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<document ID-DOI="http://dx.doi.org/10.3897/zsE.95.36788" ID-GBIF-Dataset="bc2389bd-52b8-4411-9ef5-f37d12db4898" ID-Pensoft-Pub="1860-0743-2-507" ID-Pensoft-UUID="F3BFD206564F59C4967CE6267990A52E" ID-ZooBank="10.3897/zsE.95.36788" ModsDocID="1860-0743-2-507" checkinTime="1571900144293" checkinUser="pensoft" docAuthor="Brito, Pamella Silva de, Guimaraes, Erick Cristofore, Carvalho-Costa, Luis Fernando &amp; Ottoni, Felipe Polivanov" docDate="2019" docId="00D38DF4C439548A9820A10076073CBC" docLanguage="en" docName="ZoosystEvol 95(2): 507-516" docOrigin="Zoosystematics and Evolution 95 (2)" docSource="http://dx.doi.org/" docTitle="Aphyocharax brevicaudatus Brito, Guimaraes, Carvalho-Costa &amp; Ottoni, 2019, sp. nov." docType="treatment" docUuid="C5D86CB2-B51B-4B45-AFF7-6E483533B680" docUuidSource="ZooBank" docVersion="5" id="F3BFD206564F59C4967CE6267990A52E" lastPageNumber="507" masterDocId="F3BFD206564F59C4967CE6267990A52E" masterDocTitle="A new species of Aphyocharax Guenther, 1868 (Characiformes, Characidae) from the Maracacume river basin, eastern Amazon" masterLastPageNumber="516" masterPageNumber="507" pageNumber="507" updateTime="1684346104069" updateUser="felipe">
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<mods:title id="0F0E96F347B5F012C2D0E075CD6963D4">A new species of Aphyocharax Guenther, 1868 (Characiformes, Characidae) from the Maracacume river basin, eastern Amazon</mods:title>
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<mods:namePart id="7B18270537C70F511DC522943B136F68">Brito, Pamella Silva de</mods:namePart>
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<mods:namePart id="6CB2ADE7ADBD971A165AF03AB9D0B095">Guimaraes, Erick Cristofore</mods:namePart>
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<mods:namePart id="1A0662BF16D122F611A4DFA0839023D5">Carvalho-Costa, Luis Fernando</mods:namePart>
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<mods:namePart id="922F5011B3AD33F41CC76B06E6D612A9">Ottoni, Felipe Polivanov</mods:namePart>
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<treatment id="00D38DF4C439548A9820A10076073CBC" ID-GBIF-Taxon="159834676" LSID="urn:lsid:zoobank.org:act:C5D86CB2-B51B-4B45-AFF7-6E483533B680" httpUri="http://treatment.plazi.org/id/00D38DF4C439548A9820A10076073CBC" lastPageNumber="507" pageId="0" pageNumber="507">
<subSubSection id="A57C2284006F9C89461DC8112339910C" pageId="0" pageNumber="507" type="nomenclature">
<paragraph id="8494002C4C4841B137849BFC390C4C06" pageId="0" pageNumber="507">
<taxonomicName id="FD74FF10FAC918BF7A324C018AFBAC94" LSID="00D38DF4-C439-548A-9820-A10076073CBC" family="Characidae" genus="Aphyocharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphyocharax brevicaudatus" order="Characiformes" pageId="0" pageNumber="507" phylum="Chordata" rank="species" species="brevicaudatus">Aphyocharax brevicaudatus</taxonomicName>
<taxonomicNameLabel id="E91FFDE63A84C546268419CB712413BA" pageId="0" pageNumber="507">sp. nov.</taxonomicNameLabel>
<figureCitation id="9F74653C034A608F6CD73769A3F949A3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Aphyocharax brevicaudatus sp. nov. a. CICCAA 02293, holotype (male), 35.9 mm SL; b. CICCAA 02294, paratype (female), 32.4 mm SL, Brazil: Maranhao state: Maracacume river basin. (Photographed by Erick Guimaraes)." figureDoi=".figure1" httpUri="https://binary.pensoft.net/fig/349754" pageId="0" pageNumber="507" tableDoi=".figure1">Figs 1</figureCitation>
,
<figureCitation id="2A746E8FC6FB897ED0D3C3E84D9AB0A3" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Caudal-fin of Aphyocharax brevicaudatus, holotype, CICCAA 02293, (male)." figureDoi=".figure2" httpUri="https://binary.pensoft.net/fig/349755" pageId="0" pageNumber="507" tableDoi=".figure2">2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C78DE35C59435F60CEE713AE1D641A13" pageId="0" pageNumber="507" type="holotype">
<paragraph id="3BB28BA17D661CB9754AE91B6D05757E" pageId="0" pageNumber="507">Holotype.</paragraph>
<paragraph id="DE6BC11B8DE81ADFB08FCDEE1563FB10" pageId="0" pageNumber="507">
CICCAA 02293, (male) 35.9 mm SL, Brazil,
<normalizedToken id="D322C797608C22C1C80B1077EB60AD11" originalValue="Maranhão">Maranhao</normalizedToken>
state,
<normalizedToken id="377680C7D06A2E04E9D48F570B02926B" originalValue="Maracaçumé">Maracacume</normalizedToken>
municipality,
<normalizedToken id="9160EFEA6C5FFA0571A25D537BFE6AC7" originalValue="Maracaçumé">Maracacume</normalizedToken>
River,
<geoCoordinate id="79A9AB3FC3205C30B4EDBB6968A5B085" degrees="2" direction="south" minutes="3" orientation="latitude" precision="15" seconds="14" value="-2.0538888">2°3'14&quot;S</geoCoordinate>
,
<geoCoordinate id="F0083FD158E0BC1DEA8E8997B455CFC5" degrees="45" direction="west" minutes="57" orientation="longitude" precision="15" seconds="16" value="-45.954445">45°57'16&quot;W</geoCoordinate>
; 29 Jun 2018, E.C.
<normalizedToken id="5A1CF120BBC07D7700575E82BBEB563D" originalValue="Guimarães">Guimaraes</normalizedToken>
and P.S. Brito.
</paragraph>
</subSubSection>
<subSubSection id="EF4BB1E7B2D8B90C290EF0DB7902A2D6" pageId="0" pageNumber="507" type="paratypes">
<paragraph id="78062F9D362FA1A394A62BA027DF19EC" pageId="0" pageNumber="507">Paratypes.</paragraph>
<paragraph id="51DA5B2C485A058EB4187676228045B6" pageId="0" pageNumber="507">
All from Brazil,
<normalizedToken id="BACB731FC7E7251E4264685300CCA8E8" originalValue="Maranhão">Maranhao</normalizedToken>
state: CICCAA 02294, 1 (female), 32.4 mm SL, CICCAA 02295, 35 (males), 20.9-31.7 mm SL,CICCAA 02296, 94 (females), 21-32.1 mm SL, CICCAA 02297, 30 (females) C&amp;S, 22.2-30.8 mm SL, CICCAA 02312, 2 (males) C&amp;S, 28.3-32.1 mm SL, UFRJ 11746, 10 (female), 24.2-30.2 mm SL; all collected with holotype.
</paragraph>
</subSubSection>
<subSubSection id="7200EB4DEEE6024BEB83889F21589460" pageId="0" pageNumber="507" type="diagnosis">
<paragraph id="0E818BDF46F85F782E7239C0AFC64BF6" pageId="0" pageNumber="507">Diagnosis (PAA).</paragraph>
<paragraph id="1657CB081B5B953ED4FCD8A59581DADC" pageId="0" pageNumber="507">
<taxonomicName id="A415FE05C561A2E4AAD958DC1569215F" family="Characidae" genus="Aphyocharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphyocharax brevicaudatus" order="Characiformes" pageId="0" pageNumber="507" phylum="Chordata" rank="species" species="brevicaudatus">
<emphasis id="29DF9D1F056BEEDA79067FD5267887B7" italics="true" pageId="0" pageNumber="507">Aphyocharax brevicaudatus</emphasis>
</taxonomicName>
sp. nov. differs from all its congeners by possessing the upper lobe of the caudal fin longer than the lower lobe in mature males (vs upper and lower lobes similar in length, see
<figureCitation id="B6799F49E1B2B2707F9937388A363E1D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Aphyocharax brevicaudatus sp. nov. a. CICCAA 02293, holotype (male), 35.9 mm SL; b. CICCAA 02294, paratype (female), 32.4 mm SL, Brazil: Maranhao state: Maracacume river basin. (Photographed by Erick Guimaraes)." figureDoi=".figure1" httpUri="https://binary.pensoft.net/fig/349754" pageId="0" pageNumber="507" tableDoi=".figure1">Figs 1</figureCitation>
,
<figureCitation id="014EBF749AD0D8A0BC7374EBF77CBF78" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Caudal-fin of Aphyocharax brevicaudatus, holotype, CICCAA 02293, (male)." figureDoi=".figure2" httpUri="https://binary.pensoft.net/fig/349755" pageId="0" pageNumber="507" tableDoi=".figure2">2</figureCitation>
;
<bibRefCitation id="D8D6B9D471B7B776CADBFF8A545ED5C8" DOI="https://doi.org/10.1016/j.ympev.2012.04.007" author="Tagliacollo, VA" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="507" pagination="297 - 307" refId="B36" refString="Tagliacollo, VA, Souza-Lima, R, Benine, RC, Oliveira, C, 2012. Molecular phylogeny of Aphyocharacinae (Characiformes, Characidae) with morphological diagnoses for the subfamily and recognized genera. Molecular Phylogenetics and Evolution 64: 297 - 307" title="Molecular phylogeny of Aphyocharacinae (Characiformes, Characidae) with morphological diagnoses for the subfamily and recognized genera." url="https://doi.org/10.1016/j.ympev.2012.04.007" volume="64" year="2012">Tagliacollo et al. 2012</bibRefCitation>
: fig.4). Additionally, the new species is distinguished from
<taxonomicName id="9B575EF194A51A88CB8CFE526D73B5BF" family="Characidae" genus="Aphyocharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphyocharax avary" order="Characiformes" pageId="0" pageNumber="507" phylum="Chordata" rank="species" species="avary">
<emphasis id="45163771AFF07488648834412A3DD389" italics="true" pageId="0" pageNumber="507">Aphyocharax avary</emphasis>
</taxonomicName>
and
<taxonomicName id="01E4ADFBFC3DB25FAFF439A6D1451396" lsidName="A. pusillus" pageId="0" pageNumber="507" rank="species" species="pusillus">
<emphasis id="ED1486E6718DD5BA1AA27A2E17CDCE2F" italics="true" pageId="0" pageNumber="507">A. pusillus</emphasis>
</taxonomicName>
by having hyaline middle caudal-fin rays (vs black or dark brown middle caudal-fin rays,
<bibRefCitation id="A1C560BEFBA74043E056CB900403F111" DOI="https://doi.org/10.3897/zse.94.28201" author="Brito, PS" journalOrPublisher="Zoosystematics and Evolution" pageId="0" pageNumber="507" pagination="393 - 399" refId="B3" refString="Brito, PS, Guimaraes, EC, Katz, AM, Piorski, NM, Ottoni, FP, 2018. Taxonomic status of Aphyocharaxavary Fowler 1913, Aphyocharaxpusillus Guenther 1868 and Chirodonalburnus Guenther 1869 (Characiformes: Characidae). Zoosystematics and Evolution 94: 393 - 399" title="Taxonomic status of Aphyocharaxavary Fowler 1913, Aphyocharaxpusillus Guenther 1868 and Chirodonalburnus Guenther 1869 (Characiformes: Characidae)." url="https://doi.org/10.3897/zse.94.28201" volume="94" year="2018">Brito et al. 2018</bibRefCitation>
: fig. 3); from
<taxonomicName id="A4A73EF5ADF7685C5EF9C26DA1BCAD20" family="Characidae" genus="Aphyocharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphyocharax colifax" order="Characiformes" pageId="0" pageNumber="507" phylum="Chordata" rank="species" species="colifax">
<emphasis id="414C15C4CB8973E5EAB6590297032ACA" italics="true" pageId="0" pageNumber="507">Aphyocharax colifax</emphasis>
</taxonomicName>
,
<taxonomicName id="FEDF3D8472E081C80B2DE6B32F3F3D0A" lsidName="A. yekwanae" pageId="0" pageNumber="507" rank="species" species="yekwanae">
<emphasis id="84C6735173C766BF870CB81500B1AA37" italics="true" pageId="0" pageNumber="507">A. yekwanae</emphasis>
</taxonomicName>
, and
<taxonomicName id="6F4F38CDB79209B4904688BE41118EB9" lsidName="A. rathbuni" pageId="0" pageNumber="507" rank="species" species="rathbuni">
<emphasis id="1E633C6939EB235BD9007EC5E48DB527" italics="true" pageId="0" pageNumber="507">A. rathbuni</emphasis>
</taxonomicName>
by having caudal-fin light red colouration never surpassing the vertical line of the adipose-fin (vs red colouration extending to the lateral midline of body,
<bibRefCitation id="A194C8527D323E847699741383513605" author="Willink, PW" journalOrPublisher="Ichthyological Exploration of Freshwaters" pageId="0" pageNumber="507" pagination="1 - 8" refId="B41" refString="Willink, PW, Chernoff, B, Machado-Allison, A, Provenzano, F, Petry, P, 2003. Aphyocharaxyekwanae, a new species of bloodfin tetra (Teleostei: Characiformes: Characidae) from the Guyana Shield of Venezuela. Ichthyological Exploration of Freshwaters 14: 1 - 8" title="Aphyocharaxyekwanae, a new species of bloodfin tetra (Teleostei: Characiformes: Characidae) from the Guyana Shield of Venezuela." volume="14" year="2003">Willink et al. 2003</bibRefCitation>
: fig. 1); from
<taxonomicName id="01DEC1433CB3A6C64FCD3AA0AF81FA16" lsidName="A. gracilis" pageId="0" pageNumber="507" rank="species" species="gracilis">
<emphasis id="465A0212C14DD842C591B46860C9979D" italics="true" pageId="0" pageNumber="507">A. gracilis</emphasis>
</taxonomicName>
by having a larger body depth at dorsal-fin origin (body depth), 24.5-29.2% SL (vs 20.1-20.6% SL); and from
<taxonomicName id="D3C32A18B603290B7D52598B21229352" lsidName="A. pusillus" pageId="0" pageNumber="507" rank="species" species="pusillus">
<emphasis id="8C3A66AF1265B44BDD8F2E6539473E09" italics="true" pageId="0" pageNumber="507">A. pusillus</emphasis>
</taxonomicName>
by having teeth along 2/3 of the maxillary extension (vs along proximal half of the bone,
<bibRefCitation id="FCCAE11C941C01371468F3694430D22A" DOI="https://doi.org/10.3897/zse.94.28201" author="Brito, PS" journalOrPublisher="Zoosystematics and Evolution" pageId="0" pageNumber="507" pagination="393 - 399" refId="B3" refString="Brito, PS, Guimaraes, EC, Katz, AM, Piorski, NM, Ottoni, FP, 2018. Taxonomic status of Aphyocharaxavary Fowler 1913, Aphyocharaxpusillus Guenther 1868 and Chirodonalburnus Guenther 1869 (Characiformes: Characidae). Zoosystematics and Evolution 94: 393 - 399" title="Taxonomic status of Aphyocharaxavary Fowler 1913, Aphyocharaxpusillus Guenther 1868 and Chirodonalburnus Guenther 1869 (Characiformes: Characidae)." url="https://doi.org/10.3897/zse.94.28201" volume="94" year="2018">Brito et al. 2018</bibRefCitation>
: fig. 4).
</paragraph>
<caption id="2B53E4BCBFBFB79E48E326EBB78BB9B4" doi=".figure1" httpUri="https://binary.pensoft.net/fig/349754" pageId="0" pageNumber="507" start="Figure 1" startId="F1">
<paragraph id="C9D61CAAC26F9A843E1002AACE86416E" pageId="0" pageNumber="507">
<emphasis id="D501900C8F618FAF6BC88CCC242633ED" bold="true" pageId="0" pageNumber="507">Figure 1.</emphasis>
<taxonomicName id="C8085F651B6CF0CF8EDC227C694A8871" family="Characidae" genus="Aphyocharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphyocharax brevicaudatus" order="Characiformes" pageId="0" pageNumber="507" phylum="Chordata" rank="species" species="brevicaudatus">
<emphasis id="B04CF2F0B57EBDCDDB4EF04ABC65E7FA" italics="true" pageId="0" pageNumber="507">Aphyocharax brevicaudatus</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="F136A7E3FF37540400EB731E55821230" bold="true" pageId="0" pageNumber="507">a.</emphasis>
CICCAA 02293, holotype (male), 35.9 mm SL;
<emphasis id="E3FB18FF2652F85536A346F603672405" bold="true" pageId="0" pageNumber="507">b.</emphasis>
CICCAA 02294, paratype (female), 32.4 mm SL, Brazil:
<normalizedToken id="9BF9388C959E4586A69B962782ED6FC3" originalValue="Maranhão">Maranhao</normalizedToken>
state:
<normalizedToken id="9974E0BEC4CFDD1A70CD0F84D605436E" originalValue="Maracaçumé">Maracacume</normalizedToken>
river basin. (Photographed by Erick
<normalizedToken id="89A7F7B94C9DF304C884FEA1F1C41C3E" originalValue="Guimarães">Guimaraes</normalizedToken>
).
</paragraph>
</caption>
<caption id="A400B703D2C5CDA673EBFD5B51C82890" doi=".figure2" httpUri="https://binary.pensoft.net/fig/349755" pageId="0" pageNumber="507" start="Figure 2" startId="F2">
<paragraph id="8F85D203CD449C03833775A6A831C7C5" pageId="0" pageNumber="507">
<emphasis id="BEFCC1CE9C72B9D756CAD5613852FE30" bold="true" pageId="0" pageNumber="507">Figure 2.</emphasis>
Caudal-fin of
<taxonomicName id="F6CA3986DA4E526CB70E432AEC2B6B6A" family="Characidae" genus="Aphyocharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphyocharax brevicaudatus" order="Characiformes" pageId="0" pageNumber="507" phylum="Chordata" rank="species" species="brevicaudatus">
<emphasis id="B706BAB8FE72FD61EF16ABDE60EDEFA8" italics="true" pageId="0" pageNumber="507">Aphyocharax brevicaudatus</emphasis>
</taxonomicName>
, holotype, CICCAA 02293, (male).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="7D46339F9431662F2A552035AEC1CFDC" pageId="0" pageNumber="507" type="description">
<paragraph id="61CC8AF3230E098F20D21CA53F5FDD53" pageId="0" pageNumber="507">Description.</paragraph>
<paragraph id="979552420CC52862BCF7E2120909BF7A" pageId="0" pageNumber="507">
Morphometric data is presented in Table
<tableCitation id="539B61AB92A38437335562E7960EA53F" captionStartId="T2" httpUri="http://table.plazi.org/id/3EFCC811FE6F0397D03F2972FBFDBF6D" pageId="0" pageNumber="507" tableUuid="3EFCC811FE6F0397D03F2972FBFDBF6D">2</tableCitation>
. Body shape is generally fusiform, slightly elongate, greatest body depth slightly anterior to dorsal-fin base; dorsal body profile straight or slightly convex from snout to vertical through anterior nostrils; straight or slightly convex from posterior nostrils to tip of supraoccipital bone; straight or slightly convex from this point to dorsal-fin origin; slightly convex along dorsal-fin base; postdorsal profile straight from base of last dorsal-fin ray to adipose-fin origin; slightly concave from adipose-fin to end of caudal peduncle; ventral profile convex from snout to pelvic-fin insertion; straight or slightly convex from this point to anal-fin origin; straight along anal-fin base; long snout, with its length larger than orbital diameter; five infraorbital bones; fourth infraorbital absent and sixth infraorbital reduced; posterior border of maxilla rounded, extending vertically through anterior margin of orbit, not reaching third infraorbital.
</paragraph>
<caption id="3EFCC811FE6F0397D03F2972FBFDBF6D" ID-Table-UUID="3EFCC811FE6F0397D03F2972FBFDBF6D" httpUri="http://table.plazi.org/id/3EFCC811FE6F0397D03F2972FBFDBF6D" pageId="0" pageNumber="507" start="Table 2" startId="T2">
<paragraph id="81A0ECCF6127BDFFF59344931E521C5C" pageId="0" pageNumber="507">
<emphasis id="36909077EC16B231A71F0C0320BEEFB0" bold="true" pageId="0" pageNumber="507">Table 2.</emphasis>
Morphometric data (
<emphasis id="B169C40EA406262A1234934B52D68A08" italics="true" pageId="0" pageNumber="507">N</emphasis>
= 141) of the holotype and paratypes of
<taxonomicName id="65EBCDB3A635376DCB43C8A9FC6E6406" family="Characidae" genus="Aphyocharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphyocharax brevicaudatus" order="Characiformes" pageId="0" pageNumber="507" phylum="Chordata" rank="species" species="brevicaudatus">
<emphasis id="56D10EB63130B12478C9A28DE6570B4C" italics="true" pageId="0" pageNumber="507">Aphyocharax brevicaudatus</emphasis>
</taxonomicName>
sp. nov. from the
<normalizedToken id="F04CA18725A62268CA73CDAF03B01DB9" originalValue="Maracaçumé">Maracacume</normalizedToken>
river basin. SD: Standard deviation.
</paragraph>
</caption>
<paragraph id="5D768F5380907A1B93581D1228879D6D" pageId="0" pageNumber="507">
<table id="949628357E538E11D5ECEBC0BE564E39" pageId="0" pageNumber="507">
<tr id="B84EC7143D5E648F3B84B063756648FD" pageId="0" pageNumber="507">
<th id="1714897CEF053F9B15B839D33C528B4C" colspan="1" isEmpty="true" pageId="0" pageNumber="507" rowspan="1">-</th>
<th id="59AD5104114B30E9774F6A2B8A87D368" colspan="1" pageId="0" pageNumber="507" rowspan="1">Holotype (Male)</th>
<th id="D179455BF7E76CD253D23C648B53FFFB" colspan="1" pageId="0" pageNumber="507" rowspan="1">Paratypes (Male) N = 35</th>
<th id="0A99022D6CA9599FFA9AFD2A3E130C85" colspan="1" pageId="0" pageNumber="507" rowspan="1">Mean</th>
<th id="C6275D1D47E4E3A4181EC3A2DE9FB378" colspan="1" pageId="0" pageNumber="507" rowspan="1">SD</th>
<th id="EBE0D51741826910BB74CE0221FD5DA4" colspan="1" pageId="0" pageNumber="507" rowspan="1">Paratypes (Female) N = 105</th>
<th id="7B96AE70451EC1B662F13B909D339663" colspan="1" pageId="0" pageNumber="507" rowspan="1">Mean</th>
<th id="85EBED9CC4C75F62C6166FEDB180F833" colspan="1" pageId="0" pageNumber="507" rowspan="1">SD</th>
</tr>
<tr id="DB26187A9F36C395EE2A45D1C68089B0" pageId="0" pageNumber="507">
<td id="6E25E47C2F9C8FAE323C199F6FDCCF28" colspan="1" pageId="0" pageNumber="507" rowspan="1">Standard length (mm)</td>
<td id="60BCBE21066244E0D6619E5D599A692F" colspan="1" pageId="0" pageNumber="507" rowspan="1">35.9</td>
<td id="4E0C64979B6D6173617062157B31F4ED" colspan="1" pageId="0" pageNumber="507" rowspan="1">20.9-35.9</td>
<td id="AE93AA7403CBE93C378FE09C1EF20717" colspan="1" pageId="0" pageNumber="507" rowspan="1">26.6</td>
<td id="2106CACF55DA8F47AB4F0641C06586C6" colspan="1" pageId="0" pageNumber="507" rowspan="1">-</td>
<td id="FA43A842F045A70D822D0DECA1AF6197" colspan="1" pageId="0" pageNumber="507" rowspan="1">21.0-32.4</td>
<td id="BDDAB9874979BBD2DCBBB8A13D258ECC" colspan="1" pageId="0" pageNumber="507" rowspan="1">28.0</td>
<td id="07FE5DC7465B2CEA8DBEF68265939B67" colspan="1" pageId="0" pageNumber="507" rowspan="1">-</td>
</tr>
<tr id="00EAD0C7586D760A2F053BB1FCF2855B" pageId="0" pageNumber="507">
<td id="A8FBFD87CCABDAE4ADD5E5B6A6788061" colspan="8" pageId="0" pageNumber="507" rowspan="1">
<emphasis id="2C16FC0BC5CA05B59C0E484BBE4E5B0E" bold="true" pageId="0" pageNumber="507">Percentages of standard length</emphasis>
</td>
</tr>
<tr id="F2181D4D559CC8824E8C32AE2E5E9F20" pageId="0" pageNumber="507">
<td id="515D06AD0AD5992AB45100E16CEF343E" colspan="1" pageId="0" pageNumber="507" rowspan="1">Depth at dorsal-fin origin (body depth)</td>
<td id="9FB384451163908ADD71966F0F124096" colspan="1" pageId="0" pageNumber="507" rowspan="1">25.4</td>
<td id="DD2B67FBB517AD5D5BEFF2A4DE123F03" colspan="1" pageId="0" pageNumber="507" rowspan="1">24.5-28.7</td>
<td id="5BB30145AF5EDB8D03E368534C64C6BC" colspan="1" pageId="0" pageNumber="507" rowspan="1">25.9</td>
<td id="955D77E82A3BC92913A6E58C7BD74CB7" colspan="1" pageId="0" pageNumber="507" rowspan="1">1.0</td>
<td id="DAC69A31CBC3E23177E521E4C3F03B30" colspan="1" pageId="0" pageNumber="507" rowspan="1">25.6-29.1</td>
<td id="A1FB7D6A6ECEE4E3B2BEFC3DCCBD1804" colspan="1" pageId="0" pageNumber="507" rowspan="1">26.3</td>
<td id="05D3A58C66A3604BE866CB15DF8516E3" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.8</td>
</tr>
<tr id="74A247774CC7172EB6E9E4DD0429606D" pageId="0" pageNumber="507">
<td id="A4A49F246140B1ACC9FF2FE5DF128BCC" colspan="1" pageId="0" pageNumber="507" rowspan="1">Snout to dorsal-fin origin</td>
<td id="A3A641A676EB60118F1B1BC409024464" colspan="1" pageId="0" pageNumber="507" rowspan="1">53.1</td>
<td id="3EB8C8E83EBA11C5091E0064C72FD998" colspan="1" pageId="0" pageNumber="507" rowspan="1">51.9-55.6</td>
<td id="0DE7357885FEAB57865381109F6782DA" colspan="1" pageId="0" pageNumber="507" rowspan="1">52.6</td>
<td id="275BE8DEA50A3700E6AC3C1E413C7662" colspan="1" pageId="0" pageNumber="507" rowspan="1">1.1</td>
<td id="DC57293DECD2632D3356C227A2708D5E" colspan="1" pageId="0" pageNumber="507" rowspan="1">51.8-54.5</td>
<td id="E9E3B5435B5AC2E3842289E3FA101470" colspan="1" pageId="0" pageNumber="507" rowspan="1">52.1</td>
<td id="36370DA8401B2C099261580F7B0D93BA" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.7</td>
</tr>
<tr id="8B25BFC6C2A53FF7E41A5CFE4C39CDF0" pageId="0" pageNumber="507">
<td id="BD0A2AFC6F29A2CF2DD21BB740EEF20F" colspan="1" pageId="0" pageNumber="507" rowspan="1">Snout to pectoral-fin origin</td>
<td id="5523F0F105BAA676644508C742D7FFBF" colspan="1" pageId="0" pageNumber="507" rowspan="1">23.2</td>
<td id="25367EE6CA673642FABBB11B328CAA82" colspan="1" pageId="0" pageNumber="507" rowspan="1">23.0-27.7</td>
<td id="21EAE4AD7D83A0B2F5E7239A420A9C2F" colspan="1" pageId="0" pageNumber="507" rowspan="1">23.9</td>
<td id="805C7BF3F4B574B2CE764A4C7136D360" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.9</td>
<td id="5AA2B8F006AD085199BADA3AE1EBFED0" colspan="1" pageId="0" pageNumber="507" rowspan="1">22.6-25.2</td>
<td id="0E68DBB31C9666EF922083682E4422CE" colspan="1" pageId="0" pageNumber="507" rowspan="1">23.5</td>
<td id="1FD41F8F459E553AE44A148CC9288A44" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.6</td>
</tr>
<tr id="AA4E74761A30D18599125862B64C00FB" pageId="0" pageNumber="507">
<td id="4FE8C42E15BCF7578FC48CBF94259911" colspan="1" pageId="0" pageNumber="507" rowspan="1">Snout to pelvic-fin origin</td>
<td id="278043B74F1F7538BF4B75C3077DC9D5" colspan="1" pageId="0" pageNumber="507" rowspan="1">46.3</td>
<td id="60D05759A111D412603939D515BB8A76" colspan="1" pageId="0" pageNumber="507" rowspan="1">45.1-49.4</td>
<td id="35141E206523A7C034CB653EDFFC2740" colspan="1" pageId="0" pageNumber="507" rowspan="1">45.2</td>
<td id="41ECCFC4DD3B6092DF5AAE8F7E89F44A" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.6</td>
<td id="9BA5B6404897DDD61AE557337163ACD9" colspan="1" pageId="0" pageNumber="507" rowspan="1">44.2-47.1</td>
<td id="BB1FB2A56AB511CAB49A4DC137D542DA" colspan="1" pageId="0" pageNumber="507" rowspan="1">44.6</td>
<td id="EADEEAD5FBC5FF7358858A10264F1E31" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.9</td>
</tr>
<tr id="FE1A743D136D1B3D9ED675037F044FCB" pageId="0" pageNumber="507">
<td id="EBC28DC720099F654A68909926FE089B" colspan="1" pageId="0" pageNumber="507" rowspan="1">Snout to anal-fin origin</td>
<td id="5EDA417F65B8B5905466FECA0EA9B907" colspan="1" pageId="0" pageNumber="507" rowspan="1">67.4</td>
<td id="6CA114C151CE6F18344CF8148A0A996B" colspan="1" pageId="0" pageNumber="507" rowspan="1">63.9-68.6</td>
<td id="685CC3D9289365DDCE9FBCEB622200A1" colspan="1" pageId="0" pageNumber="507" rowspan="1">64.4</td>
<td id="3BC9E46EB45587B3980D0F565E2ED9CC" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.9</td>
<td id="C281B028A639FEC5C5D7CC7560CBBECB" colspan="1" pageId="0" pageNumber="507" rowspan="1">64.0-68.5</td>
<td id="65E1275E4A586168D7BFCFB7FA26B910" colspan="1" pageId="0" pageNumber="507" rowspan="1">64.2</td>
<td id="9E2A31CA0A0E545062CEE3FB7DC86EC6" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.7</td>
</tr>
<tr id="0D016187F8829217C7B428EA3FA28E81" pageId="0" pageNumber="507">
<td id="9C8382D74EDFDB896A94FD3A99629FDB" colspan="1" pageId="0" pageNumber="507" rowspan="1">Caudal peduncle depth</td>
<td id="6D7B31754EB4AC2A157B30B5BE49FBC0" colspan="1" pageId="0" pageNumber="507" rowspan="1">10.8</td>
<td id="668C427B7A74AEE069DEFB2630F29354" colspan="1" pageId="0" pageNumber="507" rowspan="1">10.1-12.5</td>
<td id="E552F0127521D6BDA8CFF794E3D558A0" colspan="1" pageId="0" pageNumber="507" rowspan="1">11.3</td>
<td id="69C75A792377B052A4D42D60AA6A9082" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.5</td>
<td id="1371201EDA44F8056381033AA6401188" colspan="1" pageId="0" pageNumber="507" rowspan="1">10.9-12.2</td>
<td id="6F2A84CF7EAFC253D050D203C86877B7" colspan="1" pageId="0" pageNumber="507" rowspan="1">11.3</td>
<td id="638ADE417E9F8AFDE54CAFFA5506A158" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.3</td>
</tr>
<tr id="0137C278708D507DD98FF6E593E85D58" pageId="0" pageNumber="507">
<td id="9BB1C2C5FB59795538A6FA0C7E7B0FD4" colspan="1" pageId="0" pageNumber="507" rowspan="1">Caudal peduncle length</td>
<td id="D65454636011A101415B5B2D487C35DF" colspan="1" pageId="0" pageNumber="507" rowspan="1">13.2</td>
<td id="40A9E5F8A0A692AA8D7F6FC516341D90" colspan="1" pageId="0" pageNumber="507" rowspan="1">12.2-17.2</td>
<td id="B44B0778C801A7E0CFA4325B1028DBF1" colspan="1" pageId="0" pageNumber="507" rowspan="1">14.0</td>
<td id="E5E2F829490A79FA66F74DA449868661" colspan="1" pageId="0" pageNumber="507" rowspan="1">1.2</td>
<td id="820BE4D2C37354E2A492E9A89BC1F970" colspan="1" pageId="0" pageNumber="507" rowspan="1">12.2-14.9</td>
<td id="BE2E9E51BE6F2265E1090549AF160357" colspan="1" pageId="0" pageNumber="507" rowspan="1">13.1</td>
<td id="7C91971912A99559EC0550CFE4F39C90" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.7</td>
</tr>
<tr id="022B845FCE0D342AF327B054BE82F530" pageId="0" pageNumber="507">
<td id="C40D15F7554B82A1B890C73EA6C881F8" colspan="1" pageId="0" pageNumber="507" rowspan="1">Pectoral-fin length</td>
<td id="015DD6206826F2F0D4C93020BD17E05B" colspan="1" pageId="0" pageNumber="507" rowspan="1">20.4</td>
<td id="B63868BF76E77665CAE04EA559CF5CDA" colspan="1" pageId="0" pageNumber="507" rowspan="1">17.9-22.5</td>
<td id="F2FF0FD668967E47E8F7FB698CA12376" colspan="1" pageId="0" pageNumber="507" rowspan="1">19.7</td>
<td id="CADD912C8ACCFDAF4AA0A3E8C0D5A61E" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.3</td>
<td id="DB8A576EE53A857DD5BEDAD2094C9156" colspan="1" pageId="0" pageNumber="507" rowspan="1">18.6-21.1</td>
<td id="E79AEF4D943FE3AFB5C8BF7E5331BB84" colspan="1" pageId="0" pageNumber="507" rowspan="1">19.3</td>
<td id="1BE7CF571F29C7AAACA1154EE39DD790" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.6</td>
</tr>
<tr id="5FCFAF8EF7F8CD558B685E2911A157D4" pageId="0" pageNumber="507">
<td id="7F63940273BEF90548F7CD732375DA85" colspan="1" pageId="0" pageNumber="507" rowspan="1">Pelvic-fin length</td>
<td id="15C713EB93FA1EA276E2731B92BFAD0A" colspan="1" pageId="0" pageNumber="507" rowspan="1">15.9</td>
<td id="DD4A5506EA7DE67D494290AAC1366FF4" colspan="1" pageId="0" pageNumber="507" rowspan="1">14.6-20.6</td>
<td id="7210A940BAC022C50096A476540CA98B" colspan="1" pageId="0" pageNumber="507" rowspan="1">15.6</td>
<td id="A33DBAE13CC1F0954879594901EF2114" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.5</td>
<td id="260CB9C68EB9AFFB090C333A9C85315D" colspan="1" pageId="0" pageNumber="507" rowspan="1">14.0-17.1</td>
<td id="DD3A0A67744E36863C44848883A7DD25" colspan="1" pageId="0" pageNumber="507" rowspan="1">15.3</td>
<td id="EEB18DAB7B1CC02E9384EF927DDA5462" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.7</td>
</tr>
<tr id="9059B6CD7DA50AA21D2D7D6826B4C596" pageId="0" pageNumber="507">
<td id="CC804AD4C710B75DA5081A7FC12FCA99" colspan="1" pageId="0" pageNumber="507" rowspan="1">Dorsal-fin base length</td>
<td id="7D3CD35E4B3D88FD3DC61D7241277CA0" colspan="1" pageId="0" pageNumber="507" rowspan="1">11.6</td>
<td id="65B184C1802450C93CE755F5CC713F5D" colspan="1" pageId="0" pageNumber="507" rowspan="1">9.5-13.4</td>
<td id="9F3295CC667F695BF5C5045CC9519F72" colspan="1" pageId="0" pageNumber="507" rowspan="1">11.3</td>
<td id="878ECD9539226BE0ABA8044ECD4634B3" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.5</td>
<td id="FD115A300BA6655E860B6798645A54D5" colspan="1" pageId="0" pageNumber="507" rowspan="1">10.8-13.0</td>
<td id="FC08D9405A2D31F2CD76B82CB68B8790" colspan="1" pageId="0" pageNumber="507" rowspan="1">11.8</td>
<td id="A0C4D9A52D6ACBFD5AAB1AC385E5195A" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.5</td>
</tr>
<tr id="287C03D3FF3C4DA6ECF595B5DD67DE2D" pageId="0" pageNumber="507">
<td id="CA848489FD7E1B390FDCD1F30DD43211" colspan="1" pageId="0" pageNumber="507" rowspan="1">Dorsal-fin height</td>
<td id="2AAE4BF1A1E02D355396C63B42F78957" colspan="1" pageId="0" pageNumber="507" rowspan="1">23.1</td>
<td id="B4D83166D6E7B91FDEE7DEF411156EB8" colspan="1" pageId="0" pageNumber="507" rowspan="1">21.2-24.8</td>
<td id="68F812680B69886ACC595F47D781A1C9" colspan="1" pageId="0" pageNumber="507" rowspan="1">22.4</td>
<td id="7E309179A727250557FCEA61BA8C2478" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.5</td>
<td id="CAAA2FED4C54A3629E045E00444F6A85" colspan="1" pageId="0" pageNumber="507" rowspan="1">20.8-24.0</td>
<td id="0D4E1BE433343A0D8E0B69A171098A23" colspan="1" pageId="0" pageNumber="507" rowspan="1">22.3</td>
<td id="0DA131FDB6140715C1CC980209094138" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.7</td>
</tr>
<tr id="054C607AD5505BF21987D2CDAF5ADEF2" pageId="0" pageNumber="507">
<td id="EE15CFDE142BA89F5B7BC292F9B4A598" colspan="1" pageId="0" pageNumber="507" rowspan="1">Anal-fin base length</td>
<td id="1E66F33D90A7A711971ECF753B6804D0" colspan="1" pageId="0" pageNumber="507" rowspan="1">18.9</td>
<td id="1592E8CCA19CB84EB7FCF1326A3920CF" colspan="1" pageId="0" pageNumber="507" rowspan="1">16.7-21.1</td>
<td id="7873617097CC7CBB8767EF4EA970771E" colspan="1" pageId="0" pageNumber="507" rowspan="1">18.1</td>
<td id="062C9C4177AA917757662CA6948815E7" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.4</td>
<td id="BC0BE15BA30604CBB4849BBB584CC705" colspan="1" pageId="0" pageNumber="507" rowspan="1">16.8-20.7</td>
<td id="76E74D6572F1840BC777034F0649C438" colspan="1" pageId="0" pageNumber="507" rowspan="1">18.3</td>
<td id="1F70ADAAB36D7D7982B64D4953D30644" colspan="1" pageId="0" pageNumber="507" rowspan="1">1.0</td>
</tr>
<tr id="D652A2A7CBD070F2538133F2232604DA" pageId="0" pageNumber="507">
<td id="A41C3D741FE47280D0E843FCCA45C989" colspan="1" pageId="0" pageNumber="507" rowspan="1">Eye to dorsal-fin origin</td>
<td id="E850D26DDF2B7227CED3D3E208A68C22" colspan="1" pageId="0" pageNumber="507" rowspan="1">42.6</td>
<td id="220AC9E708DE34734E2B439401D7AB48" colspan="1" pageId="0" pageNumber="507" rowspan="1">40.6-54.6</td>
<td id="B421CDF8E47807A0FE8A918B609DB817" colspan="1" pageId="0" pageNumber="507" rowspan="1">42.1</td>
<td id="F5B6D2908FFA296D692814A4FB497818" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.6</td>
<td id="37931B5F8E2A81F067DFF466EF3134F6" colspan="1" pageId="0" pageNumber="507" rowspan="1">41.4-52.4</td>
<td id="65C991557079CFF13E84CB992E966D9A" colspan="1" pageId="0" pageNumber="507" rowspan="1">41.8</td>
<td id="946F6FB6D16AA3B287E6BB4AE96D00B5" colspan="1" pageId="0" pageNumber="507" rowspan="1">1.9</td>
</tr>
<tr id="157F8F06D57F8D1B2B3B16A10EB634D6" pageId="0" pageNumber="507">
<td id="AD195A535228B2BAB9C4BA2A61749688" colspan="1" pageId="0" pageNumber="507" rowspan="1">Dorsal-fin origin to caudal-fin base</td>
<td id="18C2C9988B2B9F0FE7EE1375416F812B" colspan="1" pageId="0" pageNumber="507" rowspan="1">47.6</td>
<td id="655D923D307A7206504FD82D553B4B71" colspan="1" pageId="0" pageNumber="507" rowspan="1">46.5-49.5</td>
<td id="7EF01D02E785CCAD52226E08B8467CBC" colspan="1" pageId="0" pageNumber="507" rowspan="1">46.5</td>
<td id="FC79B6BB3459BD74A8340F332CE024C9" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.7</td>
<td id="6962DFC30841AB8D4874C91B17796B5D" colspan="1" pageId="0" pageNumber="507" rowspan="1">46.4-49.4</td>
<td id="3559940794A4C03D377C8DB8B591B1DC" colspan="1" pageId="0" pageNumber="507" rowspan="1">46.5</td>
<td id="029DE403425095353EC9B1B518102D0F" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.7</td>
</tr>
<tr id="42013D27D684FF35DF955C1FE606B178" pageId="0" pageNumber="507">
<td id="EB394E63E04CA4A16F649E7AD20488F4" colspan="1" pageId="0" pageNumber="507" rowspan="1">Head length</td>
<td id="A7318BEA821EAFCB06C9265BF9938C90" colspan="1" pageId="0" pageNumber="507" rowspan="1">24.0</td>
<td id="F181F3638CB35504CDFABBEE1B7969D0" colspan="1" pageId="0" pageNumber="507" rowspan="1">22.3-26.6</td>
<td id="8D3ED807A99A14D13E270B05AECDEED5" colspan="1" pageId="0" pageNumber="507" rowspan="1">24.0</td>
<td id="28C12C7C04F99ACD21B986976A0D750A" colspan="1" pageId="0" pageNumber="507" rowspan="1">1.7</td>
<td id="3318F6617F7BDA0878E690CEB6BD065F" colspan="1" pageId="0" pageNumber="507" rowspan="1">22.3-24.9</td>
<td id="B54955D2F2698FFDEB407404E337A52F" colspan="1" pageId="0" pageNumber="507" rowspan="1">23.1</td>
<td id="788BD16E7953A76C1A322359A1B23352" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.6</td>
</tr>
<tr id="E229658E05E50CACFB8A2BB888BDD100" pageId="0" pageNumber="507">
<td id="B4B12BC6BBC065465E7CB878D69B3088" colspan="8" pageId="0" pageNumber="507" rowspan="1">
<emphasis id="559A4DDEB36DC4E8E0D410F7AE4347FB" bold="true" pageId="0" pageNumber="507">Percentages of head length</emphasis>
</td>
</tr>
<tr id="4D965DD1CCC8EA08E52E7ECD61FFFE46" pageId="0" pageNumber="507">
<td id="79211A942A80C4C68E6E1E7E24E1ED71" colspan="1" pageId="0" pageNumber="507" rowspan="1">Horizontal eye diameter</td>
<td id="C6171F9BC97F0AE05FDCB6CFA98317E9" colspan="1" pageId="0" pageNumber="507" rowspan="1">30.2</td>
<td id="6A5FFB70685663C4E4885516C9DA7B17" colspan="1" pageId="0" pageNumber="507" rowspan="1">28.7 -36.0</td>
<td id="36ECAFCC9C0B481CABEB17335106BDA2" colspan="1" pageId="0" pageNumber="507" rowspan="1">31.4</td>
<td id="09A1AC0780E10863FCEB3E0E6562D423" colspan="1" pageId="0" pageNumber="507" rowspan="1">1.5</td>
<td id="67ECE8C7EEF18E4BDBA45DB9DE505B2E" colspan="1" pageId="0" pageNumber="507" rowspan="1">29.5-34.8</td>
<td id="FB12EA527BB1551461AE42A0067391E8" colspan="1" pageId="0" pageNumber="507" rowspan="1">31.6</td>
<td id="FE8314E4DAC1C522F3C0ED5FE858549A" colspan="1" pageId="0" pageNumber="507" rowspan="1">1.4</td>
</tr>
<tr id="AFB5ECE061A0975B92DBD92D48C232F8" pageId="0" pageNumber="507">
<td id="765D92DA8B0F252B626B9AF1DF1AC204" colspan="1" pageId="0" pageNumber="507" rowspan="1">Snout length</td>
<td id="CB0309AF585A923B8005C96931B995CA" colspan="1" pageId="0" pageNumber="507" rowspan="1">24.2</td>
<td id="D9449B4557C18D2C74B10909B903E361" colspan="1" pageId="0" pageNumber="507" rowspan="1">19.7 -28.8</td>
<td id="7A47FE9EE8B4FCE9DADB08669194A7E3" colspan="1" pageId="0" pageNumber="507" rowspan="1">23.5</td>
<td id="6B32B8500FB050624F6C346512F2554E" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.6</td>
<td id="92A78C95CCD84EC267A2ED7D15867283" colspan="1" pageId="0" pageNumber="507" rowspan="1">22.8-29.3</td>
<td id="BF96C08971A53672309A11AF8FCB4FD7" colspan="1" pageId="0" pageNumber="507" rowspan="1">25.4</td>
<td id="8AE85640E349B17EB0B1C12977F9EDD8" colspan="1" pageId="0" pageNumber="507" rowspan="1">1.2</td>
</tr>
<tr id="ECD365D1109DA625FE2F5D9C849D7853" pageId="0" pageNumber="507">
<td id="A5DF337923B432B3C1D7E6BB7BDE43B2" colspan="1" pageId="0" pageNumber="507" rowspan="1">Least interorbital width</td>
<td id="D2E6A977206D33E9675900793E610F37" colspan="1" pageId="0" pageNumber="507" rowspan="1">36.8</td>
<td id="E29A6A4EB29C8D91D0BA21F2A0156ABD" colspan="1" pageId="0" pageNumber="507" rowspan="1">32.7 -38.9</td>
<td id="AA53C674E7B00D0D1D34A6156E4982F3" colspan="1" pageId="0" pageNumber="507" rowspan="1">34.1</td>
<td id="87407EB570D625E9F724B97EB7E54341" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.1</td>
<td id="503262C8D413F8D0E7F96B1689A2B0D0" colspan="1" pageId="0" pageNumber="507" rowspan="1">32.9 -37.0</td>
<td id="5B9B89D7A3D60D73644F7764D58D9E5F" colspan="1" pageId="0" pageNumber="507" rowspan="1">11.1</td>
<td id="047E19016DEBE8279D9BC0AB2A4D5CD7" colspan="1" pageId="0" pageNumber="507" rowspan="1">1.1</td>
</tr>
<tr id="E8634A0D93DB2C5FAA18C83E33E0D8DA" pageId="0" pageNumber="507">
<td id="8F93AAE3A8274D493EDBE7A4BCCCA9B3" colspan="1" pageId="0" pageNumber="507" rowspan="1">Upper jaw length</td>
<td id="502F5C763CE2E61F0B337A700C01F260" colspan="1" pageId="0" pageNumber="507" rowspan="1">34.2</td>
<td id="03DE6E334993B407BFD0A3F508E099A4" colspan="1" pageId="0" pageNumber="507" rowspan="1">31.9 -37.3</td>
<td id="972951336F75D384EF44DE2F84284A57" colspan="1" pageId="0" pageNumber="507" rowspan="1">33.4</td>
<td id="90C7C4A76C6C53E040E0098E61BCE166" colspan="1" pageId="0" pageNumber="507" rowspan="1">0.2</td>
<td id="6E688B50FE4B6B0EE1FC477724836B2E" colspan="1" pageId="0" pageNumber="507" rowspan="1">32.7-39.9</td>
<td id="1BC7AFFA514C94DDBD3383AA8AD1962D" colspan="1" pageId="0" pageNumber="507" rowspan="1">33.9</td>
<td id="D8081D43D5BFB08FC21E32E47F268D34" colspan="1" pageId="0" pageNumber="507" rowspan="1">1.4</td>
</tr>
</table>
</paragraph>
<paragraph id="94BAD326026AC02A7393776F9E267F95" pageId="0" pageNumber="507">All teeth unicuspid or tricuspid and lateral cusps, when present, much smaller; premaxillary teeth in one rows with 6(9), 7(23) tricuspid teeth; maxilla with 11(3), 12(12), 13(14), or 14(3) unicuspid teeth; dentary with 6 (2) or 7 (30) larger tricuspid teeth followed by 6(26) or 7(6) smaller tricuspid teeth.</paragraph>
<paragraph id="38F2131A1A154507057D8E69F813B9F0" pageId="0" pageNumber="507">Scales cycloid and same size over entire body generally. Predorsal scales mostly regular, but sometimes irregular just posterior to supraoccipital and/or slightly anterior to dorsal-fin. Scales covering anterior third of caudal-fin, with up to two, three, or four scales beyond posterior margin of hypural plate. Lateral line interrupted; last scale on caudal-fin base, with 9+1(12),10+1(74), 11+1(50), or 12+1(5). Longitudinal scales series including lateral-line scales 35(3), 36(3), 37(56), 38(49), or 39(30). Longitudinal scales rows between dorsal-fin origin and lateral line 5(1), 6(93) or 7(47). Horizontal scale rows between lateral line and pelvic-fin origin 4 (141), Axillary scale present. Scales in median series between tip of supraoccipital spine and dorsal-fin origin 13+1(24),14+1(65), 15+1(26), or 16+1(26). Circumpeduncular scales 13(18), 14(115), or 15(8).</paragraph>
<paragraph id="3A6071BC842E55828EC52302814716A4" pageId="0" pageNumber="507">
Dorsal-fin rays i+10(99) or ii+10(42). Dorsal-fin origin situated posterior to vertical through pelvic-fin insertion, near middle of body. First dorsal-fin pterygiophore main body located of 8th and 9th vertebrae. Adipose-fin present. Anal-fin i+14(20), iii+15(18), ii+16(61), iii+16(24), ii+17(10), iii+17(5), ii+18 (3). Anteriormost anal-fin pterygiophore inserting at 14th and 15th vertebrae. Anterior anal-fin margin slightly convex, with anteriormost rays more elongate and slightly more thickened than remaining rays, forming a distinct lobe. Remaining rays smaller with straight distal margin. Pectoral-fin rays i+9(8), i+10(113), or i+11(20). Tip of pectoral-fin not reaching pelvic-fin origin, when adpressed. Pelvic-fin rays i+7(120) or ii+7(21). Tip of pelvic-fin not reaching anal-fin origin, when adpressed. Caudal-fin with a sexually dimorphic pattern, described below (
<figureCitation id="6BF9F0373733ACCC4B3EEF399FE00748" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Aphyocharax brevicaudatus sp. nov. a. CICCAA 02293, holotype (male), 35.9 mm SL; b. CICCAA 02294, paratype (female), 32.4 mm SL, Brazil: Maranhao state: Maracacume river basin. (Photographed by Erick Guimaraes)." figureDoi=".figure1" httpUri="https://binary.pensoft.net/fig/349754" pageId="0" pageNumber="507" tableDoi=".figure1">Fig. 1</figureCitation>
). Principal caudal-fin rays 10+9(130) or 10+10(11); dorsal procurrent rays 8(2), 9(3) or 10(27) and ventral procurrent rays 7(2), 8(3) or 9(27).
</paragraph>
<paragraph id="44133E6A9013BE4CD25499F3B5CE7AC5" pageId="0" pageNumber="507">Branchiostegal rays 4(32). Supraneurals 6(4) 7(27) or 8(1). Total vertebrae 31 (1), 32(30) or 33(1).</paragraph>
</subSubSection>
<subSubSection id="B4DEBF1A9F96F9F3132F7BE9271B2F7C" pageId="0" pageNumber="507" type="colour in alcohol">
<paragraph id="A5B1099A16E5578E50D000179275E5CE" pageId="0" pageNumber="507">Colour in alcohol.</paragraph>
<paragraph id="8B8F9BC4B50AF4F6E9BA7995CC239AD5" pageId="0" pageNumber="507">Ground colouration light brown to yellowish brown. Inconspicuous light brown to light gray stripe from humeral spot to caudal-fin base, more conspicuous on posterior half. Humeral region with one conspicuous dark brown to black humeral spot. Smaller dark brown or black chromatophores homogeneously scattered. Smaller dark brown or black chromatophores homogeneously scattered along body, except on chest. Head ground colouration similar to trunk, with dark brown chromatophores present on jaws, tip of snout, opercle, and dorsal portion of head. Dorsal, adipose, anal, caudal, pectoral, and pelvic fins hyaline to light brown.</paragraph>
</subSubSection>
<subSubSection id="E2BBA13A07510B94708163BD31C56806" pageId="0" pageNumber="507" type="sexual dimorphism">
<paragraph id="C4674D279C50C5AD269BDB6F2DE51046" pageId="0" pageNumber="507">Sexual dimorphism.</paragraph>
<paragraph id="25A772AC15C6F468C38FD84BA355F533" pageId="0" pageNumber="507">
Caudal-fin of mature males with upper lobe longer (about 2/3 longer) than lower one, while both cauldal-fin lobes have similar leght in females (
<figureCitation id="60D998F0B0A4A4E562C5EEDBB875A2D3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Aphyocharax brevicaudatus sp. nov. a. CICCAA 02293, holotype (male), 35.9 mm SL; b. CICCAA 02294, paratype (female), 32.4 mm SL, Brazil: Maranhao state: Maracacume river basin. (Photographed by Erick Guimaraes)." figureDoi=".figure1" httpUri="https://binary.pensoft.net/fig/349754" pageId="0" pageNumber="507" tableDoi=".figure1">Fig. 1</figureCitation>
). Gill glands were found in all analyzed mature males of
<taxonomicName id="AEC4BAADB1CC43D7D3C5F2683E15C954" family="Characidae" genus="Aphyocharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphyocharax brevicaudatus" order="Characiformes" pageId="0" pageNumber="507" phylum="Chordata" rank="species" species="brevicaudatus">
<emphasis id="1067BFD582ABC007789EB2442CA00BA5" italics="true" pageId="0" pageNumber="507">Aphyocharax brevicaudatus</emphasis>
</taxonomicName>
sp. nov. and were always absent in females. They were always located on anteriormost portion of lower branch of first gill arch, extending posteriorly through variable number of gill filaments.
</paragraph>
</subSubSection>
<subSubSection id="962407BAF6D848F50885C7B743B52ADD" pageId="0" pageNumber="507" type="etymology">
<paragraph id="54EA3AAFA0756AA69E97945D5D29862E" pageId="0" pageNumber="507">Etymology.</paragraph>
<paragraph id="23FFAA4F9F52989DBED935354F66A4CB" pageId="0" pageNumber="507">
The name
<taxonomicName id="C0E01867B95B148B6A8A5DE1AD521E8B" lsidName="brevicaudatus" pageId="0" pageNumber="507" rank="species" species="brevicaudatus">
<emphasis id="4F149630F50DCF124757CF0BAF0DDE93" italics="true" pageId="0" pageNumber="507">brevicaudatus</emphasis>
</taxonomicName>
is a contraction of the
<taxonomicName id="4725710DB797C89426DFC6246DCB2A77" genus="Latin" lsidName="Latin" pageId="0" pageNumber="507" rank="genus">
<emphasis id="6C1E9A4C51EAC1761797F6762C0119A4" italics="true" pageId="0" pageNumber="507">Latin</emphasis>
</taxonomicName>
words
<emphasis id="50EBAD29997167F683CC959B9AF59CA9" italics="true" pageId="0" pageNumber="507">brevis</emphasis>
meaning
<normalizedToken id="21F5ED1B757AF689608769DE336D4C49" originalValue="“short”">&quot;short&quot;</normalizedToken>
and
<emphasis id="CB73E1DD17BAA13C8DC4A3B2A19A3F66" italics="true" pageId="0" pageNumber="507">cauda</emphasis>
meaning
<normalizedToken id="5B224B3ED4C089E32BD26ACC300A8F3D" originalValue="“tail”">&quot;tail&quot;</normalizedToken>
, an allusion to the shorter caudal-fin lower lobe in the mature males of the new species.
</paragraph>
</subSubSection>
<subSubSection id="B3115941FAE04B031C74CEABE609FF46" pageId="0" pageNumber="507" type="distribution">
<paragraph id="C6AC7FA05E080D1C8EA5E9CDD6988007" pageId="0" pageNumber="507">Geographic distribution.</paragraph>
<paragraph id="BA94F5A806A42F3A5E3335A0C01B996E" pageId="0" pageNumber="507">
<taxonomicName id="5658E19C210AE54F63B2BDF8B5DFBB97" family="Characidae" genus="Aphyocharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphyocharax brevicaudatus" order="Characiformes" pageId="0" pageNumber="507" phylum="Chordata" rank="species" species="brevicaudatus">
<emphasis id="910871E111F68B72965223604FFC4F59" italics="true" pageId="0" pageNumber="507">Aphyocharax brevicaudatus</emphasis>
</taxonomicName>
sp. nov. is currently known only from a single locality, the
<normalizedToken id="13216ED74ED6878581EEC0AF2C90871D" originalValue="Maracaçumé">Maracacume</normalizedToken>
river basin, a small and isolated coastal river basin of the eastern Amazon region (
<figureCitation id="B1D4BAEA2695CE77AEE36B09732FEB63" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Type locality of Aphyocharax brevicaudatus sp. nov." figureDoi=".figure3" httpUri="https://binary.pensoft.net/fig/349756" pageId="0" pageNumber="507" tableDoi=".figure3">Fig. 3</figureCitation>
).
</paragraph>
<caption id="FA485D3C129933051084F0EEB5FBC4F8" doi=".figure3" httpUri="https://binary.pensoft.net/fig/349756" pageId="0" pageNumber="507" start="Figure 3" startId="F3">
<paragraph id="40ACB8ED1BCB1C0D74583B76F64217A4" pageId="0" pageNumber="507">
<emphasis id="4660275D148162B4432253CA1E56DA2A" bold="true" pageId="0" pageNumber="507">Figure 3.</emphasis>
Type locality of
<taxonomicName id="EA353186EAC711CC592FFE4404F9C0BE" family="Characidae" genus="Aphyocharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphyocharax brevicaudatus" order="Characiformes" pageId="0" pageNumber="507" phylum="Chordata" rank="species" species="brevicaudatus">
<emphasis id="DBE3BDA32FFAD577AAB3682B0DB92AD8" italics="true" pageId="0" pageNumber="507">Aphyocharax brevicaudatus</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

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<taxonomicName id="973C3245874ECAE4BE670441AECB0B3D" ID-CoL="73MYF" LSID-ZBK="urn:lsid:zoobank.org:act:E7DA85A9-DDF7-44BF-BD07-1F567EE506ED" family="Rivulidae" genus="Moema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Moema apurinan Costa 2004:3" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="species" species="apurinan">Moema apurinan</taxonomicName>
<taxonomicNameLabel id="1B32940E1DE9B929F257B25E989CD362" rank="species">new species</taxonomicNameLabel>
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<paragraph id="ADB085817B5B9B8121E377A3A908272C" pageNumber="3">(Figs. 1-2)</paragraph>
</subSubSection>
<subSubSection id="8359171B0EE620A295636F246240C7C6" type="nomenclature">
<paragraph id="21C70CFCD5B6F2EF9D94A94A29F94554" pageNumber="3">
<materialsCitation id="6FDDDAA0487EB48F574D787F0389DABE" ID-GBIF-Occurrence="923899155" country="Brazil" stateProvince="Estado do Acre">
<typeStatus id="2F2CEDA72277952BF1EC1370C6577E4F">Holotype</typeStatus>
.
<collectionCode id="6493980EA34CC46970C35563BD76D407" collectionName="UFRJ">UFRJ</collectionCode>
5980, male, 52.5 mm SL;
<collectingCountry id="263B3D54E557789C95B509C301539F2F">Brazil</collectingCountry>
:
<collectingRegion id="A2EA44670A59848B5B4919801C0205D1">Estado do Acre</collectingRegion>
: Porto Acre, temporary pool within forest, Rio Acre floodplains, Rio Purus drainage, Rio Amazonas basin (9o35’12.2”S 67o31’50.6”W; altitude 132 m); W. J. E. M. Costa, S. Lima and A. Pinto,
<date id="106B7CF7E43B46561AAE889358DBE754" day="10" month="06" value="2003-06-10" year="2003">10 June 2003</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="75FD4956C7A32DA5AB2F0900BCB4B356" type="nomenclature">
<paragraph id="724B23B77F82927ED2E59FE8081A8765" country="Brazil" pageNumber="3">
<materialsCitation id="C8881AABFBD08B6FA3B343918CE004DE" ID-GBIF-Occurrence="923899153" country="Brazil" stateProvince="Estado do Acre">
<typeStatus id="6EA634A60E91FC9AB083971E36C32E03">Paratypes</typeStatus>
.
<collectionCode id="0DC5BE2DF1BF3840CE0556CC376B83EC" collectionName="UFRJ">UFRJ</collectionCode>
5981, 4 males, 50.2-55.4 mm SL, 5 females, 42.7-51.1 mm SL
</materialsCitation>
;
<materialsCitation id="93B2818F41415AFC7BF75BD2BE337D13" ID-GBIF-Occurrence="923899152" country="Brazil" stateProvince="Estado do Acre">
<collectionCode id="8EF8EE44A43AF5CACABF7A90CB7C9AF9" collectionName="UFRJ">UFRJ</collectionCode>
5982, 4 males, 48.2-54.4 mm SL, 3 females, 46.8-47.9 mm SL (c&amp;s); collected with
<typeStatus id="6C7ED22FB48058E7AAF255BD883042D4">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="97C1359F2867F41775AAA6590AA93B9E" type="diagnosis">
<paragraph id="FE95C133650247AED278BBFF401DAEED" pageNumber="3">
Diagnosis: The new species differs from all other species of the genus in having fewer caudal-fin rays (30-31, vs. 32-35). It is similar to
<taxonomicName id="827B5EF82649DB2BC67CBE002F83601E" family="Rivulidae" genus="Moema" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="species" species="staecki">M. staecki (Seegers)</taxonomicName>
, and distinguished from all other congeners by possessing contact organs on scales of the flank of the male (vs. contact organs absent), caudal fin acuminate (vs. rounded to subtruncate), fewer pectoral-fin rays (14, vs. 15-17), fewer vertebrae (33-35, vs. 36-39), and narrower rostral cartilage (width about 75-80% of rostral cartilage length, vs. 90-100% in the remaining species of the genus, except in
<taxonomicName id="F7F3DFB1A6426C8840C3BFBDF009E0D2" LSID-ZBK="urn:lsid:zoobank.org:act:07C0B958-A31B-47C3-9BF1-4D91025A23FA" family="Rivulidae" genus="Moema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Moema ortegai Costa 2003:7" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="species" species="ortegai">M. ortegai Costa</taxonomicName>
, about 145%). The new species differs from
<taxonomicName id="8F888F42FC0D1F872E54F328B97B1D3A" family="Rivulidae" genus="Moema" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="species" species="staecki">M. staecki</taxonomicName>
by having a longer pectoral fin (tip reaching between the base of 8th and 13th anal fin rays, vs. reaching urogenital papilla), and wider basihyal (width about 75% of basihyal length, vs. 55 %).
</paragraph>
</subSubSection>
<subSubSection id="CC0A45B0FE1F8EBBE3BB8DADD57E55AE" type="description">
<paragraph id="B0B2F33567AE74E74913F6A8866EE718" pageNumber="4">Description: Morphometric data given in Table 1. Male larger than female, largest male 55.4 mm SL. Dorsal profile gently convex from snout to end of dorsal-fin base, approximately straight to slightly concave on caudal peduncle. Ventral profile slightly convex on head, almost straight from anterior portion of venter to end of anal-fin base, nearly straight on caudal peduncle. Body slender, subcylindrical anteriorly, deeper than wide, to compressed posteriorly. Greatest body depth at level of pelvic-fin base. Jaws long, snout pointed.</paragraph>
<paragraph id="0C4494637653ABB44D7ACC7BB9053F2A" pageNumber="4">Tip of dorsal fin pointed, with short filamentous ray in male, rounded to slightly pointed in female. Anal fin approximately rectangular, longer than deep; tip pointed, long, reaching vertical through middle of caudal fin in male, rounded and short in female. Caudal fin long, acuminate, with short posterior filamentous rays in male, short and elliptical in female. Pectoral fin lanceolate, terminating in long tip, reaching between base of 8th and 13th anal fin rays in male, slightly pointed and short, reaching between pelvic-fin base and anus in female. Pelvic fin rounded, tip reaching between base of 1st and 3rd anal-fin rays in male, and between anus and urogenital papilla in female. Pelvic-fin bases in close proximity. Dorsal-fin origin on vertical through base of 9th or 10th anal-fin ray, and between neural spines of 20th and 22nd vertebrae. Anal-fin origin between pleural ribs of 16th and 17th vertebrae. Dorsal-fin rays 10-11; anal-fin rays 16-17; caudal-fin rays 30- 31; pectoral-fin rays 14; pelvic-fin rays 7.</paragraph>
<paragraph id="061EBB5E74E67CB366F98D2E95A39977" lastPageNumber="5" pageNumber="4">Scales large, cycloid. Body and head entirely scaled, except anterior ventral surface of head. Body scales extending over caudal-fin base; no scales on dorsal and anal fins. Frontal squamation E-patterned; E-scales not overlapping medially; scales arranged in irregular circular pattern around A-scale without exposed margins. Longitudinal series of scales 34- 35; transverse series of scales 8; scale rows around caudal peduncle 16. Ctenii-like contact organ on each scale of median ventral portion of male flank. Supraorbital neuromasts 3 + 3.</paragraph>
<paragraph id="523D1B2DE87F81BB23EBEFA746576253" pageNumber="5">Interhyal vestigial, not ossified. Rostral cartilage longer than wide, width 75-80% length. Basihyal subtriangular, width 75% length; basihyal cartilage 45% basihyal length. Six branchiostegal rays. Second pharyngobranchial with no, one or two small teeth. Gillrakers of first branchial arch 1 + 10. Vomerine teeth 4-11. Ventral process of posttemporal present. Total vertebrae 33-35.</paragraph>
<paragraph id="B136C21754BC919712C93A7A86965C8C" lastPageNumber="6" pageNumber="5">Coloration: Male: Side of body greenish golden, with three purplish brown stripes alternating with narrow dark golden stripes; dark red crimson dots simultaneously arranged both in longitudinal rows, on two dorsalmost purplish brown stripes, and in oblique rows crossing flank. Dorsum light brown with brown dots, venter white. Side of head greenish golden with dark brown blotches. Jaws light brown. Infraorbital region and ventral surface of head white. Iris yellowish green, with dark brown bar crossing center of eye. Dorsal fin pale greenish yellow with small dark reddish brown spots. Anal fin light greenish yellow, basal region light blue, with small dark reddish brown spots. Caudal fin pale greenish yellow with small dark reddish brown spots on dorsal portion and short brown stripes or dots parallel to fin rays on median portion; broad light orange stripe with bright yellow dorsal contour, dorsally outlined by dark brown stripe, on ventral margin of fin. Pelvic fin light greenish yellow, with transverse rows of small brown spots. Pectoral fin yellowish hyaline.</paragraph>
<paragraph id="A125F01CCC31760985762542D1A7E6FA" pageNumber="6">Female: Side of body light metallic green, with six longitudinal rows of brown dots. Dorsum light brown with brown dots, venter white. Side of head pale metallic green with faint brown spots. Jaws gray. Infraorbital region and ventral surface of head white. Iris yellow, with pale brown bar crossing center of eye. Dorsal and anal fins pale greenish yellow with small dark brown spots. Caudal fin pale greenish yellow with small dark brown spots on dorsal portion, and faint small gray spots on middle. Paired fins hyaline.</paragraph>
</subSubSection>
<subSubSection id="648FB67FC20C6821F4F8119C44C4FEDC" type="distribution">
<paragraph id="C433CD61D1F38FE50AA7A8A789EDDC0D" pageNumber="6">Distribution: Known only from the type locality, Rio Acre floodplains, Rio Purus drainage, Rio Amazonas basin, northwestern Brazil (Fig. 3).</paragraph>
</subSubSection>
<subSubSection id="30D84C77730F045793D32A2960724410" type="biology_ecology">
<paragraph id="2A0733CBCB5308BEE4DAFA36982FF0EC" pageNumber="7">Habitat notes: The type series was collected in a temporary pool within the forest. The pool was small, about 7 m of long and about 3 m wide, shallow, about 0.40-0.60 m deep. The water was dark brown. No aquatic vegetation.</paragraph>
</subSubSection>
<subSubSection id="0EC0F19A17EA845470AF3A79947AA8D9" type="etymology">
<paragraph id="BD1B52415FECF14199E37C0CABCBE3C9" pageNumber="7">
Etymology: The specific name apurinan (
<normalizedToken id="A5E3873A66323FE4020DD6DFF9111BC0" originalValue="Apurinã">Apurina</normalizedToken>
) is the Portuguese name of an indigenous tribe of people, also known as
<normalizedToken id="3927323B0F716B7ADCBAF3BF73C22500" originalValue="Ipurinã">Ipurina</normalizedToken>
,
<normalizedToken id="D0B3A156B17BBD160AC4D5E363962B08" originalValue="Popingaré">Popingare</normalizedToken>
and
<normalizedToken id="7D8FA93E00B9F9E80324711885022F6E" originalValue="Kangitê">Kangite</normalizedToken>
, with own language derived from the linguistic group Aruak, inhabiting lands drained by the Rio Purus basin in the region of the type locality of the new species.
</paragraph>
</subSubSection>
</treatment>
</document>