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<document id="B8C539869C11DFEF986ADE341D4376BF" ID-DOI="10.11646/zootaxa.3796.1.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="4915127" ID-ZooBank="31C2F235-AAC7-4BBE-ABEB-793AFF63C80C" IM.bibliography_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1623198400352" checkinUser="felipe" docAuthor="Botero-Trujillo, Ricardo & Flórez, Eduardo" docDate="2014" docId="E219AA01FFCC893DFF766DC8D1C1C59F" docLanguage="en" docName="zt03796p120.pdf" docOrigin="Zootaxa 3796 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tityus (Tityus) grottoedensis Botero-Trujillo & Flórez 2014, sp. nov." docType="treatment" docVersion="1" lastPageNumber="116" masterDocId="1E20D279FFCE8934FFE16D5FD016C54A" masterDocTitle="A new species of Tityus (Scorpiones, Buthidae) from El Edén Cave, Colombia" masterLastPageNumber="120" masterPageNumber="108" pageNumber="110" updateTime="1731061025806" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:mods id="2A4A7640A1E94BBB43DF685CE34053F6" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="BE696F4D9591F58B0BFE574E67CCA12D">
|
||||
<mods:title id="90D03C06709CC5E60F6C461C0EC31B84">A new species of Tityus (Scorpiones, Buthidae) from El Edén Cave, Colombia</mods:title>
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||||
</mods:titleInfo>
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<mods:name id="A6033AAB483BCD7198B0E8759B918908" type="personal">
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<mods:role id="94519E4DF77B877F87F48B7741CA5358">
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<mods:roleTerm id="2D01AAA08F98976203BF1D9AD4F46791">Author</mods:roleTerm>
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||||
</mods:role>
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<mods:namePart id="549D19FDB9FDF96E8AF726C59E590F1A">Botero-Trujillo, Ricardo</mods:namePart>
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||||
<mods:affiliation id="23CD9E60B29746E1124F0A007D75B9DD">Associate researcher. Laboratorio de Entomología, Unidad de Ecología y Sistemática-UNESIS, Departamento de Biología, Pontificia Universidad Javeriana, Bogotá, Colombia</mods:affiliation>
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<mods:roleTerm id="62C16B755E0EBA59D9A0D4F4E2F06718">Author</mods:roleTerm>
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||||
</mods:role>
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||||
<mods:namePart id="8C43AC1112F1A379C9E4303732232931">Flórez, Eduardo</mods:namePart>
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||||
<mods:affiliation id="8C443F68BDCCE01ECF7BC74F82A00249">Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá, Colombia. E-mail: aeflorezd @ unal. edu. co</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="E23931E138E6C9A0F8A9D5A385F0F3B3">text</mods:typeOfResource>
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<mods:title id="7FA8137D72119925B9BE995C3CEAAAF2">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B69384041421E2F085D18117EE821FC0">
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<mods:date id="0907E495306B8F56435B0911CE8EFF13">2014</mods:date>
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<mods:number id="89549ABBDA2956895D5753BD79998355">2014-05-16</mods:number>
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<mods:detail id="0E53F12A85618E0A3FE0FC569DCABBA4" type="volume">
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<mods:number id="65B0C2AFCA69401841DE9F079095CE60">3796</mods:number>
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<mods:detail id="2ED5ACC5B23FAB4DBC930275E098481C" type="issue">
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<mods:number id="B4782F23DF735133E09CD77061345E9A">1</mods:number>
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<mods:start id="B2D9878E7D6DB569BD4F1EF586C34621">108</mods:start>
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<mods:classification id="D80DA2FC74E049CF11336FE143CA84DC">journal article</mods:classification>
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<mods:identifier id="9AE5E0F95669110E53D5292EA938F293" type="DOI">10.11646/zootaxa.3796.1.5</mods:identifier>
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<mods:identifier id="A8CC5E52ED1611A944D731060302DD8E" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="FDB7836CC8B3312FA74FDE411F38948B" type="Zenodo-Dep">4915127</mods:identifier>
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<mods:identifier id="335550989E41F8DB68DBC04D0B231504" type="ZooBank">31C2F235-AAC7-4BBE-ABEB-793AFF63C80C</mods:identifier>
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</mods:mods>
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||||
<treatment id="E219AA01FFCC893DFF766DC8D1C1C59F" LSID="urn:lsid:plazi:treatment:E219AA01FFCC893DFF766DC8D1C1C59F" httpUri="http://treatment.plazi.org/id/E219AA01FFCC893DFF766DC8D1C1C59F" lastPageId="9" lastPageNumber="116" pageId="2" pageNumber="110">
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||||
<subSubSection id="22AA489CFFCC8936FF766DC8D22DC5E5" box="[151,571,151,176]" pageId="2" pageNumber="110" type="nomenclature">
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<paragraph id="6A0F1B17FFCC8936FF766DC8D22DC5E5" blockId="2.[151,571,151,213]" box="[151,571,151,176]" pageId="2" pageNumber="110">
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||||
<taxonomicName id="ADB06094FFCC8936FF766DC8D1CAC5FA" ID-CoL="7CDMG" authority="Botero-Trujillo & Flórez, 2014" authorityName="Botero-Trujillo & Flórez" authorityYear="2014" box="[151,476,151,176]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="110" phylum="Arthropoda" rank="species" species="grottoedensis" status="sp. nov." subGenus="Tityus">
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||||
<emphasis id="58C4C705FFCC8936FF766DC8D1CAC5FA" bold="true" box="[151,476,151,176]" italics="true" pageId="2" pageNumber="110">Tityus (Tityus) grottoedensis</emphasis>
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||||
</taxonomicName>
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||||
<taxonomicNameLabel id="43F77A7EFFCC8936FE056DC7D22DC5E5" box="[484,571,152,175]" pageId="2" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
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||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="22AA489CFFCC8936FF766DE3D19DC59F" box="[151,395,188,213]" pageId="2" pageNumber="110" type="description">
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<paragraph id="6A0F1B17FFCC8936FF766DE3D19DC59F" blockId="2.[151,571,151,213]" box="[151,395,188,213]" pageId="2" pageNumber="110">
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||||
<figureCitation id="F28B0792FFCC8936FF766DE3D13BC59F" box="[151,301,188,213]" captionStart-0="FIGURES 1–3" captionStart-1="FIGURES 4–7" captionStart-2="FIGURES 8–13" captionStart-3="FIGURES 14–16" captionStartId-0="2.[151,264,1638,1661]" captionStartId-1="5.[151,264,1830,1853]" captionStartId-2="6.[151,264,1320,1343]" captionStartId-3="7.[151,264,1801,1824]" captionTargetBox-0="[151,1433,640,1604]" captionTargetBox-1="[178,1424,213,1778]" captionTargetBox-2="[159,1432,213,1269]" captionTargetBox-3="[158,1428,456,1773]" captionTargetId-0="figure-343@2.[151,1436,625,1617]" captionTargetId-1="figure-60@5.[151,1436,193,1809]" captionTargetId-2="figure-295@6.[151,1436,193,1299]" captionTargetId-3="figure-223@7.[151,1436,445,1780]" captionTargetPageId-0="2" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURES 1–3. Distribution of Tityus (Tityus) grottoedensis sp. nov. 1. Map showing the location of El Edén Cave in Colombia. 2. Entrance to El Edén Cave from the opposite riverside. 3. Interior passage of El Edén cave. Photographs courtesy of Miguel A. Medrano (Fig. 2) and Iván C. Gonzalez (Fig. 3)." captionText-1="FIGURES 4–7. Tityus (Tityus) grottoedensis sp. nov., habitus. 4–5. Subadult paratype male, dorsal and ventral aspects (ICN- As-905). 6–7. One paratype female, dorsal and ventral aspects (MACN-Ar 30879). Scale bars = 10 mm." captionText-2="FIGURES 8–13. Tityus (Tityus) grottoedensis sp. nov., subadult paratype male (ICN-As-905), trichobothrial distribution in right pedipalp. 8. Femur, dorsal aspect. 9. Femur, internal aspect. 10. Patella, dorsal aspect. 11. Patella, external aspect. 12. Chela, external aspect. 13. Chela, ventral aspect (palm). Scale bars = 1 mm." captionText-3="FIGURES 14–16. Tityus (Tityus) grottoedensis sp. nov. 14. Subadult paratype male (ICN-As-905), right lateral aspect of metasoma and telson. 15–16. Holotype female (ICN-As-977). 15. Left pectine. 16. Sternites IV–VII; most granulation details of the sternites not depicted, emphasis made on carinae. Scale bars = 3 mm (Figs. 14, 16), 1 mm (Fig. 15)." figureDoi-0="http://doi.org/10.5281/zenodo.4915129" figureDoi-1="http://doi.org/10.5281/zenodo.4915131" figureDoi-2="http://doi.org/10.5281/zenodo.4915133" figureDoi-3="http://doi.org/10.5281/zenodo.4915135" httpUri-0="https://zenodo.org/record/4915129/files/figure.png" httpUri-1="https://zenodo.org/record/4915131/files/figure.png" httpUri-2="https://zenodo.org/record/4915133/files/figure.png" httpUri-3="https://zenodo.org/record/4915135/files/figure.png" pageId="2" pageNumber="110">Figures 1–16</figureCitation>
|
||||
;
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||||
<tableCitation id="27322EACFFCC8936FED86DE3D19DC59F" box="[313,395,188,213]" captionStart="TABLE 1" captionStartId="4.[147,230,454,475]" captionTargetPageId="4" captionText="TABLE 1. Meristic data for Tityus (Tityus) grottoedensis sp. nov. Measurements in millimeters. 1Sum of carapace, mesosoma and metasoma. 2Sum of tergites I– VII. 3Sum of metasomal segments I–V and telson. 4Sum of femur, patella and chela. 5Measured from the commissure of the junction with the movable finger to the finger tip. 6Measured along the marginal lamella." httpUri="http://table.plazi.org/id/3ECF4B9FFFCA8930FF726C99D281C75E" pageId="2" pageNumber="110" tableUuid="3ECF4B9FFFCA8930FF726C99D281C75E">Table 1</tableCitation>
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||||
</paragraph>
|
||||
</subSubSection>
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||||
<subSubSection id="22AA489CFFCC8936FF766C5BD2DAC777" pageId="2" pageNumber="110" type="materials_examined">
|
||||
<paragraph id="6A0F1B17FFCC8936FF766C5BD2DAC777" blockId="2.[151,1436,260,573]" pageId="2" pageNumber="110">
|
||||
<emphasis id="58C4C705FFCC8936FF766C5BD150C457" bold="true" box="[151,326,260,285]" inLineHeading="true" pageId="2" pageNumber="110" reason="1">Type material.</emphasis>
|
||||
<materialsCitation id="DAD8114AFFCC8936FEAE6C5BD224C42F" collectingDate="2010-02-06" collectorName="A. Perez Gonzalez" country="Colombia" county="Cunday" elevation="650" latitude="4.0119667" location="Vereda La Camelia" longLatPrecision="1" longitude="-74.75515" pageId="2" pageNumber="110" specimenCode="ICN-As-977" specimenCount="2" specimenCount-adult="1" specimenCount-female="1" stateProvince="Tolima department" typeStatus="holotype">
|
||||
<collectingCountry id="12A75B87FFCC8936FEAE6C5BD1F1C457" box="[335,487,260,285]" name="Colombia" pageId="2" pageNumber="110">COLOMBIA</collectingCountry>
|
||||
,
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||||
<collectingRegion id="A874D5F5FFCC8936FE156C5BD2DBC457" box="[500,717,260,285]" country="Colombia" name="Tolima" pageId="2" pageNumber="110">Tolima department</collectingRegion>
|
||||
,
|
||||
<collectingCounty id="836E639BFFCC8936FD386C5BD327C457" box="[729,817,260,285]" pageId="2" pageNumber="110">Cunday</collectingCounty>
|
||||
,
|
||||
<location id="6F6F4DCCFFCC8936FCDF6C5BD40BC457" LSID="urn:lsid:plazi:treatment:E219AA01FFCC893DFF766DC8D1C1C59F:6F6F4DCCFFCC8936FCDF6C5BD40BC457" box="[830,1053,260,285]" country="Colombia" county="Cunday" latitude="4.0119667" longLatPrecision="1" longitude="-74.75515" name="Vereda La Camelia" pageId="2" pageNumber="110" stateProvince="Tolima department">Vereda La Camelia</location>
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||||
:
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||||
<emphasis id="58C4C705FFCC8936FBC86C5BD556C457" box="[1065,1344,260,285]" italics="true" pageId="2" pageNumber="110">
|
||||
<typeStatus id="B50BA5B5FFCC8936FBC86C5BD499C457" box="[1065,1167,260,285]" pageId="2" pageNumber="110" type="holotype">Holotype</typeStatus>
|
||||
<specimenCount id="7CB6D09EFFCC8936FB796C5BD4F5C457" box="[1176,1251,260,285]" pageId="2" pageNumber="110" type="female">female</specimenCount>
|
||||
(
|
||||
<specimenCount id="7CB6D09EFFCC8936FB156C5BD524C457" box="[1268,1330,260,285]" pageId="2" pageNumber="110" type="adult">adult</specimenCount>
|
||||
):
|
||||
</emphasis>
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aphotic zone of
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||||
<location id="6F6F4DCCFFCC8936FF136C78D19BC40A" LSID="urn:lsid:plazi:treatment:E219AA01FFCC893DFF766DC8D1C1C59F:6F6F4DCCFFCC8936FF136C78D19BC40A" box="[242,397,295,320]" country="Colombia" county="Cunday" latitude="4.0119667" longLatPrecision="1" longitude="-74.75515" municipality="Vereda La Camelia" name="El Eden Cave" pageId="2" pageNumber="110" stateProvince="Tolima department">El Edén Cave</location>
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||||
at
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||||
<quantity id="AD48B6F2FFCC8936FE516C78D1EEC475" box="[432,504,295,320]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="110" unit="m" value="200.0">200 m</quantity>
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depth, under a stone,
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<geoCoordinate id="0F847DD0FFCC8936FD106C78D391C475" box="[753,903,295,320]" degrees="04" direction="north" minutes="00.718" orientation="latitude" pageId="2" pageNumber="110" precision="1" value="4.0119667">04º00.718´N</geoCoordinate>
|
||||
<geoCoordinate id="0F847DD0FFCC8936FC6E6C78D43AC475" box="[911,1068,295,320]" degrees="74" direction="west" minutes="45.309" orientation="longitude" pageId="2" pageNumber="110" precision="1" value="-74.75515">74º45.309´W</geoCoordinate>
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||||
, elev.
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||||
<quantity id="AD48B6F2FFCC8936FB926C78D4AAC475" box="[1139,1212,295,320]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="110" unit="m" value="650.0">
|
||||
<elevation id="E19DFC24FFCC8936FB926C78D4AAC475" box="[1139,1212,295,320]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="110" unit="m" value="650.0">650 m</elevation>
|
||||
</quantity>
|
||||
,
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||||
<date id="1E0E3DD7FFCC8936FB286C78D58EC40A" box="[1225,1432,295,320]" pageId="2" pageNumber="110" value="2010-02-06">
|
||||
<collectingDate id="0E4AC43FFFCC8936FB286C78D58EC40A" box="[1225,1432,295,320]" pageId="2" pageNumber="110" value="2010-02-06">06-February-2010</collectingDate>
|
||||
</date>
|
||||
;
|
||||
<collectorName id="C7457EC1FFCC8936FF766C13D16DC42F" box="[151,379,332,357]" pageId="2" pageNumber="110">A. Pérez González</collectorName>
|
||||
(
|
||||
<specimenCode id="3A16B36CFFCC8936FE726C13D230C42F" box="[403,550,332,357]" pageId="2" pageNumber="110">ICN-As-977</specimenCode>
|
||||
).
|
||||
</materialsCitation>
|
||||
<materialsCitation id="DAD8114AFFCC8936FDA36C12D0FAC4C2" pageId="2" pageNumber="110" specimenCode="MACN-Ar 30879" specimenCount="2" specimenCount-adult="1" specimenCount-female="1" typeStatus="paratype">
|
||||
<emphasis id="58C4C705FFCC8936FDA36C12D2AAC42E" box="[578,700,333,356]" italics="true" pageId="2" pageNumber="110">
|
||||
<typeStatus id="B50BA5B5FFCC8936FDA36C12D2A1C42E" box="[578,695,333,356]" pageId="2" pageNumber="110" type="paratype">Paratypes</typeStatus>
|
||||
:
|
||||
</emphasis>
|
||||
One
|
||||
<specimenCount id="7CB6D09EFFCC8936FCEC6C13D353C42F" box="[781,837,332,357]" pageId="2" pageNumber="110" type="adult">adult</specimenCount>
|
||||
<specimenCount id="7CB6D09EFFCC8936FCB46C13D3B7C42F" box="[853,929,332,357]" pageId="2" pageNumber="110" type="female">female</specimenCount>
|
||||
with same data of holotype (
|
||||
<specimenCode id="3A16B36CFFCC8936FAFE6C13D0F6C4C2" pageId="2" pageNumber="110">MACN-Ar 30879</specimenCode>
|
||||
).
|
||||
</materialsCitation>
|
||||
<materialsCitation id="DAD8114AFFCC8936FF0A6C30D470C4E7" collectingDate="2010-02-06" collectorName="A. Perez Gonzalez & L. Benavides & D. A. Luna" elevation="650" latitude="4.0119667" location="El Eden Cave" longLatPrecision="1" longitude="-74.75515" pageId="2" pageNumber="110" specimenCode="ICN-As-905" specimenCount="2" specimenCount-male="1" specimenCount-subadult="1">
|
||||
One
|
||||
<specimenCount id="7CB6D09EFFCC8936FEC56C30D195C4C2" box="[292,387,367,392]" pageId="2" pageNumber="110" type="subadult">subadult</specimenCount>
|
||||
<specimenCount id="7CB6D09EFFCC8936FE6A6C30D1D7C4C2" box="[395,449,367,392]" pageId="2" pageNumber="110" type="male">male</specimenCount>
|
||||
from outside of
|
||||
<location id="6F6F4DCCFFCC8936FD616C30D309C4C2" LSID="urn:lsid:plazi:treatment:E219AA01FFCC893DFF766DC8D1C1C59F:6F6F4DCCFFCC8936FD616C30D309C4C2" box="[640,799,367,392]" country="Colombia" county="Cunday" latitude="4.0119667" longLatPrecision="1" longitude="-74.75515" municipality="Vereda La Camelia" name="El Eden Cave" pageId="2" pageNumber="110" stateProvince="Tolima">El Edén Cave</location>
|
||||
,
|
||||
<emphasis id="58C4C705FFCC8936FCCD6C30D36EC4C2" box="[812,888,367,392]" italics="true" pageId="2" pageNumber="110">ad hoc</emphasis>
|
||||
,
|
||||
<geoCoordinate id="0F847DD0FFCC8936FC666C30D409C4CD" box="[903,1055,367,392]" degrees="04" direction="north" minutes="00.718" orientation="latitude" pageId="2" pageNumber="110" precision="1" value="4.0119667">04º00.718´N</geoCoordinate>
|
||||
<geoCoordinate id="0F847DD0FFCC8936FBC66C30D4D3C4CD" box="[1063,1221,367,392]" degrees="74" direction="west" minutes="45.309" orientation="longitude" pageId="2" pageNumber="110" precision="1" value="-74.75515">74º45.309´W</geoCoordinate>
|
||||
, at night, elev.
|
||||
<quantity id="AD48B6F2FFCC8936FA936C30D0B9C4E1" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="110" unit="m" value="650.0">
|
||||
<elevation id="E19DFC24FFCC8936FA936C30D0B9C4E1" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="110" unit="m" value="650.0">650 m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<date id="1E0E3DD7FFCC8936FF5D6CCBD19AC4E7" box="[188,396,404,429]" pageId="2" pageNumber="110" value="2010-02-06">
|
||||
<collectingDate id="0E4AC43FFFCC8936FF5D6CCBD19AC4E7" box="[188,396,404,429]" pageId="2" pageNumber="110" value="2010-02-06">06-February-2010</collectingDate>
|
||||
</date>
|
||||
;
|
||||
<collectorName id="C7457EC1FFCC8936FE796CCBD279C4E7" box="[408,623,404,429]" pageId="2" pageNumber="110">A. Pérez González</collectorName>
|
||||
,
|
||||
<collectorName id="C7457EC1FFCC8936FD9A6CCBD307C4E7" box="[635,785,404,429]" pageId="2" pageNumber="110">L. Benavides</collectorName>
|
||||
&
|
||||
<collectorName id="C7457EC1FFCC8936FCD66CCBD3AEC4E1" box="[823,952,404,429]" pageId="2" pageNumber="110">D. A. Luna</collectorName>
|
||||
(
|
||||
<specimenCode id="3A16B36CFFCC8936FC296CCBD44DC4E7" box="[968,1115,404,429]" pageId="2" pageNumber="110">ICN-As-905</specimenCode>
|
||||
).
|
||||
</materialsCitation>
|
||||
<materialsCitation id="DAD8114AFFCC8936FB8F6CCBD479C4BF" collectingDate="2010-09-07" collectorName="D. Luna" elevation="650" latitude="4.0119667" location="El Eden Cave" longLatPrecision="1" longitude="-74.75515" pageId="2" pageNumber="110" specimenCode="ICN-As-872" specimenCount="2" specimenCount-adult="1" specimenCount-female="1">
|
||||
One
|
||||
<specimenCount id="7CB6D09EFFCC8936FB446CCBD4CBC4E7" box="[1189,1245,404,429]" pageId="2" pageNumber="110" type="adult">adult</specimenCount>
|
||||
pregnant
|
||||
<specimenCount id="7CB6D09EFFCC8936FAB16CCBD58AC4E7" box="[1360,1436,404,429]" pageId="2" pageNumber="110" type="female">female</specimenCount>
|
||||
and 19 (subsequently born) newborns from the aphotic zone of
|
||||
<location id="6F6F4DCCFFCC8936FC8E6CE8D419C49A" LSID="urn:lsid:plazi:treatment:E219AA01FFCC893DFF766DC8D1C1C59F:6F6F4DCCFFCC8936FC8E6CE8D419C49A" box="[879,1039,439,464]" country="Colombia" county="Cunday" latitude="4.0119667" longLatPrecision="1" longitude="-74.75515" municipality="Vereda La Camelia" name="El Eden Cave" pageId="2" pageNumber="110" stateProvince="Tolima">El Edén Cave</location>
|
||||
,
|
||||
<emphasis id="58C4C705FFCC8936FBFD6CE8D47CC49A" box="[1052,1130,439,464]" italics="true" pageId="2" pageNumber="110">ad hoc</emphasis>
|
||||
while walking on the soil,
|
||||
<geoCoordinate id="0F847DD0FFCC8936FF766C83D138C4B9" box="[151,302,476,501]" degrees="04" direction="north" minutes="00.718" orientation="latitude" pageId="2" pageNumber="110" precision="1" value="4.0119667">04º00.718´N</geoCoordinate>
|
||||
<geoCoordinate id="0F847DD0FFCC8936FED66C83D1C3C4B9" box="[311,469,476,501]" degrees="74" direction="west" minutes="45.309" orientation="longitude" pageId="2" pageNumber="110" precision="1" value="-74.75515">74º45.309´W</geoCoordinate>
|
||||
, elev.
|
||||
<quantity id="AD48B6F2FFCC8936FDFC6C83D27EC4B9" box="[541,616,476,501]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="110" unit="m" value="650.0">
|
||||
<elevation id="E19DFC24FFCC8936FDFC6C83D27EC4B9" box="[541,616,476,501]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="110" unit="m" value="650.0">650 m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<date id="1E0E3DD7FFCC8936FD946C83D340C4BF" box="[629,854,476,501]" pageId="2" pageNumber="110" value="2010-09-07">
|
||||
<collectingDate id="0E4AC43FFFCC8936FD946C83D340C4BF" box="[629,854,476,501]" pageId="2" pageNumber="110" value="2010-09-07">07-September-2010</collectingDate>
|
||||
</date>
|
||||
;
|
||||
<collectorName id="C7457EC1FFCC8936FC826C83D3D6C4B9" box="[867,960,476,499]" pageId="2" pageNumber="110">D. Luna</collectorName>
|
||||
(
|
||||
<specimenCode id="3A16B36CFFCC8936FC316C83D475C4BF" box="[976,1123,476,501]" pageId="2" pageNumber="110">ICN-As-872</specimenCode>
|
||||
).
|
||||
</materialsCitation>
|
||||
<materialsCitation id="DAD8114AFFCC8936FB976C83D2DAC777" collectingDate="2012-11-16" collectionCode="ICN" collectorName="W. Galvis & A. Garcia & M. Medrano & C. Romero" elevation="550" location="El Eden Cave" pageId="2" pageNumber="110" specimenCode="ICN- As-837" specimenCount="1" specimenCount-juvenile="1">
|
||||
One
|
||||
<specimenCount id="7CB6D09EFFCC8936FB4F6C83D51EC4BF" box="[1198,1288,476,501]" pageId="2" pageNumber="110" type="juvenile">juvenile</specimenCount>
|
||||
(presumably male) from
|
||||
<location id="6F6F4DCCFFCC8936FEFC6CA0D1AAC752" LSID="urn:lsid:plazi:treatment:E219AA01FFCC893DFF766DC8D1C1C59F:6F6F4DCCFFCC8936FEFC6CA0D1AAC752" box="[285,444,511,536]" country="Colombia" county="Cunday" latitude="4.0119667" longLatPrecision="1" longitude="-74.75515" municipality="Vereda La Camelia" name="El Eden Cave" pageId="2" pageNumber="110" stateProvince="Tolima">El Edén Cave</location>
|
||||
, “
|
||||
<quantity id="AD48B6F2FFCC8936FE326CA0D236C752" box="[467,544,511,536]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="110" unit="m" value="550.0">
|
||||
<elevation id="E19DFC24FFCC8936FE326CA0D236C752" box="[467,544,511,536]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="110" unit="m" value="550.0">550 m</elevation>
|
||||
</quantity>
|
||||
”,
|
||||
<date id="1E0E3DD7FFCC8936FDD86CA0D30FC752" box="[569,793,511,536]" pageId="2" pageNumber="110" value="2012-11-16">
|
||||
<collectingDate id="0E4AC43FFFCC8936FDD86CA0D30FC752" box="[569,793,511,536]" pageId="2" pageNumber="110" value="2012-11-16">16-November-2012</collectingDate>
|
||||
</date>
|
||||
;
|
||||
<collectorName id="C7457EC1FFCC8936FCC46F5FD38EC752" box="[805,920,511,536]" pageId="2" pageNumber="110">W. Galvis</collectorName>
|
||||
,
|
||||
<collectorName id="C7457EC1FFCC8936FC446CA0D400C752" box="[933,1046,511,536]" pageId="2" pageNumber="110">A. García</collectorName>
|
||||
,
|
||||
<collectorName id="C7457EC1FFCC8936FBC36F5FD4B9C752" box="[1058,1199,511,536]" pageId="2" pageNumber="110">M. Medrano</collectorName>
|
||||
&
|
||||
<collectorName id="C7457EC1FFCC8936FB346CA0D547C75D" box="[1237,1361,511,536]" pageId="2" pageNumber="110">C. Romero</collectorName>
|
||||
(
|
||||
<specimenCode id="3A16B36CFFCC8936FA806CA0D0F8C777" pageId="2" pageNumber="110">ICN- As-837</specimenCode>
|
||||
). All specimens preserved in 70% ethanol.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="3ECF4B9FFFCC8936FF766B39D31EC3F0" ID-DOI="http://doi.org/10.5281/zenodo.4915129" ID-Zenodo-Dep="4915129" httpUri="https://zenodo.org/record/4915129/files/figure.png" pageId="2" pageNumber="110" startId="2.[151,264,1638,1661]" targetBox="[151,1433,640,1604]" targetPageId="2">
|
||||
<paragraph id="6A0F1B17FFCC8936FF766B39D31EC3F0" blockId="2.[151,1436,1638,1722]" pageId="2" pageNumber="110">
|
||||
<emphasis id="58C4C705FFCC8936FF766B39D129C336" bold="true" box="[151,319,1638,1661]" pageId="2" pageNumber="110">FIGURES 1–3.</emphasis>
|
||||
Distribution of
|
||||
<taxonomicName id="ADB06094FFCC8936FE106B38D30EC336" authority="Botero-Trujillo & Flórez, 2014" authorityName="Botero-Trujillo & Flórez" authorityYear="2014" box="[497,792,1638,1660]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="110" phylum="Arthropoda" rank="species" species="grottoedensis" status="sp. nov." subGenus="Tityus">
|
||||
<emphasis id="58C4C705FFCC8936FE106B38D30EC336" box="[497,792,1638,1660]" italics="true" pageId="2" pageNumber="110">Tityus (Tityus) grottoedensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="58C4C705FFCC8936FCC56B38D38FC336" bold="true" box="[804,921,1639,1660]" pageId="2" pageNumber="110">
|
||||
<taxonomicNameLabel id="43F77A7EFFCC8936FCC56B38D36CC336" box="[804,890,1639,1660]" pageId="2" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
1.
|
||||
</emphasis>
|
||||
Map showing the location of El Edén Cave in Colombia.
|
||||
<emphasis id="58C4C705FFCC8936FEE96BDAD10DC3D0" bold="true" box="[264,283,1669,1690]" pageId="2" pageNumber="110">2.</emphasis>
|
||||
Entrance to El Edén Cave from the opposite riverside.
|
||||
<emphasis id="58C4C705FFCC8936FCAE6BDAD374C3D0" bold="true" box="[847,866,1669,1690]" pageId="2" pageNumber="110">3.</emphasis>
|
||||
Interior passage of El Edén cave. Photographs courtesy of Miguel A. Medrano (Fig. 2) and Iván C. Gonzalez (Fig. 3).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="22AA489CFFCC8936FF266BBED1BDC208" pageId="2" pageNumber="110" type="etymology">
|
||||
<paragraph id="6A0F1B17FFCC8936FF266BBED1BDC208" blockId="2.[151,1437,1761,2036]" pageId="2" pageNumber="110">
|
||||
<emphasis id="58C4C705FFCC8936FF266BBED158C3B0" bold="true" box="[199,334,1761,1786]" inLineHeading="true" pageId="2" pageNumber="110" reason="1">Etymology.</emphasis>
|
||||
The species name results from the words
|
||||
<emphasis id="58C4C705FFCC8936FCA16BBDD392C3B3" box="[832,900,1762,1785]" italics="true" pageId="2" pageNumber="110">grotto</emphasis>
|
||||
(alternate term for referring to a cave; Italian “grotta” and French “grotte”) and
|
||||
<emphasis id="58C4C705FFCC8936FDF76A5AD246C254" box="[534,592,1797,1822]" italics="true" pageId="2" pageNumber="110">Edén</emphasis>
|
||||
. It is allusive to the name of the cavern where most of the specimens of the new species were found.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="22AA489CFFCC8937FF266A12D37EC78F" lastPageId="3" lastPageNumber="111" pageId="2" pageNumber="110" type="diagnosis">
|
||||
<paragraph id="6A0F1B17FFCC8937FF266A12D111C4E3" blockId="2.[151,1437,1761,2036]" lastBlockId="3.[151,1438,151,2024]" lastPageId="3" lastPageNumber="111" pageId="2" pageNumber="110">
|
||||
<emphasis id="58C4C705FFCC8936FF266A12D232C22C" bold="true" box="[199,548,1869,1894]" inLineHeading="true" pageId="2" pageNumber="110" reason="1">Diagnosis (male and female).</emphasis>
|
||||
The following combination of morphological features makes the new species unique among other species of
|
||||
<taxonomicName id="ADB06094FFCC8936FDED6A2ED2A6C2C2" authority="(Tityus)" box="[524,688,1905,1928]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="110" phylum="Arthropoda" rank="subGenus" subGenus="Tityus">
|
||||
<emphasis id="58C4C705FFCC8936FDED6A2ED2A6C2C2" box="[524,688,1905,1928]" italics="true" pageId="2" pageNumber="110">Tityus (Tityus)</emphasis>
|
||||
</taxonomicName>
|
||||
. Species of moderate size (~42.51 mm male; 50.87–54.21 mm female), relatively thin-bodied with a slender metasoma, and coloration predominantly yellow-to-chestnut (
|
||||
<figureCitation id="F28B0792FFCC8936FA856ACBD0D3C29A" captionStart="FIGURES 4–7" captionStartId="5.[151,264,1830,1853]" captionTargetBox="[178,1424,213,1778]" captionTargetId="figure-60@5.[151,1436,193,1809]" captionTargetPageId="5" captionText="FIGURES 4–7. Tityus (Tityus) grottoedensis sp. nov., habitus. 4–5. Subadult paratype male, dorsal and ventral aspects (ICN- As-905). 6–7. One paratype female, dorsal and ventral aspects (MACN-Ar 30879). Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4915131" httpUri="https://zenodo.org/record/4915131/files/figure.png" pageId="2" pageNumber="110">Figs. 4–7</figureCitation>
|
||||
). Dorsal surface with three yellowish longitudinal stripes crossing the tergites and pre-tergites (
|
||||
<figureCitation id="F28B0792FFCC8936FAFA6AE8D59BC29A" box="[1307,1421,1975,2000]" captionStart="FIGURES 4–7" captionStartId="5.[151,264,1830,1853]" captionTargetBox="[178,1424,213,1778]" captionTargetId="figure-60@5.[151,1436,193,1809]" captionTargetPageId="5" captionText="FIGURES 4–7. Tityus (Tityus) grottoedensis sp. nov., habitus. 4–5. Subadult paratype male, dorsal and ventral aspects (ICN- As-905). 6–7. One paratype female, dorsal and ventral aspects (MACN-Ar 30879). Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4915131" httpUri="https://zenodo.org/record/4915131/files/figure.png" pageId="2" pageNumber="110">Figs. 4, 6</figureCitation>
|
||||
); pedipalp hand width slightly greater than pedipalp patella width (male Hw/Pw = 1,03/1,00; female Hw/Pw = 1,06–1,11/1,00); pedipalp segments with carinae strong and granulose; movable finger of pedipalp chela with 14 oblique rows of granules (including the short apical row and invariable in the sample studied); basal piece of the middle lamella of the pectines dilated in male and female (
|
||||
<figureCitation id="F28B0792FFCD8937FCCE6D81D395C5BD" box="[815,899,222,247]" captionStart="FIGURES 14–16" captionStartId="7.[151,264,1801,1824]" captionTargetBox="[158,1428,456,1773]" captionTargetId="figure-223@7.[151,1436,445,1780]" captionTargetPageId="7" captionText="FIGURES 14–16. Tityus (Tityus) grottoedensis sp. nov. 14. Subadult paratype male (ICN-As-905), right lateral aspect of metasoma and telson. 15–16. Holotype female (ICN-As-977). 15. Left pectine. 16. Sternites IV–VII; most granulation details of the sternites not depicted, emphasis made on carinae. Scale bars = 3 mm (Figs. 14, 16), 1 mm (Fig. 15)." figureDoi="http://doi.org/10.5281/zenodo.4915135" httpUri="https://zenodo.org/record/4915135/files/figure.png" pageId="3" pageNumber="111">Fig. 15</figureCitation>
|
||||
); pectinal teeth count
|
||||
<quantity id="AD48B6F2FFCD8937FB9C6D81D4F7C5BD" box="[1149,1249,222,247]" metricMagnitude="-1" metricUnit="m" metricValue="4.953" metricValueMax="5.08" metricValueMin="4.826" pageId="3" pageNumber="111" unit="in" value="19.5" valueMax="20.0" valueMin="19.0">19–20 in</quantity>
|
||||
male and
|
||||
<quantity id="AD48B6F2FFCD8937FAB76D81D0BBC451" metricMagnitude="-1" metricUnit="m" metricValue="4.699" metricValueMax="4.826" metricValueMin="4.572" pageId="3" pageNumber="111" unit="in" value="18.5" valueMax="19.0" valueMin="18.0">18–19 in</quantity>
|
||||
female; sternites IV to VI with visible lateral longitudinal carinae beneath the book lung spiracles (
|
||||
<figureCitation id="F28B0792FFCD8937FAD96C5DD598C451" box="[1336,1422,258,283]" captionStart="FIGURES 14–16" captionStartId="7.[151,264,1801,1824]" captionTargetBox="[158,1428,456,1773]" captionTargetId="figure-223@7.[151,1436,445,1780]" captionTargetPageId="7" captionText="FIGURES 14–16. Tityus (Tityus) grottoedensis sp. nov. 14. Subadult paratype male (ICN-As-905), right lateral aspect of metasoma and telson. 15–16. Holotype female (ICN-As-977). 15. Left pectine. 16. Sternites IV–VII; most granulation details of the sternites not depicted, emphasis made on carinae. Scale bars = 3 mm (Figs. 14, 16), 1 mm (Fig. 15)." figureDoi="http://doi.org/10.5281/zenodo.4915135" httpUri="https://zenodo.org/record/4915135/files/figure.png" pageId="3" pageNumber="111">Fig. 16</figureCitation>
|
||||
); metasomal carinae strong, granulose; dorsosubmedian carinae of metasomal segments I–IV with granules subequal in size, without any spinoid or enlarged terminal granule (
|
||||
<figureCitation id="F28B0792FFCD8937FCC06C16D363C428" box="[801,885,329,354]" captionStart="FIGURES 14–16" captionStartId="7.[151,264,1801,1824]" captionTargetBox="[158,1428,456,1773]" captionTargetId="figure-223@7.[151,1436,445,1780]" captionTargetPageId="7" captionText="FIGURES 14–16. Tityus (Tityus) grottoedensis sp. nov. 14. Subadult paratype male (ICN-As-905), right lateral aspect of metasoma and telson. 15–16. Holotype female (ICN-As-977). 15. Left pectine. 16. Sternites IV–VII; most granulation details of the sternites not depicted, emphasis made on carinae. Scale bars = 3 mm (Figs. 14, 16), 1 mm (Fig. 15)." figureDoi="http://doi.org/10.5281/zenodo.4915135" httpUri="https://zenodo.org/record/4915135/files/figure.png" pageId="3" pageNumber="111">Fig. 14</figureCitation>
|
||||
); ventrosubmedian carinae complete and parallel in segments I through IV; metasomal segment V and telson not markedly darkened in comparison to the preceding segments.
|
||||
</paragraph>
|
||||
<paragraph id="6A0F1B17FFCD8937FF266CECD37EC78F" blockId="3.[151,1438,151,2024]" pageId="3" pageNumber="111">
|
||||
<emphasis id="58C4C705FFCD8937FF266CECD17BC486" bold="true" box="[199,365,435,460]" inLineHeading="true" pageId="3" pageNumber="111" reason="1">Comparisons.</emphasis>
|
||||
Among members of
|
||||
<taxonomicName id="ADB06094FFCD8937FDBE6CEBD2E9C481" authority="(Tityus)" box="[607,767,436,459]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="111" phylum="Arthropoda" rank="subGenus" subGenus="Tityus">
|
||||
<emphasis id="58C4C705FFCD8937FDBE6CEBD2E9C481" box="[607,767,436,459]" italics="true" pageId="3" pageNumber="111">Tityus (Tityus)</emphasis>
|
||||
</taxonomicName>
|
||||
, and based on characters from the external morphology,
|
||||
<taxonomicName id="ADB06094FFCD8937FA666CEBD13BC4BA" authority="Botero-Trujillo & Flórez, 2014" authorityName="Botero-Trujillo & Flórez" authorityYear="2014" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="111" phylum="Arthropoda" rank="species" species="grottoedensis" status="sp. nov.">
|
||||
<emphasis id="58C4C705FFCD8937FA666CEBD13BC4BA" italics="true" pageId="3" pageNumber="111">T. grottoedensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="58C4C705FFCD8937FED76C88D184C4BA" bold="true" box="[310,402,471,496]" pageId="3" pageNumber="111">
|
||||
<taxonomicNameLabel id="43F77A7EFFCD8937FED76C88D184C4BA" box="[310,402,471,496]" pageId="3" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
seems to mostly resemble
|
||||
<taxonomicName id="ADB06094FFCD8937FD2A6C87D403C4BA" authority="Lourenco, 1981" authorityName="Lourenco" authorityYear="1981" box="[715,1045,471,496]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="111" phylum="Arthropoda" rank="species" species="demangei">
|
||||
<emphasis id="58C4C705FFCD8937FD2A6C87D343C4BA" box="[715,853,471,496]" italics="true" pageId="3" pageNumber="111">T. demangei</emphasis>
|
||||
<bibRefCitation id="0E2166E6FFCD8937FCBF6C87D403C4BA" author="Lourenco, W. R." box="[862,1045,471,496]" pageId="3" pageNumber="111" pagination="635 - 662" refId="ref7131" refString="Lourenco, W. R. (1981) Scorpions cavernicoles de l'Equateur: Tityus demangei n. sp. et Ananteris ashmolei n. sp. (Buthidae); Troglotayosicus vachoni n. gen., n. sp. (Chactidae), scorpion troglobie. Bulletin du Museum national d'Histoire Naturelle Paris (Zoologie, Biologie et Ecologie Animale), 3, 635 - 662." type="journal article" year="1981">Lourenço, 1981</bibRefCitation>
|
||||
</taxonomicName>
|
||||
. This species, described from Los Tayos Cave in
|
||||
<collectingCountry id="12A75B87FFCD8937FEDF6CA5D18BC759" box="[318,413,506,531]" name="Ecuador" pageId="3" pageNumber="111">Ecuador</collectingCountry>
|
||||
, is similar in that it measures 54.2 mm (male), exhibits a general yellowish coloration with three light longitudinal stripes in the tergites, has metasomal and pedipalpal carinae well developed (granulose), and the movable finger of pedipalp chela bears 14–15 oblique rows of granules (
|
||||
<bibRefCitation id="0E2166E6FFCD8937FB646F1ED528C710" author="Lourenco, W. R." box="[1157,1342,577,602]" pageId="3" pageNumber="111" pagination="635 - 662" refId="ref7131" refString="Lourenco, W. R. (1981) Scorpions cavernicoles de l'Equateur: Tityus demangei n. sp. et Ananteris ashmolei n. sp. (Buthidae); Troglotayosicus vachoni n. gen., n. sp. (Chactidae), scorpion troglobie. Bulletin du Museum national d'Histoire Naturelle Paris (Zoologie, Biologie et Ecologie Animale), 3, 635 - 662." type="journal article" year="1981">Lourenço 1981</bibRefCitation>
|
||||
).
|
||||
<taxonomicName id="ADB06094FFCD8937FAB86F1DD13BC734" authority="Botero-Trujillo & Flórez, 2014" authorityName="Botero-Trujillo & Flórez" authorityYear="2014" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="111" phylum="Arthropoda" rank="species" species="grottoedensis" status="sp. nov.">
|
||||
<emphasis id="58C4C705FFCD8937FAB86F1DD13BC734" italics="true" pageId="3" pageNumber="111">Tityus grottoedensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="58C4C705FFCD8937FED46F3AD187C734" bold="true" box="[309,401,613,638]" pageId="3" pageNumber="111">
|
||||
<taxonomicNameLabel id="43F77A7EFFCD8937FED46F3AD187C734" box="[309,401,613,638]" pageId="3" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
can be separated from
|
||||
<taxonomicName id="ADB06094FFCD8937FD7F6F39D33FC734" authorityName="Lourenco" authorityYear="1981" box="[670,809,613,638]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="111" phylum="Arthropoda" rank="species" species="demangei">
|
||||
<emphasis id="58C4C705FFCD8937FD7F6F39D33FC734" box="[670,809,613,638]" italics="true" pageId="3" pageNumber="111">T. demangei</emphasis>
|
||||
</taxonomicName>
|
||||
, as the later has metasomal segments IV–V and telson noticeably darkened, the basal piece of the middle lamella of the pectines is not dilated in male, and the pectinal teeth count is slightly lower (16–18, mode 17) (
|
||||
<bibRefCitation id="0E2166E6FFCD8937FD4A6FF3D34DC78F" author="Lourenco, W. R." box="[683,859,684,709]" pageId="3" pageNumber="111" pagination="635 - 662" refId="ref7131" refString="Lourenco, W. R. (1981) Scorpions cavernicoles de l'Equateur: Tityus demangei n. sp. et Ananteris ashmolei n. sp. (Buthidae); Troglotayosicus vachoni n. gen., n. sp. (Chactidae), scorpion troglobie. Bulletin du Museum national d'Histoire Naturelle Paris (Zoologie, Biologie et Ecologie Animale), 3, 635 - 662." type="journal article" year="1981">Lourenço 1981</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="22AA489CFFCD8937FF266F90D44DC0C7" pageId="3" pageNumber="111" type="discussion">
|
||||
<paragraph id="6A0F1B17FFCD8937FF266F90D2D5C160" blockId="3.[151,1438,151,2024]" pageId="3" pageNumber="111">
|
||||
A recent collection of an adult male and a female of
|
||||
<taxonomicName id="ADB06094FFCD8937FCA26F8FD3C6C7A2" authorityName="Lourenco" authorityYear="1981" box="[835,976,719,744]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="111" phylum="Arthropoda" rank="species" species="demangei">
|
||||
<emphasis id="58C4C705FFCD8937FCA26F8FD3C6C7A2" box="[835,976,719,744]" italics="true" pageId="3" pageNumber="111">T. demangei</emphasis>
|
||||
</taxonomicName>
|
||||
from the area of the
|
||||
<typeStatus id="B50BA5B5FFCD8937FB356F8FD512C7A2" box="[1236,1284,720,744]" pageId="3" pageNumber="111">type</typeStatus>
|
||||
locality (see Methods) allowed confirming the above-referred differences. In addition, it was revealed that the median yellowish longitudinal stripe on the tergites of
|
||||
<taxonomicName id="ADB06094FFCD8937FDA36E48D2D8C665" authorityName="Lourenco" authorityYear="1981" box="[578,718,790,815]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="111" phylum="Arthropoda" rank="species" species="demangei">
|
||||
<emphasis id="58C4C705FFCD8937FDA36E48D2D8C665" box="[578,718,790,815]" italics="true" pageId="3" pageNumber="111">T. demangei</emphasis>
|
||||
</taxonomicName>
|
||||
is not continued in the pre-tergites (where the median part is brownish), as opposed to
|
||||
<taxonomicName id="ADB06094FFCD8937FE246E65D26CC618" authority="Botero-Trujillo & Flórez, 2014" authorityName="Botero-Trujillo & Flórez" authorityYear="2014" box="[453,634,825,850]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="111" phylum="Arthropoda" rank="species" species="grottoedensis" status="sp. nov.">
|
||||
<emphasis id="58C4C705FFCD8937FE246E65D26CC618" box="[453,634,825,850]" italics="true" pageId="3" pageNumber="111">T. grottoedensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="58C4C705FFCD8937FD646E66D2F5C618" bold="true" box="[645,739,825,850]" pageId="3" pageNumber="111">
|
||||
<taxonomicNameLabel id="43F77A7EFFCD8937FD646E66D2F5C618" box="[645,739,825,850]" pageId="3" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
in which it is continuous in tergites and pre-tergites. Other difference is that the ventral surface of metasomal segments I to V and the latero-distal region of the telson vesicle of
|
||||
<taxonomicName id="ADB06094FFCD8937FF586EDED153C6D3" authorityName="Lourenco" authorityYear="1981" box="[185,325,896,921]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="111" phylum="Arthropoda" rank="species" species="demangei">
|
||||
<emphasis id="58C4C705FFCD8937FF586EDED153C6D3" box="[185,325,896,921]" italics="true" pageId="3" pageNumber="111">T. demangei</emphasis>
|
||||
</taxonomicName>
|
||||
exhibit large conspicuous brown areas, absent in the new species. Finally, the newly collected specimens revealed that, even though
|
||||
<bibRefCitation id="0E2166E6FFCD8937FDAD6EFAD350C6F7" author="Lourenco, W. R." box="[588,838,932,957]" pageId="3" pageNumber="111" pagination="635 - 662" refId="ref7131" refString="Lourenco, W. R. (1981) Scorpions cavernicoles de l'Equateur: Tityus demangei n. sp. et Ananteris ashmolei n. sp. (Buthidae); Troglotayosicus vachoni n. gen., n. sp. (Chactidae), scorpion troglobie. Bulletin du Museum national d'Histoire Naturelle Paris (Zoologie, Biologie et Ecologie Animale), 3, 635 - 662." type="journal article" year="1981">Lourenço (1981: 642)</bibRefCitation>
|
||||
addressed for
|
||||
<taxonomicName id="ADB06094FFCD8937FC116EFAD46CC6F7" authorityName="Lourenco" authorityYear="1981" box="[1008,1146,932,957]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="111" phylum="Arthropoda" rank="species" species="demangei">
|
||||
<emphasis id="58C4C705FFCD8937FC116EFAD46CC6F7" box="[1008,1146,932,957]" italics="true" pageId="3" pageNumber="111">T. demangei</emphasis>
|
||||
</taxonomicName>
|
||||
that
|
||||
<emphasis id="58C4C705FFCD8937FB576EFAD27DC6A8" italics="true" pageId="3" pageNumber="111">
|
||||
“aucune trace de la carène intermédiaire sur le II
|
||||
<superScript id="9DC5B65FFFCD8937FE146E98D1EAC69F" attach="left" box="[501,508,967,981]" fontSize="6" pageId="3" pageNumber="111">e</superScript>
|
||||
anneau”
|
||||
</emphasis>
|
||||
[no trace of the intermediate (=median lateral) carina on the second metasomal segment], that species does indeed have one or two posterior granules as vestiges of the median lateral carinae, as it happens to occur in the new species.
|
||||
</paragraph>
|
||||
<paragraph id="6A0F1B17FFCD8937FF266969D44DC0C7" blockId="3.[151,1438,151,2024]" pageId="3" pageNumber="111">
|
||||
<taxonomicName id="ADB06094FFCD8937FF266969D1BCC104" authority="Botero-Trujillo & Flórez, 2014" authorityName="Botero-Trujillo & Flórez" authorityYear="2014" box="[199,426,1077,1102]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="111" phylum="Arthropoda" rank="species" species="grottoedensis" status="sp. nov.">
|
||||
<emphasis id="58C4C705FFCD8937FF266969D1BCC104" box="[199,426,1077,1102]" italics="true" pageId="3" pageNumber="111">Tityus grottoedensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="58C4C705FFCD8937FE55696AD207C104" bold="true" box="[436,529,1077,1102]" pageId="3" pageNumber="111">
|
||||
<taxonomicNameLabel id="43F77A7EFFCD8937FE55696AD207C104" box="[436,529,1077,1102]" pageId="3" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, can easily be differentiated from the Colombian con-subgeneric species with close geographical occurrence in the following respects:
|
||||
<taxonomicName id="ADB06094FFCD8937FCDD6906D471C13B" authority="Lourenco, 1994" authorityName="Lourenco" authorityYear="1994" box="[828,1127,1112,1137]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="111" phylum="Arthropoda" rank="species" species="blanci">
|
||||
<emphasis id="58C4C705FFCD8937FCDD6906D3B5C13B" box="[828,931,1112,1137]" italics="true" pageId="3" pageNumber="111">T. blanci</emphasis>
|
||||
Lourenço, 1994
|
||||
</taxonomicName>
|
||||
bears spinoid granules in dorsosubmedian carinae of metasomal segments II–IV;
|
||||
<taxonomicName id="ADB06094FFCD8937FCF06923D4B5C1DE" authority="Lourenco & Florez, 1990" authorityName="Lourenco & Florez" authorityYear="1990" box="[785,1187,1147,1172]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="111" phylum="Arthropoda" rank="species" species="sastrei">
|
||||
<emphasis id="58C4C705FFCD8937FCF06923D36EC1D9" box="[785,888,1148,1171]" italics="true" pageId="3" pageNumber="111">T. sastrei</emphasis>
|
||||
<bibRefCitation id="0E2166E6FFCD8937FC606923D4B5C1DE" author="Lourenco, W. R. & Florez, D. E." box="[897,1187,1147,1172]" pageId="3" pageNumber="111" pagination="119 - 134" refId="ref7691" refString="Lourenco, W. R. & Florez, D. E. (1990) Scorpions (Chelicerata) de Colombie. III. The scorpion fauna of the Pacific region (Choco), with some biogeographic considerations. Amazoniana, 11 (2), 119 - 134." type="journal article" year="1990">Lourenço & Flórez, 1990</bibRefCitation>
|
||||
</taxonomicName>
|
||||
possesses 16 rows of granules in the movable finger of pedipalp chela and metasomal carinae built-up by weak sparse granules;
|
||||
<taxonomicName id="ADB06094FFCD8937FA6669FFD1B7C191" authority="Lourenco, 1997" authorityName="Lourenco" authorityYear="1997" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="111" phylum="Arthropoda" rank="species" species="rebieri">
|
||||
<emphasis id="58C4C705FFCD8937FA6669FFD0F4C191" italics="true" pageId="3" pageNumber="111">T. rebieri</emphasis>
|
||||
<bibRefCitation id="0E2166E6FFCD8937FF0D699CD1B7C191" author="Lourenco, W. R." box="[236,417,1218,1243]" pageId="3" pageNumber="111" pagination="61 - 94" refId="ref7322" refString="Lourenco, W. R. (1997) Synopsis de la faune de scorpions de Colombie, avec des considerations sur la systematique et la biogeographie des especes. Revue Suisse de Zoologie, 104 (1), 61 - 94." type="journal article" year="1997">Lourenço, 1997</bibRefCitation>
|
||||
</taxonomicName>
|
||||
has ventrosubmedian carinae of metasomal segments III and IV converging proximally forming a ‘Y’;
|
||||
<taxonomicName id="ADB06094FFCD8937FEA869B8D278C1B5" authority="Lourenco, 2000" authorityName="Lourenco" authorityYear="2000" box="[329,622,1254,1279]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="111" phylum="Arthropoda" rank="species" species="gaffini">
|
||||
<emphasis id="58C4C705FFCD8937FEA869B8D1A6C1B5" box="[329,432,1254,1279]" italics="true" pageId="3" pageNumber="111">T. gaffini</emphasis>
|
||||
<bibRefCitation id="0E2166E6FFCD8937FE5B69B8D278C1B5" author="Lourenco, W. R." box="[442,622,1254,1279]" pageId="3" pageNumber="111" pagination="449 - 461" refId="ref7366" refString="Lourenco, W. R. (2000) Synopsis of the Colombian species of Tityus Koch (Chelicerata, Scorpiones, Buthidae), with descriptions of three new species. Journal of Natural History, 34, 449 - 461. http: // dx. doi. org / 10.1080 / 002229300299561" type="journal article" year="2000">Lourenço, 2000</bibRefCitation>
|
||||
</taxonomicName>
|
||||
exhibits a smaller number of pectinal teeth (15–17) and very subdued metasomal carination;
|
||||
<taxonomicName id="ADB06094FFCD8937FE7E6855D2D0C068" authority="Pocock, 1902" authorityName="Pocock" authorityYear="1902" box="[415,710,1289,1314]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="111" phylum="Arthropoda" rank="species" species="engelkei">
|
||||
<emphasis id="58C4C705FFCD8937FE7E6855D20BC068" box="[415,541,1289,1314]" italics="true" pageId="3" pageNumber="111">T. engelkei</emphasis>
|
||||
<bibRefCitation id="0E2166E6FFCD8937FDC86856D2D0C068" author="Pocock, R. I." box="[553,710,1289,1314]" pageId="3" pageNumber="111" pagination="364 - 380" refId="ref8112" refString="Pocock, R. I. (1902) A contribution to the systematics of scorpions. I. Some corrections in nomenclature. II. Notes on some species of Parabuthus contained in the British Museum. III. Descriptions of some new and old species. Annals and Magazine of Natural History, 7 (10), 364 - 380." type="journal article" year="1902">Pocock, 1902</bibRefCitation>
|
||||
</taxonomicName>
|
||||
is yellow in color without any reticulation pattern and has a smaller pectinal teeth count (15–16);
|
||||
<taxonomicName id="ADB06094FFCD8937FDD66871D346C00C" authority="Florez, 1996" authorityName="Florez" authorityYear="1996" box="[567,848,1325,1350]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="111" phylum="Arthropoda" rank="species" species="lourencoi">
|
||||
<emphasis id="58C4C705FFCD8937FDD66871D2AAC00C" box="[567,700,1325,1350]" italics="true" pageId="3" pageNumber="111">T. lourençoi</emphasis>
|
||||
<bibRefCitation id="0E2166E6FFCD8937FD226872D346C00C" author="Florez, E." box="[707,848,1325,1350]" pageId="3" pageNumber="111" pagination="683 - 685" refId="ref6668" refString="Florez, E. (1996) Tityus lourencoi, a new species of scorpion from the Cordillera Oriental, Colombia (Scorpiones, Buthidae). Revista de Biologia Tropical, 44 (2), 683 - 685." type="journal article" year="1996">Flórez, 1996</bibRefCitation>
|
||||
</taxonomicName>
|
||||
bears a posterior spinoid granule in dorsosubmedian carinae of metasomal segments and has weak carinae on the pedipalp chela (
|
||||
<bibRefCitation id="0E2166E6FFCD8937FBE3680FD48BC023" author="Florez, E." box="[1026,1181,1360,1385]" pageId="3" pageNumber="111" refId="ref6742" refString="Florez, E. (2001 b) Sinopsis de los escorpiones de la familia Buthidae en Colombia. Magister Thesis. Universidad Nacional de Colombia, Bogota, Colombia, 102 pp." type="book" year="2001">Flórez 2001b</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="0E2166E6FFCD8937FB4B680FD581C023" author="Botero-Trujillo, R." box="[1194,1431,1360,1385]" pageId="3" pageNumber="111" pagination="29 - 34" refId="ref6141" refString="Botero-Trujillo, R. (2008) Redescription of Tityus (Tityus) gaffini Lourenco, 2000 (Scorpiones: Buthidae) and new locality records. Revista Iberica de Aracnologia, 16, 29 - 34." type="journal article" year="2008">Botero-Trujillo 2008</bibRefCitation>
|
||||
; and personally examined material from the arachnological collection of the ICN-UN).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="22AA489CFFCD893CFF2668C8D13CC047" lastPageId="8" lastPageNumber="116" pageId="3" pageNumber="111" type="description">
|
||||
<paragraph id="6A0F1B17FFCD8937FF2668C8D3BDC09E" blockId="3.[151,1438,151,2024]" pageId="3" pageNumber="111">
|
||||
<emphasis id="58C4C705FFCD8937FF2668C8D14EC0FA" bold="true" box="[199,344,1431,1456]" inLineHeading="true" pageId="3" pageNumber="111" reason="1">Description.</emphasis>
|
||||
(based on adult females and subadult male only; juvenile male not considered in the description unless otherwise specified; newborns not included in this description).
|
||||
</paragraph>
|
||||
<paragraph id="6A0F1B17FFCD8937FF266881D246C2A2" blockId="3.[151,1438,151,2024]" pageId="3" pageNumber="111">
|
||||
<emphasis id="58C4C705FFCD8937FF266881D11BC0BD" bold="true" box="[199,269,1502,1527]" inLineHeading="true" italics="true" pageId="3" pageNumber="111" reason="1">Color.</emphasis>
|
||||
General color yellowish (male) to light chestnut (
|
||||
<figureCitation id="F28B0792FFCD8937FCDE6881D3BBC0BD" box="[831,941,1502,1527]" captionStart="FIGURES 4–7" captionStartId="5.[151,264,1830,1853]" captionTargetBox="[178,1424,213,1778]" captionTargetId="figure-60@5.[151,1436,193,1809]" captionTargetPageId="5" captionText="FIGURES 4–7. Tityus (Tityus) grottoedensis sp. nov., habitus. 4–5. Subadult paratype male, dorsal and ventral aspects (ICN- As-905). 6–7. One paratype female, dorsal and ventral aspects (MACN-Ar 30879). Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4915131" httpUri="https://zenodo.org/record/4915131/files/figure.png" pageId="3" pageNumber="111">Figs. 4–7</figureCitation>
|
||||
). Differences in color between the male and the females are presumably due to the male being a subadult. Carapace with variegated brownish pigmentation; median ocular tubercle brownish, not noticeably darkened; lateral ocular tubercles black. Chelicerae with coxa yellow; hand with complete reticular pattern on dorsal surface; movable and fixed fingers entirely brown. Tergites predominantly brown, with three conspicuous yellowish longitudinal stripes crossing tergites and pre-tergites I through VI in the position of medial and dorsolateral carinae; tergite VII predominantly yellow (male) or chestnut (female), without any special coloration design. Coxosternal region, genital operculum, pectinal basal piece and sternites III to V uniformly yellow (male) or chestnut (female) and without pigmentation; pectines whitish yellow; sternite VI with minute brownish area in posterior margin; sternite VII with conspicuous brownish areas medially. Metasomal segments base color same as the rest of the body, with segment V and telson only slightly darker than other segments (female); dorsal and lateral surfaces of segments I–IV and all surfaces of segment V and telson uniformly colored and spotless; segments I–IV ventral surface with brownish mottling (less evident in IV), almost undistinguishable in female. Pedipalps uniformly yellow (male) or chestnut (female) throughout, except for movable and fixed fingers which are blackish with yellowish tips. Legs with or without inconspicuous brownish mottling in patella, tibia and basitarsus.
|
||||
</paragraph>
|
||||
<caption id="3ECF4B9FFFCA8930FF726C99D281C75E" ID-Table-UUID="3ECF4B9FFFCA8930FF726C99D281C75E" httpUri="http://table.plazi.org/id/3ECF4B9FFFCA8930FF726C99D281C75E" pageId="4" pageNumber="112" startId="4.[147,230,454,475]" targetBox="[160,1691,564,1710]" targetIsTable="true" targetPageId="4">
|
||||
<paragraph id="6A0F1B17FFCA8930FF726C99D281C75E" blockId="4.[147,1734,448,532]" pageId="4" pageNumber="112">
|
||||
<emphasis id="58C4C705FFCA8930FF726C99D116C491" bold="true" box="[147,256,454,475]" pageId="4" pageNumber="112">TABLE 1.</emphasis>
|
||||
Meristic data for
|
||||
<taxonomicName id="ADB06094FFCA8930FE516C99D2DEC491" authority="Botero-Trujillo & Flórez, 2014" authorityName="Botero-Trujillo & Flórez" authorityYear="2014" box="[432,712,454,475]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="112" phylum="Arthropoda" rank="species" species="grottoedensis" status="sp. nov." subGenus="Tityus">
|
||||
<emphasis id="58C4C705FFCA8930FE516C99D2DEC491" box="[432,712,454,475]" italics="true" pageId="4" pageNumber="112">Tityus (Tityus) grottoedensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="58C4C705FFCA8930FD2F6C99D308C491" bold="true" box="[718,798,454,475]" pageId="4" pageNumber="112">
|
||||
<taxonomicNameLabel id="43F77A7EFFCA8930FD2F6C99D308C491" box="[718,798,454,475]" pageId="4" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
Measurements in millimeters.
|
||||
<superScript id="9DC5B65FFFCA8930FBAE6C9FD441C484" attach="right" box="[1103,1111,448,462]" fontSize="6" pageId="4" pageNumber="112">1</superScript>
|
||||
Sum of carapace, mesosoma and metasoma.
|
||||
<superScript id="9DC5B65FFFCA8930F9EF6C9FD600C484" attach="right" box="[1550,1558,448,462]" fontSize="6" pageId="4" pageNumber="112">2</superScript>
|
||||
Sum of tergites I– VII.
|
||||
<superScript id="9DC5B65FFFCA8930FF236C83D0DCC4A0" attach="right" box="[194,202,476,490]" fontSize="6" pageId="4" pageNumber="112">3</superScript>
|
||||
Sum of metasomal segments I–V and telson.
|
||||
<superScript id="9DC5B65FFFCA8930FD6C6C83D282C4A0" attach="right" box="[653,660,476,490]" fontSize="6" pageId="4" pageNumber="112">4</superScript>
|
||||
Sum of femur, patella and chela.
|
||||
<superScript id="9DC5B65FFFCA8930FC3E6C83D3F0C4A0" attach="right" box="[991,998,476,490]" fontSize="6" pageId="4" pageNumber="112">5</superScript>
|
||||
Measured from the commissure of the junction with the movable finger to the finger tip.
|
||||
<superScript id="9DC5B65FFFCA8930FEFC6CA6D133C74D" attach="right" box="[285,293,505,519]" fontSize="6" pageId="4" pageNumber="112">6</superScript>
|
||||
Measured along the marginal lamella.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="6A0F1B17FFCA8930FBCC6F6BD67EC3E4" pageId="4" pageNumber="112">
|
||||
<table id="18B0E9B7FFCA76CBFF416F6BD68DC3E4" box="[160,1691,564,1710]" gridcols="7" gridrows="41" pageId="4" pageNumber="112">
|
||||
<tr id="D4801955FFCA76CBFF416F6BD68DC703" box="[160,1691,564,585]" gridrow="0" pageId="4" pageNumber="112" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-5="1">
|
||||
<th id="97517029FFCA76CBFC046F6BD530C703" box="[997,1318,564,585]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="4" pageNumber="112">Adult females</th>
|
||||
<th id="97517029FFCA76CBF9EE6F6BD68DC703" box="[1551,1691,564,585]" gridcol="6" gridrow="0" pageId="4" pageNumber="112">Subadult male</th>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416F0ED68DC72C" box="[160,1691,593,614]" gridrow="1" pageId="4" pageNumber="112" rowspan-0="1" rowspan-1="1">
|
||||
<td id="97517029FFCA76CBFCFA6F0ED3B0C72C" box="[795,934,593,614]" gridcol="2" gridrow="1" pageId="4" pageNumber="112">(ICN-As-977)</td>
|
||||
<td id="97517029FFCA76CBFC046F0ED479C72C" box="[997,1135,593,614]" gridcol="3" gridrow="1" pageId="4" pageNumber="112">(ICN-As-872)</td>
|
||||
<td id="97517029FFCA76CBFB536F0ED530C72C" box="[1202,1318,593,614]" gridcol="4" gridrow="1" pageId="4" pageNumber="112">(MACN-Ar</td>
|
||||
<td id="97517029FFCA76CBFAB46F0ED5C3C72C" box="[1365,1493,593,614]" gridcol="5" gridrow="1" pageId="4" pageNumber="112">Mean female</td>
|
||||
<td id="97517029FFCA76CBF9EE6F0ED68DC72C" box="[1551,1691,593,614]" gridcol="6" gridrow="1" pageId="4" pageNumber="112">(ICN-As-905)</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416F32D68DC7C8" box="[160,1691,621,642]" gridrow="2" pageId="4" pageNumber="112" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-6="1">
|
||||
<td id="97517029FFCA76CBFB536F32D530C7C8" box="[1202,1318,621,642]" gridcol="4" gridrow="2" pageId="4" pageNumber="112">30879)</td>
|
||||
<td id="97517029FFCA76CBFAB46F32D5C3C7C8" box="[1365,1493,621,642]" gridcol="5" gridrow="2" pageId="4" pageNumber="112">(n=3)</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416FDBD68DC7EA" box="[160,1691,644,672]" gridrow="3" pageId="4" pageNumber="112" rowspan-1="1">
|
||||
<th id="97517029FFCA76CBFF416FDBD19AC7EA" box="[160,396,644,672]" gridcol="0" gridrow="3" pageId="4" pageNumber="112">Total body L: 1</th>
|
||||
<td id="97517029FFCA76CBFCFA6FDBD3B0C7EA" box="[795,934,644,672]" gridcol="2" gridrow="3" pageId="4" pageNumber="112">54.21</td>
|
||||
<td id="97517029FFCA76CBFC046FDBD479C7EA" box="[997,1135,644,672]" gridcol="3" gridrow="3" pageId="4" pageNumber="112">51.15</td>
|
||||
<td id="97517029FFCA76CBFB536FDBD530C7EA" box="[1202,1318,644,672]" gridcol="4" gridrow="3" pageId="4" pageNumber="112">50.87</td>
|
||||
<td id="97517029FFCA76CBFAB46FDBD5C3C7EA" box="[1365,1493,644,672]" gridcol="5" gridrow="3" pageId="4" pageNumber="112">52.08</td>
|
||||
<td id="97517029FFCA76CBF9EE6FDBD68DC7EA" box="[1551,1691,644,672]" gridcol="6" gridrow="3" pageId="4" pageNumber="112">42.51</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416FF9D68DC7F1" box="[160,1691,678,699]" gridrow="4" pageId="4" pageNumber="112">
|
||||
<th id="97517029FFCA76CBFF416FF9D19AC7F1" box="[160,396,678,699]" gridcol="0" gridrow="4" pageId="4" pageNumber="112">Carapace:</th>
|
||||
<td id="97517029FFCA76CBFDF56FF9D2D1C7F1" box="[532,711,678,699]" gridcol="1" gridrow="4" pageId="4" pageNumber="112">L:</td>
|
||||
<td id="97517029FFCA76CBFCFA6FF9D3B0C7F1" box="[795,934,678,699]" gridcol="2" gridrow="4" pageId="4" pageNumber="112">6.04</td>
|
||||
<td id="97517029FFCA76CBFC046FF9D479C7F1" box="[997,1135,678,699]" gridcol="3" gridrow="4" pageId="4" pageNumber="112">5.90</td>
|
||||
<td id="97517029FFCA76CBFB536FF9D530C7F1" box="[1202,1318,678,699]" gridcol="4" gridrow="4" pageId="4" pageNumber="112">5.88</td>
|
||||
<td id="97517029FFCA76CBFAB46FF9D5C3C7F1" box="[1365,1493,678,699]" gridcol="5" gridrow="4" pageId="4" pageNumber="112">5.94</td>
|
||||
<td id="97517029FFCA76CBF9EE6FF9D68DC7F1" box="[1551,1691,678,699]" gridcol="6" gridrow="4" pageId="4" pageNumber="112">4.91</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416F9CD68DC792" box="[160,1691,707,728]" gridrow="5" pageId="4" pageNumber="112" rowspan-0="1">
|
||||
<td id="97517029FFCA76CBFDF56F9CD2D1C792" box="[532,711,707,728]" gridcol="1" gridrow="5" pageId="4" pageNumber="112">Anterior W:</td>
|
||||
<td id="97517029FFCA76CBFCFA6F9CD3B0C792" box="[795,934,707,728]" gridcol="2" gridrow="5" pageId="4" pageNumber="112">4.16</td>
|
||||
<td id="97517029FFCA76CBFC046F9CD479C792" box="[997,1135,707,728]" gridcol="3" gridrow="5" pageId="4" pageNumber="112">4.10</td>
|
||||
<td id="97517029FFCA76CBFB536F9CD530C792" box="[1202,1318,707,728]" gridcol="4" gridrow="5" pageId="4" pageNumber="112">3.97</td>
|
||||
<td id="97517029FFCA76CBFAB46F9CD5C3C792" box="[1365,1493,707,728]" gridcol="5" gridrow="5" pageId="4" pageNumber="112">4.08</td>
|
||||
<td id="97517029FFCA76CBF9EE6F9CD68DC792" box="[1551,1691,707,728]" gridcol="6" gridrow="5" pageId="4" pageNumber="112">3.32</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416F80D68DC7BE" box="[160,1691,735,756]" gridrow="6" pageId="4" pageNumber="112" rowspan-0="1">
|
||||
<td id="97517029FFCA76CBFDF56F80D2D1C7BE" box="[532,711,735,756]" gridcol="1" gridrow="6" pageId="4" pageNumber="112">Posterior W:</td>
|
||||
<td id="97517029FFCA76CBFCFA6F80D3B0C7BE" box="[795,934,735,756]" gridcol="2" gridrow="6" pageId="4" pageNumber="112">6.04</td>
|
||||
<td id="97517029FFCA76CBFC046F80D479C7BE" box="[997,1135,735,756]" gridcol="3" gridrow="6" pageId="4" pageNumber="112">5.85</td>
|
||||
<td id="97517029FFCA76CBFB536F80D530C7BE" box="[1202,1318,735,756]" gridcol="4" gridrow="6" pageId="4" pageNumber="112">5.60</td>
|
||||
<td id="97517029FFCA76CBFAB46F80D5C3C7BE" box="[1365,1493,735,756]" gridcol="5" gridrow="6" pageId="4" pageNumber="112">5.83</td>
|
||||
<td id="97517029FFCA76CBF9EE6F80D68DC7BE" box="[1551,1691,735,756]" gridcol="6" gridrow="6" pageId="4" pageNumber="112">4.70</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416FAAD68DC65A" box="[160,1691,757,784]" gridrow="7" pageId="4" pageNumber="112">
|
||||
<th id="97517029FFCA76CBFF416FAAD19AC65A" box="[160,396,757,784]" gridcol="0" gridrow="7" pageId="4" pageNumber="112">Mesosoma:</th>
|
||||
<td id="97517029FFCA76CBFDF56FAAD2D1C65A" box="[532,711,757,784]" gridcol="1" gridrow="7" pageId="4" pageNumber="112">Total L:2</td>
|
||||
<td id="97517029FFCA76CBFCFA6FAAD3B0C65A" box="[795,934,757,784]" gridcol="2" gridrow="7" pageId="4" pageNumber="112">15.20</td>
|
||||
<td id="97517029FFCA76CBFC046FAAD479C65A" box="[997,1135,757,784]" gridcol="3" gridrow="7" pageId="4" pageNumber="112">13.21</td>
|
||||
<td id="97517029FFCA76CBFB536FAAD530C65A" box="[1202,1318,757,784]" gridcol="4" gridrow="7" pageId="4" pageNumber="112">13.51</td>
|
||||
<td id="97517029FFCA76CBFAB46FAAD5C3C65A" box="[1365,1493,757,784]" gridcol="5" gridrow="7" pageId="4" pageNumber="112">13.97</td>
|
||||
<td id="97517029FFCA76CBF9EE6FAAD68DC65A" box="[1551,1691,757,784]" gridcol="6" gridrow="7" pageId="4" pageNumber="112">11.28</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416E4ED68DC666" box="[160,1691,785,812]" gridrow="8" pageId="4" pageNumber="112">
|
||||
<th id="97517029FFCA76CBFF416E4ED19AC666" box="[160,396,785,812]" gridcol="0" gridrow="8" pageId="4" pageNumber="112">Metasoma:</th>
|
||||
<td id="97517029FFCA76CBFDF56E4ED2D1C666" box="[532,711,785,812]" gridcol="1" gridrow="8" pageId="4" pageNumber="112">Total L:3</td>
|
||||
<td id="97517029FFCA76CBFCFA6E4ED3B0C666" box="[795,934,785,812]" gridcol="2" gridrow="8" pageId="4" pageNumber="112">32.97</td>
|
||||
<td id="97517029FFCA76CBFC046E4ED479C666" box="[997,1135,785,812]" gridcol="3" gridrow="8" pageId="4" pageNumber="112">32.04</td>
|
||||
<td id="97517029FFCA76CBFB536E4ED530C666" box="[1202,1318,785,812]" gridcol="4" gridrow="8" pageId="4" pageNumber="112">31.48</td>
|
||||
<td id="97517029FFCA76CBFAB46E4ED5C3C666" box="[1365,1493,785,812]" gridcol="5" gridrow="8" pageId="4" pageNumber="112">32.16</td>
|
||||
<td id="97517029FFCA76CBF9EE6E4ED68DC666" box="[1551,1691,785,812]" gridcol="6" gridrow="8" pageId="4" pageNumber="112">26.32</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416E6CD68DC602" box="[160,1691,819,840]" gridrow="9" pageId="4" pageNumber="112">
|
||||
<th id="97517029FFCA76CBFF416E6CD19AC602" box="[160,396,819,840]" gridcol="0" gridrow="9" pageId="4" pageNumber="112">Metasomal segment I:</th>
|
||||
<td id="97517029FFCA76CBFDF56E6CD2D1C602" box="[532,711,819,840]" gridcol="1" gridrow="9" pageId="4" pageNumber="112">L:</td>
|
||||
<td id="97517029FFCA76CBFCFA6E6CD3B0C602" box="[795,934,819,840]" gridcol="2" gridrow="9" pageId="4" pageNumber="112">3.98</td>
|
||||
<td id="97517029FFCA76CBFC046E6CD479C602" box="[997,1135,819,840]" gridcol="3" gridrow="9" pageId="4" pageNumber="112">3.91</td>
|
||||
<td id="97517029FFCA76CBFB536E6CD530C602" box="[1202,1318,819,840]" gridcol="4" gridrow="9" pageId="4" pageNumber="112">3.78</td>
|
||||
<td id="97517029FFCA76CBFAB46E6CD5C3C602" box="[1365,1493,819,840]" gridcol="5" gridrow="9" pageId="4" pageNumber="112">3.89</td>
|
||||
<td id="97517029FFCA76CBF9EE6E6CD68DC602" box="[1551,1691,819,840]" gridcol="6" gridrow="9" pageId="4" pageNumber="112">3.10</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416E10D68DC62E" box="[160,1691,847,868]" gridrow="10" pageId="4" pageNumber="112" rowspan-0="1">
|
||||
<td id="97517029FFCA76CBFDF56E10D2D1C62E" box="[532,711,847,868]" gridcol="1" gridrow="10" pageId="4" pageNumber="112">W:</td>
|
||||
<td id="97517029FFCA76CBFCFA6E10D3B0C62E" box="[795,934,847,868]" gridcol="2" gridrow="10" pageId="4" pageNumber="112">2.80</td>
|
||||
<td id="97517029FFCA76CBFC046E10D479C62E" box="[997,1135,847,868]" gridcol="3" gridrow="10" pageId="4" pageNumber="112">2.76</td>
|
||||
<td id="97517029FFCA76CBFB536E10D530C62E" box="[1202,1318,847,868]" gridcol="4" gridrow="10" pageId="4" pageNumber="112">2.62</td>
|
||||
<td id="97517029FFCA76CBFAB46E10D5C3C62E" box="[1365,1493,847,868]" gridcol="5" gridrow="10" pageId="4" pageNumber="112">2.73</td>
|
||||
<td id="97517029FFCA76CBF9EE6E10D68DC62E" box="[1551,1691,847,868]" gridcol="6" gridrow="10" pageId="4" pageNumber="112">2.41</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416E34D68DC6CA" box="[160,1691,875,896]" gridrow="11" pageId="4" pageNumber="112" rowspan-0="1">
|
||||
<td id="97517029FFCA76CBFDF56E34D2D1C6CA" box="[532,711,875,896]" gridcol="1" gridrow="11" pageId="4" pageNumber="112">H:</td>
|
||||
<td id="97517029FFCA76CBFCFA6E34D3B0C6CA" box="[795,934,875,896]" gridcol="2" gridrow="11" pageId="4" pageNumber="112">2.67</td>
|
||||
<td id="97517029FFCA76CBFC046E34D479C6CA" box="[997,1135,875,896]" gridcol="3" gridrow="11" pageId="4" pageNumber="112">2.62</td>
|
||||
<td id="97517029FFCA76CBFB536E34D530C6CA" box="[1202,1318,875,896]" gridcol="4" gridrow="11" pageId="4" pageNumber="112">2.46</td>
|
||||
<td id="97517029FFCA76CBFAB46E34D5C3C6CA" box="[1365,1493,875,896]" gridcol="5" gridrow="11" pageId="4" pageNumber="112">2.58</td>
|
||||
<td id="97517029FFCA76CBF9EE6E34D68DC6CA" box="[1551,1691,875,896]" gridcol="6" gridrow="11" pageId="4" pageNumber="112">2.30</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416ED8D68DC6D6" box="[160,1691,903,924]" gridrow="12" pageId="4" pageNumber="112">
|
||||
<th id="97517029FFCA76CBFF416ED8D19AC6D6" box="[160,396,903,924]" gridcol="0" gridrow="12" pageId="4" pageNumber="112">Metasomal segment II:</th>
|
||||
<td id="97517029FFCA76CBFDF56ED8D2D1C6D6" box="[532,711,903,924]" gridcol="1" gridrow="12" pageId="4" pageNumber="112">L:</td>
|
||||
<td id="97517029FFCA76CBFCFA6ED8D3B0C6D6" box="[795,934,903,924]" gridcol="2" gridrow="12" pageId="4" pageNumber="112">4.91</td>
|
||||
<td id="97517029FFCA76CBFC046ED8D479C6D6" box="[997,1135,903,924]" gridcol="3" gridrow="12" pageId="4" pageNumber="112">4.80</td>
|
||||
<td id="97517029FFCA76CBFB536ED8D530C6D6" box="[1202,1318,903,924]" gridcol="4" gridrow="12" pageId="4" pageNumber="112">4.69</td>
|
||||
<td id="97517029FFCA76CBFAB46ED8D5C3C6D6" box="[1365,1493,903,924]" gridcol="5" gridrow="12" pageId="4" pageNumber="112">4.80</td>
|
||||
<td id="97517029FFCA76CBF9EE6ED8D68DC6D6" box="[1551,1691,903,924]" gridcol="6" gridrow="12" pageId="4" pageNumber="112">3.90</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416EFCD68DC6F2" box="[160,1691,931,952]" gridrow="13" pageId="4" pageNumber="112" rowspan-0="1">
|
||||
<td id="97517029FFCA76CBFDF56EFCD2D1C6F2" box="[532,711,931,952]" gridcol="1" gridrow="13" pageId="4" pageNumber="112">W:</td>
|
||||
<td id="97517029FFCA76CBFCFA6EFCD3B0C6F2" box="[795,934,931,952]" gridcol="2" gridrow="13" pageId="4" pageNumber="112">2.69</td>
|
||||
<td id="97517029FFCA76CBFC046EFCD479C6F2" box="[997,1135,931,952]" gridcol="3" gridrow="13" pageId="4" pageNumber="112">2.69</td>
|
||||
<td id="97517029FFCA76CBFB536EFCD530C6F2" box="[1202,1318,931,952]" gridcol="4" gridrow="13" pageId="4" pageNumber="112">2.59</td>
|
||||
<td id="97517029FFCA76CBFAB46EFCD5C3C6F2" box="[1365,1493,931,952]" gridcol="5" gridrow="13" pageId="4" pageNumber="112">2.66</td>
|
||||
<td id="97517029FFCA76CBF9EE6EFCD68DC6F2" box="[1551,1691,931,952]" gridcol="6" gridrow="13" pageId="4" pageNumber="112">2.19</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416EE0D68DC69E" box="[160,1691,959,980]" gridrow="14" pageId="4" pageNumber="112" rowspan-0="1">
|
||||
<td id="97517029FFCA76CBFDF56EE0D2D1C69E" box="[532,711,959,980]" gridcol="1" gridrow="14" pageId="4" pageNumber="112">H:</td>
|
||||
<td id="97517029FFCA76CBFCFA6EE0D3B0C69E" box="[795,934,959,980]" gridcol="2" gridrow="14" pageId="4" pageNumber="112">2.47</td>
|
||||
<td id="97517029FFCA76CBFC046EE0D479C69E" box="[997,1135,959,980]" gridcol="3" gridrow="14" pageId="4" pageNumber="112">2.52</td>
|
||||
<td id="97517029FFCA76CBFB536EE0D530C69E" box="[1202,1318,959,980]" gridcol="4" gridrow="14" pageId="4" pageNumber="112">2.45</td>
|
||||
<td id="97517029FFCA76CBFAB46EE0D5C3C69E" box="[1365,1493,959,980]" gridcol="5" gridrow="14" pageId="4" pageNumber="112">2.48</td>
|
||||
<td id="97517029FFCA76CBF9EE6EE0D68DC69E" box="[1551,1691,959,980]" gridcol="6" gridrow="14" pageId="4" pageNumber="112">2.20</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416E84D68DC6BA" box="[160,1691,987,1008]" gridrow="15" pageId="4" pageNumber="112">
|
||||
<th id="97517029FFCA76CBFF416E84D19AC6BA" box="[160,396,987,1008]" gridcol="0" gridrow="15" pageId="4" pageNumber="112">Metasomal segment III:</th>
|
||||
<td id="97517029FFCA76CBFDF56E84D2D1C6BA" box="[532,711,987,1008]" gridcol="1" gridrow="15" pageId="4" pageNumber="112">L:</td>
|
||||
<td id="97517029FFCA76CBFCFA6E84D3B0C6BA" box="[795,934,987,1008]" gridcol="2" gridrow="15" pageId="4" pageNumber="112">5.36</td>
|
||||
<td id="97517029FFCA76CBFC046E84D479C6BA" box="[997,1135,987,1008]" gridcol="3" gridrow="15" pageId="4" pageNumber="112">5.17</td>
|
||||
<td id="97517029FFCA76CBFB536E84D530C6BA" box="[1202,1318,987,1008]" gridcol="4" gridrow="15" pageId="4" pageNumber="112">5.11</td>
|
||||
<td id="97517029FFCA76CBFAB46E84D5C3C6BA" box="[1365,1493,987,1008]" gridcol="5" gridrow="15" pageId="4" pageNumber="112">5.21</td>
|
||||
<td id="97517029FFCA76CBF9EE6E84D68DC6BA" box="[1551,1691,987,1008]" gridcol="6" gridrow="15" pageId="4" pageNumber="112">4.21</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416EA8D68DC146" box="[160,1691,1015,1036]" gridrow="16" pageId="4" pageNumber="112" rowspan-0="1">
|
||||
<td id="97517029FFCA76CBFDF56EA8D2D1C146" box="[532,711,1015,1036]" gridcol="1" gridrow="16" pageId="4" pageNumber="112">W:</td>
|
||||
<td id="97517029FFCA76CBFCFA6EA8D3B0C146" box="[795,934,1015,1036]" gridcol="2" gridrow="16" pageId="4" pageNumber="112">2.69</td>
|
||||
<td id="97517029FFCA76CBFC046EA8D479C146" box="[997,1135,1015,1036]" gridcol="3" gridrow="16" pageId="4" pageNumber="112">2.65</td>
|
||||
<td id="97517029FFCA76CBFB536EA8D530C146" box="[1202,1318,1015,1036]" gridcol="4" gridrow="16" pageId="4" pageNumber="112">2.67</td>
|
||||
<td id="97517029FFCA76CBFAB46EA8D5C3C146" box="[1365,1493,1015,1036]" gridcol="5" gridrow="16" pageId="4" pageNumber="112">2.67</td>
|
||||
<td id="97517029FFCA76CBF9EE6EA8D68DC146" box="[1551,1691,1015,1036]" gridcol="6" gridrow="16" pageId="4" pageNumber="112">2.25</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF41694CD68DC162" box="[160,1691,1043,1064]" gridrow="17" pageId="4" pageNumber="112" rowspan-0="1">
|
||||
<td id="97517029FFCA76CBFDF5694CD2D1C162" box="[532,711,1043,1064]" gridcol="1" gridrow="17" pageId="4" pageNumber="112">H:</td>
|
||||
<td id="97517029FFCA76CBFCFA694CD3B0C162" box="[795,934,1043,1064]" gridcol="2" gridrow="17" pageId="4" pageNumber="112">2.46</td>
|
||||
<td id="97517029FFCA76CBFC04694CD479C162" box="[997,1135,1043,1064]" gridcol="3" gridrow="17" pageId="4" pageNumber="112">2.57</td>
|
||||
<td id="97517029FFCA76CBFB53694CD530C162" box="[1202,1318,1043,1064]" gridcol="4" gridrow="17" pageId="4" pageNumber="112">2.47</td>
|
||||
<td id="97517029FFCA76CBFAB4694CD5C3C162" box="[1365,1493,1043,1064]" gridcol="5" gridrow="17" pageId="4" pageNumber="112">2.50</td>
|
||||
<td id="97517029FFCA76CBF9EE694CD68DC162" box="[1551,1691,1043,1064]" gridcol="6" gridrow="17" pageId="4" pageNumber="112">2.30</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416970D68DC10E" box="[160,1691,1071,1092]" gridrow="18" pageId="4" pageNumber="112">
|
||||
<th id="97517029FFCA76CBFF416970D19AC10E" box="[160,396,1071,1092]" gridcol="0" gridrow="18" pageId="4" pageNumber="112">Metasomal segment IV:</th>
|
||||
<td id="97517029FFCA76CBFDF56970D2D1C10E" box="[532,711,1071,1092]" gridcol="1" gridrow="18" pageId="4" pageNumber="112">L:</td>
|
||||
<td id="97517029FFCA76CBFCFA6970D3B0C10E" box="[795,934,1071,1092]" gridcol="2" gridrow="18" pageId="4" pageNumber="112">5.91</td>
|
||||
<td id="97517029FFCA76CBFC046970D479C10E" box="[997,1135,1071,1092]" gridcol="3" gridrow="18" pageId="4" pageNumber="112">5.68</td>
|
||||
<td id="97517029FFCA76CBFB536970D530C10E" box="[1202,1318,1071,1092]" gridcol="4" gridrow="18" pageId="4" pageNumber="112">5.64</td>
|
||||
<td id="97517029FFCA76CBFAB46970D5C3C10E" box="[1365,1493,1071,1092]" gridcol="5" gridrow="18" pageId="4" pageNumber="112">5.74</td>
|
||||
<td id="97517029FFCA76CBF9EE6970D68DC10E" box="[1551,1691,1071,1092]" gridcol="6" gridrow="18" pageId="4" pageNumber="112">4.70</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416914D68DC12A" box="[160,1691,1099,1120]" gridrow="19" pageId="4" pageNumber="112" rowspan-0="1">
|
||||
<td id="97517029FFCA76CBFDF56914D2D1C12A" box="[532,711,1099,1120]" gridcol="1" gridrow="19" pageId="4" pageNumber="112">W:</td>
|
||||
<td id="97517029FFCA76CBFCFA6914D3B0C12A" box="[795,934,1099,1120]" gridcol="2" gridrow="19" pageId="4" pageNumber="112">2.77</td>
|
||||
<td id="97517029FFCA76CBFC046914D479C12A" box="[997,1135,1099,1120]" gridcol="3" gridrow="19" pageId="4" pageNumber="112">2.73</td>
|
||||
<td id="97517029FFCA76CBFB536914D530C12A" box="[1202,1318,1099,1120]" gridcol="4" gridrow="19" pageId="4" pageNumber="112">2.61</td>
|
||||
<td id="97517029FFCA76CBFAB46914D5C3C12A" box="[1365,1493,1099,1120]" gridcol="5" gridrow="19" pageId="4" pageNumber="112">2.70</td>
|
||||
<td id="97517029FFCA76CBF9EE6914D68DC12A" box="[1551,1691,1099,1120]" gridcol="6" gridrow="19" pageId="4" pageNumber="112">2.27</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416938D68DC136" box="[160,1691,1127,1148]" gridrow="20" pageId="4" pageNumber="112" rowspan-0="1">
|
||||
<td id="97517029FFCA76CBFDF56938D2D1C136" box="[532,711,1127,1148]" gridcol="1" gridrow="20" pageId="4" pageNumber="112">H:</td>
|
||||
<td id="97517029FFCA76CBFCFA6938D3B0C136" box="[795,934,1127,1148]" gridcol="2" gridrow="20" pageId="4" pageNumber="112">2.62</td>
|
||||
<td id="97517029FFCA76CBFC046938D479C136" box="[997,1135,1127,1148]" gridcol="3" gridrow="20" pageId="4" pageNumber="112">2.56</td>
|
||||
<td id="97517029FFCA76CBFB536938D530C136" box="[1202,1318,1127,1148]" gridcol="4" gridrow="20" pageId="4" pageNumber="112">2.52</td>
|
||||
<td id="97517029FFCA76CBFAB46938D5C3C136" box="[1365,1493,1127,1148]" gridcol="5" gridrow="20" pageId="4" pageNumber="112">2.57</td>
|
||||
<td id="97517029FFCA76CBF9EE6938D68DC136" box="[1551,1691,1127,1148]" gridcol="6" gridrow="20" pageId="4" pageNumber="112">2.20</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF4169DBD68DC1D3" box="[160,1691,1156,1177]" gridrow="21" pageId="4" pageNumber="112">
|
||||
<th id="97517029FFCA76CBFF4169DBD19AC1D3" box="[160,396,1156,1177]" gridcol="0" gridrow="21" pageId="4" pageNumber="112">Metasomal segment V:</th>
|
||||
<td id="97517029FFCA76CBFDF569DBD2D1C1D3" box="[532,711,1156,1177]" gridcol="1" gridrow="21" pageId="4" pageNumber="112">L:</td>
|
||||
<td id="97517029FFCA76CBFCFA69DBD3B0C1D3" box="[795,934,1156,1177]" gridcol="2" gridrow="21" pageId="4" pageNumber="112">7.04</td>
|
||||
<td id="97517029FFCA76CBFC0469DBD479C1D3" box="[997,1135,1156,1177]" gridcol="3" gridrow="21" pageId="4" pageNumber="112">6.85</td>
|
||||
<td id="97517029FFCA76CBFB5369DBD530C1D3" box="[1202,1318,1156,1177]" gridcol="4" gridrow="21" pageId="4" pageNumber="112">6.74</td>
|
||||
<td id="97517029FFCA76CBFAB469DBD5C3C1D3" box="[1365,1493,1156,1177]" gridcol="5" gridrow="21" pageId="4" pageNumber="112">6.88</td>
|
||||
<td id="97517029FFCA76CBF9EE69DBD68DC1D3" box="[1551,1691,1156,1177]" gridcol="6" gridrow="21" pageId="4" pageNumber="112">5.71</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF4169FFD68DC1FF" box="[160,1691,1184,1205]" gridrow="22" pageId="4" pageNumber="112" rowspan-0="1">
|
||||
<td id="97517029FFCA76CBFDF569FFD2D1C1FF" box="[532,711,1184,1205]" gridcol="1" gridrow="22" pageId="4" pageNumber="112">W:</td>
|
||||
<td id="97517029FFCA76CBFCFA69FFD3B0C1FF" box="[795,934,1184,1205]" gridcol="2" gridrow="22" pageId="4" pageNumber="112">2.77</td>
|
||||
<td id="97517029FFCA76CBFC0469FFD479C1FF" box="[997,1135,1184,1205]" gridcol="3" gridrow="22" pageId="4" pageNumber="112">2.76</td>
|
||||
<td id="97517029FFCA76CBFB5369FFD530C1FF" box="[1202,1318,1184,1205]" gridcol="4" gridrow="22" pageId="4" pageNumber="112">2.62</td>
|
||||
<td id="97517029FFCA76CBFAB469FFD5C3C1FF" box="[1365,1493,1184,1205]" gridcol="5" gridrow="22" pageId="4" pageNumber="112">2.72</td>
|
||||
<td id="97517029FFCA76CBF9EE69FFD68DC1FF" box="[1551,1691,1184,1205]" gridcol="6" gridrow="22" pageId="4" pageNumber="112">2.21</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF4169E3D68DC19B" box="[160,1691,1212,1233]" gridrow="23" pageId="4" pageNumber="112" rowspan-0="1">
|
||||
<td id="97517029FFCA76CBFDF569E3D2D1C19B" box="[532,711,1212,1233]" gridcol="1" gridrow="23" pageId="4" pageNumber="112">H:</td>
|
||||
<td id="97517029FFCA76CBFCFA69E3D3B0C19B" box="[795,934,1212,1233]" gridcol="2" gridrow="23" pageId="4" pageNumber="112">2.86</td>
|
||||
<td id="97517029FFCA76CBFC0469E3D479C19B" box="[997,1135,1212,1233]" gridcol="3" gridrow="23" pageId="4" pageNumber="112">2.62</td>
|
||||
<td id="97517029FFCA76CBFB5369E3D530C19B" box="[1202,1318,1212,1233]" gridcol="4" gridrow="23" pageId="4" pageNumber="112">2.73</td>
|
||||
<td id="97517029FFCA76CBFAB469E3D5C3C19B" box="[1365,1493,1212,1233]" gridcol="5" gridrow="23" pageId="4" pageNumber="112">2.74</td>
|
||||
<td id="97517029FFCA76CBF9EE69E3D68DC19B" box="[1551,1691,1212,1233]" gridcol="6" gridrow="23" pageId="4" pageNumber="112">2.30</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416988D68DC1A6" box="[160,1691,1239,1260]" gridrow="24" pageId="4" pageNumber="112">
|
||||
<th id="97517029FFCA76CBFF416988D19AC1A6" box="[160,396,1239,1260]" gridcol="0" gridrow="24" pageId="4" pageNumber="112">Telson:</th>
|
||||
<td id="97517029FFCA76CBFDF56988D2D1C1A6" box="[532,711,1239,1260]" gridcol="1" gridrow="24" pageId="4" pageNumber="112">L:</td>
|
||||
<td id="97517029FFCA76CBFCFA6988D3B0C1A6" box="[795,934,1239,1260]" gridcol="2" gridrow="24" pageId="4" pageNumber="112">5.77</td>
|
||||
<td id="97517029FFCA76CBFC046988D479C1A6" box="[997,1135,1239,1260]" gridcol="3" gridrow="24" pageId="4" pageNumber="112">5.63</td>
|
||||
<td id="97517029FFCA76CBFB536988D530C1A6" box="[1202,1318,1239,1260]" gridcol="4" gridrow="24" pageId="4" pageNumber="112">5.52</td>
|
||||
<td id="97517029FFCA76CBFAB46988D5C3C1A6" box="[1365,1493,1239,1260]" gridcol="5" gridrow="24" pageId="4" pageNumber="112">5.64</td>
|
||||
<td id="97517029FFCA76CBF9EE6988D68DC1A6" box="[1551,1691,1239,1260]" gridcol="6" gridrow="24" pageId="4" pageNumber="112">4.70</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF4169ABD68DC043" box="[160,1691,1268,1289]" gridrow="25" pageId="4" pageNumber="112" rowspan-0="1">
|
||||
<td id="97517029FFCA76CBFDF569ABD2D1C043" box="[532,711,1268,1289]" gridcol="1" gridrow="25" pageId="4" pageNumber="112">Vesicle W:</td>
|
||||
<td id="97517029FFCA76CBFCFA69ABD3B0C043" box="[795,934,1268,1289]" gridcol="2" gridrow="25" pageId="4" pageNumber="112">2.53</td>
|
||||
<td id="97517029FFCA76CBFC0469ABD479C043" box="[997,1135,1268,1289]" gridcol="3" gridrow="25" pageId="4" pageNumber="112">2.44</td>
|
||||
<td id="97517029FFCA76CBFB5369ABD530C043" box="[1202,1318,1268,1289]" gridcol="4" gridrow="25" pageId="4" pageNumber="112">2.37</td>
|
||||
<td id="97517029FFCA76CBFAB469ABD5C3C043" box="[1365,1493,1268,1289]" gridcol="5" gridrow="25" pageId="4" pageNumber="112">2.45</td>
|
||||
<td id="97517029FFCA76CBF9EE69ABD68DC043" box="[1551,1691,1268,1289]" gridcol="6" gridrow="25" pageId="4" pageNumber="112">1.90</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF41684FD68DC06F" box="[160,1691,1296,1317]" gridrow="26" pageId="4" pageNumber="112" rowspan-0="1">
|
||||
<td id="97517029FFCA76CBFDF5684FD2D1C06F" box="[532,711,1296,1317]" gridcol="1" gridrow="26" pageId="4" pageNumber="112">Vesicle H:</td>
|
||||
<td id="97517029FFCA76CBFCFA684FD3B0C06F" box="[795,934,1296,1317]" gridcol="2" gridrow="26" pageId="4" pageNumber="112">2.38</td>
|
||||
<td id="97517029FFCA76CBFC04684FD479C06F" box="[997,1135,1296,1317]" gridcol="3" gridrow="26" pageId="4" pageNumber="112">2.34</td>
|
||||
<td id="97517029FFCA76CBFB53684FD530C06F" box="[1202,1318,1296,1317]" gridcol="4" gridrow="26" pageId="4" pageNumber="112">2.28</td>
|
||||
<td id="97517029FFCA76CBFAB4684FD5C3C06F" box="[1365,1493,1296,1317]" gridcol="5" gridrow="26" pageId="4" pageNumber="112">2.33</td>
|
||||
<td id="97517029FFCA76CBF9EE684FD68DC06F" box="[1551,1691,1296,1317]" gridcol="6" gridrow="26" pageId="4" pageNumber="112">1.90</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416879D68DC00B" box="[160,1691,1318,1345]" gridrow="27" pageId="4" pageNumber="112">
|
||||
<th id="97517029FFCA76CBFF416879D19AC00B" box="[160,396,1318,1345]" gridcol="0" gridrow="27" pageId="4" pageNumber="112">Pedipalp:</th>
|
||||
<td id="97517029FFCA76CBFDF56879D2D1C00B" box="[532,711,1318,1345]" gridcol="1" gridrow="27" pageId="4" pageNumber="112">Total L:4</td>
|
||||
<td id="97517029FFCA76CBFCFA6879D3B0C00B" box="[795,934,1318,1345]" gridcol="2" gridrow="27" pageId="4" pageNumber="112">25.49</td>
|
||||
<td id="97517029FFCA76CBFC046879D479C00B" box="[997,1135,1318,1345]" gridcol="3" gridrow="27" pageId="4" pageNumber="112">25.10</td>
|
||||
<td id="97517029FFCA76CBFB536879D530C00B" box="[1202,1318,1318,1345]" gridcol="4" gridrow="27" pageId="4" pageNumber="112">24.50</td>
|
||||
<td id="97517029FFCA76CBFAB46879D5C3C00B" box="[1365,1493,1318,1345]" gridcol="5" gridrow="27" pageId="4" pageNumber="112">25.03</td>
|
||||
<td id="97517029FFCA76CBF9EE6879D68DC00B" box="[1551,1691,1318,1345]" gridcol="6" gridrow="27" pageId="4" pageNumber="112">20.25</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416817D68DC017" box="[160,1691,1352,1373]" gridrow="28" pageId="4" pageNumber="112">
|
||||
<th id="97517029FFCA76CBFF416817D19AC017" box="[160,396,1352,1373]" gridcol="0" gridrow="28" pageId="4" pageNumber="112">Pedipalp femur:</th>
|
||||
<td id="97517029FFCA76CBFDF56817D2D1C017" box="[532,711,1352,1373]" gridcol="1" gridrow="28" pageId="4" pageNumber="112">L:</td>
|
||||
<td id="97517029FFCA76CBFCFA6817D3B0C017" box="[795,934,1352,1373]" gridcol="2" gridrow="28" pageId="4" pageNumber="112">6.27</td>
|
||||
<td id="97517029FFCA76CBFC046817D479C017" box="[997,1135,1352,1373]" gridcol="3" gridrow="28" pageId="4" pageNumber="112">6.24</td>
|
||||
<td id="97517029FFCA76CBFB536817D530C017" box="[1202,1318,1352,1373]" gridcol="4" gridrow="28" pageId="4" pageNumber="112">6.02</td>
|
||||
<td id="97517029FFCA76CBFAB46817D5C3C017" box="[1365,1493,1352,1373]" gridcol="5" gridrow="28" pageId="4" pageNumber="112">6.18</td>
|
||||
<td id="97517029FFCA76CBF9EE6817D68DC017" box="[1551,1691,1352,1373]" gridcol="6" gridrow="28" pageId="4" pageNumber="112">5.19</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF41683BD68DC033" box="[160,1691,1380,1401]" gridrow="29" pageId="4" pageNumber="112" rowspan-0="1">
|
||||
<td id="97517029FFCA76CBFDF5683BD2D1C033" box="[532,711,1380,1401]" gridcol="1" gridrow="29" pageId="4" pageNumber="112">W:</td>
|
||||
<td id="97517029FFCA76CBFCFA683BD3B0C033" box="[795,934,1380,1401]" gridcol="2" gridrow="29" pageId="4" pageNumber="112">1.82</td>
|
||||
<td id="97517029FFCA76CBFC04683BD479C033" box="[997,1135,1380,1401]" gridcol="3" gridrow="29" pageId="4" pageNumber="112">1.81</td>
|
||||
<td id="97517029FFCA76CBFB53683BD530C033" box="[1202,1318,1380,1401]" gridcol="4" gridrow="29" pageId="4" pageNumber="112">1.74</td>
|
||||
<td id="97517029FFCA76CBFAB4683BD5C3C033" box="[1365,1493,1380,1401]" gridcol="5" gridrow="29" pageId="4" pageNumber="112">1.79</td>
|
||||
<td id="97517029FFCA76CBF9EE683BD68DC033" box="[1551,1691,1380,1401]" gridcol="6" gridrow="29" pageId="4" pageNumber="112">1.50</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF4168DFD68DC0DF" box="[160,1691,1408,1429]" gridrow="30" pageId="4" pageNumber="112" rowspan-0="1">
|
||||
<td id="97517029FFCA76CBFDF568DFD2D1C0DF" box="[532,711,1408,1429]" gridcol="1" gridrow="30" pageId="4" pageNumber="112">H:</td>
|
||||
<td id="97517029FFCA76CBFCFA68DFD3B0C0DF" box="[795,934,1408,1429]" gridcol="2" gridrow="30" pageId="4" pageNumber="112">1.21</td>
|
||||
<td id="97517029FFCA76CBFC0468DFD479C0DF" box="[997,1135,1408,1429]" gridcol="3" gridrow="30" pageId="4" pageNumber="112">1.18</td>
|
||||
<td id="97517029FFCA76CBFB5368DFD530C0DF" box="[1202,1318,1408,1429]" gridcol="4" gridrow="30" pageId="4" pageNumber="112">1.19</td>
|
||||
<td id="97517029FFCA76CBFAB468DFD5C3C0DF" box="[1365,1493,1408,1429]" gridcol="5" gridrow="30" pageId="4" pageNumber="112">1.19</td>
|
||||
<td id="97517029FFCA76CBF9EE68DFD68DC0DF" box="[1551,1691,1408,1429]" gridcol="6" gridrow="30" pageId="4" pageNumber="112">1.10</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF4168C3D68DC0FB" box="[160,1691,1436,1457]" gridrow="31" pageId="4" pageNumber="112">
|
||||
<th id="97517029FFCA76CBFF4168C3D19AC0FB" box="[160,396,1436,1457]" gridcol="0" gridrow="31" pageId="4" pageNumber="112">Pedipalp patella:</th>
|
||||
<td id="97517029FFCA76CBFDF568C3D2D1C0FB" box="[532,711,1436,1457]" gridcol="1" gridrow="31" pageId="4" pageNumber="112">L:</td>
|
||||
<td id="97517029FFCA76CBFCFA68C3D3B0C0FB" box="[795,934,1436,1457]" gridcol="2" gridrow="31" pageId="4" pageNumber="112">7.02</td>
|
||||
<td id="97517029FFCA76CBFC0468C3D479C0FB" box="[997,1135,1436,1457]" gridcol="3" gridrow="31" pageId="4" pageNumber="112">6.86</td>
|
||||
<td id="97517029FFCA76CBFB5368C3D530C0FB" box="[1202,1318,1436,1457]" gridcol="4" gridrow="31" pageId="4" pageNumber="112">6.68</td>
|
||||
<td id="97517029FFCA76CBFAB468C3D5C3C0FB" box="[1365,1493,1436,1457]" gridcol="5" gridrow="31" pageId="4" pageNumber="112">6.85</td>
|
||||
<td id="97517029FFCA76CBF9EE68C3D68DC0FB" box="[1551,1691,1436,1457]" gridcol="6" gridrow="31" pageId="4" pageNumber="112">5.65</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF4168E7D68DC087" box="[160,1691,1464,1485]" gridrow="32" pageId="4" pageNumber="112" rowspan-0="1">
|
||||
<td id="97517029FFCA76CBFDF568E7D2D1C087" box="[532,711,1464,1485]" gridcol="1" gridrow="32" pageId="4" pageNumber="112">W:</td>
|
||||
<td id="97517029FFCA76CBFCFA68E7D3B0C087" box="[795,934,1464,1485]" gridcol="2" gridrow="32" pageId="4" pageNumber="112">2.39</td>
|
||||
<td id="97517029FFCA76CBFC0468E7D479C087" box="[997,1135,1464,1485]" gridcol="3" gridrow="32" pageId="4" pageNumber="112">2.35</td>
|
||||
<td id="97517029FFCA76CBFB5368E7D530C087" box="[1202,1318,1464,1485]" gridcol="4" gridrow="32" pageId="4" pageNumber="112">2.29</td>
|
||||
<td id="97517029FFCA76CBFAB468E7D5C3C087" box="[1365,1493,1464,1485]" gridcol="5" gridrow="32" pageId="4" pageNumber="112">2.34</td>
|
||||
<td id="97517029FFCA76CBF9EE68E7D68DC087" box="[1551,1691,1464,1485]" gridcol="6" gridrow="32" pageId="4" pageNumber="112">1.88</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF41688BD68DC0A3" box="[160,1691,1492,1513]" gridrow="33" pageId="4" pageNumber="112" rowspan-0="1">
|
||||
<td id="97517029FFCA76CBFDF5688BD2D1C0A3" box="[532,711,1492,1513]" gridcol="1" gridrow="33" pageId="4" pageNumber="112">H:</td>
|
||||
<td id="97517029FFCA76CBFCFA688BD3B0C0A3" box="[795,934,1492,1513]" gridcol="2" gridrow="33" pageId="4" pageNumber="112">1.80</td>
|
||||
<td id="97517029FFCA76CBFC04688BD479C0A3" box="[997,1135,1492,1513]" gridcol="3" gridrow="33" pageId="4" pageNumber="112">1.79</td>
|
||||
<td id="97517029FFCA76CBFB53688BD530C0A3" box="[1202,1318,1492,1513]" gridcol="4" gridrow="33" pageId="4" pageNumber="112">1.77</td>
|
||||
<td id="97517029FFCA76CBFAB4688BD5C3C0A3" box="[1365,1493,1492,1513]" gridcol="5" gridrow="33" pageId="4" pageNumber="112">1.79</td>
|
||||
<td id="97517029FFCA76CBF9EE688BD68DC0A3" box="[1551,1691,1492,1513]" gridcol="6" gridrow="33" pageId="4" pageNumber="112">1.40</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF4168AFD68DC34F" box="[160,1691,1520,1541]" gridrow="34" pageId="4" pageNumber="112">
|
||||
<th id="97517029FFCA76CBFF4168AFD19AC34F" box="[160,396,1520,1541]" gridcol="0" gridrow="34" pageId="4" pageNumber="112">Pedipalp chela:</th>
|
||||
<td id="97517029FFCA76CBFDF568AFD2D1C34F" box="[532,711,1520,1541]" gridcol="1" gridrow="34" pageId="4" pageNumber="112">L:</td>
|
||||
<td id="97517029FFCA76CBFCFA68AFD3B0C34F" box="[795,934,1520,1541]" gridcol="2" gridrow="34" pageId="4" pageNumber="112">12.20</td>
|
||||
<td id="97517029FFCA76CBFC0468AFD479C34F" box="[997,1135,1520,1541]" gridcol="3" gridrow="34" pageId="4" pageNumber="112">12.00</td>
|
||||
<td id="97517029FFCA76CBFB5368AFD530C34F" box="[1202,1318,1520,1541]" gridcol="4" gridrow="34" pageId="4" pageNumber="112">11.80</td>
|
||||
<td id="97517029FFCA76CBFAB468AFD5C3C34F" box="[1365,1493,1520,1541]" gridcol="5" gridrow="34" pageId="4" pageNumber="112">12.00</td>
|
||||
<td id="97517029FFCA76CBF9EE68AFD68DC34F" box="[1551,1691,1520,1541]" gridcol="6" gridrow="34" pageId="4" pageNumber="112">9.41</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416B53D68DC36B" box="[160,1691,1548,1569]" gridrow="35" pageId="4" pageNumber="112" rowspan-0="1">
|
||||
<td id="97517029FFCA76CBFDF56B53D2D1C36B" box="[532,711,1548,1569]" gridcol="1" gridrow="35" pageId="4" pageNumber="112">W:</td>
|
||||
<td id="97517029FFCA76CBFCFA6B53D3B0C36B" box="[795,934,1548,1569]" gridcol="2" gridrow="35" pageId="4" pageNumber="112">2.53</td>
|
||||
<td id="97517029FFCA76CBFC046B53D479C36B" box="[997,1135,1548,1569]" gridcol="3" gridrow="35" pageId="4" pageNumber="112">2.62</td>
|
||||
<td id="97517029FFCA76CBFB536B53D530C36B" box="[1202,1318,1548,1569]" gridcol="4" gridrow="35" pageId="4" pageNumber="112">2.47</td>
|
||||
<td id="97517029FFCA76CBFAB46B53D5C3C36B" box="[1365,1493,1548,1569]" gridcol="5" gridrow="35" pageId="4" pageNumber="112">2.54</td>
|
||||
<td id="97517029FFCA76CBF9EE6B53D68DC36B" box="[1551,1691,1548,1569]" gridcol="6" gridrow="35" pageId="4" pageNumber="112">1.94</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416B77D68DC377" box="[160,1691,1576,1597]" gridrow="36" pageId="4" pageNumber="112" rowspan-0="1">
|
||||
<td id="97517029FFCA76CBFDF56B77D2D1C377" box="[532,711,1576,1597]" gridcol="1" gridrow="36" pageId="4" pageNumber="112">H:</td>
|
||||
<td id="97517029FFCA76CBFCFA6B77D3B0C377" box="[795,934,1576,1597]" gridcol="2" gridrow="36" pageId="4" pageNumber="112">2.38</td>
|
||||
<td id="97517029FFCA76CBFC046B77D479C377" box="[997,1135,1576,1597]" gridcol="3" gridrow="36" pageId="4" pageNumber="112">2.49</td>
|
||||
<td id="97517029FFCA76CBFB536B77D530C377" box="[1202,1318,1576,1597]" gridcol="4" gridrow="36" pageId="4" pageNumber="112">2.34</td>
|
||||
<td id="97517029FFCA76CBFAB46B77D5C3C377" box="[1365,1493,1576,1597]" gridcol="5" gridrow="36" pageId="4" pageNumber="112">2.40</td>
|
||||
<td id="97517029FFCA76CBF9EE6B77D68DC377" box="[1551,1691,1576,1597]" gridcol="6" gridrow="36" pageId="4" pageNumber="112">1.83</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416B61D68DC313" box="[160,1691,1598,1625]" gridrow="37" pageId="4" pageNumber="112" rowspan-0="1">
|
||||
<td id="97517029FFCA76CBFDF56B61D2D1C313" box="[532,711,1598,1625]" gridcol="1" gridrow="37" pageId="4" pageNumber="112">Fixed finger L:5</td>
|
||||
<td id="97517029FFCA76CBFCFA6B61D3B0C313" box="[795,934,1598,1625]" gridcol="2" gridrow="37" pageId="4" pageNumber="112">7.16</td>
|
||||
<td id="97517029FFCA76CBFC046B61D479C313" box="[997,1135,1598,1625]" gridcol="3" gridrow="37" pageId="4" pageNumber="112">6.91</td>
|
||||
<td id="97517029FFCA76CBFB536B61D530C313" box="[1202,1318,1598,1625]" gridcol="4" gridrow="37" pageId="4" pageNumber="112">6.73</td>
|
||||
<td id="97517029FFCA76CBFAB46B61D5C3C313" box="[1365,1493,1598,1625]" gridcol="5" gridrow="37" pageId="4" pageNumber="112">6.93</td>
|
||||
<td id="97517029FFCA76CBF9EE6B61D68DC313" box="[1551,1691,1598,1625]" gridcol="6" gridrow="37" pageId="4" pageNumber="112">5.40</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416B3FD68DC33F" box="[160,1691,1632,1653]" gridrow="38" pageId="4" pageNumber="112" rowspan-0="1">
|
||||
<td id="97517029FFCA76CBFDF56B3FD2D1C33F" box="[532,711,1632,1653]" gridcol="1" gridrow="38" pageId="4" pageNumber="112">Movable finger L:</td>
|
||||
<td id="97517029FFCA76CBFCFA6B3FD3B0C33F" box="[795,934,1632,1653]" gridcol="2" gridrow="38" pageId="4" pageNumber="112">8.03</td>
|
||||
<td id="97517029FFCA76CBFC046B3FD479C33F" box="[997,1135,1632,1653]" gridcol="3" gridrow="38" pageId="4" pageNumber="112">7.78</td>
|
||||
<td id="97517029FFCA76CBFB536B3FD530C33F" box="[1202,1318,1632,1653]" gridcol="4" gridrow="38" pageId="4" pageNumber="112">7.67</td>
|
||||
<td id="97517029FFCA76CBFAB46B3FD5C3C33F" box="[1365,1493,1632,1653]" gridcol="5" gridrow="38" pageId="4" pageNumber="112">7.83</td>
|
||||
<td id="97517029FFCA76CBF9EE6B3FD68DC33F" box="[1551,1691,1632,1653]" gridcol="6" gridrow="38" pageId="4" pageNumber="112">6.30</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416B22D68DC3D8" box="[160,1691,1661,1682]" gridrow="39" pageId="4" pageNumber="112" rowspan-0="1">
|
||||
<td id="97517029FFCA76CBFDF56B22D2D1C3D8" box="[532,711,1661,1682]" gridcol="1" gridrow="39" pageId="4" pageNumber="112">Palm L:</td>
|
||||
<td id="97517029FFCA76CBFCFA6B22D3B0C3D8" box="[795,934,1661,1682]" gridcol="2" gridrow="39" pageId="4" pageNumber="112">4.09</td>
|
||||
<td id="97517029FFCA76CBFC046B22D479C3D8" box="[997,1135,1661,1682]" gridcol="3" gridrow="39" pageId="4" pageNumber="112">4.20</td>
|
||||
<td id="97517029FFCA76CBFB536B22D530C3D8" box="[1202,1318,1661,1682]" gridcol="4" gridrow="39" pageId="4" pageNumber="112">4.16</td>
|
||||
<td id="97517029FFCA76CBFAB46B22D5C3C3D8" box="[1365,1493,1661,1682]" gridcol="5" gridrow="39" pageId="4" pageNumber="112">4.15</td>
|
||||
<td id="97517029FFCA76CBF9EE6B22D68DC3D8" box="[1551,1691,1661,1682]" gridcol="6" gridrow="39" pageId="4" pageNumber="112">3.20</td>
|
||||
</tr>
|
||||
<tr id="D4801955FFCA76CBFF416BCCD68DC3E4" box="[160,1691,1683,1710]" gridrow="40" pageId="4" pageNumber="112">
|
||||
<th id="97517029FFCA76CBFF416BCCD19AC3E4" box="[160,396,1683,1710]" gridcol="0" gridrow="40" pageId="4" pageNumber="112">Pectines:</th>
|
||||
<td id="97517029FFCA76CBFDF56BCCD2D1C3E4" box="[532,711,1683,1710]" gridcol="1" gridrow="40" pageId="4" pageNumber="112">L:6</td>
|
||||
<td id="97517029FFCA76CBFCFA6BCCD3B0C3E4" box="[795,934,1683,1710]" gridcol="2" gridrow="40" pageId="4" pageNumber="112">3.89</td>
|
||||
<td id="97517029FFCA76CBFC046BCCD479C3E4" box="[997,1135,1683,1710]" gridcol="3" gridrow="40" pageId="4" pageNumber="112">3.60</td>
|
||||
<td id="97517029FFCA76CBFB536BCCD530C3E4" box="[1202,1318,1683,1710]" gridcol="4" gridrow="40" pageId="4" pageNumber="112">3.67</td>
|
||||
<td id="97517029FFCA76CBFAB46BCCD5C3C3E4" box="[1365,1493,1683,1710]" gridcol="5" gridrow="40" pageId="4" pageNumber="112">3.72</td>
|
||||
<td id="97517029FFCA76CBF9EE6BCCD68DC3E4" box="[1551,1691,1683,1710]" gridcol="6" gridrow="40" pageId="4" pageNumber="112">3.00</td>
|
||||
</tr>
|
||||
</table>
|
||||
</paragraph>
|
||||
<caption id="3ECF4B9FFFCB8931FF766A79D4A0C211" ID-DOI="http://doi.org/10.5281/zenodo.4915131" ID-Zenodo-Dep="4915131" httpUri="https://zenodo.org/record/4915131/files/figure.png" pageId="5" pageNumber="113" startId="5.[151,264,1830,1853]" targetBox="[178,1424,213,1778]" targetPageId="5">
|
||||
<paragraph id="6A0F1B17FFCB8931FF766A79D4A0C211" blockId="5.[151,1436,1830,1883]" pageId="5" pageNumber="113">
|
||||
<emphasis id="58C4C705FFCB8931FF766A79D12DC276" bold="true" box="[151,315,1830,1853]" pageId="5" pageNumber="113">FIGURES 4–7.</emphasis>
|
||||
<taxonomicName id="ADB06094FFCB8931FEA36A78D274C276" authority="Botero-Trujillo & Flórez, 2014" authorityName="Botero-Trujillo & Flórez" authorityYear="2014" box="[322,610,1830,1852]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="113" phylum="Arthropoda" rank="species" species="grottoedensis" status="sp. nov." subGenus="Tityus">
|
||||
<emphasis id="58C4C705FFCB8931FEA36A78D274C276" box="[322,610,1830,1852]" italics="true" pageId="5" pageNumber="113">Tityus (Tityus) grottoedensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="58C4C705FFCB8931FD886A78D2ACC276" bold="true" box="[617,698,1831,1852]" pageId="5" pageNumber="113">
|
||||
<taxonomicNameLabel id="43F77A7EFFCB8931FD886A78D2ACC276" box="[617,698,1831,1852]" pageId="5" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, habitus.
|
||||
<emphasis id="58C4C705FFCB8931FCFE6A78D35DC276" bold="true" box="[799,843,1831,1852]" pageId="5" pageNumber="113">4–5.</emphasis>
|
||||
Subadult paratype male, dorsal and ventral aspects (ICN- As-905).
|
||||
<emphasis id="58C4C705FFCB8931FF176A1AD135C210" bold="true" box="[246,291,1861,1882]" pageId="5" pageNumber="113">6–7.</emphasis>
|
||||
One paratype female, dorsal and ventral aspects (MACN-Ar 30879). Scale bars = 10 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="3ECF4B9FFFC88932FF766877D380C036" ID-DOI="http://doi.org/10.5281/zenodo.4915133" ID-Zenodo-Dep="4915133" httpUri="https://zenodo.org/record/4915133/files/figure.png" pageId="6" pageNumber="114" startId="6.[151,264,1320,1343]" targetBox="[159,1432,213,1269]" targetPageId="6">
|
||||
<paragraph id="6A0F1B17FFC88932FF766877D380C036" blockId="6.[151,1436,1320,1404]" pageId="6" pageNumber="114">
|
||||
<emphasis id="58C4C705FFC88932FF766877D15CC074" bold="true" box="[151,330,1320,1343]" pageId="6" pageNumber="114">FIGURES 8–13.</emphasis>
|
||||
<taxonomicName id="ADB06094FFC88932FEB36875D263C075" authority="Botero-Trujillo & Flórez, 2014" authorityName="Botero-Trujillo & Flórez" authorityYear="2014" box="[338,629,1321,1343]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="114" phylum="Arthropoda" rank="species" species="grottoedensis" status="sp. nov." subGenus="Tityus">
|
||||
<emphasis id="58C4C705FFC88932FEB36875D263C075" box="[338,629,1321,1343]" italics="true" pageId="6" pageNumber="114">Tityus (Tityus) grottoedensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="58C4C705FFC88932FD9F6876D2C7C074" bold="true" box="[638,721,1321,1342]" pageId="6" pageNumber="114">
|
||||
<taxonomicNameLabel id="43F77A7EFFC88932FD9F6876D2C7C074" box="[638,721,1321,1342]" pageId="6" pageNumber="114" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, subadult paratype male (ICN-As-905), trichobothrial distribution in right pedipalp.
|
||||
<emphasis id="58C4C705FFC88932FED76818D15EC016" bold="true" box="[310,328,1351,1372]" pageId="6" pageNumber="114">8.</emphasis>
|
||||
Femur, dorsal aspect.
|
||||
<emphasis id="58C4C705FFC88932FDD66818D25FC016" bold="true" box="[567,585,1351,1372]" pageId="6" pageNumber="114">9.</emphasis>
|
||||
Femur, internal aspect.
|
||||
<emphasis id="58C4C705FFC88932FCA66818D371C016" bold="true" box="[839,871,1351,1372]" pageId="6" pageNumber="114">10.</emphasis>
|
||||
Patella, dorsal aspect.
|
||||
<emphasis id="58C4C705FFC88932FBB96818D461C016" bold="true" box="[1112,1143,1351,1372]" pageId="6" pageNumber="114">11.</emphasis>
|
||||
Patella, external aspect.
|
||||
<emphasis id="58C4C705FFC88932FA9D6818D58AC016" bold="true" box="[1404,1436,1351,1372]" pageId="6" pageNumber="114">12.</emphasis>
|
||||
Chela, external aspect.
|
||||
<emphasis id="58C4C705FFC88932FE636839D1B4C031" bold="true" box="[386,418,1382,1403]" pageId="6" pageNumber="114">13.</emphasis>
|
||||
Chela, ventral aspect (palm). Scale bars = 1 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="6A0F1B17FFC88932FF2668FCD294C0F6" blockId="6.[151,1437,1443,2008]" box="[199,642,1443,1468]" pageId="6" pageNumber="114">
|
||||
<emphasis id="58C4C705FFC88932FF2668FCD14CC0F6" bold="true" box="[199,346,1443,1468]" inLineHeading="true" italics="true" pageId="6" pageNumber="114" reason="1">Morphology.</emphasis>
|
||||
Measurements in
|
||||
<tableCitation id="27322EACFFC88932FDC968FCD26AC0F6" box="[552,636,1443,1468]" captionStart="TABLE 1" captionStartId="4.[147,230,454,475]" captionTargetPageId="4" captionText="TABLE 1. Meristic data for Tityus (Tityus) grottoedensis sp. nov. Measurements in millimeters. 1Sum of carapace, mesosoma and metasoma. 2Sum of tergites I– VII. 3Sum of metasomal segments I–V and telson. 4Sum of femur, patella and chela. 5Measured from the commissure of the junction with the movable finger to the finger tip. 6Measured along the marginal lamella." httpUri="http://table.plazi.org/id/3ECF4B9FFFCA8930FF726C99D281C75E" pageId="6" pageNumber="114" tableUuid="3ECF4B9FFFCA8930FF726C99D281C75E">Table 1</tableCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="6A0F1B17FFC88932FF266897D580C33A" blockId="6.[151,1437,1443,2008]" pageId="6" pageNumber="114">
|
||||
<emphasis id="58C4C705FFC88932FF266897D128C095" box="[199,318,1480,1503]" italics="true" pageId="6" pageNumber="114">Carapace:</emphasis>
|
||||
Trapezoidal, narrowing anteriorly and granulose throughout; anterior margin subtly emarginated; median ocular tubercle anterior to the centre of the carapace, with pair of medium (normal) sized ocelli; lateral ocular tubercles each with three ocelli; lateral ocular carinae present; anterior median and superciliary carinae continuous, as well as central median and posterior median carinae; all carinae pronounced and elevated; anterior median, median ocular, central median, posterior median, posterior marginal and posterior lateral furrows evident.
|
||||
</paragraph>
|
||||
<paragraph id="6A0F1B17FFC88932FF266B23D40CC396" blockId="6.[151,1437,1443,2008]" pageId="6" pageNumber="114">
|
||||
<emphasis id="58C4C705FFC88932FF266B23D13EC3D9" box="[199,296,1660,1683]" italics="true" pageId="6" pageNumber="114">Tergites:</emphasis>
|
||||
With similar granulation to that of the carapace; three strong longitudinal carinae (medial; paired dorsolateral) present on the posterior half of tergites I–VI; tergite VII with dorsolateral and lateral carinae complete, and medial carina represented by a granulose anteromedian elevation.
|
||||
</paragraph>
|
||||
<paragraph id="6A0F1B17FFC88932FF266BB8D383C202" blockId="6.[151,1437,1443,2008]" pageId="6" pageNumber="114">
|
||||
<emphasis id="58C4C705FFC88932FF266BB8D1A2C3B5" box="[199,436,1767,1792]" italics="true" pageId="6" pageNumber="114">Coxosternal region:</emphasis>
|
||||
Sternum longer than wide, not markedly compressed anteriorly thus giving a subpentagonal appearance, with deep median furrow; all the components of this region smooth, devoid of granules except for the anterior margin of coxae which bear granulose carina.
|
||||
</paragraph>
|
||||
<paragraph id="6A0F1B17FFC88933FF266A0CD0C6C59F" blockId="6.[151,1437,1443,2008]" lastBlockId="7.[151,1437,151,392]" lastPageId="7" lastPageNumber="115" pageId="6" pageNumber="114">
|
||||
<emphasis id="58C4C705FFC88932FF266A0CD229C221" box="[199,575,1875,1900]" italics="true" pageId="6" pageNumber="114">Genital operculum and pectines:</emphasis>
|
||||
Genital operculum divided longitudinally, formed by subtriangular plates; pectines small, not reaching the level of junction of coxae IV with the trochanter; pectinal marginal lamellae with 3/3 pieces; middle lamellae with 8–11 (female) and 10/11 (male); fulcra with 16–18 (female) and 17/19 (male); teeth count 18–19 (female, n=6, mode 19) and 19–20 (male, n=4 including juvenile, mode 19); basal piece of the middle lamella enlarged and oval in both male and female (
|
||||
<figureCitation id="F28B0792FFC98933FCD26DC8D391C5FA" box="[819,903,151,176]" captionStart="FIGURES 14–16" captionStartId="7.[151,264,1801,1824]" captionTargetBox="[158,1428,456,1773]" captionTargetId="figure-223@7.[151,1436,445,1780]" captionTargetPageId="7" captionText="FIGURES 14–16. Tityus (Tityus) grottoedensis sp. nov. 14. Subadult paratype male (ICN-As-905), right lateral aspect of metasoma and telson. 15–16. Holotype female (ICN-As-977). 15. Left pectine. 16. Sternites IV–VII; most granulation details of the sternites not depicted, emphasis made on carinae. Scale bars = 3 mm (Figs. 14, 16), 1 mm (Fig. 15)." figureDoi="http://doi.org/10.5281/zenodo.4915135" httpUri="https://zenodo.org/record/4915135/files/figure.png" pageId="7" pageNumber="115">Fig. 15</figureCitation>
|
||||
); pectinal basal piece approximately as wide as long.
|
||||
</paragraph>
|
||||
<paragraph id="6A0F1B17FFC98933FF266DBFD2CBC4C2" blockId="7.[151,1437,151,392]" pageId="7" pageNumber="115">
|
||||
<emphasis id="58C4C705FFC98933FF266DBFD127C5BD" box="[199,305,224,247]" italics="true" pageId="7" pageNumber="115">Sternites:</emphasis>
|
||||
Thoroughly covered with fine granulation; submedian longitudinal carinae strong, present on posterior two thirds of sternite VII and posterior half of sternite VI, obsolete on V, vestigial on IV, absent on III; lateral longitudinal carinae present medially on sternite VII but not complete, and small but visible medially in sternites IV to VI just beside the book lung spiracles; sternite V with a rather slit-like posterior median hyaline area (gland); book lung spiracles oval elongate (
|
||||
<figureCitation id="F28B0792FFC98933FD9C6C30D2C6C4C2" box="[637,720,367,392]" captionStart="FIGURES 14–16" captionStartId="7.[151,264,1801,1824]" captionTargetBox="[158,1428,456,1773]" captionTargetId="figure-223@7.[151,1436,445,1780]" captionTargetPageId="7" captionText="FIGURES 14–16. Tityus (Tityus) grottoedensis sp. nov. 14. Subadult paratype male (ICN-As-905), right lateral aspect of metasoma and telson. 15–16. Holotype female (ICN-As-977). 15. Left pectine. 16. Sternites IV–VII; most granulation details of the sternites not depicted, emphasis made on carinae. Scale bars = 3 mm (Figs. 14, 16), 1 mm (Fig. 15)." figureDoi="http://doi.org/10.5281/zenodo.4915135" httpUri="https://zenodo.org/record/4915135/files/figure.png" pageId="7" pageNumber="115">Fig. 16</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="3ECF4B9FFFC98933FF766A56D4DCC217" ID-DOI="http://doi.org/10.5281/zenodo.4915135" ID-Zenodo-Dep="4915135" httpUri="https://zenodo.org/record/4915135/files/figure.png" pageId="7" pageNumber="115" startId="7.[151,264,1801,1824]" targetBox="[158,1428,456,1773]" targetPageId="7">
|
||||
<paragraph id="6A0F1B17FFC98933FF766A56D4DCC217" blockId="7.[151,1436,1801,1885]" pageId="7" pageNumber="115">
|
||||
<emphasis id="58C4C705FFC98933FF766A56D141C255" bold="true" box="[151,343,1801,1824]" pageId="7" pageNumber="115">FIGURES 14–16.</emphasis>
|
||||
<taxonomicName id="ADB06094FFC98933FE806A54D290C26A" authority="Botero-Trujillo & Flórez, 2014" authorityName="Botero-Trujillo & Flórez" authorityYear="2014" box="[353,646,1802,1824]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="115" phylum="Arthropoda" rank="species" species="grottoedensis" status="sp. nov." subGenus="Tityus">
|
||||
<emphasis id="58C4C705FFC98933FE806A54D290C26A" box="[353,646,1802,1824]" italics="true" pageId="7" pageNumber="115">Tityus (Tityus) grottoedensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="58C4C705FFC98933FD706A55D319C255" bold="true" box="[657,783,1802,1823]" pageId="7" pageNumber="115">
|
||||
<taxonomicNameLabel id="43F77A7EFFC98933FD706A55D2F3C255" box="[657,741,1802,1823]" pageId="7" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
14.
|
||||
</emphasis>
|
||||
Subadult paratype male (ICN-As-905), right lateral aspect of metasoma and telson.
|
||||
<emphasis id="58C4C705FFC98933FE9A6A77D1D7C277" bold="true" box="[379,449,1832,1853]" pageId="7" pageNumber="115">15–16.</emphasis>
|
||||
Holotype female (ICN-As-977).
|
||||
<emphasis id="58C4C705FFC98933FCF66A77D321C277" bold="true" box="[791,823,1832,1853]" pageId="7" pageNumber="115">15.</emphasis>
|
||||
Left pectine.
|
||||
<emphasis id="58C4C705FFC98933FC266A77D3F0C277" bold="true" box="[967,998,1832,1853]" pageId="7" pageNumber="115">16.</emphasis>
|
||||
Sternites IV–VII; most granulation details of the sternites not depicted, emphasis made on carinae. Scale bars = 3 mm (Figs. 14, 16), 1 mm (Fig. 15).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="6A0F1B17FFC9893CFF266AD4D3FAC49A" blockId="7.[151,1436,1930,2027]" lastBlockId="8.[151,1438,151,2013]" lastPageId="8" lastPageNumber="116" pageId="7" pageNumber="115">
|
||||
<emphasis id="58C4C705FFC98933FF266AD4D153C2E8" box="[199,325,1931,1954]" italics="true" pageId="7" pageNumber="115">Metasoma:</emphasis>
|
||||
Overall metasomal appearance not displaying any remarkable sexual dimorphism (thought to be due to the male specimen being a subadult), segments and telson slender, similar in shape in male and female (
|
||||
<figureCitation id="F28B0792FFC98933FA856AF2D0D2C2A1" captionStart="FIGURES 4–7" captionStartId="5.[151,264,1830,1853]" captionTargetBox="[178,1424,213,1778]" captionTargetId="figure-60@5.[151,1436,193,1809]" captionTargetPageId="5" captionText="FIGURES 4–7. Tityus (Tityus) grottoedensis sp. nov., habitus. 4–5. Subadult paratype male, dorsal and ventral aspects (ICN- As-905). 6–7. One paratype female, dorsal and ventral aspects (MACN-Ar 30879). Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4915131" httpUri="https://zenodo.org/record/4915131/files/figure.png" pageId="7" pageNumber="115">Figs. 4–7</figureCitation>
|
||||
); segment I with ten complete parallel carinae (paired ventrosubmedian, ventrolateral, median lateral, dorsolateral, and dorsosubmedian carinae); segment II with same carination as segment I but with median lateral carinae represented only by one to three elevated granules on the posterior margin (
|
||||
<figureCitation id="F28B0792FFC6893CFBA36DE3D480C59F" box="[1090,1174,188,213]" captionStart="FIGURES 14–16" captionStartId="7.[151,264,1801,1824]" captionTargetBox="[158,1428,456,1773]" captionTargetId="figure-223@7.[151,1436,445,1780]" captionTargetPageId="7" captionText="FIGURES 14–16. Tityus (Tityus) grottoedensis sp. nov. 14. Subadult paratype male (ICN-As-905), right lateral aspect of metasoma and telson. 15–16. Holotype female (ICN-As-977). 15. Left pectine. 16. Sternites IV–VII; most granulation details of the sternites not depicted, emphasis made on carinae. Scale bars = 3 mm (Figs. 14, 16), 1 mm (Fig. 15)." figureDoi="http://doi.org/10.5281/zenodo.4915135" httpUri="https://zenodo.org/record/4915135/files/figure.png" pageId="8" pageNumber="116">Fig. 14</figureCitation>
|
||||
); segments III–IV with eight carinae (median lateral entirely absent); segment V with five carinae (ventromedian, paired ventrolateral and dorsolateral carinae); dorsosubmedian carinae of metasomal segments I–IV with granules subequal in size, without any spinoid or enlarged terminal granule; ventrosubmedian carinae complete and parallel in segments I through IV; all carinae strong and granulose, made up of elevated granules arranged in straight rows; intercarinal spaces with minute granulation similar to that found in the sternites. Telson with evidence of ventromedian, ventrolateral and dorsolateral carinae, and some conspicuous but dispersed (not in row) granules in-between; subaculear tubercle strong and spinoid, with two small dorsal granules; aculeus strongly curved.
|
||||
</paragraph>
|
||||
<paragraph id="6A0F1B17FFC6893CFF266C83D19DC7ED" blockId="8.[151,1438,151,2013]" pageId="8" pageNumber="116">
|
||||
<emphasis id="58C4C705FFC6893CFF266C83D15CC4BF" box="[199,330,476,501]" italics="true" pageId="8" pageNumber="116">Chelicerae:</emphasis>
|
||||
Cheliceral dentition characteristic of the family
|
||||
<taxonomicName id="ADB06094FFC6893CFC4B6C83D4C7C4BF" authority="(Vachon 1963)" baseAuthorityName="Vachon" baseAuthorityYear="1963" box="[938,1233,476,501]" class="Arachnida" family="Buthidae" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="116" phylum="Arthropoda" rank="family">
|
||||
Buthidae (
|
||||
<bibRefCitation id="0E2166E6FFC6893CFBC66C83D4DFC4BF" author="Vachon, M." box="[1063,1225,476,501]" pageId="8" pageNumber="116" pagination="161 - 166" refId="ref8678" refString="Vachon, M. (1963) De l'utilite, en systematique, d'une nomenclature des dents des cheliceres chez les scorpions. Bulletin du Museum national de Histoire naturelle Paris, Serie 2 e, 35 (2), 161 - 166." type="journal article" year="1963">Vachon 1963</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
. Movable finger externally with two small basal teeth, one median pronounced tooth, one subdistal tooth slightly shorter than the median, and one distal tooth. Fixed finger externally with one basal and one median tooth mounted onto a bicuspid, one subdistal, and one distal tooth. Fixed finger ventrally with only one subdistal tooth slightly basal to the dorsal subdistal. Movable finger ventrally with three strong teeth, one basal, one median, and one distal tooth larger than its dorsal counterpart.
|
||||
</paragraph>
|
||||
<paragraph id="6A0F1B17FFC6893CFF266FEBD3A7C137" blockId="8.[151,1438,151,2013]" pageId="8" pageNumber="116">
|
||||
<emphasis id="58C4C705FFC6893CFF266FEBD156C787" box="[199,320,692,717]" italics="true" pageId="8" pageNumber="116">Pedipalps:</emphasis>
|
||||
Similar in male and female, not bulkier in male or displaying any remarkable sexual dimorphism (
|
||||
<figureCitation id="F28B0792FFC6893CFF7E6F88D11DC7BA" box="[159,267,727,752]" captionStart="FIGURES 4–7" captionStartId="5.[151,264,1830,1853]" captionTargetBox="[178,1424,213,1778]" captionTargetId="figure-60@5.[151,1436,193,1809]" captionTargetPageId="5" captionText="FIGURES 4–7. Tityus (Tityus) grottoedensis sp. nov., habitus. 4–5. Subadult paratype male, dorsal and ventral aspects (ICN- As-905). 6–7. One paratype female, dorsal and ventral aspects (MACN-Ar 30879). Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4915131" httpUri="https://zenodo.org/record/4915131/files/figure.png" pageId="8" pageNumber="116">Figs. 4–7</figureCitation>
|
||||
) (thought to be due to the male specimen being a subadult); pedipalp hand width (Hw) slightly greater than pedipalp patella width (Pw) (male Hw/Pw = 1,03/1,00; female Hw/Pw = 1,06–1,11/1,00); chela wider than high; pedipalp surfaces with minute granulation similar to that of the sternites and metasoma. Femur with proximomarginal carina and five complete longitudinal carinae (dorso-interior, dorso-exterior, ventro-exterior, ventrointerior carinae, and an additional carina in internal median position); all carinae strong and granulose, made up of elevated granules arranged in straight rows; patella with seven strong longitudinal carinae (dorso-interior, dorsal, dorso-median, dorso-exterior, exterior-median, ventro-exterior and ventro-interior), made up of granules of similar size arranged in straight rows, except for the dorso-interior carina which is discontinuous and has rather spiniform tubercles; chela with nine longitudinal carinae, of similar characteristics to those of the patella except for those on the internal surface which are weaker. Movable finger of pedipalp chelae (male and female, both sides) with 14 oblique rows of granules (including the short apical row); fixed finger with 12 rows. Pedipalps orthobothriotaxic
|
||||
<typeStatus id="B50BA5B5FFC6893CFF76693BD0D9C136" box="[151,207,1124,1148]" pageId="8" pageNumber="116">Type</typeStatus>
|
||||
A (Vachon 1974, 1975); femur with α configuration (
|
||||
<figureCitation id="F28B0792FFC6893CFCC8693BD3B3C137" box="[809,933,1124,1149]" captionStart="FIGURES 8–13" captionStartId="6.[151,264,1320,1343]" captionTargetBox="[159,1432,213,1269]" captionTargetId="figure-295@6.[151,1436,193,1299]" captionTargetPageId="6" captionText="FIGURES 8–13. Tityus (Tityus) grottoedensis sp. nov., subadult paratype male (ICN-As-905), trichobothrial distribution in right pedipalp. 8. Femur, dorsal aspect. 9. Femur, internal aspect. 10. Patella, dorsal aspect. 11. Patella, external aspect. 12. Chela, external aspect. 13. Chela, ventral aspect (palm). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4915133" httpUri="https://zenodo.org/record/4915133/files/figure.png" pageId="8" pageNumber="116">Figs. 8–13</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="6A0F1B17FFC6893CFF2669D6D4F8C189" blockId="8.[151,1438,151,2013]" pageId="8" pageNumber="116">
|
||||
<emphasis id="58C4C705FFC6893CFF2669D6D112C1EA" box="[199,260,1161,1184]" italics="true" pageId="8" pageNumber="116">Legs:</emphasis>
|
||||
Basitarsus and telotarsus with relatively few setae arranged in rather straight parallel rows (mostly evident in legs III and IV); tibial spur absent; prolateral and retrolateral pedal spurs present in I–IV.
|
||||
</paragraph>
|
||||
<paragraph id="6A0F1B17FFC6893CFF266990D13CC047" blockId="8.[151,1438,151,2013]" pageId="8" pageNumber="116">
|
||||
<emphasis id="58C4C705FFC6893CFF266990D1A3C1A2" box="[199,437,1231,1256]" italics="true" pageId="8" pageNumber="116">Hemispermatophore:</emphasis>
|
||||
Male code ICN-As-905 was dissected but no hemispermatophores/paraxial organs were found inside.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="22AA489CFFC6893CFF266848D54DC3D2" pageId="8" pageNumber="116" type="discussion">
|
||||
<paragraph id="6A0F1B17FFC6893CFF266848D111C0D7" blockId="8.[151,1438,151,2013]" pageId="8" pageNumber="116">
|
||||
<emphasis id="58C4C705FFC6893CFF266848D12DC065" bold="true" box="[199,315,1303,1328]" inLineHeading="true" pageId="8" pageNumber="116" reason="1">Remarks.</emphasis>
|
||||
Even though the largest male specimen available closely resembles the size of the females, it does not display any evident secondary sexual characters. Based on this and on the absence of hemispermatophores/ paraxial organs, this specimen is a subadult. The sexual dimorphism in the new species, if present, is therefore unknown.
|
||||
</paragraph>
|
||||
<paragraph id="6A0F1B17FFC6893CFF2668F8D54DC3D2" blockId="8.[151,1438,151,2013]" pageId="8" pageNumber="116">
|
||||
<emphasis id="58C4C705FFC6893CFF2668F8D1BFC08A" bold="true" box="[199,425,1447,1472]" inLineHeading="true" pageId="8" pageNumber="116" reason="1">Field observations.</emphasis>
|
||||
Most of the specimens of the new species herein referred were collected inside the cave; however, this species is also present outside of it. One of the females (ICN-As-872) was collected from the aphotic zone of the cave. This specimen subsequently gave birth to
|
||||
<specimenCount id="7CB6D09EFFC6893CFC8368B0D3F9C342" box="[866,1007,1519,1544]" pageId="8" pageNumber="116" type="juvenile">19 juveniles</specimenCount>
|
||||
in the laboratory. In addition to the specimens documented here from the cave, additional anecdotal accounts (alleged to be) for this species have been made by visitors. Outside of it, individuals have been observed under rotten logs. Based on these observations, it seems likely that
|
||||
<taxonomicName id="ADB06094FFC6893CFEBA6B02D21AC33F" authority="Botero-Trujillo & Flórez, 2014" authorityName="Botero-Trujillo & Flórez" authorityYear="2014" box="[347,524,1628,1653]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="116" phylum="Arthropoda" rank="species" species="grottoedensis" status="sp. nov.">
|
||||
<emphasis id="58C4C705FFC6893CFEBA6B02D21AC33F" box="[347,524,1628,1653]" italics="true" pageId="8" pageNumber="116">T. grottoedensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="58C4C705FFC6893CFDF56B03D278C33F" bold="true" box="[532,622,1628,1653]" pageId="8" pageNumber="116">
|
||||
<taxonomicNameLabel id="43F77A7EFFC6893CFDF56B03D278C33F" box="[532,622,1628,1653]" pageId="8" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
be a eutroglophile or subtroglophile, instead of a trogloxene. Additional studies are needed to determine the degree of association/adaptation of the new species to the cave, however.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="22AA489CFFC6893DFF266BFBD1C1C59F" lastPageId="9" lastPageNumber="117" pageId="8" pageNumber="116" type="distribution">
|
||||
<paragraph id="6A0F1B17FFC6893CFF266BFBD382C207" blockId="8.[151,1438,151,2013]" pageId="8" pageNumber="116">
|
||||
<emphasis id="58C4C705FFC6893CFF266BFBD176C3F7" bold="true" box="[199,352,1700,1725]" inLineHeading="true" pageId="8" pageNumber="116" reason="1">Distribution.</emphasis>
|
||||
Known only from the
|
||||
<typeStatus id="B50BA5B5FFC6893CFD806BFBD287C3F6" box="[609,657,1700,1724]" pageId="8" pageNumber="116">type</typeStatus>
|
||||
locality, El Edén Cave (
|
||||
<figureCitation id="F28B0792FFC6893CFC456BFBD406C3F7" box="[932,1040,1700,1725]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1638,1661]" captionTargetBox="[151,1433,640,1604]" captionTargetId="figure-343@2.[151,1436,625,1617]" captionTargetPageId="2" captionText="FIGURES 1–3. Distribution of Tityus (Tityus) grottoedensis sp. nov. 1. Map showing the location of El Edén Cave in Colombia. 2. Entrance to El Edén Cave from the opposite riverside. 3. Interior passage of El Edén cave. Photographs courtesy of Miguel A. Medrano (Fig. 2) and Iván C. Gonzalez (Fig. 3)." figureDoi="http://doi.org/10.5281/zenodo.4915129" httpUri="https://zenodo.org/record/4915129/files/figure.png" pageId="8" pageNumber="116">Figs. 1–3</figureCitation>
|
||||
). This cave is located at an altitude of
|
||||
<quantity id="AD48B6F2FFC6893CFF576B98D0EBC395" box="[182,253,1735,1760]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="8" pageNumber="116" unit="m" value="650.0">650 m</quantity>
|
||||
in the municipality of Cunday,
|
||||
<collectingRegion id="A874D5F5FFC6893CFD856B98D32AC3AA" box="[612,828,1735,1760]" country="Colombia" name="Tolima" pageId="8" pageNumber="116">Tolima department</collectingRegion>
|
||||
. This municipality is found in the geographical valley of the
|
||||
<collectingRegion id="A874D5F5FFC6893CFF016BB3D14BC24F" box="[224,349,1772,1797]" country="Colombia" name="Magdalena" pageId="8" pageNumber="116">Magdalena</collectingRegion>
|
||||
River, close to the region known as “Middle
|
||||
<collectingRegion id="A874D5F5FFC6893CFC856BB3D3F0C24F" box="[868,998,1772,1797]" country="Colombia" name="Magdalena" pageId="8" pageNumber="116">Magdalena</collectingRegion>
|
||||
”. The cave system has an approximate length of
|
||||
<quantity id="AD48B6F2FFC6893CFEE96A50D144C26D" box="[264,338,1807,1832]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="8" pageNumber="116" unit="m" value="850.0">850 m</quantity>
|
||||
(
|
||||
<bibRefCitation id="0E2166E6FFC6893CFE836A50D216C262" author="Cabrera, W. O." box="[354,512,1807,1832]" pageId="8" pageNumber="116" pagination="1 - 11" refId="ref6207" refString="Cabrera, W. O. (1970) La Cueva de Cunday. Boletin de la Sociedad Geografica de Colombia, 27 (12), 1 - 11." type="journal article" year="1970">Cabrera 1970</bibRefCitation>
|
||||
) and geographical coordinates
|
||||
<geoCoordinate id="0F847DD0FFC6893CFC886A50D3D0C26D" box="[873,966,1807,1832]" degrees="4" direction="north" minutes="00" orientation="latitude" pageId="8" pageNumber="116" precision="925" value="4.0">4°00´N</geoCoordinate>
|
||||
<geoCoordinate id="0F847DD0FFC6893CFC316A50D457C26D" box="[976,1089,1807,1832]" degrees="74" direction="west" minutes="45" orientation="longitude" pageId="8" pageNumber="116" precision="925" value="-74.75">74°45´W</geoCoordinate>
|
||||
. The ecosystem in which it is found is considered as tropical humid forest (
|
||||
<bibRefCitation id="0E2166E6FFC6893CFD706A6BD39EC207" author="Castaneda, E. & Cadena, A. & Aguilera, A. & Sanin, L. H. & Ajello, L. & Weeks, R. & Corredor, V. & Restrepo, M." box="[657,904,1844,1869]" pageId="8" pageNumber="116" pagination="208 - 212" refId="ref6341" refString="Castaneda, E., Cadena, A., Aguilera, A., Sanin, L. H., Ajello, L., Weeks, R., Corredor, V. & Restrepo, M. (1981) Histoplasmosis epidemica. II. Hallazgos en la Cueva del Eden, Cunday, Tolima. Biomedica, 1 (4), 208 - 212." type="journal article" year="1981">
|
||||
Castañeda
|
||||
<emphasis id="58C4C705FFC6893CFCED6A6AD353C207" box="[780,837,1844,1869]" italics="true" pageId="8" pageNumber="116">et al.</emphasis>
|
||||
1981
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="6A0F1B17FFC6893DFF266A08D1C1C59F" blockId="8.[151,1438,151,2013]" lastBlockId="9.[151,1436,151,213]" lastPageId="9" lastPageNumber="117" pageId="8" pageNumber="116">
|
||||
The scorpion fauna of the area where Cunday is embedded has been relatively well investigated, for it has a central position in
|
||||
<collectingCountry id="12A75B87FFC6893CFE896A23D1C1C2DF" box="[360,471,1916,1941]" name="Colombia" pageId="8" pageNumber="116">Colombia</collectingCountry>
|
||||
close to major cities like
|
||||
<collectingRegion id="A874D5F5FFC6893CFD166A23D35EC2DF" box="[759,840,1916,1941]" country="Colombia" name="Bogota" pageId="8" pageNumber="116">Bogotá</collectingRegion>
|
||||
(capital city), Ibagué (capital of
|
||||
<collectingRegion id="A874D5F5FFC6893CFB596A23D587C2DF" box="[1208,1425,1916,1941]" country="Colombia" name="Tolima" pageId="8" pageNumber="116">Tolima department</collectingRegion>
|
||||
), and the municipalities of Melgar and Girardot (both very touristic destinations). The scorpion fauna of the region is relatively well represented in national biological collections (i.e. ICN-UN), and no other specimens of
|
||||
<taxonomicName id="ADB06094FFC6893DFA666A9AD13BC5FA" authority="Botero-Trujillo & Flórez, 2014" authorityName="Botero-Trujillo & Flórez" authorityYear="2014" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" lastPageId="9" lastPageNumber="117" order="Scorpiones" pageId="8" pageNumber="116" phylum="Arthropoda" rank="species" species="grottoedensis" status="sp. nov.">
|
||||
<emphasis id="58C4C705FFC6893DFA666A9AD13BC5FA" italics="true" lastPageId="9" lastPageNumber="117" pageId="8" pageNumber="116">T. grottoedensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="58C4C705FFC7893DFED76DC8D187C5FA" bold="true" box="[310,401,151,176]" pageId="9" pageNumber="117">
|
||||
<taxonomicNameLabel id="43F77A7EFFC7893DFED76DC8D187C5FA" box="[310,401,151,176]" pageId="9" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
have so far been found. Therefore, the new species is probably an endemic element to El Edén Cave and its vicinities.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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Reference in a new issue