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Genus
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<bibRefCitation id="7666ACF87068FF9F29A8F9A88832F543" author="Schubart" box="[411,600,1618,1644]" pageId="1" pageNumber="56" refString="Schubart, O. (1960) Eine neue cavernicole Stylodesmide aus Marokko (Diplopoda, Proterospermophora). Bulletin de la societe des sciences naturelles du Maroc, 40 (1), 27 - 32." type="journal article" year="1960">Schubart, 1960</bibRefCitation>
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: 27
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.
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species:
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<bibRefCitation id="7666ACF87068FF9F2BBFF9638E4CF580" author="Schubart" box="[908,1062,1689,1711]" pageId="1" pageNumber="56" refString="Schubart, O. (1960) Eine neue cavernicole Stylodesmide aus Marokko (Diplopoda, Proterospermophora). Bulletin de la societe des sciences naturelles du Maroc, 40 (1), 27 - 32." type="journal article" year="1960">Schubart, 1960</bibRefCitation>
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(
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)
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by original designation.
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<emphasis id="20830D1B7068FF9F28A4F9218B60F5DB" bold="true" box="[151,266,1755,1780]" pageId="1" pageNumber="56">Diagnosis</emphasis>
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(Based on
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<bibRefCitation id="7666ACF87068FF9F29BCF9268852F5DB" author="Schubart" box="[399,568,1756,1780]" pageId="1" pageNumber="56" refString="Schubart, O. (1960) Eine neue cavernicole Stylodesmide aus Marokko (Diplopoda, Proterospermophora). Bulletin de la societe des sciences naturelles du Maroc, 40 (1), 27 - 32." type="journal article" year="1960">Schubart 1960</bibRefCitation>
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). Body with 20 ‘segments’ (18 podous + 1 apodous body rings + telson) in both sexes; collum with 10 lobes on the front margin and concealing the head from above, body rings strongly arched, terga with 2 paramedian, 2 dorsolateral and 2 lateral rows of tubercles (and 2 indistinctly delimited ventrolateral rows of smaller ones); paraterga with two lobes and an additional one on the poriferous segments; ozopores on rings 5, 7, 9, 10, 13, 15 and 16, preanal ring with 6 lobes. Gonopods with large, hemispherical coxae completely covering the lateral side of the telopodites, prefemur transverse and setose, tibiotarsus distally bifurcating into two branches: one uniformly broad and furrowed at the tip, the second distally extended and apically divided into a downturned solenomerite and an accessory processes.
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215
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|
||||
<figureCitation id="8ACCCD8C706BFF9C28A4FF408B6CF3FD" box="[151,262,186,210]" captionStart-0="FIGURES 1 – 6" captionStart-1="FIGURES 13 – 16" captionStart-2="FIGURE 17" captionStartId-0="2.[151,264,1803,1826]" captionStartId-1="5.[151,264,1413,1436]" captionStartId-2="7.[151,250,1720,1743]" captionTargetBox-0="[215,1372,518,1779]" captionTargetBox-1="[202,1375,195,1388]" captionTargetBox-2="[375,1211,1095,1695]" captionTargetId-0="figure@2.[215,1372,518,1779]" captionTargetId-1="figure@5.[202,1385,195,1391]" captionTargetId-2="figure@7.[375,1212,1095,1695]" captionTargetPageId-0="2" captionTargetPageId-1="5" captionTargetPageId-2="7" captionText-0="FIGURES 1 – 6. Rharodesmus tabarkensis n. sp., adult, morphological details: Fig. 1 Head and first two body rings, adult male, anterior view; Fig. 2 Collum and three subsequent tergites, adult female; Fig. 3 Prozona detail: cellular scaly structure (left), transverse ridge of tooth-shaped cuticular ‘ cells’ and knobby area next to metazona (right), adult female; Fig. 4 Body rings 3 and 4, showing microsculpture of prozona and tubercles of metazona, adult female; Fig. 5 Tubercles on metazona 7 and 8, adult female; Fig 6 Limbus, adult male. Abbreviations: lb: posterior lobe; o: ozopore; p: porostele; t: caudolateral tubercle." captionText-1="FIGURES 13 – 16. Rharodesmus tabarkensis n. sp., gonopodal structures: Fig. 13 Gonopods in situ, profile; Fig. 14 Right gonopod, mesal view; Fig. 15 Close up of coxal distal process and hemispherical hump; Fig. 16 Left gonopod, antero-mesal view. Abbreviations: A: process A; B: process B; c: coxa, ca: cannula; cp: conical process, h: setose coxal hump; r: rounded process; so: solenomere; p: prefemoral part; sp: smooth distal process." captionText-2="FIGURE 17. Rharodesmus tabarkensis n. sp. mandible. Arrow pointing at the molar hook." httpUri-0="https://zenodo.org/record/202202/files/figure.png" httpUri-1="https://zenodo.org/record/202203/files/figure.png" httpUri-2="https://zenodo.org/record/202204/files/figure.png" pageId="2" pageNumber="57">Figs 1–17</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="5AED8282706BFF9C28A4FEF88E64F24D" pageId="2" pageNumber="57" type="materials_examined">
|
||||
<paragraph id="1248D109706BFF9C28A4FEF88E64F24D" blockId="2.[151,1437,258,462]" pageId="2" pageNumber="57">
|
||||
<emphasis id="20830D1B706BFF9C28A4FEF88BEBF234" bold="true" box="[151,385,258,283]" pageId="2" pageNumber="57">Material examined.</emphasis>
|
||||
13 3, 8 ƤƤ,
|
||||
<specimenCount id="04F11A80706BFF9C2A07FEF888D5F235" box="[564,703,258,282]" pageId="2" pageNumber="57" type="juvenile">14 juveniles</specimenCount>
|
||||
,
|
||||
<collectingCountry id="6AE09199706BFF9C2AF8FEF88954F235" box="[715,830,258,282]" name="Tunisia" pageId="2" pageNumber="57">TUNISIA</collectingCountry>
|
||||
, Jendouba Governorate, Tabarka, N36°57.8, E8°44.6, alt. <
|
||||
<quantity id="D50F7CEC706BFF9C28E4FEDC8B78F211" box="[215,274,294,318]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="57" unit="m" value="40.0">40 m</quantity>
|
||||
, coastal slope below the Genoese fort, under stones,
|
||||
<date id="6649F7C9706BFF9C2B51FEDC89BAF211" box="[866,976,294,318]" pageId="2" pageNumber="57" value="2009-03-09">9.iii.2009</date>
|
||||
, N. Akkari & H. Enghoff leg. (
|
||||
<collectionCode id="74E649CC706BFF9C2D1FFEDC8FE9F211" box="[1324,1411,294,318]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="2" pageNumber="57">ZMUC</collectionCode>
|
||||
).
|
||||
<emphasis id="20830D1B706BFF9C28F4FEB18B24F24C" bold="true" box="[199,334,331,355]" pageId="2" pageNumber="57">Etymology.</emphasis>
|
||||
Named after the
|
||||
<typeStatus id="CD4C6FAB706BFF9C2A3CFEB18855F24C" box="[527,575,331,355]" pageId="2" pageNumber="57">type</typeStatus>
|
||||
locality, Tabarka (northwestern
|
||||
<collectingCountry id="6AE09199706BFF9C2B99FEB08E68F24D" box="[938,1026,330,354]" name="Tunisia" pageId="2" pageNumber="57">Tunisia</collectingCountry>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="5AED8282706BFF9A28F4FE9489AEF412" lastPageId="4" lastPageNumber="59" pageId="2" pageNumber="57" type="description">
|
||||
<paragraph id="1248D109706BFF9C28F4FE9488EBF2E1" blockId="2.[151,1437,258,462]" pageId="2" pageNumber="57">
|
||||
<emphasis id="20830D1B706BFF9C28F4FE948B38F2A8" bold="true" box="[199,338,366,391]" pageId="2" pageNumber="57">Description</emphasis>
|
||||
(all measurements in mm). Whitish to light brownish, 20 body rings in both sexes; length: 7–7.9 (males) and 7–8.2 (females); width of prozona 0.40–0.47 (males) and 0.37–0.52 (females); width of metazona: 0.88–0.95 (males) and 0.80–0.95 (females).
|
||||
</paragraph>
|
||||
<caption id="46888181706BFF9C28A4F8F18F56F4B3" httpUri="https://zenodo.org/record/202202/files/figure.png" pageId="2" pageNumber="57" targetBox="[215,1372,518,1779]" targetPageId="2">
|
||||
<paragraph id="1248D109706BFF9C28A4F8F18F56F4B3" blockId="2.[151,1436,1803,1948]" pageId="2" pageNumber="57">
|
||||
<emphasis id="20830D1B706BFF9C28A4F8F18B53F40E" bold="true" box="[151,313,1803,1826]" pageId="2" pageNumber="57">FIGURES 1–6.</emphasis>
|
||||
<taxonomicName id="D5F7AA8A706BFF9C290CF8F68828F40D" box="[319,578,1804,1826]" class="Diplopoda" family="Pyrgodesmidae" genus="Rharodesmus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="tabarkensis" status="sp. nov.">
|
||||
<emphasis id="20830D1B706BFF9C290CF8F68828F40D" box="[319,578,1804,1826]" italics="true" pageId="2" pageNumber="57">Rharodesmus tabarkensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="20830D1B706BFF9C2A7BF8F688EBF40E" bold="true" box="[584,641,1804,1825]" pageId="2" pageNumber="57">
|
||||
<taxonomicNameLabel id="3BB0B060706BFF9C2A7BF8F688EBF40E" box="[584,641,1804,1825]" pageId="2" pageNumber="57" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, adult, morphological details: Fig. 1 Head and first two body rings, adult male, anterior view; Fig. 2 Collum and three subsequent tergites, adult female; Fig. 3 Prozona detail: cellular scaly structure (left), transverse ridge of tooth-shaped cuticular ‘cells’ and knobby area next to metazona (right), adult female; Fig. 4 Body rings 3 and 4, showing microsculpture of prozona and tubercles of metazona, adult female; Fig.5 Tubercles on metazona 7 and 8, adult female; Fig 6 Limbus, adult male. Abbreviations:
|
||||
<emphasis id="20830D1B706BFF9C2AA1F87D88CDF4B3" bold="true" box="[658,679,1927,1948]" pageId="2" pageNumber="57">lb</emphasis>
|
||||
: posterior lobe;
|
||||
<emphasis id="20830D1B706BFF9C2B7EF87D8933F4B3" bold="true" box="[845,857,1927,1948]" pageId="2" pageNumber="57">o</emphasis>
|
||||
: ozopore;
|
||||
<emphasis id="20830D1B706BFF9C2BF5F87D89BEF4B3" bold="true" box="[966,980,1927,1948]" pageId="2" pageNumber="57">p</emphasis>
|
||||
: porostele;
|
||||
<emphasis id="20830D1B706BFF9C2C7FF87D8E3EF4B3" bold="true" box="[1100,1108,1927,1948]" pageId="2" pageNumber="57">t</emphasis>
|
||||
: caudolateral tubercle.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="1248D109706AFF9D28F6FF6D8847F233" blockId="3.[151,1437,151,861]" pageId="3" pageNumber="58">
|
||||
Head transverse; dorsal surface microgranulate, epistome with short scattered setae, labral and supra-labral surface with longer ones (
|
||||
<figureCitation id="8ACCCD8C706AFF9D29AEFF468B8AF3FB" box="[413,480,188,212]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1803,1826]" captionTargetBox="[215,1372,518,1779]" captionTargetId="figure@2.[215,1372,518,1779]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Rharodesmus tabarkensis n. sp., adult, morphological details: Fig. 1 Head and first two body rings, adult male, anterior view; Fig. 2 Collum and three subsequent tergites, adult female; Fig. 3 Prozona detail: cellular scaly structure (left), transverse ridge of tooth-shaped cuticular ‘ cells’ and knobby area next to metazona (right), adult female; Fig. 4 Body rings 3 and 4, showing microsculpture of prozona and tubercles of metazona, adult female; Fig. 5 Tubercles on metazona 7 and 8, adult female; Fig 6 Limbus, adult male. Abbreviations: lb: posterior lobe; o: ozopore; p: porostele; t: caudolateral tubercle." httpUri="https://zenodo.org/record/202202/files/figure.png" pageId="3" pageNumber="58">Fig. 1</figureCitation>
|
||||
). Antennae claviform and stout (
|
||||
<figureCitation id="8ACCCD8C706AFF9D2B6FFF4689CAF3FB" box="[860,928,188,212]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1803,1826]" captionTargetBox="[215,1372,518,1779]" captionTargetId="figure@2.[215,1372,518,1779]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Rharodesmus tabarkensis n. sp., adult, morphological details: Fig. 1 Head and first two body rings, adult male, anterior view; Fig. 2 Collum and three subsequent tergites, adult female; Fig. 3 Prozona detail: cellular scaly structure (left), transverse ridge of tooth-shaped cuticular ‘ cells’ and knobby area next to metazona (right), adult female; Fig. 4 Body rings 3 and 4, showing microsculpture of prozona and tubercles of metazona, adult female; Fig. 5 Tubercles on metazona 7 and 8, adult female; Fig 6 Limbus, adult male. Abbreviations: lb: posterior lobe; o: ozopore; p: porostele; t: caudolateral tubercle." httpUri="https://zenodo.org/record/202202/files/figure.png" pageId="3" pageNumber="58">Fig. 1</figureCitation>
|
||||
), length of antennomeres: 5=6>3>2=4>7>1. Collum (
|
||||
<figureCitation id="8ACCCD8C706AFF9D28C9FF258B2AF3D8" box="[250,320,223,247]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1803,1826]" captionTargetBox="[215,1372,518,1779]" captionTargetId="figure@2.[215,1372,518,1779]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Rharodesmus tabarkensis n. sp., adult, morphological details: Fig. 1 Head and first two body rings, adult male, anterior view; Fig. 2 Collum and three subsequent tergites, adult female; Fig. 3 Prozona detail: cellular scaly structure (left), transverse ridge of tooth-shaped cuticular ‘ cells’ and knobby area next to metazona (right), adult female; Fig. 4 Body rings 3 and 4, showing microsculpture of prozona and tubercles of metazona, adult female; Fig. 5 Tubercles on metazona 7 and 8, adult female; Fig 6 Limbus, adult male. Abbreviations: lb: posterior lobe; o: ozopore; p: porostele; t: caudolateral tubercle." httpUri="https://zenodo.org/record/202202/files/figure.png" pageId="3" pageNumber="58">Fig. 2</figureCitation>
|
||||
): anterior margin with 10 distinct lobes; dorsal surface domed with numerous small tubercles and 4 bigger ones on the posterior margin.
|
||||
</paragraph>
|
||||
<paragraph id="1248D109706AFF9D28F6FEDD8E6AF283" blockId="3.[151,1437,151,861]" pageId="3" pageNumber="58">
|
||||
Body rings: Prozona with fine longitudinal stripes on the anterior margin, followed by a scaly cellular structure covering most of the surface (
|
||||
<figureCitation id="8ACCCD8C706AFF9D29D4FEB68827F24B" box="[487,589,332,356]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1803,1826]" captionTargetBox="[215,1372,518,1779]" captionTargetId="figure@2.[215,1372,518,1779]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Rharodesmus tabarkensis n. sp., adult, morphological details: Fig. 1 Head and first two body rings, adult male, anterior view; Fig. 2 Collum and three subsequent tergites, adult female; Fig. 3 Prozona detail: cellular scaly structure (left), transverse ridge of tooth-shaped cuticular ‘ cells’ and knobby area next to metazona (right), adult female; Fig. 4 Body rings 3 and 4, showing microsculpture of prozona and tubercles of metazona, adult female; Fig. 5 Tubercles on metazona 7 and 8, adult female; Fig 6 Limbus, adult male. Abbreviations: lb: posterior lobe; o: ozopore; p: porostele; t: caudolateral tubercle." httpUri="https://zenodo.org/record/202202/files/figure.png" pageId="3" pageNumber="58">Figs 3, 4</figureCitation>
|
||||
). Posterior limit of the latter area marked by a row of tooth- shaped (dorsal side) or rectangular (ventral side) cuticular ‘cells’ forming a distinct transverse ridge. Prozona behind ridge (ca. ¼ of the length of prozona) with a regular covering of subhemisphaerical knobs.
|
||||
</paragraph>
|
||||
<paragraph id="1248D109706AFF9D28F6FE428E32F073" blockId="3.[151,1437,151,861]" pageId="3" pageNumber="58">
|
||||
Metaterga (following the terminology of
|
||||
<bibRefCitation id="7666ACF8706AFF9D2AABFE4D8954F2E0" author="Hoffman" box="[664,830,439,463]" pageId="3" pageNumber="58" refString="Hoffman, R. L. (1976) A new lophodesmid millipid from Guatemalan cave, with notes on related forms (Polydesmida: Pyrgodesmidae). Revue Suisse de Zoologie, 83 (2), 307 - 316." type="journal article" year="1976">Hoffman 1976</bibRefCitation>
|
||||
) with 4 longitudinal–two paramedian and two dorso-lateral—rows of big tubercles (
|
||||
<figureCitation id="8ACCCD8C706AFF9D29C4FE268851F2DB" box="[503,571,476,500]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1803,1826]" captionTargetBox="[215,1372,518,1779]" captionTargetId="figure@2.[215,1372,518,1779]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Rharodesmus tabarkensis n. sp., adult, morphological details: Fig. 1 Head and first two body rings, adult male, anterior view; Fig. 2 Collum and three subsequent tergites, adult female; Fig. 3 Prozona detail: cellular scaly structure (left), transverse ridge of tooth-shaped cuticular ‘ cells’ and knobby area next to metazona (right), adult female; Fig. 4 Body rings 3 and 4, showing microsculpture of prozona and tubercles of metazona, adult female; Fig. 5 Tubercles on metazona 7 and 8, adult female; Fig 6 Limbus, adult male. Abbreviations: lb: posterior lobe; o: ozopore; p: porostele; t: caudolateral tubercle." httpUri="https://zenodo.org/record/202202/files/figure.png" pageId="3" pageNumber="58">Fig. 4</figureCitation>
|
||||
) and additionally smaller ones irregularly scattered over the lateral surfaces. Paraterga surmounted of a pair of big caudolateral tubercles (
|
||||
<figureCitation id="8ACCCD8C706AFF9D2B63FE0589DDF138" box="[848,951,511,535]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1803,1826]" captionTargetBox="[215,1372,518,1779]" captionTargetId="figure@2.[215,1372,518,1779]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Rharodesmus tabarkensis n. sp., adult, morphological details: Fig. 1 Head and first two body rings, adult male, anterior view; Fig. 2 Collum and three subsequent tergites, adult female; Fig. 3 Prozona detail: cellular scaly structure (left), transverse ridge of tooth-shaped cuticular ‘ cells’ and knobby area next to metazona (right), adult female; Fig. 4 Body rings 3 and 4, showing microsculpture of prozona and tubercles of metazona, adult female; Fig. 5 Tubercles on metazona 7 and 8, adult female; Fig 6 Limbus, adult male. Abbreviations: lb: posterior lobe; o: ozopore; p: porostele; t: caudolateral tubercle." httpUri="https://zenodo.org/record/202202/files/figure.png" pageId="3" pageNumber="58">Figs 4, 5</figureCitation>
|
||||
, 7,
|
||||
<emphasis id="20830D1B706AFF9D2BD1FDFA8981F137" bold="true" box="[994,1003,512,536]" pageId="3" pageNumber="58">t</emphasis>
|
||||
). The three first body rings following the collum with 2 (sometimes 3) transversal tergal rows (
|
||||
<figureCitation id="8ACCCD8C706AFF9D2B23FDDE8939F113" box="[784,851,548,572]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1803,1826]" captionTargetBox="[215,1372,518,1779]" captionTargetId="figure@2.[215,1372,518,1779]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Rharodesmus tabarkensis n. sp., adult, morphological details: Fig. 1 Head and first two body rings, adult male, anterior view; Fig. 2 Collum and three subsequent tergites, adult female; Fig. 3 Prozona detail: cellular scaly structure (left), transverse ridge of tooth-shaped cuticular ‘ cells’ and knobby area next to metazona (right), adult female; Fig. 4 Body rings 3 and 4, showing microsculpture of prozona and tubercles of metazona, adult female; Fig. 5 Tubercles on metazona 7 and 8, adult female; Fig 6 Limbus, adult male. Abbreviations: lb: posterior lobe; o: ozopore; p: porostele; t: caudolateral tubercle." httpUri="https://zenodo.org/record/202202/files/figure.png" pageId="3" pageNumber="58">Fig. 4</figureCitation>
|
||||
), subsequent body rings with 3 rows (
|
||||
<figureCitation id="8ACCCD8C706AFF9D2CCAFDDE8F57F113" box="[1273,1341,548,572]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1803,1826]" captionTargetBox="[215,1372,518,1779]" captionTargetId="figure@2.[215,1372,518,1779]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Rharodesmus tabarkensis n. sp., adult, morphological details: Fig. 1 Head and first two body rings, adult male, anterior view; Fig. 2 Collum and three subsequent tergites, adult female; Fig. 3 Prozona detail: cellular scaly structure (left), transverse ridge of tooth-shaped cuticular ‘ cells’ and knobby area next to metazona (right), adult female; Fig. 4 Body rings 3 and 4, showing microsculpture of prozona and tubercles of metazona, adult female; Fig. 5 Tubercles on metazona 7 and 8, adult female; Fig 6 Limbus, adult male. Abbreviations: lb: posterior lobe; o: ozopore; p: porostele; t: caudolateral tubercle." httpUri="https://zenodo.org/record/202202/files/figure.png" pageId="3" pageNumber="58">Fig. 5</figureCitation>
|
||||
). Tubercles of the posteriormost body rings generally much bigger and strongly protruding caudad. Limbus narrow with regularly denticulate margin (
|
||||
<figureCitation id="8ACCCD8C706AFF9D29D8FD968859F1AB" box="[491,563,620,644]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1803,1826]" captionTargetBox="[215,1372,518,1779]" captionTargetId="figure@2.[215,1372,518,1779]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Rharodesmus tabarkensis n. sp., adult, morphological details: Fig. 1 Head and first two body rings, adult male, anterior view; Fig. 2 Collum and three subsequent tergites, adult female; Fig. 3 Prozona detail: cellular scaly structure (left), transverse ridge of tooth-shaped cuticular ‘ cells’ and knobby area next to metazona (right), adult female; Fig. 4 Body rings 3 and 4, showing microsculpture of prozona and tubercles of metazona, adult female; Fig. 5 Tubercles on metazona 7 and 8, adult female; Fig 6 Limbus, adult male. Abbreviations: lb: posterior lobe; o: ozopore; p: porostele; t: caudolateral tubercle." httpUri="https://zenodo.org/record/202202/files/figure.png" pageId="3" pageNumber="58">Fig. 6</figureCitation>
|
||||
). Paraterga well expanded with 3 lateral lobes on body ring 2 (
|
||||
<figureCitation id="8ACCCD8C706AFF9D2D27FD968F30F1AB" box="[1300,1370,620,644]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1803,1826]" captionTargetBox="[215,1372,518,1779]" captionTargetId="figure@2.[215,1372,518,1779]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Rharodesmus tabarkensis n. sp., adult, morphological details: Fig. 1 Head and first two body rings, adult male, anterior view; Fig. 2 Collum and three subsequent tergites, adult female; Fig. 3 Prozona detail: cellular scaly structure (left), transverse ridge of tooth-shaped cuticular ‘ cells’ and knobby area next to metazona (right), adult female; Fig. 4 Body rings 3 and 4, showing microsculpture of prozona and tubercles of metazona, adult female; Fig. 5 Tubercles on metazona 7 and 8, adult female; Fig 6 Limbus, adult male. Abbreviations: lb: posterior lobe; o: ozopore; p: porostele; t: caudolateral tubercle." httpUri="https://zenodo.org/record/202202/files/figure.png" pageId="3" pageNumber="58">Fig. 2</figureCitation>
|
||||
) then becoming bilobed (
|
||||
<figureCitation id="8ACCCD8C706AFF9D295CFD758BB9F188" box="[367,467,655,679]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1803,1826]" captionTargetBox="[215,1372,518,1779]" captionTargetId="figure@2.[215,1372,518,1779]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Rharodesmus tabarkensis n. sp., adult, morphological details: Fig. 1 Head and first two body rings, adult male, anterior view; Fig. 2 Collum and three subsequent tergites, adult female; Fig. 3 Prozona detail: cellular scaly structure (left), transverse ridge of tooth-shaped cuticular ‘ cells’ and knobby area next to metazona (right), adult female; Fig. 4 Body rings 3 and 4, showing microsculpture of prozona and tubercles of metazona, adult female; Fig. 5 Tubercles on metazona 7 and 8, adult female; Fig 6 Limbus, adult male. Abbreviations: lb: posterior lobe; o: ozopore; p: porostele; t: caudolateral tubercle." httpUri="https://zenodo.org/record/202202/files/figure.png" pageId="3" pageNumber="58">Figs 2, 4</figureCitation>
|
||||
), except on the poriferous rings. In latter rings, the posterior lobe bifurcating into a big lobe forming the porostele and a smaller posterior one (
|
||||
<figureCitation id="8ACCCD8C706AFF9D2B23FD4E8933F1E3" box="[784,857,692,716]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1803,1826]" captionTargetBox="[215,1372,518,1779]" captionTargetId="figure@2.[215,1372,518,1779]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Rharodesmus tabarkensis n. sp., adult, morphological details: Fig. 1 Head and first two body rings, adult male, anterior view; Fig. 2 Collum and three subsequent tergites, adult female; Fig. 3 Prozona detail: cellular scaly structure (left), transverse ridge of tooth-shaped cuticular ‘ cells’ and knobby area next to metazona (right), adult female; Fig. 4 Body rings 3 and 4, showing microsculpture of prozona and tubercles of metazona, adult female; Fig. 5 Tubercles on metazona 7 and 8, adult female; Fig 6 Limbus, adult male. Abbreviations: lb: posterior lobe; o: ozopore; p: porostele; t: caudolateral tubercle." httpUri="https://zenodo.org/record/202202/files/figure.png" pageId="3" pageNumber="58">Figs 5</figureCitation>
|
||||
, 7, p). Ozopores opening on porosteles, present on body rings 5, 7, 9, 10, 12, 13, 15 and 16 (
|
||||
<figureCitation id="8ACCCD8C706AFF9D2A56FD2D88C7F1C0" box="[613,685,727,751]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1803,1826]" captionTargetBox="[215,1372,518,1779]" captionTargetId="figure@2.[215,1372,518,1779]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Rharodesmus tabarkensis n. sp., adult, morphological details: Fig. 1 Head and first two body rings, adult male, anterior view; Fig. 2 Collum and three subsequent tergites, adult female; Fig. 3 Prozona detail: cellular scaly structure (left), transverse ridge of tooth-shaped cuticular ‘ cells’ and knobby area next to metazona (right), adult female; Fig. 4 Body rings 3 and 4, showing microsculpture of prozona and tubercles of metazona, adult female; Fig. 5 Tubercles on metazona 7 and 8, adult female; Fig 6 Limbus, adult male. Abbreviations: lb: posterior lobe; o: ozopore; p: porostele; t: caudolateral tubercle." httpUri="https://zenodo.org/record/202202/files/figure.png" pageId="3" pageNumber="58">Figs 5</figureCitation>
|
||||
, 7,
|
||||
<emphasis id="20830D1B706AFF9D2AE5FD22888EF1DF" bold="true" box="[726,740,728,752]" pageId="3" pageNumber="58">o</emphasis>
|
||||
). Legs almost invisible from above, length of podomeres: tarsus>femur>prefemur>postfemur= tibia>claw. Telson (Fig. 8): epiproct short and subtruncate, ventrally with scattered marginal setae and 4 spinnerets (Fig. 8,
|
||||
<emphasis id="20830D1B706AFF9D2A92FCDA88C6F017" bold="true" box="[673,684,800,824]" pageId="3" pageNumber="58">s</emphasis>
|
||||
) (cf.
|
||||
<bibRefCitation id="7666ACF8706AFF9D2ADEFCE5891BF018" author="Shear" box="[749,881,799,823]" pageId="3" pageNumber="58" refString="Shear, W. A. (2008) Spinnerets in the milliped order Polydesmida, and the phylogenetic significance of spinnerets in millipeds (Diplopoda). International Journal of Myriapodology, 2, 123 - 146." type="journal article" year="2008">Shear 2008</bibRefCitation>
|
||||
); paraprocts each with 2 setae, both located well away from mesal margin; hypoproct subtriangular with 2 strong setae on caudal edge.
|
||||
</paragraph>
|
||||
<paragraph id="1248D109706AFF9D28A4FA888F30F688" blockId="3.[151,1436,1392,1447]" pageId="3" pageNumber="58">
|
||||
<emphasis id="20830D1B706AFF9D28A4FA888B50F6A7" bold="true" box="[151,314,1394,1417]" pageId="3" pageNumber="58">FIGURES 7–8.</emphasis>
|
||||
<taxonomicName id="D5F7AA8A706AFF9D290CFA898829F6A6" box="[319,579,1395,1417]" class="Diplopoda" family="Pyrgodesmidae" genus="Rharodesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="tabarkensis" status="sp. nov.">
|
||||
<emphasis id="20830D1B706AFF9D290CFA898829F6A6" box="[319,579,1395,1417]" italics="true" pageId="3" pageNumber="58">Rharodesmus tabarkensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="20830D1B706AFF9D2A7BFA8988EBF6A7" bold="true" box="[584,641,1395,1416]" pageId="3" pageNumber="58">
|
||||
<taxonomicNameLabel id="3BB0B060706AFF9D2A7BFA8988EBF6A7" box="[584,641,1395,1416]" pageId="3" pageNumber="58" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, close-up of ozopore and Telson: Fig. 7 Ozopore on the 16th tergite, adult male; Fig. 8 Telson, adult female. Abbreviations: lb: posterior lobe; o: ozopore, s: spinneret-like setae; t: caudolateral tubercle.
|
||||
</paragraph>
|
||||
<paragraph id="1248D109706AFF9D28F4F9E589DBF587" blockId="3.[151,1436,1564,1704]" pageId="3" pageNumber="58">
|
||||
<emphasis id="20830D1B706AFF9D28F4F9E58B89F518" bold="true" box="[199,483,1567,1591]" pageId="3" pageNumber="58">Male sexual characters.</emphasis>
|
||||
Tarsus of 1st and 2nd pairs of legs with ca 17 stout, spatulate setae; tibia of these legpairs with a few similar setae (Fig. 9). Coxal gonapophyses on 2nd pair of legs flattened (Fig. 10). Prefemur modified on the 7th and 8th pairs of male legs (Fig. 11): apically protruding into a pointed knob surmounted by a stout and contorted seta emerging from a densely poriferous basal area (Fig. 12).
|
||||
</paragraph>
|
||||
<paragraph id="1248D109706DFF9A28A4FAE98897F643" blockId="4.[151,1437,1299,1388]" pageId="4" pageNumber="59">
|
||||
<emphasis id="20830D1B706DFF9A28A4FAE98B2DF606" bold="true" box="[151,327,1299,1322]" pageId="4" pageNumber="59">FIGURES 9–12.</emphasis>
|
||||
<taxonomicName id="D5F7AA8A706DFF9A297DFAEE8839F605" box="[334,595,1300,1322]" class="Diplopoda" family="Pyrgodesmidae" genus="Rharodesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="tabarkensis" status="sp. nov.">
|
||||
<emphasis id="20830D1B706DFF9A297DFAEE8839F605" box="[334,595,1300,1322]" italics="true" pageId="4" pageNumber="59">Rharodesmus tabarkensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="20830D1B706DFF9A2A69FAEE88F9F606" bold="true" box="[602,659,1300,1321]" pageId="4" pageNumber="59">
|
||||
<taxonomicNameLabel id="3BB0B060706DFF9A2A69FAEE88F9F606" box="[602,659,1300,1321]" pageId="4" pageNumber="59" rank="species">n. sp.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, secondary sexual characters in male: Fig. 9 Spatulate setae on right second leg; Fig. 10 Flattened coxal gonapophyses on second leg pair; Fig. 11 Modified prefemora on the 7th and 8th pairs of legs; Fig. 12 close up of prefemoral process and seta on the 7th left leg.
|
||||
</paragraph>
|
||||
<paragraph id="1248D109706DFF9A28F6FA4789AEF412" blockId="4.[151,1436,1469,1853]" pageId="4" pageNumber="59">
|
||||
Gonopods: in situ with crossing telopodites (Figs 11, 13); coxa voluminous, covering most of the lateral side of the telopodite (Figs 11, 13,
|
||||
<emphasis id="20830D1B706DFF9A29FCFA188BB1F6D5" bold="true" box="[463,475,1506,1530]" pageId="4" pageNumber="59">c</emphasis>
|
||||
), hemispherical with a granulate surface and a rounded-triangular smooth distal process (
|
||||
<figureCitation id="8ACCCD8C706DFF9A28E5F9FF8B39F532" box="[214,339,1541,1565]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,1413,1436]" captionTargetBox="[202,1375,195,1388]" captionTargetId="figure@5.[202,1385,195,1391]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Rharodesmus tabarkensis n. sp., gonopodal structures: Fig. 13 Gonopods in situ, profile; Fig. 14 Right gonopod, mesal view; Fig. 15 Close up of coxal distal process and hemispherical hump; Fig. 16 Left gonopod, antero-mesal view. Abbreviations: A: process A; B: process B; c: coxa, ca: cannula; cp: conical process, h: setose coxal hump; r: rounded process; so: solenomere; p: prefemoral part; sp: smooth distal process." httpUri="https://zenodo.org/record/202203/files/figure.png" pageId="4" pageNumber="59">Fig. 14, 15</figureCitation>
|
||||
,
|
||||
<emphasis id="20830D1B706DFF9A2953F9FF8B10F532" bold="true" box="[352,378,1541,1565]" pageId="4" pageNumber="59">sp</emphasis>
|
||||
) surmounting a small setose hump (
|
||||
<figureCitation id="8ACCCD8C706DFF9A2B26F9FF89F9F532" box="[789,915,1541,1565]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,1413,1436]" captionTargetBox="[202,1375,195,1388]" captionTargetId="figure@5.[202,1385,195,1391]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Rharodesmus tabarkensis n. sp., gonopodal structures: Fig. 13 Gonopods in situ, profile; Fig. 14 Right gonopod, mesal view; Fig. 15 Close up of coxal distal process and hemispherical hump; Fig. 16 Left gonopod, antero-mesal view. Abbreviations: A: process A; B: process B; c: coxa, ca: cannula; cp: conical process, h: setose coxal hump; r: rounded process; so: solenomere; p: prefemoral part; sp: smooth distal process." httpUri="https://zenodo.org/record/202203/files/figure.png" pageId="4" pageNumber="59">Fig. 14, 15</figureCitation>
|
||||
,
|
||||
<emphasis id="20830D1B706DFF9A2B93F9FF89C5F532" bold="true" box="[928,943,1541,1565]" pageId="4" pageNumber="59">h</emphasis>
|
||||
), median margin with scattered short setae. Basal part of telopodite consisting of a big subrectangular, transverse, strongly granulated and setose ‘prefemoral’ part (
|
||||
<figureCitation id="8ACCCD8C706DFF9A28E1F9B78B4EF54A" box="[210,292,1613,1637]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,1413,1436]" captionTargetBox="[202,1375,195,1388]" captionTargetId="figure@5.[202,1385,195,1391]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Rharodesmus tabarkensis n. sp., gonopodal structures: Fig. 13 Gonopods in situ, profile; Fig. 14 Right gonopod, mesal view; Fig. 15 Close up of coxal distal process and hemispherical hump; Fig. 16 Left gonopod, antero-mesal view. Abbreviations: A: process A; B: process B; c: coxa, ca: cannula; cp: conical process, h: setose coxal hump; r: rounded process; so: solenomere; p: prefemoral part; sp: smooth distal process." httpUri="https://zenodo.org/record/202203/files/figure.png" pageId="4" pageNumber="59">Fig. 14</figureCitation>
|
||||
,
|
||||
<emphasis id="20830D1B706DFF9A2904F9B78B2CF54A" bold="true" box="[311,326,1613,1637]" pageId="4" pageNumber="59">p</emphasis>
|
||||
) linked to the coxa by a small cannula (
|
||||
<figureCitation id="8ACCCD8C706DFF9A2B34F9B78933F54A" box="[775,857,1613,1637]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,1413,1436]" captionTargetBox="[202,1375,195,1388]" captionTargetId="figure@5.[202,1385,195,1391]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Rharodesmus tabarkensis n. sp., gonopodal structures: Fig. 13 Gonopods in situ, profile; Fig. 14 Right gonopod, mesal view; Fig. 15 Close up of coxal distal process and hemispherical hump; Fig. 16 Left gonopod, antero-mesal view. Abbreviations: A: process A; B: process B; c: coxa, ca: cannula; cp: conical process, h: setose coxal hump; r: rounded process; so: solenomere; p: prefemoral part; sp: smooth distal process." httpUri="https://zenodo.org/record/202203/files/figure.png" pageId="4" pageNumber="59">Fig. 14</figureCitation>
|
||||
,
|
||||
<emphasis id="20830D1B706DFF9A2B56F9B789EAF54A" bold="true" box="[869,896,1613,1637]" pageId="4" pageNumber="59">ca</emphasis>
|
||||
); median margin of ‘prefemoral’ part with stout setae, becoming longer anteriad (
|
||||
<figureCitation id="8ACCCD8C706DFF9A2A23F98B880EF5A6" box="[528,612,1649,1673]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,1413,1436]" captionTargetBox="[202,1375,195,1388]" captionTargetId="figure@5.[202,1385,195,1391]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Rharodesmus tabarkensis n. sp., gonopodal structures: Fig. 13 Gonopods in situ, profile; Fig. 14 Right gonopod, mesal view; Fig. 15 Close up of coxal distal process and hemispherical hump; Fig. 16 Left gonopod, antero-mesal view. Abbreviations: A: process A; B: process B; c: coxa, ca: cannula; cp: conical process, h: setose coxal hump; r: rounded process; so: solenomere; p: prefemoral part; sp: smooth distal process." httpUri="https://zenodo.org/record/202203/files/figure.png" pageId="4" pageNumber="59">Fig. 14</figureCitation>
|
||||
). ‘Post-prefemoral’ part of telopodite elongated and distally divided in a short membranous rounded cylindrical branch (
|
||||
<figureCitation id="8ACCCD8C706DFF9A2A9DF96F895AF582" box="[686,816,1685,1709]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,1413,1436]" captionTargetBox="[202,1375,195,1388]" captionTargetId="figure@5.[202,1385,195,1391]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Rharodesmus tabarkensis n. sp., gonopodal structures: Fig. 13 Gonopods in situ, profile; Fig. 14 Right gonopod, mesal view; Fig. 15 Close up of coxal distal process and hemispherical hump; Fig. 16 Left gonopod, antero-mesal view. Abbreviations: A: process A; B: process B; c: coxa, ca: cannula; cp: conical process, h: setose coxal hump; r: rounded process; so: solenomere; p: prefemoral part; sp: smooth distal process." httpUri="https://zenodo.org/record/202203/files/figure.png" pageId="4" pageNumber="59">Figs 14, 16</figureCitation>
|
||||
,
|
||||
<emphasis id="20830D1B706DFF9A2B0EF96F8920F582" bold="true" box="[829,842,1685,1709]" pageId="4" pageNumber="59">r</emphasis>
|
||||
) and 2 processes of the same length: process A subrectangular (
|
||||
<figureCitation id="8ACCCD8C706DFF9A2910F9438BC9F5FE" box="[291,419,1721,1745]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,1413,1436]" captionTargetBox="[202,1375,195,1388]" captionTargetId="figure@5.[202,1385,195,1391]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Rharodesmus tabarkensis n. sp., gonopodal structures: Fig. 13 Gonopods in situ, profile; Fig. 14 Right gonopod, mesal view; Fig. 15 Close up of coxal distal process and hemispherical hump; Fig. 16 Left gonopod, antero-mesal view. Abbreviations: A: process A; B: process B; c: coxa, ca: cannula; cp: conical process, h: setose coxal hump; r: rounded process; so: solenomere; p: prefemoral part; sp: smooth distal process." httpUri="https://zenodo.org/record/202203/files/figure.png" pageId="4" pageNumber="59">Figs 14, 16</figureCitation>
|
||||
,
|
||||
<emphasis id="20830D1B706DFF9A299CF9438BA9F5FD" bold="true" box="[431,451,1721,1746]" pageId="4" pageNumber="59">A</emphasis>
|
||||
), folded interolaterad and distally serrated, especially on the margin; process B abruptly expanded distad (
|
||||
<figureCitation id="8ACCCD8C706DFF9A2951F9278BD2F5DA" box="[354,440,1757,1781]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,1413,1436]" captionTargetBox="[202,1375,195,1388]" captionTargetId="figure@5.[202,1385,195,1391]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Rharodesmus tabarkensis n. sp., gonopodal structures: Fig. 13 Gonopods in situ, profile; Fig. 14 Right gonopod, mesal view; Fig. 15 Close up of coxal distal process and hemispherical hump; Fig. 16 Left gonopod, antero-mesal view. Abbreviations: A: process A; B: process B; c: coxa, ca: cannula; cp: conical process, h: setose coxal hump; r: rounded process; so: solenomere; p: prefemoral part; sp: smooth distal process." httpUri="https://zenodo.org/record/202203/files/figure.png" pageId="4" pageNumber="59">Fig. 16</figureCitation>
|
||||
,
|
||||
<emphasis id="20830D1B706DFF9A29F4F9278BB0F5DA" bold="true" box="[455,474,1757,1781]" pageId="4" pageNumber="59">B</emphasis>
|
||||
) and bifurcating into an elongated smooth conical process (
|
||||
<figureCitation id="8ACCCD8C706DFF9A2CA4F9278F75F5DA" box="[1175,1311,1757,1781]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,1413,1436]" captionTargetBox="[202,1375,195,1388]" captionTargetId="figure@5.[202,1385,195,1391]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Rharodesmus tabarkensis n. sp., gonopodal structures: Fig. 13 Gonopods in situ, profile; Fig. 14 Right gonopod, mesal view; Fig. 15 Close up of coxal distal process and hemispherical hump; Fig. 16 Left gonopod, antero-mesal view. Abbreviations: A: process A; B: process B; c: coxa, ca: cannula; cp: conical process, h: setose coxal hump; r: rounded process; so: solenomere; p: prefemoral part; sp: smooth distal process." httpUri="https://zenodo.org/record/202203/files/figure.png" pageId="4" pageNumber="59">Figs 14, 16</figureCitation>
|
||||
, cp) and a shorter truncated one (
|
||||
<figureCitation id="8ACCCD8C706DFF9A29A0F8FB8B8FF436" box="[403,485,1793,1817]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,1413,1436]" captionTargetBox="[202,1375,195,1388]" captionTargetId="figure@5.[202,1385,195,1391]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Rharodesmus tabarkensis n. sp., gonopodal structures: Fig. 13 Gonopods in situ, profile; Fig. 14 Right gonopod, mesal view; Fig. 15 Close up of coxal distal process and hemispherical hump; Fig. 16 Left gonopod, antero-mesal view. Abbreviations: A: process A; B: process B; c: coxa, ca: cannula; cp: conical process, h: setose coxal hump; r: rounded process; so: solenomere; p: prefemoral part; sp: smooth distal process." httpUri="https://zenodo.org/record/202203/files/figure.png" pageId="4" pageNumber="59">Fig. 16</figureCitation>
|
||||
,
|
||||
<emphasis id="20830D1B706DFF9A29C2F8F88860F435" bold="true" box="[497,522,1794,1818]" pageId="4" pageNumber="59">so</emphasis>
|
||||
) forming the solenomere; seminal groove arising from the bases of processes (A) and (B) and running into a lateral furrow up to the tip of the solenomere.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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