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<document id="4A284B15C7FC9CBF919A6573C22F1A73" ID-DOI="10.11646/zootaxa.5556.1.18" ID-ISSN="1175-5326" ID-Zenodo-Dep="14595171" ID-ZooBank="2B04A638-E500-441C-80C4-A0502F137BAB" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1735930996841" checkinUser="plazi" docAuthor="Kolesnikov, Vasiliy B. &amp; Leonov, Vladislav D." docDate="2024" docId="6D6187A43673F448C5CC7593FA0FF840" docLanguage="en" docName="zootaxa.5556.1.18.pdf" docOrigin="Zootaxa 5556 (1)" docSource="https://doi.org/10.11646/zootaxa.5556.1.18" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.23:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="23" docTitle="Galumnella Berlese 1916" docType="key" docVersion="1" lastPageNumber="250" masterDocId="9158FFDC367BF440C55B7504FF92FFE6" masterDocTitle="A new species of Galumnella (Acari, Oribatida, Galumnellidae) from Vietnam" masterLastPageNumber="251" masterPageNumber="242" pageNumber="250" updateTime="1736011292691" updateUser="felipe" zenodo-license-document="CLOSED">
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<mods:title id="27959E43D665CC303ABC6D4768501C72">A new species of Galumnella (Acari, Oribatida, Galumnellidae) from Vietnam</mods:title>
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<mods:namePart id="455A301F98A44F3ECD12C814E0D26E91">Kolesnikov, Vasiliy B.</mods:namePart>
<mods:affiliation id="DB3AAB7D91273E9D95CF1336A161E283">Papanin Institute for Biology of Inland Waters, Russian Academy of Sciences, 152742 Borok, Yaroslavl Province, Russia. &amp; Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, 625003 Tyumen, Russia.</mods:affiliation>
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<mods:namePart id="3AFFF5CF4F61A904743BD7858D003F74">Leonov, Vladislav D.</mods:namePart>
<mods:affiliation id="E6FBD8518DAAFEC97E11F1D547331A6E">A. N. Severtsov Institute of Ecology and Evolution RAS, 119071, Russia</mods:affiliation>
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<subSubSection id="ADD265393673F448C5CC7593FC21FF57" box="[151,947,151,178]" pageId="8" pageNumber="250" type="nomenclature">
<paragraph id="E57736B23673F448C5CC7593FC21FF57" blockId="8.[151,947,151,178]" box="[151,947,151,178]" pageId="8" pageNumber="250">
<heading id="BE3F81DE3673F448C5CC7593FC21FF57" bold="true" box="[151,947,151,178]" fontSize="11" level="1" pageId="8" pageNumber="250" reason="1">
<emphasis id="D7BCEAA03673F448C5CC7593FC21FF57" bold="true" box="[151,947,151,178]" pageId="8" pageNumber="250">
Key to known species of
<taxonomicName id="22C84D313673F448C4917593FDCBFF54" ID-CoL="4LZ7" authority="Berlese, 1916" authorityName="Berlese" authorityYear="1916" box="[458,601,151,178]" class="Arachnida" family="Galumnellidae" genus="Galumnella" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="250" phylum="Arthropoda" rank="genus">
<emphasis id="D7BCEAA03673F448C4917593FDCBFF54" bold="true" box="[458,601,151,178]" italics="true" pageId="8" pageNumber="250">Galumnella</emphasis>
</taxonomicName>
from Asia and New
<collectingCountry id="9DDF76223673F448C6037593FC21FF57" box="[856,947,151,177]" name="Guinea-Bissau" pageId="8" pageNumber="250">Guinea</collectingCountry>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="ADD265393673F448C5CC75DEFACAFEFE" pageId="8" pageNumber="250" type="multiple">
<paragraph id="E57736B23673F448C5CC75DEFACAFEFE" blockId="8.[151,1436,218,280]" pageId="8" pageNumber="250">
For the Asian region, 18 species have been described (
<bibRefCitation id="81594B433673F448C64075DEFB1BFF12" author="Corpuz-Raros, L. &amp; Ermilov, S. G." box="[795,1161,218,244]" pageId="8" pageNumber="250" pagination="1 - 216" refId="ref6638" refString="Corpuz-Raros, L. &amp; Ermilov, S. G. (2020) Catalogue of oribatid mites (Acari: Oribatida) from Continental Southeast Asia. Zootaxa, 4893 (1), 1 - 216. https: // doi. org / 10.11646 / zootaxa. 4893.1.1" type="journal article" year="2020">Corpuz-Raros &amp; Ermilov 2020</bibRefCitation>
;
<bibRefCitation id="81594B433673F448C1CC75DEFABBFF12" author="Subias, L. S." box="[1175,1321,218,244]" pageId="8" pageNumber="250" pagination="1 - 538" refId="ref7210" refString="Subias, L. S. (2022) Listado sistematico, sinonimico y biogeografico de los acaros oribatidos (Acariformes: Oribatida) del mundo (excepto fosiles). Monografias Electronicas Sociedad Entomologica Aragonesa, 12, 1 - 538." type="journal article" year="2022">Subías 2022</bibRefCitation>
), and one Madagascan species,
<taxonomicName id="22C84D313673F448C4D275FAFD3AFEFE" authority="Balogh, 1961" authorityName="Balogh" authorityYear="1961" box="[393,680,254,280]" class="Arachnida" family="Galumnellidae" genus="Galumnella" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="250" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="D7BCEAA03673F448C4D275FAFD9AFEFE" box="[393,520,254,280]" italics="true" pageId="8" pageNumber="250">G. pauliani</emphasis>
Balogh, 1961
</taxonomicName>
, has additionally been recorded for
<collectingCountry id="9DDF76223673F448C16275FAFB0AFEFE" box="[1081,1176,254,280]" name="Vietnam" pageId="8" pageNumber="250">Vietnam</collectingCountry>
(Vu
<emphasis id="D7BCEAA03673F448C18B75FBFA9AFEFE" box="[1232,1288,254,280]" italics="true" pageId="8" pageNumber="250">et al.</emphasis>
2014).
</paragraph>
</subSubSection>
<subSubSection id="ADD265393673F448C5CC7447FA0FF840" pageId="8" pageNumber="250" type="key">
<key id="915961833673F448C5CC7447FA0BF840" pageId="8" pageNumber="250">
<keyStep id="5E3C2E173673F448C5CC7447FA0EFE93" pageId="8" pageNumber="250">
<paragraph id="E57736B23673F448C5CC7447FA0EFEBE" blockId="8.[151,1437,323,1958]" box="[151,1436,323,345]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC7447FA0EFEBE" box="[151,1436,323,345]" pageId="8" pageNumber="250">1 Notogaster without network pattern....................................................................... 2</keyLead>
</paragraph>
<paragraph id="E57736B23673F448C5CC7465FA0EFE93" blockId="8.[151,1437,323,1958]" box="[151,1436,352,374]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC7465FA0EFE93" box="[151,1436,352,374]" pageId="8" pageNumber="250">- Notogaster with network pattern......................................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="5E3C2E173673F448C5CC747AFA0EFE2B" pageId="8" pageNumber="250">
<paragraph id="E57736B23673F448C5CC747AFA0FFE57" blockId="8.[151,1437,323,1958]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC747AFA0FFE57" pageId="8" pageNumber="250">
2 Posterior edge of notogaster with two small notches, body size: 380 × 290...............................................................................
<taxonomicName id="22C84D313673F448C7EC749FFC3DFE57" authority="Balogh, 1960" authorityName="Balogh" authorityYear="1960" box="[695,943,411,433]" class="Arachnida" family="Galumnellidae" genus="Galumnella" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="250" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="D7BCEAA03673F448C7EC749FFCB6FE56" box="[695,804,411,432]" italics="true" pageId="8" pageNumber="250">G. pauliani</emphasis>
Balogh, 1960
</taxonomicName>
. Distribution:
<collectingCountry id="9DDF76223673F448C163749FFB3EFE57" box="[1080,1196,411,433]" name="Madagascar" pageId="8" pageNumber="250">Madagascar</collectingCountry>
(
<typeStatus id="3A7388103673F448C1E2749FFB76FE56" box="[1209,1252,411,432]" pageId="8" pageNumber="250">type</typeStatus>
locality),
<collectingCountry id="9DDF76223673F448C01E749FFA0BFE57" box="[1349,1433,411,433]" name="Vietnam" pageId="8" pageNumber="250">Vietnam</collectingCountry>
.
</keyLead>
</paragraph>
<paragraph id="E57736B23673F448C5CC74BDFA0EFE2B" blockId="8.[151,1437,323,1958]" box="[151,1436,440,462]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC74BDFA0EFE2B" box="[151,1436,440,462]" pageId="8" pageNumber="250">- Posterior edge of notogaster rounded, without two small notches................................................ 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="5E3C2E173673F448C5CC74D2FA0FFDA5" pageId="8" pageNumber="250">
<paragraph id="E57736B23673F448C5CC74D2FA0FFDEF" blockId="8.[151,1437,323,1958]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC74D2FA0FFDEF" pageId="8" pageNumber="250">
3 Prodorsum with two large teeth laterally; rostrum elongate, its length exceeds half the length of the prodorsum from its base to the level of the lamellar setae; body size: 464457 × 348398..
<taxonomicName id="22C84D313673F448C60A74F7FB2EFDEE" authority="Ermilov &amp; Anichkin, 2013" authorityName="Ermilov &amp; Anichkin" authorityYear="2013" box="[849,1212,499,521]" class="Arachnida" family="Galumnellidae" genus="Galumnella" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="250" phylum="Arthropoda" rank="species" species="tiunovi">
<emphasis id="D7BCEAA03673F448C60A74F7FC20FDEE" box="[849,946,499,520]" italics="true" pageId="8" pageNumber="250">G. tiunovi</emphasis>
Ermilov &amp; Anichkin, 2013
</taxonomicName>
. Distribution:
<collectingCountry id="9DDF76223673F448C01E74F7FA0BFDEF" box="[1349,1433,499,521]" name="Vietnam" pageId="8" pageNumber="250">Vietnam</collectingCountry>
.
</keyLead>
</paragraph>
<paragraph id="E57736B23673F448C5CC7715FA0FFDA5" blockId="8.[151,1437,323,1958]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC7715FA0FFDA5" pageId="8" pageNumber="250">
- Prodorsum without two large teeth laterally; rostrum shorter than half the length of the prodorsum from its base to the level of the lamellar setae; body size: 400416 × 288256.............
<taxonomicName id="22C84D313673F448C633772AFB17FDA5" authority="Balakrishnan, 1989" authorityName="Balakrishnan" authorityYear="1989" box="[872,1157,557,579]" class="Arachnida" family="Galumnellidae" genus="Galumnella" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="250" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="D7BCEAA03673F448C633772AFC50FDA4" box="[872,962,557,579]" italics="true" pageId="8" pageNumber="250">G. indica</emphasis>
Balakrishnan, 1989
</taxonomicName>
. Distribution:
<collectingCountry id="9DDF76223673F448C0547729FAD2FDA5" box="[1295,1344,557,579]" name="India" pageId="8" pageNumber="250">India</collectingCountry>
(
<collectingRegion id="270CF8503673F448C0167729FA00FDA5" box="[1357,1426,557,579]" country="India" name="Kerala" pageId="8" pageNumber="250">Kerala</collectingRegion>
).
</keyLead>
</paragraph>
</keyStep>
<keyStep id="5E3C2E173673F448C5CC774FFA0EFD9B" pageId="8" pageNumber="250">
<paragraph id="E57736B23673F448C5CC774FFA0EFD87" blockId="8.[151,1437,323,1958]" box="[151,1436,587,609]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC774FFA0EFD87" box="[151,1436,587,609]" pageId="8" pageNumber="250">4 Prodorsum with network pattern or irregular rugae.......................................................... 5</keyLead>
</paragraph>
<paragraph id="E57736B23673F448C5CC776DFA0EFD9B" blockId="8.[151,1437,323,1958]" box="[151,1436,616,638]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC776DFA0EFD9B" box="[151,1436,616,638]" pageId="8" pageNumber="250">- Prodorsum without network pattern or irregular rugae........................................................ 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="5E3C2E173673F448C5CC7781FA0EFD30" pageId="8" pageNumber="250">
<paragraph id="E57736B23673F448C5CC7781FA0FFD5E" blockId="8.[151,1437,323,1958]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC7781FA0FFD5E" pageId="8" pageNumber="250">
5 Anterior half of prodorsum and marginal area of pteromorphs with irregular rugae; body size: 415460 × 305370......................................................................
<taxonomicName id="22C84D313673F448C69777A7FB44FD5E" authority="Aoki, 1970" authorityName="Aoki" authorityYear="1970" box="[972,1238,675,697]" class="Arachnida" family="Galumnellidae" genus="Galumnella" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="250" phylum="Arthropoda" rank="species" species="angustifrons">
<emphasis id="D7BCEAA03673F448C69777A7FBF3FD5E" box="[972,1121,675,696]" italics="true" pageId="8" pageNumber="250">G. angustifrons</emphasis>
<bibRefCitation id="81594B433673F448C13C77A7FB44FD5E" author="Aoki, J." box="[1127,1238,675,697]" pageId="8" pageNumber="250" pagination="395 - 442" refId="ref6604" refString="Aoki, J. (1970) The oribatid mites of the Islands of Tsushima. Bulletin of the National Science Museum, Tokyo, 13 (3), 395 - 442." type="journal article" year="1970">Aoki, 1970</bibRefCitation>
</taxonomicName>
. Distribution:
<collectingCountry id="9DDF76223673F448C03B77A7FA08FD5E" box="[1376,1434,675,696]" name="Japan" pageId="8" pageNumber="250">Japan</collectingCountry>
.
</keyLead>
</paragraph>
<paragraph id="E57736B23673F448C5CC77C5FA0EFD30" blockId="8.[151,1437,323,1958]" box="[151,1436,704,726]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC77C5FA0EFD30" box="[151,1436,704,726]" pageId="8" pageNumber="250">- Anterior half of prodorsum and marginal area of pteromorphs without irregular rugae............................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="5E3C2E173673F448C5CC77D9FA0EFCC8" pageId="8" pageNumber="250">
<paragraph id="E57736B23673F448C5CC77D9FA0FFCF7" blockId="8.[151,1437,323,1958]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC77D9FA0FFCF7" pageId="8" pageNumber="250">
6 Pteromorphs without network pattern; posterior edge of notogaster corrugated; body size: 381398 × 265282...........................................
<taxonomicName id="22C84D313673F448C70177FFFC5EFCF7" authority="Ermilov et Kaluz" authorityName="Ermilov et Kaluz" box="[602,972,763,785]" class="Arachnida" family="Galumnellidae" genus="Galumnella" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="250" phylum="Arthropoda" rank="species" species="parageographica">
<emphasis id="D7BCEAA03673F448C70177FFFC8CFCF6" box="[602,798,763,784]" italics="true" pageId="8" pageNumber="250">G. parageographica</emphasis>
Ermilov
<emphasis id="D7BCEAA03673F448C62277FFFC18FCF6" box="[889,906,763,784]" italics="true" pageId="8" pageNumber="250">et</emphasis>
Kalúz
</taxonomicName>
,
<superScript id="12BD9BFA3673F448C68E77FFFB9EFCE0" attach="none" box="[981,1036,762,784]" fontSize="5" pageId="8" pageNumber="250">20131</superScript>
. Distribution:
<collectingCountry id="9DDF76223673F448C1C277FFFB58FCF7" box="[1177,1226,763,785]" name="India" pageId="8" pageNumber="250">India</collectingCountry>
(
<collectingRegion id="270CF8503673F448C18C77FFFA01FCF7" box="[1239,1427,763,785]" country="India" name="Arunachal Pradesh" pageId="8" pageNumber="250">Arunachal Pradesh</collectingRegion>
).
</keyLead>
</paragraph>
<paragraph id="E57736B23673F448C5CC761DFA0EFCC8" blockId="8.[151,1437,323,1958]" box="[151,1436,792,814]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC761DFA0EFCC8" box="[151,1436,792,814]" pageId="8" pageNumber="250">- Pteromorphs with network pattern........................................................................ 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="5E3C2E173673F448C5CC7632FA0EFC63" pageId="8" pageNumber="250">
<paragraph id="E57736B23673F448C5CC7632FA0FFC8F" blockId="8.[151,1437,323,1958]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC7632FA0FFC8F" pageId="8" pageNumber="250">
7 Interlamellar ridge on prodorsum present, distinct; adanal seta
<emphasis id="D7BCEAA03673F448C6197631FCC8FCAC" box="[834,858,821,842]" italics="true" pageId="8" pageNumber="250">ad</emphasis>
3 located anterolateral to anal aperture; postanal porose area absent; body size: 381398 × 265282....................................
<taxonomicName id="22C84D313673F448C1507657FBFEFC8E" authorityName="Kolesnikov &amp; Leonov" authorityYear="2024" box="[1035,1132,851,872]" class="Arachnida" family="Galumnellidae" genus="Galumnella" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="250" phylum="Arthropoda" rank="species" species="subiasi" status="sp. nov.">
<emphasis id="D7BCEAA03673F448C1507657FBFEFC8E" box="[1035,1132,851,872]" italics="true" pageId="8" pageNumber="250">G. subiasi</emphasis>
</taxonomicName>
<emphasis id="D7BCEAA03673F448C1297657FB2DFC8E" bold="true" box="[1138,1215,851,872]" pageId="8" pageNumber="250">
<taxonomicNameLabel id="CC8F57DB3673F448C1297657FB2DFC8E" box="[1138,1215,851,872]" pageId="8" pageNumber="250" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Distribution:
<collectingCountry id="9DDF76223673F448C01E7657FA0BFC8F" box="[1349,1433,851,873]" name="Vietnam" pageId="8" pageNumber="250">Vietnam</collectingCountry>
.
</keyLead>
</paragraph>
<paragraph id="E57736B23673F448C5CC7675FA0EFC63" blockId="8.[151,1437,323,1958]" box="[151,1436,880,907]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC7675FA0EFC63" box="[151,1436,880,907]" pageId="8" pageNumber="250">
- Interlamellar ridge on prodorsum absent; adanal seta
<emphasis id="D7BCEAA03673F448C7BA7674FD6BFC63" box="[737,761,880,901]" italics="true" pageId="8" pageNumber="250">ad</emphasis>
<subScript id="794C34F73673F448C7A2767BFC92FC6D" attach="left" box="[761,768,895,907]" fontSize="5" pageId="8" pageNumber="250">3</subScript>
paraanal; postanal porose area present....................... 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="5E3C2E173673F448C5CC768AFA0FFC1D" pageId="8" pageNumber="250">
<paragraph id="E57736B23673F448C5CC768AFA0FFC27" blockId="8.[151,1437,323,1958]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC768AFA0FFC27" pageId="8" pageNumber="250">
8 Lamellar seta setiform; rostrum projecting, but not strongly narrowed anteriorly; postanal porose area small; body size: 374 415 × 299327......................
<taxonomicName id="22C84D313673F448C72576AFFC26FC27" authority="Mahunka, 1995" authorityName="Mahunka" authorityYear="1995" box="[638,948,939,961]" class="Arachnida" family="Galumnellidae" genus="Galumnella" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="250" phylum="Arthropoda" rank="species" species="geographica">
<emphasis id="D7BCEAA03673F448C72576AFFC86FC26" box="[638,788,939,960]" italics="true" pageId="8" pageNumber="250">G. geographica</emphasis>
Mahunka, 1995
</taxonomicName>
. Distribution: Borneo (
<typeStatus id="3A7388103673F448C1CA76AFFB2EFC26" box="[1169,1212,939,960]" pageId="8" pageNumber="250">type</typeStatus>
location) and
<collectingCountry id="9DDF76223673F448C01E76AFFA0BFC27" box="[1349,1433,939,961]" name="Vietnam" pageId="8" pageNumber="250">Vietnam</collectingCountry>
.
</keyLead>
</paragraph>
<paragraph id="E57736B23673F448C5CC76CDFA0FFC1D" blockId="8.[151,1437,323,1958]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC76CDFA0FFC1D" pageId="8" pageNumber="250">
- Lamellar setae alveolar; rostrum strongly narrowed anteriorly; postanal porose area very small; body size: 302355 × 242 288............................................
<taxonomicName id="22C84D313673F448C64776E2FBC1FC1D" authority="(Aoki &amp; Hu, 1993)" baseAuthorityName="Aoki &amp; Hu" baseAuthorityYear="1993" box="[796,1107,997,1019]" class="Arachnida" family="Galumnellidae" genus="Galumnella" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="250" phylum="Arthropoda" rank="species" species="pulchella">
<emphasis id="D7BCEAA03673F448C64776E2FC01FC1C" box="[796,915,997,1019]" italics="true" pageId="8" pageNumber="250">G. pulchella</emphasis>
(Aoki &amp; Hu, 1993)
</taxonomicName>
. Distribution:
<collectingCountry id="9DDF76223673F448C18676E1FA84FC1D" box="[1245,1302,997,1019]" name="China" pageId="8" pageNumber="250">China</collectingCountry>
and
<collectingCountry id="9DDF76223673F448C01E76E1FA0BFC1D" box="[1349,1433,997,1019]" name="Vietnam" pageId="8" pageNumber="250">Vietnam</collectingCountry>
.
</keyLead>
</paragraph>
</keyStep>
<keyStep id="5E3C2E173673F448C5CC7107FA0EFBD3" pageId="8" pageNumber="250">
<paragraph id="E57736B23673F448C5CC7107FA0EFBFE" blockId="8.[151,1437,323,1958]" box="[151,1436,1027,1049]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC7107FA0EFBFE" box="[151,1436,1027,1049]" pageId="8" pageNumber="250">9 Pteromorphs with network pattern....................................................................... 10</keyLead>
</paragraph>
<paragraph id="E57736B23673F448C5CC7125FA0EFBD3" blockId="8.[151,1437,323,1958]" box="[151,1436,1056,1078]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC7125FA0EFBD3" box="[151,1436,1056,1078]" pageId="8" pageNumber="250">- Pteromorphs without network pattern.................................................................... 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="5E3C2E173673F448C5CC713AFA0EFB96" pageId="8" pageNumber="250">
<paragraph id="E57736B23673F448C5CC713AFA0FFBB5" blockId="8.[151,1437,323,1958]" box="[151,1437,1085,1107]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC713AFA0FFBB5" box="[151,1437,1085,1107]" pageId="8" pageNumber="250">
10 Bothridial seta setiform; body size: 348381 × 265282...
<taxonomicName id="22C84D313673F448C647713AFB2EFBB5" authority="Ermilov &amp; Anichkin, 2011" authorityName="Ermilov &amp; Anichkin" authorityYear="2011" box="[796,1212,1085,1107]" class="Arachnida" family="Galumnellidae" genus="Galumnella" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="250" phylum="Arthropoda" rank="species" species="microporosa">
<emphasis id="D7BCEAA03673F448C647713AFC20FBB5" box="[796,946,1086,1107]" italics="true" pageId="8" pageNumber="250">G. microporosa</emphasis>
<bibRefCitation id="81594B433673F448C6E37139FB2EFBB5" author="Ermilov, S. G. &amp; Anichkin, A. E." box="[952,1212,1085,1107]" pageId="8" pageNumber="250" pagination="242 - 251" refId="ref6731" refString="Ermilov, S. G. &amp; Anichkin, A. E. (2011) New oribatid mites of the genera Pergalumna and Galumnella (Acari, Oribatida, Galumnoidea) from Vietnam. Acarina, 19 (2), 242 - 251. https: // doi. org / 10.1080 / 01647954.2010.528799" type="journal article" year="2011">Ermilov &amp; Anichkin, 2011</bibRefCitation>
</taxonomicName>
. Distribution:
<collectingCountry id="9DDF76223673F448C01E7139FA0BFBB5" box="[1349,1433,1085,1107]" name="Vietnam" pageId="8" pageNumber="250">Vietnam</collectingCountry>
.
</keyLead>
</paragraph>
<paragraph id="E57736B23673F448C5CC715FFA0EFB96" blockId="8.[151,1437,323,1958]" box="[151,1436,1115,1137]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC715FFA0EFB96" box="[151,1436,1115,1137]" pageId="8" pageNumber="250">- Bothridial setae with head dilated unilaterally.............................................................. 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="5E3C2E173673F448C5CC717CFA0EFB4D" pageId="8" pageNumber="250">
<paragraph id="E57736B23673F448C5CC717CFA0EFB6B" blockId="8.[151,1437,323,1958]" box="[151,1436,1144,1166]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC717CFA0EFB6B" box="[151,1436,1144,1166]" pageId="8" pageNumber="250">11 Bothridial setae smooth............................................................................... 12</keyLead>
</paragraph>
<paragraph id="E57736B23673F448C5CC7192FA0EFB4D" blockId="8.[151,1437,323,1958]" box="[151,1436,1173,1195]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC7192FA0EFB4D" box="[151,1436,1173,1195]" pageId="8" pageNumber="250">- Bothridial setae barbed................................................................................ 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="5E3C2E173673F448C5CC71B7FA0FFAC7" pageId="8" pageNumber="250">
<paragraph id="E57736B23673F448C5CC71B7FA0FFB00" blockId="8.[151,1437,323,1958]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC71B7FA0FFB00" pageId="8" pageNumber="250">
12 Pteromorphs with large foveolae; postanal porose area present; body size: 275 × 195...................................................................................
<taxonomicName id="22C84D313673F448C61B71D4FB2EFB00" authority="Balogh &amp; Mahunka, 1967" authorityName="Balogh &amp; Mahunka" authorityYear="1967" box="[832,1212,1232,1254]" class="Arachnida" family="Galumnellidae" genus="Galumnella" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="250" phylum="Arthropoda" rank="species" species="cellularis">
<emphasis id="D7BCEAA03673F448C61B71D4FC2AFB03" box="[832,952,1232,1253]" italics="true" pageId="8" pageNumber="250">G. cellularis</emphasis>
Balogh &amp; Mahunka, 1967
</taxonomicName>
. Distribution:
<collectingCountry id="9DDF76223673F448C01E71D4FA0BFB00" box="[1349,1433,1232,1254]" name="Vietnam" pageId="8" pageNumber="250">Vietnam</collectingCountry>
.
</keyLead>
</paragraph>
<paragraph id="E57736B23673F448C5CC71EAFA0FFAC7" blockId="8.[151,1437,323,1958]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC71EAFA0FFAC7" pageId="8" pageNumber="250">
- Pteromorphs with microfoveolae; postanal porose area absent; body size: 350410 × 260280....................................................................
<taxonomicName id="22C84D313673F448C7B1700FFB41FAC6" authority="Liang, Yang, Ren &amp; Q. Zheng, 2019" authorityName="Liang, Yang, Ren &amp; Q. Zheng" authorityYear="2019" box="[746,1235,1291,1313]" class="Arachnida" family="Galumnellidae" genus="Galumnella" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="250" phylum="Arthropoda" rank="species" species="nonporosa">
<emphasis id="D7BCEAA03673F448C7B1700FFCFCFAC6" box="[746,878,1291,1312]" italics="true" pageId="8" pageNumber="250">G. nonporosa</emphasis>
Liang, Yang, Ren &amp; Q. Zheng, 2019
</taxonomicName>
. Distribution:
<collectingCountry id="9DDF76223673F448C006700FFA0BFAC7" box="[1373,1433,1291,1313]" name="China" pageId="8" pageNumber="250">China</collectingCountry>
.
</keyLead>
</paragraph>
</keyStep>
<keyStep id="5E3C2E173673F448C5CC702CFA0FFA70" pageId="8" pageNumber="250">
<paragraph id="E57736B23673F448C5CC702CFA0FFABD" blockId="8.[151,1437,323,1958]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC702CFA0FFABD" pageId="8" pageNumber="250">
13 Tarsi with one claw; notogaster with polygonal network pattern; body size: 370450 × 280320.................................................................
<taxonomicName id="22C84D313673F448C7877042FB41FABD" authority="Liang, Yang, Ren &amp; Q. Zheng, 2019" authorityName="Liang, Yang, Ren &amp; Q. Zheng" authorityYear="2019" box="[732,1235,1349,1371]" class="Arachnida" family="Galumnellidae" genus="Galumnella" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="250" phylum="Arthropoda" rank="species" species="sidorchukae">
<emphasis id="D7BCEAA03673F448C7877042FCFFFABC" box="[732,877,1349,1371]" italics="true" pageId="8" pageNumber="250">G. sidorchukae</emphasis>
Liang, Yang, Ren &amp; Q. Zheng, 2019
</taxonomicName>
. Distribution:
<collectingCountry id="9DDF76223673F448C0067041FA0BFABD" box="[1373,1433,1349,1371]" name="China" pageId="8" pageNumber="250">China</collectingCountry>
.
</keyLead>
</paragraph>
<paragraph id="E57736B23673F448C5CC7067FA0FFA70" blockId="8.[151,1437,323,1958]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC7067FA0FFA70" pageId="8" pageNumber="250">
- Tarsi with two claws; notogaster with irregular network pattern; body size: 294316 × 223240.......................................................
<taxonomicName id="22C84D313673F448C73A7084FBAAFA73" authority="(Mahunka, 1994)" baseAuthorityName="Mahunka" baseAuthorityYear="1994" box="[609,1080,1408,1430]" class="Arachnida" family="Galumnellidae" genus="Galumnella" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="250" phylum="Arthropoda" rank="species" species="csavasorum" subGenus="Bigalumnella">
<emphasis id="D7BCEAA03673F448C73A7084FC1BFA70" box="[609,905,1408,1430]" italics="true" pageId="8" pageNumber="250">G. (Bigalumnella) csavasorum</emphasis>
(Mahunka, 1994)
</taxonomicName>
. Distribution:
<collectingCountry id="9DDF76223673F448C19A7084FA84FA70" box="[1217,1302,1408,1430]" name="Thailand" pageId="8" pageNumber="250">Thailand</collectingCountry>
and
<collectingCountry id="9DDF76223673F448C01E7084FA0BFA70" box="[1349,1433,1408,1430]" name="Vietnam" pageId="8" pageNumber="250">Vietnam</collectingCountry>
.
</keyLead>
</paragraph>
</keyStep>
<keyStep id="5E3C2E173673F448C5CC709AFA0EFA37" pageId="8" pageNumber="250">
<paragraph id="E57736B23673F448C5CC709AFA0EFA55" blockId="8.[151,1437,323,1958]" box="[151,1436,1437,1459]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC709AFA0EFA55" box="[151,1436,1437,1459]" pageId="8" pageNumber="250">14 Interlamellar ridge on prodorsum present................................................................. 15</keyLead>
</paragraph>
<paragraph id="E57736B23673F448C5CC70BFFA0EFA37" blockId="8.[151,1437,323,1958]" box="[151,1436,1467,1489]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC70BFFA0EFA37" box="[151,1436,1467,1489]" pageId="8" pageNumber="250">- Interlamellar median ridge on prodorsum absent or poorly visible.............................................. 16</keyLead>
</paragraph>
</keyStep>
<keyStep id="5E3C2E173673F448C5CC70DCFA0FF9A0" pageId="8" pageNumber="250">
<paragraph id="E57736B23673F448C5CC70DCFA0FF9ED" blockId="8.[151,1437,323,1958]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC70DCFA0FF9ED" pageId="8" pageNumber="250">
15 Bothridial setae barbed; ventral shield without longitudinal ridge; body size: 298 × 235........................................................................................
<taxonomicName id="22C84D313673F448C6C870F2FB06F9ED" authority="Balogh, 1970" authorityName="Balogh" authorityYear="1970" box="[915,1172,1525,1547]" class="Arachnida" family="Galumnellidae" genus="Galumnella" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="250" phylum="Arthropoda" rank="species" species="woschitzi">
<emphasis id="D7BCEAA03673F448C6C870F2FB9BF9EC" box="[915,1033,1525,1547]" italics="true" pageId="8" pageNumber="250">G. woschitzi</emphasis>
Balogh, 1970
</taxonomicName>
. Distribution: New
<collectingCountry id="9DDF76223673F448C00A70F1FA0BF9ED" box="[1361,1433,1525,1547]" name="Guinea-Bissau" pageId="8" pageNumber="250">Guinea</collectingCountry>
.
</keyLead>
</paragraph>
<paragraph id="E57736B23673F448C5CC7317FA0FF9A0" blockId="8.[151,1437,323,1958]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC7317FA0FF9A0" pageId="8" pageNumber="250">
- Bothridial setae smooth; ventral shield with longitudinal ridge; body size: 398431 × 282315..................................................................
<taxonomicName id="22C84D313673F448C78D7334FB33F9A3" authority="Ermilov &amp; Corpuz-Raros, 2017" authorityName="Ermilov &amp; Corpuz-Raros" authorityYear="2017" box="[726,1185,1584,1606]" class="Arachnida" family="Galumnellidae" genus="Galumnella" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="250" phylum="Arthropoda" rank="species" species="quezonensis">
<emphasis id="D7BCEAA03673F448C78D7334FCFAF9A0" box="[726,872,1584,1606]" italics="true" pageId="8" pageNumber="250">G. quezonensis</emphasis>
Ermilov &amp; Corpuz-Raros, 2017
</taxonomicName>
. Distribution:
<collectingCountry id="9DDF76223673F448C0717334FA0BF9A0" box="[1322,1433,1584,1606]" name="Philippines" pageId="8" pageNumber="250">Philippines</collectingCountry>
.
</keyLead>
</paragraph>
</keyStep>
<keyStep id="5E3C2E173673F448C5CC7349FA0EF97B" pageId="8" pageNumber="250">
<paragraph id="E57736B23673F448C5CC7349FA0FF967" blockId="8.[151,1437,323,1958]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC7349FA0FF967" pageId="8" pageNumber="250">
16 Bothridial setae short, with strongly swollеn hеad; body size: 280300 × 220230........................................................................................
<taxonomicName id="22C84D313673F448C63D736FFBF3F966" authority="Aoki, 2009" authorityName="Aoki" authorityYear="2009" box="[870,1121,1643,1665]" class="Arachnida" family="Galumnellidae" genus="Galumnella" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="250" phylum="Arthropoda" rank="species" species="okinawana">
<emphasis id="D7BCEAA03673F448C63D736FFC7EF966" box="[870,1004,1643,1664]" italics="true" pageId="8" pageNumber="250">G. okinawana</emphasis>
Aoki, 2009
</taxonomicName>
. Distribution:
<collectingCountry id="9DDF76223673F448C1B0736FFAB0F966" box="[1259,1314,1643,1664]" name="Japan" pageId="8" pageNumber="250">Japan</collectingCountry>
and
<collectingCountry id="9DDF76223673F448C00B736FFA0BF967" box="[1360,1433,1643,1665]" name="Taiwan" pageId="8" pageNumber="250">Taiwan</collectingCountry>
.
</keyLead>
</paragraph>
<paragraph id="E57736B23673F448C5CC738DFA0EF97B" blockId="8.[151,1437,323,1958]" box="[151,1436,1672,1694]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC738DFA0EF97B" box="[151,1436,1672,1694]" pageId="8" pageNumber="250">- Bothridial setae long, with lanceolate head or setiform, barbed or smooth........................................ 17</keyLead>
</paragraph>
</keyStep>
<keyStep id="5E3C2E173673F448C5CC73A2FA0EF913" pageId="8" pageNumber="250">
<paragraph id="E57736B23673F448C5CC73A2FA0FF93F" blockId="8.[151,1437,323,1958]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC73A2FA0FF93F" pageId="8" pageNumber="250">
17 Bothridial setae smooth; anterior margin of ventral shield with triangular projection; body size: 280300 × 220230....................................................
<taxonomicName id="22C84D313673F448C7AD73C7FB33F93E" authority="Ermilov &amp; Corpuz-Raros, 2017" authorityName="Ermilov &amp; Corpuz-Raros" authorityYear="2017" box="[758,1185,1731,1753]" class="Arachnida" family="Galumnellidae" genus="Galumnella" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="250" phylum="Arthropoda" rank="species" species="processa">
<emphasis id="D7BCEAA03673F448C7AD73C7FCF5F93E" box="[758,871,1731,1752]" italics="true" pageId="8" pageNumber="250">G. processa</emphasis>
Ermilov &amp; Corpuz-Raros, 2017
</taxonomicName>
. Distribution:
<collectingCountry id="9DDF76223673F448C07173C7FA0BF93F" box="[1322,1433,1731,1753]" name="Philippines" pageId="8" pageNumber="250">Philippines</collectingCountry>
.
</keyLead>
</paragraph>
<paragraph id="E57736B23673F448C5CC73E5FA0EF913" blockId="8.[151,1437,323,1958]" box="[151,1436,1760,1782]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC73E5FA0EF913" box="[151,1436,1760,1782]" pageId="8" pageNumber="250">- Bothridial setae barbed; anterior margin of ventral shield without triangular projection............................. 18</keyLead>
</paragraph>
</keyStep>
<keyStep id="5E3C2E173673F448C5CC73FAFA0FF840" pageId="8" pageNumber="250">
<paragraph id="E57736B23673F448C5CC73FAFA0FF8A8" blockId="8.[151,1437,323,1958]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC73FAFA0FF8A8" pageId="8" pageNumber="250">
18 Prodorsum with a dorsal longitudinal ridge and lateral tooth-like projections; anterior tectum of epimere I well developed, forming trapezoidal and distally slightly concave anterior ledge; lines
<emphasis id="D7BCEAA03673F448C6D8721FFC02F8D6" box="[899,912,1819,1840]" italics="true" pageId="8" pageNumber="250">L</emphasis>
and
<emphasis id="D7BCEAA03673F448C6E4721FFC59F8D6" box="[959,971,1819,1840]" italics="true" pageId="8" pageNumber="250">S</emphasis>
distally divergent; barbs on bothridial seta same length; body size: 315381 × 249298..
<taxonomicName id="22C84D313673F448C723723CFB33F8AB" authority="Ermilov, Corpuz-Raros &amp; Shimano, 2014" authorityName="Ermilov, Corpuz-Raros &amp; Shimano" authorityYear="2014" box="[632,1185,1848,1870]" class="Arachnida" family="Galumnellidae" genus="Galumnella" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="250" phylum="Arthropoda" rank="species" species="junichiaokii">
<emphasis id="D7BCEAA03673F448C723723CFC9AF8AB" box="[632,776,1848,1869]" italics="true" pageId="8" pageNumber="250">G. junichiaokii</emphasis>
<bibRefCitation id="81594B433673F448C655723CFB33F8AB" author="Ermilov S. G. &amp; Corpuz-Raros L. &amp; Shimano S." box="[782,1185,1848,1870]" pageId="8" pageNumber="250" pagination="85 - 91" refId="ref6849" refString="Ermilov S. G., Corpuz-Raros L. &amp; Shimano S. (2014) First reports of Galumnellidae (Acari, Oribatida) from the Philippines, with description of Galumnella junichiaokii sp. n. Acarina, 22, (2), 85 - 91." type="journal article" year="2014">Ermilov, Corpuz-Raros &amp; Shimano, 2014</bibRefCitation>
</taxonomicName>
. Distribution:
<collectingCountry id="9DDF76223673F448C071723CFA0BF8A8" box="[1322,1433,1848,1870]" name="Philippines" pageId="8" pageNumber="250">Philippines</collectingCountry>
.
</keyLead>
</paragraph>
<paragraph id="E57736B23673F448C5CC7252FA0FF840" blockId="8.[151,1437,323,1958]" pageId="8" pageNumber="250">
<keyLead id="5E3995873673F448C5CC7252FA0FF840" pageId="8" pageNumber="250">
- Prodorsum without a dorsal longitudinal ridge and lateral tooth-like projections; anterior tectum of epimere I absent; lines
<emphasis id="D7BCEAA03673F448C03C7252FAE6F88D" box="[1383,1396,1878,1899]" italics="true" pageId="8" pageNumber="250">L</emphasis>
and
<emphasis id="D7BCEAA03673F448C5B97277FF7CF86E" box="[226,238,1907,1928]" italics="true" pageId="8" pageNumber="250">S</emphasis>
parallel; barbs on bothridial seta longer at the base of the head and gradually shorten towards its end; body size: 315436 × 249336.............
<taxonomicName id="22C84D313673F448C4837294FCB5F843" authority="Suzuki &amp; Aoki, 1970" authorityName="Suzuki &amp; Aoki" authorityYear="1970" box="[472,807,1936,1958]" class="Arachnida" family="Galumnellidae" genus="Galumnella" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="250" phylum="Arthropoda" rank="species" species="nipponica">
<emphasis id="D7BCEAA03673F448C4837294FDC7F840" box="[472,597,1936,1958]" italics="true" pageId="8" pageNumber="250">G. nipponica</emphasis>
<bibRefCitation id="81594B433673F448C7007294FCB5F843" author="Suzuki, K. &amp; Aoki, J." box="[603,807,1936,1958]" pageId="8" pageNumber="250" pagination="166 - 169" refId="ref7253" refString="Suzuki, K. &amp; Aoki, J. (1970) A new species of oribatid mite, Galumnella nipponica, from central Japan (Acari: Cryptostigmata). Annotationes Zoologicae Japonenses, 43 (3), 166 - 169." type="journal article" year="1970">Suzuki &amp; Aoki, 1970</bibRefCitation>
</taxonomicName>
(see also
<bibRefCitation id="81594B433673F448C6DD7294FBDDF840" author="Fujikawa, T. &amp; Ishikawa, K. &amp; Shiba, M. &amp; Ono, H. &amp; Morikawa, K. &amp; Tamura, H. &amp; Nakamura, Y." box="[902,1103,1936,1958]" pageId="8" pageNumber="250" pagination="1 - 22" refId="ref7019" refString="Fujikawa, T., Ishikawa, K., Shiba, M., Ono, H., Morikawa, K., Tamura, H. &amp; Nakamura, Y. (2006) Soil animals from 88 temples in Shikoku Island 4. Morphological variation in the nineteen known species of oribatid mites. Edaphologia, 79, 1 - 22." type="journal article" year="2006">
Fujikawa
<emphasis id="D7BCEAA03673F448C6BD7295FB84F843" box="[998,1046,1936,1958]" italics="true" pageId="8" pageNumber="250">et al.</emphasis>
2006
</bibRefCitation>
). Distribution:
<collectingCountry id="9DDF76223673F448C1BB7295FA85F840" box="[1248,1303,1937,1958]" name="Japan" pageId="8" pageNumber="250">Japan</collectingCountry>
and
<collectingCountry id="9DDF76223673F448C01E7294FA0BF840" box="[1349,1433,1936,1958]" name="Vietnam" pageId="8" pageNumber="250">Vietnam</collectingCountry>
.
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>