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httpUri="http://treatment.plazi.org/id/0325879DF232FFDDFF60FB4EFCC9D80B" lastPageId="14" lastPageNumber="15" pageId="2" pageNumber="3"> <subSubSection id="C3966500F232FFD1FF60FB4EFCC4D9ED" box="[156,772,1200,1225]" pageId="2" pageNumber="3" type="nomenclature"> <paragraph id="8B33368BF232FFD1FF60FB4EFCC4D9ED" blockId="2.[129,778,1013,1569]" box="[156,772,1200,1225]" pageId="2" pageNumber="3"> <taxonomicName id="4C8C4D08F232FFD1FF60FB4EFD70D9ED" authority="Delfino, Cohen & Macaluso" authorityName="Delfino, Cohen & Macaluso" box="[156,688,1200,1225]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="langebaanwegi"> @@ -112,7 +115,7 @@ Delfino, Cohen & Macaluso <subSubSection id="C3966500F232FFD1FF60FB2EFED5D9CC" box="[156,277,1232,1256]" pageId="2" pageNumber="3" type="description"> <paragraph id="8B33368BF232FFD1FF60FB2EFED5D9CC" blockId="2.[129,778,1013,1569]" box="[156,277,1232,1256]" pageId="2" pageNumber="3"> ( -<figureCitation id="13B72A0EF232FFD1FF5AFB2EFECAD9CC" box="[166,266,1232,1256]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStart-5="Figure 6" captionStart-6="Figure 7" captionStart-7="Figure 8" captionStart-8="Figure 9" captionStart-9="Figure 10" captionStartId-0="3.[113,178,1312,1336]" captionStartId-1="4.[129,194,1394,1418]" captionStartId-2="5.[113,178,1047,1071]" captionStartId-3="6.[129,194,1420,1444]" captionStartId-4="7.[113,178,1626,1650]" captionStartId-5="8.[129,194,1836,1860]" captionStartId-6="9.[114,179,1706,1730]" captionStartId-7="10.[129,194,1839,1863]" captionStartId-8="11.[113,178,1714,1738]" captionStartId-9="12.[129,194,1334,1358]" captionTargetBox-0="[117,1455,147,1281]" captionTargetBox-1="[130,1471,145,1364]" captionTargetBox-2="[147,1424,145,1017]" captionTargetBox-3="[131,1471,146,1390]" captionTargetBox-4="[115,1456,145,1596]" captionTargetBox-5="[169,1432,146,1806]" captionTargetBox-6="[115,1436,145,1676]" captionTargetBox-7="[137,1465,146,1809]" captionTargetBox-8="[131,1423,182,1679]" captionTargetBox-9="[132,1471,147,1304]" captionTargetId-0="figure-340@3.[114,1458,144,1284]" captionTargetId-1="figure-300@4.[129,1473,144,1366]" captionTargetId-2="figure-538@5.[146,1426,144,1019]" captionTargetId-3="figure-285@6.[129,1473,144,1392]" captionTargetId-4="figure-177@7.[114,1458,144,1598]" captionTargetId-5="figure-8@8.[168,1434,144,1808]" captionTargetId-6="figure-121@9.[114,1458,144,1678]" captionTargetId-7="figure-7@10.[136,1467,144,1811]" captionTargetId-8="figure-113@11.[114,1458,144,1687]" captionTargetId-9="figure-312@12.[129,1473,144,1306]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionTargetPageId-8="11" captionTargetPageId-9="12" captionText-0="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." captionText-1="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." captionText-2="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." captionText-3="Figure 4. Chersina langebaanwegi, paratype SAM-PQL-20925: male plastron in ventral view (A) and detail of the anterior lobe of the plastron in dorsal view (B); posterior lobe of the plastron in ventral view (C); pygal area in posterior view (D). Scale bars equal 5 cm." captionText-4="Figure 5. Chersina langebaanwegi, paratype SAM-PQL-21109, male: pygal area in posterior view (A); posterior portion of the shell in right lateral view (B); anterior lobe of the plastron in ventral and dorsal views (C, D); posterior lobe in ventral and dorsal views (E,F). Scale bars equal 5 cm." captionText-5="Figure 6. Chersina langebaanwegi, paratype SAM-PQL-PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm." captionText-6="Figure 7. Chersina langebaanwegi, paratype SAM-PQL-21749: nuchal in external view (A); pygal in external (B) and internal (C) views; anterior portion of the plastron (and part of the nuchal) in ventral view (D); xiphiplastra in ventral (E) and dorsal (F) views. Scale bars equal 7 cm in (A, D) and 5 cm in (B, C, E, F)." captionText-7="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." captionText-8="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." captionText-9="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." pageId="2" pageNumber="3">Figs 1–10</figureCitation> +<figureCitation id="13B72A0EF232FFD1FF5AFB2EFECAD9CC" box="[166,266,1232,1256]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStart-5="Figure 6" captionStart-6="Figure 7" captionStart-7="Figure 8" captionStart-8="Figure 9" captionStart-9="Figure 10" captionStartId-0="3.[113,178,1312,1336]" captionStartId-1="4.[129,194,1394,1418]" captionStartId-2="5.[113,178,1047,1071]" captionStartId-3="6.[129,194,1420,1444]" captionStartId-4="7.[113,178,1626,1650]" captionStartId-5="8.[129,194,1836,1860]" captionStartId-6="9.[114,179,1706,1730]" captionStartId-7="10.[129,194,1839,1863]" captionStartId-8="11.[113,178,1714,1738]" captionStartId-9="12.[129,194,1334,1358]" captionTargetBox-0="[117,1455,147,1281]" captionTargetBox-1="[130,1471,145,1364]" captionTargetBox-2="[147,1424,145,1017]" captionTargetBox-3="[131,1471,146,1390]" captionTargetBox-4="[115,1456,145,1596]" captionTargetBox-5="[169,1432,146,1806]" captionTargetBox-6="[115,1436,145,1676]" captionTargetBox-7="[137,1465,146,1809]" captionTargetBox-8="[131,1423,182,1679]" captionTargetBox-9="[132,1471,147,1304]" captionTargetId-0="figure-340@3.[114,1458,144,1284]" captionTargetId-1="figure-300@4.[129,1473,144,1366]" captionTargetId-2="figure-538@5.[146,1426,144,1019]" captionTargetId-3="figure-285@6.[129,1473,144,1392]" captionTargetId-4="figure-177@7.[114,1458,144,1598]" captionTargetId-5="figure-8@8.[168,1434,144,1808]" captionTargetId-6="figure-121@9.[114,1458,144,1678]" captionTargetId-7="figure-7@10.[136,1467,144,1811]" captionTargetId-8="figure-113@11.[114,1458,144,1687]" captionTargetId-9="figure-312@12.[129,1473,144,1306]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionTargetPageId-8="11" captionTargetPageId-9="12" captionText-0="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." captionText-1="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." captionText-2="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." captionText-3="Figure 4. Chersina langebaanwegi, paratype SAM-PQL-20925: male plastron in ventral view (A) and detail of the anterior lobe of the plastron in dorsal view (B); posterior lobe of the plastron in ventral view (C); pygal area in posterior view (D). Scale bars equal 5 cm." captionText-4="Figure 5. Chersina langebaanwegi, paratype SAM-PQL-21109, male: pygal area in posterior view (A); posterior portion of the shell in right lateral view (B); anterior lobe of the plastron in ventral and dorsal views (C, D); posterior lobe in ventral and dorsal views (E,F). Scale bars equal 5 cm." captionText-5="Figure 6. Chersina langebaanwegi, paratype SAM-PQL-PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm." captionText-6="Figure 7. Chersina langebaanwegi, paratype SAM-PQL-21749: nuchal in external view (A); pygal in external (B) and internal (C) views; anterior portion of the plastron (and part of the nuchal) in ventral view (D); xiphiplastra in ventral (E) and dorsal (F) views. Scale bars equal 7 cm in (A, D) and 5 cm in (B, C, E, F)." captionText-7="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." captionText-8="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." captionText-9="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." figureDoi-0="http://doi.org/10.5281/zenodo.15014591" figureDoi-1="http://doi.org/10.5281/zenodo.15014595" figureDoi-2="http://doi.org/10.5281/zenodo.15014597" figureDoi-3="http://doi.org/10.5281/zenodo.15014599" figureDoi-4="http://doi.org/10.5281/zenodo.15014601" figureDoi-5="http://doi.org/10.5281/zenodo.15014603" figureDoi-6="http://doi.org/10.5281/zenodo.15014607" figureDoi-7="http://doi.org/10.5281/zenodo.15014609" figureDoi-8="http://doi.org/10.5281/zenodo.15014611" figureDoi-9="http://doi.org/10.5281/zenodo.15014613" httpUri-0="https://zenodo.org/record/15014591/files/figure.png" httpUri-1="https://zenodo.org/record/15014595/files/figure.png" httpUri-2="https://zenodo.org/record/15014597/files/figure.png" httpUri-3="https://zenodo.org/record/15014599/files/figure.png" httpUri-4="https://zenodo.org/record/15014601/files/figure.png" httpUri-5="https://zenodo.org/record/15014603/files/figure.png" httpUri-6="https://zenodo.org/record/15014607/files/figure.png" httpUri-7="https://zenodo.org/record/15014609/files/figure.png" httpUri-8="https://zenodo.org/record/15014611/files/figure.png" httpUri-9="https://zenodo.org/record/15014613/files/figure.png" pageId="2" pageNumber="3">Figs 1–10</figureCitation> ) </paragraph> </subSubSection> @@ -221,7 +224,7 @@ urn:lsid:zoobank.org:pub:A8B93307-034E - 4D7F -96BC -B17F8F32B122. : nearly complete <specimenCount id="9D8AFD02F232FFD1FAC3FF6EFA43DD8C" box="[1343,1411,144,168]" pageId="2" pageNumber="3" type="female">female</specimenCount> shell ( -<figureCitation id="13B72A0EF232FFD1FCB8FF51FC5CDDE3" box="[836,924,175,199]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="3.[113,178,1312,1336]" captionStartId-1="4.[129,194,1394,1418]" captionStartId-2="5.[113,178,1047,1071]" captionTargetBox-0="[117,1455,147,1281]" captionTargetBox-1="[130,1471,145,1364]" captionTargetBox-2="[147,1424,145,1017]" captionTargetId-0="figure-340@3.[114,1458,144,1284]" captionTargetId-1="figure-300@4.[129,1473,144,1366]" captionTargetId-2="figure-538@5.[146,1426,144,1019]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." captionText-1="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." captionText-2="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." pageId="2" pageNumber="3">Figs 1–3</figureCitation> +<figureCitation id="13B72A0EF232FFD1FCB8FF51FC5CDDE3" box="[836,924,175,199]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="3.[113,178,1312,1336]" captionStartId-1="4.[129,194,1394,1418]" captionStartId-2="5.[113,178,1047,1071]" captionTargetBox-0="[117,1455,147,1281]" captionTargetBox-1="[130,1471,145,1364]" captionTargetBox-2="[147,1424,145,1017]" captionTargetId-0="figure-340@3.[114,1458,144,1284]" captionTargetId-1="figure-300@4.[129,1473,144,1366]" captionTargetId-2="figure-538@5.[146,1426,144,1019]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." captionText-1="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." captionText-2="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." figureDoi-0="http://doi.org/10.5281/zenodo.15014591" figureDoi-1="http://doi.org/10.5281/zenodo.15014595" figureDoi-2="http://doi.org/10.5281/zenodo.15014597" httpUri-0="https://zenodo.org/record/15014591/files/figure.png" httpUri-1="https://zenodo.org/record/15014595/files/figure.png" httpUri-2="https://zenodo.org/record/15014597/files/figure.png" pageId="2" pageNumber="3">Figs 1–3</figureCitation> ) </materialsCitation> . @@ -238,7 +241,7 @@ shell ( : partial <specimenCount id="9D8AFD02F232FFD1FB3AFF16FB37DC24" box="[1222,1271,232,256]" pageId="2" pageNumber="3" type="male">male</specimenCount> shell with a nearly complete plastron, several peripherals and pygal ( -<figureCitation id="13B72A0EF232FFD1FA91FEF6FA70DC04" box="[1389,1456,264,288]" captionStart="Figure 4" captionStartId="6.[129,194,1420,1444]" captionTargetBox="[131,1471,146,1390]" captionTargetId="figure-285@6.[129,1473,144,1392]" captionTargetPageId="6" captionText="Figure 4. Chersina langebaanwegi, paratype SAM-PQL-20925: male plastron in ventral view (A) and detail of the anterior lobe of the plastron in dorsal view (B); posterior lobe of the plastron in ventral view (C); pygal area in posterior view (D). Scale bars equal 5 cm." pageId="2" pageNumber="3">Fig. 4</figureCitation> +<figureCitation id="13B72A0EF232FFD1FA91FEF6FA70DC04" box="[1389,1456,264,288]" captionStart="Figure 4" captionStartId="6.[129,194,1420,1444]" captionTargetBox="[131,1471,146,1390]" captionTargetId="figure-285@6.[129,1473,144,1392]" captionTargetPageId="6" captionText="Figure 4. Chersina langebaanwegi, paratype SAM-PQL-20925: male plastron in ventral view (A) and detail of the anterior lobe of the plastron in dorsal view (B); posterior lobe of the plastron in ventral view (C); pygal area in posterior view (D). Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014599" httpUri="https://zenodo.org/record/15014599/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation> ); </materialsCitation> <materialsCitation id="3BE43CD6F232FFD1FCC5FED9FA02DC7B" country="South Africa" latitude="-32.966667" longLatPrecision="1207" longitude="18.15" municipality="Cape Town" pageId="2" pageNumber="3" specimenCode="SAM-PQL-21109" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="paratype"> @@ -248,7 +251,7 @@ shell with a nearly complete plastron, several peripherals and pygal ( : partially preserved shell of a <specimenCount id="9D8AFD02F232FFD1FAA5FED9FA4ADC1B" box="[1369,1418,295,319]" pageId="2" pageNumber="3" type="male">male</specimenCount> spe - cimen missing most of the costals and all the neurals ( -<figureCitation id="13B72A0EF232FFD1FA89FEB8FA70DC7A" box="[1397,1456,326,350]" captionStart="Figure 5" captionStartId="7.[113,178,1626,1650]" captionTargetBox="[115,1456,145,1596]" captionTargetId="figure-177@7.[114,1458,144,1598]" captionTargetPageId="7" captionText="Figure 5. Chersina langebaanwegi, paratype SAM-PQL-21109, male: pygal area in posterior view (A); posterior portion of the shell in right lateral view (B); anterior lobe of the plastron in ventral and dorsal views (C, D); posterior lobe in ventral and dorsal views (E,F). Scale bars equal 5 cm." pageId="2" pageNumber="3">Fig. 5</figureCitation> +<figureCitation id="13B72A0EF232FFD1FA89FEB8FA70DC7A" box="[1397,1456,326,350]" captionStart="Figure 5" captionStartId="7.[113,178,1626,1650]" captionTargetBox="[115,1456,145,1596]" captionTargetId="figure-177@7.[114,1458,144,1598]" captionTargetPageId="7" captionText="Figure 5. Chersina langebaanwegi, paratype SAM-PQL-21109, male: pygal area in posterior view (A); posterior portion of the shell in right lateral view (B); anterior lobe of the plastron in ventral and dorsal views (C, D); posterior lobe in ventral and dorsal views (E,F). Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014601" httpUri="https://zenodo.org/record/15014601/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation> ); </materialsCitation> <materialsCitation id="3BE43CD6F232FFD1FCC5FE9BFB59DC99" country="South Africa" latitude="-32.966667" longLatPrecision="1207" longitude="18.15" municipality="Cape Town" pageId="2" pageNumber="3" specimenCode="SAM-PQL-21174" specimenCount="1" stateProvince="Western Cape" typeStatus="paratype"> @@ -256,7 +259,7 @@ spe - cimen missing most of the costals and all the neurals ( <specimenCode id="DB2A9EF0F232FFD1FCC5FE9BFC3CDC59" box="[825,1020,357,381]" pageId="2" pageNumber="3">SAM-PQL-21174</specimenCode> </emphasis> : highly fragmentary, incomplete shell whose best -preserved elements are the nuchal, fifth neural, pygal, and, above all, both epiplastra ( -<figureCitation id="13B72A0EF232FFD1FBB1FE5AFB47DC98" box="[1101,1159,420,444]" captionStart="Figure 6" captionStartId="8.[129,194,1836,1860]" captionTargetBox="[169,1432,146,1806]" captionTargetId="figure-8@8.[168,1434,144,1808]" captionTargetPageId="8" captionText="Figure 6. Chersina langebaanwegi, paratype SAM-PQL-PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm." pageId="2" pageNumber="3">Fig. 6</figureCitation> +<figureCitation id="13B72A0EF232FFD1FBB1FE5AFB47DC98" box="[1101,1159,420,444]" captionStart="Figure 6" captionStartId="8.[129,194,1836,1860]" captionTargetBox="[169,1432,146,1806]" captionTargetId="figure-8@8.[168,1434,144,1808]" captionTargetPageId="8" captionText="Figure 6. Chersina langebaanwegi, paratype SAM-PQL-PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014603" httpUri="https://zenodo.org/record/15014603/files/figure.png" pageId="2" pageNumber="3">Fig. 6</figureCitation> ); </materialsCitation> <materialsCitation id="3BE43CD6F232FFD1FB5CFE5AFB67DF3F" country="South Africa" latitude="-32.966667" longLatPrecision="1207" longitude="18.15" municipality="Cape Town" pageId="2" pageNumber="3" specimenCode="SAM-PQL-21749" specimenCount="1" stateProvince="Western Cape" typeStatus="paratype"> @@ -264,7 +267,7 @@ spe - cimen missing most of the costals and all the neurals ( <specimenCode id="DB2A9EF0F232FFD1FB5CFE5AFAA2DC98" box="[1184,1378,420,444]" pageId="2" pageNumber="3">SAM-PQL-21749</specimenCode> </emphasis> : crushed but nearly complete shell with partially articulated elements, including well -preserved and well -exposed nuchal, pygal, epiplastra, and xiphiplastra ( -<figureCitation id="13B72A0EF232FFD1FBA0FDFCFB55DF3E" box="[1116,1173,514,538]" captionStart="Figure 7" captionStartId="9.[114,179,1706,1730]" captionTargetBox="[115,1436,145,1676]" captionTargetId="figure-121@9.[114,1458,144,1678]" captionTargetPageId="9" captionText="Figure 7. Chersina langebaanwegi, paratype SAM-PQL-21749: nuchal in external view (A); pygal in external (B) and internal (C) views; anterior portion of the plastron (and part of the nuchal) in ventral view (D); xiphiplastra in ventral (E) and dorsal (F) views. Scale bars equal 7 cm in (A, D) and 5 cm in (B, C, E, F)." pageId="2" pageNumber="3">Fig. 7</figureCitation> +<figureCitation id="13B72A0EF232FFD1FBA0FDFCFB55DF3E" box="[1116,1173,514,538]" captionStart="Figure 7" captionStartId="9.[114,179,1706,1730]" captionTargetBox="[115,1436,145,1676]" captionTargetId="figure-121@9.[114,1458,144,1678]" captionTargetPageId="9" captionText="Figure 7. Chersina langebaanwegi, paratype SAM-PQL-21749: nuchal in external view (A); pygal in external (B) and internal (C) views; anterior portion of the plastron (and part of the nuchal) in ventral view (D); xiphiplastra in ventral (E) and dorsal (F) views. Scale bars equal 7 cm in (A, D) and 5 cm in (B, C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.15014607" httpUri="https://zenodo.org/record/15014607/files/figure.png" pageId="2" pageNumber="3">Fig. 7</figureCitation> ); </materialsCitation> <materialsCitation id="3BE43CD6F232FFD1FB51FDFCFB99DF1E" country="South Africa" latitude="-32.966667" longLatPrecision="1207" longitude="18.15" municipality="Cape Town" pageId="2" pageNumber="3" specimenCode="SAM-PQL-22076" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="paratype"> @@ -274,7 +277,7 @@ spe - cimen missing most of the costals and all the neurals ( : a com - plete <specimenCount id="9D8AFD02F232FFD1FC8EFDDCFC63DF1E" box="[882,931,546,570]" pageId="2" pageNumber="3" type="male">male</specimenCount> plastron ( -<figureCitation id="13B72A0EF232FFD1FBF2FDDCFB87DF1E" box="[1038,1095,546,570]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." pageId="2" pageNumber="3">Fig. 8</figureCitation> +<figureCitation id="13B72A0EF232FFD1FBF2FDDCFB87DF1E" box="[1038,1095,546,570]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014609" httpUri="https://zenodo.org/record/15014609/files/figure.png" pageId="2" pageNumber="3">Fig. 8</figureCitation> ); </materialsCitation> <materialsCitation id="3BE43CD6F232FFD1FB9CFDDFFC17DF93" country="South Africa" latitude="-32.966667" longLatPrecision="1207" longitude="18.15" municipality="Cape Town" pageId="2" pageNumber="3" specimenCode="SAM-PQL-35023d" specimenCount="1" stateProvince="Western Cape" typeStatus="paratype"> @@ -284,7 +287,7 @@ plastron ( : partially pre - served shell of a very large <specimenCount id="9D8AFD02F232FFD1FBA8FDBFFB75DF7D" box="[1108,1205,577,601]" pageId="2" pageNumber="3" type="generic">specimen</specimenCount> represented mostly by an epiplastra, entoplastron, left hyoplastron, three neurals, pygal with the surrounding fragmentary peripherals, and three peri - pherals ( -<figureCitation id="13B72A0EF232FFD1FC6FFD61FC0BDF93" box="[915,971,671,695]" captionStart="Figure 9" captionStartId="11.[113,178,1714,1738]" captionTargetBox="[131,1423,182,1679]" captionTargetId="figure-113@11.[114,1458,144,1687]" captionTargetPageId="11" captionText="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." pageId="2" pageNumber="3">Fig. 9</figureCitation> +<figureCitation id="13B72A0EF232FFD1FC6FFD61FC0BDF93" box="[915,971,671,695]" captionStart="Figure 9" captionStartId="11.[113,178,1714,1738]" captionTargetBox="[131,1423,182,1679]" captionTargetId="figure-113@11.[114,1458,144,1687]" captionTargetPageId="11" captionText="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014611" httpUri="https://zenodo.org/record/15014611/files/figure.png" pageId="2" pageNumber="3">Fig. 9</figureCitation> ) </materialsCitation> . @@ -660,7 +663,7 @@ Jiangzuo ; and literature therein). </paragraph> </subSubSection> -<caption id="DFF36603F233FFD0FF8DFADEFCF4D88C" pageId="3" pageNumber="4" startId="3.[113,178,1312,1336]" targetBox="[117,1455,147,1281]" targetPageId="3" targetType="figure"> +<caption id="DFF36603F233FFD0FF8DFADEFCF4D88C" ID-DOI="http://doi.org/10.5281/zenodo.15014591" ID-Zenodo-Dep="15014591" httpUri="https://zenodo.org/record/15014591/files/figure.png" pageId="3" pageNumber="4" startId="3.[113,178,1312,1336]" targetBox="[117,1455,147,1281]" targetPageId="3" targetType="figure"> <paragraph id="8B33368BF233FFD0FF8DFADEFCF4D88C" blockId="3.[113,1459,1312,1449]" pageId="3" pageNumber="4"> <emphasis id="B9F8EA99F233FFD0FF8DFADEFF09D81C" bold="true" box="[113,201,1312,1336]" pageId="3" pageNumber="4">Figure 1.</emphasis> <taxonomicName id="4C8C4D08F233FFD0FF32FADEFE67D81C" box="[206,423,1312,1336]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="langebaanwegi"> @@ -692,47 +695,47 @@ regarding the following characters (see Supporting information for the compariso <emphasis id="B9F8EA99F233FFD0FE91F894FE20DAA6" box="[365,480,1898,1922]" italics="true" pageId="3" pageNumber="4">C. angulata</emphasis> </taxonomicName> ( -<figureCitation id="13B72A0EF233FFD0FE0CF894FD95DAA5" box="[496,597,1898,1922]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="3" pageNumber="4">Fig. 1E, F</figureCitation> +<figureCitation id="13B72A0EF233FFD0FE0CF894FD95DAA5" box="[496,597,1898,1922]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014591" httpUri="https://zenodo.org/record/15014591/files/figure.png" pageId="3" pageNumber="4">Fig. 1E, F</figureCitation> ; Supporting in - formation, -<figureCitation id="13B72A0EF233FFD0FF18F877FEEADA85" box="[228,298,1929,1953]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="3" pageNumber="4">Fig. S1</figureCitation> +<figureCitation id="13B72A0EF233FFD0FF18F877FEEADA85" box="[228,298,1929,1953]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014591" httpUri="https://zenodo.org/record/15014591/files/figure.png" pageId="3" pageNumber="4">Fig. S1</figureCitation> ); the nuchal is not as significantly arched (in anterior view) as that of <taxonomicName id="4C8C4D08F233FFD0FBCFFA26FB68D8D4" box="[1075,1192,1496,1520]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="angulata"> <emphasis id="B9F8EA99F233FFD0FBCFFA26FB68D8D4" box="[1075,1192,1496,1520]" italics="true" pageId="3" pageNumber="4">C. angulata</emphasis> </taxonomicName> ( -<figureCitation id="13B72A0EF233FFD0FB47FA26FACAD8D4" box="[1211,1290,1496,1520]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="3" pageNumber="4">Fig. 1G</figureCitation> +<figureCitation id="13B72A0EF233FFD0FB47FA26FACAD8D4" box="[1211,1290,1496,1520]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014591" httpUri="https://zenodo.org/record/15014591/files/figure.png" pageId="3" pageNumber="4">Fig. 1G</figureCitation> ; Supporting in - formation, -<figureCitation id="13B72A0EF233FFD0FC62FA09FC27DB2B" box="[926,999,1527,1551]" captionStart="Figure 2" captionStartId="4.[129,194,1394,1418]" captionTargetBox="[130,1471,145,1364]" captionTargetId="figure-300@4.[129,1473,144,1366]" captionTargetPageId="4" captionText="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="3" pageNumber="4">Fig. S2</figureCitation> +<figureCitation id="13B72A0EF233FFD0FC62FA09FC27DB2B" box="[926,999,1527,1551]" captionStart="Figure 2" captionStartId="4.[129,194,1394,1418]" captionTargetBox="[130,1471,145,1364]" captionTargetId="figure-300@4.[129,1473,144,1366]" captionTargetPageId="4" captionText="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014595" httpUri="https://zenodo.org/record/15014595/files/figure.png" pageId="3" pageNumber="4">Fig. S2</figureCitation> ) and does not develop a ventral transversal ridge with medial notch and a dorsal shelf as in <taxonomicName id="4C8C4D08F233FFD0FAF0F9E9FABDDB0B" box="[1292,1405,1559,1583]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="angulata"> <emphasis id="B9F8EA99F233FFD0FAF0F9E9FABDDB0B" box="[1292,1405,1559,1583]" italics="true" pageId="3" pageNumber="4">C. angulata</emphasis> </taxonomicName> ( -<figureCitation id="13B72A0EF233FFD0FA71F9E9FC87DB6A" captionStart="Figure 6" captionStartId="8.[129,194,1836,1860]" captionTargetBox="[169,1432,146,1806]" captionTargetId="figure-8@8.[168,1434,144,1808]" captionTargetPageId="8" captionText="Figure 6. Chersina langebaanwegi, paratype SAM-PQL-PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm." pageId="3" pageNumber="4">Fig. 6B</figureCitation> +<figureCitation id="13B72A0EF233FFD0FA71F9E9FC87DB6A" captionStart="Figure 6" captionStartId="8.[129,194,1836,1860]" captionTargetBox="[169,1432,146,1806]" captionTargetId="figure-8@8.[168,1434,144,1808]" captionTargetPageId="8" captionText="Figure 6. Chersina langebaanwegi, paratype SAM-PQL-PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014603" httpUri="https://zenodo.org/record/15014603/files/figure.png" pageId="3" pageNumber="4">Fig. 6B</figureCitation> ; Supporting information, -<figureCitation id="13B72A0EF233FFD0FBAFF9C8FB03DB69" box="[1107,1219,1590,1614]" captionStart="Figure 3" captionStartId="5.[113,178,1047,1071]" captionTargetBox="[147,1424,145,1017]" captionTargetId="figure-538@5.[146,1426,144,1019]" captionTargetPageId="5" captionText="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." pageId="3" pageNumber="4">Fig. S3A, B</figureCitation> +<figureCitation id="13B72A0EF233FFD0FBAFF9C8FB03DB69" box="[1107,1219,1590,1614]" captionStart="Figure 3" captionStartId="5.[113,178,1047,1071]" captionTargetBox="[147,1424,145,1017]" captionTargetId="figure-538@5.[146,1426,144,1019]" captionTargetPageId="5" captionText="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." figureDoi="http://doi.org/10.5281/zenodo.15014597" httpUri="https://zenodo.org/record/15014597/files/figure.png" pageId="3" pageNumber="4">Fig. S3A, B</figureCitation> ); the cervical is, on the dorsal surface, much broader and proportionally longer than that of <taxonomicName id="4C8C4D08F233FFD0FC87F98BFC31DBA8" box="[891,1009,1652,1676]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="angulata"> <emphasis id="B9F8EA99F233FFD0FC87F98BFC31DBA8" box="[891,1009,1652,1676]" italics="true" pageId="3" pageNumber="4">C. angulata</emphasis> </taxonomicName> ( -<figureCitation id="13B72A0EF233FFD0FBFAF98AFBB4DBA8" box="[1030,1140,1652,1677]" captionStart="Figure 2" captionStartId="4.[129,194,1394,1418]" captionTargetBox="[130,1471,145,1364]" captionTargetId="figure-300@4.[129,1473,144,1366]" captionTargetPageId="4" captionText="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="3" pageNumber="4">Fig. 2A, B</figureCitation> +<figureCitation id="13B72A0EF233FFD0FBFAF98AFBB4DBA8" box="[1030,1140,1652,1677]" captionStart="Figure 2" captionStartId="4.[129,194,1394,1418]" captionTargetBox="[130,1471,145,1364]" captionTargetId="figure-300@4.[129,1473,144,1366]" captionTargetPageId="4" captionText="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014595" httpUri="https://zenodo.org/record/15014595/files/figure.png" pageId="3" pageNumber="4">Fig. 2A, B</figureCitation> ; Supporting information, -<figureCitation id="13B72A0EF233FFD0FA71F98AFCB4DB8F" captionStart="Figure 3" captionStartId="5.[113,178,1047,1071]" captionTargetBox="[147,1424,145,1017]" captionTargetId="figure-538@5.[146,1426,144,1019]" captionTargetPageId="5" captionText="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." pageId="3" pageNumber="4">Fig. S3C, D</figureCitation> +<figureCitation id="13B72A0EF233FFD0FA71F98AFCB4DB8F" captionStart="Figure 3" captionStartId="5.[113,178,1047,1071]" captionTargetBox="[147,1424,145,1017]" captionTargetId="figure-538@5.[146,1426,144,1019]" captionTargetPageId="5" captionText="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." figureDoi="http://doi.org/10.5281/zenodo.15014597" httpUri="https://zenodo.org/record/15014597/files/figure.png" pageId="3" pageNumber="4">Fig. S3C, D</figureCitation> ); the first neural is hexagonal and not rectangular as in <taxonomicName id="4C8C4D08F233FFD0FA61F96AFC40DBEF" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="angulata"> <emphasis id="B9F8EA99F233FFD0FA61F96AFC40DBEF" italics="true" pageId="3" pageNumber="4">C. angulata</emphasis> </taxonomicName> ( -<figureCitation id="13B72A0EF233FFD0FC6DF94DFBD3DBEF" box="[913,1043,1715,1739]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="3" pageNumber="4">Figure 1A, B</figureCitation> +<figureCitation id="13B72A0EF233FFD0FC6DF94DFBD3DBEF" box="[913,1043,1715,1739]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014591" httpUri="https://zenodo.org/record/15014591/files/figure.png" pageId="3" pageNumber="4">Figure 1A, B</figureCitation> ; Supporting information, -<figureCitation id="13B72A0EF233FFD0FADDF94DFAA7DBEF" box="[1313,1383,1715,1739]" captionStart="Figure 4" captionStartId="6.[129,194,1420,1444]" captionTargetBox="[131,1471,146,1390]" captionTargetId="figure-285@6.[129,1473,144,1392]" captionTargetPageId="6" captionText="Figure 4. Chersina langebaanwegi, paratype SAM-PQL-20925: male plastron in ventral view (A) and detail of the anterior lobe of the plastron in dorsal view (B); posterior lobe of the plastron in ventral view (C); pygal area in posterior view (D). Scale bars equal 5 cm." pageId="3" pageNumber="4">Fig. S4</figureCitation> +<figureCitation id="13B72A0EF233FFD0FADDF94DFAA7DBEF" box="[1313,1383,1715,1739]" captionStart="Figure 4" captionStartId="6.[129,194,1420,1444]" captionTargetBox="[131,1471,146,1390]" captionTargetId="figure-285@6.[129,1473,144,1392]" captionTargetPageId="6" captionText="Figure 4. Chersina langebaanwegi, paratype SAM-PQL-20925: male plastron in ventral view (A) and detail of the anterior lobe of the plastron in dorsal view (B); posterior lobe of the plastron in ventral view (C); pygal area in posterior view (D). Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014599" httpUri="https://zenodo.org/record/15014599/files/figure.png" pageId="3" pageNumber="4">Fig. S4</figureCitation> ); most of the neurals host the ribheads ( -<figureCitation id="13B72A0EF233FFD0FB80F92CFB0ADBCF" box="[1148,1226,1746,1771]" captionStart="Figure 3" captionStartId="5.[113,178,1047,1071]" captionTargetBox="[147,1424,145,1017]" captionTargetId="figure-538@5.[146,1426,144,1019]" captionTargetPageId="5" captionText="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." pageId="3" pageNumber="4">Figs 3A</figureCitation> +<figureCitation id="13B72A0EF233FFD0FB80F92CFB0ADBCF" box="[1148,1226,1746,1771]" captionStart="Figure 3" captionStartId="5.[113,178,1047,1071]" captionTargetBox="[147,1424,145,1017]" captionTargetId="figure-538@5.[146,1426,144,1019]" captionTargetPageId="5" captionText="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." figureDoi="http://doi.org/10.5281/zenodo.15014597" httpUri="https://zenodo.org/record/15014597/files/figure.png" pageId="3" pageNumber="4">Figs 3A</figureCitation> , -<figureCitation id="13B72A0EF233FFD0FB2AF92DFB32DBCF" box="[1238,1266,1747,1771]" captionStart="Figure 9" captionStartId="11.[113,178,1714,1738]" captionTargetBox="[131,1423,182,1679]" captionTargetId="figure-113@11.[114,1458,144,1687]" captionTargetPageId="11" captionText="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." pageId="3" pageNumber="4">9B</figureCitation> +<figureCitation id="13B72A0EF233FFD0FB2AF92DFB32DBCF" box="[1238,1266,1747,1771]" captionStart="Figure 9" captionStartId="11.[113,178,1714,1738]" captionTargetBox="[131,1423,182,1679]" captionTargetId="figure-113@11.[114,1458,144,1687]" captionTargetPageId="11" captionText="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014611" httpUri="https://zenodo.org/record/15014611/files/figure.png" pageId="3" pageNumber="4">9B</figureCitation> ; Supporting infor - mation, -<figureCitation id="13B72A0EF233FFD0FC7CF90CFC06DA2E" box="[896,966,1778,1802]" captionStart="Figure 5" captionStartId="7.[113,178,1626,1650]" captionTargetBox="[115,1456,145,1596]" captionTargetId="figure-177@7.[114,1458,144,1598]" captionTargetPageId="7" captionText="Figure 5. Chersina langebaanwegi, paratype SAM-PQL-21109, male: pygal area in posterior view (A); posterior portion of the shell in right lateral view (B); anterior lobe of the plastron in ventral and dorsal views (C, D); posterior lobe in ventral and dorsal views (E,F). Scale bars equal 5 cm." pageId="3" pageNumber="4">Fig. S5</figureCitation> +<figureCitation id="13B72A0EF233FFD0FC7CF90CFC06DA2E" box="[896,966,1778,1802]" captionStart="Figure 5" captionStartId="7.[113,178,1626,1650]" captionTargetBox="[115,1456,145,1596]" captionTargetId="figure-177@7.[114,1458,144,1598]" captionTargetPageId="7" captionText="Figure 5. Chersina langebaanwegi, paratype SAM-PQL-21109, male: pygal area in posterior view (A); posterior portion of the shell in right lateral view (B); anterior lobe of the plastron in ventral and dorsal views (C, D); posterior lobe in ventral and dorsal views (E,F). Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014601" httpUri="https://zenodo.org/record/15014601/files/figure.png" pageId="3" pageNumber="4">Fig. S5</figureCitation> ) and not regularly the costals as in <taxonomicName id="4C8C4D08F233FFD0FAC5F90CFA6CDA2E" box="[1337,1452,1778,1802]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="angulata"> <emphasis id="B9F8EA99F233FFD0FAC5F90CFA6CDA2E" box="[1337,1452,1778,1802]" italics="true" pageId="3" pageNumber="4">C. angulata</emphasis> @@ -742,75 +745,75 @@ regarding the following characters (see Supporting information for the compariso <emphasis id="B9F8EA99F233FFD0FC9DF8CFFC14DA6C" box="[865,980,1840,1864]" italics="true" pageId="3" pageNumber="4">C. angulata</emphasis> </taxonomicName> ( -<figureCitation id="13B72A0EF233FFD0FC19F8CEFB8ADA6C" box="[997,1098,1840,1865]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="3" pageNumber="4">Fig. 1E, F</figureCitation> +<figureCitation id="13B72A0EF233FFD0FC19F8CEFB8ADA6C" box="[997,1098,1840,1865]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014591" httpUri="https://zenodo.org/record/15014591/files/figure.png" pageId="3" pageNumber="4">Fig. 1E, F</figureCitation> ; Supporting information, -<figureCitation id="13B72A0EF233FFD0FAA6F8CEFA61DA6C" box="[1370,1441,1840,1864]" captionStart="Figure 6" captionStartId="8.[129,194,1836,1860]" captionTargetBox="[169,1432,146,1806]" captionTargetId="figure-8@8.[168,1434,144,1808]" captionTargetPageId="8" captionText="Figure 6. Chersina langebaanwegi, paratype SAM-PQL-PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm." pageId="3" pageNumber="4">Fig. S6</figureCitation> +<figureCitation id="13B72A0EF233FFD0FAA6F8CEFA61DA6C" box="[1370,1441,1840,1864]" captionStart="Figure 6" captionStartId="8.[129,194,1836,1860]" captionTargetBox="[169,1432,146,1806]" captionTargetId="figure-8@8.[168,1434,144,1808]" captionTargetPageId="8" captionText="Figure 6. Chersina langebaanwegi, paratype SAM-PQL-PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014603" httpUri="https://zenodo.org/record/15014603/files/figure.png" pageId="3" pageNumber="4">Fig. S6</figureCitation> ); the anterior protrusion of the plastron (the gular protrusion of the epiplastra) is comparatively more developed than in <taxonomicName id="4C8C4D08F233FFD0FA61F88EFC40DA82" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="angulata"> <emphasis id="B9F8EA99F233FFD0FA61F88EFC40DA82" italics="true" pageId="3" pageNumber="4">C. angulata</emphasis> </taxonomicName> (especially in males; -<figureCitation id="13B72A0EF233FFD0FBA9F870FB7BDA82" box="[1109,1211,1934,1959]" captionStart="Figure 4" captionStartId="6.[129,194,1420,1444]" captionTargetBox="[131,1471,146,1390]" captionTargetId="figure-285@6.[129,1473,144,1392]" captionTargetPageId="6" captionText="Figure 4. Chersina langebaanwegi, paratype SAM-PQL-20925: male plastron in ventral view (A) and detail of the anterior lobe of the plastron in dorsal view (B); posterior lobe of the plastron in ventral view (C); pygal area in posterior view (D). Scale bars equal 5 cm." pageId="3" pageNumber="4">Figs 4A, B</figureCitation> +<figureCitation id="13B72A0EF233FFD0FBA9F870FB7BDA82" box="[1109,1211,1934,1959]" captionStart="Figure 4" captionStartId="6.[129,194,1420,1444]" captionTargetBox="[131,1471,146,1390]" captionTargetId="figure-285@6.[129,1473,144,1392]" captionTargetPageId="6" captionText="Figure 4. Chersina langebaanwegi, paratype SAM-PQL-20925: male plastron in ventral view (A) and detail of the anterior lobe of the plastron in dorsal view (B); posterior lobe of the plastron in ventral view (C); pygal area in posterior view (D). Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014599" httpUri="https://zenodo.org/record/15014599/files/figure.png" pageId="3" pageNumber="4">Figs 4A, B</figureCitation> , -<figureCitation id="13B72A0EF233FFD0FB39F870FAC0DA82" box="[1221,1280,1934,1958]" captionStart="Figure 5" captionStartId="7.[113,178,1626,1650]" captionTargetBox="[115,1456,145,1596]" captionTargetId="figure-177@7.[114,1458,144,1598]" captionTargetPageId="7" captionText="Figure 5. Chersina langebaanwegi, paratype SAM-PQL-21109, male: pygal area in posterior view (A); posterior portion of the shell in right lateral view (B); anterior lobe of the plastron in ventral and dorsal views (C, D); posterior lobe in ventral and dorsal views (E,F). Scale bars equal 5 cm." pageId="3" pageNumber="4">5C, D</figureCitation> +<figureCitation id="13B72A0EF233FFD0FB39F870FAC0DA82" box="[1221,1280,1934,1958]" captionStart="Figure 5" captionStartId="7.[113,178,1626,1650]" captionTargetBox="[115,1456,145,1596]" captionTargetId="figure-177@7.[114,1458,144,1598]" captionTargetPageId="7" captionText="Figure 5. Chersina langebaanwegi, paratype SAM-PQL-21109, male: pygal area in posterior view (A); posterior portion of the shell in right lateral view (B); anterior lobe of the plastron in ventral and dorsal views (C, D); posterior lobe in ventral and dorsal views (E,F). Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014601" httpUri="https://zenodo.org/record/15014601/files/figure.png" pageId="3" pageNumber="4">5C, D</figureCitation> , -<figureCitation id="13B72A0EF233FFD0FAF6F870FA82DA82" box="[1290,1346,1934,1958]" captionStart="Figure 6" captionStartId="8.[129,194,1836,1860]" captionTargetBox="[169,1432,146,1806]" captionTargetId="figure-8@8.[168,1434,144,1808]" captionTargetPageId="8" captionText="Figure 6. Chersina langebaanwegi, paratype SAM-PQL-PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm." pageId="3" pageNumber="4">6D, E</figureCitation> +<figureCitation id="13B72A0EF233FFD0FAF6F870FA82DA82" box="[1290,1346,1934,1958]" captionStart="Figure 6" captionStartId="8.[129,194,1836,1860]" captionTargetBox="[169,1432,146,1806]" captionTargetId="figure-8@8.[168,1434,144,1808]" captionTargetPageId="8" captionText="Figure 6. Chersina langebaanwegi, paratype SAM-PQL-PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014603" httpUri="https://zenodo.org/record/15014603/files/figure.png" pageId="3" pageNumber="4">6D, E</figureCitation> , -<figureCitation id="13B72A0EF233FFD0FAB0F871FA44DA82" box="[1356,1412,1935,1959]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." pageId="3" pageNumber="4">8A, B</figureCitation> +<figureCitation id="13B72A0EF233FFD0FAB0F871FA44DA82" box="[1356,1412,1935,1959]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014609" httpUri="https://zenodo.org/record/15014609/files/figure.png" pageId="3" pageNumber="4">8A, B</figureCitation> , -<figureCitation id="13B72A0EF233FFD7FA72F871FF53DB01" captionStart="Figure 9" captionStartId="11.[113,178,1714,1738]" captionTargetBox="[131,1423,182,1679]" captionTargetId="figure-113@11.[114,1458,144,1687]" captionTargetPageId="11" captionText="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">9C, D</figureCitation> +<figureCitation id="13B72A0EF233FFD7FA72F871FF53DB01" captionStart="Figure 9" captionStartId="11.[113,178,1714,1738]" captionTargetBox="[131,1423,182,1679]" captionTargetId="figure-113@11.[114,1458,144,1687]" captionTargetPageId="11" captionText="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014611" httpUri="https://zenodo.org/record/15014611/files/figure.png" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">9C, D</figureCitation> , -<figureCitation id="13B72A0EF234FFD7FF61F9F0FF77DB02" box="[157,183,1550,1574]" captionStart="Figure 10" captionStartId="12.[129,194,1334,1358]" captionTargetBox="[132,1471,147,1304]" captionTargetId="figure-312@12.[129,1473,144,1306]" captionTargetPageId="12" captionText="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." pageId="4" pageNumber="5">10</figureCitation> +<figureCitation id="13B72A0EF234FFD7FF61F9F0FF77DB02" box="[157,183,1550,1574]" captionStart="Figure 10" captionStartId="12.[129,194,1334,1358]" captionTargetBox="[132,1471,147,1304]" captionTargetId="figure-312@12.[129,1473,144,1306]" captionTargetPageId="12" captionText="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." figureDoi="http://doi.org/10.5281/zenodo.15014613" httpUri="https://zenodo.org/record/15014613/files/figure.png" pageId="4" pageNumber="5">10</figureCitation> ; Supporting information, -<figureCitation id="13B72A0EF234FFD7FE43F9F0FDC4DB02" box="[447,516,1550,1574]" captionStart="Figure 7" captionStartId="9.[114,179,1706,1730]" captionTargetBox="[115,1436,145,1676]" captionTargetId="figure-121@9.[114,1458,144,1678]" captionTargetPageId="9" captionText="Figure 7. Chersina langebaanwegi, paratype SAM-PQL-21749: nuchal in external view (A); pygal in external (B) and internal (C) views; anterior portion of the plastron (and part of the nuchal) in ventral view (D); xiphiplastra in ventral (E) and dorsal (F) views. Scale bars equal 7 cm in (A, D) and 5 cm in (B, C, E, F)." pageId="4" pageNumber="5">Fig. S7</figureCitation> +<figureCitation id="13B72A0EF234FFD7FE43F9F0FDC4DB02" box="[447,516,1550,1574]" captionStart="Figure 7" captionStartId="9.[114,179,1706,1730]" captionTargetBox="[115,1436,145,1676]" captionTargetId="figure-121@9.[114,1458,144,1678]" captionTargetPageId="9" captionText="Figure 7. Chersina langebaanwegi, paratype SAM-PQL-21749: nuchal in external view (A); pygal in external (B) and internal (C) views; anterior portion of the plastron (and part of the nuchal) in ventral view (D); xiphiplastra in ventral (E) and dorsal (F) views. Scale bars equal 7 cm in (A, D) and 5 cm in (B, C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.15014607" httpUri="https://zenodo.org/record/15014607/files/figure.png" pageId="4" pageNumber="5">Fig. S7</figureCitation> ); the dorsal surface of the epiplastral lips is nearly flat or very slightly convex and not vari - ably concave as in <taxonomicName id="4C8C4D08F234FFD7FEC0F9B3FE6EDB40" box="[316,430,1612,1636]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="angulata"> <emphasis id="B9F8EA99F234FFD7FEC0F9B3FE6EDB40" box="[316,430,1612,1636]" italics="true" pageId="4" pageNumber="5">C. angulata</emphasis> </taxonomicName> ( -<figureCitation id="13B72A0EF234FFD7FE41F9B2FDC3DB41" box="[445,515,1612,1637]" captionStart="Figure 10" captionStartId="12.[129,194,1334,1358]" captionTargetBox="[132,1471,147,1304]" captionTargetId="figure-312@12.[129,1473,144,1306]" captionTargetPageId="12" captionText="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." pageId="4" pageNumber="5">Fig. 10</figureCitation> +<figureCitation id="13B72A0EF234FFD7FE41F9B2FDC3DB41" box="[445,515,1612,1637]" captionStart="Figure 10" captionStartId="12.[129,194,1334,1358]" captionTargetBox="[132,1471,147,1304]" captionTargetId="figure-312@12.[129,1473,144,1306]" captionTargetPageId="12" captionText="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." figureDoi="http://doi.org/10.5281/zenodo.15014613" httpUri="https://zenodo.org/record/15014613/files/figure.png" pageId="4" pageNumber="5">Fig. 10</figureCitation> ; Supporting information, -<figureCitation id="13B72A0EF234FFD7FF7DF992FF06DBA0" box="[129,198,1644,1668]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." pageId="4" pageNumber="5">Fig. S8</figureCitation> +<figureCitation id="13B72A0EF234FFD7FF7DF992FF06DBA0" box="[129,198,1644,1668]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014609" httpUri="https://zenodo.org/record/15014609/files/figure.png" pageId="4" pageNumber="5">Fig. S8</figureCitation> ); the epiplastral lips in most of the cases end posteriorly with a nearly straight or moderately convex profile (in most cases not producing any significant pocket) and not with a very strong convexity as in <taxonomicName id="4C8C4D08F234FFD7FEE5F934FE4ADBC6" box="[281,394,1738,1762]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="angulata"> <emphasis id="B9F8EA99F234FFD7FEE5F934FE4ADBC6" box="[281,394,1738,1762]" italics="true" pageId="4" pageNumber="5">C. angulata</emphasis> </taxonomicName> (which regularly has a very deep gular pocket) ( -<figureCitation id="13B72A0EF234FFD7FF1DF917FEECDA25" box="[225,300,1769,1793]" captionStart="Figure 3" captionStartId="5.[113,178,1047,1071]" captionTargetBox="[147,1424,145,1017]" captionTargetId="figure-538@5.[146,1426,144,1019]" captionTargetPageId="5" captionText="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." pageId="4" pageNumber="5">Figs 3B</figureCitation> +<figureCitation id="13B72A0EF234FFD7FF1DF917FEECDA25" box="[225,300,1769,1793]" captionStart="Figure 3" captionStartId="5.[113,178,1047,1071]" captionTargetBox="[147,1424,145,1017]" captionTargetId="figure-538@5.[146,1426,144,1019]" captionTargetPageId="5" captionText="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." figureDoi="http://doi.org/10.5281/zenodo.15014597" httpUri="https://zenodo.org/record/15014597/files/figure.png" pageId="4" pageNumber="5">Figs 3B</figureCitation> , -<figureCitation id="13B72A0EF234FFD7FEC4F917FE96DA25" box="[312,342,1769,1793]" captionStart="Figure 5" captionStartId="7.[113,178,1626,1650]" captionTargetBox="[115,1456,145,1596]" captionTargetId="figure-177@7.[114,1458,144,1598]" captionTargetPageId="7" captionText="Figure 5. Chersina langebaanwegi, paratype SAM-PQL-21109, male: pygal area in posterior view (A); posterior portion of the shell in right lateral view (B); anterior lobe of the plastron in ventral and dorsal views (C, D); posterior lobe in ventral and dorsal views (E,F). Scale bars equal 5 cm." pageId="4" pageNumber="5">5D</figureCitation> +<figureCitation id="13B72A0EF234FFD7FEC4F917FE96DA25" box="[312,342,1769,1793]" captionStart="Figure 5" captionStartId="7.[113,178,1626,1650]" captionTargetBox="[115,1456,145,1596]" captionTargetId="figure-177@7.[114,1458,144,1598]" captionTargetPageId="7" captionText="Figure 5. Chersina langebaanwegi, paratype SAM-PQL-21109, male: pygal area in posterior view (A); posterior portion of the shell in right lateral view (B); anterior lobe of the plastron in ventral and dorsal views (C, D); posterior lobe in ventral and dorsal views (E,F). Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014601" httpUri="https://zenodo.org/record/15014601/files/figure.png" pageId="4" pageNumber="5">5D</figureCitation> , -<figureCitation id="13B72A0EF234FFD7FE9EF917FE41DA25" box="[354,385,1769,1793]" captionStart="Figure 6" captionStartId="8.[129,194,1836,1860]" captionTargetBox="[169,1432,146,1806]" captionTargetId="figure-8@8.[168,1434,144,1808]" captionTargetPageId="8" captionText="Figure 6. Chersina langebaanwegi, paratype SAM-PQL-PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm." pageId="4" pageNumber="5">6D</figureCitation> +<figureCitation id="13B72A0EF234FFD7FE9EF917FE41DA25" box="[354,385,1769,1793]" captionStart="Figure 6" captionStartId="8.[129,194,1836,1860]" captionTargetBox="[169,1432,146,1806]" captionTargetId="figure-8@8.[168,1434,144,1808]" captionTargetPageId="8" captionText="Figure 6. Chersina langebaanwegi, paratype SAM-PQL-PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014603" httpUri="https://zenodo.org/record/15014603/files/figure.png" pageId="4" pageNumber="5">6D</figureCitation> , -<figureCitation id="13B72A0EF234FFD7FE71F917FE69DA25" box="[397,425,1769,1793]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." pageId="4" pageNumber="5">8B</figureCitation> +<figureCitation id="13B72A0EF234FFD7FE71F917FE69DA25" box="[397,425,1769,1793]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014609" httpUri="https://zenodo.org/record/15014609/files/figure.png" pageId="4" pageNumber="5">8B</figureCitation> , -<figureCitation id="13B72A0EF234FFD7FE48F917FE13DA25" box="[436,467,1769,1793]" captionStart="Figure 9" captionStartId="11.[113,178,1714,1738]" captionTargetBox="[131,1423,182,1679]" captionTargetId="figure-113@11.[114,1458,144,1687]" captionTargetPageId="11" captionText="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." pageId="4" pageNumber="5">9D</figureCitation> +<figureCitation id="13B72A0EF234FFD7FE48F917FE13DA25" box="[436,467,1769,1793]" captionStart="Figure 9" captionStartId="11.[113,178,1714,1738]" captionTargetBox="[131,1423,182,1679]" captionTargetId="figure-113@11.[114,1458,144,1687]" captionTargetPageId="11" captionText="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014611" httpUri="https://zenodo.org/record/15014611/files/figure.png" pageId="4" pageNumber="5">9D</figureCitation> , -<figureCitation id="13B72A0EF234FFD7FE23F917FDE9DA25" box="[479,553,1769,1793]" captionStart="Figure 10" captionStartId="12.[129,194,1334,1358]" captionTargetBox="[132,1471,147,1304]" captionTargetId="figure-312@12.[129,1473,144,1306]" captionTargetPageId="12" captionText="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." pageId="4" pageNumber="5">10A, D</figureCitation> +<figureCitation id="13B72A0EF234FFD7FE23F917FDE9DA25" box="[479,553,1769,1793]" captionStart="Figure 10" captionStartId="12.[129,194,1334,1358]" captionTargetBox="[132,1471,147,1304]" captionTargetId="figure-312@12.[129,1473,144,1306]" captionTargetPageId="12" captionText="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." figureDoi="http://doi.org/10.5281/zenodo.15014613" httpUri="https://zenodo.org/record/15014613/files/figure.png" pageId="4" pageNumber="5">10A, D</figureCitation> ; Supporting informa - tion, -<figureCitation id="13B72A0EF234FFD7FF46F8F6FEC4DA04" box="[186,260,1800,1824]" captionStart="Figure 9" captionStartId="11.[113,178,1714,1738]" captionTargetBox="[131,1423,182,1679]" captionTargetId="figure-113@11.[114,1458,144,1687]" captionTargetPageId="11" captionText="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." pageId="4" pageNumber="5">Fig. S9</figureCitation> +<figureCitation id="13B72A0EF234FFD7FF46F8F6FEC4DA04" box="[186,260,1800,1824]" captionStart="Figure 9" captionStartId="11.[113,178,1714,1738]" captionTargetBox="[131,1423,182,1679]" captionTargetId="figure-113@11.[114,1458,144,1687]" captionTargetPageId="11" captionText="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014611" httpUri="https://zenodo.org/record/15014611/files/figure.png" pageId="4" pageNumber="5">Fig. S9</figureCitation> ); the entoplastron is wider than long instead of being longer than wide, or circular, as in <taxonomicName id="4C8C4D08F234FFD7FDEDF8D6FD42DA64" box="[529,642,1832,1856]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="angulata"> <emphasis id="B9F8EA99F234FFD7FDEDF8D6FD42DA64" box="[529,642,1832,1856]" italics="true" pageId="4" pageNumber="5">C. angulata</emphasis> </taxonomicName> ( -<figureCitation id="13B72A0EF234FFD7FD6DF8D6FD1EDA64" box="[657,734,1832,1856]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="4" pageNumber="5">Figs 1D</figureCitation> +<figureCitation id="13B72A0EF234FFD7FD6DF8D6FD1EDA64" box="[657,734,1832,1856]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014591" httpUri="https://zenodo.org/record/15014591/files/figure.png" pageId="4" pageNumber="5">Figs 1D</figureCitation> , -<figureCitation id="13B72A0EF234FFD7FD14F8D6FCC4DA64" box="[744,772,1832,1856]" captionStart="Figure 3" captionStartId="5.[113,178,1047,1071]" captionTargetBox="[147,1424,145,1017]" captionTargetId="figure-538@5.[146,1426,144,1019]" captionTargetPageId="5" captionText="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." pageId="4" pageNumber="5">3B</figureCitation> +<figureCitation id="13B72A0EF234FFD7FD14F8D6FCC4DA64" box="[744,772,1832,1856]" captionStart="Figure 3" captionStartId="5.[113,178,1047,1071]" captionTargetBox="[147,1424,145,1017]" captionTargetId="figure-538@5.[146,1426,144,1019]" captionTargetPageId="5" captionText="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." figureDoi="http://doi.org/10.5281/zenodo.15014597" httpUri="https://zenodo.org/record/15014597/files/figure.png" pageId="4" pageNumber="5">3B</figureCitation> , -<figureCitation id="13B72A0EF234FFD7FF7CF8B9FF79DA7A" box="[128,185,1863,1887]" captionStart="Figure 4" captionStartId="6.[129,194,1420,1444]" captionTargetBox="[131,1471,146,1390]" captionTargetId="figure-285@6.[129,1473,144,1392]" captionTargetPageId="6" captionText="Figure 4. Chersina langebaanwegi, paratype SAM-PQL-20925: male plastron in ventral view (A) and detail of the anterior lobe of the plastron in dorsal view (B); posterior lobe of the plastron in ventral view (C); pygal area in posterior view (D). Scale bars equal 5 cm." pageId="4" pageNumber="5">4A, B</figureCitation> +<figureCitation id="13B72A0EF234FFD7FF7CF8B9FF79DA7A" box="[128,185,1863,1887]" captionStart="Figure 4" captionStartId="6.[129,194,1420,1444]" captionTargetBox="[131,1471,146,1390]" captionTargetId="figure-285@6.[129,1473,144,1392]" captionTargetPageId="6" captionText="Figure 4. Chersina langebaanwegi, paratype SAM-PQL-20925: male plastron in ventral view (A) and detail of the anterior lobe of the plastron in dorsal view (B); posterior lobe of the plastron in ventral view (C); pygal area in posterior view (D). Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014599" httpUri="https://zenodo.org/record/15014599/files/figure.png" pageId="4" pageNumber="5">4A, B</figureCitation> , -<figureCitation id="13B72A0EF234FFD7FF38F8B9FEC0DA7A" box="[196,256,1863,1887]" captionStart="Figure 5" captionStartId="7.[113,178,1626,1650]" captionTargetBox="[115,1456,145,1596]" captionTargetId="figure-177@7.[114,1458,144,1598]" captionTargetPageId="7" captionText="Figure 5. Chersina langebaanwegi, paratype SAM-PQL-21109, male: pygal area in posterior view (A); posterior portion of the shell in right lateral view (B); anterior lobe of the plastron in ventral and dorsal views (C, D); posterior lobe in ventral and dorsal views (E,F). Scale bars equal 5 cm." pageId="4" pageNumber="5">5C, D</figureCitation> +<figureCitation id="13B72A0EF234FFD7FF38F8B9FEC0DA7A" box="[196,256,1863,1887]" captionStart="Figure 5" captionStartId="7.[113,178,1626,1650]" captionTargetBox="[115,1456,145,1596]" captionTargetId="figure-177@7.[114,1458,144,1598]" captionTargetPageId="7" captionText="Figure 5. Chersina langebaanwegi, paratype SAM-PQL-21109, male: pygal area in posterior view (A); posterior portion of the shell in right lateral view (B); anterior lobe of the plastron in ventral and dorsal views (C, D); posterior lobe in ventral and dorsal views (E,F). Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014601" httpUri="https://zenodo.org/record/15014601/files/figure.png" pageId="4" pageNumber="5">5C, D</figureCitation> , -<figureCitation id="13B72A0EF234FFD7FEF7F8B9FE83DA7A" box="[267,323,1863,1887]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." pageId="4" pageNumber="5">8A, B</figureCitation> +<figureCitation id="13B72A0EF234FFD7FEF7F8B9FE83DA7A" box="[267,323,1863,1887]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014609" httpUri="https://zenodo.org/record/15014609/files/figure.png" pageId="4" pageNumber="5">8A, B</figureCitation> , -<figureCitation id="13B72A0EF234FFD7FEB2F8B9FE4BDA7A" box="[334,395,1863,1887]" captionStart="Figure 9" captionStartId="11.[113,178,1714,1738]" captionTargetBox="[131,1423,182,1679]" captionTargetId="figure-113@11.[114,1458,144,1687]" captionTargetPageId="11" captionText="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." pageId="4" pageNumber="5">9C, D</figureCitation> +<figureCitation id="13B72A0EF234FFD7FEB2F8B9FE4BDA7A" box="[334,395,1863,1887]" captionStart="Figure 9" captionStartId="11.[113,178,1714,1738]" captionTargetBox="[131,1423,182,1679]" captionTargetId="figure-113@11.[114,1458,144,1687]" captionTargetPageId="11" captionText="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014611" httpUri="https://zenodo.org/record/15014611/files/figure.png" pageId="4" pageNumber="5">9C, D</figureCitation> ; Supporting information, -<figureCitation id="13B72A0EF234FFD7FD6BF8B9FD29DA7B" box="[663,745,1863,1887]" captionStart="Figure 10" captionStartId="12.[129,194,1334,1358]" captionTargetBox="[132,1471,147,1304]" captionTargetId="figure-312@12.[129,1473,144,1306]" captionTargetPageId="12" captionText="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." pageId="4" pageNumber="5">Fig. S10</figureCitation> +<figureCitation id="13B72A0EF234FFD7FD6BF8B9FD29DA7B" box="[663,745,1863,1887]" captionStart="Figure 10" captionStartId="12.[129,194,1334,1358]" captionTargetBox="[132,1471,147,1304]" captionTargetId="figure-312@12.[129,1473,144,1306]" captionTargetPageId="12" captionText="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." figureDoi="http://doi.org/10.5281/zenodo.15014613" httpUri="https://zenodo.org/record/15014613/files/figure.png" pageId="4" pageNumber="5">Fig. S10</figureCitation> ). </paragraph> </subSubSection> -<caption id="DFF36603F234FFD7FF7DFA8CFF36D8F9" pageId="4" pageNumber="5" startId="4.[129,194,1394,1418]" targetBox="[130,1471,145,1364]" targetPageId="4" targetType="figure"> +<caption id="DFF36603F234FFD7FF7DFA8CFF36D8F9" ID-DOI="http://doi.org/10.5281/zenodo.15014595" ID-Zenodo-Dep="15014595" httpUri="https://zenodo.org/record/15014595/files/figure.png" pageId="4" pageNumber="5" startId="4.[129,194,1394,1418]" targetBox="[130,1471,145,1364]" targetPageId="4" targetType="figure"> <paragraph id="8B33368BF234FFD7FF7DFA8CFF36D8F9" blockId="4.[129,1468,1394,1502]" pageId="4" pageNumber="5"> <emphasis id="B9F8EA99F234FFD7FF7DFA8CFF18D8AE" bold="true" box="[129,216,1394,1418]" pageId="4" pageNumber="5">Figure 2.</emphasis> <taxonomicName id="4C8C4D08F234FFD7FF21FA8CFE76D8AE" box="[221,438,1394,1418]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="langebaanwegi"> @@ -825,7 +828,7 @@ regarding the following characters (see Supporting information for the compariso If not otherwise stated, the following descrip - tion is mostly based on the <typeStatus id="54378829F234FFD7FE58F85EFDC5DA9C" box="[420,517,1952,1976]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus> SAM -PQL -20734 ( -<figureCitation id="13B72A0EF234FFD7FD22F85EFCA2DB02" box="[734,866,1550,1976]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="3.[113,178,1312,1336]" captionStartId-1="4.[129,194,1394,1418]" captionStartId-2="5.[113,178,1047,1071]" captionTargetBox-0="[117,1455,147,1281]" captionTargetBox-1="[130,1471,145,1364]" captionTargetBox-2="[147,1424,145,1017]" captionTargetId-0="figure-340@3.[114,1458,144,1284]" captionTargetId-1="figure-300@4.[129,1473,144,1366]" captionTargetId-2="figure-538@5.[146,1426,144,1019]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." captionText-1="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." captionText-2="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." pageId="4" pageNumber="5">Figs 1–3</figureCitation> +<figureCitation id="13B72A0EF234FFD7FD22F85EFCA2DB02" box="[734,866,1550,1976]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="3.[113,178,1312,1336]" captionStartId-1="4.[129,194,1394,1418]" captionStartId-2="5.[113,178,1047,1071]" captionTargetBox-0="[117,1455,147,1281]" captionTargetBox-1="[130,1471,145,1364]" captionTargetBox-2="[147,1424,145,1017]" captionTargetId-0="figure-340@3.[114,1458,144,1284]" captionTargetId-1="figure-300@4.[129,1473,144,1366]" captionTargetId-2="figure-538@5.[146,1426,144,1019]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." captionText-1="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." captionText-2="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." figureDoi-0="http://doi.org/10.5281/zenodo.15014591" figureDoi-1="http://doi.org/10.5281/zenodo.15014595" figureDoi-2="http://doi.org/10.5281/zenodo.15014597" httpUri-0="https://zenodo.org/record/15014591/files/figure.png" httpUri-1="https://zenodo.org/record/15014595/files/figure.png" httpUri-2="https://zenodo.org/record/15014597/files/figure.png" pageId="4" pageNumber="5">Figs 1–3</figureCitation> ) as it is the only nearly complete and mounted shell so far available. There are no cranial, axial, or appendicular elements associated with the <typeStatus id="54378829F234FFD7FB69F9B2FB36DB40" box="[1173,1270,1612,1636]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus> and the referred specimens. @@ -841,24 +844,24 @@ The nearly complete female shell SAM -PQL -20734 ( , width 15.7, height <quantity id="4C749B6EF234FFD7FB43F8FBFAE2DA39" box="[1215,1314,1797,1821]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="4" pageNumber="5" unit="mm" value="14.7">14.7 mm</quantity> ), with a nearly smooth dorsal profile both in lateral and anterior/posterior views. The posterior border is relatively curved inwards and not significantly flared. Very weak bosses are visible in lateral view ( -<figureCitation id="13B72A0EF234FFD7FCB8F87CFC71DABE" box="[836,945,1922,1946]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="4" pageNumber="5">Fig. 1C, D</figureCitation> +<figureCitation id="13B72A0EF234FFD7FCB8F87CFC71DABE" box="[836,945,1922,1946]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014591" httpUri="https://zenodo.org/record/15014591/files/figure.png" pageId="4" pageNumber="5">Fig. 1C, D</figureCitation> ) that correspond to five vertebral scutes. In dorsal view, the anterior edge of the shell ( -<figureCitation id="13B72A0EF234FFD7FB46F85CFAE4DA9D" box="[1210,1316,1954,1978]" captionStart="Figure 2" captionStartId="4.[129,194,1394,1418]" captionTargetBox="[130,1471,145,1364]" captionTargetId="figure-300@4.[129,1473,144,1366]" captionTargetPageId="4" captionText="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="4" pageNumber="5">Fig. 2A, B</figureCitation> +<figureCitation id="13B72A0EF234FFD7FB46F85CFAE4DA9D" box="[1210,1316,1954,1978]" captionStart="Figure 2" captionStartId="4.[129,194,1394,1418]" captionTargetBox="[130,1471,145,1364]" captionTargetId="figure-300@4.[129,1473,144,1366]" captionTargetPageId="4" captionText="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014595" httpUri="https://zenodo.org/record/15014595/files/figure.png" pageId="4" pageNumber="5">Fig. 2A, B</figureCitation> ) has a modest notch laterally delimited by the small protrusion of the first peripherals (corresponding to the sulcus between the first and second marginal) and, moreover, the central area of the carapace, corresponding to the bridge, is somewhat swollen if compared to the areas anterior and posterior to it. The anterior opening is kidney shaped because the anterior lobe of the plastron is ele - vated in the dorsal direction ( -<figureCitation id="13B72A0EF235FFD6FE66FAC9FE22D86B" box="[410,482,1335,1359]" captionStart="Figure 3" captionStartId="5.[113,178,1047,1071]" captionTargetBox="[147,1424,145,1017]" captionTargetId="figure-538@5.[146,1426,144,1019]" captionTargetPageId="5" captionText="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." pageId="5" pageNumber="6">Fig. 3A</figureCitation> +<figureCitation id="13B72A0EF235FFD6FE66FAC9FE22D86B" box="[410,482,1335,1359]" captionStart="Figure 3" captionStartId="5.[113,178,1047,1071]" captionTargetBox="[147,1424,145,1017]" captionTargetId="figure-538@5.[146,1426,144,1019]" captionTargetPageId="5" captionText="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." figureDoi="http://doi.org/10.5281/zenodo.15014597" httpUri="https://zenodo.org/record/15014597/files/figure.png" pageId="5" pageNumber="6">Fig. 3A</figureCitation> ; see description of the plas - tron). The posterior opening is rather broad ( -<figureCitation id="13B72A0EF235FFD6FDAFFAA8FD01D84A" box="[595,705,1366,1390]" captionStart="Figure 2" captionStartId="4.[129,194,1394,1418]" captionTargetBox="[130,1471,145,1364]" captionTargetId="figure-300@4.[129,1473,144,1366]" captionTargetPageId="4" captionText="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="5" pageNumber="6">Fig. 2C, D</figureCitation> +<figureCitation id="13B72A0EF235FFD6FDAFFAA8FD01D84A" box="[595,705,1366,1390]" captionStart="Figure 2" captionStartId="4.[129,194,1394,1418]" captionTargetBox="[130,1471,145,1364]" captionTargetId="figure-300@4.[129,1473,144,1366]" captionTargetPageId="4" captionText="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014595" httpUri="https://zenodo.org/record/15014595/files/figure.png" pageId="5" pageNumber="6">Fig. 2C, D</figureCitation> ) and markedly much broader than in extant <taxonomicName id="4C8C4D08F235FFD6FDF3FA8BFD05D8A9" box="[527,709,1397,1421]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="angulata"> <emphasis id="B9F8EA99F235FFD6FDF3FA8BFD05D8A9" box="[527,709,1397,1421]" italics="true" pageId="5" pageNumber="6">Chersina angulata</emphasis> </taxonomicName> . The carapace has no hinge and moreover there is no hinge between the carapace and plastron [as also indicated, among other, by the partial plastron SAM -PQL -20925 ( -<figureCitation id="13B72A0EF235FFD6FE19FA2DFDE1D8C8" box="[485,545,1491,1516]" captionStart="Figure 4" captionStartId="6.[129,194,1420,1444]" captionTargetBox="[131,1471,146,1390]" captionTargetId="figure-285@6.[129,1473,144,1392]" captionTargetPageId="6" captionText="Figure 4. Chersina langebaanwegi, paratype SAM-PQL-20925: male plastron in ventral view (A) and detail of the anterior lobe of the plastron in dorsal view (B); posterior lobe of the plastron in ventral view (C); pygal area in posterior view (D). Scale bars equal 5 cm." pageId="5" pageNumber="6">Fig. 4</figureCitation> +<figureCitation id="13B72A0EF235FFD6FE19FA2DFDE1D8C8" box="[485,545,1491,1516]" captionStart="Figure 4" captionStartId="6.[129,194,1420,1444]" captionTargetBox="[131,1471,146,1390]" captionTargetId="figure-285@6.[129,1473,144,1392]" captionTargetPageId="6" captionText="Figure 4. Chersina langebaanwegi, paratype SAM-PQL-20925: male plastron in ventral view (A) and detail of the anterior lobe of the plastron in dorsal view (B); posterior lobe of the plastron in ventral view (C); pygal area in posterior view (D). Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014599" httpUri="https://zenodo.org/record/15014599/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation> ) that is still sutured with some bridge peripherals] or within the plastron (see also SAM -PQL -22076— -<figureCitation id="13B72A0EF235FFD6FEB9F9ECFEBDDB0E" box="[325,381,1554,1578]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." pageId="5" pageNumber="6">Fig. 8</figureCitation> +<figureCitation id="13B72A0EF235FFD6FEB9F9ECFEBDDB0E" box="[325,381,1554,1578]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014609" httpUri="https://zenodo.org/record/15014609/files/figure.png" pageId="5" pageNumber="6">Fig. 8</figureCitation> ). </paragraph> -<caption id="DFF36603F235FFD6FF8DFBE9FA48D96E" pageId="5" pageNumber="6" startId="5.[113,178,1047,1071]" targetBox="[147,1424,145,1017]" targetPageId="5" targetType="figure"> +<caption id="DFF36603F235FFD6FF8DFBE9FA48D96E" ID-DOI="http://doi.org/10.5281/zenodo.15014597" ID-Zenodo-Dep="15014597" httpUri="https://zenodo.org/record/15014597/files/figure.png" pageId="5" pageNumber="6" startId="5.[113,178,1047,1071]" targetBox="[147,1424,145,1017]" targetPageId="5" targetType="figure"> <paragraph id="8B33368BF235FFD6FF8DFBE9FA48D96E" blockId="5.[113,1432,1047,1099]" pageId="5" pageNumber="6"> <emphasis id="B9F8EA99F235FFD6FF8DFBE9FF09D90B" bold="true" box="[113,201,1047,1071]" pageId="5" pageNumber="6">Figure 3.</emphasis> <taxonomicName id="4C8C4D08F235FFD6FF32FBE9FE67D90B" box="[206,423,1047,1071]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="langebaanwegi"> @@ -884,17 +887,17 @@ SAM -PQL -20734), with borders that can be nearly straight (SAM -PQL -21174) or <emphasis id="B9F8EA99F235FFD6FCBAFB44FC7BD9F5" box="[838,955,1209,1233]" italics="true" pageId="5" pageNumber="6">C. angulata</emphasis> </taxonomicName> that develops a sort of dorsal shelf (Supporting information, -<figureCitation id="13B72A0EF235FFD6FC4DFB27FBE3D9D4" box="[945,1059,1241,1265]" captionStart="Figure 3" captionStartId="5.[113,178,1047,1071]" captionTargetBox="[147,1424,145,1017]" captionTargetId="figure-538@5.[146,1426,144,1019]" captionTargetPageId="5" captionText="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." pageId="5" pageNumber="6">Fig. S3A, B</figureCitation> +<figureCitation id="13B72A0EF235FFD6FC4DFB27FBE3D9D4" box="[945,1059,1241,1265]" captionStart="Figure 3" captionStartId="5.[113,178,1047,1071]" captionTargetBox="[147,1424,145,1017]" captionTargetId="figure-538@5.[146,1426,144,1019]" captionTargetPageId="5" captionText="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." figureDoi="http://doi.org/10.5281/zenodo.15014597" httpUri="https://zenodo.org/record/15014597/files/figure.png" pageId="5" pageNumber="6">Fig. S3A, B</figureCitation> ). </paragraph> <paragraph id="8B33368BF235FFD5FCB9FB06FE75DA64" blockId="5.[810,1459,1147,1954]" lastBlockId="6.[129,777,1520,1951]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6"> There are eight neural elements and the neural formula is: 6,8,4,8,4,6A,4,6A. The first neural of SAM -PQL -20734 is anteroposteriorly elongated and hexagonal with anterolateral sides approximately equal to the posterolateral ones. Neurals from two to seven are wider than long (in absence of the proper state, the third has been scored as square but it is in fact rect - angular). The eighth is longer than wide and, at least on the better -preserved right side, it seems to be hexagonal (with a short anterolateral side). On the visceral surface of the neurals from two to five there is clear evidence for the emergence of very thin costal processes from the neural and not from the corres - ponding costals. In addition, the sixth neural may show the same condition as indicated by a small irregularity located close to the posterior border (and by the absence of a costal process on the corresponding costal). The costovertebral tunnel is, therefore, very narrow (much narrower than the width of the neurals). The two joined neurals of SAM -PQL -35023d ( -<figureCitation id="13B72A0EF235FFD6FB13F913FAEBDA21" box="[1263,1323,1773,1797]" captionStart="Figure 9" captionStartId="11.[113,178,1714,1738]" captionTargetBox="[131,1423,182,1679]" captionTargetId="figure-113@11.[114,1458,144,1687]" captionTargetPageId="11" captionText="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." pageId="5" pageNumber="6">Fig. 9</figureCitation> +<figureCitation id="13B72A0EF235FFD6FB13F913FAEBDA21" box="[1263,1323,1773,1797]" captionStart="Figure 9" captionStartId="11.[113,178,1714,1738]" captionTargetBox="[131,1423,182,1679]" captionTargetId="figure-113@11.[114,1458,144,1687]" captionTargetPageId="11" captionText="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014611" httpUri="https://zenodo.org/record/15014611/files/figure.png" pageId="5" pageNumber="6">Fig. 9</figureCitation> ) have a sort of tubercle on the visceral surface close to the lateral edge. The preservation of the ventral surface of the isolated neural of SAM - PQL -21174 does not allow confirmation of this morphology in another specimen but its irregular surface could indicate the presence of costal processes ( -<figureCitation id="13B72A0EF235FFD6FB9BF874FB64DA86" box="[1127,1188,1930,1954]" captionStart="Figure 6" captionStartId="8.[129,194,1836,1860]" captionTargetBox="[169,1432,146,1806]" captionTargetId="figure-8@8.[168,1434,144,1808]" captionTargetPageId="8" captionText="Figure 6. Chersina langebaanwegi, paratype SAM-PQL-PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm." pageId="5" pageNumber="6">Fig. 6</figureCitation> +<figureCitation id="13B72A0EF235FFD6FB9BF874FB64DA86" box="[1127,1188,1930,1954]" captionStart="Figure 6" captionStartId="8.[129,194,1836,1860]" captionTargetBox="[169,1432,146,1806]" captionTargetId="figure-8@8.[168,1434,144,1808]" captionTargetPageId="8" captionText="Figure 6. Chersina langebaanwegi, paratype SAM-PQL-PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014603" httpUri="https://zenodo.org/record/15014603/files/figure.png" pageId="5" pageNumber="6">Fig. 6</figureCitation> ). Noteworthy is that the three preserved costals of SAM -PQL -21109 do not have a costal process on the medial area of their visceral surface. There is only one suprapygal that is trapezoidal in shape; its posterior edge is approximately straight. The female pygal of SAM -PQL -20734 is quadrangular and shows a small notch centrally located on the free, ventral edge; it is only moderately convex. The male pygal SAM -PQL -20925 is also quadrangular. The male pygal of SAM - PQL -21109 and likely also the partly preserved large male pygal SAM -PQL -35023d are hexagonal, with short anterolateral sides and long, concave posterolateral sides; the free, ventral edge is broadly concave in both cases. </paragraph> -<caption id="DFF36603F236FFD5FF7DFA72FACED89B" pageId="6" pageNumber="7" startId="6.[129,194,1420,1444]" targetBox="[131,1471,146,1390]" targetPageId="6" targetType="figure"> +<caption id="DFF36603F236FFD5FF7DFA72FACED89B" ID-DOI="http://doi.org/10.5281/zenodo.15014599" ID-Zenodo-Dep="15014599" httpUri="https://zenodo.org/record/15014599/files/figure.png" pageId="6" pageNumber="7" startId="6.[129,194,1420,1444]" targetBox="[131,1471,146,1390]" targetPageId="6" targetType="figure"> <paragraph id="8B33368BF236FFD5FF7DFA72FACED89B" blockId="6.[129,1455,1420,1472]" pageId="6" pageNumber="7"> <emphasis id="B9F8EA99F236FFD5FF7DFA72FF18D880" bold="true" box="[129,216,1420,1444]" pageId="6" pageNumber="7">Figure 4.</emphasis> <taxonomicName id="4C8C4D08F236FFD5FF21FA72FE76D880" box="[221,438,1420,1444]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="langebaanwegi"> @@ -913,7 +916,7 @@ There are 11 peripherals. Those from three to seven are involved in the bridge; <typeStatus id="54378829F236FFD5FCC5F8D7FC53DA65" box="[825,915,1833,1857]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus> SAM -PQL -20734 the first and second peripherals de - velop a small, distal projection corresponding to the intermarginal sulcus, but there are no intermarginal projections or interperipheral serrations in the posterior area. Conversely, in SAM -PQL -21109 there is a marked intermarginal projection on the ninth periph - eral (best preserved on the right side). The third peripheral hosts a small axillary scute that develops on the hyoplastron buttress as well. The seventh peripheral hosts part of the large inguinal scute (that is mostly developed on the hypoplastron). </paragraph> -<caption id="DFF36603F237FFD4FF8DF9A4FF1EDB8E" pageId="7" pageNumber="8" startId="7.[113,178,1626,1650]" targetBox="[115,1456,145,1596]" targetPageId="7" targetType="figure"> +<caption id="DFF36603F237FFD4FF8DF9A4FF1EDB8E" ID-DOI="http://doi.org/10.5281/zenodo.15014601" ID-Zenodo-Dep="15014601" httpUri="https://zenodo.org/record/15014601/files/figure.png" pageId="7" pageNumber="8" startId="7.[113,178,1626,1650]" targetBox="[115,1456,145,1596]" targetPageId="7" targetType="figure"> <paragraph id="8B33368BF237FFD4FF8DF9A4FF1EDB8E" blockId="7.[113,1414,1626,1706]" pageId="7" pageNumber="8"> <emphasis id="B9F8EA99F237FFD4FF8DF9A4FF08DB56" bold="true" box="[113,200,1626,1650]" pageId="7" pageNumber="8">Figure 5.</emphasis> <taxonomicName id="4C8C4D08F237FFD4FF31F9A4FE67DB56" box="[205,423,1626,1650]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="langebaanwegi"> @@ -927,12 +930,12 @@ SAM -PQL -20734 the first and second peripherals de - velop a small, distal proj The cervical scute is rather large on both the dorsal and ventral surface of the nuchal. On the dorsal surface, it can be only slightly longer than broad (in SAM -PQL -20734, respect - ively 15.0 and <quantity id="4C749B6EF237FFD4FC40F907FBD9DA35" box="[956,1049,1785,1809]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="7" pageNumber="8" unit="mm" value="14.3">14.3 mm</quantity> ; -<figureCitation id="13B72A0EF237FFD4FBDFF907FB45DA35" box="[1059,1157,1785,1809]" captionStart="Figure 2" captionStartId="4.[129,194,1394,1418]" captionTargetBox="[130,1471,145,1364]" captionTargetId="figure-300@4.[129,1473,144,1366]" captionTargetPageId="4" captionText="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="7" pageNumber="8">Fig. 2A, B</figureCitation> +<figureCitation id="13B72A0EF237FFD4FBDFF907FB45DA35" box="[1059,1157,1785,1809]" captionStart="Figure 2" captionStartId="4.[129,194,1394,1418]" captionTargetBox="[130,1471,145,1364]" captionTargetId="figure-300@4.[129,1473,144,1366]" captionTargetPageId="4" captionText="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014595" httpUri="https://zenodo.org/record/15014595/files/figure.png" pageId="7" pageNumber="8">Fig. 2A, B</figureCitation> ), and it is slightly longer than 25% of the nuchal. The cervical scute of SAM -PQL -21174 is distinctly longer than broad but its length corresponds to about 25% of the nuchal. On the ventral surface of the nuchal, the cer - vical is very long and has nearly parallel margins. The first pleural scutes slightly overlap the nuchal in SAM -PQL -20734 ( -<figureCitation id="13B72A0EF237FFD4FA9FF868FCF8DAE9" captionStart="Figure 2" captionStartId="4.[129,194,1394,1418]" captionTargetBox="[130,1471,145,1364]" captionTargetId="figure-300@4.[129,1473,144,1366]" captionTargetPageId="4" captionText="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="7" pageNumber="8">Fig. 2A, B</figureCitation> +<figureCitation id="13B72A0EF237FFD4FA9FF868FCF8DAE9" captionStart="Figure 2" captionStartId="4.[129,194,1394,1418]" captionTargetBox="[130,1471,145,1364]" captionTargetId="figure-300@4.[129,1473,144,1366]" captionTargetPageId="4" captionText="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014595" httpUri="https://zenodo.org/record/15014595/files/figure.png" pageId="7" pageNumber="8">Fig. 2A, B</figureCitation> ) even less in SAM -PQL -21174 (where only the left side is well preserved) but not in SAM -PQL -21109 (that also preserves only the left side). </paragraph> -<caption id="DFF36603F238FFDBFF7DF8D2FA45DA7B" pageId="8" pageNumber="9" startId="8.[129,194,1836,1860]" targetBox="[169,1432,146,1806]" targetPageId="8" targetType="figure"> +<caption id="DFF36603F238FFDBFF7DF8D2FA45DA7B" ID-DOI="http://doi.org/10.5281/zenodo.15014603" ID-Zenodo-Dep="15014603" httpUri="https://zenodo.org/record/15014603/files/figure.png" pageId="8" pageNumber="9" startId="8.[129,194,1836,1860]" targetBox="[169,1432,146,1806]" targetPageId="8" targetType="figure"> <paragraph id="8B33368BF238FFDBFF7DF8D2FA45DA7B" blockId="8.[129,1438,1836,1888]" pageId="8" pageNumber="9"> <emphasis id="B9F8EA99F238FFDBFF7DF8D2FF18DA60" bold="true" box="[129,216,1836,1860]" pageId="8" pageNumber="9">Figure 6.</emphasis> <taxonomicName id="4C8C4D08F238FFDBFF21F8D2FE76DA60" box="[221,438,1836,1860]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="langebaanwegi"> @@ -941,7 +944,7 @@ The cervical scute is rather large on both the dorsal and ventral surface of the , paratype SAM -PQL -PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm. </paragraph> </caption> -<caption id="DFF36603F239FFDAFF8EF954FE14DBDE" pageId="9" pageNumber="10" startId="9.[114,179,1706,1730]" targetBox="[115,1436,145,1676]" targetPageId="9" targetType="figure"> +<caption id="DFF36603F239FFDAFF8EF954FE14DBDE" ID-DOI="http://doi.org/10.5281/zenodo.15014607" ID-Zenodo-Dep="15014607" httpUri="https://zenodo.org/record/15014607/files/figure.png" pageId="9" pageNumber="10" startId="9.[114,179,1706,1730]" targetBox="[115,1436,145,1676]" targetPageId="9" targetType="figure"> <paragraph id="8B33368BF239FFDAFF8EF954FE14DBDE" blockId="9.[113,1451,1706,1786]" pageId="9" pageNumber="10"> <emphasis id="B9F8EA99F239FFDAFF8EF954FF0ADBE6" bold="true" box="[114,202,1706,1730]" pageId="9" pageNumber="10">Figure 7.</emphasis> <taxonomicName id="4C8C4D08F239FFDAFF32F954FE68DBE6" box="[206,424,1706,1730]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="langebaanwegi"> @@ -963,10 +966,10 @@ posterior to the suture on the pygal of SAM -PQL -20734), this character was sco <emphasis id="B9F8EA99F23CFFDFFDABFA49FD0BD8EA" box="[599,715,1462,1486]" italics="true" pageId="12" pageNumber="13">C. angulata</emphasis> </taxonomicName> ). The tenth marginal scute does not contact the fifth vertebral scute. The posterior sulcus of the fifth vertebral scute is placed slightly posterior to the suprapygal -pygal suture in SAM -PQL -20734 ( -<figureCitation id="13B72A0EF23CFFDFFF77F9CDFF32DB6F" box="[139,242,1587,1612]" captionStart="Figure 2" captionStartId="4.[129,194,1394,1418]" captionTargetBox="[130,1471,145,1364]" captionTargetId="figure-300@4.[129,1473,144,1366]" captionTargetPageId="4" captionText="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="12" pageNumber="13">Fig. 2C, D</figureCitation> +<figureCitation id="13B72A0EF23CFFDFFF77F9CDFF32DB6F" box="[139,242,1587,1612]" captionStart="Figure 2" captionStartId="4.[129,194,1394,1418]" captionTargetBox="[130,1471,145,1364]" captionTargetId="figure-300@4.[129,1473,144,1366]" captionTargetPageId="4" captionText="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014595" httpUri="https://zenodo.org/record/15014595/files/figure.png" pageId="12" pageNumber="13">Fig. 2C, D</figureCitation> ) and even more posteriorly (therefore, clearly on the pygal) in SAM -PQL -20925, SAM -PQL -21109, and, above all, in the very large male SAM -PQL -35023d. The twelfth marginal scutes are fused in a single supracaudal scute. </paragraph> -<caption id="DFF36603F23AFFD9FF7DF8D1FE4EDA46" pageId="10" pageNumber="11" startId="10.[129,194,1839,1863]" targetBox="[137,1465,146,1809]" targetPageId="10" targetType="figure"> +<caption id="DFF36603F23AFFD9FF7DF8D1FE4EDA46" ID-DOI="http://doi.org/10.5281/zenodo.15014609" ID-Zenodo-Dep="15014609" httpUri="https://zenodo.org/record/15014609/files/figure.png" pageId="10" pageNumber="11" startId="10.[129,194,1839,1863]" targetBox="[137,1465,146,1809]" targetPageId="10" targetType="figure"> <paragraph id="8B33368BF23AFFD9FF7DF8D1FE4EDA46" blockId="10.[129,1453,1839,1891]" pageId="10" pageNumber="11"> <emphasis id="B9F8EA99F23AFFD9FF7DF8D1FF18DA63" bold="true" box="[129,216,1839,1863]" pageId="10" pageNumber="11">Figure 8.</emphasis> <taxonomicName id="4C8C4D08F23AFFD9FF21F8D1FE76DA63" box="[221,438,1839,1863]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="langebaanwegi"> @@ -975,7 +978,7 @@ posterior to the suture on the pygal of SAM -PQL -20734), this character was sco , paratype SAM -PQL -PQL -22076: male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views. Scale bars equal 5 cm. </paragraph> </caption> -<caption id="DFF36603F23BFFD8FF8DF94CFCCCDBC2" pageId="11" pageNumber="12" startId="11.[113,178,1714,1738]" targetBox="[131,1423,182,1679]" targetPageId="11" targetType="figure"> +<caption id="DFF36603F23BFFD8FF8DF94CFCCCDBC2" ID-DOI="http://doi.org/10.5281/zenodo.15014611" ID-Zenodo-Dep="15014611" httpUri="https://zenodo.org/record/15014611/files/figure.png" pageId="11" pageNumber="12" startId="11.[113,178,1714,1738]" targetBox="[131,1423,182,1679]" targetPageId="11" targetType="figure"> <paragraph id="8B33368BF23BFFD8FF8DF94CFCCCDBC2" blockId="11.[113,1454,1714,1767]" pageId="11" pageNumber="12"> <emphasis id="B9F8EA99F23BFFD8FF8DF94CFF09DBEE" bold="true" box="[113,201,1714,1738]" pageId="11" pageNumber="12">Figure 9.</emphasis> <taxonomicName id="4C8C4D08F23BFFD8FF32F94DFE67DBEF" box="[206,423,1715,1739]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="langebaanwegi"> @@ -984,7 +987,7 @@ posterior to the suture on the pygal of SAM -PQL -20734), this character was sco , paratype SAM -PQL -35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm. </paragraph> </caption> -<caption id="DFF36603F23CFFDFFF7DFAC8FC01D8A2" pageId="12" pageNumber="13" startId="12.[129,194,1334,1358]" targetBox="[132,1471,147,1304]" targetPageId="12" targetType="figure"> +<caption id="DFF36603F23CFFDFFF7DFAC8FC01D8A2" ID-DOI="http://doi.org/10.5281/zenodo.15014613" ID-Zenodo-Dep="15014613" httpUri="https://zenodo.org/record/15014613/files/figure.png" pageId="12" pageNumber="13" startId="12.[129,194,1334,1358]" targetBox="[132,1471,147,1304]" targetPageId="12" targetType="figure"> <paragraph id="8B33368BF23CFFDFFF7DFAC8FC01D8A2" blockId="12.[129,1463,1334,1414]" pageId="12" pageNumber="13"> <emphasis id="B9F8EA99F23CFFDFFF7DFAC8FF25D86A" bold="true" box="[129,229,1334,1358]" pageId="12" pageNumber="13">Figure 10.</emphasis> <taxonomicName id="4C8C4D08F23CFFDFFF15FAC8FE03D86A" box="[233,451,1334,1358]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="langebaanwegi"> @@ -1000,7 +1003,7 @@ posterior to the suture on the pygal of SAM -PQL -20734), this character was sco The large ( <quantity id="4C749B6EF23CFFDFFF0FF90EFE90DA23" box="[243,336,1776,1800]" metricMagnitude="-1" metricUnit="m" metricValue="2.93" pageId="12" pageNumber="13" unit="mm" value="293.0">293 mm</quantity> long) and complete male plastron SAM - PQL -22076 ( -<figureCitation id="13B72A0EF23CFFDFFEEDF8F1FE8DDA03" box="[273,333,1807,1831]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." pageId="12" pageNumber="13">Fig. 8</figureCitation> +<figureCitation id="13B72A0EF23CFFDFFEEDF8F1FE8DDA03" box="[273,333,1807,1831]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014609" httpUri="https://zenodo.org/record/15014609/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation> ) provides a unique opportunity to assess the general morphology of this shell portion that otherwise is not entirely preserved in a single specimen. The anterior lobe is somewhat trilobed, longer than the posterior lobe (93 and <quantity id="4C749B6EF23CFFDFFD13F893FF6CDA80" metricMagnitude="-2" metricUnit="m" metricValue="6.8" pageId="12" pageNumber="13" unit="mm" value="68.0">68 mm</quantity> , respectively) and does not host any hinge. In dorsal (or ventral) view, the gularo -humeral sulcus corresponds to a dis - tinct constriction in the lateral profile of the element. Because of the anterodorsal orientation of the hyoplastra, the epiplastra are raised from the horizontal plane and their anterior protru - sion is anteriorly directed and approximately parallel to the hori - zontal plane but about @@ -1012,24 +1015,24 @@ SAM -PQL -20734 pre - serves a nearly complete plastron (only the left hypoplast <paragraph id="8B33368BF23DFFDEFF8DFF17FF0BDF1E" blockId="13.[113,763,233,1979]" pageId="13" pageNumber="14"> <emphasis id="B9F8EA99F23DFFDEFF8DFF17FF28DC25" box="[113,232,233,257]" italics="true" pageId="13" pageNumber="14">Bony plates:</emphasis> The epiplastra are characterized by anterior projec - tions that in males can be very long, accounting in most cases (SAM -PQL -21174; -<figureCitation id="13B72A0EF23DFFDEFEB4FED9FE47DC1B" box="[328,391,295,319]" captionStart="Figure 6" captionStartId="8.[129,194,1836,1860]" captionTargetBox="[169,1432,146,1806]" captionTargetId="figure-8@8.[168,1434,144,1808]" captionTargetPageId="8" captionText="Figure 6. Chersina langebaanwegi, paratype SAM-PQL-PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm." pageId="13" pageNumber="14">Fig. 6</figureCitation> +<figureCitation id="13B72A0EF23DFFDEFEB4FED9FE47DC1B" box="[328,391,295,319]" captionStart="Figure 6" captionStartId="8.[129,194,1836,1860]" captionTargetBox="[169,1432,146,1806]" captionTargetId="figure-8@8.[168,1434,144,1808]" captionTargetPageId="8" captionText="Figure 6. Chersina langebaanwegi, paratype SAM-PQL-PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014603" httpUri="https://zenodo.org/record/15014603/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation> ) for about half of the anteropos - terior length of this element, but in SAM -PQL -35023d they are even longer and represent nearly 80% of the total length (total length <quantity id="4C749B6EF23DFFDEFF40FE7BFECBDCB9" box="[188,267,389,413]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" pageId="13" pageNumber="14" unit="mm" value="95.0">95 mm</quantity> ; projection length <quantity id="4C749B6EF23DFFDEFE2AFE7BFDE5DCB9" box="[470,549,389,413]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="13" pageNumber="14" unit="mm" value="75.0">75 mm</quantity> ; -<figureCitation id="13B72A0EF23DFFDEFDCEFE7BFDAFDCB9" box="[562,623,389,413]" captionStart="Figure 9" captionStartId="11.[113,178,1714,1738]" captionTargetBox="[131,1423,182,1679]" captionTargetId="figure-113@11.[114,1458,144,1687]" captionTargetPageId="11" captionText="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." pageId="13" pageNumber="14">Fig. 9</figureCitation> +<figureCitation id="13B72A0EF23DFFDEFDCEFE7BFDAFDCB9" box="[562,623,389,413]" captionStart="Figure 9" captionStartId="11.[113,178,1714,1738]" captionTargetBox="[131,1423,182,1679]" captionTargetId="figure-113@11.[114,1458,144,1687]" captionTargetPageId="11" captionText="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014611" httpUri="https://zenodo.org/record/15014611/files/figure.png" pageId="13" pageNumber="14">Fig. 9</figureCitation> ). In females, the projection is much less developed as shown by the <typeStatus id="54378829F23DFFDEFD62FE5BFD3ADC99" box="[670,762,421,445]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus> SAM -PQL -20734 and, among the referred materials, by SAM - PQL -73878 (compare male epiplastron SAM -PQL -12750c, -<figureCitation id="13B72A0EF23DFFDEFF8DFDFDFF2EDF3F" box="[113,238,515,539]" captionStart="Figure 10" captionStartId="12.[129,194,1334,1358]" captionTargetBox="[132,1471,147,1304]" captionTargetId="figure-312@12.[129,1473,144,1306]" captionTargetPageId="12" captionText="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." pageId="13" pageNumber="14">Fig. 10A–C</figureCitation> +<figureCitation id="13B72A0EF23DFFDEFF8DFDFDFF2EDF3F" box="[113,238,515,539]" captionStart="Figure 10" captionStartId="12.[129,194,1334,1358]" captionTargetBox="[132,1471,147,1304]" captionTargetId="figure-312@12.[129,1473,144,1306]" captionTargetPageId="12" captionText="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." figureDoi="http://doi.org/10.5281/zenodo.15014613" httpUri="https://zenodo.org/record/15014613/files/figure.png" pageId="13" pageNumber="14">Fig. 10A–C</figureCitation> , with female epiplastron SAM -PQL -73878, -<figureCitation id="13B72A0EF23DFFDEFD28FDFDFF7ADF1E" captionStart="Figure 10" captionStartId="12.[129,194,1334,1358]" captionTargetBox="[132,1471,147,1304]" captionTargetId="figure-312@12.[129,1473,144,1306]" captionTargetPageId="12" captionText="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." pageId="13" pageNumber="14">Fig. 10D–F</figureCitation> +<figureCitation id="13B72A0EF23DFFDEFD28FDFDFF7ADF1E" captionStart="Figure 10" captionStartId="12.[129,194,1334,1358]" captionTargetBox="[132,1471,147,1304]" captionTargetId="figure-312@12.[129,1473,144,1306]" captionTargetPageId="12" captionText="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." figureDoi="http://doi.org/10.5281/zenodo.15014613" httpUri="https://zenodo.org/record/15014613/files/figure.png" pageId="13" pageNumber="14">Fig. 10D–F</figureCitation> ). </paragraph> <paragraph id="8B33368BF23DFFDEFF71FDBCFBC2DC86" blockId="13.[113,763,233,1979]" lastBlockId="13.[810,1460,143,1765]" pageId="13" pageNumber="14"> In most cases, the projections are quite asymmetric (one of the two is much more developed than the other one), and, therefore, the interepiplastral suture is not located sagittally but deviates laterally. The long and broad dorsal surface of the lip is generally rather flat (SAM -PQL -21174) or, very rarely, modestly convex (SAM -PLQ -24826a). In dorsal view, the pos - terior edge of the gular lip is nearly straight or modestly convex but in some cases it can be moderately overhanging. There are however some exceptions such as the female right epiplastra SAM -PQL -73749 and SAM -PQL -73878 ( -<figureCitation id="13B72A0EF23DFFDEFDCDFCA5FD69DE57" box="[561,681,859,883]" captionStart="Figure 10" captionStartId="12.[129,194,1334,1358]" captionTargetBox="[132,1471,147,1304]" captionTargetId="figure-312@12.[129,1473,144,1306]" captionTargetPageId="12" captionText="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." pageId="13" pageNumber="14">Fig. 10D–F</figureCitation> +<figureCitation id="13B72A0EF23DFFDEFDCDFCA5FD69DE57" box="[561,681,859,883]" captionStart="Figure 10" captionStartId="12.[129,194,1334,1358]" captionTargetBox="[132,1471,147,1304]" captionTargetId="figure-312@12.[129,1473,144,1306]" captionTargetPageId="12" captionText="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." figureDoi="http://doi.org/10.5281/zenodo.15014613" httpUri="https://zenodo.org/record/15014613/files/figure.png" pageId="13" pageNumber="14">Fig. 10D–F</figureCitation> ) (both fitting with <taxonomicName id="4C8C4D08F23DFFDEFF11FC84FE1ADEB6" box="[237,474,890,914]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="langebaanwegi"> <emphasis id="B9F8EA99F23DFFDEFF11FC84FE1ADEB6" box="[237,474,890,914]" italics="true" pageId="13" pageNumber="14">Chersina langebaanwegi</emphasis> @@ -1047,27 +1050,27 @@ wide and 22.0 mm long, and much more in SAM -PQL -22076 where it is wide and <quantity id="4C749B6EF23DFFDEFF8DF9F2FF10DB00" box="[113,208,1548,1572]" metricMagnitude="-2" metricUnit="m" metricValue="3.63" pageId="13" pageNumber="14" unit="mm" value="36.3">36.3 mm</quantity> long. The visceral surface of the epiplastron hosts a Y -shaped ridge whose anterior rami originates from the above described tubercles on the epiplastra. The central depression between the anterior rami is in all cases very modest. The ven - tral surface of the element is weakly concave. The hyoplastra are slightly longer than wide. They develop a rather robust and long buttress that reaches the first costal plate. Their medial sector is sexually dimorphic, being concave in males and flat in females. The hypoplastra are approximately rectangular. They develop a very robust posterior buttress that contacts the fifth and sixth costal plate. These elements show the same sexual dimorphism of the hyoplastra. The xipiplastra are not significantly dimorphic, with the exception of the presence of a medial concavity devel - oped in hyo -, hypo -, and xiphyplastra of males (the plastron is rather flat in females). In both sexes, the lateral edge is slightly convex with nearly no (SAM -PQL -22076; -<figureCitation id="13B72A0EF23DFFDEFB0AFF51FAF4DDE3" box="[1270,1332,175,199]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." pageId="13" pageNumber="14">Fig. 8</figureCitation> +<figureCitation id="13B72A0EF23DFFDEFB0AFF51FAF4DDE3" box="[1270,1332,175,199]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014609" httpUri="https://zenodo.org/record/15014609/files/figure.png" pageId="13" pageNumber="14">Fig. 8</figureCitation> ) or a very modest (SAM -PQL -20734; -<figureCitation id="13B72A0EF23DFFDEFBB2FF31FB48DDC3" box="[1102,1160,207,231]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="13" pageNumber="14">Fig. 1</figureCitation> +<figureCitation id="13B72A0EF23DFFDEFBB2FF31FB48DDC3" box="[1102,1160,207,231]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014591" httpUri="https://zenodo.org/record/15014591/files/figure.png" pageId="13" pageNumber="14">Fig. 1</figureCitation> ) notch corresponding to the femoroanal sulcus. The anal notch is in both sexes rather shallow and delimiting an open angle of about 125–140°. However, the male xipiplastra of SAM -PQL -21109 ( -<figureCitation id="13B72A0EF23DFFDEFB49FED3FB2EDC60" box="[1205,1262,300,325]" captionStart="Figure 5" captionStartId="7.[113,178,1626,1650]" captionTargetBox="[115,1456,145,1596]" captionTargetId="figure-177@7.[114,1458,144,1598]" captionTargetPageId="7" captionText="Figure 5. Chersina langebaanwegi, paratype SAM-PQL-21109, male: pygal area in posterior view (A); posterior portion of the shell in right lateral view (B); anterior lobe of the plastron in ventral and dorsal views (C, D); posterior lobe in ventral and dorsal views (E,F). Scale bars equal 5 cm." pageId="13" pageNumber="14">Fig. 5</figureCitation> +<figureCitation id="13B72A0EF23DFFDEFB49FED3FB2EDC60" box="[1205,1262,300,325]" captionStart="Figure 5" captionStartId="7.[113,178,1626,1650]" captionTargetBox="[115,1456,145,1596]" captionTargetId="figure-177@7.[114,1458,144,1598]" captionTargetPageId="7" captionText="Figure 5. Chersina langebaanwegi, paratype SAM-PQL-21109, male: pygal area in posterior view (A); posterior portion of the shell in right lateral view (B); anterior lobe of the plastron in ventral and dorsal views (C, D); posterior lobe in ventral and dorsal views (E,F). Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014601" httpUri="https://zenodo.org/record/15014601/files/figure.png" pageId="13" pageNumber="14">Fig. 5</figureCitation> ) are different from those of the equally large male SAM -PQL -22076 ( -<figureCitation id="13B72A0EF23DFFDEFABFFEB2FABFDC40" box="[1347,1407,332,356]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." pageId="13" pageNumber="14">Fig. 8</figureCitation> +<figureCitation id="13B72A0EF23DFFDEFABFFEB2FABFDC40" box="[1347,1407,332,356]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014609" httpUri="https://zenodo.org/record/15014609/files/figure.png" pageId="13" pageNumber="14">Fig. 8</figureCitation> ) be - cause their tip is bent upwards and the anal notch is much deeper and narrower (115°). </paragraph> <paragraph id="8B33368BF23DFFDEFCB9FE55FAB6D888" blockId="13.[810,1460,143,1765]" pageId="13" pageNumber="14"> <emphasis id="B9F8EA99F23DFFDEFCB9FE55FC02DCE6" box="[837,962,427,450]" italics="true" pageId="13" pageNumber="14">Horny scutes</emphasis> : The gular scutes are fused in a single unit that, in all cases ( -<figureCitation id="13B72A0EF23DFFDEFC4BFE37FBECDCC5" box="[951,1068,457,481]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="13" pageNumber="14">Fig. 1C, D</figureCitation> +<figureCitation id="13B72A0EF23DFFDEFC4BFE37FBECDCC5" box="[951,1068,457,481]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014591" httpUri="https://zenodo.org/record/15014591/files/figure.png" pageId="13" pageNumber="14">Fig. 1C, D</figureCitation> ), except one, reaches posteriorly to the anterior edge of the entoplastron (without entering it). In SAM -PQL -21109 ( -<figureCitation id="13B72A0EF23DFFDEFC0BFDF6FBF3DF04" box="[1015,1075,520,544]" captionStart="Figure 5" captionStartId="7.[113,178,1626,1650]" captionTargetBox="[115,1456,145,1596]" captionTargetId="figure-177@7.[114,1458,144,1598]" captionTargetPageId="7" captionText="Figure 5. Chersina langebaanwegi, paratype SAM-PQL-21109, male: pygal area in posterior view (A); posterior portion of the shell in right lateral view (B); anterior lobe of the plastron in ventral and dorsal views (C, D); posterior lobe in ventral and dorsal views (E,F). Scale bars equal 5 cm." pageId="13" pageNumber="14">Fig. 5</figureCitation> +<figureCitation id="13B72A0EF23DFFDEFC0BFDF6FBF3DF04" box="[1015,1075,520,544]" captionStart="Figure 5" captionStartId="7.[113,178,1626,1650]" captionTargetBox="[115,1456,145,1596]" captionTargetId="figure-177@7.[114,1458,144,1598]" captionTargetPageId="7" captionText="Figure 5. Chersina langebaanwegi, paratype SAM-PQL-21109, male: pygal area in posterior view (A); posterior portion of the shell in right lateral view (B); anterior lobe of the plastron in ventral and dorsal views (C, D); posterior lobe in ventral and dorsal views (E,F). Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014601" httpUri="https://zenodo.org/record/15014601/files/figure.png" pageId="13" pageNumber="14">Fig. 5</figureCitation> ), however, it stops anteriorly to the entoplastron as shown by the incomplete, but informative, left epiplastron. The gularohumeral sulcus is laterally concave on the dorsal surface and weakly anteromedially convex on the ven - tral surface. The angle at the junction of the two gularohumeral sulci (the right and the left) varies in relation to the form of the gular projection: from slightly less than 90° (SAM -PQL -21174), to approximately 90° (SAM -PQL -20734, 20925, 35023d), to slightly more than 90° (SAM -PQL -22076). The ratio between the length of the gulars and the humerals (the median antero - posterior length of the humerals) seems to vary according to the development of the gular projections and, therefore, the sex of the specimen: the female SAM -PQL -20734 has a gular as long as the humerals, whereas all the male plastral have humerals that are at least slightly longer than the gular. The humeropectoral su - ture regularly passes posterior to the entoplastron (that, there - fore, is crossed only by the interhumeral suture), and is regularly concave in the anterior direction but has a rather variable trend: it is much more open in the males SAM -PQL -20925 ( -<figureCitation id="13B72A0EF23DFFDEFABAFBC2FABDD970" box="[1350,1405,1084,1108]" captionStart="Figure 4" captionStartId="6.[129,194,1420,1444]" captionTargetBox="[131,1471,146,1390]" captionTargetId="figure-285@6.[129,1473,144,1392]" captionTargetPageId="6" captionText="Figure 4. Chersina langebaanwegi, paratype SAM-PQL-20925: male plastron in ventral view (A) and detail of the anterior lobe of the plastron in dorsal view (B); posterior lobe of the plastron in ventral view (C); pygal area in posterior view (D). Scale bars equal 5 cm." pageId="13" pageNumber="14">Fig. 4</figureCitation> +<figureCitation id="13B72A0EF23DFFDEFABAFBC2FABDD970" box="[1350,1405,1084,1108]" captionStart="Figure 4" captionStartId="6.[129,194,1420,1444]" captionTargetBox="[131,1471,146,1390]" captionTargetId="figure-285@6.[129,1473,144,1392]" captionTargetPageId="6" captionText="Figure 4. Chersina langebaanwegi, paratype SAM-PQL-20925: male plastron in ventral view (A) and detail of the anterior lobe of the plastron in dorsal view (B); posterior lobe of the plastron in ventral view (C); pygal area in posterior view (D). Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014599" httpUri="https://zenodo.org/record/15014599/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation> ) and SAM -PQL -22076 ( -<figureCitation id="13B72A0EF23DFFDEFC0FFBA5FBEAD957" box="[1011,1066,1115,1139]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." pageId="13" pageNumber="14">Fig. 8</figureCitation> +<figureCitation id="13B72A0EF23DFFDEFC0FFBA5FBEAD957" box="[1011,1066,1115,1139]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014609" httpUri="https://zenodo.org/record/15014609/files/figure.png" pageId="13" pageNumber="14">Fig. 8</figureCitation> ), where the medial sector of the sulcus is nearly straight and perpendicular to the sagittal axis of the plastron, than in the holotypic female shell SAM -PQL -20734 ( -<figureCitation id="13B72A0EF23DFFDEFCC9FB47FC61D9F5" box="[821,929,1209,1233]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="13" pageNumber="14">Fig. 1C, D</figureCitation> +<figureCitation id="13B72A0EF23DFFDEFCC9FB47FC61D9F5" box="[821,929,1209,1233]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." figureDoi="http://doi.org/10.5281/zenodo.15014591" httpUri="https://zenodo.org/record/15014591/files/figure.png" pageId="13" pageNumber="14">Fig. 1C, D</figureCitation> ). The pectoroabdominal sulci are modestly convex in the anterior direction and reach medially the mid of the interhyoplastral suture. The abdominofemoral sulci are located close to the hypoxiphyplastral suture but do not reach it medi - ally. They are laterally slightly convex in the anterior direction and medially nearly straight. The femoroanal sulci are mod - erately sigmoid and reach, with an acute posterior angle, the interxiphyplastral suture slightly anterior to its midlength. </paragraph> <paragraph id="8B33368BF23DFFDEFCB9FA4AFC3CDBC1" blockId="13.[810,1460,143,1765]" pageId="13" pageNumber="14">Axillary and inguinal scutes are present. The first ones (one per side) are relatively small and develop mostly on the third peripheral but also on the hyoplastron. The large inguinal scutes develop mainly at the posterolateral corner of each hypoplastron as well as partly on the seventh peripheral. The inguinal scute contacts the femoral scute both on the ventral and (especially) on the lateral surface of the hypoplastron. In visceral view, there is a significant covering of the plastral scutes on the dorsal sur - face of the bony elements delimiting the anterior and posterior aperture of the shell.</paragraph> @@ -1091,7 +1094,7 @@ The tentative referral of the female right epiplastron SAM - PQL -24826b to <emphasis id="B9F8EA99F23EFFDDFE0EFEF0FD5FDC01" box="[498,671,269,293]" italics="true" pageId="14" pageNumber="15">C. langebaanwegi</emphasis> </taxonomicName> . A convex epiplastral lip also characterizes the male left epiplastron SAM - PQL -24827a which shows a straight posterior edge of the gular lip. The same condition (convex epiplastral lip poster - iorly straight) is shown by the right female epiplastron SAM - PQL -73749. Both SAM -PQL -24826b and SAM -PQL -24827a have the tip of the gular protrusion slightly pointing in the anteroventral direction. Among the referred material, a convex epiplastral lip associated with a variably shaped posterior edge of the lip (and, therefore, a variably developed pocket) is pre - sent in the male epiplastra SAM -PQL -73750 and SAM -PQL - 12740c. The female right epiplastron SAM -PQL -73878 shows an intermediate condition in having a rather flat gular lip that ends posteriorly with a marked convexity that gives origin to a moderate pocket ( -<figureCitation id="13B72A0EF23EFFDDFEBDFD3AFE57DFF8" box="[321,407,708,732]" captionStart="Figure 10" captionStartId="12.[129,194,1334,1358]" captionTargetBox="[132,1471,147,1304]" captionTargetId="figure-312@12.[129,1473,144,1306]" captionTargetPageId="12" captionText="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." pageId="14" pageNumber="15">Fig. 10F</figureCitation> +<figureCitation id="13B72A0EF23EFFDDFEBDFD3AFE57DFF8" box="[321,407,708,732]" captionStart="Figure 10" captionStartId="12.[129,194,1334,1358]" captionTargetBox="[132,1471,147,1304]" captionTargetId="figure-312@12.[129,1473,144,1306]" captionTargetPageId="12" captionText="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." figureDoi="http://doi.org/10.5281/zenodo.15014613" httpUri="https://zenodo.org/record/15014613/files/figure.png" pageId="14" pageNumber="15">Fig. 10F</figureCitation> ). </paragraph> </subSubSection> @@ -1101,14 +1104,14 @@ The tentative referral of the female right epiplastron SAM - PQL -24826b to </paragraph> <paragraph id="8B33368BF23EFFDDFF7DFCBCFEB7D931" blockId="14.[128,779,763,1327]" pageId="14" pageNumber="15"> The phylogenetic analysis resulted in the consensus trees re - ported in -<figureCitation id="13B72A0EF23EFFDDFF17FC9FFE90DE5D" box="[235,336,865,889]" captionStart="Figure 11" captionStartId="15.[113,178,1447,1471]" captionTargetBox="[117,1455,147,1416]" captionTargetId="figure-213@15.[114,1458,144,1419]" captionTargetPageId="15" captionText="Figure 11. Strict consensus trees without (A) and with (B) a molecular scaffold to constrain the position of extant taxa (according to the topologies of Thomson et al.2021 and Fritz et al. 2022). Both the topologies result from maximum parsimony phylogenetic analyses under implied weights (k = 6) of 42 extant taxa and four extinct taxa (highlighted with diamonds), using 171 osteological characters.The analysis is based on the character matrix and character definitions of Vlachos and Rabi (2018) but all extinct taxa, except the African ones coming from South of the Equator (Impregnochelys pachytectis, Mesochersus orangeus, and Namibchersus namaquensis), were removed and four extant taxa (Astrochelys yniphora, Chersina angulata, Chersobius signatus, and Pyois arachnoides), scored for shell characters only, were added." pageId="14" pageNumber="15">Figure 11</figureCitation> +<figureCitation id="13B72A0EF23EFFDDFF17FC9FFE90DE5D" box="[235,336,865,889]" captionStart="Figure 11" captionStartId="15.[113,178,1447,1471]" captionTargetBox="[117,1455,147,1416]" captionTargetId="figure-213@15.[114,1458,144,1419]" captionTargetPageId="15" captionText="Figure 11. Strict consensus trees without (A) and with (B) a molecular scaffold to constrain the position of extant taxa (according to the topologies of Thomson et al.2021 and Fritz et al. 2022). Both the topologies result from maximum parsimony phylogenetic analyses under implied weights (k = 6) of 42 extant taxa and four extinct taxa (highlighted with diamonds), using 171 osteological characters.The analysis is based on the character matrix and character definitions of Vlachos and Rabi (2018) but all extinct taxa, except the African ones coming from South of the Equator (Impregnochelys pachytectis, Mesochersus orangeus, and Namibchersus namaquensis), were removed and four extant taxa (Astrochelys yniphora, Chersina angulata, Chersobius signatus, and Pyois arachnoides), scored for shell characters only, were added." figureDoi="http://doi.org/10.5281/zenodo.15014615" httpUri="https://zenodo.org/record/15014615/files/figure.png" pageId="14" pageNumber="15">Figure 11</figureCitation> . The first round of the analysis without a molecular scaffold ( -<figureCitation id="13B72A0EF23EFFDDFEACFC7EFE69DEBC" box="[336,425,896,920]" captionStart="Figure 11" captionStartId="15.[113,178,1447,1471]" captionTargetBox="[117,1455,147,1416]" captionTargetId="figure-213@15.[114,1458,144,1419]" captionTargetPageId="15" captionText="Figure 11. Strict consensus trees without (A) and with (B) a molecular scaffold to constrain the position of extant taxa (according to the topologies of Thomson et al.2021 and Fritz et al. 2022). Both the topologies result from maximum parsimony phylogenetic analyses under implied weights (k = 6) of 42 extant taxa and four extinct taxa (highlighted with diamonds), using 171 osteological characters.The analysis is based on the character matrix and character definitions of Vlachos and Rabi (2018) but all extinct taxa, except the African ones coming from South of the Equator (Impregnochelys pachytectis, Mesochersus orangeus, and Namibchersus namaquensis), were removed and four extant taxa (Astrochelys yniphora, Chersina angulata, Chersobius signatus, and Pyois arachnoides), scored for shell characters only, were added." pageId="14" pageNumber="15">Fig. 11A</figureCitation> +<figureCitation id="13B72A0EF23EFFDDFEACFC7EFE69DEBC" box="[336,425,896,920]" captionStart="Figure 11" captionStartId="15.[113,178,1447,1471]" captionTargetBox="[117,1455,147,1416]" captionTargetId="figure-213@15.[114,1458,144,1419]" captionTargetPageId="15" captionText="Figure 11. Strict consensus trees without (A) and with (B) a molecular scaffold to constrain the position of extant taxa (according to the topologies of Thomson et al.2021 and Fritz et al. 2022). Both the topologies result from maximum parsimony phylogenetic analyses under implied weights (k = 6) of 42 extant taxa and four extinct taxa (highlighted with diamonds), using 171 osteological characters.The analysis is based on the character matrix and character definitions of Vlachos and Rabi (2018) but all extinct taxa, except the African ones coming from South of the Equator (Impregnochelys pachytectis, Mesochersus orangeus, and Namibchersus namaquensis), were removed and four extant taxa (Astrochelys yniphora, Chersina angulata, Chersobius signatus, and Pyois arachnoides), scored for shell characters only, were added." figureDoi="http://doi.org/10.5281/zenodo.15014615" httpUri="https://zenodo.org/record/15014615/files/figure.png" pageId="14" pageNumber="15">Fig. 11A</figureCitation> ; Consistency Index (CI) = 0.293; Retention Index (RI) = 0.620; Most Parsimonious Trees (MPT) = 47.997) found two most parsimonious trees, and the second round of tree bisection and reconnection retained one most parsimonious tree. </paragraph> <paragraph id="8B33368BF23EFFDDFF60FBE3FCC9D80B" blockId="14.[128,779,763,1327]" pageId="14" pageNumber="15"> In the analysis with the molecular scaffold ( -<figureCitation id="13B72A0EF23EFFDDFD5EFBE3FCC2D911" box="[674,770,1053,1077]" captionStart="Figure 11" captionStartId="15.[113,178,1447,1471]" captionTargetBox="[117,1455,147,1416]" captionTargetId="figure-213@15.[114,1458,144,1419]" captionTargetPageId="15" captionText="Figure 11. Strict consensus trees without (A) and with (B) a molecular scaffold to constrain the position of extant taxa (according to the topologies of Thomson et al.2021 and Fritz et al. 2022). Both the topologies result from maximum parsimony phylogenetic analyses under implied weights (k = 6) of 42 extant taxa and four extinct taxa (highlighted with diamonds), using 171 osteological characters.The analysis is based on the character matrix and character definitions of Vlachos and Rabi (2018) but all extinct taxa, except the African ones coming from South of the Equator (Impregnochelys pachytectis, Mesochersus orangeus, and Namibchersus namaquensis), were removed and four extant taxa (Astrochelys yniphora, Chersina angulata, Chersobius signatus, and Pyois arachnoides), scored for shell characters only, were added." pageId="14" pageNumber="15">Fig. 11B</figureCitation> +<figureCitation id="13B72A0EF23EFFDDFD5EFBE3FCC2D911" box="[674,770,1053,1077]" captionStart="Figure 11" captionStartId="15.[113,178,1447,1471]" captionTargetBox="[117,1455,147,1416]" captionTargetId="figure-213@15.[114,1458,144,1419]" captionTargetPageId="15" captionText="Figure 11. Strict consensus trees without (A) and with (B) a molecular scaffold to constrain the position of extant taxa (according to the topologies of Thomson et al.2021 and Fritz et al. 2022). Both the topologies result from maximum parsimony phylogenetic analyses under implied weights (k = 6) of 42 extant taxa and four extinct taxa (highlighted with diamonds), using 171 osteological characters.The analysis is based on the character matrix and character definitions of Vlachos and Rabi (2018) but all extinct taxa, except the African ones coming from South of the Equator (Impregnochelys pachytectis, Mesochersus orangeus, and Namibchersus namaquensis), were removed and four extant taxa (Astrochelys yniphora, Chersina angulata, Chersobius signatus, and Pyois arachnoides), scored for shell characters only, were added." figureDoi="http://doi.org/10.5281/zenodo.15014615" httpUri="https://zenodo.org/record/15014615/files/figure.png" pageId="14" pageNumber="15">Fig. 11B</figureCitation> ; CI = 0.268, RI = 0.572; MPT 53.1423), the New Technology Search found two most parsimonious trees, and the second round of tree bisection and reconnection retained two most par - simonious trees. Both the analyses confirm that the new extinct species from Langebaanweg is the sister taxon of <taxonomicName id="4C8C4D08F23EFFDDFD73FB44FCC4D9F5" box="[655,772,1209,1233]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="angulata"> <emphasis id="B9F8EA99F23EFFDDFD73FB44FCC4D9F5" box="[655,772,1209,1233]" italics="true" pageId="14" pageNumber="15">C. angulata</emphasis>