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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.131.1921" ID-GBIF-Dataset="72a81528-63ef-4531-9f9c-ade551a651d8" ID-PMC="PMC3208436" ID-Pensoft-Pub="1313-2970-131-1" ID-PubMed="22140329" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-131-1" ModsDocOrigin="ZooKeys 131" ModsDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" checkinTime="1451249839150" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. &amp; Panha, Somsak" docDate="2011" docId="9A6E243C61EB46267E19D35B46B56C17" docLanguage="en" docName="ZooKeys 131: 1-161" docOrigin="ZooKeys 131" docSource="http://dx.doi.org/10.3897/zookeys.131.1921" docTitle="Orthomorpha rotundicollis Attems 1937" docType="treatment" docVersion="6" lastPageNumber="54" masterDocId="FFFF0F54FF88BD1C371C965D22453928" masterDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="161" masterPageNumber="1" pageNumber="49" updateTime="1668152297771" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Likhitrakarn, Natdanai</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Golovatch, Sergei I.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Panha, Somsak</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>131</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>161</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.131.1921</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.131.1921</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-131-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152031111" LSID="urn:lsid:plazi:treatment:9A6E243C61EB46267E19D35B46B56C17" httpUri="http://treatment.plazi.org/id/9A6E243C61EB46267E19D35B46B56C17" lastPageId="53" lastPageNumber="54" pageId="48" pageNumber="49">
<subSubSection pageId="48" pageNumber="49" type="nomenclature">
<paragraph pageId="48" pageNumber="49">
<taxonomicName LSID="http://species-id.net/wiki/Orthomorpha_rotundicollis" authority="Attems, 1937" authorityName="Attems" authorityYear="1937" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha rotundicollis" order="Polydesmida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="rotundicollis">Orthomorpha rotundicollis (Attems, 1937)</taxonomicName>
Figs 4346
</paragraph>
</subSubSection>
<subSubSection pageId="48" pageNumber="49" type="reference_group">
<paragraph pageId="48" pageNumber="49">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus rotundicollis" order="Polydesmida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="rotundicollis">Pratinus rotundicollis</taxonomicName>
<bibRefCitation pageId="48" pageNumber="49">Attems 1937</bibRefCitation>
: 118 (D).
</paragraph>
<paragraph pageId="48" pageNumber="49">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus rotundicollis" order="Polydesmida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="rotundicollis">Pratinus rotundicollis</taxonomicName>
-
<bibRefCitation author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Nouvelle Serie," pageId="114" pageNumber="115" pagination="187 - 321" title="Die von Dr. C. Dawydoff in Franzoesisch Indochina gesammelten Myriopoden." volume="6" year="1938">Attems 1938</bibRefCitation>
: 217 (D);
<bibRefCitation author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie" pageId="114" pageNumber="115" pagination="133 - 230" title="Myriopoden von Indochina. Expedition von Dr. C. Dawydoff (1938 - 1939)." volume="5" year="1953">1953</bibRefCitation>
: 170 (R).
</paragraph>
<paragraph pageId="48" pageNumber="49">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus tuberculatus" order="Polydesmida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="tuberculatus">Pratinus tuberculatus</taxonomicName>
<bibRefCitation pageId="48" pageNumber="49">Attems 1937</bibRefCitation>
: 119 (D), syn. n.
</paragraph>
<paragraph pageId="48" pageNumber="49">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus tuberculatus" order="Polydesmida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="tuberculatus">Pratinus tuberculatus</taxonomicName>
-
<bibRefCitation author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Nouvelle Serie," pageId="114" pageNumber="115" pagination="187 - 321" title="Die von Dr. C. Dawydoff in Franzoesisch Indochina gesammelten Myriopoden." volume="6" year="1938">Attems 1938</bibRefCitation>
: 219 (D).
</paragraph>
<paragraph pageId="48" pageNumber="49">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha rotundicollis" order="Polydesmida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="rotundicollis">Orthomorpha rotundicollis</taxonomicName>
-
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="205 - 283" title="Paradoxosomatidae from Borneo (Diplopoda, Polydesmida)." volume="106" year="1963">Jeekel 1963</bibRefCitation>
: 265 (M);
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="355 - 364" title="A new species of Orthomorpha Bollman from Thailand observed in migration, with taxonomic notes on the genus (Diplopoda)." volume="107" year="1964">1964</bibRefCitation>
: 361 (M, D);
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Academisch Proefschrift, Rotterdam" pageId="114" pageNumber="115" title="On the classification and geographical distribution of the family Paradoxosomatidae (Diplopoda, Polydesmida)." year="1968">1968</bibRefCitation>
: 56 (M);
<bibRefCitation author="Hoffman, RL" journalOrPublisher="Journal of Natural History" pageId="114" pageNumber="115" pagination="361 - 389" title="Descriptions and allocation of new or poorly known genera and species of Paradoxosomatidae from south-eastern Asia (Diplopoda: Polydesmida)." url="doi: 10.1080/00222937300770281" volume="7" year="1973">Hoffman 1973</bibRefCitation>
: 363 (M);
<bibRefCitation author="Hoffman, RL" journalOrPublisher="Revue suisse de Zoologie" pageId="114" pageNumber="115" pagination="699 - 719" title="Diplopoda from Malayan caves, collected by M. Pierre Strinati." volume="84" year="1977">1977</bibRefCitation>
: 700 (M);
<bibRefCitation pageId="48" pageNumber="49">Golovatch 1998</bibRefCitation>
: 42 (M, D);
<bibRefCitation author="Enghoff, H" journalOrPublisher="Arthropoda Selecta" pageId="114" pageNumber="115" pagination="29 - 43" title="A review of the millipede fauna of Vietnam (Diplopoda)." volume="13" year="2004">Enghoff et al. 2004</bibRefCitation>
: 38 (M, R).
</paragraph>
<paragraph pageId="48" pageNumber="49">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha tuberculata" order="Polydesmida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="tuberculata">Orthomorpha tuberculata</taxonomicName>
-
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="205 - 283" title="Paradoxosomatidae from Borneo (Diplopoda, Polydesmida)." volume="106" year="1963">Jeekel 1963</bibRefCitation>
: 265 (M);
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="355 - 364" title="A new species of Orthomorpha Bollman from Thailand observed in migration, with taxonomic notes on the genus (Diplopoda)." volume="107" year="1964">1964</bibRefCitation>
: 361 (M, D);
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Academisch Proefschrift, Rotterdam" pageId="114" pageNumber="115" title="On the classification and geographical distribution of the family Paradoxosomatidae (Diplopoda, Polydesmida)." year="1968">1968</bibRefCitation>
: 56 (M);
<bibRefCitation author="Hoffman, RL" journalOrPublisher="Revue suisse de Zoologie" pageId="114" pageNumber="115" pagination="699 - 719" title="Diplopoda from Malayan caves, collected by M. Pierre Strinati." volume="84" year="1977">Hoffman 1977</bibRefCitation>
: 700 (M);
<bibRefCitation pageId="48" pageNumber="49">Golovatch 1998</bibRefCitation>
: 42 (M, D);
<bibRefCitation author="Enghoff, H" journalOrPublisher="Arthropoda Selecta" pageId="114" pageNumber="115" pagination="29 - 43" title="A review of the millipede fauna of Vietnam (Diplopoda)." volume="13" year="2004">Enghoff et al. 2004</bibRefCitation>
: 39 (M, R).
</paragraph>
</subSubSection>
<subSubSection pageId="48" pageNumber="49" type="lectotype">
<paragraph pageId="48" pageNumber="49">Lectotype.</paragraph>
<paragraph pageId="48" pageNumber="49">
♂ of
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus rotundicollis" order="Polydesmida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="rotundicollis">Pratinus rotundicollis</taxonomicName>
(NHMW-3515), Vietnam, Lamdong Prov., Lang Biang,
<normalizedToken originalValue="Arbre-Broyé">Arbre-Broye</normalizedToken>
, 1500 m, 01.02.1931, leg. C. Dawydoff.
</paragraph>
</subSubSection>
<subSubSection pageId="48" pageNumber="49" type="paralectotype">
<paragraph pageId="48" pageNumber="49">Paralectotype.</paragraph>
<paragraph pageId="48" pageNumber="49">
1 ♂ of
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus rotundicollis" order="Polydesmida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="rotundicollis">Pratinus rotundicollis</taxonomicName>
(NHMW-3515), Vietnam, Lamdong Prov., Dalat, 1500 m, 02.1933, leg. C. Dawydoff.
</paragraph>
</subSubSection>
<subSubSection pageId="48" pageNumber="49" type="non-type material">
<paragraph pageId="48" pageNumber="49">Non-type material.</paragraph>
<paragraph pageId="48" pageNumber="49">
1 ♀ of
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus rotundicollis" order="Polydesmida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="rotundicollis">Pratinus rotundicollis</taxonomicName>
(NHMW-8000), Laos, Xiangkhoang Prov., Xiangkhoang Plateau, Xiang Kuang, no date, leg. C. Dawydoff, det. Attems.
</paragraph>
</subSubSection>
<subSubSection lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49" type="syntypes">
<paragraph pageId="48" pageNumber="49">Syntypes.</paragraph>
<paragraph pageId="48" pageNumber="49">
2 ♂, 1 ♀, 1 juv. of
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus tuberculatus" order="Polydesmida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="tuberculatus">Pratinus tuberculatus</taxonomicName>
(NHMW-3519), Vietnam, Lamdong Prov., Peak Lang Biang,
<normalizedToken originalValue="Arbre-Broyé">Arbre-Broye</normalizedToken>
, 1500 m, 01.02.1931, leg. C. Dawydoff.
</paragraph>
<paragraph lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
Lectotype designation proposed herewith is necessary to ensure the species is based on a complete male coming from a certain locality, because (1) (
<bibRefCitation pageId="48" pageNumber="49">Attems (1937</bibRefCitation>
,
<bibRefCitation author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Nouvelle Serie," pageId="114" pageNumber="115" pagination="187 - 321" title="Die von Dr. C. Dawydoff in Franzoesisch Indochina gesammelten Myriopoden." volume="6" year="1938">1938</bibRefCitation>
)
<pageBreakToken pageId="49" pageNumber="50" start="start">had</pageBreakToken>
provided no information on the number of syntypes, and (2) he stated their provenance to have been both from
<normalizedToken originalValue="Arbre-Broyé">Arbre-Broye</normalizedToken>
and Dalat, Vietnam.
</paragraph>
</subSubSection>
<subSubSection lastPageId="53" lastPageNumber="54" pageId="49" pageNumber="50" type="redescription">
<paragraph pageId="49" pageNumber="50">Redescription.</paragraph>
<paragraph lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
Length 29-31 mm (♂), ca 31-35 mm (♀), width of midbody pro- and metazona 2.5-3.0 and 3.7-4.1 mm (♂), 3.4 and 4.5 mm (♀), respectively (vs 2.5 and 4.0 mm in width, as given in the available descriptions (
<bibRefCitation pageId="49" pageNumber="50">Attems 1937</bibRefCitation>
,
<bibRefCitation author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Nouvelle Serie," pageId="114" pageNumber="115" pagination="187 - 321" title="Die von Dr. C. Dawydoff in Franzoesisch Indochina gesammelten Myriopoden." volume="6" year="1938">1938</bibRefCitation>
)). Lectotype ca 31 mm long, 2.9 and 4.1 mm wide on midbody pro- and metazona, respectively. Coloration of alcohol material upon long-term preservation rather uniformly light grey-brown (Figs 43 &amp; 45) (vs with a cingulate pattern of castaneous brown head and collum, as well as regions of pro- and metazona adjacent to stricture
<pageBreakToken pageId="50" pageNumber="51" start="start">alternating</pageBreakToken>
with light brownish remaining parts, including antennae and legs, as given in the descriptions (
<bibRefCitation pageId="50" pageNumber="51">Attems 1937</bibRefCitation>
,
<bibRefCitation author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Nouvelle Serie," pageId="114" pageNumber="115" pagination="187 - 321" title="Die von Dr. C. Dawydoff in Franzoesisch Indochina gesammelten Myriopoden." volume="6" year="1938">1938</bibRefCitation>
)).
</paragraph>
<paragraph lastPageId="52" lastPageNumber="53" pageId="50" pageNumber="51">
Head usual, clypeolabral region densely setose, surface of vertex smooth, epicranial suture distinct. Antennae rather long and slender (Figs 43J, 45B &amp; J), extending behind body segment 4 (♂) or 3 (♀) dorsally. Head in width &lt;collum &lt;segments 3 and 4 &lt;segment 2 &lt;segments 5-17 (♂, ♀), gently and gradually tapering thereafter. Collum smooth, with three transverse rows of setae, 4+4 anterior, 2+2 intermediate, and 3+3 posterior setae; caudal corner of paraterga subrectangular, narrowly rounded (Figs 43A, B, J, 45A, B &amp; J). Tegument smooth and shining; metaterga very faintly rugulose, each postcollum one with one caudal row of 3(4)+3(4) very small tubercles growing a little larger laterally, as well as with 2+2 either minute or fully obliterated knobs with insertion points of abraded setae in front row; prozona very finely shagreened, surface below paraterga finely microgranulate. Axial line visible, starting from collum. Paraterga very strongly developed (Figs 43A-G, J-L, 45A-G &amp; J-L), all subhorizontal
<pageBreakToken pageId="51" pageNumber="52" start="start">and</pageBreakToken>
lying below dorsum, thin in lateral view, like blunt blades, a little thicker only on pore-bearing segments, on postcollum segments extending increasingly beyond rear tergal margin, nearly pointed to pointed, caudal tip on paraterga 16-19 evidently curved mesad. Calluses on delimited by a sulcus only dorsally, rather narrow. Paraterga 2 broad, anterior edge angulate, lateral edge with three small incisions in anterior half; posterior edge evidently concave (Figs 43A, B, J, 45A, B &amp; J). Paraterga 3 and 4 subequal, like subsequent paraterga, anterior edge broadly rounded, bordered and fused to callus, lateral edge with two small incisions in anterior half on poreless segments, with
<pageBreakToken pageId="52" pageNumber="53" start="start">only</pageBreakToken>
one incision near front 1/3 on pore-bearing ones. Ozopores evident, lateral, lying inside an ovoid groove at about 1/4 metazonital length. Transverse sulcus complete on metaterga 5-18, incomplete on metatergum 4, shallow, not reaching bases of paraterga, beaded at bottom, slightly sinuate anteromedially (Figs 43A- D, F, J-L, 45A-F &amp; J-L). Stricture between pro- and metazona rather narrow, deep, ribbed at bottom down to base of paraterga. Pleurosternal carinae complete crests only on segments 2-4 (Figs 43B, D, E, J-K, 45A-F &amp; J-L), each with an evident sharp denticle caudally, thereafter increasingly strongly reduced until segment 10 (♂, ♀). Epiproct (Figs 43E-G, 45E-G) conical, flattened dorsoventrally, apical papillae small; tip subtruncate; pre-apical papillae small, but visible, lying close to tip. Hypoproct (Figs 43G, 45G) roundly subtriangular, setiferous knobs at caudal margin small and well-separated.
</paragraph>
<paragraph lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
Sterna sparsely setose, without modifications, but with two rather large, rounded, fully separated, but nearly contiguous, setose cones between ♂ coxae 4 (Figs 43H, I, 45H &amp; I). A paramedian pair of small tubercles in front of gonopod aperture. Legs
<pageBreakToken pageId="53" pageNumber="54" start="start">long</pageBreakToken>
and slender, midbody ones ca 1.2-1.3 (♂) or 0.8-0.9 (♀) as long as midbody height, prefemora without modifications, tarsal brushes present until ♂ legs 4.
</paragraph>
<paragraph pageId="53" pageNumber="54">
Gonopods (Figs 44 &amp; 46) simple. Coxa long and slender, with several setae distodorsally. Prefemur densely setose, nearly 3 times shorter than femorite +
<normalizedToken originalValue="“postfemoral”">&quot;postfemoral&quot;</normalizedToken>
part. Femorite very slender, evidently curved, very slightly enlarged distad,
<normalizedToken originalValue="“postfemoral”">&quot;postfemoral&quot;</normalizedToken>
part demarcated by an oblique lateral sulcus; tip of solenophore small, trifid, with two subequal denticles (one terminal, the other middle) and a broader subterminal lobule.
</paragraph>
<caption pageId="53" pageNumber="54">
<paragraph pageId="53" pageNumber="54">
Figure 43.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha rotundicollis" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rotundicollis">Orthomorpha rotundicollis</taxonomicName>
Attems, 1937, ♂ lectotype (
<normalizedToken originalValue="A–I">A-I</normalizedToken>
), ♀ (
<normalizedToken originalValue="J–L">J-L</normalizedToken>
) of
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus rotundicollis" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rotundicollis">Pratinus rotundicollis</taxonomicName>
Attems, 1937. A, B, J anterior part of body, dorsal, lateral and dorsal views, respectively C, D, K segments 10 and 11, dorsal, lateral and dorsal views, respectively
<normalizedToken originalValue="E–G">E-G</normalizedToken>
, L posterior part of body, lateral, dorsal, ventral and dorsal views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively.
</paragraph>
</caption>
<caption pageId="53" pageNumber="54">
<paragraph pageId="53" pageNumber="54">
Figure 44.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha rotundicollis" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rotundicollis">Orthomorpha rotundicollis</taxonomicName>
Attems, 1937, ♂ lectotype of
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus rotundicollis" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rotundicollis">Pratinus rotundicollis</taxonomicName>
Attems, 1937. A, B right gonopod, lateral and mesal views, respectively.
</paragraph>
</caption>
<caption pageId="53" pageNumber="54">
<paragraph pageId="53" pageNumber="54">
Figure 45.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha rotundicollis" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rotundicollis">Orthomorpha rotundicollis</taxonomicName>
Attems, 1937, ♂ syntype (
<normalizedToken originalValue="A–I">A-I</normalizedToken>
), ♀ syntype (
<normalizedToken originalValue="J–L">J-L</normalizedToken>
) of
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus tuberculatus" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tuberculatus">Pratinus tuberculatus</taxonomicName>
<bibRefCitation pageId="53" pageNumber="54">Attems 1937</bibRefCitation>
. A, B, J anterior part of body, dorsal, lateral and dorsal views, respectively C, D segments 11 and 12, dorsal and lateral veiws, respectively K segments 10 and 11, dorsal view
<normalizedToken originalValue="E–F">E-F</normalizedToken>
, L posterior part of body, lateral, dorsal, ventral and dorsal views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively.
</paragraph>
</caption>
<caption pageId="53" pageNumber="54">
<paragraph pageId="53" pageNumber="54">
Figure 46.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha rotundicollis" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rotundicollis">Orthomorpha rotundicollis</taxonomicName>
Attems, 1937, ♂ syntype of
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus tuberculatus" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tuberculatus">Pratinus tuberculatus</taxonomicName>
<bibRefCitation pageId="53" pageNumber="54">Attems 1937</bibRefCitation>
. A, B right gonopod, lateral and mesal views, respectively.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="53" pageNumber="54" type="remark">
<paragraph pageId="53" pageNumber="54">Remark.</paragraph>
<paragraph pageId="53" pageNumber="54">
This species is known from Lamdong Province, southern Vietnam and Luang Prabang Province, Laos (
<bibRefCitation author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Nouvelle Serie," pageId="114" pageNumber="115" pagination="187 - 321" title="Die von Dr. C. Dawydoff in Franzoesisch Indochina gesammelten Myriopoden." volume="6" year="1938">Attems 1938</bibRefCitation>
,
<bibRefCitation author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie" pageId="114" pageNumber="115" pagination="133 - 230" title="Myriopoden von Indochina. Expedition von Dr. C. Dawydoff (1938 - 1939)." volume="5" year="1953">1953</bibRefCitation>
). The type locality of both
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha rotundicollis" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rotundicollis">Orthomorpha rotundicollis</taxonomicName>
and
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha tuberculata" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tuberculata">Orthomorpha tuberculata</taxonomicName>
being exactly the same (
<normalizedToken originalValue="Arbre-Broyé">Arbre-Broye</normalizedToken>
at Peak Lang Biang), and the differences between them so minor, rather to be treated as infraspecific variations, we are inclined to formally synonymize these species. Indeed, distinctions are observed only in
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha rotundicollis" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rotundicollis">Orthomorpha rotundicollis</taxonomicName>
showing a slightly more rounded caudal corner of the paraterga on the collum (vs slightly more angular in
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha tuberculata" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tuberculata">Orthomorpha tuberculata</taxonomicName>
), in the front row of metatergal setae (= insertion points) borne on virtually fully obliterated knobs (vs on minute knobs in
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha tuberculata" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tuberculata">Orthomorpha tuberculata</taxonomicName>
), and in the apical papillae on the epiproct directed more caudally than ventrally (vs more ventrally than caudally in
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha tuberculata" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tuberculata">Orthomorpha tuberculata</taxonomicName>
) (
<bibRefCitation pageId="53" pageNumber="54">Attems 1937</bibRefCitation>
,
<bibRefCitation author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Nouvelle Serie," pageId="114" pageNumber="115" pagination="187 - 321" title="Die von Dr. C. Dawydoff in Franzoesisch Indochina gesammelten Myriopoden." volume="6" year="1938">1938</bibRefCitation>
,
<bibRefCitation author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie" pageId="114" pageNumber="115" pagination="133 - 230" title="Myriopoden von Indochina. Expedition von Dr. C. Dawydoff (1938 - 1939)." volume="5" year="1953">1953</bibRefCitation>
). We can also add another two differences: paraterga 18 and 19 seem to be a little shorter in
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha rotundicollis" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rotundicollis">Orthomorpha rotundicollis</taxonomicName>
than in
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha tuberculata" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tuberculata">Orthomorpha tuberculata</taxonomicName>
, and the caudal row of tubercles on a few caudalmost metaterga demonstrating a pattern of 3+4 or 4+3 in
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha tuberculata" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tuberculata">Orthomorpha tuberculata</taxonomicName>
(vs invariably 3+3 in
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha rotundicollis" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rotundicollis">Orthomorpha rotundicollis</taxonomicName>
).
</paragraph>
<paragraph pageId="53" pageNumber="54">The above is a combined redescription, based on material of both nominate species.</paragraph>
</subSubSection>
</treatment>
</document>

142
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<subSubSection id="5ADD7E83FFD5FF9CFF0AFDA8043CFDF0" pageId="1" pageNumber="142" type="nomenclature">
<paragraph id="12782D08FFD5FF9CFF0AFDA80728FD90" blockId="1.[151,624,532,591]" box="[151,624,532,559]" pageId="1" pageNumber="142">
<heading id="49309A64FFD5FF9CFF0AFDA80728FD90" bold="true" box="[151,624,532,559]" fontSize="11" level="1" pageId="1" pageNumber="142" reason="1">
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<taxonomicName id="D5C7568BFFD5FF9CFF0AFDA8075DFD90" authority="Yakovlev" authorityName="Yakovlev" box="[151,517,532,559]" class="Insecta" family="Cossidae" genus="Isocossus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="zolotuhini" status="sp. nov.">
<emphasis id="20B3F11AFFD5FF9CFF0AFDA804D4FD91" bold="true" box="[151,396,532,558]" italics="true" pageId="1" pageNumber="142">Isocossus zolotuhini</emphasis>
Yakovlev
</taxonomicName>
,
<taxonomicNameLabel id="3B804C61FFD5FF9CFD8FFDA90728FD90" box="[530,624,533,559]" pageId="1" pageNumber="142" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="12782D08FFD5FF9CFF0AFD84043CFDF0" blockId="1.[151,624,532,591]" box="[151,356,568,591]" pageId="1" pageNumber="142">
(
<figureCitation id="8AFC318DFFD5FF9CFF03FD8405BAFDF0" box="[158,226,568,591]" captionStart="FIGURES 1 − 11. 1" captionStartId="2.[151,258,1798,1820]" captionTargetBox="[211,1311,205,1726]" captionTargetId="figure@2.[204,1332,194,1744]" captionTargetPageId="2" captionText="FIGURES 1 − 11. 1. Isocossus zolotuhini Yakovlev, sp. nov., holotype, male. 2. Isocossus cruciatus (Holloway, 1986), holotype, male. 3. Isocossus seria (Holloway, 1986), holotype, male. 4. Isocossus stroehli Yakovlev, 2006, holotype, male. 5. Isocossus stroehli Yakovlev, 2006, paratype, male, Malaysia, Cameron Highlands, SW Ringlet (Brücke), 650 m, 4 ° 21 ’ 36 ’’ N; 101 ° 20 ’ 52 ’’ E, 12. – 21. 0 7. 2001. leg. Ströhle (MSW). 6. Isocossus retak (Holloway, 1986), holotype, male. 7. Isocossus rufipecten (Holloway, 1986), holotype, male. 8. Isocossus telisai (Holloway, 1986), holotype, male. 9. Isocossus telisai (Holloway, 1986), male, Borneo, Sabah prov., Trus Madi bei Apin Apin, 1080 m, 21.03. – 12.04.2006, leg. K. Martini (MWM). 10. Isocossus vandeldeni Roepke, 1957, holotype, male. 11. Isocossus vandeldeni Roepke, 1957, male, Borneo, Sabah prov., Trus Madi bei Apin Apin, 1080 m, 21.03. – 12. 0 4. 2006, leg. K. Martini (MWM)." httpUri="https://zenodo.org/record/234109/files/figure.png" pageId="1" pageNumber="142">Figs 1</figureCitation>
,
<figureCitation id="8AFC318DFFD5FF9CFF72FD840404FDF0" box="[239,348,568,591]" captionStart="FIGURES 12 − 14. 12" captionStartId="3.[151,258,1881,1903]" captionTargetBox="[171,1425,208,1848]" captionTargetId="figure@3.[151,1436,191,1859]" captionTargetPageId="3" captionText="FIGURES 12 − 14. 12. Genitalia of holotype of Isocossus zolotuhini Yakovlev, sp. nov. 13. Distribution of Genus Isocossus Roepke, 1957. 14. Type locality (Thue Thien Hue Prov., A Ruang) of Isocossus zolotuhini Yakovlev, sp. nov. (photo by Vadim Zolotuhin)" httpUri="https://zenodo.org/record/234110/files/figure.png" pageId="1" pageNumber="142">12, 13−14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5ADD7E83FFD5FF9CFF0AFDC101B9FD08" pageId="1" pageNumber="142" type="materials_examined">
<paragraph id="12782D08FFD5FF9CFF0AFDC101B9FD08" blockId="1.[151,1436,637,1460]" pageId="1" pageNumber="142">
<emphasis id="20B3F11AFFD5FF9CFF0AFDC1046EFD2B" bold="true" box="[151,310,637,660]" pageId="1" pageNumber="142">
<typeStatus id="CD7C93AAFFD5FF9CFF0AFDC10597FD2B" box="[151,207,637,660]" pageId="1" pageNumber="142">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="CD7C93AAFFD5FF9CFED8FDC104F0FD2B" box="[325,424,637,660]" pageId="1" pageNumber="142" type="holotype">Holotype</typeStatus>
: male, C[entral].
<collectingCountry id="6AD06D98FFD5FF9CFDF4FDC1079EFD2B" box="[617,710,637,660]" name="Vietnam" pageId="1" pageNumber="142">Vietnam</collectingCountry>
, Thue Thien Hue Prov., A Ruang,
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,
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;
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, 24– 27.0 4.2009, leg. V. Zolotuhin &amp; A. Gurkovich (MWM);
<typeStatus id="CD7C93AAFFD5FF9CFD69FD1C060CFD08" box="[756,852,672,695]" pageId="1" pageNumber="142" type="paratype">paratype</typeStatus>
: male, same locality and data (RYB).
</paragraph>
</subSubSection>
<subSubSection id="5ADD7E83FFD5FF9CFF5AFD7E0746FCFE" pageId="1" pageNumber="142" type="diagnosis">
<paragraph id="12782D08FFD5FF9CFF5AFD7E0746FCFE" blockId="1.[151,1436,637,1460]" pageId="1" pageNumber="142">
<emphasis id="20B3F11AFFD5FF9CFF5AFD7E046CFD66" bold="true" box="[199,308,706,729]" pageId="1" pageNumber="142">Diagnosis</emphasis>
. The new species most closely resembles
<taxonomicName id="D5C7568BFFD5FF9CFC98FD7E060CFD66" box="[773,852,706,729]" class="Insecta" family="Cossidae" genus="Cossus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="retak">
<emphasis id="20B3F11AFFD5FF9CFC98FD7E060CFD66" box="[773,852,706,729]" italics="true" pageId="1" pageNumber="142">I. retak</emphasis>
</taxonomicName>
and
<taxonomicName id="D5C7568BFFD5FF9CFC11FD7E0141FD66" box="[908,1049,706,729]" class="Insecta" family="Cossidae" genus="Isocossus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="vandeldeni">
<emphasis id="20B3F11AFFD5FF9CFC11FD7E0141FD66" box="[908,1049,706,729]" italics="true" pageId="1" pageNumber="142">I. vandeldeni</emphasis>
</taxonomicName>
, but is readily distinguished by the structure of the male genitalia. It differs from
<taxonomicName id="D5C7568BFFD5FF9CFD1DFD590794FD43" box="[640,716,741,764]" class="Insecta" family="Cossidae" genus="Cossus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="retak">
<emphasis id="20B3F11AFFD5FF9CFD1DFD590794FD43" box="[640,716,741,764]" italics="true" pageId="1" pageNumber="142">I. retak</emphasis>
</taxonomicName>
by more developed sacculus along with slightly curved costal edge of valva and more massive processes of transtilla, and from
<taxonomicName id="D5C7568BFFD5FF9CFC8FFCB406C4FCA0" box="[786,924,776,799]" class="Insecta" family="Cossidae" genus="Isocossus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="vandeldeni">
<emphasis id="20B3F11AFFD5FF9CFC8FFCB406C4FCA0" box="[786,924,776,799]" italics="true" pageId="1" pageNumber="142">I. vandeldeni</emphasis>
</taxonomicName>
by longer processes of the transtilla, the smooth dorsal edge of valve, and thin uncus.
</paragraph>
</subSubSection>
<subSubSection id="5ADD7E83FFD5FF9CFF5AFCF104DFFAF3" pageId="1" pageNumber="142" type="description">
<paragraph id="12782D08FFD5FF9CFF5AFCF101AAFBAE" blockId="1.[151,1436,637,1460]" pageId="1" pageNumber="142">
<emphasis id="20B3F11AFFD5FF9CFF5AFCF10413FCDB" bold="true" box="[199,331,845,868]" pageId="1" pageNumber="142">Description</emphasis>
. Forewing length
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in the
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(
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in the
<typeStatus id="CD7C93AAFFD5FF9CFC3FFCF1015AFCDB" box="[930,1026,845,868]" pageId="1" pageNumber="142" type="paratype">paratype</typeStatus>
), wingspan
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in the
<typeStatus id="CD7C93AAFFD5FF9CFA89FCF10029FCDB" box="[1300,1393,845,868]" pageId="1" pageNumber="142" type="holotype">holotype</typeStatus>
(
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in the
<typeStatus id="CD7C93AAFFD5FF9CFE95FCCC0431FC38" box="[264,361,880,903]" pageId="1" pageNumber="142" type="paratype">paratype</typeStatus>
). Antennae bipectinate, processes of antenna strongly cut in distal third (in comparison with length of processes of proximal part of antenna). Tegula pale brown. Thorax with spot of flour white hairs from above. Forewing rather narrow, with rounded apex, flour white with fine grey pattern, base of forewing with rounded black spot on costal edge. All area of forewing with fine grey wavy pattern. Cubital zone of forewing discal area with light brown field. Hindwing grey without pattern. Fringe of both wings checkered, dark near veins and pale between them.
</paragraph>
<paragraph id="12782D08FFD5FF9CFF5AFBA107A7FA96" blockId="1.[151,1436,637,1460]" pageId="1" pageNumber="142">Male genitalia typical for the genus, uncus stout with slightly developed incisure; tegumen medium-sized; branches of gnathos rather stout, medium-sized; gnathos stout, densely covered by small thorns; processes of transtilla massive, hook-like, with thick bases and pointed tips, as long as 1/3 of valva; juxta small, heavily sclerotized, with long oar-like lateral processes situated at 100º to each other, length of juxta processes equal to length of transtilla processes; valvae rather broad, with well-developed sacculus, their distal fourth membranous, transition zone between proximal sclerotized part and distal membranous part unsharp, costal edge of valva slightly curved; saccus semicircular, rather massive. Aedeagus thick, strongly arched at medium third, slightly shorter than valva, pore of vesica in dorso-apical position and occupying 2/5 of aedeagus length, vesica without cornuti.</paragraph>
<paragraph id="12782D08FFD5FF9CFF5AFA8904DFFAF3" blockId="1.[151,1436,637,1460]" box="[199,391,1333,1356]" pageId="1" pageNumber="142">Female unknown.</paragraph>
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<emphasis id="20B3F11AFFD5FF9CFF5AFAE40418FAD0" bold="true" box="[199,320,1368,1391]" pageId="1" pageNumber="142">Etymology</emphasis>
. The new species is named after a well known Russian lepidopterist Dr. Vadim Zolotuhin (Ulyanovsk), a specialist in Macroheterocera.
</paragraph>
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<emphasis id="20B3F11AFFD5FF9CFF5AFA210478FA0B" bold="true" box="[199,288,1437,1460]" pageId="1" pageNumber="142">Ecology</emphasis>
. Specimens were attracted by light trap at the end of April, in a primary tropical forest at
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.
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1.
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Fabr.
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<paragraph id="03A147F65CE9A9D6ABC4F886829895DE" pageNumber="720">Im Mus. C. Vienn. aus Brasilien und Mexico, ein Exemplar ist mit einem Zettel versehen, auf welchem Folgendes verzeichnet ist: „ Kommt vor in der Schomburgia tibicinis in den Pseudobulben, ihre Verwundung sehr schmerzhaft, Stadt Veracruz, am Fusse des Orizabas &quot;. Ferner aus Rio Janeiro von der Novara-Expedition mitgebracht, und in meiner Sammlung ebenfalls aus Brasilien.</paragraph>
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<paragraph pageId="11" pageNumber="62">Taxon classification Animalia Coleoptera Staphylinidae</paragraph>
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<taxonomicName authority="Bernhauer" class="Insecta" family="Staphylinidae" genus="Atheta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atheta (Alaobia) ventricosa" order="Coleoptera" pageId="11" pageNumber="62" phylum="Arthropoda" rank="species" species="ventricosa" subGenus="Alaobia">Atheta (Alaobia) ventricosa Bernhauer</taxonomicName>
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(for diagnosis and illustrations, see
<bibRefCitation author="Klimaszewski, J" journalOrPublisher="Pensoft Publishers, Sofia-Moscow" pageId="32" pageNumber="83" title="Aleocharine beetles (Coleoptera, Staphylinidae) of the province of Newfoundland and Labrador, Canada." year="2011">Klimaszewski et al. 2011</bibRefCitation>
)
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<paragraph pageId="11" pageNumber="62">Distribution.</paragraph>
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Distribution of
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<td colspan="1" pageId="11" pageNumber="62" rowspan="1">SK</td>
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<td colspan="1" pageId="11" pageNumber="62" rowspan="1">Saskatchewan: 49.6704°, -109.5005°</td>
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<bibRefCitation author="Gusarov, VI" journalOrPublisher="Zootaxa" pageId="30" pageNumber="81" pagination="1 - 134" title="Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes." volume="353" year="2003 a">Gusarov 2003a</bibRefCitation>
<bibRefCitation author="Klimaszewski, J" journalOrPublisher="The Canadian Entomologist" pageId="31" pageNumber="82" pagination="1 - 48" title="Rove beetles (Coleoptera: Staphylinidae) in red spruce stands, eastern Canada: diversity, abundance, and descriptions of new species." url="10.4039/n03-123" volume="137" year="2005">Klimaszewski et al. 2005</bibRefCitation>
<bibRefCitation author="Klimaszewski, J" journalOrPublisher="The Canadian Entomologist" pageId="31" pageNumber="82" pagination="265 - 291" title="Six new species and records of aleocharine beetles from Yukon and Alaska (Coleoptera: Staphylinidae: Aleocharinae)." url="10.4039/n07-054" volume="140" year="2008">2008</bibRefCitation>
<bibRefCitation author="Klimaszewski, J" journalOrPublisher="Pensoft Publishers, Sofia-Moscow" pageId="32" pageNumber="83" title="Aleocharine beetles (Coleoptera, Staphylinidae) of the province of Newfoundland and Labrador, Canada." year="2011">2011</bibRefCitation>
<bibRefCitation author="Gouix, N" journalOrPublisher="Pensoft Publishers, Sofia-Moscow" pageId="30" pageNumber="81" title="Catalogue of aleocharine rove beetles of Canada and Alaska (Coleoptera; Staphylinidae; Aleocharinae)." year="2007">Gouix and Klimaszewski 2007</bibRefCitation>
<bibRefCitation author="Majka, CG" editor="Majka, CG" journalOrPublisher="ZooKeys" pageId="32" pageNumber="83" pagination="85 - 114" title="New records of Canadian Aleocharinae (Coleoptera: Staphylinidae)." url="10.3897/zookeys.2.7" volume="2" volumeTitle="Biodiversity, Biosystematics, and Ecology of Canadian Coleoptera." year="2008">Majka and Klimaszewski 2008</bibRefCitation>
<bibRefCitation author="Majka, CG" journalOrPublisher="ZooKeys" pageId="32" pageNumber="83" pagination="15 - 39" title="Contributions to the knowledge of the Aleocharinae (Coleoptera, Staphylinidae) in the Maritime Provinces of Canada." url="10.3897/zookeys.46.413" volume="46" year="2010">2010</bibRefCitation>
<bibRefCitation author="Bousquet, Y" journalOrPublisher="Pensoft Publishers, Sofia-Moscow" pageId="30" pageNumber="81" title="Checklist of beetles (Coleoptera) of Canada and Alaska." year="2013">Bousquet et al. 2013</bibRefCitation>
</td>
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<paragraph pageId="11" pageNumber="62">Natural history.</paragraph>
<paragraph pageId="11" pageNumber="62">
In Saskatchewan, one male was captured in the riparian zone of a pond. In Newfoundland, adults were collected in unbaited and carrion-baited pitfall traps and flight intercept traps in various coniferous and mixedwood forest types (
<bibRefCitation author="Klimaszewski, J" journalOrPublisher="Pensoft Publishers, Sofia-Moscow" pageId="32" pageNumber="83" title="Aleocharine beetles (Coleoptera, Staphylinidae) of the province of Newfoundland and Labrador, Canada." year="2011">Klimaszewski et al. 2011</bibRefCitation>
). Elsewhere, adults were found in coniferous forests and in organic litter in mixed forests (
<bibRefCitation author="Klimaszewski, J" journalOrPublisher="The Canadian Entomologist" pageId="31" pageNumber="82" pagination="1 - 48" title="Rove beetles (Coleoptera: Staphylinidae) in red spruce stands, eastern Canada: diversity, abundance, and descriptions of new species." url="10.4039/n03-123" volume="137" year="2005">Klimaszewski et al. 2005</bibRefCitation>
,
<bibRefCitation author="Majka, CG" editor="Majka, CG" journalOrPublisher="ZooKeys" pageId="32" pageNumber="83" pagination="85 - 114" title="New records of Canadian Aleocharinae (Coleoptera: Staphylinidae)." url="10.3897/zookeys.2.7" volume="2" volumeTitle="Biodiversity, Biosystematics, and Ecology of Canadian Coleoptera." year="2008">Majka and Klimaszewski 2008</bibRefCitation>
).The adults were collected from May to October.
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