Add updates up until 2024-10-24 22:00:43

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@ -43,18 +43,18 @@
<paragraph id="ACD780FCE38D2C3F0C819B466041648E" pageNumber="126">Fig. 28.</paragraph>
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[[worker]]. - Long. 2.2-2,3 mill. Noire. Mandibules, les deux tiers basilaires du scape et tarses roussatres. Reste des antennes bruna- tre. Hanches, pattes et abdomen lisses et luisants, le reste subopaque, densement strie en long, la tete et le pronotum bien plus finement que le reste du thorax qui est en outre un peu ride en long. Pilosite dressee noire, setiforme, plus longue que chez
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in sp., assez abondante sur le gastre, plus dispersee ailleurs, y compris les pattes. Pubescence assez rare, clairsemee, sauf sur les pattes et les antennes ou elle est un peu plus copieuse. Tete presque aussi longue que large, assez convexe derriere et sur les cotes. Les yeux, peu convexes, sont places un peu en arriere du milieu des cotes, dont ils occupent un peu plus du tiers; la distance qui les separe de la base des mandibules est a peu pres egale a leur grand diametre. Epistome carene, finement reticule et luisant, aire frontale distincte. Mandibules lisses, ponctuees, de 4 a 5. dents. Le scape depasse le bord occipital de la moitie de leur longueur. Deuxieme article du funicule long comme la moitie du premier et le troisieme comme les trois quarts. Pronotum un peu plus large que long, legerement concave en dessus, d'un cote a l'autre; cotes tres saillants en dehors et un peu releves au milieu. Suture promesonotale tres distincte. Etranglement du mesonotum plus marque que chez
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Devant du mesonotum un peu convexe; les saillies portant les stigmates occupent le tiers posterieur, elles sont triangulaires, plus larges a leur base que hautes, le pan anterieur egal au posterieur et mousse au sommet. L'epinotum est coupe a pic au-devant de l'incisure metanotale. La face basale forme deux protuberances coniques s'elevant en arriere et en dehors, mousses a l'extremite, contiguos a leur base que limite un sillon median. Vu de devant, l'intervalle est fortement concave. Face declive de l'epinotum plus longue, fortement oblique dans les deux tiers superieurs, horizontale dans le tiers restant. Ecaille plus haute que la longueur de son petiole poste- rieur, presque trois fois aussi haute qu'epaisse, faiblement ovale, avec deux petites dents au sommet, separees par une echancrure mediane mediocre. Gastre court et spherique.
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Espece tres caracteristique par son pronotum et sa sculpture qui la distinguent de
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et
<taxonomicName id="09A418554F33326779A46A089E3BF0A9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139218" class="Magnoliopsida" family="Asteraceae" genus="Acantholepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acantholepis carbonaria Emery" lsidName-HNS="Acantholepis carbonaria Emery" order="Asterales" pageNumber="127" phylum="Tracheophyta" rank="species" species="carbonaria">carbonaria Emery</taxonomicName>
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. Voisine de
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<taxonomicName id="89F8A44D3C507AD14F0F8F88D4C2D2DF" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139224" class="Insecta" family="Formicidae" genus="Acantholepis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Acantholepis deplanatus Stitz" lsidName-HNS="Acantholepis deplanatus Stitz" order="Hymenoptera" pageNumber="127" phylum="Arthropoda" rank="species" species="deplanatus">A. deplanatus Stitz</taxonomicName>
, mais la couleur et la sculpture different.
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P. Schmalzi
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[[ worker ]] Ferrugineo-testacea, col capo e parte dell' addome piu scuri, le mandibole e le zampe piu chiari; coperta di pubescenza gialla, mediocremente abbondante e con brevi peli ritti; questi ultimi mancano sulle tibie. Capo piu lungo che largo, alquanto ristretto innanzi, col margine posteriore dritto; gli occhi distano dal margine anteriore piu che il loro diametro e contano nel loro maggior diametro 6 grosse faccette. La punteggiatura del capo ricorda quella della
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, ma una sottoscultura microscopica rende la superficie opaca; ii solco frontale si prolunga fino a meta della lunghezza del capo, le lamine frontali sono brevemente ciliate. Le mandibole sono lucide e sparse di punti sottili, armate di 11 denti alternamente piu grandi e piu piccoli, gli anteriori ben marcati, i posteriori rudimentali. Lo scapo delle antenne raggiunge l'occipite, gli articoli 2 - 6 del flagello sono piccoli e fortemente trasversi, 7 - 10 molto piu grandi dei precedenti e un poco piu larghi che lunghi. Torace, peduncolo e addome sono lucidi, finamente punteggiati; ii dorso e subrettilineo, pero alquanto ondulato e con le suture ben marcate; sul profilo, il metanoto forma un angolo ottuso, ma ben distinto; la faccia declive e debolmente marginata sui lati. Le mesopleure hanno un dente verso i 2 / 5 superiori del loro margine anteriore. Il peduncolo ha una squama alta quanto il metanoto, poco piu larga di esso, poco meno di due volte alta quanto e grossa alla base, piana indietro e appena convessa innanzi, meno larga che alta, e alquanto assottigliata superiormente; inferiormente, il pedunculo porta in avanti una laminetta ottusa. Il 2. ° segmento e ottusamente troncato in avanti; la strozzatura alla base del 3. ° segmento e mediocremente marcata. L. 3 1 / 3 — 3 2 / 3 mm. Joinville in S. Catharina.
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<mods:title id="FDCE61F653D4DDA80698625AA1C0D515">A new genus and species of barnacle (Cirripedia, Verrucomorpha) associated with vents of the Lau Back-Arc Basin: its gross morphology, inferred first juvenile stage and affinities</mods:title>
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SPECIES. —
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<emphasis id="B95CDA13FFB4D020FEE1FDB37A479D40" box="[284,541,529,553]" italics="true" pageId="3" pageNumber="74">Imbricaverruca yamaguchii</emphasis>
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ETYMOLOGY. — From the Latin
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(to cover with tiles or scales) and
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, in reference to its otherwise
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-like appearance.
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DIAGNOSIS. — A neoverrucid differing from
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in numerous ways including having 1) a more verrucid-like operculum despite having the included median latus well developed rather than vestigial or absent, 2) a fixed scutum and tergum much wider than high rather than at least as high as wide, and 3) in retaining a complete set of well developed imbricating plates on the movable side of the wall.
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<heading id="D0D081A0FFEEE40EFEBFF8A7FD04F83F" box="[263,645,1905,1932]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="501" reason="2">
Genus
<taxonomicName id="4C274D4FFFEEE40EFEECF8A7FDAEF838" authority="Merle" authorityName="Merle" authorityYear="2024" box="[340,559,1905,1931]" class="Gastropoda" family="Muricidae" genus="Paleochicoreus" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="501" phylum="Mollusca" rank="genus" status="n. gen.">
<emphasis id="B953EADEFFEEE40EFEECF8A7FE67F838" box="[340,486,1905,1931]" italics="true" pageId="8" pageNumber="501">Paleochicoreus</emphasis>
Merle
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFEEE40EFD82F8A4FD04F83F" box="[570,645,1906,1932]" pageId="8" pageNumber="501" rank="genus">n. gen.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFEEE40EFE26F847FC86F852" pageId="8" pageNumber="501" type="description">
<paragraph id="8B9836CCFFEEE40EFE26F847FE6CF818" blockId="8.[263,645,1905,1963]" box="[414,493,1937,1963]" pageId="8" pageNumber="501">
(
<figureCitation id="131C2A49FFEEE40EFE1FF847FE65F818" box="[423,484,1937,1963]" captionStart="FIG" captionStartId="8.[132,143,1516,1533]" captionTargetBox="[132,1455,215,1472]" captionTargetId="figure-286@8.[698,1267,753,1294]" captionTargetPageId="8" captionText="FIG. 2. — Spiral cords and internal denticles of the outer lip in Paleochicoreus Merle, n. gen.: A, B, Paleoochicoreus initialis (Vokes, 1990), n. comb., holotype, USNM450376, Nacodoches, Texas (United States), middle Eocene (Lutetian); C, P. newtoni (Eames, 1957), holotype, BMNH G41982, Ombralla, Nigeria, middle Eocene;D, E, P. bicostatus (Deshayes,1835), n. comb., holotype, UCBL-EM 33437 (Deshayes coll.), Valmondois,Val dOise, Paris Basin, France, middle Eocene (Bartonian). Scale bars: 10 mm. Credits: A, B, Y. Villacampa and J. Jorstad (USNM); C, S. Tracey (NHM); D, E, P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985764" httpUri="https://zenodo.org/record/13985764/files/figure.png" pageId="8" pageNumber="501">Fig. 2</figureCitation>
)
</paragraph>
<paragraph id="8B9836CCFFEEE40EFF3CF806FC86F852" blockId="8.[132,775,2000,2017]" box="[132,775,2000,2017]" pageId="8" pageNumber="501">
<uri id="FFB63ACEFFEEE40EFF3CF806FC86F852" box="[132,775,2000,2017]" pageId="8" pageNumber="501">
urn:lsid:zoobank.org:act:
<uuid id="FF810C19FFEEE40EFEE3F806FC86F852" box="[347,775,2000,2017]" pageId="8" pageNumber="501">3E5850A7-582D-48BF-B126-BA40739DDC00</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFEEE40EFC95F947FB80F858" pageId="8" pageNumber="501" type="materials_examined">
<paragraph id="8B9836CCFFEEE40EFC95F947FABDF977" blockId="8.[810,1456,1681,2027]" pageId="8" pageNumber="501">
<smallCapsWord id="8D7EA010FFEEE40EFC95F947FCDCF914" baselines="1698,1699" box="[813,861,1681,1704]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="8" pageNumber="501">
<typeStatus id="549C886EFFEEE40EFC95F947FCDCF914" box="[813,861,1681,1704]" pageId="8" pageNumber="501">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D7EA010FFEEE40EFCD9F940FC27F914" baselines="1699" box="[865,934,1686,1703]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="species" pageId="8" pageNumber="501">SPECIES</smallCapsWord>
. —
<taxonomicName id="4C274D4FFFEEE40EFC73F947FAE2F91A" authority="Vokes, 1990" authorityName="Vokes" authorityYear="1990" box="[971,1379,1681,1705]" class="Gastropoda" family="Muricidae" genus="Chicoreus" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="501" phylum="Mollusca" rank="species" species="initialis" subGenus="Phyllonotus">
<emphasis id="B953EADEFFEEE40EFC73F947FBA2F91A" box="[971,1059,1681,1705]" italics="true" pageId="8" pageNumber="501">Chicoreus</emphasis>
(
<emphasis id="B953EADEFFEEE40EFB89F947FB18F91A" box="[1073,1177,1681,1705]" italics="true" pageId="8" pageNumber="501">Phyllonotus</emphasis>
)
<emphasis id="B953EADEFFEEE40EFB1DF947FB6AF91A" box="[1189,1259,1681,1705]" italics="true" pageId="8" pageNumber="501">initialis</emphasis>
<bibRefCitation id="EFB64B3DFFEEE40EFB55F947FAE2F91A" author="VOKES E. H." box="[1261,1379,1681,1705]" pageId="8" pageNumber="501" pagination="1 - 96" refId="ref48944" refString="VOKES E. H. 1990. - Cenozoic Muricidae of the Western Atlantic region, Part VIII - Murex s. s., Haustellum, Chicoreus, Hexaplex; additions and corrections. Tulane Studies in Geology and Paleontology 23 (1 - 3): 1 - 96." type="journal article" year="1990">Vokes, 1990</bibRefCitation>
</taxonomicName>
.
<collectingCountry id="F330765CFFEEE40EFAD3F947FCE7F977" name="United States of America" pageId="8" pageNumber="501">United States</collectingCountry>
,
<collectingRegion id="49E3F82EFFEEE40EFCD6F97AFC27F970" box="[878,934,1708,1731]" country="United States of America" name="Texas" pageId="8" pageNumber="501">Texas</collectingRegion>
, middle Eocene (Bartonian) (
<figureCitation id="131C2A49FFEEE40EFB71F97DFAB1F977" box="[1225,1328,1707,1732]" captionStart="FIG" captionStartId="8.[132,143,1516,1533]" captionTargetBox="[132,1455,215,1472]" captionTargetId="figure-286@8.[698,1267,753,1294]" captionTargetPageId="8" captionText="FIG. 2. — Spiral cords and internal denticles of the outer lip in Paleochicoreus Merle, n. gen.: A, B, Paleoochicoreus initialis (Vokes, 1990), n. comb., holotype, USNM450376, Nacodoches, Texas (United States), middle Eocene (Lutetian); C, P. newtoni (Eames, 1957), holotype, BMNH G41982, Ombralla, Nigeria, middle Eocene;D, E, P. bicostatus (Deshayes,1835), n. comb., holotype, UCBL-EM 33437 (Deshayes coll.), Valmondois,Val dOise, Paris Basin, France, middle Eocene (Bartonian). Scale bars: 10 mm. Credits: A, B, Y. Villacampa and J. Jorstad (USNM); C, S. Tracey (NHM); D, E, P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985764" httpUri="https://zenodo.org/record/13985764/files/figure.png" pageId="8" pageNumber="501">Fig. 2A, B</figureCitation>
).
</paragraph>
<paragraph id="8B9836CCFFEEE40EFC95F903FB80F858" blockId="8.[810,1456,1681,2027]" pageId="8" pageNumber="501">
<smallCapsWord id="8D7EA010FFEEE40EFC95F903FCF4F95F" baselines="1767,1768" box="[813,885,1749,1773]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Species" pageId="8" pageNumber="501">SPECIES</smallCapsWord>
<smallCapsWord id="8D7EA010FFEEE40EFCC1F90DFC5BF95F" baselines="1768" box="[889,986,1755,1772]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="included" pageId="8" pageNumber="501">INCLUDED</smallCapsWord>
. —
<taxonomicName id="4C274D4FFFEEE40EFBB8F900FAB2F95E" authority="Dickerson, 1915" authorityName="Dickerson" authorityYear="1915" box="[1024,1331,1749,1773]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="501" phylum="Mollusca" rank="species" species="packardi">
<emphasis id="B953EADEFFEEE40EFBB8F900FB0EF95E" box="[1024,1167,1749,1773]" italics="true" pageId="8" pageNumber="501">Murex packardi</emphasis>
<bibRefCitation id="EFB64B3DFFEEE40EFB2BF903FAB2F95E" author="DICKERSON R. E." box="[1171,1331,1749,1773]" pageId="8" pageNumber="501" pagination="33 - 98" refId="ref44126" refString="DICKERSON R. E. 1915. - Fauna of the type Tejon: its relation to the Cowlitz phase of the Tejon group of Washington. Proceedings of the California Academy of Sciences, 4 th series, 5 (3): 33 - 98." type="journal article" year="1915">Dickerson, 1915</bibRefCitation>
</taxonomicName>
,
<collectingRegion id="49E3F82EFFEEE40EFA83F903FA2FF95E" box="[1339,1454,1749,1773]" country="United States of America" name="Washington" pageId="8" pageNumber="501">Washington</collectingRegion>
(
<collectingCountry id="F330765CFFEEE40EFC89F926FC30F8BB" box="[817,945,1776,1800]" name="United States of America" pageId="8" pageNumber="501">United States</collectingCountry>
), middle Eocene;
<taxonomicName id="4C274D4FFFEEE40EFBEBF926FCE3F891" authority="Vokes, 1990" authorityName="Vokes" authorityYear="1990" class="Gastropoda" family="Muricidae" genus="Chicoreus" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="501" phylum="Mollusca" rank="species" species="initialis" subGenus="Phyllonotus">
<emphasis id="B953EADEFFEEE40EFBEBF926FB2BF8BB" box="[1107,1194,1776,1800]" italics="true" pageId="8" pageNumber="501">Chicoreus</emphasis>
(
<emphasis id="B953EADEFFEEE40EFB0FF926FA9CF8BB" box="[1207,1309,1776,1800]" italics="true" pageId="8" pageNumber="501">Phyllonotus</emphasis>
)
<emphasis id="B953EADEFFEEE40EFA91F926FAEFF8BB" box="[1321,1390,1776,1800]" italics="true" pageId="8" pageNumber="501">initialis</emphasis>
<bibRefCitation id="EFB64B3DFFEEE40EFAC8F926FCE3F891" author="VOKES E. H." pageId="8" pageNumber="501" pagination="1 - 96" refId="ref48944" refString="VOKES E. H. 1990. - Cenozoic Muricidae of the Western Atlantic region, Part VIII - Murex s. s., Haustellum, Chicoreus, Hexaplex; additions and corrections. Tulane Studies in Geology and Paleontology 23 (1 - 3): 1 - 96." type="journal article" year="1990">Vokes, 1990</bibRefCitation>
</taxonomicName>
,
<collectingRegion id="49E3F82EFFEEE40EFCD3F8DDFC21F891" box="[875,928,1803,1826]" country="United States of America" name="Texas" pageId="8" pageNumber="501">Texas</collectingRegion>
(
<collectingCountry id="F330765CFFEEE40EFC15F8DCFBB4F891" box="[941,1077,1802,1826]" name="United States of America" pageId="8" pageNumber="501">United States</collectingCountry>
), middle Eocene (Lutetian);
<emphasis id="B953EADEFFEEE40EFAF4F8DDFA2EF891" box="[1356,1455,1803,1826]" italics="true" pageId="8" pageNumber="501">Pterynoyus</emphasis>
<taxonomicName id="4C274D4FFFEEE40EFC95F8F0FC7AF88E" authority="Eames, 1957" authorityName="Eames" authorityYear="1957" box="[813,1019,1829,1853]" class="Gastropoda" family="Muricidae" genus="Paleoochicoreus" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="501" phylum="Mollusca" rank="species" species="newtoni">
<emphasis id="B953EADEFFEEE40EFC95F8F0FCF9F88E" box="[813,888,1830,1853]" italics="true" pageId="8" pageNumber="501">newtoni</emphasis>
<bibRefCitation id="EFB64B3DFFEEE40EFCC5F8F3FC7AF88E" author="EAMES F. E." box="[893,1019,1829,1853]" pageId="8" pageNumber="501" pagination="25 - 70" refId="ref44211" refString="EAMES F. E. 1957. - Eocene Mollusca from Nigeria: a revision. Bulletin of the British Museum of the Natural History (Geology) 3 (2): 25 - 70. https: // doi. org / 10.5962 / p. 143489" type="journal article" year="1957">Eames, 1957</bibRefCitation>
</taxonomicName>
[new name for “
<emphasis id="B953EADEFFEEE40EFB23F8F0FA96F88E" box="[1179,1303,1830,1853]" italics="true" pageId="8" pageNumber="501">Pteropurpura</emphasis>
<taxonomicName id="4C274D4FFFEEE40EFA9EF8F3FC3FF8EB" authority="Newton, 1922" authorityName="Newton" authorityYear="1922" class="Gastropoda" family="Muricidae" genus="Murex" isUncertain="true" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="501" phylum="Mollusca" rank="species" species="tricarinatus">
cf.
<emphasis id="B953EADEFFEEE40EFAFAF8F0FA2EF88E" box="[1346,1455,1830,1853]" italics="true" pageId="8" pageNumber="501">tricarinatus</emphasis>
<bibRefCitation id="EFB64B3DFFEEE40EFC95F896FC3FF8EB" author="NEWTON R. B." box="[813,958,1856,1880]" pageId="8" pageNumber="501" pagination="7 - 114" refId="ref47686" refString="NEWTON R. B. 1922. - Eocene Mollusca from Nigeria. Geological Survey of Nigeria Bulletin 3: 7 - 114. https: // www. biodiversitylibrary. org / page / 22642662" type="journal article" year="1922">Newton, 1922</bibRefCitation>
</taxonomicName>
],
<collectingCountry id="F330765CFFEEE40EFC76F8E9FB98F8E4" box="[974,1049,1855,1879]" name="Nigeria" pageId="8" pageNumber="501">Nigeria</collectingCountry>
, middle Eocene,
<figureCitation id="131C2A49FFEEE40EFB04F8E9FA86F8EB" box="[1212,1287,1855,1880]" captionStart="FIG" captionStartId="8.[132,143,1516,1533]" captionTargetBox="[132,1455,215,1472]" captionTargetId="figure-286@8.[698,1267,753,1294]" captionTargetPageId="8" captionText="FIG. 2. — Spiral cords and internal denticles of the outer lip in Paleochicoreus Merle, n. gen.: A, B, Paleoochicoreus initialis (Vokes, 1990), n. comb., holotype, USNM450376, Nacodoches, Texas (United States), middle Eocene (Lutetian); C, P. newtoni (Eames, 1957), holotype, BMNH G41982, Ombralla, Nigeria, middle Eocene;D, E, P. bicostatus (Deshayes,1835), n. comb., holotype, UCBL-EM 33437 (Deshayes coll.), Valmondois,Val dOise, Paris Basin, France, middle Eocene (Bartonian). Scale bars: 10 mm. Credits: A, B, Y. Villacampa and J. Jorstad (USNM); C, S. Tracey (NHM); D, E, P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985764" httpUri="https://zenodo.org/record/13985764/files/figure.png" pageId="8" pageNumber="501">Fig. 2C</figureCitation>
;
<taxonomicName id="4C274D4FFFEEE40EFAA9F896FC32F8C1" authority="Vincent, 1913" authorityName="Vincent" authorityYear="1913" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="501" phylum="Mollusca" rank="species" species="diderrichi">
<emphasis id="B953EADEFFEEE40EFAA9F896FA2EF8E4" box="[1297,1455,1855,1879]" italics="true" pageId="8" pageNumber="501">Murex diderrichi</emphasis>
<bibRefCitation id="EFB64B3DFFEEE40EFC95F88CFC32F8C1" author="VINCENT G." box="[813,947,1882,1906]" pageId="8" pageNumber="501" pagination="1 - 46" refId="ref48789" refString="VINCENT G. 1913. - La faune Paleocene de Landana. Annales du Musee du Congo Belge. Geologie, Paleontologie, Mineralogie serie 3, 1 (1): 1 - 46." type="journal article" year="1913">Vincent, 1913</bibRefCitation>
</taxonomicName>
[?synonym of
<taxonomicName id="4C274D4FFFEEE40EFB8FF88DFA22F8C1" authority="Oppenheim, 1904" authorityName="Oppenheim" authorityYear="1904" box="[1079,1443,1882,1906]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="501" phylum="Mollusca" rank="species" species="camerunensis">
<emphasis id="B953EADEFFEEE40EFB8FF88DFB6FF8C1" box="[1079,1262,1883,1906]" italics="true" pageId="8" pageNumber="501">Murex camerunensis</emphasis>
<bibRefCitation id="EFB64B3DFFEEE40EFB49F88CFA22F8C1" author="OPPENHEIM P." box="[1265,1443,1882,1906]" pageId="8" pageNumber="501" pagination="243 - 283" refId="ref47867" refString="OPPENHEIM P. 1904. - Uber Tertiarfossilien, warscheinlich Eocanen Altes von Kamerun. Beitrage zur Geologie von Kamerun 3: 243 - 283." type="journal article" year="1904">Oppenheim, 1904</bibRefCitation>
</taxonomicName>
],
<collectingCountry id="F330765CFFEEE40EFC93F8A2FCF2F83F" box="[811,883,1908,1932]" name="Angola" pageId="8" pageNumber="501">Angola</collectingCountry>
,
<collectingCountry id="F330765CFFEEE40EFCC6F8A3FC68F83E" box="[894,1001,1909,1933]" name="Cameroon" pageId="8" pageNumber="501">Cameroon</collectingCountry>
, late Paleocene-early Eocene;
<taxonomicName id="4C274D4FFFEEE40EFAAAF8A3FC4BF814" authority="Deshayes, 1835" authorityName="Deshayes" authorityYear="1835" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="501" phylum="Mollusca" rank="species" species="bicostatus">
<emphasis id="B953EADEFFEEE40EFAAAF8A3FA31F83F" box="[1298,1456,1908,1932]" italics="true" pageId="8" pageNumber="501">Murex bicostatus</emphasis>
<bibRefCitation id="EFB64B3DFFEEE40EFC96F859FC4BF814" author="DESHAYES G. - P." box="[814,970,1935,1959]" pageId="8" pageNumber="501" pagination="79 - 106" refId="ref43930" refString="DESHAYES G. - P. 1835. - Description des coquilles fossiles des environs de Paris. Tome 2. Livraisons 40 - 45, l'auteur, Bechet Jeune, Baudouin freres, Treuttel &amp; Wurtz, Paris: 499 - 780; Atlas 2, Levrault, Paris: pl. 79 - 106. https: // doi. org / 10.5962 / bhl. title. 52303" type="book chapter" year="1835">Deshayes, 1835</bibRefCitation>
</taxonomicName>
,
<collectingCountry id="F330765CFFEEE40EFC6CF846FB99F814" box="[980,1048,1936,1959]" name="France" pageId="8" pageNumber="501">France</collectingCountry>
, middle Eocene (Bartonian) (
<figureCitation id="131C2A49FFEEE40EFA80F859FA23F814" box="[1336,1442,1935,1959]" captionStart="FIG" captionStartId="8.[132,143,1516,1533]" captionTargetBox="[132,1455,215,1472]" captionTargetId="figure-286@8.[698,1267,753,1294]" captionTargetPageId="8" captionText="FIG. 2. — Spiral cords and internal denticles of the outer lip in Paleochicoreus Merle, n. gen.: A, B, Paleoochicoreus initialis (Vokes, 1990), n. comb., holotype, USNM450376, Nacodoches, Texas (United States), middle Eocene (Lutetian); C, P. newtoni (Eames, 1957), holotype, BMNH G41982, Ombralla, Nigeria, middle Eocene;D, E, P. bicostatus (Deshayes,1835), n. comb., holotype, UCBL-EM 33437 (Deshayes coll.), Valmondois,Val dOise, Paris Basin, France, middle Eocene (Bartonian). Scale bars: 10 mm. Credits: A, B, Y. Villacampa and J. Jorstad (USNM); C, S. Tracey (NHM); D, E, P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985764" httpUri="https://zenodo.org/record/13985764/files/figure.png" pageId="8" pageNumber="501">Fig. 2D, E</figureCitation>
).
<smallCapsWord id="8D7EA010FFEEE40EFC95F86FFC27F87C" baselines="1994,1995" box="[813,934,1977,2000]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="8" pageNumber="501">ETYMOLOGY</smallCapsWord>
. — Name formed by
<emphasis id="B953EADEFFEEE40EFBC7F86FFB3EF862" box="[1151,1215,1977,2001]" italics="true" pageId="8" pageNumber="501">palaiós</emphasis>
(meaning old in ancient Greek) and
<taxonomicName id="4C274D4FFFEEE40EFC19F805FC7AF858" authorityName="Montfort" authorityYear="1810" box="[929,1019,2003,2027]" class="Gastropoda" family="Muricidae" genus="Chicoreus" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="501" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFEEE40EFC19F805FC7AF858" box="[929,1019,2003,2027]" italics="true" pageId="8" pageNumber="501">Chicoreus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFEFE40FFF3CFF07FE57FE64" pageId="9" pageNumber="502" type="diagnosis">
<paragraph id="8B9836CCFFEFE40FFF3CFF07FE57FE64" blockId="9.[131,777,209,471]" pageId="9" pageNumber="502">
<smallCapsWord id="8D7EA010FFEFE40FFF3CFF07FF6EFF54" baselines="226,227" box="[132,239,209,232]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="9" pageNumber="502">DIAGNOSIS</smallCapsWord>
. — Biconic and middle-sized shell with
<taxonomicName id="4C274D4FFFEFE40FFDD3FF07FD45FF5A" authorityName="Montfort" authorityYear="1810" box="[619,708,209,233]" class="Gastropoda" family="Muricidae" genus="Chicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFEFE40FFDD3FF07FD45FF5A" box="[619,708,209,233]" italics="true" pageId="9" pageNumber="502">Chicoreus</emphasis>
</taxonomicName>
shape. Well marked primary cords. Spire: IP (infrasutural ramp), P1 to P3 (convex part of whorl). Last whorl: IP, P1 to P6 (convex part of whorl), P6 not atrophied, ADP, MP and ABP (siphonal canal). Nodulose protovarices on early whorls. Three varices with one or two intervarices on succeeding whorls. Varices slightly winged on last whorl. No cord spine even on P1. Shoulder sinus very short. Aperture ovate with strong internal denticles within outer lip: ID, D1 to D6. Columellar lip smooth. Parietal callus and anal notch present. Aragonite microstructure.
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFEFE40FFF3CFE21FC89F97E" pageId="9" pageNumber="502" type="discussion">
<paragraph id="8B9836CCFFEFE40FFF3CFE21FF69FDBC" blockId="9.[130,778,503,1744]" box="[132,232,503,529]" pageId="9" pageNumber="502">
<smallCapsWord id="8D7EA010FFEFE40FFF3CFE21FF69FDBC" baselines="522,522" box="[132,232,503,529]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="9" pageNumber="502">REMARKS</smallCapsWord>
</paragraph>
<paragraph id="8B9836CCFFEFE40FFF3CFDC0FC89F97E" blockId="9.[130,778,503,1744]" pageId="9" pageNumber="502">
During the Eocene several species of
<taxonomicName id="4C274D4FFFEFE40FFD9DFDC0FC82FD83" authority="Merle" authorityName="Merle" authorityYear="2024" box="[549,771,534,560]" class="Gastropoda" family="Muricidae" genus="Paleochicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B953EADEFFEFE40FFD9DFDC0FD38FD83" box="[549,697,534,560]" italics="true" pageId="9" pageNumber="502">Paleochicoreus</emphasis>
Merle
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFEFE40FFF3EFDE1FF50FDE2" box="[134,209,567,593]" pageId="9" pageNumber="502" rank="genus">n. gen.</taxonomicNameLabel>
appeared in Eastern Pacific (North American coast), Western Atlantic (North American coast) and Eastern Atlantic (West Africa and Europe), which have been attributed in the literature to
<taxonomicName id="4C274D4FFFEFE40FFEB9FD41FD97FD03" authority="Swainson, 1833" authorityName="Swainson" authorityYear="1833" box="[257,534,662,688]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFEFE40FFEB9FD41FEEBFD03" box="[257,362,663,688]" italics="true" pageId="9" pageNumber="502">Pterynotus</emphasis>
Swainson, 1833
</taxonomicName>
or
<taxonomicName id="4C274D4FFFEFE40FFD8EFD40FF3DFD63" authority="Montfort, 1810" authorityName="Montfort" authorityYear="1810" class="Gastropoda" family="Muricidae" genus="Chicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFEFE40FFD8EFD40FD16FD03" box="[566,663,662,688]" italics="true" pageId="9" pageNumber="502">Chicoreus</emphasis>
Montfort, 1810
</taxonomicName>
(
<bibRefCitation id="EFB64B3DFFEFE40FFF74FD60FED3FD63" author="EAMES F. E." box="[204,338,693,720]" pageId="9" pageNumber="502" pagination="25 - 70" refId="ref44211" refString="EAMES F. E. 1957. - Eocene Mollusca from Nigeria: a revision. Bulletin of the British Museum of the Natural History (Geology) 3 (2): 25 - 70. https: // doi. org / 10.5962 / p. 143489" type="journal article" year="1957">Eames 1957</bibRefCitation>
;
<bibRefCitation id="EFB64B3DFFEFE40FFEE7FD60FE5FFD63" author="VOKES E. H." box="[351,478,694,720]" pageId="9" pageNumber="502" pagination="1 - 141" refId="ref48907" refString="VOKES E. H. 1971. - Catalogue of the genus Murex Linne (Mollusca: Gastropoda); Muricinae, Ocenebrinae. Bulletins of American Paleontology 61 (268): 1 - 141." type="journal article" year="1971">Vokes 1971</bibRefCitation>
,
<bibRefCitation id="EFB64B3DFFEFE40FFE54FD60FDA5FD63" author="VOKES E. H." box="[492,548,694,720]" pageId="9" pageNumber="502" pagination="1 - 96" refId="ref48944" refString="VOKES E. H. 1990. - Cenozoic Muricidae of the Western Atlantic region, Part VIII - Murex s. s., Haustellum, Chicoreus, Hexaplex; additions and corrections. Tulane Studies in Geology and Paleontology 23 (1 - 3): 1 - 96." type="journal article" year="1990">1990</bibRefCitation>
;
<bibRefCitation id="EFB64B3DFFEFE40FFD8AFD60FD6AFD63" author="MERLE D. &amp; GARRIGUES B. &amp; POINTIER J. - P." box="[562,747,694,720]" pageId="9" pageNumber="502" refId="ref47234" refString="MERLE D., GARRIGUES B. &amp; POINTIER J. - P. 2011. - Fossil &amp; Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
<taxonomicName id="4C274D4FFFEFE40FFD8AFD60FDF0FD63" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[562,625,694,720]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
<emphasis id="B953EADEFFEFE40FFDC1FD60FD2CFD63" box="[633,685,694,720]" italics="true" pageId="9" pageNumber="502">et al.</emphasis>
2011
</bibRefCitation>
as?
<taxonomicName id="4C274D4FFFEFE40FFF35FD00FF6CFD43" authorityName="Montfort" authorityYear="1810" box="[141,237,726,752]" class="Gastropoda" family="Muricidae" genus="Chicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFEFE40FFF35FD00FF6CFD43" box="[141,237,726,752]" italics="true" pageId="9" pageNumber="502">Chicoreus</emphasis>
</taxonomicName>
). With
<emphasis id="B953EADEFFEFE40FFE82FD00FE23FD43" box="[314,418,726,752]" italics="true" pageId="9" pageNumber="502">
<taxonomicName id="4C274D4FFFEFE40FFE82FD00FE1CFD43" authorityName="Montfort" authorityYear="1810" box="[314,413,726,752]" class="Gastropoda" family="Muricidae" genus="Chicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus">Chicoreus</taxonomicName>
,
</emphasis>
they share a biconic shell, medium length of siphonal canal, trivaricate morphology, nodulose protovarices, an anal notch, and a parietal callus. However, these features are also shared with the genera
<taxonomicName id="4C274D4FFFEFE40FFDD0FCE0FE9CFCDC" authority="Jousseaume, 1880" authorityName="Jousseaume" authorityYear="1880" class="Gastropoda" family="Muricidae" genus="Naquetia" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFEFE40FFDD0FCE0FD48FCFC" box="[616,713,822,847]" italics="true" pageId="9" pageNumber="502">Naquetia</emphasis>
<bibRefCitation id="EFB64B3DFFEFE40FFD68FCE0FE9CFCDC" author="JOUSSEAUME F. P." pageId="9" pageNumber="502" pagination="335 - 338" refId="ref45378" refString="JOUSSEAUME F. P. 1880. - Division methodique de la famille des Purpurides. Le Naturaliste 2 (42): 335 - 338. http: // biodiversitylibrary. org / page / 33928744" type="journal article" year="1880">Jousseaume, 1880</bibRefCitation>
</taxonomicName>
,
<emphasis id="B953EADEFFEFE40FFE91FC83FE21FCDC" box="[297,416,853,879]" italics="true" pageId="9" pageNumber="502">Phyllonotus</emphasis>
Swainson, 1833 and
<taxonomicName id="4C274D4FFFEFE40FFD33FC83FEB1FC3C" authority="Arakawa, 1964" authorityName="Arakawa" authorityYear="1964" class="Gastropoda" family="Muricidae" genus="Chicomurex" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFEFE40FFD33FC83FC89FCDC" box="[651,776,853,879]" italics="true" pageId="9" pageNumber="502">Chicomurex</emphasis>
Arakawa, 1964
</taxonomicName>
.
<taxonomicName id="4C274D4FFFEFE40FFE87FCA3FDA1FC3C" authority="Merle" authorityName="Merle" authorityYear="2024" box="[319,544,885,911]" class="Gastropoda" family="Muricidae" genus="Paleochicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B953EADEFFEFE40FFE87FCA3FE54FC3C" box="[319,469,885,911]" italics="true" pageId="9" pageNumber="502">Paleochicoreus</emphasis>
Merle
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFEFE40FFD95FCA0FDFAFC23" box="[557,635,886,912]" pageId="9" pageNumber="502" rank="genus">n. gen.</taxonomicNameLabel>
differs from these three genera by having a smooth columellar lip. For example, according to
<bibRefCitation id="EFB64B3DFFEFE40FFECEFC63FD87FC7C" author="VOKES E. H." box="[374,518,949,975]" pageId="9" pageNumber="502" pagination="1 - 96" refId="ref48944" refString="VOKES E. H. 1990. - Cenozoic Muricidae of the Western Atlantic region, Part VIII - Murex s. s., Haustellum, Chicoreus, Hexaplex; additions and corrections. Tulane Studies in Geology and Paleontology 23 (1 - 3): 1 - 96." type="journal article" year="1990">Vokes (1990)</bibRefCitation>
,
<taxonomicName id="4C274D4FFFEFE40FFDACFC63FF55FC5C" authorityName="Vokes" authorityYear="1990" class="Gastropoda" family="Muricidae" genus="Chicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="species" species="initialis" subGenus="Phyllonotus">
<emphasis id="B953EADEFFEFE40FFDACFC63FDF9FC7C" box="[532,632,949,975]" italics="true" pageId="9" pageNumber="502">Chicoreus</emphasis>
(
<emphasis id="B953EADEFFEFE40FFD3FFC63FD7FFC7C" box="[647,766,949,975]" italics="true" pageId="9" pageNumber="502">Phyllonotus</emphasis>
)
<emphasis id="B953EADEFFEFE40FFF3CFC03FF55FC5C" box="[132,212,981,1007]" italics="true" pageId="9" pageNumber="502">initialis</emphasis>
</taxonomicName>
(herein
<taxonomicName id="4C274D4FFFEFE40FFE95FC03FD95FC5C" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" baseAuthorityName="Vokes" baseAuthorityYear="1990" box="[301,532,981,1007]" class="Gastropoda" family="Muricidae" genus="Paleochicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="species" species="initialis" status="comb. nov.">
<emphasis id="B953EADEFFEFE40FFE95FC03FD95FC5C" box="[301,532,981,1007]" italics="true" pageId="9" pageNumber="502">Paleochicoreus initialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFEFE40FFDA2FC03FDFAFC5C" box="[538,635,981,1007]" pageId="9" pageNumber="502" rank="species">n. comb.</taxonomicNameLabel>
) (
<figureCitation id="131C2A49FFEFE40FFD29FC03FD7EFC5C" box="[657,767,981,1007]" captionStart="FIG" captionStartId="8.[132,143,1516,1533]" captionTargetBox="[132,1455,215,1472]" captionTargetId="figure-286@8.[698,1267,753,1294]" captionTargetPageId="8" captionText="FIG. 2. — Spiral cords and internal denticles of the outer lip in Paleochicoreus Merle, n. gen.: A, B, Paleoochicoreus initialis (Vokes, 1990), n. comb., holotype, USNM450376, Nacodoches, Texas (United States), middle Eocene (Lutetian); C, P. newtoni (Eames, 1957), holotype, BMNH G41982, Ombralla, Nigeria, middle Eocene;D, E, P. bicostatus (Deshayes,1835), n. comb., holotype, UCBL-EM 33437 (Deshayes coll.), Valmondois,Val dOise, Paris Basin, France, middle Eocene (Bartonian). Scale bars: 10 mm. Credits: A, B, Y. Villacampa and J. Jorstad (USNM); C, S. Tracey (NHM); D, E, P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985764" httpUri="https://zenodo.org/record/13985764/files/figure.png" pageId="9" pageNumber="502">Fig. 2A, B</figureCitation>
) is most similar to members of
<emphasis id="B953EADEFFEFE40FFE74FC23FDC3FBBC" box="[460,578,1013,1039]" italics="true" pageId="9" pageNumber="502">Phyllonotus</emphasis>
, because of developed internal denticles of the outer lip, but it lacks the derived characters of
<emphasis id="B953EADEFFEFE40FFEB3FBE2FE01FBFD" box="[267,384,1076,1102]" italics="true" pageId="9" pageNumber="502">Phyllonotus</emphasis>
(rugae and expanded parietal shield).
<taxonomicName id="4C274D4FFFEFE40FFF3AFB82FEEAFBDD" baseAuthorityName="Eames" baseAuthorityYear="1957" box="[130,363,1108,1134]" class="Gastropoda" family="Muricidae" genus="Paleochicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="species" species="newtoni">
<emphasis id="B953EADEFFEFE40FFF3AFB82FEEAFBDD" box="[130,363,1108,1134]" italics="true" pageId="9" pageNumber="502">Paleochicoreus newtoni</emphasis>
</taxonomicName>
(
<figureCitation id="131C2A49FFEFE40FFEC1FB82FE49FBDC" box="[377,456,1108,1135]" captionStart="FIG" captionStartId="8.[132,143,1516,1533]" captionTargetBox="[132,1455,215,1472]" captionTargetId="figure-286@8.[698,1267,753,1294]" captionTargetPageId="8" captionText="FIG. 2. — Spiral cords and internal denticles of the outer lip in Paleochicoreus Merle, n. gen.: A, B, Paleoochicoreus initialis (Vokes, 1990), n. comb., holotype, USNM450376, Nacodoches, Texas (United States), middle Eocene (Lutetian); C, P. newtoni (Eames, 1957), holotype, BMNH G41982, Ombralla, Nigeria, middle Eocene;D, E, P. bicostatus (Deshayes,1835), n. comb., holotype, UCBL-EM 33437 (Deshayes coll.), Valmondois,Val dOise, Paris Basin, France, middle Eocene (Bartonian). Scale bars: 10 mm. Credits: A, B, Y. Villacampa and J. Jorstad (USNM); C, S. Tracey (NHM); D, E, P. Loubry (MNHN/CNRS)." figureDoi="http://doi.org/10.5281/zenodo.13985764" httpUri="https://zenodo.org/record/13985764/files/figure.png" pageId="9" pageNumber="502">Fig. 2C</figureCitation>
) possesses an expanded varical flange, like that in
<taxonomicName id="4C274D4FFFEFE40FFEEBFBA2FDBDFB3D" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" baseAuthorityName="Vokes" baseAuthorityYear="1990" box="[339,572,1140,1166]" class="Gastropoda" family="Muricidae" genus="Paleochicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="species" species="initialis" status="comb. nov.">
<emphasis id="B953EADEFFEFE40FFEEBFBA2FDBDFB3D" box="[339,572,1140,1166]" italics="true" pageId="9" pageNumber="502">Paleochicoreus initialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFEFE40FFDFBFBA3FD26FB3D" box="[579,679,1140,1167]" pageId="9" pageNumber="502" rank="species">n. comb.</taxonomicNameLabel>
, but this character is lacking in many
<taxonomicName id="4C274D4FFFEFE40FFE78FB42FDFDFB1D" box="[448,636,1172,1198]" class="Gastropoda" family="Muricidae" genus="Chicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B953EADEFFEFE40FFE78FB42FDA5FB1D" box="[448,548,1172,1198]" italics="true" pageId="9" pageNumber="502">Chicoreus</emphasis>
species.
</taxonomicName>
Members of
<taxonomicName id="4C274D4FFFEFE40FFF3EFB62FECAFB7D" authorityName="G. Perry" authorityYear="1810" box="[134,331,1204,1230]" class="Gastropoda" family="Muricidae" genus="Chicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="subGenus" subGenus="Triplex">
<emphasis id="B953EADEFFEFE40FFF3EFB62FF6BFB7D" box="[134,234,1204,1230]" italics="true" pageId="9" pageNumber="502">Chicoreus</emphasis>
(
<emphasis id="B953EADEFFEFE40FFF41FB62FEC3FB7D" box="[249,322,1204,1230]" italics="true" pageId="9" pageNumber="502">Triplex</emphasis>
)
</taxonomicName>
and
<emphasis id="B953EADEFFEFE40FFE3CFB62FE1CFB7D" box="[388,413,1204,1230]" italics="true" pageId="9" pageNumber="502">C.</emphasis>
(
<taxonomicName id="4C274D4FFFEFE40FFE15FB62FDE1FB7D" authorityName="Oyama" authorityYear="1950" box="[429,608,1204,1230]" class="Gastropoda" family="Muricidae" genus="Rhizophorimurex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFEFE40FFE15FB62FDE1FB7D" box="[429,608,1204,1230]" italics="true" pageId="9" pageNumber="502">Rhizophorimurex</emphasis>
</taxonomicName>
) also bear similarities with those of
<taxonomicName id="4C274D4FFFEFE40FFED6FB02FDC8FB5D" authority="Merle" authorityName="Merle" authorityYear="2024" box="[366,585,1236,1262]" class="Gastropoda" family="Muricidae" genus="Paleochicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B953EADEFFEFE40FFED6FB02FD81FB5D" box="[366,512,1236,1262]" italics="true" pageId="9" pageNumber="502">Paleochicoreus</emphasis>
Merle
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFEFE40FFDECFB03FD20FB5C" box="[596,673,1237,1263]" pageId="9" pageNumber="502" rank="genus">n. gen.</taxonomicNameLabel>
, but they lack an expanded varical flange, their internal denticles are reduced or missing, and they often develop spiny processes.
<taxonomicName id="4C274D4FFFEFE40FFF3DFAE2FF68FAFD" authorityName="Montfort" authorityYear="1810" box="[133,233,1332,1358]" class="Gastropoda" family="Muricidae" genus="Chicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFEFE40FFF3DFAE2FF68FAFD" box="[133,233,1332,1358]" italics="true" pageId="9" pageNumber="502">Chicoreus</emphasis>
</taxonomicName>
(
<emphasis id="B953EADEFFEFE40FFF40FAE2FE94FAFE" box="[248,277,1332,1357]" italics="true" pageId="9" pageNumber="502">s.s.</emphasis>
), which is rather similar to
<emphasis id="B953EADEFFEFE40FFD84FAE2FDD4FAFD" box="[572,597,1332,1358]" italics="true" pageId="9" pageNumber="502">C.</emphasis>
(
<emphasis id="B953EADEFFEFE40FFDDDFAE2FD2FFAFD" box="[613,686,1332,1358]" italics="true" pageId="9" pageNumber="502">Triplex</emphasis>
), differs by having a labral groove spine placed between P4 and P5 (
<bibRefCitation id="EFB64B3DFFEFE40FFF36FAA5FED0FA3D" author="MERLE D. &amp; GARRIGUES B. &amp; POINTIER J. - P." box="[142,337,1395,1422]" pageId="9" pageNumber="502" refId="ref47234" refString="MERLE D., GARRIGUES B. &amp; POINTIER J. - P. 2011. - Fossil &amp; Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
<taxonomicName id="4C274D4FFFEFE40FFF36FAA5FF4EFA3E" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[142,207,1395,1421]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
<emphasis id="B953EADEFFEFE40FFF60FAA2FE8EFA3E" box="[216,271,1395,1421]" italics="true" pageId="9" pageNumber="502">et al.</emphasis>
2011
</bibRefCitation>
: fig. 37A, B), lacking in
<taxonomicName id="4C274D4FFFEFE40FFDCBFAA5FF47FA1E" authority="Merle" authorityName="Merle" authorityYear="2024" class="Gastropoda" family="Muricidae" genus="Paleochicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B953EADEFFEFE40FFDCBFAA5FC88FA3E" box="[627,777,1395,1421]" italics="true" pageId="9" pageNumber="502">Paleochicoreus</emphasis>
Merle
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFEFE40FFF77FA42FE99FA1D" box="[207,280,1428,1454]" pageId="9" pageNumber="502" rank="genus">n. gen.</taxonomicNameLabel>
Species such as
<taxonomicName id="4C274D4FFFEFE40FFE04FA45FD22FA1E" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" baseAuthorityName="Vokes" baseAuthorityYear="1990" box="[444,675,1427,1453]" class="Gastropoda" family="Muricidae" genus="Paleochicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="species" species="initialis" status="comb. nov.">
<emphasis id="B953EADEFFEFE40FFE04FA45FD22FA1E" box="[444,675,1427,1453]" italics="true" pageId="9" pageNumber="502">Paleochicoreus initialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFEFE40FFD11FA42FC89FA1E" box="[681,776,1427,1454]" pageId="9" pageNumber="502" rank="species">n. comb.</taxonomicNameLabel>
and
<taxonomicName id="4C274D4FFFEFE40FFF09FA62FE9FFA7E" baseAuthorityName="Eames" baseAuthorityYear="1957" box="[177,286,1460,1486]" class="Gastropoda" family="Muricidae" genus="Paleochicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="species" species="newtoni">
<emphasis id="B953EADEFFEFE40FFF09FA62FF41FA7E" box="[177,192,1460,1485]" italics="true" pageId="9" pageNumber="502">P</emphasis>
.
<emphasis id="B953EADEFFEFE40FFF74FA62FE9FFA7E" box="[204,286,1460,1485]" italics="true" pageId="9" pageNumber="502">newtoni</emphasis>
</taxonomicName>
represent ideal ancestral lineages for
<taxonomicName id="4C274D4FFFEFE40FFD23FA65FD7FFA7E" authorityName="Montfort" authorityYear="1810" box="[667,766,1459,1485]" class="Gastropoda" family="Muricidae" genus="Chicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFEFE40FFD23FA65FD7FFA7E" box="[667,766,1459,1485]" italics="true" pageId="9" pageNumber="502">Chicoreus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C274D4FFFEFE40FFF3CFA05FE80FA5E" authorityName="Arakawa" authorityYear="1964" box="[132,257,1491,1517]" class="Gastropoda" family="Muricidae" genus="Chicomurex" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFEFE40FFF3CFA05FE80FA5E" box="[132,257,1491,1517]" italics="true" pageId="9" pageNumber="502">Chicomurex</emphasis>
</taxonomicName>
or
<emphasis id="B953EADEFFEFE40FFE90FA05FE1EFA5E" box="[296,415,1491,1517]" italics="true" pageId="9" pageNumber="502">Phyllonotus</emphasis>
(
<bibRefCitation id="EFB64B3DFFEFE40FFE08FA05FDB1FA5E" author="VOKES E. H." box="[432,560,1491,1517]" pageId="9" pageNumber="502" pagination="1 - 96" refId="ref48944" refString="VOKES E. H. 1990. - Cenozoic Muricidae of the Western Atlantic region, Part VIII - Murex s. s., Haustellum, Chicoreus, Hexaplex; additions and corrections. Tulane Studies in Geology and Paleontology 23 (1 - 3): 1 - 96." type="journal article" year="1990">Vokes 1990</bibRefCitation>
) but cannot readily be assign to one of these branches. This comment is true for other species attributed here to
<taxonomicName id="4C274D4FFFEFE40FFE6FF9C5FD33F99E" authority="Merle" authorityName="Merle" authorityYear="2024" box="[471,690,1555,1581]" class="Gastropoda" family="Muricidae" genus="Paleochicoreus" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B953EADEFFEFE40FFE6FF9C5FDE8F99E" box="[471,617,1555,1581]" italics="true" pageId="9" pageNumber="502">Paleochicoreus</emphasis>
Merle
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFEFE40FFD05F9C2FC86F99D" box="[701,775,1556,1582]" pageId="9" pageNumber="502" rank="genus">n. gen.</taxonomicNameLabel>
Here, we figure and give a new description of the
<typeStatus id="549C886EFFEFE40FFD10F9E5FC86F9FE" box="[680,775,1587,1613]" pageId="9" pageNumber="502" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C274D4FFFEFE40FFF1CF982FE73F9DD" authority="(Deshayes, 1835)" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" baseAuthorityName="Deshayes" baseAuthorityYear="1835" box="[164,498,1619,1646]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="502" phylum="Mollusca" rank="species" species="bicostatus" status="n. comb.">
<emphasis id="B953EADEFFEFE40FFF1CF982FEA5F9DE" box="[164,292,1619,1645]" italics="true" pageId="9" pageNumber="502">P. bicostatus</emphasis>
(
<bibRefCitation id="EFB64B3DFFEFE40FFE8DF985FE6BF9DD" author="DESHAYES G. - P." box="[309,490,1619,1646]" pageId="9" pageNumber="502" pagination="79 - 106" refId="ref43930" refString="DESHAYES G. - P. 1835. - Description des coquilles fossiles des environs de Paris. Tome 2. Livraisons 40 - 45, l'auteur, Bechet Jeune, Baudouin freres, Treuttel &amp; Wurtz, Paris: 499 - 780; Atlas 2, Levrault, Paris: pl. 79 - 106. https: // doi. org / 10.5962 / bhl. title. 52303" type="book chapter" year="1835">Deshayes, 1835</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A26057A5FFEFE40FFDB8F985FDE4F9DE" box="[512,613,1619,1645]" pageId="9" pageNumber="502" rank="species">n. comb.</taxonomicNameLabel>
This very rare species has not been found subsequent to its original description. Information given hereunder is based on the
<typeStatus id="549C886EFFEFE40FFD10F944FC86F91F" box="[680,775,1682,1708]" pageId="9" pageNumber="502" type="holotype">holotype</typeStatus>
retrieved from the collections of the École des Mines, Lyon.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,480 @@
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<mods:title id="CFDA5C52D281337F7A2F69D80C66A029">A new cave-dwelling hadzioid amphipod (Senticaudata, Hadzioidea, Melitidae) from sulFIdic groundwaters in Iran</mods:title>
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<mods:namePart id="D5ED5B911E0001BDD72273B41E92358F">Kuntner, Matjaž</mods:namePart>
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<mods:namePart id="0B8FC0C086AC0812AD917F5050E988E1">Fišer, Cene</mods:namePart>
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<mods:date id="B90B605E274CBE6B0C0081C9AADA75CE">2024</mods:date>
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<subSubSection id="C3208BCD6A440C7CFF34FAAEFE70FA05" box="[129,441,1392,1419]" pageId="6" pageNumber="113" type="nomenclature">
<paragraph id="8B85D8466A440C7CFF34FAAEFE70FA05" blockId="6.[129,640,1141,1741]" box="[129,441,1392,1419]" pageId="6" pageNumber="113">
<taxonomicName id="4C3AA3C56A440C7CFF34FAAEFEA8FA05" authorityName="Malek-Hosseini &amp; Brad &amp; Fatemi &amp; Kuntner &amp; Fišer" authorityYear="2024" box="[129,353,1392,1419]" class="Malacostraca" family="Melitidae" genus="Tegano" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="tashanensis" status="sp. nov.">
<emphasis id="B94E04546A440C7CFF34FAAEFEA8FA05" bold="true" box="[129,353,1392,1419]" italics="true" pageId="6" pageNumber="113">Tegano tashanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DB92F6A440C7CFED3FAAEFE70FA05" box="[358,441,1393,1419]" pageId="6" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C3208BCD6A440C7CFF34FA4BFF25FA78" pageId="6" pageNumber="113" type="description">
<paragraph id="8B85D8466A440C7CFF34FA4BFD49FA20" blockId="6.[129,640,1141,1741]" box="[129,640,1428,1454]" pageId="6" pageNumber="113">https://zoobank.org/NomenclaturalActs/614E</paragraph>
<paragraph id="8B85D8466A440C7CFF34FA67FDD6FA5D" blockId="6.[129,640,1141,1741]" box="[129,543,1464,1491]" pageId="6" pageNumber="113">536F-598E-4B12-8818-DEAF2F6026F7</paragraph>
<paragraph id="8B85D8466A440C7CFF34FA03FF25FA78" blockId="6.[129,640,1141,1741]" box="[129,236,1500,1526]" pageId="6" pageNumber="113">
<figureCitation id="1301C4C36A440C7CFF34FA03FF0FFA78" box="[129,198,1500,1526]" captionStart="FIGURE 4" captionStartId="10.[130,197,1662,1683]" captionTargetBox="[129,646,1055,1632]" captionTargetId="figure-305@10.[129,642,1055,1632]" captionTargetPageId="10" captionText="FIGURE 4 Photographs of Tegano tashanensis sp. nov.:male (A) and female (B) living in Chah Kabootari Cave, Iran." pageId="6" pageNumber="113">Figs. 4</figureCitation>
10
</paragraph>
</subSubSection>
<subSubSection id="C3208BCD6A440C7CFF2AF9DEFD12F943" pageId="6" pageNumber="113" type="diagnosis">
<paragraph id="8B85D8466A440C7CFF2AF9DEFD12F943" blockId="6.[129,640,1141,1741]" lastBlockId="6.[676,1219,1141,1746]" pageId="6" pageNumber="113">
<emphasis id="B94E04546A440C7CFF2AF9DEFEC2F994" box="[159,267,1537,1562]" italics="true" pageId="6" pageNumber="113">Diagnosis</emphasis>
. Small-sized stygobiontic amphipod, with robust appearance and clear troglomorphic features. Eyeless, unpigmented, least setose, long antennae and legs relative to body length, labium without inner lobes, mandibles without mandibular palps, exopodite
<heading id="D0CD6F2A6A440C7CFD11FBAAFB6AFB01" box="[676,1187,1141,1167]" fontSize="11" level="2" pageId="6" pageNumber="113" reason="2">of uropod III unsegmented. Pleonites 2 and</heading>
3 with 2 small setae on the postero-dorsal sides. One seta on each postero-dorsal side of the first two urosomites. Antenna I two thirds of the total body length. Outer lobe of maxilla I with 8 comb-like spines. Inner lobe of maxilliped with 3 stout, 3 serrated and 3 regular setae located apically and 3 serrated setae subapically. Gnathopods scarcely setose, highly dissimilar, propodite of gnathopod I less than half length of propodite of gnathopod II. Uropods I and II with slightly shorter exopodites than corresponding endopodites. Pereopods VI and VII of equal size, more than half of body length. Telson as long as wide, with 4 short apical setae on each lobe.
</paragraph>
</subSubSection>
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Downloaded from Brill.com
<date id="FF84FE866A440C7CFC4CF91CFB90F95C" box="[1017,1113,1731,1746]" pageId="6" pageNumber="113" value="2024-10-07">07/10/2024</date>
01:59:59PM
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</paragraph>
<paragraph id="8B85D8466A440C7CFE47F907FB0AF89C" blockId="6.[498,1220,1752,1810]" pageId="6" pageNumber="113">via Open Access. This is an open access article distributed under the terms of the CC BY 4.0 license. https://creativecommons.org/licenses/by/4.0/</paragraph>
<subSubSection id="C3208BCD6A450C7DFF34FF74FE70FE6B" pageId="7" pageNumber="114" type="materials_examined">
<paragraph id="8B85D8466A450C7DFF34FF74FE60FEF7" blockId="7.[98,610,171,1706]" pageId="7" pageNumber="114">
<emphasis id="B94E04546A450C7DFF34FF74FE94FF4B" box="[129,349,171,197]" italics="true" pageId="7" pageNumber="114">Material examined.</emphasis>
<materialsCitation id="3B52D21B6A450C7DFEDDFF74FE60FEF7" collectingDate="2018-03-17" country="Iran" county="Zagros Mountains" elevation="539" latitude="30.8694444" location="Chah Kabootari Cave" longitude="50.166944" municipality="Behbahan county" pageId="7" pageNumber="114" specimenCount="1" specimenCount-male="1" stateProvince="Khuzestan Province" typeStatus="holotype">
<emphasis id="B94E04546A450C7DFEDDFF74FE00FF4B" box="[360,457,171,197]" italics="true" pageId="7" pageNumber="114">
<typeStatus id="548166E46A450C7DFEDDFF74FE00FF4B" box="[360,457,171,197]" pageId="7" pageNumber="114" type="holotype">Holotype</typeStatus>
</emphasis>
:
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(fig. 4A),
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,
<location id="8EE58E9D6A450C7DFE31FF10FD94FF67" LSID="urn:lsid:plazi:treatment:039369506A440C70FF34FAAEFEDDFD6E:8EE58E9D6A450C7DFE31FF10FD94FF67" box="[388,605,207,233]" country="Iran" county="Zagros Mountains" municipality="Behbahan county" name="Sarjowshar village" pageId="7" pageNumber="114" stateProvince="Khuzestan Province">Sarjowshar village</location>
,
<location id="8EE58E9D6A450C7DFFD7FF2CFEC2FE83" LSID="urn:lsid:plazi:treatment:039369506A440C70FF34FAAEFEDDFD6E:8EE58E9D6A450C7DFFD7FF2CFEC2FE83" box="[98,267,243,269]" country="Iran" county="Zagros Mountains" municipality="Behbahan county" name="Tashan district" pageId="7" pageNumber="114" stateProvince="Khuzestan Province">Tashan district</location>
,
<collectingMunicipality id="6BE1423C6A450C7DFEA2FF2CFE16FE80" box="[279,479,243,270]" pageId="7" pageNumber="114">Behbahan county</collectingMunicipality>
,
<collectingRegion id="49FE16A46A450C7DFE5FFF2CFF0FFEBF" country="Iran" name="Khuzestan" pageId="7" pageNumber="114">Khuzestan Province</collectingRegion>
,
<collectingCounty id="62E4A0CA6A450C7DFF69FEC7FE7CFEBF" box="[220,437,279,306]" pageId="7" pageNumber="114">Zagros Mountains</collectingCounty>
, southwestern
<collectingCountry id="F32D98D66A450C7DFFD7FEE3FF59FED8" box="[98,144,316,342]" name="Iran" pageId="7" pageNumber="114">Iran</collectingCountry>
(30°5210N, 50°1001E, altitude
<elevation id="00173F756A450C7DFDADFEE4FF5BFEF7" metricMagnitude="2" metricUnit="m" metricValue="5.39" pageId="7" pageNumber="114" unit="m" value="539.0">
<quantity id="4CC275A36A450C7DFDADFEE4FDABFED8" box="[536,610,315,342]" metricMagnitude="2" metricUnit="m" metricValue="5.39" pageId="7" pageNumber="114" unit="m" value="539.0">539 m</quantity>
a.s.l.
</elevation>
), collected on
<date id="FF84FE866A450C7DFE83FE80FE6CFEF7" box="[310,421,351,377]" pageId="7" pageNumber="114" value="2018-03-17">
<collectingDate id="EFC0076E6A450C7DFE83FE80FE6CFEF7" box="[310,421,351,377]" pageId="7" pageNumber="114" value="2018-03-17">17.03.2018</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
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<emphasis id="B94E04546A450C7DFF34FE5BFF24FE13" box="[129,237,388,413]" italics="true" pageId="7" pageNumber="114">
<typeStatus id="548166E46A450C7DFF34FE5BFF24FE13" box="[129,237,388,413]" pageId="7" pageNumber="114" type="paratype">Paratypes</typeStatus>
</emphasis>
: Locality as same as
<typeStatus id="548166E46A450C7DFE4DFE5CFD94FE13" box="[504,605,387,413]" pageId="7" pageNumber="114" type="holotype">holotype</typeStatus>
, one dissected
<specimenCount id="9D3C13CF6A450C7DFEB2FE78FEDEFE4F" box="[263,279,423,449]" count="1" pageId="7" pageNumber="114" type="female"></specimenCount>
(fig. 4B),
<specimenCount id="9D3C13CF6A450C7DFE3CFE78FE67FE4F" box="[393,430,423,449]" count="1" pageId="7" pageNumber="114" type="male">1 ♂</specimenCount>
and
<specimenCount id="9D3C13CF6A450C7DFE59FE78FDD8FE4F" box="[492,529,423,449]" count="2" pageId="7" pageNumber="114" type="female">2 ♀</specimenCount>
undissected; collected on
<date id="FF84FE866A450C7DFEF1FE14FE7CFE6B" box="[324,437,459,485]" pageId="7" pageNumber="114" value="2018-08-12">08.12.2018</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C3208BCD6A450C7DFF34FE30FE56FD16" pageId="7" pageNumber="114" type="discussion">
<paragraph id="8B85D8466A450C7DFF34FE30FE56FD16" blockId="7.[98,610,171,1706]" pageId="7" pageNumber="114">
<emphasis id="B94E04546A450C7DFF34FE30FF1FFD87" box="[129,214,495,521]" italics="true" pageId="7" pageNumber="114">Remark</emphasis>
. The morphological description was performed on the body appendages belonging to the dissected
<typeStatus id="548166E46A450C7DFE04FDE9FDDCFDDE" box="[433,533,566,592]" pageId="7" pageNumber="114" type="holotype">holotype</typeStatus>
male. Sexual dimorphism was described by inspection of
<specimenCount id="9D3C13CF6A450C7DFF06FDA0FE52FD16" box="[179,411,638,665]" count="1" pageId="7" pageNumber="114" type="female" typeStatus="paratype">one paratype female</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="C3208BCD6A450C73FF34FD7CFB4CFEF7" lastPageId="9" lastPageNumber="116" pageId="7" pageNumber="114" type="description">
<paragraph id="8B85D8466A450C7DFF34FD7CFEB9FCC2" blockId="7.[98,610,171,1706]" pageId="7" pageNumber="114">
<emphasis id="B94E04546A450C7DFF34FD7CFE9EFD32" box="[129,343,674,700]" italics="true" pageId="7" pageNumber="114">Description (male):</emphasis>
The total body length of the male is
<quantity id="4CC275A36A450C7DFEB9FD19FE94FD6F" box="[268,349,710,737]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="7" pageNumber="114" unit="mm" value="4.7">4.7 mm</quantity>
(
<figureCitation id="1301C4C36A450C7DFEDBFD19FE74FD6F" box="[366,445,710,737]" captionStart="FIGURE 5" captionStartId="10.[676,743,1662,1683]" captionTargetBox="[653,1183,1055,1632]" captionTargetId="figure-325@10.[676,1183,1096,1620]" captionText="FIGURE 5 Tegano tashanensis sp.nov.:(A) holotype, male (4.7 mm), habitus, lateral view; (B Downloaded) epimeralfrom Brill plates.com I-III." pageId="7" pageNumber="114">fig. 5A</figureCitation>
). A chart containing detailed measurements on all diagnosis-relevant body appendages is presented in supplementary table S4.
</paragraph>
<paragraph id="8B85D8466A450C7DFF34FC89FF02FC36" blockId="7.[98,610,171,1706]" pageId="7" pageNumber="114">
<emphasis id="B94E04546A450C7DFF34FC89FF75FCFE" box="[129,188,854,880]" italics="true" pageId="7" pageNumber="114">Head</emphasis>
. Head (
<figureCitation id="1301C4C36A450C7DFEAFFC89FEA1FCFE" box="[282,360,853,880]" captionStart="FIGURE 5" captionStartId="10.[676,743,1662,1683]" captionTargetBox="[653,1183,1055,1632]" captionTargetId="figure-325@10.[676,1183,1096,1620]" captionText="FIGURE 5 Tegano tashanensis sp.nov.:(A) holotype, male (4.7 mm), habitus, lateral view; (B Downloaded) epimeralfrom Brill plates.com I-III." pageId="7" pageNumber="114">fig. 5A</figureCitation>
) with no rostrum; 10 % of the total body length (supplementary table S4).
</paragraph>
<paragraph id="8B85D8466A450C7DFF34FC1DFF25F994" blockId="7.[98,610,171,1706]" pageId="7" pageNumber="114">
<emphasis id="B94E04546A450C7DFF34FC1DFF22FC52" box="[129,235,962,988]" italics="true" pageId="7" pageNumber="114">Antennae</emphasis>
. Antenna I (fig. 6A) about two thirds of the total body length, with main flagellum formed of 20 articles. Antenna I articles with short aestethascs, one third of the respective article. Antenna I peduncle 41 % of the total length of antenna I. Accessory flagellum formed of 1 article and does not exceed the length of the first article of the main flagellum (fig. 6C). Accessory flagellum with 6 apical setae and 1 serrate seta located subapically. A more frequent shape of the accessory flagellum is that present in the
<typeStatus id="548166E46A450C7DFE75FA92FDEAFAE9" box="[448,547,1357,1383]" pageId="7" pageNumber="114" type="paratype">paratype</typeStatus>
male and females (fig. 6D). Here, accessory flagellum is longer, and it is formed of 2 articles.The distal article represents only 5 % of the total length of the accessory flagellum and it bears 2 long setae.
</paragraph>
<paragraph id="8B85D8466A450C7DFF34F9FBFE4AF924" blockId="7.[98,610,171,1706]" pageId="7" pageNumber="114">Antenna II (fig. 6A) with the flagellum formed of 10 articles; less than half of the total length of antenna I. The peduncle 60 % of the total length of antenna II.</paragraph>
<paragraph id="8B85D8466A450C7DFD11FF74FC12F927" blockId="7.[645,1157,171,1705]" pageId="7" pageNumber="114">
<emphasis id="B94E04546A450C7DFD11FF74FCECFF4B" box="[676,805,171,197]" italics="true" pageId="7" pageNumber="114">Mouthparts</emphasis>
. Labrum (fig. 6G) trapezoidal, with the width twice as much as its depth. Labium (fig. 6H) without inner lobes. The outer lobes each with one row of fine setae on the outer side. Maxilla I (fig. 6E) with 6 apical and 2 subapical setae on the distal article of the palp. The outer lobe with 8 comb-like spines apically. Inner lobe with 6 apical serrate setae and a row of lateral fine setae. Maxilla II (fig. 6F) with the outer lobe ¾ of the inner lobe length. Both lobes with a row of apical setae of various lengths. Outer lobe with a row of fine setae on the outer margin. The outer margin of inner lobe with one row of fine setae and one row of plumose setae. Mandibles lack palps. Left mandible (figs. 7I7J) with the incisor process formed of two lamellas with 3 and 4 teeth. Lacinia mobilis with 4 teeth; 4 strong serrated setae between lacinia mobilis and molar process. Right mandible (fig. 6K) with 6 teeth on incisor process. Spiniform lacinia mobilis with 2 teeth. 4 larger serrated setae and a row of fine setae between lacinia mobilis and molar process. The molar process present 12 toothlike structures and an adjacent serrated seta. Maxilliped (fig. 7H) with the palp formed of 4 articles little setose. Article 1 with 1 seta on the inner margin. Article 2 with 13 setae (of which one serrated) aligned along the inner margin over the whole length of the article, and one seta located subapically on the outer margin. Article 3 with one group of 4 apical setae and two groups of 5 setae each on both inner and outer margins. Article 4 with a group of 4 fine setae on the ventral margin and 1 seta on the dorsal margin. The outer lobe of maxilliped (fig. 7I) with 4 serrated setae and 3 flattened spines located apically, a row of 7 setae and 7 denticles aligned over the inner margin. The inner lobe is provided apically with 4 serrated setae, 3 flattened spines and 3 more regular setae. 2 serrated setae located subapically on the inner margin of inner lobe.
</paragraph>
<paragraph id="8B85D8466A4A0C72FF2AFF74FEEBFC39" blockId="8.[129,640,171,1706]" pageId="8" pageNumber="115">
<emphasis id="B94E04546A4A0C72FF2AFF74FEECFF4B" box="[159,293,171,197]" italics="true" pageId="8" pageNumber="115">Gnathopods</emphasis>
. Gnathopod I (fig. 7A) with the coxal plate in the shape of a rectangular trapezoid. The depth of coxal plate I larger than its width (ratio depth:width 1.0:0.5). Basipodite length:width ratio 1.0:0.3. Ischiopodite with one posteroventral group of 4 setae. Basipodite length:carpopodite length ratio 1.0:0.6. Meropodite with a row of fine setae along the whole ventral margin and 4 longer apical setae. Carpopodite with a row of 8 setae located posteroventral, a group of 4 setae located apically and 1 anterodorsal seta. Propodite (fig. 7C) with the length: width ratio 1.0:0.6. Propodite least setose, 2 setae on the ventral margin, a group of 4 setae on the dorsal margin and one group of 6 setae located antero-apically. 2 setae on the lateral surface and 2 small setae in the vicinity of the palmar corner. Palmar spines absent. Dactylopodite with the nail one quarter of the total dactylopodite length. 1 seta on the outer margin of dactylopodite.
</paragraph>
<paragraph id="8B85D8466A4A0C72FF2AFC1EFC80FE6B" blockId="8.[129,640,171,1706]" lastBlockId="8.[676,1188,171,1706]" pageId="8" pageNumber="115">
Gnathopod II (fig. 7B) highly different compared to gnathopod I. Ovoid coxal plate with the depth: width ratio 1.0:0.6. Basipodite length:width ratio 1.0:0.35. Ischiopodite with on posteroventral group of 3 small setae. Basipodite: carpopodite length ratio 1.0:0.55. Carpopodite with 5 groups of 24 setae along its ventral margin, 1 and 2 setae located apically on each side, 1 small seta on the dorsal margin. Long propodite (fig. 7D), length:width ratio1.0:0.7.Propodite gnathopod II:propodite gnathopod I length ratio 1.0:0.25. Propodite with eight groups of 12 setae in its ventral margin, four groups of 23 setae on the dorsal margin and one group of 4 setae located antero-apical. On its surface, the propodite has two median groups of 2 setae, and one group of 2 setae close to the palmar corner. Palmar spines missing. Short dactylopodite nail, representing 15% of the total dactylopodite length. One seta on the outer margin of dactylopodite.
<emphasis id="B94E04546A4A0C72FD77FF74FCF8FF4B" box="[706,817,171,197]" italics="true" pageId="8" pageNumber="115">Pereopods</emphasis>
. Pereopod III (fig. 7E) with a relative rectangular coxal plate, ratio depth:width 1.0:0.55. Dactylopodite with the nail measuring one third of the total length of the dactylopodite, one dorsal seta with plumose tip and one fine seta at the nail insertion. Propodite: dactylopodite length ratio 1.0:0.35. Pereopod III nearly equal in length to pereopod IV (ratio 1.0:0.97).
</paragraph>
<paragraph id="8B85D8466A4A0C72FD77FE31FD32FC8B" blockId="8.[676,1188,171,1706]" pageId="8" pageNumber="115">Pereopod IV (fig. 7F). Coxal plate with depth:width ratio 1.0:0.87. Gill ovoid, almost as large as the coxal plate. Dactylopodite (fig. 7G) with nail measuring one third of the total dactylopodite length, with one dorsal seta with plumose tip and one seta at nail insertion. Propodite:dactylopodite length ratio 1.0:0.35.</paragraph>
<paragraph id="8B85D8466A4A0C72FD77FCD1FD32FBE5" blockId="8.[676,1188,171,1706]" pageId="8" pageNumber="115">Pereopod V (fig. 8A). Coxal plate bilobed. Basipodite ovoid, length:width ratio 1.0:0.83. Short and thick setae present along the anterior and posterior margins of basipodite. Dactylopodite (fig. 8B) with one seta with plumose end on the outer margin and one seta al the insertion of the nail, which represents one quarter of the total dactylopodite length. Propodite:dactylopodite length ratio 1.0:0.38.</paragraph>
<paragraph id="8B85D8466A4A0C72FD77FBAAFC65FA5D" blockId="8.[676,1188,171,1706]" pageId="8" pageNumber="115">Pereopod VI (fig. 8C). Coxal plate of a relatively triangular shape, with a deep concavity in its dorsal margin. Basipodite (length:width ratio 1.0:0.82) with an ovoid shape, 7 groups of 2 setae on the anterior margin and 17 setae on the posterior margin. Dactylopodite (fig. 8D) with 1 plumose seta on the outer margin. The nail length represents one quarter of the total dactylopodite length. Ratio propodite:dactylopodite length 1.0:0.35.</paragraph>
<paragraph id="8B85D8466A4A0C72FD77FA03FB6DF927" blockId="8.[676,1188,171,1706]" pageId="8" pageNumber="115">Pereopod VII (fig. 8E). The coxal plate is trapezoidal. Basipodite with an ovoid to trapezoidal shape, ratio length:width 1.0:0.75. Basipodite with nine groups of 2 setae each on the anterior margin, and 21 small setae over the entire length of posterior margin.</paragraph>
<paragraph id="8B85D8466A4A0C73FD11F96CFEAEFEBF" blockId="8.[676,1219,1715,1746]" lastBlockId="9.[98,610,171,1706]" lastPageId="9" lastPageNumber="116" pageId="8" pageNumber="115">
<collectorName id="26CFBD906A4A0C72FD11F96CFC96F95C" box="[676,863,1715,1746]" pageId="8" pageNumber="115">Dactylopodite Downloaded</collectorName>
(fig. from 8F)
<collectorName id="26CFBD906A4A0C72FC2CF91CFC76F95C" box="[921,959,1731,1746]" pageId="8" pageNumber="115">Brill</collectorName>
with.com one07/10/ plumose 2024 01:59 seta on outer margin.
<collectorName id="26CFBD906A4B0C73FE86FF74FEA8FF4B" box="[307,353,171,197]" pageId="9" pageNumber="116">Nail</collectorName>
one fifth of the total dactylopodite length.
<collectorName id="26CFBD906A4B0C73FED4FF10FE54FF67" box="[353,413,207,233]" pageId="9" pageNumber="116">Ratio</collectorName>
propodite:dactylopodite length 1.0:0.37.
<collectorName id="26CFBD906A4B0C73FE3CFF2CFDD3FE80" box="[393,538,243,270]" pageId="9" pageNumber="116">Pereopods V</collectorName>
:VI:VII length ratio as 1.0:1.2:1.2
</paragraph>
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<paragraph id="8B85D8466A4A0C72FE47F907FD47F969" blockId="8.[129,654,1715,1768]" box="[498,654,1752,1768]" pageId="8" pageNumber="115">via Open Access.</paragraph>
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<collectorName id="26CFBD906A4A0C72FD23F907FD75F969" box="[662,700,1752,1767]" pageId="8" pageNumber="115">This</collectorName>
is an open access article distributed under the terms of the CC BY 4.0 license. https://creativecommons.org/licenses/by/4.0/
</paragraph>
<paragraph id="8B85D8466A4B0C73FF34FEE4FEFFFE13" blockId="9.[98,610,171,1706]" pageId="9" pageNumber="116">
<collectorName id="26CFBD906A4B0C73FF34FEE4FED2FED8" box="[129,283,315,342]" pageId="9" pageNumber="116">Pereopods VI</collectorName>
and VII of equal size, the longest legs of
<taxonomicName id="4C3AA3C56A4B0C73FEAEFEBFFE3EFEF7" authorityName="Malek-Hosseini &amp; Brad &amp; Fatemi &amp; Kuntner &amp; Fišer" authorityYear="2024" box="[283,503,351,377]" class="Malacostraca" family="Melitidae" genus="Tegano" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="116" phylum="Arthropoda" rank="species" species="tashanensis" status="sp. nov.">
<emphasis id="B94E04546A4B0C73FEAEFEBFFE3EFEF7" box="[283,503,351,377]" italics="true" pageId="9" pageNumber="116">Tegano tashanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DB92F6A4B0C73FDB6FEBFFD97FEF4" box="[515,606,352,378]" pageId="9" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
, measuring
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.
</paragraph>
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<collectorName id="26CFBD906A4B0C73FF34FE78FEACFE4F" box="[129,357,423,449]" pageId="9" pageNumber="116">
<emphasis id="B94E04546A4B0C73FF34FE78FF2BFE4F" box="[129,226,423,449]" italics="true" pageId="9" pageNumber="116">Pleopods</emphasis>
. Pleopods I
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, II and III highly similar (fig. 8G), with unequal rami and 2 retinacula (fig. 8H).
</paragraph>
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<emphasis id="B94E04546A4B0C73FF34FDCDFF17FDA2" box="[129,222,530,556]" italics="true" pageId="9" pageNumber="116">Uropods</emphasis>
. Urosomites I and II with 1 dorso-lateral seta on each side; dorsal margin of urosomites III with no seta.
<collectorName id="26CFBD906A4B0C73FEDBFD85FE54FDFA" box="[366,413,602,628]" pageId="9" pageNumber="116">One</collectorName>
spiniform seta on the posteroventral angle of the first urosomite near the insertion of uropod
<collectorName id="26CFBD906A4B0C73FE04FD7CFDD7FD32" box="[433,542,674,701]" pageId="9" pageNumber="116">I. Uropod</collectorName>
I (fig. 9A) little setose.
<collectorName id="26CFBD906A4B0C73FE9BFD19FE7BFD6E" box="[302,434,710,736]" pageId="9" pageNumber="116">Endopodite</collectorName>
slightly longer than the exopodite; ratio endopodite:exopodite 1.0:0.87.
<collectorName id="26CFBD906A4B0C73FF42FCD1FEA0FCA6" box="[247,361,782,808]" pageId="9" pageNumber="116">Uropod II</collectorName>
(fig. 9B) little setose, endopodite longer than the exopodite; ratio endopodite:exopodite 1.0:0.81.
<collectorName id="26CFBD906A4B0C73FE6CFC89FDE6FCFE" box="[473,559,854,880]" pageId="9" pageNumber="116">Uropod</collectorName>
III (fig. 9c) short, approximately 20% of the total body length.
<collectorName id="26CFBD906A4B0C73FEB5FC42FEBEFC39" box="[256,375,925,951]" pageId="9" pageNumber="116">Basipodite</collectorName>
with 2 apical stout setae on posterior margin and 2 smaller setae on anterior margin.
<collectorName id="26CFBD906A4B0C73FE8BFC3AFEB2FC71" box="[318,379,997,1023]" pageId="9" pageNumber="116">Short</collectorName>
endopodite, half the length of basipodite, with 1 short apical seta.
<collectorName id="26CFBD906A4B0C73FFD7FBF2FEE0FBC9" box="[98,297,1069,1095]" pageId="9" pageNumber="116">Single-segmented</collectorName>
exopodite, with 3 groups of 3 setae on the posterior margin, 5 groups of 23 setae on the anterior margin, six robust apical setae (fig. 9E).
</paragraph>
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<collectorName id="26CFBD906A4B0C73FF34FB62FF2DFB59" box="[129,228,1213,1239]" pageId="9" pageNumber="116">Epimeral</collectorName>
plates
</emphasis>
.
<collectorName id="26CFBD906A4B0C73FE82FB62FE54FB59" box="[311,413,1213,1239]" pageId="9" pageNumber="116">Epimeral</collectorName>
plates (
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) of different shape and size.
<collectorName id="26CFBD906A4B0C73FE32FB3FFE04FB74" box="[391,461,1248,1274]" pageId="9" pageNumber="116">Depth</collectorName>
ratio of
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plates I, II and III is 1.0:1.3:1.2. Epimeral plates with right postero-ventral angle and convex ventral margin. Epimeral plates II and III with 4 setae on the posterior margin. Pleonites II and III with 1 dorsolateral seta on each side.
</paragraph>
<paragraph id="8B85D8466A4B0C73FF34FA03FDF8F9EC" blockId="9.[98,610,171,1706]" pageId="9" pageNumber="116">
<emphasis id="B94E04546A4B0C73FF34FA03FF0FFA78" box="[129,198,1500,1526]" italics="true" pageId="9" pageNumber="116">Telson</emphasis>
(fig. 9F) as long as wide. 4 short apical spines on each lobe. The longest spine ¼ of total telson length. One mesial spine on each lobe, 12 lateral spines on each lobe.
</paragraph>
<paragraph id="8B85D8466A4B0C73FFD7F950FE89F924" blockId="9.[98,610,171,1706]" box="[98,320,1679,1706]" pageId="9" pageNumber="116">
<emphasis id="B94E04546A4B0C73FFD7F950FE89F924" bold="true" box="[98,320,1679,1706]" italics="true" pageId="9" pageNumber="116">Sexual dimorphism</emphasis>
</paragraph>
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Sexual dimorphism in
<taxonomicName id="4C3AA3C56A4B0C73FE34F96BFDABF943" authorityName="Malek-Hosseini &amp; Brad &amp; Fatemi &amp; Kuntner &amp; Fišer" authorityYear="2024" box="[385,610,1715,1741]" class="Malacostraca" family="Melitidae" genus="Tegano" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="116" phylum="Arthropoda" rank="species" species="tashanensis">
<emphasis id="B94E04546A4B0C73FE34F96BFDABF943" box="[385,610,1715,1741]" italics="true" pageId="9" pageNumber="116">Tegano tashanensis</emphasis>
</taxonomicName>
sp. nov. is negligible. While the male via Open seems stouter and with slightly longer antennae and pereopods, the diagnostic characters are invariable in the inspected male and female.
<collectorName id="26CFBD906A4B0C73FD30FEC7FD69FEBC" box="[645,672,280,306]" pageId="9" pageNumber="116">To</collectorName>
test for differences in quantitative traits, like body length (female of
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vs. male of.
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), we would need more individuals.
</paragraph>
</subSubSection>
<subSubSection id="C3208BCD6A4B0C73FD30FE79FBF2FDFA" pageId="9" pageNumber="116" type="etymology">
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<emphasis id="B94E04546A4B0C73FD30FE79FD37FE4F" bold="true" box="[645,766,422,449]" italics="true" pageId="9" pageNumber="116">
<collectorName id="26CFBD906A4B0C73FD30FE79FD37FE4F" box="[645,766,422,449]" pageId="9" pageNumber="116">Etymology</collectorName>
</emphasis>
</paragraph>
<paragraph id="8B85D8466A4B0C73FD30FE15FBF2FDFA" blockId="9.[645,1159,171,1706]" pageId="9" pageNumber="116">
<collectorName id="26CFBD906A4B0C73FD30FE15FD79FE6A" box="[645,688,458,484]" pageId="9" pageNumber="116">The</collectorName>
species is named after the name of
<collectorName id="26CFBD906A4B0C73FD30FE31FC46FD86" box="[645,911,494,520]" pageId="9" pageNumber="116">Tashan-Chah Kabootari</collectorName>
aquifer which
<collectorName id="26CFBD906A4B0C73FB80FE31FB4CFD86" box="[1077,1157,494,520]" pageId="9" pageNumber="116">Tashan</collectorName>
and
<collectorName id="26CFBD906A4B0C73FD0EFDCDFCBAFDA2" box="[699,883,530,556]" pageId="9" pageNumber="116">Chah Kabootari</collectorName>
caves are parts of this unique ecosystem.
<collectorName id="26CFBD906A4B0C73FCD7FDE9FC44FDDE" box="[866,909,566,592]" pageId="9" pageNumber="116">The</collectorName>
species epithet is an adjective meaning “of the Tashan cave”.
</paragraph>
</subSubSection>
<subSubSection id="C3208BCD6A4B0C70FD30FD7DFEDDFD6E" lastPageId="10" lastPageNumber="117" pageId="9" pageNumber="116" type="discussion">
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<emphasis id="B94E04546A4B0C73FD30FD7DFCAEFD32" bold="true" box="[645,871,673,700]" italics="true" pageId="9" pageNumber="116">
<collectorName id="26CFBD906A4B0C73FD30FD7DFCCDFD32" box="[645,772,674,700]" pageId="9" pageNumber="116">Taxonomic</collectorName>
remarks
</emphasis>
</paragraph>
<paragraph id="8B85D8466A4B0C73FD30FD19FCC4FB1E" blockId="9.[645,1159,171,1706]" pageId="9" pageNumber="116">
<collectorName id="26CFBD906A4B0C73FD30FD19FD79FD6E" box="[645,688,710,736]" pageId="9" pageNumber="116">The</collectorName>
genus
<taxonomicName id="4C3AA3C56A4B0C73FCB8FD18FC95FD6E" authorityName="Barnard &amp; Karaman" authorityYear="1982" box="[781,860,711,736]" class="Malacostraca" family="Melitidae" genus="Tegano" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B94E04546A4B0C73FCB8FD18FC95FD6E" box="[781,860,711,736]" italics="true" pageId="9" pageNumber="116">Tegano</emphasis>
</taxonomicName>
was erected by
<collectorName id="26CFBD906A4B0C73FB9FFD19FB4DFD6E" box="[1066,1156,710,736]" pageId="9" pageNumber="116">Barnard</collectorName>
and
<collectorName id="26CFBD906A4B0C73FD0CFD35FCE9FC8A" box="[697,800,746,772]" pageId="9" pageNumber="116">Karaman</collectorName>
(1982), on the basis of a single species,
<taxonomicName id="4C3AA3C56A4B0C73FD5EFCD1FC61FCA9" authorityName="Bousfield" authorityYear="1970" box="[747,936,782,808]" class="Malacostraca" family="Melitidae" genus="Melita" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="116" phylum="Arthropoda" rank="species" species="seticornis">
<emphasis id="B94E04546A4B0C73FD5EFCD1FC61FCA9" box="[747,936,782,808]" italics="true" pageId="9" pageNumber="116">Melita seticornis</emphasis>
</taxonomicName>
, primarily due to the reduced mandibular palp.
<collectorName id="26CFBD906A4B0C73FC55FCEDFB9DFCC2" box="[992,1108,818,844]" pageId="9" pageNumber="116">Nowadays</collectorName>
, the genus
<taxonomicName id="4C3AA3C56A4B0C73FD7AFC89FCD7FCE1" authorityName="Barnard &amp; Karaman" authorityYear="1982" box="[719,798,854,879]" class="Malacostraca" family="Melitidae" genus="Tegano" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B94E04546A4B0C73FD7AFC89FCD7FCE1" box="[719,798,854,879]" italics="true" pageId="9" pageNumber="116">Tegano</emphasis>
</taxonomicName>
comprises 12 described species distributed from
<collectingCountry id="F32D98D66A4B0C73FCFCFCA6FC79FC1D" box="[841,944,889,915]" name="Australia" pageId="9" pageNumber="116">Australia</collectingCountry>
,
<collectingCountry id="F32D98D66A4B0C73FC09FCA6FBE1FC1D" box="[956,1064,889,915]" name="Sri Lanka" pageId="9" pageNumber="116">Sri Lanka</collectingCountry>
(
<collectorName id="26CFBD906A4B0C73FB8CFCA6FB4CFC1D" box="[1081,1157,889,915]" pageId="9" pageNumber="116">Indian</collectorName>
<collectorName id="26CFBD906A4B0C73FD30FC42FD18FC39" box="[645,721,925,951]" pageId="9" pageNumber="116">Ocean</collectorName>
), and
<collectingCountry id="F32D98D66A4B0C73FC93FC42FCACFC39" box="[806,869,925,951]" name="Palau" pageId="9" pageNumber="116">Palau</collectingCountry>
, the Philippine Islands, and the
<collectingCountry id="F32D98D66A4B0C73FD5AFC1EFC79FC55" box="[751,944,961,987]" name="Solomon Islands" pageId="9" pageNumber="116">Solomon Islands</collectingCountry>
(all in the tropical
<location id="8EE58E9D6A4B0C73FD1AFC39FC78FC71" LSID="urn:lsid:plazi:treatment:039369506A440C70FF34FAAEFEDDFD6E:8EE58E9D6A4B0C73FD1AFC39FC78FC71" box="[687,945,997,1024]" country="Australia" name="western Pacific Ocean" pageId="9" pageNumber="116" stateProvince="Norfolk">western Pacific Ocean</location>
),
<collectingRegion id="49FE16A46A4B0C73FC73FC3AFBD2FC71" box="[966,1051,997,1023]" country="United Kingdom" name="Norfolk" pageId="9" pageNumber="116">Norfolk</collectingRegion>
<collectorName id="26CFBD906A4B0C73FB91FC3AFBAAFC71" box="[1060,1123,997,1023]" pageId="9" pageNumber="116">Coast</collectorName>
of
<collectingCountry id="F32D98D66A4B0C73FD30FBD6FC8EFBAD" box="[645,839,1033,1059]" name="United Kingdom" pageId="9" pageNumber="116">United Kingdom</collectingCountry>
and
<collectingCountry id="F32D98D66A4B0C73FC3EFBD6FBF3FBAD" box="[907,1082,1033,1059]" name="Curacao Island" pageId="9" pageNumber="116">Curaçao Island</collectingCountry>
(both
<collectorName id="26CFBD906A4B0C73FD30FBF2FCFCFBC9" box="[645,821,1069,1095]" pageId="9" pageNumber="116">Atlantic Ocean</collectorName>
).
<taxonomicName id="4C3AA3C56A4B0C73FCFFFBF1FC50FBC9" authorityName="Barnard &amp; Karaman" authorityYear="1982" box="[842,921,1070,1095]" class="Malacostraca" family="Melitidae" genus="Tegano" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B94E04546A4B0C73FCFFFBF1FC50FBC9" box="[842,921,1070,1095]" italics="true" pageId="9" pageNumber="116">Tegano</emphasis>
</taxonomicName>
includes some obligate cave-dwelling representatives (
<collectorName id="26CFBD906A4B0C73FB98FB8EFB4CFBE5" box="[1069,1157,1105,1131]" pageId="9" pageNumber="116">Sawicki</collectorName>
et al., 2005).
</paragraph>
<paragraph id="8B85D8466A4B0C70FD11FB46FDA7FF64" blockId="9.[645,1159,171,1706]" lastBlockId="10.[129,640,171,736]" lastPageId="10" lastPageNumber="117" pageId="9" pageNumber="116">
<collectorName id="26CFBD906A4B0C73FD11FB46FD06FB3D" box="[676,719,1177,1203]" pageId="9" pageNumber="116">The</collectorName>
newly described species belongs to a group of
<taxonomicName id="4C3AA3C56A4B0C73FCA5FB62FC75FB58" box="[784,956,1212,1238]" pageId="9" pageNumber="116">
<emphasis id="B94E04546A4B0C73FCA5FB62FC96FB58" box="[784,863,1213,1238]" italics="true" pageId="9" pageNumber="116">Tegano</emphasis>
species
</taxonomicName>
with completely reduced mandibular palps. There are three species in this group,
<taxonomicName id="4C3AA3C56A4B0C73FC2AFADAFB4CFA90" baseAuthorityName="G. Karaman" baseAuthorityYear="1984" box="[927,1157,1284,1310]" class="Malacostraca" family="Melitidae" genus="Tegano" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="116" phylum="Arthropoda" rank="species" species="vagabundus">
<emphasis id="B94E04546A4B0C73FC2AFADAFB4CFA90" box="[927,1157,1284,1310]" italics="true" pageId="9" pageNumber="116">Tegano vagabundus</emphasis>
</taxonomicName>
from
<collectingCountry id="F32D98D66A4B0C73FD7CFAF7FCF5FACC" box="[713,828,1320,1346]" name="Sri Lanka" pageId="9" pageNumber="116">Sri Lanka</collectingCountry>
, where mandibular palp is replaced by a long seta (Karaman, 1984), and
<taxonomicName id="4C3AA3C56A4B0C73FD30FAAEFC95FA04" authorityName="Malek-Hosseini &amp; Brad &amp; Fatemi &amp; Kuntner &amp; Fišer" authorityYear="2024" box="[645,860,1392,1418]" class="Malacostraca" family="Melitidae" genus="Tegano" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="116" phylum="Arthropoda" rank="species" species="tashanensis" status="sp. nov.">
<emphasis id="B94E04546A4B0C73FD30FAAEFC95FA04" box="[645,860,1392,1418]" italics="true" pageId="9" pageNumber="116">Tegano tashanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DB92F6A4B0C73FCD4FAAEFC7AFA05" box="[865,947,1393,1419]" pageId="9" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
in
<collectorName id="26CFBD906A4B0C73FC66FAAFFB4DFA04" box="[979,1156,1392,1418]" pageId="9" pageNumber="116">Chah Kabootari</collectorName>
Cave (
<collectingCountry id="F32D98D66A4B0C73FD67FA4BFCCAFA20" box="[722,771,1428,1454]" name="Iran" pageId="9" pageNumber="116">Iran</collectingCountry>
) and
<taxonomicName id="4C3AA3C56A4B0C73FCE6FA4AFB89FA20" authorityName="Sawicki &amp; Holsinger" authorityYear="2005" box="[851,1088,1428,1454]" class="Malacostraca" family="Melitidae" genus="Tegano" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="116" phylum="Arthropoda" rank="species" species="panglaoensis">
<emphasis id="B94E04546A4B0C73FCE6FA4AFB89FA20" box="[851,1088,1428,1454]" italics="true" pageId="9" pageNumber="116">Tegano panglaoensis</emphasis>
</taxonomicName>
from
<collectingCountry id="F32D98D66A4B0C73FD30FA67FCCFFA5C" box="[645,774,1464,1490]" name="Philippines" pageId="9" pageNumber="116">Philippines</collectingCountry>
, where the mandibular palp is completely absent. The latter species differs from herein described
<taxonomicName id="4C3AA3C56A4B0C73FC3EF9DFFBE1F994" authorityName="Malek-Hosseini &amp; Brad &amp; Fatemi &amp; Kuntner &amp; Fišer" authorityYear="2024" box="[907,1064,1536,1562]" class="Malacostraca" family="Melitidae" genus="Tegano" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="116" phylum="Arthropoda" rank="species" species="tashanensis" status="sp. nov.">
<emphasis id="B94E04546A4B0C73FC3EF9DFFBE1F994" box="[907,1064,1536,1562]" italics="true" pageId="9" pageNumber="116">T. tashanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DB92F6A4B0C73FB85F9DFFB4CF994" box="[1072,1157,1536,1562]" pageId="9" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
by presence of eyes and labial lobes, narrow bases of pereopods V-VII (much broader in
<taxonomicName id="4C3AA3C56A4B0C73FBC4F997FB4EF9EF" authorityName="Malek-Hosseini &amp; Brad &amp; Fatemi &amp; Kuntner &amp; Fišer" authorityYear="2024" box="[1137,1159,1608,1633]" pageId="9" pageNumber="116" status="sp. nov.">
<emphasis id="B94E04546A4B0C73FBC4F997FB4EF9EF" box="[1137,1159,1608,1633]" italics="true" pageId="9" pageNumber="116">T.</emphasis>
</taxonomicName>
<emphasis id="B94E04546A4B0C73FD30F9B4FCCEF90B" box="[645,775,1643,1669]" italics="true" pageId="9" pageNumber="116">tashanensis</emphasis>
<taxonomicNameLabel id="A27DB92F6A4B0C73FCA5F9B3FCA1F908" box="[784,872,1644,1670]" pageId="9" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
), single basiofacial spine on uropod I (two in
<taxonomicName id="4C3AA3C56A4B0C73FCCCF94FFBDFF927" authorityName="Malek-Hosseini &amp; Brad &amp; Fatemi &amp; Kuntner &amp; Fišer" authorityYear="2024" box="[889,1046,1679,1705]" class="Malacostraca" family="Melitidae" genus="Tegano" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="116" phylum="Arthropoda" rank="species" species="tashanensis" status="sp. nov.">
<emphasis id="B94E04546A4B0C73FCCCF94FFBDFF927" box="[889,1046,1679,1705]" italics="true" pageId="9" pageNumber="116">T. tashanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DB92F6A4B0C73FB95F94FFBB0F927" box="[1056,1145,1679,1706]" pageId="9" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
), longer distal Downloaded article from on Brill.com exopodite
<date id="FF84FE866A4B0C73FC4CF91CFB90F95C" pageId="9" pageNumber="116" value="2024-10-07">07/10/of 2024</date>
01 uro-:59:59PM. pod This III is (an short open in access
<taxonomicName id="4C3AA3C56A4B0C73FCFFF907FC2FF97F" authorityName="Malek-Hosseini &amp; Brad &amp; Fatemi &amp; Kuntner &amp; Fišer" authorityYear="2024" class="Malacostraca" family="Melitidae" genus="Tegano" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="116" phylum="Arthropoda" rank="species" species="tashanensis">
<emphasis id="B94E04546A4B0C73FCFFF907FC2FF97F" italics="true" pageId="9" pageNumber="116">T. article tashanensis</emphasis>
</taxonomicName>
distributed sp. nov.) under, and the terms more pointed telson lobes (subrounded in
<taxonomicName id="4C3AA3C56A480C70FF34FF0FFED2FF67" authorityName="Malek-Hosseini &amp; Brad &amp; Fatemi &amp; Kuntner &amp; Fišer" authorityYear="2024" box="[129,283,207,233]" class="Malacostraca" family="Melitidae" genus="Tegano" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="117" phylum="Arthropoda" rank="species" species="tashanensis" status="sp. nov.">
<emphasis id="B94E04546A480C70FF34FF0FFED2FF67" box="[129,283,207,233]" italics="true" pageId="10" pageNumber="117">T. tashanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DB92F6A480C70FE95FF0FFEBDFF67" box="[288,372,207,234]" pageId="10" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
) (
<bibRefCitation id="EFABA5B76A480C70FE3CFF10FDABFF64" author="Sawicki, T. R. &amp; Holsinger, J. R. &amp; Iliffe, T. M." box="[393,610,207,234]" pageId="10" pageNumber="117" pagination="49 - 74" refId="ref11728" refString="Sawicki, T. R., Holsinger, J. R. &amp; Iliffe, T. M. (2005) New species of amphipod crustaceans in the genera Tegano and Melita (Hadzioidea: Melitidae) from subterranean groundwaters in Guam, Palau, and the Philippines. J. Crustac. Biol. 25: 49 - 74." type="journal article" year="2005">Sawicki et al., 2005</bibRefCitation>
).
</paragraph>
<paragraph id="8B85D8466A4B0C73FDFCF907FD43F969" blockId="9.[98,650,1715,1778]" box="[585,650,1752,1767]" pageId="9" pageNumber="116">Access</paragraph>
<paragraph id="8B85D8466A4B0C73FC67F932FB0DF972" blockId="9.[796,1220,1773,1810]" box="[978,1220,1773,1789]" pageId="9" pageNumber="116">of the CC BY 4.0 license.</paragraph>
<paragraph id="8B85D8466A4B0C73FCA9F8DCFB0AF89C" blockId="9.[796,1220,1773,1810]" box="[796,1219,1795,1810]" pageId="9" pageNumber="116">https://creativecommons.org/licenses/by/4.0/</paragraph>
<paragraph id="8B85D8466A480C70FF2AFF2CFEDDFD6E" blockId="10.[129,640,171,736]" pageId="10" pageNumber="117">
There is an unclear relationship to hadzioid genus
<taxonomicName id="4C3AA3C56A480C70FF7FFEC8FF32FEBF" authorityName="Stock" authorityYear="1988" box="[202,251,279,305]" class="Malacostraca" family="Melitidae" genus="Fiha" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B94E04546A480C70FF7FFEC8FF32FEBF" box="[202,251,279,305]" italics="true" pageId="10" pageNumber="117">Fiha</emphasis>
</taxonomicName>
, which we could not analyze phylogenetically, with reduced mandibular palp and inner labial lobes. However,
<taxonomicName id="4C3AA3C56A480C70FE4BFE80FDE6FEF7" authorityName="Stock" authorityYear="1988" box="[510,559,351,377]" class="Malacostraca" family="Melitidae" genus="Fiha" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B94E04546A480C70FE4BFE80FDE6FEF7" box="[510,559,351,377]" italics="true" pageId="10" pageNumber="117">Fiha</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C3AA3C56A480C70FF0BFE5BFE5EFE13" authorityName="Malek-Hosseini &amp; Brad &amp; Fatemi &amp; Kuntner &amp; Fišer" authorityYear="2024" box="[190,407,387,413]" class="Malacostraca" family="Melitidae" genus="Tegano" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="117" phylum="Arthropoda" rank="species" species="tashanensis" status="sp. nov.">
<emphasis id="B94E04546A480C70FF0BFE5BFE5EFE13" box="[190,407,387,413]" italics="true" pageId="10" pageNumber="117">Tegano tashanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DB92F6A480C70FE2BFE5CFE3BFE13" box="[414,498,387,413]" pageId="10" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
in slenderer body (more stout in
<taxonomicName id="4C3AA3C56A480C70FEC6FE77FDD8FE4F" authorityName="Malek-Hosseini &amp; Brad &amp; Fatemi &amp; Kuntner &amp; Fišer" authorityYear="2024" box="[371,529,423,449]" class="Malacostraca" family="Melitidae" genus="Tegano" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="117" phylum="Arthropoda" rank="species" species="tashanensis" status="sp. nov.">
<emphasis id="B94E04546A480C70FEC6FE77FDD8FE4F" box="[371,529,423,449]" italics="true" pageId="10" pageNumber="117">T. tashanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DB92F6A480C70FDAEFE78FDBDFE4F" box="[539,628,423,449]" pageId="10" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
), accessory flagellum of antenna I (34 articles in
<emphasis id="B94E04546A480C70FF15FE31FF1EFD86" box="[160,215,494,520]" italics="true" pageId="10" pageNumber="117">
<taxonomicName id="4C3AA3C56A480C70FF15FE31FF1AFD86" authorityName="Stock" authorityYear="1988" box="[160,211,494,520]" class="Malacostraca" family="Melitidae" genus="Fiha" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="117" phylum="Arthropoda" rank="genus">Fiha</taxonomicName>
,
</emphasis>
<quantity id="4CC275A36A480C70FF62FE30FED2FD86" box="[215,283,494,521]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="5.08" metricValueMin="2.54" pageId="10" pageNumber="117" unit="in" value="1.5" valueMax="2.0" valueMin="1.0">12 in</quantity>
<taxonomicName id="4C3AA3C56A480C70FE91FE30FE0BFD86" authorityName="Malek-Hosseini &amp; Brad &amp; Fatemi &amp; Kuntner &amp; Fišer" authorityYear="2024" box="[292,450,494,520]" class="Malacostraca" family="Melitidae" genus="Tegano" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="117" phylum="Arthropoda" rank="species" species="tashanensis" status="sp. nov.">
<emphasis id="B94E04546A480C70FE91FE30FE0BFD86" box="[292,450,494,520]" italics="true" pageId="10" pageNumber="117">T. tashanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DB92F6A480C70FE7FFE30FDEAFD87" box="[458,547,495,521]" pageId="10" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
), longer propodus in gnathopod I (relatively shorter in
<taxonomicName id="4C3AA3C56A480C70FF2BFDE8FEF0FDDE" authorityName="Malek-Hosseini &amp; Brad &amp; Fatemi &amp; Kuntner &amp; Fišer" authorityYear="2024" box="[158,313,566,592]" class="Malacostraca" family="Melitidae" genus="Tegano" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="117" phylum="Arthropoda" rank="species" species="tashanensis" status="sp. nov.">
<emphasis id="B94E04546A480C70FF2BFDE8FEF0FDDE" box="[158,313,566,592]" italics="true" pageId="10" pageNumber="117">T. tashanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DB92F6A480C70FEF5FDE8FE5FFDDF" box="[320,406,567,593]" pageId="10" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
), coxal plate IV shallow (deeply incised in
<taxonomicName id="4C3AA3C56A480C70FEC8FD84FDD1FDFA" authorityName="Malek-Hosseini &amp; Brad &amp; Fatemi &amp; Kuntner &amp; Fišer" authorityYear="2024" box="[381,536,602,628]" class="Malacostraca" family="Melitidae" genus="Tegano" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="117" phylum="Arthropoda" rank="species" species="tashanensis" status="sp. nov.">
<emphasis id="B94E04546A480C70FEC8FD84FDD1FDFA" box="[381,536,602,628]" italics="true" pageId="10" pageNumber="117">T. tashanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DB92F6A480C70FDABFD84FDBDFDFA" box="[542,628,602,629]" pageId="10" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
), and shallower and more open telson (longer lobes close together in
<taxonomicName id="4C3AA3C56A480C70FE36FD7CFDD7FD32" authorityName="Malek-Hosseini &amp; Brad &amp; Fatemi &amp; Kuntner &amp; Fišer" authorityYear="2024" box="[387,542,674,700]" class="Malacostraca" family="Melitidae" genus="Tegano" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="117" phylum="Arthropoda" rank="species" species="tashanensis" status="sp. nov.">
<emphasis id="B94E04546A480C70FE36FD7CFDD7FD32" box="[387,542,674,700]" italics="true" pageId="10" pageNumber="117">T. tashanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DB92F6A480C70FD91FD7CFDB1FD32" box="[548,632,674,701]" pageId="10" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
) (
<bibRefCitation id="EFABA5B76A480C70FF3DFD19FEC0FD6E" author="Stock, J. H." box="[136,265,710,736]" pageId="10" pageNumber="117" pagination="79 - 100" refId="ref11854" refString="Stock, J. H. (1988) Two new Stygobiont Amphipoda (Crustacea) from Polynesia. Stygologia 4: 79 - 100." type="journal article" year="1988">Stock, 1988</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,40 +1,40 @@
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<mods:mods id="C6B0A9DBA43C43E8365D4D5D0061B2A4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E2D0DB3B322EA604AAFA06EE48BDFE8F">
<mods:title id="8FE6B5B14AC1774416B82C514BBA5815">Formicidae.</mods:title>
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<taxonomicName id="2FFDC2FD096B468F50782EE2AF5B8C16" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:179413" class="Magnoliopsida" family="Asteraceae" genus="Acantholepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acantholepis capensis hirsuta Santschi" lsidName-HNS="Acantholepis capensis hirsuta Santschi" order="Asterales" pageNumber="124" phylum="Tracheophyta" rank="subSpecies" species="capensis" subSpecies="hirsuta">A. capensis, st. hirsuta</taxonomicName>
<taxonomicName id="2FFDC2FD096B468F50782EE2AF5B8C16" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:179413" class="Insecta" family="Formicidae" genus="Acantholepis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Acantholepis capensis hirsuta Santschi" lsidName-HNS="Acantholepis capensis hirsuta Santschi" order="Hymenoptera" pageNumber="124" phylum="Arthropoda" rank="subSpecies" species="capensis" subSpecies="hirsuta">A. capensis, st. hirsuta</taxonomicName>
, nov.
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@ -42,16 +42,16 @@
<paragraph id="C853F6F1C59A93179828A7556773895F" pageNumber="124">Fig. 26.</paragraph>
<paragraph id="8B56F343A8913CD5FF48F4FB04E0AC07" pageNumber="124">
[[worker]]. - Long. 2,7 a 3 mill. Plus grande et plus fortement sculptee sur les cotes et le dessus de l'epinotum que chez
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<taxonomicName id="5B5AA12D56104902778304F1FB041322" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:179050" class="Insecta" family="Formicidae" genus="Acantholepis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Acantholepis capensis canescens Emery" lsidName-HNS="Acantholepis capensis canescens Emery" order="Hymenoptera" pageNumber="124" phylum="Arthropoda" rank="species" species="canescens">canescens</taxonomicName>
. La pilosite dressee blanche est encore plus abondante et plus
</paragraph>
<paragraph id="70A09E6A676CBE20E2AF30F5FF762BCE" pageNumber="124">
touffue que chez cette derniere; de longueur irreguliere, elle forme sur les pattes et les joues une pilosite plus fine. Pubescence plus courte que les soies, mais bien plus longue que la pubescence adjacente de
<taxonomicName id="72A5E912E8CFE5686AA83F77A786506F" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:179050" class="Magnoliopsida" family="Asteraceae" genus="Acantholepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acantholepis capensis canescens Emery" lsidName-HNS="Acantholepis capensis canescens Emery" order="Asterales" pageNumber="124" phylum="Tracheophyta" rank="species" species="canescens">canescens</taxonomicName>
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; elle est aussi plus relevee. Sur les scapes, la pubescence qui est tres adjacente sur les autres races de
<taxonomicName id="ECD3CDE6F263125DE9D633C427D875BF" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139215" class="Magnoliopsida" family="Asteraceae" genus="Acantholepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acantholepis capensis Mayr" lsidName-HNS="Acantholepis capensis Mayr" order="Asterales" pageNumber="124" phylum="Tracheophyta" rank="species" species="capensis">capensis</taxonomicName>
<taxonomicName id="ECD3CDE6F263125DE9D633C427D875BF" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139215" class="Insecta" family="Formicidae" genus="Acantholepis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Acantholepis capensis Mayr" lsidName-HNS="Acantholepis capensis Mayr" order="Hymenoptera" pageNumber="124" phylum="Arthropoda" rank="species" species="capensis">capensis</taxonomicName>
est ici tres relevee et plus longue. Promesonotum, comme chez
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<taxonomicName id="9A103B8C319DAA86B0FCAD9472946A82" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:179050" class="Insecta" family="Formicidae" genus="Acantholepis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Acantholepis capensis canescens Emery" lsidName-HNS="Acantholepis capensis canescens Emery" order="Hymenoptera" pageNumber="124" phylum="Arthropoda" rank="species" species="canescens">A. canescens</taxonomicName>
, plus elance que chez le type, le pronotum plus convexe transversalement en avant. Ecaille faiblement dentee, assez mince au sommet. Noire, moitie basale du funicule et tarses roussatres.
</paragraph>
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@ -107,7 +107,7 @@ and allied genera (
, and probably several other genera of which only males and females are known). Temporary social parasitism is probably the rule in some species of
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and in the Malagasy and Indomalayan subgenus
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of
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fig. 5.
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@ -49,7 +49,7 @@ fig. 5.
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Per la descrizione dell'altra specie (
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), giuntami dopo che queste pagine erano gia composte, veggasi in fine del lavoro.
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@ -1,38 +1,38 @@
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<taxonomicName id="0241524D2B74734D13A34B30B60A43F9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:179050" class="Magnoliopsida" family="Asteraceae" genus="Acantholepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acantholepis capensis canescens Emery" lsidName-HNS="Acantholepis capensis canescens Emery" order="Asterales" pageNumber="124" phylum="Tracheophyta" rank="subSpecies" species="capensis" subSpecies="canescens">A. capensis Mayr, st. canescens Emery</taxonomicName>
<taxonomicName id="0241524D2B74734D13A34B30B60A43F9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:179050" class="Insecta" family="Formicidae" genus="Acantholepis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Acantholepis capensis canescens Emery" lsidName-HNS="Acantholepis capensis canescens Emery" order="Hymenoptera" pageNumber="124" phylum="Arthropoda" rank="subSpecies" species="capensis" subSpecies="canescens">A. capensis Mayr, st. canescens Emery</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="4515A5F052388C0E1430A4B3B4A20B4E" type="reference_group">
<paragraph id="26C2CA65A87569C01DF3FD704933FA64" pageNumber="124">
Mayr, Verb. zool.-bot. Ges. Wien, vol. 12, p. 699 (1862) (
<taxonomicName id="6C0975CEDFAAF1605162E4D61C119449" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139215" class="Magnoliopsida" family="Asteraceae" genus="Acantholepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acantholepis capensis Mayr" lsidName-HNS="Acantholepis capensis Mayr" order="Asterales" pageNumber="124" phylum="Tracheophyta" rank="species" species="capensis">A. capensis</taxonomicName>
<taxonomicName id="6C0975CEDFAAF1605162E4D61C119449" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139215" class="Insecta" family="Formicidae" genus="Acantholepis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Acantholepis capensis Mayr" lsidName-HNS="Acantholepis capensis Mayr" order="Hymenoptera" pageNumber="124" phylum="Arthropoda" rank="species" species="capensis">A. capensis</taxonomicName>
). - Emery, Ann. Mus. civ. Genova, vol. 38, p. 601 (1897), [[worker]] (
<taxonomicName id="4E9DE876B34FE61405B1B62D2FB412F5" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:179050" class="Magnoliopsida" family="Asteraceae" genus="Acantholepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acantholepis capensis canescens Emery" lsidName-HNS="Acantholepis capensis canescens Emery" order="Asterales" pageNumber="124" phylum="Tracheophyta" rank="subSpecies" species="capensis" subSpecies="canescens">st. canescens</taxonomicName>
<taxonomicName id="4E9DE876B34FE61405B1B62D2FB412F5" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:179050" class="Insecta" family="Formicidae" genus="Acantholepis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Acantholepis capensis canescens Emery" lsidName-HNS="Acantholepis capensis canescens Emery" order="Hymenoptera" pageNumber="124" phylum="Arthropoda" rank="subSpecies" species="capensis" subSpecies="canescens">st. canescens</taxonomicName>
).
</paragraph>
</subSubSection>

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<mods:title id="8B2AC0AFBF894418B03252A4FF5DBF13">New data on embryonic development of Halisarca dujardini Johnston, 1842 (Demospongiae, Halisarcida)</mods:title>
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<treatment id="173387ACFFB3957B2157EB2DAE5AE481" LSID="urn:lsid:plazi:treatment:173387ACFFB3957B2157EB2DAE5AE481" httpUri="http://treatment.plazi.org/id/173387ACFFB3957B2157EB2DAE5AE481" lastPageId="7" lastPageNumber="362" pageId="2" pageNumber="357">
<subSubSection id="D7806531FFB3957E2157EB2DABC0E444" box="[114,509,1586,1612]" pageId="2" pageNumber="357" type="nomenclature">
<paragraph id="9F2536BAFFB3957E2157EB2DABC0E444" blockId="2.[114,647,1586,1704]" box="[114,509,1586,1612]" pageId="2" pageNumber="357">
<taxonomicName id="589A4D39FFB3957E2157EB2DABC0E444" authority="Johnston, 1842" authorityName="Johnston. Vestnik" authorityYear="1842" box="[114,509,1586,1612]" class="Demospongiae" family="Halisarcidae" genus="Halisarca" kingdom="Animalia" order="Chondrillida" pageId="2" pageNumber="357" phylum="Porifera" rank="species" species="dujardini">
<emphasis id="ADEEEAA8FFB3957E2157EB2DAB79E444" box="[114,324,1586,1612]" italics="true" pageId="2" pageNumber="357">Halisarca dujardini</emphasis>
Johnston, 1842
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="D7806531FFB3957B232FEB2DAE5AE481" lastPageId="7" lastPageNumber="362" pageId="2" pageNumber="357" type="description">
<paragraph id="9F2536BAFFB3957E232FEB2DAE6EE425" blockId="2.[114,647,1586,1704]" lastBlockId="2.[677,1211,849,1704]" pageId="2" pageNumber="357">
(
<taxonomicName id="589A4D39FFB3957E2334EB2DAAF7E463" authorityName="Lévi" authorityYear="1953" class="Demospongiae" kingdom="Animalia" pageId="2" pageNumber="367" phylum="Porifera" rank="subClass" subClass="Ceractinomorpha">Ceractinomorpha</taxonomicName>
,
<taxonomicName id="589A4D39FFB3957E21F6EB4EAB73E463" authorityName="Topsent" authorityYear="1901" box="[211,334,1617,1643]" class="Demospongiae" higherTaxonomySource="CoL" kingdom="Animalia" order="Halisarcida" pageId="2" pageNumber="357" phylum="Porifera" rank="order">Halisarcida</taxonomicName>
) is a dioecious sponge. Males begin to produce spermatocysts with spermatocytes in approximately mid-December at water temperature of about 0.1 °C. Early oocytes differentiate in the last decade of December when water temperature fluctuates around 0.6 °C. Vitellogenesis starts in May at nearly + 2 °C. Eggs maturate by the end of May. Cleavage and larval development occur from the second half of June until the end of July when water temperature averages 10-12 °C. The volume of reproductive elements (embryos and prelarvae) reaches its maximum by the end of June beginning of July and reaches to about 69.5% of sponge volume (Ereskovsky 2000). It is the time when mesohyl destruction and complete disorder of central and basal choanosome, which are largely filled with developing larvae, is taking place. Normal tissue organization remains only in the thin periphery of a sponge. In the studied population, larval emergence is swift, lasting for one and a half to two weeks in the second half of July at about 12 °C. Subsequently slow development of the new generation of sponges and postreproductive revival of maternal specimens is going on until December. The general scheme of the
<taxonomicName id="589A4D39FFB3957E22E8E8EBAE69E407" authorityName="Johnston. Vestnik" authorityYear="1842" box="[973,1108,1524,1551]" class="Demospongiae" family="Halisarcidae" genus="Halisarca" kingdom="Animalia" order="Chondrillida" pageId="2" pageNumber="357" phylum="Porifera" rank="species" species="dujardini">
<emphasis id="ADEEEAA8FFB3957E22E8E8EBAE69E407" box="[973,1108,1524,1551]" italics="true" pageId="2" pageNumber="357">H. dujardini</emphasis>
</taxonomicName>
life cycle in the White Sea is illustrated on
<figureCitation id="07A12A3FFFB3957E252BEB0CAE71E425" box="[1038,1100,1555,1581]" captionStart="FIG" captionStartId="2.[677,688,759,776]" captionTargetBox="[680,1209,206,735]" captionTargetId="figure-530@2.[668,1218,194,746]" captionTargetPageId="2" captionText="FIG. 1. — Life-history scheme of Halisarca dujardini in the White Sea." figureDoi="http://doi.org/10.5281/zenodo.5402178" httpUri="https://zenodo.org/record/5402178/files/figure.png" pageId="2" pageNumber="357">Fig. 1</figureCitation>
.
</paragraph>
<paragraph id="9F2536BAFFB3957E2380EB2DA994E4A0" blockId="2.[677,1211,849,1704]" pageId="2" pageNumber="357">
Embryogenesis, larval development and metamorphosis of the studied population of
<taxonomicName id="589A4D39FFB3957E2570EB4FA8EDE481" authorityName="Johnston. Vestnik" authorityYear="1842" class="Demospongiae" family="Halisarcidae" genus="Halisarca" kingdom="Animalia" order="Chondrillida" pageId="2" pageNumber="357" phylum="Porifera" rank="species" species="dujardini">
<emphasis id="ADEEEAA8FFB3957E2570EB4FAE57E461" box="[1109,1130,1616,1641]" italics="true" pageId="2" pageNumber="357">H</emphasis>
.
<emphasis id="ADEEEAA8FFB3957E255DEB4EA8EDE481" italics="true" pageId="2" pageNumber="357">dujardini</emphasis>
</taxonomicName>
is continuing generally for four weeks from late June till end of July.
</paragraph>
<paragraph id="9F2536BAFFB2957F2154EDD1A842E07F" blockId="3.[113,647,205,1704]" pageId="3" pageNumber="358">
The oocyte development of this viviparous species occurs inside temporary embryonic capsules formed before maturation divisions of oocytes at the expense of dedifferentiated choanocytes (
<bibRefCitation id="FB0B4B4BFFB2957F21FDEC56A812E36B" author="Korotkova G. P. &amp; Apalkova L. V." box="[216,559,329,355]" editor="Tokin I. B." pageId="3" pageNumber="358" pagination="9 - 26" refId="ref8156" refString="Korotkova G. P. &amp; Apalkova L. V. 1975. - Oogenesis of the Barents Sea sponge Halisarca dujardini: 9 - 26, in Tokin I. B. (ed.), Comparative and Experimental Morphology of the Sea Organisms. Kolske filial Academy of Sciences Apatiti (in Russian)." title="Oogenesis of the Barents Sea sponge Halisarca dujardini" type="book chapter" volumeTitle="Comparative and Experimental Morphology of the Sea Organisms. Kolske filial Academy of Sciences Apatiti (in Russian)." year="1975">Korotkova &amp; Apalkova 1975</bibRefCitation>
). Every capsule has a double layer structure: the cells of the embryonic capsule stretching along the embryo surface and the layer of collagen filaments extending parallel to the cell surface on the outside. The thickness of fiber layer is 1 ± 0.5 µm. Distal parts of embryonic capsule cells overlay each other forming a continuous cell layer. No special contacts between individual cells have been recorded (
<bibRefCitation id="FB0B4B4BFFB2957F206BEF42A84FE07F" author="Sizova N. A. &amp; Ereskovsky A. V." box="[334,626,605,631]" editor="Ereskovsky A. &amp; Keupp H. &amp; Kohring R." pageId="3" pageNumber="358" pagination="103 - 113" refId="ref8710" refString="Sizova N. A. &amp; Ereskovsky A. V. 1997. - Ultrastructural peculiarities of the early embryogenesis in a White Sea Sponge Halisarca dujardini (Demosponge, Dendroceratida): 103 - 113, in Ereskovsky A., Keupp H. &amp; Kohring R. (eds), Modern Problems of Poriferan Biology. Freie Universitat, Berlin." title="Ultrastructural peculiarities of the early embryogenesis in a White Sea Sponge Halisarca dujardini (Demosponge, Dendroceratida)" type="book chapter" volumeTitle="Modern Problems of Poriferan Biology. Freie Universitat, Berlin" year="1997">Sizova &amp; Ereskovsky 1997</bibRefCitation>
).
</paragraph>
<paragraph id="9F2536BAFFB2957F2154EF64AAE7E145" blockId="3.[113,647,205,1704]" pageId="3" pageNumber="358">
Before cleavage cell division, a mature egg is a slightly oval body of 129 × 105 µm size. A spherical nucleus about 28 µm in diameter with a distinctly marked nucleolus (8 to 12 µm) is located in the centre of an egg. Vitelline granules are evenly spread over the whole volume of egg (
<figureCitation id="07A12A3FFFB2957F215CEE2CAAF0E145" box="[121,205,819,845]" captionStart="FIG" captionStartId="8.[114,125,789,806]" captionTargetBox="[124,1201,204,764]" captionTargetId="figure-453@8.[114,1229,196,767]" captionTargetPageId="8" captionText="FIG. 5. — Scheme of H. dujardini embryonic development; A, mature egg; B, early cleavage; C, beginning of blastomere polarization; D, multipolar immigration; E, stage larval polarization; F, formation of internal sphere and migration of maternal eosinophilic granular amoebocytes into larva; G, disphaerula, larva of H. dujardini. Abbreviations: bi, blastocoel; ec, embryonic capsule; efa, eosinophilic amoebocytes; ic, internal cells; isp, internal sphere; mc, migrated cells; ppc, posterior pole cells; tc, transition cells; yg, yolk granules." figureDoi="http://doi.org/10.5281/zenodo.5402186" httpUri="https://zenodo.org/record/5402186/files/figure.png" pageId="3" pageNumber="358">Fig. 5A</figureCitation>
).
</paragraph>
<paragraph id="9F2536BAFFB2957F2154EE4EAB07E730" blockId="3.[113,647,205,1704]" pageId="3" pageNumber="358">
Eggs cleavage in
<taxonomicName id="589A4D39FFB2957F2008EE4EAB8BE164" authorityName="Johnston. Vestnik" authorityYear="1842" box="[301,438,849,876]" class="Demospongiae" family="Halisarcidae" genus="Halisarca" kingdom="Animalia" order="Chondrillida" pageId="3" pageNumber="358" phylum="Porifera" rank="species" species="dujardini">
<emphasis id="ADEEEAA8FFB2957F2008EE4EAB8BE164" box="[301,438,849,876]" italics="true" pageId="3" pageNumber="358">H. dujardini</emphasis>
</taxonomicName>
is complete, equal and asynchronous. If the egg pole where maturation divisions are taking place is conventionally considered to be the animal one, the first cleavage division in
<taxonomicName id="589A4D39FFB2957F21C8EED3AB49E1EF" authorityName="Johnston. Vestnik" authorityYear="1842" box="[237,372,972,999]" class="Demospongiae" family="Halisarcidae" genus="Halisarca" kingdom="Animalia" order="Chondrillida" pageId="3" pageNumber="358" phylum="Porifera" rank="species" species="dujardini">
<emphasis id="ADEEEAA8FFB2957F21C8EED3AB49E1EF" box="[237,372,972,999]" italics="true" pageId="3" pageNumber="358">H. dujardini</emphasis>
</taxonomicName>
occurs in the meridional plane. The size of the first two blastomeres is about 75.3 × 105.6 µm. The nucleus diameter is 16.4 µm. During all cleavage stages, the interphase blastomeres nuclei contain one to three pronucleolar bodies of 3 to 8 µm diameter at their periphery. As a rule these structures are of rounded shape. During the second cleavage division, the relative position of spindles varies from nearly parallel to perpendicular to one to another, while the relative position of blastomeres resembles a tetrahedron.
</paragraph>
<paragraph id="9F2536BAFFB2957F2154E822ABE7E407" blockId="3.[113,647,205,1704]" pageId="3" pageNumber="358">
Similarly during the third cleavage division the spindle position within the embryo is also varying. As a result of asynchronous cleavage, embryos at the stage of five, six and seven cells might be found in one sponge. The octacellular embryo is rounded or slightly oval-shaped in the cross section. All the blastomeres are oval (
<figureCitation id="07A12A3FFFB2957F205FE8EAABF3E407" box="[378,462,1525,1551]" captionStart="FIG" captionStartId="4.[114,125,1616,1633]" captionTargetBox="[139,1177,204,1569]" captionTargetId="graphics-114@4.[139,1159,204,1693]" captionTargetPageId="4" captionText="FIG. 2. — A, cleaving embryos in 8-cell stage; B, blastula after immigration of external cells; C, stage of invagination of flagellate cells forming internal sphere; D, disphaerula of H. dujardini. Abbreviations: ec, embryonic capsule; ic, internal cells; n, nucleus; efc, external flagellate cells; b, blastomere; bi, blastocoel; ifc, internal flagellate cells; is, internal sphere; ppc, posterior pole cells; tc, transition cells. Scale bars: A, 20 µm; B, 15 µm; C, 20 µm; D, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5402180" httpUri="https://zenodo.org/record/5402180/files/figure.png" pageId="3" pageNumber="358">Fig. 2A</figureCitation>
).
</paragraph>
<paragraph id="9F2536BAFFB2957F2154EB0CA946E1C0" blockId="3.[113,647,205,1704]" lastBlockId="3.[677,1210,205,1704]" pageId="3" pageNumber="358">
Subsequent cleavage continues to be asynchronous but cleavage spindles in this case are positioned in parallel to the embryo surface; as a result division planes become radial. A single-layered coeloblastula consisting of 18 to 24 cells surrounding a small cavity is thus formed. Polarity of the embryo is indistinct. Large nuclei with nucleoli are located in the central part of the cells. Rounded vitelline granules of heterogeneous content and ranging between 0.2 to 2.4 µm in diameter occupy most of the volume of blastomere cytoplasm. These structures are pyroninephilous, their dimensions slightly increase with the distance from the nucleus. Beginning after the fourth to fifth cleavage cycle, polarization of the blastomeres occurs with the movement of nuclei in apical direction and vitelline granules in the basal direction (
<figureCitation id="07A12A3FFFB2957F2240EF21A9F9E050" box="[869,964,574,600]" captionStart="FIG" captionStartId="8.[114,125,789,806]" captionTargetBox="[124,1201,204,764]" captionTargetId="figure-453@8.[114,1229,196,767]" captionTargetPageId="8" captionText="FIG. 5. — Scheme of H. dujardini embryonic development; A, mature egg; B, early cleavage; C, beginning of blastomere polarization; D, multipolar immigration; E, stage larval polarization; F, formation of internal sphere and migration of maternal eosinophilic granular amoebocytes into larva; G, disphaerula, larva of H. dujardini. Abbreviations: bi, blastocoel; ec, embryonic capsule; efa, eosinophilic amoebocytes; ic, internal cells; isp, internal sphere; mc, migrated cells; ppc, posterior pole cells; tc, transition cells; yg, yolk granules." figureDoi="http://doi.org/10.5281/zenodo.5402186" httpUri="https://zenodo.org/record/5402186/files/figure.png" pageId="3" pageNumber="358">Fig. 5C</figureCitation>
). Cleavage planes of polarized blastomeres are radially directed resulting in the formation of blastula of 130 to 155 µm diameter with a small cavity (nearly 30 µm in diameter) limited by wedge-form cells. At the stage of about 100 cells, the apical-basal polarity of blastomeres is clearly indicated (
<figureCitation id="07A12A3FFFB2957F253FEFE9AE4CE118" box="[1050,1137,758,784]" captionStart="FIG" captionStartId="8.[114,125,789,806]" captionTargetBox="[124,1201,204,764]" captionTargetId="figure-453@8.[114,1229,196,767]" captionTargetPageId="8" captionText="FIG. 5. — Scheme of H. dujardini embryonic development; A, mature egg; B, early cleavage; C, beginning of blastomere polarization; D, multipolar immigration; E, stage larval polarization; F, formation of internal sphere and migration of maternal eosinophilic granular amoebocytes into larva; G, disphaerula, larva of H. dujardini. Abbreviations: bi, blastocoel; ec, embryonic capsule; efa, eosinophilic amoebocytes; ic, internal cells; isp, internal sphere; mc, migrated cells; ppc, posterior pole cells; tc, transition cells; yg, yolk granules." figureDoi="http://doi.org/10.5281/zenodo.5402186" httpUri="https://zenodo.org/record/5402186/files/figure.png" pageId="3" pageNumber="358">Fig. 5D</figureCitation>
). Such cells are about 70 µm in length and their apical breadth is nearly 25 µm; the rounded nucleus (about 9 µm in diameter) contains a nucleolus (about 2.5 µm). Sparse cisterns of Golgi apparatus and minute mitochondria are arranged around the nucleus.
</paragraph>
<paragraph id="9F2536BAFFB2957F2380EED2A93CE79C" blockId="3.[677,1210,205,1704]" pageId="3" pageNumber="358">
At this stage some cells of a blastula migrate into the cavity. This process is apolar. Before the beginning of migration, the nucleus is shifted to the basal part of the cell which in this case begins to expand (
<figureCitation id="07A12A3FFFB2957F2203E958A9BDE669" box="[806,896,1095,1121]" captionStart="FIG" captionStartId="8.[114,125,789,806]" captionTargetBox="[124,1201,204,764]" captionTargetId="figure-453@8.[114,1229,196,767]" captionTargetPageId="8" captionText="FIG. 5. — Scheme of H. dujardini embryonic development; A, mature egg; B, early cleavage; C, beginning of blastomere polarization; D, multipolar immigration; E, stage larval polarization; F, formation of internal sphere and migration of maternal eosinophilic granular amoebocytes into larva; G, disphaerula, larva of H. dujardini. Abbreviations: bi, blastocoel; ec, embryonic capsule; efa, eosinophilic amoebocytes; ic, internal cells; isp, internal sphere; mc, migrated cells; ppc, posterior pole cells; tc, transition cells; yg, yolk granules." figureDoi="http://doi.org/10.5281/zenodo.5402186" httpUri="https://zenodo.org/record/5402186/files/figure.png" pageId="3" pageNumber="358">Fig. 5C</figureCitation>
). The examination of serial sections has indicated that two to four cells migrate simultaneously. In the central cavity of the blastula those cells become spherical (about 27 µm in diameter), the nucleus with nucleolus occupying the central position (
<figureCitation id="07A12A3FFFB2957F253FE9FEAE49E6F3" box="[1050,1140,1249,1275]" captionStart="FIG" captionStartId="4.[114,125,1616,1633]" captionTargetBox="[139,1177,204,1569]" captionTargetId="graphics-114@4.[139,1159,204,1693]" captionTargetPageId="4" captionText="FIG. 2. — A, cleaving embryos in 8-cell stage; B, blastula after immigration of external cells; C, stage of invagination of flagellate cells forming internal sphere; D, disphaerula of H. dujardini. Abbreviations: ec, embryonic capsule; ic, internal cells; n, nucleus; efc, external flagellate cells; b, blastomere; bi, blastocoel; ifc, internal flagellate cells; is, internal sphere; ppc, posterior pole cells; tc, transition cells. Scale bars: A, 20 µm; B, 15 µm; C, 20 µm; D, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5402180" httpUri="https://zenodo.org/record/5402180/files/figure.png" pageId="3" pageNumber="358">Fig. 2B</figureCitation>
). The inner cells of embryo do not form a dense mass. Subsequently they start dividing with the ensuing differentiation into amoebocytes. At the same time flagella are formed at the apical end of external cells.
</paragraph>
<paragraph id="9F2536BAFFB2957A2380E886AB66E36A" blockId="3.[677,1210,205,1704]" lastBlockId="6.[114,647,205,1704]" lastPageId="6" lastPageNumber="361" pageId="3" pageNumber="358">
Flagellate cells continue to proliferate actively. Before its division, the cell becomes rounded and migrates to the periphery of the body where it starts to divide. The division plane is perpendicular to the larval surface (
<figureCitation id="07A12A3FFFB2957F2282EB0CA9C0E425" box="[935,1021,1555,1581]" captionStart="FIG" captionStartId="5.[113,124,1616,1633]" captionTargetBox="[143,1185,204,1571]" captionTargetId="graphics-25@5.[184,1209,204,1780]" captionTargetPageId="5" captionText="FIG. 3. — A, division of external flagellate cells in prelarva; B, maternal eosinophilic granular amoebocyte between internal cells of disphaerula; C, posterior pole of larva; D, external flagellate epithelium of larva. Abbreviations: ba, bacteria; ic, internal cells; ec, embryonic capsule; ppc, posterior pole cells; tc, transition cells; dc, divided cell; ecc, embryonic capsule cell; f, flagellum; n, nucleus; yg, yolk granules. Scale bars: A, 5 µm; B, 5 µm; C, 10 µm; D, 5 µm." figureDoi="http://doi.org/10.5281/zenodo.5402182" httpUri="https://zenodo.org/record/5402182/files/figure.png" pageId="3" pageNumber="358">Figs 3A</figureCitation>
;
<figureCitation id="07A12A3FFFB2957F252DEB0CAE7BE424" box="[1032,1094,1554,1581]" captionStart="FIG" captionStartId="8.[114,125,789,806]" captionTargetBox="[124,1201,204,764]" captionTargetId="figure-453@8.[114,1229,196,767]" captionTargetPageId="8" captionText="FIG. 5. — Scheme of H. dujardini embryonic development; A, mature egg; B, early cleavage; C, beginning of blastomere polarization; D, multipolar immigration; E, stage larval polarization; F, formation of internal sphere and migration of maternal eosinophilic granular amoebocytes into larva; G, disphaerula, larva of H. dujardini. Abbreviations: bi, blastocoel; ec, embryonic capsule; efa, eosinophilic amoebocytes; ic, internal cells; isp, internal sphere; mc, migrated cells; ppc, posterior pole cells; tc, transition cells; yg, yolk granules." figureDoi="http://doi.org/10.5281/zenodo.5402186" httpUri="https://zenodo.org/record/5402186/files/figure.png" pageId="3" pageNumber="358">5E, F</figureCitation>
). After the division, the daughter cells elongate and embed themselves into the columnar epithelium. At that time the epithelial cells are about 35 µm long and 5.6 µm wide near the nucleus, which itself is about 4.3 µm in diameter. Differentiation of cells of the posterior pole is occurring simultaneously with differentiation of anterior-lateral cells (
<figureCitation id="07A12A3FFFB7957A215CEC35AAF2E34C" box="[121,207,298,324]" captionStart="FIG" captionStartId="8.[114,125,789,806]" captionTargetBox="[124,1201,204,764]" captionTargetId="figure-453@8.[114,1229,196,767]" captionTargetPageId="8" captionText="FIG. 5. — Scheme of H. dujardini embryonic development; A, mature egg; B, early cleavage; C, beginning of blastomere polarization; D, multipolar immigration; E, stage larval polarization; F, formation of internal sphere and migration of maternal eosinophilic granular amoebocytes into larva; G, disphaerula, larva of H. dujardini. Abbreviations: bi, blastocoel; ec, embryonic capsule; efa, eosinophilic amoebocytes; ic, internal cells; isp, internal sphere; mc, migrated cells; ppc, posterior pole cells; tc, transition cells; yg, yolk granules." figureDoi="http://doi.org/10.5281/zenodo.5402186" httpUri="https://zenodo.org/record/5402186/files/figure.png" pageId="6" pageNumber="361">Fig. 5E</figureCitation>
). No division of internal cells has been recorded at this stage.
</paragraph>
<caption id="CBE56632FFB595782157EB4FA841E4A9" ID-DOI="http://doi.org/10.5281/zenodo.5402180" ID-Zenodo-Dep="5402180" httpUri="https://zenodo.org/record/5402180/files/figure.png" pageId="4" pageNumber="359" startId="4.[114,125,1616,1633]" targetBox="[139,1177,204,1569]" targetPageId="4">
<paragraph id="9F2536BAFFB595782157EB4FA841E4A9" blockId="4.[114,1210,1616,1697]" pageId="4" pageNumber="359">
FIG. 2. —
<emphasis id="ADEEEAA8FFB5957821E5EB4FAAF0E469" bold="true" box="[192,205,1616,1633]" pageId="4" pageNumber="359">A</emphasis>
, cleaving embryos in 8-cell stage;
<emphasis id="ADEEEAA8FFB5957820C9EB4FABC4E469" bold="true" box="[492,505,1616,1633]" pageId="4" pageNumber="359">B</emphasis>
, blastula after immigration of external cells;
<emphasis id="ADEEEAA8FFB595782242EB4FA948E469" bold="true" box="[871,885,1616,1633]" pageId="4" pageNumber="359">C</emphasis>
, stage of invagination of flagellate cells forming internal sphere;
<emphasis id="ADEEEAA8FFB59578201EEB7AAB74E47E" bold="true" box="[315,329,1637,1654]" pageId="4" pageNumber="359">D</emphasis>
, disphaerula of
<taxonomicName id="589A4D39FFB5957820E8EB7AA816E47E" authorityName="Johnston. Vestnik" authorityYear="1842" box="[461,555,1637,1654]" class="Demospongiae" family="Halisarcidae" genus="Halisarca" kingdom="Animalia" order="Chondrillida" pageId="4" pageNumber="359" phylum="Porifera" rank="species" species="dujardini">
<emphasis id="ADEEEAA8FFB5957820E8EB7AA816E47E" box="[461,555,1637,1654]" italics="true" pageId="4" pageNumber="359">H. dujardini</emphasis>
</taxonomicName>
. Abbreviations:
<emphasis id="ADEEEAA8FFB595782397EB7AA8FAE47E" bold="true" box="[690,711,1637,1654]" pageId="4" pageNumber="359">ec</emphasis>
, embryonic capsule;
<emphasis id="ADEEEAA8FFB59578225CEB7AA9B4E47E" bold="true" box="[889,905,1637,1654]" pageId="4" pageNumber="359">ic</emphasis>
, internal cells;
<emphasis id="ADEEEAA8FFB595782526EB7AAE33E47E" bold="true" box="[1027,1038,1637,1654]" pageId="4" pageNumber="359">n</emphasis>
, nucleus;
<emphasis id="ADEEEAA8FFB595782546EB7AAE42E47E" bold="true" box="[1123,1151,1637,1654]" pageId="4" pageNumber="359">efc</emphasis>
, external flagellate cells;
<emphasis id="ADEEEAA8FFB595782034EB64AB21E484" bold="true" box="[273,284,1659,1676]" pageId="4" pageNumber="359">b</emphasis>
, blastomere;
<emphasis id="ADEEEAA8FFB5957820B5EB64AB9DE484" bold="true" box="[400,416,1659,1676]" pageId="4" pageNumber="359">bi</emphasis>
, blastocoel;
<emphasis id="ADEEEAA8FFB595782329EB64A81FE484" bold="true" box="[524,546,1659,1676]" pageId="4" pageNumber="359">ifc</emphasis>
, internal flagellate cells;
<emphasis id="ADEEEAA8FFB5957823D5EB64A8C2E484" bold="true" box="[752,767,1659,1676]" pageId="4" pageNumber="359">is</emphasis>
, internal sphere;
<emphasis id="ADEEEAA8FFB5957822B4EB64A98EE484" bold="true" box="[913,947,1659,1676]" pageId="4" pageNumber="359">ppc</emphasis>
, posterior pole cells;
<emphasis id="ADEEEAA8FFB59578254DEB64AE44E484" bold="true" box="[1128,1145,1659,1676]" pageId="4" pageNumber="359">tc</emphasis>
, transition cells. Scale bars: A, 20 µm; B, 15 µm; C, 20 µm; D, 10 µm.
</paragraph>
</caption>
<caption id="CBE56632FFB495792154EB4FA882E4A9" ID-DOI="http://doi.org/10.5281/zenodo.5402182" ID-Zenodo-Dep="5402182" httpUri="https://zenodo.org/record/5402182/files/figure.png" pageId="5" pageNumber="360" startId="5.[113,124,1616,1633]" targetBox="[143,1185,204,1571]" targetPageId="5">
<paragraph id="9F2536BAFFB495792154EB4FA882E4A9" blockId="5.[113,1209,1616,1697]" pageId="5" pageNumber="360">
FIG. 3. —
<emphasis id="ADEEEAA8FFB4957921E0EB4FAAEFE469" bold="true" box="[197,210,1616,1633]" pageId="5" pageNumber="360">A</emphasis>
, division of external flagellate cells in prelarva;
<emphasis id="ADEEEAA8FFB495792347EB4FA852E469" bold="true" box="[610,623,1616,1633]" pageId="5" pageNumber="360">B</emphasis>
, maternal eosinophilic granular amoebocyte between internal cells of disphaerula;
<emphasis id="ADEEEAA8FFB4957921FAEB7AAAD0E47E" bold="true" box="[223,237,1637,1654]" pageId="5" pageNumber="360">C</emphasis>
, posterior pole of larva;
<emphasis id="ADEEEAA8FFB49579209AEB7AABF0E47E" bold="true" box="[447,461,1637,1654]" pageId="5" pageNumber="360">D</emphasis>
, external flagellate epithelium of larva. Abbreviations:
<emphasis id="ADEEEAA8FFB4957922BEEB7AA98CE47E" bold="true" box="[923,945,1637,1654]" pageId="5" pageNumber="360">ba</emphasis>
, bacteria;
<emphasis id="ADEEEAA8FFB495792528EB7AAE20E47E" bold="true" box="[1037,1053,1637,1654]" pageId="5" pageNumber="360">ic</emphasis>
, internal cells;
<emphasis id="ADEEEAA8FFB4957925BAEB7AAE89E47E" bold="true" box="[1183,1204,1637,1654]" pageId="5" pageNumber="360">ec</emphasis>
, embryonic capsule;
<emphasis id="ADEEEAA8FFB49579203FEB64AB01E484" bold="true" box="[282,316,1659,1676]" pageId="5" pageNumber="360">ppc</emphasis>
, posterior pole cells;
<emphasis id="ADEEEAA8FFB4957920CBEB64ABC2E484" bold="true" box="[494,511,1659,1676]" pageId="5" pageNumber="360">tc</emphasis>
, transition cells;
<emphasis id="ADEEEAA8FFB4957923A9EB64A89FE484" bold="true" box="[652,674,1659,1676]" pageId="5" pageNumber="360">dc</emphasis>
, divided cell;
<emphasis id="ADEEEAA8FFB495792230EB64A908E484" bold="true" box="[789,821,1659,1676]" pageId="5" pageNumber="360">ecc</emphasis>
, embryonic capsule cell;
<emphasis id="ADEEEAA8FFB49579252FEB64AE2DE484" bold="true" box="[1034,1040,1659,1676]" pageId="5" pageNumber="360">f</emphasis>
, flagellum;
<emphasis id="ADEEEAA8FFB495792555EB64AE46E484" bold="true" box="[1136,1147,1659,1676]" pageId="5" pageNumber="360">n</emphasis>
, nucleus;
<emphasis id="ADEEEAA8FFB4957921AAEB8FAA99E4A9" bold="true" box="[143,164,1680,1697]" pageId="5" pageNumber="360">yg</emphasis>
, yolk granules. Scale bars: A, 5 µm; B, 5 µm; C, 10 µm; D, 5 µm.
</paragraph>
</caption>
<paragraph id="9F2536BAFFB7957A2157EC78ABEBE050" blockId="6.[114,647,205,1704]" pageId="6" pageNumber="361">
During the period of active proliferation of the external cells, the larval surface becomes corrugated. The internal part of larva is a cavity surrounded by basal parts of flagellate cells. At the same developmental stage, granular eosinophilic (fuchsinophilic) amoebocytes migrate from the maternal sponge mesohyl into the larva via the embryonic capsule (
<figureCitation id="07A12A3FFFB7957A206DEF21ABA0E050" box="[328,413,574,600]" captionStart="FIG" captionStartId="5.[113,124,1616,1633]" captionTargetBox="[143,1185,204,1571]" captionTargetId="graphics-25@5.[184,1209,204,1780]" captionTargetPageId="5" captionText="FIG. 3. — A, division of external flagellate cells in prelarva; B, maternal eosinophilic granular amoebocyte between internal cells of disphaerula; C, posterior pole of larva; D, external flagellate epithelium of larva. Abbreviations: ba, bacteria; ic, internal cells; ec, embryonic capsule; ppc, posterior pole cells; tc, transition cells; dc, divided cell; ecc, embryonic capsule cell; f, flagellum; n, nucleus; yg, yolk granules. Scale bars: A, 5 µm; B, 5 µm; C, 10 µm; D, 5 µm." figureDoi="http://doi.org/10.5281/zenodo.5402182" httpUri="https://zenodo.org/record/5402182/files/figure.png" pageId="6" pageNumber="361">Figs 3B</figureCitation>
;
<figureCitation id="07A12A3FFFB7957A208CEF21ABF4E050" box="[425,457,574,600]" captionStart="FIG" captionStartId="8.[114,125,789,806]" captionTargetBox="[124,1201,204,764]" captionTargetId="figure-453@8.[114,1229,196,767]" captionTargetPageId="8" captionText="FIG. 5. — Scheme of H. dujardini embryonic development; A, mature egg; B, early cleavage; C, beginning of blastomere polarization; D, multipolar immigration; E, stage larval polarization; F, formation of internal sphere and migration of maternal eosinophilic granular amoebocytes into larva; G, disphaerula, larva of H. dujardini. Abbreviations: bi, blastocoel; ec, embryonic capsule; efa, eosinophilic amoebocytes; ic, internal cells; isp, internal sphere; mc, migrated cells; ppc, posterior pole cells; tc, transition cells; yg, yolk granules." figureDoi="http://doi.org/10.5281/zenodo.5402186" httpUri="https://zenodo.org/record/5402186/files/figure.png" pageId="6" pageNumber="361">5F</figureCitation>
).
</paragraph>
<paragraph id="9F2536BAFFB7957A2157EF42AB7BE60D" blockId="6.[114,647,205,1704]" pageId="6" pageNumber="361">
Gradually the larval surface becomes flattened, proliferation is discontinued and the cells acquire the shape and size characteristic of a mature larva. Before the surface becomes finally smoothed, a deep and narrow invagination of the flagellate layer occurs, closing near its mouth (
<figureCitation id="07A12A3FFFB7957A2338EFE9A844E118" box="[541,633,758,784]" captionStart="FIG" captionStartId="4.[114,125,1616,1633]" captionTargetBox="[139,1177,204,1569]" captionTargetId="graphics-114@4.[139,1159,204,1693]" captionTargetPageId="4" captionText="FIG. 2. — A, cleaving embryos in 8-cell stage; B, blastula after immigration of external cells; C, stage of invagination of flagellate cells forming internal sphere; D, disphaerula of H. dujardini. Abbreviations: ec, embryonic capsule; ic, internal cells; n, nucleus; efc, external flagellate cells; b, blastomere; bi, blastocoel; ifc, internal flagellate cells; is, internal sphere; ppc, posterior pole cells; tc, transition cells. Scale bars: A, 20 µm; B, 15 µm; C, 20 µm; D, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5402180" httpUri="https://zenodo.org/record/5402180/files/figure.png" pageId="6" pageNumber="361">Fig. 2C</figureCitation>
). Invagination of this layer proceeds perpendicular or at an angle to the anterior-posterior axis of the larva. A single-layered closed structure surrounding a small cavity is formed by the internalized flagellate cells as a result of this process. The blastocoel is reduced, persisting in the form of narrow gaps between the internal and external flagellate cell layers.
</paragraph>
<paragraph id="9F2536BAFFB7957A2157E915AE33E3A8" blockId="6.[114,647,205,1704]" lastBlockId="6.[677,1210,205,1704]" pageId="6" pageNumber="361">
The form and shape of the closed structures newly formed by flagellate cells inside the larva may be widely variable. These formations have mostly the form of C- or ∑- shaped, curved cylinders; sometimes they are oval or spherical; their cross-sections are tubular-shaped. The inner cavity diameter ranges between 8.5 µm and 12.5 µm while inner cavity length varies from 28 µm to 61 µm. One or two cells may be often found inside the cavity.This stage terminates the larval development. The walls of embryonic capsules which contain newly formed larvae fuse with the walls of exhalant channels and larvae escape through the osculum. Milk white-colored larvae of
<taxonomicName id="589A4D39FFB7957A204CE887ABD0E7BB" authorityName="Johnston. Vestnik" authorityYear="1842" box="[361,493,1432,1459]" class="Demospongiae" family="Halisarcidae" genus="Halisarca" kingdom="Animalia" order="Chondrillida" pageId="6" pageNumber="361" phylum="Porifera" rank="species" species="dujardini">
<emphasis id="ADEEEAA8FFB7957A204CE887ABD0E7BB" box="[361,493,1432,1459]" italics="true" pageId="6" pageNumber="361">H. dujardini</emphasis>
</taxonomicName>
emerged from the maternal organism are oval or have the shape of a spheroid slightly depressed in the anterior-posterior direction with flat or concave posterior pole (
<figureCitation id="07A12A3FFFB7957A215CEB0CAAEEE425" box="[121,211,1555,1581]" captionStart="FIG" captionStartId="4.[114,125,1616,1633]" captionTargetBox="[139,1177,204,1569]" captionTargetId="graphics-114@4.[139,1159,204,1693]" captionTargetPageId="4" captionText="FIG. 2. — A, cleaving embryos in 8-cell stage; B, blastula after immigration of external cells; C, stage of invagination of flagellate cells forming internal sphere; D, disphaerula of H. dujardini. Abbreviations: ec, embryonic capsule; ic, internal cells; n, nucleus; efc, external flagellate cells; b, blastomere; bi, blastocoel; ifc, internal flagellate cells; is, internal sphere; ppc, posterior pole cells; tc, transition cells. Scale bars: A, 20 µm; B, 15 µm; C, 20 µm; D, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5402180" httpUri="https://zenodo.org/record/5402180/files/figure.png" pageId="6" pageNumber="361">Figs 2D</figureCitation>
;
<figureCitation id="07A12A3FFFB7957A21FAEB0CAB38E425" box="[223,261,1555,1581]" captionStart="FIG" captionStartId="8.[114,125,789,806]" captionTargetBox="[124,1201,204,764]" captionTargetId="figure-453@8.[114,1229,196,767]" captionTargetPageId="8" captionText="FIG. 5. — Scheme of H. dujardini embryonic development; A, mature egg; B, early cleavage; C, beginning of blastomere polarization; D, multipolar immigration; E, stage larval polarization; F, formation of internal sphere and migration of maternal eosinophilic granular amoebocytes into larva; G, disphaerula, larva of H. dujardini. Abbreviations: bi, blastocoel; ec, embryonic capsule; efa, eosinophilic amoebocytes; ic, internal cells; isp, internal sphere; mc, migrated cells; ppc, posterior pole cells; tc, transition cells; yg, yolk granules." figureDoi="http://doi.org/10.5281/zenodo.5402186" httpUri="https://zenodo.org/record/5402186/files/figure.png" pageId="6" pageNumber="361">5G</figureCitation>
). They are completely covered with flagella, however in the posterior pole region, the flagella are sparse. The diameter of flagella-sparse cell layer is about 45 µm. Anterio-lateral larval flagellate cells are nearly cylindrical, 38 to 45 µm long and 2.6 to 3.0 µm wide in the nucleus region. The prolate nucleus (2.6 × 6.6 µm) with small nucleolus and the flagellum emerging from a pocketshaped cytoplasm depression are located in their apical part (
<figureCitation id="07A12A3FFFB7957A2204EC56A94AE36B" box="[801,887,329,355]" captionStart="FIG" captionStartId="5.[113,124,1616,1633]" captionTargetBox="[143,1185,204,1571]" captionTargetId="graphics-25@5.[184,1209,204,1780]" captionTargetPageId="5" captionText="FIG. 3. — A, division of external flagellate cells in prelarva; B, maternal eosinophilic granular amoebocyte between internal cells of disphaerula; C, posterior pole of larva; D, external flagellate epithelium of larva. Abbreviations: ba, bacteria; ic, internal cells; ec, embryonic capsule; ppc, posterior pole cells; tc, transition cells; dc, divided cell; ecc, embryonic capsule cell; f, flagellum; n, nucleus; yg, yolk granules. Scale bars: A, 5 µm; B, 5 µm; C, 10 µm; D, 5 µm." figureDoi="http://doi.org/10.5281/zenodo.5402182" httpUri="https://zenodo.org/record/5402182/files/figure.png" pageId="6" pageNumber="361">Fig. 3D</figureCitation>
). The basal two thirds of these cells is filled with abundant vitelline granules. This cell layer lacks a basal membrane.
</paragraph>
<paragraph id="9F2536BAFFB7957A2380ECBBA9EAE1A1" blockId="6.[677,1210,205,1704]" pageId="6" pageNumber="361">
In longitudinal section the flagellate cells of the posterior pole are wedge-shaped; they are shorter and wider than the anterio-lateral cells (26 µm long and 6-7 µm wide in the nucleus region). Numerous vitelline granules are spread not only in basal and central parts of the cell, but near the nucleus as well. Rounded or drop-shaped nuclei are about 5.3 × 4.2 µm in diameter and contain large nucleoli (near 1.6 µm) (
<figureCitation id="07A12A3FFFB7957A2223EF85A964E0BC" box="[774,857,666,692]" captionStart="FIG" captionStartId="5.[113,124,1616,1633]" captionTargetBox="[143,1185,204,1571]" captionTargetId="graphics-25@5.[184,1209,204,1780]" captionTargetPageId="5" captionText="FIG. 3. — A, division of external flagellate cells in prelarva; B, maternal eosinophilic granular amoebocyte between internal cells of disphaerula; C, posterior pole of larva; D, external flagellate epithelium of larva. Abbreviations: ba, bacteria; ic, internal cells; ec, embryonic capsule; ppc, posterior pole cells; tc, transition cells; dc, divided cell; ecc, embryonic capsule cell; f, flagellum; n, nucleus; yg, yolk granules. Scale bars: A, 5 µm; B, 5 µm; C, 10 µm; D, 5 µm." figureDoi="http://doi.org/10.5281/zenodo.5402182" httpUri="https://zenodo.org/record/5402182/files/figure.png" pageId="6" pageNumber="361">Fig. 3C</figureCitation>
). A ring of tapering flagellate cells is arranged between cells of the posterior pole and covering flagellate cells. The length of transitional cells fluctuates around 30 µm, the width in the nucleus region is about 4 µm; nuclei with nucleoli are oval-shaped (3.5 × 4.9 µm) (
<figureCitation id="07A12A3FFFB7957A2505EE2CAE4AE145" box="[1056,1143,819,845]" captionStart="FIG" captionStartId="5.[113,124,1616,1633]" captionTargetBox="[143,1185,204,1571]" captionTargetId="graphics-25@5.[184,1209,204,1780]" captionTargetPageId="5" captionText="FIG. 3. — A, division of external flagellate cells in prelarva; B, maternal eosinophilic granular amoebocyte between internal cells of disphaerula; C, posterior pole of larva; D, external flagellate epithelium of larva. Abbreviations: ba, bacteria; ic, internal cells; ec, embryonic capsule; ppc, posterior pole cells; tc, transition cells; dc, divided cell; ecc, embryonic capsule cell; f, flagellum; n, nucleus; yg, yolk granules. Scale bars: A, 5 µm; B, 5 µm; C, 10 µm; D, 5 µm." figureDoi="http://doi.org/10.5281/zenodo.5402182" httpUri="https://zenodo.org/record/5402182/files/figure.png" pageId="6" pageNumber="361">Fig. 3C</figureCitation>
). The number of cells in the transitional zone does not exceed four to five. There are no specialized contacts between the larval cells.
</paragraph>
<paragraph id="9F2536BAFFB7957A2380EEB1A9FEE7F8" blockId="6.[677,1210,205,1704]" pageId="6" pageNumber="361">
A most striking feature of the larval structure of
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<emphasis id="ADEEEAA8FFB7957A2380EED3A913E1EF" box="[677,814,972,999]" italics="true" pageId="6" pageNumber="361">H. dujardini</emphasis>
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is the presence of a large closed formation composed of flagellate cells (derivative of the prelarval flagellate layer) inside the larva. This structure is spherical and has a cavity delimited by the apical tips of the internalized flagellate cells. The external diameter (over basal cell apexes) of spherical layer ranges from 49 µm to 54 and the diameter of its cavity from 13.8 µm to 17.2 µm (
<figureCitation id="07A12A3FFFB7957A2388E9DDA931E6D4" box="[685,780,1218,1244]" captionStart="FIG" captionStartId="4.[114,125,1616,1633]" captionTargetBox="[139,1177,204,1569]" captionTargetId="graphics-114@4.[139,1159,204,1693]" captionTargetPageId="4" captionText="FIG. 2. — A, cleaving embryos in 8-cell stage; B, blastula after immigration of external cells; C, stage of invagination of flagellate cells forming internal sphere; D, disphaerula of H. dujardini. Abbreviations: ec, embryonic capsule; ic, internal cells; n, nucleus; efc, external flagellate cells; b, blastomere; bi, blastocoel; ifc, internal flagellate cells; is, internal sphere; ppc, posterior pole cells; tc, transition cells. Scale bars: A, 20 µm; B, 15 µm; C, 20 µm; D, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5402180" httpUri="https://zenodo.org/record/5402180/files/figure.png" pageId="6" pageNumber="361">Fig. 2D</figureCitation>
). The cells forming this structure are wedge-shaped with the extended basal part concentrating numerous vitelline granules. the flagellum, submerged into a pocket-shaped depression, and the nucleus are located near the narrow apical pole. Cell length varies from 13.8 µm to 17.2 µm, cells width from 3.4 µm to 5.7 µm. Drop-shaped or egg-form nuclei (2.3 µm × 5,8 µm) in these cell contain clearly distinguished nucleoli of about 0.9 µm diameter (
<figureCitation id="07A12A3FFFB7957A2246E8C9A98AE7F8" box="[867,951,1494,1520]" captionStart="FIG" captionStartId="7.[113,124,1616,1633]" captionTargetBox="[132,627,205,1588]" captionTargetId="figure-413@7.[130,629,205,897]" captionTargetPageId="7" captionText="FIG. 4. — A, internal flagellate cells of disphaerula; B, flagellated cell that migrated from internal sphere. Abbreviations: f, flagellum; isc, internal sphere cavity; n, nucleus; yg, yolk granules; ba, bacteria. Scale bars: A, 4 µm; B, 1 µm." figureDoi="http://doi.org/10.5281/zenodo.5402184" httpUri="https://zenodo.org/record/5402184/files/figure.png" pageId="6" pageNumber="361">Fig. 4A</figureCitation>
).
</paragraph>
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Before larval emergence out of maternal tissues, the interior chambers become spherical and their dimensions decrease. The process of cell ejection seems to be the cause of these changes. Numerous intermediate cells are located among internal cells in the space between the internal and external epithelia. They are oval-shaped, one pole has a flagellum, near which a rounded nucleus is located. The cytoplasm contains numerous vitelline granules (
<figureCitation id="07A12A3FFFB6957B2229EC35A95DE34C" box="[780,864,298,324]" captionStart="FIG" captionStartId="7.[113,124,1616,1633]" captionTargetBox="[132,627,205,1588]" captionTargetId="figure-413@7.[130,629,205,897]" captionTargetPageId="7" captionText="FIG. 4. — A, internal flagellate cells of disphaerula; B, flagellated cell that migrated from internal sphere. Abbreviations: f, flagellum; isc, internal sphere cavity; n, nucleus; yg, yolk granules; ba, bacteria. Scale bars: A, 4 µm; B, 1 µm." figureDoi="http://doi.org/10.5281/zenodo.5402184" httpUri="https://zenodo.org/record/5402184/files/figure.png" pageId="7" pageNumber="362">Fig. 4B</figureCitation>
).
</paragraph>
<paragraph id="9F2536BAFFB6957B2380EC57AE79E031" blockId="7.[677,1210,205,1673]" pageId="7" pageNumber="362">Interspaces between outer and inner flagellate cells are occupied by the internal cells of the larva: nucleolate amoebocytes. In addition, granular eosinophilic cells of maternal origin are included into the larva. They may be located in different zones: apical, central, basal regions of flagellate layer, in the central part of larva, between the cells of the posterior pole.</paragraph>
<paragraph id="9F2536BAFFB6957B2380EF22A903E0F9" blockId="7.[677,1210,205,1673]" pageId="7" pageNumber="362">Larvae have a spiral swimming movement counter-clockwise at the same time rotating themselves clockwise around the anterior-posterior axis. Sometimes they perform twirling movements. The free-living period of the larva lasts for 4 to 36 hours.</paragraph>
<paragraph id="9F2536BAFFB6957B2380EFEAAEA4E183" blockId="7.[677,1210,205,1673]" pageId="7" pageNumber="362">Larvae settle on a substrate with their anterior pole. However, even on a substrate, larvae continue to rotate around their anterior-posterior axis for 30 to 45 minutes before attachment. This rotation is finished after complete attachment.</paragraph>
<paragraph id="9F2536BAFFB6957B2380EE90A9ECE688" blockId="7.[677,1210,205,1673]" pageId="7" pageNumber="362">Preparation for metamorphosis begins already during the swimming period as evidenced in larval behaviour and structure. Premetamorphic larvae are slower in their swimming, may attach themselves to surface film or sink and slowly move over the substrate. Whether attached to surface film or moving over the substrate, the larvae proceed slowly rotating.</paragraph>
<paragraph id="9F2536BAFFB6957B2380E99AA9E7E77D" blockId="7.[677,1210,205,1673]" pageId="7" pageNumber="362">Shape variation of the external flagellate cells and violation of their mutual contacts may primarily be observed on ultrastructural level. The lateral surfaces of cells become corrugated and the cells are contracted in the apical-basal direction. However, flagellate cells remain interconnected in their apical parts. In this state larvae settle down on substrate and attach to it.</paragraph>
<paragraph id="9F2536BAFFB6957B2380E865AEADE444" blockId="7.[677,1210,205,1673]" pageId="7" pageNumber="362">During the first stages of settling and metamorphosis, the larva slightly flattens in the anterior-posterior direction; its anterior pole flattens on the substrate, while the posterior one remains rounded. The total disintegration of the internal spherical chamber and external flagellate layer takes place during the first 12 hours after the attachment.</paragraph>
<caption id="CBE56632FFB6957B2154EB4FABE8E4A9" ID-DOI="http://doi.org/10.5281/zenodo.5402184" ID-Zenodo-Dep="5402184" httpUri="https://zenodo.org/record/5402184/files/figure.png" pageId="7" pageNumber="362" startId="7.[113,124,1616,1633]" targetBox="[132,627,205,1588]" targetPageId="7">
<paragraph id="9F2536BAFFB6957B2154EB4FABE8E4A9" blockId="7.[113,648,1616,1697]" pageId="7" pageNumber="362">
FIG. 4. —
<emphasis id="ADEEEAA8FFB6957B21E7EB4FAAF2E469" bold="true" box="[194,207,1616,1633]" pageId="7" pageNumber="362">A</emphasis>
, internal flagellate cells of disphaerula;
<emphasis id="ADEEEAA8FFB6957B233CEB4FA81BE469" bold="true" box="[537,550,1616,1633]" pageId="7" pageNumber="362">B</emphasis>
, flagellated cell that migrated from internal sphere. Abbreviations:
<emphasis id="ADEEEAA8FFB6957B2365EB7AA87BE47E" bold="true" box="[576,582,1637,1654]" pageId="7" pageNumber="362">f</emphasis>
, flagellum;
<emphasis id="ADEEEAA8FFB6957B21B8EB64AA8AE484" bold="true" box="[157,183,1659,1676]" pageId="7" pageNumber="362">isc</emphasis>
, internal sphere cavity;
<emphasis id="ADEEEAA8FFB6957B20ADEB64ABAEE484" bold="true" box="[392,403,1659,1676]" pageId="7" pageNumber="362">n</emphasis>
, nucleus;
<emphasis id="ADEEEAA8FFB6957B20C8EB64A83FE484" bold="true" box="[493,514,1659,1676]" pageId="7" pageNumber="362">yg</emphasis>
, yolk granules;
<emphasis id="ADEEEAA8FFB6957B2154EB8FAABAE4A9" bold="true" box="[113,135,1680,1697]" pageId="7" pageNumber="362">ba</emphasis>
, bacteria. Scale bars: A, 4 µm; B, 1 µm.
</paragraph>
</caption>
<paragraph id="9F2536BAFFB6957B2380EB4EAE5AE481" blockId="7.[677,1210,205,1673]" pageId="7" pageNumber="362">Subsequent processes associated with metamorphosis will be described in a special study.</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="1958F60B13970BA9E4283A338F13310E" pageNumber="277">
Camponotus (Myrmotrema) Perrisi For.
<taxonomicName id="BDEFCB490607CDEBC1A455F98C71946E" authority="Santschi" authorityName="Santschi" rank="subSpecies" subSpecies="nigeriensis">stirps nigeriensis Santschi</taxonomicName>
(Bayeri For.
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<taxonomicName id="5C13E54C8BD45FA63681FA23EEF01548" authority="Santschi" authorityName="Santschi" class="Collembola" family="Paronellidae" genus="Camponotus" kingdom="Animalia" order="Entomobryomorpha" phylum="Arthropoda" rank="subSpecies" species="perrisi" subGenus="Myrmotrema" subSpecies="nigeriensis">Camponotus (Myrmotrema) perrisi For. stirps nigeriensis Santschi</taxonomicName>
(
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, - Bull. lab. Zool. gen. e agr. Portici, VIII [1914]).
</paragraph>
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1.
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<taxonomicName id="6304F3BB51EF04E8894EF7DFBB8D8123" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:133571" class="Eurotiomycetes" family="Verrucariaceae" genus="Stigmatomma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName-HNS="Stigmatomma bruni r. juergi Forel" order="Verrucariales" pageNumber="87" phylum="Ascomycota" rank="subSpecies" species="bruni" subSpecies="juergi">Stigmatomma bruni For. r. juergi</taxonomicName>
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<taxonomicNameLabel id="23F9E31223C661B35F0FA5C1E1119612" rank="subSpecies">n. st.</taxonomicNameLabel>
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Longueur: Au moins 5 mm. Differe comme suit de la
<taxonomicName id="E18A762D84DCF8D70C2C3D86E85A60AB" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:133570" class="Eurotiomycetes" family="Verrucariaceae" genus="Stigmatomma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName-HNS="Stigmatomma bruni Forel" order="Verrucariales" pageNumber="87" phylum="Ascomycota" rank="species" species="bruni">St. bruni For.</taxonomicName>
<taxonomicName id="E18A762D84DCF8D70C2C3D86E85A60AB" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:133570" class="Insecta" family="Formicidae" genus="Stigmatomma" kingdom="Animalia" lsidName-HNS="Stigmatomma bruni Forel" order="Hymenoptera" pageNumber="87" phylum="Arthropoda" rank="species" species="bruni">St. bruni For.</taxonomicName>
s. str. Les mandibules soyeuses ont un bord terminal plus ou moins distinct, mais leurs dents sont analogues; elles sont densement striees-ridees en long. Epistome sans dents a son bord inferieur. La tete est plus large a son bord anterieur qu'elle n'est longue; son bord posterieur vu de face, est a peu pres droit, a angles plus arrondis. Les yeux ont plus de facettes. Le mesonotum est encore plus etroit. La face basale de l'epi- notum est encore plus large que chez le type et la face declive bien plus bordee que chez lui. Le petiole est beaucoup plus large, presque aussi large que le postpetiole, moins convexe de cote et devant; mais les etranglements sont les memes. La sculpture est plus faible. Corps assez lisse, et luisant sur le thorax et l'abdomen qui n'ont que des points epars. La tete sub-opaque est plutot faiblement ridee en travers et pas en long.
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Meme localite que la
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<taxonomicName id="9375A6D0DE9B4A4D6A1EE6439C60D9F5" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:133570" class="Insecta" family="Formicidae" genus="Stigmatomma" kingdom="Animalia" lsidName-HNS="Stigmatomma bruni Forel" order="Hymenoptera" pageNumber="87" phylum="Arthropoda" rank="species" species="bruni">St. bruni</taxonomicName>
type: Pilam Formose (Sauter).
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S. Clytemnestra
<taxonomicName id="CC4995C752142368C732BB59528C42DB" ID-CoL="4Y6D4" class="Insecta" family="Formicidae" genus="Solenopsis" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="clytemnestra">S. clytemnestra</taxonomicName>
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Camponotus [Myrmoturba) maculatus F.
<taxonomicName id="DAE0DFB4A7D00FCAEC6489AFC1541FCA" authority="Santschi." authorityName="Santschi." rank="subSpecies" subSpecies="melanocnemis">stirps melanocnemis Santschi.</taxonomicName>
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— (Fig. 14 a).
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<taxonomicName id="E758EDFF87AEA96FCBD90672DEA7D926" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:138659" class="Sordariomycetes" genus="Alfaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Alfaria minuta Emery" lsidName-HNS="Alfaria minuta Emery" order="Hypocreales" pageNumber="74" phylum="Ascomycota" rank="species" species="minuta">Alfaria minuta</taxonomicName>
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<taxonomicNameLabel id="8294B49C9F210C0B447A374E722F9EB8" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="1F9D5D2A6B4FC31783A16170B099A006" pageNumber="74">
[[ queen ]] Bruno-ferrugineo, le zampe, antenne e mandibole put chiare; opaca, anche le zampe; il 2.0 e il 3.0 segmento dell'addome meno opachi del resto. Tutto l'insetto, con le zampe e Io scapo e irto di peli fini, lunghi e obliqui. Il tegumento non e striato, ma coperto di una punteggiatura fondamentale poco fitta, meno distinta sull' addome ad esclusione del peduncolo; evvi pure una sottoscultura microscopica. Capo, torace e peduncolo hanno inoltre grosse fossette piligere, profonde e alquanto confiuenti sul capo; esse sono piu. deboli sul 2.0 segmento dell'addome, piu deboli ancora e rudimentali sul 3. °. Le antenne sono piu grosse e piu corte che nell'
<taxonomicName id="6CB0BF5D57A1D42FBD6A13D21C8E3FF7" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:138690" class="Sordariomycetes" genus="Alfaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Alfaria simulans Emery" lsidName-HNS="Alfaria simulans Emery" order="Hypocreales" pageNumber="74" phylum="Ascomycota" rank="species" species="simulans">Alfaria simulans</taxonomicName>
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e lo scapo non raggiunge interarnente l'occipite. Le mandibole sono striate e un poco lucide al margine masticatorio. Il torace e robusto e largo, rotondato in avanti. Lo scutello poco prominente, per cui il profilo forma una curva appena sinuata dietro lo scutello e un poco gibbosa tra la faccia basale e la faccia declive del metanoto. Le tibie sono molto corte e grosse. Le ali sono debolmente affumicate, con venatura e stigma bruno scuro. Il ramo esterno della costa cubitale e largamente interrotto alla base per cui la venatura dell' ala anteriore ricorda il genere
<taxonomicName id="0C72FFA4DE51CC8B3A0CFA4B0911E031" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24868" class="Insecta" family="Formicidae" genus="Myrmica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myrmica Latreille" lsidName-HNS="Myrmica Latreille" order="Hymenoptera" pageNumber="74" phylum="Arthropoda" rank="genus">Myrmica</taxonomicName>
<taxonomicName id="0C72FFA4DE51CC8B3A0CFA4B0911E031" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24868" class="Insecta" family="Formicidae" genus="Myrmica" kingdom="Animalia" lsidName="Myrmica Latreille" lsidName-HNS="Myrmica Latreille" order="Hymenoptera" pageNumber="74" phylum="Arthropoda" rank="genus">Myrmica</taxonomicName>
(un vestigio di quella interruzione si osserva pure nel presunto [[ male ]] dell'
<taxonomicName id="0E4BA89F377702E0BBBD67F68A5BC033" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:138690" class="Sordariomycetes" genus="Alfaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Alfaria simulans Emery" lsidName-HNS="Alfaria simulans Emery" order="Hypocreales" pageNumber="74" phylum="Ascomycota" rank="species" species="simulans">A. simulans</taxonomicName>
<taxonomicName id="0E4BA89F377702E0BBBD67F68A5BC033" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:138690" class="Insecta" family="Formicidae" genus="Alfaria" kingdom="Animalia" lsidName="Alfaria simulans Emery" lsidName-HNS="Alfaria simulans Emery" order="Hymenoptera" pageNumber="74" phylum="Arthropoda" rank="species" species="simulans">A. simulans</taxonomicName>
e in altri
<taxonomicName id="039AA15DEB76AB0281E583CF8D29E624" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2307" lsidName-HNS="Ponerini Lepeletier" pageNumber="74" rank="tribe">Ponerini</taxonomicName>
<taxonomicName id="039AA15DEB76AB0281E583CF8D29E624" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2307" class="Insecta" family="Formicidae" kingdom="Animalia" lsidName-HNS="Ponerini Lepeletier" order="Hymenoptera" pageNumber="74" phylum="Arthropoda" rank="tribe">Ponerini</taxonomicName>
). Peduncolo e resto dell'addome conformati come nell'
<taxonomicName id="5903F7997E89BD290DB9F84E182CA718" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:138690" class="Sordariomycetes" genus="Alfaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Alfaria simulans Emery" lsidName-HNS="Alfaria simulans Emery" order="Hypocreales" pageNumber="74" phylum="Ascomycota" rank="species" species="simulans">A. simulans</taxonomicName>
<taxonomicName id="5903F7997E89BD290DB9F84E182CA718" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:138690" class="Insecta" family="Formicidae" genus="Alfaria" kingdom="Animalia" lsidName="Alfaria simulans Emery" lsidName-HNS="Alfaria simulans Emery" order="Hymenoptera" pageNumber="74" phylum="Arthropoda" rank="species" species="simulans">A. simulans</taxonomicName>
[[ queen ]] L. 4 1 / 4 mm.
</paragraph>
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Longueur: 8 mm, 2 - 9 mm, 1. La forme est intermediaire entre les deux especes, comme la taille, mais l'epaisseur du petiole est un peu plus rapprochee de celle de la
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. Ressemble aussi a la
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., mais les cuisses et les scapes sont bruns et non rouges.
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Ent. Syst., vol. 2, p. 354 (1793), [[worker]] (
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@ -74,7 +74,7 @@ For. mais l'epinotum plus anguleux et la suture mesoepinotal plus enfoncee. Je p
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, 1925, Cat. Gen. Insc.
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@ -42,7 +42,7 @@ Emery,
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1925. Cat. Gen. Insect.
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1
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<taxonomicNameLabel id="A68E41C3D0FC8022F6901D0A5A44E961" rank="variety">n. var.</taxonomicNameLabel>
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☿, Uebergangsform zu
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<taxonomicName id="473CF8B04187AD3610DCF25A6E866B9A" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139209" class="Insecta" family="Formicidae" genus="Acantholepis" kingdom="Animalia" lsidName-HNS="Acantholepis simplex Forel" order="Hymenoptera" phylum="Arthropoda" rank="species" species="simplex">simplex</taxonomicName>
Forel, mit stumpfen, breiten Metanotumzaehnen und nur ausgebildeter, kaum gezaehnter Schuppe. Basutoland, von Herrn Rob. Wroughton erhalten. Diese Form beweist, dass
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<taxonomicName id="EDE78B49CD7F2775F88DD3C4D23BE0DD" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139209" class="Insecta" family="Formicidae" genus="Acantholepis" kingdom="Animalia" lsidName-HNS="Acantholepis simplex Forel" order="Hymenoptera" phylum="Arthropoda" rank="species" species="simplex">simplex</taxonomicName>
nur eine Subspezies von
<taxonomicName id="149FD51427A66C21101BA7F40265AA94" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139215" class="Magnoliopsida" family="Asteraceae" genus="Acantholepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName-HNS="Acantholepis capensis Mayr" order="Asterales" phylum="Tracheophyta" rank="species" species="capensis">capensis</taxonomicName>
<taxonomicName id="149FD51427A66C21101BA7F40265AA94" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139215" class="Insecta" family="Formicidae" genus="Acantholepis" kingdom="Animalia" lsidName-HNS="Acantholepis capensis Mayr" order="Hymenoptera" phylum="Arthropoda" rank="species" species="capensis">capensis</taxonomicName>
ist.
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Sous-genres
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@ -55,7 +55,7 @@ avait ainsi un aspect particulier et, a l'exception de quelques especes mal etud
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Mais ce dernier vient de modifier tout cela dans son Catalogue des
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(1925), ne reservant pour le sous-genre
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que la

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(Fig.. 5.)
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typique que par l'absence complete des tachetures en gouttelettes qui bariolent la tete, etc Sauf les mandibules, les scapes et l'extremite des segments abdominaux qui sont bruns, la Fourmi est entiere- ment d'un jaune d'ocre, terne, mais clair.
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<paragraph id="7A4CCADD6BD524A09F82AF9FFD5B0D9C" pageNumber="75">Megaloponera foetens Fab.</paragraph>
<paragraph id="7A4CCADD6BD524A09F82AF9FFD5B0D9C" pageNumber="75">
<taxonomicName id="33BC084278FD8C2D7B85C757263FE84C" authority="Fab." authorityName="Fab." class="Insecta" family="Formicidae" genus="Megaloponera" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="foetens">Megaloponera foetens Fab.</taxonomicName>
</paragraph>
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<subSubSection id="51321CC4498CA3E36D17627439098CD1" type="materials_examined">
<paragraph id="0CC2BC9A937EF19FA5500EB22081A4A8" pageNumber="75">Suedabessinien (Hg).</paragraph>

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<paragraph id="E19105732C7969C89940D356416B041C" pageNumber="9">
<taxonomicName id="5686A75D33EEBEDA9C0AD022D256EA93" ID-CoL="628QF" ID-ENA="1508249" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:146792" class="Sordariomycetes" genus="Alfaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Alfaria Emery" lsidName-HNS="Alfaria Emery" order="Hypocreales" pageNumber="9" phylum="Ascomycota" rank="genus">Alfaria</taxonomicName>
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.
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@ -43,34 +43,34 @@
<paragraph id="4C8AC62578511D176201D54187BB418D" pageNumber="9">[[ worker ]] Capo subquadrato; mandibole trigone, col margine masticatorio munito di denti piccoli e numerosi; lamine frontali distanti Tuna dall'altra, dilatate e prolungate quasi fino alla meta del capo; clipeo trasversalmente convesso, col margine anteriore distante dalle mandibole; occhi piuttosto piccoli, ma con molte faccette, situati nel mezzo dei lati del capo; antenne di 12 articoli, senza clava distinta. Torace inerme, senza suture. Segmento peduncolare dell'addome appena ingrossato nel mezzo, col contorno dorsale arcuato, col contorno ventrale incavato, attaccato al segmento seguente per quasi tutta la sua grossezza; 2. ° segmento campaniforme, separato dal 3. ° per una forte strozzatura; questo ricopre quasi tutto il resto dell'addome, e subgloboso e fortemente curvato in basso, in modo che il piano del suo margine apicale forma con quello del margine basale un angolo quasi retto; i segmenti seguenti che costituiscono la punta dell'addome sono rivolti in basso e in avanti. Tibie medie e posteriori con un solo sperone. Unghie semplici.</paragraph>
<paragraph id="019C8D70FDA9B9A96E29069C4849288B" lastPageNumber="10" pageNumber="9">
[[ queen ]] Rassomiglia alla [[ worker ]]; quella dell'
<taxonomicName id="875F4FD84FB2C3BBAAFB72D16EB2D2D3" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:138690" class="Sordariomycetes" genus="Alfaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Alfaria simulans Emery" lsidName-HNS="Alfaria simulans Emery" order="Hypocreales" pageNumber="9" phylum="Ascomycota" rank="species" species="simulans">A. simulans</taxonomicName>
<taxonomicName id="875F4FD84FB2C3BBAAFB72D16EB2D2D3" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:138690" class="Insecta" family="Formicidae" genus="Alfaria" kingdom="Animalia" lsidName="Alfaria simulans Emery" lsidName-HNS="Alfaria simulans Emery" order="Hymenoptera" pageNumber="9" phylum="Arthropoda" rank="species" species="simulans">A. simulans</taxonomicName>
ne differisce per gli occhi piu. grandi, un ocello sul vertice; le suture distinte fra i 3 segmenti del torace; lo scutello non e distinto dal resto del mesonoto; nessun vestigio di ali; peduncolo subgloboso; unghie con un piccolo dente. Nell'altra specie, vi sono tre ocelli, il torace e normale e le unghie non sono dentate.
</paragraph>
<paragraph id="79ACC1FE00A4E12291A147D999DB3455" pageNumber="10">? [[ male ]] Capo allungato, col clipeo piu sporgente che nella [[ worker ]]; mandibole trigone, allungate, un poco curvate; lamine frontali debolissime; scapo lungo circa 2 volte quanto il 1. ° articolo del flagello, piu breve del 2. °. Torace con profondi solchi parapsidiali. Peduncolo subcilindrico, un poco curvato; i due segmenti seguenti di forma ordinaria, subeguali, ricoprenti circa i 2 / 3 dell' addome, il primo di essi subcampaniforme. Tibie con un solo sperone; unghie con un forte dente. Ali con due celle cubitali.</paragraph>
<paragraph id="F1CDD0F47544B27F424DD5AB7F48A0F1" pageNumber="10">
Questo genere e molto affine a
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<taxonomicName id="AF787BB7AB288CBF5E7E030BED413CF4" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2493" class="Insecta" family="Formicidae" genus="Ectatomma" kingdom="Animalia" lsidName="Ectatomma Smith" lsidName-HNS="Ectatomma Smith" order="Hymenoptera" pageNumber="10" phylum="Arthropoda" rank="genus">Ectatomma</taxonomicName>
e particolarmente al sottogenere
<taxonomicName id="C3A934914A04C5C34BA1D4CEE68DB33B" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2497" class="Insecta" family="Formicidae" genus="Rhytidoponera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytidoponera Mayr" lsidName-HNS="Rhytidoponera Mayr" order="Hymenoptera" pageNumber="10" phylum="Arthropoda" rank="genus">Rhytidoponera</taxonomicName>
<taxonomicName id="C3A934914A04C5C34BA1D4CEE68DB33B" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2497" class="Insecta" family="Formicidae" genus="Rhytidoponera" kingdom="Animalia" lsidName="Rhytidoponera Mayr" lsidName-HNS="Rhytidoponera Mayr" order="Hymenoptera" pageNumber="10" phylum="Arthropoda" rank="genus">Rhytidoponera</taxonomicName>
: la forma, singolare del 3. ° segmento addominale che ricorda i generi
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<taxonomicName id="DA47AF7BED14C9489B048C9364C40612" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24854" class="Insecta" family="Formicidae" genus="Proceratium" kingdom="Animalia" lsidName="Proceratium Roger" lsidName-HNS="Proceratium Roger" order="Hymenoptera" pageNumber="10" phylum="Arthropoda" rank="genus">Proceratium</taxonomicName>
e
<taxonomicName id="7FA5426FD1946AF9941D23E98E358FC2" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:160189" genus="Sysphincta" lsidName="Sysphincta Mayr" lsidName-HNS="Sysphincta Mayr" pageNumber="10" rank="genus">Sysphincta</taxonomicName>
<taxonomicName id="7FA5426FD1946AF9941D23E98E358FC2" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:160189" class="Insecta" family="Formicidae" genus="Sysphincta" kingdom="Animalia" lsidName="Sysphincta Mayr" lsidName-HNS="Sysphincta Mayr" order="Hymenoptera" pageNumber="10" phylum="Arthropoda" rank="genus">Sysphincta</taxonomicName>
si trova gia distintamente accennata in varie specie di
<taxonomicName id="FECD3CA8574A2ABB1E2228F56C5787D9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2493" class="Insecta" family="Formicidae" genus="Ectatomma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectatomma Smith" lsidName-HNS="Ectatomma Smith" order="Hymenoptera" pageNumber="10" phylum="Arthropoda" rank="genus">Ectatomma</taxonomicName>
<taxonomicName id="FECD3CA8574A2ABB1E2228F56C5787D9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2493" class="Insecta" family="Formicidae" genus="Ectatomma" kingdom="Animalia" lsidName="Ectatomma Smith" lsidName-HNS="Ectatomma Smith" order="Hymenoptera" pageNumber="10" phylum="Arthropoda" rank="genus">Ectatomma</taxonomicName>
; anche la forma del peduncolo non e molto dissimile da quella dell'
<taxonomicName id="B5AAD9E10030602C85FC41806FDCA21D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:186534" class="Insecta" family="Formicidae" genus="Ectatomma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectatomma bispinosum Emery" lsidName-HNS="Ectatomma bispinosum Emery" order="Hymenoptera" pageNumber="10" phylum="Arthropoda" rank="species" species="bispinosum">E. bispinosum</taxonomicName>
<taxonomicName id="B5AAD9E10030602C85FC41806FDCA21D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:186534" class="Insecta" family="Formicidae" genus="Ectatomma" kingdom="Animalia" lsidName="Ectatomma bispinosum Emery" lsidName-HNS="Ectatomma bispinosum Emery" order="Hymenoptera" pageNumber="10" phylum="Arthropoda" rank="species" species="bispinosum">E. bispinosum</taxonomicName>
, mentre l'assenza di suture si osserva in alcune
<taxonomicName id="FC00B682DC238254DD87D9814F4B9C3E" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2495" class="Insecta" family="Formicidae" genus="Gnamptogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnamptogenys Roger" lsidName-HNS="Gnamptogenys Roger" order="Hymenoptera" pageNumber="10" phylum="Arthropoda" rank="genus">Gnamptoyenys</taxonomicName>
<taxonomicName id="FC00B682DC238254DD87D9814F4B9C3E" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2495" class="Insecta" family="Formicidae" genus="Gnamptogenys" kingdom="Animalia" lsidName="Gnamptogenys Roger" lsidName-HNS="Gnamptogenys Roger" order="Hymenoptera" pageNumber="10" phylum="Arthropoda" rank="genus">Gnamptoyenys</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="98C4A01A284620063E414CF1B6037C60" type="discussion">
<paragraph id="0CF62AA354EC619B10C1C5FA330A194A" pageNumber="10">
Il [[ male ]] non differisce da quello delle
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<taxonomicName id="92ABA537CD749DFD28DD785C8D58B5FF" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2497" class="Insecta" family="Formicidae" genus="Rhytidoponera" kingdom="Animalia" lsidName="Rhytidoponera Mayr" lsidName-HNS="Rhytidoponera Mayr" order="Hymenoptera" pageNumber="10" phylum="Arthropoda" rank="genus">Rhytidoponera</taxonomicName>
: non sono sicuro che sia stato preso con la [[ worker ]] e [[ queen ]]; lo trovai in un tubo che racchiudeva alcune [[ worker ]] di
<taxonomicName id="3FBF64604C62EA6BE955C343F6D7D866" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:146792" class="Sordariomycetes" genus="Alfaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Alfaria Emery" lsidName-HNS="Alfaria Emery" order="Hypocreales" pageNumber="10" phylum="Ascomycota" rank="genus">Alfaria</taxonomicName>
<taxonomicName id="3FBF64604C62EA6BE955C343F6D7D866" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:146792" class="Insecta" family="Formicidae" genus="Alfaria" kingdom="Animalia" lsidName="Alfaria Emery" lsidName-HNS="Alfaria Emery" order="Hymenoptera" pageNumber="10" phylum="Arthropoda" rank="genus">Alfaria</taxonomicName>
con altre formiche.
</paragraph>
</subSubSection>

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@ -43,7 +43,7 @@
F. Smith, Journ. Proc. Linn. Soc. Zool., vol. 6, p. 38 (1861), [[worker]] [
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, p. 43 (1912), [[worker]], [[male]], [[queen]].
</paragraph>
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View file

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2), [[ worker ]]. L. 7 a 9,5 mill. Peu dimorphe, a ce qu'il parait. Ressemble a la race
<taxonomicName genus="Myrmococystus" rank="variety" species="viaticus" variety="megalocola">megalocola</taxonomicName>
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Foerst. Mais il est plus petit, la tete, le thorax et l'ecaille un peu plus luisants, a sculpture plus faible et plus fine, et d'un rouge plus terne, tandis que l'abdomen est au contraire moins luisant, avec une sculpture plus forte, comme celle du reste du corps. Le noeud du pedicule est plus long, distinctement plus long que large. Du reste de la forme du
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, dont il a la tete carree, mais un peu plus convexe derriere et plutot plus longue que large, ainsi que les pattes etroites, avec une pubescence adjacente fine et line rangee soies ou piquants obliques et ecartes au bord interne. Pubescence du metanotum plutot faible. Stigmates du mesonotum tres proeminents.
</paragraph>
<paragraph pageNumber="15">[[ queen ]]. L. 7 a 8 mill. Etroite et remarquablement petite. La tete est plus etroite que chez la grande [[ worker ]] et le thorax tres etroit. Du reste comme l'ouvriere. Ailes manquent.</paragraph>
<paragraph id="9C7E0094B7CFB11D70753B1676B4150A" pageNumber="15">[[ queen ]]. L. 7 a 8 mill. Etroite et remarquablement petite. La tete est plus etroite que chez la grande [[ worker ]] et le thorax tres etroit. Du reste comme l'ouvriere. Ailes manquent.</paragraph>
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,
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(
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J'ai recu la meme race, avec les deux seules [[
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<specimenCount id="1298A8D2962C651255D7E52DCC731CD6" type="worker">worker</specimenCount>
]] de
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d'Abyssinie, de
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et de
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.
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View file

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@ -42,7 +42,7 @@ Mayr.
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Mayr., 1861, Europ. Formicid. p. 42. Emery, 1925, Gen. Insect.,
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<paragraph pageNumber="17">Buchara orient. (Tabi-dara - Zagyrdescht, 2 [[ queen ]], 6 [[ worker ]], 17. VI. 1897. Kaznakov!); Ferghana (Montagne Transalai, Bachmir, 9000 ' h., 3 [[ worker ]], 3. VII. 1895. Korzinsku!); Ussuri merid. (Sidemi, 1 [[ queen ]], 30. VII. 1897. Jankovskij!); Chine occident. (Sze- tschwan, Lunganfu, gorge de Chodzy-gou, 6000 ' h., 1 [[ worker ]], VI. 1893. Berezovsku!; Maotsch-zou - Mataigi, 1 [[ queen ]], 27. VIII. 1893. Potanin!),</paragraph>
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<taxonomicName id="C3FC4FE7606E81BF2628BD421AFCB883" ID-CoL="QDP8" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:26393" class="Collembola" family="Paronellidae" genus="Camponotus" kingdom="Animalia" lsidName="Camponotus curviscapus Emery" lsidName-HNS="Camponotus curviscapus Emery" order="Entomobryomorpha" pageNumber="71" phylum="Arthropoda" rank="species" species="curviscapus">Camponotus curviscapus</taxonomicName>
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fig. 20 — a, b.
</paragraph>

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<taxonomicName genus="Tetramonium" rank="variety" species="caespitum" variety="forte">Tetramonium caespitum L. var. forte</taxonomicName>
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<paragraph pageNumber="4">[[ worker ]]. L. 3 a 3,6 mill. Outre sa grande taille, cette variete se distingue par sa tete et son thorax grossierement et regulierement stries en long) son 2 me noeud deux fois plus large que long et, chez les exemplaires tout-a-fait typiques, par son metanotum arme de deux epines pointues, longues comme les 2 / 3 de leur intervalle.</paragraph>
<paragraph pageNumber="4">Je rapporte avec doute a cette variete des [[ queen ]] et [[ male ]] de Crimee et de Transcaucasie qui se distinguent de notre type de l'espece de l'Europe centrale et septentrionale par leur thorax plus etroit et par la tete de la [[ queen ]] un peu plus large, surtout derriere, ou elle est plus large que devant. Longueur de la [[ queen ]] 8,2, du [[ male ]] 6,0 mill.</paragraph>
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<paragraph id="1EF8E0FE6BEA358987C885165A1DD7B4" pageNumber="4">[[ worker ]]. L. 3 a 3,6 mill. Outre sa grande taille, cette variete se distingue par sa tete et son thorax grossierement et regulierement stries en long) son 2 me noeud deux fois plus large que long et, chez les exemplaires tout-a-fait typiques, par son metanotum arme de deux epines pointues, longues comme les 2 / 3 de leur intervalle.</paragraph>
<paragraph id="F7DD295803DB95CF25FEA7905D71363C" pageNumber="4">Je rapporte avec doute a cette variete des [[ queen ]] et [[ male ]] de Crimee et de Transcaucasie qui se distinguent de notre type de l'espece de l'Europe centrale et septentrionale par leur thorax plus etroit et par la tete de la [[ queen ]] un peu plus large, surtout derriere, ou elle est plus large que devant. Longueur de la [[ queen ]] 8,2, du [[ male ]] 6,0 mill.</paragraph>
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<paragraph pageNumber="4">Cette variete se trouve dans le midi de l'Europe (Camargue, Nice, Drome, Palavas pres Montpellier). Au Musee Zoolog. Ac. Imp. Sc., elle Se trouve de Crimee (Alupka, 1 [[ queen ]], 2. VI. 1899; mont Kastei, 1 [[ queen ]], 26. VI. 1900; Aluschta, 15 [[ worker ]], 13 - 15. V. 1900. N. Kuznecov!) et de Transcaucasie (les [[ queen ]] et [[ male ]] douteux): (Gouv. Kutais, Artwin, 1 [[ queen ]], 23. VI. 1898; K. Derjugin!).</paragraph>
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<paragraph id="47DA73D0EA07962A1D59670D7FFD57F1" pageNumber="4">Cette variete se trouve dans le midi de l'Europe (Camargue, Nice, Drome, Palavas pres Montpellier). Au Musee Zoolog. Ac. Imp. Sc., elle Se trouve de Crimee (Alupka, 1 [[ queen ]], 2. VI. 1899; mont Kastei, 1 [[ queen ]], 26. VI. 1900; Aluschta, 15 [[ worker ]], 13 - 15. V. 1900. N. Kuznecov!) et de Transcaucasie (les [[ queen ]] et [[ male ]] douteux): (Gouv. Kutais, Artwin, 1 [[ queen ]], 23. VI. 1898; K. Derjugin!).</paragraph>
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<paragraph id="0826593670D51E584566C08F6AC44E2F" pageNumber="4">
J'ai observe dans la Camargue ses nids qui ressemblaient a des nids de
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et qui etaient assez considerables. Dans les pays mediterraneens elle semble a premiere vue etre une espece differente des petites varietes (punicum, meridionale etc.).
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2)
<taxonomicName id="238F11734BD3FED74C6FD1B4E1C82939" authority="Santschi" authorityName="Santschi" rank="variety" variety="semispicata">var. semispicata Santschi</taxonomicName>
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:
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View file

@ -1,41 +1,42 @@
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Crematogaster oacozela Santschi = Werneri Mayr
<taxonomicName id="E89540C31BAC5C379A2957CBA138E3D0" authority="Santschi." rank="variety" variety="cacozela">var. cacozela Santschi.</taxonomicName>
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=
<taxonomicName id="0868190326B8804C88E7F1E84649BFEC" authority="Santschi." authorityName="Santschi." class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="variety" species="werneri" variety="cacozela">werneri Mayr var. cacozela Santschi.</taxonomicName>
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View file

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6.
<taxonomicName id="DD1F6B9B825D863EB472A1C73EFFE481" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139209" class="Magnoliopsida" family="Asteraceae" genus="Acantholepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acantholepis simplex Forel" lsidName-HNS="Acantholepis simplex Forel" order="Asterales" pageNumber="350" phylum="Tracheophyta" rank="species" species="simplex">Acantholepis simplex Forel</taxonomicName>
<taxonomicName id="DD1F6B9B825D863EB472A1C73EFFE481" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139209" class="Insecta" family="Formicidae" genus="Acantholepis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Acantholepis simplex Forel" lsidName-HNS="Acantholepis simplex Forel" order="Hymenoptera" pageNumber="350" phylum="Arthropoda" rank="species" species="simplex">Acantholepis simplex Forel</taxonomicName>
(Ann. soc. ent. de Belgique 6. Fevrier 1892).
</paragraph>
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@ -50,23 +50,23 @@ sehr nahe stehend, jedoch zarter, schmaeler und vor allem mit staerker ein-gesch
sind die beiden Hinterecken lappen- oder fluegelfoermig, stumpf drei-eckig vorgestreckt, eine bedeutende Concavitaet zwischen sich lassend. Bei unserer Art sind diese Hinterecken einfach recht-eckig (von der Seite gesehen) und durch eine gerade oder kaum concave Querlinie mit einander verbunden, welche zugleich die (der Laenge nach gerundete) Grenze zwischen basale und ab-schuessige Metanotumflaeche bildet. Diese Form des Metanotum erinnert sehr an diejenige gewisser
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- Arten und bildet geradezu einen Uebergang zu derselben. Die Schuppe ist viel niedriger als bei
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und etwas niedriger als bei
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, oben nur schwach ausgerandet, ohne Zaehne oder Dornen zu bilden. Die drei Stirnoccllen sind sehr deut-lich und die Schildgrube ist von der Fuehlergrube scharf getrennt, so dass diese Art doch als aechte
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zu betrachten ist.
</paragraph>
<paragraph id="6B6C588C3AD1C7D9B373E5B19E43961F" pageNumber="351">
Stark glaenzend und fast glatt; nur der Hinterleib sehr schwach genetzt. (Die
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hat einen ziemlich grob genetzten Metathorax und Mesothorax; bei
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<taxonomicName id="129990B86289890F8954045F8DE7EE2A" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139218" class="Insecta" family="Formicidae" genus="Acantholepis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aphaenogaster carbonaria Pergande" lsidName-HNS="Aphaenogaster carbonaria Pergande" order="Hymenoptera" pageNumber="351" phylum="Arthropoda" rank="species" species="carbonaria">A. carbonaria</taxonomicName>
sind Kopf und Thorax fast glanzlos, ziemlich dicht fingerhutartig punktirt.)
</paragraph>
<paragraph id="D2BDB935EA46C154A9E5035E5B7F7706" pageNumber="351">
Fast ohne Spur von anliegender Behaarung (bei
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ist der Hinterleib ziemlich stark pubescent). Abstehende Haare hellgelb, sehr zerstreut, viel zerstreuter und feiner als bei
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.
@ -91,24 +91,24 @@ zeigt der Thorax, besonders das Metanotum, eine Sculptur). Sparsam behaart. Schu
<subSubSection id="CDEEE6CF83AC5994ACB0AAD72C73520D" type="discussion">
<paragraph id="2D610FABA4D1BDD5D03E9DD66D7A0FB6" pageNumber="351">
In der citirten Arbeit (Ann. soc. out. Belg.) habe ich die Arten der Gattung
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<taxonomicName id="66B0F236D123632E9DFB915E9AF91FD8" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:146781" class="Insecta" family="Formicidae" genus="Acantholepis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Acantholepis Mayr" lsidName-HNS="Acantholepis Mayr" order="Hymenoptera" pageNumber="351" phylum="Arthropoda" rank="genus">Acantholepis</taxonomicName>
synoptisch zusammengestellt und dabei zwei neue Varietaeten der
<taxonomicName id="F7FD71E2D07BEE85C3A8BD73A561C78D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:178143" class="Magnoliopsida" family="Asteraceae" genus="Acantholepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="A. Frauenfeldi" lsidName-HNS="A. Frauenfeldi" order="Asterales" pageNumber="351" phylum="Tracheophyta" rank="species" species="frauenfeldi">A. Frauenfeldi</taxonomicName>
<taxonomicName id="F7FD71E2D07BEE85C3A8BD73A561C78D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:178143" class="Insecta" family="Formicidae" genus="Acantholepis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. Frauenfeldi" lsidName-HNS="A. Frauenfeldi" order="Hymenoptera" pageNumber="351" phylum="Arthropoda" rank="species" species="frauenfeldi">A. Frauenfeldi</taxonomicName>
aufgestellt. Die indische Varietaet
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<taxonomicName id="DAF784874455B92A41A52691B9E6F710" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139308" class="Insecta" family="Formicidae" genus="Acantholepis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="sericea" lsidName-HNS="sericea" order="Hymenoptera" pageNumber="351" phylum="Arthropoda" rank="variety" species="frauenfeldi" variety="sericea">sericea</taxonomicName>
ist sehr schlank und in Folge der genetzt-gerunzelten, ziemlich dichten Sculptur ueberall seidenglaenzend. Sie ist dunkelbraun, mit etwas roethlichem mittlerem Theil des Metanotum und hell-braeunlich-gelblichen Fuehlern, Schienen und Tarsen; nur der etwas verdickte End-theil des Fuehlerschaftes ist braeunlich. Der Kopf ist etwas abgeflacht. Die anliegende und abstehende Behaarung ist sehr spaerlich.
</paragraph>
</subSubSection>
<subSubSection id="C9333F0B1DC5B82B0EC36602FDD3F0FC" type="nomenclature">
<paragraph id="A1F2ADFE5F8765EB530DD8FB8902B8C4" pageNumber="351">
Als
<taxonomicName id="0AF259ACBE0CC7CF986B8C85D65AE9C8" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139246" class="Magnoliopsida" family="Asteraceae" genus="Acantholepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acantholepis frauenfeldi var. pubescens Forel" lsidName-HNS="Acantholepis frauenfeldi var. pubescens Forel" order="Asterales" pageNumber="351" phylum="Tracheophyta" rank="variety" species="frauenfeldi" variety="pubescens">Var. pubescens</taxonomicName>
<taxonomicName id="0AF259ACBE0CC7CF986B8C85D65AE9C8" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139246" class="Insecta" family="Formicidae" genus="Acantholepis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Acantholepis frauenfeldi var. pubescens Forel" lsidName-HNS="Acantholepis frauenfeldi var. pubescens Forel" order="Hymenoptera" pageNumber="351" phylum="Arthropoda" rank="variety" species="frauenfeldi" variety="pubescens">Var. pubescens</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="4477BE0A21651AC81FB8493C4325FE1A" type="description">
<paragraph id="E510FE55B3FBC8CB922D4AE12EA3242E" lastPageNumber="352" pageNumber="351">
habe ich eine sehr grosse, 3,2 - 4 mm. lange, schwarze Varietaet bezeichnet, die ich selbst in Suedtunesien s. Z. bei Gabes und Oued Mela gesammelt und mit
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verwechselt hatte. Diese Varietaet ist sehr characteristisch und duerfte wohl als Rasse taxirt werden. Sie ist robuster als die Stammart; der Kopf ist breit, hinten noch breiter und in der Mitte deutlich ausgebuchtet. Das Metanotum ist hinten etwas breiter mit ziemlich stumpfen, an der Basis breiten Zaehnen oder Dornen. Die Schuppe hat oben zwei Zaehne oder Dornen, die einander genaehert sind, so dass die Raender der Schuppe die Basis der Dornen seitlich deutlich ueberragen; die Schuppe ist ueberhaupt unten breiter als bei anderen Formen. Der Hinterleib ist ziemlich gross und weich. Glaenzend, schwach genetzt zerstreut abstehend, aber ziemlich reichlich und ziemlich grob ueberall anliegend behaart. Die Schienen und Schaften sind reichlich, ziemlich kurz, schief abstehend behaart. Schwarz; Mitte des Mesonotum und Mandibeln roethlich. Schienen, Tarsen und die basalen 2 / 3 der Fuehlerschaften roethlichgelb. Rest der Beine und der Fuehler braeunlich. Das [[ queen ]] ist 6 - 6,5 mm. lang, unterscheidet sich im Uebrigen wenig von demjenigen der Stammart.
</paragraph>
</subSubSection>

View file

@ -1,42 +1,42 @@
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47.
<taxonomicName id="2EEAF018AAE3222B73BA3478D15497EA" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139207" class="Magnoliopsida" family="Asteraceae" genus="Acantholepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acantholepis lunaris Emery" lsidName-HNS="Acantholepis lunaris Emery" order="Asterales" pageNumber="12" phylum="Tracheophyta" rank="species" species="lunaris">Acantholepis lunaris</taxonomicName>
<taxonomicName id="2EEAF018AAE3222B73BA3478D15497EA" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139207" class="Insecta" family="Formicidae" genus="Acantholepis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Acantholepis lunaris Emery" lsidName-HNS="Acantholepis lunaris Emery" order="Hymenoptera" pageNumber="12" phylum="Arthropoda" rank="species" species="lunaris" status="sp. nov.">Acantholepis lunaris</taxonomicName>
,
<taxonomicNameLabel id="165BFB5305A2FF99803A859D3B83EB5D" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
@ -50,9 +50,9 @@
<subSubSection id="0343DF9BA1317C672FD8CF846A9D34CB" type="discussion">
<paragraph id="E1E8AB7B46AFD3C251E7AD9F8C625320" pageNumber="12">
Elle a la forme trapue de
<taxonomicName id="17CE205B40EA76A125B4DEAD06FF36D0" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139215" class="Magnoliopsida" family="Asteraceae" genus="Acantholepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acantholepis capensis Mayr" lsidName-HNS="Acantholepis capensis Mayr" order="Asterales" pageNumber="12" phylum="Tracheophyta" rank="species" species="capensis">A. capensis Mayr</taxonomicName>
<taxonomicName id="17CE205B40EA76A125B4DEAD06FF36D0" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139215" class="Insecta" family="Formicidae" genus="Acantholepis" kingdom="Animalia" lsidName="Acantholepis capensis Mayr" lsidName-HNS="Acantholepis capensis Mayr" order="Hymenoptera" pageNumber="12" phylum="Arthropoda" rank="species" species="capensis">A. capensis Mayr</taxonomicName>
, dont elle differe surtout par la construction du metanotum: ce segment (fig. 12) se prolonge de chaque cote en une sorte de corne epaisse et pointue, courbee en arriere, dont le plan superieur (qui forme sur le profil une ligne droite) est continu avec la face basale; vu par dessus, le metanotum a la forme d'un croissant dont le creux et les cornes sont tournes en arriere. L'ecaille est haute et echancree en arc en dessus, terminee par deux dents ou epines tres aigues et un peu repliees en arriere. Sculpture a peu pres comme chez
<taxonomicName id="7A7A3473F1D7C244B640B9F3AA459FF7" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139215" class="Magnoliopsida" family="Asteraceae" genus="Acantholepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acantholepis capensis Mayr" lsidName-HNS="Acantholepis capensis Mayr" order="Asterales" pageNumber="12" phylum="Tracheophyta" rank="species" species="capensis">A. capensis</taxonomicName>
<taxonomicName id="7A7A3473F1D7C244B640B9F3AA459FF7" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139215" class="Insecta" family="Formicidae" genus="Acantholepis" kingdom="Animalia" lsidName="Acantholepis capensis Mayr" lsidName-HNS="Acantholepis capensis Mayr" order="Hymenoptera" pageNumber="12" phylum="Arthropoda" rank="species" species="capensis">A. capensis</taxonomicName>
, mais le tegument est un peu plus lisse; abdomen tres luisant; pubescence excessivement tenue et rare, poils dresses gros, obtus, peu nombreux.
</paragraph>
</subSubSection>

View file

@ -1,40 +1,40 @@
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,
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@ -49,9 +49,9 @@
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Cette espece se rapproche d'
<taxonomicName id="F99B96E7D0E0AD06B627331AD4298422" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139225" class="Magnoliopsida" family="Asteraceae" genus="Acantholepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acantholepis depressa Santschi" lsidName-HNS="Acantholepis depressa Santschi" order="Asterales" pageNumber="262" phylum="Tracheophyta" rank="species" species="depressa">A. depressa Santschi</taxonomicName>
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et d'
<taxonomicName id="1B61B7899BF8F02F80F45590305D38A0" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:179286" class="Magnoliopsida" family="Asteraceae" genus="Acantholepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acantholepis deplanata Stitz" lsidName-HNS="Acantholepis deplanata Stitz" order="Asterales" pageNumber="262" phylum="Tracheophyta" rank="species" species="deplanata">A. deplanata</taxonomicName>
<taxonomicName id="1B61B7899BF8F02F80F45590305D38A0" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:179286" class="Insecta" family="Formicidae" genus="Acantholepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acantholepis deplanata Stitz" lsidName-HNS="Acantholepis deplanata Stitz" order="Hymenoptera" pageNumber="262" phylum="Arthropoda" rank="species" species="deplanata">A. deplanata</taxonomicName>
Stitz par son pronotum concave, mais s'en distingue par son mesonotum allonge et son ecaille.
</paragraph>
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View file

@ -1,35 +1,35 @@
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@ -43,7 +43,7 @@ Santschi.
Ann. Soc. ent. France, vol. 79, p. 351 (1010), [[worker]], [[queen]] (
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.). - Emery, in Wytsman, Genera Insectorum, Formicidae, subfam.
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, faso. CXII. p. 108 (1911).
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Cremastogaster taediosa,
<taxonomicName id="7AA4FB3E7F726C9F70AD8433F7211B11" class="Insecta" family="Formicidae" genus="Cremastogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="taediosa" status="sp. nov.">Cremastogaster taediosa</taxonomicName>
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<taxonomicName genus="Tetramonium" rank="subSpecies" species="caespitum" subSpecies="semilaeve">Tetramonium caespitum L. r. semilaeve Andre</taxonomicName>
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<taxonomicName id="688872BB87C32107E79B690107AEB310" authority="Andre" authorityName="Andre" class="Insecta" family="Formicidae" genus="Tetramonium" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subSpecies" species="caespitum" subSpecies="semilaeve">Tetramonium caespitum L. r. semilaeve Andre</taxonomicName>
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<paragraph pageNumber="4">Transcaucasie: vallee d'Araxes ([[ worker ]], [[ queen ]] Reitter!) [variete claire]; gouv. de Baku, defile de Bum (2 [[ worker ]], 1892, Schelkovnikov!); Gouv. Elizabethpol (Geok-tapa, steppe, 1 [[ queen ]], 2. VII. 1901; plateau de Sarudza, 400 m. h., steppe, 1 [[ queen ]], 2. VII. 01, R. Schmidt!) [v. noire].</paragraph>
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<paragraph id="69D5E7B73B3B47F6046887F68089F0E1" pageNumber="4">Transcaucasie: vallee d'Araxes ([[ worker ]], [[ queen ]] Reitter!) [variete claire]; gouv. de Baku, defile de Bum (2 [[ worker ]], 1892, Schelkovnikov!); Gouv. Elizabethpol (Geok-tapa, steppe, 1 [[ queen ]], 2. VII. 1901; plateau de Sarudza, 400 m. h., steppe, 1 [[ queen ]], 2. VII. 01, R. Schmidt!) [v. noire].</paragraph>
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<taxonomicName id="1D211EDBB105780EDD70D02E249CD3AF" LSID-HNS="urn:lsid:biosci.ohio-state.edu:139261" class="Magnoliopsida" family="Asteraceae" genus="Acantholepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName-HNS="Acantholepis imperfecta Santschi" order="Asterales" pageNumber="257" phylum="Tracheophyta" rank="species" species="imperfecta" subGenus="Pseudacantholepis">A. (Ps.) imperfecta</taxonomicName>
<taxonomicName id="1D211EDBB105780EDD70D02E249CD3AF" LSID-HNS="urn:lsid:biosci.ohio-state.edu:139261" class="Insecta" family="Formicidae" genus="Acantholepis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName-HNS="Acantholepis imperfecta Santschi" order="Hymenoptera" pageNumber="257" phylum="Arthropoda" rank="species" species="imperfecta" subGenus="Pseudacantholepis">A. (Ps.) imperfecta</taxonomicName>
SANT.
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2)
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nov.
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[[ worker ]]. L. 6 a 8 mill. Tres semblable au
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, mais avec la couleur et la sculpture de l´
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. Aden (Dr. Bis!), ma coll.
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<taxonomicName id="6B90BE1B003D377BB140657EFD82D550" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:230406" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" lsidName="Camponotus barbarossa arminius Forel" lsidName-HNS="Camponotus barbarossa arminius Forel" order="Hymenoptera" pageNumber="21" phylum="Arthropoda" rank="subSpecies" species="arminius" subGenus="Myrmamblys" subSpecies="barbarossa">Camponotus (Myrmamblys) arminius st. barbarossa Emery</taxonomicName>
, Cat. Gen. Insect.
<taxonomicName id="E2EF3EACBCADDD175CC41FC1CE05CB06" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2254" lsidName-HNS="Formicinae Latreille" pageNumber="21" rank="subFamily">Formicinae</taxonomicName>
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, p. 142.
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@ -45,7 +45,7 @@ Mus. Ludov. Ulr., p. 412 (1764), [[male]] (
). - Fred; Smith, Cat. Hymenopt. Brit. Mus., vol. 6, p. 112 (1858),.[[worker]] [
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, fase. CII, p. 10 (1910), [[queen]], [[worker]],[[male]].
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Monomorium (
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) modestum Santschi.
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.
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View file

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<taxonomicName id="73F8BF85F0C1CD245B7B9F06EFB90928" LSID-HNS="urn:lsid:biosci.ohio-state.edu:146781" class="Magnoliopsida" family="Asteraceae" genus="Acantholepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName-HNS="Acantholepis Mayr" order="Asterales" pageNumber="257" phylum="Tracheophyta" rank="genus">Acantholepis</taxonomicName>
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<taxonomicNameLabel id="0E671721D09F8CD59A14CC8C917BB999" rank="species">n.sp.</taxonomicNameLabel>
?
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@ -42,9 +42,9 @@
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Une ouvriere et une [[queen]] ailee, etiquetees &quot; Nimba &quot; sont voisins de la forme equatoriale
<taxonomicName id="BD1EB603C864B8135A2E4C2D6B0A77C4" LSID-HNS="urn:lsid:biosci.ohio-state.edu:139218" class="Magnoliopsida" family="Asteraceae" genus="Acantholepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName-HNS="Acantholepis carbonaria Emery" order="Asterales" pageNumber="257" phylum="Tracheophyta" rank="species" species="carbonaria">A. carbonaria</taxonomicName>
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Em., mais specifiquement distincts. Les
<taxonomicName id="5F5644A9C2098D12D8A97E402D494670" LSID-HNS="urn:lsid:biosci.ohio-state.edu:146781" class="Magnoliopsida" family="Asteraceae" genus="Acantholepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName-HNS="Acantholepis Mayr" order="Asterales" pageNumber="257" phylum="Tracheophyta" rank="genus">Acantholepis</taxonomicName>
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etant tres variables individuellement, il vaut mieux attendre davantage d'exemplaires pour les decrire.
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@ -42,16 +42,16 @@
<paragraph id="D764380D427C8CFA05323A21A30875EB" pageNumber="13">— Les soies noires sont encore plus denses bien qu'un peu plus fines et plus pointues que chez le type des Monts Rouwenzori. Congo belge: Ubanghi, Banzyville (R. P. Augustin).</paragraph>
<paragraph id="0FD7CB13107CA9436D29EABF02B8CE7A" pageNumber="13">
Une variete du Benguela (Crughet) les a, au contraire, tres clairsemees et plus courtes: c'est la var.
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<taxonomicNameLabel id="31C7B98684AA51D4D3D9DC76EC38A339" rank="variety">n. var.</taxonomicNameLabel>
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Une autre variete nouvelle, que je rattache a
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., a le gastre assez luisant, bien moins toutefois que chez
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Sm.; je l'ai recue du Congo francais, c'est la variete
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<taxonomicNameLabel id="05F2F4ED614199917A1357763AC280D8" rank="variety">n. var.</taxonomicNameLabel>
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1. Subgenus
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, n. subg.
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Camponotus {Myrmotrema) Grandidieri For.
<taxonomicName id="1762809826699A55EBA7C68E3FF3F5EB" authority="Em." rank="subSpecies" subSpecies="mendax">stirps mendax Em.</taxonomicName>
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@ -48,7 +47,7 @@ M. Emery m'ecrit que l'ouvriere minor sur laquelle il a etabli cette sous-espece
Je possede deux ouvrieres (media et minor) du Natal, dont la tete est plus longue et plus triangulaire que chez
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, la couleur comme chez la
<taxonomicName id="6ABB15EB7E7D8764E5E91AA3A7E9A791" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="variety" species="grandidieri" variety="comorensis">var. comorensis</taxonomicName>
<taxonomicName id="6ABB15EB7E7D8764E5E91AA3A7E9A791" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="variety" species="grandidieri" variety="comorensis">var. comorensis</taxonomicName>
. Mais, alors que chez la petite [[ worker ]] il n'y a pas de ligne glabre, celle-ci est passablement distincte chez l'ouvriere moyenne. Il parait donc assez probable qu'elle existe chez la [[ worker ]] major, encore inconnue.
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<treatment id="C8E6029EF0D858CBA4B07674E530DD5B" ID-DOI="http://doi.org/10.5281/zenodo.6289347" ID-GBIF-Taxon="100128783" ID-Zenodo-Dep="6289347" LSID="urn:lsid:plazi:treatment:C8E6029EF0D858CBA4B07674E530DD5B" httpUri="http://treatment.plazi.org/id/C8E6029EF0D858CBA4B07674E530DD5B" lastPageNumber="101" pageNumber="100">
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<paragraph id="BE3DCB9C6399DE2D4F235EEFE874C169" pageNumber="100">
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<paragraph id="76FF4CDEFCC47CD16DF4A0612968829A" pageNumber="100">
[[ worker ]] minor. Longueur: environ 8 mm. Voisin du
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.
</paragraph>
<paragraph id="C93C5A7602EA46CBAB46F443EAB923E3" pageNumber="100">Mandibules assez fortement courbees, courtes, lisses et luisantes, avec des points espaces, armees de six dents. Tete sans les mandibules 1 1 / 2 fois plus longue que large, a cotes tres peu convexes, presque paralleles; elle est presque aussi large devant que derriere. Derriere les yeux, plutot petits mais fort convexes, situes au tiers posterieur, les bords de la tete forment une simple convexite posterieure, sans trace d'angles, et vont directement rejoindre le bord occipital articulaire. Epistome, carene, avec un court lobe anterieur. Les longues aretes frontales sont aussi rapprochees l'une de l'autre devant que derriere, mais eloignees a leur milieu par une concavite. Les tres longs scapes depassent le bord occipital de 2 1 / 2 fois leur longueur. Promesonotum fort convexe, en dos de Chameau; face basale de l'epinotum bien plus longue que la face declive, aussi longue que le mesonotum, mais presque pas convexe. Le pronotum est aussi long ' que le mesonotum. L'ecaille du petiole est tres longue (epaisse), plus longue vers sa base que sa hauteur et que sa largeur, un peu inclinee en avant; son sommet est arrondi. Pattes tres longues; tibias sans piquants.</paragraph>
@ -54,15 +54,15 @@
<subSubSection id="465C48149F69DD9517874BC7B1C5EA89" type="discussion">
<paragraph id="6A3DFB9A30C8392B2B8AD18A0442BD15" pageNumber="101">
Dans mes sous genres des
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decrits en 1914, dans la Revue suisse de Zoologie, j'ai eu le grand tort de donner le
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<taxonomicName id="7150437C28B930B1BDCDEC3D6990F47E" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:26445" class="Collembola" family="Paronellidae" genus="Camponotus" kingdom="Animalia" lsidName-HNS="Camponotus ephippium (Smith)" order="Entomobryomorpha" pageNumber="101" phylum="Arthropoda" rank="species" species="ephippium">C. ephippium</taxonomicName>
comme type du sous-genre
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, sans connaitre sa [[ worker ]] major, qui s'est revelee des lors etre une
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. Je fais peccavi a cet egard et je reconnais en meme temps, avec M. Emery, le
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<taxonomicName id="5637676A7ED685A4A8BE3B786398B6FF" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:26311" class="Collembola" family="Paronellidae" genus="Camponotus" kingdom="Animalia" lsidName-HNS="Camponotus capito Mayr" order="Entomobryomorpha" pageNumber="101" phylum="Arthropoda" rank="species" species="capito">C. capito</taxonomicName>
comme type des
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. Mais est-ce la une raison suffisante pour supprimer simplement le sous-genre
@ -70,9 +70,9 @@ comme type des
relativement a celles des especes australiennes dont le vertex n'est nullement renfle chez la grande [[ worker ]]? Je me permets d'en clouler et de maintenir mon sous-genre
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en lui donnant comme type le
<taxonomicName id="620F08E3F40D9EB58E38FF3EBDDE2A32" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:26733" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus gambeyi Emery" order="Hymenoptera" pageNumber="101" phylum="Arthropoda" rank="species" species="gambeyi">C. gambeyi Em</taxonomicName>
<taxonomicName id="620F08E3F40D9EB58E38FF3EBDDE2A32" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:26733" class="Collembola" family="Paronellidae" genus="Camponotus" kingdom="Animalia" lsidName-HNS="Camponotus gambeyi Emery" order="Entomobryomorpha" pageNumber="101" phylum="Arthropoda" rank="species" species="gambeyi">C. gambeyi Em</taxonomicName>
. dont la grande [[ worker ]] est connue. Quant au
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<taxonomicName id="FBD9D09ECCAEEFA76BE745AB746E76A3" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:26751" class="Collembola" family="Paronellidae" genus="Camponotus" kingdom="Animalia" lsidName-HNS="Camponotus gouldianus Forel" order="Entomobryomorpha" pageNumber="101" phylum="Arthropoda" rank="species" species="gouldianus">C. gouldianus</taxonomicName>
, attendons pour nous prononcer que sa grande [[ worker ]] soit decouverte.
</paragraph>
<paragraph id="5F1F8E7A96014E13C7091CA7F1E0D2EF" pageNumber="101">En terminant ce petit article je prends conge de mes chers collegues en myrmecologie, apres avoir publie mon premier travail en 1869, donc il y a 53 ans; mais il y a bien 67 ans que j'etudie les Fourmis. Aujourd'hui mes yeux ne me le permettent plus. Neanmoins, les cinq volumes de mon ouvrage biologique sur à « Le Monde special des Fourmis à » sont tous ecrits et n'attendent que leur impression.</paragraph>

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, 1925, Cat. Gon. Ins.
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, p. 142.
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<taxonomicName id="4CB9B8947460B4EDA34EF88164A51CEA" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139220" class="Magnoliopsida" family="Asteraceae" genus="Acantholepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acantholepis crinita Mayr" lsidName-HNS="Acantholepis crinita Mayr" order="Asterales" pageNumber="149" phylum="Tracheophyta" rank="species" species="crinita">A. crinita</taxonomicName>
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Arbeiter. Laenge 2.4 Mm. Schwarz, die Endhaelfte der Mandibeln, die Fuehler, ausser dem letzten Viertel des Schaftes und der Endhaelfte der Geissei und die Tarsen mehr oder weniger hell rothbraun oder braungelb. Der ganze Koerper ist mit sehr langen, weisslichen, nicht dicken ,, abstehenden Borstenhaaren reichlich besetzt, die Fuehler noch reichlicher, aber kurz und schief abstehend behaart, die Schenkel und Tibien mit mittellangen abstehenden Haaren reichlich besetzt. Eine anliegende Pubescenz ist nicht zu erkennen. Kopf, Pronotum und Mesonotum glaenzend und nahezu glatt, theilweise naemlich sehr seicht und fein lederartig gerunzelt, das Postscutellum und die Seiten des Mesothorax dort, wo der Thorax eingeschnuert ist, grob runzelig-laengsgestreift; das ganze Metanotum grob gerunzelt und ziemlich glanzlos; Petiolus und Abdomen glatt und glaenzend. Der Koerper ist staemmiger als bei
<taxonomicName id="099E166760368D175615104F100ABA8D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139215" class="Magnoliopsida" family="Asteraceae" genus="Acantholepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acantholepis capensis Mayr" lsidName-HNS="Acantholepis capensis Mayr" order="Asterales" pageNumber="149" phylum="Tracheophyta" rank="species" species="capensis">A. capensis Mayr</taxonomicName>
<taxonomicName id="099E166760368D175615104F100ABA8D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139215" class="Insecta" family="Formicidae" genus="Acantholepis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Acantholepis capensis Mayr" lsidName-HNS="Acantholepis capensis Mayr" order="Hymenoptera" pageNumber="149" phylum="Arthropoda" rank="species" species="capensis">A. capensis Mayr</taxonomicName>
. Der ovale, hinten massig ausgerandete Kopf hat die Form wie bei
<taxonomicName id="4241B7AFBF299A60F969DEB25D5E413F" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139215" class="Magnoliopsida" family="Asteraceae" genus="Acantholepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acantholepis capensis Mayr" lsidName-HNS="Acantholepis capensis Mayr" order="Asterales" pageNumber="149" phylum="Tracheophyta" rank="species" species="capensis">A. capensis</taxonomicName>
<taxonomicName id="4241B7AFBF299A60F969DEB25D5E413F" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139215" class="Insecta" family="Formicidae" genus="Acantholepis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Acantholepis capensis Mayr" lsidName-HNS="Acantholepis capensis Mayr" order="Hymenoptera" pageNumber="149" phylum="Arthropoda" rank="species" species="capensis">A. capensis</taxonomicName>
. Der Fuehlerschaft ueberragt den Hinterrand des Kopfes etwa um 1 / 4 seiner Laenge, zweites Geisseiglied kaum 1 1 / 2 so lang als dick, die anderen Glieder laenger. Das Pronotum ist so wie bei
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<taxonomicName id="52396A942878581F3982AE1DA4E8C5A0" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139215" class="Insecta" family="Formicidae" genus="Acantholepis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Acantholepis capensis Mayr" lsidName-HNS="Acantholepis capensis Mayr" order="Hymenoptera" pageNumber="149" phylum="Arthropoda" rank="species" species="capensis">A. capensis</taxonomicName>
nicht stark gewoelbt. Die Einschnuerung des Thorax ist wohl
<pageStartToken id="269FFA3EA9661AE9AA91C7E489730DE6" pageNumber="150">deutlich</pageStartToken>
aber kurz, staerker als bei
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<taxonomicName id="557B14568D012095E3764E61ED930D4F" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139215" class="Insecta" family="Formicidae" genus="Acantholepis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Acantholepis capensis Mayr" lsidName-HNS="Acantholepis capensis Mayr" order="Hymenoptera" pageNumber="150" phylum="Arthropoda" rank="species" species="capensis">A. capensis</taxonomicName>
. Das Metanotum mit zwei schief nach aussen, oben und hinten gerichteten, am Ende abgerundeten Kegeln. Die Schuppe des Petiolus oben mit zwei spitzigen Zaehnen, zwischen diesen ist der obere Rand der Schuppe bogig ausgerandet.
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Ph. Anastasii
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<taxonomicNameLabel id="4777ACDBFDCDC34295DEC5666E346F4F" rank="species">n. sp.</taxonomicNameLabel>
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@ -48,11 +48,11 @@ Ph. Anastasii
<subSubSection id="C1A6CA90BD90C0AAC00CC2066C55711C" type="discussion">
<paragraph id="A30E58BEB4BC1B3D30ACFD266DF5A02C" pageNumber="45">
Rassomiglia alla Ph. Goeldii For, da cui il [[ soldier ]] differisce per l'addome in parte opaco e per la forma del 2. ° segmento del peduncolo. D'altronde queste due specie sono fra loro molto affini, e forse converrebbe di considerarle piuttosto corne sottospecie di una medesima specie; sono pure affini alle
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e
<taxonomicName id="A627302BD5E671424D8DC2EE2ECF2864" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33747" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole floridana Emery" lsidName-HNS="Pheidole floridana Emery" order="Hymenoptera" pageNumber="45" phylum="Arthropoda" rank="species" species="floridana">floridana</taxonomicName>
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Emery. Quest' ultima, che avevo descritta come sottospecie di
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<taxonomicName id="A116018C7D1E9FFC4C492E14722AB52D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33744" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" lsidName="Pheidole flavens Roger" lsidName-HNS="Pheidole flavens Roger" order="Hymenoptera" pageNumber="45" phylum="Arthropoda" rank="species" species="flavens">flavens Rog</taxonomicName>
., mi sembra, dopo di avere ristudiato l'intero gruppo, dover formare specie a se.
</paragraph>
</subSubSection>

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[[ worker ]]. Longueur: 9 mm, 3 - 11 mm. Un peu voisin a la fois de
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et de
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, de ce dernier par sa stature ramassee; voisin aussi du
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.
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<paragraph id="7CE0603D8D54E39EC3522C49A6DDC5DC" pageNumber="99">
Mandibules assez fortement courbees vers l'extremite, armees d'environ 7 dents, subopaques, avec de nombreux gros points peu espaces et densement reticulees-ridees dans leur entredeux. Tete presque rectangulaire; sans les mandibules, elle est un peu plus longue que large, tres peu elargie en arriere, a cotes tres peu convexes, fortement et largement echancres au Lord posterieur. Lobe anterieur de l'epistome plutot court, rectangulaire; la carene mediane ne va pas jusqu'au lobe. Arenes frontales assez divergentes. Yeux situes au tiers posterieur de la tete. Le scape ne depasse le bord posterieur de la tete de guere plus que de son epaisseur. Le thorax est mediocrement convexe; la face declive de l'epinotum est plus courte que sa face basale. L'ecaille est bien plus haute et plus mince que chez
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, tranchante a son sommet; les pattes sont plus courtes.
</paragraph>
<paragraph id="FE31A227DAA566E3CAE0776D22E8A8A8" pageNumber="99">Tout le corps est tres finement et plus ou moins densement reticule-ride et, selon cette densite variable, luisant sur l'abdomen, subopaque sur le thorax et en partie l'un ou l'autre sur la tete. Les membres sont plutot subopaques. Les scapes et les joues ont de nombreux gros points enfonces qui sont epars ailleurs. Pilosite dressee et pubescence extremement eparses sur le corps, nulle sur les membres. Une rangee de petits piquants sur la face inferieure des tibias.</paragraph>

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<paragraph id="085FE4AD421F396C6DE34B4D2100163A" pageNumber="261">Var. semispicata SANT.</paragraph>
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<taxonomicName id="8FF189ACF68B44F7FAC34101853CC0B2" authority="SANT." authorityName="Sant." class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" species="maculatus" variety="semispicata">Var. semispicata SANT.</taxonomicName>
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<paragraph id="4BF0076738A7CA73163B8DA9072C5A1B" pageNumber="261">Mont To, 1.600 m., une ouvriere.</paragraph>

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<taxonomicName id="1A4D799FEDEB94D9B4D47DE1053F0CEA" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:224684" class="Collembola" family="Paronellidae" genus="Camponotus" kingdom="Animalia" lsidName-HNS="Camponotus (Myrmothrix) abdominalis r. stercorarius var. wagneri Forel" order="Entomobryomorpha" pageNumber="99" phylum="Arthropoda" rank="variety" species="abdominalis" subGenus="Myrmothrix" subSpecies="stercorarius" variety="wagneri">Camponotus (Myrmothrix) abdominalis F. r. stercorarius For. v. wagneri</taxonomicName>
<taxonomicNameLabel id="71C3386CDC7459EBF419E6F4406345F4" rank="variety">n. v.</taxonomicNameLabel>
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Plus elance que le
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typique; pattes et antennes plus longues. Le scape de la grande [[ worker ]] depasse le bord posterieur de la tete de deux fois son epaisseur (d'une fois seulement chez le type de la race). La tete est aussi moins foncee, roussatre, d'un brun presque noir chez le
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<taxonomicName id="581E073890F7E011486E84B5004FA613" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:193729" class="Collembola" family="Paronellidae" genus="Camponotus" kingdom="Animalia" lsidName-HNS="Camponotus stercorarius Forel" order="Entomobryomorpha" pageNumber="100" phylum="Arthropoda" rank="species" species="stercorarius">stercorarius</taxonomicName>
typique. Du reste de meme taille, mais tres nettement plus svelte. Perou.
</paragraph>
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<taxonomicName id="C9FD79DD24A55BDF05AF7C74E8943A67" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:178143" class="Insecta" family="Formicidae" genus="Acantholepis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Acantholepis frauenfeldi Mayr" lsidName-HNS="Acantholepis frauenfeldi Mayr" order="Hymenoptera" pageNumber="18" phylum="Arthropoda" rank="species" species="frauenfeldi">Acantholepis Frauenfeldi</taxonomicName>
Mayr.
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View file

@ -1,35 +1,35 @@
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@ -44,7 +44,7 @@ Paltothyreus
Supplem. Ent. Syst., p. 280 (1798), [[worker]] (
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). - Emery, in P. Wytsman, Genera Insectorum, Formicidae, subfam.
<taxonomicName id="F7B6CCF53044AE97F430673B2797BAA3" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2261" lsidName="Ponerinae Lepeletier" lsidName-HNS="Ponerinae Lepeletier" pageNumber="47" rank="subFamily" subFamily="Ponerinae">Ponerinae</taxonomicName>
<taxonomicName id="F7B6CCF53044AE97F430673B2797BAA3" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2261" class="Insecta" family="Formicidae" kingdom="Animalia" lsidName="Ponerinae Lepeletier" lsidName-HNS="Ponerinae Lepeletier" order="Hymenoptera" pageNumber="47" phylum="Arthropoda" rank="subFamily" subFamily="Ponerinae">Ponerinae</taxonomicName>
, p. 62 (1911).
</paragraph>
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