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<document id="CE5D9957154882338A174292202DBCFE" ID-DOI="10.5852/ejt.2020.671" ID-GBIF-Dataset="6f6af3d0-601d-47dc-849d-8ad21170e570" ID-ISSN="2118-9773" ID-Zenodo-Dep="3906756" ID-ZooBank="9EF19154-D2FE-4009-985C-1EB62CC9ACB0" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1593020698292" checkinUser="valdenar" docAuthor="D., Guinot & de, Mazancourt V." docDate="2020" docId="0393774D4008B22364E51B7DFB42F8FE" docLanguage="en" docName="ejt-671_guinot_mazancourt.pdf.imf" docOrigin="European Journal of Taxonomy 671 (671)" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Richerius marqueti Isyeb 2020, gen. et sp. nov." docType="treatment" docVersion="1" lastPageNumber="14" masterDocId="FFAA0F35400DB22E64581F42FF88FFBC" masterDocTitle="A new freshwater crab of the family Hymenosomatidae MacLeay, 1838 from New Caledonia (Crustacea, Decapoda, Brachyura) and an updated review of the hymenosomatid fauna of New Caledonia" masterLastPageNumber="29" masterPageNumber="1" pageNumber="6" updateTime="1733486495029" updateUser="guilherme" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="22B55FCAE6D94FBC32A185A8AE7193D0">A new freshwater crab of the family Hymenosomatidae MacLeay, 1838 from New Caledonia (Crustacea, Decapoda, Brachyura) and an updated review of the hymenosomatid fauna of New Caledonia</mods:title>
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<mods:namePart id="5DB9DABC48FA3DC3F74750D23A868B76">D., Guinot</mods:namePart>
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<mods:affiliation id="272BB0E582F742AE6988767D449D39B1">ISYEB (CNRS, MNHN, EPHE, Sorbonne Université), Institut Systématique Évolution Biodiversité, Muséum national d’histoire naturelle, case postale 53, 57 rue Cuvier, 75231 Paris cedex 05, France</mods:affiliation>
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<mods:nameIdentifier id="927A8ABCB4F4F4DFA0B8DA46F5D8252D" type="email">guinot@mnhn.fr</mods:nameIdentifier>
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<mods:roleTerm id="BF431CB64DF76D56ED0CB530016673C5">Author</mods:roleTerm>
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<mods:namePart id="611620FA8E44AD07BD01BCD749EB1106">de, Mazancourt V.</mods:namePart>
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<mods:affiliation id="CE12AB91CBAC3D950C03BF2C02834304">Unité Biologie des organismes et écosystèmes aquatiques (BOREA), Muséum national d’histoire naturelle, Sorbonne Université, Université de Caen Normandie, Université des Antilles, CNRS, IRD, CP 26, 57 rue Cuvier, 75231 Paris cedex 05, France & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Invalidenstrasse 43, 10115 Berlin. Germany.</mods:affiliation>
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<mods:nameIdentifier id="BA4102C9C7E5593B079E96886C195583" type="email">valentin.seizilles-de-mazancourt@edu.mnhn.fr</mods:nameIdentifier>
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<mods:title id="F7EDFDA278B887AE1187A9C4A780A034">European Journal of Taxonomy</mods:title>
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<mods:part id="49361D2AFC87FA431EFF703ECFA76354">
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<mods:date id="565A7C99AAC20772CB926B1ABFF0CD53">2020</mods:date>
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<mods:number id="12A82EFCA0618CC7BF51C134F9FE4EAC">2020-06-22</mods:number>
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<treatment id="0393774D4008B22364E51B7DFB42F8FE" ID-GBIF-Taxon="164723367" ID-Zenodo-Dep="7433864" LSID="urn:lsid:plazi:treatment:0393774D4008B22364E51B7DFB42F8FE" httpUri="http://treatment.plazi.org/id/0393774D4008B22364E51B7DFB42F8FE" includeIn="http://doi.org/10.5281/zenodo.7433864" lastPageId="13" lastPageNumber="14" pageId="5" pageNumber="6" scope_family="Hymenosomatidae" scope_infraOrder="Brachyura" scope_order="Decapoda" scope_subFamily="Odiomarinae" scope_superFamily="Hymenosomatoidea">
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<subSubSection id="C32095D04008B22564E51B7DFBCAFE60" lastPageId="11" lastPageNumber="12" pageId="5" pageNumber="6" type="description">
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<paragraph id="8B85C65B4008B22B64E51B7DFEC6FBE5" blockId="5.[189,334,1087,1113]" box="[189,334,1087,1113]" pageId="5" pageNumber="6">
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<heading id="D0CD71374008B22B64E51B7DFEC6FBE5" bold="true" box="[189,334,1087,1113]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
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<emphasis id="B94E1A494008B22B64E51B7DFEC6FBE5" bold="true" box="[189,334,1087,1113]" pageId="5" pageNumber="6">Description</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B85C65B4008B22B64E51B2CFE5BFA8B" blockId="5.[189,1399,1133,1335]" pageId="5" pageNumber="6">
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Carapace (
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<figureCitation id="1301DADE4008B22B65661B2FFE2CFB34" box="[318,420,1133,1160]" captionStart="Fig" captionStartId="6.[189,232,1782,1808]" captionTargetBox="[326,1260,292,1710]" captionTargetId="figure@6.[292,1296,265,1741]" captionTargetPageId="6" captionText="Fig. 1. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Carapace, dorsal view. B. Cephalothorax, ventral view, anterior part, with unfolded antennules, the arrow shows antennule with folded second article. Abbreviations: a1 = antennule; a2 = antenna; e = epistome; p.k. = postorbital knob; r = rostrum; u.a. = antennal urinary article; v.r. = ventral expansion of rostrum; 2/3 = thoracic sternal suture 2/3. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3906758" httpUri="https://zenodo.org/record/3906758/files/figure.png" pageId="5" pageNumber="6">Figs 1A</figureCitation>
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,
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<figureCitation id="1301DADE4008B22B65EC1B2FFE5FFB34" box="[436,471,1133,1160]" captionStart="Fig" captionStartId="10.[189,232,1855,1881]" captionTargetBox="[189,1398,901,1815]" captionTargetId="figure@10.[189,1398,901,1816]" captionTargetPageId="10" captionText="Fig. 5. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Type locality, stream where several specimens of the species were collected. B–D. Views of the holotype in vivo: dorsal (B), ventral (C) and frontal (D)." figureDoi="http://doi.org/10.5281/zenodo.3906766" httpUri="https://zenodo.org/record/3906766/files/figure.png" pageId="5" pageNumber="6">5B</figureCitation>
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) approximately circular to oval, slightly broader than long; dorsal carapace surface slightly concave, not strongly outlined by grooves, only with well defined gastrocardiac and thoracic grooves, approaching but not reaching antero- and posterolateral margins, respectively; no row of setae along lateral margins, except for some regularly mid-spaced setae in males; carapace angles not well marked. Anterolateral margin entire, without crenulations, lobes or teeth. Branchiostegite only weakly visible dorsally.
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</paragraph>
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<paragraph id="8B85C65B4008B22B64E51A1DFBC2F90A" blockId="5.[189,1399,1375,1718]" pageId="5" pageNumber="6">
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Eyes visible dorsally. Antennules (
|
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<figureCitation id="1301DADE4008B22B66041A1DFD30FAC5" box="[604,696,1375,1401]" captionStart="Fig" captionStartId="6.[189,232,1782,1808]" captionTargetBox="[326,1260,292,1710]" captionTargetId="figure@6.[292,1296,265,1741]" captionTargetPageId="6" captionText="Fig. 1. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Carapace, dorsal view. B. Cephalothorax, ventral view, anterior part, with unfolded antennules, the arrow shows antennule with folded second article. Abbreviations: a1 = antennule; a2 = antenna; e = epistome; p.k. = postorbital knob; r = rostrum; u.a. = antennal urinary article; v.r. = ventral expansion of rostrum; 2/3 = thoracic sternal suture 2/3. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3906758" httpUri="https://zenodo.org/record/3906758/files/figure.png" pageId="5" pageNumber="6">Fig. 1B</figureCitation>
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) obliquely folded along hollowed ventral parts of rostrum and entirely hidden dorsally. Antennae well separated from antennules, at least at their bases; urinary article at level of epistome. Rostrum (
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<figureCitation id="1301DADE4008B22B662C1AE7FD36FA03" box="[628,702,1445,1471]" captionStart="Fig" captionStartId="6.[189,232,1782,1808]" captionTargetBox="[326,1260,292,1710]" captionTargetId="figure@6.[292,1296,265,1741]" captionTargetPageId="6" captionText="Fig. 1. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Carapace, dorsal view. B. Cephalothorax, ventral view, anterior part, with unfolded antennules, the arrow shows antennule with folded second article. Abbreviations: a1 = antennule; a2 = antenna; e = epistome; p.k. = postorbital knob; r = rostrum; u.a. = antennal urinary article; v.r. = ventral expansion of rostrum; 2/3 = thoracic sternal suture 2/3. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3906758" httpUri="https://zenodo.org/record/3906758/files/figure.png" pageId="5" pageNumber="6">Figs 1</figureCitation>
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,
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<figureCitation id="1301DADE4008B22B66941AE7FD55FA03" box="[716,733,1445,1471]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[208,1380,266,1860]" captionTargetId="figure@9.[208,1380,266,1862]" captionTargetPageId="9" captionText="Fig. 4. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Dorsal view. B. Ventral view. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3906764" httpUri="https://zenodo.org/record/3906764/files/figure.png" pageId="5" pageNumber="6">4</figureCitation>
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,
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<figureCitation id="1301DADE4008B22B66B11AE7FD7EFA7C" box="[745,758,1445,1472]" captionStart="Fig" captionStartId="10.[189,232,1855,1881]" captionTargetBox="[189,1398,901,1815]" captionTargetId="figure@10.[189,1398,901,1816]" captionTargetPageId="10" captionText="Fig. 5. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Type locality, stream where several specimens of the species were collected. B–D. Views of the holotype in vivo: dorsal (B), ventral (C) and frontal (D)." figureDoi="http://doi.org/10.5281/zenodo.3906766" httpUri="https://zenodo.org/record/3906766/files/figure.png" pageId="5" pageNumber="6">5</figureCitation>
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B–D) broadly rounded, spade-shaped, barely deflexed; dorsal surface spatulate with marked depression; carapace rim as a small ridge continuous across behind rostrum. Proepistome represented by triangular ventral expansion of rostrum (
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<figureCitation id="1301DADE4008B22B60D11AAEFB62F9BA" box="[1161,1258,1516,1542]" captionStart="Fig" captionStartId="6.[189,232,1782,1808]" captionTargetBox="[326,1260,292,1710]" captionTargetId="figure@6.[292,1296,265,1741]" captionTargetPageId="6" captionText="Fig. 1. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Carapace, dorsal view. B. Cephalothorax, ventral view, anterior part, with unfolded antennules, the arrow shows antennule with folded second article. Abbreviations: a1 = antennule; a2 = antenna; e = epistome; p.k. = postorbital knob; r = rostrum; u.a. = antennal urinary article; v.r. = ventral expansion of rostrum; 2/3 = thoracic sternal suture 2/3. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3906758" httpUri="https://zenodo.org/record/3906758/files/figure.png" pageId="5" pageNumber="6">Fig. 1B</figureCitation>
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). Epistome moderately developed; anterior margin undulated. Lower orbital margin with one conspicuous knob, not visible dorsally. Pterygostomial regions with setae, distinctly separated from subhepatic area by marked ridge. Mxp3 moderately gaping, broad; merus and ischium broad, short; midlength of merus slightly longer than that of ischium (
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<figureCitation id="1301DADE4008B22B6654193AFDE3F92F" box="[524,619,1656,1683]" captionStart="Fig" captionStartId="6.[189,232,1782,1808]" captionTargetBox="[326,1260,292,1710]" captionTargetId="figure@6.[292,1296,265,1741]" captionTargetPageId="6" captionText="Fig. 1. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Carapace, dorsal view. B. Cephalothorax, ventral view, anterior part, with unfolded antennules, the arrow shows antennule with folded second article. Abbreviations: a1 = antennule; a2 = antenna; e = epistome; p.k. = postorbital knob; r = rostrum; u.a. = antennal urinary article; v.r. = ventral expansion of rostrum; 2/3 = thoracic sternal suture 2/3. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3906758" httpUri="https://zenodo.org/record/3906758/files/figure.png" pageId="5" pageNumber="6">Figs 1B</figureCitation>
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,
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<figureCitation id="1301DADE4008B22B6621193BFD16F92F" box="[633,670,1657,1683]" captionStart="Fig" captionStartId="7.[189,232,1846,1872]" captionTargetBox="[260,1308,282,1788]" captionTargetId="figure@7.[248,1339,265,1807]" captionTargetPageId="7" captionText="Fig. 2. Richerius marqueti gen. et sp. nov., holotype, ♂, 4.9 × 5.0 mm (MNHN-IU-2014-21500), same collection data as Fig. 1. A. Thoracic sternum. B. Mxp3. C. Sternopleonal cavity with G1s. D. G1, two views. Abbreviations: G1 = gonopod 1; G2 = gonopod 2; i = intercalated platelets; p = pleotelson; p.b. = press-button; s4 = expansion of sternite 4; 1–8 = thoracic sternites 1–8; 2/3–7/8 = thoracic sternal sutures 2/3–7/8. Scale bars: A = 2 mm; B–C = 1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3906760" httpUri="https://zenodo.org/record/3906760/files/figure.png" pageId="5" pageNumber="6">2B</figureCitation>
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). Sternum/pterygostome junction substantially developed due to extension of sternite 4. Milne-Edwards openings separated from chelipeds.
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</paragraph>
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<paragraph id="8B85C65B4008B22B64E5199FFC02F876" blockId="5.[189,1399,1757,1995]" pageId="5" pageNumber="6">
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Male chelipeds much stouter than walking legs, particularly in large males (
|
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<figureCitation id="1301DADE4008B22B601D199FFB07F94B" box="[1093,1167,1757,1783]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[208,1380,266,1860]" captionTargetId="figure@9.[208,1380,266,1862]" captionTargetPageId="9" captionText="Fig. 4. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Dorsal view. B. Ventral view. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3906764" httpUri="https://zenodo.org/record/3906764/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
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); merus and carpus with stiff, regularly spaced setae; propodus very inflated, covered with long soft setae partially extending on fingers; fingers with finely denticulate cutting edges, not gaping. Female chelipeds narrow, propodus moderately inflated, devoid of long setae; fingers proportionally rather long, with with finely denticulate cutting edges completely joined. Walking legs proportionally rather long, with margins bearing stiff, regularly spaced, scattered setae; dactyli slender but not distinctly longer than respective propodi, smoothly curved, setose, without teeth, ending in pointed tip.
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</paragraph>
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<caption id="DF4596D3400BB22864E519B4FC0AF858" ID-DOI="http://doi.org/10.5281/zenodo.3906758" ID-Zenodo-Dep="3906758" httpUri="https://zenodo.org/record/3906758/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1782,1808]" targetBox="[326,1260,292,1710]" targetPageId="6">
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<paragraph id="8B85C65B400BB22864E519B4FC0AF858" blockId="6.[189,1399,1780,2020]" pageId="6" pageNumber="7">
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<emphasis id="B94E1A49400BB22864E519B4FE83F8AC" bold="true" box="[189,267,1782,1808]" pageId="6" pageNumber="7">Fig. 1.</emphasis>
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<taxonomicName id="4C3ABDD8400BB228654A19B4FE7BF8AC" ID-CoL="84H9D" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[274,499,1782,1808]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A49400BB228654A19B4FE7BF8AC" box="[274,499,1782,1808]" italics="true" pageId="6" pageNumber="7">Richerius marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA732400BB22865A219B4FD25F8AC" box="[506,685,1782,1808]" pageId="6" pageNumber="7" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
, holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500).
|
||||
<emphasis id="B94E1A49400BB22864E5181DFF5AF8C5" bold="true" box="[189,210,1887,1913]" pageId="6" pageNumber="7">A</emphasis>
|
||||
. Carapace, dorsal view.
|
||||
<emphasis id="B94E1A49400BB22865AE1822FD82F8C6" bold="true" box="[502,522,1888,1914]" pageId="6" pageNumber="7">B</emphasis>
|
||||
. Cephalothorax, ventral view, anterior part, with unfolded antennules, the arrow shows antennule with folded second article. Abbreviations: a1 = antennule; a2 = antenna; e = epistome; p.k. = postorbital knob; r = rostrum; u.a. = antennal urinary article; v.r. = ventral expansion of rostrum; 2/3 = thoracic sternal suture 2/3. Scale bar = 2 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF4596D3400AB22964E51874FCE7F862" ID-DOI="http://doi.org/10.5281/zenodo.3906760" ID-Zenodo-Dep="3906760" httpUri="https://zenodo.org/record/3906760/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1846,1872]" targetBox="[260,1308,282,1788]" targetPageId="7">
|
||||
<paragraph id="8B85C65B400AB22964E51874FCE7F862" blockId="7.[189,1399,1846,2014]" pageId="7" pageNumber="8">
|
||||
<emphasis id="B94E1A49400AB22964E51874FE83F8EC" bold="true" box="[189,267,1846,1872]" pageId="7" pageNumber="8">Fig. 2.</emphasis>
|
||||
<taxonomicName id="4C3ABDD8400AB229654B1874FE7CF8EC" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[275,500,1846,1872]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A49400AB229654B1874FE7CF8EC" box="[275,500,1846,1872]" italics="true" pageId="7" pageNumber="8">Richerius marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA732400AB22965A41875FD38F8ED" box="[508,688,1847,1873]" pageId="7" pageNumber="8" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
, holotype, ♂, 4.9 × 5.0 mm (MNHN-IU-2014-21500), same collection data as Fig. 1.
|
||||
<emphasis id="B94E1A49400AB22965BC181BFE71F8CF" bold="true" box="[484,505,1881,1907]" pageId="7" pageNumber="8">A</emphasis>
|
||||
. Thoracic sternum.
|
||||
<emphasis id="B94E1A49400AB22966BD1818FD71F8C8" bold="true" box="[741,761,1882,1908]" pageId="7" pageNumber="8">B</emphasis>
|
||||
. Mxp3.
|
||||
<emphasis id="B94E1A49400AB2296704181BFCF9F8CF" bold="true" box="[860,881,1881,1907]" pageId="7" pageNumber="8">C</emphasis>
|
||||
. Sternopleonal cavity with G1s.
|
||||
<emphasis id="B94E1A49400AB22960AC1818FA81F8C8" bold="true" box="[1268,1289,1882,1908]" pageId="7" pageNumber="8">D</emphasis>
|
||||
. G1, two views. Abbreviations: G1 = gonopod 1; G2 = gonopod 2; i = intercalated platelets; p = pleotelson; p.b. = press-button; s4 = expansion of sternite 4; 1–8 = thoracic sternites 1–8; 2/3–7/8 = thoracic sternal sutures 2/3–7/8. Scale bars: A = 2 mm; B–C = 1 mm; D = 0.5 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF4596D34005B22664E51853FD60F860" ID-DOI="http://doi.org/10.5281/zenodo.3906762" ID-Zenodo-Dep="3906762" httpUri="https://zenodo.org/record/3906762/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,1809,1835]" targetBox="[189,1344,300,1745]" targetPageId="8">
|
||||
<paragraph id="8B85C65B4005B22664E51853FD60F860" blockId="8.[189,1399,1809,2012]" pageId="8" pageNumber="9">
|
||||
<emphasis id="B94E1A494005B22664E51853FE85F897" bold="true" box="[189,269,1809,1835]" pageId="8" pageNumber="9">Fig. 3.</emphasis>
|
||||
<taxonomicName id="4C3ABDD84005B226654F1853FE73F897" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[279,507,1809,1835]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494005B226654F1853FE73F897" box="[279,507,1809,1835]" italics="true" pageId="8" pageNumber="9">Richerius marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324005B226665D1850FD49F890" box="[517,705,1810,1836]" pageId="8" pageNumber="9" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
, paratype, ovigerous ♀, New Caledonia: North Province, 21º21.890ʹ S, 165º32.683ʹ E, Creek stream, tributary of the Böua at level of the Néaoua dam, Houaïlou township, st. HYNC 1823, 476 m a.s.l., “Our Planet Reviewed”, Hydrobio Expedition, Nicolas Charpin leg., 9 Oct. 2017, 7.0 × 7.1 mm (MNHN-IU-2014-21501).
|
||||
<emphasis id="B94E1A494005B22667D01839FC15F829" bold="true" box="[904,925,1915,1941]" pageId="8" pageNumber="9">A</emphasis>
|
||||
.Thoracic sternum, with lowered pleon.
|
||||
<emphasis id="B94E1A494005B22664E518DCFF59F804" bold="true" box="[189,209,1950,1976]" pageId="8" pageNumber="9">B</emphasis>
|
||||
. Pleon. Abbreviations: b.a. = branchiosternal aperture; e = egg; s4 = expansion of sternite 4; v = vulva; 4–8 = thoracic sternites 4–8. Scale bar = 3 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF4596D34004B22764E51826FCAAF855" ID-DOI="http://doi.org/10.5281/zenodo.3906764" ID-Zenodo-Dep="3906764" httpUri="https://zenodo.org/record/3906764/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,232,1892,1918]" targetBox="[208,1380,266,1860]" targetPageId="9">
|
||||
<paragraph id="8B85C65B4004B22764E51826FCAAF855" blockId="9.[189,1399,1891,2025]" pageId="9" pageNumber="10">
|
||||
<emphasis id="B94E1A494004B22764E51826FE82F8C2" bold="true" box="[189,266,1892,1918]" pageId="9" pageNumber="10">Fig. 4.</emphasis>
|
||||
<taxonomicName id="4C3ABDD84004B227654A1827FE7BF8C2" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[274,499,1893,1919]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494004B227654A1827FE7BF8C2" box="[274,499,1893,1919]" italics="true" pageId="9" pageNumber="10">Richerius marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324004B22765A21827FD25F8C3" box="[506,685,1893,1919]" pageId="9" pageNumber="10" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
, holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500).
|
||||
<emphasis id="B94E1A494004B22764E5188CFF5AF854" bold="true" box="[189,210,1998,2024]" pageId="9" pageNumber="10">A</emphasis>
|
||||
. Dorsal view.
|
||||
<emphasis id="B94E1A494004B2276524188CFE18F854" bold="true" box="[380,400,1998,2024]" pageId="9" pageNumber="10">B</emphasis>
|
||||
. Ventral view. Scale bars = 2 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B85C65B4007B22464E51E49FAC1FE0E" blockId="10.[189,1399,267,435]" pageId="10" pageNumber="11">
|
||||
Thoracic sternum of male (
|
||||
<figureCitation id="1301DADE4007B224665F1E49FDEEFE99" box="[519,614,267,293]" captionStart="Fig" captionStartId="7.[189,232,1846,1872]" captionTargetBox="[260,1308,282,1788]" captionTargetId="figure@7.[248,1339,265,1807]" captionTargetPageId="7" captionText="Fig. 2. Richerius marqueti gen. et sp. nov., holotype, ♂, 4.9 × 5.0 mm (MNHN-IU-2014-21500), same collection data as Fig. 1. A. Thoracic sternum. B. Mxp3. C. Sternopleonal cavity with G1s. D. G1, two views. Abbreviations: G1 = gonopod 1; G2 = gonopod 2; i = intercalated platelets; p = pleotelson; p.b. = press-button; s4 = expansion of sternite 4; 1–8 = thoracic sternites 1–8; 2/3–7/8 = thoracic sternal sutures 2/3–7/8. Scale bars: A = 2 mm; B–C = 1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3906760" httpUri="https://zenodo.org/record/3906760/files/figure.png" pageId="10" pageNumber="11">Fig. 2A</figureCitation>
|
||||
) with sternites 4–8 considerably enlarged, with suture 4/5 to 7/8 laterally confined. Sternites 1 and 2 not separated by suture but their demarcation visible on lateral margins; sternite 2 separated from sternite 3 by complete suture; sternite 3 as narrow pentagonal band; suture 3/4 tiny, only lateral, just at level of extension of sternite 4. No longitudinal line. Paired branchiosternal canal apertures located posterolaterally on sternite 8, concealed by pleon (
|
||||
<figureCitation id="1301DADE4007B22460851EDAFAB3FE0E" box="[1245,1339,408,434]" captionStart="Fig" captionStartId="8.[189,232,1809,1835]" captionTargetBox="[189,1344,300,1745]" captionTargetId="figure@8.[189,1398,265,1759]" captionTargetPageId="8" captionText="Fig. 3. Richerius marqueti gen. et sp. nov., paratype, ovigerous ♀, New Caledonia: North Province, 21º21.890ʹ S, 165º32.683ʹ E, Creek stream, tributary of the Böua at level of the Néaoua dam, Houaïlou township, st. HYNC 1823, 476 m a.s.l., “Our Planet Reviewed”, Hydrobio Expedition, Nicolas Charpin leg., 9 Oct. 2017, 7.0 × 7.1 mm (MNHN-IU-2014-21501). A.Thoracic sternum, with lowered pleon. B. Pleon. Abbreviations: b.a. = branchiosternal aperture; e = egg; s4 = expansion of sternite 4; v = vulva; 4–8 = thoracic sternites 4–8. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3906762" httpUri="https://zenodo.org/record/3906762/files/figure.png" pageId="10" pageNumber="11">Fig. 3A</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4007B22464E51E9BFBD1FD3C" blockId="10.[189,1399,473,641]" pageId="10" pageNumber="11">
|
||||
Sternopleonal cavity of male short, triangular, deep (
|
||||
<figureCitation id="1301DADE4007B22467431E9BFCFEFE48" box="[795,886,473,500]" captionStart="Fig" captionStartId="7.[189,232,1846,1872]" captionTargetBox="[260,1308,282,1788]" captionTargetId="figure@7.[248,1339,265,1807]" captionTargetPageId="7" captionText="Fig. 2. Richerius marqueti gen. et sp. nov., holotype, ♂, 4.9 × 5.0 mm (MNHN-IU-2014-21500), same collection data as Fig. 1. A. Thoracic sternum. B. Mxp3. C. Sternopleonal cavity with G1s. D. G1, two views. Abbreviations: G1 = gonopod 1; G2 = gonopod 2; i = intercalated platelets; p = pleotelson; p.b. = press-button; s4 = expansion of sternite 4; 1–8 = thoracic sternites 1–8; 2/3–7/8 = thoracic sternal sutures 2/3–7/8. Scale bars: A = 2 mm; B–C = 1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3906760" httpUri="https://zenodo.org/record/3906760/files/figure.png" pageId="10" pageNumber="11">Fig. 2C</figureCitation>
|
||||
). Pleons in both sexes (
|
||||
<figureCitation id="1301DADE4007B22460D91E9BFB68FE48" box="[1153,1248,473,500]" captionStart="Fig" captionStartId="7.[189,232,1846,1872]" captionTargetBox="[260,1308,282,1788]" captionTargetId="figure@7.[248,1339,265,1807]" captionTargetPageId="7" captionText="Fig. 2. Richerius marqueti gen. et sp. nov., holotype, ♂, 4.9 × 5.0 mm (MNHN-IU-2014-21500), same collection data as Fig. 1. A. Thoracic sternum. B. Mxp3. C. Sternopleonal cavity with G1s. D. G1, two views. Abbreviations: G1 = gonopod 1; G2 = gonopod 2; i = intercalated platelets; p = pleotelson; p.b. = press-button; s4 = expansion of sternite 4; 1–8 = thoracic sternites 1–8; 2/3–7/8 = thoracic sternal sutures 2/3–7/8. Scale bars: A = 2 mm; B–C = 1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3906760" httpUri="https://zenodo.org/record/3906760/files/figure.png" pageId="10" pageNumber="11">Figs 2A</figureCitation>
|
||||
,
|
||||
<figureCitation id="1301DADE4007B22460B31E98FA86FE48" box="[1259,1294,474,500]" captionStart="Fig" captionStartId="8.[189,232,1809,1835]" captionTargetBox="[189,1344,300,1745]" captionTargetId="figure@8.[189,1398,265,1759]" captionTargetPageId="8" captionText="Fig. 3. Richerius marqueti gen. et sp. nov., paratype, ovigerous ♀, New Caledonia: North Province, 21º21.890ʹ S, 165º32.683ʹ E, Creek stream, tributary of the Böua at level of the Néaoua dam, Houaïlou township, st. HYNC 1823, 476 m a.s.l., “Our Planet Reviewed”, Hydrobio Expedition, Nicolas Charpin leg., 9 Oct. 2017, 7.0 × 7.1 mm (MNHN-IU-2014-21501). A.Thoracic sternum, with lowered pleon. B. Pleon. Abbreviations: b.a. = branchiosternal aperture; e = egg; s4 = expansion of sternite 4; v = vulva; 4–8 = thoracic sternites 4–8. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3906762" httpUri="https://zenodo.org/record/3906762/files/figure.png" pageId="10" pageNumber="11">3B</figureCitation>
|
||||
) without fused somites, except 6 fused to telson, thus five somites plus pleotelson. Male pleon widely triangular, regularly widening from base to pleotelson; pleotelson widely triangular, somewhat trilobed, with intercalated plates partially delineated, salient and visible laterally at base, partially included. Pressbutton on lateral side of sternopleonal cavity (
|
||||
<figureCitation id="1301DADE4007B22466801D24FCBDFD3C" box="[728,821,614,640]" captionStart="Fig" captionStartId="7.[189,232,1846,1872]" captionTargetBox="[260,1308,282,1788]" captionTargetId="figure@7.[248,1339,265,1807]" captionTargetPageId="7" captionText="Fig. 2. Richerius marqueti gen. et sp. nov., holotype, ♂, 4.9 × 5.0 mm (MNHN-IU-2014-21500), same collection data as Fig. 1. A. Thoracic sternum. B. Mxp3. C. Sternopleonal cavity with G1s. D. G1, two views. Abbreviations: G1 = gonopod 1; G2 = gonopod 2; i = intercalated platelets; p = pleotelson; p.b. = press-button; s4 = expansion of sternite 4; 1–8 = thoracic sternites 1–8; 2/3–7/8 = thoracic sternal sutures 2/3–7/8. Scale bars: A = 2 mm; B–C = 1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3906760" httpUri="https://zenodo.org/record/3906760/files/figure.png" pageId="10" pageNumber="11">Fig. 2C</figureCitation>
|
||||
). Male gonopore sternal.
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4007B22564E51DEAFD8AFED0" blockId="10.[189,1399,680,777]" lastBlockId="11.[189,1399,267,364]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
|
||||
Female pleon (
|
||||
<figureCitation id="1301DADE4007B224652C1DEAFE33FD7E" box="[372,443,680,706]" captionStart="Fig" captionStartId="8.[189,232,1809,1835]" captionTargetBox="[189,1344,300,1745]" captionTargetId="figure@8.[189,1398,265,1759]" captionTargetPageId="8" captionText="Fig. 3. Richerius marqueti gen. et sp. nov., paratype, ovigerous ♀, New Caledonia: North Province, 21º21.890ʹ S, 165º32.683ʹ E, Creek stream, tributary of the Böua at level of the Néaoua dam, Houaïlou township, st. HYNC 1823, 476 m a.s.l., “Our Planet Reviewed”, Hydrobio Expedition, Nicolas Charpin leg., 9 Oct. 2017, 7.0 × 7.1 mm (MNHN-IU-2014-21501). A.Thoracic sternum, with lowered pleon. B. Pleon. Abbreviations: b.a. = branchiosternal aperture; e = egg; s4 = expansion of sternite 4; v = vulva; 4–8 = thoracic sternites 4–8. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3906762" httpUri="https://zenodo.org/record/3906762/files/figure.png" pageId="10" pageNumber="11">Fig. 3</figureCitation>
|
||||
A–B) oval to expanded in ovigerous females. Female pleopods 2–5 biramous. Ovigerous females with very few and large eggs measuring
|
||||
<quantity id="4CC26BBE4007B22467C21D89FB8DFD5A" box="[922,1029,715,742]" metricMagnitude="-4" metricUnit="m" metricValue="9.6" pageId="10" pageNumber="11" unit="mm" value="0.96">0.96 mm</quantity>
|
||||
in diameter (
|
||||
<figureCitation id="1301DADE4007B22460F31D89FA84FD59" box="[1195,1292,715,741]" captionStart="Fig" captionStartId="8.[189,232,1809,1835]" captionTargetBox="[189,1344,300,1745]" captionTargetId="figure@8.[189,1398,265,1759]" captionTargetPageId="8" captionText="Fig. 3. Richerius marqueti gen. et sp. nov., paratype, ovigerous ♀, New Caledonia: North Province, 21º21.890ʹ S, 165º32.683ʹ E, Creek stream, tributary of the Böua at level of the Néaoua dam, Houaïlou township, st. HYNC 1823, 476 m a.s.l., “Our Planet Reviewed”, Hydrobio Expedition, Nicolas Charpin leg., 9 Oct. 2017, 7.0 × 7.1 mm (MNHN-IU-2014-21501). A.Thoracic sternum, with lowered pleon. B. Pleon. Abbreviations: b.a. = branchiosternal aperture; e = egg; s4 = expansion of sternite 4; v = vulva; 4–8 = thoracic sternites 4–8. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3906762" httpUri="https://zenodo.org/record/3906762/files/figure.png" pageId="10" pageNumber="11">Fig. 3A</figureCitation>
|
||||
). Vulvae located on undivided portion of thoracic sternum corresponding to sternite 6, thus not deplaced anteriorly. First gonopods stout, practically filling most of sternopleonal cavity (
|
||||
<figureCitation id="1301DADE4006B225604B1E49FBFDFE99" box="[1043,1141,267,293]" captionStart="Fig" captionStartId="7.[189,232,1846,1872]" captionTargetBox="[260,1308,282,1788]" captionTargetId="figure@7.[248,1339,265,1807]" captionTargetPageId="7" captionText="Fig. 2. Richerius marqueti gen. et sp. nov., holotype, ♂, 4.9 × 5.0 mm (MNHN-IU-2014-21500), same collection data as Fig. 1. A. Thoracic sternum. B. Mxp3. C. Sternopleonal cavity with G1s. D. G1, two views. Abbreviations: G1 = gonopod 1; G2 = gonopod 2; i = intercalated platelets; p = pleotelson; p.b. = press-button; s4 = expansion of sternite 4; 1–8 = thoracic sternites 1–8; 2/3–7/8 = thoracic sternal sutures 2/3–7/8. Scale bars: A = 2 mm; B–C = 1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3906760" httpUri="https://zenodo.org/record/3906760/files/figure.png" pageId="11" pageNumber="12">Fig. 2C</figureCitation>
|
||||
). G1 curved at base, otherwise with little curvature, with terminal portion bearing fine setae, terminating in one lobe, without corneous process (
|
||||
<figureCitation id="1301DADE4006B22565CE1E10FE7DFED0" box="[406,501,338,364]" captionStart="Fig" captionStartId="7.[189,232,1846,1872]" captionTargetBox="[260,1308,282,1788]" captionTargetId="figure@7.[248,1339,265,1807]" captionTargetPageId="7" captionText="Fig. 2. Richerius marqueti gen. et sp. nov., holotype, ♂, 4.9 × 5.0 mm (MNHN-IU-2014-21500), same collection data as Fig. 1. A. Thoracic sternum. B. Mxp3. C. Sternopleonal cavity with G1s. D. G1, two views. Abbreviations: G1 = gonopod 1; G2 = gonopod 2; i = intercalated platelets; p = pleotelson; p.b. = press-button; s4 = expansion of sternite 4; 1–8 = thoracic sternites 1–8; 2/3–7/8 = thoracic sternal sutures 2/3–7/8. Scale bars: A = 2 mm; B–C = 1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3906760" httpUri="https://zenodo.org/record/3906760/files/figure.png" pageId="11" pageNumber="12">Fig. 2D</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="DF4596D34007B22464E5187DFCD8F85A" ID-DOI="http://doi.org/10.5281/zenodo.3906766" ID-Zenodo-Dep="3906766" httpUri="https://zenodo.org/record/3906766/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,232,1855,1881]" targetBox="[189,1398,901,1815]" targetPageId="10">
|
||||
<paragraph id="8B85C65B4007B22464E5187DFCD8F85A" blockId="10.[189,1399,1853,2022]" pageId="10" pageNumber="11">
|
||||
<emphasis id="B94E1A494007B22464E5187DFE82F8E5" bold="true" box="[189,266,1855,1881]" pageId="10" pageNumber="11">Fig. 5.</emphasis>
|
||||
<taxonomicName id="4C3ABDD84007B224654A187DFE7BF8E5" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[274,499,1855,1881]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494007B224654A187DFE7BF8E5" box="[274,499,1855,1881]" italics="true" pageId="10" pageNumber="11">Richerius marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324007B22465A2187DFD25F8E5" box="[506,685,1855,1881]" pageId="10" pageNumber="11" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
, holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500).
|
||||
<emphasis id="B94E1A494007B22464E518EAFF5AF87E" bold="true" box="[189,210,1960,1986]" pageId="10" pageNumber="11">A</emphasis>
|
||||
. Type locality, stream where several specimens of the species were collected.
|
||||
<emphasis id="B94E1A494007B22460CA18EBFB41F87F" bold="true" box="[1170,1225,1961,1987]" pageId="10" pageNumber="11">B–D</emphasis>
|
||||
. Views of the holotype in vivo: dorsal (B), ventral (C) and frontal (D).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B85C65B4006B22564E51ED1FE9DFE11" blockId="11.[189,277,403,429]" box="[189,277,403,429]" pageId="11" pageNumber="12">
|
||||
<heading id="D0CD71374006B22564E51ED1FE9DFE11" bold="true" box="[189,277,403,429]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
|
||||
<emphasis id="B94E1A494006B22564E51ED1FE9DFE11" bold="true" box="[189,277,403,429]" pageId="11" pageNumber="12">Colour</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4006B22564E51E80FBCAFE60" blockId="11.[189,1090,449,476]" box="[189,1090,449,476]" pageId="11" pageNumber="12">
|
||||
In life (
|
||||
<figureCitation id="1301DADE4006B225654C1E80FED1FE60" box="[276,345,449,476]" captionStart="Fig" captionStartId="10.[189,232,1855,1881]" captionTargetBox="[189,1398,901,1815]" captionTargetId="figure@10.[189,1398,901,1816]" captionTargetPageId="10" captionText="Fig. 5. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Type locality, stream where several specimens of the species were collected. B–D. Views of the holotype in vivo: dorsal (B), ventral (C) and frontal (D)." figureDoi="http://doi.org/10.5281/zenodo.3906766" httpUri="https://zenodo.org/record/3906766/files/figure.png" pageId="11" pageNumber="12">Fig. 5</figureCitation>
|
||||
B–D), the colour is overall brown, mottled with yellowish dots.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C32095D04006B22564E51D41FEDAFCFE" pageId="11" pageNumber="12" type="distribution">
|
||||
<paragraph id="8B85C65B4006B22564E51D41FEDEFDA1" blockId="11.[189,342,515,541]" box="[189,342,515,541]" pageId="11" pageNumber="12">
|
||||
<heading id="D0CD71374006B22564E51D41FEDEFDA1" bold="true" box="[189,342,515,541]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
|
||||
<emphasis id="B94E1A494006B22564E51D41FEDEFDA1" bold="true" box="[189,342,515,541]" pageId="11" pageNumber="12">Distribution</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4006B22564E51D70FEDAFCFE" blockId="11.[189,1399,562,834]" pageId="11" pageNumber="12">
|
||||
<taxonomicName id="4C3ABDD84006B22564E51D70FE29FDF0" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[189,417,562,588]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494006B22564E51D70FE29FDF0" box="[189,417,562,588]" italics="true" pageId="11" pageNumber="12">Richerius marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324006B22565F31D70FDEDFDF0" box="[427,613,562,588]" pageId="11" pageNumber="12" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
is exclusively known from freshwaters in small flowing streams where it lives among the aquatic vegetation (
|
||||
<figureCitation id="1301DADE4006B225668A1D17FCB9FDCC" box="[722,817,597,624]" captionStart="Fig" captionStartId="10.[189,232,1855,1881]" captionTargetBox="[189,1398,901,1815]" captionTargetId="figure@10.[189,1398,901,1816]" captionTargetPageId="10" captionText="Fig. 5. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Type locality, stream where several specimens of the species were collected. B–D. Views of the holotype in vivo: dorsal (B), ventral (C) and frontal (D)." figureDoi="http://doi.org/10.5281/zenodo.3906766" httpUri="https://zenodo.org/record/3906766/files/figure.png" pageId="11" pageNumber="12">Fig. 5A</figureCitation>
|
||||
). It is found at an altitude of up
|
||||
<quantity id="4CC26BBE4006B22560EA1D17FB77FDD3" box="[1202,1279,597,624]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="12" unit="m" value="500.0">500 m</quantity>
|
||||
, therefore further from the sea than
|
||||
<taxonomicName id="4C3ABDD84006B22565B21D3AFD4AFD2E" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[490,706,632,658]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494006B22565B21D3AFD4AFD2E" box="[490,706,632,658]" italics="true" pageId="11" pageNumber="12">Odiomaris pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
(that is usually encountered in the lower course of rivers). By comparison,
|
||||
<taxonomicName id="4C3ABDD84006B22565D91DDEFCA9FD09" authority="(Chilton, 1882)" baseAuthorityName="Chilton" baseAuthorityYear="1882" box="[385,801,667,694]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="lacustris">
|
||||
<emphasis id="B94E1A494006B22565D91DDEFDEAFD09" box="[385,610,667,694]" italics="true" pageId="11" pageNumber="12">Amarinus lacustris</emphasis>
|
||||
(Chilton, 1882)
|
||||
</taxonomicName>
|
||||
has been reported from
|
||||
<collectingCountry id="F32D86CB4006B225601F1DDEFB6EFD09" box="[1095,1254,667,694]" name="New Zealand" pageId="11" pageNumber="12">New Zealand</collectingCountry>
|
||||
freshwaters from Lord Howe Island at altitudes between
|
||||
<quantity id="4CC26BBE4006B22566B91DFCFCC4FD65" box="[737,844,702,729]" metricMagnitude="1" metricUnit="m" metricValue="7.65" metricValueMax="9.2" metricValueMin="6.1" pageId="11" pageNumber="12" unit="m" value="76.5" valueMax="92.0" valueMin="61.0">61–92 m</quantity>
|
||||
(
|
||||
<bibRefCitation id="EFABBBAA4006B22567061DFCFB91FD65" author="Etheridge R." box="[862,1049,702,729]" pageId="11" pageNumber="12" pagination="1 - 4" refId="ref12979" refString="Etheridge R. 1889. The general zoology of Lord Howe Island; containing also an account of the collections made by the Australian Museum Collecting Party, Aug. - Sept., 1887. Australian Museum Memoir 2 (1): 1 - 4. https: // doi. org / 10.3853 / j. 0067 - 1967.2.1889.479" type="journal article" year="1889">
|
||||
Etheridge
|
||||
<date id="FF84E09B4006B22567851DFDFB91FD65" box="[989,1049,703,729]" pageId="11" pageNumber="12" value="1889">1889</date>
|
||||
</bibRefCitation>
|
||||
) and even about
|
||||
<quantity id="4CC26BBE4006B22560B61DFDFAFFFD65" box="[1262,1399,703,729]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="11" pageNumber="12" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">800–900 m</quantity>
|
||||
(
|
||||
<bibRefCitation id="EFABBBAA4006B225649D1DA0FEEDFD40" author="Chilton C." box="[197,357,737,764]" pageId="11" pageNumber="12" pagination="316 - 320" refId="ref12420" refString="Chilton C. 1915. A freshwater crab and its distribution in Australia and New Zealand. Transactions and Proceedings of the New Zealand Institute 47: 316 - 320. Available from http: // rsnz. natlib. govt. nz / volume / rsnz _ 47 / rsnz _ 47 _ 00 _ 005860. html [accessed 27 May 2020]." type="journal article" year="1915">
|
||||
Chilton
|
||||
<date id="FF84E09B4006B225657F1DA3FEEDFD40" box="[295,357,737,764]" pageId="11" pageNumber="12" value="1915">1915</date>
|
||||
</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFABBBAA4006B22565291DA0FD95FD41" author="Holthuis L. B." box="[369,541,738,765]" pageId="11" pageNumber="12" pagination="109 - 121" refId="ref13478" refString="Holthuis L. B. 1968. On Hymenosomatidae (Crustacea Decapoda Brachyura) from fresh water, with the description of a new species. Beaufortia 15 (195): 109 - 121. Available from https: // www. repository. naturalis. nl / record / 504776 [accessed 27 May 2020]." type="journal article" year="1968">
|
||||
Holthuis
|
||||
<date id="FF84E09B4006B22565861DA0FD95FD41" box="[478,541,738,765]" pageId="11" pageNumber="12" value="1968">1968</date>
|
||||
</bibRefCitation>
|
||||
);
|
||||
<taxonomicName id="4C3ABDD84006B225666B1DA0FCF5FD40" authority="Holthuis, 1968" authorityName="Holthuis" authorityYear="1968" box="[563,893,738,764]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="angelicus">
|
||||
<emphasis id="B94E1A494006B225666B1DA0FD4CFD40" box="[563,708,738,764]" italics="true" pageId="11" pageNumber="12">A. angelicus</emphasis>
|
||||
<bibRefCitation id="EFABBBAA4006B22566941DA0FCF5FD40" author="Holthuis L. B." box="[716,893,738,764]" pageId="11" pageNumber="12" pagination="109 - 121" refId="ref13478" refString="Holthuis L. B. 1968. On Hymenosomatidae (Crustacea Decapoda Brachyura) from fresh water, with the description of a new species. Beaufortia 15 (195): 109 - 121. Available from https: // www. repository. naturalis. nl / record / 504776 [accessed 27 May 2020]." type="journal article" year="1968">Holthuis, 1968</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, from the central mountain range of
|
||||
<collectingCountry id="F32D86CB4006B22561771DA0FEDBFCA3" name="Papua New Guinea" pageId="11" pageNumber="12">Papua New Guinea</collectingCountry>
|
||||
, was collected in pure freshwater of a watercress swamp at an altitude of
|
||||
<quantity id="4CC26BBE4006B22560F11C47FA89FCA3" box="[1193,1281,773,799]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="12" unit="m" value="1600.0">1600 m</quantity>
|
||||
(
|
||||
<bibRefCitation id="EFABBBAA4006B22561481C47FF72FCFE" author="Holthuis L. B." pageId="11" pageNumber="12" pagination="109 - 121" refId="ref13478" refString="Holthuis L. B. 1968. On Hymenosomatidae (Crustacea Decapoda Brachyura) from fresh water, with the description of a new species. Beaufortia 15 (195): 109 - 121. Available from https: // www. repository. naturalis. nl / record / 504776 [accessed 27 May 2020]." type="journal article" year="1968">
|
||||
Holthuis
|
||||
<date id="FF84E09B4006B22564E51C6AFF72FCFE" box="[189,250,808,834]" pageId="11" pageNumber="12" value="1968">1968</date>
|
||||
</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EFABBBAA4006B225655E1C6AFECDFCFE" author="Holthuis L. B." box="[262,325,808,834]" pageId="11" pageNumber="12" pagination="603 - 619" refId="ref13537" refString="Holthuis L. B. 1982. Freshwater Crustacea Decapoda of New Guinea. In: Gressitt J. L. (ed.) Biogeography and Ecology of New Guinea. Vol. 2, Pt. 6. Monographiae Biologicae 42: 603 - 619. https: // doi. org / 10.1007 / 978 - 94 - 009 - 8632 - 9 _ 28" type="journal article" year="1982">
|
||||
<date id="FF84E09B4006B225655E1C6AFECDFCFE" box="[262,325,808,834]" pageId="11" pageNumber="12" value="1982">1982</date>
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C32095D04006B22364E51C28FB42F8FE" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" type="discussion">
|
||||
<paragraph id="8B85C65B4006B22564E51C28FEA7FC38" blockId="11.[189,303,874,900]" box="[189,303,874,900]" pageId="11" pageNumber="12">
|
||||
<heading id="D0CD71374006B22564E51C28FEA7FC38" bold="true" box="[189,303,874,900]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
|
||||
<emphasis id="B94E1A494006B22564E51C28FEA7FC38" bold="true" box="[189,303,874,900]" pageId="11" pageNumber="12">Remarks</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4006B22564E51CDAFC25FB53" blockId="11.[189,1399,920,1263]" pageId="11" pageNumber="12">
|
||||
The subfamily
|
||||
<taxonomicName id="4C3ABDD84006B22565241CDAFD9DFC0E" authorityName="Guinot" authorityYear="2011" box="[380,533,920,946]" class="Malacostraca" family="Hymenosomatidae" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subFamily" subFamily="Odiomarinae">Odiomarinae</taxonomicName>
|
||||
was erected by
|
||||
<bibRefCitation id="EFABBBAA4006B22566B51CDAFC2EFC0E" author="Guinot D." box="[749,934,920,946]" pageId="11" pageNumber="12" pagination="20 - 32" refId="ref13311" refString="Guinot D. 2011 a. Odiomarinae nov. subfam., a new subfamily for two primitive genera of the Hymenosomatidae MacLeay, 1838 (Crustacea, Decapoda, Brachyura). Zootaxa 2732: 20 - 32. https: // doi. org / 10.11646 / zootaxa. 2732.1.2" type="journal article" year="2011">Guinot (2011a)</bibRefCitation>
|
||||
to receive two genera of the family
|
||||
<taxonomicName id="4C3ABDD84006B22564E51CF9FE11FC69" authorityName="MacLeay" authorityYear="1838" box="[189,409,955,981]" class="Malacostraca" family="Hymenosomatidae" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="family">Hymenosomatidae</taxonomicName>
|
||||
characterised by the presence on the male pleon of intercalated platelets, either articulated and moveable (
|
||||
<bibRefCitation id="EFABBBAA4006B22565A31C9CFD2BFC45" author="Guinot D." box="[507,675,990,1017]" pageId="11" pageNumber="12" pagination="20 - 32" refId="ref13311" refString="Guinot D. 2011 a. Odiomarinae nov. subfam., a new subfamily for two primitive genera of the Hymenosomatidae MacLeay, 1838 (Crustacea, Decapoda, Brachyura). Zootaxa 2732: 20 - 32. https: // doi. org / 10.11646 / zootaxa. 2732.1.2" type="journal article" year="2011">Guinot 2011a</bibRefCitation>
|
||||
: fig. 2) or relatively less well-demarcated:
|
||||
<taxonomicName id="4C3ABDD84006B22560F21C9CFE46FBA0" authority="Ng & Richer de Forges, 1996" authorityName="Ng & Richer de Forges" authorityYear="1996" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494006B22560F21C9CFAAFFC44" box="[1194,1319,990,1016]" italics="true" pageId="11" pageNumber="12">Odiomaris</emphasis>
|
||||
<bibRefCitation id="EFABBBAA4006B225616A1C9DFE46FBA0" author="Ng P. K. L. & Richer de Forges B." pageId="11" pageNumber="12" pagination="263 - 276" refId="ref14618" refString="Ng P. K. L. & Richer de Forges B. 1996. The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of New Caledonia, with descriptions of two new genera and two new species. Memoirs of the Queensland Museum 39 (2): 263 - 276. Available from https: // www. biodiversitylibrary. org / part / 50813 # / summary [accessed 27 May 2020]." type="journal article" year="1996">Ng & Richer de Forges, 1996</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, endemic to
|
||||
<collectingCountry id="F32D86CB4006B22566381B40FC90FBA0" box="[608,792,1026,1052]" name="New Caledonia" pageId="11" pageNumber="12">New Caledonia</collectingCountry>
|
||||
, and
|
||||
<emphasis id="B94E1A494006B225670C1B40FC33FBA0" box="[852,955,1026,1052]" italics="true" pageId="11" pageNumber="12">Amarinu</emphasis>
|
||||
(at least
|
||||
<emphasis id="B94E1A494006B225607A1B41FB07FBA0" box="[1058,1167,1027,1052]" italics="true" pageId="11" pageNumber="12">pro parte</emphasis>
|
||||
), mostly from fresh and estuarine waters of the Indo-West Pacific region.
|
||||
<emphasis id="B94E1A494006B225676E1B67FC13FB83" box="[822,923,1061,1087]" italics="true" pageId="11" pageNumber="12">Richeriu</emphasis>
|
||||
s gen. nov. shows several plesiomorphic characters that include the male and female pleons without fused somites (except for pleotelson), thus consisting of six elements (namely the maximum of somites existing in
|
||||
<taxonomicName id="4C3ABDD84006B22560041B29FAB5FB3A" authorityName="MacLeay" authorityYear="1838" box="[1116,1341,1131,1158]" class="Malacostraca" family="Hymenosomatidae" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="family">Hymenosomatidae</taxonomicName>
|
||||
), the prominent, partially demarcated intercalated platelets, the thoracic sternum with the anterior sternites forming a small produced plate, the vulvae not anteriorly displaced, the G1 only gently curved and simple.
|
||||
<taxonomicName id="4C3ABDD84006B22565421B97FE02FB53" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[282,394,1237,1263]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B94E1A494006B22565421B97FE02FB53" box="[282,394,1237,1263]" italics="true" pageId="11" pageNumber="12">Richerius</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324006B22565CA1B97FE75FB53" box="[402,509,1237,1263]" pageId="11" pageNumber="12" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
can be assigned to the
|
||||
<taxonomicName id="4C3ABDD84006B22567541B97FC21FB53" authorityName="Guinot" authorityYear="2011" box="[780,937,1237,1263]" class="Malacostraca" family="Hymenosomatidae" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subFamily" subFamily="Odiomarinae">Odiomarinae</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4006B22564E51A54FEBCF9F6" blockId="11.[189,1399,1302,1611]" pageId="11" pageNumber="12">
|
||||
<taxonomicName id="4C3ABDD84006B22564E51A54FEB2FA8C" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[189,314,1302,1328]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494006B22564E51A54FEB2FA8C" box="[189,314,1302,1328]" italics="true" pageId="11" pageNumber="12">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
is known from two species: the
|
||||
<typeStatus id="548178F94006B22566811A55FC84FA8D" box="[729,780,1303,1329]" pageId="11" pageNumber="12">type</typeStatus>
|
||||
species
|
||||
<taxonomicName id="4C3ABDD84006B22567211A54FAF9FA8C" authority="A. Milne-Edwards, 1873" authorityName="A. Milne-Edwards" authorityYear="1873" box="[889,1393,1302,1329]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pilosa">
|
||||
<emphasis id="B94E1A494006B22567211A54FBBEFA8C" box="[889,1078,1302,1328]" italics="true" pageId="11" pageNumber="12">Elamena pilosa</emphasis>
|
||||
A. Milne-Edwards, 1873
|
||||
</taxonomicName>
|
||||
, referred to as
|
||||
<taxonomicName id="4C3ABDD84006B22565341A78FD2CFAEF" authority="White, 1846" authorityName="White" authorityYear="1846" box="[364,676,1337,1364]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494006B22565341A78FD8CFAE8" box="[364,516,1338,1364]" italics="true" pageId="11" pageNumber="12">Halicarcinus</emphasis>
|
||||
White, 1846
|
||||
</taxonomicName>
|
||||
by
|
||||
<bibRefCitation id="EFABBBAA4006B22566801A7BFC12FAE8" author="Holthuis L. B." box="[728,922,1337,1364]" pageId="11" pageNumber="12" pagination="109 - 121" refId="ref13478" refString="Holthuis L. B. 1968. On Hymenosomatidae (Crustacea Decapoda Brachyura) from fresh water, with the description of a new species. Beaufortia 15 (195): 109 - 121. Available from https: // www. repository. naturalis. nl / record / 504776 [accessed 27 May 2020]." type="journal article" year="1968">Holthuis (1968)</bibRefCitation>
|
||||
, as
|
||||
<taxonomicName id="4C3ABDD84006B22567961A78FBC8FAE8" authorityName="Lucas" authorityYear="1980" box="[974,1088,1338,1364]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494006B22567961A78FBC8FAE8" box="[974,1088,1338,1364]" italics="true" pageId="11" pageNumber="12">Amarinus</emphasis>
|
||||
</taxonomicName>
|
||||
by
|
||||
<bibRefCitation id="EFABBBAA4006B225602D1A78FA9CFAE8" author="Lucas J. S." box="[1141,1300,1338,1364]" pageId="11" pageNumber="12" pagination="148 - 247" refId="ref14163" refString="Lucas J. S. 1980. Spider crabs of the family Hymenosomatidae (Crustacea: Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum 33 (4): 148 - 247. https: // doi. org / 10.3853 / j. 0067 - 1975.33.1980.199" type="journal article" year="1980">Lucas (1980)</bibRefCitation>
|
||||
then as
|
||||
<taxonomicName id="4C3ABDD84006B22564E51A1FFEB2FACB" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[189,314,1373,1399]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494006B22564E51A1FFEB2FACB" box="[189,314,1373,1399]" italics="true" pageId="11" pageNumber="12">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
by
|
||||
<bibRefCitation id="EFABBBAA4006B22565301A1FFD5FFAC4" author="Ng P. K. L. & Richer de Forges B." box="[360,727,1373,1400]" pageId="11" pageNumber="12" pagination="263 - 276" refId="ref14618" refString="Ng P. K. L. & Richer de Forges B. 1996. The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of New Caledonia, with descriptions of two new genera and two new species. Memoirs of the Queensland Museum 39 (2): 263 - 276. Available from https: // www. biodiversitylibrary. org / part / 50813 # / summary [accessed 27 May 2020]." type="journal article" year="1996">Ng & Richer de Forges (1996)</bibRefCitation>
|
||||
, mainly freshwater but also euryhaline; and
|
||||
<taxonomicName id="4C3ABDD84006B22560BE1A1FFDD9FA26" authority="Davie & Richer de Forges, 1996" authorityName="Davie & Richer de Forges" authorityYear="1996" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494006B22560BE1A1FFAFFFACB" box="[1254,1399,1373,1399]" italics="true" pageId="11" pageNumber="12">O. estuarius</emphasis>
|
||||
<bibRefCitation id="EFABBBAA4006B22564E51AC2FDD9FA26" author="Davie P. J. F. & Richer de Forges B." box="[189,593,1408,1434]" pageId="11" pageNumber="12" pagination="257 - 262" refId="ref12668" refString="Davie P. J. F. & Richer de Forges B. 1996. Two new species of hymenosomatid crabs from New Caledonia. Memoirs of the Queensland Museum 39 (2): 257 - 262." type="journal article" year="1996">Davie & Richer de Forges, 1996</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, exclusively brackish. An important difference between these two species is the rostrum: it is lowered ventrally as a triangular, V-shaped projection in
|
||||
<taxonomicName id="4C3ABDD84006B22560C61AE6FA9DFA01" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[1182,1301,1443,1470]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494006B22560C61AE6FA9DFA01" box="[1182,1301,1443,1470]" italics="true" pageId="11" pageNumber="12">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
so as to be positioned between the antennules (
|
||||
<figureCitation id="1301DADE4006B22566D71A84FD65FA5D" box="[655,749,1478,1505]" captionStart="Fig" captionStartId="16.[189,232,1919,1945]" captionTargetBox="[402,1193,1524,1866]" captionTargetId="figure@16.[385,1203,1509,1878]" captionTargetPageId="16" captionText="Fig. 7. Odiomaris pilosus A. Milne-Edwards, 1873, syntype, ♂, New Caledonia, freshwater, B. Balansa leg. (RMNH D 32). A. Ventral anterior part. B. G1 (after Holthuis 1968: fig. 3a–b). Abbreviations: a1 = antennule, with folded second article; a2 = antenna; e = epistome: p.k. = postorbital knobs; r = rostrum." figureDoi="http://doi.org/10.5281/zenodo.3906770" httpUri="https://zenodo.org/record/3906770/files/figure.png" pageId="11" pageNumber="12">Fig. 7A</figureCitation>
|
||||
) (A. Milne-Edwards 1873: pl. 18, fig. 6a, as
|
||||
<taxonomicName id="4C3ABDD84006B22561481A84FE8DF9B8" authorityName="A.Milne-Edwards" authorityYear="1873" class="Malacostraca" family="Hymenosomatidae" genus="Elamene" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pilosa">
|
||||
<emphasis id="B94E1A494006B22561481A84FE8DF9B8" italics="true" pageId="11" pageNumber="12">Elamene pilosa</emphasis>
|
||||
</taxonomicName>
|
||||
;
|
||||
<bibRefCitation id="EFABBBAA4006B225654E1AA8FDFCF9BF" author="Ng P. K. L. & Richer de Forges B." box="[278,628,1513,1540]" pageId="11" pageNumber="12" pagination="263 - 276" refId="ref14618" refString="Ng P. K. L. & Richer de Forges B. 1996. The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of New Caledonia, with descriptions of two new genera and two new species. Memoirs of the Queensland Museum 39 (2): 263 - 276. Available from https: // www. biodiversitylibrary. org / part / 50813 # / summary [accessed 27 May 2020]." type="journal article" year="1996">Ng & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 6c–d), whereas it is spatulate and does not extend to form a ventral projection between the antennules in
|
||||
<taxonomicName id="4C3ABDD84006B225669F194FFCDFF99B" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[711,855,1549,1575]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494006B225669F194FFCDFF99B" box="[711,855,1549,1575]" italics="true" pageId="11" pageNumber="12">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1301DADE4006B225673E194FFC21F99B" box="[870,937,1549,1575]" captionStart="Fig" captionStartId="18.[189,232,1892,1918]" captionTargetBox="[189,1398,466,1853]" captionTargetId="figure@18.[189,1398,466,1853]" captionTargetPageId="18" captionText="Fig. 8. Odiomaris estuarius Davie & Richer de Forges, 1996, holotype, ♂, New Caledonia, Dumbea estuary, Davie & Richer de Forges leg., 8 Dec. 1993, 4.5 × 4.3 mm (MNHN-IU-2014-11870 = MNHN-B25278). A. Dorsal view. B. Ventral view. Credit: RECOLNAT (ANR-11-INBS-0004) - Noémy MOLLARET – 2017." figureDoi="http://doi.org/10.5281/zenodo.3906772" httpUri="https://zenodo.org/record/3906772/files/figure.png" pageId="11" pageNumber="12">Fig. 8</figureCitation>
|
||||
A–B) (
|
||||
<bibRefCitation id="EFABBBAA4006B22567A2194FFAFBF99B" author="Davie P. J. F. & Richer de Forges B." box="[1018,1395,1549,1575]" pageId="11" pageNumber="12" pagination="257 - 262" refId="ref12668" refString="Davie P. J. F. & Richer de Forges B. 1996. Two new species of hymenosomatid crabs from New Caledonia. Memoirs of the Queensland Museum 39 (2): 257 - 262." type="journal article" year="1996">Davie & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 1a–b).
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4006B22264E51933FBAFFED0" blockId="11.[189,1399,1649,1993]" lastBlockId="12.[189,1399,267,364]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
|
||||
<taxonomicName id="4C3ABDD84006B22564E51933FEA5F937" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[189,301,1649,1675]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B94E1A494006B22564E51933FEA5F937" box="[189,301,1649,1675]" italics="true" pageId="11" pageNumber="12">Richerius</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324006B225656F1930FE2DF930" box="[311,421,1650,1676]" pageId="11" pageNumber="12" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
shares with
|
||||
<taxonomicName id="4C3ABDD84006B22566181933FD35F937" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[576,701,1649,1675]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494006B22566181933FD35F937" box="[576,701,1649,1675]" italics="true" pageId="11" pageNumber="12">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
the same arrangement of the cephalic appendices. But its rostrum is spatulate without folding down ventrally (
|
||||
<figureCitation id="1301DADE4006B225677519D6FCF0F913" box="[813,888,1684,1711]" captionStart="Fig" captionStartId="6.[189,232,1782,1808]" captionTargetBox="[326,1260,292,1710]" captionTargetId="figure@6.[292,1296,265,1741]" captionTargetPageId="6" captionText="Fig. 1. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Carapace, dorsal view. B. Cephalothorax, ventral view, anterior part, with unfolded antennules, the arrow shows antennule with folded second article. Abbreviations: a1 = antennule; a2 = antenna; e = epistome; p.k. = postorbital knob; r = rostrum; u.a. = antennal urinary article; v.r. = ventral expansion of rostrum; 2/3 = thoracic sternal suture 2/3. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3906758" httpUri="https://zenodo.org/record/3906758/files/figure.png" pageId="11" pageNumber="12">Figs 1</figureCitation>
|
||||
,
|
||||
<figureCitation id="1301DADE4006B22567DF19D7FC10F913" box="[903,920,1685,1711]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[208,1380,266,1860]" captionTargetId="figure@9.[208,1380,266,1862]" captionTargetPageId="9" captionText="Fig. 4. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Dorsal view. B. Ventral view. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3906764" httpUri="https://zenodo.org/record/3906764/files/figure.png" pageId="11" pageNumber="12">4</figureCitation>
|
||||
,
|
||||
<figureCitation id="1301DADE4006B22567FE19D6FC44F913" box="[934,972,1684,1711]" captionStart="Fig" captionStartId="10.[189,232,1855,1881]" captionTargetBox="[189,1398,901,1815]" captionTargetId="figure@10.[189,1398,901,1816]" captionTargetPageId="10" captionText="Fig. 5. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Type locality, stream where several specimens of the species were collected. B–D. Views of the holotype in vivo: dorsal (B), ventral (C) and frontal (D)." figureDoi="http://doi.org/10.5281/zenodo.3906766" httpUri="https://zenodo.org/record/3906766/files/figure.png" pageId="11" pageNumber="12">5D</figureCitation>
|
||||
), instead of being lowered ventrally as a V-shaped projection located between the antennules as in
|
||||
<taxonomicName id="4C3ABDD84006B22567D719FAFBEEF96E" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[911,1126,1720,1746]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494006B22567D719FAFBEEF96E" box="[911,1126,1720,1746]" italics="true" pageId="11" pageNumber="12">Odiomaris pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1301DADE4006B225602D19FAFB5CF96E" box="[1141,1236,1720,1746]" captionStart="Fig" captionStartId="16.[189,232,1919,1945]" captionTargetBox="[402,1193,1524,1866]" captionTargetId="figure@16.[385,1203,1509,1878]" captionTargetPageId="16" captionText="Fig. 7. Odiomaris pilosus A. Milne-Edwards, 1873, syntype, ♂, New Caledonia, freshwater, B. Balansa leg. (RMNH D 32). A. Ventral anterior part. B. G1 (after Holthuis 1968: fig. 3a–b). Abbreviations: a1 = antennule, with folded second article; a2 = antenna; e = epistome: p.k. = postorbital knobs; r = rostrum." figureDoi="http://doi.org/10.5281/zenodo.3906770" httpUri="https://zenodo.org/record/3906770/files/figure.png" pageId="11" pageNumber="12">Fig. 7A</figureCitation>
|
||||
). In
|
||||
<taxonomicName id="4C3ABDD84006B225615E19FAFAFEF96E" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[1286,1398,1720,1746]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B94E1A494006B225615E19FAFAFEF96E" box="[1286,1398,1720,1746]" italics="true" pageId="11" pageNumber="12">Richerius</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324006B22564E5199EFEA3F94A" box="[189,299,1756,1782]" pageId="11" pageNumber="12" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
(
|
||||
<figureCitation id="1301DADE4006B22565651999FE12F949" box="[317,410,1755,1781]" captionStart="Fig" captionStartId="6.[189,232,1782,1808]" captionTargetBox="[326,1260,292,1710]" captionTargetId="figure@6.[292,1296,265,1741]" captionTargetPageId="6" captionText="Fig. 1. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Carapace, dorsal view. B. Cephalothorax, ventral view, anterior part, with unfolded antennules, the arrow shows antennule with folded second article. Abbreviations: a1 = antennule; a2 = antenna; e = epistome; p.k. = postorbital knob; r = rostrum; u.a. = antennal urinary article; v.r. = ventral expansion of rostrum; 2/3 = thoracic sternal suture 2/3. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3906758" httpUri="https://zenodo.org/record/3906758/files/figure.png" pageId="11" pageNumber="12">Fig. 1B</figureCitation>
|
||||
), the proepistome is represented by a ventral expansion of the rostrum; therefore, the antennules are located along the proepistome, whereas in
|
||||
<taxonomicName id="4C3ABDD84006B22567EB19BCFB07F8A4" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[947,1167,1790,1816]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494006B22567EB19BCFB07F8A4" box="[947,1167,1790,1816]" italics="true" pageId="11" pageNumber="12">Odiomaris pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
the antennules are obliquely folded in a fossa hollowed all along with the lateral parts of the V-shaped rostrum and the eye.
|
||||
<taxonomicName id="4C3ABDD84006B22564AE1807FE50F8E2" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[246,472,1861,1887]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494006B22564AE1807FE50F8E2" box="[246,472,1861,1887]" italics="true" pageId="11" pageNumber="12">Richerius marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324006B22565B91807FD1DF8E3" box="[481,661,1861,1887]" pageId="11" pageNumber="12" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
actually has a rostrum and proepistome more similar to those of
|
||||
<taxonomicName id="4C3ABDD84006B2256487182AFEFAF83E" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[223,370,1896,1922]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494006B2256487182AFEFAF83E" box="[223,370,1896,1922]" italics="true" pageId="11" pageNumber="12">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1301DADE4006B22565DC182AFE44F83E" box="[388,460,1896,1922]" captionStart="Fig" captionStartId="18.[189,232,1892,1918]" captionTargetBox="[189,1398,466,1853]" captionTargetId="figure@18.[189,1398,466,1853]" captionTargetPageId="18" captionText="Fig. 8. Odiomaris estuarius Davie & Richer de Forges, 1996, holotype, ♂, New Caledonia, Dumbea estuary, Davie & Richer de Forges leg., 8 Dec. 1993, 4.5 × 4.3 mm (MNHN-IU-2014-11870 = MNHN-B25278). A. Dorsal view. B. Ventral view. Credit: RECOLNAT (ANR-11-INBS-0004) - Noémy MOLLARET – 2017." figureDoi="http://doi.org/10.5281/zenodo.3906772" httpUri="https://zenodo.org/record/3906772/files/figure.png" pageId="11" pageNumber="12">Fig. 8</figureCitation>
|
||||
) (
|
||||
<bibRefCitation id="EFABBBAA4006B22565BF182AFCE7F83E" author="Davie P. J. F. & Richer de Forges B." box="[487,879,1896,1922]" pageId="11" pageNumber="12" pagination="257 - 262" refId="ref12668" refString="Davie P. J. F. & Richer de Forges B. 1996. Two new species of hymenosomatid crabs from New Caledonia. Memoirs of the Queensland Museum 39 (2): 257 - 262." type="journal article" year="1996">Davie & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 1) than to those of
|
||||
<taxonomicName id="4C3ABDD84006B22560CE182AFA98F83E" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[1174,1296,1896,1922]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494006B22560CE182AFA98F83E" box="[1174,1296,1896,1922]" italics="true" pageId="11" pageNumber="12">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
. In fact, the genus-level differences between
|
||||
<taxonomicName id="4C3ABDD84006B22566DB18C9FD7BF819" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[643,755,1931,1957]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B94E1A494006B22566DB18C9FD7BF819" box="[643,755,1931,1957]" italics="true" pageId="11" pageNumber="12">Richerius</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324006B225675A18CEFCFBF81A" box="[770,883,1932,1958]" pageId="11" pageNumber="12" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
and
|
||||
<taxonomicName id="4C3ABDD84006B22567E118C9FBBEF819" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[953,1078,1931,1957]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494006B22567E118C9FBBEF819" box="[953,1078,1931,1957]" italics="true" pageId="11" pageNumber="12">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
remain unclear. Actually,
|
||||
<taxonomicName id="4C3ABDD84006B22564E518EDFEC7F874" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[189,335,1967,1993]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494006B22564E518EDFEC7F874" box="[189,335,1967,1993]" italics="true" pageId="11" pageNumber="12">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
seems morphologically closer to
|
||||
<taxonomicName id="4C3ABDD84006B22566BC18ECFCDCF874" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[740,852,1966,1992]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B94E1A494006B22566BC18ECFCDCF874" box="[740,852,1966,1992]" italics="true" pageId="11" pageNumber="12">Richerius</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324006B225670618EDFC43F875" box="[862,971,1967,1993]" pageId="11" pageNumber="12" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
than to
|
||||
<taxonomicName id="4C3ABDD84006B225606918EDFB22F874" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[1073,1194,1966,1993]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494006B225606918EDFB22F874" box="[1073,1194,1966,1993]" italics="true" pageId="11" pageNumber="12">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
, at least in some characteristics. A possible transfer of
|
||||
<taxonomicName id="4C3ABDD84001B222662D1E4EFC8DFE99" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[629,773,268,294]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494001B222662D1E4EFC8DFE99" box="[629,773,268,294]" italics="true" pageId="12" pageNumber="13">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
to
|
||||
<taxonomicName id="4C3ABDD84001B22267721E49FC12FE99" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[810,922,267,293]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B94E1A494001B22267721E49FC12FE99" box="[810,922,267,293]" italics="true" pageId="12" pageNumber="13">Richerius</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324001B22267FA1E4EFB84FE9A" box="[930,1036,268,294]" pageId="12" pageNumber="13" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
has been considered but needs to be decided later, especially when the status of
|
||||
<taxonomicName id="4C3ABDD84001B22266B71E6DFC7BFEF5" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[751,1011,302,329]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B22266B71E6DFCE4FEF5" box="[751,876,303,329]" italics="true" pageId="12" pageNumber="13">Odiomaris</emphasis>
|
||||
aff.
|
||||
<emphasis id="B94E1A494001B22267F81E6DFC7BFEF5" box="[928,1011,303,329]" italics="true" pageId="12" pageNumber="13">pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
is resolved (see below, ‘Remarks on
|
||||
<taxonomicName id="4C3ABDD84001B22264BA1E10FE60FED0" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[226,488,338,364]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B22264BA1E10FED7FED0" box="[226,351,338,364]" italics="true" pageId="12" pageNumber="13">Odiomaris</emphasis>
|
||||
aff.
|
||||
<emphasis id="B94E1A494001B22265CD1E10FE60FED0" box="[405,488,338,364]" italics="true" pageId="12" pageNumber="13">pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
from the Iouanga River and two other streams’).
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4001B22264E51ED1FAD8FA17" blockId="12.[189,1399,403,1451]" pageId="12" pageNumber="13">
|
||||
Along the lower orbital margin there are two salient knobs in both species of
|
||||
<taxonomicName id="4C3ABDD84001B22260601ED1FB3DFE11" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[1080,1205,403,429]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B22260601ED1FB3DFE11" box="[1080,1205,403,429]" italics="true" pageId="12" pageNumber="13">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
, both prominent in dorsal view in
|
||||
<taxonomicName id="4C3ABDD84001B22265DA1EF5FE7FFE6C" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[386,503,438,465]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B22265DA1EF5FE7FFE6C" box="[386,503,438,465]" italics="true" pageId="12" pageNumber="13">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
(A. Milne-Edwards 1873: pl. 18, fig. 6a;
|
||||
<bibRefCitation id="EFABBBAA4001B22267881EF5FA96FE6C" author="Ng P. K. L. & Richer de Forges B." box="[976,1310,438,465]" pageId="12" pageNumber="13" pagination="263 - 276" refId="ref14618" refString="Ng P. K. L. & Richer de Forges B. 1996. The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of New Caledonia, with descriptions of two new genera and two new species. Memoirs of the Queensland Museum 39 (2): 263 - 276. Available from https: // www. biodiversitylibrary. org / part / 50813 # / summary [accessed 27 May 2020]." type="journal article" year="1996">Ng & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 6a, d, not figured in fig. 6c;
|
||||
<bibRefCitation id="EFABBBAA4001B22265BA1E98FCE7FE48" author="Richer de Forges B. & Jamieson B. G. M. & Guinot D. & Tudge C. C." box="[482,879,474,500]" pageId="12" pageNumber="13" pagination="233 - 242" refId="ref15566" refString="Richer de Forges B., Jamieson B. G. M., Guinot D. & Tudge C. C. 1997. Ultrastructure of the spermatozoa of Hymenosomatidae (Crustacea: Brachyura) and the relationships of the family. Marine Biology 130: 233 - 242. https: // doi. org / 10.1007 / s 002270050243" type="journal article" year="1997">Guinot & Richer de Forges 1997</bibRefCitation>
|
||||
: figs 1a, c, 2c), but shorter and not dorsally visible in
|
||||
<taxonomicName id="4C3ABDD84001B22265681EBFFE37FDAB" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[304,447,509,535]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494001B22265681EBFFE37FDAB" box="[304,447,509,535]" italics="true" pageId="12" pageNumber="13">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<bibRefCitation id="EFABBBAA4001B22265961EBFFCC0FDAB" author="Davie P. J. F. & Richer de Forges B." box="[462,840,509,535]" pageId="12" pageNumber="13" pagination="257 - 262" refId="ref12668" refString="Davie P. J. F. & Richer de Forges B. 1996. Two new species of hymenosomatid crabs from New Caledonia. Memoirs of the Queensland Museum 39 (2): 257 - 262." type="journal article" year="1996">Davie & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 1a; not figured in fig. 1b). In contrast, there is only one, smaller knob in
|
||||
<taxonomicName id="4C3ABDD84001B22266401D62FD00FD86" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[536,648,544,570]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B94E1A494001B22266401D62FD00FD86" box="[536,648,544,570]" italics="true" pageId="12" pageNumber="13">Richerius</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324001B22266CA1D63FD77FD87" box="[658,767,545,571]" pageId="12" pageNumber="13" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
(
|
||||
<figureCitation id="1301DADE4001B22267491D62FCE6FD86" box="[785,878,544,570]" captionStart="Fig" captionStartId="6.[189,232,1782,1808]" captionTargetBox="[326,1260,292,1710]" captionTargetId="figure@6.[292,1296,265,1741]" captionTargetPageId="6" captionText="Fig. 1. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Carapace, dorsal view. B. Cephalothorax, ventral view, anterior part, with unfolded antennules, the arrow shows antennule with folded second article. Abbreviations: a1 = antennule; a2 = antenna; e = epistome; p.k. = postorbital knob; r = rostrum; u.a. = antennal urinary article; v.r. = ventral expansion of rostrum; 2/3 = thoracic sternal suture 2/3. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3906758" httpUri="https://zenodo.org/record/3906758/files/figure.png" pageId="12" pageNumber="13">Fig. 1B</figureCitation>
|
||||
). The cheliped palm of
|
||||
<taxonomicName id="4C3ABDD84001B22260C91D63FA95FD86" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[1169,1309,545,570]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494001B22260C91D63FA95FD86" box="[1169,1309,545,570]" italics="true" pageId="12" pageNumber="13">R. marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324001B222617E1D63FE9FFDE2" pageId="12" pageNumber="13" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
is covered with long, flexible setae (
|
||||
<figureCitation id="1301DADE4001B222669C1D01FC80FDE2" box="[708,776,579,606]" captionStart="Fig" captionStartId="10.[189,232,1855,1881]" captionTargetBox="[189,1398,901,1815]" captionTargetId="figure@10.[189,1398,901,1816]" captionTargetPageId="10" captionText="Fig. 5. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Type locality, stream where several specimens of the species were collected. B–D. Views of the holotype in vivo: dorsal (B), ventral (C) and frontal (D)." figureDoi="http://doi.org/10.5281/zenodo.3906766" httpUri="https://zenodo.org/record/3906766/files/figure.png" pageId="12" pageNumber="13">Fig. 5</figureCitation>
|
||||
C–D), instead of numerous spinules (or stiff setae) in
|
||||
<taxonomicName id="4C3ABDD84001B22264E51D25FEBCFD3C" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[189,308,614,641]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B22264E51D25FEBCFD3C" box="[189,308,614,641]" italics="true" pageId="12" pageNumber="13">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1301DADE4001B222651D1D24FE03FD3D" box="[325,395,614,641]" captionStart="Fig" captionStartId="15.[189,232,1921,1947]" captionTargetBox="[189,1398,300,1857]" captionTargetId="figure@15.[189,1398,296,1857]" captionTargetPageId="15" captionText="Fig. 6. Odiomaris pilosus A. Milne-Edwards, 1873, living specimens. A. Collected from the river Hiit (North Province, Hienghène township). B. Collected from the Baie Nord creek (South Province, Mont- Dore township). Credit: Nicolas Charpin – Photographies Vies d’Ô douce." figureDoi="http://doi.org/10.5281/zenodo.3906768" httpUri="https://zenodo.org/record/3906768/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
|
||||
) (A. Milne-Edwards, 1873: pl. 18, fig. 6d;
|
||||
<bibRefCitation id="EFABBBAA4001B22267D01D25FB69FD3C" author="Ng P. K. L. & Richer de Forges B." box="[904,1249,614,641]" pageId="12" pageNumber="13" pagination="263 - 276" refId="ref14618" refString="Ng P. K. L. & Richer de Forges B. 1996. The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of New Caledonia, with descriptions of two new genera and two new species. Memoirs of the Queensland Museum 39 (2): 263 - 276. Available from https: // www. biodiversitylibrary. org / part / 50813 # / summary [accessed 27 May 2020]." type="journal article" year="1996">Ng & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 6f), and sparse, short setae in
|
||||
<taxonomicName id="4C3ABDD84001B22265EA1DC8FDC9FD1F" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[434,577,650,676]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494001B22265EA1DC8FDC9FD1F" box="[434,577,650,676]" italics="true" pageId="12" pageNumber="13">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1301DADE4001B22266171DC8FD1DFD18" box="[591,661,650,676]" captionStart="Fig" captionStartId="18.[189,232,1892,1918]" captionTargetBox="[189,1398,466,1853]" captionTargetId="figure@18.[189,1398,466,1853]" captionTargetPageId="18" captionText="Fig. 8. Odiomaris estuarius Davie & Richer de Forges, 1996, holotype, ♂, New Caledonia, Dumbea estuary, Davie & Richer de Forges leg., 8 Dec. 1993, 4.5 × 4.3 mm (MNHN-IU-2014-11870 = MNHN-B25278). A. Dorsal view. B. Ventral view. Credit: RECOLNAT (ANR-11-INBS-0004) - Noémy MOLLARET – 2017." figureDoi="http://doi.org/10.5281/zenodo.3906772" httpUri="https://zenodo.org/record/3906772/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
|
||||
). In fact, the whole body (including the margins of the rostrum) and legs of
|
||||
<taxonomicName id="4C3ABDD84001B222651A1DEFFE30FD7B" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[322,440,685,711]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B222651A1DEFFE30FD7B" box="[322,440,685,711]" italics="true" pageId="12" pageNumber="13">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
are covered with spiniform, stiff setae that give a bristle appearance (
|
||||
<bibRefCitation id="EFABBBAA4001B22260871DEFFEFEFD56" author="Ng P. K. L. & Richer de Forges B." pageId="12" pageNumber="13" pagination="263 - 276" refId="ref14618" refString="Ng P. K. L. & Richer de Forges B. 1996. The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of New Caledonia, with descriptions of two new genera and two new species. Memoirs of the Queensland Museum 39 (2): 263 - 276. Available from https: // www. biodiversitylibrary. org / part / 50813 # / summary [accessed 27 May 2020]." type="journal article" year="1996">Ng & Richer de Forges 1996</bibRefCitation>
|
||||
: figs 5a–b, 6a, d;
|
||||
<bibRefCitation id="EFABBBAA4001B222661E1D92FC45FD56" author="Richer de Forges B. & Jamieson B. G. M. & Guinot D. & Tudge C. C." box="[582,973,720,747]" pageId="12" pageNumber="13" pagination="233 - 242" refId="ref15566" refString="Richer de Forges B., Jamieson B. G. M., Guinot D. & Tudge C. C. 1997. Ultrastructure of the spermatozoa of Hymenosomatidae (Crustacea: Brachyura) and the relationships of the family. Marine Biology 130: 233 - 242. https: // doi. org / 10.1007 / s 002270050243" type="journal article" year="1997">Guinot & Richer de Forges 1997</bibRefCitation>
|
||||
: fig. 1a), whereas
|
||||
<taxonomicName id="4C3ABDD84001B22260FB1D92FABBFD56" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[1187,1331,720,746]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494001B22260FB1D92FABBFD56" box="[1187,1331,720,746]" italics="true" pageId="12" pageNumber="13">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
has a carapace with soft, short setae and legs with longer setae (
|
||||
<bibRefCitation id="EFABBBAA4001B222673B1DB1FB55FCB1" author="Davie P. J. F. & Richer de Forges B." box="[867,1245,755,782]" pageId="12" pageNumber="13" pagination="257 - 262" refId="ref12668" refString="Davie P. J. F. & Richer de Forges B. 1996. Two new species of hymenosomatid crabs from New Caledonia. Memoirs of the Queensland Museum 39 (2): 257 - 262." type="journal article" year="1996">Davie & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 1a). The male cheliped fingers are practically joining in
|
||||
<taxonomicName id="4C3ABDD84001B22266B41C55FCFEFC8C" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[748,886,791,816]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494001B22266B41C55FCFEFC8C" box="[748,886,791,816]" italics="true" pageId="12" pageNumber="13">R. marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324001B22267261C55FBB9FC8D" box="[894,1073,791,817]" pageId="12" pageNumber="13" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
(
|
||||
<figureCitation id="1301DADE4001B22260191C54FB16FC8D" box="[1089,1182,790,817]" captionStart="Fig" captionStartId="10.[189,232,1855,1881]" captionTargetBox="[189,1398,901,1815]" captionTargetId="figure@10.[189,1398,901,1816]" captionTargetPageId="10" captionText="Fig. 5. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Type locality, stream where several specimens of the species were collected. B–D. Views of the holotype in vivo: dorsal (B), ventral (C) and frontal (D)." figureDoi="http://doi.org/10.5281/zenodo.3906766" httpUri="https://zenodo.org/record/3906766/files/figure.png" pageId="12" pageNumber="13">Fig. 5C</figureCitation>
|
||||
), but with a broad proximal gap in
|
||||
<taxonomicName id="4C3ABDD84001B22265DC1C78FE75FCE8" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[388,509,826,852]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B22265DC1C78FE75FCE8" box="[388,509,826,852]" italics="true" pageId="12" pageNumber="13">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1301DADE4001B22266481C78FDDFFCE8" box="[528,599,825,852]" captionStart="Fig" captionStartId="15.[189,232,1921,1947]" captionTargetBox="[189,1398,300,1857]" captionTargetId="figure@15.[189,1398,296,1857]" captionTargetPageId="15" captionText="Fig. 6. Odiomaris pilosus A. Milne-Edwards, 1873, living specimens. A. Collected from the river Hiit (North Province, Hienghène township). B. Collected from the Baie Nord creek (South Province, Mont- Dore township). Credit: Nicolas Charpin – Photographies Vies d’Ô douce." figureDoi="http://doi.org/10.5281/zenodo.3906768" httpUri="https://zenodo.org/record/3906768/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
|
||||
A–B) (A. Milne-Edwards 1873: 322, pl. 18, fig. 6, 6d, as
|
||||
<taxonomicName id="4C3ABDD84001B22261481C78FE8DFCCB" authorityName="A.Milne-Edwards" authorityYear="1873" class="Malacostraca" family="Hymenosomatidae" genus="Elamene" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosa">
|
||||
<emphasis id="B94E1A494001B22261481C78FE8DFCCB" italics="true" pageId="12" pageNumber="13">Elamene pilosa</emphasis>
|
||||
</taxonomicName>
|
||||
;
|
||||
<bibRefCitation id="EFABBBAA4001B222654C1C1FFDE5FCCB" author="Ng P. K. L. & Richer de Forges B." box="[276,621,861,887]" pageId="12" pageNumber="13" pagination="263 - 276" refId="ref14618" refString="Ng P. K. L. & Richer de Forges B. 1996. The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of New Caledonia, with descriptions of two new genera and two new species. Memoirs of the Queensland Museum 39 (2): 263 - 276. Available from https: // www. biodiversitylibrary. org / part / 50813 # / summary [accessed 27 May 2020]." type="journal article" year="1996">Ng & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 6f); in
|
||||
<taxonomicName id="4C3ABDD84001B22266AF1C1FFC0FFCCB" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[759,903,861,887]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494001B22266AF1C1FFC0FFCCB" box="[759,903,861,887]" italics="true" pageId="12" pageNumber="13">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
the chelae are narrower, with long fingers without marked gap (
|
||||
<figureCitation id="1301DADE4001B22265EF1CC2FE73FC26" box="[439,507,896,922]" captionStart="Fig" captionStartId="18.[189,232,1892,1918]" captionTargetBox="[189,1398,466,1853]" captionTargetId="figure@18.[189,1398,466,1853]" captionTargetPageId="18" captionText="Fig. 8. Odiomaris estuarius Davie & Richer de Forges, 1996, holotype, ♂, New Caledonia, Dumbea estuary, Davie & Richer de Forges leg., 8 Dec. 1993, 4.5 × 4.3 mm (MNHN-IU-2014-11870 = MNHN-B25278). A. Dorsal view. B. Ventral view. Credit: RECOLNAT (ANR-11-INBS-0004) - Noémy MOLLARET – 2017." figureDoi="http://doi.org/10.5281/zenodo.3906772" httpUri="https://zenodo.org/record/3906772/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
|
||||
A–B) (
|
||||
<bibRefCitation id="EFABBBAA4001B22266151CC2FC40FC26" author="Davie P. J. F. & Richer de Forges B." box="[589,968,896,923]" pageId="12" pageNumber="13" pagination="257 - 262" refId="ref12668" refString="Davie P. J. F. & Richer de Forges B. 1996. Two new species of hymenosomatid crabs from New Caledonia. Memoirs of the Queensland Museum 39 (2): 257 - 262." type="journal article" year="1996">Davie & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 1). The grooves on the carapace dorsal surface are very distinct in
|
||||
<taxonomicName id="4C3ABDD84001B222661B1CE6FD31FC01" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[579,697,931,958]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B222661B1CE6FD31FC01" box="[579,697,931,958]" italics="true" pageId="12" pageNumber="13">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
, well marked in
|
||||
<taxonomicName id="4C3ABDD84001B22267221CE6FB8BFC01" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[890,1027,932,957]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494001B22267221CE6FB8BFC01" box="[890,1027,932,957]" italics="true" pageId="12" pageNumber="13">R. marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324001B22260501CE6FB3FFC02" box="[1032,1207,932,958]" pageId="12" pageNumber="13" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
, whereas almost indiscernible in
|
||||
<taxonomicName id="4C3ABDD84001B22265201C85FD8FFC5C" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[376,519,967,993]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494001B22265201C85FD8FFC5C" box="[376,519,967,993]" italics="true" pageId="12" pageNumber="13">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
. The sterno-pleonal cavity is short in
|
||||
<taxonomicName id="4C3ABDD84001B22267E31C84FBA3FC5C" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[955,1067,966,992]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B94E1A494001B22267E31C84FBA3FC5C" box="[955,1067,966,992]" italics="true" pageId="12" pageNumber="13">Richerius</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324001B222606A1C85FB15FC5C" box="[1074,1181,967,993]" pageId="12" pageNumber="13" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
and in
|
||||
<taxonomicName id="4C3ABDD84001B22260AA1C84FAE7FC5C" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[1266,1391,966,992]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B22260AA1C84FAE7FC5C" box="[1266,1391,966,992]" italics="true" pageId="12" pageNumber="13">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
, and accordingly the pleon also. The male pleon is as a narrow triangle, with a long pleotelson in both
|
||||
<taxonomicName id="4C3ABDD84001B22264E51B4FFEB2FB9B" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[189,314,1037,1063]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B22264E51B4FFEB2FB9B" box="[189,314,1037,1063]" italics="true" pageId="12" pageNumber="13">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
species (A. Milne-Edwards 1873: pl. 18, fig. 6b, as
|
||||
<taxonomicName id="4C3ABDD84001B22267C71B4FFBDDFB9B" authorityName="A.Milne-Edwards" authorityYear="1873" box="[927,1109,1037,1063]" class="Malacostraca" family="Hymenosomatidae" genus="Elamene" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosa">
|
||||
<emphasis id="B94E1A494001B22267C71B4FFBDDFB9B" box="[927,1109,1037,1063]" italics="true" pageId="12" pageNumber="13">Elamene pilosa</emphasis>
|
||||
</taxonomicName>
|
||||
;
|
||||
<bibRefCitation id="EFABBBAA4001B222603D1B4FFF74FBF6" author="Ng P. K. L. & Richer de Forges B." pageId="12" pageNumber="13" pagination="263 - 276" refId="ref14618" refString="Ng P. K. L. & Richer de Forges B. 1996. The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of New Caledonia, with descriptions of two new genera and two new species. Memoirs of the Queensland Museum 39 (2): 263 - 276. Available from https: // www. biodiversitylibrary. org / part / 50813 # / summary [accessed 27 May 2020]." type="journal article" year="1996">Ng & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 7a–b;
|
||||
<bibRefCitation id="EFABBBAA4001B22265271B72FC80FBF6" author="Richer de Forges B. & Jamieson B. G. M. & Guinot D. & Tudge C. C." box="[383,776,1072,1099]" pageId="12" pageNumber="13" pagination="233 - 242" refId="ref15566" refString="Richer de Forges B., Jamieson B. G. M., Guinot D. & Tudge C. C. 1997. Ultrastructure of the spermatozoa of Hymenosomatidae (Crustacea: Brachyura) and the relationships of the family. Marine Biology 130: 233 - 242. https: // doi. org / 10.1007 / s 002270050243" type="journal article" year="1997">Guinot & Richer de Forges 1997</bibRefCitation>
|
||||
: fig. 2a:
|
||||
<taxonomicName id="4C3ABDD84001B22267361B72FC6DFBF6" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[878,997,1072,1098]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B22267361B72FC6DFBF6" box="[878,997,1072,1098]" italics="true" pageId="12" pageNumber="13">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
;
|
||||
<bibRefCitation id="EFABBBAA4001B22267AE1B72FAFBFBF6" author="Davie P. J. F. & Richer de Forges B." box="[1014,1395,1072,1099]" pageId="12" pageNumber="13" pagination="257 - 262" refId="ref12668" refString="Davie P. J. F. & Richer de Forges B. 1996. Two new species of hymenosomatid crabs from New Caledonia. Memoirs of the Queensland Museum 39 (2): 257 - 262." type="journal article" year="1996">Davie & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 2c:
|
||||
<taxonomicName id="4C3ABDD84001B222654D1B16FE2CFBD1" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[277,420,1108,1134]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494001B222654D1B16FE2CFBD1" box="[277,420,1108,1134]" italics="true" pageId="12" pageNumber="13">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
), markedly wider and with a much shorter pleotelson in
|
||||
<taxonomicName id="4C3ABDD84001B22260601B16FB49FBD1" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[1080,1217,1108,1133]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494001B22260601B16FB49FBD1" box="[1080,1217,1108,1133]" italics="true" pageId="12" pageNumber="13">R. marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324001B22260901B16FAFFFBD2" box="[1224,1399,1108,1134]" pageId="12" pageNumber="13" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
(
|
||||
<figureCitation id="1301DADE4001B222649E1B34FEAEFB2D" box="[198,294,1142,1169]" captionStart="Fig" captionStartId="7.[189,232,1846,1872]" captionTargetBox="[260,1308,282,1788]" captionTargetId="figure@7.[248,1339,265,1807]" captionTargetPageId="7" captionText="Fig. 2. Richerius marqueti gen. et sp. nov., holotype, ♂, 4.9 × 5.0 mm (MNHN-IU-2014-21500), same collection data as Fig. 1. A. Thoracic sternum. B. Mxp3. C. Sternopleonal cavity with G1s. D. G1, two views. Abbreviations: G1 = gonopod 1; G2 = gonopod 2; i = intercalated platelets; p = pleotelson; p.b. = press-button; s4 = expansion of sternite 4; 1–8 = thoracic sternites 1–8; 2/3–7/8 = thoracic sternal sutures 2/3–7/8. Scale bars: A = 2 mm; B–C = 1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3906760" httpUri="https://zenodo.org/record/3906760/files/figure.png" pageId="12" pageNumber="13">Fig. 2A</figureCitation>
|
||||
). The intercalated platelets, which are completely demarcated and moveable in
|
||||
<taxonomicName id="4C3ABDD84001B22260A31B35FAFEFB2C" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[1275,1398,1142,1169]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B22260A31B35FAFEFB2C" box="[1275,1398,1142,1169]" italics="true" pageId="12" pageNumber="13">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<bibRefCitation id="EFABBBAA4001B222649D1BD8FEE3FB08" author="Guinot D." box="[197,363,1178,1204]" pageId="12" pageNumber="13" pagination="20 - 32" refId="ref13311" refString="Guinot D. 2011 a. Odiomarinae nov. subfam., a new subfamily for two primitive genera of the Hymenosomatidae MacLeay, 1838 (Crustacea, Decapoda, Brachyura). Zootaxa 2732: 20 - 32. https: // doi. org / 10.11646 / zootaxa. 2732.1.2" type="journal article" year="2011">Guinot 2011a</bibRefCitation>
|
||||
: fig. 2), are only partially delineated in
|
||||
<taxonomicName id="4C3ABDD84001B22267651BD8FC4FFB0F" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[829,967,1178,1203]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494001B22267651BD8FC4FFB0F" box="[829,967,1178,1203]" italics="true" pageId="12" pageNumber="13">R. marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324001B22267971BD8FB0BFB08" box="[975,1155,1178,1204]" pageId="12" pageNumber="13" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
, and not detached in
|
||||
<emphasis id="B94E1A494001B22264E51BFFFEDCFB6B" box="[189,340,1213,1239]" italics="true" pageId="12" pageNumber="13">
|
||||
<taxonomicName id="4C3ABDD84001B22264E51BFFFEC7FB6B" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[189,335,1213,1239]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="estuarius">O. estuarius</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
The female pleon is rather similar in
|
||||
<taxonomicName id="4C3ABDD84001B22267521BFFFCF2FB6B" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[778,890,1213,1239]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B94E1A494001B22267521BFFFCF2FB6B" box="[778,890,1213,1239]" italics="true" pageId="12" pageNumber="13">Richerius</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324001B22267D91BFFFC63FB6B" box="[897,1003,1213,1239]" pageId="12" pageNumber="13" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
and
|
||||
<taxonomicName id="4C3ABDD84001B222607B1BFFFB28FB6B" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[1059,1184,1213,1239]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B222607B1BFFFB28FB6B" box="[1059,1184,1213,1239]" italics="true" pageId="12" pageNumber="13">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
, with five somites plus pleotelson. The G1 of
|
||||
<taxonomicName id="4C3ABDD84001B22266541BA2FDF4FB46" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[524,636,1248,1274]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B94E1A494001B22266541BA2FDF4FB46" box="[524,636,1248,1274]" italics="true" pageId="12" pageNumber="13">Richerius</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324001B22266DE1BA3FD7CFB47" box="[646,756,1249,1275]" pageId="12" pageNumber="13" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
(
|
||||
<figureCitation id="1301DADE4001B222675F1BA2FCEEFB46" box="[775,870,1248,1274]" captionStart="Fig" captionStartId="7.[189,232,1846,1872]" captionTargetBox="[260,1308,282,1788]" captionTargetId="figure@7.[248,1339,265,1807]" captionTargetPageId="7" captionText="Fig. 2. Richerius marqueti gen. et sp. nov., holotype, ♂, 4.9 × 5.0 mm (MNHN-IU-2014-21500), same collection data as Fig. 1. A. Thoracic sternum. B. Mxp3. C. Sternopleonal cavity with G1s. D. G1, two views. Abbreviations: G1 = gonopod 1; G2 = gonopod 2; i = intercalated platelets; p = pleotelson; p.b. = press-button; s4 = expansion of sternite 4; 1–8 = thoracic sternites 1–8; 2/3–7/8 = thoracic sternal sutures 2/3–7/8. Scale bars: A = 2 mm; B–C = 1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3906760" httpUri="https://zenodo.org/record/3906760/files/figure.png" pageId="12" pageNumber="13">Fig. 2D</figureCitation>
|
||||
) is curved at the base, otherwise with little curvature, with a terminal portion bearing fine setae and ending in one lobe without corneous process, whereas the G1 of
|
||||
<taxonomicName id="4C3ABDD84001B22265CE1A64FD9BFAFC" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[406,531,1318,1344]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B22265CE1A64FD9BFAFC" box="[406,531,1318,1344]" italics="true" pageId="12" pageNumber="13">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
is characterised by two distinct distal processes, a longer corneous process and a shorter lobular elongation of the stem (
|
||||
<figureCitation id="1301DADE4001B222669F1A08FCACFAD8" box="[711,804,1354,1380]" captionStart="Fig" captionStartId="16.[189,232,1919,1945]" captionTargetBox="[402,1193,1524,1866]" captionTargetId="figure@16.[385,1203,1509,1878]" captionTargetPageId="16" captionText="Fig. 7. Odiomaris pilosus A. Milne-Edwards, 1873, syntype, ♂, New Caledonia, freshwater, B. Balansa leg. (RMNH D 32). A. Ventral anterior part. B. G1 (after Holthuis 1968: fig. 3a–b). Abbreviations: a1 = antennule, with folded second article; a2 = antenna; e = epistome: p.k. = postorbital knobs; r = rostrum." figureDoi="http://doi.org/10.5281/zenodo.3906770" httpUri="https://zenodo.org/record/3906770/files/figure.png" pageId="12" pageNumber="13">Fig. 7B</figureCitation>
|
||||
:
|
||||
<taxonomicName id="4C3ABDD84001B22267771A08FC2DFAD8" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[815,933,1354,1380]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B22267771A08FC2DFAD8" box="[815,933,1354,1380]" italics="true" pageId="12" pageNumber="13">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
) (
|
||||
<bibRefCitation id="EFABBBAA4001B22267E61A08FA9CFADF" author="Ng P. K. L. & Richer de Forges B." box="[958,1300,1353,1380]" pageId="12" pageNumber="13" pagination="263 - 276" refId="ref14618" refString="Ng P. K. L. & Richer de Forges B. 1996. The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of New Caledonia, with descriptions of two new genera and two new species. Memoirs of the Queensland Museum 39 (2): 263 - 276. Available from https: // www. biodiversitylibrary. org / part / 50813 # / summary [accessed 27 May 2020]." type="journal article" year="1996">Ng & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 7c– d:
|
||||
<taxonomicName id="4C3ABDD84001B22264851A2FFEDFFA3B" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[221,343,1389,1415]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B22264851A2FFEDFFA3B" box="[221,343,1389,1415]" italics="true" pageId="12" pageNumber="13">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
;
|
||||
<bibRefCitation id="EFABBBAA4001B22265311A2FFD66FA3B" author="Davie P. J. F. & Richer de Forges B." box="[361,750,1389,1415]" pageId="12" pageNumber="13" pagination="257 - 262" refId="ref12668" refString="Davie P. J. F. & Richer de Forges B. 1996. Two new species of hymenosomatid crabs from New Caledonia. Memoirs of the Queensland Museum 39 (2): 257 - 262." type="journal article" year="1996">Davie & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 2c:
|
||||
<taxonomicName id="4C3ABDD84001B22267011A2FFC63FA3B" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[857,1003,1389,1415]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494001B22267011A2FFC63FA3B" box="[857,1003,1389,1415]" italics="true" pageId="12" pageNumber="13">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
).
|
||||
<taxonomicName id="4C3ABDD84001B222605E1A2FFB10FA3B" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[1030,1176,1389,1415]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494001B222605E1A2FFB10FA3B" box="[1030,1176,1389,1415]" italics="true" pageId="12" pageNumber="13">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
is a much smaller species than
|
||||
<taxonomicName id="4C3ABDD84001B222650B1AD2FE42FA16" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[339,458,1424,1450]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B222650B1AD2FE42FA16" box="[339,458,1424,1450]" italics="true" pageId="12" pageNumber="13">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="4C3ABDD84001B22266521AD3FD1CFA16" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[522,660,1425,1450]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494001B22266521AD3FD1CFA16" box="[522,660,1425,1450]" italics="true" pageId="12" pageNumber="13">R. marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324001B22266C31AD3FCC3FA17" box="[667,843,1425,1451]" pageId="12" pageNumber="13" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
seems to be nearly as small as
|
||||
<taxonomicName id="4C3ABDD84001B22260E11AD2FAC1FA16" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[1209,1353,1424,1450]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494001B22260E11AD2FAC1FA16" box="[1209,1353,1424,1450]" italics="true" pageId="12" pageNumber="13">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4001B22264E51A93FA91F9C4" blockId="12.[189,1399,1489,1657]" pageId="12" pageNumber="13">
|
||||
The main differences to distinguish the two genera
|
||||
<taxonomicName id="4C3ABDD84001B22267421A93FC1FFA57" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[794,919,1489,1515]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B22267421A93FC1FFA57" box="[794,919,1489,1515]" italics="true" pageId="12" pageNumber="13">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C3ABDD84001B22267891A90FBCBFA50" authorityName="Lucas" authorityYear="1980" box="[977,1091,1490,1516]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B22267891A90FBCBFA50" box="[977,1091,1490,1516]" italics="true" pageId="12" pageNumber="13">Amarinus</emphasis>
|
||||
</taxonomicName>
|
||||
stated by
|
||||
<bibRefCitation id="EFABBBAA4001B22260E41A93FE36F9B3" author="Davie P. J. F. & Richer de Forges B." pageId="12" pageNumber="13" pagination="257 - 262" refId="ref12668" refString="Davie P. J. F. & Richer de Forges B. 1996. Two new species of hymenosomatid crabs from New Caledonia. Memoirs of the Queensland Museum 39 (2): 257 - 262." type="journal article" year="1996">Davie & Richer de Forges (1996: 259)</bibRefCitation>
|
||||
were: 1) in
|
||||
<taxonomicName id="4C3ABDD84001B22266121AB7FD4FF9B3" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[586,711,1525,1551]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B22266121AB7FD4FF9B3" box="[586,711,1525,1551]" italics="true" pageId="12" pageNumber="13">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
, the G1 (see
|
||||
<figureCitation id="1301DADE4001B22267381AB7FC30F9B3" box="[864,952,1525,1551]" captionStart="Fig" captionStartId="16.[189,232,1919,1945]" captionTargetBox="[402,1193,1524,1866]" captionTargetId="figure@16.[385,1203,1509,1878]" captionTargetPageId="16" captionText="Fig. 7. Odiomaris pilosus A. Milne-Edwards, 1873, syntype, ♂, New Caledonia, freshwater, B. Balansa leg. (RMNH D 32). A. Ventral anterior part. B. G1 (after Holthuis 1968: fig. 3a–b). Abbreviations: a1 = antennule, with folded second article; a2 = antenna; e = epistome: p.k. = postorbital knobs; r = rostrum." figureDoi="http://doi.org/10.5281/zenodo.3906770" httpUri="https://zenodo.org/record/3906770/files/figure.png" pageId="12" pageNumber="13">Fig. 7B</figureCitation>
|
||||
) more slender, with two distinct distal processes, a longer corneous process and a shorter lobular elongation of the stem than in
|
||||
<taxonomicName id="4C3ABDD84001B2226099195AFABBF98E" authorityName="Lucas" authorityYear="1980" box="[1217,1331,1560,1586]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B2226099195AFABBF98E" box="[1217,1331,1560,1586]" italics="true" pageId="12" pageNumber="13">Amarinus</emphasis>
|
||||
</taxonomicName>
|
||||
; 2) in
|
||||
<taxonomicName id="4C3ABDD84001B22264E51979FEB2F9E9" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[189,314,1595,1621]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B22264E51979FEB2F9E9" box="[189,314,1595,1621]" italics="true" pageId="12" pageNumber="13">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
, the elongated triangular telson of the male pleon is significantly longer than wide at base, whereas in
|
||||
<taxonomicName id="4C3ABDD84001B222651B191DFE3DF9C5" authorityName="Lucas" authorityYear="1980" box="[323,437,1631,1657]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B222651B191DFE3DF9C5" box="[323,437,1631,1657]" italics="true" pageId="12" pageNumber="13">Amarinus</emphasis>
|
||||
</taxonomicName>
|
||||
the telson is more or less rounded and short, being much wider than long.
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4001B22364E519E2FDEBFEF4" blockId="12.[189,1399,1696,2004]" lastBlockId="13.[189,1398,267,328]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
|
||||
The genus
|
||||
<taxonomicName id="4C3ABDD84001B222651D19E2FDDEF906" authority="Lucas, 1980" authorityName="Lucas" authorityYear="1980" box="[325,598,1696,1722]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B222651D19E2FE3FF906" box="[325,439,1696,1722]" italics="true" pageId="12" pageNumber="13">Amarinus</emphasis>
|
||||
<bibRefCitation id="EFABBBAA4001B222659B19E2FDDEF906" author="Lucas J. S." box="[451,598,1696,1722]" pageId="12" pageNumber="13" pagination="148 - 247" refId="ref14163" refString="Lucas J. S. 1980. Spider crabs of the family Hymenosomatidae (Crustacea: Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum 33 (4): 148 - 247. https: // doi. org / 10.3853 / j. 0067 - 1975.33.1980.199" type="journal article" year="1980">Lucas, 1980</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(
|
||||
<typeStatus id="548178F94001B222663119E2FD15F906" box="[617,669,1696,1722]" pageId="12" pageNumber="13">type</typeStatus>
|
||||
species by original designation:
|
||||
<taxonomicName id="4C3ABDD84001B222606B19E2FF71F961" authority="Chilton, 1882" authorityName="Chilton" authorityYear="1882" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="lacustris">
|
||||
<emphasis id="B94E1A494001B222606B19E2FA84F906" box="[1075,1292,1696,1722]" italics="true" pageId="12" pageNumber="13">Elamena lacustris</emphasis>
|
||||
Chilton, 1882
|
||||
</taxonomicName>
|
||||
) is known from more than ten species. They all inhabit low salinity environments, from brackish habitats to pure freshwater waters permanently (streams, lakes, swamps), and have a large distribution (
|
||||
<collectingCountry id="F32D86CB4001B222649D184BFEE0F89F" box="[197,360,1801,1827]" name="New Zealand" pageId="12" pageNumber="13">New Zealand</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="F32D86CB4001B222652D184BFE6CF89F" box="[373,484,1801,1827]" name="Australia" pageId="12" pageNumber="13">Australia</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="F32D86CB4001B22265A9184BFDEEF89F" box="[497,614,1801,1827]" name="Indonesia" pageId="12" pageNumber="13">Indonesia</collectingCountry>
|
||||
, the
|
||||
<collectingCountry id="F32D86CB4001B22266F8184BFCAFF89F" box="[672,807,1801,1827]" name="Philippines" pageId="12" pageNumber="13">Philippines</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="F32D86CB4001B222676D1848FBA9F898" box="[821,1057,1801,1828]" name="Papua New Guinea" pageId="12" pageNumber="13">Papua New Guinea</collectingCountry>
|
||||
). Cases where
|
||||
<taxonomicName id="4C3ABDD84001B222608C1848FACEF898" authorityName="Lucas" authorityYear="1980" box="[1236,1350,1802,1828]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B222608C1848FACEF898" box="[1236,1350,1802,1828]" italics="true" pageId="12" pageNumber="13">Amarinus</emphasis>
|
||||
</taxonomicName>
|
||||
has been reported from
|
||||
<collectingCountry id="F32D86CB4001B22265F2186FFDEDF8FA" box="[426,613,1836,1863]" name="New Caledonia" pageId="12" pageNumber="13">New Caledonia</collectingCountry>
|
||||
, as in
|
||||
<bibRefCitation id="EFABBBAA4001B22266EC186EFC53F8FB" author="Chuang C. T. N. & Ng P. K. L." box="[692,987,1836,1863]" pageId="12" pageNumber="13" pagination="85 - 92" refId="ref12538" refString="Chuang C. T. N. & Ng P. K. L. 1994. The ecology and biology of Southeast Asian false spider crabs (Crustacea: Decapoda: Brachyura: Hymenosomatidae). In: Sasekumar A., Marshall N. & Macintosh D. J. (eds) Ecology and Conservation of Southeast Asian Marine and Freshwater Environments including Wetlands. Hydrobiologia 285: 85 - 92. https: // doi. org / 10.1007 / BF 00005656" type="journal article" year="1994">Chuang & Ng (1994: 87</bibRefCitation>
|
||||
, 90, table 1, under
|
||||
<taxonomicName id="4C3ABDD84001B222609C186FFAB1F8FA" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[1220,1337,1836,1863]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B222609C186FFAB1F8FA" box="[1220,1337,1836,1863]" italics="true" pageId="12" pageNumber="13">A. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
) and erroneously as in
|
||||
<bibRefCitation id="EFABBBAA4001B22265D51812FDE7F8D6" author="Guinot D." box="[397,623,1872,1898]" pageId="12" pageNumber="13" pagination="20 - 32" refId="ref13311" refString="Guinot D. 2011 a. Odiomarinae nov. subfam., a new subfamily for two primitive genera of the Hymenosomatidae MacLeay, 1838 (Crustacea, Decapoda, Brachyura). Zootaxa 2732: 20 - 32. https: // doi. org / 10.11646 / zootaxa. 2732.1.2" type="journal article" year="2011">Guinot (2011a: 23)</bibRefCitation>
|
||||
, are attributable to the fact that the species
|
||||
<taxonomicName id="4C3ABDD84001B22260331812FB36F8D6" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[1131,1214,1872,1898]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B22260331812FB36F8D6" box="[1131,1214,1872,1898]" italics="true" pageId="12" pageNumber="13">pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
was previously associated with the genus
|
||||
<taxonomicName id="4C3ABDD84001B22265AB1831FDEDF831" authorityName="Lucas" authorityYear="1980" box="[499,613,1907,1933]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B22265AB1831FDEDF831" box="[499,613,1907,1933]" italics="true" pageId="12" pageNumber="13">Amarinus</emphasis>
|
||||
</taxonomicName>
|
||||
(see
|
||||
<bibRefCitation id="EFABBBAA4001B22266FD1831FCA7F831" author="Lucas J. S." box="[677,815,1907,1933]" pageId="12" pageNumber="13" pagination="148 - 247" refId="ref14163" refString="Lucas J. S. 1980. Spider crabs of the family Hymenosomatidae (Crustacea: Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum 33 (4): 148 - 247. https: // doi. org / 10.3853 / j. 0067 - 1975.33.1980.199" type="journal article" year="1980">Lucas 1980</bibRefCitation>
|
||||
) until
|
||||
<bibRefCitation id="EFABBBAA4001B22267261831FB61F832" author="Ng P. K. L. & Richer de Forges B." box="[894,1257,1907,1934]" pageId="12" pageNumber="13" pagination="263 - 276" refId="ref14618" refString="Ng P. K. L. & Richer de Forges B. 1996. The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of New Caledonia, with descriptions of two new genera and two new species. Memoirs of the Queensland Museum 39 (2): 263 - 276. Available from https: // www. biodiversitylibrary. org / part / 50813 # / summary [accessed 27 May 2020]." type="journal article" year="1996">Ng & Richer de Forges (1996)</bibRefCitation>
|
||||
made it the
|
||||
<typeStatus id="548178F94001B22264E518D5FF78F80D" box="[189,240,1943,1969]" pageId="12" pageNumber="13">type</typeStatus>
|
||||
species of their new genus
|
||||
<taxonomicName id="4C3ABDD84001B222666418D4FD31F80C" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[572,697,1942,1968]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B222666418D4FD31F80C" box="[572,697,1942,1968]" italics="true" pageId="12" pageNumber="13">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
. So far, no species of
|
||||
<emphasis id="B94E1A494001B222679D18D5FBA4F80D" box="[965,1068,1943,1969]" italics="true" pageId="12" pageNumber="13">Amarinu</emphasis>
|
||||
s has actually been reported in
|
||||
<collectingCountry id="F32D86CB4001B222648418F8FE1FF86F" box="[220,407,1977,2004]" name="New Caledonia" pageId="12" pageNumber="13">New Caledonia</collectingCountry>
|
||||
. Suspecting that our new species might belong to the genus
|
||||
<taxonomicName id="4C3ABDD84001B222603918F8FB5BF868" authorityName="Lucas" authorityYear="1980" box="[1121,1235,1978,2004]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B222603918F8FB5BF868" box="[1121,1235,1978,2004]" italics="true" pageId="12" pageNumber="13">Amarinus</emphasis>
|
||||
</taxonomicName>
|
||||
, we therefore carefully compared
|
||||
<taxonomicName id="4C3ABDD84000B22365F21E49FD92FE99" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[426,538,267,293]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B94E1A494000B22365F21E49FD92FE99" box="[426,538,267,293]" italics="true" pageId="13" pageNumber="14">Richerius</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324000B223667B1E4EFD07FE9A" box="[547,655,268,294]" pageId="13" pageNumber="14" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
to
|
||||
<taxonomicName id="4C3ABDD84000B22366EF1E4EFCA1FE9A" authorityName="Lucas" authorityYear="1980" box="[695,809,268,294]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494000B22366EF1E4EFCA1FE9A" box="[695,809,268,294]" italics="true" pageId="13" pageNumber="14">Amarinus</emphasis>
|
||||
</taxonomicName>
|
||||
, and in particular to its
|
||||
<typeStatus id="548178F94000B22360101E4EFBF3FE9A" box="[1096,1147,268,294]" pageId="13" pageNumber="14">type</typeStatus>
|
||||
species,
|
||||
<taxonomicName id="4C3ABDD84000B22360B01E4EFAE7FE99" authorityName="Chilton" authorityYear="1882" box="[1256,1391,267,294]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lacustris">
|
||||
<emphasis id="B94E1A494000B22360B01E4EFAE7FE99" box="[1256,1391,267,294]" italics="true" pageId="13" pageNumber="14">A. lacustris</emphasis>
|
||||
</taxonomicName>
|
||||
, another freshwater hymenosomatid.
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4000B22364E51E32FAC3FDAB" blockId="13.[189,1399,368,536]" pageId="13" pageNumber="14">
|
||||
<taxonomicName id="4C3ABDD84000B22364E51E32FE17FE36" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[189,415,368,394]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494000B22364E51E32FE17FE36" box="[189,415,368,394]" italics="true" pageId="13" pageNumber="14">Richerius marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324000B22365F21E33FDECFE37" box="[426,612,369,395]" pageId="13" pageNumber="14" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
can be distinguished from
|
||||
<taxonomicName id="4C3ABDD84000B22367E91E32FB19FE36" baseAuthorityName="Chilton" baseAuthorityYear="1882" box="[945,1169,368,394]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lacustris">
|
||||
<emphasis id="B94E1A494000B22367E91E32FB19FE36" box="[945,1169,368,394]" italics="true" pageId="13" pageNumber="14">Amarinus lacustris</emphasis>
|
||||
</taxonomicName>
|
||||
by: the wide male pleon, with prominent marks corresponding to intercalated platelets (
|
||||
<figureCitation id="1301DADE4000B223678C1ED1FBA5FE11" box="[980,1069,403,429]" captionStart="Fig" captionStartId="7.[189,232,1846,1872]" captionTargetBox="[260,1308,282,1788]" captionTargetId="figure@7.[248,1339,265,1807]" captionTargetPageId="7" captionText="Fig. 2. Richerius marqueti gen. et sp. nov., holotype, ♂, 4.9 × 5.0 mm (MNHN-IU-2014-21500), same collection data as Fig. 1. A. Thoracic sternum. B. Mxp3. C. Sternopleonal cavity with G1s. D. G1, two views. Abbreviations: G1 = gonopod 1; G2 = gonopod 2; i = intercalated platelets; p = pleotelson; p.b. = press-button; s4 = expansion of sternite 4; 1–8 = thoracic sternites 1–8; 2/3–7/8 = thoracic sternal sutures 2/3–7/8. Scale bars: A = 2 mm; B–C = 1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3906760" httpUri="https://zenodo.org/record/3906760/files/figure.png" pageId="13" pageNumber="14">Fig. 2A</figureCitation>
|
||||
) (versus narrow and without intercalated platelets in
|
||||
<taxonomicName id="4C3ABDD84000B22365841EF5FDECFE6C" baseAuthorityName="Chilton" baseAuthorityYear="1882" box="[476,612,438,465]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lacustris">
|
||||
<emphasis id="B94E1A494000B22365841EF5FDECFE6C" box="[476,612,438,465]" italics="true" pageId="13" pageNumber="14">A. lacustris</emphasis>
|
||||
</taxonomicName>
|
||||
, see Melrose 1975: fig.
|
||||
<quantity id="4CC26BBE4000B22367D71EF5FC36FE6D" box="[911,958,439,465]" metricMagnitude="-2" metricUnit="kg" metricValue="4.2" pageId="13" pageNumber="14" unit="g" value="42.0">42g</quantity>
|
||||
;
|
||||
<bibRefCitation id="EFABBBAA4000B22367951EF5FBD4FE6D" author="Lucas J. S." box="[973,1116,439,465]" pageId="13" pageNumber="14" pagination="148 - 247" refId="ref14163" refString="Lucas J. S. 1980. Spider crabs of the family Hymenosomatidae (Crustacea: Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum 33 (4): 148 - 247. https: // doi. org / 10.3853 / j. 0067 - 1975.33.1980.199" type="journal article" year="1980">Lucas 1980</bibRefCitation>
|
||||
: fig. 7b); the G1 rather narrow and with one distal lobe without corneous process (
|
||||
<figureCitation id="1301DADE4000B22367351E98FC40FE48" box="[877,968,474,500]" captionStart="Fig" captionStartId="7.[189,232,1846,1872]" captionTargetBox="[260,1308,282,1788]" captionTargetId="figure@7.[248,1339,265,1807]" captionTargetPageId="7" captionText="Fig. 2. Richerius marqueti gen. et sp. nov., holotype, ♂, 4.9 × 5.0 mm (MNHN-IU-2014-21500), same collection data as Fig. 1. A. Thoracic sternum. B. Mxp3. C. Sternopleonal cavity with G1s. D. G1, two views. Abbreviations: G1 = gonopod 1; G2 = gonopod 2; i = intercalated platelets; p = pleotelson; p.b. = press-button; s4 = expansion of sternite 4; 1–8 = thoracic sternites 1–8; 2/3–7/8 = thoracic sternal sutures 2/3–7/8. Scale bars: A = 2 mm; B–C = 1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3906760" httpUri="https://zenodo.org/record/3906760/files/figure.png" pageId="13" pageNumber="14">Fig. 2D</figureCitation>
|
||||
) (versus stout and without lobes, see Melrose 1975: fig. 42h–i;
|
||||
<bibRefCitation id="EFABBBAA4000B22365B31EBFFDFDFDAB" author="Lucas J. S." box="[491,629,509,535]" pageId="13" pageNumber="14" pagination="148 - 247" refId="ref14163" refString="Lucas J. S. 1980. Spider crabs of the family Hymenosomatidae (Crustacea: Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum 33 (4): 148 - 247. https: // doi. org / 10.3853 / j. 0067 - 1975.33.1980.199" type="journal article" year="1980">Lucas 1980</bibRefCitation>
|
||||
: fig. 10e); and by the arrangement of antennules and antennae.
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4000B22364E51D7DFCA0FB13" blockId="13.[189,1399,574,1199]" pageId="13" pageNumber="14">
|
||||
In providing a key of the Southeast Asian hymenosomatids,
|
||||
<bibRefCitation id="EFABBBAA4000B22367FA1D7DFB57FDE5" author="Ng P. K. L. & Chuang C. T. N." box="[930,1247,574,601]" pageId="13" pageNumber="14" pagination="1 - 82" refId="ref14547" refString="Ng P. K. L. & Chuang C. T. N. 1996 The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of Southeast Asia, with notes on other species. Raffles Bulletin of Zoology Supplement 3: 1 - 82. Available from https: // decapoda. nhm. org / pdfs / 12225 / 12225. pdf [accessed 27 May 2020]." type="journal article" year="1996">Ng & Chuang (1996: 3–5</bibRefCitation>
|
||||
, 6–12) have shown the presence of several groups of species within
|
||||
<taxonomicName id="4C3ABDD84000B22367001D20FC42FDC0" authorityName="Lucas" authorityYear="1980" box="[856,970,610,636]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494000B22367001D20FC42FDC0" box="[856,970,610,636]" italics="true" pageId="13" pageNumber="14">Amarinus</emphasis>
|
||||
</taxonomicName>
|
||||
, suggesting that the genus could be paraphyletic. Today, another problem arises, especially regarding the antennular morphology. Melrose (1975: 84, 87, figs 41–42, as
|
||||
<taxonomicName id="4C3ABDD84000B22366761DEAFCBFFD7E" box="[558,823,680,706]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lacustris">
|
||||
<emphasis id="B94E1A494000B22366761DEAFCBFFD7E" box="[558,823,680,706]" italics="true" pageId="13" pageNumber="14">Halicarcinus lacustris</emphasis>
|
||||
</taxonomicName>
|
||||
), who has thoroughly studied the
|
||||
<typeStatus id="548178F94000B22360BB1DEAFA9EFD7E" box="[1251,1302,680,706]" pageId="13" pageNumber="14">type</typeStatus>
|
||||
species
|
||||
<taxonomicName id="4C3ABDD84000B22364E51D89FECDFD59" box="[189,325,715,741]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lacustris">
|
||||
<emphasis id="B94E1A494000B22364E51D89FECDFD59" box="[189,325,715,741]" italics="true" pageId="13" pageNumber="14">A. lacustris</emphasis>
|
||||
</taxonomicName>
|
||||
, confined to lakes and non-tidal rivers of
|
||||
<collectingCountry id="F32D86CB4000B22367191D8EFC68FD59" box="[833,992,715,742]" name="New Zealand" pageId="13" pageNumber="14">New Zealand</collectingCountry>
|
||||
and southeastern
|
||||
<collectingCountry id="F32D86CB4000B22360E11D89FAA0FD59" box="[1209,1320,715,741]" name="Australia" pageId="13" pageNumber="14">Australia</collectingCountry>
|
||||
, states that the antennules are “small, not visible dorsally when folded”. Her figure 41d actually seems to show a folded antennule, only with the broad basal article and the second cylindrical article, without the short, supposedly folded flagellum being seen [the frontal view of fig. 41e in Melrose (1975) shows a complete, unfolded antennule]. Our examination of
|
||||
<taxonomicName id="4C3ABDD84000B223677B1C1AFB8AFCCE" baseAuthorityName="Chilton" baseAuthorityYear="1882" box="[803,1026,856,882]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lacustris">
|
||||
<emphasis id="B94E1A494000B223677B1C1AFB8AFCCE" box="[803,1026,856,882]" italics="true" pageId="13" pageNumber="14">Amarinus lacustris</emphasis>
|
||||
</taxonomicName>
|
||||
, on the other hand, reveals that both antennules and antennae are inserted very closely together at their bases and remain parallel, with both flagella visible dorsally, as represented by
|
||||
<bibRefCitation id="EFABBBAA4000B22366871CDDFCFCFC04" author="Lucas J. S." box="[735,884,926,953]" pageId="13" pageNumber="14" pagination="148 - 247" refId="ref14163" refString="Lucas J. S. 1980. Spider crabs of the family Hymenosomatidae (Crustacea: Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum 33 (4): 148 - 247. https: // doi. org / 10.3853 / j. 0067 - 1975.33.1980.199" type="journal article" year="1980">Lucas (1980</bibRefCitation>
|
||||
: fig. 1d) for a generalised hymenosomatid or by Melrose (1975: fig. 43c) for
|
||||
<taxonomicName id="4C3ABDD84000B223667E1C83FC2DFC60" authority="Melrose, 1975" authorityName="Melrose" authorityYear="1975" box="[550,933,961,988]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tongi">
|
||||
<emphasis id="B94E1A494000B223667E1C83FD7CFC67" box="[550,756,961,987]" italics="true" pageId="13" pageNumber="14">Halicarinus tongi</emphasis>
|
||||
Melrose, 1975
|
||||
</taxonomicName>
|
||||
. In fact, if it is possible for the antennule of
|
||||
<taxonomicName id="4C3ABDD84000B22364861CA7FEEDFC43" baseAuthorityName="Chilton" baseAuthorityYear="1882" box="[222,357,997,1023]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lacustris">
|
||||
<emphasis id="B94E1A494000B22364861CA7FEEDFC43" box="[222,357,997,1023]" italics="true" pageId="13" pageNumber="14">A. lacustris</emphasis>
|
||||
</taxonomicName>
|
||||
to fold itself down [but not obliquely along the rostrum as in
|
||||
<taxonomicName id="4C3ABDD84000B22360141CA7FB41FC43" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[1100,1225,997,1023]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494000B22360141CA7FB41FC43" box="[1100,1225,997,1023]" italics="true" pageId="13" pageNumber="14">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1301DADE4000B22360831CA7FAB2FC43" box="[1243,1338,997,1023]" captionStart="Fig" captionStartId="16.[189,232,1919,1945]" captionTargetBox="[402,1193,1524,1866]" captionTargetId="figure@16.[385,1203,1509,1878]" captionTargetPageId="16" captionText="Fig. 7. Odiomaris pilosus A. Milne-Edwards, 1873, syntype, ♂, New Caledonia, freshwater, B. Balansa leg. (RMNH D 32). A. Ventral anterior part. B. G1 (after Holthuis 1968: fig. 3a–b). Abbreviations: a1 = antennule, with folded second article; a2 = antenna; e = epistome: p.k. = postorbital knobs; r = rostrum." figureDoi="http://doi.org/10.5281/zenodo.3906770" httpUri="https://zenodo.org/record/3906770/files/figure.png" pageId="13" pageNumber="14">Fig. 7A</figureCitation>
|
||||
), see A. Milne-Edwards 1873: pl. 18, fig. 6a], this does not correspond to the resting position. In the seven specimens of
|
||||
<taxonomicName id="4C3ABDD84000B223653C1B69FE65FBF9" baseAuthorityName="Chilton" baseAuthorityYear="1882" box="[356,493,1067,1093]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lacustris">
|
||||
<emphasis id="B94E1A494000B223653C1B69FE65FBF9" box="[356,493,1067,1093]" italics="true" pageId="13" pageNumber="14">A. lacustris</emphasis>
|
||||
</taxonomicName>
|
||||
examined, the antennule and antenna are very closely inserted and remain parallel, both being stretched forward; the unfolded flagellum is visible dorsally in all individuals. In any event, the disposition of the antennule and antenna differs from that
|
||||
<taxonomicName id="4C3ABDD84000B223605A1B33FBF7FB37" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[1026,1151,1137,1163]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494000B223605A1B33FBF7FB37" box="[1026,1151,1137,1163]" italics="true" pageId="13" pageNumber="14">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1301DADE4000B22360D71B33FB65FB30" box="[1167,1261,1137,1164]" captionStart="Fig" captionStartId="16.[189,232,1919,1945]" captionTargetBox="[402,1193,1524,1866]" captionTargetId="figure@16.[385,1203,1509,1878]" captionTargetPageId="16" captionText="Fig. 7. Odiomaris pilosus A. Milne-Edwards, 1873, syntype, ♂, New Caledonia, freshwater, B. Balansa leg. (RMNH D 32). A. Ventral anterior part. B. G1 (after Holthuis 1968: fig. 3a–b). Abbreviations: a1 = antennule, with folded second article; a2 = antenna; e = epistome: p.k. = postorbital knobs; r = rostrum." figureDoi="http://doi.org/10.5281/zenodo.3906770" httpUri="https://zenodo.org/record/3906770/files/figure.png" pageId="13" pageNumber="14">Fig. 7A</figureCitation>
|
||||
:
|
||||
<taxonomicName id="4C3ABDD84000B22360AF1B30FAE5FB37" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[1271,1389,1137,1164]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494000B22360AF1B30FAE5FB37" box="[1271,1389,1137,1164]" italics="true" pageId="13" pageNumber="14">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
) (
|
||||
<bibRefCitation id="EFABBBAA4000B223649D1BD7FD96FB13" author="Ng P. K. L. & Richer de Forges B." box="[197,542,1173,1199]" pageId="13" pageNumber="14" pagination="263 - 276" refId="ref14618" refString="Ng P. K. L. & Richer de Forges B. 1996. The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of New Caledonia, with descriptions of two new genera and two new species. Memoirs of the Queensland Museum 39 (2): 263 - 276. Available from https: // www. biodiversitylibrary. org / part / 50813 # / summary [accessed 27 May 2020]." type="journal article" year="1996">Ng & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 6c–d:
|
||||
<taxonomicName id="4C3ABDD84000B22366F81BD7FC9EFB13" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[672,790,1173,1199]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494000B22366F81BD7FC9EFB13" box="[672,790,1173,1199]" italics="true" pageId="13" pageNumber="14">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4000B22364E51B94FECAFA5B" blockId="13.[189,1399,1238,1511]" pageId="13" pageNumber="14">
|
||||
A picture of the cephalic region in ventral view of
|
||||
<taxonomicName id="4C3ABDD84000B223677F1B95FB9BFB4C" authorityName="Holthuis" authorityYear="1968" box="[807,1043,1238,1265]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="angelicus">
|
||||
<emphasis id="B94E1A494000B223677F1B95FB9BFB4C" box="[807,1043,1238,1265]" italics="true" pageId="13" pageNumber="14">Amarinus angelicus</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<bibRefCitation id="EFABBBAA4000B223607E1B94FA99FB4C" author="Holthuis L. B." box="[1062,1297,1238,1264]" pageId="13" pageNumber="14" pagination="109 - 121" refId="ref13478" refString="Holthuis L. B. 1968. On Hymenosomatidae (Crustacea Decapoda Brachyura) from fresh water, with the description of a new species. Beaufortia 15 (195): 109 - 121. Available from https: // www. repository. naturalis. nl / record / 504776 [accessed 27 May 2020]." type="journal article" year="1968">Holthuis 1968: 114</bibRefCitation>
|
||||
, fig. 2b) shows an antennule “entirely hidden below the rostrum”, obliquely folded, which does not seem to correspond to the condition of
|
||||
<taxonomicName id="4C3ABDD84000B223666C1A5FFD34FA8A" baseAuthorityName="Chilton" baseAuthorityYear="1882" box="[564,700,1308,1335]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lacustris">
|
||||
<emphasis id="B94E1A494000B223666C1A5FFD34FA8A" box="[564,700,1308,1335]" italics="true" pageId="13" pageNumber="14">A. lacustris</emphasis>
|
||||
</taxonomicName>
|
||||
. This character deserves to be reviewed for all species of
|
||||
<taxonomicName id="4C3ABDD84000B22364E51A02FEA7FAE6" authorityName="Lucas" authorityYear="1980" box="[189,303,1344,1370]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494000B22364E51A02FEA7FAE6" box="[189,303,1344,1370]" italics="true" pageId="13" pageNumber="14">Amarinus</emphasis>
|
||||
</taxonomicName>
|
||||
, and the allocation of
|
||||
<taxonomicName id="4C3ABDD84000B223666D1A02FD2FFAE6" authorityName="Lucas" authorityYear="1980" box="[565,679,1344,1370]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494000B223666D1A02FD2FFAE6" box="[565,679,1344,1370]" italics="true" pageId="13" pageNumber="14">Amarinus</emphasis>
|
||||
</taxonomicName>
|
||||
(including the
|
||||
<typeStatus id="548178F94000B22367001A02FC03FAE6" box="[856,907,1344,1370]" pageId="13" pageNumber="14">type</typeStatus>
|
||||
species and other included species) to the
|
||||
<taxonomicName id="4C3ABDD84000B22364E51A21FEDEFAC1" authorityName="Guinot" authorityYear="2011" box="[189,342,1379,1405]" class="Malacostraca" family="Hymenosomatidae" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subFamily" subFamily="Odiomarinae">Odiomarinae</taxonomicName>
|
||||
needs further investigation. In the new species described here,
|
||||
<taxonomicName id="4C3ABDD84000B22360001A21FAB3FAC1" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[1112,1339,1379,1405]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494000B22360001A21FAB3FAC1" box="[1112,1339,1379,1405]" italics="true" pageId="13" pageNumber="14">Richerius marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324000B223611D1A21FEB1FA1D" pageId="13" pageNumber="14" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
(
|
||||
<figureCitation id="1301DADE4000B22365131AC4FE2FFA1C" box="[331,423,1414,1440]" captionStart="Fig" captionStartId="6.[189,232,1782,1808]" captionTargetBox="[326,1260,292,1710]" captionTargetId="figure@6.[292,1296,265,1741]" captionTargetPageId="6" captionText="Fig. 1. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Carapace, dorsal view. B. Cephalothorax, ventral view, anterior part, with unfolded antennules, the arrow shows antennule with folded second article. Abbreviations: a1 = antennule; a2 = antenna; e = epistome; p.k. = postorbital knob; r = rostrum; u.a. = antennal urinary article; v.r. = ventral expansion of rostrum; 2/3 = thoracic sternal suture 2/3. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3906758" httpUri="https://zenodo.org/record/3906758/files/figure.png" pageId="13" pageNumber="14">Fig. 1B</figureCitation>
|
||||
), the antennule and antenna are separated at least at their insertion site, and only the antennal flagellum is visible dorsally; thus it shares an arrangement substantially similar to that of
|
||||
<taxonomicName id="4C3ABDD84000B22364E51A8EFEB2FA5A" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[189,314,1484,1510]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494000B22364E51A8EFEB2FA5A" box="[189,314,1484,1510]" italics="true" pageId="13" pageNumber="14">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4000B22364E5194CFB42F8FE" blockId="13.[189,1399,1550,1858]" pageId="13" pageNumber="14">
|
||||
It was the study of spermatozoa carried out first in the two species of
|
||||
<taxonomicName id="4C3ABDD84000B223606A194CFB27F994" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[1074,1199,1550,1576]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494000B223606A194CFB27F994" box="[1074,1199,1550,1576]" italics="true" pageId="13" pageNumber="14">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C3ABDD84000B223609C194CFAC8F994" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[1220,1344,1550,1576]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494000B223609C194CFAC8F994" box="[1220,1344,1550,1576]" italics="true" pageId="13" pageNumber="14">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C3ABDD84000B22364E51970FEC4F9F7" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[189,332,1586,1612]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494000B22364E51970FEC4F9F7" box="[189,332,1586,1612]" italics="true" pageId="13" pageNumber="14">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
, by
|
||||
<bibRefCitation id="EFABBBAA4000B22365261973FD53F9F7" author="Richer de Forges B. & Jamieson B. G. M. & Guinot D. & Tudge C. C." box="[382,731,1585,1612]" pageId="13" pageNumber="14" pagination="233 - 242" refId="ref15566" refString="Richer de Forges B., Jamieson B. G. M., Guinot D. & Tudge C. C. 1997. Ultrastructure of the spermatozoa of Hymenosomatidae (Crustacea: Brachyura) and the relationships of the family. Marine Biology 130: 233 - 242. https: // doi. org / 10.1007 / s 002270050243" type="journal article" year="1997">
|
||||
Richer de Forges
|
||||
<emphasis id="B94E1A494000B22366141970FD0EF9F7" box="[588,646,1585,1611]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
|
||||
(1997)
|
||||
</bibRefCitation>
|
||||
and later in
|
||||
<taxonomicName id="4C3ABDD84000B22367331973FB94F9F7" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[875,1052,1585,1611]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="vesca">
|
||||
<emphasis id="B94E1A494000B22367331973FB94F9F7" box="[875,1052,1585,1611]" italics="true" pageId="13" pageNumber="14">Elamena vesca</emphasis>
|
||||
</taxonomicName>
|
||||
by
|
||||
<bibRefCitation id="EFABBBAA4000B223601F1973FAFEF9F7" author="Jamieson B. G. M. & Tudge C. C." box="[1095,1398,1585,1611]" pageId="13" pageNumber="14" pagination="1 - 95" refId="ref13652" refString="Jamieson B. G. M. & Tudge C. C. 2000. Crustacea-Decapoda. In: Adiyodi K. G. & Adiyodi R. G. (ser. eds), Jamieson B. G. M. (vol. ed.) Reproductive Biology of Invertebrates. Volume IX, Part C. Progress in Male Gamete Ultrastructure and Phylogeny. John Wiley and Sons. Chichester: 1 - 95." type="book chapter" year="2000">Jamieson & Tudge (2000)</bibRefCitation>
|
||||
that showed the very particular nature of the hymenosomatid sperm within the
|
||||
<taxonomicName id="4C3ABDD84000B22360041916FB52F9D2" authorityName="Latreille" authorityYear="1802" box="[1116,1242,1620,1646]" infraOrder="Brachyura" pageId="13" pageNumber="3" rank="infraOrder">Brachyura</taxonomicName>
|
||||
. Indeed, they are distinguished by at least nine major characteristics from those of all the other
|
||||
<taxonomicName id="4C3ABDD84000B223603C1935FB56F92D" authorityName="Latreille" authorityYear="1802" box="[1124,1246,1655,1681]" infraOrder="Brachyura" pageId="13" pageNumber="3" rank="infraOrder">Brachyura</taxonomicName>
|
||||
taxa studied, particularly the groups with which they have been associated, the Majoidea Samouelle, 1819 and the Thoracotremata Guinot, 1977. The ‘hymenosomatid-type of spermatozoon’ is unique within
|
||||
<taxonomicName id="4C3ABDD84000B22360A419FCFAFEF964" authorityName="Latreille" authorityYear="1802" box="[1276,1398,1726,1752]" infraOrder="Brachyura" pageId="13" pageNumber="3" rank="infraOrder">Brachyura</taxonomicName>
|
||||
(
|
||||
<bibRefCitation id="EFABBBAA4000B223649D19A3FE1BF947" author="Tudge C. & Scheltinga D. M. & Jamieson B. G. M. & Guinot D. & Richer de Forges B." box="[197,403,1761,1787]" pageId="13" pageNumber="14" pagination="1 - 20" refId="ref15940" refString="Tudge C. C, Scheltinga D. M., Jamieson B. G. M., Guinot D. & Richer de Forges B. 2014. Comparative ultrastructure of the spermatozoa of the Majoidea (Crustacea, Decapoda, Brachyura) with new data on six species in five genera. Acta Zoologica 95: 1 - 20. https: // doi. org / 10.1111 / azo. 12005" type="journal article" year="2014">
|
||||
Tudge
|
||||
<emphasis id="B94E1A494000B223654E19A0FEC6F947" box="[278,334,1761,1787]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
|
||||
2014
|
||||
</bibRefCitation>
|
||||
). The highly developed projection of the acrosome from the nucleus in hymenosomatid spermatozoon recalls the totally emergent acrosome of Podotremata Guinot, 1977 and may represent the plesiomorphic condition in the Eubrachyura Saint Laurent, 1980 (
|
||||
<bibRefCitation id="EFABBBAA4000B22367981865FBECF8FE" author="Guinot D." box="[960,1124,1831,1858]" pageId="13" pageNumber="14" pagination="20 - 32" refId="ref13311" refString="Guinot D. 2011 a. Odiomarinae nov. subfam., a new subfamily for two primitive genera of the Hymenosomatidae MacLeay, 1838 (Crustacea, Decapoda, Brachyura). Zootaxa 2732: 20 - 32. https: // doi. org / 10.11646 / zootaxa. 2732.1.2" type="journal article" year="2011">Guinot 2011a</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EFABBBAA4000B22360281865FB35F8FE" author="Guinot D." box="[1136,1213,1831,1858]" pageId="13" pageNumber="14" pagination="40 - 52" refId="ref13363" refString="Guinot D. 2011 b. The position of the Hymenosomatidae MacLeay, 1838, within the Brachyura (Crustacea, Decapoda). Zootaxa 2890: 40 - 52. https: // doi. org / 10.11646 / zootaxa. 2890.1.4" type="journal article" year="2011">2011b</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,54 +1,54 @@
|
|||
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<mods:identifier id="528FF2A226EABCA8EB374A0F0ABA4211" type="ISSN">2118-9773</mods:identifier>
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<mods:identifier id="01E7FBD008A49BA5764D07833E69C8A0" type="Zenodo-Dep">3906756</mods:identifier>
|
||||
<mods:identifier id="AABFBAFD6C46312B39409BE032BC6986" type="ZooBank">9EF19154-D2FE-4009-985C-1EB62CC9ACB0</mods:identifier>
|
||||
<mods:classification id="A119A29E6DB7962305FA1271B9A56781">journal article</mods:classification>
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||||
<mods:identifier id="0EDF838891B9534981C756EA852B447A" type="DOI">10.5852/ejt.2020.671</mods:identifier>
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|
||||
</mods:mods>
|
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<treatment id="0393774D400EB22A66111BB8FBFFFDFC" ID-DOI="http://doi.org/10.5281/zenodo.7433864" ID-GBIF-Taxon="164723367" ID-Zenodo-Dep="7433864" LSID="urn:lsid:plazi:treatment:0393774D400EB22A66111BB8FBFFFDFC" httpUri="http://treatment.plazi.org/id/0393774D400EB22A66111BB8FBFFFDFC" lastPageId="4" lastPageNumber="4" pageId="3" pageNumber="4" scope_family="Hymenosomatidae" scope_infraOrder="Brachyura" scope_order="Decapoda" scope_subFamily="Odiomarinae" scope_superFamily="Hymenosomatoidea">
|
||||
<treatment id="0393774D400EB22A66111BB8FBFFFDFC" ID-DOI="http://doi.org/10.5281/zenodo.7433864" ID-GBIF-Taxon="164723367" ID-Zenodo-Dep="7433864" LSID="urn:lsid:plazi:treatment:0393774D400EB22A66111BB8FBFFFDFC" httpUri="http://treatment.plazi.org/id/0393774D400EB22A66111BB8FBFFFDFC" lastPageId="4" lastPageNumber="14" pageId="3" pageNumber="4" scope_family="Hymenosomatidae" scope_infraOrder="Brachyura" scope_order="Decapoda" scope_subFamily="Odiomarinae" scope_superFamily="Hymenosomatoidea">
|
||||
<subSubSection id="C32095D0400EB22D66111BB8FC63FAA9" box="[585,1003,1274,1301]" pageId="3" pageNumber="4" type="nomenclature">
|
||||
<paragraph id="8B85C65B400EB22D66111BB8FC63FAA9" blockId="3.[370,1217,1274,1372]" box="[585,1003,1274,1301]" pageId="3" pageNumber="4">
|
||||
<heading id="D0CD7137400EB22D66111BB8FC63FAA9" box="[585,1003,1274,1301]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="4" reason="2">
|
||||
|
|
@ -258,5 +258,904 @@ specs; same collection data as for preceding; DNA voucher: CA2187; GenBak:
|
|||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C32095D04008B22564E51B7DFBCAFE60" lastPageId="11" lastPageNumber="12" pageId="5" pageNumber="6" type="description">
|
||||
<paragraph id="8B85C65B4008B22B64E51B7DFEC6FBE5" blockId="5.[189,334,1087,1113]" box="[189,334,1087,1113]" pageId="5" pageNumber="6">
|
||||
<heading id="D0CD71374008B22B64E51B7DFEC6FBE5" bold="true" box="[189,334,1087,1113]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
|
||||
<emphasis id="B94E1A494008B22B64E51B7DFEC6FBE5" bold="true" box="[189,334,1087,1113]" pageId="5" pageNumber="6">Description</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4008B22B64E51B2CFE5BFA8B" blockId="5.[189,1399,1133,1335]" pageId="5" pageNumber="6">
|
||||
Carapace (
|
||||
<figureCitation id="1301DADE4008B22B65661B2FFE2CFB34" box="[318,420,1133,1160]" captionStart="Fig" captionStartId="6.[189,232,1782,1808]" captionTargetBox="[326,1260,292,1710]" captionTargetId="figure@6.[292,1296,265,1741]" captionTargetPageId="6" captionText="Fig. 1. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Carapace, dorsal view. B. Cephalothorax, ventral view, anterior part, with unfolded antennules, the arrow shows antennule with folded second article. Abbreviations: a1 = antennule; a2 = antenna; e = epistome; p.k. = postorbital knob; r = rostrum; u.a. = antennal urinary article; v.r. = ventral expansion of rostrum; 2/3 = thoracic sternal suture 2/3. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3906758" httpUri="https://zenodo.org/record/3906758/files/figure.png" pageId="5" pageNumber="6">Figs 1A</figureCitation>
|
||||
,
|
||||
<figureCitation id="1301DADE4008B22B65EC1B2FFE5FFB34" box="[436,471,1133,1160]" captionStart="Fig" captionStartId="10.[189,232,1855,1881]" captionTargetBox="[189,1398,901,1815]" captionTargetId="figure@10.[189,1398,901,1816]" captionTargetPageId="10" captionText="Fig. 5. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Type locality, stream where several specimens of the species were collected. B–D. Views of the holotype in vivo: dorsal (B), ventral (C) and frontal (D)." figureDoi="http://doi.org/10.5281/zenodo.3906766" httpUri="https://zenodo.org/record/3906766/files/figure.png" pageId="5" pageNumber="6">5B</figureCitation>
|
||||
) approximately circular to oval, slightly broader than long; dorsal carapace surface slightly concave, not strongly outlined by grooves, only with well defined gastrocardiac and thoracic grooves, approaching but not reaching antero- and posterolateral margins, respectively; no row of setae along lateral margins, except for some regularly mid-spaced setae in males; carapace angles not well marked. Anterolateral margin entire, without crenulations, lobes or teeth. Branchiostegite only weakly visible dorsally.
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4008B22B64E51A1DFBC2F90A" blockId="5.[189,1399,1375,1718]" pageId="5" pageNumber="6">
|
||||
Eyes visible dorsally. Antennules (
|
||||
<figureCitation id="1301DADE4008B22B66041A1DFD30FAC5" box="[604,696,1375,1401]" captionStart="Fig" captionStartId="6.[189,232,1782,1808]" captionTargetBox="[326,1260,292,1710]" captionTargetId="figure@6.[292,1296,265,1741]" captionTargetPageId="6" captionText="Fig. 1. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Carapace, dorsal view. B. Cephalothorax, ventral view, anterior part, with unfolded antennules, the arrow shows antennule with folded second article. Abbreviations: a1 = antennule; a2 = antenna; e = epistome; p.k. = postorbital knob; r = rostrum; u.a. = antennal urinary article; v.r. = ventral expansion of rostrum; 2/3 = thoracic sternal suture 2/3. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3906758" httpUri="https://zenodo.org/record/3906758/files/figure.png" pageId="5" pageNumber="6">Fig. 1B</figureCitation>
|
||||
) obliquely folded along hollowed ventral parts of rostrum and entirely hidden dorsally. Antennae well separated from antennules, at least at their bases; urinary article at level of epistome. Rostrum (
|
||||
<figureCitation id="1301DADE4008B22B662C1AE7FD36FA03" box="[628,702,1445,1471]" captionStart="Fig" captionStartId="6.[189,232,1782,1808]" captionTargetBox="[326,1260,292,1710]" captionTargetId="figure@6.[292,1296,265,1741]" captionTargetPageId="6" captionText="Fig. 1. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Carapace, dorsal view. B. Cephalothorax, ventral view, anterior part, with unfolded antennules, the arrow shows antennule with folded second article. Abbreviations: a1 = antennule; a2 = antenna; e = epistome; p.k. = postorbital knob; r = rostrum; u.a. = antennal urinary article; v.r. = ventral expansion of rostrum; 2/3 = thoracic sternal suture 2/3. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3906758" httpUri="https://zenodo.org/record/3906758/files/figure.png" pageId="5" pageNumber="6">Figs 1</figureCitation>
|
||||
,
|
||||
<figureCitation id="1301DADE4008B22B66941AE7FD55FA03" box="[716,733,1445,1471]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[208,1380,266,1860]" captionTargetId="figure@9.[208,1380,266,1862]" captionTargetPageId="9" captionText="Fig. 4. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Dorsal view. B. Ventral view. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3906764" httpUri="https://zenodo.org/record/3906764/files/figure.png" pageId="5" pageNumber="6">4</figureCitation>
|
||||
,
|
||||
<figureCitation id="1301DADE4008B22B66B11AE7FD7EFA7C" box="[745,758,1445,1472]" captionStart="Fig" captionStartId="10.[189,232,1855,1881]" captionTargetBox="[189,1398,901,1815]" captionTargetId="figure@10.[189,1398,901,1816]" captionTargetPageId="10" captionText="Fig. 5. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Type locality, stream where several specimens of the species were collected. B–D. Views of the holotype in vivo: dorsal (B), ventral (C) and frontal (D)." figureDoi="http://doi.org/10.5281/zenodo.3906766" httpUri="https://zenodo.org/record/3906766/files/figure.png" pageId="5" pageNumber="6">5</figureCitation>
|
||||
B–D) broadly rounded, spade-shaped, barely deflexed; dorsal surface spatulate with marked depression; carapace rim as a small ridge continuous across behind rostrum. Proepistome represented by triangular ventral expansion of rostrum (
|
||||
<figureCitation id="1301DADE4008B22B60D11AAEFB62F9BA" box="[1161,1258,1516,1542]" captionStart="Fig" captionStartId="6.[189,232,1782,1808]" captionTargetBox="[326,1260,292,1710]" captionTargetId="figure@6.[292,1296,265,1741]" captionTargetPageId="6" captionText="Fig. 1. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Carapace, dorsal view. B. Cephalothorax, ventral view, anterior part, with unfolded antennules, the arrow shows antennule with folded second article. Abbreviations: a1 = antennule; a2 = antenna; e = epistome; p.k. = postorbital knob; r = rostrum; u.a. = antennal urinary article; v.r. = ventral expansion of rostrum; 2/3 = thoracic sternal suture 2/3. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3906758" httpUri="https://zenodo.org/record/3906758/files/figure.png" pageId="5" pageNumber="6">Fig. 1B</figureCitation>
|
||||
). Epistome moderately developed; anterior margin undulated. Lower orbital margin with one conspicuous knob, not visible dorsally. Pterygostomial regions with setae, distinctly separated from subhepatic area by marked ridge. Mxp3 moderately gaping, broad; merus and ischium broad, short; midlength of merus slightly longer than that of ischium (
|
||||
<figureCitation id="1301DADE4008B22B6654193AFDE3F92F" box="[524,619,1656,1683]" captionStart="Fig" captionStartId="6.[189,232,1782,1808]" captionTargetBox="[326,1260,292,1710]" captionTargetId="figure@6.[292,1296,265,1741]" captionTargetPageId="6" captionText="Fig. 1. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Carapace, dorsal view. B. Cephalothorax, ventral view, anterior part, with unfolded antennules, the arrow shows antennule with folded second article. Abbreviations: a1 = antennule; a2 = antenna; e = epistome; p.k. = postorbital knob; r = rostrum; u.a. = antennal urinary article; v.r. = ventral expansion of rostrum; 2/3 = thoracic sternal suture 2/3. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3906758" httpUri="https://zenodo.org/record/3906758/files/figure.png" pageId="5" pageNumber="6">Figs 1B</figureCitation>
|
||||
,
|
||||
<figureCitation id="1301DADE4008B22B6621193BFD16F92F" box="[633,670,1657,1683]" captionStart="Fig" captionStartId="7.[189,232,1846,1872]" captionTargetBox="[260,1308,282,1788]" captionTargetId="figure@7.[248,1339,265,1807]" captionTargetPageId="7" captionText="Fig. 2. Richerius marqueti gen. et sp. nov., holotype, ♂, 4.9 × 5.0 mm (MNHN-IU-2014-21500), same collection data as Fig. 1. A. Thoracic sternum. B. Mxp3. C. Sternopleonal cavity with G1s. D. G1, two views. Abbreviations: G1 = gonopod 1; G2 = gonopod 2; i = intercalated platelets; p = pleotelson; p.b. = press-button; s4 = expansion of sternite 4; 1–8 = thoracic sternites 1–8; 2/3–7/8 = thoracic sternal sutures 2/3–7/8. Scale bars: A = 2 mm; B–C = 1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3906760" httpUri="https://zenodo.org/record/3906760/files/figure.png" pageId="5" pageNumber="6">2B</figureCitation>
|
||||
). Sternum/pterygostome junction substantially developed due to extension of sternite 4. Milne-Edwards openings separated from chelipeds.
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4008B22B64E5199FFC02F876" blockId="5.[189,1399,1757,1995]" pageId="5" pageNumber="6">
|
||||
Male chelipeds much stouter than walking legs, particularly in large males (
|
||||
<figureCitation id="1301DADE4008B22B601D199FFB07F94B" box="[1093,1167,1757,1783]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[208,1380,266,1860]" captionTargetId="figure@9.[208,1380,266,1862]" captionTargetPageId="9" captionText="Fig. 4. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Dorsal view. B. Ventral view. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3906764" httpUri="https://zenodo.org/record/3906764/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
|
||||
); merus and carpus with stiff, regularly spaced setae; propodus very inflated, covered with long soft setae partially extending on fingers; fingers with finely denticulate cutting edges, not gaping. Female chelipeds narrow, propodus moderately inflated, devoid of long setae; fingers proportionally rather long, with with finely denticulate cutting edges completely joined. Walking legs proportionally rather long, with margins bearing stiff, regularly spaced, scattered setae; dactyli slender but not distinctly longer than respective propodi, smoothly curved, setose, without teeth, ending in pointed tip.
|
||||
</paragraph>
|
||||
<caption id="DF4596D3400BB22864E519B4FC0AF858" ID-DOI="http://doi.org/10.5281/zenodo.3906758" ID-Zenodo-Dep="3906758" httpUri="https://zenodo.org/record/3906758/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1782,1808]" targetBox="[326,1260,292,1710]" targetPageId="6">
|
||||
<paragraph id="8B85C65B400BB22864E519B4FC0AF858" blockId="6.[189,1399,1780,2020]" pageId="6" pageNumber="7">
|
||||
<emphasis id="B94E1A49400BB22864E519B4FE83F8AC" bold="true" box="[189,267,1782,1808]" pageId="6" pageNumber="7">Fig. 1.</emphasis>
|
||||
<taxonomicName id="4C3ABDD8400BB228654A19B4FE7BF8AC" ID-CoL="84H9D" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[274,499,1782,1808]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A49400BB228654A19B4FE7BF8AC" box="[274,499,1782,1808]" italics="true" pageId="6" pageNumber="7">Richerius marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA732400BB22865A219B4FD25F8AC" box="[506,685,1782,1808]" pageId="6" pageNumber="7" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
, holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500).
|
||||
<emphasis id="B94E1A49400BB22864E5181DFF5AF8C5" bold="true" box="[189,210,1887,1913]" pageId="6" pageNumber="7">A</emphasis>
|
||||
. Carapace, dorsal view.
|
||||
<emphasis id="B94E1A49400BB22865AE1822FD82F8C6" bold="true" box="[502,522,1888,1914]" pageId="6" pageNumber="7">B</emphasis>
|
||||
. Cephalothorax, ventral view, anterior part, with unfolded antennules, the arrow shows antennule with folded second article. Abbreviations: a1 = antennule; a2 = antenna; e = epistome; p.k. = postorbital knob; r = rostrum; u.a. = antennal urinary article; v.r. = ventral expansion of rostrum; 2/3 = thoracic sternal suture 2/3. Scale bar = 2 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF4596D3400AB22964E51874FCE7F862" ID-DOI="http://doi.org/10.5281/zenodo.3906760" ID-Zenodo-Dep="3906760" httpUri="https://zenodo.org/record/3906760/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1846,1872]" targetBox="[260,1308,282,1788]" targetPageId="7">
|
||||
<paragraph id="8B85C65B400AB22964E51874FCE7F862" blockId="7.[189,1399,1846,2014]" pageId="7" pageNumber="8">
|
||||
<emphasis id="B94E1A49400AB22964E51874FE83F8EC" bold="true" box="[189,267,1846,1872]" pageId="7" pageNumber="8">Fig. 2.</emphasis>
|
||||
<taxonomicName id="4C3ABDD8400AB229654B1874FE7CF8EC" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[275,500,1846,1872]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A49400AB229654B1874FE7CF8EC" box="[275,500,1846,1872]" italics="true" pageId="7" pageNumber="8">Richerius marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA732400AB22965A41875FD38F8ED" box="[508,688,1847,1873]" pageId="7" pageNumber="8" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
, holotype, ♂, 4.9 × 5.0 mm (MNHN-IU-2014-21500), same collection data as Fig. 1.
|
||||
<emphasis id="B94E1A49400AB22965BC181BFE71F8CF" bold="true" box="[484,505,1881,1907]" pageId="7" pageNumber="8">A</emphasis>
|
||||
. Thoracic sternum.
|
||||
<emphasis id="B94E1A49400AB22966BD1818FD71F8C8" bold="true" box="[741,761,1882,1908]" pageId="7" pageNumber="8">B</emphasis>
|
||||
. Mxp3.
|
||||
<emphasis id="B94E1A49400AB2296704181BFCF9F8CF" bold="true" box="[860,881,1881,1907]" pageId="7" pageNumber="8">C</emphasis>
|
||||
. Sternopleonal cavity with G1s.
|
||||
<emphasis id="B94E1A49400AB22960AC1818FA81F8C8" bold="true" box="[1268,1289,1882,1908]" pageId="7" pageNumber="8">D</emphasis>
|
||||
. G1, two views. Abbreviations: G1 = gonopod 1; G2 = gonopod 2; i = intercalated platelets; p = pleotelson; p.b. = press-button; s4 = expansion of sternite 4; 1–8 = thoracic sternites 1–8; 2/3–7/8 = thoracic sternal sutures 2/3–7/8. Scale bars: A = 2 mm; B–C = 1 mm; D = 0.5 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF4596D34005B22664E51853FD60F860" ID-DOI="http://doi.org/10.5281/zenodo.3906762" ID-Zenodo-Dep="3906762" httpUri="https://zenodo.org/record/3906762/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,1809,1835]" targetBox="[189,1344,300,1745]" targetPageId="8">
|
||||
<paragraph id="8B85C65B4005B22664E51853FD60F860" blockId="8.[189,1399,1809,2012]" pageId="8" pageNumber="9">
|
||||
<emphasis id="B94E1A494005B22664E51853FE85F897" bold="true" box="[189,269,1809,1835]" pageId="8" pageNumber="9">Fig. 3.</emphasis>
|
||||
<taxonomicName id="4C3ABDD84005B226654F1853FE73F897" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[279,507,1809,1835]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494005B226654F1853FE73F897" box="[279,507,1809,1835]" italics="true" pageId="8" pageNumber="9">Richerius marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324005B226665D1850FD49F890" box="[517,705,1810,1836]" pageId="8" pageNumber="9" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
, paratype, ovigerous ♀, New Caledonia: North Province, 21º21.890ʹ S, 165º32.683ʹ E, Creek stream, tributary of the Böua at level of the Néaoua dam, Houaïlou township, st. HYNC 1823, 476 m a.s.l., “Our Planet Reviewed”, Hydrobio Expedition, Nicolas Charpin leg., 9 Oct. 2017, 7.0 × 7.1 mm (MNHN-IU-2014-21501).
|
||||
<emphasis id="B94E1A494005B22667D01839FC15F829" bold="true" box="[904,925,1915,1941]" pageId="8" pageNumber="9">A</emphasis>
|
||||
.Thoracic sternum, with lowered pleon.
|
||||
<emphasis id="B94E1A494005B22664E518DCFF59F804" bold="true" box="[189,209,1950,1976]" pageId="8" pageNumber="9">B</emphasis>
|
||||
. Pleon. Abbreviations: b.a. = branchiosternal aperture; e = egg; s4 = expansion of sternite 4; v = vulva; 4–8 = thoracic sternites 4–8. Scale bar = 3 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF4596D34004B22764E51826FCAAF855" ID-DOI="http://doi.org/10.5281/zenodo.3906764" ID-Zenodo-Dep="3906764" httpUri="https://zenodo.org/record/3906764/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,232,1892,1918]" targetBox="[208,1380,266,1860]" targetPageId="9">
|
||||
<paragraph id="8B85C65B4004B22764E51826FCAAF855" blockId="9.[189,1399,1891,2025]" pageId="9" pageNumber="10">
|
||||
<emphasis id="B94E1A494004B22764E51826FE82F8C2" bold="true" box="[189,266,1892,1918]" pageId="9" pageNumber="10">Fig. 4.</emphasis>
|
||||
<taxonomicName id="4C3ABDD84004B227654A1827FE7BF8C2" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[274,499,1893,1919]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494004B227654A1827FE7BF8C2" box="[274,499,1893,1919]" italics="true" pageId="9" pageNumber="10">Richerius marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324004B22765A21827FD25F8C3" box="[506,685,1893,1919]" pageId="9" pageNumber="10" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
, holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500).
|
||||
<emphasis id="B94E1A494004B22764E5188CFF5AF854" bold="true" box="[189,210,1998,2024]" pageId="9" pageNumber="10">A</emphasis>
|
||||
. Dorsal view.
|
||||
<emphasis id="B94E1A494004B2276524188CFE18F854" bold="true" box="[380,400,1998,2024]" pageId="9" pageNumber="10">B</emphasis>
|
||||
. Ventral view. Scale bars = 2 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B85C65B4007B22464E51E49FAC1FE0E" blockId="10.[189,1399,267,435]" pageId="10" pageNumber="11">
|
||||
Thoracic sternum of male (
|
||||
<figureCitation id="1301DADE4007B224665F1E49FDEEFE99" box="[519,614,267,293]" captionStart="Fig" captionStartId="7.[189,232,1846,1872]" captionTargetBox="[260,1308,282,1788]" captionTargetId="figure@7.[248,1339,265,1807]" captionTargetPageId="7" captionText="Fig. 2. Richerius marqueti gen. et sp. nov., holotype, ♂, 4.9 × 5.0 mm (MNHN-IU-2014-21500), same collection data as Fig. 1. A. Thoracic sternum. B. Mxp3. C. Sternopleonal cavity with G1s. D. G1, two views. Abbreviations: G1 = gonopod 1; G2 = gonopod 2; i = intercalated platelets; p = pleotelson; p.b. = press-button; s4 = expansion of sternite 4; 1–8 = thoracic sternites 1–8; 2/3–7/8 = thoracic sternal sutures 2/3–7/8. Scale bars: A = 2 mm; B–C = 1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3906760" httpUri="https://zenodo.org/record/3906760/files/figure.png" pageId="10" pageNumber="11">Fig. 2A</figureCitation>
|
||||
) with sternites 4–8 considerably enlarged, with suture 4/5 to 7/8 laterally confined. Sternites 1 and 2 not separated by suture but their demarcation visible on lateral margins; sternite 2 separated from sternite 3 by complete suture; sternite 3 as narrow pentagonal band; suture 3/4 tiny, only lateral, just at level of extension of sternite 4. No longitudinal line. Paired branchiosternal canal apertures located posterolaterally on sternite 8, concealed by pleon (
|
||||
<figureCitation id="1301DADE4007B22460851EDAFAB3FE0E" box="[1245,1339,408,434]" captionStart="Fig" captionStartId="8.[189,232,1809,1835]" captionTargetBox="[189,1344,300,1745]" captionTargetId="figure@8.[189,1398,265,1759]" captionTargetPageId="8" captionText="Fig. 3. Richerius marqueti gen. et sp. nov., paratype, ovigerous ♀, New Caledonia: North Province, 21º21.890ʹ S, 165º32.683ʹ E, Creek stream, tributary of the Böua at level of the Néaoua dam, Houaïlou township, st. HYNC 1823, 476 m a.s.l., “Our Planet Reviewed”, Hydrobio Expedition, Nicolas Charpin leg., 9 Oct. 2017, 7.0 × 7.1 mm (MNHN-IU-2014-21501). A.Thoracic sternum, with lowered pleon. B. Pleon. Abbreviations: b.a. = branchiosternal aperture; e = egg; s4 = expansion of sternite 4; v = vulva; 4–8 = thoracic sternites 4–8. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3906762" httpUri="https://zenodo.org/record/3906762/files/figure.png" pageId="10" pageNumber="11">Fig. 3A</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4007B22464E51E9BFBD1FD3C" blockId="10.[189,1399,473,641]" pageId="10" pageNumber="11">
|
||||
Sternopleonal cavity of male short, triangular, deep (
|
||||
<figureCitation id="1301DADE4007B22467431E9BFCFEFE48" box="[795,886,473,500]" captionStart="Fig" captionStartId="7.[189,232,1846,1872]" captionTargetBox="[260,1308,282,1788]" captionTargetId="figure@7.[248,1339,265,1807]" captionTargetPageId="7" captionText="Fig. 2. Richerius marqueti gen. et sp. nov., holotype, ♂, 4.9 × 5.0 mm (MNHN-IU-2014-21500), same collection data as Fig. 1. A. Thoracic sternum. B. Mxp3. C. Sternopleonal cavity with G1s. D. G1, two views. Abbreviations: G1 = gonopod 1; G2 = gonopod 2; i = intercalated platelets; p = pleotelson; p.b. = press-button; s4 = expansion of sternite 4; 1–8 = thoracic sternites 1–8; 2/3–7/8 = thoracic sternal sutures 2/3–7/8. Scale bars: A = 2 mm; B–C = 1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3906760" httpUri="https://zenodo.org/record/3906760/files/figure.png" pageId="10" pageNumber="11">Fig. 2C</figureCitation>
|
||||
). Pleons in both sexes (
|
||||
<figureCitation id="1301DADE4007B22460D91E9BFB68FE48" box="[1153,1248,473,500]" captionStart="Fig" captionStartId="7.[189,232,1846,1872]" captionTargetBox="[260,1308,282,1788]" captionTargetId="figure@7.[248,1339,265,1807]" captionTargetPageId="7" captionText="Fig. 2. Richerius marqueti gen. et sp. nov., holotype, ♂, 4.9 × 5.0 mm (MNHN-IU-2014-21500), same collection data as Fig. 1. A. Thoracic sternum. B. Mxp3. C. Sternopleonal cavity with G1s. D. G1, two views. Abbreviations: G1 = gonopod 1; G2 = gonopod 2; i = intercalated platelets; p = pleotelson; p.b. = press-button; s4 = expansion of sternite 4; 1–8 = thoracic sternites 1–8; 2/3–7/8 = thoracic sternal sutures 2/3–7/8. Scale bars: A = 2 mm; B–C = 1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3906760" httpUri="https://zenodo.org/record/3906760/files/figure.png" pageId="10" pageNumber="11">Figs 2A</figureCitation>
|
||||
,
|
||||
<figureCitation id="1301DADE4007B22460B31E98FA86FE48" box="[1259,1294,474,500]" captionStart="Fig" captionStartId="8.[189,232,1809,1835]" captionTargetBox="[189,1344,300,1745]" captionTargetId="figure@8.[189,1398,265,1759]" captionTargetPageId="8" captionText="Fig. 3. Richerius marqueti gen. et sp. nov., paratype, ovigerous ♀, New Caledonia: North Province, 21º21.890ʹ S, 165º32.683ʹ E, Creek stream, tributary of the Böua at level of the Néaoua dam, Houaïlou township, st. HYNC 1823, 476 m a.s.l., “Our Planet Reviewed”, Hydrobio Expedition, Nicolas Charpin leg., 9 Oct. 2017, 7.0 × 7.1 mm (MNHN-IU-2014-21501). A.Thoracic sternum, with lowered pleon. B. Pleon. Abbreviations: b.a. = branchiosternal aperture; e = egg; s4 = expansion of sternite 4; v = vulva; 4–8 = thoracic sternites 4–8. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3906762" httpUri="https://zenodo.org/record/3906762/files/figure.png" pageId="10" pageNumber="11">3B</figureCitation>
|
||||
) without fused somites, except 6 fused to telson, thus five somites plus pleotelson. Male pleon widely triangular, regularly widening from base to pleotelson; pleotelson widely triangular, somewhat trilobed, with intercalated plates partially delineated, salient and visible laterally at base, partially included. Pressbutton on lateral side of sternopleonal cavity (
|
||||
<figureCitation id="1301DADE4007B22466801D24FCBDFD3C" box="[728,821,614,640]" captionStart="Fig" captionStartId="7.[189,232,1846,1872]" captionTargetBox="[260,1308,282,1788]" captionTargetId="figure@7.[248,1339,265,1807]" captionTargetPageId="7" captionText="Fig. 2. Richerius marqueti gen. et sp. nov., holotype, ♂, 4.9 × 5.0 mm (MNHN-IU-2014-21500), same collection data as Fig. 1. A. Thoracic sternum. B. Mxp3. C. Sternopleonal cavity with G1s. D. G1, two views. Abbreviations: G1 = gonopod 1; G2 = gonopod 2; i = intercalated platelets; p = pleotelson; p.b. = press-button; s4 = expansion of sternite 4; 1–8 = thoracic sternites 1–8; 2/3–7/8 = thoracic sternal sutures 2/3–7/8. Scale bars: A = 2 mm; B–C = 1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3906760" httpUri="https://zenodo.org/record/3906760/files/figure.png" pageId="10" pageNumber="11">Fig. 2C</figureCitation>
|
||||
). Male gonopore sternal.
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4007B22564E51DEAFD8AFED0" blockId="10.[189,1399,680,777]" lastBlockId="11.[189,1399,267,364]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
|
||||
Female pleon (
|
||||
<figureCitation id="1301DADE4007B224652C1DEAFE33FD7E" box="[372,443,680,706]" captionStart="Fig" captionStartId="8.[189,232,1809,1835]" captionTargetBox="[189,1344,300,1745]" captionTargetId="figure@8.[189,1398,265,1759]" captionTargetPageId="8" captionText="Fig. 3. Richerius marqueti gen. et sp. nov., paratype, ovigerous ♀, New Caledonia: North Province, 21º21.890ʹ S, 165º32.683ʹ E, Creek stream, tributary of the Böua at level of the Néaoua dam, Houaïlou township, st. HYNC 1823, 476 m a.s.l., “Our Planet Reviewed”, Hydrobio Expedition, Nicolas Charpin leg., 9 Oct. 2017, 7.0 × 7.1 mm (MNHN-IU-2014-21501). A.Thoracic sternum, with lowered pleon. B. Pleon. Abbreviations: b.a. = branchiosternal aperture; e = egg; s4 = expansion of sternite 4; v = vulva; 4–8 = thoracic sternites 4–8. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3906762" httpUri="https://zenodo.org/record/3906762/files/figure.png" pageId="10" pageNumber="11">Fig. 3</figureCitation>
|
||||
A–B) oval to expanded in ovigerous females. Female pleopods 2–5 biramous. Ovigerous females with very few and large eggs measuring
|
||||
<quantity id="4CC26BBE4007B22467C21D89FB8DFD5A" box="[922,1029,715,742]" metricMagnitude="-4" metricUnit="m" metricValue="9.6" pageId="10" pageNumber="11" unit="mm" value="0.96">0.96 mm</quantity>
|
||||
in diameter (
|
||||
<figureCitation id="1301DADE4007B22460F31D89FA84FD59" box="[1195,1292,715,741]" captionStart="Fig" captionStartId="8.[189,232,1809,1835]" captionTargetBox="[189,1344,300,1745]" captionTargetId="figure@8.[189,1398,265,1759]" captionTargetPageId="8" captionText="Fig. 3. Richerius marqueti gen. et sp. nov., paratype, ovigerous ♀, New Caledonia: North Province, 21º21.890ʹ S, 165º32.683ʹ E, Creek stream, tributary of the Böua at level of the Néaoua dam, Houaïlou township, st. HYNC 1823, 476 m a.s.l., “Our Planet Reviewed”, Hydrobio Expedition, Nicolas Charpin leg., 9 Oct. 2017, 7.0 × 7.1 mm (MNHN-IU-2014-21501). A.Thoracic sternum, with lowered pleon. B. Pleon. Abbreviations: b.a. = branchiosternal aperture; e = egg; s4 = expansion of sternite 4; v = vulva; 4–8 = thoracic sternites 4–8. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3906762" httpUri="https://zenodo.org/record/3906762/files/figure.png" pageId="10" pageNumber="11">Fig. 3A</figureCitation>
|
||||
). Vulvae located on undivided portion of thoracic sternum corresponding to sternite 6, thus not deplaced anteriorly. First gonopods stout, practically filling most of sternopleonal cavity (
|
||||
<figureCitation id="1301DADE4006B225604B1E49FBFDFE99" box="[1043,1141,267,293]" captionStart="Fig" captionStartId="7.[189,232,1846,1872]" captionTargetBox="[260,1308,282,1788]" captionTargetId="figure@7.[248,1339,265,1807]" captionTargetPageId="7" captionText="Fig. 2. Richerius marqueti gen. et sp. nov., holotype, ♂, 4.9 × 5.0 mm (MNHN-IU-2014-21500), same collection data as Fig. 1. A. Thoracic sternum. B. Mxp3. C. Sternopleonal cavity with G1s. D. G1, two views. Abbreviations: G1 = gonopod 1; G2 = gonopod 2; i = intercalated platelets; p = pleotelson; p.b. = press-button; s4 = expansion of sternite 4; 1–8 = thoracic sternites 1–8; 2/3–7/8 = thoracic sternal sutures 2/3–7/8. Scale bars: A = 2 mm; B–C = 1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3906760" httpUri="https://zenodo.org/record/3906760/files/figure.png" pageId="11" pageNumber="12">Fig. 2C</figureCitation>
|
||||
). G1 curved at base, otherwise with little curvature, with terminal portion bearing fine setae, terminating in one lobe, without corneous process (
|
||||
<figureCitation id="1301DADE4006B22565CE1E10FE7DFED0" box="[406,501,338,364]" captionStart="Fig" captionStartId="7.[189,232,1846,1872]" captionTargetBox="[260,1308,282,1788]" captionTargetId="figure@7.[248,1339,265,1807]" captionTargetPageId="7" captionText="Fig. 2. Richerius marqueti gen. et sp. nov., holotype, ♂, 4.9 × 5.0 mm (MNHN-IU-2014-21500), same collection data as Fig. 1. A. Thoracic sternum. B. Mxp3. C. Sternopleonal cavity with G1s. D. G1, two views. Abbreviations: G1 = gonopod 1; G2 = gonopod 2; i = intercalated platelets; p = pleotelson; p.b. = press-button; s4 = expansion of sternite 4; 1–8 = thoracic sternites 1–8; 2/3–7/8 = thoracic sternal sutures 2/3–7/8. Scale bars: A = 2 mm; B–C = 1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3906760" httpUri="https://zenodo.org/record/3906760/files/figure.png" pageId="11" pageNumber="12">Fig. 2D</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="DF4596D34007B22464E5187DFCD8F85A" ID-DOI="http://doi.org/10.5281/zenodo.3906766" ID-Zenodo-Dep="3906766" httpUri="https://zenodo.org/record/3906766/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,232,1855,1881]" targetBox="[189,1398,901,1815]" targetPageId="10">
|
||||
<paragraph id="8B85C65B4007B22464E5187DFCD8F85A" blockId="10.[189,1399,1853,2022]" pageId="10" pageNumber="11">
|
||||
<emphasis id="B94E1A494007B22464E5187DFE82F8E5" bold="true" box="[189,266,1855,1881]" pageId="10" pageNumber="11">Fig. 5.</emphasis>
|
||||
<taxonomicName id="4C3ABDD84007B224654A187DFE7BF8E5" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[274,499,1855,1881]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494007B224654A187DFE7BF8E5" box="[274,499,1855,1881]" italics="true" pageId="10" pageNumber="11">Richerius marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324007B22465A2187DFD25F8E5" box="[506,685,1855,1881]" pageId="10" pageNumber="11" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
, holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500).
|
||||
<emphasis id="B94E1A494007B22464E518EAFF5AF87E" bold="true" box="[189,210,1960,1986]" pageId="10" pageNumber="11">A</emphasis>
|
||||
. Type locality, stream where several specimens of the species were collected.
|
||||
<emphasis id="B94E1A494007B22460CA18EBFB41F87F" bold="true" box="[1170,1225,1961,1987]" pageId="10" pageNumber="11">B–D</emphasis>
|
||||
. Views of the holotype in vivo: dorsal (B), ventral (C) and frontal (D).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8B85C65B4006B22564E51ED1FE9DFE11" blockId="11.[189,277,403,429]" box="[189,277,403,429]" pageId="11" pageNumber="12">
|
||||
<heading id="D0CD71374006B22564E51ED1FE9DFE11" bold="true" box="[189,277,403,429]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
|
||||
<emphasis id="B94E1A494006B22564E51ED1FE9DFE11" bold="true" box="[189,277,403,429]" pageId="11" pageNumber="12">Colour</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4006B22564E51E80FBCAFE60" blockId="11.[189,1090,449,476]" box="[189,1090,449,476]" pageId="11" pageNumber="12">
|
||||
In life (
|
||||
<figureCitation id="1301DADE4006B225654C1E80FED1FE60" box="[276,345,449,476]" captionStart="Fig" captionStartId="10.[189,232,1855,1881]" captionTargetBox="[189,1398,901,1815]" captionTargetId="figure@10.[189,1398,901,1816]" captionTargetPageId="10" captionText="Fig. 5. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Type locality, stream where several specimens of the species were collected. B–D. Views of the holotype in vivo: dorsal (B), ventral (C) and frontal (D)." figureDoi="http://doi.org/10.5281/zenodo.3906766" httpUri="https://zenodo.org/record/3906766/files/figure.png" pageId="11" pageNumber="12">Fig. 5</figureCitation>
|
||||
B–D), the colour is overall brown, mottled with yellowish dots.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C32095D04006B22564E51D41FEDAFCFE" pageId="11" pageNumber="12" type="distribution">
|
||||
<paragraph id="8B85C65B4006B22564E51D41FEDEFDA1" blockId="11.[189,342,515,541]" box="[189,342,515,541]" pageId="11" pageNumber="12">
|
||||
<heading id="D0CD71374006B22564E51D41FEDEFDA1" bold="true" box="[189,342,515,541]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
|
||||
<emphasis id="B94E1A494006B22564E51D41FEDEFDA1" bold="true" box="[189,342,515,541]" pageId="11" pageNumber="12">Distribution</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4006B22564E51D70FEDAFCFE" blockId="11.[189,1399,562,834]" pageId="11" pageNumber="12">
|
||||
<taxonomicName id="4C3ABDD84006B22564E51D70FE29FDF0" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[189,417,562,588]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494006B22564E51D70FE29FDF0" box="[189,417,562,588]" italics="true" pageId="11" pageNumber="12">Richerius marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324006B22565F31D70FDEDFDF0" box="[427,613,562,588]" pageId="11" pageNumber="12" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
is exclusively known from freshwaters in small flowing streams where it lives among the aquatic vegetation (
|
||||
<figureCitation id="1301DADE4006B225668A1D17FCB9FDCC" box="[722,817,597,624]" captionStart="Fig" captionStartId="10.[189,232,1855,1881]" captionTargetBox="[189,1398,901,1815]" captionTargetId="figure@10.[189,1398,901,1816]" captionTargetPageId="10" captionText="Fig. 5. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Type locality, stream where several specimens of the species were collected. B–D. Views of the holotype in vivo: dorsal (B), ventral (C) and frontal (D)." figureDoi="http://doi.org/10.5281/zenodo.3906766" httpUri="https://zenodo.org/record/3906766/files/figure.png" pageId="11" pageNumber="12">Fig. 5A</figureCitation>
|
||||
). It is found at an altitude of up
|
||||
<quantity id="4CC26BBE4006B22560EA1D17FB77FDD3" box="[1202,1279,597,624]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="12" unit="m" value="500.0">500 m</quantity>
|
||||
, therefore further from the sea than
|
||||
<taxonomicName id="4C3ABDD84006B22565B21D3AFD4AFD2E" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[490,706,632,658]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494006B22565B21D3AFD4AFD2E" box="[490,706,632,658]" italics="true" pageId="11" pageNumber="12">Odiomaris pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
(that is usually encountered in the lower course of rivers). By comparison,
|
||||
<taxonomicName id="4C3ABDD84006B22565D91DDEFCA9FD09" authority="(Chilton, 1882)" baseAuthorityName="Chilton" baseAuthorityYear="1882" box="[385,801,667,694]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="lacustris">
|
||||
<emphasis id="B94E1A494006B22565D91DDEFDEAFD09" box="[385,610,667,694]" italics="true" pageId="11" pageNumber="12">Amarinus lacustris</emphasis>
|
||||
(Chilton, 1882)
|
||||
</taxonomicName>
|
||||
has been reported from
|
||||
<collectingCountry id="F32D86CB4006B225601F1DDEFB6EFD09" box="[1095,1254,667,694]" name="New Zealand" pageId="11" pageNumber="12">New Zealand</collectingCountry>
|
||||
freshwaters from Lord Howe Island at altitudes between
|
||||
<quantity id="4CC26BBE4006B22566B91DFCFCC4FD65" box="[737,844,702,729]" metricMagnitude="1" metricUnit="m" metricValue="7.65" metricValueMax="9.2" metricValueMin="6.1" pageId="11" pageNumber="12" unit="m" value="76.5" valueMax="92.0" valueMin="61.0">61–92 m</quantity>
|
||||
(
|
||||
<bibRefCitation id="EFABBBAA4006B22567061DFCFB91FD65" author="Etheridge R." box="[862,1049,702,729]" pageId="11" pageNumber="12" pagination="1 - 4" refId="ref12979" refString="Etheridge R. 1889. The general zoology of Lord Howe Island; containing also an account of the collections made by the Australian Museum Collecting Party, Aug. - Sept., 1887. Australian Museum Memoir 2 (1): 1 - 4. https: // doi. org / 10.3853 / j. 0067 - 1967.2.1889.479" type="journal article" year="1889">
|
||||
Etheridge
|
||||
<date id="FF84E09B4006B22567851DFDFB91FD65" box="[989,1049,703,729]" pageId="11" pageNumber="12" value="1889">1889</date>
|
||||
</bibRefCitation>
|
||||
) and even about
|
||||
<quantity id="4CC26BBE4006B22560B61DFDFAFFFD65" box="[1262,1399,703,729]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="11" pageNumber="12" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">800–900 m</quantity>
|
||||
(
|
||||
<bibRefCitation id="EFABBBAA4006B225649D1DA0FEEDFD40" author="Chilton C." box="[197,357,737,764]" pageId="11" pageNumber="12" pagination="316 - 320" refId="ref12420" refString="Chilton C. 1915. A freshwater crab and its distribution in Australia and New Zealand. Transactions and Proceedings of the New Zealand Institute 47: 316 - 320. Available from http: // rsnz. natlib. govt. nz / volume / rsnz _ 47 / rsnz _ 47 _ 00 _ 005860. html [accessed 27 May 2020]." type="journal article" year="1915">
|
||||
Chilton
|
||||
<date id="FF84E09B4006B225657F1DA3FEEDFD40" box="[295,357,737,764]" pageId="11" pageNumber="12" value="1915">1915</date>
|
||||
</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EFABBBAA4006B22565291DA0FD95FD41" author="Holthuis L. B." box="[369,541,738,765]" pageId="11" pageNumber="12" pagination="109 - 121" refId="ref13478" refString="Holthuis L. B. 1968. On Hymenosomatidae (Crustacea Decapoda Brachyura) from fresh water, with the description of a new species. Beaufortia 15 (195): 109 - 121. Available from https: // www. repository. naturalis. nl / record / 504776 [accessed 27 May 2020]." type="journal article" year="1968">
|
||||
Holthuis
|
||||
<date id="FF84E09B4006B22565861DA0FD95FD41" box="[478,541,738,765]" pageId="11" pageNumber="12" value="1968">1968</date>
|
||||
</bibRefCitation>
|
||||
);
|
||||
<taxonomicName id="4C3ABDD84006B225666B1DA0FCF5FD40" authority="Holthuis, 1968" authorityName="Holthuis" authorityYear="1968" box="[563,893,738,764]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="angelicus">
|
||||
<emphasis id="B94E1A494006B225666B1DA0FD4CFD40" box="[563,708,738,764]" italics="true" pageId="11" pageNumber="12">A. angelicus</emphasis>
|
||||
<bibRefCitation id="EFABBBAA4006B22566941DA0FCF5FD40" author="Holthuis L. B." box="[716,893,738,764]" pageId="11" pageNumber="12" pagination="109 - 121" refId="ref13478" refString="Holthuis L. B. 1968. On Hymenosomatidae (Crustacea Decapoda Brachyura) from fresh water, with the description of a new species. Beaufortia 15 (195): 109 - 121. Available from https: // www. repository. naturalis. nl / record / 504776 [accessed 27 May 2020]." type="journal article" year="1968">Holthuis, 1968</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, from the central mountain range of
|
||||
<collectingCountry id="F32D86CB4006B22561771DA0FEDBFCA3" name="Papua New Guinea" pageId="11" pageNumber="12">Papua New Guinea</collectingCountry>
|
||||
, was collected in pure freshwater of a watercress swamp at an altitude of
|
||||
<quantity id="4CC26BBE4006B22560F11C47FA89FCA3" box="[1193,1281,773,799]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="12" unit="m" value="1600.0">1600 m</quantity>
|
||||
(
|
||||
<bibRefCitation id="EFABBBAA4006B22561481C47FF72FCFE" author="Holthuis L. B." pageId="11" pageNumber="12" pagination="109 - 121" refId="ref13478" refString="Holthuis L. B. 1968. On Hymenosomatidae (Crustacea Decapoda Brachyura) from fresh water, with the description of a new species. Beaufortia 15 (195): 109 - 121. Available from https: // www. repository. naturalis. nl / record / 504776 [accessed 27 May 2020]." type="journal article" year="1968">
|
||||
Holthuis
|
||||
<date id="FF84E09B4006B22564E51C6AFF72FCFE" box="[189,250,808,834]" pageId="11" pageNumber="12" value="1968">1968</date>
|
||||
</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EFABBBAA4006B225655E1C6AFECDFCFE" author="Holthuis L. B." box="[262,325,808,834]" pageId="11" pageNumber="12" pagination="603 - 619" refId="ref13537" refString="Holthuis L. B. 1982. Freshwater Crustacea Decapoda of New Guinea. In: Gressitt J. L. (ed.) Biogeography and Ecology of New Guinea. Vol. 2, Pt. 6. Monographiae Biologicae 42: 603 - 619. https: // doi. org / 10.1007 / 978 - 94 - 009 - 8632 - 9 _ 28" type="journal article" year="1982">
|
||||
<date id="FF84E09B4006B225655E1C6AFECDFCFE" box="[262,325,808,834]" pageId="11" pageNumber="12" value="1982">1982</date>
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C32095D04006B22364E51C28FB42F8FE" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" type="discussion">
|
||||
<paragraph id="8B85C65B4006B22564E51C28FEA7FC38" blockId="11.[189,303,874,900]" box="[189,303,874,900]" pageId="11" pageNumber="12">
|
||||
<heading id="D0CD71374006B22564E51C28FEA7FC38" bold="true" box="[189,303,874,900]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
|
||||
<emphasis id="B94E1A494006B22564E51C28FEA7FC38" bold="true" box="[189,303,874,900]" pageId="11" pageNumber="12">Remarks</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4006B22564E51CDAFC25FB53" blockId="11.[189,1399,920,1263]" pageId="11" pageNumber="12">
|
||||
The subfamily
|
||||
<taxonomicName id="4C3ABDD84006B22565241CDAFD9DFC0E" authorityName="Guinot" authorityYear="2011" box="[380,533,920,946]" class="Malacostraca" family="Hymenosomatidae" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subFamily" subFamily="Odiomarinae">Odiomarinae</taxonomicName>
|
||||
was erected by
|
||||
<bibRefCitation id="EFABBBAA4006B22566B51CDAFC2EFC0E" author="Guinot D." box="[749,934,920,946]" pageId="11" pageNumber="12" pagination="20 - 32" refId="ref13311" refString="Guinot D. 2011 a. Odiomarinae nov. subfam., a new subfamily for two primitive genera of the Hymenosomatidae MacLeay, 1838 (Crustacea, Decapoda, Brachyura). Zootaxa 2732: 20 - 32. https: // doi. org / 10.11646 / zootaxa. 2732.1.2" type="journal article" year="2011">Guinot (2011a)</bibRefCitation>
|
||||
to receive two genera of the family
|
||||
<taxonomicName id="4C3ABDD84006B22564E51CF9FE11FC69" authorityName="MacLeay" authorityYear="1838" box="[189,409,955,981]" class="Malacostraca" family="Hymenosomatidae" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="family">Hymenosomatidae</taxonomicName>
|
||||
characterised by the presence on the male pleon of intercalated platelets, either articulated and moveable (
|
||||
<bibRefCitation id="EFABBBAA4006B22565A31C9CFD2BFC45" author="Guinot D." box="[507,675,990,1017]" pageId="11" pageNumber="12" pagination="20 - 32" refId="ref13311" refString="Guinot D. 2011 a. Odiomarinae nov. subfam., a new subfamily for two primitive genera of the Hymenosomatidae MacLeay, 1838 (Crustacea, Decapoda, Brachyura). Zootaxa 2732: 20 - 32. https: // doi. org / 10.11646 / zootaxa. 2732.1.2" type="journal article" year="2011">Guinot 2011a</bibRefCitation>
|
||||
: fig. 2) or relatively less well-demarcated:
|
||||
<taxonomicName id="4C3ABDD84006B22560F21C9CFE46FBA0" authority="Ng & Richer de Forges, 1996" authorityName="Ng & Richer de Forges" authorityYear="1996" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494006B22560F21C9CFAAFFC44" box="[1194,1319,990,1016]" italics="true" pageId="11" pageNumber="12">Odiomaris</emphasis>
|
||||
<bibRefCitation id="EFABBBAA4006B225616A1C9DFE46FBA0" author="Ng P. K. L. & Richer de Forges B." pageId="11" pageNumber="12" pagination="263 - 276" refId="ref14618" refString="Ng P. K. L. & Richer de Forges B. 1996. The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of New Caledonia, with descriptions of two new genera and two new species. Memoirs of the Queensland Museum 39 (2): 263 - 276. Available from https: // www. biodiversitylibrary. org / part / 50813 # / summary [accessed 27 May 2020]." type="journal article" year="1996">Ng & Richer de Forges, 1996</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, endemic to
|
||||
<collectingCountry id="F32D86CB4006B22566381B40FC90FBA0" box="[608,792,1026,1052]" name="New Caledonia" pageId="11" pageNumber="12">New Caledonia</collectingCountry>
|
||||
, and
|
||||
<emphasis id="B94E1A494006B225670C1B40FC33FBA0" box="[852,955,1026,1052]" italics="true" pageId="11" pageNumber="12">Amarinu</emphasis>
|
||||
(at least
|
||||
<emphasis id="B94E1A494006B225607A1B41FB07FBA0" box="[1058,1167,1027,1052]" italics="true" pageId="11" pageNumber="12">pro parte</emphasis>
|
||||
), mostly from fresh and estuarine waters of the Indo-West Pacific region.
|
||||
<emphasis id="B94E1A494006B225676E1B67FC13FB83" box="[822,923,1061,1087]" italics="true" pageId="11" pageNumber="12">Richeriu</emphasis>
|
||||
s gen. nov. shows several plesiomorphic characters that include the male and female pleons without fused somites (except for pleotelson), thus consisting of six elements (namely the maximum of somites existing in
|
||||
<taxonomicName id="4C3ABDD84006B22560041B29FAB5FB3A" authorityName="MacLeay" authorityYear="1838" box="[1116,1341,1131,1158]" class="Malacostraca" family="Hymenosomatidae" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="family">Hymenosomatidae</taxonomicName>
|
||||
), the prominent, partially demarcated intercalated platelets, the thoracic sternum with the anterior sternites forming a small produced plate, the vulvae not anteriorly displaced, the G1 only gently curved and simple.
|
||||
<taxonomicName id="4C3ABDD84006B22565421B97FE02FB53" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[282,394,1237,1263]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B94E1A494006B22565421B97FE02FB53" box="[282,394,1237,1263]" italics="true" pageId="11" pageNumber="12">Richerius</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324006B22565CA1B97FE75FB53" box="[402,509,1237,1263]" pageId="11" pageNumber="12" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
can be assigned to the
|
||||
<taxonomicName id="4C3ABDD84006B22567541B97FC21FB53" authorityName="Guinot" authorityYear="2011" box="[780,937,1237,1263]" class="Malacostraca" family="Hymenosomatidae" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subFamily" subFamily="Odiomarinae">Odiomarinae</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4006B22564E51A54FEBCF9F6" blockId="11.[189,1399,1302,1611]" pageId="11" pageNumber="12">
|
||||
<taxonomicName id="4C3ABDD84006B22564E51A54FEB2FA8C" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[189,314,1302,1328]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494006B22564E51A54FEB2FA8C" box="[189,314,1302,1328]" italics="true" pageId="11" pageNumber="12">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
is known from two species: the
|
||||
<typeStatus id="548178F94006B22566811A55FC84FA8D" box="[729,780,1303,1329]" pageId="11" pageNumber="12">type</typeStatus>
|
||||
species
|
||||
<taxonomicName id="4C3ABDD84006B22567211A54FAF9FA8C" authority="A. Milne-Edwards, 1873" authorityName="A. Milne-Edwards" authorityYear="1873" box="[889,1393,1302,1329]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pilosa">
|
||||
<emphasis id="B94E1A494006B22567211A54FBBEFA8C" box="[889,1078,1302,1328]" italics="true" pageId="11" pageNumber="12">Elamena pilosa</emphasis>
|
||||
A. Milne-Edwards, 1873
|
||||
</taxonomicName>
|
||||
, referred to as
|
||||
<taxonomicName id="4C3ABDD84006B22565341A78FD2CFAEF" authority="White, 1846" authorityName="White" authorityYear="1846" box="[364,676,1337,1364]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494006B22565341A78FD8CFAE8" box="[364,516,1338,1364]" italics="true" pageId="11" pageNumber="12">Halicarcinus</emphasis>
|
||||
White, 1846
|
||||
</taxonomicName>
|
||||
by
|
||||
<bibRefCitation id="EFABBBAA4006B22566801A7BFC12FAE8" author="Holthuis L. B." box="[728,922,1337,1364]" pageId="11" pageNumber="12" pagination="109 - 121" refId="ref13478" refString="Holthuis L. B. 1968. On Hymenosomatidae (Crustacea Decapoda Brachyura) from fresh water, with the description of a new species. Beaufortia 15 (195): 109 - 121. Available from https: // www. repository. naturalis. nl / record / 504776 [accessed 27 May 2020]." type="journal article" year="1968">Holthuis (1968)</bibRefCitation>
|
||||
, as
|
||||
<taxonomicName id="4C3ABDD84006B22567961A78FBC8FAE8" authorityName="Lucas" authorityYear="1980" box="[974,1088,1338,1364]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494006B22567961A78FBC8FAE8" box="[974,1088,1338,1364]" italics="true" pageId="11" pageNumber="12">Amarinus</emphasis>
|
||||
</taxonomicName>
|
||||
by
|
||||
<bibRefCitation id="EFABBBAA4006B225602D1A78FA9CFAE8" author="Lucas J. S." box="[1141,1300,1338,1364]" pageId="11" pageNumber="12" pagination="148 - 247" refId="ref14163" refString="Lucas J. S. 1980. Spider crabs of the family Hymenosomatidae (Crustacea: Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum 33 (4): 148 - 247. https: // doi. org / 10.3853 / j. 0067 - 1975.33.1980.199" type="journal article" year="1980">Lucas (1980)</bibRefCitation>
|
||||
then as
|
||||
<taxonomicName id="4C3ABDD84006B22564E51A1FFEB2FACB" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[189,314,1373,1399]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494006B22564E51A1FFEB2FACB" box="[189,314,1373,1399]" italics="true" pageId="11" pageNumber="12">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
by
|
||||
<bibRefCitation id="EFABBBAA4006B22565301A1FFD5FFAC4" author="Ng P. K. L. & Richer de Forges B." box="[360,727,1373,1400]" pageId="11" pageNumber="12" pagination="263 - 276" refId="ref14618" refString="Ng P. K. L. & Richer de Forges B. 1996. The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of New Caledonia, with descriptions of two new genera and two new species. Memoirs of the Queensland Museum 39 (2): 263 - 276. Available from https: // www. biodiversitylibrary. org / part / 50813 # / summary [accessed 27 May 2020]." type="journal article" year="1996">Ng & Richer de Forges (1996)</bibRefCitation>
|
||||
, mainly freshwater but also euryhaline; and
|
||||
<taxonomicName id="4C3ABDD84006B22560BE1A1FFDD9FA26" authority="Davie & Richer de Forges, 1996" authorityName="Davie & Richer de Forges" authorityYear="1996" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494006B22560BE1A1FFAFFFACB" box="[1254,1399,1373,1399]" italics="true" pageId="11" pageNumber="12">O. estuarius</emphasis>
|
||||
<bibRefCitation id="EFABBBAA4006B22564E51AC2FDD9FA26" author="Davie P. J. F. & Richer de Forges B." box="[189,593,1408,1434]" pageId="11" pageNumber="12" pagination="257 - 262" refId="ref12668" refString="Davie P. J. F. & Richer de Forges B. 1996. Two new species of hymenosomatid crabs from New Caledonia. Memoirs of the Queensland Museum 39 (2): 257 - 262." type="journal article" year="1996">Davie & Richer de Forges, 1996</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, exclusively brackish. An important difference between these two species is the rostrum: it is lowered ventrally as a triangular, V-shaped projection in
|
||||
<taxonomicName id="4C3ABDD84006B22560C61AE6FA9DFA01" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[1182,1301,1443,1470]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494006B22560C61AE6FA9DFA01" box="[1182,1301,1443,1470]" italics="true" pageId="11" pageNumber="12">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
so as to be positioned between the antennules (
|
||||
<figureCitation id="1301DADE4006B22566D71A84FD65FA5D" box="[655,749,1478,1505]" captionStart="Fig" captionStartId="16.[189,232,1919,1945]" captionTargetBox="[402,1193,1524,1866]" captionTargetId="figure@16.[385,1203,1509,1878]" captionTargetPageId="16" captionText="Fig. 7. Odiomaris pilosus A. Milne-Edwards, 1873, syntype, ♂, New Caledonia, freshwater, B. Balansa leg. (RMNH D 32). A. Ventral anterior part. B. G1 (after Holthuis 1968: fig. 3a–b). Abbreviations: a1 = antennule, with folded second article; a2 = antenna; e = epistome: p.k. = postorbital knobs; r = rostrum." figureDoi="http://doi.org/10.5281/zenodo.3906770" httpUri="https://zenodo.org/record/3906770/files/figure.png" pageId="11" pageNumber="12">Fig. 7A</figureCitation>
|
||||
) (A. Milne-Edwards 1873: pl. 18, fig. 6a, as
|
||||
<taxonomicName id="4C3ABDD84006B22561481A84FE8DF9B8" authorityName="A.Milne-Edwards" authorityYear="1873" class="Malacostraca" family="Hymenosomatidae" genus="Elamene" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pilosa">
|
||||
<emphasis id="B94E1A494006B22561481A84FE8DF9B8" italics="true" pageId="11" pageNumber="12">Elamene pilosa</emphasis>
|
||||
</taxonomicName>
|
||||
;
|
||||
<bibRefCitation id="EFABBBAA4006B225654E1AA8FDFCF9BF" author="Ng P. K. L. & Richer de Forges B." box="[278,628,1513,1540]" pageId="11" pageNumber="12" pagination="263 - 276" refId="ref14618" refString="Ng P. K. L. & Richer de Forges B. 1996. The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of New Caledonia, with descriptions of two new genera and two new species. Memoirs of the Queensland Museum 39 (2): 263 - 276. Available from https: // www. biodiversitylibrary. org / part / 50813 # / summary [accessed 27 May 2020]." type="journal article" year="1996">Ng & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 6c–d), whereas it is spatulate and does not extend to form a ventral projection between the antennules in
|
||||
<taxonomicName id="4C3ABDD84006B225669F194FFCDFF99B" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[711,855,1549,1575]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494006B225669F194FFCDFF99B" box="[711,855,1549,1575]" italics="true" pageId="11" pageNumber="12">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1301DADE4006B225673E194FFC21F99B" box="[870,937,1549,1575]" captionStart="Fig" captionStartId="18.[189,232,1892,1918]" captionTargetBox="[189,1398,466,1853]" captionTargetId="figure@18.[189,1398,466,1853]" captionTargetPageId="18" captionText="Fig. 8. Odiomaris estuarius Davie & Richer de Forges, 1996, holotype, ♂, New Caledonia, Dumbea estuary, Davie & Richer de Forges leg., 8 Dec. 1993, 4.5 × 4.3 mm (MNHN-IU-2014-11870 = MNHN-B25278). A. Dorsal view. B. Ventral view. Credit: RECOLNAT (ANR-11-INBS-0004) - Noémy MOLLARET – 2017." figureDoi="http://doi.org/10.5281/zenodo.3906772" httpUri="https://zenodo.org/record/3906772/files/figure.png" pageId="11" pageNumber="12">Fig. 8</figureCitation>
|
||||
A–B) (
|
||||
<bibRefCitation id="EFABBBAA4006B22567A2194FFAFBF99B" author="Davie P. J. F. & Richer de Forges B." box="[1018,1395,1549,1575]" pageId="11" pageNumber="12" pagination="257 - 262" refId="ref12668" refString="Davie P. J. F. & Richer de Forges B. 1996. Two new species of hymenosomatid crabs from New Caledonia. Memoirs of the Queensland Museum 39 (2): 257 - 262." type="journal article" year="1996">Davie & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 1a–b).
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4006B22264E51933FBAFFED0" blockId="11.[189,1399,1649,1993]" lastBlockId="12.[189,1399,267,364]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
|
||||
<taxonomicName id="4C3ABDD84006B22564E51933FEA5F937" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[189,301,1649,1675]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B94E1A494006B22564E51933FEA5F937" box="[189,301,1649,1675]" italics="true" pageId="11" pageNumber="12">Richerius</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324006B225656F1930FE2DF930" box="[311,421,1650,1676]" pageId="11" pageNumber="12" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
shares with
|
||||
<taxonomicName id="4C3ABDD84006B22566181933FD35F937" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[576,701,1649,1675]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494006B22566181933FD35F937" box="[576,701,1649,1675]" italics="true" pageId="11" pageNumber="12">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
the same arrangement of the cephalic appendices. But its rostrum is spatulate without folding down ventrally (
|
||||
<figureCitation id="1301DADE4006B225677519D6FCF0F913" box="[813,888,1684,1711]" captionStart="Fig" captionStartId="6.[189,232,1782,1808]" captionTargetBox="[326,1260,292,1710]" captionTargetId="figure@6.[292,1296,265,1741]" captionTargetPageId="6" captionText="Fig. 1. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Carapace, dorsal view. B. Cephalothorax, ventral view, anterior part, with unfolded antennules, the arrow shows antennule with folded second article. Abbreviations: a1 = antennule; a2 = antenna; e = epistome; p.k. = postorbital knob; r = rostrum; u.a. = antennal urinary article; v.r. = ventral expansion of rostrum; 2/3 = thoracic sternal suture 2/3. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3906758" httpUri="https://zenodo.org/record/3906758/files/figure.png" pageId="11" pageNumber="12">Figs 1</figureCitation>
|
||||
,
|
||||
<figureCitation id="1301DADE4006B22567DF19D7FC10F913" box="[903,920,1685,1711]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[208,1380,266,1860]" captionTargetId="figure@9.[208,1380,266,1862]" captionTargetPageId="9" captionText="Fig. 4. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Dorsal view. B. Ventral view. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3906764" httpUri="https://zenodo.org/record/3906764/files/figure.png" pageId="11" pageNumber="12">4</figureCitation>
|
||||
,
|
||||
<figureCitation id="1301DADE4006B22567FE19D6FC44F913" box="[934,972,1684,1711]" captionStart="Fig" captionStartId="10.[189,232,1855,1881]" captionTargetBox="[189,1398,901,1815]" captionTargetId="figure@10.[189,1398,901,1816]" captionTargetPageId="10" captionText="Fig. 5. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Type locality, stream where several specimens of the species were collected. B–D. Views of the holotype in vivo: dorsal (B), ventral (C) and frontal (D)." figureDoi="http://doi.org/10.5281/zenodo.3906766" httpUri="https://zenodo.org/record/3906766/files/figure.png" pageId="11" pageNumber="12">5D</figureCitation>
|
||||
), instead of being lowered ventrally as a V-shaped projection located between the antennules as in
|
||||
<taxonomicName id="4C3ABDD84006B22567D719FAFBEEF96E" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[911,1126,1720,1746]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494006B22567D719FAFBEEF96E" box="[911,1126,1720,1746]" italics="true" pageId="11" pageNumber="12">Odiomaris pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1301DADE4006B225602D19FAFB5CF96E" box="[1141,1236,1720,1746]" captionStart="Fig" captionStartId="16.[189,232,1919,1945]" captionTargetBox="[402,1193,1524,1866]" captionTargetId="figure@16.[385,1203,1509,1878]" captionTargetPageId="16" captionText="Fig. 7. Odiomaris pilosus A. Milne-Edwards, 1873, syntype, ♂, New Caledonia, freshwater, B. Balansa leg. (RMNH D 32). A. Ventral anterior part. B. G1 (after Holthuis 1968: fig. 3a–b). Abbreviations: a1 = antennule, with folded second article; a2 = antenna; e = epistome: p.k. = postorbital knobs; r = rostrum." figureDoi="http://doi.org/10.5281/zenodo.3906770" httpUri="https://zenodo.org/record/3906770/files/figure.png" pageId="11" pageNumber="12">Fig. 7A</figureCitation>
|
||||
). In
|
||||
<taxonomicName id="4C3ABDD84006B225615E19FAFAFEF96E" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[1286,1398,1720,1746]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B94E1A494006B225615E19FAFAFEF96E" box="[1286,1398,1720,1746]" italics="true" pageId="11" pageNumber="12">Richerius</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324006B22564E5199EFEA3F94A" box="[189,299,1756,1782]" pageId="11" pageNumber="12" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
(
|
||||
<figureCitation id="1301DADE4006B22565651999FE12F949" box="[317,410,1755,1781]" captionStart="Fig" captionStartId="6.[189,232,1782,1808]" captionTargetBox="[326,1260,292,1710]" captionTargetId="figure@6.[292,1296,265,1741]" captionTargetPageId="6" captionText="Fig. 1. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Carapace, dorsal view. B. Cephalothorax, ventral view, anterior part, with unfolded antennules, the arrow shows antennule with folded second article. Abbreviations: a1 = antennule; a2 = antenna; e = epistome; p.k. = postorbital knob; r = rostrum; u.a. = antennal urinary article; v.r. = ventral expansion of rostrum; 2/3 = thoracic sternal suture 2/3. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3906758" httpUri="https://zenodo.org/record/3906758/files/figure.png" pageId="11" pageNumber="12">Fig. 1B</figureCitation>
|
||||
), the proepistome is represented by a ventral expansion of the rostrum; therefore, the antennules are located along the proepistome, whereas in
|
||||
<taxonomicName id="4C3ABDD84006B22567EB19BCFB07F8A4" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[947,1167,1790,1816]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494006B22567EB19BCFB07F8A4" box="[947,1167,1790,1816]" italics="true" pageId="11" pageNumber="12">Odiomaris pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
the antennules are obliquely folded in a fossa hollowed all along with the lateral parts of the V-shaped rostrum and the eye.
|
||||
<taxonomicName id="4C3ABDD84006B22564AE1807FE50F8E2" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[246,472,1861,1887]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494006B22564AE1807FE50F8E2" box="[246,472,1861,1887]" italics="true" pageId="11" pageNumber="12">Richerius marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324006B22565B91807FD1DF8E3" box="[481,661,1861,1887]" pageId="11" pageNumber="12" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
actually has a rostrum and proepistome more similar to those of
|
||||
<taxonomicName id="4C3ABDD84006B2256487182AFEFAF83E" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[223,370,1896,1922]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494006B2256487182AFEFAF83E" box="[223,370,1896,1922]" italics="true" pageId="11" pageNumber="12">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1301DADE4006B22565DC182AFE44F83E" box="[388,460,1896,1922]" captionStart="Fig" captionStartId="18.[189,232,1892,1918]" captionTargetBox="[189,1398,466,1853]" captionTargetId="figure@18.[189,1398,466,1853]" captionTargetPageId="18" captionText="Fig. 8. Odiomaris estuarius Davie & Richer de Forges, 1996, holotype, ♂, New Caledonia, Dumbea estuary, Davie & Richer de Forges leg., 8 Dec. 1993, 4.5 × 4.3 mm (MNHN-IU-2014-11870 = MNHN-B25278). A. Dorsal view. B. Ventral view. Credit: RECOLNAT (ANR-11-INBS-0004) - Noémy MOLLARET – 2017." figureDoi="http://doi.org/10.5281/zenodo.3906772" httpUri="https://zenodo.org/record/3906772/files/figure.png" pageId="11" pageNumber="12">Fig. 8</figureCitation>
|
||||
) (
|
||||
<bibRefCitation id="EFABBBAA4006B22565BF182AFCE7F83E" author="Davie P. J. F. & Richer de Forges B." box="[487,879,1896,1922]" pageId="11" pageNumber="12" pagination="257 - 262" refId="ref12668" refString="Davie P. J. F. & Richer de Forges B. 1996. Two new species of hymenosomatid crabs from New Caledonia. Memoirs of the Queensland Museum 39 (2): 257 - 262." type="journal article" year="1996">Davie & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 1) than to those of
|
||||
<taxonomicName id="4C3ABDD84006B22560CE182AFA98F83E" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[1174,1296,1896,1922]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494006B22560CE182AFA98F83E" box="[1174,1296,1896,1922]" italics="true" pageId="11" pageNumber="12">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
. In fact, the genus-level differences between
|
||||
<taxonomicName id="4C3ABDD84006B22566DB18C9FD7BF819" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[643,755,1931,1957]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B94E1A494006B22566DB18C9FD7BF819" box="[643,755,1931,1957]" italics="true" pageId="11" pageNumber="12">Richerius</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324006B225675A18CEFCFBF81A" box="[770,883,1932,1958]" pageId="11" pageNumber="12" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
and
|
||||
<taxonomicName id="4C3ABDD84006B22567E118C9FBBEF819" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[953,1078,1931,1957]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494006B22567E118C9FBBEF819" box="[953,1078,1931,1957]" italics="true" pageId="11" pageNumber="12">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
remain unclear. Actually,
|
||||
<taxonomicName id="4C3ABDD84006B22564E518EDFEC7F874" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[189,335,1967,1993]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494006B22564E518EDFEC7F874" box="[189,335,1967,1993]" italics="true" pageId="11" pageNumber="12">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
seems morphologically closer to
|
||||
<taxonomicName id="4C3ABDD84006B22566BC18ECFCDCF874" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[740,852,1966,1992]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B94E1A494006B22566BC18ECFCDCF874" box="[740,852,1966,1992]" italics="true" pageId="11" pageNumber="12">Richerius</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324006B225670618EDFC43F875" box="[862,971,1967,1993]" pageId="11" pageNumber="12" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
than to
|
||||
<taxonomicName id="4C3ABDD84006B225606918EDFB22F874" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[1073,1194,1966,1993]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494006B225606918EDFB22F874" box="[1073,1194,1966,1993]" italics="true" pageId="11" pageNumber="12">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
, at least in some characteristics. A possible transfer of
|
||||
<taxonomicName id="4C3ABDD84001B222662D1E4EFC8DFE99" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[629,773,268,294]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494001B222662D1E4EFC8DFE99" box="[629,773,268,294]" italics="true" pageId="12" pageNumber="13">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
to
|
||||
<taxonomicName id="4C3ABDD84001B22267721E49FC12FE99" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[810,922,267,293]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B94E1A494001B22267721E49FC12FE99" box="[810,922,267,293]" italics="true" pageId="12" pageNumber="13">Richerius</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324001B22267FA1E4EFB84FE9A" box="[930,1036,268,294]" pageId="12" pageNumber="13" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
has been considered but needs to be decided later, especially when the status of
|
||||
<taxonomicName id="4C3ABDD84001B22266B71E6DFC7BFEF5" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[751,1011,302,329]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B22266B71E6DFCE4FEF5" box="[751,876,303,329]" italics="true" pageId="12" pageNumber="13">Odiomaris</emphasis>
|
||||
aff.
|
||||
<emphasis id="B94E1A494001B22267F81E6DFC7BFEF5" box="[928,1011,303,329]" italics="true" pageId="12" pageNumber="13">pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
is resolved (see below, ‘Remarks on
|
||||
<taxonomicName id="4C3ABDD84001B22264BA1E10FE60FED0" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[226,488,338,364]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B22264BA1E10FED7FED0" box="[226,351,338,364]" italics="true" pageId="12" pageNumber="13">Odiomaris</emphasis>
|
||||
aff.
|
||||
<emphasis id="B94E1A494001B22265CD1E10FE60FED0" box="[405,488,338,364]" italics="true" pageId="12" pageNumber="13">pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
from the Iouanga River and two other streams’).
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4001B22264E51ED1FAD8FA17" blockId="12.[189,1399,403,1451]" pageId="12" pageNumber="13">
|
||||
Along the lower orbital margin there are two salient knobs in both species of
|
||||
<taxonomicName id="4C3ABDD84001B22260601ED1FB3DFE11" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[1080,1205,403,429]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B22260601ED1FB3DFE11" box="[1080,1205,403,429]" italics="true" pageId="12" pageNumber="13">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
, both prominent in dorsal view in
|
||||
<taxonomicName id="4C3ABDD84001B22265DA1EF5FE7FFE6C" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[386,503,438,465]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B22265DA1EF5FE7FFE6C" box="[386,503,438,465]" italics="true" pageId="12" pageNumber="13">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
(A. Milne-Edwards 1873: pl. 18, fig. 6a;
|
||||
<bibRefCitation id="EFABBBAA4001B22267881EF5FA96FE6C" author="Ng P. K. L. & Richer de Forges B." box="[976,1310,438,465]" pageId="12" pageNumber="13" pagination="263 - 276" refId="ref14618" refString="Ng P. K. L. & Richer de Forges B. 1996. The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of New Caledonia, with descriptions of two new genera and two new species. Memoirs of the Queensland Museum 39 (2): 263 - 276. Available from https: // www. biodiversitylibrary. org / part / 50813 # / summary [accessed 27 May 2020]." type="journal article" year="1996">Ng & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 6a, d, not figured in fig. 6c;
|
||||
<bibRefCitation id="EFABBBAA4001B22265BA1E98FCE7FE48" author="Richer de Forges B. & Jamieson B. G. M. & Guinot D. & Tudge C. C." box="[482,879,474,500]" pageId="12" pageNumber="13" pagination="233 - 242" refId="ref15566" refString="Richer de Forges B., Jamieson B. G. M., Guinot D. & Tudge C. C. 1997. Ultrastructure of the spermatozoa of Hymenosomatidae (Crustacea: Brachyura) and the relationships of the family. Marine Biology 130: 233 - 242. https: // doi. org / 10.1007 / s 002270050243" type="journal article" year="1997">Guinot & Richer de Forges 1997</bibRefCitation>
|
||||
: figs 1a, c, 2c), but shorter and not dorsally visible in
|
||||
<taxonomicName id="4C3ABDD84001B22265681EBFFE37FDAB" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[304,447,509,535]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494001B22265681EBFFE37FDAB" box="[304,447,509,535]" italics="true" pageId="12" pageNumber="13">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<bibRefCitation id="EFABBBAA4001B22265961EBFFCC0FDAB" author="Davie P. J. F. & Richer de Forges B." box="[462,840,509,535]" pageId="12" pageNumber="13" pagination="257 - 262" refId="ref12668" refString="Davie P. J. F. & Richer de Forges B. 1996. Two new species of hymenosomatid crabs from New Caledonia. Memoirs of the Queensland Museum 39 (2): 257 - 262." type="journal article" year="1996">Davie & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 1a; not figured in fig. 1b). In contrast, there is only one, smaller knob in
|
||||
<taxonomicName id="4C3ABDD84001B22266401D62FD00FD86" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[536,648,544,570]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B94E1A494001B22266401D62FD00FD86" box="[536,648,544,570]" italics="true" pageId="12" pageNumber="13">Richerius</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324001B22266CA1D63FD77FD87" box="[658,767,545,571]" pageId="12" pageNumber="13" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
(
|
||||
<figureCitation id="1301DADE4001B22267491D62FCE6FD86" box="[785,878,544,570]" captionStart="Fig" captionStartId="6.[189,232,1782,1808]" captionTargetBox="[326,1260,292,1710]" captionTargetId="figure@6.[292,1296,265,1741]" captionTargetPageId="6" captionText="Fig. 1. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Carapace, dorsal view. B. Cephalothorax, ventral view, anterior part, with unfolded antennules, the arrow shows antennule with folded second article. Abbreviations: a1 = antennule; a2 = antenna; e = epistome; p.k. = postorbital knob; r = rostrum; u.a. = antennal urinary article; v.r. = ventral expansion of rostrum; 2/3 = thoracic sternal suture 2/3. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3906758" httpUri="https://zenodo.org/record/3906758/files/figure.png" pageId="12" pageNumber="13">Fig. 1B</figureCitation>
|
||||
). The cheliped palm of
|
||||
<taxonomicName id="4C3ABDD84001B22260C91D63FA95FD86" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[1169,1309,545,570]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494001B22260C91D63FA95FD86" box="[1169,1309,545,570]" italics="true" pageId="12" pageNumber="13">R. marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324001B222617E1D63FE9FFDE2" pageId="12" pageNumber="13" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
is covered with long, flexible setae (
|
||||
<figureCitation id="1301DADE4001B222669C1D01FC80FDE2" box="[708,776,579,606]" captionStart="Fig" captionStartId="10.[189,232,1855,1881]" captionTargetBox="[189,1398,901,1815]" captionTargetId="figure@10.[189,1398,901,1816]" captionTargetPageId="10" captionText="Fig. 5. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Type locality, stream where several specimens of the species were collected. B–D. Views of the holotype in vivo: dorsal (B), ventral (C) and frontal (D)." figureDoi="http://doi.org/10.5281/zenodo.3906766" httpUri="https://zenodo.org/record/3906766/files/figure.png" pageId="12" pageNumber="13">Fig. 5</figureCitation>
|
||||
C–D), instead of numerous spinules (or stiff setae) in
|
||||
<taxonomicName id="4C3ABDD84001B22264E51D25FEBCFD3C" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[189,308,614,641]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B22264E51D25FEBCFD3C" box="[189,308,614,641]" italics="true" pageId="12" pageNumber="13">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1301DADE4001B222651D1D24FE03FD3D" box="[325,395,614,641]" captionStart="Fig" captionStartId="15.[189,232,1921,1947]" captionTargetBox="[189,1398,300,1857]" captionTargetId="figure@15.[189,1398,296,1857]" captionTargetPageId="15" captionText="Fig. 6. Odiomaris pilosus A. Milne-Edwards, 1873, living specimens. A. Collected from the river Hiit (North Province, Hienghène township). B. Collected from the Baie Nord creek (South Province, Mont- Dore township). Credit: Nicolas Charpin – Photographies Vies d’Ô douce." figureDoi="http://doi.org/10.5281/zenodo.3906768" httpUri="https://zenodo.org/record/3906768/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
|
||||
) (A. Milne-Edwards, 1873: pl. 18, fig. 6d;
|
||||
<bibRefCitation id="EFABBBAA4001B22267D01D25FB69FD3C" author="Ng P. K. L. & Richer de Forges B." box="[904,1249,614,641]" pageId="12" pageNumber="13" pagination="263 - 276" refId="ref14618" refString="Ng P. K. L. & Richer de Forges B. 1996. The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of New Caledonia, with descriptions of two new genera and two new species. Memoirs of the Queensland Museum 39 (2): 263 - 276. Available from https: // www. biodiversitylibrary. org / part / 50813 # / summary [accessed 27 May 2020]." type="journal article" year="1996">Ng & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 6f), and sparse, short setae in
|
||||
<taxonomicName id="4C3ABDD84001B22265EA1DC8FDC9FD1F" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[434,577,650,676]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494001B22265EA1DC8FDC9FD1F" box="[434,577,650,676]" italics="true" pageId="12" pageNumber="13">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1301DADE4001B22266171DC8FD1DFD18" box="[591,661,650,676]" captionStart="Fig" captionStartId="18.[189,232,1892,1918]" captionTargetBox="[189,1398,466,1853]" captionTargetId="figure@18.[189,1398,466,1853]" captionTargetPageId="18" captionText="Fig. 8. Odiomaris estuarius Davie & Richer de Forges, 1996, holotype, ♂, New Caledonia, Dumbea estuary, Davie & Richer de Forges leg., 8 Dec. 1993, 4.5 × 4.3 mm (MNHN-IU-2014-11870 = MNHN-B25278). A. Dorsal view. B. Ventral view. Credit: RECOLNAT (ANR-11-INBS-0004) - Noémy MOLLARET – 2017." figureDoi="http://doi.org/10.5281/zenodo.3906772" httpUri="https://zenodo.org/record/3906772/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
|
||||
). In fact, the whole body (including the margins of the rostrum) and legs of
|
||||
<taxonomicName id="4C3ABDD84001B222651A1DEFFE30FD7B" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[322,440,685,711]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B222651A1DEFFE30FD7B" box="[322,440,685,711]" italics="true" pageId="12" pageNumber="13">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
are covered with spiniform, stiff setae that give a bristle appearance (
|
||||
<bibRefCitation id="EFABBBAA4001B22260871DEFFEFEFD56" author="Ng P. K. L. & Richer de Forges B." pageId="12" pageNumber="13" pagination="263 - 276" refId="ref14618" refString="Ng P. K. L. & Richer de Forges B. 1996. The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of New Caledonia, with descriptions of two new genera and two new species. Memoirs of the Queensland Museum 39 (2): 263 - 276. Available from https: // www. biodiversitylibrary. org / part / 50813 # / summary [accessed 27 May 2020]." type="journal article" year="1996">Ng & Richer de Forges 1996</bibRefCitation>
|
||||
: figs 5a–b, 6a, d;
|
||||
<bibRefCitation id="EFABBBAA4001B222661E1D92FC45FD56" author="Richer de Forges B. & Jamieson B. G. M. & Guinot D. & Tudge C. C." box="[582,973,720,747]" pageId="12" pageNumber="13" pagination="233 - 242" refId="ref15566" refString="Richer de Forges B., Jamieson B. G. M., Guinot D. & Tudge C. C. 1997. Ultrastructure of the spermatozoa of Hymenosomatidae (Crustacea: Brachyura) and the relationships of the family. Marine Biology 130: 233 - 242. https: // doi. org / 10.1007 / s 002270050243" type="journal article" year="1997">Guinot & Richer de Forges 1997</bibRefCitation>
|
||||
: fig. 1a), whereas
|
||||
<taxonomicName id="4C3ABDD84001B22260FB1D92FABBFD56" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[1187,1331,720,746]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494001B22260FB1D92FABBFD56" box="[1187,1331,720,746]" italics="true" pageId="12" pageNumber="13">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
has a carapace with soft, short setae and legs with longer setae (
|
||||
<bibRefCitation id="EFABBBAA4001B222673B1DB1FB55FCB1" author="Davie P. J. F. & Richer de Forges B." box="[867,1245,755,782]" pageId="12" pageNumber="13" pagination="257 - 262" refId="ref12668" refString="Davie P. J. F. & Richer de Forges B. 1996. Two new species of hymenosomatid crabs from New Caledonia. Memoirs of the Queensland Museum 39 (2): 257 - 262." type="journal article" year="1996">Davie & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 1a). The male cheliped fingers are practically joining in
|
||||
<taxonomicName id="4C3ABDD84001B22266B41C55FCFEFC8C" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[748,886,791,816]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494001B22266B41C55FCFEFC8C" box="[748,886,791,816]" italics="true" pageId="12" pageNumber="13">R. marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324001B22267261C55FBB9FC8D" box="[894,1073,791,817]" pageId="12" pageNumber="13" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
(
|
||||
<figureCitation id="1301DADE4001B22260191C54FB16FC8D" box="[1089,1182,790,817]" captionStart="Fig" captionStartId="10.[189,232,1855,1881]" captionTargetBox="[189,1398,901,1815]" captionTargetId="figure@10.[189,1398,901,1816]" captionTargetPageId="10" captionText="Fig. 5. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Type locality, stream where several specimens of the species were collected. B–D. Views of the holotype in vivo: dorsal (B), ventral (C) and frontal (D)." figureDoi="http://doi.org/10.5281/zenodo.3906766" httpUri="https://zenodo.org/record/3906766/files/figure.png" pageId="12" pageNumber="13">Fig. 5C</figureCitation>
|
||||
), but with a broad proximal gap in
|
||||
<taxonomicName id="4C3ABDD84001B22265DC1C78FE75FCE8" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[388,509,826,852]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B22265DC1C78FE75FCE8" box="[388,509,826,852]" italics="true" pageId="12" pageNumber="13">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1301DADE4001B22266481C78FDDFFCE8" box="[528,599,825,852]" captionStart="Fig" captionStartId="15.[189,232,1921,1947]" captionTargetBox="[189,1398,300,1857]" captionTargetId="figure@15.[189,1398,296,1857]" captionTargetPageId="15" captionText="Fig. 6. Odiomaris pilosus A. Milne-Edwards, 1873, living specimens. A. Collected from the river Hiit (North Province, Hienghène township). B. Collected from the Baie Nord creek (South Province, Mont- Dore township). Credit: Nicolas Charpin – Photographies Vies d’Ô douce." figureDoi="http://doi.org/10.5281/zenodo.3906768" httpUri="https://zenodo.org/record/3906768/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
|
||||
A–B) (A. Milne-Edwards 1873: 322, pl. 18, fig. 6, 6d, as
|
||||
<taxonomicName id="4C3ABDD84001B22261481C78FE8DFCCB" authorityName="A.Milne-Edwards" authorityYear="1873" class="Malacostraca" family="Hymenosomatidae" genus="Elamene" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosa">
|
||||
<emphasis id="B94E1A494001B22261481C78FE8DFCCB" italics="true" pageId="12" pageNumber="13">Elamene pilosa</emphasis>
|
||||
</taxonomicName>
|
||||
;
|
||||
<bibRefCitation id="EFABBBAA4001B222654C1C1FFDE5FCCB" author="Ng P. K. L. & Richer de Forges B." box="[276,621,861,887]" pageId="12" pageNumber="13" pagination="263 - 276" refId="ref14618" refString="Ng P. K. L. & Richer de Forges B. 1996. The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of New Caledonia, with descriptions of two new genera and two new species. Memoirs of the Queensland Museum 39 (2): 263 - 276. Available from https: // www. biodiversitylibrary. org / part / 50813 # / summary [accessed 27 May 2020]." type="journal article" year="1996">Ng & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 6f); in
|
||||
<taxonomicName id="4C3ABDD84001B22266AF1C1FFC0FFCCB" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[759,903,861,887]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494001B22266AF1C1FFC0FFCCB" box="[759,903,861,887]" italics="true" pageId="12" pageNumber="13">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
the chelae are narrower, with long fingers without marked gap (
|
||||
<figureCitation id="1301DADE4001B22265EF1CC2FE73FC26" box="[439,507,896,922]" captionStart="Fig" captionStartId="18.[189,232,1892,1918]" captionTargetBox="[189,1398,466,1853]" captionTargetId="figure@18.[189,1398,466,1853]" captionTargetPageId="18" captionText="Fig. 8. Odiomaris estuarius Davie & Richer de Forges, 1996, holotype, ♂, New Caledonia, Dumbea estuary, Davie & Richer de Forges leg., 8 Dec. 1993, 4.5 × 4.3 mm (MNHN-IU-2014-11870 = MNHN-B25278). A. Dorsal view. B. Ventral view. Credit: RECOLNAT (ANR-11-INBS-0004) - Noémy MOLLARET – 2017." figureDoi="http://doi.org/10.5281/zenodo.3906772" httpUri="https://zenodo.org/record/3906772/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
|
||||
A–B) (
|
||||
<bibRefCitation id="EFABBBAA4001B22266151CC2FC40FC26" author="Davie P. J. F. & Richer de Forges B." box="[589,968,896,923]" pageId="12" pageNumber="13" pagination="257 - 262" refId="ref12668" refString="Davie P. J. F. & Richer de Forges B. 1996. Two new species of hymenosomatid crabs from New Caledonia. Memoirs of the Queensland Museum 39 (2): 257 - 262." type="journal article" year="1996">Davie & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 1). The grooves on the carapace dorsal surface are very distinct in
|
||||
<taxonomicName id="4C3ABDD84001B222661B1CE6FD31FC01" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[579,697,931,958]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B222661B1CE6FD31FC01" box="[579,697,931,958]" italics="true" pageId="12" pageNumber="13">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
, well marked in
|
||||
<taxonomicName id="4C3ABDD84001B22267221CE6FB8BFC01" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[890,1027,932,957]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494001B22267221CE6FB8BFC01" box="[890,1027,932,957]" italics="true" pageId="12" pageNumber="13">R. marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324001B22260501CE6FB3FFC02" box="[1032,1207,932,958]" pageId="12" pageNumber="13" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
, whereas almost indiscernible in
|
||||
<taxonomicName id="4C3ABDD84001B22265201C85FD8FFC5C" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[376,519,967,993]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494001B22265201C85FD8FFC5C" box="[376,519,967,993]" italics="true" pageId="12" pageNumber="13">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
. The sterno-pleonal cavity is short in
|
||||
<taxonomicName id="4C3ABDD84001B22267E31C84FBA3FC5C" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[955,1067,966,992]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B94E1A494001B22267E31C84FBA3FC5C" box="[955,1067,966,992]" italics="true" pageId="12" pageNumber="13">Richerius</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324001B222606A1C85FB15FC5C" box="[1074,1181,967,993]" pageId="12" pageNumber="13" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
and in
|
||||
<taxonomicName id="4C3ABDD84001B22260AA1C84FAE7FC5C" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[1266,1391,966,992]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B22260AA1C84FAE7FC5C" box="[1266,1391,966,992]" italics="true" pageId="12" pageNumber="13">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
, and accordingly the pleon also. The male pleon is as a narrow triangle, with a long pleotelson in both
|
||||
<taxonomicName id="4C3ABDD84001B22264E51B4FFEB2FB9B" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[189,314,1037,1063]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B22264E51B4FFEB2FB9B" box="[189,314,1037,1063]" italics="true" pageId="12" pageNumber="13">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
species (A. Milne-Edwards 1873: pl. 18, fig. 6b, as
|
||||
<taxonomicName id="4C3ABDD84001B22267C71B4FFBDDFB9B" authorityName="A.Milne-Edwards" authorityYear="1873" box="[927,1109,1037,1063]" class="Malacostraca" family="Hymenosomatidae" genus="Elamene" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosa">
|
||||
<emphasis id="B94E1A494001B22267C71B4FFBDDFB9B" box="[927,1109,1037,1063]" italics="true" pageId="12" pageNumber="13">Elamene pilosa</emphasis>
|
||||
</taxonomicName>
|
||||
;
|
||||
<bibRefCitation id="EFABBBAA4001B222603D1B4FFF74FBF6" author="Ng P. K. L. & Richer de Forges B." pageId="12" pageNumber="13" pagination="263 - 276" refId="ref14618" refString="Ng P. K. L. & Richer de Forges B. 1996. The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of New Caledonia, with descriptions of two new genera and two new species. Memoirs of the Queensland Museum 39 (2): 263 - 276. Available from https: // www. biodiversitylibrary. org / part / 50813 # / summary [accessed 27 May 2020]." type="journal article" year="1996">Ng & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 7a–b;
|
||||
<bibRefCitation id="EFABBBAA4001B22265271B72FC80FBF6" author="Richer de Forges B. & Jamieson B. G. M. & Guinot D. & Tudge C. C." box="[383,776,1072,1099]" pageId="12" pageNumber="13" pagination="233 - 242" refId="ref15566" refString="Richer de Forges B., Jamieson B. G. M., Guinot D. & Tudge C. C. 1997. Ultrastructure of the spermatozoa of Hymenosomatidae (Crustacea: Brachyura) and the relationships of the family. Marine Biology 130: 233 - 242. https: // doi. org / 10.1007 / s 002270050243" type="journal article" year="1997">Guinot & Richer de Forges 1997</bibRefCitation>
|
||||
: fig. 2a:
|
||||
<taxonomicName id="4C3ABDD84001B22267361B72FC6DFBF6" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[878,997,1072,1098]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B22267361B72FC6DFBF6" box="[878,997,1072,1098]" italics="true" pageId="12" pageNumber="13">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
;
|
||||
<bibRefCitation id="EFABBBAA4001B22267AE1B72FAFBFBF6" author="Davie P. J. F. & Richer de Forges B." box="[1014,1395,1072,1099]" pageId="12" pageNumber="13" pagination="257 - 262" refId="ref12668" refString="Davie P. J. F. & Richer de Forges B. 1996. Two new species of hymenosomatid crabs from New Caledonia. Memoirs of the Queensland Museum 39 (2): 257 - 262." type="journal article" year="1996">Davie & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 2c:
|
||||
<taxonomicName id="4C3ABDD84001B222654D1B16FE2CFBD1" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[277,420,1108,1134]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494001B222654D1B16FE2CFBD1" box="[277,420,1108,1134]" italics="true" pageId="12" pageNumber="13">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
), markedly wider and with a much shorter pleotelson in
|
||||
<taxonomicName id="4C3ABDD84001B22260601B16FB49FBD1" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[1080,1217,1108,1133]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494001B22260601B16FB49FBD1" box="[1080,1217,1108,1133]" italics="true" pageId="12" pageNumber="13">R. marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324001B22260901B16FAFFFBD2" box="[1224,1399,1108,1134]" pageId="12" pageNumber="13" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
(
|
||||
<figureCitation id="1301DADE4001B222649E1B34FEAEFB2D" box="[198,294,1142,1169]" captionStart="Fig" captionStartId="7.[189,232,1846,1872]" captionTargetBox="[260,1308,282,1788]" captionTargetId="figure@7.[248,1339,265,1807]" captionTargetPageId="7" captionText="Fig. 2. Richerius marqueti gen. et sp. nov., holotype, ♂, 4.9 × 5.0 mm (MNHN-IU-2014-21500), same collection data as Fig. 1. A. Thoracic sternum. B. Mxp3. C. Sternopleonal cavity with G1s. D. G1, two views. Abbreviations: G1 = gonopod 1; G2 = gonopod 2; i = intercalated platelets; p = pleotelson; p.b. = press-button; s4 = expansion of sternite 4; 1–8 = thoracic sternites 1–8; 2/3–7/8 = thoracic sternal sutures 2/3–7/8. Scale bars: A = 2 mm; B–C = 1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3906760" httpUri="https://zenodo.org/record/3906760/files/figure.png" pageId="12" pageNumber="13">Fig. 2A</figureCitation>
|
||||
). The intercalated platelets, which are completely demarcated and moveable in
|
||||
<taxonomicName id="4C3ABDD84001B22260A31B35FAFEFB2C" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[1275,1398,1142,1169]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B22260A31B35FAFEFB2C" box="[1275,1398,1142,1169]" italics="true" pageId="12" pageNumber="13">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<bibRefCitation id="EFABBBAA4001B222649D1BD8FEE3FB08" author="Guinot D." box="[197,363,1178,1204]" pageId="12" pageNumber="13" pagination="20 - 32" refId="ref13311" refString="Guinot D. 2011 a. Odiomarinae nov. subfam., a new subfamily for two primitive genera of the Hymenosomatidae MacLeay, 1838 (Crustacea, Decapoda, Brachyura). Zootaxa 2732: 20 - 32. https: // doi. org / 10.11646 / zootaxa. 2732.1.2" type="journal article" year="2011">Guinot 2011a</bibRefCitation>
|
||||
: fig. 2), are only partially delineated in
|
||||
<taxonomicName id="4C3ABDD84001B22267651BD8FC4FFB0F" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[829,967,1178,1203]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494001B22267651BD8FC4FFB0F" box="[829,967,1178,1203]" italics="true" pageId="12" pageNumber="13">R. marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324001B22267971BD8FB0BFB08" box="[975,1155,1178,1204]" pageId="12" pageNumber="13" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
, and not detached in
|
||||
<emphasis id="B94E1A494001B22264E51BFFFEDCFB6B" box="[189,340,1213,1239]" italics="true" pageId="12" pageNumber="13">
|
||||
<taxonomicName id="4C3ABDD84001B22264E51BFFFEC7FB6B" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[189,335,1213,1239]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="estuarius">O. estuarius</taxonomicName>
|
||||
.
|
||||
</emphasis>
|
||||
The female pleon is rather similar in
|
||||
<taxonomicName id="4C3ABDD84001B22267521BFFFCF2FB6B" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[778,890,1213,1239]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B94E1A494001B22267521BFFFCF2FB6B" box="[778,890,1213,1239]" italics="true" pageId="12" pageNumber="13">Richerius</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324001B22267D91BFFFC63FB6B" box="[897,1003,1213,1239]" pageId="12" pageNumber="13" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
and
|
||||
<taxonomicName id="4C3ABDD84001B222607B1BFFFB28FB6B" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[1059,1184,1213,1239]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B222607B1BFFFB28FB6B" box="[1059,1184,1213,1239]" italics="true" pageId="12" pageNumber="13">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
, with five somites plus pleotelson. The G1 of
|
||||
<taxonomicName id="4C3ABDD84001B22266541BA2FDF4FB46" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[524,636,1248,1274]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B94E1A494001B22266541BA2FDF4FB46" box="[524,636,1248,1274]" italics="true" pageId="12" pageNumber="13">Richerius</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324001B22266DE1BA3FD7CFB47" box="[646,756,1249,1275]" pageId="12" pageNumber="13" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
(
|
||||
<figureCitation id="1301DADE4001B222675F1BA2FCEEFB46" box="[775,870,1248,1274]" captionStart="Fig" captionStartId="7.[189,232,1846,1872]" captionTargetBox="[260,1308,282,1788]" captionTargetId="figure@7.[248,1339,265,1807]" captionTargetPageId="7" captionText="Fig. 2. Richerius marqueti gen. et sp. nov., holotype, ♂, 4.9 × 5.0 mm (MNHN-IU-2014-21500), same collection data as Fig. 1. A. Thoracic sternum. B. Mxp3. C. Sternopleonal cavity with G1s. D. G1, two views. Abbreviations: G1 = gonopod 1; G2 = gonopod 2; i = intercalated platelets; p = pleotelson; p.b. = press-button; s4 = expansion of sternite 4; 1–8 = thoracic sternites 1–8; 2/3–7/8 = thoracic sternal sutures 2/3–7/8. Scale bars: A = 2 mm; B–C = 1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3906760" httpUri="https://zenodo.org/record/3906760/files/figure.png" pageId="12" pageNumber="13">Fig. 2D</figureCitation>
|
||||
) is curved at the base, otherwise with little curvature, with a terminal portion bearing fine setae and ending in one lobe without corneous process, whereas the G1 of
|
||||
<taxonomicName id="4C3ABDD84001B22265CE1A64FD9BFAFC" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[406,531,1318,1344]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B22265CE1A64FD9BFAFC" box="[406,531,1318,1344]" italics="true" pageId="12" pageNumber="13">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
is characterised by two distinct distal processes, a longer corneous process and a shorter lobular elongation of the stem (
|
||||
<figureCitation id="1301DADE4001B222669F1A08FCACFAD8" box="[711,804,1354,1380]" captionStart="Fig" captionStartId="16.[189,232,1919,1945]" captionTargetBox="[402,1193,1524,1866]" captionTargetId="figure@16.[385,1203,1509,1878]" captionTargetPageId="16" captionText="Fig. 7. Odiomaris pilosus A. Milne-Edwards, 1873, syntype, ♂, New Caledonia, freshwater, B. Balansa leg. (RMNH D 32). A. Ventral anterior part. B. G1 (after Holthuis 1968: fig. 3a–b). Abbreviations: a1 = antennule, with folded second article; a2 = antenna; e = epistome: p.k. = postorbital knobs; r = rostrum." figureDoi="http://doi.org/10.5281/zenodo.3906770" httpUri="https://zenodo.org/record/3906770/files/figure.png" pageId="12" pageNumber="13">Fig. 7B</figureCitation>
|
||||
:
|
||||
<taxonomicName id="4C3ABDD84001B22267771A08FC2DFAD8" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[815,933,1354,1380]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B22267771A08FC2DFAD8" box="[815,933,1354,1380]" italics="true" pageId="12" pageNumber="13">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
) (
|
||||
<bibRefCitation id="EFABBBAA4001B22267E61A08FA9CFADF" author="Ng P. K. L. & Richer de Forges B." box="[958,1300,1353,1380]" pageId="12" pageNumber="13" pagination="263 - 276" refId="ref14618" refString="Ng P. K. L. & Richer de Forges B. 1996. The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of New Caledonia, with descriptions of two new genera and two new species. Memoirs of the Queensland Museum 39 (2): 263 - 276. Available from https: // www. biodiversitylibrary. org / part / 50813 # / summary [accessed 27 May 2020]." type="journal article" year="1996">Ng & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 7c– d:
|
||||
<taxonomicName id="4C3ABDD84001B22264851A2FFEDFFA3B" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[221,343,1389,1415]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B22264851A2FFEDFFA3B" box="[221,343,1389,1415]" italics="true" pageId="12" pageNumber="13">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
;
|
||||
<bibRefCitation id="EFABBBAA4001B22265311A2FFD66FA3B" author="Davie P. J. F. & Richer de Forges B." box="[361,750,1389,1415]" pageId="12" pageNumber="13" pagination="257 - 262" refId="ref12668" refString="Davie P. J. F. & Richer de Forges B. 1996. Two new species of hymenosomatid crabs from New Caledonia. Memoirs of the Queensland Museum 39 (2): 257 - 262." type="journal article" year="1996">Davie & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 2c:
|
||||
<taxonomicName id="4C3ABDD84001B22267011A2FFC63FA3B" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[857,1003,1389,1415]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494001B22267011A2FFC63FA3B" box="[857,1003,1389,1415]" italics="true" pageId="12" pageNumber="13">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
).
|
||||
<taxonomicName id="4C3ABDD84001B222605E1A2FFB10FA3B" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[1030,1176,1389,1415]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494001B222605E1A2FFB10FA3B" box="[1030,1176,1389,1415]" italics="true" pageId="12" pageNumber="13">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
is a much smaller species than
|
||||
<taxonomicName id="4C3ABDD84001B222650B1AD2FE42FA16" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[339,458,1424,1450]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B222650B1AD2FE42FA16" box="[339,458,1424,1450]" italics="true" pageId="12" pageNumber="13">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="4C3ABDD84001B22266521AD3FD1CFA16" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[522,660,1425,1450]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494001B22266521AD3FD1CFA16" box="[522,660,1425,1450]" italics="true" pageId="12" pageNumber="13">R. marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324001B22266C31AD3FCC3FA17" box="[667,843,1425,1451]" pageId="12" pageNumber="13" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
seems to be nearly as small as
|
||||
<taxonomicName id="4C3ABDD84001B22260E11AD2FAC1FA16" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[1209,1353,1424,1450]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494001B22260E11AD2FAC1FA16" box="[1209,1353,1424,1450]" italics="true" pageId="12" pageNumber="13">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4001B22264E51A93FA91F9C4" blockId="12.[189,1399,1489,1657]" pageId="12" pageNumber="13">
|
||||
The main differences to distinguish the two genera
|
||||
<taxonomicName id="4C3ABDD84001B22267421A93FC1FFA57" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[794,919,1489,1515]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B22267421A93FC1FFA57" box="[794,919,1489,1515]" italics="true" pageId="12" pageNumber="13">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C3ABDD84001B22267891A90FBCBFA50" authorityName="Lucas" authorityYear="1980" box="[977,1091,1490,1516]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B22267891A90FBCBFA50" box="[977,1091,1490,1516]" italics="true" pageId="12" pageNumber="13">Amarinus</emphasis>
|
||||
</taxonomicName>
|
||||
stated by
|
||||
<bibRefCitation id="EFABBBAA4001B22260E41A93FE36F9B3" author="Davie P. J. F. & Richer de Forges B." pageId="12" pageNumber="13" pagination="257 - 262" refId="ref12668" refString="Davie P. J. F. & Richer de Forges B. 1996. Two new species of hymenosomatid crabs from New Caledonia. Memoirs of the Queensland Museum 39 (2): 257 - 262." type="journal article" year="1996">Davie & Richer de Forges (1996: 259)</bibRefCitation>
|
||||
were: 1) in
|
||||
<taxonomicName id="4C3ABDD84001B22266121AB7FD4FF9B3" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[586,711,1525,1551]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B22266121AB7FD4FF9B3" box="[586,711,1525,1551]" italics="true" pageId="12" pageNumber="13">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
, the G1 (see
|
||||
<figureCitation id="1301DADE4001B22267381AB7FC30F9B3" box="[864,952,1525,1551]" captionStart="Fig" captionStartId="16.[189,232,1919,1945]" captionTargetBox="[402,1193,1524,1866]" captionTargetId="figure@16.[385,1203,1509,1878]" captionTargetPageId="16" captionText="Fig. 7. Odiomaris pilosus A. Milne-Edwards, 1873, syntype, ♂, New Caledonia, freshwater, B. Balansa leg. (RMNH D 32). A. Ventral anterior part. B. G1 (after Holthuis 1968: fig. 3a–b). Abbreviations: a1 = antennule, with folded second article; a2 = antenna; e = epistome: p.k. = postorbital knobs; r = rostrum." figureDoi="http://doi.org/10.5281/zenodo.3906770" httpUri="https://zenodo.org/record/3906770/files/figure.png" pageId="12" pageNumber="13">Fig. 7B</figureCitation>
|
||||
) more slender, with two distinct distal processes, a longer corneous process and a shorter lobular elongation of the stem than in
|
||||
<taxonomicName id="4C3ABDD84001B2226099195AFABBF98E" authorityName="Lucas" authorityYear="1980" box="[1217,1331,1560,1586]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B2226099195AFABBF98E" box="[1217,1331,1560,1586]" italics="true" pageId="12" pageNumber="13">Amarinus</emphasis>
|
||||
</taxonomicName>
|
||||
; 2) in
|
||||
<taxonomicName id="4C3ABDD84001B22264E51979FEB2F9E9" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[189,314,1595,1621]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B22264E51979FEB2F9E9" box="[189,314,1595,1621]" italics="true" pageId="12" pageNumber="13">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
, the elongated triangular telson of the male pleon is significantly longer than wide at base, whereas in
|
||||
<taxonomicName id="4C3ABDD84001B222651B191DFE3DF9C5" authorityName="Lucas" authorityYear="1980" box="[323,437,1631,1657]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B222651B191DFE3DF9C5" box="[323,437,1631,1657]" italics="true" pageId="12" pageNumber="13">Amarinus</emphasis>
|
||||
</taxonomicName>
|
||||
the telson is more or less rounded and short, being much wider than long.
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4001B22364E519E2FDEBFEF4" blockId="12.[189,1399,1696,2004]" lastBlockId="13.[189,1398,267,328]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
|
||||
The genus
|
||||
<taxonomicName id="4C3ABDD84001B222651D19E2FDDEF906" authority="Lucas, 1980" authorityName="Lucas" authorityYear="1980" box="[325,598,1696,1722]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B222651D19E2FE3FF906" box="[325,439,1696,1722]" italics="true" pageId="12" pageNumber="13">Amarinus</emphasis>
|
||||
<bibRefCitation id="EFABBBAA4001B222659B19E2FDDEF906" author="Lucas J. S." box="[451,598,1696,1722]" pageId="12" pageNumber="13" pagination="148 - 247" refId="ref14163" refString="Lucas J. S. 1980. Spider crabs of the family Hymenosomatidae (Crustacea: Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum 33 (4): 148 - 247. https: // doi. org / 10.3853 / j. 0067 - 1975.33.1980.199" type="journal article" year="1980">Lucas, 1980</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(
|
||||
<typeStatus id="548178F94001B222663119E2FD15F906" box="[617,669,1696,1722]" pageId="12" pageNumber="13">type</typeStatus>
|
||||
species by original designation:
|
||||
<taxonomicName id="4C3ABDD84001B222606B19E2FF71F961" authority="Chilton, 1882" authorityName="Chilton" authorityYear="1882" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="lacustris">
|
||||
<emphasis id="B94E1A494001B222606B19E2FA84F906" box="[1075,1292,1696,1722]" italics="true" pageId="12" pageNumber="13">Elamena lacustris</emphasis>
|
||||
Chilton, 1882
|
||||
</taxonomicName>
|
||||
) is known from more than ten species. They all inhabit low salinity environments, from brackish habitats to pure freshwater waters permanently (streams, lakes, swamps), and have a large distribution (
|
||||
<collectingCountry id="F32D86CB4001B222649D184BFEE0F89F" box="[197,360,1801,1827]" name="New Zealand" pageId="12" pageNumber="13">New Zealand</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="F32D86CB4001B222652D184BFE6CF89F" box="[373,484,1801,1827]" name="Australia" pageId="12" pageNumber="13">Australia</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="F32D86CB4001B22265A9184BFDEEF89F" box="[497,614,1801,1827]" name="Indonesia" pageId="12" pageNumber="13">Indonesia</collectingCountry>
|
||||
, the
|
||||
<collectingCountry id="F32D86CB4001B22266F8184BFCAFF89F" box="[672,807,1801,1827]" name="Philippines" pageId="12" pageNumber="13">Philippines</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="F32D86CB4001B222676D1848FBA9F898" box="[821,1057,1801,1828]" name="Papua New Guinea" pageId="12" pageNumber="13">Papua New Guinea</collectingCountry>
|
||||
). Cases where
|
||||
<taxonomicName id="4C3ABDD84001B222608C1848FACEF898" authorityName="Lucas" authorityYear="1980" box="[1236,1350,1802,1828]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B222608C1848FACEF898" box="[1236,1350,1802,1828]" italics="true" pageId="12" pageNumber="13">Amarinus</emphasis>
|
||||
</taxonomicName>
|
||||
has been reported from
|
||||
<collectingCountry id="F32D86CB4001B22265F2186FFDEDF8FA" box="[426,613,1836,1863]" name="New Caledonia" pageId="12" pageNumber="13">New Caledonia</collectingCountry>
|
||||
, as in
|
||||
<bibRefCitation id="EFABBBAA4001B22266EC186EFC53F8FB" author="Chuang C. T. N. & Ng P. K. L." box="[692,987,1836,1863]" pageId="12" pageNumber="13" pagination="85 - 92" refId="ref12538" refString="Chuang C. T. N. & Ng P. K. L. 1994. The ecology and biology of Southeast Asian false spider crabs (Crustacea: Decapoda: Brachyura: Hymenosomatidae). In: Sasekumar A., Marshall N. & Macintosh D. J. (eds) Ecology and Conservation of Southeast Asian Marine and Freshwater Environments including Wetlands. Hydrobiologia 285: 85 - 92. https: // doi. org / 10.1007 / BF 00005656" type="journal article" year="1994">Chuang & Ng (1994: 87</bibRefCitation>
|
||||
, 90, table 1, under
|
||||
<taxonomicName id="4C3ABDD84001B222609C186FFAB1F8FA" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[1220,1337,1836,1863]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B222609C186FFAB1F8FA" box="[1220,1337,1836,1863]" italics="true" pageId="12" pageNumber="13">A. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
) and erroneously as in
|
||||
<bibRefCitation id="EFABBBAA4001B22265D51812FDE7F8D6" author="Guinot D." box="[397,623,1872,1898]" pageId="12" pageNumber="13" pagination="20 - 32" refId="ref13311" refString="Guinot D. 2011 a. Odiomarinae nov. subfam., a new subfamily for two primitive genera of the Hymenosomatidae MacLeay, 1838 (Crustacea, Decapoda, Brachyura). Zootaxa 2732: 20 - 32. https: // doi. org / 10.11646 / zootaxa. 2732.1.2" type="journal article" year="2011">Guinot (2011a: 23)</bibRefCitation>
|
||||
, are attributable to the fact that the species
|
||||
<taxonomicName id="4C3ABDD84001B22260331812FB36F8D6" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[1131,1214,1872,1898]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494001B22260331812FB36F8D6" box="[1131,1214,1872,1898]" italics="true" pageId="12" pageNumber="13">pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
was previously associated with the genus
|
||||
<taxonomicName id="4C3ABDD84001B22265AB1831FDEDF831" authorityName="Lucas" authorityYear="1980" box="[499,613,1907,1933]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B22265AB1831FDEDF831" box="[499,613,1907,1933]" italics="true" pageId="12" pageNumber="13">Amarinus</emphasis>
|
||||
</taxonomicName>
|
||||
(see
|
||||
<bibRefCitation id="EFABBBAA4001B22266FD1831FCA7F831" author="Lucas J. S." box="[677,815,1907,1933]" pageId="12" pageNumber="13" pagination="148 - 247" refId="ref14163" refString="Lucas J. S. 1980. Spider crabs of the family Hymenosomatidae (Crustacea: Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum 33 (4): 148 - 247. https: // doi. org / 10.3853 / j. 0067 - 1975.33.1980.199" type="journal article" year="1980">Lucas 1980</bibRefCitation>
|
||||
) until
|
||||
<bibRefCitation id="EFABBBAA4001B22267261831FB61F832" author="Ng P. K. L. & Richer de Forges B." box="[894,1257,1907,1934]" pageId="12" pageNumber="13" pagination="263 - 276" refId="ref14618" refString="Ng P. K. L. & Richer de Forges B. 1996. The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of New Caledonia, with descriptions of two new genera and two new species. Memoirs of the Queensland Museum 39 (2): 263 - 276. Available from https: // www. biodiversitylibrary. org / part / 50813 # / summary [accessed 27 May 2020]." type="journal article" year="1996">Ng & Richer de Forges (1996)</bibRefCitation>
|
||||
made it the
|
||||
<typeStatus id="548178F94001B22264E518D5FF78F80D" box="[189,240,1943,1969]" pageId="12" pageNumber="13">type</typeStatus>
|
||||
species of their new genus
|
||||
<taxonomicName id="4C3ABDD84001B222666418D4FD31F80C" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[572,697,1942,1968]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B222666418D4FD31F80C" box="[572,697,1942,1968]" italics="true" pageId="12" pageNumber="13">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
. So far, no species of
|
||||
<emphasis id="B94E1A494001B222679D18D5FBA4F80D" box="[965,1068,1943,1969]" italics="true" pageId="12" pageNumber="13">Amarinu</emphasis>
|
||||
s has actually been reported in
|
||||
<collectingCountry id="F32D86CB4001B222648418F8FE1FF86F" box="[220,407,1977,2004]" name="New Caledonia" pageId="12" pageNumber="13">New Caledonia</collectingCountry>
|
||||
. Suspecting that our new species might belong to the genus
|
||||
<taxonomicName id="4C3ABDD84001B222603918F8FB5BF868" authorityName="Lucas" authorityYear="1980" box="[1121,1235,1978,2004]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494001B222603918F8FB5BF868" box="[1121,1235,1978,2004]" italics="true" pageId="12" pageNumber="13">Amarinus</emphasis>
|
||||
</taxonomicName>
|
||||
, we therefore carefully compared
|
||||
<taxonomicName id="4C3ABDD84000B22365F21E49FD92FE99" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[426,538,267,293]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||||
<emphasis id="B94E1A494000B22365F21E49FD92FE99" box="[426,538,267,293]" italics="true" pageId="13" pageNumber="14">Richerius</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324000B223667B1E4EFD07FE9A" box="[547,655,268,294]" pageId="13" pageNumber="14" rank="genus">gen. nov.</taxonomicNameLabel>
|
||||
to
|
||||
<taxonomicName id="4C3ABDD84000B22366EF1E4EFCA1FE9A" authorityName="Lucas" authorityYear="1980" box="[695,809,268,294]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494000B22366EF1E4EFCA1FE9A" box="[695,809,268,294]" italics="true" pageId="13" pageNumber="14">Amarinus</emphasis>
|
||||
</taxonomicName>
|
||||
, and in particular to its
|
||||
<typeStatus id="548178F94000B22360101E4EFBF3FE9A" box="[1096,1147,268,294]" pageId="13" pageNumber="14">type</typeStatus>
|
||||
species,
|
||||
<taxonomicName id="4C3ABDD84000B22360B01E4EFAE7FE99" authorityName="Chilton" authorityYear="1882" box="[1256,1391,267,294]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lacustris">
|
||||
<emphasis id="B94E1A494000B22360B01E4EFAE7FE99" box="[1256,1391,267,294]" italics="true" pageId="13" pageNumber="14">A. lacustris</emphasis>
|
||||
</taxonomicName>
|
||||
, another freshwater hymenosomatid.
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4000B22364E51E32FAC3FDAB" blockId="13.[189,1399,368,536]" pageId="13" pageNumber="14">
|
||||
<taxonomicName id="4C3ABDD84000B22364E51E32FE17FE36" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[189,415,368,394]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494000B22364E51E32FE17FE36" box="[189,415,368,394]" italics="true" pageId="13" pageNumber="14">Richerius marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324000B22365F21E33FDECFE37" box="[426,612,369,395]" pageId="13" pageNumber="14" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
can be distinguished from
|
||||
<taxonomicName id="4C3ABDD84000B22367E91E32FB19FE36" baseAuthorityName="Chilton" baseAuthorityYear="1882" box="[945,1169,368,394]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lacustris">
|
||||
<emphasis id="B94E1A494000B22367E91E32FB19FE36" box="[945,1169,368,394]" italics="true" pageId="13" pageNumber="14">Amarinus lacustris</emphasis>
|
||||
</taxonomicName>
|
||||
by: the wide male pleon, with prominent marks corresponding to intercalated platelets (
|
||||
<figureCitation id="1301DADE4000B223678C1ED1FBA5FE11" box="[980,1069,403,429]" captionStart="Fig" captionStartId="7.[189,232,1846,1872]" captionTargetBox="[260,1308,282,1788]" captionTargetId="figure@7.[248,1339,265,1807]" captionTargetPageId="7" captionText="Fig. 2. Richerius marqueti gen. et sp. nov., holotype, ♂, 4.9 × 5.0 mm (MNHN-IU-2014-21500), same collection data as Fig. 1. A. Thoracic sternum. B. Mxp3. C. Sternopleonal cavity with G1s. D. G1, two views. Abbreviations: G1 = gonopod 1; G2 = gonopod 2; i = intercalated platelets; p = pleotelson; p.b. = press-button; s4 = expansion of sternite 4; 1–8 = thoracic sternites 1–8; 2/3–7/8 = thoracic sternal sutures 2/3–7/8. Scale bars: A = 2 mm; B–C = 1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3906760" httpUri="https://zenodo.org/record/3906760/files/figure.png" pageId="13" pageNumber="14">Fig. 2A</figureCitation>
|
||||
) (versus narrow and without intercalated platelets in
|
||||
<taxonomicName id="4C3ABDD84000B22365841EF5FDECFE6C" baseAuthorityName="Chilton" baseAuthorityYear="1882" box="[476,612,438,465]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lacustris">
|
||||
<emphasis id="B94E1A494000B22365841EF5FDECFE6C" box="[476,612,438,465]" italics="true" pageId="13" pageNumber="14">A. lacustris</emphasis>
|
||||
</taxonomicName>
|
||||
, see Melrose 1975: fig.
|
||||
<quantity id="4CC26BBE4000B22367D71EF5FC36FE6D" box="[911,958,439,465]" metricMagnitude="-2" metricUnit="kg" metricValue="4.2" pageId="13" pageNumber="14" unit="g" value="42.0">42g</quantity>
|
||||
;
|
||||
<bibRefCitation id="EFABBBAA4000B22367951EF5FBD4FE6D" author="Lucas J. S." box="[973,1116,439,465]" pageId="13" pageNumber="14" pagination="148 - 247" refId="ref14163" refString="Lucas J. S. 1980. Spider crabs of the family Hymenosomatidae (Crustacea: Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum 33 (4): 148 - 247. https: // doi. org / 10.3853 / j. 0067 - 1975.33.1980.199" type="journal article" year="1980">Lucas 1980</bibRefCitation>
|
||||
: fig. 7b); the G1 rather narrow and with one distal lobe without corneous process (
|
||||
<figureCitation id="1301DADE4000B22367351E98FC40FE48" box="[877,968,474,500]" captionStart="Fig" captionStartId="7.[189,232,1846,1872]" captionTargetBox="[260,1308,282,1788]" captionTargetId="figure@7.[248,1339,265,1807]" captionTargetPageId="7" captionText="Fig. 2. Richerius marqueti gen. et sp. nov., holotype, ♂, 4.9 × 5.0 mm (MNHN-IU-2014-21500), same collection data as Fig. 1. A. Thoracic sternum. B. Mxp3. C. Sternopleonal cavity with G1s. D. G1, two views. Abbreviations: G1 = gonopod 1; G2 = gonopod 2; i = intercalated platelets; p = pleotelson; p.b. = press-button; s4 = expansion of sternite 4; 1–8 = thoracic sternites 1–8; 2/3–7/8 = thoracic sternal sutures 2/3–7/8. Scale bars: A = 2 mm; B–C = 1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3906760" httpUri="https://zenodo.org/record/3906760/files/figure.png" pageId="13" pageNumber="14">Fig. 2D</figureCitation>
|
||||
) (versus stout and without lobes, see Melrose 1975: fig. 42h–i;
|
||||
<bibRefCitation id="EFABBBAA4000B22365B31EBFFDFDFDAB" author="Lucas J. S." box="[491,629,509,535]" pageId="13" pageNumber="14" pagination="148 - 247" refId="ref14163" refString="Lucas J. S. 1980. Spider crabs of the family Hymenosomatidae (Crustacea: Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum 33 (4): 148 - 247. https: // doi. org / 10.3853 / j. 0067 - 1975.33.1980.199" type="journal article" year="1980">Lucas 1980</bibRefCitation>
|
||||
: fig. 10e); and by the arrangement of antennules and antennae.
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4000B22364E51D7DFCA0FB13" blockId="13.[189,1399,574,1199]" pageId="13" pageNumber="14">
|
||||
In providing a key of the Southeast Asian hymenosomatids,
|
||||
<bibRefCitation id="EFABBBAA4000B22367FA1D7DFB57FDE5" author="Ng P. K. L. & Chuang C. T. N." box="[930,1247,574,601]" pageId="13" pageNumber="14" pagination="1 - 82" refId="ref14547" refString="Ng P. K. L. & Chuang C. T. N. 1996 The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of Southeast Asia, with notes on other species. Raffles Bulletin of Zoology Supplement 3: 1 - 82. Available from https: // decapoda. nhm. org / pdfs / 12225 / 12225. pdf [accessed 27 May 2020]." type="journal article" year="1996">Ng & Chuang (1996: 3–5</bibRefCitation>
|
||||
, 6–12) have shown the presence of several groups of species within
|
||||
<taxonomicName id="4C3ABDD84000B22367001D20FC42FDC0" authorityName="Lucas" authorityYear="1980" box="[856,970,610,636]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494000B22367001D20FC42FDC0" box="[856,970,610,636]" italics="true" pageId="13" pageNumber="14">Amarinus</emphasis>
|
||||
</taxonomicName>
|
||||
, suggesting that the genus could be paraphyletic. Today, another problem arises, especially regarding the antennular morphology. Melrose (1975: 84, 87, figs 41–42, as
|
||||
<taxonomicName id="4C3ABDD84000B22366761DEAFCBFFD7E" box="[558,823,680,706]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lacustris">
|
||||
<emphasis id="B94E1A494000B22366761DEAFCBFFD7E" box="[558,823,680,706]" italics="true" pageId="13" pageNumber="14">Halicarcinus lacustris</emphasis>
|
||||
</taxonomicName>
|
||||
), who has thoroughly studied the
|
||||
<typeStatus id="548178F94000B22360BB1DEAFA9EFD7E" box="[1251,1302,680,706]" pageId="13" pageNumber="14">type</typeStatus>
|
||||
species
|
||||
<taxonomicName id="4C3ABDD84000B22364E51D89FECDFD59" box="[189,325,715,741]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lacustris">
|
||||
<emphasis id="B94E1A494000B22364E51D89FECDFD59" box="[189,325,715,741]" italics="true" pageId="13" pageNumber="14">A. lacustris</emphasis>
|
||||
</taxonomicName>
|
||||
, confined to lakes and non-tidal rivers of
|
||||
<collectingCountry id="F32D86CB4000B22367191D8EFC68FD59" box="[833,992,715,742]" name="New Zealand" pageId="13" pageNumber="14">New Zealand</collectingCountry>
|
||||
and southeastern
|
||||
<collectingCountry id="F32D86CB4000B22360E11D89FAA0FD59" box="[1209,1320,715,741]" name="Australia" pageId="13" pageNumber="14">Australia</collectingCountry>
|
||||
, states that the antennules are “small, not visible dorsally when folded”. Her figure 41d actually seems to show a folded antennule, only with the broad basal article and the second cylindrical article, without the short, supposedly folded flagellum being seen [the frontal view of fig. 41e in Melrose (1975) shows a complete, unfolded antennule]. Our examination of
|
||||
<taxonomicName id="4C3ABDD84000B223677B1C1AFB8AFCCE" baseAuthorityName="Chilton" baseAuthorityYear="1882" box="[803,1026,856,882]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lacustris">
|
||||
<emphasis id="B94E1A494000B223677B1C1AFB8AFCCE" box="[803,1026,856,882]" italics="true" pageId="13" pageNumber="14">Amarinus lacustris</emphasis>
|
||||
</taxonomicName>
|
||||
, on the other hand, reveals that both antennules and antennae are inserted very closely together at their bases and remain parallel, with both flagella visible dorsally, as represented by
|
||||
<bibRefCitation id="EFABBBAA4000B22366871CDDFCFCFC04" author="Lucas J. S." box="[735,884,926,953]" pageId="13" pageNumber="14" pagination="148 - 247" refId="ref14163" refString="Lucas J. S. 1980. Spider crabs of the family Hymenosomatidae (Crustacea: Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum 33 (4): 148 - 247. https: // doi. org / 10.3853 / j. 0067 - 1975.33.1980.199" type="journal article" year="1980">Lucas (1980</bibRefCitation>
|
||||
: fig. 1d) for a generalised hymenosomatid or by Melrose (1975: fig. 43c) for
|
||||
<taxonomicName id="4C3ABDD84000B223667E1C83FC2DFC60" authority="Melrose, 1975" authorityName="Melrose" authorityYear="1975" box="[550,933,961,988]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tongi">
|
||||
<emphasis id="B94E1A494000B223667E1C83FD7CFC67" box="[550,756,961,987]" italics="true" pageId="13" pageNumber="14">Halicarinus tongi</emphasis>
|
||||
Melrose, 1975
|
||||
</taxonomicName>
|
||||
. In fact, if it is possible for the antennule of
|
||||
<taxonomicName id="4C3ABDD84000B22364861CA7FEEDFC43" baseAuthorityName="Chilton" baseAuthorityYear="1882" box="[222,357,997,1023]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lacustris">
|
||||
<emphasis id="B94E1A494000B22364861CA7FEEDFC43" box="[222,357,997,1023]" italics="true" pageId="13" pageNumber="14">A. lacustris</emphasis>
|
||||
</taxonomicName>
|
||||
to fold itself down [but not obliquely along the rostrum as in
|
||||
<taxonomicName id="4C3ABDD84000B22360141CA7FB41FC43" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[1100,1225,997,1023]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494000B22360141CA7FB41FC43" box="[1100,1225,997,1023]" italics="true" pageId="13" pageNumber="14">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1301DADE4000B22360831CA7FAB2FC43" box="[1243,1338,997,1023]" captionStart="Fig" captionStartId="16.[189,232,1919,1945]" captionTargetBox="[402,1193,1524,1866]" captionTargetId="figure@16.[385,1203,1509,1878]" captionTargetPageId="16" captionText="Fig. 7. Odiomaris pilosus A. Milne-Edwards, 1873, syntype, ♂, New Caledonia, freshwater, B. Balansa leg. (RMNH D 32). A. Ventral anterior part. B. G1 (after Holthuis 1968: fig. 3a–b). Abbreviations: a1 = antennule, with folded second article; a2 = antenna; e = epistome: p.k. = postorbital knobs; r = rostrum." figureDoi="http://doi.org/10.5281/zenodo.3906770" httpUri="https://zenodo.org/record/3906770/files/figure.png" pageId="13" pageNumber="14">Fig. 7A</figureCitation>
|
||||
), see A. Milne-Edwards 1873: pl. 18, fig. 6a], this does not correspond to the resting position. In the seven specimens of
|
||||
<taxonomicName id="4C3ABDD84000B223653C1B69FE65FBF9" baseAuthorityName="Chilton" baseAuthorityYear="1882" box="[356,493,1067,1093]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lacustris">
|
||||
<emphasis id="B94E1A494000B223653C1B69FE65FBF9" box="[356,493,1067,1093]" italics="true" pageId="13" pageNumber="14">A. lacustris</emphasis>
|
||||
</taxonomicName>
|
||||
examined, the antennule and antenna are very closely inserted and remain parallel, both being stretched forward; the unfolded flagellum is visible dorsally in all individuals. In any event, the disposition of the antennule and antenna differs from that
|
||||
<taxonomicName id="4C3ABDD84000B223605A1B33FBF7FB37" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[1026,1151,1137,1163]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494000B223605A1B33FBF7FB37" box="[1026,1151,1137,1163]" italics="true" pageId="13" pageNumber="14">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="1301DADE4000B22360D71B33FB65FB30" box="[1167,1261,1137,1164]" captionStart="Fig" captionStartId="16.[189,232,1919,1945]" captionTargetBox="[402,1193,1524,1866]" captionTargetId="figure@16.[385,1203,1509,1878]" captionTargetPageId="16" captionText="Fig. 7. Odiomaris pilosus A. Milne-Edwards, 1873, syntype, ♂, New Caledonia, freshwater, B. Balansa leg. (RMNH D 32). A. Ventral anterior part. B. G1 (after Holthuis 1968: fig. 3a–b). Abbreviations: a1 = antennule, with folded second article; a2 = antenna; e = epistome: p.k. = postorbital knobs; r = rostrum." figureDoi="http://doi.org/10.5281/zenodo.3906770" httpUri="https://zenodo.org/record/3906770/files/figure.png" pageId="13" pageNumber="14">Fig. 7A</figureCitation>
|
||||
:
|
||||
<taxonomicName id="4C3ABDD84000B22360AF1B30FAE5FB37" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[1271,1389,1137,1164]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494000B22360AF1B30FAE5FB37" box="[1271,1389,1137,1164]" italics="true" pageId="13" pageNumber="14">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
) (
|
||||
<bibRefCitation id="EFABBBAA4000B223649D1BD7FD96FB13" author="Ng P. K. L. & Richer de Forges B." box="[197,542,1173,1199]" pageId="13" pageNumber="14" pagination="263 - 276" refId="ref14618" refString="Ng P. K. L. & Richer de Forges B. 1996. The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of New Caledonia, with descriptions of two new genera and two new species. Memoirs of the Queensland Museum 39 (2): 263 - 276. Available from https: // www. biodiversitylibrary. org / part / 50813 # / summary [accessed 27 May 2020]." type="journal article" year="1996">Ng & Richer de Forges 1996</bibRefCitation>
|
||||
: fig. 6c–d:
|
||||
<taxonomicName id="4C3ABDD84000B22366F81BD7FC9EFB13" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[672,790,1173,1199]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494000B22366F81BD7FC9EFB13" box="[672,790,1173,1199]" italics="true" pageId="13" pageNumber="14">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4000B22364E51B94FECAFA5B" blockId="13.[189,1399,1238,1511]" pageId="13" pageNumber="14">
|
||||
A picture of the cephalic region in ventral view of
|
||||
<taxonomicName id="4C3ABDD84000B223677F1B95FB9BFB4C" authorityName="Holthuis" authorityYear="1968" box="[807,1043,1238,1265]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="angelicus">
|
||||
<emphasis id="B94E1A494000B223677F1B95FB9BFB4C" box="[807,1043,1238,1265]" italics="true" pageId="13" pageNumber="14">Amarinus angelicus</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<bibRefCitation id="EFABBBAA4000B223607E1B94FA99FB4C" author="Holthuis L. B." box="[1062,1297,1238,1264]" pageId="13" pageNumber="14" pagination="109 - 121" refId="ref13478" refString="Holthuis L. B. 1968. On Hymenosomatidae (Crustacea Decapoda Brachyura) from fresh water, with the description of a new species. Beaufortia 15 (195): 109 - 121. Available from https: // www. repository. naturalis. nl / record / 504776 [accessed 27 May 2020]." type="journal article" year="1968">Holthuis 1968: 114</bibRefCitation>
|
||||
, fig. 2b) shows an antennule “entirely hidden below the rostrum”, obliquely folded, which does not seem to correspond to the condition of
|
||||
<taxonomicName id="4C3ABDD84000B223666C1A5FFD34FA8A" baseAuthorityName="Chilton" baseAuthorityYear="1882" box="[564,700,1308,1335]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="lacustris">
|
||||
<emphasis id="B94E1A494000B223666C1A5FFD34FA8A" box="[564,700,1308,1335]" italics="true" pageId="13" pageNumber="14">A. lacustris</emphasis>
|
||||
</taxonomicName>
|
||||
. This character deserves to be reviewed for all species of
|
||||
<taxonomicName id="4C3ABDD84000B22364E51A02FEA7FAE6" authorityName="Lucas" authorityYear="1980" box="[189,303,1344,1370]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494000B22364E51A02FEA7FAE6" box="[189,303,1344,1370]" italics="true" pageId="13" pageNumber="14">Amarinus</emphasis>
|
||||
</taxonomicName>
|
||||
, and the allocation of
|
||||
<taxonomicName id="4C3ABDD84000B223666D1A02FD2FFAE6" authorityName="Lucas" authorityYear="1980" box="[565,679,1344,1370]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494000B223666D1A02FD2FFAE6" box="[565,679,1344,1370]" italics="true" pageId="13" pageNumber="14">Amarinus</emphasis>
|
||||
</taxonomicName>
|
||||
(including the
|
||||
<typeStatus id="548178F94000B22367001A02FC03FAE6" box="[856,907,1344,1370]" pageId="13" pageNumber="14">type</typeStatus>
|
||||
species and other included species) to the
|
||||
<taxonomicName id="4C3ABDD84000B22364E51A21FEDEFAC1" authorityName="Guinot" authorityYear="2011" box="[189,342,1379,1405]" class="Malacostraca" family="Hymenosomatidae" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subFamily" subFamily="Odiomarinae">Odiomarinae</taxonomicName>
|
||||
needs further investigation. In the new species described here,
|
||||
<taxonomicName id="4C3ABDD84000B22360001A21FAB3FAC1" authority="Isyeb, 2020" authorityName="Isyeb" authorityYear="2020" box="[1112,1339,1379,1405]" class="Malacostraca" family="Hymenosomatidae" genus="Richerius" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="marqueti" status="gen. et sp. nov.">
|
||||
<emphasis id="B94E1A494000B22360001A21FAB3FAC1" box="[1112,1339,1379,1405]" italics="true" pageId="13" pageNumber="14">Richerius marqueti</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A27DA7324000B223611D1A21FEB1FA1D" pageId="13" pageNumber="14" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||||
(
|
||||
<figureCitation id="1301DADE4000B22365131AC4FE2FFA1C" box="[331,423,1414,1440]" captionStart="Fig" captionStartId="6.[189,232,1782,1808]" captionTargetBox="[326,1260,292,1710]" captionTargetId="figure@6.[292,1296,265,1741]" captionTargetPageId="6" captionText="Fig. 1. Richerius marqueti gen. et sp. nov., holotype, ♂, New Caledonia, South Province, 21º26.326ʹ S, 165º31.909ʹ E, Pouéo River, tributary of the Néra, Bouïrou village, Bourail township, 180 m a.s.l., Valentin de Mazancourt and Gérard Marquet leg., 28 Sep. 2016, 4.9 × 5.0 mm (MNHN-IU-2014-21500). A. Carapace, dorsal view. B. Cephalothorax, ventral view, anterior part, with unfolded antennules, the arrow shows antennule with folded second article. Abbreviations: a1 = antennule; a2 = antenna; e = epistome; p.k. = postorbital knob; r = rostrum; u.a. = antennal urinary article; v.r. = ventral expansion of rostrum; 2/3 = thoracic sternal suture 2/3. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3906758" httpUri="https://zenodo.org/record/3906758/files/figure.png" pageId="13" pageNumber="14">Fig. 1B</figureCitation>
|
||||
), the antennule and antenna are separated at least at their insertion site, and only the antennal flagellum is visible dorsally; thus it shares an arrangement substantially similar to that of
|
||||
<taxonomicName id="4C3ABDD84000B22364E51A8EFEB2FA5A" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[189,314,1484,1510]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494000B22364E51A8EFEB2FA5A" box="[189,314,1484,1510]" italics="true" pageId="13" pageNumber="14">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8B85C65B4000B22364E5194CFB42F8FE" blockId="13.[189,1399,1550,1858]" pageId="13" pageNumber="14">
|
||||
It was the study of spermatozoa carried out first in the two species of
|
||||
<taxonomicName id="4C3ABDD84000B223606A194CFB27F994" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[1074,1199,1550,1576]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B94E1A494000B223606A194CFB27F994" box="[1074,1199,1550,1576]" italics="true" pageId="13" pageNumber="14">Odiomaris</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C3ABDD84000B223609C194CFAC8F994" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[1220,1344,1550,1576]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pilosus">
|
||||
<emphasis id="B94E1A494000B223609C194CFAC8F994" box="[1220,1344,1550,1576]" italics="true" pageId="13" pageNumber="14">O. pilosus</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C3ABDD84000B22364E51970FEC4F9F7" authorityName="Davie & Richer de Forges" authorityYear="1996" box="[189,332,1586,1612]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="estuarius">
|
||||
<emphasis id="B94E1A494000B22364E51970FEC4F9F7" box="[189,332,1586,1612]" italics="true" pageId="13" pageNumber="14">O. estuarius</emphasis>
|
||||
</taxonomicName>
|
||||
, by
|
||||
<bibRefCitation id="EFABBBAA4000B22365261973FD53F9F7" author="Richer de Forges B. & Jamieson B. G. M. & Guinot D. & Tudge C. C." box="[382,731,1585,1612]" pageId="13" pageNumber="14" pagination="233 - 242" refId="ref15566" refString="Richer de Forges B., Jamieson B. G. M., Guinot D. & Tudge C. C. 1997. Ultrastructure of the spermatozoa of Hymenosomatidae (Crustacea: Brachyura) and the relationships of the family. Marine Biology 130: 233 - 242. https: // doi. org / 10.1007 / s 002270050243" type="journal article" year="1997">
|
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Richer de Forges
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<emphasis id="B94E1A494000B22366141970FD0EF9F7" box="[588,646,1585,1611]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
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(1997)
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</bibRefCitation>
|
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and later in
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||||
<taxonomicName id="4C3ABDD84000B22367331973FB94F9F7" authorityName="Ng & Richer de Forges" authorityYear="1996" box="[875,1052,1585,1611]" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="vesca">
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<emphasis id="B94E1A494000B22367331973FB94F9F7" box="[875,1052,1585,1611]" italics="true" pageId="13" pageNumber="14">Elamena vesca</emphasis>
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</taxonomicName>
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by
|
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<bibRefCitation id="EFABBBAA4000B223601F1973FAFEF9F7" author="Jamieson B. G. M. & Tudge C. C." box="[1095,1398,1585,1611]" pageId="13" pageNumber="14" pagination="1 - 95" refId="ref13652" refString="Jamieson B. G. M. & Tudge C. C. 2000. Crustacea-Decapoda. In: Adiyodi K. G. & Adiyodi R. G. (ser. eds), Jamieson B. G. M. (vol. ed.) Reproductive Biology of Invertebrates. Volume IX, Part C. Progress in Male Gamete Ultrastructure and Phylogeny. John Wiley and Sons. Chichester: 1 - 95." type="book chapter" year="2000">Jamieson & Tudge (2000)</bibRefCitation>
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that showed the very particular nature of the hymenosomatid sperm within the
|
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<taxonomicName id="4C3ABDD84000B22360041916FB52F9D2" authorityName="Latreille" authorityYear="1802" box="[1116,1242,1620,1646]" infraOrder="Brachyura" pageId="13" pageNumber="3" rank="infraOrder">Brachyura</taxonomicName>
|
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. Indeed, they are distinguished by at least nine major characteristics from those of all the other
|
||||
<taxonomicName id="4C3ABDD84000B223603C1935FB56F92D" authorityName="Latreille" authorityYear="1802" box="[1124,1246,1655,1681]" infraOrder="Brachyura" pageId="13" pageNumber="3" rank="infraOrder">Brachyura</taxonomicName>
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taxa studied, particularly the groups with which they have been associated, the Majoidea Samouelle, 1819 and the Thoracotremata Guinot, 1977. The ‘hymenosomatid-type of spermatozoon’ is unique within
|
||||
<taxonomicName id="4C3ABDD84000B22360A419FCFAFEF964" authorityName="Latreille" authorityYear="1802" box="[1276,1398,1726,1752]" infraOrder="Brachyura" pageId="13" pageNumber="3" rank="infraOrder">Brachyura</taxonomicName>
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(
|
||||
<bibRefCitation id="EFABBBAA4000B223649D19A3FE1BF947" author="Tudge C. & Scheltinga D. M. & Jamieson B. G. M. & Guinot D. & Richer de Forges B." box="[197,403,1761,1787]" pageId="13" pageNumber="14" pagination="1 - 20" refId="ref15940" refString="Tudge C. C, Scheltinga D. M., Jamieson B. G. M., Guinot D. & Richer de Forges B. 2014. Comparative ultrastructure of the spermatozoa of the Majoidea (Crustacea, Decapoda, Brachyura) with new data on six species in five genera. Acta Zoologica 95: 1 - 20. https: // doi. org / 10.1111 / azo. 12005" type="journal article" year="2014">
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Tudge
|
||||
<emphasis id="B94E1A494000B223654E19A0FEC6F947" box="[278,334,1761,1787]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
|
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2014
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</bibRefCitation>
|
||||
). The highly developed projection of the acrosome from the nucleus in hymenosomatid spermatozoon recalls the totally emergent acrosome of Podotremata Guinot, 1977 and may represent the plesiomorphic condition in the Eubrachyura Saint Laurent, 1980 (
|
||||
<bibRefCitation id="EFABBBAA4000B22367981865FBECF8FE" author="Guinot D." box="[960,1124,1831,1858]" pageId="13" pageNumber="14" pagination="20 - 32" refId="ref13311" refString="Guinot D. 2011 a. Odiomarinae nov. subfam., a new subfamily for two primitive genera of the Hymenosomatidae MacLeay, 1838 (Crustacea, Decapoda, Brachyura). Zootaxa 2732: 20 - 32. https: // doi. org / 10.11646 / zootaxa. 2732.1.2" type="journal article" year="2011">Guinot 2011a</bibRefCitation>
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,
|
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<bibRefCitation id="EFABBBAA4000B22360281865FB35F8FE" author="Guinot D." box="[1136,1213,1831,1858]" pageId="13" pageNumber="14" pagination="40 - 52" refId="ref13363" refString="Guinot D. 2011 b. The position of the Hymenosomatidae MacLeay, 1838, within the Brachyura (Crustacea, Decapoda). Zootaxa 2890: 40 - 52. https: // doi. org / 10.11646 / zootaxa. 2890.1.4" type="journal article" year="2011">2011b</bibRefCitation>
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).
|
||||
</paragraph>
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</subSubSection>
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</treatment>
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|
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@ -1,66 +1,66 @@
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|
||||
<treatment id="A13C03190B32FFB6FD93FBACB239FC10" ID-DOI="http://doi.org/10.5281/zenodo.4328677" ID-GBIF-Taxon="164870717" ID-Zenodo-Dep="4328677" LSID="urn:lsid:plazi:treatment:A13C03190B32FFB6FD93FBACB239FC10" httpUri="http://treatment.plazi.org/id/A13C03190B32FFB6FD93FBACB239FC10" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" scope_family="Gesneriaceae">
|
||||
<subSubSection id="618FE1840B32FFB8FD93FBACB359FB24" box="[540,1047,1149,1176]" pageId="6" pageNumber="7" type="nomenclature">
|
||||
<paragraph id="292AB20F0B32FFB8FD93FBACB359FB24" blockId="6.[540,1047,1149,1210]" box="[540,1047,1149,1176]" pageId="6" pageNumber="7">
|
||||
<heading id="726205630B32FFB8FD93FBACB359FB24" box="[540,1047,1149,1176]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="7" reason="2">
|
||||
|
|
@ -91,14 +91,14 @@ Merr. (
|
|||
</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="292AB20F0B32FFB8FF32FAE5B6E5FACD" blockId="6.[189,1398,1296,1393]" pageId="6" pageNumber="7">
|
||||
<materialsCitation id="99FDB8520B32FFB8FF32FAE5B6E5FACD" ID-GBIF-Occurrence="2804098323" collectingDate="1913-12-09" collectionCode="BM" collectorName="Phil. Pl. Ramos" country="Philippines" county="Mt Cauayan" location="Luzon" municipality="Camarines" pageId="6" pageNumber="7" specimenCount="1" typeStatus="holotype">
|
||||
<materialsCitation id="99FDB8520B32FFB8FF32FAE5B6E5FACD" ID-GBIF-Occurrence="2804098323" collectingDate="1913-12-09" collectionCode="BM, GH, NY, P, US" collectorName="Phil. Pl. Ramos" country="Philippines" county="Mt Cauayan" location="Luzon" municipality="Camarines" pageId="6" pageNumber="7" specimenCount="1" typeStatus="syntype">
|
||||
<collectingCountry id="5182F29F0B32FFB8FF32FAE5B624FAF2" box="[189,362,1332,1358]" name="Philippines" pageId="6" pageNumber="7">PHILIPPINES</collectingCountry>
|
||||
<emphasis id="1BE16E1D0B32FFB8FEFDFAE5B632FAF2" bold="true" box="[370,380,1332,1358]" pageId="6" pageNumber="7">•</emphasis>
|
||||
<location id="2C4AE4D40B32FFB8FE0AFAE5B69DFAF2" LSID="urn:lsid:plazi:treatment:A13C03190B32FFB6FD93FBACB239FC10:2C4AE4D40B32FFB8FE0AFAE5B69DFAF2" box="[389,467,1332,1358]" country="Philippines" county="Mt Cauayan" municipality="Camarines" name="Luzon" pageId="6" pageNumber="7">Luzon</location>
|
||||
,
|
||||
<collectingMunicipality id="C94E28750B32FFB8FE6FFAE2B52EFAF1" box="[480,608,1331,1357]" pageId="6" pageNumber="7">Camarines</collectingMunicipality>
|
||||
,
|
||||
<collectingCounty id="C04BCA830B32FFB8FDE2FAE5B44CFAF2" box="[621,770,1332,1358]" pageId="6" pageNumber="7">Mt Cauayan</collectingCounty>
|
||||
<location id="2C4AE4D40B32FFB8FDE2FAE5B44CFAF2" LSID="urn:lsid:plazi:treatment:A13C03190B32FFB6FD93FBACB239FC10:2C4AE4D40B32FFB8FDE2FAE5B44CFAF2" box="[621,770,1332,1358]" country="Philippines" county="Mt Cauayan" municipality="Camarines" name="Mt Cauayan" pageId="6" pageNumber="7">Mt Cauayan</location>
|
||||
;
|
||||
<date id="5D2B94CF0B32FFB8FC80FAE5B4EDFAF2" box="[783,931,1332,1358]" pageId="6" pageNumber="7" value="1913-12-09">
|
||||
<collectingDate id="4D6F6D270B32FFB8FC80FAE5B4EDFAF2" box="[783,931,1332,1358]" pageId="6" pageNumber="7" value="1913-12-09">9 Dec. 1913</collectingDate>
|
||||
|
|
@ -108,14 +108,26 @@ Merr. (
|
|||
<collectorName id="8460D7D90B32FFB8FC3FFAE2B325FAF1" box="[944,1131,1331,1357]" pageId="6" pageNumber="7">Phil. Pl. Ramos</collectorName>
|
||||
1548
|
||||
</emphasis>
|
||||
(syntypes: BM!, GH!, NY!, P!, US!).
|
||||
(
|
||||
<typeStatus id="F62E0CAD0B32FFB8FB30FAE5B262FAF2" box="[1215,1324,1332,1358]" pageId="6" pageNumber="7">syntypes</typeStatus>
|
||||
:
|
||||
<collectionCode id="4F842ACA0B32FFB8FAB7FAE5B220FAF2" box="[1336,1390,1332,1358]" pageId="6" pageNumber="7">BM</collectionCode>
|
||||
!,
|
||||
<collectionCode id="4F842ACA0B32FFB8FF32FA86B7A1FACD" box="[189,239,1367,1393]" pageId="6" pageNumber="7">GH</collectionCode>
|
||||
!,
|
||||
<collectionCode id="4F842ACA0B32FFB8FE8FFA86B67DFACD" box="[256,307,1367,1393]" pageId="6" pageNumber="7">NY</collectionCode>
|
||||
!,
|
||||
<collectionCode id="4F842ACA0B32FFB8FECCFA86B614FACD" box="[323,346,1367,1393]" pageId="6" pageNumber="7">P</collectionCode>
|
||||
!,
|
||||
<collectionCode id="4F842ACA0B32FFB8FEE4FA86B6D7FACD" box="[363,409,1367,1393]" pageId="6" pageNumber="7">US</collectionCode>
|
||||
!).
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="292AB20F0B32FFB8FF65FA49B6AFFA0E" blockId="6.[189,1399,1432,1670]" box="[234,481,1432,1458]" pageId="6" pageNumber="7">
|
||||
<emphasis id="1BE16E1D0B32FFB8FF65FA49B6AFFA0E" bold="true" box="[234,481,1432,1458]" pageId="6" pageNumber="7">Additional material</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="292AB20F0B32FFB8FF32FA6AB3EDF93A" blockId="6.[189,1399,1432,1670]" pageId="6" pageNumber="7">
|
||||
<materialsCitation id="99FDB8520B32FFB8FF32FA6AB3F8FA69" ID-GBIF-Occurrence="2804098301" box="[189,1206,1467,1494]" collectingDate="1915-08" collectingDateMax="1997-03-22" collectingDateMin="1915-08" collectorName="Mendum" country="Philippines" location="Mt Isarog" municipality="Camarines Sur" pageId="6" pageNumber="7" specimenCount="1">
|
||||
<materialsCitation id="99FDB8520B32FFB8FF32FA6AB3F8FA69" ID-GBIF-Occurrence="2804098301" box="[189,1206,1467,1494]" collectingDate="1915-08" collectingDateMax="1997-03-22" collectingDateMin="1915-08" collectionCode="US" collectorName="Ramos" country="Philippines" location="Mt Isarog" municipality="Camarines Sur" pageId="6" pageNumber="7" specimenCode="US00081328" specimenCount="1">
|
||||
<collectingCountry id="5182F29F0B32FFB8FF32FA6AB624FA69" box="[189,362,1467,1493]" name="Philippines" pageId="6" pageNumber="7">PHILIPPINES</collectingCountry>
|
||||
<emphasis id="1BE16E1D0B32FFB8FEFCFA6AB633FA69" bold="true" box="[371,381,1467,1493]" pageId="6" pageNumber="7">•</emphasis>
|
||||
<location id="2C4AE4D40B32FFB8FE08FA6DB6B0FA6A" LSID="urn:lsid:plazi:treatment:A13C03190B32FFB6FD93FBACB239FC10:2C4AE4D40B32FFB8FE08FA6DB6B0FA6A" box="[391,510,1468,1494]" country="Philippines" municipality="Camarines Sur" name="Mt Isarog" pageId="6" pageNumber="7">Mt Isarog</location>
|
||||
|
|
@ -126,11 +138,16 @@ Merr. (
|
|||
<collectingDate id="4D6F6D270B32FFB8FD43FA6AB403FA6A" box="[716,845,1467,1494]" pageId="6" pageNumber="7" value="1915-08">Aug. 1915</collectingDate>
|
||||
</date>
|
||||
;
|
||||
<emphasis id="1BE16E1D0B32FFB8FCD4FA6DB4B0FA69" box="[859,1022,1467,1493]" italics="true" pageId="6" pageNumber="7">Ramos 23554</emphasis>
|
||||
(US00081328)
|
||||
<emphasis id="1BE16E1D0B32FFB8FCD4FA6DB4B0FA69" box="[859,1022,1467,1493]" italics="true" pageId="6" pageNumber="7">
|
||||
<collectorName id="8460D7D90B32FFB8FCD4FA6DB4E5FA69" box="[859,939,1468,1493]" pageId="6" pageNumber="7">Ramos</collectorName>
|
||||
23554
|
||||
</emphasis>
|
||||
(
|
||||
<specimenCode id="79331A740B32FFB8FB80FA6AB3E0FA69" box="[1039,1198,1467,1493]" pageId="6" pageNumber="7">US00081328</specimenCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
<emphasis id="1BE16E1D0B32FFB8FB30FA6AB387FA69" bold="true" box="[1215,1225,1467,1493]" pageId="6" pageNumber="7">•</emphasis>
|
||||
<materialsCitation id="99FDB8520B32FFB8FB5CFA6AB5E9FA45" ID-GBIF-Occurrence="2804098307" collectingDate="1915-08" collectingDateMax="1997-03-22" collectingDateMin="1915-08" collectorName="Mendum" country="Philippines" location="Mt Isarog" municipality="Camarines Sur" pageId="6" pageNumber="7" specimenCount="1">
|
||||
<materialsCitation id="99FDB8520B32FFB8FB5CFA6AB5E9FA45" ID-GBIF-Occurrence="2804098307" collectingDate="1997-03-22" collectingDateMax="1997-03-22" collectingDateMin="1915-08" collectionCode="E" collectorName="Mendum" country="Philippines" location="Mt Isarog" municipality="Camarines Sur" pageId="6" pageNumber="7" specimenCode="E00057041" specimenCount="1">
|
||||
ibid.;
|
||||
<date id="5D2B94CF0B32FFB8FA96FA6AB7B5FA45" pageId="6" pageNumber="7" value="1997-03-22">
|
||||
<collectingDate id="4D6F6D270B32FFB8FA96FA6AB7B5FA45" pageId="6" pageNumber="7" value="1997-03-22">22 Mar. 1997</collectingDate>
|
||||
|
|
@ -140,10 +157,12 @@ ibid.;
|
|||
<collectorName id="8460D7D90B32FFB8FE87FA0FB621FA44" box="[264,367,1502,1528]" pageId="6" pageNumber="7">Mendum</collectorName>
|
||||
et al. 29182
|
||||
</emphasis>
|
||||
(E00057041)
|
||||
(
|
||||
<specimenCode id="79331A740B32FFB8FD9AFA0FB5EEFA45" box="[533,672,1502,1529]" pageId="6" pageNumber="7">E00057041</specimenCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
<emphasis id="1BE16E1D0B32FFB8FD3FFA0EB5F4FA45" bold="true" box="[688,698,1503,1529]" pageId="6" pageNumber="7">•</emphasis>
|
||||
<materialsCitation id="99FDB8520B32FFB8FD4CFA0FB277FA45" ID-GBIF-Occurrence="2804098310" box="[707,1337,1502,1529]" collectingDate="1997-03-23" collectorName="Argent" pageId="6" pageNumber="7" specimenCount="1">
|
||||
<materialsCitation id="99FDB8520B32FFB8FD4CFA0FB277FA45" ID-GBIF-Occurrence="2804098310" box="[707,1337,1502,1529]" collectingDate="1997-03-23" collectingDateMax="1997-03-22" collectingDateMin="1915-08" collectionCode="L" collectorName="Argent" country="Philippines" location="Mt Isarog" municipality="Camarines Sur" pageId="6" pageNumber="7" specimenCode="L.3805694" specimenCount="1">
|
||||
ibid.;
|
||||
<date id="5D2B94CF0B32FFB8FC86FA0EB4E2FA45" box="[777,940,1503,1529]" pageId="6" pageNumber="7" value="1997-03-23">
|
||||
<collectingDate id="4D6F6D270B32FFB8FC86FA0EB4E2FA45" box="[777,940,1503,1529]" pageId="6" pageNumber="7" value="1997-03-23">23 Mar. 1997</collectingDate>
|
||||
|
|
@ -153,10 +172,12 @@ ibid.;
|
|||
<collectorName id="8460D7D90B32FFB8FC36FA0EB346FA45" box="[953,1032,1503,1529]" pageId="6" pageNumber="7">Argent</collectorName>
|
||||
et al. 20182
|
||||
</emphasis>
|
||||
(L.3805694)
|
||||
(
|
||||
<specimenCode id="79331A740B32FFB8FB22FA0FB27FFA45" box="[1197,1329,1502,1529]" pageId="6" pageNumber="7">L.3805694</specimenCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
<emphasis id="1BE16E1D0B32FFB8FACEFA0EB205FA45" bold="true" box="[1345,1355,1503,1529]" pageId="6" pageNumber="7">•</emphasis>
|
||||
<materialsCitation id="99FDB8520B32FFB8FADBFA0EB239F9A0" ID-GBIF-Occurrence="2804098328" collectingDate="1915-12" collectorName="Elmer" location="Mt Bulusan" municipality="Sorsogon" pageId="6" pageNumber="7" specimenCount="1">
|
||||
<materialsCitation id="99FDB8520B32FFB8FADBFA0EB239F9A0" ID-GBIF-Occurrence="2804098328" collectingDate="1915-12" collectionCode="L, US, U, P" collectorName="Elmer" location="Mt Bulusan" municipality="Sorsogon" pageId="6" pageNumber="7" specimenCode="L.2818244, US00081329, U.1341267, P03884333" specimenCount="1">
|
||||
<location id="2C4AE4D40B32FFB8FADBFA0EB66EF9A0" LSID="urn:lsid:plazi:treatment:A13C03190B32FFB6FD93FBACB239FC10:2C4AE4D40B32FFB8FADBFA0EB66EF9A0" municipality="Sorsogon" name="Mt Bulusan" pageId="6" pageNumber="7">Mt Bulusan</location>
|
||||
,
|
||||
<collectingMunicipality id="C94E28750B32FFB8FEBCF9D3B6E8F9A0" box="[307,422,1538,1564]" pageId="6" pageNumber="7">Sorsogon</collectingMunicipality>
|
||||
|
|
@ -169,10 +190,18 @@ et al. 20182
|
|||
<collectorName id="8460D7D90B32FFB8FDC0F9D3B5D9F9A0" box="[591,663,1538,1564]" pageId="6" pageNumber="7">Elmer</collectorName>
|
||||
16074
|
||||
</emphasis>
|
||||
(L.2818244, US00081329, U.1341267, P03884333)
|
||||
(
|
||||
<specimenCode id="79331A740B32FFB8FC89F9D3B4C4F9A0" box="[774,906,1538,1564]" pageId="6" pageNumber="7">L.2818244</specimenCode>
|
||||
,
|
||||
<specimenCode id="79331A740B32FFB8FC12F9D3B375F9A0" box="[925,1083,1538,1564]" pageId="6" pageNumber="7">US00081329</specimenCode>
|
||||
,
|
||||
<specimenCode id="79331A740B32FFB8FBC1F9D3B39AF9A0" box="[1102,1236,1538,1564]" pageId="6" pageNumber="7">U.1341267</specimenCode>
|
||||
,
|
||||
<specimenCode id="79331A740B32FFB8FB68F9D3B220F9A0" box="[1255,1390,1538,1564]" pageId="6" pageNumber="7">P03884333</specimenCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
<emphasis id="1BE16E1D0B32FFB8FF32F9F4B789F983" bold="true" box="[189,199,1573,1599]" pageId="6" pageNumber="7">•</emphasis>
|
||||
<materialsCitation id="99FDB8520B32FFB8FF42F9F4B5BEF9FC" ID-GBIF-Occurrence="2804098333" box="[205,752,1573,1600]" collectingDate="1958-06-19" collectorName="Sinclair" pageId="6" pageNumber="7" specimenCount="1">
|
||||
<materialsCitation id="99FDB8520B32FFB8FF42F9F4B5BEF9FC" ID-GBIF-Occurrence="2804098333" box="[205,752,1573,1600]" collectingDate="1958-06-19" collectionCode="E" collectorName="Sinclair" location="Mt Bulusan" municipality="Sorsogon" pageId="6" pageNumber="7" specimenCode="E00631523" specimenCount="1">
|
||||
ibid.;
|
||||
<date id="5D2B94CF0B32FFB8FE9FF9F4B6E9F9FC" box="[272,423,1573,1600]" pageId="6" pageNumber="7" value="1958-06-19">
|
||||
<collectingDate id="4D6F6D270B32FFB8FE9FF9F4B6E9F9FC" box="[272,423,1573,1600]" pageId="6" pageNumber="7" value="1958-06-19">19 Jun. 1958</collectingDate>
|
||||
|
|
@ -182,13 +211,15 @@ ibid.;
|
|||
<collectorName id="8460D7D90B32FFB8FE3DF9F4B541F983" box="[434,527,1573,1599]" pageId="6" pageNumber="7">Sinclair</collectorName>
|
||||
9624
|
||||
</emphasis>
|
||||
(E00631523)
|
||||
(
|
||||
<specimenCode id="79331A740B32FFB8FDD2F9F4B5A6F9FC" box="[605,744,1573,1600]" pageId="6" pageNumber="7">E00631523</specimenCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
<emphasis id="1BE16E1D0B32FFB8FD79F9F4B44EF983" bold="true" box="[758,768,1573,1599]" pageId="6" pageNumber="7">•</emphasis>
|
||||
<materialsCitation id="99FDB8520B32FFB8FC89F9F4B54FF9DF" ID-GBIF-Occurrence="2804098334" collectingDate="1991-11-15" collectorName="Reynoso & Romero & Fuentes" location="Mt Mayon" pageId="6" pageNumber="7" specimenCount="1" stateProvince="Albay">
|
||||
<location id="2C4AE4D40B32FFB8FC89F9F4B4CAF983" LSID="urn:lsid:plazi:treatment:A13C03190B32FFB6FD93FBACB239FC10:2C4AE4D40B32FFB8FC89F9F4B4CAF983" box="[774,900,1573,1599]" name="Mt Mayon" pageId="6" pageNumber="7" stateProvince="Albay">Mt Mayon</location>
|
||||
<materialsCitation id="99FDB8520B32FFB8FC89F9F4B54FF9DF" ID-GBIF-Occurrence="2804098334" collectingDate="1991-11-15" collectionCode="E" collectorName="Reynoso & Romero & Fuentes" location="Mt Mayon" municipality="Albay" pageId="6" pageNumber="7" specimenCode="E00316099" specimenCount="1" stateProvince="Albay">
|
||||
<location id="2C4AE4D40B32FFB8FC89F9F4B4CAF983" LSID="urn:lsid:plazi:treatment:A13C03190B32FFB6FD93FBACB239FC10:2C4AE4D40B32FFB8FC89F9F4B4CAF983" box="[774,900,1573,1599]" municipality="Albay" name="Mt Mayon" pageId="6" pageNumber="7" stateProvince="Albay">Mt Mayon</location>
|
||||
,
|
||||
<collectingRegion id="EB517CED0B32FFB8FC02F9F4B496F983" box="[909,984,1573,1599]" pageId="6" pageNumber="7">Albay</collectingRegion>
|
||||
<collectingMunicipality id="C94E28750B32FFB8FC02F9F4B496F983" box="[909,984,1573,1599]" pageId="6" pageNumber="7">Albay</collectingMunicipality>
|
||||
;
|
||||
<date id="5D2B94CF0B32FFB8FC6DF9F4B3CFF983" box="[994,1153,1573,1600]" pageId="6" pageNumber="7" value="1991-11-15">
|
||||
<collectingDate id="4D6F6D270B32FFB8FC6DF9F4B3CFF983" box="[994,1153,1573,1600]" pageId="6" pageNumber="7" value="1991-11-15">15 Nov. 1991</collectingDate>
|
||||
|
|
@ -199,14 +230,18 @@ ibid.;
|
|||
</emphasis>
|
||||
,
|
||||
<emphasis id="1BE16E1D0B32FFB8FB73F9F7B610F9DE" italics="true" pageId="6" pageNumber="7">
|
||||
<collectorName id="8460D7D90B32FFB8FB73F9F7B654F9DE" pageId="6" pageNumber="7">Romero & Fuentes</collectorName>
|
||||
<collectorName id="8460D7D90B32FFB8FB73F9F7B216F983" box="[1276,1368,1574,1599]" pageId="6" pageNumber="7">Romero</collectorName>
|
||||
&
|
||||
<collectorName id="8460D7D90B32FFB8FF32F998B654F9DE" box="[189,282,1609,1634]" pageId="6" pageNumber="7">Fuentes</collectorName>
|
||||
3584
|
||||
</emphasis>
|
||||
(E00316099)
|
||||
(
|
||||
<specimenCode id="79331A740B32FFB8FEE1F999B6B7F9DF" box="[366,505,1608,1635]" pageId="6" pageNumber="7">E00316099</specimenCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
<emphasis id="1BE16E1D0B32FFB8FD85F999B55AF9DE" bold="true" box="[522,532,1608,1634]" pageId="6" pageNumber="7">•</emphasis>
|
||||
<materialsCitation id="99FDB8520B32FFB8FD92F999B22DF9DE" ID-GBIF-Occurrence="2804098326" box="[541,1379,1608,1635]" collectingDate="1917-12-11" collectorName="Ramos" location="Catanduanes" pageId="6" pageNumber="7" specimenCount="1" stateProvince="Catanduanes">
|
||||
<collectingRegion id="EB517CED0B32FFB8FD92F999B5F9F9DE" box="[541,695,1608,1634]" pageId="6" pageNumber="7">Catanduanes</collectingRegion>
|
||||
<materialsCitation id="99FDB8520B32FFB8FD92F999B22DF9DE" ID-GBIF-Occurrence="2804098326" box="[541,1379,1608,1635]" collectingDate="1917-12-11" collectionCode="US, P" collectorName="Ramos" location="Catanduanes" municipality="Catanduanes" pageId="6" pageNumber="7" specimenCode="US00081330, P03884332" specimenCount="1" stateProvince="Catanduanes">
|
||||
<collectingMunicipality id="C94E28750B32FFB8FD92F999B5F9F9DE" box="[541,695,1608,1634]" pageId="6" pageNumber="7">Catanduanes</collectingMunicipality>
|
||||
;
|
||||
<date id="5D2B94CF0B32FFB8FD4BF999B428F9DE" box="[708,870,1608,1635]" pageId="6" pageNumber="7" value="1917-12-11">
|
||||
<collectingDate id="4D6F6D270B32FFB8FD4BF999B428F9DE" box="[708,870,1608,1635]" pageId="6" pageNumber="7" value="1917-12-11">11 Dec. 1917</collectingDate>
|
||||
|
|
@ -216,13 +251,17 @@ ibid.;
|
|||
<collectorName id="8460D7D90B32FFB8FCFBF998B48AF9DE" box="[884,964,1609,1634]" pageId="6" pageNumber="7">Ramos</collectorName>
|
||||
30288
|
||||
</emphasis>
|
||||
(US00081330, P03884332)
|
||||
(
|
||||
<specimenCode id="79331A740B32FFB8FBA8F999B389F9DE" box="[1063,1223,1608,1634]" pageId="6" pageNumber="7">US00081330</specimenCode>
|
||||
,
|
||||
<specimenCode id="79331A740B32FFB8FB5BF999B212F9DE" box="[1236,1372,1608,1634]" pageId="6" pageNumber="7">P03884332</specimenCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
<emphasis id="1BE16E1D0B32FFB8FAE3F999B238F9DE" bold="true" box="[1388,1398,1608,1634]" pageId="6" pageNumber="7">•</emphasis>
|
||||
<materialsCitation id="99FDB8520B32FFB8FF32F9BDB3EDF93A" ID-GBIF-Occurrence="2804098330" box="[189,1187,1643,1670]" collectingDate="1995-10-29" collectorName="Reynoso & Sagcal & Fernando" location="Mt Malinao" pageId="6" pageNumber="7" specimenCount="1" stateProvince="Albay">
|
||||
<location id="2C4AE4D40B32FFB8FF32F9BDB602F939" LSID="urn:lsid:plazi:treatment:A13C03190B32FFB6FD93FBACB239FC10:2C4AE4D40B32FFB8FF32F9BDB602F939" box="[189,332,1643,1670]" name="Mt Malinao" pageId="6" pageNumber="7" stateProvince="Albay">Mt Malinao</location>
|
||||
<materialsCitation id="99FDB8520B32FFB8FF32F9BDB3EDF93A" ID-GBIF-Occurrence="2804098330" box="[189,1187,1643,1670]" collectingDate="1995-10-29" collectionCode="L" collectorName="Reynoso & Sagcal & Fernando" location="Mt Malinao" municipality="Albay" pageId="6" pageNumber="7" specimenCode="L.3805666" specimenCount="1" stateProvince="Albay">
|
||||
<location id="2C4AE4D40B32FFB8FF32F9BDB602F939" LSID="urn:lsid:plazi:treatment:A13C03190B32FFB6FD93FBACB239FC10:2C4AE4D40B32FFB8FF32F9BDB602F939" box="[189,332,1643,1670]" municipality="Albay" name="Mt Malinao" pageId="6" pageNumber="7" stateProvince="Albay">Mt Malinao</location>
|
||||
,
|
||||
<collectingRegion id="EB517CED0B32FFB8FEDAF9BAB6EEF939" box="[341,416,1643,1669]" pageId="6" pageNumber="7">Albay</collectingRegion>
|
||||
<collectingMunicipality id="C94E28750B32FFB8FEDAF9BAB6EEF939" box="[341,416,1643,1669]" pageId="6" pageNumber="7">Albay</collectingMunicipality>
|
||||
;
|
||||
<date id="5D2B94CF0B32FFB8FE23F9BAB506F93A" box="[428,584,1643,1670]" pageId="6" pageNumber="7" value="1995-10-29">
|
||||
<collectingDate id="4D6F6D270B32FFB8FE23F9BAB506F93A" box="[428,584,1643,1670]" pageId="6" pageNumber="7" value="1995-10-29">29 Oct. 1995</collectingDate>
|
||||
|
|
@ -238,7 +277,9 @@ ibid.;
|
|||
<collectorName id="8460D7D90B32FFB8FCB3F9BAB4FEF939" box="[828,944,1643,1669]" pageId="6" pageNumber="7">Fernando</collectorName>
|
||||
21406
|
||||
</emphasis>
|
||||
(L.3805666).
|
||||
(
|
||||
<specimenCode id="79331A740B32FFB8FB9FF9BAB3D8F93A" box="[1040,1174,1643,1670]" pageId="6" pageNumber="7">L.3805666</specimenCode>
|
||||
).
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,69 +1,68 @@
|
|||
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|
||||
<treatment id="A13C03190B37FFB8FE4EFBE7B5B7FB83" ID-GBIF-Taxon="164870716" LSID="urn:lsid:plazi:treatment:A13C03190B37FFB8FE4EFBE7B5B7FB83" httpUri="http://treatment.plazi.org/id/A13C03190B37FFB8FE4EFBE7B5B7FB83" lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4" scope_family="Gesneriaceae">
|
||||
<subSubSection id="618FE1840B37FFBDFE4EFBE7B33DFBED" box="[449,1139,1078,1105]" pageId="3" pageNumber="4" type="nomenclature">
|
||||
<paragraph id="292AB20F0B37FFBDFE4EFBE7B33DFBED" blockId="3.[449,1139,1078,1175]" box="[449,1139,1078,1105]" pageId="3" pageNumber="4">
|
||||
<heading id="726205630B37FFBDFE4EFBE7B33DFBED" box="[449,1139,1078,1105]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="4" reason="2">
|
||||
<taxonomicName id="EE95C98C0B37FFBDFE4EFBE7B35FFBED" ID-CoL="7VLVW" authority="Olivar, H.J.Atkins & Muellner" authorityName="Olivar, H. J. Atkins & Muellner" authorityYear="2020" box="[449,1041,1078,1105]" class="Magnoliopsida" family="Gesneriaceae" genus="Cyrtandra" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="argentii" status="sp. nov.">
|
||||
<emphasis id="1BE16E1D0B37FFBDFE4EFBE7B5EDFBED" bold="true" box="[449,675,1078,1105]" italics="true" pageId="3" pageNumber="4">Cyrtandra argentii</emphasis>
|
||||
|
|
@ -71,7 +70,11 @@ Olivar, H.J.Atkins & Muellner
|
|||
</taxonomicName>
|
||||
<taxonomicNameLabel id="00D2D3660B37FFBDFB97FBE6B33DFBED" box="[1048,1139,1079,1105]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</heading>
|
||||
urn:lsid:ipni.org:names:77209562-1
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="618FE1840B37FFBDFDC9FB8BB4CEFB2B" pageId="3" pageNumber="4" type="description">
|
||||
<paragraph id="292AB20F0B37FFBDFDC9FB8BB4A3FBC9" blockId="3.[449,1139,1078,1175]" box="[582,1005,1114,1141]" pageId="3" pageNumber="4">urn:lsid:ipni.org:names:77209562-1</paragraph>
|
||||
<paragraph id="292AB20F0B37FFBDFD3AFBACB4CEFB2B" blockId="3.[449,1139,1078,1175]" box="[693,896,1149,1175]" pageId="3" pageNumber="4">
|
||||
<figureCitation id="B1AEAE8A0B37FFBDFD3AFBACB5B0FB2B" box="[693,766,1149,1175]" captionStart="Fig" captionStartId="4.[189,232,1919,1945]" captionTargetBox="[260,1327,1089,1876]" captionTargetId="figure@4.[249,1339,1083,1884]" captionTargetPageId="4" captionText="Fig. 1. Map of the Philippines showing known distributions of Cyrtandra argentii Olivar, H.J.Atkins & Muellner sp. nov. (*), C. ferruginea Merr. (■), C. hirtigera H.J.Atkins & Cronk (◆), and C. villosissima Merr. (●) based on collection localities. A single point may represent more than one collection." figureDoi="http://doi.org/10.5281/zenodo.3928634" httpUri="https://zenodo.org/record/3928634/files/figure.png" pageId="3" pageNumber="4">Figs 1</figureCitation>
|
||||
, 2, 3A, 4A
|
||||
</paragraph>
|
||||
|
|
@ -116,14 +119,14 @@ locality. George’s contribution to our knowledge of the Philippine flora is un
|
|||
</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="292AB20F0B37FFBDFF32F970B647F8BD" blockId="3.[189,1399,1661,1793]" pageId="3" pageNumber="4">
|
||||
<materialsCitation id="99FDB8520B37FFBDFF32F970B647F8BD" ID-GBIF-Occurrence="2804098305" collectingDate="1997-03-13" collectionCode="PNH" collectorName="Mendum, M. & Argent, C. G. C. & Pennington, R. T. & Wilkie, P. & Reynoso, E. J. & Gaerlan, F." country="Philippines" elevation="600" location="Oriental Province" municipality="Mindoro Island" pageId="3" pageNumber="4" specimenCount="1" typeStatus="holotype">
|
||||
<materialsCitation id="99FDB8520B37FFBDFF32F970B25EF962" ID-GBIF-Occurrence="2804098305" collectingDate="1997-03-13" collectionCode="E" collectorName="Mendum, M. & Argent, C. G. C. & Pennington, R. T. & Wilkie, P. & Reynoso, E. J. & Gaerlan, F." country="Philippines" elevation="600" location="Mindoro Island" municipality="Mindoro Island" pageId="3" pageNumber="4" specimenCount="1" stateProvince="Oriental Province" typeStatus="holotype">
|
||||
<collectingCountry id="5182F29F0B37FFBDFF32F970B624F907" box="[189,362,1697,1723]" name="Philippines" pageId="3" pageNumber="4">PHILIPPINES</collectingCountry>
|
||||
<emphasis id="1BE16E1D0B37FFBDFEFCF970B633F907" bold="true" box="[371,381,1697,1723]" pageId="3" pageNumber="4">•</emphasis>
|
||||
<collectingMunicipality id="C94E28750B37FFBDFE08F971B50DF906" box="[391,579,1696,1722]" pageId="3" pageNumber="4">Mindoro Island</collectingMunicipality>
|
||||
<location id="2C4AE4D40B37FFBDFE08F971B50DF906" LSID="urn:lsid:plazi:treatment:A13C03190B37FFB8FE4EFBE7B5B7FB83:2C4AE4D40B37FFBDFE08F971B50DF906" box="[391,579,1696,1722]" country="Philippines" municipality="Mindoro Island" name="Mindoro Island" pageId="3" pageNumber="4" stateProvince="Oriental Province">Mindoro Island</location>
|
||||
,
|
||||
<location id="2C4AE4D40B37FFBDFDDFF971B46BF906" LSID="urn:lsid:plazi:treatment:A13C03190B37FFB8FE4EFBE7B5B7FB83:2C4AE4D40B37FFBDFDDFF971B46BF906" box="[592,805,1696,1722]" country="Philippines" municipality="Mindoro Island" name="Oriental Province" pageId="3" pageNumber="4">Oriental Province</location>
|
||||
<collectingRegion id="EB517CED0B37FFBDFDDFF971B46BF906" box="[592,805,1696,1722]" pageId="3" pageNumber="4">Oriental Province</collectingRegion>
|
||||
,
|
||||
<location id="2C4AE4D40B37FFBDFCBCF970B4F9F906" LSID="urn:lsid:plazi:treatment:A13C03190B37FFB8FE4EFBE7B5B7FB83:2C4AE4D40B37FFBDFCBCF970B4F9F906" box="[819,951,1696,1723]" country="Philippines" municipality="Mindoro Island" name="Mt Halcon" pageId="3" pageNumber="4">Mt Halcon</location>
|
||||
<location id="2C4AE4D40B37FFBDFCBCF970B4F9F906" LSID="urn:lsid:plazi:treatment:A13C03190B37FFB8FE4EFBE7B5B7FB83:2C4AE4D40B37FFBDFCBCF970B4F9F906" box="[819,951,1696,1723]" country="Philippines" municipality="Mindoro Island" name="Mt Halcon" pageId="3" pageNumber="4" stateProvince="Oriental Province">Mt Halcon</location>
|
||||
;
|
||||
<quantity id="EE6D1FEA0B37FFBDFC49F971B35BF907" box="[966,1045,1696,1723]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="4" unit="m" value="600.0">
|
||||
<elevation id="A2B8553C0B37FFBDFC49F971B35BF907" box="[966,1045,1696,1723]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="4" unit="m" value="600.0">600 m</elevation>
|
||||
|
|
@ -170,8 +173,14 @@ locality. George’s contribution to our knowledge of the Philippine flora is un
|
|||
</collectorName>
|
||||
29053 (
|
||||
<typeStatus id="F62E0CAD0B37FFBDFBFCF915B3AEF962" box="[1139,1248,1732,1758]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
|
||||
: E!; isotype:
|
||||
<collectionCode id="4F842ACA0B37FFBDFF32F936B7B3F8BD" box="[189,253,1767,1793]" country="Philippines" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15314" lsid="urn:lsid:biocol.org:col:15314" name="National Museum" pageId="3" pageNumber="4" type="Herbarium">PNH</collectionCode>
|
||||
:
|
||||
<collectionCode id="4F842ACA0B37FFBDFB62F915B248F962" box="[1261,1286,1732,1758]" pageId="3" pageNumber="4">E</collectionCode>
|
||||
!;
|
||||
</materialsCitation>
|
||||
<materialsCitation id="99FDB8520B37FFBDFA95F915B647F8BD" ID-GBIF-Occurrence="2804098305" collectingDate="1997-03-13" collectionCode="PNH" collectorName="Mendum, M. & Argent, C. G. C. & Pennington, R. T. & Wilkie, P. & Reynoso, E. J. & Gaerlan, F." country="Philippines" elevation="600" location="Mindoro Island" municipality="Mindoro Island" pageId="3" pageNumber="4" specimenCount="1" stateProvince="Oriental Province" typeStatus="isotype">
|
||||
<typeStatus id="F62E0CAD0B37FFBDFA95F915B23DF962" box="[1306,1395,1732,1758]" pageId="3" pageNumber="4">isotype</typeStatus>
|
||||
:
|
||||
<collectionCode id="4F842ACA0B37FFBDFF32F936B7B3F8BD" box="[189,253,1767,1793]" collectionName="PNH" country="Philippines" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15314" lsid="urn:lsid:biocol.org:col:15314" name="National Museum" pageId="3" pageNumber="4" type="Herbarium">PNH</collectionCode>
|
||||
).
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
|
|
@ -179,10 +188,10 @@ locality. George’s contribution to our knowledge of the Philippine flora is un
|
|||
<emphasis id="1BE16E1D0B37FFBDFF65F8F9B6AFF8FE" bold="true" box="[234,481,1832,1858]" pageId="3" pageNumber="4">Additional material</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="292AB20F0B37FFBDFF32F89DB239F873" blockId="3.[189,1399,1832,2000]" pageId="3" pageNumber="4">
|
||||
<materialsCitation id="99FDB8520B37FFBDFF32F89DB238F8DA" ID-GBIF-Occurrence="2804098322" box="[189,1398,1867,1894]" collectingDate="1986-05-12" collectorName="C. E. Ridsdale" country="Philippines" location="Mindoro" municipality="Mt Halcon" pageId="3" pageNumber="4" specimenCount="1">
|
||||
<materialsCitation id="99FDB8520B37FFBDFF32F89DB238F8DA" ID-GBIF-Occurrence="2804098322" box="[189,1398,1867,1894]" collectingDate="1986-05-12" collectionCode="K, L" collectorName="C. E. Ridsdale" country="Philippines" location="Mindoro" municipality="Mt Halcon" pageId="3" pageNumber="4" specimenCode="K000223279, L.2822762" specimenCount="1">
|
||||
<collectingCountry id="5182F29F0B37FFBDFF32F89DB624F8DA" box="[189,362,1868,1894]" name="Philippines" pageId="3" pageNumber="4">PHILIPPINES</collectingCountry>
|
||||
<emphasis id="1BE16E1D0B37FFBDFEFBF89DB630F8DA" bold="true" box="[372,382,1868,1894]" pageId="3" pageNumber="4">•</emphasis>
|
||||
<collectingMunicipality id="C94E28750B37FFBDFE07F89DB542F8D9" box="[392,524,1867,1894]" pageId="3" pageNumber="4">Mt Halcon</collectingMunicipality>
|
||||
<collectingMunicipality id="C94E28750B37FFBDFE07F89DB542F8D9" box="[392,524,1867,1894]" country="Philippines" municipality="Mt Halcon" name="Mt Halcon" pageId="3" pageNumber="4">Mt Halcon</collectingMunicipality>
|
||||
,
|
||||
<location id="2C4AE4D40B37FFBDFD95F89AB5CAF8D9" LSID="urn:lsid:plazi:treatment:A13C03190B37FFB8FE4EFBE7B5B7FB83:2C4AE4D40B37FFBDFD95F89AB5CAF8D9" box="[538,644,1867,1893]" country="Philippines" municipality="Mt Halcon" name="Mindoro" pageId="3" pageNumber="4">Mindoro</location>
|
||||
;
|
||||
|
|
@ -194,48 +203,65 @@ locality. George’s contribution to our knowledge of the Philippine flora is un
|
|||
<collectorName id="8460D7D90B37FFBDFCC9F89DB4A5F8D9" box="[838,1003,1867,1894]" pageId="3" pageNumber="4">C.E. Ridsdale</collectorName>
|
||||
1762
|
||||
</emphasis>
|
||||
(K000223279, L.2822762)
|
||||
(
|
||||
<specimenCode id="79331A740B37FFBDFBCFF89DB391F8DA" box="[1088,1247,1868,1894]" pageId="3" pageNumber="4">K000223279</specimenCode>
|
||||
,
|
||||
<specimenCode id="79331A740B37FFBDFB62F89AB221F8DA" box="[1261,1391,1867,1894]" pageId="3" pageNumber="4">L.2822762</specimenCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
<emphasis id="1BE16E1D0B37FFBDFF32F8BEB789F835" bold="true" box="[189,199,1903,1929]" pageId="3" pageNumber="4">•</emphasis>
|
||||
<materialsCitation id="99FDB8520B37FFBDFF43F8BEB388F835" ID-GBIF-Occurrence="2804098308" box="[204,1222,1902,1929]" collectingDate="1991-04-01" collectingDateMax="1997-03-13" collectingDateMin="1991-04-01" location="Gaerlan" municipality="Argent" pageId="3" pageNumber="4" specimenCount="1">
|
||||
<materialsCitation id="99FDB8520B37FFBDFF43F8BEB388F835" ID-GBIF-Occurrence="2804098308" box="[204,1222,1902,1929]" collectingDate="1991-04-01" collectingDateMax="1997-03-13" collectingDateMin="1991-04-01" collectionCode="K, L, US" collectorName="Stone & Reynoso & Sagcal" country="Philippines" location="Mindoro" municipality="Argent" pageId="3" pageNumber="4" specimenCode="K000223280, L.2822797, US00737625" specimenCount="1">
|
||||
ibid.;
|
||||
<date id="5D2B94CF0B37FFBDFE80F8BEB6D6F835" box="[271,408,1903,1929]" pageId="3" pageNumber="4" value="1991-04-01">
|
||||
<collectingDate id="4D6F6D270B37FFBDFE80F8BEB6D6F835" box="[271,408,1903,1929]" pageId="3" pageNumber="4" value="1991-04-01">1Apr. 1991</collectingDate>
|
||||
</date>
|
||||
;
|
||||
<emphasis id="1BE16E1D0B37FFBDFE2DF8BEB6ADF835" box="[418,483,1903,1929]" italics="true" pageId="3" pageNumber="4">Stone</emphasis>
|
||||
<emphasis id="1BE16E1D0B37FFBDFE2DF8BEB6ADF835" box="[418,483,1903,1929]" italics="true" pageId="3" pageNumber="4">
|
||||
<collectorName id="8460D7D90B37FFBDFE2DF8BEB6ADF835" box="[418,483,1903,1929]" pageId="3" pageNumber="4">Stone</collectorName>
|
||||
</emphasis>
|
||||
,
|
||||
<emphasis id="1BE16E1D0B37FFBDFE60F8A1B51CF835" box="[495,594,1904,1929]" italics="true" pageId="3" pageNumber="4">Reynoso</emphasis>
|
||||
<emphasis id="1BE16E1D0B37FFBDFE60F8A1B51CF835" box="[495,594,1904,1929]" italics="true" pageId="3" pageNumber="4">
|
||||
<collectorName id="8460D7D90B37FFBDFE60F8A1B51CF835" box="[495,594,1904,1929]" pageId="3" pageNumber="4">Reynoso</collectorName>
|
||||
</emphasis>
|
||||
,
|
||||
<emphasis id="1BE16E1D0B37FFBDFDD0F8BEB5AEF835" box="[607,736,1903,1929]" italics="true" pageId="3" pageNumber="4">Sagcal 504</emphasis>
|
||||
(K000223280, L.2822797, US00737625)
|
||||
<emphasis id="1BE16E1D0B37FFBDFDD0F8BEB5AEF835" box="[607,736,1903,1929]" italics="true" pageId="3" pageNumber="4">
|
||||
<collectorName id="8460D7D90B37FFBDFDD0F8BEB5E1F835" box="[607,687,1903,1929]" pageId="3" pageNumber="4">Sagcal</collectorName>
|
||||
504
|
||||
</emphasis>
|
||||
(
|
||||
<specimenCode id="79331A740B37FFBDFD62F8BEB4C2F835" box="[749,908,1903,1929]" pageId="3" pageNumber="4">K000223280</specimenCode>
|
||||
,
|
||||
<specimenCode id="79331A740B37FFBDFC1AF8BEB356F835" box="[917,1048,1903,1929]" pageId="3" pageNumber="4">L.2822797</specimenCode>
|
||||
,
|
||||
<specimenCode id="79331A740B37FFBDFBADF8BFB3F0F835" box="[1058,1214,1902,1929]" pageId="3" pageNumber="4">US00737625</specimenCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
<emphasis id="1BE16E1D0B37FFBDFB44F8BEB39BF835" bold="true" box="[1227,1237,1903,1929]" pageId="3" pageNumber="4">•</emphasis>
|
||||
<materialsCitation id="99FDB8520B37FFBDFB54F8BEB469F811" ID-GBIF-Occurrence="2804098314" collectingDate="1991-04-01" collectingDateMax="1997-03-13" collectingDateMin="1991-04-01" location="Gaerlan" municipality="Argent" pageId="3" pageNumber="4" specimenCount="1">
|
||||
<materialsCitation id="99FDB8520B37FFBDFB54F8BEB469F811" ID-GBIF-Occurrence="2804098314" collectingDate="1997-03-13" collectingDateMax="1997-03-13" collectingDateMin="1991-04-01" collectionCode="L" collectorName="Argent & Gaerlan & Reynoso" country="Philippines" location="Mindoro" municipality="Argent" pageId="3" pageNumber="4" specimenCode="L.3805692" specimenCount="1">
|
||||
ibid.;
|
||||
<date id="5D2B94CF0B37FFBDFA92F8BEB7B5F810" pageId="3" pageNumber="4" value="1997-03-13">
|
||||
<collectingDate id="4D6F6D270B37FFBDFA92F8BEB7B5F810" pageId="3" pageNumber="4" value="1997-03-13">13 Mar. 1997</collectingDate>
|
||||
</date>
|
||||
;
|
||||
<emphasis id="1BE16E1D0B37FFBDFE87F843B619F810" box="[264,343,1938,1964]" italics="true" pageId="3" pageNumber="4">
|
||||
<collectingMunicipality id="C94E28750B37FFBDFE87F843B619F810" box="[264,343,1938,1964]" pageId="3" pageNumber="4">Argent</collectingMunicipality>
|
||||
<collectorName id="8460D7D90B37FFBDFE87F843B619F810" box="[264,343,1938,1964]" pageId="3" pageNumber="4">Argent</collectorName>
|
||||
</emphasis>
|
||||
,
|
||||
<emphasis id="1BE16E1D0B37FFBDFEE9F843B686F810" box="[358,456,1938,1964]" italics="true" pageId="3" pageNumber="4">
|
||||
<location id="2C4AE4D40B37FFBDFEE9F843B686F810" LSID="urn:lsid:plazi:treatment:A13C03190B37FFB8FE4EFBE7B5B7FB83:2C4AE4D40B37FFBDFEE9F843B686F810" box="[358,456,1938,1964]" municipality="Argent" name="Gaerlan" pageId="3" pageNumber="4">Gaerlan</location>
|
||||
<collectorName id="8460D7D90B37FFBDFEE9F843B686F810" box="[358,456,1938,1964]" pageId="3" pageNumber="4">Gaerlan</collectorName>
|
||||
</emphasis>
|
||||
,
|
||||
<emphasis id="1BE16E1D0B37FFBDFE58F842B5C5F810" box="[471,651,1938,1964]" italics="true" pageId="3" pageNumber="4">
|
||||
<location id="2C4AE4D40B37FFBDFE58F842B574F810" LSID="urn:lsid:plazi:treatment:A13C03190B37FFB8FE4EFBE7B5B7FB83:2C4AE4D40B37FFBDFE58F842B574F810" box="[471,570,1939,1964]" municipality="Argent" name="Reynoso" pageId="3" pageNumber="4">Reynoso</location>
|
||||
<collectorName id="8460D7D90B37FFBDFE58F842B574F810" box="[471,570,1939,1964]" pageId="3" pageNumber="4">Reynoso</collectorName>
|
||||
20053
|
||||
</emphasis>
|
||||
(L.3805692)
|
||||
(
|
||||
<specimenCode id="79331A740B37FFBDFD13F843B451F811" box="[668,799,1938,1965]" pageId="3" pageNumber="4">L.3805692</specimenCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
<emphasis id="1BE16E1D0B37FFBDFCA0F843B477F810" bold="true" box="[815,825,1938,1964]" pageId="3" pageNumber="4">•</emphasis>
|
||||
<materialsCitation id="99FDB8520B37FFBDFCCDF843B527F86C" ID-GBIF-Occurrence="2804098316" collectingDate="1968-03-25" collectorName="Jacobs, M." location="Baler Aurora" municipality="Sierra Madre Mountains" pageId="3" pageNumber="4" specimenCount="1">
|
||||
<collectingMunicipality id="C94E28750B37FFBDFCCDF843B32BF810" box="[834,1125,1938,1964]" pageId="3" pageNumber="4">Sierra Madre Mountains</collectingMunicipality>
|
||||
,
|
||||
<location id="2C4AE4D40B37FFBDFBFEF843B240F810" LSID="urn:lsid:plazi:treatment:A13C03190B37FFB8FE4EFBE7B5B7FB83:2C4AE4D40B37FFBDFBFEF843B240F810" box="[1137,1294,1938,1964]" municipality="Sierra Madre Mountains" name="Baler Aurora" pageId="3" pageNumber="4">Baler Aurora</location>
|
||||
<materialsCitation id="99FDB8520B37FFBDFCCDF843B527F86C" ID-GBIF-Occurrence="2804098316" collectingDate="1968-03-25" collectionCode="L" collectorName="Jacobs, M." location="Sierra Madre Mountains, Baler Aurora" municipality="Sierra Madre Mountains" pageId="3" pageNumber="4" specimenCode="L.2822694" specimenCount="1">
|
||||
<location id="2C4AE4D40B37FFBDFCCDF843B240F810" LSID="urn:lsid:plazi:treatment:A13C03190B37FFB8FE4EFBE7B5B7FB83:2C4AE4D40B37FFBDFCCDF843B240F810" box="[834,1294,1938,1964]" municipality="Sierra Madre Mountains" name="Sierra Madre Mountains, Baler Aurora" pageId="3" pageNumber="4">Sierra Madre Mountains, Baler Aurora</location>
|
||||
;
|
||||
<date id="5D2B94CF0B37FFBDFA95F843B7B5F873" pageId="3" pageNumber="4" value="1968-03-25">
|
||||
<collectingDate id="4D6F6D270B37FFBDFA95F843B7B5F873" pageId="3" pageNumber="4" value="1968-03-25">25 Mar. 1968</collectingDate>
|
||||
|
|
@ -246,11 +272,13 @@ ibid.;
|
|||
,
|
||||
<emphasis id="1BE16E1D0B37FFBDFEE5F867B6C4F873" box="[362,394,1974,1999]" italics="true" pageId="3" pageNumber="4">M.</emphasis>
|
||||
</collectorName>
|
||||
8002 (L.2822694)
|
||||
8002 (
|
||||
<specimenCode id="79331A740B37FFBDFE51F864B52FF86C" box="[478,609,1973,2000]" pageId="3" pageNumber="4">L.2822694</specimenCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
<emphasis id="1BE16E1D0B37FFBDFDFDF864B532F873" bold="true" box="[626,636,1973,1999]" pageId="3" pageNumber="4">•</emphasis>
|
||||
<materialsCitation id="99FDB8520B37FFBDFD0AF864B239F873" ID-GBIF-Occurrence="2804098320" box="[645,1399,1973,2000]" collectingDate="1993-03-08" collectorName="Barbon & Garcia & Fernando" location="Aurora National Park" municipality="Aurora National Park" pageId="3" pageNumber="4" specimenCount="1">
|
||||
<collectingMunicipality id="C94E28750B37FFBDFD0AF864B4C7F873" box="[645,905,1973,1999]" pageId="3" pageNumber="4">Aurora National Park</collectingMunicipality>
|
||||
<location id="2C4AE4D40B37FFBDFD0AF864B4C7F873" LSID="urn:lsid:plazi:treatment:A13C03190B37FFB8FE4EFBE7B5B7FB83:2C4AE4D40B37FFBDFD0AF864B4C7F873" box="[645,905,1973,1999]" municipality="Aurora National Park" name="Aurora National Park" pageId="3" pageNumber="4">Aurora National Park</location>
|
||||
;
|
||||
<date id="5D2B94CF0B37FFBDFC19F864B365F873" box="[918,1067,1973,2000]" pageId="3" pageNumber="4" value="1993-03-08">
|
||||
<collectingDate id="4D6F6D270B37FFBDFC19F864B365F873" box="[918,1067,1973,2000]" pageId="3" pageNumber="4" value="1993-03-08">8 Mar. 1993</collectingDate>
|
||||
|
|
@ -275,7 +303,7 @@ ibid.;
|
|||
<emphasis id="1BE16E1D0B30FFBAFF32FEDDB7B6FE9A" box="[189,248,268,294]" italics="true" pageId="4" pageNumber="5">9121</emphasis>
|
||||
(K000223281, L.3794225)
|
||||
<emphasis id="1BE16E1D0B30FFBAFDCAFEDDB501FE9A" bold="true" box="[581,591,268,294]" pageId="4" pageNumber="5">•</emphasis>
|
||||
<materialsCitation id="99FDB8520B30FFBAFDD7FEDAB616FEF5" ID-GBIF-Occurrence="2804098318" collectingDate="1955-06-25" collectorName="Lagrimas, M." location="Llavac" municipality="Infanta" pageId="4" pageNumber="5" specimenCount="1" stateProvince="Quezon Province">
|
||||
<materialsCitation id="99FDB8520B30FFBAFDD7FEDAB616FEF5" ID-GBIF-Occurrence="2804098318" collectingDate="1955-06-25" collectionCode="L" collectorName="Lagrimas, M." location="Llavac" municipality="Infanta" pageId="4" pageNumber="5" specimenCode="L.2822646" specimenCount="1" stateProvince="Quezon Province">
|
||||
<location id="2C4AE4D40B30FFBAFDD7FEDAB5E5FE99" LSID="urn:lsid:plazi:treatment:A13C03190B37FFB8FE4EFBE7B5B7FB83:2C4AE4D40B30FFBAFDD7FEDAB5E5FE99" box="[600,683,267,293]" municipality="Infanta" name="Llavac" pageId="4" pageNumber="5" stateProvince="Quezon Province">Llavac</location>
|
||||
,
|
||||
<collectingMunicipality id="C94E28750B30FFBAFD37FEDAB440FE99" box="[696,782,267,293]" pageId="4" pageNumber="5">Infanta</collectingMunicipality>
|
||||
|
|
@ -291,7 +319,9 @@ ibid.;
|
|||
,
|
||||
<emphasis id="1BE16E1D0B30FFBAFAADFEDDB20CFE99" box="[1314,1346,268,293]" italics="true" pageId="4" pageNumber="5">M.</emphasis>
|
||||
</collectorName>
|
||||
521 (L.2822646).
|
||||
521 (
|
||||
<specimenCode id="79331A740B30FFBAFF4AFEFFB602FEF5" box="[197,332,302,329]" pageId="4" pageNumber="5">L.2822646</specimenCode>
|
||||
).
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
@ -504,5 +534,4 @@ Bramley
|
|||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</subSection>
|
||||
</document>
|
||||
Loading…
Reference in a new issue