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<document id="4FB45C2408938061028A730542C79F25" ID-DOI="10.1016/j.rbe.2018.01.002" ID-ISSN="1806-9665" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722640665602" checkinUser="felipe" docAuthor="Kohlmann, Bert &amp; Vaz-de-Mello, Fernando Z." docDate="2018" docId="327E87E1D879FFEDFCC0FAEFFF44B0C5" docLanguage="en" docName="RevBrasEntomol.62.2.131-134.pdf" docOrigin="Revista Brasileira de Entomologia 62 (2)" docSource="http://dx.doi.org/10.1016/j.rbe.2018.01.002" docStyle="DocumentStyle:0303D37C566E4E2A42C6591146119864.3:RevBrasEntomol.2015-.journal_article" docStyleId="0303D37C566E4E2A42C6591146119864" docStyleName="RevBrasEntomol.2015-.journal_article" docStyleVersion="3" docTitle="Ateuchus tuza Kohlmann and Vaz de Mello 2018, sp. nov." docType="treatment" docVersion="1" lastPageNumber="133" masterDocId="CE47FF99D878FFEEFFE4FFE8FFE1B002" masterDocTitle="A new key for the species of Ateuchus Weber (Coleoptera: Scarabaeidae: Scarabaeinae) occurring in Mexico, with a description of the first North American inquiline species from a rodent burrow (Rodentia: Geomydae) and new distribution records" masterLastPageNumber="134" masterPageNumber="131" pageNumber="132" updateTime="1722684135551" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0"> <document id="37BFF805D55ED96C1F04E82B32AD9CE3" ID-DOI="10.1016/j.rbe.2018.01.002" ID-ISSN="1806-9665" ID-Zenodo-Dep="13198486" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722640665602" checkinUser="felipe" docAuthor="Kohlmann, Bert &amp; Vaz-de-Mello, Fernando Z." docDate="2018" docId="327E87E1D879FFEDFCC0FAEFFF44B0C5" docLanguage="en" docName="RevBrasEntomol.62.2.131-134.pdf" docOrigin="Revista Brasileira de Entomologia 62 (2)" docSource="http://dx.doi.org/10.1016/j.rbe.2018.01.002" docStyle="DocumentStyle:0303D37C566E4E2A42C6591146119864.3:RevBrasEntomol.2015-.journal_article" docStyleId="0303D37C566E4E2A42C6591146119864" docStyleName="RevBrasEntomol.2015-.journal_article" docStyleVersion="3" docTitle="Ateuchus tuza Kohlmann and Vaz de Mello 2018, sp. nov." docType="treatment" docVersion="2" lastPageNumber="133" masterDocId="CE47FF99D878FFEEFFE4FFE8FFE1B002" masterDocTitle="A new key for the species of Ateuchus Weber (Coleoptera: Scarabaeidae: Scarabaeinae) occurring in Mexico, with a description of the first North American inquiline species from a rodent burrow (Rodentia: Geomydae) and new distribution records" masterLastPageNumber="134" masterPageNumber="131" pageNumber="132" updateTime="1722688723938" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="10EB576EB191D7C82B993B6AA956666A">A new key for the species of Ateuchus Weber (Coleoptera: Scarabaeidae: Scarabaeinae) occurring in Mexico, with a description of the first North American inquiline species from a rodent burrow (Rodentia: Geomydae) and new distribution records</mods:title> <mods:title id="00A61656C370A95F3A5454DF894B3D2B">A new key for the species of Ateuchus Weber (Coleoptera: Scarabaeidae: Scarabaeinae) occurring in Mexico, with a description of the first North American inquiline species from a rodent burrow (Rodentia: Geomydae) and new distribution records</mods:title>
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<mods:namePart id="FE02B2C0DDD4AD1058D7FCD1FBA460E9">Kohlmann, Bert</mods:namePart> <mods:namePart id="AAC34C2A2E5146FA349F963FC7E86709">Kohlmann, Bert</mods:namePart>
<mods:affiliation id="700BA985F08BC401021C1DAB7531D973">EARTH University, San José, Costa Rica</mods:affiliation> <mods:affiliation id="33924B3699551F2F491ADFB530AA98FB">EARTH University, San José, Costa Rica</mods:affiliation>
</mods:name> </mods:name>
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<mods:namePart id="866F51669BD72A2E754D8D09FB7D4108">Vaz-de-Mello, Fernando Z.</mods:namePart> <mods:namePart id="4E8528C5FF16969C89D9731F3820375F">Vaz-de-Mello, Fernando Z.</mods:namePart>
<mods:affiliation id="A48F7275E4E06FD190F20F81DC945AF8">Universidade Federal de Mato Grosso, Instituto de Biociências, Departamento de Biologia e Zoologia, Cuiabá, MT, Brazil</mods:affiliation> <mods:affiliation id="CC582B8804C3A838403104287BE6072E">Universidade Federal de Mato Grosso, Instituto de Biociências, Departamento de Biologia e Zoologia, Cuiabá, MT, Brazil</mods:affiliation>
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<treatment id="327E87E1D879FFEDFCC0FAEFFF44B0C5" LSID="urn:lsid:plazi:treatment:327E87E1D879FFEDFCC0FAEFFF44B0C5" httpUri="http://treatment.plazi.org/id/327E87E1D879FFEDFCC0FAEFFF44B0C5" lastPageId="3" lastPageNumber="133" pageId="1" pageNumber="132"> <treatment id="327E87E1D879FFEDFCC0FAEFFF44B0C5" LSID="urn:lsid:plazi:treatment:327E87E1D879FFEDFCC0FAEFFF44B0C5" httpUri="http://treatment.plazi.org/id/327E87E1D879FFEDFCC0FAEFFF44B0C5" lastPageId="3" lastPageNumber="133" pageId="1" pageNumber="132">
<subSubSection id="F2CD657CD879FFEFFCC0FAEFFAA5B518" box="[804,1348,1287,1306]" pageId="1" pageNumber="132" type="nomenclature"> <subSubSection id="F2CD657CD879FFEFFCC0FAEFFAA5B518" box="[804,1348,1287,1306]" pageId="1" pageNumber="132" type="nomenclature">
@ -64,7 +65,7 @@ Kohlmann and Vaz de Mello
<subSubSection id="F2CD657CD879FFEFFCC0FACBFC6CB534" box="[804,909,1315,1334]" pageId="1" pageNumber="132" type="description"> <subSubSection id="F2CD657CD879FFEFFCC0FACBFC6CB534" box="[804,909,1315,1334]" pageId="1" pageNumber="132" type="description">
<paragraph id="BA6836F7D879FFEFFCC0FACBFC6CB534" blockId="1.[804,1475,1315,1809]" box="[804,909,1315,1334]" pageId="1" pageNumber="132"> <paragraph id="BA6836F7D879FFEFFCC0FACBFC6CB534" blockId="1.[804,1475,1315,1809]" box="[804,909,1315,1334]" pageId="1" pageNumber="132">
( (
<figureCitation id="22EC2A72D879FFEFFCC8FACBFC64B534" box="[812,901,1315,1334]" captionStart-0="Figures 1–3" captionStart-1="Figures 4–5" captionStartId-0="1.[804,867,1208,1222]" captionStartId-1="2.[833,896,1056,1070]" captionTargetBox-0="[839,1442,150,1177]" captionTargetBox-1="[867,1471,150,1024]" captionTargetId-0="figure-1443@1.[835,1443,150,1177]" captionTargetId-1="figure-619@2.[864,1472,149,1026]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Figures 1–3. Ateuchus tuza sp. nov.; 1, dorsal habitus of holotype; 2, ventral habitus of holotype; 3, wrinkled area at the base of the head of a male." captionText-1="Figures 4–5. A. tuza sp. nov.; 4, drawing of the internal sac, line equals 1 mm; 5, A. hornai (Balthasar, 1939), dorsal habitus of the female holotype." pageId="1" pageNumber="132">Figs. 1–4</figureCitation> <figureCitation id="22EC2A72D879FFEFFCC8FACBFC64B534" box="[812,901,1315,1334]" captionStart-0="Figures 1–3" captionStart-1="Figures 4–5" captionStartId-0="1.[804,867,1208,1222]" captionStartId-1="2.[833,896,1056,1070]" captionTargetBox-0="[839,1442,150,1177]" captionTargetBox-1="[867,1471,150,1024]" captionTargetId-0="figure-1443@1.[835,1443,150,1177]" captionTargetId-1="figure-619@2.[864,1472,149,1026]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Figures 1–3. Ateuchus tuza sp. nov.; 1, dorsal habitus of holotype; 2, ventral habitus of holotype; 3, wrinkled area at the base of the head of a male." captionText-1="Figures 4–5. A. tuza sp. nov.; 4, drawing of the internal sac, line equals 1 mm; 5, A. hornai (Balthasar, 1939), dorsal habitus of the female holotype." figureDoi-0="http://doi.org/10.5281/zenodo.13198488" figureDoi-1="http://doi.org/10.5281/zenodo.13198490" httpUri-0="https://zenodo.org/record/13198488/files/figure.png" httpUri-1="https://zenodo.org/record/13198490/files/figure.png" pageId="1" pageNumber="132">Figs. 1–4</figureCitation>
) )
</paragraph> </paragraph>
</subSubSection> </subSubSection>
@ -128,13 +129,13 @@ by the following combination of characters: Long and slender body; dorsum glossy
<emphasis id="88A3EAE5D879FFEFFCC0F99AFC44B687" bold="true" box="[804,933,1650,1669]" pageId="1" pageNumber="132">Description.</emphasis> <emphasis id="88A3EAE5D879FFEFFCC0F99AFC44B687" bold="true" box="[804,933,1650,1669]" pageId="1" pageNumber="132">Description.</emphasis>
<typeStatus id="656C8855D879FFEFFC4EF99AFBE6B687" box="[938,1031,1650,1669]" pageId="1" pageNumber="132" type="holotype">Holotype</typeStatus> <typeStatus id="656C8855D879FFEFFC4EF99AFBE6B687" box="[938,1031,1650,1669]" pageId="1" pageNumber="132" type="holotype">Holotype</typeStatus>
male ( male (
<figureCitation id="22EC2A72D879FFEFFBA8F99AFB45B687" box="[1100,1188,1650,1669]" captionStart-0="Figures 1–3" captionStart-1="Figures 4–5" captionStartId-0="1.[804,867,1208,1222]" captionStartId-1="2.[833,896,1056,1070]" captionTargetBox-0="[839,1442,150,1177]" captionTargetBox-1="[867,1471,150,1024]" captionTargetId-0="figure-1443@1.[835,1443,150,1177]" captionTargetId-1="figure-619@2.[864,1472,149,1026]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Figures 1–3. Ateuchus tuza sp. nov.; 1, dorsal habitus of holotype; 2, ventral habitus of holotype; 3, wrinkled area at the base of the head of a male." captionText-1="Figures 4–5. A. tuza sp. nov.; 4, drawing of the internal sac, line equals 1 mm; 5, A. hornai (Balthasar, 1939), dorsal habitus of the female holotype." pageId="1" pageNumber="132">Figs. 1–4</figureCitation> <figureCitation id="22EC2A72D879FFEFFBA8F99AFB45B687" box="[1100,1188,1650,1669]" captionStart-0="Figures 1–3" captionStart-1="Figures 4–5" captionStartId-0="1.[804,867,1208,1222]" captionStartId-1="2.[833,896,1056,1070]" captionTargetBox-0="[839,1442,150,1177]" captionTargetBox-1="[867,1471,150,1024]" captionTargetId-0="figure-1443@1.[835,1443,150,1177]" captionTargetId-1="figure-619@2.[864,1472,149,1026]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Figures 1–3. Ateuchus tuza sp. nov.; 1, dorsal habitus of holotype; 2, ventral habitus of holotype; 3, wrinkled area at the base of the head of a male." captionText-1="Figures 4–5. A. tuza sp. nov.; 4, drawing of the internal sac, line equals 1 mm; 5, A. hornai (Balthasar, 1939), dorsal habitus of the female holotype." figureDoi-0="http://doi.org/10.5281/zenodo.13198488" figureDoi-1="http://doi.org/10.5281/zenodo.13198490" httpUri-0="https://zenodo.org/record/13198488/files/figure.png" httpUri-1="https://zenodo.org/record/13198490/files/figure.png" pageId="1" pageNumber="132">Figs. 1–4</figureCitation>
). ).
<emphasis id="88A3EAE5D879FFEFFB53F99AFB13B687" bold="true" box="[1207,1266,1650,1669]" pageId="1" pageNumber="132">Body.</emphasis> <emphasis id="88A3EAE5D879FFEFFB53F99AFB13B687" bold="true" box="[1207,1266,1650,1669]" pageId="1" pageNumber="132">Body.</emphasis>
Elongate ( Elongate (
<figureCitation id="22EC2A72D879FFEFFAB9F99AFA54B687" box="[1373,1461,1650,1669]" captionStart="Figures 1–3" captionStartId="1.[804,867,1208,1222]" captionTargetBox="[839,1442,150,1177]" captionTargetId="figure-1443@1.[835,1443,150,1177]" captionTargetPageId="1" captionText="Figures 1–3. Ateuchus tuza sp. nov.; 1, dorsal habitus of holotype; 2, ventral habitus of holotype; 3, wrinkled area at the base of the head of a male." pageId="1" pageNumber="132">Figs. 1–2</figureCitation> <figureCitation id="22EC2A72D879FFEFFAB9F99AFA54B687" box="[1373,1461,1650,1669]" captionStart="Figures 1–3" captionStartId="1.[804,867,1208,1222]" captionTargetBox="[839,1442,150,1177]" captionTargetId="figure-1443@1.[835,1443,150,1177]" captionTargetPageId="1" captionText="Figures 1–3. Ateuchus tuza sp. nov.; 1, dorsal habitus of holotype; 2, ventral habitus of holotype; 3, wrinkled area at the base of the head of a male." figureDoi="http://doi.org/10.5281/zenodo.13198488" httpUri="https://zenodo.org/record/13198488/files/figure.png" pageId="1" pageNumber="132">Figs. 1–2</figureCitation>
), convex dorsally ( ), convex dorsally (
<figureCitation id="22EC2A72D879FFEFFC3FF966FBF7B6A3" box="[987,1046,1678,1697]" captionStart="Figures 1–3" captionStartId="1.[804,867,1208,1222]" captionTargetBox="[839,1442,150,1177]" captionTargetId="figure-1443@1.[835,1443,150,1177]" captionTargetPageId="1" captionText="Figures 1–3. Ateuchus tuza sp. nov.; 1, dorsal habitus of holotype; 2, ventral habitus of holotype; 3, wrinkled area at the base of the head of a male." pageId="1" pageNumber="132">Fig. 1</figureCitation> <figureCitation id="22EC2A72D879FFEFFC3FF966FBF7B6A3" box="[987,1046,1678,1697]" captionStart="Figures 1–3" captionStartId="1.[804,867,1208,1222]" captionTargetBox="[839,1442,150,1177]" captionTargetId="figure-1443@1.[835,1443,150,1177]" captionTargetPageId="1" captionText="Figures 1–3. Ateuchus tuza sp. nov.; 1, dorsal habitus of holotype; 2, ventral habitus of holotype; 3, wrinkled area at the base of the head of a male." figureDoi="http://doi.org/10.5281/zenodo.13198488" httpUri="https://zenodo.org/record/13198488/files/figure.png" pageId="1" pageNumber="132">Fig. 1</figureCitation>
). ).
<emphasis id="88A3EAE5D879FFEFFBCAF965FBBCB6A2" bold="true" box="[1070,1117,1677,1696]" pageId="1" pageNumber="132">Size.</emphasis> <emphasis id="88A3EAE5D879FFEFFBCAF965FBBCB6A2" bold="true" box="[1070,1117,1677,1696]" pageId="1" pageNumber="132">Size.</emphasis>
Total length 10.0 mm. Maximum width 5.5 mm. Total length 10.0 mm. Maximum width 5.5 mm.
@ -142,33 +143,33 @@ Total length 10.0 mm. Maximum width 5.5 mm.
Dark reddish brown to black, lacking metallic sheen. Dark reddish brown to black, lacking metallic sheen.
<emphasis id="88A3EAE5D879FFEFFC7DF92EFC34B6DB" bold="true" box="[921,981,1734,1753]" pageId="1" pageNumber="132">Head.</emphasis> <emphasis id="88A3EAE5D879FFEFFC7DF92EFC34B6DB" bold="true" box="[921,981,1734,1753]" pageId="1" pageNumber="132">Head.</emphasis>
Clypeal margin anteriorly broadly V-shaped ( Clypeal margin anteriorly broadly V-shaped (
<figureCitation id="22EC2A72D879FFEFFCC8F909FC6AB6F6" box="[812,907,1761,1780]" captionStart="Figures 1–3" captionStartId="1.[804,867,1208,1222]" captionTargetBox="[839,1442,150,1177]" captionTargetId="figure-1443@1.[835,1443,150,1177]" captionTargetPageId="1" captionText="Figures 1–3. Ateuchus tuza sp. nov.; 1, dorsal habitus of holotype; 2, ventral habitus of holotype; 3, wrinkled area at the base of the head of a male." pageId="1" pageNumber="132">Figs. 1–2</figureCitation> <figureCitation id="22EC2A72D879FFEFFCC8F909FC6AB6F6" box="[812,907,1761,1780]" captionStart="Figures 1–3" captionStartId="1.[804,867,1208,1222]" captionTargetBox="[839,1442,150,1177]" captionTargetId="figure-1443@1.[835,1443,150,1177]" captionTargetPageId="1" captionText="Figures 1–3. Ateuchus tuza sp. nov.; 1, dorsal habitus of holotype; 2, ventral habitus of holotype; 3, wrinkled area at the base of the head of a male." figureDoi="http://doi.org/10.5281/zenodo.13198488" httpUri="https://zenodo.org/record/13198488/files/figure.png" pageId="1" pageNumber="132">Figs. 1–2</figureCitation>
); anterior margin slightly upturned, gena and frons coarsely wrinkled, vertex coarsely punctate, base of head with a small wrinkled area in the center ( ); anterior margin slightly upturned, gena and frons coarsely wrinkled, vertex coarsely punctate, base of head with a small wrinkled area in the center (
<figureCitation id="22EC2A72D87AFFECFE35FF70FDE6B0A9" box="[465,519,152,171]" captionStart="Figures 1–3" captionStartId="1.[804,867,1208,1222]" captionTargetBox="[839,1442,150,1177]" captionTargetId="figure-1443@1.[835,1443,150,1177]" captionTargetPageId="1" captionText="Figures 1–3. Ateuchus tuza sp. nov.; 1, dorsal habitus of holotype; 2, ventral habitus of holotype; 3, wrinkled area at the base of the head of a male." pageId="2" pageNumber="133">Fig. 3</figureCitation> <figureCitation id="22EC2A72D87AFFECFE35FF70FDE6B0A9" box="[465,519,152,171]" captionStart="Figures 1–3" captionStartId="1.[804,867,1208,1222]" captionTargetBox="[839,1442,150,1177]" captionTargetId="figure-1443@1.[835,1443,150,1177]" captionTargetPageId="1" captionText="Figures 1–3. Ateuchus tuza sp. nov.; 1, dorsal habitus of holotype; 2, ventral habitus of holotype; 3, wrinkled area at the base of the head of a male." figureDoi="http://doi.org/10.5281/zenodo.13198488" httpUri="https://zenodo.org/record/13198488/files/figure.png" pageId="2" pageNumber="133">Fig. 3</figureCitation>
), eyes viewed from above six times longer than wide. ), eyes viewed from above six times longer than wide.
<emphasis id="88A3EAE5D87AFFECFE77FF5CFDE7B0C5" bold="true" box="[403,518,180,199]" pageId="2" pageNumber="133">Pronotum.</emphasis> <emphasis id="88A3EAE5D87AFFECFE77FF5CFDE7B0C5" bold="true" box="[403,518,180,199]" pageId="2" pageNumber="133">Pronotum.</emphasis>
Transverse, strongly convex, surface glossy; anterior pronotal margin complete; midline weakly impressed at base; pronotal surface punctate throughout, a group of coarse punctures at the center of the pronotal base ( Transverse, strongly convex, surface glossy; anterior pronotal margin complete; midline weakly impressed at base; pronotal surface punctate throughout, a group of coarse punctures at the center of the pronotal base (
<figureCitation id="22EC2A72D87AFFECFF4AFECBFF02B134" box="[174,227,291,310]" captionStart="Figures 1–3" captionStartId="1.[804,867,1208,1222]" captionTargetBox="[839,1442,150,1177]" captionTargetId="figure-1443@1.[835,1443,150,1177]" captionTargetPageId="1" captionText="Figures 1–3. Ateuchus tuza sp. nov.; 1, dorsal habitus of holotype; 2, ventral habitus of holotype; 3, wrinkled area at the base of the head of a male." pageId="2" pageNumber="133">Fig. 1</figureCitation> <figureCitation id="22EC2A72D87AFFECFF4AFECBFF02B134" box="[174,227,291,310]" captionStart="Figures 1–3" captionStartId="1.[804,867,1208,1222]" captionTargetBox="[839,1442,150,1177]" captionTargetId="figure-1443@1.[835,1443,150,1177]" captionTargetPageId="1" captionText="Figures 1–3. Ateuchus tuza sp. nov.; 1, dorsal habitus of holotype; 2, ventral habitus of holotype; 3, wrinkled area at the base of the head of a male." figureDoi="http://doi.org/10.5281/zenodo.13198488" httpUri="https://zenodo.org/record/13198488/files/figure.png" pageId="2" pageNumber="133">Fig. 1</figureCitation>
); lateral pronotal fossae shallow. ); lateral pronotal fossae shallow.
<emphasis id="88A3EAE5D87AFFECFDDFFECBFD61B134" bold="true" box="[571,640,291,310]" pageId="2" pageNumber="133">Elytra.</emphasis> <emphasis id="88A3EAE5D87AFFECFDDFFECBFD61B134" bold="true" box="[571,640,291,310]" pageId="2" pageNumber="133">Elytra.</emphasis>
( (
<figureCitation id="22EC2A72D87AFFECFD6AFECBFD22B134" box="[654,707,291,310]" captionStart="Figures 1–3" captionStartId="1.[804,867,1208,1222]" captionTargetBox="[839,1442,150,1177]" captionTargetId="figure-1443@1.[835,1443,150,1177]" captionTargetPageId="1" captionText="Figures 1–3. Ateuchus tuza sp. nov.; 1, dorsal habitus of holotype; 2, ventral habitus of holotype; 3, wrinkled area at the base of the head of a male." pageId="2" pageNumber="133">Fig. 1</figureCitation> <figureCitation id="22EC2A72D87AFFECFD6AFECBFD22B134" box="[654,707,291,310]" captionStart="Figures 1–3" captionStartId="1.[804,867,1208,1222]" captionTargetBox="[839,1442,150,1177]" captionTargetId="figure-1443@1.[835,1443,150,1177]" captionTargetPageId="1" captionText="Figures 1–3. Ateuchus tuza sp. nov.; 1, dorsal habitus of holotype; 2, ventral habitus of holotype; 3, wrinkled area at the base of the head of a male." figureDoi="http://doi.org/10.5281/zenodo.13198488" httpUri="https://zenodo.org/record/13198488/files/figure.png" pageId="2" pageNumber="133">Fig. 1</figureCitation>
). Striae fine, shallowly impressed on disk, becoming well defined and deeply impressed on apical declivity; strial punctures elongatecrenulate, slightly wider than stria, separated by 2–3 diameters on disk and apical declivity. Interstriae slightly convex on disk, surface minutely punctate throughout. ). Striae fine, shallowly impressed on disk, becoming well defined and deeply impressed on apical declivity; strial punctures elongatecrenulate, slightly wider than stria, separated by 2–3 diameters on disk and apical declivity. Interstriae slightly convex on disk, surface minutely punctate throughout.
<emphasis id="88A3EAE5D87AFFECFDE9FE47FD5BB1C0" bold="true" box="[525,698,431,450]" pageId="2" pageNumber="133">Thoracic sterna.</emphasis> <emphasis id="88A3EAE5D87AFFECFDE9FE47FD5BB1C0" bold="true" box="[525,698,431,450]" pageId="2" pageNumber="133">Thoracic sterna.</emphasis>
( (
<figureCitation id="22EC2A72D87AFFECFD2FFE47FCE5B1C0" box="[715,772,431,450]" captionStart="Figures 1–3" captionStartId="1.[804,867,1208,1222]" captionTargetBox="[839,1442,150,1177]" captionTargetId="figure-1443@1.[835,1443,150,1177]" captionTargetPageId="1" captionText="Figures 1–3. Ateuchus tuza sp. nov.; 1, dorsal habitus of holotype; 2, ventral habitus of holotype; 3, wrinkled area at the base of the head of a male." pageId="2" pageNumber="133">Fig. 2</figureCitation> <figureCitation id="22EC2A72D87AFFECFD2FFE47FCE5B1C0" box="[715,772,431,450]" captionStart="Figures 1–3" captionStartId="1.[804,867,1208,1222]" captionTargetBox="[839,1442,150,1177]" captionTargetId="figure-1443@1.[835,1443,150,1177]" captionTargetPageId="1" captionText="Figures 1–3. Ateuchus tuza sp. nov.; 1, dorsal habitus of holotype; 2, ventral habitus of holotype; 3, wrinkled area at the base of the head of a male." figureDoi="http://doi.org/10.5281/zenodo.13198488" httpUri="https://zenodo.org/record/13198488/files/figure.png" pageId="2" pageNumber="133">Fig. 2</figureCitation>
). Proepisternum excavate anteriorly, surface of excavated portion granulate, with fine and rather long setae, bordered posteriorly by a well-defined keel. Proepimeron finely wrinkled. Sternellum glabrous and forming a keel at its center. Mesometasternal suture arched medially, marginal bead fine. Mesosternum coarsely punctured forming rugulae. Mesepisternum flat, surface-forming rugulae. Metasternum evenly convex, disk evidently punctate, surface glossy between punctures, lateral lobes forming rugulae; midline clearly impressed along three fourths of its length. ). Proepisternum excavate anteriorly, surface of excavated portion granulate, with fine and rather long setae, bordered posteriorly by a well-defined keel. Proepimeron finely wrinkled. Sternellum glabrous and forming a keel at its center. Mesometasternal suture arched medially, marginal bead fine. Mesosternum coarsely punctured forming rugulae. Mesepisternum flat, surface-forming rugulae. Metasternum evenly convex, disk evidently punctate, surface glossy between punctures, lateral lobes forming rugulae; midline clearly impressed along three fourths of its length.
<emphasis id="88A3EAE5D87AFFECFD39FD42FCF0B2BF" bold="true" box="[733,785,682,701]" pageId="2" pageNumber="133">Legs.</emphasis> <emphasis id="88A3EAE5D87AFFECFD39FD42FCF0B2BF" bold="true" box="[733,785,682,701]" pageId="2" pageNumber="133">Legs.</emphasis>
( (
<figureCitation id="22EC2A72D87AFFECFF9EFD2EFF33B2DB" box="[122,210,710,729]" captionStart="Figures 1–3" captionStartId="1.[804,867,1208,1222]" captionTargetBox="[839,1442,150,1177]" captionTargetId="figure-1443@1.[835,1443,150,1177]" captionTargetPageId="1" captionText="Figures 1–3. Ateuchus tuza sp. nov.; 1, dorsal habitus of holotype; 2, ventral habitus of holotype; 3, wrinkled area at the base of the head of a male." pageId="2" pageNumber="133">Figs. 1–2</figureCitation> <figureCitation id="22EC2A72D87AFFECFF9EFD2EFF33B2DB" box="[122,210,710,729]" captionStart="Figures 1–3" captionStartId="1.[804,867,1208,1222]" captionTargetBox="[839,1442,150,1177]" captionTargetId="figure-1443@1.[835,1443,150,1177]" captionTargetPageId="1" captionText="Figures 1–3. Ateuchus tuza sp. nov.; 1, dorsal habitus of holotype; 2, ventral habitus of holotype; 3, wrinkled area at the base of the head of a male." figureDoi="http://doi.org/10.5281/zenodo.13198488" httpUri="https://zenodo.org/record/13198488/files/figure.png" pageId="2" pageNumber="133">Figs. 1–2</figureCitation>
). Foretibia with four teeth on outer margin, the basal one weakly developed; foretibial spur expanded and truncate apically; foretibiae and forefemora long and slender; forefemur smooth, finely punctate throughout. Metatibiae obliquely truncate apically; apical spur spiniform. ). Foretibia with four teeth on outer margin, the basal one weakly developed; foretibial spur expanded and truncate apically; foretibiae and forefemora long and slender; forefemur smooth, finely punctate throughout. Metatibiae obliquely truncate apically; apical spur spiniform.
<emphasis id="88A3EAE5D87AFFECFEB6FCDDFE5EB34A" bold="true" box="[338,447,821,840]" pageId="2" pageNumber="133">Abdomen.</emphasis> <emphasis id="88A3EAE5D87AFFECFEB6FCDDFE5EB34A" bold="true" box="[338,447,821,840]" pageId="2" pageNumber="133">Abdomen.</emphasis>
( (
<figureCitation id="22EC2A72D87AFFECFE29FCDDFDE3B34B" box="[461,514,821,841]" captionStart="Figures 1–3" captionStartId="1.[804,867,1208,1222]" captionTargetBox="[839,1442,150,1177]" captionTargetId="figure-1443@1.[835,1443,150,1177]" captionTargetPageId="1" captionText="Figures 1–3. Ateuchus tuza sp. nov.; 1, dorsal habitus of holotype; 2, ventral habitus of holotype; 3, wrinkled area at the base of the head of a male." pageId="2" pageNumber="133">Fig. 2</figureCitation> <figureCitation id="22EC2A72D87AFFECFE29FCDDFDE3B34B" box="[461,514,821,841]" captionStart="Figures 1–3" captionStartId="1.[804,867,1208,1222]" captionTargetBox="[839,1442,150,1177]" captionTargetId="figure-1443@1.[835,1443,150,1177]" captionTargetPageId="1" captionText="Figures 1–3. Ateuchus tuza sp. nov.; 1, dorsal habitus of holotype; 2, ventral habitus of holotype; 3, wrinkled area at the base of the head of a male." figureDoi="http://doi.org/10.5281/zenodo.13198488" httpUri="https://zenodo.org/record/13198488/files/figure.png" pageId="2" pageNumber="133">Fig. 2</figureCitation>
). Sternites 3–7 crenulately punctate along their anterior borders. Last abdominal segment slender. Pygidium slightly convex and lightly shagreen with indistinct punctuation; basal sulcus fine and deep throughout; marginal bead continuous. ). Sternites 3–7 crenulately punctate along their anterior borders. Last abdominal segment slender. Pygidium slightly convex and lightly shagreen with indistinct punctuation; basal sulcus fine and deep throughout; marginal bead continuous.
<emphasis id="88A3EAE5D87AFFECFEC8FC4DFE2DB3BA" bold="true" box="[300,460,933,952]" pageId="2" pageNumber="133">Male genitalia.</emphasis> <emphasis id="88A3EAE5D87AFFECFEC8FC4DFE2DB3BA" bold="true" box="[300,460,933,952]" pageId="2" pageNumber="133">Male genitalia.</emphasis>
Parameres simple, tapering to apex in lateral view, apical portion rounded in dorsal view. The internal sac of the aedeagus with three hooks, two spine-like and one spoon-like ( Parameres simple, tapering to apex in lateral view, apical portion rounded in dorsal view. The internal sac of the aedeagus with three hooks, two spine-like and one spoon-like (
<figureCitation id="22EC2A72D87AFFECFEF3FC11FEADB40E" box="[279,332,1017,1036]" captionStart="Figures 4–5" captionStartId="2.[833,896,1056,1070]" captionTargetBox="[867,1471,150,1024]" captionTargetId="figure-619@2.[864,1472,149,1026]" captionTargetPageId="2" captionText="Figures 4–5. A. tuza sp. nov.; 4, drawing of the internal sac, line equals 1 mm; 5, A. hornai (Balthasar, 1939), dorsal habitus of the female holotype." pageId="2" pageNumber="133">Fig. 4</figureCitation> <figureCitation id="22EC2A72D87AFFECFEF3FC11FEADB40E" box="[279,332,1017,1036]" captionStart="Figures 4–5" captionStartId="2.[833,896,1056,1070]" captionTargetBox="[867,1471,150,1024]" captionTargetId="figure-619@2.[864,1472,149,1026]" captionTargetPageId="2" captionText="Figures 4–5. A. tuza sp. nov.; 4, drawing of the internal sac, line equals 1 mm; 5, A. hornai (Balthasar, 1939), dorsal habitus of the female holotype." figureDoi="http://doi.org/10.5281/zenodo.13198490" httpUri="https://zenodo.org/record/13198490/files/figure.png" pageId="2" pageNumber="133">Fig. 4</figureCitation>
). ).
</paragraph> </paragraph>
<paragraph id="BA6836F7D87AFFECFF96FBFCFE99B4B1" blockId="2.[114,786,152,1985]" pageId="2" pageNumber="133"> <paragraph id="BA6836F7D87AFFECFF96FBFCFE99B4B1" blockId="2.[114,786,152,1985]" pageId="2" pageNumber="133">
@ -178,7 +179,7 @@ Parameres simple, tapering to apex in lateral view, apical portion rounded in do
</emphasis> </emphasis>
Female. Total length 9.5 mm. Maximum width 5.0 mm. Same as male with the following sexual differences: clypeal margin anteriorly, not so broadly V-shaped; lateral pronotal margin not arched; last abdominal segment broader medially; foretibiae and forefemora shorter and foretibial spurs slender and slightly bent at tip; pygidium not as long. Female. Total length 9.5 mm. Maximum width 5.0 mm. Same as male with the following sexual differences: clypeal margin anteriorly, not so broadly V-shaped; lateral pronotal margin not arched; last abdominal segment broader medially; foretibiae and forefemora shorter and foretibial spurs slender and slightly bent at tip; pygidium not as long.
</paragraph> </paragraph>
<caption id="EEA8667FD87AFFECFCA5FBC8FAA1B447" pageId="2" pageNumber="133" startId="2.[833,896,1056,1070]" targetBox="[867,1471,150,1024]" targetPageId="2" targetType="figure"> <caption id="EEA8667FD87AFFECFCA5FBC8FAA1B447" ID-DOI="http://doi.org/10.5281/zenodo.13198490" ID-Zenodo-Dep="13198490" httpUri="https://zenodo.org/record/13198490/files/figure.png" pageId="2" pageNumber="133" startId="2.[833,896,1056,1070]" targetBox="[867,1471,150,1024]" targetPageId="2" targetType="figure">
<paragraph id="BA6836F7D87AFFECFCA5FBC8FAA1B447" blockId="2.[833,1505,1055,1093]" pageId="2" pageNumber="133"> <paragraph id="BA6836F7D87AFFECFCA5FBC8FAA1B447" blockId="2.[833,1505,1055,1093]" pageId="2" pageNumber="133">
<emphasis id="88A3EAE5D87AFFECFCA5FBC8FC46B42C" bold="true" box="[833,935,1056,1070]" pageId="2" pageNumber="133">Figures 4–5.</emphasis> <emphasis id="88A3EAE5D87AFFECFCA5FBC8FC46B42C" bold="true" box="[833,935,1056,1070]" pageId="2" pageNumber="133">Figures 4–5.</emphasis>
<taxonomicName id="7DD74D74D87AFFECFC54FBF7FC07B42C" authorityName="Kohlmann &amp; Vaz-de-Mello" authorityYear="2018" box="[944,998,1055,1070]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="species" species="tuza" status="sp. nov."> <taxonomicName id="7DD74D74D87AFFECFC54FBF7FC07B42C" authorityName="Kohlmann &amp; Vaz-de-Mello" authorityYear="2018" box="[944,998,1055,1070]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="species" species="tuza" status="sp. nov.">

73
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@ -1,52 +1,53 @@
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<mods:title id="4C52C097FDB0439F8BF9E4D5279E6C61">A new key for the species of Ateuchus Weber (Coleoptera: Scarabaeidae: Scarabaeinae) occurring in Mexico, with a description of the first North American inquiline species from a rodent burrow (Rodentia: Geomydae) and new distribution records</mods:title> <mods:title id="59E858D13A91E61D3BBB406CF5895FD6">A new key for the species of Ateuchus Weber (Coleoptera: Scarabaeidae: Scarabaeinae) occurring in Mexico, with a description of the first North American inquiline species from a rodent burrow (Rodentia: Geomydae) and new distribution records</mods:title>
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<mods:affiliation id="415E037813DE2644C63DA4CA0599FD19">Universidade Federal de Mato Grosso, Instituto de Biociências, Departamento de Biologia e Zoologia, Cuiabá, MT, Brazil</mods:affiliation> <mods:affiliation id="5B12AAFFC1439DC90BDD2BDBBFAB9FB2">Universidade Federal de Mato Grosso, Instituto de Biociências, Departamento de Biologia e Zoologia, Cuiabá, MT, Brazil</mods:affiliation>
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( (
<figureCitation id="22EC2A72D87BFFEDFFB9FDC4FF72B23D" box="[93,147,556,575]" captionStart="Figures 4–5" captionStartId="2.[833,896,1056,1070]" captionTargetBox="[867,1471,150,1024]" captionTargetId="figure-619@2.[864,1472,149,1026]" captionTargetPageId="2" captionText="Figures 4–5. A. tuza sp. nov.; 4, drawing of the internal sac, line equals 1 mm; 5, A. hornai (Balthasar, 1939), dorsal habitus of the female holotype." pageId="3" pageNumber="134">Fig. 5</figureCitation> <figureCitation id="22EC2A72D87BFFEDFFB9FDC4FF72B23D" box="[93,147,556,575]" captionStart="Figures 4–5" captionStartId="2.[833,896,1056,1070]" captionTargetBox="[867,1471,150,1024]" captionTargetId="figure-619@2.[864,1472,149,1026]" captionTargetPageId="2" captionText="Figures 4–5. A. tuza sp. nov.; 4, drawing of the internal sac, line equals 1 mm; 5, A. hornai (Balthasar, 1939), dorsal habitus of the female holotype." figureDoi="http://doi.org/10.5281/zenodo.13198490" httpUri="https://zenodo.org/record/13198490/files/figure.png" pageId="3" pageNumber="134">Fig. 5</figureCitation>
) )
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