diff --git a/data/03/A1/0B/03A10B47FFE8FF95FD03F978FD08FBAD.xml b/data/03/A1/0B/03A10B47FFE8FF95FD03F978FD08FBAD.xml index d1cd8346574..3193ce9b5f2 100644 --- a/data/03/A1/0B/03A10B47FFE8FF95FD03F978FD08FBAD.xml +++ b/data/03/A1/0B/03A10B47FFE8FF95FD03F978FD08FBAD.xml @@ -1,40 +1,41 @@ - - - -A mountain of millipedes VII: The genus Eviulisoma Silvestri, 1910, in the Udzungwa Mountains, Tanzania, and related species from other Eastern Arc Mountains. With notes on Eoseviulisoma Brolemann, 1920, and Suohelisoma Hoffman, 1963 (Diplopoda, Polydesmida, Paradoxosomatidae) + + + +A mountain of millipedes VII: The genus Eviulisoma Silvestri, 1910, in the Udzungwa Mountains, Tanzania, and related species from other Eastern Arc Mountains. With notes on Eoseviulisoma Brolemann, 1920, and Suohelisoma Hoffman, 1963 (Diplopoda, Polydesmida, Paradoxosomatidae) - - -Author + + +Author -Enghoff, Henrik +Enghoff, Henrik -text - - -European Journal of Taxonomy +text + + +European Journal of Taxonomy - -2018 - -2018-06-19 + +2018 + +2018-06-19 - -445 + +445 - -1 -90 + +1 +90 -journal article -10.5852/ejt.2018.445 -5dfd4ff4-6c8b-47f4-9aa7-0f6b1b2af967 -1489598 -852A3F68-B728-413A-B12E-56F306D56C35 +journal article +10.5852/ejt.2018.445 +5dfd4ff4-6c8b-47f4-9aa7-0f6b1b2af967 +2118-9773 +1489598 +852A3F68-B728-413A-B12E-56F306D56C35 - + @@ -48,7 +49,7 @@ group - + Diagnosis @@ -490,9 +491,9 @@ The included species furthermore share a hypoproct with three prominent marginal sp. nov. , which is also larger than the others, having a distinct ringed colour pattern. -Included species: - + +Included species: E. @@ -506,8 +507,6 @@ The included species furthermore share a hypoproct with three prominent marginal sp. nov. - - E. aequilobatum @@ -574,8 +573,6 @@ The included species furthermore share a hypoproct with three prominent marginal sp. nov. - - E. alluaudi diff --git a/data/03/A1/0B/03A10B47FFFEFF80FDB7FE71FB7AFA5B.xml b/data/03/A1/0B/03A10B47FFFEFF80FDB7FE71FB7AFA5B.xml new file mode 100644 index 00000000000..66fdd1ce3a2 --- /dev/null +++ b/data/03/A1/0B/03A10B47FFFEFF80FDB7FE71FB7AFA5B.xml @@ -0,0 +1,185 @@ + + + +A mountain of millipedes VII: The genus Eviulisoma Silvestri, 1910, in the Udzungwa Mountains, Tanzania, and related species from other Eastern Arc Mountains. With notes on Eoseviulisoma Brolemann, 1920, and Suohelisoma Hoffman, 1963 (Diplopoda, Polydesmida, Paradoxosomatidae) + + + +Author + +Enghoff, Henrik + +text + + +European Journal of Taxonomy + + +2018 + +2018-06-19 + + +445 + + +1 +90 + + + +journal article +10.5852/ejt.2018.445 +5dfd4ff4-6c8b-47f4-9aa7-0f6b1b2af967 +2118-9773 +1489598 +852A3F68-B728-413A-B12E-56F306D56C35 + + + + +Tribe + +Eviulisomatini +Brölemann, 1916 + + + + + + +Diagnosis + + + +Paradoxosomatidae +in which paranota are very strongly reduced or virtually absent, and the ʻfemoriteʼ of the gonopod is strongly reduced, such that solenomere, solenophore and up to two acropodital processes seem to originate directly from the end of the prefemoral part. + + + + + +Eviulisoma + +is classified in the tribe +Eviulisomatini +which, according to +Nguyen & Sierwald (2013) +and +Reboleira & Enghoff (2013) +, also contains + +Boreviulisoma +Brolemann, 1928 + +(SW Palaearctic: NW Africa, SW Europe), + +Eoseviulisoma +Brolemann, 1920 +(Afrotropical) + +, + +Onciurosoma +Silvestri, 1932 (Neotropical) + +, + +Scolodesmus +Cook, 1896 (Afrotropical) + +, + +Suohelisoma +Hoffman, 1964 +(Afrotropical) + +and + +Wubidesmus +Chamberlin, 1927 (Afrotropical) + +. To these should be added + +Jeekelosoma +Mauriès, 1985 + +, raised to full generic status from subgenus under + +Eviulisoma + +by Enghoff & Reboleira (in prep.), and possibly + +Stirosoma +Attems, 1953 +(Afrotropical) + +. See below concerning + +Eoseviulisoma + +and + +Suohelisoma + +. In the key to +Central +African eviulisomatinine genera by +Hoffman (1971) +, + +Eviulisoma + +keys out together with + +Suohelisoma + +in couplet 1 because of the ʻdorsalʼ (actually cranio-distal) lobe on the mesal side of the gonopod coxa. These two nominal genera also share the strongly excavated sternum 6 (absent in a few species of + +Eviulisoma + +, however), a character also present in + +Wubidesmus +. + +Variously excavated sterna 6 have been described for certain other paradoxosomatids, e.g., + +Luzonomorpha pallidula +Jeekel, 2000 + +, and + +Montesecaria golovatchi +Jeekel, 2002 + +( +Jeekel 2000 +, +2002 +), but as far as can be deduced from the descriptions, these excavations are quite different from those seen in + +Eviulisoma + +spp. Most eviulisomatinine genera share a strongly reduced body ring setation: two rows of very thin setae on the collum and only one row on each of the following body rings. The majority of paradoxosomatid genera have two or three rows of setae on postcollar body rings, and this character state is shared by the eviulisomatinine genera + +Boreviulisoma +( +Reboleira & Enghoff 2013 +) + +and + +Jeekelosoma + +( +Mauriès 1985; Enghoff & Reboleira in prep. +). + + +Table 1 +summarizes some important characters across the eviulisomatinine genera. + + + + \ No newline at end of file diff --git a/data/03/A1/0B/03A10B47FFFEFF96FDD6F99BFCC4F972.xml b/data/03/A1/0B/03A10B47FFFEFF96FDD6F99BFCC4F972.xml index 72cf21e4509..3f8108cce5c 100644 --- a/data/03/A1/0B/03A10B47FFFEFF96FDD6F99BFCC4F972.xml +++ b/data/03/A1/0B/03A10B47FFFEFF96FDD6F99BFCC4F972.xml @@ -1,40 +1,41 @@ - - - -A mountain of millipedes VII: The genus Eviulisoma Silvestri, 1910, in the Udzungwa Mountains, Tanzania, and related species from other Eastern Arc Mountains. With notes on Eoseviulisoma Brolemann, 1920, and Suohelisoma Hoffman, 1963 (Diplopoda, Polydesmida, Paradoxosomatidae) + + + +A mountain of millipedes VII: The genus Eviulisoma Silvestri, 1910, in the Udzungwa Mountains, Tanzania, and related species from other Eastern Arc Mountains. With notes on Eoseviulisoma Brolemann, 1920, and Suohelisoma Hoffman, 1963 (Diplopoda, Polydesmida, Paradoxosomatidae) - - -Author + + +Author -Enghoff, Henrik +Enghoff, Henrik -text - - -European Journal of Taxonomy +text + + +European Journal of Taxonomy - -2018 - -2018-06-19 + +2018 + +2018-06-19 - -445 + +445 - -1 -90 + +1 +90 -journal article -10.5852/ejt.2018.445 -5dfd4ff4-6c8b-47f4-9aa7-0f6b1b2af967 -1489598 -852A3F68-B728-413A-B12E-56F306D56C35 +journal article +10.5852/ejt.2018.445 +5dfd4ff4-6c8b-47f4-9aa7-0f6b1b2af967 +2118-9773 +1489598 +852A3F68-B728-413A-B12E-56F306D56C35 - + @@ -46,9 +47,9 @@ Genus - - + + Eviulisoma @@ -56,6 +57,10 @@ Genus : 463 + + + + (type species: Iulidesmus cavallii @@ -65,10 +70,11 @@ Genus , by original designation). - + + + - Eoseviulisoma @@ -93,7 +99,6 @@ species: , by original designation). - @@ -144,18 +149,17 @@ species: - + Included species - - See Table 2. + Table 1. @@ -1077,6 +1081,7 @@ Attems, 1937 + Mauriès (1985) described a subgenus of @@ -1097,7 +1102,7 @@ described a subgenus of will again become an endemic Afrotropical genus. - + Diagnosis @@ -1128,7 +1133,7 @@ may be diagnosed as follows: A genus of -Paradoxosomatidae +Paradoxosomatidae in which: - paranota are missing, or at most present as tiny keels on ring 2 only @@ -1147,6 +1152,8 @@ in which: Gonopod terminology + + Jeekel (2003) gave detailed redescriptions of many species of @@ -1351,10 +1358,8 @@ spp. - Anal valves (paraprocts) with margin raised as narrow lips. - Subanal scale (hypoproct) variable, sometimes semicircular, sometimes trapezoid, sometimes with three more or less conspicuous tubercles at distal margin. -- Legs 0.8–1.5 × as long as body diameter. Relative lengths of podomeres variable; femur the longest in -most cases, but in some short-legged species, prefemur is as long as femur, and in a few long-legged -ones, tarsus is as long as or longer than femur. - +- Legs 0.8–1.5 × as long as body diameter. Relative lengths of podomeres variable; femur the longest in most cases, but in some short-legged species, prefemur is as long as femur, and in a few long-legged ones, tarsus is as long as or longer than femur. + - Non-gonopodal sexual characters: - Sternum 5 usually with a tongue-shaped /subtriangular / subrectangular process ( Fig. 6 ) between anterior legs (pair 4). One species ( @@ -1364,7 +1369,14 @@ spp. sp. nov. ) without a process ( Fig. 36 -) and one +) and one ( + +E. biquintum + +sp. nov. +) with a small tubercle between the 4th leg pair and a similar one between the 5th ( +Fig. 35 +). @@ -1402,27 +1414,17 @@ sternal cone; = terminal tubercle. Scale bars: 0.1 mm. - -( - -E. biquintum - -sp. nov. -) with a small tubercle between the 4th leg pair and a similar one between the 5th ( -Fig. 35 -). - - Sternum 6 usually deeply excavated ( Fig. 6 ), anterior margin of excavation curved, with a row of long setae. Fig. 6 shows a sternum 6 excavation as it appears in most species; deviating morphologies occur in certain species ( -Figs 26 +Figs 26 , -28 +28 , -30 +30 , 34 ). @@ -1524,15 +1526,10 @@ spp., scopulae on postfemur ( , paratype from Mito Mitatu. Scale bars: A, C, E = 0.05 mm; B, F = 0.01 mm; D = 0.1 mm. -- Scopulae (dense coverings of modified setae) usually present on the ventral side of femur, postfemur, - -tibia and tarsus on anterior legs ( + +- Scopulae (dense coverings of modified setae) usually present on the ventral side of femur, postfemur, tibia and tarsus on anterior legs ( Fig. 7 -), decreasing in size from anterior to posterior, often disappearing - -from (femur and) postfemur on posteriormost legs (scopulae only on tibia and tarsus in some non- - -Udzungwan species). +), decreasing in size from anterior to posterior, often disappearing from (femur and) postfemur on posteriormost legs (scopulae only on tibia and tarsus in some non-Udzungwan species). E. kangense @@ -1542,10 +1539,8 @@ and E. sternale sp. nov. -furthermore with the ventral surfaces +furthermore with the ventral surfaces of prefemora and femora flattened and hairless on some leg pairs. No podomeres with swellings or processes. -of prefemora and femora flattened and hairless on some leg pairs. No podomeres with swellings or -processes. - Gonopods ( Fig. 8 @@ -1580,7 +1575,7 @@ Fig. 8 ) originating on lateral side of gonopod, very variable in shape, sometimes a simple rolled sheet, sometimes with a distinct ʻconductorʼ process guiding the solenomere; in species with an ‘open’ solenophore, an area with parallel ridges (e.g., Figs 12–13 , -18 +18 ) is sometimes seen – whether similar ridges are present in rolled-up solenophores is unknown. diff --git a/data/42/B5/14/42B51489FA575CB497D653CBD8BBF811.xml b/data/42/B5/14/42B51489FA575CB497D653CBD8BBF811.xml new file mode 100644 index 00000000000..d3e7824cd7d --- /dev/null +++ b/data/42/B5/14/42B51489FA575CB497D653CBD8BBF811.xml @@ -0,0 +1,472 @@ + + + +Revision of the millipede family Dalodesmidae in Madagascar, with descriptions of two new Malagasy species of Dalodesmus Cook, 1896 (Diplopoda, Polydesmida) + + + +Author + +Wesener, Thomas +0000-0002-2028-3541 +Zoological Research Museum Alexander Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, D- 53113 Bonn, Germany + + + +Author + +Akkari, Nesrine +0000-0001-5019-4833 +Third Zoological Department, Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria + + + +Author + +Golovatch, Sergei I. +Institute of Ecology and Evolution, Russian Academy of Sciences, Leninsky prospekt 33, Moscow 119071, Russia + +text + + +ZooKeys + + +2025 + +2025-01-08 + + +1223 + + +185 +220 + + + +journal article +10.3897/zookeys.1223.139346 +451749E9-009E-43E9-A6F1-892035BDF1B0 + + + + + +Dalodesmus speophilus + +sp. nov. + + + + +Figs 15 +, +16 +, +17 +, +18 + + + + +Type material. + + + + +Holotype + +• +♂ +( + +MZUF + +), +Madagascar +, + +Grotta +di Anjohibe + +, + +15 ° 32 ' 33.08 " S +, +46 ° 53 ' 5.99 " E + +, + +12. ix. 1989 + +, +L. Bartolozzi +& +S. Taiti +leg. + +; + +Paratypes + +: +6 ♂ +, +12 ♀ +, +12 juv. +( + +MZUF + +); 1 M, 1 F ( + +ZFMK + +), same data as +holotype +. + + + + +Diagnosis. + + +Length < +20 mm +in both sexes, the smallest species of the genus (all other species with males> +21 mm +). Tips of paraterga not projecting past posterior tergal margin, like in + +D. hova + +, + +D. tectus + +, + +D. kompantsevi + +sp. nov. +, vs sharper and projecting beyond margin in + +D. hamatus + +, + +D. odontopezus + +and + +D. orator + +. Differs from + +D. hova + +, + +D. tectus + +, and + +D. kompantsevi + +sp. nov. +in the contrasting yellow paratergal tips (uniformly brown in the other species). Gonopods unique for the genus + +Dalodesmus + +in the main branch being subdivided into an apical and a subapical branch. See also Key below. + + + + +Etymology. + + +From +speophilus +, a noun in apposition, after the +type +locality, a gigantic cave. + + + + +Description. + + +Length in male ~ +16.5–17.5 mm +( +n += 3), width of midbody pro- and metazona 1.5 and +2.8–2.9 mm +( +n += 3), respectively, female length +16.5–19 mm +( +n += 4), width of prozona +1.6 mm +( +n += 1), of metazona +3.7–4.1 mm +( +n += 4). + + +Colouration in alcohol light brown to beige, paraterga light, head brown, epicranium grey, legs light, except for a few tarsi pale grey; antennae dark brown (Fig. +15 A – F +). A live photograph of a potential + +D. speophilus + +sp. nov. +, +70 km +away from the +type +locality, shows dull, dark brown to blackish tergites with pale white to almost yellow paratergal tips, legs white to pale grey (Fig. +16 D +). + + + + + + + +Dalodesmus speophilus + +sp. nov. +, ♂ holotype ( +MZUF +), multi-layer photographs +A +habitus, dorsal view +B +midbody rings, dorsal view +C +posterior body rings, dorsal view +D +head, ventral view +E +posterior body rings with telson, dorsal view +F +telson, ventral view. Not to scale. + + + + + + + + +Dalodesmus speophilus + +sp. nov. +, +A – C, E +♂ holotype ( +MZUF +), multi-layer photographs +D +photograph of live specimen of + +Dalodesmus +cf. +speophilus + +, courtesy Justin Gerlach. +A +Anterior part of body with gonopods, ventral view +B +gonopods, posterior view +C +gonopods, lateral view +D +live specimen +E +habitus, lateral view. Not to scale. + + + +Body with 20 rings. Tegument mainly dull, microgranulate to microtuberculate throughout, even surfaces of prozona and of metazona below paraterga finely microgranulate, sterna granulate. Head also densely microtuberculate or granulate throughout, micropilose up to level of antennae; epicranial suture thin, but distinct; genae squarish, set off ventrally from gnathochilarial stipes by a small, but evident ridge (Figs +15 D +, +16 A +). Interantennal isthmus ~ 2 × diameter of antennal socket (Fig. +15 D +). Antennae short and rather clavate, in situ reaching in both sexes back past ring 3 when stretched dorsally, very densely setose and microgranulate. In length, antennomere 6> 2> 5> 4> 2> 1 = 7; antennomere 6 the largest and the highest, antennomeres 5 and 6 each with a small, round, distodorsal knob, most likely beset with sensory cones. In width, collum ≤ head <ring 3 <2 <4–16; thereafter body gradually tapering towards telson (Fig. +15 A +). Collum transversely suboval, regularly and broadly rounded laterally, densely tuberculate, most tuberculations being slightly oblong-oval, evident, equipped with very short, mostly subclavate setae and arranged in 20–22 lateral, 7–8 transverse, rather irregular, arcuated rows. Metaterga 2 and 3 narrow, each with four similar, transverse, arcuated rows of setigerous tubercles (Fig. +17 C +), following metaterga each largely with 5–6 such rows (Fig. +15 A +). Paraterga well-developed, set high (mostly at upper ¼ body), largely slightly upturned to subhorizontal, thus leaving the dorsum only faintly convex (Fig. +15 A – C +); anterior and posterior margins of paraterga 2 and 3 clearly drawn both forward and caudad, following paraterga drawn increasingly only caudad, but caudal corners produced past rear tergal margin only on rings 15–19; posterior margins clearly bisinuate, well concave behind paraterga (Fig. +15 E +). Lateral and caudal margins of paraterga beset with numerous, similarly oblong and usually subequal, setigerous tubercles / lobulations, caudolateral lobulation being nearly sharp (Fig. +17 A, B +). Ozopores inconspicuous (Fig. +17 E, F +), opening dorsally near penultimate lateral lobulation on pore-baring rings 5, 7, 9, 10, 12, 13, 15–19. Strictures between pro- and metazona narrow and rather deep, nearly smooth (Fig. +17 A +). + + + + + + + +Dalodesmus speophilus + +sp. nov. +, ♂ holotype ( +MZUF +), ♀ paratype ( +ZFMK +), +SEM +micrographs +A +midbody ring with pro- and metazonite, dorsal view +B +surface of metazonite +C +detail of surface structures of metazonite +D +endotergum +E +paratergum with ozopore +F +ozopore +G +gonopod, apical view +H +female vulva; +I +endotergum, ventral view +J +male leg pair 1, posterior view +K +tarsus 1 +L +midbody legs. Scale bars: 200 mm ( +A, L +); 100 µm ( +B, E, G, H, J, K +); 10 µm ( +C; D, I +); 20 µm ( +F +). + + + +Telson: Epiproct small, conical and subtruncate at tip. Hypoproct trapeziform, with 1 + 1 setae borne on distinct oblong knobs at caudal margin. Paraprocts with 2 + 2 setae on triangular, projecting knobs (Figs +15 E, F +, +16 E +). + + +Limbus very thin, small, and entire. Neither an axial line nor pleurosternal carinae (Fig. +15 A – C +). Endotergum inconspicuous, posterior margin of metazona projecting into dense, long sharp, apically microdenticulate, triangular projections (Fig. +15 D, I +). + + +Gonopodal aperture roundly pentagonal, relatively small, taking up ~ 1 / 2 width of metazonum 7, clearly open and drawn into metazonum 6 (Fig. +16 A +). + + +First +♂ +leg-pair shorter and wider than other legs, with long coxae, large; size of its podomeres: tarsus> coxa> prefemur = femur> postfemur = tibia (Fig. +17 J +), tarsus with sharp claw, ventral spines, and numerous long setae (Fig. +17 K +). Midbody legs incrassate, rather long. 1.4–1.5 × as long as body height (Fig. +17 L +), with small, stout, abundant, and usually curved setae with admixture of sphaerotrichomes ventrally on all podomeres ( +♂ +); gonopores on +♂ +coxae 2 inconspicuous, each borne on a very small swelling ( +♂ +); prefemora not bulged laterally; claws very small; in length, tarsus> femur> prefemur> tibia> postfemur> coxa. + + +♀ +vulva setose, symmetrical, lateral and inner plates of same width. Operculum large, each side with three or four very long setae, longest reaching the apical margin of vulva (Fig. +17 H +). + + +Gonopods (Figs +16 B, C +, +17 G +, +18 A – L +) very slender and long, tips in situ reaching anteriorly until coxae 5 (Fig. +16 A, B +). Both coxites and prefemoral (= densely setose parts) of telopodites equally very short and stout, fused medially, the former fully and the latter mostly hidden inside gonopodal aperture. Femorites (fe) contiguous medially in basal 1 / 3, sparsely setose almost all along, both slightly flattened dorsoventrally and diverging distad towards acropodites. Apical portions of telopodite (= acropodites) clearly diverging, rather complex and compact, curved ventrad and clearly divided into four unequal branches: a short, submesal, tubiform, simple and non-sigmoid solenomere (sl) flanked by a rather elaborate, tri-branched solenophore, this latter being represented by an immediately adjacent, short, lobe-shaped, lateral branch (lb), a large, subacuminate, slender, twisted, and membranous medial branch (mb) with a denticle near base, and an apical (ab) and a subapical branch (sb), both latter membranous, curved / bent ventrad and ragged / irregular in shape. + + + + + + + +Dalodesmus speophilus + +sp. nov. +, ♂ holotype ( +MZUF +), +SEM +micrographs, right gonopod +A +anterior view +B +postero-mesal view +C +mesal view +D +posterior view +E +lateral view +F +lateral view +G +acropodite, anterior view +H +acropodite, postero-mesal view +I +acropodite, mesal view +J +acropodite, posterior view +K +acropodite, lateral view +L +acropodite, lateral view. Abbreviations: ab = apical branch; fe = femorite; lb = lateral branch; mb = mesal branch; sb = subapical branch; sl = solenomere branch. Scale bars: 100 µm. + + + + + \ No newline at end of file diff --git a/data/4F/9F/CB/4F9FCBD8F2765C5BB032EDB13FA1CEE1.xml b/data/4F/9F/CB/4F9FCBD8F2765C5BB032EDB13FA1CEE1.xml new file mode 100644 index 00000000000..a9736f2b4f4 --- /dev/null +++ b/data/4F/9F/CB/4F9FCBD8F2765C5BB032EDB13FA1CEE1.xml @@ -0,0 +1,165 @@ + + + +Revision of the millipede family Dalodesmidae in Madagascar, with descriptions of two new Malagasy species of Dalodesmus Cook, 1896 (Diplopoda, Polydesmida) + + + +Author + +Wesener, Thomas +0000-0002-2028-3541 +Zoological Research Museum Alexander Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, D- 53113 Bonn, Germany + + + +Author + +Akkari, Nesrine +0000-0001-5019-4833 +Third Zoological Department, Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria + + + +Author + +Golovatch, Sergei I. +Institute of Ecology and Evolution, Russian Academy of Sciences, Leninsky prospekt 33, Moscow 119071, Russia + +text + + +ZooKeys + + +2025 + +2025-01-08 + + +1223 + + +185 +220 + + + +journal article +10.3897/zookeys.1223.139346 +451749E9-009E-43E9-A6F1-892035BDF1B0 + + + + + +Dalodesmus orator +Hoffman, 1974 + + + + + +Fig. 10 + + + + + + + +Dalodesmus orator + +Hoffman, 1974: 225 +, figs 1–9 (D); + +Golovatch and Hoffman 1989: 162 + +(L); + +Enghoff 2003: 623 + +(L); + +Wesener and Enghoff 2022: 926 + +(L). + + + + + + + + +Remark. + + +This species was described from a +♂ +holotype +, +two ♂ +and +two ♀ +paratypes +, all coming from Ambohimitombo, central +Madagascar +( +Hoffman 1974 +), in the collection of the Natural History Museum in London, +United Kingdom +. + + + + +Brief description. + + +(After +Hoffman 1974 +.) +♂ +holotype +~ +26.5 mm +long and up to 2.3 and +5.3 mm +wide on pro- and metazona, respectively; +♀ +up to +4.5 mm +wide. Colouration light reddish brown with yellow paratergal apices and legs. Paraterga largely clearly upturned above dorsum and directed more laterad than caudad, at ca 45 °, tips elongate and sharpened caudally; dorsal surface between paraterga mostly vaguely areate (Fig. +10 A +). + + + + + + + +Dalodesmus orator +Hoffman, 1974 + +, ♂ holotype, modified from +Hoffman 1974 +A +midbody tergite, dorsal view +B +gonopod, posterior view +C +acropodite, mesal view +D +gonopod, lateral view. Abbreviations: lb = lateral branch; mb = mesal branch; pfe = postfemorite; sl = solenomere branch. Not to scale. + + + +Gonopods (Fig. +10 B – D +) showing bare femorites (fe), coupled with rather complex acropodites: an unusually short and small solenomere (sl) directed ventrad, lying between the highest, suberect, distally unequally bifid mesal branch (mb) and a much shorter and strongly folded lateral branch (lb) directed ventrad. + + + + \ No newline at end of file diff --git a/data/B7/7B/E4/B77BE40B1CC852998AFAF927C052F425.xml b/data/B7/7B/E4/B77BE40B1CC852998AFAF927C052F425.xml new file mode 100644 index 00000000000..66ff3c48235 --- /dev/null +++ b/data/B7/7B/E4/B77BE40B1CC852998AFAF927C052F425.xml @@ -0,0 +1,324 @@ + + + +Revision of the millipede family Dalodesmidae in Madagascar, with descriptions of two new Malagasy species of Dalodesmus Cook, 1896 (Diplopoda, Polydesmida) + + + +Author + +Wesener, Thomas +0000-0002-2028-3541 +Zoological Research Museum Alexander Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, D- 53113 Bonn, Germany + + + +Author + +Akkari, Nesrine +0000-0001-5019-4833 +Third Zoological Department, Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria + + + +Author + +Golovatch, Sergei I. +Institute of Ecology and Evolution, Russian Academy of Sciences, Leninsky prospekt 33, Moscow 119071, Russia + +text + + +ZooKeys + + +2025 + +2025-01-08 + + +1223 + + +185 +220 + + + +journal article +10.3897/zookeys.1223.139346 +451749E9-009E-43E9-A6F1-892035BDF1B0 + + + + + +Dalodesmus kompantsevi + +sp. nov. + + + + +Figs 13 +, +14 + + + + +Type material. + + + + +Holotype + +• +♂ +( + +ZMUM + +), +Northern +Madagascar +, +Antsiranana Prov. +, + +4 km +SW of Joffreville + +(= Ambohitra), +Parc National Montagne d’Ambre +, + +12.51358 ° S +, +49.183001 ° E + +, + +900–1000 m +a. s. l. + +, tropical forest, + +16–18. XII. 2018 + +, +A. Kompantsev +leg. + + + + + +Other material. + + +• + +1 ♀ +, ( + +MZUF + +Fi- 30 A), +Madagascar +, +Montagne d’Ambre +, + +900 m + +, c / o grande cascade, leg. + +26 Sept. 1989 + +, +L. Bartolozzi +& +S. Taiti + +. + + + + +Diagnosis. + + +Tips of paraterga not projecting past posterior tergal margin (Fig. +13 A – C +), like in + +D. hova + +, + +D. tectus + +and + +D. speophilus + +sp. nov. +, vs sharper and projecting beyond margin in + +D. hamatus + +, + +D. odontopezus + +and + +D. orator + +. Differs from all other species of the genus primarily by the light colouration, coupled with the unusually compact, short, and clearly trifid acropodite, this being divided into three subequally long and upright branches: a middle, subacuminate and non-sigmoid solenomere (sl) flanked by a securiform, axe-shaped mesal branch (mb) and a nearly finger-shaped lateral branch (lb) of the solenophore (Fig. +13 E, F +). See also Key below. + + + + + + + +Dalodesmus kompantsevi + +sp. nov. +, ♂ holotype: +A – C +habitus, dorsal, lateral, and ventral views, respectively +D +body ring 7 with gonopods in situ, ventral view +E, F +gonopods, dorsal and ventral views, respectively. Abbreviations: fe femorite, lb lateral branch, mb mesal branch, sl solenomere. Photographs by K. Makarov, taken not to scale. + + + + + +Etymology. + + +To honour the late Aleksandr Kompantsev ( +Russia +), the collector. + + + + +Description. + + +Length of +holotype +~ +23 mm +, width of midbody pro- and metazona 1.5 and +2.9 mm +, respectively. Width of prozona +2.2 mm +, of metazona +4.2 mm +in +♀ +non-type. + + +Colouration in alcohol uniformly pale brown to beige, axial stripe on prozona thin, vague, and grey, antennae somewhat infuscate, increasingly brown distad (Fig. +13 A – C +). + + +Body with 20 rings. Tegument mainly dull to only slightly shining, microgranulate to microtuberculate throughout, even surfaces of prozona and of metazona below paraterga finely microgranulate, sterna granulate. Head also densely microtuberculate or granulate throughout, micropilose up to level of antennae; epicranial suture thin, but distinct; genae squarish, set off ventrally from gnathochilarial stipes by a small, but evident ridge (Fig. +13 A – C +). Interantennal isthmus ~ 2 × diameter of antennal socket (Fig. +13 C +). Antennae short and rather clavate, in situ reaching back past ring 3 when stretched dorsally, very densely setose and microgranulate. In length, antennomere 6> 5> 3> 4> 2> 1 = 7; antennomere 6 the largest and the highest, antennomeres 5 and 6 each with a small, round, distodorsal knob. In width, collum <head <ring 3 <2 = 4–16; thereafter body gradually tapering towards telson (Fig. +13 A +). Collum transversely suboval, regularly and broadly rounded laterally, densely tuberculate, most tuberculations being slightly oblong-oval, evident, equipped with very short, mostly subclavate setae and arranged in 7–8 transverse, rather irregular, arcuate rows. Metaterga 2 and 3 each with four similar transverse arcuated rows of setiferous tubercles, following metaterga each largely with 5–6 such rows (Figs +13 A +, +14 A, B +). Paraterga well-developed, set high (mostly at upper ¼ body), largely slightly upturned to subhorizontal, thus leaving the dorsum only faintly convex (Figs +13 B +, +14 B +); anterior and posterior margins of paraterga 2 and 3 clearly drawn both forward and caudad, following paraterga drawn increasingly only caudad, but caudal corners produced past rear tergal margin only on rings 17–19; posterior margins clearly bisinuate, well concave behind paraterga (Fig. +13 A – C +). Lateral and caudal margins of paraterga beset with numerous, similarly oblong, and usually subequal, setigerous tubercles / lobulations, caudolateral lobulation nearly sharp. Ozopores (Fig. +14 B, D +) inconspicuous, opening dorsally near penultimate lateral lobulation on pore-baring rings 5, 7, 9, 10, 12, 13, 15–19. Strictures between pro- and metazona narrow and rather deep, nearly smooth (Fig. +14 C +). Epiproct small, conical and subtruncate at tip. Hypoproct trapeziform, with 1 + 1 setae borne on distinct oblong knobs at caudal margin. Limbus very thin, small, and entire. Neither an axial line nor pleurosternal carinae (Fig. +13 A – C +). Endotergum inconspicuous, posterior margin of metazona projecting into long, sharp, apically microdenticulate, triangular projections (Fig. +14 E +). + + + + + + + +Dalodesmus kompantsevi + +sp. nov. +, ♀ ( +MZUF +Fi 30 A). +SEM +micrographs +A +midbody ring with pro- and metazonite, dorsal view +B +midbody ring, paranotum with ozopore +C +detail of surface structures of prozonite and metazonite +D +ozopore +E +endotergum +F +midbody legs. Scale bars: 1000 µm ( +A +); 100 µm ( +B, C +); 10 µm ( +D, E +); 200 µm ( +F +). + + + +Sterna mostly densely setose, deep, and narrow between coxae 1–3, increasingly broad thereafter, clearly excavate between coxae 5 and 6 ( +♂ +); postgonopodal sterna rather flat, devoid of modifications, cross-impressions faint (Fig. +13 C +). Gonopodal aperture roundly pentagonal, relatively small, taking up ~ 1 / 3 width of metazonum 7, clearly open and drawn into metazonum 6 (Fig. +13 D +). Legs incrassate, rather long. 1.4–1.5 × as long as body height, with small, stout, abundant and usual curved setae (Fig. +14 F +) with admixture of sphaerotrichomes ventrally on all podomeres ( +♂ +); gonopores on coxae 2 inconspicuous, each borne on a very small swelling ( +♂ +); prefemora not bulged laterally; claws very small; in length, tarsus> femur> prefemur> tibia> postfemur> coxa. + + +Gonopods (Fig. +13 D – F +) very slender and long, tips in situ reaching anteriorly beyond coxae 6 (Fig. +13 C, D +). Both coxites and clearly prefemoral (= densely setose) parts of telopodites equally very short and stout, fused medially, the former fully and the latter mostly hidden inside gonopodal aperture. Femorites (fe) contiguous medially in basal 1 / 3, basically bare and only one at base with a distinct lateral seta, both very faintly attenuating and diverging distad. Acropodites of both telopodites clearly diverging, but unusually compact, short, and clearly trifid, being divided into three subequally short and upright branches: a middle, subacuminate and non-sigmoid solenomere (sl) flanked by a solenophore represented by a securiform, axe-shaped mesal branch (mb) and a nearly finger-shaped lateral branch (lb) with a mesal knob parabasally. + + + + \ No newline at end of file