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<document ID-DOI="http://dx.doi.org/10.3897/BDJ.9.e65970" ID-Pensoft-Pub="1314-2828-9-e65970" ID-Pensoft-UUID="A559676C573554B8A4CFB45D00F7A876" ModsDocID="1314-2828-9-e65970" checkinTime="1633362685362" checkinUser="pensoft" docAuthor="Fassbender, Nico, Stefanoudis, Paris V, Filander, Zoleka Nontlantla, Gendron, Gilberte, Mah, Christopher L, Mattio, Lydiane, Mortimer, Jeanne A, Moura, Carlos J, Samaai, Toufiek, Samimi-Namin, Kaveh, Wagner, Daniel, Walton, Rowana &amp; Woodall, Lucy C" docDate="2021" docId="7D277E8A7EB05488831F789BC5F3787F" docLanguage="en" docName="BiodivDatJour 9: e65970" docOrigin="Biodiversity Data Journal 9" docPubDate="2021-08-27" docSource="http://dx.doi.org/10.3897/BDJ.9.e65970" docTitle="Theonella swinhoei subsp. f. fm. Creamy brown encrusting sponges" docType="treatment" docVersion="2" id="A559676C573554B8A4CFB45D00F7A876" lastPageNumber="65970" masterDocId="A559676C573554B8A4CFB45D00F7A876" masterDocTitle="Reef benthos of Seychelles - A field guide" masterLastPageNumber="65970" masterPageNumber="65970" pageNumber="65970" updateTime="1635278358358" updateUser="pensoft">
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<mods:title>Reef benthos of Seychelles - A field guide</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Fassbender, Nico</mods:namePart>
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<mods:namePart>Stefanoudis, Paris V</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4040-8364</mods:nameIdentifier>
<mods:affiliation>Department of Zoology, University of Oxford, Oxford, United Kingdom &amp; Nekton Foundation, Oxford, United Kingdom</mods:affiliation>
</mods:name>
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<mods:role>
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<mods:namePart>Filander, Zoleka Nontlantla</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6905-4440</mods:nameIdentifier>
<mods:affiliation>Department of Forestry, Fisheries and Environment, Branch Oceans and Coasts, Cape Town, South Africa</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gendron, Gilberte</mods:namePart>
<mods:affiliation>Sustainable Ocean Seychelles, Victoria, Seychelles</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mah, Christopher L</mods:namePart>
<mods:affiliation>Smithsonian Institution National Museum of Natural History, Washington, United States of America</mods:affiliation>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Mattio, Lydiane</mods:namePart>
<mods:affiliation>University of Cape Town, Rondebosch, Cape Town, South Africa &amp; blue [c] weed, Brest, France</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mortimer, Jeanne A</mods:namePart>
<mods:affiliation>Seychelles' Conservation &amp; Climate Adaptation Trust (SeyCCAT), Victoria, Mahe, Seychelles &amp; Department of Biology, University of Florida, Gainesville, Florida, United States of America &amp; Island Conservation Society (ICS), Point Larue, Mahe, Seychelles</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Moura, Carlos J</mods:namePart>
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<mods:affiliation>OKEANOS / DOP, University of the Azores, Horta, Portugal</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Samaai, Toufiek</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7269-293X</mods:nameIdentifier>
<mods:affiliation>Department of Forestry, Fisheries and Environment, Branch Oceans and Coasts, Cape Town, South Africa &amp; University of Cape Town, Rondebosch, Cape Town, South Africa &amp; iZiko Museums of South Africa, Cape Town, South Africa &amp; University of the Western Cape, Bellville, Cape Town, South Africa</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Samimi-Namin, Kaveh</mods:namePart>
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<mods:affiliation>Naturalis Biodiversity Center, Leiden, Netherlands</mods:affiliation>
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<mods:namePart>Wagner, Daniel</mods:namePart>
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<mods:name type="personal">
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<mods:namePart>Walton, Rowana</mods:namePart>
<mods:affiliation>James Michel Blue Economy Research Institute, University of Seychelles, Anse Royale, Mahe ́, Seychelles</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Woodall, Lucy C</mods:namePart>
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<mods:affiliation>Department of Zoology, University of Oxford, Oxford, United Kingdom &amp; Nekton Foundation, Oxford, United Kingdom</mods:affiliation>
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<mods:date>2021</mods:date>
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<mods:number>2021-08-27</mods:number>
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<treatment LSID="urn:lsid:plazi:treatment:7D277E8A7EB05488831F789BC5F3787F" httpUri="http://treatment.plazi.org/id/7D277E8A7EB05488831F789BC5F3787F" lastPageNumber="65970" pageId="0" pageNumber="65970">
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<taxonomicName LSID="7D277E8A-7EB0-5488-831F-789BC5F3787F" class="Hexactinellida" family="Theonellidae" form="Creamy brown encrusting sponges" genus="Theonella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="subsp. f." order="Amphidiscosida" pageId="0" pageNumber="65970" phylum="Porifera" rank="form" species="Swinhoei" subSpecies="f.">&quot;class Demospongiae&quot; order indet. sp. 1 f. Creamy brown encrusting sponges</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65970" type="materials_examined">
<paragraph pageId="0" pageNumber="65970">Materials</paragraph>
<paragraph pageId="0" pageNumber="65970">
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectingMethod="Submersible OR Remotely Operated Vehicle OR SCUBA" collectorName="Taxon, Demospongiae, Animalia, Porifera, Location, Indian Ocean, Aldabra W, Alphonse N, Astove W, D'Arros N, Desroches S, Poivre E, Submersible OR Remotely Operated Vehicle OR" country="Seychelles" county="Human" determinerName="Nico Fassbender, Toufiek Samaai, Paris Stefanoudis" elevation="10" location="Aldabra W 1, Alphonse N 1, Astove W 1, D'Arros N 1, Desroches S 1, Poivre E 1" municipality="Record Level" pageId="0" pageNumber="65970" specimenCount="1" typeStatus="Other material">
<emphasis bold="true" pageId="0" pageNumber="65970">Type status:</emphasis>
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectingMethod="Submersible OR Remotely Operated Vehicle OR SCUBA" collectorName="Taxon, Demospongiae, Animalia, Porifera, Location, Indian Ocean, Aldabra W, Alphonse N, Astove W, D'Arros N, Desroches S, Poivre E, Submersible OR Remotely Operated Vehicle OR" country="Seychelles" county="Human" determinerName="Nico Fassbender, Toufiek Samaai, Paris Stefanoudis" elevation="10" location="Aldabra W 1, Alphonse N 1, Astove W 1, D'Arros N 1, Desroches S 1, Poivre E 1" municipality="Record Level" specimenCount="1" typeStatus="Other material">
<typeStatus pageId="0" pageNumber="65970">Other material</typeStatus>
.
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<collectorName>Taxon</collectorName>
:
</emphasis>
scientificName:
<collectorName>Demospongiae</collectorName>
; kingdom:
<collectorName>Animalia</collectorName>
; phylum:
<collectorName>Porifera</collectorName>
; class:
<collectorName>Demospongiae</collectorName>
;
<emphasis bold="true" pageId="0" pageNumber="65970">
<collectorName>Location</collectorName>
:
</emphasis>
waterBody:
<collectorName>Indian Ocean</collectorName>
; country:
<collectingCountry name="Seychelles" pageId="0" pageNumber="65970">Seychelles</collectingCountry>
; locality:
<location LSID="urn:lsid:plazi:treatment:7D277E8A7EB05488831F789BC5F3787F:E07B1DA8B9421F3DBEB5BF9B6C6059BD" country="Seychelles" county="Human" municipality="Record Level" name="Aldabra W 1, Alphonse N 1, Astove W 1, D'Arros N 1, Desroches S 1, Poivre E 1" pageId="0" pageNumber="65970">
<collectorName>Aldabra W</collectorName>
1,
<collectorName>Alphonse N</collectorName>
1,
<collectorName>Astove W</collectorName>
1,
<collectorName>D'Arros N</collectorName>
1,
<collectorName>Desroches S</collectorName>
1,
<collectorName>Poivre E</collectorName>
1
</location>
; minimumDepthInMeters:
<quantity metricMagnitude="0" metricUnit="m" metricValue="9.5" unit="m" value="9.5">
<elevation metricMagnitude="0" metricUnit="m" metricValue="9.5" unit="m" value="9.5">9.5 m</elevation>
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; maximumDepthInMeters:
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.283" unit="m" value="128.3">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.283" unit="m" value="128.3">128.3 m</elevation>
</quantity>
; locationRemarks:
<location LSID="urn:lsid:plazi:treatment:7D277E8A7EB05488831F789BC5F3787F:2EEC1F82E88E5025B8CE0697011DC97C" country="Seychelles" county="Human" municipality="Record Level" name="First Descent">First Descent</location>
:
<collectingCountry name="Seychelles">Seychelles</collectingCountry>
<location LSID="urn:lsid:plazi:treatment:7D277E8A7EB05488831F789BC5F3787F:FEE6119B006571A59FE5EA9C7D1DECF5" country="Seychelles" county="Human" municipality="Record Level" name="Expedition">Expedition</location>
;
<emphasis bold="true" pageId="0" pageNumber="65970">
<location LSID="urn:lsid:plazi:treatment:7D277E8A7EB05488831F789BC5F3787F:E81B8CA04FBADBAD256FEA742971E536" country="Seychelles" county="Human" municipality="Record Level" name="Identification">Identification</location>
:
</emphasis>
identifiedBy:
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<location LSID="urn:lsid:plazi:treatment:7D277E8A7EB05488831F789BC5F3787F:37A0FA913FC219A15D079A5573B56C7D" country="Seychelles" county="Human" municipality="Record Level" name="Nico Fassbender">Nico Fassbender</location>
,
<location LSID="urn:lsid:plazi:treatment:7D277E8A7EB05488831F789BC5F3787F:EEC028985078C65FE08234689514DC4B" country="Seychelles" county="Human" municipality="Record Level" name="Toufiek Samaai">Toufiek Samaai</location>
,
<location LSID="urn:lsid:plazi:treatment:7D277E8A7EB05488831F789BC5F3787F:F3138617EC9963F38A410F383802CDC7" country="Seychelles" county="Human" municipality="Record Level" name="Paris Stefanoudis">Paris Stefanoudis</location>
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; dateIdentified: 2019, 2020; identificationRemarks: identified only from imagery;
<emphasis bold="true" pageId="0" pageNumber="65970">
<location LSID="urn:lsid:plazi:treatment:7D277E8A7EB05488831F789BC5F3787F:92E6B81CD91ED061511366A8C6455FCF" country="Seychelles" county="Human" municipality="Record Level" name="Event">Event</location>
:
</emphasis>
samplingProtocol:
<collectingMethod pageId="0" pageNumber="65970">
<collectorName>Submersible OR Remotely Operated Vehicle OR</collectorName>
SCUBA
</collectingMethod>
;
<emphasis bold="true" pageId="0" pageNumber="65970">
<collectingMunicipality>Record Level</collectingMunicipality>
:
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basisOfRecord:
<collectingCounty>Human</collectingCounty>
observation
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<paragraph pageId="0" pageNumber="65970">Notes</paragraph>
<paragraph pageId="0" pageNumber="65970">
Encrusting sponge with a velvety smooth surface texture. Colouration creamy brown, sometimes mottled with yellow patches. This group likely contains a variety of species belonging to different families; however, consistent positive identification was not always possible (e.g. due to distance from the camera) and, in most cases, requires microscopic examination. Observed genera included
<taxonomicName authorityName="Gray" authorityYear="1867" class="Demospongiae" family="Suberitidae" genus="Aaptos" kingdom="Animalia" lsidName="Aaptos" order="Suberitida" pageId="0" pageNumber="65970" phylum="Porifera" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="65970">Aaptos</emphasis>
</taxonomicName>
sp. (Fig.
<figureCitation captionStartId="F6744596" pageId="0" pageNumber="65970">177</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="C7FAEAD8AE72BB694898ED85FC9641F8" bold="true" box="[151,789,151,178]" pageId="9" pageNumber="138">
Taxon—
<taxonomicName id="328E4D49AE72BB69490EED8AFC9641F8" authority="Boulenger, 1908" authorityName="Boulenger" authorityYear="1908" box="[257,789,151,178]" class="Reptilia" family="Agamidae" genus="Malayodracon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="robinsonii">
<emphasis id="C7FAEAD8AE72BB69490EED8AFDB741FB" bold="true" box="[257,564,151,177]" italics="true" pageId="9" pageNumber="138">Malayodracon robinsonii</emphasis>
(
<bibRefCitation id="911F4B3BAE72BB694A4CED85FC8E41F8" author="Boulenger" box="[579,781,152,178]" pageId="9" pageNumber="138" refString="Boulenger, G. A. (1908) Report on the Gunong Tahan expedition, May - Sept. 1905. III. Fishes, batrachians and reptiles. Journal of the Federated Malay States Museums, 3, 61 - 69." type="journal article" year="1908">Boulenger, 1908</bibRefCitation>
)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="F53136CAAE72BB694898EDC3FE9B41BE" blockId="9.[151,1437,222,490]" box="[151,280,222,244]" pageId="9" pageNumber="138">Chresonymy</paragraph>
</subSubSection>
<subSubSection id="BD946541AE72BB694898EDE0FD5640A0" pageId="9" pageNumber="138" type="reference_group">
<paragraph id="F53136CAAE72BB694898EDE0FCD94059" blockId="9.[151,1437,222,490]" box="[151,858,253,275]" pageId="9" pageNumber="138">
<treatmentCitationGroup id="D59E11E4AE72BB694898EDE0FCD94059" box="[151,858,253,275]" pageId="9" pageNumber="138">
<taxonomicName id="328E4D49AE72BB694898EDE0FE214059" ID-CoL="8GQXS" box="[151,418,253,275]" class="Squamata" family="Agamidae" genus="Gonyocephalus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="robinsonii">
<emphasis id="C7FAEAD8AE72BB694898EDE0FE214059" box="[151,418,253,275]" italics="true" pageId="9" pageNumber="138">Gonyocephalus robinsonii</emphasis>
</taxonomicName>
—
<treatmentCitation id="742F10DBAE72BB6949B3EDE0FD044059" author="Boulenger" box="[444,647,253,275]" page="65" pageId="9" pageNumber="138" year="1908">
<bibRefCitation id="911F4B3BAE72BB6949B3EDE0FDE24059" author="Boulenger" box="[444,609,253,275]" pageId="9" pageNumber="138" refString="Boulenger, G. A. (1908) Report on the Gunong Tahan expedition, May - Sept. 1905. III. Fishes, batrachians and reptiles. Journal of the Federated Malay States Museums, 3, 61 - 69." type="journal article" year="1908">Boulenger 1908</bibRefCitation>
: 65
</treatmentCitation>
;
<treatmentCitation id="742F10DBAE72BB694A9DEDE0FCD94059" author="Boulenger" box="[658,858,253,275]" page="67" pageId="9" pageNumber="138" year="1912">
<bibRefCitation id="911F4B3BAE72BB694A9DEDE0FCB44059" author="Boulenger" box="[658,823,253,275]" pageId="9" pageNumber="138" refString="Boulenger, G. A. (1912) A vertebrate fauna of the Malay Peninsula from the Isthmus of Kra to Singapore incl. the adjacent islands. Reptilia and Amphibia. Taylor &amp; Francis, London, XIII + 298 pp." type="book" year="1912">Boulenger 1912</bibRefCitation>
: 67
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="F53136CAAE72BB694898EC01FDD94078" blockId="9.[151,1437,222,490]" box="[151,602,284,306]" pageId="9" pageNumber="138">
<treatmentCitationGroup id="D59E11E4AE72BB694898EC01FDD94078" box="[151,602,284,306]" pageId="9" pageNumber="138">
<emphasis id="C7FAEAD8AE72BB694898EC01FE324078" box="[151,433,284,306]" italics="true" pageId="9" pageNumber="138">
<taxonomicName id="328E4D49AE72BB694898EC01FEB74078" box="[151,308,284,306]" class="Squamata" family="Agamidae" genus="Gonyocephalus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="138" phylum="Chordata" rank="genus">Gonyocephalus</taxonomicName>
robinsoni—
</emphasis>
<treatmentCitation id="742F10DBAE72BB6949BEEC01FDD94078" author="Smith" box="[433,602,284,306]" page="269" pageId="9" pageNumber="138" year="1922">
<bibRefCitation id="911F4B3BAE72BB6949BEEC01FDA84078" author="Smith" box="[433,555,284,306]" pageId="9" pageNumber="138" refString="Smith, M. A. (1922) On a collection of reptiles and batrachians from the mountains of Pahang, Malay Peninsula. Journal of the Federated Malay States Museums, 10, 263 - 282." type="journal article" year="1922">Smith 1922</bibRefCitation>
: 269
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="F53136CAAE72BB694898EC27FCA2401A" blockId="9.[151,1437,222,490]" box="[151,801,314,336]" pageId="9" pageNumber="138">
<treatmentCitationGroup id="D59E11E4AE72BB694898EC27FCA2401A" box="[151,801,314,336]" pageId="9" pageNumber="138">
<taxonomicName id="328E4D49AE72BB694898EC27FE0C401A" box="[151,399,314,336]" class="Reptilia" family="Agamidae" genus="Gonocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="robinsoni">
<emphasis id="C7FAEAD8AE72BB694898EC27FE0C401A" box="[151,399,314,336]" italics="true" pageId="9" pageNumber="138">Gonocephalus robinsoni</emphasis>
</taxonomicName>
—
<treatmentCitation id="742F10DBAE72BB6949A6EC27FDCB401A" author="Smith" box="[425,584,314,336]" page="24" pageId="9" pageNumber="138" year="1930">
<bibRefCitation id="911F4B3BAE72BB6949A6EC27FDA2401A" author="Smith" box="[425,545,314,336]" pageId="9" pageNumber="138" refString="Smith, M. A. (1930) The Reptilia and Amphibia of the Malay Peninsula. Bulletin of the Raffles Museum, 3, 1 - 149." type="journal article" year="1930">Smith 1930</bibRefCitation>
: 24
</treatmentCitation>
;
<treatmentCitation id="742F10DBAE72BB694A5DEC27FCA2401A" author="Smirnova" box="[594,801,314,336]" page="128" pageId="9" pageNumber="138" year="2003">
<bibRefCitation id="911F4B3BAE72BB694A5DEC27FD72401A" author="Smirnova" box="[594,753,314,336]" pageId="9" pageNumber="138" refString="Smirnova, Y. A. (2003) Dermal papillous scales at the Rock Agamas of the genus Laudakia (Sauria, Agamidae). Modern Herpetology. Collected Papers, 2, 124 - 136. [in Russian]" type="journal article" year="2003">Smirnova 2003</bibRefCitation>
: 128
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="F53136CAAE72BB694898EC44FB8A4025" blockId="9.[151,1437,222,490]" box="[151,1033,345,367]" pageId="9" pageNumber="138">
<treatmentCitationGroup id="D59E11E4AE72BB694898EC44FB8A4025" box="[151,1033,345,367]" pageId="9" pageNumber="138">
<taxonomicName id="328E4D49AE72BB694898EC44FE154025" box="[151,406,345,367]" class="Squamata" family="Agamidae" genus="Goniocephalus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="robinsoni">
<emphasis id="C7FAEAD8AE72BB694898EC44FE154025" box="[151,406,345,367]" italics="true" pageId="9" pageNumber="138">Goniocephalus robinsoni</emphasis>
</taxonomicName>
—
<treatmentCitation id="742F10DBAE72BB6949BFEC44FDDF4025" author="Smith" box="[432,604,345,367]" page="133" pageId="9" pageNumber="138" year="1935">
<bibRefCitation id="911F4B3BAE72BB6949BFEC44FDAA4025" author="Smith" box="[432,553,345,367]" pageId="9" pageNumber="138" refString="Smith, M. A. (1935) The Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese subregion. Reptilia and Amphibia. Vol. II. Sauria. Taylor &amp; Francis, London, xiii + 440 pp." type="book" year="1935">Smith 1935</bibRefCitation>
: 133
</treatmentCitation>
;
<treatmentCitation id="742F10DBAE72BB694A69EC44FD744025" author="Sly" box="[614,759,345,367]" page="156" pageId="9" pageNumber="138" year="1976">
<bibRefCitation id="911F4B3BAE72BB694A69EC44FD474025" author="Sly" box="[614,708,345,367]" pageId="9" pageNumber="138" refString="Sly, G. R. (1976) New locality records for some Peninsular Malaysia amphibia, reptilia and mammalia. Malayan Nature Journal, 29 (3), 155 - 157." type="journal article" year="1976">Sly 1976</bibRefCitation>
: 156
</treatmentCitation>
;
<bibRefCitation id="911F4B3BAE72BB694B0EEC47FC0B4025" author="Bourret" box="[769,904,345,367]" pageId="9" pageNumber="138" refString="Bourret, R. (1947 [2009]) Les Lezards de l'Indochine. Ed. Chimaira, Frankfurt / M., 624 pp." type="book" year="1947">Bourret 1947</bibRefCitation>
“2009”: 211
</treatmentCitationGroup>
</paragraph>
<paragraph id="F53136CAAE72BB694898EC65FADA4081" blockId="9.[151,1437,222,490]" pageId="9" pageNumber="138">
<treatmentCitationGroup id="D59E11E4AE72BB694898EC65FADA4081" pageId="9" pageNumber="138">
<taxonomicName id="328E4D49AE72BB694898EC65FE1F40C4" box="[151,412,376,398]" class="Reptilia" family="Agamidae" genus="Gonocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="robinsonii">
<emphasis id="C7FAEAD8AE72BB694898EC65FE1F40C4" box="[151,412,376,398]" italics="true" pageId="9" pageNumber="138">Gonocephalus robinsonii</emphasis>
</taxonomicName>
—
<treatmentCitation id="742F10DBAE72BB6949B9EC65FD0A40C4" author="Smedley" box="[438,649,376,398]" page="110" pageId="9" pageNumber="138" year="1931">
<bibRefCitation id="911F4B3BAE72BB6949B9EC65FDD240C4" author="Smedley" box="[438,593,376,398]" pageId="9" pageNumber="138" refString="Smedley, N. (1931) Amphibians and reptiles from the Cameron Highlands, Malay Peninsula. Bulletin of the Raffles Museum, 6, 105 - 123." type="journal article" year="1931">Smedley 1931</bibRefCitation>
: 110
</treatmentCitation>
;
<treatmentCitation id="742F10DBAE72BB694A95EC65FCE540C4" author="Wermuth" box="[666,870,376,398]" page="61" pageId="9" pageNumber="138" year="1967">
<bibRefCitation id="911F4B3BAE72BB694A95EC65FCB940C4" author="Wermuth" box="[666,826,376,398]" pageId="9" pageNumber="138" refString="Wermuth, H. (1967) Liste der rezenten Amphibien und Reptilien: Agamidae. Das Tierreich, 86, i - xiv + 1 - 127." type="journal article" year="1967">Wermuth 1967</bibRefCitation>
: 61
</treatmentCitation>
;
<treatmentCitation id="742F10DBAE72BB694B78EC65FBBF40C4" author="Moody" box="[887,1084,376,398]" page="299" pageId="9" pageNumber="138" year="1980">
<bibRefCitation id="911F4B3BAE72BB694B78EC65FB8140C4" author="Moody" box="[887,1026,376,398]" pageId="9" pageNumber="138" refString="Moody, S. M. (1980) Phylogenetic and historical biogeographical relationship of the genera in the family Agamidae (Reptilia: Lacertilia). Ph. D. Thesis, University of Michigan, Ann Arbor, Michigan, 373 pp." type="book" year="1980">Moody 1980</bibRefCitation>
: 299
</treatmentCitation>
; Denzer &amp; Manthey 1991: 312;
<treatmentCitation id="742F10DBAE72BB6948C8EC8BFE7D40E6" author="Manthey" box="[199,510,406,428]" page="16" pageId="9" pageNumber="138" year="1992">
<bibRefCitation id="911F4B3BAE72BB6948C8EC8BFE5440E6" author="Manthey" box="[199,471,406,428]" pageId="9" pageNumber="138" refString="Manthey, U. &amp; Denzer, W. (1992 a) Die Echten Winkelkopfagamen der Gattung Gonocephalus Kaup (Sauria: Agamidae) IV. Gonocephalus mjoebergi Smith, 1925 und Gonocephalus robinsonii Boulenger, 1908. Sauria, 14 (1), 15 - 19." type="journal article" year="1992" yearSuffix="a">Manthey &amp; Denzer 1992a</bibRefCitation>
: 16
</treatmentCitation>
;
<treatmentCitation id="742F10DBAE72BB694A05EC8BFCC640E6" author="Manthey" box="[522,837,406,428]" page="65" pageId="9" pageNumber="138" year="1992">
<bibRefCitation id="911F4B3BAE72BB694A05EC8BFC9E40E6" author="Manthey" box="[522,797,406,428]" pageId="9" pageNumber="138" refString="Manthey, U. &amp; Schuster, N. (1992) Agamen. Herpetologischer Fachverlag, Munster, 120 pp." type="book" year="1992">Manthey &amp; Schuster 1992</bibRefCitation>
: 65
</treatmentCitation>
;
<treatmentCitation id="742F10DBAE72BB694B5EEC8BFB3740E6" author="Manthey" box="[849,1204,406,428]" page="189" pageId="9" pageNumber="138" year="1997">
<bibRefCitation id="911F4B3BAE72BB694B5EEC8BFBFC40E6" author="Manthey" box="[849,1151,406,428]" pageId="9" pageNumber="138" refString="Manthey, U. &amp; Grossmann, W. (1997) Amphibien &amp; Reptilien Sudostasiens. Natur und Tier Verlag, Munster, 512 pp." type="book" year="1997">Manthey &amp; Grossmann 1997</bibRefCitation>
: 189
</treatmentCitation>
;
<treatmentCitation id="742F10DBAE72BB694CCFEC8BFF734081" author="Chan-ard" page="102" pageId="9" pageNumber="138" year="1999">
<bibRefCitation id="911F4B3BAE72BB694CCFEC8BFA1B40E6" author="Chan-ard" box="[1216,1432,406,429]" pageId="9" pageNumber="138" refString="Chan-ard, T., Grossmann, W., Gumprecht, A. &amp; Schulz, K. D. (1999) Amphibians and reptiles of peninsular Malaysia and Thailand-an illustrated checklist [bilingual English and German]. Bushmaster Publications, Wurselen, 240 pp." type="book" year="1999">
Chan-ard
<emphasis id="C7FAEAD8AE72BB694D29EC85FAD840E6" box="[1318,1371,406,429]" italics="true" pageId="9" pageNumber="138">et al.</emphasis>
1999
</bibRefCitation>
: 102
</treatmentCitation>
;
<treatmentCitation id="742F10DBAE72BB6948F5ECA8FE564081" author="Diong" box="[250,469,437,459]" page="73" pageId="9" pageNumber="138" year="2000">
<bibRefCitation id="911F4B3BAE72BB6948F5ECA8FE334081" author="Diong" box="[250,432,437,459]" pageId="9" pageNumber="138" refString="Diong, C. - H., Low, M. - H., Tan, E. - C., Yong, H. - S., Hikida, T. &amp; Ota, H. (2000) On the monophyly of the agamid genus Gonocephalus Kaup, 1825 (Reptilia: Agamidae): a chromosomal perspective. Current Herpetology, 19, 71 - 79. http: // dx. doi. org / 10.5358 / hsj. 19.71" type="journal article" year="2000">
Diong
<emphasis id="C7FAEAD8AE72BB69494FECABFEF04081" box="[320,371,437,459]" italics="true" pageId="9" pageNumber="138">et al.</emphasis>
2000
</bibRefCitation>
: 73
</treatmentCitation>
;
<bibRefCitation id="911F4B3BAE72BB6949D0ECA8FD1B4081" author="Honda" box="[479,664,437,459]" pageId="9" pageNumber="138" refString="Honda, M., Ota, H., Sengoku, S., Yong, H. - S. &amp; Hikida, T. (2002) Molecular Evaluation of Phylogenetic Significances in the Highly Divergent Karyotypes of the Genus Gonocephalus (Reptilia: Agamidae) from Tropical Asia. Zoological Science, 19, 129 - 133. http: // dx. doi. org / 10.2108 / zsj. 19.129" type="journal article" year="2002">
Honda
<emphasis id="C7FAEAD8AE72BB694A26ECABFDDF4081" box="[553,604,437,459]" italics="true" pageId="9" pageNumber="138">et al.</emphasis>
2002
</bibRefCitation>
;
<treatmentCitation id="742F10DBAE72BB694AADECABFC564081" author="Denzer" box="[674,981,437,459]" page="256" pageId="9" pageNumber="138" year="2009">
<bibRefCitation id="911F4B3BAE72BB694AADECABFC214081" author="Denzer" box="[674,930,437,459]" pageId="9" pageNumber="138" refString="Denzer, W. &amp; Manthey, U. (2009) Remarks on the type specimen of Gonocephalus mjobergi Smith, 1925 (Sauria: Agamidae). Bonner Zoologische Beitrage, 56 (4), 255 - 258." type="journal article" year="2009">Denzer &amp; Manthey 2009</bibRefCitation>
: 256
</treatmentCitation>
;
<treatmentCitation id="742F10DBAE72BB694BD0ECA8FB234081" author="Grismer" box="[991,1184,437,459]" page="258" pageId="9" pageNumber="138" year="2011">
<bibRefCitation id="911F4B3BAE72BB694BD0ECA8FBEE4081" author="Grismer" box="[991,1133,437,459]" pageId="9" pageNumber="138" refString="Grismer, L. L. (2011) Lizards of Peninsular Malaysia, Singapore and their adjacent archipelagos. Edition Chimaira, Frankfurt / M., 728 pp." type="book" year="2011">Grismer 2011</bibRefCitation>
: 258
</treatmentCitation>
;
<bibRefCitation id="911F4B3BAE72BB694CA4ECABFADA4081" author="Pyron" box="[1195,1369,437,459]" pageId="9" pageNumber="138" refString="Pyron, R. A., Burbrink, F. T. &amp; Wiens, J. J. (2013) A phylogeny and revised classification of Squamata, including 4161 species of lizards and snakes. BMC Evolutionary Biology, 13, 93. http: // dx. doi. org / 10.1186 / 1471 - 2148 - 13 - 93" type="journal article" year="2013">
Pyron
<emphasis id="C7FAEAD8AE72BB694CE2ECABFAA24081" box="[1261,1313,437,459]" italics="true" pageId="9" pageNumber="138">et al.</emphasis>
2013
</bibRefCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="F53136CAAE72BB694898ECC9FD5640A0" blockId="9.[151,1437,222,490]" box="[151,725,468,490]" pageId="9" pageNumber="138">
<treatmentCitationGroup id="D59E11E4AE72BB694898ECC9FD5640A0" box="[151,725,468,490]" pageId="9" pageNumber="138">
(
<emphasis id="C7FAEAD8AE72BB694890ECC9FE0F40A0" box="[159,396,468,490]" italics="true" pageId="9" pageNumber="138">
<taxonomicName id="328E4D49AE72BB694890ECC9FEDA40A0" box="[159,345,468,490]" class="Reptilia" family="Agamidae" genus="Gonocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="inc">Gonocephalus inc</taxonomicName>
. sed.
</emphasis>
)
<taxonomicName id="328E4D49AE72BB694995ECC9FD8240A0" box="[410,513,468,490]" class="Reptilia" family="Agamidae" genus="Gonocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="robinsonii">
<emphasis id="C7FAEAD8AE72BB694995ECC9FD8240A0" box="[410,513,468,490]" italics="true" pageId="9" pageNumber="138">robinsonii</emphasis>
</taxonomicName>
—
<treatmentCitation id="742F10DBAE72BB694A14ECC9FD5640A0" author="Manthey" box="[539,725,468,490]" page="46" pageId="9" pageNumber="138" year="2010">
<bibRefCitation id="911F4B3BAE72BB694A14ECC9FD3140A0" author="Manthey" box="[539,690,468,490]" pageId="9" pageNumber="138" refString="Manthey, U. (2010) Agamid Lizards of Southern Asia - Draconinae 2, Leiolepidinae." type="journal volume" year="2010">Manthey 2010</bibRefCitation>
: 46
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BD946541AE72BB694898EF0BFA9A4525" pageId="9" pageNumber="138" type="diagnosis">
<paragraph id="F53136CAAE72BB694898EF0BFB9443F4" blockId="9.[151,1436,534,1999]" pageId="9" pageNumber="138">
<emphasis id="C7FAEAD8AE72BB694898EF0BFE924365" bold="true" box="[151,273,534,559]" pageId="9" pageNumber="138">Diagnosis.</emphasis>
Medium sized lizard; males: SVL
<quantity id="32769B2FAE72BB694AA3EF0BFCC64364" box="[684,837,534,559]" metricMagnitude="-1" metricUnit="m" metricValue="1.365" metricValueMax="1.58" metricValueMin="1.15" pageId="9" pageNumber="138" unit="mm" value="136.5" valueMax="158.0" valueMin="115.0">115–158 mm</quantity>
, TL
<quantity id="32769B2FAE72BB694B8FEF0BFB984364" box="[896,1051,534,559]" metricMagnitude="-1" metricUnit="m" metricValue="3.025" metricValueMax="3.2" metricValueMin="2.85" pageId="9" pageNumber="138" unit="mm" value="302.5" valueMax="320.0" valueMin="285.0">285–320 mm</quantity>
; females: SVL
<quantity id="32769B2FAE72BB694CDDEF0BFAE94364" box="[1234,1386,534,559]" metricMagnitude="-1" metricUnit="m" metricValue="1.2349999999999999" metricValueMax="1.3199999999999998" metricValueMin="1.15" pageId="9" pageNumber="138" unit="mm" value="123.5" valueMax="132.0" valueMin="115.0">115–132 mm</quantity>
, TL
<quantity id="32769B2FAE72BB694898EF26FEAF4318" box="[151,300,571,595]" metricMagnitude="-1" metricUnit="m" metricValue="2.95" metricValueMax="3.0" metricValueMin="2.9" pageId="9" pageNumber="138" unit="mm" value="295.0" valueMax="300.0" valueMin="290.0">290–300 mm</quantity>
(
<typeStatus id="2A358868AE72BB694935EF26FEEF4319" box="[314,364,571,595]" pageId="9" pageNumber="138">type</typeStatus>
specimen, male, SVL
<quantity id="32769B2FAE72BB694A65EF27FD4A4318" box="[618,713,570,595]" metricMagnitude="-1" metricUnit="m" metricValue="1.52" pageId="9" pageNumber="138" unit="mm" value="152.0">152 mm</quantity>
, TL
<quantity id="32769B2FAE72BB694AF2EF26FCDD4319" box="[765,862,571,595]" metricMagnitude="-1" metricUnit="m" metricValue="3.2" pageId="9" pageNumber="138" unit="mm" value="320.0">320 mm</quantity>
), TL up to 2.6 times SVL; body triangular in crosssection; no patagia; head large, elongated and pointed; bony protuberances on the occipital region; sharp canthus rostralis and rounded superciliary edge; tympanum exposed; antehumeral fold present, transverse gular fold absent; large gular sac; nuchal and dorsal crest continuous; no preanal or femoral pores.
</paragraph>
<paragraph id="F53136CAAE72BB6948C8EFD7FE124205" blockId="9.[151,1436,534,1999]" pageId="9" pageNumber="138">
<emphasis id="C7FAEAD8AE72BB6948C8EFD7FD2843A9" bold="true" box="[199,683,714,739]" pageId="9" pageNumber="138">Comparison to other draconine genera.</emphasis>
Morphologically
<taxonomicName id="328E4D49AE72BB694B73EFD7FB1D43A9" box="[892,1182,714,739]" class="Reptilia" family="Agamidae" genus="Malayodracon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="robinsonii">
<emphasis id="C7FAEAD8AE72BB694B73EFD7FB1D43A9" box="[892,1182,714,739]" italics="true" pageId="9" pageNumber="138">Malayodracon robinsonii</emphasis>
</taxonomicName>
differs from all other agamid lizards by a combination of the following characters: visible tympanum, bony ridge in the occipital region, large gular sac with a rounded tip extending onto the chest, small weakly keeled, muricate dorsal scales and triangular crest scales.
</paragraph>
<paragraph id="F53136CAAE72BB6948C8EE46FA9A4525" blockId="9.[151,1436,534,1999]" pageId="9" pageNumber="138">
From its former congeners of the genus of
<taxonomicName id="328E4D49AE72BB694AD8EE47FCFA4239" box="[727,889,858,883]" class="Reptilia" family="Agamidae" genus="Gonocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="genus" sensu="lato">
<emphasis id="C7FAEAD8AE72BB694AD8EE47FCFA4239" box="[727,889,858,883]" italics="true" pageId="9" pageNumber="138">Gonocephalus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DCC957A3AE72BB694B89EE46FC374239" box="[902,948,858,883]" pageId="9" pageNumber="138" sensu="lato">s. l.</taxonomicNameLabel>
it differs additionally by the absence of supporting scales along the nuchal crest (vs. present) and the absence of a distinct transverse gular fold (vs. present). In particular
<taxonomicName id="328E4D49AE72BB69499BEEBFFD3742F1" box="[404,692,930,955]" class="Reptilia" family="Agamidae" genus="Malayodracon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="robinsonii">
<emphasis id="C7FAEAD8AE72BB69499BEEBFFD3742F1" box="[404,692,930,955]" italics="true" pageId="9" pageNumber="138">Malayodracon robinsonii</emphasis>
</taxonomicName>
differs from the
<taxonomicName id="328E4D49AE72BB694B77EEBFFBDF42F1" box="[888,1116,930,955]" class="Reptilia" family="Agamidae" genus="Gonocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="bellii">
<emphasis id="C7FAEAD8AE72BB694B77EEBFFBDF42F1" box="[888,1116,930,955]" italics="true" pageId="9" pageNumber="138">Gonocephalus bellii</emphasis>
</taxonomicName>
Group by the possession of triangular nuchal and dorsal crest scales (vs. spines), from the
<taxonomicName id="328E4D49AE72BB694B6CEEDAFC754295" box="[867,1014,966,991]" class="Reptilia" family="Agamidae" genus="Gonocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="megalepis">
<emphasis id="C7FAEAD8AE72BB694B6CEEDAFC754295" box="[867,1014,966,991]" italics="true" pageId="9" pageNumber="138">G. megalepis</emphasis>
</taxonomicName>
Group by the lack of a large conical scale at the posterior joint of mandibular and maxillar region (vs. present in
<emphasis id="C7FAEAD8AE72BB694BE6EEF7FB5A4549" box="[1001,1241,1002,1027]" italics="true" pageId="9" pageNumber="138">
<taxonomicName id="328E4D49AE72BB694BE6EEF7FBDF4549" box="[1001,1116,1002,1027]" class="Reptilia" family="Agamidae" genus="Gonocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="lacunosus">lacunosus</taxonomicName>
,
<taxonomicName id="328E4D49AE72BB694C68EEF7FB5A4549" box="[1127,1241,1002,1027]" class="Reptilia" family="Agamidae" genus="Gonocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="megalepis">megalepis</taxonomicName>
</emphasis>
and
<taxonomicName id="328E4D49AE72BB694D01EEF7FACD4549" box="[1294,1358,1002,1027]" class="Reptilia" family="Agamidae" genus="Gonocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="klossi">
<emphasis id="C7FAEAD8AE72BB694D01EEF7FACD4549" box="[1294,1358,1002,1027]" italics="true" pageId="9" pageNumber="138">klossi</emphasis>
</taxonomicName>
), from the
<taxonomicName id="328E4D49AE72BB6948CFE912FE1E456D" box="[192,413,1038,1063]" class="Reptilia" family="Agamidae" genus="Gonocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="chamaeleontinus">
<emphasis id="C7FAEAD8AE72BB6948CFE912FE1E456D" box="[192,413,1038,1063]" italics="true" pageId="9" pageNumber="138">G. chamaeleontinus</emphasis>
</taxonomicName>
Group by the shape of the supraciliary edge (rounded in
<taxonomicName id="328E4D49AE72BB694C10E912FB34456D" box="[1055,1207,1038,1063]" class="Reptilia" family="Agamidae" genus="Malayodracon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="robinsonii">
<emphasis id="C7FAEAD8AE72BB694C10E912FB34456D" box="[1055,1207,1038,1063]" italics="true" pageId="9" pageNumber="138">M. robinsonii</emphasis>
</taxonomicName>
vs. angular) and the form of the nuchal crest (single triangular scales in
<taxonomicName id="328E4D49AE72BB694ADDE929FCEB4501" box="[722,872,1074,1099]" class="Reptilia" family="Agamidae" genus="Malayodracon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="robinsonii">
<emphasis id="C7FAEAD8AE72BB694ADDE929FCEB4501" box="[722,872,1074,1099]" italics="true" pageId="9" pageNumber="138">M. robinsonii</emphasis>
</taxonomicName>
vs. erected nuchal crest consisting of several scale rows) and
<taxonomicName id="328E4D49AE72BB694902E94AFE014525" box="[269,386,1110,1135]" class="Reptilia" family="Agamidae" genus="Gonocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="grandis">
<emphasis id="C7FAEAD8AE72BB694902E94AFE014525" box="[269,386,1110,1135]" italics="true" pageId="9" pageNumber="138">G. grandis</emphasis>
</taxonomicName>
by the shape of the nuchal crest (crest scales of
<taxonomicName id="328E4D49AE72BB694B92E94AFB924525" box="[925,1041,1110,1135]" class="Reptilia" family="Agamidae" genus="Gonocephalus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="grandis">
<emphasis id="C7FAEAD8AE72BB694B92E94AFB924525" box="[925,1041,1110,1135]" italics="true" pageId="9" pageNumber="138">G. grandis</emphasis>
</taxonomicName>
fused similar to a sail).
</paragraph>
</subSubSection>
<subSubSection id="BD946541AE72BB6B48C8E967FBB543E2" lastPageId="11" lastPageNumber="140" pageId="9" pageNumber="138" type="reference_group">
<paragraph id="F53136CAAE72BB6948C8E967FE6444B0" blockId="9.[151,1436,534,1999]" pageId="9" pageNumber="138">
<taxonomicName id="328E4D49AE72BB6948C8E967FE6645D9" box="[199,485,1146,1171]" class="Reptilia" family="Agamidae" genus="Malayodracon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="robinsonii">
<emphasis id="C7FAEAD8AE72BB6948C8E967FE6645D9" box="[199,485,1146,1171]" italics="true" pageId="9" pageNumber="138">Malayodracon robinsonii</emphasis>
</taxonomicName>
differs from the species that appear in the same node of our phylogenetic analysis as follows: from all
<taxonomicName id="328E4D49AE72BB694953E983FE7A45FD" box="[348,505,1182,1207]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="genus">
<emphasis id="C7FAEAD8AE72BB694953E983FE7A45FD" box="[348,505,1182,1207]" italics="true" pageId="9" pageNumber="138">Acanthosaura</emphasis>
</taxonomicName>
by a lack of a large spine behind the orbit (vs. present in
<taxonomicName id="328E4D49AE72BB694C88E983FAA745FD" box="[1159,1316,1182,1207]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="genus">
<emphasis id="C7FAEAD8AE72BB694C88E983FAA745FD" box="[1159,1316,1182,1207]" italics="true" pageId="9" pageNumber="138">Acanthosaura</emphasis>
</taxonomicName>
) and from all
<taxonomicName id="328E4D49AE72BB6948B6E9DFFEBD4591" box="[185,318,1218,1243]" class="Reptilia" family="Agamidae" genus="Phoxophrys" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="genus">
<emphasis id="C7FAEAD8AE72BB6948B6E9DFFEBD4591" box="[185,318,1218,1243]" italics="true" pageId="9" pageNumber="138">Phoxophrys</emphasis>
</taxonomicName>
by an exposed tympanum and the absence of a blue mouth. From
<taxonomicName id="328E4D49AE72BB694C2DE9DFFA884591" box="[1058,1291,1218,1243]" class="Reptilia" family="Agamidae" genus="Japalura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="polygonata">
<emphasis id="C7FAEAD8AE72BB694C2DE9DFFA884591" box="[1058,1291,1218,1243]" italics="true" pageId="9" pageNumber="138">Japalura polygonata</emphasis>
</taxonomicName>
and
<taxonomicName id="328E4D49AE72BB694D4DE9D9FA054591" box="[1346,1414,1218,1243]" class="Reptilia" family="Agamidae" genus="Japalura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="luei">
<emphasis id="C7FAEAD8AE72BB694D4DE9D9FA054591" box="[1346,1414,1218,1243]" italics="true" pageId="9" pageNumber="138">J. luei</emphasis>
</taxonomicName>
it can be distinguished by the possession of small dorsal scales (vs. large heterogeneous scalation) as well as the possession of a dorsal crest. From all other species of
<taxonomicName id="328E4D49AE72BB694AF0E817FCE74469" box="[767,868,1290,1315]" class="Reptilia" family="Agamidae" genus="Japalura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="genus">
<emphasis id="C7FAEAD8AE72BB694AF0E817FCE74469" box="[767,868,1290,1315]" italics="true" pageId="9" pageNumber="138">Japalura</emphasis>
</taxonomicName>
it can be distinguished by its gular sac pholidosis which is small and smooth in
<taxonomicName id="328E4D49AE72BB6949E4E832FD00440D" box="[491,643,1326,1351]" class="Reptilia" family="Agamidae" genus="Malayodracon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="robinsonii">
<emphasis id="C7FAEAD8AE72BB6949E4E832FD00440D" box="[491,643,1326,1351]" italics="true" pageId="9" pageNumber="138">M. robinsonii</emphasis>
</taxonomicName>
and enlarged and often keeled in
<taxonomicName id="328E4D49AE72BB694C0CE833FBEB440D" box="[1027,1128,1326,1351]" class="Reptilia" family="Agamidae" genus="Japalura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="genus">
<emphasis id="C7FAEAD8AE72BB694C0CE833FBEB440D" box="[1027,1128,1326,1351]" italics="true" pageId="9" pageNumber="138">Japalura</emphasis>
</taxonomicName>
species. Additionally most
<taxonomicName id="328E4D49AE72BB694898E84FFF7F4421" box="[151,252,1362,1387]" class="Reptilia" family="Agamidae" genus="Japalura" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="genus">
<emphasis id="C7FAEAD8AE72BB694898E84FFF7F4421" box="[151,252,1362,1387]" italics="true" pageId="9" pageNumber="138">Japalura</emphasis>
</taxonomicName>
spp. have a concealed tympanum (exceptions are some Indian and Nepalese species formerly referred to as
<taxonomicName id="328E4D49AE72BB6948B9E86AFDD144C5" authority="Kastle &amp; Schleich 1998" authorityName="Kastle &amp; Schleich" authorityYear="1998" box="[182,594,1398,1423]" class="Squamata" family="Agamidae" genus="Oriotiaris" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="138" phylum="Chordata" rank="genus">
<emphasis id="C7FAEAD8AE72BB6948B9E86AFEA544C4" box="[182,294,1399,1422]" italics="true" pageId="9" pageNumber="138">Oriotiaris</emphasis>
(
<bibRefCitation id="911F4B3BAE72BB694939E86BFDCA44C5" author="Kastle" box="[310,585,1398,1423]" pageId="9" pageNumber="138" refString="Kastle, W. &amp; Schleich, H. H. (1998) Notes on Comparative Ethology and Taxonomy of the Genus Japalura. In: Schleich, H. H. &amp; Kastle, W. (Eds.), Contributions to the Herpetology of South-Asia (Nepal India). Veroffentlichungen aus dem Fuhlrott- Museum, Wuppertal, pp. 233 - 246." type="book chapter" year="1998">Kästle &amp; Schleich 1998</bibRefCitation>
)
</taxonomicName>
. From
<taxonomicName id="328E4D49AE72BB694AACE86BFCF144C5" box="[675,882,1398,1423]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="versicolor">
<emphasis id="C7FAEAD8AE72BB694AACE86BFCF144C5" box="[675,882,1398,1423]" italics="true" pageId="9" pageNumber="138">Calotes versicolor</emphasis>
</taxonomicName>
and all other species of that genus
<taxonomicName id="328E4D49AE72BB694D0BE86AFA1F44C5" box="[1284,1436,1398,1423]" class="Reptilia" family="Agamidae" genus="Malayodracon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="robinsonii">
<emphasis id="C7FAEAD8AE72BB694D0BE86AFA1F44C5" box="[1284,1436,1398,1423]" italics="true" pageId="9" pageNumber="138">M. robinsonii</emphasis>
</taxonomicName>
differs in pholidotic characters such as small dorsal and gular scales (vs enlarged dorsals in
<taxonomicName id="328E4D49AE72BB694CC5E886FADC44F9" box="[1226,1375,1434,1459]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="versicolor">
<emphasis id="C7FAEAD8AE72BB694CC5E886FADC44F9" box="[1226,1375,1434,1459]" italics="true" pageId="9" pageNumber="138">C. versicolor</emphasis>
</taxonomicName>
) and dorsal scales smaller than ventrals (vs. dorsal scales larger than ventral scales in
<taxonomicName id="328E4D49AE72BB694C13E8A2FB2C449D" box="[1052,1199,1470,1495]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="species" species="versicolor">
<emphasis id="C7FAEAD8AE72BB694C13E8A2FB2C449D" box="[1052,1199,1470,1495]" italics="true" pageId="9" pageNumber="138">C. versicolor</emphasis>
</taxonomicName>
) as well as the shape of the nuchal and dorsal crest.
</paragraph>
<paragraph id="F53136CAAE72BB6948C8EB1BFAEE4755" blockId="9.[151,1436,534,1999]" box="[199,1389,1542,1567]" pageId="9" pageNumber="138">
Furthermore it differs from the remaining Southeast Asian genera in the subfamily
<taxonomicName id="328E4D49AE72BB694C65EB1BFB684755" box="[1130,1259,1542,1567]" class="Reptilia" family="Agamidae" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="subFamily" subFamily="Draconinae">Draconinae</taxonomicName>
as follows:
</paragraph>
<paragraph id="F53136CAAE72BB6948C8EB37FEB647C1" blockId="9.[151,1436,534,1999]" pageId="9" pageNumber="138">
<taxonomicName id="328E4D49AE72BB6948C8EB37FE524709" authority="Peters, 1864" authorityName="Peters" authorityYear="1864" box="[199,465,1578,1603]" class="Reptilia" family="Agamidae" genus="Aphaniotis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="genus">
<emphasis id="C7FAEAD8AE72BB6948C8EB37FEC34709" box="[199,320,1578,1603]" italics="true" pageId="9" pageNumber="138">Aphaniotis</emphasis>
<bibRefCitation id="911F4B3BAE72BB694948EB36FE524709" author="Peters" box="[327,465,1578,1603]" pageId="9" pageNumber="138" refString="Peters, W. (1864) Uber einige neue Saugethiere (Mormops, Macrotus, Vesperus, Molossus, Capromys), Amphibien (Plathydactylus, Otocryptis, Euprepes, Ungalia, Dromicus, Tropidonotus, Xenodon, Hylodes), und Fische (Sillago, Sebastes, Channa, Myctophum, Carassius, Barbus). Monatsberichte der Koniglich Preussischen Akademie der Wissenschaften zu Berlin, 1864, 381 - 399." type="journal article" year="1864">Peters, 1864</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="328E4D49AE72BB694A09EB37FCA24709" authority="Manthey" authorityName="Manthey" box="[518,801,1578,1603]" class="Reptilia" family="Agamidae" genus="Pseudocophotis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="genus">
<emphasis id="C7FAEAD8AE72BB694A09EB37FD3B4709" box="[518,696,1578,1603]" italics="true" pageId="9" pageNumber="138">Pseudocophotis</emphasis>
Manthey
</taxonomicName>
in
<bibRefCitation id="911F4B3BAE72BB694B4BEB37FB0C4709" author="Manthey" box="[836,1167,1578,1603]" pageId="9" pageNumber="138" refString="Manthey, U. &amp; Grossmann, W. (1997) Amphibien &amp; Reptilien Sudostasiens. Natur und Tier Verlag, Munster, 512 pp." type="book" year="1997">Manthey &amp; Grossmann, 1997</bibRefCitation>
by a visible tympanum; additionally by absence of a blue mouth or a rostral appendage and from
<taxonomicName id="328E4D49AE72BB694BF5EB53FB28472D" box="[1018,1195,1614,1639]" class="Reptilia" family="Agamidae" genus="Pseudocophotis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="genus">
<emphasis id="C7FAEAD8AE72BB694BF5EB53FB28472D" box="[1018,1195,1614,1639]" italics="true" pageId="9" pageNumber="138">Pseudocophotis</emphasis>
</taxonomicName>
by the absence of a prehensile tail
</paragraph>
<paragraph id="F53136CAAE72BB6948C8EB8BFB1247E5" blockId="9.[151,1436,534,1999]" box="[199,1169,1686,1711]" pageId="9" pageNumber="138">
<taxonomicName id="328E4D49AE72BB6948C8EB8BFE5D47E5" authority="Kaup, 1827" authorityName="Kaup" authorityYear="1827" box="[199,478,1686,1711]" class="Reptilia" family="Agamidae" genus="Bronchocela" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="genus">
<emphasis id="C7FAEAD8AE72BB6948C8EB8BFED647E5" box="[199,341,1686,1711]" italics="true" pageId="9" pageNumber="138">Bronchocela</emphasis>
<bibRefCitation id="911F4B3BAE72BB694953EB8AFE5D47E5" author="Kaup" box="[348,478,1687,1711]" pageId="9" pageNumber="138" refString="Kaup, F. (1827) Zoologische Monographien. Isis von Oken, 20, 610 - 625, pls. 1 - 2." type="journal article" year="1827">Kaup, 1827</bibRefCitation>
</taxonomicName>
by the absence of a lateral skin fold on both sides of the neck
</paragraph>
<paragraph id="F53136CAAE72BB6948C8EBA7FAF64799" blockId="9.[151,1436,534,1999]" box="[199,1397,1722,1747]" pageId="9" pageNumber="138">
<taxonomicName id="328E4D49AE72BB6948C8EBA7FE3A4799" authority="Manthey" authorityName="Manthey" box="[199,441,1722,1747]" class="Reptilia" family="Agamidae" genus="Complicitus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="genus">
<emphasis id="C7FAEAD8AE72BB6948C8EBA7FECC4799" box="[199,335,1722,1747]" italics="true" pageId="9" pageNumber="138">Complicitus</emphasis>
Manthey
</taxonomicName>
in
<bibRefCitation id="911F4B3BAE72BB6949D2EBA7FCAA4799" author="Manthey" box="[477,809,1722,1747]" pageId="9" pageNumber="138" refString="Manthey, U. &amp; Grossmann, W. (1997) Amphibien &amp; Reptilien Sudostasiens. Natur und Tier Verlag, Munster, 512 pp." type="book" year="1997">Manthey &amp; Grossmann, 1997</bibRefCitation>
by the absence of lateral pockets on the gular pouch
</paragraph>
<paragraph id="F53136CAAE72BB6948C8EBC3FF4F4650" blockId="9.[151,1436,534,1999]" pageId="9" pageNumber="138">
<taxonomicName id="328E4D49AE72BB6948C8EBC3FE6347BD" authority="Fitzinger" authorityName="Fitzinger" box="[199,480,1758,1783]" class="Reptilia" family="Agamidae" genus="Coryphophylax" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="genus">
<emphasis id="C7FAEAD8AE72BB6948C8EBC3FEF747BD" box="[199,372,1758,1783]" italics="true" pageId="9" pageNumber="138">Coryphophylax</emphasis>
Fitzinger
</taxonomicName>
in
<bibRefCitation id="911F4B3BAE72BB694A09EBC3FD5E47BD" author="Steindachner" box="[518,733,1758,1783]" pageId="9" pageNumber="138" refString="Steindachner, F. (1867) Reise der osterreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wullerstorf-Urbair. Zoologischer Theil. Repilien. K. K. Hof- und Staatsdruckerei, Wien, 3 pls., 98 pp." type="book" year="1867">Steindachner, 1867</bibRefCitation>
by the absence of a nuchal sail and the possession of a dorsal crest
</paragraph>
<paragraph id="F53136CAAE72BB6948C8EA3BFC784675" blockId="9.[151,1436,534,1999]" box="[199,1019,1830,1855]" pageId="9" pageNumber="138">
<taxonomicName id="328E4D49AE72BB6948C8EA3BFE644675" authority="Doria, 1888" authorityName="Doria" authorityYear="1888" box="[199,487,1830,1855]" class="Reptilia" family="Agamidae" genus="Dendragama" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="genus">
<emphasis id="C7FAEAD8AE72BB6948C8EA3BFED84675" box="[199,347,1830,1855]" italics="true" pageId="9" pageNumber="138">Dendragama</emphasis>
<bibRefCitation id="911F4B3BAE72BB69496DEA3BFE644675" author="Doria" box="[354,487,1830,1855]" pageId="9" pageNumber="138" refString="Doria, G. (1888) Note Erpetologiche - Alcuni nuovi Sauri raccolti in Sumatra dal Dr. O. Beccari. Annali del Museo Civico di Storia Naturale di Genova, 2, Series 6, 646 - 652." type="journal article" year="1888">Doria, 1888</bibRefCitation>
</taxonomicName>
by cranioskeletal features and gular pouch size
</paragraph>
<paragraph id="F53136CAAE72BB6948C8EA51FC3B4629" blockId="9.[151,1436,534,1999]" box="[199,952,1866,1891]" pageId="9" pageNumber="138">
<taxonomicName id="328E4D49AE72BB6948C8EA51FE414629" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[199,450,1866,1891]" class="Reptilia" family="Agamidae" genus="Draco" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="genus">
<emphasis id="C7FAEAD8AE72BB6948C8EA51FE8C4629" box="[199,271,1868,1891]" italics="true" pageId="9" pageNumber="138">Draco</emphasis>
Linnaeus, 1758
</taxonomicName>
by the absence of elongated ribs and patagia
</paragraph>
<paragraph id="F53136CAAE72BB6948C8EA72FAEB46CD" blockId="9.[151,1436,534,1999]" box="[199,1384,1902,1927]" pageId="9" pageNumber="138">
<taxonomicName id="328E4D49AE72BB6948C8EA72FD9946CD" authority="Boulenger, 1885" authorityName="Boulenger" authorityYear="1885" box="[199,538,1902,1927]" class="Reptilia" family="Agamidae" genus="Harpesaurus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="genus">
<emphasis id="C7FAEAD8AE72BB6948C8EA72FEDA46CC" box="[199,345,1903,1926]" italics="true" pageId="9" pageNumber="138">Harpesaurus</emphasis>
<bibRefCitation id="911F4B3BAE72BB69496FEA73FD9946CD" author="Boulenger" box="[352,538,1902,1927]" pageId="9" pageNumber="138" refString="Boulenger, G. A. (1885) Catalogue of the Lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. Trustees of the British Museum, London, 450 pp." type="book" year="1885">Boulenger, 1885</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="328E4D49AE72BB694A5FEA73FC3F46CD" authority="Parker, 1924" authorityName="Parker" authorityYear="1924" box="[592,956,1902,1927]" class="Reptilia" family="Agamidae" genus="Thaumatorhynchus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="genus">
<emphasis id="C7FAEAD8AE72BB694A5FEA73FCAB46CD" box="[592,808,1902,1927]" italics="true" pageId="9" pageNumber="138">Thaumatorhynchus</emphasis>
<bibRefCitation id="911F4B3BAE72BB694B20EA73FC3F46CD" author="Parker" box="[815,956,1902,1927]" pageId="9" pageNumber="138" refString="Parker, H. W. (1924) Description of a new agamid lizard from Sumatra. The Annals and Magazine of Natural History, Series 9, 14, 624 - 625. http: // dx. doi. org / 10.1080 / 00222932408633173" type="journal article" year="1924">Parker, 1924</bibRefCitation>
</taxonomicName>
by the absence of a rostral appendage
</paragraph>
<paragraph id="F53136CAAE72BB6948C8EA8FFCCE4685" blockId="9.[151,1436,534,1999]" pageId="9" pageNumber="138">
<taxonomicName id="328E4D49AE72BB6948C8EA8FFDFA46E1" authority="Manthey &amp; Denzer, 2000" authorityName="Manthey &amp; Denzer" authorityYear="2000" box="[199,633,1938,1963]" class="Reptilia" family="Agamidae" genus="Hypsicalotes" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="138" phylum="Chordata" rank="genus">
<emphasis id="C7FAEAD8AE72BB6948C8EA8FFEDB46E1" box="[199,344,1938,1963]" italics="true" pageId="9" pageNumber="138">Hypsicalotes</emphasis>
<bibRefCitation id="911F4B3BAE72BB694951EA8FFDFA46E1" author="Manthey" box="[350,633,1938,1963]" pageId="9" pageNumber="138" refString="Manthey, U. &amp; Denzer, W. (2000) Description of a new genus, Hypsicalotes gen nov. (Sauria: Agamidae) from Mount Kinabalu, North Borneo, with remarks on the generic identity of Gonocephalus schultzewestrumi Urban, 1999. Hamadryad, 25 (1), 13 - 20." type="journal article" year="2000">Manthey &amp; Denzer, 2000</bibRefCitation>
</taxonomicName>
by the absence of large plates on either side of the head and the absence of large lanceolate scales along the midline of the gular pouch
</paragraph>
<paragraph id="F53136CAAE71BB6A48C8ED8AFC4B419F" blockId="10.[151,1437,151,357]" pageId="10" pageNumber="139">
<taxonomicName id="328E4D49AE71BB6A48C8ED8AFDF141FA" authority="Ananjeva &amp; Stuart, 2001" authorityName="Ananjeva &amp; Stuart" authorityYear="2001" box="[199,626,151,176]" class="Reptilia" family="Agamidae" genus="Mantheyus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="139" phylum="Chordata" rank="genus">
<emphasis id="C7FAEAD8AE71BB6A48C8ED8AFEC141FA" box="[199,322,151,176]" italics="true" pageId="10" pageNumber="139">Mantheyus</emphasis>
<bibRefCitation id="911F4B3BAE71BB6A4943ED8AFDF141FA" author="Ananjeva" box="[332,626,151,176]" pageId="10" pageNumber="139" refString="Ananjeva, N. B. &amp; Stuart, B. L. (2001) The agamid lizard Ptyctolaemus phuwuanensis Manthey and Nabhitabhata, 1991 from Thailand and Laos represents a new genus. Russian Journal of Herpetology, 8 (3), 165 - 170." type="journal article" year="2001">Ananjeva &amp; Stuart, 2001</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="328E4D49AE71BB6A4A8EED8AFC3241FA" authority="Peters, 1864" authorityName="Peters" authorityYear="1864" box="[641,945,151,176]" class="Reptilia" family="Agamidae" genus="Ptyctolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="139" phylum="Chordata" rank="genus">
<emphasis id="C7FAEAD8AE71BB6A4A8EED8AFC9B41FA" box="[641,792,151,176]" italics="true" pageId="10" pageNumber="139">Ptyctolaemus</emphasis>
<bibRefCitation id="911F4B3BAE71BB6A4B2CED85FC3241FA" author="Peters" box="[803,945,151,176]" pageId="10" pageNumber="139" refString="Peters, W. (1864) Uber einige neue Saugethiere (Mormops, Macrotus, Vesperus, Molossus, Capromys), Amphibien (Plathydactylus, Otocryptis, Euprepes, Ungalia, Dromicus, Tropidonotus, Xenodon, Hylodes), und Fische (Sillago, Sebastes, Channa, Myctophum, Carassius, Barbus). Monatsberichte der Koniglich Preussischen Akademie der Wissenschaften zu Berlin, 1864, 381 - 399." type="journal article" year="1864">Peters, 1864</bibRefCitation>
</taxonomicName>
and “
<taxonomicName id="328E4D49AE71BB6A4BF4ED8AFB1E41FA" box="[1019,1181,151,176]" class="Reptilia" family="Agamidae" genus="Gonocephalus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="139" phylum="Chordata" rank="genus">
<emphasis id="C7FAEAD8AE71BB6A4BF4ED8AFB1E41FA" box="[1019,1181,151,176]" italics="true" pageId="10" pageNumber="139">Gonocephalus</emphasis>
</taxonomicName>
” (Genus A)
<taxonomicName id="328E4D49AE71BB6A4D37ED8AFEA3419F" authority="Smith, 1925" authorityName="Smith" authorityYear="1925" class="Reptilia" family="Agamidae" genus="Gonocephalus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="139" phylum="Chordata" rank="species" species="mjobergi">
<emphasis id="C7FAEAD8AE71BB6A4D37ED8AFA1F41FA" box="[1336,1436,151,176]" italics="true" pageId="10" pageNumber="139">mjobergi</emphasis>
<bibRefCitation id="911F4B3BAE71BB6A4898EDA1FEA3419F" author="Smith" box="[151,288,188,213]" pageId="10" pageNumber="139" refString="Smith, M. A. (1925) On a collection of reptiles and amphibians from Mt. Murud, Borneo. Sarawak Museum Journal, 8, 5 - 14." type="journal article" year="1925">Smith, 1925</bibRefCitation>
</taxonomicName>
by the absence of longitudinal skin folds on the gular region
</paragraph>
<paragraph id="F53136CAAE71BB6A48C8EDC2FC3541B2" blockId="10.[151,1437,151,357]" box="[199,950,223,248]" pageId="10" pageNumber="139">
<taxonomicName id="328E4D49AE71BB6A48C8EDC2FE7141B2" authority="Kaup, 1827" authorityName="Kaup" authorityYear="1827" box="[199,498,223,248]" class="Reptilia" family="Agamidae" genus="Pseudocalotes" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="139" phylum="Chordata" rank="genus">
<emphasis id="C7FAEAD8AE71BB6A48C8EDC2FEEA41B2" box="[199,361,223,248]" italics="true" pageId="10" pageNumber="139">Pseudocalotes</emphasis>
<bibRefCitation id="911F4B3BAE71BB6A4960EDFDFE7141B2" author="Kaup" box="[367,498,224,248]" pageId="10" pageNumber="139" refString="Kaup, F. (1827) Zoologische Monographien. Isis von Oken, 20, 610 - 625, pls. 1 - 2." type="journal article" year="1827">Kaup, 1827</bibRefCitation>
</taxonomicName>
by the absence of rhombic dorsal scales
</paragraph>
<paragraph id="F53136CAAE71BB6A48C8EC19FB63402E" blockId="10.[151,1437,151,357]" pageId="10" pageNumber="139">
<emphasis id="C7FAEAD8AE71BB6A48C8EC19FE384057" bold="true" box="[199,443,260,285]" pageId="10" pageNumber="139">General description.</emphasis>
The
<typeStatus id="2A358868AE71BB6A49FBEC19FDD54057" box="[500,598,260,285]" pageId="10" pageNumber="139" type="holotype">holotype</typeStatus>
(see
<figureCitation id="6DB52A4FAE71BB6A4A9DEC19FD6D4056" box="[658,750,260,285]" captionStart="FIGURE 1" captionStartId="3.[151,250,1424,1446]" captionTargetBox="[153,1433,445,1403]" captionTargetId="figure@3.[153,1433,445,1403]" captionTargetPageId="3" captionText="FIGURE 1. Ventrolateral view of the head, gular and pectoral regions (“ Gonocephalus ” robinsonii holotype BMNH 1946.8.14.81)" httpUri="https://zenodo.org/record/234817/files/figure.png" pageId="10" pageNumber="139">Figure 1</figureCitation>
) has been described in
<bibRefCitation id="911F4B3BAE71BB6A4BF8EC19FB3D4056" author="Boulenger" box="[1015,1214,260,285]" pageId="10" pageNumber="139" refString="Boulenger, G. A. (1908) Report on the Gunong Tahan expedition, May - Sept. 1905. III. Fishes, batrachians and reptiles. Journal of the Federated Malay States Museums, 3, 61 - 69." type="journal article" year="1908">Boulenger (1908)</bibRefCitation>
and several aspects including variation were added by
<bibRefCitation id="911F4B3BAE71BB6A4A20EC3AFD65400A" author="Smedley" box="[559,742,295,320]" pageId="10" pageNumber="139" refString="Smedley, N. (1931) Amphibians and reptiles from the Cameron Highlands, Malay Peninsula. Bulletin of the Raffles Museum, 6, 105 - 123." type="journal article" year="1931">Smedley (1931)</bibRefCitation>
after additional material became available. A very detailed description of
<taxonomicName id="328E4D49AE71BB6A4937EC51FDD5402F" box="[312,598,332,357]" class="Reptilia" family="Agamidae" genus="Malayodracon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="139" phylum="Chordata" rank="species" species="robinsonii">
<emphasis id="C7FAEAD8AE71BB6A4937EC51FDD5402F" box="[312,598,332,357]" italics="true" pageId="10" pageNumber="139">Malayodracon robinsonii</emphasis>
</taxonomicName>
(as
<taxonomicName id="328E4D49AE71BB6A4A8AEC51FC23402F" box="[645,928,332,357]" class="Reptilia" family="Agamidae" genus="Gonocephalus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="139" phylum="Chordata" rank="species" species="robinsonii">
<emphasis id="C7FAEAD8AE71BB6A4A8AEC51FC23402F" box="[645,928,332,357]" italics="true" pageId="10" pageNumber="139">Gonocephalus robinsonii</emphasis>
</taxonomicName>
) is given in
<bibRefCitation id="911F4B3BAE71BB6A4C23EC51FB58402E" author="Grismer" box="[1068,1243,332,357]" pageId="10" pageNumber="139" refString="Grismer, L. L. (2011) Lizards of Peninsular Malaysia, Singapore and their adjacent archipelagos. Edition Chimaira, Frankfurt / M., 728 pp." type="book" year="2011">Grismer (2011)</bibRefCitation>
.
</paragraph>
<caption id="A1F16642AE71BB6A4898EA34FA9A4675" box="[151,1305,1833,1855]" httpUri="https://zenodo.org/record/234821/files/figure.png" pageId="10" pageNumber="139" targetBox="[340,1246,392,1811]" targetPageId="10">
<paragraph id="F53136CAAE71BB6A4898EA34FA9A4675" blockId="10.[151,1305,1833,1855]" box="[151,1305,1833,1855]" pageId="10" pageNumber="139">
<emphasis id="C7FAEAD8AE71BB6A4898EA34FE914674" bold="true" box="[151,274,1833,1855]" pageId="10" pageNumber="139">FIGURE 5.</emphasis>
<taxonomicName id="328E4D49AE71BB6A4916EA34FD984675" box="[281,539,1833,1855]" class="Reptilia" family="Agamidae" genus="Malayodracon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="139" phylum="Chordata" rank="species" species="robinsonii">
<emphasis id="C7FAEAD8AE71BB6A4916EA34FD984675" box="[281,539,1833,1855]" italics="true" pageId="10" pageNumber="139">Malayodracon robinsonii</emphasis>
</taxonomicName>
from Tanah Rata, Cameron Highlands, West Malaysia. Photo: U. Manthey.
</paragraph>
</caption>
<paragraph id="F53136CAAE71BB6A48C8EA76FA1E46BA" blockId="10.[151,1437,1899,2032]" pageId="10" pageNumber="139">Head scales small; scales on the supraciliary edge longer than wide; nasal in contact with first supralabial, sometimes also with second; typically 8 supra- and 8 infralabial scales (maximum 10 each). Slightly enlarged scales on the prominent v-shaped protrusion in the occipital region; large conical scale on either side of occiput. Gular pouch large, reaching the chest, tip rounded; gular scales small, homogeneous, smooth or slightly keeled.</paragraph>
<paragraph id="F53136CAAE70BB6B4898ED8AFD1340C2" blockId="11.[151,1437,151,1400]" pageId="11" pageNumber="140">Body elongated, laterally slightly compressed; dorsal scales nearly homogeneous, small, keeled, directed backwards and upwards intermixed with several—usually five or six—parallel oblique rows of enlarged scales. Scale numbers around midbody sometimes exceeding 100. Nuchal and dorsal crests continuous consisting of a single row of flat triangular—in some specimens lanceolate—scales touching at the base and decreasing in size caudad. Ventrals significantly larger than dorsals, keeled. Subdigital lamellae keeled, 31–36 on fourth toe. Tail long, roundish in cross-section. Enlarged scales along the mid ridge of the tail, continuous with dorsal crest. Underside of tail with strongly keeled scales.</paragraph>
<paragraph id="F53136CAAE70BB6B48C8EC89FBB543E2" blockId="11.[151,1437,151,1400]" pageId="11" pageNumber="140">
Adults in life typically dark green dorsally and dorsolaterally, dirty white ventrally; sometimes ground colour yellowish green (see
<figureCitation id="6DB52A4FAE70BB6B499AECAAFE78409A" box="[405,507,439,464]" captionStart="FIGURE 5" captionStartId="10.[151,250,1833,1855]" captionTargetBox="[340,1246,392,1811]" captionTargetId="figure@10.[340,1247,365,1811]" captionTargetPageId="10" captionText="FIGURE 5. Malayodracon robinsonii from Tanah Rata, Cameron Highlands, West Malaysia. Photo: U. Manthey." httpUri="https://zenodo.org/record/234821/files/figure.png" pageId="11" pageNumber="140">Figure 5</figureCitation>
). Gular sac typically reddish brown, greyish or black, continuously darkening towards the tip. A dark shoulder patch and dark oblique cross bands often visible. Enlarged dorsal scales (if present) often coloured much lighter than surrounding body colouration, forming bands. Labials and area below tympanum (plus tympanum) conspicuously white. Colour dependent upon age. Juveniles typically yellowish brown ground colour with brown cross bands. Labials in juveniles rather dirty white to pale brown, shoulder patch brown. In alcohol the colour fades and grey tones are prevalent (s. photograph of the
<typeStatus id="2A358868AE70BB6B4C6BEF70FB1743CF" box="[1124,1172,621,645]" pageId="11" pageNumber="140">type</typeStatus>
specimen in
<figureCitation id="6DB52A4FAE70BB6B4D23EF71FA0843CE" box="[1324,1419,620,645]" captionStart="FIGURE 1" captionStartId="3.[151,250,1424,1446]" captionTargetBox="[153,1433,445,1403]" captionTargetId="figure@3.[153,1433,445,1403]" captionTargetPageId="3" captionText="FIGURE 1. Ventrolateral view of the head, gular and pectoral regions (“ Gonocephalus ” robinsonii holotype BMNH 1946.8.14.81)" httpUri="https://zenodo.org/record/234817/files/figure.png" pageId="11" pageNumber="140">Figure 1</figureCitation>
). However, the black shoulder patch and dark dorsal cross bands remain discernible.
</paragraph>
</subSubSection>
<subSubSection id="BD946541AE70BB6B48C8EFA9FA1942CA" pageId="11" pageNumber="140" type="description">
<paragraph id="F53136CAAE70BB6B48C8EFA9FA1942CA" blockId="11.[151,1437,151,1400]" pageId="11" pageNumber="140">
<emphasis id="C7FAEAD8AE70BB6B48C8EFA9FEB84387" bold="true" box="[199,315,692,717]" pageId="11" pageNumber="140">Variation.</emphasis>
Hitherto known specimens from the
<typeStatus id="2A358868AE70BB6B4AFBEFA8FCA74387" box="[756,804,693,717]" pageId="11" pageNumber="140">type</typeStatus>
locality (Gunung Tahan) do not show enlarged dorsal scales arranged in oblique rows (
<bibRefCitation id="911F4B3BAE70BB6B4A18EFCAFD5143BA" author="Boulenger" box="[535,722,727,752]" pageId="11" pageNumber="140" refString="Boulenger, G. A. (1908) Report on the Gunong Tahan expedition, May - Sept. 1905. III. Fishes, batrachians and reptiles. Journal of the Federated Malay States Museums, 3, 61 - 69." type="journal article" year="1908">Boulenger 1908</bibRefCitation>
,
<bibRefCitation id="911F4B3BAE70BB6B4AEEEFCAFCCA43BA" author="Sly" box="[737,841,727,752]" pageId="11" pageNumber="140" refString="Sly, G. R. (1976) New locality records for some Peninsular Malaysia amphibia, reptilia and mammalia. Malayan Nature Journal, 29 (3), 155 - 157." type="journal article" year="1976">Sly 1976</bibRefCitation>
) as it can be seen in specimens from the
<collectingCountry id="8D99765AAE70BB6B4D3AEFC5FA1F43BA" box="[1333,1436,728,752]" name="Cameroon" pageId="11" pageNumber="140">Cameron</collectingCountry>
Highland region. It is conceivable that these two populations have been separated for a long time and constitute subspecies. However, in order to establish consistency of this character more material from the remote mountain ranges of central
<collectingCountry id="8D99765AAE70BB6B496AEE59FE4F4217" box="[357,460,836,861]" name="Malaysia" pageId="11" pageNumber="140">Malaysia</collectingCountry>
is needed. Additionally there exists a photographic record of a specimen from the
<collectingCountry id="8D99765AAE70BB6B4898EE75FF7D42CA" box="[151,254,872,896]" name="Cameroon" pageId="11" pageNumber="140">Cameron</collectingCountry>
Highlands without apparent enlarged scales across the dorsum rendering the above observation doubtful.
</paragraph>
</subSubSection>
<subSubSection id="BD946541AE70BB6B48C8EE91FF6045A2" pageId="11" pageNumber="140" type="distribution">
<paragraph id="F53136CAAE70BB6B48C8EE91FD124536" blockId="11.[151,1437,151,1400]" pageId="11" pageNumber="140">
<emphasis id="C7FAEAD8AE70BB6B48C8EE91FEE342EF" bold="true" box="[199,352,908,933]" pageId="11" pageNumber="140">Distribution.</emphasis>
<taxonomicName id="328E4D49AE70BB6B4966EE91FD0A42EF" box="[361,649,908,933]" class="Reptilia" family="Agamidae" genus="Malayodracon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="140" phylum="Chordata" rank="species" species="robinsonii">
<emphasis id="C7FAEAD8AE70BB6B4966EE91FD0A42EF" box="[361,649,908,933]" italics="true" pageId="11" pageNumber="140">Malayodracon robinsonii</emphasis>
</taxonomicName>
is restricted in its distribution from mid to high altitude areas of the peninsular Malaysian highlands (approx.
<quantity id="32769B2FAE70BB6B4A89EEB2FC98428D" box="[646,795,943,968]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.5" metricValueMin="0.6" pageId="11" pageNumber="140" unit="m" value="1050.0" valueMax="1500.0" valueMin="600.0">600–1500 m</quantity>
asl) inhabiting submontane and montane forests.
<taxonomicName id="328E4D49AE70BB6B4D72EEACFE8A42A7" class="Reptilia" family="Agamidae" genus="Malayodracon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="140" phylum="Chordata" rank="species" species="robinsonii">
<emphasis id="C7FAEAD8AE70BB6B4D72EEACFE8A42A7" italics="true" pageId="11" pageNumber="140">M. robinsonii</emphasis>
</taxonomicName>
shares this isolated distribution with several other reptile species that are restricted to the
<collectingCountry id="8D99765AAE70BB6B4D3AEEC9FA1F42A6" box="[1333,1436,980,1004]" name="Cameroon" pageId="11" pageNumber="140">Cameron</collectingCountry>
Highlands or the central mountain ranges of
<collectingCountry id="8D99765AAE70BB6B4AB8EEEAFC9E455A" box="[695,797,1015,1040]" name="Malaysia" pageId="11" pageNumber="140">Malaysia</collectingCountry>
(Fraser and Larut Hills; Genting Highlands) such as
<taxonomicName id="328E4D49AE70BB6B4898E900FCA4457E" authority="Vogel, David &amp; Pauwels, 2004" authorityName="Vogel, David &amp; Pauwels" authorityYear="2004" box="[151,807,1052,1077]" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="140" phylum="Chordata" rank="species" species="nebularis">
<emphasis id="C7FAEAD8AE70BB6B4898E900FE20457F" box="[151,419,1052,1077]" italics="true" pageId="11" pageNumber="140">Trimeresurus nebularis</emphasis>
<bibRefCitation id="911F4B3BAE70BB6B49BFE901FCA4457E" author="Vogel" box="[432,807,1052,1077]" pageId="11" pageNumber="140" refString="Vogel, G., David, P. &amp; Pauwels, O. S. G. (2004) A review of morphological variation in Trimeresurus popeiorum (Serpentes: Viperidae: Crotalinae), with the description of two new species. Zootaxa, 727, 1 - 63." type="journal article" year="2004">Vogel, David &amp; Pauwels, 2004</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="328E4D49AE70BB6B4B36E901FB60457E" authority="Smedley, 1931" authorityName="Smedley" authorityYear="1931" box="[825,1251,1052,1077]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="140" phylum="Chordata" rank="species" species="sanguineus">
<emphasis id="C7FAEAD8AE70BB6B4B36E901FB97457E" box="[825,1044,1052,1077]" italics="true" pageId="11" pageNumber="140">Hebius sanguineus</emphasis>
(
<bibRefCitation id="911F4B3BAE70BB6B4C26E901FB59457E" author="Smedley" box="[1065,1242,1052,1077]" pageId="11" pageNumber="140" refString="Smedley, N. (1931) Amphibians and reptiles from the Cameron Highlands, Malay Peninsula. Bulletin of the Raffles Museum, 6, 105 - 123." type="journal article" year="1931">Smedley, 1931</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="328E4D49AE70BB6B4CFAE901FEF94512" authority="Lim, 1963" authorityName="Lim" authorityYear="1963" class="Reptilia" family="Colubridae" genus="Macrocalamus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="140" phylum="Chordata" rank="species" species="tweediei">
<emphasis id="C7FAEAD8AE70BB6B4CFAE901FF774512" italics="true" pageId="11" pageNumber="140">Macrocalamus tweediei</emphasis>
<bibRefCitation id="911F4B3BAE70BB6B490FE922FEF94512" author="Lim" box="[256,378,1087,1112]" pageId="11" pageNumber="140" refString="Lim, B. L. (1963) Macrocalamus tweediei, a new species of Reed Snake from Malaya. Bulletin of the National Museum Singapore, 32, 99 - 102." type="journal article" year="1963">Lim, 1963</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="328E4D49AE70BB6B49B5E922FC2E4512" authority="Smedley, 1931" authorityName="Smedley" authorityYear="1931" box="[442,941,1087,1112]" class="Reptilia" family="Colubridae" genus="Collorhabdium" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="140" phylum="Chordata" rank="species" species="williamsoni">
<emphasis id="C7FAEAD8AE70BB6B49B5E922FD704512" box="[442,755,1087,1112]" italics="true" pageId="11" pageNumber="140">Collorhabdium williamsoni</emphasis>
<bibRefCitation id="911F4B3BAE70BB6B4AF0E922FC2E4512" author="Smedley" box="[767,941,1087,1112]" pageId="11" pageNumber="140" refString="Smedley, N. (1931) Amphibians and reptiles from the Cameron Highlands, Malay Peninsula. Bulletin of the Raffles Museum, 6, 105 - 123." type="journal article" year="1931">Smedley, 1931</bibRefCitation>
</taxonomicName>
. Endemism on genus and species level is comparatively high in this geographical area.
</paragraph>
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<taxonomicName id="328E4D49AE70BB6B48C8E99AFE6545EA" box="[199,486,1159,1184]" class="Reptilia" family="Agamidae" genus="Malayodracon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="140" phylum="Chordata" rank="species" species="robinsonii">
<emphasis id="C7FAEAD8AE70BB6B48C8E99AFE6545EA" box="[199,486,1159,1184]" italics="true" pageId="11" pageNumber="140">Malayodracon robinsonii</emphasis>
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has been reported from Genting Highlands (Ulu Kali),
<collectingCountry id="8D99765AAE70BB6B4C6BE995FB4845EA" box="[1124,1227,1160,1184]" name="Cameroon" pageId="11" pageNumber="140">Cameron</collectingCountry>
Highlands (Tanah Rata, Gunung Brinchang, Gunung Beremban, Gunung Jesar) and further north from its
<typeStatus id="2A358868AE70BB6B4CA9E9B0FB55458F" box="[1190,1238,1197,1221]" pageId="11" pageNumber="140">type</typeStatus>
locality Gunung Tahan.
</paragraph>
</subSubSection>
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<emphasis id="C7FAEAD8AE70BB6B48C8E9E9FECD4447" bold="true" box="[199,334,1268,1293]" pageId="11" pageNumber="140">Etymology.</emphasis>
The name
<taxonomicName id="328E4D49AE70BB6B49C5E9E9FDEC4447" box="[458,623,1268,1293]" class="Reptilia" family="Agamidae" genus="Malayodracon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="137" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="C7FAEAD8AE70BB6B49C5E9E9FDEC4447" box="[458,623,1268,1293]" italics="true" pageId="11" pageNumber="140">Malayodracon</emphasis>
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was chosen to express that the
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and currently only known species of the genus is restricted to
<collectingCountry id="8D99765AAE70BB6B49A2E80AFD90447A" box="[429,531,1303,1328]" name="Malaysia" pageId="11" pageNumber="140">Malaysia</collectingCountry>
(latinized malaya, male form malayo owing to the gender of the ending -dracon) and constitutes a genus belonging to the subfamily
<taxonomicName id="328E4D49AE70BB6B4ADFE821FCD2441F" box="[720,849,1340,1365]" class="Reptilia" family="Agamidae" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="140" phylum="Chordata" rank="subFamily" subFamily="Draconinae">Draconinae</taxonomicName>
(gr. drakon / δράκων; a serpent in Greek mythology; latinized dracon = engl. dragon).
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<emphasis id="C7FAEAD8AE73BB68493AEA13FDDE466D" box="[309,605,1806,1831]" italics="true" pageId="8" pageNumber="137">Gonyocephalus robinsonii</emphasis>
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</taxonomicName>
;
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locality: Gunong (=Gunung / Mt.) Tahan (
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1946.8.14.81 (previously
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1906.2.28.8); collected by H.C. Robinson.
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170
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<collectingCountry id="8D990FEEFFA9E7070A35FA90FC65FA97" box="[821,976,1316,1339]" name="Colombia" pageId="0" pageNumber="61">COLOMBIA</collectingCountry>
, Nariño, Tumaco, Finca Mar Agrícola, Universidad de Nariño,
<quantity id="3276E29BFFA9E7070898FAF0FE6BFAF7" box="[408,478,1348,1371]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="3.0" metricValueMin="0.0" pageId="0" pageNumber="61" unit="m" value="1.5" valueMax="3.0" valueMin="0.0">0–3 m</quantity>
, in a pond,
<date id="813069BEFFA9E7070B5BFAF0FD59FAF7" box="[603,748,1348,1371]" pageId="0" pageNumber="61" value="2010-06-22">22 June 2010</date>
, leg. D. Padilla, (
<collectionCode id="939FD7BBFFA9E7070AA6FAF0FC6CFAF7" box="[934,985,1348,1371]" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="0" pageNumber="61">ICN</collectionCode>
).
<typeStatus id="2A35F1DCFFA9E7070AEBFAF0FB34FAF7" box="[1003,1153,1348,1371]" pageId="0" pageNumber="61" type="paratype">PARATYPES</typeStatus>
, same data as
<typeStatus id="2A35F1DCFFA9E7070C1BFAF0FACEFAF7" box="[1307,1403,1348,1371]" pageId="0" pageNumber="61" type="holotype">holotype</typeStatus>
, of these the following have been distributed: 1 brach M, 1 brach F (
<collectionCode id="939FD7BBFFA9E7070A4AFAD0FCC8FAD7" box="[842,893,1380,1403]" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="0" pageNumber="61">ICN</collectionCode>
), 1 macr M, 1 macr F (PSO-CZ).
</paragraph>
</subSubSection>
<subSubSection id="BD941CF5FFA9E70609C5FA37FCD4FEA3" lastPageId="1" lastPageNumber="62" pageId="0" pageNumber="61" type="description">
<paragraph id="F5314F7EFFA9E70709C5FA37FED5FA77" blockId="0.[151,1437,1315,2011]" pageId="0" pageNumber="61">
<emphasis id="C7FA936CFFA9E70709C5FA37FEE7FA37" bold="true" box="[197,338,1411,1435]" pageId="0" pageNumber="61">Description.</emphasis>
Macropterous form. Dimensions. Length M 7.60−7.86, F 7.60−7.70; greatest body width M 1.88; F 2.16. Brachypterous form, length M 7.0, F 6.9; greatest body width M 1.76; F 2.08. Width of head M (all) and F macr, 1.62, F brach 1.56.
</paragraph>
<paragraph id="F5314F7EFFA9E70709C5FA50FDF2F9F7" blockId="0.[151,1437,1315,2011]" pageId="0" pageNumber="61">Color. General body bright white, dark eyes. Pleura, inner margin with a fringe of short and black hair. Abdominal venter black except keel and portions of connexivum, these yellow; abdominal dorsum red with irregular areas of black and black apex. Along median longitudinal axis of head, pronotum, and hemelytra covered with thin, long, transparent setae. Membrane, however, not covered.</paragraph>
<paragraph id="F5314F7EFFA9E70609C5F9D0FE7AFF63" blockId="0.[151,1437,1315,2011]" lastBlockId="1.[151,1437,152,623]" lastPageId="1" lastPageNumber="62" pageId="0" pageNumber="61">
Male structural characteristics: As viewed from above, outline of head rounded with anterior margin of vertex continuous with that of eyes. Also, vertex slightly indented. Width of head 4.7 times anterior width of vertex (1.62/0.34) and smaller than humeral width of pronotum; synthlipsis 1.8 times anterior width of vertex (0.18/0.34); along median longitudinal axis, head is 2.2 times shorter than that of pronotum; tylus inflated, and with thin, long, transparent setae; labrum’s width twice its length, displaying a tuft of transparent, hair, and a truncate apex; rostral prong (
<figureCitation id="6DB553FBFFA9E7070C06F950FAF0F957" box="[1286,1349,1764,1787]" captionStart="FIGURES 1 − 6" captionStartId="1.[151,272,1952,1976]" captionTargetBox="[154,1416,689,1906]" captionTargetId="figure@1.[151,1436,677,1929]" captionTargetPageId="1" captionText="FIGURES 1 − 6. Buenoa uselus sp. n. male. 1 Rostrum in lateral view; 2 Fore leg; 3 Femoral comb (400 X); 4 Tibial comb (400 X); 5 Preapical structure of fore tarsus II; 6 Dorsal apex of middle tibia." httpUri="https://zenodo.org/record/198115/files/figure.png" pageId="0" pageNumber="61">Fig. 1</figureCitation>
) longer than third rostral segment, with base originating laterally and protruding anteriorly at distal end of third rostral segment, rounded apex. Pronotum tricarinate; lateral margins slightly divergent; posterior margin convex, medially concave; median length 1.3 times its width (1.72/1.3). Scutellum short, pronotum with median length 1.6 times median length of scutellum. Fore femur wide, somewhat thickened basally and slightly thinner apically (
<figureCitation id="6DB553FBFFA9E7070D34F8D0FBC3F8D7" box="[1076,1142,1892,1915]" captionStart="FIGURES 1 − 6" captionStartId="1.[151,272,1952,1976]" captionTargetBox="[154,1416,689,1906]" captionTargetId="figure@1.[151,1436,677,1929]" captionTargetPageId="1" captionText="FIGURES 1 − 6. Buenoa uselus sp. n. male. 1 Rostrum in lateral view; 2 Fore leg; 3 Femoral comb (400 X); 4 Tibial comb (400 X); 5 Preapical structure of fore tarsus II; 6 Dorsal apex of middle tibia." httpUri="https://zenodo.org/record/198115/files/figure.png" pageId="0" pageNumber="61">Fig. 2</figureCitation>
); stridulatory area with 17– 20 weak ridges, with apical small and irregular teeth (
<figureCitation id="6DB553FBFFA9E7070BDEF830FC95F837" box="[734,800,1924,1947]" captionStart="FIGURES 1 − 6" captionStartId="1.[151,272,1952,1976]" captionTargetBox="[154,1416,689,1906]" captionTargetId="figure@1.[151,1436,677,1929]" captionTargetPageId="1" captionText="FIGURES 1 − 6. Buenoa uselus sp. n. male. 1 Rostrum in lateral view; 2 Fore leg; 3 Femoral comb (400 X); 4 Tibial comb (400 X); 5 Preapical structure of fore tarsus II; 6 Dorsal apex of middle tibia." httpUri="https://zenodo.org/record/198115/files/figure.png" pageId="0" pageNumber="61">Fig. 3</figureCitation>
); dorsal and ventral to stridulatory area with parallel striae visible. Fore tibia with stridulatory comb consisting of approximately 40 teeth (
<figureCitation id="6DB553FBFFA9E7070AE0F810FB94F817" box="[992,1057,1956,1979]" captionStart="FIGURES 1 − 6" captionStartId="1.[151,272,1952,1976]" captionTargetBox="[154,1416,689,1906]" captionTargetId="figure@1.[151,1436,677,1929]" captionTargetPageId="1" captionText="FIGURES 1 − 6. Buenoa uselus sp. n. male. 1 Rostrum in lateral view; 2 Fore leg; 3 Femoral comb (400 X); 4 Tibial comb (400 X); 5 Preapical structure of fore tarsus II; 6 Dorsal apex of middle tibia." httpUri="https://zenodo.org/record/198115/files/figure.png" pageId="0" pageNumber="61">Fig. 4</figureCitation>
), with 10 long basal teeth, followed by ~30 shorter teeth. Fore and middle legs have dark preapical structure in tarsus II (
<figureCitation id="6DB553FBFFA9E7070D21F870FBD7F877" box="[1057,1122,1988,2011]" captionStart="FIGURES 1 − 6" captionStartId="1.[151,272,1952,1976]" captionTargetBox="[154,1416,689,1906]" captionTargetId="figure@1.[151,1436,677,1929]" captionTargetPageId="1" captionText="FIGURES 1 − 6. Buenoa uselus sp. n. male. 1 Rostrum in lateral view; 2 Fore leg; 3 Femoral comb (400 X); 4 Tibial comb (400 X); 5 Preapical structure of fore tarsus II; 6 Dorsal apex of middle tibia." httpUri="https://zenodo.org/record/198115/files/figure.png" pageId="0" pageNumber="61">Fig. 5</figureCitation>
). Dorsal apex of middle tibia with two rows of four transparent setae (
<figureCitation id="6DB553FBFFA8E7060B48FF2CFD3FFF03" box="[584,650,152,175]" captionStart="FIGURES 1 − 6" captionStartId="1.[151,272,1952,1976]" captionTargetBox="[154,1416,689,1906]" captionTargetId="figure@1.[151,1436,677,1929]" captionTargetPageId="1" captionText="FIGURES 1 − 6. Buenoa uselus sp. n. male. 1 Rostrum in lateral view; 2 Fore leg; 3 Femoral comb (400 X); 4 Tibial comb (400 X); 5 Preapical structure of fore tarsus II; 6 Dorsal apex of middle tibia." httpUri="https://zenodo.org/record/198115/files/figure.png" pageId="1" pageNumber="62">Fig. 6</figureCitation>
). Male genital claspers normal. Spine from caudo-sinistral margin curved and very narrow 2/3 apically.
</paragraph>
<paragraph id="F5314F7EFFA8E70609C5FF6CFCD4FEA3" blockId="1.[151,1437,152,623]" pageId="1" pageNumber="62">Female structural characteristics: female has the setae characteristic of the middle tibiae and the preapical structure in tarsus II. Also, hind femur has 25 spines along posterior margin.</paragraph>
</subSubSection>
<subSubSection id="BD941CF5FFA8E70609C5FEA3FD0FFDC3" pageId="1" pageNumber="62" type="etymology">
<paragraph id="F5314F7EFFA8E70609C5FEA3FD41FE83" blockId="1.[151,1437,152,623]" box="[197,756,279,303]" pageId="1" pageNumber="62">
<emphasis id="C7FA936CFFA8E70609C5FEA3FEFCFE83" bold="true" box="[197,329,279,303]" pageId="1" pageNumber="62">Etymology:</emphasis>
<taxonomicName id="328E34FDFFA8E706084FFEACFE2EFE83" box="[335,411,280,303]" class="Insecta" family="Notonectidae" genus="Buenoa" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="uselus" status="sp. nov.">Uselus</taxonomicName>
. A Greek noun meaning “lofty.”
</paragraph>
<paragraph id="F5314F7EFFA8E70609C5FE83FDA3FE03" blockId="1.[151,1437,152,623]" pageId="1" pageNumber="62">
<emphasis id="C7FA936CFFA8E70609C5FE83FE10FEE3" bold="true" box="[197,421,311,335]" pageId="1" pageNumber="62">Comparative notes:</emphasis>
<taxonomicName id="328E34FDFFA8E70608ABFE8CFDBEFEE3" box="[427,523,312,335]" class="Insecta" family="Notonectidae" genus="Buenoa" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="uselus">
<emphasis id="C7FA936CFFA8E70608ABFE8CFDBEFEE3" box="[427,523,312,335]" italics="true" pageId="1" pageNumber="62">B. uselus</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="328E34FDFFA8E7060B95FE8CFC25FEE3" authority="Fieber" authorityName="Fieber" box="[661,912,312,335]" class="Insecta" family="Notonectidae" genus="Buenoa" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="platycnemis">
<emphasis id="C7FA936CFFA8E7060B95FE8CFD10FEE3" box="[661,677,312,335]" italics="true" pageId="1" pageNumber="62">B</emphasis>
.
<emphasis id="C7FA936CFFA8E7060BB3FE8CFC87FEE3" box="[691,818,312,335]" italics="true" pageId="1" pageNumber="62">platycnemis</emphasis>
(Fieber)
</taxonomicName>
, but
<taxonomicName id="328E34FDFFA8E7060AC4FE8CFBD5FEE3" box="[964,1120,312,335]" class="Insecta" family="Notonectidae" genus="Buenoa" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="platycnemis">
<emphasis id="C7FA936CFFA8E7060AC4FE8CFC61FEE3" box="[964,980,312,335]" italics="true" pageId="1" pageNumber="62">B</emphasis>
.
<emphasis id="C7FA936CFFA8E7060AE1FE8CFBD5FEE3" box="[993,1120,312,335]" italics="true" pageId="1" pageNumber="62">platycnemis</emphasis>
</taxonomicName>
is smaller (length M 4.6−5.7, F 4.9−5.1), its measures morphometric are different, the fore femur has 11–14 sclerotized ridges and it also lacks the preapical structure in the anterior and middle tarsus II.
<taxonomicName id="328E34FDFFA8E7060BDFFECCFCF5FE23" box="[735,832,376,399]" class="Insecta" family="Notonectidae" genus="Buenoa" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="uselus">
<emphasis id="C7FA936CFFA8E7060BDFFECCFCF5FE23" box="[735,832,376,399]" italics="true" pageId="1" pageNumber="62">B. uselus</emphasis>
</taxonomicName>
, male and female have long setae on the outer surface of the middle tibia at the apex (
<figureCitation id="6DB553FBFFA8E70608C7FE2CFDBCFE03" box="[455,521,408,431]" captionStart="FIGURES 1 − 6" captionStartId="1.[151,272,1952,1976]" captionTargetBox="[154,1416,689,1906]" captionTargetId="figure@1.[151,1436,677,1929]" captionTargetPageId="1" captionText="FIGURES 1 − 6. Buenoa uselus sp. n. male. 1 Rostrum in lateral view; 2 Fore leg; 3 Femoral comb (400 X); 4 Tibial comb (400 X); 5 Preapical structure of fore tarsus II; 6 Dorsal apex of middle tibia." httpUri="https://zenodo.org/record/198115/files/figure.png" pageId="1" pageNumber="62">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="F5314F7EFFA8E70609C5FE03FBFFFDA3" blockId="1.[151,1437,152,623]" pageId="1" pageNumber="62">
<emphasis id="C7FA936CFFA8E70609C5FE03FE9CFE63" bold="true" box="[197,297,439,463]" pageId="1" pageNumber="62">Habitat.</emphasis>
The specimens were collected from a permanent lake of more than a meter deep, with surrounding vegetation and open waters; tropical humid forest encompasses the lake. The habitat is characterized by air temperature 32 ºC, water temperature 26 ºC, pH 7, conductivity 621 S/m, and dissolved oxygen 50%.
</paragraph>
<paragraph id="F5314F7EFFA8E70609C5FDACFD0FFDC3" blockId="1.[151,1437,152,623]" pageId="1" pageNumber="62">
<taxonomicName id="328E34FDFFA8E70609C5FDACFE9BFD83" box="[197,302,536,559]" class="Insecta" family="Notonectidae" genus="Buenoa" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="uselus" status="sp. nov.">
<emphasis id="C7FA936CFFA8E70609C5FDACFE9BFD83" box="[197,302,536,559]" italics="true" pageId="1" pageNumber="62">B. uselus</emphasis>
</taxonomicName>
<emphasis id="C7FA936CFFA8E7060836FDA3FECCFD83" bold="true" box="[310,377,535,559]" pageId="1" pageNumber="62">
<taxonomicNameLabel id="DCC92E17FFA8E7060836FDA3FECCFD83" box="[310,377,535,559]" pageId="1" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is found with
<taxonomicName id="328E34FDFFA8E7060B23FDACFC6AFD83" authority="Westwood, 1834" authorityName="Westwood" authorityYear="1834" box="[547,991,536,559]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="pulchella">
<emphasis id="C7FA936CFFA8E7060B23FDACFCA1FD83" box="[547,788,536,559]" italics="true" pageId="1" pageNumber="62">Microvelia pulchella</emphasis>
Westwood, 1834
</taxonomicName>
,
<taxonomicName id="328E34FDFFA8E7060AECFDACFAD2FD83" authority="Uhler, 1894" authorityName="Uhler" authorityYear="1894" box="[1004,1383,536,559]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="C7FA936CFFA8E7060AECFDACFB66FD83" box="[1004,1235,536,559]" italics="true" pageId="1" pageNumber="62">Microvelia longipes</emphasis>
Uhler, 1894
</taxonomicName>
, and
<taxonomicName id="328E34FDFFA8E7060997FD8CFD9DFDE3" authority="Kirkaldy, 1899" authorityName="Kirkaldy" authorityYear="1899" box="[151,552,568,591]" class="Insecta" family="Gerridae" genus="Trepobates" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="taylori">
<emphasis id="C7FA936CFFA8E7060997FD8CFED6FDE3" box="[151,355,568,591]" italics="true" pageId="1" pageNumber="62">Trepobates taylori</emphasis>
(Kirkaldy, 1899)
</taxonomicName>
; in another lake with
<taxonomicName id="328E34FDFFA8E7060A1CFD8CFAB2FDE3" authority="Drake &amp; Harris, 1933" authorityName="Drake &amp; Harris" authorityYear="1933" box="[796,1287,568,591]" class="Insecta" family="Gerridae" genus="Limnogonus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="aduncus">
<emphasis id="C7FA936CFFA8E7060A1CFD8CFBBCFDE3" box="[796,1033,568,591]" italics="true" pageId="1" pageNumber="62">Limnogonus aduncus</emphasis>
Drake &amp; Harris, 1933
</taxonomicName>
,
<taxonomicName id="328E34FDFFA8E7060C14FD8CFE10FDC3" authority="Stal, 1859" authorityName="Stal" authorityYear="1859" class="Insecta" family="Gerridae" genus="Limnogonus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="franciscanus">
<emphasis id="C7FA936CFFA8E7060C14FD8CFEAAFDC3" italics="true" pageId="1" pageNumber="62">Limnogonus franciscanus</emphasis>
(Stål, 1859)
</taxonomicName>
, and
<taxonomicName id="328E34FDFFA8E70608DDFDECFD07FDC3" box="[477,690,600,623]" class="Insecta" family="Veliidae" genus="Microvelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="C7FA936CFFA8E70608DDFDECFD07FDC3" box="[477,690,600,623]" italics="true" pageId="1" pageNumber="62">Microvelia longipes</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

294
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Key to determine the present species of genus
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<emphasis id="C7FA936CFFABE7050BA1FF2CFD42FF1C" bold="true" box="[673,759,152,176]" italics="true" pageId="2" pageNumber="63">Buenoa</emphasis>
</taxonomicName>
from
<collectingCountry id="8D990FEEFFABE7050A3CFF23FC1EFF03" box="[828,939,151,175]" name="Colombia" pageId="2" pageNumber="63">Colombia</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="F5314F7EFFABE7050997FF0CFEBBFF63" blockId="2.[151,939,151,207]" box="[151,270,184,207]" pageId="2" pageNumber="63">Males only</paragraph>
</subSubSection>
<subSubSection id="BD941CF5FFABE7050997FF4CFA29F823" pageId="2" pageNumber="63" type="key">
<keyStep id="4E7A57DBFFABE7050997FF4CFA2EFE83" pageId="2" pageNumber="63">
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<keyLead id="4E7FEC4BFFABE7050997FF4CFA2EFEA3" box="[151,1435,248,271]" pageId="2" pageNumber="63">1 Fore femur with stridulatory area present. Fore tibia without parallel margins ........................................................... 2</keyLead>
</paragraph>
<paragraph id="F5314F7EFFABE7050997FEACFA2EFE83" blockId="2.[151,1436,248,1935]" box="[151,1435,280,303]" pageId="2" pageNumber="63">
<keyLead id="4E7FEC4BFFABE7050997FEACFA2EFE83" box="[151,1435,280,303]" pageId="2" pageNumber="63">- Fore femur with stridulatory area absent. Fore tibia with almost parallel margins .................................................... 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="4E7A57DBFFABE7050997FE8CFA20FE03" pageId="2" pageNumber="63">
<paragraph id="F5314F7EFFABE7050997FE8CFA29FE23" blockId="2.[151,1436,248,1935]" pageId="2" pageNumber="63">
<keyLead id="4E7FEC4BFFABE7050997FE8CFA29FE23" pageId="2" pageNumber="63">
2 Stout species, male L/W: 8.5/2.32; rostral prong longer than third rostral segment, with base originating laterally and protruding anteriorly at distal end of third rostral segment; fore femur has 15 sclerotized ridges, fore tibiae 30 teeth (see Padilla-Gil, 2010c,
<figureCitation id="6DB553FBFFABE70508C9FECCFDBBFE23" box="[457,526,376,399]" captionStart="FIGURES 1 − 6" captionStartId="1.[151,272,1952,1976]" captionTargetBox="[154,1416,689,1906]" captionTargetId="figure@1.[151,1436,677,1929]" captionTargetPageId="1" captionText="FIGURES 1 − 6. Buenoa uselus sp. n. male. 1 Rostrum in lateral view; 2 Fore leg; 3 Femoral comb (400 X); 4 Tibial comb (400 X); 5 Preapical structure of fore tarsus II; 6 Dorsal apex of middle tibia." httpUri="https://zenodo.org/record/198115/files/figure.png" pageId="2" pageNumber="63">Figs. 1</figureCitation>
–7)
<emphasis id="C7FA936CFFABE7050B36FECCFA82FE23" box="[566,1335,376,399]" italics="true" pageId="2" pageNumber="63">
................................................................................................................
<taxonomicName id="328E34FDFFABE7050C27FECCFA82FE23" ID-CoL="NNSP" box="[1319,1335,376,399]" class="Insecta" family="Notonectidae" genus="Buenoa" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="funensis">B</taxonomicName>
</emphasis>
.
<emphasis id="C7FA936CFFABE7050C44FECCFA29FE23" box="[1348,1436,376,399]" italics="true" pageId="2" pageNumber="63">funensis</emphasis>
</keyLead>
</paragraph>
<paragraph id="F5314F7EFFABE7050997FE2CFA20FE03" blockId="2.[151,1436,248,1935]" box="[151,1429,408,431]" pageId="2" pageNumber="63">
<keyLead id="4E7FEC4BFFABE7050997FE2CFA20FE03" box="[151,1429,408,431]" pageId="2" pageNumber="63">- Slender species, male L less than 8.2, W less than 2 ...................................................................................................3</keyLead>
</paragraph>
</keyStep>
<keyStep id="4E7A57DBFFABE7050997FE0CFA29FE43" pageId="2" pageNumber="63">
<paragraph id="F5314F7EFFABE7050997FE0CFA2EFE63" blockId="2.[151,1436,248,1935]" box="[151,1435,440,463]" pageId="2" pageNumber="63">
<keyLead id="4E7FEC4BFFABE7050997FE0CFA2EFE63" box="[151,1435,440,463]" pageId="2" pageNumber="63">3. Synthlipsis longer or equal than two-fifths anterior width of vertex............................................................................ 7</keyLead>
</paragraph>
<paragraph id="F5314F7EFFABE7050997FE6CFA29FE43" blockId="2.[151,1436,248,1935]" box="[151,1436,472,495]" pageId="2" pageNumber="63">
<keyLead id="4E7FEC4BFFABE7050997FE6CFA29FE43" box="[151,1436,472,495]" pageId="2" pageNumber="63">- Synthlipsis shorter than two-fifths anterior width of vertex .........................................................................................4</keyLead>
</paragraph>
</keyStep>
<keyStep id="4E7A57DBFFABE7050997FE4CFA29FD83" pageId="2" pageNumber="63">
<paragraph id="F5314F7EFFABE7050997FE4CFA2EFDA3" blockId="2.[151,1436,248,1935]" box="[151,1435,504,527]" pageId="2" pageNumber="63">
<keyLead id="4E7FEC4BFFABE7050997FE4CFA2EFDA3" box="[151,1435,504,527]" pageId="2" pageNumber="63">4. Synthlipsis four times anterior width of vertex ............................................................................................................ 5</keyLead>
</paragraph>
<paragraph id="F5314F7EFFABE7050997FDACFA29FD83" blockId="2.[151,1436,248,1935]" box="[151,1436,536,559]" pageId="2" pageNumber="63">
<keyLead id="4E7FEC4BFFABE7050997FDACFA29FD83" box="[151,1436,536,559]" pageId="2" pageNumber="63">- Synthlipsis three times anterior width of vertex ...........................................................................................................6</keyLead>
</paragraph>
</keyStep>
<keyStep id="4E7A57DBFFABE7050997FD8CFA2EFD03" pageId="2" pageNumber="63">
<paragraph id="F5314F7EFFABE7050997FD8CFA29FDC3" blockId="2.[151,1436,248,1935]" pageId="2" pageNumber="63">
<keyLead id="4E7FEC4BFFABE7050997FD8CFA29FDC3" pageId="2" pageNumber="63">
5 Smaller species, male L 5.3–6.8. Fore tibia with 3–4 peg-like setae on inner surface near to apex. Fore and middle legs lacking preapical structure of tarsus II
<emphasis id="C7FA936CFFABE7050B93FDECFA29FDC3" box="[659,1436,600,623]" italics="true" pageId="2" pageNumber="63">
...................................................................................................
<taxonomicName id="328E34FDFFABE7050C2DFDECFA29FDC3" ID-CoL="NNSR" box="[1325,1436,600,623]" class="Insecta" family="Notonectidae" genus="Buenoa" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="gracilis">B. gracilis</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F5314F7EFFABE7050997FDCCFA2EFD03" blockId="2.[151,1436,248,1935]" pageId="2" pageNumber="63">
<keyLead id="4E7FEC4BFFABE7050997FDCCFA2EFD03" pageId="2" pageNumber="63">
- Larger species, male L 8.0. Fore tibia lacking setae on inner surface close to apex. Fore and middle legs with preapical structure of tarsus II (see
<bibRefCitation id="911F328FFFABE70508FCFD2CFD76FD03" author="Padilla-Gil" box="[508,707,664,687]" pageId="2" pageNumber="63" refString="Padilla-Gil, D. N. (2010 a) Five new species of Buenoa (Hemiptera: Heteroptera: Notonectidae) from Colombia. Zootaxa, 2411, 22 - 32." type="journal article" year="2010" yearSuffix="a">Padilla-Gil, 2010a</bibRefCitation>
, fig. 3e) .........................................................................
<taxonomicName id="328E34FDFFABE7050C10FD2CFA2EFD03" ID-CoL="NNTK" box="[1296,1435,664,687]" class="Insecta" family="Notonectidae" genus="Buenoa" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="prosthetus">
<emphasis id="C7FA936CFFABE7050C10FD2CFA2EFD03" box="[1296,1435,664,687]" italics="true" pageId="2" pageNumber="63">B. prosthetus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="4E7A57DBFFABE7050997FD0CFA29FC83" pageId="2" pageNumber="63">
<paragraph id="F5314F7EFFABE7050997FD0CFA29FD43" blockId="2.[151,1436,248,1935]" pageId="2" pageNumber="63">
<keyLead id="4E7FEC4BFFABE7050997FD0CFA29FD43" pageId="2" pageNumber="63">
6 Smaller species, male L 5.9. Fore tibia lacking setae on outer surface close to apex. Fore and middle legs lacking preapical structure of tarsus II
<emphasis id="C7FA936CFFABE7050B07FD6CFA29FD43" box="[519,1436,728,751]" italics="true" pageId="2" pageNumber="63">
......................................................................................................................
<taxonomicName id="328E34FDFFABE7050C1EFD6CFA29FD43" ID-CoL="694KD" box="[1310,1436,728,751]" class="Insecta" family="Notonectidae" genus="Buenoa" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="amazona">B. amazona</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F5314F7EFFABE7050997FD4CFA29FC83" blockId="2.[151,1436,248,1935]" pageId="2" pageNumber="63">
<keyLead id="4E7FEC4BFFABE7050997FD4CFA29FC83" pageId="2" pageNumber="63">
- Larger species, male L 8.0. Fore tibia with three short, peg-like setae on outer surface close to apex. Fore and middle legs with preapical structure of tarsus II (see
<bibRefCitation id="911F328FFFABE7050BA7FCACFCDFFC83" author="Padilla-Gil" box="[679,874,792,815]" pageId="2" pageNumber="63" refString="Padilla-Gil, D. N. (2010 a) Five new species of Buenoa (Hemiptera: Heteroptera: Notonectidae) from Colombia. Zootaxa, 2411, 22 - 32." type="journal article" year="2010" yearSuffix="a">Padilla-Gil, 2010a</bibRefCitation>
, fig. 3d–e).................................................
<taxonomicName id="328E34FDFFABE7050C2BFCACFA29FC83" ID-CoL="NNSC" box="[1323,1436,792,815]" class="Insecta" family="Notonectidae" genus="Buenoa" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="dactylis">
<emphasis id="C7FA936CFFABE7050C2BFCACFA8EFC83" box="[1323,1339,792,815]" italics="true" pageId="2" pageNumber="63">B</emphasis>
.
<emphasis id="C7FA936CFFABE7050C49FCACFA29FC83" box="[1353,1436,792,815]" italics="true" pageId="2" pageNumber="63">dactylis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="4E7A57DBFFABE7050997FC8CFA2EFCC3" pageId="2" pageNumber="63">
<paragraph id="F5314F7EFFABE7050997FC8CFA2EFCE3" blockId="2.[151,1436,248,1935]" box="[151,1435,824,847]" pageId="2" pageNumber="63">
<keyLead id="4E7FEC4BFFABE7050997FC8CFA2EFCE3" box="[151,1435,824,847]" pageId="2" pageNumber="63">7 Stridulatory comb of fore tibia with less than 21 teeth................................................................................................. 8</keyLead>
</paragraph>
<paragraph id="F5314F7EFFABE7050997FCECFA2EFCC3" blockId="2.[151,1436,248,1935]" box="[151,1435,856,879]" pageId="2" pageNumber="63">
<keyLead id="4E7FEC4BFFABE7050997FCECFA2EFCC3" box="[151,1435,856,879]" pageId="2" pageNumber="63">- Stridulatory comb of fore tibia with more than 22 teeth............................................................................................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="4E7A57DBFFABE7050997FCCCFA29FC43" pageId="2" pageNumber="63">
<paragraph id="F5314F7EFFABE7050997FCCCFA29FC03" blockId="2.[151,1436,248,1935]" pageId="2" pageNumber="63">
<keyLead id="4E7FEC4BFFABE7050997FCCCFA29FC03" pageId="2" pageNumber="63">
8 Smaller species, male L 5.5–6.8. Stridulatory comb of fore femur with 17−21 teeth. Fore and middle legs lacking preapical structure of tarsus II
<emphasis id="C7FA936CFFABE7050B07FC2CFA29FC03" box="[519,1436,920,943]" italics="true" pageId="2" pageNumber="63">
...............................................................................................................
<taxonomicName id="328E34FDFFABE7050DEFFC2CFA29FC03" ID-CoL="NNSB" box="[1263,1436,920,943]" class="Insecta" family="Notonectidae" genus="Buenoa" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="cucunubensis">B. cucunubensis</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F5314F7EFFABE7050997FC0CFA29FC43" blockId="2.[151,1436,248,1935]" pageId="2" pageNumber="63">
<keyLead id="4E7FEC4BFFABE7050997FC0CFA29FC43" pageId="2" pageNumber="63">
- Longer species, male L 8.2. Stridulatory comb of fore femur with 5 teeth. Fore and middle legs with preapical structure of tarsus II (see
<bibRefCitation id="911F328FFFABE70508A3FC6CFDDCFC43" author="Padilla-Gil" box="[419,617,984,1007]" pageId="2" pageNumber="63" refString="Padilla-Gil, D. N. (2010 a) Five new species of Buenoa (Hemiptera: Heteroptera: Notonectidae) from Colombia. Zootaxa, 2411, 22 - 32." type="journal article" year="2010" yearSuffix="a">Padilla-Gil, 2010a</bibRefCitation>
, fig. 3e)
<emphasis id="C7FA936CFFABE7050BC2FC6CFA29FC43" box="[706,1436,984,1007]" italics="true" pageId="2" pageNumber="63">
..............................................................................................
<taxonomicName id="328E34FDFFABE7050C3AFC6CFA29FC43" ID-CoL="NNS6" box="[1338,1436,984,1007]" class="Insecta" family="Notonectidae" genus="Buenoa" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="burtsa">B. burtsa</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="4E7A57DBFFABE7050997FC4CFA2EFBE3" pageId="2" pageNumber="63">
<paragraph id="F5314F7EFFABE7050997FC4CFA29FB83" blockId="2.[151,1436,248,1935]" pageId="2" pageNumber="63">
<keyLead id="4E7FEC4BFFABE7050997FC4CFA29FB83" pageId="2" pageNumber="63">
9 Stridulatory comb of fore tibia with 50 irregular teeth; rostral prong originates in distal end of third rostral segment, laterally stands out on terminal segment and has a round apex (see
<bibRefCitation id="911F328FFFABE7050A90FBACFBE2FB83" author="Padilla-Gil" box="[912,1111,1048,1071]" pageId="2" pageNumber="63" refString="Padilla-Gil, D. N. (2010 a) Five new species of Buenoa (Hemiptera: Heteroptera: Notonectidae) from Colombia. Zootaxa, 2411, 22 - 32." type="journal article" year="2010" yearSuffix="a">Padilla-Gil, 2010a</bibRefCitation>
, figs. 1a–c) .....
<taxonomicName id="328E34FDFFABE7050DF9FBACFA29FB83" ID-CoL="NNTZ" box="[1273,1436,1048,1071]" class="Insecta" family="Notonectidae" genus="Buenoa" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="tumaquensis">
<emphasis id="C7FA936CFFABE7050DF9FBACFA29FB83" box="[1273,1436,1048,1071]" italics="true" pageId="2" pageNumber="63">B. tumaquensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="F5314F7EFFABE7050997FB8CFA2EFBE3" blockId="2.[151,1436,248,1935]" box="[151,1435,1080,1103]" pageId="2" pageNumber="63">
<keyLead id="4E7FEC4BFFABE7050997FB8CFA2EFBE3" box="[151,1435,1080,1103]" pageId="2" pageNumber="63">- Stridulatory comb of fore tibia with 24–38 teeth........................................................................................................ 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="4E7A57DBFFABE7050997FBECFA2EFB23" pageId="2" pageNumber="63">
<paragraph id="F5314F7EFFABE7050997FBECFA23FBC3" blockId="2.[151,1436,248,1935]" box="[151,1430,1112,1135]" pageId="2" pageNumber="63">
<keyLead id="4E7FEC4BFFABE7050997FBECFA23FBC3" box="[151,1430,1112,1135]" pageId="2" pageNumber="63">
10 Rostral prong is projected anteriorly at distal end of third rostral segment (
<figureCitation id="6DB553FBFFABE7050AE1FBECFBAEFBC3" box="[993,1051,1112,1135]" captionStart="FIGURES 1 − 6" captionStartId="1.[151,272,1952,1976]" captionTargetBox="[154,1416,689,1906]" captionTargetId="figure@1.[151,1436,677,1929]" captionTargetPageId="1" captionText="FIGURES 1 − 6. Buenoa uselus sp. n. male. 1 Rostrum in lateral view; 2 Fore leg; 3 Femoral comb (400 X); 4 Tibial comb (400 X); 5 Preapical structure of fore tarsus II; 6 Dorsal apex of middle tibia." httpUri="https://zenodo.org/record/198115/files/figure.png" pageId="2" pageNumber="63">Fig. 1</figureCitation>
).................................................. 11
</keyLead>
</paragraph>
<paragraph id="F5314F7EFFABE7050997FBCCFA2EFB23" blockId="2.[151,1436,248,1935]" box="[151,1435,1144,1167]" pageId="2" pageNumber="63">
<keyLead id="4E7FEC4BFFABE7050997FBCCFA2EFB23" box="[151,1435,1144,1167]" pageId="2" pageNumber="63">- Rostral prong is not projected anteriorly at distal end of third rostral segment.......................................................... 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="4E7A57DBFFABE7050997FB2CFA2EFAE3" pageId="2" pageNumber="63">
<paragraph id="F5314F7EFFABE7050997FB2CFA2EFB43" blockId="2.[151,1436,248,1935]" pageId="2" pageNumber="63">
<keyLead id="4E7FEC4BFFABE7050997FB2CFA2EFB43" pageId="2" pageNumber="63">
11 Longer species, male L 7.6−7.8. Stridulatory area of fore tibia with 30 teeth; stridulatory comb of fore femur have 17−20 teeth; dorsal apex of middle tibiae with two rows of four transparent setae (
<figureCitation id="6DB553FBFFABE7050D37FB0CFB2CFB63" box="[1079,1177,1208,1231]" captionStart="FIGURES 1 − 6" captionStartId="1.[151,272,1952,1976]" captionTargetBox="[154,1416,689,1906]" captionTargetId="figure@1.[151,1436,677,1929]" captionTargetPageId="1" captionText="FIGURES 1 − 6. Buenoa uselus sp. n. male. 1 Rostrum in lateral view; 2 Fore leg; 3 Femoral comb (400 X); 4 Tibial comb (400 X); 5 Preapical structure of fore tarsus II; 6 Dorsal apex of middle tibia." httpUri="https://zenodo.org/record/198115/files/figure.png" pageId="2" pageNumber="63">Figs 1−6</figureCitation>
); fore and middle tarsus bearing preapical structure (
<figureCitation id="6DB553FBFFABE70508F2FB6CFD87FB43" box="[498,562,1240,1263]" captionStart="FIGURES 1 − 6" captionStartId="1.[151,272,1952,1976]" captionTargetBox="[154,1416,689,1906]" captionTargetId="figure@1.[151,1436,677,1929]" captionTargetPageId="1" captionText="FIGURES 1 − 6. Buenoa uselus sp. n. male. 1 Rostrum in lateral view; 2 Fore leg; 3 Femoral comb (400 X); 4 Tibial comb (400 X); 5 Preapical structure of fore tarsus II; 6 Dorsal apex of middle tibia." httpUri="https://zenodo.org/record/198115/files/figure.png" pageId="2" pageNumber="63">Fig. 5</figureCitation>
)
<emphasis id="C7FA936CFFABE7050B3CFB6CFAECFB43" box="[572,1369,1240,1263]" italics="true" pageId="2" pageNumber="63">
.........................................................................................................
<taxonomicName id="328E34FDFFABE7050DFDFB6CFAECFB43" ID-CoL="5WSKH" box="[1277,1369,1240,1263]" class="Insecta" family="Notonectidae" genus="Buenoa" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="uselus" status="sp. nov.">B. uselus</taxonomicName>
</emphasis>
<emphasis id="C7FA936CFFABE7050C60FB63FA2EFB43" bold="true" box="[1376,1435,1239,1263]" pageId="2" pageNumber="63">
<taxonomicNameLabel id="DCC92E17FFABE7050C60FB63FA2EFB43" box="[1376,1435,1239,1263]" pageId="2" pageNumber="63" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="F5314F7EFFABE7050997FB4CFA2EFAE3" blockId="2.[151,1436,248,1935]" pageId="2" pageNumber="63">
<keyLead id="4E7FEC4BFFABE7050997FB4CFA2EFAE3" pageId="2" pageNumber="63">
- Smaller species, male L 4.6–5.7. Stridulatory area of fore tibia with 35 teeth; stridulatory comb of fore femur has 11−14 teeth; dorsal apex of middle tibiae lacking of setae; fore and middle tarsus lacking preapical structure............
<emphasis id="C7FA936CFFABE70509CEFA8CFA2EFAE3" box="[206,1435,1336,1359]" italics="true" pageId="2" pageNumber="63">
................................................................................................................................................................
<taxonomicName id="328E34FDFFABE7050DFFFA8CFA2EFAE3" ID-CoL="NNTJ" box="[1279,1435,1336,1359]" class="Insecta" family="Notonectidae" genus="Buenoa" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="platycnemis">B. platycnemis</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="4E7A57DBFFABE7050997FAECFA2EFA63" pageId="2" pageNumber="63">
<paragraph id="F5314F7EFFABE7050997FAECFA2EFA23" blockId="2.[151,1436,248,1935]" pageId="2" pageNumber="63">
<keyLead id="4E7FEC4BFFABE7050997FAECFA2EFA23" pageId="2" pageNumber="63">12 Rostral prong originate in half or basal part from third rostral segment; stridulatory area of fore femur with 17 ridges .................................................................................................................................................................................... 13</keyLead>
</paragraph>
<paragraph id="F5314F7EFFABE705099DFA2CFA2EFA63" blockId="2.[151,1436,248,1935]" pageId="2" pageNumber="63">
<keyLead id="4E7FEC4BFFABE705099DFA2CFA2EFA63" pageId="2" pageNumber="63">- Rostral prong originate in apex or 1/3 apical from third rostral segment, stridulatory area of fore femur with 5−6 ridges........................................................................................................................................................................... 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="4E7A57DBFFABE7050997FA6CFA29F923" pageId="2" pageNumber="63">
<paragraph id="F5314F7EFFABE7050997FA6CFA29F983" blockId="2.[151,1436,248,1935]" pageId="2" pageNumber="63">
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13 Rostral prong distinctly longer than third rostral segment, with base originating laterally near to base of third rostral segment. Scutellum with median length equal or less than that of pronotum. Fore femur somewhat thickened at apex
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<taxonomicName id="328E34FDFFABE7050C2AF9ACFA29F983" ID-CoL="NNTF" box="[1322,1436,1560,1583]" class="Insecta" family="Notonectidae" genus="Buenoa" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="pallipes">B. pallipes</taxonomicName>
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- Rostral prong is slightly shorter or similar to long of third rostral segment, with base originating laterally midway of third segment. Scutellum with median length greater than that of pronotum. Fore femur neither wide nor thickened at apex
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<taxonomicName id="328E34FDFFABE7050C31F9CCFA29F923" ID-CoL="5WSKB" box="[1329,1436,1656,1679]" class="Insecta" family="Notonectidae" genus="Buenoa" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="pallens">B. pallens</taxonomicName>
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14 Short species L: 6.7−7.5; stridulatory comb of fore tibia with 26 teeth. Rostral prong originates in 1/3 apical of third rostral segment and has a round apex
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<taxonomicName id="328E34FDFFABE7050C3AF90CFA29F963" ID-CoL="NNT9" box="[1338,1436,1720,1743]" class="Insecta" family="Notonectidae" genus="Buenoa" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="nieseri">B. nieseri</taxonomicName>
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- Large species L: 7.7−8.1; stridulatory comb of fore tibia show 35 teeth. Rostral prong originates in proximal end of third rostral segment and has a round apex (see
<bibRefCitation id="911F328FFFABE7050BC3F94CFC3FF8A3" author="Padilla-Gil" box="[707,906,1784,1807]" pageId="2" pageNumber="63" refString="Padilla-Gil, D. N. (2010 a) Five new species of Buenoa (Hemiptera: Heteroptera: Notonectidae) from Colombia. Zootaxa, 2411, 22 - 32." type="journal article" year="2010" yearSuffix="a">Padilla-Gil, 2010a</bibRefCitation>
,
<figureCitation id="6DB553FBFFABE7050A94F94CFC69F8A3" box="[916,988,1784,1807]" captionStart="FIGURES 1 − 6" captionStartId="1.[151,272,1952,1976]" captionTargetBox="[154,1416,689,1906]" captionTargetId="figure@1.[151,1436,677,1929]" captionTargetPageId="1" captionText="FIGURES 1 − 6. Buenoa uselus sp. n. male. 1 Rostrum in lateral view; 2 Fore leg; 3 Femoral comb (400 X); 4 Tibial comb (400 X); 5 Preapical structure of fore tarsus II; 6 Dorsal apex of middle tibia." httpUri="https://zenodo.org/record/198115/files/figure.png" pageId="2" pageNumber="63">Figs. 2</figureCitation>
a–c) ........................................
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<emphasis id="C7FA936CFFABE7050C21F94CFA29F8A3" box="[1313,1436,1784,1807]" italics="true" pageId="2" pageNumber="63">B. anomala</emphasis>
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15 Inner margins of eyes convergent towards posterior part; tylus not inflated. Fore tibia with stridulatory comb of 18– 20 teeth, and all have nearly same size
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<taxonomicName id="328E34FDFFABE7050C39F88CFA2EF8E3" ID-CoL="NNTN" box="[1337,1435,1848,1871]" class="Insecta" family="Notonectidae" genus="Buenoa" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="salutis">B. salutis</taxonomicName>
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- Inner margins of eyes nearly straight; tylus very inflated. Fore tibia with stridulatory comb of 24–32 teeth of different sizes
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<taxonomicName id="328E34FDFFABE7050C39F8CCFA29F823" ID-CoL="NNTR" box="[1337,1436,1912,1935]" class="Insecta" family="Notonectidae" genus="Buenoa" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="similis">B. similis</taxonomicName>
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