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<mods:title id="0FDDEAFB08B149F5C8CE47DB21681122">New data on the wolf spiders from theAcantholycosa-complex (Araneae: Lycosidae) from the South Siberia</mods:title>
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<mods:affiliation id="B49CD1871960D26FC8A9E174DA39209F">Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia. &amp; Tomsk State University, Lenina Pr., 36, Tomsk, RF- 634050, Russia.</mods:affiliation>
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<taxonomicName id="9D892EA59B13A40ABFED8EE3FE2CFC15" ID-CoL="64GBZ" authorityName="Marusik, Hippa &amp; Koponen" authorityYear="1996" box="[151,435,926,953]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="568" phylum="Arthropoda" rank="species" species="azyuzini">
<emphasis id="68FD89349B13A40ABFED8EE3FE2CFC15" bold="true" box="[151,435,926,953]" italics="true" pageId="1" pageNumber="568">Acantholycosa azyuzini</emphasis>
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group
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<taxonomicName id="9D892EA59B13A40ABFED8E9EFE19FC57" ID-CoL="64GBZ" authorityName="Marusik, Hippa &amp; Koponen" authorityYear="1996" box="[151,390,995,1019]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="568" phylum="Arthropoda" rank="species" species="azyuzini">
<emphasis id="68FD89349B13A40ABFED8E9EFE19FC57" box="[151,390,995,1019]" italics="true" pageId="1" pageNumber="568">Acantholycosa azyuzini</emphasis>
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group—
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<bibRefCitation id="3E1828D79B13A40ABE9D8E9EFD7BFC57" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." box="[487,740,995,1019]" pageId="1" pageNumber="568" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">
Marusik
<emphasis id="68FD89349B13A40ABD388E99FDF1FC57" box="[578,622,995,1019]" italics="true" pageId="1" pageNumber="568">et al</emphasis>
. 2004: 105
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.
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<emphasis id="68FD89349B13A40ABFED8954FE8DFBEF" bold="true" box="[151,274,1065,1091]" pageId="1" pageNumber="568">Diagnosis.</emphasis>
The males of the species belonging to this group can be recognized by their broad and almost straight embolus in combination with claw-like or triangular paleal outgrowth and spine-like terminal apophysis. The females of the species belonging to the
<taxonomicName id="9D892EA59B13A40ABD2C890FFD2DFB27" authorityName="Marusik, Hippa &amp; Koponen" authorityYear="1996" box="[598,690,1138,1163]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="568" phylum="Arthropoda" rank="species" species="azyuzini">
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-group can be diagnosed by a diamond-shaped epigynal fovea and a pentagonal septum.
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<emphasis id="68FD89349B13A40ABFBD89C4FEF8FB7F" bold="true" box="[199,359,1209,1235]" pageId="1" pageNumber="568">Composition.</emphasis>
This group includes eight species:
<taxonomicName id="9D892EA59B13A40ABC7289C7FACFFB7F" authority="Marusik, Azarkina &amp; Koponen, 2004" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[776,1360,1209,1236]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="568" phylum="Arthropoda" rank="species" species="altaiensis">
<emphasis id="68FD89349B13A40ABC7289C7FC0AFB7F" box="[776,917,1209,1236]" italics="true" pageId="1" pageNumber="568">A. altaiensis</emphasis>
<bibRefCitation id="3E1828D79B13A40ABCDA89C4FACFFB7F" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." box="[928,1360,1209,1236]" pageId="1" pageNumber="568" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">Marusik, Azarkina &amp; Koponen, 2004</bibRefCitation>
</taxonomicName>
(
<collectingRegion id="984D9BC49B13A40ABA1989C4FA02FB7F" box="[1379,1437,1209,1235]" country="Russia" name="Altay" pageId="1" pageNumber="568">Altai</collectingRegion>
Mountains),
<taxonomicName id="9D892EA59B13A40ABE5389A3FCA1FB5B" authority="Marusik, Hippa &amp; Koponen, 1996" authorityName="Marusik, Hippa &amp; Koponen" authorityYear="1996" box="[297,830,1245,1272]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="568" phylum="Arthropoda" rank="species" species="azyuzini">
<emphasis id="68FD89349B13A40ABE5389A3FE37FB5B" box="[297,424,1246,1272]" italics="true" pageId="1" pageNumber="568">A. azyuzini</emphasis>
<bibRefCitation id="3E1828D79B13A40ABEC889A0FCA1FB5B" author="Marusik, Y. M. &amp; Hippa, H. &amp; Koponen, S." box="[434,830,1245,1272]" pageId="1" pageNumber="568" pagination="11 - 45" refId="ref10406" refString="Marusik, Y. M., Hippa, H. &amp; Koponen, S. (1996) Spiders (Araneae) from the Altai area, Southern Siberia. Acta Zoologica Fennica, 201, 11 - 45." type="journal article" year="1996">Marusik, Hippa &amp; Koponen, 1996</bibRefCitation>
</taxonomicName>
(
<collectingRegion id="984D9BC49B13A40ABC2A89A0FC15FB5B" box="[848,906,1245,1271]" country="Russia" name="Altay" pageId="1" pageNumber="568">Altai</collectingRegion>
Mountains),
<taxonomicName id="9D892EA59B13A40ABB5C89A3FEFDFAB7" authority="Marusik, Azarkina &amp; Koponen, 2004" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="568" phylum="Arthropoda" rank="species" species="katunensis">
<emphasis id="68FD89349B13A40ABB5C89A3FB21FB5B" box="[1062,1214,1245,1272]" italics="true" pageId="1" pageNumber="568">A. katunensis</emphasis>
<bibRefCitation id="3E1828D79B13A40ABBB389A0FEFDFAB7" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." pageId="1" pageNumber="568" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">Marusik, Azarkina &amp; Koponen, 2004</bibRefCitation>
</taxonomicName>
(
<collectingRegion id="984D9BC49B13A40ABE08887CFE33FAB7" box="[370,428,1281,1307]" country="Russia" name="Altay" pageId="1" pageNumber="568">Altai</collectingRegion>
Mountains),
<taxonomicName id="9D892EA59B13A40ABD39887FFBBFFAB7" authority="Fomichev &amp; Marusik, 2018" authorityName="Fomichev &amp; Marusik" authorityYear="2018" box="[579,1056,1281,1308]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="568" phylum="Arthropoda" rank="species" species="kronestedti">
<emphasis id="68FD89349B13A40ABD39887FFD41FAB7" box="[579,734,1281,1308]" italics="true" pageId="1" pageNumber="568">A. kronestedti</emphasis>
<bibRefCitation id="3E1828D79B13A40ABD9D887CFBBFFAB7" author="Fomichev, A. A. &amp; Marusik, Y. M." box="[743,1056,1281,1307]" pageId="1" pageNumber="568" pagination="271 - 284" refId="ref9976" refString="Fomichev, A. A. &amp; Marusik, Y. M. (2018) Five new species of the Acantholycosa - complex (Araneae: Lycosidae) from Mongolia. Zootaxa, 4497 (2), 271 - 284. https: // doi. org / 10.11646 / zootaxa. 4497.2.7" type="journal article" year="2018">Fomichev &amp; Marusik, 2018</bibRefCitation>
</taxonomicName>
(Mongolian
<collectingRegion id="984D9BC49B13A40ABBC9887CFB70FAB0" box="[1203,1263,1281,1308]" country="Russia" name="Altay" pageId="1" pageNumber="568">Altai</collectingRegion>
),
<taxonomicName id="9D892EA59B13A40ABA79887FFDB6FA93" authority="Marusik, Azarkina &amp; Koponen, 2004" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="568" phylum="Arthropoda" rank="species" species="sayanensis">
<emphasis id="68FD89349B13A40ABA79887FFA03FAB7" box="[1283,1436,1282,1308]" italics="true" pageId="1" pageNumber="568">A. sayanensis</emphasis>
<bibRefCitation id="3E1828D79B13A40ABFED8858FDB6FA93" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." box="[151,553,1317,1344]" pageId="1" pageNumber="568" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">Marusik,Azarkina&amp;Koponen,2004</bibRefCitation>
</taxonomicName>
(Western Sayan Mountains),
<taxonomicName id="9D892EA59B13A40ABC12885BFC41FA93" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[872,990,1317,1344]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="568" phylum="Arthropoda" rank="species" species="sidorovi" status="sp. nov.">
<emphasis id="68FD89349B13A40ABC12885BFC41FA93" box="[872,990,1317,1344]" italics="true" pageId="1" pageNumber="568">A.sidorovi</emphasis>
</taxonomicName>
<emphasis id="68FD89349B13A40ABC9B8858FB82FA93" bold="true" box="[993,1053,1317,1343]" pageId="1" pageNumber="568">
<taxonomicNameLabel id="73CE344F9B13A40ABC9B8858FB82FA93" box="[993,1053,1317,1343]" pageId="1" pageNumber="568" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<collectingRegion id="984D9BC49B13A40ABB528858FBFDFA93" box="[1064,1122,1317,1343]" country="Russia" name="Altay" pageId="1" pageNumber="568">Altai</collectingRegion>
Mountains),
<taxonomicName id="9D892EA59B13A40ABB8A885BFE5EFACF" authority="Marusik &amp; Logunov, 2011" authorityName="Marusik &amp; Logunov" authorityYear="2011" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="568" phylum="Arthropoda" rank="species" species="tarbagataica">
<emphasis id="68FD89349B13A40ABB8A885BFA02FA93" box="[1264,1437,1317,1344]" italics="true" pageId="1" pageNumber="568">A.tarbagataica</emphasis>
Marusik &amp; Logunov, 2011
</taxonomicName>
(Tarbagatai Mt. Range),
<taxonomicName id="9D892EA59B13A40ABDA18837FB05FACF" authority="Fomichev &amp; Marusik, 2018" authorityName="Fomichev &amp; Marusik" authorityYear="2018" box="[731,1178,1353,1380]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="568" phylum="Arthropoda" rank="species" species="vahterae">
<emphasis id="68FD89349B13A40ABDA18837FCC3FACF" box="[731,860,1353,1380]" italics="true" pageId="1" pageNumber="568">A. vahterae</emphasis>
<bibRefCitation id="3E1828D79B13A40ABC198834FB05FACF" author="Fomichev, A. A. &amp; Marusik, Y. M." box="[867,1178,1353,1379]" pageId="1" pageNumber="568" pagination="271 - 284" refId="ref9976" refString="Fomichev, A. A. &amp; Marusik, Y. M. (2018) Five new species of the Acantholycosa - complex (Araneae: Lycosidae) from Mongolia. Zootaxa, 4497 (2), 271 - 284. https: // doi. org / 10.11646 / zootaxa. 4497.2.7" type="journal article" year="2018">Fomichev &amp; Marusik, 2018</bibRefCitation>
</taxonomicName>
(Mongolian
<collectingRegion id="984D9BC49B13A40ABA518834FAF7FAC8" box="[1323,1384,1353,1380]" country="Russia" name="Altay" pageId="1" pageNumber="568">Altai</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="5D54EAB694E233F67F576ACC6EA52028">New data on the wolf spiders from theAcantholycosa-complex (Araneae: Lycosidae) from the South Siberia</mods:title>
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,
<collectorName id="F77C30F09B13A40ABBAE8BE1FA07F91A" box="[1236,1432,1692,1718]" pageId="1" pageNumber="568">S.E. Chernyshov</collectorName>
</materialsCitation>
;
<materialsCitation id="EAE15F7B9B13A40ABFED8BBDFE58F977" box="[151,455,1728,1755]" collectionCode="ISEA" country="Russia" pageId="1" pageNumber="568" specimenCount="14" specimenCount-female="3" specimenCount-male="11" stateProvince="Krasnoyarsk" typeStatus="holotype">
<specimenCount id="4C8F9EAF9B13A40ABFED8BBDFF58F976" box="[151,199,1728,1754]" pageId="1" pageNumber="568" type="male">11♂</specimenCount>
<specimenCount id="4C8F9EAF9B13A40ABFAA8BBDFF6DF976" box="[208,242,1728,1754]" pageId="1" pageNumber="568" type="female">3♀</specimenCount>
(
<collectionCode id="3C98CDE39B13A40ABE798BBDFEDAF976" box="[259,325,1728,1754]" pageId="1" pageNumber="568">ISEA</collectionCode>
, 001.8651)
</materialsCitation>
,
<materialsCitation id="EAE15F7B9B13A40ABEAE8BBDFBE9F952" collectingDate="2020-07-16" collectionCode="ISEA" collectorName="A. A. Fomichev" country="Russia" county="Borus Mt. Range" elevation="1900" location="Bol'shoi Borus" municipality="Malyi Borus" pageId="1" pageNumber="568" specimenCount="1" specimenCount-female="1" stateProvince="Krasnoyarsk" typeStatus="holotype">
<specimenCount id="4C8F9EAF9B13A40ABEAE8BBDFE68F976" box="[468,503,1728,1754]" pageId="1" pageNumber="568" type="female">1♀</specimenCount>
(
<collectionCode id="3C98CDE39B13A40ABD7D8BBDFDD6F976" box="[519,585,1728,1754]" pageId="1" pageNumber="568">ISEA</collectionCode>
, 001.8721),
<collectingCounty id="B3572DAA9B13A40ABDA28BBCFC02F977" box="[728,925,1729,1755]" pageId="1" pageNumber="568">Borus Mt. Range</collectingCounty>
,
<collectingMunicipality id="BA52CF5C9B13A40ABCD08BBDFBA7F977" box="[938,1080,1728,1755]" pageId="1" pageNumber="568">Malyi Borus</collectingMunicipality>
and
<location id="5F5603FD9B13A40ABB088BBDFA83F977" LSID="urn:lsid:plazi:treatment:D220E4309B13A40EBFED88C8FE1CFBA7:5F5603FD9B13A40ABB088BBDFA83F977" box="[1138,1308,1728,1755]" country="Russia" county="Borus Mt. Range" municipality="Malyi Borus" name="Bol'shoi Borus" pageId="1" pageNumber="568" stateProvince="Krasnoyarsk">Bol’shoi Borus</location>
(=
<location id="5F5603FD9B13A40ABA418BBDFA0BF977" LSID="urn:lsid:plazi:treatment:D220E4309B13A40EBFED88C8FE1CFBA7:5F5603FD9B13A40ABA418BBDFA0BF977" box="[1339,1428,1728,1755]" country="Russia" county="Borus Mt. Range" municipality="Malyi Borus" name="Poilova" pageId="1" pageNumber="568" stateProvince="Krasnoyarsk">Poilova</location>
)
<location id="5F5603FD9B13A40ABFED8B99FE8DF952" LSID="urn:lsid:plazi:treatment:D220E4309B13A40EBFED88C8FE1CFBA7:5F5603FD9B13A40ABFED8B99FE8DF952" box="[151,274,1764,1790]" country="Russia" county="Borus Mt. Range" municipality="Malyi Borus" name="Mountains" pageId="1" pageNumber="568" stateProvince="Krasnoyarsk">Mountains</location>
, 52°47–48’N, 91°31–33’E, scree,
<quantity id="9D71F8C39B13A40ABDE88B99FCB0F953" box="[658,815,1764,1791]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.3" metricValueMin="1.5" pageId="1" pageNumber="568" unit="m" value="1900.0" valueMax="2300.0" valueMin="1500.0">
<elevation id="D1A4B2159B13A40ABDE88B99FCB0F953" box="[658,815,1764,1791]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.3" metricValueMin="1.5" pageId="1" pageNumber="568" unit="m" value="1900.0" valueMax="2300.0" valueMin="1500.0">1500–2300 m</elevation>
</quantity>
,
<date id="2E3773E69B13A40ABC408B99FC24F952" box="[826,955,1764,1790]" pageId="1" pageNumber="568" value="2020-07-16">
<collectingDate id="3E738A0E9B13A40ABC408B99FC24F952" box="[826,955,1764,1790]" pageId="1" pageNumber="568" value="2020-07-16">16.07.2020</collectingDate>
</date>
,
<collectorName id="F77C30F09B13A40ABCBF8B99FBE9F952" box="[965,1142,1764,1790]" pageId="1" pageNumber="568">A.A. Fomichev</collectorName>
</materialsCitation>
;
<materialsCitation id="EAE15F7B9B13A40ABBFA8B99FAD0F952" box="[1152,1359,1764,1791]" collectionCode="ZMMU" country="Russia" pageId="1" pageNumber="568" specimenCount="17" specimenCount-female="12" specimenCount-male="5" stateProvince="Krasnoyarsk" typeStatus="holotype">
<specimenCount id="4C8F9EAF9B13A40ABBFA8B99FB3CF953" box="[1152,1187,1764,1791]" pageId="1" pageNumber="568" type="male">5♂</specimenCount>
<specimenCount id="4C8F9EAF9B13A40ABBD08B99FB44F952" box="[1194,1243,1764,1790]" pageId="1" pageNumber="568" type="female">12♀</specimenCount>
(
<collectionCode id="3C98CDE39B13A40ABB908B99FAD8F952" box="[1258,1351,1764,1790]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="1" pageNumber="568" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="EAE15F7B9B13A40ABA238B98FA07F88E" collectingDate="2020-07-15" collectorName="A. A. Fomichev &amp; V. V. Sergeev" country="Russia" elevation="1300" latitude="52.816666" location="Pelekhova Mt." longLatPrecision="1081" longitude="91.51667" municipality="Borus Mt. Range" pageId="1" pageNumber="568" specimenCount="1" stateProvince="Krasnoyarsk" typeStatus="holotype">
<collectingMunicipality id="BA52CF5C9B13A40ABA238B98FE8EF88F" pageId="1" pageNumber="568">Borus Mt. Range</collectingMunicipality>
,
<location id="5F5603FD9B13A40ABE648A75FE5CF88F" LSID="urn:lsid:plazi:treatment:D220E4309B13A40EBFED88C8FE1CFBA7:5F5603FD9B13A40ABE648A75FE5CF88F" box="[286,451,1800,1827]" country="Russia" latitude="52.816666" longLatPrecision="1081" longitude="91.51667" municipality="Borus Mt. Range" name="Pelekhova Mt." pageId="1" pageNumber="568" stateProvince="Krasnoyarsk">Pelekhova Mt.</location>
Pass,
<geoCoordinate id="3FBD33E19B13A40ABD778A75FDEFF88F" box="[525,624,1800,1827]" degrees="52" direction="north" minutes="49" orientation="latitude" pageId="1" pageNumber="568" precision="925" value="52.816666">52°49’N</geoCoordinate>
,
<geoCoordinate id="3FBD33E19B13A40ABD078A75FD42F88E" box="[637,733,1800,1826]" degrees="91" direction="east" minutes="31" orientation="longitude" pageId="1" pageNumber="568" precision="925" value="91.51667">91°31’E</geoCoordinate>
, scree,
<quantity id="9D71F8C39B13A40ABC498A75FC14F88F" box="[819,907,1800,1827]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="568" unit="m" value="1300.0">
<elevation id="D1A4B2159B13A40ABC498A75FC14F88F" box="[819,907,1800,1827]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="568" unit="m" value="1300.0">1300 m</elevation>
</quantity>
, 13,
<date id="2E3773E69B13A40ABCCD8A75FBA3F88F" box="[951,1084,1800,1827]" pageId="1" pageNumber="568" value="2020-07-15">
<collectingDate id="3E738A0E9B13A40ABCCD8A75FBA3F88F" box="[951,1084,1800,1827]" pageId="1" pageNumber="568" value="2020-07-15">15.07.2020</collectingDate>
</date>
,
<collectorName id="F77C30F09B13A40ABB3D8A75FB68F88E" box="[1095,1271,1800,1826]" pageId="1" pageNumber="568">A.A. Fomichev</collectorName>
,
<collectorName id="F77C30F09B13A40ABA798A74FA07F88E" box="[1283,1432,1800,1827]" pageId="1" pageNumber="568">V.V. Sergeev</collectorName>
</materialsCitation>
;
<materialsCitation id="EAE15F7B9B13A40ABFED8A51FEDCF8EA" box="[151,323,1836,1862]" country="Russia" location="Republic" pageId="1" pageNumber="568" specimenCount="1" stateProvince="Tyva" typeStatus="holotype">
<emphasis id="68FD89349B13A40ABFED8A51FEDCF8EA" bold="true" box="[151,323,1836,1862]" pageId="1" pageNumber="568">
<collectingRegion id="984D9BC49B13A40ABFED8A51FF4CF8EA" box="[151,211,1836,1862]" country="Russia" name="Tyva" pageId="1" pageNumber="568">Tuva</collectingRegion>
<location id="5F5603FD9B13A40ABFA28A51FEDCF8EA" LSID="urn:lsid:plazi:treatment:D220E4309B13A40EBFED88C8FE1CFBA7:5F5603FD9B13A40ABFA28A51FEDCF8EA" box="[216,323,1836,1862]" country="Russia" name="Republic" pageId="1" pageNumber="568" stateProvince="Tyva">Republic</location>
</emphasis>
</materialsCitation>
:
<materialsCitation id="EAE15F7B9B13A40ABE2A8A51FB2FF8C6" collectingDate="2017-07-06" collectionCode="ISEA" collectorName="A. A. Fomichev" country="Russia" county="Western Sayan Mt. Range" elevation="2500" latitude="51.716667" location="Sabalkias Mt." longLatPrecision="1088" longitude="89.9" municipality="Sayanskyi Mt." pageId="1" pageNumber="568" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Tyva" typeStatus="holotype">
<specimenCount id="4C8F9EAF9B13A40ABE2A8A51FEECF8EA" box="[336,371,1836,1862]" pageId="1" pageNumber="568" type="male">3♂</specimenCount>
<specimenCount id="4C8F9EAF9B13A40ABE028A51FE05F8EA" box="[376,410,1836,1862]" pageId="1" pageNumber="568" type="female">3♀</specimenCount>
(
<collectionCode id="3C98CDE39B13A40ABEDD8A51FE76F8EA" box="[423,489,1836,1862]" pageId="1" pageNumber="568">ISEA</collectionCode>
, 001.8652),
<collectingCounty id="B3572DAA9B13A40ABD0B8A50FC0FF8EB" box="[625,912,1836,1863]" pageId="1" pageNumber="568">Western Sayan Mt. Range</collectingCounty>
,
<collectingMunicipality id="BA52CF5C9B13A40ABCE08A51FBA5F8EB" box="[922,1082,1836,1863]" pageId="1" pageNumber="568">Sayanskyi Mt.</collectingMunicipality>
Pass,
<quantity id="9D71F8C39B13A40ABB068A51FB2CF8EA" box="[1148,1203,1836,1863]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="568" unit="km" value="5.0">5 km</quantity>
SSW from
<location id="5F5603FD9B13A40ABA4B8A51FF5DF8C7" LSID="urn:lsid:plazi:treatment:D220E4309B13A40EBFED88C8FE1CFBA7:5F5603FD9B13A40ABA4B8A51FF5DF8C7" country="Russia" county="Western Sayan Mt. Range" latitude="51.716667" longLatPrecision="1088" longitude="89.9" municipality="Sayanskyi Mt." name="Sabalkias Mt." pageId="1" pageNumber="568" stateProvince="Tyva">Sabalkias Mt.</location>
,
<geoCoordinate id="3FBD33E19B13A40ABFB78A2DFEAFF8C7" box="[205,304,1872,1899]" degrees="51" direction="north" minutes="43" orientation="latitude" pageId="1" pageNumber="568" precision="925" value="51.716667">51°43’N</geoCoordinate>
,
<geoCoordinate id="3FBD33E19B13A40ABE468A2DFE03F8C7" box="[316,412,1872,1899]" degrees="89" direction="east" minutes="54" orientation="longitude" pageId="1" pageNumber="568" precision="925" value="89.9">89°54’E</geoCoordinate>
, scree and alpine meadow,
<quantity id="9D71F8C39B13A40ABDB48A2DFCF4F8C7" box="[718,875,1872,1899]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="2.6" metricValueMin="2.4" pageId="1" pageNumber="568" unit="m" value="2500.0" valueMax="2600.0" valueMin="2400.0">
<elevation id="D1A4B2159B13A40ABDB48A2DFCF4F8C7" box="[718,875,1872,1899]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="2.6" metricValueMin="2.4" pageId="1" pageNumber="568" unit="m" value="2500.0" valueMax="2600.0" valueMin="2400.0">2400–2600 m</elevation>
</quantity>
,
<date id="2E3773E69B13A40ABC0C8A2DFC67F8C6" box="[886,1016,1872,1898]" pageId="1" pageNumber="568" value="2017-07-06">
<collectingDate id="3E738A0E9B13A40ABC0C8A2DFC67F8C6" box="[886,1016,1872,1898]" pageId="1" pageNumber="568" value="2017-07-06">07.06.2017</collectingDate>
</date>
,
<collectorName id="F77C30F09B13A40ABB7B8A2DFB2FF8C6" box="[1025,1200,1872,1898]" pageId="1" pageNumber="568">A.A. Fomichev</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="5A3655269B13A40ABFBD8A09FAB6F87A" blockId="1.[151,1437,1656,2007]" pageId="1" pageNumber="568">
<emphasis id="68FD89349B13A40ABFBD8A09FE43F822" bold="true" box="[199,476,1908,1934]" pageId="1" pageNumber="568">Comparative material:</emphasis>
<taxonomicName id="9D892EA59B13A40ABE9C8A09FB1BF822" authority="Marusik, Hippa &amp; Koponen, 1996" authorityName="Marusik, Hippa &amp; Koponen" authorityYear="1996" box="[486,1156,1908,1935]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="568" phylum="Arthropoda" rank="species" species="azyuzini">
<emphasis id="68FD89349B13A40ABE9C8A09FD6FF822" box="[486,752,1908,1934]" italics="true" pageId="1" pageNumber="568">Acantholycosa azyuzini</emphasis>
<materialsCitation id="EAE15F7B9B13A40ABD838A09FB1BF822" box="[761,1156,1908,1935]" collectingDate="2019-07-26" collectionCode="ISEA" collectorName="A. A. Fomichev &amp; E. A. Nepaeva &amp; Yu. V. Dyachkov" country="Russia" county="Katunskyi Mt. Range" elevation="2400" latitude="49.98365" location="Kuyguk River" longLatPrecision="1" longitude="85.8516" municipality="Mul'ta River" pageId="1" pageNumber="568" specimenCount="1" specimenCount-female="1" stateProvince="Altay">
<bibRefCitation id="3E1828D79B13A40ABD838A09FB1BF822" author="Marusik, Y. M. &amp; Hippa, H. &amp; Koponen, S." box="[761,1156,1908,1935]" pageId="1" pageNumber="568" pagination="11 - 45" refId="ref10406" refString="Marusik, Y. M., Hippa, H. &amp; Koponen, S. (1996) Spiders (Araneae) from the Altai area, Southern Siberia. Acta Zoologica Fennica, 201, 11 - 45." type="journal article" year="1996">Marusik, Hippa &amp; Koponen, 1996</bibRefCitation>
</materialsCitation>
</taxonomicName>
(
<figureCitation id="C2B249A39B13A40ABBEF8A09FA83F823" box="[1173,1308,1908,1935]" captionStart="FIGURES 54–62" captionStartId="8.[151,264,1578,1602]" captionTargetBox="[151,1436,181,1554]" captionTargetId="figure-33@8.[151,1436,181,1554]" captionTargetPageId="8" captionText="FIGURES 54–62. Epigyne and female internal genitalia of Acantholycosa sayanensis (54–56), A. azyuzini (57–59) and A. sidorovi sp. n. (60–62). 54, 57, 60—intact, ventral; 55, 58, 61—macerated, ventral; 56, 59, 62—macerated, dorsal. Scale bars=0.2 mm. Abbreviations: Ap apical pocket, Cd copulatory ducts, Fo epigynal fovea, Li lip of the epigyne, Re receptacle, Sb septal base, Ss septal stem." figureDoi="http://doi.org/10.5281/zenodo.5300802" httpUri="https://zenodo.org/record/5300802/files/figure.png" pageId="1" pageNumber="568">Figs 57–59</figureCitation>
):
<collectingCountry id="229E15B69B13A40ABA4B8A09FA06F822" box="[1329,1433,1908,1934]" name="Russia" pageId="1" pageNumber="568">RUSSIA</collectingCountry>
:
<emphasis id="68FD89349B13A40ABFED8AE5FED5F81E" bold="true" box="[151,330,1944,1970]" pageId="1" pageNumber="568">
<collectingRegion id="984D9BC49B13A40ABFED8AE5FED5F81E" box="[151,330,1944,1970]" country="Russia" name="Altay" pageId="1" pageNumber="568">Altai Republic</collectingRegion>
</emphasis>
:
<specimenCount id="4C8F9EAF9B13A40ABE258AE5FE1EF81E" box="[351,385,1944,1970]" pageId="1" pageNumber="568" type="female">1♀</specimenCount>
(
<collectionCode id="3C98CDE39B13A40ABEEC8AE5FE47F81E" box="[406,472,1944,1970]" pageId="1" pageNumber="568">ISEA</collectionCode>
, 001.8663)
<collectingCounty id="B3572DAA9B13A40ABD128AE5FCF9F81F" box="[616,870,1944,1971]" pageId="1" pageNumber="568">Katunskyi Mt. Range</collectingCounty>
, watershed of
<collectingMunicipality id="BA52CF5C9B13A40ABB638AE5FB30F81E" box="[1049,1199,1944,1970]" pageId="1" pageNumber="568">Mul’ta River</collectingMunicipality>
and
<location id="5F5603FD9B13A40ABB8B8AE5FA07F81E" LSID="urn:lsid:plazi:treatment:D220E4309B13A40EBFED88C8FE1CFBA7:5F5603FD9B13A40ABB8B8AE5FA07F81E" box="[1265,1432,1944,1970]" country="Russia" county="Katunskyi Mt. Range" latitude="49.98365" longLatPrecision="1" longitude="85.8516" municipality="Mul'ta River" name="Kuyguk River" pageId="1" pageNumber="568" stateProvince="Altay">Kuyguk River</location>
,
<geoCoordinate id="3FBD33E19B13A40ABFED8AC1FEB3F87B" box="[151,300,1980,2007]" degrees="49" direction="north" minutes="59.019" orientation="latitude" pageId="1" pageNumber="568" precision="1" value="49.98365">49°59.019’N</geoCoordinate>
,
<geoCoordinate id="3FBD33E19B13A40ABE4D8AC1FE56F87B" box="[311,457,1980,2007]" degrees="85" direction="east" minutes="51.096" orientation="longitude" pageId="1" pageNumber="568" precision="1" value="85.8516">85°51.096’E</geoCoordinate>
, scree,
<quantity id="9D71F8C39B13A40ABD618AC1FDEDF87B" box="[539,626,1980,2007]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="1" pageNumber="568" unit="m" value="2400.0">
<elevation id="D1A4B2159B13A40ABD618AC1FDEDF87B" box="[539,626,1980,2007]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="1" pageNumber="568" unit="m" value="2400.0">2400 m</elevation>
</quantity>
,
<date id="2E3773E69B13A40ABD048AC1FD60F87A" box="[638,767,1980,2006]" pageId="1" pageNumber="568" value="2019-07-26">
<collectingDate id="3E738A0E9B13A40ABD048AC1FD60F87A" box="[638,767,1980,2006]" pageId="1" pageNumber="568" value="2019-07-26">26.07.2019</collectingDate>
</date>
,
<collectorName id="F77C30F09B13A40ABC728AC1FC28F87A" box="[776,951,1980,2006]" pageId="1" pageNumber="568">A.A. Fomichev</collectorName>
,
<collectorName id="F77C30F09B13A40ABCB88AC1FBFEF87B" box="[962,1121,1980,2007]" pageId="1" pageNumber="568">E.A. Nepaeva</collectorName>
,
<collectorName id="F77C30F09B13A40ABB118AC0FABAF87A" box="[1131,1317,1980,2007]" pageId="1" pageNumber="568">Yu.V. Dyachkov</collectorName>
.
</paragraph>
<caption id="0EF605AE9B10A409BFED8A55FDEFF824" ID-DOI="http://doi.org/10.5281/zenodo.5300794" ID-Zenodo-Dep="5300794" httpUri="https://zenodo.org/record/5300794/files/figure.png" pageId="2" pageNumber="569" startId="2.[151,264,1832,1856]" targetBox="[157,1436,189,1799]" targetPageId="2">
<paragraph id="5A3655269B10A409BFED8A55FDEFF824" blockId="2.[151,1436,1832,1929]" pageId="2" pageNumber="569">
<emphasis id="68FD89349B10A409BFED8A55FEA2F8EC" bold="true" box="[151,317,1832,1856]" pageId="2" pageNumber="569">FIGURES 1–9.</emphasis>
General appearance of
<taxonomicName id="9D892EA59B10A409BD4F8A54FCDEF8EC" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[565,833,1833,1857]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="569" phylum="Arthropoda" rank="species" species="sayanensis">
<emphasis id="68FD89349B10A409BD4F8A54FCDEF8EC" box="[565,833,1833,1857]" italics="true" pageId="2" pageNumber="569">Acantholycosa sayanensis</emphasis>
</taxonomicName>
(1–2),
<taxonomicName id="9D892EA59B10A409BCF58A54FC62F8ED" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[911,1021,1833,1857]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="569" phylum="Arthropoda" rank="species" species="kulikovi" status="sp. nov.">
<emphasis id="68FD89349B10A409BCF58A54FC62F8ED" box="[911,1021,1833,1857]" italics="true" pageId="2" pageNumber="569">A. kulikovi</emphasis>
</taxonomicName>
<emphasis id="68FD89349B10A409BB7F8A54FBDFF8ED" bold="true" box="[1029,1088,1833,1857]" pageId="2" pageNumber="569">
<taxonomicNameLabel id="73CE344F9B10A409BB7F8A54FBDFF8ED" box="[1029,1088,1833,1857]" pageId="2" pageNumber="569" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(3),
<taxonomicName id="9D892EA59B10A409BB0F8A54FB43F8ED" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[1141,1244,1833,1857]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="569" phylum="Arthropoda" rank="species" species="levinae">
<emphasis id="68FD89349B10A409BB0F8A54FB43F8ED" box="[1141,1244,1833,1857]" italics="true" pageId="2" pageNumber="569">A. levinae</emphasis>
</taxonomicName>
(4–5),
<taxonomicName id="9D892EA59B10A409BA518A54FA03F8EC" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[1323,1436,1833,1856]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="569" phylum="Arthropoda" rank="species" species="sergeevi" status="sp. nov.">
<emphasis id="68FD89349B10A409BA518A54FA03F8EC" box="[1323,1436,1833,1856]" italics="true" pageId="2" pageNumber="569">A. sergeevi</emphasis>
</taxonomicName>
<emphasis id="68FD89349B10A409BFED8A30FF4FF8C9" bold="true" box="[151,208,1869,1893]" pageId="2" pageNumber="569">
<taxonomicNameLabel id="73CE344F9B10A409BFED8A30FF4FF8C9" box="[151,208,1869,1893]" pageId="2" pageNumber="569" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(6),
<taxonomicName id="9D892EA59B10A409BE7B8A30FE24F8C9" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[257,443,1869,1893]" class="Arachnida" family="Lycosidae" genus="Mongolicosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="569" phylum="Arthropoda" rank="species" species="lizae" status="sp. nov.">
<emphasis id="68FD89349B10A409BE7B8A30FE24F8C9" box="[257,443,1869,1893]" italics="true" pageId="2" pageNumber="569">Mongolicosa lizae</emphasis>
</taxonomicName>
<emphasis id="68FD89349B10A409BEB88A30FE64F8C9" bold="true" box="[450,507,1869,1893]" pageId="2" pageNumber="569">
<taxonomicNameLabel id="73CE344F9B10A409BEB88A30FE64F8C9" box="[450,507,1869,1893]" pageId="2" pageNumber="569" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(8–9) and leg I of
<taxonomicName id="9D892EA59B10A409BDC38A30FCB7F8C8" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[697,808,1869,1892]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="569" phylum="Arthropoda" rank="species" species="sergeevi" status="sp. nov.">
<emphasis id="68FD89349B10A409BDC38A30FCB7F8C8" box="[697,808,1869,1892]" italics="true" pageId="2" pageNumber="569">A. sergeevi</emphasis>
</taxonomicName>
<emphasis id="68FD89349B10A409BC558A30FCF7F8C9" bold="true" box="[815,872,1869,1893]" pageId="2" pageNumber="569">
<taxonomicNameLabel id="73CE344F9B10A409BC558A30FCF7F8C9" box="[815,872,1869,1893]" pageId="2" pageNumber="569" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(7). 1, 4, 6, 8—male, dorsal; 2, 3, 5, 9—female, dorsal; 7—leg I of male, prolateral. Scale bars=2 mm.
</paragraph>
</caption>
<caption id="0EF605AE9B11A408BFED8A44FD11F835" ID-DOI="http://doi.org/10.5281/zenodo.5300796" ID-Zenodo-Dep="5300796" httpUri="https://zenodo.org/record/5300796/files/figure.png" pageId="3" pageNumber="570" startId="3.[151,264,1849,1873]" targetBox="[155,1425,188,1810]" targetPageId="3">
<paragraph id="5A3655269B11A408BFED8A44FD11F835" blockId="3.[151,1437,1849,1945]" pageId="3" pageNumber="570">
<emphasis id="68FD89349B11A408BFED8A44FEC9F8FD" bold="true" box="[151,342,1849,1873]" pageId="3" pageNumber="570">FIGURES 10–19.</emphasis>
Male palp (10–13) and bulb (14–19) of
<taxonomicName id="9D892EA59B11A408BD868A44FC73F8FD" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[764,1004,1849,1873]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="570" phylum="Arthropoda" rank="species" species="sergeevi" status="sp. nov.">
<emphasis id="68FD89349B11A408BD868A44FC73F8FD" box="[764,1004,1849,1873]" italics="true" pageId="3" pageNumber="570">Acantholycosa sergeevi</emphasis>
</taxonomicName>
<emphasis id="68FD89349B11A408BC8E8A44FBB0F8FD" bold="true" box="[1012,1071,1849,1873]" pageId="3" pageNumber="570">
<taxonomicNameLabel id="73CE344F9B11A408BC8E8A44FBB0F8FD" box="[1012,1071,1849,1873]" pageId="3" pageNumber="570" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(10–11),
<taxonomicName id="9D892EA59B11A408BBEE8A44FABFF8FD" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[1172,1312,1849,1873]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="570" phylum="Arthropoda" rank="species" species="sayanensis">
<emphasis id="68FD89349B11A408BBEE8A44FABFF8FD" box="[1172,1312,1849,1873]" italics="true" pageId="3" pageNumber="570">A. sayanensis</emphasis>
</taxonomicName>
(14–15),
<taxonomicName id="9D892EA59B11A408BAFC8A44FF76F8D9" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="570" phylum="Arthropoda" rank="species" species="sidorovi" status="sp. nov.">
<emphasis id="68FD89349B11A408BAFC8A44FF76F8D9" italics="true" pageId="3" pageNumber="570">A. sidorovi</emphasis>
</taxonomicName>
<emphasis id="68FD89349B11A408BF888A20FEB2F8D9" bold="true" box="[242,301,1885,1909]" pageId="3" pageNumber="570">
<taxonomicNameLabel id="73CE344F9B11A408BF888A20FEB2F8D9" box="[242,301,1885,1909]" pageId="3" pageNumber="570" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(16–17),
<taxonomicName id="9D892EA59B11A408BEEF8A20FE63F8D9" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[405,508,1885,1909]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="570" phylum="Arthropoda" rank="species" species="levinae">
<emphasis id="68FD89349B11A408BEEF8A20FE63F8D9" box="[405,508,1885,1909]" italics="true" pageId="3" pageNumber="570">A. levinae</emphasis>
</taxonomicName>
(18-19) and
<taxonomicName id="9D892EA59B11A408BDFD8A20FCDBF8D9" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[647,836,1885,1909]" class="Arachnida" family="Lycosidae" genus="Mongolicosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="570" phylum="Arthropoda" rank="species" species="lizae" status="sp. nov.">
<emphasis id="68FD89349B11A408BDFD8A20FCDBF8D9" box="[647,836,1885,1909]" italics="true" pageId="3" pageNumber="570">Mongolicosa lizae</emphasis>
</taxonomicName>
<emphasis id="68FD89349B11A408BC368A20FC17F8D9" bold="true" box="[844,904,1885,1909]" pageId="3" pageNumber="570">
<taxonomicNameLabel id="73CE344F9B11A408BC368A20FC17F8D9" box="[844,904,1885,1909]" pageId="3" pageNumber="570" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(12–13). 10, 12—retrolateral; 11, 13, 14, 16, 18— ventral; 15, 17, 19—anterior. Scale bars=0.2 mm.
</paragraph>
</caption>
<paragraph id="5A3655269B16A40FBFED8B0CFE51F95A" blockId="4.[151,1437,1649,1782]" pageId="4" pageNumber="571">
<emphasis id="68FD89349B16A40FBFED8B0CFEC8F926" bold="true" box="[151,343,1649,1674]" pageId="4" pageNumber="571">FIGURES 20–31.</emphasis>
Bulb (20, 24, 28) and embolic division (21–23, 25–27, 29–31) of
<taxonomicName id="9D892EA59B16A40FBB698B0FFA80F925" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[1043,1311,1650,1674]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="571" phylum="Arthropoda" rank="species" species="sayanensis">
<emphasis id="68FD89349B16A40FBB698B0FFA80F925" box="[1043,1311,1650,1674]" italics="true" pageId="4" pageNumber="571">Acantholycosa sayanensis</emphasis>
</taxonomicName>
(20–23),
<taxonomicName id="9D892EA59B16A40FBAFC8B0FFF76F902" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="571" phylum="Arthropoda" rank="species" species="sidorovi" status="sp. nov.">
<emphasis id="68FD89349B16A40FBAFC8B0FFF76F902" italics="true" pageId="4" pageNumber="571">A. sidorovi</emphasis>
</taxonomicName>
<emphasis id="68FD89349B16A40FBF8B8BEBFEB4F902" bold="true" box="[241,299,1686,1710]" pageId="4" pageNumber="571">
<taxonomicNameLabel id="73CE344F9B16A40FBF8B8BEBFEB4F902" box="[241,299,1686,1710]" pageId="4" pageNumber="571" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(24–27) and
<taxonomicName id="9D892EA59B16A40FBECC8BEBFD83F902" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[438,540,1686,1710]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="571" phylum="Arthropoda" rank="species" species="levinae">
<emphasis id="68FD89349B16A40FBECC8BEBFD83F902" box="[438,540,1686,1710]" italics="true" pageId="4" pageNumber="571">A. levinae</emphasis>
</taxonomicName>
(28–31). 20, 23–24, 27–28, 31—retrolateral; 21, 25, 29—ventral, 22, 26, 30—anterior. Scale bars=
<quantity id="9D71F8C39B16A40FBE748BC7FEFEF97D" box="[270,353,1722,1746]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="571" unit="mm" value="0.2">0.2 mm</quantity>
. Abbreviations:
<emphasis id="68FD89349B16A40FBD728BC7FDB5F97D" box="[520,554,1722,1745]" italics="true" pageId="4" pageNumber="571">Em</emphasis>
embolus,
<emphasis id="68FD89349B16A40FBDEF8BC7FD33F97E" box="[661,684,1722,1746]" italics="true" pageId="4" pageNumber="571">Pl</emphasis>
palea,
<emphasis id="68FD89349B16A40FBD8D8BC7FC8CF97D" box="[759,787,1722,1745]" italics="true" pageId="4" pageNumber="571">Po</emphasis>
paleal outgrowth,
<emphasis id="68FD89349B16A40FBCAF8BC7FC71F97D" box="[981,1006,1722,1745]" italics="true" pageId="4" pageNumber="571">Sp</emphasis>
spine,
<emphasis id="68FD89349B16A40FBB408BC7FBCDF97D" box="[1082,1106,1722,1745]" italics="true" pageId="4" pageNumber="571">Ta</emphasis>
terminal apophysis,
<emphasis id="68FD89349B16A40FBA538BC7FAA0F97D" box="[1321,1343,1722,1745]" italics="true" pageId="4" pageNumber="571">Ts</emphasis>
triangleshaped projection of the palea.
</paragraph>
</subSubSection>
<subSubSection id="129306AD9B16A40EBFBD8A59FE1CFE2B" lastPageId="5" lastPageNumber="572" pageId="4" pageNumber="571" type="diagnosis">
<paragraph id="5A3655269B16A40EBFBD8A59FE1CFE2B" blockId="4.[151,1437,1828,2034]" lastBlockId="5.[151,1437,151,1035]" lastPageId="5" lastPageNumber="572" pageId="4" pageNumber="571">
<emphasis id="68FD89349B16A40FBFBD8A59FEDDF892" bold="true" box="[199,322,1828,1854]" pageId="4" pageNumber="571">Diagnosis.</emphasis>
<taxonomicName id="9D892EA59B16A40FBE308A59FDEFF892" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[330,624,1828,1854]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="571" phylum="Arthropoda" rank="species" species="sayanensis">
<emphasis id="68FD89349B16A40FBE308A59FDEFF892" box="[330,624,1828,1854]" italics="true" pageId="4" pageNumber="571">Acantholycosa sayanensis</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="9D892EA59B16A40FBC738A59FC0BF892" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[777,916,1828,1854]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="571" phylum="Arthropoda" rank="species" species="altaiensis">
<emphasis id="68FD89349B16A40FBC738A59FC0BF892" box="[777,916,1828,1854]" italics="true" pageId="4" pageNumber="571">A. altaiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="9D892EA59B16A40FBCB48A59FBD4F892" authorityName="Marusik, Hippa &amp; Koponen" authorityYear="1996" box="[974,1099,1828,1854]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="571" phylum="Arthropoda" rank="species" species="azyuzini">
<emphasis id="68FD89349B16A40FBCB48A59FBD4F892" box="[974,1099,1828,1854]" italics="true" pageId="4" pageNumber="571">A. azyuzini</emphasis>
</taxonomicName>
. The female of
<taxonomicName id="9D892EA59B16A40FBA788A59FA03F892" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[1282,1436,1828,1854]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="571" phylum="Arthropoda" rank="species" species="sayanensis">
<emphasis id="68FD89349B16A40FBA788A59FA03F892" box="[1282,1436,1828,1854]" italics="true" pageId="4" pageNumber="571">A. sayanensis</emphasis>
</taxonomicName>
is similar to these species in having diamond-shaped epigynal fovea (
<emphasis id="68FD89349B16A40FBCBF8A34FC7BF8CE" box="[965,996,1865,1890]" italics="true" pageId="4" pageNumber="571">Fo</emphasis>
). The female of
<taxonomicName id="9D892EA59B16A40FBBD78A35FAD6F8CE" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[1197,1353,1864,1890]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="571" phylum="Arthropoda" rank="species" species="sayanensis">
<emphasis id="68FD89349B16A40FBBD78A35FAD6F8CE" box="[1197,1353,1864,1890]" italics="true" pageId="4" pageNumber="571">A. sayanensis</emphasis>
</taxonomicName>
differs from those of
<taxonomicName id="9D892EA59B16A40FBE478A11FE55F82A" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[317,458,1900,1926]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="571" phylum="Arthropoda" rank="species" species="altaiensis">
<emphasis id="68FD89349B16A40FBE478A11FE55F82A" box="[317,458,1900,1926]" italics="true" pageId="4" pageNumber="571">A. altaiensis</emphasis>
</taxonomicName>
by having septal base (
<emphasis id="68FD89349B16A40FBD9A8A11FD63F82A" box="[736,764,1900,1926]" italics="true" pageId="4" pageNumber="571">Sb</emphasis>
) almost as large as the epigynal fovea (vs. much smaller) and diverging receptacles (
<emphasis id="68FD89349B16A40FBEB38AECFE78F806" box="[457,487,1937,1962]" italics="true" pageId="4" pageNumber="571">Re</emphasis>
) (cf.
<figureCitation id="C2B249A39B16A40FBD5D8AEDFD39F807" box="[551,678,1936,1963]" captionStart="FIGURES 54–62" captionStartId="8.[151,264,1578,1602]" captionTargetBox="[151,1436,181,1554]" captionTargetId="figure-33@8.[151,1436,181,1554]" captionTargetPageId="8" captionText="FIGURES 54–62. Epigyne and female internal genitalia of Acantholycosa sayanensis (54–56), A. azyuzini (57–59) and A. sidorovi sp. n. (60–62). 54, 57, 60—intact, ventral; 55, 58, 61—macerated, ventral; 56, 59, 62—macerated, dorsal. Scale bars=0.2 mm. Abbreviations: Ap apical pocket, Cd copulatory ducts, Fo epigynal fovea, Li lip of the epigyne, Re receptacle, Sb septal base, Ss septal stem." figureDoi="http://doi.org/10.5281/zenodo.5300802" httpUri="https://zenodo.org/record/5300802/files/figure.png" pageId="4" pageNumber="571">Figs 54–56</figureCitation>
and figs 8, 9 in
<bibRefCitation id="3E1828D79B16A40FBC1E8AEDFBD6F806" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." box="[868,1097,1936,1962]" pageId="4" pageNumber="571" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">
Marusik
<emphasis id="68FD89349B16A40FBCB08AECFB9CF806" box="[970,1027,1936,1962]" italics="true" pageId="4" pageNumber="571">et al.</emphasis>
2004
</bibRefCitation>
). It can be distinguished from the female of
<taxonomicName id="9D892EA59B16A40FBE428AC9FE29F862" authorityName="Marusik, Hippa &amp; Koponen" authorityYear="1996" box="[312,438,1972,1998]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="571" phylum="Arthropoda" rank="species" species="azyuzini">
<emphasis id="68FD89349B16A40FBE428AC9FE29F862" box="[312,438,1972,1998]" italics="true" pageId="4" pageNumber="571">A. azyuzini</emphasis>
</taxonomicName>
by the rugose (vs. smooth) surface of the septal base (cf.
<figureCitation id="C2B249A39B16A40FBB2F8AC9FB45F863" box="[1109,1242,1972,1999]" captionStart="FIGURES 54–62" captionStartId="8.[151,264,1578,1602]" captionTargetBox="[151,1436,181,1554]" captionTargetId="figure-33@8.[151,1436,181,1554]" captionTargetPageId="8" captionText="FIGURES 54–62. Epigyne and female internal genitalia of Acantholycosa sayanensis (54–56), A. azyuzini (57–59) and A. sidorovi sp. n. (60–62). 54, 57, 60—intact, ventral; 55, 58, 61—macerated, ventral; 56, 59, 62—macerated, dorsal. Scale bars=0.2 mm. Abbreviations: Ap apical pocket, Cd copulatory ducts, Fo epigynal fovea, Li lip of the epigyne, Re receptacle, Sb septal base, Ss septal stem." figureDoi="http://doi.org/10.5281/zenodo.5300802" httpUri="https://zenodo.org/record/5300802/files/figure.png" pageId="4" pageNumber="571">Figs 54, 57</figureCitation>
). The male of
<taxonomicName id="9D892EA59B16A40FBAFE8AC9FE8FF85E" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="571" phylum="Arthropoda" rank="species" species="sayanensis">
<emphasis id="68FD89349B16A40FBAFE8AC9FE8FF85E" italics="true" pageId="4" pageNumber="571">A. sayanensis</emphasis>
</taxonomicName>
is similar to those of
<taxonomicName id="9D892EA59B16A40FBD7E8AA5FD11F85E" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[516,654,2008,2034]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="571" phylum="Arthropoda" rank="species" species="altaiensis">
<emphasis id="68FD89349B16A40FBD7E8AA5FD11F85E" box="[516,654,2008,2034]" italics="true" pageId="4" pageNumber="571">A. altaiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="9D892EA59B16A40FBDBF8AA5FCDFF85E" authorityName="Marusik, Hippa &amp; Koponen" authorityYear="1996" box="[709,832,2008,2034]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="571" phylum="Arthropoda" rank="species" species="azyuzini">
<emphasis id="68FD89349B16A40FBDBF8AA5FCDFF85E" box="[709,832,2008,2034]" italics="true" pageId="4" pageNumber="571">A. azyuzini</emphasis>
</taxonomicName>
in having broad and straight embolus (
<emphasis id="68FD89349B16A40FBB8D8AA4FA83F85E" box="[1271,1308,2009,2034]" italics="true" pageId="4" pageNumber="571">Em</emphasis>
), claw-like paleal outgrowth (
<emphasis id="68FD89349B17A40EBE1D8DEAFE19FF1C" box="[359,390,151,176]" italics="true" pageId="5" pageNumber="572">Po</emphasis>
) and spine-like terminal apophysis (
<emphasis id="68FD89349B17A40EBC528DEAFCDCFF1C" box="[808,835,151,176]" italics="true" pageId="5" pageNumber="572">Ta</emphasis>
). The males of
<taxonomicName id="9D892EA59B17A40EBC838DEAFB0CFF1C" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[1017,1171,151,177]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="572" phylum="Arthropoda" rank="species" species="sayanensis">
<emphasis id="68FD89349B17A40EBC838DEAFB0CFF1C" box="[1017,1171,151,177]" italics="true" pageId="5" pageNumber="572">A. sayanensis</emphasis>
</taxonomicName>
differ from those of
<taxonomicName id="9D892EA59B17A40EBAFE8DEAFE9DFF79" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="572" phylum="Arthropoda" rank="species" species="altaiensis">
<emphasis id="68FD89349B17A40EBAFE8DEAFE9DFF79" italics="true" pageId="5" pageNumber="572">A. altaiensis</emphasis>
</taxonomicName>
in having rounded and solid anterior part of the tegular apophysis (
<emphasis id="68FD89349B17A40EBC968DC6FB95FF78" box="[1004,1034,187,212]" italics="true" pageId="5" pageNumber="572">Tg</emphasis>
) (vs. triangular and laminar anterior part), a small, almost indistinct triangular spine on the base of the embolus (vs. large outgrowth) and terminal apophysis bent outward (vs. directed inward) (cf. Figs 21–22, 32 and figs 2–3, 6 in
<bibRefCitation id="3E1828D79B17A40EBB398C7FFAA3FEB0" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." box="[1091,1340,258,285]" pageId="5" pageNumber="572" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">Marusik et al. (2004))</bibRefCitation>
. Finally, the male of
<taxonomicName id="9D892EA59B17A40EBE5E8C5BFE20FEEC" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[292,447,294,320]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="572" phylum="Arthropoda" rank="species" species="sayanensis">
<emphasis id="68FD89349B17A40EBE5E8C5BFE20FEEC" box="[292,447,294,320]" italics="true" pageId="5" pageNumber="572">A. sayanensis</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="9D892EA59B17A40EBD868C5BFCE5FEEC" authorityName="Marusik, Hippa &amp; Koponen" authorityYear="1996" box="[764,890,294,320]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="572" phylum="Arthropoda" rank="species" species="azyuzini">
<emphasis id="68FD89349B17A40EBD868C5BFCE5FEEC" box="[764,890,294,320]" italics="true" pageId="5" pageNumber="572">A. azyuzini</emphasis>
</taxonomicName>
by the embolus with a widened distal part (vs. pointed) and by the presence of a small spine on the base of embolus (vs. absence, cf. Figs 21–22 and fig.
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<bibRefCitation id="3E1828D79B17A40EBFED8C10FEE8FE2B" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." box="[151,375,365,391]" pageId="5" pageNumber="572" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">
Marusik
<emphasis id="68FD89349B17A40EBF868C13FEABFE2B" box="[252,308,365,391]" italics="true" pageId="5" pageNumber="572">et al.</emphasis>
2004
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="129306AD9B17A40EBFBD8CECFBA7FC33" pageId="5" pageNumber="572" type="description">
<paragraph id="5A3655269B17A40EBFBD8CECFDE0FD2E" blockId="5.[151,1437,151,1035]" pageId="5" pageNumber="572">
<emphasis id="68FD89349B17A40EBFBD8CECFEC6FE07" bold="true" box="[199,345,401,427]" pageId="5" pageNumber="572">Description.</emphasis>
Female. Total length 9.6. Carapace: 4.4 long, 3.6 wide. General appearance as in
<figureCitation id="C2B249A39B17A40EBBA88CECFA8BFE07" box="[1234,1300,401,427]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1832,1856]" captionTargetBox="[157,1436,189,1799]" captionTargetId="figure-33@2.[151,1436,181,1809]" captionTargetPageId="2" captionText="FIGURES 1–9. General appearance of Acantholycosa sayanensis (1–2), A. kulikovi sp. n. (3), A. levinae (4–5), A. sergeevi sp. n. (6), Mongolicosa lizae sp. n. (8–9) and leg I of A. sergeevi sp. n. (7). 1, 4, 6, 8—male, dorsal; 2, 3, 5, 9—female, dorsal; 7—leg I of male, prolateral. Scale bars=2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300794" httpUri="https://zenodo.org/record/5300794/files/figure.png" pageId="5" pageNumber="572">Fig. 2</figureCitation>
. Coloration. Carapace, chelicerae and sternum dark brown. Eye field black. Endites and labium yellow-brown. Coxae and palps brown with yellow spots. Legs dark brown. Femora with yellow-brown annulation. Abdomen dark gray, almost black dorsally, with dark brown cardiac mark. Venter of abdomen yellow-gray. Spinnerets dark gray, almost black. Spination of leg I: Fe d1-1-1 p0-0-2 r1-0-1; Pa r1; Ti p1-0-0 r1-0-0 v2-2-2-2-2-2; Mt p1-1-0 r1-1-0 v2-2-0. Leg measurements: I 4.35, 1.95, 4.05, 3.8, 1.8 (15.95); II 4.4, 1.95, 3.75, 3.95, 1.75 (15.8); III 4.3, 1.8, 3.5, 4.8, 1.9 (16.3); IV 5.25, 1.95, 4.6, 7.25, 2.55 (21.6).
</paragraph>
<paragraph id="5A3655269B17A40EBFBD8FF6FD5DFCBC" blockId="5.[151,1437,151,1035]" pageId="5" pageNumber="572">
Epigyne and internal genitalia as in
<figureCitation id="C2B249A39B17A40EBD2C8FF6FD49FD0A" box="[598,726,651,678]" captionStart="FIGURES 54–62" captionStartId="8.[151,264,1578,1602]" captionTargetBox="[151,1436,181,1554]" captionTargetId="figure-33@8.[151,1436,181,1554]" captionTargetPageId="8" captionText="FIGURES 54–62. Epigyne and female internal genitalia of Acantholycosa sayanensis (54–56), A. azyuzini (57–59) and A. sidorovi sp. n. (60–62). 54, 57, 60—intact, ventral; 55, 58, 61—macerated, ventral; 56, 59, 62—macerated, dorsal. Scale bars=0.2 mm. Abbreviations: Ap apical pocket, Cd copulatory ducts, Fo epigynal fovea, Li lip of the epigyne, Re receptacle, Sb septal base, Ss septal stem." figureDoi="http://doi.org/10.5281/zenodo.5300802" httpUri="https://zenodo.org/record/5300802/files/figure.png" pageId="5" pageNumber="572">Figs 54–56</figureCitation>
. Fovea as wide as long. Septal base fills almost the entire fovea. Septal stem (
<emphasis id="68FD89349B17A40EBE578FD2FED9FD65" box="[301,326,687,713]" italics="true" pageId="5" pageNumber="572">Ss</emphasis>
) as long as septal base. Lips of the epigyne (
<emphasis id="68FD89349B17A40EBC348FCDFCFAFD65" box="[846,869,688,713]" italics="true" pageId="5" pageNumber="572">Li</emphasis>
) not touching each other. Apical pocket (
<emphasis id="68FD89349B17A40EBA3B8FD2FAFFFD65" box="[1345,1376,687,713]" italics="true" pageId="5" pageNumber="572">Ap</emphasis>
) two times narrower than septal stem length, with one hood. Receptacles not clearly delineated from copulatory ducts (
<emphasis id="68FD89349B17A40EBFDB8F8BFF5DFCBC" box="[161,194,758,784]" italics="true" pageId="5" pageNumber="572">Cd</emphasis>
). Copulatory ducts as long as the receptacles.
</paragraph>
<paragraph id="5A3655269B17A40EBFBD8E67FBA7FC33" blockId="5.[151,1437,151,1035]" pageId="5" pageNumber="572">
Male. See
<bibRefCitation id="3E1828D79B17A40EBE3B8E67FDABFC98" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." box="[321,564,794,820]" pageId="5" pageNumber="572" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">
Marusik
<emphasis id="68FD89349B17A40EBEDD8E66FE7EFC98" box="[423,481,794,820]" italics="true" pageId="5" pageNumber="572">et al.</emphasis>
(2004)
</bibRefCitation>
for description. General appearance as in
<figureCitation id="C2B249A39B17A40EBB628E67FBC3FC98" box="[1048,1116,794,820]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1832,1856]" captionTargetBox="[157,1436,189,1799]" captionTargetId="figure-33@2.[151,1436,181,1809]" captionTargetPageId="2" captionText="FIGURES 1–9. General appearance of Acantholycosa sayanensis (1–2), A. kulikovi sp. n. (3), A. levinae (4–5), A. sergeevi sp. n. (6), Mongolicosa lizae sp. n. (8–9) and leg I of A. sergeevi sp. n. (7). 1, 4, 6, 8—male, dorsal; 2, 3, 5, 9—female, dorsal; 7—leg I of male, prolateral. Scale bars=2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300794" httpUri="https://zenodo.org/record/5300794/files/figure.png" pageId="5" pageNumber="572">Fig. 1</figureCitation>
. Palp as in
<figureCitation id="C2B249A39B17A40EBB9E8E67FAFAFC99" box="[1252,1381,794,821]" captionStart="FIGURES 10–19" captionStartId="3.[151,264,1849,1873]" captionTargetBox="[155,1425,188,1810]" captionTargetId="figure-15@3.[151,1436,181,1825]" captionTargetPageId="3" captionText="FIGURES 10–19. Male palp (10–13) and bulb (14–19) of Acantholycosa sergeevi sp. n. (10–11), A. sayanensis (14–15), A. sidorovi sp. n. (16–17), A. levinae (18-19) and Mongolicosa lizae sp. n. (12–13). 10, 12—retrolateral; 11, 13, 14, 16, 18— ventral; 15, 17, 19—anterior. Scale bars=0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300796" httpUri="https://zenodo.org/record/5300796/files/figure.png" pageId="5" pageNumber="572">Figs 14–15</figureCitation>
, 20– 23, 32–33. Tegular apophysis hammer-shaped, without apical arm and with a short basal arm (
<emphasis id="68FD89349B17A40EBB958E42FA91FCF4" box="[1263,1294,831,856]" italics="true" pageId="5" pageNumber="572">Ba</emphasis>
). Palea (
<emphasis id="68FD89349B17A40EBA008E43FA0CFCF4" box="[1402,1427,830,856]" italics="true" pageId="5" pageNumber="572">Pl</emphasis>
) rounded. Paleal outgrowth large, hooked. Terminal apophysis spine-like in the apical part, almost touching the paleal outgrowth. Embolus expanded distally, with a hook-like tip in anterior view.
</paragraph>
</subSubSection>
<subSubSection id="129306AD9B17A40EBFBD8ED4FE1CFBA7" pageId="5" pageNumber="572" type="distribution">
<paragraph id="5A3655269B17A40EBFBD8ED4FE1CFBA7" blockId="5.[151,1437,151,1035]" pageId="5" pageNumber="572">
<emphasis id="68FD89349B17A40EBFBD8ED4FEFEFC6F" bold="true" box="[199,353,937,963]" pageId="5" pageNumber="572">Distribution.</emphasis>
The species is known from three localities in the Western Sayan Mountains, which are located more than
<quantity id="9D71F8C39B17A40EBE6B8EB0FEF9FC4B" box="[273,358,973,999]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="572" unit="km" value="100.0">100 km</quantity>
apart (
<figureCitation id="C2B249A39B17A40EBECF8EB0FDA5FC44" box="[437,570,973,1000]" captionStart="FIGURES 72–77" captionStartId="13.[151,264,1676,1700]" captionTargetBox="[151,1436,309,1652]" captionTargetId="figure-76@13.[151,1436,309,1652]" captionTargetPageId="13" captionText="FIGURES 72–77. Male of Acantholycosa sayanensis from Borus Mt. Range (72), female of Acantholycosa sp. (A. azyuzini or A. levinae) with egg sac, from Katunskyi Mt. Range (73), habitat of A. sayanensis and A. sergeevi sp. n. in Borus Mt. Range (74), collecting localities of studied species from the Acantholycosa-complex (75–76) and map showing the main mountain systems of South Siberia and adjacent regions (77). Diamond—A. sayanensis, square—A. sidorovi sp. n., triangle—A. kulikovi sp. n., star—A. levinae, circle—A. sergeevi sp. n., inverted triangle—Mongolicosa lizae sp. n. The frame of Fig. 76 refers to the content of Fig. 75." figureDoi="http://doi.org/10.5281/zenodo.5300806" httpUri="https://zenodo.org/record/5300806/files/figure.png" pageId="5" pageNumber="572">Figs 74–76</figureCitation>
). This species appears to have one of the broadest ranges among its endemic to Siberia congeners.
</paragraph>
</subSubSection>
</treatment>
</document>

347
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<mods:title id="5EA08E312746C752A3B6A6A02D89EABE">New data on the wolf spiders from theAcantholycosa-complex (Araneae: Lycosidae) from the South Siberia</mods:title>
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<heading id="017EE24A9B17A40EBFED8932FE65FBC6" bold="true" box="[151,506,1103,1130]" fontSize="11" level="1" pageId="5" pageNumber="572" reason="1">
<emphasis id="68FD89349B17A40EBFED8932FE65FBC6" bold="true" box="[151,506,1103,1130]" pageId="5" pageNumber="572">
<taxonomicName id="9D892EA59B17A40EBFED8932FE2FFBC6" ID-CoL="999ML" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[151,432,1103,1130]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="572" phylum="Arthropoda" rank="species" species="sidorovi" status="sp. nov.">
<emphasis id="68FD89349B17A40EBFED8932FE2FFBC6" bold="true" box="[151,432,1103,1130]" italics="true" pageId="5" pageNumber="572">Acantholycosa sidorovi</emphasis>
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<figureCitation id="C2B249A39B17A40EBFED890EFE88FB21" box="[151,279,1139,1165]" captionStart="FIGURES 10–19" captionStartId="3.[151,264,1849,1873]" captionTargetBox="[155,1425,188,1810]" captionTargetId="figure-15@3.[151,1436,181,1825]" captionTargetPageId="3" captionText="FIGURES 10–19. Male palp (10–13) and bulb (14–19) of Acantholycosa sergeevi sp. n. (10–11), A. sayanensis (14–15), A. sidorovi sp. n. (16–17), A. levinae (18-19) and Mongolicosa lizae sp. n. (12–13). 10, 12—retrolateral; 11, 13, 14, 16, 18— ventral; 15, 17, 19—anterior. Scale bars=0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300796" httpUri="https://zenodo.org/record/5300796/files/figure.png" pageId="5" pageNumber="572">Figs 16–17</figureCitation>
, 24–27, 34–35, 60–62, 75–76
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<paragraph id="5A3655269B17A40EBFED89C9FBB2FA95" blockId="5.[151,1437,1204,2041]" pageId="5" pageNumber="572">
<emphasis id="68FD89349B17A40EBFED89C9FF7BFB62" bold="true" box="[151,228,1204,1230]" pageId="5" pageNumber="572">Types.</emphasis>
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<collectingRegion id="984D9BC49B17A40EBE2589C9FE45FB62" box="[351,474,1204,1230]" country="Russia" name="Altay" pageId="5" pageNumber="572">Altai Krai</collectingRegion>
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:
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(
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, 001.8653),
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, without precise locality and date, collector unknown (donated by
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.
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. Sidorov)
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;
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:
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(
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, 001.8654),
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(
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, 001.8724), together with the
</materialsCitation>
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(
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, 001.8655),
<collectingMunicipality id="BA52CF5C9B17A40EBDF48987FCC1FAB9" box="[654,862,1274,1301]" pageId="5" pageNumber="572">Tigirek Mt. Range</collectingMunicipality>
, the upper reaches of
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,
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,
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,
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<elevation id="D1A4B2159B17A40EBE798860FE3FFA94" box="[259,416,1309,1336]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="5" pageNumber="572" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">1400–1500 m</elevation>
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,
<date id="2E3773E69B17A40EBED68860FD80FA9B" box="[428,543,1309,1335]" pageId="5" pageNumber="572" value="2016-07-09">
<collectingDate id="3E738A0E9B17A40EBED68860FD80FA9B" box="[428,543,1309,1335]" pageId="5" pageNumber="572" value="2016-07-09">9.07.2016</collectingDate>
</date>
, collector unknown (donated by
<collectionCode id="3C98CDE39B17A40EBCF78863FC3EFA94" box="[909,929,1310,1336]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="572" type="Museum">V</collectionCode>
.
<collectionCode id="3C98CDE39B17A40EBCDF8863FC26FA94" box="[933,953,1310,1336]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="572" type="Museum">V</collectionCode>
. Sidorov)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="129306AD9B17A40EBFBD883CFD4EFAD2" pageId="5" pageNumber="572" type="etymology">
<paragraph id="5A3655269B17A40EBFBD883CFD4EFAD2" blockId="5.[151,1437,1204,2041]" pageId="5" pageNumber="572">
<emphasis id="68FD89349B17A40EBFBD883CFED1FAF7" bold="true" box="[199,334,1345,1371]" pageId="5" pageNumber="572">Etymology.</emphasis>
The specific name is a patronym given in honor of my friend, Vladislav V. Sidorov (Barnaul,
<collectingCountry id="229E15B69B17A40EBFED8819FF79FAD3" box="[151,230,1380,1407]" name="Russia" pageId="5" pageNumber="572">Russia</collectingCountry>
), who donated the
<typeStatus id="8532EB849B17A40EBEC38818FE6BFAD3" box="[441,500,1381,1407]" pageId="5" pageNumber="572">types</typeStatus>
of the new species.
</paragraph>
</subSubSection>
<subSubSection id="129306AD9B17A40EBFBD88FAFD29F934" pageId="5" pageNumber="572" type="diagnosis">
<paragraph id="5A3655269B17A40EBFBD88FAFD29F934" blockId="5.[151,1437,1204,2041]" pageId="5" pageNumber="572">
<emphasis id="68FD89349B17A40EBFBD88FAFEDDFA0D" bold="true" box="[199,322,1415,1441]" pageId="5" pageNumber="572">Diagnosis.</emphasis>
The new species is closely related to
<taxonomicName id="9D892EA59B17A40EBDA588F5FCF7FA0D" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[735,872,1415,1442]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="572" phylum="Arthropoda" rank="species" species="altaiensis">
<emphasis id="68FD89349B17A40EBDA588F5FCF7FA0D" box="[735,872,1415,1442]" italics="true" pageId="5" pageNumber="572">A. altaiensis</emphasis>
</taxonomicName>
, which is also known from the Western
<collectingRegion id="984D9BC49B17A40EBA5B88FAFAC2FA0D" box="[1313,1373,1415,1441]" country="Russia" name="Altay" pageId="5" pageNumber="572">Altai</collectingRegion>
. Both species have a spine (
<emphasis id="68FD89349B17A40EBEF388D6FE3AFA69" box="[393,421,1451,1477]" italics="true" pageId="5" pageNumber="572">Sp</emphasis>
) near base of the embolus (
<emphasis id="68FD89349B17A40EBDA688D6FC9EFA68" box="[732,769,1451,1476]" italics="true" pageId="5" pageNumber="572">Em</emphasis>
), laminar apical arm (
<emphasis id="68FD89349B17A40EBC8088D6FB86FA69" box="[1018,1049,1451,1477]" italics="true" pageId="5" pageNumber="572">Aa</emphasis>
) of the tegular apophysis (
<emphasis id="68FD89349B17A40EBA3C88D6FAFBFA68" box="[1350,1380,1451,1476]" italics="true" pageId="5" pageNumber="572">Tg</emphasis>
) and diamond-shaped epigynal fovea (
<emphasis id="68FD89349B17A40EBD7788B3FDB3FA4B" box="[525,556,1486,1511]" italics="true" pageId="5" pageNumber="572">Fo</emphasis>
) with pentagonal septal base (
<emphasis id="68FD89349B17A40EBCFB88B3FC02FA44" box="[897,925,1486,1512]" italics="true" pageId="5" pageNumber="572">Sb</emphasis>
). The male of
<taxonomicName id="9D892EA59B17A40EBB3B88B3FB24FA44" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[1089,1211,1486,1512]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="572" phylum="Arthropoda" rank="species" species="sidorovi" status="sp. nov.">
<emphasis id="68FD89349B17A40EBB3B88B3FB24FA44" box="[1089,1211,1486,1512]" italics="true" pageId="5" pageNumber="572">A. sidorovi</emphasis>
</taxonomicName>
<emphasis id="68FD89349B17A40EBBB888B3FA9EFA44" bold="true" box="[1218,1281,1486,1512]" pageId="5" pageNumber="572">
<taxonomicNameLabel id="73CE344F9B17A40EBBB888B3FA9EFA44" box="[1218,1281,1486,1512]" pageId="5" pageNumber="572" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs by the enlarged and more curved paleal outgrowth (
<emphasis id="68FD89349B17A40EBDE3888FFD27F9A7" box="[665,696,1522,1547]" italics="true" pageId="5" pageNumber="572">Po</emphasis>
), more prominent spine near base of the embolus (in retrolateral view) and by underdeveloped apical arm of the tegular apophysis (cf. Figs 25, 27, 34 and figs 2, 4,
<quantity id="9D71F8C39B17A40EBB8B8B69FA83F982" box="[1265,1308,1556,1582]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="5" pageNumber="572" unit="in" value="6.0">6 in</quantity>
<bibRefCitation id="3E1828D79B17A40EBA598B69FF68F9FD" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." pageId="5" pageNumber="572" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">
Marusik
<emphasis id="68FD89349B17A40EBAF28B68FF32F9FD" italics="true" pageId="5" pageNumber="572">et al</emphasis>
. 2004
</bibRefCitation>
). The female of the new species can be separated from the female of
<taxonomicName id="9D892EA59B17A40EBC8F8B45FBE1F9FD" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[1013,1150,1591,1618]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="572" phylum="Arthropoda" rank="species" species="altaiensis">
<emphasis id="68FD89349B17A40EBC8F8B45FBE1F9FD" box="[1013,1150,1591,1618]" italics="true" pageId="5" pageNumber="572">A. altaiensis</emphasis>
</taxonomicName>
by the apical pocket (
<emphasis id="68FD89349B17A40EBA0E8B45FA0CF9FE" box="[1396,1427,1592,1618]" italics="true" pageId="5" pageNumber="572">Ap</emphasis>
) almost as wide as the epigynal fovea (vs. significantly narrower) and by the longer and thinner receptacles (
<emphasis id="68FD89349B17A40EBA2E8B26FAEDF9D8" box="[1364,1394,1627,1652]" italics="true" pageId="5" pageNumber="572">Re</emphasis>
, cf.
<figureCitation id="C2B249A39B17A40EBFED8B03FE8BF93B" box="[151,276,1661,1688]" captionStart="FIGURES 54–62" captionStartId="8.[151,264,1578,1602]" captionTargetBox="[151,1436,181,1554]" captionTargetId="figure-33@8.[151,1436,181,1554]" captionTargetPageId="8" captionText="FIGURES 54–62. Epigyne and female internal genitalia of Acantholycosa sayanensis (54–56), A. azyuzini (57–59) and A. sidorovi sp. n. (60–62). 54, 57, 60—intact, ventral; 55, 58, 61—macerated, ventral; 56, 59, 62—macerated, dorsal. Scale bars=0.2 mm. Abbreviations: Ap apical pocket, Cd copulatory ducts, Fo epigynal fovea, Li lip of the epigyne, Re receptacle, Sb septal base, Ss septal stem." figureDoi="http://doi.org/10.5281/zenodo.5300802" httpUri="https://zenodo.org/record/5300802/files/figure.png" pageId="5" pageNumber="572">Figs 60, 62</figureCitation>
and figs
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<bibRefCitation id="3E1828D79B17A40EBEB08B03FD35F934" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." box="[458,682,1662,1688]" pageId="5" pageNumber="572" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">
Marusik
<emphasis id="68FD89349B17A40EBD548B03FDC0F934" box="[558,607,1662,1688]" italics="true" pageId="5" pageNumber="572">et al</emphasis>
. 2004
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="129306AD9B17A402BFBD8BDCFB18FF79" lastPageId="9" lastPageNumber="576" pageId="5" pageNumber="572" type="description">
<paragraph id="5A3655269B17A40EBFBD8BDCFF6FF823" blockId="5.[151,1437,1204,2041]" pageId="5" pageNumber="572">
<emphasis id="68FD89349B17A40EBFBD8BDCFEC6F917" bold="true" box="[199,345,1697,1723]" pageId="5" pageNumber="572">Description.</emphasis>
Male (
<typeStatus id="8532EB849B17A40EBEDC8BDCFD90F910" box="[422,527,1697,1724]" pageId="5" pageNumber="572" type="holotype">holotype</typeStatus>
). Total length 7.5. Carapace: 3.8 long, 3.2 wide. Coloration. Carapace dark brown. Eye field almost black. Chelicerae, sternum and palps brown. Labium, endites and coxae yellow-gray. Legs yellowbrown. Femora of legs with brown transverse stripes and longitudinal yellow stripes. Abdomen yellow-brown dorsally. Cardiac mark almost indistinct. Venter of the abdomen yellow-gray. Spinnerets dark gray. Spination of leg I: Fe d1-1-1 p0-0-2 r0-1-1; Ti p1-0-0 r1-0-0 v2-2-2-2-1-2; Mt p1-1-0 r0-1-0 v2-2-0. Leg measurements: I 4.4, 1.7, 4.2, 4.6, 1.8 (16.7); II 4.3, 1.7, 4.0, 4.7, 1.85 (16.55); III 4.2, 1.55, 3.6, 5.45, 1.85 (16.65); IV 4.9, 1.6, 4.3, 6.3, 1.55 (18.65).
</paragraph>
<paragraph id="5A3655269B17A40EBFBD8AEAFDB4F855" blockId="5.[151,1437,1204,2041]" pageId="5" pageNumber="572">
Palp as in
<figureCitation id="C2B249A39B17A40EBE428AEAFE28F81D" box="[312,439,1943,1969]" captionStart="FIGURES 10–19" captionStartId="3.[151,264,1849,1873]" captionTargetBox="[155,1425,188,1810]" captionTargetId="figure-15@3.[151,1436,181,1825]" captionTargetPageId="3" captionText="FIGURES 10–19. Male palp (10–13) and bulb (14–19) of Acantholycosa sergeevi sp. n. (10–11), A. sayanensis (14–15), A. sidorovi sp. n. (16–17), A. levinae (18-19) and Mongolicosa lizae sp. n. (12–13). 10, 12—retrolateral; 11, 13, 14, 16, 18— ventral; 15, 17, 19—anterior. Scale bars=0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300796" httpUri="https://zenodo.org/record/5300796/files/figure.png" pageId="5" pageNumber="572">Figs 16–17</figureCitation>
, 24–27, 34–35. Tegular apophysis almost square, with a poorly developed basal arm (
<emphasis id="68FD89349B17A40EBA178AE5FA13F81D" box="[1389,1420,1944,1969]" italics="true" pageId="5" pageNumber="572">Ba</emphasis>
). Palea (
<emphasis id="68FD89349B17A40EBF9C8AC7FF60F878" box="[230,255,1978,2004]" italics="true" pageId="5" pageNumber="572">Pl</emphasis>
) rounded. Paleal outgrowth strongly bent. Terminal apophysis (
<emphasis id="68FD89349B17A40EBCA08AC6FC6AF878" box="[986,1013,1979,2004]" italics="true" pageId="5" pageNumber="572">Ta</emphasis>
) straight, spine-like. Embolus with a semicircular notch in the distal part.
</paragraph>
<caption id="0EF605AE9B14A40DBFED8BB3FDBBF8DB" ID-DOI="http://doi.org/10.5281/zenodo.5300798" ID-Zenodo-Dep="5300798" httpUri="https://zenodo.org/record/5300798/files/figure.png" pageId="6" pageNumber="573" startId="6.[151,264,1742,1766]" targetBox="[154,1435,189,1719]" targetPageId="6">
<paragraph id="5A3655269B14A40DBFED8BB3FDBBF8DB" blockId="6.[151,1437,1742,1911]" pageId="6" pageNumber="573">
<emphasis id="68FD89349B14A40DBFED8BB3FECBF94B" bold="true" box="[151,340,1742,1767]" pageId="6" pageNumber="573">FIGURES 32–43.</emphasis>
Tegulum and subtegulum (32–37), bulb (38–40) and embolic division (41–43) of
<taxonomicName id="9D892EA59B14A40DBBE98BB2FA03F94A" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[1171,1436,1743,1767]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="573" phylum="Arthropoda" rank="species" species="sayanensis">
<emphasis id="68FD89349B14A40DBBE98BB2FA03F94A" box="[1171,1436,1743,1767]" italics="true" pageId="6" pageNumber="573">Acantholycosa sayanensis</emphasis>
</taxonomicName>
(32–33),
<taxonomicName id="9D892EA59B14A40DBF8F8B8EFEFCF8A7" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[245,355,1779,1803]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="573" phylum="Arthropoda" rank="species" species="sidorovi" status="sp. nov.">
<emphasis id="68FD89349B14A40DBF8F8B8EFEFCF8A7" box="[245,355,1779,1803]" italics="true" pageId="6" pageNumber="573">A. sidorovi</emphasis>
</taxonomicName>
<emphasis id="68FD89349B14A40DBE118B8EFE3BF8A7" bold="true" box="[363,420,1779,1803]" pageId="6" pageNumber="573">
<taxonomicNameLabel id="73CE344F9B14A40DBE118B8EFE3BF8A7" box="[363,420,1779,1803]" pageId="6" pageNumber="573" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(34–35),
<taxonomicName id="9D892EA59B14A40DBD738B8EFDF1F8A7" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[521,622,1779,1803]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="573" phylum="Arthropoda" rank="species" species="levinae">
<emphasis id="68FD89349B14A40DBD738B8EFDF1F8A7" box="[521,622,1779,1803]" italics="true" pageId="6" pageNumber="573">A. levinae</emphasis>
</taxonomicName>
(36–37) and
<taxonomicName id="9D892EA59B14A40DBD828B8EFCF7F8A6" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[760,872,1779,1802]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="573" phylum="Arthropoda" rank="species" species="sergeevi" status="sp. nov.">
<emphasis id="68FD89349B14A40DBD828B8EFCF7F8A6" box="[760,872,1779,1802]" italics="true" pageId="6" pageNumber="573">A. sergeevi</emphasis>
</taxonomicName>
<emphasis id="68FD89349B14A40DBC0A8B8EFC36F8A7" bold="true" box="[880,937,1779,1803]" pageId="6" pageNumber="573">
<taxonomicNameLabel id="73CE344F9B14A40DBC0A8B8EFC36F8A7" box="[880,937,1779,1803]" pageId="6" pageNumber="573" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(38–43). 32, 34, 36, 38, 41—ventral; 33, 35, 37, 39, 42—anterior; 40, 43—retrolateral. Scale bars=0.2 mm. Abbreviations:
<emphasis id="68FD89349B14A40DBCE58A6AFC24F882" box="[927,955,1815,1838]" italics="true" pageId="6" pageNumber="573">Aa</emphasis>
apical arm of the tegular apophysis,
<emphasis id="68FD89349B14A40DBA398A6AFAC0F882" box="[1347,1375,1815,1838]" italics="true" pageId="6" pageNumber="573">Ba</emphasis>
basal arm of the tegular apophysis,
<emphasis id="68FD89349B14A40DBEBF8A46FE78F8FE" box="[453,487,1851,1874]" italics="true" pageId="6" pageNumber="573">Em</emphasis>
embolus,
<emphasis id="68FD89349B14A40DBD2A8A46FDF4F8FE" box="[592,619,1851,1874]" italics="true" pageId="6" pageNumber="573">Pe</emphasis>
extension of the palea,
<emphasis id="68FD89349B14A40DBC268A46FCECF8FF" box="[860,883,1851,1875]" italics="true" pageId="6" pageNumber="573">Pl</emphasis>
palea,
<emphasis id="68FD89349B14A40DBCC18A46FC4CF8FE" box="[955,979,1851,1874]" italics="true" pageId="6" pageNumber="573">Ta</emphasis>
terminal apophysis,
<emphasis id="68FD89349B14A40DBBDD8A46FB5DF8FE" box="[1191,1218,1851,1874]" italics="true" pageId="6" pageNumber="573">Tg</emphasis>
tegular apophysis,
<emphasis id="68FD89349B14A40DBAFD8A46FA02F8FE" box="[1415,1437,1851,1874]" italics="true" pageId="6" pageNumber="573">Ts</emphasis>
triangle-shaped projection of the palea.
</paragraph>
</caption>
<caption id="0EF605AE9B15A40CBFED8A7FFC7CF82A" ID-DOI="http://doi.org/10.5281/zenodo.5300800" ID-Zenodo-Dep="5300800" httpUri="https://zenodo.org/record/5300800/files/figure.png" pageId="7" pageNumber="574" startId="7.[151,264,1794,1818]" targetBox="[163,1427,188,1770]" targetPageId="7">
<paragraph id="5A3655269B15A40CBFED8A7FFC7CF82A" blockId="7.[151,1436,1794,1926]" pageId="7" pageNumber="574">
<emphasis id="68FD89349B15A40CBFED8A7FFEC8F8B6" bold="true" box="[151,343,1794,1818]" pageId="7" pageNumber="574">FIGURES 44–53.</emphasis>
Tegulum and subtegulum (44–45, 52–53), bulb (46–48) and embolic division (49–51) of
<taxonomicName id="9D892EA59B15A40CBA7D8A7FFF75F892" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="574" phylum="Arthropoda" rank="species" species="sergeevi" status="sp. nov.">
<emphasis id="68FD89349B15A40CBA7D8A7FFF75F892" italics="true" pageId="7" pageNumber="574">Acantholycosa sergeevi</emphasis>
</taxonomicName>
<emphasis id="68FD89349B15A40CBF8E8A5BFEAFF892" bold="true" box="[244,304,1830,1854]" pageId="7" pageNumber="574">
<taxonomicNameLabel id="73CE344F9B15A40CBF8E8A5BFEAFF892" box="[244,304,1830,1854]" pageId="7" pageNumber="574" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(44–45) and
<taxonomicName id="9D892EA59B15A40CBEBB8A5BFDE0F892" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[449,639,1830,1854]" class="Arachnida" family="Lycosidae" genus="Mongolicosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="574" phylum="Arthropoda" rank="species" species="lizae" status="sp. nov.">
<emphasis id="68FD89349B15A40CBEBB8A5BFDE0F892" box="[449,639,1830,1854]" italics="true" pageId="7" pageNumber="574">Mongolicosa lizae</emphasis>
</taxonomicName>
<emphasis id="68FD89349B15A40CBDF38A5BFD5AF892" bold="true" box="[649,709,1830,1854]" pageId="7" pageNumber="574">
<taxonomicNameLabel id="73CE344F9B15A40CBDF38A5BFD5AF892" box="[649,709,1830,1854]" pageId="7" pageNumber="574" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(46–53). 44, 46, 49, 52—ventral; 45, 47, 50, 53—anterior; 48, 51— retrolateral. Scale bars=0.2 mm. Abbreviations:
<emphasis id="68FD89349B15A40CBDFA8A37FD03F8CD" box="[640,668,1866,1889]" italics="true" pageId="7" pageNumber="574">Aa</emphasis>
apical arm of the tegular apophysis,
<emphasis id="68FD89349B15A40CBB6C8A36FBADF8CE" box="[1046,1074,1867,1890]" italics="true" pageId="7" pageNumber="574">Ba</emphasis>
basal arm of the tegular apophysis,
<emphasis id="68FD89349B15A40CBFED8A12FF26F82A" box="[151,185,1903,1926]" italics="true" pageId="7" pageNumber="574">Em</emphasis>
embolus,
<emphasis id="68FD89349B15A40CBE588A13FEA6F82A" box="[290,313,1902,1926]" italics="true" pageId="7" pageNumber="574">Pl</emphasis>
palea,
<emphasis id="68FD89349B15A40CBEFB8A13FE05F82A" box="[385,410,1902,1926]" italics="true" pageId="7" pageNumber="574">Sd</emphasis>
sperm duct,
<emphasis id="68FD89349B15A40CBD678A12FDAAF82A" box="[541,565,1903,1926]" italics="true" pageId="7" pageNumber="574">Ta</emphasis>
terminal apophysis,
<emphasis id="68FD89349B15A40CBC728A12FCBCF82A" box="[776,803,1903,1926]" italics="true" pageId="7" pageNumber="574">Tg</emphasis>
tegular apophysis.
</paragraph>
</caption>
<caption id="0EF605AE9B1AA403BFED8B57FE37F902" ID-DOI="http://doi.org/10.5281/zenodo.5300802" ID-Zenodo-Dep="5300802" httpUri="https://zenodo.org/record/5300802/files/figure.png" pageId="8" pageNumber="575" startId="8.[151,264,1578,1602]" targetBox="[151,1436,181,1554]" targetPageId="8">
<paragraph id="5A3655269B1AA403BFED8B57FE37F902" blockId="8.[151,1437,1578,1711]" pageId="8" pageNumber="575">
<emphasis id="68FD89349B1AA403BFED8B57FEC8F9EF" bold="true" box="[151,343,1578,1603]" pageId="8" pageNumber="575">FIGURES 54–62.</emphasis>
Epigyne and female internal genitalia of
<taxonomicName id="9D892EA59B1AA403BC758B56FB84F9EE" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[783,1051,1579,1603]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="575" phylum="Arthropoda" rank="species" species="sayanensis">
<emphasis id="68FD89349B1AA403BC758B56FB84F9EE" box="[783,1051,1579,1603]" italics="true" pageId="8" pageNumber="575">Acantholycosa sayanensis</emphasis>
</taxonomicName>
(54–56),
<taxonomicName id="9D892EA59B1AA403BBFE8B56FB69F9EE" authorityName="Marusik, Hippa &amp; Koponen" authorityYear="1996" box="[1156,1270,1579,1602]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="575" phylum="Arthropoda" rank="species" species="azyuzini">
<emphasis id="68FD89349B1AA403BBFE8B56FB69F9EE" box="[1156,1270,1579,1602]" italics="true" pageId="8" pageNumber="575">A. azyuzini</emphasis>
</taxonomicName>
(57–59) and
<taxonomicName id="9D892EA59B1AA403BAFC8B56FF76F9CB" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="575" phylum="Arthropoda" rank="species" species="sidorovi" status="sp. nov.">
<emphasis id="68FD89349B1AA403BAFC8B56FF76F9CB" italics="true" pageId="8" pageNumber="575">A. sidorovi</emphasis>
</taxonomicName>
<emphasis id="68FD89349B1AA403BF898B32FEAFF9CB" bold="true" box="[243,304,1615,1639]" pageId="8" pageNumber="575">
<taxonomicNameLabel id="73CE344F9B1AA403BF898B32FEAFF9CB" box="[243,304,1615,1639]" pageId="8" pageNumber="575" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(60–62). 54, 57, 60—intact, ventral; 55, 58, 61—macerated, ventral; 56, 59, 62—macerated, dorsal. Scale bars=0.2 mm. Abbreviations:
<emphasis id="68FD89349B1AA403BEBF8B0EFE7EF926" box="[453,481,1651,1674]" italics="true" pageId="8" pageNumber="575">Ap</emphasis>
apical pocket,
<emphasis id="68FD89349B1AA403BD038B0EFD08F927" box="[633,663,1651,1675]" italics="true" pageId="8" pageNumber="575">Cd</emphasis>
copulatory ducts,
<emphasis id="68FD89349B1AA403BC288B0EFCF1F926" box="[850,878,1651,1674]" italics="true" pageId="8" pageNumber="575">Fo</emphasis>
epigynal fovea,
<emphasis id="68FD89349B1AA403BB6C8B0EFBB4F926" box="[1046,1067,1651,1674]" italics="true" pageId="8" pageNumber="575">Li</emphasis>
lip of the epigyne,
<emphasis id="68FD89349B1AA403BB958B0EFA95F926" box="[1263,1290,1651,1674]" italics="true" pageId="8" pageNumber="575">Re</emphasis>
receptacle,
<emphasis id="68FD89349B1AA403BAF98B0EFA03F927" box="[1411,1436,1651,1675]" italics="true" pageId="8" pageNumber="575">Sb</emphasis>
septal base,
<emphasis id="68FD89349B1AA403BE688BEAFEB6F902" box="[274,297,1687,1710]" italics="true" pageId="8" pageNumber="575">Ss</emphasis>
septal stem.
</paragraph>
</caption>
<paragraph id="5A3655269B1AA403BFBD8A7CFCACF87C" blockId="8.[151,1437,1793,2035]" pageId="8" pageNumber="575">Female. Total length 8.7. Carapace: 3.4 long, 2.75 wide. Coloration. Carapace dark brown. Eye field almost black. Chelicerae, sternum and labium brown. Endites and coxae yellow-gray. Legs and palps yellow-gray with brown annulation. Abdomen brown-gray dorsally. Cardiac mark dark brown. Venter of the abdomen yellow. Spinnerets yellow-gray. Spination of leg I: Fe d1-1-1 p0-0-2 r0-0-1; Ti p1-0-0 r1-0-0 v2-2-2-2-1-2; Mt p0-1-0 r0- 1-0 v2-2-0. Leg measurements: I 3.15, 1.45, 2.8, 2.75, 1.35 (11.5); II 3.15, 1.4, 2.65, 2.85, 1.3 (11.35); III 2.9, 1.25, 2.45, 3.45, 1.4 (11.45); IV 3.75, 1.4, 3.25, 5.15, 1.95 (15.5).</paragraph>
<paragraph id="5A3655269B1AA402BFBD8AA4FB18FF79" blockId="8.[151,1437,1793,2035]" lastBlockId="9.[151,1436,151,249]" lastPageId="9" lastPageNumber="576" pageId="8" pageNumber="575">
Epigyne and internal genitalia as in
<figureCitation id="C2B249A39B1AA403BD258AA4FD7EF85F" box="[607,737,2009,2035]" captionStart="FIGURES 54–62" captionStartId="8.[151,264,1578,1602]" captionTargetBox="[151,1436,181,1554]" captionTargetId="figure-33@8.[151,1436,181,1554]" captionTargetPageId="8" captionText="FIGURES 54–62. Epigyne and female internal genitalia of Acantholycosa sayanensis (54–56), A. azyuzini (57–59) and A. sidorovi sp. n. (60–62). 54, 57, 60—intact, ventral; 55, 58, 61—macerated, ventral; 56, 59, 62—macerated, dorsal. Scale bars=0.2 mm. Abbreviations: Ap apical pocket, Cd copulatory ducts, Fo epigynal fovea, Li lip of the epigyne, Re receptacle, Sb septal base, Ss septal stem." figureDoi="http://doi.org/10.5281/zenodo.5300802" httpUri="https://zenodo.org/record/5300802/files/figure.png" pageId="8" pageNumber="575">Figs 60–62</figureCitation>
. Fovea slightly longer than wide. Septal base smooth, 2 times longer than wide. Septal stem undeveloped. Lips of the epigyne (
<emphasis id="68FD89349B1BA402BC1C8DEAFCE2FF1C" box="[870,893,151,176]" italics="true" pageId="9" pageNumber="576">Li</emphasis>
) not touching each other. Apical pocket with one hood. Receptacles fusiform. Copulatory ducts (
<emphasis id="68FD89349B1BA402BDD38DC6FD55FF79" box="[681,714,187,213]" italics="true" pageId="9" pageNumber="576">Cd</emphasis>
) almost straight, as long as receptacles.
</paragraph>
</subSubSection>
<subSubSection id="129306AD9B1BA402BFBD8DA3FB13FF55" box="[199,1164,222,249]" pageId="9" pageNumber="576" type="distribution">
<paragraph id="5A3655269B1BA402BFBD8DA3FB13FF55" blockId="9.[151,1436,151,249]" box="[199,1164,222,249]" pageId="9" pageNumber="576">
<emphasis id="68FD89349B1BA402BFBD8DA3FEFEFF54" bold="true" box="[199,353,222,248]" pageId="9" pageNumber="576">Distribution.</emphasis>
Known only from the
<typeStatus id="8532EB849B1BA402BD1A8DA2FD0FFF55" box="[608,656,223,249]" pageId="9" pageNumber="576">type</typeStatus>
locality in the
<collectingRegion id="984D9BC49B1BA402BC4D8DA2FCF0FF55" box="[823,879,223,249]" country="Russia" name="Altay" pageId="9" pageNumber="576">Altai</collectingRegion>
Mountains (
<figureCitation id="C2B249A39B1BA402BC878DA2FB1FFF55" box="[1021,1152,222,249]" captionStart="FIGURES 72–77" captionStartId="13.[151,264,1676,1700]" captionTargetBox="[151,1436,309,1652]" captionTargetId="figure-76@13.[151,1436,309,1652]" captionTargetPageId="13" captionText="FIGURES 72–77. Male of Acantholycosa sayanensis from Borus Mt. Range (72), female of Acantholycosa sp. (A. azyuzini or A. levinae) with egg sac, from Katunskyi Mt. Range (73), habitat of A. sayanensis and A. sergeevi sp. n. in Borus Mt. Range (74), collecting localities of studied species from the Acantholycosa-complex (75–76) and map showing the main mountain systems of South Siberia and adjacent regions (77). Diamond—A. sayanensis, square—A. sidorovi sp. n., triangle—A. kulikovi sp. n., star—A. levinae, circle—A. sergeevi sp. n., inverted triangle—Mongolicosa lizae sp. n. The frame of Fig. 76 refers to the content of Fig. 75." figureDoi="http://doi.org/10.5281/zenodo.5300806" httpUri="https://zenodo.org/record/5300806/files/figure.png" pageId="9" pageNumber="576">Figs 75–76</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="8575B447B1469C8A382EE694195A575C">New data on the wolf spiders from theAcantholycosa-complex (Araneae: Lycosidae) from the South Siberia</mods:title>
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<emphasis id="68FD89349B18A401BFED8FACFE39FD40" bold="true" box="[151,422,721,748]" italics="true" pageId="10" pageNumber="577">Acantholycosa levinae</emphasis>
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<figureCitation id="C2B249A39B18A401BFED8F8BFF64FCBD" box="[151,251,758,785]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1832,1856]" captionTargetBox="[157,1436,189,1799]" captionTargetId="figure-33@2.[151,1436,181,1809]" captionTargetPageId="2" captionText="FIGURES 1–9. General appearance of Acantholycosa sayanensis (1–2), A. kulikovi sp. n. (3), A. levinae (4–5), A. sergeevi sp. n. (6), Mongolicosa lizae sp. n. (8–9) and leg I of A. sergeevi sp. n. (7). 1, 4, 6, 8—male, dorsal; 2, 3, 5, 9—female, dorsal; 7—leg I of male, prolateral. Scale bars=2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300794" httpUri="https://zenodo.org/record/5300794/files/figure.png" pageId="10" pageNumber="577">Figs 4–5</figureCitation>
,
<figureCitation id="C2B249A39B18A401BE7C8F8BFED0FCBC" box="[262,335,758,784]" captionStart="FIGURES 10–19" captionStartId="3.[151,264,1849,1873]" captionTargetBox="[155,1425,188,1810]" captionTargetId="figure-15@3.[151,1436,181,1825]" captionTargetPageId="3" captionText="FIGURES 10–19. Male palp (10–13) and bulb (14–19) of Acantholycosa sergeevi sp. n. (10–11), A. sayanensis (14–15), A. sidorovi sp. n. (16–17), A. levinae (18-19) and Mongolicosa lizae sp. n. (12–13). 10, 12—retrolateral; 11, 13, 14, 16, 18— ventral; 15, 17, 19—anterior. Scale bars=0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300796" httpUri="https://zenodo.org/record/5300796/files/figure.png" pageId="10" pageNumber="577">18–19</figureCitation>
, 28–31, 36–37, 66–68, 75–76
</paragraph>
</subSubSection>
<subSubSection id="129306AD9B18A401BFED8E46FD5FFCDD" pageId="10" pageNumber="577" type="reference_group">
<paragraph id="5A3655269B18A401BFED8E46FCDFFCFE" blockId="10.[151,832,826,881]" box="[151,832,826,850]" pageId="10" pageNumber="577">
<treatmentCitationGroup id="7A9972089B18A401BFED8E46FCDFFCFE" box="[151,832,826,850]" pageId="10" pageNumber="577">
<treatmentCitation id="DB2873379B18A401BFED8E46FD2FFCFE" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." box="[151,688,826,850]" page="112" pageId="10" pageNumber="577" year="2004">
<taxonomicName id="9D892EA59B18A401BFED8E46FD2FFCFE" ID-CoL="64GDM" authority="Marusik, Azarkina &amp; Koponen, 2004: 112" authorityName="Marusik, Azarkina &amp; Koponen" authorityPageNumber="112" authorityYear="2004" box="[151,688,826,850]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="levinae">
<emphasis id="68FD89349B18A401BFED8E46FF63FCFE" box="[151,252,826,850]" italics="true" pageId="10" pageNumber="577">A. levinae</emphasis>
<bibRefCitation id="3E1828D79B18A401BE798E47FD2FFCFE" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." box="[259,688,826,850]" pageId="10" pageNumber="577" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">Marusik, Azarkina &amp; Koponen, 2004: 112</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, f. 61–65 (
<specimenCount id="4C8F9EAF9B18A401BC648E47FCACFCFE" box="[798,819,826,850]" pageId="10" pageNumber="577" type="male">♂</specimenCount>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="5A3655269B18A401BFED8E24FD5FFCDD" blockId="10.[151,832,826,881]" box="[151,704,857,881]" pageId="10" pageNumber="577">
<treatmentCitationGroup id="7A9972089B18A401BFED8E24FD5FFCDD" box="[151,704,857,881]" pageId="10" pageNumber="577">
<taxonomicName id="9D892EA59B18A401BFED8E24FF63FCDD" ID-CoL="64GDM" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[151,252,857,881]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="levinae">
<emphasis id="68FD89349B18A401BFED8E24FF63FCDD" box="[151,252,857,881]" italics="true" pageId="10" pageNumber="577">A. levinae</emphasis>
</taxonomicName>
—
<treatmentCitation id="DB2873379B18A401BE668E24FDC8FCDD" author="Marusik, Y. M. &amp; Omelko, M. M." box="[284,599,857,881]" page="597" pageId="10" pageNumber="577" year="2017">
<bibRefCitation id="3E1828D79B18A401BE668E24FDC8FCDD" author="Marusik, Y. M. &amp; Omelko, M. M." box="[284,599,857,881]" pageId="10" pageNumber="577" pagination="597 - 600" refId="ref10588" refString="Marusik, Y. M. &amp; Omelko, M. M. (2017) A new species of Acantholycosa baltoroi - group (Araneae: Lycosidae, Pardosinae) from the Russian Far East. Zootaxa, 4232 (4), 597 - 600. https: // doi. org / 10.11646 / zootaxa. 4232.4.12" type="journal article" year="2017">Marusik &amp; Omelko, 2017: 597</bibRefCitation>
</treatmentCitation>
, f. 11 (
<specimenCount id="4C8F9EAF9B18A401BDE48E24FD2CFCDD" box="[670,691,857,881]" pageId="10" pageNumber="577" type="male">♂</specimenCount>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="129306AD9B18A401BFED8EE2FE53FBC1" pageId="10" pageNumber="577" type="materials_examined">
<paragraph id="5A3655269B18A401BFED8EE2FE53FBC1" blockId="10.[151,1437,927,2034]" pageId="10" pageNumber="577">
<emphasis id="68FD89349B18A401BFED8EE2FE1CFC15" bold="true" box="[151,387,927,953]" pageId="10" pageNumber="577">Material examined.</emphasis>
<materialsCitation id="EAE15F7B9B18A401BEF68EE2FC7AFC16" box="[396,997,927,954]" collectionCode="ISEA" country="Russia" location="Altai Republic" pageId="10" pageNumber="577" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Altay">
<collectingCountry id="229E15B69B18A401BEF68EE2FE6BFC15" box="[396,500,927,953]" name="Russia" pageId="10" pageNumber="577">RUSSIA</collectingCountry>
:
<emphasis id="68FD89349B18A401BD7A8EE2FD30FC15" bold="true" box="[512,687,927,953]" pageId="10" pageNumber="577">
<collectingRegion id="984D9BC49B18A401BD7A8EE2FD30FC15" box="[512,687,927,953]" country="Russia" name="Altay" pageId="10" pageNumber="577">Altai Republic</collectingRegion>
</emphasis>
:
<specimenCount id="4C8F9EAF9B18A401BDBA8EE2FD7CFC15" box="[704,739,927,953]" pageId="10" pageNumber="577" type="male">1♂</specimenCount>
<specimenCount id="4C8F9EAF9B18A401BD968EE2FC90FC15" box="[748,783,927,953]" pageId="10" pageNumber="577" type="female">1♀</specimenCount>
(
<collectionCode id="3C98CDE39B18A401BC5A8EE2FCFDFC15" box="[800,866,927,953]" pageId="10" pageNumber="577">ISEA</collectionCode>
, 001.8658)
</materialsCitation>
,
<materialsCitation id="EAE15F7B9B18A401BC888EE2FE4EFBAD" collectingDate="2019-07-27" collectionCode="ISEA" collectorName="A. A. Fomichev &amp; Yu. V. Dyachkov" country="Russia" elevation="1705" latitude="49.983334" location="Srednee Mul'tinskoe Lake" longLatPrecision="1100" longitude="85.833336" municipality="Katunskyi Mt. Range" pageId="10" pageNumber="577" specimenCount="1" specimenCount-female="1" stateProvince="Altay">
<specimenCount id="4C8F9EAF9B18A401BC888EE2FB8AFC15" box="[1010,1045,927,953]" pageId="10" pageNumber="577" type="female">1♀</specimenCount>
(
<collectionCode id="3C98CDE39B18A401BB5C8EE2FBF7FC15" box="[1062,1128,927,953]" pageId="10" pageNumber="577">ISEA</collectionCode>
, 001.8722),
<collectingMunicipality id="BA52CF5C9B18A401BB828EE2FF7EFC72" pageId="10" pageNumber="577">Katunskyi Mt. Range</collectingMunicipality>
,
<location id="5F5603FD9B18A401BF918EBEFD8BFC71" LSID="urn:lsid:plazi:treatment:D220E4309B18A400BFED8FACFA1BFE7D:5F5603FD9B18A401BF918EBEFD8BFC71" box="[235,532,963,989]" country="Russia" latitude="49.983334" longLatPrecision="1100" longitude="85.833336" municipality="Katunskyi Mt. Range" name="Srednee Mul'tinskoe Lake" pageId="10" pageNumber="577" stateProvince="Altay">Srednee Mul’tinskoe Lake</location>
,
<geoCoordinate id="3FBD33E19B18A401BD648EBEFD1EFC72" box="[542,641,963,990]" degrees="49" direction="north" minutes="59" orientation="latitude" pageId="10" pageNumber="577" precision="925" value="49.983334">49°59’N</geoCoordinate>
,
<geoCoordinate id="3FBD33E19B18A401BDF18EBEFD73FC72" box="[651,748,963,990]" degrees="85" direction="east" minutes="50" orientation="longitude" pageId="10" pageNumber="577" precision="925" value="85.833336">85°50’E</geoCoordinate>
, scree and lichen pillows,
<quantity id="9D71F8C39B18A401BB768EBEFB37FC72" box="[1036,1192,963,990]" metricMagnitude="3" metricUnit="m" metricValue="1.705" metricValueMax="1.76" metricValueMin="1.65" pageId="10" pageNumber="577" unit="m" value="1705.0" valueMax="1760.0" valueMin="1650.0">
<elevation id="D1A4B2159B18A401BB768EBEFB37FC72" box="[1036,1192,963,990]" metricMagnitude="3" metricUnit="m" metricValue="1.705" metricValueMax="1.76" metricValueMin="1.65" pageId="10" pageNumber="577" unit="m" value="1705.0" valueMax="1760.0" valueMin="1650.0">1650–1760 m</elevation>
</quantity>
, 23–
<date id="2E3773E69B18A401BBA68EBEFAC1FC71" box="[1244,1374,963,989]" pageId="10" pageNumber="577" value="2019-07-27">
<collectingDate id="3E738A0E9B18A401BBA68EBEFAC1FC71" box="[1244,1374,963,989]" pageId="10" pageNumber="577" value="2019-07-27">27.07.2019</collectingDate>
</date>
,
<collectorName id="F77C30F09B18A401BA1C8EBEFE97FBAD" pageId="10" pageNumber="577">A.A. Fomichev</collectorName>
,
<collectorName id="F77C30F09B18A401BE6E8E95FE4EFBAD" box="[276,465,999,1026]" pageId="10" pageNumber="577">Yu.V. Dyachkov</collectorName>
</materialsCitation>
;
<materialsCitation id="EAE15F7B9B18A401BEA78E9AFAF0FB8A" collectingDate="2019-07-26" collectionCode="ISEA" collectorName="A. A. Fomichev &amp; Yu. V. Dyachkov, E. A &amp; Nepaeva" country="Russia" county="Katunskyi Mt. Range" elevation="2400" latitude="49.98365" location="Kuiguk River" longLatPrecision="1" longitude="85.8516" municipality="Mul'ta River" pageId="10" pageNumber="577" specimenCount="2" specimenCount-female="2" stateProvince="Altay">
<specimenCount id="4C8F9EAF9B18A401BEA78E9AFD9FFBAD" box="[477,512,999,1025]" pageId="10" pageNumber="577" type="female">2♀</specimenCount>
(
<collectionCode id="3C98CDE39B18A401BD6A8E9AFDCDFBAD" box="[528,594,999,1025]" pageId="10" pageNumber="577">ISEA</collectionCode>
, 001.8659),
<collectingCounty id="B3572DAA9B18A401BD9B8E9AFC49FBAE" box="[737,982,999,1026]" pageId="10" pageNumber="577">Katunskyi Mt. Range</collectingCounty>
, watershed of
<collectingMunicipality id="BA52CF5C9B18A401BB078E9AFA90FBAD" box="[1149,1295,999,1025]" pageId="10" pageNumber="577">Mul’ta River</collectingMunicipality>
and
<location id="5F5603FD9B18A401BA328E9AFF48FB89" LSID="urn:lsid:plazi:treatment:D220E4309B18A400BFED8FACFA1BFE7D:5F5603FD9B18A401BA328E9AFF48FB89" country="Russia" county="Katunskyi Mt. Range" latitude="49.98365" longLatPrecision="1" longitude="85.8516" municipality="Mul'ta River" name="Kuiguk River" pageId="10" pageNumber="577" stateProvince="Altay">Kuiguk River</location>
,
<geoCoordinate id="3FBD33E19B18A401BF988976FEE8FB8A" box="[226,375,1035,1062]" degrees="49" direction="north" minutes="59.019" orientation="latitude" pageId="10" pageNumber="577" precision="1" value="49.98365">49°59.019’N</geoCoordinate>
,
<geoCoordinate id="3FBD33E19B18A401BEF88976FD8CFB8A" box="[386,531,1035,1062]" degrees="85" direction="east" minutes="51.096" orientation="longitude" pageId="10" pageNumber="577" precision="1" value="85.8516">85°51.096’E</geoCoordinate>
, scree,
<quantity id="9D71F8C39B18A401BD1C8976FD23FB8A" box="[614,700,1035,1062]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="577" unit="m" value="2400.0">
<elevation id="D1A4B2159B18A401BD1C8976FD23FB8A" box="[614,700,1035,1062]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="577" unit="m" value="2400.0">2400 m</elevation>
</quantity>
,
<date id="2E3773E69B18A401BDB28976FCD6FB89" box="[712,841,1035,1061]" pageId="10" pageNumber="577" value="2019-07-26">
<collectingDate id="3E738A0E9B18A401BDB28976FCD6FB89" box="[712,841,1035,1061]" pageId="10" pageNumber="577" value="2019-07-26">26.07.2019</collectingDate>
</date>
,
<collectorName id="F77C30F09B18A401BC288976FB9EFB89" box="[850,1025,1035,1061]" pageId="10" pageNumber="577">A.A. Fomichev</collectorName>
,
<collectorName id="F77C30F09B18A401BB718971FB61FB89" box="[1035,1278,1035,1062]" pageId="10" pageNumber="577">Yu.V. Dyachkov, E.A</collectorName>
.
<collectorName id="F77C30F09B18A401BA738971FAF0FB8A" box="[1289,1391,1036,1062]" pageId="10" pageNumber="577">Nepaeva</collectorName>
</materialsCitation>
;
<materialsCitation id="EAE15F7B9B18A401BA038976FEB1FBE5" collectionCode="ZMMU" country="Russia" pageId="10" pageNumber="577" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Altay">
<specimenCount id="4C8F9EAF9B18A401BA038976FA02FB89" box="[1401,1437,1035,1061]" pageId="10" pageNumber="577" type="male">3♂</specimenCount>
<specimenCount id="4C8F9EAF9B18A401BFED8952FF25FBE5" box="[151,186,1071,1097]" pageId="10" pageNumber="577" type="female">1♀</specimenCount>
(
<collectionCode id="3C98CDE39B18A401BFB38952FEB9FBE5" box="[201,294,1071,1097]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="10" pageNumber="577" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="EAE15F7B9B18A401BE438952FE57FBC1" collectingDate="2018-07-21" collectorName="A. E. Naidenov &amp; K. E. Naidenova" country="Russia" elevation="1942" latitude="49.916668" location="Poperechnoe Lake" longLatPrecision="1100" longitude="85.88333" municipality="Katunskyi Mt. Range" pageId="10" pageNumber="577" specimenCount="1" stateProvince="Altay">
<collectingMunicipality id="BA52CF5C9B18A401BE438952FDB4FBE6" box="[313,555,1071,1098]" pageId="10" pageNumber="577">Katunskyi Mt. Range</collectingMunicipality>
,
<location id="5F5603FD9B18A401BD4D8952FC96FBE5" LSID="urn:lsid:plazi:treatment:D220E4309B18A400BFED8FACFA1BFE7D:5F5603FD9B18A401BD4D8952FC96FBE5" box="[567,777,1071,1097]" country="Russia" latitude="49.916668" longLatPrecision="1100" longitude="85.88333" municipality="Katunskyi Mt. Range" name="Poperechnoe Lake" pageId="10" pageNumber="577" stateProvince="Altay">Poperechnoe Lake</location>
,
<geoCoordinate id="3FBD33E19B18A401BC6E8952FCE8FBE6" box="[788,887,1071,1098]" degrees="49" direction="north" minutes="55" orientation="latitude" pageId="10" pageNumber="577" precision="925" value="49.916668">49°55’N</geoCoordinate>
,
<geoCoordinate id="3FBD33E19B18A401BCF98952FC7CFBE6" box="[899,995,1071,1098]" degrees="85" direction="east" minutes="53" orientation="longitude" pageId="10" pageNumber="577" precision="925" value="85.88333">85°53’E</geoCoordinate>
, scree,
<quantity id="9D71F8C39B18A401BB4C8952FB4CFBE6" box="[1078,1235,1071,1098]" metricMagnitude="3" metricUnit="m" metricValue="1.9425" metricValueMax="2.0" metricValueMin="1.885" pageId="10" pageNumber="577" unit="m" value="1942.5" valueMax="2000.0" valueMin="1885.0">
<elevation id="D1A4B2159B18A401BB4C8952FB4CFBE6" box="[1078,1235,1071,1098]" metricMagnitude="3" metricUnit="m" metricValue="1.9425" metricValueMax="2.0" metricValueMin="1.885" pageId="10" pageNumber="577" unit="m" value="1942.5" valueMax="2000.0" valueMin="1885.0">1885–2000 m</elevation>
</quantity>
,
<date id="2E3773E69B18A401BBA58952FAC0FBE5" box="[1247,1375,1071,1097]" pageId="10" pageNumber="577" value="2018-07-21">
<collectingDate id="3E738A0E9B18A401BBA58952FAC0FBE5" box="[1247,1375,1071,1097]" pageId="10" pageNumber="577" value="2018-07-21">21.07.2018</collectingDate>
</date>
,
<collectorName id="F77C30F09B18A401BA138952FE9AFBC1" pageId="10" pageNumber="577">A.E. Naidenov</collectorName>
,
<collectorName id="F77C30F09B18A401BE6A8929FE57FBC1" box="[272,456,1107,1134]" pageId="10" pageNumber="577">K.E. Naidenova</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="129306AD9B18A401BFBD890AFADEF9C9" pageId="10" pageNumber="577" type="diagnosis">
<paragraph id="5A3655269B18A401BFBD890AFADEF9C9" blockId="10.[151,1437,927,2034]" pageId="10" pageNumber="577">
<emphasis id="68FD89349B18A401BFBD890AFEDDFB3D" bold="true" box="[199,322,1143,1169]" pageId="10" pageNumber="577">Diagnosis.</emphasis>
<taxonomicName id="9D892EA59B18A401BE32890AFDDBFB3D" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[328,580,1143,1169]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="levinae">
<emphasis id="68FD89349B18A401BE32890AFDDBFB3D" box="[328,580,1143,1169]" italics="true" pageId="10" pageNumber="577">Acantholycosa levinae</emphasis>
</taxonomicName>
resembles
<taxonomicName id="9D892EA59B18A401BDBB890AFCA7FB3D" baseAuthorityName="Caporiacco" baseAuthorityYear="1935" box="[705,824,1143,1169]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="baltoroi">
<emphasis id="68FD89349B18A401BDBB890AFCA7FB3D" box="[705,824,1143,1169]" italics="true" pageId="10" pageNumber="577">A. baltoroi</emphasis>
</taxonomicName>
. Both species have protruding palea (
<emphasis id="68FD89349B18A401BBA7890AFB69FB3D" box="[1245,1270,1143,1169]" italics="true" pageId="10" pageNumber="577">Pl</emphasis>
) with triangleshaped projection (
<emphasis id="68FD89349B18A401BE1789E1FE1AFB19" box="[365,389,1180,1205]" italics="true" pageId="10" pageNumber="577">Ts</emphasis>
), robust terminal apophysis (
<emphasis id="68FD89349B18A401BDB689E1FD78FB19" box="[716,743,1180,1205]" italics="true" pageId="10" pageNumber="577">Ta</emphasis>
) and relatively straight embolus (
<emphasis id="68FD89349B18A401BB2589E1FB1BFB19" box="[1119,1156,1180,1205]" italics="true" pageId="10" pageNumber="577">Em</emphasis>
). The males of these two species differ by the tip of the embolus (bilobated in
<taxonomicName id="9D892EA59B18A401BD8989C2FCFBFB75" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[755,868,1215,1241]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="levinae">
<emphasis id="68FD89349B18A401BD8989C2FCFBFB75" box="[755,868,1215,1241]" italics="true" pageId="10" pageNumber="577">A. levinae</emphasis>
</taxonomicName>
, and single-blade in
<taxonomicName id="9D892EA59B18A401BB2889C2FB53FB75" baseAuthorityName="Caporiacco" baseAuthorityYear="1935" box="[1106,1228,1215,1241]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="baltoroi">
<emphasis id="68FD89349B18A401BB2889C2FB53FB75" box="[1106,1228,1215,1241]" italics="true" pageId="10" pageNumber="577">A. baltoroi</emphasis>
</taxonomicName>
) and by the shape of the terminal apophysis (curved and bifurcated in
<taxonomicName id="9D892EA59B18A401BDA6899EFCD2FB51" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[732,845,1251,1277]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="levinae">
<emphasis id="68FD89349B18A401BDA6899EFCD2FB51" box="[732,845,1251,1277]" italics="true" pageId="10" pageNumber="577">A. levinae</emphasis>
</taxonomicName>
, and straight and blunt in
<taxonomicName id="9D892EA59B18A401BB09899EFB74FB51" baseAuthorityName="Caporiacco" baseAuthorityYear="1935" box="[1139,1259,1251,1277]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="baltoroi">
<emphasis id="68FD89349B18A401BB09899EFB74FB51" box="[1139,1259,1251,1277]" italics="true" pageId="10" pageNumber="577">A. baltoroi</emphasis>
</taxonomicName>
, cf. Fig. 29 and fig.
<quantity id="9D71F8C39B18A401BFBF887AFF61FA8D" box="[197,254,1287,1313]" metricMagnitude="0" metricUnit="m" metricValue="1.524" pageId="10" pageNumber="577" unit="in" value="60.0">60 in</quantity>
<bibRefCitation id="3E1828D79B18A401BE7C887AFE77FA8D" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." box="[262,488,1287,1313]" pageId="10" pageNumber="577" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">
Marusik
<emphasis id="68FD89349B18A401BE118875FE3BFA8D" box="[363,420,1287,1313]" italics="true" pageId="10" pageNumber="577">et al.</emphasis>
2004
</bibRefCitation>
). The female of
<taxonomicName id="9D892EA59B18A401BDDA887AFC8EFA8D" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[672,785,1287,1313]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="levinae">
<emphasis id="68FD89349B18A401BDDA887AFC8EFA8D" box="[672,785,1287,1313]" italics="true" pageId="10" pageNumber="577">A. levinae</emphasis>
</taxonomicName>
differs from those of
<taxonomicName id="9D892EA59B18A401BB72887AFB1FFA8D" baseAuthorityName="Caporiacco" baseAuthorityYear="1935" box="[1032,1152,1287,1313]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="baltoroi">
<emphasis id="68FD89349B18A401BB72887AFB1FFA8D" box="[1032,1152,1287,1313]" italics="true" pageId="10" pageNumber="577">A. baltoroi</emphasis>
</taxonomicName>
by the shape of epigynal fovea (
<emphasis id="68FD89349B18A401BF9D8851FE99FAE9" box="[231,262,1324,1349]" italics="true" pageId="10" pageNumber="577">Fo</emphasis>
), which is rectangular in the former species (vs. heart-shaped) and by rectangular septal base (
<emphasis id="68FD89349B18A401BA3F8856FAFEFAE9" box="[1349,1377,1323,1349]" italics="true" pageId="10" pageNumber="577">Sb</emphasis>
) (vs. anchor-shaped, cf.
<figureCitation id="C2B249A39B18A401BE0A8832FE5DFAC5" box="[368,450,1359,1385]" captionStart="FIGURES 63–71" captionStartId="11.[151,264,1861,1885]" captionTargetBox="[151,1436,489,1838]" captionTargetId="figure-147@11.[151,1436,489,1838]" captionTargetPageId="11" captionText="FIGURES 63–71. Epigyne and female internal genitalia of Acantholycosa kulikovi sp. n. (63–65), A. levinae (66–68) and Mongolicosa lizae sp. n. (69–71). 63, 66, 69—intact, ventral; 64, 67, 70—macerated, ventral; 65, 68, 71—macerated, dorsal. Scale bars=0.2 mm. Abbreviations: Al apical lips of the epigyne, Ap apical pocket, Cd copulatory ducts, Fo epigynal fovea, Li lip of the epigyne, Re receptacle, Sa septal base arm, Sb septal base, Ss septal stem." figureDoi="http://doi.org/10.5281/zenodo.5300804" httpUri="https://zenodo.org/record/5300804/files/figure.png" pageId="10" pageNumber="577">Fig. 66</figureCitation>
and fig. 2D in
<bibRefCitation id="3E1828D79B18A401BD0C8832FC92FAC6" author="Buchar, J." box="[630,781,1359,1386]" pageId="10" pageNumber="577" pagination="201 - 227" refId="ref9724" refString="Buchar, J. (1976) Uber einige Lycosiden (Araneae) aus Nepal. Ergebnisse des Forschungsunternehmens Nepal Himalaya, 5, 201 - 227." type="journal article" year="1976">Buchar 1976</bibRefCitation>
). The epigyne of
<taxonomicName id="9D892EA59B18A401BCA38832FBD4FAC5" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[985,1099,1359,1385]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="levinae">
<emphasis id="68FD89349B18A401BCA38832FBD4FAC5" box="[985,1099,1359,1385]" italics="true" pageId="10" pageNumber="577">A. levinae</emphasis>
</taxonomicName>
is similar to those of species from the
<taxonomicName id="9D892EA59B18A401BF85880EFEC0FA21" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[255,351,1395,1421]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="plumalis">
<emphasis id="68FD89349B18A401BF85880EFEC0FA21" box="[255,351,1395,1421]" italics="true" pageId="10" pageNumber="577">plumalis</emphasis>
</taxonomicName>
-group:
<taxonomicName id="9D892EA59B18A401BEC3880EFC78FA21" authority="Marusik, Azarkina &amp; Koponen, 2004" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[441,999,1395,1422]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="logunovi">
<emphasis id="68FD89349B18A401BEC3880EFDA5FA21" box="[441,570,1395,1421]" italics="true" pageId="10" pageNumber="577">A. logunovi</emphasis>
<bibRefCitation id="3E1828D79B18A401BD38880EFC78FA21" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." box="[578,999,1395,1422]" pageId="10" pageNumber="577" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">Marusik, Azarkina &amp; Koponen, 2004</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="9D892EA59B18A401BC88880EFEDAFA1D" authority="Marusik, Azarkina &amp; Koponen, 2004" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="paraplumalis">
<emphasis id="68FD89349B18A401BC88880EFB38FA21" box="[1010,1191,1395,1421]" italics="true" pageId="10" pageNumber="577">A. paraplumalis</emphasis>
<bibRefCitation id="3E1828D79B18A401BBD4880EFEDAFA1D" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." pageId="10" pageNumber="577" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">Marusik, Azarkina &amp; Koponen, 2004</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="9D892EA59B18A401BE0788EAFC32FA1D" authority="Marusik, Azarkina &amp; Koponen, 2004" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[381,941,1431,1458]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="plumalis">
<emphasis id="68FD89349B18A401BE0788EAFE62FA1D" box="[381,509,1431,1457]" italics="true" pageId="10" pageNumber="577">A. plumalis</emphasis>
<bibRefCitation id="3E1828D79B18A401BD7F88EAFC32FA1D" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." box="[517,941,1431,1458]" pageId="10" pageNumber="577" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">Marusik, Azarkina &amp; Koponen, 2004</bibRefCitation>
</taxonomicName>
. The similarity includes a rectangular shape of the fovea as well as septal base and apical pocket (
<emphasis id="68FD89349B18A401BD8C88C6FC8AFA79" box="[758,789,1467,1493]" italics="true" pageId="10" pageNumber="577">Ap</emphasis>
) consisting of one hood. The female of
<taxonomicName id="9D892EA59B18A401BBA188C6FAD3FA79" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[1243,1356,1467,1493]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="levinae">
<emphasis id="68FD89349B18A401BBA188C6FAD3FA79" box="[1243,1356,1467,1493]" italics="true" pageId="10" pageNumber="577">A. levinae</emphasis>
</taxonomicName>
differs from all aforementioned species by a reduced septal stem. In addition, it can be separated from
<taxonomicName id="9D892EA59B18A401BBB188A2FAD2FA55" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[1227,1357,1503,1529]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="logunovi">
<emphasis id="68FD89349B18A401BBB188A2FAD2FA55" box="[1227,1357,1503,1529]" italics="true" pageId="10" pageNumber="577">A. logunovi</emphasis>
</taxonomicName>
and
<taxonomicName id="9D892EA59B18A401BAFE88A2FEB3F9B1" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="paraplumalis">
<emphasis id="68FD89349B18A401BAFE88A2FEB3F9B1" italics="true" pageId="10" pageNumber="577">A. paraplumalis</emphasis>
</taxonomicName>
by having the apical pocket as wide as the septal base (vs. twice narrower). Finally, the female of
<taxonomicName id="9D892EA59B18A401BAFE8B7EFF77F9ED" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="levinae">
<emphasis id="68FD89349B18A401BAFE8B7EFF77F9ED" italics="true" pageId="10" pageNumber="577">A. levinae</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="9D892EA59B18A401BD5A8B5AFD3EF9ED" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[544,673,1575,1601]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="plumalis">
<emphasis id="68FD89349B18A401BD5A8B5AFD3EF9ED" box="[544,673,1575,1601]" italics="true" pageId="10" pageNumber="577">A. plumalis</emphasis>
</taxonomicName>
by having receptacles (
<emphasis id="68FD89349B18A401BCC88B55FC4FF9ED" box="[946,976,1576,1601]" italics="true" pageId="10" pageNumber="577">Re</emphasis>
) almost touching each other (vs. widely spaced, cf.
<figureCitation id="C2B249A39B18A401BE6F8B36FE11F9C9" box="[277,398,1611,1637]" captionStart="FIGURES 63–71" captionStartId="11.[151,264,1861,1885]" captionTargetBox="[151,1436,489,1838]" captionTargetId="figure-147@11.[151,1436,489,1838]" captionTargetPageId="11" captionText="FIGURES 63–71. Epigyne and female internal genitalia of Acantholycosa kulikovi sp. n. (63–65), A. levinae (66–68) and Mongolicosa lizae sp. n. (69–71). 63, 66, 69—intact, ventral; 64, 67, 70—macerated, ventral; 65, 68, 71—macerated, dorsal. Scale bars=0.2 mm. Abbreviations: Al apical lips of the epigyne, Ap apical pocket, Cd copulatory ducts, Fo epigynal fovea, Li lip of the epigyne, Re receptacle, Sa septal base arm, Sb septal base, Ss septal stem." figureDoi="http://doi.org/10.5281/zenodo.5300804" httpUri="https://zenodo.org/record/5300804/files/figure.png" pageId="10" pageNumber="577">Fig. 66, 68</figureCitation>
and fig.
<quantity id="9D71F8C39B18A401BE888B36FD82F9C9" box="[498,541,1611,1637]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="10" pageNumber="577" unit="in" value="3.0">3 in</quantity>
<bibRefCitation id="3E1828D79B18A401BD5E8B36FCC9F9C9" author="Fomichev, A. A. &amp; Marusik, Y. M." box="[548,854,1611,1637]" pageId="10" pageNumber="577" pagination="65 - 68" refId="ref9802" refString="Fomichev, A. A. &amp; Marusik, Y. M. (2011) First description of the female of Acantholycosa logunovi (Araneae: Lycosidae). Zootaxa, 2813 (1), 65 - 68. https: // doi. org / 10.11646 / zootaxa. 2813.1.3" type="journal article" year="2011">Fomichev &amp; Marusik 2011</bibRefCitation>
, and figs 47, 50–
<quantity id="9D71F8C39B18A401BB6C8B36FBD1F9C9" box="[1046,1102,1611,1638]" metricMagnitude="0" metricUnit="m" metricValue="1.3208" pageId="10" pageNumber="577" unit="in" value="52.0">52 in</quantity>
<bibRefCitation id="3E1828D79B18A401BB2F8B36FAAAF9C9" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." box="[1109,1333,1611,1637]" pageId="10" pageNumber="577" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">
Marusik
<emphasis id="68FD89349B18A401BBC38B31FB6DF9C9" box="[1209,1266,1611,1637]" italics="true" pageId="10" pageNumber="577">et al.</emphasis>
2004
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="129306AD9B18A400BFBD8B12FDB8FF55" lastPageId="11" lastPageNumber="578" pageId="10" pageNumber="577" type="description">
<paragraph id="5A3655269B18A401BFBD8B12FABBF892" blockId="10.[151,1437,927,2034]" pageId="10" pageNumber="577">
<emphasis id="68FD89349B18A401BFBD8B12FEC6F925" bold="true" box="[199,345,1647,1673]" pageId="10" pageNumber="577">Description.</emphasis>
Female.Total length 8.6. Carapace: 4.15 long, 3.3 wide. General appearance as in
<figureCitation id="C2B249A39B18A401BBAE8B12FA8BF926" box="[1236,1300,1647,1674]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1832,1856]" captionTargetBox="[157,1436,189,1799]" captionTargetId="figure-33@2.[151,1436,181,1809]" captionTargetPageId="2" captionText="FIGURES 1–9. General appearance of Acantholycosa sayanensis (1–2), A. kulikovi sp. n. (3), A. levinae (4–5), A. sergeevi sp. n. (6), Mongolicosa lizae sp. n. (8–9) and leg I of A. sergeevi sp. n. (7). 1, 4, 6, 8—male, dorsal; 2, 3, 5, 9—female, dorsal; 7—leg I of male, prolateral. Scale bars=2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300794" httpUri="https://zenodo.org/record/5300794/files/figure.png" pageId="10" pageNumber="577">Fig. 5</figureCitation>
.Coloration. Carapace, sternum and labium dark brown. Eye field black. Chelicerae, endites and palps yellow-brown. Coxae yellow-gray. Legs yellow-brown, femora with transverse dark gray stripes. Abdomen dark gray dorsally, with dark brown cardiac mark. Venter of abdomen yellow-gray. Spinnerets dark gray. Spination of leg I: Fe d1-1-1 p0-0-2 r0- 1-1; Ti p1-0-0 r1-0-0 v1-2-2-2-2-2; Mt p0-1-0 r0-1-0 v2-2-0. Leg measurements: I 3.9, 1.8, 3.65, 3.45, 1.7 (14.5); II 3.9, 1.75, 3.4, 3.55, 1.75 (14.35); III 3.8, 1.6, 3.15, 4.3, 1.85 (14.7); IV 4.85, 1.8, 4.3, 6.85, 2.55 (20.35).
</paragraph>
<paragraph id="5A3655269B18A401BFBD8A3AFC47F805" blockId="10.[151,1437,927,2034]" pageId="10" pageNumber="577">
Epigyne and internal genitalia as shown in
<figureCitation id="C2B249A39B18A401BDCB8A3AFCACF8CD" box="[689,819,1863,1889]" captionStart="FIGURES 63–71" captionStartId="11.[151,264,1861,1885]" captionTargetBox="[151,1436,489,1838]" captionTargetId="figure-147@11.[151,1436,489,1838]" captionTargetPageId="11" captionText="FIGURES 63–71. Epigyne and female internal genitalia of Acantholycosa kulikovi sp. n. (63–65), A. levinae (66–68) and Mongolicosa lizae sp. n. (69–71). 63, 66, 69—intact, ventral; 64, 67, 70—macerated, ventral; 65, 68, 71—macerated, dorsal. Scale bars=0.2 mm. Abbreviations: Al apical lips of the epigyne, Ap apical pocket, Cd copulatory ducts, Fo epigynal fovea, Li lip of the epigyne, Re receptacle, Sa septal base arm, Sb septal base, Ss septal stem." figureDoi="http://doi.org/10.5281/zenodo.5300804" httpUri="https://zenodo.org/record/5300804/files/figure.png" pageId="10" pageNumber="577">Figs 66–68</figureCitation>
. Fovea as wide as long. Septal base smoothly declines anteriorly. Lips of the epigyne (
<emphasis id="68FD89349B18A401BE808A11FD8EF829" box="[506,529,1900,1925]" italics="true" pageId="10" pageNumber="577">Li</emphasis>
) separated by one of their wide. Receptacles clavate, not clearly delineated from copulatory ducts (
<emphasis id="68FD89349B18A401BE1B8AF2FE1DF805" box="[353,386,1935,1961]" italics="true" pageId="10" pageNumber="577">Cd</emphasis>
). Copulatory ducts 1.5 times longer than receptacles.
</paragraph>
<paragraph id="5A3655269B18A400BFBD8ACEFDB8FF55" blockId="10.[151,1437,927,2034]" lastBlockId="11.[151,1437,151,465]" lastPageId="11" lastPageNumber="578" pageId="10" pageNumber="577">
Male. See
<bibRefCitation id="3E1828D79B18A401BE448ACEFDB4F861" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." box="[318,555,1971,1997]" pageId="10" pageNumber="577" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">
Marusik
<emphasis id="68FD89349B18A401BED88AC9FE45F861" box="[418,474,1971,1997]" italics="true" pageId="10" pageNumber="577">et al.</emphasis>
(2004)
</bibRefCitation>
for description. General appearance as in
<figureCitation id="C2B249A39B18A401BB788ACEFBD9F861" box="[1026,1094,1971,1997]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1832,1856]" captionTargetBox="[157,1436,189,1799]" captionTargetId="figure-33@2.[151,1436,181,1809]" captionTargetPageId="2" captionText="FIGURES 1–9. General appearance of Acantholycosa sayanensis (1–2), A. kulikovi sp. n. (3), A. levinae (4–5), A. sergeevi sp. n. (6), Mongolicosa lizae sp. n. (8–9) and leg I of A. sergeevi sp. n. (7). 1, 4, 6, 8—male, dorsal; 2, 3, 5, 9—female, dorsal; 7—leg I of male, prolateral. Scale bars=2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300794" httpUri="https://zenodo.org/record/5300794/files/figure.png" pageId="10" pageNumber="577">Fig. 4</figureCitation>
. Palp as in
<figureCitation id="C2B249A39B18A401BBBE8ACEFADBF861" box="[1220,1348,1971,1997]" captionStart="FIGURES 10–19" captionStartId="3.[151,264,1849,1873]" captionTargetBox="[155,1425,188,1810]" captionTargetId="figure-15@3.[151,1436,181,1825]" captionTargetPageId="3" captionText="FIGURES 10–19. Male palp (10–13) and bulb (14–19) of Acantholycosa sergeevi sp. n. (10–11), A. sayanensis (14–15), A. sidorovi sp. n. (16–17), A. levinae (18-19) and Mongolicosa lizae sp. n. (12–13). 10, 12—retrolateral; 11, 13, 14, 16, 18— ventral; 15, 17, 19—anterior. Scale bars=0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300796" httpUri="https://zenodo.org/record/5300796/files/figure.png" pageId="10" pageNumber="577">Figs 18–19</figureCitation>
, 28–31, 36–37. Tegular apophysis (
<emphasis id="68FD89349B18A401BEBB8AA5FE40F85D" box="[449,479,2008,2033]" italics="true" pageId="10" pageNumber="577">Tg</emphasis>
) almost oval, with reduced apical and basal arms. Palea with abrupt retrolateral margin. Triangle shaped projection almost straight, with a hooked apical part. Terminal apophysis massive, subdivided into two lobes. Embolus with a spine (
<emphasis id="68FD89349B19A400BD2B8DC6FDF2FF79" box="[593,621,187,213]" italics="true" pageId="11" pageNumber="578">Sp</emphasis>
) in the basal part. Distal part of the embolus subdivided into triangular secondary part and embolus proper.
</paragraph>
</subSubSection>
<subSubSection id="129306AD9B19A400BFBD8C7FFF5BFEEE" pageId="11" pageNumber="578" type="distribution">
<paragraph id="5A3655269B19A400BFBD8C7FFF5BFEEE" blockId="11.[151,1437,151,465]" pageId="11" pageNumber="578">
<emphasis id="68FD89349B19A400BFBD8C7FFEFEFEB0" bold="true" box="[199,353,258,284]" pageId="11" pageNumber="578">Distribution.</emphasis>
Known from several close localities in the Katunskyi Mt. Range in
<collectingRegion id="984D9BC49B19A400BB158C7EFB38FEB1" box="[1135,1191,259,285]" country="Russia" name="Altay" pageId="11" pageNumber="578">Altai</collectingRegion>
Mountains (
<figureCitation id="C2B249A39B19A400BA428C7EFF28FEEE" captionStart="FIGURES 72–77" captionStartId="13.[151,264,1676,1700]" captionTargetBox="[151,1436,309,1652]" captionTargetId="figure-76@13.[151,1436,309,1652]" captionTargetPageId="13" captionText="FIGURES 72–77. Male of Acantholycosa sayanensis from Borus Mt. Range (72), female of Acantholycosa sp. (A. azyuzini or A. levinae) with egg sac, from Katunskyi Mt. Range (73), habitat of A. sayanensis and A. sergeevi sp. n. in Borus Mt. Range (74), collecting localities of studied species from the Acantholycosa-complex (75–76) and map showing the main mountain systems of South Siberia and adjacent regions (77). Diamond—A. sayanensis, square—A. sidorovi sp. n., triangle—A. kulikovi sp. n., star—A. levinae, circle—A. sergeevi sp. n., inverted triangle—Mongolicosa lizae sp. n. The frame of Fig. 76 refers to the content of Fig. 75." figureDoi="http://doi.org/10.5281/zenodo.5300806" httpUri="https://zenodo.org/record/5300806/files/figure.png" pageId="11" pageNumber="578">Figs 75– 76</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="129306AD9B19A400BFBD8C37FA1BFE7D" pageId="11" pageNumber="578" type="discussion">
<paragraph id="5A3655269B19A400BFBD8C37FA1BFE7D" blockId="11.[151,1437,151,465]" pageId="11" pageNumber="578">
<emphasis id="68FD89349B19A400BFBD8C37FED0FEC8" bold="true" box="[199,335,330,356]" pageId="11" pageNumber="578">Comments.</emphasis>
There are some doubts if the females match with males due to the morphology of the epigyne. Since
<taxonomicName id="9D892EA59B19A400BFED8C12FE97FE25" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[151,264,367,393]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="578" phylum="Arthropoda" rank="species" species="levinae">
<emphasis id="68FD89349B19A400BFED8C12FE97FE25" box="[151,264,367,393]" italics="true" pageId="11" pageNumber="578">A. levinae</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="9D892EA59B19A400BEC28C12FD8EFE25" baseAuthorityName="Caporiacco" baseAuthorityYear="1935" box="[440,529,367,393]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="578" phylum="Arthropoda" rank="species" species="baltoroi">
<emphasis id="68FD89349B19A400BEC28C12FD8EFE25" box="[440,529,367,393]" italics="true" pageId="11" pageNumber="578">baltoroi</emphasis>
</taxonomicName>
-group, this species epigyne would be expected to have an anchor-shaped septal base. Instead, the epigyne of the females described herein are similar to those of species from the
<taxonomicName id="9D892EA59B19A400BB908CEEFAD5FE01" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[1258,1354,403,429]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="578" phylum="Arthropoda" rank="species" species="plumalis">
<emphasis id="68FD89349B19A400BB908CEEFAD5FE01" box="[1258,1354,403,429]" italics="true" pageId="11" pageNumber="578">plumalis</emphasis>
</taxonomicName>
-group. Nevertheless, males and females were collected together in two different places, thus I treat them as conspecific.
</paragraph>
</subSubSection>
</treatment>
</document>

147
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<mods:title id="D26D01D45254E0D6F6638864ECA658D1">New data on the wolf spiders from theAcantholycosa-complex (Araneae: Lycosidae) from the South Siberia</mods:title>
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<emphasis id="68FD89349B18A401BFED8DEAFD9FFF1D" bold="true" box="[151,512,151,178]" pageId="10" pageNumber="577">
<taxonomicName id="9D892EA59B18A401BFED8DEAFE30FF1E" ID-CoL="64GBY" baseAuthorityName="Caporiacco" baseAuthorityYear="1935" box="[151,431,151,178]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="baltoroi">
<emphasis id="68FD89349B18A401BFED8DEAFE30FF1E" bold="true" box="[151,431,151,178]" italics="true" pageId="10" pageNumber="577">Acantholycosa baltoroi</emphasis>
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group
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<paragraph id="5A3655269B18A401BFED8DA1FD74FF58" blockId="10.[151,1437,220,640]" box="[151,747,220,244]" pageId="10" pageNumber="577">
<treatmentCitationGroup id="7A9972089B18A401BFED8DA1FD74FF58" box="[151,747,220,244]" pageId="10" pageNumber="577">
<taxonomicName id="9D892EA59B18A401BFED8DA1FE1BFF58" ID-CoL="64GBY" baseAuthorityName="Caporiacco" baseAuthorityYear="1935" box="[151,388,220,244]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="baltoroi">
<emphasis id="68FD89349B18A401BFED8DA1FE1BFF58" box="[151,388,220,244]" italics="true" pageId="10" pageNumber="577">Acantholycosa baltoroi</emphasis>
</taxonomicName>
group —
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<bibRefCitation id="3E1828D79B18A401BE918DA1FD78FF58" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." box="[491,743,220,244]" pageId="10" pageNumber="577" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">
Marusik
<emphasis id="68FD89349B18A401BD3C8DA1FDE6FF58" box="[582,633,220,244]" italics="true" pageId="10" pageNumber="577">et al.</emphasis>
2004: 112
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.
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</paragraph>
</subSubSection>
<subSubSection id="129306AD9B18A401BFED8D80FEBFFDB9" pageId="10" pageNumber="577" type="diagnosis">
<paragraph id="5A3655269B18A401BFED8D80FC4CFECC" blockId="10.[151,1437,220,640]" pageId="10" pageNumber="577">
<emphasis id="68FD89349B18A401BFED8D80FE8DFEBB" bold="true" box="[151,274,253,279]" pageId="10" pageNumber="577">Diagnosis.</emphasis>
Members of this group can be easily distinguished based on the following features: palea with a prominent extension in the terminal part and a triangle-shaped projection on the posterior edge, robust terminal apophysis with bifurcate or truncate tip, and anchor-shaped septum (
<bibRefCitation id="3E1828D79B18A401BD9F8C3BFC58FECC" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." box="[741,967,326,352]" pageId="10" pageNumber="577" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">
Marusik
<emphasis id="68FD89349B18A401BC318C3AFC1BFECC" box="[843,900,326,352]" italics="true" pageId="10" pageNumber="577">et al.</emphasis>
2004
</bibRefCitation>
).
</paragraph>
<paragraph id="5A3655269B18A401BFBD8C14FEBFFDB9" blockId="10.[151,1437,220,640]" pageId="10" pageNumber="577">
<emphasis id="68FD89349B18A401BFBD8C14FEF8FE2F" bold="true" box="[199,359,361,387]" pageId="10" pageNumber="577">Composition.</emphasis>
This species group includes six species:
<taxonomicName id="9D892EA59B18A401BC388C17FB3EFE29" authority="(Caporiacco, 1935)" baseAuthorityName="Caporiacco" baseAuthorityYear="1935" box="[834,1185,362,389]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="baltoroi">
<emphasis id="68FD89349B18A401BC388C17FC22FE28" box="[834,957,362,388]" italics="true" pageId="10" pageNumber="577">A. baltoroi</emphasis>
(Caporiacco, 1935)
</taxonomicName>
(Himalaya,
<collectingRegion id="984D9BC49B18A401BA4A8C17FAF5FE28" box="[1328,1386,362,388]" country="China" name="Xizang" pageId="10" pageNumber="577">Tibet</collectingRegion>
and Central, Northern and Northeastern
<collectingCountry id="229E15B69B18A401BD4D8CF3FDE1FE05" box="[567,638,398,425]" name="China" pageId="10" pageNumber="577">China</collectingCountry>
),
<taxonomicName id="9D892EA59B18A401BDEE8CF3FB20FE04" authority="Marusik, Azarkina &amp; Koponen, 2004" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[660,1215,398,424]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="levinae">
<emphasis id="68FD89349B18A401BDEE8CF3FC98FE04" box="[660,775,398,424]" italics="true" pageId="10" pageNumber="577">A. levinae</emphasis>
<bibRefCitation id="3E1828D79B18A401BC6B8CF3FB20FE04" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." box="[785,1215,398,424]" pageId="10" pageNumber="577" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">Marusik, Azarkina &amp; Koponen, 2004</bibRefCitation>
</taxonomicName>
(Altai Mountains),
<taxonomicName id="9D892EA59B18A401BFED8CCFFE88FE60" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[151,279,434,460]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="sergeevi" status="sp. nov.">
<emphasis id="68FD89349B18A401BFED8CCFFE88FE60" box="[151,279,434,460]" italics="true" pageId="10" pageNumber="577">A. sergeevi</emphasis>
</taxonomicName>
<emphasis id="68FD89349B18A401BE588CCFFEF9FE60" bold="true" box="[290,358,434,460]" pageId="10" pageNumber="577">
<taxonomicNameLabel id="73CE344F9B18A401BE588CCFFEF9FE60" box="[290,358,434,460]" pageId="10" pageNumber="577" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(Western Sayan Mountains),
<taxonomicName id="9D892EA59B18A401BDBE8CCFFBA2FE61" authority="(Levi &amp; Levi, 1951)" baseAuthorityName="Levi &amp; Levi" baseAuthorityYear="1951" box="[708,1085,434,461]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="solituda">
<emphasis id="68FD89349B18A401BDBE8CCFFCDEFE60" box="[708,833,434,460]" italics="true" pageId="10" pageNumber="577">A. solituda</emphasis>
(Levi &amp; Levi, 1951)
</taxonomicName>
(Rocky Mountains,
<collectingCountry id="229E15B69B18A401BA4A8CCFFAF3FE60" box="[1328,1388,434,460]" name="United States of America" pageId="10" pageNumber="577">USA</collectingCountry>
),
<taxonomicName id="9D892EA59B18A401BAFE8CCFFE35FE5C" authority="(Marusik, 1993)" baseAuthorityName="Marusik" baseAuthorityYear="1993" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="sterneri">
<emphasis id="68FD89349B18A401BAFE8CCFFF71FE5C" italics="true" pageId="10" pageNumber="577">A. sterneri</emphasis>
(
<bibRefCitation id="3E1828D79B18A401BF868CABFE3DFE5C" author="Marusik, Y. M." box="[252,418,470,496]" pageId="10" pageNumber="577" pagination="77 - 81" refId="ref10315" refString="Marusik, Y. M. (1993) Three new wandering spider species (Aranei Lycosidae Gnaphosidae) from Mongolia. Arthropoda Selecta, 2 (1), 77 - 81." type="journal article" year="1993">Marusik, 1993</bibRefCitation>
)
</taxonomicName>
(Southern Siberia and
<collectingCountry id="229E15B69B18A401BDD38CABFC87FE5D" box="[681,792,470,497]" name="Mongolia" pageId="10" pageNumber="577">Mongolia</collectingCountry>
) and
<taxonomicName id="9D892EA59B18A401BC2E8CABFB63FE5C" authority="Marusik &amp; Omelko, 2017" authorityName="Marusik &amp; Omelko" authorityYear="2017" box="[852,1276,470,496]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="zonsteini">
<emphasis id="68FD89349B18A401BC2E8CABFC48FE5C" box="[852,983,470,496]" italics="true" pageId="10" pageNumber="577">A. zonsteini</emphasis>
<bibRefCitation id="3E1828D79B18A401BCA78CABFB63FE5C" author="Marusik, Y. M. &amp; Omelko, M. M." box="[989,1276,470,496]" pageId="10" pageNumber="577" pagination="597 - 600" refId="ref10588" refString="Marusik, Y. M. &amp; Omelko, M. M. (2017) A new species of Acantholycosa baltoroi - group (Araneae: Lycosidae, Pardosinae) from the Russian Far East. Zootaxa, 4232 (4), 597 - 600. https: // doi. org / 10.11646 / zootaxa. 4232.4.12" type="journal article" year="2017">Marusik &amp; Omelko, 2017</bibRefCitation>
</taxonomicName>
(Sikhote-Alin Mountains).
</paragraph>
</subSubSection>
<subSubSection id="129306AD9B18A401BFBD8F60FE80FD2C" pageId="10" pageNumber="577" type="discussion">
<paragraph id="5A3655269B18A401BFBD8F60FE80FD2C" blockId="10.[151,1437,220,640]" pageId="10" pageNumber="577">
<emphasis id="68FD89349B18A401BFBD8F60FED0FD9B" bold="true" box="[199,335,541,567]" pageId="10" pageNumber="577">Comments.</emphasis>
It should be noted that the
<taxonomicName id="9D892EA59B18A401BDFE8F63FD42FD94" baseAuthorityName="Caporiacco" baseAuthorityYear="1935" box="[644,733,542,568]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="577" phylum="Arthropoda" rank="species" species="baltoroi">
<emphasis id="68FD89349B18A401BDFE8F63FD42FD94" box="[644,733,542,568]" italics="true" pageId="10" pageNumber="577">baltoroi</emphasis>
</taxonomicName>
-group might represent a separate genus due to the unique cutlike palea with a relatively small, triangle-shaped projection, instead of well-developed paleal apophysis (
<bibRefCitation id="3E1828D79B18A401BA478F3FFE8CFD2C" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." pageId="10" pageNumber="577" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">
Marusik
<emphasis id="68FD89349B18A401BFED8F1AFF50FD2C" box="[151,207,614,640]" italics="true" pageId="10" pageNumber="577">et al.</emphasis>
2004
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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<emphasis id="68FD89349B1BA402BFED8C37FE2FFEC9" bold="true" box="[151,432,330,357]" italics="true" pageId="9" pageNumber="576">Acantholycosa lignaria</emphasis>
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Marusik
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<emphasis id="68FD89349B1BA402BFED8CA8FE8DFE43" bold="true" box="[151,274,469,495]" pageId="9" pageNumber="576">Diagnosis.</emphasis>
Representatives of this group can be recognized only from male characters. The males from
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- group possess a laminar paleal outgrowth, strong terminal apophysis, and reduced apical arm of tegular apophysis (
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Marusik
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). Females of species assigned to this species group have very heterogeneous epigynes.
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<emphasis id="68FD89349B1BA402BFBD8F3CFEF8FDF7" bold="true" box="[199,359,577,603]" pageId="9" pageNumber="576">Composition.</emphasis>
Four species belong to this species group:
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Zyuzin &amp; Marusik, 1988
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(from Central Aimag in Mongolia to the Kolyma River and Sikhote-Alin Mt. Range),
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(Sikhote-Alin Mt. Range),
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Mountains) and
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<figureCitation id="C2B249A39B1BA402BFED8E40FF42FCFB" box="[151,221,829,855]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1832,1856]" captionTargetBox="[157,1436,189,1799]" captionTargetId="figure-33@2.[151,1436,181,1809]" captionTargetPageId="2" captionText="FIGURES 1–9. General appearance of Acantholycosa sayanensis (1–2), A. kulikovi sp. n. (3), A. levinae (4–5), A. sergeevi sp. n. (6), Mongolicosa lizae sp. n. (8–9) and leg I of A. sergeevi sp. n. (7). 1, 4, 6, 8—male, dorsal; 2, 3, 5, 9—female, dorsal; 7—leg I of male, prolateral. Scale bars=2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300794" httpUri="https://zenodo.org/record/5300794/files/figure.png" pageId="9" pageNumber="576">Figs 3</figureCitation>
,
<figureCitation id="C2B249A39B1BA402BF908E40FEABFCF4" box="[234,308,829,856]" captionStart="FIGURES 63–71" captionStartId="11.[151,264,1861,1885]" captionTargetBox="[151,1436,489,1838]" captionTargetId="figure-147@11.[151,1436,489,1838]" captionTargetPageId="11" captionText="FIGURES 63–71. Epigyne and female internal genitalia of Acantholycosa kulikovi sp. n. (63–65), A. levinae (66–68) and Mongolicosa lizae sp. n. (69–71). 63, 66, 69—intact, ventral; 64, 67, 70—macerated, ventral; 65, 68, 71—macerated, dorsal. Scale bars=0.2 mm. Abbreviations: Al apical lips of the epigyne, Ap apical pocket, Cd copulatory ducts, Fo epigynal fovea, Li lip of the epigyne, Re receptacle, Sa septal base arm, Sb septal base, Ss septal stem." figureDoi="http://doi.org/10.5281/zenodo.5300804" httpUri="https://zenodo.org/record/5300804/files/figure.png" pageId="9" pageNumber="576">63–65</figureCitation>
,
<figureCitation id="C2B249A39B1BA402BE448E40FE1BFCF4" box="[318,388,829,856]" captionStart="FIGURES 72–77" captionStartId="13.[151,264,1676,1700]" captionTargetBox="[151,1436,309,1652]" captionTargetId="figure-76@13.[151,1436,309,1652]" captionTargetPageId="13" captionText="FIGURES 72–77. Male of Acantholycosa sayanensis from Borus Mt. Range (72), female of Acantholycosa sp. (A. azyuzini or A. levinae) with egg sac, from Katunskyi Mt. Range (73), habitat of A. sayanensis and A. sergeevi sp. n. in Borus Mt. Range (74), collecting localities of studied species from the Acantholycosa-complex (75–76) and map showing the main mountain systems of South Siberia and adjacent regions (77). Diamond—A. sayanensis, square—A. sidorovi sp. n., triangle—A. kulikovi sp. n., star—A. levinae, circle—A. sergeevi sp. n., inverted triangle—Mongolicosa lizae sp. n. The frame of Fig. 76 refers to the content of Fig. 75." figureDoi="http://doi.org/10.5281/zenodo.5300806" httpUri="https://zenodo.org/record/5300806/files/figure.png" pageId="9" pageNumber="576">75–76</figureCitation>
</paragraph>
<paragraph id="5A3655269B1BA402BFED8EF8FDB5FBA7" blockId="9.[151,1437,901,1900]" pageId="9" pageNumber="576">
<emphasis id="68FD89349B1BA402BFED8EF8FF7BFC33" bold="true" box="[151,228,901,927]" pageId="9" pageNumber="576">Types.</emphasis>
<materialsCitation id="EAE15F7B9B1BA402BF898EF8FB2AFC6F" collectingDate="2015-05-09" collectionCode="ISEA" collectorName="A. A. Fomichev" country="Kazakhstan" elevation="1550" latitude="50.25" location="Kazachikha River valley" longLatPrecision="1098" longitude="83.76667" municipality="Ivanovskyi Mt. Range" pageId="9" pageNumber="576" specimenCount="1" specimenCount-female="1" stateProvince="Shyghys Qazaqstan Oblysy" typeStatus="holotype">
<collectingCountry id="229E15B69B1BA402BF898EF8FE2AFC33" box="[243,437,901,927]" name="Kazakhstan" pageId="9" pageNumber="576">KAZAKHSTAN</collectingCountry>
:
<emphasis id="68FD89349B1BA402BEBD8EF8FD67FC33" bold="true" box="[455,760,901,927]" pageId="9" pageNumber="576">
<collectingRegion id="984D9BC49B1BA402BEBD8EF8FD67FC33" box="[455,760,901,927]" country="Kazakhstan" name="Shyghys Qazaqstan Oblysy" pageId="9" pageNumber="576">East Kazakhstan Oblast</collectingRegion>
</emphasis>
:
<typeStatus id="8532EB849B1BA402BC6A8EF8FCEDFC33" box="[784,882,901,927]" pageId="9" pageNumber="576" type="holotype">holotype</typeStatus>
<specimenCount id="4C8F9EAF9B1BA402BCFB8EFBFC09FC0C" box="[897,918,902,928]" pageId="9" pageNumber="576" type="female">♀</specimenCount>
(
<collectionCode id="3C98CDE39B1BA402BCD78EF8FC70FC33" box="[941,1007,901,927]" pageId="9" pageNumber="576">ISEA</collectionCode>
, 001.8656),
<collectingMunicipality id="BA52CF5C9B1BA402BBF18EF8FA07FC0C" box="[1163,1432,901,928]" pageId="9" pageNumber="576">Ivanovskyi Mt. Range</collectingMunicipality>
,
<location id="5F5603FD9B1BA402BFED8ED4FE34FC6F" LSID="urn:lsid:plazi:treatment:D220E4309B1BA402BFED8E64FF67F8C0:5F5603FD9B1BA402BFED8ED4FE34FC6F" box="[151,427,937,963]" country="Kazakhstan" latitude="50.25" longLatPrecision="1098" longitude="83.76667" municipality="Ivanovskyi Mt. Range" name="Kazachikha River valley" pageId="9" pageNumber="576" stateProvince="Shyghys Qazaqstan Oblysy">Kazachikha River valley</location>
,
<geoCoordinate id="3FBD33E19B1BA402BECF8ED4FD87FC68" box="[437,536,937,964]" degrees="50" direction="north" minutes="15" orientation="latitude" pageId="9" pageNumber="576" precision="925" value="50.25">50°15’N</geoCoordinate>
,
<geoCoordinate id="3FBD33E19B1BA402BD598ED4FD1CFC6F" box="[547,643,937,963]" degrees="83" direction="east" minutes="46" orientation="longitude" pageId="9" pageNumber="576" precision="925" value="83.76667">83°46’E</geoCoordinate>
, scree,
<quantity id="9D71F8C39B1BA402BDAE8ED4FCEFFC68" box="[724,880,937,964]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.7" metricValueMin="1.4" pageId="9" pageNumber="576" unit="m" value="1550.0" valueMax="1700.0" valueMin="1400.0">
<elevation id="D1A4B2159B1BA402BDAE8ED4FCEFFC68" box="[724,880,937,964]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.7" metricValueMin="1.4" pageId="9" pageNumber="576" unit="m" value="1550.0" valueMax="1700.0" valueMin="1400.0">1400–1700 m</elevation>
</quantity>
,
<date id="2E3773E69B1BA402BC018ED4FC64FC68" box="[891,1019,937,964]" pageId="9" pageNumber="576" value="2015-05-09">
<collectingDate id="3E738A0E9B1BA402BC018ED4FC64FC68" box="[891,1019,937,964]" pageId="9" pageNumber="576" value="2015-05-09">05.09.2015</collectingDate>
</date>
,
<collectorName id="F77C30F09B1BA402BB7E8ED7FB2AFC6F" box="[1028,1205,937,964]" pageId="9" pageNumber="576">A.A. Fomichev</collectorName>
</materialsCitation>
;
<materialsCitation id="EAE15F7B9B1BA402BBC48ED7FDB9FBA7" collectingDate="2015-09-03" collectionCode="ISEA" collectorName="A. A. Fomichev" country="Kazakhstan" county="Ivanovskyi Mt. Range" elevation="2000" latitude="50.3" location="Vyshe-Ivanovskyi Belok Mt." longLatPrecision="1098" longitude="83.86667" municipality="Voroshilova Peak" pageId="9" pageNumber="576" specimenCount="1" specimenCount-female="1" stateProvince="Shyghys Qazaqstan Oblysy" typeStatus="paratype">
<typeStatus id="8532EB849B1BA402BBC48ED7FA81FC68" box="[1214,1310,938,964]" pageId="9" pageNumber="576" type="paratype">paratype</typeStatus>
<specimenCount id="4C8F9EAF9B1BA402BA5F8ED4FAD7FC6F" box="[1317,1352,937,963]" pageId="9" pageNumber="576" type="female">1♀</specimenCount>
(
<collectionCode id="3C98CDE39B1BA402BA2C8ED4FA07FC6F" box="[1366,1432,937,963]" pageId="9" pageNumber="576">ISEA</collectionCode>
, 001.8657),
<collectingCounty id="B3572DAA9B1BA402BE628EB0FD8BFC44" box="[280,532,973,1000]" pageId="9" pageNumber="576">Ivanovskyi Mt. Range</collectingCounty>
,
<collectingMunicipality id="BA52CF5C9B1BA402BD658EB0FD7CFC4B" box="[543,739,973,999]" pageId="9" pageNumber="576">Voroshilova Peak</collectingMunicipality>
(=
<location id="5F5603FD9B1BA402BC7B8EB0FBD5FC44" LSID="urn:lsid:plazi:treatment:D220E4309B1BA402BFED8E64FF67F8C0:5F5603FD9B1BA402BC7B8EB0FBD5FC44" box="[769,1098,973,1000]" country="Kazakhstan" county="Ivanovskyi Mt. Range" latitude="50.3" longLatPrecision="1098" longitude="83.86667" municipality="Voroshilova Peak" name="Vyshe-Ivanovskyi Belok Mt." pageId="9" pageNumber="576" stateProvince="Shyghys Qazaqstan Oblysy">Vyshe-Ivanovskyi Belok Mt.</location>
),
<geoCoordinate id="3FBD33E19B1BA402BB258EB0FB5DFC44" box="[1119,1218,973,1000]" degrees="50" direction="north" minutes="18" orientation="latitude" pageId="9" pageNumber="576" precision="925" value="50.3">50°18’N</geoCoordinate>
,
<geoCoordinate id="3FBD33E19B1BA402BBB48EB0FAB1FC44" box="[1230,1326,973,1000]" degrees="83" direction="east" minutes="52" orientation="longitude" pageId="9" pageNumber="576" precision="925" value="83.86667">83°52’E</geoCoordinate>
, moraine,
<quantity id="9D71F8C39B1BA402BFED8E8CFF71FBA0" box="[151,238,1009,1036]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="576" unit="m" value="2000.0">
<elevation id="D1A4B2159B1BA402BFED8E8CFF71FBA0" box="[151,238,1009,1036]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="576" unit="m" value="2000.0">2000 m</elevation>
</quantity>
,
<date id="2E3773E69B1BA402BF808E8CFEF3FBA0" box="[250,364,1009,1036]" pageId="9" pageNumber="576" value="2015-09-03">
<collectingDate id="3E738A0E9B1BA402BF808E8CFEF3FBA0" box="[250,364,1009,1036]" pageId="9" pageNumber="576" value="2015-09-03">3.09.2015</collectingDate>
</date>
,
<collectorName id="F77C30F09B1BA402BE0C8E8FFDB9FBA7" box="[374,550,1009,1036]" pageId="9" pageNumber="576">A.A. Fomichev</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="129306AD9B1BA402BFBD8968FEDEFBDB" pageId="9" pageNumber="576" type="etymology">
<paragraph id="5A3655269B1BA402BFBD8968FEDEFBDB" blockId="9.[151,1437,901,1900]" pageId="9" pageNumber="576">
<emphasis id="68FD89349B1BA402BFBD8968FED1FB83" bold="true" box="[199,334,1045,1071]" pageId="9" pageNumber="576">Etymology.</emphasis>
The specific name is a patronym given in honor of my friend, Evgeniy Yu. Kulikov (Rubtsovsk,
<collectingCountry id="229E15B69B1BA402BFED8944FF79FBF8" box="[151,230,1081,1108]" name="Russia" pageId="9" pageNumber="576">Russia</collectingCountry>
), who helped in the organization of the field trip to Ivanovskyi Mt. Range, where the
<typeStatus id="8532EB849B1BA402BBEB8947FB53FBF8" box="[1169,1228,1082,1108]" pageId="9" pageNumber="576">types</typeStatus>
of the new species were collected.
</paragraph>
</subSubSection>
<subSubSection id="129306AD9B1BA402BFBD89FCFE3FFAAB" pageId="9" pageNumber="576" type="diagnosis">
<paragraph id="5A3655269B1BA402BFBD89FCFE3FFAAB" blockId="9.[151,1437,901,1900]" pageId="9" pageNumber="576">
<emphasis id="68FD89349B1BA402BFBD89FCFEDDFB37" bold="true" box="[199,322,1153,1179]" pageId="9" pageNumber="576">Diagnosis.</emphasis>
The new species is similar to the widespread
<taxonomicName id="9D892EA59B1BA402BC2A89FFFC55FB37" baseAuthorityName="Clerck" baseAuthorityYear="1757" box="[848,970,1153,1180]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="576" phylum="Arthropoda" rank="species" species="lignaria">
<emphasis id="68FD89349B1BA402BC2A89FFFC55FB37" box="[848,970,1153,1180]" italics="true" pageId="9" pageNumber="576">A. lignaria</emphasis>
</taxonomicName>
in having triangular epigynal fovea (
<emphasis id="68FD89349B1BA402BA0E89FFFA0CFB37" box="[1396,1427,1154,1179]" italics="true" pageId="9" pageNumber="576">Fo</emphasis>
) and quadrangular septal base (
<emphasis id="68FD89349B1BA402BE9689D8FD97FB13" box="[492,520,1189,1215]" italics="true" pageId="9" pageNumber="576">Sb</emphasis>
), but differs from this species by the apical pocket (
<emphasis id="68FD89349B1BA402BB2889DBFBEEFB6C" box="[1106,1137,1190,1216]" italics="true" pageId="9" pageNumber="576">Ap</emphasis>
) with one hood (vs. apical pocket with two hoods) and by distinct apical lips of the epigyne (
<emphasis id="68FD89349B1BA402BC0D89B4FC0FFB4F" box="[887,912,1225,1251]" italics="true" pageId="9" pageNumber="576">Al</emphasis>
) (vs. indistinguishable, cf.
<figureCitation id="C2B249A39B1BA402BBBB89B4FA8FFB4F" box="[1217,1296,1225,1251]" captionStart="FIGURES 63–71" captionStartId="11.[151,264,1861,1885]" captionTargetBox="[151,1436,489,1838]" captionTargetId="figure-147@11.[151,1436,489,1838]" captionTargetPageId="11" captionText="FIGURES 63–71. Epigyne and female internal genitalia of Acantholycosa kulikovi sp. n. (63–65), A. levinae (66–68) and Mongolicosa lizae sp. n. (69–71). 63, 66, 69—intact, ventral; 64, 67, 70—macerated, ventral; 65, 68, 71—macerated, dorsal. Scale bars=0.2 mm. Abbreviations: Al apical lips of the epigyne, Ap apical pocket, Cd copulatory ducts, Fo epigynal fovea, Li lip of the epigyne, Re receptacle, Sa septal base arm, Sb septal base, Ss septal stem." figureDoi="http://doi.org/10.5281/zenodo.5300804" httpUri="https://zenodo.org/record/5300804/files/figure.png" pageId="9" pageNumber="576">Fig. 63</figureCitation>
and fig.
<quantity id="9D71F8C39B1BA402BA0889B4FF32FAAB" metricMagnitude="0" metricUnit="m" metricValue="3.048" pageId="9" pageNumber="576" unit="in" value="120.0">120 in</quantity>
<bibRefCitation id="3E1828D79B1BA402BFCE8990FE0BFAAB" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." box="[180,404,1261,1287]" pageId="9" pageNumber="576" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">
Marusik
<emphasis id="68FD89349B1BA402BE628993FECEFAAB" box="[280,337,1261,1287]" italics="true" pageId="9" pageNumber="576">et al.</emphasis>
2004
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="129306AD9B1BA402BFBD886CFEE7F91B" pageId="9" pageNumber="576" type="description">
<paragraph id="5A3655269B1BA402BFBD886CFDB3F9A8" blockId="9.[151,1437,901,1900]" pageId="9" pageNumber="576">
<emphasis id="68FD89349B1BA402BFBD886CFEC6FA87" bold="true" box="[199,345,1297,1323]" pageId="9" pageNumber="576">Description.</emphasis>
Female (
<typeStatus id="8532EB849B1BA402BEB8886CFDB4FA80" box="[450,555,1297,1324]" pageId="9" pageNumber="576" type="holotype">holotype</typeStatus>
). Total length 8.3. Carapace: 4.2 long, 3.5 wide. General appearance as in
<figureCitation id="C2B249A39B1BA402BA0A886CFF38FAE3" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1832,1856]" captionTargetBox="[157,1436,189,1799]" captionTargetId="figure-33@2.[151,1436,181,1809]" captionTargetPageId="2" captionText="FIGURES 1–9. General appearance of Acantholycosa sayanensis (1–2), A. kulikovi sp. n. (3), A. levinae (4–5), A. sergeevi sp. n. (6), Mongolicosa lizae sp. n. (8–9) and leg I of A. sergeevi sp. n. (7). 1, 4, 6, 8—male, dorsal; 2, 3, 5, 9—female, dorsal; 7—leg I of male, prolateral. Scale bars=2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300794" httpUri="https://zenodo.org/record/5300794/files/figure.png" pageId="9" pageNumber="576">Fig. 3</figureCitation>
. Coloration. Carapace and chelicerae dark brown. Eye field black. Sternum and labium brown. Endites, coxae and palps yellow-brown. Legs yellow-brown with yellow stripes. Femora III–IV with indistinct gray annulation. Abdomen dark gray dorsally, with dark brown cardiac mark. Venter of abdomen yellow-gray. Spinnerets dark gray. Spination of leg I: Fe d1-1-1 p0-0-2 r1-0-1; Pa r1; Ti p1-0-0 r1-0-0 v2-2-2-2-2-2; Mt p0-1-0 r0-1-0 v2-2-0. Leg measurements: I 4.1, 1.8, 3.9, 3.7, 1.65 (15.15); II 4.05, 1.8, 3.7, 3.85, 1.6 (15.0); III 4.0, 1.65, 3.5, 4.8, 1.8 (15.75); IV 4.95, 1.85, 4.5, 6.9, 2.45 (20.65).
</paragraph>
<paragraph id="5A3655269B1BA402BFBD8B70FE38F93F" blockId="9.[151,1437,901,1900]" pageId="9" pageNumber="576">
Epigyne and internal genitalia as in
<figureCitation id="C2B249A39B1BA402BD2E8B70FD4BF984" box="[596,724,1549,1576]" captionStart="FIGURES 63–71" captionStartId="11.[151,264,1861,1885]" captionTargetBox="[151,1436,489,1838]" captionTargetId="figure-147@11.[151,1436,489,1838]" captionTargetPageId="11" captionText="FIGURES 63–71. Epigyne and female internal genitalia of Acantholycosa kulikovi sp. n. (63–65), A. levinae (66–68) and Mongolicosa lizae sp. n. (69–71). 63, 66, 69—intact, ventral; 64, 67, 70—macerated, ventral; 65, 68, 71—macerated, dorsal. Scale bars=0.2 mm. Abbreviations: Al apical lips of the epigyne, Ap apical pocket, Cd copulatory ducts, Fo epigynal fovea, Li lip of the epigyne, Re receptacle, Sa septal base arm, Sb septal base, Ss septal stem." figureDoi="http://doi.org/10.5281/zenodo.5300804" httpUri="https://zenodo.org/record/5300804/files/figure.png" pageId="9" pageNumber="576">Figs 63–65</figureCitation>
. Fovea is very narrow. Septal base bulging fills almost the entire fovea. Septal stem undeveloped. Lips of the epigyne (
<emphasis id="68FD89349B1BA402BC7D8B4FFC81F9E7" box="[775,798,1586,1611]" italics="true" pageId="9" pageNumber="576">Li</emphasis>
) separated by twice their width. Apical pocket as wide as septal base. Receptacles (
<emphasis id="68FD89349B1BA402BEA08B2BFE67F9C3" box="[474,504,1622,1647]" italics="true" pageId="9" pageNumber="576">Re</emphasis>
) clavate, clearly delineated from copulatory ducts (
<emphasis id="68FD89349B1BA402BB3F8B28FBF9F9C3" box="[1093,1126,1621,1647]" italics="true" pageId="9" pageNumber="576">Cd</emphasis>
). Copulatory ducts 2 times shorter than receptacles.
</paragraph>
<paragraph id="5A3655269B1BA402BFBD8BE0FEE7F91B" blockId="9.[151,1437,901,1900]" box="[199,376,1693,1719]" pageId="9" pageNumber="576">Male unknown.</paragraph>
</subSubSection>
<subSubSection id="129306AD9B1BA402BFBD8BBCFEB0F8AC" pageId="9" pageNumber="576" type="distribution">
<paragraph id="5A3655269B1BA402BFBD8BBCFEB0F8AC" blockId="9.[151,1437,901,1900]" pageId="9" pageNumber="576">
<emphasis id="68FD89349B1BA402BFBD8BBCFEFEF977" bold="true" box="[199,353,1729,1755]" pageId="9" pageNumber="576">Distribution.</emphasis>
Known from two closely spaced localities in the Ivanovskyi Mt. Range of the
<collectingRegion id="984D9BC49B1BA402BB9E8BBCFA83F977" box="[1252,1308,1729,1755]" country="Russia" name="Altay" pageId="9" pageNumber="576">Altai</collectingRegion>
Mountains (
<figureCitation id="C2B249A39B1BA402BFE58B98FEBCF8AC" box="[159,291,1765,1792]" captionStart="FIGURES 72–77" captionStartId="13.[151,264,1676,1700]" captionTargetBox="[151,1436,309,1652]" captionTargetId="figure-76@13.[151,1436,309,1652]" captionTargetPageId="13" captionText="FIGURES 72–77. Male of Acantholycosa sayanensis from Borus Mt. Range (72), female of Acantholycosa sp. (A. azyuzini or A. levinae) with egg sac, from Katunskyi Mt. Range (73), habitat of A. sayanensis and A. sergeevi sp. n. in Borus Mt. Range (74), collecting localities of studied species from the Acantholycosa-complex (75–76) and map showing the main mountain systems of South Siberia and adjacent regions (77). Diamond—A. sayanensis, square—A. sidorovi sp. n., triangle—A. kulikovi sp. n., star—A. levinae, circle—A. sergeevi sp. n., inverted triangle—Mongolicosa lizae sp. n. The frame of Fig. 76 refers to the content of Fig. 75." figureDoi="http://doi.org/10.5281/zenodo.5300806" httpUri="https://zenodo.org/record/5300806/files/figure.png" pageId="9" pageNumber="576">Figs 75–76</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="129306AD9B1BA402BFBD8A74FF67F8C0" pageId="9" pageNumber="576" type="discussion">
<paragraph id="5A3655269B1BA402BFBD8A74FF67F8C0" blockId="9.[151,1437,901,1900]" pageId="9" pageNumber="576">
<emphasis id="68FD89349B1BA402BFBD8A74FED0F88F" bold="true" box="[199,335,1801,1827]" pageId="9" pageNumber="576">Comments.</emphasis>
Members of
<taxonomicName id="9D892EA59B1BA402BE928A74FD71F88F" baseAuthorityName="Clerck" baseAuthorityYear="1757" box="[488,750,1801,1827]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="576" phylum="Arthropoda" rank="species" species="lignaria">
<emphasis id="68FD89349B1BA402BE928A74FD71F88F" box="[488,750,1801,1827]" italics="true" pageId="9" pageNumber="576">Acantholycosa lignaria</emphasis>
</taxonomicName>
-group can be confidently recognized by males only. Despite this,
<taxonomicName id="9D892EA59B1BA402BFB48A53FED8F8EB" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[206,327,1837,1864]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="576" phylum="Arthropoda" rank="species" species="kulikovi" status="sp. nov.">
<emphasis id="68FD89349B1BA402BFB48A53FED8F8EB" box="[206,327,1837,1864]" italics="true" pageId="9" pageNumber="576">A. kulikovi</emphasis>
</taxonomicName>
<emphasis id="68FD89349B1BA402BE348A50FE12F8EB" bold="true" box="[334,397,1837,1863]" pageId="9" pageNumber="576">
<taxonomicNameLabel id="73CE344F9B1BA402BE348A50FE12F8EB" box="[334,397,1837,1863]" pageId="9" pageNumber="576" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is herein placed in this group based on strong similarity between its epigyne and those of
<taxonomicName id="9D892EA59B1BA402BAFE8A53FF6EF8C7" baseAuthorityName="Clerck" baseAuthorityYear="1757" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="576" phylum="Arthropoda" rank="species" species="lignaria">
<emphasis id="68FD89349B1BA402BAFE8A53FF6EF8C7" italics="true" pageId="9" pageNumber="576">A. lignaria</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

260
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<mods:title id="C50042FE5ABC5984DB69270BC627199A">New data on the wolf spiders from theAcantholycosa-complex (Araneae: Lycosidae) from the South Siberia</mods:title>
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<emphasis id="68FD89349B1CA405BFED8DEAFEEDFF1E" bold="true" box="[151,370,151,178]" italics="true" pageId="14" pageNumber="581">Mongolicosa lizae</emphasis>
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<figureCitation id="C2B249A39B1CA405BFED8DC6FF65FF79" box="[151,250,187,213]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1832,1856]" captionTargetBox="[157,1436,189,1799]" captionTargetId="figure-33@2.[151,1436,181,1809]" captionTargetPageId="2" captionText="FIGURES 1–9. General appearance of Acantholycosa sayanensis (1–2), A. kulikovi sp. n. (3), A. levinae (4–5), A. sergeevi sp. n. (6), Mongolicosa lizae sp. n. (8–9) and leg I of A. sergeevi sp. n. (7). 1, 4, 6, 8—male, dorsal; 2, 3, 5, 9—female, dorsal; 7—leg I of male, prolateral. Scale bars=2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300794" httpUri="https://zenodo.org/record/5300794/files/figure.png" pageId="14" pageNumber="581">Figs 8–9</figureCitation>
,
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,
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,
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<collectingCounty id="B3572DAA9B1CA405BFED8C7EFF7BFEB1" box="[151,228,259,285]" pageId="14" pageNumber="581">Types.</collectingCounty>
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NNE from
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,
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, mountain stony tundra and scree,
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(
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, 001.8723), together with the
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(
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), same locality, alpine meadow with scree,
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.
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</subSubSection>
<subSubSection id="129306AD9B1CA405BFBD8CCAFD35FE59" pageId="14" pageNumber="581" type="etymology">
<paragraph id="5A3655269B1CA405BFBD8CCAFD35FE59" blockId="14.[151,1437,259,1618]" pageId="14" pageNumber="581">
<emphasis id="68FD89349B1CA405BFBD8CCAFED1FE7D" bold="true" box="[199,334,439,465]" pageId="14" pageNumber="581">Etymology.</emphasis>
The specific name is a matronym in honor of my wife, Elizaveta A. Nepaeva (Barnaul,
<collectingCountry id="229E15B69B1CA405BA3B8CCAFA0FFE7E" box="[1345,1424,439,466]" name="Russia" pageId="14" pageNumber="581">Russia</collectingCountry>
), who helped in spider collection for this project.
</paragraph>
</subSubSection>
<subSubSection id="129306AD9B1CA405BFBD8C82FE1CFD5D" pageId="14" pageNumber="581" type="diagnosis">
<paragraph id="5A3655269B1CA405BFBD8C82FE1CFD5D" blockId="14.[151,1437,259,1618]" pageId="14" pageNumber="581">
<emphasis id="68FD89349B1CA405BFBD8C82FEDDFDB5" bold="true" box="[199,322,511,537]" pageId="14" pageNumber="581">Diagnosis.</emphasis>
<taxonomicName id="9D892EA59B1CA405BE308C82FD85FDB5" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[330,538,511,537]" class="Arachnida" family="Lycosidae" genus="Mongolicosa" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="581" phylum="Arthropoda" rank="species" species="lizae" status="sp. nov.">
<emphasis id="68FD89349B1CA405BE308C82FD85FDB5" box="[330,538,511,537]" italics="true" pageId="14" pageNumber="581">Mongolicosa lizae</emphasis>
</taxonomicName>
<emphasis id="68FD89349B1CA405BD598C82FDFBFDB5" bold="true" box="[547,612,511,537]" pageId="14" pageNumber="581">
<taxonomicNameLabel id="73CE344F9B1CA405BD598C82FDFBFDB5" box="[547,612,511,537]" pageId="14" pageNumber="581" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is most similar to
<taxonomicName id="9D892EA59B1CA405BC478F7DFC50FDB5" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[829,975,511,537]" class="Arachnida" family="Lycosidae" genus="Mongolicosa" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="581" phylum="Arthropoda" rank="species" species="buryatica">
<emphasis id="68FD89349B1CA405BC478F7DFC50FDB5" box="[829,975,511,537]" italics="true" pageId="14" pageNumber="581">M. buryatica</emphasis>
</taxonomicName>
in having a tegular apophysis (
<emphasis id="68FD89349B1CA405BA418F7DFAC6FDB5" box="[1339,1369,512,537]" italics="true" pageId="14" pageNumber="581">Tg</emphasis>
) with arms equal in length to each other, a smoothly curved embolus (
<emphasis id="68FD89349B1CA405BC108F59FC10FD91" box="[874,911,548,573]" italics="true" pageId="14" pageNumber="581">Em</emphasis>
) without a basal spine, and lips of the epigyne (
<emphasis id="68FD89349B1CA405BFDB8F35FF27FDCD" box="[161,184,584,609]" italics="true" pageId="14" pageNumber="581">Li</emphasis>
) the same height as septal base arms (
<emphasis id="68FD89349B1CA405BD108F3AFD19FDCD" box="[618,646,583,609]" italics="true" pageId="14" pageNumber="581">Sa</emphasis>
). The males of the new species can be distinguished from those of
<taxonomicName id="9D892EA59B1CA405BA048F35FE9DFD29" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" class="Arachnida" family="Lycosidae" genus="Mongolicosa" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="581" phylum="Arthropoda" rank="species" species="buryatica">
<emphasis id="68FD89349B1CA405BA048F35FE9DFD29" italics="true" pageId="14" pageNumber="581">M. buryatica</emphasis>
</taxonomicName>
by an elongated terminal apophysis (
<emphasis id="68FD89349B1CA405BDD78F11FD57FD29" box="[685,712,620,645]" italics="true" pageId="14" pageNumber="581">Ta</emphasis>
), which surpasses the embolus tip, and thinner basal part of the embolus (cf.
<figureCitation id="C2B249A39B1CA405BE548FF2FE30FD06" box="[302,431,655,682]" captionStart="FIGURES 44–53" captionStartId="7.[151,264,1794,1818]" captionTargetBox="[163,1427,188,1770]" captionTargetId="figure-15@7.[151,1436,181,1770]" captionTargetPageId="7" captionText="FIGURES 44–53. Tegulum and subtegulum (44–45, 52–53), bulb (46–48) and embolic division (49–51) of Acantholycosa sergeevi sp. n. (44–45) and Mongolicosa lizae sp. n. (46–53). 44, 46, 49, 52—ventral; 45, 47, 50, 53—anterior; 48, 51— retrolateral. Scale bars=0.2 mm. Abbreviations: Aa apical arm of the tegular apophysis, Ba basal arm of the tegular apophysis, Em embolus, Pl palea, Sd sperm duct, Ta terminal apophysis, Tg tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.5300800" httpUri="https://zenodo.org/record/5300800/files/figure.png" pageId="14" pageNumber="581">Figs 46, 50</figureCitation>
and fig.
<quantity id="9D71F8C39B1CA405BD628FF2FD06FD05" box="[536,665,655,681]" metricMagnitude="0" metricUnit="m" metricValue="4.7625" metricValueMax="4.7752" metricValueMin="4.7498" pageId="14" pageNumber="581" unit="in" value="187.5" valueMax="188.0" valueMin="187.0">187–188 in</quantity>
<bibRefCitation id="3E1828D79B1CA405BDD88FF2FC17FD05" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." box="[674,904,655,681]" pageId="14" pageNumber="581" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">
Marusik
<emphasis id="68FD89349B1CA405BC728FEDFCDCFD05" box="[776,835,655,681]" italics="true" pageId="14" pageNumber="581">et al.</emphasis>
2004
</bibRefCitation>
). The females can be distinguished from those of
<taxonomicName id="9D892EA59B1CA405BFCC8FC9FED8FD61" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[182,327,691,717]" class="Arachnida" family="Lycosidae" genus="Mongolicosa" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="581" phylum="Arthropoda" rank="species" species="buryatica">
<emphasis id="68FD89349B1CA405BFCC8FC9FED8FD61" box="[182,327,691,717]" italics="true" pageId="14" pageNumber="581">M. buryatica</emphasis>
</taxonomicName>
by having distinct apical pocket (
<emphasis id="68FD89349B1CA405BDBC8FCEFD7AFD61" box="[710,741,691,717]" italics="true" pageId="14" pageNumber="581">Ap</emphasis>
) composed by two separate hoods (cf.
<figureCitation id="C2B249A39B1CA405BBDA8FCEFB6FFD61" box="[1184,1264,691,717]" captionStart="FIGURES 63–71" captionStartId="11.[151,264,1861,1885]" captionTargetBox="[151,1436,489,1838]" captionTargetId="figure-147@11.[151,1436,489,1838]" captionTargetPageId="11" captionText="FIGURES 63–71. Epigyne and female internal genitalia of Acantholycosa kulikovi sp. n. (63–65), A. levinae (66–68) and Mongolicosa lizae sp. n. (69–71). 63, 66, 69—intact, ventral; 64, 67, 70—macerated, ventral; 65, 68, 71—macerated, dorsal. Scale bars=0.2 mm. Abbreviations: Al apical lips of the epigyne, Ap apical pocket, Cd copulatory ducts, Fo epigynal fovea, Li lip of the epigyne, Re receptacle, Sa septal base arm, Sb septal base, Ss septal stem." figureDoi="http://doi.org/10.5281/zenodo.5300804" httpUri="https://zenodo.org/record/5300804/files/figure.png" pageId="14" pageNumber="581">Fig. 69</figureCitation>
and fig.
<quantity id="9D71F8C39B1CA405BA2F8FCEFA03FD61" box="[1365,1436,691,717]" metricMagnitude="0" metricUnit="m" metricValue="5.4102" pageId="14" pageNumber="581" unit="in" value="213.0">213 in</quantity>
<bibRefCitation id="3E1828D79B1CA405BFED8FAAFEE8FD5D" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." box="[151,375,727,753]" pageId="14" pageNumber="581" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">
Marusik
<emphasis id="68FD89349B1CA405BF868FA5FEABFD5D" box="[252,308,727,753]" italics="true" pageId="14" pageNumber="581">et al.</emphasis>
2004
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="129306AD9B1CA405BFBD8F86FCFCF981" pageId="14" pageNumber="581" type="description">
<paragraph id="5A3655269B1CA405BFBD8F86FDB3FBBE" blockId="14.[151,1437,259,1618]" pageId="14" pageNumber="581">
<emphasis id="68FD89349B1CA405BFBD8F86FEC6FCB9" bold="true" box="[199,345,763,789]" pageId="14" pageNumber="581">Description.</emphasis>
Male (
<typeStatus id="8532EB849B1CA405BED08F86FD8CFCBA" box="[426,531,763,790]" pageId="14" pageNumber="581" type="holotype">holotype</typeStatus>
). Total length 8.2. Carapace: 4.2 long, 3.25 wide. General appearance as in
<figureCitation id="C2B249A39B1CA405BA0A8F86FF38FC95" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1832,1856]" captionTargetBox="[157,1436,189,1799]" captionTargetId="figure-33@2.[151,1436,181,1809]" captionTargetPageId="2" captionText="FIGURES 1–9. General appearance of Acantholycosa sayanensis (1–2), A. kulikovi sp. n. (3), A. levinae (4–5), A. sergeevi sp. n. (6), Mongolicosa lizae sp. n. (8–9) and leg I of A. sergeevi sp. n. (7). 1, 4, 6, 8—male, dorsal; 2, 3, 5, 9—female, dorsal; 7—leg I of male, prolateral. Scale bars=2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300794" httpUri="https://zenodo.org/record/5300794/files/figure.png" pageId="14" pageNumber="581">Fig. 8</figureCitation>
. Coloration. Carapace dark brown, almost black. Eye field black. Sternum dark gray. Labium and endites yellowgray. Chelicerae dark brown. Palps: Fe–Pt dark brown, Ti and cymbium black. Coxae yellow-gray with yellow spots. Legs dark brown dorsally, yellow-gray ventrally. Fe of all legs darker than others segments. Abdomen dark gray dorsally, almost black. Cardiac mark dark brown. Venter of the abdomen gray. Spinnerets black. Spination of leg I: Fe d1-1-1 p0-1-2 r1-1-1; Pa p1 r1; Ti d1-0-1 p1-0-0 r1-0-1 v2-2-2-1-2; Mt p0-1-0 r1-1-0 v2-2-0. Leg measurements: I 3.3, 1.6, 3.0, 3.1, 1.65 (12.65); II 3.3, 1.6, 2.85, 3.15, 1.6 (12.5); III 3.35, 1.55, 2.7, 3.7, 1.6 (12.9); IV 4.15, 1.7, 3.6, 5.55, 2.15 (17.15).
</paragraph>
<paragraph id="5A3655269B1CA405BFBD8966FD3FFB69" blockId="14.[151,1437,259,1618]" pageId="14" pageNumber="581">
Palp as in
<figureCitation id="C2B249A39B1CA405BE468966FE24FB99" box="[316,443,1051,1077]" captionStart="FIGURES 10–19" captionStartId="3.[151,264,1849,1873]" captionTargetBox="[155,1425,188,1810]" captionTargetId="figure-15@3.[151,1436,181,1825]" captionTargetPageId="3" captionText="FIGURES 10–19. Male palp (10–13) and bulb (14–19) of Acantholycosa sergeevi sp. n. (10–11), A. sayanensis (14–15), A. sidorovi sp. n. (16–17), A. levinae (18-19) and Mongolicosa lizae sp. n. (12–13). 10, 12—retrolateral; 11, 13, 14, 16, 18— ventral; 15, 17, 19—anterior. Scale bars=0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300796" httpUri="https://zenodo.org/record/5300796/files/figure.png" pageId="14" pageNumber="581">Figs 12–13</figureCitation>
,
<figureCitation id="C2B249A39B1CA405BEBD8966FD8EFB9A" box="[455,529,1051,1078]" captionStart="FIGURES 44–53" captionStartId="7.[151,264,1794,1818]" captionTargetBox="[163,1427,188,1770]" captionTargetId="figure-15@7.[151,1436,181,1770]" captionTargetPageId="7" captionText="FIGURES 44–53. Tegulum and subtegulum (44–45, 52–53), bulb (46–48) and embolic division (49–51) of Acantholycosa sergeevi sp. n. (44–45) and Mongolicosa lizae sp. n. (46–53). 44, 46, 49, 52—ventral; 45, 47, 50, 53—anterior; 48, 51— retrolateral. Scale bars=0.2 mm. Abbreviations: Aa apical arm of the tegular apophysis, Ba basal arm of the tegular apophysis, Em embolus, Pl palea, Sd sperm duct, Ta terminal apophysis, Tg tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.5300800" httpUri="https://zenodo.org/record/5300800/files/figure.png" pageId="14" pageNumber="581">46–53</figureCitation>
. Tibia and cymbium covered with dense black setae. Tegular apophysis with two arms, of which the apical arm (
<emphasis id="68FD89349B1CA405BE8C8942FD8AFBF5" box="[502,533,1087,1113]" italics="true" pageId="14" pageNumber="581">Aa</emphasis>
) sharply pointed and basal arm (
<emphasis id="68FD89349B1CA405BCF2893DFC38FBF5" box="[904,935,1088,1113]" italics="true" pageId="14" pageNumber="581">Ba</emphasis>
) rounded in ventral view. Palea (
<emphasis id="68FD89349B1CA405BA678942FAA9FBF5" box="[1309,1334,1087,1113]" italics="true" pageId="14" pageNumber="581">Pl</emphasis>
) without developed apophysis but with a small sclerotized part retrolaterally. Terminal apophysis long, thin, and slightly curved. Embolus relatively narrow, almost 3 times longer than wide. Embolic tip rounded. Sperm duct (
<emphasis id="68FD89349B1CA405BA4489FAFAC5FB0D" box="[1342,1370,1159,1185]" italics="true" pageId="14" pageNumber="581">Sd</emphasis>
) runs closer to the inner (dorsal) margin of embolus.
</paragraph>
<paragraph id="5A3655269B1CA405BFBD89B2FAEBFA32" blockId="14.[151,1437,259,1618]" pageId="14" pageNumber="581">
Female. Total length 8.8. Carapace: 4.4 long, 3.55 wide. General appearance as in
<figureCitation id="C2B249A39B1CA405BB2589B2FB3BFB45" box="[1119,1188,1231,1257]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1832,1856]" captionTargetBox="[157,1436,189,1799]" captionTargetId="figure-33@2.[151,1436,181,1809]" captionTargetPageId="2" captionText="FIGURES 1–9. General appearance of Acantholycosa sayanensis (1–2), A. kulikovi sp. n. (3), A. levinae (4–5), A. sergeevi sp. n. (6), Mongolicosa lizae sp. n. (8–9) and leg I of A. sergeevi sp. n. (7). 1, 4, 6, 8—male, dorsal; 2, 3, 5, 9—female, dorsal; 7—leg I of male, prolateral. Scale bars=2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300794" httpUri="https://zenodo.org/record/5300794/files/figure.png" pageId="14" pageNumber="581">Fig. 9</figureCitation>
. Coloration. Carapace dark brown. Sternum, labium and endites yellow-gray. Chelicerae brown. Coxae yellow-gray with yellow spots. Palps brown. Legs yellow-brown. Fe of all legs darker than other segments. Abdomen dark gray dorsally, almost black. Cardiac mark dark brown. Venter of the abdomen yellow-gray. Spinnerets black. Spination of leg I: Fe d1-1-1 p0-1-2 r1-1-1; Ti p1-0-0 r1-0-1 v2-2-2-1-2; Mt p0-1-0 r0-1-0 v2-2-0. Leg measurements: I 3.65, 1.8, 3.3, 3.05, 1.7 (13.5); II 3.7, 1.75, 3.1, 3.1, 1.7 (13.35); III 3.65, 1.65, 2.95, 3.8, 1.7 (13.75); IV 4.6, 1.8, 4.0, 6.0, 2.35 (18.75).
</paragraph>
<paragraph id="5A3655269B1CA405BFBD88DAFCFCF981" blockId="14.[151,1437,259,1618]" pageId="14" pageNumber="581">
Epigyne and internal genitalia as in
<figureCitation id="C2B249A39B1CA405BD2688DAFD42FA6D" box="[604,733,1447,1473]" captionStart="FIGURES 63–71" captionStartId="11.[151,264,1861,1885]" captionTargetBox="[151,1436,489,1838]" captionTargetId="figure-147@11.[151,1436,489,1838]" captionTargetPageId="11" captionText="FIGURES 63–71. Epigyne and female internal genitalia of Acantholycosa kulikovi sp. n. (63–65), A. levinae (66–68) and Mongolicosa lizae sp. n. (69–71). 63, 66, 69—intact, ventral; 64, 67, 70—macerated, ventral; 65, 68, 71—macerated, dorsal. Scale bars=0.2 mm. Abbreviations: Al apical lips of the epigyne, Ap apical pocket, Cd copulatory ducts, Fo epigynal fovea, Li lip of the epigyne, Re receptacle, Sa septal base arm, Sb septal base, Ss septal stem." figureDoi="http://doi.org/10.5281/zenodo.5300804" httpUri="https://zenodo.org/record/5300804/files/figure.png" pageId="14" pageNumber="581">Figs 69–71</figureCitation>
. Fovea (
<emphasis id="68FD89349B1CA405BC4788D5FCC3FA6D" box="[829,860,1448,1473]" italics="true" pageId="14" pageNumber="581">Fo</emphasis>
) pear-shaped, as wide as long. Apical pocket deep. Septal stem (
<emphasis id="68FD89349B1CA405BE4A88B6FED6FA49" box="[304,329,1483,1509]" italics="true" pageId="14" pageNumber="581">Ss</emphasis>
) thickened posteriorly. Septal base (
<emphasis id="68FD89349B1CA405BD8B88B6FC92FA49" box="[753,781,1483,1509]" italics="true" pageId="14" pageNumber="581">Sb</emphasis>
) anchor-shaped, 3 times wider than high, posterior edge triangular. Lips of the epigyne closely spaced. Receptacles (
<emphasis id="68FD89349B1CA405BC39888DFCFEF9A5" box="[835,865,1520,1545]" italics="true" pageId="14" pageNumber="581">Re</emphasis>
) clavate, clearly delineated from copulatory ducts (
<emphasis id="68FD89349B1CA405BFDB8B6EFF5DF981" box="[161,194,1555,1581]" italics="true" pageId="14" pageNumber="581">Cd</emphasis>
). Copulatory ducts as long as receptacles, smoothly curved.
</paragraph>
</subSubSection>
<subSubSection id="129306AD9B1CA405BFBD8B4AFB53F9FE" box="[199,1228,1591,1618]" pageId="14" pageNumber="581" type="distribution">
<paragraph id="5A3655269B1CA405BFBD8B4AFB53F9FE" blockId="14.[151,1437,259,1618]" box="[199,1228,1591,1618]" pageId="14" pageNumber="581">
<emphasis id="68FD89349B1CA405BFBD8B4AFEFEF9FD" bold="true" box="[199,353,1591,1617]" pageId="14" pageNumber="581">Distribution.</emphasis>
Known only from the
<typeStatus id="8532EB849B1CA405BD1A8B45FD0FF9FE" box="[608,656,1592,1618]" pageId="14" pageNumber="581">type</typeStatus>
locality in the Tannu-Ola Mountains (
<figureCitation id="C2B249A39B1CA405BB478B4AFB5FF9FE" box="[1085,1216,1591,1618]" captionStart="FIGURES 72–77" captionStartId="13.[151,264,1676,1700]" captionTargetBox="[151,1436,309,1652]" captionTargetId="figure-76@13.[151,1436,309,1652]" captionTargetPageId="13" captionText="FIGURES 72–77. Male of Acantholycosa sayanensis from Borus Mt. Range (72), female of Acantholycosa sp. (A. azyuzini or A. levinae) with egg sac, from Katunskyi Mt. Range (73), habitat of A. sayanensis and A. sergeevi sp. n. in Borus Mt. Range (74), collecting localities of studied species from the Acantholycosa-complex (75–76) and map showing the main mountain systems of South Siberia and adjacent regions (77). Diamond—A. sayanensis, square—A. sidorovi sp. n., triangle—A. kulikovi sp. n., star—A. levinae, circle—A. sergeevi sp. n., inverted triangle—Mongolicosa lizae sp. n. The frame of Fig. 76 refers to the content of Fig. 75." figureDoi="http://doi.org/10.5281/zenodo.5300806" httpUri="https://zenodo.org/record/5300806/files/figure.png" pageId="14" pageNumber="581">Figs 75–76</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="51AFEFD4228AFDF9111D30EFB3ECB6C5">New data on the wolf spiders from theAcantholycosa-complex (Araneae: Lycosidae) from the South Siberia</mods:title>
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<emphasis id="68FD89349B1EA407BFED8B04FD8FF938" bold="true" box="[151,528,1657,1684]" italics="true" pageId="12" pageNumber="579">Mongolicosa pseudoferruginea</emphasis>
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<emphasis id="68FD89349B1EA407BFED8BC3FE4AF97A" box="[151,469,1726,1750]" italics="true" pageId="12" pageNumber="579">Mongolicosa pseudoferruginea</emphasis>
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group—
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Marusik
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2004: 131
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<emphasis id="68FD89349B1EA407BFED8A79FE8DF8B2" bold="true" box="[151,274,1796,1822]" pageId="12" pageNumber="579">Diagnosis.</emphasis>
Species from this group can be distinguished by the touching lips of the epigyne (
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Marusik
<emphasis id="68FD89349B1EA407BA758A78FAD8F8B2" box="[1295,1351,1796,1822]" italics="true" pageId="12" pageNumber="579">et al.</emphasis>
2004
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<emphasis id="68FD89349B1EA407BFBD8A55FEF8F8EE" bold="true" box="[199,359,1832,1858]" pageId="12" pageNumber="579">Composition.</emphasis>
This species group includes eight species:
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<emphasis id="68FD89349B1EA407BC2D8A54FC75F8EE" box="[855,1002,1832,1858]" italics="true" pageId="12" pageNumber="579">M. buryatica</emphasis>
<bibRefCitation id="3E1828D79B1EA407BC898A55FA03F8EF" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." box="[1011,1436,1832,1859]" pageId="12" pageNumber="579" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">Marusik, Azarkina &amp; Koponen, 2004</bibRefCitation>
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(Eastern Sayan Mountains),
<taxonomicName id="9D892EA59B1EA407BEA18A30FC4FF8CB" authority="Fomichev &amp; Marusik, 2018" authorityName="Fomichev &amp; Marusik" authorityYear="2018" box="[475,976,1868,1895]" class="Arachnida" family="Lycosidae" genus="Mongolicosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="579" phylum="Arthropoda" rank="species" species="cherepanovi">
<emphasis id="68FD89349B1EA407BEA18A30FD14F8CA" box="[475,651,1868,1894]" italics="true" pageId="12" pageNumber="579">M. cherepanovi</emphasis>
<bibRefCitation id="3E1828D79B1EA407BDEE8A31FC4FF8CB" author="Fomichev, A. A. &amp; Marusik, Y. M." box="[660,976,1868,1895]" pageId="12" pageNumber="579" pagination="271 - 284" refId="ref9976" refString="Fomichev, A. A. &amp; Marusik, Y. M. (2018) Five new species of the Acantholycosa - complex (Araneae: Lycosidae) from Mongolia. Zootaxa, 4497 (2), 271 - 284. https: // doi. org / 10.11646 / zootaxa. 4497.2.7" type="journal article" year="2018">Fomichev &amp; Marusik, 2018</bibRefCitation>
</taxonomicName>
(Mongolian
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),
<taxonomicName id="9D892EA59B1EA407BBCE8A30FE49F827" authority="Marusik, Azarkina &amp; Koponen, 2004" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" class="Arachnida" family="Lycosidae" genus="Mongolicosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="579" phylum="Arthropoda" rank="species" species="glupovi">
<emphasis id="68FD89349B1EA407BBCE8A30FAB0F8CA" box="[1204,1327,1868,1894]" italics="true" pageId="12" pageNumber="579">M. glupovi</emphasis>
<bibRefCitation id="3E1828D79B1EA407BA428A31FE49F827" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." pageId="12" pageNumber="579" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">Marusik, Azarkina &amp; Koponen, 2004</bibRefCitation>
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(
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Mountains),
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<emphasis id="68FD89349B1EA407BDC78A0CFC85F826" box="[701,794,1904,1930]" italics="true" pageId="12" pageNumber="579">M. lizae</emphasis>
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<taxonomicNameLabel id="73CE344F9B1EA407BC598A0CFCF9F827" box="[803,870,1905,1931]" pageId="12" pageNumber="579" rank="species">sp. n.</taxonomicNameLabel>
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(Eastern Tannu-Ola Mountains),
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<emphasis id="68FD89349B1EA407BB928A0CFA03F826" box="[1256,1436,1904,1930]" italics="true" pageId="12" pageNumber="579">M. mongolensis</emphasis>
<bibRefCitation id="3E1828D79B1EA407BFED8AE9FDDFF803" author="Marusik, Y. M. &amp; Azarkina, G. N. &amp; Koponen, S." box="[151,576,1940,1967]" pageId="12" pageNumber="579" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. &amp; Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">Marusik, Azarkina &amp; Koponen, 2004</bibRefCitation>
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(Khangai Mountains)
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<emphasis id="68FD89349B1EA407BC3E8AE8FC56F802" box="[836,969,1940,1966]" italics="true" pageId="12" pageNumber="579">M. ozkutuki</emphasis>
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(Mongolian
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),
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<emphasis id="68FD89349B1EA407BF9D8AC4FE4CF87E" box="[231,467,1976,2002]" italics="true" pageId="12" pageNumber="579">M. pseudoferruginea</emphasis>
(Schenkel, 1936)
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(Eastern Tian Shan Mountains) and
<taxonomicName id="9D892EA59B1EA407BB428AC4FF50F85B" authority="Fomichev &amp; Marusik, 2017" authorityName="Fomichev &amp; Marusik" authorityYear="2017" class="Arachnida" family="Lycosidae" genus="Mongolicosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="579" phylum="Arthropoda" rank="species" species="uncia">
<emphasis id="68FD89349B1EA407BB428AC4FB04F87E" box="[1080,1179,1977,2002]" italics="true" pageId="12" pageNumber="579">M. uncia</emphasis>
Fomichev &amp;
<bibRefCitation id="3E1828D79B1EA407BA428AC5FF50F85B" author="Marusik, Y. M. &amp; Omelko, M. M." pageId="12" pageNumber="579" pagination="597 - 600" refId="ref10588" refString="Marusik, Y. M. &amp; Omelko, M. M. (2017) A new species of Acantholycosa baltoroi - group (Araneae: Lycosidae, Pardosinae) from the Russian Far East. Zootaxa, 4232 (4), 597 - 600. https: // doi. org / 10.11646 / zootaxa. 4232.4.12" type="journal article" year="2017">Marusik, 2017</bibRefCitation>
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(Mongolian
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).
</paragraph>
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<emphasis id="68FD89349B1FA406BFBD8DEBFED0FF1C" bold="true" box="[199,335,150,176]" pageId="13" pageNumber="580">Comments.</emphasis>
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Marusik
<emphasis id="68FD89349B1FA406BEC78DEAFE69FF1D" box="[445,502,151,177]" italics="true" pageId="13" pageNumber="580">et al.</emphasis>
(2004)
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split the genus into two groups on the basis of the epigynal shape, without taking into account the morphology of the male palp. This was due to the fact that males are not known for many
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<emphasis id="68FD89349B1FA406BFED8DA2FEB6FF55" box="[151,297,223,249]" italics="true" pageId="13" pageNumber="580">Mongolicosa</emphasis>
species
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; until now, about half of species of this genus are known from females only (
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).
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<emphasis id="68FD89349B1EA407BFED88B6FEACFA4A" bold="true" box="[151,307,1483,1510]" italics="true" pageId="12" pageNumber="579">Mongolicosa</emphasis>
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<emphasis id="68FD89349B1EA407BFED8DEAFE2DFF1D" bold="true" box="[151,434,151,178]" italics="true" pageId="12" pageNumber="579">Acantholycosa sergeevi</emphasis>
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<figureCitation id="C2B249A39B1EA407BFED8DC6FF64FF79" box="[151,251,187,213]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1832,1856]" captionTargetBox="[157,1436,189,1799]" captionTargetId="figure-33@2.[151,1436,181,1809]" captionTargetPageId="2" captionText="FIGURES 1–9. General appearance of Acantholycosa sayanensis (1–2), A. kulikovi sp. n. (3), A. levinae (4–5), A. sergeevi sp. n. (6), Mongolicosa lizae sp. n. (8–9) and leg I of A. sergeevi sp. n. (7). 1, 4, 6, 8—male, dorsal; 2, 3, 5, 9—female, dorsal; 7—leg I of male, prolateral. Scale bars=2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300794" httpUri="https://zenodo.org/record/5300794/files/figure.png" pageId="12" pageNumber="579">Figs 6–7</figureCitation>
,
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,
<figureCitation id="C2B249A39B1EA407BE238DC6FE3DFF79" box="[345,418,187,213]" captionStart="FIGURES 32–43" captionStartId="6.[151,264,1742,1766]" captionTargetBox="[154,1435,189,1719]" captionTargetId="figure-33@6.[151,1437,181,1719]" captionTargetPageId="6" captionText="FIGURES 32–43. Tegulum and subtegulum (32–37), bulb (38–40) and embolic division (41–43) of Acantholycosa sayanensis (32–33), A. sidorovi sp. n. (34–35), A. levinae (36–37) and A. sergeevi sp. n. (38–43). 32, 34, 36, 38, 41—ventral; 33, 35, 37, 39, 42—anterior; 40, 43—retrolateral. Scale bars=0.2 mm. Abbreviations: Aa apical arm of the tegular apophysis, Ba basal arm of the tegular apophysis, Em embolus, Pe extension of the palea, Pl palea, Ta terminal apophysis, Tg tegular apophysis, Ts triangle-shaped projection of the palea." figureDoi="http://doi.org/10.5281/zenodo.5300798" httpUri="https://zenodo.org/record/5300798/files/figure.png" pageId="12" pageNumber="579">38–43</figureCitation>
,
<figureCitation id="C2B249A39B1EA407BED78DC6FE69FF7A" box="[429,502,187,214]" captionStart="FIGURES 44–53" captionStartId="7.[151,264,1794,1818]" captionTargetBox="[163,1427,188,1770]" captionTargetId="figure-15@7.[151,1436,181,1770]" captionTargetPageId="7" captionText="FIGURES 44–53. Tegulum and subtegulum (44–45, 52–53), bulb (46–48) and embolic division (49–51) of Acantholycosa sergeevi sp. n. (44–45) and Mongolicosa lizae sp. n. (46–53). 44, 46, 49, 52—ventral; 45, 47, 50, 53—anterior; 48, 51— retrolateral. Scale bars=0.2 mm. Abbreviations: Aa apical arm of the tegular apophysis, Ba basal arm of the tegular apophysis, Em embolus, Pl palea, Sd sperm duct, Ta terminal apophysis, Tg tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.5300800" httpUri="https://zenodo.org/record/5300800/files/figure.png" pageId="12" pageNumber="579">44–45</figureCitation>
,
<figureCitation id="C2B249A39B1EA407BD7B8DC6FDD7FF79" box="[513,584,187,213]" captionStart="FIGURES 72–77" captionStartId="13.[151,264,1676,1700]" captionTargetBox="[151,1436,309,1652]" captionTargetId="figure-76@13.[151,1436,309,1652]" captionTargetPageId="13" captionText="FIGURES 72–77. Male of Acantholycosa sayanensis from Borus Mt. Range (72), female of Acantholycosa sp. (A. azyuzini or A. levinae) with egg sac, from Katunskyi Mt. Range (73), habitat of A. sayanensis and A. sergeevi sp. n. in Borus Mt. Range (74), collecting localities of studied species from the Acantholycosa-complex (75–76) and map showing the main mountain systems of South Siberia and adjacent regions (77). Diamond—A. sayanensis, square—A. sidorovi sp. n., triangle—A. kulikovi sp. n., star—A. levinae, circle—A. sergeevi sp. n., inverted triangle—Mongolicosa lizae sp. n. The frame of Fig. 76 refers to the content of Fig. 75." figureDoi="http://doi.org/10.5281/zenodo.5300806" httpUri="https://zenodo.org/record/5300806/files/figure.png" pageId="12" pageNumber="579">74–76</figureCitation>
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<emphasis id="68FD89349B1EA407BFED8C7EFF46FEB1" bold="true" box="[151,217,259,285]" pageId="12" pageNumber="579">Type.</emphasis>
<materialsCitation id="EAE15F7B9B1EA407BF9B8C7EFAA1FEED" collectingDate="2020-07-16" collectionCode="ISEA" collectorName="A. A. Fomichev" country="Russia" elevation="1900" location="Borus Mt. Range" pageId="12" pageNumber="579" specimenCount="1" specimenCount-male="1" stateProvince="Krasnoyarsk" typeStatus="holotype">
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<collectingRegion id="984D9BC49B1EA407BE2F8C7EFDAEFEB1" box="[341,561,259,285]" country="Russia" name="Krasnoyarsk" pageId="12" pageNumber="579">Krasnoyarsk Krai</collectingRegion>
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:
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(
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, 001.8660),
<location id="5F5603FD9B1EA407BCD98C79FBF7FEB2" LSID="urn:lsid:plazi:treatment:D220E4309B1EA407BFED8DEAFD64FAD6:5F5603FD9B1EA407BCD98C79FBF7FEB2" box="[931,1128,260,286]" country="Russia" name="Borus Mt. Range" pageId="12" pageNumber="579" stateProvince="Krasnoyarsk">Borus Mt. Range</location>
, Malyi Borus and Bol’shoi Borus (=Poilova) Mountains, 52°47–48’N, 91°31–33’E, scree,
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,
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<collectingDate id="3E738A0E9B1EA407BB7F8C5AFB1AFEED" box="[1029,1157,295,321]" pageId="12" pageNumber="579" value="2020-07-16">16.07.2020</collectingDate>
</date>
,
<collectorName id="F77C30F09B1EA407BBF58C5AFAA1FEED" box="[1167,1342,295,321]" pageId="12" pageNumber="579">A.A. Fomichev</collectorName>
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.
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<emphasis id="68FD89349B1EA407BFBD8C36FED1FEC9" bold="true" box="[199,334,331,357]" pageId="12" pageNumber="579">Etymology.</emphasis>
The specific name is a patronym given in honor of my friend, Vasiliy V. Sergeev (
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,
<collectingCountry id="229E15B69B1EA407BFED8C12FF79FE26" box="[151,230,367,394]" name="Russia" pageId="12" pageNumber="579">Russia</collectingCountry>
), who helped in spider collection and in organization of the field trip to Borus Mt. Range, where the
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of this new species was collected.
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<subSubSection id="129306AD9B1EA407BFBD8CCAFD45FC95" pageId="12" pageNumber="579" type="diagnosis">
<paragraph id="5A3655269B1EA407BFBD8CCAFD45FC95" blockId="12.[151,1437,259,1402]" pageId="12" pageNumber="579">
<emphasis id="68FD89349B1EA407BFBD8CCAFEDDFE7D" bold="true" box="[199,322,439,465]" pageId="12" pageNumber="579">Diagnosis.</emphasis>
The new species is similar to the Siberian species
<taxonomicName id="9D892EA59B1EA407BC0C8CCAFC73FE7D" baseAuthorityName="Marusik" baseAuthorityYear="1993" box="[886,1004,439,465]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="579" phylum="Arthropoda" rank="species" species="sterneri">
<emphasis id="68FD89349B1EA407BC0C8CCAFC73FE7D" box="[886,1004,439,465]" italics="true" pageId="12" pageNumber="579">A. sterneri</emphasis>
</taxonomicName>
as well as
<taxonomicName id="9D892EA59B1EA407BB1C8CCAFB40FE7D" baseAuthorityName="Levi &amp; Levi" baseAuthorityYear="1951" box="[1126,1247,439,465]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="579" phylum="Arthropoda" rank="species" species="solituda">
<emphasis id="68FD89349B1EA407BB1C8CCAFB40FE7D" box="[1126,1247,439,465]" italics="true" pageId="12" pageNumber="579">A. solituda</emphasis>
</taxonomicName>
, from the Rocky Mountains (
<collectingCountry id="229E15B69B1EA407BE5B8CA6FEC0FE59" box="[289,351,475,501]" name="United States of America" pageId="12" pageNumber="579">USA</collectingCountry>
). All three species share the following features: palea (
<emphasis id="68FD89349B1EA407BCA78CA6FC69FE59" box="[989,1014,475,501]" italics="true" pageId="12" pageNumber="579">Pl</emphasis>
) with a prominent extension (
<emphasis id="68FD89349B1EA407BA2C8CA1FAEBFE59" box="[1366,1396,476,501]" italics="true" pageId="12" pageNumber="579">Pe</emphasis>
) in the terminal part and the triangle shaped projection (
<emphasis id="68FD89349B1EA407BD918F7DFC9CFDB5" box="[747,771,512,537]" italics="true" pageId="12" pageNumber="579">Ts</emphasis>
) on its posterior edge, strong terminal apophysis (
<emphasis id="68FD89349B1EA407BA448F7DFAC6FDB5" box="[1342,1369,512,537]" italics="true" pageId="12" pageNumber="579">Ta</emphasis>
) with bifurcate tip and thick, cotton-like pubescence on legs I and II.
<taxonomicName id="9D892EA59B1EA407BCF98F5EFB11FD91" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[899,1166,547,573]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="579" phylum="Arthropoda" rank="species" species="sergeevi" status="sp. nov.">
<emphasis id="68FD89349B1EA407BCF98F5EFB11FD91" box="[899,1166,547,573]" italics="true" pageId="12" pageNumber="579">Acantholycosa sergeevi</emphasis>
</taxonomicName>
<emphasis id="68FD89349B1EA407BBE38F5EFB43FD91" bold="true" box="[1177,1244,547,573]" pageId="12" pageNumber="579">
<taxonomicNameLabel id="73CE344F9B1EA407BBE38F5EFB43FD91" box="[1177,1244,547,573]" pageId="12" pageNumber="579" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs from the both aforementioned species by the more developed and pointed triangle shaped projection on the posterior edge of the palea (cf.
<figureCitation id="C2B249A39B1EA407BE2B8F16FE50FD29" box="[337,463,619,645]" captionStart="FIGURES 32–43" captionStartId="6.[151,264,1742,1766]" captionTargetBox="[154,1435,189,1719]" captionTargetId="figure-33@6.[151,1437,181,1719]" captionTargetPageId="6" captionText="FIGURES 32–43. Tegulum and subtegulum (32–37), bulb (38–40) and embolic division (41–43) of Acantholycosa sayanensis (32–33), A. sidorovi sp. n. (34–35), A. levinae (36–37) and A. sergeevi sp. n. (38–43). 32, 34, 36, 38, 41—ventral; 33, 35, 37, 39, 42—anterior; 40, 43—retrolateral. Scale bars=0.2 mm. Abbreviations: Aa apical arm of the tegular apophysis, Ba basal arm of the tegular apophysis, Em embolus, Pe extension of the palea, Pl palea, Ta terminal apophysis, Tg tegular apophysis, Ts triangle-shaped projection of the palea." figureDoi="http://doi.org/10.5281/zenodo.5300798" httpUri="https://zenodo.org/record/5300798/files/figure.png" pageId="12" pageNumber="579">Figs 41–42</figureCitation>
and fig 3, 7 in
<bibRefCitation id="3E1828D79B1EA407BD048F16FC5FFD29" author="Kronestedt, T. &amp; Marusik, Y. M." box="[638,960,619,645]" pageId="12" pageNumber="579" pagination="63 - 71" refId="ref10150" refString="Kronestedt, T. &amp; Marusik, Y. M. (2002) On Acantholycosa solituda (Levi &amp; Levi) and A. sterneri (Marusik) (Araneae: Lycosidae), a pair of geographically distant allied species. Acta Arachnologica, 51 (1), 63 - 71. https: // doi. org / 10.2476 / asjaa. 51.63" type="journal article" year="2002">Kronestedt &amp; Marusik 2002</bibRefCitation>
). The new species can be differed from
<taxonomicName id="9D892EA59B1EA407BAFE8F16FF71FD05" baseAuthorityName="Marusik" baseAuthorityYear="1993" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="579" phylum="Arthropoda" rank="species" species="sterneri">
<emphasis id="68FD89349B1EA407BAFE8F16FF71FD05" italics="true" pageId="12" pageNumber="579">A. sterneri</emphasis>
</taxonomicName>
by the terminal part of the palea less extended, the embolus (
<emphasis id="68FD89349B1EA407BCEF8FEDFC25FD05" box="[917,954,656,681]" italics="true" pageId="12" pageNumber="579">Em</emphasis>
) directed at 2:30 o’clock (vs. 3:30 o’clock) and by the almost straight anterior edge of the tegular apophysis (
<emphasis id="68FD89349B1EA407BCF18FC9FC36FD61" box="[907,937,692,717]" italics="true" pageId="12" pageNumber="579">Tg</emphasis>
) (vs. triangular, cf.
<figureCitation id="C2B249A39B1EA407BBF58FCEFA8DFD61" box="[1167,1298,691,717]" captionStart="FIGURES 32–43" captionStartId="6.[151,264,1742,1766]" captionTargetBox="[154,1435,189,1719]" captionTargetId="figure-33@6.[151,1437,181,1719]" captionTargetPageId="6" captionText="FIGURES 32–43. Tegulum and subtegulum (32–37), bulb (38–40) and embolic division (41–43) of Acantholycosa sayanensis (32–33), A. sidorovi sp. n. (34–35), A. levinae (36–37) and A. sergeevi sp. n. (38–43). 32, 34, 36, 38, 41—ventral; 33, 35, 37, 39, 42—anterior; 40, 43—retrolateral. Scale bars=0.2 mm. Abbreviations: Aa apical arm of the tegular apophysis, Ba basal arm of the tegular apophysis, Em embolus, Pe extension of the palea, Pl palea, Ta terminal apophysis, Tg tegular apophysis, Ts triangle-shaped projection of the palea." figureDoi="http://doi.org/10.5281/zenodo.5300798" httpUri="https://zenodo.org/record/5300798/files/figure.png" pageId="12" pageNumber="579">Figs 38, 41</figureCitation>
,
<figureCitation id="C2B249A39B1EA407BA5A8FCEFAA3FD61" box="[1312,1340,691,717]" captionStart="FIGURES 44–53" captionStartId="7.[151,264,1794,1818]" captionTargetBox="[163,1427,188,1770]" captionTargetId="figure-15@7.[151,1436,181,1770]" captionTargetPageId="7" captionText="FIGURES 44–53. Tegulum and subtegulum (44–45, 52–53), bulb (46–48) and embolic division (49–51) of Acantholycosa sergeevi sp. n. (44–45) and Mongolicosa lizae sp. n. (46–53). 44, 46, 49, 52—ventral; 45, 47, 50, 53—anterior; 48, 51— retrolateral. Scale bars=0.2 mm. Abbreviations: Aa apical arm of the tegular apophysis, Ba basal arm of the tegular apophysis, Em embolus, Pl palea, Sd sperm duct, Ta terminal apophysis, Tg tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.5300800" httpUri="https://zenodo.org/record/5300800/files/figure.png" pageId="12" pageNumber="579">44</figureCitation>
and fig.
<quantity id="9D71F8C39B1EA407BFED8FA5FF5CFD5D" box="[151,195,727,754]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="12" pageNumber="579" unit="in" value="7.0">7 in</quantity>
<bibRefCitation id="3E1828D79B1EA407BFB18FAAFD91FD5D" author="Kronestedt, T. &amp; Marusik, Y. M." box="[203,526,727,753]" pageId="12" pageNumber="579" pagination="63 - 71" refId="ref10150" refString="Kronestedt, T. &amp; Marusik, Y. M. (2002) On Acantholycosa solituda (Levi &amp; Levi) and A. sterneri (Marusik) (Araneae: Lycosidae), a pair of geographically distant allied species. Acta Arachnologica, 51 (1), 63 - 71. https: // doi. org / 10.2476 / asjaa. 51.63" type="journal article" year="2002">Kronestedt &amp; Marusik 2002</bibRefCitation>
).
<taxonomicName id="9D892EA59B1EA407BD5B8FAAFCB5FD5D" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[545,810,727,753]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="579" phylum="Arthropoda" rank="species" species="sergeevi" status="sp. nov.">
<emphasis id="68FD89349B1EA407BD5B8FAAFCB5FD5D" box="[545,810,727,753]" italics="true" pageId="12" pageNumber="579">Acantholycosa sergeevi</emphasis>
</taxonomicName>
<emphasis id="68FD89349B1EA407BC488FAAFCEDFD5D" bold="true" box="[818,882,727,753]" pageId="12" pageNumber="579">
<taxonomicNameLabel id="73CE344F9B1EA407BC488FAAFCEDFD5D" box="[818,882,727,753]" pageId="12" pageNumber="579" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="9D892EA59B1EA407BBDD8FAAFABEFD5D" baseAuthorityName="Levi &amp; Levi" baseAuthorityYear="1951" box="[1191,1313,727,753]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="579" phylum="Arthropoda" rank="species" species="solituda">
<emphasis id="68FD89349B1EA407BBDD8FAAFABEFD5D" box="[1191,1313,727,753]" italics="true" pageId="12" pageNumber="579">A. solituda</emphasis>
</taxonomicName>
by thinner embolus (in anterior view) and by a straight basal arm (
<emphasis id="68FD89349B1EA407BC758F81FCB1FCB9" box="[783,814,764,789]" italics="true" pageId="12" pageNumber="579">Ba</emphasis>
) of the tegular apophysis (vs. bent posteriorly, cf.
<figureCitation id="C2B249A39B1EA407BA168F86FF29FC95" captionStart="FIGURES 32–43" captionStartId="6.[151,264,1742,1766]" captionTargetBox="[154,1435,189,1719]" captionTargetId="figure-33@6.[151,1437,181,1719]" captionTargetPageId="6" captionText="FIGURES 32–43. Tegulum and subtegulum (32–37), bulb (38–40) and embolic division (41–43) of Acantholycosa sayanensis (32–33), A. sidorovi sp. n. (34–35), A. levinae (36–37) and A. sergeevi sp. n. (38–43). 32, 34, 36, 38, 41—ventral; 33, 35, 37, 39, 42—anterior; 40, 43—retrolateral. Scale bars=0.2 mm. Abbreviations: Aa apical arm of the tegular apophysis, Ba basal arm of the tegular apophysis, Em embolus, Pe extension of the palea, Pl palea, Ta terminal apophysis, Tg tegular apophysis, Ts triangle-shaped projection of the palea." figureDoi="http://doi.org/10.5281/zenodo.5300798" httpUri="https://zenodo.org/record/5300798/files/figure.png" pageId="12" pageNumber="579">Figs 42</figureCitation>
,
<figureCitation id="C2B249A39B1EA407BFBB8E62FF42FC95" box="[193,221,799,825]" captionStart="FIGURES 44–53" captionStartId="7.[151,264,1794,1818]" captionTargetBox="[163,1427,188,1770]" captionTargetId="figure-15@7.[151,1436,181,1770]" captionTargetPageId="7" captionText="FIGURES 44–53. Tegulum and subtegulum (44–45, 52–53), bulb (46–48) and embolic division (49–51) of Acantholycosa sergeevi sp. n. (44–45) and Mongolicosa lizae sp. n. (46–53). 44, 46, 49, 52—ventral; 45, 47, 50, 53—anterior; 48, 51— retrolateral. Scale bars=0.2 mm. Abbreviations: Aa apical arm of the tegular apophysis, Ba basal arm of the tegular apophysis, Em embolus, Pl palea, Sd sperm duct, Ta terminal apophysis, Tg tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.5300800" httpUri="https://zenodo.org/record/5300800/files/figure.png" pageId="12" pageNumber="579">44</figureCitation>
and fig.
<quantity id="9D71F8C39B1EA407BE3B8E62FE17FC95" box="[321,392,799,826]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.27" metricValueMin="1.016" pageId="12" pageNumber="579" unit="in" value="4.5" valueMax="5.0" valueMin="4.0">4–5 in</quantity>
<bibRefCitation id="3E1828D79B1EA407BEF58E62FD51FC95" author="Kronestedt, T. &amp; Marusik, Y. M." box="[399,718,799,825]" pageId="12" pageNumber="579" pagination="63 - 71" refId="ref10150" refString="Kronestedt, T. &amp; Marusik, Y. M. (2002) On Acantholycosa solituda (Levi &amp; Levi) and A. sterneri (Marusik) (Araneae: Lycosidae), a pair of geographically distant allied species. Acta Arachnologica, 51 (1), 63 - 71. https: // doi. org / 10.2476 / asjaa. 51.63" type="journal article" year="2002">Kronestedt &amp; Marusik 2002</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="129306AD9B1EA407BFBD8E3EFE0EFB45" pageId="12" pageNumber="579" type="description">
<paragraph id="5A3655269B1EA407BFBD8E3EFB0EFB9A" blockId="12.[151,1437,259,1402]" pageId="12" pageNumber="579">
<emphasis id="68FD89349B1EA407BFBD8E3EFEC6FCF1" bold="true" box="[199,345,835,861]" pageId="12" pageNumber="579">Description.</emphasis>
Male (
<typeStatus id="8532EB849B1EA407BECA8E3EFD86FCF2" box="[432,537,835,862]" pageId="12" pageNumber="579" type="holotype">holotype</typeStatus>
). Total length 10.3. Carapace: 5.1 long, 4.25 wide. General appearance as in
<figureCitation id="C2B249A39B1EA407BFED8E1AFF40FC2D" box="[151,223,871,897]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1832,1856]" captionTargetBox="[157,1436,189,1799]" captionTargetId="figure-33@2.[151,1436,181,1809]" captionTargetPageId="2" captionText="FIGURES 1–9. General appearance of Acantholycosa sayanensis (1–2), A. kulikovi sp. n. (3), A. levinae (4–5), A. sergeevi sp. n. (6), Mongolicosa lizae sp. n. (8–9) and leg I of A. sergeevi sp. n. (7). 1, 4, 6, 8—male, dorsal; 2, 3, 5, 9—female, dorsal; 7—leg I of male, prolateral. Scale bars=2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300794" httpUri="https://zenodo.org/record/5300794/files/figure.png" pageId="12" pageNumber="579">Fig. 6</figureCitation>
. Coloration. Carapace and chelicerae black. Sternum and labium dark-brown. Endites yellow-gray. Palps black. Coxae yellow-gray with yellow spots. Legs dark brown, almost black. Legs I–III have a thick cotton-like pubescence of curled hairs. Abdomen dark gray dorsally, almost black. Cardiac mark dark brown. Venter of the abdomen yellow-gray. Spinnerets black. Spination of leg I: Fe d1-1-1 p0-0-3 r0-1-1; Pa p1 r1; Ti d1-0-1 p1-0-0 v2- 2-2-2-2-2-2-2; Mt p1-1-0 r1-1-0 v2-2-0. Leg measurements: I 4.6, 2.2, 4.35, 4.15, 1.85 (17.15); II 4.55, 2.15, 4.2, 4.25, 1.95 (17.1); III 4.6, 2.0, 3.85, 5.45, 2.05 (17.95); IV 5.35, 2.05, 4.8, 7.85, 2.55 (22.6).
</paragraph>
<paragraph id="5A3655269B1EA407BFBD8942FC43FB69" blockId="12.[151,1437,259,1402]" pageId="12" pageNumber="579">
Palp as in
<figureCitation id="C2B249A39B1EA407BE478942FE22FBF5" box="[317,445,1087,1113]" captionStart="FIGURES 10–19" captionStartId="3.[151,264,1849,1873]" captionTargetBox="[155,1425,188,1810]" captionTargetId="figure-15@3.[151,1436,181,1825]" captionTargetPageId="3" captionText="FIGURES 10–19. Male palp (10–13) and bulb (14–19) of Acantholycosa sergeevi sp. n. (10–11), A. sayanensis (14–15), A. sidorovi sp. n. (16–17), A. levinae (18-19) and Mongolicosa lizae sp. n. (12–13). 10, 12—retrolateral; 11, 13, 14, 16, 18— ventral; 15, 17, 19—anterior. Scale bars=0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300796" httpUri="https://zenodo.org/record/5300796/files/figure.png" pageId="12" pageNumber="579">Figs 10–11</figureCitation>
,
<figureCitation id="C2B249A39B1EA407BEB38942FD8DFBF5" box="[457,530,1087,1113]" captionStart="FIGURES 32–43" captionStartId="6.[151,264,1742,1766]" captionTargetBox="[154,1435,189,1719]" captionTargetId="figure-33@6.[151,1437,181,1719]" captionTargetPageId="6" captionText="FIGURES 32–43. Tegulum and subtegulum (32–37), bulb (38–40) and embolic division (41–43) of Acantholycosa sayanensis (32–33), A. sidorovi sp. n. (34–35), A. levinae (36–37) and A. sergeevi sp. n. (38–43). 32, 34, 36, 38, 41—ventral; 33, 35, 37, 39, 42—anterior; 40, 43—retrolateral. Scale bars=0.2 mm. Abbreviations: Aa apical arm of the tegular apophysis, Ba basal arm of the tegular apophysis, Em embolus, Pe extension of the palea, Pl palea, Ta terminal apophysis, Tg tegular apophysis, Ts triangle-shaped projection of the palea." figureDoi="http://doi.org/10.5281/zenodo.5300798" httpUri="https://zenodo.org/record/5300798/files/figure.png" pageId="12" pageNumber="579">38–43</figureCitation>
,
<figureCitation id="C2B249A39B1EA407BD648942FDF7FBF6" box="[542,616,1087,1114]" captionStart="FIGURES 44–53" captionStartId="7.[151,264,1794,1818]" captionTargetBox="[163,1427,188,1770]" captionTargetId="figure-15@7.[151,1436,181,1770]" captionTargetPageId="7" captionText="FIGURES 44–53. Tegulum and subtegulum (44–45, 52–53), bulb (46–48) and embolic division (49–51) of Acantholycosa sergeevi sp. n. (44–45) and Mongolicosa lizae sp. n. (46–53). 44, 46, 49, 52—ventral; 45, 47, 50, 53—anterior; 48, 51— retrolateral. Scale bars=0.2 mm. Abbreviations: Aa apical arm of the tegular apophysis, Ba basal arm of the tegular apophysis, Em embolus, Pl palea, Sd sperm duct, Ta terminal apophysis, Tg tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.5300800" httpUri="https://zenodo.org/record/5300800/files/figure.png" pageId="12" pageNumber="579">44–45</figureCitation>
. Tegular apophysis hammer-shaped, with reduced apical arm. Palea with abrupt retrolateral margin. Triangle-shaped projection straight and sharply pointed, bearing conical tubercle on the retrolateral edge. Terminal apophysis massive, subdivided into two lobes. Embolus completely straight. Distal part of the embolus bears a rectangular laminar outgrowth on the anterior edge.
</paragraph>
<paragraph id="5A3655269B1EA407BFBD89B2FE0EFB45" blockId="12.[151,1437,259,1402]" box="[199,401,1231,1257]" pageId="12" pageNumber="579">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="129306AD9B1EA407BFBD898EFB63FAA2" box="[199,1276,1267,1294]" pageId="12" pageNumber="579" type="distribution">
<paragraph id="5A3655269B1EA407BFBD898EFB63FAA2" blockId="12.[151,1437,259,1402]" box="[199,1276,1267,1294]" pageId="12" pageNumber="579">
<emphasis id="68FD89349B1EA407BFBD898EFEFEFAA1" bold="true" box="[199,353,1267,1293]" pageId="12" pageNumber="579">Distribution.</emphasis>
Known only from the
<typeStatus id="8532EB849B1EA407BD1A8989FD0FFAA2" box="[608,656,1268,1294]" pageId="12" pageNumber="579">type</typeStatus>
locality in the Western Sayan Mountains (
<figureCitation id="C2B249A39B1EA407BB16898EFB6FFAA2" box="[1132,1264,1267,1294]" captionStart="FIGURES 72–77" captionStartId="13.[151,264,1676,1700]" captionTargetBox="[151,1436,309,1652]" captionTargetId="figure-76@13.[151,1436,309,1652]" captionTargetPageId="13" captionText="FIGURES 72–77. Male of Acantholycosa sayanensis from Borus Mt. Range (72), female of Acantholycosa sp. (A. azyuzini or A. levinae) with egg sac, from Katunskyi Mt. Range (73), habitat of A. sayanensis and A. sergeevi sp. n. in Borus Mt. Range (74), collecting localities of studied species from the Acantholycosa-complex (75–76) and map showing the main mountain systems of South Siberia and adjacent regions (77). Diamond—A. sayanensis, square—A. sidorovi sp. n., triangle—A. kulikovi sp. n., star—A. levinae, circle—A. sergeevi sp. n., inverted triangle—Mongolicosa lizae sp. n. The frame of Fig. 76 refers to the content of Fig. 75." figureDoi="http://doi.org/10.5281/zenodo.5300806" httpUri="https://zenodo.org/record/5300806/files/figure.png" pageId="12" pageNumber="579">Figs 74–76</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="129306AD9B1EA407BFBD886AFD64FAD6" pageId="12" pageNumber="579" type="discussion">
<paragraph id="5A3655269B1EA407BFBD886AFD64FAD6" blockId="12.[151,1437,259,1402]" pageId="12" pageNumber="579">
<emphasis id="68FD89349B1EA407BFBD886AFED0FA9D" bold="true" box="[199,335,1303,1329]" pageId="12" pageNumber="579">Comments.</emphasis>
This species was found in the same locality and in the same scree patches as
<taxonomicName id="9D892EA59B1EA407BB8D886AFE8FFAF9" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="579" phylum="Arthropoda" rank="species" species="sayanensis">
<emphasis id="68FD89349B1EA407BB8D886AFE8FFAF9" italics="true" pageId="12" pageNumber="579">Acantholycosa sayanensis</emphasis>
</taxonomicName>
(see above). Interestingly,
<taxonomicName id="9D892EA59B1EA407BD3F8846FD5DFAF9" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[581,706,1339,1365]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="579" phylum="Arthropoda" rank="species" species="sergeevi" status="sp. nov.">
<emphasis id="68FD89349B1EA407BD3F8846FD5DFAF9" box="[581,706,1339,1365]" italics="true" pageId="12" pageNumber="579">A. sergeevi</emphasis>
</taxonomicName>
<emphasis id="68FD89349B1EA407BDB08846FC94FAF9" bold="true" box="[714,779,1339,1365]" pageId="12" pageNumber="579">
<taxonomicNameLabel id="73CE344F9B1EA407BDB08846FC94FAF9" box="[714,779,1339,1365]" pageId="12" pageNumber="579" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is much rarer than
<taxonomicName id="9D892EA59B1EA407BC918846FB1BFAF9" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[1003,1156,1339,1365]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="579" phylum="Arthropoda" rank="species" species="sayanensis">
<emphasis id="68FD89349B1EA407BC918846FB1BFAF9" box="[1003,1156,1339,1365]" italics="true" pageId="12" pageNumber="579">A. sayanensis</emphasis>
</taxonomicName>
. The
<typeStatus id="8532EB849B1EA407BBBD8846FAB6FAF9" box="[1223,1321,1339,1365]" pageId="12" pageNumber="579" type="holotype">holotype</typeStatus>
male was collected together with
<specimenCount id="4C8F9EAF9B1EA407BEE58822FDA8FAD5" box="[415,567,1375,1401]" pageId="12" pageNumber="579" type="generic">30 specimens</specimenCount>
of
<taxonomicName id="9D892EA59B1EA407BD268822FD6BFAD5" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" box="[604,756,1375,1401]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="579" phylum="Arthropoda" rank="species" species="sayanensis">
<emphasis id="68FD89349B1EA407BD268822FD6BFAD5" box="[604,756,1375,1401]" italics="true" pageId="12" pageNumber="579">A. sayanensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>