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<document id="53C9611B9A114A1BB6CDE4E07D23BD77" ID-DOI="10.11646/phytotaxa.510.3.4" ID-ISSN="1179-3163" ID-Zenodo-Dep="5426421" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="operationResults" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1630397611626" checkinUser="marcus" docAuthor="Bunchalee, Pasakorn, Leeratiwong, Charan &amp; Johnson, David M." docDate="2021" docId="1E1287B5FF91FFC2FF37FC4FFDA1F8AF" docLanguage="en" docName="phytotaxa.510.3.4.pdf" docOrigin="Phytotaxa 510 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.510.3.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Polyalthia pumila Ridl. J. Straits Branch Roy. Asiat. Soc." docType="treatment" docVersion="1" lastPageNumber="248" masterDocId="E22BFFCDFF96FFCBFFBFFFDDFFAFFFD6" masterDocTitle="Two new species and a new record of the genus Polyalthia (Annonaceae) from Peninsular Thailand" masterLastPageNumber="250" masterPageNumber="239" pageNumber="246" updateTime="1732197805484" updateUser="guilherme" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="2D65D2C7029989D728B51839CF3CA7F7">
<mods:title id="13679E200F7CDAB05BC5434B0CE79332">Two new species and a new record of the genus Polyalthia (Annonaceae) from Peninsular Thailand</mods:title>
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<mods:namePart id="92D6CAA371DE1DD2B79BAA0CD35E03A8">Bunchalee, Pasakorn</mods:namePart>
<mods:nameIdentifier id="E5E3981F87495B6549FFB0A5FEFCDE4C" type="ORCID">0000-0001-8644-3360</mods:nameIdentifier>
<mods:affiliation id="4814549C80473E9EC64C3E53D790B6EC">Department of Biology, Faculty of Science, Mahasarakham University, Kantarawichai, Maha Sarakham, 44150, Thailand. Palaeontological Research and Education Center, Mahasarakham University, Kantarawichai, Maha Sarakham, 44150, Thailand. Division of Biological Science, Faculty of Science, Prince of Songkla University, Hat Yai, Songkhla 90112, Thailand. &amp; pasakorn. b @ msu. ac. th; https: // orcid. org / 0000 - 0001 - 8644 - 3360</mods:affiliation>
<mods:nameIdentifier id="430119F950A6F45E0B434FA67EB1892F" type="email">pasakorn.b@msu.ac.th</mods:nameIdentifier>
</mods:name>
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<mods:role id="41703E256FF74160B03ABDFB99017620">
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<mods:namePart id="6B269CE8A434EDA73ED59A5B162121BB">Leeratiwong, Charan</mods:namePart>
<mods:nameIdentifier id="6C7F53E3B66ED42D34DCC5A3F96AC6A5" type="ORCID">0000-0002-0752-4275</mods:nameIdentifier>
<mods:affiliation id="118D35B18936651F40EAED2BCA7351E6">charan. leeratiwong @ gmail. com; https: // orcid. org / 0000 - 0002 - 0752 - 4275</mods:affiliation>
<mods:nameIdentifier id="7432FC1B4747FBFEAD3D3854773A850D" type="email">charan.leeratiwong@gmail.com</mods:nameIdentifier>
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<mods:namePart id="25FD4305CE9ECED7E5A3ED221519DE54">Johnson, David M.</mods:namePart>
<mods:nameIdentifier id="AF57DB655FE1E32DA36C26DD188751C6" type="ORCID">0000-0003-2896-7419</mods:nameIdentifier>
<mods:affiliation id="8369087FEA8EA0C88A9D82BABBB439CB">Department of Botany and Microbiology, Ohio Wesleyan University, Delaware, Ohio, 43015, USA &amp; dmjohnso @ owu. edu; https: // orcid. org / 0000 - 0003 - 2896 - 7419</mods:affiliation>
<mods:nameIdentifier id="84C9981DF52A13A56A9783187B2F8858" type="email">dmjohnso@owu.edu</mods:nameIdentifier>
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<mods:title id="7DABB31286EBDE76C9A911BDE4136C05">Phytotaxa</mods:title>
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<mods:date id="E37DD39F89A74F75861BE309785FCEE5">2021</mods:date>
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<mods:number id="45089848365E43D890009FE092DE1995">2021-07-09</mods:number>
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<mods:number id="08424FF170A9D06493E3412D3F3007D7">510</mods:number>
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<treatment id="1E1287B5FF91FFC2FF37FC4FFDA1F8AF" LSID="urn:lsid:plazi:treatment:1E1287B5FF91FFC2FF37FC4FFDA1F8AF" httpUri="http://treatment.plazi.org/id/1E1287B5FF91FFC2FF37FC4FFDA1F8AF" lastPageId="9" lastPageNumber="248" pageId="7" pageNumber="246">
<subSubSection id="DEA16528FF91FFCCFF37FC4FFC02FC7A" box="[136,941,914,941]" pageId="7" pageNumber="246" type="nomenclature">
<paragraph id="960436A3FF91FFCCFF37FC4FFC02FC7A" blockId="7.[136,1452,914,1013]" box="[136,941,914,941]" pageId="7" pageNumber="246">
<taxonomicName id="51BB4D20FF91FFCCFF37FC4FFCAAFC7A" ID-CoL="4KZQR" authority="Ridl. J. Straits Branch Roy. Asiat. Soc." box="[136,773,914,941]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="7" pageNumber="246" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="A4CFEAB1FF91FFCCFF37FC4FFEF5FC7A" bold="true" box="[136,346,914,940]" pageId="7" pageNumber="246">Polyalthia pumila</emphasis>
Ridl. J. Straits Branch Roy.Asiat. Soc.
</taxonomicName>
54: 12. (1910).
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF91FFCCFC0EFC4EFC9DFC23" pageId="7" pageNumber="246" type="materials_examined">
<paragraph id="960436A3FF91FFCCFC0EFC4EFC9DFC23" blockId="7.[136,1452,914,1013]" pageId="7" pageNumber="246">
TYPE:
<materialsCitation id="26D33CFEFF91FFCCFBBBFC4FFE88FC22" collectingDate="1896-03" collectionCode="SING" collectorName="Ridley" country="Malaysia" location="Sera Woods" municipality="Dindings" pageId="7" pageNumber="246" specimenCode="SING0059298" specimenCount="1" stateProvince="Perak" typeStatus="lectotype">
Peninsular
<collectingCountry id="EEAC7633FF91FFCCFBC0FC4FFB47FC7A" box="[1151,1256,914,940]" name="Malaysia" pageId="7" pageNumber="246">Malaysia</collectingCountry>
,
<collectingRegion id="547FF841FF91FFCCFB4DFC4FFA9AFC7A" box="[1266,1333,914,940]" country="Malaysia" name="Perak" pageId="7" pageNumber="246">Perak</collectingRegion>
,
<collectingMunicipality id="7660ACD9FF91FFCCFA81FC4FFA07FC7A" box="[1342,1448,914,940]" pageId="7" pageNumber="246">Dindings</collectingMunicipality>
,
<location id="93646078FF91FFCCFF37FC6BFEBFFC06" LSID="urn:lsid:plazi:treatment:1E1287B5FF91FFC2FF37FC4FFDA1F8AF:93646078FF91FFCCFF37FC6BFEBFFC06" box="[136,272,950,976]" country="Malaysia" county="Peninsular" municipality="Dindings" name="Sera Woods" pageId="7" pageNumber="246" stateProvince="Perak">Sera Woods</location>
,
<date id="E2051063FF91FFCCFEA4FC6BFE0AFC06" box="[283,421,950,976]" pageId="7" pageNumber="246" value="1896-03">
<collectingDate id="F241E98BFF91FFCCFEA4FC6BFE0AFC06" box="[283,421,950,976]" pageId="7" pageNumber="246" value="1896-03">March 1896</collectingDate>
</date>
,
<emphasis id="A4CFEAB1FF91FFCCFE0FFC6BFD99FC06" box="[432,566,950,976]" italics="true" pageId="7" pageNumber="246">
<collectorName id="3B4E5375FF91FFCCFE0FFC6BFE57FC06" box="[432,504,950,976]" pageId="7" pageNumber="246">Ridley</collectorName>
7996
</emphasis>
(
<typeStatus id="49008801FF91FFCCFDFBFC6BFD02FC06" box="[580,685,950,976]" pageId="7" pageNumber="246" type="lectotype">lectotype</typeStatus>
<collectionCode id="F0AAAE66FF91FFCCFD0CFC6BFD5BFC06" box="[691,756,950,976]" collectionName="SING" pageId="7" pageNumber="246">SING</collectionCode>
[
<specimenCode id="C61D9ED8FF91FFCCFCBEFC6BFC01FC07" box="[769,942,950,977]" pageId="7" pageNumber="246">SING0059298</specimenCode>
], designated by Turner, Gard. Bull. Singapore 64: 241. 2012
</materialsCitation>
;
<materialsCitation id="26D33CFEFF91FFCCFE8DFC07FD21FC23" box="[306,654,986,1013]" collectingDate="1896-03" collectionCode="K" collectorName="Ridley" country="Malaysia" location="Sera Woods" municipality="Dindings" pageId="7" pageNumber="246" specimenCode="K000691442" specimenCount="1" stateProvince="Perak" typeStatus="isolectotype">
<typeStatus id="49008801FF91FFCCFE8DFC07FE16FC22" box="[306,441,986,1012]" pageId="7" pageNumber="246">isolectotype</typeStatus>
<collectionCode id="F0AAAE66FF91FFCCFE7FFC07FE75FC22" box="[448,474,986,1012]" pageId="7" pageNumber="246">K</collectionCode>
! [
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]
</materialsCitation>
). (
<figureCitation id="0E802A26FF91FFCCFD16FC07FD59FC23" box="[681,758,986,1013]" captionStart="FIGURE 5" captionStartId="8.[136,229,1949,1971]" captionTargetBox="[177,1387,203,1911]" captionTargetId="figure-21@8.[164,1423,187,1927]" captionTargetPageId="8" captionText="FIGURE 5. Polyalthia pumila: A. Habit; B. Inflorescence, showing supra-axillary attachment; C. Base of flower, with petals removed to show stamens and carpels; D. Sepals (outside below, inside above); E. Outside of outer petal; F. Outside of inner petal; G. Stamens (inside view on left, outside view on right); H. Carpel; I. Fruit; J. Monocarp. The illustration was drawn by Thadanai Saensuk from Leeratiwong 20-1528 (PSU)." figureDoi="http://doi.org/10.5281/zenodo.5426431" httpUri="https://zenodo.org/record/5426431/files/figure.png" pageId="7" pageNumber="246">Figs. 5</figureCitation>
&amp;
<figureCitation id="0E802A26FF91FFCCFCA5FC07FC86FC23" box="[794,809,986,1013]" captionStart="FIGURE 6" captionStartId="9.[136,229,1531,1553]" captionTargetBox="[167,1427,205,1480]" captionTargetId="figure-18@9.[151,1436,190,1507]" captionTargetPageId="9" captionText="FIGURE 6. Polyalthia pumila: A. Twig and flowers; B. Flowers and a single monocarp; C. Upper surface of leaf base; D. Lower surface of leaf; E. Fruit; F. Seed (Side view). Photographs in A &amp; B were taken Charan Leeratiwong in Narathiwat, Cha Nae District, Dusong Yo Subdistrict, Ruepoh village, Mo Tae Mountain, 4 April 2020. C, D, E &amp; F are from Leeratiwong 20-1528 (PSU); photographs by Pasakorn Bunchalee." figureDoi="http://doi.org/10.5281/zenodo.5426433" httpUri="https://zenodo.org/record/5426433/files/figure.png" pageId="7" pageNumber="246">6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF91FFCCFF37FBFFFCD4F8DA" pageId="7" pageNumber="246" type="description">
<paragraph id="960436A3FF91FFCCFF37FBFFFCD4F8DA" blockId="7.[136,1452,1058,2057]" pageId="7" pageNumber="246">
Shrubs
<quantity id="51439B46FF91FFCCFF61FBFFFEFEFBEB" box="[222,337,1058,1085]" metricMagnitude="0" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="7" pageNumber="246" unit="m" value="1.0" valueMax="1.2" valueMin="0.8">0.81.2 m</quantity>
tall, bark smooth, blackish brown, primary branching spiral.
<emphasis id="A4CFEAB1FF91FFCCFBB1FBFEFB6DFBEA" box="[1038,1218,1058,1084]" italics="true" pageId="7" pageNumber="246">Young branches</emphasis>
rusty brown to dark brown tomentose, the hairs erect and ca. 1.0 mm long, with moderately prominent lenticels.
<emphasis id="A4CFEAB1FF91FFCCFB3AFB9AFB7CFBB6" box="[1157,1235,1095,1120]" italics="true" pageId="7" pageNumber="246">Leaves</emphasis>
with petioles stout, 3.06.0 mm long, flat above, rustybrown tomentose; lamina coriaceous, smooth, symmetrically or asymmetrically elliptic, oblanceolate or obovate, (12.0)15.030.0 × (4.5)6.011.0 cm, base asymmetrically cordate with auricles 2.0 3.0 mm long, apex acute to acuminate with acumen 7.015.0 mm long, upper side glabrous except on base of midrib, which is sparsely puberulous, lower side glabrescent, puberulous along midrib; midrib grooved above; lateral veins (10)1216 per side, fairly straight, diverging at 5565° from the midrib, simple loop-forming 3.04.0 mm from the margins; intersecondary veins present; tertiary veins reticulate.
<emphasis id="A4CFEAB1FF91FFCCFCF5FAC3FC44FAEE" box="[842,1003,1310,1336]" italics="true" pageId="7" pageNumber="246">Inflorescences</emphasis>
supra-axillary on leafy branches, 1(3)- flowered; pedicels
<quantity id="51439B46FF91FFCCFEE5FA9FFE4FFA8B" box="[346,480,1346,1373]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.5" metricValueMin="1.5" pageId="7" pageNumber="246" unit="mm" value="2.5" valueMax="3.5" valueMin="1.5">1.53.5 mm</quantity>
long,
<quantity id="51439B46FF91FFCCFD9BFA9FFD06FA8B" box="[548,681,1346,1373]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="7" pageNumber="246" unit="mm" value="1.65" valueMax="1.8" valueMin="1.5">1.51.8 mm</quantity>
thick, pubescent; bract solitary, linear, 1.52.5 × ca.
<quantity id="51439B46FF91FFCCFB52FA9FFAEBFA8B" box="[1261,1348,1346,1373]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="246" unit="mm" value="0.8">0.8 mm</quantity>
, attached at pedicel base, sparsely pubescent outside and glabrous inside; bud lanceolate, petals separate at base, valvate at apex.
<emphasis id="A4CFEAB1FF91FFCCFF37FA57FF7EFA72" box="[136,209,1418,1444]" italics="true" pageId="7" pageNumber="246">Sepals</emphasis>
imbricate, brownish-green
<emphasis id="A4CFEAB1FF91FFCCFDB3FA56FDF5FA72" box="[524,602,1419,1444]" italics="true" pageId="7" pageNumber="246">in vivo</emphasis>
, ovate, 5.06.0 × 4.05.0 mm, apex acute, sparsely pubescent outside and glabrous inside.
<emphasis id="A4CFEAB1FF91FFCCFEFDFA73FE27FA1E" box="[322,392,1454,1480]" italics="true" pageId="7" pageNumber="246">Petals</emphasis>
orangish red to red, coriaceous, separate, erect, apex acute; outer petals lanceolate, 23.030.0 × 3.05.0 mm; inner petals linear, 25.033.0 × 2.03.0 mm.
<emphasis id="A4CFEAB1FF91FFCCFC89FA0EFC3CFA3B" box="[822,915,1491,1517]" italics="true" pageId="7" pageNumber="246">Stamens</emphasis>
orangish red to red, slightly cuneate, 1.31.5 ×
<quantity id="51439B46FF91FFCCFF37FA2BFEBEF9C7" box="[136,273,1526,1553]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="7" pageNumber="246" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.60.7 mm</quantity>
, filament ca.
<quantity id="51439B46FF91FFCCFE19FA2BFE53F9C7" box="[422,508,1526,1553]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="246" unit="mm" value="0.2">0.2 mm</quantity>
long, anthers ca. 1.0 mm long; anther connective apex truncate; androecium 5.06.0 mm in diameter.
<emphasis id="A4CFEAB1FF91FFCCFEF0F9C7FE07F9E2" box="[335,424,1562,1588]" italics="true" pageId="7" pageNumber="246">Carpels</emphasis>
1015 per flower, 2.22.5 ×
<quantity id="51439B46FF91FFCCFCBEF9C7FC20F9E3" box="[769,911,1562,1589]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="7" pageNumber="246" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0.30.4 mm</quantity>
, pubescent, style rudimentary, stigmas clavate, 1.0
<quantity id="51439B46FF91FFCCFF05F9E3FEBFF98F" box="[186,272,1598,1625]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="246" unit="mm" value="1.2">1.2 mm</quantity>
, pubescent, exceeding the height of the anther connective apices; ovules ca. 2 per carpel, attached laterally.
<emphasis id="A4CFEAB1FF91FFCCFF37F9BEFF68F9AA" box="[136,199,1635,1660]" italics="true" pageId="7" pageNumber="246">Torus</emphasis>
cushion-shaped and slightly concave at top, 2.53.0 mm in diameter, 1.0
<quantity id="51439B46FF91FFCCFBA8F9BFFBC1F9AB" box="[1047,1134,1634,1661]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="246" unit="mm" value="1.2">1.2 mm</quantity>
thick, glabrous.
<emphasis id="A4CFEAB1FF91FFCCFA92F9BEFADEF9AA" box="[1325,1393,1635,1660]" italics="true" pageId="7" pageNumber="246">Fruits</emphasis>
with 712 monocarps on pedicels 18.025.0 mm long, 1.82.0 mm thick, pubescent.
<emphasis id="A4CFEAB1FF91FFCCFB98F95AFB08F976" box="[1063,1191,1671,1696]" italics="true" pageId="7" pageNumber="246">Monocarps</emphasis>
greenish yellow when ripe, globose to subglobose or suboblongoid, 9.012.0 × 8.09.0 mm, base rounded, apex mucronate, glabrous, stipes 1.02.0 mm long, 1.0
<quantity id="51439B46FF91FFCCFE2FF913FE47F93F" box="[400,488,1742,1769]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="246" unit="mm" value="1.2">1.2 mm</quantity>
thick.
<emphasis id="A4CFEAB1FF91FFCCFD85F913FDD5F93E" box="[570,634,1742,1768]" italics="true" pageId="7" pageNumber="246">Seeds</emphasis>
plano-convex, 12 per monocarp, ca. 8.0 × 7.0 × 4.0 mm, grooved in horizontal view, surface rugulate and sparsely pitted, slightly shiny.
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF91FFCCFF02F8CBFB2AF882" pageId="7" pageNumber="246" type="distribution">
<paragraph id="960436A3FF91FFCCFF02F8CBFB2AF882" blockId="7.[136,1452,1058,2057]" pageId="7" pageNumber="246">
<emphasis id="A4CFEAB1FF91FFCCFF02F8CBFE5BF8E6" bold="true" box="[189,500,1814,1840]" pageId="7" pageNumber="246">Distribution and Ecology:</emphasis>
—Understory of tropical rain forest in
<collectingRegion id="547FF841FF91FFCCFC15F8CBFB3DF8E6" box="[938,1170,1814,1840]" country="Thailand" name="Narathiwat" pageId="7" pageNumber="246">Narathiwat Province</collectingRegion>
near the southern border of Peninsular
<collectingCountry id="EEAC7633FF91FFCCFE9CF8E7FE28F882" box="[291,391,1850,1876]" name="Thailand" pageId="7" pageNumber="246">Thailand</collectingCountry>
(
<figureCitation id="0E802A26FF91FFCCFE29F8E7FE75F883" box="[406,474,1850,1877]" captionStart="FIGURE 7" captionStartId="10.[136,229,1555,1577]" captionTargetBox="[178,1417,221,1505]" captionTargetId="figure-21@10.[151,1436,190,1531]" captionTargetPageId="10" captionText="FIGURE 7. A. &amp; B. Maps showing distributions of Polyalthia heliopetala (●), P. taweensis (▲) and P. pumila (■) in southern Peninsular Thailand. Map in B is an enlargement of the area shown in the rectangle in A. Maps by Pasakorn Bunchalee." figureDoi="http://doi.org/10.5281/zenodo.5426436" httpUri="https://zenodo.org/record/5426436/files/figure.png" pageId="7" pageNumber="246">Fig. 7</figureCitation>
), south to
<collectingRegion id="547FF841FF91FFCCFDEFF8E7FD3FF882" box="[592,656,1850,1876]" country="Malaysia" name="Perak" pageId="7" pageNumber="246">Perak</collectingRegion>
and
<collectingRegion id="547FF841FF91FFCCFD79F8E7FCABF882" box="[710,772,1850,1876]" country="Malaysia" name="Johor" pageId="7" pageNumber="246">Johor</collectingRegion>
States of
<collectingCountry id="EEAC7633FF91FFCCFCCBF8E7FC74F882" box="[884,987,1850,1876]" name="Malaysia" pageId="7" pageNumber="246">Malaysia</collectingCountry>
(
<bibRefCitation id="F22A4B52FF91FFCCFC56F8E7FBD7F882" author="Ridley, H. N." box="[1001,1144,1850,1876]" pageId="7" pageNumber="246" pagination="1 - 61" refId="ref5232" refString="Ridley, H. N. (1910) New or rare Malayan plants. Series V. Journal of the Straits Branch of the Royal Asiatic Society 54: 1 - 61." type="journal article" year="1910">Ridley 1910</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF91FFCCFF02F883FD6DF8AE" box="[189,706,1886,1912]" pageId="7" pageNumber="246" type="biology_ecology">
<paragraph id="960436A3FF91FFCCFF02F883FD6DF8AE" blockId="7.[136,1452,1058,2057]" box="[189,706,1886,1912]" pageId="7" pageNumber="246">
<emphasis id="A4CFEAB1FF91FFCCFF02F883FEECF8AE" bold="true" box="[189,323,1886,1912]" pageId="7" pageNumber="246">Phenology:</emphasis>
—Flowering and fruiting in April.
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF91FFCCFF02F85FFDE7F84B" box="[189,584,1922,1949]" pageId="7" pageNumber="246" type="vernacular_names">
<paragraph id="960436A3FF91FFCCFF02F85FFDE7F84B" blockId="7.[136,1452,1058,2057]" box="[189,584,1922,1949]" pageId="7" pageNumber="246">
<emphasis id="A4CFEAB1FF91FFCCFF02F85FFEFDF84A" bold="true" box="[189,338,1922,1948]" pageId="7" pageNumber="246">Local name:</emphasis>
—Thai: Saed sri nara.
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF91FFCCFF02F87BFC9DF7DE" pageId="7" pageNumber="246" type="conservation">
<paragraph id="960436A3FF91FFCCFF02F87BFC9DF7DE" blockId="7.[136,1452,1058,2057]" pageId="7" pageNumber="246">
<emphasis id="A4CFEAB1FF91FFCCFF02F87BFDAEF816" bold="true" box="[189,513,1958,1984]" pageId="7" pageNumber="246">IUCN Conservation status:</emphasis>
—The plant is known from single locality in
<collectingCountry id="EEAC7633FF91FFCCFC4CF87BFBF5F816" box="[1011,1114,1958,1984]" name="Thailand" pageId="7" pageNumber="246">Thailand</collectingCountry>
. In Peninsular
<collectingCountry id="EEAC7633FF91FFCCFB42F87BFACBF816" box="[1277,1380,1958,1984]" name="Malaysia" pageId="7" pageNumber="246">Malaysia</collectingCountry>
it was reported from two widely separated localities in the original description but it has not been re-collected since 1896. Therefore, this species is considered as Data Deficient (DD).
</paragraph>
</subSubSection>
<caption id="C2C4662BFF9EFFC3FF37F840FE8EF7F6" ID-DOI="http://doi.org/10.5281/zenodo.5426431" ID-Zenodo-Dep="5426431" httpUri="https://zenodo.org/record/5426431/files/figure.png" pageId="8" pageNumber="247" startId="8.[136,229,1949,1971]" targetBox="[177,1387,203,1911]" targetPageId="8">
<paragraph id="960436A3FF9EFFC3FF37F840FE8EF7F6" blockId="8.[136,1452,1949,2080]" pageId="8" pageNumber="247">
<emphasis id="A4CFEAB1FF9EFFC3FF37F840FF52F865" bold="true" box="[136,253,1949,1971]" pageId="8" pageNumber="247">FIGURE 5.</emphasis>
<taxonomicName id="51BB4D20FF9EFFC3FEBCF840FDBEF865" authority=": A. Habit" authorityName="A. Habit" box="[259,529,1949,1971]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="8" pageNumber="247" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="A4CFEAB1FF9EFFC3FEBCF840FE1FF864" box="[259,432,1949,1970]" italics="true" pageId="8" pageNumber="247">Polyalthia pumila</emphasis>
: A. Habit
</taxonomicName>
; B. Inflorescence, showing supra-axillary attachment; C. Base of flower, with petals removed to show stamens and carpels; D. Sepals (outside below, inside above); E. Outside of outer petal; F. Outside of inner petal; G. Stamens (inside view on left, outside view on right); H. Carpel; I. Fruit; J. Monocarp. The illustration was drawn by Thadanai Saensuk from
<emphasis id="A4CFEAB1FF9EFFC3FA8DF83BFF77F7C9" italics="true" pageId="8" pageNumber="247">Leeratiwong 20-1528</emphasis>
(PSU).
</paragraph>
</caption>
<caption id="C2C4662BFF9FFFC2FF37FA26FF5AF9AB" ID-DOI="http://doi.org/10.5281/zenodo.5426433" ID-Zenodo-Dep="5426433" httpUri="https://zenodo.org/record/5426433/files/figure.png" pageId="9" pageNumber="248" startId="9.[136,229,1531,1553]" targetBox="[167,1427,205,1480]" targetPageId="9">
<paragraph id="960436A3FF9FFFC2FF37FA26FF5AF9AB" blockId="9.[136,1451,1531,1661]" pageId="9" pageNumber="248">
<emphasis id="A4CFEAB1FF9FFFC2FF37FA26FF52F9C6" bold="true" box="[136,253,1531,1553]" pageId="9" pageNumber="248">FIGURE 6.</emphasis>
<taxonomicName id="51BB4D20FF9FFFC2FEBCFA26FDA6F9C7" authority=": A. Twig" authorityName="A. Twig" box="[259,521,1531,1553]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="9" pageNumber="248" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="A4CFEAB1FF9FFFC2FEBCFA26FE1FF9C6" box="[259,432,1531,1552]" italics="true" pageId="9" pageNumber="248">Polyalthia pumila</emphasis>
: A. Twig
</taxonomicName>
and flowers; B. Flowers and a single monocarp; C. Upper surface of leaf base; D. Lower surface of leaf; E. Fruit; F. Seed (Side view). Photographs in A &amp; B were taken Charan Leeratiwong in Narathiwat, Cha Nae District, Dusong Yo Subdistrict, Ruepoh village, Mo Tae Mountain, 4 April 2020. C, D, E &amp; F are from
<emphasis id="A4CFEAB1FF9FFFC2FC21F999FBC2F98E" box="[926,1133,1603,1625]" italics="true" pageId="9" pageNumber="248">Leeratiwong 20-1528</emphasis>
(PSU); photographs by Pasakorn Bunchalee.
</paragraph>
</caption>
<subSubSection id="DEA16528FF9FFFC2FF02F976FB1FF93C" pageId="9" pageNumber="248" type="materials_examined">
<paragraph id="960436A3FF9FFFC2FF02F976FB1FF93C" blockId="9.[136,1452,1707,1913]" pageId="9" pageNumber="248">
<emphasis id="A4CFEAB1FF9FFFC2FF02F976FE17F913" bold="true" box="[189,440,1707,1733]" pageId="9" pageNumber="248">Specimen examined:</emphasis>
<materialsCitation id="26D33CFEFF9FFFC2FE6BF971FB03F93C" collectingDate="2020-04-04" country="Thailand" county="Cha Nae District" location="Ruepoh village" municipality="Dusong Yo Subdistrict" pageId="9" pageNumber="248" specimenCount="1" stateProvince="Narathiwat">
<collectingCountry id="EEAC7633FF9FFFC2FE6BF971FDC7F910" box="[468,616,1708,1734]" name="Thailand" pageId="9" pageNumber="248">THAILAND</collectingCountry>
. PENINSULAR:
<collectingRegion id="547FF841FF9FFFC2FC86F976FC17F913" box="[825,952,1707,1733]" country="Thailand" name="Narathiwat" pageId="9" pageNumber="248">Narathiwat</collectingRegion>
,
<collectingCounty id="7F654E2FFF9FFFC2FC78F976FB20F913" box="[967,1167,1707,1734]" pageId="9" pageNumber="248">Cha Nae District</collectingCounty>
,
<collectingMunicipality id="7660ACD9FF9FFFC2FB20F971FA08F913" box="[1183,1447,1707,1734]" pageId="9" pageNumber="248">Dusong Yo Subdistrict</collectingMunicipality>
,
<location id="93646078FF9FFFC2FF37F912FE9AF93F" LSID="urn:lsid:plazi:treatment:1E1287B5FF91FFC2FF37FC4FFDA1F8AF:93646078FF9FFFC2FF37F912FE9AF93F" box="[136,309,1743,1769]" country="Thailand" county="Cha Nae District" municipality="Dusong Yo Subdistrict" name="Ruepoh village" pageId="9" pageNumber="248" stateProvince="Narathiwat">Ruepoh village</location>
,
<location id="93646078FF9FFFC2FEFFF90DFDA2F93F" LSID="urn:lsid:plazi:treatment:1E1287B5FF91FFC2FF37FC4FFDA1F8AF:93646078FF9FFFC2FEFFF90DFDA2F93F" box="[320,525,1743,1770]" country="Thailand" county="Cha Nae District" municipality="Dusong Yo Subdistrict" name="Mo Tae Mountain" pageId="9" pageNumber="248" stateProvince="Narathiwat">Mo Tae Mountain</location>
,
<date id="E2051063FF9FFFC2FDA7F90DFD0AF93C" box="[536,677,1743,1770]" pageId="9" pageNumber="248" value="2020-04-04">
<collectingDate id="F241E98BFF9FFFC2FDA7F90DFD0AF93C" box="[536,677,1743,1770]" pageId="9" pageNumber="248" value="2020-04-04">4 April 2020</collectingDate>
</date>
(flower &amp; fruit),
<emphasis id="A4CFEAB1FF9FFFC2FCD5F90DFBF4F93C" box="[874,1115,1744,1770]" italics="true" pageId="9" pageNumber="248">Leeratiwong 20-1528</emphasis>
(PSU)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF9FFFC2FF02F929FDA1F8AF" pageId="9" pageNumber="248" type="discussion">
<paragraph id="960436A3FF9FFFC2FF02F929FDA1F8AF" blockId="9.[136,1452,1707,1913]" pageId="9" pageNumber="248">
<emphasis id="A4CFEAB1FF9FFFC2FF02F929FEA6F8D8" bold="true" box="[189,265,1780,1806]" pageId="9" pageNumber="248">Notes:</emphasis>
—This is the first report of
<taxonomicName id="51BB4D20FF9FFFC2FD87F92EFCAEF8DB" box="[568,769,1779,1805]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="9" pageNumber="248" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="A4CFEAB1FF9FFFC2FD87F92EFCAEF8DB" box="[568,769,1779,1805]" italics="true" pageId="9" pageNumber="248">Polyalthia pumila</emphasis>
</taxonomicName>
for
<collectingCountry id="EEAC7633FF9FFFC2FC92F92EFC3BF8DB" box="[813,916,1779,1805]" name="Thailand" pageId="9" pageNumber="248">Thailand</collectingCountry>
. The species is notable for its stature, among the shortest in the family (
<bibRefCitation id="F22A4B52FF9FFFC2FE3BF8CAFD8FF8E4" author="Sinclair, J." box="[388,544,1815,1842]" pageId="9" pageNumber="248" pagination="149 - 516" refId="ref5266" refString="Sinclair, J. (1955) A revision of the Malayan Annonaceae. Gardens' Bulletin, Singapore 14: 149 - 516." type="journal article" year="1955">Sinclair 1955</bibRefCitation>
). It is similar to
<taxonomicName id="51BB4D20FF9FFFC2FD67F8C5FCC7F8E7" authorityName=", Turner" authorityYear="2014" box="[728,872,1815,1841]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="9" pageNumber="248" phylum="Tracheophyta" rank="species" species="motleyana">
<emphasis id="A4CFEAB1FF9FFFC2FD67F8C5FCC7F8E7" box="[728,872,1815,1841]" italics="true" pageId="9" pageNumber="248">P. motleyana</emphasis>
</taxonomicName>
and its relatives in having branches with distichous phyllotaxis, a lamina with a more or less bullate surface, the leaf base more or less cordate, and secondary veins with simple brochidodromous venation.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,391 @@
<document id="0D0A6A769FD3C5BA84742468ED2E32D1" ID-DOI="10.11646/phytotaxa.510.3.4" ID-ISSN="1179-3163" ID-Zenodo-Dep="5426421" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="operationResults" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1630397611626" checkinUser="marcus" docAuthor="Bunchalee, Pasakorn, Leeratiwong, Charan &amp; Johnson, David M." docDate="2021" docId="1E1287B5FF92FFCCFF37FCF1FAE6FCB2" docLanguage="en" docName="phytotaxa.510.3.4.pdf" docOrigin="Phytotaxa 510 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.510.3.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Polyalthia taweensis Bunchalee &amp; Leerat. 2021, sp. nov." docType="treatment" docVersion="1" lastPageNumber="246" masterDocId="E22BFFCDFF96FFCBFFBFFFDDFFAFFFD6" masterDocTitle="Two new species and a new record of the genus Polyalthia (Annonaceae) from Peninsular Thailand" masterLastPageNumber="250" masterPageNumber="239" pageNumber="243" updateTime="1732197805484" updateUser="guilherme" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="866934010240641D99C5AB9AFED89450">Two new species and a new record of the genus Polyalthia (Annonaceae) from Peninsular Thailand</mods:title>
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<mods:name id="640C148DF5BD03F3C45978F978E6F81E" type="personal">
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<mods:namePart id="E1499E017F372F382E20926FFD577F46">Bunchalee, Pasakorn</mods:namePart>
<mods:nameIdentifier id="7F0A1F2235EFC516B3A4CF7B34E3F26F" type="ORCID">0000-0001-8644-3360</mods:nameIdentifier>
<mods:affiliation id="7C9A8C4EC11095A30FD21612A408F56B">Department of Biology, Faculty of Science, Mahasarakham University, Kantarawichai, Maha Sarakham, 44150, Thailand. Palaeontological Research and Education Center, Mahasarakham University, Kantarawichai, Maha Sarakham, 44150, Thailand. Division of Biological Science, Faculty of Science, Prince of Songkla University, Hat Yai, Songkhla 90112, Thailand. &amp; pasakorn. b @ msu. ac. th; https: // orcid. org / 0000 - 0001 - 8644 - 3360</mods:affiliation>
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<mods:namePart id="23AB548C9F4747FDC8031B23EE1F50C3">Leeratiwong, Charan</mods:namePart>
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<mods:affiliation id="3338383B36CB1CA75D009161D9464995">Department of Botany and Microbiology, Ohio Wesleyan University, Delaware, Ohio, 43015, USA &amp; dmjohnso @ owu. edu; https: // orcid. org / 0000 - 0003 - 2896 - 7419</mods:affiliation>
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<treatment id="1E1287B5FF92FFCCFF37FCF1FAE6FCB2" LSID="urn:lsid:plazi:treatment:1E1287B5FF92FFCCFF37FCF1FAE6FCB2" httpUri="http://treatment.plazi.org/id/1E1287B5FF92FFCCFF37FCF1FAE6FCB2" lastPageId="7" lastPageNumber="246" pageId="4" pageNumber="243">
<subSubSection id="DEA16528FF92FFCFFF37FCF1FCCCFC91" box="[136,867,812,839]" pageId="4" pageNumber="243" type="nomenclature">
<paragraph id="960436A3FF92FFCFFF37FCF1FCCCFC91" blockId="4.[136,867,812,839]" box="[136,867,812,839]" pageId="4" pageNumber="243">
<taxonomicName id="51BB4D20FF92FFCFFF37FCF1FDC2FC91" ID-CoL="C8352" authority="Bunchalee &amp; Leerat." authorityName="Bunchalee &amp; Leerat." authorityYear="2021" box="[136,621,812,839]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="4" pageNumber="243" phylum="Tracheophyta" rank="species" species="taweensis" status="sp. nov.">
<emphasis id="A4CFEAB1FF92FFCFFF37FCF1FED5FC90" bold="true" box="[136,378,812,838]" pageId="4" pageNumber="243">Polyalthia taweensis</emphasis>
Bunchalee &amp; Leerat.
</taxonomicName>
,
<emphasis id="A4CFEAB1FF92FFCFFDC8FCF3FD63FC91" box="[631,716,814,839]" italics="true" pageId="4" pageNumber="243">
<taxonomicNameLabel id="BFFC57CAFF92FFCFFDC8FCF3FD63FC91" box="[631,716,814,839]" pageId="4" pageNumber="243" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="0E802A26FF92FFCFFD64FCF0FC87FC91" box="[731,808,813,839]" captionStart="FIGURE 3" captionStartId="5.[136,229,1955,1977]" captionTargetBox="[209,1375,206,1919]" captionTargetId="figure-18@5.[173,1415,187,1933]" captionTargetPageId="5" captionText="FIGURE 3. Polyalthia taweensis: A. Habit; B. Close-up of petiole and leaf base (lower side); C. Inflorescence; D. Flower; E. Sepals (outside above, inside below); F. Outside of outer petal; G. Inside of outer petal, showing raised rugose area (arrow); H. Outside of inner petal; I. Inside of inner petal; J. Inner petal, side view showing inward curvature to midpoint (arrow) and erect apex; K. Carpel; L. Stamen, outer view. The illustration was drawn by Nattapong Chomlok from Leeratiwong 20-1534 (BKF)." figureDoi="http://doi.org/10.5281/zenodo.5426427" httpUri="https://zenodo.org/record/5426427/files/figure.png" pageId="4" pageNumber="243">Figs. 3</figureCitation>
&amp;
<figureCitation id="0E802A26FF92FFCFFCF3FCF0FCF4FC91" box="[844,859,813,839]" captionStart="FIGURE 4" captionStartId="6.[136,229,1918,1940]" captionTargetBox="[170,1423,199,1885]" captionTargetId="figure-21@6.[151,1436,187,1894]" captionTargetPageId="6" captionText="FIGURE 4. Polyalthia taweensis: A. Leaves and inflorescences; B. Upper surface of leaf base; C. Lower surface of leaf base; D. Inflorescence; E. Sepals (outside on left, inside on light); F. Outsides of petals (outer petal on left, inner petal on right); G. Insides of petals (outer petal showing raised rugose border on left, inner petal on right). A, B &amp; C are from Middleton, Phuphat, Pooma &amp; Williams 2900 (BKF); E, F &amp; G are from Leeratiwong 20-1534 (PSU, spirit material); all photographs by Pasakorn Bunchalee. Photograph in D was taken by Yamilah Sama-ae in Narathiwat, Ra Ngae District, Bo Ngo Subdistrict, Ta We Mountain, 20 April 2020." figureDoi="http://doi.org/10.5281/zenodo.5426429" httpUri="https://zenodo.org/record/5426429/files/figure.png" pageId="4" pageNumber="243">4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF92FFCFFF37FCA5FEB4FB94" pageId="4" pageNumber="243" type="diagnosis">
<paragraph id="960436A3FF92FFCFFF37FCA5FEB4FB94" blockId="4.[136,1451,888,1162]" pageId="4" pageNumber="243">
<taxonomicName id="51BB4D20FF92FFCFFF37FCA5FEFFFC58" box="[136,336,888,910]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="4" pageNumber="243" phylum="Tracheophyta" rank="species" species="taweensis">
<emphasis id="A4CFEAB1FF92FFCFFF37FCA5FEFFFC58" box="[136,336,888,910]" italics="true" pageId="4" pageNumber="243">Polyalthia taweensis</emphasis>
</taxonomicName>
resembles
<taxonomicName id="51BB4D20FF92FFCFFE02FCA4FDCDFC58" authority="Hook." authorityName="Hook." box="[445,610,888,910]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="4" pageNumber="243" phylum="Tracheophyta" rank="species" species="obliqua">
<emphasis id="A4CFEAB1FF92FFCFFE02FCA4FDB1FC5B" box="[445,542,888,910]" italics="true" pageId="4" pageNumber="243">P. obliqua</emphasis>
Hook.
</taxonomicName>
f. &amp; Thomson (1855: 138) in leaf shape and size and the shape of the petals, but differs in having supra-axillary or leaf-opposed inflorescences, greenish yellow petals, a differentiated border along the margin on the inner surface of the outer petals, sepals 4.05.0 mm long, and one pedicellar bract. In contrast,
<taxonomicName id="51BB4D20FF92FFCFFB9CFC1CFB2AFC03" box="[1059,1157,960,982]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="4" pageNumber="243" phylum="Tracheophyta" rank="species" species="obliqua">
<emphasis id="A4CFEAB1FF92FFCFFB9CFC1CFB2AFC03" box="[1059,1157,960,982]" italics="true" pageId="4" pageNumber="243">P. obliqua</emphasis>
</taxonomicName>
has axillary inflorescences on leafy or leafless branches, white petals, the inner with a rosy flush at the base (
<bibRefCitation id="F22A4B52FF92FFCFFC03FC39FBEDFC2C" author="Sinclair, J." box="[956,1090,996,1018]" pageId="4" pageNumber="243" pagination="149 - 516" refId="ref5266" refString="Sinclair, J. (1955) A revision of the Malayan Annonaceae. Gardens' Bulletin, Singapore 14: 149 - 516." type="journal article" year="1955">Sinclair 1955</bibRefCitation>
), the inner surface of the outer petals uniform in texture, sepals ca. 2.0 mm long, and two pedicellar bracts, one attached at the pedicel base and the other at the pedicel midpoint.
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF92FFCFFF37FB8CFE5EFB5C" pageId="4" pageNumber="243" type="materials_examined">
<paragraph id="960436A3FF92FFCFFF37FB8CFE5EFB5C" blockId="4.[136,1451,888,1162]" pageId="4" pageNumber="243">
<collectingCounty id="7F654E2FFF92FFCFFF37FB8CFF15FBB0" box="[136,186,1105,1126]" pageId="4" pageNumber="243">Type</collectingCounty>
:—
<materialsCitation id="26D33CFEFF92FFCFFF69FB8CFEF8FB5F" collectingDate="2020-04-20" collectionCode="PSU" collectorName="Leeratiwong" country="Thailand" location="Bo Ngo Subdistrict" municipality="Ra-Ngae District" pageId="4" pageNumber="243" specimenCount="1" stateProvince="Narathiwat" typeStatus="holotype">
<collectingCountry id="EEAC7633FF92FFCFFF69FB8CFEF9FBB0" box="[214,342,1105,1126]" name="Thailand" pageId="4" pageNumber="243">THAILAND</collectingCountry>
.
<collectingRegion id="547FF841FF92FFCFFEE1FB8DFE64FBB0" box="[350,459,1104,1126]" country="Thailand" name="Narathiwat" pageId="4" pageNumber="243">Narathiwat</collectingRegion>
,
<collectingMunicipality id="7660ACD9FF92FFCFFE6AFB8CFDD4FBB0" box="[469,635,1104,1126]" pageId="4" pageNumber="243">Ra-Ngae District</collectingMunicipality>
,
<location id="93646078FF92FFCFFD3BFB8CFCEEFBB0" LSID="urn:lsid:plazi:treatment:1E1287B5FF92FFCCFF37FCF1FAE6FCB2:93646078FF92FFCFFD3BFB8CFCEEFBB0" box="[644,833,1104,1126]" country="Thailand" county="Type" municipality="Ra-Ngae District" name="Bo Ngo Subdistrict" pageId="4" pageNumber="243" stateProvince="Narathiwat">Bo Ngo Subdistrict</location>
,
<location id="93646078FF92FFCFFCF5FB8CFC43FBB0" LSID="urn:lsid:plazi:treatment:1E1287B5FF92FFCCFF37FCF1FAE6FCB2:93646078FF92FFCFFCF5FB8CFC43FBB0" box="[842,1004,1104,1126]" country="Thailand" county="Type" municipality="Ra-Ngae District" name="Ta We Mountain" pageId="4" pageNumber="243" stateProvince="Narathiwat">Ta We Mountain</location>
,
<date id="E2051063FF92FFCFFC49FB8DFBD4FBB3" box="[1014,1147,1104,1126]" pageId="4" pageNumber="243" value="2020-04-20">
<collectingDate id="F241E98BFF92FFCFFC49FB8DFBD4FBB3" box="[1014,1147,1104,1126]" pageId="4" pageNumber="243" value="2020-04-20">20 April 2020</collectingDate>
</date>
(flower),
<emphasis id="A4CFEAB1FF92FFCFFB63FB8CFA04FBB3" box="[1244,1451,1104,1126]" italics="true" pageId="4" pageNumber="243">
<collectorName id="3B4E5375FF92FFCFFB63FB8CFAFAFBB0" box="[1244,1365,1105,1126]" pageId="4" pageNumber="243">Leeratiwong</collectorName>
20-1534
</emphasis>
(
<typeStatus id="49008801FF92FFCFFF7BFBA9FEB5FB5C" box="[196,282,1140,1162]" pageId="4" pageNumber="243" type="holotype">holotype</typeStatus>
<collectionCode id="F0AAAE66FF92FFCFFEA0FBA9FEFCFB5F" box="[287,339,1140,1161]" pageId="4" pageNumber="243">PSU</collectionCode>
!
</materialsCitation>
;
<materialsCitation id="26D33CFEFF92FFCFFEDFFBA9FE49FB5C" box="[352,486,1140,1162]" collectionCode="BKF" pageId="4" pageNumber="243" specimenCount="1" typeStatus="isotype">
<typeStatus id="49008801FF92FFCFFEDFFBA9FE0AFB5C" box="[352,421,1140,1162]" pageId="4" pageNumber="243">isotype</typeStatus>
<collectionCode id="F0AAAE66FF92FFCFFE14FBA9FE4DFB5C" box="[427,482,1140,1162]" collectionName="BKF" pageId="4" pageNumber="243">BKF</collectionCode>
!
</materialsCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF92FFCFFF37FB64FEDBF7D9" pageId="4" pageNumber="243" type="description">
<paragraph id="960436A3FF92FFCFFF37FB64FEDBF7D9" blockId="4.[136,1452,1209,2063]" pageId="4" pageNumber="243">
Shrubs or trees 2.012.0 m tall, d.b.h. up to 18.0 cm, bark smooth, greyish black or grey-brown, primary branching spiral.
<emphasis id="A4CFEAB1FF92FFCFFF69FB03FE25FB21" box="[214,394,1245,1271]" italics="true" pageId="4" pageNumber="243">Young branches</emphasis>
glabrous with prominent lenticels.
<emphasis id="A4CFEAB1FF92FFCFFCA3FB03FCC5FB21" box="[796,874,1246,1271]" italics="true" pageId="4" pageNumber="243">Leaves</emphasis>
with petioles 1.53.0 mm long, 1.52.0 mm thick, sunken above; lamina chartaceous, symmetrically or asymmetrically elliptic to slightly oblanceolate and curved inward, 13.020.0 ×
<quantity id="51439B46FF92FFCFFEA6FAF8FE34FAE9" box="[281,411,1317,1344]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="7.5" metricValueMin="3.5" pageId="4" pageNumber="243" unit="cm" value="5.5" valueMax="7.5" valueMin="3.5">3.57.5 cm</quantity>
, base asymmetrically cordate with auricles
<quantity id="51439B46FF92FFCFFC31FAF8FBB9FAE9" box="[910,1046,1317,1343]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="4" pageNumber="243" unit="mm" value="1.0" valueMax="1.2" valueMin="0.8">0.81.2 mm</quantity>
long, apex acuminate with acumen 10.015.0 mm long, upper side glabrous, lower side glabrescent; midrib prominently grooved above; lateral veins 14 16 per side, arched towards the apex, diverging at 6070° from the midrib, festooned loop-forming 2.05.0 mm from the margins; intersecondary veins indistinct; tertiary veins reticulate.
<emphasis id="A4CFEAB1FF92FFCFFC30FA4CFB9FFA7D" box="[911,1072,1425,1451]" italics="true" pageId="4" pageNumber="243">Inflorescences</emphasis>
supra-axillary or leaf-opposed on leafy branches, 12-flowered; peduncle up to
<quantity id="51439B46FF92FFCFFD34FA68FD63FA19" box="[651,716,1461,1487]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="243" unit="mm" value="1.0">1 mm</quantity>
long, pedicels 7.010.0 mm long, 0.81.0 mm thick, puberulous; bract solitary, triangular or lanceolate, 3.04.0 ×
<quantity id="51439B46FF92FFCFFD10FA04FC96FA25" box="[687,825,1497,1524]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="4" pageNumber="243" unit="mm" value="1.35" valueMax="1.5" valueMin="1.2">1.21.5 mm</quantity>
, attached from near the pedicel base to near the pedicel apex, apex acute, puberulous outside and glabrous inside.
<emphasis id="A4CFEAB1FF92FFCFFCADFA20FCF4F9C1" box="[786,859,1533,1559]" italics="true" pageId="4" pageNumber="243">Sepals</emphasis>
valvate, light green
<emphasis id="A4CFEAB1FF92FFCFFBFEFA23FB23F9C1" box="[1089,1164,1534,1559]" italics="true" pageId="4" pageNumber="243">in vivo</emphasis>
, thinly coriaceous, ovate, 4.05.0 × 3.04.0 mm, apex acute to slightly acuminate, puberulous outside and glabrous inside.
<emphasis id="A4CFEAB1FF92FFCFFB43F9FCFAEDF9ED" box="[1276,1346,1569,1595]" italics="true" pageId="4" pageNumber="243">Petals</emphasis>
greenish yellow
<emphasis id="A4CFEAB1FF92FFCFFF61F99BFE82F989" box="[222,301,1606,1631]" italics="true" pageId="4" pageNumber="243">in vivo</emphasis>
, erect or slightly spreading, coriaceous, apex acute, pubescent outside, glabrous inside; outer petals oblong or ovate, 7.08.0 × 4.05.0 mm, smooth outside, with a raised rugose border along margin inside (
<figureCitation id="0E802A26FF92FFCFFAFDF9B4FA31F955" box="[1346,1438,1641,1667]" captionStart="FIGURE 3" captionStartId="5.[136,229,1955,1977]" captionTargetBox="[209,1375,206,1919]" captionTargetId="figure-18@5.[173,1415,187,1933]" captionTargetPageId="5" captionText="FIGURE 3. Polyalthia taweensis: A. Habit; B. Close-up of petiole and leaf base (lower side); C. Inflorescence; D. Flower; E. Sepals (outside above, inside below); F. Outside of outer petal; G. Inside of outer petal, showing raised rugose area (arrow); H. Outside of inner petal; I. Inside of inner petal; J. Inner petal, side view showing inward curvature to midpoint (arrow) and erect apex; K. Carpel; L. Stamen, outer view. The illustration was drawn by Nattapong Chomlok from Leeratiwong 20-1534 (BKF)." figureDoi="http://doi.org/10.5281/zenodo.5426427" httpUri="https://zenodo.org/record/5426427/files/figure.png" pageId="4" pageNumber="243">Fig. 3G</figureCitation>
); inner petals elliptic, 8.09.0 × 3.04.0 mm, curved inward to the midpoint but with the apex erect (
<figureCitation id="0E802A26FF92FFCFFB57F950FA97F971" box="[1256,1336,1677,1703]" captionStart="FIGURE 3" captionStartId="5.[136,229,1955,1977]" captionTargetBox="[209,1375,206,1919]" captionTargetId="figure-18@5.[173,1415,187,1933]" captionTargetPageId="5" captionText="FIGURE 3. Polyalthia taweensis: A. Habit; B. Close-up of petiole and leaf base (lower side); C. Inflorescence; D. Flower; E. Sepals (outside above, inside below); F. Outside of outer petal; G. Inside of outer petal, showing raised rugose area (arrow); H. Outside of inner petal; I. Inside of inner petal; J. Inner petal, side view showing inward curvature to midpoint (arrow) and erect apex; K. Carpel; L. Stamen, outer view. The illustration was drawn by Nattapong Chomlok from Leeratiwong 20-1534 (BKF)." figureDoi="http://doi.org/10.5281/zenodo.5426427" httpUri="https://zenodo.org/record/5426427/files/figure.png" pageId="4" pageNumber="243">Fig. 3J</figureCitation>
).
<emphasis id="A4CFEAB1FF92FFCFFAF1F950FA04F971" box="[1358,1451,1677,1703]" italics="true" pageId="4" pageNumber="243">Stamens</emphasis>
pale yellow, cuneate, 1.01.2 × 0.81.0 mm, filament ca.
<quantity id="51439B46FF92FFCFFCA7F96CFCC0F91D" box="[792,879,1713,1739]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="243" unit="mm" value="0.2">0.2 mm</quantity>
long; anthers
<quantity id="51439B46FF92FFCFFBADF96CFB34F91D" box="[1042,1179,1713,1739]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="4" pageNumber="243" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">0.60.8 mm</quantity>
long, anther connective apex flat; androecium 3.04.0 mm in diameter.
<emphasis id="A4CFEAB1FF92FFCFFD06F908FCBDF939" box="[697,786,1749,1775]" italics="true" pageId="4" pageNumber="243">Carpels</emphasis>
57 per flower, 1.61.8 ×
<quantity id="51439B46FF92FFCFFBE9F908FB4AF939" box="[1110,1253,1749,1776]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="4" pageNumber="243" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.50.6 mm</quantity>
, pubescent, style rudimentary, stigma obovoid,
<quantity id="51439B46FF92FFCFFE60F924FDC5F8C5" box="[479,618,1785,1812]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="4" pageNumber="243" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.50.6 mm</quantity>
, pubescent, exceeding the height of the anther connective apices; ovules 2 per carpel, attached laterally.
<emphasis id="A4CFEAB1FF92FFCFFE52F8C3FD83F8E1" box="[493,556,1822,1847]" italics="true" pageId="4" pageNumber="243">Torus</emphasis>
convex, flat at the apex, 3.0
<quantity id="51439B46FF92FFCFFC3FF8C0FC79F8E1" box="[896,982,1821,1848]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="4" pageNumber="243" unit="mm" value="3.5">3.5 mm</quantity>
in diameter, 1.0
<quantity id="51439B46FF92FFCFFB21F8C0FB5AF8E1" box="[1182,1269,1821,1847]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="243" unit="mm" value="1.2">1.2 mm</quantity>
thick, glabrous.
<emphasis id="A4CFEAB1FF92FFCFFF37F89FFF6DF88D" box="[136,194,1858,1883]" italics="true" pageId="4" pageNumber="243">Fruit</emphasis>
of ca. 12 monocarps borne on a pedicel ca. 17.0 mm long,
<quantity id="51439B46FF92FFCFFCDEF89CFC17F88D" box="[865,952,1857,1883]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="243" unit="mm" value="1.8">1.8 mm</quantity>
thick.
<emphasis id="A4CFEAB1FF92FFCFFBB9F89FFB29F88D" box="[1030,1158,1858,1883]" italics="true" pageId="4" pageNumber="243">Monocarps</emphasis>
green
<emphasis id="A4CFEAB1FF92FFCFFB6CF89FFAB0F88D" box="[1235,1311,1858,1883]" italics="true" pageId="4" pageNumber="243">in vivo</emphasis>
(immature), subglobose, 12.016.0 × 14.016.0 mm, slightly puckered and verrucose, glabrate, apex rounded, occasionally with an apiculum &lt;1.0 mm long, base rounded, stipes ca. 14.0 mm long,
<quantity id="51439B46FF92FFCFFC20F854FC58F875" box="[927,1015,1929,1956]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="243" unit="mm" value="1.5">1.5 mm</quantity>
thick, rugulose, sparsely pubescent to glabrate.
<emphasis id="A4CFEAB1FF92FFCFFF4FF870FE9FF811" box="[240,304,1965,1991]" italics="true" pageId="4" pageNumber="243">Seeds</emphasis>
12 per monocarp, attached laterally in a single row, disk-shaped, 9.812.0 × 8.011.0 ×
<quantity id="51439B46FF92FFCFFAA1F870FA08F811" box="[1310,1447,1965,1991]" metricMagnitude="-3" metricUnit="m" metricValue="5.449999999999999" metricValueMax="6.1" metricValueMin="4.8" pageId="4" pageNumber="243" unit="mm" value="5.449999999999999" valueMax="6.1" valueMin="4.8">4.86.1 mm</quantity>
, light brown, slightly shiny, surface densely wrinkled, with a distinct circumferential groove; endosperm rumination pattern not observed.
</paragraph>
</subSubSection>
<caption id="C2C4662BFF93FFCEFF37F87EFB99F7F0" ID-DOI="http://doi.org/10.5281/zenodo.5426427" ID-Zenodo-Dep="5426427" httpUri="https://zenodo.org/record/5426427/files/figure.png" pageId="5" pageNumber="244" startId="5.[136,229,1955,1977]" targetBox="[209,1375,206,1919]" targetPageId="5">
<paragraph id="960436A3FF93FFCEFF37F87EFB99F7F0" blockId="5.[136,1452,1955,2086]" pageId="5" pageNumber="244">
<emphasis id="A4CFEAB1FF93FFCEFF37F87EFF50F86E" bold="true" box="[136,255,1955,1977]" pageId="5" pageNumber="244">FIGURE 3.</emphasis>
<taxonomicName id="51BB4D20FF93FFCEFEB8F87EFD9AF86F" authority=": A. Habit" authorityName="A. Habit" box="[263,565,1955,1977]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="5" pageNumber="244" phylum="Tracheophyta" rank="species" species="taweensis">
<emphasis id="A4CFEAB1FF93FFCEFEB8F87EFE7FF86F" box="[263,464,1955,1977]" italics="true" pageId="5" pageNumber="244">Polyalthia taweensis</emphasis>
: A. Habit
</taxonomicName>
; B. Close-up of petiole and leaf base (lower side); C. Inflorescence; D. Flower; E. Sepals (outside above, inside below); F. Outside of outer petal; G. Inside of outer petal, showing raised rugose area (arrow); H. Outside of inner petal; I. Inside of inner petal; J. Inner petal, side view showing inward curvature to midpoint (arrow) and erect apex; K. Carpel; L. Stamen, outer view. The illustration was drawn by Nattapong Chomlok from
<emphasis id="A4CFEAB1FF93FFCEFCA4F7CDFC45F7F3" box="[795,1002,2064,2085]" italics="true" pageId="5" pageNumber="244">Leeratiwong 20-1534</emphasis>
(BKF).
</paragraph>
</caption>
<caption id="C2C4662BFF90FFCDFF37F8A3FB67F7F2" ID-DOI="http://doi.org/10.5281/zenodo.5426429" ID-Zenodo-Dep="5426429" httpUri="https://zenodo.org/record/5426429/files/figure.png" pageId="6" pageNumber="245" startId="6.[136,229,1918,1940]" targetBox="[170,1423,199,1885]" targetPageId="6">
<paragraph id="960436A3FF90FFCDFF37F8A3FB67F7F2" blockId="6.[136,1452,1918,2084]" pageId="6" pageNumber="245">
<emphasis id="A4CFEAB1FF90FFCDFF37F8A3FEAEF845" bold="true" box="[136,257,1918,1940]" pageId="6" pageNumber="245">FIGURE 4.</emphasis>
<taxonomicName id="51BB4D20FF90FFCDFEB4F8A3FDE5F842" authority=": A. Leaves" authorityName="A. Leaves" box="[267,586,1918,1940]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="6" pageNumber="245" phylum="Tracheophyta" rank="species" species="taweensis">
<emphasis id="A4CFEAB1FF90FFCDFEB4F8A3FE79F842" box="[267,470,1918,1940]" italics="true" pageId="6" pageNumber="245">Polyalthia taweensis</emphasis>
: A. Leaves
</taxonomicName>
and inflorescences; B. Upper surface of leaf base; C. Lower surface of leaf base; D. Inflorescence; E. Sepals (outside on left, inside on light); F. Outsides of petals (outer petal on left, inner petal on right); G. Insides of petals (outer petal showing raised rugose border on left, inner petal on right). A, B &amp; C are from
<emphasis id="A4CFEAB1FF90FFCDFC4EF81BFA04F80A" box="[1009,1451,1990,2012]" italics="true" pageId="6" pageNumber="245">Middleton, Phuphat, Pooma &amp; Williams 2900</emphasis>
(BKF); E, F &amp; G are from
<emphasis id="A4CFEAB1FF90FFCDFE2AF836FDCAF7D6" box="[405,613,2027,2048]" italics="true" pageId="6" pageNumber="245">Leeratiwong 20-1534</emphasis>
(PSU, spirit material); all photographs by Pasakorn Bunchalee. Photograph in D was taken by Yamilah Sama-ae in Narathiwat, Ra Ngae District, Bo Ngo Subdistrict, Ta We Mountain, 20 April 2020.
</paragraph>
</caption>
<subSubSection id="DEA16528FF91FFCCFF02FF43FD97FF0B" pageId="7" pageNumber="246" type="distribution">
<paragraph id="960436A3FF91FFCCFF02FF43FD97FF0B" blockId="7.[136,1452,158,869]" pageId="7" pageNumber="246">
<emphasis id="A4CFEAB1FF91FFCCFF02FF43FE58FF6E" bold="true" box="[189,503,158,184]" pageId="7" pageNumber="246">Distribution and Ecology:</emphasis>
—Occurs in tropical rain forest at
<quantity id="51439B46FF91FFCCFC3CFF43FBABFF6F" box="[899,1028,158,185]" metricMagnitude="2" metricUnit="m" metricValue="1.85" metricValueMax="2.5" metricValueMin="1.2" pageId="7" pageNumber="246" unit="m" value="185.0" valueMax="250.0" valueMin="120.0">120250 m</quantity>
in
<collectingRegion id="547FF841FF91FFCCFB93FF43FB07FF6E" box="[1068,1192,158,184]" country="Thailand" name="Narathiwat" pageId="7" pageNumber="246">Narathiwat</collectingRegion>
and
<collectingRegion id="547FF841FF91FFCCFB5DFF43FABBFF6E" box="[1250,1300,158,184]" country="Thailand" name="Yala" pageId="7" pageNumber="246">Yala</collectingRegion>
Provinces of southern Peninsular
<collectingCountry id="EEAC7633FF91FFCCFED3FF1FFE7CFF0A" box="[364,467,194,220]" name="Thailand" pageId="7" pageNumber="246">Thailand</collectingCountry>
. (
<figureCitation id="0E802A26FF91FFCCFE59FF1FFD85FF0B" box="[486,554,194,221]" captionStart="FIGURE 7" captionStartId="10.[136,229,1555,1577]" captionTargetBox="[178,1417,221,1505]" captionTargetId="figure-21@10.[151,1436,190,1531]" captionTargetPageId="10" captionText="FIGURE 7. A. &amp; B. Maps showing distributions of Polyalthia heliopetala (●), P. taweensis (▲) and P. pumila (■) in southern Peninsular Thailand. Map in B is an enlargement of the area shown in the rectangle in A. Maps by Pasakorn Bunchalee." figureDoi="http://doi.org/10.5281/zenodo.5426436" httpUri="https://zenodo.org/record/5426436/files/figure.png" pageId="7" pageNumber="246">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF91FFCCFF02FF3BFC13FED6" box="[189,956,230,256]" pageId="7" pageNumber="246" type="biology_ecology">
<paragraph id="960436A3FF91FFCCFF02FF3BFC13FED6" blockId="7.[136,1452,158,869]" box="[189,956,230,256]" pageId="7" pageNumber="246">
<emphasis id="A4CFEAB1FF91FFCCFF02FF3BFEECFED6" bold="true" box="[189,323,230,256]" pageId="7" pageNumber="246">Phenology:</emphasis>
—Flowering in February and April, fruiting in February.
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF91FFCCFF02FED7FDFCFEF2" box="[189,595,266,293]" pageId="7" pageNumber="246" type="vernacular_names">
<paragraph id="960436A3FF91FFCCFF02FED7FDFCFEF2" blockId="7.[136,1452,158,869]" box="[189,595,266,293]" pageId="7" pageNumber="246">
<emphasis id="A4CFEAB1FF91FFCCFF02FED7FEFDFEF2" bold="true" box="[189,338,266,292]" pageId="7" pageNumber="246">Local name:</emphasis>
—Thai: Nara morakot.
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF91FFCCFF02FEF3FEA6FEBA" pageId="7" pageNumber="246" type="etymology">
<paragraph id="960436A3FF91FFCCFF02FEF3FEA6FEBA" blockId="7.[136,1452,158,869]" pageId="7" pageNumber="246">
<emphasis id="A4CFEAB1FF91FFCCFF02FEF3FEE7FE9E" bold="true" box="[189,328,302,328]" pageId="7" pageNumber="246">Etymology:</emphasis>
—The specific epithet refers to the locality, Ta We Mountain, where the species was first discovered.
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF91FFCCFF02FEABFADBFE62" pageId="7" pageNumber="246" type="biology_ecology">
<paragraph id="960436A3FF91FFCCFF02FEABFADBFE62" blockId="7.[136,1452,158,869]" pageId="7" pageNumber="246">
<emphasis id="A4CFEAB1FF91FFCCFF02FEABFDADFE46" bold="true" box="[189,514,374,400]" pageId="7" pageNumber="246">IUCN Conservation status:</emphasis>
—The species has been collected from only three localities. There is no information regarding population size at any of the localities. Therefore, the species is considered to be Data Deficient (DD).
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF91FFCCFF02FE63FED1FDBE" pageId="7" pageNumber="246" type="materials_examined">
<paragraph id="960436A3FF91FFCCFF02FE63FED1FDBE" blockId="7.[136,1452,158,869]" pageId="7" pageNumber="246">
<emphasis id="A4CFEAB1FF91FFCCFF02FE63FD91FE0E" bold="true" box="[189,574,446,472]" pageId="7" pageNumber="246">Additional specimens examined:</emphasis>
<materialsCitation id="26D33CFEFF91FFCCFDE5FE63FCB6FDF6" collectingDate="2004-02-10" collectionCode="A, BKF" collectorName="Middleton &amp; Phuphat &amp; Pooma &amp; Williams" country="Thailand" elevation="250" latitude="6.0833335" location="Ban Chulapon Phattana" longLatPrecision="1305" longitude="101.38333" municipality="Than To" pageId="7" pageNumber="246" specimenCode="L.3729224, L.3729128" specimenCount="1" stateProvince="Yala">
<collectingCountry id="EEAC7633FF91FFCCFDE5FE63FD41FE0E" box="[602,750,446,472]" name="Thailand" pageId="7" pageNumber="246">THAILAND</collectingCountry>
. PENINSULAR:
<collectingRegion id="547FF841FF91FFCCFC0EFE63FC49FE0E" box="[945,998,446,472]" country="Thailand" name="Yala" pageId="7" pageNumber="246">Yala</collectingRegion>
,
<collectingMunicipality id="7660ACD9FF91FFCCFC50FE63FBE1FE0F" box="[1007,1102,446,473]" pageId="7" pageNumber="246">Than To</collectingMunicipality>
,
<location id="93646078FF91FFCCFBE8FE62FAF3FE0E" LSID="urn:lsid:plazi:treatment:1E1287B5FF92FFCCFF37FCF1FAE6FCB2:93646078FF91FFCCFBE8FE62FAF3FE0E" box="[1111,1372,446,473]" country="Thailand" latitude="6.0833335" longLatPrecision="1305" longitude="101.38333" municipality="Than To" name="Ban Chulapon Phattana" pageId="7" pageNumber="246" stateProvince="Yala">Ban Chulapon Phattana</location>
7 area, trail from substation up along ridge,
<geoCoordinate id="F38F5064FF91FFCCFDA0FE3FFD2EFE2B" box="[543,641,482,509]" degrees="6" direction="north" minutes="05" orientation="latitude" pageId="7" pageNumber="246" precision="925" value="6.0833335">6° 05´N</geoCoordinate>
,
<geoCoordinate id="F38F5064FF91FFCCFD33FE3FFCA9FE2B" box="[652,774,482,509]" degrees="101" direction="east" minutes="23" orientation="longitude" pageId="7" pageNumber="246" precision="925" value="101.38333">101° 23´E</geoCoordinate>
, alt.
<quantity id="51439B46FF91FFCCFC84FE3FFC2BFE2B" box="[827,900,482,509]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="246" unit="m" value="250.0">
<elevation id="1D96D190FF91FFCCFC84FE3FFC2BFE2B" box="[827,900,482,509]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="246" unit="m" value="250.0">250 m</elevation>
</quantity>
,
<date id="E2051063FF91FFCCFC30FE3FFBFAFE2A" box="[911,1109,482,508]" pageId="7" pageNumber="246" value="2004-02-10">
<collectingDate id="F241E98BFF91FFCCFC30FE3FFBFAFE2A" box="[911,1109,482,508]" pageId="7" pageNumber="246" value="2004-02-10">10 February 2004</collectingDate>
</date>
(flower),
<emphasis id="A4CFEAB1FF91FFCCFB7AFE3FFE36FDF6" italics="true" pageId="7" pageNumber="246">
<collectorName id="3B4E5375FF91FFCCFB7AFE3FFA94FE2A" box="[1221,1339,482,508]" pageId="7" pageNumber="246">Middleton</collectorName>
,
<collectorName id="3B4E5375FF91FFCCFAFAFE3FFA08FE2A" box="[1349,1447,482,508]" pageId="7" pageNumber="246">Phuphat</collectorName>
,
<collectorName id="3B4E5375FF91FFCCFF37FDDAFF78FDF6" box="[136,215,519,544]" pageId="7" pageNumber="246">Pooma</collectorName>
&amp;
<collectorName id="3B4E5375FF91FFCCFF46FDDBFEF4FDF6" box="[249,347,518,544]" pageId="7" pageNumber="246">Williams</collectorName>
2900
</emphasis>
(
<collectionCode id="F0AAAE66FF91FFCCFE17FDDBFE10FDF6" box="[424,447,518,544]" pageId="7" pageNumber="246">A</collectionCode>
,
<collectionCode id="F0AAAE66FF91FFCCFE75FDDAFDAEFDF7" box="[458,513,519,545]" pageId="7" pageNumber="246">BKF</collectionCode>
,
<specimenCode id="C61D9ED8FF91FFCCFDB4FDDBFD27FDF6" box="[523,648,518,544]" pageId="7" pageNumber="246">L.3729224</specimenCode>
,
<specimenCode id="C61D9ED8FF91FFCCFD2DFDDBFCBEFDF6" box="[658,785,518,544]" pageId="7" pageNumber="246">L.3729128</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="26D33CFEFF91FFCCFC9DFDDBFED5FDBE" collectingDate="2004-02-14" collectionCode="A" collectorName="Middleton &amp; Phuphat &amp; Pooma &amp; Williams" elevation="120" location="Hala Bala Wildlife Sanctuary" municipality="Than To" pageId="7" pageNumber="246" specimenCode="L.3729130" specimenCount="1" stateProvince="Yala">
<collectingRegion id="547FF841FF91FFCCFC9DFDDBFCF8FDF6" box="[802,855,518,544]" country="Thailand" name="Yala" pageId="7" pageNumber="246">Yala</collectingRegion>
,
<collectingMunicipality id="7660ACD9FF91FFCCFCDEFDDBFC6FFDF7" box="[865,960,518,545]" pageId="7" pageNumber="246">Than To</collectingMunicipality>
,
<location id="93646078FF91FFCCFC75FDDBFABEFDF6" LSID="urn:lsid:plazi:treatment:1E1287B5FF92FFCCFF37FCF1FAE6FCB2:93646078FF91FFCCFC75FDDBFABEFDF6" box="[970,1297,518,544]" municipality="Than To" name="Hala Bala Wildlife Sanctuary" pageId="7" pageNumber="246" stateProvince="Yala">Hala Bala Wildlife Sanctuary</location>
,
<location id="93646078FF91FFCCFAA4FDDBFF01FD92" LSID="urn:lsid:plazi:treatment:1E1287B5FF92FFCCFF37FCF1FAE6FCB2:93646078FF91FFCCFAA4FDDBFF01FD92" municipality="Than To" name="Khlong Nam Sai" pageId="7" pageNumber="246" stateProvince="Yala">Khlong Nam Sai</location>
, 6° 01´70˝ N, 101° 24´E, alt.
<quantity id="51439B46FF91FFCCFDB8FDF7FDFFFD93" box="[519,592,554,581]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="246" unit="m" value="120.0">
<elevation id="1D96D190FF91FFCCFDB8FDF7FDFFFD93" box="[519,592,554,581]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="246" unit="m" value="120.0">120 m</elevation>
</quantity>
,
<date id="E2051063FF91FFCCFDE3FDF7FC8CFD92" box="[604,803,554,580]" pageId="7" pageNumber="246" value="2004-02-14">
<collectingDate id="F241E98BFF91FFCCFDE3FDF7FC8CFD92" box="[604,803,554,580]" pageId="7" pageNumber="246" value="2004-02-14">14 February 2004</collectingDate>
</date>
(flower &amp; fruit),
<emphasis id="A4CFEAB1FF91FFCCFC57FDF7FF6FFDBE" italics="true" pageId="7" pageNumber="246">
<collectorName id="3B4E5375FF91FFCCFC57FDF7FBF1FD92" box="[1000,1118,554,580]" pageId="7" pageNumber="246">Middleton</collectorName>
,
<collectorName id="3B4E5375FF91FFCCFBD6FDF7FB64FD92" box="[1129,1227,554,580]" pageId="7" pageNumber="246">Phuphat</collectorName>
,
<collectorName id="3B4E5375FF91FFCCFB69FDF6FA8AFD92" box="[1238,1317,555,580]" pageId="7" pageNumber="246">Pooma</collectorName>
&amp;
<collectorName id="3B4E5375FF91FFCCFAF6FDF7FA04FD92" box="[1353,1451,554,580]" pageId="7" pageNumber="246">Williams</collectorName>
3083
</emphasis>
(
<collectionCode id="F0AAAE66FF91FFCCFF6FFD93FF48FDBE" box="[208,231,590,616]" pageId="7" pageNumber="246">A</collectionCode>
,
<specimenCode id="C61D9ED8FF91FFCCFF4CFD93FEDDFDBE" box="[243,370,590,616]" pageId="7" pageNumber="246">L.3729130</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF91FFCCFF02FDAFFAE6FCB2" pageId="7" pageNumber="246" type="discussion">
<paragraph id="960436A3FF91FFCCFF02FDAFFAE6FCB2" blockId="7.[136,1452,158,869]" pageId="7" pageNumber="246">
<emphasis id="A4CFEAB1FF91FFCCFF02FDAFFEA6FD5A" bold="true" box="[189,265,626,652]" pageId="7" pageNumber="246">Notes:</emphasis>
—Comparative data on
<taxonomicName id="51BB4D20FF91FFCCFDA2FDAEFD3EFD5A" box="[541,657,626,652]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="7" pageNumber="246" phylum="Tracheophyta" rank="species" species="obliqua">
<emphasis id="A4CFEAB1FF91FFCCFDA2FDAEFD3EFD5A" box="[541,657,626,652]" italics="true" pageId="7" pageNumber="246">P. obliqua</emphasis>
</taxonomicName>
used in the diagnosis were taken from
<bibRefCitation id="F22A4B52FF91FFCCFBE6FDAFFAA7FD5B" author="Sinclair, J." box="[1113,1288,626,653]" pageId="7" pageNumber="246" pagination="149 - 516" refId="ref5266" refString="Sinclair, J. (1955) A revision of the Malayan Annonaceae. Gardens' Bulletin, Singapore 14: 149 - 516." type="journal article" year="1955">Sinclair (1955)</bibRefCitation>
. Inflorescence position in
<taxonomicName id="51BB4D20FF91FFCCFEBEFD4BFE24FD66" box="[257,395,662,688]" class="Magnoliopsida" family="Annonaceae" kingdom="Plantae" order="Magnoliales" pageId="7" pageNumber="246" phylum="Tracheophyta" rank="family">Annonaceae</taxonomicName>
is usually a conservative feature indicating distinction at the generic level or a marker for groups of related species within a genus, so it is surprising to have this distinction between two otherwise similar species.
<bibRefCitation id="F22A4B52FF91FFCCFF37FD02FE89FD2E" author="Turner, I. M." box="[136,294,734,761]" pageId="7" pageNumber="246" pagination="77 - 98" refId="ref5290" refString="Turner, I. M. (2008) New species and new records for Polyalthia (Annonaceae) in Borneo. Folia Malaysiana 9: 77 - 98." type="journal article" year="2008">Turner (2008)</bibRefCitation>
documented the same difference, however, between
<taxonomicName id="51BB4D20FF91FFCCFCC3FD03FA04FD2E" authority="(Blume) Blume (1830: 71)" authorityName="Blume" authorityPageNumber="71" authorityYear="1830" baseAuthorityName="Blume" box="[892,1451,734,761]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="7" pageNumber="246" phylum="Tracheophyta" rank="species" species="subcordata">
<emphasis id="A4CFEAB1FF91FFCCFCC3FD03FBD9FD2E" box="[892,1142,734,760]" italics="true" pageId="7" pageNumber="246">Polyalthia subcordata</emphasis>
(Blume)
<bibRefCitation id="F22A4B52FF91FFCCFB5CFD03FA04FD2E" author="Blume, C. L." box="[1251,1451,734,760]" pageId="7" pageNumber="246" pagination="1 - 108" refId="ref4930" refString="Blume, C. L. (1830) Flora Javae, volume 2: Anonaceae. Frank, Brussels, pp. 1 - 108 + 53 plates. https: // doi. org / 10.5962 / bhl. title. 48445" type="book chapter" year="1830">Blume (1830: 71)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="51BB4D20FF91FFCCFF07FCDFFD68FCCA" authority="(Blume) Blume (1830: 73)" authorityName="Blume" authorityPageNumber="73" authorityYear="1830" baseAuthorityName="Blume" box="[184,711,770,797]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="7" pageNumber="246" phylum="Tracheophyta" rank="species" species="elliptica">
<emphasis id="A4CFEAB1FF91FFCCFF07FCDFFE3FFCCA" box="[184,400,770,796]" italics="true" pageId="7" pageNumber="246">Polyalthia elliptica</emphasis>
(Blume)
<bibRefCitation id="F22A4B52FF91FFCCFE43FCDFFD68FCCA" author="Blume, C. L." box="[508,711,770,796]" pageId="7" pageNumber="246" pagination="1 - 108" refId="ref4930" refString="Blume, C. L. (1830) Flora Javae, volume 2: Anonaceae. Frank, Brussels, pp. 1 - 108 + 53 plates. https: // doi. org / 10.5962 / bhl. title. 48445" type="book chapter" year="1830">Blume (1830: 73)</bibRefCitation>
</taxonomicName>
, so inflorescence position appears to be a more variable character within
<taxonomicName id="51BB4D20FF91FFCCFF68FCFBFEE3FC96" authorityName="Blume" authorityYear="1830" box="[215,332,806,832]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="7" pageNumber="246" phylum="Tracheophyta" rank="genus">
<emphasis id="A4CFEAB1FF91FFCCFF68FCFBFEE3FC96" box="[215,332,806,832]" italics="true" pageId="7" pageNumber="246">Polyalthia</emphasis>
</taxonomicName>
than in other
<taxonomicName id="51BB4D20FF91FFCCFE56FCFBFDDCFC96" box="[489,627,806,832]" class="Magnoliopsida" family="Annonaceae" kingdom="Plantae" order="Magnoliales" pageId="7" pageNumber="246" phylum="Tracheophyta" rank="family">Annonaceae</taxonomicName>
genera. The differentiated border of the outer petals in
<taxonomicName id="51BB4D20FF91FFCCFB59FCFAFAC0FC96" box="[1254,1391,807,832]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="7" pageNumber="246" phylum="Tracheophyta" rank="species" species="taweensis">
<emphasis id="A4CFEAB1FF91FFCCFB59FCFAFAC0FC96" box="[1254,1391,807,832]" italics="true" pageId="7" pageNumber="246">P. taweensis</emphasis>
</taxonomicName>
is an unusual feature; in other
<taxonomicName id="51BB4D20FF91FFCCFE21FC97FD87FCB2" box="[414,552,842,868]" class="Magnoliopsida" family="Annonaceae" kingdom="Plantae" order="Magnoliales" pageId="7" pageNumber="246" phylum="Tracheophyta" rank="family">Annonaceae</taxonomicName>
such modifications occur only on the inner petals (
<bibRefCitation id="F22A4B52FF91FFCCFBD9FC97FA93FCB2" author="Van Heusden, E. C. H." box="[1126,1340,842,868]" pageId="7" pageNumber="246" pagination="1 - 218" refId="ref5482" refString="Van Heusden, E. C. H. (1992) Flowers of Annonaceae: morphology, classification, and evolution. Blumea Supplement 7: 1 - 218." type="journal article" year="1992">van Heusden 1992</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,343 @@
<document id="1E3D42D3C34BCC391C8DB7AA2FB8E3A3" ID-DOI="10.11646/phytotaxa.510.3.4" ID-ISSN="1179-3163" ID-Zenodo-Dep="5426421" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="operationResults" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1630397611626" checkinUser="marcus" docAuthor="Bunchalee, Pasakorn, Leeratiwong, Charan &amp; Johnson, David M." docDate="2021" docId="1E1287B5FF97FFCAFF37FF29FF46F87C" docLanguage="en" docName="phytotaxa.510.3.4.pdf" docOrigin="Phytotaxa 510 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.510.3.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Polyalthia heliopetala Leerat. &amp; Bunchalee 2021, sp. nov." docType="treatment" docVersion="1" lastPageNumber="240" masterDocId="E22BFFCDFF96FFCBFFBFFFDDFFAFFFD6" masterDocTitle="Two new species and a new record of the genus Polyalthia (Annonaceae) from Peninsular Thailand" masterLastPageNumber="250" masterPageNumber="239" pageNumber="240" updateTime="1732197805484" updateUser="guilherme" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="1AA24A92156D01652C762080F6C6DCB9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="2733BB02C0BD149C12C9C50B33680A92">Two new species and a new record of the genus Polyalthia (Annonaceae) from Peninsular Thailand</mods:title>
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<mods:name id="5BC23E3FABE56A8F6EDD8B678F1DEF7E" type="personal">
<mods:role id="A4783150A82E6288C681FE010DA45F3E">
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<mods:namePart id="0C1F80502C2BA03FB3626D7D8DEB282D">Bunchalee, Pasakorn</mods:namePart>
<mods:nameIdentifier id="8A17A26A5FDBC79C1858A61246B813C2" type="ORCID">0000-0001-8644-3360</mods:nameIdentifier>
<mods:affiliation id="F7B3BF24EFD53FBE0F3DAA7C6BB34AF8">Department of Biology, Faculty of Science, Mahasarakham University, Kantarawichai, Maha Sarakham, 44150, Thailand. Palaeontological Research and Education Center, Mahasarakham University, Kantarawichai, Maha Sarakham, 44150, Thailand. Division of Biological Science, Faculty of Science, Prince of Songkla University, Hat Yai, Songkhla 90112, Thailand. &amp; pasakorn. b @ msu. ac. th; https: // orcid. org / 0000 - 0001 - 8644 - 3360</mods:affiliation>
<mods:nameIdentifier id="C07477CEA2EE53401679CCE320842171" type="email">pasakorn.b@msu.ac.th</mods:nameIdentifier>
</mods:name>
<mods:name id="3CEB5F55A61587D3BFD53C53DDBEBF28" type="personal">
<mods:role id="DF794510C15F1BA6209DA06F594A8B54">
<mods:roleTerm id="01823E228BD8BC5DAC75908B45BD498E">Author</mods:roleTerm>
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<mods:namePart id="B7A4780516F12E539673CF74EB6B2F72">Leeratiwong, Charan</mods:namePart>
<mods:nameIdentifier id="AF8FE601206713DDF456703D5865063A" type="ORCID">0000-0002-0752-4275</mods:nameIdentifier>
<mods:affiliation id="FE7A8A9EF81065C9C7E46BC4A6C5A10C">charan. leeratiwong @ gmail. com; https: // orcid. org / 0000 - 0002 - 0752 - 4275</mods:affiliation>
<mods:nameIdentifier id="F3647556ECE9775EC2C7B9DBA2DB8DF7" type="email">charan.leeratiwong@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="748F8C1943439179117AB42FED34ECCD" type="personal">
<mods:role id="2BF0DF5F6C0549B0BA1464718E89C469">
<mods:roleTerm id="CA8096E67342BB82240C6256D5FD0407">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F6A17C5062AFB2F82E391159445A9BB5">Johnson, David M.</mods:namePart>
<mods:nameIdentifier id="0E05C83EA2B1A950D0B1617C3915BC0E" type="ORCID">0000-0003-2896-7419</mods:nameIdentifier>
<mods:affiliation id="BFD31DA53B0E2D8FA84FB15D10555AC3">Department of Botany and Microbiology, Ohio Wesleyan University, Delaware, Ohio, 43015, USA &amp; dmjohnso @ owu. edu; https: // orcid. org / 0000 - 0003 - 2896 - 7419</mods:affiliation>
<mods:nameIdentifier id="A7F38E6F327CF406F1F2134A634F8EA0" type="email">dmjohnso@owu.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="80B81BAC3291513B6BC90258269E3311">text</mods:typeOfResource>
<mods:relatedItem id="345462DF03DB55BD4B43877F74778860" type="host">
<mods:titleInfo id="13CE11396C1E19BDF668EA46EADD331A">
<mods:title id="BD7244BC76F7089C77C04B9BA83E126B">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="C8D33A9D377025A9B400F4C11B05E253">
<mods:date id="22F789699F3754C2412EB77F0B1E83FF">2021</mods:date>
<mods:detail id="213CACBE1E8FAAA05EF477FD7E8013BA" type="pubDate">
<mods:number id="B602D7965E28A9FCF840DBF95C16A930">2021-07-09</mods:number>
</mods:detail>
<mods:detail id="DE37D9778A701BD0C6832635FAC3BBFB" type="volume">
<mods:number id="534B62135615E25EA71D78511C4AABBA">510</mods:number>
</mods:detail>
<mods:detail id="C92A161B0FBDBFD5F3CD10832E43D826" type="issue">
<mods:number id="8443B755E53A8AA3372F9BBB0D8F0AE2">3</mods:number>
</mods:detail>
<mods:extent id="DFB91939B4B7B2E3E6A6603DF2CCB1FD" unit="page">
<mods:start id="703FBAAB9A9F4D66E333D7C7A0D38F6A">239</mods:start>
<mods:end id="CC4FA48AC147124C63244BB3683E0ED2">250</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="A6A19C2ED595DE318E8D28C3156F689C">
<mods:url id="D42CD019EA2C4E5688FC0B7368B015FC">http://dx.doi.org/10.11646/phytotaxa.510.3.4</mods:url>
</mods:location>
<mods:classification id="0B09E45E80DA015BDAE2F9104A54DF69">journal article</mods:classification>
<mods:identifier id="A3F9AB9CE84FDF642D95D4EF86F4FBDC" type="DOI">10.11646/phytotaxa.510.3.4</mods:identifier>
<mods:identifier id="6C86E87A1A5D8DC014F1E46C822B4C9A" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="A3D92B07CD3DDB7731ED1FA9F880BE7F" type="Zenodo-Dep">5426421</mods:identifier>
</mods:mods>
<treatment id="1E1287B5FF97FFCAFF37FF29FF46F87C" LSID="urn:lsid:plazi:treatment:1E1287B5FF97FFCAFF37FF29FF46F87C" httpUri="http://treatment.plazi.org/id/1E1287B5FF97FFCAFF37FF29FF46F87C" lastPageNumber="240" pageId="1" pageNumber="240">
<subSubSection id="DEA16528FF97FFCAFF37FF29FCDAFED8" box="[136,885,244,271]" pageId="1" pageNumber="240" type="nomenclature">
<paragraph id="960436A3FF97FFCAFF37FF29FCDAFED8" blockId="1.[136,885,244,271]" box="[136,885,244,271]" pageId="1" pageNumber="240">
<taxonomicName id="51BB4D20FF97FFCAFF37FF29FDD2FED8" ID-CoL="C832T" authority="Leerat. &amp; Bunchalee" authorityName="Leerat. &amp; Bunchalee" authorityYear="2021" box="[136,637,244,271]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="240" phylum="Tracheophyta" rank="species" species="heliopetala" status="sp. nov.">
<emphasis id="A4CFEAB1FF97FFCAFF37FF29FE24FED8" bold="true" box="[136,395,244,270]" pageId="1" pageNumber="240">Polyalthia heliopetala</emphasis>
Leerat. &amp; Bunchalee
</taxonomicName>
,
<emphasis id="A4CFEAB1FF97FFCAFD37FF28FD72FED8" box="[648,733,245,270]" italics="true" pageId="1" pageNumber="240">
<taxonomicNameLabel id="BFFC57CAFF97FFCAFD37FF28FD72FED8" box="[648,733,245,270]" pageId="1" pageNumber="240" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="0E802A26FF97FFCAFD53FF29FC96FED8" box="[748,825,244,270]" captionStart="FIGURE 1" captionStartId="2.[136,229,1916,1938]" captionTargetBox="[199,1380,218,1873]" captionTargetId="figure-21@2.[178,1409,203,1890]" captionTargetPageId="2" captionText="FIGURE 1. Polyalthia heliopetala: A. Habit; B. Leaf bases (upper side) and inflorescences, with point of inflorescence attachments to stem indicated by arrows; C. Flower showing recurved petals; D. Flower, top view showing the embossed patch on the inner base of the petals; E. Outer surfaces of outer petals; F. Outer surfaces of inner petals; G. Stamens and persistent sepal on pedicel; H. Torus with some stamens removed to show attachment of stamens and carpels on the convex torus surface; I. Carpel; J. Stamens (outside on left, inside on right); K. Monocarps. The illustration was drawn by Nattapong Chomlok from Leeratiwong 20-1527 (BKF)." figureDoi="http://doi.org/10.5281/zenodo.5426423" httpUri="https://zenodo.org/record/5426423/files/figure.png" pageId="1" pageNumber="240">Figs. 1</figureCitation>
&amp;
<figureCitation id="0E802A26FF97FFCAFCE2FF29FCC3FED8" box="[861,876,244,270]" captionStart="FIGURE 2" captionStartId="3.[136,229,1920,1942]" captionTargetBox="[179,1406,197,1886]" captionTargetId="figure-18@3.[164,1423,187,1898]" captionTargetPageId="3" captionText="FIGURE 2. Polyalthia heliopetala: A. Leaves and inflorescences; B. Lower surface of leaf base and flower; C. Flower showing the embossed basal patch of the inner surface of the petals; D. Upper surface of leaves and flower; E. Lower surface of leaf showing venation; F. Monocarps.A, B, D and E are from Leeratiwong 20-1527(BKF); F is from Puudjaa 626 (BKF)]; all photographs by Pasakorn Bunchalee. Photograph in C was taken by Yamilah Sama-ae in Narathiwat, Ra Ngae District, Bo Ngo Subdistrict, Bue Chaeng Village, 7 April 2020." figureDoi="http://doi.org/10.5281/zenodo.5426425" httpUri="https://zenodo.org/record/5426425/files/figure.png" pageId="1" pageNumber="240">2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF97FFCAFF37FE9DFCE5FE14" pageId="1" pageNumber="240" type="reference_group">
<paragraph id="960436A3FF97FFCAFF37FE9DFCE5FE14" blockId="1.[136,1453,320,522]" pageId="1" pageNumber="240">
<taxonomicName id="51BB4D20FF97FFCAFF37FE9DFECEFE83" authorityName="Leerat. &amp; Bunchalee" authorityYear="2021" box="[136,353,320,341]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="240" phylum="Tracheophyta" rank="species" species="heliopetala">
<emphasis id="A4CFEAB1FF97FFCAFF37FE9DFECEFE83" box="[136,353,320,341]" italics="true" pageId="1" pageNumber="240">Polyalthia heliopetala</emphasis>
</taxonomicName>
resembles
<taxonomicName id="51BB4D20FF97FFCAFE6CFE9DFD49FE80" authority="King (1892: 65)" authorityName=", King" authorityPageNumber="65" authorityYear="1892" box="[467,742,320,342]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="240" phylum="Tracheophyta" rank="species" species="oblonga">
<emphasis id="A4CFEAB1FF97FFCAFE6CFE9DFD92FE83" box="[467,573,320,341]" italics="true" pageId="1" pageNumber="240">P. oblonga</emphasis>
<bibRefCitation id="F22A4B52FF97FFCAFDFAFE9DFD49FE80" author="King, G." box="[581,742,320,342]" pageId="1" pageNumber="240" pagination="1 - 130" refId="ref5104" refString="King, G. (1892) Materials for a flora of the Malay Peninsula, no. 4. Journal of the Asiatic Society of Bengal, Part 2, Natural History 61: 1 - 130." type="journal article" year="1892">King (1892: 65)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="51BB4D20FF97FFCAFCA4FE9DFC36FE83" authorityName=", Turner" authorityYear="2014" box="[795,921,320,341]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="240" phylum="Tracheophyta" rank="species" species="motleyana">
<emphasis id="A4CFEAB1FF97FFCAFCA4FE9DFC36FE83" box="[795,921,320,341]" italics="true" pageId="1" pageNumber="240">P. motleyana</emphasis>
</taxonomicName>
(Hook.f.)
<bibRefCitation id="F22A4B52FF97FFCAFBBCFE9DFB44FE80" author="Airy Shaw, H. K." box="[1027,1259,320,342]" pageId="1" pageNumber="240" pagination="275 - 290" refId="ref4875" refString="Airy Shaw, H. K. (1939) XXXII - Additions to the flora of Borneo and other Malay islands: XII. The Annonaceae of the Oxford University Expedition to Sarawak, 1932. Kew Bulletin 1939: 275 - 290. https: // doi. org / 10.2307 / 4111738" type="journal article" year="1939">Airy Shaw (1939: 280)</bibRefCitation>
but differs from
<taxonomicName id="51BB4D20FF97FFCAFA25FE9DFEA3FEAF" authorityName=", King" authorityYear="1892" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="240" phylum="Tracheophyta" rank="species" species="oblonga">
<emphasis id="A4CFEAB1FF97FFCAFA25FE9DFEA3FEAF" italics="true" pageId="1" pageNumber="240">P. oblonga</emphasis>
</taxonomicName>
in the flower pedicels 7.010.0 versus 31.057.0 mm long and lanceolate versus oblong inner petals and from
<taxonomicName id="51BB4D20FF97FFCAFA8FFEB9FA04FEAF" authorityName=", Turner" authorityYear="2014" box="[1328,1451,356,377]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="240" phylum="Tracheophyta" rank="species" species="motleyana">
<emphasis id="A4CFEAB1FF97FFCAFA8FFEB9FA04FEAF" box="[1328,1451,356,377]" italics="true" pageId="1" pageNumber="240">P. motleyana</emphasis>
</taxonomicName>
in the oblong to lanceolate rather than ovate petals and the monocarp stipes 18.025.0 versus ca. 10.0 mm long. It differs from both species in having embossed patches on the inner bases of the petals.
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF97FFCAFF37FE0DFDE8FDDC" pageId="1" pageNumber="240" type="materials_examined">
<paragraph id="960436A3FF97FFCAFF37FE0DFDE8FDDC" blockId="1.[136,1453,320,522]" pageId="1" pageNumber="240">
<collectingCounty id="7F654E2FFF97FFCAFF37FE0DFF15FE33" box="[136,186,464,485]" pageId="1" pageNumber="240">Type</collectingCounty>
:—
<materialsCitation id="26D33CFEFF97FFCAFF69FE0DFE02FDDF" collectingDate="2020-04-07" collectionCode="PSU" collectorName="Leeratiwong" country="Thailand" location="Bo Ngo Subdistrict" municipality="Ra Ngae District" pageId="1" pageNumber="240" specimenCount="1" stateProvince="Narathiwat" typeStatus="holotype">
<collectingCountry id="EEAC7633FF97FFCAFF69FE0DFEF9FE33" box="[214,342,464,485]" name="Thailand" pageId="1" pageNumber="240">THAILAND</collectingCountry>
.
<collectingRegion id="547FF841FF97FFCAFEE1FE0DFE64FE30" box="[350,459,464,486]" country="Thailand" name="Narathiwat" pageId="1" pageNumber="240">Narathiwat</collectingRegion>
,
<collectingMunicipality id="7660ACD9FF97FFCAFE6BFE0DFDD7FE30" box="[468,632,464,486]" pageId="1" pageNumber="240">Ra Ngae District</collectingMunicipality>
,
<location id="93646078FF97FFCAFD3EFE0DFC92FE30" LSID="urn:lsid:plazi:treatment:1E1287B5FF97FFCAFF37FF29FF46F87C:93646078FF97FFCAFD3EFE0DFC92FE30" box="[641,829,464,486]" country="Thailand" county="Type" municipality="Ra Ngae District" name="Bo Ngo Subdistrict" pageId="1" pageNumber="240" stateProvince="Narathiwat">Bo Ngo Subdistrict</location>
,
<location id="93646078FF97FFCAFCF9FE0DFBA6FE30" LSID="urn:lsid:plazi:treatment:1E1287B5FF97FFCAFF37FF29FF46F87C:93646078FF97FFCAFCF9FE0DFBA6FE30" box="[838,1033,464,486]" country="Thailand" county="Type" municipality="Ra Ngae District" name="Bue Chaeng Village" pageId="1" pageNumber="240" stateProvince="Narathiwat">Bue Chaeng Village</location>
,
<date id="E2051063FF97FFCAFBADFE0DFB25FE33" box="[1042,1162,464,486]" pageId="1" pageNumber="240" value="2020-04-07">
<collectingDate id="F241E98BFF97FFCAFBADFE0DFB25FE33" box="[1042,1162,464,486]" pageId="1" pageNumber="240" value="2020-04-07">7 April 2020</collectingDate>
</date>
(flower &amp; fruit),
<emphasis id="A4CFEAB1FF97FFCAFA8DFE0DFEA1FDDF" italics="true" pageId="1" pageNumber="240">
<collectorName id="3B4E5375FF97FFCAFA8DFE0DFA04FE33" box="[1330,1451,464,485]" pageId="1" pageNumber="240">Leeratiwong</collectorName>
20-1527
</emphasis>
(
<typeStatus id="49008801FF97FFCAFEA5FE29FEDFFDDC" box="[282,368,500,522]" pageId="1" pageNumber="240" type="holotype">holotype</typeStatus>
<collectionCode id="F0AAAE66FF97FFCAFECAFE29FE06FDDF" box="[373,425,500,521]" pageId="1" pageNumber="240">PSU</collectionCode>
!
</materialsCitation>
;
<materialsCitation id="26D33CFEFF97FFCAFE09FE29FD93FDDC" box="[438,572,500,522]" collectingDate="2020-04-07" collectionCode="BKF" collectorName="Leeratiwong" country="Thailand" location="Bo Ngo Subdistrict" municipality="Ra Ngae District" pageId="1" pageNumber="240" specimenCount="1" stateProvince="Narathiwat" typeStatus="isotype">
<typeStatus id="49008801FF97FFCAFE09FE29FE54FDDC" box="[438,507,500,522]" pageId="1" pageNumber="240">isotype</typeStatus>
<collectionCode id="F0AAAE66FF97FFCAFDBEFE29FD97FDDC" box="[513,568,500,522]" collectionName="BKF" pageId="1" pageNumber="240">BKF</collectionCode>
!
</materialsCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF97FFCAFF37FDE5FF43FA91" pageId="1" pageNumber="240" type="description">
<paragraph id="960436A3FF97FFCAFF37FDE5FF43FA91" blockId="1.[136,1452,568,1962]" pageId="1" pageNumber="240">
Shrubs 2.04.0 m tall, bark smooth, greyish black, primary branching spiral.
<emphasis id="A4CFEAB1FF97FFCAFC58FDE4FB34FD84" box="[999,1179,568,594]" italics="true" pageId="1" pageNumber="240">Young branches</emphasis>
rusty-brown tomentose, the hairs erect, 0.71.0 mm long, with moderately prominent lenticels.
<emphasis id="A4CFEAB1FF97FFCAFC11FD80FC53FDA0" box="[942,1020,605,630]" italics="true" pageId="1" pageNumber="240">Leaves</emphasis>
with petioles stout, 2.03.0 mm long, 3.0
<quantity id="51439B46FF97FFCAFF05FD5DFEA1FD4D" box="[186,270,640,667]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="240" unit="mm" value="3.5">3.5 mm</quantity>
thick, flat above, rustybrown tomentose; lamina chartaceous, asymmetrically elliptic to oblanceolate and slightly curved inward, 18.032.0 × 5.010.0 cm, base asymmetrically cordate with auricles 1.02.0 mm long, apex cuspidate to acuminate with an acumen 8.010.0 (12.0) mm long, upper side glabrous except for the sparsely tomentose midrib, lower side glabrous except for hairs along midrib and lateral veins; midrib grooved above; lateral veins 1416 per side, slightly arched towards the apex, at midpoint diverging at 6070° from the midrib, festooned loop-forming 1.52.0 mm from the margins; intersecondary veins present; tertiary veins weakly scalariform.
<emphasis id="A4CFEAB1FF97FFCAFB02FCE9FAF1FC98" box="[1213,1374,820,846]" italics="true" pageId="1" pageNumber="240">Inflorescences</emphasis>
1(3)- flowered, supra-axillary or terminal on leafy branches; pedicels 7.010.0 mm long, 1.52.0 mm thick, puberulous; bracts 2, attached at the pedicel base, lanceolate, 1.82.0 × 1.0
<quantity id="51439B46FF97FFCAFCEAFCA1FC02FC41" box="[853,941,892,919]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="240" unit="mm" value="1.2">1.2 mm</quantity>
, apex acute, puberulous outside and glabrous inside.
<emphasis id="A4CFEAB1FF97FFCAFF64FC7DFE8BFC6C" box="[219,292,928,954]" italics="true" pageId="1" pageNumber="240">Sepals</emphasis>
valvate, brownish yellow
<emphasis id="A4CFEAB1FF97FFCAFDEBFC7CFD0EFC6C" box="[596,673,929,954]" italics="true" pageId="1" pageNumber="240">in vivo</emphasis>
, ovate, 5.06.0 × 3.04.0 mm, apex acuminate with acumen ca. 1.0 mm long, puberulous outside and glabrous inside.
<emphasis id="A4CFEAB1FF97FFCAFD05FC19FCAFFC08" box="[698,768,964,990]" italics="true" pageId="1" pageNumber="240">Petals</emphasis>
yellow, orange, or brownish orange
<emphasis id="A4CFEAB1FF97FFCAFB28FC18FB4CFC08" box="[1175,1251,965,990]" italics="true" pageId="1" pageNumber="240">in vivo</emphasis>
, coriaceous, erect or slightly recurved, apex acute, inner surface embossed with a shallowly ovate basal patch (most visible in fresh condition), pubescent outside, glabrous inside; outer petals oblong or lanceolate, 15.018.0 × 5.07.0 mm; inner petals lanceolate, 17.022.0 × 3.05.0 mm.
<emphasis id="A4CFEAB1FF97FFCAFDCBFBEDFD7EFB9C" box="[628,721,1072,1098]" italics="true" pageId="1" pageNumber="240">Stamens</emphasis>
yellow
<emphasis id="A4CFEAB1FF97FFCAFC92FBECFCD6FB9C" box="[813,889,1073,1098]" italics="true" pageId="1" pageNumber="240">in vivo</emphasis>
, slightly cuneate, 2.02.2 ×
<quantity id="51439B46FF97FFCAFB06FBEDFAEDFB9D" box="[1209,1346,1072,1099]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="1" pageNumber="240" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.50.6 mm</quantity>
, filament ca.
<quantity id="51439B46FF97FFCAFF0DFB89FEA5FBB9" box="[178,266,1108,1135]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="240" unit="mm" value="0.3">0.3 mm</quantity>
long; anthers
<quantity id="51439B46FF97FFCAFE0EFB89FD94FBB9" box="[433,571,1108,1135]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="1" pageNumber="240" unit="mm" value="1.7" valueMax="1.8" valueMin="1.6">1.61.8 mm</quantity>
long, anther connective apex truncate; androecium 7.09.0 mm in diameter.
<emphasis id="A4CFEAB1FF97FFCAFF37FBA5FF4EFB44" box="[136,225,1144,1170]" italics="true" pageId="1" pageNumber="240">Carpels</emphasis>
1215 per flower, 2.62.8 ×
<quantity id="51439B46FF97FFCAFD91FBA5FD15FB45" box="[558,698,1144,1171]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="1" pageNumber="240" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.50.6 mm</quantity>
, pubescent, style rudimentary, stigmas obovoid or oblong,
<quantity id="51439B46FF97FFCAFAE8FBA5FF19FB61" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="1" pageNumber="240" unit="mm" value="0.6" valueMax="0.7" valueMin="0.5">0.50.7 mm</quantity>
, pubescent, equal in height to the anther connective apices; ovules 2 per carpel, attached laterally.
<emphasis id="A4CFEAB1FF97FFCAFAB2FB40FAE3FB60" box="[1293,1356,1181,1206]" italics="true" pageId="1" pageNumber="240">Torus</emphasis>
convex, slightly concave at the apex, 3.04.0 mm in diameter, 1.82.0 mm thick, glabrous.
<emphasis id="A4CFEAB1FF97FFCAFB87FB1CFBD3FB0C" box="[1080,1148,1217,1242]" italics="true" pageId="1" pageNumber="240">Fruits</emphasis>
with 2228 monocarps on pedicels 10.015.0 mm long,
<quantity id="51439B46FF97FFCAFE70FB39FDF9FB29" box="[463,598,1252,1279]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="1" pageNumber="240" unit="mm" value="1.65" valueMax="1.8" valueMin="1.5">1.51.8 mm</quantity>
thick, glabrous.
<emphasis id="A4CFEAB1FF97FFCAFCB1FB38FC21FB28" box="[782,910,1253,1278]" italics="true" pageId="1" pageNumber="240">Monocarps</emphasis>
green
<emphasis id="A4CFEAB1FF97FFCAFC68FB38FB8EFB28" box="[983,1057,1253,1278]" italics="true" pageId="1" pageNumber="240">in vivo</emphasis>
(immature), ellipsoid to oblongoid, 8.010.0 × 7.08.0 mm, base obtuse, apex mucronate, stipes 18.025.0 mm long, 1.0
<quantity id="51439B46FF97FFCAFBE7FAD5FB01FAF5" box="[1112,1198,1288,1315]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="240" unit="mm" value="1.2">1.2 mm</quantity>
thick, glabrous.
<emphasis id="A4CFEAB1FF97FFCAFAD3FAD5FA03FAF4" box="[1388,1452,1288,1314]" italics="true" pageId="1" pageNumber="240">Seeds</emphasis>
not seen.
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF97FFCAFF02FA8DFCE9FA59" pageId="1" pageNumber="240" type="distribution">
<paragraph id="960436A3FF97FFCAFF02FA8DFCE9FA59" blockId="1.[136,1452,568,1962]" pageId="1" pageNumber="240">
<emphasis id="A4CFEAB1FF97FFCAFF02FA8DFE58FABC" bold="true" box="[189,503,1360,1386]" pageId="1" pageNumber="240">Distribution and Ecology:</emphasis>
—Scattered in shaded areas of tropical rain forest, in
<collectingRegion id="547FF841FF97FFCAFBE1FA8DFAE8FABC" box="[1118,1351,1360,1386]" country="Thailand" name="Narathiwat" pageId="1" pageNumber="240">Narathiwat Province</collectingRegion>
near the southern border of Peninsular
<collectingCountry id="EEAC7633FF97FFCAFE64FAA9FDEEFA58" box="[475,577,1396,1422]" name="Thailand" pageId="1" pageNumber="240">Thailand</collectingCountry>
, ca.
<quantity id="51439B46FF97FFCAFDCCFAA9FD15FA59" box="[627,698,1396,1423]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="240" unit="m" value="200.0">200 m</quantity>
alt. (
<figureCitation id="0E802A26FF97FFCAFD4CFAA9FC98FA59" box="[755,823,1396,1423]" captionStart="FIGURE 7" captionStartId="10.[136,229,1555,1577]" captionTargetBox="[178,1417,221,1505]" captionTargetId="figure-21@10.[151,1436,190,1531]" captionTargetPageId="10" captionText="FIGURE 7. A. &amp; B. Maps showing distributions of Polyalthia heliopetala (●), P. taweensis (▲) and P. pumila (■) in southern Peninsular Thailand. Map in B is an enlargement of the area shown in the rectangle in A. Maps by Pasakorn Bunchalee." figureDoi="http://doi.org/10.5281/zenodo.5426436" httpUri="https://zenodo.org/record/5426436/files/figure.png" pageId="1" pageNumber="240">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF97FFCAFF02FA45FC17FA64" box="[189,952,1432,1458]" pageId="1" pageNumber="240" type="biology_ecology">
<paragraph id="960436A3FF97FFCAFF02FA45FC17FA64" blockId="1.[136,1452,568,1962]" box="[189,952,1432,1458]" pageId="1" pageNumber="240">
<emphasis id="A4CFEAB1FF97FFCAFF02FA45FEECFA64" bold="true" box="[189,323,1432,1458]" pageId="1" pageNumber="240">Phenology:</emphasis>
—Flowering MarchApril, fruiting in April and August.
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF97FFCAFF02FA61FDF8FA00" box="[189,599,1468,1495]" pageId="1" pageNumber="240" type="vernacular_names">
<paragraph id="960436A3FF97FFCAFF02FA61FDF8FA00" blockId="1.[136,1452,568,1962]" box="[189,599,1468,1495]" pageId="1" pageNumber="240">
<emphasis id="A4CFEAB1FF97FFCAFF02FA61FEFDFA00" bold="true" box="[189,338,1468,1494]" pageId="1" pageNumber="240">Local name:</emphasis>
—Thai: Lueang suriya.
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF97FFCAFF02FA3DFA93FA2D" box="[189,1340,1504,1531]" pageId="1" pageNumber="240" type="etymology">
<paragraph id="960436A3FF97FFCAFF02FA3DFA93FA2D" blockId="1.[136,1452,568,1962]" box="[189,1340,1504,1531]" pageId="1" pageNumber="240">
<emphasis id="A4CFEAB1FF97FFCAFF02FA3DFEE7FA2C" bold="true" box="[189,328,1504,1530]" pageId="1" pageNumber="240">Etymology:</emphasis>
—The specific epithet refers to the spreading orange petals resembling the rays of the sun.
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF97FFCAFF02F9D9FDF6F9B0" pageId="1" pageNumber="240" type="conservation">
<paragraph id="960436A3FF97FFCAFF02F9D9FDF6F9B0" blockId="1.[136,1452,568,1962]" pageId="1" pageNumber="240">
<emphasis id="A4CFEAB1FF97FFCAFF02F9D9FDA5F9C8" bold="true" box="[189,522,1540,1566]" pageId="1" pageNumber="240">IUCN Conservation status:</emphasis>
—Known from two localities along the southern border of Peninsular
<collectingCountry id="EEAC7633FF97FFCAFAFEF9D9FA07F9C8" box="[1345,1448,1540,1566]" name="Thailand" pageId="1" pageNumber="240">Thailand</collectingCountry>
. However, there is insufficient information regarding the number of populations and population size. Therefore, the species is treated as Data Deficient (DD).
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF97FFCAFF02F9ADFCEAF904" pageId="1" pageNumber="240" type="materials_examined">
<paragraph id="960436A3FF97FFCAFF02F9ADFCEAF904" blockId="1.[136,1452,568,1962]" pageId="1" pageNumber="240">
<emphasis id="A4CFEAB1FF97FFCAFF02F9ADFDFBF95C" bold="true" box="[189,596,1648,1674]" pageId="1" pageNumber="240">Additional specimens examined:</emphasis>
<materialsCitation id="26D33CFEFF97FFCAFDD0F9ADFAB4F978" collectingDate="2003-03-18" collectionCode="BKF" collectorName="Koonkhunthod &amp; Chasuwan &amp; Berg" country="Thailand" elevation="200" location="Hala-Bala Wildlife Sanctuary" municipality="Waeng District" pageId="1" pageNumber="240" specimenCount="1" stateProvince="Narathiwat">
<collectingCountry id="EEAC7633FF97FFCAFDD0F9ADFCACF95C" box="[623,771,1648,1674]" name="Thailand" pageId="1" pageNumber="240">THAILAND</collectingCountry>
. PENINSULAR:
<collectingRegion id="547FF841FF97FFCAFC61F9ADFBF2F95C" box="[990,1117,1648,1674]" country="Thailand" name="Narathiwat" pageId="1" pageNumber="240">Narathiwat</collectingRegion>
,
<collectingMunicipality id="7660ACD9FF97FFCAFBCEF9ACFA89F95C" box="[1137,1318,1648,1675]" pageId="1" pageNumber="240">Waeng District</collectingMunicipality>
,
<location id="93646078FF97FFCAFA85F9ADFEF1F978" LSID="urn:lsid:plazi:treatment:1E1287B5FF97FFCAFF37FF29FF46F87C:93646078FF97FFCAFA85F9ADFEF1F978" country="Thailand" municipality="Waeng District" name="Hala-Bala Wildlife Sanctuary" pageId="1" pageNumber="240" stateProvince="Narathiwat">Hala-Bala Wildlife Sanctuary</location>
, alt.
<quantity id="51439B46FF97FFCAFE2AF949FE4FF979" box="[405,480,1684,1711]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="240" unit="m" value="200.0">
<elevation id="1D96D190FF97FFCAFE2AF949FE4FF979" box="[405,480,1684,1711]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="240" unit="m" value="200.0">200 m</elevation>
</quantity>
,
<date id="E2051063FF97FFCAFE52F949FD35F978" box="[493,666,1684,1710]" pageId="1" pageNumber="240" value="2003-03-18">
<collectingDate id="F241E98BFF97FFCAFE52F949FD35F978" box="[493,666,1684,1710]" pageId="1" pageNumber="240" value="2003-03-18">18 March 2003</collectingDate>
</date>
(flower),
<emphasis id="A4CFEAB1FF97FFCAFCB1F949FB69F978" box="[782,1222,1684,1710]" italics="true" pageId="1" pageNumber="240">
<collectorName id="3B4E5375FF97FFCAFCB1F949FC19F978" box="[782,950,1684,1710]" pageId="1" pageNumber="240">Koonkhunthod</collectorName>
,
<collectorName id="3B4E5375FF97FFCAFC7DF949FB97F978" box="[962,1080,1684,1710]" pageId="1" pageNumber="240">Chasuwan</collectorName>
&amp;
<collectorName id="3B4E5375FF97FFCAFBE0F948FB3BF978" box="[1119,1172,1685,1710]" pageId="1" pageNumber="240">Berg</collectorName>
393
</emphasis>
(
<collectionCode id="F0AAAE66FF97FFCAFB68F949FABCF978" box="[1239,1299,1684,1710]" collectionName="BKF" pageId="1" pageNumber="240">BKF</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="26D33CFEFF97FFCAFA97F949FCEEF904" collectingDate="1999-08-03" collectionCode="BKF" collectorName="Puudjaa" location="Waeng" pageId="1" pageNumber="240" specimenCount="1" stateProvince="Narathiwat">
<collectingRegion id="547FF841FF97FFCAFA97F949FA08F978" box="[1320,1447,1684,1710]" country="Thailand" name="Narathiwat" pageId="1" pageNumber="240">Narathiwat</collectingRegion>
,
<location id="93646078FF97FFCAFF37F964FF77F905" LSID="urn:lsid:plazi:treatment:1E1287B5FF97FFCAFF37FF29FF46F87C:93646078FF97FFCAFF37F964FF77F905" box="[136,216,1721,1747]" name="Waeng" pageId="1" pageNumber="240" stateProvince="Narathiwat">Waeng</location>
,
<location id="93646078FF97FFCAFF5CF965FEF8F904" LSID="urn:lsid:plazi:treatment:1E1287B5FF97FFCAFF37FF29FF46F87C:93646078FF97FFCAFF5CF965FEF8F904" box="[227,343,1720,1746]" name="Bala-Hala" pageId="1" pageNumber="240" stateProvince="Narathiwat">Bala-Hala</location>
,
<date id="E2051063FF97FFCAFEDCF965FDA9F904" box="[355,518,1720,1746]" pageId="1" pageNumber="240" value="1999-08-03">
<collectingDate id="F241E98BFF97FFCAFEDCF965FDA9F904" box="[355,518,1720,1746]" pageId="1" pageNumber="240" value="1999-08-03">3 August 1999</collectingDate>
</date>
(fruit),
<emphasis id="A4CFEAB1FF97FFCAFDE1F965FD41F904" box="[606,750,1720,1746]" italics="true" pageId="1" pageNumber="240">
<collectorName id="3B4E5375FF97FFCAFDE1F965FD12F904" box="[606,701,1720,1746]" pageId="1" pageNumber="240">Puudjaa</collectorName>
626
</emphasis>
(
<collectionCode id="F0AAAE66FF97FFCAFD42F965FC96F904" box="[765,825,1720,1746]" collectionName="BKF" pageId="1" pageNumber="240">BKF</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DEA16528FF97FFCAFF02F901FF46F87C" pageId="1" pageNumber="240" type="discussion">
<paragraph id="960436A3FF97FFCAFF02F901FF46F87C" blockId="1.[136,1452,568,1962]" pageId="1" pageNumber="240">
<emphasis id="A4CFEAB1FF97FFCAFF02F901FEA6F920" bold="true" box="[189,265,1756,1782]" pageId="1" pageNumber="240">Notes:</emphasis>
<taxonomicName id="51BB4D20FF97FFCAFE99F901FDB1F920" authorityName="Leerat. &amp; Bunchalee" authorityYear="2021" box="[294,542,1756,1782]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="240" phylum="Tracheophyta" rank="species" species="heliopetala">
<emphasis id="A4CFEAB1FF97FFCAFE99F901FDB1F920" box="[294,542,1756,1782]" italics="true" pageId="1" pageNumber="240">Polyalthia heliopetala</emphasis>
</taxonomicName>
is contrasted with
<taxonomicName id="51BB4D20FF97FFCAFD51F900FCCBF920" authorityName=", King" authorityYear="1892" box="[750,868,1756,1782]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="240" phylum="Tracheophyta" rank="species" species="oblonga">
<emphasis id="A4CFEAB1FF97FFCAFD51F900FCCBF920" box="[750,868,1756,1782]" italics="true" pageId="1" pageNumber="240">P. oblonga</emphasis>
</taxonomicName>
and
<taxonomicName id="51BB4D20FF97FFCAFC26F900FB87F920" authorityName=", Turner" authorityYear="2014" box="[921,1064,1756,1782]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="240" phylum="Tracheophyta" rank="species" species="motleyana">
<emphasis id="A4CFEAB1FF97FFCAFC26F900FB87F920" box="[921,1064,1756,1782]" italics="true" pageId="1" pageNumber="240">P. motleyana</emphasis>
</taxonomicName>
in
<tableCitation id="DB390318FF97FFCAFBF6F901FB33F920" box="[1097,1180,1756,1782]" captionStart="TABLE 1" captionStartId="4.[136,224,159,183]" captionTargetBox="[144,1442,232,755]" captionText="TABLE 1 Character comparison of Polyalthia heliopetala, P. oblonga, and P. motleyana s.s. Measurements from King converted from inches used in the original description." pageId="1" pageNumber="240">Table 1</tableCitation>
.
<bibRefCitation id="F22A4B52FF97FFCAFB18F900FAEBF920" author="Turner, I. M." box="[1191,1348,1756,1783]" pageId="1" pageNumber="240" refId="ref5369" refString="Turner, I. M. (2014) Annonaceae. In: Soepadmo, E., Saw, L. G., Chung, R. C. K. &amp; Kiew, R. (Eds.) Tree flora of Sabah and Sarawak, Volume 8. Forest Research Institute Malaysia (FRIM), Kepong, Malaysia." type="book" year="2014">Turner (2014)</bibRefCitation>
accepted
<taxonomicName id="51BB4D20FF97FFCAFF37F8DCFEACF8CC" authorityName=", King" authorityYear="1892" box="[136,259,1792,1818]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="240" phylum="Tracheophyta" rank="species" species="oblonga">
<emphasis id="A4CFEAB1FF97FFCAFF37F8DCFEACF8CC" box="[136,259,1792,1818]" italics="true" pageId="1" pageNumber="240">P. oblonga</emphasis>
</taxonomicName>
and
<taxonomicName id="51BB4D20FF97FFCAFEFFF8DCFE7CF8CC" authorityName=", Turner" authorityYear="2014" box="[320,467,1792,1818]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="240" phylum="Tracheophyta" rank="species" species="motleyana">
<emphasis id="A4CFEAB1FF97FFCAFEFFF8DCFE7CF8CC" box="[320,467,1792,1818]" italics="true" pageId="1" pageNumber="240">P. motleyana</emphasis>
</taxonomicName>
as distinct species, but his description of
<taxonomicName id="51BB4D20FF97FFCAFC00F8DCFB95F8CC" authorityName=", King" authorityYear="1892" box="[959,1082,1792,1818]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="240" phylum="Tracheophyta" rank="species" species="oblonga">
<emphasis id="A4CFEAB1FF97FFCAFC00F8DCFB95F8CC" box="[959,1082,1792,1818]" italics="true" pageId="1" pageNumber="240">P. oblonga</emphasis>
</taxonomicName>
differs sufficiently from Kings protologue that data are given separately from the two sources. For
<taxonomicName id="51BB4D20FF97FFCAFCCEF8F8FBAFF8E8" authorityName=", Turner" authorityYear="2014" box="[881,1024,1828,1854]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="240" phylum="Tracheophyta" rank="species" species="motleyana">
<emphasis id="A4CFEAB1FF97FFCAFCCEF8F8FBAFF8E8" box="[881,1024,1828,1854]" italics="true" pageId="1" pageNumber="240">P. motleyana</emphasis>
</taxonomicName>
the description of
<bibRefCitation id="F22A4B52FF97FFCAFB72F8F8FAC6F8E8" author="Turner, I. M." box="[1229,1385,1828,1855]" pageId="1" pageNumber="240" refId="ref5369" refString="Turner, I. M. (2014) Annonaceae. In: Soepadmo, E., Saw, L. G., Chung, R. C. K. &amp; Kiew, R. (Eds.) Tree flora of Sabah and Sarawak, Volume 8. Forest Research Institute Malaysia (FRIM), Kepong, Malaysia." type="book" year="2014">Turner (2014)</bibRefCitation>
alone was used for comparison. A distinctive feature of the petals of
<taxonomicName id="51BB4D20FF97FFCAFCF5F894FC4BF8B4" authorityName="Leerat. &amp; Bunchalee" authorityYear="2021" box="[842,996,1864,1890]" class="Magnoliopsida" family="Annonaceae" genus="Polyalthia" kingdom="Plantae" order="Magnoliales" pageId="1" pageNumber="240" phylum="Tracheophyta" rank="species" species="heliopetala">
<emphasis id="A4CFEAB1FF97FFCAFCF5F894FC4BF8B4" box="[842,996,1864,1890]" italics="true" pageId="1" pageNumber="240">P. heliopetala</emphasis>
</taxonomicName>
that has not been described for either of the other two species is the raised and thickened patch of tissue at the base of both the outer and the inner petals (
<figureCitation id="0E802A26FF97FFCAFACCF8B1FF03F87C" captionStart="FIGURE 1" captionStartId="2.[136,229,1916,1938]" captionTargetBox="[199,1380,218,1873]" captionTargetId="figure-21@2.[178,1409,203,1890]" captionTargetPageId="2" captionText="FIGURE 1. Polyalthia heliopetala: A. Habit; B. Leaf bases (upper side) and inflorescences, with point of inflorescence attachments to stem indicated by arrows; C. Flower showing recurved petals; D. Flower, top view showing the embossed patch on the inner base of the petals; E. Outer surfaces of outer petals; F. Outer surfaces of inner petals; G. Stamens and persistent sepal on pedicel; H. Torus with some stamens removed to show attachment of stamens and carpels on the convex torus surface; I. Carpel; J. Stamens (outside on left, inside on right); K. Monocarps. The illustration was drawn by Nattapong Chomlok from Leeratiwong 20-1527 (BKF)." figureDoi="http://doi.org/10.5281/zenodo.5426423" httpUri="https://zenodo.org/record/5426423/files/figure.png" pageId="1" pageNumber="240">Figs. 1D</figureCitation>
,
<figureCitation id="0E802A26FF97FFCAFF07F84DFF72F87C" box="[184,221,1936,1962]" captionStart="FIGURE 2" captionStartId="3.[136,229,1920,1942]" captionTargetBox="[179,1406,197,1886]" captionTargetId="figure-18@3.[164,1423,187,1898]" captionTargetPageId="3" captionText="FIGURE 2. Polyalthia heliopetala: A. Leaves and inflorescences; B. Lower surface of leaf base and flower; C. Flower showing the embossed basal patch of the inner surface of the petals; D. Upper surface of leaves and flower; E. Lower surface of leaf showing venation; F. Monocarps.A, B, D and E are from Leeratiwong 20-1527(BKF); F is from Puudjaa 626 (BKF)]; all photographs by Pasakorn Bunchalee. Photograph in C was taken by Yamilah Sama-ae in Narathiwat, Ra Ngae District, Bo Ngo Subdistrict, Bue Chaeng Village, 7 April 2020." figureDoi="http://doi.org/10.5281/zenodo.5426425" httpUri="https://zenodo.org/record/5426425/files/figure.png" pageId="1" pageNumber="240">2C</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>