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|
||||
<bibRefCitation id="EFB34B5DFFD0FFC9FF5FFD1AFE87FD42" author="Lamarck JB" box="[129,316,723,747]" pageId="12" pageNumber="13" pagination="63 - 91" refId="ref19100" refString="Lamarck JB. Prodrome d'une nouvelle classification des coquilles, comprenant une redaction appropriee des caracteres generiques, et l'etablissement d'un grand nombre de genres nouveaux. Lu a l'Institut national, le 21 frimaire an VII. Memoires de la Societe d'Histoire Naturelle de Paris 1799; 1: 63 - 91." type="journal article" year="1799">Lamarck 1799: 75</bibRefCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3386527FFD0FFC9FF5FFCC4FD67FC8C" box="[129,732,780,805]" pageId="12" pageNumber="13" type="type_taxon">
|
||||
<paragraph id="8B9D36ACFFD0FFC9FF5FFCC4FD67FC8C" blockId="12.[129,732,780,805]" box="[129,732,780,805]" pageId="12" pageNumber="13">
|
||||
<emphasis id="B956EABEFFD0FFC9FF5FFCC4FF47FC8D" box="[129,252,781,804]" italics="true" pageId="12" pageNumber="13">
|
||||
<typeStatus id="5499880EFFD0FFC9FF5FFCC4FF0AFC8D" box="[129,177,781,804]" pageId="12" pageNumber="13">Type</typeStatus>
|
||||
species:
|
||||
</emphasis>
|
||||
<taxonomicName id="4C224D2FFFD0FFC9FEDFFCC4FD8AFC8D" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[257,561,780,805]" class="Gastropoda" family="Lymnaeidae" genus="Helix" kingdom="Animalia" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="stagnalis">
|
||||
<emphasis id="B956EABEFFD0FFC9FEDFFCC4FE2BFC8C" box="[257,400,781,805]" italics="true" pageId="12" pageNumber="13">Helix stagnalis</emphasis>
|
||||
Linnaeus, 1758
|
||||
</taxonomicName>
|
||||
(by monotypy).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3386527FFD0FFC9FF5FFC8EFDA2FB90" pageId="12" pageNumber="13" type="diagnosis">
|
||||
<paragraph id="8B9D36ACFFD0FFC9FF5FFC8EFDA2FB90" blockId="12.[128,777,838,1081]" pageId="12" pageNumber="13">
|
||||
<emphasis id="B956EABEFFD0FFC9FF5FFC8EFF5CFCF7" box="[129,231,839,862]" italics="true" pageId="12" pageNumber="13">Diagnosis:</emphasis>
|
||||
Shell large (≤
|
||||
<quantity id="4CDA9B49FFD0FFC9FE57FC8FFE6FFCF4" box="[393,468,838,862]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="13" unit="mm" value="70.0">70 mm</quantity>
|
||||
height, usually
|
||||
<quantity id="4CDA9B49FFD0FFC9FD54FC8FFD45FCF7" box="[650,766,838,862]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="12" pageNumber="13" unit="mm" value="45.0" valueMax="50.0" valueMin="40.0">40–50 mm</quantity>
|
||||
), ovate-conical to ear-shaped (
|
||||
<figureCitation id="13192A29FFD0FFC9FE71FCACFE49FCD4" box="[431,498,869,893]" captionStart="Figure 3" captionStartId="9.[113,178,1839,1863]" captionTargetBox="[181,1406,146,1795]" captionTargetId="figure-7@9.[160,1412,144,1811]" captionTargetPageId="9" captionText="Figure 3. Shells of the Lymnaeinae species. A, Dallirhytis atkaensis (USA, Alaska, Birch Lake; RMBH). B, Dallirhytis atkaensis (Russia, the Chukchi Peninsula, an unnamed lake; ZIN). C, Galba sibirica (Mongolia, the Teellin-gol River; after Vinarski et al. 2017b). D, Galba truncatula (Russia, Irkutsk Region, vicinities of Kirensk Town; LMBI). E, Ladislavella liogyra (Russia, Amur basin, vicinities of Nikol’sk-Ussuriysky Town; ZIN). F, Walhiana arctica (Canada, Ontario, Hudson’s Bay; NHMUK). G, Walhiana catascopium (Canada, British Columbia, Aintworth hot springs; RMBH). H, Walhiana catascopium (Russia, Kamchatka, Azabachye Lake; RMBH). I, Lymnaea jugularis (USA, Michigan, the Rouge River; NHMUK). J, Lymnaea sorensis (Russia, Lake Baikal, a syntype; ZIN). K, Lymnaea jugularis (USA, Minnesota, a neotype with label; ZIN). Scale bars: 2 mm in C, D, F–H; 5 mm in A, B, E, I–K.Photographs: Maxim V.Vinarski (A–F, I–K) and Olga V.Aksenova (G, H)." pageId="12" pageNumber="13">Fig. 3J</figureCitation>
|
||||
); both shell size and shell proportions are prone to high variability, both within and among populations. Aperture relatively large, with the body whorl moderately to greatly inflated. Prostate with numerous small folds inside. Praeputium is thick and sac-like, its length exceeds the penis sheath length by 2.5–7.0 times (
|
||||
<figureCitation id="13192A29FFD0FFC9FDDAFBCBFDF0FBB3" box="[516,587,1026,1050]" captionStart="Figure 4" captionStartId="10.[130,195,1678,1702]" captionTargetBox="[134,1471,149,1640]" captionTargetId="figure-11@10.[129,1476,145,1650]" captionTargetPageId="10" captionText="Figure 4. The copulatory apparatuses of the pond snails discussed in the taxonomic account. A, Dallirhytis atkaensis (Russia, the Chukchi Peninsula, Lake Vaaliuchio). B, Galba sibirica (Mongolia, the Teellin-gol River; after Vinarski et al. 2017b). C, Galba truncatula (Abkhazia, Gagra District, wet shore of Inkit Lake; after Vinarski et al.2017b). D, Ladislavella liogyra (Russia, Primorye, a pool near Lake Lebedinoye).E, Walhiana catascopium (Russia, Kamchatka, Azabachye Lake; RMBH). F, Lymnaea sorensis (Russia, Kamchatka, Kamchatka River; RMBH). G, Kamtschaticana kamtschatica (Russia, Magadan Region, a water reservoir; after Vinarski et al. 2021). H, Orientogalba ollula (Russia, Primorye Territory, near Possiet Settlement; after Kruglov 2005). I, Radix auricularia (Russia, Tyumen’ Region, Vylposl channel near Labytnangi Town). J, Radix plicatula (China, Bejing, an artificial pond in the former Emperor’s summer palace; after Vinarski et al. 2020). K, Radix cf.hamadai (Japan, Matsumae; after Vinarski et al. 2020). Scale bars: 1 mm.Photographs: Maxim V. Vinarski (A–D, G, I–K) and Olga V. Aksenova (E, F)." pageId="12" pageNumber="13">Fig. 4F</figureCitation>
|
||||
). The latter is very short and narrow. Penial knot is present.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3386527FFD0FFC9FF5FFB98FD7EFACA" pageId="12" pageNumber="13" type="discussion">
|
||||
<paragraph id="8B9D36ACFFD0FFC9FF5FFB98FD7EFACA" blockId="12.[129,778,1105,1379]" pageId="12" pageNumber="13">
|
||||
<emphasis id="B956EABEFFD0FFC9FF5FFB98FEA5FBC0" box="[129,286,1105,1129]" italics="true" pageId="12" pageNumber="13">Species richness:</emphasis>
|
||||
Most authors include a single species, the great pond snail,
|
||||
<taxonomicName id="4C224D2FFFD0FFC9FF2BFBB8FE12FB21" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[245,425,1136,1160]" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="stagnalis">
|
||||
<emphasis id="B956EABEFFD0FFC9FF2BFBB8FE12FB21" box="[245,425,1136,1160]" italics="true" pageId="12" pageNumber="13">Lymnaea stagnalis</emphasis>
|
||||
</taxonomicName>
|
||||
, which is thought to be widely distributed in Eurasia and North America [but see
|
||||
<bibRefCitation id="EFB34B5DFFD0FFC9FD59FB46FEE0FB6F" author="Kruglov ND & Starobogatov YI" pageId="12" pageNumber="13" pagination="21 - 35" refId="ref18822" refString="Kruglov ND, Starobogatov YI. Methods of experimental hybridization and some results of its applications in the taxonomy of Lymnaeidae. Malacological Review 1985 a; 18: 21 - 35." type="journal article" year="1985">Kruglov and Starobogatov (1985a</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EFB34B5DFFD0FFC9FEB5FB66FE24FB6E" author="Kruglov ND & Starobogatov YI" box="[363,415,1199,1223]" pageId="12" pageNumber="13" pagination="65 - 92" refId="ref18930" refString="Kruglov ND, Starobogatov YI. Annotated and illustrated catalogue of species of the family Lymnaeidae (Gastropoda Pulmonata Lymnaeiformes) of Palaearctic and adjacent river drainage areas. Part I. Ruthinica 1993; 3: 65 - 92." type="journal article" year="1993">1993</bibRefCitation>
|
||||
), who split
|
||||
<emphasis id="B956EABEFFD0FFC9FDFBFB66FD79FB6E" box="[549,706,1199,1223]" italics="true" pageId="12" pageNumber="13">
|
||||
<taxonomicName id="4C224D2FFFD0FFC9FDFBFB66FD1BFB6E" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[549,672,1199,1223]" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="stagnalis">Ly. stagnalis</taxonomicName>
|
||||
<taxonomicNameLabel id="A26557C5FFD0FFC9FD77FB66FD79FB6E" box="[681,706,1199,1223]" pageId="12" pageNumber="13" sensu="lato">s.l.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
into a series of morphospecies, five of them living in North America and six in Eurasia]. Our phylogenetic analysis supports the hypothesis that the great pond snail might be a series of morphologically cryptic species, virtually indistinguishable both conchologically and anatomically (
|
||||
<bibRefCitation id="EFB34B5DFFD0FFC9FE3BFA82FD0FFACA" author="Aksenova OV & Bolotov IN & Khrebtova IS" box="[485,692,1355,1379]" pageId="12" pageNumber="13" pagination="67 - 101" refId="ref17294" refString="Aksenova OV, Bolotov IN, Khrebtova IS et al. Phylogeny and taxonomy of the family Lymnaeidae. In: Vinarski MV, Vazquez AA (eds.), The Lymnaeidae. Zoological Monographs, Vol. 7. Cham: Springer, 2023, 67 - 101. https: // doi. org / 10.1007 / 978 - 3 - 031 - 30292 - 3 _ 3" type="book chapter" year="2023">
|
||||
Aksenova
|
||||
<emphasis id="B956EABEFFD0FFC9FD90FA85FDC7FACA" box="[590,636,1355,1379]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
|
||||
2023
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3386527FFD0FFC9FF5FFA4CFEEAF818" pageId="12" pageNumber="13" type="distribution">
|
||||
<paragraph id="8B9D36ACFFD0FFC9FF5FFA4CFEEAF818" blockId="12.[129,778,1413,1969]" pageId="12" pageNumber="13">
|
||||
<emphasis id="B956EABEFFD0FFC9FF5FFA4CFEB9FA34" box="[129,258,1413,1437]" italics="true" pageId="12" pageNumber="13">Distribution:</emphasis>
|
||||
Holarctic; introduced to some tropical regions of the world. Most of the species belonging to this genus occur in Eurasia, whereas North America is probably inhabited by a single species,
|
||||
<taxonomicName id="4C224D2FFFD0FFC9FE9DFA2AFDC6FA52" authority="Say, 1817" authorityName="Say" authorityYear="1817" box="[323,637,1507,1531]" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="jugularis">
|
||||
<emphasis id="B956EABEFFD0FFC9FE9DFA2AFDB8FA52" box="[323,515,1507,1531]" italics="true" pageId="12" pageNumber="13">Lymnaea jugularis</emphasis>
|
||||
Say, 1817
|
||||
</taxonomicName>
|
||||
(=
|
||||
<taxonomicName id="4C224D2FFFD0FFC9FD73FA2AFEFEF9B3" authority="Say, 1821" authorityName="Say" authorityYear="1821" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="appressa">
|
||||
<emphasis id="B956EABEFFD0FFC9FD73FA2AFF6CF9B3" italics="true" pageId="12" pageNumber="13">Lymnaea appressa</emphasis>
|
||||
Say, 1821
|
||||
</taxonomicName>
|
||||
) (
|
||||
<figureCitation id="13192A29FFD0FFC9FE81F9CBFE18F9B3" box="[351,419,1538,1562]" captionStart="Figure 3" captionStartId="9.[113,178,1839,1863]" captionTargetBox="[181,1406,146,1795]" captionTargetId="figure-7@9.[160,1412,144,1811]" captionTargetPageId="9" captionText="Figure 3. Shells of the Lymnaeinae species. A, Dallirhytis atkaensis (USA, Alaska, Birch Lake; RMBH). B, Dallirhytis atkaensis (Russia, the Chukchi Peninsula, an unnamed lake; ZIN). C, Galba sibirica (Mongolia, the Teellin-gol River; after Vinarski et al. 2017b). D, Galba truncatula (Russia, Irkutsk Region, vicinities of Kirensk Town; LMBI). E, Ladislavella liogyra (Russia, Amur basin, vicinities of Nikol’sk-Ussuriysky Town; ZIN). F, Walhiana arctica (Canada, Ontario, Hudson’s Bay; NHMUK). G, Walhiana catascopium (Canada, British Columbia, Aintworth hot springs; RMBH). H, Walhiana catascopium (Russia, Kamchatka, Azabachye Lake; RMBH). I, Lymnaea jugularis (USA, Michigan, the Rouge River; NHMUK). J, Lymnaea sorensis (Russia, Lake Baikal, a syntype; ZIN). K, Lymnaea jugularis (USA, Minnesota, a neotype with label; ZIN). Scale bars: 2 mm in C, D, F–H; 5 mm in A, B, E, I–K.Photographs: Maxim V.Vinarski (A–F, I–K) and Olga V.Aksenova (G, H)." pageId="12" pageNumber="13">Fig. 3I</figureCitation>
|
||||
). In the East Asian region, we accept a single species, which is widely distributed throughout the Altai, East Siberia, the extreme northeast of Asia, West
|
||||
<collectingCountry id="F335763CFFD0FFC9FF5FF9A9FF7AF9D1" box="[129,193,1632,1656]" name="China" pageId="12" pageNumber="13">China</collectingCountry>
|
||||
(
|
||||
<collectingRegion id="49E6F84EFFD0FFC9FF0FF9A9FE8FF9D1" box="[209,308,1632,1656]" country="China" name="Xinjiang Uygur" pageId="12" pageNumber="13">Xinjiang</collectingRegion>
|
||||
), and, probably,
|
||||
<collectingCountry id="F335763CFFD0FFC9FE38F9A9FDF6F9D1" box="[486,589,1632,1656]" name="Mongolia" pageId="12" pageNumber="13">Mongolia</collectingCountry>
|
||||
. The oldest available name for this species is
|
||||
<taxonomicName id="4C224D2FFFD0FFC9FE65F949FF03F91E" authority="W. Dybowski, 1912" authorityName="W. Dybowski" authorityYear="1912" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="sorensis">
|
||||
<emphasis id="B956EABEFFD0FFC9FE65F949FDD0F93E" box="[443,619,1664,1687]" italics="true" pageId="12" pageNumber="13">Lymnaea sorensis</emphasis>
|
||||
W.
|
||||
<bibRefCitation id="EFB34B5DFFD0FFC9FD47F9B6FF03F91E" author="Dybowski B" pageId="12" pageNumber="13" pagination="165 - 218" refId="ref18162" refString="Dybowski B. Bemerkungen und Zusatze zu der Arbeit von Dr. W. Dybowski ' Mollusken der Uferregion des Baikalsees'. Annuaire du Musee Zoologique de l'Academie Imperiale des Sciences de Saint- Petersbourg 1912; 17: 165 - 218." type="journal article" year="1912">Dybowski, 1912</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, based on its
|
||||
<typeStatus id="5499880EFFD0FFC9FE9DF956FEC9F91E" box="[323,370,1695,1719]" pageId="12" pageNumber="13">type</typeStatus>
|
||||
locality of Lake Baikal. The
|
||||
<typeStatus id="5499880EFFD0FFC9FD44F956FD72F91E" box="[666,713,1695,1719]" pageId="12" pageNumber="13">type</typeStatus>
|
||||
series of this species is housed in the
|
||||
<collectionCode id="ED33AE69FFD0FFC9FE63F977FE50F97C" box="[445,491,1726,1749]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="12" pageNumber="13" type="Museum">ZIN</collectionCode>
|
||||
collection (
|
||||
<figureCitation id="13192A29FFD0FFC9FDB6F977FD11F97F" box="[616,682,1726,1750]" captionStart="Figure 3" captionStartId="9.[113,178,1839,1863]" captionTargetBox="[181,1406,146,1795]" captionTargetId="figure-7@9.[160,1412,144,1811]" captionTargetPageId="9" captionText="Figure 3. Shells of the Lymnaeinae species. A, Dallirhytis atkaensis (USA, Alaska, Birch Lake; RMBH). B, Dallirhytis atkaensis (Russia, the Chukchi Peninsula, an unnamed lake; ZIN). C, Galba sibirica (Mongolia, the Teellin-gol River; after Vinarski et al. 2017b). D, Galba truncatula (Russia, Irkutsk Region, vicinities of Kirensk Town; LMBI). E, Ladislavella liogyra (Russia, Amur basin, vicinities of Nikol’sk-Ussuriysky Town; ZIN). F, Walhiana arctica (Canada, Ontario, Hudson’s Bay; NHMUK). G, Walhiana catascopium (Canada, British Columbia, Aintworth hot springs; RMBH). H, Walhiana catascopium (Russia, Kamchatka, Azabachye Lake; RMBH). I, Lymnaea jugularis (USA, Michigan, the Rouge River; NHMUK). J, Lymnaea sorensis (Russia, Lake Baikal, a syntype; ZIN). K, Lymnaea jugularis (USA, Minnesota, a neotype with label; ZIN). Scale bars: 2 mm in C, D, F–H; 5 mm in A, B, E, I–K.Photographs: Maxim V.Vinarski (A–F, I–K) and Olga V.Aksenova (G, H)." pageId="12" pageNumber="13">Fig. 3J</figureCitation>
|
||||
). In addition, we studied numerous topotypic specimens of this species collected in
|
||||
<date id="FF9C106CFFD0FFC9FEF3F934FE23F8BC" box="[301,408,1789,1813]" pageId="12" pageNumber="13" value="2021-06">June 2021</date>
|
||||
(
|
||||
<collectionCode id="ED33AE69FFD0FFC9FE7BF934FE52F8BC" box="[421,489,1789,1813]" pageId="12" pageNumber="13">LMBI</collectionCode>
|
||||
collection). Based on these considerations, we adopt this name for the clade of sequences from Northeast and Central Asia. Conchologically and anatomically, it corresponds fully to the great pond snail, which is common in Europe and West Siberia. There are no similar species in
|
||||
<collectingCountry id="F335763CFFD0FFC9FF35F853FEF6F818" box="[235,333,1945,1969]" name="China" pageId="12" pageNumber="13">East Asia</collectingCountry>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3386527FFD0FFC9FCE7FF59FB86FC7E" pageId="12" pageNumber="13" type="discussion">
|
||||
<paragraph id="8B9D36ACFFD0FFC9FCE7FF59FB86FC7E" blockId="12.[825,1476,144,983]" pageId="12" pageNumber="13">
|
||||
<emphasis id="B956EABEFFD0FFC9FCE7FF59FBA4FF01" box="[825,1055,144,168]" italics="true" pageId="12" pageNumber="13">Nomenclatural remark:</emphasis>
|
||||
<bibRefCitation id="EFB34B5DFFD0FFC9FBF6FF59FB1CFF01" author="Say T" box="[1064,1191,144,168]" pageId="12" pageNumber="13" refId="ref20249" refString="Say T. Conchology. In: Nicholson W (ed.), American Edition of the British Encyclopedia, or, Dictionary of Arts and Sciences Comprising an Accurate and Popular View of the Present Improved State of Human Knowledge; First Edition A- 3 - C- 6. Philadelphia: Mitchell, Ames, White, 1817 a." type="book" year="1817">Say (1817a)</bibRefCitation>
|
||||
described
|
||||
<taxonomicName id="4C224D2FFFD0FFC9FAC4FF58FA28FF01" authorityName="Say" authorityYear="1817" box="[1306,1427,144,168]" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="jugularis">
|
||||
<emphasis id="B956EABEFFD0FFC9FAC4FF58FA28FF01" box="[1306,1427,144,168]" italics="true" pageId="12" pageNumber="13">Ly. jugularis</emphasis>
|
||||
</taxonomicName>
|
||||
as a species of ‘great pond snail’ in North America. Some authors accepted this name, treating
|
||||
<taxonomicName id="4C224D2FFFD0FFC9FBBBFF06FB66FF4F" authorityName="Say" authorityYear="1821" box="[1125,1245,207,230]" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="appressa">
|
||||
<emphasis id="B956EABEFFD0FFC9FBBBFF06FB66FF4F" box="[1125,1245,207,230]" italics="true" pageId="12" pageNumber="13">Ly. appressa</emphasis>
|
||||
</taxonomicName>
|
||||
as its junior synonym (e.g.
|
||||
<bibRefCitation id="EFB34B5DFFD0FFC9FCB7FF27FB11FEAF" author="Kruglov ND & Starobogatov YI" box="[873,1194,238,262]" pageId="12" pageNumber="13" pagination="65 - 92" refId="ref18930" refString="Kruglov ND, Starobogatov YI. Annotated and illustrated catalogue of species of the family Lymnaeidae (Gastropoda Pulmonata Lymnaeiformes) of Palaearctic and adjacent river drainage areas. Part I. Ruthinica 1993; 3: 65 - 92." type="journal article" year="1993">Kruglov and Starobogatov 1993</bibRefCitation>
|
||||
). In contrast,
|
||||
<bibRefCitation id="EFB34B5DFFD0FFC9FAEBFF27FA7AFEAF" author="Burch JB" box="[1333,1473,238,262]" pageId="12" pageNumber="13" refId="ref17812" refString="Burch JB. North American Freshwater Snails. Hamburg, MI: Malacological Publications, 1989." type="book" year="1989">Burch (1989)</bibRefCitation>
|
||||
explicitly ignored
|
||||
<taxonomicName id="4C224D2FFFD0FFC9FC31FEC7FBDEFE8C" authorityName="Say" authorityYear="1817" box="[1007,1125,269,293]" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="jugularis">
|
||||
<emphasis id="B956EABEFFD0FFC9FC31FEC7FBDEFE8C" box="[1007,1125,269,293]" italics="true" pageId="12" pageNumber="13">Ly. jugularis</emphasis>
|
||||
</taxonomicName>
|
||||
as an available name, without providing an explanation.
|
||||
<bibRefCitation id="EFB34B5DFFD0FFC9FBEDFEE4FA80FEEC" author="Clarke AH" box="[1075,1339,301,325]" pageId="12" pageNumber="13" pagination="1 - 509" refId="ref17971" refString="Clarke AH. The freshwater molluscs of the Canadian Interior Basin. Malacologia 1973; 13: 1 - 509." type="journal article" year="1973">Clarke (1973: 300–301)</bibRefCitation>
|
||||
argued that
|
||||
<taxonomicName id="4C224D2FFFD0FFC9FCE7FE84FC08FECD" authorityName="Say" authorityYear="1817" box="[825,947,332,356]" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="jugularis">
|
||||
<emphasis id="B956EABEFFD0FFC9FCE7FE84FC08FECD" box="[825,947,332,356]" italics="true" pageId="12" pageNumber="13">Ly. jugularis</emphasis>
|
||||
</taxonomicName>
|
||||
is unrecognizable because it was based on a very short and incomplete description that was not accompanied by a figure. He characterized this species as ‘a doubtful inhabitant of the United States’.
|
||||
<bibRefCitation id="EFB34B5DFFD0FFC9FB87FE63FB4AFE6B" author="Clarke AH" box="[1113,1265,426,450]" pageId="12" pageNumber="13" pagination="1 - 509" refId="ref17971" refString="Clarke AH. The freshwater molluscs of the Canadian Interior Basin. Malacologia 1973; 13: 1 - 509." type="journal article" year="1973">Clarke (1973)</bibRefCitation>
|
||||
decided to use the name
|
||||
<taxonomicName id="4C224D2FFFD0FFC9FCA7FE03FC4BFE48" authorityName="Say" authorityYear="1821" box="[889,1008,458,481]" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="appressa">
|
||||
<emphasis id="B956EABEFFD0FFC9FCA7FE03FC4BFE48" box="[889,1008,458,481]" italics="true" pageId="12" pageNumber="13">Ly. appressa</emphasis>
|
||||
</taxonomicName>
|
||||
for this species. Later,
|
||||
<bibRefCitation id="EFB34B5DFFD0FFC9FB3EFE00FACEFE48" author="Clarke AH" box="[1248,1397,457,481]" pageId="12" pageNumber="13" refId="ref17992" refString="Clarke AH. The Freshwater Molluscs of Canada. Ottawa: National Museum of Natural Sciences, National Museums of Canada, 1981." type="book" year="1981">Clarke (1981)</bibRefCitation>
|
||||
opined that ‘Say’s first (1817) description of
|
||||
<taxonomicName id="4C224D2FFFD0FFC9FB18FE20FAA1FDA8" authorityName="Say" authorityYear="1817" box="[1222,1306,489,513]" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="jugularis">
|
||||
<emphasis id="B956EABEFFD0FFC9FB18FE20FAA1FDA8" box="[1222,1306,489,513]" italics="true" pageId="12" pageNumber="13">jugularis</emphasis>
|
||||
</taxonomicName>
|
||||
is recognizable’ and that
|
||||
<taxonomicName id="4C224D2FFFD0FFC9FC49FDC0FC50FD89" authorityName="Say" authorityYear="1821" box="[919,1003,521,544]" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="appressa">
|
||||
<emphasis id="B956EABEFFD0FFC9FC49FDC0FC50FD89" box="[919,1003,521,544]" italics="true" pageId="12" pageNumber="13">appressa</emphasis>
|
||||
</taxonomicName>
|
||||
was its junior synonym. The type specimens of
|
||||
<taxonomicName id="4C224D2FFFD0FFC9FC88FDE1FC6BFD96" authorityName="Say" authorityYear="1817" box="[854,976,551,575]" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="jugularis">
|
||||
<emphasis id="B956EABEFFD0FFC9FC88FDE1FC6BFD96" box="[854,976,551,575]" italics="true" pageId="12" pageNumber="13">Ly. jugularis</emphasis>
|
||||
</taxonomicName>
|
||||
are lost, which might prevent researchers from reaching an agreement on the identity of this name. In contrast, the type series of
|
||||
<taxonomicName id="4C224D2FFFD0FFC9FC28FDAEFBD4FDD7" authorityName="Say" authorityYear="1821" box="[1014,1135,615,638]" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="appressa">
|
||||
<emphasis id="B956EABEFFD0FFC9FC28FDAEFBD4FDD7" box="[1014,1135,615,638]" italics="true" pageId="12" pageNumber="13">Ly. appressa</emphasis>
|
||||
</taxonomicName>
|
||||
is extant (see
|
||||
<bibRefCitation id="EFB34B5DFFD0FFC9FAD4FDAFFA01FDD7" author="Baker FC" box="[1290,1466,614,638]" pageId="12" pageNumber="13" pagination="1 - 539" refId="ref17618" refString="Baker FC. The Lymnaeidae of North and Middle America. Special Publications of the Chicago Academy of Sciences 1911; 3: 1 - 539." type="journal article" year="1911">Baker 1911: 141</bibRefCitation>
|
||||
) and available to identify this species. To resolve this confusing situation, we here designate a
|
||||
<typeStatus id="5499880EFFD0FFC9FBB8FD6CFB01FD14" box="[1126,1210,677,701]" pageId="12" pageNumber="13" type="neotype">neotype</typeStatus>
|
||||
for
|
||||
<taxonomicName id="4C224D2FFFD0FFC9FB3CFD6CFCD4FD75" authority="Say, 1817" authorityName="Say" authorityYear="1817" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="jugularis">
|
||||
<emphasis id="B956EABEFFD0FFC9FB3CFD6CFA2EFD14" box="[1250,1429,677,701]" italics="true" pageId="12" pageNumber="13">Lymnaea jugularis</emphasis>
|
||||
Say, 1817
|
||||
</taxonomicName>
|
||||
. This specimen was collected from an unknown locality in
|
||||
<collectingRegion id="49E6F84EFFD0FFC9FCE4FD2AFC11FD52" box="[826,938,739,763]" country="United States of America" name="Minnesota" pageId="12" pageNumber="13">Minnesota</collectingRegion>
|
||||
, North America, in the mid-19th century and is kept in
|
||||
<collectionCode id="ED33AE69FFD0FFC9FC8BFCCAFC39FCB3" box="[853,898,771,794]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="12" pageNumber="13" type="Museum">ZIN</collectionCode>
|
||||
(no.
|
||||
<quantity id="4CDA9B49FFD0FFC9FC64FCCAFC59FCB2" box="[954,994,771,795]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="12" pageNumber="13" unit="in" value="1.0">1 in</quantity>
|
||||
the systematic catalogue). The exact sampling date is unknown; we know only that this sample was obtained from the
|
||||
<collectingCountry id="F335763CFFD0FFC9FC46FC88FB9BFCF0" box="[920,1056,833,857]" name="United States of America" pageId="12" pageNumber="13">United States</collectingCountry>
|
||||
National Museum in the end of the 19th century (
|
||||
<figureCitation id="13192A29FFD0FFC9FC42FCA8FC51FCD0" box="[924,1002,865,889]" captionStart="Figure 3" captionStartId="9.[113,178,1839,1863]" captionTargetBox="[181,1406,146,1795]" captionTargetId="figure-7@9.[160,1412,144,1811]" captionTargetPageId="9" captionText="Figure 3. Shells of the Lymnaeinae species. A, Dallirhytis atkaensis (USA, Alaska, Birch Lake; RMBH). B, Dallirhytis atkaensis (Russia, the Chukchi Peninsula, an unnamed lake; ZIN). C, Galba sibirica (Mongolia, the Teellin-gol River; after Vinarski et al. 2017b). D, Galba truncatula (Russia, Irkutsk Region, vicinities of Kirensk Town; LMBI). E, Ladislavella liogyra (Russia, Amur basin, vicinities of Nikol’sk-Ussuriysky Town; ZIN). F, Walhiana arctica (Canada, Ontario, Hudson’s Bay; NHMUK). G, Walhiana catascopium (Canada, British Columbia, Aintworth hot springs; RMBH). H, Walhiana catascopium (Russia, Kamchatka, Azabachye Lake; RMBH). I, Lymnaea jugularis (USA, Michigan, the Rouge River; NHMUK). J, Lymnaea sorensis (Russia, Lake Baikal, a syntype; ZIN). K, Lymnaea jugularis (USA, Minnesota, a neotype with label; ZIN). Scale bars: 2 mm in C, D, F–H; 5 mm in A, B, E, I–K.Photographs: Maxim V.Vinarski (A–F, I–K) and Olga V.Aksenova (G, H)." pageId="12" pageNumber="13">Fig. 3K</figureCitation>
|
||||
). This specimen shows all characters typical for the Nearctic
|
||||
<emphasis id="B956EABEFFD0FFC9FC3AFC48FB02FC31" box="[996,1209,896,920]" italics="true" pageId="12" pageNumber="13">
|
||||
<taxonomicName id="4C224D2FFFD0FFC9FC3AFC48FB21FC31" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[996,1178,896,920]" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" kingdom="Animalia" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" sensu="lato" species="stagnalis">Lymnaea stagnalis</taxonomicName>
|
||||
<taxonomicNameLabel id="A26557C5FFD0FFC9FB7EFC49FB02FC31" box="[1184,1209,896,920]" pageId="12" pageNumber="13" sensu="lato">s.l.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
as they were described in papers by
|
||||
<bibRefCitation id="EFB34B5DFFD0FFC9FC78FC56FB8BFC1E" author="Baker FC" box="[934,1072,927,951]" pageId="12" pageNumber="13" pagination="1 - 539" refId="ref17618" refString="Baker FC. The Lymnaeidae of North and Middle America. Special Publications of the Chicago Academy of Sciences 1911; 3: 1 - 539." type="journal article" year="1911">Baker (1911)</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EFB34B5DFFD0FFC9FBE3FC56FB7DFC1E" author="Clarke AH" box="[1085,1222,927,951]" pageId="12" pageNumber="13" pagination="1 - 509" refId="ref17971" refString="Clarke AH. The freshwater molluscs of the Canadian Interior Basin. Malacologia 1973; 13: 1 - 509." type="journal article" year="1973">Clarke (1973</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EFB34B5DFFD0FFC9FB0DFC56FABCFC1E" author="Clarke AH" box="[1235,1287,927,951]" pageId="12" pageNumber="13" refId="ref17992" refString="Clarke AH. The Freshwater Molluscs of Canada. Ottawa: National Museum of Natural Sciences, National Museums of Canada, 1981." type="book" year="1981">1981</bibRefCitation>
|
||||
), and some other North American authors.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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Reference in a new issue