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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.133.37773" ID-GBIF-Dataset="a6caa96b-fb3a-4877-9e16-b2f519188af6" ID-PMC="PMC6791900" ID-Pensoft-Pub="1314-2003-133-1" ID-Pensoft-UUID="99F75C9803D65C4880141BE7B6589FB5" ID-PubMed="31632179" ModsDocID="1314-2003-133-1" checkinTime="1570497209971" checkinUser="pensoft" docAuthor="Shalimov, Aleksandr Petrovich, Wu, Yu-Dong &amp; Zhang, Xian-Chun" docDate="2019" docId="000E4F3F4E555417938A3EA17449CDDB" docLanguage="en" docName="PhytoKeys 133: 1-76" docOrigin="PhytoKeys 133" docSource="http://dx.doi.org/10.3897/phytokeys.133.37773" docTitle="Selaginella chrysorrhizos Spring 1850" docType="treatment" docVersion="6" id="99F75C9803D65C4880141BE7B6589FB5" lastPageNumber="1" masterDocId="99F75C9803D65C4880141BE7B6589FB5" masterDocTitle="A taxonomic revision of the genus Selaginella (Selaginellaceae) from Nepal" masterLastPageNumber="76" masterPageNumber="1" pageNumber="1" updateTime="1668139527481" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A taxonomic revision of the genus Selaginella (Selaginellaceae) from Nepal</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Shalimov, Aleksandr Petrovich</mods:namePart>
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<mods:namePart>Wu, Yu-Dong</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Zhang, Xian-Chun</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>PhytoKeys</mods:title>
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<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>133</mods:number>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.133.37773</mods:identifier>
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<treatment ID-GBIF-Taxon="159654801" LSID="urn:lsid:plazi:treatment:000E4F3F4E555417938A3EA17449CDDB" httpUri="http://treatment.plazi.org/id/000E4F3F4E555417938A3EA17449CDDB" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<taxonomicName LSID="000E4F3F-4E55-5417-938A-3EA17449CDDB" authority="Spring" authorityName="Spring" authorityYear="1850" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Selaginella chrysorrhizos" order="Selaginellales" pageId="0" pageNumber="1" phylum="Pteridophyta" rank="species" species="chrysorrhizos">Selaginella chrysorrhizos Spring</taxonomicName>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Morphological diversity of the leaves of Nepalese Sealginella species 1 A-C S. repanda (Nakaike 3708, PE) 2 A-C S. vaginata (Nakaike 1102, PE) 3 A-C S. chrysorrhizos (Nakaike 3708, PE). A - Axillary leaves, B - Dorsal leaves, C - Ventral leaves. Scale bars: 0.2 mm." figureDoi="10.3897/phytokeys.133.37773.figure7" httpUri="https://binary.pensoft.net/fig/344903" pageId="0" pageNumber="1" tableDoi="10.3897/phytokeys.133.37773.figure7">Figs 7</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diversity strobili of Nepalese Selaginella species A S. chrysorrhizos (Nakaike 3708, PE) B S. reticulata (Nakaike 1760, PE) C S. subdiaphana (Zhang 5, PE) D S. tenuifolia (PE-Xizang Exped. PE 6280, PE). Scale bars: 1 mm (A, D), 2 mm (B, C)." figureDoi="10.3897/phytokeys.133.37773.figure11" httpUri="https://binary.pensoft.net/fig/344907" pageId="0" pageNumber="1" tableDoi="10.3897/phytokeys.133.37773.figure11">
(3
<normalizedToken originalValue="A–C">A-C</normalizedToken>
), 11A
</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Selaginella reticulata (Hook. &amp; Grev.) Spring A (A 1) Habit, lower surface B strobilus, lower surface C fragment of the lower surface of the lateral branches D fragment of the upper surface of the lateral branches (T. Nakaike 1760, PE)." figureDoi="10.3897/phytokeys.133.37773.figure32" httpUri="https://binary.pensoft.net/fig/344928" pageId="0" pageNumber="1" tableDoi="10.3897/phytokeys.133.37773.figure32">, 32</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Spring" authorityYear="1850" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Selaginella chrysorrhizos" order="Selaginellales" pageId="0" pageNumber="1" phylum="Pteridophyta" rank="species" species="chrysorrhizos">Selaginella chrysorrhizos</taxonomicName>
Spring, Monogr. Lycop. 2: 251, no. 189. 1850, p. p.;
<bibRefCitation author="Iwatsuki, K" editor="Ohashi, H" journalOrPublisher="Vol. 8. The University Museum, the University of Tokyo, Tokyo" pageId="0" pageNumber="1" pagination="166 - 205" refId="B12" refString="Iwatsuki, K, 1975. Pteridophyta. In: Ohashi, H, Ed., Flora of Eastern Himalaya. Vol. 8. The University Museum, the University of Tokyo, Tokyo: 166 - 205" title="Pteridophyta." volumeTitle="Flora of Eastern Himalaya." year="1975">Iwatsuki 1975</bibRefCitation>
;
<bibRefCitation author="Dixit, RD" journalOrPublisher="Dehra Dun, Bishen Singh Mahendra Pal Singh" pageId="0" pageNumber="1" refId="B4" refString="Dixit, RD, 1992. Selaginellaceae of India. Dehra Dun, Bishen Singh Mahendra Pal Singh" title="Selaginellaceae of India." year="1992">Dixit 1992</bibRefCitation>
;
<bibRefCitation author="Thapa, N" journalOrPublisher="Department Plant Resources" pageId="0" pageNumber="1" pagination="1 - 175" refId="B27" refString="Thapa, N, 2002. Pteridophytes of Nepal. Bull. Department Plant Resources 19: 1 - 175" title="Pteridophytes of Nepal. Bull." volume="19" year="2002">Thapa 2002</bibRefCitation>
;
<bibRefCitation author="Fraser-Jenkins, CR" journalOrPublisher="National Herbarium and Plant Laboratories, Department of Plant Resources, Ministry of Forests and Soil Conservation, Kathmandu" pageId="0" pageNumber="1" refId="B7" refString="Fraser-Jenkins, CR, Kandel, DR, Pariyar, S, 2015. Ferns and fern-allies of Nepal. Vol. 1. National Herbarium and Plant Laboratories, Department of Plant Resources, Ministry of Forests and Soil Conservation, Kathmandu" title="Ferns and fern-allies of Nepal. Vol. 1." year="2015">Fraser-Jenkins et al. 2015</bibRefCitation>
;
<bibRefCitation author="Fraser-Jenkins, CR" journalOrPublisher="National Herbarium and Plant Laboratories, Department of Plant Resources, Ministry of Forests and Soil Conservation, Kathmandu" pageId="0" pageNumber="1" refId="B8" refString="Fraser-Jenkins, CR, Gandhi, KN, Kholia, BS, Benniamin, A, 2017. An annotated checklist of Indian Pteridophytes, Part - I (Lycopodiaceae to Thelypteridace). Bishen Singh Mahendra Pal Singh Dehra Dun, 562 pp." title="An annotated checklist of Indian Pteridophytes, Part - I (Lycopodiaceae to Thelypteridace). Bishen Singh Mahendra Pal Singh Dehra Dun, 562 pp." year="2017">Fraser-Jenkins et al. 2017</bibRefCitation>
.
<emphasis bold="true" pageId="0" pageNumber="1">Type.</emphasis>
(lectotype, designated by
<bibRefCitation author="Fraser-Jenkins, CR" journalOrPublisher="National Herbarium and Plant Laboratories, Department of Plant Resources, Ministry of Forests and Soil Conservation, Kathmandu" pageId="0" pageNumber="1" refId="B8" refString="Fraser-Jenkins, CR, Gandhi, KN, Kholia, BS, Benniamin, A, 2017. An annotated checklist of Indian Pteridophytes, Part - I (Lycopodiaceae to Thelypteridace). Bishen Singh Mahendra Pal Singh Dehra Dun, 562 pp." title="An annotated checklist of Indian Pteridophytes, Part - I (Lycopodiaceae to Thelypteridace). Bishen Singh Mahendra Pal Singh Dehra Dun, 562 pp." year="2017">Fraser-Jenkins et al. 2017</bibRefCitation>
) Assam,
<emphasis italics="true" pageId="0" pageNumber="1">Griffith 141</emphasis>
(K).
</paragraph>
<paragraph pageId="0" pageNumber="1">
=
<taxonomicName authorityName="R.D.Dixit" authorityYear="1985" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Selaginella panchghaniana" order="Selaginellales" pageId="0" pageNumber="1" phylum="Pteridophyta" rank="species" species="panchghaniana">Selaginella panchghaniana</taxonomicName>
R.D. Dixit, Bull. Bot. Surv. India 25(1-4): 226, t. 1, f. 3. 1985;
<bibRefCitation author="Dixit, RD" journalOrPublisher="Dehra Dun, Bishen Singh Mahendra Pal Singh" pageId="0" pageNumber="1" refId="B4" refString="Dixit, RD, 1992. Selaginellaceae of India. Dehra Dun, Bishen Singh Mahendra Pal Singh" title="Selaginellaceae of India." year="1992">Dixit 1992</bibRefCitation>
.
<emphasis bold="true" pageId="0" pageNumber="1">Type.</emphasis>
INDIA. Maharashtra-Mahabaleshwar: Panchghani, 7 Nov 1966,
<emphasis italics="true" pageId="0" pageNumber="1">Panigrahi 11739</emphasis>
(holotype: BSA).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
Stems 8-12 cm, evergreen or seasonally green, suberect. Rhizophores restricted to base of stem. Main stems branched from near base or from lower part upward, in basal part main stem 0.5-1.1 mm in diam. Stems glabrous, glossy yellow, sulcate, primary leafy branches 5-8 pairs, forked or once or twice pinnately branched. Axillary leaves ovate-oblong, 1.0-2.0
<normalizedToken originalValue="×">x</normalizedToken>
0.5-0.8 in base slightly cuneate, margin denticulate, apex obtuse. Ventral leaves ovate-oblong, 0.6-0.8
<normalizedToken originalValue="×">x</normalizedToken>
1-2 mm, ascending, acroscopic base ovate-oblong, slightly dilated, imbricate, distantly rotundate at the base, margin denticulate, basiscopic base, entire, except apices, apex obtuse. Dorsal leaves ovate, 1.8-2
<normalizedToken originalValue="×">x</normalizedToken>
0.5-0.8 mm, subfalcate, margin denticulate, apex shortly cuspidate. Strobili solitary, terminal, compact, 3-7
<normalizedToken originalValue="×">x</normalizedToken>
1.5-2 mm. Sporophylls dimorphic, ventral sporophylls ovate, aristate, margin ciliate; dorsal sporophylls oblong, obtuse, margin ciliate-dentate. Megaspores dark-brown, surface verrucate; microspores pale-brown, surface verrucate.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="ecology">
<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="1">On banks and on large stones. Alt. 200-2000 m.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution in Nepal.</paragraph>
<paragraph pageId="0" pageNumber="1">W, C, E.</paragraph>
<paragraph pageId="0" pageNumber="1">Nepalese threatened status: not available data.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">General distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">BANGLADESH, BHUTAN, INDIA (Assam, Kerala, Madhya Pradesh, Manipur, Meghalaya, Mizoram, Sikkim, West Bengal), LAOS, MYANMAR, THAILAND, VIETNAM.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="chromosome number">
<paragraph pageId="0" pageNumber="1">Chromosome number.</paragraph>
<paragraph pageId="0" pageNumber="1">Not available data.</paragraph>
<paragraph pageId="0" pageNumber="1">Selected specimens examined:</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">C Nepal</emphasis>
: KASKI: &quot;Bhoot Bridge, between Butwal and Pokhara, alt. c. 300 m, 9 Nov 1988,
<emphasis italics="true" pageId="0" pageNumber="1">T. Nakaike 3723</emphasis>
&quot; (PE).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">MAKAWANPUR</emphasis>
: &quot;Pisulin [Fishling], near Muglin, Gorkha [Makawanpur District], alt. c. 300 m, 12 Nov 1988,
<emphasis italics="true" pageId="0" pageNumber="1">T. Nakaike 3831</emphasis>
&quot; (PE).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">E Nepal: SOLUKHUMBHU</emphasis>
: &quot;between Junbesi and Rachowa, Solukumbu, alt. 2600-3400 m, 12 Dec 1988,
<emphasis italics="true" pageId="0" pageNumber="1">T. Nakaike 3304</emphasis>
&quot; (PE).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">SANKHUWASABHA</emphasis>
: Simbu,
<geoCoordinate degrees="27" direction="north" minutes="22" orientation="latitude" precision="925" value="27.366667">27°22'N</geoCoordinate>
,
<geoCoordinate degrees="87" direction="east" minutes="47" orientation="longitude" precision="925" value="87.78333">87°47'E</geoCoordinate>
, alt. 1800 m, 5 Dec 1971,
<emphasis italics="true" pageId="0" pageNumber="1">J.F. Dobremez</emphasis>
DBR NEP
<emphasis italics="true" pageId="0" pageNumber="1">1335</emphasis>
&quot; (E00754784); &quot;Sunaturi, W. of Hetauda, Makawanpur, alt. c. 200 m, 8 Nov 1988,
<emphasis italics="true" pageId="0" pageNumber="1">T. Nakaike 3702</emphasis>
&quot; (PE); &quot; l.c.
<emphasis italics="true" pageId="0" pageNumber="1">3698</emphasis>
&quot; (PE); &quot;l.c.
<emphasis italics="true" pageId="0" pageNumber="1">3720</emphasis>
&quot; (PE); &quot;l.c.
<emphasis italics="true" pageId="0" pageNumber="1">3706</emphasis>
&quot; (PE); &quot;Simbu,
<geoCoordinate degrees="27" direction="north" minutes="22" orientation="latitude" precision="925" value="27.366667">27°22'N</geoCoordinate>
,
<geoCoordinate degrees="87" direction="east" minutes="47" orientation="longitude" precision="925" value="87.78333">87°47'E</geoCoordinate>
, alt. 2000 m, 5 Dec 1971,
<emphasis italics="true" pageId="0" pageNumber="1">J.F. Dobremez</emphasis>
DBR NEP
<emphasis italics="true" pageId="0" pageNumber="1">1333</emphasis>
&quot; (E00670601, E00754800); &quot;Simbu,
<geoCoordinate degrees="27" direction="north" minutes="22" orientation="latitude" precision="925" value="27.366667">27°22'N</geoCoordinate>
,
<geoCoordinate degrees="87" direction="east" minutes="47" orientation="longitude" precision="925" value="87.78333">87°47'E</geoCoordinate>
, alt. 1800 m, 5 Oct 1971,
<emphasis italics="true" pageId="0" pageNumber="1">J.F. Dobremez</emphasis>
DBR NEP
<emphasis italics="true" pageId="0" pageNumber="1">1335</emphasis>
&quot; (E00670602).
</paragraph>
<caption ID-Zenodo-Dep="3489998" doi="10.3897/phytokeys.133.37773.figure7" httpUri="https://binary.pensoft.net/fig/344903" pageId="0" pageNumber="1" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 7.</emphasis>
Morphological diversity of the leaves of Nepalese
<taxonomicName genus="Sealginella" lsidName="Sealginella" pageId="0" pageNumber="1" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Sealginella</emphasis>
</taxonomicName>
species
<emphasis bold="true" pageId="0" pageNumber="1">
1
<normalizedToken originalValue="A–C">A-C</normalizedToken>
</emphasis>
<taxonomicName family="Selaginellaceae" kingdom="Plantae" lsidName="S. repanda" order="Selaginellales" pageId="0" pageNumber="1" rank="species" species="repanda">
<emphasis italics="true" pageId="0" pageNumber="1">S. repanda</emphasis>
</taxonomicName>
(
<emphasis italics="true" pageId="0" pageNumber="1">Nakaike 3708</emphasis>
, PE)
<emphasis bold="true" pageId="0" pageNumber="1">
2
<normalizedToken originalValue="A–C">A-C</normalizedToken>
</emphasis>
<taxonomicName family="Selaginellaceae" kingdom="Plantae" lsidName="S. vaginata" order="Selaginellales" pageId="0" pageNumber="1" rank="species" species="vaginata">
<emphasis italics="true" pageId="0" pageNumber="1">S. vaginata</emphasis>
</taxonomicName>
(
<emphasis italics="true" pageId="0" pageNumber="1">Nakaike 1102</emphasis>
, PE)
<emphasis bold="true" pageId="0" pageNumber="1">
3
<normalizedToken originalValue="A–C">A-C</normalizedToken>
</emphasis>
<taxonomicName family="Selaginellaceae" kingdom="Plantae" lsidName="S. chrysorrhizos" order="Selaginellales" pageId="0" pageNumber="1" rank="species" species="chrysorrhizos">
<emphasis italics="true" pageId="0" pageNumber="1">S. chrysorrhizos</emphasis>
</taxonomicName>
(
<emphasis italics="true" pageId="0" pageNumber="1">Nakaike 3708</emphasis>
, PE). A - Axillary leaves, B - Dorsal leaves, C - Ventral leaves. Scale bars: 0.2 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.950.51298" ID-GBIF-Dataset="21975627-96b7-45c7-9c80-0a73a850169a" ID-PMC="PMC7387366" ID-Pensoft-Pub="1313-2970-950-51" ID-Pensoft-UUID="7F08092C01585A70A7948EA57A2A8E2E" ID-PubMed="32774101" ID-ZooBank="3E805885D3354FB3AE8DFA443FAD82AE" ModsDocID="1313-2970-950-51" checkinTime="1595285263451" checkinUser="pensoft" docAuthor="Takaoka, Hiroyuki, Srisuka, Wichai, Fukuda, Masako &amp; Saeung, Atiporn" docDate="2020" docId="000E73CAE2B75781B3B3D18E00195374" docLanguage="en" docName="ZooKeys 950: 51-152" docOrigin="ZooKeys 950" docSource="http://dx.doi.org/10.3897/zookeys.950.51298" docTitle="Simulium (Gomphostilbia) maelanoiense Takaoka, Srisuka &amp; Saeung 2020, sp. nov." docType="treatment" docUuid="5776AAC8-0AE0-45E1-8873-B482DD48F13E" docUuidSource="ZooBank" docVersion="4" id="7F08092C01585A70A7948EA57A2A8E2E" lastPageNumber="51" masterDocId="7F08092C01585A70A7948EA57A2A8E2E" masterDocTitle="Twenty-one new species of the Simulium (Gomphostilbia) asakoae species group (Diptera, Simuliidae) in Thailand, with their genetic relationships" masterLastPageNumber="152" masterPageNumber="51" pageNumber="51" updateTime="1668169112888" updateUser="ExternalLinkService">
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<mods:title>Twenty-one new species of the Simulium (Gomphostilbia) asakoae species group (Diptera, Simuliidae) in Thailand, with their genetic relationships</mods:title>
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<mods:namePart>Takaoka, Hiroyuki</mods:namePart>
<mods:affiliation>Tropical Infectious Diseases Research and Education Centre (TIDREC), University of Malaya, 50603, Kuala Lumpur, Malaysia</mods:affiliation>
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<mods:namePart>Srisuka, Wichai</mods:namePart>
<mods:affiliation>Entomology Section, Queen Sirikit Botanic Garden, P. O. Box 7, Maerim, Chiang Mai 50180, Thailand</mods:affiliation>
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<mods:namePart>Fukuda, Masako</mods:namePart>
<mods:affiliation>Institute for Research Promotion, Oita University, Idaigaoka 1 - 1, Hasama, Yufu City, Oita, 879 - 5593, Japan</mods:affiliation>
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<mods:namePart>Saeung, Atiporn</mods:namePart>
<mods:affiliation>Center of Insect Vector Study, Department of Parasitology, Faculty of Medicine, Chiang Mai University, Chiang Mai 50200, Thailand</mods:affiliation>
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<mods:identifier type="Pensoft-UUID">7F08092C01585A70A7948EA57A2A8E2E</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="165843381" LSID="urn:lsid:zoobank.org:act:5776AAC8-0AE0-45E1-8873-B482DD48F13E" httpUri="http://treatment.plazi.org/id/000E73CAE2B75781B3B3D18E00195374" lastPageNumber="51" pageId="0" pageNumber="51">
<subSubSection pageId="0" pageNumber="51" type="nomenclature">
<paragraph pageId="0" pageNumber="51">
<taxonomicName LSID="000E73CA-E2B7-5781-B3B3-D18E00195374" authority="Takaoka, Srisuka &amp; Saeung" authorityName="Takaoka, Srisuka &amp; Saeung" authorityYear="2020" class="Insecta" family="Simuliidae" genus="Simulium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Simulium (Gomphostilbia) maelanoiense" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="maelanoiense" status="sp. nov." subGenus="Gomphostilbia">Simulium (Gomphostilbia) maelanoiense Takaoka, Srisuka &amp; Saeung</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="51">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female, male and pupa of S. maelanoiense sp. nov. A female B-F male G-I pupa. A mandible (left side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (right side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view). Scale bars: 1.0 mm (I); 0.1 mm (B, G), 0.02 mm (C-F); 0.01 mm (A, H)." figureDoi="10.3897/zookeys.950.51298.figure9" httpUri="https://binary.pensoft.net/fig/433873" pageId="0" pageNumber="51">Fig. 9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="materials_examined">
<paragraph pageId="0" pageNumber="51">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Holotype</emphasis>
: Male (with its associated pupal exuviae and cocoon) (in 80% ethanol) labeled as &quot;Holotype:
<taxonomicName authorityName="Takaoka, Srisuka &amp; Saeung" authorityYear="2020" class="Insecta" family="Simuliidae" genus="Simulium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Simulium maelanoiense" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="maelanoiense">
<emphasis italics="true" pageId="0" pageNumber="51">Simulium maelanoiense</emphasis>
</taxonomicName>
male, QSBG col. no. 105, Thailand, 14-VII-2017, by W. Srisuka&quot;, collected from a small stream (width 25 cm, depth 2.5 cm, bed sandy, moderate flow, pH 6.8, 21.9 °C, partially shaded, elevation 974 m,
<geoCoordinate degrees="18" direction="north" minutes="20" orientation="latitude" precision="1" seconds="35.8" value="18.343279">18°20'35.8&quot;N</geoCoordinate>
,
<geoCoordinate degrees="98" direction="east" minutes="01" orientation="longitude" precision="1" seconds="23.8" value="98.023285">98°01'23.8&quot;E</geoCoordinate>
), at Tee La Chan Waterfall, Mae La Noi District, Mae Hong Son Province, Thailand, 14-VII-2017, by W. Srisuka (Coll. No. 105).
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Paratypes</emphasis>
: Two females (thoraces for DNA analysis) (with its associated pupal exuviae and cocoon) (in 80% ethanol), collected from a stream (width 110 cm, depth 12 cm, bed sandy, moderate flow, pH 6.9, 18.3 °C, partially shaded, elevation 1,446 m,
<geoCoordinate degrees="18" direction="north" minutes="51" orientation="latitude" precision="1" seconds="38.8" value="18.860779">18°51'38.8&quot;N</geoCoordinate>
,
<geoCoordinate degrees="99" direction="east" minutes="22" orientation="longitude" precision="1" seconds="15.2" value="99.37089">99°22'15.2&quot;E</geoCoordinate>
), at Kiew Fin, Muang Pan District, Lampang Province, Thailand, 6-IV-2018, by W. Srisuka (Coll. No. 36); two males (thorax of one male for DNA analysis) (with their associated pupal exuviae and cocoons) (in 80% ethanol), same data as for holotype.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="diagnosis">
<paragraph pageId="0" pageNumber="51">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="51">
Female: mandible with one tooth on the outer margin (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female, male and pupa of S. maelanoiense sp. nov. A female B-F male G-I pupa. A mandible (left side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (right side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view). Scale bars: 1.0 mm (I); 0.1 mm (B, G), 0.02 mm (C-F); 0.01 mm (A, H)." figureDoi="10.3897/zookeys.950.51298.figure9" httpUri="https://binary.pensoft.net/fig/433873" pageId="0" pageNumber="51">9A</figureCitation>
). Male: small number of upper-eye facets in eleven vertical columns and 13 horizontal rows, and hind basitarsus (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female, male and pupa of S. maelanoiense sp. nov. A female B-F male G-I pupa. A mandible (left side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (right side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view). Scale bars: 1.0 mm (I); 0.1 mm (B, G), 0.02 mm (C-F); 0.01 mm (A, H)." figureDoi="10.3897/zookeys.950.51298.figure9" httpUri="https://binary.pensoft.net/fig/433873" pageId="0" pageNumber="51">9B</figureCitation>
) 1.0-1.1 times as wide as the hind femur.
</paragraph>
<caption doi="10.3897/zookeys.950.51298.figure9" httpUri="https://binary.pensoft.net/fig/433873" pageId="0" pageNumber="51" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" pageId="0" pageNumber="51">Figure 9.</emphasis>
Female, male and pupa of
<taxonomicName lsidName="S. maelanoiense" pageId="0" pageNumber="51" rank="species" species="maelanoiense">
<emphasis italics="true" pageId="0" pageNumber="51">S. maelanoiense</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="51">A</emphasis>
female
<emphasis bold="true" pageId="0" pageNumber="51">B-F</emphasis>
male
<emphasis bold="true" pageId="0" pageNumber="51">G-I</emphasis>
pupa.
<emphasis bold="true" pageId="0" pageNumber="51">A</emphasis>
mandible (left side)
<emphasis bold="true" pageId="0" pageNumber="51">B</emphasis>
hind basitarsus and second tarsomere (left side; lateral view)
<emphasis bold="true" pageId="0" pageNumber="51">C</emphasis>
coxites, styles and ventral plate (ventral view)
<emphasis bold="true" pageId="0" pageNumber="51">D</emphasis>
style (right side; ventrolateral view)
<emphasis bold="true" pageId="0" pageNumber="51">E</emphasis>
ventral plate and median sclerite (lateral view)
<emphasis bold="true" pageId="0" pageNumber="51">F</emphasis>
ventral plate (caudal view)
<emphasis bold="true" pageId="0" pageNumber="51">G</emphasis>
gill filaments (right side; lateral view)
<emphasis bold="true" pageId="0" pageNumber="51">H</emphasis>
terminal hooks (caudal view)
<emphasis bold="true" pageId="0" pageNumber="51">I</emphasis>
cocoon (dorsal view). Scale bars: 1.0 mm (
<emphasis bold="true" pageId="0" pageNumber="51">I</emphasis>
); 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="51">B, G</emphasis>
), 0.02 mm (
<emphasis bold="true" pageId="0" pageNumber="51">C-F</emphasis>
); 0.01 mm (
<emphasis bold="true" pageId="0" pageNumber="51">A, H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="description">
<paragraph pageId="0" pageNumber="51">Description.</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" pageId="0" pageNumber="51">Female</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="51">N</emphasis>
= 2). Body length 2.0 mm.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Head.</emphasis>
Frontal ratio 1.9:1.0:2.7; frons:head ratio 1.0:4.9. Labrum 0.70 times length of clypeus. Maxillary palpus: proportional lengths of third, fourth, and fifth palpal segments 1.0:1.1:2.1; sensory vesicle 0.26-0.32 times length of third palpal segment. Lacinia with ten or eleven inner and 12 or 13 outer teeth. Mandible (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female, male and pupa of S. maelanoiense sp. nov. A female B-F male G-I pupa. A mandible (left side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (right side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view). Scale bars: 1.0 mm (I); 0.1 mm (B, G), 0.02 mm (C-F); 0.01 mm (A, H)." figureDoi="10.3897/zookeys.950.51298.figure9" httpUri="https://binary.pensoft.net/fig/433873" pageId="0" pageNumber="51">9A</figureCitation>
) with 20 inner teeth and one outer tooth at some distance from tip.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Legs.</emphasis>
Foreleg: basitarsus moderately dilated, 6.6 times as long as its greatest width. Midleg: tarsus dark brown to brownish black though basal half of basitarsus dark yellow (its border not well defined). Hind leg: coxa light brown; tibia yellowish white on basal three-fifths and light brown to brownish black on rest; tarsus brownish black except basal three-fifths (though base light brown) and basal half of second tarsomere yellowish white; basitarsus 6.9 times as long as wide, and 0.7 and 0.5 times as wide as greatest widths of tibia and femur, respectively; calcipala 0.6 times as wide as greatest width of basitarsus; claw with large basal tooth 0.44 times length of claw.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Wing.</emphasis>
Length 2.0 mm.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Abdomen.</emphasis>
Dorsal surface of abdomen medium to dark brown except anterior four-fifths of segment 2 ochreous.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Terminalia.</emphasis>
Sternite 8 with 27 medium-long to long hairs together with two to four slender short hairs on each side. Ovipositor valves each moderately covered with microsetae interspersed with one to three short hairs. Paraproct in ventral view with four sensilla on anteromedial surface; paraproct in lateral view 0.6 times as long as wide, with 19 or 20 medium-long to long hairs on ventral and lateral surfaces.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" pageId="0" pageNumber="51">Male</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="51">N</emphasis>
= 3). Body length 2.3-2.4 mm.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Head.</emphasis>
Somewhat wider than thorax. Upper eye medium brown, consisting of large facets in eleven (rarely ten) vertical columns and 13 horizontal rows on each side. Antenna light to medium brown except scape, pedicel, and base of first flagellomere whitish yellow; first flagellomere elongate, 1.9 times length of second. Maxillary palpus: proportional lengths of third, fourth, and fifth palpal segments 1.0:1.2:2.6; sensory vesicle 0.16-0.21 times length of third palpal segment.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Legs.</emphasis>
Foreleg: tibia yellowish white except apical three-tenths dark brown; basitarsus moderately dilated, 6.6-7.8 times as long as its greatest width. Hind leg: coxa light brown; tibia dark brown to brownish black except little less than basal half yellowish white; tarsus (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female, male and pupa of S. maelanoiense sp. nov. A female B-F male G-I pupa. A mandible (left side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (right side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view). Scale bars: 1.0 mm (I); 0.1 mm (B, G), 0.02 mm (C-F); 0.01 mm (A, H)." figureDoi="10.3897/zookeys.950.51298.figure9" httpUri="https://binary.pensoft.net/fig/433873" pageId="0" pageNumber="51">9B</figureCitation>
) brownish black except little more than basal two-fifths of basitarsus and basal one-third of second tarsomere yellowish white; basitarsus (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female, male and pupa of S. maelanoiense sp. nov. A female B-F male G-I pupa. A mandible (left side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (right side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view). Scale bars: 1.0 mm (I); 0.1 mm (B, G), 0.02 mm (C-F); 0.01 mm (A, H)." figureDoi="10.3897/zookeys.950.51298.figure9" httpUri="https://binary.pensoft.net/fig/433873" pageId="0" pageNumber="51">9B</figureCitation>
) 3.6-3.8 times as long as wide, and 1.0 and 1.0-1.1 times as wide as greatest width of tibia and femur, respectively; calcipala (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female, male and pupa of S. maelanoiense sp. nov. A female B-F male G-I pupa. A mandible (left side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (right side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view). Scale bars: 1.0 mm (I); 0.1 mm (B, G), 0.02 mm (C-F); 0.01 mm (A, H)." figureDoi="10.3897/zookeys.950.51298.figure9" httpUri="https://binary.pensoft.net/fig/433873" pageId="0" pageNumber="51">9B</figureCitation>
) slightly shorter than basal width, and 0.28 times as wide as greatest width of basitarsus.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Wing.</emphasis>
Length 2.0-2.1 mm. Subcosta with 2-13 hairs, though rarely without hair.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Genitalia.</emphasis>
Style in ventrolateral view (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female, male and pupa of S. maelanoiense sp. nov. A female B-F male G-I pupa. A mandible (left side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (right side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view). Scale bars: 1.0 mm (I); 0.1 mm (B, G), 0.02 mm (C-F); 0.01 mm (A, H)." figureDoi="10.3897/zookeys.950.51298.figure9" httpUri="https://binary.pensoft.net/fig/433873" pageId="0" pageNumber="51">9D</figureCitation>
) slightly tapered toward apex, with round apex. Ventral plate in ventral view (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female, male and pupa of S. maelanoiense sp. nov. A female B-F male G-I pupa. A mandible (left side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (right side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view). Scale bars: 1.0 mm (I); 0.1 mm (B, G), 0.02 mm (C-F); 0.01 mm (A, H)." figureDoi="10.3897/zookeys.950.51298.figure9" httpUri="https://binary.pensoft.net/fig/433873" pageId="0" pageNumber="51">9C</figureCitation>
) with basal arms of moderate length, nearly parallel-sided, then convergent apically; ventral plate in caudal view (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female, male and pupa of S. maelanoiense sp. nov. A female B-F male G-I pupa. A mandible (left side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (right side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view). Scale bars: 1.0 mm (I); 0.1 mm (B, G), 0.02 mm (C-F); 0.01 mm (A, H)." figureDoi="10.3897/zookeys.950.51298.figure9" httpUri="https://binary.pensoft.net/fig/433873" pageId="0" pageNumber="51">9F</figureCitation>
) with ventral margin nearly straight. Cercus with 13-15 hairs.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" pageId="0" pageNumber="51">Pupa</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="51">N</emphasis>
= 5). Body length 2.5-3.0 mm.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Head.</emphasis>
Integument yellow.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Thorax.</emphasis>
Integument yellow, moderately covered with round tubercles except dorsal surface of posterior half sparsely covered with tubercles, and dorsolateral surface of posterior half almost bare. Gill (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female, male and pupa of S. maelanoiense sp. nov. A female B-F male G-I pupa. A mandible (left side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (right side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view). Scale bars: 1.0 mm (I); 0.1 mm (B, G), 0.02 mm (C-F); 0.01 mm (A, H)." figureDoi="10.3897/zookeys.950.51298.figure9" httpUri="https://binary.pensoft.net/fig/433873" pageId="0" pageNumber="51">9G</figureCitation>
) composed of eight slender thread-like filaments, arranged as (3+3)+2 (or rarely [3+(1+2)]+2) from dorsal to ventral; common basal stalk 0.6-0.7 times length of interspiracular trunk; dorsal triplet composed of three individual filaments arising at same level, and with extremely short stalk in some pupae, middle triplet mostly composed of three individual filaments arising at same level, or rarely composed of one individual and two paired filaments with extremely short secondary stalk; stalk of ventral pair of filaments variable in length, 0.6-1.3 times length of common basal stalk, and 0.5-1.0 times length of interspiracular trunk; primary stalk of dorsal triplet lying against that of lower pair at angle of 60-80° when viewed laterally; filaments of dorsal triplet subequal in length (1.5-2.0 mm) and thickness to one another; filaments of middle triplet subequal in length (1.7-2.5 mm) and thickness, and two filaments of ventral pair subequal in length (2.5-3.6 mm) and thickness to each other and 1.4-1.8 times as thick as six other filaments of dorsal and middle triplets when compared basally.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Abdomen.</emphasis>
Dorsally, segments 1 and 2 without minute tubercles; segment 9 with pair of wide flat terminal hooks (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female, male and pupa of S. maelanoiense sp. nov. A female B-F male G-I pupa. A mandible (left side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (right side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view). Scale bars: 1.0 mm (I); 0.1 mm (B, G), 0.02 mm (C-F); 0.01 mm (A, H)." figureDoi="10.3897/zookeys.950.51298.figure9" httpUri="https://binary.pensoft.net/fig/433873" pageId="0" pageNumber="51">9H</figureCitation>
), of which outer margin 2.1-2.3 times length of inner margin and crenulated when viewed caudally.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Cocoon</emphasis>
(Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female, male and pupa of S. maelanoiense sp. nov. A female B-F male G-I pupa. A mandible (left side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (right side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view). Scale bars: 1.0 mm (I); 0.1 mm (B, G), 0.02 mm (C-F); 0.01 mm (A, H)." figureDoi="10.3897/zookeys.950.51298.figure9" httpUri="https://binary.pensoft.net/fig/433873" pageId="0" pageNumber="51">9I</figureCitation>
). Dark brown, slipper-shaped, moderately woven, moderately extended ventrolaterally; anterior margin thickly woven medially, often with bulge; 3.0-3.8 mm long by 2.0-2.5 mm wide.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" pageId="0" pageNumber="51">Mature larva.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="etymology">
<paragraph pageId="0" pageNumber="51">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="51">
The species name,
<taxonomicName lsidName="maelanoiense" pageId="0" pageNumber="51" rank="species" species="maelanoiense">
<emphasis italics="true" pageId="0" pageNumber="51">maelanoiense</emphasis>
</taxonomicName>
, refers to the district, Mae La Noi, where this species was collected.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="distribution">
<paragraph pageId="0" pageNumber="51">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="51">Thailand (Lampang and Mae Hong Son).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="discussion">
<paragraph pageId="0" pageNumber="51">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="51">
<taxonomicName authorityName="Takaoka, Srisuka &amp; Saeung" authorityYear="2020" class="Insecta" family="Simuliidae" genus="Simulium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Simulium maelanoiense" order="Diptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="maelanoiense">
<emphasis italics="true" pageId="0" pageNumber="51">Simulium maelanoiense</emphasis>
</taxonomicName>
sp. nov. is most similar in the male and pupa to
<taxonomicName lsidName="S. sutheppuiense" pageId="0" pageNumber="51" rank="species" species="sutheppuiense">
<emphasis italics="true" pageId="0" pageNumber="51">S. sutheppuiense</emphasis>
</taxonomicName>
sp. nov. in many characters including the number of male upper-eye facets, relative width of the hind basitarsus compared to the hind tibia and femur, shape of the ventral plate when viewed caudally, and arrangement of the pupal gill. However, this new species is barely distinguished in the female from
<taxonomicName lsidName="S. sutheppuiense" pageId="0" pageNumber="51" rank="species" species="sutheppuiense">
<emphasis italics="true" pageId="0" pageNumber="51">S. sutheppuiense</emphasis>
</taxonomicName>
sp. nov. by the number of the outer teeth of the mandible (one tooth in this new species versus three teeth in
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<emphasis italics="true" pageId="0" pageNumber="51">S. sutheppuiense</emphasis>
</taxonomicName>
sp. nov.) and the length ratio of the labrum against the clypeus (0.7 in this new species versus 0.6 in
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sp. nov.).
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64
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Vashkevich, 1924
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<figureCitation id="109C5F81FF804A06FD69FE96BD43FE89" box="[703,778,338,364]" captionStart="Fig" captionStartId="2.[189,232,1966,1992]" captionTargetBox="[199,1398,1100,1897]" captionTargetId="figure@2.[194,1482,1044,1958]" captionTargetPageId="2" captionText="Fig. 2. Map of the Udzungwa Mts showing collection sites for gomphodesmids. Based on Marshall et al. (2010: fig. 1)." figureDoi="http://doi.org/10.5281/zenodo.3928330" httpUri="https://zenodo.org/record/3928330/files/figure.png" pageId="10" pageNumber="11">Figs 2</figureCitation>
,
<figureCitation id="109C5F81FF804A06FCC0FE96BD75FE88" box="[790,828,338,365]" captionStart="Fig" captionStartId="6.[189,232,1500,1526]" captionTargetBox="[217,1367,909,1445]" captionTargetId="figure@6.[207,1377,905,1453]" captionTargetPageId="6" captionText="Fig. 5. Sternal process on body ring 6. A. Emphysemastix spp. B. Agrophogonus spp. except A. hamulifer Olsen &amp; Enghoff sp. nov. C. Agrophogonus hamulifer Olsen &amp; Enghoff sp. nov. D. Pogoropsis prolixopes Rosenmejer &amp; Enghoff gen. et sp. nov. E. Pogoro siren Rosenmejer &amp; Enghoff sp. nov. F. Pogoro scharffi Hoffman, 2005. G. Pogoro alopias Rosenmejer &amp; Enghoff sp. nov. Not drawn to scale." figureDoi="http://doi.org/10.5281/zenodo.3928336" httpUri="https://zenodo.org/record/3928336/files/figure.png" pageId="10" pageNumber="11">5G</figureCitation>
,
<figureCitation id="109C5F81FF804A06FC9FFE96BD3DFE89" box="[841,884,338,364]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="10.[189,232,1999,2025]" captionStartId-1="11.[189,232,1932,1958]" captionTargetBox-0="[254,1332,1428,1949]" captionTargetBox-1="[369,1206,265,1908]" captionTargetId-0="figure@10.[249,1337,1413,1958]" captionTargetId-1="figure@11.[369,1216,265,1908]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="Fig. 8. Pogoro alopias Rosenmejer &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621670). Scale bar: 5 mm." captionText-1="Fig. 9. Pogoro alopias Rosenmejer &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621670), right gonopod. A–B. Mesal view. C–D. Lateral view. E–F. Dorsal view. Abbreviations: L = Process L; M = Process M; tl = triangular lobe. Scale bars: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3928342" figureDoi-1="http://doi.org/10.5281/zenodo.3928344" httpUri-0="https://zenodo.org/record/3928342/files/figure.png" httpUri-1="https://zenodo.org/record/3928344/files/figure.png" pageId="10" pageNumber="11">8–9</figureCitation>
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<paragraph id="88184304FF804A06FF6BFE06BAD0FE1A" blockId="10.[189,1398,450,512]" pageId="10" pageNumber="11">Differs from other species of the genus by the shape of the postnodal telopodite, which resembles a thresher shark’s tail, and by the triangular shape of the sternal process on segment 6.</paragraph>
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The name is a noun in apposition, referring to the thresher shark,
<emphasis id="BAD39F16FF804A06FC15FD91BACEFD8A" box="[963,1159,597,623]" italics="true" pageId="10" pageNumber="11">Alopias vulpinus</emphasis>
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<paragraph id="88184304FF804A06FF6BFD7EBFE3FD31" blockId="10.[189,426,698,724]" box="[189,426,698,724]" pageId="10" pageNumber="11">
<heading id="D350F468FF804A06FF6BFD7EBFE3FD31" bold="true" box="[189,426,698,724]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="BAD39F16FF804A06FF6BFD7EBFE3FD31" bold="true" box="[189,426,698,724]" pageId="10" pageNumber="11">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF804A06FF3CFD2DBF15FCE6" blockId="10.[189,1398,745,876]" box="[234,348,745,771]" pageId="10" pageNumber="11">
<emphasis id="BAD39F16FF804A06FF3CFD2DBF15FCE6" bold="true" box="[234,348,745,771]" pageId="10" pageNumber="11">
<typeStatus id="571CFDA6FF804A06FF3CFD2DBF15FCE6" box="[234,348,745,771]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="88184304FF804A06FF6BFCC8BC4FFC89" blockId="10.[189,1398,745,876]" pageId="10" pageNumber="11">
<materialsCitation id="38CF4959FF804A06FF6BFCC8BC4FFC89" ID-GBIF-Occurrence="2803530316" collectingDate="1997-05-17" collectingDateMax="1997-05-27" collectingDateMin="1997-05-17" collectionCode="ZMUC" collectorName="SI Exp." country="Tanzania" elevation="1800" latitude="-8.36825" location="Kihanga Stream" longLatPrecision="1" longitude="35.97822" municipality="Udzungwa Scarp Forest Reserve" pageId="10" pageNumber="11" specimenCode="NHMD 621670" specimenCount="1" stateProvince="Iringa District" typeStatus="holotype">
<collectingCountry id="F0B00394FF804A06FF6BFCC8BF18FCC3" box="[189,337,780,806]" name="Tanzania" pageId="10" pageNumber="11">TANZANIA</collectingCountry>
•
<specimenCount id="9EA1888DFF804A06FEA7FCC8BFC0FCC3" box="[369,393,780,806]" pageId="10" pageNumber="11" type="male">♂</specimenCount>
;
<collectingRegion id="4A638DE6FF804A06FE4DFCCFBC0EFCC0" box="[411,583,779,805]" country="Tanzania" name="Iringa" pageId="10" pageNumber="11">Iringa District</collectingRegion>
,
<collectingMunicipality id="687CD97EFF804A06FD81FCCFBDACFCC3" box="[599,997,779,806]" pageId="10" pageNumber="11">Udzungwa Scarp Forest Reserve</collectingMunicipality>
,
<locationDeviation id="99AA5A26FF804A06FC23FCC8BB3AFCC0" box="[1013,1395,779,806]" location="Masisiwe Village" pageId="10" pageNumber="11">
<quantity id="4F5FEEE1FF804A06FC23FCC8BA0BFCC0" box="[1013,1090,779,806]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="11" unit="km" value="11.0">11 km</quantity>
SE of Masisiwe Village
</locationDeviation>
,
<location id="8D7815DFFF804A06FF6BFCEBBFCCFCAC" LSID="urn:lsid:plazi:treatment:000EF212FF804A00FDD4FEC8BDB6FDB9:8D7815DFFF804A06FF6BFCEBBFCCFCAC" box="[189,389,815,841]" country="Tanzania" latitude="-8.36825" longLatPrecision="1" longitude="35.97822" municipality="Udzungwa Scarp Forest Reserve" name="Kihanga Stream" pageId="10" pageNumber="11" stateProvince="Iringa District">Kihanga Stream</location>
;
<geoCoordinate id="ED9325C3FF804A06FE4EFCE9BC0DFCAC" box="[408,580,813,841]" degrees="08" direction="south" minutes="22" orientation="latitude" pageId="10" pageNumber="11" precision="1" seconds="05.7" value="-8.36825">08°22′05.7″ S</geoCoordinate>
,
<geoCoordinate id="ED9325C3FF804A06FD8EFCE9BD4EFCAC" box="[600,775,813,841]" degrees="35" direction="east" minutes="58" orientation="longitude" pageId="10" pageNumber="11" precision="1" seconds="41.6" value="35.97822">35°58′41.6″ E</geoCoordinate>
,
<elevation id="038AA437FF804A06FCCCFCEBBD89FCAC" box="[794,960,815,841]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="10" pageNumber="11" unit="m" value="1800.0">
<quantity id="4F5FEEE1FF804A06FCCCFCEBBD33FCAC" box="[794,890,815,841]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="10" pageNumber="11" unit="m" value="1800.0">1800 m</quantity>
a.s.l.
</elevation>
;
<date id="FC1965C4FF804A06FC03FCEBBAE7FCAC" box="[981,1198,815,841]" pageId="10" pageNumber="11" value="1997-05-17" valueMax="1997-05-27" valueMin="1997-05-17">
<collectingDate id="EC5D9C2CFF804A06FC03FCEBBAE7FCAC" box="[981,1198,815,841]" pageId="10" pageNumber="11" value="1997-05-17" valueMax="1997-05-27" valueMin="1997-05-17">17–27 May 1997</collectingDate>
</date>
;
<collectionCode id="EEB6DBC1FF804A06FB17FCEBBB5FFCAC" box="[1217,1302,815,841]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="10" pageNumber="11">ZMUC</collectionCode>
and SI Exp. 1997;
<specimenCode id="D801EB7FFF804A06FE93FC96BC4BFC89" box="[325,514,850,876]" collectionCode="NHMD" pageId="10" pageNumber="11">NHMD 621670</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="88184304FF804A06FF3CFC57BFEFFC48" blockId="10.[189,1052,915,977]" box="[234,422,915,941]" pageId="10" pageNumber="11">
<emphasis id="BAD39F16FF804A06FF3CFC57BFEFFC48" bold="true" box="[234,422,915,941]" pageId="10" pageNumber="11">Other material</emphasis>
</paragraph>
<paragraph id="88184304FF804A06FF6BFC73BA55FC35" blockId="10.[189,1052,915,977]" box="[189,1052,950,977]" pageId="10" pageNumber="11">
<materialsCitation id="38CF4959FF804A06FF6BFC73BA51FC35" ID-GBIF-Occurrence="2803530310" box="[189,1048,950,977]" collectingDate="1997-05-17" collectingDateMax="1997-05-27" collectingDateMin="1997-05-17" collectionCode="NHMD" collectorName="SI Exp." country="Tanzania" elevation="1800" latitude="-8.36825" location="TANZANIA" longLatPrecision="1" longitude="35.97822" municipality="Udzungwa Scarp Forest Reserve" pageId="10" pageNumber="11" specimenCode="NHMD 621671" specimenCount="1" specimenCount-female="1" stateProvince="Iringa">
<collectingCountry id="F0B00394FF804A06FF6BFC73BF18FC34" box="[189,337,951,977]" name="Tanzania" pageId="10" pageNumber="11">TANZANIA</collectingCountry>
•
<specimenCount id="9EA1888DFF804A06FEBFFC73BFDFFC34" box="[361,406,951,977]" pageId="10" pageNumber="11" type="female">1 ♀</specimenCount>
;
<geologicalTimeScale id="FA87C3AFFF804A06FE73FC73BD06FC35" box="[421,847,950,977]" pageId="10" pageNumber="11">same collecting data as for holotype</geologicalTimeScale>
;
<specimenCode id="D801EB7FFF804A06FC8DFC73BA51FC35" box="[859,1048,950,977]" collectionCode="NHMD" pageId="10" pageNumber="11">NHMD 621671</specimenCode>
</materialsCitation>
.
</paragraph>
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<subSubSection id="C0BD108FFF804A00FF6BFC3CBCCAFE15" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11" type="description">
<paragraph id="88184304FF804A06FF6BFC3CBF07FBF7" blockId="10.[189,334,1016,1042]" box="[189,334,1016,1042]" pageId="10" pageNumber="11">
<heading id="D350F468FF804A06FF6BFC3CBF07FBF7" bold="true" box="[189,334,1016,1042]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="BAD39F16FF804A06FF6BFC3CBF07FBF7" bold="true" box="[189,334,1016,1042]" pageId="10" pageNumber="11">Description</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF804A06FF3CFBE3BF63FBA4" blockId="10.[189,1398,1063,1159]" box="[234,298,1063,1089]" pageId="10" pageNumber="11">
<emphasis id="BAD39F16FF804A06FF3CFBE3BF63FBA4" bold="true" box="[234,298,1063,1089]" pageId="10" pageNumber="11">Male</emphasis>
</paragraph>
<paragraph id="88184304FF804A06FF6BFB8EBC7BFB62" blockId="10.[189,1398,1063,1159]" pageId="10" pageNumber="11">
SIZE AND SHAPE. Body length ca
<quantity id="4F5FEEE1FF804A06FDE8FB8DBCDAFB81" box="[574,659,1097,1124]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" pageId="10" pageNumber="11" unit="mm" value="53.0">53 mm</quantity>
. Maximum width
<quantity id="4F5FEEE1FF804A06FCBCFB8DBD8EFB81" box="[874,967,1097,1124]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="10" pageNumber="11" unit="mm" value="6.9">6.9 mm</quantity>
. W/L ratio 13%. Height of body ring eight
<quantity id="4F5FEEE1FF804A06FF29FBA9BF15FB62" box="[255,348,1133,1159]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="10" pageNumber="11" unit="mm" value="4.9">4.9 mm</quantity>
. H/W ratio= 72%.
</paragraph>
<paragraph id="88184304FF804A06FF6BFB6FBF59FB0D" blockId="10.[189,1398,1194,1256]" pageId="10" pageNumber="11">
COLOUR (
<figureCitation id="109C5F81FF804A06FEE6FB6EBF3CFB20" box="[304,373,1194,1221]" captionStart="Fig" captionStartId="10.[189,232,1999,2025]" captionTargetBox="[254,1332,1428,1949]" captionTargetId="figure@10.[249,1337,1413,1958]" captionTargetPageId="10" captionText="Fig. 8. Pogoro alopias Rosenmejer &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621670). Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928342" httpUri="https://zenodo.org/record/3928342/files/figure.png" pageId="10" pageNumber="11">Fig. 8</figureCitation>
). Face, collum, prozona, sterna and metaterga yellow/brown, legs and antennae slightly lighter.
</paragraph>
<paragraph id="88184304FF804A06FF6BFACFBB2FFAC3" blockId="10.[189,1382,1291,1318]" box="[189,1382,1291,1318]" pageId="10" pageNumber="11">
STERNAL PROCESS ON BODY RING 6 (
<figureCitation id="109C5F81FF804A06FDB4FACFBC89FAC3" box="[610,704,1291,1318]" captionStart="Fig" captionStartId="6.[189,232,1500,1526]" captionTargetBox="[217,1367,909,1445]" captionTargetId="figure@6.[207,1377,905,1453]" captionTargetPageId="6" captionText="Fig. 5. Sternal process on body ring 6. A. Emphysemastix spp. B. Agrophogonus spp. except A. hamulifer Olsen &amp; Enghoff sp. nov. C. Agrophogonus hamulifer Olsen &amp; Enghoff sp. nov. D. Pogoropsis prolixopes Rosenmejer &amp; Enghoff gen. et sp. nov. E. Pogoro siren Rosenmejer &amp; Enghoff sp. nov. F. Pogoro scharffi Hoffman, 2005. G. Pogoro alopias Rosenmejer &amp; Enghoff sp. nov. Not drawn to scale." figureDoi="http://doi.org/10.5281/zenodo.3928336" httpUri="https://zenodo.org/record/3928336/files/figure.png" pageId="10" pageNumber="11">Fig. 5G</figureCitation>
). Triangular with ‘wavy’ dark brown edge and long setae.
</paragraph>
<caption id="DCD8138CFF804A06FF6BF80BBB3EF80F" ID-DOI="http://doi.org/10.5281/zenodo.3928342" ID-Zenodo-Dep="3928342" box="[189,1399,1999,2026]" httpUri="https://zenodo.org/record/3928342/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,232,1999,2025]" targetBox="[254,1332,1428,1949]" targetPageId="10">
<paragraph id="88184304FF804A06FF6BF80BBB3EF80F" blockId="10.[189,1399,1999,2026]" box="[189,1399,1999,2026]" pageId="10" pageNumber="11">
<emphasis id="BAD39F16FF804A06FF6BF80BBF43F80C" bold="true" box="[189,266,1999,2025]" pageId="10" pageNumber="11">Fig. 8.</emphasis>
<taxonomicName id="4FA73887FF804A06FEC6F814BC92F80C" authority="Rosenmejer &amp; Enghoff" authorityName="Rosenmejer &amp; Enghoff" authorityYear="2020" box="[272,731,1999,2025]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoro" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="alopias" status="sp. nov.">
<emphasis id="BAD39F16FF804A06FEC6F814BF8DF80C" box="[272,452,1999,2025]" italics="true" pageId="10" pageNumber="11">Pogoro alopias</emphasis>
Rosenmejer &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF804A06FD34F814BD77F80F" box="[738,830,2000,2026]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NHMD 621670). Scale bar: 5 mm.
</paragraph>
</caption>
<caption id="DCD8138CFF814A07FF6BF848BCD9F808" ID-DOI="http://doi.org/10.5281/zenodo.3928344" ID-Zenodo-Dep="3928344" httpUri="https://zenodo.org/record/3928344/files/figure.png" pageId="11" pageNumber="12" startId="11.[189,232,1932,1958]" targetBox="[369,1206,265,1908]" targetPageId="11">
<paragraph id="88184304FF814A07FF6BF848BCD9F808" blockId="11.[189,1399,1932,2030]" pageId="11" pageNumber="12">
<emphasis id="BAD39F16FF814A07FF6BF848BF45F843" bold="true" box="[189,268,1932,1958]" pageId="11" pageNumber="12">Fig. 9.</emphasis>
<taxonomicName id="4FA73887FF814A07FEC2F849BCAEF843" authority="Rosenmejer &amp; Enghoff" authorityName="Rosenmejer &amp; Enghoff" authorityYear="2020" box="[276,743,1932,1959]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoro" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="alopias" status="sp. nov.">
<emphasis id="BAD39F16FF814A07FEC2F849BF83F843" box="[276,458,1932,1958]" italics="true" pageId="11" pageNumber="12">Pogoro alopias</emphasis>
Rosenmejer &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF814A07FD39F849BD04F842" box="[751,845,1933,1959]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NHMD 621670), right gonopod.
<emphasis id="BAD39F16FF814A07FF6BF874BEBDF82F" bold="true" box="[189,244,1968,1994]" pageId="11" pageNumber="12">A–B</emphasis>
. Mesal view.
<emphasis id="BAD39F16FF814A07FE40F874BF86F82F" bold="true" box="[406,463,1968,1994]" pageId="11" pageNumber="12">C–D</emphasis>
. Lateral view.
<emphasis id="BAD39F16FF814A07FDABF874BCF8F82F" bold="true" box="[637,689,1968,1994]" pageId="11" pageNumber="12">E–F</emphasis>
. Dorsal view. Abbreviations:
<emphasis id="BAD39F16FF814A07FBDBF874BA54F82C" box="[1037,1053,1968,1993]" italics="true" pageId="11" pageNumber="12">L</emphasis>
= Process L;
<emphasis id="BAD39F16FF814A07FB6BF874BA9CF82C" box="[1213,1237,1968,1993]" italics="true" pageId="11" pageNumber="12">M</emphasis>
= Process M;
<emphasis id="BAD39F16FF814A07FF6BF817BE84F808" box="[189,205,2003,2029]" italics="true" pageId="11" pageNumber="12">tl</emphasis>
= triangular lobe. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="88184304FF864A00FF6BFEC8BF9FFEAC" blockId="12.[189,1399,267,329]" pageId="12" pageNumber="13">HYPOPROCT. Almost semicircular, with small round median projection, paramedian tubercles projecting beyond edge of sclerite.</paragraph>
<paragraph id="88184304FF864A00FF6BFEA8BCCAFE15" blockId="12.[189,1398,364,496]" pageId="12" pageNumber="13">
GONOPOD (
<figureCitation id="109C5F81FF864A00FE93FEA8BFC3FE63" box="[325,394,364,391]" captionStart="Fig" captionStartId="11.[189,232,1932,1958]" captionTargetBox="[369,1206,265,1908]" captionTargetId="figure@11.[369,1216,265,1908]" captionTargetPageId="11" captionText="Fig. 9. Pogoro alopias Rosenmejer &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621670), right gonopod. A–B. Mesal view. C–D. Lateral view. E–F. Dorsal view. Abbreviations: L = Process L; M = Process M; tl = triangular lobe. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928344" httpUri="https://zenodo.org/record/3928344/files/figure.png" pageId="12" pageNumber="13">Fig. 9</figureCitation>
). Gonocoxa with paracannular setae. Telopodite endonodal. Process M and L similar in shape and size, both curved laterad. Postnodal teleopodite wide at base, narrowing shortly after, relatively short, curved dorsad. With narrow triangular subapical lobe on lateral side. Solenomere of telopodite narrow and curved anteriad.
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF864A00FF6BFDD0BDB6FDB9" box="[189,1023,532,605]" pageId="12" pageNumber="13" type="distribution">
<paragraph id="88184304FF864A00FF6BFDD0BF1FFDCB" blockId="12.[189,342,532,558]" box="[189,342,532,558]" pageId="12" pageNumber="13">
<heading id="D350F468FF864A00FF6BFDD0BF1FFDCB" bold="true" box="[189,342,532,558]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="BAD39F16FF864A00FF6BFDD0BF1FFDCB" bold="true" box="[189,342,532,558]" pageId="12" pageNumber="13">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF864A00FF6BFD87BDB6FDB9" blockId="12.[189,1023,578,605]" box="[189,1023,578,605]" pageId="12" pageNumber="13">
Known only from the
<typeStatus id="571CFDA6FF864A00FE14FD87BFBCFDB8" box="[450,501,579,605]" pageId="12" pageNumber="13">type</typeStatus>
locality in the Udzungwa Scarp FR (
<figureCitation id="109C5F81FF864A00FC7FFD86BDB8FDB9" box="[937,1009,578,604]" captionStart="Fig" captionStartId="2.[189,232,1966,1992]" captionTargetBox="[199,1398,1100,1897]" captionTargetId="figure@2.[194,1482,1044,1958]" captionTargetPageId="2" captionText="Fig. 2. Map of the Udzungwa Mts showing collection sites for gomphodesmids. Based on Marshall et al. (2010: fig. 1)." figureDoi="http://doi.org/10.5281/zenodo.3928330" httpUri="https://zenodo.org/record/3928330/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

149
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<mods:title id="E2635406A9471931D1C383313C64A49C">A mountain of millipedes IX: Species of the family Gomphodesmidae from the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida)</mods:title>
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<mods:namePart id="DCEAA0A132C77645CF54B53F57F7971D">Olsen, Sissel Anna</mods:namePart>
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<mods:affiliation id="7E43FEB469EF80A58F9A251D02DDF3BC">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 København Ø, Denmark</mods:affiliation>
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<mods:namePart id="DB998BFAB2E7C693DF97E44CAE4A48C3">Rosenmejer, Trine</mods:namePart>
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species
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<emphasis id="BAD39F16FF844A02FF6BFAADBF8DFA67" box="[189,452,1384,1410]" italics="true" pageId="14" pageNumber="15">Pogoropsis prolixopes</emphasis>
Rosenmejer &amp; Enghoff
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Moderate/small-sized (length
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), convex gomphodesmids, pore formula: 5, 7, 9–19.
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Four apical sensory cones on antennae, fossa present on 5
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and 6
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antennomeres. Paranota slightly declivent. Stricture poorly defined, with visible edge only in front of anterior spiracle. Pleurosternal carinae present on body rings 1–17. Anterior legs without ventral tubercles, with sparse setation. Setae on dorsal side of tarsi shorter and stouter than the rest. Subonychial pads on legs 1–6. Sharing most of these characters with
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<emphasis id="BAD39F16FF844A02FE37F957BC71F949" box="[481,568,1683,1708]" italics="true" pageId="14" pageNumber="15">Pogoro</emphasis>
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but differing by: sternal process of segment 6 longer than wide, with triangular tip. Gonopod telopodite relatively long and slender, making a hairpin bend on the middle, so the postnodal telepodite lies close to the prefemoral region. Nodal process L and paracannular setae absent.
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The genus name refers to the similarity to the genus
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.
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Gender feminine.
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<emphasis id="BAD39F16FF844A02FF6BF866BFC0F859" bold="true" box="[189,393,1954,1980]" pageId="14" pageNumber="15">Included species</emphasis>
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<heading id="D350F468FF854A03FE7BFECFBACEFEC3" box="[429,1159,266,294]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="16" reason="2">
<taxonomicName id="4FA73887FF854A03FE7BFECFBD99FEC0" ID-CoL="6eb43d6a-9930-4256-99a9-aecb646378b8" authority="Rosenmejer &amp; Enghoff" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[429,976,266,293]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoropsis" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="prolixopes" status="gen. et sp. nov.">
<emphasis id="BAD39F16FF854A03FE7BFECFBCFEFEC0" bold="true" box="[429,695,266,293]" italics="true" pageId="15" pageNumber="16">Pogoropsis prolixopes</emphasis>
Rosenmejer &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF854A03FC01FEC8BACEFEC3" box="[983,1159,268,294]" pageId="15" pageNumber="16" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF854A03FEBBFEEABDCDFE89" pageId="15" pageNumber="16" type="description">
<paragraph id="88184304FF854A03FEBBFEEABA8FFEAC" blockId="15.[365,1222,266,364]" box="[365,1222,302,329]" pageId="15" pageNumber="16">
<uri id="FC364F06FF854A03FEBBFEEABA8FFEAC" box="[365,1222,302,329]" pageId="15" pageNumber="16">
urn:lsid:zoobank.org:act:
<uuid id="FC0179D1FF854A03FD5CFEEABA8FFEAC" box="[650,1222,302,329]" pageId="15" pageNumber="16">D35FD14F-0709-4AF2-945F-AEDA96FF91E3</uuid>
</uri>
</paragraph>
<paragraph id="88184304FF854A03FD66FE95BDCDFE89" blockId="15.[365,1222,266,364]" box="[688,900,337,364]" pageId="15" pageNumber="16">
<figureCitation id="109C5F81FF854A03FD66FE95BCB2FE89" box="[688,763,337,364]" captionStart="Fig" captionStartId="2.[189,232,1966,1992]" captionTargetBox="[199,1398,1100,1897]" captionTargetId="figure@2.[194,1482,1044,1958]" captionTargetPageId="2" captionText="Fig. 2. Map of the Udzungwa Mts showing collection sites for gomphodesmids. Based on Marshall et al. (2010: fig. 1)." figureDoi="http://doi.org/10.5281/zenodo.3928330" httpUri="https://zenodo.org/record/3928330/files/figure.png" pageId="15" pageNumber="16">Figs 2</figureCitation>
,
<figureCitation id="109C5F81FF854A03FCD1FE95BD67FE89" box="[775,814,337,364]" captionStart="Fig" captionStartId="6.[189,232,1500,1526]" captionTargetBox="[217,1367,909,1445]" captionTargetId="figure@6.[207,1377,905,1453]" captionTargetPageId="6" captionText="Fig. 5. Sternal process on body ring 6. A. Emphysemastix spp. B. Agrophogonus spp. except A. hamulifer Olsen &amp; Enghoff sp. nov. C. Agrophogonus hamulifer Olsen &amp; Enghoff sp. nov. D. Pogoropsis prolixopes Rosenmejer &amp; Enghoff gen. et sp. nov. E. Pogoro siren Rosenmejer &amp; Enghoff sp. nov. F. Pogoro scharffi Hoffman, 2005. G. Pogoro alopias Rosenmejer &amp; Enghoff sp. nov. Not drawn to scale." figureDoi="http://doi.org/10.5281/zenodo.3928336" httpUri="https://zenodo.org/record/3928336/files/figure.png" pageId="15" pageNumber="16">5D</figureCitation>
,
<figureCitation id="109C5F81FF854A03FCECFE96BDCDFE89" box="[826,900,338,364]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="15.[189,232,1931,1957]" captionStartId-1="16.[189,232,1929,1955]" captionTargetBox-0="[534,1043,1079,1897]" captionTargetBox-1="[202,1375,266,1896]" captionTargetId-0="figure@15.[516,1062,1073,1897]" captionTargetId-1="figure@16.[202,1384,265,1896]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="Fig. 12. Pogoropsis prolixopes Rosenmejer &amp; Enghoff gen. et sp. nov., holotype, ♂ (NHMD 621672). A. Habitus. B. Front end, showing gonopods, ventral view. C. Hind end, ventral view. Scale bars: A = 5 mm; B = 3 mm; C = 1 mm." captionText-1="Fig. 13. Pogoropsis prolixopes Rosenmejer &amp; Enghoff gen. et sp. nov., paratype, ♂, left gonopod (NHMD 621673). A–B. Mesal view. C. Apex of solenomere. D–E. Lateral view. F–G. Dorsal view. Abbreviations: M = Process M; stl = slender triangular lobe; tl = triangular lobe. Scale bars: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3928350" figureDoi-1="http://doi.org/10.5281/zenodo.3928352" httpUri-0="https://zenodo.org/record/3928350/files/figure.png" httpUri-1="https://zenodo.org/record/3928352/files/figure.png" pageId="15" pageNumber="16">12–13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF854A03FF6BFE57BC38FE1B" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="88184304FF854A03FF6BFE57BF7FFE48" blockId="15.[189,310,403,429]" box="[189,310,403,429]" pageId="15" pageNumber="16">
<heading id="D350F468FF854A03FF6BFE57BF7FFE48" bold="true" box="[189,310,403,429]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="BAD39F16FF854A03FF6BFE57BF7FFE48" bold="true" box="[189,310,403,429]" pageId="15" pageNumber="16">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF854A03FF6BFE06BC38FE1B" blockId="15.[189,1398,449,510]" pageId="15" pageNumber="16">As for the genus. Differs conspicuously from other Udzungwan gomphodesmids species by the very long and slender gonopod telopodite.</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF854A03FF6BFDE2BF7EFD77" pageId="15" pageNumber="16" type="etymology">
<paragraph id="88184304FF854A03FF6BFDE2BF0DFDA5" blockId="15.[189,324,550,576]" box="[189,324,550,576]" pageId="15" pageNumber="16">
<heading id="D350F468FF854A03FF6BFDE2BF0DFDA5" bold="true" box="[189,324,550,576]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="BAD39F16FF854A03FF6BFDE2BF0DFDA5" bold="true" box="[189,324,550,576]" pageId="15" pageNumber="16">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF854A03FF6BFD90BF7EFD77" blockId="15.[189,1398,596,658]" pageId="15" pageNumber="16">The name is a Latin noun in apposition meaning “long leg/foot” and refers to the unusually long gonopods.</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF854A03FF6BFD7DBAB8FC11" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="88184304FF854A03FF6BFD7DBFE3FD36" blockId="15.[189,426,697,723]" box="[189,426,697,723]" pageId="15" pageNumber="16">
<heading id="D350F468FF854A03FF6BFD7DBFE3FD36" bold="true" box="[189,426,697,723]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="BAD39F16FF854A03FF6BFD7DBFE3FD36" bold="true" box="[189,426,697,723]" pageId="15" pageNumber="16">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF854A03FF3CFD2CBF15FCE7" blockId="15.[189,1399,744,841]" box="[234,348,744,770]" pageId="15" pageNumber="16">
<emphasis id="BAD39F16FF854A03FF3CFD2CBF15FCE7" bold="true" box="[234,348,744,770]" pageId="15" pageNumber="16">
<typeStatus id="571CFDA6FF854A03FF3CFD2CBF15FCE7" box="[234,348,744,770]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="88184304FF854A03FF6BFCCFBB3EFCAD" blockId="15.[189,1399,744,841]" pageId="15" pageNumber="16">
<materialsCitation id="38CF4959FF854A03FF6BFCCFBB3EFCAD" ID-GBIF-Occurrence="2803530302" collectingDate="2000-12-07" collectingMethod="open woodland" collectionCode="NHMD" collectorName="Frontier Tanzania UMBS" country="Tanzania" elevation="1510" latitude="-7.7595" location="West Kilombero Scarp Forest" longLatPrecision="1" longitude="36.44372" pageId="15" pageNumber="16" specimenCode="NHMD 621672" specimenCount="1" stateProvince="Iringa region" typeStatus="holotype">
<collectingCountry id="F0B00394FF854A03FF6BFCCFBF18FCC0" box="[189,337,779,805]" name="Tanzania" pageId="15" pageNumber="16">TANZANIA</collectingCountry>
•
<specimenCount id="9EA1888DFF854A03FEB8FCCFBFCFFCC0" box="[366,390,779,805]" pageId="15" pageNumber="16" type="male">♂</specimenCount>
;
<collectingRegion id="4A638DE6FF854A03FE40FCCFBC78FCC0" box="[406,561,779,805]" country="Tanzania" name="Iringa" pageId="15" pageNumber="16">Iringa region</collectingRegion>
/district,
<location id="8D7815DFFF854A03FD4DFCCFBA48FCC0" LSID="urn:lsid:plazi:treatment:000EF212FF854A1DFE7BFECFBFABFA5A:8D7815DFFF854A03FD4DFCCFBA48FCC0" box="[667,1025,779,805]" country="Tanzania" latitude="-7.7595" longLatPrecision="1" longitude="36.44372" name="West Kilombero Scarp Forest" pageId="15" pageNumber="16" stateProvince="Iringa region">West Kilombero Scarp Forest</location>
;
<geoCoordinate id="ED9325C3FF854A03FBD9FCCDBAFEFCC0" box="[1039,1207,777,805]" degrees="07" direction="south" minutes="45" orientation="latitude" pageId="15" pageNumber="16" precision="1" seconds="34.2" value="-7.7595">07°45′34.2″ S</geoCoordinate>
,
<geoCoordinate id="ED9325C3FF854A03FB10FCCDBB38FCC0" box="[1222,1393,777,805]" degrees="36" direction="east" minutes="26" orientation="longitude" pageId="15" pageNumber="16" precision="1" seconds="37.4" value="36.44372">36°26′37.4″ E</geoCoordinate>
;
<elevation id="038AA437FF854A03FF6BFCEABF1DFCAD" box="[189,340,814,841]" metricMagnitude="3" metricUnit="m" metricValue="1.51" pageId="15" pageNumber="16" unit="m" value="1510.0">
<quantity id="4F5FEEE1FF854A03FF6BFCEABF5CFCAC" box="[189,277,814,841]" metricMagnitude="3" metricUnit="m" metricValue="1.51" pageId="15" pageNumber="16" unit="m" value="1510.0">1510 m</quantity>
a.s.l.
</elevation>
;
<date id="FC1965C4FF854A03FEB6FCEBBFB9FCAD" box="[352,496,814,841]" pageId="15" pageNumber="16" value="2000-12-07">
<collectingDate id="EC5D9C2CFF854A03FEB6FCEBBFB9FCAD" box="[352,496,814,841]" pageId="15" pageNumber="16" value="2000-12-07">7 Dec. 2000</collectingDate>
</date>
;
<collectorName id="255226D2FF854A03FE2DFCEABD6BFCAD" box="[507,802,814,840]" pageId="15" pageNumber="16">Frontier Tanzania UMBS</collectorName>
leg.;
<collectingMethod id="51E63B13FF854A03FCB5FCEBBA50FCAD" box="[867,1049,814,841]" pageId="15" pageNumber="16">open woodland</collectingMethod>
, plot
<taxonomicName id="4FA73887FF854A03FB81FCEABAE1FCAD" box="[1111,1192,814,840]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="genus">
<emphasis id="BAD39F16FF854A03FB81FCEABAE1FCAD" box="[1111,1192,814,840]" italics="true" pageId="15" pageNumber="16">Acacia</emphasis>
</taxonomicName>
;
<specimenCode id="D801EB7FFF854A03FB61FCEBBB3AFCAD" box="[1207,1395,814,841]" collectionCode="NHMD" pageId="15" pageNumber="16">NHMD 621672</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="88184304FF854A03FF3CFCB4BF2EFC6F" blockId="15.[189,1399,880,1012]" box="[234,359,880,906]" pageId="15" pageNumber="16">
<emphasis id="BAD39F16FF854A03FF3CFCB4BF2EFC6F" bold="true" box="[234,359,880,906]" pageId="15" pageNumber="16">
<typeStatus id="571CFDA6FF854A03FF3CFCB4BF2EFC6F" box="[234,359,880,906]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="88184304FF854A03FF6BFC57BAB8FC11" blockId="15.[189,1399,880,1012]" pageId="15" pageNumber="16">
<materialsCitation id="38CF4959FF854A03FF6BFC57BAC3FC48" ID-GBIF-Occurrence="2803530313" box="[189,1162,914,942]" collectingDate="2000-12-07" collectionCode="NHMD" country="Tanzania" elevation="1510" latitude="-7.7595" location="TANZANIA" longLatPrecision="1" longitude="36.44372" pageId="15" pageNumber="16" specimenCode="NHMD 621673" specimenCount="21" specimenCount-female="2" specimenCount-male="19" stateProvince="Iringa" typeStatus="paratype">
<collectingCountry id="F0B00394FF854A03FF6BFC57BF1BFC48" box="[189,338,915,941]" name="Tanzania" pageId="15" pageNumber="16">TANZANIA</collectingCountry>
•
<specimenCount id="9EA1888DFF854A03FEBDFC57BFF2FC4B" box="[363,443,914,942]" pageId="15" pageNumber="16" type="male">19 ♂♂</specimenCount>
,
<specimenCount id="9EA1888DFF854A03FE1CFC57BC44FC48" box="[458,525,915,941]" pageId="15" pageNumber="16" type="female">2 ♀♀</specimenCount>
;
<geologicalTimeScale id="FA87C3AFFF854A03FDCDFC57BD8CFC48" box="[539,965,915,941]" pageId="15" pageNumber="16">same collecting data as for holotype</geologicalTimeScale>
;
<specimenCode id="D801EB7FFF854A03FC07FC57BAC3FC48" box="[977,1162,915,941]" collectionCode="NHMD" pageId="15" pageNumber="16">NHMD 621673</specimenCode>
</materialsCitation>
•
<materialsCitation id="38CF4959FF854A03FB75FC57BAA4FC11" ID-GBIF-Occurrence="2803530314" collectedFrom="montane forest" collectingDate="2000-11-23" collectionCode="NHMD" collectorName="Frontier Tanzania UMBS" country="Tanzania" elevation="1111" latitude="-7.88875" location="West Kilombero Scarp Forest" longLatPrecision="1" longitude="36.38656" pageId="15" pageNumber="16" specimenCode="NHMD 621685" specimenCount="5" specimenCount-female="2" specimenCount-male="1" stateProvince="Iringa region" typeStatus="paratype">
<specimenCount id="9EA1888DFF854A03FB75FC57BA87FC4B" box="[1187,1230,914,942]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
,
<specimenCount id="9EA1888DFF854A03FB0BFC57BB69FC48" box="[1245,1312,915,941]" pageId="15" pageNumber="16" type="female">2 ♀♀</specimenCount>
,
<specimenCount id="9EA1888DFF854A03FAFBFC57BB3BFC48" box="[1325,1394,915,941]" pageId="15" pageNumber="16" type="generic">2 juv.</specimenCount>
;
<collectingRegion id="4A638DE6FF854A03FF6BFC72BF11FC35" box="[189,344,950,976]" country="Tanzania" name="Iringa" pageId="15" pageNumber="16">Iringa region</collectingRegion>
/district,
<location id="8D7815DFFF854A03FE15FC73BD62FC34" LSID="urn:lsid:plazi:treatment:000EF212FF854A1DFE7BFECFBFABFA5A:8D7815DFFF854A03FE15FC73BD62FC34" box="[451,811,950,977]" country="Tanzania" latitude="-7.88875" longLatPrecision="1" longitude="36.38656" name="West Kilombero Scarp Forest" pageId="15" pageNumber="16" stateProvince="Iringa region">West Kilombero Scarp Forest</location>
;
<geoCoordinate id="ED9325C3FF854A03FCECFC70BDAAFC35" box="[826,995,948,976]" degrees="07" direction="south" minutes="53" orientation="latitude" pageId="15" pageNumber="16" precision="1" seconds="19.5" value="-7.88875">07°53′19.5″ S</geoCoordinate>
,
<geoCoordinate id="ED9325C3FF854A03FC25FC70BAD4FC34" box="[1011,1181,948,977]" degrees="36" direction="east" minutes="23" orientation="longitude" pageId="15" pageNumber="16" precision="1" seconds="11.6" value="36.38656">36°23′11.6″ E</geoCoordinate>
;
<elevation id="038AA437FF854A03FB7BFC72BB00FC35" box="[1197,1353,950,977]" metricMagnitude="3" metricUnit="m" metricValue="1.111" pageId="15" pageNumber="16" unit="m" value="1111.0">
<quantity id="4F5FEEE1FF854A03FB7BFC72BB4FFC34" box="[1197,1286,950,977]" metricMagnitude="3" metricUnit="m" metricValue="1.111" pageId="15" pageNumber="16" unit="m" value="1111.0">1111 m</quantity>
a.s.l.
</elevation>
;
<date id="FC1965C4FF854A03FA8FFC72BF73FC16" pageId="15" pageNumber="16" value="2000-11-23">
<collectingDate id="EC5D9C2CFF854A03FA8FFC72BF73FC16" pageId="15" pageNumber="16" value="2000-11-23">23 Nov. 2000</collectingDate>
</date>
;
<collectorName id="255226D2FF854A03FE90FC1DBC26FC16" box="[326,623,985,1011]" pageId="15" pageNumber="16">Frontier Tanzania UMBS</collectorName>
leg.;
<collectedFrom id="3BBB00F9FF854A03FD64FC1EBD2DFC16" box="[690,868,985,1012]" pageId="15" pageNumber="16">montane forest</collectedFrom>
, plot 17, Ukami;
<specimenCode id="D801EB7FFF854A03FBE6FC1EBAA4FC11" box="[1072,1261,985,1012]" collectionCode="NHMD" pageId="15" pageNumber="16">NHMD 621685</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DCD8138CFF854A03FF6BF84FBC5DF809" ID-DOI="http://doi.org/10.5281/zenodo.3928350" ID-Zenodo-Dep="3928350" httpUri="https://zenodo.org/record/3928350/files/figure.png" pageId="15" pageNumber="16" startId="15.[189,232,1931,1957]" targetBox="[534,1043,1079,1897]" targetPageId="15">
<paragraph id="88184304FF854A03FF6BF84FBC5DF809" blockId="15.[189,1399,1931,2028]" pageId="15" pageNumber="16">
<emphasis id="BAD39F16FF854A03FF6BF84FBF53F840" bold="true" box="[189,282,1931,1957]" pageId="15" pageNumber="16">Fig. 12.</emphasis>
<taxonomicName id="4FA73887FF854A03FEF4F848BD0FF840" authority="Rosenmejer &amp; Enghoff" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[290,838,1931,1957]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoropsis" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="prolixopes" status="gen. et sp. nov.">
<emphasis id="BAD39F16FF854A03FEF4F848BC63F840" box="[290,554,1931,1957]" italics="true" pageId="15" pageNumber="16">Pogoropsis prolixopes</emphasis>
Rosenmejer &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF854A03FC98F848BA4AF843" box="[846,1027,1932,1958]" pageId="15" pageNumber="16" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NHMD 621672).
<emphasis id="BAD39F16FF854A03FF6BF86ABE9BF82D" bold="true" box="[189,210,1966,1992]" pageId="15" pageNumber="16">A</emphasis>
. Habitus.
<emphasis id="BAD39F16FF854A03FE9BF86BBF28F82C" bold="true" box="[333,353,1967,1993]" pageId="15" pageNumber="16">B</emphasis>
. Front end, showing gonopods, ventral view.
<emphasis id="BAD39F16FF854A03FC56F86ABDDCF82D" bold="true" box="[896,917,1966,1992]" pageId="15" pageNumber="16">C</emphasis>
. Hind end, ventral view. Scale bars: A = 5 mm; B = 3 mm; C = 1 mm.
</paragraph>
</caption>
<caption id="DCD8138CFF9A4A1CFF6BF84DBB13F80F" ID-DOI="http://doi.org/10.5281/zenodo.3928352" ID-Zenodo-Dep="3928352" httpUri="https://zenodo.org/record/3928352/files/figure.png" pageId="16" pageNumber="17" startId="16.[189,232,1929,1955]" targetBox="[202,1375,266,1896]" targetPageId="16">
<paragraph id="88184304FF9A4A1CFF6BF84DBB13F80F" blockId="16.[189,1399,1928,2026]" pageId="16" pageNumber="17">
<emphasis id="BAD39F16FF9A4A1CFF6BF84DBF57F846" bold="true" box="[189,286,1929,1955]" pageId="16" pageNumber="17">Fig. 13.</emphasis>
<taxonomicName id="4FA73887FF9A4A1CFEFCF84DBD14F846" authority="Rosenmejer &amp; Enghoff" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[298,861,1928,1955]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoropsis" kingdom="Animalia" order="Polydesmida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="prolixopes" status="gen. et sp. nov.">
<emphasis id="BAD39F16FF9A4A1CFEFCF84DBC7FF847" box="[298,566,1928,1954]" italics="true" pageId="16" pageNumber="17">Pogoropsis prolixopes</emphasis>
Rosenmejer &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF9A4A1CFCBFF84DBA63F846" box="[873,1066,1929,1955]" pageId="16" pageNumber="17" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, paratype, ♂, left gonopod (NHMD 621673).
<emphasis id="BAD39F16FF9A4A1CFE4BF868BF9DF823" bold="true" box="[413,468,1964,1990]" pageId="16" pageNumber="17">A–B</emphasis>
. Mesal view.
<emphasis id="BAD39F16FF9A4A1CFD56F868BCDCF823" bold="true" box="[640,661,1964,1990]" pageId="16" pageNumber="17">C</emphasis>
. Apex of solenomere.
<emphasis id="BAD39F16FF9A4A1CFC7FF868BDA9F823" bold="true" box="[937,992,1964,1990]" pageId="16" pageNumber="17">D–E</emphasis>
. Lateral view.
<emphasis id="BAD39F16FF9A4A1CFB41F868BA87F823" bold="true" box="[1175,1230,1964,1990]" pageId="16" pageNumber="17">F–G</emphasis>
. Dorsal view. Abbreviations:
<emphasis id="BAD39F16FF9A4A1CFEA5F814BFC2F80C" box="[371,395,2000,2025]" italics="true" pageId="16" pageNumber="17">M</emphasis>
= Process M;
<emphasis id="BAD39F16FF9A4A1CFDE3F80BBC18F80C" box="[565,593,1999,2025]" italics="true" pageId="16" pageNumber="17">stl</emphasis>
= slender triangular lobe;
<emphasis id="BAD39F16FF9A4A1CFC51F80BBDDEF80C" box="[903,919,1999,2025]" italics="true" pageId="16" pageNumber="17">tl</emphasis>
= triangular lobe. Scale bars: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C0BD108FFF9B4A1DFF6BFECFBD33FCBA" pageId="17" pageNumber="18" type="description">
<paragraph id="88184304FF9B4A1DFF6BFECFBF07FEC0" blockId="17.[189,334,267,293]" box="[189,334,267,293]" pageId="17" pageNumber="18">
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<emphasis id="BAD39F16FF9B4A1DFF6BFECFBF07FEC0" bold="true" box="[189,334,267,293]" pageId="17" pageNumber="18">Description</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF9B4A1DFF3CFEF8BF7CFEB3" blockId="17.[189,1398,316,418]" box="[234,309,316,342]" pageId="17" pageNumber="18">
<emphasis id="BAD39F16FF9B4A1DFF3CFEF8BF7CFEB3" bold="true" box="[234,309,316,342]" pageId="17" pageNumber="18">Males</emphasis>
</paragraph>
<paragraph id="88184304FF9B4A1DFF6BFEA6BCE2FE47" blockId="17.[189,1398,316,418]" pageId="17" pageNumber="18">
SIZE AND SHAPE. Body length ca
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. Maximum width 6.0–
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. W/L ratio 20%. Height of body ring eight
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. H/W ratio= 68%.
</paragraph>
<paragraph id="88184304FF9B4A1DFF6BFE0CBD7EFE07" blockId="17.[189,823,456,482]" box="[189,823,456,482]" pageId="17" pageNumber="18">
COLOUR (
<figureCitation id="109C5F81FF9B4A1DFEE6FE0CBFD5FE07" box="[304,412,456,482]" captionStart="Fig" captionStartId="15.[189,232,1931,1957]" captionTargetBox="[534,1043,1079,1897]" captionTargetId="figure@15.[516,1062,1073,1897]" captionTargetPageId="15" captionText="Fig. 12. Pogoropsis prolixopes Rosenmejer &amp; Enghoff gen. et sp. nov., holotype, ♂ (NHMD 621672). A. Habitus. B. Front end, showing gonopods, ventral view. C. Hind end, ventral view. Scale bars: A = 5 mm; B = 3 mm; C = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3928350" httpUri="https://zenodo.org/record/3928350/files/figure.png" pageId="17" pageNumber="18">Fig. 12A</figureCitation>
). Uniformly yellowish/light brown
</paragraph>
<paragraph id="88184304FF9B4A1DFF6BFDCCBA1BFDC7" blockId="17.[189,1106,520,547]" box="[189,1106,520,547]" pageId="17" pageNumber="18">
STERNAL PROCESS ON BODY RING 6 (
<figureCitation id="109C5F81FF9B4A1DFDB4FDCCBC89FDC6" box="[610,704,520,547]" captionStart="Fig" captionStartId="6.[189,232,1500,1526]" captionTargetBox="[217,1367,909,1445]" captionTargetId="figure@6.[207,1377,905,1453]" captionTargetPageId="6" captionText="Fig. 5. Sternal process on body ring 6. A. Emphysemastix spp. B. Agrophogonus spp. except A. hamulifer Olsen &amp; Enghoff sp. nov. C. Agrophogonus hamulifer Olsen &amp; Enghoff sp. nov. D. Pogoropsis prolixopes Rosenmejer &amp; Enghoff gen. et sp. nov. E. Pogoro siren Rosenmejer &amp; Enghoff sp. nov. F. Pogoro scharffi Hoffman, 2005. G. Pogoro alopias Rosenmejer &amp; Enghoff sp. nov. Not drawn to scale." figureDoi="http://doi.org/10.5281/zenodo.3928336" httpUri="https://zenodo.org/record/3928336/files/figure.png" pageId="17" pageNumber="18">Fig. 5D</figureCitation>
). Longer than wide, triangular tip.
</paragraph>
<paragraph id="88184304FF9B4A1DFF6BFD8CBCF4FD6D" blockId="17.[189,1398,584,649]" pageId="17" pageNumber="18">
HYPOPROCT. Suboval in shape, without median projection (
<figureCitation id="109C5F81FF9B4A1DFCADFD8CBDA1FD87" box="[891,1000,584,610]" captionStart="Fig" captionStartId="15.[189,232,1931,1957]" captionTargetBox="[534,1043,1079,1897]" captionTargetId="figure@15.[516,1062,1073,1897]" captionTargetPageId="15" captionText="Fig. 12. Pogoropsis prolixopes Rosenmejer &amp; Enghoff gen. et sp. nov., holotype, ♂ (NHMD 621672). A. Habitus. B. Front end, showing gonopods, ventral view. C. Hind end, ventral view. Scale bars: A = 5 mm; B = 3 mm; C = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3928350" httpUri="https://zenodo.org/record/3928350/files/figure.png" pageId="17" pageNumber="18">Fig. 12C</figureCitation>
). Paramedian tubercles of normal size, not projecting beyond edge of sclerite.
</paragraph>
<paragraph id="88184304FF9B4A1DFF6BFD6ABD33FCBA" blockId="17.[189,1399,686,864]" pageId="17" pageNumber="18">
GONOPOD (
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). Gonocoxa without paracannular setae. Telopodite mesonodal. Nodus with few small triangular processes. Process M thin, narrow at base, and closely following postnodal telopodite. Process L absent. Postnodal teleopodite relatively long, curved dorsomesad, closely following prefemoral region. Solenomere with triangular lobe, followed by longer and more slender triangular lobe. Apex thin and curved, projecting at right angles before end of solenomere.
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF9B4A1DFF6BFC42BD56FBF9" pageId="17" pageNumber="18" type="distribution">
<paragraph id="88184304FF9B4A1DFF6BFC42BF1FFC45" blockId="17.[189,342,902,928]" box="[189,342,902,928]" pageId="17" pageNumber="18">
<heading id="D350F468FF9B4A1DFF6BFC42BF1FFC45" bold="true" box="[189,342,902,928]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="BAD39F16FF9B4A1DFF6BFC42BF1FFC45" bold="true" box="[189,342,902,928]" pageId="17" pageNumber="18">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF9B4A1DFF6BFC73BD56FBF9" blockId="17.[189,1398,951,1053]" pageId="17" pageNumber="18">
Known only from the
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locality and one further site in the Udzungwa Mts, West Kilombero Scarp, montane forest and open woodland (
<figureCitation id="109C5F81FF9B4A1DFDB5FC18BCE3FC12" box="[611,682,988,1015]" captionStart="Fig" captionStartId="2.[189,232,1966,1992]" captionTargetBox="[199,1398,1100,1897]" captionTargetId="figure@2.[194,1482,1044,1958]" captionTargetPageId="2" captionText="Fig. 2. Map of the Udzungwa Mts showing collection sites for gomphodesmids. Based on Marshall et al. (2010: fig. 1)." figureDoi="http://doi.org/10.5281/zenodo.3928330" httpUri="https://zenodo.org/record/3928330/files/figure.png" pageId="17" pageNumber="18">Fig. 2</figureCitation>
). The two sites are ca
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apart and are situated in different forested areas (cf.
<bibRefCitation id="EC363EF5FF9B4A1DFE40FBC6BC9FFBF9" author="Frontier Tanzania &amp; Doody K. Z. &amp; Howell K. M. &amp; Fanning E." box="[406,726,1026,1052]" pageId="17" pageNumber="18" refId="ref11841" refString="Frontier Tanzania 2001. West Kilombero Scarp Forest Reserve - Zoological Report. In: Doody K. Z., Howell K. M. &amp; Fanning E. (eds) Report for the Udzungwa Mountains Forest Management and Biodiversity Conservation Project, MEMA, Iringa, Tanzania. Available from http: // www. easternarc. or. tz / groups / webcontent / documents / pdf / WKSFR % 20 Zoological % 20 Report. PDF [accessed 27 May 2020]." type="book" year="2001">
Frontier
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2001: 26
</bibRefCitation>
, 174).
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF9B4A1DFF6BFB83BFABFA5A" pageId="17" pageNumber="18" type="discussion">
<paragraph id="88184304FF9B4A1DFF6BFB83BF66FB84" blockId="17.[189,303,1095,1121]" box="[189,303,1095,1121]" pageId="17" pageNumber="18">
<heading id="D350F468FF9B4A1DFF6BFB83BF66FB84" bold="true" box="[189,303,1095,1121]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="BAD39F16FF9B4A1DFF6BFB83BF66FB84" bold="true" box="[189,303,1095,1121]" pageId="17" pageNumber="18">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF9B4A1DFF6BFBB3BFABFA5A" blockId="17.[189,1399,1142,1471]" pageId="17" pageNumber="18">
The shape of the sternal process on the 6
<superScript id="7FD2EE4CFF9B4A1DFD4DFBB2BCE1FB61" attach="left" box="[667,680,1142,1156]" fontSize="6" pageId="17" pageNumber="18">th</superScript>
body ring is more similar to that seen in species of the genus
<taxonomicName id="4FA73887FF9B4A1DFF6BFB59BC4AFB5D" authority="Hoffman, 2005" authorityName="Hoffman" authorityYear="2005" box="[189,515,1181,1208]" class="Diplopoda" family="Gomphodesmidae" genus="Ngurubates" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="BAD39F16FF9B4A1DFF6BFB59BF0FFB52" box="[189,326,1181,1207]" italics="true" pageId="17" pageNumber="18">Ngurubates</emphasis>
<bibRefCitation id="EC363EF5FF9B4A1DFE9BFB59BC4AFB5D" author="Hoffman R. L." box="[333,515,1181,1208]" pageId="17" pageNumber="18" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman, 2005</bibRefCitation>
</taxonomicName>
, than to that of species of
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<emphasis id="BAD39F16FF9B4A1DFCE0FB5ABDC4FB52" box="[822,909,1182,1207]" italics="true" pageId="17" pageNumber="18">Pogoro</emphasis>
</taxonomicName>
. In the key to genera of
<taxonomicName id="4FA73887FF9B4A1DFB7AFB59BFC8FB3B" authority=", Hoffman (2005)" authorityName="Hoffman" authorityYear="2005" class="Diplopoda" family="Gomphodesmidae" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="tribe" tribe="Gomphodesmini">
Gomphodesmini,
<bibRefCitation id="EC363EF5FF9B4A1DFF6BFB07BFC8FB3B" author="Hoffman R. L." box="[189,385,1219,1246]" pageId="17" pageNumber="18" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman (2005)</bibRefCitation>
</taxonomicName>
actually uses this character to distinguish between
<taxonomicName id="4FA73887FF9B4A1DFBD5FB00BA13FB38" authorityName="Hoffman" authorityYear="2005" box="[1027,1114,1220,1245]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoro" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="BAD39F16FF9B4A1DFBD5FB00BA13FB38" box="[1027,1114,1220,1245]" italics="true" pageId="17" pageNumber="18">Pogoro</emphasis>
</taxonomicName>
and
<taxonomicName id="4FA73887FF9B4A1DFB4BFB07BB6FFB38" authorityName="Hoffman" authorityYear="2005" box="[1181,1318,1219,1245]" class="Diplopoda" family="Gomphodesmidae" genus="Ngurubates" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="BAD39F16FF9B4A1DFB4BFB07BB6FFB38" box="[1181,1318,1219,1245]" italics="true" pageId="17" pageNumber="18">Ngurubates</emphasis>
</taxonomicName>
. After examining both genus descriptions, however, it became clear that
<taxonomicName id="4FA73887FF9B4A1DFC6AFB2DBA8BFAE6" authority="Olsen &amp; Rosenmejer &amp; Enghoff, 2020" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[956,1218,1257,1283]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoropsis" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="prolixopes" status="gen. et sp. nov.">
<emphasis id="BAD39F16FF9B4A1DFC6AFB2DBA8BFAE6" box="[956,1218,1257,1283]" italics="true" pageId="17" pageNumber="18">Pogoropsis prolixopes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF9B4A1DFB1FFB2DBB3EFAE6" box="[1225,1399,1257,1283]" pageId="17" pageNumber="18" rank="species">gen. et sp. nov.</taxonomicNameLabel>
fits into neither of these genera.
<taxonomicName id="4FA73887FF9B4A1DFDEBFACABC8FFACD" authorityName="Hoffman" authorityYear="2005" box="[573,710,1294,1320]" class="Diplopoda" family="Gomphodesmidae" genus="Ngurubates" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="BAD39F16FF9B4A1DFDEBFACABC8FFACD" box="[573,710,1294,1320]" italics="true" pageId="17" pageNumber="18">Ngurubates</emphasis>
</taxonomicName>
species have subonychial pads only on legs 1–5 (
<taxonomicName id="4FA73887FF9B4A1DFAC1FACBBB27FACD" authorityName="Hoffman" authorityYear="2005" box="[1303,1390,1295,1320]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoro" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="BAD39F16FF9B4A1DFAC1FACBBB27FACD" box="[1303,1390,1295,1320]" italics="true" pageId="17" pageNumber="18">Pogoro</emphasis>
</taxonomicName>
: 1–6), and ventral tubercles on anterior legs are present (
<taxonomicName id="4FA73887FF9B4A1DFC8AFAF1BDFAFAAB" authorityName="Hoffman" authorityYear="2005" box="[860,947,1333,1358]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoro" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="BAD39F16FF9B4A1DFC8AFAF1BDFAFAAB" box="[860,947,1333,1358]" italics="true" pageId="17" pageNumber="18">Pogoro</emphasis>
</taxonomicName>
: absent). Almost all characteristics of the new species fit
<taxonomicName id="4FA73887FF9B4A1DFE75FA9FBFB3FA91" authorityName="Hoffman" authorityYear="2005" box="[419,506,1371,1396]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoro" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="BAD39F16FF9B4A1DFE75FA9FBFB3FA91" box="[419,506,1371,1396]" italics="true" pageId="17" pageNumber="18">Pogoro</emphasis>
</taxonomicName>
. The only differences from the
<taxonomicName id="4FA73887FF9B4A1DFCA1FA9FBD87FA91" authorityName="Hoffman" authorityYear="2005" box="[887,974,1371,1396]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoro" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="BAD39F16FF9B4A1DFCA1FA9FBD87FA91" box="[887,974,1371,1396]" italics="true" pageId="17" pageNumber="18">Pogoro</emphasis>
</taxonomicName>
species description are the absence of paracannular setae, the absence of gonopodal process L and in particular the very elongated, hairpinbent gonopod telopodite.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="57BC0C002978F0D56DFDA0522F3DD439">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="1C04C7E2D16190F82F3735D8B22ADCE5">
<mods:date id="6AC088939913DD293B0AC072AA49B934">2020</mods:date>
<mods:detail id="ED59EABEE189BC513AC46BA66F02181D" type="pubDate">
<mods:number id="585A2E00AC7AE34AB58A1A80B314C769">2020-06-29</mods:number>
</mods:detail>
<mods:detail id="BDF9E06AF7A78D81A76E41DF621B38A7" type="volume">
<mods:number id="D73B93B1CCCAFFD63736EEC0C8A060D5">675</mods:number>
</mods:detail>
<mods:extent id="D8A50D578B63DDC62673FBFC838929FE" unit="page">
<mods:start id="3ED9893EBAA2892BA2E0C506D0CEA22D">1</mods:start>
<mods:end id="7E975476631ABD39531FF5A260B74F7A">35</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="AAFA3C3A8905556DD45470A5B4189509">journal article</mods:classification>
<mods:identifier id="F3FB0B6EE147B99A613DAA29694DE154" type="CLB-Dataset">21573</mods:identifier>
<mods:identifier id="56719713447FB85506A83BD196E07205" type="DOI">10.5852/ejt.2020.675</mods:identifier>
<mods:identifier id="F631A876B4B9B15E9C89EC087827D634" type="GBIF-Dataset">39cc6284-8ad7-47f0-bf5d-2122174905ed</mods:identifier>
<mods:identifier id="620E6A2203B0D1C7B601DB543D48FBEE" type="Zenodo-Dep">3928326</mods:identifier>
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<treatment id="000EF212FF864A02FDD9FD5FBDB6FB46" ID-DOI="http://doi.org/10.5281/zenodo.4332905" ID-GBIF-Taxon="164870668" ID-Zenodo-Dep="4332905" LSID="urn:lsid:zoobank.org:act:90399199-DF61-47B0-8D5E-B01AAAF16DE2" httpUri="http://treatment.plazi.org/id/000EF212FF864A02FDD9FD5FBDB6FB46" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
<subSubSection id="C0BD108FFF864A00FDD9FD5FBA6CFD50" box="[527,1061,666,693]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="88184304FF864A00FDD9FD5FBA6CFD50" blockId="12.[363,1224,666,763]" box="[527,1061,666,693]" pageId="12" pageNumber="13">
<heading id="D350F468FF864A00FDD9FD5FBA6CFD50" box="[527,1061,666,693]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="13" reason="2">
<taxonomicName id="4FA73887FF864A00FDD9FD5FBD8AFD51" ID-CoL="f82ea711-b87c-48b5-aed2-1654e66295d0" authority="Rosenmejer &amp; Enghoff" authorityName="Rosenmejer &amp; Enghoff" authorityYear="2020" box="[527,963,666,693]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoro" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="siren" status="sp. nov.">
<emphasis id="BAD39F16FF864A00FDD9FD5FBCE3FD51" bold="true" box="[527,682,666,693]" italics="true" pageId="12" pageNumber="13">Pogoro siren</emphasis>
Rosenmejer &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF864A00FC1CFD5FBA6CFD50" box="[970,1061,667,693]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF864A00FEBDFD79BDC8FD1E" pageId="12" pageNumber="13" type="description">
<paragraph id="88184304FF864A00FEBDFD79BA81FD3D" blockId="12.[363,1224,666,763]" box="[363,1224,701,728]" pageId="12" pageNumber="13">
<uri id="FC364F06FF864A00FEBDFD79BA81FD3D" box="[363,1224,701,728]" pageId="12" pageNumber="13">
urn:lsid:zoobank.org:act:
<uuid id="FC0179D1FF864A00FD5DFD79BA81FD3D" box="[651,1224,701,728]" pageId="12" pageNumber="13">90399199-DF61-47B0-8D5E-B01AAAF16DE2</uuid>
</uri>
</paragraph>
<paragraph id="88184304FF864A00FD64FD25BDC8FD1E" blockId="12.[363,1224,666,763]" box="[690,897,736,763]" pageId="12" pageNumber="13">
<figureCitation id="109C5F81FF864A00FD64FD25BCB4FD1E" box="[690,765,737,763]" captionStart="Fig" captionStartId="2.[189,232,1966,1992]" captionTargetBox="[199,1398,1100,1897]" captionTargetId="figure@2.[194,1482,1044,1958]" captionTargetPageId="2" captionText="Fig. 2. Map of the Udzungwa Mts showing collection sites for gomphodesmids. Based on Marshall et al. (2010: fig. 1)." figureDoi="http://doi.org/10.5281/zenodo.3928330" httpUri="https://zenodo.org/record/3928330/files/figure.png" pageId="12" pageNumber="13">
Figs
<geoCoordinate id="ED9325C3FF864A00FD3AFD25BCB4FD1E" box="[748,765,737,763]" degrees="2, 5" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="5555" value="2.5">2</geoCoordinate>
</figureCitation>
,
<figureCitation id="109C5F81FF864A00FCDCFD24BD64FD1E" box="[778,813,736,763]" captionStart="Fig" captionStartId="6.[189,232,1500,1526]" captionTargetBox="[217,1367,909,1445]" captionTargetId="figure@6.[207,1377,905,1453]" captionTargetPageId="6" captionText="Fig. 5. Sternal process on body ring 6. A. Emphysemastix spp. B. Agrophogonus spp. except A. hamulifer Olsen &amp; Enghoff sp. nov. C. Agrophogonus hamulifer Olsen &amp; Enghoff sp. nov. D. Pogoropsis prolixopes Rosenmejer &amp; Enghoff gen. et sp. nov. E. Pogoro siren Rosenmejer &amp; Enghoff sp. nov. F. Pogoro scharffi Hoffman, 2005. G. Pogoro alopias Rosenmejer &amp; Enghoff sp. nov. Not drawn to scale." figureDoi="http://doi.org/10.5281/zenodo.3928336" httpUri="https://zenodo.org/record/3928336/files/figure.png" pageId="12" pageNumber="13">5E</figureCitation>
,
<figureCitation id="109C5F81FF864A00FCEFFD25BDC8FD1E" box="[825,897,737,763]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="12.[189,232,2000,2026]" captionStartId-1="13.[189,232,1929,1955]" captionTargetBox-0="[372,1217,1203,1949]" captionTargetBox-1="[334,1230,265,1883]" captionTargetId-0="figure@12.[244,1341,1199,1954]" captionTargetId-1="figure@13.[334,1254,265,1885]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="Fig. 10. Pogoro siren Rosenmejer &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621674). Scale bar: 5 mm." captionText-1="Fig. 11. Pogoro siren Rosenmejer &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621674), right gonopod. A–B. Mesal view. C–D. Lateral view. E–F. Dorsal view. Abbreviations: L = Process L; M = Process M; sl = subapical lobe; ul = uncate lobe. Scale bars: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3928346" figureDoi-1="http://doi.org/10.5281/zenodo.3928348" httpUri-0="https://zenodo.org/record/3928346/files/figure.png" httpUri-1="https://zenodo.org/record/3928348/files/figure.png" pageId="12" pageNumber="13">10–11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF864A00FF6BFCE6BFB4FC6B" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="88184304FF864A00FF6BFCE6BF7FFCD9" blockId="12.[189,310,802,828]" box="[189,310,802,828]" pageId="12" pageNumber="13">
<heading id="D350F468FF864A00FF6BFCE6BF7FFCD9" bold="true" box="[189,310,802,828]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="BAD39F16FF864A00FF6BFCE6BF7FFCD9" bold="true" box="[189,310,802,828]" pageId="12" pageNumber="13">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF864A00FF6BFC95BFB4FC6B" blockId="12.[189,1398,849,910]" pageId="12" pageNumber="13">Differs from other species of the genus by the shape of the postnodal telopodite and by the non-enlarged tubercles on the hypoproct.</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF864A00FF6BFC72BD06FBC4" pageId="12" pageNumber="13" type="etymology">
<paragraph id="88184304FF864A00FF6BFC72BF0DFC35" blockId="12.[189,324,950,976]" box="[189,324,950,976]" pageId="12" pageNumber="13">
<heading id="D350F468FF864A00FF6BFC72BF0DFC35" bold="true" box="[189,324,950,976]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="BAD39F16FF864A00FF6BFC72BF0DFC35" bold="true" box="[189,324,950,976]" pageId="12" pageNumber="13">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF864A00FF6BFC20BD06FBC4" blockId="12.[189,1398,996,1058]" pageId="12" pageNumber="13">The name is a Latin noun in apposition, meaning mermaid, and refers to the mermaid tail-like shape of the postnodal telopodite when seen from the mesal side.</paragraph>
</subSubSection>
<caption id="DCD8138CFF864A00FF6BF814BB3AF80E" ID-DOI="http://doi.org/10.5281/zenodo.3928346" ID-Zenodo-Dep="3928346" box="[189,1395,2000,2027]" httpUri="https://zenodo.org/record/3928346/files/figure.png" pageId="12" pageNumber="13" startId="12.[189,232,2000,2026]" targetBox="[372,1217,1203,1949]" targetPageId="12">
<paragraph id="88184304FF864A00FF6BF814BB3AF80E" blockId="12.[189,1395,2000,2027]" box="[189,1395,2000,2027]" pageId="12" pageNumber="13">
<emphasis id="BAD39F16FF864A00FF6BF814BF50F80F" bold="true" box="[189,281,2000,2026]" pageId="12" pageNumber="13">Fig. 10.</emphasis>
<taxonomicName id="4FA73887FF864A00FEF6F815BC98F80F" authority="Rosenmejer &amp; Enghoff" authorityName="Rosenmejer &amp; Enghoff" authorityYear="2020" box="[288,721,2000,2027]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoro" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="siren" status="sp. nov.">
<emphasis id="BAD39F16FF864A00FEF6F815BFF0F80F" box="[288,441,2001,2026]" italics="true" pageId="12" pageNumber="13">Pogoro siren</emphasis>
Rosenmejer &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF864A00FD0EF815BD7CF80E" box="[728,821,2001,2027]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NHMD 621674). Scale bar: 5 mm.
</paragraph>
</caption>
<caption id="DCD8138CFF874A01FF6BF84DBD15F80F" ID-DOI="http://doi.org/10.5281/zenodo.3928348" ID-Zenodo-Dep="3928348" httpUri="https://zenodo.org/record/3928348/files/figure.png" pageId="13" pageNumber="14" startId="13.[189,232,1929,1955]" targetBox="[334,1230,265,1883]" targetPageId="13">
<paragraph id="88184304FF874A01FF6BF84DBD15F80F" blockId="13.[189,1399,1929,2026]" pageId="13" pageNumber="14">
<emphasis id="BAD39F16FF874A01FF6BF84DBF53F846" bold="true" box="[189,282,1929,1955]" pageId="13" pageNumber="14">Fig. 11.</emphasis>
<taxonomicName id="4FA73887FF874A01FEF5F84EBC94F846" authority="Rosenmejer &amp; Enghoff" authorityName="Rosenmejer &amp; Enghoff" authorityYear="2020" box="[291,733,1929,1955]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoro" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="siren" status="sp. nov.">
<emphasis id="BAD39F16FF874A01FEF5F84EBFF7F846" box="[291,446,1930,1955]" italics="true" pageId="13" pageNumber="14">Pogoro siren</emphasis>
Rosenmejer &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF874A01FD31F84DBD0FF846" box="[743,838,1929,1955]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NHMD 621674), right gonopod.
<emphasis id="BAD39F16FF874A01FF6BF868BEBDF823" bold="true" box="[189,244,1964,1990]" pageId="13" pageNumber="14">A–B</emphasis>
. Mesal view.
<emphasis id="BAD39F16FF874A01FE40F868BF86F823" bold="true" box="[406,463,1964,1990]" pageId="13" pageNumber="14">C–D</emphasis>
. Lateral view.
<emphasis id="BAD39F16FF874A01FDABF868BCF8F823" bold="true" box="[637,689,1964,1990]" pageId="13" pageNumber="14">E–F</emphasis>
. Dorsal view. Abbreviations:
<emphasis id="BAD39F16FF874A01FBDBF869BA54F823" box="[1037,1053,1965,1990]" italics="true" pageId="13" pageNumber="14">L</emphasis>
= Process L;
<emphasis id="BAD39F16FF874A01FB6BF869BA9CF823" box="[1213,1237,1965,1990]" italics="true" pageId="13" pageNumber="14">M</emphasis>
= Process M;
<emphasis id="BAD39F16FF874A01FF6BF80BBE98F80C" box="[189,209,1999,2025]" italics="true" pageId="13" pageNumber="14">sl</emphasis>
= subapical lobe;
<emphasis id="BAD39F16FF874A01FE7EF80BBFF6F80C" box="[424,447,1999,2025]" italics="true" pageId="13" pageNumber="14">ul</emphasis>
= uncate lobe. Scale bars: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C0BD108FFF844A02FF6BFECFBF37FE54" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="88184304FF844A02FF6BFECFBFE3FEC0" blockId="14.[189,1399,267,434]" box="[189,426,267,293]" pageId="14" pageNumber="15">
<heading id="D350F468FF844A02FF6BFECFBFE3FEC0" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="BAD39F16FF844A02FF6BFECFBFE3FEC0" bold="true" box="[189,426,267,293]" pageId="14" pageNumber="15">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF844A02FF3CFEF3BF15FEB4" blockId="14.[189,1399,267,434]" box="[234,348,311,337]" pageId="14" pageNumber="15">
<emphasis id="BAD39F16FF844A02FF3CFEF3BF15FEB4" bold="true" box="[234,348,311,337]" pageId="14" pageNumber="15">
<typeStatus id="571CFDA6FF844A02FF3CFEF3BF15FEB4" box="[234,348,311,337]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="88184304FF844A02FF6BFE93BF37FE54" blockId="14.[189,1399,267,434]" pageId="14" pageNumber="15">
<materialsCitation id="38CF4959FF844A02FF6BFE93BF37FE54" ID-GBIF-Occurrence="2803530319" collectingDate="2014-11-02" collectingMethod="pitfall traps" collectionCode="NHMD" collectorName="J. Malumbres-Olarte" country="Tanzania" elevation="1531" latitude="-8.48875" location="Chita" longLatPrecision="1" longitude="35.907585" municipality="Udzungwa Scarp Catchment Forest Reserve" pageId="14" pageNumber="15" specimenCode="NHMD 621674" specimenCount="1" specimenCount-male="1" stateProvince="Morogoro Region" typeStatus="holotype">
<collectingCountry id="F0B00394FF844A02FF6BFE93BF18FE94" box="[189,337,343,369]" name="Tanzania" pageId="14" pageNumber="15">TANZANIA</collectingCountry>
•
<specimenCount id="9EA1888DFF844A02FEA0FE93BFC7FE94" box="[374,398,343,369]" pageId="14" pageNumber="15" type="male">♂</specimenCount>
;
<collectingRegion id="4A638DE6FF844A02FE77FE93BC34FE94" box="[417,637,343,369]" country="Tanzania" name="Morogoro" pageId="14" pageNumber="15">Morogoro Region</collectingRegion>
,
<collectingMunicipality id="687CD97EFF844A02FD59FE93BAF9FE94" box="[655,1200,343,369]" pageId="14" pageNumber="15">Udzungwa Scarp Catchment Forest Reserve</collectingMunicipality>
,
<location id="8D7815DFFF844A02FB14FE93BB4CFE94" LSID="urn:lsid:plazi:treatment:000EF212FF864A02FDD9FD5FBDB6FB46:8D7815DFFF844A02FB14FE93BB4CFE94" box="[1218,1285,343,369]" country="Tanzania" latitude="-8.48875" longLatPrecision="1" longitude="35.907585" municipality="Udzungwa Scarp Catchment Forest Reserve" name="Chita" pageId="14" pageNumber="15" stateProvince="Morogoro Region">Chita</location>
, plot 18;
<geoCoordinate id="ED9325C3FF844A02FF6BFEB1BF21FE74" box="[189,360,373,401]" degrees="08" direction="south" minutes="29" orientation="latitude" pageId="14" pageNumber="15" precision="1" seconds="19.5" value="-8.48875">08°29′19.5″ S</geoCoordinate>
,
<geoCoordinate id="ED9325C3FF844A02FEACFEB1BC6EFE77" box="[378,551,373,402]" degrees="35" direction="east" minutes="54" orientation="longitude" pageId="14" pageNumber="15" precision="1" seconds="27.3" value="35.907585">35°54′27.3″ E</geoCoordinate>
,
<elevation id="038AA437FF844A02FDEFFEB3BC95FE74" box="[569,732,375,402]" metricMagnitude="3" metricUnit="m" metricValue="1.531" pageId="14" pageNumber="15" unit="m" value="1531.0">
<quantity id="4F5FEEE1FF844A02FDEFFEB3BCDEFE77" box="[569,663,375,402]" metricMagnitude="3" metricUnit="m" metricValue="1.531" pageId="14" pageNumber="15" unit="m" value="1531.0">1531 m</quantity>
a.s.l.
</elevation>
;
<date id="FC1965C4FF844A02FD39FEB3BDC4FE74" box="[751,909,375,402]" pageId="14" pageNumber="15" value="2014-11-02">
<collectingDate id="EC5D9C2CFF844A02FD39FEB3BDC4FE74" box="[751,909,375,402]" pageId="14" pageNumber="15" value="2014-11-02">2 Nov. 2014</collectingDate>
</date>
;
<collectorName id="255226D2FF844A02FC48FEBCBADFFE74" box="[926,1174,375,402]" pageId="14" pageNumber="15">J. Malumbres-Olarte</collectorName>
leg.;
<collectingMethod id="51E63B13FF844A02FB35FEB3BB3BFE77" box="[1251,1394,375,402]" pageId="14" pageNumber="15">pitfall traps</collectingMethod>
;
<specimenCode id="D801EB7FFF844A02FF6BFE5CBF33FE54" box="[189,378,407,434]" collectionCode="NHMD" pageId="14" pageNumber="15">NHMD 621674</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF844A02FF6BFE12BEBEFBDD" pageId="14" pageNumber="15" type="description">
<paragraph id="88184304FF844A02FF6BFE12BF07FE15" blockId="14.[189,1398,470,605]" box="[189,334,470,496]" pageId="14" pageNumber="15">
<heading id="D350F468FF844A02FF6BFE12BF07FE15" bold="true" box="[189,334,470,496]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="BAD39F16FF844A02FF6BFE12BF07FE15" bold="true" box="[189,334,470,496]" pageId="14" pageNumber="15">Description</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF844A02FF3CFDC6BC70FDB9" blockId="14.[189,1398,470,605]" pageId="14" pageNumber="15">
<emphasis id="BAD39F16FF844A02FF3CFDC6BF63FDF9" bold="true" box="[234,298,514,540]" pageId="14" pageNumber="15">Male</emphasis>
SIZE AND SHAPE. Body length ca
<quantity id="4F5FEEE1FF844A02FDE8FDE6BCDAFDD9" box="[574,659,546,573]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="14" pageNumber="15" unit="mm" value="55.0">55 mm</quantity>
. Maximum width
<quantity id="4F5FEEE1FF844A02FCBCFDE6BD8EFDD9" box="[874,967,546,572]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="14" pageNumber="15" unit="mm" value="7.8">7.8 mm</quantity>
. W/L ratio 13%. Height of body ring eight
<quantity id="4F5FEEE1FF844A02FF29FD86BF15FDB8" box="[255,348,578,605]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="14" pageNumber="15" unit="mm" value="6.5">6.5 mm</quantity>
. H/W ratio = 83%.
</paragraph>
<paragraph id="88184304FF844A02FF6BFD45BD42FD5E" blockId="14.[189,1398,641,700]" pageId="14" pageNumber="15">
COLOUR (
<figureCitation id="109C5F81FF844A02FEF8FD45BFCCFD7E" box="[302,389,641,667]" captionStart="Fig" captionStartId="12.[189,232,2000,2026]" captionTargetBox="[372,1217,1203,1949]" captionTargetId="figure@12.[244,1341,1199,1954]" captionTargetPageId="12" captionText="Fig. 10. Pogoro siren Rosenmejer &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621674). Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928346" httpUri="https://zenodo.org/record/3928346/files/figure.png" pageId="14" pageNumber="15">Fig. 10</figureCitation>
). Collum and dorsal surface of metaterga dark brown. Face, legs and antennae medium brown. Prozona and ventral surface lighter brown.
</paragraph>
<paragraph id="88184304FF844A02FF6BFD24BEB6FCFF" blockId="14.[189,1398,735,794]" pageId="14" pageNumber="15">
STERNAL PROCESS ON BODY RING 6 (
<figureCitation id="109C5F81FF844A02FD89FD24BCF3FD1F" box="[607,698,735,762]" captionStart="Fig" captionStartId="6.[189,232,1500,1526]" captionTargetBox="[217,1367,909,1445]" captionTargetId="figure@6.[207,1377,905,1453]" captionTargetPageId="6" captionText="Fig. 5. Sternal process on body ring 6. A. Emphysemastix spp. B. Agrophogonus spp. except A. hamulifer Olsen &amp; Enghoff sp. nov. C. Agrophogonus hamulifer Olsen &amp; Enghoff sp. nov. D. Pogoropsis prolixopes Rosenmejer &amp; Enghoff gen. et sp. nov. E. Pogoro siren Rosenmejer &amp; Enghoff sp. nov. F. Pogoro scharffi Hoffman, 2005. G. Pogoro alopias Rosenmejer &amp; Enghoff sp. nov. Not drawn to scale." figureDoi="http://doi.org/10.5281/zenodo.3928336" httpUri="https://zenodo.org/record/3928336/files/figure.png" pageId="14" pageNumber="15">Fig. 5E</figureCitation>
). Transverse across sternum with dark brown edge and long setae.
</paragraph>
<paragraph id="88184304FF844A02FF6BFCFBBF9FFC9C" blockId="14.[189,1398,830,889]" pageId="14" pageNumber="15">HYPOPROCT. Almost semicircular, without median projection, paramedian tubercles not projecting beyond edge of sclerite.</paragraph>
<paragraph id="88184304FF844A02FF6BFC59BEBEFBDD" blockId="14.[189,1399,925,1080]" pageId="14" pageNumber="15">
GONOPOD (
<figureCitation id="109C5F81FF844A02FE91FC59BFE8FC52" box="[327,417,925,951]" captionStart="Fig" captionStartId="13.[189,232,1929,1955]" captionTargetBox="[334,1230,265,1883]" captionTargetId="figure@13.[334,1254,265,1885]" captionTargetPageId="13" captionText="Fig. 11. Pogoro siren Rosenmejer &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621674), right gonopod. A–B. Mesal view. C–D. Lateral view. E–F. Dorsal view. Abbreviations: L = Process L; M = Process M; sl = subapical lobe; ul = uncate lobe. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928348" httpUri="https://zenodo.org/record/3928348/files/figure.png" pageId="14" pageNumber="15">Fig. 11</figureCitation>
). Gonocoxa with paracannular setae. Telopodite mesonodal. Process M twice as wide at base and slightly longer than process L; both curved laterad. Postnodal teleopodite relatively short, curved dorsomesad, with curved narrow subapical lobe with rounded apex on lateral side. Uncate subapical lobe on mesal side. Solenomere of telopodite curved inwards against apex of uncate subapical lobe.
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF844A02FF6BFB99BDB6FB46" box="[189,1023,1117,1187]" pageId="14" pageNumber="15" type="distribution">
<paragraph id="88184304FF844A02FF6BFB99BF1FFB92" blockId="14.[189,1023,1117,1187]" box="[189,342,1117,1143]" pageId="14" pageNumber="15">
<heading id="D350F468FF844A02FF6BFB99BF1FFB92" bold="true" box="[189,342,1117,1143]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="BAD39F16FF844A02FF6BFB99BF1FFB92" bold="true" box="[189,342,1117,1143]" pageId="14" pageNumber="15">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF844A02FF6BFB4DBDB6FB46" blockId="14.[189,1023,1117,1187]" box="[189,1023,1160,1187]" pageId="14" pageNumber="15">
Known only from the
<typeStatus id="571CFDA6FF844A02FE14FB4DBFBCFB46" box="[450,501,1161,1187]" pageId="14" pageNumber="15">type</typeStatus>
locality in the Udzungwa Scarp FR (
<figureCitation id="109C5F81FF844A02FC7FFB4CBDB8FB46" box="[937,1009,1160,1187]" captionStart="Fig" captionStartId="2.[189,232,1966,1992]" captionTargetBox="[199,1398,1100,1897]" captionTargetId="figure@2.[194,1482,1044,1958]" captionTargetPageId="2" captionText="Fig. 2. Map of the Udzungwa Mts showing collection sites for gomphodesmids. Based on Marshall et al. (2010: fig. 1)." figureDoi="http://doi.org/10.5281/zenodo.3928330" httpUri="https://zenodo.org/record/3928330/files/figure.png" pageId="14" pageNumber="15">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

215
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<paragraph id="88184304FF8D4A0BFD89F8DABD9CF8DC" blockId="7.[607,981,1821,1883]" box="[607,981,1821,1849]" pageId="7" pageNumber="8">
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<taxonomicName id="4FA73887FF8D4A0BFD89F8DABD9CF8DC" ID-CoL="4KWC7" authority="Hoffman, 2005" authorityName="Hoffman" authorityYear="2005" box="[607,981,1821,1849]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoro" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="scharffi">
<emphasis id="BAD39F16FF8D4A0BFD89F8DABD52F8DD" bold="true" box="[607,795,1821,1848]" italics="true" pageId="7" pageNumber="8">Pogoro scharffi</emphasis>
<bibRefCitation id="EC363EF5FF8D4A0BFCF4F8DABD9CF8DC" author="Hoffman R. L." box="[802,981,1822,1849]" pageId="7" pageNumber="8" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman, 2005</bibRefCitation>
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<paragraph id="88184304FF8D4A0BFD14F884BD38F8BF" blockId="7.[607,981,1821,1883]" box="[706,881,1856,1883]" pageId="7" pageNumber="8">
<figureCitation id="109C5F81FF8D4A0BFD14F884BD44F8BF" box="[706,781,1856,1882]" captionStart="Fig" captionStartId="2.[189,232,1966,1992]" captionTargetBox="[199,1398,1100,1897]" captionTargetId="figure@2.[194,1482,1044,1958]" captionTargetPageId="2" captionText="Fig. 2. Map of the Udzungwa Mts showing collection sites for gomphodesmids. Based on Marshall et al. (2010: fig. 1)." figureDoi="http://doi.org/10.5281/zenodo.3928330" httpUri="https://zenodo.org/record/3928330/files/figure.png" pageId="7" pageNumber="8">Figs 2</figureCitation>
,
<figureCitation id="109C5F81FF8D4A0BFCCCF884BD70F8BE" box="[794,825,1856,1883]" captionStart="Fig" captionStartId="6.[189,232,1500,1526]" captionTargetBox="[217,1367,909,1445]" captionTargetId="figure@6.[207,1377,905,1453]" captionTargetPageId="6" captionText="Fig. 5. Sternal process on body ring 6. A. Emphysemastix spp. B. Agrophogonus spp. except A. hamulifer Olsen &amp; Enghoff sp. nov. C. Agrophogonus hamulifer Olsen &amp; Enghoff sp. nov. D. Pogoropsis prolixopes Rosenmejer &amp; Enghoff gen. et sp. nov. E. Pogoro siren Rosenmejer &amp; Enghoff sp. nov. F. Pogoro scharffi Hoffman, 2005. G. Pogoro alopias Rosenmejer &amp; Enghoff sp. nov. Not drawn to scale." figureDoi="http://doi.org/10.5281/zenodo.3928336" httpUri="https://zenodo.org/record/3928336/files/figure.png" pageId="7" pageNumber="8">5F</figureCitation>
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<figureCitation id="109C5F81FF8D4A0BFC93F884BD38F8BF" box="[837,881,1856,1882]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="8.[189,232,2000,2026]" captionStartId-1="9.[189,232,1929,1955]" captionTargetBox-0="[399,1200,1299,1965]" captionTargetBox-1="[297,1266,266,1882]" captionTargetId-0="figure@8.[387,1209,1286,1973]" captionTargetId-1="figure@9.[297,1280,265,1882]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Fig. 6. Pogoro scharffi Hoffman, 2005, paratype, ♂ (NHMD 621669). Scale bar: 5 mm." captionText-1="Fig. 7. Pogoro scharffi Hoffman, 2005, paratype, ♂, left gonopod (NHMD 621669). A–B. Mesal view. C–D. Lateral view. E–F. Dorsal view. Abbreviations: L = Process L; M = Process M; tl = triangular lobe; ul = uncate lobe. Scale bars: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3928338" figureDoi-1="http://doi.org/10.5281/zenodo.3928340" httpUri-0="https://zenodo.org/record/3928338/files/figure.png" httpUri-1="https://zenodo.org/record/3928340/files/figure.png" pageId="7" pageNumber="8">6–7</figureCitation>
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<heading id="D350F468FF8D4A0BFF6BF845BF7FF87E" bold="true" box="[189,310,1921,1947]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="BAD39F16FF8D4A0BFF6BF845BF7FF87E" bold="true" box="[189,310,1921,1947]" pageId="7" pageNumber="8">Diagnosis</emphasis>
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</paragraph>
<paragraph id="88184304FF8D4A0BFF6BF86ABD43F80E" blockId="7.[189,1398,1966,2027]" pageId="7" pageNumber="8">Differs from other species of the genus by the shape of the subapical lobes on the postnodal telepodite, and by the shape and curvature of the solenomere.</paragraph>
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<heading id="D350F468FF824A04FF6BFECFBFE3FEC0" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="BAD39F16FF824A04FF6BFECFBFE3FEC0" bold="true" box="[189,426,267,293]" pageId="8" pageNumber="9">Material examined</emphasis>
</heading>
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</emphasis>
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<paragraph id="88184304FF824A04FF6BFE9DBB5BFE70" blockId="8.[189,1399,312,405]" pageId="8" pageNumber="9">
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<collectingCountry id="F0B00394FF824A04FF6BFE9DBF18FE96" box="[189,337,345,371]" name="Tanzania" pageId="8" pageNumber="9">TANZANIA</collectingCountry>
•
<specimenCount id="9EA1888DFF824A04FEB8FE9CBFCDFE91" box="[366,388,344,372]" pageId="8" pageNumber="9" type="male">♂</specimenCount>
;
<collectingRegion id="4A638DE6FF824A04FE40FE9DBC77FE96" box="[406,574,345,371]" country="Tanzania" name="Iringa" pageId="8" pageNumber="9">Iringa Region</collectingRegion>
,
<collectingMunicipality id="687CD97EFF824A04FD9DFE9DBD4DFE96" box="[587,772,345,371]" pageId="8" pageNumber="9">Udzungwa Mts</collectingMunicipality>
,
<collectorName id="255226D2FF824A04FCC5FE9DBA46FE96" box="[787,1039,345,371]" pageId="8" pageNumber="9">Udzungwa Scarp FR</collectorName>
,
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;
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a.s.l.
</elevation>
;
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;
<collectorName id="255226D2FF824A04FE01FEBEBC18FE71" box="[471,593,378,404]" pageId="8" pageNumber="9">N. Scharff</collectorName>
leg.;
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in
<collectedFrom id="3BBB00F9FF824A04FCE8FEBEBA1DFE71" box="[830,1108,378,404]" pageId="8" pageNumber="9">intermediate rain forest</collectedFrom>
;
<specimenCode id="D801EB7FFF824A04FBB6FEBEBB47FE70" box="[1120,1294,378,405]" collectionCode="NHMD" pageId="8" pageNumber="9">NHMD 50501</specimenCode>
.
</materialsCitation>
</paragraph>
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<typeStatus id="571CFDA6FF824A04FF3CFE7EBF2EFE31" box="[234,359,442,468]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
</emphasis>
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<collectingCountry id="F0B00394FF824A04FF6BFE1FBF18FE10" box="[189,337,475,501]" name="Tanzania" pageId="8" pageNumber="9">TANZANIA</collectingCountry>
•
<specimenCount id="9EA1888DFF824A04FEBCFE1FBFE7FE10" box="[362,430,475,501]" pageId="8" pageNumber="9" type="male">9 ♂♂</specimenCount>
,
<specimenCount id="9EA1888DFF824A04FE6DFE1FBFB7FE10" box="[443,510,475,501]" pageId="8" pageNumber="9" type="female">2 ♀♀</specimenCount>
;
<collectingRegion id="4A638DE6FF824A04FDDBFE1EBCFAFE11" box="[525,691,474,500]" country="Tanzania" name="Iringa" pageId="8" pageNumber="9">Iringa Region</collectingRegion>
,
<collectingMunicipality id="687CD97EFF824A04FD68FE1EBD3CFE10" box="[702,885,474,501]" pageId="8" pageNumber="9">Udzungwa Mts</collectingMunicipality>
,
<collectorName id="255226D2FF824A04FC54FE1EBA33FE10" box="[898,1146,474,501]" pageId="8" pageNumber="9">Udzungwa Scarp FR</collectorName>
,
<locationDeviation id="99AA5A26FF824A04FB53FE1EBB3BFE11" box="[1157,1394,474,500]" pageId="8" pageNumber="9">above Chita Village</locationDeviation>
;
<elevation id="038AA437FF824A04FF6BFE38BFE0FDF3" box="[189,425,508,535]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.65" metricValueMin="1.05" pageId="8" pageNumber="9" unit="m" value="1350.0" valueMax="1650.0" valueMin="1050.0">
<quantity id="4F5FEEE1FF824A04FF6BFE38BF2DFDF3" box="[189,356,508,535]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.65" metricValueMin="1.05" pageId="8" pageNumber="9" unit="m" value="1350.0" valueMax="1650.0" valueMin="1050.0">1050–1650 m</quantity>
a.s.l.
</elevation>
;
<date id="FC1965C4FF824A04FE6DFE38BC99FDF3" box="[443,720,508,534]" pageId="8" pageNumber="9" value="1984-10-26" valueMax="1984-11-13" valueMin="1984-10-26">
<collectingDate id="EC5D9C2CFF824A04FE6DFE38BC99FDF3" box="[443,720,508,534]" pageId="8" pageNumber="9" value="1984-10-26" valueMax="1984-11-13" valueMin="1984-10-26">26 Oct.–13 Nov. 1984</collectingDate>
</date>
;
<collectorName id="255226D2FF824A04FD37FE38BD29FDF3" box="[737,864,508,534]" pageId="8" pageNumber="9">N. Scharff</collectorName>
leg.;
<collectedFrom id="3BBB00F9FF824A04FC7DFE38BB3AFDF3" box="[939,1395,508,534]" pageId="8" pageNumber="9">intermediate and montane rain forest</collectedFrom>
;
<specimenCode id="D801EB7FFF824A04FF6BFDDABF33FDD2" box="[189,378,541,568]" collectionCode="NHMD" pageId="8" pageNumber="9">NHMD 621669</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF824A04FF6BFD98BD98FB65" pageId="8" pageNumber="9" type="description">
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<emphasis id="BAD39F16FF824A04FF6BFD98BF07FD93" bold="true" box="[189,334,604,630]" pageId="8" pageNumber="9">Description</emphasis>
(modified after
<bibRefCitation id="EC363EF5FF824A04FDDAFD98BCF1FD92" author="Hoffman R. L." box="[524,696,604,631]" pageId="8" pageNumber="9" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman 2005</bibRefCitation>
)
<emphasis id="BAD39F16FF824A04FF3CFD4DBF63FD46" bold="true" box="[234,298,649,675]" pageId="8" pageNumber="9">Male</emphasis>
</paragraph>
<paragraph id="88184304FF824A04FF6BFD6EBC6AFD03" blockId="8.[189,1398,651,742]" pageId="8" pageNumber="9">
SIZE AND SHAPE. Body length ca
<quantity id="4F5FEEE1FF824A04FDE8FD6EBCDAFD20" box="[574,659,682,709]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="9" unit="mm" value="40.0">40 mm</quantity>
. Maximum width
<quantity id="4F5FEEE1FF824A04FCBCFD6FBD8EFD20" box="[874,967,683,709]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="8" pageNumber="9" unit="mm" value="7.7">7.7 mm</quantity>
. W/L ratio 19%. Height of body ring eight
<quantity id="4F5FEEE1FF824A04FF29FD0FBF0FFD03" box="[255,326,715,742]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="9" unit="mm" value="5.0">5 mm</quantity>
. H/W ratio = 65%.
</paragraph>
<paragraph id="88184304FF824A04FF6BFCCDBFC5FCA3" blockId="8.[189,1398,777,838]" pageId="8" pageNumber="9">
COLOUR (
<figureCitation id="109C5F81FF824A04FEE6FCCDBF3FFCC1" box="[304,374,777,804]" captionStart="Fig" captionStartId="8.[189,232,2000,2026]" captionTargetBox="[399,1200,1299,1965]" captionTargetId="figure@8.[387,1209,1286,1973]" captionTargetPageId="8" captionText="Fig. 6. Pogoro scharffi Hoffman, 2005, paratype, ♂ (NHMD 621669). Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928338" httpUri="https://zenodo.org/record/3928338/files/figure.png" pageId="8" pageNumber="9">Fig. 6</figureCitation>
). Face, collum, metaterga medium brown, legs and antennae lighter, prozona and sterna nearly colourless.
</paragraph>
<paragraph id="88184304FF824A04FF6BFCACBB1DFC67" blockId="8.[189,1364,872,898]" box="[189,1364,872,898]" pageId="8" pageNumber="9">STERNAL PROCESS ON BODY RING 6. Transverse across sternum with dark brown edge and long setae.</paragraph>
<paragraph id="88184304FF824A04FF6BFC60BCA2FC3A" blockId="8.[189,1398,931,991]" pageId="8" pageNumber="9">HYPOPROCT. Almost semicircular, with moderate round median projection, paramedian tubercles above normal size, projecting beyond edge of sclerite.</paragraph>
<paragraph id="88184304FF824A04FF6BFBC5BD98FB65" blockId="8.[189,1398,1024,1153]" pageId="8" pageNumber="9">
GONOPOD (
<figureCitation id="109C5F81FF824A04FE92FBC4BFC2FBFE" box="[324,395,1024,1051]" captionStart="Fig" captionStartId="9.[189,232,1929,1955]" captionTargetBox="[297,1266,266,1882]" captionTargetId="figure@9.[297,1280,265,1882]" captionTargetPageId="9" captionText="Fig. 7. Pogoro scharffi Hoffman, 2005, paratype, ♂, left gonopod (NHMD 621669). A–B. Mesal view. C–D. Lateral view. E–F. Dorsal view. Abbreviations: L = Process L; M = Process M; tl = triangular lobe; ul = uncate lobe. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928340" httpUri="https://zenodo.org/record/3928340/files/figure.png" pageId="8" pageNumber="9">Fig. 7</figureCitation>
). Gonocoxa with paracannular setae. Telopodite mesonodal. Process M twice as wide at base and slightly longer than process L, both curved laterad. Postnodal teleopodite relatively short, curved dorsomesad, with subapical triangular lobe on lateral side and uncate lobe on mesal. Solenomere of telopodite curved away from uncate subapical lobe (
<figureCitation id="109C5F81FF824A04FC95FBA2BDC1FB65" box="[835,904,1126,1152]" captionStart="Fig" captionStartId="9.[189,232,1929,1955]" captionTargetBox="[297,1266,266,1882]" captionTargetId="figure@9.[297,1280,265,1882]" captionTargetPageId="9" captionText="Fig. 7. Pogoro scharffi Hoffman, 2005, paratype, ♂, left gonopod (NHMD 621669). A–B. Mesal view. C–D. Lateral view. E–F. Dorsal view. Abbreviations: L = Process L; M = Process M; tl = triangular lobe; ul = uncate lobe. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928340" httpUri="https://zenodo.org/record/3928340/files/figure.png" pageId="8" pageNumber="9">Fig. 7</figureCitation>
A–B).
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF824A04FF6BFB60BA61FB09" box="[189,1064,1188,1261]" pageId="8" pageNumber="9" type="distribution">
<paragraph id="88184304FF824A04FF6BFB60BF1FFB5B" blockId="8.[189,342,1188,1214]" box="[189,342,1188,1214]" pageId="8" pageNumber="9">
<heading id="D350F468FF824A04FF6BFB60BF1FFB5B" bold="true" box="[189,342,1188,1214]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="BAD39F16FF824A04FF6BFB60BF1FFB5B" bold="true" box="[189,342,1188,1214]" pageId="8" pageNumber="9">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF824A04FF6BFB17BA61FB09" blockId="8.[189,1064,1234,1261]" box="[189,1064,1234,1261]" pageId="8" pageNumber="9">
Known only from two very close sites in the Udzungwa Scarp FR (
<figureCitation id="109C5F81FF824A04FC04FB16BA53FB09" box="[978,1050,1234,1260]" captionStart="Fig" captionStartId="2.[189,232,1966,1992]" captionTargetBox="[199,1398,1100,1897]" captionTargetId="figure@2.[194,1482,1044,1958]" captionTargetPageId="2" captionText="Fig. 2. Map of the Udzungwa Mts showing collection sites for gomphodesmids. Based on Marshall et al. (2010: fig. 1)." figureDoi="http://doi.org/10.5281/zenodo.3928330" httpUri="https://zenodo.org/record/3928330/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

175
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<paragraph id="88184304FF8D4A0BFDBEFDE6BD85FDD8" blockId="7.[616,972,546,573]" box="[616,972,546,573]" pageId="7" pageNumber="8">
<heading id="D350F468FF8D4A0BFDBEFDE6BD85FDD8" box="[616,972,546,573]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="8" reason="2">
Genus
<taxonomicName id="4FA73887FF8D4A0BFD6CFDE6BD85FDD8" ID-CoL="7PV5L" authority="Hoffman, 2005" authorityName="Hoffman" authorityYear="2005" box="[698,972,546,573]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoro" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="BAD39F16FF8D4A0BFD6CFDE6BD5BFDD9" bold="true" box="[698,786,546,572]" italics="true" pageId="7" pageNumber="8">Pogoro</emphasis>
<bibRefCitation id="EC363EF5FF8D4A0BFCCFFDE6BD85FDD8" author="Hoffman R. L." box="[793,972,546,573]" pageId="7" pageNumber="8" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman, 2005</bibRefCitation>
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<emphasis id="BAD39F16FF8D4A0BFF6BFDA7BF7FFD98" bold="true" box="[189,310,611,637]" pageId="7" pageNumber="8">Diagnosis</emphasis>
(after
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, modified)
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Moderate/small-sized (length
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), convex gomphodesmids, pore formula: 5, 7, 9–19. Four apical sensory cones on antennae, fossa present on 5
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and 6
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antennomeres. Collum transversely suboval, lateral ends strongly declivent. Paranota slightly declivent. Stricture poorly defined, with visible edge only in front of anterior spiracle. Pleurosternal carinae present on body rings 1–17. Anterior legs without ventral tubercles, with sparse setation. Setae on dorsal side of tarsi shorter and stouter than the rest. Subonychial pads on legs 1–6. Surface of sides of metazona slightly more uneven than sides of prozona. Postgonopodal sterna without transverse ridges, set with long silky setae. 15
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body ring without paxillus. Caudal edge of gonopod aperture elevated in front of coxae, medially obliterated.
</paragraph>
<paragraph id="88184304FF8D4A0BFF6BFC05BF31FBA6" blockId="7.[189,1399,961,1091]" pageId="7" pageNumber="8">
<bibRefCitation id="EC363EF5FF8D4A0BFF6BFC05BF33FC39" author="Hoffman R. L." box="[189,378,961,988]" pageId="7" pageNumber="8" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman (2005)</bibRefCitation>
described the sides of the metazona as being “coarsely rugovermiculose”; this character was, however, not observed on any of the type specimens of
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<emphasis id="BAD39F16FF8D4A0BFC58FC20BA4AFC1B" box="[910,1027,996,1022]" italics="true" pageId="7" pageNumber="8">P. scharffi</emphasis>
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. Hoffman also stated that there are no paracannular setae in
<taxonomicName id="4FA73887FF8D4A0BFDDFFBC3BCF6FBC5" authorityName="Hoffman" authorityYear="2005" box="[521,703,1030,1056]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoro" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="scharffi">
<emphasis id="BAD39F16FF8D4A0BFDDFFBC3BCF6FBC5" box="[521,703,1030,1056]" italics="true" pageId="7" pageNumber="8">Pogoro scharffi</emphasis>
</taxonomicName>
, but some such setae were observed on the
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and all of the
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</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF8D4A0BFF6BFBADBC86FB13" pageId="7" pageNumber="8" type="reference_group">
<paragraph id="88184304FF8D4A0BFF6BFBADBFC0FB66" blockId="7.[189,393,1129,1155]" box="[189,393,1129,1155]" pageId="7" pageNumber="8">
<heading id="D350F468FF8D4A0BFF6BFBADBFC0FB66" bold="true" box="[189,393,1129,1155]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="BAD39F16FF8D4A0BFF6BFBADBFC0FB66" bold="true" box="[189,393,1129,1155]" pageId="7" pageNumber="8">Included species</emphasis>
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<taxonomicName id="4FA73887FF8D4A0BFF6BFB53BC67FB54" ID-CoL="4KWC7" authority="Hoffman, 2005" authorityName="Hoffman" authorityYear="2005" box="[189,558,1174,1201]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoro" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="scharffi">
<emphasis id="BAD39F16FF8D4A0BFF6BFB53BF3AFB54" box="[189,371,1175,1201]" italics="true" pageId="7" pageNumber="8">Pogoro scharffi</emphasis>
<bibRefCitation id="EC363EF5FF8D4A0BFEADFB53BC67FB54" author="Hoffman R. L." box="[379,558,1174,1201]" pageId="7" pageNumber="8" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman, 2005</bibRefCitation>
</taxonomicName>
(
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species)
</paragraph>
<paragraph id="88184304FF8D4A0BFF6BFB7EBCA5FB31" blockId="7.[189,748,1174,1270]" box="[189,748,1209,1236]" pageId="7" pageNumber="8">
<taxonomicName id="4FA73887FF8D4A0BFF6BFB7EBCC2FB36" ID-CoL="41a2c411-2db0-40a6-a338-7fed1c85dc32" authority="Rosenmejer &amp; Enghoff" authorityName="Rosenmejer &amp; Enghoff" authorityYear="2020" box="[189,651,1209,1235]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoro" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="alopias" status="sp. nov.">
<emphasis id="BAD39F16FF8D4A0BFF6BFB7EBF38FB36" box="[189,369,1209,1235]" italics="true" pageId="7" pageNumber="8">Pogoro alopias</emphasis>
Rosenmejer &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF8D4A0BFD47FB7EBCA5FB31" box="[657,748,1210,1236]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="88184304FF8D4A0BFF6BFB18BC86FB13" blockId="7.[189,748,1174,1270]" box="[189,719,1243,1270]" pageId="7" pageNumber="8">
<taxonomicName id="4FA73887FF8D4A0BFF6BFB18BC27FB10" ID-CoL="f82ea711-b87c-48b5-aed2-1654e66295d0" authority="Rosenmejer &amp; Enghoff" authorityName="Rosenmejer &amp; Enghoff" authorityYear="2020" box="[189,622,1243,1270]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoro" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="siren" status="sp. nov.">
<emphasis id="BAD39F16FF8D4A0BFF6BFB18BF1CFB10" box="[189,341,1244,1269]" italics="true" pageId="7" pageNumber="8">Pogoro siren</emphasis>
Rosenmejer &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF8D4A0BFDA3FB18BC86FB13" box="[629,719,1244,1270]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
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<paragraph id="88184304FF8D4A0BFF6BFAD8BF1FFAD3" blockId="7.[189,342,1308,1334]" box="[189,342,1308,1334]" pageId="7" pageNumber="8">
<heading id="D350F468FF8D4A0BFF6BFAD8BF1FFAD3" bold="true" box="[189,342,1308,1334]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="BAD39F16FF8D4A0BFF6BFAD8BF1FFAD3" bold="true" box="[189,342,1308,1334]" pageId="7" pageNumber="8">Distribution</emphasis>
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Udzungwa Mts,
<collectingCountry id="F0B00394FF8D4A0BFEA9FA8EBFA5FA81" box="[383,492,1354,1380]" name="Tanzania" pageId="7" pageNumber="8">Tanzania</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF8D4A0BFF6BFA4EBC10FA22" pageId="7" pageNumber="8" type="multiple">
<paragraph id="88184304FF8D4A0BFF6BFA4EBC10FA22" blockId="7.[189,1399,1418,1479]" pageId="7" pageNumber="8">
As all known species of
<taxonomicName id="4FA73887FF8D4A0BFE3AFA4FBC0AFA41" authorityName="Hoffman" authorityYear="2005" box="[492,579,1419,1444]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoro" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="BAD39F16FF8D4A0BFE3AFA4FBC0AFA41" box="[492,579,1419,1444]" italics="true" pageId="7" pageNumber="8">Pogoro</emphasis>
</taxonomicName>
live in the Udzungwa Mts, they can be keyed out using the key to Udzungwa gomphodesmids above.
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF8D4A0BFF6BFA2ABD17F904" pageId="7" pageNumber="8" type="discussion">
<paragraph id="88184304FF8D4A0BFF6BFA2ABF66F9ED" blockId="7.[189,303,1518,1544]" box="[189,303,1518,1544]" pageId="7" pageNumber="8">
<heading id="D350F468FF8D4A0BFF6BFA2ABF66F9ED" bold="true" box="[189,303,1518,1544]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="BAD39F16FF8D4A0BFF6BFA2ABF66F9ED" bold="true" box="[189,303,1518,1544]" pageId="7" pageNumber="8">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF8D4A0BFF6BF9DFBD17F904" blockId="7.[189,1399,1563,1761]" pageId="7" pageNumber="8">
All three species of
<taxonomicName id="4FA73887FF8D4A0BFE76F9D8BFBEF9D0" authorityName="Hoffman" authorityYear="2005" box="[416,503,1564,1589]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoro" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="BAD39F16FF8D4A0BFE76F9D8BFBEF9D0" box="[416,503,1564,1589]" italics="true" pageId="7" pageNumber="8">Pogoro</emphasis>
</taxonomicName>
have been collected within a relatively small area in the Udzungwa Scarp FR; the collecting sites are hardly more than
<quantity id="4F5FEEE1FF8D4A0BFD44F9F9BC92F9B2" box="[658,731,1597,1623]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="8" unit="km" value="10.0">10 km</quantity>
apart (cf.
<bibRefCitation id="EC363EF5FF8D4A0BFC87F9FABAC3F9BD" author="Menegon M. &amp; Salvidio S." box="[849,1162,1597,1624]" pageId="7" pageNumber="8" pagination="205 - 212" refId="ref12202" refString="Menegon M. &amp; Salvidio S. 2005. Amphibian and reptile diversity in the southern Udzungwa Scarp Forest Reserve, south-eastern Tanzania. In: Huber B. A., Sinclair B. J. &amp; Lampe K. - H. (eds) African Biodiversity: Molecules, Organisms, Ecosystems. Proceedings of the 5 th International Symposium of Tropical Biology, Museum Koenig, Bonn: 205 - 212. Springer Verlag, Boston. https: // doi. org / 10.1007 / 0 - 387 - 24320 - 8 _ 18" type="book chapter" year="2005">Menegon &amp; Salvidio 2005</bibRefCitation>
: fig. 1). Considering that the species are very similar to each other, and that only one male specimen of each is known for two of them, the possibility that they all represent one variable species cannot be excluded. Additional collecting is needed to elucidate this, but for the time being we consider the differences mentioned in the key and description as warranting separate species status.
</paragraph>
</subSubSection>
</treatment>
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Key to species of
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from the Udzungwa Mts
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The key also includes
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<emphasis id="BAD39F16FF8F4A09FE15FE85BCA0FEBE" box="[451,745,321,347]" italics="true" pageId="5" pageNumber="6">Emphysemastix dracarys</emphasis>
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from
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town.
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<keyLead id="3356E031FF8F4A09FF6BFE46BB3EFE06" pageId="5" pageNumber="6">
1. Sternal process of body ring 6 ca 1½ times as long as wide, with median apiculus (
<figureCitation id="109C5F81FF8F4A09FB42FE46BAB9FE78" box="[1172,1264,386,413]" captionStart="Fig" captionStartId="6.[189,232,1500,1526]" captionTargetBox="[217,1367,909,1445]" captionTargetId="figure@6.[207,1377,905,1453]" captionTargetPageId="6" captionText="Fig. 5. Sternal process on body ring 6. A. Emphysemastix spp. B. Agrophogonus spp. except A. hamulifer Olsen &amp; Enghoff sp. nov. C. Agrophogonus hamulifer Olsen &amp; Enghoff sp. nov. D. Pogoropsis prolixopes Rosenmejer &amp; Enghoff gen. et sp. nov. E. Pogoro siren Rosenmejer &amp; Enghoff sp. nov. F. Pogoro scharffi Hoffman, 2005. G. Pogoro alopias Rosenmejer &amp; Enghoff sp. nov. Not drawn to scale." figureDoi="http://doi.org/10.5281/zenodo.3928336" httpUri="https://zenodo.org/record/3928336/files/figure.png" pageId="5" pageNumber="6">Fig. 5A</figureCitation>
). Postnodal telopodite with a subglobose enlargement along the outer curvature...(Genus
<taxonomicName id="4FA73887FF8F4A09FB6EFE61BFD4FE06" ID-CoL="7PD53" authority="Hoffman, 1966" authorityName="Hoffman" authorityYear="1966" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="BAD39F16FF8F4A09FB6EFE61BB3EFE25" bold="true" box="[1208,1399,421,448]" italics="true" pageId="5" pageNumber="6">Emphysemastix</emphasis>
<bibRefCitation id="EC363EF5FF8F4A09FF3CFE0DBFD4FE06" author="Hoffman R. L." box="[234,413,457,483]" pageId="5" pageNumber="6" pagination="193 - 208" refId="ref11929" refString="Hoffman R. L. 1966. Notes on Aulodesmus and the proposal of a related new genus (Polydesmida: Gomphodesmidae). Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut 63: 193 - 208." type="journal article" year="1966">Hoffman, 1966</bibRefCitation>
</taxonomicName>
).................................................................................................................................. 2
</keyLead>
</paragraph>
<paragraph id="88184304FF8F4A09FF17FE29BB3EFDA9" blockId="5.[189,1399,384,589]" pageId="5" pageNumber="6">
<keyLead id="3356E031FF8F4A09FF17FE29BB3EFDA9" pageId="5" pageNumber="6">
– Sternal process of body ring 6 different (the sternal process of
<taxonomicName id="4FA73887FF8F4A09FC7DFE28BF03FDCC" ID-CoL="304e4a5f-84a3-48a8-a050-0ffa499f6690" authority="Olsen &amp; Enghoff" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="pusillokiellandi" status="sp. nov.">
<emphasis id="BAD39F16FF8F4A09FC7DFE28BB5BFDE3" box="[939,1298,492,518]" italics="true" pageId="5" pageNumber="6">Agrophogonus pusillokiellandi</emphasis>
Olsen &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF8F4A09FE80FDD4BFFFFDCF" box="[342,438,528,554]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
is broken in the unique
<typeStatus id="571CFDA6FF8F4A09FD38FDCBBD1EFDCF" box="[750,855,527,554]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
). Postnodal telopodite without a subglobose enlargement........................................................................................................................................ 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="33535BA1FF8F4A09FF6BFDB0BB3EFD1D" pageId="5" pageNumber="6">
<paragraph id="88184304FF8F4A09FF6BFDB0BB3EFD57" blockId="5.[189,1399,628,760]" pageId="5" pageNumber="6">
<keyLead id="3356E031FF8F4A09FF6BFDB0BB3EFD57" pageId="5" pageNumber="6">
2. Paxillus (mid-ventral process on body ring 15) triangular (as
<figureCitation id="109C5F81FF8F4A09FC6EFDB0BA5FFD6B" box="[952,1046,628,654]" captionStart="Fig" captionStartId="5.[189,232,1803,1829]" captionTargetBox="[196,1394,1386,1769]" captionTargetId="figure@5.[189,1398,1376,1774]" captionTargetPageId="5" captionText="Fig. 4. Particular gomphodesmid structural elements. A. Paxillus – a mid-ventral, most often triangular process on body ring 15 in most gomphodesmid males, from Hoffman (2005), modified. B. Torus – a low convexity of variable size and shape, found on the ventral surface of the poriferous paranota in males of many gomphodesmid species. Not drawn to scale." figureDoi="http://doi.org/10.5281/zenodo.3928334" httpUri="https://zenodo.org/record/3928334/files/figure.png" pageId="5" pageNumber="6">Fig. 4A</figureCitation>
). Nodal process M absent ..... ..........................................................................
<taxonomicName id="4FA73887FF8F4A09FCC0FD52BB5CFD54" ID-CoL="16d88827-f339-4af8-be47-f1d00a209c13" authority="Olsen &amp; Enghoff" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[790,1301,662,689]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="dracarys" status="sp. nov.">
<emphasis id="BAD39F16FF8F4A09FCC0FD52BA0DFD54" bold="true" box="[790,1092,662,689]" italics="true" pageId="5" pageNumber="6">Emphysemastix dracarys</emphasis>
Olsen &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF8F4A09FACAFD5CBB3EFD57" box="[1308,1399,664,690]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="88184304FF8F4A09FF6BFD7FBB3EFD1D" blockId="5.[189,1399,628,760]" pageId="5" pageNumber="6">
<keyLead id="3356E031FF8F4A09FF6BFD7FBB3EFD1D" pageId="5" pageNumber="6">
– Paxillus rounded. Nodal processes M and L both present .................................................................. ..............................................................................
<taxonomicName id="4FA73887FF8F4A09FCFBFD19BB5CFD12" ID-CoL="4d033a7e-aff2-449c-94cf-8fa82cca2e17" authority="Olsen &amp; Enghoff" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[813,1301,733,760]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="frampt" status="sp. nov.">
<emphasis id="BAD39F16FF8F4A09FCFBFD19BA0DFD12" bold="true" box="[813,1092,733,760]" italics="true" pageId="5" pageNumber="6">Emphysemastix frampt</emphasis>
Olsen &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF8F4A09FACAFD1ABB3EFD1D" box="[1308,1399,734,760]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="33535BA1FF8F4A09FF6BFCDBBB3EFC23" pageId="5" pageNumber="6">
<paragraph id="88184304FF8F4A09FF6BFCDBBB3EFC46" blockId="5.[189,1399,799,966]" pageId="5" pageNumber="6">
<keyLead id="3356E031FF8F4A09FF6BFCDBBB3EFC46" pageId="5" pageNumber="6">
3. Sternal process on body ring 6 longer than wide with triangular tip (
<figureCitation id="109C5F81FF8F4A09FBF8FCDBBAD9FCDF" box="[1070,1168,799,826]" captionStart="Fig" captionStartId="6.[189,232,1500,1526]" captionTargetBox="[217,1367,909,1445]" captionTargetId="figure@6.[207,1377,905,1453]" captionTargetPageId="6" captionText="Fig. 5. Sternal process on body ring 6. A. Emphysemastix spp. B. Agrophogonus spp. except A. hamulifer Olsen &amp; Enghoff sp. nov. C. Agrophogonus hamulifer Olsen &amp; Enghoff sp. nov. D. Pogoropsis prolixopes Rosenmejer &amp; Enghoff gen. et sp. nov. E. Pogoro siren Rosenmejer &amp; Enghoff sp. nov. F. Pogoro scharffi Hoffman, 2005. G. Pogoro alopias Rosenmejer &amp; Enghoff sp. nov. Not drawn to scale." figureDoi="http://doi.org/10.5281/zenodo.3928336" httpUri="https://zenodo.org/record/3928336/files/figure.png" pageId="5" pageNumber="6">Fig. 5D</figureCitation>
). Nodal process M present, L absent. Gonopod telopodite making a hairpin bend, postnodal telopodite very long and straight, running close to and parallel with coxa (
<figureCitation id="109C5F81FF8F4A09FCFFFCA1BDC9FC9A" box="[809,896,869,895]" captionStart="Fig" captionStartId="16.[189,232,1929,1955]" captionTargetBox="[202,1375,266,1896]" captionTargetId="figure@16.[202,1384,265,1896]" captionTargetPageId="16" captionText="Fig. 13. Pogoropsis prolixopes Rosenmejer &amp; Enghoff gen. et sp. nov., paratype, ♂, left gonopod (NHMD 621673). A–B. Mesal view. C. Apex of solenomere. D–E. Lateral view. F–G. Dorsal view. Abbreviations: M = Process M; stl = slender triangular lobe; tl = triangular lobe. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928352" httpUri="https://zenodo.org/record/3928352/files/figure.png" pageId="5" pageNumber="6">Fig. 13</figureCitation>
). Paracannular setae absent ....................... ..........................................................
<taxonomicName id="4FA73887FF8F4A09FD4AFC4CBA89FC47" ID-CoL="6eb43d6a-9930-4256-99a9-aecb646378b8" authority="Rosenmejer &amp; Enghoff" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[668,1216,904,931]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoropsis" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="prolixopes" status="gen. et sp. nov.">
<emphasis id="BAD39F16FF8F4A09FD4AFC4CBDEFFC46" bold="true" box="[668,934,904,931]" italics="true" pageId="5" pageNumber="6">Pogoropsis prolixopes</emphasis>
Rosenmejer &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF8F4A09FB11FC4DBB3EFC46" box="[1223,1399,905,931]" pageId="5" pageNumber="6" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="88184304FF8F4A09FF1EFC68BB3EFC23" blockId="5.[189,1399,799,966]" box="[200,1399,940,966]" pageId="5" pageNumber="6">
<keyLead id="3356E031FF8F4A09FF1EFC68BB3EFC23" box="[200,1399,940,966]" pageId="5" pageNumber="6">– Sternal process of body ring 6 and postnodal telopodite different .................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="33535BA1FF8F4A09FF6BFC29BB3EFB94" pageId="5" pageNumber="6">
<paragraph id="88184304FF8F4A09FF6BFC29BB3EFBCE" blockId="5.[189,1399,1005,1137]" pageId="5" pageNumber="6">
<keyLead id="3356E031FF8F4A09FF6BFC29BB3EFBCE" pageId="5" pageNumber="6">
4. Sternal process of body ring 6 broad and flat, or triangular with rounded tip (
<figureCitation id="109C5F81FF8F4A09FB43FC29BA95FBED" box="[1173,1244,1005,1032]" captionStart="Fig" captionStartId="6.[189,232,1500,1526]" captionTargetBox="[217,1367,909,1445]" captionTargetId="figure@6.[207,1377,905,1453]" captionTargetPageId="6" captionText="Fig. 5. Sternal process on body ring 6. A. Emphysemastix spp. B. Agrophogonus spp. except A. hamulifer Olsen &amp; Enghoff sp. nov. C. Agrophogonus hamulifer Olsen &amp; Enghoff sp. nov. D. Pogoropsis prolixopes Rosenmejer &amp; Enghoff gen. et sp. nov. E. Pogoro siren Rosenmejer &amp; Enghoff sp. nov. F. Pogoro scharffi Hoffman, 2005. G. Pogoro alopias Rosenmejer &amp; Enghoff sp. nov. Not drawn to scale." figureDoi="http://doi.org/10.5281/zenodo.3928336" httpUri="https://zenodo.org/record/3928336/files/figure.png" pageId="5" pageNumber="6">Fig. 5</figureCitation>
E–G). Nodal process M present...(Genus
<taxonomicName id="4FA73887FF8F4A09FDE4FBD5BD0DFBCE" ID-CoL="7PV5L" authority="Hoffman, 2005" authorityName="Hoffman" authorityYear="2005" box="[562,836,1040,1067]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoro" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="BAD39F16FF8F4A09FDE4FBD5BCC3FBCE" bold="true" box="[562,650,1041,1067]" italics="true" pageId="5" pageNumber="6">Pogoro</emphasis>
<bibRefCitation id="EC363EF5FF8F4A09FD47FBD4BD0DFBCE" author="Hoffman R. L." box="[657,836,1040,1067]" pageId="5" pageNumber="6" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman, 2005</bibRefCitation>
</taxonomicName>
)........................................................................ 5
</keyLead>
</paragraph>
<paragraph id="88184304FF8F4A09FF17FBF0BB3EFB94" blockId="5.[189,1399,1005,1137]" pageId="5" pageNumber="6">
<keyLead id="3356E031FF8F4A09FF17FBF0BB3EFB94" pageId="5" pageNumber="6">
– Sternal process of body ring 6 more or less elongate, apically rounded (
<figureCitation id="109C5F81FF8F4A09FF25FB93BF71FB94" box="[243,312,1110,1137]" captionStart="Fig" captionStartId="6.[189,232,1500,1526]" captionTargetBox="[217,1367,909,1445]" captionTargetId="figure@6.[207,1377,905,1453]" captionTargetPageId="6" captionText="Fig. 5. Sternal process on body ring 6. A. Emphysemastix spp. B. Agrophogonus spp. except A. hamulifer Olsen &amp; Enghoff sp. nov. C. Agrophogonus hamulifer Olsen &amp; Enghoff sp. nov. D. Pogoropsis prolixopes Rosenmejer &amp; Enghoff gen. et sp. nov. E. Pogoro siren Rosenmejer &amp; Enghoff sp. nov. F. Pogoro scharffi Hoffman, 2005. G. Pogoro alopias Rosenmejer &amp; Enghoff sp. nov. Not drawn to scale." figureDoi="http://doi.org/10.5281/zenodo.3928336" httpUri="https://zenodo.org/record/3928336/files/figure.png" pageId="5" pageNumber="6">Fig. 5</figureCitation>
B–C). Nodal process M absent...(Genus
<taxonomicName id="4FA73887FF8F4A09FCC2FB92BAC8FB94" ID-CoL="7NNCP" authority="Hoffman, 2005" authorityName="Hoffman" authorityYear="2005" box="[788,1153,1110,1137]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="BAD39F16FF8F4A09FCC2FB92BD8EFB94" bold="true" box="[788,967,1110,1137]" italics="true" pageId="5" pageNumber="6">Agrophogonus</emphasis>
<bibRefCitation id="EC363EF5FF8F4A09FC18FB93BAC8FB94" author="Hoffman R. L." box="[974,1153,1110,1137]" pageId="5" pageNumber="6" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman, 2005</bibRefCitation>
</taxonomicName>
)............................. 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="33535BA1FF8F4A09FF6BFB5CBB3EFB1C" pageId="5" pageNumber="6">
<paragraph id="88184304FF8F4A09FF6BFB5CBB3EFB57" blockId="5.[189,1399,1176,1273]" box="[189,1399,1176,1203]" pageId="5" pageNumber="6">
<keyLead id="3356E031FF8F4A09FF6BFB5CBB3EFB57" box="[189,1399,1176,1203]" pageId="5" pageNumber="6">5. Enlarged tubercles on hypoproct projecting beyond edge of sclerite................................................ 6</keyLead>
</paragraph>
<paragraph id="88184304FF8F4A09FF6BFB78BB3EFB1C" blockId="5.[189,1399,1176,1273]" pageId="5" pageNumber="6">
<keyLead id="3356E031FF8F4A09FF6BFB78BB3EFB1C" pageId="5" pageNumber="6">
– Tubercles on hypoproct not projecting beyond edge of sclerite.......................................................... .....................................................................................
<taxonomicName id="4FA73887FF8F4A09FCB7FB1BBB5CFB1D" ID-CoL="f82ea711-b87c-48b5-aed2-1654e66295d0" authority="Rosenmejer &amp; Enghoff" authorityName="Rosenmejer &amp; Enghoff" authorityYear="2020" box="[865,1301,1246,1273]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoro" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="siren" status="sp. nov.">
<emphasis id="BAD39F16FF8F4A09FCB7FB1BBDB5FB1D" bold="true" box="[865,1020,1246,1273]" italics="true" pageId="5" pageNumber="6">Pogoro siren</emphasis>
Rosenmejer &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF8F4A09FACAFB1BBB3EFB1C" box="[1308,1399,1247,1273]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<caption id="DCD8138CFF8F4A09FF6BF8CFBD3CF86A" ID-DOI="http://doi.org/10.5281/zenodo.3928334" ID-Zenodo-Dep="3928334" httpUri="https://zenodo.org/record/3928334/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1803,1829]" targetBox="[196,1394,1386,1769]" targetPageId="5">
<paragraph id="88184304FF8F4A09FF6BF8CFBD3CF86A" blockId="5.[189,1399,1803,1935]" pageId="5" pageNumber="6">
<emphasis id="BAD39F16FF8F4A09FF6BF8CFBF43F8C0" bold="true" box="[189,266,1803,1829]" pageId="5" pageNumber="6">Fig. 4.</emphasis>
Particular gomphodesmid structural elements.
<emphasis id="BAD39F16FF8F4A09FCE3F8CFBD03F8C0" bold="true" box="[821,842,1803,1829]" pageId="5" pageNumber="6">A</emphasis>
. Paxillus – a mid-ventral, most often triangular process on body ring 15 in most gomphodesmid males, from
<bibRefCitation id="EC363EF5FF8F4A09FC4DF8EABA17F8AC" author="Hoffman R. L." box="[923,1118,1838,1865]" pageId="5" pageNumber="6" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman (2005)</bibRefCitation>
, modified.
<emphasis id="BAD39F16FF8F4A09FB32F8EBBAB1F8AC" bold="true" box="[1252,1272,1839,1865]" pageId="5" pageNumber="6">B</emphasis>
. Torus – a low convexity of variable size and shape, found on the ventral surface of the poriferous paranota in males of many gomphodesmid species. Not drawn to scale.
</paragraph>
</caption>
<keyStep id="33535BA1FF8C4A0AFF6BFECFBB3EFE75" pageId="6" pageNumber="7">
<paragraph id="88184304FF8C4A0AFF6BFECFBB3EFEAC" blockId="6.[189,1399,267,400]" pageId="6" pageNumber="7">
<keyLead id="3356E031FF8C4A0AFF6BFECFBB3EFEAC" pageId="6" pageNumber="7">
6. Two subapical lobes on postnodal telopodite, a triangular lateral lobe and an uncate mesal lobe (
<figureCitation id="109C5F81FF8C4A0AFF25FEEBBF70FEAC" box="[243,313,303,329]" captionStart="Fig" captionStartId="9.[189,232,1929,1955]" captionTargetBox="[297,1266,266,1882]" captionTargetId="figure@9.[297,1280,265,1882]" captionTargetPageId="9" captionText="Fig. 7. Pogoro scharffi Hoffman, 2005, paratype, ♂, left gonopod (NHMD 621669). A–B. Mesal view. C–D. Lateral view. E–F. Dorsal view. Abbreviations: L = Process L; M = Process M; tl = triangular lobe; ul = uncate lobe. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928340" httpUri="https://zenodo.org/record/3928340/files/figure.png" pageId="6" pageNumber="7">Fig. 7</figureCitation>
) ..............................................................................................
<taxonomicName id="4FA73887FF8C4A0AFBD6FEEBBB3EFEAC" ID-CoL="4KWC7" authority="Hoffman, 2005" authorityName="Hoffman" authorityYear="2005" box="[1024,1399,302,329]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoro" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="scharffi">
<emphasis id="BAD39F16FF8C4A0AFBD6FEEBBAF5FEAC" bold="true" box="[1024,1212,302,329]" italics="true" pageId="6" pageNumber="7">Pogoro scharffi</emphasis>
<bibRefCitation id="EC363EF5FF8C4A0AFB12FEEBBB3EFEAC" author="Hoffman R. L." box="[1220,1399,302,329]" pageId="6" pageNumber="7" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman, 2005</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="88184304FF8C4A0AFF17FE96BB3EFE75" blockId="6.[189,1399,267,400]" pageId="6" pageNumber="7">
<keyLead id="3356E031FF8C4A0AFF17FE96BB3EFE75" pageId="6" pageNumber="7">
– Only one subapical lobe, of a narrow triangular shape, on postnodal telopodite (
<figureCitation id="109C5F81FF8C4A0AFF25FEB1BF70FE6A" box="[243,313,373,399]" captionStart="Fig" captionStartId="11.[189,232,1932,1958]" captionTargetBox="[369,1206,265,1908]" captionTargetId="figure@11.[369,1216,265,1908]" captionTargetPageId="11" captionText="Fig. 9. Pogoro alopias Rosenmejer &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621670), right gonopod. A–B. Mesal view. C–D. Lateral view. E–F. Dorsal view. Abbreviations: L = Process L; M = Process M; tl = triangular lobe. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928344" httpUri="https://zenodo.org/record/3928344/files/figure.png" pageId="6" pageNumber="7">Fig. 9</figureCitation>
) .....................................................................
<taxonomicName id="4FA73887FF8C4A0AFC91FEB1BB5CFE6A" ID-CoL="41a2c411-2db0-40a6-a338-7fed1c85dc32" authority="Rosenmejer &amp; Enghoff" authorityName="Rosenmejer &amp; Enghoff" authorityYear="2020" box="[839,1301,373,400]" class="Diplopoda" family="Gomphodesmidae" genus="Pogoro" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="alopias" status="sp. nov.">
<emphasis id="BAD39F16FF8C4A0AFC91FEB1BDB5FE75" bold="true" box="[839,1020,373,400]" italics="true" pageId="6" pageNumber="7">Pogoro alopias</emphasis>
Rosenmejer &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF8C4A0AFACAFEB2BB3EFE75" box="[1308,1399,374,400]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="33535BA1FF8C4A0AFF6BFE73BB3EFE11" pageId="6" pageNumber="7">
<paragraph id="88184304FF8C4A0AFF6BFE73BB3EFE34" blockId="6.[189,1399,438,500]" box="[189,1399,438,465]" pageId="6" pageNumber="7">
<keyLead id="3356E031FF8C4A0AFF6BFE73BB3EFE34" box="[189,1399,438,465]" pageId="6" pageNumber="7">
7. Postnodal telopodite short,flattened,laminate(
<figureCitation id="109C5F81FF8C4A0AFCD6FE72BD18FE34" box="[768,849,438,465]" captionStart="Fig" captionStartId="29.[190,233,1929,1955]" captionTargetBox="[349,1236,266,1886]" captionTargetId="figure@29.[347,1237,265,1887]" captionTargetPageId="29" captionText="Fig. 19. Agrophogonus mwanihana Hoffman, 2005, specimen from Udzungwa Mts National Park, Sanje Kati camp &amp; plot (NHMD 621682), right gonopod. A–B. Mesal view. C–D. Lateral view. E–F. Dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928364" httpUri="https://zenodo.org/record/3928364/files/figure.png" pageId="6" pageNumber="7">Fig.19</figureCitation>
)…
<taxonomicName id="4FA73887FF8C4A0AFCA8FE72BB3EFE34" ID-CoL="B8S4" authority="Hoffman, 2995" authorityName="Hoffman" authorityYear="2995" box="[894,1399,438,465]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="mwanihana">
<emphasis id="BAD39F16FF8C4A0AFCA8FE72BA8DFE34" bold="true" box="[894,1220,438,465]" italics="true" pageId="6" pageNumber="7">Agrophogonus mwanihana</emphasis>
Hoffman, 2995
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<keyLead id="3356E031FF8C4A0AFF17FE1EBB3EFE11" box="[193,1399,474,500]" pageId="6" pageNumber="7">
<emphasis id="BAD39F16FF8C4A0AFF17FE1EBE99FE16" box="[193,208,474,499]" italics="true" pageId="6" pageNumber="7">–</emphasis>
Postnodal telopodite longer, slender, evenly attenuate to apex ......................................................... 8
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<paragraph id="88184304FF8C4A0AFF6BFDDFBB3EFDBC" blockId="6.[189,1399,537,671]" pageId="6" pageNumber="7">
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8. Postnodal telopodite very long, forming a subcircular ¾ loop, evenly curved, no sharp angular twist. (
<figureCitation id="109C5F81FF8C4A0AFF25FDFABF01FDBC" box="[243,328,574,601]" captionStart="Fig" captionStartId="26.[189,232,1929,1955]" captionTargetBox="[317,1259,268,1898]" captionTargetId="figure@26.[317,1268,268,1898]" captionTargetPageId="26" captionText="Fig. 18. Agrophogonus hamulifer Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621678), left gonopod. A–B. Mesal view. C–D. Lateral view. E–F. Dorsal view. Abbreviations: A = Process A; B = Process B; C = Process C; L = Process L. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928362" httpUri="https://zenodo.org/record/3928362/files/figure.png" pageId="6" pageNumber="7">Fig. 18</figureCitation>
) ............................................................
<taxonomicName id="4FA73887FF8C4A0AFCC7FDFABB5CFDBD" ID-CoL="9f1516ed-177e-4f9a-b6fc-d2c3dc1f0513" authority="Olsen &amp; Enghoff" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[785,1301,574,601]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="hamulifer" status="sp. nov.">
<emphasis id="BAD39F16FF8C4A0AFCC7FDFABA0DFDBC" bold="true" box="[785,1092,574,601]" italics="true" pageId="6" pageNumber="7">Agrophogonus hamulifer</emphasis>
Olsen &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF8C4A0AFACAFDFBBB3EFDBC" box="[1308,1399,575,601]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
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<keyLead id="3356E031FF8C4A0AFF17FDA6BB3EFD7A" pageId="6" pageNumber="7">
– Postnodal telopodite shorter, with a sharp twist in a dorsal direction, perpendicular to prefemoral region, not forming a ¾ circle (
<figureCitation id="109C5F81FF8C4A0AFD8CFD41BCE7FD7A" box="[602,686,645,671]" captionStart="Fig" captionStartId="31.[189,232,1895,1921]" captionTargetBox="[236,1337,265,1863]" captionTargetId="figure@31.[234,1350,265,1863]" captionTargetPageId="31" captionText="Fig. 21. Agrophogonus pusillokiellandi Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621683). A–D, F–G. Left gonopod. A–B. Mesal view. C–D. Lateral view. F–G. Dorsal view. E. Right gonopod, lateral view. H. Sternal process on ring 6. Abbreviations: A = process A; B = process B; C = process C; L = process L. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928368" httpUri="https://zenodo.org/record/3928368/files/figure.png" pageId="6" pageNumber="7">Fig. 21</figureCitation>
) ............................................................................................ 9
</keyLead>
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<keyStep id="33535BA1FF8C4A0AFF6BFD02BB3EFCAF" pageId="6" pageNumber="7">
<paragraph id="88184304FF8C4A0AFF6BFD02BB3EFCE1" blockId="6.[189,1399,710,842]" pageId="6" pageNumber="7">
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9. Body width
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. Nodal process C very long, slender and curved (
<bibRefCitation id="EC363EF5FF8C4A0AFBCEFD02BA83FD04" author="Hoffman R. L." box="[1048,1226,710,737]" pageId="6" pageNumber="7" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman 2005</bibRefCitation>
: fig. 451) ....... ............................................................................................
<taxonomicName id="4FA73887FF8C4A0AFC4EFD2DBB3EFCE1" ID-CoL="B8S2" authority="Hoffman, 2005" authorityName="Hoffman" authorityYear="2005" box="[920,1399,745,772]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="kiellandi">
<emphasis id="BAD39F16FF8C4A0AFC4EFD2DBAF5FCE1" bold="true" box="[920,1212,745,772]" italics="true" pageId="6" pageNumber="7">Agrophogonus kiellandi</emphasis>
<bibRefCitation id="EC363EF5FF8C4A0AFB12FD2DBB3EFCE1" author="Hoffman R. L." box="[1220,1399,745,772]" pageId="6" pageNumber="7" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman, 2005</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="88184304FF8C4A0AFF17FCC9BB3EFCAF" blockId="6.[189,1399,710,842]" pageId="6" pageNumber="7">
<keyLead id="3356E031FF8C4A0AFF17FCC9BB3EFCAF" pageId="6" pageNumber="7">
– Body width
<quantity id="4F5FEEE1FF8C4A0AFEADFCC9BF91FCC2" box="[379,472,781,807]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="6" pageNumber="7" unit="mm" value="4.1">4.1 mm</quantity>
. Nodal process C shorter and less curved (
<figureCitation id="109C5F81FF8C4A0AFC1DFCC9BA56FCC2" box="[971,1055,781,807]" captionStart="Fig" captionStartId="31.[189,232,1895,1921]" captionTargetBox="[236,1337,265,1863]" captionTargetId="figure@31.[234,1350,265,1863]" captionTargetPageId="31" captionText="Fig. 21. Agrophogonus pusillokiellandi Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621683). A–D, F–G. Left gonopod. A–B. Mesal view. C–D. Lateral view. F–G. Dorsal view. E. Right gonopod, lateral view. H. Sternal process on ring 6. Abbreviations: A = process A; B = process B; C = process C; L = process L. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928368" httpUri="https://zenodo.org/record/3928368/files/figure.png" pageId="6" pageNumber="7">Fig. 21</figureCitation>
) ............................................ .................................................................
<taxonomicName id="4FA73887FF8C4A0AFD18FCEBBB5CFCAF" ID-CoL="304e4a5f-84a3-48a8-a050-0ffa499f6690" authority="Olsen &amp; Enghoff" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[718,1301,815,842]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="pusillokiellandi" status="sp. nov.">
<emphasis id="BAD39F16FF8C4A0AFD18FCEBBA0DFCAF" bold="true" box="[718,1092,815,842]" italics="true" pageId="6" pageNumber="7">Agrophogonus pusillokiellandi</emphasis>
Olsen &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF8C4A0AFACAFCF4BB3EFCAF" box="[1308,1399,816,842]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
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</key>
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</treatment>
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110
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<emphasis id="BAD39F16FF914A17FDFCF944BD06F97E" bold="true" box="[554,847,1664,1691]" italics="true" pageId="27" pageNumber="28">Agrophogonus kiellandi</emphasis>
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<emphasis id="BAD39F16FF914A17FF6BF900BF9EF93B" box="[189,471,1732,1758]" italics="true" pageId="27" pageNumber="28">Agrophogonus kiellandi</emphasis>
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.
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<emphasis id="BAD39F16FF914A17FF6BF8C3BFE3F8C4" bold="true" box="[189,426,1799,1825]" pageId="27" pageNumber="28">Material examined</emphasis>
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<paragraph id="88184304FF914A17FF6BF8FCBF4DF8B7" blockId="27.[189,260,1848,1874]" box="[189,260,1848,1874]" pageId="27" pageNumber="28">None.</paragraph>
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<paragraph id="88184304FF914A17FF6BF8BFBF1FF870" blockId="27.[189,342,1915,1941]" box="[189,342,1915,1941]" pageId="27" pageNumber="28">
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Known only from the
<typeStatus id="571CFDA6FF914A17FE1BF868BC49F823" box="[461,512,1964,1990]" pageId="27" pageNumber="28">type</typeStatus>
locality,
<collectingRegion id="4A638DE6FF914A17FDA7F868BD50F823" box="[625,793,1964,1990]" country="Tanzania" name="Iringa" pageId="27" pageNumber="28">Iringa Region</collectingRegion>
,
<collectingRegion id="4A638DE6FF914A17FCF1F868BD9BF823" box="[807,978,1964,1990]" country="Tanzania" name="Iringa" pageId="27" pageNumber="28">Iringa District</collectingRegion>
, Udzungwa Mts, Mt. Nyumbenito,
<quantity id="4F5FEEE1FF914A17FF6BF815BF5FF80E" box="[189,278,2001,2027]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="27" pageNumber="28" unit="m" value="2000.0">2000 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="DBE10C3EC1E38B4F3BD47BC5C4F45E89">A mountain of millipedes IX: Species of the family Gomphodesmidae from the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida)</mods:title>
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<heading id="D350F468FF924A14FDECFCF6BDB3FCA9" box="[570,1018,817,844]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="25" reason="2">
Genus
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<emphasis id="BAD39F16FF924A14FD5AFCF5BD76FCA9" bold="true" box="[652,831,817,844]" italics="true" pageId="24" pageNumber="25">Agrophogonus</emphasis>
<bibRefCitation id="EC363EF5FF924A14FC91FCF5BDB3FCA9" author="Hoffman R. L." box="[839,1018,817,844]" pageId="24" pageNumber="25" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman, 2005</bibRefCitation>
</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF924A14FF6BFCB1BC94FA08" pageId="24" pageNumber="25" type="diagnosis">
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<emphasis id="BAD39F16FF924A14FF6BFCB1BF7FFC6A" bold="true" box="[189,310,885,911]" pageId="24" pageNumber="25">Diagnosis</emphasis>
(after
<bibRefCitation id="EC363EF5FF924A14FE52FCB1BC7BFC75" author="Hoffman R. L." box="[388,562,885,912]" pageId="24" pageNumber="25" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman 2005</bibRefCitation>
, modified)
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Small to medium-sized (length
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) gomphodesmids. Four apical sensory cones on antennae, pore formula: 5, 7, 9–19. Terga smooth. Legs 1–7 of males with large and conspicuous subonychial pads; coxae of 6
<superScript id="7FD2EE4CFF924A14FEE3FC34BF0BFC1B" attach="left" box="[309,322,1008,1022]" fontSize="6" pageId="24" pageNumber="25">th</superScript>
pair conically produced; sternum of ring 6 with elongate, apically rounded process; sterna of several postgonopodal rings with indistinct transverse carinae (apparently absent in
<taxonomicName id="4FA73887FF924A14FB48FBD3BB3EFBD4" authority="Olsen &amp; Rosenmejer &amp; Enghoff, 2020" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[1182,1399,1047,1073]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pusillokiellandi" status="sp. nov.">
<emphasis id="BAD39F16FF924A14FB48FBD3BB3EFBD4" box="[1182,1399,1047,1073]" italics="true" pageId="24" pageNumber="25">A. pusillokiellandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF924A14FF6BFBF9BF55FBB2" box="[189,284,1085,1111]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
); no paxillus on sternum 15. Side of metazona smooth, pleurosternal carinae present. No torus evident.
</paragraph>
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Gonopod aperture oval, posterior rim elevated into triangular lobes immediately behind each gonocoxa, cut down to level of sternum medially. Gonopods relatively long and slender, coxae in contact medially, without projection over base of cannula; telepodite set against coxa at a right angle, prefemur slender, nearly straight, gradually narrowed distally, without basal excavation on either side. Nodus entirely endonodal, without trace of median process M, lateral process L present or absent; inner edge of nodus produced into three or four acute projections of varying shape; postnodal telepodite relatively short, curved laterad (longer and curved in a ¾ circle in
<taxonomicName id="4FA73887FF924A14FCF0FA4CBDF2FA44" authority="Olsen &amp; Rosenmejer &amp; Enghoff, 2020" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[806,955,1415,1442]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="hamulifer" status="sp. nov.">
<emphasis id="BAD39F16FF924A14FCF0FA4CBDF2FA44" box="[806,955,1415,1442]" italics="true" pageId="24" pageNumber="25">A. hamulifer</emphasis>
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<taxonomicNameLabel id="A1E0226DFF924A14FC10FA4CBA6EFA47" box="[966,1063,1416,1442]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
), distally either slender and acuminate or laminately flattened, rather abruptly flexed immediately beyond nodus; prostatic groove visible over most of its length in mesal aspect.
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<paragraph id="88184304FF924A14FF6BF9D3BFC0F9D4" blockId="24.[189,393,1559,1585]" box="[189,393,1559,1585]" pageId="24" pageNumber="25">
<heading id="D350F468FF924A14FF6BF9D3BFC0F9D4" bold="true" box="[189,393,1559,1585]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="BAD39F16FF924A14FF6BF9D3BFC0F9D4" bold="true" box="[189,393,1559,1585]" pageId="24" pageNumber="25">Included species</emphasis>
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<taxonomicName id="4FA73887FF924A14FF6BF983BCECF987" ID-CoL="B8S3" authority="Hoffman, 2005" authorityName="Hoffman" authorityYear="2005" box="[189,677,1607,1634]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="mangalisa">
<emphasis id="BAD39F16FF924A14FF6BF983BFA3F984" box="[189,490,1607,1633]" italics="true" pageId="24" pageNumber="25">Agrophogonus mangalisa</emphasis>
<bibRefCitation id="EC363EF5FF924A14FE24F983BCECF987" author="Hoffman R. L." box="[498,677,1607,1634]" pageId="24" pageNumber="25" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman, 2005</bibRefCitation>
</taxonomicName>
(
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species)
</paragraph>
<paragraph id="88184304FF924A14FF6BF9A9BD5CF96D" blockId="24.[189,857,1607,1865]" box="[189,789,1645,1672]" pageId="24" pageNumber="25">
<taxonomicName id="4FA73887FF924A14FF6BF9A9BCFAF962" ID-CoL="9f1516ed-177e-4f9a-b6fc-d2c3dc1f0513" authority="Olsen &amp; Enghoff" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[189,691,1645,1671]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="hamulifer" status="sp. nov.">
<emphasis id="BAD39F16FF924A14FF6BF9A9BFABF962" box="[189,482,1645,1671]" italics="true" pageId="24" pageNumber="25">Agrophogonus hamulifer</emphasis>
Olsen &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF924A14FD6CF9AABD5CF96D" box="[698,789,1646,1672]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="88184304FF924A14FF6BF957BCC2F948" blockId="24.[189,857,1607,1865]" box="[189,651,1683,1709]" pageId="24" pageNumber="25">
<taxonomicName id="4FA73887FF924A14FF6BF957BCC2F948" ID-CoL="B8RZ" authority="(Verhoeff, 1941)" baseAuthorityName="Verhoeff" baseAuthorityYear="1941" box="[189,651,1683,1709]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="harrisi">
<emphasis id="BAD39F16FF924A14FF6BF957BF89F948" box="[189,448,1683,1709]" italics="true" pageId="24" pageNumber="25">Agrophogonus harrisi</emphasis>
(Verhoeff, 1941)
</taxonomicName>
</paragraph>
<paragraph id="88184304FF924A14FF6BF97CBCD8F936" blockId="24.[189,857,1607,1865]" box="[189,657,1720,1747]" pageId="24" pageNumber="25">
<taxonomicName id="4FA73887FF924A14FF6BF97CBCD8F936" ID-CoL="B8S2" authority="Hoffman, 2005" authorityName="Hoffman" authorityYear="2005" box="[189,657,1720,1747]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="kiellandi">
<emphasis id="BAD39F16FF924A14FF6BF97CBF9EF937" box="[189,471,1720,1746]" italics="true" pageId="24" pageNumber="25">Agrophogonus kiellandi</emphasis>
<bibRefCitation id="EC363EF5FF924A14FE08F97CBCD8F936" author="Hoffman R. L." box="[478,657,1720,1747]" pageId="24" pageNumber="25" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman, 2005</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="88184304FF924A14FF6BF91ABCFAF91C" blockId="24.[189,857,1607,1865]" box="[189,691,1758,1785]" pageId="24" pageNumber="25">
<taxonomicName id="4FA73887FF924A14FF6BF91ABCFAF91C" ID-CoL="B8S4" authority="Hoffman, 2005" authorityName="Hoffmann" authorityYear="2005" box="[189,691,1758,1785]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="mwanihana">
<emphasis id="BAD39F16FF924A14FF6BF91ABFB0F91D" box="[189,505,1758,1784]" italics="true" pageId="24" pageNumber="25">Agrophogonus mwanihana</emphasis>
<bibRefCitation id="EC363EF5FF924A14FDD6F91ABCFAF91C" author="Hoffman R. L." box="[512,691,1758,1785]" pageId="24" pageNumber="25" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman, 2005</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="88184304FF924A14FF6BF8C2BD10F8C4" blockId="24.[189,857,1607,1865]" box="[189,857,1798,1825]" pageId="24" pageNumber="25">
<taxonomicName id="4FA73887FF924A14FF6BF8C2BCB1F8C5" ID-CoL="304e4a5f-84a3-48a8-a050-0ffa499f6690" authority="Olsen &amp; Enghoff" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[189,760,1798,1824]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pusillokiellandi" status="sp. nov.">
<emphasis id="BAD39F16FF924A14FF6BF8C2BC6EF8C5" box="[189,551,1798,1824]" italics="true" pageId="24" pageNumber="25">Agrophogonus pusillokiellandi</emphasis>
Olsen &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF924A14FD29F8C3BD10F8C4" box="[767,857,1799,1825]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="88184304FF924A14FF6BF8EABC35F8AC" blockId="24.[189,857,1607,1865]" box="[189,636,1838,1865]" pageId="24" pageNumber="25">
<taxonomicName id="4FA73887FF924A14FF6BF8EABC35F8AC" ID-CoL="B8S5" authority="Hoffman, 2005" authorityName="Hoffman" authorityYear="2005" box="[189,636,1838,1865]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="BAD39F16FF924A14FF6BF8EABF88F8AD" box="[189,449,1838,1864]" italics="true" pageId="24" pageNumber="25">Agrophogonus tridens</emphasis>
<bibRefCitation id="EC363EF5FF924A14FE1FF8EABC35F8AC" author="Hoffman R. L." box="[457,636,1838,1865]" pageId="24" pageNumber="25" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman, 2005</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF924A14FF6BF8B1BFD6F80E" pageId="24" pageNumber="25" type="distribution">
<paragraph id="88184304FF924A14FF6BF8B1BF1FF86A" blockId="24.[189,342,1909,1935]" box="[189,342,1909,1935]" pageId="24" pageNumber="25">
<heading id="D350F468FF924A14FF6BF8B1BF1FF86A" bold="true" box="[189,342,1909,1935]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="BAD39F16FF924A14FF6BF8B1BF1FF86A" bold="true" box="[189,342,1909,1935]" pageId="24" pageNumber="25">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF924A14FF6BF86CBFD6F80E" blockId="24.[189,1399,1960,2027]" pageId="24" pageNumber="25">
<collectingCountry id="F0B00394FF924A14FF6BF86CBF63F827" box="[189,298,1960,1986]" name="Tanzania" pageId="24" pageNumber="25">Tanzania</collectingCountry>
. Two of the previously described species are known from the Udzungwa Mts, viz.,
<taxonomicName id="4FA73887FF924A14FB26F86DBB3EF827" authorityName="Hoffman" authorityYear="2005" box="[1264,1399,1960,1987]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="kiellandi">
<emphasis id="BAD39F16FF924A14FB26F86DBB3EF827" box="[1264,1399,1960,1987]" italics="true" pageId="24" pageNumber="25">A. kiellandi</emphasis>
</taxonomicName>
and
<taxonomicName id="4FA73887FF924A14FF39F815BFD1F80E" authorityName="Hoffmann" authorityYear="2005" box="[239,408,2001,2027]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="mwanihana">
<emphasis id="BAD39F16FF924A14FF39F815BFD1F80E" box="[239,408,2001,2027]" italics="true" pageId="24" pageNumber="25">A. mwanihana</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF934A15FF6BFEC8BB3EFABB" pageId="25" pageNumber="26" type="key">
<paragraph id="88184304FF934A15FF6BFEC8BDF3FEC3" blockId="25.[189,954,267,294]" box="[189,954,267,294]" pageId="25" pageNumber="26">
<emphasis id="BAD39F16FF934A15FF6BFEC8BC0EFEC3" bold="true" box="[189,583,267,294]" pageId="25" pageNumber="26">
Key to species of
<taxonomicName id="4FA73887FF934A15FE42FECFBC0EFEC3" ID-CoL="7NNCP" authorityName="Hoffman" authorityYear="2005" box="[404,583,267,294]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="BAD39F16FF934A15FE42FECFBC0EFEC3" bold="true" box="[404,583,267,294]" italics="true" pageId="25" pageNumber="26">Agrophogonus</emphasis>
</taxonomicName>
</emphasis>
(modified after
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)
</paragraph>
<key id="FC361435FF934A15FF6BFE87BB3EFABB" pageId="25" pageNumber="26">
<keyStep id="33535BA1FF934A15FF6BFE87BB3EFE6D" pageId="25" pageNumber="26">
<paragraph id="88184304FF934A15FF6BFE87BB3EFEB8" blockId="25.[189,1399,322,393]" box="[189,1399,322,349]" pageId="25" pageNumber="26">
<keyLead id="3356E031FF934A15FF6BFE87BB3EFEB8" box="[189,1399,322,349]" pageId="25" pageNumber="26">
1. Postnodal telepodite short, flattened, laminate (as
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)............................................................... 2
</keyLead>
</paragraph>
<paragraph id="88184304FF934A15FF17FEABBB3EFE6D" blockId="25.[189,1399,322,393]" box="[193,1399,366,393]" pageId="25" pageNumber="26">
<keyLead id="3356E031FF934A15FF17FEABBB3EFE6D" box="[193,1399,366,393]" pageId="25" pageNumber="26">
– Postnodal telepodite mostly longer, slender, evenly attenuate to apex (as
<figureCitation id="109C5F81FF934A15FBE9FEAABAD1FE6D" box="[1087,1176,366,392]" captionStart="Fig" captionStartId="26.[189,232,1929,1955]" captionTargetBox="[317,1259,268,1898]" captionTargetId="figure@26.[317,1268,268,1898]" captionTargetPageId="26" captionText="Fig. 18. Agrophogonus hamulifer Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621678), left gonopod. A–B. Mesal view. C–D. Lateral view. E–F. Dorsal view. Abbreviations: A = Process A; B = Process B; C = Process C; L = Process L. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928362" httpUri="https://zenodo.org/record/3928362/files/figure.png" pageId="25" pageNumber="26">Figs 18</figureCitation>
,
<figureCitation id="109C5F81FF934A15FB73FEAABA8AFE6D" box="[1189,1219,366,392]" captionStart="Fig" captionStartId="31.[189,232,1895,1921]" captionTargetBox="[236,1337,265,1863]" captionTargetId="figure@31.[234,1350,265,1863]" captionTargetPageId="31" captionText="Fig. 21. Agrophogonus pusillokiellandi Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621683). A–D, F–G. Left gonopod. A–B. Mesal view. C–D. Lateral view. F–G. Dorsal view. E. Right gonopod, lateral view. H. Sternal process on ring 6. Abbreviations: A = process A; B = process B; C = process C; L = process L. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928368" httpUri="https://zenodo.org/record/3928368/files/figure.png" pageId="25" pageNumber="26">21</figureCitation>
).................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="33535BA1FF934A15FF6BFE7CBB3EFDB3" pageId="25" pageNumber="26">
<paragraph id="88184304FF934A15FF6BFE7CBB3EFE1B" blockId="25.[189,1399,440,598]" pageId="25" pageNumber="26">
<keyLead id="3356E031FF934A15FF6BFE7CBB3EFE1B" pageId="25" pageNumber="26">
2. Nodus with four primary dentations (
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: fig. 450), not deeply constricted at base ......... ..................................................................................................................
<taxonomicName id="4FA73887FF934A15FBEEFE20BB3EFE1B" ID-CoL="B8RZ" authority="(Verhoeff, 1941)" baseAuthorityName="Verhoeff" baseAuthorityYear="1941" box="[1080,1399,483,510]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="harrisi">
<emphasis id="BAD39F16FF934A15FBEEFE20BAE4FE1B" bold="true" box="[1080,1197,483,510]" italics="true" pageId="25" pageNumber="26">A. harrisi</emphasis>
(Verhoeff, 1941)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="88184304FF934A15FF17FDD4BB3EFDB3" blockId="25.[189,1399,440,598]" pageId="25" pageNumber="26">
<keyLead id="3356E031FF934A15FF17FDD4BB3EFDB3" pageId="25" pageNumber="26">
– Nodus with three primary dentations (
<figureCitation id="109C5F81FF934A15FD02FDCBBD7AFDCF" box="[724,819,527,554]" captionStart="Fig" captionStartId="29.[190,233,1929,1955]" captionTargetBox="[349,1236,266,1886]" captionTargetId="figure@29.[347,1237,265,1887]" captionTargetPageId="29" captionText="Fig. 19. Agrophogonus mwanihana Hoffman, 2005, specimen from Udzungwa Mts National Park, Sanje Kati camp &amp; plot (NHMD 621682), right gonopod. A–B. Mesal view. C–D. Lateral view. E–F. Dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928364" httpUri="https://zenodo.org/record/3928364/files/figure.png" pageId="25" pageNumber="26">Fig. 19</figureCitation>
;
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: fig. 449), basally with deep constriction .........................................................................................
<taxonomicName id="4FA73887FF934A15FBDDFDFFBB3EFDB3" ID-CoL="B8S4" authority="Hoffman, 2005" authorityName="Hoffmann" authorityYear="2005" box="[1035,1399,571,598]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="mwanihana">
<emphasis id="BAD39F16FF934A15FBDDFDFFBAF5FDB3" bold="true" box="[1035,1212,571,598]" italics="true" pageId="25" pageNumber="26">A. mwanihana</emphasis>
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</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="33535BA1FF934A15FF6BFD41BB3EFCC7" pageId="25" pageNumber="26">
<paragraph id="88184304FF934A15FF6BFD41BB3EFD2E" blockId="25.[189,1399,645,803]" pageId="25" pageNumber="26">
<keyLead id="3356E031FF934A15FF6BFD41BB3EFD2E" pageId="25" pageNumber="26">
3. Nodal process A broad, apically tridentate; ectal nodal surface distinctly convex; processes B and C robust, subequal in size and shape (
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: fig. 453).....................
<taxonomicName id="4FA73887FF934A15FB91FD75BB3EFD2E" ID-CoL="B8S5" authority="Hoffman, 2005" authorityName="Hoffman" authorityYear="2005" box="[1095,1399,688,715]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="BAD39F16FF934A15FB91FD75BAF5FD2E" bold="true" box="[1095,1212,688,715]" italics="true" pageId="25" pageNumber="26">A. tridens</emphasis>
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</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="88184304FF934A15FF17FD19BB3EFCC7" blockId="25.[189,1399,645,803]" pageId="25" pageNumber="26">
<keyLead id="3356E031FF934A15FF17FD19BB3EFCC7" pageId="25" pageNumber="26">
– Nodal process A acute and spiniform; ectal nodal surface flattened (in mesal aspect,
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: fig. 451).............................................................................................................................................. 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="33535BA1FF934A15FF6BFC96BB3EFC21" pageId="25" pageNumber="26">
<paragraph id="88184304FF934A15FF6BFC96BB3EFC7D" blockId="25.[189,1399,850,964]" pageId="25" pageNumber="26">
<keyLead id="3356E031FF934A15FF6BFC96BB3EFC7D" pageId="25" pageNumber="26">
4. Nodal process A with a small, bifid accessory process at base on lateral side (
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: figs 456–457, see remark to
<taxonomicName id="4FA73887FF934A15FDE1FCBABCA9FC72" ID-CoL="B8S4" authorityName="Hoffmann" authorityYear="2005" box="[567,736,893,920]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="mwanihana">
<emphasis id="BAD39F16FF934A15FDE1FCBABCA9FC72" box="[567,736,893,920]" italics="true" pageId="25" pageNumber="26">A. mwanihana</emphasis>
</taxonomicName>
, below)
<emphasis id="BAD39F16FF934A15FC94FCBABA5EFC72" box="[834,1047,894,919]" italics="true" pageId="25" pageNumber="26">.............................</emphasis>
<taxonomicName id="4FA73887FF934A15FBCBFCB9BB3EFC7D" ID-CoL="B8S3" authority="Hoffman, 2005" authorityName="Hoffman" authorityYear="2005" box="[1053,1399,893,920]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="mangalisa">
<emphasis id="BAD39F16FF934A15FBCBFCB9BAF4FC7D" bold="true" box="[1053,1213,893,920]" italics="true" pageId="25" pageNumber="26">A. mangalisa</emphasis>
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</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="88184304FF934A15FF17FC6EBB3EFC21" blockId="25.[189,1399,850,964]" box="[193,1399,937,964]" pageId="25" pageNumber="26">
<keyLead id="3356E031FF934A15FF17FC6EBB3EFC21" box="[193,1399,937,964]" pageId="25" pageNumber="26">– Nodal process A without an accessory basal process on lateral side ................................................. 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="33535BA1FF934A15FF6BFC37BB3EFB74" pageId="25" pageNumber="26">
<paragraph id="88184304FF934A15FF6BFC37BB3EFBDC" blockId="25.[189,1399,1011,1169]" pageId="25" pageNumber="26">
<keyLead id="3356E031FF934A15FF6BFC37BB3EFBDC" pageId="25" pageNumber="26">5. Postnodal telepodite relatively short, curved abruptly dorsad, almost perpendicular to prefemoral region ................................................................................................................................................. 6</keyLead>
</paragraph>
<paragraph id="88184304FF934A15FF6BFB8FBB3EFB74" blockId="25.[189,1399,1011,1169]" pageId="25" pageNumber="26">
<keyLead id="3356E031FF934A15FF6BFB8FBB3EFB74" pageId="25" pageNumber="26">
– Postnodal telepodite long, evenly curved dorsad, appearing hook-like (
<figureCitation id="109C5F81FF934A15FBFEFB8EBA34FB80" box="[1064,1149,1098,1125]" captionStart="Fig" captionStartId="26.[189,232,1929,1955]" captionTargetBox="[317,1259,268,1898]" captionTargetId="figure@26.[317,1268,268,1898]" captionTargetPageId="26" captionText="Fig. 18. Agrophogonus hamulifer Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621678), left gonopod. A–B. Mesal view. C–D. Lateral view. E–F. Dorsal view. Abbreviations: A = Process A; B = Process B; C = Process C; L = Process L. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928362" httpUri="https://zenodo.org/record/3928362/files/figure.png" pageId="25" pageNumber="26">Fig. 18</figureCitation>
) ............................... ...............................................................................................
<taxonomicName id="4FA73887FF934A15FC7FFBB2BB5CFB75" ID-CoL="9f1516ed-177e-4f9a-b6fc-d2c3dc1f0513" authority="Olsen &amp; Enghoff" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[937,1301,1142,1169]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="hamulifer" status="sp. nov.">
<emphasis id="BAD39F16FF934A15FC7FFBB2BA0DFB74" bold="true" box="[937,1092,1142,1169]" italics="true" pageId="25" pageNumber="26">A. hamulifer</emphasis>
Olsen &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF934A15FACAFBB3BB3EFB74" box="[1308,1399,1143,1169]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="33535BA1FF934A15FF6BFB04BB3EFABB" pageId="25" pageNumber="26">
<paragraph id="88184304FF934A15FF6BFB04BB3EFAE2" blockId="25.[189,1399,1216,1374]" pageId="25" pageNumber="26">
<keyLead id="3356E031FF934A15FF6BFB04BB3EFAE2" pageId="25" pageNumber="26">
6. Width
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. Nodal process C very long, slender and curved (
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: fig. 451) ................ .................................................................................................................
<taxonomicName id="4FA73887FF934A15FBE6FB28BB3EFAE2" ID-CoL="B8S2" authority="Hoffman, 2005" authorityName="Hoffman" authorityYear="2005" box="[1072,1399,1259,1287]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kiellandi">
<emphasis id="BAD39F16FF934A15FBE6FB28BAF5FAE3" bold="true" box="[1072,1212,1259,1286]" italics="true" pageId="25" pageNumber="26">A. kiellandi</emphasis>
<bibRefCitation id="EC363EF5FF934A15FB12FB28BB3EFAE2" author="Hoffman R. L." box="[1220,1399,1260,1287]" pageId="25" pageNumber="26" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman, 2005</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="88184304FF934A15FF6BFADCBB3EFABB" blockId="25.[189,1399,1216,1374]" pageId="25" pageNumber="26">
<keyLead id="3356E031FF934A15FF6BFADCBB3EFABB" pageId="25" pageNumber="26">
– Width
<quantity id="4F5FEEE1FF934A15FEECFADCBFDDFAD7" box="[314,404,1304,1330]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="25" pageNumber="26" unit="mm" value="4.1">4.1 mm</quantity>
(
<figureCitation id="109C5F81FF934A15FE75FAD3BC65FAD7" box="[419,556,1303,1330]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="29.[190,233,1929,1955]" captionStartId-1="30.[189,232,1965,1991]" captionTargetBox-0="[349,1236,266,1886]" captionTargetBox-1="[418,1171,1411,1922]" captionTargetId-0="figure@29.[347,1237,265,1887]" captionTargetId-1="figure@30.[418,1171,1411,1922]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionText-0="Fig. 19. Agrophogonus mwanihana Hoffman, 2005, specimen from Udzungwa Mts National Park, Sanje Kati camp &amp; plot (NHMD 621682), right gonopod. A–B. Mesal view. C–D. Lateral view. E–F. Dorsal view. Scale bars: 0.5 mm." captionText-1="Fig. 20. Agrophogonus pusillokiellandi Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621683). Scale bar: 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3928364" figureDoi-1="http://doi.org/10.5281/zenodo.3928366" httpUri-0="https://zenodo.org/record/3928364/files/figure.png" httpUri-1="https://zenodo.org/record/3928366/files/figure.png" pageId="25" pageNumber="26">Figs 19–20</figureCitation>
). Nodal process C shorter and less curved (
<figureCitation id="109C5F81FF934A15FBF1FAD3BA35FAD7" box="[1063,1148,1303,1330]" captionStart="Fig" captionStartId="31.[189,232,1895,1921]" captionTargetBox="[236,1337,265,1863]" captionTargetId="figure@31.[234,1350,265,1863]" captionTargetPageId="31" captionText="Fig. 21. Agrophogonus pusillokiellandi Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621683). A–D, F–G. Left gonopod. A–B. Mesal view. C–D. Lateral view. F–G. Dorsal view. E. Right gonopod, lateral view. H. Sternal process on ring 6. Abbreviations: A = process A; B = process B; C = process C; L = process L. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928368" httpUri="https://zenodo.org/record/3928368/files/figure.png" pageId="25" pageNumber="26">Fig. 21</figureCitation>
)................................ .....................................................................................
<taxonomicName id="4FA73887FF934A15FCB0FA87BB5CFAB8" ID-CoL="304e4a5f-84a3-48a8-a050-0ffa499f6690" authority="Olsen &amp; Enghoff" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[870,1301,1347,1373]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pusillokiellandi" status="sp. nov.">
<emphasis id="BAD39F16FF934A15FCB0FA87BA0DFAB8" bold="true" box="[870,1092,1347,1373]" italics="true" pageId="25" pageNumber="26">A. pusillokiellandi</emphasis>
Olsen &amp; Enghoff
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<taxonomicNameLabel id="A1E0226DFF934A15FACAFA80BB3EFABB" box="[1308,1399,1348,1374]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>

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<mods:title id="999FF8B79B5FA0BADDF9FEF36AA22F7E">A mountain of millipedes IX: Species of the family Gomphodesmidae from the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida)</mods:title>
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<mods:namePart id="1CBD5192A9DA1C950C7513C7F7C4FE7A">Olsen, Sissel Anna</mods:namePart>
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<heading id="D350F468FF934A15FE31FA67BA04FA5B" box="[487,1101,1443,1470]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="26" reason="2">
<taxonomicName id="4FA73887FF934A15FE31FA67BDA2FA5B" ID-CoL="9f1516ed-177e-4f9a-b6fc-d2c3dc1f0513" authority="Olsen &amp; Enghoff" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[487,1003,1443,1470]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="hamulifer" status="sp. nov.">
<emphasis id="BAD39F16FF934A15FE31FA67BD52FA5B" bold="true" box="[487,795,1443,1470]" italics="true" pageId="25" pageNumber="26">Agrophogonus hamulifer</emphasis>
Olsen &amp; Enghoff
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<figureCitation id="109C5F81FF934A15FD11FA3FBD59F9F0" box="[711,784,1531,1557]" captionStart="Fig" captionStartId="1.[189,232,1966,1992]" captionTargetBox="[361,1222,1253,1923]" captionTargetId="figure@1.[355,1226,1247,1927]" captionTargetPageId="1" captionText="Fig. 1. Agrophogonus hamulifer Olsen &amp; Enghoff sp. nov., paratype, ♂ (NHMD 621681), showing the typical gomphodesmid habitus. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3928328" httpUri="https://zenodo.org/record/3928328/files/figure.png" pageId="25" pageNumber="26">Figs 1</figureCitation>
,
<figureCitation id="109C5F81FF934A15FCC8FA3FBD0AF9F3" box="[798,835,1531,1558]" captionStart="Fig" captionStartId="6.[189,232,1500,1526]" captionTargetBox="[217,1367,909,1445]" captionTargetId="figure@6.[207,1377,905,1453]" captionTargetPageId="6" captionText="Fig. 5. Sternal process on body ring 6. A. Emphysemastix spp. B. Agrophogonus spp. except A. hamulifer Olsen &amp; Enghoff sp. nov. C. Agrophogonus hamulifer Olsen &amp; Enghoff sp. nov. D. Pogoropsis prolixopes Rosenmejer &amp; Enghoff gen. et sp. nov. E. Pogoro siren Rosenmejer &amp; Enghoff sp. nov. F. Pogoro scharffi Hoffman, 2005. G. Pogoro alopias Rosenmejer &amp; Enghoff sp. nov. Not drawn to scale." figureDoi="http://doi.org/10.5281/zenodo.3928336" httpUri="https://zenodo.org/record/3928336/files/figure.png" pageId="25" pageNumber="26">5C</figureCitation>
,
<figureCitation id="109C5F81FF934A15FC99FA3FBD25F9F0" box="[847,876,1531,1557]" captionStart="Fig" captionStartId="26.[189,232,1929,1955]" captionTargetBox="[317,1259,268,1898]" captionTargetId="figure@26.[317,1268,268,1898]" captionTargetPageId="26" captionText="Fig. 18. Agrophogonus hamulifer Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621678), left gonopod. A–B. Mesal view. C–D. Lateral view. E–F. Dorsal view. Abbreviations: A = Process A; B = Process B; C = Process C; L = Process L. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928362" httpUri="https://zenodo.org/record/3928362/files/figure.png" pageId="25" pageNumber="26">18</figureCitation>
</paragraph>
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<subSubSection id="C0BD108FFF934A15FF6BF981BC46F8A0" pageId="25" pageNumber="26" type="diagnosis">
<paragraph id="88184304FF934A15FF6BF981BF7FF9BA" blockId="25.[189,310,1605,1631]" box="[189,310,1605,1631]" pageId="25" pageNumber="26">
<heading id="D350F468FF934A15FF6BF981BF7FF9BA" bold="true" box="[189,310,1605,1631]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="BAD39F16FF934A15FF6BF981BF7FF9BA" bold="true" box="[189,310,1605,1631]" pageId="25" pageNumber="26">Diagnosis</emphasis>
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</paragraph>
<paragraph id="88184304FF934A15FF6BF9B8BC46F8A0" blockId="25.[189,1399,1660,1861]" pageId="25" pageNumber="26">
Resembles
<taxonomicName id="4FA73887FF934A15FE93F9B8BF84F973" authorityName="Hoffman" authorityYear="2005" box="[325,461,1660,1686]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kiellandi">
<emphasis id="BAD39F16FF934A15FE93F9B8BF84F973" box="[325,461,1660,1686]" italics="true" pageId="25" pageNumber="26">A. kiellandi</emphasis>
</taxonomicName>
and
<taxonomicName id="4FA73887FF934A15FDDEF9B8BCA8F973" authority="Olsen &amp; Rosenmejer &amp; Enghoff, 2020" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[520,737,1660,1686]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pusillokiellandi" status="sp. nov.">
<emphasis id="BAD39F16FF934A15FDDEF9B8BCA8F973" box="[520,737,1660,1686]" italics="true" pageId="25" pageNumber="26">A. pusillokiellandi</emphasis>
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in having the postnodal telopodite slender, and having nodal process A spiniform, without an accessory basal process. Differs from these species in having the postnodal telopodite very long and evenly curved. Differs further from
<taxonomicName id="4FA73887FF934A15FBA7F910BABEF908" authorityName="Hoffman" authorityYear="2005" box="[1137,1271,1747,1774]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kiellandi">
<emphasis id="BAD39F16FF934A15FBA7F910BABEF908" box="[1137,1271,1747,1774]" italics="true" pageId="25" pageNumber="26">A. kiellandi</emphasis>
</taxonomicName>
by smaller body size (width
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vs
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in
<taxonomicName id="4FA73887FF934A15FD19F93BBD10F8FC" authorityName="Hoffman" authorityYear="2005" box="[719,857,1791,1817]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kiellandi">
<emphasis id="BAD39F16FF934A15FD19F93BBD10F8FC" box="[719,857,1791,1817]" italics="true" pageId="25" pageNumber="26">A. kiellandi</emphasis>
</taxonomicName>
) and from
<taxonomicName id="4FA73887FF934A15FC34F93BBAF2F8FC" authority="Olsen &amp; Rosenmejer &amp; Enghoff, 2020" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[994,1211,1791,1817]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pusillokiellandi" status="sp. nov.">
<emphasis id="BAD39F16FF934A15FC34F93BBAF2F8FC" box="[994,1211,1791,1817]" italics="true" pageId="25" pageNumber="26">A. pusillokiellandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF934A15FB12F8C4BB68F8FF" box="[1220,1313,1792,1818]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
(width
<quantity id="4F5FEEE1FF934A15FF6BF8EFBF50F8A0" box="[189,281,1835,1861]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="25" pageNumber="26" unit="mm" value="4.1">4.1 mm</quantity>
) by larger body size.
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF934A15FF6BF8B7BDC3F80E" pageId="25" pageNumber="26" type="etymology">
<paragraph id="88184304FF934A15FF6BF8B7BF0DF868" blockId="25.[189,324,1907,1933]" box="[189,324,1907,1933]" pageId="25" pageNumber="26">
<heading id="D350F468FF934A15FF6BF8B7BF0DF868" bold="true" box="[189,324,1907,1933]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="BAD39F16FF934A15FF6BF8B7BF0DF868" bold="true" box="[189,324,1907,1933]" pageId="25" pageNumber="26">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF934A15FF6BF863BDC3F80E" blockId="25.[189,1399,1959,2027]" pageId="25" pageNumber="26">The name is a Latin noun in apposition, meaning ‘carrying a small hook’ and refers to the resemblance of the postnodal telepodite to the curvature of a fishing hook.</paragraph>
</subSubSection>
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<paragraph id="88184304FF904A16FF6BF84DBD44F80F" blockId="26.[189,1399,1929,2026]" pageId="26" pageNumber="27">
<emphasis id="BAD39F16FF904A16FF6BF84DBF51F846" bold="true" box="[189,280,1929,1955]" pageId="26" pageNumber="27">Fig. 18.</emphasis>
<taxonomicName id="4FA73887FF904A16FECBF84DBD45F846" authority="Olsen &amp; Enghoff" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[285,780,1929,1955]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="hamulifer" status="sp. nov.">
<emphasis id="BAD39F16FF904A16FECBF84DBC09F846" box="[285,576,1929,1955]" italics="true" pageId="26" pageNumber="27">Agrophogonus hamulifer</emphasis>
Olsen &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF904A16FCC4F84EBD24F841" box="[786,877,1930,1956]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NHMD 621678), left gonopod.
<emphasis id="BAD39F16FF904A16FF6BF868BEBDF823" bold="true" box="[189,244,1964,1990]" pageId="26" pageNumber="27">A–B</emphasis>
. Mesal view.
<emphasis id="BAD39F16FF904A16FE4EF868BF98F823" bold="true" box="[408,465,1964,1990]" pageId="26" pageNumber="27">C–D</emphasis>
. Lateral view.
<emphasis id="BAD39F16FF904A16FDA9F869BCFAF822" bold="true" box="[639,691,1965,1991]" pageId="26" pageNumber="27">E–F</emphasis>
. Dorsal view. Abbreviations:
<emphasis id="BAD39F16FF904A16FBC7F869BA74F823" box="[1041,1085,1965,1991]" italics="true" pageId="26" pageNumber="27">A =</emphasis>
Process A;
<emphasis id="BAD39F16FF904A16FB1FF869BA92F823" box="[1225,1243,1965,1990]" italics="true" pageId="26" pageNumber="27">B</emphasis>
= Process B;
<emphasis id="BAD39F16FF904A16FF6BF814BE98F80F" box="[189,209,2000,2026]" italics="true" pageId="26" pageNumber="27">C</emphasis>
= Process C;
<emphasis id="BAD39F16FF904A16FEA2F814BFCDF80C" box="[372,388,2000,2025]" italics="true" pageId="26" pageNumber="27">L</emphasis>
= Process L. Scale bars: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C0BD108FFF914A17FF6BFECFBFBBFD78" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="88184304FF914A17FF6BFECFBFE3FEC0" blockId="27.[189,426,267,293]" box="[189,426,267,293]" pageId="27" pageNumber="28">
<heading id="D350F468FF914A17FF6BFECFBFE3FEC0" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="BAD39F16FF914A17FF6BFECFBFE3FEC0" bold="true" box="[189,426,267,293]" pageId="27" pageNumber="28">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF914A17FF3CFEF8BF15FEB3" blockId="27.[189,1399,316,453]" box="[234,348,316,342]" pageId="27" pageNumber="28">
<emphasis id="BAD39F16FF914A17FF3CFEF8BF15FEB3" bold="true" box="[234,348,316,342]" pageId="27" pageNumber="28">
<typeStatus id="571CFDA6FF914A17FF3CFEF8BF15FEB3" box="[234,348,316,342]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="88184304FF914A17FF6BFEA5BF54FE20" blockId="27.[189,1399,316,453]" pageId="27" pageNumber="28">
<materialsCitation id="38CF4959FF914A17FF6BFEA5BF54FE20" ID-GBIF-Occurrence="2803530317" collectedFrom="Ukami, montane forest" collectingDate="2000-11-25" collectionCode="NHMD" collectorName="Frontier Tanzania UMBS" country="Tanzania" elevation="1115" latitude="-7.88875" location="Iringa Region" longLatPrecision="1" longitude="36.38656" pageId="27" pageNumber="28" specimenCode="NHMD 621678" specimenCount="1" stateProvince="Iringa Region" typeStatus="holotype">
<collectingCountry id="F0B00394FF914A17FF6BFEA5BF18FE9E" box="[189,337,353,379]" name="Tanzania" pageId="27" pageNumber="28">TANZANIA</collectingCountry>
•
<specimenCount id="9EA1888DFF914A17FEA5FEA5BFC2FE9E" box="[371,395,353,379]" pageId="27" pageNumber="28" type="male">♂</specimenCount>
;
<collectingRegion id="4A638DE6FF914A17FE4BFEA5BC0DFE9E" box="[413,580,353,379]" country="Tanzania" name="Iringa" pageId="27" pageNumber="28">Iringa Region</collectingRegion>
/district,
<locationDeviation id="99AA5A26FF914A17FD67FEA5BDB1FE9E" box="[689,1016,353,379]" pageId="27" pageNumber="28">West Kilombero Scarp FR</locationDeviation>
;
<geoCoordinate id="ED9325C3FF914A17FBDFFE9BBAFAFE9E" box="[1033,1203,351,379]" degrees="07" direction="south" minutes="53" orientation="latitude" pageId="27" pageNumber="28" precision="1" seconds="19.5" value="-7.88875">07°53′19.5″ S</geoCoordinate>
,
<geoCoordinate id="ED9325C3FF914A17FB13FE9BBB38FE9E" box="[1221,1393,351,379]" degrees="36" direction="east" minutes="23" orientation="longitude" pageId="27" pageNumber="28" precision="1" seconds="11.6" value="36.38656">36°23′11.6″ E</geoCoordinate>
;
<elevation id="038AA437FF914A17FF6BFE42BF11FE45" box="[189,344,390,417]" metricMagnitude="3" metricUnit="m" metricValue="1.115" pageId="27" pageNumber="28" unit="m" value="1115.0">
<quantity id="4F5FEEE1FF914A17FF6BFE42BF5FFE45" box="[189,278,390,417]" metricMagnitude="3" metricUnit="m" metricValue="1.115" pageId="27" pageNumber="28" unit="m" value="1115.0">1115 m</quantity>
a.s.l.
</elevation>
;
<date id="FC1965C4FF914A17FEBEFE42BC46FE45" box="[360,527,390,417]" pageId="27" pageNumber="28" value="2000-11-25">
<collectingDate id="EC5D9C2CFF914A17FEBEFE42BC46FE45" box="[360,527,390,417]" pageId="27" pageNumber="28" value="2000-11-25">25 Nov. 2000</collectingDate>
</date>
;
<collectorName id="255226D2FF914A17FDC8FE42BD05FE45" box="[542,844,390,416]" pageId="27" pageNumber="28">Frontier Tanzania UMBS</collectorName>
leg.; plot 17,
<collectedFrom id="3BBB00F9FF914A17FC2EFE42BB46FE45" box="[1016,1295,390,416]" pageId="27" pageNumber="28">Ukami, montane forest</collectedFrom>
;
<specimenCode id="D801EB7FFF914A17FACBFE42BF50FE20" collectionCode="NHMD" pageId="27" pageNumber="28">NHMD 621678</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="88184304FF914A17FF3CFE2BBF2EFDEC" blockId="27.[189,1399,495,669]" box="[234,359,495,521]" pageId="27" pageNumber="28">
<emphasis id="BAD39F16FF914A17FF3CFE2BBF2EFDEC" bold="true" box="[234,359,495,521]" pageId="27" pageNumber="28">
<typeStatus id="571CFDA6FF914A17FF3CFE2BBF2EFDEC" box="[234,359,495,521]" pageId="27" pageNumber="28" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="88184304FF914A17FF6BFDD0BFBBFD78" blockId="27.[189,1399,495,669]" pageId="27" pageNumber="28">
<materialsCitation id="38CF4959FF914A17FF6BFDD0BB42FDC8" ID-GBIF-Occurrence="2803530312" box="[189,1291,531,558]" collectingDate="2000-11-25" collectionCode="NHMD" collectorName="West Kilombero Scarp FR" country="Tanzania" elevation="1115" latitude="-7.88875" location="TANZANIA" longLatPrecision="1" longitude="36.38656" pageId="27" pageNumber="28" specimenCode="NHMD 621679" specimenCount="47" specimenCount-female="7" specimenCount-male="11" stateProvince="Iringa" typeStatus="paratype">
<collectingCountry id="F0B00394FF914A17FF6BFDD0BF18FDCB" box="[189,337,532,558]" name="Tanzania" pageId="27" pageNumber="28">TANZANIA</collectingCountry>
•
<specimenCount id="9EA1888DFF914A17FEB3FDD0BFF3FDCB" box="[357,442,532,558]" pageId="27" pageNumber="28" type="male">11 ♂♂</specimenCount>
,
<specimenCount id="9EA1888DFF914A17FE1CFDD0BC59FDCB" box="[458,528,532,558]" pageId="27" pageNumber="28" type="female">7 ♀♀</specimenCount>
,
<specimenCount id="9EA1888DFF914A17FDF7FDD0BC3EFDC8" box="[545,631,531,558]" pageId="27" pageNumber="28" type="generic">29 juv.</specimenCount>
; same collecting data as for holotype;
<specimenCode id="D801EB7FFF914A17FB99FDD0BB42FDC8" box="[1103,1291,531,558]" collectionCode="NHMD" pageId="27" pageNumber="28">NHMD 621679</specimenCode>
</materialsCitation>
•
<materialsCitation id="38CF4959FF914A17FAFCFDD0BB6EFDB7" ID-GBIF-Occurrence="2803530311" collectingDate="2000-11-30" collectionCode="NHMD" collectorName="West Kilombero Scarp FR" country="Tanzania" elevation="1115" latitude="-7.88875" location="West Kilombero Scarp" longLatPrecision="1" longitude="36.38656" pageId="27" pageNumber="28" specimenCode="NHMD 621680" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Iringa" typeStatus="paratype">
<specimenCount id="9EA1888DFF914A17FAFCFDD0BB38FDCB" box="[1322,1393,532,558]" pageId="27" pageNumber="28" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9EA1888DFF914A17FF6BFDFDBF48FDB6" box="[189,257,569,595]" pageId="27" pageNumber="28" type="female">2 ♀♀</specimenCount>
; same collecting data as for holotype;
<elevation id="038AA437FF914A17FD1AFDFDBDE7FDB7" box="[716,942,568,595]" metricMagnitude="3" metricUnit="m" metricValue="1.115" metricValueMax="1.13" metricValueMin="1.1" pageId="27" pageNumber="28" unit="m" value="1115.0" valueMax="1130.0" valueMin="1100.0">
<quantity id="4F5FEEE1FF914A17FD1AFDFDBD24FDB6" box="[716,877,569,595]" metricMagnitude="3" metricUnit="m" metricValue="1.115" metricValueMax="1.13" metricValueMin="1.1" pageId="27" pageNumber="28" unit="m" value="1115.0" valueMax="1130.0" valueMin="1100.0">1100–1130 m</quantity>
a.s.l.
</elevation>
;
<date id="FC1965C4FF914A17FC6AFDFDBA29FDB6" box="[956,1120,569,595]" pageId="27" pageNumber="28" value="2000-11-30">
<collectingDate id="EC5D9C2CFF914A17FC6AFDFDBA29FDB6" box="[956,1120,569,595]" pageId="27" pageNumber="28" value="2000-11-30">30 Nov. 2000</collectingDate>
</date>
;
<specimenCode id="D801EB7FFF914A17FBBBFDFDBB6EFDB7" box="[1133,1319,568,595]" collectionCode="NHMD" pageId="27" pageNumber="28">NHMD 621680</specimenCode>
</materialsCitation>
•
<materialsCitation id="38CF4959FF914A17FA94FDFDBFA7FD78" ID-GBIF-Occurrence="2803530309" collectingDate="2000-11-25" collectionCode="NHMD" collectorName="West Kilombero Scarp FR" country="Tanzania" elevation="1400" latitude="-7.8440003" location="West Kilombero Scarp" longLatPrecision="1" longitude="36.371555" pageId="27" pageNumber="28" specimenCode="NHMD 621681" specimenCount="2" specimenCount-male="1" stateProvince="Iringa" typeStatus="paratype">
<specimenCount id="9EA1888DFF914A17FA94FDFDBB39FDB6" box="[1346,1392,569,595]" pageId="27" pageNumber="28" type="male">1 ♂</specimenCount>
,
<specimenCount id="9EA1888DFF914A17FF6BFD9ABF4BFD9D" box="[189,258,606,632]" pageId="27" pageNumber="28" type="generic">1 juv.</specimenCount>
; same collecting data as for holotype but
<geoCoordinate id="ED9325C3FF914A17FD3BFD98BDDDFD9D" box="[749,916,604,632]" degrees="07" direction="south" minutes="50" orientation="latitude" pageId="27" pageNumber="28" precision="1" seconds="38.4" value="-7.8440003">07°50′38.4″ S</geoCoordinate>
,
<geoCoordinate id="ED9325C3FF914A17FC77FD98BA02FD9D" box="[929,1099,604,632]" degrees="36" direction="east" minutes="22" orientation="longitude" pageId="27" pageNumber="28" precision="1" seconds="17.6" value="36.371555">36°22′17.6″ E</geoCoordinate>
;
<elevation id="038AA437FF914A17FB8EFD9ABB72FD9D" box="[1112,1339,606,632]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.41" metricValueMin="1.39" pageId="27" pageNumber="28" unit="m" value="1400.0" valueMax="1410.0" valueMin="1390.0">
<quantity id="4F5FEEE1FF914A17FB8EFD9ABAB3FD9D" box="[1112,1274,606,632]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.41" metricValueMin="1.39" pageId="27" pageNumber="28" unit="m" value="1400.0" valueMax="1410.0" valueMin="1390.0">1390–1410 m</quantity>
a.s.l.
</elevation>
; plot Paradiso;
<specimenCode id="D801EB7FFF914A17FEE7FD47BFA7FD78" box="[305,494,643,669]" collectionCode="NHMD" pageId="27" pageNumber="28">NHMD 621681</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF914A17FF6BFD02BF42FA27" pageId="27" pageNumber="28" type="description">
<paragraph id="88184304FF914A17FF6BFD02BF07FD05" blockId="27.[189,334,710,736]" box="[189,334,710,736]" pageId="27" pageNumber="28">
<heading id="D350F468FF914A17FF6BFD02BF07FD05" bold="true" box="[189,334,710,736]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="BAD39F16FF914A17FF6BFD02BF07FD05" bold="true" box="[189,334,710,736]" pageId="27" pageNumber="28">Description</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF914A17FF3CFD33BF7CFCF4" blockId="27.[189,1223,759,823]" box="[234,309,759,785]" pageId="27" pageNumber="28">
<emphasis id="BAD39F16FF914A17FF3CFD33BF7CFCF4" bold="true" box="[234,309,759,785]" pageId="27" pageNumber="28">Males</emphasis>
</paragraph>
<paragraph id="88184304FF914A17FF6BFCD8BA8EFCD3" blockId="27.[189,1223,759,823]" box="[189,1223,796,823]" pageId="27" pageNumber="28">
SIZE AND SHAPE. Body length
<quantity id="4F5FEEE1FF914A17FDCBFCD8BCD6FCD3" box="[541,671,796,822]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" metricValueMax="3.1" metricValueMin="2.7" pageId="27" pageNumber="28" unit="mm" value="29.0" valueMax="31.0" valueMin="27.0">27–31 mm</quantity>
. Maximum width
<quantity id="4F5FEEE1FF914A17FCA0FCD8BA4FFCD3" box="[886,1030,796,823]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" metricValueMax="6.2" metricValueMin="5.2" pageId="27" pageNumber="28" unit="mm" value="5.7" valueMax="6.2" valueMin="5.2">5.2–6.2 mm</quantity>
. W/L ratio 20%.
</paragraph>
<paragraph id="88184304FF914A17FF6BFC98BA48FC7F" blockId="27.[189,1398,859,923]" pageId="27" pageNumber="28">
COLOUR (
<figureCitation id="109C5F81FF914A17FEE6FC9FBF3AFC93" box="[304,371,859,886]" captionStart="Fig" captionStartId="1.[189,232,1966,1992]" captionTargetBox="[361,1222,1253,1923]" captionTargetId="figure@1.[355,1226,1247,1927]" captionTargetPageId="1" captionText="Fig. 1. Agrophogonus hamulifer Olsen &amp; Enghoff sp. nov., paratype, ♂ (NHMD 621681), showing the typical gomphodesmid habitus. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3928328" httpUri="https://zenodo.org/record/3928328/files/figure.png" pageId="27" pageNumber="28">Fig. 1</figureCitation>
). Very pale brown, darker around base of setae. Specimen stored in ethanol since 2000; thus, any pigmentation that may have been present in life is now faded.
</paragraph>
<paragraph id="88184304FF914A17FF6BFC04BFE3FC1A" blockId="27.[189,1399,960,1023]" pageId="27" pageNumber="28">
STERNAL PROCESS ON BODY RING 6 (
<figureCitation id="109C5F81FF914A17FD84FC04BCE1FC3E" box="[594,680,960,987]" captionStart="Fig" captionStartId="6.[189,232,1500,1526]" captionTargetBox="[217,1367,909,1445]" captionTargetId="figure@6.[207,1377,905,1453]" captionTargetPageId="6" captionText="Fig. 5. Sternal process on body ring 6. A. Emphysemastix spp. B. Agrophogonus spp. except A. hamulifer Olsen &amp; Enghoff sp. nov. C. Agrophogonus hamulifer Olsen &amp; Enghoff sp. nov. D. Pogoropsis prolixopes Rosenmejer &amp; Enghoff gen. et sp. nov. E. Pogoro siren Rosenmejer &amp; Enghoff sp. nov. F. Pogoro scharffi Hoffman, 2005. G. Pogoro alopias Rosenmejer &amp; Enghoff sp. nov. Not drawn to scale." figureDoi="http://doi.org/10.5281/zenodo.3928336" httpUri="https://zenodo.org/record/3928336/files/figure.png" pageId="27" pageNumber="28">Fig. 5C</figureCitation>
). Long and slender, but broader towards basis and anterior end. Tip evenly rounded.
</paragraph>
<paragraph id="88184304FF914A17FF6BFBE1BCFFFBDA" blockId="27.[189,694,1061,1087]" box="[189,694,1061,1087]" pageId="27" pageNumber="28">BODY RINGS. No transverse sternal carinae.</paragraph>
<paragraph id="88184304FF914A17FF6BFBA1BDF2FB2C" blockId="27.[189,1399,1124,1225]" pageId="27" pageNumber="28">TELSON. Epiproct flattened at posterior end, forming a small vertical plane, with four protruding setae. Hypoproct semicircular to slightly triangular, paramedian tubercles large, extending beyond edge of sclerite. Setae on paraproct protruding from clear basal tubercles.</paragraph>
<paragraph id="88184304FF914A17FF6BFB2ABF42FA27" blockId="27.[189,1399,1262,1474]" pageId="27" pageNumber="28">
GONOPOD (
<figureCitation id="109C5F81FF914A17FE93FB2ABFD2FAED" box="[325,411,1262,1289]" captionStart="Fig" captionStartId="26.[189,232,1929,1955]" captionTargetBox="[317,1259,268,1898]" captionTargetId="figure@26.[317,1268,268,1898]" captionTargetPageId="26" captionText="Fig. 18. Agrophogonus hamulifer Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621678), left gonopod. A–B. Mesal view. C–D. Lateral view. E–F. Dorsal view. Abbreviations: A = Process A; B = Process B; C = Process C; L = Process L. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928362" httpUri="https://zenodo.org/record/3928362/files/figure.png" pageId="27" pageNumber="28">Fig. 18</figureCitation>
). Gonopod aperture suboval, extended back between second pair of legs on seventh segment, elevated in front of coxae, eliminated medially. Paracannular setae present. Nodal process L present, process M absent. Processes A and B slender and curved, process C longer than A and B. Processes B and C forcipulate in their curvature and position to each other. Postnodal telepodite very long, slender and acuminate, longer than in other members of the genus, without an abrupt flexure after nodus.
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF914A17FF6BFA2CBEBBF9BD" pageId="27" pageNumber="28" type="distribution">
<paragraph id="88184304FF914A17FF6BFA2CBF1FF9E7" blockId="27.[189,342,1512,1538]" box="[189,342,1512,1538]" pageId="27" pageNumber="28">
<heading id="D350F468FF914A17FF6BFA2CBF1FF9E7" bold="true" box="[189,342,1512,1538]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="BAD39F16FF914A17FF6BFA2CBF1FF9E7" bold="true" box="[189,342,1512,1538]" pageId="27" pageNumber="28">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF914A17FF6BF9DCBEBBF9BD" blockId="27.[189,1398,1560,1624]" pageId="27" pageNumber="28">Found at two separate locations within the same forest (Ukami) in the West Kilombero FR, Udzungwa Mts.</paragraph>
</subSubSection>
</treatment>
</document>

186
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<mods:title id="1F94D4F8251DEB95B8598C1C2CF7C4B9">A mountain of millipedes IX: Species of the family Gomphodesmidae from the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida)</mods:title>
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<mods:date id="21A0217642373B52F924AC18AAFB511F">2020</mods:date>
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<emphasis id="BAD39F16FF964A10FDCEFECFBD28FEC3" bold="true" box="[536,865,267,294]" italics="true" pageId="28" pageNumber="29">Agrophogonus mwanihana</emphasis>
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<emphasis id="BAD39F16FF964A10FF6BFEB6BFB0FE69" box="[189,505,370,396]" italics="true" pageId="28" pageNumber="29">Agrophogonus mwanihana</emphasis>
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<emphasis id="BAD39F16FF964A10FF6BFE70BFE3FE2B" bold="true" box="[189,426,436,462]" pageId="28" pageNumber="29">Material examined</emphasis>
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<materialsCitation id="38CF4959FF964A10FF6BFE20BF50FDA3" ID-GBIF-Occurrence="2803530307" collectingDate="2014-02-07" collectingMethod="pitfall traps" collectionCode="NHMD" collectorName="J. Malumbres-Olarte" country="Tanzania" elevation="850" latitude="-7.763222" location="Sanje Kati" longLatPrecision="1" longitude="36.886223" municipality="Udzungwa Mts National Park" pageId="28" pageNumber="29" specimenCode="NHMD 621682" specimenCount="13" specimenCount-female="5" stateProvince="Morogoro Region">
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•
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,
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;
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,
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camp and plot;
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,
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;
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a.s.l.
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leg.;
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;
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.
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<emphasis id="BAD39F16FF964A10FF6BFDAABF1FFD6D" bold="true" box="[189,342,622,648]" pageId="28" pageNumber="29">Distribution</emphasis>
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Known from the
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locality:
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,
<collectingRegion id="4A638DE6FF964A10FD30FD59BDD9FD52" box="[742,912,669,695]" country="Tanzania" name="Iringa" pageId="28" pageNumber="29">Iringa District</collectingRegion>
, Udzungwa Mts, Mwanihana FR,
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a.s.l. (corresponds to
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a.s.l.), Sanje
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valley, and now also from a very close site at a slightly lower elevation (
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).
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<emphasis id="BAD39F16FF964A10FF6BFCECBF66FCA7" bold="true" box="[189,303,808,834]" pageId="28" pageNumber="29">Remarks</emphasis>
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The new specimens agree with the description by
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. It should be noticed that the drawings on p.
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Hoffman’s book, although the caption states “Figs 454–455: Gonopods,
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<emphasis id="BAD39F16FF964A10FB1DFCBFBF0FFC5C" italics="true" pageId="28" pageNumber="29">Agrophogonus mwanihana</emphasis>
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….”, actually are figs 456–457 and show
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<emphasis id="BAD39F16FF964A10FCE2FC64BD87FC5C" box="[820,974,927,954]" italics="true" pageId="28" pageNumber="29">A. mangalisa</emphasis>
</taxonomicName>
.
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We here present new illustrations of the gonopod of
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<emphasis id="BAD39F16FF964A10FCFCFC26BD9DFC19" box="[810,980,994,1020]" italics="true" pageId="28" pageNumber="29">A. mwanihana</emphasis>
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, based on one of the newly studied specimens (
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).
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<mods:title id="BCC50CB4690844B7F74E9A42574A725F">A mountain of millipedes IX: Species of the family Gomphodesmidae from the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida)</mods:title>
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<mods:namePart id="04F5B68550CF16F1BFAD01978EDE108F">Olsen, Sissel Anna</mods:namePart>
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<mods:affiliation id="C780F6A235E344FC7C96A5FAF26B6BEE">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 København Ø, Denmark</mods:affiliation>
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<mods:namePart id="47EEE4B3E0E568A40E39A2D3964F93CD">Rosenmejer, Trine</mods:namePart>
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<mods:affiliation id="69C45938FD31079F2DADB2088488F062">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 København Ø, Denmark</mods:affiliation>
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<mods:namePart id="C2B5EC387CB0F38C951EC55FE6435564">Enghoff, Henrik</mods:namePart>
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<mods:affiliation id="6DCAEA1ADE3B0B23407C234255786E1E">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 København Ø, Denmark</mods:affiliation>
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<heading id="D350F468FF964A10FE13FB9ABA27FB9C" box="[453,1134,1118,1145]" centered="true" fontSize="11" level="2" pageId="28" pageNumber="29" reason="2">
<taxonomicName id="4FA73887FF964A10FE13FB9ABA44FB9C" ID-CoL="304e4a5f-84a3-48a8-a050-0ffa499f6690" authority="Olsen &amp; Enghoff" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[453,1037,1118,1145]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="pusillokiellandi" status="sp. nov.">
<emphasis id="BAD39F16FF964A10FE13FB9ABD72FB9C" bold="true" box="[453,827,1118,1145]" italics="true" pageId="28" pageNumber="29">Agrophogonus pusillokiellandi</emphasis>
Olsen &amp; Enghoff
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<figureCitation id="109C5F81FF964A10FD0EFB63BD12FB24" box="[728,859,1191,1217]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="30.[189,232,1965,1991]" captionStartId-1="31.[189,232,1895,1921]" captionTargetBox-0="[418,1171,1411,1922]" captionTargetBox-1="[236,1337,265,1863]" captionTargetId-0="figure@30.[418,1171,1411,1922]" captionTargetId-1="figure@31.[234,1350,265,1863]" captionTargetPageId-0="30" captionTargetPageId-1="31" captionText-0="Fig. 20. Agrophogonus pusillokiellandi Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621683). Scale bar: 5 mm." captionText-1="Fig. 21. Agrophogonus pusillokiellandi Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621683). A–D, F–G. Left gonopod. A–B. Mesal view. C–D. Lateral view. F–G. Dorsal view. E. Right gonopod, lateral view. H. Sternal process on ring 6. Abbreviations: A = process A; B = process B; C = process C; L = process L. Scale bars: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3928366" figureDoi-1="http://doi.org/10.5281/zenodo.3928368" httpUri-0="https://zenodo.org/record/3928366/files/figure.png" httpUri-1="https://zenodo.org/record/3928368/files/figure.png" pageId="28" pageNumber="29">Figs 20–21</figureCitation>
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<emphasis id="BAD39F16FF964A10FF6BFB2DBF7FFAE6" bold="true" box="[189,310,1257,1283]" pageId="28" pageNumber="29">Diagnosis</emphasis>
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Differs from the other species of the genus by the combination of small size (width
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), a slender postnodal telopodite, and lack of a bifid accessory process laterally on nodus. Virtually similar to
<taxonomicName id="4FA73887FF964A10FF6BFAA5BF0CFA9E" authorityName="Hoffman" authorityYear="2005" box="[189,325,1377,1403]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="kiellandi">
<emphasis id="BAD39F16FF964A10FF6BFAA5BF0CFA9E" box="[189,325,1377,1403]" italics="true" pageId="28" pageNumber="29">A. kiellandi</emphasis>
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except for size and for having process C of the gonopod telopodite shorter and broader.
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<emphasis id="BAD39F16FF964A10FF6BFA60BF0DFA5B" bold="true" box="[189,324,1444,1470]" pageId="28" pageNumber="29">Etymology</emphasis>
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The specific name is derived from the Latin ‘pusillus’, meaning ‘small’, and ‘kiellandi’, to emphasize the strong similarity with
<taxonomicName id="4FA73887FF964A10FE3AFA33BC3DF9F4" authorityName="Hoffman" authorityYear="2005" box="[492,628,1527,1553]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="kiellandi">
<emphasis id="BAD39F16FF964A10FE3AFA33BC3DF9F4" box="[492,628,1527,1553]" italics="true" pageId="28" pageNumber="29">A. kiellandi</emphasis>
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, except for size.
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<subSubSection id="C0BD108FFF964A10FF6BF9FDBAF3F8B0" pageId="28" pageNumber="29" type="materials_examined">
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<heading id="D350F468FF964A10FF6BF9FDBFE3F9B6" bold="true" box="[189,426,1593,1619]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="BAD39F16FF964A10FF6BF9FDBFE3F9B6" bold="true" box="[189,426,1593,1619]" pageId="28" pageNumber="29">Material examined</emphasis>
</heading>
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<emphasis id="BAD39F16FF964A10FF3CF9ADBF15F966" bold="true" box="[234,348,1641,1667]" pageId="28" pageNumber="29">
<typeStatus id="571CFDA6FF964A10FF3CF9ADBF15F966" box="[234,348,1641,1667]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
</emphasis>
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<materialsCitation id="38CF4959FF964A10FF6BF949BF37F90A" ID-GBIF-Occurrence="2803530318" collectingDate="2000-12-08" collectionCode="NHMD" collectorName="Frontier Tanzania" country="Tanzania" elevation="1510" latitude="-7.7595" location="West Kilombero Scarp Forest Reserve" longLatPrecision="1" longitude="36.44372" pageId="28" pageNumber="29" specimenCode="NHMD 621683" specimenCount="1" stateProvince="Iringa region" typeStatus="holotype">
<collectingCountry id="F0B00394FF964A10FF6BF949BF18F942" box="[189,337,1677,1703]" name="Tanzania" pageId="28" pageNumber="29">TANZANIA</collectingCountry>
•
<specimenCount id="9EA1888DFF964A10FEABF949BFDCF942" box="[381,405,1677,1703]" pageId="28" pageNumber="29" type="male">♂</specimenCount>
;
<collectingRegion id="4A638DE6FF964A10FE7AF948BC07F943" box="[428,590,1676,1702]" country="Tanzania" name="Iringa" pageId="28" pageNumber="29">Iringa region</collectingRegion>
/district,
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;
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,
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;
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<quantity id="4F5FEEE1FF964A10FEA9F974BF97F92E" box="[383,478,1712,1739]" metricMagnitude="3" metricUnit="m" metricValue="1.51" pageId="28" pageNumber="29" unit="m" value="1510.0">1510 m</quantity>
a.s.l.
</elevation>
;
<date id="FC1965C4FF964A10FDEEF975BC9EF92E" box="[568,727,1713,1739]" pageId="28" pageNumber="29" value="2000-12-08">
<collectingDate id="EC5D9C2CFF964A10FDEEF975BC9EF92E" box="[568,727,1713,1739]" pageId="28" pageNumber="29" value="2000-12-08">8 Dec. 2000</collectingDate>
</date>
;
<collectorName id="255226D2FF964A10FD3CF974BDF7F92F" box="[746,958,1712,1738]" pageId="28" pageNumber="29">Frontier Tanzania</collectorName>
leg.; plot
<taxonomicName id="4FA73887FF964A10FB9EF974BAD5F92F" box="[1096,1180,1712,1738]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="28" pageNumber="16" phylum="Tracheophyta" rank="genus">Acacia</taxonomicName>
, (open)woodland;
<specimenCode id="D801EB7FFF964A10FF6BF911BF33F90A" box="[189,378,1749,1775]" collectionCode="NHMD" pageId="28" pageNumber="29">NHMD 621683</specimenCode>
.
</materialsCitation>
</paragraph>
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<emphasis id="BAD39F16FF964A10FF3CF8D3BFEFF8D4" bold="true" box="[234,422,1815,1841]" pageId="28" pageNumber="29">Other material</emphasis>
</paragraph>
<paragraph id="88184304FF964A10FF6BF8FFBAF3F8B0" blockId="28.[189,1210,1815,1878]" box="[189,1210,1851,1878]" pageId="28" pageNumber="29">
<materialsCitation id="38CF4959FF964A10FF6BF8FFBAFFF8B0" ID-GBIF-Occurrence="2803530308" box="[189,1206,1851,1878]" collectingDate="2000-12-05" collectionCode="NHMD" collectorName="Frontier Tanzania" country="Tanzania" elevation="1510" latitude="-7.7595" location="TANZANIA" longLatPrecision="1" longitude="36.44372" pageId="28" pageNumber="29" specimenCode="NHMD 621684" specimenCount="1" specimenCount-female="1" stateProvince="Iringa">
<collectingCountry id="F0B00394FF964A10FF6BF8FFBF18F8B0" box="[189,337,1851,1877]" name="Tanzania" pageId="28" pageNumber="29">TANZANIA</collectingCountry>
•
<specimenCount id="9EA1888DFF964A10FEBFF8FFBFDFF8B3" box="[361,406,1851,1878]" pageId="28" pageNumber="29" type="female">1 ♀</specimenCount>
; same collecting data as for holotype;
<date id="FC1965C4FF964A10FC8DF8FFBDA4F8B0" box="[859,1005,1851,1878]" pageId="28" pageNumber="29" value="2000-12-05">
<collectingDate id="EC5D9C2CFF964A10FC8DF8FFBDA4F8B0" box="[859,1005,1851,1878]" pageId="28" pageNumber="29" value="2000-12-05">5 Dec. 2000</collectingDate>
</date>
;
<specimenCode id="D801EB7FFF964A10FC2FF8F8BAFFF8B0" box="[1017,1206,1851,1878]" collectionCode="NHMD" pageId="28" pageNumber="29">NHMD 621684</specimenCode>
</materialsCitation>
.
</paragraph>
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<emphasis id="BAD39F16FF964A10FF6BF8B9BF07F872" bold="true" box="[189,334,1917,1943]" pageId="28" pageNumber="29">Description</emphasis>
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</paragraph>
<paragraph id="88184304FF964A10FF6BF869BC65F80E" blockId="28.[189,1398,1965,2027]" pageId="28" pageNumber="29">
SIZE AND SHAPE. Body length
<quantity id="4F5FEEE1FF964A10FDC9F869BCC1F822" box="[543,648,1965,1991]" metricMagnitude="-2" metricUnit="m" metricValue="2.03" pageId="28" pageNumber="29" unit="mm" value="20.3">20.3 mm</quantity>
(approximate, specimen broken into several pieces). Maximum width
<quantity id="4F5FEEE1FF964A10FED1F815BF2DF80E" box="[263,356,2001,2027]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="28" pageNumber="29" unit="mm" value="4.1">4.1 mm</quantity>
. W/L ratio 20 %.
</paragraph>
<caption id="DCD8138CFF974A11FF68F84DBFA5F80F" ID-DOI="http://doi.org/10.5281/zenodo.3928364" ID-Zenodo-Dep="3928364" httpUri="https://zenodo.org/record/3928364/files/figure.png" pageId="29" pageNumber="30" startId="29.[190,233,1929,1955]" targetBox="[349,1236,266,1886]" targetPageId="29">
<paragraph id="88184304FF974A11FF68F84DBFA5F80F" blockId="29.[190,1399,1929,2026]" pageId="29" pageNumber="30">
<emphasis id="BAD39F16FF974A11FF68F84DBF51F846" bold="true" box="[190,280,1929,1955]" pageId="29" pageNumber="30">Fig. 19.</emphasis>
<taxonomicName id="4FA73887FF974A11FEC8F84DBD5AF841" authority="Hoffman, 2005" authorityName="Hoffmann" authorityYear="2005" box="[286,787,1929,1956]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="mwanihana">
<emphasis id="BAD39F16FF974A11FEC8F84DBC11F846" box="[286,600,1929,1955]" italics="true" pageId="29" pageNumber="30">Agrophogonus mwanihana</emphasis>
<bibRefCitation id="EC363EF5FF974A11FD88F84DBD5AF841" author="Hoffman R. L." box="[606,787,1929,1956]" pageId="29" pageNumber="30" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman, 2005</bibRefCitation>
</taxonomicName>
, specimen from Udzungwa Mts National Park, Sanje Kati camp &amp; plot (NHMD 621682), right gonopod.
<emphasis id="BAD39F16FF974A11FCF4F868BD10F823" bold="true" box="[802,857,1964,1990]" pageId="29" pageNumber="30">A–B</emphasis>
. Mesal view.
<emphasis id="BAD39F16FF974A11FC2BF868BA7FF823" bold="true" box="[1021,1078,1964,1990]" pageId="29" pageNumber="30">C–D</emphasis>
. Lateral view.
<emphasis id="BAD39F16FF974A11FB30F869BB53F822" bold="true" box="[1254,1306,1965,1991]" pageId="29" pageNumber="30">E–F</emphasis>
. Dorsal view. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="88184304FF944A12FF6BFEC8BD81FEC3" blockId="30.[189,968,267,294]" box="[189,968,267,294]" pageId="30" pageNumber="31">
COLOUR. After 18 years in alcohol without pigmentation (
<figureCitation id="109C5F81FF944A12FCB5FECFBDF2FEC3" box="[867,955,267,294]" captionStart="Fig" captionStartId="30.[189,232,1965,1991]" captionTargetBox="[418,1171,1411,1922]" captionTargetId="figure@30.[418,1171,1411,1922]" captionTargetPageId="30" captionText="Fig. 20. Agrophogonus pusillokiellandi Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621683). Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928366" httpUri="https://zenodo.org/record/3928366/files/figure.png" pageId="30" pageNumber="31">Fig. 20</figureCitation>
).
</paragraph>
<paragraph id="88184304FF944A12FF6BFE89BD35FE6F" blockId="30.[189,1398,333,395]" pageId="30" pageNumber="31">
BODY RINGS. Sternal process on ring 6 probably damaged, anterior edge very unevenly rounded and jagged (
<figureCitation id="109C5F81FF944A12FECDFEB4BFC0FE6F" box="[283,393,368,394]" captionStart="Fig" captionStartId="31.[189,232,1895,1921]" captionTargetBox="[236,1337,265,1863]" captionTargetId="figure@31.[234,1350,265,1863]" captionTargetPageId="31" captionText="Fig. 21. Agrophogonus pusillokiellandi Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621683). A–D, F–G. Left gonopod. A–B. Mesal view. C–D. Lateral view. F–G. Dorsal view. E. Right gonopod, lateral view. H. Sternal process on ring 6. Abbreviations: A = process A; B = process B; C = process C; L = process L. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928368" httpUri="https://zenodo.org/record/3928368/files/figure.png" pageId="30" pageNumber="31">Fig. 21H</figureCitation>
). Apparently no transverse sternal carinae.
</paragraph>
<paragraph id="88184304FF944A12FF6BFE76BAE9FDF7" blockId="30.[189,1399,433,530]" pageId="30" pageNumber="31">TELSON. Epiproct flattened at posterior end, forming a small, vertical flat surface with four protruding setae. Hypoproct semicircular with slightly triangular edge, paramedian tubercles large, extended beyond edge of sclerite. Setae on paraprocts protruding from distinct basal tubercles.</paragraph>
<paragraph id="88184304FF944A12FF6BFDFEBB63FC32" blockId="30.[189,1399,569,983]" pageId="30" pageNumber="31">
GONOPOD (
<figureCitation id="109C5F81FF944A12FE90FDFDBFD2FDB1" box="[326,411,569,596]" captionStart="Fig" captionStartId="31.[189,232,1895,1921]" captionTargetBox="[236,1337,265,1863]" captionTargetId="figure@31.[234,1350,265,1863]" captionTargetPageId="31" captionText="Fig. 21. Agrophogonus pusillokiellandi Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621683). A–D, F–G. Left gonopod. A–B. Mesal view. C–D. Lateral view. F–G. Dorsal view. E. Right gonopod, lateral view. H. Sternal process on ring 6. Abbreviations: A = process A; B = process B; C = process C; L = process L. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928368" httpUri="https://zenodo.org/record/3928368/files/figure.png" pageId="30" pageNumber="31">Fig. 21</figureCitation>
A–G). Gonopod aperture suboval, extending back between second pair of legs on seventh ring, elevated in front of coxae, eliminated medially. Gonocoxa with paracannular setae. Prefemur with deep basal fossa on lateral side. Nodal process L present, process M absent. Processes A and B slender and curved, process C broader and longer than A and B. Processes B and C somewhat forcipulate in their curvature and position to each other. Postnodal telepodite slender and acuminate, slightly flattened, but not laminate as in
<taxonomicName id="4FA73887FF944A12FD95FD2EBCA6FCE6" authorityName="Hoffmann" authorityYear="2005" box="[579,751,745,772]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="mwanihana">
<emphasis id="BAD39F16FF944A12FD95FD2EBCA6FCE6" box="[579,751,745,772]" italics="true" pageId="30" pageNumber="31">A. mwanihana</emphasis>
</taxonomicName>
and
<taxonomicName id="4FA73887FF944A12FCFBFD2EBDE9FCE6" baseAuthorityName="Verhoeff" baseAuthorityYear="1941" box="[813,928,745,772]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="harrisi">
<emphasis id="BAD39F16FF944A12FCFBFD2EBDE9FCE6" box="[813,928,745,772]" italics="true" pageId="30" pageNumber="31">A. harrisi</emphasis>
</taxonomicName>
. The right gonopod has a considerably higher number of small spikes and outgrowths than the left, and whereas process L on the left gonopod is slender and lance-shaped (
<figureCitation id="109C5F81FF944A12FE27FCF4BC15FCAF" box="[497,604,816,842]" captionStart="Fig" captionStartId="31.[189,232,1895,1921]" captionTargetBox="[236,1337,265,1863]" captionTargetId="figure@31.[234,1350,265,1863]" captionTargetPageId="31" captionText="Fig. 21. Agrophogonus pusillokiellandi Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621683). A–D, F–G. Left gonopod. A–B. Mesal view. C–D. Lateral view. F–G. Dorsal view. E. Right gonopod, lateral view. H. Sternal process on ring 6. Abbreviations: A = process A; B = process B; C = process C; L = process L. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928368" httpUri="https://zenodo.org/record/3928368/files/figure.png" pageId="30" pageNumber="31">Fig. 21D</figureCitation>
), on the right gonopod it is laminate and has a secondary tip halfway along its length (
<figureCitation id="109C5F81FF944A12FE57FC97BFA5FC88" box="[385,492,851,877]" captionStart="Fig" captionStartId="31.[189,232,1895,1921]" captionTargetBox="[236,1337,265,1863]" captionTargetId="figure@31.[234,1350,265,1863]" captionTargetPageId="31" captionText="Fig. 21. Agrophogonus pusillokiellandi Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621683). A–D, F–G. Left gonopod. A–B. Mesal view. C–D. Lateral view. F–G. Dorsal view. E. Right gonopod, lateral view. H. Sternal process on ring 6. Abbreviations: A = process A; B = process B; C = process C; L = process L. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928368" httpUri="https://zenodo.org/record/3928368/files/figure.png" pageId="30" pageNumber="31">Fig. 21E</figureCitation>
). The angle between the prefemoral region and postnodal telepodite is almost 90° on the left gonopod, but the postnodal telopodite is more slanting posteriad on the right gonopod. The gonopods are almost identical to those of
<taxonomicName id="4FA73887FF944A12FD35FC5EBD25FC56" authorityName="Hoffman" authorityYear="2005" box="[739,876,921,948]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="kiellandi">
<emphasis id="BAD39F16FF944A12FD35FC5EBD25FC56" box="[739,876,921,948]" italics="true" pageId="30" pageNumber="31">A. kiellandi</emphasis>
</taxonomicName>
, but nodal process C is shorter, broader and less curved in
<taxonomicName id="4FA73887FF944A12FEB0FC79BC74FC33" authority="Olsen &amp; Rosenmejer &amp; Enghoff, 2020" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[358,573,956,983]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="pusillokiellandi" status="sp. nov.">
<emphasis id="BAD39F16FF944A12FEB0FC79BC74FC33" box="[358,573,956,983]" italics="true" pageId="30" pageNumber="31">A. pusillokiellandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF944A12FD92FC79BCD6FC32" box="[580,671,957,983]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="109C5F81FF944A12FD78FC79BD4AFC32" box="[686,771,957,983]" captionStart="Fig" captionStartId="31.[189,232,1895,1921]" captionTargetBox="[236,1337,265,1863]" captionTargetId="figure@31.[234,1350,265,1863]" captionTargetPageId="31" captionText="Fig. 21. Agrophogonus pusillokiellandi Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621683). A–D, F–G. Left gonopod. A–B. Mesal view. C–D. Lateral view. F–G. Dorsal view. E. Right gonopod, lateral view. H. Sternal process on ring 6. Abbreviations: A = process A; B = process B; C = process C; L = process L. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928368" httpUri="https://zenodo.org/record/3928368/files/figure.png" pageId="30" pageNumber="31">Fig. 21</figureCitation>
) than in
<taxonomicName id="4FA73887FF944A12FCBAFC79BDBAFC33" authorityName="Hoffman" authorityYear="2005" box="[876,1011,956,983]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="kiellandi">
<emphasis id="BAD39F16FF944A12FCBAFC79BDBAFC33" box="[876,1011,956,983]" italics="true" pageId="30" pageNumber="31">A. kiellandi</emphasis>
</taxonomicName>
(
<bibRefCitation id="EC363EF5FF944A12FBD4FC79BAFDFC32" author="Hoffman R. L." box="[1026,1204,956,983]" pageId="30" pageNumber="31" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman 2005</bibRefCitation>
: fig. 451).
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF944A12FF6BFC3ABF67FB8F" pageId="30" pageNumber="31" type="distribution">
<paragraph id="88184304FF944A12FF6BFC3ABF1FFBFD" blockId="30.[189,342,1022,1048]" box="[189,342,1022,1048]" pageId="30" pageNumber="31">
<heading id="D350F468FF944A12FF6BFC3ABF1FFBFD" bold="true" box="[189,342,1022,1048]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="BAD39F16FF944A12FF6BFC3ABF1FFBFD" bold="true" box="[189,342,1022,1048]" pageId="30" pageNumber="31">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF944A12FF6BFBE9BF67FB8F" blockId="30.[189,1399,1068,1130]" pageId="30" pageNumber="31">
Known only from the
<typeStatus id="571CFDA6FF944A12FE03FBE9BC41FBA2" box="[469,520,1069,1095]" pageId="30" pageNumber="31">type</typeStatus>
locality, West Kilombero Scarp Forest Reserve, in the Udzungwa Mts,
<collectingCountry id="F0B00394FF944A12FF6BFB94BF63FB8F" box="[189,298,1104,1130]" name="Tanzania" pageId="30" pageNumber="31">Tanzania</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF944A2CFF6BFB56BB55FBF5" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31" type="discussion">
<paragraph id="88184304FF944A12FF6BFB56BF66FB49" blockId="30.[189,303,1170,1196]" box="[189,303,1170,1196]" pageId="30" pageNumber="31">
<heading id="D350F468FF944A12FF6BFB56BF66FB49" bold="true" box="[189,303,1170,1196]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="BAD39F16FF944A12FF6BFB56BF66FB49" bold="true" box="[189,303,1170,1196]" pageId="30" pageNumber="31">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF944A2CFF6BFB04BB55FBF5" blockId="30.[189,1399,1216,1348]" lastBlockId="32.[189,1399,267,1040]" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31">
The right-left differences observed on the gonopods of the unique specimen of
<taxonomicName id="4FA73887FF944A12FB92FB04BB53FB3F" authority="Olsen &amp; Rosenmejer &amp; Enghoff, 2020" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[1092,1306,1216,1242]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="pusillokiellandi" status="sp. nov.">
<emphasis id="BAD39F16FF944A12FB92FB04BB53FB3F" box="[1092,1306,1216,1242]" italics="true" pageId="30" pageNumber="31">A. pusillokiellandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF944A12FAC8FB04BB3EFB3F" box="[1310,1399,1216,1242]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
emphasize the desirability of having more than a single individual at hand when describing a new species, since with only one specimen it is impossible to estimate the level of intraspecific variation. Basing a new species on a single specimen which is almost identical to a known species (also known only from a single specimen), except for its body size, might seem ill-advised. The size difference in question, however, is dramatic: the width of the
<typeStatus id="571CFDA6FFAA4A2CFE27FEF5BC11FEAE" box="[497,600,305,331]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4FA73887FFAA4A2CFDABFEF5BD4DFEAE" authorityName="Hoffman" authorityYear="2005" box="[637,772,305,331]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="kiellandi">
<emphasis id="BAD39F16FFAA4A2CFDABFEF5BD4DFEAE" box="[637,772,305,331]" italics="true" pageId="32" pageNumber="33">A. kiellandi</emphasis>
</taxonomicName>
is 2.1 times that of the
<typeStatus id="571CFDA6FFAA4A2CFBC5FEF5BA33FEAE" box="[1043,1146,305,331]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4FA73887FFAA4A2CFB76FEF5BB3EFEAE" authority="Olsen &amp; Rosenmejer &amp; Enghoff, 2020" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[1184,1399,305,331]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="pusillokiellandi" status="sp. nov.">
<emphasis id="BAD39F16FFAA4A2CFB76FEF5BB3EFEAE" box="[1184,1399,305,331]" italics="true" pageId="32" pageNumber="33">A. pusillokiellandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFFAA4A2CFF6BFE93BF52FE94" box="[189,283,343,369]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
, corresponding to an almost tenfold volume difference. Large intraspecific size differences are not very frequent among polydesmidan millipedes, but there are some cases: in the
<taxonomicName id="4FA73887FFAA4A2CFB4CFEBFBB3AFE70" authorityName="Cook" authorityYear="1896" box="[1178,1395,379,405]" class="Diplopoda" family="Gomphodesmidae" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="family">Gomphodesmidae</taxonomicName>
, the most variable species known to date is
<taxonomicName id="4FA73887FFAA4A2CFD6DFE65BA33FE5E" authority="(Gerstacker, 1873)" baseAuthorityName="Gerstacker" baseAuthorityYear="1873" box="[699,1146,417,443]" class="Diplopoda" family="Gomphodesmidae" genus="Astrodesmus" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="laxus">
<emphasis id="BAD39F16FFAA4A2CFD6DFE65BDDFFE5E" box="[699,918,417,443]" italics="true" pageId="32" pageNumber="33">Astrodesmus laxus</emphasis>
(Gerstäcker, 1873)
</taxonomicName>
in which male width varies from
<quantity id="4F5FEEE1FFAA4A2CFE99FE02BFA9FE04" box="[335,480,454,481]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.5" metricValueMin="0.6" pageId="32" pageNumber="33" unit="mm" value="10.5" valueMax="15.0" valueMin="6.0">6 to 15 mm</quantity>
. The smallest form was originally described as a separate species,
<taxonomicName id="4FA73887FFAA4A2CFAD3FE02BF0EFDE0" authority="Cook, 1899" authorityName="Cook" authorityYear="1899" class="Diplopoda" family="Gomphodesmidae" genus="Astrodesmus" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="petilus">
<emphasis id="BAD39F16FFAA4A2CFAD3FE02BB3FFE05" box="[1285,1398,454,480]" italics="true" pageId="32" pageNumber="33">A. petilus</emphasis>
Cook, 1899
</taxonomicName>
, but was ranked as a subspecies of
<taxonomicName id="4FA73887FFAA4A2CFD0AFE28BD70FDE0" baseAuthorityName="Gerstacker" baseAuthorityYear="1873" box="[732,825,491,518]" class="Diplopoda" family="Gomphodesmidae" genus="Astrodesmus" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="laxus">
<emphasis id="BAD39F16FFAA4A2CFD0AFE28BD70FDE0" box="[732,825,491,518]" italics="true" pageId="32" pageNumber="33">A. laxus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EC363EF5FFAA4A2CFCB5FE2FBA6DFDE3" author="Hoffman R. L." box="[867,1060,491,518]" pageId="32" pageNumber="33" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman (2005)</bibRefCitation>
. Other subspecies of
<taxonomicName id="4FA73887FFAA4A2CFACFFE28BB3EFDE0" baseAuthorityName="Gerstacker" baseAuthorityYear="1873" box="[1305,1399,491,518]" class="Diplopoda" family="Gomphodesmidae" genus="Astrodesmus" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="laxus">
<emphasis id="BAD39F16FFAA4A2CFACFFE28BB3EFDE0" box="[1305,1399,491,518]" italics="true" pageId="32" pageNumber="33">A. laxus</emphasis>
</taxonomicName>
have male widths varying from
<quantity id="4F5FEEE1FFAA4A2CFDE2FDD5BC9BFDCE" box="[564,722,528,555]" metricMagnitude="-2" metricUnit="m" metricValue="1.215" metricValueMax="1.5" metricValueMin="0.93" pageId="32" pageNumber="33" unit="mm" value="12.15" valueMax="15.0" valueMin="9.3">9.3 to 15 mm</quantity>
(
<bibRefCitation id="EC363EF5FFAA4A2CFD37FDD5BDDDFDCE" author="Hoffman R. L." box="[737,916,528,555]" pageId="32" pageNumber="33" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman 2005</bibRefCitation>
).
<bibRefCitation id="EC363EF5FFAA4A2CFC71FDD5BA02FDCE" author="Decker P." box="[935,1099,529,555]" pageId="32" pageNumber="33" pagination="201 - 218" refId="ref11353" refString="Decker P. 2016. Integrative taxonomic revision of the polymorphic flat-millipede genera Oncocladosoma and Somethus in South Australia (Diplopoda: Polydesmida: Paradoxosomatidae). Invertebrate Systematics 30: 201 - 218. https: // doi. org / 10.1071 / IS 15047" type="journal article" year="2016">Decker (2016</bibRefCitation>
, pers.comm.) found male width to range from
<quantity id="4F5FEEE1FFAA4A2CFE6CFDF2BC2FFDB4" box="[442,614,566,593]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="4.3" metricValueMin="1.5" pageId="32" pageNumber="33" unit="mm" value="2.9" valueMax="4.3" valueMin="1.5">1.5 to 4.3 mm</quantity>
in
<taxonomicName id="4FA73887FFAA4A2CFD47FDF2BB0BFDB4" authority="(Attems, 1944)" baseAuthorityName="Attems" baseAuthorityYear="1944" box="[657,1346,566,593]" class="Diplopoda" family="Paradoxosomatidae" genus="Somethus" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis id="BAD39F16FFAA4A2CFD47FDF2BDC9FDB5" box="[657,896,566,592]" italics="true" pageId="32" pageNumber="33">Somethus castaneus</emphasis>
(Attems, 1944) (Paradoxosomatidae)
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and from
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in
<taxonomicName id="4FA73887FFAA4A2CFE1FFD98BD48FD90" authority="Jeekel, 2002" authorityName="Jeekel" authorityYear="2002" box="[457,769,603,630]" class="Diplopoda" family="Paradoxosomatidae" genus="Somethus" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="lancearius">
<emphasis id="BAD39F16FFAA4A2CFE1FFD98BC2AFD90" box="[457,611,603,630]" italics="true" pageId="32" pageNumber="33">S. lancearius</emphasis>
Jeekel, 2002
</taxonomicName>
, in the latter case even within one small conservation park.
<bibRefCitation id="EC363EF5FFAA4A2CFF29FD45BFFAFD7E" author="Mesibov R." box="[255,435,641,667]" pageId="32" pageNumber="33" pagination="103 - 146" refId="ref12301" refString="Mesibov R. 2006. The millipede genus Lissodesmus Chamberlin, 1920 (Diplopoda: Polydesmida: Dalodesmidae) from Tasmania and Victoria, with descriptions of a new genus and 24 new species. Memoirs of Museum Victoria 62: 103 - 146. https: // doi. org / 10.24199 / j. mmv. 2005.62.4" type="journal article" year="2006">Mesibov (2006</bibRefCitation>
, pers. comm.) found male widths between 1.2 and
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in
<taxonomicName id="4FA73887FFAA4A2CFBAFFD44BC7BFD24" authority="Mesibov, 2006" authorityName="Mesibov" authorityYear="2006" class="Diplopoda" family="Dalodesmidae" genus="Lissodesmus" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="BAD39F16FFAA4A2CFBAFFD44BB3FFD7F" box="[1145,1398,640,666]" italics="true" pageId="32" pageNumber="33">Lissodesmus hamatus</emphasis>
<bibRefCitation id="EC363EF5FFAA4A2CFF6BFD62BF24FD25" author="Mesibov R." box="[189,365,678,704]" pageId="32" pageNumber="33" pagination="103 - 146" refId="ref12301" refString="Mesibov R. 2006. The millipede genus Lissodesmus Chamberlin, 1920 (Diplopoda: Polydesmida: Dalodesmidae) from Tasmania and Victoria, with descriptions of a new genus and 24 new species. Memoirs of Museum Victoria 62: 103 - 146. https: // doi. org / 10.24199 / j. mmv. 2005.62.4" type="journal article" year="2006">Mesibov, 2006</bibRefCitation>
(Dalodesmidae)
</taxonomicName>
and on https://myriapodology.org/
<taxonomicName id="4FA73887FFAA4A2CFC1BFD62BA2FFD25" authorityName="Leach" authorityYear="1815" box="[973,1126,678,704]" class="Diplopoda" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="35" phylum="Arthropoda" rank="order">polydesmida</taxonomicName>
/size.html he published a striking photo of two female
<taxonomicName id="4FA73887FFAA4A2CFD91FD08BC87FD00" authorityName="Mesibov" authorityYear="2006" box="[583,718,715,741]" class="Diplopoda" family="Dalodesmidae" genus="Lissodesmus" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="BAD39F16FFAA4A2CFD91FD08BC87FD00" box="[583,718,715,741]" italics="true" pageId="32" pageNumber="33">L. hamatus</emphasis>
</taxonomicName>
, one a “dwarf”, the other a “giant”. Boyan Vagalinski (pers. comm.) has seen size differences on the same scale in
<taxonomicName id="4FA73887FFAA4A2CFC4EFD34BF30FCD4" authority="Daday, 1889" authorityName="Daday" authorityYear="1889" class="Diplopoda" family="Polydesmidae" genus="Polydesmus" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="mediterraneus">
<emphasis id="BAD39F16FFAA4A2CFC4EFD34BA9EFCEF" box="[920,1239,752,778]" italics="true" pageId="32" pageNumber="33">Polydesmus mediterraneus</emphasis>
Daday, 1889 (Polydesmidae)
</taxonomicName>
. We nevertheless choose to describe our small
<taxonomicName id="4FA73887FFAA4A2CFC60FCD2BA2BFCD5" authorityName="Hoffman" authorityYear="2005" box="[950,1122,790,816]" class="Diplopoda" family="Gomphodesmidae" genus="Agrophogonus" kingdom="Animalia" order="Polydesmida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="BAD39F16FFAA4A2CFC60FCD2BA2BFCD5" box="[950,1122,790,816]" italics="true" pageId="32" pageNumber="33">Agrophogonus</emphasis>
</taxonomicName>
as a new species, also being under the impression of the recent finding of two taxa of the genus
<taxonomicName id="4FA73887FFAA4A2CFB96FCFFBB3EFCB0" authority="Pocock, 1894" authorityName="Pocock" authorityYear="1894" box="[1088,1399,827,853]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="BAD39F16FFAA4A2CFB96FCFFBA80FCB0" box="[1088,1225,827,853]" italics="true" pageId="32" pageNumber="33">Thyropygus</emphasis>
Pocock, 1894
</taxonomicName>
(family
<taxonomicName id="4FA73887FFAA4A2CFECCFCA4BCC3FC9E" authority="Attems, 1909" authorityName="Attems" authorityYear="1909" box="[282,650,864,891]" class="Diplopoda" family="Harpagophoridae" kingdom="Animalia" order="Spirostreptida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="family">Harpagophoridae Attems, 1909</taxonomicName>
) with identical gonopods, but of different sizes and molecularly sufficiently different to be regarded as separate species (
<bibRefCitation id="EC363EF5FFAA4A2CFCB8FC42BA3EFC45" author="Pimvichai P. &amp; Enghoff H. &amp; Panha S." box="[878,1143,902,928]" pageId="32" pageNumber="33" pagination="47 - 68" refId="ref12537" refString="Pimvichai P., Enghoff H. &amp; Panha S. 2011. A revision of the Thyropygus allevatus group. Part 3: the T. induratus subgroup (Diplopoda: Spirostreptida: Harpagophoridae). Zootaxa 2941: 47 - 68. https: // doi. org / 10.11646 / zootaxa. 2941.1.3" type="journal article" year="2011">
Pimvichai
<emphasis id="BAD39F16FFAA4A2CFC24FC43BA78FC45" box="[1010,1073,902,928]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
2011
</bibRefCitation>
). The coexistence of several pairs of morphotypes, interpreted as species, of the genus
<emphasis id="BAD39F16FFAA4A2CFC2FFC6FBAFAFC20" box="[1017,1203,939,965]" italics="true" pageId="32" pageNumber="33">Nepalmatojulus</emphasis>
, Mauriès, 1983 (family
<taxonomicName id="4FA73887FFAA4A2CFECCFC14BC4FFC0E" authority="Leach, 1814" authorityName="Leach" authorityYear="1814" box="[282,518,976,1003]" class="Diplopoda" family="Julidae" kingdom="Animalia" order="Julida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="family">Julidae Leach, 1814</taxonomicName>
) with identical or extremely similar gonopods in the mountains of northern
<collectingCountry id="F0B00394FFAA4A2CFF6BFC32BF6CFBF5" box="[189,293,1014,1040]" name="Thailand" pageId="32" pageNumber="33">Thailand</collectingCountry>
(
<bibRefCitation id="EC363EF5FFAA4A2CFEE2FC32BF90FBF5" author="Enghoff H." box="[308,473,1014,1040]" pageId="32" pageNumber="33" pagination="241 - 331" refId="ref11399" refString="Enghoff H. 1987. Revision of Nepalmatoiuluus Mauries, 1983 - a Southeast Asian genus of millipedes (Diplopoda: Julida: Julidae). Courier Forschungsinstitut Senckenberg 93: 241 - 331." type="journal article" year="1987">Enghoff 1987</bibRefCitation>
) also suggests that size plays an important role in millipede speciation.
</paragraph>
</subSubSection>
</treatment>
</document>

236
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<emphasis id="BAD39F16FF994A1FFE3FFE0ABD5EFE0C" bold="true" box="[489,791,462,489]" italics="true" pageId="19" pageNumber="20">Emphysemastix dracarys</emphasis>
Olsen &amp; Enghoff
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,
<figureCitation id="109C5F81FF994A1FFCC1FDD1BD7EFDCA" box="[791,823,533,559]" captionStart="Fig" captionStartId="20.[189,232,1930,1956]" captionTargetBox="[259,1314,265,1901]" captionTargetId="figure@20.[259,1331,265,1901]" captionTargetPageId="20" captionText="Fig. 14. Emphysemastic dracarys Olsen &amp; Enghoff sp. nov., holotype, ♂, left gonopod (NHMD 621675). A–B. Mesal view. C–D. Lateral view. E–F. Dorsal view. Abbreviations: L = Process L; rl = rounded lobe; sg = subglobose enlargement; sp = subterminal process. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928354" httpUri="https://zenodo.org/record/3928354/files/figure.png" pageId="19" pageNumber="20">14</figureCitation>
,
<figureCitation id="109C5F81FF994A1FFC95FDD1BD3CFDD5" box="[835,885,533,560]" captionStart="Fig" captionStartId="21.[189,232,1930,1956]" captionTargetBox="[354,1230,1369,1895]" captionTargetId="figure@21.[343,1238,1364,1901]" captionTargetPageId="21" captionText="Fig. 15. Emphysemastix spp., gonopod tip. A. E. dracarys Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621675). B. E. frampt Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621677). Abbreviations: so = solenomere; sp = subterminal process. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3928356" httpUri="https://zenodo.org/record/3928356/files/figure.png" pageId="19" pageNumber="20">15A</figureCitation>
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<emphasis id="BAD39F16FF994A1FFF6BFD93BF7FFD94" bold="true" box="[189,310,599,625]" pageId="19" pageNumber="20">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF994A1FFF6BFD42BB3EFD02" blockId="19.[189,1399,646,743]" pageId="19" pageNumber="20">Differs from the other species of the genus by the combination of having a small, not very distinct subtriangular ventral lobe on the gonopod prefemur, missing nodal process M, possessing a welldeveloped process L, and having indentures on the postnodal telepodite on the outer side of the curvature.</paragraph>
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<paragraph id="88184304FF994A1FFF6BFCCBBF0DFCCC" blockId="19.[189,324,783,809]" box="[189,324,783,809]" pageId="19" pageNumber="20">
<heading id="D350F468FF994A1FFF6BFCCBBF0DFCCC" bold="true" box="[189,324,783,809]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="BAD39F16FF994A1FFF6BFCCBBF0DFCCC" bold="true" box="[189,324,783,809]" pageId="19" pageNumber="20">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF994A1FFF6BFCF9BD48FC7B" blockId="19.[189,1399,829,927]" pageId="19" pageNumber="20">The name is to be treated as a noun in apposition. The tip of the gonopod looks like the gaping mouth of a dragon when seen from the dorsal side. The word “dracarys” is a command used in the TV series “Game of Thrones” to make dragons breathe fire.</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF994A1FFF6BFC02BA57FB5E" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="88184304FF994A1FFF6BFC02BFE3FC05" blockId="19.[189,426,966,992]" box="[189,426,966,992]" pageId="19" pageNumber="20">
<heading id="D350F468FF994A1FFF6BFC02BFE3FC05" bold="true" box="[189,426,966,992]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="BAD39F16FF994A1FFF6BFC02BFE3FC05" bold="true" box="[189,426,966,992]" pageId="19" pageNumber="20">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF994A1FFF3CFC31BF15FBEA" blockId="19.[189,1398,1013,1110]" box="[234,348,1013,1039]" pageId="19" pageNumber="20">
<emphasis id="BAD39F16FF994A1FFF3CFC31BF15FBEA" bold="true" box="[234,348,1013,1039]" pageId="19" pageNumber="20">
<typeStatus id="571CFDA6FF994A1FFF3CFC31BF15FBEA" box="[234,348,1013,1039]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="88184304FF994A1FFF6BFBDCBF37FBB3" blockId="19.[189,1398,1013,1110]" pageId="19" pageNumber="20">
<materialsCitation id="38CF4959FF994A1FFF6BFBDCBF37FBB3" ID-GBIF-Occurrence="2803530304" collectingDate="1996-03" collectingDateMax="1996-04-31" collectingDateMin="1996-03" collectionCode="NHMD" collectorName="L. Sorensen" country="Tanzania" latitude="-7.7666664" location="Iringa City" longLatPrecision="1303" longitude="35.7" pageId="19" pageNumber="20" specimenCode="NHMD 621675" specimenCount="1" stateProvince="Iringa Region" typeStatus="holotype">
<collectingCountry id="F0B00394FF994A1FFF6BFBDCBF18FBD7" box="[189,337,1048,1074]" name="Tanzania" pageId="19" pageNumber="20">TANZANIA</collectingCountry>
•
<specimenCount id="9EA1888DFF994A1FFEA4FBDDBFC3FBD6" box="[370,394,1049,1075]" pageId="19" pageNumber="20" type="male">♂</specimenCount>
;
<collectingRegion id="4A638DE6FF994A1FFE4DFBDCBC0DFBD7" box="[411,580,1048,1074]" country="Tanzania" name="Iringa" pageId="19" pageNumber="20">Iringa Region</collectingRegion>
,
<location id="8D7815DFFF994A1FFD82FBDCBC92FBD7" LSID="urn:lsid:plazi:treatment:000EF212FF994A19FE3FFE0ABC9DFDC7:8D7815DFFF994A1FFD82FBDCBC92FBD7" box="[596,731,1048,1074]" country="Tanzania" latitude="-7.7666664" longLatPrecision="1303" longitude="35.7" name="Iringa City" pageId="19" pageNumber="20" stateProvince="Iringa Region">Iringa City</location>
;
<geoCoordinate id="ED9325C3FF994A1FFD3DFBD2BD0EFBD7" box="[747,839,1046,1074]" degrees="7" direction="south" minutes="46" orientation="latitude" pageId="19" pageNumber="20" precision="925" value="-7.7666664">7°46′ S</geoCoordinate>
,
<geoCoordinate id="ED9325C3FF994A1FFC8EFBD2BD8CFBD6" box="[856,965,1046,1075]" degrees="35" direction="east" minutes="42" orientation="longitude" pageId="19" pageNumber="20" precision="925" value="35.7">35°42′ E</geoCoordinate>
;
<date id="FC1965C4FF994A1FFC03FBDCBADEFBD7" box="[981,1175,1048,1074]" pageId="19" pageNumber="20" value="1996-03" valueMax="1996-04-31" valueMin="1996-03-00">
<collectingDate id="EC5D9C2CFF994A1FFC03FBDCBADEFBD7" box="[981,1175,1048,1074]" pageId="19" pageNumber="20" value="1996-03" valueMax="1996-04-31" valueMin="1996-03-00">Mar.–Apr. 1996</collectingDate>
</date>
;
<collectorName id="255226D2FF994A1FFB7EFBDDBB71FBD7" box="[1192,1336,1048,1075]" pageId="19" pageNumber="20">L. Sørensen</collectorName>
leg.;
<specimenCode id="D801EB7FFF994A1FFF6BFBF8BF33FBB3" box="[189,378,1083,1110]" collectionCode="NHMD" pageId="19" pageNumber="20">NHMD 621675</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="88184304FF994A1FFF3CFBB9BFEFFB72" blockId="19.[189,1054,1149,1211]" box="[234,422,1149,1175]" pageId="19" pageNumber="20">
<emphasis id="BAD39F16FF994A1FFF3CFBB9BFEFFB72" bold="true" box="[234,422,1149,1175]" pageId="19" pageNumber="20">Other material</emphasis>
</paragraph>
<paragraph id="88184304FF994A1FFF6BFB65BA57FB5E" blockId="19.[189,1054,1149,1211]" box="[189,1054,1185,1211]" pageId="19" pageNumber="20">
<materialsCitation id="38CF4959FF994A1FFF6BFB65BA53FB5E" ID-GBIF-Occurrence="2803530306" box="[189,1050,1185,1211]" collectingDate="1996-03" collectingDateMax="1996-04-31" collectingDateMin="1996-03" collectionCode="NHMD" collectorName="L. Sorensen" country="Tanzania" latitude="-7.7666664" location="TANZANIA" longLatPrecision="1303" longitude="35.7" pageId="19" pageNumber="20" specimenCode="NHMD 621676" specimenCount="1" specimenCount-female="1" stateProvince="Iringa">
<collectingCountry id="F0B00394FF994A1FFF6BFB65BF1BFB5E" box="[189,338,1185,1211]" name="Tanzania" pageId="19" pageNumber="20">TANZANIA</collectingCountry>
•
<specimenCount id="9EA1888DFF994A1FFEBDFB65BFD1FB5E" box="[363,408,1185,1211]" pageId="19" pageNumber="20" type="female">1 ♀</specimenCount>
; same collecting data as for holotype;
<specimenCode id="D801EB7FFF994A1FFC8BFB65BA53FB5E" box="[861,1050,1185,1211]" collectionCode="NHMD" pageId="19" pageNumber="20">NHMD 621676</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF994A19FF6BFB26BDCEFE6A" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20" type="description">
<paragraph id="88184304FF994A1FFF6BFB26BF07FB19" blockId="19.[189,334,1250,1276]" box="[189,334,1250,1276]" pageId="19" pageNumber="20">
<heading id="D350F468FF994A1FFF6BFB26BF07FB19" bold="true" box="[189,334,1250,1276]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="BAD39F16FF994A1FFF6BFB26BF07FB19" bold="true" box="[189,334,1250,1276]" pageId="19" pageNumber="20">Description</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF994A1FFF3CFAD6BF63FAC9" blockId="19.[189,1399,1298,1395]" box="[234,298,1298,1324]" pageId="19" pageNumber="20">
<emphasis id="BAD39F16FF994A1FFF3CFAD6BF63FAC9" bold="true" box="[234,298,1298,1324]" pageId="19" pageNumber="20">Male</emphasis>
</paragraph>
<paragraph id="88184304FF994A1FFF6BFAF1BF48FA97" blockId="19.[189,1399,1298,1395]" pageId="19" pageNumber="20">
SIZE AND SHAPE. A rather slender species. Body length ca
<quantity id="4F5FEEE1FF994A1FFCBDFAF1BD9EFAAA" box="[875,983,1333,1360]" metricMagnitude="-2" metricUnit="m" metricValue="4.65" pageId="19" pageNumber="20" unit="mm" value="46.5">46.5 mm</quantity>
. Maximum width
<quantity id="4F5FEEE1FF994A1FFB67FAF1BAB1FAAA" box="[1201,1272,1333,1359]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="19" pageNumber="20" unit="mm" value="7.0">7 mm</quantity>
. W/L ratio 15 %.
</paragraph>
<paragraph id="88184304FF994A1FFF6BFA52BC31FA31" blockId="19.[189,1399,1430,1492]" pageId="19" pageNumber="20">COLOUR. Dorsum of metazona darker brown than sides, sternum and prozona; paranota light brown. Antennae very dark, legs dark brown.</paragraph>
<paragraph id="88184304FF994A1FFF6BFA3CBC32F97A" blockId="19.[189,1399,1527,1696]" pageId="19" pageNumber="20">BODY RINGS. Paranotum of rings 2–4 extended anteriad, on rings 4–19 extended gradually more posteriad. Two transverse carinae on the posterior end of rings 8 and 9, and 4 transverse carinae on rings 10–19. Torus present as clear cones. Stricture relatively deep on dorsal side, with sharp edge in front of anterior spiracles and pleurosternal carinae. Pleurosternal carinae present on rings 2–17, but very small on ring 2. Paxillus triangular, pointed.</paragraph>
<paragraph id="88184304FF994A1FFF6BF907BF36F8E4" blockId="19.[189,1398,1731,1793]" pageId="19" pageNumber="20">
LEGS. Coxae and prefemora with fewer setae, and basal tubercles smaller than in
<taxonomicName id="4FA73887FF994A1FFBA3F900BAAAF938" authority="Olsen &amp; Rosenmejer &amp; Enghoff, 2020" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[1141,1251,1731,1757]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="frampt" status="sp. nov.">
<emphasis id="BAD39F16FF994A1FFBA3F900BAAAF938" box="[1141,1251,1731,1757]" italics="true" pageId="19" pageNumber="20">E. frampt</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF994A1FFB3DF900BB0CF93B" box="[1259,1349,1732,1758]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4FA73887FF994A1FFF6BF92CBF30F8E4" baseAuthorityName="Carl" baseAuthorityYear="1909" box="[189,377,1767,1793]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="flavosignatus">
<emphasis id="BAD39F16FF994A1FFF6BF92CBF30F8E4" box="[189,377,1767,1793]" italics="true" pageId="19" pageNumber="20">E. flavosignatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="88184304FF994A1FFF6BF8E1BF75F887" blockId="19.[189,1398,1829,1890]" pageId="19" pageNumber="20">HYPOPROCT. Paramedian tubercles large, extended beyond edge of sclerite, but not beyond end of median projection.</paragraph>
<paragraph id="88184304FF994A19FF6BF84EBDCEFE6A" blockId="19.[189,1398,1930,2027]" lastBlockId="21.[189,1399,267,400]" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20">
GONOPOD (
<figureCitation id="109C5F81FF994A1FFE91F84EBF98F840" box="[327,465,1930,1957]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="20.[189,232,1930,1956]" captionStartId-1="21.[189,232,1930,1956]" captionTargetBox-0="[259,1314,265,1901]" captionTargetBox-1="[354,1230,1369,1895]" captionTargetId-0="figure@20.[259,1331,265,1901]" captionTargetId-1="figure@21.[343,1238,1364,1901]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionText-0="Fig. 14. Emphysemastic dracarys Olsen &amp; Enghoff sp. nov., holotype, ♂, left gonopod (NHMD 621675). A–B. Mesal view. C–D. Lateral view. E–F. Dorsal view. Abbreviations: L = Process L; rl = rounded lobe; sg = subglobose enlargement; sp = subterminal process. Scale bars: 0.5 mm." captionText-1="Fig. 15. Emphysemastix spp., gonopod tip. A. E. dracarys Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621675). B. E. frampt Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621677). Abbreviations: so = solenomere; sp = subterminal process. Scale bars: 0.25 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3928354" figureDoi-1="http://doi.org/10.5281/zenodo.3928356" httpUri-0="https://zenodo.org/record/3928354/files/figure.png" httpUri-1="https://zenodo.org/record/3928356/files/figure.png" pageId="19" pageNumber="20">Figs 14–15</figureCitation>
). Gonocoxae with lateral row of setae across distal edge of anterior side of gonocoxae, no paracannular setae. Prefemoral part with a small rounded ventral lobe. Telopodite endonodal, process L present, process M absent. Postnodal telepodite long, curved ventrad almost perpendicular to setose prefemoral region and slightly mesad, then sharply dorsad and expanding into subglobose, hollow enlargement, then curving almost 270° mesad and posteriad, extending laterad beyond gonocoxa and prefemoral region. Anterior and mesal side of this curvature with indentures. Subterminal process narrow, triangular, curved ventromesad.
</paragraph>
</subSubSection>
<caption id="DCD8138CFF9E4A18FF6BF84EBA37F80E" ID-DOI="http://doi.org/10.5281/zenodo.3928354" ID-Zenodo-Dep="3928354" httpUri="https://zenodo.org/record/3928354/files/figure.png" pageId="20" pageNumber="21" startId="20.[189,232,1930,1956]" targetBox="[259,1314,265,1901]" targetPageId="20">
<paragraph id="88184304FF9E4A18FF6BF84EBA37F80E" blockId="20.[189,1399,1930,2027]" pageId="20" pageNumber="21">
<emphasis id="BAD39F16FF9E4A18FF6BF84EBF50F841" bold="true" box="[189,281,1930,1956]" pageId="20" pageNumber="21">Fig. 14.</emphasis>
<taxonomicName id="4FA73887FF9E4A18FEC9F84EBD5AF841" authority="Olsen &amp; Enghoff" authorityName="Olsen &amp; Enghoff" authorityYear="2020" box="[287,787,1930,1956]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastic" kingdom="Animalia" order="Polydesmida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="dracarys" status="sp. nov.">
<emphasis id="BAD39F16FF9E4A18FEC9F84EBC0DF841" box="[287,580,1930,1956]" italics="true" pageId="20" pageNumber="21">Emphysemastic dracarys</emphasis>
Olsen &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF9E4A18FCCFF84EBD3AF841" box="[793,883,1930,1956]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂, left gonopod (NHMD 621675).
<emphasis id="BAD39F16FF9E4A18FF6BF869BEBDF822" bold="true" box="[189,244,1965,1991]" pageId="20" pageNumber="21">A–B</emphasis>
. Mesal view.
<emphasis id="BAD39F16FF9E4A18FE48F869BF9EF822" bold="true" box="[414,471,1965,1991]" pageId="20" pageNumber="21">C–D</emphasis>
. Lateral view.
<emphasis id="BAD39F16FF9E4A18FD5AF869BC89F822" bold="true" box="[652,704,1965,1991]" pageId="20" pageNumber="21">E–F</emphasis>
. Dorsal view. Abbreviations:
<emphasis id="BAD39F16FF9E4A18FBF0F86ABA7FF822" box="[1062,1078,1966,1991]" italics="true" pageId="20" pageNumber="21">L</emphasis>
= Process L;
<emphasis id="BAD39F16FF9E4A18FB36F869BABDF822" box="[1248,1268,1965,1991]" italics="true" pageId="20" pageNumber="21">rl</emphasis>
= rounded lobe;
<emphasis id="BAD39F16FF9E4A18FF29F815BF50F80F" box="[255,281,2001,2026]" italics="true" pageId="20" pageNumber="21">sg</emphasis>
= subglobose enlargement;
<emphasis id="BAD39F16FF9E4A18FDB2F815BC37F80F" box="[612,638,2001,2026]" italics="true" pageId="20" pageNumber="21">sp</emphasis>
= subterminal process. Scale bars: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C0BD108FFF9F4A19FF6BFE72BC9DFDC7" pageId="21" pageNumber="22" type="distribution">
<paragraph id="88184304FF9F4A19FF6BFE72BF1FFE35" blockId="21.[189,342,438,464]" box="[189,342,438,464]" pageId="21" pageNumber="22">
<heading id="D350F468FF9F4A19FF6BFE72BF1FFE35" bold="true" box="[189,342,438,464]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="BAD39F16FF9F4A19FF6BFE72BF1FFE35" bold="true" box="[189,342,438,464]" pageId="21" pageNumber="22">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF9F4A19FF6BFE22BC9DFDC7" blockId="21.[189,1399,485,547]" pageId="21" pageNumber="22">
Known only from the
<typeStatus id="571CFDA6FF9F4A19FE1CFE22BFB4FDE5" box="[458,509,486,512]" pageId="21" pageNumber="22">type</typeStatus>
locality,
<collectingRegion id="4A638DE6FF9F4A19FDBBFE21BCFAFE1A" box="[621,691,485,511]" country="Tanzania" name="Iringa" pageId="21" pageNumber="22">Iringa</collectingRegion>
city, some
<quantity id="4F5FEEE1FF9F4A19FCE9FE21BDC2FE1A" box="[831,907,485,512]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="21" pageNumber="22" unit="km" value="50.0">50 km</quantity>
NW of the Udzungwa Mts (
<figureCitation id="109C5F81FF9F4A19FB31FE21BB78FE1A" box="[1255,1329,485,511]" captionStart="Fig" captionStartId="2.[189,232,1966,1992]" captionTargetBox="[199,1398,1100,1897]" captionTargetId="figure@2.[194,1482,1044,1958]" captionTargetPageId="2" captionText="Fig. 2. Map of the Udzungwa Mts showing collection sites for gomphodesmids. Based on Marshall et al. (2010: fig. 1)." figureDoi="http://doi.org/10.5281/zenodo.3928330" httpUri="https://zenodo.org/record/3928330/files/figure.png" pageId="21" pageNumber="22">Fig. 2</figureCitation>
). The altitude of the
<typeStatus id="571CFDA6FF9F4A19FEB1FDCDBFD3FDC6" box="[359,410,521,547]" pageId="21" pageNumber="22">type</typeStatus>
locality is ca
<quantity id="4F5FEEE1FF9F4A19FDEBFDCCBCDFFDC6" box="[573,662,520,547]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="21" pageNumber="22" unit="m" value="1600.0">1600 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="D0DF3B87CF7E55EF67B412E07F694766">A mountain of millipedes IX: Species of the family Gomphodesmidae from the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida)</mods:title>
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<mods:namePart id="F5D4DA8863D6D406E9318CC6722E4206">Olsen, Sissel Anna</mods:namePart>
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<mods:affiliation id="1FA2F9E2B55AC4169E7AD6048879ADF9">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 København Ø, Denmark</mods:affiliation>
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<mods:namePart id="106D9C62DDBB04A1319A45B421849824">Rosenmejer, Trine</mods:namePart>
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<mods:namePart id="8724E459E00C1239C76E2F7170B876D5">Enghoff, Henrik</mods:namePart>
<mods:nameIdentifier id="869981932EC763619EEED9BC81936185" type="LSID">FB09A817-000D-43C3-BCC4-2BC1E5373635</mods:nameIdentifier>
<mods:affiliation id="62454725E5F4A0E9027FCBF534442973">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 København Ø, Denmark</mods:affiliation>
<mods:nameIdentifier id="A53EABAD46CAD04A776C4C082A34B15F" type="email">henghoff@snm.ku.dk</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="F9936BBA1214ADFE486C8B298EF18245">text</mods:typeOfResource>
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<mods:titleInfo id="3E3D10290B3C2C936601C031B66A4BB3">
<mods:title id="2000F3CBB760CDA21849BEF45B81B673">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="11DC39F2E2AEF4BC9CF69BD5543F88C7">
<mods:date id="DFA68111E226417AF9D7F90582082A90">2020</mods:date>
<mods:detail id="DB1EE01ADEB7780BF336260EA9A128FF" type="pubDate">
<mods:number id="9CE13777745A5FF4D1B3C86F3BDD6581">2020-06-29</mods:number>
</mods:detail>
<mods:detail id="E2AC8B656AA4D2CD1992EBF059B1A12E" type="volume">
<mods:number id="37F9110BE3DFFB0867193631E85313DC">675</mods:number>
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<mods:start id="6155D629EA6FED7154AB8136C06A0369">1</mods:start>
<mods:end id="C7E431A46567CE82E4E742CD33FDE797">35</mods:end>
</mods:extent>
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<mods:classification id="66BDA37361C8A935C3A8139BDE7583E8">journal article</mods:classification>
<mods:identifier id="BB8B42FD61FDDFD2CAD7D6498B139402" type="CLB-Dataset">21573</mods:identifier>
<mods:identifier id="1A561968B29BC279A81B97D8B1C8C45F" type="DOI">10.5852/ejt.2020.675</mods:identifier>
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<treatment id="000EF212FF9B4A1FFDE2F9EABB3EFE75" ID-DOI="http://doi.org/10.5281/zenodo.4332915" ID-GBIF-Taxon="164870651" ID-Zenodo-Dep="4332915" LSID="urn:lsid:plazi:treatment:000EF212FF9B4A1FFDE2F9EABB3EFE75" httpUri="http://treatment.plazi.org/id/000EF212FF9B4A1FFDE2F9EABB3EFE75" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
<subSubSection id="C0BD108FFF9B4A1DFDE2F9EABA49F9A2" box="[564,1024,1581,1608]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="88184304FF9B4A1DFDE2F9EABA49F9A2" blockId="17.[564,1024,1581,1608]" box="[564,1024,1581,1608]" pageId="17" pageNumber="18">
<heading id="D350F468FF9B4A1DFDE2F9EABA49F9A2" box="[564,1024,1581,1608]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="18" reason="2">
Genus
<taxonomicName id="4FA73887FF9B4A1DFD50F9E9BA49F9A2" ID-CoL="7PD53" authority="Hoffman, 1966" authorityName="Hoffman" authorityYear="1966" box="[646,1024,1581,1608]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="BAD39F16FF9B4A1DFD50F9E9BD0CF9AD" bold="true" box="[646,837,1581,1608]" italics="true" pageId="17" pageNumber="18">Emphysemastix</emphasis>
<bibRefCitation id="EC363EF5FF9B4A1DFC9BF9E9BA49F9A2" author="Hoffman R. L." box="[845,1024,1581,1607]" pageId="17" pageNumber="18" pagination="193 - 208" refId="ref11929" refString="Hoffman R. L. 1966. Notes on Aulodesmus and the proposal of a related new genus (Polydesmida: Gomphodesmidae). Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut 63: 193 - 208." type="journal article" year="1966">Hoffman, 1966</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF9B4A1EFF6BF9B5BF83FDB5" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="88184304FF9B4A1DFF6BF9B5BCF9F969" blockId="17.[189,688,1649,1676]" box="[189,688,1649,1676]" pageId="17" pageNumber="18">
<emphasis id="BAD39F16FF9B4A1DFF6BF9B5BF7FF96E" bold="true" box="[189,310,1649,1675]" pageId="17" pageNumber="18">Diagnosis</emphasis>
(after
<bibRefCitation id="EC363EF5FF9B4A1DFE52F9B5BC7BF969" author="Hoffman R. L." box="[388,562,1649,1676]" pageId="17" pageNumber="18" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman 2005</bibRefCitation>
, modified)
</paragraph>
<paragraph id="88184304FF9B4A1EFF6BF967BF83FDB5" blockId="17.[189,1399,1698,2028]" lastBlockId="18.[189,1399,266,592]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
Small to moderate-sized (length
<quantity id="4F5FEEE1FF9B4A1DFDFBF966BCE0F958" box="[557,681,1698,1725]" metricMagnitude="-2" metricUnit="m" metricValue="4.85" metricValueMax="6.5" metricValueMin="3.2" pageId="17" pageNumber="18" unit="mm" value="48.5" valueMax="65.0" valueMin="32.0">32–65mm</quantity>
)gomphodesmids,pore formula:5,7, 9–19. Four apical sensory cones on antennae, clear, transverse fossa present on 5
<superScript id="7FD2EE4CFF9B4A1DFC9FF903BD1FF930" attach="left" box="[841,854,1735,1749]" fontSize="6" pageId="17" pageNumber="18">th</superScript>
and 6
<superScript id="7FD2EE4CFF9B4A1DFC74F903BDE6F930" attach="left" box="[930,943,1735,1749]" fontSize="6" pageId="17" pageNumber="18">th</superScript>
antennomere. Collum asymmetrically narrowed laterally, with a shallow emargination in the posterior edge. Paranota placed high on the sides, moderate to large in size, dorsum only slightly convex. Sterna broad, with two transverse carinae on posterior end of body ring 8, four transverse carinae on rings 9–19 (except
<taxonomicName id="4FA73887FF9B4A1DFBF8F8FEBB3EF8B0" authority="Hoffman, 2005" authorityName="Hoffman" authorityYear="2005" box="[1070,1399,1850,1877]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="congdoni">
<emphasis id="BAD39F16FF9B4A1DFBF8F8FEBAF5F8B1" box="[1070,1212,1850,1876]" italics="true" pageId="17" pageNumber="18">E. congdoni</emphasis>
<bibRefCitation id="EC363EF5FF9B4A1DFB12F8FEBB3EF8B0" author="Hoffman R. L." box="[1220,1399,1850,1877]" pageId="17" pageNumber="18" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman, 2005</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4FA73887FF9B4A1DFF38F8A4BF3FF89C" authority="Olsen &amp; Rosenmejer &amp; Enghoff, 2020" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[238,374,1887,1913]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="dracarys" status="sp. nov.">
<emphasis id="BAD39F16FF9B4A1DFF38F8A4BF3FF89C" box="[238,374,1887,1913]" italics="true" pageId="17" pageNumber="18">E. dracarys</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF9B4A1DFEABF8A4BF92F89F" box="[381,475,1888,1914]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
). Coxae and prefemora with long hairs originating in clear basal tubercles on at least posterior legs. Sides of metazona smooth. Torus (
<figureCitation id="109C5F81FF9B4A1DFC93F841BDE9F87A" box="[837,928,1925,1951]" captionStart="Fig" captionStartId="5.[189,232,1803,1829]" captionTargetBox="[196,1394,1386,1769]" captionTargetId="figure@5.[189,1398,1376,1774]" captionTargetPageId="5" captionText="Fig. 4. Particular gomphodesmid structural elements. A. Paxillus – a mid-ventral, most often triangular process on body ring 15 in most gomphodesmid males, from Hoffman (2005), modified. B. Torus – a low convexity of variable size and shape, found on the ventral surface of the poriferous paranota in males of many gomphodesmid species. Not drawn to scale." figureDoi="http://doi.org/10.5281/zenodo.3928334" httpUri="https://zenodo.org/record/3928334/files/figure.png" pageId="17" pageNumber="18">Fig. 4B</figureCitation>
) present as small, distinct, cone-shaped projections. Subonychial pads on male legs 1–6. Sternal process of segment 6 long and slender, with median apiculus (
<figureCitation id="109C5F81FF9B4A1DFE5EF815BFACF809" box="[392,485,2001,2028]" captionStart="Fig" captionStartId="6.[189,232,1500,1526]" captionTargetBox="[217,1367,909,1445]" captionTargetId="figure@6.[207,1377,905,1453]" captionTargetPageId="6" captionText="Fig. 5. Sternal process on body ring 6. A. Emphysemastix spp. B. Agrophogonus spp. except A. hamulifer Olsen &amp; Enghoff sp. nov. C. Agrophogonus hamulifer Olsen &amp; Enghoff sp. nov. D. Pogoropsis prolixopes Rosenmejer &amp; Enghoff gen. et sp. nov. E. Pogoro siren Rosenmejer &amp; Enghoff sp. nov. F. Pogoro scharffi Hoffman, 2005. G. Pogoro alopias Rosenmejer &amp; Enghoff sp. nov. Not drawn to scale." figureDoi="http://doi.org/10.5281/zenodo.3928336" httpUri="https://zenodo.org/record/3928336/files/figure.png" pageId="17" pageNumber="18">Fig. 5A</figureCitation>
). Paxillus (
<figureCitation id="109C5F81FF9B4A1DFDB2F815BCF7F80E" box="[612,702,2001,2027]" captionStart="Fig" captionStartId="5.[189,232,1803,1829]" captionTargetBox="[196,1394,1386,1769]" captionTargetId="figure@5.[189,1398,1376,1774]" captionTargetPageId="5" captionText="Fig. 4. Particular gomphodesmid structural elements. A. Paxillus – a mid-ventral, most often triangular process on body ring 15 in most gomphodesmid males, from Hoffman (2005), modified. B. Torus – a low convexity of variable size and shape, found on the ventral surface of the poriferous paranota in males of many gomphodesmid species. Not drawn to scale." figureDoi="http://doi.org/10.5281/zenodo.3928334" httpUri="https://zenodo.org/record/3928334/files/figure.png" pageId="17" pageNumber="18">Fig. 4A</figureCitation>
) present on sternum of body ring 15 (except in
<taxonomicName id="4FA73887FF9B4A1DFB0CF816BB2CF80E" authorityName="Hoffman" authorityYear="2005" box="[1242,1381,2001,2027]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="congdoni">
<emphasis id="BAD39F16FF9B4A1DFB0CF816BB2CF80E" box="[1242,1381,2001,2027]" italics="true" pageId="17" pageNumber="18">E. congdoni</emphasis>
</taxonomicName>
). Gonopod aperture asymmetrically oval, extended back between the 8
<superScript id="7FD2EE4CFF984A1EFC34FECEBDA6FEFD" attach="left" box="[994,1007,266,280]" fontSize="6" pageId="18" pageNumber="19">th</superScript>
pair of legs, quite large, posterior edge elevated behind the gonocoxae, but not in front of coxae of the 8
<superScript id="7FD2EE4CFF984A1EFC2EFEEBBA4CFED8" attach="left" box="[1016,1029,303,317]" fontSize="6" pageId="18" pageNumber="19">th</superScript>
pair of legs. Rim of aperture is thickened, variously emarginate medially. Gonocoxa slanting and shortened as usual, not produced over base of cannula. Prefemoral region without basal fossa on the median side, but deeply impressed on lateral side (where surface is membranous), making prefemur seem almost like carina. Telepodite endonodal. Process M present, varies in both size and shape (exception:
<taxonomicName id="4FA73887FF984A1EFCAEFE02BDB6FE3A" authority="Olsen &amp; Rosenmejer &amp; Enghoff, 2020" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[888,1023,453,479]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="dracarys" status="sp. nov.">
<emphasis id="BAD39F16FF984A1EFCAEFE02BDB6FE3A" box="[888,1023,453,479]" italics="true" pageId="18" pageNumber="19">E. dracarys</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF984A1EFBD3FE02BA2AFE05" box="[1029,1123,454,480]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
). Process L can be quite small, but is larger in some species. Postnodal telopodite curved dorsomesad, with a subglobose, hollow enlargement along the outer curvature. Apex of telopodite with a subterminal process of variable shape and size (e.g.,
<figureCitation id="109C5F81FF984A1EFEB0FDF1BFF5FDB5" box="[358,444,565,592]" captionStart="Fig" captionStartId="21.[189,232,1930,1956]" captionTargetBox="[354,1230,1369,1895]" captionTargetId="figure@21.[343,1238,1364,1901]" captionTargetPageId="21" captionText="Fig. 15. Emphysemastix spp., gonopod tip. A. E. dracarys Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621675). B. E. frampt Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621677). Abbreviations: so = solenomere; sp = subterminal process. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3928356" httpUri="https://zenodo.org/record/3928356/files/figure.png" pageId="18" pageNumber="19">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF984A1EFF6BFDBDBA8EFBF2" pageId="18" pageNumber="19" type="reference_group">
<paragraph id="88184304FF984A1EFF6BFDBDBFC0FD77" blockId="18.[189,393,632,659]" box="[189,393,632,659]" pageId="18" pageNumber="19">
<heading id="D350F468FF984A1EFF6BFDBDBFC0FD77" bold="true" box="[189,393,632,659]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="BAD39F16FF984A1EFF6BFDBDBFC0FD77" bold="true" box="[189,393,632,659]" pageId="18" pageNumber="19">Included species</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF984A1EFF6BFD6DBD02FD21" blockId="18.[189,843,681,931]" box="[189,843,681,708]" pageId="18" pageNumber="19">
<taxonomicName id="4FA73887FF984A1EFF6BFD6DBCEBFD26" ID-CoL="39HQZ" authority="(Attems, 1931)" baseAuthorityName="Attems" baseAuthorityYear="1931" box="[189,674,681,707]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="trepidans">
<emphasis id="BAD39F16FF984A1EFF6BFD6DBFA0FD26" box="[189,489,681,707]" italics="true" pageId="18" pageNumber="19">Emphysemastix trepidans</emphasis>
(
<bibRefCitation id="EC363EF5FF984A1EFE2EFD6DBCD0FD26" author="Attems C." box="[504,665,681,707]" pageId="18" pageNumber="19" pagination="1 - 150" refId="ref11191" refString="Attems C. 1931. Die Familie Leptodesmidae und andere Polydesmiden. Zoologica, Stuttgart 30 (70): 1 - 150." type="journal article" year="1931">Attems, 1931</bibRefCitation>
)
</taxonomicName>
(
<typeStatus id="571CFDA6FF984A1EFD67FD6DBCACFD26" box="[689,741,681,707]" pageId="18" pageNumber="19">type</typeStatus>
species)
</paragraph>
<paragraph id="88184304FF984A1EFF6BFD0ABCEBFD0C" blockId="18.[189,843,681,931]" box="[189,674,718,745]" pageId="18" pageNumber="19">
<taxonomicName id="4FA73887FF984A1EFF6BFD0ABCEBFD0C" ID-CoL="39HQV" authority="Hoffman, 2005" authorityName="Hoffman" authorityYear="2005" box="[189,674,718,745]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="congdoni">
<emphasis id="BAD39F16FF984A1EFF6BFD0ABFA1FD0D" box="[189,488,718,744]" italics="true" pageId="18" pageNumber="19">Emphysemastix congdoni</emphasis>
<bibRefCitation id="EC363EF5FF984A1EFE39FD0ABCEBFD0C" author="Hoffman R. L." box="[495,674,718,745]" pageId="18" pageNumber="19" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman, 2005</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="88184304FF984A1EFF6BFD37BD5FFCEB" blockId="18.[189,843,681,931]" box="[189,790,755,782]" pageId="18" pageNumber="19">
<taxonomicName id="4FA73887FF984A1EFF6BFD37BCFDFCE8" ID-CoL="16d88827-f339-4af8-be47-f1d00a209c13" authority="Olsen &amp; Enghoff" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[189,692,755,781]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="dracarys" status="sp. nov.">
<emphasis id="BAD39F16FF984A1EFF6BFD37BFAAFCE8" box="[189,483,755,781]" italics="true" pageId="18" pageNumber="19">Emphysemastix dracarys</emphasis>
Olsen &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF984A1EFD6DFD30BD5FFCEB" box="[699,790,756,782]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="88184304FF984A1EFF6BFCDCBCE5FCD6" blockId="18.[189,843,681,931]" box="[189,684,792,819]" pageId="18" pageNumber="19">
<taxonomicName id="4FA73887FF984A1EFF6BFCDCBCE5FCD6" ID-CoL="39HQW" authority="(Carl, 1909)" baseAuthorityName="Carl" baseAuthorityYear="1909" box="[189,684,792,819]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="flavosignatus">
<emphasis id="BAD39F16FF984A1EFF6BFCDCBC5FFCD7" box="[189,534,792,818]" italics="true" pageId="18" pageNumber="19">Emphysemastix flavosignatus</emphasis>
(
<bibRefCitation id="EC363EF5FF984A1EFDF3FCDCBCEDFCD6" author="Carl J." box="[549,676,792,819]" pageId="18" pageNumber="19" pagination="281 - 365" refId="ref11215" refString="Carl J. 1909. Reise von Dr. J. Carl im nordlichen Zentral-Afrikanischen Seengebiet. Diplopoden. Revue suisse de Zoologie 17: 281 - 365. Available from https: // www. biodiversitylibrary. org / item / 40663 # page / 293 / mode / 1 up [accessed 27 May 2020]." type="journal article" year="1909">Carl, 1909</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="88184304FF984A1EFF6BFCFABCB5FCBD" blockId="18.[189,843,681,931]" box="[189,764,830,856]" pageId="18" pageNumber="19">
<taxonomicName id="4FA73887FF984A1EFF6BFCFABCD3FCBD" ID-CoL="4d033a7e-aff2-449c-94cf-8fa82cca2e17" authority="Olsen &amp; Enghoff" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[189,666,830,856]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="frampt" status="sp. nov.">
<emphasis id="BAD39F16FF984A1EFF6BFCFABF80FCBD" box="[189,457,830,856]" italics="true" pageId="18" pageNumber="19">Emphysemastix frampt</emphasis>
Olsen &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF984A1EFD77FCFABCB5FCBD" box="[673,764,830,856]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="88184304FF984A1EFF6BFCA7BC34FC9B" blockId="18.[189,843,681,931]" box="[189,637,867,894]" pageId="18" pageNumber="19">
<taxonomicName id="4FA73887FF984A1EFF6BFCA7BC34FC9B" ID-CoL="39HQX" authority="Hoffman, 2005" authorityName="Hoffman" authorityYear="2005" box="[189,637,867,894]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="image">
<emphasis id="BAD39F16FF984A1EFF6BFCA7BF8AFC98" box="[189,451,867,893]" italics="true" pageId="18" pageNumber="19">Emphysemastix image</emphasis>
<bibRefCitation id="EC363EF5FF984A1EFE1CFCA7BC34FC9B" author="Hoffman R. L." box="[458,637,867,894]" pageId="18" pageNumber="19" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman, 2005</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="88184304FF984A1EFF6BFC4CBC88FC46" blockId="18.[189,843,681,931]" box="[189,705,904,931]" pageId="18" pageNumber="19">
<taxonomicName id="4FA73887FF984A1EFF6BFC4CBC88FC46" ID-CoL="39HQY" authority="Hoffman, 2005" authorityName="Hoffman" authorityYear="2005" box="[189,705,904,931]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="magnifrater">
<emphasis id="BAD39F16FF984A1EFF6BFC4CBC4EFC47" box="[189,519,904,930]" italics="true" pageId="18" pageNumber="19">Emphysemastix magnifrater</emphasis>
<bibRefCitation id="EC363EF5FF984A1EFDD8FC4CBC88FC46" author="Hoffman R. L." box="[526,705,904,931]" pageId="18" pageNumber="19" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman, 2005</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="88184304FF984A1EFF6BFC0FBF1FFC00" blockId="18.[189,342,971,997]" box="[189,342,971,997]" pageId="18" pageNumber="19">
<heading id="D350F468FF984A1EFF6BFC0FBF1FFC00" bold="true" box="[189,342,971,997]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="BAD39F16FF984A1EFF6BFC0FBF1FFC00" bold="true" box="[189,342,971,997]" pageId="18" pageNumber="19">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF984A1EFF6BFC38BA8EFBF2" blockId="18.[189,1223,1020,1047]" box="[189,1223,1020,1047]" pageId="18" pageNumber="19">
<collectingCountry id="F0B00394FF984A1EFF6BFC38BF63FBF3" box="[189,298,1020,1046]" name="Tanzania" pageId="18" pageNumber="19">Tanzania</collectingCountry>
. None of the previously described species are known from the Udzungwa Mts.
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF984A1FFF6BFB84BB3EFE75" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="key">
<paragraph id="88184304FF984A1EFF6BFB84BD8FFBBF" blockId="18.[189,966,1087,1114]" box="[189,966,1087,1114]" pageId="18" pageNumber="19">
<emphasis id="BAD39F16FF984A1EFF6BFB84BC1AFBBF" bold="true" box="[189,595,1087,1114]" pageId="18" pageNumber="19">
Key to species of
<taxonomicName id="4FA73887FF984A1EFE42FBFBBC1AFBBF" ID-CoL="7PD53" authorityName="Hoffman" authorityYear="1966" box="[404,595,1087,1114]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="BAD39F16FF984A1EFE42FBFBBC1AFBBF" bold="true" box="[404,595,1087,1114]" italics="true" pageId="18" pageNumber="19">Emphysemastix</emphasis>
</taxonomicName>
</emphasis>
(modified after
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)
</paragraph>
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<paragraph id="88184304FF984A1EFF6BFBB4BB3EFB4A" blockId="18.[189,1399,1136,1274]" pageId="18" pageNumber="19">
<key id="FC361435FF984A1EFF6BFBB4BB3EFB4A" pageId="18" pageNumber="19">
<keyLead id="3356E031FF984A1EFF6BFBB4BB3EFB4A" pageId="18" pageNumber="19">
1. Basal setose side of gonopod prefemur unmodified, as seen in mesal aspect not extended ventrad as a distinct subtriangular lobe (
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: fig. 201) ..................................................................... 2
</keyLead>
</key>
</paragraph>
<paragraph id="88184304FF984A1EFF17FB7FBB3EFB1F" blockId="18.[189,1399,1136,1274]" pageId="18" pageNumber="19">
<keyLead id="3356E031FF984A1EFF17FB7FBB3EFB1F" pageId="18" pageNumber="19">
– Basal setose side of prefemur projected ventrad as a more (
<bibRefCitation id="EC363EF5FF984A1EFC4EFB7EBA00FB30" author="Hoffman R. L." box="[920,1097,1210,1237]" pageId="18" pageNumber="19" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman 2005</bibRefCitation>
: fig. 204) or less (
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,
<figureCitation id="109C5F81FF984A1EFF3CFB24BF41FB1F" box="[234,264,1248,1274]" captionStart="Fig" captionStartId="23.[189,232,1930,1956]" captionTargetBox="[288,1288,265,1886]" captionTargetId="figure@23.[288,1299,265,1886]" captionTargetPageId="23" captionText="Fig. 17. Emphysemastix frampt Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621677), left gonopod. A–B. Mesal view. C–D. Lateral view. E–F. Dorsal view. Abbreviations: L = Process L; M = Process M; rl = rounded lobe; sg = subglobose enlargement. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3928360" httpUri="https://zenodo.org/record/3928360/files/figure.png" pageId="18" pageNumber="19">17</figureCitation>
) distinct subtriangular lobe ........................................................................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="33535BA1FF984A1EFF6BFAE7BB3EFA1D" pageId="18" pageNumber="19">
<paragraph id="88184304FF984A1EFF6BFAE7BB3EFA6D" blockId="18.[189,1399,1315,1528]" pageId="18" pageNumber="19">
<keyLead id="3356E031FF984A1EFF6BFAE7BB3EFA6D" pageId="18" pageNumber="19">
2. Postnodal telopodite very short, scarcely longer than distal lobe of nodus; nodal process M long and slender, projecting dorsally well beyond telopodite (
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: fig. 201); sterna with sharp transverse carinae, body ring 15 with sternal paxillus............................
<taxonomicName id="4FA73887FF984A1EFBFDFAAABB3EFA6D" ID-CoL="39HQZ" authority="(Attems, 1931)" baseAuthorityName="Attems" baseAuthorityYear="1931" box="[1067,1399,1389,1416]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="trepidans">
<emphasis id="BAD39F16FF984A1EFBFDFAAABAF4FA6D" bold="true" box="[1067,1213,1389,1416]" italics="true" pageId="18" pageNumber="19">E. trepidans</emphasis>
(
<bibRefCitation id="EC363EF5FF984A1EFB1BFAAABB27FA6D" author="Attems C." box="[1229,1390,1390,1416]" pageId="18" pageNumber="19" pagination="1 - 150" refId="ref11191" refString="Attems C. 1931. Die Familie Leptodesmidae und andere Polydesmiden. Zoologica, Stuttgart 30 (70): 1 - 150." type="journal article" year="1931">Attems, 1931</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="88184304FF984A1EFF17FA57BB3EFA1D" blockId="18.[189,1399,1315,1528]" pageId="18" pageNumber="19">
<keyLead id="3356E031FF984A1EFF17FA57BB3EFA1D" pageId="18" pageNumber="19">
– Postnodal telopodite 3–4 × as long as distal lobe of nodus; nodal process M small and short, not extending beyond edge of telopodite (
<bibRefCitation id="EC363EF5FF984A1EFD66FA7CBD2DFA36" author="Hoffman R. L." box="[688,868,1464,1491]" pageId="18" pageNumber="19" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman 2005</bibRefCitation>
: fig. 202); sterna without transverse carinae, 15
<superScript id="7FD2EE4CFF984A1EFEDEFA1FBF5CFA0C" attach="left" box="[264,277,1499,1513]" fontSize="6" pageId="18" pageNumber="19">th</superScript>
body ring without paxillus ...............................................................
<taxonomicName id="4FA73887FF984A1EFBFCFA19BB3EFA1D" ID-CoL="39HQV" authority="Hoffman, 2005" authorityName="Hoffman" authorityYear="2005" box="[1066,1399,1500,1528]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="congdoni">
<emphasis id="BAD39F16FF984A1EFBFCFA19BAF5FA12" bold="true" box="[1066,1212,1500,1527]" italics="true" pageId="18" pageNumber="19">E. congdoni</emphasis>
<bibRefCitation id="EC363EF5FF984A1EFB12FA19BB3EFA1D" author="Hoffman R. L." box="[1220,1399,1501,1528]" pageId="18" pageNumber="19" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman, 2005</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="33535BA1FF984A1EFF6BF9E5BB3EF985" pageId="18" pageNumber="19">
<paragraph id="88184304FF984A1EFF6BF9E5BB3EF9DE" blockId="18.[189,1399,1568,1632]" box="[189,1399,1568,1595]" pageId="18" pageNumber="19">
<keyLead id="3356E031FF984A1EFF6BF9E5BB3EF9DE" box="[189,1399,1568,1595]" pageId="18" pageNumber="19">
3. Nodal process M absent (
<figureCitation id="109C5F81FF984A1EFDC3F9E4BC23F9DE" box="[533,618,1568,1595]" captionStart="Fig" captionStartId="20.[189,232,1930,1956]" captionTargetBox="[259,1314,265,1901]" captionTargetId="figure@20.[259,1331,265,1901]" captionTargetPageId="20" captionText="Fig. 14. Emphysemastic dracarys Olsen &amp; Enghoff sp. nov., holotype, ♂, left gonopod (NHMD 621675). A–B. Mesal view. C–D. Lateral view. E–F. Dorsal view. Abbreviations: L = Process L; rl = rounded lobe; sg = subglobose enlargement; sp = subterminal process. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928354" httpUri="https://zenodo.org/record/3928354/files/figure.png" pageId="18" pageNumber="19">Fig. 14</figureCitation>
)............................................
<taxonomicName id="4FA73887FF984A1EFC6CF9E5BB5CF9DF" ID-CoL="16d88827-f339-4af8-be47-f1d00a209c13" authority="Olsen &amp; Enghoff" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[954,1301,1568,1595]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="dracarys" status="sp. nov.">
<emphasis id="BAD39F16FF984A1EFC6CF9E5BA0DF9DE" bold="true" box="[954,1092,1568,1595]" italics="true" pageId="18" pageNumber="19">E. dracarys</emphasis>
Olsen &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF984A1EFACAF9E5BB3EF9DE" box="[1308,1399,1569,1595]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="88184304FF984A1EFF17F982BB3EF985" blockId="18.[189,1399,1568,1632]" box="[193,1399,1606,1632]" pageId="18" pageNumber="19">
<keyLead id="3356E031FF984A1EFF17F982BB3EF985" box="[193,1399,1606,1632]" pageId="18" pageNumber="19">– Nodal process M present ................................................................................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="33535BA1FF984A1EFF6BF94DBB3EF8DC" pageId="18" pageNumber="19">
<paragraph id="88184304FF984A1EFF6BF94DBB3EF90B" blockId="18.[189,1399,1673,1849]" pageId="18" pageNumber="19">
<keyLead id="3356E031FF984A1EFF6BF94DBB3EF90B" pageId="18" pageNumber="19">
4. Nodal process M broadly laminate, apical edge truncate and serrate (
<bibRefCitation id="EC363EF5FF984A1EFB67F94DBB3AF941" author="Hoffman R. L." box="[1201,1395,1673,1700]" pageId="18" pageNumber="19" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman 2005</bibRefCitation>
: fig. 205); distal nodal lobe elongate and subrectangular in mesal aspect; body width
<quantity id="4F5FEEE1FF984A1EFF3CF910BF74F90B" box="[234,317,1748,1774]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="18" pageNumber="19" unit="mm" value="14.0">14 mm</quantity>
................................................................................................
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<emphasis id="BAD39F16FF984A1EFBDFF910BAF5F908" bold="true" box="[1033,1212,1747,1774]" italics="true" pageId="18" pageNumber="19">E. magnifrater</emphasis>
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</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="88184304FF984A1EFF17F93DBB3EF8DC" blockId="18.[189,1399,1673,1849]" pageId="18" pageNumber="19">
<keyLead id="3356E031FF984A1EFF17F93DBB3EF8DC" pageId="18" pageNumber="19">– Nodal process M slender, subterete, gradually acuminate; distal nodal lobe shorter and subovoid; body narrower.................................................................................................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="33535BA1FF984A1EFF6BF8A5BB3EF809" pageId="18" pageNumber="19">
<paragraph id="88184304FF984A1EFF6BF8A5BB3EF845" blockId="18.[189,1399,1889,2028]" pageId="18" pageNumber="19">
<keyLead id="3356E031FF984A1EFF6BF8A5BB3EF845" pageId="18" pageNumber="19">5. Nodal process L long, visible in mesal aspect; postnodal telopodite of moderate length, not extending proximad beyond base of cannula ..................................................................................................... 6</keyLead>
</paragraph>
<paragraph id="88184304FF984A1EFF17F868BB3EF809" blockId="18.[189,1399,1889,2028]" pageId="18" pageNumber="19">
<keyLead id="3356E031FF984A1EFF17F868BB3EF809" pageId="18" pageNumber="19">
– Nodal process L very small, not visible in mesal aspect; postnodal telopodite very long, extending proximad far beyond base of cannula (
<bibRefCitation id="EC363EF5FF984A1EFD7BF815BD16F809" author="Hoffman R. L." box="[685,863,2001,2028]" pageId="18" pageNumber="19" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman 2005</bibRefCitation>
: fig. 207).................
<taxonomicName id="4FA73887FF984A1EFB87F815BB3EF809" ID-CoL="39HQX" authority="Hoffman, 2005" authorityName="Hoffman" authorityYear="2005" box="[1105,1399,2001,2028]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="image">
<emphasis id="BAD39F16FF984A1EFB87F815BAF5F80E" bold="true" box="[1105,1212,2001,2027]" italics="true" pageId="18" pageNumber="19">E. image</emphasis>
<bibRefCitation id="EC363EF5FF984A1EFB12F815BB3EF809" author="Hoffman R. L." box="[1220,1399,2001,2028]" pageId="18" pageNumber="19" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman, 2005</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="33535BA1FF994A1FFF6BFECFBB3EFE75" pageId="19" pageNumber="20">
<paragraph id="88184304FF994A1FFF6BFECFBB3FFEAC" blockId="19.[189,1399,267,400]" pageId="19" pageNumber="20">
<keyLead id="3356E031FF994A1FFF6BFECFBB3FFEAC" pageId="19" pageNumber="20">
6. Body width 9.0–
<quantity id="4F5FEEE1FF994A1FFE7BFEC8BC4EFEC3" box="[429,519,268,294]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="19" pageNumber="20" unit="mm" value="9.3">9.3 mm</quantity>
. Paxillus triangular, pointed. Gonopodal nodus subrectangular, processes M and L relatively short, straight (
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: fig. 206).....................
<taxonomicName id="4FA73887FF994A1FFBC8FEEBBB3FFEAC" ID-CoL="39HQW" authority="(Carl, 1909)" baseAuthorityName="Carl" baseAuthorityYear="1909" box="[1054,1398,302,329]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="flavosignatus">
<emphasis id="BAD39F16FF994A1FFBC8FEEBBAA9FEAC" bold="true" box="[1054,1248,302,329]" italics="true" pageId="19" pageNumber="20">E. flavosignatus</emphasis>
(
<bibRefCitation id="EC363EF5FF994A1FFB39FEEBBB27FEAC" author="Carl J." box="[1263,1390,303,329]" pageId="19" pageNumber="20" pagination="281 - 365" refId="ref11215" refString="Carl J. 1909. Reise von Dr. J. Carl im nordlichen Zentral-Afrikanischen Seengebiet. Diplopoden. Revue suisse de Zoologie 17: 281 - 365. Available from https: // www. biodiversitylibrary. org / item / 40663 # page / 293 / mode / 1 up [accessed 27 May 2020]." type="journal article" year="1909">Carl, 1909</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="88184304FF994A1FFF17FE97BB3EFE75" blockId="19.[189,1399,267,400]" pageId="19" pageNumber="20">
<keyLead id="3356E031FF994A1FFF17FE97BB3EFE75" pageId="19" pageNumber="20">
– Body width
<quantity id="4F5FEEE1FF994A1FFEABFE97BFA3FE88" box="[381,490,339,365]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="19" pageNumber="20" unit="mm" value="13.1">13.1 mm</quantity>
. Paxillus rounded. Gonopodal nodus with slightly diverging sides, processes M and L longer, more curved (
<figureCitation id="109C5F81FF994A1FFD84FEB2BCEFFE75" box="[594,678,374,400]" captionStart="Fig" captionStartId="23.[189,232,1930,1956]" captionTargetBox="[288,1288,265,1886]" captionTargetId="figure@23.[288,1299,265,1886]" captionTargetPageId="23" captionText="Fig. 17. Emphysemastix frampt Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621677), left gonopod. A–B. Mesal view. C–D. Lateral view. E–F. Dorsal view. Abbreviations: L = Process L; M = Process M; rl = rounded lobe; sg = subglobose enlargement. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3928360" httpUri="https://zenodo.org/record/3928360/files/figure.png" pageId="19" pageNumber="20">Fig. 17</figureCitation>
).......................................
<taxonomicName id="4FA73887FF994A1FFC06FEB2BB5CFE75" ID-CoL="4d033a7e-aff2-449c-94cf-8fa82cca2e17" authority="Olsen &amp; Enghoff" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[976,1301,373,400]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="frampt" status="sp. nov.">
<emphasis id="BAD39F16FF994A1FFC06FEB2BA0DFE6A" bold="true" box="[976,1092,373,400]" italics="true" pageId="19" pageNumber="20">E. frampt</emphasis>
Olsen &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF994A1FFACAFEB2BB3EFE75" box="[1308,1399,374,400]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>

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<document id="D677908E5B893E88D8834A1B96F5E690" ID-CLB-Dataset="21573" ID-DOI="10.5852/ejt.2020.675" ID-GBIF-Dataset="39cc6284-8ad7-47f0-bf5d-2122174905ed" ID-Zenodo-Dep="3928326" ID-ZooBank="88FFA67B-C2DE-43C6-ACB1-44EDCF119EBA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1593696236104" checkinUser="valdenar" docAuthor="Olsen, Sissel Anna, Rosenmejer, Trine &amp; Enghoff, Henrik" docDate="2020" docId="000EF212FF9F4A14FE23FB8EBA11FD28" docLanguage="en" docName="ejt_675_olsen_rosenmejer_enghoff.pdf" docOrigin="European Journal of Taxonomy 675" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Emphysemastix frampt Olsen &amp; Rosenmejer &amp; Enghoff 2020, sp. nov." docType="treatment" docUuid="CBF0C445-4D5A-48D1-83EA-C509E0567476" docUuidSource="ZooBank" docVersion="8" lastPageNumber="25" masterDocId="FC378A6AFF8A4A0CFFD6FFC4BE49FFE5" masterDocTitle="A mountain of millipedes IX: Species of the family Gomphodesmidae from the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida)" masterLastPageNumber="35" masterPageNumber="1" pageNumber="22" updateTime="1698842977217" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9B8E504DE360C1156C75F31396EBA130">A mountain of millipedes IX: Species of the family Gomphodesmidae from the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida)</mods:title>
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<mods:namePart id="AE46DB42569E88891F806D84F3202164">Olsen, Sissel Anna</mods:namePart>
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<mods:affiliation id="EC7D6D4A4D88D78E09CD7ABB0DBB7113">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 København Ø, Denmark</mods:affiliation>
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<mods:namePart id="FFEFF21F9BDF72273E7FFBF3231C4B85">Rosenmejer, Trine</mods:namePart>
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<mods:affiliation id="887349BC326AC94D678FC0904A4ABFA7">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 København Ø, Denmark</mods:affiliation>
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<mods:namePart id="22CDED9D4547E8340D6292C5F9F5F30F">Enghoff, Henrik</mods:namePart>
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<mods:affiliation id="F6E32CDC6EF87C4B74F3F5A4DBE75DFF">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 København Ø, Denmark</mods:affiliation>
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<treatment id="000EF212FF9F4A14FE23FB8EBA11FD28" ID-DOI="http://doi.org/10.5281/zenodo.4332925" ID-GBIF-Taxon="164870653" ID-Zenodo-Dep="4332925" LSID="urn:lsid:zoobank.org:act:CBF0C445-4D5A-48D1-83EA-C509E0567476" httpUri="http://treatment.plazi.org/id/000EF212FF9F4A14FE23FB8EBA11FD28" lastPageId="24" lastPageNumber="25" pageId="21" pageNumber="22">
<subSubSection id="C0BD108FFF9F4A19FE23FB8EBA76FB83" box="[501,1087,1098,1126]" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="88184304FF9F4A19FE23FB8EBA76FB83" blockId="21.[366,1222,1098,1196]" box="[501,1087,1098,1126]" pageId="21" pageNumber="22">
<heading id="D350F468FF9F4A19FE23FB8EBA76FB83" box="[501,1087,1098,1126]" centered="true" fontSize="11" level="2" pageId="21" pageNumber="22" reason="2">
<taxonomicName id="4FA73887FF9F4A19FE23FB8EBD97FB80" ID-CoL="4d033a7e-aff2-449c-94cf-8fa82cca2e17" authority="Olsen &amp; Enghoff" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[501,990,1098,1125]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="frampt" status="sp. nov.">
<emphasis id="BAD39F16FF9F4A19FE23FB8EBD45FB81" bold="true" box="[501,780,1098,1125]" italics="true" pageId="21" pageNumber="22">Emphysemastix frampt</emphasis>
Olsen &amp; Enghoff
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<taxonomicNameLabel id="A1E0226DFF9F4A19FC32FB88BA76FB83" box="[996,1087,1100,1126]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<paragraph id="88184304FF9F4A19FD7CFB55BDC3FB4E" blockId="21.[366,1222,1098,1196]" box="[682,906,1169,1196]" pageId="21" pageNumber="22">
<figureCitation id="109C5F81FF9F4A19FD7CFB55BCBCFB49" box="[682,757,1169,1196]" captionStart="Fig" captionStartId="2.[189,232,1966,1992]" captionTargetBox="[199,1398,1100,1897]" captionTargetId="figure@2.[194,1482,1044,1958]" captionTargetPageId="2" captionText="Fig. 2. Map of the Udzungwa Mts showing collection sites for gomphodesmids. Based on Marshall et al. (2010: fig. 1)." figureDoi="http://doi.org/10.5281/zenodo.3928330" httpUri="https://zenodo.org/record/3928330/files/figure.png" pageId="21" pageNumber="22">Figs 2</figureCitation>
,
<figureCitation id="109C5F81FF9F4A19FCD7FB55BD7DFB49" box="[769,820,1169,1196]" captionStart="Fig" captionStartId="21.[189,232,1930,1956]" captionTargetBox="[354,1230,1369,1895]" captionTargetId="figure@21.[343,1238,1364,1901]" captionTargetPageId="21" captionText="Fig. 15. Emphysemastix spp., gonopod tip. A. E. dracarys Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621675). B. E. frampt Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621677). Abbreviations: so = solenomere; sp = subterminal process. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3928356" httpUri="https://zenodo.org/record/3928356/files/figure.png" pageId="21" pageNumber="22">15B</figureCitation>
,
<figureCitation id="109C5F81FF9F4A19FC96FB55BDC3FB4E" box="[832,906,1169,1195]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="22.[189,232,1964,1990]" captionStartId-1="23.[189,232,1930,1956]" captionTargetBox-0="[476,1116,1456,1914]" captionTargetBox-1="[288,1288,265,1886]" captionTargetId-0="figure@22.[462,1124,1450,1924]" captionTargetId-1="figure@23.[288,1299,265,1886]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionText-0="Fig. 16. Emphysemastix frampt Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621677). Scale bar: 5 mm." captionText-1="Fig. 17. Emphysemastix frampt Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621677), left gonopod. A–B. Mesal view. C–D. Lateral view. E–F. Dorsal view. Abbreviations: L = Process L; M = Process M; rl = rounded lobe; sg = subglobose enlargement. Scale bars: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3928358" figureDoi-1="http://doi.org/10.5281/zenodo.3928360" httpUri-0="https://zenodo.org/record/3928358/files/figure.png" httpUri-1="https://zenodo.org/record/3928360/files/figure.png" pageId="21" pageNumber="22">16–17</figureCitation>
</paragraph>
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<subSubSection id="C0BD108FFF9F4A19FF6BFB17BC71FADA" pageId="21" pageNumber="22" type="diagnosis">
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<heading id="D350F468FF9F4A19FF6BFB17BF7FFB08" bold="true" box="[189,310,1235,1261]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="BAD39F16FF9F4A19FF6BFB17BF7FFB08" bold="true" box="[189,310,1235,1261]" pageId="21" pageNumber="22">Diagnosis</emphasis>
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<paragraph id="88184304FF9F4A19FF6BFAC5BC71FADA" blockId="21.[189,1399,1281,1343]" pageId="21" pageNumber="22">Differs from the other species of the genus by having a rounded, rather than triangular paxillus. Nodal processes M and L both present.</paragraph>
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<caption id="DCD8138CFF9F4A19FF6BF84EBDFFF80E" ID-DOI="http://doi.org/10.5281/zenodo.3928356" ID-Zenodo-Dep="3928356" httpUri="https://zenodo.org/record/3928356/files/figure.png" pageId="21" pageNumber="22" startId="21.[189,232,1930,1956]" targetBox="[354,1230,1369,1895]" targetPageId="21">
<paragraph id="88184304FF9F4A19FF6BF84EBDFFF80E" blockId="21.[189,1399,1930,2027]" pageId="21" pageNumber="22">
<emphasis id="BAD39F16FF9F4A19FF6BF84EBF57F841" bold="true" box="[189,286,1930,1956]" pageId="21" pageNumber="22">Fig. 15.</emphasis>
<taxonomicName id="4FA73887FF9F4A19FEFCF84EBFA9F841" authorityName="Hoffman" authorityYear="1966" box="[298,480,1930,1956]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="BAD39F16FF9F4A19FEFCF84EBFA9F841" box="[298,480,1930,1956]" italics="true" pageId="21" pageNumber="22">Emphysemastix</emphasis>
</taxonomicName>
spp., gonopod tip.
<emphasis id="BAD39F16FF9F4A19FD03F84EBCA3F841" bold="true" box="[725,746,1930,1956]" pageId="21" pageNumber="22">A</emphasis>
.
<taxonomicName id="4FA73887FF9F4A19FD2BF84FBA23F841" authority="Olsen &amp; Enghoff" authorityName="Olsen &amp; Enghoff" authorityYear="2020" box="[765,1130,1930,1956]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastic" kingdom="Animalia" order="Polydesmida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="dracarys" status="sp. nov.">
<emphasis id="BAD39F16FF9F4A19FD2BF84FBDC2F841" box="[765,907,1930,1956]" italics="true" pageId="21" pageNumber="22">E. dracarys</emphasis>
Olsen &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF9F4A19FBA0F84EBA9CF841" box="[1142,1237,1930,1956]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NHMD 621675).
<emphasis id="BAD39F16FF9F4A19FE40F869BFE3F822" bold="true" box="[406,426,1965,1991]" pageId="21" pageNumber="22">B</emphasis>
.
<taxonomicName id="4FA73887FF9F4A19FE6EF86ABCBDF822" authority="Olsen &amp; Enghoff" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[440,756,1965,1991]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="frampt" status="sp. nov.">
<emphasis id="BAD39F16FF9F4A19FE6EF86ABC6CF822" box="[440,549,1965,1991]" italics="true" pageId="21" pageNumber="22">E. frampt</emphasis>
Olsen &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF9F4A19FD2CF86ABD1DF82D" box="[762,852,1966,1992]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NHMD 621677). Abbreviations:
<emphasis id="BAD39F16FF9F4A19FF6BF815BE9EF80F" box="[189,215,2001,2026]" italics="true" pageId="21" pageNumber="22">so</emphasis>
= solenomere;
<emphasis id="BAD39F16FF9F4A19FE5BF815BFEEF80F" box="[397,423,2001,2026]" italics="true" pageId="21" pageNumber="22">sp</emphasis>
= subterminal process. Scale bars: 0.25 mm.
</paragraph>
</caption>
<subSubSection id="C0BD108FFF9C4A1AFF6BFECFBA9AFE92" pageId="22" pageNumber="23" type="etymology">
<paragraph id="88184304FF9C4A1AFF6BFECFBF0DFEC0" blockId="22.[189,324,267,293]" box="[189,324,267,293]" pageId="22" pageNumber="23">
<heading id="D350F468FF9C4A1AFF6BFECFBF0DFEC0" bold="true" box="[189,324,267,293]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="BAD39F16FF9C4A1AFF6BFECFBF0DFEC0" bold="true" box="[189,324,267,293]" pageId="22" pageNumber="23">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF9C4A1AFF6BFEFDBA9AFE92" blockId="22.[189,1399,313,375]" pageId="22" pageNumber="23">The species is named after Kingseeker Frampt from the Dark Souls video game series, due to the gonopods’ resemblance to the creature. The name is to be treated as a noun in apposition.</paragraph>
</subSubSection>
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<paragraph id="88184304FF9C4A1AFF6BFE5ABFE3FE5D" blockId="22.[189,426,414,440]" box="[189,426,414,440]" pageId="22" pageNumber="23">
<heading id="D350F468FF9C4A1AFF6BFE5ABFE3FE5D" bold="true" box="[189,426,414,440]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="BAD39F16FF9C4A1AFF6BFE5ABFE3FE5D" bold="true" box="[189,426,414,440]" pageId="22" pageNumber="23">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF9C4A1AFF3CFE09BF15FE02" blockId="22.[189,1399,461,593]" box="[234,348,461,487]" pageId="22" pageNumber="23">
<emphasis id="BAD39F16FF9C4A1AFF3CFE09BF15FE02" bold="true" box="[234,348,461,487]" pageId="22" pageNumber="23">
<typeStatus id="571CFDA6FF9C4A1AFF3CFE09BF15FE02" box="[234,348,461,487]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="88184304FF9C4A1AFF6BFE34BF37FDB5" blockId="22.[189,1399,461,593]" pageId="22" pageNumber="23">
<materialsCitation id="38CF4959FF9C4A1AFF6BFE34BF37FDB5" ID-GBIF-Occurrence="2803530305" collectingDate="1993-12" collectionCode="NHMD" collectorName="L. A. Hansen &amp; J. O. Svendsen" country="Tanzania" county="Mahenge District" elevation="1700" latitude="-7.8" location="Udekwa Village" longLatPrecision="1302" longitude="36.35" municipality="West Kilombero Scarp" pageId="22" pageNumber="23" specimenCode="NHMD 621677" specimenCount="1" stateProvince="Iringa Region" typeStatus="holotype">
<collectingCountry id="F0B00394FF9C4A1AFF6BFE34BF1BFDEF" box="[189,338,496,522]" name="Tanzania" pageId="22" pageNumber="23">TANZANIA</collectingCountry>
•
<specimenCount id="9EA1888DFF9C4A1AFEBDFE2BBFC8FDEE" box="[363,385,495,523]" pageId="22" pageNumber="23" type="male">♂</specimenCount>
;
<collectingRegion id="4A638DE6FF9C4A1AFE47FE34BC7EFDEF" box="[401,567,496,522]" country="Tanzania" name="Iringa" pageId="22" pageNumber="23">Iringa Region</collectingRegion>
,
<collectingCounty id="61793B88FF9C4A1AFD94FE34BD5BFDEF" box="[578,786,496,522]" pageId="22" pageNumber="23">Mahenge District</collectingCounty>
,
<locationDeviation id="99AA5A26FF9C4A1AFCCBFE34BA1EFDEF" box="[797,1111,496,522]" pageId="22" pageNumber="23">West Kilombero Scarp FR</locationDeviation>
, Nyumbanitu Mts, S of?
<location id="8D7815DFFF9C4A1AFF6BFDD7BFC9FDC8" LSID="urn:lsid:plazi:treatment:000EF212FF9F4A14FE23FB8EBA11FD28:8D7815DFFF9C4A1AFF6BFDD7BFC9FDC8" box="[189,384,531,557]" country="Tanzania" county="Mahenge District" latitude="-7.8" longLatPrecision="1302" longitude="36.35" municipality="West Kilombero Scarp" name="Udekwa Village" pageId="22" pageNumber="23" stateProvince="Iringa Region">Udekwa Village</location>
;
<geoCoordinate id="ED9325C3FF9C4A1AFE58FDD5BFBEFDC8" box="[398,503,529,557]" degrees="07" direction="south" minutes="48" orientation="latitude" pageId="22" pageNumber="23" precision="925" value="-7.8">07°48′ S</geoCoordinate>
,
<geoCoordinate id="ED9325C3FF9C4A1AFDD0FDD5BC3BFDCB" box="[518,626,529,558]" degrees="36" direction="east" minutes="21" orientation="longitude" pageId="22" pageNumber="23" precision="925" value="36.35">36°21′ E</geoCoordinate>
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a.s.l.
</elevation>
;
<date id="FC1965C4FF9C4A1AFCF8FDD0BDE5FDC8" box="[814,940,531,558]" pageId="22" pageNumber="23" value="1993-12">
<collectingDate id="EC5D9C2CFF9C4A1AFCF8FDD0BDE5FDC8" box="[814,940,531,558]" pageId="22" pageNumber="23" value="1993-12">Dec. 1993</collectingDate>
</date>
;
<collectorName id="255226D2FF9C4A1AFC6DFDD7BA1BFDCB" box="[955,1106,531,558]" pageId="22" pageNumber="23">L.A. Hansen</collectorName>
and
<collectorName id="255226D2FF9C4A1AFB46FDD7BB70FDC8" box="[1168,1337,531,557]" pageId="22" pageNumber="23">J.O. Svendsen</collectorName>
leg.;
<specimenCode id="D801EB7FFF9C4A1AFF6BFDF3BF33FDB5" box="[189,378,566,593]" collectionCode="NHMD" pageId="22" pageNumber="23">NHMD 621677</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF9C4A14FF6BFDBCBDAAFE7D" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23" type="description">
<paragraph id="88184304FF9C4A1AFF6BFDBCBF07FD77" blockId="22.[189,334,632,658]" box="[189,334,632,658]" pageId="22" pageNumber="23">
<heading id="D350F468FF9C4A1AFF6BFDBCBF07FD77" bold="true" box="[189,334,632,658]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="BAD39F16FF9C4A1AFF6BFDBCBF07FD77" bold="true" box="[189,334,632,658]" pageId="22" pageNumber="23">Description</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF9C4A1AFF3CFD63BF63FD24" blockId="22.[189,1399,679,775]" box="[234,298,679,705]" pageId="22" pageNumber="23">
<emphasis id="BAD39F16FF9C4A1AFF3CFD63BF63FD24" bold="true" box="[234,298,679,705]" pageId="22" pageNumber="23">Male</emphasis>
</paragraph>
<paragraph id="88184304FF9C4A1AFF6BFD0EBC9EFCE2" blockId="22.[189,1399,679,775]" pageId="22" pageNumber="23">
SIZE AND SHAPE. A large member of the genus, rather wide compared to length. Body length ca
<quantity id="4F5FEEE1FF9C4A1AFACBFD0DBB3BFD01" box="[1309,1394,713,740]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="22" pageNumber="23" unit="mm" value="55.0">55 mm</quantity>
. Maximum width
<quantity id="4F5FEEE1FF9C4A1AFE5FFD29BFBDFCE2" box="[393,500,749,775]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="22" pageNumber="23" unit="mm" value="13.1">13.1 mm</quantity>
. W/L ratio ca 24%.
</paragraph>
<paragraph id="88184304FF9C4A1AFF6BFCEEBA9DFC8D" blockId="22.[189,1398,810,872]" pageId="22" pageNumber="23">
COLOUR (
<figureCitation id="109C5F81FF9C4A1AFEE7FCEEBFC1FCA0" box="[305,392,810,837]" captionStart="Fig" captionStartId="22.[189,232,1964,1990]" captionTargetBox="[476,1116,1456,1914]" captionTargetId="figure@22.[462,1124,1450,1924]" captionTargetPageId="22" captionText="Fig. 16. Emphysemastix frampt Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621677). Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928358" httpUri="https://zenodo.org/record/3928358/files/figure.png" pageId="22" pageNumber="23">Fig. 16</figureCitation>
). Metazona orange-brown, with lighter colour on posterior part and on paranota; legs darker and more intensely/vividly orange. Prozona as metazona but without lighter areas.
</paragraph>
<paragraph id="88184304FF9C4A1AFF6BFC4FBA29FBB0" blockId="22.[189,1399,907,1110]" pageId="22" pageNumber="23">BODY RINGS. Paranota set high on sides, dorsum moderately convex. Anterior edge of ring 2–4 straight, anterior edge of rings 5–20 with gradually larger posterior-facing edge. Torus present, but small and indistinct. Stricture poorly defined, but with sharp edge in front of anterior spiracles. Pleurosternal carinae present, knobby, distinct from ring 3, increasing in size towards ring 8–9, afterwards decreasing in size towards posterior end, indistinct on ring 18. Transverse sternal carinae as typical of the genus, but very small on anterior of ring 9. Anterior edge of paxillus rounded, with ridges.</paragraph>
<paragraph id="88184304FF9C4A1AFF6BFBBDBC79FB71" blockId="22.[189,560,1145,1172]" box="[189,560,1145,1172]" pageId="22" pageNumber="23">
LEGS. Relatively long (
<figureCitation id="109C5F81FF9C4A1AFE1DFBBDBC6AFB71" box="[459,547,1145,1172]" captionStart="Fig" captionStartId="22.[189,232,1964,1990]" captionTargetBox="[476,1116,1456,1914]" captionTargetId="figure@22.[462,1124,1450,1924]" captionTargetPageId="22" captionText="Fig. 16. Emphysemastix frampt Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621677). Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3928358" httpUri="https://zenodo.org/record/3928358/files/figure.png" pageId="22" pageNumber="23">Fig. 16</figureCitation>
).
</paragraph>
<paragraph id="88184304FF9C4A1AFF6BFB73BF20FB11" blockId="22.[189,1398,1206,1268]" pageId="22" pageNumber="23">HYPOPROCT. Paramediean tubercles small, not extended past edge of sclerite. Median projection large and elongated.</paragraph>
<paragraph id="88184304FF9C4A14FF6BFAD3BDAAFE7D" blockId="22.[189,1399,1303,1400]" lastBlockId="24.[189,1399,267,408]" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23">
GONOPOD (
<figureCitation id="109C5F81FF9C4A1AFE92FAD3BFF8FAD7" box="[324,433,1303,1330]" captionStart="Fig" captionStartId="21.[189,232,1930,1956]" captionTargetBox="[354,1230,1369,1895]" captionTargetId="figure@21.[343,1238,1364,1901]" captionTargetPageId="21" captionText="Fig. 15. Emphysemastix spp., gonopod tip. A. E. dracarys Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621675). B. E. frampt Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621677). Abbreviations: so = solenomere; sp = subterminal process. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3928356" httpUri="https://zenodo.org/record/3928356/files/figure.png" pageId="22" pageNumber="23">Figs 15B</figureCitation>
,
<figureCitation id="109C5F81FF9C4A1AFE6BFAD3BF94FAD4" box="[445,477,1303,1329]" captionStart="Fig" captionStartId="23.[189,232,1930,1956]" captionTargetBox="[288,1288,265,1886]" captionTargetId="figure@23.[288,1299,265,1886]" captionTargetPageId="23" captionText="Fig. 17. Emphysemastix frampt Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621677), left gonopod. A–B. Mesal view. C–D. Lateral view. E–F. Dorsal view. Abbreviations: L = Process L; M = Process M; rl = rounded lobe; sg = subglobose enlargement. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3928360" httpUri="https://zenodo.org/record/3928360/files/figure.png" pageId="22" pageNumber="23">17</figureCitation>
). Gonocoxa with some paracannular setae, row of setae continuing on anterior side of distal end of gonocoxa. Prefemoral part with rounded ventral lobe. Telopodite endonodal. Nodal processes M and L similar in size; process L slightly curved mesad; process M slightly longer, distal end curved mesad. Postnodal telopodite relatively long, curved ventrad almost perpendicular to setose prefemoral region and directed first slightly mesad, then dorsad and expanding into subglobose, hollow enlargement, then curving mesad and forming a half-circle for the rest of its length. Subterminal process narrow and curved ventrad, solenomere curved dorsad, then ventrad.
</paragraph>
</subSubSection>
<caption id="DCD8138CFF9C4A1AFF6BF868BF41F80F" ID-DOI="http://doi.org/10.5281/zenodo.3928358" ID-Zenodo-Dep="3928358" httpUri="https://zenodo.org/record/3928358/files/figure.png" pageId="22" pageNumber="23" startId="22.[189,232,1964,1990]" targetBox="[476,1116,1456,1914]" targetPageId="22">
<paragraph id="88184304FF9C4A1AFF6BF868BF41F80F" blockId="22.[189,1399,1964,2026]" pageId="22" pageNumber="23">
<emphasis id="BAD39F16FF9C4A1AFF6BF868BF55F823" bold="true" box="[189,284,1964,1990]" pageId="22" pageNumber="23">Fig. 16.</emphasis>
<taxonomicName id="4FA73887FF9C4A1AFEF0F868BD46F823" authority="Olsen &amp; Enghoff" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[294,783,1964,1990]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="frampt" status="sp. nov.">
<emphasis id="BAD39F16FF9C4A1AFEF0F868BC7CF823" box="[294,565,1964,1990]" italics="true" pageId="22" pageNumber="23">Emphysemastix frampt</emphasis>
Olsen &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF9C4A1AFCCFF868BD3FF823" box="[793,886,1964,1990]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NHMD 621677). Scale bar: 5 mm.
</paragraph>
</caption>
<caption id="DCD8138CFF9D4A1BFF6BF84EBD99F80E" ID-DOI="http://doi.org/10.5281/zenodo.3928360" ID-Zenodo-Dep="3928360" httpUri="https://zenodo.org/record/3928360/files/figure.png" pageId="23" pageNumber="24" startId="23.[189,232,1930,1956]" targetBox="[288,1288,265,1886]" targetPageId="23">
<paragraph id="88184304FF9D4A1BFF6BF84EBD99F80E" blockId="23.[189,1399,1930,2027]" pageId="23" pageNumber="24">
<emphasis id="BAD39F16FF9D4A1BFF6BF84EBF50F841" bold="true" box="[189,281,1930,1956]" pageId="23" pageNumber="24">Fig. 17.</emphasis>
<taxonomicName id="4FA73887FF9D4A1BFEF6F84EBCB7F841" authority="Olsen &amp; Enghoff" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[288,766,1930,1957]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="frampt" status="sp. nov.">
<emphasis id="BAD39F16FF9D4A1BFEF6F84EBC65F841" box="[288,556,1930,1956]" italics="true" pageId="23" pageNumber="24">Emphysemastix frampt</emphasis>
Olsen &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF9D4A1BFCD3F84FBD29F840" box="[773,864,1931,1957]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NHMD 621677), left gonopod.
<emphasis id="BAD39F16FF9D4A1BFF6BF86ABEBDF82D" bold="true" box="[189,244,1966,1992]" pageId="23" pageNumber="24">A–B</emphasis>
. Mesal view.
<emphasis id="BAD39F16FF9D4A1BFE40F86ABF86F82D" bold="true" box="[406,463,1966,1992]" pageId="23" pageNumber="24">C–D</emphasis>
. Lateral view.
<emphasis id="BAD39F16FF9D4A1BFDA8F86ABCFBF82D" bold="true" box="[638,690,1966,1992]" pageId="23" pageNumber="24">E–F</emphasis>
. Dorsal view. Abbreviations:
<emphasis id="BAD39F16FF9D4A1BFBD8F86ABA57F822" box="[1038,1054,1966,1991]" italics="true" pageId="23" pageNumber="24">L</emphasis>
= Process L;
<emphasis id="BAD39F16FF9D4A1BFB6BF86ABA9CF822" box="[1213,1237,1966,1991]" italics="true" pageId="23" pageNumber="24">M</emphasis>
= Process M;
<emphasis id="BAD39F16FF9D4A1BFF6BF815BE98F80E" box="[189,209,2001,2027]" italics="true" pageId="23" pageNumber="24">rl</emphasis>
= rounded lobe;
<emphasis id="BAD39F16FF9D4A1BFE4FF816BFFAF80E" box="[409,435,2002,2027]" italics="true" pageId="23" pageNumber="24">sg</emphasis>
= subglobose enlargement. Scale bars: 1 mm.
</paragraph>
</caption>
<subSubSection id="C0BD108FFF924A14FF6BFE7ABA83FDEF" box="[189,1226,446,522]" pageId="24" pageNumber="25" type="distribution">
<paragraph id="88184304FF924A14FF6BFE7ABF1FFE3D" blockId="24.[189,342,446,472]" box="[189,342,446,472]" pageId="24" pageNumber="25">
<heading id="D350F468FF924A14FF6BFE7ABF1FFE3D" bold="true" box="[189,342,446,472]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="BAD39F16FF924A14FF6BFE7ABF1FFE3D" bold="true" box="[189,342,446,472]" pageId="24" pageNumber="25">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF924A14FF6BFE34BA83FDEF" blockId="24.[189,1226,496,522]" box="[189,1226,496,522]" pageId="24" pageNumber="25">
Known only from the
<typeStatus id="571CFDA6FF924A14FE14FE34BFBCFDEF" box="[450,501,496,522]" pageId="24" pageNumber="25">type</typeStatus>
locality in the Nyambanitu Mts, West Kilombero FR (
<figureCitation id="109C5F81FF924A14FBA2FE34BAF5FDEF" box="[1140,1212,496,522]" captionStart="Fig" captionStartId="2.[189,232,1966,1992]" captionTargetBox="[199,1398,1100,1897]" captionTargetId="figure@2.[194,1482,1044,1958]" captionTargetPageId="2" captionText="Fig. 2. Map of the Udzungwa Mts showing collection sites for gomphodesmids. Based on Marshall et al. (2010: fig. 1)." figureDoi="http://doi.org/10.5281/zenodo.3928330" httpUri="https://zenodo.org/record/3928330/files/figure.png" pageId="24" pageNumber="25">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C0BD108FFF924A14FF6BFDF1BA11FD28" pageId="24" pageNumber="25" type="discussion">
<paragraph id="88184304FF924A14FF6BFDF1BF66FDAA" blockId="24.[189,303,565,591]" box="[189,303,565,591]" pageId="24" pageNumber="25">
<heading id="D350F468FF924A14FF6BFDF1BF66FDAA" bold="true" box="[189,303,565,591]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="BAD39F16FF924A14FF6BFDF1BF66FDAA" bold="true" box="[189,303,565,591]" pageId="24" pageNumber="25">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="88184304FF924A14FF6BFDA2BA11FD28" blockId="24.[189,1398,614,717]" pageId="24" pageNumber="25">
The gonopods of
<taxonomicName id="4FA73887FF924A14FE5FFDA2BFBFFD65" authority="Olsen &amp; Rosenmejer &amp; Enghoff, 2020" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[393,502,614,640]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="frampt" status="sp. nov.">
<emphasis id="BAD39F16FF924A14FE5FFDA2BFBFFD65" box="[393,502,614,640]" italics="true" pageId="24" pageNumber="25">E. frampt</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF924A14FE2BFDA2BC1EFD65" box="[509,599,614,640]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
are particularly similar to those of
<taxonomicName id="4FA73887FF924A14FC38FDA2BAE0FD65" baseAuthorityName="Carl" baseAuthorityYear="1909" box="[1006,1193,614,640]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="flavosignatus">
<emphasis id="BAD39F16FF924A14FC38FDA2BAE0FD65" box="[1006,1193,614,640]" italics="true" pageId="24" pageNumber="25">E. flavosignatus</emphasis>
</taxonomicName>
(compare
<figureCitation id="109C5F81FF924A14FAF2FDA2BB3FFD65" box="[1316,1398,614,640]" captionStart="Fig" captionStartId="23.[189,232,1930,1956]" captionTargetBox="[288,1288,265,1886]" captionTargetId="figure@23.[288,1299,265,1886]" captionTargetPageId="23" captionText="Fig. 17. Emphysemastix frampt Olsen &amp; Enghoff sp. nov., holotype, ♂ (NHMD 621677), left gonopod. A–B. Mesal view. C–D. Lateral view. E–F. Dorsal view. Abbreviations: L = Process L; M = Process M; rl = rounded lobe; sg = subglobose enlargement. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3928360" httpUri="https://zenodo.org/record/3928360/files/figure.png" pageId="24" pageNumber="25">Fig. 17</figureCitation>
with
<bibRefCitation id="EC363EF5FF924A14FF2FFD48BFE3FD42" author="Hoffman R. L." box="[249,426,652,679]" pageId="24" pageNumber="25" refId="ref11967" refString="Hoffman R. L. 2005. Monograph of the Gomphodesmidae: A Family of African Polydesmoid Millipeds. Museum of Natural History, Vienna." type="book" year="2005">Hoffman 2005</bibRefCitation>
: fig. 206). Processes M and L in
<taxonomicName id="4FA73887FF924A14FCE4FD49BDE8FD43" authority="Olsen &amp; Rosenmejer &amp; Enghoff, 2020" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[818,929,652,678]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="frampt" status="sp. nov.">
<emphasis id="BAD39F16FF924A14FCE4FD49BDE8FD43" box="[818,929,652,678]" italics="true" pageId="24" pageNumber="25">E. frampt</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF924A14FC7CFD48BA4FFD43" box="[938,1030,652,678]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
are, however, longer and more curved, and the nodus is larger, with more divergent sides in
<taxonomicName id="4FA73887FF924A14FC51FD77BDBCFD29" authority="Olsen &amp; Rosenmejer &amp; Enghoff, 2020" authorityName="Olsen &amp; Rosenmejer &amp; Enghoff" authorityYear="2020" box="[903,1013,690,716]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="frampt" status="sp. nov.">
<emphasis id="BAD39F16FF924A14FC51FD77BDBCFD29" box="[903,1013,690,716]" italics="true" pageId="24" pageNumber="25">E. frampt</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1E0226DFF924A14FC2BFD77BA11FD28" box="[1021,1112,691,717]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>

143
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<mods:title id="1769BB81A02A80CFFE1296B13A6411DE">A mountain of millipedes IX: Species of the family Gomphodesmidae from the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida)</mods:title>
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<emphasis id="BAD39F16FF9F4A19FDCDFD8DBDC8FD81" bold="true" box="[539,897,585,612]" italics="true" pageId="21" pageNumber="22">Emphysemastix flavosignatus</emphasis>
(
<bibRefCitation id="EC363EF5FF9F4A19FC47FD8EBA59FD81" author="Carl J." box="[913,1040,586,612]" pageId="21" pageNumber="22" pagination="281 - 365" refId="ref11215" refString="Carl J. 1909. Reise von Dr. J. Carl im nordlichen Zentral-Afrikanischen Seengebiet. Diplopoden. Revue suisse de Zoologie 17: 281 - 365. Available from https: // www. biodiversitylibrary. org / item / 40663 # page / 293 / mode / 1 up [accessed 27 May 2020]." type="journal article" year="1909">Carl, 1909</bibRefCitation>
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<taxonomicName id="4FA73887FF9F4A19FF6BFD48BCD7FD40" ID-CoL="3D8DM" authority="Carl, 1909: 307" authorityName="Carl" authorityPageNumber="307" authorityYear="1909" box="[189,670,651,677]" class="Diplopoda" family="Gomphodesmidae" genus="Euryzonus" kingdom="Animalia" order="Polydesmida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="flavosignatus">
<emphasis id="BAD39F16FF9F4A19FF6BFD48BF92FD40" box="[189,475,651,677]" italics="true" pageId="21" pageNumber="22">Euryzonus flavosignatus</emphasis>
<bibRefCitation id="EC363EF5FF9F4A19FE34FD4FBCD7FD40" author="Carl J." box="[482,670,651,677]" pageId="21" pageNumber="22" pagination="281 - 365" refId="ref11215" refString="Carl J. 1909. Reise von Dr. J. Carl im nordlichen Zentral-Afrikanischen Seengebiet. Diplopoden. Revue suisse de Zoologie 17: 281 - 365. Available from https: // www. biodiversitylibrary. org / item / 40663 # page / 293 / mode / 1 up [accessed 27 May 2020]." type="journal article" year="1909">Carl, 1909: 307</bibRefCitation>
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<taxonomicName id="4FA73887FF9F4A19FF6BFD08BFA4FD03" ID-CoL="JS92" baseAuthorityName="Carl" baseAuthorityYear="1909" box="[189,493,716,742]" class="Diplopoda" family="Gomphodesmidae" genus="Aulodesmus" kingdom="Animalia" order="Polydesmida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="flavosignatus">
<emphasis id="BAD39F16FF9F4A19FF6BFD08BFA4FD03" box="[189,493,716,742]" italics="true" pageId="21" pageNumber="22">Aulodesmus flavosignatus</emphasis>
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–
<treatmentCitation id="09066515FF9F4A19FDDFFD09BC97FD02" author="Attems C." box="[521,734,717,743]" page="102" pageId="21" pageNumber="22" year="1931">
<bibRefCitation id="EC363EF5FF9F4A19FDDFFD09BC97FD02" author="Attems C." box="[521,734,717,743]" pageId="21" pageNumber="22" pagination="1 - 150" refId="ref11191" refString="Attems C. 1931. Die Familie Leptodesmidae und andere Polydesmiden. Zoologica, Stuttgart 30 (70): 1 - 150." type="journal article" year="1931">Attems 1931: 102</bibRefCitation>
</treatmentCitation>
.
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<paragraph id="88184304FF9F4A19FF6BFD34BD68FCEF" blockId="21.[189,801,716,778]" box="[189,801,752,778]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="A8B7642AFF9F4A19FF6BFD34BD68FCEF" box="[189,801,752,778]" pageId="21" pageNumber="22">
<taxonomicName id="4FA73887FF9F4A19FF6BFD34BC5FFCEF" ID-CoL="39HQW" baseAuthorityName="Carl" baseAuthorityYear="1909" box="[189,534,752,778]" class="Diplopoda" family="Gomphodesmidae" genus="Emphysemastix" kingdom="Animalia" order="Polydesmida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="flavosignatus">
<emphasis id="BAD39F16FF9F4A19FF6BFD34BC5FFCEF" box="[189,534,752,778]" italics="true" pageId="21" pageNumber="22">Emphysemastix flavosignatus</emphasis>
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–
<treatmentCitation id="09066515FF9F4A19FDE5FD34BD57FCEF" author="Hoffman R. L." box="[563,798,752,778]" page="208" pageId="21" pageNumber="22" year="1966">
<bibRefCitation id="EC363EF5FF9F4A19FDE5FD34BD57FCEF" author="Hoffman R. L." box="[563,798,752,778]" pageId="21" pageNumber="22" pagination="193 - 208" refId="ref11929" refString="Hoffman R. L. 1966. Notes on Aulodesmus and the proposal of a related new genus (Polydesmida: Gomphodesmidae). Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut 63: 193 - 208." type="journal article" year="1966">Hoffman 1966: 208</bibRefCitation>
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<emphasis id="BAD39F16FF9F4A19FF6BFCF5BFE3FCAE" bold="true" box="[189,426,817,843]" pageId="21" pageNumber="22">Material examined</emphasis>
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<emphasis id="BAD39F16FF9F4A19FF3CFCA4BF15FC9F" bold="true" box="[234,348,864,890]" pageId="21" pageNumber="22">
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<paragraph id="88184304FF9F4A19FF6BFC47BA77FC7B" blockId="21.[189,1086,864,926]" box="[189,1086,899,926]" pageId="21" pageNumber="22">
<materialsCitation id="38CF4959FF9F4A19FF6BFC47BA73FC7B" ID-GBIF-Occurrence="2803530315" box="[189,1082,899,926]" collectionCode="NHMD" country="Tanzania" location="TANZANIA" pageId="21" pageNumber="22" specimenCode="NHMD 3853" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectingCountry id="F0B00394FF9F4A19FF6BFC47BF18FC78" box="[189,337,899,925]" name="Tanzania" pageId="21" pageNumber="22">TANZANIA</collectingCountry>
•
<specimenCount id="9EA1888DFF9F4A19FEBFFC47BFDEFC78" box="[361,407,899,925]" pageId="21" pageNumber="22" type="male">1 ♂</specimenCount>
; “Karagwe, D.-O.-Afr.; Carl ded.; cotype”;
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.
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<heading id="D350F468FF9F4A19FF6BFC01BF6DFC3A" bold="true" box="[189,292,965,991]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="BAD39F16FF9F4A19FF6BFC01BF6DFC3A" bold="true" box="[189,292,965,991]" pageId="21" pageNumber="22">Remark</emphasis>
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<collectingRegion id="4A638DE6FF9F4A19FF6BFC37BEA0FBE8" box="[189,233,1011,1037]" country="Cameroon" name="North" pageId="21" pageNumber="22">Not</collectingRegion>
reported from the Udzungwa Mts, but studied for comparison with the new species.
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74
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Family
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Tarsiidae
<bibRefCitation id="99ACDCF13723720652ECC39E55128366" author="Gray, J. E." refId="ref127941" refString="Gray, J. E. 1825. Outline of an attempt at the disposition of the Mammalia into tribes and families with a list of the genera apparently appertaining to each tribe. Annals of Philosophy, n. s., ser. 2, 10: 337 - 344." year="1825">Gray 1825</bibRefCitation>
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(1 genus with 7 species and 3 subspecies)
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<emphasis id="F902A0FB7B98F90EFB7970C763A54EB0" inLineHeading="true" pageNumber="127">Discussion:</emphasis>
Almost invariably placed in a group with
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