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<mods:title id="D77ADBDA32635EED55F76BA7A74CECFF">New species and new records of Cosmolaelaps Berlese (Acari: Laelapidae) from Russia, with a review of the Russian species of the genus</mods:title>
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<mods:namePart id="EE08ED77A32F81705BBAA256FD466BA2">Döker, Ismail</mods:namePart>
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<mods:affiliation id="D18F81AB987D4805C7215691716F635A">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Semakova Str. 10, 625003 Tyumen, Russia. &amp; Cukurova University, Agricultural Faculty, Department of Plant Protection, Acarology Laboratory, 01330, Adana, Turkey. https: // orcid. org / 0000 - 0002 - 2741 - 4946 (Joharchi) https: // orcid. org / 0000 - 0002 - 1412 - 1554 (Döker) https: // orcid. org / 0000 - 0003 - 2831 - 7213 (Khaustov)</mods:affiliation>
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<paragraph id="5A4C6132FFA1983F01C7AAA3712CF8C9" blockId="1.[151,521,1790,1853]" box="[151,521,1790,1817]" pageId="1" pageNumber="487">
<heading id="0104D65EFFA1983F01C7AAA3712CF8C9" bold="true" box="[151,521,1790,1817]" fontSize="11" level="1" pageId="1" pageNumber="487" reason="1">
<taxonomicName id="9DF31AB1FFA1983F01C7AAA3712CF8C9" authority="(Gu)" baseAuthorityName="Gu" box="[151,521,1790,1817]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="487" phylum="Arthropoda" rank="species" species="chianensis">
<emphasis id="6887BD20FFA1983F01C7AAA3712CF8C9" bold="true" box="[151,521,1790,1817]" pageId="1" pageNumber="487">
<emphasis id="6887BD20FFA1983F01C7AAA372E2F8C8" bold="true" box="[151,455,1790,1817]" italics="true" pageId="1" pageNumber="487">Cosmolaelaps chianensis</emphasis>
(Gu)
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<subSubSection id="12E932B9FFA1983F01C7AB7E720AF8EC" box="[151,303,1827,1853]" pageId="1" pageNumber="487" type="description">
<paragraph id="5A4C6132FFA1983F01C7AB7E720AF8EC" blockId="1.[151,521,1790,1853]" box="[151,303,1827,1853]" pageId="1" pageNumber="487">
(
<figureCitation id="C2C87DB7FFA1983F01CFAB7E7203F8EC" box="[159,294,1827,1853]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1944,1968]" captionTargetBox="[182,1405,181,1920]" captionTargetId="figure-18@2.[182,1405,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–7. DIC micrographs of Cosmolaelaps chianensis (Gu), female. 1, idiosoma in dorsal view, enlarged section not to scale; 2, idiosoma in ventral view; 3, sternal shield; 4, general view of opisthogastric area; 5, chelicera; 6, dorsal aspect of femur, genu and tibia leg I; 7, ventral aspect of femur, genu and tibia leg I." figureDoi="http://doi.org/10.5281/zenodo.6530960" httpUri="https://zenodo.org/record/6530960/files/figure.png" pageId="1" pageNumber="487">Figures 1–7</figureCitation>
)
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<paragraph id="5A4C6132FFA1983F01C7AB357137F8AE" blockId="1.[151,1381,1895,2042]" box="[151,530,1895,1919]" pageId="1" pageNumber="487">
<treatmentCitationGroup id="7AE3461CFFA1983F01C7AB357137F8AE" box="[151,530,1895,1919]" pageId="1" pageNumber="487">
<treatmentCitation id="DB524723FFA1983F01C7AB35712BF8AE" author="Gu, Y. - M." box="[151,526,1895,1919]" page="441" pageId="1" pageNumber="487" year="1990">
<taxonomicName id="9DF31AB1FFA1983F01C7AB35712BF8AE" authority="Gu, 1990: 441" authorityName="Gu" authorityPageNumber="441" authorityYear="1990" box="[151,526,1895,1919]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="487" phylum="Arthropoda" rank="species" species="chianensis">
<emphasis id="6887BD20FFA1983F01C7AB357256F8AE" box="[151,371,1895,1919]" italics="true" pageId="1" pageNumber="487">Hypoaspis chianensis</emphasis>
<bibRefCitation id="3E621CC3FFA1983F002AAB3A712BF8AE" author="Gu, Y. - M." box="[378,526,1895,1919]" pageId="1" pageNumber="487" pagination="441 - 443" refId="ref14592" refString="Gu, Y. - M. (1990) A new species of the genus Hypoaspis (Acari: Laelapidae). Acta Zootaxonomica Sinica, 15, 441 - 443. [in Chinese]" type="journal article" year="1990">Gu, 1990: 441</bibRefCitation>
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.
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<paragraph id="5A4C6132FFA1983F01C7ABDB702CF86D" blockId="1.[151,1381,1895,2042]" pageId="1" pageNumber="487">
<treatmentCitationGroup id="7AE3461CFFA1983F01C7ABDB702CF86D" pageId="1" pageNumber="487">
<treatmentCitation id="DB524723FFA1983F01C7ABDB7022F84F" author="Xu, X. &amp; Liang, L." box="[151,775,1926,1950]" page="193" pageId="1" pageNumber="487" year="1996">
<taxonomicName id="9DF31AB1FFA1983F01C7ABDB7022F84F" authority="Xu &amp; Liang, 1996: 193" authorityName="Xu &amp; Liang" authorityPageNumber="193" authorityYear="1996" box="[151,775,1926,1950]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="487" phylum="Arthropoda" rank="species" species="hefeiensis" subGenus="Cosmolaelaps">
<emphasis id="6887BD20FFA1983F01C7ABDB7227F84C" box="[151,258,1926,1949]" italics="true" pageId="1" pageNumber="487">Hypoaspis</emphasis>
(
<emphasis id="6887BD20FFA1983F0041ABDB7284F84F" box="[273,417,1926,1950]" italics="true" pageId="1" pageNumber="487">Cosmolaelaps</emphasis>
)
<emphasis id="6887BD20FFA1983F00FFABDB7136F84F" box="[431,531,1926,1950]" italics="true" pageId="1" pageNumber="487">hefeiensis</emphasis>
<bibRefCitation id="3E621CC3FFA1983F0349ABDB7022F84F" author="Xu, X. &amp; Liang, L." box="[537,775,1926,1950]" pageId="1" pageNumber="487" pagination="189 - 197" refId="ref16841" refString="Xu, X. &amp; Liang, L. (1996) Four new species of the Hypoaspidinae (Acari: Laelapidae) from moss in China. Systematic and Applied Acarology, 1, 189 - 197. [in Chinese] https: // doi. org / 10.11158 / saa. 1.1.23" type="journal article" year="1996">Xu &amp; Liang, 1996: 193</bibRefCitation>
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(junior synonymy by
<treatmentCitation id="DB524723FFA1983F02B8ABDB7757F84F" author="Ma, L. - M." box="[1000,1138,1926,1950]" page="23" pageId="1" pageNumber="487" year="2006">
<bibRefCitation id="3E621CC3FFA1983F02B8ABDB7757F84F" author="Ma, L. - M." box="[1000,1138,1926,1950]" pageId="1" pageNumber="487" pagination="23 - 26" refId="ref15730" refString="Ma, L. - M. (2006) New synonyms of gamasid mites (Acari: Mesostigmata). Acta Arachnologica Sinica, 15, 23 - 26. [in Chinese]" type="journal article" year="2006">Ma, 2006: 23</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="DB524723FFA1983F052DABDB767CF84F" author="Bai, X. - L. &amp; Ma, L. - M." box="[1149,1369,1926,1950]" page="558" pageId="1" pageNumber="487" year="2012">
<bibRefCitation id="3E621CC3FFA1983F052DABDB767CF84F" author="Bai, X. - L. &amp; Ma, L. - M." box="[1149,1369,1926,1950]" pageId="1" pageNumber="487" pagination="555 - 558" refId="ref13456" refString="Bai, X. - L. &amp; Ma, L. - M. (2012) A new species of the genus Hypoaspis from Ningxia, China with supplementary description of Hypoaspis subpictus Gu et Bai (Acari: Laelapidae). Acta Zootaxonomica Sinica, 37, 555 - 558. [in Chinese]" type="journal article" year="2012">Bai &amp; Ma, 2012: 558</bibRefCitation>
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).
<taxonomicName id="9DF31AB1FFA1983F01C7ABF87136F86D" authorityName="Xu &amp; Liang" authorityYear="1996" box="[151,531,1956,1981]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="487" phylum="Arthropoda" rank="species" species="hefeiensis" subGenus="Cosmolaelaps">
<emphasis id="6887BD20FFA1983F01C7ABF87227F86D" box="[151,258,1957,1980]" italics="true" pageId="1" pageNumber="487">Hypoaspis</emphasis>
(
<emphasis id="6887BD20FFA1983F0041ABF97284F86D" box="[273,417,1956,1980]" italics="true" pageId="1" pageNumber="487">Cosmolaelaps</emphasis>
)
<emphasis id="6887BD20FFA1983F00FFABF97136F86D" box="[431,531,1956,1980]" italics="true" pageId="1" pageNumber="487">hefeiensis</emphasis>
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.—
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<bibRefCitation id="3E621CC3FFA1983F0362ABF97020F86D" author="Bei, N. - X. &amp; Shi, C. - M. &amp; Yin, S. - G." box="[562,773,1956,1981]" pageId="1" pageNumber="487" pagination="648 - 650" refId="ref13627" refString="Bei, N. - X., Shi, C. - M. &amp; Yin, S. - G. (2003) A new species of the genus Hypoaspis from China (Acari, Laelapidae). Acta Zootaxonomica Sinica, 28, 648 - 650. [in Chinese]" type="journal article" year="2003">
Bei
<emphasis id="6887BD20FFA1983F030CABF871ACF86D" box="[604,649,1956,1980]" italics="true" pageId="1" pageNumber="487">et al</emphasis>
., 2003: 648
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<paragraph id="5A4C6132FFA1983F01C7AB9971A7F80A" blockId="1.[151,1381,1895,2042]" box="[151,642,1987,2011]" pageId="1" pageNumber="487">
<treatmentCitationGroup id="7AE3461CFFA1983F01C7AB9971A7F80A" box="[151,642,1987,2011]" pageId="1" pageNumber="487">
<taxonomicName id="9DF31AB1FFA1983F01C7AB997256F80A" authorityName="Gu" authorityYear="1990" box="[151,371,1987,2011]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="487" phylum="Arthropoda" rank="species" species="chianensis">
<emphasis id="6887BD20FFA1983F01C7AB997256F80A" box="[151,371,1987,2011]" italics="true" pageId="1" pageNumber="487">Hypoaspis chianensis</emphasis>
</taxonomicName>
.—
<treatmentCitation id="DB524723FFA1983F00C1AB99715BF80A" author="Ren, T. - G. &amp; Guo, X. - G." box="[401,638,1987,2011]" page="329" pageId="1" pageNumber="487" year="2008">
<bibRefCitation id="3E621CC3FFA1983F00C1AB99715BF80A" author="Ren, T. - G. &amp; Guo, X. - G." box="[401,638,1987,2011]" pageId="1" pageNumber="487" pagination="326 - 332" refId="ref16327" refString="Ren, T. - G. &amp; Guo, X. - G. (2008) Preliminary study on Laelapidae fauna in China (Acari: Gamasidna: Laelapidae). Chinese Journal of Vector Biology and Control, 19 (4), 326 - 332." type="journal article" year="2008">Ren &amp; Guo, 2008: 329</bibRefCitation>
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.
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<paragraph id="5A4C6132FFA1983F01C7ABBF7096F82B" blockId="1.[151,1381,1895,2042]" box="[151,947,2018,2042]" pageId="1" pageNumber="487">
<treatmentCitationGroup id="7AE3461CFFA1983F01C7ABBF7096F82B" box="[151,947,2018,2042]" pageId="1" pageNumber="487">
<taxonomicName id="9DF31AB1FFA1983F01C7ABBF72BDF82B" authority="(Gu)" box="[151,408,2018,2042]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="487" phylum="Arthropoda" rank="species" species="chianensis">
<emphasis id="6887BD20FFA1983F01C7ABBF72BDF82B" box="[151,408,2018,2042]" italics="true" pageId="1" pageNumber="487">Cosmolaelaps chianensis</emphasis>
</taxonomicName>
.—
<treatmentCitation id="DB524723FFA1983F00E5ABBF719DF82B" author="Moreira, G. F. &amp; Klompen, H. &amp; de Moraes, G. J." box="[437,696,2018,2042]" page="319" pageId="1" pageNumber="487" year="2014">
<bibRefCitation id="3E621CC3FFA1983F00E5ABBF719DF82B" author="Moreira, G. F. &amp; Klompen, H. &amp; de Moraes, G. J." box="[437,696,2018,2042]" pageId="1" pageNumber="487" pagination="317 - 346" refId="ref15991" refString="Moreira, G. F., Klompen, H. &amp; de Moraes, G. J. (2014) Redefinition of Cosmolaelaps Berlese (Acari: Laelapidae) and description of five new species from Brazil. Zootaxa, 3764 (3), 317 - 346. https: // doi. org / 10.11646 / zootaxa. 3764.3.4" type="journal article" year="2014">
Moreira
<emphasis id="6887BD20FFA1983F0340ABBE7119F82B" box="[528,572,2018,2042]" italics="true" pageId="1" pageNumber="487">et al</emphasis>
., 2014: 319
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;
<treatmentCitation id="DB524723FFA1983F0392ABBF708AF82B" ID-DOI="http://doi.org/10.5281/zenodo.6003548" ID-Zenodo-Dep="6003548" author="Keum, E. &amp; Jung, C. &amp; Joharchi, O." box="[706,943,2018,2042]" httpUri="http://treatment.plazi.org/id/9C301B0A4804FF88D097FA88FB94FE4D" page="486" pageId="1" pageNumber="487" year="2017">
<bibRefCitation id="3E621CC3FFA1983F0392ABBF708AF82B" author="Keum, E. &amp; Jung, C. &amp; Joharchi, O." box="[706,943,2018,2042]" pageId="1" pageNumber="487" pagination="485 - 505" refId="ref15495" refString="Keum, E., Jung, C. &amp; Joharchi, O. (2017) New species and new records of the family Laelapidae (Acari: Mesostigmata) from Republic of Korea. Zootaxa, 4353 (3), 485 - 505. https: // doi. org / 10.11646 / zootaxa. 4353.3.5" type="journal article" year="2017">
Keum
<emphasis id="6887BD20FFA1983F0257ABBE7016F82B" box="[775,819,2018,2042]" italics="true" pageId="1" pageNumber="487">et al</emphasis>
., 2017: 486
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<paragraph id="5A4C6132FFA2983C01C7ABC57064F829" blockId="2.[151,1437,1944,2041]" pageId="2" pageNumber="488">
<emphasis id="6887BD20FFA2983C01C7ABC5721FF861" bold="true" box="[151,314,1944,1968]" pageId="2" pageNumber="488">FIGURES 1–7.</emphasis>
DIC micrographs of
<taxonomicName id="9DF31AB1FFA2983C0341ABC5706CF861" authority="(Gu)" baseAuthorityName="Gu" box="[529,841,1944,1968]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="488" phylum="Arthropoda" rank="species" species="chianensis">
<emphasis id="6887BD20FFA2983C0341ABC57034F861" box="[529,785,1944,1968]" italics="true" pageId="2" pageNumber="488">Cosmolaelaps chianensis</emphasis>
(Gu)
</taxonomicName>
, female. 1, idiosoma in dorsal view, enlarged section not to scale; 2, idiosoma in ventral view; 3, sternal shield; 4, general view of opisthogastric area; 5, chelicera; 6, dorsal aspect of femur, genu and tibia leg I; 7, ventral aspect of femur, genu and tibia leg I.
</paragraph>
</caption>
<paragraph id="5A4C6132FFA3983D01C7ACCB7624FF04" blockId="3.[151,1437,150,1041]" pageId="3" pageNumber="489">
<emphasis id="6887BD20FFA3983D01C7ACCB72A9FF61" bold="true" box="[151,396,150,176]" pageId="3" pageNumber="489">Specimens examined</emphasis>
.Ten females and
<specimenCount id="4CF5AABBFFA3983D031BACCA719BFF60" box="[587,702,151,177]" pageId="3" pageNumber="489" type="male">five males</specimenCount>
, National forest,
<collectingRegion id="9837AFD0FFA3983D0224ACCA70F3FF60" box="[884,982,151,177]" country="Russia" name="Sakhalin" pageId="3" pageNumber="489">Sakhalin</collectingRegion>
Island,
<collectingCountry id="22E421A2FFA3983D0579ACCA7753FF60" box="[1065,1142,151,177]" name="Russia" pageId="3" pageNumber="489">Russia</collectingCountry>
,
<geoCoordinate id="3FC707F5FFA3983D052FACCB7627FF60" box="[1151,1282,150,177]" degrees="46" direction="north" minutes="57" orientation="latitude" pageId="3" pageNumber="489" precision="15" seconds="39" value="46.960835">46°57'39&quot;N</geoCoordinate>
<geoCoordinate id="3FC707F5FFA3983D0456ACCB76B2FF60" box="[1286,1431,150,177]" degrees="142" direction="east" minutes="45" orientation="longitude" pageId="3" pageNumber="489" precision="15" seconds="28" value="142.75778">142°45'28&quot;E</geoCoordinate>
,
<date id="2E4D47F2FFA3983D01C7ACE6721BFF04" box="[151,318,187,213]" pageId="3" pageNumber="489" value="2021-08-09">9 August 2021</date>
, O. Joharchi coll., in the nest of
<taxonomicName id="9DF31AB1FFA3983D03F8ACE67011FF04" box="[680,820,187,213]" class="Insecta" family="Formicidae" genus="Myrmica" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="489" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="6887BD20FFA3983D03F8ACE67028FF05" box="[680,781,187,212]" italics="true" pageId="3" pageNumber="489">Myrmica</emphasis>
sp.
</taxonomicName>
(
<taxonomicName id="9DF31AB1FFA3983D0213ACE670C7FF04" ID-CoL="HYM" box="[835,994,187,213]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="489" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
:
<taxonomicName id="9DF31AB1FFA3983D02BBACE67748FF07" ID-CoL="623T5" box="[1003,1133,187,214]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="489" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
) (in TUMZ).
</paragraph>
</subSubSection>
<subSubSection id="12E932B9FFA3983D0197AC827626FBC0" pageId="3" pageNumber="489" type="discussion">
<paragraph id="5A4C6132FFA3983D0197AC827626FBC0" blockId="3.[151,1437,150,1041]" pageId="3" pageNumber="489">
<emphasis id="6887BD20FFA3983D0197AC827211FF28" bold="true" box="[199,308,223,249]" pageId="3" pageNumber="489">Remarks</emphasis>
.
<taxonomicName id="9DF31AB1FFA3983D0013AC827145FF28" box="[323,608,223,249]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="489" phylum="Arthropoda" rank="species" species="chianensis">
<emphasis id="6887BD20FFA3983D0013AC827145FF28" box="[323,608,223,249]" italics="true" pageId="3" pageNumber="489">Cosmolaelaps chianensis</emphasis>
</taxonomicName>
was described from
<collectingCountry id="22E421A2FFA3983D021DAC8270B5FF28" box="[845,912,223,249]" name="China" pageId="3" pageNumber="489">China</collectingCountry>
(
<bibRefCitation id="3E621CC3FFA3983D02F0AC82772BFF28" author="Gu, Y. - M." box="[928,1038,223,249]" pageId="3" pageNumber="489" pagination="441 - 443" refId="ref14592" refString="Gu, Y. - M. (1990) A new species of the genus Hypoaspis (Acari: Laelapidae). Acta Zootaxonomica Sinica, 15, 441 - 443. [in Chinese]" type="journal article" year="1990">Gu, 1990</bibRefCitation>
). It has been found associated with
<taxonomicName id="9DF31AB1FFA3983D01C7AD5E72A4FECC" authority="Thomas" authorityName="Thomas" authorityYear="1916" box="[151,385,259,285]" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" pageId="3" pageNumber="489" phylum="Chordata" rank="species" species="pahari">
<emphasis id="6887BD20FFA3983D01C7AD5E7238FECC" box="[151,285,259,285]" italics="true" pageId="3" pageNumber="489">Mus pahari</emphasis>
Thomas
</taxonomicName>
(
<taxonomicName id="9DF31AB1FFA3983D00C3AD5E72DBFECC" authorityName="Bowdich" authorityYear="1821" box="[403,510,259,285]" class="Mammalia" kingdom="Animalia" order="Rodentia" pageId="3" pageNumber="489" phylum="Chordata" rank="order">Rodentia</taxonomicName>
:
<taxonomicName id="9DF31AB1FFA3983D035DAD5E7154FECF" authorityName="Illiger" authorityYear="1811" box="[525,625,259,286]" class="Mammalia" family="Muridae" kingdom="Animalia" order="Rodentia" pageId="3" pageNumber="489" phylum="Chordata" rank="family">Muridae</taxonomicName>
). The description of this species is brief and both the description and illustrations lack many important details.
<bibRefCitation id="3E621CC3FFA3983D0332AD7A71FDFE93" author="Ma, L. - M." box="[610,728,295,322]" pageId="3" pageNumber="489" pagination="23 - 26" refId="ref15730" refString="Ma, L. - M. (2006) New synonyms of gamasid mites (Acari: Mesostigmata). Acta Arachnologica Sinica, 15, 23 - 26. [in Chinese]" type="journal article" year="2006">Ma (2006)</bibRefCitation>
considered that
<taxonomicName id="9DF31AB1FFA3983D02DCAD7A77BAFE90" box="[908,1183,295,321]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="489" phylum="Arthropoda" rank="species" species="hefeiensis">
<emphasis id="6887BD20FFA3983D02DCAD7A77BAFE90" box="[908,1183,295,321]" italics="true" pageId="3" pageNumber="489">Cosmolaelaps hefeiensis</emphasis>
</taxonomicName>
is a junior synonym of
<taxonomicName id="9DF31AB1FFA3983D01C7AD16720BFEB4" box="[151,302,331,357]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="489" phylum="Arthropoda" rank="species" species="chianensis">
<emphasis id="6887BD20FFA3983D01C7AD16738FFEB4" box="[151,170,331,357]" italics="true" pageId="3" pageNumber="489">C</emphasis>
.
<emphasis id="6887BD20FFA3983D01E8AD16720BFEB4" box="[184,302,331,357]" italics="true" pageId="3" pageNumber="489">chianensis</emphasis>
</taxonomicName>
. However,
<bibRefCitation id="3E621CC3FFA3983D00FFAD167100FEB7" author="Ma, L. - M." box="[431,549,331,358]" pageId="3" pageNumber="489" pagination="23 - 26" refId="ref15730" refString="Ma, L. - M. (2006) New synonyms of gamasid mites (Acari: Mesostigmata). Acta Arachnologica Sinica, 15, 23 - 26. [in Chinese]" type="journal article" year="2006">Ma (2006)</bibRefCitation>
did not provide any explanation for this decision, nor did he provide the details of the examined specimens.
<taxonomicName id="9DF31AB1FFA3983D008DAD3271D6FE58" box="[477,755,367,393]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="489" phylum="Arthropoda" rank="species" species="hefeiensis">
<emphasis id="6887BD20FFA3983D008DAD3271D6FE58" box="[477,755,367,393]" italics="true" pageId="3" pageNumber="489">Cosmolaelaps hefeiensis</emphasis>
</taxonomicName>
was also described from
<collectingCountry id="22E421A2FFA3983D054BAD32777BFE58" box="[1051,1118,367,393]" name="China" pageId="3" pageNumber="489">China</collectingCountry>
(
<bibRefCitation id="3E621CC3FFA3983D053EAD327662FE5B" author="Xu, X. &amp; Liang, L." box="[1134,1351,367,394]" pageId="3" pageNumber="489" pagination="189 - 197" refId="ref16841" refString="Xu, X. &amp; Liang, L. (1996) Four new species of the Hypoaspidinae (Acari: Laelapidae) from moss in China. Systematic and Applied Acarology, 1, 189 - 197. [in Chinese] https: // doi. org / 10.11158 / saa. 1.1.23" type="journal article" year="1996">Xu &amp; Liang, 1996</bibRefCitation>
). It has been found from moss (
<bibRefCitation id="3E621CC3FFA3983D00F3ADCE7151FE7F" author="Xu, X. &amp; Liang, L." box="[419,628,403,430]" pageId="3" pageNumber="489" pagination="189 - 197" refId="ref16841" refString="Xu, X. &amp; Liang, L. (1996) Four new species of the Hypoaspidinae (Acari: Laelapidae) from moss in China. Systematic and Applied Acarology, 1, 189 - 197. [in Chinese] https: // doi. org / 10.11158 / saa. 1.1.23" type="journal article" year="1996">Xu &amp; Liang, 1996</bibRefCitation>
) and this species also has been recorded from soil in
<collectingCountry id="22E421A2FFA3983D059DADCE76B9FE7C" box="[1229,1436,403,429]" name="South Korea" pageId="3" pageNumber="489">Republic of Korea</collectingCountry>
(
<bibRefCitation id="3E621CC3FFA3983D01CFADEA724CFE00" author="Keum, E. &amp; Jung, C. &amp; Joharchi, O." box="[159,361,439,465]" pageId="3" pageNumber="489" pagination="485 - 505" refId="ref15495" refString="Keum, E., Jung, C. &amp; Joharchi, O. (2017) New species and new records of the family Laelapidae (Acari: Mesostigmata) from Republic of Korea. Zootaxa, 4353 (3), 485 - 505. https: // doi. org / 10.11646 / zootaxa. 4353.3.5" type="journal article" year="2017">
Keum
<emphasis id="6887BD20FFA3983D01BBADEA723EFE00" box="[235,283,439,465]" italics="true" pageId="3" pageNumber="489">et al</emphasis>
., 2017
</bibRefCitation>
) and
<collectingCountry id="22E421A2FFA3983D00F5ADEA72D5FE00" box="[421,496,439,465]" name="Russia" pageId="3" pageNumber="489">Russia</collectingCountry>
(
<bibRefCitation id="3E621CC3FFA3983D00AEADEA71ECFE00" author="Marchenko, I. I." box="[510,713,439,465]" pageId="3" pageNumber="489" pagination="151 - 157" refId="ref15880" refString="Marchenko, I. I. (2017) Soil gamasid mites (Acari, Mesostigmata) of Sokhondinskii Nature Reserve, Zabaikalskii Krai, Russia. Euroasian Entomological Journal, 16, 151 - 157. [in Russian]" type="journal article" year="2017">Marchenko, 2017</bibRefCitation>
). By comparing the descriptions and figures of these two species, we found some distinguishing morphological differences: (1) dorsal shield setae
<emphasis id="6887BD20FFA3983D0577AD867767FE24" box="[1063,1090,475,501]" italics="true" pageId="3" pageNumber="489">J5</emphasis>
and
<emphasis id="6887BD20FFA3983D052AAD8677B2FE24" box="[1146,1175,475,501]" italics="true" pageId="3" pageNumber="489">Z5</emphasis>
more or less similar in length, and
<emphasis id="6887BD20FFA3983D0049ADA2720BFDC8" box="[281,302,511,537]" italics="true" pageId="3" pageNumber="489">j1</emphasis>
considerably shorter than
<emphasis id="6887BD20FFA3983D0304ADA2714CFDC8" box="[596,617,511,537]" italics="true" pageId="3" pageNumber="489">
j
<quantity id="9D0BCCD7FFA3983D030EADA2714CFDC8" box="[606,617,511,537]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="3" pageNumber="489" unit="in" value="2.0">2</quantity>
</emphasis>
in
<taxonomicName id="9DF31AB1FFA3983D03DBADA2703CFDC8" box="[651,793,511,537]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="489" phylum="Arthropoda" rank="species" species="hefeiensis">
<emphasis id="6887BD20FFA3983D03DBADA271BBFDC8" box="[651,670,511,537]" italics="true" pageId="3" pageNumber="489">C</emphasis>
.
<emphasis id="6887BD20FFA3983D03FBADA2703CFDC8" box="[683,793,511,537]" italics="true" pageId="3" pageNumber="489">hefeiensis</emphasis>
</taxonomicName>
(see
<bibRefCitation id="3E621CC3FFA3983D0203ADA27738FDC8" author="Xu, X. &amp; Liang, L." box="[851,1053,511,537]" pageId="3" pageNumber="489" pagination="189 - 197" refId="ref16841" refString="Xu, X. &amp; Liang, L. (1996) Four new species of the Hypoaspidinae (Acari: Laelapidae) from moss in China. Systematic and Applied Acarology, 1, 189 - 197. [in Chinese] https: // doi. org / 10.11158 / saa. 1.1.23" type="journal article" year="1996">Xu &amp; Liang, 1996</bibRefCitation>
and
<figureCitation id="C2C87DB7FFA3983D0502ADA277B7FDC8" box="[1106,1170,511,537]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1944,1968]" captionTargetBox="[182,1405,181,1920]" captionTargetId="figure-18@2.[182,1405,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–7. DIC micrographs of Cosmolaelaps chianensis (Gu), female. 1, idiosoma in dorsal view, enlarged section not to scale; 2, idiosoma in ventral view; 3, sternal shield; 4, general view of opisthogastric area; 5, chelicera; 6, dorsal aspect of femur, genu and tibia leg I; 7, ventral aspect of femur, genu and tibia leg I." figureDoi="http://doi.org/10.5281/zenodo.6530960" httpUri="https://zenodo.org/record/6530960/files/figure.png" pageId="3" pageNumber="489">Fig. 1</figureCitation>
of current study), while in
<taxonomicName id="9DF31AB1FFA3983D01E3AE7E726CFDEC" box="[179,329,547,573]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="489" phylum="Arthropoda" rank="species" species="chianensis">
<emphasis id="6887BD20FFA3983D01E3AE7E73E3FDEC" box="[179,198,547,573]" italics="true" pageId="3" pageNumber="489">C</emphasis>
.
<emphasis id="6887BD20FFA3983D0183AE7E726CFDEC" box="[211,329,547,573]" italics="true" pageId="3" pageNumber="489">chianensis</emphasis>
</taxonomicName>
setae
<emphasis id="6887BD20FFA3983D00DDAE7E728FFDEC" box="[397,426,547,573]" italics="true" pageId="3" pageNumber="489">Z5</emphasis>
are obviously longer than
<emphasis id="6887BD20FFA3983D0380AE7E71CEFDEC" box="[720,747,547,573]" italics="true" pageId="3" pageNumber="489">J5</emphasis>
and
<emphasis id="6887BD20FFA3983D024FAE7E7011FDEC" box="[799,820,547,573]" italics="true" pageId="3" pageNumber="489">j1</emphasis>
is similar length to
<emphasis id="6887BD20FFA3983D055FAE7E7701FDEC" box="[1039,1060,547,573]" italics="true" pageId="3" pageNumber="489">j2</emphasis>
(see
<figureCitation id="C2C87DB7FFA3983D050DAE7E77BBFDEC" box="[1117,1182,547,573]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1944,1968]" captionTargetBox="[182,1405,181,1920]" captionTargetId="figure-18@2.[182,1405,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–7. DIC micrographs of Cosmolaelaps chianensis (Gu), female. 1, idiosoma in dorsal view, enlarged section not to scale; 2, idiosoma in ventral view; 3, sternal shield; 4, general view of opisthogastric area; 5, chelicera; 6, dorsal aspect of femur, genu and tibia leg I; 7, ventral aspect of femur, genu and tibia leg I." figureDoi="http://doi.org/10.5281/zenodo.6530960" httpUri="https://zenodo.org/record/6530960/files/figure.png" pageId="3" pageNumber="489">
Fig.
<quantity id="9D0BCCD7FFA3983D05C0AE7E77BBFDEC" box="[1168,1182,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="489" unit="in" value="1.0">1</quantity>
</figureCitation>
in
<bibRefCitation id="3E621CC3FFA3983D05EFAE7E760EFDEC" author="Gu, Y. - M." box="[1215,1323,547,573]" pageId="3" pageNumber="489" pagination="441 - 443" refId="ref14592" refString="Gu, Y. - M. (1990) A new species of the genus Hypoaspis (Acari: Laelapidae). Acta Zootaxonomica Sinica, 15, 441 - 443. [in Chinese]" type="journal article" year="1990">Gu, 1990</bibRefCitation>
); (2) body size 395 long, 255 wide in
<taxonomicName id="9DF31AB1FFA3983D009FAE1A7142FDB0" box="[463,615,583,609]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="489" phylum="Arthropoda" rank="species" species="chianensis">
<emphasis id="6887BD20FFA3983D009FAE1A72C7FDB0" box="[463,482,583,609]" italics="true" pageId="3" pageNumber="489">C</emphasis>
.
<emphasis id="6887BD20FFA3983D00A1AE1A7142FDB0" box="[497,615,583,609]" italics="true" pageId="3" pageNumber="489">chianensis</emphasis>
</taxonomicName>
(see
<bibRefCitation id="3E621CC3FFA3983D03F5AE1A7036FDB0" author="Gu, Y. - M." box="[677,787,583,609]" pageId="3" pageNumber="489" pagination="441 - 443" refId="ref14592" refString="Gu, Y. - M. (1990) A new species of the genus Hypoaspis (Acari: Laelapidae). Acta Zootaxonomica Sinica, 15, 441 - 443. [in Chinese]" type="journal article" year="1990">Gu, 1990</bibRefCitation>
), while in
<taxonomicName id="9DF31AB1FFA3983D02DDAE1A7738FDB0" box="[909,1053,583,609]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="489" phylum="Arthropoda" rank="species" species="hefeiensis">
<emphasis id="6887BD20FFA3983D02DDAE1A7085FDB0" box="[909,928,583,609]" italics="true" pageId="3" pageNumber="489">C</emphasis>
.
<emphasis id="6887BD20FFA3983D02FFAE1A7738FDB0" box="[943,1053,583,609]" italics="true" pageId="3" pageNumber="489">hefeiensis</emphasis>
</taxonomicName>
size of body larger, especially its width (484–560 long, 345–391 wide) (see
<bibRefCitation id="3E621CC3FFA3983D0324AE367060FD57" author="Xu, X. &amp; Liang, L." box="[628,837,619,646]" pageId="3" pageNumber="489" pagination="189 - 197" refId="ref16841" refString="Xu, X. &amp; Liang, L. (1996) Four new species of the Hypoaspidinae (Acari: Laelapidae) from moss in China. Systematic and Applied Acarology, 1, 189 - 197. [in Chinese] https: // doi. org / 10.11158 / saa. 1.1.23" type="journal article" year="1996">Xu &amp; Liang, 1996</bibRefCitation>
). Our efforts to see
<typeStatus id="8548DF90FFA3983D0572AE367777FD54" box="[1058,1106,619,645]" pageId="3" pageNumber="489">type</typeStatus>
material of these two species were not successful so, in this study we follow
<bibRefCitation id="3E621CC3FFA3983D03F9AED27017FD7B" author="Ma, L. - M." box="[681,818,655,682]" pageId="3" pageNumber="489" pagination="23 - 26" refId="ref15730" refString="Ma, L. - M. (2006) New synonyms of gamasid mites (Acari: Mesostigmata). Acta Arachnologica Sinica, 15, 23 - 26. [in Chinese]" type="journal article" year="2006">Ma’s (2006)</bibRefCitation>
treatment and provisionally retain these two species as synonyms until further comparative studies clarify their relationship. Our concept of the species is based on that of
<bibRefCitation id="3E621CC3FFA3983D01E6AE8A72B5FD23" author="Xu, X. &amp; Liang, L." box="[182,400,727,754]" pageId="3" pageNumber="489" pagination="189 - 197" refId="ref16841" refString="Xu, X. &amp; Liang, L. (1996) Four new species of the Hypoaspidinae (Acari: Laelapidae) from moss in China. Systematic and Applied Acarology, 1, 189 - 197. [in Chinese] https: // doi. org / 10.11158 / saa. 1.1.23" type="journal article" year="1996">Xu &amp; Liang (1996)</bibRefCitation>
and the diagnosis given as follows is based primarily on specimens from
<collectingCountry id="22E421A2FFA3983D059EAE8A763EFD20" box="[1230,1307,727,753]" name="Russia" pageId="3" pageNumber="489">Russia</collectingCountry>
, but also in comparison with the original description of
<taxonomicName id="9DF31AB1FFA3983D032FAEA67028FCC4" box="[639,781,763,789]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="489" phylum="Arthropoda" rank="species" species="hefeiensis">
<emphasis id="6887BD20FFA3983D032FAEA671B7FCC4" box="[639,658,763,789]" italics="true" pageId="3" pageNumber="489">C</emphasis>
.
<emphasis id="6887BD20FFA3983D03CFAEA67028FCC4" box="[671,781,763,789]" italics="true" pageId="3" pageNumber="489">hefeiensis</emphasis>
</taxonomicName>
. Our specimens agree very well with the description given by
<bibRefCitation id="3E621CC3FFA3983D01EBAF4272AFFCE8" author="Xu, X. &amp; Liang, L." box="[187,394,799,825]" pageId="3" pageNumber="489" pagination="189 - 197" refId="ref16841" refString="Xu, X. &amp; Liang, L. (1996) Four new species of the Hypoaspidinae (Acari: Laelapidae) from moss in China. Systematic and Applied Acarology, 1, 189 - 197. [in Chinese] https: // doi. org / 10.11158 / saa. 1.1.23" type="journal article" year="1996">Xu &amp; Liang, 1996</bibRefCitation>
for
<taxonomicName id="9DF31AB1FFA3983D00EBAF4271EAFCE8" box="[443,719,799,825]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="489" phylum="Arthropoda" rank="species" species="hefeiensis">
<emphasis id="6887BD20FFA3983D00EBAF4271EAFCE8" box="[443,719,799,825]" italics="true" pageId="3" pageNumber="489">Cosmolaelaps hefeiensis</emphasis>
</taxonomicName>
. The species is easily recognised by the long and thick setae of the dorsal shield (most setae long enough to reach well past the base of next posterior seta) (
<figureCitation id="C2C87DB7FFA3983D05DEAF1E77EBFC8C" box="[1166,1230,835,861]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1944,1968]" captionTargetBox="[182,1405,181,1920]" captionTargetId="figure-18@2.[182,1405,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–7. DIC micrographs of Cosmolaelaps chianensis (Gu), female. 1, idiosoma in dorsal view, enlarged section not to scale; 2, idiosoma in ventral view; 3, sternal shield; 4, general view of opisthogastric area; 5, chelicera; 6, dorsal aspect of femur, genu and tibia leg I; 7, ventral aspect of femur, genu and tibia leg I." figureDoi="http://doi.org/10.5281/zenodo.6530960" httpUri="https://zenodo.org/record/6530960/files/figure.png" pageId="3" pageNumber="489">Fig. 1</figureCitation>
),
<emphasis id="6887BD20FFA3983D05B5AF1E77DEFC8C" box="[1253,1275,835,861]" italics="true" pageId="3" pageNumber="489">j1</emphasis>
,
<emphasis id="6887BD20FFA3983D0458AF1E7604FC8C" box="[1288,1313,835,861]" italics="true" pageId="3" pageNumber="489">z1</emphasis>
,
<emphasis id="6887BD20FFA3983D047EAF1E7669FC8C" box="[1326,1356,835,861]" italics="true" pageId="3" pageNumber="489">Z5</emphasis>
simple and without knob at their bases,
<emphasis id="6887BD20FFA3983D00ABAF3A7135FC50" box="[507,528,871,897]" italics="true" pageId="3" pageNumber="489">j1</emphasis>
apically bent (hook shaped) and
<emphasis id="6887BD20FFA3983D022DAF3A70BFFC50" box="[893,922,871,897]" italics="true" pageId="3" pageNumber="489">Z5</emphasis>
shorter than
<emphasis id="6887BD20FFA3983D0579AF3A7761FC50" box="[1065,1092,871,897]" italics="true" pageId="3" pageNumber="489">J5</emphasis>
(
<emphasis id="6887BD20FFA3983D0503AF3A7748FC50" box="[1107,1133,871,897]" italics="true" pageId="3" pageNumber="489">J5</emphasis>
≈ 1.5 x
<emphasis id="6887BD20FFA3983D0595AF3A77C7FC50" box="[1221,1250,871,897]" italics="true" pageId="3" pageNumber="489">Z5</emphasis>
) (
<figureCitation id="C2C87DB7FFA3983D05AAAF3A761FFC50" box="[1274,1338,871,897]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1944,1968]" captionTargetBox="[182,1405,181,1920]" captionTargetId="figure-18@2.[182,1405,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–7. DIC micrographs of Cosmolaelaps chianensis (Gu), female. 1, idiosoma in dorsal view, enlarged section not to scale; 2, idiosoma in ventral view; 3, sternal shield; 4, general view of opisthogastric area; 5, chelicera; 6, dorsal aspect of femur, genu and tibia leg I; 7, ventral aspect of femur, genu and tibia leg I." figureDoi="http://doi.org/10.5281/zenodo.6530960" httpUri="https://zenodo.org/record/6530960/files/figure.png" pageId="3" pageNumber="489">Fig. 1</figureCitation>
); sternal setae short, at most reaching base of next setae (
<figureCitation id="C2C87DB7FFA3983D03E4AFD6703DFC74" box="[692,792,907,933]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1944,1968]" captionTargetBox="[182,1405,181,1920]" captionTargetId="figure-18@2.[182,1405,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–7. DIC micrographs of Cosmolaelaps chianensis (Gu), female. 1, idiosoma in dorsal view, enlarged section not to scale; 2, idiosoma in ventral view; 3, sternal shield; 4, general view of opisthogastric area; 5, chelicera; 6, dorsal aspect of femur, genu and tibia leg I; 7, ventral aspect of femur, genu and tibia leg I." figureDoi="http://doi.org/10.5281/zenodo.6530960" httpUri="https://zenodo.org/record/6530960/files/figure.png" pageId="3" pageNumber="489">Figs 2, 3</figureCitation>
), soft opisthogastric cuticle bearing 16 pairs setae,
<emphasis id="6887BD20FFA3983D0409AFD676B9FC74" box="[1369,1436,907,933]" italics="true" pageId="3" pageNumber="489">Jv4–5</emphasis>
and
<emphasis id="6887BD20FFA3983D0199AFF2722BFC18" box="[201,270,943,969]" italics="true" pageId="3" pageNumber="489">Zv4–5</emphasis>
thickened (
<figureCitation id="C2C87DB7FFA3983D00C5AFF272DBFC18" box="[405,510,943,969]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1944,1968]" captionTargetBox="[182,1405,181,1920]" captionTargetId="figure-18@2.[182,1405,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–7. DIC micrographs of Cosmolaelaps chianensis (Gu), female. 1, idiosoma in dorsal view, enlarged section not to scale; 2, idiosoma in ventral view; 3, sternal shield; 4, general view of opisthogastric area; 5, chelicera; 6, dorsal aspect of femur, genu and tibia leg I; 7, ventral aspect of femur, genu and tibia leg I." figureDoi="http://doi.org/10.5281/zenodo.6530960" httpUri="https://zenodo.org/record/6530960/files/figure.png" pageId="3" pageNumber="489">Figs 2, 4</figureCitation>
), post-anal seta slightly thicker and longer than para-anal setae (
<figureCitation id="C2C87DB7FFA3983D05B3AFF2766EFC18" box="[1251,1355,943,969]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1944,1968]" captionTargetBox="[182,1405,181,1920]" captionTargetId="figure-18@2.[182,1405,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–7. DIC micrographs of Cosmolaelaps chianensis (Gu), female. 1, idiosoma in dorsal view, enlarged section not to scale; 2, idiosoma in ventral view; 3, sternal shield; 4, general view of opisthogastric area; 5, chelicera; 6, dorsal aspect of femur, genu and tibia leg I; 7, ventral aspect of femur, genu and tibia leg I." figureDoi="http://doi.org/10.5281/zenodo.6530960" httpUri="https://zenodo.org/record/6530960/files/figure.png" pageId="3" pageNumber="489">Figs 2, 4</figureCitation>
), fixed digit of chelicera of female with six teeth, including three large proximal teeth (posterior to pilus dentilis) (
<figureCitation id="C2C87DB7FFA3983D0419AF8E76A8FC3C" box="[1353,1421,978,1005]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1944,1968]" captionTargetBox="[182,1405,181,1920]" captionTargetId="figure-18@2.[182,1405,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–7. DIC micrographs of Cosmolaelaps chianensis (Gu), female. 1, idiosoma in dorsal view, enlarged section not to scale; 2, idiosoma in ventral view; 3, sternal shield; 4, general view of opisthogastric area; 5, chelicera; 6, dorsal aspect of femur, genu and tibia leg I; 7, ventral aspect of femur, genu and tibia leg I." figureDoi="http://doi.org/10.5281/zenodo.6530960" httpUri="https://zenodo.org/record/6530960/files/figure.png" pageId="3" pageNumber="489">Fig. 5</figureCitation>
), spatulate seta on legs absent, genu and tibia of leg I without conspicuously thickened seta (
<figureCitation id="C2C87DB7FFA3983D05C0AFAA77D0FBC0" box="[1168,1269,1015,1041]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1944,1968]" captionTargetBox="[182,1405,181,1920]" captionTargetId="figure-18@2.[182,1405,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–7. DIC micrographs of Cosmolaelaps chianensis (Gu), female. 1, idiosoma in dorsal view, enlarged section not to scale; 2, idiosoma in ventral view; 3, sternal shield; 4, general view of opisthogastric area; 5, chelicera; 6, dorsal aspect of femur, genu and tibia leg I; 7, ventral aspect of femur, genu and tibia leg I." figureDoi="http://doi.org/10.5281/zenodo.6530960" httpUri="https://zenodo.org/record/6530960/files/figure.png" pageId="3" pageNumber="489">Figs 6, 7</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

129
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Genus
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<emphasis id="6887BD20FFA1983F01BEA99772B3FA34" bold="true" box="[238,406,1482,1509]" italics="true" pageId="1" pageNumber="487">Cosmolaelaps</emphasis>
Berlese
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<subSubSection id="12E932B9FFA1983F01C7AA52712CF997" pageId="1" pageNumber="487" type="reference_group">
<paragraph id="5A4C6132FFA1983F01C7AA527168F9F6" blockId="1.[151,883,1551,1637]" box="[151,589,1551,1575]" pageId="1" pageNumber="487">
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<emphasis id="6887BD20FFA1983F01C7AA5273C2F9F6" box="[151,231,1551,1575]" italics="true" pageId="1" pageNumber="487">Laelaps</emphasis>
(
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)
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.
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<taxonomicName id="9DF31AB1FFA1983F01C7AA737202F997" authorityName="Berlese" authorityYear="1903" box="[151,295,1582,1606]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="487" phylum="Arthropoda" rank="genus">
<emphasis id="6887BD20FFA1983F01C7AA737202F997" box="[151,295,1582,1606]" italics="true" pageId="1" pageNumber="487">Cosmolaelaps</emphasis>
</taxonomicName>
.
<emphasis id="6887BD20FFA1983F007DAA727261F997" box="[301,324,1583,1606]" italics="true" pageId="1" pageNumber="487">—</emphasis>
<treatmentCitation id="DB524723FFA1983F0014AA737120F997" author="Berlese, A." box="[324,517,1582,1606]" page="157" pageId="1" pageNumber="487" year="1920">
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.
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<paragraph id="5A4C6132FFA1983F01C7AA107056F9B5" blockId="1.[151,883,1551,1637]" box="[151,883,1612,1637]" pageId="1" pageNumber="487">
<typeStatus id="8548DF90FFA1983F01C7AA1073ECF9B4" box="[151,201,1613,1637]" pageId="1" pageNumber="487">Type</typeStatus>
species
<taxonomicName id="9DF31AB1FFA1983F004FAA117152F9B4" authority="Berlese, 1883: 2" authorityName="Berlese" authorityPageNumber="2" authorityYear="1883" box="[287,631,1612,1637]" class="Arachnida" family="Laelapidae" genus="Laelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="487" phylum="Arthropoda" rank="species" species="claviger">
<emphasis id="6887BD20FFA1983F004FAA1172ECF9B5" box="[287,457,1612,1636]" italics="true" pageId="1" pageNumber="487">Laelaps claviger</emphasis>
<bibRefCitation id="3E621CC3FFA1983F009FAA117152F9B4" author="Berlese, A." box="[463,631,1612,1637]" pageId="1" pageNumber="487" pagination="2" refId="ref13682" refString="Berlese, A. (1883) s. n. Acari, Myriapoda et Scorpiones Hucusque in Italia Reperta, 4, 2. https: // doi. org / 10.5962 / bhl. title. 69269" type="journal article" year="1883">Berlese, 1883: 2</bibRefCitation>
</taxonomicName>
; by original designation.
</paragraph>
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<paragraph id="5A4C6132FFA1983F01C7AACE7793F97C" blockId="1.[151,1206,1683,1709]" box="[151,1206,1683,1709]" pageId="1" pageNumber="487">
<emphasis id="6887BD20FFA1983F01C7AACE722FF97C" bold="true" box="[151,266,1683,1709]" italics="true" pageId="1" pageNumber="487">Diagnosis</emphasis>
. The diagnosis of
<taxonomicName id="9DF31AB1FFA1983F0089AACE715DF97C" authorityName="Berlese" authorityYear="1903" box="[473,632,1683,1709]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="487" phylum="Arthropoda" rank="genus">
<emphasis id="6887BD20FFA1983F0089AACE715DF97C" box="[473,632,1683,1709]" italics="true" pageId="1" pageNumber="487">Cosmolaelaps</emphasis>
</taxonomicName>
used here is based on that of
<bibRefCitation id="3E621CC3FFA1983F0293AACE7797F97C" author="Moreira, G. F. &amp; Klompen, H. &amp; de Moraes, G. J." box="[963,1202,1683,1709]" pageId="1" pageNumber="487" pagination="317 - 346" refId="ref15991" refString="Moreira, G. F., Klompen, H. &amp; de Moraes, G. J. (2014) Redefinition of Cosmolaelaps Berlese (Acari: Laelapidae) and description of five new species from Brazil. Zootaxa, 3764 (3), 317 - 346. https: // doi. org / 10.11646 / zootaxa. 3764.3.4" type="journal article" year="2014">
Moreira
<emphasis id="6887BD20FFA1983F0575AACE7773F97C" box="[1061,1110,1683,1709]" italics="true" pageId="1" pageNumber="487">et al</emphasis>
. (2014)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="CFF54445B5B95D5F33BB59EB57392D57">New species and new records of Cosmolaelaps Berlese (Acari: Laelapidae) from Russia, with a review of the Russian species of the genus</mods:title>
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<mods:namePart id="E71A064C8BE9C20CE99DE1C93C5BC37D">Döker, Ismail</mods:namePart>
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<mods:affiliation id="A48F3551AE81BBD549E8029ECC6D1273">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Semakova Str. 10, 625003 Tyumen, Russia. &amp; Cukurova University, Agricultural Faculty, Department of Plant Protection, Acarology Laboratory, 01330, Adana, Turkey. https: // orcid. org / 0000 - 0002 - 2741 - 4946 (Joharchi) https: // orcid. org / 0000 - 0002 - 1412 - 1554 (Döker) https: // orcid. org / 0000 - 0003 - 2831 - 7213 (Khaustov)</mods:affiliation>
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<subSubSection id="12E932B9FFA3983D01C7A83F7180FBAC" box="[151,677,1122,1149]" pageId="3" pageNumber="489" type="nomenclature">
<paragraph id="5A4C6132FFA3983D01C7A83F7180FBAC" blockId="3.[151,677,1122,1185]" box="[151,677,1122,1149]" pageId="3" pageNumber="489">
<heading id="0104D65EFFA3983D01C7A83F7180FBAC" bold="true" box="[151,677,1122,1149]" fontSize="11" level="1" pageId="3" pageNumber="489" reason="1">
<emphasis id="6887BD20FFA3983D01C7A83F7180FBAC" bold="true" box="[151,677,1122,1149]" pageId="3" pageNumber="489">
<taxonomicName id="9DF31AB1FFA3983D01C7A83F7165FBAD" authority="Joharchi" authorityName="Joharchi" authorityYear="2022" box="[151,576,1122,1149]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="489" phylum="Arthropoda" rank="species" species="lasiophilus" status="sp. nov.">
<emphasis id="6887BD20FFA3983D01C7A83F72EFFBAC" bold="true" box="[151,458,1122,1149]" italics="true" pageId="3" pageNumber="489">Cosmolaelaps lasiophilus</emphasis>
Joharchi
</taxonomicName>
<taxonomicNameLabel id="73B4005BFFA3983D0317A83E7180FBAC" box="[583,677,1123,1149]" pageId="3" pageNumber="489" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="12E932B9FFA3983D01C7A8DA7218FB70" box="[151,317,1159,1185]" pageId="3" pageNumber="489" type="description">
<paragraph id="5A4C6132FFA3983D01C7A8DA7218FB70" blockId="3.[151,677,1122,1185]" box="[151,317,1159,1185]" pageId="3" pageNumber="489">
(
<figureCitation id="C2C87DB7FFA3983D01CFA8DA7211FB70" box="[159,308,1159,1185]" captionStart-0="FIGURES 8–12" captionStart-1="FIGURES 13–20" captionStart-2="FIGURE 21–24" captionStartId-0="4.[151,264,1943,1967]" captionStartId-1="5.[151,264,1908,1932]" captionStartId-2="6.[151,250,1926,1950]" captionTargetBox-0="[181,1397,196,1901]" captionTargetBox-1="[198,1388,181,1884]" captionTargetBox-2="[161,1425,197,1887]" captionTargetId-0="figure-18@4.[167,1419,181,1919]" captionTargetId-1="figure-19@5.[198,1389,181,1884]" captionTargetId-2="figure-18@6.[152,1436,181,1902]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." captionText-1="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." captionText-2="FIGURE 21–24. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 21, leg I (trochanter-tibia); 22, leg II; 23, leg III; 24, leg IV." figureDoi-0="http://doi.org/10.5281/zenodo.6530964" figureDoi-1="http://doi.org/10.5281/zenodo.6530968" figureDoi-2="http://doi.org/10.5281/zenodo.6530972" httpUri-0="https://zenodo.org/record/6530964/files/figure.png" httpUri-1="https://zenodo.org/record/6530968/files/figure.png" httpUri-2="https://zenodo.org/record/6530972/files/figure.png" pageId="3" pageNumber="489">Figures 8–24</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="12E932B9FFA3983D01C7A892700BFAE0" pageId="3" pageNumber="489" type="materials_examined">
<paragraph id="5A4C6132FFA3983D01C7A892700BFAE0" blockId="3.[151,1437,1230,2013]" pageId="3" pageNumber="489">
<emphasis id="6887BD20FFA3983D01C7A8927263FB39" bold="true" box="[151,326,1230,1257]" pageId="3" pageNumber="489">Type material.</emphasis>
<materialsCitation id="EA9B6B6FFFA3983D001DA89276B8FADC" ID-GBIF-Occurrence="3763889304" collectedFrom="in the nest of Lasius niger (L.) (Hymenoptera: Formicidae)" collectingDate="2021-10-18" collectionCode="TUMZ" collectorName="Alexander A. Khaustov" country="Russia" latitude="57.066666" location="Uspenka" longLatPrecision="1053" longitude="65.066666" municipality="Uspenka" pageId="3" pageNumber="489" specimenCount="1" specimenCount-female="1" stateProvince="Tyumen Province" typeStatus="holotype">
<typeStatus id="8548DF90FFA3983D001DA892729DFB38" box="[333,440,1231,1257]" pageId="3" pageNumber="489" type="holotype">Holotype</typeStatus>
,
<specimenCount id="4CF5AABBFFA3983D0094A8927136FB38" box="[452,531,1231,1257]" pageId="3" pageNumber="489" type="female">female</specimenCount>
, vicinities of
<collectingMunicipality id="BA28FB48FFA3983D03FCA8927034FB38" box="[684,785,1231,1257]" pageId="3" pageNumber="489">Uspenka</collectingMunicipality>
,
<collectingRegion id="9837AFD0FFA3983D024DA89270C3FB38" box="[797,998,1231,1257]" country="Russia" name="Tyumen'" pageId="3" pageNumber="489">Tyumen Province</collectingRegion>
,
<collectingCountry id="22E421A2FFA3983D02A2A892771AFB38" box="[1010,1087,1231,1257]" name="Russia" pageId="3" pageNumber="489">Russia</collectingCountry>
,
<geoCoordinate id="3FC707F5FFA3983D051BA893778EFB38" box="[1099,1195,1230,1257]" degrees="57" direction="north" minutes="04" orientation="latitude" pageId="3" pageNumber="489" precision="925" value="57.066666">57°04'N</geoCoordinate>
,
<geoCoordinate id="3FC707F5FFA3983D05E6A8937637FB38" box="[1206,1298,1230,1257]" degrees="65" direction="east" minutes="04" orientation="longitude" pageId="3" pageNumber="489" precision="925" value="65.066666">65°04'E</geoCoordinate>
,
<date id="2E4D47F2FFA3983D044EA89273F7FADC" pageId="3" pageNumber="489" value="2021-10-18">
<collectingDate id="3E09BE1AFFA3983D044EA89273F7FADC" pageId="3" pageNumber="489" value="2021-10-18">18 October 2021</collectingDate>
</date>
,
<collectorName id="F70604E4FFA3983D01B1A8AE72D6FADC" box="[225,499,1267,1293]" pageId="3" pageNumber="489">Alexander A. Khaustov</collectorName>
coll.,
<collectedFrom id="E9EF22CFFFA3983D0314A8AE7620FADF" box="[580,1285,1267,1294]" pageId="3" pageNumber="489">
in the nest of
<taxonomicName id="9DF31AB1FFA3983D03A2A8AE70A6FADD" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[754,899,1267,1292]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="489" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="6887BD20FFA3983D03A2A8AE70A6FADD" box="[754,899,1267,1292]" italics="true" pageId="3" pageNumber="489">Lasius niger</emphasis>
</taxonomicName>
(L.) (
<taxonomicName id="9DF31AB1FFA3983D029EA8AE7748FADC" box="[974,1133,1267,1293]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="489" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
:
<taxonomicName id="9DF31AB1FFA3983D052CA8AE77DBFADF" box="[1148,1278,1267,1294]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="489" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
)
</collectedFrom>
(in
<collectionCode id="3CE2F9F7FFA3983D046DA8AE76B7FADC" box="[1341,1426,1267,1293]" pageId="3" pageNumber="489">TUMZ</collectionCode>
).
</materialsCitation>
<materialsCitation id="EA9B6B6FFFA3983D01C7A94A700BFAE0" ID-GBIF-Occurrence="3763889301" box="[151,814,1303,1329]" pageId="3" pageNumber="489" typeStatus="paratype">
<typeStatus id="8548DF90FFA3983D01C7A94A722DFAE0" box="[151,264,1303,1329]" pageId="3" pageNumber="489" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="4CF5AABBFFA3983D0043A94A72BAFAE0" box="[275,415,1303,1329]" pageId="3" pageNumber="489" type="female">five females</specimenCount>
, same data as
<typeStatus id="8548DF90FFA3983D036FA94A7184FAE0" box="[575,673,1303,1329]" pageId="3" pageNumber="489" type="holotype">holotype</typeStatus>
(in
<collectionCode id="3CE2F9F7FFA3983D039DA94A7007FAE0" box="[717,802,1303,1329]" pageId="3" pageNumber="489">TUMZ</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="12E932B9FFA3983D0197A967773FF9A4" pageId="3" pageNumber="489" type="diagnosis">
<paragraph id="5A4C6132FFA3983D0197A967773FF9A4" blockId="3.[151,1437,1230,2013]" pageId="3" pageNumber="489">
<emphasis id="6887BD20FFA3983D0197A96772D2FA85" bold="true" box="[199,503,1338,1364]" pageId="3" pageNumber="489">Diagnosis (adult female).</emphasis>
Dorsal shield with weak reticulate ornamentation, bearing 39 pairs of relatively long (sub-equal), knife-shaped setae, mostly reaching base of next posterior setae in series,
<emphasis id="6887BD20FFA3983D05C0A902778BFAA8" box="[1168,1198,1375,1401]" italics="true" pageId="3" pageNumber="489">Z5</emphasis>
shorter than
<emphasis id="6887BD20FFA3983D046FA902767CFAA8" box="[1343,1369,1375,1401]" italics="true" pageId="3" pageNumber="489">J5</emphasis>
(ratio of
<emphasis id="6887BD20FFA3983D01E7A9DE73F7FA4C" box="[183,210,1411,1437]" italics="true" pageId="3" pageNumber="489">J5</emphasis>
/
<emphasis id="6887BD20FFA3983D018AA9DE73D2FA4C" box="[218,247,1411,1437]" italics="true" pageId="3" pageNumber="489">Z5</emphasis>
length ≃1.5); presternal area lightly sclerotised, anterior margin of shield hardly conspicuous, posterior margin distinctly concave, ratio of shield length/width (at broadest level) ≃ 0.8; genital shield ratio of length/ width (at broadest level) ≃ 2.4, shield ornamented posteromedially by 7–8 cells flanked by a median Λ-shaped ornamentation, anal shield ratio of length/width (at broadest level) ≃ 1.2, circum-anal setae smooth and subequal, post-anal seta thickened; distance between genital and anal shields is shorter than length of anal shield. Fixed digit of chelicera with six teeth of various sizes. Legs without spatulate setae but most setae conspicuously thickened and some inserted on small tubercles. Legs I and IV shorter than length of idiosoma.
</paragraph>
</subSubSection>
<subSubSection id="12E932B9FFA398360197AA2372ADFE90" lastPageId="8" lastPageNumber="494" pageId="3" pageNumber="489" type="description">
<paragraph id="5A4C6132FFA3983D0197AA2372D9F94B" blockId="3.[151,1437,1230,2013]" box="[199,508,1662,1690]" pageId="3" pageNumber="489">
<emphasis id="6887BD20FFA3983D0197AA237292F948" bold="true" box="[199,439,1662,1689]" pageId="3" pageNumber="489">Description. Female</emphasis>
(n=6)
</paragraph>
<paragraph id="5A4C6132FFA3983D0197AAFE7167F80C" blockId="3.[151,1437,1230,2013]" pageId="3" pageNumber="489">
<emphasis id="6887BD20FFA3983D0197AAFE72A6F96C" box="[199,387,1699,1725]" italics="true" pageId="3" pageNumber="489">Dorsal idiosoma</emphasis>
(
<figureCitation id="C2C87DB7FFA3983D00C1AAFE72FCF96C" box="[401,473,1699,1725]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="3" pageNumber="489">Figs 8</figureCitation>
,
<figureCitation id="C2C87DB7FFA3983D00B6AAFE717DF96F" box="[486,600,1698,1726]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="3" pageNumber="489">13, 15, 16</figureCitation>
). Dorsal shield oval shaped, small, 389–406 long, 256–265 wide, covering entire dorsal idiosoma, shield with weak reticulation, more distinct in opisthonotal and lateral regions. Shield with 39 pairs of almost uniform length setae (28–40), including two pairs of
<emphasis id="6887BD20FFA3983D029AAAB670C3F8D5" box="[970,998,1771,1796]" italics="true" pageId="3" pageNumber="489">Zx</emphasis>
setae between
<emphasis id="6887BD20FFA3983D05C5AAB67784F8D5" box="[1173,1185,1771,1796]" italics="true" pageId="3" pageNumber="489">J</emphasis>
and
<emphasis id="6887BD20FFA3983D058BAAB677CEF8D5" box="[1243,1259,1771,1796]" italics="true" pageId="3" pageNumber="489">Z</emphasis>
setae and three unpaired supernumerary seta
<emphasis id="6887BD20FFA3983D00B7AB527125F8F9" box="[487,512,1807,1832]" italics="true" pageId="3" pageNumber="489">Jx</emphasis>
between
<emphasis id="6887BD20FFA3983D033EAB5271ADF8F8" box="[622,648,1807,1833]" italics="true" pageId="3" pageNumber="489">J2</emphasis>
and
<emphasis id="6887BD20FFA3983D0392AB5271F8F8F8" box="[706,733,1807,1833]" italics="true" pageId="3" pageNumber="489">J4</emphasis>
, except
<emphasis id="6887BD20FFA3983D026EAB527071F8F8" box="[830,852,1807,1833]" italics="true" pageId="3" pageNumber="489">j1</emphasis>
,
<emphasis id="6887BD20FFA3983D0234AB5270A4F8F8" box="[868,897,1807,1833]" italics="true" pageId="3" pageNumber="489">Z5</emphasis>
(14–18) simple and thickened,
<emphasis id="6887BD20FFA3983D05BCAB527627F8F8" box="[1260,1282,1807,1833]" italics="true" pageId="3" pageNumber="489">j1</emphasis>
apically bent (hook shaped),
<emphasis id="6887BD20FFA3983D0014AB6E7279F89C" box="[324,348,1843,1869]" italics="true" pageId="3" pageNumber="489">z1</emphasis>
(15–18) and
<emphasis id="6887BD20FFA3983D00A1AB6E7129F89C" box="[497,524,1843,1869]" italics="true" pageId="3" pageNumber="489">J5</emphasis>
(21–24) shorter (
<figureCitation id="C2C87DB7FFA3983D039DAB6E706BF89F" box="[717,846,1842,1870]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="3" pageNumber="489">Figs 15, 16</figureCitation>
). All setae slightly swollen near the base and pointed at tip (knife-shaped), reaching base of following setae, setae of central area shorter (not reaching base of following setae) (
<figureCitation id="C2C87DB7FFA3983D01B6AB267209F844" box="[230,300,1915,1941]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="3" pageNumber="489">Figs 8</figureCitation>
,
<figureCitation id="C2C87DB7FFA3983D0067AB267282F847" box="[311,423,1914,1942]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="3" pageNumber="489">13, 15, 16</figureCitation>
). Dorsal shield with 22 pairs of pore-like structures, including seven pairs of gland openings (
<emphasis id="6887BD20FFA3983D01F1ABC273EFF868" box="[161,202,1951,1977]" italics="true" pageId="3" pageNumber="489">gd1</emphasis>
,
<emphasis id="6887BD20FFA3983D0189ABC27226F868" box="[217,259,1951,1977]" italics="true" pageId="3" pageNumber="489">gd2</emphasis>
,
<emphasis id="6887BD20FFA3983D0042ABC27219F868" box="[274,316,1951,1977]" italics="true" pageId="3" pageNumber="489">gd4</emphasis>
,
<emphasis id="6887BD20FFA3983D001BABC27250F868" box="[331,373,1951,1977]" italics="true" pageId="3" pageNumber="489">gd5</emphasis>
,
<emphasis id="6887BD20FFA3983D00D3ABC37288F869" box="[387,429,1950,1976]" italics="true" pageId="3" pageNumber="489">gd6</emphasis>
,
<emphasis id="6887BD20FFA3983D00ECABC272C3F868" box="[444,486,1951,1977]" italics="true" pageId="3" pageNumber="489">gd8</emphasis>
,
<emphasis id="6887BD20FFA3983D00A5ABC2713BF868" box="[501,542,1951,1977]" italics="true" pageId="3" pageNumber="489">gd9</emphasis>
) and 15 pairs of poroids,
<emphasis id="6887BD20FFA3983D0213ABC37042F869" box="[835,871,1950,1976]" italics="true" pageId="3" pageNumber="489">id6</emphasis>
indistinct. Shape, position and relative length and shape of setae shown in
<figureCitation id="C2C87DB7FFA3983D00F8AB9E7137F80C" box="[424,530,1987,2013]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="3" pageNumber="489">Figures 8</figureCitation>
,
<figureCitation id="C2C87DB7FFA3983D034FAB9E711BF80C" box="[543,574,1987,2013]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="3" pageNumber="489">13</figureCitation>
.
</paragraph>
<caption id="0E8C31BAFFA4983A01C7ABCA7004F805" ID-DOI="http://doi.org/10.5281/zenodo.6530964" ID-Zenodo-Dep="6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="4" pageNumber="490" startId="4.[151,264,1943,1967]" targetBox="[181,1397,196,1901]" targetPageId="4">
<paragraph id="5A4C6132FFA4983A01C7ABCA7004F805" blockId="4.[151,1437,1943,2004]" pageId="4" pageNumber="490">
<emphasis id="6887BD20FFA4983A01C7ABCA726FF87E" bold="true" box="[151,330,1943,1967]" pageId="4" pageNumber="490">FIGURES 8–12.</emphasis>
<taxonomicName id="9DF31AB1FFA4983A0004ABC5719EF861" authority="Joharchi" authorityName="Joharchi" authorityYear="2022" box="[340,699,1944,1968]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="490" phylum="Arthropoda" rank="species" species="lasiophilus" status="sp. nov.">
<emphasis id="6887BD20FFA4983A0004ABC5717EF861" box="[340,603,1944,1968]" italics="true" pageId="4" pageNumber="490">Cosmolaelaps lasiophilus</emphasis>
Joharchi
</taxonomicName>
<taxonomicNameLabel id="73B4005BFFA4983A0394ABC5703CF87E" box="[708,793,1944,1968]" pageId="4" pageNumber="490" rank="species">
<emphasis id="6887BD20FFA4983A0394ABC57036F861" bold="true" box="[708,787,1944,1968]" pageId="4" pageNumber="490">sp. nov</emphasis>
.
</taxonomicNameLabel>
, female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera.
</paragraph>
</caption>
<caption id="0E8C31BAFFA5983B01C7AB297224F829" ID-DOI="http://doi.org/10.5281/zenodo.6530968" ID-Zenodo-Dep="6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="5" pageNumber="491" startId="5.[151,264,1908,1932]" targetBox="[198,1388,181,1884]" targetPageId="5">
<paragraph id="5A4C6132FFA5983B01C7AB297224F829" blockId="5.[151,1437,1908,2040]" pageId="5" pageNumber="491">
<emphasis id="6887BD20FFA5983B01C7AB297270F85D" bold="true" box="[151,341,1908,1932]" pageId="5" pageNumber="491">FIGURES 13–20.</emphasis>
DIC micrographs of
<taxonomicName id="9DF31AB1FFA5983B0362AB2970B3F85D" authority="Joharchi" authorityName="Joharchi" authorityYear="2022" box="[562,918,1908,1932]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="491" phylum="Arthropoda" rank="species" species="lasiophilus" status="sp. nov.">
<emphasis id="6887BD20FFA5983B0362AB29701DF85D" box="[562,824,1908,1932]" italics="true" pageId="5" pageNumber="491">Cosmolaelaps lasiophilus</emphasis>
Joharchi
</taxonomicName>
<taxonomicNameLabel id="73B4005BFFA5983B02CDAB2870D5F85D" box="[925,1008,1909,1933]" pageId="5" pageNumber="491" rank="species">
<emphasis id="6887BD20FFA5983B02CDAB2870CFF85C" bold="true" box="[925,1002,1909,1933]" pageId="5" pageNumber="491">sp. nov</emphasis>
.
</taxonomicNameLabel>
, female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures.
</paragraph>
</caption>
<caption id="0E8C31BAFFA6983801C7ABDB7393F813" ID-DOI="http://doi.org/10.5281/zenodo.6530972" ID-Zenodo-Dep="6530972" httpUri="https://zenodo.org/record/6530972/files/figure.png" pageId="6" pageNumber="492" startId="6.[151,250,1926,1950]" targetBox="[161,1425,197,1887]" targetPageId="6">
<paragraph id="5A4C6132FFA6983801C7ABDB7393F813" blockId="6.[151,1436,1926,1986]" pageId="6" pageNumber="492">
<emphasis id="6887BD20FFA6983801C7ABDB7263F84F" bold="true" box="[151,326,1926,1950]" pageId="6" pageNumber="492">FIGURE 21–24.</emphasis>
<taxonomicName id="9DF31AB1FFA69838001CABDB718BF84F" authority="Joharchi" authorityName="Joharchi" authorityYear="2022" box="[332,686,1926,1950]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="492" phylum="Arthropoda" rank="species" species="lasiophilus" status="sp. nov.">
<emphasis id="6887BD20FFA69838001CABDB7174F84F" box="[332,593,1926,1950]" italics="true" pageId="6" pageNumber="492">Cosmolaelaps lasiophilus</emphasis>
Joharchi
</taxonomicName>
<taxonomicNameLabel id="73B4005BFFA6983803E5ABDB7022F84F" box="[693,775,1926,1950]" pageId="6" pageNumber="492" rank="species">
<emphasis id="6887BD20FFA6983803E5ABDB7025F84F" bold="true" box="[693,768,1926,1950]" pageId="6" pageNumber="492">sp. nov</emphasis>
.
</taxonomicNameLabel>
, female. 21, leg I (trochanter-tibia); 22, leg II; 23, leg III; 24, leg IV.
</paragraph>
</caption>
<paragraph id="5A4C6132FFA798390197ACCA7050FBAC" blockId="7.[151,1437,150,2014]" pageId="7" pageNumber="493">
<emphasis id="6887BD20FFA798390197ACCA72AEFF60" box="[199,395,151,177]" italics="true" pageId="7" pageNumber="493">Ventral idiosoma</emphasis>
(
<figureCitation id="C2C87DB7FFA7983900CFACCA72CBFF60" box="[415,494,151,177]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="7" pageNumber="493">Figs 9</figureCitation>
,
<figureCitation id="C2C87DB7FFA7983900AFACCA7168FF60" box="[511,589,151,177]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="7" pageNumber="493">14, 17</figureCitation>
). Tritosternum with paired pilose laciniae (64–68), fused basally (3–5), columnar base 23–27 × 12–15 wide; presternal area lightly sclerotised, with a few transverse curved lines, merged to sternal shield. Sternal shield (length 92–96) narrowest between coxae II (74–80), widest (114–118), anterior margin of shield hardly conspicuous, posterior margin distinctly concave, anterolateral corners narrowly extending between coxae I–II, distally bearing gland pores
<emphasis id="6887BD20FFA798390331AD7A71ACFE90" box="[609,649,295,321]" italics="true" pageId="7" pageNumber="493">gvb</emphasis>
; endopodal elements between coxae II and III fused with shield and endopodal elements III/IV elongate, narrow and curved. Sternal shield with three pairs of smooth setae (
<emphasis id="6887BD20FFA798390478AD16766DFEB4" box="[1320,1352,331,357]" italics="true" pageId="7" pageNumber="493">st1</emphasis>
31–33,
<emphasis id="6887BD20FFA7983901C7AD327392FE58" box="[151,183,367,393]" italics="true" pageId="7" pageNumber="493">st2</emphasis>
28–30,
<emphasis id="6887BD20FFA798390046AD327212FE58" box="[278,311,367,393]" italics="true" pageId="7" pageNumber="493">st3</emphasis>
24–27) and two pairs of slit-shaped pore-like structures (
<emphasis id="6887BD20FFA798390299AD3270C9FE58" box="[969,1004,367,393]" italics="true" pageId="7" pageNumber="493">iv1</emphasis>
adjacent to setae
<emphasis id="6887BD20FFA7983905E9AD3277FFFE58" box="[1209,1242,367,393]" italics="true" pageId="7" pageNumber="493">st1</emphasis>
;
<emphasis id="6887BD20FFA7983905BBAD327628FE58" box="[1259,1293,367,393]" italics="true" pageId="7" pageNumber="493">iv2</emphasis>
between
<emphasis id="6887BD20FFA79839042CAD3276B9FE58" box="[1404,1436,367,393]" italics="true" pageId="7" pageNumber="493">st2</emphasis>
and
<emphasis id="6887BD20FFA798390197ADCE73CDFE7C" box="[199,232,403,429]" italics="true" pageId="7" pageNumber="493">st3</emphasis>
), setae not reaching base of following setae, shield with lineate-reticulate ornamentation on lateral surface, longitudinally median part smooth (or faintly reticulated) (
<figureCitation id="C2C87DB7FFA79839024DADEA7040FE00" box="[797,869,439,465]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="7" pageNumber="493">Figs 9</figureCitation>
,
<figureCitation id="C2C87DB7FFA79839023FADEA70ABFE00" box="[879,910,439,465]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="7" pageNumber="493">14</figureCitation>
). Metasternal setae
<emphasis id="6887BD20FFA798390539ADEA77ACFE00" box="[1129,1161,439,465]" italics="true" pageId="7" pageNumber="493">st4</emphasis>
(24–27) and metasternal poroids located on soft integument; metasternal platelets absent. Genital shield flask-shaped, slightly expanded laterally past level of setae
<emphasis id="6887BD20FFA798390097ADA272C2FDC8" box="[455,487,511,537]" italics="true" pageId="7" pageNumber="493">st5</emphasis>
, length 165–180, maximum width 71–75, posterior margin rounded, anterior hyaline margin of shield irregularly convex, overlapping posterior area of sternal shield (extending beyond base of setae
<emphasis id="6887BD20FFA7983901C7AE1A739DFDB0" box="[151,184,583,609]" italics="true" pageId="7" pageNumber="493">st3</emphasis>
), shield ornamented posteromedially by 7–8 cells flanked by a median Λ-shaped ornamentation, bearing a pair of simple setae
<emphasis id="6887BD20FFA798390001AE367254FD54" box="[337,369,619,645]" italics="true" pageId="7" pageNumber="493">st5</emphasis>
(22–24) (
<figureCitation id="C2C87DB7FFA7983900B7AE367116FD54" box="[487,563,619,645]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="7" pageNumber="493">Figs 9</figureCitation>
,
<figureCitation id="C2C87DB7FFA798390313AE367147FD54" box="[579,610,619,645]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="7" pageNumber="493">14</figureCitation>
). Shield flanked by two pairs of minute, elongate paragenital platelets; paragenital poroids
<emphasis id="6887BD20FFA79839002CAED272BBFD78" box="[380,414,655,681]" italics="true" pageId="7" pageNumber="493">iv5</emphasis>
located on soft cuticle lateral to shield near seta
<emphasis id="6887BD20FFA79839028FAED27725FD78" box="[991,1024,655,681]" italics="true" pageId="7" pageNumber="493">st5</emphasis>
. Anal shield subtriangular, rounded anteriorly, length 65–70, width 52–55, anterior half lineate-reticulate, para-anal setae and post-anal seta subequal in length (15–18), post-anal seta thicker, cribrum consisting of a terminal tuft with 3–4 irregular rows of spicules, anteriormost rows of spicules extending slightly beyond post-anal seta (
<figureCitation id="C2C87DB7FFA798390294AEA6772BFCC4" box="[964,1038,763,789]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="7" pageNumber="493">Figs 9</figureCitation>
,
<figureCitation id="C2C87DB7FFA79839054BAEA67740FCC4" box="[1051,1125,763,789]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="7" pageNumber="493">14, 17</figureCitation>
); anal gland pores (
<emphasis id="6887BD20FFA798390416AEA6764AFCC4" box="[1350,1391,763,789]" italics="true" pageId="7" pageNumber="493">gv3</emphasis>
) on anterolateral margin of anal shield. Soft opisthogastric cuticle surrounding genital and anal shields with one pair of suboval metapodal plates (17–19 long × 4–6 wide), six pairs of poroids (five
<emphasis id="6887BD20FFA798390556AF1E770DFC8D" box="[1030,1064,835,860]" italics="true" pageId="7" pageNumber="493">ivo</emphasis>
;
<emphasis id="6887BD20FFA798390568AF1E777FFC8D" box="[1080,1114,835,860]" italics="true" pageId="7" pageNumber="493">ivp</emphasis>
) and 16 pairs of moderately thick setae (
<emphasis id="6887BD20FFA798390075AF3A72A4FC50" box="[293,385,871,897]" italics="true" pageId="7" pageNumber="493">Jv1–Jv5</emphasis>
,
<emphasis id="6887BD20FFA7983900C2AF3A72D5FC51" box="[402,496,871,897]" italics="true" pageId="7" pageNumber="493">Zv1– Zv</emphasis>
5,
<emphasis id="6887BD20FFA79839035FAF3A7179FC50" box="[527,604,871,897]" italics="true" pageId="7" pageNumber="493">R1–R4</emphasis>
,
<emphasis id="6887BD20FFA79839033DAF3A71C4FC50" box="[621,737,871,897]" italics="true" pageId="7" pageNumber="493">UR2–UR3</emphasis>
); almost all setae uniform in length (11–16), and arising on small sclerotised platelet, except
<emphasis id="6887BD20FFA798390340AFD67176FC74" box="[528,595,907,933]" italics="true" pageId="7" pageNumber="493">Jv1–3</emphasis>
and
<emphasis id="6887BD20FFA7983903DDAFD67192FC74" box="[653,695,907,933]" italics="true" pageId="7" pageNumber="493">Zv1</emphasis>
longer (20–24) and setiform (
<figureCitation id="C2C87DB7FFA798390540AFD6777FFC74" box="[1040,1114,907,933]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="7" pageNumber="493">Figs 9</figureCitation>
,
<figureCitation id="C2C87DB7FFA798390537AFD67797FC74" box="[1127,1202,907,933]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="7" pageNumber="493">14, 17</figureCitation>
). Peritrematal shield well developed laterad peritremes, bearing one gland pore and one poroid at level near coxae II–III, post-stigmatic extension of shield relatively wide, not reaching posterior level of coxae IV, with two pairs of poroids and one pair of gland pores; anterior region of peritrematal shield fused to anterior margin of dorsal shield behind setae
<emphasis id="6887BD20FFA7983901C7A8467395FBE4" box="[151,176,1051,1077]" italics="true" pageId="7" pageNumber="493">z1</emphasis>
. Peritremes long, extending to mid-level of coxa I (near
<emphasis id="6887BD20FFA79839026DA8467072FBE4" box="[829,855,1051,1077]" italics="true" pageId="7" pageNumber="493">s1</emphasis>
). Two exopodal platelets between coxae II–III and III–IV present, anterior platelet small and subtriangular, those between coxae III–IV slightly extending posteriorly; parapodal platelets strip-like, bearing gland pore
<emphasis id="6887BD20FFA7983903EDA83E71C0FBAC" box="[701,741,1123,1149]" italics="true" pageId="7" pageNumber="493">gv2</emphasis>
(
<figureCitation id="C2C87DB7FFA7983903A4A83E7018FBAC" box="[756,829,1123,1149]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="7" pageNumber="493">Figs 9</figureCitation>
,
<figureCitation id="C2C87DB7FFA798390219A83E704DFBAC" box="[841,872,1123,1149]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="7" pageNumber="493">14</figureCitation>
).
</paragraph>
<paragraph id="5A4C6132FFA798390197A8DA779AF9FC" blockId="7.[151,1437,150,2014]" pageId="7" pageNumber="493">
<taxonomicName id="9DF31AB1FFA798390197A8DA7273FB70" authorityName="Krefft" authorityYear="1871" box="[199,342,1159,1185]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="7" pageNumber="493" phylum="Nematoda" rank="genus">
<emphasis id="6887BD20FFA798390197A8DA7273FB70" box="[199,342,1159,1185]" italics="true" pageId="7" pageNumber="493">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="C2C87DB7FFA798390037A8DA72CFFB70" box="[359,490,1159,1185]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="7" pageNumber="493">Figs 10–12</figureCitation>
,
<figureCitation id="C2C87DB7FFA7983900A7A8DA7167FB70" box="[503,578,1159,1185]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="7" pageNumber="493">18, 19</figureCitation>
). Anterior margin of epistome convex, irregularly denticulate, with smooth median spike (
<figureCitation id="C2C87DB7FFA798390015A8F672BEFB14" box="[325,411,1195,1221]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="7" pageNumber="493">Fig. 11</figureCitation>
). Hypostomal groove with six transverse rows of denticles, each row with 8–19 small denticles, with smooth posterior transverse line, groove wider anteriorly, progressively narrowing from anterior to posterior, anteriormost row slightly convex (
<figureCitation id="C2C87DB7FFA7983903DAA8AE71C5FADC" box="[650,736,1267,1293]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="7" pageNumber="493">Figs 10</figureCitation>
,
<figureCitation id="C2C87DB7FFA7983903BCA8AE702EFADC" box="[748,779,1267,1293]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="7" pageNumber="493">18</figureCitation>
). Corniculi horn-like, internal malae longer than corniculi, comprising a pair of pilose median projections, and 7–9 lateral, longer, tentacle-like, smooth projections (
<figureCitation id="C2C87DB7FFA79839046AA94A76ABFAE0" box="[1338,1422,1303,1329]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="7" pageNumber="493">Fig. 10</figureCitation>
); labrum acuminate, considerably longer than internal malae, fringed; hypostomal and capitular setae smooth,
<emphasis id="6887BD20FFA7983904D0A96676B9FA84" box="[1408,1436,1339,1365]" italics="true" pageId="7" pageNumber="493">h3</emphasis>
(34–40)&gt;
<emphasis id="6887BD20FFA79839005EA902720FFAA8" box="[270,298,1375,1401]" italics="true" pageId="7" pageNumber="493">h1</emphasis>
(28–31)&gt;
<emphasis id="6887BD20FFA7983900F8A90272E7FAA9" box="[424,450,1375,1400]" italics="true" pageId="7" pageNumber="493">pc</emphasis>
(21–25)&gt;
<emphasis id="6887BD20FFA798390310A9027179FAA8" box="[576,604,1375,1401]" italics="true" pageId="7" pageNumber="493">h2</emphasis>
(11–14) (
<figureCitation id="C2C87DB7FFA798390399A9027005FAA8" box="[713,800,1375,1401]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="7" pageNumber="493">Figs 10</figureCitation>
,
<figureCitation id="C2C87DB7FFA79839027CA902706EFAA8" box="[812,843,1375,1401]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="7" pageNumber="493">18</figureCitation>
). Chaetotaxy of palps: trochanter 2, femur 5, genu 6, tibia 14, tarsus 15, all setae smooth except palpfemur with seta
<emphasis id="6887BD20FFA798390208A9DE7051FA4C" box="[856,884,1411,1437]" italics="true" pageId="7" pageNumber="493">d3</emphasis>
thickened and inserted on small tubercles, seta
<emphasis id="6887BD20FFA7983904D6A9DE76B9FA4C" box="[1414,1436,1411,1437]" italics="true" pageId="7" pageNumber="493">al</emphasis>
thickened and apically subspatulate; palpgenu with
<emphasis id="6887BD20FFA79839038EA9FA7027FA10" box="[734,770,1447,1473]" italics="true" pageId="7" pageNumber="493">al1</emphasis>
stout,
<emphasis id="6887BD20FFA79839021EA9FA7057FA10" box="[846,882,1447,1473]" italics="true" pageId="7" pageNumber="493">al2</emphasis>
paddle-like; palp tarsal claw two-tined (
<figureCitation id="C2C87DB7FFA79839046BA9FA76AAFA10" box="[1339,1423,1447,1473]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="7" pageNumber="493">Fig. 19</figureCitation>
). Supralabral process indistinct. Fixed digit of chelicera with an offset and subapical tooth (gabelzhan), followed by five teeth, a setaceous pilus dentilis, dorsal cheliceral seta prostrate, arthrodial membrane with a rounded flap and normal filaments; cheliceral lyrifissures distinct, movable digit of chelicera bidentate (
<figureCitation id="C2C87DB7FFA79839050EAA4E7797F9FC" box="[1118,1202,1555,1581]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="7" pageNumber="493">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="5A4C6132FFA798390197AA6A7613F948" blockId="7.[151,1437,150,2014]" pageId="7" pageNumber="493">
<emphasis id="6887BD20FFA798390197AA6A72F5F981" box="[199,464,1591,1616]" italics="true" pageId="7" pageNumber="493">Insemination structures</emphasis>
(
<figureCitation id="C2C87DB7FFA7983900B1AA6A7113F980" box="[481,566,1591,1617]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="7" pageNumber="493">Fig. 20</figureCitation>
). Laelapid-type sperm access system, tubulus long, wider at the solenostome level of coxa III and entering sacculus via a pair of circular openings. Sacculus irregular, the proximal ends of the tubulus slightly swollen at the junction with ramus, ends of ramus with considerably horn-like projection.
</paragraph>
<paragraph id="5A4C6132FFA798360197AAFE7677FECC" blockId="7.[151,1437,150,2014]" lastBlockId="8.[151,1437,151,1725]" lastPageId="8" lastPageNumber="494" pageId="7" pageNumber="493">
<emphasis id="6887BD20FFA798390197AAFE73D9F96D" box="[199,252,1699,1724]" italics="true" pageId="7" pageNumber="493">Legs</emphasis>
(
<figureCitation id="C2C87DB7FFA79839005CAAFE72AAF96C" box="[268,399,1699,1725]" captionStart="FIGURE 21–24" captionStartId="6.[151,250,1926,1950]" captionTargetBox="[161,1425,197,1887]" captionTargetId="figure-18@6.[152,1436,181,1902]" captionTargetPageId="6" captionText="FIGURE 21–24. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 21, leg I (trochanter-tibia); 22, leg II; 23, leg III; 24, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530972" httpUri="https://zenodo.org/record/6530972/files/figure.png" pageId="7" pageNumber="493">Figs 21–24</figureCitation>
). Legs II and III short (260–278, 240–255), I and IV longer (365–385, 347–364) (excluding pre-tarsus). Chaetotaxy normal for free-living
<taxonomicName id="9DF31AB1FFA7983903F0AA9A703EF930" box="[672,795,1735,1761]" class="Arachnida" family="Laelapidae" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="493" phylum="Arthropoda" rank="family">Laelapidae</taxonomicName>
(
<emphasis id="6887BD20FFA79839027BAA9A704CF931" box="[811,873,1735,1760]" italics="true" pageId="7" pageNumber="493">sensu</emphasis>
<bibRefCitation id="3E621CC3FFA798390221AA9A7729F933" author="Evans, G. O." box="[881,1036,1735,1762]" pageId="7" pageNumber="493" pagination="275 - 303" refId="ref14188" refString="Evans, G. O. (1963 a) Observations on the chaetotaxy of the legs in the free-living Gamasina (Acari: Mesostigmata). Bulletin of the British Museum (Natural History) Zoology, 10 (5), 275 - 303. https: // doi. org / 10.5962 / bhl. part. 20528" type="journal article" year="1963">Evans, 1963a</bibRefCitation>
): Leg I (
<figureCitation id="C2C87DB7FFA79839053CAA9A77E5F930" box="[1132,1216,1735,1761]" captionStart="FIGURE 21–24" captionStartId="6.[151,250,1926,1950]" captionTargetBox="[161,1425,197,1887]" captionTargetId="figure-18@6.[152,1436,181,1902]" captionTargetPageId="6" captionText="FIGURE 21–24. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 21, leg I (trochanter-tibia); 22, leg II; 23, leg III; 24, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530972" httpUri="https://zenodo.org/record/6530972/files/figure.png" pageId="7" pageNumber="493">Fig. 21</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-1/1, 0/2-1 (
<emphasis id="6887BD20FFA7983900FCAAB672E7F8D4" box="[428,450,1771,1797]" italics="true" pageId="7" pageNumber="493">al</emphasis>
and
<emphasis id="6887BD20FFA7983900ADAAB6713CF8D4" box="[509,537,1771,1797]" italics="true" pageId="7" pageNumber="493">ad</emphasis>
thickened), femur 2-3/1, 2/3-2 (
<emphasis id="6887BD20FFA7983902DFAAB67091F8D4" box="[911,948,1771,1797]" italics="true" pageId="7" pageNumber="493">al1</emphasis>
,
<emphasis id="6887BD20FFA798390294AAB670CBF8D4" box="[964,1006,1771,1797]" italics="true" pageId="7" pageNumber="493">ad3</emphasis>
and
<emphasis id="6887BD20FFA798390579AAB67776F8D4" box="[1065,1107,1771,1797]" italics="true" pageId="7" pageNumber="493">pd2</emphasis>
thickened), genu 2-3/2, 3/1- 2 (dorsals and laterals thickened and
<emphasis id="6887BD20FFA798390319AB5271B5F8F8" box="[585,656,1807,1833]" italics="true" pageId="7" pageNumber="493">ad1–3</emphasis>
inserted on small tubercles), tibia 2-3/2, 3/1-2 (dorsals and laterals thickened). Leg II (
<figureCitation id="C2C87DB7FFA798390026AB6E72E9F89C" box="[374,460,1843,1869]" captionStart="FIGURE 21–24" captionStartId="6.[151,250,1926,1950]" captionTargetBox="[161,1425,197,1887]" captionTargetId="figure-18@6.[152,1436,181,1902]" captionTargetPageId="6" captionText="FIGURE 21–24. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 21, leg I (trochanter-tibia); 22, leg II; 23, leg III; 24, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530972" httpUri="https://zenodo.org/record/6530972/files/figure.png" pageId="7" pageNumber="493">Fig. 22</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-0/1, 0/2-1 (
<emphasis id="6887BD20FFA798390299AB6E70FAF89C" box="[969,991,1843,1869]" italics="true" pageId="7" pageNumber="493">al</emphasis>
slightly thickened), femur 2-3/1, 2/2-1 (
<emphasis id="6887BD20FFA7983901F1AB0A73EFF8A0" box="[161,202,1879,1905]" italics="true" pageId="7" pageNumber="493">ad1</emphasis>
,
<emphasis id="6887BD20FFA798390188AB0A7227F8A0" box="[216,258,1879,1905]" italics="true" pageId="7" pageNumber="493">pd1</emphasis>
and
<emphasis id="6887BD20FFA798390068AB0A7244F8A0" box="[312,353,1879,1905]" italics="true" pageId="7" pageNumber="493">pd2</emphasis>
thickened and inserted on small tubercles,
<emphasis id="6887BD20FFA798390212AB0A7049F8A0" box="[834,876,1879,1905]" italics="true" pageId="7" pageNumber="493">pd2</emphasis>
longest), genu 2-3/1, 2/1-2 (dorsals thickened and inserted on small tubercles), tibia 2-2/1, 2/1-2. Leg III (
<figureCitation id="C2C87DB7FFA7983903A2AB267060F844" box="[754,837,1915,1941]" captionStart="FIGURE 21–24" captionStartId="6.[151,250,1926,1950]" captionTargetBox="[161,1425,197,1887]" captionTargetId="figure-18@6.[152,1436,181,1902]" captionTargetPageId="6" captionText="FIGURE 21–24. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 21, leg I (trochanter-tibia); 22, leg II; 23, leg III; 24, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530972" httpUri="https://zenodo.org/record/6530972/files/figure.png" pageId="7" pageNumber="493">Fig. 23</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-1/1, 0/2-0 (
<emphasis id="6887BD20FFA798390477AB267666F844" box="[1319,1347,1915,1941]" italics="true" pageId="7" pageNumber="493">ad</emphasis>
slightly thickened), femur 1-2/1, 1/0-1 (
<emphasis id="6887BD20FFA7983900A5ABC2713AF868" box="[501,543,1951,1977]" italics="true" pageId="7" pageNumber="493">ad1</emphasis>
,
<emphasis id="6887BD20FFA79839037CABC2716DF868" box="[556,584,1951,1977]" italics="true" pageId="7" pageNumber="493">pd</emphasis>
and
<emphasis id="6887BD20FFA79839032CABC271B7F868" box="[636,658,1951,1977]" italics="true" pageId="7" pageNumber="493">pl</emphasis>
thickened,
<emphasis id="6887BD20FFA798390241ABC2701EF868" box="[785,827,1951,1977]" italics="true" pageId="7" pageNumber="493">ad1</emphasis>
inserted on small tubercle), genu 2-2/1, 2/1-1 (all setae on segment slightly thickened), tibia: 2-1/1, 2/1-1 (all setae on segment slightly thickened). Leg IV (
<figureCitation id="C2C87DB7FFA7983905AEAB9E7676F80C" box="[1278,1363,1987,2013]" captionStart="FIGURE 21–24" captionStartId="6.[151,250,1926,1950]" captionTargetBox="[161,1425,197,1887]" captionTargetId="figure-18@6.[152,1436,181,1902]" captionTargetPageId="6" captionText="FIGURE 21–24. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 21, leg I (trochanter-tibia); 22, leg II; 23, leg III; 24, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530972" httpUri="https://zenodo.org/record/6530972/files/figure.png" pageId="7" pageNumber="493">Fig. 24</figureCitation>
): coxa 0-0/1, 0/0-0, trochanter 1-1/1, 0/2-0 (
<emphasis id="6887BD20FFA898360361ACCA7168FF60" box="[561,589,151,177]" italics="true" pageId="8" pageNumber="494">ad</emphasis>
thickened), femur 1-2/1, 1/0-1 (
<emphasis id="6887BD20FFA8983602E0ACCA70FFFF60" box="[944,986,151,177]" italics="true" pageId="8" pageNumber="494">ad1</emphasis>
and
<emphasis id="6887BD20FFA89836055EACCA771CFF60" box="[1038,1081,151,177]" italics="true" pageId="8" pageNumber="494">ad2</emphasis>
thickened and inserted on small tubercles), genu 2-2/1, 3/0-1 (dorsals thickened), tibia 2-1/1, 3/1-2 (dorsals and laterals thickened,
<emphasis id="6887BD20FFA8983605A4ACE67632FF04" box="[1268,1303,187,213]" italics="true" pageId="8" pageNumber="494">al2</emphasis>
inserted on small tubercle). Tarsi II–IV with 18 setae (3- 3/2, 3/2-3 +
<emphasis id="6887BD20FFA898360278AC82706CFF29" box="[808,841,223,248]" italics="true" pageId="8" pageNumber="494">mv</emphasis>
,
<emphasis id="6887BD20FFA898360207AC82705CFF28" box="[855,889,223,249]" italics="true" pageId="8" pageNumber="494">md</emphasis>
); almost all setae on tarsus II–IV thickened, see
<figureCitation id="C2C87DB7FFA8983601C7AD5E7219FECC" box="[151,316,259,285]" captionStart="FIGURE 21–24" captionStartId="6.[151,250,1926,1950]" captionTargetBox="[161,1425,197,1887]" captionTargetId="figure-18@6.[152,1436,181,1902]" captionTargetPageId="6" captionText="FIGURE 21–24. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 21, leg I (trochanter-tibia); 22, leg II; 23, leg III; 24, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530972" httpUri="https://zenodo.org/record/6530972/files/figure.png" pageId="8" pageNumber="494">Figures 22–24</figureCitation>
. All pretarsi with well-developed paired claws, rounded pulvilli and normal ambulacral stalk.
</paragraph>
<paragraph id="5A4C6132FFA898360197AD7A72ADFE90" blockId="8.[151,1437,151,1725]" box="[199,392,295,321]" pageId="8" pageNumber="494">
<emphasis id="6887BD20FFA898360197AD7A722EFE90" bold="true" box="[199,267,295,321]" pageId="8" pageNumber="494">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="12E932B9FFA898360197AD1670BEFE58" pageId="8" pageNumber="494" type="etymology">
<paragraph id="5A4C6132FFA898360197AD1670BEFE58" blockId="8.[151,1437,151,1725]" pageId="8" pageNumber="494">
<emphasis id="6887BD20FFA898360197AD16726DFEB4" bold="true" box="[199,328,331,357]" pageId="8" pageNumber="494">Etymology</emphasis>
. The specific name of the new species is derived from its occurrence in nests of ants of the genus
<taxonomicName id="9DF31AB1FFA8983601C7AD3273C5FE59" authorityName="Fabricius" authorityYear="1804" box="[151,224,367,392]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="494" phylum="Arthropoda" rank="genus">
<emphasis id="6887BD20FFA8983601C7AD3273C5FE59" box="[151,224,367,392]" italics="true" pageId="8" pageNumber="494">Lasius</emphasis>
</taxonomicName>
and Greek
<emphasis id="6887BD20FFA898360032AD3372B9FE58" box="[354,412,366,393]" italics="true" pageId="8" pageNumber="494">ΦΙΛία</emphasis>
(philia), meaning &quot;friendship&quot; or &quot;fondness&quot;.
</paragraph>
</subSubSection>
<subSubSection id="12E932B9FFA898360197ADCF70F6F96C" pageId="8" pageNumber="494" type="diagnosis">
<paragraph id="5A4C6132FFA898360197ADCF762AFDB0" blockId="8.[151,1437,151,1725]" pageId="8" pageNumber="494">
<emphasis id="6887BD20FFA898360197ADCF72F5FE7D" bold="true" box="[199,464,402,428]" pageId="8" pageNumber="494">Differential diagnosis.</emphasis>
<taxonomicName id="9DF31AB1FFA89836008AADCE71D8FE7C" authorityName="Joharchi" authorityYear="2022" box="[474,765,403,429]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="lasiophilus">
<emphasis id="6887BD20FFA89836008AADCE71D8FE7C" box="[474,765,403,429]" italics="true" pageId="8" pageNumber="494">Cosmolaelaps lasiophilus</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="9DF31AB1FFA8983602A1ADCE72E4FE03" authority="(Michael, 1891) sensu Evans &amp; Till (1966)" authorityName="sensu Evans &amp; Till" authorityYear="1966" baseAuthorityName="Michael" baseAuthorityYear="1891" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="vacua">
<emphasis id="6887BD20FFA8983602A1ADCE77F9FE7D" box="[1009,1244,403,429]" italics="true" pageId="8" pageNumber="494">Cosmolaelaps vacua</emphasis>
(
<bibRefCitation id="3E621CC3FFA8983605BDADCE76B1FE7C" author="Michael, A. D." box="[1261,1428,403,429]" pageId="8" pageNumber="494" pagination="638 - 654" refId="ref15960" refString="Michael, A. D. (1891) On the association of gamasids with ants. Proceedings of the Zoological Society of London, 4, 638 - 654." type="journal article" year="1891">Michael, 1891</bibRefCitation>
)
<emphasis id="6887BD20FFA8983601C7ADEA73F0FE01" box="[151,213,439,464]" italics="true" pageId="8" pageNumber="494">sensu</emphasis>
<bibRefCitation id="3E621CC3FFA89836018DADEA72E4FE03" author="Evans, G. O. &amp; Till, W. M." box="[221,449,439,466]" pageId="8" pageNumber="494" pagination="107 - 370" refId="ref14300" refString="Evans, G. O. &amp; Till, W. M. (1966) Studies on the British Dermanyssidae (Acari: Mesostigmata). Part II. Classification. Bulletin of the British Museum (Natural History) Zoology, 14, 107 - 370." type="journal article" year="1966">Evans &amp; Till (1966)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="9DF31AB1FFA89836009DADEA71D9FE00" authority="Zeman, 1982" authorityName="Zeman" authorityYear="1982" box="[461,764,439,465]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="thysanifer">
<emphasis id="6887BD20FFA89836009DADEA72C5FE00" box="[461,480,439,465]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA8983600BEADEA717BFE00" box="[494,606,439,465]" italics="true" pageId="8" pageNumber="494">thysanifer</emphasis>
<bibRefCitation id="3E621CC3FFA898360335ADEA71D9FE00" author="Zeman, P." box="[613,764,439,465]" pageId="8" pageNumber="494" pagination="231 - 237" refId="ref16896" refString="Zeman, P. (1982) Two new species of Hypoaspidinae (Acari: Mesostigmata: Dermanysside) associated with ants. Vestnik Ceskoslovenske Spolecnosti Zoologicke, 46, 231 - 237." type="journal article" year="1982">Zeman, 1982</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="9DF31AB1FFA898360258ADEA7770FE00" authority="Ryke, 1963" authorityName="Ryke" authorityYear="1963" box="[776,1109,439,465]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="transvaalensis">
<emphasis id="6887BD20FFA898360258ADEA703EFE00" box="[776,795,439,465]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA898360279ADEA70EEFE00" box="[809,971,439,465]" italics="true" pageId="8" pageNumber="494">transvaalensis</emphasis>
<bibRefCitation id="3E621CC3FFA898360282ADEA7770FE00" author="Ryke, P. A. J." box="[978,1109,439,465]" pageId="8" pageNumber="494" pagination="1 - 15" refId="ref16407" refString="Ryke, P. A. J. (1963) Some free-living Hypoaspidinae (Acari: Mesostigmata) from South Africa. Revista de Biologia, 5, 1 - 15." type="journal article" year="1963">Ryke, 1963</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="9DF31AB1FFA898360530ADEA76B2FE00" authority="(Berlese, 1905)" baseAuthorityName="Berlese" baseAuthorityYear="1905" box="[1120,1431,438,465]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="6887BD20FFA898360530ADEA7756FE00" box="[1120,1139,439,465]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA8983605D1ADEA77C5FE00" box="[1153,1248,439,465]" italics="true" pageId="8" pageNumber="494">robustus</emphasis>
(
<bibRefCitation id="3E621CC3FFA8983605A0ADEA76AAFE00" author="Berlese, A." box="[1264,1423,438,465]" pageId="8" pageNumber="494" pagination="154 - 176" refId="ref13741" refString="Berlese, A. (1905) Acari nuovi. Manipulus IV. Redia, 2, 154 - 176." type="journal article" year="1905">Berlese, 1905</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="9DF31AB1FFA8983601C7AD8672FEFE24" authority="Joharchi 2019" authorityName="Joharchi" authorityYear="2019" box="[151,475,475,501]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="siberiensis">
<emphasis id="6887BD20FFA8983601C7AD86738FFE24" box="[151,170,475,501]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA8983601EAAD867217FE24" box="[186,306,475,501]" italics="true" pageId="8" pageNumber="494">siberiensis</emphasis>
<bibRefCitation id="3E621CC3FFA89836006BAD8672FEFE24" author="Joharchi, O &amp; Trach, V. A." box="[315,475,475,501]" pageId="8" pageNumber="494" pagination="486 - 494" refId="ref14677" refString="Joharchi, O &amp; Trach, V. A. (2019) A new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Ukraine. Zootaxa, 4647 (1), 486 - 494. https: // doi. org / 10.11646 / zootaxa. 4647.1.30" type="journal article" year="2019">Joharchi 2019</bibRefCitation>
</taxonomicName>
(in
<bibRefCitation id="3E621CC3FFA89836035DAD8671D9FE24" author="Joharchi, O &amp; Trach, V. A." box="[525,764,475,501]" pageId="8" pageNumber="494" pagination="486 - 494" refId="ref14677" refString="Joharchi, O &amp; Trach, V. A. (2019) A new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Ukraine. Zootaxa, 4647 (1), 486 - 494. https: // doi. org / 10.11646 / zootaxa. 4647.1.30" type="journal article" year="2019">
Joharchi
<emphasis id="6887BD20FFA898360325AD86718CFE24" box="[629,681,475,501]" italics="true" pageId="8" pageNumber="494">et al</emphasis>
., 2019
</bibRefCitation>
),
<taxonomicName id="9DF31AB1FFA898360242AD86777BFE27" authority="(Shcherbak, 1971 b)" baseAuthorityName="Shcherbak" baseAuthorityYear="1971" box="[786,1118,475,502]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="6887BD20FFA898360242AD867000FE24" box="[786,805,475,501]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA898360265AD867056FE25" box="[821,883,475,500]" italics="true" pageId="8" pageNumber="494">mixta</emphasis>
(
<bibRefCitation id="3E621CC3FFA8983602D4AD867770FE27" author="Shcherbak, G. I." box="[900,1109,475,502]" pageId="8" pageNumber="494" pagination="20 - 30" refId="ref16506" refString="Shcherbak, G. I. (1971 b) Species of Gamasoidea mites from the genus Hypoaspis Can. (Parasitiformes, Gamasoidea) new for the USSR fauna. Collection of papers from the Zoological Museum of the Institute of Zoology of SSR Ukraine Academy of Sciences, 34, 20 - 30. [in Ukrainian]" type="journal article" year="1971">Shcherbak, 1971b</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="9DF31AB1FFA89836053CAD867651FE24" authority="(Karg, 1988)" baseAuthorityName="Karg" baseAuthorityYear="1988" box="[1132,1396,475,501]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="pumili">
<emphasis id="6887BD20FFA89836053CAD86775AFE24" box="[1132,1151,475,501]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA8983605DFAD8677F2FE24" box="[1167,1239,475,501]" italics="true" pageId="8" pageNumber="494">pumili</emphasis>
(
<bibRefCitation id="3E621CC3FFA8983605B8AD867649FE24" author="Karg, W." box="[1256,1388,475,501]" pageId="8" pageNumber="494" pagination="509 - 525" refId="ref15324" refString="Karg, W. (1988) Die Arten der Raubmilbengattung Cosmolaelaps Berlese, 1903 (Gattung Hypoaspis Canestrini, 1884, Acarina, Parasitiformes). Zoologische Jahrbucher Abteilung fur Systematik, Okologie und Geographie der Tiere, 115, 509 - 525." type="journal article" year="1988">Karg, 1988</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="9DF31AB1FFA8983604D3AD86725EFDC8" authority="(Karg, 1987)" baseAuthorityName="Karg" baseAuthorityYear="1987" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="lingua">
<emphasis id="6887BD20FFA8983604D3AD8676B3FE24" box="[1411,1430,475,501]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA8983601C7ADA273FAFDC8" box="[151,223,511,537]" italics="true" pageId="8" pageNumber="494">lingua</emphasis>
(
<bibRefCitation id="3E621CC3FFA8983601BFADA27256FDC8" author="Karg, W." box="[239,371,511,537]" pageId="8" pageNumber="494" pagination="289 - 302" refId="ref15285" refString="Karg, W. (1987) Neue Raubmilbenarten der Gattung Hypoaspis Can. 1884 (Acari: Parasitiformes). Zoologische Jahrbucher Abteilung fur Systematik, Okologie und Geographie der Tiere, 114, 289 - 302." type="journal article" year="1987">Karg, 1987</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="9DF31AB1FFA8983600D8ADA271B4FDCB" authority="(Ma, 1996)" baseAuthorityName="Ma" baseAuthorityYear="1996" box="[392,657,511,538]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="sungaris">
<emphasis id="6887BD20FFA8983600D8ADA272BEFDC8" box="[392,411,511,537]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA8983600F9ADA2712CFDC9" box="[425,521,511,536]" italics="true" pageId="8" pageNumber="494">sungaris</emphasis>
(
<bibRefCitation id="3E621CC3FFA89836034AADA271ACFDCB" author="Ma, L. - M." box="[538,649,511,538]" pageId="8" pageNumber="494" pagination="48 - 54" refId="ref15688" refString="Ma, L. - M. (1996) Three new species of the genus Hypoaspis from Jilin Province, China (Acari: Laelapidae). Acta Zootaxonomica Sinica, 21, 48 - 54. [in Chinese]" type="journal article" year="1996">Ma, 1996</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="9DF31AB1FFA89836039AADA2777CFDC8" authority="(Bei et al., 2003)" baseAuthorityName="Bei" baseAuthorityYear="2003" box="[714,1113,511,537]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="shenyangensis">
<emphasis id="6887BD20FFA89836039AADA271F8FDC8" box="[714,733,511,537]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA8983603BCADA270ABFDC8" box="[748,910,511,537]" italics="true" pageId="8" pageNumber="494">shenyangensis</emphasis>
(
<bibRefCitation id="3E621CC3FFA8983602CEADA27774FDC8" author="Bei, N. - X. &amp; Shi, C. - M. &amp; Yin, S. - G." box="[926,1105,511,537]" pageId="8" pageNumber="494" pagination="648 - 650" refId="ref13627" refString="Bei, N. - X., Shi, C. - M. &amp; Yin, S. - G. (2003) A new species of the genus Hypoaspis from China (Acari, Laelapidae). Acta Zootaxonomica Sinica, 28, 648 - 650. [in Chinese]" type="journal article" year="2003">
Bei
<emphasis id="6887BD20FFA89836029FADA27724FDC8" box="[975,1025,511,537]" italics="true" pageId="8" pageNumber="494">et al</emphasis>
., 2003
</bibRefCitation>
)
</taxonomicName>
in the shape of dorsal setae, which are slightly swollen near the base and pointed at tip (knife-shaped) (see
<figureCitation id="C2C87DB7FFA898360552AE7E7744FDEC" box="[1026,1121,547,573]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="8" pageNumber="494">Figure 8</figureCitation>
), with all dorsal setae of the same shape and sub-equal in length (at most reaching base of following setae), except setae
<emphasis id="6887BD20FFA8983605CEAE1A7791FDB0" box="[1182,1204,583,609]" italics="true" pageId="8" pageNumber="494">j1</emphasis>
and
<emphasis id="6887BD20FFA8983605BAAE1A762DFDB0" box="[1258,1288,583,609]" italics="true" pageId="8" pageNumber="494">Z5</emphasis>
.
</paragraph>
<paragraph id="5A4C6132FFA898360197AE3670F6F96C" blockId="8.[151,1437,151,1725]" pageId="8" pageNumber="494">
<taxonomicName id="9DF31AB1FFA898360197AE3672CAFD54" authorityName="Joharchi" authorityYear="2022" box="[199,495,619,645]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="lasiophilus">
<emphasis id="6887BD20FFA898360197AE3672CAFD54" box="[199,495,619,645]" italics="true" pageId="8" pageNumber="494">Cosmolaelaps lasiophilus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="9DF31AB1FFA8983603C7AE367024FD55" authorityName="sensu Evans &amp; Till" authorityYear="1966" baseAuthorityName="Michael" baseAuthorityYear="1891" box="[663,769,619,645]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="vacua">
<emphasis id="6887BD20FFA8983603C7AE36718FFD54" box="[663,682,619,645]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA8983603EEAE367024FD55" box="[702,769,619,644]" italics="true" pageId="8" pageNumber="494">vacua</emphasis>
</taxonomicName>
by lacking spatulate setae on legs, most setae on legs conspicuously thickened and some inserted on small tubercles, while in
<taxonomicName id="9DF31AB1FFA898360282AED27712FD79" authorityName="sensu Evans &amp; Till" authorityYear="1966" baseAuthorityName="Michael" baseAuthorityYear="1891" box="[978,1079,655,681]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="vacua">
<emphasis id="6887BD20FFA898360282AED270C0FD78" box="[978,997,655,681]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA8983602A4AED27712FD79" box="[1012,1079,655,680]" italics="true" pageId="8" pageNumber="494">vacua</emphasis>
</taxonomicName>
two spatulate setae are present on femora II and IV.
<bibRefCitation id="3E621CC3FFA8983600D9AEEE714BFD1F" author="Evans, G. O. &amp; Till, W. M." box="[393,622,691,718]" pageId="8" pageNumber="494" pagination="107 - 370" refId="ref14300" refString="Evans, G. O. &amp; Till, W. M. (1966) Studies on the British Dermanyssidae (Acari: Mesostigmata). Part II. Classification. Bulletin of the British Museum (Natural History) Zoology, 14, 107 - 370." type="journal article" year="1966">Evans &amp; Till (1966)</bibRefCitation>
did not provide any information about the shape of the leg setae for
<taxonomicName id="9DF31AB1FFA8983604D3AEEE73FFFD21" authorityName="sensu Evans &amp; Till" authorityYear="1966" baseAuthorityName="Michael" baseAuthorityYear="1891" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="vacua">
<emphasis id="6887BD20FFA8983604D3AEEE76B3FD1C" box="[1411,1430,691,717]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA8983601C7AE8A73FFFD21" box="[151,218,727,752]" italics="true" pageId="8" pageNumber="494">vacua</emphasis>
</taxonomicName>
, but according to the original description (
<bibRefCitation id="3E621CC3FFA8983603E4AE8A707DFD20" author="Michael, A. D." box="[692,856,727,753]" pageId="8" pageNumber="494" pagination="638 - 654" refId="ref15960" refString="Michael, A. D. (1891) On the association of gamasids with ants. Proceedings of the Zoological Society of London, 4, 638 - 654." type="journal article" year="1891">Michael, 1891</bibRefCitation>
) of
<taxonomicName id="9DF31AB1FFA8983602D5AE8A7748FD21" authorityName="sensu Evans &amp; Till" authorityYear="1966" baseAuthorityName="Michael" baseAuthorityYear="1891" box="[901,1133,727,753]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="vacua">
<emphasis id="6887BD20FFA8983602D5AE8A7748FD21" box="[901,1133,727,753]" italics="true" pageId="8" pageNumber="494">Cosmolaelaps vacua</emphasis>
</taxonomicName>
, femora II and IV bear two spatulate setae (
<bibRefCitation id="3E621CC3FFA89836001DAEA672F5FCC4" author="Karg, W." box="[333,464,763,789]" pageId="8" pageNumber="494" pagination="213 - 232" refId="ref15239" refString="Karg, W. (1981) Die Untergattung Cosmolaelaps Berlese, 1903 der Raubmilbengattung Hypoaspis Canestrini, 1884. Mitteilungen aus dem Zoologischen Museum in Berlin, 57, 213 - 232. https: // doi. org / 10.1002 / mmnz. 19810570203" type="journal article" year="1981">Karg, 1981</bibRefCitation>
,
<figureCitation id="C2C87DB7FFA89836008EAEA67114FCC4" box="[478,561,763,789]" captionStart="FIGURES 8–12" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1397,196,1901]" captionTargetId="figure-18@4.[167,1419,181,1919]" captionTargetPageId="4" captionText="FIGURES 8–12. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 8, dorsal idiosoma, enlarged section not to scale; 9, ventral idiosoma; 10, subcapitulum; 11, epistome; 12, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530964" httpUri="https://zenodo.org/record/6530964/files/figure.png" pageId="8" pageNumber="494">Fig. 8a</figureCitation>
, also illustrated this character). It differs from
<taxonomicName id="9DF31AB1FFA89836051BAEA677FBFCC4" authorityName="Zeman" authorityYear="1982" box="[1099,1246,763,789]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="thysanifer">
<emphasis id="6887BD20FFA89836051BAEA6777BFCC4" box="[1099,1118,763,789]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA89836053EAEA677FBFCC4" box="[1134,1246,763,789]" italics="true" pageId="8" pageNumber="494">thysanifer</emphasis>
</taxonomicName>
by setae
<emphasis id="6887BD20FFA89836041DAEA6764EFCC4" box="[1357,1387,763,789]" italics="true" pageId="8" pageNumber="494">Z5</emphasis>
and some opisthogastric setae thickened, versus setae
<emphasis id="6887BD20FFA898360399AF4271C3FCE8" box="[713,742,799,825]" italics="true" pageId="8" pageNumber="494">Z5</emphasis>
brush-like and all opisthogastric setae simple and setiform in
<taxonomicName id="9DF31AB1FFA8983601C7AF1E720FFC8C" authorityName="Zeman" authorityYear="1982" box="[151,298,835,861]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="thysanifer">
<emphasis id="6887BD20FFA8983601C7AF1E738FFC8C" box="[151,170,835,861]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA8983601EAAF1E720FFC8C" box="[186,298,835,861]" italics="true" pageId="8" pageNumber="494">thysanifer</emphasis>
</taxonomicName>
. It differs from
<taxonomicName id="9DF31AB1FFA8983600B4AF1E7142FC8D" baseAuthorityName="Ma" baseAuthorityYear="1996" box="[484,615,835,861]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="sungaris">
<emphasis id="6887BD20FFA8983600B4AF1E72D2FC8C" box="[484,503,835,861]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA898360357AF1E7142FC8D" box="[519,615,835,860]" italics="true" pageId="8" pageNumber="494">sungaris</emphasis>
</taxonomicName>
and
<taxonomicName id="9DF31AB1FFA8983603F2AF1E7043FC8C" box="[674,870,835,861]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="shenyangensis">
<emphasis id="6887BD20FFA8983603F2AF1E7190FC8C" box="[674,693,835,861]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA898360394AF1E7043FC8C" box="[708,870,835,861]" italics="true" pageId="8" pageNumber="494">shenyangensis</emphasis>
</taxonomicName>
by its much smaller size of body (389–406 long, 256–265wide), versus body size much larger in both of
<taxonomicName id="9DF31AB1FFA89836025AAF3A70AEFC51" baseAuthorityName="Ma" baseAuthorityYear="1996" box="[778,907,871,897]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="sungaris">
<emphasis id="6887BD20FFA89836025AAF3A7038FC50" box="[778,797,871,897]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA89836027BAF3A70AEFC51" box="[811,907,871,896]" italics="true" pageId="8" pageNumber="494">sungaris</emphasis>
</taxonomicName>
and
<taxonomicName id="9DF31AB1FFA898360293AF3A77A3FC50" box="[963,1158,871,897]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="shenyangensis">
<emphasis id="6887BD20FFA898360293AF3A70F3FC50" box="[963,982,871,897]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA8983602B4AF3A77A3FC50" box="[996,1158,871,897]" italics="true" pageId="8" pageNumber="494">shenyangensis</emphasis>
</taxonomicName>
(566 long × 383 wide in
<taxonomicName id="9DF31AB1FFA8983601C7AFD67232FC75" baseAuthorityName="Ma" baseAuthorityYear="1996" box="[151,279,907,933]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="sungaris">
<emphasis id="6887BD20FFA8983601C7AFD6738FFC74" box="[151,170,907,933]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA8983601E7AFD67232FC75" box="[183,279,907,932]" italics="true" pageId="8" pageNumber="494">sungaris</emphasis>
</taxonomicName>
, and 567 long × 413 wide in
<taxonomicName id="9DF31AB1FFA89836030AAFD671D9FC74" box="[602,764,907,933]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="shenyangensis">
<emphasis id="6887BD20FFA89836030AAFD671D9FC74" box="[602,764,907,933]" italics="true" pageId="8" pageNumber="494">shenyangensis</emphasis>
</taxonomicName>
) and also differs from
<taxonomicName id="9DF31AB1FFA8983602A8AFD6775DFC75" baseAuthorityName="Ma" baseAuthorityYear="1996" box="[1016,1144,907,933]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="sungaris">
<emphasis id="6887BD20FFA8983602A8AFD6772EFC74" box="[1016,1035,907,933]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA898360548AFD6775DFC75" box="[1048,1144,907,932]" italics="true" pageId="8" pageNumber="494">sungaris</emphasis>
</taxonomicName>
by its longer dorsal shield setae (reaching base of following setae), versus dorsal shield setae much shorter in
<taxonomicName id="9DF31AB1FFA898360507AFF277FFFC19" baseAuthorityName="Ma" baseAuthorityYear="1996" box="[1111,1242,943,969]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="sungaris">
<emphasis id="6887BD20FFA898360507AFF2774FFC18" box="[1111,1130,943,969]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA89836052AAFF277FFFC19" box="[1146,1242,943,968]" italics="true" pageId="8" pageNumber="494">sungaris</emphasis>
</taxonomicName>
(barely reaching more than half of distance of following setae in series).
<taxonomicName id="9DF31AB1FFA898360251AF8E7707FC3C" authorityName="Joharchi" authorityYear="2022" box="[769,1058,979,1005]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="lasiophilus">
<emphasis id="6887BD20FFA898360251AF8E7707FC3C" box="[769,1058,979,1005]" italics="true" pageId="8" pageNumber="494">Cosmolaelaps lasiophilus</emphasis>
</taxonomicName>
can be readily distinguished from
<taxonomicName id="9DF31AB1FFA8983601C7AFAA7279FBC0" authorityName="Ryke" authorityYear="1963" box="[151,348,1015,1041]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="transvaalensis">
<emphasis id="6887BD20FFA8983601C7AFAA738FFBC0" box="[151,170,1015,1041]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA8983601EAAFAA7279FBC0" box="[186,348,1015,1041]" italics="true" pageId="8" pageNumber="494">transvaalensis</emphasis>
</taxonomicName>
by some opisthogastric setae thickened and most setae on legs conspicuously thickened (some inserted on small tubercles), versus setae on both opisthogastric cuticle and legs simple in
<taxonomicName id="9DF31AB1FFA8983605F2A8467642FBE4" authorityName="Ryke" authorityYear="1963" box="[1186,1383,1051,1077]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="transvaalensis">
<emphasis id="6887BD20FFA8983605F2A8467790FBE4" box="[1186,1205,1051,1077]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA898360595A8467642FBE4" box="[1221,1383,1051,1077]" italics="true" pageId="8" pageNumber="494">transvaalensis</emphasis>
</taxonomicName>
(see
<bibRefCitation id="3E621CC3FFA8983601C7A862723EFB8B" author="Ryke, P. A. J." box="[151,283,1087,1114]" pageId="8" pageNumber="494" pagination="1 - 15" refId="ref16407" refString="Ryke, P. A. J. (1963) Some free-living Hypoaspidinae (Acari: Mesostigmata) from South Africa. Revista de Biologia, 5, 1 - 15." type="journal article" year="1963">Ryke, 1963</bibRefCitation>
). It differs from
<taxonomicName id="9DF31AB1FFA898360082A862711FFB88" baseAuthorityName="Karg" baseAuthorityYear="1988" box="[466,570,1087,1113]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="pumili">
<emphasis id="6887BD20FFA898360082A86272C0FB88" box="[466,485,1087,1113]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA8983600A2A862711FFB88" box="[498,570,1087,1113]" italics="true" pageId="8" pageNumber="494">pumili</emphasis>
</taxonomicName>
because setae
<emphasis id="6887BD20FFA89836038EA86271D9FB88" box="[734,764,1087,1113]" italics="true" pageId="8" pageNumber="494">Z5</emphasis>
are thickened, shorter than
<emphasis id="6887BD20FFA898360560A862776EFB88" box="[1072,1099,1087,1113]" italics="true" pageId="8" pageNumber="494">J5</emphasis>
and legs I, IV are shorter than length of idiosoma, while in
<taxonomicName id="9DF31AB1FFA898360082A83E711CFBAC" baseAuthorityName="Karg" baseAuthorityYear="1988" box="[466,569,1123,1149]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="pumili">
<emphasis id="6887BD20FFA898360082A83E72C0FBAC" box="[466,485,1123,1149]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA8983600A1A83E711CFBAC" box="[497,569,1123,1149]" italics="true" pageId="8" pageNumber="494">pumili</emphasis>
</taxonomicName>
setae
<emphasis id="6887BD20FFA89836032BA83E71BCFBAC" box="[635,665,1123,1149]" italics="true" pageId="8" pageNumber="494">Z5</emphasis>
setiform, longer than
<emphasis id="6887BD20FFA8983602DBA83E7080FBAC" box="[907,933,1123,1149]" italics="true" pageId="8" pageNumber="494">J5</emphasis>
and legs I, IV longer than length of idiosoma.
<taxonomicName id="9DF31AB1FFA8983601C7A8DA7211FB70" authorityName="Joharchi" authorityYear="2022" box="[151,308,1159,1185]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="lasiophilus">
<emphasis id="6887BD20FFA8983601C7A8DA738FFB70" box="[151,170,1159,1185]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA8983601E9A8DA7211FB70" box="[185,308,1159,1185]" italics="true" pageId="8" pageNumber="494">lasiophilus</emphasis>
</taxonomicName>
can be easily distinguished from
<taxonomicName id="9DF31AB1FFA8983603E2A8DA7017FB70" baseAuthorityName="Berlese" baseAuthorityYear="1905" box="[690,818,1159,1185]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="6887BD20FFA8983603E2A8DA71E0FB70" box="[690,709,1159,1185]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA898360383A8DA7017FB70" box="[723,818,1159,1185]" italics="true" pageId="8" pageNumber="494">robustus</emphasis>
</taxonomicName>
by the smaller size of the genital shield (there are two pairs of setae between genital and anal shields) and presternal area lightly sclerotised (with a few transverse curved lines and merged to the sternal shield), while genital shield expanded posteriorly in
<taxonomicName id="9DF31AB1FFA898360516A89277E3FB38" baseAuthorityName="Berlese" baseAuthorityYear="1905" box="[1094,1222,1231,1257]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="6887BD20FFA898360516A892777CFB38" box="[1094,1113,1231,1257]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA898360537A89277E3FB38" box="[1127,1222,1231,1257]" italics="true" pageId="8" pageNumber="494">robustus</emphasis>
</taxonomicName>
, setae
<emphasis id="6887BD20FFA898360444A892761BFB38" box="[1300,1342,1231,1257]" italics="true" pageId="8" pageNumber="494">Zv1</emphasis>
and
<emphasis id="6887BD20FFA898360425A89276B9FB38" box="[1397,1436,1231,1257]" italics="true" pageId="8" pageNumber="494">Jv1</emphasis>
inserted on lateral margins of shield (in addition of
<emphasis id="6887BD20FFA89836038AA8AE71DEFADC" box="[730,763,1267,1293]" italics="true" pageId="8" pageNumber="494">st5</emphasis>
) and two triangular presternal plates present in
<taxonomicName id="9DF31AB1FFA898360445A8AE76B0FADC" baseAuthorityName="Berlese" baseAuthorityYear="1905" box="[1301,1429,1267,1293]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="6887BD20FFA898360445A8AE760DFADC" box="[1301,1320,1267,1293]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA898360466A8AE76B0FADC" box="[1334,1429,1267,1293]" italics="true" pageId="8" pageNumber="494">robustus</emphasis>
</taxonomicName>
.
<taxonomicName id="9DF31AB1FFA8983601C7A94A7293FAE0" authorityName="Joharchi" authorityYear="2022" box="[151,438,1303,1329]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="lasiophilus">
<emphasis id="6887BD20FFA8983601C7A94A7293FAE0" box="[151,438,1303,1329]" italics="true" pageId="8" pageNumber="494">Cosmolaelaps lasiophilus</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="9DF31AB1FFA89836038FA94A7062FAE0" baseAuthorityName="Karg" baseAuthorityYear="1987" box="[735,839,1303,1329]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="lingua">
<emphasis id="6887BD20FFA89836038FA94A71D7FAE0" box="[735,754,1303,1329]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA8983603AFA94A7062FAE0" box="[767,839,1303,1329]" italics="true" pageId="8" pageNumber="494">lingua</emphasis>
</taxonomicName>
by its small body size (389–406 long, 256–265 wide), thick and short opisthogastric setae and presternal area lightly sclerotised (anterior margin of sternal shield hardly conspicuous), versus body larger (480–500 long, 300–330 wide), opisthogastric setae simple (setiform) and long, sclerotised platelets absent and presternal plates joined to sternal shield in
<taxonomicName id="9DF31AB1FFA8983602B6A9DE7775FA4C" baseAuthorityName="Karg" baseAuthorityYear="1987" box="[998,1104,1411,1437]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="lingua">
<emphasis id="6887BD20FFA8983602B6A9DE70DCFA4C" box="[998,1017,1411,1437]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA898360558A9DE7775FA4C" box="[1032,1104,1411,1437]" italics="true" pageId="8" pageNumber="494">lingua</emphasis>
</taxonomicName>
.
<taxonomicName id="9DF31AB1FFA89836050FA9DE76A4FA4C" authorityName="Joharchi" authorityYear="2022" box="[1119,1409,1411,1437]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="lasiophilus">
<emphasis id="6887BD20FFA89836050FA9DE76A4FA4C" box="[1119,1409,1411,1437]" italics="true" pageId="8" pageNumber="494">Cosmolaelaps lasiophilus</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="9DF31AB1FFA89836001BA9FA72C0FA10" authorityName="Joharchi" authorityYear="2019" box="[331,485,1447,1473]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="siberiensis">
<emphasis id="6887BD20FFA89836001BA9FA727BFA10" box="[331,350,1447,1473]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA89836003DA9FA72C0FA10" box="[365,485,1447,1473]" italics="true" pageId="8" pageNumber="494">siberiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="9DF31AB1FFA89836034EA9FA715BFA11" baseAuthorityName="Shcherbak" baseAuthorityYear="1971" box="[542,638,1447,1473]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="6887BD20FFA89836034EA9FA7114FA10" box="[542,561,1447,1473]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA898360310A9FA715BFA11" box="[576,638,1447,1472]" italics="true" pageId="8" pageNumber="494">mixta</emphasis>
</taxonomicName>
, but can be readily distinguished from these two species by its longer dorsal setae, some of them reach to the base of next setae, while in both
<taxonomicName id="9DF31AB1FFA898360281A996774EFA34" authorityName="Joharchi" authorityYear="2019" box="[977,1131,1483,1509]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="siberiensis">
<emphasis id="6887BD20FFA898360281A99670C1FA34" box="[977,996,1483,1509]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA8983602A3A996774EFA34" box="[1011,1131,1483,1509]" italics="true" pageId="8" pageNumber="494">siberiensis</emphasis>
</taxonomicName>
<emphasis id="6887BD20FFA898360523A99677B8FA34" box="[1139,1181,1483,1509]" italics="true" pageId="8" pageNumber="494">and</emphasis>
<taxonomicName id="9DF31AB1FFA8983605F5A9967620FA35" baseAuthorityName="Shcherbak" baseAuthorityYear="1971" box="[1189,1285,1483,1509]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="6887BD20FFA8983605F5A996779DFA34" box="[1189,1208,1483,1509]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA898360597A9967620FA35" box="[1223,1285,1483,1508]" italics="true" pageId="8" pageNumber="494">mixta</emphasis>
</taxonomicName>
dorsal shield setae considerably shorter and none of them reach base of next. Overall
<taxonomicName id="9DF31AB1FFA898360299A9B27740F9D8" authorityName="Joharchi" authorityYear="2022" box="[969,1125,1519,1545]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="lasiophilus">
<emphasis id="6887BD20FFA898360299A9B270F9F9D8" box="[969,988,1519,1545]" italics="true" pageId="8" pageNumber="494">C</emphasis>
.
<emphasis id="6887BD20FFA8983602BAA9B27740F9D8" box="[1002,1125,1519,1545]" italics="true" pageId="8" pageNumber="494">lasiophilus</emphasis>
</taxonomicName>
can be easily distinguished from all other congeners by the following combination of characters: (1) dorsal shield bearing 39 pairs of relatively long setae (some reach base of next posterior setae in series),
<emphasis id="6887BD20FFA898360200AA6A7043F980" box="[848,870,1591,1617]" italics="true" pageId="8" pageNumber="494">j1</emphasis>
,
<emphasis id="6887BD20FFA898360225AA6A70ABF980" box="[885,910,1591,1617]" italics="true" pageId="8" pageNumber="494">z1</emphasis>
and
<emphasis id="6887BD20FFA898360295AA6A70C6F980" box="[965,995,1591,1617]" italics="true" pageId="8" pageNumber="494">Z5</emphasis>
simple,
<emphasis id="6887BD20FFA898360514AA6A7747F980" box="[1092,1122,1591,1617]" italics="true" pageId="8" pageNumber="494">Z5</emphasis>
shorter than
<emphasis id="6887BD20FFA8983605A6AA6A7634F980" box="[1270,1297,1591,1617]" italics="true" pageId="8" pageNumber="494">J5</emphasis>
(ratio of J5/ Z5 length ≃ 1.5); (2) post-anal seta thickened; (3) almost all opisthogastric setae thickened and arising on minute sclerotised platelet; (4) fixed digit of chelicera with six various size of teeth; (5) legs without spatulate setae but most setae conspicuously thickened and some inserted on small tubercles.
</paragraph>
</subSubSection>
</treatment>
</document>

458
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@ -0,0 +1,458 @@
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<mods:title id="9CB1D51572B46A59EDE71101786EDEA1">New species and new records of Cosmolaelaps Berlese (Acari: Laelapidae) from Russia, with a review of the Russian species of the genus</mods:title>
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<mods:affiliation id="E2973162971C9E9EFAE72C11133A20FF">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Semakova Str. 10, 625003 Tyumen, Russia. &amp; Cukurova University, Agricultural Faculty, Department of Plant Protection, Acarology Laboratory, 01330, Adana, Turkey. https: // orcid. org / 0000 - 0002 - 2741 - 4946 (Joharchi) https: // orcid. org / 0000 - 0002 - 1412 - 1554 (Döker) https: // orcid. org / 0000 - 0003 - 2831 - 7213 (Khaustov)</mods:affiliation>
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<treatment id="D25AD024FFA8983201C7AB5372D3FCC4" ID-DOI="http://doi.org/10.5281/zenodo.6530966" ID-GBIF-Taxon="195277957" ID-Zenodo-Dep="6530966" LSID="urn:lsid:plazi:treatment:D25AD024FFA8983201C7AB5372D3FCC4" httpUri="http://treatment.plazi.org/id/D25AD024FFA8983201C7AB5372D3FCC4" lastPageId="12" lastPageNumber="498" pageId="8" pageNumber="494">
<subSubSection id="12E932B9FFA8983601C7AB537159F8F8" box="[151,636,1806,1833]" pageId="8" pageNumber="494" type="nomenclature">
<paragraph id="5A4C6132FFA8983601C7AB537159F8F8" blockId="8.[151,636,1806,1869]" box="[151,636,1806,1833]" pageId="8" pageNumber="494">
<heading id="0104D65EFFA8983601C7AB537159F8F8" bold="true" box="[151,636,1806,1833]" fontSize="11" level="1" pageId="8" pageNumber="494" reason="1">
<emphasis id="6887BD20FFA8983601C7AB537159F8F8" bold="true" box="[151,636,1806,1833]" pageId="8" pageNumber="494">
<taxonomicName id="9DF31AB1FFA8983601C7AB537132F8F9" authority="Joharchi" authorityName="Joharchi" authorityYear="2022" box="[151,535,1806,1833]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="latisetis" status="sp. nov.">
<emphasis id="6887BD20FFA8983601C7AB537284F8F8" bold="true" box="[151,417,1806,1833]" italics="true" pageId="8" pageNumber="494">Cosmolaelaps latisetis</emphasis>
Joharchi
</taxonomicName>
<taxonomicNameLabel id="73B4005BFFA89836034FAB527159F8F8" box="[543,636,1807,1833]" pageId="8" pageNumber="494" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="12E932B9FFA8983601C7AB6E726EF89C" box="[151,331,1842,1869]" pageId="8" pageNumber="494" type="description">
<paragraph id="5A4C6132FFA8983601C7AB6E726EF89C" blockId="8.[151,636,1806,1869]" box="[151,331,1842,1869]" pageId="8" pageNumber="494">
(
<figureCitation id="C2C87DB7FFA8983601CFAB6E7266F89C" box="[159,323,1842,1869]" captionStart-0="FIGURES 25–26" captionStart-1="FIGURES 27–29" captionStart-2="FIGURES 30–40" captionStart-3="FIGURE 41–44" captionStartId-0="9.[151,264,1943,1967]" captionStartId-1="10.[151,264,1468,1492]" captionStartId-2="13.[151,264,1874,1898]" captionStartId-3="14.[151,250,1943,1967]" captionTargetBox-0="[193,1391,190,1899]" captionTargetBox-1="[171,1417,193,1432]" captionTargetBox-2="[214,1374,184,1849]" captionTargetBox-3="[183,1406,196,1906]" captionTargetId-0="figure-19@9.[179,1408,181,1919]" captionTargetId-1="figure-18@10.[151,1436,181,1444]" captionTargetId-2="figure-19@13.[212,1375,181,1851]" captionTargetId-3="figure-18@14.[175,1413,181,1919]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="13" captionTargetPageId-3="14" captionText-0="FIGURES 25–26. Cosmolaelaps latisetis Joharchi sp. nov., female.25, dorsal idiosoma; 26, ventral idiosoma. Enlarged sections not to scale." captionText-1="FIGURES 27–29. Cosmolaelaps latisetis Joharchi sp. nov., female. 27, subcapitulum; 28, epistome; 29, chelicera." captionText-2="FIGURES 30–40. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." captionText-3="FIGURE 41–44. Cosmolaelaps latisetis Joharchi sp. nov., female. 41, leg I (trochanter-tibia); 42, leg II; 43, leg III; 44, leg IV." figureDoi-0="http://doi.org/10.5281/zenodo.6530976" figureDoi-1="http://doi.org/10.5281/zenodo.6530978" figureDoi-2="http://doi.org/10.5281/zenodo.6531011" figureDoi-3="http://doi.org/10.5281/zenodo.6530984" httpUri-0="https://zenodo.org/record/6530976/files/figure.png" httpUri-1="https://zenodo.org/record/6530978/files/figure.png" httpUri-2="https://zenodo.org/record/6531011/files/figure.png" httpUri-3="https://zenodo.org/record/6530984/files/figure.png" pageId="8" pageNumber="494">Figures 25–44</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="12E932B9FFA8983601C7AB26776DF80C" pageId="8" pageNumber="494" type="materials_examined">
<paragraph id="5A4C6132FFA8983601C7AB26776DF80C" blockId="8.[151,1437,1914,2014]" pageId="8" pageNumber="494">
<emphasis id="6887BD20FFA8983601C7AB267261F845" bold="true" box="[151,324,1914,1941]" pageId="8" pageNumber="494">Type material.</emphasis>
<materialsCitation id="EA9B6B6FFFA898360019AB26728CF80C" ID-GBIF-Occurrence="3763889303" collectedFrom="in the nest of Lasius flavus (Fabricius, 1782) (Hymenoptera: Formicidae)" collectingDate="2021-08-11" collectionCode="TUMZ" collectorName="O. Joharchi" country="Russia" latitude="46.71375" location="on the coast of the Sea of Japan (East Sea)" longLatPrecision="10" longitude="141.88722" pageId="8" pageNumber="494" specimenCount="1" specimenCount-female="1" stateProvince="Sakhalin Island" typeStatus="holotype">
<typeStatus id="8548DF90FFA898360019AB267291F844" box="[329,436,1915,1941]" pageId="8" pageNumber="494" type="holotype">Holotype</typeStatus>
,
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,
<location id="5F2C37E9FFA898360347AB2670CCF844" LSID="urn:lsid:plazi:treatment:D25AD024FFA8983201C7AB5372D3FCC4:5F2C37E9FFA898360347AB2670CCF844" box="[535,1001,1915,1941]" country="Russia" latitude="46.71375" longLatPrecision="10" longitude="141.88722" name="on the coast of the Sea of Japan (East Sea)" pageId="8" pageNumber="494" stateProvince="Sakhalin Island">on the coast of the Sea of Japan (East Sea)</location>
,
<collectingRegion id="9837AFD0FFA8983602A3AB267787F844" box="[1011,1186,1915,1941]" pageId="8" pageNumber="494">Sakhalin Island</collectingRegion>
,
<collectingCountry id="22E421A2FFA8983605FCAB2677DCF844" box="[1196,1273,1915,1941]" name="Russia" pageId="8" pageNumber="494">Russia</collectingCountry>
,
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<geoCoordinate id="3FC707F5FFA8983601C7ABC3720DF868" box="[151,296,1950,1977]" degrees="141" direction="east" minutes="53" orientation="longitude" pageId="8" pageNumber="494" precision="15" seconds="14" value="141.88722">141°53'14&quot;E</geoCoordinate>
,
<date id="2E4D47F2FFA898360068ABC272D1F868" box="[312,500,1951,1977]" pageId="8" pageNumber="494" value="2021-08-11">
<collectingDate id="3E09BE1AFFA898360068ABC272D1F868" box="[312,500,1951,1977]" pageId="8" pageNumber="494" value="2021-08-11">11 August 2021</collectingDate>
</date>
,
<collectorName id="F70604E4FFA898360353ABC271ADF868" box="[515,648,1951,1977]" pageId="8" pageNumber="494">O. Joharchi</collectorName>
coll.,
<collectedFrom id="E9EF22CFFFA898360387ABC27205F80F" pageId="8" pageNumber="494">
in the nest of
<taxonomicName id="9DF31AB1FFA8983602D0ABC277CDF868" authority="(Fabricius, 1782)" baseAuthorityName="Fabricius" baseAuthorityYear="1782" box="[896,1256,1951,1977]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="flavus">
<emphasis id="6887BD20FFA8983602D0ABC27732F868" box="[896,1047,1951,1977]" italics="true" pageId="8" pageNumber="494">Lasius flavus</emphasis>
(Fabricius, 1782)
</taxonomicName>
(
<taxonomicName id="9DF31AB1FFA8983605AAABC276BCF868" box="[1274,1433,1951,1977]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="494" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
:
<taxonomicName id="9DF31AB1FFA8983601C7AB9E723CF80F" box="[151,281,1987,2014]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="494" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
)
</collectedFrom>
(in
<collectionCode id="3CE2F9F7FFA89836001DAB9E7287F80C" box="[333,418,1987,2013]" pageId="8" pageNumber="494">TUMZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="EA9B6B6FFFA8983600E3AB9E7760F80C" ID-GBIF-Occurrence="3763889302" box="[435,1093,1987,2013]" pageId="8" pageNumber="494" typeStatus="paratype">
<typeStatus id="8548DF90FFA8983600E3AB9E7101F80C" box="[435,548,1987,2013]" pageId="8" pageNumber="494" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="4CF5AABBFFA89836037FAB9E719FF80C" box="[559,698,1987,2013]" pageId="8" pageNumber="494" type="female">two females</specimenCount>
, same data as
<typeStatus id="8548DF90FFA89836020AAB9E7099F80C" box="[858,956,1987,2013]" pageId="8" pageNumber="494" type="holotype">holotype</typeStatus>
(in
<collectionCode id="3CE2F9F7FFA8983602B8AB9E7718F80C" box="[1000,1085,1987,2013]" pageId="8" pageNumber="494">TUMZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="0E8C31BAFFA9983701C7ABCA7237F805" ID-DOI="http://doi.org/10.5281/zenodo.6530976" ID-Zenodo-Dep="6530976" httpUri="https://zenodo.org/record/6530976/files/figure.png" pageId="9" pageNumber="495" startId="9.[151,264,1943,1967]" targetBox="[193,1391,190,1899]" targetPageId="9">
<paragraph id="5A4C6132FFA9983701C7ABCA7237F805" blockId="9.[151,1437,1943,2004]" pageId="9" pageNumber="495">
<emphasis id="6887BD20FFA9983701C7ABCA7276F87E" bold="true" box="[151,339,1943,1967]" pageId="9" pageNumber="495">FIGURES 25–26.</emphasis>
<taxonomicName id="9DF31AB1FFA998370007ABC571B1F861" authority="Joharchi" authorityName="Joharchi" authorityYear="2022" box="[343,660,1944,1968]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="495" phylum="Arthropoda" rank="species" species="latisetis" status="sp. nov.">
<emphasis id="6887BD20FFA998370007ABC5711FF861" box="[343,570,1944,1968]" italics="true" pageId="9" pageNumber="495">Cosmolaelaps latisetis</emphasis>
Joharchi
</taxonomicName>
<taxonomicNameLabel id="73B4005BFFA9983703C9ABC571CCF87E" box="[665,745,1944,1968]" pageId="9" pageNumber="495" rank="species">
<emphasis id="6887BD20FFA9983703C9ABC571C6F861" bold="true" box="[665,739,1944,1968]" pageId="9" pageNumber="495">sp. nov</emphasis>
.
</taxonomicNameLabel>
, female. 25, dorsal idiosoma; 26, ventral idiosoma. Enlarged sections not to scale.
</paragraph>
</caption>
<caption id="0E8C31BAFFAA983401C7A9E17603FA05" ID-DOI="http://doi.org/10.5281/zenodo.6530978" ID-Zenodo-Dep="6530978" box="[151,1318,1468,1492]" httpUri="https://zenodo.org/record/6530978/files/figure.png" pageId="10" pageNumber="496" startId="10.[151,264,1468,1492]" targetBox="[171,1417,193,1432]" targetPageId="10">
<paragraph id="5A4C6132FFAA983401C7A9E17603FA05" blockId="10.[151,1318,1468,1492]" box="[151,1318,1468,1492]" pageId="10" pageNumber="496">
<emphasis id="6887BD20FFAA983401C7A9E17271FA05" bold="true" box="[151,340,1468,1492]" pageId="10" pageNumber="496">FIGURES 27–29.</emphasis>
<taxonomicName id="9DF31AB1FFAA9834000AA9E171BEFA05" authority="Joharchi" authorityName="Joharchi" authorityYear="2022" box="[346,667,1468,1492]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="496" phylum="Arthropoda" rank="species" species="latisetis" status="sp. nov.">
<emphasis id="6887BD20FFAA9834000AA9E1711AFA05" box="[346,575,1468,1492]" italics="true" pageId="10" pageNumber="496">Cosmolaelaps latisetis</emphasis>
Joharchi
</taxonomicName>
<taxonomicNameLabel id="73B4005BFFAA983403F1A9E171D6FA05" box="[673,755,1468,1492]" pageId="10" pageNumber="496" rank="species">
<emphasis id="6887BD20FFAA983403F1A9E171C8FA05" bold="true" box="[673,749,1468,1492]" pageId="10" pageNumber="496">sp. nov</emphasis>
.
</taxonomicNameLabel>
, female. 27, subcapitulum; 28, epistome; 29, chelicera.
</paragraph>
</caption>
<subSubSection id="12E932B9FFAA98340197AA5F71BBF855" pageId="10" pageNumber="496" type="diagnosis">
<paragraph id="5A4C6132FFAA98340197AA5F71BBF855" blockId="10.[151,1437,1538,2033]" pageId="10" pageNumber="496">
<emphasis id="6887BD20FFAA98340197AA5F721FF9CD" bold="true" box="[199,314,1538,1564]" pageId="10" pageNumber="496">Diagnosis</emphasis>
(adult female). Dorsal shield well reticulate, with 39 pairs of scimitar-like setae, reaching well past base of next posterior setae,
<emphasis id="6887BD20FFAA983400B3AA7A7124F990" box="[483,513,1575,1601]" italics="true" pageId="10" pageNumber="496">Z5</emphasis>
shorter than
<emphasis id="6887BD20FFAA983403CBAA7A7193F990" box="[667,694,1575,1601]" italics="true" pageId="10" pageNumber="496">J5</emphasis>
(ratio of J5/Z5 ≈ 1.5); presternal area poorly sclerotised, sternal shield with lineate-reticulate ornamentation on lateral surface, bearing three pairs of long setae, extending slightly beyond of next posterior setae, ratio of shield length/width (at broadest level) ≃ 0.65; genital shield ratio of length/ width (at broadest level) ≃ 1.9, shield ornamented posteromedially by seven cells flanked by a median Λ-shaped ornamentation, anal shield ratio of length/width (at broadest level) ≃ 1.2, post-anal seta thickened, sparsely barbed, apically bent (hook shaped); distance between genital and anal shields is shorter than length of anal shield, almost all opisthogastric setae well thickened (especially
<emphasis id="6887BD20FFAA9834038BAAA27026F8C8" box="[731,771,1791,1817]" italics="true" pageId="10" pageNumber="496">Jv5</emphasis>
, apically bent) and arising on minute sclerotised platelet, except
<emphasis id="6887BD20FFAA983401B7AB7F720FF8ED" box="[231,298,1826,1853]" italics="true" pageId="10" pageNumber="496">Jv1–2</emphasis>
and
<emphasis id="6887BD20FFAA9834000FAB7F72ACF8ED" box="[351,393,1826,1852]" italics="true" pageId="10" pageNumber="496">Zv1</emphasis>
. Fixed digit of chelicera with five teeth, including two large proximal teeth (posterior to pilus dentilis). Legs without spatulate setae but most setae conspicuously thickened and some inserted on small tubercles. Legs I and IV shorter than length of idiosoma.
</paragraph>
</subSubSection>
<subSubSection id="12E932B9FFAA98320197ABD372ADFD1C" lastPageId="12" lastPageNumber="498" pageId="10" pageNumber="496" type="description">
<paragraph id="5A4C6132FFAA98340197ABD372D8F879" blockId="10.[151,1437,1538,2033]" box="[199,509,1934,1960]" pageId="10" pageNumber="496">
<emphasis id="6887BD20FFAA98340197ABD37292F879" bold="true" box="[199,439,1934,1960]" pageId="10" pageNumber="496">Description. Female</emphasis>
(n=3)
</paragraph>
<paragraph id="5A4C6132FFAA98350197ABEF7075FEB4" blockId="10.[151,1437,1538,2033]" lastBlockId="11.[151,1437,151,2014]" lastPageId="11" lastPageNumber="497" pageId="10" pageNumber="496">
<emphasis id="6887BD20FFAA98340197ABEF72A6F81D" box="[199,387,1970,1996]" italics="true" pageId="10" pageNumber="496">Dorsal idiosoma</emphasis>
(
<figureCitation id="C2C87DB7FFAA983400C2ABEF72CCF81C" box="[402,489,1970,1997]" captionStart="FIGURES 25–26" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[193,1391,190,1899]" captionTargetId="figure-19@9.[179,1408,181,1919]" captionTargetPageId="9" captionText="FIGURES 25–26. Cosmolaelaps latisetis Joharchi sp. nov., female.25, dorsal idiosoma; 26, ventral idiosoma. Enlarged sections not to scale." figureDoi="http://doi.org/10.5281/zenodo.6530976" httpUri="https://zenodo.org/record/6530976/files/figure.png" pageId="10" pageNumber="496">Figs 25</figureCitation>
,
<figureCitation id="C2C87DB7FFAA983400A6ABEF714CF81D" box="[502,617,1970,1996]" captionStart="FIGURES 30–40" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 30–40. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="10" pageNumber="496">30, 32, 33</figureCitation>
). Dorsal shield semi-circular, large, reticulate throughout, 498–512 long, 404–410 wide, covering entire dorsal idiosoma. Shield with 39 pairs of very long scimitar-like setae, 22 podonotal, 17 opisthonotal, including two pairs of
<emphasis id="6887BD20FFAB98350301ACCA7148FF61" box="[593,621,151,176]" italics="true" pageId="11" pageNumber="497">Zx</emphasis>
setae between
<emphasis id="6887BD20FFAB98350247ACCA7006FF61" box="[791,803,151,176]" italics="true" pageId="11" pageNumber="497">J</emphasis>
and
<emphasis id="6887BD20FFAB9835020AACCA704FFF61" box="[858,874,151,176]" italics="true" pageId="11" pageNumber="497">Z</emphasis>
setae, and three unpaired setae
<emphasis id="6887BD20FFAB9835059EACCA77C2FF61" box="[1230,1255,151,176]" italics="true" pageId="11" pageNumber="497">Jx</emphasis>
between
<emphasis id="6887BD20FFAB98350402ACCA7649FF60" box="[1362,1388,151,177]" italics="true" pageId="11" pageNumber="497">J2</emphasis>
and
<emphasis id="6887BD20FFAB983501C7ACE67397FF04" box="[151,178,187,213]" italics="true" pageId="11" pageNumber="497">J4</emphasis>
, setae reaching well past base of next posterior setae, mostly subequal in length (60–73), all dorsal setae very thick and slightly swollen at base, except
<emphasis id="6887BD20FFAB98350329AC8271ABFF28" box="[633,654,223,249]" italics="true" pageId="11" pageNumber="497">j1</emphasis>
(20–23) and
<emphasis id="6887BD20FFAB9835027BAC827061FF28" box="[811,836,223,249]" italics="true" pageId="11" pageNumber="497">z1</emphasis>
(17–20) simple and shortest, ratio of
<emphasis id="6887BD20FFAB983505A9AC827631FF28" box="[1273,1300,223,249]" italics="true" pageId="11" pageNumber="497">J5</emphasis>
(37–40)/Z5 (22–25) ≈ 1.5,
<emphasis id="6887BD20FFAB9835006BAD5E727CFECC" box="[315,345,259,285]" italics="true" pageId="11" pageNumber="497">Z5</emphasis>
sparsely barbed (
<figureCitation id="C2C87DB7FFAB9835034BAD5E7154FECC" box="[539,625,258,285]" captionStart="FIGURES 25–26" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[193,1391,190,1899]" captionTargetId="figure-19@9.[179,1408,181,1919]" captionTargetPageId="9" captionText="FIGURES 25–26. Cosmolaelaps latisetis Joharchi sp. nov., female.25, dorsal idiosoma; 26, ventral idiosoma. Enlarged sections not to scale." figureDoi="http://doi.org/10.5281/zenodo.6530976" httpUri="https://zenodo.org/record/6530976/files/figure.png" pageId="11" pageNumber="497">Figs 25</figureCitation>
,
<figureCitation id="C2C87DB7FFAB9835032CAD5E71C8FECC" box="[636,749,259,285]" captionStart="FIGURES 30–40" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 30–40. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">30, 32, 33</figureCitation>
). Dorsal shield with 23 pairs of pore-like structures, including seven pairs of gland openings (
<emphasis id="6887BD20FFAB98350351AD7A710EFE90" box="[513,555,295,321]" italics="true" pageId="11" pageNumber="497">gd1</emphasis>
,
<emphasis id="6887BD20FFAB9835036CAD7A7143FE90" box="[572,614,295,321]" italics="true" pageId="11" pageNumber="497">gd2</emphasis>
,
<emphasis id="6887BD20FFAB98350326AD7A7184FE90" box="[630,673,295,321]" italics="true" pageId="11" pageNumber="497">gd4</emphasis>
,
<emphasis id="6887BD20FFAB983503E1AD7A71FEFE90" box="[689,731,295,321]" italics="true" pageId="11" pageNumber="497">gd5</emphasis>
,
<emphasis id="6887BD20FFAB983503BCAD7B7033FE91" box="[748,790,294,320]" italics="true" pageId="11" pageNumber="497">gd6</emphasis>
,
<emphasis id="6887BD20FFAB98350276AD7A7075FE90" box="[806,848,295,321]" italics="true" pageId="11" pageNumber="497">gd8</emphasis>
,
<emphasis id="6887BD20FFAB98350231AD7A70AFFE90" box="[865,906,295,321]" italics="true" pageId="11" pageNumber="497">gd9</emphasis>
) and 16 pairs of poroids. Shape, position and relative length and shape of setae shown in
<figureCitation id="C2C87DB7FFAB983503D0AD1671D2FEB4" box="[640,759,330,357]" captionStart="FIGURES 25–26" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[193,1391,190,1899]" captionTargetId="figure-19@9.[179,1408,181,1919]" captionTargetPageId="9" captionText="FIGURES 25–26. Cosmolaelaps latisetis Joharchi sp. nov., female.25, dorsal idiosoma; 26, ventral idiosoma. Enlarged sections not to scale." figureDoi="http://doi.org/10.5281/zenodo.6530976" httpUri="https://zenodo.org/record/6530976/files/figure.png" pageId="11" pageNumber="497">Figures 25</figureCitation>
and
<figureCitation id="C2C87DB7FFAB9835027DAD167069FEB4" box="[813,844,331,357]" captionStart="FIGURES 30–40" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 30–40. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">30</figureCitation>
.
</paragraph>
<paragraph id="5A4C6132FFAB98350197AD327089FA4C" blockId="11.[151,1437,151,2014]" pageId="11" pageNumber="497">
<emphasis id="6887BD20FFAB98350197AD3272A2FE58" box="[199,391,367,393]" italics="true" pageId="11" pageNumber="497">Ventral idiosoma</emphasis>
(
<figureCitation id="C2C87DB7FFAB983500C7AD3272CAFE58" box="[407,495,367,393]" captionStart="FIGURES 25–26" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[193,1391,190,1899]" captionTargetId="figure-19@9.[179,1408,181,1919]" captionTargetPageId="9" captionText="FIGURES 25–26. Cosmolaelaps latisetis Joharchi sp. nov., female.25, dorsal idiosoma; 26, ventral idiosoma. Enlarged sections not to scale." figureDoi="http://doi.org/10.5281/zenodo.6530976" httpUri="https://zenodo.org/record/6530976/files/figure.png" pageId="11" pageNumber="497">Figs 26</figureCitation>
,
<figureCitation id="C2C87DB7FFAB983500ADAD327156FE58" box="[509,627,367,393]" captionStart="FIGURES 30–40" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 30–40. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">31, 34, 40</figureCitation>
). Tritosternum with paired pilose laciniae (77–80), fused basally (6–8), columnar base 27–29 × 15–17 wide; presternal area poorly sclerotised. Sternal shield (length 90–95) narrowest between coxae II (94–97), widest (143–147), anterior margin of shield hardly conspicuous, posterior margin distinctly concave, anterolateral corners narrowly slightly extending between coxae I–II, gland pores
<emphasis id="6887BD20FFAB983505AEAD867603FE24" box="[1278,1318,475,501]" italics="true" pageId="11" pageNumber="497">gvb</emphasis>
indistinct; endopodal elements between coxae II and III fused with shield and endopodal elements III/IV relatively small, subtriangular, posteriorly not extended. Sternal shield with three pairs of subequal smooth setae (
<emphasis id="6887BD20FFAB9835059DAE7E7620FDEC" box="[1229,1285,547,573]" italics="true" pageId="11" pageNumber="497">st1-3</emphasis>
) (42–46) and two pairs of large slit-shaped pore-like structures (
<emphasis id="6887BD20FFAB98350395AE1A71C2FDB0" box="[709,743,583,609]" italics="true" pageId="11" pageNumber="497">iv1</emphasis>
adjacent to setae
<emphasis id="6887BD20FFAB983502FAAE1A70EEFDB0" box="[938,971,583,609]" italics="true" pageId="11" pageNumber="497">st1</emphasis>
;
<emphasis id="6887BD20FFAB98350288AE1A70DEFDB0" box="[984,1019,583,609]" italics="true" pageId="11" pageNumber="497">iv2</emphasis>
between
<emphasis id="6887BD20FFAB98350534AE1A77A1FDB0" box="[1124,1156,583,609]" italics="true" pageId="11" pageNumber="497">st2</emphasis>
and
<emphasis id="6887BD20FFAB983505E9AE1A77FFFDB0" box="[1209,1242,583,609]" italics="true" pageId="11" pageNumber="497">st3</emphasis>
), setae extending slightly beyond base of following setae; shield with lineate-reticulate ornamentation on lateral surface, longitudinally median part smooth (or faintly reticulated) (
<figureCitation id="C2C87DB7FFAB983503C2AED271CEFD78" box="[658,747,655,681]" captionStart="FIGURES 25–26" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[193,1391,190,1899]" captionTargetId="figure-19@9.[179,1408,181,1919]" captionTargetPageId="9" captionText="FIGURES 25–26. Cosmolaelaps latisetis Joharchi sp. nov., female.25, dorsal idiosoma; 26, ventral idiosoma. Enlarged sections not to scale." figureDoi="http://doi.org/10.5281/zenodo.6530976" httpUri="https://zenodo.org/record/6530976/files/figure.png" pageId="11" pageNumber="497">Figs 26</figureCitation>
,
<figureCitation id="C2C87DB7FFAB983503AAAED2703CFD78" box="[762,793,655,681]" captionStart="FIGURES 30–40" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 30–40. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">31</figureCitation>
). Metasternal setae
<emphasis id="6887BD20FFAB983502AFAED27705FD78" box="[1023,1056,655,681]" italics="true" pageId="11" pageNumber="497">st4</emphasis>
(42–44) and metasternal poroids located on soft integument; metasternal platelets absent. Genital shield flask-shaped, slightly expanded laterally past level of setae
<emphasis id="6887BD20FFAB98350062AE8A7276FD20" box="[306,339,727,753]" italics="true" pageId="11" pageNumber="497">st5</emphasis>
, length 208–212, maximum width 108–111, posterior margin rounded, anterior hyaline margin of shield irregularly convex, overlapping posterior area of sternal shield (extending beyond base of setae
<emphasis id="6887BD20FFAB98350443AEA67611FCC4" box="[1299,1332,763,789]" italics="true" pageId="11" pageNumber="497">st3</emphasis>
), surface ornamented posteromedially by seven cells flanked by a median Λ-shaped ornamentation; bearing a pair of simple setae
<emphasis id="6887BD20FFAB98350188AF1E73DCFC8C" box="[216,249,835,861]" italics="true" pageId="11" pageNumber="497">st5</emphasis>
(38–40) (
<figureCitation id="C2C87DB7FFAB9835003BAF1E72E6FC8C" box="[363,451,835,861]" captionStart="FIGURES 25–26" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[193,1391,190,1899]" captionTargetId="figure-19@9.[179,1408,181,1919]" captionTargetPageId="9" captionText="FIGURES 25–26. Cosmolaelaps latisetis Joharchi sp. nov., female.25, dorsal idiosoma; 26, ventral idiosoma. Enlarged sections not to scale." figureDoi="http://doi.org/10.5281/zenodo.6530976" httpUri="https://zenodo.org/record/6530976/files/figure.png" pageId="11" pageNumber="497">Figs 26</figureCitation>
,
<figureCitation id="C2C87DB7FFAB98350081AF1E7138FC8C" box="[465,541,835,861]" captionStart="FIGURES 30–40" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 30–40. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">31, 34</figureCitation>
). Shield flanked by two pairs of minute, oval paragenital platelets; paragenital poroids
<emphasis id="6887BD20FFAB983501A1AF3A7236FC50" box="[241,275,871,897]" italics="true" pageId="11" pageNumber="497">iv5</emphasis>
located on soft cuticle lateral to shield near seta
<emphasis id="6887BD20FFAB9835027BAF3A7069FC50" box="[811,844,871,897]" italics="true" pageId="11" pageNumber="497">st5</emphasis>
. Anal shield subtriangular, rounded anteriorly, length 91–95, width 75–78, anterior half faintly lineate-reticulate, bearing a pair of simple para-anal setae (23–26), postanal seta (30–32) thickened, sparsely barbed, apically bent (hook shaped) (see
<figureCitation id="C2C87DB7FFAB9835055FAFF27741FC18" box="[1039,1124,943,969]" captionStart="FIGURES 25–26" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[193,1391,190,1899]" captionTargetId="figure-19@9.[179,1408,181,1919]" captionTargetPageId="9" captionText="FIGURES 25–26. Cosmolaelaps latisetis Joharchi sp. nov., female.25, dorsal idiosoma; 26, ventral idiosoma. Enlarged sections not to scale." figureDoi="http://doi.org/10.5281/zenodo.6530976" httpUri="https://zenodo.org/record/6530976/files/figure.png" pageId="11" pageNumber="497">Figs 26</figureCitation>
,
<figureCitation id="C2C87DB7FFAB98350521AFF277C6FC18" box="[1137,1251,943,969]" captionStart="FIGURES 30–40" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 30–40. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">31, 34, 40</figureCitation>
) and with a pair of marginal pores (
<emphasis id="6887BD20FFAB9835003AAF8E72B7FC3C" box="[362,402,979,1005]" italics="true" pageId="11" pageNumber="497">gv3</emphasis>
); cribrum consisting of a terminal tuft with 3–4 irregular rows of spicules, anteriormost rows of spicules extending slightly beyond post-anal seta (
<figureCitation id="C2C87DB7FFAB983503A3AFAA706EFBC0" box="[755,843,1015,1041]" captionStart="FIGURES 25–26" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[193,1391,190,1899]" captionTargetId="figure-19@9.[179,1408,181,1919]" captionTargetPageId="9" captionText="FIGURES 25–26. Cosmolaelaps latisetis Joharchi sp. nov., female.25, dorsal idiosoma; 26, ventral idiosoma. Enlarged sections not to scale." figureDoi="http://doi.org/10.5281/zenodo.6530976" httpUri="https://zenodo.org/record/6530976/files/figure.png" pageId="11" pageNumber="497">Figs 26</figureCitation>
,
<figureCitation id="C2C87DB7FFAB98350209AFAA70F5FBC0" box="[857,976,1015,1041]" captionStart="FIGURES 30–40" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 30–40. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">31, 34, 40</figureCitation>
). Soft opisthogastric cuticle surrounding genital and anal shields with seven pairs of poroids, including one pair of paragenital poroids
<emphasis id="6887BD20FFAB983505FCA84677EBFBE4" box="[1196,1230,1051,1077]" italics="true" pageId="11" pageNumber="497">iv5</emphasis>
,
<emphasis id="6887BD20FFAB9835058CA84677DBFBE5" box="[1244,1278,1051,1076]" italics="true" pageId="11" pageNumber="497">ivp</emphasis>
and five pairs of
<emphasis id="6887BD20FFAB983501E5A86273F2FB89" box="[181,215,1087,1112]" italics="true" pageId="11" pageNumber="497">ivo</emphasis>
, inguinal gland pores
<emphasis id="6887BD20FFAB9835009FA86272D2FB88" box="[463,503,1087,1113]" italics="true" pageId="11" pageNumber="497">gv2</emphasis>
on soft cuticle close to parapodals and one pair of suboval metapodal plates (13–16 long × 8–10 wide) present, soft cuticle with two minute paragenital platelets between metapodals and genital shield, bearing 16 pairs of thick setae (
<emphasis id="6887BD20FFAB983500A8A8DA7171FB70" box="[504,596,1159,1185]" italics="true" pageId="11" pageNumber="497">Jv1–Jv5</emphasis>
,
<emphasis id="6887BD20FFAB98350331A8DA7199FB71" box="[609,700,1159,1185]" italics="true" pageId="11" pageNumber="497">Zv1– Zv</emphasis>
5,
<emphasis id="6887BD20FFAB98350388A8DA7001FB70" box="[728,804,1159,1185]" italics="true" pageId="11" pageNumber="497">R1–R4</emphasis>
,
<emphasis id="6887BD20FFAB98350262A8DA7082FB70" box="[818,935,1159,1185]" italics="true" pageId="11" pageNumber="497">UR2–UR3</emphasis>
); almost all setae uniform in length (20–30), and arising on small sclerotised platelet, except
<emphasis id="6887BD20FFAB983503E8A8F671DFFB14" box="[696,762,1195,1221]" italics="true" pageId="11" pageNumber="497">Jv1–2</emphasis>
(33–35) and
<emphasis id="6887BD20FFAB983502C4A8F6709BFB14" box="[916,958,1195,1221]" italics="true" pageId="11" pageNumber="497">Zv1</emphasis>
(38–40) longer and setiform (
<figureCitation id="C2C87DB7FFAB98350444A8F67649FB14" box="[1300,1388,1195,1221]" captionStart="FIGURES 25–26" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[193,1391,190,1899]" captionTargetId="figure-19@9.[179,1408,181,1919]" captionTargetPageId="9" captionText="FIGURES 25–26. Cosmolaelaps latisetis Joharchi sp. nov., female.25, dorsal idiosoma; 26, ventral idiosoma. Enlarged sections not to scale." figureDoi="http://doi.org/10.5281/zenodo.6530976" httpUri="https://zenodo.org/record/6530976/files/figure.png" pageId="11" pageNumber="497">Figs 26</figureCitation>
,
<figureCitation id="C2C87DB7FFAB98350429A8F67392FB38" captionStart="FIGURES 30–40" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 30–40. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">31, 34</figureCitation>
). Peritrematal shield well developed laterad peritremes, bearing one gland pore and one poroid at level near coxae II–III, post-stigmatic extension of shield relatively wide, not reaching posterior level of coxae IV, with two pairs of poroids and one pair of gland pores; anterior region of peritrematal shield fused to anterior margin of dorsal shield behind setae
<emphasis id="6887BD20FFAB98350021A96672AFFA84" box="[369,394,1339,1365]" italics="true" pageId="11" pageNumber="497">z1</emphasis>
. Peritremes long, extending to mid-level of coxa I (near
<emphasis id="6887BD20FFAB983502ADA9667704FA84" box="[1021,1057,1339,1365]" italics="true" pageId="11" pageNumber="497">id1</emphasis>
). Two exopodal platelets between coxae II–III and III–IV present, anterior platelet small and subtriangular, those between coxae III–IV linear, narrow, more or less abutting to parapodal more or less abutting to parapodals.
</paragraph>
<paragraph id="5A4C6132FFAB98350197A9FA700CF8A0" blockId="11.[151,1437,151,2014]" pageId="11" pageNumber="497">
<taxonomicName id="9DF31AB1FFAB98350197A9FA7273FA10" authorityName="Krefft" authorityYear="1871" box="[199,342,1447,1473]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="11" pageNumber="497" phylum="Nematoda" rank="genus">
<emphasis id="6887BD20FFAB98350197A9FA7273FA10" box="[199,342,1447,1473]" italics="true" pageId="11" pageNumber="497">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="C2C87DB7FFAB98350038A9FA7135FA10" box="[360,528,1447,1473]" captionStart="FIGURES 27–29" captionStartId="10.[151,264,1468,1492]" captionTargetBox="[171,1417,193,1432]" captionTargetId="figure-18@10.[151,1436,181,1444]" captionTargetPageId="10" captionText="FIGURES 27–29. Cosmolaelaps latisetis Joharchi sp. nov., female. 27, subcapitulum; 28, epistome; 29, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530978" httpUri="https://zenodo.org/record/6530978/files/figure.png" pageId="11" pageNumber="497">Figures 27–29</figureCitation>
,
<figureCitation id="C2C87DB7FFAB9835034EA9FB71B2FA10" box="[542,663,1446,1473]" captionStart="FIGURES 30–40" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 30–40. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">35, 36, 39</figureCitation>
). Anterior margin of epistome convex, irregularly denticulate, with smooth median spike, lineate posteriorly (
<figureCitation id="C2C87DB7FFAB983503D5A99671FAFA34" box="[645,735,1483,1509]" captionStart="FIGURES 27–29" captionStartId="10.[151,264,1468,1492]" captionTargetBox="[171,1417,193,1432]" captionTargetId="figure-18@10.[151,1436,181,1444]" captionTargetPageId="10" captionText="FIGURES 27–29. Cosmolaelaps latisetis Joharchi sp. nov., female. 27, subcapitulum; 28, epistome; 29, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530978" httpUri="https://zenodo.org/record/6530978/files/figure.png" pageId="11" pageNumber="497">Fig. 28</figureCitation>
). Hypostomal groove with six transverse rows of denticles, each row with 6–16 small denticles, except anteriormost row widest, slightly convex, with 28–30 small denticles, groove with smooth posterior transverse line and progressively narrowing from anterior to posterior (
<figureCitation id="C2C87DB7FFAB9835046CAA4E76B2F9FC" box="[1340,1431,1555,1581]" captionStart="FIGURES 27–29" captionStartId="10.[151,264,1468,1492]" captionTargetBox="[171,1417,193,1432]" captionTargetId="figure-18@10.[151,1436,181,1444]" captionTargetPageId="10" captionText="FIGURES 27–29. Cosmolaelaps latisetis Joharchi sp. nov., female. 27, subcapitulum; 28, epistome; 29, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530978" httpUri="https://zenodo.org/record/6530978/files/figure.png" pageId="11" pageNumber="497">Figs 27</figureCitation>
,
<figureCitation id="C2C87DB7FFAB983501C7AA6A7392F983" box="[151,183,1591,1618]" captionStart="FIGURES 30–40" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 30–40. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">36</figureCitation>
). Corniculi horn-like, internal malae longer than corniculi, comprising a pair of pilose median projections, and 10–12 lateral, shorter, tentacle-like, smooth projections (
<figureCitation id="C2C87DB7FFAB9835024FAA067053F9A4" box="[799,886,1627,1653]" captionStart="FIGURES 27–29" captionStartId="10.[151,264,1468,1492]" captionTargetBox="[171,1417,193,1432]" captionTargetId="figure-18@10.[151,1436,181,1444]" captionTargetPageId="10" captionText="FIGURES 27–29. Cosmolaelaps latisetis Joharchi sp. nov., female. 27, subcapitulum; 28, epistome; 29, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530978" httpUri="https://zenodo.org/record/6530978/files/figure.png" pageId="11" pageNumber="497">Fig. 27</figureCitation>
); labrum acuminate, longer than internal malae, fringed; hypostomal and capitular setae smooth,
<emphasis id="6887BD20FFAB983503E8AA2271F1F948" box="[696,724,1663,1689]" italics="true" pageId="11" pageNumber="497">h3</emphasis>
(48–51)&gt;
<emphasis id="6887BD20FFAB98350202AA227048F948" box="[850,877,1663,1689]" italics="true" pageId="11" pageNumber="497">h1</emphasis>
(36–39)&gt;
<emphasis id="6887BD20FFAB983502BBAA227720F949" box="[1003,1029,1663,1688]" italics="true" pageId="11" pageNumber="497">pc</emphasis>
(28–30)&gt;
<emphasis id="6887BD20FFAB983505D3AA2277BAF948" box="[1155,1183,1663,1689]" italics="true" pageId="11" pageNumber="497">h2</emphasis>
(17–20) (
<figureCitation id="C2C87DB7FFAB9835045DAA227641F948" box="[1293,1380,1663,1689]" captionStart="FIGURES 27–29" captionStartId="10.[151,264,1468,1492]" captionTargetBox="[171,1417,193,1432]" captionTargetId="figure-18@10.[151,1436,181,1444]" captionTargetPageId="10" captionText="FIGURES 27–29. Cosmolaelaps latisetis Joharchi sp. nov., female. 27, subcapitulum; 28, epistome; 29, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530978" httpUri="https://zenodo.org/record/6530978/files/figure.png" pageId="11" pageNumber="497">Figs 27</figureCitation>
,
<figureCitation id="C2C87DB7FFAB98350420AA2276B5F94B" box="[1392,1424,1663,1690]" captionStart="FIGURES 30–40" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 30–40. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">36</figureCitation>
). Chaetotaxy of palps: trochanter 2, femur 5, genu 6, tibia 14, tarsus 15, all setae smooth except palpfemur with seta
<emphasis id="6887BD20FFAB983501C7AA9A7396F930" box="[151,179,1735,1761]" italics="true" pageId="11" pageNumber="497">d3</emphasis>
thickened and inserted on small tubercles, seta
<emphasis id="6887BD20FFAB98350396AA9A71F9F930" box="[710,732,1735,1761]" italics="true" pageId="11" pageNumber="497">al</emphasis>
thickened; palpgenu with
<emphasis id="6887BD20FFAB98350554AA9A770DF930" box="[1028,1064,1735,1761]" italics="true" pageId="11" pageNumber="497">al1</emphasis>
stout,
<emphasis id="6887BD20FFAB98350522AA9A77B3F930" box="[1138,1174,1735,1761]" italics="true" pageId="11" pageNumber="497">al2</emphasis>
paddle-like; palp tarsal claw two-tined (
<figureCitation id="C2C87DB7FFAB9835001FAAB67286F8D4" box="[335,419,1770,1797]" captionStart="FIGURES 30–40" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 30–40. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">Fig. 35</figureCitation>
). Supralabral process indistinct. Fixed digit of chelicera with an offset and subapical tooth (gabelzhan), followed by four teeth, including two large proximal teeth posterior to a setaceous pilus dentilis, dorsal cheliceral seta prostrate, arthrodial membrane with a rounded flap and normal filaments; cheliceral lyrifissures distinct, movable digit of chelicera bidentate (
<figureCitation id="C2C87DB7FFAB983503C9AB0A71D5F8A0" box="[665,752,1879,1905]" captionStart="FIGURES 27–29" captionStartId="10.[151,264,1468,1492]" captionTargetBox="[171,1417,193,1432]" captionTargetId="figure-18@10.[151,1436,181,1444]" captionTargetPageId="10" captionText="FIGURES 27–29. Cosmolaelaps latisetis Joharchi sp. nov., female. 27, subcapitulum; 28, epistome; 29, chelicera." figureDoi="http://doi.org/10.5281/zenodo.6530978" httpUri="https://zenodo.org/record/6530978/files/figure.png" pageId="11" pageNumber="497">Figs 29</figureCitation>
,
<figureCitation id="C2C87DB7FFAB983503ACAB0A7039F8A0" box="[764,796,1879,1905]" captionStart="FIGURES 30–40" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 30–40. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="11" pageNumber="497">39</figureCitation>
).
</paragraph>
<paragraph id="5A4C6132FFAB98350197AB267042F844" blockId="11.[151,1437,151,2014]" box="[199,871,1915,1941]" pageId="11" pageNumber="497">
<emphasis id="6887BD20FFAB98350197AB2672EAF845" box="[199,463,1915,1940]" italics="true" pageId="11" pageNumber="497">Insemination structures</emphasis>
. Not seen, apparently unsclerotised.
</paragraph>
<paragraph id="5A4C6132FFAB98320197ABC27288FD78" blockId="11.[151,1437,151,2014]" lastBlockId="12.[151,1437,151,789]" lastPageId="12" lastPageNumber="498" pageId="11" pageNumber="497">
<emphasis id="6887BD20FFAB98350197ABC273D9F869" box="[199,252,1951,1976]" italics="true" pageId="11" pageNumber="497">Legs</emphasis>
(
<figureCitation id="C2C87DB7FFAB9835005AABC272A8F868" box="[266,397,1951,1977]" captionStart="FIGURE 41–44" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[183,1406,196,1906]" captionTargetId="figure-18@14.[175,1413,181,1919]" captionTargetPageId="14" captionText="FIGURE 41–44. Cosmolaelaps latisetis Joharchi sp. nov., female. 41, leg I (trochanter-tibia); 42, leg II; 43, leg III; 44, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530984" httpUri="https://zenodo.org/record/6530984/files/figure.png" pageId="11" pageNumber="497">Figs 41–44</figureCitation>
). Legs II and III short (352–356, 343–348), I and IV longer (465–470) (excluding pre-tarsus). Chaetotaxy normal for free-living
<taxonomicName id="9DF31AB1FFAB9835034DAB9E71BDF80C" box="[541,664,1987,2013]" class="Arachnida" family="Laelapidae" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="497" phylum="Arthropoda" rank="family">Laelapidae</taxonomicName>
(
<emphasis id="6887BD20FFAB983503FAAB9E71CDF80D" box="[682,744,1987,2012]" italics="true" pageId="11" pageNumber="497">sensu</emphasis>
<bibRefCitation id="3E621CC3FFAB983503A0AB9E70A8F80F" author="Evans, G. O." box="[752,909,1987,2014]" pageId="11" pageNumber="497" pagination="275 - 303" refId="ref14188" refString="Evans, G. O. (1963 a) Observations on the chaetotaxy of the legs in the free-living Gamasina (Acari: Mesostigmata). Bulletin of the British Museum (Natural History) Zoology, 10 (5), 275 - 303. https: // doi. org / 10.5962 / bhl. part. 20528" type="journal article" year="1963">Evans, 1963a</bibRefCitation>
): Leg I (
<figureCitation id="C2C87DB7FFAB983502BFAB9E7761F80C" box="[1007,1092,1987,2013]" captionStart="FIGURE 41–44" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[183,1406,196,1906]" captionTargetId="figure-18@14.[175,1413,181,1919]" captionTargetPageId="14" captionText="FIGURE 41–44. Cosmolaelaps latisetis Joharchi sp. nov., female. 41, leg I (trochanter-tibia); 42, leg II; 43, leg III; 44, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530984" httpUri="https://zenodo.org/record/6530984/files/figure.png" pageId="11" pageNumber="497">Fig. 41</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-1/1, 0/2-1 (
<emphasis id="6887BD20FFAC98320079ACCA721AFF60" box="[297,319,151,177]" italics="true" pageId="12" pageNumber="498">al</emphasis>
and
<emphasis id="6887BD20FFAC98320022ACCA72ABFF60" box="[370,398,151,177]" italics="true" pageId="12" pageNumber="498">ad</emphasis>
thickened and inserted on small tubercles), femur 2-3/1, 2/3-2 (all setae on segment thickened except
<emphasis id="6887BD20FFAC983201B9ACE67229FF04" box="[233,268,187,213]" italics="true" pageId="12" pageNumber="498">al2</emphasis>
,
<emphasis id="6887BD20FFAC9832004CACE67213FF05" box="[284,310,187,212]" italics="true" pageId="12" pageNumber="498">av</emphasis>
,
<emphasis id="6887BD20FFAC98320016ACE6724FFF04" box="[326,362,187,213]" italics="true" pageId="12" pageNumber="498">pl1</emphasis>
,
<emphasis id="6887BD20FFAC98320029ACE672B8FF04" box="[377,413,187,213]" italics="true" pageId="12" pageNumber="498">pl2</emphasis>
,
<emphasis id="6887BD20FFAC983200FCACE672F2FF04" box="[428,471,187,213]" italics="true" pageId="12" pageNumber="498">ad2</emphasis>
and
<emphasis id="6887BD20FFAC98320340ACE67111FF04" box="[528,564,187,213]" italics="true" pageId="12" pageNumber="498">al1</emphasis>
,
<emphasis id="6887BD20FFAC98320313ACE67148FF04" box="[579,621,187,213]" italics="true" pageId="12" pageNumber="498">ad1</emphasis>
,
<emphasis id="6887BD20FFAC9832032DACE67182FF04" box="[637,679,187,213]" italics="true" pageId="12" pageNumber="498">ad3</emphasis>
inserted on small tubercles, see
<figureCitation id="C2C87DB7FFAC9832054BACE677E7FF04" box="[1051,1218,187,213]" captionStart="FIGURES 30–40" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 30–40. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="12" pageNumber="498">Figures 37, 38</figureCitation>
), genu 2-3/2, 3/1-2 (all setae on segment thickened and dorsals inserted on small tubercles, see
<figureCitation id="C2C87DB7FFAC9832055AAC8277E7FF28" box="[1034,1218,223,249]" captionStart="FIGURES 30–40" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 30–40. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="12" pageNumber="498">Figures 37&amp; 38</figureCitation>
), tibia 2-3/2, 3/1-2 (all setae on segment thickened and dorsals inserted on small tubercles, see
<figureCitation id="C2C87DB7FFAC983202BEAD5E77B1FECC" box="[1006,1172,259,285]" captionStart="FIGURES 30–40" captionStartId="13.[151,264,1874,1898]" captionTargetBox="[214,1374,184,1849]" captionTargetId="figure-19@13.[212,1375,181,1851]" captionTargetPageId="13" captionText="FIGURES 30–40. DIC micrographs of Cosmolaelaps latisetis Joharchi sp. nov., female. 30, idiosoma in dorsal view; 31, idiosoma in ventral view; 32, general view of some podonotal setae; 33, general view of some opisthonotal setae; 34, general view of opisthogastric area; 35, distal portion of palp, with a focus on tarsal claw; 36, subcapitulum; 37, dorsal aspect of femur, genu and tibia leg I; 38, ventral aspect of femur, genu and tibia leg I; 39, chelicera; 40, anal shield." figureDoi="http://doi.org/10.5281/zenodo.6531011" httpUri="https://zenodo.org/record/6531011/files/figure.png" pageId="12" pageNumber="498">Figures 37, 38</figureCitation>
). Leg II (
<figureCitation id="C2C87DB7FFAC983205AEAD5E7671FECC" box="[1278,1364,259,285]" captionStart="FIGURE 41–44" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[183,1406,196,1906]" captionTargetId="figure-18@14.[175,1413,181,1919]" captionTargetPageId="14" captionText="FIGURE 41–44. Cosmolaelaps latisetis Joharchi sp. nov., female. 41, leg I (trochanter-tibia); 42, leg II; 43, leg III; 44, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530984" httpUri="https://zenodo.org/record/6530984/files/figure.png" pageId="12" pageNumber="498">Fig. 42</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-0/1, 0/2-1 (
<emphasis id="6887BD20FFAC9832036BAD7A7174FE90" box="[571,593,295,321]" italics="true" pageId="12" pageNumber="498">al</emphasis>
thickened), femur 2-3/1, 2/2-1 (all setae on segment thickened except
<emphasis id="6887BD20FFAC98320421AD7A76B0FE90" box="[1393,1429,295,321]" italics="true" pageId="12" pageNumber="498">al2</emphasis>
, and
<emphasis id="6887BD20FFAC98320197AD1673D4FEB4" box="[199,241,331,357]" italics="true" pageId="12" pageNumber="498">ad1</emphasis>
,
<emphasis id="6887BD20FFAC98320050AD16720EFEB4" box="[256,299,331,357]" italics="true" pageId="12" pageNumber="498">pd1</emphasis>
,
<emphasis id="6887BD20FFAC98320069AD167246FEB4" box="[313,355,331,357]" italics="true" pageId="12" pageNumber="498">pd2</emphasis>
inserted on small tubercles,
<emphasis id="6887BD20FFAC983203F8AD1671F7FEB4" box="[680,722,331,357]" italics="true" pageId="12" pageNumber="498">pd2</emphasis>
longest), genu 2-3/1, 2/1-2 (dorsals and ventrals thickened and dorsals inserted on small tubercles), tibia 2-2/1, 2/1-2 (all setae on segment thickened except anterolaterals). Leg III (
<figureCitation id="C2C87DB7FFAC98320192ADCE7232FE7C" box="[194,279,403,429]" captionStart="FIGURE 41–44" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[183,1406,196,1906]" captionTargetId="figure-18@14.[175,1413,181,1919]" captionTargetPageId="14" captionText="FIGURE 41–44. Cosmolaelaps latisetis Joharchi sp. nov., female. 41, leg I (trochanter-tibia); 42, leg II; 43, leg III; 44, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530984" httpUri="https://zenodo.org/record/6530984/files/figure.png" pageId="12" pageNumber="498">Fig. 43</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-1/1, 0/2-0 (
<emphasis id="6887BD20FFAC98320255ADCE703EFE7C" box="[773,795,403,429]" italics="true" pageId="12" pageNumber="498">al</emphasis>
and
<emphasis id="6887BD20FFAC98320201ADCE7048FE7C" box="[849,877,403,429]" italics="true" pageId="12" pageNumber="498">ad</emphasis>
thickened and inserted on small tubercles), femur 1-2/1, 1/0-1 (dorsals and anterolateral thickened,
<emphasis id="6887BD20FFAC9832039DADEA71D2FE00" box="[717,759,439,465]" italics="true" pageId="12" pageNumber="498">ad1</emphasis>
and
<emphasis id="6887BD20FFAC98320263ADEA7078FE00" box="[819,861,439,465]" italics="true" pageId="12" pageNumber="498">ad2</emphasis>
inserted on small tubercle), genu 2-2/1, 2/1-1 (all setae on segment thickened), tibia: 2-1/1, 2/1-1 (all setae on segment thickened). Leg IV (
<figureCitation id="C2C87DB7FFAC983205FDAD867620FE24" box="[1197,1285,475,501]" captionStart="FIGURE 41–44" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[183,1406,196,1906]" captionTargetId="figure-18@14.[175,1413,181,1919]" captionTargetPageId="14" captionText="FIGURE 41–44. Cosmolaelaps latisetis Joharchi sp. nov., female. 41, leg I (trochanter-tibia); 42, leg II; 43, leg III; 44, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530984" httpUri="https://zenodo.org/record/6530984/files/figure.png" pageId="12" pageNumber="498">Fig. 44</figureCitation>
): coxa 0-0/1, 0/0-0, trochanter 1-1/1, 0/2-0 (
<emphasis id="6887BD20FFAC983200A3ADA2712CFDC8" box="[499,521,511,537]" italics="true" pageId="12" pageNumber="498">al</emphasis>
and
<emphasis id="6887BD20FFAC98320312ADA2717BFDC8" box="[578,606,511,537]" italics="true" pageId="12" pageNumber="498">ad</emphasis>
thickened and inserted on small tubercles), femur 1-2/1, 1/0-1 (
<emphasis id="6887BD20FFAC98320469ADA27646FDC8" box="[1337,1379,511,537]" italics="true" pageId="12" pageNumber="498">ad1</emphasis>
,
<emphasis id="6887BD20FFAC98320422ADA276B9FDC8" box="[1394,1436,511,537]" italics="true" pageId="12" pageNumber="498">ad2</emphasis>
thickened and inserted on small tubercles), genu 2-2/1, 3/0-1 (all setae on segment slightly thickened), tibia 2-1/1, 3/1-2 (all setae on segment slightly thickened). Tarsi II–IV with 18 setae (3- 3/2, 3/2-3 +
<emphasis id="6887BD20FFAC983205D4AE1A7780FDB1" box="[1156,1189,583,608]" italics="true" pageId="12" pageNumber="498">mv</emphasis>
,
<emphasis id="6887BD20FFAC983205E4AE1A77F3FDB0" box="[1204,1238,583,609]" italics="true" pageId="12" pageNumber="498">md</emphasis>
); almost all setae on tarsus II–IV thickened, see
<figureCitation id="C2C87DB7FFAC983200A3AE3671BDFD54" box="[499,664,619,645]" captionStart="FIGURE 41–44" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[183,1406,196,1906]" captionTargetId="figure-18@14.[175,1413,181,1919]" captionTargetPageId="14" captionText="FIGURE 41–44. Cosmolaelaps latisetis Joharchi sp. nov., female. 41, leg I (trochanter-tibia); 42, leg II; 43, leg III; 44, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530984" httpUri="https://zenodo.org/record/6530984/files/figure.png" pageId="12" pageNumber="498">Figures 42–44</figureCitation>
. All pretarsi with well-developed paired claws, rounded pulvilli and normal ambulacral stalk.
</paragraph>
<paragraph id="5A4C6132FFAC98320197AEEE72ADFD1C" blockId="12.[151,1437,151,789]" box="[199,392,691,717]" pageId="12" pageNumber="498">
<emphasis id="6887BD20FFAC98320197AEEE722EFD1C" bold="true" box="[199,267,691,717]" pageId="12" pageNumber="498">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="12E932B9FFAC98320197AE8A72D3FCC4" pageId="12" pageNumber="498" type="etymology">
<paragraph id="5A4C6132FFAC98320197AE8A72D3FCC4" blockId="12.[151,1437,151,789]" pageId="12" pageNumber="498">
<emphasis id="6887BD20FFAC98320197AE8A726BFD20" bold="true" box="[199,334,727,753]" pageId="12" pageNumber="498">Etymology.</emphasis>
The specific name
<taxonomicName id="9DF31AB1FFAC9832037CAE8A71A6FD20" authorityName="Joharchi" authorityYear="2022" box="[556,643,727,753]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="498" phylum="Arthropoda" rank="species" species="latisetis">
<emphasis id="6887BD20FFAC9832037CAE8A71A6FD20" box="[556,643,727,753]" italics="true" pageId="12" pageNumber="498">latisetis</emphasis>
</taxonomicName>
, from the Latin &quot;
<emphasis id="6887BD20FFAC98320219AE8A704BFD20" box="[841,878,727,753]" italics="true" pageId="12" pageNumber="498">lati</emphasis>
-&quot; meaning broad, thick and &quot;
<emphasis id="6887BD20FFAC983205EDAE8A77CAFD21" box="[1213,1263,727,752]" italics="true" pageId="12" pageNumber="498">setis</emphasis>
&quot; meaning hair, refers to the broad dorsal setae.
</paragraph>
</subSubSection>
</treatment>
</document>

177
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@ -0,0 +1,177 @@
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<mods:title id="863B657B6DE43DEEDF53B2815699395A">New species and new records of Cosmolaelaps Berlese (Acari: Laelapidae) from Russia, with a review of the Russian species of the genus</mods:title>
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<subSubSection id="12E932B9FFAF983101C7AAB971CCF92F" box="[151,745,1764,1791]" pageId="15" pageNumber="501" type="nomenclature">
<paragraph id="5A4C6132FFAF983101C7AAB971CCF92F" blockId="15.[151,745,1764,1791]" box="[151,745,1764,1791]" pageId="15" pageNumber="501">
<heading id="0104D65EFFAF983101C7AAB971CCF92F" bold="true" box="[151,745,1764,1791]" fontSize="11" level="1" pageId="15" pageNumber="501" reason="1">
<taxonomicName id="9DF31AB1FFAF983101C7AAB971CCF92F" authority="Keum, Jung and Joharchi" authorityName="Keum, Jung and Joharchi" box="[151,745,1764,1791]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="15" pageNumber="501" phylum="Arthropoda" rank="species" species="sejongi">
<emphasis id="6887BD20FFAF983101C7AAB971CCF92F" bold="true" box="[151,745,1764,1791]" pageId="15" pageNumber="501">
<emphasis id="6887BD20FFAF983101C7AAB972B9F92F" bold="true" box="[151,412,1764,1791]" italics="true" pageId="15" pageNumber="501">Cosmolaelaps sejongi</emphasis>
Keum, Jung and Joharchi
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="12E932B9FFAF983101C7AB74726EF8B7" pageId="15" pageNumber="501" type="reference_group">
<paragraph id="5A4C6132FFAF983101C7AB7471DDF890" blockId="15.[151,760,1833,1894]" box="[151,760,1833,1857]" pageId="15" pageNumber="501">
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<emphasis id="6887BD20FFAF983101C7AB747253F890" box="[151,374,1833,1857]" italics="true" pageId="15" pageNumber="501">Cosmolaelaps sejongi</emphasis>
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.
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</paragraph>
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(
<figureCitation id="C2C87DB7FFAF983101CFAB167261F8B7" box="[159,324,1867,1894]" captionStart="FIGURES 45–49" captionStartId="16.[151,264,1943,1967]" captionTargetBox="[180,1406,181,1920]" captionTargetId="figure-18@16.[180,1407,181,1920]" captionTargetPageId="16" captionText="FIGURES 45–49. DIC micrographs of Cosmolaelaps sejongi Keum, Jung and Joharchi, female. 45, idiosoma in dorsal view; 46, idiosoma in ventral view; 47, general view of some opisthonotal setae; 48, sternal and genital shields; 49, general view of opisthogastric area." figureDoi="http://doi.org/10.5281/zenodo.6530986" httpUri="https://zenodo.org/record/6530986/files/figure.png" pageId="15" pageNumber="501">Figures 45–49</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="12E932B9FFAF983101C7ABCE77F8F803" pageId="15" pageNumber="501" type="description">
<paragraph id="5A4C6132FFAF983101C7ABCE77F8F803" blockId="15.[151,1436,1939,2002]" pageId="15" pageNumber="501">
<emphasis id="6887BD20FFAF983101C7ABCE72B0F87C" bold="true" box="[151,405,1939,1965]" pageId="15" pageNumber="501">Specimens examined</emphasis>
. Six females,
<collectingRegion id="9837AFD0FFAF98310315ABCE7182F87C" box="[581,679,1939,1965]" country="Russia" name="Sakhalin" pageId="15" pageNumber="501">Sakhalin</collectingRegion>
Island,
<collectingCountry id="22E421A2FFAF9831025CABCE707CF87C" box="[780,857,1939,1965]" name="Russia" pageId="15" pageNumber="501">Russia</collectingCountry>
,
<geoCoordinate id="3FC707F5FFAF9831023AABCE70CBF87C" box="[874,1006,1939,1965]" degrees="46" direction="north" minutes="44" orientation="latitude" pageId="15" pageNumber="501" precision="15" seconds="06" value="46.735">46°44'06&quot;N</geoCoordinate>
<geoCoordinate id="3FC707F5FFAF983102AAABCE7785F87F" box="[1018,1184,1939,1966]" degrees="142" direction="east" minutes="12" orientation="longitude" pageId="15" pageNumber="501" precision="1" seconds="17.5" value="142.20486">142°12'17.5&quot;E</geoCoordinate>
,
<date id="2E4D47F2FFAF983105E2ABCE7655F87C" box="[1202,1392,1939,1965]" pageId="15" pageNumber="501" value="2021-08-11">11 August 2021</date>
, O. Joharchi coll., in the nest of
<taxonomicName id="9DF31AB1FFAF98310084ABE57034F803" authority="(Nylander)" baseAuthorityName="Nylander" baseAuthorityYear="1846" box="[468,785,1975,2002]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="501" phylum="Arthropoda" rank="species" species="umbratus">
<emphasis id="6887BD20FFAF98310084ABE571AAF800" box="[468,655,1975,2001]" italics="true" pageId="15" pageNumber="501">Lasius umbratus</emphasis>
(Nylander)
</taxonomicName>
(
<taxonomicName id="9DF31AB1FFAF9831024FABEA709BF800" box="[799,958,1975,2001]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="501" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
:
<taxonomicName id="9DF31AB1FFAF98310298ABEA776FF803" box="[968,1098,1975,2002]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="501" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
) (in TUMZ).
</paragraph>
</subSubSection>
<caption id="0E8C31BAFFB0982E01C7ABCA7278F826" ID-DOI="http://doi.org/10.5281/zenodo.6530986" ID-Zenodo-Dep="6530986" httpUri="https://zenodo.org/record/6530986/files/figure.png" pageId="16" pageNumber="502" startId="16.[151,264,1943,1967]" targetBox="[180,1406,181,1920]" targetPageId="16">
<paragraph id="5A4C6132FFB0982E01C7ABCA7278F826" blockId="16.[151,1436,1943,2040]" pageId="16" pageNumber="502">
<emphasis id="6887BD20FFB0982E01C7ABCA7270F87E" bold="true" box="[151,341,1943,1967]" pageId="16" pageNumber="502">FIGURES 45–49.</emphasis>
DIC micrographs of
<taxonomicName id="9DF31AB1FFB0982E0360ABC57739F861" authority="Keum, Jung and Joharchi" authorityName="Keum, Jung and Joharchi" box="[560,1052,1944,1968]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="16" pageNumber="502" phylum="Arthropoda" rank="species" species="sejongi">
<emphasis id="6887BD20FFB0982E0360ABC5702AF87E" box="[560,783,1944,1968]" italics="true" pageId="16" pageNumber="502">Cosmolaelaps sejongi</emphasis>
Keum, Jung and Joharchi
</taxonomicName>
, female. 45, idiosoma in dorsal view; 46, idiosoma in ventral view; 47, general view of some opisthonotal setae; 48, sternal and genital shields; 49, general view of opisthogastric area.
</paragraph>
</caption>
<subSubSection id="12E932B9FFB1982F0197ACCA7601FE7C" pageId="17" pageNumber="503" type="discussion">
<paragraph id="5A4C6132FFB1982F0197ACCA7601FE7C" blockId="17.[151,1437,151,429]" pageId="17" pageNumber="503">
<emphasis id="6887BD20FFB1982F0197ACCA7211FF60" bold="true" box="[199,308,151,177]" pageId="17" pageNumber="503">Remarks</emphasis>
.
<taxonomicName id="9DF31AB1FFB1982F0012ACCA711DFF61" box="[322,568,151,177]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="503" phylum="Arthropoda" rank="species" species="sejongi">
<emphasis id="6887BD20FFB1982F0012ACCA711DFF61" box="[322,568,151,177]" italics="true" pageId="17" pageNumber="503">Cosmolaelaps sejongi</emphasis>
</taxonomicName>
was described from
<collectingCountry id="22E421A2FFB1982F024FACCA70C8FF60" box="[799,1005,151,177]" name="South Korea" pageId="17" pageNumber="503">Republic of Korea</collectingCountry>
(
<bibRefCitation id="3E621CC3FFB1982F02ABACCA77EFFF60" author="Keum, E. &amp; Jung, C. &amp; Joharchi, O." box="[1019,1226,151,177]" pageId="17" pageNumber="503" pagination="485 - 505" refId="ref15495" refString="Keum, E., Jung, C. &amp; Joharchi, O. (2017) New species and new records of the family Laelapidae (Acari: Mesostigmata) from Republic of Korea. Zootaxa, 4353 (3), 485 - 505. https: // doi. org / 10.11646 / zootaxa. 4353.3.5" type="journal article" year="2017">
Keum
<emphasis id="6887BD20FFB1982F0518ACCA775CFF60" box="[1096,1145,151,177]" italics="true" pageId="17" pageNumber="503">et al</emphasis>
., 2017
</bibRefCitation>
). It has been found from soil around roots of an oak tree (
<bibRefCitation id="3E621CC3FFB1982F0368ACE67020FF04" author="Keum, E. &amp; Jung, C. &amp; Joharchi, O." box="[568,773,187,213]" pageId="17" pageNumber="503" pagination="485 - 505" refId="ref15495" refString="Keum, E., Jung, C. &amp; Joharchi, O. (2017) New species and new records of the family Laelapidae (Acari: Mesostigmata) from Republic of Korea. Zootaxa, 4353 (3), 485 - 505. https: // doi. org / 10.11646 / zootaxa. 4353.3.5" type="journal article" year="2017">
Keum
<emphasis id="6887BD20FFB1982F03D5ACE67190FF04" box="[645,693,187,213]" italics="true" pageId="17" pageNumber="503">et al</emphasis>
., 2017
</bibRefCitation>
), and is now recorded in
<collectingCountry id="22E421A2FFB1982F0549ACE67741FF04" box="[1049,1124,187,213]" name="Russia" pageId="17" pageNumber="503">Russia</collectingCountry>
for the first time, in the nest of
<taxonomicName id="9DF31AB1FFB1982F01E6AC82721AFF28" baseAuthorityName="Nylander" baseAuthorityYear="1846" box="[182,319,223,249]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="503" phylum="Arthropoda" rank="species" species="umbratus">
<emphasis id="6887BD20FFB1982F01E6AC8273E3FF29" box="[182,198,223,248]" italics="true" pageId="17" pageNumber="503">L</emphasis>
.
<emphasis id="6887BD20FFB1982F0185AC82721AFF28" box="[213,319,223,249]" italics="true" pageId="17" pageNumber="503">umbratus</emphasis>
</taxonomicName>
. Our specimens agree well with the description given by
<bibRefCitation id="3E621CC3FFB1982F029DAC82778CFF28" author="Keum, E. &amp; Jung, C. &amp; Joharchi, O." box="[973,1193,223,249]" pageId="17" pageNumber="503" pagination="485 - 505" refId="ref15495" refString="Keum, E., Jung, C. &amp; Joharchi, O. (2017) New species and new records of the family Laelapidae (Acari: Mesostigmata) from Republic of Korea. Zootaxa, 4353 (3), 485 - 505. https: // doi. org / 10.11646 / zootaxa. 4353.3.5" type="journal article" year="2017">
Keum
<emphasis id="6887BD20FFB1982F054AAC827769FF28" box="[1050,1100,223,249]" italics="true" pageId="17" pageNumber="503">et al</emphasis>
. (2017)
</bibRefCitation>
. The species is easily recognised by the distinctive shape of the post-anal seta (thickened, sparsely barbed and apically hook shaped), which is inserted on a small tubercle (
<figureCitation id="C2C87DB7FFB1982F031AAD7A71F5FE90" box="[586,720,295,321]" captionStart="FIGURES 45–49" captionStartId="16.[151,264,1943,1967]" captionTargetBox="[180,1406,181,1920]" captionTargetId="figure-18@16.[180,1407,181,1920]" captionTargetPageId="16" captionText="FIGURES 45–49. DIC micrographs of Cosmolaelaps sejongi Keum, Jung and Joharchi, female. 45, idiosoma in dorsal view; 46, idiosoma in ventral view; 47, general view of some opisthonotal setae; 48, sternal and genital shields; 49, general view of opisthogastric area." figureDoi="http://doi.org/10.5281/zenodo.6530986" httpUri="https://zenodo.org/record/6530986/files/figure.png" pageId="17" pageNumber="503">Figs 46, 49</figureCitation>
); dorsal shield setae thick and long, most setae long enough to reach well past the base of next posterior seta, and
<emphasis id="6887BD20FFB1982F03B4AD167027FEB4" box="[740,770,331,357]" italics="true" pageId="17" pageNumber="503">Z5</emphasis>
sparsely barbed, longer and thicker than
<emphasis id="6887BD20FFB1982F058DAD1677DDFEB4" box="[1245,1272,331,357]" italics="true" pageId="17" pageNumber="503">J5</emphasis>
(
<figureCitation id="C2C87DB7FFB1982F0459AD1676AAFEB4" box="[1289,1423,330,357]" captionStart="FIGURES 45–49" captionStartId="16.[151,264,1943,1967]" captionTargetBox="[180,1406,181,1920]" captionTargetId="figure-18@16.[180,1407,181,1920]" captionTargetPageId="16" captionText="FIGURES 45–49. DIC micrographs of Cosmolaelaps sejongi Keum, Jung and Joharchi, female. 45, idiosoma in dorsal view; 46, idiosoma in ventral view; 47, general view of some opisthonotal setae; 48, sternal and genital shields; 49, general view of opisthogastric area." figureDoi="http://doi.org/10.5281/zenodo.6530986" httpUri="https://zenodo.org/record/6530986/files/figure.png" pageId="17" pageNumber="503">Figs 45, 47</figureCitation>
); sternal setae short, not reaching base of following setae (
<figureCitation id="C2C87DB7FFB1982F024FAD327081FE58" box="[799,932,367,393]" captionStart="FIGURES 45–49" captionStartId="16.[151,264,1943,1967]" captionTargetBox="[180,1406,181,1920]" captionTargetId="figure-18@16.[180,1407,181,1920]" captionTargetPageId="16" captionText="FIGURES 45–49. DIC micrographs of Cosmolaelaps sejongi Keum, Jung and Joharchi, female. 45, idiosoma in dorsal view; 46, idiosoma in ventral view; 47, general view of some opisthonotal setae; 48, sternal and genital shields; 49, general view of opisthogastric area." figureDoi="http://doi.org/10.5281/zenodo.6530986" httpUri="https://zenodo.org/record/6530986/files/figure.png" pageId="17" pageNumber="503">Figs 46, 48</figureCitation>
); soft opisthogastric cuticle bearing 16 pairs setae,
<emphasis id="6887BD20FFB1982F018DADCE7221FE7C" box="[221,260,403,429]" italics="true" pageId="17" pageNumber="503">Jv5</emphasis>
well thickened, apically hook shaped and subequal in length with post-anal seta (
<figureCitation id="C2C87DB7FFB1982F05C5ADCE763DFE7C" box="[1173,1304,403,429]" captionStart="FIGURES 45–49" captionStartId="16.[151,264,1943,1967]" captionTargetBox="[180,1406,181,1920]" captionTargetId="figure-18@16.[180,1407,181,1920]" captionTargetPageId="16" captionText="FIGURES 45–49. DIC micrographs of Cosmolaelaps sejongi Keum, Jung and Joharchi, female. 45, idiosoma in dorsal view; 46, idiosoma in ventral view; 47, general view of some opisthonotal setae; 48, sternal and genital shields; 49, general view of opisthogastric area." figureDoi="http://doi.org/10.5281/zenodo.6530986" httpUri="https://zenodo.org/record/6530986/files/figure.png" pageId="17" pageNumber="503">Figs 46, 49</figureCitation>
).
</paragraph>
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</treatment>
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Sheals
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<emphasis id="6887BD20FFAF983101C7A8C57290FB61" box="[151,437,1176,1200]" italics="true" pageId="15" pageNumber="501">Cosmolaelaps rectangularis</emphasis>
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<emphasis id="6887BD20FFAF983101C7A8EA72B5FB1F" box="[151,400,1206,1230]" italics="true" pageId="15" pageNumber="501">Hypoaspis rectangularis</emphasis>
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.
<emphasis id="6887BD20FFAF983100C6A8EA7288FB1F" box="[406,429,1207,1230]" italics="true" pageId="15" pageNumber="501">—</emphasis>
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<emphasis id="6887BD20FFAF983101C7A88B7227FB3C" box="[151,258,1238,1261]" italics="true" pageId="15" pageNumber="501">Hypoaspis</emphasis>
(
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)
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</taxonomicName>
.—
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;
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;
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;
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<bibRefCitation id="3E621CC3FFAF98310197A8A97294FADD" author="Faraji, F. &amp; Abedi, L. &amp; Ostovan, H." box="[199,433,1268,1292]" pageId="15" pageNumber="501" pagination="205 - 209" refId="ref14412" refString="Faraji, F., Abedi, L. &amp; Ostovan, H. (2008) A new species of Hypoaspis Canestrini from Iran with a key to the Iranian species of Hypoaspis (Acari, Gamasina, Hypoaspididae). Zoosystematics and Evolution, 84, 205 - 209. https: // doi. org / 10.1002 / zoos. 200800005" type="journal article" year="2008">
Faraji
<emphasis id="6887BD20FFAF98310059A8A97210FADD" box="[265,309,1268,1292]" italics="true" pageId="15" pageNumber="501">et al</emphasis>
., 2008: 207
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<emphasis id="6887BD20FFAF983101C7A94F7290FAFB" box="[151,437,1298,1322]" italics="true" pageId="15" pageNumber="501">Cosmolaelaps rectangularis</emphasis>
</taxonomicName>
.—
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<bibRefCitation id="3E621CC3FFAF98310082A94F71F0FAFB" author="Moreira, G. F. &amp; Klompen, H. &amp; de Moraes, G. J." box="[466,725,1298,1322]" pageId="15" pageNumber="501" pagination="317 - 346" refId="ref15991" refString="Moreira, G. F., Klompen, H. &amp; de Moraes, G. J. (2014) Redefinition of Cosmolaelaps Berlese (Acari: Laelapidae) and description of five new species from Brazil. Zootaxa, 3764 (3), 317 - 346. https: // doi. org / 10.11646 / zootaxa. 3764.3.4" type="journal article" year="2014">
Moreira
<emphasis id="6887BD20FFAF9831037CA94E717CFAFB" box="[556,601,1298,1322]" italics="true" pageId="15" pageNumber="501">et al</emphasis>
., 2014: 320
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<emphasis id="6887BD20FFAF983101C7A90572B4FAA3" bold="true" box="[151,401,1368,1394]" pageId="15" pageNumber="501">Specimens examined</emphasis>
. Four females, vicinity of Uspenka,
<collectingRegion id="9837AFD0FFAF98310266A9047724FAA3" box="[822,1025,1368,1395]" country="Russia" name="Tyumen'" pageId="15" pageNumber="501">Tyumen Province</collectingRegion>
,
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,
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,
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, 20 May, 2020, O. Joharchi coll., in the nest of
<taxonomicName id="9DF31AB1FFAF9831036DA92071ECFA46" box="[573,713,1405,1431]" class="Insecta" family="Formicidae" genus="Myrmica" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="501" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="6887BD20FFAF9831036DA9207187FA47" box="[573,674,1405,1430]" italics="true" pageId="15" pageNumber="501">Myrmica</emphasis>
sp.
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(
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:
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).
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<emphasis id="6887BD20FFAF98310197A9FC721EFA6A" bold="true" box="[199,315,1441,1467]" pageId="15" pageNumber="501">Remarks.</emphasis>
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described
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<emphasis id="6887BD20FFAF98310302A9FD70A8FA6B" box="[594,909,1440,1466]" italics="true" pageId="15" pageNumber="501">Cosmolaelaps rectangularis</emphasis>
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as a new species from Patagonia,
<collectingCountry id="22E421A2FFAF98310454A9FD7653FA6B" box="[1284,1398,1440,1466]" name="Argentina" pageId="15" pageNumber="501">Argentina</collectingCountry>
, on the basis of
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specimen collected on an unspecified substrate. Subsequently,
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redescribed the species based on female specimens and described the male of this species for the first time from
<collectingCountry id="22E421A2FFAF98310598A9B5762FF9D3" box="[1224,1290,1512,1538]" name="Israel" pageId="15" pageNumber="501">Israel</collectingCountry>
. This species has been recorded from North America, Europe and Asia from soil-litter or ant nests and is now recorded in
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for the first time, from a nest of
<taxonomicName id="9DF31AB1FFAF98310351AA6C71ADF99A" box="[513,648,1585,1611]" class="Insecta" family="Formicidae" genus="Myrmica" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="501" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="6887BD20FFAF98310351AA6C7143F99B" box="[513,614,1585,1610]" italics="true" pageId="15" pageNumber="501">Myrmica</emphasis>
sp
</taxonomicName>
.. Our newly collected material agrees very well with the redescription given by
<bibRefCitation id="3E621CC3FFAF98310054AA0872BEF9BE" author="Costa, M." box="[260,411,1620,1647]" pageId="15" pageNumber="501" pagination="1 - 30" refId="ref13901" refString="Costa, M. (1968) Little known and new litter-inhabiting Laelapine mites (Acari, Mesostigmata) from Israel. Israel Journal of Zoology, 17, 1 - 30." type="journal article" year="1968">Costa (1968)</bibRefCitation>
. This species can be readily recognised by the 39 pairs of spatulate-mucronate-tricarinate dorsal setae.
</paragraph>
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</treatment>
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315
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<subSubSection id="12E932B9FFAF983101C7ACCA714CFF60" box="[151,617,151,178]" pageId="15" pageNumber="501" type="nomenclature">
<paragraph id="5A4C6132FFAF983101C7ACCA714CFF60" blockId="15.[151,617,151,178]" box="[151,617,151,178]" pageId="15" pageNumber="501">
<heading id="0104D65EFFAF983101C7ACCA714CFF60" bold="true" box="[151,617,151,178]" fontSize="11" level="1" pageId="15" pageNumber="501" reason="1">
<taxonomicName id="9DF31AB1FFAF983101C7ACCA714CFF60" authority="(Shcherbak)" baseAuthorityName="Shcherbak" baseAuthorityYear="1971" box="[151,617,151,178]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="15" pageNumber="501" phylum="Arthropoda" rank="species" species="lutegiensis">
<emphasis id="6887BD20FFAF983101C7ACCA714CFF60" bold="true" box="[151,617,151,178]" pageId="15" pageNumber="501">
<emphasis id="6887BD20FFAF983101C7ACCA72E2FF63" bold="true" box="[151,455,151,178]" italics="true" pageId="15" pageNumber="501">Cosmolaelaps lutegiensis</emphasis>
(Shcherbak)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
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<treatmentCitation id="DB524723FFAF983101C7AC81717FFF22" author="Shcherbak, G. I." box="[151,602,219,244]" page="76" pageId="15" pageNumber="501" year="1971">
<taxonomicName id="9DF31AB1FFAF983101C7AC81717FFF22" authority="Shcherbak, 1971 a: 76" authorityName="Shcherbak" authorityPageNumber="76" authorityYear="1971" box="[151,602,219,244]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="15" pageNumber="501" phylum="Arthropoda" rank="species" species="lutegiensis">
<emphasis id="6887BD20FFAF983101C7AC817251FF25" box="[151,372,220,244]" italics="true" pageId="15" pageNumber="501">Hypoaspis lutegiensis</emphasis>
<bibRefCitation id="3E621CC3FFAF9831002BAC81717FFF22" author="Shcherbak, G. I." box="[379,602,219,244]" pageId="15" pageNumber="501" pagination="76 - 79" refId="ref16468" refString="Shcherbak, G. I. (1971 a) New species of gamasid from the genus Hypoaspis Canestrini (Acarina, Gamasoidea). Vestnik Zoologii, 5, 76 - 79. [in Russian]" type="journal article" year="1971">Shcherbak, 1971a: 76</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
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<treatmentCitationGroup id="7AE3461CFFAF983101C7ACA77265FEE0" pageId="15" pageNumber="501">
<taxonomicName id="9DF31AB1FFAF983101C7ACA772BFFEC3" baseAuthorityName="Shcherbak" baseAuthorityYear="1971" box="[151,410,250,274]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="15" pageNumber="501" phylum="Arthropoda" rank="species" species="lutegiensis">
<emphasis id="6887BD20FFAF983101C7ACA772BFFEC3" box="[151,410,250,274]" italics="true" pageId="15" pageNumber="501">Cosmolaelaps lutegiensis</emphasis>
</taxonomicName>
.—
<treatmentCitation id="DB524723FFAF983100E7ACA77192FEC3" author="Moreira, G. F. &amp; Klompen, H. &amp; de Moraes, G. J." box="[439,695,250,274]" page="319" pageId="15" pageNumber="501" year="2014">
<bibRefCitation id="3E621CC3FFAF983100E7ACA77192FEC3" author="Moreira, G. F. &amp; Klompen, H. &amp; de Moraes, G. J." box="[439,695,250,274]" pageId="15" pageNumber="501" pagination="317 - 346" refId="ref15991" refString="Moreira, G. F., Klompen, H. &amp; de Moraes, G. J. (2014) Redefinition of Cosmolaelaps Berlese (Acari: Laelapidae) and description of five new species from Brazil. Zootaxa, 3764 (3), 317 - 346. https: // doi. org / 10.11646 / zootaxa. 3764.3.4" type="journal article" year="2014">
Moreira
<emphasis id="6887BD20FFAF98310343ACA67165FEC3" box="[531,576,250,274]" italics="true" pageId="15" pageNumber="501">et al</emphasis>
. 2014: 319
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="DB524723FFAF98310392ACA770F6FEC3" author="Ramroodi, S. &amp; Hajizadeh, J. &amp; Joharchi, O." box="[706,979,250,274]" page="542" pageId="15" pageNumber="501" year="2014">
<bibRefCitation id="3E621CC3FFAF98310392ACA770F6FEC3" author="Ramroodi, S. &amp; Hajizadeh, J. &amp; Joharchi, O." box="[706,979,250,274]" pageId="15" pageNumber="501" pagination="533 - 544" refId="ref16273" refString="Ramroodi, S., Hajizadeh, J. &amp; Joharchi, O. (2014) Two new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Iran. Zootaxa, 3847 (4), 533 - 544. https: // doi. org / 10.11646 / zootaxa. 3847.4.3" type="journal article" year="2014">
Ramroodi
<emphasis id="6887BD20FFAF9831027FACA67079FEC3" box="[815,860,250,274]" italics="true" pageId="15" pageNumber="501">et al</emphasis>
. 2014: 542
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="DB524723FFAF9831028EACA77624FEC3" author="Joharchi, O &amp; Trach, V. A." box="[990,1281,250,274]" page="486" pageId="15" pageNumber="501" year="2019">
<bibRefCitation id="3E621CC3FFAF9831028EACA77624FEC3" author="Joharchi, O &amp; Trach, V. A." box="[990,1281,250,274]" pageId="15" pageNumber="501" pagination="486 - 494" refId="ref14677" refString="Joharchi, O &amp; Trach, V. A. (2019) A new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Ukraine. Zootaxa, 4647 (1), 486 - 494. https: // doi. org / 10.11646 / zootaxa. 4647.1.30" type="journal article" year="2019">Joharchi &amp; Trach 2019: 486</bibRefCitation>
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;
<treatmentCitation id="DB524723FFAF9831045CACA7721EFEE0" ID-DOI="http://doi.org/10.5281/zenodo.4441023" ID-Zenodo-Dep="4441023" author="Joharchi, O. &amp; Issakova, A. K. &amp; Asyamova, O. S. &amp; Abbasi-Sarcheshmeh, M. &amp; Tolstikov, A. V." httpUri="http://treatment.plazi.org/id/039287E3B00F985EF0815624FC47FDAA" page="479" pageId="15" pageNumber="501" year="2020">
<bibRefCitation id="3E621CC3FFAF9831045CACA7721EFEE0" author="Joharchi, O. &amp; Issakova, A. K. &amp; Asyamova, O. S. &amp; Abbasi-Sarcheshmeh, M. &amp; Tolstikov, A. V." pageId="15" pageNumber="501" pagination="473 - 498" refId="ref14853" refString="Joharchi, O., Issakova, A. K., Asyamova, O. S., Abbasi-Sarcheshmeh, M. &amp; Tolstikov, A. V. (2020 a) Some soil-inhabiting mites (Acari: Mesostigmata) from Kazakhstan, with description of a new species of Gaeolaelaps Evans &amp; Till (Acari: Laelapidae). Zootaxa, 4819 (3), 473 - 498. https: // doi. org / 10.11646 / zootaxa. 4819.3.3" type="journal article" year="2020">
Joharchi
<emphasis id="6887BD20FFAF98310439ACA676B3FEC3" box="[1385,1430,250,274]" italics="true" pageId="15" pageNumber="501">et al</emphasis>
. 2020a: 479
</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="5A4C6132FFAF983101C7AD02708DFE4C" blockId="15.[151,1437,351,1026]" pageId="15" pageNumber="501">
<emphasis id="6887BD20FFAF983101C7AD0272A8FEA8" bold="true" box="[151,397,351,377]" pageId="15" pageNumber="501">Specimens examined</emphasis>
.
<materialsCitation id="EA9B6B6FFFAF983100C8AD027081FE4C" ID-GBIF-Occurrence="3763889305" collectingDate="2021-06-13" collectionCode="TSUMZ" collectorName="O. Joharchi" country="Russia" county="Western Crimea" latitude="45.191944" location="Five" longLatPrecision="1" longitude="33.333054" municipality="Yevpatoria" pageId="15" pageNumber="501" specimenCount="1">
<location id="5F2C37E9FFAF983100C8AD0272EFFEA8" LSID="urn:lsid:plazi:treatment:D25AD024FFAF983101C7ACCA701EFBD0:5F2C37E9FFAF983100C8AD0272EFFEA8" box="[408,458,351,377]" country="Russia" county="Western Crimea" latitude="45.191944" longLatPrecision="1" longitude="33.333054" municipality="Yevpatoria" name="Five" pageId="15" pageNumber="501">Five</location>
females, near the
<location id="5F2C37E9FFAF983103DDAD027040FEA8" LSID="urn:lsid:plazi:treatment:D25AD024FFAF983101C7ACCA701EFBD0:5F2C37E9FFAF983103DDAD027040FEA8" box="[653,869,351,377]" country="Russia" county="Western Crimea" latitude="45.191944" longLatPrecision="1" longitude="33.333054" municipality="Yevpatoria" name="Moinakskoye Lake" pageId="15" pageNumber="501">Moinakskoye Lake</location>
,
<collectingMunicipality id="BA28FB48FFAF9831023CAD0270CCFEA8" box="[876,1001,351,377]" pageId="15" pageNumber="501">Yevpatoria</collectingMunicipality>
,
<collectingCounty id="B32D19BEFFAF983102A1AD3D7780FEA8" box="[1009,1189,351,378]" pageId="15" pageNumber="501">Western Crimea</collectingCounty>
,
<collectingCountry id="22E421A2FFAF983105FEAD0277DEFEA8" box="[1198,1275,351,377]" name="Russia" pageId="15" pageNumber="501">Russia</collectingCountry>
,
<geoCoordinate id="3FC707F5FFAF98310454AD0276B9FEAB" box="[1284,1436,351,378]" degrees="45" direction="north" minutes="11" orientation="latitude" pageId="15" pageNumber="501" precision="1" seconds="31.0" value="45.191944">45°11'31.0&quot;N</geoCoordinate>
<geoCoordinate id="3FC707F5FFAF983101C7ADDE720AFE4F" box="[151,303,387,414]" degrees="33" direction="east" minutes="19" orientation="longitude" pageId="15" pageNumber="501" precision="1" seconds="59.0" value="33.333054">33°19'59.0&quot;E</geoCoordinate>
,
<date id="2E4D47F2FFAF9831006BADDE72F6FE4C" box="[315,467,387,414]" pageId="15" pageNumber="501" value="2021-06-13">
<collectingDate id="3E09BE1AFFAF9831006BADDE72F6FE4C" box="[315,467,387,414]" pageId="15" pageNumber="501" value="2021-06-13">13 June 2021</collectingDate>
</date>
,
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coll., from soil (in
<collectionCode id="3CE2F9F7FFAF98310268ADDE70B9FE4C" box="[824,924,387,413]" pageId="15" pageNumber="501">TSUMZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="12E932B9FFAF98310197ADFA701EFBD0" pageId="15" pageNumber="501" type="discussion">
<paragraph id="5A4C6132FFAF98310197ADFA701EFBD0" blockId="15.[151,1437,351,1026]" pageId="15" pageNumber="501">
<emphasis id="6887BD20FFAF98310197ADFA721EFE10" bold="true" box="[199,315,423,449]" pageId="15" pageNumber="501">Remarks.</emphasis>
<taxonomicName id="9DF31AB1FFAF98310012ADFA7145FE10" baseAuthorityName="Shcherbak" baseAuthorityYear="1971" box="[322,608,423,449]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="15" pageNumber="501" phylum="Arthropoda" rank="species" species="lutegiensis">
<emphasis id="6887BD20FFAF98310012ADFA7145FE10" box="[322,608,423,449]" italics="true" pageId="15" pageNumber="501">Cosmolaelaps lutegiensis</emphasis>
</taxonomicName>
was described from
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,
<collectingCountry id="22E421A2FFAF983102DEADFA70CDFE10" box="[910,1000,423,449]" name="Ukraine" pageId="15" pageNumber="501">Ukraine</collectingCountry>
(
<bibRefCitation id="3E621CC3FFAF983102A7ADFA77E3FE10" author="Shcherbak, G. I." box="[1015,1222,423,449]" pageId="15" pageNumber="501" pagination="76 - 79" refId="ref16468" refString="Shcherbak, G. I. (1971 a) New species of gamasid from the genus Hypoaspis Canestrini (Acarina, Gamasoidea). Vestnik Zoologii, 5, 76 - 79. [in Russian]" type="journal article" year="1971">Shcherbak, 1971a</bibRefCitation>
). It has been found from soil-litter of a pine forest and is now recorded from
<collectingCountry id="22E421A2FFAF9831025FAD96707FFE34" box="[783,858,459,485]" name="Russia" pageId="15" pageNumber="501">Russia</collectingCountry>
for the first time, from the soil. Our newly collected material agrees very well with the description given by
<bibRefCitation id="3E621CC3FFAF9831024DADB270DDFDD8" author="Shcherbak, G. I." box="[797,1016,495,521]" pageId="15" pageNumber="501" pagination="76 - 79" refId="ref16468" refString="Shcherbak, G. I. (1971 a) New species of gamasid from the genus Hypoaspis Canestrini (Acarina, Gamasoidea). Vestnik Zoologii, 5, 76 - 79. [in Russian]" type="journal article" year="1971">Shcherbak (1971a)</bibRefCitation>
.
<bibRefCitation id="3E621CC3FFAF98310555ADB27635FDD8" author="Ramroodi, S. &amp; Hajizadeh, J. &amp; Joharchi, O." box="[1029,1296,495,522]" pageId="15" pageNumber="501" pagination="533 - 544" refId="ref16273" refString="Ramroodi, S., Hajizadeh, J. &amp; Joharchi, O. (2014) Two new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Iran. Zootaxa, 3847 (4), 533 - 544. https: // doi. org / 10.11646 / zootaxa. 3847.4.3" type="journal article" year="2014">
Ramroodi
<emphasis id="6887BD20FFAF9831052FADAD7797FDD8" box="[1151,1202,495,521]" italics="true" pageId="15" pageNumber="501">et al</emphasis>
. (2014)
</bibRefCitation>
,
<bibRefCitation id="3E621CC3FFAF9831044EADB2720FFDFC" author="Joharchi, O &amp; Trach, V. A." pageId="15" pageNumber="501" pagination="486 - 494" refId="ref14677" refString="Joharchi, O &amp; Trach, V. A. (2019) A new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Ukraine. Zootaxa, 4647 (1), 486 - 494. https: // doi. org / 10.11646 / zootaxa. 4647.1.30" type="journal article" year="2019">Joharchi &amp; Trach (2019)</bibRefCitation>
, and
<bibRefCitation id="3E621CC3FFAF98310033AE4E717EFDFC" author="Joharchi, O. &amp; Issakova, A. K. &amp; Asyamova, O. S. &amp; Abbasi-Sarcheshmeh, M. &amp; Tolstikov, A. V." box="[355,603,531,558]" pageId="15" pageNumber="501" pagination="473 - 498" refId="ref14853" refString="Joharchi, O., Issakova, A. K., Asyamova, O. S., Abbasi-Sarcheshmeh, M. &amp; Tolstikov, A. V. (2020 a) Some soil-inhabiting mites (Acari: Mesostigmata) from Kazakhstan, with description of a new species of Gaeolaelaps Evans &amp; Till (Acari: Laelapidae). Zootaxa, 4819 (3), 473 - 498. https: // doi. org / 10.11646 / zootaxa. 4819.3.3" type="journal article" year="2020">
Joharchi
<emphasis id="6887BD20FFAF98310097AE4972D2FDFC" box="[455,503,531,557]" italics="true" pageId="15" pageNumber="501">et al</emphasis>
. (2020a)
</bibRefCitation>
have reported the species from
<collectingCountry id="22E421A2FFAF983102E9AE4970CCFDFF" box="[953,1001,532,558]" name="Iran" pageId="15" pageNumber="501">Iran</collectingCountry>
,
<collectingCountry id="22E421A2FFAF983102A2AE4E7769FDFC" box="[1010,1100,531,557]" name="Ukraine" pageId="15" pageNumber="501">Ukraine</collectingCountry>
and
<collectingCountry id="22E421A2FFAF983105D0AE4E7623FDFC" box="[1152,1286,531,557]" name="Kazakhstan" pageId="15" pageNumber="501">Kazakhstan</collectingCountry>
, respectively. They stated inaccurately that the dorsal shield has 38 pairs of lancet-like setae, but there are 39 pairs (including two pairs of
<emphasis id="6887BD20FFAF98310073AE01721AFDA4" box="[291,319,604,629]" italics="true" pageId="15" pageNumber="501">Zx</emphasis>
setae between
<emphasis id="6887BD20FFAF983100B8AE0172D1FDA4" box="[488,500,604,629]" italics="true" pageId="15" pageNumber="501">J</emphasis>
and
<emphasis id="6887BD20FFAF9831037BAE01711EFDA4" box="[555,571,604,629]" italics="true" pageId="15" pageNumber="501">Z</emphasis>
setae, and three unpaired setae
<emphasis id="6887BD20FFAF983102CCAE017090FDA4" box="[924,949,604,629]" italics="true" pageId="15" pageNumber="501">Jx</emphasis>
between
<emphasis id="6887BD20FFAF9831054FAE06771FFDA4" box="[1055,1082,603,629]" italics="true" pageId="15" pageNumber="501">J3</emphasis>
and
<emphasis id="6887BD20FFAF9831053FAE0177AFFDA7" box="[1135,1162,604,630]" italics="true" pageId="15" pageNumber="501">J5</emphasis>
) as stated in the original description (see
<bibRefCitation id="3E621CC3FFAF9831001DAE22713DFD48" author="Shcherbak, G. I." box="[333,536,639,665]" pageId="15" pageNumber="501" pagination="76 - 79" refId="ref16468" refString="Shcherbak, G. I. (1971 a) New species of gamasid from the genus Hypoaspis Canestrini (Acarina, Gamasoidea). Vestnik Zoologii, 5, 76 - 79. [in Russian]" type="journal article" year="1971">Shcherbak, 1971a</bibRefCitation>
).
<bibRefCitation id="3E621CC3FFAF9831037BAEDD71CBFD48" author="Bregetova, N. G." box="[555,750,639,666]" pageId="15" pageNumber="501" pagination="483 - 554" refId="ref13808" refString="Bregetova, N. G. (1977) Family Laelaptidae Berlese, 1892. In: Ghilyarov, M. S. &amp; Bregetova, N. G. (Eds.) Key to the Soil Inhabiting Mites. Mesostigmata. Nauka, Leningrad, pp. 483 - 554. [in Russian]" type="book chapter" year="1977">Bregetova (1977)</bibRefCitation>
suspected
<taxonomicName id="9DF31AB1FFAF98310237AE2277A1FD48" baseAuthorityName="Shcherbak" baseAuthorityYear="1971" box="[871,1156,639,665]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="15" pageNumber="501" phylum="Arthropoda" rank="species" species="lutegiensis">
<emphasis id="6887BD20FFAF98310237AE2277A1FD48" box="[871,1156,639,665]" italics="true" pageId="15" pageNumber="501">Cosmolaelaps lutegiensis</emphasis>
</taxonomicName>
maybe a junior synonym of
<taxonomicName id="9DF31AB1FFAF983101E6AEF972DCFD6F" authority="Samsinak, 1958" authorityName="Samsinak" authorityYear="1958" box="[182,505,675,702]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="15" pageNumber="501" phylum="Arthropoda" rank="species" species="helianthi">
<emphasis id="6887BD20FFAF983101E6AEF973EFFD6C" box="[182,202,676,701]" italics="true" pageId="15" pageNumber="501">H</emphasis>
.
<emphasis id="6887BD20FFAF98310188AEFE7219FD6C" box="[216,316,675,701]" italics="true" pageId="15" pageNumber="501">helianthi</emphasis>
<bibRefCitation id="3E621CC3FFAF98310013AEFE72DCFD6F" author="Samsinak, K." box="[323,505,675,702]" pageId="15" pageNumber="501" pagination="185 - 198" refId="ref16440" refString="Samsinak, K. (1958) Roztoci ze slunecnicovych semen (Acari). Ceskoslovenska Parasitologie, 5, 185 - 198. [in Czech]" type="journal article" year="1958">Samšiňák, 1958</bibRefCitation>
</taxonomicName>
. We have not had the opportunity to examine the
<typeStatus id="8548DF90FFAF98310577AEF97772FD6F" box="[1063,1111,676,702]" pageId="15" pageNumber="501">type</typeStatus>
specimens of
<taxonomicName id="9DF31AB1FFAF983105A9AEF976A5FD6C" authorityName="Samsinak" authorityYear="1958" box="[1273,1408,675,702]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="15" pageNumber="501" phylum="Arthropoda" rank="species" species="helianthi">
<emphasis id="6887BD20FFAF983105A9AEF97628FD6C" box="[1273,1293,676,701]" italics="true" pageId="15" pageNumber="501">H</emphasis>
.
<emphasis id="6887BD20FFAF9831044CAEFE76A5FD6C" box="[1308,1408,675,701]" italics="true" pageId="15" pageNumber="501">helianthi</emphasis>
</taxonomicName>
in order to confirm that synonymy, but by comparing the descriptions and figures of these two species, we found some distinguishing morphological differences: (1) sternal seta
<emphasis id="6887BD20FFAF9831024DAEB6701BFCD4" box="[797,830,747,773]" italics="true" pageId="15" pageNumber="501">st1</emphasis>
is off the sternal shield in
<taxonomicName id="9DF31AB1FFAF98310536AEB177C9FCD4" authorityName="Samsinak" authorityYear="1958" box="[1126,1260,747,774]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="15" pageNumber="501" phylum="Arthropoda" rank="species" species="helianthi">
<emphasis id="6887BD20FFAF98310536AEB1775FFCD4" box="[1126,1146,748,773]" italics="true" pageId="15" pageNumber="501">H</emphasis>
.
<emphasis id="6887BD20FFAF983105D8AEB677C9FCD4" box="[1160,1260,747,773]" italics="true" pageId="15" pageNumber="501">helianthi</emphasis>
</taxonomicName>
(see
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Figure 4-
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</figureCitation>
in
<bibRefCitation id="3E621CC3FFAF983101E5AF52724EFCFB" author="Samsinak, K." box="[181,363,783,810]" pageId="15" pageNumber="501" pagination="185 - 198" refId="ref16440" refString="Samsinak, K. (1958) Roztoci ze slunecnicovych semen (Acari). Ceskoslovenska Parasitologie, 5, 185 - 198. [in Czech]" type="journal article" year="1958">Samšiňák, 1958</bibRefCitation>
), while in
<taxonomicName id="9DF31AB1FFAF983100B4AF4D715BFCF8" baseAuthorityName="Shcherbak" baseAuthorityYear="1971" box="[484,638,783,810]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="15" pageNumber="501" phylum="Arthropoda" rank="species" species="lutegiensis">
<emphasis id="6887BD20FFAF983100B4AF4D72D2FCFB" box="[484,503,784,810]" italics="true" pageId="15" pageNumber="501">C</emphasis>
.
<emphasis id="6887BD20FFAF98310356AF52715BFCF8" box="[518,638,783,809]" italics="true" pageId="15" pageNumber="501">lutegiensis</emphasis>
</taxonomicName>
it is obviously on the shield (see
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Fig.
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</figureCitation>
in
<bibRefCitation id="3E621CC3FFAF98310520AF52764DFCF8" author="Joharchi, O. &amp; Issakova, A. K. &amp; Asyamova, O. S. &amp; Abbasi-Sarcheshmeh, M. &amp; Tolstikov, A. V." box="[1136,1384,783,810]" pageId="15" pageNumber="501" pagination="473 - 498" refId="ref14853" refString="Joharchi, O., Issakova, A. K., Asyamova, O. S., Abbasi-Sarcheshmeh, M. &amp; Tolstikov, A. V. (2020 a) Some soil-inhabiting mites (Acari: Mesostigmata) from Kazakhstan, with description of a new species of Gaeolaelaps Evans &amp; Till (Acari: Laelapidae). Zootaxa, 4819 (3), 473 - 498. https: // doi. org / 10.11646 / zootaxa. 4819.3.3" type="journal article" year="2020">
Joharchi
<emphasis id="6887BD20FFAF98310587AF4D762CFCF8" box="[1239,1289,783,809]" italics="true" pageId="15" pageNumber="501">et al</emphasis>
., 2020a
</bibRefCitation>
); (2) dorsal setae smooth and mostly do not reach to base of next setae in series (see
<figureCitation id="C2C87DB7FFAF98310550AF6E7751FC9F" box="[1024,1140,819,846]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1944,1968]" captionTargetBox="[182,1405,181,1920]" captionTargetId="figure-18@2.[182,1405,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–7. DIC micrographs of Cosmolaelaps chianensis (Gu), female. 1, idiosoma in dorsal view, enlarged section not to scale; 2, idiosoma in ventral view; 3, sternal shield; 4, general view of opisthogastric area; 5, chelicera; 6, dorsal aspect of femur, genu and tibia leg I; 7, ventral aspect of femur, genu and tibia leg I." figureDoi="http://doi.org/10.5281/zenodo.6530960" httpUri="https://zenodo.org/record/6530960/files/figure.png" pageId="15" pageNumber="501">
Figure 5-
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</figureCitation>
in
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), while in
<taxonomicName id="9DF31AB1FFAF983101E5AF05726AFCA0" baseAuthorityName="Shcherbak" baseAuthorityYear="1971" box="[181,335,855,882]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="15" pageNumber="501" phylum="Arthropoda" rank="species" species="lutegiensis">
<emphasis id="6887BD20FFAF983101E5AF0573EDFCA3" box="[181,200,856,882]" italics="true" pageId="15" pageNumber="501">C</emphasis>
.
<emphasis id="6887BD20FFAF98310187AF0A726AFCA0" box="[215,335,855,881]" italics="true" pageId="15" pageNumber="501">lutegiensis</emphasis>
</taxonomicName>
all setae have a small basal protuberance and are mostly long enough to reach base of next setae in series (see
<figureCitation id="C2C87DB7FFAF98310062AF267288FC44" box="[306,429,891,917]" captionStart="FIGURES 13–20" captionStartId="5.[151,264,1908,1932]" captionTargetBox="[198,1388,181,1884]" captionTargetId="figure-19@5.[198,1389,181,1884]" captionTargetPageId="5" captionText="FIGURES 13–20. DIC micrographs of Cosmolaelaps lasiophilus Joharchi sp. nov., female. 13, idiosoma in dorsal view; 14, idiosoma in ventral view; 15, general view of some podonotal setae; 16, general view of some opisthonotal setae; 17, general view of part of opisthogastric area; 18, subcapitulum; 19, distal portion of palp, with focus on tarsal claw; 20, insemination structures." figureDoi="http://doi.org/10.5281/zenodo.6530968" httpUri="https://zenodo.org/record/6530968/files/figure.png" pageId="15" pageNumber="501">
Figures
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</figureCitation>
,
<figureCitation id="C2C87DB7FFAF983100E9AF2672F0FC44" box="[441,469,891,917]" captionStart="FIGURE 21–24" captionStartId="6.[151,250,1926,1950]" captionTargetBox="[161,1425,197,1887]" captionTargetId="figure-18@6.[152,1436,181,1902]" captionTargetPageId="6" captionText="FIGURE 21–24. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 21, leg I (trochanter-tibia); 22, leg II; 23, leg III; 24, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530972" httpUri="https://zenodo.org/record/6530972/files/figure.png" pageId="15" pageNumber="501">22</figureCitation>
in
<bibRefCitation id="3E621CC3FFAF983100ABAF2671D6FC44" author="Joharchi, O. &amp; Issakova, A. K. &amp; Asyamova, O. S. &amp; Abbasi-Sarcheshmeh, M. &amp; Tolstikov, A. V." box="[507,755,891,918]" pageId="15" pageNumber="501" pagination="473 - 498" refId="ref14853" refString="Joharchi, O., Issakova, A. K., Asyamova, O. S., Abbasi-Sarcheshmeh, M. &amp; Tolstikov, A. V. (2020 a) Some soil-inhabiting mites (Acari: Mesostigmata) from Kazakhstan, with description of a new species of Gaeolaelaps Evans &amp; Till (Acari: Laelapidae). Zootaxa, 4819 (3), 473 - 498. https: // doi. org / 10.11646 / zootaxa. 4819.3.3" type="journal article" year="2020">
Joharchi
<emphasis id="6887BD20FFAF98310332AF2171B1FC44" box="[610,660,891,917]" italics="true" pageId="15" pageNumber="501">et al</emphasis>
., 2020a
</bibRefCitation>
); (3) sternal shield with reticulate ornamentation throughout
<taxonomicName id="9DF31AB1FFAF983101C7AFFD723AFC68" authorityName="Samsinak" authorityYear="1958" box="[151,287,927,954]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="15" pageNumber="501" phylum="Arthropoda" rank="species" species="helianthi">
<emphasis id="6887BD20FFAF983101C7AFFD738EFC68" box="[151,171,928,953]" italics="true" pageId="15" pageNumber="501">H</emphasis>
.
<emphasis id="6887BD20FFAF983101EBAFC2723AFC68" box="[187,287,927,953]" italics="true" pageId="15" pageNumber="501">helianthi</emphasis>
</taxonomicName>
(see
<figureCitation id="C2C87DB7FFAF9831000DAFC272F1FC68" box="[349,468,927,953]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1944,1968]" captionTargetBox="[182,1405,181,1920]" captionTargetId="figure-18@2.[182,1405,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–7. DIC micrographs of Cosmolaelaps chianensis (Gu), female. 1, idiosoma in dorsal view, enlarged section not to scale; 2, idiosoma in ventral view; 3, sternal shield; 4, general view of opisthogastric area; 5, chelicera; 6, dorsal aspect of femur, genu and tibia leg I; 7, ventral aspect of femur, genu and tibia leg I." figureDoi="http://doi.org/10.5281/zenodo.6530960" httpUri="https://zenodo.org/record/6530960/files/figure.png" pageId="15" pageNumber="501">
Figure 4-
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</figureCitation>
in
<bibRefCitation id="3E621CC3FFAF983100AAAFC27197FC6B" author="Samsinak, K." box="[506,690,927,954]" pageId="15" pageNumber="501" pagination="185 - 198" refId="ref16440" refString="Samsinak, K. (1958) Roztoci ze slunecnicovych semen (Acari). Ceskoslovenska Parasitologie, 5, 185 - 198. [in Czech]" type="journal article" year="1958">Samšiňák, 1958</bibRefCitation>
), while in
<taxonomicName id="9DF31AB1FFAF9831027DAFFD70E2FC68" baseAuthorityName="Shcherbak" baseAuthorityYear="1971" box="[813,967,927,954]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="15" pageNumber="501" phylum="Arthropoda" rank="species" species="lutegiensis">
<emphasis id="6887BD20FFAF9831027DAFFD7065FC6B" box="[813,832,928,954]" italics="true" pageId="15" pageNumber="501">C</emphasis>
.
<emphasis id="6887BD20FFAF9831021FAFC270E2FC68" box="[847,967,927,953]" italics="true" pageId="15" pageNumber="501">lutegiensis</emphasis>
</taxonomicName>
shield smooth almost throughout, except a longitudinally lateral part between
<emphasis id="6887BD20FFAF98310365AF9E7172FC0C" box="[565,599,963,989]" italics="true" pageId="15" pageNumber="501">iv1</emphasis>
and
<emphasis id="6887BD20FFAF983103DEAF9E718AFC0C" box="[654,687,963,989]" italics="true" pageId="15" pageNumber="501">st3</emphasis>
faintly reticulated with curved lines (see
<figureCitation id="C2C87DB7FFAF983105D2AF9E77F7FC0C" box="[1154,1234,963,989]" captionStart="FIGURE 21–24" captionStartId="6.[151,250,1926,1950]" captionTargetBox="[161,1425,197,1887]" captionTargetId="figure-18@6.[152,1436,181,1902]" captionTargetPageId="6" captionText="FIGURE 21–24. Cosmolaelaps lasiophilus Joharchi sp. nov., female. 21, leg I (trochanter-tibia); 22, leg II; 23, leg III; 24, leg IV." figureDoi="http://doi.org/10.5281/zenodo.6530972" httpUri="https://zenodo.org/record/6530972/files/figure.png" pageId="15" pageNumber="501">
Fig.
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</figureCitation>
in
<bibRefCitation id="3E621CC3FFAF983105A6AF9E73C4FBD0" author="Joharchi, O. &amp; Issakova, A. K. &amp; Asyamova, O. S. &amp; Abbasi-Sarcheshmeh, M. &amp; Tolstikov, A. V." pageId="15" pageNumber="501" pagination="473 - 498" refId="ref14853" refString="Joharchi, O., Issakova, A. K., Asyamova, O. S., Abbasi-Sarcheshmeh, M. &amp; Tolstikov, A. V. (2020 a) Some soil-inhabiting mites (Acari: Mesostigmata) from Kazakhstan, with description of a new species of Gaeolaelaps Evans &amp; Till (Acari: Laelapidae). Zootaxa, 4819 (3), 473 - 498. https: // doi. org / 10.11646 / zootaxa. 4819.3.3" type="journal article" year="2020">
Joharchi
<emphasis id="6887BD20FFAF9831040DAF9976ABFC0C" box="[1373,1422,963,989]" italics="true" pageId="15" pageNumber="501">et al</emphasis>
., 2020a
</bibRefCitation>
). Therefore, we believe these are two distinct species.
</paragraph>
</subSubSection>
</treatment>
</document>

444
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<mods:title id="C8D3A8CB5BD671092999099B25A2A5D3">New species and new records of Cosmolaelaps Berlese (Acari: Laelapidae) from Russia, with a review of the Russian species of the genus</mods:title>
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<mods:identifier id="6344EEDEE1057DF8396E431A5C8D931F" type="GBIF-Dataset">0cd92d63-eeac-48d6-9c9d-270f866de7d3</mods:identifier>
<mods:identifier id="E87D50BE72779251AAD96C00DB38B629" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5C485484E4609B7804F61EFDC144D0CE" type="Zenodo-Dep">6530956</mods:identifier>
<mods:identifier id="A6153671BDD46895DDF8E1B09C6F41A0" type="ZooBank">BD6A434A-21D7-49FC-A09F-FEF4BC740B14</mods:identifier>
</mods:mods>
<treatment id="D25AD024FFB2982D01C7AF4276B9FEF3" ID-DOI="http://doi.org/10.5281/zenodo.6533201" ID-GBIF-Taxon="195277951" ID-Zenodo-Dep="6533201" LSID="urn:lsid:plazi:treatment:D25AD024FFB2982D01C7AF4276B9FEF3" httpUri="http://treatment.plazi.org/id/D25AD024FFB2982D01C7AF4276B9FEF3" lastPageId="19" lastPageNumber="505" pageId="18" pageNumber="504">
<subSubSection id="12E932B9FFB2982C01C7AF427065FCE9" box="[151,832,798,825]" pageId="18" pageNumber="504" type="nomenclature">
<paragraph id="5A4C6132FFB2982C01C7AF427065FCE9" blockId="18.[151,832,798,825]" box="[151,832,798,825]" pageId="18" pageNumber="504">
<heading id="0104D65EFFB2982C01C7AF427065FCE9" bold="true" box="[151,832,798,825]" fontSize="11" level="1" pageId="18" pageNumber="504" reason="1">
<emphasis id="6887BD20FFB2982C01C7AF427065FCE9" bold="true" box="[151,832,798,825]" pageId="18" pageNumber="504">
Key to species of
<taxonomicName id="9DF31AB1FFB2982C003EAF437133FCE8" authorityName="Berlese" authorityYear="1903" box="[366,534,798,825]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="504" phylum="Arthropoda" rank="genus">
<emphasis id="6887BD20FFB2982C003EAF437133FCE8" bold="true" box="[366,534,798,825]" italics="true" pageId="18" pageNumber="504">Cosmolaelaps</emphasis>
</taxonomicName>
occurring in the
<collectingCountry id="22E421A2FFB2982C03BDAF437065FCE9" box="[749,832,798,824]" name="Russia" pageId="18" pageNumber="504">Russia</collectingCountry>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="12E932B9FFB2982D01C7AF0076B9FEF3" lastPageId="19" lastPageNumber="505" pageId="18" pageNumber="504" type="key">
<keyStep id="E1077997FFB2982C01C7AF0076B9FC5E" pageId="18" pageNumber="504">
<paragraph id="5A4C6132FFB2982C01C7AF0076B9FCA3" blockId="18.[151,1437,861,2027]" box="[151,1436,861,883]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01C7AF0076B9FCA3" box="[151,1436,861,883]" pageId="18" pageNumber="504">1. Dorsal setae spatulate with a small basal protuberance........................................................ 2</keyLead>
</paragraph>
<paragraph id="5A4C6132FFB2982C01CDAF2676B9FC5E" blockId="18.[151,1437,861,2027]" box="[157,1436,890,912]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01CDAF2676B9FC5E" box="[157,1436,890,912]" pageId="18" pageNumber="504">– Dorsal setae in other forms (e.g., cuneiform, lancet, knife or scimitar-like)........................................ 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="E1077997FFB2982C01C7AFC576B9FBD7" pageId="18" pageNumber="504">
<paragraph id="5A4C6132FFB2982C01C7AFC576B8FC1B" blockId="18.[151,1437,861,2027]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01C7AFC576B8FC1B" pageId="18" pageNumber="504">
2. Anal shield nearly rounded (semi-circular anterior margin), wider than long, anus small, length of anal shield exceeding length of anal opening by 6 or more times..............
<taxonomicName id="9DF31AB1FFB2982C03B3AFE8777FFC1A" authority="(Berlese, 1883)" baseAuthorityName="Berlese" baseAuthorityYear="1883" box="[739,1114,949,971]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="504" phylum="Arthropoda" rank="species" species="claviger">
<emphasis id="6887BD20FFB2982C03B3AFE870E5FC1B" box="[739,960,949,970]" italics="true" pageId="18" pageNumber="504">Cosmolaelaps claviger</emphasis>
(
<bibRefCitation id="3E621CC3FFB2982C029CAFE87776FC1A" author="Berlese, A." box="[972,1107,949,971]" pageId="18" pageNumber="504" pagination="2" refId="ref13682" refString="Berlese, A. (1883) s. n. Acari, Myriapoda et Scorpiones Hucusque in Italia Reperta, 4, 2. https: // doi. org / 10.5962 / bhl. title. 69269" type="journal article" year="1883">Berlese, 1883</bibRefCitation>
)
</taxonomicName>
[=
<taxonomicName id="9DF31AB1FFB2982C052BAFE876B3FC1B" authority="Pirianyk, 1959" authorityName="Pirianyk" authorityYear="1959" box="[1147,1430,948,971]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="504" phylum="Arthropoda" rank="species" species="bregetovae">
<emphasis id="6887BD20FFB2982C052BAFE877AEFC1B" box="[1147,1163,949,970]" italics="true" pageId="18" pageNumber="504">C</emphasis>
.
<emphasis id="6887BD20FFB2982C05C7AFE87624FC1B" box="[1175,1281,949,970]" italics="true" pageId="18" pageNumber="504">bregetovae</emphasis>
<bibRefCitation id="3E621CC3FFB2982C0457AFE876B3FC1B" author="Pirianyk, G. I." box="[1287,1430,948,971]" pageId="18" pageNumber="504" pagination="97 - 105" refId="ref16171" refString="Pirianyk, G. I. (1959) New species of gamasid mites (Parasitiformes, Gamasoidea). Pratsi Instytut Zoologii Kiev, 15, 97 - 105. [in Ukrainian]" type="journal article" year="1959">Pirianyk, 1959</bibRefCitation>
</taxonomicName>
]
</keyLead>
</paragraph>
<paragraph id="5A4C6132FFB2982C01CDAF8E76B9FBD7" blockId="18.[151,1437,861,2027]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01CDAF8E76B9FBD7" pageId="18" pageNumber="504">
– Anal shield nearly subtriangular, as long as wide, anus normal, length of anal shield exceeding length of anal opening by 3-4 times........................................................................
<taxonomicName id="9DF31AB1FFB2982C052FAFAD76B9FBD7" authority="Sheals, 1962" authorityName="Sheals" authorityYear="1962" box="[1151,1436,1008,1030]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="504" phylum="Arthropoda" rank="species" species="rectangularis">
<emphasis id="6887BD20FFB2982C052FAFAD77AAFBD4" box="[1151,1167,1008,1029]" italics="true" pageId="18" pageNumber="504">C</emphasis>
.
<emphasis id="6887BD20FFB2982C05CBAFAD763EFBD4" box="[1179,1307,1008,1029]" italics="true" pageId="18" pageNumber="504">rectangularis</emphasis>
<bibRefCitation id="3E621CC3FFB2982C0472AFAD76B9FBD7" author="Sheals, J. G." box="[1314,1436,1008,1030]" pageId="18" pageNumber="504" pagination="83 - 110" refId="ref16566" refString="Sheals, J. G. (1962) Mesostigmata: Gamasida (Acari). Biologie de L'Amerique Australe, 1, 83 - 110." type="journal article" year="1962">Sheals, 1962</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E1077997FFB2982C01C7A85076B9FB91" pageId="18" pageNumber="504">
<paragraph id="5A4C6132FFB2982C01C7A85076B9FBF3" blockId="18.[151,1437,861,2027]" box="[151,1436,1037,1059]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01C7A85076B9FBF3" box="[151,1436,1037,1059]" pageId="18" pageNumber="504">3. Dorsal setae cuneiform................................................................................ 4</keyLead>
</paragraph>
<paragraph id="5A4C6132FFB2982C01CDA87676B9FB91" blockId="18.[151,1437,861,2027]" box="[157,1436,1066,1088]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01CDA87676B9FB91" box="[157,1436,1066,1088]" pageId="18" pageNumber="504">– Dorsal setae lancet, knife or scimitar-like.................................................................. 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="E1077997FFB2982C01C7A81576B9FBAA" pageId="18" pageNumber="504">
<paragraph id="5A4C6132FFB2982C01C7A81576B9FB8F" blockId="18.[151,1437,861,2027]" box="[151,1436,1096,1118]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01C7A81576B9FB8F" box="[151,1436,1096,1118]" pageId="18" pageNumber="504">
4. Post-anal seta spatulate .....................................................
<taxonomicName id="9DF31AB1FFB2982C0512A81576B9FB8F" authority="(Evans &amp; Till, 1966)" baseAuthorityName="Evans &amp; Till" baseAuthorityYear="1966" box="[1090,1436,1096,1118]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="504" phylum="Arthropoda" rank="species" species="neocuneifer">
<emphasis id="6887BD20FFB2982C0512A8157777FB8C" box="[1090,1106,1096,1117]" italics="true" pageId="18" pageNumber="504">C</emphasis>
.
<emphasis id="6887BD20FFB2982C050EA81577EAFB8C" box="[1118,1231,1096,1117]" italics="true" pageId="18" pageNumber="504">neocuneifer</emphasis>
(
<bibRefCitation id="3E621CC3FFB2982C058CA81576B0FB8F" author="Evans, G. O. &amp; Till, W. M." box="[1244,1429,1096,1118]" pageId="18" pageNumber="504" pagination="107 - 370" refId="ref14300" refString="Evans, G. O. &amp; Till, W. M. (1966) Studies on the British Dermanyssidae (Acari: Mesostigmata). Part II. Classification. Bulletin of the British Museum (Natural History) Zoology, 14, 107 - 370." type="journal article" year="1966">Evans &amp; Till, 1966</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="5A4C6132FFB2982C01CDA83876B9FBAA" blockId="18.[151,1437,861,2027]" box="[157,1436,1125,1147]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01CDA83876B9FBAA" box="[157,1436,1125,1147]" pageId="18" pageNumber="504">
– Post-anal seta simple............................................................
<taxonomicName id="9DF31AB1FFB2982C05C1A83876B9FBAA" authority="(Michael, 1891)" baseAuthorityName="Michael" baseAuthorityYear="1891" box="[1169,1436,1125,1147]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="504" phylum="Arthropoda" rank="species" species="cuneifer">
<emphasis id="6887BD20FFB2982C05C1A8387784FBAB" box="[1169,1185,1125,1146]" italics="true" pageId="18" pageNumber="504">C</emphasis>
.
<emphasis id="6887BD20FFB2982C05FDA83877D9FBAB" box="[1197,1276,1125,1146]" italics="true" pageId="18" pageNumber="504">cuneifer</emphasis>
(
<bibRefCitation id="3E621CC3FFB2982C0458A83876B0FBAA" author="Michael, A. D." box="[1288,1429,1125,1147]" pageId="18" pageNumber="504" pagination="638 - 654" refId="ref15960" refString="Michael, A. D. (1891) On the association of gamasids with ants. Proceedings of the Zoological Society of London, 4, 638 - 654." type="journal article" year="1891">Michael, 1891</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E1077997FFB2982C01C7A8DF76B9FB64" pageId="18" pageNumber="504">
<paragraph id="5A4C6132FFB2982C01C7A8DF76B9FB49" blockId="18.[151,1437,861,2027]" box="[151,1436,1154,1176]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01C7A8DF76B9FB49" box="[151,1436,1154,1176]" pageId="18" pageNumber="504">5. Dorsal setae slender, lancet-like, with small basal protuberance................................................ 6</keyLead>
</paragraph>
<paragraph id="5A4C6132FFB2982C01CDA8FD76B9FB64" blockId="18.[151,1437,861,2027]" box="[157,1436,1184,1206]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01CDA8FD76B9FB64" box="[157,1436,1184,1206]" pageId="18" pageNumber="504">– Dorsal setae relatively expanded, knife or scimitar-like, with conspicuous basal protuberance......................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="E1077997FFB2982C01C7A8E076B9FADC" pageId="18" pageNumber="504">
<paragraph id="5A4C6132FFB2982C01C7A8E076B9FB03" blockId="18.[151,1437,861,2027]" box="[151,1436,1213,1235]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01C7A8E076B9FB03" box="[151,1436,1213,1235]" pageId="18" pageNumber="504">6. Movable digit with normal two teeth...................................................................... 7</keyLead>
</paragraph>
<paragraph id="5A4C6132FFB2982C01CDA88676B9FADC" blockId="18.[151,1437,861,2027]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01CDA88676B9FADC" pageId="18" pageNumber="504">
– Two teeth of movable digit separated by a row of smaller teeth.......................................................
<taxonomicName id="9DF31AB1FFB2982C007CA8A5717DFADF" authority="(Berlese, 1910)" baseAuthorityName="Berlese" baseAuthorityYear="1910" box="[300,600,1272,1294]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="504" phylum="Arthropoda" rank="species" species="gryllotalpae">
<emphasis id="6887BD20FFB2982C007CA8A57219FADC" box="[300,316,1272,1293]" italics="true" pageId="18" pageNumber="504">C</emphasis>
.
<emphasis id="6887BD20FFB2982C0018A8A57298FADC" box="[328,445,1272,1293]" italics="true" pageId="18" pageNumber="504">gryllotalpae</emphasis>
(
<bibRefCitation id="3E621CC3FFB2982C009AA8A57175FADF" author="Berlese, A." box="[458,592,1272,1294]" pageId="18" pageNumber="504" pagination="242 - 271" refId="ref13762" refString="Berlese, A. (1910) Lista di nuove specie e nuovi generi di Acari. Redia, 6, 242 - 271." type="journal article" year="1910">Berlese, 1910</bibRefCitation>
)
</taxonomicName>
[=
<taxonomicName id="9DF31AB1FFB2982C0323A8A571C8FADC" box="[627,749,1272,1293]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="504" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="6887BD20FFB2982C0323A8A571A1FADC" box="[627,644,1272,1293]" italics="true" pageId="18" pageNumber="504">H</emphasis>
.
<emphasis id="6887BD20FFB2982C03C0A8A571C8FADC" box="[656,749,1272,1293]" italics="true" pageId="18" pageNumber="504">elongatus</emphasis>
</taxonomicName>
,
<taxonomicName id="9DF31AB1FFB2982C03AAA8A570A7FADC" authorityName="Teng" authorityYear="1982" box="[762,898,1272,1293]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="504" phylum="Arthropoda" rank="species" species="acutiscutus">
<emphasis id="6887BD20FFB2982C03AAA8A5702FFADC" box="[762,778,1272,1293]" italics="true" pageId="18" pageNumber="504">C</emphasis>
.
<emphasis id="6887BD20FFB2982C0246A8A570A7FADC" box="[790,898,1272,1293]" italics="true" pageId="18" pageNumber="504">acutiscutus</emphasis>
</taxonomicName>
,
<taxonomicName id="9DF31AB1FFB2982C02DEA8A57712FADC" authorityName="Bai &amp; Wang" authorityYear="2005" box="[910,1079,1272,1293]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="504" phylum="Arthropoda" rank="species" species="subacutiscutus">
<emphasis id="6887BD20FFB2982C02DEA8A570BBFADC" box="[910,926,1272,1293]" italics="true" pageId="18" pageNumber="504">C</emphasis>
.
<emphasis id="6887BD20FFB2982C02FAA8A57712FADC" box="[938,1079,1272,1293]" italics="true" pageId="18" pageNumber="504">subacutiscutus</emphasis>
</taxonomicName>
,
<taxonomicName id="9DF31AB1FFB2982C0513A8A57791FADC" authorityName="Sklyar" authorityYear="2012" box="[1091,1204,1272,1293]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="504" phylum="Arthropoda" rank="species" species="mirificus">
<emphasis id="6887BD20FFB2982C0513A8A57776FADC" box="[1091,1107,1272,1293]" italics="true" pageId="18" pageNumber="504">C</emphasis>
.
<emphasis id="6887BD20FFB2982C050FA8A57791FADC" box="[1119,1204,1272,1293]" italics="true" pageId="18" pageNumber="504">mirificus</emphasis>
</taxonomicName>
and
<taxonomicName id="9DF31AB1FFB2982C05B3A8A576B1FADC" authorityName="Willmann" authorityYear="1951" box="[1251,1428,1272,1293]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="504" phylum="Arthropoda" rank="species" species="angustiscutatus">
<emphasis id="6887BD20FFB2982C05B3A8A577D6FADC" box="[1251,1267,1272,1293]" italics="true" pageId="18" pageNumber="504">C</emphasis>
.
<emphasis id="6887BD20FFB2982C05AFA8A576B1FADC" box="[1279,1428,1272,1293]" italics="true" pageId="18" pageNumber="504">angustiscutatus</emphasis>
</taxonomicName>
]
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E1077997FFB2982C01C7A94876B9FA96" pageId="18" pageNumber="504">
<paragraph id="5A4C6132FFB2982C01C7A94876B9FAFA" blockId="18.[151,1437,861,2027]" box="[151,1436,1301,1323]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01C7A94876B9FAFA" box="[151,1436,1301,1323]" pageId="18" pageNumber="504">
7. Dorsal shield with 37 pairs of setae (
<emphasis id="6887BD20FFB2982C0312A948717DFAFB" box="[578,600,1301,1322]" italics="true" pageId="18" pageNumber="504">z3</emphasis>
and
<emphasis id="6887BD20FFB2982C03D6A94871B9FAFB" box="[646,668,1301,1322]" italics="true" pageId="18" pageNumber="504">s1</emphasis>
absent) .....................................
<taxonomicName id="9DF31AB1FFB2982C05FAA94876B9FAFA" authority="(Michael, 1891)" baseAuthorityName="Michael" baseAuthorityYear="1891" box="[1194,1436,1301,1323]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="504" phylum="Arthropoda" rank="species" species="acuta">
<emphasis id="6887BD20FFB2982C05FAA948779FFAFB" box="[1194,1210,1301,1322]" italics="true" pageId="18" pageNumber="504">C</emphasis>
.
<emphasis id="6887BD20FFB2982C0596A94B77DEFAFA" box="[1222,1275,1302,1323]" italics="true" pageId="18" pageNumber="504">acuta</emphasis>
(
<bibRefCitation id="3E621CC3FFB2982C0458A94876B0FAFA" author="Michael, A. D." box="[1288,1429,1301,1323]" pageId="18" pageNumber="504" pagination="638 - 654" refId="ref15960" refString="Michael, A. D. (1891) On the association of gamasids with ants. Proceedings of the Zoological Society of London, 4, 638 - 654." type="journal article" year="1891">Michael, 1891</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="5A4C6132FFB2982C01CDA96E76B9FA96" blockId="18.[151,1437,861,2027]" box="[157,1436,1330,1352]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01CDA96E76B9FA96" box="[157,1436,1330,1352]" pageId="18" pageNumber="504">– Dorsal shield with 39–40 pairs of setae.................................................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="E1077997FFB2982C01C7A90D76B9FA52" pageId="18" pageNumber="504">
<paragraph id="5A4C6132FFB2982C01C7A90D76B9FAB7" blockId="18.[151,1437,861,2027]" box="[151,1436,1360,1382]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01C7A90D76B9FAB7" box="[151,1436,1360,1382]" pageId="18" pageNumber="504">
8. Dorsal shield with 39 pairs of setae (including two pairs of
<emphasis id="6887BD20FFB2982C024CA90D7011FAB4" box="[796,820,1360,1381]" italics="true" pageId="18" pageNumber="504">Zx</emphasis>
setae) ...................
<taxonomicName id="9DF31AB1FFB2982C0508A90D76B9FAB7" authority="(Shcherbak, 1971 a)" baseAuthorityName="Shcherbak" baseAuthorityYear="1971" box="[1112,1436,1360,1382]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="504" phylum="Arthropoda" rank="species" species="lutegiensis">
<emphasis id="6887BD20FFB2982C0508A90D774DFAB4" box="[1112,1128,1360,1381]" italics="true" pageId="18" pageNumber="504">C</emphasis>
.
<emphasis id="6887BD20FFB2982C0524A90D77FEFAB4" box="[1140,1243,1360,1381]" italics="true" pageId="18" pageNumber="504">lutegiensis</emphasis>
(
<bibRefCitation id="3E621CC3FFB2982C05B7A90D76B0FAB7" author="Shcherbak, G. I." box="[1255,1429,1360,1382]" pageId="18" pageNumber="504" pagination="76 - 79" refId="ref16468" refString="Shcherbak, G. I. (1971 a) New species of gamasid from the genus Hypoaspis Canestrini (Acarina, Gamasoidea). Vestnik Zoologii, 5, 76 - 79. [in Russian]" type="journal article" year="1971">Shcherbak, 1971a</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="5A4C6132FFB2982C01CDA93076B9FA52" blockId="18.[151,1437,861,2027]" box="[157,1436,1389,1411]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01CDA93076B9FA52" box="[157,1436,1389,1411]" pageId="18" pageNumber="504">
– Dorsal shield with 40 pairs of setae (including three pairs of
<emphasis id="6887BD20FFB2982C0278A9337065FA52" box="[808,832,1390,1411]" italics="true" pageId="18" pageNumber="504">Zx</emphasis>
setae) ................. ..
<taxonomicName id="9DF31AB1FFB2982C050DA93076B9FA52" authority="(Pirianyk, 1959)" baseAuthorityName="Pirianyk" baseAuthorityYear="1959" box="[1117,1436,1389,1411]" class="Arachnida" family="Laelapidae" genus="Laelaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="504" phylum="Arthropoda" rank="species" species="markewitschi">
<emphasis id="6887BD20FFB2982C050DA9307748FA53" box="[1117,1133,1389,1410]" italics="true" pageId="18" pageNumber="504">C</emphasis>
.
<emphasis id="6887BD20FFB2982C0529A93077DCFA53" box="[1145,1273,1389,1410]" italics="true" pageId="18" pageNumber="504">markewitschi</emphasis>
(
<bibRefCitation id="3E621CC3FFB2982C0455A93076B0FA52" author="Pirianyk, G. I." box="[1285,1429,1389,1411]" pageId="18" pageNumber="504" pagination="97 - 105" refId="ref16171" refString="Pirianyk, G. I. (1959) New species of gamasid mites (Parasitiformes, Gamasoidea). Pratsi Instytut Zoologii Kiev, 15, 97 - 105. [in Ukrainian]" type="journal article" year="1959">Pirianyk, 1959</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E1077997FFB2982C01C7A9D776B9FA6C" pageId="18" pageNumber="504">
<paragraph id="5A4C6132FFB2982C01C7A9D776B9FA4E" blockId="18.[151,1437,861,2027]" box="[151,1436,1418,1440]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01C7A9D776B9FA4E" box="[151,1436,1418,1440]" pageId="18" pageNumber="504">9. Dorsal shield with very long setae, almost all setae reaching well past base of following setae (scimitar-like)............ 10</keyLead>
</paragraph>
<paragraph id="5A4C6132FFB2982C01CDA9F576B9FA6C" blockId="18.[151,1437,861,2027]" box="[157,1436,1448,1470]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01CDA9F576B9FA6C" box="[157,1436,1448,1470]" pageId="18" pageNumber="504">– Dorsal shield setae shorter, at most reaching base of following setae (knife-like) ................................... 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="E1077997FFB2982C01C7A99876B9FA26" pageId="18" pageNumber="504">
<paragraph id="5A4C6132FFB2982C01C7A99876B9FA0A" blockId="18.[151,1437,861,2027]" box="[151,1436,1477,1499]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01C7A99876B9FA0A" box="[151,1436,1477,1499]" pageId="18" pageNumber="504">
10. Dorsal shield setae
<emphasis id="6887BD20FFB2982C00CDA9987293FA0B" box="[413,438,1477,1498]" italics="true" pageId="18" pageNumber="504">Z5</emphasis>
longer than
<emphasis id="6887BD20FFB2982C037FA9987163FA0B" box="[559,582,1477,1498]" italics="true" pageId="18" pageNumber="504">J5</emphasis>
...............................................................
<taxonomicName id="9DF31AB1FFB2982C046BA99876B9FA0A" box="[1339,1436,1477,1499]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="504" phylum="Arthropoda" rank="species" species="sejongi">
<emphasis id="6887BD20FFB2982C046BA998766EFA0B" box="[1339,1355,1477,1498]" italics="true" pageId="18" pageNumber="504">C</emphasis>
.
<emphasis id="6887BD20FFB2982C0407A99B76B9FA0A" box="[1367,1436,1478,1499]" italics="true" pageId="18" pageNumber="504">sejongi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="5A4C6132FFB2982C01CDA9BE76B9FA26" blockId="18.[151,1437,861,2027]" box="[157,1436,1506,1528]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01CDA9BE76B9FA26" box="[157,1436,1506,1528]" pageId="18" pageNumber="504">
– Dorsal shield setae
<emphasis id="6887BD20FFB2982C00CDA9BE7293FA29" box="[413,438,1507,1528]" italics="true" pageId="18" pageNumber="504">Z5</emphasis>
shorter than
<emphasis id="6887BD20FFB2982C0364A9BE716EFA29" box="[564,587,1507,1528]" italics="true" pageId="18" pageNumber="504">J5</emphasis>
(ratio of
<emphasis id="6887BD20FFB2982C03F5A9BE7199FA29" box="[677,700,1507,1528]" italics="true" pageId="18" pageNumber="504">J5</emphasis>
/
<emphasis id="6887BD20FFB2982C0392A9BE71F9FA29" box="[706,732,1507,1528]" italics="true" pageId="18" pageNumber="504">Z5</emphasis>
≈ 1.5)................................................... 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E1077997FFB2982C01C7AA5D76B9F9BC" pageId="18" pageNumber="504">
<paragraph id="5A4C6132FFB2982C01C7AA5D76B9F9E3" blockId="18.[151,1437,861,2027]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01C7AA5D76B9F9E3" pageId="18" pageNumber="504">
11. Post-anal seta well thickened, sparsely barbed and apically hook shaped, sternal setae long, extending beyond base of following setae.........................................................................
<taxonomicName id="9DF31AB1FFB2982C05DAAA40766DF9E2" authority="Joharchi" authorityName="Joharchi" authorityYear="2022" box="[1162,1352,1565,1587]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="504" phylum="Arthropoda" rank="species" species="latisetis" status="sp. nov.">
<emphasis id="6887BD20FFB2982C05DAAA4077BFF9E3" box="[1162,1178,1565,1586]" italics="true" pageId="18" pageNumber="504">C</emphasis>
.
<emphasis id="6887BD20FFB2982C05F6AA4077D4F9E3" box="[1190,1265,1565,1586]" italics="true" pageId="18" pageNumber="504">latisetis</emphasis>
Joharchi
</taxonomicName>
<taxonomicNameLabel id="73B4005BFFB2982C041EAA4076B9F9E3" box="[1358,1436,1565,1586]" pageId="18" pageNumber="504" rank="species">
<emphasis id="6887BD20FFB2982C041EAA4076B3F9E3" bold="true" box="[1358,1430,1565,1586]" pageId="18" pageNumber="504">sp. nov</emphasis>
.
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="5A4C6132FFB2982C01CDAA6676B9F9BC" blockId="18.[151,1437,861,2027]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01CDAA6676B9F9BC" pageId="18" pageNumber="504">
– Post-anal seta slightly thickened, simple, sternal setae short, at most reaching base of following setae ................................................................................................
<taxonomicName id="9DF31AB1FFB2982C052DAA0577DBF9BC" box="[1149,1278,1624,1645]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="504" phylum="Arthropoda" rank="species" species="chianensis">
<emphasis id="6887BD20FFB2982C052DAA0577A8F9BC" box="[1149,1165,1624,1645]" italics="true" pageId="18" pageNumber="504">C</emphasis>
.
<emphasis id="6887BD20FFB2982C05C9AA0577DBF9BC" box="[1177,1278,1624,1645]" italics="true" pageId="18" pageNumber="504">chianensis</emphasis>
</taxonomicName>
[=
<taxonomicName id="9DF31AB1FFB2982C044AAA0576B0F9BC" box="[1306,1429,1624,1645]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="504" phylum="Arthropoda" rank="species" species="hefeiensis">
<emphasis id="6887BD20FFB2982C044AAA05760FF9BC" box="[1306,1322,1624,1645]" italics="true" pageId="18" pageNumber="504">C</emphasis>
.
<emphasis id="6887BD20FFB2982C0466AA0576B0F9BC" box="[1334,1429,1624,1645]" italics="true" pageId="18" pageNumber="504">hefeiensis</emphasis>
</taxonomicName>
]
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E1077997FFB2982C01C7AA2876B9F979" pageId="18" pageNumber="504">
<paragraph id="5A4C6132FFB2982C01C7AA2876B9F95B" blockId="18.[151,1437,861,2027]" box="[151,1436,1653,1675]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01C7AA2876B9F95B" box="[151,1436,1653,1675]" pageId="18" pageNumber="504">12. Femur IV with spatulate setae ........................................................................... 13</keyLead>
</paragraph>
<paragraph id="5A4C6132FFB2982C01CDAACE76B9F979" blockId="18.[151,1437,861,2027]" box="[157,1436,1682,1704]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01CDAACE76B9F979" box="[157,1436,1682,1704]" pageId="18" pageNumber="504">– Femur IV without spatulate setae........................................................................ 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="E1077997FFB2982C01C7AAED76B9F933" pageId="18" pageNumber="504">
<paragraph id="5A4C6132FFB2982C01C7AAED76B9F914" blockId="18.[151,1437,861,2027]" box="[151,1436,1711,1734]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01C7AAED76B9F914" box="[151,1436,1711,1734]" pageId="18" pageNumber="504">
13. Presternal area with two triangular plates and obviously separated from sternal shield..........
<taxonomicName id="9DF31AB1FFB2982C05C4AAED76B9F914" authority="(Berlese, 1905)" baseAuthorityName="Berlese" baseAuthorityYear="1905" box="[1172,1436,1711,1734]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="504" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="6887BD20FFB2982C05C4AAED7781F914" box="[1172,1188,1712,1733]" italics="true" pageId="18" pageNumber="504">C</emphasis>
.
<emphasis id="6887BD20FFB2982C05E0AAED7627F914" box="[1200,1282,1712,1733]" italics="true" pageId="18" pageNumber="504">robustus</emphasis>
(
<bibRefCitation id="3E621CC3FFB2982C045FAAED76B0F914" author="Berlese, A." box="[1295,1429,1711,1734]" pageId="18" pageNumber="504" pagination="154 - 176" refId="ref13741" refString="Berlese, A. (1905) Acari nuovi. Manipulus IV. Redia, 2, 154 - 176." type="journal article" year="1905">Berlese, 1905</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="5A4C6132FFB2982C01CDAA9076B9F933" blockId="18.[151,1437,861,2027]" box="[157,1436,1741,1763]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01CDAA9076B9F933" box="[157,1436,1741,1763]" pageId="18" pageNumber="504">– Presternal area lightly sclerotised, with a few transverse curved lines and merged to sternal shield, without distinct plates.. 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="E1077997FFB2982C01C7AAB776B9F886" pageId="18" pageNumber="504">
<paragraph id="5A4C6132FFB2982C01C7AAB776B9F8CC" blockId="18.[151,1437,861,2027]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01C7AAB776B9F8CC" pageId="18" pageNumber="504">
14. Dorsal setae relatively short and thin, mostly not reach to base of next setae in series, femora II and IV with spatulate setae.....................................
<taxonomicName id="9DF31AB1FFB2982C03DDAB5570A6F8CF" authority="(Michael, 1891)" authorityName="sensu Evans &amp; Till" authorityYear="1966" baseAuthorityName="Michael" baseAuthorityYear="1891" box="[653,899,1800,1822]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="504" phylum="Arthropoda" rank="species" species="vacua">
<emphasis id="6887BD20FFB2982C03DDAB5571B8F8CC" box="[653,669,1800,1821]" italics="true" pageId="18" pageNumber="504">C</emphasis>
.
<emphasis id="6887BD20FFB2982C03F9AB5571C7F8CC" box="[681,738,1800,1821]" italics="true" pageId="18" pageNumber="504">vacua</emphasis>
(
<bibRefCitation id="3E621CC3FFB2982C03BFAB557059F8CF" author="Michael, A. D." box="[751,892,1800,1822]" pageId="18" pageNumber="504" pagination="638 - 654" refId="ref15960" refString="Michael, A. D. (1891) On the association of gamasids with ants. Proceedings of the Zoological Society of London, 4, 638 - 654." type="journal article" year="1891">Michael, 1891</bibRefCitation>
)
</taxonomicName>
[=
<taxonomicName id="9DF31AB1FFB2982C02CEAB5577BCF8CF" authority="Oudemans" authorityName="Oudemans" authorityYear="1902" box="[926,1177,1800,1822]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="504" phylum="Arthropoda" rank="species" species="militiformis">
<emphasis id="6887BD20FFB2982C02CEAB55708AF8CC" box="[926,943,1800,1821]" italics="true" pageId="18" pageNumber="504">H</emphasis>
.
<emphasis id="6887BD20FFB2982C02ECAB557709F8CC" box="[956,1068,1800,1821]" italics="true" pageId="18" pageNumber="504">militiformis</emphasis>
Oudemans
</taxonomicName>
and
<emphasis id="6887BD20FFB2982C0597AB5577FDF8CC" box="[1223,1240,1800,1821]" italics="true" pageId="18" pageNumber="504">H</emphasis>
. (
<emphasis id="6887BD20FFB2982C05BCAB5577D9F8CC" box="[1260,1276,1800,1821]" italics="true" pageId="18" pageNumber="504">C</emphasis>
.)
<emphasis id="6887BD20FFB2982C0440AB5576B1F8CC" box="[1296,1428,1800,1821]" italics="true" pageId="18" pageNumber="504">serratosimilis</emphasis>
]
</keyLead>
</paragraph>
<paragraph id="5A4C6132FFB2982C01CDAB7876B9F886" blockId="18.[151,1437,861,2027]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01CDAB7876B9F886" pageId="18" pageNumber="504">
– Dorsal setae much thicker, mostly long enough to reach base of next setae in series, femora II–IV with spatulate setae...............................................................................
<taxonomicName id="9DF31AB1FFB2982C050EAB1E76B9F886" authority="Davydova, 1977" authorityName="Davydova" authorityYear="1977" box="[1118,1436,1858,1880]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="504" phylum="Arthropoda" rank="species" species="dendrophilus">
<emphasis id="6887BD20FFB2982C050EAB1E774BF889" box="[1118,1134,1859,1880]" italics="true" pageId="18" pageNumber="504">C</emphasis>
.
<emphasis id="6887BD20FFB2982C052AAB1F77DDF886" box="[1146,1272,1858,1879]" italics="true" pageId="18" pageNumber="504">dendrophilus</emphasis>
<bibRefCitation id="3E621CC3FFB2982C05AEAB1F76B9F886" author="Davydova, M. S." box="[1278,1436,1858,1880]" pageId="18" pageNumber="504" pagination="23 - 29" refId="ref13934" refString="Davydova, M. S. (1977) New and a little-known species of free-living gamasid mites (Parasitiformes, Gamasoidea). In: Cherepanov, A. I. (Ed.), Taxa of the Siberian Fauna, Nauka, Novosibirsk, pp. 23 - 29. [in Russian]" type="book chapter" year="1977">Davydova, 1977</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E1077997FFB2982D01C7AB0276B9FF7C" lastPageId="19" lastPageNumber="505" pageId="18" pageNumber="504">
<paragraph id="5A4C6132FFB2982C01C7AB0276B9F861" blockId="18.[151,1437,861,2027]" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982C01C7AB0276B9F861" pageId="18" pageNumber="504">15. Dorsal setae long enough to reach base of following setae, opisthogastric setae arising on small sclerotised platelets and moderately thick, distance between genital and anal shields is shorter than length of anal shield, anal shield longer than wide .................................................................................................. 16</keyLead>
</paragraph>
<paragraph id="5A4C6132FFB2982D01CDABE576B9FF7C" blockId="18.[151,1437,861,2027]" lastBlockId="19.[151,1437,151,291]" lastPageId="19" lastPageNumber="505" pageId="18" pageNumber="504">
<keyLead id="E102C207FFB2982D01CDABE576B9FF7C" lastPageId="19" lastPageNumber="505" pageId="18" pageNumber="504">
– Dorsal setae barely reaching more than half of distance of following setae in series, opisthogastric setae thin and without any such small sclerotised platelets, distance between genital and anal shields is about 1.5 times the length of anal shield, anal shield wider than long ...................................................... ..
<taxonomicName id="9DF31AB1FFB3982D0546ACCA76B9FF7C" authority="(Shcherbak, 1971 b)" baseAuthorityName="Shcherbak" baseAuthorityYear="1971" box="[1046,1436,151,173]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="19" pageNumber="505" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="6887BD20FFB3982D0546ACCA77FCFF7C" box="[1046,1241,151,173]" italics="true" pageId="19" pageNumber="505">Cosmolaelaps mixta</emphasis>
(
<bibRefCitation id="3E621CC3FFB3982D05B5ACCA76B3FF7C" author="Shcherbak, G. I." box="[1253,1430,151,173]" pageId="19" pageNumber="505" pagination="20 - 30" refId="ref16506" refString="Shcherbak, G. I. (1971 b) Species of Gamasoidea mites from the genus Hypoaspis Can. (Parasitiformes, Gamasoidea) new for the USSR fauna. Collection of papers from the Zoological Museum of the Institute of Zoology of SSR Ukraine Academy of Sciences, 34, 20 - 30. [in Ukrainian]" type="journal article" year="1971">Shcherbak, 1971b</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E1077997FFB3982D01C7ACE876B9FEF3" pageId="19" pageNumber="505">
<paragraph id="5A4C6132FFB3982D01C7ACE876B9FF36" blockId="19.[151,1437,151,291]" pageId="19" pageNumber="505">
<keyLead id="E102C207FFB3982D01C7ACE876B9FF36" pageId="19" pageNumber="505">
16. Genital shield tapering posteriorly, anterior margin of sternal shield with a median notch, fixed digit of chelicera with three teeth..............................................................................
<taxonomicName id="9DF31AB1FFB3982D05D6AC8E76B9FF36" authority="Joharchi, 2019" authorityName="Joharchi" authorityYear="2019" box="[1158,1436,210,232]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="19" pageNumber="505" phylum="Arthropoda" rank="species" species="siberiensis">
<emphasis id="6887BD20FFB3982D05D6AC8E77B3FF39" box="[1158,1174,211,232]" italics="true" pageId="19" pageNumber="505">C</emphasis>
.
<emphasis id="6887BD20FFB3982D05F2AC8F762CFF36" box="[1186,1289,210,231]" italics="true" pageId="19" pageNumber="505">siberiensis</emphasis>
<bibRefCitation id="3E621CC3FFB3982D045FAC8F76B9FF36" author="Joharchi, O &amp; Trach, V. A." box="[1295,1436,210,232]" pageId="19" pageNumber="505" pagination="486 - 494" refId="ref14677" refString="Joharchi, O &amp; Trach, V. A. (2019) A new species of Cosmolaelaps Berlese (Acari: Laelapidae) from Ukraine. Zootaxa, 4647 (1), 486 - 494. https: // doi. org / 10.11646 / zootaxa. 4647.1.30" type="journal article" year="2019">Joharchi, 2019</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="5A4C6132FFB3982D01CDACAD76B9FEF3" blockId="19.[151,1437,151,291]" pageId="19" pageNumber="505">
<keyLead id="E102C207FFB3982D01CDACAD76B9FEF3" pageId="19" pageNumber="505">
– Genital shield rounded posteriorly, anterior margin of sternal shield hardly conspicuous, fixed digit of chelicera with six various size of teeth............................................................... ..
<taxonomicName id="9DF31AB1FFB3982D053CAD50766DFEF2" authority="Joharchi" authorityName="Joharchi" authorityYear="2022" box="[1132,1352,269,291]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="19" pageNumber="505" phylum="Arthropoda" rank="species" species="lasiophilus" status="sp. nov.">
<emphasis id="6887BD20FFB3982D053CAD507759FEF3" box="[1132,1148,269,290]" italics="true" pageId="19" pageNumber="505">C</emphasis>
.
<emphasis id="6887BD20FFB3982D05D8AD5077D4FEF3" box="[1160,1265,269,290]" italics="true" pageId="19" pageNumber="505">lasiophilus</emphasis>
Joharchi
</taxonomicName>
<taxonomicNameLabel id="73B4005BFFB3982D041EAD5076B9FEF3" box="[1358,1436,269,290]" pageId="19" pageNumber="505" rank="species">
<emphasis id="6887BD20FFB3982D041EAD5076B3FEF3" bold="true" box="[1358,1430,269,290]" pageId="19" pageNumber="505">sp. nov</emphasis>
.
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>