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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1091.78986" ID-GBIF-Dataset="c7d745cb-f0b3-43e7-b541-8107ba4d9941" ID-PMC="PMC9005470" ID-Pensoft-Pub="1313-2970-1091-57" ID-Pensoft-UUID="BE91F5636B7C53138A1ACDB7B85AAA6A" ID-PubMed="35586019" ID-ZooBank="9A83F1CA292E41FCA3217B4719C51E7B" ModsDocID="1313-2970-1091-57" checkinTime="1648741034311" checkinUser="pensoft" docAuthor="Guo, Xiao-Jiang, Cheng, Rui, Jiang, Shan, Xue, Da-Yong &amp; Han, Hong-Xiang" docDate="2022" docId="D212121A7958533DA6406A697BF7B111" docLanguage="en" docName="ZooKeys 1091: 57-98" docOrigin="ZooKeys 1091" docPubDate="2022-03-31" docSource="http://dx.doi.org/10.3897/zookeys.1091.78986" docTitle="Ditrigona quinaria subsp. leucophaea Wilkinson 1968" docType="treatment" docVersion="4" id="BE91F5636B7C53138A1ACDB7B85AAA6A" lastPageNumber="57" masterDocId="BE91F5636B7C53138A1ACDB7B85AAA6A" masterDocTitle="Four new species of Ditrigona Moore (Lepidoptera, Drepanidae) in China and an annotated catalogue" masterLastPageNumber="98" masterPageNumber="57" pageNumber="57" updateTime="1668151582051" updateUser="ExternalLinkService">
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<mods:title>Four new species of Ditrigona Moore (Lepidoptera, Drepanidae) in China and an annotated catalogue</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Guo, Xiao-Jiang</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; College of Life Science, Hebei University, Baoding, Hebei 071000, China</mods:affiliation>
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<mods:namePart>Cheng, Rui</mods:namePart>
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<mods:namePart>Jiang, Shan</mods:namePart>
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<mods:namePart>Xue, Da-Yong</mods:namePart>
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<mods:namePart>Han, Hong-Xiang</mods:namePart>
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<taxonomicName LSID="D212121A-7958-533D-A640-6A697BF7B111" authority="Wilkinson, 1968" authorityName="Wilkinson" authorityYear="1968" class="Insecta" family="Drepanidae" genus="Ditrigona" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ditrigona quinaria subsp. leucophaea" order="Lepidoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="subSpecies" species="quinaria" subSpecies="leucophaea">Ditrigona quinaria leucophaea Wilkinson, 1968</taxonomicName>
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<figureCitation captionStart="Figures 127" captionStartId="F1" captionText="Figures 1 - 27. Adults of Ditrigona 1 D. derocina, male 2 D. diana, male 3 D. crystalla, holotype, male 4 D. spilota, male, ZFMK 5 D. furvicosta, male 6 D. jardanaria, male 7 D. media, paratype, male, ZFMK 8 D. sericea, male 9 D. sericea, male (Auzatella pentesticha Chu &amp; Wang, allotype) 10 D. pentesticha, holotype, female 11 D. q. erminea, holotype, male, ZFMK 12 D. q. spodia, holotype, male, ZFMK 13 D. q. leucophaea, holotype, male, ZFMK 14 D. obliquilinea thibetaria, male 15 D. triangularia, lectotype, male 16 D. uniuncusa, male 17 D. tenuiata, holotype, male 18 D. regularis, male 19, 20 D. sinespina sp. nov., 19 holotype, male 20 paratype, female 21, 22 D. parva sp. nov., 21 holotype, male 22 paratype, female 23 D. concava sp. nov., holotype, male 24 D. titana, holotype, male, ZFMK 25 D. pomenaria male 26 D. polyobotaria, holotype, female, ZFMK 27 D. typhodes, male. Scale bar: 1 cm." figureDoi="10.3897/zookeys.1091.78986.figures1-27" httpUri="https://binary.pensoft.net/fig/666820" pageId="0" pageNumber="57">Fig. 13</figureCitation>
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<taxonomicName authorityName="Wilkinson" authorityYear="1968" class="Insecta" family="Drepanidae" genus="Ditrigona" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ditrigona quinaria subsp. leucophaea" order="Lepidoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="subSpecies" species="quinaria" subSpecies="leucophaea">Ditrigona quinaria leucophaea</taxonomicName>
Wilkinson, 1968: 443. Holotype ♂, China: Tibet [Sichuan], Batang (ZFMK).
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<subSubSection pageId="0" pageNumber="57" type="materials_examined">
<paragraph pageId="0" pageNumber="57">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="57">
<materialsCitation collectingDate="1936-05-16" collectorName="H. Hoene, No." country="China" county="Batang" elevation="2800" location="Yantze" municipality="Im Tal" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="holotype">
<emphasis bold="true" pageId="0" pageNumber="57">
<collectingCountry name="China">China</collectingCountry>
:
<collectingRegion country="China" name="Sichuan">Sichuan</collectingRegion>
</emphasis>
:
<specimenCount type="male">1♂</specimenCount>
(ZFMK),
<typeStatus>holotype</typeStatus>
,
<collectingCounty>Batang</collectingCounty>
(
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),
<collectingMunicipality>Im Tal</collectingMunicipality>
dea
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, ca.
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.8" unit="m" value="2800.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.8" unit="m" value="2800.0">2800 m</elevation>
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,
<collectingDate value="1936-05-16">16.V.1936</collectingDate>
,
<collectorName>
H.
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</collectorName>
,
<taxonomicName class="Insecta" family="Drepanidae" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="family">Drepanidae</taxonomicName>
genitalia slide
<collectorName>No.</collectorName>
1486, moth photograph examined
</materialsCitation>
.
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<paragraph pageId="0" pageNumber="57">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="57">China (Sichuan).</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="A14470100812FD48BD3CD246932867F6" pageId="0" pageNumber="6604">FLO; FAI*; GRA*; SMG</paragraph>
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Occasional Migrant; Occasional Wintering.
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<mods:identifier id="10AFB8F09B5E4C12ABB4277A2DD7A9DA" type="CLB-Dataset">29772</mods:identifier>
<mods:identifier id="6898B133523A788FC40A17B7003AB297" type="DOI">10.11646/zootaxa.4442.1.1</mods:identifier>
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<mods:identifier id="BCC4C5B30FF82348BD8B105943C3E959" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="638D3A51E20D082BCF2DB60ADCB5950B" type="Zenodo-Dep">1302713</mods:identifier>
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<subSection id="33348ED92A46D840FF3E7A671A9FB1C6" pageId="7" pageNumber="8" type="key">
<treatment id="D21226242A46D840FF3E7A671A9FB1C6" ID-DOI="http://doi.org/10.5281/zenodo.5950843" ID-GBIF-Taxon="146043490" ID-Zenodo-Dep="5950843" LSID="urn:lsid:plazi:treatment:D21226242A46D840FF3E7A671A9FB1C6" httpUri="http://treatment.plazi.org/id/D21226242A46D840FF3E7A671A9FB1C6" lastPageNumber="8" pageId="7" pageNumber="8">
<subSubSection id="12A1C4B92A46D840FF3E7A671D98B5A7" box="[151,667,871,898]" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="5A0497322A46D840FF3E7A671D98B5A7" blockId="7.[151,667,871,898]" box="[151,667,871,898]" pageId="7" pageNumber="8">
<heading id="014C205E2A46D840FF3E7A671D98B5A7" bold="true" box="[151,667,871,898]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<emphasis id="68CF4B202A46D840FF3E7A671D98B5A7" bold="true" box="[151,667,871,898]" pageId="7" pageNumber="8">
Key to the Asiatic
<taxonomicName id="9DBBECB12A46D840FED57A681EE2B5A4" ID-CoL="8KVFM" authorityName="Latreille" authorityYear="1824" box="[380,481,871,897]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A46D840FED57A681EE2B5A4" bold="true" box="[380,481,871,897]" italics="true" pageId="7" pageNumber="8">Calleida</emphasis>
</taxonomicName>
species groups
</emphasis>
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</paragraph>
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<subSubSection id="12A1C4B92A46D840FF3E7AA11A9FB1C6" pageId="7" pageNumber="8" type="key">
<key id="2E2AC0032A46D840FF3E7AA11A9FB1C6" pageId="7" pageNumber="8">
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<paragraph id="5A0497322A46D840FF3E7AA11A98B5E6" blockId="7.[151,1436,942,2019]" box="[151,1435,942,963]" pageId="7" pageNumber="8">
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1. Abdominal sternum VII with two or more setae on each side in males (
<figureCitation id="C2808BB72A46D840FC207AA11CE3B5E6" box="[905,992,942,963]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="7" pageNumber="8">figs. 23</figureCitation>
), three or more setae in females (
<figureCitation id="C2808BB72A46D840FAB77AA11A77B5E6" box="[1310,1396,942,963]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="7" pageNumber="8">figs. 56</figureCitation>
). 2
</keyLead>
</paragraph>
<paragraph id="5A0497322A46D840FF3E7AC31A9FB5C4" blockId="7.[151,1436,942,2019]" box="[151,1436,972,993]" pageId="7" pageNumber="8">
<keyLead id="E14A34072A46D840FF3E7AC31A9FB5C4" box="[151,1436,972,993]" pageId="7" pageNumber="8">
- Abdominal sternum VII with one seta on each side in males (
<figureCitation id="C2808BB72A46D840FC827AC31C5DB5C4" box="[811,862,972,993]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="7" pageNumber="8">fig. 1</figureCitation>
), two setae in females (
<figureCitation id="C2808BB72A46D840FBEA7AC31B74B5C4" box="[1091,1143,972,993]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="7" pageNumber="8">fig. 4</figureCitation>
)...................... 3
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</paragraph>
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<keyStep id="E14F8F972A46D840FF3E7AE61A9FB273" pageId="7" pageNumber="8">
<paragraph id="5A0497322A46D840FF3E7AE61A9FB23E" blockId="7.[151,1436,942,2019]" pageId="7" pageNumber="8">
<keyLead id="E14A34072A46D840FF3E7AE61A9FB23E" pageId="7" pageNumber="8">
2. Pronotum coarsely punctate (
<figureCitation id="C2808BB72A46D840FE537AE61D3FB5DB" box="[506,572,1001,1022]" captionStart="FIGURES 2229" captionStartId="13.[151,264,1822,1844]" captionTargetBox="[158,1420,311,1777]" captionTargetId="figure@13.[151,1436,301,1801]" captionTargetPageId="13" captionText="FIGURES 2229. Calleida puncticollis sp. n. 22 habitus and labels of holotype (male, Hubei, IZAS), scale bar = 2.0 mm; 23 habitus of paratype (female, Shaanxi, IZAS), scale bar = 2.0 mm; 24 pronotum (Shaanxi, China), [coarsely punctate], scale bar = 0.5 mm; 25 ovipositor (Hubei, China), scale bar = 0.1 mm; 26 female reproductive tract (Hubei, China), scale bar = 0.5 mm; 27 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm; 28 left and right parameres of aedeagus (holotype), scale bar = 0.2 mm; 29 male sternum IX (holotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302721/files/figure.png" pageId="7" pageNumber="8">fig. 24</figureCitation>
). Male with abdominal sternum VII not notched (
<figureCitation id="C2808BB72A46D840FBB17AE61B4EB5DB" box="[1048,1101,1001,1022]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="7" pageNumber="8">fig. 3</figureCitation>
); endophallus of aedeagus without chitinized copulatory pieces (
<figureCitation id="C2808BB72A46D840FE527D091D38B23E" box="[507,571,1030,1051]" captionStart="FIGURES 2229" captionStartId="13.[151,264,1822,1844]" captionTargetBox="[158,1420,311,1777]" captionTargetId="figure@13.[151,1436,301,1801]" captionTargetPageId="13" captionText="FIGURES 2229. Calleida puncticollis sp. n. 22 habitus and labels of holotype (male, Hubei, IZAS), scale bar = 2.0 mm; 23 habitus of paratype (female, Shaanxi, IZAS), scale bar = 2.0 mm; 24 pronotum (Shaanxi, China), [coarsely punctate], scale bar = 0.5 mm; 25 ovipositor (Hubei, China), scale bar = 0.1 mm; 26 female reproductive tract (Hubei, China), scale bar = 0.5 mm; 27 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm; 28 left and right parameres of aedeagus (holotype), scale bar = 0.2 mm; 29 male sternum IX (holotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302721/files/figure.png" pageId="7" pageNumber="8">fig. 27</figureCitation>
).............................................. 2.
<taxonomicName id="9DBBECB12A46D840FB2E7D091A10B23E" ID-CoL="8M9ZJ" box="[1159,1299,1030,1051]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="68CF4B202A46D840FB2E7D091A10B23E" box="[1159,1299,1030,1051]" italics="true" pageId="7" pageNumber="8">C. puncticollis</emphasis>
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species group
</keyLead>
</paragraph>
<paragraph id="5A0497322A46D840FF3E7D2B1A9FB273" blockId="7.[151,1436,942,2019]" pageId="7" pageNumber="8">
<keyLead id="E14A34072A46D840FF3E7D2B1A9FB273" pageId="7" pageNumber="8">
- Pronotum very finely punctate or impunctate. Male with notched abdominal sternum VII (
<figureCitation id="C2808BB72A46D840FBF77D2B1B95B21C" box="[1118,1174,1060,1081]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="7" pageNumber="8">fig. 2</figureCitation>
); endophallus of aedeagus with one or a pair of copulatory pieces............................................. 3.
<taxonomicName id="9DBBECB12A46D840FB267D4E1A10B273" ID-CoL="5WTZV" authorityName="Heller in Mindanao" authorityYear="1921" box="[1167,1299,1089,1110]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="discoidalis">
<emphasis id="68CF4B202A46D840FB267D4E1A10B273" box="[1167,1299,1089,1110]" italics="true" pageId="7" pageNumber="8">C. discoidalis</emphasis>
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species group
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<keyStep id="E14F8F972A46D840FF3E7D511A98B2B4" pageId="7" pageNumber="8">
<paragraph id="5A0497322A46D840FF3E7D511A98B256" blockId="7.[151,1436,942,2019]" box="[151,1435,1118,1139]" pageId="7" pageNumber="8">
<keyLead id="E14A34072A46D840FF3E7D511A98B256" box="[151,1435,1118,1139]" pageId="7" pageNumber="8">
3. Anterior angles of pronotum without setae (
<figureCitation id="C2808BB72A46D840FD2D7D511DC6B256" box="[644,709,1118,1139]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="7" pageNumber="8">fig. 14</figureCitation>
)........................................................... 4
</keyLead>
</paragraph>
<paragraph id="5A0497322A46D840FF3E7D731A98B2B4" blockId="7.[151,1436,942,2019]" box="[151,1435,1148,1169]" pageId="7" pageNumber="8">
<keyLead id="E14A34072A46D840FF3E7D731A98B2B4" box="[151,1435,1148,1169]" pageId="7" pageNumber="8">
- Anterior angles of pronotum each with some, erect setae (
<figureCitation id="C2808BB72A46D840FCA27D731C74B2B4" box="[779,887,1148,1169]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="7" pageNumber="8">figs. 1113</figureCitation>
)............................................ 8
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<paragraph id="5A0497322A46D840FF3E7D961A9FB2EE" blockId="7.[151,1436,942,2019]" pageId="7" pageNumber="8">
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4. Outer apical angle of elytra acute, forming sharp tooth (
<figureCitation id="C2808BB72A46D840FD5E7D961C3BB28B" box="[759,824,1177,1198]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="7" pageNumber="8">fig. 15</figureCitation>
). Median lobe of aedeagus in lateral aspect with a distinct, very prominent ventral lobe (
<figureCitation id="C2808BB72A46D840FE617DB91D10B2EE" box="[456,531,1206,1227]" captionStart="FIGURES 4447" captionStartId="23.[151,263,1096,1118]" captionTargetBox="[179,1384,197,1065]" captionTargetId="figure@23.[151,1436,193,1074]" captionTargetPageId="23" captionText="FIGURES 4447. Calleida terminata Waterhouse. 44 habitus and labels of lectotype (male, Sarawak, Malaysia, NHML), scale bar = 2.0 mm; 45 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (lectotype), scale bar = 0.5 mm; 46 left and right parameres of aedeagus (lectotype), scale bar = 0.2 mm; 47 male sternum IX (lectotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302733/files/figure.png" pageId="7" pageNumber="8">figs. 45</figureCitation>
,
<figureCitation id="C2808BB72A46D840FDB47DB91D34B2EE" box="[541,567,1206,1227]" captionStart="FIGURES 4855" captionStartId="26.[151,264,1681,1703]" captionTargetBox="[156,1417,287,1659]" captionTargetId="figure@26.[151,1436,265,1660]" captionTargetPageId="26" captionText="FIGURES 4855. Calleida corporaali Andrewes. 48 habitus (female, Sabah, Malaysia), scale bar = 2.0 mm; 49 habitus and labels of lectotype for C. corporaali Andrewes (male, Sumatra, NHML), scale bar = 2.0 mm; 50 habitus and labels of lectotype for C. lieftincki Louwerens (female, Borneo, NNML), scale bar = 2.0 mm; 51 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (lectotype), scale bar = 0.5 mm; 52 left and right parameres of aedeagus (lectotype), scale bar = 0.2 mm; 53 male sternum IX (lectotype), scale bar = 0.5 mm; 54 ovipositor (Sabah, Malaysia), scale bar = 0.2 mm; 55 female reproductive tract (Sabah, Malaysia), scale bar = 0.5 mm; abbreviations: sp spermatheca, pd spermathecal pedicel, gd spermathecal gland duct, ga glandular area, at atrium, bpl basal sclerotized plate." httpUri="https://zenodo.org/record/1302735/files/figure.png" pageId="7" pageNumber="8">51</figureCitation>
); abdominal sternum VII deeply notched in males........ 5.
<taxonomicName id="9DBBECB12A46D840FB307DB91A11B2EE" ID-CoL="PLBR" box="[1177,1298,1206,1227]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="terminata">
<emphasis id="68CF4B202A46D840FB307DB91A11B2EE" box="[1177,1298,1206,1227]" italics="true" pageId="7" pageNumber="8">C. terminata</emphasis>
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species group
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<paragraph id="5A0497322A46D840FF3E7DDB1A98B323" blockId="7.[151,1436,942,2019]" pageId="7" pageNumber="8">
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- Outer apical angle of elytra square or obtusely rounded, not forming tooth (
<figureCitation id="C2808BB72A46D840FC677DDB1B43B2CC" box="[974,1088,1236,1257]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="7" pageNumber="8">figs. 1618</figureCitation>
). Median lobe of aedeagus without prominent ventral lobe; abdominal sternum VII shallowly notched in males....................................... 5
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<paragraph id="5A0497322A46D840FF3E7C011A9FB37B" blockId="7.[151,1436,942,2019]" pageId="7" pageNumber="8">
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5. Pronotal disc and all parts of legs piceous to black (
<figureCitation id="C2808BB72A46D840FD7E7C011C4BB306" box="[727,840,1294,1315]" captionStart="FIGURES 6776" captionStartId="36.[151,264,213,235]" captionText="FIGURES 6776. Calleida doriae Bates. 67 habitus and labels of lectotype (female, Bhamo, Myanmar, MSNG), scale bar = 2.0 mm; 68 habitus of paralectotype (male, Assam, India, MNHN), scale bar = 2.0 mm; 69 habitus (male, Yunnan, China), scale bar = 2.0 mm; 70 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (Yunnan, China), scale bar = 0.5 mm; 71 left and right parameres of aedeagus (Yunnan, China), scale bar = 0.2 mm; 72 male sternum IX (Yunnan, China), scale bar = 0.5 mm; 73 ovipositor (Yunnan, China), scale bar = 0.2 mm; 74 female reproductive tract (Yunnan, China), scale bar = 0.5 mm; 75 median lobe of aedeagus, ventral view (paralectotype, Assam), scale bar = 0.5 mm; 76 median lobe of aedeagus, ventral view (Guangxi, China), scale bar = 0.5 mm." pageId="7" pageNumber="8">figs. 6769</figureCitation>
), sometimes fourth and fifth tarsomeres dark brown (
<figureCitation id="C2808BB72A46D840FAE47C011A8DB306" box="[1357,1422,1294,1315]" captionStart="FIGURES 7778" captionStartId="36.[151,264,428,450]" captionText="FIGURES 7778. Calleida rapax Andrewes. 77 median lobe of aedeagus, ventral view (paratype, Landsdowne, Inida, NHML), scale bar = 0.5 mm; 78 habitus and labels of holotype (male, Dehra Dun, India, NHML), scale bar = 2.0 mm." pageId="7" pageNumber="8">fig. 78</figureCitation>
). Species larger sized (L:
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). Median lobe of aedeagus markedly concaved ventrally, without ventral ridges or lobes................................................................................. 7.
<taxonomicName id="9DBBECB12A46D840FB117C461A10B37B" ID-CoL="68TZV" authorityName="Bates Lectotypus" authorityYear="1892" box="[1208,1299,1353,1374]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="doriae">
<emphasis id="68CF4B202A46D840FB117C461A10B37B" box="[1208,1299,1353,1374]" italics="true" pageId="7" pageNumber="8">C. doriae</emphasis>
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species group
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- Pronotal disc and legs usually reddish yellow to brown, if legs partly dark, at least all tarsomeres reddish brown (
<figureCitation id="C2808BB72A46D840FAB57C691A5CB35E" box="[1308,1375,1382,1403]" captionStart="FIGURES 6166" captionStartId="31.[151,264,1853,1875]" captionTargetBox="[159,1432,203,1814]" captionTargetId="figure@31.[151,1436,193,1832]" captionTargetPageId="31" captionText="FIGURES 6166. Calleida borneensis sp. n. 61 habitus and labels of holotype (male, Sabah, Malaysia, NHML), scale bar = 2.0 mm; 62 habitus of paratype (male, Sarawak, Malaysia, NHML), scale bar = 2.0 mm; 63 female reproductive tract (Sabah, Malaysia), scale bar = 0.5 mm; abbreviations: sp spermatheca, pd spermathecal pedicel, gd spermathecal gland duct, ga glandular area, at atrium, bpj spermathecal basal projection, apt pedicel apical protuberance; 64 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm; 65 left and right parameres of aedeagus (holotype), scale bar = 0.2 mm; 66 ovipositor (Sabah, Malaysia), scale bar = 0.2 mm." httpUri="https://zenodo.org/record/1302741/files/figure.png" pageId="7" pageNumber="8">fig. 62</figureCitation>
). Species smaller sized (L:
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). Median lobe of aedeagus simple or strongly ridged ventrally......................6
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<paragraph id="5A0497322A46D840FF3E7CAE1A9FB3F6" blockId="7.[151,1436,942,2019]" pageId="7" pageNumber="8">
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6. Elytral apex strongly curved, outer apical angle markedly widened (
<figureCitation id="C2808BB72A46D840FCC27CAE1CAFB393" box="[875,940,1441,1462]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="7" pageNumber="8">fig. 16</figureCitation>
). Median lobe of aedeagus with apical lamina elongate, widened and ax-shaped (
<figureCitation id="C2808BB72A46D840FDAA7CB11D45B3F6" box="[515,582,1470,1491]" captionStart="FIGURES 6166" captionStartId="31.[151,264,1853,1875]" captionTargetBox="[159,1432,203,1814]" captionTargetId="figure@31.[151,1436,193,1832]" captionTargetPageId="31" captionText="FIGURES 6166. Calleida borneensis sp. n. 61 habitus and labels of holotype (male, Sabah, Malaysia, NHML), scale bar = 2.0 mm; 62 habitus of paratype (male, Sarawak, Malaysia, NHML), scale bar = 2.0 mm; 63 female reproductive tract (Sabah, Malaysia), scale bar = 0.5 mm; abbreviations: sp spermatheca, pd spermathecal pedicel, gd spermathecal gland duct, ga glandular area, at atrium, bpj spermathecal basal projection, apt pedicel apical protuberance; 64 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm; 65 left and right parameres of aedeagus (holotype), scale bar = 0.2 mm; 66 ovipositor (Sabah, Malaysia), scale bar = 0.2 mm." httpUri="https://zenodo.org/record/1302741/files/figure.png" pageId="7" pageNumber="8">fig. 64</figureCitation>
).............................................. 6.
<taxonomicName id="9DBBECB12A46D840FB267CB11A10B3F6" ID-CoL="8M5HW" box="[1167,1299,1470,1491]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis id="68CF4B202A46D840FB267CB11A10B3F6" box="[1167,1299,1470,1491]" italics="true" pageId="7" pageNumber="8">C. borneensis</emphasis>
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species group
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<paragraph id="5A0497322A46D840FF3E7CD31A98B02B" blockId="7.[151,1436,942,2019]" pageId="7" pageNumber="8">
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- Elytral apex straight or weakly curved, outer apical angle not or slightly widened (
<figureCitation id="C2808BB72A46D840FC507CD31B6AB3D4" box="[1017,1129,1500,1521]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="7" pageNumber="8">figs. 1718</figureCitation>
). Median lobe of aedeagus with apical lamina short and simple........................................................................... 7
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7. Body size generally greater than
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; elytra with an indistinct bluish purple patch on disc; metacoxae with few accessory fine setae. Median lobe of aedeagus peculiar in form, irregularly explanate and ridged ventrally...............................................................................................8.
<taxonomicName id="9DBBECB12A46D840FBE17F5E1BD7B043" ID-CoL="5WTZW" authorityName="Dejean" authorityYear="1831" box="[1096,1236,1617,1638]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="chloroptera">
<emphasis id="68CF4B202A46D840FBE17F5E1BD7B043" box="[1096,1236,1617,1638]" italics="true" pageId="7" pageNumber="8">C. chloroptera</emphasis>
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species group (pars)
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- Body size generally smaller than
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; elytra completely metallic greenish, or disc with reddish patch; metacoxae only with 3 or 4 primary setae. Median lobe of aedeagus simple................................. 9.
<taxonomicName id="9DBBECB12A46D840FB2E7F831A10B084" ID-CoL="PLBL" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[1159,1299,1676,1697]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="splendidula">
<emphasis id="68CF4B202A46D840FB2E7F831A10B084" box="[1159,1299,1676,1697]" italics="true" pageId="7" pageNumber="8">C. splendidula</emphasis>
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8. Dorsal surface densely and evenly pubescent (
<figureCitation id="C2808BB72A46D840FD357FA61C0EB09B" box="[668,781,1705,1726]" captionStart="FIGURES 3641" captionStartId="18.[151,264,1697,1719]" captionTargetBox="[184,1416,280,1673]" captionTargetId="figure@18.[151,1436,241,1676]" captionTargetPageId="18" captionText="FIGURES 3641. Calleida excelsa Bates. 36 habitus and labels of lectotype (female, Carin Chebà, Myanmar, MSNG), scale bar = 2.0 mm; 37 habitus of paralectotype (female, Senmigion, Myanmar, MNHN), scale bar = 2.0 mm; 38 habitus (female, Yunnan, China), scale bar = 2.0 mm; 39 female reproductive tract (Chiang Mai, Thailand), scale bar = 0.5 mm; 40 ovipositor (Yunnan, China), scale bar = 0.2 mm; 41 apex of immature aedeagus, left-lateral view, scale bar = 0.2 mm." httpUri="https://zenodo.org/record/1302727/files/figure.png" pageId="7" pageNumber="8">figs. 3638</figureCitation>
). Elytra with wide reddish patch on disc............................................................................................... 4.
<taxonomicName id="9DBBECB12A46D840FBC37FC91BD7B0FE" ID-CoL="5X83Q" authorityName="Chaudoir Lectotypus" authorityYear="1872" box="[1130,1236,1734,1755]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lativittis">
<emphasis id="68CF4B202A46D840FBC37FC91BD7B0FE" box="[1130,1236,1734,1755]" italics="true" pageId="7" pageNumber="8">C. lativittis</emphasis>
</taxonomicName>
species group (pars)
</keyLead>
</paragraph>
<paragraph id="5A0497322A46D840FF3E7FEB1A9FB0DC" blockId="7.[151,1436,942,2019]" box="[151,1436,1764,1785]" pageId="7" pageNumber="8">
<keyLead id="E14A34072A46D840FF3E7FEB1A9FB0DC" box="[151,1436,1764,1785]" pageId="7" pageNumber="8">- Dorsal surface glabrous, or with scarce pubescence restricted to some body parts. Elytra fully metallic or with reddish patch. 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="E14F8F972A46D840FF3E7E0E1A98B14B" pageId="7" pageNumber="8">
<paragraph id="5A0497322A46D840FF3E7E0E1A9FB116" blockId="7.[151,1436,942,2019]" pageId="7" pageNumber="8">
<keyLead id="E14A34072A46D840FF3E7E0E1A9FB116" pageId="7" pageNumber="8">
9. Antennomeres 13 with numerous fine setae in addition to the primary ones (
<figureCitation id="C2808BB72A46D840FC6B7E0E1CF4B133" box="[962,1015,1793,1814]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="7" pageNumber="8">fig. 7</figureCitation>
); pronotal lateral margin setose through full length (
<figureCitation id="C2808BB72A46D840FE837E111E69B116" box="[298,362,1822,1843]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="7" pageNumber="8">fig. 11</figureCitation>
). Endophallus with a long and bent chitinized flagellum (
<figureCitation id="C2808BB72A46D840FC387E111CD1B116" box="[913,978,1822,1843]" captionStart="FIGURES 1921" captionStartId="10.[151,264,1927,1949]" captionTargetBox="[221,1323,236,1868]" captionTargetId="figure@10.[204,1364,193,1894]" captionTargetPageId="10" captionText="FIGURES 1921. Calleida gressittiana sp. n. 19 habitus and labels of holotype (male, Kinabalu, Borneo, MNHN), scale bar = 2.0 mm; 20 habitus of paratype (female, Sandakan, Borneo, NHML), scale bar = 2.0 mm; 21 median lobe of aedeagus, rightlateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302717/files/figure.png" pageId="7" pageNumber="8">fig. 21</figureCitation>
)............1.
<taxonomicName id="9DBBECB12A46D840FB2F7E111A10B116" ID-CoL="8MMBN" box="[1158,1299,1822,1843]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gressittiana">
<emphasis id="68CF4B202A46D840FB2F7E111A10B116" box="[1158,1299,1822,1843]" italics="true" pageId="7" pageNumber="8">C. gressittiana</emphasis>
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species group
</keyLead>
</paragraph>
<paragraph id="5A0497322A46D840FF3E7E331A98B14B" blockId="7.[151,1436,942,2019]" pageId="7" pageNumber="8">
<keyLead id="E14A34072A46D840FF3E7E331A98B14B" pageId="7" pageNumber="8">
- Antennomeres 13 with only the primary apical setae (
<figureCitation id="C2808BB72A46D840FD477E331C27B174" box="[750,804,1852,1873]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="7" pageNumber="8">fig. 9</figureCitation>
); pronotum lateral margin with fine setae restricted to the anterior angles (
<figureCitation id="C2808BB72A46D840FE827E561E6EB14B" box="[299,365,1881,1902]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="7" pageNumber="8">fig. 13</figureCitation>
). Endophallus with a pair of copulatory pieces not modified as long flagellum......................... 10
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10. Only anterior angles of pronotum setose, elytral humeral margins without setae. Elytra very elongate, EL/EW 1.802.05. Median lobe of aedeagus simple, without ventral ridges or lobes (
<figureCitation id="C2808BB72A46D840FCFC7E9B1CC6B18C" box="[853,965,1940,1961]" captionStart="FIGURES 3035" captionStartId="17.[151,264,1228,1250]" captionTargetBox="[160,1405,496,1200]" captionTargetId="figure@17.[151,1436,481,1207]" captionTargetPageId="17" captionText="FIGURES 3035. Calleida lativittis Chaudoir. 30 habitus and labels of lectotype (female, &quot;Deccan&quot;, MNHN), scale bar = 2.0 mm; 31 habitus of paralectotype (female, &quot;Deccan&quot;, MNHN), scale bar = 2.0 mm; 3235 male genitalia, scale bar = 0.5 mm: 32 median lobe of aedeagus, dorsal view; 33 left paramere of aedeagus; 34 median lobe of aedeagus, left-lateral view; 35 male sternum IX." httpUri="https://zenodo.org/record/1302725/files/figure.png" pageId="7" pageNumber="8">figs. 32, 34</figureCitation>
)........... 4.
<taxonomicName id="9DBBECB12A46D840FBC37E9B1BD7B18C" ID-CoL="5X83Q" authorityName="Chaudoir Lectotypus" authorityYear="1872" box="[1130,1236,1940,1961]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lativittis">
<emphasis id="68CF4B202A46D840FBC37E9B1BD7B18C" box="[1130,1236,1940,1961]" italics="true" pageId="7" pageNumber="8">C. lativittis</emphasis>
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species group (pars)
</keyLead>
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<paragraph id="5A0497322A46D840FF3E7EBE1A9FB1C6" blockId="7.[151,1436,942,2019]" pageId="7" pageNumber="8">
<keyLead id="E14A34072A46D840FF3E7EBE1A9FB1C6" pageId="7" pageNumber="8">
- Both anterior angles of pronotum and elytral humeral margins setose (
<figureCitation id="C2808BB72A46D840FC2D7EBE1CC4B1E3" box="[900,967,1969,1990]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="7" pageNumber="8">fig. 10</figureCitation>
). Elytra shorter, EL/EW 1.651.75. Median lobe of aedeagus peculiar in form, irregularly explanate and ridged ventrally..............8.
<taxonomicName id="9DBBECB12A46D840FBE17EC11BD7B1C6" ID-CoL="5WTZW" authorityName="Dejean" authorityYear="1831" box="[1096,1236,1998,2019]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="chloroptera">
<emphasis id="68CF4B202A46D840FBE17EC11BD7B1C6" box="[1096,1236,1998,2019]" italics="true" pageId="7" pageNumber="8">C. chloroptera</emphasis>
</taxonomicName>
species group (pars)
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[1]
<taxonomicName id="9DBBECB12A48D84EFF697A531D60B553" ID-CoL="8MMBN" authority="Casale &amp; Shi" authorityName="Casale &amp; Shi" box="[192,611,860,886]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="gressittiana" status="sp. nov.">
<emphasis id="68CF4B202A48D84EFF697A531EBAB553" bold="true" box="[192,441,860,886]" italics="true" pageId="9" pageNumber="10">Calleida gressittiana</emphasis>
Casale &amp; Shi
</taxonomicName>
<taxonomicNameLabel id="73FCF65B2A48D84EFDC37A531DAFB553" box="[618,684,860,886]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A48D84EFF3E7A921FE9B24F" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="5A0497322A48D84EFF3E7A921C65B593" blockId="9.[151,1437,925,2030]" box="[151,870,925,950]" pageId="9" pageNumber="10">
<emphasis id="68CF4B202A48D84EFF3E7A921E3AB593" bold="true" box="[151,313,925,950]" pageId="9" pageNumber="10">
<typeStatus id="850029902A48D84EFF3E7A921FD1B593" box="[151,210,925,950]" pageId="9" pageNumber="10">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="22ACD7A22A48D84EFEE97A921EA9B593" box="[320,426,925,950]" name="Malaysia" pageId="9" pageNumber="10">Malaysia</collectingCountry>
, N Borneo:
<collectingRegion id="987F59D02A48D84EFD987A921D7BB593" box="[561,632,925,950]" country="Malaysia" name="Sabah" pageId="9" pageNumber="10">Sabah</collectingRegion>
, Kinabalu Mountain.
</paragraph>
<paragraph id="5A0497322A48D84EFF6E7ACE1FE9B24F" blockId="9.[151,1437,925,2030]" pageId="9" pageNumber="10">
<emphasis id="68CF4B202A48D84EFF6E7ACE1E75B5FF" bold="true" box="[199,374,961,986]" pageId="9" pageNumber="10">
<typeStatus id="850029902A48D84EFF6E7ACE1E01B5FF" box="[199,258,961,986]" pageId="9" pageNumber="10">Type</typeStatus>
material:
</emphasis>
<materialsCitation id="EAD39D6F2A48D84EFED47ACE1A9FB5FF" ID-GBIF-Occurrence="1852047888" box="[381,1436,961,986]" collectingDate="1903-08-05" collectorName="John Waterstradt" location="Nord Bornéo Mont Kina Balu" pageId="9" pageNumber="10" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="850029902A48D84EFED47ACE1EEBB5FF" box="[381,488,961,986]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
,
<specimenCount id="4CBD5CBB2A48D84EFE5A7ACE1D2FB5FF" box="[499,556,961,986]" pageId="9" pageNumber="10" type="male">male</specimenCount>
, “
<location id="5F64C1E92A48D84EFDE87ACE1C90B5FF" LSID="urn:lsid:plazi:treatment:D21226242A48D84CFF3E7A531E44B1CA:5F64C1E92A48D84EFDE87ACE1C90B5FF" box="[577,915,961,986]" name="Nord Bornéo Mont Kina Balu" pageId="9" pageNumber="10">
Nord Bornéo Mont
<taxonomicName id="9DBBECB12A48D84EFC887ACE1C90B5FF" authority="Balu" authorityName="Balu" box="[801,915,961,986]" class="Echinoidea" family="Schizasteridae" genus="Kina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spatangoida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="genus">Kina Balu</taxonomicName>
</location>
<collectingDate id="3E41481A2A48D84EFC337ACE1B03B5FF" box="[922,1024,961,986]" pageId="9" pageNumber="10" value="1903-08-05">
<date id="2E05B1F22A48D84EFC337ACE1B03B5FF" box="[922,1024,961,986]" pageId="9" pageNumber="10" value="1903-08-05">5-8-1903</date>
</collectingDate>
<collectorName id="F74EF2E42A48D84EFBAE7ACE1BC5B5FF" box="[1031,1222,961,986]" pageId="9" pageNumber="10">John Waterstradt</collectorName>
” (MNHN,
<figureCitation id="C2808BB72A48D84EFAEA7ACE1A8CB5FF" box="[1347,1423,961,986]" captionStart="FIGURES 1921" captionStartId="10.[151,264,1927,1949]" captionTargetBox="[221,1323,236,1868]" captionTargetId="figure@10.[204,1364,193,1894]" captionTargetPageId="10" captionText="FIGURES 1921. Calleida gressittiana sp. n. 19 habitus and labels of holotype (male, Kinabalu, Borneo, MNHN), scale bar = 2.0 mm; 20 habitus of paratype (female, Sandakan, Borneo, NHML), scale bar = 2.0 mm; 21 median lobe of aedeagus, rightlateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302717/files/figure.png" pageId="9" pageNumber="10">fig. 19</figureCitation>
).
</materialsCitation>
<typeStatus id="850029902A48D84EFF3E7AE91E0AB5DB" box="[151,265,998,1022]" pageId="9" pageNumber="10" type="paratype">Paratypes</typeStatus>
: one female “
<collectingCountry id="22ACD7A22A48D84EFE0F7AEA1D10B5DB" box="[422,531,997,1022]" name="Malaysia" pageId="9" pageNumber="10">Malaysia</collectingCountry>
: Borneo (Brit. N.) Sandakan Bay (NW) Sepilok For. Res.
<quantity id="9D433AD72A48D84EFB1B7AE91A04B5D8" box="[1202,1287,998,1022]" metricMagnitude="0" metricUnit="m" metricValue="5.5" metricValueMax="10.0" metricValueMin="1.0" pageId="9" pageNumber="10" unit="m" value="5.5" valueMax="10.0" valueMin="1.0">1-10 m</quantity>
,
<date id="2E05B1F22A48D84EFABC7AEA1A92B5DB" box="[1301,1425,997,1022]" pageId="9" pageNumber="10" value="1957-10-27">X-27-1957</date>
<collectionCode id="3CAA0FF72A48D84EFF3E7D051FECB207" box="[151,239,1034,1058]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="9" pageNumber="10">NHML</collectionCode>
); one female, same data, “J.L. Gressitt Malaise trap J.L. Gressitt collector Bishop Museum” (
<collectionCode id="3CAA0FF72A48D84EFABA7D051A69B207" box="[1299,1386,1034,1058]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="9" pageNumber="10">NHML</collectionCode>
,
<figureCitation id="C2808BB72A48D84EFADF7D061FB4B263" captionStart="FIGURES 1921" captionStartId="10.[151,264,1927,1949]" captionTargetBox="[221,1323,236,1868]" captionTargetId="figure@10.[204,1364,193,1894]" captionTargetPageId="10" captionText="FIGURES 1921. Calleida gressittiana sp. n. 19 habitus and labels of holotype (male, Kinabalu, Borneo, MNHN), scale bar = 2.0 mm; 20 habitus of paratype (female, Sandakan, Borneo, NHML), scale bar = 2.0 mm; 21 median lobe of aedeagus, rightlateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302717/files/figure.png" pageId="9" pageNumber="10">fig. 20</figureCitation>
); one female, “
<collectingCountry id="22ACD7A22A48D84EFEC37D221ED5B263" box="[362,470,1069,1094]" name="Malaysia" pageId="9" pageNumber="10">Malaysia</collectingCountry>
,
<collectingRegion id="987F59D02A48D84EFE4C7D221D68B263" box="[485,619,1069,1094]" country="Indonesia" name="Kalimantan Utara" pageId="9" pageNumber="10">Kalimantan</collectingRegion>
, Kota Kinabalu, Gunung Emas Nat. Park
<emphasis id="68CF4B202A48D84EFBF57D221BBBB263" box="[1116,1208,1069,1094]" italics="true" pageId="9" pageNumber="10">Raflesia</emphasis>
(sic!) 6-
<date id="2E05B1F22A48D84EFAB47D221A91B263" box="[1309,1426,1069,1094]" pageId="9" pageNumber="10" value="1999-10-02">10.2.1999</date>
” (
<collectionCode id="3CAA0FF72A48D84EFF367D5D1FDDB24F" box="[159,222,1106,1130]" country="Germany" httpUri="http://grbio.org/cool/mqei-3i36" name="Culture Collection of Algae at the University of Marburg" pageId="9" pageNumber="10">CCA</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A48D84EFF6E7D7A1D30B343" pageId="9" pageNumber="10" type="etymology">
<paragraph id="5A0497322A48D84EFF6E7D7A1D30B343" blockId="9.[151,1437,925,2030]" pageId="9" pageNumber="10">
<emphasis id="68CF4B202A48D84EFF6E7D7A1E8AB2AB" bold="true" box="[199,393,1141,1166]" pageId="9" pageNumber="10">Specific epithet.</emphasis>
We are pleased to dedicate this new, handsome species to Judson Linsley Gressitt, who devoted much effort and time to developing knowledge of the insect fauna of SE Asia, New
<collectingCountry id="22ACD7A22A48D84EFB317D961BEAB297" box="[1176,1257,1177,1202]" name="Guinea-Bissau" pageId="9" pageNumber="10">Guinea</collectingCountry>
and
<collectingCountry id="22ACD7A22A48D84EFAB77D961A9FB297" box="[1310,1436,1177,1202]" name="Micronesia" pageId="9" pageNumber="10">Micronesia</collectingCountry>
and collected two specimens of this species in Borneo. J. L. Gressitt was born in
<collectingRegion id="987F59D02A48D84EFB947DB21B87B2F3" box="[1085,1156,1213,1238]" country="Japan" name="Tokyo" pageId="9" pageNumber="10">Tokyo</collectingRegion>
on
<date id="2E05B1F22A48D84EFB1B7DB21A48B2F3" box="[1202,1355,1213,1238]" pageId="9" pageNumber="10" value="1914-06-16">16 June 1914</date>
and as professional entomologist and zoologist spent his academic activity among
<collectingCountry id="22ACD7A22A48D84EFC547DED1B2EB2DF" box="[1021,1069,1250,1274]" name="United States of America" pageId="9" pageNumber="10">U.S.</collectingCountry>
A,
<collectingCountry id="22ACD7A22A48D84EFBFD7DEE1B94B2DF" box="[1108,1175,1249,1274]" name="China" pageId="9" pageNumber="10">China</collectingCountry>
and
<collectingCountry id="22ACD7A22A48D84EFB7B7DED1A16B2DC" box="[1234,1301,1250,1273]" name="Japan" pageId="9" pageNumber="10">Japan</collectingCountry>
, with many biological explorations in remote areas, mountains climbed, and countless flights in small planes in several dangerous countries. He and his wife Margaret were aboard a regular commercial flight that crashed in south
<collectingCountry id="22ACD7A22A48D84EFAF07C261A9FB367" box="[1369,1436,1321,1346]" name="China" pageId="9" pageNumber="10">China</collectingCountry>
on
<date id="2E05B1F22A48D84EFF137C421E5DB343" box="[186,350,1357,1382]" pageId="9" pageNumber="10" value="1982-04-26">April 26, 1982</date>
(
<bibRefCitation id="3E2AEAC32A48D84EFEC57C421D25B343" box="[364,550,1357,1382]" pageId="9" pageNumber="10" refId="ref30797" refString="Radovsky, F. (1983) J. Linsley Gressitt (1914 - 1982). International Journal of Entomology, 25 (1), 1 - 10." type="journal article">Radovsky, 1983</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A48D84EFF6E7C7E1D8BB11F" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="5A0497322A48D84EFF6E7C7E1DF0B0A3" blockId="9.[151,1437,925,2030]" pageId="9" pageNumber="10">
<emphasis id="68CF4B202A48D84EFF6E7C7E1E42B3AF" bold="true" box="[199,321,1393,1418]" pageId="9" pageNumber="10">Diagnosis.</emphasis>
With the character states of the genus
<taxonomicName id="9DBBECB12A48D84EFCA17C7E1C6AB3AF" authorityName="Latreille" authorityYear="1824" box="[776,873,1393,1418]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A48D84EFCA17C7E1C6AB3AF" box="[776,873,1393,1418]" italics="true" pageId="9" pageNumber="10">Calleida</emphasis>
</taxonomicName>
in the widest sense, but from all Asiatic species known so far markedly characterized by the peculiar combination of the following morphological characters: medium sized (L:
<quantity id="9D433AD72A48D84EFECD7CB51EECB3F7" box="[356,495,1466,1490]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="8.8" metricValueMin="8.2" pageId="9" pageNumber="10" unit="mm" value="8.5" valueMax="8.8" valueMin="8.2">8.28.8 mm</quantity>
); antennomeres 13 with several additional fine setae besides the primary ones (
<figureCitation id="C2808BB72A48D84EFADC7CB61FA4B3D3" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="9" pageNumber="10">fig. 7</figureCitation>
); pronotum markedly cordiform, its anterior angles and lateral margins setose (
<figureCitation id="C2808BB72A48D84EFB967CD21B8EB3D3" box="[1087,1165,1501,1526]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="9" pageNumber="10">fig. 11</figureCitation>
); elytra elongate-ovate; elytral striae very deep, punctate, intervals convex. Dorsal surface markedly bicolor: head and pronotum reddish, contrasting with the metallic, dark blue-violet elytra; legs bicolor: femora and tibiae blackish, tarsomeres reddishbrown. Abdominal sternum VII with one seta on each side in males, two in females. Male genitalia with a long and bent chitinized flagellum on the endophallus (
<figureCitation id="C2808BB72A48D84EFD307F621DE5B0A3" box="[665,742,1645,1670]" captionStart="FIGURES 1921" captionStartId="10.[151,264,1927,1949]" captionTargetBox="[221,1323,236,1868]" captionTargetId="figure@10.[204,1364,193,1894]" captionTargetPageId="10" captionText="FIGURES 1921. Calleida gressittiana sp. n. 19 habitus and labels of holotype (male, Kinabalu, Borneo, MNHN), scale bar = 2.0 mm; 20 habitus of paratype (female, Sandakan, Borneo, NHML), scale bar = 2.0 mm; 21 median lobe of aedeagus, rightlateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302717/files/figure.png" pageId="9" pageNumber="10">fig. 21</figureCitation>
).
</paragraph>
<paragraph id="5A0497322A48D84EFF6E7F9E1D8BB11F" blockId="9.[151,1437,925,2030]" pageId="9" pageNumber="10">
This species is similar to some representatives of the
<taxonomicName id="9DBBECB12A48D84EFC8D7F9D1CCAB08F" authorityName="Dejean" authorityYear="1831" box="[804,969,1681,1706]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="chloroptera">
<emphasis id="68CF4B202A48D84EFC8D7F9D1CCAB08F" box="[804,969,1681,1706]" italics="true" pageId="9" pageNumber="10">C. chloroptera</emphasis>
</taxonomicName>
species group in general appearance, but can be easily distinguished by the presence of additional setae on the lateral margins of pronotum and the three basal antennomeres. Only one other species,
<taxonomicName id="9DBBECB12A48D84EFD007FD51C69B0D4" authority="Bates" authorityName="Bates" box="[681,874,1753,1778]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="excelsa">
<emphasis id="68CF4B202A48D84EFD007FD51C23B0D7" box="[681,800,1753,1778]" italics="true" pageId="9" pageNumber="10">C. excelsa</emphasis>
Bates
</taxonomicName>
(
<figureCitation id="C2808BB72A48D84EFCD77FD61CC3B0D7" box="[894,960,1753,1778]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="9" pageNumber="10">fig. 8</figureCitation>
), have accessory setae on the first three antennomeres among all Asiatic
<taxonomicName id="9DBBECB12A48D84EFDAE7FF21D64B133" authorityName="Latreille" authorityYear="1824" box="[519,615,1789,1814]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A48D84EFDAE7FF21D64B133" box="[519,615,1789,1814]" italics="true" pageId="9" pageNumber="10">Calleida</emphasis>
</taxonomicName>
, but is markedly different from the new species by its densely pubescent body (see below,
<taxonomicName id="9DBBECB12A48D84EFEF37E2D1ED4B11F" authorityName="Chaudoir Lectotypus" authorityYear="1872" box="[346,471,1825,1850]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="lativittis">
<emphasis id="68CF4B202A48D84EFEF37E2D1ED4B11F" box="[346,471,1825,1850]" italics="true" pageId="9" pageNumber="10">C. lativittis</emphasis>
</taxonomicName>
species group).
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A48D84CFF6E7E4A1D30B1A6" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10" type="description">
<paragraph id="5A0497322A48D84EFF6E7E4A1A2AB178" blockId="9.[151,1437,925,2030]" box="[199,1321,1861,1886]" pageId="9" pageNumber="10">
<emphasis id="68CF4B202A48D84EFF6E7E4A1E5AB17B" bold="true" box="[199,345,1861,1886]" pageId="9" pageNumber="10">Description.</emphasis>
<emphasis id="68CF4B202A48D84EFEF67E4A1D18B17B" box="[351,539,1861,1886]" italics="true" pageId="9" pageNumber="10">General features</emphasis>
as in
<figureCitation id="C2808BB72A48D84EFDF47E4A1DFFB17B" box="[605,764,1861,1886]" captionStart="FIGURES 1921" captionStartId="10.[151,264,1927,1949]" captionTargetBox="[221,1323,236,1868]" captionTargetId="figure@10.[204,1364,193,1894]" captionTargetPageId="10" captionText="FIGURES 1921. Calleida gressittiana sp. n. 19 habitus and labels of holotype (male, Kinabalu, Borneo, MNHN), scale bar = 2.0 mm; 20 habitus of paratype (female, Sandakan, Borneo, NHML), scale bar = 2.0 mm; 21 median lobe of aedeagus, rightlateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302717/files/figure.png" pageId="9" pageNumber="10">figures 19, 20</figureCitation>
. Medium sized: L:
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; TL: 8.0
<quantity id="9D433AD72A48D84EFB657E491A26B178" box="[1228,1317,1862,1886]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="9" pageNumber="10" unit="mm" value="8.4">8.4 mm</quantity>
.
</paragraph>
<paragraph id="5A0497322A48D84EFF6E7E661FE5B1CB" blockId="9.[151,1437,925,2030]" pageId="9" pageNumber="10">
<emphasis id="68CF4B202A48D84EFF6E7E661E15B1A7" box="[199,278,1897,1922]" italics="true" pageId="9" pageNumber="10">Colour</emphasis>
: head and pronotum reddish; basal two antennomeres and base of the third reddish; following antennomeres dark piceous. Pterothorax and abdominal sterna blackish. Elytra metallic dark blue-violet, with a touch of metallic cupreous-green at base and apex. Femora and tibiae dark piceous to black, tarsomeres reddishbrown.
</paragraph>
<caption id="0EC4C7BA2A4BD84DFF3E7E881C98B1FF" httpUri="https://zenodo.org/record/1302717/files/figure.png" pageId="10" pageNumber="11" startId="10.[151,264,1927,1949]" targetBox="[221,1323,236,1868]" targetPageId="10">
<paragraph id="5A0497322A4BD84DFF3E7E881C98B1FF" blockId="10.[151,1436,1926,2010]" pageId="10" pageNumber="11">
<emphasis id="68CF4B202A4BD84DFF3E7E881D2CB1B9" bold="true" box="[151,559,1926,1949]" pageId="10" pageNumber="11">
FIGURES 1921.
<taxonomicName id="9DBBECB12A4BD84DFEF07E891D2CB1B9" box="[345,559,1926,1948]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="gressittiana" status="sp. nov.">
<emphasis id="68CF4B202A4BD84DFEF07E891D2CB1B9" bold="true" box="[345,559,1926,1948]" italics="true" pageId="10" pageNumber="11">Calleida gressittiana</emphasis>
</taxonomicName>
</emphasis>
<taxonomicNameLabel id="73FCF65B2A4BD84DFD9C7E871D6FB1B8" box="[565,620,1928,1949]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
<emphasis id="68CF4B202A4BD84DFDDB7E881D88B1B8" bold="true" box="[626,651,1927,1949]" pageId="10" pageNumber="11">19</emphasis>
habitus and labels of holotype (male, Kinabalu, Borneo, MNHN), scale bar = 2.0 mm;
<emphasis id="68CF4B202A4BD84DFF5D7EAA1E0EB19E" bold="true" box="[244,269,1957,1979]" pageId="10" pageNumber="11">20</emphasis>
habitus of paratype (female, Sandakan, Borneo, NHML), scale bar = 2.0 mm;
<emphasis id="68CF4B202A4BD84DFB9E7EAA1B52B19E" bold="true" box="[1079,1105,1957,1979]" pageId="10" pageNumber="11">21</emphasis>
median lobe of aedeagus, rightlateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm.
</paragraph>
</caption>
<caption id="0EC4C7BA2A4AD84CFF3E7AFE1B27B222" box="[151,1060,1009,1031]" httpUri="https://zenodo.org/record/1302719/files/figure.png" pageId="11" pageNumber="12" startId="11.[151,209,1009,1031]" targetBox="[262,1324,193,988]" targetPageId="11">
<paragraph id="5A0497322A4AD84CFF3E7AFE1B27B222" blockId="11.[151,1060,1009,1031]" box="[151,1060,1009,1031]" pageId="11" pageNumber="12">
<emphasis id="68CF4B202A4AD84CFF3E7AFE1FE7B222" bold="true" box="[151,228,1009,1031]" pageId="11" pageNumber="12">MAP 1</emphasis>
Distributions for species in the
<taxonomicName id="9DBBECB12A4AD84CFD8E7AFD1DB9B222" box="[551,698,1010,1031]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="gressittiana">
<emphasis id="68CF4B202A4AD84CFD8E7AFD1DB9B222" box="[551,698,1010,1031]" italics="true" pageId="11" pageNumber="12">C. gressittiana</emphasis>
</taxonomicName>
group:
<taxonomicName id="9DBBECB12A4AD84CFCA37AFD1C9EB222" box="[778,925,1010,1031]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="gressittiana" status="sp. nov.">
<emphasis id="68CF4B202A4AD84CFCA37AFD1C9EB222" box="[778,925,1010,1031]" italics="true" pageId="11" pageNumber="12">C. gressittiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73FCF65B2A4AD84CFC0D7AFD1CD8B222" box="[932,987,1010,1031]" pageId="11" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
(blue).
</paragraph>
</caption>
<paragraph id="5A0497322A4AD84CFF6E7D3F1BE8B2AA" blockId="11.[151,1437,1070,2031]" pageId="11" pageNumber="12">
<emphasis id="68CF4B202A4AD84CFF6E7D3F1EF1B262" box="[199,498,1070,1095]" italics="true" pageId="11" pageNumber="12">Lustre and microsculpture</emphasis>
: head, pronotum and elytra shiny, with generally effaced microsculpture on head and pronotum, more evident on the elytral intervals for females, in form of transverse mesh pattern, effaced on pronotum for males. Pronotal disc and elytral intervals with fine, hardly visible micro-punctuation.
</paragraph>
<paragraph id="5A0497322A4AD84CFF6E7D951A30B2F2" blockId="11.[151,1437,1070,2031]" pageId="11" pageNumber="12">
<emphasis id="68CF4B202A4AD84CFF6E7D951E07B296" box="[199,260,1178,1203]" italics="true" pageId="11" pageNumber="12">Head</emphasis>
: wide, with evident neck constriction; genae swollen, contiguous with the posterior margin of eyes; eyes very large and prominent. Antennomeres 13 with several additional fine setae besides the primary setae.
</paragraph>
<paragraph id="5A0497322A4AD84CFF6E7DEB1A52B3AF" blockId="11.[151,1437,1070,2031]" pageId="11" pageNumber="12">
<emphasis id="68CF4B202A4AD84CFF6E7DEB1E35B2DE" box="[199,310,1252,1275]" italics="true" pageId="11" pageNumber="12">Pronotum</emphasis>
: transverse-cordiform (ratio PW/PL: 1.181.23), with lateral margins deeply sinuate and constricted to the base; anterior angles setose, lateral margins densely setose on anterior half, sporadically setose on posterior half. Lateral furrows wide, lateral reflection moderate. Disc depressed, with superficial, scarce transverse wrinkles. Anterior angles not prominent; basal angles acutely prominent outside, basal foveae wide and deep. Base straight, slightly oblique and beaded at sides only. One mid-lateral seta and one basolateral seta on each side present.
</paragraph>
<paragraph id="5A0497322A4AD84CFF6E7C991B99B01A" blockId="11.[151,1437,1070,2031]" pageId="11" pageNumber="12">
<emphasis id="68CF4B202A4AD84CFF6E7C991E0EB38A" box="[199,269,1430,1455]" italics="true" pageId="11" pageNumber="12">Elytra</emphasis>
: elongate-ovate (ratio EL/EW: 1.551.64), sub-convex, slightly widened at the apical third; humeral angles rounded; striae deeply impressed, punctate; intervals convex. Post-humeral sinuation inconspicuous, preapical callosity small, slightly distinct at apex of the sixth and seventh intervals. Apex beaded, oblique, with the outer angles fully rounded, not thickened. The third interval with two discal and one apical setiferous punctures; umbilicate series of 16 deep, foveate punctures along eighth stria, interrupted in the middle.
</paragraph>
<paragraph id="5A0497322A4AD84CFF6E7F431B23B0A2" blockId="11.[151,1437,1070,2031]" pageId="11" pageNumber="12">
<emphasis id="68CF4B202A4AD84CFF6E7F431FFFB046" box="[199,252,1612,1635]" italics="true" pageId="11" pageNumber="12">Legs</emphasis>
: basal two metatarsomeres with distinct dorsal groove; the fourth tarsomere deeply bilobed, its lobes long and widened. Tarsal claws denticulate, each with six long teeth on the inner side.
</paragraph>
<paragraph id="5A0497322A4AD84CFF6E7F9D1EBEB0EA" blockId="11.[151,1437,1070,2031]" pageId="11" pageNumber="12">
<emphasis id="68CF4B202A4AD84CFF6E7F9D1E91B08E" box="[199,402,1682,1707]" italics="true" pageId="11" pageNumber="12">Abdominal sterna</emphasis>
: sternum VII with one seta on each side in males, two setae in females; apex notched in males, straight in females.
</paragraph>
<paragraph id="5A0497322A4AD84CFF6E7FD51CA3B17A" blockId="11.[151,1437,1070,2031]" pageId="11" pageNumber="12">
<emphasis id="68CF4B202A4AD84CFF6E7FD51E6CB0D6" box="[199,367,1754,1779]" italics="true" pageId="11" pageNumber="12">Male genitalia</emphasis>
: median lobe of aedeagus (
<figureCitation id="C2808BB72A4AD84CFD1A7FD51C02B0D6" box="[691,769,1754,1779]" captionStart="FIGURES 1921" captionStartId="10.[151,264,1927,1949]" captionTargetBox="[221,1323,236,1868]" captionTargetId="figure@10.[204,1364,193,1894]" captionTargetPageId="10" captionText="FIGURES 1921. Calleida gressittiana sp. n. 19 habitus and labels of holotype (male, Kinabalu, Borneo, MNHN), scale bar = 2.0 mm; 20 habitus of paratype (female, Sandakan, Borneo, NHML), scale bar = 2.0 mm; 21 median lobe of aedeagus, rightlateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302717/files/figure.png" pageId="11" pageNumber="12">fig. 21</figureCitation>
) flattened, in lateral view slightly curved; ventral margin distinctly protruded near the middle, in ventral aspect slightly expanded before the apical orifice; apical lamina short, rounded in ventral aspect; endophallus with a long chitinized flagellum (as modification of the ventral copulatory piece), twisted and dilated at base, bent backward at apex.
</paragraph>
<paragraph id="5A0497322A4AD84CFF6E7E651D30B1A6" blockId="11.[151,1437,1070,2031]" box="[199,563,1898,1923]" pageId="11" pageNumber="12">
<emphasis id="68CF4B202A4AD84CFF6E7E651E86B1A6" box="[199,389,1898,1923]" italics="true" pageId="11" pageNumber="12">Female genitalia</emphasis>
: not examined.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A4AD84CFF6E7E811E44B1CA" pageId="11" pageNumber="12" type="discussion">
<paragraph id="5A0497322A4AD84CFF6E7E811E44B1CA" blockId="11.[151,1437,1070,2031]" pageId="11" pageNumber="12">
<emphasis id="68CF4B202A4AD84CFF6E7E811D9FB182" bold="true" box="[199,668,1934,1959]" pageId="11" pageNumber="12">Geographical distribution and habitat.</emphasis>
Known so far only from NE Borneo (
<collectingRegion id="987F59D02A4AD84CFBF17E811BA2B182" box="[1112,1185,1934,1959]" country="Malaysia" name="Sabah" pageId="11" pageNumber="12">Sabah</collectingRegion>
,
<collectingCountry id="22ACD7A22A4AD84CFB077E811A15B182" box="[1198,1302,1934,1959]" name="Malaysia" pageId="11" pageNumber="12">Malaysia</collectingCountry>
) in tropical rain forest (
<figureCitation id="C2808BB72A4AD84CFE877EBC1E7DB1EE" box="[302,382,1971,1995]" captionStart="MAP 1" captionStartId="11.[151,209,1009,1031]" captionTargetBox="[262,1324,193,988]" captionTargetId="figure@11.[262,1324,193,988]" captionTargetPageId="11" captionText="MAP 1 Distributions for species in the C. gressittiana group: C. gressittiana sp. n. (blue)." httpUri="https://zenodo.org/record/1302719/files/figure.png" pageId="11" pageNumber="12">map 1</figureCitation>
). Some specimens were obtained from Malaise trap and giant flowers of
<taxonomicName id="9DBBECB12A4AD84CFA917EBC1E47B1CA" authorityName="R.Brown" authorityYear="1821" class="Magnoliopsida" family="Rafflesiaceae" genus="Rafflesia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="genus">
<emphasis id="68CF4B202A4AD84CFA917EBC1A9FB1E9" box="[1336,1436,1971,1996]" italics="true" pageId="11" pageNumber="12">Rafflesia</emphasis>
(Rafflesiaceae)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>

View file

@ -0,0 +1,122 @@
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<mods:title id="6BF1B649E8553CAEA48158C52DBD37AF">Revision of the Oriental species of Calleida Latreille (sensu lato). Part 1: Introduction, groups of species, and species of six species groups (Coleoptera: Carabidae: Lebiini)</mods:title>
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<heading id="014C205E2A48D84EFF3E79971D58B697" bold="true" box="[151,603,151,178]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<emphasis id="68CF4B202A48D84EFF3E79971D58B697" bold="true" box="[151,603,151,178]" pageId="9" pageNumber="10">
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<keyStep id="E14F8F972A48D84EFF3E79971D58B697" box="[151,603,151,178]" pageId="9" pageNumber="10">
1.
<taxonomicName id="9DBBECB12A48D84EFF1D79981EAFB694" ID-CoL="8MMBN" box="[180,428,151,177]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="gressittiana">
<emphasis id="68CF4B202A48D84EFF1D79981EAFB694" bold="true" box="[180,428,151,177]" italics="true" pageId="9" pageNumber="10">Calleida gressittiana</emphasis>
</taxonomicName>
species group
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</heading>
</paragraph>
<paragraph id="5A0497322A48D84EFF3E79D11DABB78E" blockId="9.[151,1437,222,787]" pageId="9" pageNumber="10">
<emphasis id="68CF4B202A48D84EFF3E79D11EA3B6D2" bold="true" box="[151,416,222,247]" pageId="9" pageNumber="10">Diagnostic characters</emphasis>
: With the character states of
<taxonomicName id="9DBBECB12A48D84EFCA579D11C6FB6D2" ID-CoL="8KVFM" authorityName="Latreille" authorityYear="1824" box="[780,876,222,247]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A48D84EFCA579D11C6FB6D2" box="[780,876,222,247]" italics="true" pageId="9" pageNumber="10">Calleida</emphasis>
</taxonomicName>
(in the widest sense, see above), restricted or amplified as follows: Medium-sized species (L:
<quantity id="9D433AD72A48D84EFD61780C1C55B73E" box="[712,854,259,283]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="8.8" metricValueMin="8.2" pageId="9" pageNumber="10" unit="mm" value="8.5" valueMax="8.8" valueMin="8.2">8.28.8 mm</quantity>
). Dorsal side bicolor: head and pronotum reddish, elytra bluish, without reddish patch. Antennomeres 13 with numerous fine setae in addition to the primary ones (
<figureCitation id="C2808BB72A48D84EFF3678451FDEB746" box="[159,221,330,355]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="9" pageNumber="10">fig. 7</figureCitation>
); pronotal lateral margins setose through full length (
<figureCitation id="C2808BB72A48D84EFC9D78451C83B746" box="[820,896,330,355]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="9" pageNumber="10">fig. 11</figureCitation>
). Abdominal sternum VII with one seta on each side in males, two setae in females. Endophallus of aedeagus with a long and bent chitinized flagellum (
<figureCitation id="C2808BB72A48D84EFAE978611A8DB7A2" box="[1344,1422,366,391]" captionStart="FIGURES 1921" captionStartId="10.[151,264,1927,1949]" captionTargetBox="[221,1323,236,1868]" captionTargetId="figure@10.[204,1364,193,1894]" captionTargetPageId="10" captionText="FIGURES 1921. Calleida gressittiana sp. n. 19 habitus and labels of holotype (male, Kinabalu, Borneo, MNHN), scale bar = 2.0 mm; 20 habitus of paratype (female, Sandakan, Borneo, NHML), scale bar = 2.0 mm; 21 median lobe of aedeagus, rightlateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302717/files/figure.png" pageId="9" pageNumber="10">fig. 21</figureCitation>
), which is the modified ventral copulatory piece.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A48D84EFF6E78B91FE3B536" pageId="9" pageNumber="10" type="description">
<paragraph id="5A0497322A48D84EFF6E78B91D11B482" blockId="9.[151,1437,222,787]" pageId="9" pageNumber="10">
<emphasis id="68CF4B202A48D84EFF6E78B91E87B7EA" bold="true" box="[199,388,438,463]" pageId="9" pageNumber="10">Taxonomic note</emphasis>
: One only species,
<taxonomicName id="9DBBECB12A48D84EFDF678B81C01B7EB" box="[607,770,439,462]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="gressittiana" status="sp. nov.">
<emphasis id="68CF4B202A48D84EFDF678B81C01B7EB" box="[607,770,439,462]" italics="true" pageId="9" pageNumber="10">C. gressittiana</emphasis>
</taxonomicName>
<emphasis id="68CF4B202A48D84EFCA078B91C4BB7EA" bold="true" box="[777,840,438,463]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="73FCF65B2A48D84EFCA078B91C4BB7EA" box="[777,840,438,463]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
from Borneo (
<figureCitation id="C2808BB72A48D84EFC4578B81B31B7EA" box="[1004,1074,439,463]" captionStart="MAP 1" captionStartId="11.[151,209,1009,1031]" captionTargetBox="[262,1324,193,988]" captionTargetId="figure@11.[262,1324,193,988]" captionTargetPageId="11" captionText="MAP 1 Distributions for species in the C. gressittiana group: C. gressittiana sp. n. (blue)." httpUri="https://zenodo.org/record/1302719/files/figure.png" pageId="9" pageNumber="10">map 1</figureCitation>
), is attributed to this group. The phylogenetic position of this species group is questionable and isolated in the Asiatic fauna for its peculiar endophallus characters and the unique setation of antennae and lateral margins of pronotum. This is the only Asiatic species with a long, bent flagellum in endophallus, a feature widely represented amongst the Neotropical
<taxonomicName id="9DBBECB12A48D84EFF3E7B491FFBB47A" authorityName="Latreille" authorityYear="1824" box="[151,248,582,607]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A48D84EFF3E7B491FFBB47A" box="[151,248,582,607]" italics="true" pageId="9" pageNumber="10">Calleida</emphasis>
</taxonomicName>
species. On the contrary, all other Asiatic species have in endophallus two smaller (rarely only one in the
<taxonomicName id="9DBBECB12A48D84EFF3E7B641E24B4A6" box="[151,295,619,643]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="terminata">
<emphasis id="68CF4B202A48D84EFF3E7B641E24B4A6" box="[151,295,619,643]" italics="true" pageId="9" pageNumber="10">C. terminata</emphasis>
</taxonomicName>
species group), linear copulatory pieces, as in all
<taxonomicName id="9DBBECB12A48D84EFCC07B651CCAB4A6" authorityName="Latreille" authorityYear="1824" box="[873,969,618,643]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A48D84EFCC07B651CCAB4A6" box="[873,969,618,643]" italics="true" pageId="9" pageNumber="10">Calleida</emphasis>
</taxonomicName>
species from Africa and Madagascar, or unarmed (only in
<taxonomicName id="9DBBECB12A48D84EFEF67B801D01B482" box="[351,514,654,679]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="68CF4B202A48D84EFEF67B801D01B482" box="[351,514,654,679]" italics="true" pageId="9" pageNumber="10">C. puncticollis</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="5A0497322A48D84EFF6E7BBD1FE3B536" blockId="9.[151,1437,222,787]" pageId="9" pageNumber="10">
This only character, which could be a result of homoplasy (evolutionary convergence), does not indicate any relationship with Neotropical
<taxonomicName id="9DBBECB12A48D84EFE4C7BD91D5FB4CA" authorityName="Chaudoir" authorityYear="1873" box="[485,604,726,751]" class="Insecta" family="Carabidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="subTribe" subTribe="Calleidina">Calleidina</taxonomicName>
. It allows us, however, to isolate
<taxonomicName id="9DBBECB12A48D84EFC627BD81B6DB4CB" box="[971,1134,727,750]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="gressittiana" status="sp. nov.">
<emphasis id="68CF4B202A48D84EFC627BD81B6DB4CB" box="[971,1134,727,750]" italics="true" pageId="9" pageNumber="10">C. gressittiana</emphasis>
</taxonomicName>
<emphasis id="68CF4B202A48D84EFBDC7BD91BB7B4CA" bold="true" box="[1141,1204,726,751]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="73FCF65B2A48D84EFBDC7BD91BB7B4CA" box="[1141,1204,726,751]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
in a separate species group.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,319 @@
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<mods:title id="D59675D368FAAE626318DC81E7D260B1">Revision of the Oriental species of Calleida Latreille (sensu lato). Part 1: Introduction, groups of species, and species of six species groups (Coleoptera: Carabidae: Lebiini)</mods:title>
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<treatment id="D21226242A4DD848FF3E7A851E30B7F5" ID-GBIF-Taxon="146043511" LSID="urn:lsid:plazi:treatment:D21226242A4DD848FF3E7A851E30B7F5" httpUri="http://treatment.plazi.org/id/D21226242A4DD848FF3E7A851E30B7F5" lastPageId="15" lastPageNumber="16" pageId="12" pageNumber="13">
<subSubSection id="12A1C4B92A4DD84BFF3E7A851DB0B581" box="[151,691,906,932]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="5A0497322A4DD84BFF3E7A851DB0B581" blockId="12.[151,691,906,932]" box="[151,691,906,932]" pageId="12" pageNumber="13">
<heading id="014C205E2A4DD84BFF3E7A851DB0B581" bold="true" box="[151,691,906,932]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="68CF4B202A4DD84BFF3E7A851DB0B581" bold="true" box="[151,691,906,932]" pageId="12" pageNumber="13">
[2]
<taxonomicName id="9DBBECB12A4DD84BFF697A851D67B581" ID-CoL="8M9ZJ" authority="Shi &amp; Casale" authorityName="Shi &amp; Casale" box="[192,612,906,932]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="puncticollis" status="sp. nov.">
<emphasis id="68CF4B202A4DD84BFF697A851EBAB581" bold="true" box="[192,441,906,932]" italics="true" pageId="12" pageNumber="13">Calleida puncticollis</emphasis>
Shi &amp; Casale
</taxonomicName>
,
<taxonomicNameLabel id="73FCF65B2A4DD84BFDD87A851DB0B581" box="[625,691,906,932]" pageId="12" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A4DD84BFF3E7ADE1DA4B2C3" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="5A0497322A4DD84BFF3E7ADE1B02B5CF" blockId="12.[151,1436,977,2010]" box="[151,1025,977,1002]" pageId="12" pageNumber="13">
<emphasis id="68CF4B202A4DD84BFF3E7ADE1E3AB5CF" bold="true" box="[151,313,977,1002]" pageId="12" pageNumber="13">
<typeStatus id="850029902A4DD84BFF3E7ADE1FD1B5CF" box="[151,210,977,1002]" pageId="12" pageNumber="13">Type</typeStatus>
locality:
</emphasis>
<materialsCitation id="EAD39D6F2A4DD84BFEE97ADE1B02B5CF" ID-GBIF-Occurrence="1852048006" box="[320,1025,977,1002]" collectingDate="1994. IX. 6," collectionCode="Inst. Zool. Acad. Sin, (IZAS" collectorName="Chen Jun" country="China" county="Hubei, Zigui" elevation="110" latitude="31.034" location="Jiulingtou" longLatPrecision="72" longitude="110.791" municipality="Zigui" pageId="12" pageNumber="13" specimenCode="148278" specimenCount="1" specimenCount-male="1" stateProvince="Hubei" typeStatus="holotype">
<collectingCountry id="22ACD7A22A4DD84BFEE97ADE1E85B5CF" box="[320,390,977,1002]" name="China" pageId="12" pageNumber="13">China</collectingCountry>
, Hubei, Zigui county, Jiulingtou (
<geoCoordinate id="3F8FF1F52A4DD84BFD577ADD1C6CB5CF" box="[766,879,978,1002]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="55" value="31.034">N31.034°</geoCoordinate>
,
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).
</materialsCitation>
</paragraph>
<paragraph id="5A0497322A4DD84BFF6E7AFA1DA4B2C3" blockId="12.[151,1436,977,2010]" pageId="12" pageNumber="13">
<emphasis id="68CF4B202A4DD84BFF6E7AFA1E79B22B" bold="true" box="[199,378,1013,1038]" pageId="12" pageNumber="13">Type Material:</emphasis>
<typeStatus id="850029902A4DD84BFE287AFA1EEFB22B" box="[385,492,1013,1038]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
,
<specimenCount id="4CBD5CBB2A4DD84BFE517AFA1D32B22B" box="[504,561,1013,1038]" pageId="12" pageNumber="13" type="male">male</specimenCount>
, &quot;
<collectingCountry id="22ACD7A22A4DD84BFDEC7AFA1D8EB22B" box="[581,653,1013,1038]" name="China" pageId="12" pageNumber="13">China</collectingCountry>
,
<collectingCounty id="B365EFBE2A4DD84BFD307AFA1C29B22B" box="[665,810,1013,1038]" pageId="12" pageNumber="13">Hubei, Zigui</collectingCounty>
,
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,
<elevation id="D19670012A4DD84BFC1D7AF91CFFB22B" box="[948,1020,1014,1038]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="13" unit="m" value="110.0">
<quantity id="9D433AD72A4DD84BFC1D7AF91CFFB22B" box="[948,1020,1014,1038]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="13" unit="m" value="110.0">110 m</quantity>
</elevation>
&quot; [in Chinese]; &quot;
<collectingDate id="3E41481A2A4DD84BFB067AFA1A29B22B" box="[1199,1322,1013,1038]" pageId="12" pageNumber="13">
1994.
<date id="2E05B1F22A4DD84BFB587AFA1A27B22B" box="[1265,1316,1013,1038]" pageId="12" pageNumber="13">IX.6</date>
,
</collectingDate>
leg.
<collectorName id="F74EF2E42A4DD84BFAC87AFA1FC3B217" pageId="12" pageNumber="13">Chen Jun</collectorName>
&quot; [in Chinese]; &quot;
<specimenCode id="0A1D3F492A4DD84BFE2B7D151EDAB217" box="[386,473,1050,1074]" pageId="12" pageNumber="13">148278</specimenCode>
<collectionCode id="3CAA0FF72A4DD84BFE4C7D151DE7B217" box="[485,740,1049,1074]" pageId="12" pageNumber="13">Inst. Zool. Acad. Sin</collectionCode>
.&quot;
<collectionCode id="3CAA0FF72A4DD84BFD577D151C4BB217" box="[766,840,1050,1074]" pageId="12" pageNumber="13">(IZAS</collectionCode>
,
<figureCitation id="C2808BB72A4DD84BFCF07D161CAFB217" box="[857,940,1049,1074]" captionStart="FIGURES 2229" captionStartId="13.[151,264,1822,1844]" captionTargetBox="[158,1420,311,1777]" captionTargetId="figure@13.[151,1436,301,1801]" captionTargetPageId="13" captionText="FIGURES 2229. Calleida puncticollis sp. n. 22 habitus and labels of holotype (male, Hubei, IZAS), scale bar = 2.0 mm; 23 habitus of paratype (female, Shaanxi, IZAS), scale bar = 2.0 mm; 24 pronotum (Shaanxi, China), [coarsely punctate], scale bar = 0.5 mm; 25 ovipositor (Hubei, China), scale bar = 0.1 mm; 26 female reproductive tract (Hubei, China), scale bar = 0.5 mm; 27 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm; 28 left and right parameres of aedeagus (holotype), scale bar = 0.2 mm; 29 male sternum IX (holotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302721/files/figure.png" pageId="12" pageNumber="13">fig. 22</figureCitation>
).
<typeStatus id="850029902A4DD84BFC6C7D151B35B217" box="[965,1078,1050,1074]" pageId="12" pageNumber="13" type="paratype">Paratypes</typeStatus>
: one female, &quot;
<collectingRegion id="987F59D02A4DD84BFB4A7D161A33B217" box="[1251,1328,1049,1074]" country="China" name="Fujian" pageId="12" pageNumber="13">Fujian</collectingRegion>
, Fuzhou, 1982.
<date id="2E05B1F22A4DD84BFF7F7D311E2FB273" box="[214,300,1086,1110]" pageId="12" pageNumber="13" value="2012-08">VIII.12</date>
, leg. Chen Yuanhong&quot; (
<collectionCode id="3CAA0FF72A4DD84BFD917D311D7FB273" box="[568,636,1086,1110]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="12" pageNumber="13">IZAS</collectionCode>
); one female, &quot;
<collectingRegion id="987F59D02A4DD84BFC8D7D321C6DB273" box="[804,878,1085,1110]" country="China" name="Hubei" pageId="12" pageNumber="13">Hubei</collectingRegion>
, Zigui, Jiulingtou,
<quantity id="9D433AD72A4DD84BFBEA7D321B84B273" box="[1091,1159,1085,1110]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="13" unit="m" value="150.0">150m</quantity>
&quot;, &quot;1993.
<date id="2E05B1F22A4DD84BFB4E7D311A37B273" box="[1255,1332,1086,1110]" pageId="12" pageNumber="13" value="2013-07">VII.13</date>
, leg. Yao Jian&quot;, &quot;23357 Inst. Zool. Acad. Sin.&quot; (
<collectionCode id="3CAA0FF72A4DD84BFDE37D6D1D8DB25F" box="[586,654,1122,1146]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="12" pageNumber="13">IZAS</collectionCode>
); one female, &quot;
<collectingRegion id="987F59D02A4DD84BFC917D6D1C8BB25F" box="[824,904,1122,1146]" country="China" name="Hunan" pageId="12" pageNumber="13">Hunan</collectingRegion>
, Chenxian&quot;, &quot;1985.
<date id="2E05B1F22A4DD84BFBCC7D6E1BA6B25F" box="[1125,1189,1121,1146]" pageId="12" pageNumber="13" value="2017-04">IV.17</date>
&quot; (
<collectionCode id="3CAA0FF72A4DD84BFB167D6D1A00B25F" box="[1215,1283,1122,1146]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="12" pageNumber="13">IZAS</collectionCode>
); one female, &quot;
<collectingRegion id="987F59D02A4DD84BFF097D8A1E03B2BB" box="[160,256,1157,1182]" country="China" name="Shaanxi" pageId="12" pageNumber="13">Shaanxi</collectingRegion>
, Foping,
<quantity id="9D433AD72A4DD84BFED87D891EB7B2BB" box="[369,436,1158,1182]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="13" unit="m" value="900.0">900m</quantity>
, 1999.
<date id="2E05B1F22A4DD84BFDAA7D891D45B2BB" box="[515,582,1158,1182]" pageId="12" pageNumber="13" value="1927-06">VI.27</date>
, leg. Yao Jian&quot; (
<collectionCode id="3CAA0FF72A4DD84BFCA77D891C53B2BB" box="[782,848,1158,1182]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="12" pageNumber="13">IZAS</collectionCode>
,
<figureCitation id="C2808BB72A4DD84BFCC97D8A1CB2B2BB" box="[864,945,1157,1182]" captionStart="FIGURES 2229" captionStartId="13.[151,264,1822,1844]" captionTargetBox="[158,1420,311,1777]" captionTargetId="figure@13.[151,1436,301,1801]" captionTargetPageId="13" captionText="FIGURES 2229. Calleida puncticollis sp. n. 22 habitus and labels of holotype (male, Hubei, IZAS), scale bar = 2.0 mm; 23 habitus of paratype (female, Shaanxi, IZAS), scale bar = 2.0 mm; 24 pronotum (Shaanxi, China), [coarsely punctate], scale bar = 0.5 mm; 25 ovipositor (Hubei, China), scale bar = 0.1 mm; 26 female reproductive tract (Hubei, China), scale bar = 0.5 mm; 27 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm; 28 left and right parameres of aedeagus (holotype), scale bar = 0.2 mm; 29 male sternum IX (holotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302721/files/figure.png" pageId="12" pageNumber="13">fig. 23</figureCitation>
); one female, &quot;
<collectingRegion id="987F59D02A4DD84BFBCE7D8A1BC5B2BB" box="[1127,1222,1157,1182]" country="China" name="Shaanxi" pageId="12" pageNumber="13">Shaanxi</collectingRegion>
, Liuba, Miaotaizi,
<quantity id="9D433AD72A4DD84BFF3E7DA61FE4B2E7" box="[151,231,1193,1218]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="12" pageNumber="13" unit="m" value="1350.0">1350m</quantity>
, 1998.
<date id="2E05B1F22A4DD84BFE9E7DA51E81B2E7" box="[311,386,1194,1218]" pageId="12" pageNumber="13" value="1921-07">VII.21</date>
, leg. Yao Jian&quot; (
<collectionCode id="3CAA0FF72A4DD84BFDE07DA51D8AB2E7" box="[585,649,1194,1218]" country="Germany" httpUri="http://grbio.org/cool/mqei-3i36" name="Culture Collection of Algae at the University of Marburg" pageId="12" pageNumber="13">CCA</collectionCode>
); one ex., “China-Shaanxi, Lueyang,
<date id="2E05B1F22A4DD84BFBEB7DA51BDFB2E7" box="[1090,1244,1194,1218]" pageId="12" pageNumber="13" value="2009-05-23" valueMax="2009-05-30" valueMin="2009-05-23">23-30.V.2009</date>
, lgt. E. Kučera” (CDW); one male, “
<collectingRegion id="987F59D02A4DD84BFED17DC21EDEB2C3" box="[376,477,1229,1254]" country="China" name="Guangxi" pageId="12" pageNumber="13">Guangxi</collectingRegion>
, Guilin&quot; (
<collectionCode id="3CAA0FF72A4DD84BFDE37DC11D98B2C3" box="[586,667,1230,1254]" pageId="12" pageNumber="13">SCAU</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A4DD84BFF6E7DFE1C30B30B" pageId="12" pageNumber="13" type="etymology">
<paragraph id="5A0497322A4DD84BFF6E7DFE1C30B30B" blockId="12.[151,1436,977,2010]" pageId="12" pageNumber="13">
<emphasis id="68CF4B202A4DD84BFF6E7DFE1E85B32F" bold="true" box="[199,390,1265,1290]" pageId="12" pageNumber="13">Specific epithet.</emphasis>
In Latin,
<emphasis id="68CF4B202A4DD84BFE537DFD1D64B32C" box="[506,615,1266,1289]" italics="true" pageId="12" pageNumber="13">punctatus</emphasis>
means punctate, and
<emphasis id="68CF4B202A4DD84BFCF67DFE1C9FB32F" box="[863,924,1265,1290]" italics="true" pageId="12" pageNumber="13">collis</emphasis>
means neck; the specific name is referred to the heavily punctate neck and pronotum in the new species.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A4DD84BFF6E7C361FC6B0B0" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="5A0497322A4DD84BFF6E7C361CE0B023" blockId="12.[151,1436,977,2010]" pageId="12" pageNumber="13">
<emphasis id="68CF4B202A4DD84BFF6E7C361E42B377" bold="true" box="[199,321,1337,1362]" pageId="12" pageNumber="13">Diagnosis.</emphasis>
With the character states of the genus
<taxonomicName id="9DBBECB12A4DD84BFCA17C361C6AB377" authorityName="Latreille" authorityYear="1824" box="[776,873,1337,1362]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A4DD84BFCA17C361C6AB377" box="[776,873,1337,1362]" italics="true" pageId="12" pageNumber="13">Calleida</emphasis>
</taxonomicName>
in the widest sense, but from all Asiatic species known so far markedly characterized by the following peculiar characters: small-medium sized (L: 6.78.0 mm); pronotum slightly angulate near the middle, with heavy punctures on disc (
<figureCitation id="C2808BB72A4DD84BFBBF7C8E1B6AB3BF" box="[1046,1129,1409,1434]" captionStart="FIGURES 2229" captionStartId="13.[151,264,1822,1844]" captionTargetBox="[158,1420,311,1777]" captionTargetId="figure@13.[151,1436,301,1801]" captionTargetPageId="13" captionText="FIGURES 2229. Calleida puncticollis sp. n. 22 habitus and labels of holotype (male, Hubei, IZAS), scale bar = 2.0 mm; 23 habitus of paratype (female, Shaanxi, IZAS), scale bar = 2.0 mm; 24 pronotum (Shaanxi, China), [coarsely punctate], scale bar = 0.5 mm; 25 ovipositor (Hubei, China), scale bar = 0.1 mm; 26 female reproductive tract (Hubei, China), scale bar = 0.5 mm; 27 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm; 28 left and right parameres of aedeagus (holotype), scale bar = 0.2 mm; 29 male sternum IX (holotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302721/files/figure.png" pageId="12" pageNumber="13">fig. 24</figureCitation>
); apical labial palpomeres markedly securiform in both males and females (no sexual dimorphism); abdominal sternum VII not notched in either males and females, with eight setae in males (four on each side,
<figureCitation id="C2808BB72A4DD84BFC1A7CC61CECB3C7" box="[947,1007,1481,1506]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="12" pageNumber="13">fig. 3</figureCitation>
), ten or more setae in females (five to seven on each side,
<figureCitation id="C2808BB72A4DD84BFEDF7CE21EB0B023" box="[374,435,1517,1542]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="12" pageNumber="13">fig. 6</figureCitation>
); endophallus without copulatory pieces (
<figureCitation id="C2808BB72A4DD84BFC217CE21CD5B023" box="[904,982,1517,1542]" captionStart="FIGURES 2229" captionStartId="13.[151,264,1822,1844]" captionTargetBox="[158,1420,311,1777]" captionTargetId="figure@13.[151,1436,301,1801]" captionTargetPageId="13" captionText="FIGURES 2229. Calleida puncticollis sp. n. 22 habitus and labels of holotype (male, Hubei, IZAS), scale bar = 2.0 mm; 23 habitus of paratype (female, Shaanxi, IZAS), scale bar = 2.0 mm; 24 pronotum (Shaanxi, China), [coarsely punctate], scale bar = 0.5 mm; 25 ovipositor (Hubei, China), scale bar = 0.1 mm; 26 female reproductive tract (Hubei, China), scale bar = 0.5 mm; 27 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm; 28 left and right parameres of aedeagus (holotype), scale bar = 0.2 mm; 29 male sternum IX (holotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302721/files/figure.png" pageId="12" pageNumber="13">fig. 27</figureCitation>
).
</paragraph>
<paragraph id="5A0497322A4DD84BFF6E7F1E1FC6B0B0" blockId="12.[151,1436,977,2010]" pageId="12" pageNumber="13">
This new species, although a little similar to some members of the
<taxonomicName id="9DBBECB12A4DD84BFC7D7F1D1B78B00F" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[980,1147,1553,1578]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="splendidula">
<emphasis id="68CF4B202A4DD84BFC7D7F1D1B78B00F" box="[980,1147,1553,1578]" italics="true" pageId="12" pageNumber="13">C. splendidula</emphasis>
</taxonomicName>
species group in general appearance and coloration, such as
<taxonomicName id="9DBBECB12A4DD84BFD957F391DABB06B" box="[572,680,1589,1614]" class="Insecta" family="Tortricidae" genus="Cydia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="onoha">
<emphasis id="68CF4B202A4DD84BFD957F391DABB06B" box="[572,680,1589,1614]" italics="true" pageId="12" pageNumber="13">C. onoha</emphasis>
</taxonomicName>
, can be easily identified from all other Asiatic
<taxonomicName id="9DBBECB12A4DD84BFB4C7F3A1A45B06B" authorityName="Latreille" authorityYear="1824" box="[1253,1350,1589,1614]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A4DD84BFB4C7F3A1A45B06B" box="[1253,1350,1589,1614]" italics="true" pageId="12" pageNumber="13">Calleida</emphasis>
</taxonomicName>
by the presence of distinct, deep punctures on pronotum and the presence of additional setae on the abdominal sternum VII.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A4DD848FF6E7FAE1E30B7F5" lastPageId="15" lastPageNumber="16" pageId="12" pageNumber="13" type="description">
<paragraph id="5A0497322A4DD84BFF6E7FAE1A57B09C" blockId="12.[151,1436,977,2010]" box="[199,1364,1697,1722]" pageId="12" pageNumber="13">
<emphasis id="68CF4B202A4DD84BFF6E7FAE1E5AB09F" bold="true" box="[199,345,1697,1722]" pageId="12" pageNumber="13">Description.</emphasis>
<emphasis id="68CF4B202A4DD84BFEF67FAE1D18B09F" box="[351,539,1697,1722]" italics="true" pageId="12" pageNumber="13">General features</emphasis>
as in
<figureCitation id="C2808BB72A4DD84BFDF47FAE1DDDB09F" box="[605,734,1697,1722]" captionStart="FIGURES 2229" captionStartId="13.[151,264,1822,1844]" captionTargetBox="[158,1420,311,1777]" captionTargetId="figure@13.[151,1436,301,1801]" captionTargetPageId="13" captionText="FIGURES 2229. Calleida puncticollis sp. n. 22 habitus and labels of holotype (male, Hubei, IZAS), scale bar = 2.0 mm; 23 habitus of paratype (female, Shaanxi, IZAS), scale bar = 2.0 mm; 24 pronotum (Shaanxi, China), [coarsely punctate], scale bar = 0.5 mm; 25 ovipositor (Hubei, China), scale bar = 0.1 mm; 26 female reproductive tract (Hubei, China), scale bar = 0.5 mm; 27 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm; 28 left and right parameres of aedeagus (holotype), scale bar = 0.2 mm; 29 male sternum IX (holotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302721/files/figure.png" pageId="12" pageNumber="13">figs. 22, 23</figureCitation>
. Small-medium sized: L:
<quantity id="9D433AD72A4DD84BFC567FAE1B8AB09C" box="[1023,1161,1697,1722]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.2" metricValueMin="6.8" pageId="12" pageNumber="13" unit="mm" value="7.5" valueMax="8.2" valueMin="6.8">6.88.2 mm</quantity>
; TL:
<quantity id="9D433AD72A4DD84BFB6C7FAE1A4CB09C" box="[1221,1359,1697,1722]" metricMagnitude="-3" metricUnit="m" metricValue="7.15" metricValueMax="7.8" metricValueMin="6.5" pageId="12" pageNumber="13" unit="mm" value="7.15" valueMax="7.8" valueMin="6.5">6.57.8 mm</quantity>
.
</paragraph>
<paragraph id="5A0497322A4DD84BFF6E7FCA1DDFB16F" blockId="12.[151,1436,977,2010]" pageId="12" pageNumber="13">
<emphasis id="68CF4B202A4DD84BFF6E7FCA1E15B0FB" box="[199,278,1733,1758]" italics="true" pageId="12" pageNumber="13">Colour</emphasis>
: Head reddish brown, palpi and antennae yellow reddish; pronotum reddish brown, somewhat paler at lateral margins; scutellum reddish brown; elytra green to greenish-cupreous, with strong metallic reflection, or in some specimens greenish with reddish cupreous reflection; suture and lateral margins reddish brown; epipleura reddish brown; ventral side and legs reddish brown.
</paragraph>
<paragraph id="5A0497322A4DD84BFF6E7E591CFBB1B7" blockId="12.[151,1436,977,2010]" pageId="12" pageNumber="13">
<emphasis id="68CF4B202A4DD84BFF6E7E591D02B14B" box="[199,513,1877,1902]" italics="true" pageId="12" pageNumber="13">Lustre and microsculpture:</emphasis>
microsculpture vanished over the entire dorsal surface; pronotum shiny, with distinctly coarse and deep punctures; elytra shiny, with strong metallic lustre.
</paragraph>
<paragraph id="5A0497322A4DD84AFF6E7E921D50B6DD" blockId="12.[151,1436,977,2010]" lastBlockId="13.[151,1436,151,248]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<emphasis id="68CF4B202A4DD84BFF6E7E921E07B193" box="[199,260,1949,1974]" italics="true" pageId="12" pageNumber="13">Head</emphasis>
with evident dorsal punctures; genae very short, not swollen, obliquely and gradually narrowed towards the neck; genae as long as the half length of eyes; vertex slightly convex; frons more sparsely punctate than vertex, with irregular, distinct or very faint wrinkles; supraorbital furrows deep, vanished before the hind edge of eyes; antennae reaching the basal fifth of elytra; terminal labial palpomere strongly securiform in both male and female, terminal maxillary palpomere fusiform.
</paragraph>
<caption id="0EC4C7BA2A4CD84AFF3E7E111BD4B18A" httpUri="https://zenodo.org/record/1302721/files/figure.png" pageId="13" pageNumber="14" startId="13.[151,264,1822,1844]" targetBox="[158,1420,311,1777]" targetPageId="13">
<paragraph id="5A0497322A4CD84AFF3E7E111BD4B18A" blockId="13.[151,1436,1822,1967]" pageId="13" pageNumber="14">
<emphasis id="68CF4B202A4CD84AFF3E7E111D31B111" bold="true" box="[151,562,1822,1844]" pageId="13" pageNumber="14">
FIGURES 2229.
<taxonomicName id="9DBBECB12A4CD84AFEF27E111D31B111" box="[347,562,1822,1844]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="puncticollis" status="sp. nov.">
<emphasis id="68CF4B202A4CD84AFEF27E111D31B111" bold="true" box="[347,562,1822,1844]" italics="true" pageId="13" pageNumber="14">Calleida puncticollis</emphasis>
</taxonomicName>
</emphasis>
<taxonomicNameLabel id="73FCF65B2A4CD84AFD907E101D73B111" box="[569,624,1823,1844]" pageId="13" pageNumber="14" rank="species">sp. n.</taxonomicNameLabel>
<emphasis id="68CF4B202A4CD84AFDDE7E111D92B111" bold="true" box="[631,657,1822,1844]" pageId="13" pageNumber="14">22</emphasis>
habitus and labels of holotype (male, Hubei, IZAS), scale bar = 2.0 mm;
<emphasis id="68CF4B202A4CD84AFA2A7E111A9FB111" bold="true" box="[1411,1436,1822,1844]" pageId="13" pageNumber="14">23</emphasis>
habitus of paratype (female, Shaanxi, IZAS), scale bar = 2.0 mm;
<emphasis id="68CF4B202A4CD84AFC9D7E321C4DB176" bold="true" box="[820,846,1853,1875]" pageId="13" pageNumber="14">24</emphasis>
pronotum (Shaanxi, China), [coarsely punctate], scale bar = 0.5 mm;
<emphasis id="68CF4B202A4CD84AFEAE7E531E23B157" bold="true" box="[263,288,1884,1906]" pageId="13" pageNumber="14">25</emphasis>
ovipositor (Hubei, China), scale bar = 0.1 mm;
<emphasis id="68CF4B202A4CD84AFCA17E531C21B157" bold="true" box="[776,802,1884,1906]" pageId="13" pageNumber="14">26</emphasis>
female reproductive tract (Hubei, China), scale bar = 0.5 mm;
<emphasis id="68CF4B202A4CD84AFF3E7E741FB2B1B4" bold="true" box="[151,177,1915,1937]" pageId="13" pageNumber="14">27</emphasis>
median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm;
<emphasis id="68CF4B202A4CD84AFB527E741A16B1B4" bold="true" box="[1275,1301,1915,1937]" pageId="13" pageNumber="14">28</emphasis>
left and right parameres of aedeagus (holotype), scale bar = 0.2 mm;
<emphasis id="68CF4B202A4CD84AFD607E961DE1B18A" bold="true" box="[713,738,1945,1967]" pageId="13" pageNumber="14">29</emphasis>
male sternum IX (holotype), scale bar = 0.5 mm.
</paragraph>
</caption>
<caption id="0EC4C7BA2A4FD849FF3E7DEE1B1BB2D2" box="[151,1048,1249,1271]" httpUri="https://zenodo.org/record/1302723/files/figure.png" pageId="14" pageNumber="15" startId="14.[151,209,1249,1271]" targetBox="[151,1436,195,1227]" targetPageId="14">
<paragraph id="5A0497322A4FD849FF3E7DEE1B1BB2D2" blockId="14.[151,1048,1249,1271]" box="[151,1048,1249,1271]" pageId="14" pageNumber="15">
<emphasis id="68CF4B202A4FD849FF3E7DEE1FE7B2D2" bold="true" box="[151,228,1249,1271]" pageId="14" pageNumber="15">MAP 2</emphasis>
Distributions for species in the
<taxonomicName id="9DBBECB12A4FD849FD8E7DED1DB9B2D2" box="[551,698,1249,1271]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="68CF4B202A4FD849FD8E7DED1DB9B2D2" box="[551,698,1249,1271]" italics="true" pageId="14" pageNumber="15">C. puncticollis</emphasis>
</taxonomicName>
group:
<taxonomicName id="9DBBECB12A4FD849FCA07DED1C9EB2D2" box="[777,925,1249,1271]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="puncticollis" status="sp. nov.">
<emphasis id="68CF4B202A4FD849FCA07DED1C9EB2D2" box="[777,925,1249,1271]" italics="true" pageId="14" pageNumber="15">C. puncticollis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73FCF65B2A4FD849FC0A7DED1CD9B2D2" box="[931,986,1250,1271]" pageId="14" pageNumber="15" rank="species">sp. n.</taxonomicNameLabel>
(red).
</paragraph>
</caption>
<paragraph id="5A0497322A4FD849FF6E7C2A1CBCB3D5" blockId="14.[151,1436,1315,2024]" pageId="14" pageNumber="15">
<emphasis id="68CF4B202A4FD849FF6E7C2A1E35B319" box="[199,310,1317,1340]" italics="true" pageId="14" pageNumber="15">Pronotum</emphasis>
(
<figureCitation id="C2808BB72A4FD849FEE17C2C1E96B319" box="[328,405,1315,1340]" captionStart="FIGURES 2229" captionStartId="13.[151,264,1822,1844]" captionTargetBox="[158,1420,311,1777]" captionTargetId="figure@13.[151,1436,301,1801]" captionTargetPageId="13" captionText="FIGURES 2229. Calleida puncticollis sp. n. 22 habitus and labels of holotype (male, Hubei, IZAS), scale bar = 2.0 mm; 23 habitus of paratype (female, Shaanxi, IZAS), scale bar = 2.0 mm; 24 pronotum (Shaanxi, China), [coarsely punctate], scale bar = 0.5 mm; 25 ovipositor (Hubei, China), scale bar = 0.1 mm; 26 female reproductive tract (Hubei, China), scale bar = 0.5 mm; 27 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm; 28 left and right parameres of aedeagus (holotype), scale bar = 0.2 mm; 29 male sternum IX (holotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302721/files/figure.png" pageId="14" pageNumber="15">fig. 24</figureCitation>
) hexagonal (ratio PW/PL 1.211.27), its greatest width at about the anterior third; lateral margins forming an obtuse angle at the greatest width, slightly sinuate before the posterior angles; posterior angles almost rectangular, not distinctly pointed; lateral furrows narrow, somewhat raised; disc slightly convex, with some transverse wrinkles close to the median furrow; median furrow distinct, not reaching the anterior and basal margins; surface of the disc with distinctly coarse punctures, larger and denser along the median line, in basal foveae, in the pre-basal and anterior areas, and scattered at sides of disc.
</paragraph>
<paragraph id="5A0497322A4FD849FF6E7CF41C6EB0A5" blockId="14.[151,1436,1315,2024]" pageId="14" pageNumber="15">
<emphasis id="68CF4B202A4FD849FF6E7CF41E0EB031" box="[199,269,1531,1556]" italics="true" pageId="14" pageNumber="15">Elytra</emphasis>
relatively short (ratio EL/EW: 1.561.62), with basal ridge extending to scutellar stria; striae moderately deep, punctate, the punctures gradually weakened to the apical part; intervals flat, finely and sparsely punctate; eighth interval slightly tumid near the apex; umbilicate series of 15 pores along eighth stria; apical truncation straight or slightly concave; outer apical angle obtusely rounded.
</paragraph>
<paragraph id="5A0497322A4FD849FF6E7F841E11B135" blockId="14.[151,1436,1315,2024]" pageId="14" pageNumber="15">
<emphasis id="68CF4B202A4FD849FF6E7F841E4CB081" box="[199,335,1675,1700]" italics="true" pageId="14" pageNumber="15">Ventral side</emphasis>
: Prosternum, lateral area of metasternum, and metepisterna with short setae; abdominal sterna with sparse accessory setae, denser on sternum VII; abdominal sternum VII with four setae on each side in males, five to seven setae on each side in females; apical margin of sternum VII not notched in both males and females (
<figureCitation id="C2808BB72A4FD849FF367FF81E07B135" box="[159,260,1783,1808]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="14" pageNumber="15">figs. 3, 6</figureCitation>
).
</paragraph>
<paragraph id="5A0497322A4FD849FF6E7E141DEBB185" blockId="14.[151,1436,1315,2024]" pageId="14" pageNumber="15">
<emphasis id="68CF4B202A4FD849FF6E7E141E6EB111" box="[199,365,1819,1844]" italics="true" pageId="14" pageNumber="15">Male genitalia</emphasis>
: median lobe (
<figureCitation id="C2808BB72A4FD849FDB37E141D65B111" box="[538,614,1819,1844]" captionStart="FIGURES 2229" captionStartId="13.[151,264,1822,1844]" captionTargetBox="[158,1420,311,1777]" captionTargetId="figure@13.[151,1436,301,1801]" captionTargetPageId="13" captionText="FIGURES 2229. Calleida puncticollis sp. n. 22 habitus and labels of holotype (male, Hubei, IZAS), scale bar = 2.0 mm; 23 habitus of paratype (female, Shaanxi, IZAS), scale bar = 2.0 mm; 24 pronotum (Shaanxi, China), [coarsely punctate], scale bar = 0.5 mm; 25 ovipositor (Hubei, China), scale bar = 0.1 mm; 26 female reproductive tract (Hubei, China), scale bar = 0.5 mm; 27 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm; 28 left and right parameres of aedeagus (holotype), scale bar = 0.2 mm; 29 male sternum IX (holotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302721/files/figure.png" pageId="14" pageNumber="15">fig. 27</figureCitation>
) pleuropic left, slightly curved, strongly expanded in dorsal view in the middle, narrow in lateral view, slightly expanded before the apical orifice; apical lamina flat, wider than length, fully rounded at apex; endophallus without any chitinized piece; left paramere strongly depressed in dorsal side; right paramere slightly emarginated at apex (
<figureCitation id="C2808BB72A4FD849FD247E881DD8B185" box="[653,731,1927,1952]" captionStart="FIGURES 2229" captionStartId="13.[151,264,1822,1844]" captionTargetBox="[158,1420,311,1777]" captionTargetId="figure@13.[151,1436,301,1801]" captionTargetPageId="13" captionText="FIGURES 2229. Calleida puncticollis sp. n. 22 habitus and labels of holotype (male, Hubei, IZAS), scale bar = 2.0 mm; 23 habitus of paratype (female, Shaanxi, IZAS), scale bar = 2.0 mm; 24 pronotum (Shaanxi, China), [coarsely punctate], scale bar = 0.5 mm; 25 ovipositor (Hubei, China), scale bar = 0.1 mm; 26 female reproductive tract (Hubei, China), scale bar = 0.5 mm; 27 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm; 28 left and right parameres of aedeagus (holotype), scale bar = 0.2 mm; 29 male sternum IX (holotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302721/files/figure.png" pageId="14" pageNumber="15">fig. 28</figureCitation>
).
</paragraph>
<paragraph id="5A0497322A4FD848FF6E7EA41D1AB765" blockId="14.[151,1436,1315,2024]" lastBlockId="15.[151,1437,151,464]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="68CF4B202A4FD849FF6E7EA41E89B1E1" box="[199,394,1963,1988]" italics="true" pageId="14" pageNumber="15">Female genitalia</emphasis>
(reproductive tract
<figureCitation id="C2808BB72A4FD849FDDC7EA41DC1B1E1" box="[629,706,1963,1988]" captionStart="FIGURES 2229" captionStartId="13.[151,264,1822,1844]" captionTargetBox="[158,1420,311,1777]" captionTargetId="figure@13.[151,1436,301,1801]" captionTargetPageId="13" captionText="FIGURES 2229. Calleida puncticollis sp. n. 22 habitus and labels of holotype (male, Hubei, IZAS), scale bar = 2.0 mm; 23 habitus of paratype (female, Shaanxi, IZAS), scale bar = 2.0 mm; 24 pronotum (Shaanxi, China), [coarsely punctate], scale bar = 0.5 mm; 25 ovipositor (Hubei, China), scale bar = 0.1 mm; 26 female reproductive tract (Hubei, China), scale bar = 0.5 mm; 27 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm; 28 left and right parameres of aedeagus (holotype), scale bar = 0.2 mm; 29 male sternum IX (holotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302721/files/figure.png" pageId="14" pageNumber="15">fig. 26</figureCitation>
and ovipositor
<figureCitation id="C2808BB72A4FD849FCD77EA41CD3B1E1" box="[894,976,1963,1988]" captionStart="FIGURES 2229" captionStartId="13.[151,264,1822,1844]" captionTargetBox="[158,1420,311,1777]" captionTargetId="figure@13.[151,1436,301,1801]" captionTargetPageId="13" captionText="FIGURES 2229. Calleida puncticollis sp. n. 22 habitus and labels of holotype (male, Hubei, IZAS), scale bar = 2.0 mm; 23 habitus of paratype (female, Shaanxi, IZAS), scale bar = 2.0 mm; 24 pronotum (Shaanxi, China), [coarsely punctate], scale bar = 0.5 mm; 25 ovipositor (Hubei, China), scale bar = 0.1 mm; 26 female reproductive tract (Hubei, China), scale bar = 0.5 mm; 27 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm; 28 left and right parameres of aedeagus (holotype), scale bar = 0.2 mm; 29 male sternum IX (holotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302721/files/figure.png" pageId="14" pageNumber="15">fig. 25</figureCitation>
): spermatheca clubbed, longer than the pedicel, base without any additional structure, surface strongly whorled; spermathecal pedicel with the basal two thirds expanded, apical half slender; spermathecal gland duct inserted ventrally at base of spermatheca, a little shorter than spermatheca; glandular area about same length as gland duct, base with small and protuberant atrium. Gonocoxite I of ovipositor very wide, triangular; gonocoxite II small, subulate, at base three times longer than wide; inner margin with several setae, extending from the basal third to apex; outer and inner margin straight; apex with short membranous extension.
</paragraph>
<paragraph id="5A0497322A4ED848FF6E78431E30B7F5" blockId="15.[151,1437,151,464]" pageId="15" pageNumber="16">
<emphasis id="68CF4B202A4ED848FF6E78431D95B740" bold="true" box="[199,662,332,357]" pageId="15" pageNumber="16">Geographical distribution and habitat.</emphasis>
A widely distributed but apparently very rare species in
<collectingCountry id="22ACD7A22A4ED848FAB178431A58B740" box="[1304,1371,332,357]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
(
<figureCitation id="C2808BB72A4ED848FAC378431FA4B7AD" captionStart="MAP 2" captionStartId="14.[151,209,1249,1271]" captionTargetBox="[151,1436,195,1227]" captionTargetId="figure@14.[151,1436,195,1227]" captionTargetPageId="14" captionText="MAP 2 Distributions for species in the C. puncticollis group: C. puncticollis sp. n. (red)." httpUri="https://zenodo.org/record/1302723/files/figure.png" pageId="15" pageNumber="16">map 2</figureCitation>
), known from
<collectingRegion id="987F59D02A4ED848FEFF78601EB0B7AD" box="[342,435,367,392]" country="China" name="Shaanxi" pageId="15" pageNumber="16">Shaanxi</collectingRegion>
,
<collectingRegion id="987F59D02A4ED848FE6878601D0BB7AD" box="[449,520,367,392]" country="China" name="Hubei" pageId="15" pageNumber="16">Hubei</collectingRegion>
,
<collectingRegion id="987F59D02A4ED848FDBE787F1D67B7A2" box="[535,612,368,391]" country="China" name="Hunan" pageId="15" pageNumber="16">Hunan</collectingRegion>
,
<collectingRegion id="987F59D02A4ED848FDDB78601DD1B7AD" box="[626,722,367,392]" country="China" name="Guangxi" pageId="15" pageNumber="16">Guangxi</collectingRegion>
and
<collectingRegion id="987F59D02A4ED848FCA678601C54B7AD" box="[783,855,367,392]" country="China" name="Fujian" pageId="15" pageNumber="16">Fujian</collectingRegion>
provinces. The sampled specimens show that this species inhabits in subtropical moist broadleaf forests, and tends to prefer old-growth forest in low elevation (less than
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).
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>

View file

@ -0,0 +1,105 @@
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<emphasis id="68CF4B202A4DD84BFF3E79971D58B697" bold="true" box="[151,603,151,178]" pageId="12" pageNumber="13">
2.
<taxonomicName id="9DBBECB12A4DD84BFF1D79981EAFB694" ID-CoL="8M9ZJ" box="[180,428,151,177]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis id="68CF4B202A4DD84BFF1D79981EAFB694" bold="true" box="[180,428,151,177]" italics="true" pageId="12" pageNumber="13">Calleida puncticollis</emphasis>
</taxonomicName>
species group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A4DD84BFF3E79D11E2AB512" pageId="12" pageNumber="13" type="description">
<paragraph id="5A0497322A4DD84BFF3E79D11D38B78E" blockId="12.[151,1436,222,823]" pageId="12" pageNumber="13">
<emphasis id="68CF4B202A4DD84BFF3E79D11EA3B6D2" bold="true" box="[151,416,222,247]" pageId="12" pageNumber="13">Diagnostic characters</emphasis>
: With the character states of
<taxonomicName id="9DBBECB12A4DD84BFCA579D11C6FB6D2" authorityName="Latreille" authorityYear="1824" box="[780,876,222,247]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A4DD84BFCA579D11C6FB6D2" box="[780,876,222,247]" italics="true" pageId="12" pageNumber="13">Calleida</emphasis>
</taxonomicName>
(in the widest sense, see above), restricted or amplified as follows: small-sized species (L
<quantity id="9D433AD72A4DD84BFD3A780D1C1CB73E" box="[659,799,258,283]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.2" metricValueMin="6.8" pageId="12" pageNumber="13" unit="mm" value="7.5" valueMax="8.2" valueMin="6.8">6.88.2 mm</quantity>
). Head and pronotum brownish, elytra metallic green to cupreous, without reddish patch. Apical labial palpomeres markedly securiform in both male and female. Pronotal disc heavily punctate (
<figureCitation id="C2808BB72A4DD84BFE3D78451EE0B746" box="[404,483,330,355]" captionStart="FIGURES 2229" captionStartId="13.[151,264,1822,1844]" captionTargetBox="[158,1420,311,1777]" captionTargetId="figure@13.[151,1436,301,1801]" captionTargetPageId="13" captionText="FIGURES 2229. Calleida puncticollis sp. n. 22 habitus and labels of holotype (male, Hubei, IZAS), scale bar = 2.0 mm; 23 habitus of paratype (female, Shaanxi, IZAS), scale bar = 2.0 mm; 24 pronotum (Shaanxi, China), [coarsely punctate], scale bar = 0.5 mm; 25 ovipositor (Hubei, China), scale bar = 0.1 mm; 26 female reproductive tract (Hubei, China), scale bar = 0.5 mm; 27 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm; 28 left and right parameres of aedeagus (holotype), scale bar = 0.2 mm; 29 male sternum IX (holotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302721/files/figure.png" pageId="12" pageNumber="13">fig. 24</figureCitation>
). Abdominal sternum VII not notched at apex both in males and females (
<figureCitation id="C2808BB72A4DD84BFA8E78451A8DB746" box="[1319,1422,330,355]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="12" pageNumber="13">figs. 3, 6</figureCitation>
), with eight setae in males, ten or more setae in females. Median lobe of aedeagus stout, endophallus without any chitinized copulatory pieces (
<figureCitation id="C2808BB72A4DD84BFE49789D1D2DB78E" box="[480,558,402,427]" captionStart="FIGURES 2229" captionStartId="13.[151,264,1822,1844]" captionTargetBox="[158,1420,311,1777]" captionTargetId="figure@13.[151,1436,301,1801]" captionTargetPageId="13" captionText="FIGURES 2229. Calleida puncticollis sp. n. 22 habitus and labels of holotype (male, Hubei, IZAS), scale bar = 2.0 mm; 23 habitus of paratype (female, Shaanxi, IZAS), scale bar = 2.0 mm; 24 pronotum (Shaanxi, China), [coarsely punctate], scale bar = 0.5 mm; 25 ovipositor (Hubei, China), scale bar = 0.1 mm; 26 female reproductive tract (Hubei, China), scale bar = 0.5 mm; 27 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm; 28 left and right parameres of aedeagus (holotype), scale bar = 0.2 mm; 29 male sternum IX (holotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302721/files/figure.png" pageId="12" pageNumber="13">fig. 27</figureCitation>
).
</paragraph>
<paragraph id="5A0497322A4DD84BFF6E78B91A35B432" blockId="12.[151,1436,222,823]" pageId="12" pageNumber="13">
<emphasis id="68CF4B202A4DD84BFF6E78B91E86B7EA" bold="true" box="[199,389,438,463]" pageId="12" pageNumber="13">Taxonomic note</emphasis>
: Only one species,
<taxonomicName id="9DBBECB12A4DD84BFDCD78B81C0AB7EA" box="[612,777,438,463]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="puncticollis" status="sp. nov.">
<emphasis id="68CF4B202A4DD84BFDCD78B81C0AB7EA" box="[612,777,438,463]" italics="true" pageId="12" pageNumber="13">C. puncticollis</emphasis>
</taxonomicName>
<emphasis id="68CF4B202A4DD84BFCB978B91C53B7EA" bold="true" box="[784,848,438,463]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="73FCF65B2A4DD84BFCB978B91C53B7EA" box="[784,848,438,463]" pageId="12" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, is attributed to this group. This species is very rare in collections but recorded from several provinces in central and eastern China (
<figureCitation id="C2808BB72A4DD84BFBBF78D41B63B7D6" box="[1046,1120,475,499]" captionStart="MAP 2" captionStartId="14.[151,209,1249,1271]" captionTargetBox="[151,1436,195,1227]" captionTargetId="figure@14.[151,1436,195,1227]" captionTargetPageId="14" captionText="MAP 2 Distributions for species in the C. puncticollis group: C. puncticollis sp. n. (red)." httpUri="https://zenodo.org/record/1302723/files/figure.png" pageId="12" pageNumber="13">map 2</figureCitation>
). The systematic position of this species group is rather isolated in the Asiatic fauna because of the peculiar characters of endophallus.
</paragraph>
<paragraph id="5A0497322A4DD84BFF6E7B2D1E2AB512" blockId="12.[151,1436,222,823]" pageId="12" pageNumber="13">
The following peculiar characters isolate this group from all other known Asiatic
<taxonomicName id="9DBBECB12A4DD84BFBC57B2D1BCEB41E" authorityName="Latreille" authorityYear="1824" box="[1132,1229,546,571]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A4DD84BFBC57B2D1BCEB41E" box="[1132,1229,546,571]" italics="true" pageId="12" pageNumber="13">Calleida</emphasis>
</taxonomicName>
species known so far: abdominal sternum VII in male not notched, and endophallus without copulatory pieces. In all other Asiatic
<taxonomicName id="9DBBECB12A4DD84BFF3E7B651FF4B4A6" authorityName="Latreille" authorityYear="1824" box="[151,247,618,643]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A4DD84BFF3E7B651FF4B4A6" box="[151,247,618,643]" italics="true" pageId="12" pageNumber="13">Calleida</emphasis>
</taxonomicName>
, the male anal sternum is distinctly notched, and the endophallus has distinct chitinized copulatory pieces, composed of long flagellum or two short pieces. This is also the only Asiatic species with abdominal sternum VII with more than two apical setae in males (more than four in females) not included in the
<taxonomicName id="9DBBECB12A4DD84BFB1E7BBD1A9FB4EE" authorityName="Heller in Mindanao" authorityYear="1921" box="[1207,1436,690,715]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="discoidalis">
<emphasis id="68CF4B202A4DD84BFB1E7BBD1A9FB4EE" box="[1207,1436,690,715]" italics="true" pageId="12" pageNumber="13">Calleida discoidalis</emphasis>
</taxonomicName>
species group (see Key to the species groups), which will be treated in a separate contribution. However, the different external and male sexual characters mentioned above suggested that these two groups may not closely related at all.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,111 @@
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<mods:title id="FA7FC27DC367F28724CA5A066FB63DCA">Revision of the Oriental species of Calleida Latreille (sensu lato). Part 1: Introduction, groups of species, and species of six species groups (Coleoptera: Carabidae: Lebiini)</mods:title>
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<subSubSection id="12A1C4B92A4ED848FF3E7B111D4CB41D" box="[151,591,542,568]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="5A0497322A4ED848FF3E7B111D4CB41D" blockId="15.[151,591,542,568]" box="[151,591,542,568]" pageId="15" pageNumber="16">
<heading id="014C205E2A4ED848FF3E7B111D4CB41D" bold="true" box="[151,591,542,568]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<emphasis id="68CF4B202A4ED848FF3E7B111D4CB41D" bold="true" box="[151,591,542,568]" pageId="15" pageNumber="16">
3.
<taxonomicName id="9DBBECB12A4ED848FF1D7B111E9CB41D" ID-CoL="5WTZV" authorityName="Heller in Mindanao" authorityYear="1921" box="[180,415,542,568]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="discoidalis">
<emphasis id="68CF4B202A4ED848FF1D7B111E9CB41D" bold="true" box="[180,415,542,568]" italics="true" pageId="15" pageNumber="16">Calleida discoidalis</emphasis>
</taxonomicName>
species group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A4ED848FF3E7B6A1FECB2BF" pageId="15" pageNumber="16" type="description">
<paragraph id="5A0497322A4ED848FF3E7B6A1E60B4CF" blockId="15.[151,1436,613,1178]" pageId="15" pageNumber="16">
<emphasis id="68CF4B202A4ED848FF3E7B6A1E99B45B" bold="true" box="[151,410,613,638]" pageId="15" pageNumber="16">Diagnostic characters</emphasis>
: With the character states of Asiatic
<taxonomicName id="9DBBECB12A4ED848FC927B6A1C9FB45B" authorityName="Latreille" authorityYear="1824" box="[827,924,613,638]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A4ED848FC927B6A1C9FB45B" box="[827,924,613,638]" italics="true" pageId="15" pageNumber="16">Calleida</emphasis>
</taxonomicName>
(in the widest sense, see above), restricted as follows: abdominal sternum VII with four or more setae in males (two or more on each side), six or more setae in females (three or more on each side); sternum VII notched in males (
<figureCitation id="C2808BB72A4ED848FC7D7BA21B40B4E3" box="[980,1091,685,710]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="15" pageNumber="16">figs. 2, 5</figureCitation>
); endophallus with two short copulatory pieces.
</paragraph>
<paragraph id="5A0497322A4ED848FF6E7BFA1CA1B573" blockId="15.[151,1436,613,1178]" pageId="15" pageNumber="16">
<emphasis id="68CF4B202A4ED848FF6E7BFA1E87B52B" bold="true" box="[199,388,757,782]" pageId="15" pageNumber="16">Taxonomic note</emphasis>
: Two already described species,
<taxonomicName id="9DBBECB12A4ED848FD5F7BF91CDEB52B" authority="Heller" authorityName="Heller in Mindanao" authorityYear="1921" box="[758,989,757,782]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="discoidalis">
<emphasis id="68CF4B202A4ED848FD5F7BF91C93B52B" box="[758,912,757,782]" italics="true" pageId="15" pageNumber="16">C. discoidalis</emphasis>
Heller
</taxonomicName>
and
<taxonomicName id="9DBBECB12A4ED848FBBD7BF91BFFB52B" authority="Jedlicka" authorityName="Jedlicka" authorityYear="1953" box="[1044,1276,757,782]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="fukiensis">
<emphasis id="68CF4B202A4ED848FBBD7BF91B95B52B" box="[1044,1174,757,782]" italics="true" pageId="15" pageNumber="16">C. fukiensis</emphasis>
Jedlička
</taxonomicName>
, are attributed to this group, and several new species will be described in our next contribution (Shi &amp; Casale, submitted). Species of this group are distributed in
<collectingCountry id="22ACD7A22A4ED848FE5B7A321D3BB573" box="[498,568,829,854]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
,
<collectingCountry id="22ACD7A22A4ED848FDEA7A311D7FB570" box="[579,636,830,853]" name="Laos" pageId="15" pageNumber="16">Laos</collectingCountry>
,
<collectingCountry id="22ACD7A22A4ED848FD2E7A321DE5B573" box="[647,742,829,854]" name="Vietnam" pageId="15" pageNumber="16">Vietnam</collectingCountry>
and
<collectingCountry id="22ACD7A22A4ED848FCB57A321C9DB573" box="[796,926,829,854]" name="Philippines" pageId="15" pageNumber="16">Philippines</collectingCountry>
.
</paragraph>
<paragraph id="5A0497322A4ED848FF6E7A6E1C24B20B" blockId="15.[151,1436,613,1178]" pageId="15" pageNumber="16">
This species group is very probably not monophyletic, and relationships amongst species are unclear. In species of this group the only similarity is the presence of two short copulatory pieces in endophallus, a character common with many Asiatic, African and Madagascan
<taxonomicName id="9DBBECB12A4ED848FCA17AA61C6BB5E7" authorityName="Latreille" authorityYear="1824" box="[776,872,937,962]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A4ED848FCA17AA61C6BB5E7" box="[776,872,937,962]" italics="true" pageId="15" pageNumber="16">Calleida</emphasis>
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species. The group is so far defined by one only external morphological feature (abdominal sternum VII multisetose), which could be demonstrated as not synapomorphic in the Asiatic species: in fact, this character is also present in
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<emphasis id="68CF4B202A4ED848FBAC7AFE1B65B22F" box="[1029,1126,1009,1034]" italics="true" pageId="15" pageNumber="16">Calleida</emphasis>
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species of several different groups in Africa, Madagascar and the Neotropical Region.
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Nevertheless, at moment the multiple setation of the abdominal sternum VII is a valid diagnostic feature able to isolate this “group of convenience” from all other species of Asia (except the
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<emphasis id="68CF4B202A4ED848FBB77D521BA2B253" box="[1054,1185,1117,1142]" italics="true" pageId="15" pageNumber="16">puncticollis</emphasis>
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species group, treated before).
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4.
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<emphasis id="68CF4B202A4ED848FF1D7DE21E83B322" bold="true" box="[180,384,1261,1287]" italics="true" pageId="15" pageNumber="16">Calleida lativittis</emphasis>
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species group
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<emphasis id="68CF4B202A4ED848FF3E7C3B1E9AB368" bold="true" box="[151,409,1332,1357]" pageId="15" pageNumber="16">Diagnostic characters</emphasis>
: Medium-sized, elongate, slender species: L
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. Head, pronotum, lateral margins of elytra and ventral side yellow reddish; elytra metallic green or bluish green, with or without a large reddish patch extended from scutellum to apex. Anterior angles of pronotum with several short, erect setae; pronotum, elytral intervals and abdominal sterna punctate, glabrous or with distinct pubescence. Elytra elongate and narrow (ratio EL/EW: 1.762.02), parallel-sided or slightly widened at the posterior fifth. Abdominal sternum VII with one seta on each side in males, two setae in females. Median lobe of aedeagus simple, without prominent lobe on the ventral side; endophallus with two narrow, spine-like copulatory pieces.
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<emphasis id="68CF4B202A4ED848FF6E7F201E87B06D" bold="true" box="[199,388,1583,1608]" pageId="15" pageNumber="16">Taxonomic note</emphasis>
: Three species, including one new species, are attributed to this group, distributed from
<collectingCountry id="22ACD7A22A4ED848FACB7F201A9FB06D" box="[1378,1436,1583,1608]" name="India" pageId="15" pageNumber="16">India</collectingCountry>
to northern
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and
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(
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). Species of this group are rare and little known, and strictly allopatric according to examined materials (
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).
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This species group is considered to be monophyletic, for the similarities in the markedly elongate elytra, anterior angles of pronotum setose, and the general habitus. So far, the complete male genitalia of one only species have been examined (
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<emphasis id="68CF4B202A4ED848FE237FEA1D05B0D8" box="[394,518,1764,1789]" italics="true" pageId="15" pageNumber="16">C. lativittis</emphasis>
Chaudoir
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), and some details of two immature males of another species (
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<emphasis id="68CF4B202A4ED848FA837FEA1A9FB0D8" box="[1322,1436,1764,1789]" italics="true" pageId="15" pageNumber="16">C. excelsa</emphasis>
Bates
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) from Yunnan, which may suggest relationships with taxa of the
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Key to species of
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species group
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<paragraph id="5A0497322A4ED848FF3E7EB51A9BB1C9" blockId="15.[151,1435,1908,2028]" pageId="15" pageNumber="16">1. Elytra metallic bluish green; disc without reddish patch, only sutural interval reddish (Fig. 42). Elytra sub-convex, parallel sided. Dorsal pubescence reduced to a few, short setae at the base of pronotum. S India (Coimbatore, Bangalore)...........</paragraph>
</footnote>
</subSubSection>
<subSubSection id="12A1C4B92A51D857FF4A79981A9FB7B2" pageId="16" pageNumber="17" type="key">
<paragraph id="5A0497322A51D857FF4A79981A9FB688" blockId="16.[151,1437,151,407]" box="[227,1436,151,173]" pageId="16" pageNumber="17">
......................................................................... [5]
<taxonomicName id="9DBBECB12A51D857FBDA79981A5EB689" ID-CoL="8MCZV" authority="Casale &amp; Shi" authorityName="Casale &amp; Shi" box="[1139,1373,151,172]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="3" phylum="Arthropoda" rank="species" species="jelineki" status="sp. nov.">
<emphasis id="68CF4B202A51D857FBDA79981BD5B689" box="[1139,1238,151,172]" italics="true" pageId="16" pageNumber="17">C. jelineki</emphasis>
Casale &amp; Shi
</taxonomicName>
,
<emphasis id="68CF4B202A51D857FACF79981A9FB688" bold="true" box="[1382,1436,151,173]" pageId="16" pageNumber="17">sp. n.</emphasis>
</paragraph>
<paragraph id="5A0497322A51D857FF3E79BA1A9FB6C2" blockId="16.[151,1437,151,407]" pageId="16" pageNumber="17">- Elytra metallic green, disc with large reddish patch extended from the base to apex on the inner three to five intervals. Elytra depressed, narrow but slightly widened at the posterior fifth. Pronotal disc evenly pubescent or glabrous................. 2</paragraph>
<paragraph id="5A0497322A51D857FF3E79E01A98B71A" blockId="16.[151,1437,151,407]" pageId="16" pageNumber="17">
2. Dorsal side with dense pubescence on the head, pronotum and elytral intervals. Femora blackish at apical third or less. Head wide and depressed. Pronotum transverse-cordate, widened in the middle, with lateral furrows wide and markedly flattened. NE
<collectingCountry id="22ACD7A22A51D857FEAE78251E38B71A" box="[263,315,298,319]" name="India" pageId="16" pageNumber="17">India</collectingCountry>
, N
<collectingCountry id="22ACD7A22A51D857FEF278251EB0B71A" box="[347,435,298,319]" name="Thailand" pageId="16" pageNumber="17">Thailand</collectingCountry>
,
<collectingCountry id="22ACD7A22A51D857FE1578251D18B71A" box="[444,539,298,319]" name="Myanmar" pageId="16" pageNumber="17">Myanmar</collectingCountry>
, SW
<collectingCountry id="22ACD7A22A51D857FDE678251D8BB71A" box="[591,648,298,319]" name="China" pageId="16" pageNumber="17">China</collectingCountry>
(
<collectingRegion id="987F59D02A51D857FD3D78251DE2B71A" box="[660,737,298,319]" country="China" name="Yunnan" pageId="16" pageNumber="17">Yunnan</collectingRegion>
)......................................... [4]
<taxonomicName id="9DBBECB12A51D857FB5778251A98B71A" ID-CoL="8L4NH" authority="Bates" authorityName="Bates" box="[1278,1435,298,319]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="excelsa">
<emphasis id="68CF4B202A51D857FB5778251A62B71A" box="[1278,1377,298,319]" italics="true" pageId="16" pageNumber="17">C. excelsa</emphasis>
Bates
</taxonomicName>
</paragraph>
<paragraph id="5A0497322A51D857FF3E78481A9FB7B2" blockId="16.[151,1437,151,407]" pageId="16" pageNumber="17">
- Both pronotal disc and elytral intervals glabrous. Femora blackish at apical two thirds or more. Head narrow and convex. Pronotum elongate-cordate; disc convex, with narrow lateral furrows.
<collectingCountry id="22ACD7A22A51D857FCC8786A1C91B75F" box="[865,914,357,378]" name="India" pageId="16" pageNumber="17">India</collectingCountry>
(W and N
<collectingCountry id="22ACD7A22A51D857FC55786A1B32B75F" box="[1020,1073,357,378]" name="India" pageId="16" pageNumber="17">India</collectingCountry>
),
<collectingCountry id="22ACD7A22A51D857FBEB786A1BB7B75F" box="[1090,1204,357,378]" name="Bangladesh" pageId="16" pageNumber="17">Bangladesh</collectingCountry>
.................................................................................................... [3]
<taxonomicName id="9DBBECB12A51D857FB7B788D1A9FB7B2" ID-CoL="5X83Q" authority="CHaUdOir" authorityName="Chaudoir Lectotypus" authorityYear="1872" box="[1234,1436,386,407]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="22" phylum="Arthropoda" rank="species" species="lativittis">
<emphasis id="68CF4B202A51D857FB7B788D1A3EB7B2" box="[1234,1341,386,407]" italics="true" pageId="16" pageNumber="17">C. lativittis</emphasis>
Chaudoir
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>

View file

@ -0,0 +1,384 @@
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<mods:title id="33BD7AE6F9E55FE63AA9ED3060B262C3">Revision of the Oriental species of Calleida Latreille (sensu lato). Part 1: Introduction, groups of species, and species of six species groups (Coleoptera: Carabidae: Lebiini)</mods:title>
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<mods:namePart id="AACF2DB553099B98C05EF3679C75C512">Casale, Achille</mods:namePart>
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<mods:namePart id="F380B0E44961776A00F3864C54203B16">Shi, Hongliang</mods:namePart>
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<mods:title id="F8CDFBE36FD5763C9E42D86FA343F24A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3CC3AECDBF67D76AC998F8CED6423DB8">
<mods:date id="B955711F401616CC7EA013EBA205403E">2018</mods:date>
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<mods:number id="647B834EF0A3B0A150422D32780FBB90">2018-06-29</mods:number>
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<subSection id="33348ED92A50D854FF3E7FC11C01B1F8" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18" type="multiple">
<treatment id="D21226242A50D854FF3E7FC11C01B1F8" ID-DOI="http://doi.org/10.5281/zenodo.5950859" ID-GBIF-Taxon="146043506" ID-Zenodo-Dep="5950859" LSID="urn:lsid:plazi:treatment:D21226242A50D854FF3E7FC11C01B1F8" httpUri="http://treatment.plazi.org/id/D21226242A50D854FF3E7FC11C01B1F8" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
<subSubSection id="12A1C4B92A50D856FF3E7FC11C0AB0CD" box="[151,777,1741,1768]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="5A0497322A50D856FF3E7FC11C0AB0CD" blockId="17.[151,777,1741,1768]" box="[151,777,1741,1768]" pageId="17" pageNumber="18">
<heading id="014C205E2A50D856FF3E7FC11C0AB0CD" bold="true" box="[151,777,1741,1768]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<emphasis id="68CF4B202A50D856FF3E7FC11C0AB0CD" bold="true" box="[151,777,1741,1768]" pageId="17" pageNumber="18">
[4]
<taxonomicName id="9DBBECB12A50D856FF697FC21D18B0CD" ID-CoL="8L4NH" authority="Bates, 1892" authorityName="Bates" authorityYear="1892" box="[192,539,1741,1768]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="excelsa">
<emphasis id="68CF4B202A50D856FF697FC21E86B0C2" bold="true" box="[192,389,1741,1767]" italics="true" pageId="17" pageNumber="18">Calleida excelsa</emphasis>
<bibRefCitation id="3E2AEAC32A50D856FE257FC11D18B0CD" box="[396,539,1742,1768]" pageId="17" pageNumber="18" refId="ref29370" refString="Bates, H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of the Carabidae. Annali del Museo Civico di Storia naturale di Genova, 32, 267 - 428." type="journal article">Bates, 1892</bibRefCitation>
</taxonomicName>
, status resurrected
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A50D856FF3E7E1B1B59B16C" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="5A0497322A50D856FF3E7E1B1B59B16C" blockId="17.[151,1436,1812,1865]" pageId="17" pageNumber="18">
<taxonomicName id="9DBBECB12A50D856FF3E7E1B1EF1B10F" ID-CoL="8L4NH" authority="Bates, 1892: 422" authorityName="Bates" authorityPageNumber="422" authorityYear="1892" box="[151,498,1812,1834]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="excelsa">
<emphasis id="68CF4B202A50D856FF3E7E1B1E43B10F" box="[151,320,1812,1834]" italics="true" pageId="17" pageNumber="18">Calleida excelsa</emphasis>
<bibRefCitation id="3E2AEAC32A50D856FEEF7E1A1EC2B10F" box="[326,449,1812,1834]" pageId="17" pageNumber="18" refId="ref29370" refString="Bates, H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of the Carabidae. Annali del Museo Civico di Storia naturale di Genova, 32, 267 - 428." type="journal article">Bates, 1892</bibRefCitation>
: 422
</taxonomicName>
(
<typeStatus id="850029902A50D856FDA97E1B1D2EB10F" box="[512,557,1812,1834]" pageId="17" pageNumber="18">type</typeStatus>
locality: Carin Chebà [
<collectingRegion id="987F59D02A50D856FCB37E1B1C59B10F" box="[794,858,1812,1834]" country="Myanmar" name="Kayin" pageId="17" pageNumber="18">Kayin</collectingRegion>
State];
<typeStatus id="850029902A50D856FC007E1B1B05B10F" box="[937,1030,1812,1834]" pageId="17" pageNumber="18" type="lectotype">lectotype</typeStatus>
deposited in MSNG);
<bibRefCitation id="3E2AEAC32A50D856FB477E1B1A9BB10F" box="[1262,1432,1812,1834]" pageId="17" pageNumber="18" refId="ref28815" refString="Andrewes, H. E. (1921) Notes on Synonymy and on some Types of Oriental Carabidae in various foreign collections. Transactions of the Royal entomological Society, London, 69, 145 - 190. https: // doi. org / 10.1111 / j. 1365 - 2311.1921. tb 02805. x" type="journal article">Andrewes, 1921</bibRefCitation>
: 150. (as synonym under
<taxonomicName id="9DBBECB12A50D856FE697E3B1D32B16C" ID-CoL="5X83Q" authorityName="Chaudoir Lectotypus" authorityYear="1872" box="[448,561,1843,1865]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lativittis">
<emphasis id="68CF4B202A50D856FE697E3B1D32B16C" box="[448,561,1843,1865]" italics="true" pageId="17" pageNumber="18">C. lativittis</emphasis>
</taxonomicName>
);
<bibRefCitation id="3E2AEAC32A50D856FDEE7E3C1DDFB16C" box="[583,732,1843,1865]" pageId="17" pageNumber="18" refId="ref30044" refString="Jedlicka, A. (1963) Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae-Brachyninae (Coleoptera, Carabidae). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden, 28, 269 - 579." type="journal article">Jedlička, 1963</bibRefCitation>
: 435 (as synonym under
<taxonomicName id="9DBBECB12A50D856FC707E3B1B49B16C" ID-CoL="5X83Q" authorityName="Chaudoir Lectotypus" authorityYear="1872" box="[985,1098,1843,1865]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lativittis">
<emphasis id="68CF4B202A50D856FC707E3B1B49B16C" box="[985,1098,1843,1865]" italics="true" pageId="17" pageNumber="18">C. lativittis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A50D854FF3E7E7A1C23B739" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="5A0497322A50D856FF3E7E7A1A9FB1F2" blockId="17.[151,1436,1909,2007]" pageId="17" pageNumber="18">
<emphasis id="68CF4B202A50D856FF3E7E7A1EC3B1AB" bold="true" box="[151,448,1909,1934]" pageId="17" pageNumber="18">
<typeStatus id="850029902A50D856FF3E7E7A1FD1B1AB" box="[151,210,1909,1934]" pageId="17" pageNumber="18">Type</typeStatus>
material examined:
</emphasis>
<materialsCitation id="EAD39D6F2A50D856FE6E7E781DF6B1F2" ID-GBIF-Occurrence="1852047372" collectingDate="1988-12" collectorName="L. Fea" country="Myanmar" elevation="1000" location="Carin Chebà" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
<typeStatus id="850029902A50D856FE6E7E781D39B1AA" box="[455,570,1911,1935]" pageId="17" pageNumber="18" type="lectotype">Lectotype</typeStatus>
, designated herein,
<specimenCount id="4CBD5CBB2A50D856FCBE7E7A1C65B1AB" box="[791,870,1909,1934]" pageId="17" pageNumber="18" type="female">female</specimenCount>
, “
<location id="5F64C1E92A50D856FCD27E7A1B0AB1AB" LSID="urn:lsid:plazi:treatment:D21226242A50D854FF3E7FC11C01B1F8:5F64C1E92A50D856FCD27E7A1B0AB1AB" box="[891,1033,1909,1934]" country="Myanmar" name="Carin Chebà" pageId="17" pageNumber="18">Carin Chebà</location>
[
<collectingCountry id="22ACD7A22A50D856FBBE7E781B6AB1AB" box="[1047,1129,1911,1934]" name="Myanmar" pageId="17" pageNumber="18">Burma</collectingCountry>
]
<elevation id="D19670012A50D856FBDF7E791BFEB1AB" box="[1142,1277,1910,1934]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="17" pageNumber="18" unit="m" value="1000.0" valueMax="1100.0" valueMin="900.0">
<quantity id="9D433AD72A50D856FBDF7E791BFEB1AB" box="[1142,1277,1910,1934]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="17" pageNumber="18" unit="m" value="1000.0" valueMax="1100.0" valueMin="900.0">900-1100 m</quantity>
</elevation>
<collectorName id="F74EF2E42A50D856FAAA7E781A48B1AB" box="[1283,1355,1911,1934]" pageId="17" pageNumber="18">L. Fea</collectorName>
V
<date id="2E05B1F22A50D856FAC57E781FB5B197" pageId="17" pageNumber="18" value="1988-12">
<collectingDate id="3E41481A2A50D856FAC57E781FB5B197" pageId="17" pageNumber="18" value="1988-12">XII- 88</collectingDate>
</date>
”, “Typus”, “
<taxonomicName id="9DBBECB12A50D856FEED7E951D4BB197" authority="Bates" authorityName="Bates" box="[324,584,1946,1971]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="excelsa">Calleida excelsa Bates</taxonomicName>
” (handwritten by Bates), “
<taxonomicName id="9DBBECB12A50D856FCDF7E951B16B197" authority="Bates" authorityName="Bates" box="[886,1045,1946,1971]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="excelsa">excelsa Bates</taxonomicName>
”, “
<taxonomicName id="9DBBECB12A50D856FB9F7E951A35B197" authority="Bates" authorityName="Bates" box="[1078,1334,1946,1971]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="excelsa">Calleida excelsa Bates</taxonomicName>
(es. tip.) Bates”, “
<location id="5F64C1E92A50D856FF557EB01D22B1F2" LSID="urn:lsid:plazi:treatment:D21226242A50D854FF3E7FC11C01B1F8:5F64C1E92A50D856FF557EB01D22B1F2" box="[252,545,1982,2007]" country="Myanmar" name="Museo Civico di Genova" pageId="17" pageNumber="18">Museo Civico di Genova</location>
” (MSNG,
<figureCitation id="C2808BB72A50D856FD357EB11DEFB1F2" box="[668,748,1982,2007]" captionStart="FIGURES 3641" captionStartId="18.[151,264,1697,1719]" captionTargetBox="[184,1416,280,1673]" captionTargetId="figure@18.[151,1436,241,1676]" captionTargetPageId="18" captionText="FIGURES 3641. Calleida excelsa Bates. 36 habitus and labels of lectotype (female, Carin Chebà, Myanmar, MSNG), scale bar = 2.0 mm; 37 habitus of paralectotype (female, Senmigion, Myanmar, MNHN), scale bar = 2.0 mm; 38 habitus (female, Yunnan, China), scale bar = 2.0 mm; 39 female reproductive tract (Chiang Mai, Thailand), scale bar = 0.5 mm; 40 ovipositor (Yunnan, China), scale bar = 0.2 mm; 41 apex of immature aedeagus, left-lateral view, scale bar = 0.2 mm." httpUri="https://zenodo.org/record/1302727/files/figure.png" pageId="17" pageNumber="18">fig. 36</figureCitation>
)
</materialsCitation>
.
<typeStatus id="850029902A50D856FCA87EB11C9FB1F2" box="[769,924,1982,2007]" pageId="17" pageNumber="18" type="paralectotype">Paralectotype</typeStatus>
, one female, &quot;
<collectingCountry id="22ACD7A22A50D856FBEB7EB11BB3B1F2" box="[1090,1200,1982,2007]" name="Myanmar" pageId="17" pageNumber="18">Birmania</collectingCountry>
, Senmigion, Fea. 28
</paragraph>
<paragraph id="5A0497322A53D855FF3E79981CA4B6F1" blockId="18.[151,1436,151,213]" pageId="18" pageNumber="19">
Feb. 86&quot;, &quot;
<taxonomicName id="9DBBECB12A53D855FEA779981D13B695" authority="Bates" authorityName="Bates" box="[270,528,151,176]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="excelsa">Calleida excelsa Bates</taxonomicName>
&quot; (handwritten by Bates); &quot;Ex.Musaeo,
<bibRefCitation id="3E2AEAC32A53D855FC6279971B97B695" box="[971,1172,152,176]" pageId="18" pageNumber="19" refId="ref29370" refString="Bates, H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of the Carabidae. Annali del Museo Civico di Storia naturale di Genova, 32, 267 - 428." type="journal article">H.W. Bates, 1892</bibRefCitation>
&quot; (
<collectionCode id="3CAA0FF72A53D855FB0579971A05B695" box="[1196,1286,152,176]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="18" pageNumber="19">MNHN</collectionCode>
,
<figureCitation id="C2808BB72A53D855FABB79981A5DB695" box="[1298,1374,151,176]" captionStart="FIGURES 3641" captionStartId="18.[151,264,1697,1719]" captionTargetBox="[184,1416,280,1673]" captionTargetId="figure@18.[151,1436,241,1676]" captionTargetPageId="18" captionText="FIGURES 3641. Calleida excelsa Bates. 36 habitus and labels of lectotype (female, Carin Chebà, Myanmar, MSNG), scale bar = 2.0 mm; 37 habitus of paralectotype (female, Senmigion, Myanmar, MNHN), scale bar = 2.0 mm; 38 habitus (female, Yunnan, China), scale bar = 2.0 mm; 39 female reproductive tract (Chiang Mai, Thailand), scale bar = 0.5 mm; 40 ovipositor (Yunnan, China), scale bar = 0.2 mm; 41 apex of immature aedeagus, left-lateral view, scale bar = 0.2 mm." httpUri="https://zenodo.org/record/1302727/files/figure.png" pageId="18" pageNumber="19">fig. 37</figureCitation>
); one female, &quot;
<collectingRegion id="987F59D02A53D855FF5379B31E3BB6F1" box="[250,312,188,212]" country="Philippines" name="Naga" pageId="18" pageNumber="19">Naga</collectingRegion>
hills 3000&quot;, &quot;Ex.Musaeo,
<bibRefCitation id="3E2AEAC32A53D855FDF679B21C2BB6F1" box="[607,808,188,212]" pageId="18" pageNumber="19" refId="ref29370" refString="Bates, H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of the Carabidae. Annali del Museo Civico di Storia naturale di Genova, 32, 267 - 428." type="journal article">H.W. Bates, 1892</bibRefCitation>
&quot; (
<collectionCode id="3CAA0FF72A53D855FC9679B31C98B6F1" box="[831,923,188,212]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="18" pageNumber="19">MNHN</collectionCode>
).
</paragraph>
<caption id="0EC4C7BA2A53D855FF3E7FAE1BD3B136" httpUri="https://zenodo.org/record/1302727/files/figure.png" pageId="18" pageNumber="19" startId="18.[151,264,1697,1719]" targetBox="[184,1416,280,1673]" targetPageId="18">
<paragraph id="5A0497322A53D855FF3E7FAE1BD3B136" blockId="18.[151,1436,1697,1811]" pageId="18" pageNumber="19">
<emphasis id="68CF4B202A53D855FF3E7FAE1E56B092" bold="true" box="[151,341,1697,1719]" pageId="18" pageNumber="19">FIGURES 3641.</emphasis>
<taxonomicName id="9DBBECB12A53D855FEF57FAE1D4EB092" authority="Bates." authorityName="Bates." box="[348,589,1697,1719]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="excelsa">
<emphasis id="68CF4B202A53D855FEF57FAE1D05B092" bold="true" box="[348,518,1697,1719]" italics="true" pageId="18" pageNumber="19">Calleida excelsa</emphasis>
Bates.
</taxonomicName>
<emphasis id="68CF4B202A53D855FDFA7FAE1D6EB092" bold="true" box="[595,621,1697,1719]" pageId="18" pageNumber="19">36</emphasis>
habitus and labels of lectotype (female, Carin Chebà, Myanmar, MSNG), scale bar = 2.0 mm;
<emphasis id="68CF4B202A53D855FE9A7FCF1E4FB0F3" bold="true" box="[307,332,1728,1750]" pageId="18" pageNumber="19">37</emphasis>
habitus of paralectotype (female, Senmigion, Myanmar, MNHN), scale bar = 2.0 mm;
<emphasis id="68CF4B202A53D855FB7F7FCF1BF3B0F3" bold="true" box="[1238,1264,1728,1750]" pageId="18" pageNumber="19">38</emphasis>
habitus (female, Yunnan, China), scale bar = 2.0 mm;
<emphasis id="68CF4B202A53D855FDB17FD11D31B0D1" bold="true" box="[536,562,1758,1780]" pageId="18" pageNumber="19">39</emphasis>
female reproductive tract (Chiang Mai, Thailand), scale bar = 0.5 mm;
<emphasis id="68CF4B202A53D855FABD7FD11A2DB0D1" bold="true" box="[1300,1326,1758,1780]" pageId="18" pageNumber="19">40</emphasis>
ovipositor (Yunnan, China), scale bar = 0.2 mm;
<emphasis id="68CF4B202A53D855FDB37FF21D37B136" bold="true" box="[538,564,1789,1811]" pageId="18" pageNumber="19">41</emphasis>
apex of immature aedeagus, left-lateral view, scale bar = 0.2 mm.
</paragraph>
</caption>
<paragraph id="5A0497322A53D854FF6E7E4F1C23B739" blockId="18.[151,1436,1856,2025]" lastBlockId="19.[151,1437,151,2013]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<emphasis id="68CF4B202A53D855FF6E7E4F1D18B17C" bold="true" box="[199,539,1856,1881]" pageId="18" pageNumber="19">Non-type material examined</emphasis>
:
<emphasis id="68CF4B202A53D855FD837E4F1D69B17C" bold="true" box="[554,618,1856,1881]" pageId="18" pageNumber="19">
<collectingCountry id="22ACD7A22A53D855FD837E4F1D69B17C" box="[554,618,1856,1881]" name="India" pageId="18" pageNumber="19">India</collectingCountry>
</emphasis>
: one female “
<collectingCountry id="22ACD7A22A53D855FCAF7E4F1C46B17C" box="[774,837,1856,1881]" name="India" pageId="18" pageNumber="19">India</collectingCountry>
”, “Bowring 63.47” “
<taxonomicName id="9DBBECB12A53D855FB847E4F1A2EB17C" authority="Chd." authorityName="Chaudoir Lectotypus" authorityYear="1872" box="[1069,1325,1856,1881]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lativittis">Calleida lativittis Chd.</taxonomicName>
=
<taxonomicName id="9DBBECB12A53D855FAE37E4F1FD6B159" authority="Bates" authorityName="Bates" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="excelsa">excelsa Bates</taxonomicName>
compared with
<typeStatus id="850029902A53D855FE0A7E6A1EDDB158" box="[419,478,1893,1917]" pageId="18" pageNumber="19">types</typeStatus>
H.E.A.[Andrewes]” (
<collectionCode id="3CAA0FF72A53D855FD4D7E6A1C3EB158" box="[740,829,1893,1917]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="18" pageNumber="19">NHML</collectionCode>
);
<emphasis id="68CF4B202A53D855FCF07E6B1CD2B158" bold="true" box="[857,977,1892,1917]" pageId="18" pageNumber="19">
<collectingCountry id="22ACD7A22A53D855FCF07E6B1CD2B158" box="[857,977,1892,1917]" name="Myanmar" pageId="18" pageNumber="19">Myanmar</collectingCountry>
</emphasis>
: one female “
<collectingRegion id="987F59D02A53D855FB2C7E6A1BEEB158" box="[1157,1261,1893,1917]" country="Myanmar" name="Yangon" pageId="18" pageNumber="19">Rangoon</collectingRegion>
E.M. Janson” (
<collectionCode id="3CAA0FF72A53D855FF367E871FFBB185" box="[159,248,1928,1952]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="18" pageNumber="19">NHML</collectionCode>
); one female “
<collectingCountry id="22ACD7A22A53D855FE367E871D0FB184" box="[415,524,1928,1953]" name="Myanmar" pageId="18" pageNumber="19">Birmania</collectingCountry>
” (
<collectionCode id="3CAA0FF72A53D855FD8E7E871D64B185" box="[551,615,1928,1952]" country="Germany" httpUri="http://grbio.org/cool/mqei-3i36" name="Culture Collection of Algae at the University of Marburg" pageId="18" pageNumber="19">CCA</collectionCode>
);
<emphasis id="68CF4B202A53D855FDD57E871DEAB184" bold="true" box="[636,745,1928,1953]" pageId="18" pageNumber="19">
<collectingCountry id="22ACD7A22A53D855FDD57E871DEAB184" box="[636,745,1928,1953]" name="Thailand" pageId="18" pageNumber="19">Thailand</collectingCountry>
</emphasis>
: one female, “
<collectingCountry id="22ACD7A22A53D855FC3A7E871CFAB184" box="[915,1017,1928,1953]" name="Thailand" pageId="18" pageNumber="19">Thailand</collectingCountry>
bor. Chiang Dao env. 21.5-
<date id="2E05B1F22A53D855FA917E871A9FB184" box="[1336,1436,1928,1953]" pageId="18" pageNumber="19" value="1995-06-04">4.6.1995</date>
lgt. M. Snizek” (
<collectionCode id="3CAA0FF72A53D855FEF57EA21E98B1E0" box="[348,411,1965,1989]" country="Germany" httpUri="http://grbio.org/cool/mqei-3i36" name="Culture Collection of Algae at the University of Marburg" pageId="18" pageNumber="19">CCA</collectionCode>
); one female, “
<date id="2E05B1F22A53D855FDE67EA21DADB1E0" box="[591,686,1965,1989]" pageId="18" pageNumber="19" value="1998-01-02">2.I.1998</date>
Thaild.
<collectingRegion id="987F59D02A53D855FCBA7EA31CA3B1E0" box="[787,928,1964,1989]" country="Thailand" name="Chiang Mai" pageId="18" pageNumber="19">Chiang Mai</collectingRegion>
. Samoeng,
<geoCoordinate id="3F8FF1F52A53D855FB817EA21BDBB1E0" box="[1064,1240,1964,1989]" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="1" value="18.862556">N 018°51'45.2''</geoCoordinate>
<geoCoordinate id="3F8FF1F52A53D855FB4B7EA21A92B1E0" box="[1250,1425,1965,1989]" direction="east" orientation="longitude" pageId="18" pageNumber="19" precision="1" value="98.72186">E 098°43'18.7''</geoCoordinate>
” (
<collectionCode id="3CAA0FF72A53D855FF367EDF1FFAB1CD" box="[159,249,2000,2024]" country="Switzerland" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="18" pageNumber="19">NHMB</collectionCode>
); one female, same data (
<collectionCode id="3CAA0FF72A53D855FDBC7EDF1D57B1CD" box="[533,596,2000,2024]" country="Germany" httpUri="http://grbio.org/cool/mqei-3i36" name="Culture Collection of Algae at the University of Marburg" pageId="18" pageNumber="19">CCA</collectionCode>
);
<emphasis id="68CF4B202A53D855FDCE7EDF1DB3B1CC" bold="true" box="[615,688,2000,2025]" pageId="18" pageNumber="19">
<collectingCountry id="22ACD7A22A53D855FDCE7EDF1DB3B1CC" box="[615,688,2000,2025]" name="China" pageId="18" pageNumber="19">China</collectingCountry>
</emphasis>
: two teneral males, &quot;
<collectingRegion id="987F59D02A53D855FC327EDF1CF5B1CD" box="[923,1014,2000,2024]" country="China" name="Yunnan" pageId="18" pageNumber="19">Yunnan</collectingRegion>
, Yingjiang county, Nabang township,
<geoCoordinate id="3F8FF1F52A52D854FF3E79981E14B695" box="[151,279,151,176]" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="1" value="24.75323">N24.75323</geoCoordinate>
,
<geoCoordinate id="3F8FF1F52A52D854FE8A79981EA3B695" box="[291,416,151,176]" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="1" value="97.56032">E97.56032</geoCoordinate>
,
<quantity id="9D433AD72A52D854FE0579981EECB695" box="[428,495,151,176]" metricMagnitude="2" metricUnit="m" metricValue="2.52" pageId="19" pageNumber="20" unit="m" value="252.0">252m</quantity>
, 2008.
<date id="2E05B1F22A52D854FD9279971D6CB695" box="[571,623,152,176]" pageId="19" pageNumber="20" value="1931-05">V.31</date>
, light trap, Shi Hongliang leg.&quot; (
<collectionCode id="3CAA0FF72A52D854FC4879971B26B695" box="[993,1061,152,176]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="19" pageNumber="20">IZAS</collectionCode>
); one female &quot;
<collectingRegion id="987F59D02A52D854FB6E79971A21B695" box="[1223,1314,152,176]" country="China" name="Yunnan" pageId="19" pageNumber="20">Yunnan</collectingRegion>
, Yingjiang county, Tongbiguan, Laoxiangping,
<quantity id="9D433AD72A52D854FD9079B31D7FB6F0" box="[569,636,188,213]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="19" pageNumber="20" unit="m" value="450.0">450m</quantity>
, 2003.
<date id="2E05B1F22A52D854FD6579B31C07B6F1" box="[716,772,188,212]" pageId="19" pageNumber="20" value="1929-10">X.29</date>
, Qin Ruihao leg.&quot; (
<collectionCode id="3CAA0FF72A52D854FC4579B31B43B6F1" box="[1004,1088,188,212]" pageId="19" pageNumber="20">SWFU</collectionCode>
); one female, &quot;
<collectingRegion id="987F59D02A52D854FB5B79B31A4EB6F1" box="[1266,1357,188,212]" country="China" name="Yunnan" pageId="19" pageNumber="20">Yunnan</collectingRegion>
, Ruili, Huyu,
<quantity id="9D433AD72A52D854FF4A79EF1E25B6DD" box="[227,294,224,248]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="19" pageNumber="20" unit="m" value="900.0">900m</quantity>
, 2003.
<date id="2E05B1F22A52D854FEDA79D01EA6B6DD" box="[371,421,223,248]" pageId="19" pageNumber="20">XI.5</date>
, Qin Ruihao leg.&quot; (
<collectionCode id="3CAA0FF72A52D854FDD779EF1DD1B6DD" box="[638,722,224,248]" pageId="19" pageNumber="20">SWFU</collectionCode>
); one female &quot;
<collectingRegion id="987F59D02A52D854FCD979EF1CC8B6DD" box="[880,971,224,248]" country="China" name="Yunnan" pageId="19" pageNumber="20">Yunnan</collectingRegion>
, Menglun town, Xishuangbanna Tropical Botany Garden, 2007.
<date id="2E05B1F22A52D854FE27780B1ECEB739" box="[398,461,260,284]" pageId="19" pageNumber="20" value="1920-04">IV.20</date>
, Cao Liangming leg.&quot; (
<collectionCode id="3CAA0FF72A52D854FD7A780B1C17B739" box="[723,788,260,284]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="19" pageNumber="20">CAU</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A52D854FF6E78281BD2B7AD" pageId="19" pageNumber="20" type="discussion">
<paragraph id="5A0497322A52D854FF6E78281BD2B7AD" blockId="19.[151,1437,151,2013]" pageId="19" pageNumber="20">
<emphasis id="68CF4B202A52D854FF6E78281E8FB765" bold="true" box="[199,396,295,320]" pageId="19" pageNumber="20">Taxonomic note.</emphasis>
<bibRefCitation id="3E2AEAC32A52D854FE3D78281D52B765" box="[404,593,295,320]" pageId="19" pageNumber="20" refId="ref28815" refString="Andrewes, H. E. (1921) Notes on Synonymy and on some Types of Oriental Carabidae in various foreign collections. Transactions of the Royal entomological Society, London, 69, 145 - 190. https: // doi. org / 10.1111 / j. 1365 - 2311.1921. tb 02805. x" type="journal article">Andrewes (1921</bibRefCitation>
,
<bibRefCitation id="3E2AEAC32A52D854FDF478271D95B765" box="[605,662,296,320]" pageId="19" pageNumber="20" refId="ref28962" refString="Andrewes, H. E. (1930) Catalogue of Indian Insects. Part 18. Carabidae. Government of India, Central Publication Branch, Calcutta, 389 pp." type="book">1930</bibRefCitation>
) wrongly proposed this taxon as junior synonym of
<taxonomicName id="9DBBECB12A52D854FB4E78271A66B765" authorityName="Chaudoir Lectotypus" authorityYear="1872" box="[1255,1381,295,320]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="lativittis">
<emphasis id="68CF4B202A52D854FB4E78271A66B765" box="[1255,1381,295,320]" italics="true" pageId="19" pageNumber="20">C. lativittis</emphasis>
</taxonomicName>
, and this synonymy was accepted by
<bibRefCitation id="3E2AEAC32A52D854FDA878431DADB740" box="[513,686,332,357]" pageId="19" pageNumber="20" refId="ref30044" refString="Jedlicka, A. (1963) Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae-Brachyninae (Coleoptera, Carabidae). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden, 28, 269 - 579." type="journal article">Jedlička (1963)</bibRefCitation>
and cited by
<bibRefCitation id="3E2AEAC32A52D854FCEC78421CE9B740" box="[837,1002,332,357]" pageId="19" pageNumber="20" refId="ref30481" refString="Lorenz, W. (2005) Systematic list of extant ground beetles of the world (Insecta Coleoptera Geadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 2 nd Edition. Wolfgang Lorenz, Tutzing, 530 pp." type="book">Lorenz (2005)</bibRefCitation>
. Based on the examination of
<typeStatus id="850029902A52D854FA9578421A6FB740" box="[1340,1388,333,357]" pageId="19" pageNumber="20">type</typeStatus>
and further material of both species, we demonstrated that
<taxonomicName id="9DBBECB12A52D854FD50787F1C6FB7AD" box="[761,876,367,392]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="excelsa">
<emphasis id="68CF4B202A52D854FD50787F1C6FB7AD" box="[761,876,367,392]" italics="true" pageId="19" pageNumber="20">C. excelsa</emphasis>
</taxonomicName>
is a valid, very distinct species.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A52D854FF6E789B1EDAB4E8" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="5A0497322A52D854FF6E789B1C1DB445" blockId="19.[151,1437,151,2013]" pageId="19" pageNumber="20">
<emphasis id="68CF4B202A52D854FF6E789B1E42B788" bold="true" box="[199,321,404,429]" pageId="19" pageNumber="20">Diagnosis.</emphasis>
With the character states of
<taxonomicName id="9DBBECB12A52D854FD21789B1DEAB788" authorityName="Latreille" authorityYear="1824" box="[648,745,404,429]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A52D854FD21789B1DEAB788" box="[648,745,404,429]" italics="true" pageId="19" pageNumber="20">Calleida</emphasis>
</taxonomicName>
of the
<taxonomicName id="9DBBECB12A52D854FC95789B1C9BB788" authorityName="Chaudoir Lectotypus" authorityYear="1872" box="[828,920,404,429]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="lativittis">
<emphasis id="68CF4B202A52D854FC95789B1C9BB788" box="[828,920,404,429]" italics="true" pageId="19" pageNumber="20">lativittis</emphasis>
</taxonomicName>
species group, but from other known species distinguishable by the following character combination: dorsal side with dense pubescence all through head, pronotum and elytra. Elytra metallic green, disc with reddish patch on the inner three to five intervals; all antennomeres yellowish, or 311 somewhat darkened; femora yellowish, blackish only at extreme apex or apical third; tibiae and tarsomeres yellowish. Head wide and depressed. Pronotum transverse-cordate, widened in the middle, with lateral furrows wide and markedly flattened.
</paragraph>
<paragraph id="5A0497322A52D854FF6E7B631EDAB4E8" blockId="19.[151,1437,151,2013]" pageId="19" pageNumber="20">
Closely related and similar to
<taxonomicName id="9DBBECB12A52D854FD867B621C25B4A0" authority="Chaudoir" authorityName="Chaudoir Lectotypus" authorityYear="1872" box="[559,806,620,645]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="lativittis">
<emphasis id="68CF4B202A52D854FD867B621DB3B4A0" box="[559,688,620,645]" italics="true" pageId="19" pageNumber="20">C. lativittis</emphasis>
Chaudoir
</taxonomicName>
, but easily recognized by the head, pronotum, elytral intervals and underside distinctly pubescent, and by the elytral disc with reddish patch usually wider and blackish area of femurs much shorter.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A52D854FF6E7BD81BCEB155" pageId="19" pageNumber="20" type="description">
<paragraph id="5A0497322A52D854FF6E7BD81E84B530" blockId="19.[151,1437,151,2013]" pageId="19" pageNumber="20">
<emphasis id="68CF4B202A52D854FF6E7BD81E5AB4D5" bold="true" box="[199,345,727,752]" pageId="19" pageNumber="20">Description.</emphasis>
The original description provided by
<bibRefCitation id="3E2AEAC32A52D854FC8A7BD71CBAB4D5" box="[803,953,728,752]" pageId="19" pageNumber="20" refId="ref29370" refString="Bates, H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of the Carabidae. Annali del Museo Civico di Storia naturale di Genova, 32, 267 - 428." type="journal article">Bates (1892)</bibRefCitation>
in Latin is here completed with several additional characters.
</paragraph>
<paragraph id="5A0497322A52D854FF6E7A101D6FB5A5" blockId="19.[151,1437,151,2013]" pageId="19" pageNumber="20">
<emphasis id="68CF4B202A52D854FF6E7A101E86B51D" box="[199,389,799,824]" italics="true" pageId="19" pageNumber="20">General features</emphasis>
as in
<figureCitation id="C2808BB72A52D854FE677A101D6CB51D" box="[462,623,799,824]" captionStart="FIGURES 3641" captionStartId="18.[151,264,1697,1719]" captionTargetBox="[184,1416,280,1673]" captionTargetId="figure@18.[151,1436,241,1676]" captionTargetPageId="18" captionText="FIGURES 3641. Calleida excelsa Bates. 36 habitus and labels of lectotype (female, Carin Chebà, Myanmar, MSNG), scale bar = 2.0 mm; 37 habitus of paralectotype (female, Senmigion, Myanmar, MNHN), scale bar = 2.0 mm; 38 habitus (female, Yunnan, China), scale bar = 2.0 mm; 39 female reproductive tract (Chiang Mai, Thailand), scale bar = 0.5 mm; 40 ovipositor (Yunnan, China), scale bar = 0.2 mm; 41 apex of immature aedeagus, left-lateral view, scale bar = 0.2 mm." httpUri="https://zenodo.org/record/1302727/files/figure.png" pageId="19" pageNumber="20">figures 3638</figureCitation>
: Medium sized: L: 9.0
<quantity id="9D433AD72A52D854FC2A7A2F1CE8B512" box="[899,1003,800,824]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="19" pageNumber="20" unit="mm" value="10.8">10.8 mm</quantity>
; TL:
<quantity id="9D433AD72A52D854FB857A2F1BBBB512" box="[1068,1208,800,824]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" metricValueMax="9.8" metricValueMin="8.4" pageId="19" pageNumber="20" unit="mm" value="9.1" valueMax="9.8" valueMin="8.4">8.49.8 mm</quantity>
. Body elongate and slender, depressed; elytra slightly widened at the posterior fifth. Head, pronotum, elytral intervals, meso- and metasternum densely punctate-pubescent.
</paragraph>
<paragraph id="5A0497322A52D854FF6E7A831B63B5C8" blockId="19.[151,1437,151,2013]" pageId="19" pageNumber="20">
<emphasis id="68CF4B202A52D854FF6E7A831E15B580" box="[199,278,908,933]" italics="true" pageId="19" pageNumber="20">Colour</emphasis>
: head, pronotum, lateral margins of elytra, epipleura, underside, antennae and legs rufous; femora blackish at apex, or blackish at apical third in some individuals; elytra metallic green or bluish green; disc with a wide reddish patch, extended from the base to apex on the inner three to five intervals.
</paragraph>
<paragraph id="5A0497322A52D854FF6E7AF61D35B27D" blockId="19.[151,1437,151,2013]" pageId="19" pageNumber="20">
<emphasis id="68CF4B202A52D854FF6E7AF61EF9B235" box="[199,506,1015,1040]" italics="true" pageId="19" pageNumber="20">Lustre and microsculpture</emphasis>
: shiny; head with vanished microsculpture; pronotum mostly without distinct microsculpture, except some fine transverse meshes on disc and on middle part of the basal area; elytra with very shallow isodiametric microsculpture.
</paragraph>
<paragraph id="5A0497322A52D854FF6E7D6B1DECB2E1" blockId="19.[151,1437,151,2013]" pageId="19" pageNumber="20">
<emphasis id="68CF4B202A52D854FF6E7D6B1E07B258" box="[199,260,1124,1149]" italics="true" pageId="19" pageNumber="20">Head</emphasis>
wide and flat, narrower than pronotum, with some oblique frontal wrinkles, smooth except several fine, numerous setiferous punctures; frontal foveae very shallow. Genae rather short, tumid; eyes large and prominent; basal three antennomeres with dense additional setae.
</paragraph>
<paragraph id="5A0497322A52D854FF6E7DDE1A4CB370" blockId="19.[151,1437,151,2013]" pageId="19" pageNumber="20">
<emphasis id="68CF4B202A52D854FF6E7DDE1E35B2CD" box="[199,310,1233,1256]" italics="true" pageId="19" pageNumber="20">Pronotum</emphasis>
transverse-cordate (PW/PL: 1.121.25), widened in the middle, with lateral furrows wide, markedly flattened. Disc with shallow transverse wrinkles and numerous fine setiferous punctures. Lateral margin slightly reflexed, rounded in the anterior half, only sinuate in the basal sixth, then parallel sided to the posterior angles, which are rectangular. Anterior angles effaced, each with six to eight setae. Basal foveae deep and elongate.
</paragraph>
<paragraph id="5A0497322A52D854FF6E7C501A40B3E5" blockId="19.[151,1437,151,2013]" pageId="19" pageNumber="20">
<emphasis id="68CF4B202A52D854FF6E7C501E0EB35D" box="[199,269,1375,1400]" italics="true" pageId="19" pageNumber="20">Elytra</emphasis>
elongate and narrow (EL/EW: 1.762.02), slightly widened at the posterior fifth. Striae very deep, punctate; intervals convex, with evident setiferous punctures; eighth interval not tumid at apex. Umbilicate series of 1517 pores along the eighth stria. Apical truncation straight; outer apical angle not thickened, rounded.
</paragraph>
<paragraph id="5A0497322A52D854FF6E7CC31CB9B008" blockId="19.[151,1437,151,2013]" pageId="19" pageNumber="20">
<emphasis id="68CF4B202A52D854FF6E7CC31E4CB3C0" box="[199,335,1484,1509]" italics="true" pageId="19" pageNumber="20">Ventral side</emphasis>
: lateral areas of meso- and metasternum and abdominal sterna with long, evident pubescence; abdominal sternum VII with two (one on each side) setae in males, four (two on each side) setae in females; apical margin of abdominal sternum VII notched in males, straight in females.
</paragraph>
<paragraph id="5A0497322A52D854FF6E7F381CBAB051" blockId="19.[151,1437,151,2013]" pageId="19" pageNumber="20">
<emphasis id="68CF4B202A52D854FF6E7F381E68B075" box="[199,363,1591,1616]" italics="true" pageId="19" pageNumber="20">Male genitalia</emphasis>
: only two immature males examined, with the following characters: apical orifice opened to the left; apical lamina thickened, as long as wide, rounded at apex (
<figureCitation id="C2808BB72A52D854FCF67F531CAFB051" box="[863,940,1628,1653]" captionStart="FIGURES 3641" captionStartId="18.[151,264,1697,1719]" captionTargetBox="[184,1416,280,1673]" captionTargetId="figure@18.[151,1436,241,1676]" captionTargetPageId="18" captionText="FIGURES 3641. Calleida excelsa Bates. 36 habitus and labels of lectotype (female, Carin Chebà, Myanmar, MSNG), scale bar = 2.0 mm; 37 habitus of paralectotype (female, Senmigion, Myanmar, MNHN), scale bar = 2.0 mm; 38 habitus (female, Yunnan, China), scale bar = 2.0 mm; 39 female reproductive tract (Chiang Mai, Thailand), scale bar = 0.5 mm; 40 ovipositor (Yunnan, China), scale bar = 0.2 mm; 41 apex of immature aedeagus, left-lateral view, scale bar = 0.2 mm." httpUri="https://zenodo.org/record/1302727/files/figure.png" pageId="19" pageNumber="20">fig. 41</figureCitation>
).
</paragraph>
<paragraph id="5A0497322A52D854FF6E7F701BCEB155" blockId="19.[151,1437,151,2013]" pageId="19" pageNumber="20">
<emphasis id="68CF4B202A52D854FF6E7F701E85B0BD" box="[199,390,1663,1688]" italics="true" pageId="19" pageNumber="20">Female genitalia</emphasis>
(reproductive tract
<figureCitation id="C2808BB72A52D854FDCA7F701DAFB0BD" box="[611,684,1663,1688]" captionStart="FIGURES 3641" captionStartId="18.[151,264,1697,1719]" captionTargetBox="[184,1416,280,1673]" captionTargetId="figure@18.[151,1436,241,1676]" captionTargetPageId="18" captionText="FIGURES 3641. Calleida excelsa Bates. 36 habitus and labels of lectotype (female, Carin Chebà, Myanmar, MSNG), scale bar = 2.0 mm; 37 habitus of paralectotype (female, Senmigion, Myanmar, MNHN), scale bar = 2.0 mm; 38 habitus (female, Yunnan, China), scale bar = 2.0 mm; 39 female reproductive tract (Chiang Mai, Thailand), scale bar = 0.5 mm; 40 ovipositor (Yunnan, China), scale bar = 0.2 mm; 41 apex of immature aedeagus, left-lateral view, scale bar = 0.2 mm." httpUri="https://zenodo.org/record/1302727/files/figure.png" pageId="19" pageNumber="20">fig. 39</figureCitation>
and ovipositor
<figureCitation id="C2808BB72A52D854FCF27F701CAAB0BD" box="[859,937,1663,1688]" captionStart="FIGURES 3641" captionStartId="18.[151,264,1697,1719]" captionTargetBox="[184,1416,280,1673]" captionTargetId="figure@18.[151,1436,241,1676]" captionTargetPageId="18" captionText="FIGURES 3641. Calleida excelsa Bates. 36 habitus and labels of lectotype (female, Carin Chebà, Myanmar, MSNG), scale bar = 2.0 mm; 37 habitus of paralectotype (female, Senmigion, Myanmar, MNHN), scale bar = 2.0 mm; 38 habitus (female, Yunnan, China), scale bar = 2.0 mm; 39 female reproductive tract (Chiang Mai, Thailand), scale bar = 0.5 mm; 40 ovipositor (Yunnan, China), scale bar = 0.2 mm; 41 apex of immature aedeagus, left-lateral view, scale bar = 0.2 mm." httpUri="https://zenodo.org/record/1302727/files/figure.png" pageId="19" pageNumber="20">fig. 40</figureCitation>
): spermatheca digitiform, a little longer than the pedicel, with very short basal projection; surface faintly and finely whorled; spermathecal pedicel with base weakly expanded; spermathecal gland duct inserted at base of the basal projection, long and slender, about two times as long as spermatheca; glandular area strongly expanded, about one fifth as long as gland duct. Gonocoxite II of ovipositor sub-rectangular, as long as 2.5 times the basal width; base much wider than apex; inner margin with several setae, extending from the middle to apex; outer margin distinctly curved at middle, only slightly setose in the apical third; apex obliquely truncate, with irregular membranous extension on the outer part.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A52D854FF6E7E731C01B1F8" pageId="19" pageNumber="20" type="distribution">
<paragraph id="5A0497322A52D854FF6E7E731C01B1F8" blockId="19.[151,1437,151,2013]" pageId="19" pageNumber="20">
<emphasis id="68CF4B202A52D854FF6E7E731D9EB1B0" bold="true" box="[199,669,1916,1941]" pageId="19" pageNumber="20">Geographical Distribution and habitat.</emphasis>
Distribution range as shown in
<figureCitation id="C2808BB72A52D854FBA07E721B50B1B1" box="[1033,1107,1916,1940]" captionStart="MAP 3" captionStartId="21.[151,209,1134,1156]" captionTargetBox="[151,1436,195,1112]" captionTargetId="figure@21.[151,1436,195,1112]" captionText="MAP 3 Distributions for species in the C. lativittis group: C. lativittis Chaudoir (green); C. excelsa Bates (blue); C. jelineki sp. n. (red)." httpUri="https://zenodo.org/record/1302731/files/figure.png" pageId="19" pageNumber="20">map 3</figureCitation>
:
<collectingCountry id="22ACD7A22A52D854FBC97E731B9AB1B0" box="[1120,1177,1916,1941]" name="India" pageId="19" pageNumber="20">India</collectingCountry>
(
<collectingRegion id="987F59D02A52D854FB007E731A18B1B0" box="[1193,1307,1916,1941]" country="India" name="Nagaland" pageId="19" pageNumber="20">Nagaland</collectingRegion>
);
<collectingCountry id="22ACD7A22A52D854FA867E721A9FB1B0" box="[1327,1436,1917,1941]" name="Myanmar" pageId="19" pageNumber="20">Myanmar</collectingCountry>
(Carin Chebà, Senmigion,
<collectingRegion id="987F59D02A52D854FE6D7EAF1D1CB19D" box="[452,543,1952,1976]" country="Myanmar" name="Yangon" pageId="19" pageNumber="20">Yangon</collectingRegion>
); N
<collectingCountry id="22ACD7A22A52D854FDE77EAF1DB1B19C" box="[590,690,1952,1977]" name="Thailand" pageId="19" pageNumber="20">Thailand</collectingCountry>
(Chiang Dao,
<collectingRegion id="987F59D02A52D854FCF17EAF1CE1B19C" box="[856,994,1952,1977]" country="Thailand" name="Chiang Mai" pageId="19" pageNumber="20">Chiang Mai</collectingRegion>
); SW
<collectingCountry id="22ACD7A22A52D854FB817EAF1B6CB19C" box="[1064,1135,1952,1977]" name="China" pageId="19" pageNumber="20">China</collectingCountry>
:
<collectingRegion id="987F59D02A52D854FBD37EAE1BCCB19D" box="[1146,1231,1953,1976]" country="China" name="Yunnan" pageId="19" pageNumber="20">Yunnan</collectingRegion>
(Yingjiang, Ruili, Xishuangbanna). In deciduous and sub-tropical forests.
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>

View file

@ -0,0 +1,289 @@
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<mods:title id="D42B1828C0E0708D5E695AE5A07B6F2E">Revision of the Oriental species of Calleida Latreille (sensu lato). Part 1: Introduction, groups of species, and species of six species groups (Coleoptera: Carabidae: Lebiini)</mods:title>
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<subSubSection id="12A1C4B92A51D857FF3E78E71D50B427" box="[151,595,487,514]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="5A0497322A51D857FF3E78E71D50B427" blockId="16.[151,595,487,514]" box="[151,595,487,514]" pageId="16" pageNumber="17">
<heading id="014C205E2A51D857FF3E78E71D50B427" bold="true" box="[151,595,487,514]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="68CF4B202A51D857FF3E78E71D50B427" bold="true" box="[151,595,487,514]" pageId="16" pageNumber="17">
[3]
<taxonomicName id="9DBBECB12A51D857FF6978E81D50B427" ID-CoL="5X83Q" authority="Chaudoir, 1872" authorityName="Chaudoir Lectotypus" authorityYear="1872" box="[192,595,487,514]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="lativittis">
<emphasis id="68CF4B202A51D857FF6978E81E8EB424" bold="true" box="[192,397,487,513]" italics="true" pageId="16" pageNumber="17">Calleida lativittis</emphasis>
<bibRefCitation id="3E2AEAC32A51D857FE3D78E71D50B427" box="[404,595,488,514]" pageId="16" pageNumber="17" refId="ref29572" refString="Chaudoir, B. M. de. (1872) Monographie des Callidides. Annales de la Societe Entomologique de Belgique, 15 (1871 - 1872), 97 - 204." type="journal article">Chaudoir, 1872</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A51D857FF3E7B211A39B461" box="[151,1338,558,580]" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="5A0497322A51D857FF3E7B211A39B461" blockId="16.[151,1338,558,580]" box="[151,1338,558,580]" pageId="16" pageNumber="17">
<taxonomicName id="9DBBECB12A51D857FF3E7B211D1FB461" ID-CoL="5X83Q" authority="Chaudoir, 1872: 113" authorityName="Chaudoir Lectotypus" authorityPageNumber="113" authorityYear="1872" box="[151,540,558,580]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="lativittis">
<emphasis id="68CF4B202A51D857FF3E7B211E44B461" box="[151,327,558,580]" italics="true" pageId="16" pageNumber="17">Calleida lativittis</emphasis>
<bibRefCitation id="3E2AEAC32A51D857FEE47B211EEDB461" box="[333,494,558,580]" pageId="16" pageNumber="17" refId="ref29572" refString="Chaudoir, B. M. de. (1872) Monographie des Callidides. Annales de la Societe Entomologique de Belgique, 15 (1871 - 1872), 97 - 204." type="journal article">Chaudoir, 1872</bibRefCitation>
: 113
</taxonomicName>
(
<typeStatus id="850029902A51D857FD807B211D55B461" box="[553,598,558,580]" pageId="16" pageNumber="17">type</typeStatus>
locality: “Deccan”;
<typeStatus id="850029902A51D857FC8F7B211C80B461" box="[806,899,558,580]" pageId="16" pageNumber="17" type="lectotype">lectotype</typeStatus>
deposited in MNHN);
<bibRefCitation id="3E2AEAC32A51D857FBC47B211A00B461" box="[1133,1283,558,580]" pageId="16" pageNumber="17" refId="ref30044" refString="Jedlicka, A. (1963) Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae-Brachyninae (Coleoptera, Carabidae). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden, 28, 269 - 579." type="journal article">Jedlička, 1963</bibRefCitation>
: 435.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A51D857FF3E7B7E1B18B5A3" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="5A0497322A51D857FF3E7B7E1EDBB4F7" blockId="16.[151,1437,625,2018]" pageId="16" pageNumber="17">
<emphasis id="68CF4B202A51D857FF3E7B7E1EC1B4AF" bold="true" box="[151,450,625,650]" pageId="16" pageNumber="17">
<typeStatus id="850029902A51D857FF3E7B7E1FD1B4AF" box="[151,210,625,650]" pageId="16" pageNumber="17">Type</typeStatus>
material examined:
</emphasis>
<materialsCitation id="EAD39D6F2A51D857FE637B7E1CD7B488" ID-GBIF-Occurrence="1852047373" collectorName="A. Casale" location="Coll. Jeakes" municipality="Coll. Jeakes" pageId="16" pageNumber="17" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
<typeStatus id="850029902A51D857FE637B7E1D37B4AF" box="[458,564,625,650]" pageId="16" pageNumber="17" type="lectotype">lectotype</typeStatus>
, designated herein, female, &quot;
<collectingMunicipality id="BA600D482A51D857FCDE7B7E1B05B4AF" box="[887,1030,625,650]" pageId="16" pageNumber="17">Coll. Jeakes</collectingMunicipality>
&quot;, &quot;Ex Musaeo Chaudoir&quot;; &quot;
<taxonomicName id="9DBBECB12A51D857FA957B7E1EEBB48B" authority="Chaudoir Lectotypus" authorityName="Chaudoir Lectotypus" authorityYear="1872" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="lativittis">
<emphasis id="68CF4B202A51D857FA957B7E1FF0B48B" italics="true" pageId="16" pageNumber="17">Calleida lativittis</emphasis>
Chaudoir Lectotypus
</taxonomicName>
<specimenCount id="4CBD5CBB2A51D857FE5B7B991D04B488" box="[498,519,662,685]" pageId="16" pageNumber="17" type="female"></specimenCount>
<collectorName id="F74EF2E42A51D857FDB97B9A1D7CB488" box="[528,639,661,685]" pageId="16" pageNumber="17">A. Casale</collectorName>
des. 2010&quot; (MNHN,
<figureCitation id="C2808BB72A51D857FCDE7B9A1CC5B488" box="[887,966,661,685]" captionStart="FIGURES 3035" captionStartId="17.[151,264,1228,1250]" captionTargetBox="[160,1405,496,1200]" captionTargetId="figure@17.[151,1436,481,1207]" captionTargetPageId="17" captionText="FIGURES 3035. Calleida lativittis Chaudoir. 30 habitus and labels of lectotype (female, &quot;Deccan&quot;, MNHN), scale bar = 2.0 mm; 31 habitus of paralectotype (female, &quot;Deccan&quot;, MNHN), scale bar = 2.0 mm; 3235 male genitalia, scale bar = 0.5 mm: 32 median lobe of aedeagus, dorsal view; 33 left paramere of aedeagus; 34 median lobe of aedeagus, left-lateral view; 35 male sternum IX." httpUri="https://zenodo.org/record/1302725/files/figure.png" pageId="16" pageNumber="17">fig. 30</figureCitation>
);
</materialsCitation>
<typeStatus id="850029902A51D857FC757B9A1B75B488" box="[988,1142,661,685]" pageId="16" pageNumber="17" type="paralectotype">paralectotype</typeStatus>
, one female, &quot;Ex Musaeo Chaudoir&quot; (
<collectionCode id="3CAA0FF72A51D857FEB07BB51E70B4F7" box="[281,371,698,722]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="17">MNHN</collectionCode>
,
<figureCitation id="C2808BB72A51D857FED77BB61EC9B4F7" box="[382,458,697,722]" captionStart="FIGURES 3035" captionStartId="17.[151,264,1228,1250]" captionTargetBox="[160,1405,496,1200]" captionTargetId="figure@17.[151,1436,481,1207]" captionTargetPageId="17" captionText="FIGURES 3035. Calleida lativittis Chaudoir. 30 habitus and labels of lectotype (female, &quot;Deccan&quot;, MNHN), scale bar = 2.0 mm; 31 habitus of paralectotype (female, &quot;Deccan&quot;, MNHN), scale bar = 2.0 mm; 3235 male genitalia, scale bar = 0.5 mm: 32 median lobe of aedeagus, dorsal view; 33 left paramere of aedeagus; 34 median lobe of aedeagus, left-lateral view; 35 male sternum IX." httpUri="https://zenodo.org/record/1302725/files/figure.png" pageId="16" pageNumber="17">fig. 31</figureCitation>
).
</paragraph>
<paragraph id="5A0497322A51D857FF6E7BD21B18B5A3" blockId="16.[151,1437,625,2018]" pageId="16" pageNumber="17">
<emphasis id="68CF4B202A51D857FF6E7BD21D23B4D3" bold="true" box="[199,544,733,758]" pageId="16" pageNumber="17">Non-type material examined</emphasis>
:
<emphasis id="68CF4B202A51D857FD987BD21D79B4D3" bold="true" box="[561,634,733,758]" pageId="16" pageNumber="17">
<collectingCountry id="22ACD7A22A51D857FD987BD21D76B4D3" box="[561,629,733,758]" name="India" pageId="16" pageNumber="17">India</collectingCountry>
:
</emphasis>
one male, “Ind. Mus. Bhogaon Purneah Dist. N. Bengal.
<date id="2E05B1F22A51D857FABE7BD11A7AB4D3" box="[1303,1401,734,758]" pageId="16" pageNumber="17" value="2009-03-19">19.III.09</date>
C. Paiua&quot;, “Ind. Mus. Calcutta&quot;, “H.E. Andrewes Coll. B.M.
<date id="2E05B1F22A51D857FC817A0E1C89B53F" box="[808,906,769,794]" pageId="16" pageNumber="17" value="1945" valueMax="1997">1945-97</date>
&quot;, “
<taxonomicName id="9DBBECB12A51D857FC037A0E1A5DB53F" authority="Chd. H. E. Andrewes" authorityName="Chd. H.E. Andrewes" box="[938,1374,769,794]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="lativittis">Calleida lativittis Chd. H.E. Andrewes</taxonomicName>
det.” (
<collectionCode id="3CAA0FF72A51D857FF367A291FFBB51B" box="[159,248,806,830]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="16" pageNumber="17">NHML</collectionCode>
); one male, “Coll. Melly” (no locality) (
<collectionCode id="3CAA0FF72A51D857FD417A291C2BB51B" box="[744,808,806,830]" country="Germany" httpUri="http://grbio.org/cool/mqei-3i36" name="Culture Collection of Algae at the University of Marburg" pageId="16" pageNumber="17">CCA</collectionCode>
); two females, “
<collectingCountry id="22ACD7A22A51D857FC597A2A1B33B51B" box="[1008,1072,805,830]" name="India" pageId="16" pageNumber="17">India</collectingCountry>
”; “Muséum Paris ex Coll. M. Maindron Coll. G. Babault 1930” (one of them identified as “
<emphasis id="68CF4B202A51D857FCE97A461CA9B547" box="[832,938,841,866]" italics="true" pageId="16" pageNumber="17">rubricata</emphasis>
?”) (
<collectionCode id="3CAA0FF72A51D857FC737A451B35B547" box="[986,1078,842,866]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="17">MNHN</collectionCode>
); one male, two females “
<collectingCountry id="22ACD7A22A51D857FAFD7A461A92B547" box="[1364,1425,841,866]" name="India" pageId="16" pageNumber="17">India</collectingCountry>
” (
<collectionCode id="3CAA0FF72A51D857FF367A611FDDB5A3" box="[159,222,878,902]" country="Germany" httpUri="http://grbio.org/cool/mqei-3i36" name="Culture Collection of Algae at the University of Marburg" pageId="16" pageNumber="17">CCA</collectionCode>
).
<emphasis id="68CF4B202A51D857FF587A621E78B5A3" bold="true" box="[241,379,877,902]" pageId="16" pageNumber="17">
<collectingCountry id="22ACD7A22A51D857FF587A621E78B5A3" box="[241,379,877,902]" name="Bangladesh" pageId="16" pageNumber="17">Bangladesh</collectingCountry>
</emphasis>
: one female, “Comercolly. Indes” [=Kumerkhali] (
<collectionCode id="3CAA0FF72A51D857FC117A611B0CB5A3" box="[952,1039,878,902]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="16" pageNumber="17">MSNG</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A51D857FF6E7A9E1E9AB27B" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="5A0497322A51D857FF6E7A9E1E9AB27B" blockId="16.[151,1437,625,2018]" pageId="16" pageNumber="17">
<emphasis id="68CF4B202A51D857FF6E7A9E1E42B58F" bold="true" box="[199,321,913,938]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
With the character states of
<taxonomicName id="9DBBECB12A51D857FD2E7A9E1DEBB58F" authorityName="Latreille" authorityYear="1824" box="[647,744,913,938]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A51D857FD2E7A9E1DEBB58F" box="[647,744,913,938]" italics="true" pageId="16" pageNumber="17">Calleida</emphasis>
</taxonomicName>
of the
<taxonomicName id="9DBBECB12A51D857FC907A9E1C96B58F" box="[825,917,913,938]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="lativittis">
<emphasis id="68CF4B202A51D857FC907A9E1C96B58F" box="[825,917,913,938]" italics="true" pageId="16" pageNumber="17">lativittis</emphasis>
</taxonomicName>
species group, but distinguishable from other known species by the following character combination: dorsal side glabrous except some erect setae near the anterior angles of pronotum. Elytra metallic green, disc with reddish patch widened on the inner three intervals in the apical half; antennomeres 311 markedly darkened; femora blackish in the apical two thirds, base reddish; tibiae dark brownish in some individuals. Head narrow and convex. Pronotum elongate-cordate; disc convex, with narrow lateral furrows.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A51D857FF6E7D661C9DB2EF" pageId="16" pageNumber="17" type="discussion">
<paragraph id="5A0497322A51D857FF6E7D661C9DB2EF" blockId="16.[151,1437,625,2018]" pageId="16" pageNumber="17">
Closely related and similar to
<taxonomicName id="9DBBECB12A51D857FD817D651D9DB2A7" box="[552,670,1129,1154]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="jelineki" status="sp. nov.">
<emphasis id="68CF4B202A51D857FD817D651D9DB2A7" box="[552,670,1129,1154]" italics="true" pageId="16" pageNumber="17">C. jelineki</emphasis>
</taxonomicName>
<emphasis id="68CF4B202A51D857FD017D661DE8B2A7" bold="true" box="[680,747,1129,1154]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="73FCF65B2A51D857FD017D661DE8B2A7" box="[680,747,1129,1154]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
described below, but easily recognized by: pronotum more transverse, more widened in the middle, with lateral sides more deeply sinuate in the basal fifth, the elytral disc with larger reddish patch, and by antennae and legs mostly darkened.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A51D856FF6E7DDA1D33B788" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="description">
<paragraph id="5A0497322A51D857FF6E7DDA1E13B337" blockId="16.[151,1437,625,2018]" pageId="16" pageNumber="17">
<emphasis id="68CF4B202A51D857FF6E7DDA1E5AB2CB" bold="true" box="[199,345,1237,1262]" pageId="16" pageNumber="17">Description.</emphasis>
The original description provided by
<bibRefCitation id="3E2AEAC32A51D857FCA17DDA1CC0B2CB" box="[776,963,1237,1262]" pageId="16" pageNumber="17" refId="ref29572" refString="Chaudoir, B. M. de. (1872) Monographie des Callidides. Annales de la Societe Entomologique de Belgique, 15 (1871 - 1872), 97 - 204." type="journal article">Chaudoir (1872)</bibRefCitation>
is here completed with several additional characters.
</paragraph>
<paragraph id="5A0497322A51D857FF6E7C121B2DB35B" blockId="16.[151,1437,625,2018]" pageId="16" pageNumber="17">
<emphasis id="68CF4B202A51D857FF6E7C121E85B313" box="[199,390,1309,1334]" italics="true" pageId="16" pageNumber="17">General features</emphasis>
as in
<figureCitation id="C2808BB72A51D857FE787C121D5BB313" box="[465,600,1309,1334]" captionStart="FIGURES 3035" captionStartId="17.[151,264,1228,1250]" captionTargetBox="[160,1405,496,1200]" captionTargetId="figure@17.[151,1436,481,1207]" captionTargetPageId="17" captionText="FIGURES 3035. Calleida lativittis Chaudoir. 30 habitus and labels of lectotype (female, &quot;Deccan&quot;, MNHN), scale bar = 2.0 mm; 31 habitus of paralectotype (female, &quot;Deccan&quot;, MNHN), scale bar = 2.0 mm; 3235 male genitalia, scale bar = 0.5 mm: 32 median lobe of aedeagus, dorsal view; 33 left paramere of aedeagus; 34 median lobe of aedeagus, left-lateral view; 35 male sternum IX." httpUri="https://zenodo.org/record/1302725/files/figure.png" pageId="16" pageNumber="17">figs. 30, 31</figureCitation>
. Medium sized: L: 10.0
<quantity id="9D433AD72A51D857FCD77C111CE4B310" box="[894,999,1310,1334]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="16" pageNumber="17" unit="mm" value="10.8">10.8 mm</quantity>
; TL: 9.0
<quantity id="9D433AD72A51D857FBF27C121BB5B310" box="[1115,1206,1309,1334]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="16" pageNumber="17" unit="mm" value="9.5">9.5 mm</quantity>
. Body elongate and slender, glabrous except a very short, hardly visible pubescence on the abdominal sternum VII and some erect setae near the anterior angles of pronotum; elytra slightly widened in the posterior fifth.
</paragraph>
<paragraph id="5A0497322A51D857FF6E7C861C35B02B" blockId="16.[151,1437,625,2018]" pageId="16" pageNumber="17">
<emphasis id="68CF4B202A51D857FF6E7C861E15B387" box="[199,278,1417,1442]" italics="true" pageId="16" pageNumber="17">Colour</emphasis>
: head, pronotum, lateral margins and epipleura of elytra, and underside reddish; antennae darkened from the third antennomere to apex; femora blackish in apical two thirds or more, base reddish, tibiae reddish in some specimens, darkened in others; elytra metallic blue greenish, with wide reddish patch extended from base to apex, widened on the inner three intervals in the apical half.
</paragraph>
<paragraph id="5A0497322A51D857FF6E7F151D4DB05F" blockId="16.[151,1437,625,2018]" pageId="16" pageNumber="17">
<emphasis id="68CF4B202A51D857FF6E7F151EF9B017" box="[199,506,1561,1586]" italics="true" pageId="16" pageNumber="17">Lustre and microsculpture</emphasis>
: shiny; head with vanished microsculpture; pronotum mostly without distinct microsculpture, except some fine transverse meshes on disc and middle part of the basal area; elytra with distinct microsculpture in isodiametric meshes.
</paragraph>
<paragraph id="5A0497322A51D857FF6E7F8A1CA8B0E7" blockId="16.[151,1437,625,2018]" pageId="16" pageNumber="17">
<emphasis id="68CF4B202A51D857FF6E7F8A1E07B0BB" box="[199,260,1669,1694]" italics="true" pageId="16" pageNumber="17">Head</emphasis>
wide, sub-convex, narrower than pronotum, smooth, with some shallow oblique frontal wrinkles; frontal foveae short but deep. Genae short, convex; eyes large and prominent.
</paragraph>
<paragraph id="5A0497322A51D857FF6E7FC11BD0B174" blockId="16.[151,1437,625,2018]" pageId="16" pageNumber="17">
<emphasis id="68CF4B202A51D857FF6E7FC11E35B0C0" box="[199,310,1742,1765]" italics="true" pageId="16" pageNumber="17">Pronotum</emphasis>
cordate, moderately transverse (ratio PW/PL: 1.16), widened in the middle, with deep lateral furrows. Disc with very shallow transverse wrinkles. Basal foveae deep and elongate. Lateral sides slightly reflexed, rounded in the anterior half, only sinuate in the basal sixth, then parallel sided anteriorly to the basal angles, which are rectangular. Base straight. Anterior angles effaced, each with some short setae.
</paragraph>
<paragraph id="5A0497322A51D857FF6E7E521B99B19B" blockId="16.[151,1437,625,2018]" pageId="16" pageNumber="17">
<emphasis id="68CF4B202A51D857FF6E7E521E0EB153" box="[199,269,1885,1910]" italics="true" pageId="16" pageNumber="17">Elytra</emphasis>
elongate and narrow (EL/EW: 1.81), slightly widened at the posterior fifth. Striae deep, punctate; intervals flat or sub-convex, with evident deep punctures; interval 8 not tumid near apex. Umbilicate series of 15 16 pores along stria 8. Apical truncation straight; outer apical angle not thickened, rounded.
</paragraph>
<paragraph id="5A0497322A51D856FF6E7EC61EA4B6F0" blockId="16.[151,1437,625,2018]" lastBlockId="17.[151,1436,151,429]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<emphasis id="68CF4B202A51D857FF6E7EC61E4FB1C7" box="[199,332,1993,2018]" italics="true" pageId="16" pageNumber="17">Ventral side</emphasis>
: abdominal sternum VII with sparse, hardly visible pubescence, with two (one on each side) setae in male, four (two on each side) setae in female; apical margin of abdominal sternum VII notched in the middle in male, straight in female.
</paragraph>
<paragraph id="5A0497322A50D856FF6E79D01A98B7AD" blockId="17.[151,1436,151,429]" pageId="17" pageNumber="18">
<emphasis id="68CF4B202A50D856FF6E79D01E6FB6DD" box="[199,364,223,248]" italics="true" pageId="17" pageNumber="18">Male genitalia</emphasis>
as in
<figureCitation id="C2808BB72A50D856FE0779D01D32B6DD" box="[430,561,223,248]" captionStart="FIGURES 3035" captionStartId="17.[151,264,1228,1250]" captionTargetBox="[160,1405,496,1200]" captionTargetId="figure@17.[151,1436,481,1207]" captionTargetPageId="17" captionText="FIGURES 3035. Calleida lativittis Chaudoir. 30 habitus and labels of lectotype (female, &quot;Deccan&quot;, MNHN), scale bar = 2.0 mm; 31 habitus of paralectotype (female, &quot;Deccan&quot;, MNHN), scale bar = 2.0 mm; 3235 male genitalia, scale bar = 0.5 mm: 32 median lobe of aedeagus, dorsal view; 33 left paramere of aedeagus; 34 median lobe of aedeagus, left-lateral view; 35 male sternum IX." httpUri="https://zenodo.org/record/1302725/files/figure.png" pageId="17" pageNumber="18">figs. 3234</figureCitation>
: median lobe of aedeagus regularly curved on the dorsal side, undulate on the ventral side; apical lamina flat, developed, angulate dorsally, fully rounded ventrally; endophallus with two chitinized V-shaped copulatory pieces close to the left lateral margin of the median lobe, their apex adjacent to the base of the apical orifice, close to each other at base, the ventral one narrower, sinuate and acute at apex, the dorsal one shorter and wider. Left paramere dorsally depressed, 1.5 times as long as wide; right paramere not emarginate.
</paragraph>
<paragraph id="5A0497322A50D856FF6E789B1D33B788" blockId="17.[151,1436,151,429]" box="[199,560,404,429]" pageId="17" pageNumber="18">
<emphasis id="68CF4B202A50D856FF6E789B1E1FB788" box="[199,284,404,429]" italics="true" pageId="17" pageNumber="18">Female</emphasis>
genitalia: not examined.
</paragraph>
</subSubSection>
<caption id="0EC4C7BA2A50D856FF3E7DC31E13B31B" httpUri="https://zenodo.org/record/1302725/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,264,1228,1250]" targetBox="[160,1405,496,1200]" targetPageId="17">
<paragraph id="5A0497322A50D856FF3E7DC31E13B31B" blockId="17.[151,1437,1228,1342]" pageId="17" pageNumber="18">
<emphasis id="68CF4B202A50D856FF3E7DC31E57B2C7" bold="true" box="[151,340,1228,1250]" pageId="17" pageNumber="18">FIGURES 3035.</emphasis>
<taxonomicName id="9DBBECB12A50D856FEF27DC31D74B2C7" authority="Chaudoir." authorityName="Chaudoir Lectotypus" authorityYear="1872" box="[347,631,1228,1250]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lativittis">
<emphasis id="68CF4B202A50D856FEF27DC31D0FB2C7" bold="true" box="[347,524,1228,1250]" italics="true" pageId="17" pageNumber="18">Calleida lativittis</emphasis>
Chaudoir.
</taxonomicName>
<emphasis id="68CF4B202A50D856FDD47DC31D95B2C7" bold="true" box="[637,662,1228,1250]" pageId="17" pageNumber="18">30</emphasis>
habitus and labels of lectotype (female, &quot;Deccan&quot;, MNHN), scale bar = 2.0 mm;
<emphasis id="68CF4B202A50D856FF627DE41FE6B324" bold="true" box="[203,229,1259,1281]" pageId="17" pageNumber="18">31</emphasis>
habitus of paralectotype (female, &quot;Deccan&quot;, MNHN), scale bar = 2.0 mm;
<emphasis id="68CF4B202A50D856FC7E7DE41B15B324" bold="true" box="[983,1046,1259,1281]" pageId="17" pageNumber="18">3235</emphasis>
male genitalia, scale bar = 0.5 mm:
<emphasis id="68CF4B202A50D856FA2A7DE41A9EB324" bold="true" box="[1411,1437,1259,1281]" pageId="17" pageNumber="18">32</emphasis>
median lobe of aedeagus, dorsal view;
<emphasis id="68CF4B202A50D856FD857C051D46B305" bold="true" box="[556,581,1290,1312]" pageId="17" pageNumber="18">33</emphasis>
left paramere of aedeagus;
<emphasis id="68CF4B202A50D856FCCE7C051C83B305" bold="true" box="[871,896,1290,1312]" pageId="17" pageNumber="18">34</emphasis>
median lobe of aedeagus, left-lateral view;
<emphasis id="68CF4B202A50D856FAE07C051A61B305" bold="true" box="[1353,1378,1290,1312]" pageId="17" pageNumber="18">35</emphasis>
male sternum IX.
</paragraph>
</caption>
<subSubSection id="12A1C4B92A50D856FF6E7C6A1EFEB05E" pageId="17" pageNumber="18" type="distribution">
<paragraph id="5A0497322A50D856FF6E7C6A1A43B386" blockId="17.[151,1436,1381,1659]" pageId="17" pageNumber="18">
<emphasis id="68CF4B202A50D856FF6E7C6A1DADB35B" bold="true" box="[199,686,1381,1406]" pageId="17" pageNumber="18">Geographical distribution and habitat.</emphasis>
<collectingRegion id="987F59D02A50D856FD147C681C1BB35B" box="[701,792,1383,1406]" country="American Samoa" name="Western" pageId="17" pageNumber="18">Western</collectingRegion>
and Northern
<collectingCountry id="22ACD7A22A50D856FC7D7C6A1B11B35B" box="[980,1042,1381,1406]" name="India" pageId="17" pageNumber="18">India</collectingCountry>
: “Deccan” (
<typeStatus id="850029902A50D856FB017C691BD9B35B" box="[1192,1242,1382,1406]" pageId="17" pageNumber="18">type</typeStatus>
locality);
<collectingRegion id="987F59D02A50D856FAF77C6A1A9FB35B" box="[1374,1436,1381,1406]" country="India" name="Bihar" pageId="17" pageNumber="18">Bihar</collectingRegion>
(Purnea);
<collectingRegion id="987F59D02A50D856FEAF7C851E93B386" box="[262,400,1418,1443]" country="India" name="Uttarakhand" pageId="17" pageNumber="18">Uttarakhand</collectingRegion>
(Haldwani);
<collectingCountry id="22ACD7A22A50D856FD8F7C851DAEB386" box="[550,685,1418,1443]" name="Bangladesh" pageId="17" pageNumber="18">Bangladesh</collectingCountry>
: Kumerkhali (
<figureCitation id="C2808BB72A50D856FCE57C851C97B387" box="[844,916,1418,1442]" captionStart="MAP 3" captionStartId="21.[151,209,1134,1156]" captionTargetBox="[151,1436,195,1112]" captionTargetId="figure@21.[151,1436,195,1112]" captionText="MAP 3 Distributions for species in the C. lativittis group: C. lativittis Chaudoir (green); C. excelsa Bates (blue); C. jelineki sp. n. (red)." httpUri="https://zenodo.org/record/1302731/files/figure.png" pageId="17" pageNumber="18">map 3</figureCitation>
). Dubious locality:
<collectingRegion id="987F59D02A50D856FBDB7C851BBDB387" box="[1138,1214,1418,1442]" country="India" name="Assam" pageId="17" pageNumber="18">Assam</collectingRegion>
(Sibsagar).
</paragraph>
<paragraph id="5A0497322A50D856FF6E7CA01FEAB3CE" blockId="17.[151,1436,1381,1659]" pageId="17" pageNumber="18">No data are available on the habitat where specimens were sampled, probably in deciduous and sub-tropical forests.</paragraph>
<paragraph id="5A0497322A50D856FF6E7CFA1EFEB05E" blockId="17.[151,1436,1381,1659]" pageId="17" pageNumber="18">
The record from
<collectingRegion id="987F59D02A50D856FE2E7CF91ED0B02B" box="[391,467,1526,1550]" country="India" name="Assam" pageId="17" pageNumber="18">Assam</collectingRegion>
(Sibsagar) was only mentioned by
<bibRefCitation id="3E2AEAC32A50D856FCCD7CFA1B28B02B" box="[868,1067,1525,1550]" pageId="17" pageNumber="18" refId="ref28962" refString="Andrewes, H. E. (1930) Catalogue of Indian Insects. Part 18. Carabidae. Government of India, Central Publication Branch, Calcutta, 389 pp." type="book">Andrewes (1930)</bibRefCitation>
. We didn't examine any material from
<collectingRegion id="987F59D02A50D856FF7D7F151E23B017" box="[212,288,1562,1586]" country="India" name="Assam" pageId="17" pageNumber="18">Assam</collectingRegion>
or other adjacent localities of northeast
<collectingCountry id="22ACD7A22A50D856FD767F151C1FB016" box="[735,796,1562,1587]" name="India" pageId="17" pageNumber="18">India</collectingCountry>
. Considering the fact that
<bibRefCitation id="3E2AEAC32A50D856FB967F151A02B017" box="[1087,1281,1562,1587]" pageId="17" pageNumber="18" refId="ref28815" refString="Andrewes, H. E. (1921) Notes on Synonymy and on some Types of Oriental Carabidae in various foreign collections. Transactions of the Royal entomological Society, London, 69, 145 - 190. https: // doi. org / 10.1111 / j. 1365 - 2311.1921. tb 02805. x" type="journal article">Andrewes (1921)</bibRefCitation>
confused this species with
<taxonomicName id="9DBBECB12A50D856FE8F7F311E9BB073" box="[294,408,1597,1622]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="excelsa">
<emphasis id="68CF4B202A50D856FE8F7F311E9BB073" box="[294,408,1597,1622]" italics="true" pageId="17" pageNumber="18">C. excelsa</emphasis>
</taxonomicName>
, and
<collectingRegion id="987F59D02A50D856FE7C7F311D21B073" box="[469,546,1598,1622]" country="India" name="Assam" pageId="17" pageNumber="18">Assam</collectingRegion>
is very close to one confirmed record (
<collectingRegion id="987F59D02A50D856FC7A7F311B13B073" box="[979,1040,1598,1622]" country="Philippines" name="Naga" pageId="17" pageNumber="18">Naga</collectingRegion>
hills) of
<taxonomicName id="9DBBECB12A50D856FBDD7F311BE5B073" box="[1140,1254,1597,1622]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="excelsa">
<emphasis id="68CF4B202A50D856FBDD7F311BE5B073" box="[1140,1254,1597,1622]" italics="true" pageId="17" pageNumber="18">C. excelsa</emphasis>
</taxonomicName>
, we believe that this locality is a dubious record.
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>

View file

@ -0,0 +1,277 @@
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<mods:title id="F7A409BEB40594524C154F14702532E8">Revision of the Oriental species of Calleida Latreille (sensu lato). Part 1: Introduction, groups of species, and species of six species groups (Coleoptera: Carabidae: Lebiini)</mods:title>
</mods:titleInfo>
<mods:name id="83C6C1F817EAA035A1E2B4EDCB67B6CB" type="personal">
<mods:role id="6222CF31772ABE8D0997D7C02B158FA4">
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<mods:namePart id="188930291E8A798F5952DDD395DF3715">Casale, Achille</mods:namePart>
</mods:name>
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</mods:role>
<mods:namePart id="7B0A3AE8A9A73C11AAAB4E5B9830AC16">Shi, Hongliang</mods:namePart>
</mods:name>
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<mods:titleInfo id="8770A3AFF1451C274E72A49C7855588D">
<mods:title id="716D41FEF8E6C439F741F83D0EF44D17">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3CB40ED0BECA725435D69C5C42D9C84B">
<mods:date id="A75144FFFC5AE0AD355C759EE329BA11">2018</mods:date>
<mods:detail id="CB17DA7BED0D193AE951D8BEC2BF079A" type="pubDate">
<mods:number id="D9D6642C4665CB08045782360D0BA99D">2018-06-29</mods:number>
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</mods:detail>
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<mods:classification id="21FB67443F8DAAE2F5519A5C29853437">journal volume</mods:classification>
<mods:identifier id="FB4EAB7CC4CF7AB0EB357F45728119C3" type="CLB-Dataset">29772</mods:identifier>
<mods:identifier id="310C124FE3A2F1216404E0EE4C8E373C" type="DOI">10.11646/zootaxa.4442.1.1</mods:identifier>
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</mods:mods>
<subSection id="33348ED92A55D851FF3E79971C0FB7D0" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21" type="multiple">
<treatment id="D21226242A55D851FF3E79971C0FB7D0" ID-DOI="http://doi.org/10.5281/zenodo.5950861" ID-GBIF-Taxon="146043504" ID-Zenodo-Dep="5950861" LSID="urn:lsid:plazi:treatment:D21226242A55D851FF3E79971C0FB7D0" httpUri="http://treatment.plazi.org/id/D21226242A55D851FF3E79971C0FB7D0" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21">
<subSubSection id="12A1C4B92A55D853FF3E79971D82B697" box="[151,641,151,178]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="5A0497322A55D853FF3E79971D82B697" blockId="20.[151,641,151,178]" box="[151,641,151,178]" pageId="20" pageNumber="21">
<heading id="014C205E2A55D853FF3E79971D82B697" bold="true" box="[151,641,151,178]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<emphasis id="68CF4B202A55D853FF3E79971D82B697" bold="true" box="[151,641,151,178]" pageId="20" pageNumber="21">
[5]
<taxonomicName id="9DBBECB12A55D853FF6979981D32B697" ID-CoL="8MCZV" authority="Casale &amp; Shi" authorityName="Casale &amp; Shi" box="[192,561,151,178]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="jelineki" status="sp. nov.">
<emphasis id="68CF4B202A55D853FF6979981E84B694" bold="true" box="[192,391,151,177]" italics="true" pageId="20" pageNumber="21">Calleida jelineki</emphasis>
Casale &amp; Shi
</taxonomicName>
,
<taxonomicNameLabel id="73FCF65B2A55D853FD9679971D82B697" box="[575,641,152,178]" pageId="20" pageNumber="21" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A55D853FF3E79D11C3AB7EB" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="5A0497322A55D853FF3E79D11D31B6D2" blockId="20.[151,1436,222,787]" box="[151,562,222,247]" pageId="20" pageNumber="21">
<emphasis id="68CF4B202A55D853FF3E79D11E3AB6D2" bold="true" box="[151,313,222,247]" pageId="20" pageNumber="21">
<typeStatus id="850029902A55D853FF3E79D11FD1B6D2" box="[151,210,222,247]" pageId="20" pageNumber="21">Type</typeStatus>
locality:
</emphasis>
S.
<collectingCountry id="22ACD7A22A55D853FEF479D11E98B6D2" box="[349,411,222,247]" name="India" pageId="20" pageNumber="21">India</collectingCountry>
: Coimbatore.
</paragraph>
<paragraph id="5A0497322A55D853FF6E780D1BF0B7A2" blockId="20.[151,1436,222,787]" pageId="20" pageNumber="21">
<emphasis id="68CF4B202A55D853FF6E780D1E74B73E" bold="true" box="[199,375,258,283]" pageId="20" pageNumber="21">
<typeStatus id="850029902A55D853FF6E780D1E01B73E" box="[199,258,258,283]" pageId="20" pageNumber="21">Type</typeStatus>
material:
</emphasis>
<materialsCitation id="EAD39D6F2A55D853FED4780D1CA8B71A" ID-GBIF-Occurrence="1852047394" country="India" location="Call." municipality="Coimbatore Nathan" pageId="20" pageNumber="21" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="850029902A55D853FED4780D1EEBB73E" box="[381,488,258,283]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
,
<specimenCount id="4CBD5CBB2A55D853FE5D780D1D40B73E" box="[500,579,258,283]" pageId="20" pageNumber="21" type="female">female</specimenCount>
, &quot;S.
<collectingCountry id="22ACD7A22A55D853FDDF780D1DB3B73E" box="[630,688,258,283]" name="India" pageId="20" pageNumber="21">India</collectingCountry>
<collectingMunicipality id="BA600D482A55D853FD1E780D1C9BB73E" box="[695,920,258,283]" pageId="20" pageNumber="21">Coimbatore Nathan</collectingMunicipality>
”, “
<taxonomicName id="9DBBECB12A55D853FC1E780D1B1CB73E" authorityName="Jedlicka" authorityYear="1953" box="[951,1055,258,283]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="fukiensis">fukiensis</taxonomicName>
det. Ing. Jedlička”, “
<emphasis id="68CF4B202A55D853FAA2780D1A9FB73E" box="[1291,1436,258,283]" italics="true" pageId="20" pageNumber="21">
<location id="5F64C1E92A55D853FAA2780D1A40B73E" LSID="urn:lsid:plazi:treatment:D21226242A55D851FF3E79971C0FB7D0:5F64C1E92A55D853FAA2780D1A40B73E" box="[1291,1347,258,283]" country="India" municipality="Coimbatore Nathan" name="Call." pageId="20" pageNumber="21">
<taxonomicName id="9DBBECB12A55D853FAA2780D1A3CB73E" box="[1291,1343,258,283]" class="Insecta" family="Cerambycidae" genus="Call" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus" status="sp. nov.">Call</taxonomicName>
.
</location>
<taxonomicName id="9DBBECB12A55D853FAE3780D1A9FB73E" box="[1354,1436,258,283]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="jelineki" status="sp. nov.">jelineki</taxonomicName>
</emphasis>
<taxonomicNameLabel id="73FCF65B2A55D853FF3E78281FD4B71B" box="[151,215,295,318]" pageId="20" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
Holotypus ♀ 1997 A. Casale det.” [in litt.] (NMPC,
<figureCitation id="C2808BB72A55D853FCE478291C9DB71A" box="[845,926,294,319]" captionStart="FIGURES 4243" captionStartId="20.[151,264,1934,1956]" captionTargetBox="[211,1385,839,1887]" captionTargetId="figure@20.[151,1436,839,1912]" captionTargetPageId="20" captionText="FIGURES 4243. Calleida jelineki sp. n. 42 habitus and labels of holotype (female, Coimbatore, Inida, NMPC), scale bar = 2.0 mm; 43 female reproductive tract and ovipositor (holotype), scale bar = 0.5 mm; abbreviations: sp spermatheca, pd spermathecal pedicel, gd spermathecal gland duct, ga glandular area." httpUri="https://zenodo.org/record/1302729/files/figure.png" pageId="20" pageNumber="21">fig. 42</figureCitation>
).
</materialsCitation>
<typeStatus id="850029902A55D853FC1C78281B25B71A" box="[949,1062,295,319]" pageId="20" pageNumber="21" type="paratype">Paratypes</typeStatus>
: one female, “Samanahally près Bangalore R. B. Tabourel IV.
<date id="2E05B1F22A55D853FE4A78441D35B746" box="[483,566,331,355]" pageId="20" pageNumber="21" value="1899-05">V.1899</date>
”, “
<emphasis id="68CF4B202A55D853FDF178451DEAB746" box="[600,745,330,355]" italics="true" pageId="20" pageNumber="21">
<taxonomicName id="9DBBECB12A55D853FDF178451D88B746" box="[600,651,330,355]" class="Insecta" family="Cerambycidae" genus="Call" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus" status="sp. nov.">Call</taxonomicName>
.
<taxonomicName id="9DBBECB12A55D853FD3E78451DEAB746" box="[663,745,330,355]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="jelineki" status="sp. nov.">jelineki</taxonomicName>
</emphasis>
<taxonomicNameLabel id="73FCF65B2A55D853FD5B78441C2CB747" box="[754,815,331,354]" pageId="20" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
<typeStatus id="850029902A55D853FC9E78441CA6B746" box="[823,933,331,355]" pageId="20" pageNumber="21" type="paratype">Paratypus</typeStatus>
<specimenCount id="4CBD5CBB2A55D853FC0478441CC1B747" box="[941,962,331,354]" pageId="20" pageNumber="21"></specimenCount>
A. Casale &amp; H. Shi det. 2016” (
<collectionCode id="3CAA0FF72A55D853FA9D78441A93B746" box="[1332,1424,331,355]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="20" pageNumber="21">MNHN</collectionCode>
); one female, “E. Ind[ia]”, “
<emphasis id="68CF4B202A55D853FE1678611D4CB7A2" box="[447,591,366,391]" italics="true" pageId="20" pageNumber="21">
<taxonomicName id="9DBBECB12A55D853FE1678611EF1B7A2" box="[447,498,366,391]" class="Insecta" family="Cerambycidae" genus="Call" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus" status="sp. nov.">Call</taxonomicName>
.
<taxonomicName id="9DBBECB12A55D853FE5478611D4CB7A2" box="[509,591,366,391]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="jelineki" status="sp. nov.">jelineki</taxonomicName>
</emphasis>
<taxonomicNameLabel id="73FCF65B2A55D853FDFE78601D90B7A3" box="[599,659,367,390]" pageId="20" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
<typeStatus id="850029902A55D853FD3378601C0BB7A2" box="[666,776,367,391]" pageId="20" pageNumber="21" type="paratype">Paratypus</typeStatus>
<specimenCount id="4CBD5CBB2A55D853FCA678601C27B7A3" box="[783,804,367,390]" pageId="20" pageNumber="21"></specimenCount>
A. Casale &amp; H. Shi det. 2016” (
<collectionCode id="3CAA0FF72A55D853FB2778601BE4B7A2" box="[1166,1255,367,391]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="20" pageNumber="21">NHML</collectionCode>
).
</paragraph>
<paragraph id="5A0497322A55D853FF6E789D1C3AB7EB" blockId="20.[151,1436,222,787]" pageId="20" pageNumber="21">
<emphasis id="68CF4B202A55D853FF6E789D1D0EB78E" bold="true" box="[199,525,402,427]" pageId="20" pageNumber="21">
Notes on the
<typeStatus id="850029902A55D853FECC789D1E9BB78E" box="[357,408,402,427]" pageId="20" pageNumber="21">type</typeStatus>
material.
</emphasis>
The
<typeStatus id="850029902A55D853FDE8789D1DA0B78E" box="[577,675,402,427]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
of this new species was misidentified by Jedlička, as it appears by the identification label, as
<taxonomicName id="9DBBECB12A55D853FE1678B81D41B7EA" authorityName="Jedlicka" authorityYear="1953" box="[447,578,438,463]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="fukiensis">
<emphasis id="68CF4B202A55D853FE1678B81D41B7EA" box="[447,578,438,463]" italics="true" pageId="20" pageNumber="21">C. fukiensis</emphasis>
</taxonomicName>
described by himself.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A55D853FF6E78D51FFDB47B" pageId="20" pageNumber="21" type="etymology">
<paragraph id="5A0497322A55D853FF6E78D51FFDB47B" blockId="20.[151,1436,222,787]" pageId="20" pageNumber="21">
<emphasis id="68CF4B202A55D853FF6E78D51E8FB7D6" bold="true" box="[199,396,474,499]" pageId="20" pageNumber="21">Specific epithet.</emphasis>
This new species is dedicated to our friend and colleague Josef Jelínek, well known coleopterist and former curator at the Entomological Department of the National Museum in
<collectingRegion id="987F59D02A55D853FB4778F01A3CB432" box="[1262,1343,511,535]" country="Czech Republic" name="Praha" pageId="20" pageNumber="21">Prague</collectingRegion>
, for his invaluable help over many years to one of the authors (A.C.) in examination of entomological material in that museum.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A55D852FF6E7B651CB7B315" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="diagnosis">
<paragraph id="5A0497322A55D853FF6E7B651BDAB536" blockId="20.[151,1436,222,787]" pageId="20" pageNumber="21">
<emphasis id="68CF4B202A55D853FF6E7B651E42B4A6" bold="true" box="[199,321,618,643]" pageId="20" pageNumber="21">Diagnosis.</emphasis>
With the character states of
<taxonomicName id="9DBBECB12A55D853FD2C7B651DE5B4A6" authorityName="Latreille" authorityYear="1824" box="[645,742,618,643]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A55D853FD2C7B651DE5B4A6" box="[645,742,618,643]" italics="true" pageId="20" pageNumber="21">Calleida</emphasis>
</taxonomicName>
of the
<taxonomicName id="9DBBECB12A55D853FC9F7B651C91B4A6" authorityName="Chaudoir Lectotypus" authorityYear="1872" box="[822,914,618,643]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="lativittis">
<emphasis id="68CF4B202A55D853FC9F7B651C91B4A6" box="[822,914,618,643]" italics="true" pageId="20" pageNumber="21">lativittis</emphasis>
</taxonomicName>
species group, but distinct from the two other species of this group by the following character combination: elytra very elongate and narrow (EL/EW 1.87), parallel sided, subconvex; elytra metallic blue greenish, without discal reddish patch, with the only sutural interval reddish; femora darkened at about apical fourth, all antennomeres reddish brown; pubescence reduced to a few, short, hardly visible setae on abdominal sterna, and a few, short setae at the base of pronotal disc.
</paragraph>
<caption id="0EC4C7BA2A55D853FF3E7E811C50B1C7" httpUri="https://zenodo.org/record/1302729/files/figure.png" pageId="20" pageNumber="21" startId="20.[151,264,1934,1956]" targetBox="[211,1385,839,1887]" targetPageId="20">
<paragraph id="5A0497322A55D853FF3E7E811C50B1C7" blockId="20.[151,1436,1934,2018]" pageId="20" pageNumber="21">
<emphasis id="68CF4B202A55D853FF3E7E811D0BB181" bold="true" box="[151,520,1934,1956]" pageId="20" pageNumber="21">
FIGURES 4243.
<taxonomicName id="9DBBECB12A55D853FEF57E811D0BB181" box="[348,520,1934,1956]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="jelineki" status="sp. nov.">
<emphasis id="68CF4B202A55D853FEF57E811D0BB181" bold="true" box="[348,520,1934,1956]" italics="true" pageId="20" pageNumber="21">Calleida jelineki</emphasis>
</taxonomicName>
</emphasis>
<taxonomicNameLabel id="73FCF65B2A55D853FDA67E801D44B181" box="[527,583,1935,1956]" pageId="20" pageNumber="21" rank="species">sp. n.</taxonomicNameLabel>
<emphasis id="68CF4B202A55D853FDE77E811D6BB181" bold="true" box="[590,616,1934,1956]" pageId="20" pageNumber="21">42</emphasis>
habitus and labels of holotype (female, Coimbatore, Inida, NMPC), scale bar = 2.0 mm;
<emphasis id="68CF4B202A55D853FF527EA21E16B1E6" bold="true" box="[251,277,1965,1987]" pageId="20" pageNumber="21">43</emphasis>
female reproductive tract and ovipositor (holotype), scale bar = 0.5 mm; abbreviations:
<emphasis id="68CF4B202A55D853FB647EA21BE6B1E6" bold="true" box="[1229,1253,1965,1987]" pageId="20" pageNumber="21">sp</emphasis>
spermatheca,
<emphasis id="68CF4B202A55D853FA297EA21A9FB1E6" bold="true" box="[1408,1436,1965,1987]" pageId="20" pageNumber="21">pd</emphasis>
spermathecal pedicel,
<emphasis id="68CF4B202A55D853FED17EC31E90B1C7" bold="true" box="[376,403,1996,2018]" pageId="20" pageNumber="21">gd</emphasis>
spermathecal gland duct,
<emphasis id="68CF4B202A55D853FD357EC31DB6B1C7" bold="true" box="[668,693,1996,2018]" pageId="20" pageNumber="21">ga</emphasis>
glandular area.
</paragraph>
</caption>
<caption id="0EC4C7BA2A54D852FF3E7D611FEBB286" httpUri="https://zenodo.org/record/1302731/files/figure.png" pageId="21" pageNumber="22" startId="21.[151,209,1134,1156]" targetBox="[151,1436,195,1112]" targetPageId="21">
<paragraph id="5A0497322A54D852FF3E7D611FEBB286" blockId="21.[151,1436,1134,1187]" pageId="21" pageNumber="22">
<emphasis id="68CF4B202A54D852FF3E7D611FE7B2A1" bold="true" box="[151,228,1134,1156]" pageId="21" pageNumber="22">MAP 3</emphasis>
Distributions for species in the
<taxonomicName id="9DBBECB12A54D852FD8E7D601D9BB2A1" authorityName="Chaudoir Lectotypus" authorityYear="1872" box="[551,664,1134,1156]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="lativittis">
<emphasis id="68CF4B202A54D852FD8E7D601D9BB2A1" box="[551,664,1134,1156]" italics="true" pageId="21" pageNumber="22">C. lativittis</emphasis>
</taxonomicName>
group:
<taxonomicName id="9DBBECB12A54D852FD4E7D601CBFB2A1" authority="Chaudoir" authorityName="Chaudoir Lectotypus" authorityYear="1872" box="[743,956,1134,1156]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="lativittis">
<emphasis id="68CF4B202A54D852FD4E7D601C5BB2A1" box="[743,856,1134,1156]" italics="true" pageId="21" pageNumber="22">C. lativittis</emphasis>
Chaudoir
</taxonomicName>
(green);
<taxonomicName id="9DBBECB12A54D852FBB07D601BC3B2A1" authority="Bates" authorityName="Bates" box="[1049,1216,1134,1156]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="excelsa">
<emphasis id="68CF4B202A54D852FBB07D601B81B2A1" box="[1049,1154,1134,1156]" italics="true" pageId="21" pageNumber="22">C. excelsa</emphasis>
Bates
</taxonomicName>
(blue);
<taxonomicName id="9DBBECB12A54D852FAB97D601A7AB2A1" box="[1296,1401,1134,1156]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="jelineki" status="sp. nov.">
<emphasis id="68CF4B202A54D852FAB97D601A7AB2A1" box="[1296,1401,1134,1156]" italics="true" pageId="21" pageNumber="22">C. jelineki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73FCF65B2A54D852FAD67D601FA9B286" pageId="21" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
(red).
</paragraph>
</caption>
<paragraph id="5A0497322A54D852FF6E7DC01CB7B315" blockId="21.[151,1437,1231,2012]" pageId="21" pageNumber="22">
The new species is similar to
<taxonomicName id="9DBBECB12A54D852FD8B7DDF1C15B2CD" authority="Chaudoir" authorityName="Chaudoir Lectotypus" authorityYear="1872" box="[546,790,1231,1256]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="lativittis">
<emphasis id="68CF4B202A54D852FD8B7DDF1DA2B2CD" box="[546,673,1231,1256]" italics="true" pageId="21" pageNumber="22">C. lativittis</emphasis>
Chaudoir
</taxonomicName>
, but easily distinguishable by the pronotum narrower in front, with lateral sides less sinuate in front of the basal angles, which are obtuse, not prominent outside, the femora darkened only at apex, and by other characters specified in description.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A54D851FF6E7C341A36B789" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="description">
<paragraph id="5A0497322A54D852FF6E7C341D63B35D" blockId="21.[151,1437,1231,2012]" pageId="21" pageNumber="22">
<emphasis id="68CF4B202A54D852FF6E7C341E5AB371" bold="true" box="[199,345,1339,1364]" pageId="21" pageNumber="22">Description.</emphasis>
<emphasis id="68CF4B202A54D852FECD7C341D27B371" box="[356,548,1339,1364]" italics="true" pageId="21" pageNumber="22">General features</emphasis>
as in
<figureCitation id="C2808BB72A54D852FDDD7C341DC5B371" box="[628,710,1339,1364]" captionStart="FIGURES 4243" captionStartId="20.[151,264,1934,1956]" captionTargetBox="[211,1385,839,1887]" captionTargetId="figure@20.[151,1436,839,1912]" captionTargetPageId="20" captionText="FIGURES 4243. Calleida jelineki sp. n. 42 habitus and labels of holotype (female, Coimbatore, Inida, NMPC), scale bar = 2.0 mm; 43 female reproductive tract and ovipositor (holotype), scale bar = 0.5 mm; abbreviations: sp spermatheca, pd spermathecal pedicel, gd spermathecal gland duct, ga glandular area." httpUri="https://zenodo.org/record/1302729/files/figure.png" pageId="21" pageNumber="22">fig. 42</figureCitation>
: Medium sized: L:
<quantity id="9D433AD72A54D852FC187C341B0CB376" box="[945,1039,1339,1364]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="21" pageNumber="22" unit="mm" value="8.5">8.5 mm</quantity>
; TL: 8.0 mm. Body elongate and slender, parallel sided; elytra subconvex.
</paragraph>
<paragraph id="5A0497322A54D852FF6E7C8C1E4DB3C1" blockId="21.[151,1437,1231,2012]" pageId="21" pageNumber="22">
<emphasis id="68CF4B202A54D852FF6E7C8C1E15B3B9" box="[199,278,1411,1436]" italics="true" pageId="21" pageNumber="22">Colour</emphasis>
: head, pronotum, lateral margins and epipleura of elytra, underside, antennae and legs rufous; femora blackish at about apical fourth; elytra metallic blue greenish, without reddish patch on disc, with only sutural interval reddish.
</paragraph>
<paragraph id="5A0497322A54D852FF6E7CFE1DABB075" blockId="21.[151,1437,1231,2012]" pageId="21" pageNumber="22">
<emphasis id="68CF4B202A54D852FF6E7CFE1EF9B02D" box="[199,506,1519,1544]" italics="true" pageId="21" pageNumber="22">Lustre and microsculpture</emphasis>
: shiny; head with vanished microsculpture; pronotum mostly without distinct microsculpture, except some fine transverse meshes on disc and on middle part of the basal area; elytra with distinct microsculpture in isodiametric meshes.
</paragraph>
<paragraph id="5A0497322A54D852FF6E7F541A0BB0BD" blockId="21.[151,1437,1231,2012]" pageId="21" pageNumber="22">
<emphasis id="68CF4B202A54D852FF6E7F541E07B051" box="[199,260,1627,1652]" italics="true" pageId="21" pageNumber="22">Head</emphasis>
wide, subconvex, narrower than pronotum, smooth or with some oblique frontal wrinkles between the frontal foveae; frontal foveae very shallow. Genae rather short and convex; eyes large and prominent.
</paragraph>
<paragraph id="5A0497322A54D852FF6E7FAA1A69B10D" blockId="21.[151,1437,1231,2012]" pageId="21" pageNumber="22">
<emphasis id="68CF4B202A54D852FF6E7FAA1E35B099" box="[199,310,1701,1724]" italics="true" pageId="21" pageNumber="22">Pronotum</emphasis>
cordate, narrow (ratio PW/PL: 1.07), with lateral furrows widened in the basal fourth. Disc with shallow transverse wrinkles and several fine punctures; a few short pubescence present on the base of pronotal disc. Lateral sides slightly reflexed, rounded in the anterior half, slightly sinuate in the basal sixth. Anterior angles effaced, each with 810 setae. Basal foveae short but deep. Basal margin oblique at sides, basal angles obtuse.
</paragraph>
<paragraph id="5A0497322A54D852FF6E7E3C1A34B1B1" blockId="21.[151,1437,1231,2012]" pageId="21" pageNumber="22">
<emphasis id="68CF4B202A54D852FF6E7E3C1E0EB169" box="[199,269,1843,1868]" italics="true" pageId="21" pageNumber="22">Elytra</emphasis>
elongate and narrow (EL/EW: 1.87), parallel sided. Striae very deep, finely punctate; intervals convex in the posterior half, deeply punctate, without pubescence; eighth interval not tumid near the apex. Umbilicate series of 15 pores along eighth stria. Apical truncation straight; outer apical angle not thickened, rounded.
</paragraph>
<paragraph id="5A0497322A54D852FF6E7E901B5FB1F9" blockId="21.[151,1437,1231,2012]" pageId="21" pageNumber="22">
<emphasis id="68CF4B202A54D852FF6E7E901E4DB19D" box="[199,334,1951,1976]" italics="true" pageId="21" pageNumber="22">Ventral side</emphasis>
: abdominal sterna with sparse and short pubescence; abdominal sternum VII with four (two on each side) setae in female; apical margin of abdominal sternum VII straight in female.
</paragraph>
<paragraph id="5A0497322A57D851FF6E79981EE9B695" blockId="22.[151,1436,151,501]" box="[199,490,151,176]" pageId="22" pageNumber="23">
<emphasis id="68CF4B202A57D851FF6E79981E68B695" box="[199,363,151,176]" italics="true" pageId="22" pageNumber="23">Male genitalia</emphasis>
: unknown.
</paragraph>
<paragraph id="5A0497322A57D851FF6E79B31A36B789" blockId="22.[151,1436,151,501]" pageId="22" pageNumber="23">
<emphasis id="68CF4B202A57D851FF6E79B31E89B6F0" box="[199,394,188,213]" italics="true" pageId="22" pageNumber="23">Female genitalia</emphasis>
(reproductive tract and ovipositor,
<figureCitation id="C2808BB72A57D851FC8079B31C75B6F0" box="[809,886,188,213]" captionStart="FIGURES 4243" captionStartId="20.[151,264,1934,1956]" captionTargetBox="[211,1385,839,1887]" captionTargetId="figure@20.[151,1436,839,1912]" captionTargetPageId="20" captionText="FIGURES 4243. Calleida jelineki sp. n. 42 habitus and labels of holotype (female, Coimbatore, Inida, NMPC), scale bar = 2.0 mm; 43 female reproductive tract and ovipositor (holotype), scale bar = 0.5 mm; abbreviations: sp spermatheca, pd spermathecal pedicel, gd spermathecal gland duct, ga glandular area." httpUri="https://zenodo.org/record/1302729/files/figure.png" pageId="22" pageNumber="23">fig. 43</figureCitation>
):spermatheca digitiform, much longer than the pedicel, with very short basal projection; surface faintly and finely whorled; spermathecal pedicel short, with basal half expanded, apical half slender; spermathecal gland duct inserted at base of the projection, long and slender, a little longer than spermatheca; glandular area inflated, about one eight length as gland duct. Gonocoxite II of ovipositor sub-rectangular, as long as three times of the basal width; base slightly wider than apex; inner margin with several setae, extending from the basal third to apex; outer margin distinctly curved at middle, only slightly setose in the apical third; apex obliquely truncate, with irregular membranous extension on the outer part.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A57D851FF6E78B81C0FB7D0" pageId="22" pageNumber="23" type="distribution">
<paragraph id="5A0497322A57D851FF6E78B81C0FB7D0" blockId="22.[151,1436,151,501]" pageId="22" pageNumber="23">
<emphasis id="68CF4B202A57D851FF6E78B81DA0B7F5" bold="true" box="[199,675,439,464]" pageId="22" pageNumber="23">Geographical Distribution and habitat.</emphasis>
S.
<collectingCountry id="22ACD7A22A57D851FD6478B81C08B7F5" box="[717,779,439,464]" name="India" pageId="22" pageNumber="23">India</collectingCountry>
: Coimbatore, Bangalore, “E
<collectingCountry id="22ACD7A22A57D851FBF078B81B96B7F5" box="[1113,1173,439,464]" name="India" pageId="22" pageNumber="23">India</collectingCountry>
” (
<figureCitation id="C2808BB72A57D851FB1878B71BFEB7F5" box="[1201,1277,440,464]" captionStart="MAP 3" captionStartId="21.[151,209,1134,1156]" captionTargetBox="[151,1436,195,1112]" captionTargetId="figure@21.[151,1436,195,1112]" captionText="MAP 3 Distributions for species in the C. lativittis group: C. lativittis Chaudoir (green); C. excelsa Bates (blue); C. jelineki sp. n. (red)." httpUri="https://zenodo.org/record/1302731/files/figure.png" pageId="22" pageNumber="23">map 3</figureCitation>
). No data are available on the habitat where specimens were sampled.
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>

View file

@ -0,0 +1,83 @@
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<mods:title id="C9C6CB9F6A5C6A7F0A0551EAB31AA53C">Revision of the Oriental species of Calleida Latreille (sensu lato). Part 1: Introduction, groups of species, and species of six species groups (Coleoptera: Carabidae: Lebiini)</mods:title>
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<subSubSection id="12A1C4B92A57D851FF3E7B4D1D46B479" box="[151,581,578,604]" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="5A0497322A57D851FF3E7B4D1D46B479" blockId="22.[151,581,578,604]" box="[151,581,578,604]" pageId="22" pageNumber="23">
<heading id="014C205E2A57D851FF3E7B4D1D46B479" bold="true" box="[151,581,578,604]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<emphasis id="68CF4B202A57D851FF3E7B4D1D46B479" bold="true" box="[151,581,578,604]" pageId="22" pageNumber="23">
5.
<taxonomicName id="9DBBECB12A57D851FF1D7B4D1E96B479" ID-CoL="PLBR" box="[180,405,578,604]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="terminata">
<emphasis id="68CF4B202A57D851FF1D7B4D1E96B479" bold="true" box="[180,405,578,604]" italics="true" pageId="22" pageNumber="23">Calleida terminata</emphasis>
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species group
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<emphasis id="68CF4B202A57D851FF3E7B861EA3B487" bold="true" box="[151,416,649,674]" pageId="22" pageNumber="23">Diagnostic characters</emphasis>
: With the character states of
<taxonomicName id="9DBBECB12A57D851FCA57B861C6FB487" authorityName="Latreille" authorityYear="1824" box="[780,876,649,674]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A57D851FCA57B861C6FB487" box="[780,876,649,674]" italics="true" pageId="22" pageNumber="23">Calleida</emphasis>
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(in the widest sense, see above), restricted or amplified as follows: medium sized species (L: 8.511.0 mm). Pronotum and ventral side metallic green, or brown with marked metallic reflection, vertex with reddish patch; elytra metallic green or cupreous, with golden cupreous or purple hue at shoulders and/or near apex. Elytral outer apical angles markedly widened, forming distinctly sharp, acute angles. Abdominal sternum VII in male markedly notched at middle, with one seta on each side in males, two setae in females. Median lobe of aedeagus in lateral aspect with a distinct, very prominent ventral lobe.
</paragraph>
<paragraph id="5A0497322A57D851FF6E7A6E1EB0B5E7" blockId="22.[151,1436,649,1070]" pageId="22" pageNumber="23">
<materialsCitation id="EAD39D6F2A57D851FF6E7A6E1EB0B5E7" ID-GBIF-Occurrence="1852047922" collectorName="SE Asia." country="Vietnam" location="Sumatra" pageId="22" pageNumber="23" specimenCount="1" stateProvince="Sumatra">
<emphasis id="68CF4B202A57D851FF6E7A6E1E8BB55F" bold="true" box="[199,392,865,890]" pageId="22" pageNumber="23">Taxonomic note</emphasis>
: Three species are attributed to this group, distributed in the Malay Peninsula,
<collectingRegion id="987F59D02A57D851FA9E7A6D1A94B55F" box="[1335,1431,866,890]" country="Indonesia" name="Sumatera Utara" pageId="22" pageNumber="23">Sumatra</collectingRegion>
, Borneo and southern
<collectingCountry id="22ACD7A22A57D851FE3B7A8A1EF2B5BB" box="[402,497,901,926]" name="Vietnam" pageId="22" pageNumber="23">Vietnam</collectingCountry>
(
<figureCitation id="C2808BB72A57D851FDAA7A891D53B5BB" box="[515,592,902,926]" captionStart="MAP 4" captionStartId="29.[151,209,1076,1098]" captionTargetBox="[151,1436,195,1054]" captionTargetId="figure@29.[151,1436,195,1054]" captionTargetPageId="29" captionText="MAP 4 Distributions for species in the C. terminata group: C. terminata Waterhouse (green); C. corporaali Andrewes (blue); C. viet sp. n. (red)." httpUri="https://zenodo.org/record/1302739/files/figure.png" pageId="22" pageNumber="23">map 4</figureCitation>
). Species of this group are rare in collections, known so far from a few specimens from
<collectorName id="F74EF2E42A57D851FEF97AA51EB0B5E7" box="[336,435,937,962]" pageId="22" pageNumber="23">SE Asia.</collectorName>
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<paragraph id="5A0497322A57D851FF6E7AC21EFEB20B" blockId="22.[151,1436,649,1070]" pageId="22" pageNumber="23">The monophyly of this species group seems to be clear, for the following synapomorphic characters: elytral outer apical angles sharp, acute; male abdominal sternum VII markedly notched; median lobe of aedeagus with a distinct lobe on the ventral side.</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,117 @@
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Key to species of
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<emphasis id="68CF4B202A57D851FEC47D8E1D4CB2BE" bold="true" box="[365,591,1153,1179]" italics="true" pageId="22" pageNumber="23">Calleida terminata</emphasis>
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species group
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<keyLead id="E14A34072A57D851FF3E7DC81A9FB311" pageId="22" pageNumber="23">
1. Head and pronotum dark brownish, only with weak metallic greenish reflection on pronotal disc; femora dark brownish, with very faint metallic green reflection. Pronotal disc without or with very shallow transverse wrinkles. Outer apical angles of elytra sharp, not or only slightly prominent apically. Larger sized species (L: 9.512.0). (Distribution:
<collectingCountry id="22ACD7A22A57D851FB707C0D1A39B332" box="[1241,1338,1282,1303]" name="Indonesia" pageId="22" pageNumber="23">Indonesia</collectingCountry>
:
<collectingRegion id="987F59D02A57D851FAEC7C0D1A9BB332" box="[1349,1432,1282,1303]" country="Indonesia" name="Sumatera Utara" pageId="22" pageNumber="23">Sumatra</collectingRegion>
; Malay Peninsula and Borneo)......................................................[7]
<taxonomicName id="9DBBECB12A57D851FB1B7C101A9FB311" ID-CoL="5X5ZW" authority="Andrewes" authorityName="Andr. H. E. Andrewes" authorityYear="1929" box="[1202,1436,1311,1332]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="corporaali">
<emphasis id="68CF4B202A57D851FB1B7C101A37B311" box="[1202,1332,1311,1332]" italics="true" pageId="22" pageNumber="23">C. corporaali</emphasis>
Andrewes
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- Head, pronotum and elytra strongly metallic green or cupreous; femora with distinct metallic green reflection. Pronotal disc with distinct transverse wrinkles. Outer apical angles of elytra acutely prominent apically. Smaller sized species (L:
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)................................................................................................ 2
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2. Antennae brownish, first antennomere with weak metallic green reflection; femora and underside completely metallic green. Median lobe of aedeagus in lateral aspect with ventral margin straight beyond the main ventral lobe (
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).
<collectingCountry id="22ACD7A22A57D851FB587CBD1A4EB3E2" box="[1265,1357,1458,1479]" name="Malaysia" pageId="22" pageNumber="23">Malaysia</collectingCountry>
: Borneo (
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<collectingRegion id="987F59D02A57D851FF4E7CC01E3FB3C1" box="[231,316,1487,1508]" country="Malaysia" name="Sarawak" pageId="22" pageNumber="23">Sarawak</collectingRegion>
</collectingCountry>
)..................................................................... [6]
<taxonomicName id="9DBBECB12A57D851FB057CC01A98B3C1" ID-CoL="PLBR" authority="WaterHOUse" authorityName="Waterhouse" box="[1196,1435,1487,1508]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="30" phylum="Arthropoda" rank="species" species="terminata">
<emphasis id="68CF4B202A57D851FB057CC01A26B3C1" box="[1196,1317,1487,1508]" italics="true" pageId="22" pageNumber="23">C. terminata</emphasis>
Waterhouse
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<keyLead id="E14A34072A57D851FF3E7CE21A9FB018" pageId="22" pageNumber="23">
- Antennae completely brownish, first antennomere without metallic reflection; femora and underside brownish, with marked metallic green reflection. Median lobe of aedeagus in lateral aspect with ventral margin pointed beyond the main ventral lobe (
<figureCitation id="C2808BB72A57D851FF417F281E29B019" box="[232,298,1575,1596]" captionStart="FIGURES 5660" captionStartId="28.[151,264,1120,1142]" captionTargetBox="[159,1407,265,1093]" captionTargetId="figure@28.[151,1436,265,1099]" captionTargetPageId="28" captionText="FIGURES 5660. Calleida viet sp. n. 56 habitus and labels of holotype (male, Vung Tau, Vietnam, CCA), scale bar = 2.0 mm; 5759 male genitalia (holotype), scale bar = 0.5 mm: 57 median lobe of aedeagus, ventral view; 58 left paramere of aedeagus; 59 median lobe of aedeagus, left-lateral view; 60 female reproductive tract and ovipositor (paratype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302737/files/figure.png" pageId="22" pageNumber="23">fig. 59</figureCitation>
). Vietnam (Vung Tau).................................................... [8]
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Casale &amp; Shi
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,
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<mods:identifier id="C5E9A41C7C6EE8D269916A7785CE3453" type="DOI">10.11646/zootaxa.4442.1.1</mods:identifier>
<mods:identifier id="BBE534AFE2ED75974E5B3F79497DDF06" type="GBIF-Dataset">ded8bdb6-f8ba-4e0b-bed1-1ec08963edfd</mods:identifier>
<mods:identifier id="73BBCC4E74A8C5D035D98E65B78A3CBA" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1CE3F5FAFDE0120128F8563B873CF6D0" type="Zenodo-Dep">1302713</mods:identifier>
<mods:identifier id="F5FFB69CF48A4287DCAEE4ADBDCDF11D" type="ZooBank">195B0471-553A-4617-B901-E9DBD2323D14</mods:identifier>
</mods:mods>
<subSection id="33348ED92A57D85FFF3E7F8C1A6AB579" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23" type="multiple">
<treatment id="D21226242A57D85FFF3E7F8C1A6AB579" ID-DOI="http://doi.org/10.5281/zenodo.5950867" ID-GBIF-Taxon="146043502" ID-Zenodo-Dep="5950867" LSID="urn:lsid:plazi:treatment:D21226242A57D85FFF3E7F8C1A6AB579" httpUri="http://treatment.plazi.org/id/D21226242A57D85FFF3E7F8C1A6AB579" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23">
<subSubSection id="12A1C4B92A57D851FF3E7F8C1D8BB0B8" box="[151,648,1666,1693]" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="5A0497322A57D851FF3E7F8C1D8BB0B8" blockId="22.[151,648,1666,1693]" box="[151,648,1666,1693]" pageId="22" pageNumber="23">
<heading id="014C205E2A57D851FF3E7F8C1D8BB0B8" bold="true" box="[151,648,1666,1693]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<emphasis id="68CF4B202A57D851FF3E7F8C1D8BB0B8" bold="true" box="[151,648,1666,1693]" pageId="22" pageNumber="23">
[6]
<taxonomicName id="9DBBECB12A57D851FF697F8D1D8BB0B8" ID-CoL="PLBR" authority="Waterhouse, 1876" authorityName="Waterhouse" authorityYear="1876" box="[192,648,1666,1693]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="terminata">
<emphasis id="68CF4B202A57D851FF697F8D1EA1B0B9" bold="true" box="[192,418,1666,1692]" italics="true" pageId="22" pageNumber="23">Calleida terminata</emphasis>
<bibRefCitation id="3E2AEAC32A57D851FE007F8C1D8BB0B8" box="[425,648,1667,1693]" pageId="22" pageNumber="23" refId="ref30966" refString="Waterhouse, C. O. (1876) On various new genera and species of Coleoptera. The Transactions of the Royal Entomological Society London, 24 (1), 11 - 25. https: // doi. org / 10.1111 / j. 1365 - 2311.1876. tb 01122. x" type="journal article">Waterhouse, 1876</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A57D851FF3E7FCB1A68B0FF" box="[151,1387,1732,1754]" pageId="22" pageNumber="23" type="reference_group">
<paragraph id="5A0497322A57D851FF3E7FCB1A68B0FF" blockId="22.[151,1387,1732,1754]" box="[151,1387,1732,1754]" pageId="22" pageNumber="23">
<taxonomicName id="9DBBECB12A57D851FF3E7FCB1D2EB0FF" authority="Waterhouse, 1876: 11" authorityName="Waterhouse" authorityPageNumber="11" authorityYear="1876" box="[151,557,1732,1754]" class="Insecta" family="Carabidae" genus="Callida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="terminata">
<emphasis id="68CF4B202A57D851FF3E7FCB1E4FB0FF" box="[151,332,1732,1754]" italics="true" pageId="22" pageNumber="23">Callida terminata</emphasis>
<bibRefCitation id="3E2AEAC32A57D851FEFB7FCB1D08B0FF" box="[338,523,1732,1754]" pageId="22" pageNumber="23" refId="ref30966" refString="Waterhouse, C. O. (1876) On various new genera and species of Coleoptera. The Transactions of the Royal Entomological Society London, 24 (1), 11 - 25. https: // doi. org / 10.1111 / j. 1365 - 2311.1876. tb 01122. x" type="journal article">Waterhouse, 1876</bibRefCitation>
: 11
</taxonomicName>
(
<typeStatus id="850029902A57D851FD927FCB1D6BB0FF" box="[571,616,1732,1754]" pageId="22" pageNumber="23">type</typeStatus>
locality: Borneo,
<collectingCountry id="22ACD7A22A57D851FCB67FCB1C79B0FF" box="[799,890,1732,1754]" name="Malaysia" pageId="22" pageNumber="23">
<collectingRegion id="987F59D02A57D851FCB67FCB1C79B0FF" box="[799,890,1732,1754]" country="Malaysia" name="Sarawak" pageId="22" pageNumber="23">Sarawak</collectingRegion>
</collectingCountry>
;
<typeStatus id="850029902A57D851FC2A7FCB1CE3B0FF" box="[899,992,1732,1754]" pageId="22" pageNumber="23" type="lectotype">lectotype</typeStatus>
deposited in NHML);
<bibRefCitation id="3E2AEAC32A57D851FB617FCB1A42B0FF" box="[1224,1345,1732,1754]" pageId="22" pageNumber="23" refId="ref30886" refString="Stork, N. (1986) An annotated checklist of the Carabidae (including Cicindelinae, Rhysodinae and Paussinae) recorded from Borneo. In: Occasional Papers on Systematic Entomology. Fol. 2. British Museum (Natural History), London, pp. 1 - 24." type="book chapter">Stork, 1986</bibRefCitation>
: 16.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A57D851FF3E7E0E1CB2B1D7" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="5A0497322A57D851FF3E7E0E1D15B11B" blockId="22.[151,1436,1793,2034]" pageId="22" pageNumber="23">
<emphasis id="68CF4B202A57D851FF3E7E0E1EC1B13F" bold="true" box="[151,450,1793,1818]" pageId="22" pageNumber="23">
<typeStatus id="850029902A57D851FF3E7E0E1FD1B13F" box="[151,210,1793,1818]" pageId="22" pageNumber="23">Type</typeStatus>
material examined
</emphasis>
:
<materialsCitation id="EAD39D6F2A57D851FE7C7E0D1D15B11B" ID-GBIF-Occurrence="1852047908" location="Sarawak" pageId="22" pageNumber="23" specimenCount="1" specimenCount-male="1" stateProvince="Sarawak" typeStatus="lectotype">
<typeStatus id="850029902A57D851FE7C7E0D1D46B13F" box="[469,581,1794,1818]" pageId="22" pageNumber="23" type="lectotype">Lectotype</typeStatus>
designate herein,
<specimenCount id="4CBD5CBB2A57D851FC897E0E1C5AB13F" box="[800,857,1793,1818]" pageId="22" pageNumber="23" type="male">male</specimenCount>
, “Type H. T.”, “
<collectingRegion id="987F59D02A57D851FB887E0E1B8BB13F" box="[1057,1160,1793,1818]" country="Malaysia" name="Sarawak" pageId="22" pageNumber="23">Sarawak</collectingRegion>
”, “
<taxonomicName id="9DBBECB12A57D851FB047E0E1E42B11B" authority="C. Waterh." authorityName="C. Waterh." class="Insecta" family="Carabidae" genus="Callida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="terminata">Callida terminata C. Waterh. (Type)</taxonomicName>
” (NHML,
<figureCitation id="C2808BB72A57D851FE147E2A1D0AB11B" box="[445,521,1829,1854]" captionStart="FIGURES 4447" captionStartId="23.[151,263,1096,1118]" captionTargetBox="[179,1384,197,1065]" captionTargetId="figure@23.[151,1436,193,1074]" captionTargetPageId="23" captionText="FIGURES 4447. Calleida terminata Waterhouse. 44 habitus and labels of lectotype (male, Sarawak, Malaysia, NHML), scale bar = 2.0 mm; 45 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (lectotype), scale bar = 0.5 mm; 46 left and right parameres of aedeagus (lectotype), scale bar = 0.2 mm; 47 male sternum IX (lectotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302733/files/figure.png" pageId="22" pageNumber="23">fig. 44</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="5A0497322A57D851FF6E7E461C6CB1A3" blockId="22.[151,1436,1793,2034]" pageId="22" pageNumber="23">
<emphasis id="68CF4B202A57D851FF6E7E461D1EB147" bold="true" box="[199,541,1865,1890]" pageId="22" pageNumber="23">Non-type material examined</emphasis>
: one female, &quot;Borneo&quot;, &quot;Frey Coll., 1905.100&quot; (
<collectionCode id="3CAA0FF72A57D851FBE97E451B9AB147" box="[1088,1177,1866,1890]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="22" pageNumber="23">NHML</collectionCode>
); one female, &quot;46370&quot;, &quot;Borneo&quot;, &quot;Frey Coll., 1905.100&quot;; &quot;
<taxonomicName id="9DBBECB12A57D851FD8D7E621DF0B1A3" authority="Waterh." authorityName="Waterhouse" authorityYear="1876" box="[548,755,1901,1926]" class="Insecta" family="Carabidae" genus="Callida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="terminata">terminata Waterh.</taxonomicName>
&quot; (
<collectionCode id="3CAA0FF72A57D851FCA37E611C60B1A3" box="[778,867,1902,1926]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="22" pageNumber="23">NHML</collectionCode>
).
</paragraph>
<paragraph id="5A0497322A57D851FF6E7E9E1CB2B1D7" blockId="22.[151,1436,1793,2034]" pageId="22" pageNumber="23">
<emphasis id="68CF4B202A57D851FF6E7E9E1EE0B18F" bold="true" box="[199,483,1937,1962]" pageId="22" pageNumber="23">
Notes on
<typeStatus id="850029902A57D851FE937E9E1E6EB18F" box="[314,365,1937,1962]" pageId="22" pageNumber="23">type</typeStatus>
material.
</emphasis>
The original publication (
<bibRefCitation id="3E2AEAC32A57D851FCB97E9E1CDDB18F" box="[784,990,1937,1962]" pageId="22" pageNumber="23" refId="ref30966" refString="Waterhouse, C. O. (1876) On various new genera and species of Coleoptera. The Transactions of the Royal Entomological Society London, 24 (1), 11 - 25. https: // doi. org / 10.1111 / j. 1365 - 2311.1876. tb 01122. x" type="journal article">Waterhouse, 1876</bibRefCitation>
) didnt state or imply this species was based on a single specimen, although it seems so. According to the Recommendation 73F and Article 74.6 of
<collectionCode id="3CAA0FF72A57D851FF3E7ED51FD8B1D7" box="[151,219,2010,2034]" pageId="22" pageNumber="23">ICZN</collectionCode>
, we designated a
<typeStatus id="850029902A57D851FE097ED61D05B1D7" box="[416,518,2009,2034]" pageId="22" pageNumber="23" type="lectotype">lectotype</typeStatus>
herein rather than assume a
<typeStatus id="850029902A57D851FCEE7ED61CAEB1D7" box="[839,941,2009,2034]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<caption id="0EC4C7BA2A56D850FF3E7D471A5AB2BE" httpUri="https://zenodo.org/record/1302733/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,263,1096,1118]" targetBox="[179,1384,197,1065]" targetPageId="23">
<paragraph id="5A0497322A56D850FF3E7D471A5AB2BE" blockId="23.[151,1436,1095,1179]" pageId="23" pageNumber="24">
<emphasis id="68CF4B202A56D850FF3E7D471E52B27B" bold="true" box="[151,337,1096,1118]" pageId="23" pageNumber="24">FIGURES 4447.</emphasis>
<taxonomicName id="9DBBECB12A56D850FEF17D481D9EB27B" authority="Waterhouse." authorityName="Waterhouse." box="[344,669,1095,1118]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="terminata">
<emphasis id="68CF4B202A56D850FEF17D481D1AB278" bold="true" box="[344,537,1095,1117]" italics="true" pageId="23" pageNumber="24">Calleida terminata</emphasis>
Waterhouse.
</taxonomicName>
<emphasis id="68CF4B202A56D850FD0B7D471DB8B27B" bold="true" box="[674,699,1096,1118]" pageId="23" pageNumber="24">44</emphasis>
habitus and labels of lectotype (male, Sarawak, Malaysia, NHML), scale bar = 2.0 mm;
<emphasis id="68CF4B202A56D850FE837D691E40B259" bold="true" box="[298,323,1126,1148]" pageId="23" pageNumber="24">45</emphasis>
median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (lectotype), scale bar = 0.5 mm;
<emphasis id="68CF4B202A56D850FA2A7D691A9FB259" bold="true" box="[1411,1436,1126,1148]" pageId="23" pageNumber="24">46</emphasis>
left and right parameres of aedeagus (lectotype), scale bar = 0.2 mm;
<emphasis id="68CF4B202A56D850FCE47D8A1C65B2BE" bold="true" box="[845,870,1157,1179]" pageId="23" pageNumber="24">47</emphasis>
male sternum IX (lectotype), scale bar = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="12A1C4B92A56D850FF6E7DCD1C7AB03A" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="5A0497322A56D850FF6E7DCD1D86B396" blockId="23.[151,1437,1218,2035]" pageId="23" pageNumber="24">
<emphasis id="68CF4B202A56D850FF6E7DCD1E42B2FE" bold="true" box="[199,321,1218,1243]" pageId="23" pageNumber="24">Diagnosis.</emphasis>
With the character states of the
<taxonomicName id="9DBBECB12A56D850FD1C7DCD1C88B2FE" box="[693,907,1218,1243]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="terminata">
<emphasis id="68CF4B202A56D850FD1C7DCD1C88B2FE" box="[693,907,1218,1243]" italics="true" pageId="23" pageNumber="24">Calleida terminata</emphasis>
</taxonomicName>
species group, but distinct from the two other species of this group by the following character combination: medium sized (L:
<quantity id="9D433AD72A56D850FB907DE91BA0B2DA" box="[1081,1187,1254,1279]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="23" pageNumber="24" unit="mm" value="10.5">10.5 mm</quantity>
). Head and pronotum strongly metallic green or cupreous; elytra metallic green with wide cupreous metallic refection on apical and lateral regions; antennae, palpomeres, lateral furrows of pronotum, coxae and tarsi reddish; femora and underside strongly metallic green. Pronotum with deep transverse wrinkles. Elytra with outer apical angle acutely prominent apically. Median lobe of aedeagus in lateral aspect with ventral margin completely straight beyond the main ventral lobe; endophallus with one chitinized piece.
</paragraph>
<paragraph id="5A0497322A56D850FF6E7CB11C7AB03A" blockId="23.[151,1437,1218,2035]" pageId="23" pageNumber="24">
This species can be easily recognized among all Asiatic
<taxonomicName id="9DBBECB12A56D850FC917CB11C9BB3F2" authorityName="Latreille" authorityYear="1824" box="[824,920,1470,1495]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A56D850FC917CB11C9BB3F2" box="[824,920,1470,1495]" italics="true" pageId="23" pageNumber="24">Calleida</emphasis>
</taxonomicName>
species for the fully metallic green colour and the very shiny body on both the dorsal and the ventral side. For comparison with
<taxonomicName id="9DBBECB12A56D850FBF57CEC1A4DB3DE" authority="Casale &amp; Shi" authorityName="Casale &amp; Shi" box="[1116,1358,1506,1531]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="viet" status="sp. nov.">
<emphasis id="68CF4B202A56D850FBF57CEC1BABB3DE" box="[1116,1192,1507,1531]" italics="true" pageId="23" pageNumber="24">C. viet</emphasis>
Casale &amp; Shi
</taxonomicName>
<emphasis id="68CF4B202A56D850FAF07CED1A9FB3DE" bold="true" box="[1369,1436,1506,1531]" pageId="23" pageNumber="24">
<taxonomicNameLabel id="73FCF65B2A56D850FAF07CED1A9FB3DE" box="[1369,1436,1506,1531]" pageId="23" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
described below see diagnosis of
<taxonomicName id="9DBBECB12A56D850FDA77F081D54B03B" box="[526,599,1543,1566]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="viet" status="sp. nov.">
<emphasis id="68CF4B202A56D850FDA77F081D54B03B" box="[526,599,1543,1566]" italics="true" pageId="23" pageNumber="24">C. viet</emphasis>
</taxonomicName>
<emphasis id="68CF4B202A56D850FDF77F091D9DB03A" bold="true" box="[606,670,1542,1567]" pageId="23" pageNumber="24">
<taxonomicNameLabel id="73FCF65B2A56D850FDF77F091D9DB03A" box="[606,670,1542,1567]" pageId="23" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and key to species.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A56D85FFF6E7F251D30B51D" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="description">
<paragraph id="5A0497322A56D850FF6E7F251CFAB042" blockId="23.[151,1437,1218,2035]" pageId="23" pageNumber="24">
<emphasis id="68CF4B202A56D850FF6E7F251E5AB066" bold="true" box="[199,345,1578,1603]" pageId="23" pageNumber="24">Description.</emphasis>
<emphasis id="68CF4B202A56D850FEC87F251D1DB066" box="[353,542,1578,1603]" italics="true" pageId="23" pageNumber="24">General features</emphasis>
as in
<figureCitation id="C2808BB72A56D850FDCC7F251DB7B066" box="[613,692,1578,1603]" captionStart="FIGURES 4447" captionStartId="23.[151,263,1096,1118]" captionTargetBox="[179,1384,197,1065]" captionTargetId="figure@23.[151,1436,193,1074]" captionTargetPageId="23" captionText="FIGURES 4447. Calleida terminata Waterhouse. 44 habitus and labels of lectotype (male, Sarawak, Malaysia, NHML), scale bar = 2.0 mm; 45 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (lectotype), scale bar = 0.5 mm; 46 left and right parameres of aedeagus (lectotype), scale bar = 0.2 mm; 47 male sternum IX (lectotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302733/files/figure.png" pageId="23" pageNumber="24">fig. 44</figureCitation>
. Medium-large sized species: L: 10.5; TL: 10.0. (male
<typeStatus id="850029902A56D850FA837F251A92B066" box="[1322,1425,1578,1603]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
). Body relative wide, elytra sub-depressed, slightly widened at the apical third.
</paragraph>
<paragraph id="5A0497322A56D850FF6E7F7D1BE6B0D2" blockId="23.[151,1437,1218,2035]" pageId="23" pageNumber="24">
<emphasis id="68CF4B202A56D850FF6E7F7D1E15B0AE" box="[199,278,1650,1675]" italics="true" pageId="23" pageNumber="24">Colour</emphasis>
: head, pronotum, femora, tibiae, and ventral side bright metallic green; palpi, antennae, basal half of mandibles, lateral furrows of pronotum, coxae and trochanters reddish; first antennomere with weak metallic green reflection. Head with two distinct reddish spots on vertex. Elytra metallic green, with wide reddish purple reflection on the lateral and pre-apical regions; suture, lateral margin and epipleura metallic green.
</paragraph>
<paragraph id="5A0497322A56D850FF6E7E0B1B51B11A" blockId="23.[151,1437,1218,2035]" pageId="23" pageNumber="24">
<emphasis id="68CF4B202A56D850FF6E7E0B1EF6B13E" box="[199,501,1794,1819]" italics="true" pageId="23" pageNumber="24">Lustre and microsculpture</emphasis>
: dorsal side very shiny and polished. Microsculpture composed of isodiametric meshes, almost vanished on head, invisible on pronotum, distinct on elytral intervals.
</paragraph>
<paragraph id="5A0497322A56D850FF6E7E451EA7B18E" blockId="23.[151,1437,1218,2035]" pageId="23" pageNumber="24">
<emphasis id="68CF4B202A56D850FF6E7E451E07B146" box="[199,260,1866,1891]" italics="true" pageId="23" pageNumber="24">Head</emphasis>
wide, flat, smooth; genae markedly constricted to the neck; frontal foveae shallow, sub-striate in front, ending in a rounded depression on each side behind; eyes prominent, hemispherical; antennae reaching backwards the basal sixth of elytra.
</paragraph>
<paragraph id="5A0497322A56D85FFF6E7EB81A57B6DD" blockId="23.[151,1437,1218,2035]" lastBlockId="24.[151,1437,151,861]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<emphasis id="68CF4B202A56D850FF6E7EB81E35B1EB" box="[199,310,1975,1998]" italics="true" pageId="23" pageNumber="24">Pronotum</emphasis>
flat, cordate, wider than the head, barely wider than long (PW/PL: 1.08); posterior margin bisinuate, wider than anterior margin. Disc with deep transverse wrinkles. Anterior angles rounded, lateral margins reflexed, rounded anteriorly and sinuate posteriorly. Posterior angles reflexed and projecting laterally, sharply pointed but obtuse at apex; median furrow deep, shallower near apex; the deep basal foveae, starting from near the posterior angles, extend anteriorly on each side, becoming gradually shallower, and disappearing near anterior angles.
</paragraph>
<paragraph id="5A0497322A59D85FFF6E780B1A20B788" blockId="24.[151,1437,151,861]" pageId="24" pageNumber="25">
<emphasis id="68CF4B202A59D85FFF6E780B1E0EB738" box="[199,269,260,285]" italics="true" pageId="24" pageNumber="25">Elytra</emphasis>
: sub-depressed, wide (EL/EW: 1.67), shallowly and transversely depressed on each side behind base, slightly dilated near the apical third. Shoulders rather square, sides slightly compressed at basal third; apex obliquely truncate; outer apical angles acutely prominent apically, apex sharp. Striae fine and shallow, with fine punctures, very shallow near apex, stria l much deeper near apex; intervals slightly convex, flat on disc, interval 3 with two pores at one fourth and near apex, interval 8 dilated near apex into a smooth rounded callosity.
</paragraph>
<paragraph id="5A0497322A59D85FFF6E78B81DD4B7D0" blockId="24.[151,1437,151,861]" pageId="24" pageNumber="25">
<emphasis id="68CF4B202A59D85FFF6E78B81E3FB7F5" box="[199,316,439,464]" italics="true" pageId="24" pageNumber="25">Underside</emphasis>
: Abdominal sternum VII in male deeply notched at middle and with one seta on each side in males; straight and with two setae on each side in females.
</paragraph>
<paragraph id="5A0497322A59D85FFF6E7B0E1C38B418" blockId="24.[151,1437,151,861]" pageId="24" pageNumber="25">
<emphasis id="68CF4B202A59D85FFF6E7B0E1FFFB43D" box="[199,252,513,536]" italics="true" pageId="24" pageNumber="25">Legs</emphasis>
: Tarsi with the fourth tarsomere strongly bilobed, in male pro- and mesotarsi with the three basal tarsomeres clothed beneath with two rows of adhesive hairs.
</paragraph>
<paragraph id="5A0497322A59D85FFF6E7B481FF0B530" blockId="24.[151,1437,151,861]" pageId="24" pageNumber="25">
<emphasis id="68CF4B202A59D85FFF6E7B481E6FB445" box="[199,364,583,608]" italics="true" pageId="24" pageNumber="25">Male genitalia</emphasis>
: median lobe of aedeagus (
<figureCitation id="C2808BB72A59D85FFD0D7B481DECB445" box="[676,751,583,608]" captionStart="FIGURES 4447" captionStartId="23.[151,263,1096,1118]" captionTargetBox="[179,1384,197,1065]" captionTargetId="figure@23.[151,1436,193,1074]" captionTargetPageId="23" captionText="FIGURES 4447. Calleida terminata Waterhouse. 44 habitus and labels of lectotype (male, Sarawak, Malaysia, NHML), scale bar = 2.0 mm; 45 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (lectotype), scale bar = 0.5 mm; 46 left and right parameres of aedeagus (lectotype), scale bar = 0.2 mm; 47 male sternum IX (lectotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302733/files/figure.png" pageId="24" pageNumber="25">fig. 45</figureCitation>
) slightly bent; middle of right side strongly dilated in dorsal and ventral aspect, forming a short lobe, straight in lateral aspect; ventral margin with a very developed and prominent ventral lobe, approximately triangular shaped; ventral margin completely straight beyond the main ventral lobe; apical lamina flat and short, rounded at apex; endophallus with single short and narrow chitinized piece, curved at apex. Left paramere wide, depressed on the dorsal side; right paramere a little protruded at apex. (
<figureCitation id="C2808BB72A59D85FFF367BF31FE8B530" box="[159,235,764,789]" captionStart="FIGURES 4447" captionStartId="23.[151,263,1096,1118]" captionTargetBox="[179,1384,197,1065]" captionTargetId="figure@23.[151,1436,193,1074]" captionTargetPageId="23" captionText="FIGURES 4447. Calleida terminata Waterhouse. 44 habitus and labels of lectotype (male, Sarawak, Malaysia, NHML), scale bar = 2.0 mm; 45 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (lectotype), scale bar = 0.5 mm; 46 left and right parameres of aedeagus (lectotype), scale bar = 0.2 mm; 47 male sternum IX (lectotype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302733/files/figure.png" pageId="24" pageNumber="25">fig. 46</figureCitation>
)
</paragraph>
<paragraph id="5A0497322A59D85FFF6E7A101D30B51D" blockId="24.[151,1437,151,861]" box="[199,563,799,824]" pageId="24" pageNumber="25">
<emphasis id="68CF4B202A59D85FFF6E7A101E86B51D" box="[199,389,799,824]" italics="true" pageId="24" pageNumber="25">Female genitalia</emphasis>
: not examined.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A59D85FFF6E7A4B1A6AB579" box="[199,1385,836,861]" pageId="24" pageNumber="25" type="distribution">
<paragraph id="5A0497322A59D85FFF6E7A4B1A6AB579" blockId="24.[151,1437,151,861]" box="[199,1385,836,861]" pageId="24" pageNumber="25">
<emphasis id="68CF4B202A59D85FFF6E7A4B1D96B578" bold="true" box="[199,661,836,861]" pageId="24" pageNumber="25">Geographical distribution and habitat.</emphasis>
Only known from Borneo (
<collectingCountry id="22ACD7A22A59D85FFC657A4B1B33B578" box="[972,1072,836,861]" name="Malaysia" pageId="24" pageNumber="25">
<collectingRegion id="987F59D02A59D85FFC657A4B1B33B578" box="[972,1072,836,861]" country="Malaysia" name="Sarawak" pageId="24" pageNumber="25">Sarawak</collectingRegion>
</collectingCountry>
) in tropical forests (
<figureCitation id="C2808BB72A59D85FFABB7A4B1A58B579" box="[1298,1371,836,860]" captionStart="MAP 4" captionStartId="29.[151,209,1076,1098]" captionTargetBox="[151,1436,195,1054]" captionTargetId="figure@29.[151,1436,195,1054]" captionTargetPageId="29" captionText="MAP 4 Distributions for species in the C. terminata group: C. terminata Waterhouse (green); C. corporaali Andrewes (blue); C. viet sp. n. (red)." httpUri="https://zenodo.org/record/1302739/files/figure.png" pageId="24" pageNumber="25">map 4</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>

View file

@ -0,0 +1,512 @@
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<mods:title id="C1891948F36CEFF91CA658902B765FFB">Revision of the Oriental species of Calleida Latreille (sensu lato). Part 1: Introduction, groups of species, and species of six species groups (Coleoptera: Carabidae: Lebiini)</mods:title>
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<subSubSection id="12A1C4B92A59D85FFF3E7AAD1D7BB599" box="[151,632,930,956]" pageId="24" pageNumber="25" type="synonymic_list">
<paragraph id="5A0497322A59D85FFF3E7AAD1D7BB599" blockId="24.[151,632,930,956]" box="[151,632,930,956]" pageId="24" pageNumber="25">
<heading id="014C205E2A59D85FFF3E7AAD1D7BB599" bold="true" box="[151,632,930,956]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
<emphasis id="68CF4B202A59D85FFF3E7AAD1D7BB599" bold="true" box="[151,632,930,956]" pageId="24" pageNumber="25">
[7]
<taxonomicName id="9DBBECB12A59D85FFF697AAD1D7BB599" ID-CoL="5X5ZW" authority="Andrewes, 1929" authorityName="Andr. H. E. Andrewes" authorityYear="1929" box="[192,632,930,956]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="corporaali">
<emphasis id="68CF4B202A59D85FFF697AAD1EA9B599" bold="true" box="[192,426,930,956]" italics="true" pageId="24" pageNumber="25">Calleida corporaali</emphasis>
<bibRefCitation id="3E2AEAC32A59D85FFE187AAD1D7BB599" box="[433,632,930,956]" pageId="24" pageNumber="25" refId="ref28873" refString="Andrewes, H. E. (1929) Papers on Oriental Carabidae. XXII. The Annals and Magazine of Natural History, Series 10, 4 (22), 353 - 371. https: // doi. org / 10.1080 / 00222932908673067" type="journal article">Andrewes, 1929</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A59D85FFF3E7AE61A89B219" pageId="24" pageNumber="25" type="reference_group">
<paragraph id="5A0497322A59D85FFF3E7AE61BE1B5DA" blockId="24.[151,1436,1001,1085]" box="[151,1250,1001,1023]" pageId="24" pageNumber="25">
<taxonomicName id="9DBBECB12A59D85FFF3E7AE61D43B5DA" ID-CoL="5X5ZW" authority="Andrewes, 1929: 370" authorityName="Andr. H. E. Andrewes" authorityPageNumber="370" authorityYear="1929" box="[151,576,1001,1023]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="corporaali">
<emphasis id="68CF4B202A59D85FFF3E7AE61E62B5DA" box="[151,353,1001,1023]" italics="true" pageId="24" pageNumber="25">Calleida corporaali</emphasis>
<bibRefCitation id="3E2AEAC32A59D85FFECE7AE61D13B5DA" box="[359,528,1001,1023]" pageId="24" pageNumber="25" refId="ref28873" refString="Andrewes, H. E. (1929) Papers on Oriental Carabidae. XXII. The Annals and Magazine of Natural History, Series 10, 4 (22), 353 - 371. https: // doi. org / 10.1080 / 00222932908673067" type="journal article">Andrewes, 1929</bibRefCitation>
: 370
</taxonomicName>
(
<typeStatus id="850029902A59D85FFDE47AE61D82B5DA" box="[589,641,1001,1023]" pageId="24" pageNumber="25">Type</typeStatus>
locality:
<collectingRegion id="987F59D02A59D85FFD497AE61C34B5DA" box="[736,823,1001,1023]" country="Indonesia" name="Sumatera Utara" pageId="24" pageNumber="25">Sumatra</collectingRegion>
, Brastagi;
<typeStatus id="850029902A59D85FFC0A7AE61B03B5DA" box="[931,1024,1001,1023]" pageId="24" pageNumber="25" type="lectotype">lectotype</typeStatus>
deposited in NHML).
</paragraph>
<paragraph id="5A0497322A59D85FFF3E7D071A89B219" blockId="24.[151,1436,1001,1085]" pageId="24" pageNumber="25">
<taxonomicName id="9DBBECB12A59D85FFF3E7D071D39B23B" ID-CoL="8L4P8" authority="Louwerens, 1958: 254" authorityName="Louwerens" authorityPageNumber="254" authorityYear="1958" box="[151,570,1032,1054]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="lieftincki">
<emphasis id="68CF4B202A59D85FFF3E7D071E4DB23B" box="[151,334,1032,1054]" italics="true" pageId="24" pageNumber="25">Calleida lieftincki</emphasis>
<bibRefCitation id="3E2AEAC32A59D85FFEFC7D061D0AB23B" box="[341,521,1032,1054]" pageId="24" pageNumber="25" refId="ref30526" refString="Louwerens, C. J. (1958) On a collection of Carabidae from the Northern Moluccas (Coleoptera). Part 2. Treubia, 24, 131 - 262." type="journal article">Louwerens, 1958</bibRefCitation>
: 254
</taxonomicName>
(
<typeStatus id="850029902A59D85FFDE17D071D7FB23B" box="[584,636,1032,1054]" pageId="24" pageNumber="25">Type</typeStatus>
locality: Borneo, Pemantan River near Sampit;
<typeStatus id="850029902A59D85FFBC17D071BC3B23B" box="[1128,1216,1032,1054]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
deposited in NNML, labeled as
<taxonomicName id="9DBBECB12A59D85FFE9B7D291D5BB219" authority="Louwerens" authorityName="Louwerens" box="[306,600,1062,1085]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="wegneri">
<emphasis id="68CF4B202A59D85FFE9B7D291EE2B218" box="[306,481,1062,1085]" italics="true" pageId="24" pageNumber="25">Calleida wegneri</emphasis>
Louwerens
</taxonomicName>
(
<emphasis id="68CF4B202A59D85FFDCF7D271C0FB219" box="[614,780,1062,1085]" italics="true" pageId="24" pageNumber="25">nomen in litteris</emphasis>
);
<bibRefCitation id="3E2AEAC32A59D85FFC8B7D291C98B219" box="[802,923,1062,1084]" pageId="24" pageNumber="25" refId="ref30886" refString="Stork, N. (1986) An annotated checklist of the Carabidae (including Cicindelinae, Rhysodinae and Paussinae) recorded from Borneo. In: Occasional Papers on Systematic Entomology. Fol. 2. British Museum (Natural History), London, pp. 1 - 24." type="book chapter">Stork, 1986</bibRefCitation>
: 16 (misspelled as
<taxonomicName id="9DBBECB12A59D85FFBF27D281BD5B219" box="[1115,1238,1062,1084]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="lieftincka">
<emphasis id="68CF4B202A59D85FFBF27D281BD5B219" box="[1115,1238,1062,1084]" italics="true" pageId="24" pageNumber="25">C. lieftincka</emphasis>
</taxonomicName>
)
<emphasis id="68CF4B202A59D85FFB4C7D291A80B219" bold="true" box="[1253,1411,1062,1084]" pageId="24" pageNumber="25">new synonymy</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A59D85FFF3E7D661CD3B177" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="5A0497322A59D85FFF3E7D661C5AB37F" blockId="24.[151,1437,1129,2018]" pageId="24" pageNumber="25">
<emphasis id="68CF4B202A59D85FFF3E7D661DBEB2A7" bold="true" box="[151,701,1129,1154]" pageId="24" pageNumber="25">
<typeStatus id="850029902A59D85FFF3E7D661FD1B2A7" box="[151,210,1129,1154]" pageId="24" pageNumber="25">Type</typeStatus>
material examined:
<taxonomicName id="9DBBECB12A59D85FFE787D661DB7B2A7" authorityName="Andr. H. E. Andrewes" authorityYear="1929" box="[465,692,1129,1154]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="corporaali">
<emphasis id="68CF4B202A59D85FFE787D661DB7B2A7" bold="true" box="[465,692,1129,1154]" italics="true" pageId="24" pageNumber="25">Calleida corporaali</emphasis>
</taxonomicName>
:
</emphasis>
<materialsCitation id="EAD39D6F2A59D85FFD6E7D651BB4B2EF" collectingDate="21-02-13" collectionCode="B" collectorName="J. B. Corporaal" county="Sumatras O. K.," elevation="0" location="Brastagi" municipality="Brastagi" pageId="24" pageNumber="25" specimenCode="B.M. 1945-97" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<typeStatus id="850029902A59D85FFD6E7D651C39B2A7" box="[711,826,1130,1154]" pageId="24" pageNumber="25" type="lectotype">Lectotype</typeStatus>
, designated herein,
<specimenCount id="4CBD5CBB2A59D85FFB887D661B59B2A7" box="[1057,1114,1129,1154]" pageId="24" pageNumber="25" type="male">male</specimenCount>
, “
<collectorName id="F74EF2E42A59D85FFBDC7D651A1DB2A7" box="[1141,1310,1129,1154]" pageId="24" pageNumber="25">J.B. Corporaal</collectorName>
.
<collectingCounty id="B365EFBE2A59D85FFA857D651FD7B283" pageId="24" pageNumber="25">Sumatras O.K.,</collectingCounty>
<collectingMunicipality id="BA600D482A59D85FFF747D821E3DB283" box="[221,318,1165,1190]" pageId="24" pageNumber="25">Brastagi</collectingMunicipality>
.
<collectingDate id="3E41481A2A59D85FFEE07D811EA3B283" box="[329,416,1166,1190]" pageId="24" pageNumber="25" value="21-02-13">
<date id="2E05B1F22A59D85FFEE07D811EA3B283" box="[329,416,1166,1190]" pageId="24" pageNumber="25" value="21-02-13">13.2.21</date>
</collectingDate>
,
<elevation id="D19670012A59D85FFE047D811D0AB283" box="[429,521,1166,1190]" pageId="24" pageNumber="25">1300 M</elevation>
”, “Type”, “
<taxonomicName id="9DBBECB12A59D85FFD257D821BA1B283" authority="Corporaali Type Andr. H. E Andrewes" authorityName="Corporaali Type Andr. H.E Andrewes" box="[652,1186,1165,1190]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">Calleida Corporaali Type Andr. H.E Andrewes</taxonomicName>
det.” (handwritten by Andrewes), “H.E. Andrewes Coll.
<specimenCode id="0A1D3F492A59D85FFDB77DBD1DBCB2EF" box="[542,703,1201,1226]" pageId="24" pageNumber="25">B.M. 1945-97</specimenCode>
”, “Ex Coll. B.H. Klynstra” (NHML,
<figureCitation id="C2808BB72A59D85FFBC87DBE1BADB2EF" box="[1121,1198,1201,1226]" captionStart="FIGURES 4855" captionStartId="26.[151,264,1681,1703]" captionTargetBox="[156,1417,287,1659]" captionTargetId="figure@26.[151,1436,265,1660]" captionTargetPageId="26" captionText="FIGURES 4855. Calleida corporaali Andrewes. 48 habitus (female, Sabah, Malaysia), scale bar = 2.0 mm; 49 habitus and labels of lectotype for C. corporaali Andrewes (male, Sumatra, NHML), scale bar = 2.0 mm; 50 habitus and labels of lectotype for C. lieftincki Louwerens (female, Borneo, NNML), scale bar = 2.0 mm; 51 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (lectotype), scale bar = 0.5 mm; 52 left and right parameres of aedeagus (lectotype), scale bar = 0.2 mm; 53 male sternum IX (lectotype), scale bar = 0.5 mm; 54 ovipositor (Sabah, Malaysia), scale bar = 0.2 mm; 55 female reproductive tract (Sabah, Malaysia), scale bar = 0.5 mm; abbreviations: sp spermatheca, pd spermathecal pedicel, gd spermathecal gland duct, ga glandular area, at atrium, bpl basal sclerotized plate." httpUri="https://zenodo.org/record/1302735/files/figure.png" pageId="24" pageNumber="25">fig. 49</figureCitation>
)
</materialsCitation>
.
<typeStatus id="850029902A59D85FFB687DBE1A6AB2EF" box="[1217,1385,1201,1226]" pageId="24" pageNumber="25" type="paralectotype">Paralectotypes</typeStatus>
: one male, &quot;Malaya
<collectingRegion id="987F59D02A59D85FFEF57DDA1D0CB2C8" box="[348,527,1237,1262]" country="Malaysia" name="Kuala Lumpur" pageId="24" pageNumber="25">Kuala Lumpur</collectingRegion>
. Gombak valley,
<date id="2E05B1F22A59D85FFD507DD91CC5B2CB" box="[761,966,1237,1262]" pageId="24" pageNumber="25" value="1921-10-22">Oct. 22nd 1921</date>
&quot;, &quot;
<taxonomicName id="9DBBECB12A59D85FFC587DDA1E3FB337" authority="Corporaali Cotype Andr., H. E. Andrewes" authorityName="Corporaali Cotype Andr., H.E.Andrewes" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">Calleida Corporaali Cotype Andr., H.E.Andrewes</taxonomicName>
det.&quot;, &quot;15&quot;, &quot;Ex F.M.S. / Museum. / B.M. 1955-354.&quot; (
<collectionCode id="3CAA0FF72A59D85FFC437DF51B40B337" box="[1002,1091,1274,1298]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="24" pageNumber="25">NHML</collectionCode>
); one female, &quot;J.B.Corporaal Banddar. Baroe
<date id="2E05B1F22A59D85FFECB7C111EC9B313" box="[354,458,1310,1334]" pageId="24" pageNumber="25" value="1920-01-07">7.1.1920</date>
&quot;, &quot;Museum Leiden Ex coll. B.H.Klynstra&quot;, &quot;
<taxonomicName id="9DBBECB12A59D85FFC547C121E3FB37F" authority="Corporaali Cotyp. Andr., H. E. Andrewes" authorityName="Corporaali Cotyp. Andr., H.E.Andrewes" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">Calleida Corporaali Cotyp. Andr., H.E.Andrewes</taxonomicName>
det.&quot;, &quot;
<taxonomicName id="9DBBECB12A59D85FFE277C4E1D89B37F" authority="Andr. Corporaali" authorityName="Andr. Corporaali" box="[398,650,1345,1370]" class="Insecta" family="Cerambycidae" genus="Call" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">Call Andr. Corporaali</taxonomicName>
&quot;; &quot;
<typeStatus id="850029902A59D85FFD017C4D1DDEB37F" box="[680,733,1346,1370]" pageId="24" pageNumber="25">type</typeStatus>
&quot; (
<collectionCode id="3CAA0FF72A59D85FFD5D7C4D1C4EB37F" box="[756,845,1346,1370]" pageId="24" pageNumber="25">NNML</collectionCode>
).
</paragraph>
<paragraph id="5A0497322A59D85FFF6E7C6B1DA2B3E3" blockId="24.[151,1437,1129,2018]" pageId="24" pageNumber="25">
<taxonomicName id="9DBBECB12A59D85FFF6E7C6B1E97B358" authorityName="Louwerens" authorityYear="1958" box="[199,404,1380,1405]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="lieftincki">
<emphasis id="68CF4B202A59D85FFF6E7C6B1E97B358" bold="true" box="[199,404,1380,1405]" italics="true" pageId="24" pageNumber="25">Calleida lieftincki</emphasis>
</taxonomicName>
:
<typeStatus id="850029902A59D85FFE0A7C6A1D0DB35B" box="[419,526,1381,1406]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
, teneral female, here identified: &quot;S. Borneo,
<quantity id="9D433AD72A59D85FFC567C691B41B35B" box="[1023,1090,1382,1406]" metricMagnitude="2" metricUnit="m" metricValue="1.09" pageId="24" pageNumber="25" unit="m" value="109.0">109m</quantity>
, Pemantan-Sampit Riv. ult.
<date id="2E05B1F22A59D85FFAD37C6A1FD1B387" pageId="24" pageNumber="25" value="1953-07">Vii 1953</date>
, M.A.Lieftinck&quot;, &quot;Museum Leiden ex. Collection C.J. Louwerens rec. 1979&quot;, &quot;
<typeStatus id="850029902A59D85FFBEA7C851B89B387" box="[1091,1162,1418,1442]" pageId="24" pageNumber="25">Typus</typeStatus>
<taxonomicName id="9DBBECB12A59D85FFB387C861A9EB384" authority="Louw." authorityName="Louw." box="[1169,1437,1417,1442]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="wegneri">Calleida wegneri Louw.</taxonomicName>
det. C.J. Louwerens&quot;, &quot;
<typeStatus id="850029902A59D85FFE307CA11ECDB3E3" box="[409,462,1454,1478]" pageId="24" pageNumber="25">type</typeStatus>
&quot; (
<collectionCode id="3CAA0FF72A59D85FFE4C7CA11D3FB3E3" box="[485,572,1454,1478]" pageId="24" pageNumber="25">NNML</collectionCode>
,
<figureCitation id="C2808BB72A59D85FFDEE7CA21D97B3E3" box="[583,660,1453,1478]" captionStart="FIGURES 4855" captionStartId="26.[151,264,1681,1703]" captionTargetBox="[156,1417,287,1659]" captionTargetId="figure@26.[151,1436,265,1660]" captionTargetPageId="26" captionText="FIGURES 4855. Calleida corporaali Andrewes. 48 habitus (female, Sabah, Malaysia), scale bar = 2.0 mm; 49 habitus and labels of lectotype for C. corporaali Andrewes (male, Sumatra, NHML), scale bar = 2.0 mm; 50 habitus and labels of lectotype for C. lieftincki Louwerens (female, Borneo, NNML), scale bar = 2.0 mm; 51 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (lectotype), scale bar = 0.5 mm; 52 left and right parameres of aedeagus (lectotype), scale bar = 0.2 mm; 53 male sternum IX (lectotype), scale bar = 0.5 mm; 54 ovipositor (Sabah, Malaysia), scale bar = 0.2 mm; 55 female reproductive tract (Sabah, Malaysia), scale bar = 0.5 mm; abbreviations: sp spermatheca, pd spermathecal pedicel, gd spermathecal gland duct, ga glandular area, at atrium, bpl basal sclerotized plate." httpUri="https://zenodo.org/record/1302735/files/figure.png" pageId="24" pageNumber="25">fig. 50</figureCitation>
).
</paragraph>
<paragraph id="5A0497322A59D85FFF6E7CDE1CD3B177" blockId="24.[151,1437,1129,2018]" pageId="24" pageNumber="25">
<emphasis id="68CF4B202A59D85FFF6E7CDE1D27B3CF" bold="true" box="[199,548,1489,1514]" pageId="24" pageNumber="25">Non-type material examined</emphasis>
:
<emphasis id="68CF4B202A59D85FFD9E7CDE1D9CB3CF" bold="true" box="[567,671,1489,1514]" pageId="24" pageNumber="25">
<collectingRegion id="987F59D02A59D85FFD9E7CDE1D9CB3CF" box="[567,671,1489,1514]" country="Indonesia" name="Sumatera Utara" pageId="24" pageNumber="25">Sumatra</collectingRegion>
</emphasis>
: one female, “
<collectingRegion id="987F59D02A59D85FFCE67CDD1CACB3CF" box="[847,943,1490,1514]" country="Indonesia" name="Sumatera Utara" pageId="24" pageNumber="25">Sumatra</collectingRegion>
H. Kubale S.”, &quot;Museum fur Naturkunde Humboldt Univ.
<collectingRegion id="987F59D02A59D85FFEFF7CFA1E9FB02B" box="[342,412,1525,1550]" country="Germany" name="Berlin" pageId="24" pageNumber="25">Berlin</collectingRegion>
(
<collectionCode id="3CAA0FF72A59D85FFE057CF91D1FB02B" box="[428,540,1526,1550]" pageId="24" pageNumber="25">MNHUB</collectionCode>
)&quot;, &quot;
<taxonomicName id="9DBBECB12A59D85FFDED7CFA1C35B02B" baseAuthorityName="Andrewes" baseAuthorityYear="1929" box="[580,822,1525,1550]" class="Insecta" family="Carabidae" genus="Callidiola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="corporaali">
<emphasis id="68CF4B202A59D85FFDED7CFA1C35B02B" box="[580,822,1525,1550]" italics="true" pageId="24" pageNumber="25">Callidiola corporaali</emphasis>
</taxonomicName>
det. A. Casale&quot; (identified as
<taxonomicName id="9DBBECB12A59D85FFB257CF91A19B028" box="[1164,1306,1526,1549]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="terminata">
<emphasis id="68CF4B202A59D85FFB257CF91A19B028" box="[1164,1306,1526,1549]" italics="true" pageId="24" pageNumber="25">C. terminata</emphasis>
</taxonomicName>
) (
<collectionCode id="3CAA0FF72A59D85FFA9D7CF91A93B02B" box="[1332,1424,1526,1550]" pageId="24" pageNumber="25">MNHU</collectionCode>
); one male, “Paggar Alam,
<collectingRegion id="987F59D02A59D85FFE697F151D23B017" box="[448,544,1562,1586]" country="Indonesia" name="Sumatera Utara" pageId="24" pageNumber="25">Sumatra</collectingRegion>
“ (
<collectionCode id="3CAA0FF72A59D85FFD927F151D78B017" box="[571,635,1562,1586]" country="Germany" httpUri="http://grbio.org/cool/mqei-3i36" name="Culture Collection of Algae at the University of Marburg" pageId="24" pageNumber="25">CCA</collectionCode>
);
<emphasis id="68CF4B202A59D85FFD397F161DE4B017" bold="true" box="[656,743,1561,1586]" pageId="24" pageNumber="25">Borneo</emphasis>
: one female, &quot;Borneo:
<collectingRegion id="987F59D02A59D85FFC5C7F161B3FB017" box="[1013,1084,1561,1586]" country="Malaysia" name="Sabah" pageId="24" pageNumber="25">Sabah</collectingRegion>
, Keningau district, Jungle Girl Camp,
<geoCoordinate id="3F8FF1F52A59D85FFF417F321E4FB073" box="[232,332,1597,1622]" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="5" value="5.443">N5.4430</geoCoordinate>
,
<geoCoordinate id="3F8FF1F52A59D85FFEF17F321ED7B073" box="[344,468,1597,1622]" direction="east" orientation="longitude" pageId="24" pageNumber="25" precision="5" value="116.4512">E116.4512</geoCoordinate>
;
<quantity id="9D433AD72A59D85FFE497F311D32B073" box="[480,561,1598,1622]" metricMagnitude="3" metricUnit="m" metricValue="1.182" pageId="24" pageNumber="25" unit="m" value="1182.0">1182m</quantity>
; Shi H.L. &amp; Liu Y. lgt., light trap, Ins. Zoo.,
<collectionCode id="3CAA0FF72A59D85FFB827F311B66B073" box="[1067,1125,1598,1622]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="24" pageNumber="25">CAS</collectionCode>
. 2016.
<date id="2E05B1F22A59D85FFB067F321BEDB073" box="[1199,1262,1597,1622]" pageId="24" pageNumber="25" value="1930-04">IV.30</date>
N&quot; (
<collectionCode id="3CAA0FF72A59D85FFAB57F311A63B073" box="[1308,1376,1598,1622]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="24" pageNumber="25">IZAS</collectionCode>
); one female, same data but date 2016.
<date id="2E05B1F22A59D85FFDBB7F6E1D37B05F" box="[530,564,1633,1658]" pageId="24" pageNumber="25">V.1</date>
(
<collectionCode id="3CAA0FF72A59D85FFDEC7F6D1D8AB05F" box="[581,649,1634,1658]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="24" pageNumber="25">IZAS</collectionCode>
); one male, “
<collectingCountry id="22ACD7A22A59D85FFC897F6E1C84B05F" box="[800,903,1633,1658]" name="Malaysia" pageId="24" pageNumber="25">
<collectingRegion id="987F59D02A59D85FFC897F6E1C84B05F" box="[800,903,1633,1658]" country="Malaysia" name="Sarawak" pageId="24" pageNumber="25">Sarawak</collectingRegion>
</collectingCountry>
: Bidi
<date id="2E05B1F22A59D85FFC657F6E1B46B05F" box="[972,1093,1633,1658]" pageId="24" pageNumber="25" value="1909-03-23">23.iii.1909</date>
C.J. Brooks. B.M. 1936-681” (
<collectionCode id="3CAA0FF72A59D85FFF367F891FFBB0BB" box="[159,248,1670,1694]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="24" pageNumber="25">NHML</collectionCode>
); one female, &quot;M398&quot;, &quot;Oxford Univ. Exp. B.M. Hobby &amp; A.W. Moore. B.M. 1933-254&quot;, &quot;
<taxonomicName id="9DBBECB12A59D85FFB577F8A1E9DB0E7" authority="Andr. Det. G. E. Bryant" authorityName="Andr. Det. G.E. Bryant" class="Insecta" family="Carabidae" genus="Callida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="tenuis">Callida tenuis Andr. Det. G.E. Bryant</taxonomicName>
&quot;, &quot;
<collectingCountry id="22ACD7A22A59D85FFE157FA61D20B0E7" box="[444,547,1705,1730]" name="Malaysia" pageId="24" pageNumber="25">
<collectingRegion id="987F59D02A59D85FFE157FA61D20B0E7" box="[444,547,1705,1730]" country="Malaysia" name="Sarawak" pageId="24" pageNumber="25">Sarawak</collectingRegion>
</collectingCountry>
: R. Kapah trib. of R. Tinjar.
<date id="2E05B1F22A59D85FFCC17FA51CD3B0E7" box="[872,976,1706,1730]" pageId="24" pageNumber="25" value="1932-10-03">3.x.1932</date>
.&quot;, &quot;dear tree&quot;, &quot;
<taxonomicName id="9DBBECB12A59D85FFBD27FA61E49B0C3" authority="Andr. H. E. Andrewes" authorityName="Andr. H. E. Andrewes" authorityYear="1929" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="corporaali">Calleida corporaali Andr. H.E. Andrewes</taxonomicName>
det.&quot; (
<collectionCode id="3CAA0FF72A59D85FFE0B7FC11EF8B0C3" box="[418,507,1742,1766]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="24" pageNumber="25">NHML</collectionCode>
); one male, &quot;H.C.Siebers M.O. Borneo Exp. Long Petak
<specimenCount id="4CBD5CBB2A59D85FFB1E7FC21A01B0C3" box="[1207,1282,1741,1766]" pageId="24" pageNumber="25" type="generic">450M.</specimenCount>
IX-X.1925&quot;; &quot;Museum Leiden ex. collection C.J. Louwerens rec.1979&quot;; &quot;
<taxonomicName id="9DBBECB12A59D85FFCCE7FFE1A59B12F" authority="Corporaali Andr. H. E. Andrewes" authorityName="Corporaali Andr. H.E. Andrewes" box="[871,1370,1777,1802]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">Calleida Corporaali Andr. H.E. Andrewes</taxonomicName>
det.&quot; (
<collectionCode id="3CAA0FF72A59D85FFF367E191FFBB10B" box="[159,248,1814,1838]" pageId="24" pageNumber="25">NNML</collectionCode>
);
<emphasis id="68CF4B202A59D85FFEA27E1A1ED2B10B" bold="true" box="[267,465,1813,1838]" pageId="24" pageNumber="25">Malay peninsula</emphasis>
: one female “
<collectingCountry id="22ACD7A22A59D85FFDC57E1A1DDAB10B" box="[620,729,1813,1838]" name="Malaysia" pageId="24" pageNumber="25">Malaysia</collectingCountry>
:
<collectingRegion id="987F59D02A59D85FFD4D7E1A1C34B10B" box="[740,823,1813,1838]" country="Malaysia" name="Pahang" pageId="24" pageNumber="25">Pahang</collectingRegion>
distr.,
<quantity id="9D433AD72A59D85FFC2B7E191CCAB10B" box="[898,969,1813,1838]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="24" pageNumber="25" unit="km" value="30.0">30 km</quantity>
NE Raub, Lata Lembik, 3°56N 101°38E,
<quantity id="9D433AD72A59D85FFF3E7E351E10B174" box="[151,275,1850,1874]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="24" pageNumber="25" unit="m" value="300.0" valueMax="400.0" valueMin="200.0">200-400 m</quantity>
,
<date id="2E05B1F22A59D85FFEB67E351E8CB177" box="[287,399,1850,1874]" pageId="24" pageNumber="25" value="2002-04" valueMax="2002-05-01" valueMin="2002-04-22">22.IV-1.V</date>
&amp;
<date id="2E05B1F22A59D85FFE1A7E361D3EB177" box="[435,573,1849,1874]" pageId="24" pageNumber="25" value="2002-05-08" valueMax="2002-05-15" valueMin="2002-05-08">8-15.V.2002</date>
E. Jendek &amp; O. Šauša leg.” (
<collectionCode id="3CAA0FF72A59D85FFC2D7E351CC0B177" box="[900,963,1850,1874]" country="Germany" httpUri="http://grbio.org/cool/mqei-3i36" name="Culture Collection of Algae at the University of Marburg" pageId="24" pageNumber="25">CCA</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A59D85EFF6E7E521DC0B485" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="discussion">
<paragraph id="5A0497322A59D85FFF6E7E521E3EB1C7" blockId="24.[151,1437,1129,2018]" pageId="24" pageNumber="25">
<emphasis id="68CF4B202A59D85FFF6E7E521EE7B153" bold="true" box="[199,484,1885,1910]" pageId="24" pageNumber="25">
Notes on
<typeStatus id="850029902A59D85FFE937E521E6DB153" box="[314,366,1885,1910]" pageId="24" pageNumber="25">type</typeStatus>
material.
</emphasis>
For most species described by Andrewes, the name-baring the
<typeStatus id="850029902A59D85FFB6A7E511BF0B153" box="[1219,1267,1886,1910]" pageId="24" pageNumber="25">type</typeStatus>
specimen was originally fixed, although cited as &quot;
<typeStatus id="850029902A59D85FFD8C7E8D1D59B1BF" box="[549,602,1922,1946]" pageId="24" pageNumber="25">type</typeStatus>
&quot; (=
<typeStatus id="850029902A59D85FFD247E8E1DF2B1BF" box="[653,753,1921,1946]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
) and cotype (=
<typeStatus id="850029902A59D85FFC337E8D1B02B1BF" box="[922,1025,1922,1946]" pageId="24" pageNumber="25" type="paratype">paratype</typeStatus>
). But, for
<taxonomicName id="9DBBECB12A59D85FFBDC7E8D1A0CB1BF" authorityName="Andr. H. E. Andrewes" authorityYear="1929" box="[1141,1295,1921,1946]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="corporaali">
<emphasis id="68CF4B202A59D85FFBDC7E8D1A0CB1BF" box="[1141,1295,1921,1946]" italics="true" pageId="24" pageNumber="25">C. corporaali</emphasis>
</taxonomicName>
, there is not any original designation. We here designate the male specimen labeled as &quot;
<typeStatus id="850029902A59D85FFC747EA91B11B19B" box="[989,1042,1958,1982]" pageId="24" pageNumber="25">type</typeStatus>
&quot; by Andrewes himself as
<typeStatus id="850029902A59D85FFA9C7EAA1A9FB19B" box="[1333,1436,1957,1982]" pageId="24" pageNumber="25" type="lectotype">lectotype</typeStatus>
of this species.
</paragraph>
<paragraph id="5A0497322A58D85EFF6E79981DC0B485" blockId="25.[151,1437,151,2021]" pageId="25" pageNumber="26">
<taxonomicName id="9DBBECB12A58D85EFF6E79981D10B68A" authority="Louwerens" authorityName="Louwerens" authorityYear="1958" box="[199,531,151,176]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="lieftincki">
<emphasis id="68CF4B202A58D85EFF6E79981E8CB695" box="[199,399,151,176]" italics="true" pageId="25" pageNumber="26">Calleida lieftincki</emphasis>
Louwerens
</taxonomicName>
has been so far a mysterious and questionable taxon for us. The character state of multisetose abdominal sternum VII (“
<emphasis id="68CF4B202A58D85EFDED79B41B26B6F6" box="[580,1061,187,212]" italics="true" pageId="25" pageNumber="26">last male ventral segment 4-setose at apex</emphasis>
”), as stressed by the author, lead us previously including this taxon in the
<taxonomicName id="9DBBECB12A58D85EFDC379D01C05B6D2" authorityName="Heller in Mindanao" authorityYear="1921" box="[618,774,222,247]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="discoidalis">
<emphasis id="68CF4B202A58D85EFDC379D01C05B6D2" box="[618,774,222,247]" italics="true" pageId="25" pageNumber="26">C. discoidalis</emphasis>
</taxonomicName>
species group as here defined. The
<typeStatus id="850029902A58D85EFB0B79EF1BD1B6DD" box="[1186,1234,224,248]" pageId="25" pageNumber="26">type</typeStatus>
material with the original name has been so far impossible to find. One of the authors of the present contribution (A.C.), despite the help of colleagues of the MNHN and the NNML, was unable to find it both in the NNML and the Bogor Museum (
<collectingCountry id="22ACD7A22A58D85EFF3778461E0CB747" box="[158,271,329,354]" name="Indonesia" pageId="25" pageNumber="26">Indonesia</collectingCountry>
,
<collectingRegion id="987F59D02A58D85EFEB778461E57B744" box="[286,340,329,353]" country="Indonesia" name="Jawa Timur" pageId="25" pageNumber="26">Java</collectingRegion>
), where part of the Louwerens entomological material is preserved. Finally, during his stay in Europe in 2010, the junior author (H.S.) found in the NNML a female specimen labeled by Louwerens as “
<taxonomicName id="9DBBECB12A58D85EFA9578631FF7B78C" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="wegneri">Calleida wegneri</taxonomicName>
” (
<emphasis id="68CF4B202A58D85EFEBF789E1EDEB78C" box="[278,477,400,425]" italics="true" pageId="25" pageNumber="26">nomen in litteris</emphasis>
never published), coming from the
<typeStatus id="850029902A58D85EFC36789E1CCCB78C" box="[927,975,401,425]" pageId="25" pageNumber="26">type</typeStatus>
locality of
<taxonomicName id="9DBBECB12A58D85EFBCE789E1BF1B78C" authorityName="Louwerens" authorityYear="1958" box="[1127,1266,400,425]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="lieftincki">
<emphasis id="68CF4B202A58D85EFBCE789E1BF1B78C" box="[1127,1266,400,425]" italics="true" pageId="25" pageNumber="26">C. lieftincki</emphasis>
</taxonomicName>
and perfectly corresponding to the original description of this species provided by the author. Therefore, this female specimen has to be considered as the
<typeStatus id="850029902A58D85EFE7B78D81D37B7D5" box="[466,564,471,496]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
of this taxon. Its examination showed that the abdominal sternum VII is not “multisetose”, but bears two setae in females on each side. This specimen (
<figureCitation id="C2808BB72A58D85EFB8F78F51B74B436" box="[1062,1143,506,531]" captionStart="FIGURES 4855" captionStartId="26.[151,264,1681,1703]" captionTargetBox="[156,1417,287,1659]" captionTargetId="figure@26.[151,1436,265,1660]" captionTargetPageId="26" captionText="FIGURES 4855. Calleida corporaali Andrewes. 48 habitus (female, Sabah, Malaysia), scale bar = 2.0 mm; 49 habitus and labels of lectotype for C. corporaali Andrewes (male, Sumatra, NHML), scale bar = 2.0 mm; 50 habitus and labels of lectotype for C. lieftincki Louwerens (female, Borneo, NNML), scale bar = 2.0 mm; 51 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (lectotype), scale bar = 0.5 mm; 52 left and right parameres of aedeagus (lectotype), scale bar = 0.2 mm; 53 male sternum IX (lectotype), scale bar = 0.5 mm; 54 ovipositor (Sabah, Malaysia), scale bar = 0.2 mm; 55 female reproductive tract (Sabah, Malaysia), scale bar = 0.5 mm; abbreviations: sp spermatheca, pd spermathecal pedicel, gd spermathecal gland duct, ga glandular area, at atrium, bpl basal sclerotized plate." httpUri="https://zenodo.org/record/1302735/files/figure.png" pageId="25" pageNumber="26">fig. 50</figureCitation>
) is quite teneral, but its morphological features correspond to those of the other specimen cited above (see: non-type material examined). Furthermore, we examined a male specimen of
<taxonomicName id="9DBBECB12A58D85EFD057B4D1C46B47F" authorityName="Andr. H. E. Andrewes" authorityYear="1929" box="[684,837,577,602]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="corporaali">
<emphasis id="68CF4B202A58D85EFD057B4D1C46B47F" box="[684,837,577,602]" italics="true" pageId="25" pageNumber="26">C. corporaali</emphasis>
</taxonomicName>
from Borneo (Long Petak) in NNML, and confirmed that its male genitalia are identical to those of the
<typeStatus id="850029902A58D85EFD607B6A1C28B45B" box="[713,811,613,638]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
from
<collectingRegion id="987F59D02A58D85EFCD87B6A1CD1B458" box="[881,978,613,637]" country="Indonesia" name="Sumatera Utara" pageId="25" pageNumber="26">Sumatra</collectingRegion>
. Based on these facts, we concluded that
<taxonomicName id="9DBBECB12A58D85EFF3E7B861E1AB484" authorityName="Louwerens" authorityYear="1958" box="[151,281,648,673]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="lieftincki">
<emphasis id="68CF4B202A58D85EFF3E7B861E1AB484" box="[151,281,648,673]" italics="true" pageId="25" pageNumber="26">C. lieftincki</emphasis>
</taxonomicName>
is a junior synonym of
<taxonomicName id="9DBBECB12A58D85EFD8A7B861DBFB484" authorityName="Andr. H. E. Andrewes" authorityYear="1929" box="[547,700,648,673]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="corporaali">
<emphasis id="68CF4B202A58D85EFD8A7B861DBFB484" box="[547,700,648,673]" italics="true" pageId="25" pageNumber="26">C. corporaali</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A58D85EFF6E7BA31E61B202" pageId="25" pageNumber="26" type="diagnosis">
<paragraph id="5A0497322A58D85EFF6E7BA31B00B5BC" blockId="25.[151,1437,151,2021]" pageId="25" pageNumber="26">
<emphasis id="68CF4B202A58D85EFF6E7BA31E42B4E0" bold="true" box="[199,321,684,709]" pageId="25" pageNumber="26">Diagnosis.</emphasis>
With the character states of the
<taxonomicName id="9DBBECB12A58D85EFD1C7BA31C88B4E1" box="[693,907,684,709]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="terminata">
<emphasis id="68CF4B202A58D85EFD1C7BA31C88B4E1" box="[693,907,684,709]" italics="true" pageId="25" pageNumber="26">Calleida terminata</emphasis>
</taxonomicName>
species group, but distinct from the two other species of this group by the following character combination: medium-large sized (L: 9.512.0 mm); head and pronotum dark brownish, with faint metallic green lustre on pronotal disc; elytra metallic green with narrow cupreous metallic reflection on apical and lateral regions; femora and underside with weak metallic green reflection. Pronotal disc without or with very shallow transverse wrinkles. Outer apical angles of elytra sharp, but not or only slightly prominent apically. Median lobe of aedeagus in lateral aspect with large ventral lobe, bisinuate on the dorsal and ventral margins; endophallus with one only chitinized piece.
</paragraph>
<paragraph id="5A0497322A58D85EFF6E7AAB1E61B202" blockId="25.[151,1437,151,2021]" pageId="25" pageNumber="26">
This species is similar to
<taxonomicName id="9DBBECB12A58D85EFE497AAA1DF8B598" authority="Waterhouse" authorityName="Waterhouse" box="[480,763,932,957]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="terminata">
<emphasis id="68CF4B202A58D85EFE497AAA1EF0B599" box="[480,499,933,956]" italics="true" pageId="25" pageNumber="26">C</emphasis>
.
<emphasis id="68CF4B202A58D85EFDA97AAA1D6EB599" box="[512,621,933,956]" italics="true" pageId="25" pageNumber="26">terminata</emphasis>
Waterhouse
</taxonomicName>
, but differs in: head, pronotum and underside only with very weak metallic lustre; pronotum longer, without deep wrinkles; elytra with reddish purple reflection restricted to a narrow area at apex; outer apical angles of elytra not or only slightly prominent distally; median lobe of aedeagus different in shape.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A58D85CFF6E7D3D1E09B43D" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26" type="description">
<paragraph id="5A0497322A58D85EFF6E7D3D1D9EB248" blockId="25.[151,1437,151,2021]" pageId="25" pageNumber="26">
<emphasis id="68CF4B202A58D85EFF6E7D3D1E5AB26E" bold="true" box="[199,345,1074,1099]" pageId="25" pageNumber="26">Description.</emphasis>
The original description provided by
<bibRefCitation id="3E2AEAC32A58D85EFCBE7D3D1CE3B26F" box="[791,992,1074,1099]" pageId="25" pageNumber="26" refId="ref28873" refString="Andrewes, H. E. (1929) Papers on Oriental Carabidae. XXII. The Annals and Magazine of Natural History, Series 10, 4 (22), 353 - 371. https: // doi. org / 10.1080 / 00222932908673067" type="journal article">Andrewes (1929)</bibRefCitation>
, here completed with some additional characters, is enough to distinguish this taxon.
</paragraph>
<paragraph id="5A0497322A58D85EFF6E7D761DB1B290" blockId="25.[151,1437,151,2021]" pageId="25" pageNumber="26">
<emphasis id="68CF4B202A58D85EFF6E7D761E80B2B4" box="[199,387,1145,1170]" italics="true" pageId="25" pageNumber="26">General features</emphasis>
: Medium-large sized species: L: 9.512.0 mm; TL: 9.0
<quantity id="9D433AD72A58D85EFBAF7D761B6FB2B4" box="[1030,1132,1145,1169]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="25" pageNumber="26" unit="mm" value="11.3">11.3 mm</quantity>
. Body relative wide, elytra sub-depressed, slightly expanded at apical third.
</paragraph>
<paragraph id="5A0497322A58D85EFF6E7DB01B3EB343" blockId="25.[151,1437,151,2021]" pageId="25" pageNumber="26">
<emphasis id="68CF4B202A58D85EFF6E7DB01E15B2FD" box="[199,278,1215,1240]" italics="true" pageId="25" pageNumber="26">Colour</emphasis>
: Head, pronotum and underside dark brown to piceous, with indistinct metallic green reflection. Apex of palpi and lateral margins of pronotum testaceous; antennae reddish, darkened from antennomere 4; trochanters brown; femora and tibiae piceous, with slightly distinct metallic green reflection. Head with two reddish spots, not always divided, on vertex. Elytra metallic green, bright in front and along sides and near apex, but dull on apical two-thirds of disk, with some reddish purple reflections at shoulders and near apex.
</paragraph>
<paragraph id="5A0497322A58D85EFF6E7C7D1DC2B388" blockId="25.[151,1437,151,2021]" pageId="25" pageNumber="26">
<emphasis id="68CF4B202A58D85EFF6E7C7D1EE8B3AF" box="[199,491,1393,1418]" italics="true" pageId="25" pageNumber="26">Lustre and microsculpture</emphasis>
: meshes of the microsculpture isodiametric on elytra and head, faint on the latter; on the pronotum they are wide, but almost vanished.
</paragraph>
<paragraph id="5A0497322A58D85EFF6E7CB71EA7B03D" blockId="25.[151,1437,151,2021]" pageId="25" pageNumber="26">
<emphasis id="68CF4B202A58D85EFF6E7CB71E07B3F4" box="[199,260,1464,1489]" italics="true" pageId="25" pageNumber="26">Head</emphasis>
wide, flat, smooth; genae markedly constricted to the neck; frontal foveae shallow, sub-striate in front, ending in a rounded depression on each side behind; eyes prominent, hemispherical; antennae reaching backwards the basal sixth of elytra.
</paragraph>
<paragraph id="5A0497322A58D85EFF6E7F2B1E47B135" blockId="25.[151,1437,151,2021]" pageId="25" pageNumber="26">
<emphasis id="68CF4B202A58D85EFF6E7F2B1E35B01E" box="[199,310,1572,1595]" italics="true" pageId="25" pageNumber="26">Pronotum</emphasis>
flat, cordate, wider than head, barely wider than long (PW/PL: 1.031.11); posterior margin bisinuate, wider than the anterior margin. Disc without or with very faint transverse wrinkles. Anterior angles rounded, lateral margins reflexed, rounded anteriorly and sinuate posteriorly. Posterior angles reflexed and projecting laterally, sharply pointed but apex slightly obtuse; median furrow deep, shallower near apex; front transverse impressions very shallow, posterior one deep at sides and joining the deep basal foveae which, starting from near the posterior angles, run forwards on each side, becoming gradually shallower, and disappearing near anterior angles.
</paragraph>
<paragraph id="5A0497322A58D85EFF6E7E141C22B1C0" blockId="25.[151,1437,151,2021]" pageId="25" pageNumber="26">
<emphasis id="68CF4B202A58D85EFF6E7E141E0EB111" box="[199,269,1819,1844]" italics="true" pageId="25" pageNumber="26">Elytra</emphasis>
: sub-depressed, wide (EL/EW: 1.581.69), transversely depressed on each side behind base, faintly dilated behind the apical third. Shoulders rather square, sides slightly compressed at basal third; apex obliquely truncate; the outer apical angles strongly thickened, on each side forming a sharp tooth, not or only slightly prominent apically. Striae fine, with distinct punctures, very shallow on disk behind, first stria much deeper near apex; intervals slightly convex, flat on disc, third interval with two pores at one fourth and near apex, eighth interval dilated near apex into a smooth rounded callosity.
</paragraph>
<paragraph id="5A0497322A5BD85DFF6E79981E80B6F0" blockId="26.[151,1436,151,213]" pageId="26" pageNumber="27">
<emphasis id="68CF4B202A5BD85DFF6E79981E3FB695" box="[199,316,151,176]" italics="true" pageId="26" pageNumber="27">Underside</emphasis>
: Abdominal sternum VII in male deeply notched at middle and with one seta on each side in male; two setae in females.
</paragraph>
<caption id="0EC4C7BA2A5BD85DFF3E7F9E1CD3B17A" httpUri="https://zenodo.org/record/1302735/files/figure.png" pageId="26" pageNumber="27" startId="26.[151,264,1681,1703]" targetBox="[156,1417,287,1659]" targetPageId="26">
<paragraph id="5A0497322A5BD85DFF3E7F9E1CD3B17A" blockId="26.[151,1436,1681,1887]" pageId="26" pageNumber="27">
<emphasis id="68CF4B202A5BD85DFF3E7F9E1E56B082" bold="true" box="[151,341,1681,1703]" pageId="26" pageNumber="27">FIGURES 4855.</emphasis>
<taxonomicName id="9DBBECB12A5BD85DFEF57F9E1D9EB082" authority="Andrewes." authorityName="Andr. H. E. Andrewes" authorityYear="1929" box="[348,669,1681,1703]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="corporaali">
<emphasis id="68CF4B202A5BD85DFEF57F9E1D24B082" bold="true" box="[348,551,1681,1703]" italics="true" pageId="26" pageNumber="27">Calleida corporaali</emphasis>
Andrewes.
</taxonomicName>
<emphasis id="68CF4B202A5BD85DFD0D7F9E1DBEB082" bold="true" box="[676,701,1681,1703]" pageId="26" pageNumber="27">48</emphasis>
habitus (female, Sabah, Malaysia), scale bar = 2.0 mm;
<emphasis id="68CF4B202A5BD85DFAAC7F9E1A1CB082" bold="true" box="[1285,1311,1681,1703]" pageId="26" pageNumber="27">49</emphasis>
habitus and labels of lectotype for
<taxonomicName id="9DBBECB12A5BD85DFED37FBE1D6CB0E3" authority="Andrewes" authorityName="Andr. H. E. Andrewes" authorityYear="1929" box="[378,623,1712,1734]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="corporaali">
<emphasis id="68CF4B202A5BD85DFED37FBE1D07B0E3" box="[378,516,1712,1734]" italics="true" pageId="26" pageNumber="27">C. corporaali</emphasis>
Andrewes
</taxonomicName>
(male, Sumatra, NHML), scale bar = 2.0 mm;
<emphasis id="68CF4B202A5BD85DFBE37FBF1B60B0E3" bold="true" box="[1098,1123,1712,1734]" pageId="26" pageNumber="27">50</emphasis>
habitus and labels of lectotype for
<taxonomicName id="9DBBECB12A5BD85DFF157FC01EA8B0C1" authority="Louwerens" authorityName="Louwerens" authorityYear="1958" box="[188,427,1742,1764]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="lieftincki">
<emphasis id="68CF4B202A5BD85DFF157FC01E37B0C1" box="[188,308,1742,1764]" italics="true" pageId="26" pageNumber="27">C. lieftincki</emphasis>
Louwerens
</taxonomicName>
(female, Borneo, NNML), scale bar = 2.0 mm;
<emphasis id="68CF4B202A5BD85DFC357FC11CB6B0C1" bold="true" box="[924,949,1742,1764]" pageId="26" pageNumber="27">51</emphasis>
median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (lectotype), scale bar = 0.5 mm;
<emphasis id="68CF4B202A5BD85DFCBB7FE21C2FB126" bold="true" box="[786,812,1773,1795]" pageId="26" pageNumber="27">52</emphasis>
left and right parameres of aedeagus (lectotype), scale bar = 0.2 mm;
<emphasis id="68CF4B202A5BD85DFF5A7E031E0EB107" bold="true" box="[243,269,1804,1826]" pageId="26" pageNumber="27">53</emphasis>
male sternum IX (lectotype), scale bar = 0.5 mm;
<emphasis id="68CF4B202A5BD85DFCB87E031C29B107" bold="true" box="[785,810,1804,1826]" pageId="26" pageNumber="27">54</emphasis>
ovipositor (Sabah, Malaysia), scale bar = 0.2 mm;
<emphasis id="68CF4B202A5BD85DFA9E7E031A52B107" bold="true" box="[1335,1361,1804,1826]" pageId="26" pageNumber="27">55</emphasis>
female reproductive tract (Sabah, Malaysia), scale bar = 0.5 mm; abbreviations:
<emphasis id="68CF4B202A5BD85DFC1F7E251CCDB165" bold="true" box="[950,974,1834,1856]" pageId="26" pageNumber="27">sp</emphasis>
spermatheca,
<emphasis id="68CF4B202A5BD85DFBC27E251B84B165" bold="true" box="[1131,1159,1834,1856]" pageId="26" pageNumber="27">pd</emphasis>
spermathecal pedicel,
<emphasis id="68CF4B202A5BD85DFA287E251A98B165" bold="true" box="[1409,1435,1834,1856]" pageId="26" pageNumber="27">gd</emphasis>
spermathecal gland duct,
<emphasis id="68CF4B202A5BD85DFE307E461EB1B17A" bold="true" box="[409,434,1865,1887]" pageId="26" pageNumber="27">ga</emphasis>
glandular area,
<emphasis id="68CF4B202A5BD85DFDFF7E461D68B17A" bold="true" box="[598,619,1865,1887]" pageId="26" pageNumber="27">at</emphasis>
atrium,
<emphasis id="68CF4B202A5BD85DFD697E461DE0B17A" bold="true" box="[704,739,1865,1887]" pageId="26" pageNumber="27">bpl</emphasis>
basal sclerotized plate.
</paragraph>
</caption>
<paragraph id="5A0497322A5BD85CFF6E7EA01BECB738" blockId="26.[151,1436,1967,2029]" lastBlockId="27.[151,1437,151,717]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
<emphasis id="68CF4B202A5BD85DFF6E7EA01E6FB1ED" box="[199,364,1967,1992]" italics="true" pageId="26" pageNumber="27">Male genitalia</emphasis>
: similar to those of
<taxonomicName id="9DBBECB12A5BD85DFDE77EBF1DDFB1ED" box="[590,732,1968,1992]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="terminata">
<emphasis id="68CF4B202A5BD85DFDE77EBF1DDFB1ED" box="[590,732,1968,1992]" italics="true" pageId="26" pageNumber="27">C. terminata</emphasis>
</taxonomicName>
; median lobe of aedeagus (
<figureCitation id="C2808BB72A5BD85DFBBA7EA01B5DB1ED" box="[1043,1118,1967,1992]" captionStart="FIGURES 4855" captionStartId="26.[151,264,1681,1703]" captionTargetBox="[156,1417,287,1659]" captionTargetId="figure@26.[151,1436,265,1660]" captionTargetPageId="26" captionText="FIGURES 4855. Calleida corporaali Andrewes. 48 habitus (female, Sabah, Malaysia), scale bar = 2.0 mm; 49 habitus and labels of lectotype for C. corporaali Andrewes (male, Sumatra, NHML), scale bar = 2.0 mm; 50 habitus and labels of lectotype for C. lieftincki Louwerens (female, Borneo, NNML), scale bar = 2.0 mm; 51 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (lectotype), scale bar = 0.5 mm; 52 left and right parameres of aedeagus (lectotype), scale bar = 0.2 mm; 53 male sternum IX (lectotype), scale bar = 0.5 mm; 54 ovipositor (Sabah, Malaysia), scale bar = 0.2 mm; 55 female reproductive tract (Sabah, Malaysia), scale bar = 0.5 mm; abbreviations: sp spermatheca, pd spermathecal pedicel, gd spermathecal gland duct, ga glandular area, at atrium, bpl basal sclerotized plate." httpUri="https://zenodo.org/record/1302735/files/figure.png" pageId="26" pageNumber="27">fig. 51</figureCitation>
) stout, slightly bent; middle of right side strongly dilated in dorsal and ventral aspect, forming a short lobe near the middle; dorsal and ventral margins bisinuate in lateral aspect; ventral margin with a very developed and prominent ventral lobe and a second smaller sized obtuse, rounded lobe beyond the major one; apical lamina flat, short, rounded at apex; endophallus with one only undulate, long and narrow chitinized piece, located in the middle area near the left lateral margin. Left paramere wide, depressed on the dorsal side; right paramere a little protruded at apex. (
<figureCitation id="C2808BB72A5AD85CFB35780B1BE4B738" box="[1180,1255,260,285]" captionStart="FIGURES 4855" captionStartId="26.[151,264,1681,1703]" captionTargetBox="[156,1417,287,1659]" captionTargetId="figure@26.[151,1436,265,1660]" captionTargetPageId="26" captionText="FIGURES 4855. Calleida corporaali Andrewes. 48 habitus (female, Sabah, Malaysia), scale bar = 2.0 mm; 49 habitus and labels of lectotype for C. corporaali Andrewes (male, Sumatra, NHML), scale bar = 2.0 mm; 50 habitus and labels of lectotype for C. lieftincki Louwerens (female, Borneo, NNML), scale bar = 2.0 mm; 51 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (lectotype), scale bar = 0.5 mm; 52 left and right parameres of aedeagus (lectotype), scale bar = 0.2 mm; 53 male sternum IX (lectotype), scale bar = 0.5 mm; 54 ovipositor (Sabah, Malaysia), scale bar = 0.2 mm; 55 female reproductive tract (Sabah, Malaysia), scale bar = 0.5 mm; abbreviations: sp spermatheca, pd spermathecal pedicel, gd spermathecal gland duct, ga glandular area, at atrium, bpl basal sclerotized plate." httpUri="https://zenodo.org/record/1302735/files/figure.png" pageId="27" pageNumber="28">fig. 52</figureCitation>
)
</paragraph>
<paragraph id="5A0497322A5AD85CFF6E78281E09B43D" blockId="27.[151,1437,151,717]" pageId="27" pageNumber="28">
<emphasis id="68CF4B202A5AD85CFF6E78281E8BB765" box="[199,392,295,320]" italics="true" pageId="27" pageNumber="28">Female genitalia</emphasis>
(reproductive tract
<figureCitation id="C2808BB72A5AD85CFDC478281DBBB765" box="[621,696,295,320]" captionStart="FIGURES 4855" captionStartId="26.[151,264,1681,1703]" captionTargetBox="[156,1417,287,1659]" captionTargetId="figure@26.[151,1436,265,1660]" captionTargetPageId="26" captionText="FIGURES 4855. Calleida corporaali Andrewes. 48 habitus (female, Sabah, Malaysia), scale bar = 2.0 mm; 49 habitus and labels of lectotype for C. corporaali Andrewes (male, Sumatra, NHML), scale bar = 2.0 mm; 50 habitus and labels of lectotype for C. lieftincki Louwerens (female, Borneo, NNML), scale bar = 2.0 mm; 51 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (lectotype), scale bar = 0.5 mm; 52 left and right parameres of aedeagus (lectotype), scale bar = 0.2 mm; 53 male sternum IX (lectotype), scale bar = 0.5 mm; 54 ovipositor (Sabah, Malaysia), scale bar = 0.2 mm; 55 female reproductive tract (Sabah, Malaysia), scale bar = 0.5 mm; abbreviations: sp spermatheca, pd spermathecal pedicel, gd spermathecal gland duct, ga glandular area, at atrium, bpl basal sclerotized plate." httpUri="https://zenodo.org/record/1302735/files/figure.png" pageId="27" pageNumber="28">fig. 55</figureCitation>
and ovipositor
<figureCitation id="C2808BB72A5AD85CFCC778281CBDB765" box="[878,958,295,320]" captionStart="FIGURES 4855" captionStartId="26.[151,264,1681,1703]" captionTargetBox="[156,1417,287,1659]" captionTargetId="figure@26.[151,1436,265,1660]" captionTargetPageId="26" captionText="FIGURES 4855. Calleida corporaali Andrewes. 48 habitus (female, Sabah, Malaysia), scale bar = 2.0 mm; 49 habitus and labels of lectotype for C. corporaali Andrewes (male, Sumatra, NHML), scale bar = 2.0 mm; 50 habitus and labels of lectotype for C. lieftincki Louwerens (female, Borneo, NNML), scale bar = 2.0 mm; 51 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (lectotype), scale bar = 0.5 mm; 52 left and right parameres of aedeagus (lectotype), scale bar = 0.2 mm; 53 male sternum IX (lectotype), scale bar = 0.5 mm; 54 ovipositor (Sabah, Malaysia), scale bar = 0.2 mm; 55 female reproductive tract (Sabah, Malaysia), scale bar = 0.5 mm; abbreviations: sp spermatheca, pd spermathecal pedicel, gd spermathecal gland duct, ga glandular area, at atrium, bpl basal sclerotized plate." httpUri="https://zenodo.org/record/1302735/files/figure.png" pageId="27" pageNumber="28">fig. 54</figureCitation>
): spermatheca digitiform, about 1.7 times length as the pedicel, surface not whorled, with evident basal sclerotized plate (
<emphasis id="68CF4B202A5AD85CFBBA78431BF3B740" box="[1043,1264,332,357]" italics="true" pageId="27" pageNumber="28">annulus receptaculi</emphasis>
); spermathecal pedicel not expanded; spermathecal gland duct about half length as spermatheca, laterally inserted below the basal sclerotized plate; glandular area weakly inflated, about same length as gland duct, base with small protuberant atrium. Gonocoxite II of ovipositor about 2.5 times as long as the basal width, narrowed to apex, apex oblique truncate,; inner margin setose from the basal third to apex; outer and inner margins straight; apex with membranous extension.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A5AD85CFF6E7B2B1DE6B4E9" pageId="27" pageNumber="28" type="distribution">
<paragraph id="5A0497322A5AD85CFF6E7B2B1A1BB445" blockId="27.[151,1437,151,717]" pageId="27" pageNumber="28">
<emphasis id="68CF4B202A5AD85CFF6E7B2B1DABB418" bold="true" box="[199,680,548,573]" pageId="27" pageNumber="28">Geographical distribution and habitat.</emphasis>
<collectingCountry id="22ACD7A22A5AD85CFD1C7B2B1C21B418" box="[693,802,548,573]" name="Indonesia" pageId="27" pageNumber="28">Indonesia</collectingCountry>
and
<collectingCountry id="22ACD7A22A5AD85CFCCC7B2B1CCFB418" box="[869,972,548,573]" name="Malaysia" pageId="27" pageNumber="28">Malaysia</collectingCountry>
(
<figureCitation id="C2808BB72A5AD85CFC487B2B1B32B419" box="[993,1073,548,572]" captionStart="MAP 4" captionStartId="29.[151,209,1076,1098]" captionTargetBox="[151,1436,195,1054]" captionTargetId="figure@29.[151,1436,195,1054]" captionTargetPageId="29" captionText="MAP 4 Distributions for species in the C. terminata group: C. terminata Waterhouse (green); C. corporaali Andrewes (blue); C. viet sp. n. (red)." httpUri="https://zenodo.org/record/1302739/files/figure.png" pageId="27" pageNumber="28">map 4</figureCitation>
):
<collectingRegion id="987F59D02A5AD85CFBE47B2B1BAEB419" box="[1101,1197,548,572]" country="Indonesia" name="Sumatera Utara" pageId="27" pageNumber="28">Sumatra</collectingRegion>
, Borneo and Malay Peninsula. This species is sympatric with
<taxonomicName id="9DBBECB12A5AD85CFDC37B471DFBB445" box="[618,760,584,608]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="terminata">
<emphasis id="68CF4B202A5AD85CFDC37B471DFBB445" box="[618,760,584,608]" italics="true" pageId="27" pageNumber="28">C. terminata</emphasis>
</taxonomicName>
and
<taxonomicName id="9DBBECB12A5AD85CFC877B471B42B47A" authority="in Borneo" authorityName="in Borneo" box="[814,1089,583,608]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis id="68CF4B202A5AD85CFC877B471CCBB445" box="[814,968,583,608]" italics="true" pageId="27" pageNumber="28">C. borneensis</emphasis>
in Borneo
</taxonomicName>
, in tropical forests.
</paragraph>
<paragraph id="5A0497322A5AD85CFF6E7B631DE6B4E9" blockId="27.[151,1437,151,717]" pageId="27" pageNumber="28">
Besides examined materials,
<bibRefCitation id="3E2AEAC32A5AD85CFDA47B631DD3B4A1" box="[525,720,620,645]" pageId="27" pageNumber="28" refId="ref28873" refString="Andrewes, H. E. (1929) Papers on Oriental Carabidae. XXII. The Annals and Magazine of Natural History, Series 10, 4 (22), 353 - 371. https: // doi. org / 10.1080 / 00222932908673067" type="journal article">Andrewes (1929)</bibRefCitation>
reported this species also from
<collectingRegion id="987F59D02A5AD85CFB9B7B631BD9B4A1" box="[1074,1242,620,645]" country="Malaysia" name="Kuala Lumpur" pageId="27" pageNumber="28">Kuala Lumpur</collectingRegion>
;
<collectingRegion id="987F59D02A5AD85CFB4C7B631A49B4A0" box="[1253,1354,620,645]" country="Malaysia" name="Selangor" pageId="27" pageNumber="28">Selangor</collectingRegion>
,
<collectingRegion id="987F59D02A5AD85CFAFC7B631A9BB4A0" box="[1365,1432,620,645]" country="Malaysia" name="Perak" pageId="27" pageNumber="28">Perak</collectingRegion>
, Gunong Kledang, and Kudat (Borneo). The record of this species in Kudat (N Borneo,
<bibRefCitation id="3E2AEAC32A5AD85CFBDC7B801A2DB48D" box="[1141,1326,655,680]" pageId="27" pageNumber="28" refId="ref28873" refString="Andrewes, H. E. (1929) Papers on Oriental Carabidae. XXII. The Annals and Magazine of Natural History, Series 10, 4 (22), 353 - 371. https: // doi. org / 10.1080 / 00222932908673067" type="journal article">Andrewes, 1929</bibRefCitation>
) needs to be confirmed (see the Distribution in
<taxonomicName id="9DBBECB12A5AD85CFD927BBA1DD7B4E8" box="[571,724,692,717]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis id="68CF4B202A5AD85CFD927BBA1DD7B4E8" box="[571,724,692,717]" italics="true" pageId="27" pageNumber="28">C. borneensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>

View file

@ -0,0 +1,220 @@
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<mods:title id="5AADFF3A0340C7D2A068A99F8BC2F7A2">Revision of the Oriental species of Calleida Latreille (sensu lato). Part 1: Introduction, groups of species, and species of six species groups (Coleoptera: Carabidae: Lebiini)</mods:title>
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<mods:date id="EDF34BEDC4775CFA1A302D464493B605">2018</mods:date>
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<treatment id="D21226242A5AD85BFF3E7A2F1B9AB1A4" ID-DOI="http://doi.org/10.5281/zenodo.5950871" ID-GBIF-Taxon="146043499" ID-Zenodo-Dep="5950871" LSID="urn:lsid:plazi:treatment:D21226242A5AD85BFF3E7A2F1B9AB1A4" httpUri="http://treatment.plazi.org/id/D21226242A5AD85BFF3E7A2F1B9AB1A4" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<subSubSection id="12A1C4B92A5AD85CFF3E7A2F1D51B51F" box="[151,594,799,826]" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="5A0497322A5AD85CFF3E7A2F1D51B51F" blockId="27.[151,594,799,826]" box="[151,594,799,826]" pageId="27" pageNumber="28">
<heading id="014C205E2A5AD85CFF3E7A2F1D51B51F" bold="true" box="[151,594,799,826]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
<emphasis id="68CF4B202A5AD85CFF3E7A2F1D51B51F" bold="true" box="[151,594,799,826]" pageId="27" pageNumber="28">
[8]
<taxonomicName id="9DBBECB12A5AD85CFF697A101D01B51F" ID-CoL="8MK8T" authority="Casale &amp; Shi" authorityName="Casale &amp; Shi" box="[192,514,799,826]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="viet" status="sp. nov.">
<emphasis id="68CF4B202A5AD85CFF697A101E54B51C" bold="true" box="[192,343,799,825]" italics="true" pageId="27" pageNumber="28">Calleida viet</emphasis>
Casale &amp; Shi
</taxonomicName>
,
<taxonomicNameLabel id="73FCF65B2A5AD85CFDB97A2F1D51B51F" box="[528,594,800,826]" pageId="27" pageNumber="28" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A5AD85CFF3E7A691EE8B5E2" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="5A0497322A5AD85CFF3E7A691D21B55B" blockId="27.[151,1437,870,2011]" box="[151,546,870,895]" pageId="27" pageNumber="28">
<emphasis id="68CF4B202A5AD85CFF3E7A691E3AB55A" bold="true" box="[151,313,870,895]" pageId="27" pageNumber="28">
<typeStatus id="850029902A5AD85CFF3E7A691FD1B55A" box="[151,210,870,895]" pageId="27" pageNumber="28">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="22ACD7A22A5AD85CFEE97A691EA7B55A" box="[320,420,870,895]" name="Vietnam" pageId="27" pageNumber="28">Vietnam</collectingCountry>
: Vung Tau.
</paragraph>
<paragraph id="5A0497322A5AD85CFF6E7A851EE8B5E2" blockId="27.[151,1437,870,2011]" pageId="27" pageNumber="28">
<emphasis id="68CF4B202A5AD85CFF6E7A851E74B586" bold="true" box="[199,375,906,931]" pageId="27" pageNumber="28">
<typeStatus id="850029902A5AD85CFF6E7A851E01B586" box="[199,258,906,931]" pageId="27" pageNumber="28">Type</typeStatus>
material:
</emphasis>
<materialsCitation id="EAD39D6F2A5AD85CFED77A851B24B586" ID-GBIF-Occurrence="1852047370" box="[382,1063,906,931]" collectingDate="1988-11" country="Vietnam" location="Vung Tau" pageId="27" pageNumber="28" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="850029902A5AD85CFED77A851EEAB586" box="[382,489,906,931]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
,
<specimenCount id="4CBD5CBB2A5AD85CFE5C7A851D2DB586" box="[501,558,906,931]" pageId="27" pageNumber="28" type="male">male</specimenCount>
, “
<collectingCountry id="22ACD7A22A5AD85CFDED7A851DABB586" box="[580,680,906,931]" name="Vietnam" pageId="27" pageNumber="28">Vietnam</collectingCountry>
,
<location id="5F64C1E92A5AD85CFD1A7A841C22B587" LSID="urn:lsid:plazi:treatment:D21226242A5AD85BFF3E7A2F1B9AB1A4:5F64C1E92A5AD85CFD1A7A841C22B587" box="[691,801,907,931]" country="Vietnam" name="Vung Tau" pageId="27" pageNumber="28">Vung Tau</location>
<collectingDate id="3E41481A2A5AD85CFC817A841C69B586" box="[808,874,907,931]" pageId="27" pageNumber="28" value="1988-11">
<date id="2E05B1F22A5AD85CFC817A841C69B586" box="[808,874,907,931]" pageId="27" pageNumber="28" value="1988-11">XI.88</date>
</collectingDate>
” (CCA,
<figureCitation id="C2808BB72A5AD85CFC677A851B18B586" box="[974,1051,906,931]" captionStart="FIGURES 5660" captionStartId="28.[151,264,1120,1142]" captionTargetBox="[159,1407,265,1093]" captionTargetId="figure@28.[151,1436,265,1099]" captionTargetPageId="28" captionText="FIGURES 5660. Calleida viet sp. n. 56 habitus and labels of holotype (male, Vung Tau, Vietnam, CCA), scale bar = 2.0 mm; 5759 male genitalia (holotype), scale bar = 0.5 mm: 57 median lobe of aedeagus, ventral view; 58 left paramere of aedeagus; 59 median lobe of aedeagus, left-lateral view; 60 female reproductive tract and ovipositor (paratype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302737/files/figure.png" pageId="27" pageNumber="28">fig. 56</figureCitation>
).
</materialsCitation>
<typeStatus id="850029902A5AD85CFB867A841B96B586" box="[1071,1173,907,931]" pageId="27" pageNumber="28" type="paratype">Paratype</typeStatus>
: one female, “
<collectingCountry id="22ACD7A22A5AD85CFA9D7A851A9BB586" box="[1332,1432,906,931]" name="Vietnam" pageId="27" pageNumber="28">Vietnam</collectingCountry>
, Vung Tau 20-
<date id="2E05B1F22A5AD85CFE867AA01E84B5E2" box="[303,391,943,967]" pageId="27" pageNumber="28" value="88-07-30">30.7.88</date>
” (
<collectionCode id="3CAA0FF72A5AD85CFE097AA01EE3B5E2" box="[416,480,943,967]" country="Germany" httpUri="http://grbio.org/cool/mqei-3i36" name="Culture Collection of Algae at the University of Marburg" pageId="27" pageNumber="28">CCA</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A5AD85CFF6E7ADD1ECAB22A" pageId="27" pageNumber="28" type="etymology">
<paragraph id="5A0497322A5AD85CFF6E7ADD1ECAB22A" blockId="27.[151,1437,870,2011]" pageId="27" pageNumber="28">
<emphasis id="68CF4B202A5AD85CFF6E7ADD1E85B5CE" bold="true" box="[199,390,978,1003]" pageId="27" pageNumber="28">Specific epithet.</emphasis>
This new species is dedicated to the Viet people, living in the country where is located the
<typeStatus id="850029902A5AD85CFF3E7AF81FC4B22A" box="[151,199,1015,1039]" pageId="27" pageNumber="28">type</typeStatus>
locality of the species.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A5AD85CFF6E7D151A9EB3C6" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="5A0497322A5AD85CFF6E7D151CA7B32E" blockId="27.[151,1437,870,2011]" pageId="27" pageNumber="28">
<emphasis id="68CF4B202A5AD85CFF6E7D151E42B216" bold="true" box="[199,321,1050,1075]" pageId="27" pageNumber="28">Diagnosis.</emphasis>
With the character states of the
<taxonomicName id="9DBBECB12A5AD85CFD1C7D151C88B216" box="[693,907,1050,1075]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="terminata">
<emphasis id="68CF4B202A5AD85CFD1C7D151C88B216" box="[693,907,1050,1075]" italics="true" pageId="27" pageNumber="28">Calleida terminata</emphasis>
</taxonomicName>
species group, but distinct from the two other species of this group by the following character combination: medium sized (L:
<quantity id="9D433AD72A5AD85CFBEE7D311BA6B272" box="[1095,1189,1086,1111]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="27" pageNumber="28" unit="mm" value="9.5">9.5 mm</quantity>
); head and pronotum strongly metallic green or with cupreous reflections; elytra metallic green with wide cupreous metallic refection patch on apical and lateral regions; antennae, palpomeres, lateral furrows of pronotum, coxae and tarsi reddish; femora and underside strongly metallic green. Pronotum with deep transverse wrinkles. Elytra with outer apical angle acutely prominent apically. Median lobe of aedeagus in lateral aspect with ventral margin pointed beyond the main ventral lobe; endophallus with two chitinized copulatory pieces.
</paragraph>
<paragraph id="5A0497322A5AD85CFF6E7C191A9EB3C6" blockId="27.[151,1437,870,2011]" pageId="27" pageNumber="28">
This new species is similar to
<taxonomicName id="9DBBECB12A5AD85CFD8B7C181DB2B30B" box="[546,689,1303,1326]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="terminata">
<emphasis id="68CF4B202A5AD85CFD8B7C181DB2B30B" box="[546,689,1303,1326]" italics="true" pageId="27" pageNumber="28">C. terminata</emphasis>
</taxonomicName>
. In external features, these two species can be distinguished by a few characters: in
<taxonomicName id="9DBBECB12A5AD85CFECA7C341EAFB376" box="[355,428,1339,1363]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="viet">
<emphasis id="68CF4B202A5AD85CFECA7C341EAFB376" box="[355,428,1339,1363]" italics="true" pageId="27" pageNumber="28">C. viet</emphasis>
</taxonomicName>
, the first antennomere is without any trace of metallic reflection, the femora and underside are less distinctly metallic, and the size is smaller. But the differences in the distribution range and the characters in the male genitalia can easily separate these two species: in
<taxonomicName id="9DBBECB12A5AD85CFC937C8C1C87B3BE" box="[826,900,1411,1435]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="viet">
<emphasis id="68CF4B202A5AD85CFC937C8C1C87B3BE" box="[826,900,1411,1435]" italics="true" pageId="27" pageNumber="28">C. viet</emphasis>
</taxonomicName>
, the median lobe of aedeagus in ventral aspect has the main ventral lobe emarginated at the apex; in lateral aspect, the ventral margin beyond the main ventral lobe shows a smaller lobe, and the apex is strongly dilated at the apical orifice; endophallus with two copulatory pieces.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A5AD85BFF6E7CE11A90B11C" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="description">
<paragraph id="5A0497322A5AD85CFF6E7CE11BA4B023" blockId="27.[151,1437,870,2011]" box="[199,1191,1518,1543]" pageId="27" pageNumber="28">
<emphasis id="68CF4B202A5AD85CFF6E7CE11E5AB022" bold="true" box="[199,345,1518,1543]" pageId="27" pageNumber="28">Description.</emphasis>
<emphasis id="68CF4B202A5AD85CFEF67CE11D18B022" box="[351,539,1518,1543]" italics="true" pageId="27" pageNumber="28">General features</emphasis>
: (
<figureCitation id="C2808BB72A5AD85CFD9B7CE11D7EB022" box="[562,637,1518,1543]" captionStart="FIGURES 5660" captionStartId="28.[151,264,1120,1142]" captionTargetBox="[159,1407,265,1093]" captionTargetId="figure@28.[151,1436,265,1099]" captionTargetPageId="28" captionText="FIGURES 5660. Calleida viet sp. n. 56 habitus and labels of holotype (male, Vung Tau, Vietnam, CCA), scale bar = 2.0 mm; 5759 male genitalia (holotype), scale bar = 0.5 mm: 57 median lobe of aedeagus, ventral view; 58 left paramere of aedeagus; 59 median lobe of aedeagus, left-lateral view; 60 female reproductive tract and ovipositor (paratype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302737/files/figure.png" pageId="27" pageNumber="28">fig. 56</figureCitation>
) Medium-sized species: L:
<quantity id="9D433AD72A5AD85CFC1C7CE11B0DB023" box="[949,1038,1518,1543]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="27" pageNumber="28" unit="mm" value="9.5">9.5 mm</quantity>
; TL: 9.0 mm.
</paragraph>
<paragraph id="5A0497322A5AD85CFF6E7F1D1B14B09F" blockId="27.[151,1437,870,2011]" pageId="27" pageNumber="28">
<emphasis id="68CF4B202A5AD85CFF6E7F1D1E15B00E" box="[199,278,1554,1579]" italics="true" pageId="27" pageNumber="28">Colour</emphasis>
: Head metallic green; clypeus, labrum and mandibles blackish; frons dark metallic green, vertex with two reddish spots; antennae and tarsi reddish; tibiae dark brown, femora blackish with marked metallic greenish reflection. Pronotum metallic green or green with cupreous metallic reflection; elytra metallic green, a little cupreous at base, with wide cupreous-purple metallic refection patch on the apical and lateral regions; underside dark brown, with metallic green reflection on prosternum and abdominal sterna.
</paragraph>
<paragraph id="5A0497322A5AD85CFF6E7FC81E57B126" blockId="27.[151,1437,870,2011]" pageId="27" pageNumber="28">
<emphasis id="68CF4B202A5AD85CFF6E7FC81EEFB0FA" box="[199,492,1734,1759]" italics="true" pageId="27" pageNumber="28">Lustre and microsculpture</emphasis>
: very shiny, polished. Isodiametric meshes of the microsculpture on head and elytra almost vanished.
</paragraph>
<paragraph id="5A0497322A5AD85CFF6E7E011D6BB14A" blockId="27.[151,1437,870,2011]" pageId="27" pageNumber="28">
<emphasis id="68CF4B202A5AD85CFF6E7E011E07B102" box="[199,260,1806,1831]" italics="true" pageId="27" pageNumber="28">Head</emphasis>
wide, flat, smooth; genae markedly constricted to the neck; frontal foveae shallow, sub-striate in front; genae very short, markedly narrowed to the neck constriction; eyes prominent, hemispherical; antennae reaching backwards to the basal sixth of the elytra.
</paragraph>
<paragraph id="5A0497322A5AD85BFF6E7E731E0FB6F0" blockId="27.[151,1437,870,2011]" lastBlockId="28.[151,1436,151,213]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<emphasis id="68CF4B202A5AD85CFF6E7E731E35B1B6" box="[199,310,1916,1939]" italics="true" pageId="27" pageNumber="28">Pronotum</emphasis>
cordate, wider than long (ratio PW/PL: 1.25). Disc subconvex, with deep transverse wrinkles extended to the lateral furrows; median furrow complete, reaching the basal margin, markedly impressed. Anterior angles rounded; lateral margins rounded in front, deeply sinuate in the basal fifth. Basal margin bisinuate, oblique at sides; posterior angles reflexed and prominent laterally, sharply pointed but apex obtuse; basal foveae short and very deep.
</paragraph>
<caption id="0EC4C7BA2A5DD85BFF3E7D6F1A66B296" httpUri="https://zenodo.org/record/1302737/files/figure.png" pageId="28" pageNumber="29" startId="28.[151,264,1120,1142]" targetBox="[159,1407,265,1093]" targetPageId="28">
<paragraph id="5A0497322A5DD85BFF3E7D6F1A66B296" blockId="28.[151,1436,1120,1203]" pageId="28" pageNumber="29">
<emphasis id="68CF4B202A5DD85BFF3E7D6F1EDFB253" bold="true" box="[151,476,1120,1142]" pageId="28" pageNumber="29">
FIGURES 5660.
<taxonomicName id="9DBBECB12A5DD85BFEF07D6F1EDFB253" box="[345,476,1120,1142]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="viet" status="sp. nov.">
<emphasis id="68CF4B202A5DD85BFEF07D6F1EDFB253" bold="true" box="[345,476,1120,1142]" italics="true" pageId="28" pageNumber="29">Calleida viet</emphasis>
</taxonomicName>
</emphasis>
<taxonomicNameLabel id="73FCF65B2A5DD85BFE4B7D6E1D1BB253" box="[482,536,1121,1142]" pageId="28" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
<emphasis id="68CF4B202A5DD85BFDB77D6F1D3BB253" bold="true" box="[542,568,1120,1142]" pageId="28" pageNumber="29">56</emphasis>
habitus and labels of holotype (male, Vung Tau, Vietnam, CCA), scale bar = 2.0 mm;
<emphasis id="68CF4B202A5DD85BFF3E7D701FD5B2B0" bold="true" box="[151,214,1151,1173]" pageId="28" pageNumber="29">5759</emphasis>
male genitalia (holotype), scale bar = 0.5 mm:
<emphasis id="68CF4B202A5DD85BFD137D701DD0B2B0" bold="true" box="[698,723,1151,1173]" pageId="28" pageNumber="29">57</emphasis>
median lobe of aedeagus, ventral view;
<emphasis id="68CF4B202A5DD85BFBC67D701B8BB2B0" bold="true" box="[1135,1160,1151,1173]" pageId="28" pageNumber="29">58</emphasis>
left paramere of aedeagus;
<emphasis id="68CF4B202A5DD85BFF3E7D921FB2B296" bold="true" box="[151,177,1181,1203]" pageId="28" pageNumber="29">59</emphasis>
median lobe of aedeagus, left-lateral view;
<emphasis id="68CF4B202A5DD85BFDC57D921D85B296" bold="true" box="[620,646,1181,1203]" pageId="28" pageNumber="29">60</emphasis>
female reproductive tract and ovipositor (paratype), scale bar = 0.5 mm.
</paragraph>
</caption>
<paragraph id="5A0497322A5DD85BFF6E7DEF1A0FB3AC" blockId="28.[151,1436,1248,1921]" pageId="28" pageNumber="29">
<emphasis id="68CF4B202A5DD85BFF6E7DEF1E0EB2DC" box="[199,269,1248,1273]" italics="true" pageId="28" pageNumber="29">Elytra</emphasis>
: sub-depressed, relatively short (EL/EW: 1.68), transversely depressed on each side behind base, faintly dilated behind. Humeral angles rounded, lateral sides slightly compressed at one third from base; apex obliquely truncate, distinctly thickened at outer apical angles, which are sharp and acutely prominent apically. Striae deep, finely punctate; intervals convex at base, flattened on disc, with fine, shallow but distinct punctuation; the third interval with two discal and one apical pores. Pre-apical callosity on eighth interval small but evident.
</paragraph>
<paragraph id="5A0497322A5DD85BFF6E7C9B1E80B3F4" blockId="28.[151,1436,1248,1921]" pageId="28" pageNumber="29">
<emphasis id="68CF4B202A5DD85BFF6E7C9B1E3FB388" box="[199,316,1428,1453]" italics="true" pageId="28" pageNumber="29">Underside</emphasis>
: Abdominal sternum VII in male deeply notched at middle and with one seta on each side in male; two setae in females.
</paragraph>
<paragraph id="5A0497322A5DD85BFF6E7CD31C14B0E8" blockId="28.[151,1436,1248,1921]" pageId="28" pageNumber="29">
<emphasis id="68CF4B202A5DD85BFF6E7CD31E73B3D0" box="[199,368,1500,1525]" italics="true" pageId="28" pageNumber="29">Male genitalia</emphasis>
: Median lobe of aedeagus (
<figureCitation id="C2808BB72A5DD85BFD687CD31C48B3D0" box="[705,843,1500,1525]" captionStart="FIGURES 5660" captionStartId="28.[151,264,1120,1142]" captionTargetBox="[159,1407,265,1093]" captionTargetId="figure@28.[151,1436,265,1099]" captionTargetPageId="28" captionText="FIGURES 5660. Calleida viet sp. n. 56 habitus and labels of holotype (male, Vung Tau, Vietnam, CCA), scale bar = 2.0 mm; 5759 male genitalia (holotype), scale bar = 0.5 mm: 57 median lobe of aedeagus, ventral view; 58 left paramere of aedeagus; 59 median lobe of aedeagus, left-lateral view; 60 female reproductive tract and ovipositor (paratype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302737/files/figure.png" pageId="28" pageNumber="29">figs. 57, 59</figureCitation>
) very peculiar in shape: in ventral aspect, middle strongly dilated to the right side; in lateral aspect, dorsal margin sinuate, ventral margin with two developed lobes, the major one near the middle, very prominent, thickened and chitinized on the anterior margin, apex emarginate in ventral aspect; the minor one a little beyond the major one, smaller and obtuse; apical lamina very short, rounded at apex. Endophallus with two copulatory pieces, located in the middle area near the left lateral margin, very different in shape to each other: the left one smaller and thin, V shaped, the right one much bigger, sinuate, spine-like. Left paramere elongate, depressed on the dorsal side. (
<figureCitation id="C2808BB72A5DD85BFD6A7FBB1C0DB0E8" box="[707,782,1716,1741]" captionStart="FIGURES 5660" captionStartId="28.[151,264,1120,1142]" captionTargetBox="[159,1407,265,1093]" captionTargetId="figure@28.[151,1436,265,1099]" captionTargetPageId="28" captionText="FIGURES 5660. Calleida viet sp. n. 56 habitus and labels of holotype (male, Vung Tau, Vietnam, CCA), scale bar = 2.0 mm; 5759 male genitalia (holotype), scale bar = 0.5 mm: 57 median lobe of aedeagus, ventral view; 58 left paramere of aedeagus; 59 median lobe of aedeagus, left-lateral view; 60 female reproductive tract and ovipositor (paratype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302737/files/figure.png" pageId="28" pageNumber="29">fig. 58</figureCitation>
)
</paragraph>
<paragraph id="5A0497322A5DD85BFF6E7FD71A90B11C" blockId="28.[151,1436,1248,1921]" pageId="28" pageNumber="29">
<emphasis id="68CF4B202A5DD85BFF6E7FD71E8AB0D4" box="[199,393,1752,1777]" italics="true" pageId="28" pageNumber="29">Female genitalia</emphasis>
(reproductive tract and ovipositor,
<figureCitation id="C2808BB72A5DD85BFC8D7FD71C77B0D4" box="[804,884,1752,1777]" captionStart="FIGURES 5660" captionStartId="28.[151,264,1120,1142]" captionTargetBox="[159,1407,265,1093]" captionTargetId="figure@28.[151,1436,265,1099]" captionTargetPageId="28" captionText="FIGURES 5660. Calleida viet sp. n. 56 habitus and labels of holotype (male, Vung Tau, Vietnam, CCA), scale bar = 2.0 mm; 5759 male genitalia (holotype), scale bar = 0.5 mm: 57 median lobe of aedeagus, ventral view; 58 left paramere of aedeagus; 59 median lobe of aedeagus, left-lateral view; 60 female reproductive tract and ovipositor (paratype), scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1302737/files/figure.png" pageId="28" pageNumber="29">fig. 60</figureCitation>
): spermatheca digitiform, tumid at apex, longer than the pedicel, with an evident basal sclerotized plate
<emphasis id="68CF4B202A5DD85BFCBF7FF31B05B130" box="[790,1030,1788,1813]" italics="true" pageId="28" pageNumber="29">(annulus receptaculi)</emphasis>
, surface not whorled; spermathecal pedicel not expanded in the basal half; spermathecal gland and gonocoxites I and II as in description of the genus.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A5DD85BFF6E7E4B1B9AB1A4" pageId="28" pageNumber="29" type="distribution">
<paragraph id="5A0497322A5DD85BFF6E7E4B1B9AB1A4" blockId="28.[151,1436,1248,1921]" pageId="28" pageNumber="29">
<emphasis id="68CF4B202A5DD85BFF6E7E4B1D97B178" bold="true" box="[199,660,1860,1885]" pageId="28" pageNumber="29">Geographical distribution and habitat.</emphasis>
<collectingCountry id="22ACD7A22A5DD85BFD327E4B1DFCB178" box="[667,767,1860,1885]" name="Vietnam" pageId="28" pageNumber="29">Vietnam</collectingCountry>
: Vung Tau, known so far only from the
<typeStatus id="850029902A5DD85BFB107E4A1BEAB178" box="[1209,1257,1861,1885]" pageId="28" pageNumber="29">type</typeStatus>
locality located in the southern coast of the country (
<figureCitation id="C2808BB72A5DD85BFD997E661D79B1A4" box="[560,634,1897,1921]" captionStart="MAP 4" captionStartId="29.[151,209,1076,1098]" captionTargetBox="[151,1436,195,1054]" captionTargetId="figure@29.[151,1436,195,1054]" captionTargetPageId="29" captionText="MAP 4 Distributions for species in the C. terminata group: C. terminata Waterhouse (green); C. corporaali Andrewes (blue); C. viet sp. n. (red)." httpUri="https://zenodo.org/record/1302739/files/figure.png" pageId="28" pageNumber="29">map 4</figureCitation>
). Probably sampled in tropical secondary forest.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,120 @@
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<subSubSection id="12A1C4B92A5CD85AFF3E7DBB1D51B2EB" box="[151,594,1204,1230]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="5A0497322A5CD85AFF3E7DBB1D51B2EB" blockId="29.[151,594,1204,1230]" box="[151,594,1204,1230]" pageId="29" pageNumber="30">
<heading id="014C205E2A5CD85AFF3E7DBB1D51B2EB" bold="true" box="[151,594,1204,1230]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
<emphasis id="68CF4B202A5CD85AFF3E7DBB1D51B2EB" bold="true" box="[151,594,1204,1230]" pageId="29" pageNumber="30">
6.
<taxonomicName id="9DBBECB12A5CD85AFF1D7DBB1EA1B2EB" ID-CoL="8M5HW" box="[180,418,1204,1230]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis id="68CF4B202A5CD85AFF1D7DBB1EA1B2EB" bold="true" box="[180,418,1204,1230]" italics="true" pageId="29" pageNumber="30">Calleida borneensis</emphasis>
</taxonomicName>
species group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A5CD85AFF3E7DF51A6EB12E" pageId="29" pageNumber="30" type="description">
<paragraph id="5A0497322A5CD85AFF3E7DF51FC0B3CE" blockId="29.[151,1437,1274,1803]" pageId="29" pageNumber="30">
<emphasis id="68CF4B202A5CD85AFF3E7DF51EA3B336" bold="true" box="[151,416,1274,1299]" pageId="29" pageNumber="30">Diagnostic characters</emphasis>
: With the character states of
<taxonomicName id="9DBBECB12A5CD85AFCA57DF51C6FB336" box="[780,876,1274,1299]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A5CD85AFCA57DF51C6FB336" box="[780,876,1274,1299]" italics="true" pageId="29" pageNumber="30">Calleida</emphasis>
</taxonomicName>
(in the widest sense, see above), restricted or amplified as follows: medium sized species (L:
<quantity id="9D433AD72A5CD85AFD107C101C45B312" box="[697,838,1311,1335]" metricMagnitude="-3" metricUnit="m" metricValue="9.15" metricValueMax="9.9" metricValueMin="8.4" pageId="29" pageNumber="30" unit="mm" value="9.15" valueMax="9.9" valueMin="8.4">8.49.9 mm</quantity>
). Pronotum and the ventral side dark brownish, with very faint metallic green reflection on pronotal disc, vertex with reddish patch; elytra metallic green, with golden reddish purple hue at sides and near apex. Elytral apex strongly curved, outer apical angles markedly widened, but not forming acute angles (
<figureCitation id="C2808BB72A5CD85AFE127C851D0BB386" box="[443,520,1418,1443]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="29" pageNumber="30">fig. 16</figureCitation>
). Abdominal sternum VII in male moderately notched at middle, with one seta on each side in males, two setae in females. Median lobe of aedeagus with apical lamina capitate and securiform (
<figureCitation id="C2808BB72A5CD85AFADC7CA11FB4B3CE" captionStart="FIGURES 6166" captionStartId="31.[151,264,1853,1875]" captionTargetBox="[159,1432,203,1814]" captionTargetId="figure@31.[151,1436,193,1832]" captionTargetPageId="31" captionText="FIGURES 6166. Calleida borneensis sp. n. 61 habitus and labels of holotype (male, Sabah, Malaysia, NHML), scale bar = 2.0 mm; 62 habitus of paratype (male, Sarawak, Malaysia, NHML), scale bar = 2.0 mm; 63 female reproductive tract (Sabah, Malaysia), scale bar = 0.5 mm; abbreviations: sp spermatheca, pd spermathecal pedicel, gd spermathecal gland duct, ga glandular area, at atrium, bpj spermathecal basal projection, apt pedicel apical protuberance; 64 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm; 65 left and right parameres of aedeagus (holotype), scale bar = 0.2 mm; 66 ovipositor (Sabah, Malaysia), scale bar = 0.2 mm." httpUri="https://zenodo.org/record/1302741/files/figure.png" pageId="29" pageNumber="30">fig. 64</figureCitation>
).
</paragraph>
<paragraph id="5A0497322A5CD85AFF6E7CF91C5FB016" blockId="29.[151,1437,1274,1803]" pageId="29" pageNumber="30">
<emphasis id="68CF4B202A5CD85AFF6E7CF91E8BB02A" bold="true" box="[199,392,1526,1551]" pageId="29" pageNumber="30">Taxonomic note</emphasis>
: Only one species,
<taxonomicName id="9DBBECB12A5CD85AFDDD7CF81C11B02A" box="[628,786,1526,1551]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="borneensis" status="sp. nov.">
<emphasis id="68CF4B202A5CD85AFDDD7CF81C11B02A" box="[628,786,1526,1551]" italics="true" pageId="29" pageNumber="30">C. borneensis</emphasis>
</taxonomicName>
<emphasis id="68CF4B202A5CD85AFCB47CF91C63B02A" bold="true" box="[797,864,1526,1551]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="73FCF65B2A5CD85AFCB47CF91C63B02A" box="[797,864,1526,1551]" pageId="29" pageNumber="30" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is attributable to this group. This rare species is known so far only from a few specimens from Borneo (
<figureCitation id="C2808BB72A5CD85AFCAC7F141C4DB016" box="[773,846,1562,1587]" captionStart="MAP 5" captionStartId="32.[151,209,1330,1352]" captionTargetBox="[264,1323,517,1309]" captionTargetId="figure@32.[264,1323,517,1309]" captionTargetPageId="32" captionText="MAP 5 Distributions for species in the C. borneensis group: C. borneensis sp. n. (red)." httpUri="https://zenodo.org/record/1302743/files/figure.png" pageId="29" pageNumber="30">map 5</figureCitation>
).
</paragraph>
<paragraph id="5A0497322A5CD85AFF6E7F301A63B0C2" blockId="29.[151,1437,1274,1803]" pageId="29" pageNumber="30">
<taxonomicName id="9DBBECB12A5CD85AFF6E7F301E66B072" box="[199,357,1598,1623]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="borneensis" status="sp. nov.">
<emphasis id="68CF4B202A5CD85AFF6E7F301E66B072" box="[199,357,1598,1623]" italics="true" pageId="29" pageNumber="30">C. borneensis</emphasis>
</taxonomicName>
<emphasis id="68CF4B202A5CD85AFED97F311EB7B072" bold="true" box="[368,436,1598,1623]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="73FCF65B2A5CD85AFED97F311EB7B072" box="[368,436,1598,1623]" pageId="29" pageNumber="30" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to members of the
<taxonomicName id="9DBBECB12A5CD85AFCB17F301CA9B072" box="[792,938,1599,1623]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="terminata">
<emphasis id="68CF4B202A5CD85AFCB17F301CA9B072" box="[792,938,1599,1623]" italics="true" pageId="29" pageNumber="30">C. terminata</emphasis>
</taxonomicName>
group (especially to
<taxonomicName id="9DBBECB12A5CD85AFB0E7F301A47B072" authorityName="Andr. H. E. Andrewes" authorityYear="1929" box="[1191,1348,1598,1623]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="corporaali">
<emphasis id="68CF4B202A5CD85AFB0E7F301A47B072" box="[1191,1348,1598,1623]" italics="true" pageId="29" pageNumber="30">C. corporaali</emphasis>
</taxonomicName>
) in the general appearance, curved elytral truncation, and markedly widened elytral outer apical angles. But, compared with species of the
<taxonomicName id="9DBBECB12A5CD85AFED07F881D0AB0BA" box="[377,521,1671,1695]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="terminata">
<emphasis id="68CF4B202A5CD85AFED07F881D0AB0BA" box="[377,521,1671,1695]" italics="true" pageId="29" pageNumber="30">C. terminata</emphasis>
</taxonomicName>
group, it is markedly different in several morphological characters, such as the outer apical angles of elytra widened but not acutely toothed, the abdominal sternum VII in males only moderately notched, and the median lobe of aedeagus without prominent ventral lobe and with apical lamina axe-shaped.
</paragraph>
<paragraph id="5A0497322A5CD85AFF3E7FFD1A6EB12E" blockId="29.[151,1437,1274,1803]" box="[151,1389,1778,1803]" pageId="29" pageNumber="30">
Therefore we isolate this new species in a separate group, related to the
<taxonomicName id="9DBBECB12A5CD85AFC157FFC1B49B12E" box="[956,1098,1779,1803]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="terminata">
<emphasis id="68CF4B202A5CD85AFC157FFC1B49B12E" box="[956,1098,1779,1803]" italics="true" pageId="29" pageNumber="30">C. terminata</emphasis>
</taxonomicName>
or
<taxonomicName id="9DBBECB12A5CD85AFBC67FFC1A10B12E" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[1135,1299,1778,1803]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="splendidula">
<emphasis id="68CF4B202A5CD85AFBC67FFC1A10B12E" box="[1135,1299,1778,1803]" italics="true" pageId="29" pageNumber="30">C. splendidula</emphasis>
</taxonomicName>
groups.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,351 @@
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<mods:title id="E201538399EC63409168D5D8B9C03F4B">Revision of the Oriental species of Calleida Latreille (sensu lato). Part 1: Introduction, groups of species, and species of six species groups (Coleoptera: Carabidae: Lebiini)</mods:title>
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<emphasis id="68CF4B202A5CD85AFF3E7E5B1DA9B14B" bold="true" box="[151,682,1876,1902]" pageId="29" pageNumber="30">
[9]
<taxonomicName id="9DBBECB12A5CD85AFF697E5B1D58B14B" ID-CoL="8M5HW" authority="Shi &amp; Casale" authorityName="Shi &amp; Casale" box="[192,603,1876,1902]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="borneensis" status="sp. nov.">
<emphasis id="68CF4B202A5CD85AFF697E5B1EACB14B" bold="true" box="[192,431,1876,1902]" italics="true" pageId="29" pageNumber="30">Calleida borneensis</emphasis>
Shi &amp; Casale
</taxonomicName>
,
<taxonomicNameLabel id="73FCF65B2A5CD85AFDCE7E5B1DA9B14B" box="[615,682,1876,1902]" pageId="29" pageNumber="30" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A5CD859FF3E7E951A43B738" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="5A0497322A5CD85AFF3E7E951D44B196" blockId="29.[151,1436,1946,2007]" box="[151,583,1946,1971]" pageId="29" pageNumber="30">
<emphasis id="68CF4B202A5CD85AFF3E7E951E3AB196" bold="true" box="[151,313,1946,1971]" pageId="29" pageNumber="30">
<typeStatus id="850029902A5CD85AFF3E7E951FD1B196" box="[151,210,1946,1971]" pageId="29" pageNumber="30">Type</typeStatus>
locality:
</emphasis>
Borneo:
<collectingCountry id="22ACD7A22A5CD85AFE087E951D07B196" box="[417,516,1946,1971]" name="Malaysia" pageId="29" pageNumber="30">
<collectingRegion id="987F59D02A5CD85AFE087E951D07B196" box="[417,516,1946,1971]" country="Malaysia" name="Sarawak" pageId="29" pageNumber="30">Sarawak</collectingRegion>
</collectingCountry>
, Bidi.
</paragraph>
<paragraph id="5A0497322A5CD85AFF6E7EB11A9FB1F2" blockId="29.[151,1436,1946,2007]" box="[199,1436,1982,2007]" pageId="29" pageNumber="30">
<emphasis id="68CF4B202A5CD85AFF6E7EB11E7BB1F2" bold="true" box="[199,376,1982,2007]" pageId="29" pageNumber="30">
<typeStatus id="850029902A5CD85AFF6E7EB11E01B1F2" box="[199,258,1982,2007]" pageId="29" pageNumber="30">Type</typeStatus>
material:
</emphasis>
<materialsCitation id="EAD39D6F2A5CD85AFE297EB11A9FB1F2" ID-GBIF-Occurrence="1852047368" box="[384,1436,1982,2007]" collectingMethod="light trap" collectionCode="CAS" elevation="1182" latitude="5.443" location="Borneo" longLatPrecision="7" longitude="116.4512" municipality="Keningau district" pageId="29" pageNumber="30" specimenCode="2 016.IV.24N" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus id="850029902A5CD85AFE297EB11EE8B1F2" box="[384,491,1982,2007]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
,
<specimenCount id="4CBD5CBB2A5CD85AFE517EB11D32B1F2" box="[504,561,1982,2007]" pageId="29" pageNumber="30" type="male">male</specimenCount>
, &quot;
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:
<collectingRegion id="987F59D02A5CD85AFD027EB11DF1B1F2" box="[683,754,1982,2007]" country="Malaysia" name="Sabah" pageId="29" pageNumber="30">Sabah</collectingRegion>
,
<collectingMunicipality id="BA600D482A5CD85AFD567EB11CC6B1F2" box="[767,965,1982,2007]" pageId="29" pageNumber="30">Keningau district</collectingMunicipality>
,
<location id="5F64C1E92A5CD85AFC787EB11B9DB1F2" LSID="urn:lsid:plazi:treatment:D21226242A5CD867FF3E7E5B1B0BB7F5:5F64C1E92A5CD85AFC787EB11B9DB1F2" box="[977,1182,1982,2007]" latitude="5.443" longLatPrecision="7" longitude="116.4512" municipality="Keningau district" name="Jungle Girl Camp" pageId="29" pageNumber="30" stateProvince="Sabah">Jungle Girl Camp</location>
,
<geoCoordinate id="3F8FF1F52A5CD85AFB037EB11A0CB1F2" box="[1194,1295,1982,2007]" direction="north" orientation="latitude" pageId="29" pageNumber="30" precision="5" value="5.443">N5.4430</geoCoordinate>
,
<geoCoordinate id="3F8FF1F52A5CD85AFAB27EB11A9BB1F2" box="[1307,1432,1982,2007]" direction="east" orientation="longitude" pageId="29" pageNumber="30" precision="5" value="116.4512">E116.4512</geoCoordinate>
;
</materialsCitation>
</paragraph>
<paragraph id="5A0497322A5FD859FF3E79971A43B738" blockId="30.[151,1437,151,1977]" pageId="30" pageNumber="31">
<quantity id="9D433AD72A5FD859FF3E79971FE4B695" box="[151,231,152,176]" metricMagnitude="3" metricUnit="m" metricValue="1.182" pageId="30" pageNumber="31" unit="m" value="1182.0">
<elevation id="D19670012A5FD859FF3E79971FE4B695" box="[151,231,152,176]" metricMagnitude="3" metricUnit="m" metricValue="1.182" pageId="30" pageNumber="31" unit="m" value="1182.0">1182m</elevation>
</quantity>
;
<location id="5F64C1E92A5FD859FF5A79981EBEB68A" LSID="urn:lsid:plazi:treatment:D21226242A5CD867FF3E7E5B1B0BB7F5:5F64C1E92A5FD859FF5A79981EBEB68A" box="[243,445,151,176]" latitude="5.443" longLatPrecision="7" longitude="116.4512" municipality="Keningau district" name="Shi H. L. &amp; Liu Y." pageId="30" pageNumber="31" stateProvince="Sabah">Shi H.L. &amp; Liu Y.</location>
lgt.,
<collectingMethod id="83FAEF252A5FD859FE5E79981D5CB68A" box="[503,607,151,176]" pageId="30" pageNumber="31">light trap</collectingMethod>
, Ins. Zoo.,
<collectionCode id="3CAA0FF72A5FD859FD7779971C1BB695" box="[734,792,152,176]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="30" pageNumber="31">CAS</collectionCode>
.
<specimenCode id="0A1D3F492A5FD859FC8A79971CB6B695" box="[803,949,151,176]" pageId="30" pageNumber="31">2 016.IV.24N</specimenCode>
&quot; (NHML,
<figureCitation id="C2808BB72A5FD859FB9979981B7FB695" box="[1072,1148,151,176]" captionStart="FIGURES 6166" captionStartId="31.[151,264,1853,1875]" captionTargetBox="[159,1432,203,1814]" captionTargetId="figure@31.[151,1436,193,1832]" captionTargetPageId="31" captionText="FIGURES 6166. Calleida borneensis sp. n. 61 habitus and labels of holotype (male, Sabah, Malaysia, NHML), scale bar = 2.0 mm; 62 habitus of paratype (male, Sarawak, Malaysia, NHML), scale bar = 2.0 mm; 63 female reproductive tract (Sabah, Malaysia), scale bar = 0.5 mm; abbreviations: sp spermatheca, pd spermathecal pedicel, gd spermathecal gland duct, ga glandular area, at atrium, bpj spermathecal basal projection, apt pedicel apical protuberance; 64 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm; 65 left and right parameres of aedeagus (holotype), scale bar = 0.2 mm; 66 ovipositor (Sabah, Malaysia), scale bar = 0.2 mm." httpUri="https://zenodo.org/record/1302741/files/figure.png" pageId="30" pageNumber="31">fig. 61</figureCitation>
).
<typeStatus id="850029902A5FD859FB3879971A01B695" box="[1169,1282,152,176]" pageId="30" pageNumber="31" type="paratype">Paratypes</typeStatus>
: two females, same data as
<typeStatus id="850029902A5FD859FE9679B31EA2B6F0" box="[319,417,188,213]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
but date 2016.
<date id="2E05B1F22A5FD859FDF279B31D9BB6F0" box="[603,664,188,213]" pageId="30" pageNumber="31" value="1925-04">IV.25</date>
N and 2016.
<date id="2E05B1F22A5FD859FC8779B31C6FB6F0" box="[814,876,188,213]" pageId="30" pageNumber="31" value="1929-04">IV.29</date>
N (
<collectionCode id="3CAA0FF72A5FD859FC3E79B31CD8B6F1" box="[919,987,188,212]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="30" pageNumber="31">IZAS</collectionCode>
); one teneral male, &quot; Borneo:
<collectingRegion id="987F59D02A5FD859FAF879B31A9BB6F0" box="[1361,1432,188,213]" country="Malaysia" name="Sabah" pageId="30" pageNumber="31">Sabah</collectingRegion>
, Keningau district, Jungle Girl Camp,
<geoCoordinate id="3F8FF1F52A5FD859FDE779D01DB2B6DD" box="[590,689,223,248]" direction="north" orientation="latitude" pageId="30" pageNumber="31" precision="5" value="5.443">N5.4430</geoCoordinate>
,
<geoCoordinate id="3F8FF1F52A5FD859FD6879D01C3EB6DD" box="[705,829,223,248]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="5" value="116.4512">E116.4512</geoCoordinate>
;
<quantity id="9D433AD72A5FD859FCE579EF1C9FB6DD" box="[844,924,224,248]" metricMagnitude="3" metricUnit="m" metricValue="1.182" pageId="30" pageNumber="31" unit="m" value="1182.0">1182m</quantity>
; Yuan F. lgt., Malaise trap, Ins. Zoo.,
<collectionCode id="3CAA0FF72A5FD859FAF679EF1A9AB6DD" box="[1375,1433,224,248]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="30" pageNumber="31">CAS</collectionCode>
. 2016.
<date id="2E05B1F22A5FD859FF7F780B1E16B738" box="[214,277,260,285]" pageId="30" pageNumber="31" value="1925-04">IV.25</date>
&quot; (
<collectionCode id="3CAA0FF72A5FD859FE84780B1E72B739" box="[301,369,260,284]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="30" pageNumber="31">IZAS</collectionCode>
); one male, “
<collectingCountry id="22ACD7A22A5FD859FDAA780B1D69B738" box="[515,618,260,285]" name="Malaysia" pageId="30" pageNumber="31">
<collectingRegion id="987F59D02A5FD859FDAA780B1D69B738" box="[515,618,260,285]" country="Malaysia" name="Sarawak" pageId="30" pageNumber="31">Sarawak</collectingRegion>
</collectingCountry>
: Bidi
<date id="2E05B1F22A5FD859FD05780B1C26B738" box="[684,805,260,285]" pageId="30" pageNumber="31" value="1909-03-23">23.iii.1909</date>
C.J. Brooks. B.M. 1936-681” (
<collectionCode id="3CAA0FF72A5FD859FB2D780B1BD8B739" box="[1156,1243,260,284]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="30" pageNumber="31">NHML</collectionCode>
,
<figureCitation id="C2808BB72A5FD859FB4E780B1A30B738" box="[1255,1331,260,285]" captionStart="FIGURES 6166" captionStartId="31.[151,264,1853,1875]" captionTargetBox="[159,1432,203,1814]" captionTargetId="figure@31.[151,1436,193,1832]" captionTargetPageId="31" captionText="FIGURES 6166. Calleida borneensis sp. n. 61 habitus and labels of holotype (male, Sabah, Malaysia, NHML), scale bar = 2.0 mm; 62 habitus of paratype (male, Sarawak, Malaysia, NHML), scale bar = 2.0 mm; 63 female reproductive tract (Sabah, Malaysia), scale bar = 0.5 mm; abbreviations: sp spermatheca, pd spermathecal pedicel, gd spermathecal gland duct, ga glandular area, at atrium, bpj spermathecal basal projection, apt pedicel apical protuberance; 64 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm; 65 left and right parameres of aedeagus (holotype), scale bar = 0.2 mm; 66 ovipositor (Sabah, Malaysia), scale bar = 0.2 mm." httpUri="https://zenodo.org/record/1302741/files/figure.png" pageId="30" pageNumber="31">fig. 62</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A5FD859FF6E78281B9CB71A" box="[199,1183,295,320]" pageId="30" pageNumber="31" type="etymology">
<paragraph id="5A0497322A5FD859FF6E78281B9CB71A" blockId="30.[151,1437,151,1977]" box="[199,1183,295,320]" pageId="30" pageNumber="31">
<emphasis id="68CF4B202A5FD859FF6E78281E86B765" bold="true" box="[199,389,295,320]" pageId="30" pageNumber="31">Specific epithet.</emphasis>
This new species is named from its
<typeStatus id="850029902A5FD859FCB578271C4FB765" box="[796,844,296,320]" pageId="30" pageNumber="31">type</typeStatus>
locality, the island of Borneo.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A5FD859FF6E78431E90B4D5" pageId="30" pageNumber="31" type="diagnosis">
<paragraph id="5A0497322A5FD859FF6E78431BA8B43D" blockId="30.[151,1437,151,1977]" pageId="30" pageNumber="31">
<emphasis id="68CF4B202A5FD859FF6E78431E42B740" bold="true" box="[199,321,332,357]" pageId="30" pageNumber="31">Diagnosis.</emphasis>
The new species is distinct from all the other known Asiatic
<taxonomicName id="9DBBECB12A5FD859FB9178431B9AB740" box="[1080,1177,332,357]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A5FD859FB9178431B9AB740" box="[1080,1177,332,357]" italics="true" pageId="30" pageNumber="31">Calleida</emphasis>
</taxonomicName>
species based on the following character combination: medium sized (L:
<quantity id="9D433AD72A5FD859FD76787F1C69B7AD" box="[735,874,368,392]" metricMagnitude="-3" metricUnit="m" metricValue="9.15" metricValueMax="9.9" metricValueMin="8.4" pageId="30" pageNumber="31" unit="mm" value="9.15" valueMax="9.9" valueMin="8.4">8.49.9 mm</quantity>
). Head and pronotum piceous or dark brown, with very faint greenish reflections; elytra metallic green, with reddish purple reflections. Elytral apex strongly curved, outer apical angles markedly widened, but not forming acute angles (
<figureCitation id="C2808BB72A5FD859FC1678B81B0CB7F5" box="[959,1039,439,464]" captionStart="FIGURES 118" captionStartId="8.[151,264,1664,1686]" captionTargetBox="[216,1350,195,1626]" captionTargetId="figure@8.[216,1350,195,1626]" captionTargetPageId="8" captionText="FIGURES 118. Characters of Calleida spp. 13 sternum VII of males: 1 C. cf. splendidula (Guangdong, China), [apex notched, one seta on each side]; 2 C. fukiensis (Fujian, China), [apex notched, two setae on each side]; 3 C. puncticollis sp. n. (Holotype, Hubei, China), [apex straight, four setae on each side]. 46 sternum VII of females: 4 C. cf. splendidula (Guangdong, China), [two setae on each side]; 5 C. fukiensis (Shaanxi, China), [three setae on each side]; 6 C. puncticollis sp. n. (Hubei, China), [six and seven setae on each side]. 79 basal three antennomeres: 7 C. gressittiana sp. n. (Holotype), [with sparse additional setae]; 8 C. excelsa (Yunnan, China), [with dense additional setae]; 9 C. borneensis sp. n. (Sabah, Malaysia), [without additional setae]. 10 elytral shoulder, C. sultana (Yunnan, China), [humeral margins setose]. 1114 pronotal anterior angle: 11 C. gressittiana sp. n. (Holotype), [with erect setae]; 12 C. excelsa (Yunnan, China), [with erect setae]; 13 C. sultana (Yunnan, China), [with erect setae]; 14 C. borneensis sp. n. (Sabah, Malaysia), [without setae]. 1518 elytral outer apical angle: 15 C. corporaali (Sabah, Malaysia), [acute, forming sharp tooth]; 16 C. borneensis sp. n. (Sabah, Malaysia), [markedly widened, not forming tooth]; 17 C. doriae (Yunnan, China), [slightly widened]; 18 C. gressittiana sp. n. (Holotype), [completely round, not widened]." httpUri="https://zenodo.org/record/1302715/files/figure.png" pageId="30" pageNumber="31">fig. 16</figureCitation>
). Abdominal sternum VII in male normally notched at middle, with one seta on each side in males, two setae in females. Median lobe of aedeagus without any prominent ventral lobe, apical lamina elongate, capitate and securiform (
<figureCitation id="C2808BB72A5FD859FBF978F01B9DB43D" box="[1104,1182,511,536]" captionStart="FIGURES 6166" captionStartId="31.[151,264,1853,1875]" captionTargetBox="[159,1432,203,1814]" captionTargetId="figure@31.[151,1436,193,1832]" captionTargetPageId="31" captionText="FIGURES 6166. Calleida borneensis sp. n. 61 habitus and labels of holotype (male, Sabah, Malaysia, NHML), scale bar = 2.0 mm; 62 habitus of paratype (male, Sarawak, Malaysia, NHML), scale bar = 2.0 mm; 63 female reproductive tract (Sabah, Malaysia), scale bar = 0.5 mm; abbreviations: sp spermatheca, pd spermathecal pedicel, gd spermathecal gland duct, ga glandular area, at atrium, bpj spermathecal basal projection, apt pedicel apical protuberance; 64 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm; 65 left and right parameres of aedeagus (holotype), scale bar = 0.2 mm; 66 ovipositor (Sabah, Malaysia), scale bar = 0.2 mm." httpUri="https://zenodo.org/record/1302741/files/figure.png" pageId="30" pageNumber="31">fig. 64</figureCitation>
).
</paragraph>
<paragraph id="5A0497322A5FD859FF6E7B2A1E90B4D5" blockId="30.[151,1437,151,1977]" pageId="30" pageNumber="31">
It is very similar to and sympatric with
<taxonomicName id="9DBBECB12A5FD859FD3C7B2A1B2AB419" authority="Andrewes in Borneo" authorityName="Andrewes in Borneo" authorityYear="1929" box="[661,1065,548,573]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="corporaali">
<emphasis id="68CF4B202A5FD859FD3C7B2A1C33B418" box="[661,816,548,573]" italics="true" pageId="30" pageNumber="31">C. corporaali</emphasis>
Andrewes in Borneo
</taxonomicName>
, but distinguishable by: the size smaller; the elytral microsculpture vanished; the elytral outer apical angles obtuse, not forming sharp angles; the abdominal sternum VII moderately notched; and the very different male genitalia. The new species is also similar in coloration and sympatric with
<taxonomicName id="9DBBECB12A5FD859FDA47B9F1D71B48D" box="[525,626,656,680]" class="Insecta" family="Carabidae" genus="Callida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis id="68CF4B202A5FD859FDA47B9F1D71B48D" box="[525,626,656,680]" italics="true" pageId="30" pageNumber="31">C. tenuis</emphasis>
</taxonomicName>
(
<taxonomicName id="9DBBECB12A5FD859FD2A7B9F1C24B48D" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[643,807,655,680]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="splendidula">
<emphasis id="68CF4B202A5FD859FD2A7B9F1C24B48D" box="[643,807,655,680]" italics="true" pageId="30" pageNumber="31">C. splendidula</emphasis>
</taxonomicName>
group), but distinguishable by: body size larger; elytral microsculpture vanished; elytral apex markedly curved, outer apical angles widened; median lobe of aedeagus with capitate apical lamina.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A5FD859FF6E7BF31C57B19C" pageId="30" pageNumber="31" type="description">
<paragraph id="5A0497322A5FD859FF6E7BF31A3FB531" blockId="30.[151,1437,151,1977]" box="[199,1340,764,789]" pageId="30" pageNumber="31">
<emphasis id="68CF4B202A5FD859FF6E7BF31E5AB530" bold="true" box="[199,345,764,789]" pageId="30" pageNumber="31">Description.</emphasis>
<emphasis id="68CF4B202A5FD859FEF67BF31D18B531" box="[351,539,764,789]" italics="true" pageId="30" pageNumber="31">General features</emphasis>
: (
<figureCitation id="C2808BB72A5FD859FD9B7BF31DB1B530" box="[562,690,764,789]" captionStart="FIGURES 6166" captionStartId="31.[151,264,1853,1875]" captionTargetBox="[159,1432,203,1814]" captionTargetId="figure@31.[151,1436,193,1832]" captionTargetPageId="31" captionText="FIGURES 6166. Calleida borneensis sp. n. 61 habitus and labels of holotype (male, Sabah, Malaysia, NHML), scale bar = 2.0 mm; 62 habitus of paratype (male, Sarawak, Malaysia, NHML), scale bar = 2.0 mm; 63 female reproductive tract (Sabah, Malaysia), scale bar = 0.5 mm; abbreviations: sp spermatheca, pd spermathecal pedicel, gd spermathecal gland duct, ga glandular area, at atrium, bpj spermathecal basal projection, apt pedicel apical protuberance; 64 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm; 65 left and right parameres of aedeagus (holotype), scale bar = 0.2 mm; 66 ovipositor (Sabah, Malaysia), scale bar = 0.2 mm." httpUri="https://zenodo.org/record/1302741/files/figure.png" pageId="30" pageNumber="31">figs. 61, 62</figureCitation>
) Medium sized species: L:
<quantity id="9D433AD72A5FD859FC4E7BF31B71B531" box="[999,1138,764,788]" metricMagnitude="-3" metricUnit="m" metricValue="9.15" metricValueMax="9.9" metricValueMin="8.4" pageId="30" pageNumber="31" unit="mm" value="9.15" valueMax="9.9" valueMin="8.4">8.49.9 mm</quantity>
; TL: 8.0
<quantity id="9D433AD72A5FD859FB767BF31A3BB531" box="[1247,1336,764,788]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="30" pageNumber="31" unit="mm" value="9.4">9.4 mm</quantity>
.
</paragraph>
<paragraph id="5A0497322A5FD859FF6E7A101E7AB5E2" blockId="30.[151,1437,151,1977]" pageId="30" pageNumber="31">
<emphasis id="68CF4B202A5FD859FF6E7A101E15B51D" box="[199,278,799,824]" italics="true" pageId="30" pageNumber="31">Colour</emphasis>
: head, pronotum, femora, tibiae and underside piceous to dark brownish, head with vague reddish patch on vertex, pronotum with very faint metallic greenish reflections; apex of palpi and lateral margins of pronotum testaceous; antennomeres 13 reddish, 411 dark brown, tarsomeres and trochanters reddish brown; elytra bright metallic green, dull on apical two-thirds of inner two intervals, with reddish purple reflections at shoulders, lateral sides and near apex.
</paragraph>
<paragraph id="5A0497322A5FD859FF6E7ADA1E81B235" blockId="30.[151,1437,151,1977]" pageId="30" pageNumber="31">
<emphasis id="68CF4B202A5FD859FF6E7ADA1EF0B5C8" box="[199,499,980,1005]" italics="true" pageId="30" pageNumber="31">Lustre and microsculpture</emphasis>
: meshes of the microsculpture isodiametric, very faint on the head, vanished on pronotum and elytra.
</paragraph>
<paragraph id="5A0497322A5FD859FF6E7D131CF7B27D" blockId="30.[151,1437,151,1977]" pageId="30" pageNumber="31">
<emphasis id="68CF4B202A5FD859FF6E7D131E07B210" box="[199,260,1052,1077]" italics="true" pageId="30" pageNumber="31">Head</emphasis>
wide, flat, smooth; genae markedly constricted to the neck; frontal foveae shallow, sub-striate in front, ending in a rounded depression on each side; eyes prominent, hemispherical.
</paragraph>
<paragraph id="5A0497322A5FD859FF6E7D6A1C05B2CD" blockId="30.[151,1437,151,1977]" pageId="30" pageNumber="31">
<emphasis id="68CF4B202A5FD859FF6E7D6A1E35B259" box="[199,310,1125,1148]" italics="true" pageId="30" pageNumber="31">Pronotum</emphasis>
subquadrate-cordate (ratio PW/PL: 1.061.11); base oblique at sides, wider than anterior margin. Anterior angles rounded; lateral margins reflexed, rounded in front and slightly or markedly sinuate in front of the posterior angles which are rectangular, not projecting laterally. Disc with very faint transverse wrinkles; median furrow deep, reaching the base; basal foveae very deep.
</paragraph>
<paragraph id="5A0497322A5FD859FF6E7DFB1DB5B35D" blockId="30.[151,1437,151,1977]" pageId="30" pageNumber="31">
<emphasis id="68CF4B202A5FD859FF6E7DFB1E0EB328" box="[199,269,1268,1293]" italics="true" pageId="30" pageNumber="31">Elytra</emphasis>
depressed, elongate (ratio EL/EW: 1.691.78), faintly dilated behind. Striae fine, finely but deeply punctate; intervals slightly convex, flattened on disc; interval 8 dilated near apex into a smooth rounded callosity. Elytral apex obliquely truncate, truncation distinctly curved; the outer apical angles markedly widened, obtusely rounded, not forming acute or prominent angles.
</paragraph>
<paragraph id="5A0497322A5FD859FF6E7C8B1D82B3E5" blockId="30.[151,1437,151,1977]" pageId="30" pageNumber="31">
<emphasis id="68CF4B202A5FD859FF6E7C8B1E3FB3B8" box="[199,316,1412,1437]" italics="true" pageId="30" pageNumber="31">Underside</emphasis>
glabrous and shinning. Abdominal sternum VII in male normally notched at middle, with one seta on each side in males; two setae in females.
</paragraph>
<paragraph id="5A0497322A5FD859FF6E7CC31D81B098" blockId="30.[151,1437,151,1977]" pageId="30" pageNumber="31">
<emphasis id="68CF4B202A5FD859FF6E7CC31E6EB3C0" box="[199,365,1484,1509]" italics="true" pageId="30" pageNumber="31">Male genitalia</emphasis>
: as in
<figureCitation id="C2808BB72A5FD859FE127CC31D0AB3C0" box="[443,521,1484,1509]" captionStart="FIGURES 6166" captionStartId="31.[151,264,1853,1875]" captionTargetBox="[159,1432,203,1814]" captionTargetId="figure@31.[151,1436,193,1832]" captionTargetPageId="31" captionText="FIGURES 6166. Calleida borneensis sp. n. 61 habitus and labels of holotype (male, Sabah, Malaysia, NHML), scale bar = 2.0 mm; 62 habitus of paratype (male, Sarawak, Malaysia, NHML), scale bar = 2.0 mm; 63 female reproductive tract (Sabah, Malaysia), scale bar = 0.5 mm; abbreviations: sp spermatheca, pd spermathecal pedicel, gd spermathecal gland duct, ga glandular area, at atrium, bpj spermathecal basal projection, apt pedicel apical protuberance; 64 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm; 65 left and right parameres of aedeagus (holotype), scale bar = 0.2 mm; 66 ovipositor (Sabah, Malaysia), scale bar = 0.2 mm." httpUri="https://zenodo.org/record/1302741/files/figure.png" pageId="30" pageNumber="31">fig. 64</figureCitation>
. Median lobe of aedeagus of peculiar shape: ventral margin markedly sinuate in lateral aspect; in ventral aspect, middle part distinctly dilated, and then gradually narrowed to apex; apical lamina very developed, much longer than the basal width, capitate and axe-shaped in dorsal and ventral aspect, hooked in lateral aspect. Endophallus with two chitinized copulatory pieces close to the left lateral margin of the median lobe, close to each other at base, V-shaped; the ventral one longer, sinuate and acicular at apex, apex adjacent to the base of the apical orifice; the dorsal one shorter and wider. Left paramere dorsally depressed, two times as long as wide; right paramere slightly emarginate. (
<figureCitation id="C2808BB72A5FD859FD867FAB1D79B098" box="[559,634,1700,1725]" captionStart="FIGURES 6166" captionStartId="31.[151,264,1853,1875]" captionTargetBox="[159,1432,203,1814]" captionTargetId="figure@31.[151,1436,193,1832]" captionTargetPageId="31" captionText="FIGURES 6166. Calleida borneensis sp. n. 61 habitus and labels of holotype (male, Sabah, Malaysia, NHML), scale bar = 2.0 mm; 62 habitus of paratype (male, Sarawak, Malaysia, NHML), scale bar = 2.0 mm; 63 female reproductive tract (Sabah, Malaysia), scale bar = 0.5 mm; abbreviations: sp spermatheca, pd spermathecal pedicel, gd spermathecal gland duct, ga glandular area, at atrium, bpj spermathecal basal projection, apt pedicel apical protuberance; 64 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm; 65 left and right parameres of aedeagus (holotype), scale bar = 0.2 mm; 66 ovipositor (Sabah, Malaysia), scale bar = 0.2 mm." httpUri="https://zenodo.org/record/1302741/files/figure.png" pageId="30" pageNumber="31">fig. 65</figureCitation>
)
</paragraph>
<paragraph id="5A0497322A5FD859FF6E7FC81C57B19C" blockId="30.[151,1437,151,1977]" pageId="30" pageNumber="31">
<emphasis id="68CF4B202A5FD859FF6E7FC81E84B0C5" box="[199,391,1735,1760]" italics="true" pageId="30" pageNumber="31">Female genitalia</emphasis>
(reproductive tract
<figureCitation id="C2808BB72A5FD859FDC17FC81DB1B0C5" box="[616,690,1735,1760]" captionStart="FIGURES 6166" captionStartId="31.[151,264,1853,1875]" captionTargetBox="[159,1432,203,1814]" captionTargetId="figure@31.[151,1436,193,1832]" captionTargetPageId="31" captionText="FIGURES 6166. Calleida borneensis sp. n. 61 habitus and labels of holotype (male, Sabah, Malaysia, NHML), scale bar = 2.0 mm; 62 habitus of paratype (male, Sarawak, Malaysia, NHML), scale bar = 2.0 mm; 63 female reproductive tract (Sabah, Malaysia), scale bar = 0.5 mm; abbreviations: sp spermatheca, pd spermathecal pedicel, gd spermathecal gland duct, ga glandular area, at atrium, bpj spermathecal basal projection, apt pedicel apical protuberance; 64 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm; 65 left and right parameres of aedeagus (holotype), scale bar = 0.2 mm; 66 ovipositor (Sabah, Malaysia), scale bar = 0.2 mm." httpUri="https://zenodo.org/record/1302741/files/figure.png" pageId="30" pageNumber="31">fig. 63</figureCitation>
and ovipositor
<figureCitation id="C2808BB72A5FD859FCCC7FC81CB0B0C5" box="[869,947,1735,1760]" captionStart="FIGURES 6166" captionStartId="31.[151,264,1853,1875]" captionTargetBox="[159,1432,203,1814]" captionTargetId="figure@31.[151,1436,193,1832]" captionTargetPageId="31" captionText="FIGURES 6166. Calleida borneensis sp. n. 61 habitus and labels of holotype (male, Sabah, Malaysia, NHML), scale bar = 2.0 mm; 62 habitus of paratype (male, Sarawak, Malaysia, NHML), scale bar = 2.0 mm; 63 female reproductive tract (Sabah, Malaysia), scale bar = 0.5 mm; abbreviations: sp spermatheca, pd spermathecal pedicel, gd spermathecal gland duct, ga glandular area, at atrium, bpj spermathecal basal projection, apt pedicel apical protuberance; 64 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm; 65 left and right parameres of aedeagus (holotype), scale bar = 0.2 mm; 66 ovipositor (Sabah, Malaysia), scale bar = 0.2 mm." httpUri="https://zenodo.org/record/1302741/files/figure.png" pageId="30" pageNumber="31">fig. 66</figureCitation>
): spermatheca digitiform, one third as long as the pedicel; surface very weakly whorled, with large stout basal projection; spermathecal pedicel straight at base, then strongly bisinuate, S-shaped, with a small apical protuberance; spermathecal gland duct about as long as spermatheca, laterally inserted on the basal projection; glandular area slightly thickened, about twice length as gland duct, base with gradually protuberant atrium. Gonocoxite II of ovipositor three times as long as the basal width, slightly narrowed to apex, apex truncate, a little oblique; inner margin setose from the middle to apex; outer margin a little curved; apex with short membranous extension.
</paragraph>
</subSubSection>
<caption id="0EC4C7BA2A5ED858FF3E7E321CF5B1C8" httpUri="https://zenodo.org/record/1302741/files/figure.png" pageId="31" pageNumber="32" startId="31.[151,264,1853,1875]" targetBox="[159,1432,203,1814]" targetPageId="31">
<paragraph id="5A0497322A5ED858FF3E7E321CF5B1C8" blockId="31.[151,1437,1853,2029]" pageId="31" pageNumber="32">
<emphasis id="68CF4B202A5ED858FF3E7E321D28B176" bold="true" box="[151,555,1853,1875]" pageId="31" pageNumber="32">
FIGURES 6166.
<taxonomicName id="9DBBECB12A5ED858FEF57E321D28B176" box="[348,555,1853,1875]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="borneensis" status="sp. nov.">
<emphasis id="68CF4B202A5ED858FEF57E321D28B176" bold="true" box="[348,555,1853,1875]" italics="true" pageId="31" pageNumber="32">Calleida borneensis</emphasis>
</taxonomicName>
</emphasis>
<taxonomicNameLabel id="73FCF65B2A5ED858FD9B7E311D69B176" box="[562,618,1854,1875]" pageId="31" pageNumber="32" rank="species">sp. n.</taxonomicNameLabel>
<emphasis id="68CF4B202A5ED858FDD87E321D88B176" bold="true" box="[625,651,1853,1875]" pageId="31" pageNumber="32">61</emphasis>
habitus and labels of holotype (male, Sabah, Malaysia, NHML), scale bar = 2.0 mm;
<emphasis id="68CF4B202A5ED858FF5A7E531E0FB157" bold="true" box="[243,268,1884,1906]" pageId="31" pageNumber="32">62</emphasis>
habitus of paratype (male, Sarawak, Malaysia, NHML), scale bar = 2.0 mm;
<emphasis id="68CF4B202A5ED858FB817E531B42B157" bold="true" box="[1064,1089,1884,1906]" pageId="31" pageNumber="32">63</emphasis>
female reproductive tract (Sabah, Malaysia), scale bar = 0.5 mm; abbreviations:
<emphasis id="68CF4B202A5ED858FD227E741DA1B1B4" bold="true" box="[651,674,1915,1937]" pageId="31" pageNumber="32">sp</emphasis>
spermatheca,
<emphasis id="68CF4B202A5ED858FC947E741C5AB1B4" bold="true" box="[829,857,1915,1937]" pageId="31" pageNumber="32">pd</emphasis>
spermathecal pedicel,
<emphasis id="68CF4B202A5ED858FBE77E741B6AB1B4" bold="true" box="[1102,1129,1915,1937]" pageId="31" pageNumber="32">gd</emphasis>
spermathecal gland duct,
<emphasis id="68CF4B202A5ED858FA2A7E741A9EB1B4" bold="true" box="[1411,1437,1915,1937]" pageId="31" pageNumber="32">ga</emphasis>
glandular area,
<emphasis id="68CF4B202A5ED858FE907E961E4DB18A" bold="true" box="[313,334,1945,1967]" pageId="31" pageNumber="32">at</emphasis>
atrium,
<emphasis id="68CF4B202A5ED858FE007E961ECEB18A" bold="true" box="[425,461,1945,1967]" pageId="31" pageNumber="32">bpj</emphasis>
spermathecal basal projection,
<emphasis id="68CF4B202A5ED858FCB17E961C38B18A" bold="true" box="[792,827,1945,1967]" pageId="31" pageNumber="32">apt</emphasis>
pedicel apical protuberance;
<emphasis id="68CF4B202A5ED858FBD97E961B89B18A" bold="true" box="[1136,1162,1945,1967]" pageId="31" pageNumber="32">64</emphasis>
median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (holotype), scale bar = 0.5 mm;
<emphasis id="68CF4B202A5ED858FC557EB71B15B1EB" bold="true" box="[1020,1046,1976,1998]" pageId="31" pageNumber="32">65</emphasis>
left and right parameres of aedeagus (holotype), scale bar = 0.2 mm;
<emphasis id="68CF4B202A5ED858FE757ED81EF6B1C8" bold="true" box="[476,501,2007,2029]" pageId="31" pageNumber="32">66</emphasis>
ovipositor (Sabah, Malaysia), scale bar = 0.2 mm.
</paragraph>
</caption>
<subSubSection id="12A1C4B92A61D867FF6E79981C73B6F1" pageId="32" pageNumber="33" type="distribution">
<paragraph id="5A0497322A61D867FF6E79981C73B6F1" blockId="32.[151,1436,151,464]" pageId="32" pageNumber="33">
<emphasis id="68CF4B202A61D867FF6E79981D95B695" bold="true" box="[199,662,151,176]" pageId="32" pageNumber="33">Geographical distribution and habitat.</emphasis>
Only known from Borneo:
<collectingCountry id="22ACD7A22A61D867FC6679981B2CB695" box="[975,1071,151,176]" name="Malaysia" pageId="32" pageNumber="33">
<collectingRegion id="987F59D02A61D867FC6679981B2CB695" box="[975,1071,151,176]" country="Malaysia" name="Sarawak" pageId="32" pageNumber="33">Sarawak</collectingRegion>
</collectingCountry>
and
<collectingRegion id="987F59D02A61D867FBCF79981BAEB695" box="[1126,1197,151,176]" country="Malaysia" name="Sabah" pageId="32" pageNumber="33">Sabah</collectingRegion>
,
<collectingCountry id="22ACD7A22A61D867FB1079981A1CB695" box="[1209,1311,151,176]" name="Malaysia" pageId="32" pageNumber="33">Malaysia</collectingCountry>
(
<figureCitation id="C2808BB72A61D867FA8779971A75B695" box="[1326,1398,151,176]" captionStart="MAP 5" captionStartId="32.[151,209,1330,1352]" captionTargetBox="[264,1323,517,1309]" captionTargetId="figure@32.[264,1323,517,1309]" captionTargetPageId="32" captionText="MAP 5 Distributions for species in the C. borneensis group: C. borneensis sp. n. (red)." httpUri="https://zenodo.org/record/1302743/files/figure.png" pageId="32" pageNumber="33">map 5</figureCitation>
) in tropical rain forest. Some specimens were collected by light trap.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A61D867FF6E79D01B0BB7F5" pageId="32" pageNumber="33" type="discussion">
<paragraph id="5A0497322A61D867FF6E79D01C28B741" blockId="32.[151,1436,151,464]" pageId="32" pageNumber="33">
Remark
<emphasis id="68CF4B202A61D867FEB679D01E32B6DD" bold="true" box="[287,305,223,248]" pageId="32" pageNumber="33">s.</emphasis>
In the original description of
<taxonomicName id="9DBBECB12A61D867FD2779EF1CFDB6DD" authority=", Andrewes (1929)" authorityName=", Andrewes" authorityYear="1929" box="[654,1022,223,248]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="corporaali">
<emphasis id="68CF4B202A61D867FD2779EF1C2AB6DD" box="[654,809,223,248]" italics="true" pageId="32" pageNumber="33">C. corporaali</emphasis>
,
<bibRefCitation id="3E2AEAC32A61D867FC9079D01CFDB6DD" box="[825,1022,223,248]" pageId="32" pageNumber="33" refId="ref28873" refString="Andrewes, H. E. (1929) Papers on Oriental Carabidae. XXII. The Annals and Magazine of Natural History, Series 10, 4 (22), 353 - 371. https: // doi. org / 10.1080 / 00222932908673067" type="journal article">Andrewes (1929)</bibRefCitation>
</taxonomicName>
wrote that he examined one further specimen from
<collectingCountry id="22ACD7A22A61D867FEED780B1EE3B739" box="[324,480,260,285]" name="Malaysia" pageId="32" pageNumber="33">North Borneo</collectingCountry>
, Kudat, “which is smaller in size and narrower, with a touch of deep red at the apex of elytra, so that it approaches more nearly to
<taxonomicName id="9DBBECB12A61D867FDDE78271C07B765" box="[631,772,296,320]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="terminata">
<emphasis id="68CF4B202A61D867FDDE78271C07B765" box="[631,772,296,320]" italics="true" pageId="32" pageNumber="33">C. terminata</emphasis>
</taxonomicName>
”. This specimen was cited (
<collectingRegion id="987F59D02A61D867FB9778281B8BB765" box="[1086,1160,295,320]" country="Malaysia" name="Sabah" pageId="32" pageNumber="33">Sabah</collectingRegion>
: Kudat) as
<taxonomicName id="9DBBECB12A61D867FAAD78271A9FB765" authorityName=", Andrewes" authorityYear="1929" box="[1284,1436,295,320]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="corporaali">
<emphasis id="68CF4B202A61D867FAAD78271A9FB765" box="[1284,1436,295,320]" italics="true" pageId="32" pageNumber="33">C. corporaali</emphasis>
</taxonomicName>
by
<bibRefCitation id="3E2AEAC32A61D867FF1378431E48B740" box="[186,331,332,357]" pageId="32" pageNumber="33" refId="ref30886" refString="Stork, N. (1986) An annotated checklist of the Carabidae (including Cicindelinae, Rhysodinae and Paussinae) recorded from Borneo. In: Occasional Papers on Systematic Entomology. Fol. 2. British Museum (Natural History), London, pp. 1 - 24." type="book chapter">Stork (1986)</bibRefCitation>
. Maybe, it should belong to
<taxonomicName id="9DBBECB12A61D867FD2378421C20B740" box="[650,803,332,357]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis id="68CF4B202A61D867FD2378421C20B740" box="[650,803,332,357]" italics="true" pageId="32" pageNumber="33">C. borneensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="5A0497322A61D867FF6E78601B0BB7F5" blockId="32.[151,1436,151,464]" pageId="32" pageNumber="33">
<bibRefCitation id="3E2AEAC32A61D867FF6E78601E99B7AD" box="[199,410,367,392]" pageId="32" pageNumber="33" refId="ref28994" refString="Andrewes H. E. (1931 a) Carabidae of Mt. Kinabalu North Borneo. Journal of the federated Malay Museums, XVI (3 - 4), 431 - 485." type="book chapter">Andrewes (1931a)</bibRefCitation>
, in his monograph on the
<taxonomicName id="9DBBECB12A61D867FD6C78601C34B7AD" authorityName="" authorityYear="1802" box="[709,823,367,392]" class="Insecta" family="Carabidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
of the Mt. Kinabalu (
<collectingRegion id="987F59D02A61D867FB9878601B78B7AD" box="[1073,1147,367,392]" country="Malaysia" name="Sabah" pageId="32" pageNumber="33">Sabah</collectingRegion>
), didnt cite any
<taxonomicName id="9DBBECB12A61D867FA9578601A9FB7AD" box="[1340,1436,367,392]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A61D867FA9578601A9FB7AD" box="[1340,1436,367,392]" italics="true" pageId="32" pageNumber="33">Calleida</emphasis>
</taxonomicName>
species. But, according to the present contribution, at least four species are distributed in
<collectingRegion id="987F59D02A61D867FB2F789B1BCCB788" box="[1158,1231,404,429]" country="Malaysia" name="Sabah" pageId="32" pageNumber="33">Sabah</collectingRegion>
:
<emphasis id="68CF4B202A61D867FB70789A1E83B7F5" italics="true" pageId="32" pageNumber="33">
<taxonomicName id="9DBBECB12A61D867FB70789A1A75B788" box="[1241,1398,404,429]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="borneensis">C. borneensis</taxonomicName>
,
<taxonomicName id="9DBBECB12A61D867FA2B789A1E11B7F5" authorityName=", Andrewes" authorityYear="1929" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="corporaali">C. corporaali</taxonomicName>
,
<taxonomicName id="9DBBECB12A61D867FEB478B71E83B7F5" box="[285,384,440,464]" class="Insecta" family="Carabidae" genus="Callida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="tenuis">C. tenuis</taxonomicName>
</emphasis>
and
<taxonomicName id="9DBBECB12A61D867FE1F78B71D59B7F5" box="[438,602,440,464]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="gressittiana">
<emphasis id="68CF4B202A61D867FE1F78B71D59B7F5" box="[438,602,440,464]" italics="true" pageId="32" pageNumber="33">C. gressittiana</emphasis>
</taxonomicName>
, two of which are endemic to Borneo.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:identifier id="4CCAD22D0536C5E079BD5B54BD7C8E30" type="CLB-Dataset">29772</mods:identifier>
<mods:identifier id="86135F1FCF9C4A50397C2477AB1BB14B" type="DOI">10.11646/zootaxa.4442.1.1</mods:identifier>
<mods:identifier id="07F1E610850E1EC1A3BD3D52C2CE9C5E" type="GBIF-Dataset">ded8bdb6-f8ba-4e0b-bed1-1ec08963edfd</mods:identifier>
<mods:identifier id="E13496B46EDABEFB8C21357CDBB42BF3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="0320F7C60810CF57FFB326460D2C980C" type="Zenodo-Dep">1302713</mods:identifier>
<mods:identifier id="D60A37621FFAEF5494C58128D21C966C" type="ZooBank">195B0471-553A-4617-B901-E9DBD2323D14</mods:identifier>
</mods:mods>
<subSection id="33348ED92A60D865FF3E7B211FDDB1B0" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="multiple">
<treatment id="D21226242A60D865FF3E7B211FDDB1B0" ID-DOI="http://doi.org/10.5281/zenodo.5950879" ID-GBIF-Taxon="146043498" ID-Zenodo-Dep="5950879" LSID="urn:lsid:plazi:treatment:D21226242A60D865FF3E7B211FDDB1B0" httpUri="http://treatment.plazi.org/id/D21226242A60D865FF3E7B211FDDB1B0" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
<subSubSection id="12A1C4B92A60D866FF3E7B211D1EB46D" box="[151,541,558,584]" pageId="33" pageNumber="34" type="nomenclature">
<paragraph id="5A0497322A60D866FF3E7B211D1EB46D" blockId="33.[151,541,558,584]" box="[151,541,558,584]" pageId="33" pageNumber="34">
<heading id="014C205E2A60D866FF3E7B211D1EB46D" bold="true" box="[151,541,558,584]" fontSize="11" level="1" pageId="33" pageNumber="34" reason="1">
<emphasis id="68CF4B202A60D866FF3E7B211D1EB46D" bold="true" box="[151,541,558,584]" pageId="33" pageNumber="34">
[10]
<taxonomicName id="9DBBECB12A60D866FF667B211D1EB46D" ID-CoL="68TZV" authority="Bates, 1892" authorityName="Bates Lectotypus" authorityYear="1892" box="[207,541,558,584]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="doriae">
<emphasis id="68CF4B202A60D866FF667B211E8BB46D" bold="true" box="[207,392,558,584]" italics="true" pageId="33" pageNumber="34">Calleida doriae</emphasis>
<bibRefCitation id="3E2AEAC32A60D866FE267B211D1EB46D" box="[399,541,558,584]" pageId="33" pageNumber="34" refId="ref29370" refString="Bates, H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of the Carabidae. Annali del Museo Civico di Storia naturale di Genova, 32, 267 - 428." type="journal article">Bates, 1892</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A60D866FF3E7B7A1A42B4AE" box="[151,1345,629,651]" pageId="33" pageNumber="34" type="reference_group">
<paragraph id="5A0497322A60D866FF3E7B7A1A42B4AE" blockId="33.[151,1345,629,651]" box="[151,1345,629,651]" pageId="33" pageNumber="34">
<taxonomicName id="9DBBECB12A60D866FF3E7B7A1EEBB4AE" ID-CoL="68TZV" authority="Bates, 1892: 422" authorityName="Bates Lectotypus" authorityPageNumber="422" authorityYear="1892" box="[151,488,629,651]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="doriae">
<emphasis id="68CF4B202A60D866FF3E7B7A1E35B4AE" box="[151,310,629,651]" italics="true" pageId="33" pageNumber="34">Calleida doriae</emphasis>
<bibRefCitation id="3E2AEAC32A60D866FE947B791EBBB4AE" box="[317,440,629,651]" pageId="33" pageNumber="34" refId="ref29370" refString="Bates, H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of the Carabidae. Annali del Museo Civico di Storia naturale di Genova, 32, 267 - 428." type="journal article">Bates, 1892</bibRefCitation>
: 422
</taxonomicName>
(
<typeStatus id="850029902A60D866FE5D7B7A1D2BB4AE" box="[500,552,629,651]" pageId="33" pageNumber="34">Type</typeStatus>
locality:
<collectingCountry id="22ACD7A22A60D866FD217B791DD2B4AE" box="[648,721,630,651]" name="Myanmar" pageId="33" pageNumber="34">Burma</collectingCountry>
: Bhamo;
<typeStatus id="850029902A60D866FC987B7A1C8DB4AE" box="[817,910,629,651]" pageId="33" pageNumber="34" type="lectotype">lectotype</typeStatus>
deposited in MSNG);
<bibRefCitation id="3E2AEAC32A60D866FBDD7B7A1A09B4AE" box="[1140,1290,629,651]" pageId="33" pageNumber="34" refId="ref30044" refString="Jedlicka, A. (1963) Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae-Brachyninae (Coleoptera, Carabidae). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden, 28, 269 - 579." type="journal article">Jedlička, 1963</bibRefCitation>
: 436.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A60D866FF3E7BB81FE2B175" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="5A0497322A60D866FF3E7BB81CF5B545" blockId="33.[151,1437,695,2017]" pageId="33" pageNumber="34">
<emphasis id="68CF4B202A60D866FF3E7BB81EC5B4F5" bold="true" box="[151,454,695,720]" pageId="33" pageNumber="34">
<typeStatus id="850029902A60D866FF3E7BB81FD1B4F5" box="[151,210,695,720]" pageId="33" pageNumber="34">Type</typeStatus>
material examined
</emphasis>
:
<materialsCitation id="EAD39D6F2A60D866FE727BB71A9FB53D" ID-GBIF-Occurrence="1852047369" collectingDate="1886-06" country="Myanmar" location="Fea" municipality="Fea" pageId="33" pageNumber="34" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
<typeStatus id="850029902A60D866FE727BB71D48B4F5" box="[475,587,696,720]" pageId="33" pageNumber="34" type="lectotype">Lectotype</typeStatus>
designated herein,
<specimenCount id="4CBD5CBB2A60D866FC937BB81C8AB4F5" box="[826,905,695,720]" pageId="33" pageNumber="34" type="female">female</specimenCount>
, “Bhamò
<collectingCountry id="22ACD7A22A60D866FBAE7BB81B6CB4F5" box="[1031,1135,695,720]" name="Myanmar" pageId="33" pageNumber="34">Birmania</collectingCountry>
<collectingMunicipality id="BA600D482A60D866FBD47BB71BA5B4EA" box="[1149,1190,696,719]" pageId="33" pageNumber="34">Fea</collectingMunicipality>
<collectingDate id="3E41481A2A60D866FB1A7BB71A18B4F5" box="[1203,1307,695,720]" pageId="33" pageNumber="34" value="1886-06">
<date id="2E05B1F22A60D866FB1A7BB71A18B4F5" box="[1203,1307,695,720]" pageId="33" pageNumber="34" value="1886-06">VI 1886</date>
</collectingDate>
”, “Typus”, “
<taxonomicName id="9DBBECB12A60D866FF087BD31EA6B4D1" authority="Doriae Bates" authorityName="Doriae Bates" box="[161,421,732,757]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">Calleida Doriae Bates</taxonomicName>
” (handwritten by Bates), “
<taxonomicName id="9DBBECB12A60D866FD4D7BD31C81B4D1" authority="Bates" authorityName="Bates Lectotypus" authorityYear="1892" box="[740,898,732,757]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="38" phylum="Arthropoda" rank="species" species="doriae">Doriae Bates</taxonomicName>
”, “
<taxonomicName id="9DBBECB12A60D866FC0E7BD31CD8B4D0" box="[935,987,732,757]" class="Insecta" family="Cerambycidae" genus="Call" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="21" phylum="Arthropoda" rank="genus">Call</taxonomicName>
. Doriae typus! Bates”, “Syntypus ♀”, “
<taxonomicName id="9DBBECB12A60D866FF087BF01D26B53D" authority="Bates Lectotypus" authorityName="Bates Lectotypus" authorityYear="1892" box="[161,549,767,792]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="doriae">Calleida doriae Bates Lectotypus</taxonomicName>
♀ Casale &amp; Shi des. 2014”, “Museo Civico di Genova” (MSNG,
<figureCitation id="C2808BB72A60D866FA977BF01A8CB53D" box="[1342,1423,767,792]" captionStart="FIGURES 6776" captionStartId="36.[151,264,213,235]" captionText="FIGURES 6776. Calleida doriae Bates. 67 habitus and labels of lectotype (female, Bhamo, Myanmar, MSNG), scale bar = 2.0 mm; 68 habitus of paralectotype (male, Assam, India, MNHN), scale bar = 2.0 mm; 69 habitus (male, Yunnan, China), scale bar = 2.0 mm; 70 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (Yunnan, China), scale bar = 0.5 mm; 71 left and right parameres of aedeagus (Yunnan, China), scale bar = 0.2 mm; 72 male sternum IX (Yunnan, China), scale bar = 0.5 mm; 73 ovipositor (Yunnan, China), scale bar = 0.2 mm; 74 female reproductive tract (Yunnan, China), scale bar = 0.5 mm; 75 median lobe of aedeagus, ventral view (paralectotype, Assam), scale bar = 0.5 mm; 76 median lobe of aedeagus, ventral view (Guangxi, China), scale bar = 0.5 mm." pageId="33" pageNumber="34">fig. 67</figureCitation>
).
</materialsCitation>
<typeStatus id="850029902A60D866FF3E7A2B1E3CB518" box="[151,319,804,829]" pageId="33" pageNumber="34" type="paralectotype">Paralectotypes</typeStatus>
: one male, &quot;Nandehing Valley, Assam'', &quot;
<taxonomicName id="9DBBECB12A60D866FCB87A2B1B06B519" authority="Bates" authorityName="Bates Lectotypus" authorityYear="1892" box="[785,1029,804,829]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="doriae">Calleida doriae Bates</taxonomicName>
&quot;, &quot;Ex. Musaeo
<bibRefCitation id="3E2AEAC32A60D866FB6B7A2A1A8DB519" box="[1218,1422,804,828]" pageId="33" pageNumber="34" refId="ref29370" refString="Bates, H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of the Carabidae. Annali del Museo Civico di Storia naturale di Genova, 32, 267 - 428." type="journal article">H.W. Bates, 1892</bibRefCitation>
&quot;, &quot;
<typeStatus id="850029902A60D866FF097A471E02B545" box="[160,257,840,864]" pageId="33" pageNumber="34" type="syntype">Syntype</typeStatus>
,
<taxonomicName id="9DBBECB12A60D866FEA57A481D47B545" authority="Bates, 1892" authorityName="Bates Lectotypus" authorityYear="1892" box="[268,580,839,864]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="doriae">
Calleida doriae
<bibRefCitation id="3E2AEAC32A60D866FE147A471D47B545" box="[445,580,840,864]" pageId="33" pageNumber="34" refId="ref29370" refString="Bates, H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of the Carabidae. Annali del Museo Civico di Storia naturale di Genova, 32, 267 - 428." type="journal article">Bates, 1892</bibRefCitation>
</taxonomicName>
, det. Shi H.L. 2011&quot; (
<collectionCode id="3CAA0FF72A60D866FC9E7A471C92B545" box="[823,913,840,864]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="33" pageNumber="34">MNHN</collectionCode>
,
<figureCitation id="C2808BB72A60D866FC347A481CE9B545" box="[925,1002,839,864]" captionStart="FIGURES 6776" captionStartId="36.[151,264,213,235]" captionText="FIGURES 6776. Calleida doriae Bates. 67 habitus and labels of lectotype (female, Bhamo, Myanmar, MSNG), scale bar = 2.0 mm; 68 habitus of paralectotype (male, Assam, India, MNHN), scale bar = 2.0 mm; 69 habitus (male, Yunnan, China), scale bar = 2.0 mm; 70 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (Yunnan, China), scale bar = 0.5 mm; 71 left and right parameres of aedeagus (Yunnan, China), scale bar = 0.2 mm; 72 male sternum IX (Yunnan, China), scale bar = 0.5 mm; 73 ovipositor (Yunnan, China), scale bar = 0.2 mm; 74 female reproductive tract (Yunnan, China), scale bar = 0.5 mm; 75 median lobe of aedeagus, ventral view (paralectotype, Assam), scale bar = 0.5 mm; 76 median lobe of aedeagus, ventral view (Guangxi, China), scale bar = 0.5 mm." pageId="33" pageNumber="34">fig. 68</figureCitation>
).
</paragraph>
<paragraph id="5A0497322A60D866FF6E7A631FE2B175" blockId="33.[151,1437,695,2017]" pageId="33" pageNumber="34">
<emphasis id="68CF4B202A60D866FF6E7A631D2BB5A0" bold="true" box="[199,552,876,901]" pageId="33" pageNumber="34">Non-type material examined</emphasis>
:
<emphasis id="68CF4B202A60D866FD977A631D9EB5A0" bold="true" box="[574,669,876,901]" pageId="33" pageNumber="34">Malysia</emphasis>
: one female, “
<collectingCountry id="22ACD7A22A60D866FCFA7A631CB8B5A0" box="[851,955,876,901]" name="Malaysia" pageId="33" pageNumber="34">Malaysia</collectingCountry>
W,
<collectingRegion id="987F59D02A60D866FC5E7A631B39B5A0" box="[1015,1082,876,901]" country="Malaysia" name="Perak" pageId="33" pageNumber="34">Perak</collectingRegion>
, 30 Km SE of Ipoh,
<quantity id="9D433AD72A60D866FAEE7A631A94B5A1" box="[1351,1431,876,900]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="33" pageNumber="34" unit="m" value="900.0">900 m</quantity>
, Cameron
<collectingRegion id="987F59D02A60D866FEAE7A801E7EB58D" box="[263,381,911,936]" country="Czech Republic" name="Vysocina" pageId="33" pageNumber="34">Highlands</collectingRegion>
, Ringlet,
<date id="2E05B1F22A60D866FE477A801DA2B58D" box="[494,673,911,936]" pageId="33" pageNumber="34" value="2001-04-25" valueMax="2001-05-05" valueMin="2001-04-25">25.IV5.V.2001</date>
P. Čechovský” (
<collectingCountry id="22ACD7A22A60D866FCCB7A9F1CA0B58D" box="[866,931,912,936]" name="Chad" pageId="33" pageNumber="34">CAD</collectingCountry>
);
<emphasis id="68CF4B202A60D866FC117A801CF8B58D" bold="true" box="[952,1019,911,936]" pageId="33" pageNumber="34">
<collectingCountry id="22ACD7A22A60D866FC117A801CF5B58D" box="[952,1014,911,936]" name="Laos" pageId="33" pageNumber="34">Laos</collectingCountry>
:
</emphasis>
one male, “
<collectingCountry id="22ACD7A22A60D866FB207A9F1BC6B58D" box="[1161,1221,912,936]" name="Laos" pageId="33" pageNumber="34">Laos</collectingCountry>
: Ban Na Khua, II-
<date id="2E05B1F22A60D866FF3E7ABB1FFCB5E9" box="[151,255,948,972]" pageId="33" pageNumber="34" value="1920-07">VII-1920</date>
R.V. de Salvaza” [Ban Nam Kueung,
<collectingRegion id="987F59D02A60D866FD047ABB1DFAB5E8" box="[685,761,948,973]" country="Laos" name="Bokeo" pageId="33" pageNumber="34">Bokeo</collectingRegion>
] (
<collectionCode id="3CAA0FF72A60D866FCA67ABB1C6BB5E9" box="[783,872,948,972]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="33" pageNumber="34">NHML</collectionCode>
);
<emphasis id="68CF4B202A60D866FCD27ABB1CDCB5E8" bold="true" box="[891,991,948,973]" pageId="33" pageNumber="34">
<collectingCountry id="22ACD7A22A60D866FCD27ABB1CDCB5E8" box="[891,991,948,973]" name="Vietnam" pageId="33" pageNumber="34">Vietnam</collectingCountry>
</emphasis>
: three males and females, “N-Vietnam, Sapa (
<collectingRegion id="987F59D02A60D866FF4D7AD71E48B5D5" box="[228,331,983,1008]" country="Vietnam" name="Lao Cai" pageId="33" pageNumber="34">Lao Cai</collectingRegion>
), 22°20
<geoCoordinate id="3F8FF1F52A60D866FE197AD71D33B5D5" box="[432,560,983,1008]" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="925" value="103.833336">N 103°50</geoCoordinate>
E,
<date id="2E05B1F22A60D866FDFE7AD81C1FB5D5" box="[599,796,983,1008]" pageId="33" pageNumber="34" value="1991-05-25" valueMax="1991-06-10" valueMin="1991-05-25">25.V.-10.VI.1991</date>
, E. Jendek” (
<collectionCode id="3CAA0FF72A60D866FC617AD71B28B5D5" box="[968,1067,984,1008]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="33" pageNumber="34">NHMW</collectionCode>
);
<emphasis id="68CF4B202A60D866FBEC7AD81B8DB5D5" bold="true" box="[1093,1166,983,1008]" pageId="33" pageNumber="34">
<collectingCountry id="22ACD7A22A60D866FBEC7AD81B8DB5D5" box="[1093,1166,983,1008]" name="China" pageId="33" pageNumber="34">China</collectingCountry>
</emphasis>
?one female, &quot;
<collectingRegion id="987F59D02A60D866FA957AD71A94B5D5" box="[1340,1431,984,1008]" country="China" name="Yunnan" pageId="33" pageNumber="34">Yunnan</collectingRegion>
, Yingjiang county, Nabang, power station,
<quantity id="9D433AD72A60D866FDDF7AF31DBAB231" box="[630,697,1020,1044]" metricMagnitude="2" metricUnit="m" metricValue="4.7299999999999995" pageId="33" pageNumber="34" unit="m" value="473.0">473m</quantity>
, 2016.
<date id="2E05B1F22A60D866FCAC7AF31C38B230" box="[773,827,1020,1045]" pageId="33" pageNumber="34" value="1927-05">V.27</date>
, light trap, Yang Xiaodong leg.&quot; (
<collectionCode id="3CAA0FF72A60D866FB157AF31A0EB231" box="[1212,1293,1020,1044]" country="USA" httpUri="http://grbio.org/cool/3dng-x47a" name="Carthage College" pageId="33" pageNumber="34">CCCC</collectionCode>
); two males, one female, &quot;
<collectingCountry id="22ACD7A22A60D866FE857D101E77B21D" box="[300,372,1055,1080]" name="China" pageId="33" pageNumber="34">China</collectingCountry>
,
<collectingRegion id="987F59D02A60D866FE297D2F1ED6B212" box="[384,469,1056,1079]" country="China" name="Yunnan" pageId="33" pageNumber="34">Yunnan</collectingRegion>
Banna, Menglun G213,
<geoCoordinate id="3F8FF1F52A60D866FD597D101C73B21D" box="[752,880,1055,1080]" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="1" value="21.53833">N21.53833</geoCoordinate>
,
<geoCoordinate id="3F8FF1F52A60D866FCD57D2F1B04B21D" box="[892,1031,1056,1080]" direction="east" orientation="longitude" pageId="33" pageNumber="34" precision="1" value="101.17001">E101.17001</geoCoordinate>
, 2009.
<date id="2E05B1F22A60D866FBFD7D101B96B21D" box="[1108,1173,1055,1080]" pageId="33" pageNumber="34" value="1925-11">
XI.
<quantity id="9D433AD72A60D866FBD17D101B96B21D" box="[1144,1173,1055,1080]" metricMagnitude="4" metricUnit="m" metricValue="2.5618" pageId="33" pageNumber="34" unit="m" value="25618.0">25</quantity>
</date>
, 618m, Tang Guo, Yao Zhiyuan leg&quot; (
<collectionCode id="3CAA0FF72A60D866FE957D4B1E83B279" box="[316,384,1092,1116]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="33" pageNumber="34">IZAS</collectionCode>
); one female, &quot;
<collectingRegion id="987F59D02A60D866FD847D4B1D8BB279" box="[557,648,1092,1116]" country="China" name="Yunnan" pageId="33" pageNumber="34">Yunnan</collectingRegion>
, Xishuangbanna, Xiaomengyang,
<quantity id="9D433AD72A60D866FBA67D4B1B52B278" box="[1039,1105,1092,1117]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="33" pageNumber="34" unit="m" value="850.0">850m</quantity>
, 1957.
<date id="2E05B1F22A60D866FB097D4B1BE2B278" box="[1184,1249,1092,1117]" pageId="33" pageNumber="34" value="2017-06">VI.17</date>
, Wang Shuyong leg.&quot; (
<collectionCode id="3CAA0FF72A60D866FF727D671E1CB2A5" box="[219,287,1128,1152]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="33" pageNumber="34">IZAS</collectionCode>
); one female, &quot;
<collectingRegion id="987F59D02A60D866FE6D7D671D1CB2A5" box="[452,543,1128,1152]" country="China" name="Yunnan" pageId="33" pageNumber="34">Yunnan</collectingRegion>
, Jingdong county, Jinping town, Liyue,
<quantity id="9D433AD72A60D866FC497D681B33B2A5" box="[992,1072,1127,1152]" metricMagnitude="3" metricUnit="m" metricValue="1.63" pageId="33" pageNumber="34" unit="m" value="1630.0">1630m</quantity>
, 2001.
<date id="2E05B1F22A60D866FBD57D681BBEB2A5" box="[1148,1213,1127,1152]" pageId="33" pageNumber="34" value="2016-11">XI.16</date>
, Ou Xiaohong leg.&quot; (
<collectionCode id="3CAA0FF72A60D866FF367D831FF2B281" box="[159,241,1164,1188]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14107" name="Southwest Forestry College" pageId="33" pageNumber="34">SWFC</collectionCode>
); one male, &quot;
<collectingRegion id="987F59D02A60D866FE3E7D831EF1B281" box="[407,498,1164,1188]" country="China" name="Yunnan" pageId="33" pageNumber="34">Yunnan</collectingRegion>
, Longling county, Banglazhang,
<quantity id="9D433AD72A60D866FC2B7D831B11B280" box="[898,1042,1164,1189]" metricMagnitude="3" metricUnit="m" metricValue="1.395" metricValueMax="1.6" metricValueMin="1.19" pageId="33" pageNumber="34" unit="m" value="1395.0" valueMax="1600.0" valueMin="1190.0">1190-1600m</quantity>
, 2002.
<date id="2E05B1F22A60D866FBCF7D831BA0B281" box="[1126,1187,1164,1188]" pageId="33" pageNumber="34" value="1918-04">IV.18</date>
, Situ Yingxian leg.&quot; (
<collectionCode id="3CAA0FF72A60D866FF367DBF1FF2B2ED" box="[159,241,1200,1224]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14107" name="Southwest Forestry College" pageId="33" pageNumber="34">SWFC</collectionCode>
); one male, &quot;
<collectingRegion id="987F59D02A60D866FE297DBF1ED8B2ED" box="[384,475,1200,1224]" country="China" name="Yunnan" pageId="33" pageNumber="34">Yunnan</collectingRegion>
, Hekou county, Huayudong, 2009.
<date id="2E05B1F22A60D866FCC87DBF1CC6B2ED" box="[865,965,1200,1224]" pageId="33" pageNumber="34" value="1920-04-21">IV.20-21</date>
, Zhang Weiwei leg.&quot; (
<collectionCode id="3CAA0FF72A60D866FB177DBF1A1EB2ED" box="[1214,1309,1200,1224]" pageId="33" pageNumber="34">ZWWC</collectionCode>
); one male, &quot;
<collectingRegion id="987F59D02A60D866FF097DDB1E05B2C8" box="[160,262,1236,1261]" country="China" name="Guangxi" pageId="33" pageNumber="34">Guangxi</collectingRegion>
, Jinxiu county, Luoxiang,
<quantity id="9D433AD72A60D866FD957DDB1D7CB2C9" box="[572,639,1236,1260]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="33" pageNumber="34" unit="m" value="200.0">200m</quantity>
, 1999.
<date id="2E05B1F22A60D866FD677DDB1C00B2C8" box="[718,771,1236,1261]" pageId="33" pageNumber="34" value="2015-05">V.15</date>
, Zhang Yanzhou leg.&quot; (
<collectionCode id="3CAA0FF72A60D866FBB17DDB1B5FB2C9" box="[1048,1116,1236,1260]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="33" pageNumber="34">IZAS</collectionCode>
); one male, &quot;
<collectingRegion id="987F59D02A60D866FB5E7DDB1A54B2C8" box="[1271,1367,1236,1261]" country="China" name="Sichuan" pageId="33" pageNumber="34">Sichuan</collectingRegion>
, Miyi county, 1958.
<date id="2E05B1F22A60D866FE997DF81E7FB335" box="[304,380,1271,1296]" pageId="33" pageNumber="34" value="1920-07">VII.20</date>
&quot; (
<collectionCode id="3CAA0FF72A60D866FE3E7DF71ED8B335" box="[407,475,1272,1296]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="33" pageNumber="34">IZAS</collectionCode>
); one male, &quot;
<collectingRegion id="987F59D02A60D866FDDA7DF81DCAB335" box="[627,713,1271,1296]" country="China" name="Xizang" pageId="33" pageNumber="34">Xizang</collectingRegion>
, way between Yarang to Medog, 2006.
<date id="2E05B1F22A60D866FB247DF81BE2B335" box="[1165,1249,1271,1296]" pageId="33" pageNumber="34" value="1920-08">VIII.20</date>
, Liang Hongbin leg.&quot; (
<collectionCode id="3CAA0FF72A60D866FF727C131E1CB311" box="[219,287,1308,1332]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="33" pageNumber="34">IZAS</collectionCode>
);
<emphasis id="68CF4B202A60D866FE9D7C131E7EB310" bold="true" box="[308,381,1308,1333]" pageId="33" pageNumber="34">
<collectingCountry id="22ACD7A22A60D866FE9D7C131E7BB310" box="[308,376,1308,1333]" name="India" pageId="33" pageNumber="34">India</collectingCountry>
:
</emphasis>
one female, “NE
<collectingCountry id="22ACD7A22A60D866FDE37C131D85B310" box="[586,646,1308,1333]" name="India" pageId="33" pageNumber="34">India</collectingCountry>
,
<collectingRegion id="987F59D02A60D866FD3B7C131C12B310" box="[658,785,1308,1333]" country="India" name="Meghalaya" pageId="33" pageNumber="34">Meghalaya</collectingRegion>
,
<quantity id="9D433AD72A60D866FCB77C131C54B310" box="[798,855,1308,1333]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="33" pageNumber="34" unit="km" value="1.0">1 km</quantity>
E Tura,
<quantity id="9D433AD72A60D866FC127C131B06B311" box="[955,1029,1308,1333]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="33" pageNumber="34" unit="m" value="550.0">550 m</quantity>
, 3.
<date id="2E05B1F22A60D866FB8B7C131B83B310" box="[1058,1152,1308,1333]" pageId="33" pageNumber="34" value="1925-05-02">V.02 25</date>
° 30N, 90° 14E, Tryzna leg.”;
<specimenCount id="4CBD5CBB2A60D866FF727C301E3CB37D" box="[219,319,1343,1368]" pageId="33" pageNumber="34" type="male">25 males</specimenCount>
and females, “
<collectingCountry id="22ACD7A22A60D866FE4C7C301D22B37D" box="[485,545,1343,1368]" name="India" pageId="33" pageNumber="34">India</collectingCountry>
W. Bengal, Bhakta B., Jalpaiguri Distr.
<date id="2E05B1F22A60D866FC4B7C301B69B37D" box="[994,1130,1343,1368]" pageId="33" pageNumber="34" value="1990-04-11" valueMax="1990-04-15" valueMin="1990-04-11">11-15.IV.90</date>
”; “
<collectingCountry id="22ACD7A22A60D866FB257C301BD5B37D" box="[1164,1238,1343,1368]" name="India" pageId="33" pageNumber="34">Indien</collectingCountry>
Darjeeling D Ch. J. Rai Kalimpong
<quantity id="9D433AD72A60D866FECF7C6B1EBFB359" box="[358,444,1380,1404]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="33" pageNumber="34" unit="m" value="1100.0">1100 m</quantity>
Upp. Bombusty
<date id="2E05B1F22A60D866FDD67C6B1DFCB359" box="[639,767,1380,1404]" pageId="33" pageNumber="34" value="1987-04-29">29.IV.1987</date>
”; “
<collectingCountry id="22ACD7A22A60D866FC887C6B1C68B358" box="[801,875,1380,1405]" name="India" pageId="33" pageNumber="34">Indien</collectingCountry>
W Bengal Bhakta B. Jalpaiguri Distr. Doors
<date id="2E05B1F22A60D866FAD87C6B1FF8B385" pageId="33" pageNumber="34" value="1990-04-11" valueMax="1990-04-15" valueMin="1990-04-11">11.- 15.IV.90</date>
”; “
<collectingCountry id="22ACD7A22A60D866FEB77C881E59B385" box="[286,346,1415,1440]" name="India" pageId="33" pageNumber="34">India</collectingCountry>
1979 Darjeeling D. Bhakta B. Monshong
<quantity id="9D433AD72A60D866FCE97C871C95B385" box="[832,918,1416,1440]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="33" pageNumber="34" unit="m" value="1400.0">1400 m</quantity>
<date id="2E05B1F22A60D866FC367C881CCDB385" box="[927,974,1415,1440]" pageId="33" pageNumber="34">5.V.</date>
”; “
<collectingCountry id="22ACD7A22A60D866FC5B7C881B2DB385" box="[1010,1070,1415,1440]" name="India" pageId="33" pageNumber="34">India</collectingCountry>
Darjeeling D. Bhakta B. Chuba
<quantity id="9D433AD72A60D866FF3E7CA31FDCB3E1" box="[151,223,1452,1477]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="33" pageNumber="34" unit="m" value="670.0">670 m</quantity>
<date id="2E05B1F22A60D866FF4F7CA31E77B3E0" box="[230,372,1452,1477]" pageId="33" pageNumber="34" value="1984-07-16">16.VII.1984</date>
”; “Darjeeling D. 79
<collectingCountry id="22ACD7A22A60D866FDF77CA31D94B3E0" box="[606,663,1452,1477]" name="India" pageId="33" pageNumber="34">India</collectingCountry>
Bhakta B. Sombarey
<quantity id="9D433AD72A60D866FC3B7CA31CDAB3E1" box="[914,985,1452,1476]" metricMagnitude="2" metricUnit="m" metricValue="7.18" pageId="33" pageNumber="34" unit="m" value="718.0">718 m</quantity>
<date id="2E05B1F22A60D866FC487CA31B46B3E1" box="[993,1093,1452,1476]" pageId="33" pageNumber="34" value="1979-04-19">19.IV.79</date>
”; “
<collectingCountry id="22ACD7A22A60D866FBCE7CA31BB2B3E0" box="[1127,1201,1452,1477]" name="India" pageId="33" pageNumber="34">Indien</collectingCountry>
Darjeeling D. Ch. J. Rai Kalimpong Tirpai
<date id="2E05B1F22A60D866FE317CDF1EFDB3CD" box="[408,510,1488,1512]" pageId="33" pageNumber="34" value="1987-05-01">1.V.1987</date>
” (
<collectionCode id="3CAA0FF72A60D866FDB07CDF1D72B3CD" box="[537,625,1488,1512]" country="Switzerland" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="33" pageNumber="34">NHMB</collectionCode>
,
<collectionCode id="3CAA0FF72A60D866FDD77CDF1DBFB3CD" box="[638,700,1488,1512]" country="Germany" httpUri="http://grbio.org/cool/mqei-3i36" name="Culture Collection of Algae at the University of Marburg" pageId="33" pageNumber="34">CCA</collectionCode>
); one male, “
<collectingCountry id="22ACD7A22A60D866FCF87CC01C8EB3CD" box="[849,909,1487,1512]" name="India" pageId="33" pageNumber="34">India</collectingCountry>
(
<collectingRegion id="987F59D02A60D866FC347CC01CF3B3CD" box="[925,1008,1487,1512]" country="India" name="Sikkim" pageId="33" pageNumber="34">Sikkim</collectingRegion>
State), N.
<collectingRegion id="987F59D02A60D866FBC27CC01BBCB3CD" box="[1131,1215,1487,1512]" country="India" name="Sikkim" pageId="33" pageNumber="34">Sikkim</collectingRegion>
, Phensang
<quantity id="9D433AD72A60D866FA967CC01A94B3CD" box="[1343,1431,1487,1512]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="33" pageNumber="34" unit="m" value="1500.0">1500 m</quantity>
,
<date id="2E05B1F22A60D866FF3E7CFB1E0AB029" box="[151,265,1524,1548]" pageId="33" pageNumber="34" value="2013-05-10">10.V.2013</date>
K. Krause” (CDW); three ex. &quot;
<collectingCountry id="22ACD7A22A60D866FDC57CFB1DA3B028" box="[620,672,1524,1549]" name="India" pageId="33" pageNumber="34">Inde</collectingCountry>
Anglaise, Pedong, Region de Darjeeling.&quot; (
<collectionCode id="3CAA0FF72A60D866FB397CFB1BEFB029" box="[1168,1260,1524,1548]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="33" pageNumber="34">MNHN</collectionCode>
); 33 exs., &quot;
<collectingCountry id="22ACD7A22A60D866FAC17CFB1A9FB028" box="[1384,1436,1524,1549]" name="India" pageId="33" pageNumber="34">Inde</collectingCountry>
Anglaise, Pedong, Region de Darjeeling., Chasseurs indigenes, 1933.&quot; (
<collectionCode id="3CAA0FF72A60D866FC157F171B1BB015" box="[956,1048,1560,1584]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="33" pageNumber="34">MNHN</collectionCode>
); one ex., &quot;
<collectingCountry id="22ACD7A22A60D866FB3D7F181BCAB015" box="[1172,1225,1559,1584]" name="India" pageId="33" pageNumber="34">Inde</collectingCountry>
Anglaise, Pedong, Region de Darjeeling., Chasseurs indigenes, 1932.&quot; (
<collectionCode id="3CAA0FF72A60D866FCBB7F331C6DB071" box="[786,878,1596,1620]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="33" pageNumber="34">MNHN</collectionCode>
); one ex., &quot;
<collectingCountry id="22ACD7A22A60D866FC577F331B31B070" box="[1022,1074,1596,1621]" name="India" pageId="33" pageNumber="34">Inde</collectingCountry>
Anglaise, Pedong, Region de Darjeeling., Chasseurs indigenes, 1935.&quot; (
<collectionCode id="3CAA0FF72A60D866FDDB7F6F1DCDB05D" box="[626,718,1632,1656]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="33" pageNumber="34">MNHN</collectionCode>
); one ex., &quot;Chota-Nagpore, Falkot, R.P.Cardon, VII-VIII 1897&quot; (
<collectionCode id="3CAA0FF72A60D866FF367F8B1FF8B0B9" box="[159,251,1668,1692]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="33" pageNumber="34">MNHN</collectionCode>
); one ex. &quot;Bom-bay.&quot;, &quot;Janson, Acq. 1884&quot; (
<collectionCode id="3CAA0FF72A60D866FCA07F8B1C66B0B9" box="[777,869,1668,1692]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="33" pageNumber="34">MNHN</collectionCode>
);
<emphasis id="68CF4B202A60D866FCD57F8B1CDDB0B8" bold="true" box="[892,990,1668,1693]" pageId="33" pageNumber="34">
<collectingCountry id="22ACD7A22A60D866FCD57F8B1CDAB0B8" box="[892,985,1668,1693]" name="Bhutan" pageId="33" pageNumber="34">Bhutan</collectingCountry>
:
</emphasis>
one male, &quot;British Bootang, L. Durel 1898&quot; (
<collectionCode id="3CAA0FF72A60D866FF437FA71E45B0E5" box="[234,326,1704,1728]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="33" pageNumber="34">MNHN</collectionCode>
); one ex., &quot;British Bootan, Padong, L.Durel 1914&quot; (
<collectionCode id="3CAA0FF72A60D866FC387FA71CEEB0E5" box="[913,1005,1704,1728]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="33" pageNumber="34">MNHN</collectionCode>
); one ex., &quot;British Bootan, Padong, L. Durel 1913&quot; (
<collectionCode id="3CAA0FF72A60D866FE9A7FC31E8CB0C1" box="[307,399,1740,1764]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="33" pageNumber="34">MNHN</collectionCode>
); one ex., &quot;British Bootang, Sakiou, L. Durel&quot; (
<collectionCode id="3CAA0FF72A60D866FC047FC31B0AB0C1" box="[941,1033,1740,1764]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="33" pageNumber="34">MNHN</collectionCode>
);
<emphasis id="68CF4B202A60D866FBB47FC31B6FB0C0" bold="true" box="[1053,1132,1740,1765]" pageId="33" pageNumber="34">
<collectingCountry id="22ACD7A22A60D866FBB47FC31B64B0C0" box="[1053,1127,1740,1765]" name="Nepal" pageId="33" pageNumber="34">Nepal</collectingCountry>
:
</emphasis>
two ex., “E-Nepal:
<collectingRegion id="987F59D02A60D866FAFB7FC31A9BB0C0" box="[1362,1432,1740,1765]" country="Nepal" name="Kosi" pageId="33" pageNumber="34">Koshi</collectingRegion>
,
<quantity id="9D433AD72A60D866FF3E7FE01E3EB12D" box="[151,317,1775,1800]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="33" pageNumber="34" unit="m" value="1400.0" valueMax="1600.0" valueMin="1200.0">12001600 m</quantity>
, Phulwari-Waku,
<date id="2E05B1F22A60D866FDB17FE01D88B12D" box="[536,651,1775,1800]" pageId="33" pageNumber="34" value="1985-06-09">9.VI.1985</date>
” (
<collectionCode id="3CAA0FF72A60D866FD077FFF1C13B12D" box="[686,784,1776,1800]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="33" pageNumber="34">NHMW</collectionCode>
); one ex., “E-Nepal: Dhakuta,
<date id="2E05B1F22A60D866FB3C7FFE1BF6B12D" box="[1173,1269,1776,1800]" pageId="33" pageNumber="34" value="1983-05">V. 1983</date>
(Holzschuh)” (
<collectionCode id="3CAA0FF72A60D866FF377E1B1E03B109" box="[158,256,1812,1836]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="33" pageNumber="34">NHMW</collectionCode>
); three males and females, “E-Lumbughat-Baiseghat
<quantity id="9D433AD72A60D866FCD97E1B1CB8B109" box="[880,955,1812,1837]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="33" pageNumber="34" unit="m" value="450.0">450 m</quantity>
<date id="2E05B1F22A60D866FC6E7E1B1B29B108" box="[967,1066,1812,1837]" pageId="33" pageNumber="34" value="1985-06-15">15.VI.85</date>
<collectingRegion id="987F59D02A60D866FB9C7E1B1B7BB108" box="[1077,1144,1812,1837]" country="Nepal" name="Kosi" pageId="33" pageNumber="34">Koshi</collectingRegion>
M. Brancucci” (
<collectionCode id="3CAA0FF72A60D866FAE97E1B1A9BB109" box="[1344,1432,1812,1836]" country="Switzerland" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="33" pageNumber="34">NHMB</collectionCode>
,
<collectionCode id="3CAA0FF72A60D866FF3E7E371FD5B175" box="[151,214,1848,1872]" country="Germany" httpUri="http://grbio.org/cool/mqei-3i36" name="Culture Collection of Algae at the University of Marburg" pageId="33" pageNumber="34">CCA</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A60D865FF6E7E531B45B789" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="discussion">
<paragraph id="5A0497322A60D865FF6E7E531EFAB6F1" blockId="33.[151,1437,695,2017]" lastBlockId="34.[151,1437,151,1941]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
<emphasis id="68CF4B202A60D866FF6E7E531EDCB150" bold="true" box="[199,479,1884,1909]" pageId="33" pageNumber="34">
Notes on
<typeStatus id="850029902A60D866FE9E7E531E69B150" box="[311,362,1884,1909]" pageId="33" pageNumber="34">type</typeStatus>
material.
</emphasis>
<bibRefCitation id="3E2AEAC32A60D866FE4E7E521D7BB151" box="[487,632,1884,1908]" pageId="33" pageNumber="34" refId="ref29370" refString="Bates, H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of the Carabidae. Annali del Museo Civico di Storia naturale di Genova, 32, 267 - 428." type="journal article">Bates (1892)</bibRefCitation>
wrote that “a male example of the same fine species was taken by Mr. Chennell at Noa Dehing in
<collectingRegion id="987F59D02A60D866FE7F7E8F1D2BB1BD" box="[470,552,1920,1944]" country="India" name="Assam" pageId="33" pageNumber="34">Assam</collectingRegion>
”. This male
<typeStatus id="850029902A60D866FD137E8E1C12B1BC" box="[698,785,1921,1945]" pageId="33" pageNumber="34" type="syntype">syntype</typeStatus>
(
<figureCitation id="C2808BB72A60D866FC8A7E701C72B1BD" box="[803,881,1919,1944]" captionStart="FIGURES 6776" captionStartId="36.[151,264,213,235]" captionText="FIGURES 6776. Calleida doriae Bates. 67 habitus and labels of lectotype (female, Bhamo, Myanmar, MSNG), scale bar = 2.0 mm; 68 habitus of paralectotype (male, Assam, India, MNHN), scale bar = 2.0 mm; 69 habitus (male, Yunnan, China), scale bar = 2.0 mm; 70 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (Yunnan, China), scale bar = 0.5 mm; 71 left and right parameres of aedeagus (Yunnan, China), scale bar = 0.2 mm; 72 male sternum IX (Yunnan, China), scale bar = 0.5 mm; 73 ovipositor (Yunnan, China), scale bar = 0.2 mm; 74 female reproductive tract (Yunnan, China), scale bar = 0.5 mm; 75 median lobe of aedeagus, ventral view (paralectotype, Assam), scale bar = 0.5 mm; 76 median lobe of aedeagus, ventral view (Guangxi, China), scale bar = 0.5 mm." pageId="33" pageNumber="34">fig. 68</figureCitation>
) has its antennal colour much lighter than other examined specimens of this species, so that is somewhat similar to
<taxonomicName id="9DBBECB12A60D866FC0F7EAA1B0FB199" authorityName="Andr. Cotype H. E Andrewes" authorityYear="1933" box="[934,1036,1957,1980]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="rapax">
<emphasis id="68CF4B202A60D866FC0F7EAA1B0FB199" box="[934,1036,1957,1980]" italics="true" pageId="33" pageNumber="34">C. rapax</emphasis>
</taxonomicName>
from NW
<collectingCountry id="22ACD7A22A60D866FB277EAB1BC4B198" box="[1166,1223,1956,1981]" name="India" pageId="33" pageNumber="34">India</collectingCountry>
(see the key). We attributed it to
<taxonomicName id="9DBBECB12A60D866FEE87EC61EAEB1C4" authorityName="Bates Lectotypus" authorityYear="1892" box="[321,429,1992,2017]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="doriae">
<emphasis id="68CF4B202A60D866FEE87EC61EAEB1C4" box="[321,429,1992,2017]" italics="true" pageId="33" pageNumber="34">C. doriae</emphasis>
</taxonomicName>
, not to
<taxonomicName id="9DBBECB12A60D866FDAE7EC61D68B1C5" authorityName="Andr. Cotype H. E Andrewes" authorityYear="1933" box="[519,619,1993,2016]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="rapax">
<emphasis id="68CF4B202A60D866FDAE7EC61D68B1C5" box="[519,619,1993,2016]" italics="true" pageId="33" pageNumber="34">C. rapax</emphasis>
</taxonomicName>
, owing to: pronotum and tarsomeres distinctly darker than in
<taxonomicName id="9DBBECB12A60D866FA997EC61A97B1C5" authorityName="Andr. Cotype H. E Andrewes" authorityYear="1933" box="[1328,1428,1993,2016]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="rapax">
<emphasis id="68CF4B202A60D866FA997EC61A97B1C5" box="[1328,1428,1993,2016]" italics="true" pageId="33" pageNumber="34">C. rapax</emphasis>
</taxonomicName>
; apical lamina of aedeagus a little longer (W/L: 1.23) than in
<taxonomicName id="9DBBECB12A63D865FCC779971CD6B695" authorityName="Andr. Cotype H. E Andrewes" authorityYear="1933" box="[878,981,152,176]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="rapax">
<emphasis id="68CF4B202A63D865FCC779971CD6B695" box="[878,981,152,176]" italics="true" pageId="34" pageNumber="35">C. rapax</emphasis>
</taxonomicName>
; and distribution range closer to other confirmed records of
<taxonomicName id="9DBBECB12A63D865FE2079B21EF1B6F0" authorityName="Bates Lectotypus" authorityYear="1892" box="[393,498,188,213]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="doriae">
<emphasis id="68CF4B202A63D865FE2079B21EF1B6F0" box="[393,498,188,213]" italics="true" pageId="34" pageNumber="35">C. doriae</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="5A0497322A63D865FF6E79EF1B15B765" blockId="34.[151,1437,151,1941]" pageId="34" pageNumber="35">
Because this male
<typeStatus id="850029902A63D865FE3379EF1EF2B6DD" box="[410,497,224,248]" pageId="34" pageNumber="35" type="syntype">syntype</typeStatus>
does not have characters typical of this species, and
<bibRefCitation id="3E2AEAC32A63D865FBE079EF1BD9B6DD" box="[1097,1242,224,248]" pageId="34" pageNumber="35" refId="ref29370" refString="Bates, H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of the Carabidae. Annali del Museo Civico di Storia naturale di Genova, 32, 267 - 428." type="journal article">Bates (1892)</bibRefCitation>
clearly indicated the specimen collected in
<collectingCountry id="22ACD7A22A63D865FE65780A1D1BB739" box="[460,536,261,284]" name="Myanmar" pageId="34" pageNumber="35">Burma</collectingCountry>
as the primary
<typeStatus id="850029902A63D865FD70780A1C0AB738" box="[729,777,261,285]" pageId="34" pageNumber="35">type</typeStatus>
in a paper dedicated to the
<taxonomicName id="9DBBECB12A63D865FBF7780B1BD3B738" authorityName="" authorityYear="1802" box="[1118,1232,260,285]" class="Insecta" family="Carabidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
from
<collectingCountry id="22ACD7A22A63D865FAB4780A1A6FB739" box="[1309,1388,261,284]" name="Myanmar" pageId="34" pageNumber="35">Burma</collectingCountry>
, we designated here the female
<typeStatus id="850029902A63D865FE6078271D23B765" box="[457,544,296,320]" pageId="34" pageNumber="35" type="syntype">syntype</typeStatus>
from Bhamo (MSNG) as
<typeStatus id="850029902A63D865FCEF78281CAFB765" box="[838,940,295,320]" pageId="34" pageNumber="35" type="lectotype">lectotype</typeStatus>
(
<figureCitation id="C2808BB72A63D865FC1278281B0AB765" box="[955,1033,295,320]" captionStart="FIGURES 6776" captionStartId="36.[151,264,213,235]" captionText="FIGURES 6776. Calleida doriae Bates. 67 habitus and labels of lectotype (female, Bhamo, Myanmar, MSNG), scale bar = 2.0 mm; 68 habitus of paralectotype (male, Assam, India, MNHN), scale bar = 2.0 mm; 69 habitus (male, Yunnan, China), scale bar = 2.0 mm; 70 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (Yunnan, China), scale bar = 0.5 mm; 71 left and right parameres of aedeagus (Yunnan, China), scale bar = 0.2 mm; 72 male sternum IX (Yunnan, China), scale bar = 0.5 mm; 73 ovipositor (Yunnan, China), scale bar = 0.2 mm; 74 female reproductive tract (Yunnan, China), scale bar = 0.5 mm; 75 median lobe of aedeagus, ventral view (paralectotype, Assam), scale bar = 0.5 mm; 76 median lobe of aedeagus, ventral view (Guangxi, China), scale bar = 0.5 mm." pageId="34" pageNumber="35">fig. 67</figureCitation>
).
</paragraph>
<paragraph id="5A0497322A63D865FF6E78431B45B789" blockId="34.[151,1437,151,1941]" pageId="34" pageNumber="35">
<emphasis id="68CF4B202A63D865FF6E78431D8DB740" bold="true" box="[199,654,332,357]" pageId="34" pageNumber="35">Notes on non-type material examined.</emphasis>
Curiously, although this species is well represented in the collections of the MNHN by several specimens from the former
<collectingCountry id="22ACD7A22A63D865FD5878601C81B7AD" box="[753,898,367,392]" name="India" pageId="34" pageNumber="35">British India</collectingCountry>
,
<bibRefCitation id="3E2AEAC32A63D865FC2778601B51B7AD" box="[910,1106,367,392]" pageId="34" pageNumber="35" refId="ref28919" refString="Andrewes, H. E. (1924) Mission dans les provinces centrales de l'Inde et dans la region occidentale de l'Himalaya 1914 par Guy Babault. Carabidae. Imprimerie generale Lahure, Paris, 117 pp., 4 pls." type="book">Andrewes (1924)</bibRefCitation>
was not able to examine any
<taxonomicName id="9DBBECB12A63D865FF3E789B1FFBB788" box="[151,248,404,429]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A63D865FF3E789B1FFBB788" box="[151,248,404,429]" italics="true" pageId="34" pageNumber="35">Calleida</emphasis>
</taxonomicName>
specimen sampled by Guy Babault in his expedition to that region in 1914.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A63D865FF6E78B81A75B418" pageId="34" pageNumber="35" type="diagnosis">
<paragraph id="5A0497322A63D865FF6E78B81A75B418" blockId="34.[151,1437,151,1941]" pageId="34" pageNumber="35">
<emphasis id="68CF4B202A63D865FF6E78B81E42B7F5" bold="true" box="[199,321,439,464]" pageId="34" pageNumber="35">Diagnosis.</emphasis>
This species is easily recognizable from most Asiatic
<taxonomicName id="9DBBECB12A63D865FC1C78B81B15B7F5" box="[949,1046,439,464]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A63D865FC1C78B81B15B7F5" box="[949,1046,439,464]" italics="true" pageId="34" pageNumber="35">Calleida</emphasis>
</taxonomicName>
except
<taxonomicName id="9DBBECB12A63D865FBD878B71BD5B7F5" authorityName="Andr. Cotype H. E Andrewes" authorityYear="1933" box="[1137,1238,440,464]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="rapax">
<emphasis id="68CF4B202A63D865FBD878B71BD5B7F5" box="[1137,1238,440,464]" italics="true" pageId="34" pageNumber="35">C. rapax</emphasis>
</taxonomicName>
by the following character combination: large size (L: 12.0
<quantity id="9D433AD72A63D865FD2078D31DF2B7D1" box="[649,753,476,501]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="34" pageNumber="35" unit="mm" value="14.5">14.5 mm</quantity>
, the largest Asiatic
<taxonomicName id="9DBBECB12A63D865FC7378D31B38B7D0" box="[986,1083,476,501]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A63D865FC7378D31B38B7D0" box="[986,1083,476,501]" italics="true" pageId="34" pageNumber="35">Calleida</emphasis>
</taxonomicName>
species); body dark in colour, pronotal disc largely blackish; legs completely blackish; elytra metallic green, with faint bluish-purple discal patch; median lobe of aedeagus markedly concaved ventrally. The comparisons with
<taxonomicName id="9DBBECB12A63D865FBAD7B2A1B65B419" authorityName="Andr. Cotype H. E Andrewes" authorityYear="1933" box="[1028,1126,549,572]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="rapax">
<emphasis id="68CF4B202A63D865FBAD7B2A1B65B419" box="[1028,1126,549,572]" italics="true" pageId="34" pageNumber="35">C. rapax</emphasis>
</taxonomicName>
are provided in the key.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A63D865FF6E7B481EA2B120" pageId="34" pageNumber="35" type="description">
<paragraph id="5A0497322A63D865FF6E7B481FEAB48D" blockId="34.[151,1437,151,1941]" pageId="34" pageNumber="35">
<emphasis id="68CF4B202A63D865FF6E7B481E5AB445" bold="true" box="[199,345,583,608]" pageId="34" pageNumber="35">Description.</emphasis>
The short original description provided by
<bibRefCitation id="3E2AEAC32A63D865FCE17B471CDAB445" box="[840,985,584,608]" pageId="34" pageNumber="35" refId="ref29370" refString="Bates, H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of the Carabidae. Annali del Museo Civico di Storia naturale di Genova, 32, 267 - 428." type="journal article">Bates (1892)</bibRefCitation>
in Latin, here translated and completed with some additional characters, is enough to distinguish this species (see also the Key to the species of this species group).
</paragraph>
<paragraph id="5A0497322A63D865FF6E7BBB1C0EB4D5" blockId="34.[151,1437,151,1941]" pageId="34" pageNumber="35">
<emphasis id="68CF4B202A63D865FF6E7BBB1E87B4E9" box="[199,388,692,717]" italics="true" pageId="34" pageNumber="35">General features</emphasis>
: As in
<figureCitation id="C2808BB72A63D865FE707BBB1D52B4E8" box="[473,593,692,717]" captionStart="FIGURES 6776" captionStartId="36.[151,264,213,235]" captionText="FIGURES 6776. Calleida doriae Bates. 67 habitus and labels of lectotype (female, Bhamo, Myanmar, MSNG), scale bar = 2.0 mm; 68 habitus of paralectotype (male, Assam, India, MNHN), scale bar = 2.0 mm; 69 habitus (male, Yunnan, China), scale bar = 2.0 mm; 70 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (Yunnan, China), scale bar = 0.5 mm; 71 left and right parameres of aedeagus (Yunnan, China), scale bar = 0.2 mm; 72 male sternum IX (Yunnan, China), scale bar = 0.5 mm; 73 ovipositor (Yunnan, China), scale bar = 0.2 mm; 74 female reproductive tract (Yunnan, China), scale bar = 0.5 mm; 75 median lobe of aedeagus, ventral view (paralectotype, Assam), scale bar = 0.5 mm; 76 median lobe of aedeagus, ventral view (Guangxi, China), scale bar = 0.5 mm." pageId="34" pageNumber="35">fig. 6769</figureCitation>
. Large sized species: L: 12.015.0 mm; TL:
<quantity id="9D433AD72A63D865FBF97BBB1BF4B4E9" box="[1104,1271,692,717]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.45" metricValueMin="1.15" pageId="34" pageNumber="35" unit="mm" value="13.0" valueMax="14.5" valueMin="11.5">11.514.5 mm</quantity>
. Body relative wide, elytra depressed, slightly expanded at apical third.
</paragraph>
<paragraph id="5A0497322A63D865FF6E7BF31A44B580" blockId="34.[151,1437,151,1941]" pageId="34" pageNumber="35">
<emphasis id="68CF4B202A63D865FF6E7BF31E15B530" box="[199,278,764,789]" italics="true" pageId="34" pageNumber="35">Colour</emphasis>
: Dark brownish to black, shiny; head fully dark brownish or black, with two distinct reddish spots on vertex; one or two basal antennomeres yellowish, in sharp contrast with the following third and fourth antennomeres piceous, the remaining antennomeres brownish; disc of pronotum dark brown or blackish, with sides narrowly reddish; elytra metallic green, in some individuals golden-cupreous at apex, with large faint bluish purple discal patch; legs completely blackish, all tarsomeres blackish to dark brown; underside brownish or black.
</paragraph>
<paragraph id="5A0497322A63D865FF6E7ABE1E01B5C8" blockId="34.[151,1437,151,1941]" pageId="34" pageNumber="35">
<emphasis id="68CF4B202A63D865FF6E7ABE1EECB5ED" box="[199,495,943,968]" italics="true" pageId="34" pageNumber="35">Lustre and microsculpture</emphasis>
: meshes of the microsculpture isodiametric, vanished on head and pronotum, faint on elytra.
</paragraph>
<paragraph id="5A0497322A63D865FF6E7AF81D87B210" blockId="34.[151,1437,151,1941]" pageId="34" pageNumber="35">
<emphasis id="68CF4B202A63D865FF6E7AF81E07B235" box="[199,260,1015,1040]" italics="true" pageId="34" pageNumber="35">Head</emphasis>
wide, flat, smooth; genae markedly constricted to the neck; frontal foveae shallow and wide; antennae reaching backwards the basal fifth of elytra.
</paragraph>
<paragraph id="5A0497322A63D865FF6E7D4E1B2FB258" blockId="34.[151,1437,151,1941]" pageId="34" pageNumber="35">
<emphasis id="68CF4B202A63D865FF6E7D4E1E35B27D" box="[199,310,1089,1112]" italics="true" pageId="34" pageNumber="35">Pronotum</emphasis>
wider than head, cordate; lateral margins widely explanate, widened in front, moderately reflexed and markedly sinuate in the basal third; posterior angles rectangular; disc smooth.
</paragraph>
<paragraph id="5A0497322A63D865FF6E7D881CDAB2E0" blockId="34.[151,1437,151,1941]" pageId="34" pageNumber="35">
<emphasis id="68CF4B202A63D865FF6E7D881E0EB285" box="[199,269,1159,1184]" italics="true" pageId="34" pageNumber="35">Elytra</emphasis>
elongate and depressed; striae finely punctate; intervals smooth, flat or slightly convex; apical margin obliquely truncate, with outer angles gradually thickened but not angulate.
</paragraph>
<paragraph id="5A0497322A63D865FF6E7DC01ECBB328" blockId="34.[151,1437,151,1941]" pageId="34" pageNumber="35">
<emphasis id="68CF4B202A63D865FF6E7DC01E3FB2CD" box="[199,316,1231,1256]" italics="true" pageId="34" pageNumber="35">Underside</emphasis>
: Abdominal sternum VII in male normally notched at middle and with one seta on each side in male; two setae in females.
</paragraph>
<paragraph id="5A0497322A63D865FF6E7C181C20B02D" blockId="34.[151,1437,151,1941]" pageId="34" pageNumber="35">
<emphasis id="68CF4B202A63D865FF6E7C181E6DB315" box="[199,366,1303,1328]" italics="true" pageId="34" pageNumber="35">Male genitalia</emphasis>
: median lobe of aedeagus (
<figureCitation id="C2808BB72A63D865FD047C181DF8B315" box="[685,763,1303,1328]" captionStart="FIGURES 6776" captionStartId="36.[151,264,213,235]" captionText="FIGURES 6776. Calleida doriae Bates. 67 habitus and labels of lectotype (female, Bhamo, Myanmar, MSNG), scale bar = 2.0 mm; 68 habitus of paralectotype (male, Assam, India, MNHN), scale bar = 2.0 mm; 69 habitus (male, Yunnan, China), scale bar = 2.0 mm; 70 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (Yunnan, China), scale bar = 0.5 mm; 71 left and right parameres of aedeagus (Yunnan, China), scale bar = 0.2 mm; 72 male sternum IX (Yunnan, China), scale bar = 0.5 mm; 73 ovipositor (Yunnan, China), scale bar = 0.2 mm; 74 female reproductive tract (Yunnan, China), scale bar = 0.5 mm; 75 median lobe of aedeagus, ventral view (paralectotype, Assam), scale bar = 0.5 mm; 76 median lobe of aedeagus, ventral view (Guangxi, China), scale bar = 0.5 mm." pageId="34" pageNumber="35">fig. 70</figureCitation>
) slightly bent, its middle part strongly expanded, and then abruptly narrowed in dorsal view; dorsal and ventral margins slightly curved in lateral view, ventral surface markedly concaved in the middle; apical orifice pleuropic left, with a triangular chitinized piece; apical lamina flat, rounded at apex, expanded and thickened, basal width a little greater than length (W/L about 1.11.3); endophallus with two chitinized copulatory pieces, located in the middle area near the left lateral margin, long and narrow, close to each other at base, V-shaped; the dorsal one much shorter than the ventral one; left paramere depressed on the dorsal side; right paramere barely curved at apex (
<figureCitation id="C2808BB72A63D865FD607CE01C15B02D" box="[713,790,1519,1544]" captionStart="FIGURES 6776" captionStartId="36.[151,264,213,235]" captionText="FIGURES 6776. Calleida doriae Bates. 67 habitus and labels of lectotype (female, Bhamo, Myanmar, MSNG), scale bar = 2.0 mm; 68 habitus of paralectotype (male, Assam, India, MNHN), scale bar = 2.0 mm; 69 habitus (male, Yunnan, China), scale bar = 2.0 mm; 70 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (Yunnan, China), scale bar = 0.5 mm; 71 left and right parameres of aedeagus (Yunnan, China), scale bar = 0.2 mm; 72 male sternum IX (Yunnan, China), scale bar = 0.5 mm; 73 ovipositor (Yunnan, China), scale bar = 0.2 mm; 74 female reproductive tract (Yunnan, China), scale bar = 0.5 mm; 75 median lobe of aedeagus, ventral view (paralectotype, Assam), scale bar = 0.5 mm; 76 median lobe of aedeagus, ventral view (Guangxi, China), scale bar = 0.5 mm." pageId="34" pageNumber="35">fig. 71</figureCitation>
).
</paragraph>
<paragraph id="5A0497322A63D865FF6E7F1B1EA2B120" blockId="34.[151,1437,151,1941]" pageId="34" pageNumber="35">
<emphasis id="68CF4B202A63D865FF6E7F1B1E85B008" box="[199,390,1556,1581]" italics="true" pageId="34" pageNumber="35">Female genitalia</emphasis>
(reproductive tract
<figureCitation id="C2808BB72A63D865FDCA7F1B1DAEB009" box="[611,685,1556,1581]" captionStart="FIGURES 6776" captionStartId="36.[151,264,213,235]" captionText="FIGURES 6776. Calleida doriae Bates. 67 habitus and labels of lectotype (female, Bhamo, Myanmar, MSNG), scale bar = 2.0 mm; 68 habitus of paralectotype (male, Assam, India, MNHN), scale bar = 2.0 mm; 69 habitus (male, Yunnan, China), scale bar = 2.0 mm; 70 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (Yunnan, China), scale bar = 0.5 mm; 71 left and right parameres of aedeagus (Yunnan, China), scale bar = 0.2 mm; 72 male sternum IX (Yunnan, China), scale bar = 0.5 mm; 73 ovipositor (Yunnan, China), scale bar = 0.2 mm; 74 female reproductive tract (Yunnan, China), scale bar = 0.5 mm; 75 median lobe of aedeagus, ventral view (paralectotype, Assam), scale bar = 0.5 mm; 76 median lobe of aedeagus, ventral view (Guangxi, China), scale bar = 0.5 mm." pageId="34" pageNumber="35">fig. 74</figureCitation>
and ovipositor
<figureCitation id="C2808BB72A63D865FCF57F1B1CAAB009" box="[860,937,1556,1581]" captionStart="FIGURES 6776" captionStartId="36.[151,264,213,235]" captionText="FIGURES 6776. Calleida doriae Bates. 67 habitus and labels of lectotype (female, Bhamo, Myanmar, MSNG), scale bar = 2.0 mm; 68 habitus of paralectotype (male, Assam, India, MNHN), scale bar = 2.0 mm; 69 habitus (male, Yunnan, China), scale bar = 2.0 mm; 70 median lobe of aedeagus, right-lateral, ventral, left-lateral, and dorsal views (Yunnan, China), scale bar = 0.5 mm; 71 left and right parameres of aedeagus (Yunnan, China), scale bar = 0.2 mm; 72 male sternum IX (Yunnan, China), scale bar = 0.5 mm; 73 ovipositor (Yunnan, China), scale bar = 0.2 mm; 74 female reproductive tract (Yunnan, China), scale bar = 0.5 mm; 75 median lobe of aedeagus, ventral view (paralectotype, Assam), scale bar = 0.5 mm; 76 median lobe of aedeagus, ventral view (Guangxi, China), scale bar = 0.5 mm." pageId="34" pageNumber="35">fig. 73</figureCitation>
): spermatheca digitiform, about same length as the pedicel, surface very faintly whorled, with a fine and sharp basal projection; spermathecal pedicel straight at base, an then weakly bisinuate, with a large apical protuberance; spermathecal gland duct a little longer than spermatheca, laterally inserted on the basal projection; glandular area weakly widened, about same length as gland duct, atrium near base, slightly swollen. Gonocoxite II of ovipositor 2.5 times as long as the basal width, narrowed to apex, apex truncate, not oblique; setose from the apical third to apex; outer margin a little curved; apex with membranous extension.
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<emphasis id="68CF4B202A63D865FF6E7E001D9DB10D" bold="true" box="[199,670,1807,1832]" pageId="34" pageNumber="35">Geographical distribution and habitat.</emphasis>
Widely distributed in Oriental region, but only recorded from the mainland (
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):
<collectingCountry id="22ACD7A22A63D865FEC47E3A1EDFB168" box="[365,476,1845,1869]" name="Myanmar" pageId="34" pageNumber="35">Myanmar</collectingCountry>
,
<collectingCountry id="22ACD7A22A63D865FE4E7E3A1D23B169" box="[487,544,1845,1868]" name="Laos" pageId="34" pageNumber="35">Laos</collectingCountry>
,
<collectingCountry id="22ACD7A22A63D865FD857E3B1D8DB168" box="[556,654,1844,1869]" name="Vietnam" pageId="34" pageNumber="35">Vietnam</collectingCountry>
,
<collectingCountry id="22ACD7A22A63D865FD307E3B1C03B168" box="[665,768,1844,1869]" name="Malaysia" pageId="34" pageNumber="35">Malaysia</collectingCountry>
(Malay Peninsula), SW
<collectingCountry id="22ACD7A22A63D865FBBC7E3B1B5BB168" box="[1045,1112,1844,1869]" name="China" pageId="34" pageNumber="35">China</collectingCountry>
(
<collectingRegion id="987F59D02A63D865FBCE7E3B1BC2B169" box="[1127,1217,1844,1868]" country="China" name="Yunnan" pageId="34" pageNumber="35">Yunnan</collectingRegion>
,
<collectingRegion id="987F59D02A63D865FB657E3B1A2AB168" box="[1228,1321,1844,1869]" country="China" name="Sichuan" pageId="34" pageNumber="35">Sichuan</collectingRegion>
,
<collectingRegion id="987F59D02A63D865FA9C7E3B1A9BB168" box="[1333,1432,1844,1869]" country="China" name="Guangxi" pageId="34" pageNumber="35">Guangxi</collectingRegion>
,
<collectingRegion id="987F59D02A63D865FF3E7E581FEFB155" box="[151,236,1879,1904]" country="China" name="Xizang" pageId="34" pageNumber="35">Xizang</collectingRegion>
),
<collectingCountry id="22ACD7A22A63D865FEAB7E581E55B155" box="[258,342,1879,1904]" name="Bhutan" pageId="34" pageNumber="35">Bhutan</collectingCountry>
, E
<collectingCountry id="22ACD7A22A63D865FE297E581EC5B155" box="[384,454,1879,1904]" name="Nepal" pageId="34" pageNumber="35">Nepal</collectingCountry>
, N and NE
<collectingCountry id="22ACD7A22A63D865FDFF7E581D90B155" box="[598,659,1879,1904]" name="India" pageId="34" pageNumber="35">India</collectingCountry>
. Some individuals were collected using canopy fogging in tropical forest.
</paragraph>
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</treatment>
</subSection>
</document>

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Key to species of
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species group
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1. Basal one or two antennomeres reddish yellow, markedly contrasting in colour with the piceous antennomeres 3 and 4; subse- quent antennomeres brownish; head dark brownish or black with one or two small reddish spots on frons; pronotum black to piceous, with sides narrowly reddish; all tarsomeres blackish. Median lobe of aedeagus elongate and slenderer, with apical lamella slightly longer, W/L about 1.11.3 (
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). (
<collectingCountry id="22ACD7A22A60D866FD7C78391C36B76E" box="[725,821,310,331]" name="Myanmar" pageId="33" pageNumber="34">Myanmar</collectingCountry>
,
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,
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, Malay peninsula, SW
<collectingCountry id="22ACD7A22A60D866FB0178391BE7B76E" box="[1192,1252,310,331]" name="China" pageId="33" pageNumber="34">China</collectingCountry>
, N, NE
<collectingCountry id="22ACD7A22A60D866FA8678391A61B76E" box="[1327,1378,310,331]" name="India" pageId="33" pageNumber="34">India</collectingCountry>
,
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,
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)...........................................................................[10]
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Bates
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- Basal three or four antennomeres yellowish to reddish, moderately contrasting in colour with the brownish following ones; head mostly reddish brown; pronotum piceous to brownish, with sides widely reddish; fourth and fifth tarsomeres brownish. Median lobe of aedeagus stout, with apical lamella slightly shorter, W/L about 1.4 (
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). (NW India)............................................................................................... [11]
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<emphasis id="68CF4B202A61D867FF3E7C811D1EB38D" bold="true" box="[151,541,1422,1448]" pageId="32" pageNumber="33">
7.
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<emphasis id="68CF4B202A61D867FF1D7C811E6FB38D" bold="true" box="[180,364,1422,1448]" italics="true" pageId="32" pageNumber="33">Calleida doriae</emphasis>
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species group
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<emphasis id="68CF4B202A61D867FF3E7CDB1EA3B3C8" bold="true" box="[151,416,1492,1517]" pageId="32" pageNumber="33">Diagnostic characters</emphasis>
: With the character states of
<taxonomicName id="9DBBECB12A61D867FCA57CDB1C6FB3C8" box="[780,876,1492,1517]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A61D867FCA57CDB1C6FB3C8" box="[780,876,1492,1517]" italics="true" pageId="32" pageNumber="33">Calleida</emphasis>
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(in the widest sense, see above), restricted or amplified as follows: large sized species (L:
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); body blackish or reddish brown, all parts of legs including tarsi blackish; elytra metallic green, with vague blue-violet patch on disc near the suture; metasternum and metacoxae with pubescence; abdominal sternum VII with one seta on each side in males, two in females; median lobe of aedeagus markedly concave ventrally, without ventral ridges or lobes; apical lamina short.
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<emphasis id="68CF4B202A61D867FF6E7F861E85B087" bold="true" box="[199,390,1673,1698]" pageId="32" pageNumber="33">Taxonomic note</emphasis>
: Two very closely related species,
<taxonomicName id="9DBBECB12A61D867FCBE7F851CCBB084" authority="Bates" authorityName="Bates Lectotypus" authorityYear="1892" box="[791,968,1673,1698]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="doriae">
<emphasis id="68CF4B202A61D867FCBE7F851C81B087" box="[791,898,1673,1698]" italics="true" pageId="32" pageNumber="33">C. doriae</emphasis>
Bates
</taxonomicName>
and
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<emphasis id="68CF4B202A61D867FBAB7F851B65B084" box="[1026,1126,1674,1697]" italics="true" pageId="32" pageNumber="33">C. rapax</emphasis>
Andrewes
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are attributed to this group. The former species is known from several localities in northern
<collectingCountry id="22ACD7A22A61D867FB8D7FA21B63B0E3" box="[1060,1120,1709,1734]" name="India" pageId="32" pageNumber="33">India</collectingCountry>
, southern
<collectingCountry id="22ACD7A22A61D867FB777FA21A22B0E3" box="[1246,1313,1709,1734]" name="China" pageId="32" pageNumber="33">China</collectingCountry>
and other southeast Asian counties, whereas the latter is restricted to northwest
<collectingCountry id="22ACD7A22A61D867FC087FDE1CD8B0CF" box="[929,987,1745,1770]" name="India" pageId="32" pageNumber="33">India</collectingCountry>
(
<figureCitation id="C2808BB72A61D867FC407FDE1B31B0CF" box="[1001,1074,1745,1770]" captionStart="MAP 6" captionStartId="37.[151,209,1692,1714]" captionTargetBox="[151,1436,700,1671]" captionTargetId="figure@37.[151,1436,700,1671]" captionTargetPageId="37" captionText="MAP 6 Distributions for species in the C. doriae group: C. doriae Bates (blue); C. rapax Andrewes (red)." httpUri="https://zenodo.org/record/1302745/files/figure.png" pageId="32" pageNumber="33">map 6</figureCitation>
).
</paragraph>
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The two taxa of this group are very similar, and could be treated as synonyms or subspecies of one species. Further material from across the wide distribution range of species of this group will allow a better solution of the question. This group is considered related to the
<taxonomicName id="9DBBECB12A61D867FD127E311C5CB173" authorityName="Dejean" authorityYear="1831" box="[699,863,1853,1878]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="chloroptera">
<emphasis id="68CF4B202A61D867FD127E311C5CB173" box="[699,863,1853,1878]" italics="true" pageId="32" pageNumber="33">C. chloroptera</emphasis>
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species group for the similarities in: large size and depressed body; the presence of a vague bluish purple patch on elytral disc; and metacoxae with accessory pubescence.
</paragraph>
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<mods:title id="D192E3F2DA3A6D096BD5575DE83259CB">Revision of the Oriental species of Calleida Latreille (sensu lato). Part 1: Introduction, groups of species, and species of six species groups (Coleoptera: Carabidae: Lebiini)</mods:title>
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8.
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<emphasis id="68CF4B202A64D862FF1D7FF21EAEB132" bold="true" box="[180,429,1789,1815]" italics="true" pageId="37" pageNumber="38">Calleida chloroptera</emphasis>
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species group
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<paragraph id="5A0497322A64D862FF3E7E4B1DE4B1C8" blockId="37.[151,1436,1860,2029]" pageId="37" pageNumber="38">
<emphasis id="68CF4B202A64D862FF3E7E4B1EA3B178" bold="true" box="[151,416,1860,1885]" pageId="37" pageNumber="38">Diagnostic characters</emphasis>
: With the character states of
<taxonomicName id="9DBBECB12A64D862FCA57E4B1C6FB178" box="[780,876,1860,1885]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A64D862FCA57E4B1C6FB178" box="[780,876,1860,1885]" italics="true" pageId="37" pageNumber="38">Calleida</emphasis>
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(in the widest sense, see above), restricted or amplified as follows: medium-large sized species (L: 9.511.0 mm); body depressed; elytra metallic green to cupreous, disc usually with vague bluish purple patch; metacoxae with accessory pubescence; median lobe of aedeagus of peculiar shape, its ventral side with irregular and sharp ridges; spermathecal pedicel with large apical protuberance, about one third length as spermatheca.
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<paragraph id="5A0497322A67D861FF6E79981A9DB741" blockId="38.[151,1438,151,429]" pageId="38" pageNumber="39">
<emphasis id="68CF4B202A67D861FF6E79981E81B695" bold="true" box="[199,386,151,176]" pageId="38" pageNumber="39">Taxonomic note</emphasis>
: Several described taxa are attributed to this species group, including some widely distributed species such as
<taxonomicName id="9DBBECB12A67D861FEEA79B21D40B6F0" authority="Redtenbacher" authorityName="Redtenbacher" box="[323,579,188,213]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="lepida">
<emphasis id="68CF4B202A67D861FEEA79B21EA4B6F0" box="[323,423,188,213]" italics="true" pageId="38" pageNumber="39">C. lepida</emphasis>
Redtenbacher
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and
<taxonomicName id="9DBBECB12A67D861FDD179B21C2DB6F1" authority="Bates" authorityName="Bates" authorityYear="1892" box="[632,814,188,213]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="sultana">
<emphasis id="68CF4B202A67D861FDD179B21DEBB6F0" box="[632,744,188,213]" italics="true" pageId="38" pageNumber="39">C. sultana</emphasis>
Bates
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, but some of them are synonyms of previously described taxa, and some others are still undescribed. This species group has a wide geographical range, from the westernmost India to Sulawesi in the east. All valid species are strictly allopatric to each other, and some of them are narrowly distributed. The highly modified male genitalia define the different species, but some species are very difficult to distinguish by external features. We will review this group in our third revision of Asiatic
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species (in progress).
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The monophyly of the
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group is quite clear, for unique and peculiar characters in male genitalia. This group could be closely related to the
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<emphasis id="68CF4B202A67D861FDC7789A1DDBB788" box="[622,728,404,429]" italics="true" pageId="38" pageNumber="39">C. doriae</emphasis>
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<mods:part id="B73AA371BB60A7791D4525C46CC3064F">
<mods:date id="EB2ED3EC10C5F3D5AA6AEE5B4C224DE7">2018</mods:date>
<mods:detail id="F2632DE8265D3789338065DD3E7A85BA" type="pubDate">
<mods:number id="E170945068179FA9CB86024070F660D5">2018-06-29</mods:number>
</mods:detail>
<mods:detail id="5CD00D9272717CF1BD34CA4F4AA311FE" type="volume">
<mods:number id="C5E79855DB19A8B5C3FCBD54BFCB8EA2">4442</mods:number>
</mods:detail>
<mods:detail id="A7BFD2E5F6D4AC5A7DE893A4980D352D" type="issue">
<mods:number id="9B961B4049D5891B671DBA12ADF69D8B">1</mods:number>
</mods:detail>
</mods:part>
</mods:relatedItem>
<mods:classification id="F8BBE7DEFD0441EC3F0EE88A7048A2EC">journal volume</mods:classification>
<mods:identifier id="E64E3BA3C7F7D5D9EEB1BA591639C627" type="CLB-Dataset">29772</mods:identifier>
<mods:identifier id="D0DF41893C812D26E62B3CDD0952E033" type="DOI">10.11646/zootaxa.4442.1.1</mods:identifier>
<mods:identifier id="D90D4EFA463098036FDA7FAA00BABCAC" type="GBIF-Dataset">ded8bdb6-f8ba-4e0b-bed1-1ec08963edfd</mods:identifier>
<mods:identifier id="8B8EC5ECFD17E1B9A5AFCF572B139C52" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="16529228A0B0A77D8D2B6A868D41CFF4" type="Zenodo-Dep">1302713</mods:identifier>
<mods:identifier id="39A0475CACCA96695D8B2EDE6DAC943B" type="ZooBank">195B0471-553A-4617-B901-E9DBD2323D14</mods:identifier>
</mods:mods>
<treatment id="D21226242A65D862FF3E7B291FEFB4A0" ID-DOI="http://doi.org/10.5281/zenodo.5950881" ID-GBIF-Taxon="146043494" ID-Zenodo-Dep="5950881" LSID="urn:lsid:plazi:treatment:D21226242A65D862FF3E7B291FEFB4A0" httpUri="http://treatment.plazi.org/id/D21226242A65D862FF3E7B291FEFB4A0" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
<subSubSection id="12A1C4B92A65D863FF3E7B291D4EB465" box="[151,589,550,576]" pageId="36" pageNumber="37" type="nomenclature">
<paragraph id="5A0497322A65D863FF3E7B291D4EB465" blockId="36.[151,589,550,576]" box="[151,589,550,576]" pageId="36" pageNumber="37">
<heading id="014C205E2A65D863FF3E7B291D4EB465" bold="true" box="[151,589,550,576]" fontSize="11" level="1" pageId="36" pageNumber="37" reason="1">
<emphasis id="68CF4B202A65D863FF3E7B291D4EB465" bold="true" box="[151,589,550,576]" pageId="36" pageNumber="37">
[11]
<taxonomicName id="9DBBECB12A65D863FF647B291D4EB465" ID-CoL="PLBD" authority="Andrewes, 1933" authorityName="Andr. Cotype H. E Andrewes" authorityYear="1933" box="[205,589,550,576]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="rapax">
<emphasis id="68CF4B202A65D863FF647B291E83B465" bold="true" box="[205,384,550,576]" italics="true" pageId="36" pageNumber="37">Calleida rapax</emphasis>
<bibRefCitation id="3E2AEAC32A65D863FE2E7B291D4EB465" box="[391,589,550,576]" pageId="36" pageNumber="37" refId="ref29063" refString="Andrewes, H. E. (1933) Papers on Oriental Carabidae. XXVIII. The Annals and Magazine of Natural History, Series 10, 11 (63), 399 - 413. https: // doi. org / 10.1080 / 00222933308673670" type="journal article">Andrewes, 1933</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A65D863FF3E7B621A1DB4A6" box="[151,1310,621,643]" pageId="36" pageNumber="37" type="reference_group">
<paragraph id="5A0497322A65D863FF3E7B621A1DB4A6" blockId="36.[151,1310,621,643]" box="[151,1310,621,643]" pageId="36" pageNumber="37">
<taxonomicName id="9DBBECB12A65D863FF3E7B621D0CB4A6" ID-CoL="PLBD" authority="Andrewes, 1933: 399" authorityName="Andr. Cotype H. E Andrewes" authorityPageNumber="399" authorityYear="1933" box="[151,527,621,643]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="rapax">
<emphasis id="68CF4B202A65D863FF3E7B621E2CB4A6" box="[151,303,621,643]" italics="true" pageId="36" pageNumber="37">Calleida rapax</emphasis>
<bibRefCitation id="3E2AEAC32A65D863FE9F7B621EDCB4A6" box="[310,479,621,643]" pageId="36" pageNumber="37" refId="ref29063" refString="Andrewes, H. E. (1933) Papers on Oriental Carabidae. XXVIII. The Annals and Magazine of Natural History, Series 10, 11 (63), 399 - 413. https: // doi. org / 10.1080 / 00222933308673670" type="journal article">Andrewes, 1933</bibRefCitation>
: 399
</taxonomicName>
(
<typeStatus id="850029902A65D863FDB27B621D4CB4A6" box="[539,591,621,643]" pageId="36" pageNumber="37">Type</typeStatus>
locality:
<collectingCountry id="22ACD7A22A65D863FD067B621DE4B4A6" box="[687,743,621,643]" name="India" pageId="36" pageNumber="37">India</collectingCountry>
: Dehra
<collectingRegion id="987F59D02A65D863FC9C7B611C60B4A6" box="[821,867,622,643]" country="United Kingdom" name="Down" pageId="36" pageNumber="37">Dun</collectingRegion>
, Lachiwala;
<typeStatus id="850029902A65D863FC4D7B621B3EB4A6" box="[996,1085,621,643]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
deposited in NHML).
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A65D863FF3E7BA01A4BB5E1" pageId="36" pageNumber="37" type="materials_examined">
<paragraph id="5A0497322A65D863FF3E7BA01EFAB585" blockId="36.[151,1437,687,2009]" pageId="36" pageNumber="37">
<emphasis id="68CF4B202A65D863FF3E7BA01EBCB4ED" bold="true" box="[151,447,687,712]" pageId="36" pageNumber="37">
<typeStatus id="850029902A65D863FF3E7BA01FD1B4ED" box="[151,210,687,712]" pageId="36" pageNumber="37">Type</typeStatus>
material examined
</emphasis>
:
<materialsCitation id="EAD39D6F2A65D863FE787BA01EF7B535" ID-GBIF-Occurrence="1852047398" collectedFrom="“ Tectona grandis ”, &quot; Under bark" collectingDate="1923-04-06" collectionCode="B" collectorName="C. F. C. Beeson" country="India" county="Dehra Dun" location="Lachiwala, Dehra" pageId="36" pageNumber="37" specimenCode="B.M 1932-26" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="850029902A65D863FE787BA01D3FB4ED" box="[465,572,687,712]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
,
<specimenCount id="4CBD5CBB2A65D863FDE37BA01D80B4ED" box="[586,643,687,712]" pageId="36" pageNumber="37" type="male">male</specimenCount>
, “
<location id="5F64C1E92A65D863FD357BA01C69B4ED" LSID="urn:lsid:plazi:treatment:D21226242A65D862FF3E7B291FEFB4A0:5F64C1E92A65D863FD357BA01C69B4ED" box="[668,874,687,712]" country="India" county="Dehra Dun" name="Lachiwala, Dehra" pageId="36" pageNumber="37">Lachiwala, Dehra</location>
Dun U.P.,
<collectorName id="F74EF2E42A65D863FC467BBF1B99B4E2" box="[1007,1178,688,712]" pageId="36" pageNumber="37">C.F.C. Beeson</collectorName>
.,
<collectingDate id="3E41481A2A65D863FB057BA01A1EB4ED" box="[1196,1309,687,712]" pageId="36" pageNumber="37" value="1923-04-06">
<date id="2E05B1F22A65D863FB057BA01A1EB4ED" box="[1196,1309,687,712]" pageId="36" pageNumber="37" value="1923-04-06">6.IV.1923</date>
</collectingDate>
”,
<collectedFrom id="E9A7D4CF2A65D863FA917BBF1E90B4C8" pageId="36" pageNumber="37">“Tectona grandis”, &quot;Under bark</collectedFrom>
&quot;, &quot;326&quot;, &quot;
<collectingCountry id="22ACD7A22A65D863FE577BDB1D3EB4C8" box="[510,573,724,749]" name="India" pageId="36" pageNumber="37">India</collectingCountry>
.
<collectingCounty id="B365EFBE2A65D863FDE17BDB1DC0B4C9" box="[584,707,724,749]" pageId="36" pageNumber="37">Dehra Dun</collectingCounty>
Coll.
<specimenCode id="0A1D3F492A65D863FCA17BDA1CA0B4C8" box="[776,931,724,749]" pageId="36" pageNumber="37">B.M 1932-26</specimenCode>
&quot;, &quot;
<taxonomicName id="9DBBECB12A65D863FC697BDB1BAFB4C8" authority="Andr." authorityName="Andr. Cotype, Andrewes" authorityYear="1933" box="[960,1196,724,749]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="rapax">Calleida rapax Andr.</taxonomicName>
Type H.E. Andrewes det.&quot;, &quot;Type&quot; (NHML,
<figureCitation id="C2808BB72A65D863FE337BF81EE4B535" box="[410,487,759,784]" captionStart="FIGURES 7778" captionStartId="36.[151,264,428,450]" captionText="FIGURES 7778. Calleida rapax Andrewes. 77 median lobe of aedeagus, ventral view (paratype, Landsdowne, Inida, NHML), scale bar = 0.5 mm; 78 habitus and labels of holotype (male, Dehra Dun, India, NHML), scale bar = 2.0 mm." pageId="36" pageNumber="37">fig. 78</figureCitation>
).
</materialsCitation>
<typeStatus id="850029902A65D863FE527BF71D6FB535" box="[507,620,760,784]" pageId="36" pageNumber="37" type="paratype">Paratypes</typeStatus>
: one male, &quot;Nauri, 2,500 Landsdowne. B.M. Bathia.
<date id="2E05B1F22A65D863FB637BF71A4FB535" box="[1226,1356,759,784]" pageId="36" pageNumber="37" value="1926-03-18">18.III.1926</date>
”, “For. Res. Inst. Dehra Duna”, “dry ex Bamboo”, &quot;305&quot;, &quot;Co-type&quot;, “Andrewes coll. B.M.
<date id="2E05B1F22A65D863FBC67A131BD3B510" box="[1135,1232,796,821]" pageId="36" pageNumber="37" value="1945" valueMax="1997">1945-97</date>
”, “
<taxonomicName id="9DBBECB12A65D863FB5D7A131EDFB57D" authority="Andr. Cotype H. E Andrewes" authorityName="Andr. Cotype H. E Andrewes" authorityYear="1933" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="rapax">Calleida rapax Andr. Cotype H.E Andrewes</taxonomicName>
det.” (
<collectionCode id="3CAA0FF72A65D863FD817A4F1D82B57D" box="[552,641,832,856]" country="France" httpUri="http://grbio.org/cool/4jmm-iazx" name="Musee Guimet d'Histoire Naturelle de Lyon" pageId="36" pageNumber="37">MHNL</collectionCode>
); one female, &quot;New Forest, Dehra
<collectingRegion id="987F59D02A65D863FBAE7A4F1B39B572" box="[1031,1082,832,855]" country="United Kingdom" name="Down" pageId="36" pageNumber="37">Dun</collectingRegion>
, U.P. R.N. Mathur.
<date id="2E05B1F22A65D863FAB17A301A8DB57D" box="[1304,1422,831,856]" pageId="36" pageNumber="37" value="1932-09-06">6.IX.1932</date>
&quot;, &quot;1590&quot;, &quot;Co-type&quot;, &quot;For. Res. Inst. Dehra
<collectingRegion id="987F59D02A65D863FDD17A6B1DAEB559" box="[632,685,868,892]" country="United Kingdom" name="Down" pageId="36" pageNumber="37">Dun</collectingRegion>
&quot;, &quot;
<taxonomicName id="9DBBECB12A65D863FD657A6B1B96B558" authority="Andr. Cotype, Andrewes" authorityName="Andr. Cotype, Andrewes" authorityYear="1933" box="[716,1173,868,893]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="rapax">Calleida rapax Andr. Cotype, Andrewes</taxonomicName>
det.&quot;, &quot;H.E. Andrewes Coll. B.M.
<date id="2E05B1F22A65D863FEBC7A881E7BB585" box="[277,376,903,928]" pageId="36" pageNumber="37" value="1945" valueMax="1997">1945-97</date>
.&quot; (
<collectionCode id="3CAA0FF72A65D863FE3D7A871EEEB585" box="[404,493,904,928]" country="France" httpUri="http://grbio.org/cool/4jmm-iazx" name="Musee Guimet d'Histoire Naturelle de Lyon" pageId="36" pageNumber="37">MHNL</collectionCode>
).
</paragraph>
<paragraph id="5A0497322A65D863FF6E7AA31A4BB5E1" blockId="36.[151,1437,687,2009]" box="[199,1352,940,965]" pageId="36" pageNumber="37">
<emphasis id="68CF4B202A65D863FF6E7AA31D26B5E0" bold="true" box="[199,549,940,965]" pageId="36" pageNumber="37">Non-type material examined:</emphasis>
1 ex., “
<collectingCountry id="22ACD7A22A65D863FDD37AA31DBBB5E0" box="[634,696,940,965]" name="India" pageId="36" pageNumber="37">India</collectingCountry>
, Dehra
<collectingRegion id="987F59D02A65D863FCA67AA21C40B5E1" box="[783,835,941,964]" country="United Kingdom" name="Down" pageId="36" pageNumber="37">Dun</collectingRegion>
. G. Rogers 1906-202” (
<collectionCode id="3CAA0FF72A65D863FBE07AA31B84B5E1" box="[1097,1159,940,964]" country="Germany" httpUri="http://grbio.org/cool/mqei-3i36" name="Culture Collection of Algae at the University of Marburg" pageId="36" pageNumber="37">CCA</collectionCode>
, ex S.L Straneo).
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A65D863FF6E7AC01B99B2B8" pageId="36" pageNumber="37" type="discussion">
<paragraph id="5A0497322A65D863FF6E7AC01DEBB270" blockId="36.[151,1437,687,2009]" pageId="36" pageNumber="37">
<emphasis id="68CF4B202A65D863FF6E7AC01E88B5CD" bold="true" box="[199,395,975,1000]" pageId="36" pageNumber="37">Taxonomic note.</emphasis>
This taxon is very similar to
<taxonomicName id="9DBBECB12A65D863FD7B7ADF1C38B5CD" authorityName="Bates Lectotypus" authorityYear="1892" box="[722,827,975,1000]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="doriae">
<emphasis id="68CF4B202A65D863FD7B7ADF1C38B5CD" box="[722,827,975,1000]" italics="true" pageId="36" pageNumber="37">C. doriae</emphasis>
</taxonomicName>
. They only have small differences in colour and in the apex of the aedeagal median lobe. Considering their allopatric distribution,
<taxonomicName id="9DBBECB12A65D863FC577AFA1B61B229" authorityName="Andr. Cotype, Andrewes" authorityYear="1933" box="[1022,1122,1013,1036]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="rapax">
<emphasis id="68CF4B202A65D863FC577AFA1B61B229" box="[1022,1122,1013,1036]" italics="true" pageId="36" pageNumber="37">C. rapax</emphasis>
</taxonomicName>
could be treated as a mere northwestern isolated subspecies of
<taxonomicName id="9DBBECB12A65D863FDEA7D171DB1B215" authorityName="Bates Lectotypus" authorityYear="1892" box="[579,690,1047,1072]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="doriae">
<emphasis id="68CF4B202A65D863FDEA7D171DB1B215" box="[579,690,1047,1072]" italics="true" pageId="36" pageNumber="37">C. doriae</emphasis>
</taxonomicName>
, or a synonym of it. However, owing to the scarce examined material, at the moment we treat it as a valid species.
</paragraph>
<paragraph id="5A0497322A65D863FF6E7D6F1B99B2B8" blockId="36.[151,1437,687,2009]" pageId="36" pageNumber="37">
Its relationships suggested by
<bibRefCitation id="3E2AEAC32A65D863FD897D501DE6B25D" box="[544,741,1119,1144]" pageId="36" pageNumber="37" refId="ref29063" refString="Andrewes, H. E. (1933) Papers on Oriental Carabidae. XXVIII. The Annals and Magazine of Natural History, Series 10, 11 (63), 399 - 413. https: // doi. org / 10.1080 / 00222933308673670" type="journal article">Andrewes (1933)</bibRefCitation>
are wrong: “very closely allied to
<emphasis id="68CF4B202A65D863FBDE7D501BE2B25D" box="[1143,1249,1119,1144]" italics="true" pageId="36" pageNumber="37">femoralis</emphasis>
Chaud.,
<taxonomicName id="9DBBECB12A65D863FAE07D501FDBB2B9" authority="Bates" authorityName="Bates" authorityYear="1892" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="sultana">
<emphasis id="68CF4B202A65D863FAE07D501A9FB25D" box="[1353,1436,1119,1144]" italics="true" pageId="36" pageNumber="37">sultana</emphasis>
Bates
</taxonomicName>
, and
<taxonomicName id="9DBBECB12A65D863FEBB7D8B1EB5B2B8" authority="Andr." authorityName="Andr." box="[274,438,1156,1181]" class="Insecta" family="Gracillariidae" genus="Caloptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="68CF4B202A65D863FEBB7D8B1E68B2B8" box="[274,363,1156,1181]" italics="true" pageId="36" pageNumber="37">pallipes</emphasis>
Andr.
</taxonomicName>
”. This species was not cited by
<bibRefCitation id="3E2AEAC32A65D863FCB37D8B1CC4B2B8" box="[794,967,1156,1181]" pageId="36" pageNumber="37" refId="ref30044" refString="Jedlicka, A. (1963) Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae-Brachyninae (Coleoptera, Carabidae). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden, 28, 269 - 579." type="journal article">Jedlička (1963)</bibRefCitation>
in his monograph.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A65D863FF6E7DA81A60B29A" box="[199,1379,1191,1216]" pageId="36" pageNumber="37" type="diagnosis">
<paragraph id="5A0497322A65D863FF6E7DA81A60B29A" blockId="36.[151,1437,687,2009]" box="[199,1379,1191,1216]" pageId="36" pageNumber="37">
<emphasis id="68CF4B202A65D863FF6E7DA81E42B2E5" bold="true" box="[199,321,1191,1216]" pageId="36" pageNumber="37">Diagnosis.</emphasis>
See above, in the key to the species of the
<taxonomicName id="9DBBECB12A65D863FC8A7DA71C8EB2E5" authorityName="Bates Lectotypus" authorityYear="1892" box="[803,909,1191,1216]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="doriae">
<emphasis id="68CF4B202A65D863FC8A7DA71C8EB2E5" box="[803,909,1191,1216]" italics="true" pageId="36" pageNumber="37">C. doriae</emphasis>
</taxonomicName>
species group and diagnosis of
<taxonomicName id="9DBBECB12A65D863FB5B7DA71A5FB2E5" authorityName="Bates Lectotypus" authorityYear="1892" box="[1266,1372,1191,1216]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="doriae">
<emphasis id="68CF4B202A65D863FB5B7DA71A5FB2E5" box="[1266,1372,1191,1216]" italics="true" pageId="36" pageNumber="37">C. doriae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="12A1C4B92A65D862FF6E7DC31D30B7D0" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="description">
<paragraph id="5A0497322A65D863FF6E7DC31DB3B32D" blockId="36.[151,1437,687,2009]" pageId="36" pageNumber="37">
<emphasis id="68CF4B202A65D863FF6E7DC31E5AB2C0" bold="true" box="[199,345,1228,1253]" pageId="36" pageNumber="37">Description.</emphasis>
The original description provided by
<bibRefCitation id="3E2AEAC32A65D863FCBE7DC31CE3B2C1" box="[791,992,1228,1253]" pageId="36" pageNumber="37" refId="ref29063" refString="Andrewes, H. E. (1933) Papers on Oriental Carabidae. XXVIII. The Annals and Magazine of Natural History, Series 10, 11 (63), 399 - 413. https: // doi. org / 10.1080 / 00222933308673670" type="journal article">Andrewes (1933)</bibRefCitation>
, here completed with some additional characters, is enough to distinguish this species.
</paragraph>
<paragraph id="5A0497322A65D863FF6E7C1B1CDDB309" blockId="36.[151,1437,687,2009]" box="[199,990,1300,1325]" pageId="36" pageNumber="37">
<emphasis id="68CF4B202A65D863FF6E7C1B1E80B309" box="[199,387,1300,1325]" italics="true" pageId="36" pageNumber="37">General features</emphasis>
: As in
<figureCitation id="C2808BB72A65D863FE7C7C1B1D22B309" box="[469,545,1300,1325]" captionStart="FIGURES 7778" captionStartId="36.[151,264,428,450]" captionText="FIGURES 7778. Calleida rapax Andrewes. 77 median lobe of aedeagus, ventral view (paratype, Landsdowne, Inida, NHML), scale bar = 0.5 mm; 78 habitus and labels of holotype (male, Dehra Dun, India, NHML), scale bar = 2.0 mm." pageId="36" pageNumber="37">fig. 78</figureCitation>
. Large sized species: L: 12.013.0 mm.
</paragraph>
<paragraph id="5A0497322A65D863FF6E7C381FD8B020" blockId="36.[151,1437,687,2009]" pageId="36" pageNumber="37">
<emphasis id="68CF4B202A65D863FF6E7C381E15B375" box="[199,278,1335,1360]" italics="true" pageId="36" pageNumber="37">Colour</emphasis>
: Piceous to reddish brown, shiny; head reddish brown, occiput darker; antennomere 1 yellowish brown, antennomeres 2, 3 and basal half of 4 reddish, apical half of antennomere 4 and lateral sides of remaining antennomeres dark; base and apex of palpomeres yellowish; pronotal disc dark brown, with sides widely reddish; elytra metallic green with faint bluish purple discal patch, sometimes the marginal furrows greenish blue or violet; major parts of legs blackish, trochanters reddish, fourth and fifth tarsomeres reddish brown; underside brownish or black.
</paragraph>
<paragraph id="5A0497322A65D863FF6E7F1E1E58B068" blockId="36.[151,1437,687,2009]" pageId="36" pageNumber="37">
<emphasis id="68CF4B202A65D863FF6E7F1E1EFCB00D" box="[199,511,1551,1576]" italics="true" pageId="36" pageNumber="37">Lustre and microsculpture:</emphasis>
microsculpture vanished on head and pronotum; faint, formed by isodiametric meshes on elytra.
</paragraph>
<paragraph id="5A0497322A65D863FF6E7F581C08B0B0" blockId="36.[151,1437,687,2009]" pageId="36" pageNumber="37">
<emphasis id="68CF4B202A65D863FF6E7F581E07B055" box="[199,260,1623,1648]" italics="true" pageId="36" pageNumber="37">Head</emphasis>
flat and impunctate; frontal foveae shallow and uneven; eyes prominent; neck constriction evident; antennae reaching backwards to the basal fifth of elytra.
</paragraph>
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<emphasis id="68CF4B202A65D863FF6E7FAE1E35B09D" box="[199,310,1697,1720]" italics="true" pageId="36" pageNumber="37">Pronotum</emphasis>
rather flat and cordate, transverse (ratio PW/PL: 1.13-1.19), a little wider than the head; base oblique at sides, wider than anterior margin. Anterior angles rounded; lateral sides moderately explanate and slightly reflexed, rounded in front, then nearly straight to the posterior angles which are obtuse, not projecting laterally. Disc smooth; median furrow and basal foveae fairly deep, the foveae continued forward on each side forming shallow sulci, parallel with lateral sides.
</paragraph>
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<emphasis id="68CF4B202A65D863FF6E7E5B1E0EB148" box="[199,269,1876,1901]" italics="true" pageId="36" pageNumber="37">Elytra</emphasis>
depressed, three-fifths wider than pronotum, nearly four-fifth longer than wide; lateral sides slightly compressed behind shoulders and then a little dilated, so that the widest point is behind middle. Elytral apex obliquely truncate, the outer apical angles markedly widened, obtusely rounded, not angulate. Striae moderately deep, very finely punctate; intervals moderately convex, with some sparse minute punctuation.
</paragraph>
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<emphasis id="68CF4B202A64D862FF6E79981E3FB695" box="[199,316,151,176]" italics="true" pageId="37" pageNumber="38">Underside</emphasis>
with very fine scattered punctuation and pubescence on metasternum, more developed in males. Abdominal sternum VII in males notched at middle, with one seta on each side, two setae in females.
</paragraph>
<paragraph id="5A0497322A64D862FF6E79D01D14B7EA" blockId="37.[151,1437,151,645]" pageId="37" pageNumber="38">
<emphasis id="68CF4B202A64D862FF6E79D01E6EB6DD" box="[199,365,223,248]" italics="true" pageId="37" pageNumber="38">Male genitalia</emphasis>
very similar to that of
<taxonomicName id="9DBBECB12A64D862FDDC79EF1DE3B6DD" authorityName="Bates Lectotypus" authorityYear="1892" box="[629,736,223,248]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="doriae">
<emphasis id="68CF4B202A64D862FDDC79EF1DE3B6DD" box="[629,736,223,248]" italics="true" pageId="37" pageNumber="38">C. doriae</emphasis>
</taxonomicName>
, but apex of the median lobe of aedeagus a little shorter (
<figureCitation id="C2808BB72A64D862FADC79D01FB4B739" captionStart="FIGURES 7778" captionStartId="36.[151,264,428,450]" captionText="FIGURES 7778. Calleida rapax Andrewes. 77 median lobe of aedeagus, ventral view (paratype, Landsdowne, Inida, NHML), scale bar = 0.5 mm; 78 habitus and labels of holotype (male, Dehra Dun, India, NHML), scale bar = 2.0 mm." pageId="37" pageNumber="38">fig. 77</figureCitation>
); median lobe slightly bent, its middle part strongly expanded in dorsal view; dorsal and ventral margins slightly curved in lateral view, ventral surface markedly concaved in the middle; apical orifice pleuropic left, with a triangular chitinized piece; apical lamina flat, rounded at apex, expanded and thickened, basal width greater than length (W/L about 1.4); endophallus with two chitinized copulatory pieces, located in the middle area near the left lateral margin, long and narrow, close to each other at base, V shaped; left paramere depressed on the dorsal side; right paramere not curved at apex.
</paragraph>
<paragraph id="5A0497322A64D862FF6E78D31D30B7D0" blockId="37.[151,1437,151,645]" box="[199,563,476,501]" pageId="37" pageNumber="38">
<emphasis id="68CF4B202A64D862FF6E78D31E86B7D0" box="[199,389,476,501]" italics="true" pageId="37" pageNumber="38">Female genitalia</emphasis>
: not examined.
</paragraph>
</subSubSection>
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<emphasis id="68CF4B202A64D862FF6E78F01D9EB43D" bold="true" box="[199,669,511,536]" pageId="37" pageNumber="38">Geographical Distribution and habitat.</emphasis>
Only known from northwestern
<collectingCountry id="22ACD7A22A64D862FBBB78F01B48B43D" box="[1042,1099,511,536]" name="India" pageId="37" pageNumber="38">India</collectingCountry>
(
<figureCitation id="C2808BB72A64D862FBF27B0F1BA5B43D" box="[1115,1190,511,536]" captionStart="MAP 6" captionStartId="37.[151,209,1692,1714]" captionTargetBox="[151,1436,700,1671]" captionTargetId="figure@37.[151,1436,700,1671]" captionTargetPageId="37" captionText="MAP 6 Distributions for species in the C. doriae group: C. doriae Bates (blue); C. rapax Andrewes (red)." httpUri="https://zenodo.org/record/1302745/files/figure.png" pageId="37" pageNumber="38">map 6</figureCitation>
):
<collectingRegion id="987F59D02A64D862FB1578F01A45B43D" box="[1212,1350,511,536]" country="India" name="Uttarakhand" pageId="37" pageNumber="38">Uttarakhand</collectingRegion>
(Dehra
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, Lansdowne).
<bibRefCitation id="3E2AEAC32A64D862FED97B2B1D36B419" box="[368,565,548,573]" pageId="37" pageNumber="38" refId="ref29063" refString="Andrewes, H. E. (1933) Papers on Oriental Carabidae. XXVIII. The Annals and Magazine of Natural History, Series 10, 11 (63), 399 - 413. https: // doi. org / 10.1080 / 00222933308673670" type="journal article">Andrewes (1933)</bibRefCitation>
reported that the
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was collected under bark of
<emphasis id="68CF4B202A64D862FB077B2A1A60B418" box="[1198,1379,548,573]" italics="true" pageId="37" pageNumber="38">Tectona grandis</emphasis>
, and several
<typeStatus id="850029902A64D862FF587B471E5FB445" box="[241,348,584,608]" pageId="37" pageNumber="38" type="paratype">paratypes</typeStatus>
were in bamboo leaf roll, predaceous on
<taxonomicName id="9DBBECB12A64D862FCE77B481B6FB445" authority="(Cramer)" authorityName="Cramer" box="[846,1132,583,608]" class="Insecta" family="Hyblaeidae" genus="Hyblaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="puera">
<emphasis id="68CF4B202A64D862FCE77B481CF9B445" box="[846,1018,583,608]" italics="true" pageId="37" pageNumber="38">Hyblaea puera</emphasis>
(Cramer)
</taxonomicName>
and
<taxonomicName id="9DBBECB12A64D862FB027B481FEFB4A0" authority="Walker." authorityName="Walker." class="Insecta" family="Pyralidae" genus="Hapalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="machaeralis">
<emphasis id="68CF4B202A64D862FB027B481A9FB445" box="[1195,1436,583,608]" italics="true" pageId="37" pageNumber="38">Hapalia machaeralis</emphasis>
Walker.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="E347582936607298B3EEEAC00DE9BA68">Revision of the Oriental species of Calleida Latreille (sensu lato). Part 1: Introduction, groups of species, and species of six species groups (Coleoptera: Carabidae: Lebiini)</mods:title>
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9.
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<emphasis id="68CF4B202A67D861FF1D78F51EAFB431" bold="true" box="[180,428,506,532]" italics="true" pageId="38" pageNumber="39">Calleida splendidula</emphasis>
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species group
</emphasis>
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<subSubSection id="12A1C4B92A67D861FF3E7B4E1D21B20B" pageId="38" pageNumber="39" type="description">
<paragraph id="5A0497322A67D861FF3E7B4E1B22B573" blockId="38.[151,1437,577,1070]" pageId="38" pageNumber="39">
<emphasis id="68CF4B202A67D861FF3E7B4E1EA3B47F" bold="true" box="[151,416,577,602]" pageId="38" pageNumber="39">Diagnostic characters</emphasis>
: With the character states of
<taxonomicName id="9DBBECB12A67D861FCA57B4E1C6FB47F" box="[780,876,577,602]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A67D861FCA57B4E1C6FB47F" box="[780,876,577,602]" italics="true" pageId="38" pageNumber="39">Calleida</emphasis>
</taxonomicName>
(in the widest sense, see above), restricted or amplified as follows: small to medium sized species (L: 6.59.0 mm in general, exceptionally reaching
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). Head and pronotum piceous to reddish; legs with all tarsomeres reddish brown; elytra metallic green, cupreous or purple-reddish, often with a reddish or brownish patch on disc. Metacoxae glabrous except for the fixed three or four setae. Median lobe of aedeagus simple, not modified ventrally or apically, generally flat in form; right margin more or less sinuate in ventral aspect; apical lamina rounded; endophallus with two chitinized copulatory pieces similar in shape, close to each other, one fourth as long as one third of the median lobe. Spermathecal pedicel curved or curly in some species, spermatheca not whorled, with basal projection.
</paragraph>
<paragraph id="5A0497322A67D861FF6E7A6E1A59B5C3" blockId="38.[151,1437,577,1070]" pageId="38" pageNumber="39">
<emphasis id="68CF4B202A67D861FF6E7A6E1E87B55F" bold="true" box="[199,388,865,890]" pageId="38" pageNumber="39">Taxonomic note</emphasis>
: This is the most diverse and complicate group of Asiatic
<taxonomicName id="9DBBECB12A67D861FBB97A6E1B73B55F" box="[1040,1136,865,890]" class="Insecta" family="Carabidae" genus="Calleida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="68CF4B202A67D861FBB97A6E1B73B55F" box="[1040,1136,865,890]" italics="true" pageId="38" pageNumber="39">Calleida</emphasis>
</taxonomicName>
species. Several described taxa are attributable to this species group, distributed almost through the full Oriental Region. Some of them are very common, and some others are insufficiently described. Apart from the coloration, different species have only minute differences in external and genital characters that makes the definition of every species very difficult.
</paragraph>
<paragraph id="5A0497322A67D861FF6E7AFE1D21B20B" blockId="38.[151,1437,577,1070]" pageId="38" pageNumber="39">This group seems to be monophyletic, but many character states are inferred by plesiomorphies. The revision of this species group is in progress.</paragraph>
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Rubus
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Ph. J. Mueller et
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<paragraph id="DB146696DF39CA13CE98106B4BDED079" pageId="null" pageNumber="433">
(
<emphasis id="A8B44EC5A36E8EECD09DA6A8C483E3EA" italics="true" pageId="null" pageNumber="433">keine Abbildung</emphasis>
)
</paragraph>
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</title>
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<normalizedToken id="48446966E8B2DC3035DEDED41BB30FF1" originalValue="Schößling" pageId="null" pageNumber="433">Schoessling</normalizedToken>
bogig, kantig bis rund, zerstreut behaart bis kahl.
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5
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, oberseits
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behaart, unterseits
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, locker und abstehend behaart.
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mit nur etwa 0,2 mm lang gestielten
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, diese die abstehenden Haare nicht
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(bei den
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Arten der Gruppe
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gestielte
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); mit vielen geraden, flachen Stacheln.
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nach der
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.
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<normalizedToken id="A36A6978D4062E87F8DD222D8731DAE7" originalValue="weiß" pageId="null" pageNumber="433">weiss</normalizedToken>
oder
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. Fruchtknoten fast kahl. -
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:
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Sommer.
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<emphasis id="1695787940780ED455EEECF998B248E3" italics="true" pageId="null" pageNumber="433">Zytologische Angaben.</emphasis>
Keine Untersuchungen.
</paragraph>
<paragraph id="83EDA22BD2EF3B92064F2BFAF836A06A" pageId="null" pageNumber="433">
<emphasis id="0BC28895C9BD30BE94AE00C46DBB0601" italics="true" pageId="null" pageNumber="433">Standort.</emphasis>
Vorwiegend kollin. Humose
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.
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, Lichtungen,
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.
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Verbreitung.
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Pflanze:
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, Frankreich, Belgien, Westdeutschland, Schweiz. - Im Gebiet im Mittelland, ziemlich selten, Oberrheinische Tiefebene, Schaffhausen, Vorderrheintal.
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View file

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Syllis beneliahuae (Campoy &amp;
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, 1982)
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<paragraph id="7ADC2185694251340B173BCA03BE4E78" pageId="0" pageNumber="20997">Notes</paragraph>
<paragraph id="A1A8FA8093BDAB9AB3A2F1771A9B46F3" pageId="0" pageNumber="20997">Type locality: Mediterranean (Spain).</paragraph>
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