Add updates up until 2025-01-24 17:06:22

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<document id="9642A59CAF72C0D2D14F4B543A06C024" ID-DOI="10.11646/zootaxa.5570.2.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="14735135" ID-ZooBank="8B1BAE9E-FBD3-4AC3-9FA3-9CCC2AA81392" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737737634945" checkinUser="plazi" docAuthor="Xin, Fei-Yi, Zhong, Xiao-Tian, Guo, Wen-Chi, Zhang, Yi-Feng, Qi, Zhi-Hao &amp; Wu, Song-Qing" docDate="2025" docId="03DC8794FFA2D678FF3CFAA0C25B2B0D" docLanguage="en" docName="zootaxa.5570.2.6.pdf" docOrigin="Zootaxa 5570 (2)" docSource="https://doi.org/10.11646/zootaxa.5570.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neolucanus brochieri Bomans &amp; Miyashita 1997" docType="treatment" docVersion="2" lastPageNumber="357" masterDocId="FFE5FFECFFAED675FFABFFE1C3352F38" masterDocTitle="A taxonomic note on Chinese species of the Neolucanus fuscus group with the description of a new species (Coleoptera: Lucanidae: Lucaninae)" masterLastPageNumber="360" masterPageNumber="344" pageNumber="356" updateTime="1737738157217" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E0726F887DDC5AB544052C049D004923">A taxonomic note on Chinese species of the Neolucanus fuscus group with the description of a new species (Coleoptera: Lucanidae: Lucaninae)</mods:title>
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<mods:namePart id="E68195A7F4A24FEC313549BF862D08D1">Xin, Fei-Yi</mods:namePart>
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<mods:namePart id="C6CD99E572109DB4F597560B08978A50">Zhong, Xiao-Tian</mods:namePart>
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<mods:namePart id="3E5850FC07D82BAA23083FCA45D65FC0">Zhang, Yi-Feng</mods:namePart>
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<mods:namePart id="26DD9577C2EEBF1ED6DA9C0186069FE5">Qi, Zhi-Hao</mods:namePart>
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<mods:affiliation id="677EBFB7833F4F0A0DFA26516CB46C19">Key Laboratory of Integrated Pest Management in Ecological Forests, Fujian Province University, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Fujian Academy of Forestry, Fuzhou 350012, China.</mods:affiliation>
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<mods:namePart id="44DEF4C04BD27C345F74593A7E2290B2">Wu, Song-Qing</mods:namePart>
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<heading id="D08281EEFFA2D679FF3CFAA0C1CC2A63" bold="true" box="[151,761,1345,1371]" fontSize="11" level="1" pageId="12" pageNumber="356" reason="1">
<taxonomicName id="4C754D01FFA2D679FF3CFAA0C1CC2A63" ID-CoL="46LQW" authority="Bomans &amp; Miyashita, 1997" authorityName="Bomans &amp; Miyashita" authorityYear="1997" box="[151,761,1345,1371]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="356" phylum="Arthropoda" rank="species" species="brochieri">
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<emphasis id="B901EA90FFA2D679FF3CFAA0C2AA2A63" bold="true" box="[151,415,1345,1371]" italics="true" pageId="12" pageNumber="356">Neolucanus brochieri</emphasis>
<bibRefCitation id="EFE44B73FFA2D679FE0DFAA0C1CC2A63" author="Bomans, H. E. &amp; Miyashita, T." box="[422,761,1345,1371]" pageId="12" pageNumber="356" pagination="2 - 3" refId="ref7344" refString="Bomans, H. E. &amp; Miyashita, T. (1997) Description de trois nouvelles especes de Lucanidae du nord Birmanie. Besoiro, 4, 2 - 3." type="journal article" year="1997">Bomans &amp; Miyashita, 1997</bibRefCitation>
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Suggested Chinese common name:
<emphasis id="B901EA90FFA2D679FD8CFA87C1FA2A45" bold="true" box="[551,719,1382,1405]" pageId="12" pageNumber="356">DZae圆OiNJƤ</emphasis>
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<figureCitation id="134E2A07FFA2D679FF3CFA68C22D2A9B" box="[151,280,1417,1443]" captionStart="FIGURE 1" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[312,1275,185,1877]" captionTargetId="figure-17@3.[306,1281,179,1883]" captionTargetPageId="3" captionText="FIGURE 1. Male habitus of Neolucanus species. AB, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype, 36.0 mm, telodont (Lvchun, Yunnan); CE, N. fuscus Didier, 1926; CD, 50.9 mm, telodont (Yên Bái, Yên Bái); E, 44.8 mm, telodont (Ziyuan, Guangxi); FH, N. brochieri Bomans &amp; Miyashita, 1997; FG, 41.1mm, telodont (Gonshan, Yunnan); H, 41.0 mm, telodont (Gonshan, Yunnan). A, C, E, F, H, dorsal view; B, D, G, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735137" httpUri="https://zenodo.org/record/14735137/files/figure.png" pageId="12" pageNumber="356">Figs. 1FH</figureCitation>
;
<figureCitation id="134E2A07FFA2D679FE8FFA68C2502A9B" box="[292,357,1417,1443]" captionStart="FIGURE 2" captionStartId="4.[151,250,1771,1795]" captionTargetBox="[158,1429,186,1740]" captionTargetId="figure-18@4.[151,1436,181,1747]" captionTargetPageId="4" captionText="FIGURE 2. Male habitus of Neolucanus species. AB, N. horizontalis Xin, Zhong &amp; Qi, new species, paratype, 33.7 mm, mesodont (Jinping, Yunnan); CD, N. fuscus Didier, 1926, 34.7mm, priodont (Wuming, Guangxi); EF, N. brochieri Bomans &amp; Miyashita, 1997, 31.4 mm, priodont (Gonshan, Yunnan). A, C, E, dorsal view; B, D, F, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735139" httpUri="https://zenodo.org/record/14735139/files/figure.png" pageId="12" pageNumber="356">2EF</figureCitation>
; 3MP; 4IL; 5MP; 6D, H, L; 7D
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<taxonomicName id="4C754D01FFA2D679FF3CFA2FC1982ADE" ID-CoL="46LQW" authority="Bomans &amp; Miyashita, 1997: 2" authorityName="Bomans &amp; Miyashita" authorityPageNumber="2" authorityYear="1997" box="[151,685,1486,1510]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="356" phylum="Arthropoda" rank="species" species="brochieri">
<emphasis id="B901EA90FFA2D679FF3CFA2FC2472ADE" box="[151,370,1486,1510]" italics="true" pageId="12" pageNumber="356">Neolucanus brochieri</emphasis>
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locality.
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.
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material examined.
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None.
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<emphasis id="B901EA90FFA2D679FF6CF9BDC10F294E" bold="true" box="[199,570,1628,1654]" pageId="12" pageNumber="356">Additional material examined.</emphasis>
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<collectingCountry id="F3627612FFA2D679FDEFF9BDC1A4294E" box="[580,657,1628,1654]" name="China" pageId="12" pageNumber="356">China</collectingCountry>
:
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,
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,
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, 2011, local collector leg
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.;
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<collectingCounty id="62AB4E0EFFA2D679FF3CF945C2502986" box="[151,357,1700,1726]" pageId="12" pageNumber="356">Gongshan County</collectingCounty>
,
<location id="8EAA6059FFA2D679FEDAF945C17F2986" LSID="urn:lsid:plazi:treatment:03DC8794FFA2D678FF3CFAA0C25B2B0D:8EAA6059FFA2D679FEDAF945C17F2986" box="[369,586,1700,1726]" country="China" county="Gongshan County" municipality="Nujiang Prefecture" name="Dulongjiang valley" pageId="12" pageNumber="356" stateProvince="Yunnan">Dulongjiang valley</location>
,
<date id="FFCB1042FFA2D679FDFCF945C1D62986" box="[599,739,1700,1726]" pageId="12" pageNumber="356" value="2023-07-19">
<collectingDate id="EF8FE9AAFFA2D679FDFCF945C1D62986" box="[599,739,1700,1726]" pageId="12" pageNumber="356" value="2023-07-19">19.VII.2023</collectingDate>
</date>
,
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<elevation id="0058D1B1FFA2D679FD44F945C00A2986" box="[751,831,1700,1726]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="356" unit="m" value="1800.0">1800m</elevation>
</quantity>
,
<collectorName id="26805354FFA2D679FCE7F945C08D2986" box="[844,952,1700,1726]" pageId="12" pageNumber="356">Feiyi Xin</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B1D3CDFFFA2D679FC52F945C7BD29DA" collectingDate="2015-07" collectionCode="CZHQ" collectorName="Xinxiong Deng" country="China" county="Nujiang Prefecture" elevation="2500" location="Pianma Township" municipality="Lushui" pageId="12" pageNumber="356" specimenCount="15" specimenCount-male="15" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="9D73FD0BFFA2D679FC52F945C76B2986" box="[1017,1118,1700,1727]" count="15" pageId="12" pageNumber="356" type="male">15 males</specimenCount>
(
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),
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,
<collectingCounty id="62AB4E0EFFA2D679FF3CF929C25B29DA" box="[151,366,1736,1762]" pageId="12" pageNumber="356">Nujiang Prefecture</collectingCounty>
,
<collectingMunicipality id="6BAEACF8FFA2D679FED2F929C2F229DA" box="[377,455,1736,1762]" pageId="12" pageNumber="356">Lushui</collectingMunicipality>
city,
<location id="8EAA6059FFA2D679FDAFF929C1E529DA" LSID="urn:lsid:plazi:treatment:03DC8794FFA2D678FF3CFAA0C25B2B0D:8EAA6059FFA2D679FDAFF929C1E529DA" box="[516,720,1736,1762]" country="China" county="Nujiang Prefecture" municipality="Lushui" name="Pianma Township" pageId="12" pageNumber="356" stateProvince="Yunnan">Pianma Township</location>
,
<date id="FFCB1042FFA2D679FD71F929C07629DB" box="[730,835,1736,1763]" pageId="12" pageNumber="356" value="2015-07">
<collectingDate id="EF8FE9AAFFA2D679FD71F929C07629DB" box="[730,835,1736,1763]" pageId="12" pageNumber="356" value="2015-07">VII.2015</collectingDate>
</date>
,
<quantity id="4C8D9B67FFA2D679FCE5F929C0AA29DB" box="[846,927,1736,1763]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="356" unit="m" value="2500.0">
<elevation id="0058D1B1FFA2D679FCE5F929C0AA29DB" box="[846,927,1736,1763]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="356" unit="m" value="2500.0">2500m</elevation>
</quantity>
,
<collectorName id="26805354FFA2D679FC01F929C76D29DA" box="[938,1112,1736,1762]" pageId="12" pageNumber="356">Xinxiong Deng</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFA2D678FF6CF90DC2B82CF1" lastPageId="13" lastPageNumber="357" pageId="12" pageNumber="356" type="description">
<paragraph id="8BCA3682FFA2D679FF6CF90DC160283D" blockId="12.[151,1437,1556,2014]" box="[199,597,1771,1798]" pageId="12" pageNumber="356">
<emphasis id="B901EA90FFA2D679FF6CF90DC160283D" bold="true" box="[199,597,1771,1798]" pageId="12" pageNumber="356">Redescription of the male (n=18).</emphasis>
</paragraph>
<paragraph id="8BCA3682FFA2D679FF6CF8F1C7B92812" blockId="12.[151,1437,1556,2014]" box="[199,1164,1808,1834]" pageId="12" pageNumber="356">
Body length:
<quantity id="4C8D9B67FFA2D679FECBF8F1C1362812" box="[352,515,1808,1834]" metricMagnitude="-2" metricUnit="m" metricValue="3.875" metricValueMax="4.81" metricValueMin="2.94" pageId="12" pageNumber="356" unit="mm" value="38.75" valueMax="48.1" valueMin="29.4">29.448.1 mm</quantity>
(
<bibRefCitation id="EFE44B73FFA2D679FDBBF8F1C1A22812" author="Fujita, H." box="[528,663,1808,1834]" pageId="12" pageNumber="356" refId="ref7425" refString="Fujita, H. (2010) The Lucanid Beetles of the World. Mushi-Sha, Tokyo, 472 pp., 248 pls." type="book" year="2010">Fujita 2010</bibRefCitation>
;
<bibRefCitation id="EFE44B73FFA2D679FD0AF8F1C0662812" author="Toshiyuki, T." box="[673,851,1808,1834]" pageId="12" pageNumber="356" pagination="1 - 87" refId="ref7955" refString="Toshiyuki, T. (2021) Illustrated guide to the genus Neolucanus of the world. Be-kuwa, 78, 1 - 87." type="journal article" year="2021">Toshiyuki 2021</bibRefCitation>
;
<bibRefCitation id="EFE44B73FFA2D679FCF6F8F1C0822812" author="Yi, D." box="[861,951,1808,1834]" pageId="12" pageNumber="356" refId="ref7980" refString="Yi, D. (2023) Lucanidae of the World. Design Spacetime, Seoul, 190 pp., 160 pls." type="book" year="2023">Yi 2023</bibRefCitation>
and present paper)
</paragraph>
<paragraph id="8BCA3682FFA2D679FF6CF8D5C26728AE" blockId="12.[151,1437,1556,2014]" pageId="12" pageNumber="356">
<emphasis id="B901EA90FFA2D679FF6CF8D5C2022876" bold="true" box="[199,311,1844,1870]" pageId="12" pageNumber="356">Telodont.</emphasis>
<emphasis id="B901EA90FFA2D679FEEBF8D5C2AC2876" box="[320,409,1844,1870]" italics="true" pageId="12" pageNumber="356">Habitus</emphasis>
(
<figureCitation id="134E2A07FFA2D679FE01F8D5C1162876" box="[426,547,1844,1870]" captionStart="FIGURE 1" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[312,1275,185,1877]" captionTargetId="figure-17@3.[306,1281,179,1883]" captionTargetPageId="3" captionText="FIGURE 1. Male habitus of Neolucanus species. AB, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype, 36.0 mm, telodont (Lvchun, Yunnan); CE, N. fuscus Didier, 1926; CD, 50.9 mm, telodont (Yên Bái, Yên Bái); E, 44.8 mm, telodont (Ziyuan, Guangxi); FH, N. brochieri Bomans &amp; Miyashita, 1997; FG, 41.1mm, telodont (Gonshan, Yunnan); H, 41.0 mm, telodont (Gonshan, Yunnan). A, C, E, F, H, dorsal view; B, D, G, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735137" httpUri="https://zenodo.org/record/14735137/files/figure.png" pageId="12" pageNumber="356">Fig. 1FH</figureCitation>
). Head and pronotum black, elytra black or aurantiaceous with black borders; obviously glossy; body with invisible pubescence in dorsal view, labium with dense setae all legs with sparse yellow setae on surface.
</paragraph>
<paragraph id="8BCA3682FFA2D678FF6CF841C21E2E79" blockId="12.[151,1437,1556,2014]" lastBlockId="13.[151,1437,151,1077]" lastPageId="13" lastPageNumber="357" pageId="12" pageNumber="356">
<emphasis id="B901EA90FFA2D679FF6CF841C2312882" box="[199,260,1952,1978]" italics="true" pageId="12" pageNumber="356">Head</emphasis>
transverse. Anterior ridge raised behind mandible and widely concave on the middle. Canthus long and wide, completely surround the eye. Clypeolabrum (Fig. 3M) tiny and transverse, slightly visible in dorsal view, fused with frons. Mandible (Fig. 3MN) smooth from base to teeth on inner margin; in lateral view, clearly up-curved and thinnest at basal 1/3, a dorsal tooth behind apex forming an approximately arc-shaped and weakly protrudes beyond upper margin; in dorsal view, slightly curved, thickness almost regular from base to inner teeth, with 23 continuous small teeth. Antennal club with 3 antennomeres; antennomeres 89 subquadrilateral; antennomere 10 semi-elliptic.
</paragraph>
<paragraph id="8BCA3682FFA3D678FF6CFEAAC2032E95" blockId="13.[151,1437,151,1077]" pageId="13" pageNumber="357">
<emphasis id="B901EA90FFA3D678FF6CFEAAC2032E5C" box="[199,310,331,356]" italics="true" pageId="13" pageNumber="357">Pronotum</emphasis>
(Fig. 3O) transverse, markedly longer and wider than head with slightly raised edges.Anterior margin bisinuate, posterior margin dimly bisinuate. Anterior angles blunt and directed forwards, lateral angles and posterior angles obtuse.
</paragraph>
<paragraph id="8BCA3682FFA3D678FF6CFE56C7302EE9" blockId="13.[151,1437,151,1077]" box="[199,1029,439,465]" pageId="13" pageNumber="357">
<emphasis id="B901EA90FFA3D678FF6CFE56C2382EE9" box="[199,269,439,465]" italics="true" pageId="13" pageNumber="357">Elytra</emphasis>
elongate, anterior margin of elytra clearly narrower than pronotum.
</paragraph>
<paragraph id="8BCA3682FFA3D678FF6CFE3AC0072D05" blockId="13.[151,1437,151,1077]" pageId="13" pageNumber="357">
<emphasis id="B901EA90FFA3D678FF6CFE3AC3C92ECC" box="[199,252,475,500]" italics="true" pageId="13" pageNumber="357">Legs</emphasis>
. Protibia (Fig. 3P) with 46 conspicuous sharp and short protuberances along outer margin, apex bifurcate with branches moderately sharp at tip; inner margin with a unclear gap at base. Mesotibia and metatibia straight and outer margin with a terminal spine, without external spines.
</paragraph>
<paragraph id="8BCA3682FFA3D678FF6CFDA6C15B2C01" blockId="13.[151,1437,151,1077]" pageId="13" pageNumber="357">
<emphasis id="B901EA90FFA3D678FF6CFDA6C2D62D59" box="[199,483,583,609]" italics="true" pageId="13" pageNumber="357">Abdomen and aedeagus.</emphasis>
Abdominal tergite VIII (Fig. 5M) subsemi-circular, lateral angles poorly defined. Abdominal ventrite VIII (Fig. 5N) with smooth basal margin. Ventral plate of abdominal segment IX (Fig. 5P) slender without membranous stripe, obviously constricted at base, then widens sharply and significantly, widens gradually to about apical 1/5. Aedeagus (
<figureCitation id="134E2A07FFA3D678FDCFFD52C1CB2DF5" box="[612,766,691,717]" captionStart="FIGURE 6" captionStartId="8.[151,250,1909,1933]" captionTargetBox="[180,1408,181,1879]" captionTargetId="figure-18@8.[174,1414,181,1885]" captionTargetPageId="8" captionText="FIGURE 6. Aedeagus (not including flagellum) of Neolucanus species. AB, EF, IJ, N. horizontalis Xin, Zhong &amp; Qi, new species A, E, I, holotype (Lvchun, Yunnan); B, F, J, paratype (Jinping, Yunnan). C, G, K, N. fuscus Didier, 1926 (Ziyuan, Guangxi). D, H, L, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). AD, dorsal view; EH, ventral view; IL, lateral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735143" httpUri="https://zenodo.org/record/14735143/files/figure.png" pageId="13" pageNumber="357">Fig. 6D, H, L</figureCitation>
) with a significant and relatively slender pair of sclerotized dorsal plates; ventral plate at apical end of basal piece membranous. Paramere relatively narrow, with a obvious curvature at base, apex of paramere curved outward. Penis large, almost as long as paramere in ventral view. Flagellum long, apex of flagellum simple.
</paragraph>
<paragraph id="8BCA3682FFA3D678FF6CFCA3C2B82CF1" blockId="13.[151,1437,151,1077]" pageId="13" pageNumber="357">
<emphasis id="B901EA90FFA3D678FF6CFCA3C2F42C65" bold="true" box="[199,449,834,861]" pageId="13" pageNumber="357">Variation. Mesodont.</emphasis>
Mandible shorter and straighter in dorsal view, with a rounded tooth at base.
<emphasis id="B901EA90FFA3D678FAAEFCA3C6A92C64" bold="true" box="[1285,1436,834,860]" pageId="13" pageNumber="357">Amphiodont</emphasis>
and
<emphasis id="B901EA90FFA3D678FF63FC87C2052CB8" bold="true" box="[200,304,870,896]" pageId="13" pageNumber="357">priodont</emphasis>
(
<figureCitation id="134E2A07FFA3D678FEE9FC86C2832CB9" box="[322,438,871,897]" captionStart="FIGURE 2" captionStartId="4.[151,250,1771,1795]" captionTargetBox="[158,1429,186,1740]" captionTargetId="figure-18@4.[151,1436,181,1747]" captionTargetPageId="4" captionText="FIGURE 2. Male habitus of Neolucanus species. AB, N. horizontalis Xin, Zhong &amp; Qi, new species, paratype, 33.7 mm, mesodont (Jinping, Yunnan); CD, N. fuscus Didier, 1926, 34.7mm, priodont (Wuming, Guangxi); EF, N. brochieri Bomans &amp; Miyashita, 1997, 31.4 mm, priodont (Gonshan, Yunnan). A, C, E, dorsal view; B, D, F, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735139" httpUri="https://zenodo.org/record/14735139/files/figure.png" pageId="13" pageNumber="357">Fig. 2EF</figureCitation>
). Mandible (
<figureCitation id="134E2A07FFA3D678FDE1FC86C1872CB9" box="[586,690,871,897]" captionStart="FIGURE 4" captionStartId="6.[151,250,1582,1606]" captionTargetBox="[158,1429,188,1551]" captionTargetId="figure-18@6.[151,1436,181,1558]" captionTargetPageId="6" captionText="FIGURE 4. Male fore-head (AB, EF, IJ), pronotum (C, G, K), protibia and apical part of profemur (D, H, L) of Neolucanus species.AD, N. horizontalis Xin, Zhong &amp; Qi, new species, paratype (Jinping, Yunnan); EH, N. fuscus Didier, 1926 (Wuming, Guangxi); IL, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, dorsal view; B, F, J, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14735141" httpUri="https://zenodo.org/record/14735141/files/figure.png" pageId="13" pageNumber="357">Fig. 4IJ</figureCitation>
) slender, straight and only slightly longer than head with unclear consecutive small teeth behind apex, with almost no branch at apex in lateral view; clypeolabrum (
<figureCitation id="134E2A07FFA3D678FB58FC6AC6742C9D" box="[1267,1345,907,933]" captionStart="FIGURE 4" captionStartId="6.[151,250,1582,1606]" captionTargetBox="[158,1429,188,1551]" captionTargetId="figure-18@6.[151,1436,181,1558]" captionTargetPageId="6" captionText="FIGURE 4. Male fore-head (AB, EF, IJ), pronotum (C, G, K), protibia and apical part of profemur (D, H, L) of Neolucanus species.AD, N. horizontalis Xin, Zhong &amp; Qi, new species, paratype (Jinping, Yunnan); EH, N. fuscus Didier, 1926 (Wuming, Guangxi); IL, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, dorsal view; B, F, J, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14735141" httpUri="https://zenodo.org/record/14735141/files/figure.png" pageId="13" pageNumber="357">Fig. 4I</figureCitation>
) clearly visible in dorsal view.
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFA3D678FF6CFC33C1972CD5" box="[199,674,978,1005]" pageId="13" pageNumber="357" type="diagnosis">
<paragraph id="8BCA3682FFA3D678FF6CFC33C1972CD5" blockId="13.[151,1437,151,1077]" box="[199,674,978,1005]" pageId="13" pageNumber="357">
<emphasis id="B901EA90FFA3D678FF6CFC33C2FB2CD4" bold="true" box="[199,462,978,1004]" pageId="13" pageNumber="357">Differential diagnosis.</emphasis>
See
<tableCitation id="C6F70339FFA3D678FDA8FC32C1632CD5" box="[515,598,979,1005]" captionStart="TABLE 1" captionStartId="14.[151,239,728,752]" captionText="TABLE 1. Male main differences of external characters of Neolucanus horizontalis Xin, Zhong &amp; Qi, new species and its related species." pageId="13" pageNumber="357">Table 1</tableCitation>
and
<tableCitation id="C6F70339FFA3D678FD26FC32C1AB2CD5" box="[653,670,979,1005]" captionStart="TABLE 2" captionStartId="15.[151,239,151,175]" captionText="TABLE 2. Main differences of male abdomen and aedeagus of Neolucanus horizontalis Xin, Zhong &amp; Qi, new species and its related species." pageId="13" pageNumber="357">2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFA3D678FF6CFC17C25B2B0D" pageId="13" pageNumber="357" type="distribution">
<paragraph id="8BCA3682FFA3D678FF6CFC17C25B2B0D" blockId="13.[151,1437,151,1077]" pageId="13" pageNumber="357">
<emphasis id="B901EA90FFA3D678FF6CFC17C2542B28" bold="true" box="[199,353,1014,1040]" pageId="13" pageNumber="357">Distribution.</emphasis>
<collectingCountry id="F3627612FFA3D678FEC3FC16C29E2B29" box="[360,427,1015,1041]" name="China" pageId="13" pageNumber="357">China</collectingCountry>
(
<collectingRegion id="49B1F860FFA3D678FE12FC16C1202B29" box="[441,533,1015,1041]" country="China" name="Yunnan" pageId="13" pageNumber="357">Yunnan</collectingRegion>
);
<collectingCountry id="F3627612FFA3D678FD82FC16C1A32B29" box="[553,662,1015,1041]" name="Myanmar" pageId="13" pageNumber="357">Myanmar</collectingCountry>
(
<collectingRegion id="49B1F860FFA3D678FD0FFC16C1CD2B29" box="[676,760,1015,1041]" country="Myanmar" name="Kachin" pageId="13" pageNumber="357">Kachin</collectingRegion>
) (
<bibRefCitation id="EFE44B73FFA3D678FCA5FC16C0A02B29" author="Fujita, H." box="[782,917,1015,1041]" pageId="13" pageNumber="357" refId="ref7425" refString="Fujita, H. (2010) The Lucanid Beetles of the World. Mushi-Sha, Tokyo, 472 pp., 248 pls." type="book" year="2010">Fujita 2010</bibRefCitation>
;
<bibRefCitation id="EFE44B73FFA3D678FC0BFC16C7012B29" author="Schenk, K. D." box="[928,1076,1015,1041]" pageId="13" pageNumber="357" pagination="4 - 38" refId="ref7844" refString="Schenk, K. D. (2012) Notes to the Lucanidae of Asia and description of new taxa of the genus Neolucanus. Beetles World, 6, 4 - 38." type="journal article" year="2012">Schenk 2012</bibRefCitation>
,
<bibRefCitation id="EFE44B73FFA3D678FBEBFC16C7492B29" author="Schenk, K. D." box="[1088,1148,1015,1041]" pageId="13" pageNumber="357" pagination="2 - 8" refId="ref7878" refString="Schenk, K. D. (2014) Catalogue of Lucanidae: Genus Neolucanus Thomson, 1862. Beetles World, 10, 2 - 8." type="journal article" year="2014">2014</bibRefCitation>
;
<bibRefCitation id="EFE44B73FFA3D678FB2DFC16C60D2B29" author="Toshiyuki, T." box="[1158,1336,1015,1041]" pageId="13" pageNumber="357" pagination="1 - 87" refId="ref7955" refString="Toshiyuki, T. (2021) Illustrated guide to the genus Neolucanus of the world. Be-kuwa, 78, 1 - 87." type="journal article" year="2021">Toshiyuki 2021</bibRefCitation>
;
<bibRefCitation id="EFE44B73FFA3D678FAE8FC16C6A92B29" author="Yi, D." box="[1347,1436,1015,1041]" pageId="13" pageNumber="357" refId="ref7980" refString="Yi, D. (2023) Lucanidae of the World. Design Spacetime, Seoul, 190 pp., 160 pls." type="book" year="2023">Yi 2023</bibRefCitation>
and present paper).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,627 @@
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<mods:title id="1D043243A19DC89F55980BD126046443">A taxonomic note on Chinese species of the Neolucanus fuscus group with the description of a new species (Coleoptera: Lucanidae: Lucaninae)</mods:title>
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<mods:role id="C064A4F92EE27E6AD4C7A585FC8BEC00">
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<mods:namePart id="F39DFA16FEB19493B6B8D7C0CA2A4D2D">Xin, Fei-Yi</mods:namePart>
<mods:nameIdentifier id="197B962391C615B7DB3E439A37CE2F17" type="ORCID">0009-0002-4407-694X</mods:nameIdentifier>
<mods:affiliation id="160898452FCC4A17E2F9C9620D89EFA9">Fujian Colleges and Universities Engineering Research Institute of Conservation and Utilization of Natural Bioresources, Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Key Laboratory of Integrated Pest Management in Ecological Forests, Fujian Province University, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China.</mods:affiliation>
</mods:name>
<mods:name id="2805478CE27B121FFC47735E01637B38" type="personal">
<mods:role id="7BADEC5C33863255AD0427A364AE6E17">
<mods:roleTerm id="EE7C9A8CCDEE225B216C0F94185AC887">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F5ECFDA05BB5F91A76BFB1C02EE9C452">Zhong, Xiao-Tian</mods:namePart>
<mods:nameIdentifier id="85C93D45CBE73158D551CCC503EFC0D7" type="ORCID">0009-0003-1423-9274</mods:nameIdentifier>
<mods:affiliation id="12E3EB8E07096F2E32025163CF703BB2">School of Geoscience and Technology, Southwest Petroleum University, Chengdu 610500, China.</mods:affiliation>
</mods:name>
<mods:name id="D72AF184FA664415DB3FD0EC8199B555" type="personal">
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<mods:roleTerm id="6C9A2E140170A77C8C0BDA47C078C1F0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A11B10F848D1550FBE5D9F3BEFA1190D">Guo, Wen-Chi</mods:namePart>
<mods:nameIdentifier id="7E8AEDFBBFBB7F4B8083FF5C8FE3CF0F" type="ORCID">0009-0008-4116-4016</mods:nameIdentifier>
<mods:affiliation id="4BD8547D69524CE176ADE63AFD644922">Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China.</mods:affiliation>
<mods:nameIdentifier id="17D712B3C1432311F263BD37921FAE98" type="email">353600225@qq.com</mods:nameIdentifier>
</mods:name>
<mods:name id="9A76BA655E7A694C1B164E04774A87E6" type="personal">
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<mods:roleTerm id="14E4023D8DD7EB3C7762FCD8972F5C1E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A3D64AA6B502A84E252F58C6EBE251A3">Zhang, Yi-Feng</mods:namePart>
<mods:nameIdentifier id="7266B4C42A07D055B3B5787C8FCA8E0B" type="ORCID">0000-0003-3264-9472</mods:nameIdentifier>
<mods:affiliation id="F7DCC2462B3009FFD5E5DC07BCDA5077">Fujian Academy of Forestry, Fuzhou 350012, China.</mods:affiliation>
<mods:nameIdentifier id="145A8FD94FC1478EEB2BE3908FC0707C" type="email">zhang_yifeng@foxmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="89B6009AEEEBE49EB3175308C5E3E1BD" type="personal">
<mods:role id="FFC403D8C94BD670E97D6F91ED58CFB1">
<mods:roleTerm id="7D425DD34155806DF352BFEF77550505">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8E9C076CD9597BDED8A89893EBDBA317">Qi, Zhi-Hao</mods:namePart>
<mods:nameIdentifier id="6C320BD91FFBC715248B6304D3A26786" type="ORCID">0000-0001-8596-0913</mods:nameIdentifier>
<mods:affiliation id="51E0C6DDAE435D76E79BFFADCB89DED0">Key Laboratory of Integrated Pest Management in Ecological Forests, Fujian Province University, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Fujian Academy of Forestry, Fuzhou 350012, China.</mods:affiliation>
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</mods:role>
<mods:namePart id="7693BA36907AAF86785A22FC52CE3F79">Wu, Song-Qing</mods:namePart>
<mods:nameIdentifier id="5B1BDD88E966ABAB952E37864CE1BEAF" type="ORCID">0000-0002-2763-5378</mods:nameIdentifier>
<mods:affiliation id="DE7FA07C2B5A9D555EF19A11A4BB5BD1">Fujian Colleges and Universities Engineering Research Institute of Conservation and Utilization of Natural Bioresources, Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Key Laboratory of Integrated Pest Management in Ecological Forests, Fujian Province University, Fujian Agriculture and Forestry University, Fuzhou 350002, China. &amp; Forestry College, Fujian Agriculture and Forestry University, Fuzhou 350002, China.</mods:affiliation>
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<mods:title id="130EB74649978BF47AC1CE1555018296">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="73E5DCCA61B4911288190C6CA0AF0A9A">
<mods:date id="A33F8A231DF49A1A9CAE1981DEB7CD58">2025</mods:date>
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<mods:number id="F06AB9F641EF2EF9E4AF580F9EDA8599">2025-01-17</mods:number>
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<mods:url id="53ADD3BD8BB0F2DF209DB2550B1351CD">https://doi.org/10.11646/zootaxa.5570.2.6</mods:url>
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<mods:identifier id="E1E051035CCDBAD65C6C6E90217E5A84" type="DOI">10.11646/zootaxa.5570.2.6</mods:identifier>
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<mods:identifier id="C3F0E0FC3609AC4A3F13E7F2E1F30BF2" type="Zenodo-Dep">14735135</mods:identifier>
<mods:identifier id="D021EC5441BC84B4208881A834BE2FEF" type="ZooBank">8B1BAE9E-FBD3-4AC3-9FA3-9CCC2AA81392</mods:identifier>
</mods:mods>
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<subSubSection id="C36F6509FFACD677FF3CFF76C1602FEB" pageId="2" pageNumber="346" type="nomenclature">
<paragraph id="8BCA3682FFACD677FF3CFF76C00C2F89" blockId="2.[151,879,151,250]" box="[151,825,151,178]" pageId="2" pageNumber="346">
<heading id="D08281EEFFACD677FF3CFF76C00C2F89" bold="true" box="[151,825,151,178]" fontSize="11" level="1" pageId="2" pageNumber="346" reason="1">
<emphasis id="B901EA90FFACD677FF3CFF76C00C2F89" bold="true" box="[151,825,151,178]" pageId="2" pageNumber="346">
<taxonomicName id="4C754D01FFACD677FF3CFF76C1AE2F89" authority="Xin, Zhong &amp; Qi" authorityName="Xin, Zhong &amp; Qi" authorityYear="2025" box="[151,667,151,178]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="346" phylum="Arthropoda" rank="species" species="horizontalis" status="sp. nov.">
<emphasis id="B901EA90FFACD677FF3CFF76C28A2F8A" bold="true" box="[151,447,151,178]" italics="true" pageId="2" pageNumber="346">Neolucanus horizontalis</emphasis>
Xin, Zhong &amp; Qi
</taxonomicName>
,
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<paragraph id="8BCA3682FFACD677FF3CFF5AC1602FEB" blockId="2.[151,879,151,250]" box="[151,597,187,214]" pageId="2" pageNumber="346">
Chinese common name:
<emphasis id="B901EA90FFACD677FE06FF5DC1602FEB" bold="true" box="[429,597,188,211]" pageId="2" pageNumber="346">平®圆OiNJƤ</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFACD677FF3CFF3EC05A2FC1" box="[151,879,223,250]" pageId="2" pageNumber="346" type="description">
<paragraph id="8BCA3682FFACD677FF3CFF3EC05A2FC1" blockId="2.[151,879,151,250]" box="[151,879,223,250]" pageId="2" pageNumber="346">
<figureCitation id="134E2A07FFACD677FF3CFF3EC2292FC1" box="[151,284,223,249]" captionStart="FIGURE 1" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[312,1275,185,1877]" captionTargetId="figure-17@3.[306,1281,179,1883]" captionTargetPageId="3" captionText="FIGURE 1. Male habitus of Neolucanus species. AB, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype, 36.0 mm, telodont (Lvchun, Yunnan); CE, N. fuscus Didier, 1926; CD, 50.9 mm, telodont (Yên Bái, Yên Bái); E, 44.8 mm, telodont (Ziyuan, Guangxi); FH, N. brochieri Bomans &amp; Miyashita, 1997; FG, 41.1mm, telodont (Gonshan, Yunnan); H, 41.0 mm, telodont (Gonshan, Yunnan). A, C, E, F, H, dorsal view; B, D, G, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735137" httpUri="https://zenodo.org/record/14735137/files/figure.png" pageId="2" pageNumber="346">Figs. 1AB</figureCitation>
;
<figureCitation id="134E2A07FFACD677FE8CFF3EC2582FC1" box="[295,365,223,249]" captionStart="FIGURE 2" captionStartId="4.[151,250,1771,1795]" captionTargetBox="[158,1429,186,1740]" captionTargetId="figure-18@4.[151,1436,181,1747]" captionTargetPageId="4" captionText="FIGURE 2. Male habitus of Neolucanus species. AB, N. horizontalis Xin, Zhong &amp; Qi, new species, paratype, 33.7 mm, mesodont (Jinping, Yunnan); CD, N. fuscus Didier, 1926, 34.7mm, priodont (Wuming, Guangxi); EF, N. brochieri Bomans &amp; Miyashita, 1997, 31.4 mm, priodont (Gonshan, Yunnan). A, C, E, dorsal view; B, D, F, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735139" httpUri="https://zenodo.org/record/14735139/files/figure.png" pageId="2" pageNumber="346">2AB</figureCitation>
; 3EH; 4AD; 5AH; 6AB, EF, IJ; 7AB
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFACD677FF3CFEC6C0492EE9" pageId="2" pageNumber="346" type="materials_examined">
<paragraph id="8BCA3682FFACD677FF3CFEC6C7A42E7A" blockId="2.[151,1437,295,1905]" box="[151,1169,295,322]" pageId="2" pageNumber="346">
<materialsCitation id="3B1D3CDFFFACD677FF3CFEC6C7B82E7A" box="[151,1165,295,322]" country="China" county="Lvchun County" location="Pinghe Town" municipality="Honghe Prefecture" pageId="2" pageNumber="346" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B901EA90FFACD677FF3CFEC6C2032E79" bold="true" box="[151,310,295,321]" pageId="2" pageNumber="346">
<typeStatus id="54CE8820FFACD677FF3CFEC6C3E72E79" box="[151,210,295,321]" pageId="2" pageNumber="346">Type</typeStatus>
locality.
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<collectingCountry id="F3627612FFACD677FE97FEC6C2B72E79" box="[316,386,295,321]" name="China" pageId="2" pageNumber="346">China</collectingCountry>
,
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,
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,
<collectingCounty id="62AB4E0EFFACD677FC90FEC6C0DE2E79" box="[827,1003,295,321]" pageId="2" pageNumber="346">Lvchun County</collectingCounty>
,
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</materialsCitation>
.
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CFEAAC0492EE9" blockId="2.[151,1437,295,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CFEAAC24C2E5D" bold="true" box="[199,377,331,357]" pageId="2" pageNumber="346">Type material.</emphasis>
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<typeStatus id="54CE8820FFACD677FE28FEAAC2C72E5D" box="[387,498,331,357]" pageId="2" pageNumber="346" type="holotype">Holotype</typeStatus>
,
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male (
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: IOZ (E) 211512),
<collectingCountry id="F3627612FFACD677FC3DFEAAC0E92E5D" box="[918,988,331,357]" name="China" pageId="2" pageNumber="346">China</collectingCountry>
,
<collectingRegion id="49B1F860FFACD677FC42FEADC79A2E5D" box="[1001,1199,331,358]" country="China" name="Yunnan" pageId="2" pageNumber="346">Yunnan Province</collectingRegion>
,
<collectingMunicipality id="6BAEACF8FFACD677FB14FEAAC6AD2E5D" box="[1215,1432,331,357]" pageId="2" pageNumber="346">Honghe Prefecture</collectingMunicipality>
,
<collectingCounty id="62AB4E0EFFACD677FF3CFE8EC2792EB1" box="[151,332,367,393]" pageId="2" pageNumber="346">Lvchun County</collectingCounty>
,
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,
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,
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,
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<collectingDate id="EF8FE9AAFFACD677FCECFE8EC08E2EB1" box="[839,955,367,393]" pageId="2" pageNumber="346" value="2024-06-08">8.VI.2024</collectingDate>
</date>
,
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, local collector leg
</materialsCitation>
.
<materialsCitation id="3B1D3CDFFFACD677FAA4FE8EC04D2EE9" collectingDate="2023-06-02" collectionCode="CXTZ" country="China" county="Honghe Prefecture" elevation="2400" latitude="22.756666" location="Ma'andi Township" longLatPrecision="21" longitude="103.43917" municipality="Jinping county" pageId="2" pageNumber="346" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
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<typeStatus id="54CE8820FFACD677FAA4FE8EC64B2EB1" box="[1295,1406,367,393]" pageId="2" pageNumber="346" type="paratype">Paratype</typeStatus>
,
</emphasis>
<specimenCount id="9D73FD0BFFACD677FA25FE8EC3F82E95" count="1" pageId="2" pageNumber="346" type="male">1 male</specimenCount>
(
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),
<collectingCountry id="F3627612FFACD677FEE8FE72C2BC2E95" box="[323,393,403,429]" name="China" pageId="2" pageNumber="346">China</collectingCountry>
,
<collectingRegion id="49B1F860FFACD677FE3DFE75C16E2E95" box="[406,603,403,430]" country="China" name="Yunnan" pageId="2" pageNumber="346">Yunnan Province</collectingRegion>
,
<collectingCounty id="62AB4E0EFFACD677FDC2FE72C0742E95" box="[617,833,403,429]" pageId="2" pageNumber="346">Honghe Prefecture</collectingCounty>
,
<collectingMunicipality id="6BAEACF8FFACD677FCE5FE72C0C22E96" box="[846,1015,403,430]" pageId="2" pageNumber="346">Jinping county</collectingMunicipality>
,
<location id="8EAA6059FFACD677FBAEFE72C7EB2E95" LSID="urn:lsid:plazi:treatment:03DC8794FFACD672FF3CFF76C62A28B3:8EAA6059FFACD677FBAEFE72C7EB2E95" box="[1029,1246,403,429]" country="China" county="Honghe Prefecture" latitude="22.756666" longLatPrecision="21" longitude="103.43917" municipality="Jinping county" name="Ma'andi Township" pageId="2" pageNumber="346" stateProvince="Yunnan">Maandi Township</location>
,
<location id="8EAA6059FFACD677FB47FE72C6AD2E95" LSID="urn:lsid:plazi:treatment:03DC8794FFACD672FF3CFF76C62A28B3:8EAA6059FFACD677FB47FE72C6AD2E95" box="[1260,1432,403,429]" country="China" county="Honghe Prefecture" latitude="22.756666" longLatPrecision="21" longitude="103.43917" municipality="Jinping county" name="Sitai Mountain" pageId="2" pageNumber="346" stateProvince="Yunnan">Sitai Mountain</location>
,
<geoCoordinate id="EE415045FFACD677FF3CFE56C2162EEA" box="[151,291,439,466]" degrees="22" direction="north" minutes="45" orientation="latitude" pageId="2" pageNumber="346" precision="15" seconds="24" value="22.756666">22°4524”N</geoCoordinate>
,
<geoCoordinate id="EE415045FFACD677FE85FE56C2F02EE9" box="[302,453,439,465]" degrees="103" direction="east" minutes="26" orientation="longitude" pageId="2" pageNumber="346" precision="15" seconds="21" value="103.43917">103°2621”E</geoCoordinate>
,
<quantity id="4C8D9B67FFACD677FE7BFE56C1142EE9" box="[464,545,439,465]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="346" unit="m" value="2400.0">
<elevation id="0058D1B1FFACD677FE7BFE56C1142EE9" box="[464,545,439,465]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="346" unit="m" value="2400.0">2400m</elevation>
</quantity>
,
<date id="FFCB1042FFACD677FD87FE56C1952EE9" box="[556,672,439,465]" pageId="2" pageNumber="346" value="2023-06-02">
<collectingDate id="EF8FE9AAFFACD677FD87FE56C1952EE9" box="[556,672,439,465]" pageId="2" pageNumber="346" value="2023-06-02">2.VI.2023</collectingDate>
</date>
, local collector leg
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFACD677FF6CFE3AC29729A1" pageId="2" pageNumber="346" type="description">
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<emphasis id="B901EA90FFACD677FF6CFE3AC16D2ECD" bold="true" box="[199,600,475,501]" pageId="2" pageNumber="346">
Description of the
<typeStatus id="54CE8820FFACD677FE08FE3AC13E2ECD" box="[419,523,475,501]" pageId="2" pageNumber="346" type="holotype">holotype</typeStatus>
, male.
</emphasis>
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CFDE1C63B2D05" blockId="2.[151,1437,295,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CFDE1C2442D21" box="[199,369,512,537]" italics="true" pageId="2" pageNumber="346">Measurements.</emphasis>
Body length (36.0 mm). Length of particular body parts: head (
<quantity id="4C8D9B67FFACD677FBE1FE1EC7922D21" box="[1098,1191,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="2" pageNumber="346" unit="mm" value="3.8">3.8 mm</quantity>
), mandible (
<quantity id="4C8D9B67FFACD677FA9FFE1EC6A52D21" box="[1332,1424,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="2" pageNumber="346" unit="mm" value="8.2">8.2 mm</quantity>
), pronotum (7.0 mm), elytra (
<quantity id="4C8D9B67FFACD677FE7BFDC2C10C2D05" box="[464,569,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="2" pageNumber="346" unit="mm" value="16.3">16.3 mm</quantity>
). Width: head (
<quantity id="4C8D9B67FFACD677FD4FFDC2C00A2D05" box="[740,831,547,573]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="2" pageNumber="346" unit="mm" value="9.1">9.1 mm</quantity>
), pronotum (
<quantity id="4C8D9B67FFACD677FC65FDC2C7022D05" box="[974,1079,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="2" pageNumber="346" unit="mm" value="12.3">12.3 mm</quantity>
), elytra (
<quantity id="4C8D9B67FFACD677FB33FDC2C6372D05" box="[1176,1282,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="2" pageNumber="346" unit="mm" value="12.3">12.3 mm</quantity>
).
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CFDA6C0992DBD" blockId="2.[151,1437,295,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CFDA6C2152D59" box="[199,288,583,609]" italics="true" pageId="2" pageNumber="346">Habitus</emphasis>
(
<figureCitation id="134E2A07FFACD677FE84FDA6C29F2D59" box="[303,426,583,609]" captionStart="FIGURE 1" captionStartId="3.[151,250,1908,1932]" captionTargetBox="[312,1275,185,1877]" captionTargetId="figure-17@3.[306,1281,179,1883]" captionTargetPageId="3" captionText="FIGURE 1. Male habitus of Neolucanus species. AB, N. horizontalis Xin, Zhong &amp; Qi, new species, holotype, 36.0 mm, telodont (Lvchun, Yunnan); CE, N. fuscus Didier, 1926; CD, 50.9 mm, telodont (Yên Bái, Yên Bái); E, 44.8 mm, telodont (Ziyuan, Guangxi); FH, N. brochieri Bomans &amp; Miyashita, 1997; FG, 41.1mm, telodont (Gonshan, Yunnan); H, 41.0 mm, telodont (Gonshan, Yunnan). A, C, E, F, H, dorsal view; B, D, G, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735137" httpUri="https://zenodo.org/record/14735137/files/figure.png" pageId="2" pageNumber="346">Fig. 1AB</figureCitation>
). Head and pronotum black, elytra kermesinus, with slightly glossy; whole body with tiny and almost invisible pubescence, all legs with yellow setae on surface.
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CFD6EC6572CB9" blockId="2.[151,1437,295,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CFD6EC2312D91" box="[199,260,655,681]" italics="true" pageId="2" pageNumber="346">Head</emphasis>
transverse. Anterior ridge raised behind mandible and widely concave on the middle. Canthus long and wide, completely surround the eye. Clypeolabrum (Fig. 3E) tiny and subtriangle, visible in dorsal view, fused with frons, apex sharp. Mandible (Fig. 3EF) about 2.2 times longer than head, inner smooth from base to teeth; in lateral view, almost horizontal, a dorsal tooth behind apex forming an approximately vertical right-angle and slightly protrudes beyond upper margin; in dorsal view, slightly curved, base thick, becomes thicker firstly and then thinner from base to teeth, with 2 continuous subtriangle inner teeth behind apex, proximal one smaller, apex of mandible sharp. Antennal club with 3 antennomeres; antennomeres 89 subquadrilateral; antennomere 10 semi-elliptic.
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CFC6DC02B2CD5" blockId="2.[151,1437,295,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CFC6DC2032C9D" box="[199,310,908,933]" italics="true" pageId="2" pageNumber="346">Pronotum</emphasis>
(Fig. 3G) transverse, 1.8 times wider than long, markedly longer and wider than head with slightly raised edges. Anterior margin bisinuate, posterior margin slightly bisinuate. Anterior angles rounded and directed forwards, lateral angles obtuse and posterior angles blunt.
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CFC16C0932B29" blockId="2.[151,1437,295,1905]" box="[199,934,1015,1042]" pageId="2" pageNumber="346">
<taxonomicName id="4C754D01FFACD677FF6CFC16C2002B29" box="[199,309,1015,1041]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="2" pageNumber="346" phylum="Arthropoda" rank="genus">
<emphasis id="B901EA90FFACD677FF6CFC16C2002B29" box="[199,309,1015,1041]" italics="true" pageId="2" pageNumber="346">Scutellum</emphasis>
</taxonomicName>
subtriangular, punctate, width almost as long as length.
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CFBFDC3EF2B45" blockId="2.[151,1437,295,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CFBFDC3C92B0D" box="[199,252,1052,1077]" italics="true" pageId="2" pageNumber="346">Legs</emphasis>
. Protibia (Fig. 3H) with 2 conspicuous sharp protuberances along outer margin, apex bifurcate with branches moderately sharp at tip. Mesotibia and metatibia straight and smooth at outer margin with a terminal spine.
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CFB66C62D2B9A" blockId="2.[151,1437,295,1905]" box="[199,1304,1159,1186]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CFB66C2382B99" box="[199,269,1159,1185]" italics="true" pageId="2" pageNumber="346">Elytra</emphasis>
elongate, 1.3 times longer than wide, anterior margin significantly narrower than pronotum.
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CFB4AC11E2ADD" blockId="2.[151,1437,295,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CFB4AC2D12BFD" box="[199,484,1195,1221]" italics="true" pageId="2" pageNumber="346">Abdomen and aedeagus.</emphasis>
Abdominal tergite VIII (Fig. 5A) nearly semilune, lateral angles poorly defined. Abdominal ventrite VIII (Fig. 5B) with 2 slightly protruding tips on basal margin near middle. Ventral plate of abdominal segment IX (Fig. 5D) elongate without membranous stripe, basal part extremely narrow, then suddenly became wider, widens gradually to apex and constricted near apex. Aedeagus (
<figureCitation id="134E2A07FFACD677FBB8FAF6C7912A09" box="[1043,1188,1303,1330]" captionStart="FIGURE 6" captionStartId="8.[151,250,1909,1933]" captionTargetBox="[180,1408,181,1879]" captionTargetId="figure-18@8.[174,1414,181,1885]" captionTargetPageId="8" captionText="FIGURE 6. Aedeagus (not including flagellum) of Neolucanus species. AB, EF, IJ, N. horizontalis Xin, Zhong &amp; Qi, new species A, E, I, holotype (Lvchun, Yunnan); B, F, J, paratype (Jinping, Yunnan). C, G, K, N. fuscus Didier, 1926 (Ziyuan, Guangxi). D, H, L, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). AD, dorsal view; EH, ventral view; IL, lateral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735143" httpUri="https://zenodo.org/record/14735143/files/figure.png" pageId="2" pageNumber="346">Fig. 6A, E, I</figureCitation>
) in dorsal view about 3.2 times longer than wide. Basal piece elongated, weakly longer than parameres in dorsal view, with a significant and relatively slender pair of sclerotized dorsal plates, and almost as wide as paramere; ventral plate at apical end of basal piece membranous. Paramere relatively narrow, with a small inner curvature at base, apex of paramere curved outward. Penis about 0.9 times length of paramere in ventral view. Flagellum long, about 1.6 times longer than aedeagus, apex of flagellum simple.
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CFA0EC21B294D" blockId="2.[151,1437,295,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CFA0EC1912931" bold="true" box="[199,676,1519,1545]" pageId="2" pageNumber="346">
Variation of
<typeStatus id="54CE8820FFACD677FECEFA0EC2E52931" box="[357,464,1519,1545]" pageId="2" pageNumber="346" type="paratype">paratype</typeStatus>
(n=1). Mesodont
</emphasis>
(
<figureCitation id="134E2A07FFACD677FD1CFA0EC0012931" box="[695,820,1519,1545]" captionStart="FIGURE 2" captionStartId="4.[151,250,1771,1795]" captionTargetBox="[158,1429,186,1740]" captionTargetId="figure-18@4.[151,1436,181,1747]" captionTargetPageId="4" captionText="FIGURE 2. Male habitus of Neolucanus species. AB, N. horizontalis Xin, Zhong &amp; Qi, new species, paratype, 33.7 mm, mesodont (Jinping, Yunnan); CD, N. fuscus Didier, 1926, 34.7mm, priodont (Wuming, Guangxi); EF, N. brochieri Bomans &amp; Miyashita, 1997, 31.4 mm, priodont (Gonshan, Yunnan). A, C, E, dorsal view; B, D, F, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14735139" httpUri="https://zenodo.org/record/14735139/files/figure.png" pageId="2" pageNumber="346">Fig. 2AB</figureCitation>
). Mandible 1.8 times longer than head (
<figureCitation id="134E2A07FFACD677FABAFA0EC6BA2931" box="[1297,1423,1519,1545]" captionStart="FIGURE 4" captionStartId="6.[151,250,1582,1606]" captionTargetBox="[158,1429,188,1551]" captionTargetId="figure-18@6.[151,1436,181,1558]" captionTargetPageId="6" captionText="FIGURE 4. Male fore-head (AB, EF, IJ), pronotum (C, G, K), protibia and apical part of profemur (D, H, L) of Neolucanus species.AD, N. horizontalis Xin, Zhong &amp; Qi, new species, paratype (Jinping, Yunnan); EH, N. fuscus Didier, 1926 (Wuming, Guangxi); IL, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, dorsal view; B, F, J, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14735141" httpUri="https://zenodo.org/record/14735141/files/figure.png" pageId="2" pageNumber="346">Fig. 4AB</figureCitation>
), shorter and straighter in dorsal view, dorsal tooth almost not protrudes beyond upper margin in lateral view, with 3 indistinct smaller inner teeth and a large blunt tooth at base. Protibia (
<figureCitation id="134E2A07FFACD677FC14F9D6C7222969" box="[959,1047,1591,1617]" captionStart="FIGURE 4" captionStartId="6.[151,250,1582,1606]" captionTargetBox="[158,1429,188,1551]" captionTargetId="figure-18@6.[151,1436,181,1558]" captionTargetPageId="6" captionText="FIGURE 4. Male fore-head (AB, EF, IJ), pronotum (C, G, K), protibia and apical part of profemur (D, H, L) of Neolucanus species.AD, N. horizontalis Xin, Zhong &amp; Qi, new species, paratype (Jinping, Yunnan); EH, N. fuscus Didier, 1926 (Wuming, Guangxi); IL, N. brochieri Bomans &amp; Miyashita, 1997 (Gonshan, Yunnan). A, CD, E, GH, I, KL, dorsal view; B, F, J, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14735141" httpUri="https://zenodo.org/record/14735141/files/figure.png" pageId="2" pageNumber="346">Fig. 4D</figureCitation>
) with 3 sharp protuberances along outer margin.
</paragraph>
<paragraph id="8BCA3682FFACD677FF6CF99EC29729A1" blockId="2.[151,1437,295,1905]" box="[199,418,1663,1689]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CF99EC21029A1" bold="true" box="[199,293,1663,1689]" pageId="2" pageNumber="346">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFACD677FF6CF942C2E5283D" pageId="2" pageNumber="346" type="diagnosis">
<paragraph id="8BCA3682FFACD677FF6CF942C2E5283D" blockId="2.[151,1437,295,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CF942C2F82985" bold="true" box="[199,461,1699,1725]" pageId="2" pageNumber="346">Differential diagnosis.</emphasis>
The new species is similar to
<taxonomicName id="4C754D01FFACD677FCBCF945C7242985" authority="Didier, 1926" authorityName="Didier" authorityYear="1926" box="[791,1041,1699,1726]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="346" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B901EA90FFACD677FCBCF945C01F2985" box="[791,810,1700,1725]" italics="true" pageId="2" pageNumber="346">N</emphasis>
.
<emphasis id="B901EA90FFACD677FC9CF942C0482985" box="[823,893,1699,1725]" italics="true" pageId="2" pageNumber="346">fuscus</emphasis>
<bibRefCitation id="EFE44B73FFACD677FC28F942C7242985" author="Didier, R." box="[899,1041,1699,1725]" pageId="2" pageNumber="346" pagination="210 - 214" refId="ref7377" refString="Didier, R. (1926) Descriptions sommaires de Lucanides nouveaux, du genre Neolucanus, de la faune indo-chinoise. Bulletin de la Societe Entomologique de France, 1926, 210 - 214. https: // doi. org / 10.3406 / bsef. 1926.27736" type="journal article" year="1926">Didier, 1926</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C754D01FFACD677FBB0F945C3E729D9" authority="Bomans &amp; Miyashita, 1997" authorityName="Bomans &amp; Miyashita" authorityYear="1997" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="346" phylum="Arthropoda" rank="species" species="brochieri">
<emphasis id="B901EA90FFACD677FBB0F945C71B2985" box="[1051,1070,1700,1725]" italics="true" pageId="2" pageNumber="346">N</emphasis>
.
<emphasis id="B901EA90FFACD677FB90F942C7972985" box="[1083,1186,1699,1725]" italics="true" pageId="2" pageNumber="346">brochieri</emphasis>
<bibRefCitation id="EFE44B73FFACD677FB03F945C3E729D9" author="Bomans, H. E. &amp; Miyashita, T." pageId="2" pageNumber="346" pagination="2 - 3" refId="ref7344" refString="Bomans, H. E. &amp; Miyashita, T. (1997) Description de trois nouvelles especes de Lucanidae du nord Birmanie. Besoiro, 4, 2 - 3." type="journal article" year="1997">Bomans &amp; Miyashita, 1997</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C754D01FFACD677FF4BF929C2CD29DA" authority="Baba, 1995" authorityName="Baba" authorityYear="1995" box="[224,504,1735,1762]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="346" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B901EA90FFACD677FF4BF929C3C629D9" box="[224,243,1736,1761]" italics="true" pageId="2" pageNumber="346">N</emphasis>
.
<emphasis id="B901EA90FFACD677FEAFF926C25929D9" box="[260,364,1735,1761]" italics="true" pageId="2" pageNumber="346">lehmanni</emphasis>
<bibRefCitation id="EFE44B73FFACD677FEDDF926C2CD29DA" author="Baba, M." box="[374,504,1735,1762]" pageId="2" pageNumber="346" pagination="8 - 11" refId="ref7312" refString="Baba, M. (1995) A new species of the genus Neolucanus Tomson, 1862 from south Burma (Myanmar). Gekkan-Mushi, 292, 8 - 11." type="journal article" year="1995">
Baba,
<quantity id="4C8D9B67FFACD677FE6BF926C2CD29DA" box="[448,504,1735,1762]" metricMagnitude="1" metricUnit="m" metricValue="5.0673" pageId="2" pageNumber="346" unit="in" value="1995.0">1995</quantity>
</bibRefCitation>
</taxonomicName>
in general appearance, but the following set of characters could help to identify precisely (
<tableCitation id="C6F70339FFACD677FEA0F90AC26A283D" box="[267,351,1771,1797]" captionStart="TABLE 1" captionStartId="14.[151,239,728,752]" captionText="TABLE 1. Male main differences of external characters of Neolucanus horizontalis Xin, Zhong &amp; Qi, new species and its related species." pageId="2" pageNumber="346">Table 1</tableCitation>
,
<tableCitation id="C6F70339FFACD677FEC6F90AC2F6283D" box="[365,451,1771,1797]" captionStart="TABLE 2" captionStartId="15.[151,239,151,175]" captionText="TABLE 2. Main differences of male abdomen and aedeagus of Neolucanus horizontalis Xin, Zhong &amp; Qi, new species and its related species." pageId="2" pageNumber="346">Table 2</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFACD677FF6CF8EEC2082875" pageId="2" pageNumber="346" type="etymology">
<paragraph id="8BCA3682FFACD677FF6CF8EEC2082875" blockId="2.[151,1437,295,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CF8EEC27B2811" bold="true" box="[199,334,1807,1833]" pageId="2" pageNumber="346">Etymology.</emphasis>
This new species is named after the morphological features of its horizontal mandible when viewed in lateral view.
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFACD670FF6CF8B6C74D28C1" lastPageId="5" lastPageNumber="349" pageId="2" pageNumber="346" type="distribution">
<paragraph id="8BCA3682FFACD677FF6CF8B6C1142849" blockId="2.[151,1437,295,1905]" box="[199,545,1879,1905]" pageId="2" pageNumber="346">
<emphasis id="B901EA90FFACD677FF6CF8B6C2542849" bold="true" box="[199,353,1879,1905]" pageId="2" pageNumber="346">Distribution.</emphasis>
<collectingCountry id="F3627612FFACD677FECCF8B6C29F2849" box="[359,426,1879,1905]" name="China" pageId="2" pageNumber="346">China</collectingCountry>
(
<collectingRegion id="49B1F860FFACD677FE12F8B6C1202849" box="[441,533,1879,1905]" country="China" name="Yunnan" pageId="2" pageNumber="346">Yunnan</collectingRegion>
).
</paragraph>
<caption id="DF0A660AFFADD676FF3CF895C7AC28C0" ID-DOI="http://doi.org/10.5281/zenodo.14735137" ID-Zenodo-Dep="14735137" httpUri="https://zenodo.org/record/14735137/files/figure.png" pageId="3" pageNumber="347" startId="3.[151,250,1908,1932]" targetBox="[312,1275,185,1877]" targetPageId="3" targetType="figure">
<paragraph id="8BCA3682FFADD676FF3CF895C7AC28C0" blockId="3.[151,1436,1908,2041]" pageId="3" pageNumber="347">
<emphasis id="B901EA90FFADD676FF3CF895C22028B4" bold="true" box="[151,277,1908,1932]" pageId="3" pageNumber="347">FIGURE 1.</emphasis>
Male habitus of
<taxonomicName id="4C754D01FFADD676FE62F894C18128B4" box="[457,692,1908,1933]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="347" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="B901EA90FFADD676FE62F894C17428B5" box="[457,577,1909,1933]" italics="true" pageId="3" pageNumber="347">Neolucanus</emphasis>
species.
<emphasis id="B901EA90FFADD676FD0AF895C18128B4" bold="true" box="[673,692,1908,1932]" pageId="3" pageNumber="347">A</emphasis>
</taxonomicName>
<emphasis id="B901EA90FFADD676FD1FF895C1E228B4" bold="true" box="[692,727,1908,1932]" pageId="3" pageNumber="347">B,</emphasis>
<taxonomicName id="4C754D01FFADD676FD4BF894C70C28B5" authority="Xin, Zhong &amp; Qi" authorityName="Xin, Zhong &amp; Qi" authorityYear="2025" box="[736,1081,1909,1933]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="347" phylum="Arthropoda" rank="species" species="horizontalis" status="new species">
<emphasis id="B901EA90FFADD676FD4BF894C1C428B4" box="[736,753,1909,1932]" italics="true" pageId="3" pageNumber="347">N</emphasis>
.
<emphasis id="B901EA90FFADD676FCABF894C04C28B5" box="[768,889,1909,1933]" italics="true" pageId="3" pageNumber="347">horizontalis</emphasis>
Xin, Zhong &amp; Qi
</taxonomicName>
,
<emphasis id="B901EA90FFADD676FBEEF894C7F128B4" bold="true" box="[1093,1220,1908,1933]" pageId="3" pageNumber="347">
<taxonomicNameLabel id="A23257EBFFADD676FBEEF894C7F128B4" box="[1093,1220,1908,1933]" pageId="3" pageNumber="347" rank="species">new species</taxonomicNameLabel>
</emphasis>
, holotype, 36.0 mm, telodont (Lvchun, Yunnan);
<emphasis id="B901EA90FFADD676FE13F879C2DA2888" bold="true" box="[440,495,1944,1968]" pageId="3" pageNumber="347">CE,</emphasis>
<taxonomicName id="4C754D01FFADD676FE5DF878C1EA2889" authority="Didier, 1926" authorityName="Didier" authorityYear="1926" box="[502,735,1945,1969]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="347" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B901EA90FFADD676FE5DF878C1322888" box="[502,519,1945,1968]" italics="true" pageId="3" pageNumber="347">N</emphasis>
.
<emphasis id="B901EA90FFADD676FDBEF878C1612889" box="[533,596,1945,1969]" italics="true" pageId="3" pageNumber="347">fuscus</emphasis>
<bibRefCitation id="EFE44B73FFADD676FDF0F878C1EA2889" author="Didier, R." box="[603,735,1945,1969]" pageId="3" pageNumber="347" pagination="210 - 214" refId="ref7377" refString="Didier, R. (1926) Descriptions sommaires de Lucanides nouveaux, du genre Neolucanus, de la faune indo-chinoise. Bulletin de la Societe Entomologique de France, 1926, 210 - 214. https: // doi. org / 10.3406 / bsef. 1926.27736" type="journal article" year="1926">Didier, 1926</bibRefCitation>
</taxonomicName>
;
<emphasis id="B901EA90FFADD676FD41F879C0142888" bold="true" box="[746,801,1944,1968]" pageId="3" pageNumber="347">CD,</emphasis>
50.9 mm, telodont (Yên Bái, Yên Bái);
<emphasis id="B901EA90FFADD676FB69F878C7EC2889" bold="true" box="[1218,1241,1945,1969]" pageId="3" pageNumber="347">E,</emphasis>
44.8 mm, telodont (Ziyuan, Guangxi);
<emphasis id="B901EA90FFADD676FEC8F85CC2AC28ED" bold="true" box="[355,409,1981,2005]" pageId="3" pageNumber="347">FH,</emphasis>
<taxonomicName id="4C754D01FFADD676FE0AF85CC07728ED" authority="Bomans &amp; Miyashita, 1997" authorityName="Bomans &amp; Miyashita" authorityYear="1997" box="[417,834,1981,2005]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="347" phylum="Arthropoda" rank="species" species="brochieri">
<emphasis id="B901EA90FFADD676FE0AF85CC28728EC" box="[417,434,1981,2004]" italics="true" pageId="3" pageNumber="347">N</emphasis>
.
<emphasis id="B901EA90FFADD676FE14F85CC12928ED" box="[447,540,1981,2005]" italics="true" pageId="3" pageNumber="347">brochieri</emphasis>
<bibRefCitation id="EFE44B73FFADD676FD8FF85CC07728ED" author="Bomans, H. E. &amp; Miyashita, T." box="[548,834,1981,2005]" pageId="3" pageNumber="347" pagination="2 - 3" refId="ref7344" refString="Bomans, H. E. &amp; Miyashita, T. (1997) Description de trois nouvelles especes de Lucanidae du nord Birmanie. Besoiro, 4, 2 - 3." type="journal article" year="1997">Bomans &amp; Miyashita, 1997</bibRefCitation>
</taxonomicName>
;
<emphasis id="B901EA90FFADD676FCE6F85DC0B628EC" bold="true" box="[845,899,1980,2004]" pageId="3" pageNumber="347">FG,</emphasis>
41.1mm, telodont (Gonshan, Yunnan);
<emphasis id="B901EA90FFADD676FAB1F85CC60628ED" bold="true" box="[1306,1331,1981,2005]" pageId="3" pageNumber="347">H,</emphasis>
41.0 mm, telodont (Gonshan, Yunnan).
<emphasis id="B901EA90FFADD676FE6AF801C16528C1" bold="true" box="[449,592,2016,2041]" pageId="3" pageNumber="347">A, C, E, F, H,</emphasis>
dorsal view;
<emphasis id="B901EA90FFADD676FD71F800C00528C0" bold="true" box="[730,816,2016,2041]" pageId="3" pageNumber="347">B, D, G,</emphasis>
ventral view. Scale bar = 10.0 mm.
</paragraph>
</caption>
<caption id="DF0A660AFFAAD671FF3CF90AC6412872" ID-DOI="http://doi.org/10.5281/zenodo.14735139" ID-Zenodo-Dep="14735139" httpUri="https://zenodo.org/record/14735139/files/figure.png" pageId="4" pageNumber="348" startId="4.[151,250,1771,1795]" targetBox="[158,1429,186,1740]" targetPageId="4" targetType="figure">
<paragraph id="8BCA3682FFAAD671FF3CF90AC6412872" blockId="4.[151,1437,1771,1867]" pageId="4" pageNumber="348">
<emphasis id="B901EA90FFAAD671FF3CF90AC220283B" bold="true" box="[151,277,1771,1795]" pageId="4" pageNumber="348">FIGURE 2.</emphasis>
Male habitus of
<taxonomicName id="4C754D01FFAAD671FE61F90AC180283B" box="[458,693,1771,1795]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="348" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="B901EA90FFAAD671FE61F90AC177283B" box="[458,578,1771,1795]" italics="true" pageId="4" pageNumber="348">Neolucanus</emphasis>
species.
<emphasis id="B901EA90FFAAD671FD09F90AC180283B" bold="true" box="[674,693,1771,1795]" pageId="4" pageNumber="348">A</emphasis>
</taxonomicName>
<emphasis id="B901EA90FFAAD671FD1EF90AC1ED283B" bold="true" box="[693,728,1771,1795]" pageId="4" pageNumber="348">B,</emphasis>
<taxonomicName id="4C754D01FFAAD671FD4AF90DC70F283B" authority="Xin, Zhong &amp; Qi" authorityName="Xin, Zhong &amp; Qi" authorityYear="2025" box="[737,1082,1771,1795]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="348" phylum="Arthropoda" rank="species" species="horizontalis" status="new species">
<emphasis id="B901EA90FFAAD671FD4AF90DC1C7283B" box="[737,754,1772,1795]" italics="true" pageId="4" pageNumber="348">N</emphasis>
.
<emphasis id="B901EA90FFAAD671FCABF90AC04C283B" box="[768,889,1771,1795]" italics="true" pageId="4" pageNumber="348">horizontalis</emphasis>
Xin, Zhong &amp; Qi
</taxonomicName>
,
<emphasis id="B901EA90FFAAD671FBEDF90AC7F0283B" bold="true" box="[1094,1221,1771,1795]" pageId="4" pageNumber="348">
<taxonomicNameLabel id="A23257EBFFAAD671FBEDF90AC7F0283B" box="[1094,1221,1771,1795]" pageId="4" pageNumber="348" rank="species">new species</taxonomicNameLabel>
</emphasis>
, paratype, 33.7 mm, mesodont (Jinping, Yunnan);
<emphasis id="B901EA90FFAAD671FE14F8EEC2C2281F" bold="true" box="[447,503,1807,1831]" pageId="4" pageNumber="348">CD,</emphasis>
<taxonomicName id="4C754D01FFAAD671FE57F8F1C1EB281F" authority="Didier, 1926" authorityName="Didier" authorityYear="1926" box="[508,734,1807,1831]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="348" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B901EA90FFAAD671FE57F8F1C138281F" box="[508,525,1808,1831]" italics="true" pageId="4" pageNumber="348">N</emphasis>
.
<emphasis id="B901EA90FFAAD671FDB2F8EEC16D281F" box="[537,600,1807,1831]" italics="true" pageId="4" pageNumber="348">fuscus</emphasis>
<bibRefCitation id="EFE44B73FFAAD671FDF5F8EEC1EB281F" author="Didier, R." box="[606,734,1807,1831]" pageId="4" pageNumber="348" pagination="210 - 214" refId="ref7377" refString="Didier, R. (1926) Descriptions sommaires de Lucanides nouveaux, du genre Neolucanus, de la faune indo-chinoise. Bulletin de la Societe Entomologique de France, 1926, 210 - 214. https: // doi. org / 10.3406 / bsef. 1926.27736" type="journal article" year="1926">Didier, 1926</bibRefCitation>
</taxonomicName>
, 34.7mm, priodont (Wuming, Guangxi);
<emphasis id="B901EA90FFAAD671FBD1F8EEC79E281F" bold="true" box="[1146,1195,1807,1831]" pageId="4" pageNumber="348">EF,</emphasis>
<taxonomicName id="4C754D01FFAAD671FB1AF8F1C20B2873" authority="Bomans &amp; Miyashita, 1997" authorityName="Bomans &amp; Miyashita" authorityYear="1997" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="348" phylum="Arthropoda" rank="species" species="brochieri">
<emphasis id="B901EA90FFAAD671FB1AF8F1C7F7281F" box="[1201,1218,1808,1831]" italics="true" pageId="4" pageNumber="348">N</emphasis>
.
<emphasis id="B901EA90FFAAD671FB66F8EEC61F281F" box="[1229,1322,1807,1831]" italics="true" pageId="4" pageNumber="348">brochieri</emphasis>
<bibRefCitation id="EFE44B73FFAAD671FA9BF8EEC20B2873" author="Bomans, H. E. &amp; Miyashita, T." pageId="4" pageNumber="348" pagination="2 - 3" refId="ref7344" refString="Bomans, H. E. &amp; Miyashita, T. (1997) Description de trois nouvelles especes de Lucanidae du nord Birmanie. Besoiro, 4, 2 - 3." type="journal article" year="1997">Bomans &amp; Miyashita, 1997</bibRefCitation>
</taxonomicName>
, 31.4 mm, priodont (Gonshan, Yunnan).
<emphasis id="B901EA90FFAAD671FD77F8D2C0042873" bold="true" box="[732,817,1843,1867]" pageId="4" pageNumber="348">A, C, E,</emphasis>
dorsal view;
<emphasis id="B901EA90FFAAD671FC10F8D2C73E2873" bold="true" box="[955,1035,1843,1867]" pageId="4" pageNumber="348">B, D, F,</emphasis>
ventral view. Scale bar = 10.0 mm.
</paragraph>
</caption>
<paragraph id="8BCA3682FFABD670FF3CF895C74D28C1" blockId="5.[151,1437,1908,2041]" pageId="5" pageNumber="349">
<emphasis id="B901EA90FFABD670FF3CF895C22628B4" bold="true" box="[151,275,1908,1932]" pageId="5" pageNumber="349">FIGURE 3.</emphasis>
Male fore-head
<emphasis id="B901EA90FFABD670FE10F895C19228B4" bold="true" box="[443,679,1908,1933]" pageId="5" pageNumber="349">(AB, EF, IJ, MN)</emphasis>
, pronotum
<emphasis id="B901EA90FFABD670FCB6F895C09728B4" bold="true" box="[797,930,1908,1933]" pageId="5" pageNumber="349">(C, G, K, O)</emphasis>
, protibia and apical part of profemur
<emphasis id="B901EA90FFABD670FAB4F895C6A928B4" bold="true" box="[1311,1436,1908,1933]" pageId="5" pageNumber="349">(D, H, L, P)</emphasis>
of
<taxonomicName id="4C754D01FFABD670FF18F878C2AF2888" box="[179,410,1944,1969]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="349" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="B901EA90FFABD670FF18F878C21E2889" box="[179,299,1945,1969]" italics="true" pageId="5" pageNumber="349">Neolucanus</emphasis>
species.
<emphasis id="B901EA90FFABD670FE23F879C2AF2888" bold="true" box="[392,410,1944,1968]" pageId="5" pageNumber="349">A</emphasis>
</taxonomicName>
<emphasis id="B901EA90FFABD670FE31F879C28C2888" bold="true" box="[410,441,1944,1968]" pageId="5" pageNumber="349">D</emphasis>
,
<taxonomicName id="4C754D01FFABD670FE6DF878C18B2888" authority="Baba, 1995" authorityName="Baba" authorityYear="1995" box="[454,702,1944,1969]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="349" phylum="Arthropoda" rank="species" species="lehmanni">
<emphasis id="B901EA90FFABD670FE6DF878C2E22888" box="[454,471,1945,1968]" italics="true" pageId="5" pageNumber="349">N</emphasis>
.
<emphasis id="B901EA90FFABD670FE4EF878C1762889" box="[485,579,1945,1969]" italics="true" pageId="5" pageNumber="349">lehmanni</emphasis>
<bibRefCitation id="EFE44B73FFABD670FDE1F878C18B2888" author="Baba, M." box="[586,702,1944,1969]" pageId="5" pageNumber="349" pagination="8 - 11" refId="ref7312" refString="Baba, M. (1995) A new species of the genus Neolucanus Tomson, 1862 from south Burma (Myanmar). Gekkan-Mushi, 292, 8 - 11." type="journal article" year="1995">Baba, 1995</bibRefCitation>
</taxonomicName>
(Imitation of
<bibRefCitation id="EFE44B73FFABD670FCE6F878C0F12888" author="Baba, M." box="[845,964,1944,1969]" pageId="5" pageNumber="349" pagination="8 - 11" refId="ref7312" refString="Baba, M. (1995) A new species of the genus Neolucanus Tomson, 1862 from south Burma (Myanmar). Gekkan-Mushi, 292, 8 - 11." type="journal article" year="1995">
<collectingRegion id="49B1F860FFABD670FCE6F878C0B12889" box="[845,900,1945,1969]" country="Myanmar" name="Chin" pageId="5" pageNumber="349">Baba</collectingRegion>
, 1995
</bibRefCitation>
);
<emphasis id="B901EA90FFABD670FC7CF878C73C2889" bold="true" box="[983,1033,1945,1969]" pageId="5" pageNumber="349">EH</emphasis>
,
<taxonomicName id="4C754D01FFABD670FBBEF878C6532889" authority="Xin, Zhong &amp; Qi" authorityName="Xin, Zhong &amp; Qi" authorityYear="2025" box="[1045,1382,1945,1969]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="349" phylum="Arthropoda" rank="species" species="horizontalis" status="new species">
<emphasis id="B901EA90FFABD670FBBEF878C7132888" box="[1045,1062,1945,1968]" italics="true" pageId="5" pageNumber="349">N</emphasis>
.
<emphasis id="B901EA90FFABD670FB98F878C7992889" box="[1075,1196,1945,1969]" italics="true" pageId="5" pageNumber="349">horizontalis</emphasis>
Xin, Zhong &amp; Qi
</taxonomicName>
,
<emphasis id="B901EA90FFABD670FADAF878C3D728EC" bold="true" pageId="5" pageNumber="349">
<taxonomicNameLabel id="A23257EBFFABD670FADAF878C3D728EC" pageId="5" pageNumber="349" rank="species">new species</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="54CE8820FFABD670FF44F85CC27D28ED" box="[239,328,1981,2005]" pageId="5" pageNumber="349" type="holotype">holotype</typeStatus>
(Lvchun,
<collectingRegion id="49B1F860FFABD670FE04F85CC13428ED" box="[431,513,1981,2005]" country="China" name="Yunnan" pageId="5" pageNumber="349">Yunnan</collectingRegion>
);
<emphasis id="B901EA90FFABD670FDB8F85CC10E28ED" bold="true" box="[531,571,1981,2005]" pageId="5" pageNumber="349">IL</emphasis>
,
<taxonomicName id="4C754D01FFABD670FDECF85CC01F28EC" authority="Didier, 1926" authorityName="Didier" authorityYear="1926" box="[583,810,1980,2005]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="349" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B901EA90FFABD670FDECF85CC16D28EC" box="[583,600,1981,2004]" italics="true" pageId="5" pageNumber="349">N</emphasis>
.
<emphasis id="B901EA90FFABD670FDCEF85CC19128ED" box="[613,676,1981,2005]" italics="true" pageId="5" pageNumber="349">fuscus</emphasis>
<bibRefCitation id="EFE44B73FFABD670FD00F85CC01F28EC" author="Didier, R." box="[683,810,1980,2005]" pageId="5" pageNumber="349" pagination="210 - 214" refId="ref7377" refString="Didier, R. (1926) Descriptions sommaires de Lucanides nouveaux, du genre Neolucanus, de la faune indo-chinoise. Bulletin de la Societe Entomologique de France, 1926, 210 - 214. https: // doi. org / 10.3406 / bsef. 1926.27736" type="journal article" year="1926">Didier, 1926</bibRefCitation>
</taxonomicName>
(Ziyuan,
<collectingRegion id="49B1F860FFABD670FC26F85CC0DC28ED" box="[909,1001,1981,2005]" country="China" name="Guangxi" pageId="5" pageNumber="349">Guangxi</collectingRegion>
);
<emphasis id="B901EA90FFABD670FC51F85CC71B28ED" bold="true" box="[1018,1070,1981,2005]" pageId="5" pageNumber="349">MP</emphasis>
,
<taxonomicName id="4C754D01FFABD670FB90F85CC3FF28C1" authority="Bomans &amp; Miyashita, 1997" authorityName="Bomans &amp; Miyashita" authorityYear="1997" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="349" phylum="Arthropoda" rank="species" species="brochieri">
<emphasis id="B901EA90FFABD670FB90F85CC77928EC" box="[1083,1100,1981,2004]" italics="true" pageId="5" pageNumber="349">N</emphasis>
.
<emphasis id="B901EA90FFABD670FBF2F85CC78328ED" box="[1113,1206,1981,2005]" italics="true" pageId="5" pageNumber="349">brochieri</emphasis>
<bibRefCitation id="EFE44B73FFABD670FB16F85CC3FF28C1" author="Bomans, H. E. &amp; Miyashita, T." pageId="5" pageNumber="349" pagination="2 - 3" refId="ref7344" refString="Bomans, H. E. &amp; Miyashita, T. (1997) Description de trois nouvelles especes de Lucanidae du nord Birmanie. Besoiro, 4, 2 - 3." type="journal article" year="1997">Bomans &amp; Miyashita, 1997</bibRefCitation>
</taxonomicName>
(Gonshan,
<collectingRegion id="49B1F860FFABD670FE95F800C2A528C1" box="[318,400,2017,2041]" country="China" name="Yunnan" pageId="5" pageNumber="349">Yunnan</collectingRegion>
).
<emphasis id="B901EA90FFABD670FE0AF801C03C28C0" bold="true" box="[417,777,2016,2041]" pageId="5" pageNumber="349">A, CD, E, GH, I, KL, M, OP,</emphasis>
dorsal view;
<emphasis id="B901EA90FFABD670FC39F800C0C028C1" bold="true" box="[914,1013,2017,2041]" pageId="5" pageNumber="349">B, F, J, N</emphasis>
lateral view.
</paragraph>
</subSubSection>
<caption id="DF0A660AFFA8D673FF3CF9CFC221298A" ID-DOI="http://doi.org/10.5281/zenodo.14735141" ID-Zenodo-Dep="14735141" httpUri="https://zenodo.org/record/14735141/files/figure.png" pageId="6" pageNumber="350" startId="6.[151,250,1582,1606]" targetBox="[158,1429,188,1551]" targetPageId="6" targetType="figure">
<paragraph id="8BCA3682FFA8D673FF3CF9CFC221298A" blockId="6.[151,1437,1582,1714]" pageId="6" pageNumber="350">
<emphasis id="B901EA90FFA8D673FF3CF9CFC227297E" bold="true" box="[151,274,1582,1606]" pageId="6" pageNumber="350">FIGURE 4.</emphasis>
Male fore-head
<emphasis id="B901EA90FFA8D673FE1CF9CFC169297E" bold="true" box="[439,604,1582,1607]" pageId="6" pageNumber="350">(AB, EF, IJ)</emphasis>
, pronotum
<emphasis id="B901EA90FFA8D673FD64F9CFC004297E" bold="true" box="[719,817,1582,1606]" pageId="6" pageNumber="350">(C, G, K)</emphasis>
, protibia and apical part of profemur
<emphasis id="B901EA90FFA8D673FB0DF9CFC633297E" bold="true" box="[1190,1286,1582,1607]" pageId="6" pageNumber="350">(D, H, L)</emphasis>
of
<taxonomicName id="4C754D01FFA8D673FA8EF9CFC3C82952" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="B901EA90FFA8D673FA8EF9CFC6A8297E" box="[1317,1437,1582,1606]" italics="true" pageId="6" pageNumber="350">Neolucanus</emphasis>
species.
<emphasis id="B901EA90FFA8D673FF40F9B3C3C82952" bold="true" box="[235,253,1618,1642]" pageId="6" pageNumber="350">A</emphasis>
</taxonomicName>
<emphasis id="B901EA90FFA8D673FF56F9B3C2282952" bold="true" box="[253,285,1618,1642]" pageId="6" pageNumber="350">D</emphasis>
,
<taxonomicName id="4C754D01FFA8D673FE83F9B2C15B2952" authority="Xin, Zhong &amp; Qi" authorityName="Xin, Zhong &amp; Qi" authorityYear="2025" box="[296,622,1618,1643]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="horizontalis" status="new species">
<emphasis id="B901EA90FFA8D673FE83F9B2C20C2952" box="[296,313,1619,1642]" italics="true" pageId="6" pageNumber="350">N</emphasis>
.
<emphasis id="B901EA90FFA8D673FEEFF9B3C2882952" box="[324,445,1618,1642]" italics="true" pageId="6" pageNumber="350">horizontalis</emphasis>
Xin, Zhong &amp; Qi
</taxonomicName>
,
<emphasis id="B901EA90FFA8D673FDDDF9B2C1C72952" bold="true" box="[630,754,1618,1643]" pageId="6" pageNumber="350">
<taxonomicNameLabel id="A23257EBFFA8D673FDDDF9B2C1C72952" box="[630,754,1618,1643]" pageId="6" pageNumber="350" rank="species">new species</taxonomicNameLabel>
</emphasis>
, paratype (Jinping, Yunnan);
<emphasis id="B901EA90FFA8D673FBBCF9B2C77C2953" bold="true" box="[1047,1097,1619,1643]" pageId="6" pageNumber="350">EH</emphasis>
,
<taxonomicName id="4C754D01FFA8D673FBFFF9B2C6042952" authority="Didier, 1926" authorityName="Didier" authorityYear="1926" box="[1108,1329,1618,1643]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B901EA90FFA8D673FBFFF9B2C7502952" box="[1108,1125,1619,1642]" italics="true" pageId="6" pageNumber="350">N</emphasis>
.
<emphasis id="B901EA90FFA8D673FBDBF9B2C79A2953" box="[1136,1199,1619,1643]" italics="true" pageId="6" pageNumber="350">fuscus</emphasis>
<bibRefCitation id="EFE44B73FFA8D673FB1FF9B3C6042952" author="Didier, R." box="[1204,1329,1618,1642]" pageId="6" pageNumber="350" pagination="210 - 214" refId="ref7377" refString="Didier, R. (1926) Descriptions sommaires de Lucanides nouveaux, du genre Neolucanus, de la faune indo-chinoise. Bulletin de la Societe Entomologique de France, 1926, 210 - 214. https: // doi. org / 10.3406 / bsef. 1926.27736" type="journal article" year="1926">Didier, 1926</bibRefCitation>
</taxonomicName>
(Wuming, Guangxi);
<emphasis id="B901EA90FFA8D673FEAEF996C21829B7" bold="true" box="[261,301,1655,1679]" pageId="6" pageNumber="350">IL</emphasis>
,
<taxonomicName id="4C754D01FFA8D673FE90F996C1EF29B7" authority="Bomans &amp; Miyashita, 1997" authorityName="Bomans &amp; Miyashita" authorityYear="1997" box="[315,730,1654,1679]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="brochieri">
<emphasis id="B901EA90FFA8D673FE90F996C27929B6" box="[315,332,1655,1678]" italics="true" pageId="6" pageNumber="350">N</emphasis>
.
<emphasis id="B901EA90FFA8D673FEF2F997C28329B6" box="[345,438,1654,1678]" italics="true" pageId="6" pageNumber="350">brochieri</emphasis>
<bibRefCitation id="EFE44B73FFA8D673FE15F996C1EF29B7" author="Bomans, H. E. &amp; Miyashita, T." box="[446,730,1654,1679]" pageId="6" pageNumber="350" pagination="2 - 3" refId="ref7344" refString="Bomans, H. E. &amp; Miyashita, T. (1997) Description de trois nouvelles especes de Lucanidae du nord Birmanie. Besoiro, 4, 2 - 3." type="journal article" year="1997">Bomans &amp; Miyashita, 1997</bibRefCitation>
</taxonomicName>
(Gonshan, Yunnan).
<emphasis id="B901EA90FFA8D673FC1FF997C7F629B7" bold="true" box="[948,1219,1654,1679]" pageId="6" pageNumber="350">A, CD, E, GH, I, KL,</emphasis>
dorsal view;
<emphasis id="B901EA90FFA8D673FAFBF996C6A929B7" bold="true" box="[1360,1436,1655,1679]" pageId="6" pageNumber="350">B, F, J,</emphasis>
lateral view.
</paragraph>
</caption>
<subSubSection id="C36F6509FFA8D673FF6CF8E5C0072827" box="[199,818,1796,1823]" pageId="6" pageNumber="350" type="biology_ecology">
<paragraph id="8BCA3682FFA8D673FF6CF8E5C0072827" blockId="6.[151,1436,1796,1895]" box="[199,818,1796,1823]" pageId="6" pageNumber="350">
<emphasis id="B901EA90FFA8D673FF6CF8E5C2122826" bold="true" box="[199,295,1796,1822]" pageId="6" pageNumber="350">Biology.</emphasis>
This species can be attracted to artificial light.
</paragraph>
</subSubSection>
<subSubSection id="C36F6509FFA8D672FF6CF8C8C62A28B3" lastPageId="7" lastPageNumber="351" pageId="6" pageNumber="350" type="discussion">
<paragraph id="8BCA3682FFA8D673FF6CF8C8C6B0285F" blockId="6.[151,1436,1796,1895]" pageId="6" pageNumber="350">
<emphasis id="B901EA90FFA8D673FF6CF8C8C20E287B" bold="true" box="[199,315,1833,1859]" pageId="6" pageNumber="350">Remarks.</emphasis>
Many species within the
<taxonomicName id="4C754D01FFA8D673FDE4F8C8C181287B" authorityName="Didier" authorityYear="1926" box="[591,692,1833,1859]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="350" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B901EA90FFA8D673FDE4F8C8C157287A" box="[591,610,1833,1858]" italics="true" pageId="6" pageNumber="350">N</emphasis>
.
<emphasis id="B901EA90FFA8D673FDC5F8C8C181287B" box="[622,692,1833,1859]" italics="true" pageId="6" pageNumber="350">fuscus</emphasis>
</taxonomicName>
group exhibit individuals with elytra featuring prominent yellowish areas. Therefore, we hypothesize that similar individuals will also be found within the newly discovered species.
</paragraph>
<paragraph id="8BCA3682FFA9D672FF3CF8E6C62A28B3" blockId="7.[151,1437,1799,1931]" pageId="7" pageNumber="351">
<emphasis id="B901EA90FFA9D672FF3CF8E6C2272827" bold="true" box="[151,274,1799,1823]" pageId="7" pageNumber="351">FIGURE 5.</emphasis>
Male abdominal tergite VIII (
<emphasis id="B901EA90FFA9D672FD94F8E6C1932827" bold="true" box="[575,678,1799,1823]" pageId="7" pageNumber="351">A, E, I, M</emphasis>
), abdominal ventrite VIII (
<emphasis id="B901EA90FFA9D672FC1DF8E6C7202827" bold="true" box="[950,1045,1799,1823]" pageId="7" pageNumber="351">B, F, J, N</emphasis>
) and abdominal segment IX (
<emphasis id="B901EA90FFA9D672FA94F8E6C213287B" bold="true" pageId="7" pageNumber="351">CD, G H, KL, OP</emphasis>
) of
<taxonomicName id="4C754D01FFA9D672FEFBF8CAC103287B" box="[336,566,1835,1859]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="351" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="B901EA90FFA9D672FEFBF8CAC2FD287B" box="[336,456,1835,1859]" italics="true" pageId="7" pageNumber="351">Neolucanus</emphasis>
species.
<emphasis id="B901EA90FFA9D672FD8FF8CAC103287B" bold="true" box="[548,566,1835,1859]" pageId="7" pageNumber="351">A</emphasis>
</taxonomicName>
<emphasis id="B901EA90FFA9D672FD9CF8CAC168287B" bold="true" box="[567,605,1835,1859]" pageId="7" pageNumber="351">H,</emphasis>
<taxonomicName id="4C754D01FFA9D672FDCFF8CDC086287B" authority="Xin, Zhong &amp; Qi" authorityName="Xin, Zhong &amp; Qi" authorityYear="2025" box="[612,947,1835,1859]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="351" phylum="Arthropoda" rank="species" species="horizontalis" status="new species">
<emphasis id="B901EA90FFA9D672FDCFF8CDC140287B" box="[612,629,1836,1859]" italics="true" pageId="7" pageNumber="351">N</emphasis>
.
<emphasis id="B901EA90FFA9D672FD29F8CAC1CE287B" box="[642,763,1835,1859]" italics="true" pageId="7" pageNumber="351">horizontalis</emphasis>
Xin, Zhong &amp; Qi
</taxonomicName>
,
<emphasis id="B901EA90FFA9D672FC16F8CAC74D287B" bold="true" box="[957,1144,1835,1859]" pageId="7" pageNumber="351">
<taxonomicNameLabel id="A23257EBFFA9D672FC16F8CAC70F287B" box="[957,1082,1835,1859]" pageId="7" pageNumber="351" rank="species">new species</taxonomicNameLabel>
AD,
</emphasis>
<typeStatus id="54CE8820FFA9D672FBD5F8CAC7E2287B" box="[1150,1239,1835,1859]" pageId="7" pageNumber="351" type="holotype">holotype</typeStatus>
(Lvchun,
<collectingRegion id="49B1F860FFA9D672FA94F8CAC6A4287B" box="[1343,1425,1835,1859]" country="China" name="Yunnan" pageId="7" pageNumber="351">Yunnan</collectingRegion>
);
<emphasis id="B901EA90FFA9D672FF3CF8AEC3FA285F" bold="true" box="[151,207,1871,1895]" pageId="7" pageNumber="351">EH,</emphasis>
<typeStatus id="54CE8820FFA9D672FF7FF8B1C21E2850" box="[212,299,1872,1896]" pageId="7" pageNumber="351" type="paratype">paratype</typeStatus>
(Jinping,
<collectingRegion id="49B1F860FFA9D672FE25F8AEC2D5285F" box="[398,480,1871,1895]" country="China" name="Yunnan" pageId="7" pageNumber="351">Yunnan</collectingRegion>
).
<emphasis id="B901EA90FFA9D672FE44F8AEC128285F" bold="true" box="[495,541,1871,1895]" pageId="7" pageNumber="351">IL,</emphasis>
<taxonomicName id="4C754D01FFA9D672FD89F8B1C034285F" authority="Didier, 1926" authorityName="Didier" authorityYear="1926" box="[546,769,1871,1895]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="351" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B901EA90FFA9D672FD89F8B1C106285F" box="[546,563,1872,1895]" italics="true" pageId="7" pageNumber="351">N</emphasis>
.
<emphasis id="B901EA90FFA9D672FD95F8AEC148285F" box="[574,637,1871,1895]" italics="true" pageId="7" pageNumber="351">fuscus</emphasis>
<bibRefCitation id="EFE44B73FFA9D672FD29F8AEC034285F" author="Didier, R." box="[642,769,1871,1895]" pageId="7" pageNumber="351" pagination="210 - 214" refId="ref7377" refString="Didier, R. (1926) Descriptions sommaires de Lucanides nouveaux, du genre Neolucanus, de la faune indo-chinoise. Bulletin de la Societe Entomologique de France, 1926, 210 - 214. https: // doi. org / 10.3406 / bsef. 1926.27736" type="journal article" year="1926">Didier, 1926</bibRefCitation>
</taxonomicName>
(Ziyuan,
<collectingRegion id="49B1F860FFA9D672FCCAF8AEC088285F" box="[865,957,1871,1895]" country="China" name="Guangxi" pageId="7" pageNumber="351">Guangxi</collectingRegion>
).
<emphasis id="B901EA90FFA9D672FC67F8AEC731285F" bold="true" box="[972,1028,1871,1895]" pageId="7" pageNumber="351">MP,</emphasis>
<taxonomicName id="4C754D01FFA9D672FBA2F8B1C6A8285F" authority="Bomans &amp; Miyashita, 1997" authorityName="Bomans &amp; Miyashita" authorityYear="1997" box="[1033,1437,1871,1895]" class="Insecta" family="Lucanidae" genus="Neolucanus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="351" phylum="Arthropoda" rank="species" species="brochieri">
<emphasis id="B901EA90FFA9D672FBA2F8B1C72F285F" box="[1033,1050,1872,1895]" italics="true" pageId="7" pageNumber="351">N</emphasis>
.
<emphasis id="B901EA90FFA9D672FB8EF8AEC7B7285F" box="[1061,1154,1871,1895]" italics="true" pageId="7" pageNumber="351">brochieri</emphasis>
<bibRefCitation id="EFE44B73FFA9D672FB23F8B1C6A8285F" author="Bomans, H. E. &amp; Miyashita, T." box="[1160,1437,1871,1895]" pageId="7" pageNumber="351" pagination="2 - 3" refId="ref7344" refString="Bomans, H. E. &amp; Miyashita, T. (1997) Description de trois nouvelles especes de Lucanidae du nord Birmanie. Besoiro, 4, 2 - 3." type="journal article" year="1997">Bomans &amp; Miyashita, 1997</bibRefCitation>
</taxonomicName>
(Gonshan,
<collectingRegion id="49B1F860FFA9D672FEAEF892C26228B3" box="[261,343,1907,1931]" country="China" name="Yunnan" pageId="7" pageNumber="351">Yunnan</collectingRegion>
).
<emphasis id="B901EA90FFA9D672FEC3F892C16C28B3" bold="true" box="[360,601,1907,1931]" pageId="7" pageNumber="351">A, C, E, G, I, K, M, O,</emphasis>
dorsal view;
<emphasis id="B901EA90FFA9D672FD49F892C0F628B3" bold="true" box="[738,963,1907,1931]" pageId="7" pageNumber="351">B, D, F, H, J, L, N, P,</emphasis>
ventral view. Scale bar = 1.0 mm.
</paragraph>
</subSubSection>
</treatment>
</document>