figure 3. Schematic reconstructions of the legs of Araneagryllus dylani gen. et sp. nov. showing the tibial armature and the distribution of the colour pattern. A, prothoracic leg. B, mesothoracic leg. C, metathoracic leg. Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.4720044" httpUri="https://zenodo.org/record/4720044/files/figure.png" pageId="3" pageNumber="59">Fig. 3A, B</figureCitation>
). Procoxa large, broadly articulated with prosternum; protrochanter much smaller, largely obscured by a large bubble; profemur broad basally, narrowing very slightly towards apex, covered in fine setae and numerous irregular dark patches; numerous longer setae on the dorsal surface, particularly distally; genicular lobes prominent, inflected; protibia a little over half the length of profemur, with three dark bands and two apical spurs of equal length; tympana absent; probasitarsus three-quarters as long as protibia, with prominent apical spine; second tarsomere small; third tarsomere long, gracile. Mesocoxa approximately half the size of procoxa; mesotrochanter small, triangular; mesofemur similar to profemur, with irregular dark patches, prominent genicular lobes, but with more numerous long setae; mesotibia a little over half the length of mesofemur, with three dark bands and two apical spurs of equal length; tarsus as in the prothoracic leg. Metathoracic leg long, robust, saltatorial (
<figureCitation id="A0F32A30FFCE8D032DFCCEC42A94FB08" box="[1178,1262,1177,1199]" captionStart="Figure 3" captionStartId="5.[144,223,1132,1151]" captionTargetBox="[144,758,198,1101]" captionTargetId="figure-137@5.[144,760,196,1103]" captionTargetPageId="5" captionText="Figure 3. Schematic reconstructions of the legs of Araneagryllus dylani gen. et sp. nov. showing the tibial armature and the distribution of the colour pattern. A, prothoracic leg. B, mesothoracic leg. C, metathoracic leg. Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.4720044" httpUri="https://zenodo.org/record/4720044/files/figure.png" pageId="3" pageNumber="59">Fig. 3C</figureCitation>
); metafemur strongly inflated basally, with numerous dark patches along the outer facets and dark bands distally; metatibia equal in length to metafemur, with four dark bands; denticulate with two parallel rows of between ten and 12 short spines and four long subapical spurs; three apical spurs, with the median spur markedly longer; metabasitarsus long, with a single dark band, and one long apical spur; second tarsomere much reduced; third tarsomere long, gracile, slightly curved.
</paragraph>
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42
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28.
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. B.M.
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<paragraph id="F58E3F61124C11F59A55B30858EA4C31" pageId="168" pageNumber="168">Female. Length 3 1/2 lines.-Pale ferruginous and opake, with the scape, basal joints of the flagellum, and the legs pale; the apical joints of the antennae and the margins of the mandibles dusky. The head and thorax very evenly and delicately striated; the wings smoky, palest at their base, with the nervures fuscous; the second and following segments of the abdomen shining. Worker. Length 1 1/4 line.-Pale rufo-testaceous, not shining: head oblong-ovate, with the mandibles bent, and very acute at their apex, their inner margin furnished with acute teeth; the thorax narrowed, and nearly parallel behind the prothorax; the metathorax armed with two upright acute spines. Abdomen ovate, truncate at the base, and more or less fuscous at the apex.</paragraph>
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<paragraph id="C0D708A6DC8D495F84A303B23B0715DD" pageId="168" pageNumber="168">Hab. Rio. (Coll. Rev. Hamlet Clark.)</paragraph>
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