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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1110.80808" ID-Pensoft-Pub="1313-2970-1110-151" ID-Pensoft-UUID="40031A8835615F5D91BEB36B2B28DA1F" ID-ZooBank="7311F29E987840FE935B6B1E061262B2" ModsDocID="1313-2970-1110-151" checkinTime="1657093046334" checkinUser="pensoft" docAuthor="Cumming, Royce T. &amp; Le Tirant, Ste ́ phane" docDate="2022" docId="AAE955A2E02F54708E1ADCB6CCCE4765" docLanguage="en" docName="ZooKeys 1110: 151-200" docOrigin="ZooKeys 1110" docPubDate="2022-07-05" docSource="http://dx.doi.org/10.3897/zookeys.1110.80808" docTitle="Rakaphyllium exsectum Cumming &amp; Le Tirant 2022, comb. nov." docType="treatment" docVersion="1" id="40031A8835615F5D91BEB36B2B28DA1F" lastPageNumber="151" masterDocId="40031A8835615F5D91BEB36B2B28DA1F" masterDocTitle="Three new genera and one new species of leaf insect from Melanesia (Phasmatodea, Phylliidae)" masterLastPageNumber="200" masterPageNumber="151" pageNumber="151" updateTime="1657093046334" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Three new genera and one new species of leaf insect from Melanesia (Phasmatodea, Phylliidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cumming, Royce T.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7930-1292</mods:nameIdentifier>
<mods:affiliation>Montreal Insectarium, 4101 rue Sherbrooke est, Montre ́ al, Que ́ bec, H 1 X 2 B 2, Canada &amp; Richard Gilder Graduate School, American Museum of Natural History, New York, NY 10024, USA &amp; Biology, Graduate Center, City University of New York, NY, USA</mods:affiliation>
<mods:nameIdentifier type="email">roycecumming@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Le Tirant, Ste ́ phane</mods:namePart>
<mods:affiliation>Montreal Insectarium, 4101 rue Sherbrooke est, Montre ́ al, Que ́ bec, H 1 X 2 B 2, Canada</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-07-05</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1110</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>151</mods:start>
<mods:end>200</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1110.80808</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1110.80808</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1110-151</mods:identifier>
<mods:identifier type="ZooBank">7311F29E987840FE935B6B1E061262B2</mods:identifier>
<mods:identifier type="Pensoft-UUID">40031A8835615F5D91BEB36B2B28DA1F</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:AAE955A2E02F54708E1ADCB6CCCE4765" httpUri="http://treatment.plazi.org/id/AAE955A2E02F54708E1ADCB6CCCE4765" lastPageNumber="151" pageId="0" pageNumber="151">
<subSubSection pageId="0" pageNumber="151" type="nomenclature">
<paragraph pageId="0" pageNumber="151">
<taxonomicName LSID="AAE955A2-E02F-5470-8E1A-DCB6CCCE4765" authority="(Zompro, 2001 b)" authorityName="Cumming &amp; Le Tirant" authorityYear="2022" baseAuthorityName="Zompro" baseAuthorityYear="2001" family="Phylliidae" genus="Rakaphyllium" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rakaphyllium exsectum" order="Phasmatodea" pageId="0" pageNumber="151" rank="species" species="exsectum" status="comb. nov.">Rakaphyllium exsectum (Zompro, 2001b)</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="151">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="151" type="description">
<paragraph pageId="0" pageNumber="151">
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Rakaphyllium exsectum comb. nov. holotype female (NHMUK) A habitus, dorsal B profemora, dorsal C details of head through thorax, dorsal D genitalia, ventral E details of antenna, dorsal F associated data labels G details of head through thorax, dorsolateral. Scale bar only associated with A. Photographs A, F, G taken by Paul Brock (United Kingdom), other photographs taken by first author." figureDoi="10.3897/zookeys.1110.80808.figure9" httpUri="https://binary.pensoft.net/fig/710968" pageId="0" pageNumber="151">Fig. 9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="151" type="materials_examined">
<paragraph pageId="0" pageNumber="151">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="151">
<materialsCitation collectingDate="2000-10" determinerName="O. Zompro" location="Rawlinson Mts." specimenCount="♀" typeStatus="Holotype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="151">
<typeStatus>Holotype</typeStatus>
</emphasis>
(
<specimenCount>♀</specimenCount>
): &quot;HOLO-TYPE;
<location LSID="urn:lsid:plazi:treatment:AAE955A2E02F54708E1ADCB6CCCE4765:24131F78276FE3E64C5C328A5E77A506" name="Rawlinson Mts.">Rawlinson Mts.</location>
inland Huon Gulf. (Keysser).; Rothschild Bequest B. M.
<collectingDate bridgedPair="-" value="1939-01">1939-I.</collectingDate>
; Rawlinson Mt;
<taxonomicName authorityName="Zompro" authorityYear="2001" class="Insecta" family="Phylliidae" genus="Phyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllium exsectum" order="Phasmatodea" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="exsectum">
<emphasis italics="true" pageId="0" pageNumber="151">Phyllium exsectum</emphasis>
</taxonomicName>
<specimenCount>♀</specimenCount>
Zompro n. sp.
<typeStatus>HOLOTYPUS</typeStatus>
det.
<determinerName>O. Zompro</determinerName>
<collectingDate value="2000-10">X.2000</collectingDate>
; BMNH(E) #845235&quot; (NHMUK; Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Rakaphyllium exsectum comb. nov. holotype female (NHMUK) A habitus, dorsal B profemora, dorsal C details of head through thorax, dorsal D genitalia, ventral E details of antenna, dorsal F associated data labels G details of head through thorax, dorsolateral. Scale bar only associated with A. Photographs A, F, G taken by Paul Brock (United Kingdom), other photographs taken by first author." figureDoi="10.3897/zookeys.1110.80808.figure9" httpUri="https://binary.pensoft.net/fig/710968" pageId="0" pageNumber="151">9</figureCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="151" type="remarks">
<paragraph pageId="0" pageNumber="151">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="151">
This remarkable looking species is unfortunately only known from the singular holotype female from the little accessed Rawlinson Mountains of the Huon Peninsula (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distribution map noting all presently known records of Vaabonbonphyllium gen. nov. (denoted as star symbols) and Rakaphyllium gen. nov. (denoted as circle symbols) which were located and had data that could accurately be mapped. See Supplementary File 1 for full details for all records presented." figureDoi="10.3897/zookeys.1110.80808.figure5" httpUri="https://binary.pensoft.net/fig/710964" pageId="0" pageNumber="151">5</figureCitation>
). Even in other phylliid species with prominent lobes on the abdomen, they pale in comparison to the unique habitus of this species, which likely evolved to mimic some strongly lobed host plant. Unfortunately, as this is the only specimen known and it lacks ecological data, nothing is known about a possible plant that the unique shape of this species may have evolved to resemble through advergent evolution.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="151" type="differentiation">
<paragraph pageId="0" pageNumber="151">Differentiation.</paragraph>
<paragraph pageId="0" pageNumber="151">
At present only the female is known for this species, no tentative males have been found which could represent the opposite sex. This species can be differentiated from its single other congenic by the elaborate lobed abdomen (although some
<taxonomicName authorityName="Cumming &amp; Le Tirant" authorityYear="2022" baseAuthorityName="Giglio-Tos" baseAuthorityYear="1912" family="Phylliidae" genus="Rakaphyllium" kingdom="Animalia" lsidName="Rakaphyllium schultzei" order="Phasmatodea" pageId="0" pageNumber="151" rank="species" species="schultzei">
<emphasis italics="true" pageId="0" pageNumber="151">Rakaphyllium schultzei</emphasis>
</taxonomicName>
-like females have been observed with slight lobes suggesting morphological plasticity within that species) or more reliably by the profemoral exterior lobe, which in
<taxonomicName authorityName="Cumming &amp; Le Tirant" authorityYear="2022" baseAuthorityName="Zompro" baseAuthorityYear="2001" family="Phylliidae" genus="Rakaphyllium" kingdom="Animalia" lsidName="Rakaphyllium exsectum" order="Phasmatodea" pageId="0" pageNumber="151" rank="species" species="exsectum">
<emphasis italics="true" pageId="0" pageNumber="151">Rakaphyllium exsectum</emphasis>
</taxonomicName>
comb. nov. is right angled (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Rakaphyllium exsectum comb. nov. holotype female (NHMUK) A habitus, dorsal B profemora, dorsal C details of head through thorax, dorsal D genitalia, ventral E details of antenna, dorsal F associated data labels G details of head through thorax, dorsolateral. Scale bar only associated with A. Photographs A, F, G taken by Paul Brock (United Kingdom), other photographs taken by first author." figureDoi="10.3897/zookeys.1110.80808.figure9" httpUri="https://binary.pensoft.net/fig/710968" pageId="0" pageNumber="151">9B</figureCitation>
), versus
<taxonomicName authorityName="Cumming &amp; Le Tirant" authorityYear="2022" baseAuthorityName="Giglio-Tos" baseAuthorityYear="1912" family="Phylliidae" genus="Rakaphyllium" kingdom="Animalia" lsidName="Rakaphyllium schultzei" order="Phasmatodea" pageId="0" pageNumber="151" rank="species" species="schultzei">
<emphasis italics="true" pageId="0" pageNumber="151">Rakaphyllium schultzei</emphasis>
</taxonomicName>
which always has the profemoral exterior lobe roundly arcing from end to end without a distinct angle present (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Rakaphyllium schultzei comb. nov. holotype female. Note that the holotype female is heavily damaged in several places, especially the abdomen which has numerous holes through which the background can be seen, this is not an artifact of image postprocessing A habitus, dorsal B right front leg, dorsal C associated data labels D details of head through thorax, dorsal. Scale bar only associated with A. Photographs by Arne Koehler (SDEI) and Mandy Schroeter (SDEI)." figureDoi="10.3897/zookeys.1110.80808.figure6" httpUri="https://binary.pensoft.net/fig/710965" pageId="0" pageNumber="151">6B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="151" type="distribution">
<paragraph pageId="0" pageNumber="151">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="151">
The type locality of Rawlinson Mts. on the Huon Gulf, Morobe Province, Papua New Guinea is the only record known at present (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distribution map noting all presently known records of Vaabonbonphyllium gen. nov. (denoted as star symbols) and Rakaphyllium gen. nov. (denoted as circle symbols) which were located and had data that could accurately be mapped. See Supplementary File 1 for full details for all records presented." figureDoi="10.3897/zookeys.1110.80808.figure5" httpUri="https://binary.pensoft.net/fig/710964" pageId="0" pageNumber="151">5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:titleInfo id="FB2BF96BCBBAE48CB9C0E359AF0D8043">
<mods:title id="D16EF834864033AE632495638D5A5ECA">Chapter 7: Linnaean Plant Names and their Types (part C)</mods:title>
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<mods:role id="01BA4D03AAA19BDA15A29FA1E2D3EB5D">
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</mods:role>
<mods:namePart id="C18738E5110B69B6DCBB8AB40E5C850E">Jarvis, Charlie</mods:namePart>
<mods:affiliation id="8B4DB9032FBD3E4A4010C214658ABB61">Department of Botany, Natural History Museum, Cromwell Road, London, UK</mods:affiliation>
</mods:name>
<mods:typeOfResource id="45E835C8D489D4C96E547AFBC18EB5F0">text</mods:typeOfResource>
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<mods:title id="32997DB4B5239210E335F535DCB04349">Order out of Chaos. Linnaean Plant Types and their Types</mods:title>
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<treatment id="AAE96A0455779957D6C6F4C14FB70AAB" ID-DOI="http://doi.org/10.5281/zenodo.4379355" ID-GBIF-Taxon="164003830" ID-Zenodo-Dep="4379355" LSID="urn:lsid:plazi:treatment:AAE96A0455779957D6C6F4C14FB70AAB" httpUri="http://treatment.plazi.org/id/AAE96A0455779957D6C6F4C14FB70AAB" lastPageNumber="414" pageNumber="414">
<subSubSection id="E7AB224163487D771232BF6061A9CF2B" pageNumber="414" type="reference_group">
<paragraph id="0D028CE35BFAAD26E9AFC5C8AA00D890" pageNumber="414">
<treatmentCitation id="E28CA4AD03993882829B44DC827E9F9C" author="Linnaeus" pageNumber="563" year="1767">
<bibCitation id="46E186F0605DBC14B3AC378FBEF81B9F" author="Linnaeus" pagination="563" title="Chrysanthemum italicum" volumeTitle="Systema Naturae, ed. 12, 2" year="1767">
<taxonomicName id="1E36347BD3B27EF6BA0C700527FDBFC2" authority="Linnaeus" authorityName="Linnaeus" authorityYear="1767" class="Magnoliopsida" family="Asteraceae" genus="Chrysanthemum" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="italicum">
<emphasis id="969A32A27CA7405B00E935175DD8AE11" bold="true" pageNumber="414">Chrysanthemum italicum</emphasis>
Linnaeus
</taxonomicName>
,
<volumeTitle id="DA41AFF3745C272342C71D5247263CD0">
<emphasis id="08C02B0E9FA5B5B1AE711C14A747AC2F" italics="true" pageNumber="414">Systema Naturae</emphasis>
, ed. 12, 2
</volumeTitle>
: 563;
<emphasis id="18D88C9BBC3E44ABE4477CE43A631750" italics="true" pageNumber="414">
<volumeTitle id="032FFF21C8A5CFB963AEDEFA1B9CFB25">Mantissa Plantarum</volumeTitle>
</emphasis>
: 116. 1767
</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E245641FC60D4F0B6D4176A81D0C72A6" pageNumber="414" type="distribution">
<paragraph id="4C9DBF95ECEC2B03FD16C57D70BFD8D7" pageNumber="414">&quot;Habitat in Italia. Arduini.&quot; RCN: 6445.</paragraph>
</subSubSection>
<subSubSection id="EBF8471E9CD62171B9287A7F0FB38933" pageNumber="414" type="materials_examined">
<paragraph id="10A0769F83170460D05998BBD901B95D" pageNumber="414">Type not designated.</paragraph>
</subSubSection>
<subSubSection id="6F94C9784CB2338D707F3740999A00AF" pageNumber="414" type="materials_examined">
<paragraph id="C9831123C7E33DA9E84BE3AB9A475FCD" pageNumber="414">
Original material:
<emphasis id="3A3EBEBD910A8E8A0CAD43E8FD0AFC20" italics="true" pageNumber="414">Arduino 69</emphasis>
,
<materialsCitation id="F125B27BE2B85A5C64B5D6BCBAD29891" ID-GBIF-Occurrence="2612072839" collectionCode="LINN" specimenCode="1012.21">
Herb. Linn. No. 1012.21 (
<collectionCode id="CD9325C56066EB01F79EAB28225FD53B">LINN</collectionCode>
)
</materialsCitation>
; [icon] in Micheli, Nov. Pl. Gen.: 34, t. 29. 1729. -Voucher:
<materialsCitation id="C6B2AB1675DBCA5BE83B33189D505B81" ID-GBIF-Occurrence="2612072352" collectionCode="FI" specimenCode="2809">
Herb. Micheli 2809, III: 584, sub
<emphasis id="66CE1ED7798AC3920918610C3E7C243C" italics="true" pageNumber="414">Pyrethrum</emphasis>
, 2810 (
<collectionCode id="193749675CAAB4BEB80872334A35AF17">FI</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="949B6601654550FB9A2398686218175D" pageNumber="414" type="nomenclature">
<paragraph id="D065F3E371D91BCB5265EB4E8A8A5781" pageNumber="414">
Current name:
<taxonomicName id="C6164719F3B179DD220BC396D76C249F" ID-CoL="7N2LJ" authority="(Willd.) Briq. &amp; Cavill." authorityName="Greuter" authorityYear="2003" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" class="Magnoliopsida" family="Asteraceae" genus="Tanacetum" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="variety" species="corymbosum" variety="tenuifolium">
<emphasis id="AFEB61FBD8B98D9CF1A34931016302CE" italics="true" pageNumber="414">
<emphasis id="E587CA2F23CE708A06927AF8FBE7CE50" bold="true" italics="true" pageNumber="414">Tanacetum corymbosum</emphasis>
</emphasis>
(L.) Sch. Bip. var.
<emphasis id="D12830D7FA3A92DF0BCC89C3BCB4C131" italics="true" pageNumber="414">
<emphasis id="765224F6015ECB61F0C7D6870C8CBCF9" bold="true" italics="true" pageNumber="414">tenuifolium</emphasis>
</emphasis>
(Willd.) Briq. &amp; Cavill.
</taxonomicName>
(
<taxonomicName id="63D65925A84B425C403D343D62B3EDBB" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C32D78B80C53785187E09B1E66853484" pageNumber="414" type="discussion">
<paragraph id="E0C16B7C1A5DFC75A2C338C59E5030AF" pageNumber="414">
<emphasis id="58DABDF0731E0ED5BED88B313972E6F5" italics="true" pageNumber="414">Note:</emphasis>
The Micheli reference, an original element for this name, was also cited simultaneously in the synonymy of
<taxonomicName id="180B1DA6696147B3E3D3A52CDFAD2357" authority="L., Syst. Nat." authorityName="Linnaeus" authorityYear="1767" class="Magnoliopsida" family="Asteraceae" genus="Chrysanthemum" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="achilleae">
<emphasis id="3A40F0098587EAF7036C728F2EA0B36C" italics="true" pageNumber="414">C. achilleae</emphasis>
L.,
<emphasis id="084157EE0CC490B1FBBBEA3F40823CEF" italics="true" pageNumber="414">Syst. Nat.</emphasis>
</taxonomicName>
, ed. 12, 2: 562 (1767).
</paragraph>
</subSubSection>
</treatment>
</document>