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<document id="55123F27B7399E3EA16475D6DD53CFE4" ID-CLB-Dataset="307822" ID-DOI="10.11646/zootaxa.5566.3.4" ID-GBIF-Dataset="3b0ded88-4de2-47ab-a799-be403aa61373" ID-ISSN="1175-5326" ID-Zenodo-Dep="14703382" ID-ZooBank="E7984AE6-EE79-49A1-BA1D-E138FE7E510B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1737370770158" checkinUser="plazi" docAuthor="Patoka, Jiří, Akmal, Surya Gentha, Bláha, Martin &amp; Kouba, Antonín" docDate="2025" docId="03DB87943B48037D419A44C726243339" docLanguage="en" docName="zootaxa.5566.3.4.pdf" docOrigin="Zootaxa 5566 (3)" docSource="https://doi.org/10.11646/zootaxa.5566.3.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cherax pulverulentus Patoka, Akmal, Bláha &amp; Kouba, 2025, sp. nov." docType="treatment" docVersion="5" lastPageNumber="532" masterDocId="FFE2FFEC3B4A0377410D420425473530" masterDocTitle="Cherax pulverulentus, a new freshwater crayfish (Decapoda: Parastacidae) from Southwest Papua Province, Indonesia" masterLastPageNumber="534" masterPageNumber="522" pageNumber="524" updateTime="1737637449387" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="003849EAE3ABB9F5A8355CAA6D3BD897">Cherax pulverulentus, a new freshwater crayfish (Decapoda: Parastacidae) from Southwest Papua Province, Indonesia</mods:title>
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<mods:namePart id="BF9BC705C380223C8CAE5E699BF278ED">Patoka, Jiří</mods:namePart>
<mods:affiliation id="E2D2260D6DEC0637B5B02D6F972BF690">Department of Zoology and Fisheries, Faculty of Agrobiology, Food and Natural Resources, Czech University of Life Sciences Prague, Kamýcká 129, Prague-Suchdol 165 00, Czech Republic. &amp; Department of Preschool &amp; Primary Education, Faculty of Education, Jan Evangelista Purkyně University in Ústí nad Labem, Pasteurova 1, 400 96 Ústí nad Labem, Czech Republic.</mods:affiliation>
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<mods:namePart id="667755AAE4EDB7F416B01B78E17EF45D">Akmal, Surya Gentha</mods:namePart>
<mods:affiliation id="718DA62F24CED1A17A435A4B87CDD182">IPB University, The Institute for Research and Community Service, Centre for Coastal and Marine Resources Studies, Bogor 166 80, Indonesia.</mods:affiliation>
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<mods:namePart id="7C9BF480BCA2C043C2038678CCCCB1AA">Bláha, Martin</mods:namePart>
<mods:affiliation id="A1ADC080B7E083080C32BFB52BEF7D96">Faculty of Fisheries and Protection of Waters, South Bohemian Research Center of Aquaculture and Biodiversity of Hydrocenoses, University of South Bohemia in České Budějovice, Zátiší 728 / II, Vodňany 389 25, Czech Republic.</mods:affiliation>
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<mods:namePart id="C04F031A5DE548D528EEE7243D5C78C3">Kouba, Antonín</mods:namePart>
<mods:affiliation id="71EF48CB22A2EA785E4D41B74365CEF9">Faculty of Fisheries and Protection of Waters, South Bohemian Research Center of Aquaculture and Biodiversity of Hydrocenoses, University of South Bohemia in České Budějovice, Zátiší 728 / II, Vodňany 389 25, Czech Republic.</mods:affiliation>
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<taxonomicName id="4C724D013B480375419A44C724D933EE" authorityName="Patoka &amp; Akmal &amp; Bláha &amp; Kouba" authorityYear="2025" box="[151,414,1731,1758]" class="Malacostraca" family="Parastacidae" genus="Cherax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="524" phylum="Arthropoda" rank="species" species="pulverulentus" status="sp. nov.">
<emphasis id="B906EA903B480375419A44C724D933EE" bold="true" box="[151,414,1731,1758]" italics="true" pageId="2" pageNumber="524">Cherax pulverulentus</emphasis>
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<taxonomicNameLabel id="A23557EB3B48037540AB44C7274433ED" box="[422,515,1731,1757]" pageId="2" pageNumber="524" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BCD36823B480375419A44E324693231" blockId="2.[151,515,1731,1793]" box="[151,302,1767,1793]" pageId="2" pageNumber="524">
(
<figureCitation id="13492A073B480375419244E3244C3231" box="[159,267,1767,1793]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1427,1451]" captionStartId-1="4.[151,250,1887,1911]" captionStartId-2="5.[151,250,1963,1987]" captionTargetBox-0="[151,1435,669,1403]" captionTargetBox-1="[157,1428,425,1856]" captionTargetBox-2="[164,1424,201,1939]" captionTargetId-0="figure-256@3.[151,1435,669,1403]" captionTargetId-1="figure-119@4.[151,1436,417,1864]" captionTargetId-2="figure-23@5.[164,1424,201,1939]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Cherax pulverulentus sp. nov., holotype, adult male, MZB Cru5782, CL = 49 mm." captionText-1="FIGURE 2. Cherax pulverulentus sp. nov.: ad, f, g from holotype ♂, MZB Cru5782; e, h from allotype ♀, MZB Cru5783; (a) lateral view of carapace; (b) ventral view of right male chela; (c) dorsal view of right male chela; (d) dorsal view of carapace; (e) sternal keel ventral view of right chela; (f) epistome; (g) dorsal view of right scaphocerite; (h) dorsal view of right female chela. Scale bars = 1 cm." captionText-2="FIGURE 3. (a) Cherax pulverulentus sp. nov., allotype, adult female, MZB Cru5783, CL = 41 mm; (b) C. pulverulentus sp. nov., paratype 4 (purple form), adult male, MZB Cru5784, CL = 37 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14703384" figureDoi-1="http://doi.org/10.5281/zenodo.14703388" figureDoi-2="http://doi.org/10.5281/zenodo.14703394" httpUri-0="https://zenodo.org/record/14703384/files/figure.png" httpUri-1="https://zenodo.org/record/14703388/files/figure.png" httpUri-2="https://zenodo.org/record/14703394/files/figure.png" pageId="2" pageNumber="524">Figs. 13</figureCitation>
,
<figureCitation id="13492A073B480375401A44E324613231" box="[279,294,1767,1793]" captionStart="FIGURE 6" captionStartId="8.[151,250,1169,1193]" captionTargetBox="[151,1435,181,1144]" captionTargetId="figure-19@8.[151,1435,181,1144]" captionTargetPageId="8" captionText="FIGURE 6. Cherax pulverulentus sp. nov. (adult male) of blue form captured at a thermal spring in Hungary (after Bláha et al. 2022)." figureDoi="http://doi.org/10.5281/zenodo.14703403" httpUri="https://zenodo.org/record/14703403/files/figure.png" pageId="2" pageNumber="524">6</figureCitation>
)
</paragraph>
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<paragraph id="8BCD36823B480375419A452827823274" blockId="2.[151,709,1835,1890]" box="[151,709,1835,1860]" pageId="2" pageNumber="524">
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<taxonomicName id="4C724D013B480375419A452824143274" ID-CoL="7S74C" box="[151,339,1836,1860]" class="Malacostraca" family="Parastacidae" genus="Cherax" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="524" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="B906EA903B480375419A452825A53274" box="[151,226,1836,1860]" italics="true" pageId="2" pageNumber="524">Cherax</emphasis>
cf.
<emphasis id="B906EA903B4803754005452824143274" box="[264,339,1836,1860]" italics="true" pageId="2" pageNumber="524">pulcher</emphasis>
</taxonomicName>
<emphasis id="B906EA903B4803754059452824283274" box="[340,367,1836,1860]" italics="true" pageId="2" pageNumber="524">.—</emphasis>
<treatmentCitation id="0AD310933B480375407D4528277F3273" author="Blaha, M. &amp; Weiperth, A. &amp; Patoka, J. &amp; Szajbert, B. &amp; Balogh, E. R. &amp; Staszny, A. &amp; Ferincz, A. &amp; Lente, V. &amp; Maciaszek, R. &amp; Kouba, A." box="[368,568,1835,1860]" page="5" pageId="2" pageNumber="524" year="2022">
<bibRefCitation id="EFE34B733B480375407D4528277F3273" author="Blaha, M. &amp; Weiperth, A. &amp; Patoka, J. &amp; Szajbert, B. &amp; Balogh, E. R. &amp; Staszny, A. &amp; Ferincz, A. &amp; Lente, V. &amp; Maciaszek, R. &amp; Kouba, A." box="[368,568,1835,1860]" pageId="2" pageNumber="524" refId="ref5251" refString="Blaha, M., Weiperth, A., Patoka, J., Szajbert, B., Balogh, E. R., Staszny, A., Ferincz, A., Lente, V., Maciaszek, R. &amp; Kouba, A. (2022) The pet trade as a source of non-native decapods: the case of crayfish and shrimps in a thermal waterbody in Hungary. Environmental Monitoring and Assessment, 194, 795. https: // doi. org / 10.1007 / s 10661 - 022 - 10361 - 9" type="journal volume" year="2022">
Bláha
<emphasis id="B906EA903B48037540BC452824A23274" box="[433,485,1836,1860]" italics="true" pageId="2" pageNumber="524">et al.</emphasis>
2022: 5
</bibRefCitation>
</treatmentCitation>
, tab. 1, fig. 2.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCD36823B480375419A454E27103252" blockId="2.[151,709,1835,1890]" box="[151,599,1866,1890]" pageId="2" pageNumber="524">
<emphasis id="B906EA903B480375419A454E247B3252" box="[151,316,1866,1890]" italics="true" pageId="2" pageNumber="524">
<taxonomicName id="4C724D013B480375419A454E247F3252" ID-CoL="7S73S" authorityName="Roux" authorityYear="1911" box="[151,312,1866,1890]" class="Malacostraca" family="Parastacidae" genus="Cherax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="524" phylum="Arthropoda" rank="species" species="lorentzi">Cherax lorentzi</taxonomicName>
.
</emphasis>
<bibRefCitation id="EFE34B733B4803754059454E27483252" author="Patoka, J. &amp; Kalous, L. &amp; Kopecky, O." box="[340,527,1866,1890]" pageId="2" pageNumber="524" pagination="2489 - 2494" refId="ref6386" refString="Patoka, J., Kalous, L. &amp; Kopecky, O. (2014) Risk assessment of the crayfish pet trade based on data from the Czech Republic. Biological Invasions, 16, 2489 - 2494. https: // doi. org / 10.1007 / s 10530 - 014 - 0682 - 5" type="journal article" year="2014">
Patoka
<emphasis id="B906EA903B48037540AD454F24943252" box="[416,467,1866,1890]" italics="true" pageId="2" pageNumber="524">et al.</emphasis>
2014
</bibRefCitation>
: tab. 2.
</paragraph>
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<subSubSection id="C36865093B480374419A459524A035E5" lastPageId="3" lastPageNumber="525" pageId="2" pageNumber="524" type="materials_examined">
<paragraph id="8BCD36823B480375419A459527DF32C2" blockId="2.[151,1436,1936,2035]" pageId="2" pageNumber="524">
<emphasis id="B906EA903B480375419A4595240F329A" bold="true" box="[151,328,1936,1963]" pageId="2" pageNumber="524">Type material.</emphasis>
All from tributary streams to the Ayamaru Lake, Framu Subdistrict, Ayamaru District, Maybrat Regency, Southwest Papua Province, Bird's Head Peninsula, Indonesian New
<collectingCountry id="F36576123B480375450545B0211B32FE" box="[1032,1116,1972,1998]" name="Guinea-Bissau" pageId="2" pageNumber="524">Guinea</collectingCountry>
; collected by an anonymous Indonesian supplier. All specimens are adults.
</paragraph>
<paragraph id="8BCD36823B49037441CA429324A035E5" blockId="3.[151,1437,150,645]" pageId="3" pageNumber="525">
<materialsCitation id="3B1A3CDF3B49037441CA4293274B3581" box="[199,524,150,177]" collectionCode="MZB" pageId="3" pageNumber="525" specimenCode="Cru5782" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54C988203B49037441CA429324733581" box="[199,308,151,177]" pageId="3" pageNumber="525" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED63AE473B49037440324293243B3581" box="[319,380,151,177]" collectionName="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="3" pageNumber="525" type="Museum">MZB</collectionCode>
<specimenCode id="DBD49EF93B490374408E429224AE3581" box="[387,489,150,177]" pageId="3" pageNumber="525">Cru5782</specimenCode>
,
<specimenCount id="9D74FD0B3B49037440F94293274B3581" box="[500,524,151,177]" count="1" pageId="3" pageNumber="525" type="male"></specimenCount>
</materialsCitation>
.
<materialsCitation id="3B1A3CDF3B490374431A429326143581" box="[535,851,150,177]" collectionCode="MZB" pageId="3" pageNumber="525" specimenCode="Cru5783" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
<typeStatus id="54C988203B490374431A4293273A3581" box="[535,637,151,177]" pageId="3" pageNumber="525" type="allotype">Allotype</typeStatus>
:
<collectionCode id="ED63AE473B4903744385429327823581" box="[648,709,151,177]" collectionName="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="3" pageNumber="525" type="Museum">MZB</collectionCode>
<specimenCode id="DBD49EF93B49037443C0429226763581" box="[717,817,150,177]" pageId="3" pageNumber="525">Cru5783</specimenCode>
,
<specimenCount id="9D74FD0B3B4903744230429326143581" box="[829,851,151,177]" count="1" pageId="3" pageNumber="525" type="female"></specimenCount>
</materialsCitation>
.
<materialsCitation id="3B1A3CDF3B490374426C429325F135E5" collectionCode="MZB" location="Cru" pageId="3" pageNumber="525" specimenCode="Cru5784" specimenCount="2" specimenCount-female="2" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="54C988203B490374426C429326953581" box="[865,978,151,177]" pageId="3" pageNumber="525" type="paratype">Paratypes</typeStatus>
:
<collectionCode id="ED63AE473B49037442D34293215C3581" box="[990,1051,151,177]" collectionName="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="3" pageNumber="525" type="Museum">MZB</collectionCode>
<specimenCode id="DBD49EF93B490374452F429221C03581" box="[1058,1159,150,177]" pageId="3" pageNumber="525">Cru5784</specimenCode>
,
<specimenCount id="9D74FD0B3B490374459E4293219D3581" box="[1171,1242,151,177]" count="2" pageId="3" pageNumber="525" type="male">two ♂</specimenCount>
(paratypes 2 and 4),
</materialsCitation>
<materialsCitation id="3B1A3CDF3B49037441B042BF24A435E5" box="[189,483,187,213]" collectionCode="MZB" location="Cru" pageId="3" pageNumber="525" specimenCount="2" specimenCount-female="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9D74FD0B3B49037441B042BF244435E5" box="[189,259,187,213]" count="2" pageId="3" pageNumber="525" type="female">two ♀</specimenCount>
(
<typeStatus id="54C988203B490374401C42BF243935E5" box="[273,382,187,213]" pageId="3" pageNumber="525" type="paratype">paratypes</typeStatus>
1 and 3)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36865093B49037441CA42DA216E37B5" pageId="3" pageNumber="525" type="diagnosis">
<paragraph id="8BCD36823B49037441CA42DA216E37B5" blockId="3.[151,1437,150,645]" pageId="3" pageNumber="525">
<emphasis id="B906EA903B49037441CA42DA240535C8" bold="true" box="[199,322,222,248]" pageId="3" pageNumber="525">Diagnosis.</emphasis>
Carapace surface smooth except for 14 small spiniform tubercles posteriorly behind cervical groove on lateral sides of carapace. Eyes large, globular, and darkly pigmented, cornea slightly broader than eyestalk. Rostrum slender, 2.02.3 (x̅ = 2.2, SD = 0.1) times as long as wide, with slightly excavated margins; rostral margins carinate and with 3 prominent teeth. Postorbital ridges prominent with acute tubercle at anterior terminus. Median carina absent. Scaphocerite triangular, regularly narrowing towards apex, with single distinct spine at terminus. Antennular peduncle slightly reaching behind acumen. Antennal peduncle reaching slightly behind apex of scaphocerite. Areola 1.92.4 (x̅ = 2.0, SD = 0.3) times as long as wide at narrowest part. Laterally just behind cervical groove with 3 or 4 prominent anteriorly oriented spines present. Carapace 2.22.5 (x̅ = 2.3, SD = 0.1) times long as wide, and 1.11.5 (x̅ = 1.4, SD = 0.1) times longer than chela. Chela 1.92.3 (x̅ = 2.1, SD = 0.1) times as long as broad, and 3.95.0 (x̅ = 4.3, SD = 0.4) times height. Chelae of adult males with whitish uncalcified patch on lateral margin, extending from middle of palm, slightly behind base of fixed finger (propodus). Dactylar cutting edge with row of small granules and one large tubercle. Fingers with hooked tips.
</paragraph>
</subSubSection>
<caption id="DF0D660A3B490374419A47972127309B" ID-DOI="http://doi.org/10.5281/zenodo.14703384" ID-Zenodo-Dep="14703384" box="[151,1120,1427,1451]" httpUri="https://zenodo.org/record/14703384/files/figure.png" pageId="3" pageNumber="525" startId="3.[151,250,1427,1451]" targetBox="[151,1435,669,1403]" targetPageId="3" targetType="figure">
<paragraph id="8BCD36823B490374419A47972127309B" blockId="3.[151,1120,1427,1451]" box="[151,1120,1427,1451]" pageId="3" pageNumber="525">
<emphasis id="B906EA903B490374419A47972454309B" bold="true" box="[151,275,1427,1451]" pageId="3" pageNumber="525">FIGURE 1.</emphasis>
<taxonomicName id="4C724D013B4903744014479724B3309B" authorityName="Patoka &amp; Akmal &amp; Bláha &amp; Kouba" authorityYear="2025" box="[281,500,1427,1451]" class="Malacostraca" family="Parastacidae" genus="Cherax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="525" phylum="Arthropoda" rank="species" species="pulverulentus" status="sp. nov.">
<emphasis id="B906EA903B4903744014479724B3309B" box="[281,500,1427,1451]" italics="true" pageId="3" pageNumber="525">Cherax pulverulentus</emphasis>
</taxonomicName>
<emphasis id="B906EA903B49037440F64797270C309B" bold="true" box="[507,587,1427,1451]" pageId="3" pageNumber="525">
<taxonomicNameLabel id="A23557EB3B49037440F64797270C309B" box="[507,587,1427,1451]" pageId="3" pageNumber="525" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, adult male, MZB Cru5782, CL = 49 mm.
</paragraph>
</caption>
<subSubSection id="C36865093B49037E41CA47F927423455" lastPageId="9" lastPageNumber="531" pageId="3" pageNumber="525" type="description">
<paragraph id="8BCD36823B49037441CA47F926CE3398" blockId="3.[151,1437,1533,2027]" pageId="3" pageNumber="525">
<emphasis id="B906EA903B49037441CA47F927693327" bold="true" box="[199,558,1533,1559]" pageId="3" pageNumber="525">
Description of
<typeStatus id="54C988203B490374407047F924A33327" box="[381,484,1533,1559]" pageId="3" pageNumber="525" type="holotype">holotype</typeStatus>
male.
</emphasis>
(
<figureCitation id="13492A073B490374434D47F927E83327" box="[576,687,1533,1559]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[151,250,1427,1451]" captionStartId-1="4.[151,250,1887,1911]" captionTargetBox-0="[151,1435,669,1403]" captionTargetBox-1="[157,1428,425,1856]" captionTargetId-0="figure-256@3.[151,1435,669,1403]" captionTargetId-1="figure-119@4.[151,1436,417,1864]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1. Cherax pulverulentus sp. nov., holotype, adult male, MZB Cru5782, CL = 49 mm." captionText-1="FIGURE 2. Cherax pulverulentus sp. nov.: ad, f, g from holotype ♂, MZB Cru5782; e, h from allotype ♀, MZB Cru5783; (a) lateral view of carapace; (b) ventral view of right male chela; (c) dorsal view of right male chela; (d) dorsal view of carapace; (e) sternal keel ventral view of right chela; (f) epistome; (g) dorsal view of right scaphocerite; (h) dorsal view of right female chela. Scale bars = 1 cm." figureDoi-0="http://doi.org/10.5281/zenodo.14703384" figureDoi-1="http://doi.org/10.5281/zenodo.14703388" httpUri-0="https://zenodo.org/record/14703384/files/figure.png" httpUri-1="https://zenodo.org/record/14703388/files/figure.png" pageId="3" pageNumber="525">Figs. 12</figureCitation>
). Adult. Body and eyes pigmented. Body subcylindrical, slightly compressed laterally. Cephalothorax 1.1 times broader than pleon; surface smooth, densely pitted, with set of 3 anteriorly oriented small spiniform tubercles laterally just posteriorly to 3 cervical groove at level of antennae and below (both sides;
<figureCitation id="13492A073B4903744061446D24FA33B3" box="[364,445,1641,1667]" captionStart="FIGURE 2" captionStartId="4.[151,250,1887,1911]" captionTargetBox="[157,1428,425,1856]" captionTargetId="figure-119@4.[151,1436,417,1864]" captionTargetPageId="4" captionText="FIGURE 2. Cherax pulverulentus sp. nov.: ad, f, g from holotype ♂, MZB Cru5782; e, h from allotype ♀, MZB Cru5783; (a) lateral view of carapace; (b) ventral view of right male chela; (c) dorsal view of right male chela; (d) dorsal view of carapace; (e) sternal keel ventral view of right chela; (f) epistome; (g) dorsal view of right scaphocerite; (h) dorsal view of right female chela. Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14703388" httpUri="https://zenodo.org/record/14703388/files/figure.png" pageId="3" pageNumber="525">Fig. 2a</figureCitation>
). Areola 1.9 times as long as broad at the narrowest part (
<figureCitation id="13492A073B490374454C446D21D233B3" box="[1089,1173,1641,1667]" captionStart="FIGURE 2" captionStartId="4.[151,250,1887,1911]" captionTargetBox="[157,1428,425,1856]" captionTargetId="figure-119@4.[151,1436,417,1864]" captionTargetPageId="4" captionText="FIGURE 2. Cherax pulverulentus sp. nov.: ad, f, g from holotype ♂, MZB Cru5782; e, h from allotype ♀, MZB Cru5783; (a) lateral view of carapace; (b) ventral view of right male chela; (c) dorsal view of right male chela; (d) dorsal view of carapace; (e) sternal keel ventral view of right chela; (f) epistome; (g) dorsal view of right scaphocerite; (h) dorsal view of right female chela. Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14703388" httpUri="https://zenodo.org/record/14703388/files/figure.png" pageId="3" pageNumber="525">Fig. 2d</figureCitation>
). Length of areola 31% of CL; surface smooth, pitted. Cervical groove distinct, non-setose.
</paragraph>
<paragraph id="8BCD36823B49037441CA44B625B432DB" blockId="3.[151,1437,1533,2027]" pageId="3" pageNumber="525">
Rostrum (
<figureCitation id="13492A073B490374403744B524EB33FB" box="[314,428,1713,1739]" captionStart="FIGURE 2" captionStartId="4.[151,250,1887,1911]" captionTargetBox="[157,1428,425,1856]" captionTargetId="figure-119@4.[151,1436,417,1864]" captionTargetPageId="4" captionText="FIGURE 2. Cherax pulverulentus sp. nov.: ad, f, g from holotype ♂, MZB Cru5782; e, h from allotype ♀, MZB Cru5783; (a) lateral view of carapace; (b) ventral view of right male chela; (c) dorsal view of right male chela; (d) dorsal view of carapace; (e) sternal keel ventral view of right chela; (f) epistome; (g) dorsal view of right scaphocerite; (h) dorsal view of right female chela. Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14703388" httpUri="https://zenodo.org/record/14703388/files/figure.png" pageId="3" pageNumber="525">Fig. 2a, d</figureCitation>
) prominent, relatively slender, lanceolate in shape, 2.3 times as long as wide. Acumen with directly oriented spine at terminus. Median carina absent. Rostral margins elevated, anteriorly convergent throughout length to acumen, posteriorly continuing in rostral carinae on carapace. Lateral margins with 3 slightly upturned prominent teeth in distal half. Upper surface smooth, pitted, without setae, short sparse setae on outer rostral margins and on ventral side of rostrum. Rostral carinae prominent, extending as slight elevation posteriorly on carapace, gradually fading and indistinct behind middle of cephalothorax. Postorbital ridges (
<figureCitation id="13492A073B49037445A545612054324F" box="[1192,1299,1893,1919]" captionStart="FIGURE 2" captionStartId="4.[151,250,1887,1911]" captionTargetBox="[157,1428,425,1856]" captionTargetId="figure-119@4.[151,1436,417,1864]" captionTargetPageId="4" captionText="FIGURE 2. Cherax pulverulentus sp. nov.: ad, f, g from holotype ♂, MZB Cru5782; e, h from allotype ♀, MZB Cru5783; (a) lateral view of carapace; (b) ventral view of right male chela; (c) dorsal view of right male chela; (d) dorsal view of carapace; (e) sternal keel ventral view of right chela; (f) epistome; (g) dorsal view of right scaphocerite; (h) dorsal view of right female chela. Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14703388" httpUri="https://zenodo.org/record/14703388/files/figure.png" pageId="3" pageNumber="525">Fig. 2a, d</figureCitation>
) prominent, strongly elevated and gradually fading. Anterior terminus of postorbital ridges with slightly upturned spiniform tubercle. Eyes (
<figureCitation id="13492A073B490374404945A924E932F7" box="[324,430,1965,1991]" captionStart="FIGURE 2" captionStartId="4.[151,250,1887,1911]" captionTargetBox="[157,1428,425,1856]" captionTargetId="figure-119@4.[151,1436,417,1864]" captionTargetPageId="4" captionText="FIGURE 2. Cherax pulverulentus sp. nov.: ad, f, g from holotype ♂, MZB Cru5782; e, h from allotype ♀, MZB Cru5783; (a) lateral view of carapace; (b) ventral view of right male chela; (c) dorsal view of right male chela; (d) dorsal view of carapace; (e) sternal keel ventral view of right chela; (f) epistome; (g) dorsal view of right scaphocerite; (h) dorsal view of right female chela. Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14703388" httpUri="https://zenodo.org/record/14703388/files/figure.png" pageId="3" pageNumber="525">Fig. 2a, d</figureCitation>
) relatively large; cornea globular, darkly pigmented, about as long as eyestalk and slightly broader.
</paragraph>
<paragraph id="8BCD36823B4E037341CA4293260934B9" blockId="4.[151,1436,151,393]" pageId="4" pageNumber="526">
Antennules and antennae normal in shape; antennae with cut-off terminal parts, longer than TL in living individuals. Antennular peduncle slightly reaching behind acumen, antennal peduncle reaching slightly behind apex of scaphocerite. Coxicerite of antennal peduncle with spiniform tubercle anteriorly; basicerite with 1 lateral and 1 ventral spiniform and hooked tubercle. Scaphocerite (
<figureCitation id="13492A073B4E0373421743072629342D" box="[794,878,259,285]" captionStart="FIGURE 2" captionStartId="4.[151,250,1887,1911]" captionTargetBox="[157,1428,425,1856]" captionTargetId="figure-119@4.[151,1436,417,1864]" captionTargetPageId="4" captionText="FIGURE 2. Cherax pulverulentus sp. nov.: ad, f, g from holotype ♂, MZB Cru5782; e, h from allotype ♀, MZB Cru5783; (a) lateral view of carapace; (b) ventral view of right male chela; (c) dorsal view of right male chela; (d) dorsal view of carapace; (e) sternal keel ventral view of right chela; (f) epistome; (g) dorsal view of right scaphocerite; (h) dorsal view of right female chela. Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14703388" httpUri="https://zenodo.org/record/14703388/files/figure.png" pageId="4" pageNumber="526">
Fig.
<quantity id="4C8A9B673B4E0373425C43072629342D" box="[849,878,259,285]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="4" pageNumber="526" unit="g" value="2.0">2g</quantity>
</figureCitation>
) horizontal, triangular, with lamina 2.8 times as long as wide, broadest at midlength; reaching slightly behind end of antennular peduncle and acumen; regularly narrowing into apex; thickened outer lateral margin with prominent spiniform tubercle at apex reaching distinctly beyond lamina; inner margin strongly covered by dense setae.
</paragraph>
<caption id="DF0D660A3B4E0373419A455B24D232D3" ID-DOI="http://doi.org/10.5281/zenodo.14703388" ID-Zenodo-Dep="14703388" httpUri="https://zenodo.org/record/14703388/files/figure.png" pageId="4" pageNumber="526" startId="4.[151,250,1887,1911]" targetBox="[157,1428,425,1856]" targetPageId="4" targetType="figure">
<paragraph id="8BCD36823B4E0373419A455B24D232D3" blockId="4.[151,1437,1887,2019]" pageId="4" pageNumber="526">
<emphasis id="B906EA903B4E0373419A455B24543247" bold="true" box="[151,275,1887,1911]" pageId="4" pageNumber="526">FIGURE 2.</emphasis>
<taxonomicName id="4C724D013B4E03734014455B24B43247" authorityName="Patoka &amp; Akmal &amp; Bláha &amp; Kouba" authorityYear="2025" box="[281,499,1887,1911]" class="Malacostraca" family="Parastacidae" genus="Cherax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="526" phylum="Arthropoda" rank="species" species="pulverulentus" status="sp. nov.">
<emphasis id="B906EA903B4E03734014455B24B43247" box="[281,499,1887,1911]" italics="true" pageId="4" pageNumber="526">Cherax pulverulentus</emphasis>
</taxonomicName>
<emphasis id="B906EA903B4E037340F4455B270E3247" bold="true" box="[505,585,1887,1911]" pageId="4" pageNumber="526">
<taxonomicNameLabel id="A23557EB3B4E037340F4455B270E3247" box="[505,585,1887,1911]" pageId="4" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: ad, f, g from holotype ♂, MZB Cru5782; e, h from allotype ♀, MZB Cru5783; (a) lateral view of carapace; (b) ventral view of right male chela; (c) dorsal view of right male chela; (d) dorsal view of carapace; (e) sternal keel ventral view of right chela; (f) epistome; (g) dorsal view of right scaphocerite; (h) dorsal view of right female chela. Scale bars = 1 cm.
</paragraph>
</caption>
<caption id="DF0D660A3B4F0372419A45AF263E32D6" ID-DOI="http://doi.org/10.5281/zenodo.14703394" ID-Zenodo-Dep="14703394" httpUri="https://zenodo.org/record/14703394/files/figure.png" pageId="5" pageNumber="527" startId="5.[151,250,1963,1987]" targetBox="[164,1424,201,1939]" targetPageId="5" targetType="figure">
<paragraph id="8BCD36823B4F0372419A45AF263E32D6" blockId="5.[151,1436,1963,2023]" pageId="5" pageNumber="527">
<emphasis id="B906EA903B4F0372419A45AF245332F3" bold="true" box="[151,276,1963,1987]" pageId="5" pageNumber="527">FIGURE 3.</emphasis>
(a)
<taxonomicName id="4C724D013B4F0372404D45AF275B32F3" authorityName="Patoka &amp; Akmal &amp; Bláha &amp; Kouba" authorityYear="2025" box="[320,540,1963,1987]" class="Malacostraca" family="Parastacidae" genus="Cherax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="527" phylum="Arthropoda" rank="species" species="pulverulentus" status="sp. nov.">
<emphasis id="B906EA903B4F0372404D45AF275B32F3" box="[320,540,1963,1987]" italics="true" pageId="5" pageNumber="527">Cherax pulverulentus</emphasis>
</taxonomicName>
<emphasis id="B906EA903B4F0372432E45AF273232F3" bold="true" box="[547,629,1963,1987]" pageId="5" pageNumber="527">
<taxonomicNameLabel id="A23557EB3B4F0372432E45AF273232F3" box="[547,629,1963,1987]" pageId="5" pageNumber="527" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, allotype, adult female, MZB Cru5783, CL = 41 mm; (b)
<taxonomicName id="4C724D013B4F037245C345AF203132F3" authorityName="Patoka &amp; Akmal &amp; Bláha &amp; Kouba" authorityYear="2025" box="[1230,1398,1963,1987]" class="Malacostraca" family="Parastacidae" genus="Cherax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="527" phylum="Arthropoda" rank="species" species="pulverulentus" status="sp. nov.">
<emphasis id="B906EA903B4F037245C345AF219832F2" box="[1230,1247,1963,1986]" italics="true" pageId="5" pageNumber="527">C</emphasis>
.
<emphasis id="B906EA903B4F037245E145AF203132F3" box="[1260,1398,1963,1987]" italics="true" pageId="5" pageNumber="527">pulverulentus</emphasis>
</taxonomicName>
<emphasis id="B906EA903B4F0372447345AF258432D7" bold="true" pageId="5" pageNumber="527">
<taxonomicNameLabel id="A23557EB3B4F0372447345AF258432D7" pageId="5" pageNumber="527" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype 4 (purple form), adult male, MZB Cru5784, CL = 37 mm.
</paragraph>
</caption>
<caption id="DF0D660A3B4C0371419A469F265631E7" ID-DOI="http://doi.org/10.5281/zenodo.14703397" ID-Zenodo-Dep="14703397" httpUri="https://zenodo.org/record/14703397/files/figure.png" pageId="6" pageNumber="528" startId="6.[151,250,1179,1203]" targetBox="[151,1435,181,1154]" targetPageId="6" targetType="figure">
<paragraph id="8BCD36823B4C0371419A469F265631E7" blockId="6.[151,1437,1179,1239]" pageId="6" pageNumber="528">
<emphasis id="B906EA903B4C0371419A469F24573183" bold="true" box="[151,272,1179,1203]" pageId="6" pageNumber="528">FIGURE 4</emphasis>
. Western part of New Guinea, and the locality of
<taxonomicName id="4C724D013B4C03714229469F21453183" authorityName="Patoka &amp; Akmal &amp; Bláha &amp; Kouba" authorityYear="2025" box="[804,1026,1179,1203]" class="Malacostraca" family="Parastacidae" genus="Cherax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="528" phylum="Arthropoda" rank="species" species="pulverulentus" status="sp. nov.">
<emphasis id="B906EA903B4C03714229469F21453183" box="[804,1026,1179,1203]" italics="true" pageId="6" pageNumber="528">Cherax pulverulentus</emphasis>
</taxonomicName>
<emphasis id="B906EA903B4C03714506469F21183183" bold="true" box="[1035,1119,1179,1203]" pageId="6" pageNumber="528">
<taxonomicNameLabel id="A23557EB3B4C03714506469F21183183" box="[1035,1119,1179,1203]" pageId="6" pageNumber="528" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in the Birds Head Peninsula, Southwest Papua Province, Indonesia indicated by the red dot.
</paragraph>
</caption>
<paragraph id="8BCD36823B4C037141CA472D242E3080" blockId="6.[151,1437,1321,2032]" pageId="6" pageNumber="528">
Mouthparts typical. Epistome (
<figureCitation id="13492A073B4C0371432C472D27293073" box="[545,622,1321,1347]" captionStart="FIGURE 2" captionStartId="4.[151,250,1887,1911]" captionTargetBox="[157,1428,425,1856]" captionTargetId="figure-119@4.[151,1436,417,1864]" captionTargetPageId="4" captionText="FIGURE 2. Cherax pulverulentus sp. nov.: ad, f, g from holotype ♂, MZB Cru5782; e, h from allotype ♀, MZB Cru5783; (a) lateral view of carapace; (b) ventral view of right male chela; (c) dorsal view of right male chela; (d) dorsal view of carapace; (e) sternal keel ventral view of right chela; (f) epistome; (g) dorsal view of right scaphocerite; (h) dorsal view of right female chela. Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14703388" httpUri="https://zenodo.org/record/14703388/files/figure.png" pageId="6" pageNumber="528">Fig. 2f</figureCitation>
) broadly triangular, anteriorly with rounded lobe (projection) constricted at base and with setose margins; each lateral margin covered with 2 groups of small tubercles separated by smooth place; central part smooth with distinct fovea, not pitted; epistomial zygoma prominent and thick, moderately arched with oblique arms.
</paragraph>
<paragraph id="8BCD36823B4C037041CA47BD20373471" blockId="6.[151,1437,1321,2032]" lastBlockId="7.[151,1436,151,321]" lastPageId="7" lastPageNumber="529" pageId="6" pageNumber="528">
Male chelipeds (
<figureCitation id="13492A073B4C0371408F47BD24A830E3" box="[386,495,1465,1491]" captionStart="FIGURE 2" captionStartId="4.[151,250,1887,1911]" captionTargetBox="[157,1428,425,1856]" captionTargetId="figure-119@4.[151,1436,417,1864]" captionTargetPageId="4" captionText="FIGURE 2. Cherax pulverulentus sp. nov.: ad, f, g from holotype ♂, MZB Cru5782; e, h from allotype ♀, MZB Cru5783; (a) lateral view of carapace; (b) ventral view of right male chela; (c) dorsal view of right male chela; (d) dorsal view of carapace; (e) sternal keel ventral view of right chela; (f) epistome; (g) dorsal view of right scaphocerite; (h) dorsal view of right female chela. Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14703388" httpUri="https://zenodo.org/record/14703388/files/figure.png" pageId="6" pageNumber="528">Fig. 2b, c</figureCitation>
) subequal in form and size. Chelae 2.2 times as long as wide and 4.3 times as long as height, strongly compressed; chela surface smooth; palm 1.5 times longer than fingers; carapace 1.3 times longer than chela; with gap between each finger when closed and with hooked tips; dactyl broad at base, regularly tapering towards tip, ventral and dorsal surface of dactyl with scattered punctuation; fixed finger hooked, merging gradually into palm of chela; fixed finger 1.6 times broader than dactyl at base. Outer lateral margin of chelae with swollen soft and uncalcified patch extending from middle of palm slightly behind base of fixed finger; surface of uncalcified patch slightly pitted; entire inner lateral margin of palm covered with slender row of more than 10 bluntly topped teeth. Dactylar cutting edge with small granular teeth, and with large prominent tooth approximately in middle of cutting edge; sparsely setose on ventral surface near base. Dactylar tip with acute, hooked spine pointing outwards at angle of approximately 45°. Propodal cutting edge with numerous small granular teeth scattered throughout entire length of fixed finger; single large tooth present in middle length of cutting edge; wide fovea in basal half of fixed finger on ventral surface; fovea setose. Propodal tip with hooked spine. Propodal and dactylar tips crossing when fingers clasp. Carpus smooth, pitted; with well-developed acute and hooked spiniform tubercle in middle of dorsolateral inner margin (mentioned tubercle is characteristic for genus
<taxonomicName id="4C724D013B4C037142C44589215C3297" authorityName="Erichson" authorityYear="1846" box="[969,1051,1933,1959]" class="Malacostraca" family="Parastacidae" genus="Cherax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="528" phylum="Arthropoda" rank="genus">
<emphasis id="B906EA903B4C037142C44589215C3297" box="[969,1051,1933,1959]" italics="true" pageId="6" pageNumber="528">Cherax</emphasis>
</taxonomicName>
); terminating with almost directly oriented spiniform tubercle; inner lateral carpal surface covered with tiny setae; ventral surface with fovea; margins slightly elevated; inner margin with set of 6 small granules and acute spiniform tubercle oriented almost directly; outer margin with spiniform tubercle oriented directly. Merus laterally strongly depressed in basal part; surface smooth, pitted; single directly oriented spiniform tubercle present on dorsal surface; 2 directly oriented spiniform tubercles on ventral surface; ridge of small granules on entire inner ventrolateral margin, terminating with single almost directly oriented spiniform tubercle; chela about 1.8 times longer than merus. Ischium laterally strongly depressed, surface smooth and pitted, with 1 prominent and set of small spiniform tubercles on ventral margin.
</paragraph>
<caption id="DF0D660A3B4D0370419A458620CF328D" ID-DOI="http://doi.org/10.5281/zenodo.14703399" ID-Zenodo-Dep="14703399" httpUri="https://zenodo.org/record/14703399/files/figure.png" pageId="7" pageNumber="529" startId="7.[151,250,1922,1946]" targetBox="[159,1427,353,1891]" targetPageId="7" targetType="figure">
<paragraph id="8BCD36823B4D0370419A458620CF328D" blockId="7.[151,1437,1922,1982]" pageId="7" pageNumber="529">
<emphasis id="B906EA903B4D0370419A4586244932AA" bold="true" box="[151,270,1922,1946]" pageId="7" pageNumber="529">FIGURE 5</emphasis>
. Bayesian analysis consensus phylogram of selected species of
<taxonomicName id="4C724D013B4D037042AC458626AB32AA" authorityName="Erichson" authorityYear="1846" box="[929,1004,1922,1946]" class="Malacostraca" family="Parastacidae" genus="Cherax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="529" phylum="Arthropoda" rank="genus">
<emphasis id="B906EA903B4D037042AC458626AB32AA" box="[929,1004,1922,1946]" italics="true" pageId="7" pageNumber="529">Cherax</emphasis>
</taxonomicName>
based on combined COI and 16S dataset. Posterior probabilities and ML bootstrap values are displayed next to each node. The scale bar indicates the substitution rate.
</paragraph>
</caption>
<caption id="DF0D660A3B42037F419A469525BD31FD" ID-DOI="http://doi.org/10.5281/zenodo.14703403" ID-Zenodo-Dep="14703403" httpUri="https://zenodo.org/record/14703403/files/figure.png" pageId="8" pageNumber="530" startId="8.[151,250,1169,1193]" targetBox="[151,1435,181,1144]" targetPageId="8" targetType="figure">
<paragraph id="8BCD36823B42037F419A469525BD31FD" blockId="8.[151,1436,1169,1229]" pageId="8" pageNumber="530">
<emphasis id="B906EA903B42037F419A469524533199" bold="true" box="[151,276,1169,1193]" pageId="8" pageNumber="530">FIGURE 6.</emphasis>
<taxonomicName id="4C724D013B42037F4011469524BF3199" authorityName="Patoka &amp; Akmal &amp; Bláha &amp; Kouba" authorityYear="2025" box="[284,504,1169,1193]" class="Malacostraca" family="Parastacidae" genus="Cherax" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="530" phylum="Arthropoda" rank="species" species="pulverulentus" status="sp. nov.">
<emphasis id="B906EA903B42037F4011469524BF3199" box="[284,504,1169,1193]" italics="true" pageId="8" pageNumber="530">Cherax pulverulentus</emphasis>
</taxonomicName>
<emphasis id="B906EA903B42037F430D469527163199" bold="true" box="[512,593,1169,1193]" pageId="8" pageNumber="530">
<taxonomicNameLabel id="A23557EB3B42037F430D469527163199" box="[512,593,1169,1193]" pageId="8" pageNumber="530" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(adult male) of blue form captured at a thermal spring in Hungary (after
<bibRefCitation id="EFE34B733B42037F444A469525A831FD" author="Blaha, M. &amp; Weiperth, A. &amp; Patoka, J. &amp; Szajbert, B. &amp; Balogh, E. R. &amp; Staszny, A. &amp; Ferincz, A. &amp; Lente, V. &amp; Maciaszek, R. &amp; Kouba, A." pageId="8" pageNumber="530" refId="ref5251" refString="Blaha, M., Weiperth, A., Patoka, J., Szajbert, B., Balogh, E. R., Staszny, A., Ferincz, A., Lente, V., Maciaszek, R. &amp; Kouba, A. (2022) The pet trade as a source of non-native decapods: the case of crayfish and shrimps in a thermal waterbody in Hungary. Environmental Monitoring and Assessment, 194, 795. https: // doi. org / 10.1007 / s 10661 - 022 - 10361 - 9" type="journal volume" year="2022">
Bláha
<emphasis id="B906EA903B42037F4487469625F531FD" italics="true" pageId="8" pageNumber="530">et al.</emphasis>
2022
</bibRefCitation>
).
</paragraph>
</caption>
<paragraph id="8BCD36823B42037F41CA471B243A30B1" blockId="8.[151,1437,1311,2022]" pageId="8" pageNumber="530">Second pereiopod reaching behind apex of scaphocerite when extended. Palm as long as fingers, both sparsely setose; tips of fingers hooked. Carpus 1.7 times longer than palm. Merus 1.5 times longer than carpus and 1.5 times longer than ischium.</paragraph>
<paragraph id="8BCD36823B42037F41CA478F272C30DD" blockId="8.[151,1437,1311,2022]" pageId="8" pageNumber="530">Third pereiopod 1.2 times longer than second pereiopod. Palm 1.8 times longer than fingers. Fingers and palm sparsely setose; tips of fingers slightly hooked. Carpus 1.4 times longer than palm. Merus 1.4 times longer than carpus and 2.3 times longer than ischium.</paragraph>
<paragraph id="8BCD36823B42037F41CA47F3202F3305" blockId="8.[151,1437,1311,2022]" pageId="8" pageNumber="530">Fourth pereiopod reaching to about middle of scaphocerite. Propodus and dactyl setose. Dactyl slightly hooked. Propodus 1.3 times longer than carpus. Merus 1.4 times longer than carpus and 1.8 times longer than ischium.</paragraph>
<paragraph id="8BCD36823B42037F41CA443B21B0334D" blockId="8.[151,1437,1311,2022]" pageId="8" pageNumber="530">Fifth pereiopod not reaching base of scaphocerite. Propodus and dactyl setose. Dactyl slightly hooked. Propodus 1.8 times longer than carpus. Merus 1.5 times longer than carpus and 2.4 times longer than ischium.</paragraph>
<paragraph id="8BCD36823B42037F41CA4483241F33F5" blockId="8.[151,1437,1311,2022]" pageId="8" pageNumber="530">Dorsal surface of pleon smooth medially; pleura smooth, densely pitted; pleomeres strongly setose on ventral posterior margin.</paragraph>
<paragraph id="8BCD36823B42037F41CA44CB24C2323D" blockId="8.[151,1437,1311,2022]" pageId="8" pageNumber="530">Telson with 2 posteriorly directed spiniform tubercles caudolaterally; surface with tiny setae. Posterior half of tail fan membranous.</paragraph>
<paragraph id="8BCD36823B42037F41CA451320493249" blockId="8.[151,1437,1311,2022]" pageId="8" pageNumber="530">Uropodal protopod with single posteriorly directed spiniform tubercle on distal margin. Endopod with 2 posteriorly directed spiniform tubercles in middle and on outer margin of mesial lobe. Exopod with transverse row of posteriorly directed diminutive spiniform tubercles. The surface of the telson with tiny setose hairs.</paragraph>
<paragraph id="8BCD36823B42037E41CA458726F835E5" blockId="8.[151,1437,1311,2022]" lastBlockId="9.[151,1437,150,753]" lastPageId="9" lastPageNumber="531" pageId="8" pageNumber="530">
<emphasis id="B906EA903B42037F41CA4587277F32AD" bold="true" box="[199,568,1923,1949]" pageId="8" pageNumber="530">
Description of
<typeStatus id="54C988203B42037F40774587249E32AD" box="[378,473,1923,1949]" pageId="8" pageNumber="530" type="allotype">allotype</typeStatus>
female.
</emphasis>
(
<figureCitation id="13492A073B42037F4344458727DB32AD" box="[585,668,1923,1949]" captionStart="FIGURE 3" captionStartId="5.[151,250,1963,1987]" captionTargetBox="[164,1424,201,1939]" captionTargetId="figure-23@5.[164,1424,201,1939]" captionTargetPageId="5" captionText="FIGURE 3. (a) Cherax pulverulentus sp. nov., allotype, adult female, MZB Cru5783, CL = 41 mm; (b) C. pulverulentus sp. nov., paratype 4 (purple form), adult male, MZB Cru5784, CL = 37 mm." figureDoi="http://doi.org/10.5281/zenodo.14703394" httpUri="https://zenodo.org/record/14703394/files/figure.png" pageId="8" pageNumber="530">Fig. 3a</figureCitation>
). Adult. Differing from
<typeStatus id="54C988203B42037F42BD4587215532AD" box="[944,1042,1923,1949]" pageId="8" pageNumber="530" type="holotype">holotype</typeStatus>
follows: Soft uncalcified patch on cheliped palm absent (
<figureCitation id="13492A073B42037F409E45A324A132F1" box="[403,486,1959,1985]" captionStart="FIGURE 2" captionStartId="4.[151,250,1887,1911]" captionTargetBox="[157,1428,425,1856]" captionTargetId="figure-119@4.[151,1436,417,1864]" captionTargetPageId="4" captionText="FIGURE 2. Cherax pulverulentus sp. nov.: ad, f, g from holotype ♂, MZB Cru5782; e, h from allotype ♀, MZB Cru5783; (a) lateral view of carapace; (b) ventral view of right male chela; (c) dorsal view of right male chela; (d) dorsal view of carapace; (e) sternal keel ventral view of right chela; (f) epistome; (g) dorsal view of right scaphocerite; (h) dorsal view of right female chela. Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14703388" httpUri="https://zenodo.org/record/14703388/files/figure.png" pageId="8" pageNumber="530">Fig. 2h</figureCitation>
); chelae 2.3 times as long as broad and 4.5 times as long as height; palm of chela 1.1 times longer than fingers; large tubercles on propodal cutting edges smaller, indistinct and less prominent than in
<typeStatus id="54C988203B43037E419A429325BA3581" box="[151,253,151,177]" pageId="9" pageNumber="531" type="holotype">holotype</typeStatus>
. Carapace 1.5 times longer than chela; cervical groove with set of 3 anteriorly directed prominent tubercles on both sides of cephalothorax. Pleon almost as broad as cephalothorax.
</paragraph>
<paragraph id="8BCD36823B43037E41CA42DB27423455" blockId="9.[151,1437,150,753]" pageId="9" pageNumber="531">
Thoracic sternal keel (
<figureCitation id="13492A073B43037E40CA42DB275C35C9" box="[455,539,223,249]" captionStart="FIGURE 2" captionStartId="4.[151,250,1887,1911]" captionTargetBox="[157,1428,425,1856]" captionTargetId="figure-119@4.[151,1436,417,1864]" captionTargetPageId="4" captionText="FIGURE 2. Cherax pulverulentus sp. nov.: ad, f, g from holotype ♂, MZB Cru5782; e, h from allotype ♀, MZB Cru5783; (a) lateral view of carapace; (b) ventral view of right male chela; (c) dorsal view of right male chela; (d) dorsal view of carapace; (e) sternal keel ventral view of right chela; (f) epistome; (g) dorsal view of right scaphocerite; (h) dorsal view of right female chela. Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.14703388" httpUri="https://zenodo.org/record/14703388/files/figure.png" pageId="9" pageNumber="531">Fig. 2e</figureCitation>
): Sternite XXII with wide and sloped median keel anterior margin to base of coxa of pereiopod 1 to base of coxa of pereiopod 3. Sternite XII with rounded narrow and small lateral processes. Small median keel at coxopodite of pereiopod 4. Sternite XIII with parallel and wide lateral processes, angle of lateral margin with large scoops.
</paragraph>
</subSubSection>
<subSubSection id="C36865093B43037D41CA436B27F337FD" lastPageId="10" lastPageNumber="532" pageId="9" pageNumber="531" type="discussion">
<paragraph id="8BCD36823B43037E41CA436B243E37B5" blockId="9.[151,1437,150,753]" pageId="9" pageNumber="531">
<emphasis id="B906EA903B43037E41CA436B247C34B9" bold="true" box="[199,315,367,393]" pageId="9" pageNumber="531">Remarks.</emphasis>
<taxonomicName id="4C724D013B43037E404E436B277234B9" authorityName="Patoka &amp; Akmal &amp; Bláha &amp; Kouba" authorityYear="2025" box="[323,565,367,393]" class="Malacostraca" family="Parastacidae" genus="Cherax" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="531" phylum="Arthropoda" rank="species" species="pulverulentus" status="sp. nov.">
<emphasis id="B906EA903B43037E404E436B277234B9" box="[323,565,367,393]" italics="true" pageId="9" pageNumber="531">Cherax pulverulentus</emphasis>
</taxonomicName>
<emphasis id="B906EA903B43037E4330436B27D034B9" bold="true" box="[573,663,367,393]" pageId="9" pageNumber="531">
<taxonomicNameLabel id="A23557EB3B43037E4330436B27D034B9" box="[573,663,367,393]" pageId="9" pageNumber="531" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, is genetically and morphologically most similar to
<taxonomicName id="4C724D013B43037E45EB436B20D134B9" box="[1254,1430,367,393]" class="Malacostraca" family="Parastacidae" genus="Cherax" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="531" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="B906EA903B43037E45EB436B20D134B9" box="[1254,1430,367,393]" italics="true" pageId="9" pageNumber="531">Cherax pulcher</emphasis>
</taxonomicName>
, which is endemic to the Teminabuan Region, Southwest
<collectingRegion id="49B6F8603B43037E421A4397268E349D" box="[791,969,403,429]" country="Indonesia" name="Papua Barat" pageId="9" pageNumber="531">Papua Province</collectingRegion>
,
<collectingCountry id="F36576123B43037E42D843972105349D" box="[981,1090,403,429]" name="Indonesia" pageId="9" pageNumber="531">Indonesia</collectingCountry>
(Lukhaup 2015). Both species may be easily distinguished using sequence divergence, and differ in the following characters: in body and chela coloration; the length of the second pereiopod, which reaches behind the apex of scaphocerite in
<taxonomicName id="4C724D013B43037E45D143DF20D234C5" authorityName="Patoka &amp; Akmal &amp; Bláha &amp; Kouba" authorityYear="2025" box="[1244,1429,475,501]" class="Malacostraca" family="Parastacidae" genus="Cherax" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="531" phylum="Arthropoda" rank="species" species="pulverulentus">
<emphasis id="B906EA903B43037E45D143DF20D234C5" box="[1244,1429,475,501]" italics="true" pageId="9" pageNumber="531">C. pulverulentus</emphasis>
</taxonomicName>
, while reaching the middle of scaphocerite in
<taxonomicName id="4C724D013B43037E439643FB26543729" box="[667,787,511,537]" class="Malacostraca" family="Parastacidae" genus="Cherax" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="531" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="B906EA903B43037E439643FB27E93729" box="[667,686,511,537]" italics="true" pageId="9" pageNumber="531">C</emphasis>
.
<emphasis id="B906EA903B43037E43B043FB26543729" box="[701,787,511,537]" italics="true" pageId="9" pageNumber="531">pulcher</emphasis>
</taxonomicName>
; and chela is 1.92.3 times as long as broad and 3.95.0 times as its height in
<taxonomicName id="4C724D013B43037E408640272702370D" authorityName="Patoka &amp; Akmal &amp; Bláha &amp; Kouba" authorityYear="2025" box="[395,581,547,573]" class="Malacostraca" family="Parastacidae" genus="Cherax" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="531" phylum="Arthropoda" rank="species" species="pulverulentus">
<emphasis id="B906EA903B43037E408640272702370D" box="[395,581,547,573]" italics="true" pageId="9" pageNumber="531">C. pulverulentus</emphasis>
</taxonomicName>
versus 2.4 times as long as broad and 5.1 times as its height in
<taxonomicName id="4C724D013B43037E4413402720D1370D" box="[1310,1430,547,573]" class="Malacostraca" family="Parastacidae" genus="Cherax" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="531" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="B906EA903B43037E441340272076370D" box="[1310,1329,547,573]" italics="true" pageId="9" pageNumber="531">C</emphasis>
.
<emphasis id="B906EA903B43037E444D402720D1370D" box="[1344,1430,547,573]" italics="true" pageId="9" pageNumber="531">pulcher</emphasis>
</taxonomicName>
. Both species can be found in two color forms, purple and blue (see below), which is the main shared feature that makes them similar.
</paragraph>
<paragraph id="8BCD36823B43037E41CA408B244137C1" blockId="9.[151,1437,150,753]" pageId="9" pageNumber="531">
Selected morphological characteristics of all type specimens are given in
<tableCitation id="C6F003393B43037E4517408B21343799" box="[1050,1139,655,681]" captionStart="TABLE 2" captionStartId="9.[151,239,800,824]" captionTargetPageId="9" captionText="TABLE 2. Meristic counts and measurements (mm) of the holotype, allotype, and all paratypes of Cherax pulverulentus sp. nov." httpUri="http://table.plazi.org/id/DF0D660A3B43037E419A412425AA366D" pageId="9" pageNumber="531" tableUuid="DF0D660A3B43037E419A412425AA366D">Table 2</tableCitation>
. The
<typeStatus id="54C988203B43037E45B8408B20503799" box="[1205,1303,655,681]" pageId="9" pageNumber="531" type="holotype">holotype</typeStatus>
has cut-off second and third pereiopod on the left side within the analysis (these legs are preserved with the specimen in the same jar).
</paragraph>
<caption id="DF0D660A3B43037E419A412425AA366D" ID-Table-UUID="DF0D660A3B43037E419A412425AA366D" httpUri="http://table.plazi.org/id/DF0D660A3B43037E419A412425AA366D" pageId="9" pageNumber="531" startId="9.[151,239,800,824]" targetBox="[163,1378,909,1954]" targetIsTable="true" targetPageId="9" targetType="table">
<paragraph id="8BCD36823B43037E419A412425AA366D" blockId="9.[151,1436,800,861]" pageId="9" pageNumber="531">
<emphasis id="B906EA903B43037E419A4124244D3608" bold="true" box="[151,266,800,824]" pageId="9" pageNumber="531">TABLE 2.</emphasis>
Meristic counts and measurements (mm) of the holotype, allotype, and all paratypes of
<taxonomicName id="4C724D013B43037E45BB412420DB3608" authorityName="Patoka &amp; Akmal &amp; Bláha &amp; Kouba" authorityYear="2025" box="[1206,1436,800,824]" class="Malacostraca" family="Parastacidae" genus="Cherax" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="531" phylum="Arthropoda" rank="species" species="pulverulentus" status="sp. nov.">
<emphasis id="B906EA903B43037E45BB412420DB3608" box="[1206,1436,800,824]" italics="true" pageId="9" pageNumber="531">Cherax pulverulentus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557EB3B43037E419A414125AA366D" box="[151,237,837,861]" pageId="9" pageNumber="531" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BCD36823B43037E43004189207D3292" pageId="9" pageNumber="531">
<table id="F972C4223B43FC8841AE418920253292" box="[163,1378,909,1954]" gridcols="7" gridrows="24" pageId="9" pageNumber="531">
<tr id="354234C03B43FC8841AE418920253699" box="[163,1378,909,937]" gridrow="0" pageId="9" pageNumber="531" rowspan-0="1" rowspan-5="1" rowspan-6="1">
<th id="76935DBC3B43FC8843004189272C3699" box="[525,619,909,937]" gridcol="1" gridrow="0" pageId="9" pageNumber="531">Holotype</th>
<th id="76935DBC3B43FC8843CD4189265E3699" box="[704,793,909,937]" gridcol="2" gridrow="0" pageId="9" pageNumber="531">Allotype</th>
<th id="76935DBC3B43FC884265418921203699" box="[872,1127,909,937]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="9" pageNumber="531">Paratypes MZB Cru5784</th>
</tr>
<tr id="354234C03B43FC8841AE41B5202536F9" box="[163,1378,945,969]" gridrow="1" pageId="9" pageNumber="531" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76935DBC3B43FC8841AE41B524B036F9" box="[163,503,945,969]" gridcol="0" gridrow="1" pageId="9" pageNumber="531">Characteristic</th>
<td id="76935DBC3B43FC88430041B5272C36F9" box="[525,619,945,969]" gridcol="1" gridrow="1" pageId="9" pageNumber="531">MZB</td>
<td id="76935DBC3B43FC8843CD41B5265E36F9" box="[704,793,945,969]" gridcol="2" gridrow="1" pageId="9" pageNumber="531">MZB</td>
</tr>
<tr id="354234C03B43FC8841AE41D120253131" box="[163,1378,981,1025]" gridrow="2" pageId="9" pageNumber="531" rowspan-0="1">
<td id="76935DBC3B43FC88430041D1272C3131" box="[525,619,981,1025]" gridcol="1" gridrow="2" pageId="9" pageNumber="531">Cru5782</td>
<td id="76935DBC3B43FC8843CD41D1265E3131" box="[704,793,981,1025]" gridcol="2" gridrow="2" pageId="9" pageNumber="531">Cru5783</td>
<td id="76935DBC3B43FC88426541D126EA3131" box="[872,941,981,1025]" gridcol="3" gridrow="2" pageId="9" pageNumber="531">1</td>
<td id="76935DBC3B43FC8842E641D121203131" box="[1003,1127,981,1025]" gridcol="4" gridrow="2" pageId="9" pageNumber="531">2</td>
<td id="76935DBC3B43FC88458D41D121823131" box="[1152,1221,981,1025]" gridcol="5" gridrow="2" pageId="9" pageNumber="531">3</td>
<td id="76935DBC3B43FC88442C41D120253131" box="[1313,1378,981,1025]" gridcol="6" gridrow="2" pageId="9" pageNumber="531">4</td>
</tr>
<tr id="354234C03B43FC8841AE461A20253106" box="[163,1378,1054,1078]" gridrow="3" pageId="9" pageNumber="531">
<th id="76935DBC3B43FC8841AE461A24B03106" box="[163,503,1054,1078]" gridcol="0" gridrow="3" pageId="9" pageNumber="531">Sex</th>
<td id="76935DBC3B43FC884300461A272C3106" box="[525,619,1054,1078]" gridcol="1" gridrow="3" pageId="9" pageNumber="531">male</td>
<td id="76935DBC3B43FC8843CD461A265E3106" box="[704,793,1054,1078]" gridcol="2" gridrow="3" pageId="9" pageNumber="531">female</td>
<td id="76935DBC3B43FC884265461A26EA3106" box="[872,941,1054,1078]" gridcol="3" gridrow="3" pageId="9" pageNumber="531">female</td>
<td id="76935DBC3B43FC8842E6461A21203106" box="[1003,1127,1054,1078]" gridcol="4" gridrow="3" pageId="9" pageNumber="531">male</td>
<td id="76935DBC3B43FC88458D461A21823106" box="[1152,1221,1054,1078]" gridcol="5" gridrow="3" pageId="9" pageNumber="531">female</td>
<td id="76935DBC3B43FC88442C461A20253106" box="[1313,1378,1054,1078]" gridcol="6" gridrow="3" pageId="9" pageNumber="531">male</td>
</tr>
<tr id="354234C03B43FC8841AE464F20253153" box="[163,1378,1099,1123]" gridrow="4" pageId="9" pageNumber="531">
<th id="76935DBC3B43FC8841AE464F24B03153" box="[163,503,1099,1123]" gridcol="0" gridrow="4" pageId="9" pageNumber="531">Coloration form</th>
<td id="76935DBC3B43FC884300464F272C3153" box="[525,619,1099,1123]" gridcol="1" gridrow="4" pageId="9" pageNumber="531">blue</td>
<td id="76935DBC3B43FC8843CD464F265E3153" box="[704,793,1099,1123]" gridcol="2" gridrow="4" pageId="9" pageNumber="531">purple</td>
<td id="76935DBC3B43FC884265464F26EA3153" box="[872,941,1099,1123]" gridcol="3" gridrow="4" pageId="9" pageNumber="531">blue</td>
<td id="76935DBC3B43FC8842E6464F21203153" box="[1003,1127,1099,1123]" gridcol="4" gridrow="4" pageId="9" pageNumber="531">purple</td>
<td id="76935DBC3B43FC88458D464F21823153" box="[1152,1221,1099,1123]" gridcol="5" gridrow="4" pageId="9" pageNumber="531">blue</td>
<td id="76935DBC3B43FC88442C464F20253153" box="[1313,1378,1099,1123]" gridcol="6" gridrow="4" pageId="9" pageNumber="531">purple</td>
</tr>
<tr id="354234C03B43FC8841AE4672202531BE" box="[163,1378,1142,1166]" gridrow="5" pageId="9" pageNumber="531">
<th id="76935DBC3B43FC8841AE467224B031BE" box="[163,503,1142,1166]" gridcol="0" gridrow="5" pageId="9" pageNumber="531">Rostral teeth number*</th>
<td id="76935DBC3B43FC8843004672272C31BE" box="[525,619,1142,1166]" gridcol="1" gridrow="5" pageId="9" pageNumber="531">3/3</td>
<td id="76935DBC3B43FC8843CD4672265E31BE" box="[704,793,1142,1166]" gridcol="2" gridrow="5" pageId="9" pageNumber="531">2/3</td>
<td id="76935DBC3B43FC884265467226EA31BE" box="[872,941,1142,1166]" gridcol="3" gridrow="5" pageId="9" pageNumber="531">3/3</td>
<td id="76935DBC3B43FC8842E64672212031BE" box="[1003,1127,1142,1166]" gridcol="4" gridrow="5" pageId="9" pageNumber="531">3/3</td>
<td id="76935DBC3B43FC88458D4672218231BE" box="[1152,1221,1142,1166]" gridcol="5" gridrow="5" pageId="9" pageNumber="531">3/3</td>
<td id="76935DBC3B43FC88442C4672202531BE" box="[1313,1378,1142,1166]" gridcol="6" gridrow="5" pageId="9" pageNumber="531">3/3</td>
</tr>
<tr id="354234C03B43FC8841AE469A202531EC" box="[163,1378,1182,1244]" gridrow="6" pageId="9" pageNumber="531">
<th id="76935DBC3B43FC8841AE469A24B031EC" box="[163,503,1182,1244]" gridcol="0" gridrow="6" pageId="9" pageNumber="531">Cervical grove lateral tubercle number*</th>
<td id="76935DBC3B43FC884300469A272C31EC" box="[525,619,1182,1244]" gridcol="1" gridrow="6" pageId="9" pageNumber="531">4/4</td>
<td id="76935DBC3B43FC8843CD469A265E31EC" box="[704,793,1182,1244]" gridcol="2" gridrow="6" pageId="9" pageNumber="531">3/3</td>
<td id="76935DBC3B43FC884265469A26EA31EC" box="[872,941,1182,1244]" gridcol="3" gridrow="6" pageId="9" pageNumber="531">3/3</td>
<td id="76935DBC3B43FC8842E6469A212031EC" box="[1003,1127,1182,1244]" gridcol="4" gridrow="6" pageId="9" pageNumber="531">¾</td>
<td id="76935DBC3B43FC88458D469A218231EC" box="[1152,1221,1182,1244]" gridcol="5" gridrow="6" pageId="9" pageNumber="531">3/3</td>
<td id="76935DBC3B43FC88442C469A202531EC" box="[1313,1378,1182,1244]" gridcol="6" gridrow="6" pageId="9" pageNumber="531">4/3</td>
</tr>
<tr id="354234C03B43FC8841AE46EE20253032" box="[163,1378,1258,1282]" gridrow="7" pageId="9" pageNumber="531">
<th id="76935DBC3B43FC8841AE46EE24B03032" box="[163,503,1258,1282]" gridcol="0" gridrow="7" pageId="9" pageNumber="531">Carapace length</th>
<td id="76935DBC3B43FC88430046EE272C3032" box="[525,619,1258,1282]" gridcol="1" gridrow="7" pageId="9" pageNumber="531">49</td>
<td id="76935DBC3B43FC8843CD46EE265E3032" box="[704,793,1258,1282]" gridcol="2" gridrow="7" pageId="9" pageNumber="531">41</td>
<td id="76935DBC3B43FC88426546EE26EA3032" box="[872,941,1258,1282]" gridcol="3" gridrow="7" pageId="9" pageNumber="531">41</td>
<td id="76935DBC3B43FC8842E646EE21203032" box="[1003,1127,1258,1282]" gridcol="4" gridrow="7" pageId="9" pageNumber="531">39</td>
<td id="76935DBC3B43FC88458D46EE21823032" box="[1152,1221,1258,1282]" gridcol="5" gridrow="7" pageId="9" pageNumber="531">42</td>
<td id="76935DBC3B43FC88442C46EE20253032" box="[1313,1378,1258,1282]" gridcol="6" gridrow="7" pageId="9" pageNumber="531">37</td>
</tr>
<tr id="354234C03B43FC8841AE47102025301C" box="[163,1378,1300,1324]" gridrow="8" pageId="9" pageNumber="531">
<th id="76935DBC3B43FC8841AE471024B0301C" box="[163,503,1300,1324]" gridcol="0" gridrow="8" pageId="9" pageNumber="531">Carapace width</th>
<td id="76935DBC3B43FC8843004710272C301C" box="[525,619,1300,1324]" gridcol="1" gridrow="8" pageId="9" pageNumber="531">22</td>
<td id="76935DBC3B43FC8843CD4710265E301C" box="[704,793,1300,1324]" gridcol="2" gridrow="8" pageId="9" pageNumber="531">17</td>
<td id="76935DBC3B43FC884265471026EA301C" box="[872,941,1300,1324]" gridcol="3" gridrow="8" pageId="9" pageNumber="531">18</td>
<td id="76935DBC3B43FC8842E647102120301C" box="[1003,1127,1300,1324]" gridcol="4" gridrow="8" pageId="9" pageNumber="531">17</td>
<td id="76935DBC3B43FC88458D47102182301C" box="[1152,1221,1300,1324]" gridcol="5" gridrow="8" pageId="9" pageNumber="531">19</td>
<td id="76935DBC3B43FC88442C47102025301C" box="[1313,1378,1300,1324]" gridcol="6" gridrow="8" pageId="9" pageNumber="531">15</td>
</tr>
<tr id="354234C03B43FC8841AE473A20253066" box="[163,1378,1342,1366]" gridrow="9" pageId="9" pageNumber="531">
<th id="76935DBC3B43FC8841AE473A24B03066" box="[163,503,1342,1366]" gridcol="0" gridrow="9" pageId="9" pageNumber="531">Pleon width</th>
<td id="76935DBC3B43FC884300473A272C3066" box="[525,619,1342,1366]" gridcol="1" gridrow="9" pageId="9" pageNumber="531">20</td>
<td id="76935DBC3B43FC8843CD473A265E3066" box="[704,793,1342,1366]" gridcol="2" gridrow="9" pageId="9" pageNumber="531">16</td>
<td id="76935DBC3B43FC884265473A26EA3066" box="[872,941,1342,1366]" gridcol="3" gridrow="9" pageId="9" pageNumber="531">18</td>
<td id="76935DBC3B43FC8842E6473A21203066" box="[1003,1127,1342,1366]" gridcol="4" gridrow="9" pageId="9" pageNumber="531">15</td>
<td id="76935DBC3B43FC88458D473A21823066" box="[1152,1221,1342,1366]" gridcol="5" gridrow="9" pageId="9" pageNumber="531">19</td>
<td id="76935DBC3B43FC88442C473A20253066" box="[1313,1378,1342,1366]" gridcol="6" gridrow="9" pageId="9" pageNumber="531">14</td>
</tr>
<tr id="354234C03B43FC8841AE476C202530B0" box="[163,1378,1384,1408]" gridrow="10" pageId="9" pageNumber="531">
<th id="76935DBC3B43FC8841AE476C24B030B0" box="[163,503,1384,1408]" gridcol="0" gridrow="10" pageId="9" pageNumber="531">Rostral length</th>
<td id="76935DBC3B43FC884300476C272C30B0" box="[525,619,1384,1408]" gridcol="1" gridrow="10" pageId="9" pageNumber="531">14</td>
<td id="76935DBC3B43FC8843CD476C265E30B0" box="[704,793,1384,1408]" gridcol="2" gridrow="10" pageId="9" pageNumber="531">11</td>
<td id="76935DBC3B43FC884265476C26EA30B0" box="[872,941,1384,1408]" gridcol="3" gridrow="10" pageId="9" pageNumber="531">10</td>
<td id="76935DBC3B43FC8842E6476C212030B0" box="[1003,1127,1384,1408]" gridcol="4" gridrow="10" pageId="9" pageNumber="531">9</td>
<td id="76935DBC3B43FC88458D476C218230B0" box="[1152,1221,1384,1408]" gridcol="5" gridrow="10" pageId="9" pageNumber="531">11</td>
<td id="76935DBC3B43FC88442C476C202530B0" box="[1313,1378,1384,1408]" gridcol="6" gridrow="10" pageId="9" pageNumber="531">9</td>
</tr>
<tr id="354234C03B43FC8841AE47962025309A" box="[163,1378,1426,1450]" gridrow="11" pageId="9" pageNumber="531">
<th id="76935DBC3B43FC8841AE479624B0309A" box="[163,503,1426,1450]" gridcol="0" gridrow="11" pageId="9" pageNumber="531">Rostral width</th>
<td id="76935DBC3B43FC8843004796272C309A" box="[525,619,1426,1450]" gridcol="1" gridrow="11" pageId="9" pageNumber="531">6</td>
<td id="76935DBC3B43FC8843CD4796265E309A" box="[704,793,1426,1450]" gridcol="2" gridrow="11" pageId="9" pageNumber="531">5</td>
<td id="76935DBC3B43FC884265479626EA309A" box="[872,941,1426,1450]" gridcol="3" gridrow="11" pageId="9" pageNumber="531">5</td>
<td id="76935DBC3B43FC8842E647962120309A" box="[1003,1127,1426,1450]" gridcol="4" gridrow="11" pageId="9" pageNumber="531">4</td>
<td id="76935DBC3B43FC88458D47962182309A" box="[1152,1221,1426,1450]" gridcol="5" gridrow="11" pageId="9" pageNumber="531">5</td>
<td id="76935DBC3B43FC88442C47962025309A" box="[1313,1378,1426,1450]" gridcol="6" gridrow="11" pageId="9" pageNumber="531">4</td>
</tr>
<tr id="354234C03B43FC8841AE47B8202530E4" box="[163,1378,1468,1492]" gridrow="12" pageId="9" pageNumber="531">
<th id="76935DBC3B43FC8841AE47B824B030E4" box="[163,503,1468,1492]" gridcol="0" gridrow="12" pageId="9" pageNumber="531">Scaphocerite lamina length</th>
<td id="76935DBC3B43FC88430047B8272C30E4" box="[525,619,1468,1492]" gridcol="1" gridrow="12" pageId="9" pageNumber="531">14</td>
<td id="76935DBC3B43FC8843CD47B8265E30E4" box="[704,793,1468,1492]" gridcol="2" gridrow="12" pageId="9" pageNumber="531">10</td>
<td id="76935DBC3B43FC88426547B826EA30E4" box="[872,941,1468,1492]" gridcol="3" gridrow="12" pageId="9" pageNumber="531">10</td>
<td id="76935DBC3B43FC8842E647B8212030E4" box="[1003,1127,1468,1492]" gridcol="4" gridrow="12" pageId="9" pageNumber="531">10</td>
<td id="76935DBC3B43FC88458D47B8218230E4" box="[1152,1221,1468,1492]" gridcol="5" gridrow="12" pageId="9" pageNumber="531">13</td>
<td id="76935DBC3B43FC88442C47B8202530E4" box="[1313,1378,1468,1492]" gridcol="6" gridrow="12" pageId="9" pageNumber="531">10</td>
</tr>
<tr id="354234C03B43FC8841AE47E2202530CE" box="[163,1378,1510,1534]" gridrow="13" pageId="9" pageNumber="531">
<th id="76935DBC3B43FC8841AE47E224B030CE" box="[163,503,1510,1534]" gridcol="0" gridrow="13" pageId="9" pageNumber="531">Scaphocerite lamina width</th>
<td id="76935DBC3B43FC88430047E2272C30CE" box="[525,619,1510,1534]" gridcol="1" gridrow="13" pageId="9" pageNumber="531">5</td>
<td id="76935DBC3B43FC8843CD47E2265E30CE" box="[704,793,1510,1534]" gridcol="2" gridrow="13" pageId="9" pageNumber="531">4</td>
<td id="76935DBC3B43FC88426547E226EA30CE" box="[872,941,1510,1534]" gridcol="3" gridrow="13" pageId="9" pageNumber="531">4</td>
<td id="76935DBC3B43FC8842E647E2212030CE" box="[1003,1127,1510,1534]" gridcol="4" gridrow="13" pageId="9" pageNumber="531">4</td>
<td id="76935DBC3B43FC88458D47E2218230CE" box="[1152,1221,1510,1534]" gridcol="5" gridrow="13" pageId="9" pageNumber="531">5</td>
<td id="76935DBC3B43FC88442C47E2202530CE" box="[1313,1378,1510,1534]" gridcol="6" gridrow="13" pageId="9" pageNumber="531">3</td>
</tr>
<tr id="354234C03B43FC8841AE441420253318" box="[163,1378,1552,1576]" gridrow="14" pageId="9" pageNumber="531">
<th id="76935DBC3B43FC8841AE441424B03318" box="[163,503,1552,1576]" gridcol="0" gridrow="14" pageId="9" pageNumber="531">Areola length</th>
<td id="76935DBC3B43FC8843004414272C3318" box="[525,619,1552,1576]" gridcol="1" gridrow="14" pageId="9" pageNumber="531">15</td>
<td id="76935DBC3B43FC8843CD4414265E3318" box="[704,793,1552,1576]" gridcol="2" gridrow="14" pageId="9" pageNumber="531">12</td>
<td id="76935DBC3B43FC884265441426EA3318" box="[872,941,1552,1576]" gridcol="3" gridrow="14" pageId="9" pageNumber="531">12</td>
<td id="76935DBC3B43FC8842E6441421203318" box="[1003,1127,1552,1576]" gridcol="4" gridrow="14" pageId="9" pageNumber="531">12</td>
<td id="76935DBC3B43FC88458D441421823318" box="[1152,1221,1552,1576]" gridcol="5" gridrow="14" pageId="9" pageNumber="531">13</td>
<td id="76935DBC3B43FC88442C441420253318" box="[1313,1378,1552,1576]" gridcol="6" gridrow="14" pageId="9" pageNumber="531">11</td>
</tr>
<tr id="354234C03B43FC8841AE443E20253362" box="[163,1378,1594,1618]" gridrow="15" pageId="9" pageNumber="531">
<th id="76935DBC3B43FC8841AE443E24B03362" box="[163,503,1594,1618]" gridcol="0" gridrow="15" pageId="9" pageNumber="531">Areola width</th>
<td id="76935DBC3B43FC884300443E272C3362" box="[525,619,1594,1618]" gridcol="1" gridrow="15" pageId="9" pageNumber="531">8</td>
<td id="76935DBC3B43FC8843CD443E265E3362" box="[704,793,1594,1618]" gridcol="2" gridrow="15" pageId="9" pageNumber="531">5</td>
<td id="76935DBC3B43FC884265443E26EA3362" box="[872,941,1594,1618]" gridcol="3" gridrow="15" pageId="9" pageNumber="531">6</td>
<td id="76935DBC3B43FC8842E6443E21203362" box="[1003,1127,1594,1618]" gridcol="4" gridrow="15" pageId="9" pageNumber="531">8</td>
<td id="76935DBC3B43FC88458D443E21823362" box="[1152,1221,1594,1618]" gridcol="5" gridrow="15" pageId="9" pageNumber="531">7</td>
<td id="76935DBC3B43FC88442C443E20253362" box="[1313,1378,1594,1618]" gridcol="6" gridrow="15" pageId="9" pageNumber="531">5</td>
</tr>
<tr id="354234C03B43FC8841AE44602025334C" box="[163,1378,1636,1660]" gridrow="16" pageId="9" pageNumber="531">
<th id="76935DBC3B43FC8841AE446024B0334C" box="[163,503,1636,1660]" gridcol="0" gridrow="16" pageId="9" pageNumber="531">Chela length</th>
<td id="76935DBC3B43FC8843004460272C334C" box="[525,619,1636,1660]" gridcol="1" gridrow="16" pageId="9" pageNumber="531">39</td>
<td id="76935DBC3B43FC8843CD4460265E334C" box="[704,793,1636,1660]" gridcol="2" gridrow="16" pageId="9" pageNumber="531">27</td>
<td id="76935DBC3B43FC884265446026EA334C" box="[872,941,1636,1660]" gridcol="3" gridrow="16" pageId="9" pageNumber="531">28</td>
<td id="76935DBC3B43FC8842E644602120334C" box="[1003,1127,1636,1660]" gridcol="4" gridrow="16" pageId="9" pageNumber="531">34</td>
<td id="76935DBC3B43FC88458D44602182334C" box="[1152,1221,1636,1660]" gridcol="5" gridrow="16" pageId="9" pageNumber="531">30</td>
<td id="76935DBC3B43FC88442C44602025334C" box="[1313,1378,1636,1660]" gridcol="6" gridrow="16" pageId="9" pageNumber="531">27</td>
</tr>
<tr id="354234C03B43FC8841AE448A20253396" box="[163,1378,1678,1702]" gridrow="17" pageId="9" pageNumber="531">
<th id="76935DBC3B43FC8841AE448A24B03396" box="[163,503,1678,1702]" gridcol="0" gridrow="17" pageId="9" pageNumber="531">Chela width</th>
<td id="76935DBC3B43FC884300448A272C3396" box="[525,619,1678,1702]" gridcol="1" gridrow="17" pageId="9" pageNumber="531">18</td>
<td id="76935DBC3B43FC8843CD448A265E3396" box="[704,793,1678,1702]" gridcol="2" gridrow="17" pageId="9" pageNumber="531">12</td>
<td id="76935DBC3B43FC884265448A26EA3396" box="[872,941,1678,1702]" gridcol="3" gridrow="17" pageId="9" pageNumber="531">13</td>
<td id="76935DBC3B43FC8842E6448A21203396" box="[1003,1127,1678,1702]" gridcol="4" gridrow="17" pageId="9" pageNumber="531">17</td>
<td id="76935DBC3B43FC88458D448A21823396" box="[1152,1221,1678,1702]" gridcol="5" gridrow="17" pageId="9" pageNumber="531">13</td>
<td id="76935DBC3B43FC88442C448A20253396" box="[1313,1378,1678,1702]" gridcol="6" gridrow="17" pageId="9" pageNumber="531">14</td>
</tr>
<tr id="354234C03B43FC8841AE44BC202533E0" box="[163,1378,1720,1744]" gridrow="18" pageId="9" pageNumber="531">
<th id="76935DBC3B43FC8841AE44BC24B033E0" box="[163,503,1720,1744]" gridcol="0" gridrow="18" pageId="9" pageNumber="531">Chela height</th>
<td id="76935DBC3B43FC88430044BC272C33E0" box="[525,619,1720,1744]" gridcol="1" gridrow="18" pageId="9" pageNumber="531">9</td>
<td id="76935DBC3B43FC8843CD44BC265E33E0" box="[704,793,1720,1744]" gridcol="2" gridrow="18" pageId="9" pageNumber="531">6</td>
<td id="76935DBC3B43FC88426544BC26EA33E0" box="[872,941,1720,1744]" gridcol="3" gridrow="18" pageId="9" pageNumber="531">7</td>
<td id="76935DBC3B43FC8842E644BC212033E0" box="[1003,1127,1720,1744]" gridcol="4" gridrow="18" pageId="9" pageNumber="531">8</td>
<td id="76935DBC3B43FC88458D44BC218233E0" box="[1152,1221,1720,1744]" gridcol="5" gridrow="18" pageId="9" pageNumber="531">6</td>
<td id="76935DBC3B43FC88442C44BC202533E0" box="[1313,1378,1720,1744]" gridcol="6" gridrow="18" pageId="9" pageNumber="531">7</td>
</tr>
<tr id="354234C03B43FC8841AE44E6202533CA" box="[163,1378,1762,1786]" gridrow="19" pageId="9" pageNumber="531">
<th id="76935DBC3B43FC8841AE44E624B033CA" box="[163,503,1762,1786]" gridcol="0" gridrow="19" pageId="9" pageNumber="531">Chela palm length</th>
<td id="76935DBC3B43FC88430044E6272C33CA" box="[525,619,1762,1786]" gridcol="1" gridrow="19" pageId="9" pageNumber="531">24</td>
<td id="76935DBC3B43FC8843CD44E6265E33CA" box="[704,793,1762,1786]" gridcol="2" gridrow="19" pageId="9" pageNumber="531">15</td>
<td id="76935DBC3B43FC88426544E626EA33CA" box="[872,941,1762,1786]" gridcol="3" gridrow="19" pageId="9" pageNumber="531">15</td>
<td id="76935DBC3B43FC8842E644E6212033CA" box="[1003,1127,1762,1786]" gridcol="4" gridrow="19" pageId="9" pageNumber="531">20</td>
<td id="76935DBC3B43FC88458D44E6218233CA" box="[1152,1221,1762,1786]" gridcol="5" gridrow="19" pageId="9" pageNumber="531">17</td>
<td id="76935DBC3B43FC88442C44E6202533CA" box="[1313,1378,1762,1786]" gridcol="6" gridrow="19" pageId="9" pageNumber="531">17</td>
</tr>
<tr id="354234C03B43FC8841AE450820253214" box="[163,1378,1804,1828]" gridrow="20" pageId="9" pageNumber="531">
<th id="76935DBC3B43FC8841AE450824B03214" box="[163,503,1804,1828]" gridcol="0" gridrow="20" pageId="9" pageNumber="531">Chela dactyl length</th>
<td id="76935DBC3B43FC8843004508272C3214" box="[525,619,1804,1828]" gridcol="1" gridrow="20" pageId="9" pageNumber="531">16</td>
<td id="76935DBC3B43FC8843CD4508265E3214" box="[704,793,1804,1828]" gridcol="2" gridrow="20" pageId="9" pageNumber="531">13</td>
<td id="76935DBC3B43FC884265450826EA3214" box="[872,941,1804,1828]" gridcol="3" gridrow="20" pageId="9" pageNumber="531">14</td>
<td id="76935DBC3B43FC8842E6450821203214" box="[1003,1127,1804,1828]" gridcol="4" gridrow="20" pageId="9" pageNumber="531">13</td>
<td id="76935DBC3B43FC88458D450821823214" box="[1152,1221,1804,1828]" gridcol="5" gridrow="20" pageId="9" pageNumber="531">14</td>
<td id="76935DBC3B43FC88442C450820253214" box="[1313,1378,1804,1828]" gridcol="6" gridrow="20" pageId="9" pageNumber="531">12</td>
</tr>
<tr id="354234C03B43FC8841AE45322025327E" box="[163,1378,1846,1870]" gridrow="21" pageId="9" pageNumber="531">
<th id="76935DBC3B43FC8841AE453224B0327E" box="[163,503,1846,1870]" gridcol="0" gridrow="21" pageId="9" pageNumber="531">Chela propodus width</th>
<td id="76935DBC3B43FC8843004532272C327E" box="[525,619,1846,1870]" gridcol="1" gridrow="21" pageId="9" pageNumber="531">8</td>
<td id="76935DBC3B43FC8843CD4532265E327E" box="[704,793,1846,1870]" gridcol="2" gridrow="21" pageId="9" pageNumber="531">6</td>
<td id="76935DBC3B43FC884265453226EA327E" box="[872,941,1846,1870]" gridcol="3" gridrow="21" pageId="9" pageNumber="531">6</td>
<td id="76935DBC3B43FC8842E645322120327E" box="[1003,1127,1846,1870]" gridcol="4" gridrow="21" pageId="9" pageNumber="531">8</td>
<td id="76935DBC3B43FC88458D45322182327E" box="[1152,1221,1846,1870]" gridcol="5" gridrow="21" pageId="9" pageNumber="531">5</td>
<td id="76935DBC3B43FC88442C45322025327E" box="[1313,1378,1846,1870]" gridcol="6" gridrow="21" pageId="9" pageNumber="531">8</td>
</tr>
<tr id="354234C03B43FC8841AE456420253249" box="[163,1378,1888,1913]" gridrow="22" pageId="9" pageNumber="531">
<th id="76935DBC3B43FC8841AE456424B03249" box="[163,503,1888,1913]" gridcol="0" gridrow="22" pageId="9" pageNumber="531">Chela dactyl width</th>
<td id="76935DBC3B43FC8843004564272C3249" box="[525,619,1888,1913]" gridcol="1" gridrow="22" pageId="9" pageNumber="531">5</td>
<td id="76935DBC3B43FC8843CD4564265E3249" box="[704,793,1888,1913]" gridcol="2" gridrow="22" pageId="9" pageNumber="531">4</td>
<td id="76935DBC3B43FC884265456426EA3249" box="[872,941,1888,1913]" gridcol="3" gridrow="22" pageId="9" pageNumber="531">5</td>
<td id="76935DBC3B43FC8842E6456421203249" box="[1003,1127,1888,1913]" gridcol="4" gridrow="22" pageId="9" pageNumber="531">4</td>
<td id="76935DBC3B43FC88458D456421823249" box="[1152,1221,1888,1913]" gridcol="5" gridrow="22" pageId="9" pageNumber="531">4</td>
<td id="76935DBC3B43FC88442C456420253249" box="[1313,1378,1888,1913]" gridcol="6" gridrow="22" pageId="9" pageNumber="531">4</td>
</tr>
<tr id="354234C03B43FC8841AE458E20253292" box="[163,1378,1930,1954]" gridrow="23" pageId="9" pageNumber="531">
<th id="76935DBC3B43FC8841AE458E24B03292" box="[163,503,1930,1954]" gridcol="0" gridrow="23" pageId="9" pageNumber="531">Cheliped merus length</th>
<td id="76935DBC3B43FC884300458E272C3292" box="[525,619,1930,1954]" gridcol="1" gridrow="23" pageId="9" pageNumber="531">22</td>
<td id="76935DBC3B43FC8843CD458E265E3292" box="[704,793,1930,1954]" gridcol="2" gridrow="23" pageId="9" pageNumber="531">15</td>
<td id="76935DBC3B43FC884265458E26EA3292" box="[872,941,1930,1954]" gridcol="3" gridrow="23" pageId="9" pageNumber="531">15</td>
<td id="76935DBC3B43FC8842E6458E21203292" box="[1003,1127,1930,1954]" gridcol="4" gridrow="23" pageId="9" pageNumber="531">17</td>
<td id="76935DBC3B43FC88458D458E21823292" box="[1152,1221,1930,1954]" gridcol="5" gridrow="23" pageId="9" pageNumber="531">18</td>
<td id="76935DBC3B43FC88442C458E20253292" box="[1313,1378,1930,1954]" gridcol="6" gridrow="23" pageId="9" pageNumber="531">15</td>
</tr>
</table>
</paragraph>
<paragraph id="8BCD36823B43037E419A45B4241C32F8" blockId="9.[151,347,1968,1992]" box="[151,347,1968,1992]" pageId="9" pageNumber="531">
<tableNote id="7694370C3B43037E419A45B4241C32F8" box="[151,347,1968,1992]" pageId="9" pageNumber="531" targetBox="[163,1378,909,1954]" targetPageId="9">*left/right margin.</tableNote>
</paragraph>
<paragraph id="8BCD36823B40037D41CA4292216B3581" blockId="10.[151,1437,150,1545]" box="[199,1068,150,177]" pageId="10" pageNumber="532">
<emphasis id="B906EA903B40037D41CA429225B93580" bold="true" box="[199,254,150,176]" pageId="10" pageNumber="532">Size.</emphasis>
<typeStatus id="54C988203B40037D40084293242A3581" box="[261,365,151,177]" pageId="10" pageNumber="532" type="holotype">Holotype</typeStatus>
male, CL =
<quantity id="4C8A9B673B40037D40F74293270B3581" box="[506,588,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="4.9" pageId="10" pageNumber="532" unit="mm" value="49.0">49 mm</quantity>
.
<typeStatus id="54C988203B40037D4358429327F03581" box="[597,695,151,177]" pageId="10" pageNumber="532" type="allotype">Allotype</typeStatus>
female, CL =
<quantity id="4C8A9B673B40037D4255429326ED3581" box="[856,938,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" pageId="10" pageNumber="532" unit="mm" value="41.0">41 mm</quantity>
.
<typeStatus id="54C988203B40037D42B8429321503581" box="[949,1047,151,177]" pageId="10" pageNumber="532" type="paratype">Paratype</typeStatus>
1
</paragraph>
<paragraph id="8BCD36823B40037D41CA42BF25AA35C9" blockId="10.[151,1437,150,1545]" pageId="10" pageNumber="532">
female, CL =
<quantity id="4C8A9B673B40037D406942BF24F135E5" box="[356,438,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" pageId="10" pageNumber="532" unit="mm" value="41.0">41 mm</quantity>
.
<typeStatus id="54C988203B40037D40CC42BF276435E5" box="[449,547,187,213]" pageId="10" pageNumber="532" type="paratype">Paratype</typeStatus>
<specimenCount id="9D74FD0B3B40037D432642BF273E35E5" box="[555,633,187,213]" count="2" pageId="10" pageNumber="532" type="male">2 male</specimenCount>
, CL =
<quantity id="4C8A9B673B40037D43CA42BF265D35E5" box="[711,794,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="10" pageNumber="532" unit="mm" value="39.0">39 mm</quantity>
.
<typeStatus id="54C988203B40037D422842BF26C035E5" box="[805,903,187,213]" pageId="10" pageNumber="532" type="paratype">Paratype</typeStatus>
<specimenCount id="9D74FD0B3B40037D428342BF26B435E5" box="[910,1011,187,213]" count="3" pageId="10" pageNumber="532" type="female">3 female</specimenCount>
, CL =
<quantity id="4C8A9B673B40037D454D42BF21D435E5" box="[1088,1171,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" pageId="10" pageNumber="532" unit="mm" value="42.0">42 mm</quantity>
.
<typeStatus id="54C988203B40037D459342BF204735E5" box="[1182,1280,187,213]" pageId="10" pageNumber="532" type="paratype">Paratype</typeStatus>
<specimenCount id="9D74FD0B3B40037D440A42BF201135E5" box="[1287,1366,187,213]" count="4" pageId="10" pageNumber="532" type="male">4 male</specimenCount>
, CL =
<quantity id="4C8A9B673B40037D419A42DB25AE35C9" box="[151,233,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" pageId="10" pageNumber="532" unit="mm" value="37.0">37 mm</quantity>
.
</paragraph>
<paragraph id="8BCD36823B40037D41CA430626BC342D" blockId="10.[151,1437,150,1545]" box="[199,1019,258,285]" pageId="10" pageNumber="532">
<emphasis id="B906EA903B40037D41CA430624DD342C" bold="true" box="[199,410,258,284]" pageId="10" pageNumber="532">Coloration in life.</emphasis>
Two color forms exist sympatrically: purple and blue.
</paragraph>
<paragraph id="8BCD36823B40037D41CA4323246537B5" blockId="10.[151,1437,150,1545]" pageId="10" pageNumber="532">
<emphasis id="B906EA903B40037D41CA432324173471" box="[199,336,295,321]" italics="true" pageId="10" pageNumber="532">Purple form</emphasis>
(
<figureCitation id="13492A073B40037D406C432324F13471" box="[353,438,295,321]" captionStart="FIGURE 3" captionStartId="5.[151,250,1963,1987]" captionTargetBox="[164,1424,201,1939]" captionTargetId="figure-23@5.[164,1424,201,1939]" captionTargetPageId="5" captionText="FIGURE 3. (a) Cherax pulverulentus sp. nov., allotype, adult female, MZB Cru5783, CL = 41 mm; (b) C. pulverulentus sp. nov., paratype 4 (purple form), adult male, MZB Cru5784, CL = 37 mm." figureDoi="http://doi.org/10.5281/zenodo.14703394" httpUri="https://zenodo.org/record/14703394/files/figure.png" pageId="10" pageNumber="532">Fig. 3b</figureCitation>
): Background color of body dark purple, marbled on carapace sides with numerous tiny pale spots. Cephalothorax dorsolaterally near cervical groove with rounded light purple spot on each side. Pleon with prominent light purple spot on both lateral sides on each pleomere, ventrolaterally pale. Chelipeds deep blue with whitish joints, palm of propodus deep blue, outer lateral margin white, soft uncalcified patch in adult males also white. Fingers deep blue with black distal third and orange tips. Row of blunt spines on inner lateral margin of palm same color as palm. Ventral surface of palm and chela same color as dorsal side. Remaining pereiopods deep blue. Uropods blue with white joints. Soft distal part of the caudal fan not pigmented. Antennal and antennular peduncle blue, flagella light purple. Maxillipeds light blue. Ventral surface of cephalothorax and pleon pale, pleopods pale in basal half and light blue in distal half. Dorsal surface of sixth pleomere with more or less prominent reddish crossshaped spot.
</paragraph>
<paragraph id="8BCD36823B40037D41CA408B27F337FD" blockId="10.[151,1437,150,1545]" pageId="10" pageNumber="532">
<emphasis id="B906EA903B40037D41CA408B24703799" box="[199,311,655,681]" italics="true" pageId="10" pageNumber="532">Blue form</emphasis>
(
<figureCitation id="13492A073B40037D404A408B24D13799" box="[327,406,655,681]" captionStart="FIGURE 1" captionStartId="3.[151,250,1427,1451]" captionTargetBox="[151,1435,669,1403]" captionTargetId="figure-256@3.[151,1435,669,1403]" captionTargetPageId="3" captionText="FIGURE 1. Cherax pulverulentus sp. nov., holotype, adult male, MZB Cru5782, CL = 49 mm." figureDoi="http://doi.org/10.5281/zenodo.14703384" httpUri="https://zenodo.org/record/14703384/files/figure.png" pageId="10" pageNumber="532">Figs. 1</figureCitation>
,
<figureCitation id="13492A073B40037D40A9408B24863799" box="[420,449,655,681]" captionStart="FIGURE 3" captionStartId="5.[151,250,1963,1987]" captionTargetBox="[164,1424,201,1939]" captionTargetId="figure-23@5.[164,1424,201,1939]" captionTargetPageId="5" captionText="FIGURE 3. (a) Cherax pulverulentus sp. nov., allotype, adult female, MZB Cru5783, CL = 41 mm; (b) C. pulverulentus sp. nov., paratype 4 (purple form), adult male, MZB Cru5784, CL = 37 mm." figureDoi="http://doi.org/10.5281/zenodo.14703394" httpUri="https://zenodo.org/record/14703394/files/figure.png" pageId="10" pageNumber="532">3a</figureCitation>
,
<figureCitation id="13492A073B40037D40C3408B24983799" box="[462,479,655,681]" captionStart="FIGURE 6" captionStartId="8.[151,250,1169,1193]" captionTargetBox="[151,1435,181,1144]" captionTargetId="figure-19@8.[151,1435,181,1144]" captionTargetPageId="8" captionText="FIGURE 6. Cherax pulverulentus sp. nov. (adult male) of blue form captured at a thermal spring in Hungary (after Bláha et al. 2022)." figureDoi="http://doi.org/10.5281/zenodo.14703403" httpUri="https://zenodo.org/record/14703403/files/figure.png" pageId="10" pageNumber="532">6</figureCitation>
): Background color bluish, brownish, dark brown or grey. Purple coloration lacking. Other coloration patterns similar to purple form.
</paragraph>
</subSubSection>
<subSubSection id="C36865093B40037D41CA40D326A53609" pageId="10" pageNumber="532" type="etymology">
<paragraph id="8BCD36823B40037D41CA40D327863625" blockId="10.[151,1437,150,1545]" pageId="10" pageNumber="532">
<emphasis id="B906EA903B40037D41CA40D3240937C1" bold="true" box="[199,334,727,753]" pageId="10" pageNumber="532">Etymology.</emphasis>
The meaning of the Latin term
<taxonomicName id="4C724D013B40037D43B840D3260A37C1" authorityName="Patoka &amp; Akmal &amp; Bláha &amp; Kouba" authorityYear="2025" box="[693,845,727,753]" class="Malacostraca" family="Parastacidae" genus="Cherax" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="532" phylum="Arthropoda" rank="species" species="pulverulentus">
<emphasis id="B906EA903B40037D43B840D3260A37C1" box="[693,845,727,753]" italics="true" pageId="10" pageNumber="532">pulverulentus</emphasis>
</taxonomicName>
is “covered with dust” in reference to the many tiny dot-like spots on the carapace of the new species.
</paragraph>
<paragraph id="8BCD36823B40037D41CA411A26A53609" blockId="10.[151,1437,150,1545]" box="[199,994,798,825]" pageId="10" pageNumber="532">
<emphasis id="B906EA903B40037D41CA411A24C23609" bold="true" box="[199,389,798,825]" pageId="10" pageNumber="532">Common name.</emphasis>
The “dusty crayfish” is proposed as a common name.
</paragraph>
</subSubSection>
<subSubSection id="C36865093B40037D41CA4146200836F9" pageId="10" pageNumber="532" type="distribution">
<paragraph id="8BCD36823B40037D41CA4146200836F9" blockId="10.[151,1437,150,1545]" pageId="10" pageNumber="532">
<emphasis id="B906EA903B40037D41CA41462426366C" bold="true" box="[199,353,834,860]" pageId="10" pageNumber="532">Distribution.</emphasis>
Based on information from the supplier, the new species occurs westward in surrounding tributary streams to the Ayamaru Lake, Framu Subdistrict, Ayamaru District, Maybrat Regency, Southwest
<collectingRegion id="49B6F8603B40037D45EB416320DF36B1" box="[1254,1432,871,897]" country="Indonesia" name="Papua Barat" pageId="10" pageNumber="532">Papua Province</collectingRegion>
, Bird's Head Peninsula,
<collectingCountry id="F36576123B40037D4094418F27413695" box="[409,518,907,933]" name="Indonesia" pageId="10" pageNumber="532">Indonesia</collectingCountry>
(
<figureCitation id="13492A073B40037D4319418F271F3695" box="[532,600,907,933]" captionStart="FIGURE 4" captionStartId="6.[151,250,1179,1203]" captionTargetBox="[151,1435,181,1154]" captionTargetId="figure-19@6.[151,1435,181,1154]" captionTargetPageId="6" captionText="FIGURE 4. Western part of New Guinea, and the locality of Cherax pulverulentus sp. nov. in the Birds Head Peninsula, Southwest Papua Province, Indonesia indicated by the red dot." figureDoi="http://doi.org/10.5281/zenodo.14703397" httpUri="https://zenodo.org/record/14703397/files/figure.png" pageId="10" pageNumber="532">Fig. 4</figureCitation>
). The locality is on the Ayamaru limestone plateau about
<quantity id="4C8A9B673B40037D45C3418E20533695" box="[1230,1300,906,933]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="532" unit="m" value="250.0">250 m</quantity>
a.s.l. Future detailed surveys of the locality are recommended to improve the knowledge of the new species distribution.
</paragraph>
</subSubSection>
<subSubSection id="C36865093B40037D41CA41D626243339" pageId="10" pageNumber="532" type="discussion">
<paragraph id="8BCD36823B40037D41CA41D626243339" blockId="10.[151,1437,150,1545]" pageId="10" pageNumber="532">
<emphasis id="B906EA903B40037D41CA41D6243536DC" bold="true" box="[199,370,978,1004]" pageId="10" pageNumber="532">Phylogenetics.</emphasis>
The phylogenetic relationship inferred from two mitochondrial gene fragments (
<emphasis id="B906EA903B40037D440C41D7207636DD" box="[1281,1329,979,1005]" italics="true" pageId="10" pageNumber="532">COI</emphasis>
and
<emphasis id="B906EA903B40037D446441D620D436DC" box="[1385,1427,978,1004]" italics="true" pageId="10" pageNumber="532">16S</emphasis>
) results in a phylogram with a clearly defined species,
<taxonomicName id="4C724D013B40037D43F641F326F33121" authorityName="Patoka &amp; Akmal &amp; Bláha &amp; Kouba" authorityYear="2025" box="[763,948,1015,1041]" class="Malacostraca" family="Parastacidae" genus="Cherax" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="532" phylum="Arthropoda" rank="species" species="pulverulentus" status="sp. nov.">
<emphasis id="B906EA903B40037D43F641F326493121" box="[763,782,1015,1041]" italics="true" pageId="10" pageNumber="532">C</emphasis>
.
<emphasis id="B906EA903B40037D421141F326F33121" box="[796,948,1015,1041]" italics="true" pageId="10" pageNumber="532">pulverulentus</emphasis>
</taxonomicName>
<emphasis id="B906EA903B40037D42B041F3215F3121" bold="true" box="[957,1048,1015,1041]" pageId="10" pageNumber="532">
<taxonomicNameLabel id="A23557EB3B40037D42B041F3215F3121" box="[957,1048,1015,1041]" pageId="10" pageNumber="532" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13492A073B40037D452541F3212A3121" box="[1064,1133,1014,1041]" captionStart="FIGURE 5" captionStartId="7.[151,250,1922,1946]" captionTargetBox="[159,1427,353,1891]" captionTargetId="figure-102@7.[151,1436,345,1899]" captionTargetPageId="7" captionText="FIGURE 5. Bayesian analysis consensus phylogram of selected species of Cherax based on combined COI and 16S dataset. Posterior probabilities and ML bootstrap values are displayed next to each node. The scale bar indicates the substitution rate." figureDoi="http://doi.org/10.5281/zenodo.14703399" httpUri="https://zenodo.org/record/14703399/files/figure.png" pageId="10" pageNumber="532">Fig. 5</figureCitation>
). The new species forms a strongly supported clade sister to
<taxonomicName id="4C724D013B40037D4316461F27D33105" box="[539,660,1051,1077]" class="Malacostraca" family="Parastacidae" genus="Cherax" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="532" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="B906EA903B40037D4316461F27693105" box="[539,558,1051,1077]" italics="true" pageId="10" pageNumber="532">C</emphasis>
.
<emphasis id="B906EA903B40037D4333461F27D33105" box="[574,660,1051,1077]" italics="true" pageId="10" pageNumber="532">pulcher</emphasis>
</taxonomicName>
with model corrected differences of 3.8 and 2.1% for
<emphasis id="B906EA903B40037D440A461F20703105" box="[1287,1335,1051,1077]" italics="true" pageId="10" pageNumber="532">COI</emphasis>
and
<emphasis id="B906EA903B40037D447F461E20DB3104" box="[1394,1436,1050,1076]" italics="true" pageId="10" pageNumber="532">16S</emphasis>
genes, respectively. These two species form a clade as sister group to
<taxonomicName id="4C724D013B40037D4287463B21123169" box="[906,1109,1087,1113]" class="Malacostraca" family="Parastacidae" genus="Cherax" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="532" phylum="Arthropoda" rank="species" species="wagenknechtae">
<emphasis id="B906EA903B40037D4287463B26DA3169" box="[906,925,1087,1113]" italics="true" pageId="10" pageNumber="532">C</emphasis>
.
<emphasis id="B906EA903B40037D42A5463B21123169" box="[936,1109,1087,1113]" italics="true" pageId="10" pageNumber="532">wagenknechtae</emphasis>
</taxonomicName>
, from which
<taxonomicName id="4C724D013B40037D45E8463B20DB3169" authorityName="Patoka &amp; Akmal &amp; Bláha &amp; Kouba" authorityYear="2025" box="[1253,1436,1087,1113]" class="Malacostraca" family="Parastacidae" genus="Cherax" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="532" phylum="Arthropoda" rank="species" species="pulverulentus" status="sp. nov.">
<emphasis id="B906EA903B40037D45E8463B21BF3169" box="[1253,1272,1087,1113]" italics="true" pageId="10" pageNumber="532">C</emphasis>
.
<emphasis id="B906EA903B40037D4409463B20DB3169" box="[1284,1436,1087,1113]" italics="true" pageId="10" pageNumber="532">pulverulentus</emphasis>
</taxonomicName>
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differing by 6.8 and 3.8% for
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and
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genes, respectively. From three newly analysed specimens, two haplotypes were identified for both genes analysed (GenBank accession numbers PP751626 and PP751627 for
<emphasis id="B906EA903B40037D419A46AF258031F5" box="[151,199,1195,1221]" italics="true" pageId="10" pageNumber="532">COI</emphasis>
, PP751802 and PP751803 for
<emphasis id="B906EA903B40037D432646AE271231F4" box="[555,597,1194,1220]" italics="true" pageId="10" pageNumber="532">16S</emphasis>
). Both the moderate level of sequence divergence and the morphological differences described above warrant recognition of
<taxonomicName id="4C724D013B40037D43D546CB26D631D9" authorityName="Patoka &amp; Akmal &amp; Bláha &amp; Kouba" authorityYear="2025" box="[728,913,1231,1257]" class="Malacostraca" family="Parastacidae" genus="Cherax" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="532" phylum="Arthropoda" rank="species" species="pulverulentus" status="sp. nov.">
<emphasis id="B906EA903B40037D43D546CB27AC31D9" box="[728,747,1231,1257]" italics="true" pageId="10" pageNumber="532">C</emphasis>
.
<emphasis id="B906EA903B40037D43F446CB26D631D9" box="[761,913,1231,1257]" italics="true" pageId="10" pageNumber="532">pulverulentus</emphasis>
</taxonomicName>
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as distinct from the closely related
<taxonomicName id="4C724D013B40037D448E46CB25AA303D" class="Malacostraca" family="Parastacidae" genus="Cherax" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="532" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="B906EA903B40037D448E46CB20D131D9" box="[1411,1430,1231,1257]" italics="true" pageId="10" pageNumber="532">C</emphasis>
.
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. Phylogenetic analyses showed that previously studied individuals from Hungarian thermal waters (
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Weiperth
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2019
</bibRefCitation>
;
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Bláha
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2022
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) belong to
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<emphasis id="B906EA903B40037D4351471327283001" box="[604,623,1303,1329]" italics="true" pageId="10" pageNumber="532">C</emphasis>
.
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</taxonomicName>
<emphasis id="B906EA903B40037D4211471326313001" bold="true" box="[796,886,1303,1329]" pageId="10" pageNumber="532">
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</emphasis>
(OL806576, OL806577, OL790139 for
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and OL828274, OL828272, OL780440 for
<emphasis id="B906EA903B40037D4347473E27333064" box="[586,628,1338,1364]" italics="true" pageId="10" pageNumber="532">16S</emphasis>
). These previously analysed individuals represent three other haplotypes different from the
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sequences of newly described species (
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). Additionally, two mitochondrial sequences labelled as
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<emphasis id="B906EA903B40037D40174787246A30AD" box="[282,301,1411,1437]" italics="true" pageId="10" pageNumber="532">C</emphasis>
.
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(NC026227, KM501042) contain gene fragments sharing the same haplotype with our PP751626 (
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) and PP752802 (
<emphasis id="B906EA903B40037D430047A2277030F0" box="[525,567,1446,1472]" italics="true" pageId="10" pageNumber="532">16S</emphasis>
) haplotypes. This underlines the historical problem with proper identification of the species of
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originating from New
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, which is very difficult without comparison with other related species using basic molecular methods such as DNA barcoding.
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Genus
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An endemic genus for
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Genus
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A genus with a Gondwanan distribution, occurring in
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,
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20 species are known. It is the most abundant and widespread genus in
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, in almost every stream one can find species of this genus.
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A genus with a Gondwanan distribution occurring in
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,
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,
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and
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Genus
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Genus
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Imamura, 1984
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(
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12 species are known.
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A small genus with three species known from
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Genus
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<emphasis id="B931EA8FFFF6B92080ADFC073496FC44" bold="true" box="[238,442,979,1006]" italics="true" pageId="17" pageNumber="498">Coaustraliobates</emphasis>
Cook, 1974
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A genus occurring in
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,
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and New
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. Within
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four species are known.
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Genus
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K.O. Viets, 1978
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A genus confined to
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,
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and New
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(
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). Within
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26 species are known.
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Genus
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<emphasis id="B931EA8FFFFEB92880ADFA3A34A0F9A3" bold="true" box="[238,396,1518,1545]" italics="true" pageId="25" pageNumber="506">Kallimobates</emphasis>
K.O. Viets, 1978
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A small genus endemic for
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@ -97,24 +97,19 @@ Broadtoed feathertailed glider
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<emphasis id="82B8EAD0E02FFFBB9BFFFCFAFE87FCFE" bold="true" box="[151,346,798,825]" pageId="19" pageNumber="554">Type specimens.</emphasis>
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,
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(F), 570, three spirit preserved thinly furred pouch young, collected by
, 569 (F), 570, three spirit preserved thinly furred pouch young, collected by
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in the vicinity of
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on the
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of north
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.
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to
. According to
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@ -122,20 +117,16 @@ caught in the same tree … all three of the same age and size”; the three are
</materialsCitation>
.
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has had the skull removed but this was not located in the
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collection; it may have been sent to
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and formed the basis of his margin annotation regarding examination of a
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collection; it may have been sent to Oldfield Thomas in London and formed the basis of his margin annotation regarding examination of a
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in
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<date id="C4721002E02FFFBB99ADFC34FCBCFC2D" box="[709,865,977,1003]" pageId="19" pageNumber="554" value="1916-10">October 1916</date>
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.
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@ -160,7 +151,7 @@ unfurred ventrally, naked skin forming a prehensile pad (
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<emphasis id="82B8EAD0E02FFFBB9BAFFB0AFE84FACF" bold="true" box="[199,345,1263,1289]" pageId="19" pageNumber="554">Description.</emphasis>
Body measurements based on mature adults are presented in
<tableCitation id="FD4E0379E02FFFBB9F7EFB0AFBB1FACF" box="[1046,1132,1263,1289]" captionStart="TABLE 3" captionStartId="19.[152,244,152,176]" captionText="TABLE 3. Body measurements (mm) as mean±SD and range based on mature adults and sample sizes (N)." pageId="19" pageNumber="554">Table 3</tableCitation>
<tableCitation id="FD4E0379E02FFFBB9F7EFB0AFBB1FACF" box="[1046,1132,1263,1289]" captionStart="TABLE 3" captionStartId="19.[152,244,152,176]" captionTargetPageId="19" captionText="TABLE 3. Body measurements (mm) as mean±SD and range based on mature adults and sample sizes (N)." pageId="19" pageNumber="554">Table 3</tableCitation>
. Small body size, up to
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head-body length. Flanks with a narrow patagium, running along the limbs and flank from the elbow to the knee.
@ -224,7 +215,7 @@ throughout large areas of south-eastern
, with numerous instances of syntopy at the level of museum locational data (
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, Supplementary
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).
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View file

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<paragraph id="B07336C2E036FFA29BFFFA67FE39FA5A" blockId="10.[151,484,1410,1437]" box="[151,484,1410,1437]" pageId="10" pageNumber="545">
<heading id="EB3B81AEE036FFA29BFFFA67FE39FA5A" bold="true" box="[151,484,1410,1437]" fontSize="11" level="1" pageId="10" pageNumber="545" reason="1">
<taxonomicName id="77CC4D41E036FFA29BFFFA67FE39FA5A" ID-CoL="NB3" authority="Desmarest, 1818" authorityName="Desmarest" authorityYear="1818" box="[151,484,1410,1437]" class="Mammalia" family="Acrobatidae" genus="Acrobates" kingdom="Animalia" order="Diprotodontia" pageId="10" pageNumber="545" phylum="Chordata" rank="genus">
<emphasis id="82B8EAD0E036FFA29BFFFA67FE39FA5A" bold="true" box="[151,484,1410,1437]" pageId="10" pageNumber="545">
<emphasis id="82B8EAD0E036FFA29BFFFA67FECDFA5B" bold="true" box="[151,272,1410,1437]" italics="true" pageId="10" pageNumber="545">Acrobates</emphasis>
<bibRefCitation id="D45D4B33E036FFA29A7FFA66FE39FA5A" author="Desmarest, A. G." box="[279,484,1410,1437]" pageId="10" pageNumber="545" refId="ref16800" refString="Desmarest, A. G. (1818) n. k. Nouveau Dictionnaire d'Histoire Naturelle, New Edition, 25, 405." type="journal volume" year="1818">Desmarest, 1818</bibRefCitation>
</emphasis>
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</heading>
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<taxonomicName id="77CC4D41E036FFA29BFFFA22FE9FFA19" authority="Anon." authorityName="Anon." box="[151,322,1479,1503]" class="Mammalia" family="Acrobatidae" genus="Ascobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="10" pageNumber="545" phylum="Chordata" rank="genus">
<emphasis id="82B8EAD0E036FFA29BFFFA22FF23FA19" box="[151,254,1479,1503]" italics="true" pageId="10" pageNumber="545">Ascobates</emphasis>
Anon.
</taxonomicName>
in Knight, C. (1839)
<emphasis id="82B8EAD0E036FFA29974FA2DFD24FA19" box="[540,761,1479,1503]" italics="true" pageId="10" pageNumber="545">Penny Encyclopaedia</emphasis>
. London. Vol. 14, No. 907, p. 454. [error for
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<emphasis id="82B8EAD0E036FFA29FACFA22FAF7FA19" box="[1220,1322,1479,1503]" italics="true" pageId="10" pageNumber="545">Acrobates</emphasis>
</taxonomicName>
].
</paragraph>
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<taxonomicName id="77CC4D41E036FFA29BFFFA03FDFFFA38" authority="Gloger, C. W. L. (1841)" authorityName="Gloger, C. W. L." authorityYear="1841" box="[151,546,1510,1534]" class="Mammalia" family="Acrobatidae" genus="Cercoptenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="10" pageNumber="545" phylum="Chordata" rank="genus">
<emphasis id="82B8EAD0E036FFA29BFFFA03FECBFA3B" box="[151,278,1510,1533]" italics="true" pageId="10" pageNumber="545">Cercoptenus</emphasis>
Gloger, C. W. L. (1841)
</taxonomicName>
.
<emphasis id="82B8EAD0E036FFA29947FA03FB4FFA38" box="[559,1170,1510,1534]" italics="true" pageId="10" pageNumber="545">Gemeinnütziges Hand- und Hilfsbuch der Naturgeschichte</emphasis>
. Fur gebildete leser aller Stände. Bd I. Breslau: Schulz and Co.
</paragraph>
</subSubSection>
<subSubSection id="F8D66549E036FFA29BFFF9AEFC7BF9A3" box="[151,934,1610,1637]" pageId="10" pageNumber="545" type="type_taxon">
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<emphasis id="82B8EAD0E036FFA29BFFF9AEFEEFF9A2" bold="true" box="[151,306,1610,1637]" pageId="10" pageNumber="545">
<typeStatus id="6F778860E036FFA29BFFF9AEFF0FF9A3" box="[151,210,1611,1637]" pageId="10" pageNumber="545">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="77CC4D41E036FFA29A51F9AEFD7BF9A3" authority="Shaw, 1794" authorityName="Shaw" authorityYear="1794" box="[313,678,1611,1637]" class="Mammalia" family="Didelphidae" genus="Didelphis" kingdom="Animalia" order="Didelphimorphia" pageId="10" pageNumber="545" phylum="Chordata" rank="species" species="pygmaeus">
<emphasis id="82B8EAD0E036FFA29A51F9AEFDC0F9A2" box="[313,541,1611,1637]" italics="true" pageId="10" pageNumber="545">Didelphis pygmaeus</emphasis>
<bibRefCitation id="D45D4B33E036FFA2994CF9AEFD7BF9A3" author="Shaw, G." box="[548,678,1611,1637]" pageId="10" pageNumber="545" refId="ref20439" refString="Shaw, G. (1794) Zoology of New Holland. James Sowerby, London, 33 pp." type="book" year="1794">Shaw, 1794</bibRefCitation>
</taxonomicName>
by original monotypy.
</paragraph>
</subSubSection>
<subSubSection id="F8D66549E036FFA29BAFF98BFBBFF86F" pageId="10" pageNumber="545" type="diagnosis">
<paragraph id="B07336C2E036FFA29BAFF98BFBBFF86F" blockId="10.[151,1437,1610,1961]" pageId="10" pageNumber="545">
<emphasis id="82B8EAD0E036FFA29BAFF98BFE7EF94E" bold="true" box="[199,419,1646,1672]" pageId="10" pageNumber="545">Generic diagnosis.</emphasis>
Differing externally from
<taxonomicName id="77CC4D41E036FFA299A7F98AFCB9F94F" ID-CoL="472M" box="[719,868,1647,1673]" class="Mammalia" family="Acrobatidae" genus="Distoechurus" kingdom="Animalia" order="Diprotodontia" pageId="10" pageNumber="545" phylum="Chordata" rank="genus">
<emphasis id="82B8EAD0E036FFA299A7F98AFCB9F94F" box="[719,868,1647,1673]" italics="true" pageId="10" pageNumber="545">Distoechurus</emphasis>
</taxonomicName>
, the only other genus of acrobatid possums, in the possession of a patagial membrane between the elbow and ankle joints.
<bibRefCitation id="D45D4B33E036FFA298D2F976FB59F96B" author="Beck, R. M. D. &amp; Voss, R. S. &amp; Jansa, S. A." box="[954,1156,1683,1709]" pageId="10" pageNumber="545" pagination="1 - 350" refId="ref16290" refString="Beck, R. M. D., Voss, R. S. &amp; Jansa, S. A. (2022) Craniodental morphology and phylogeny of marsupials. Bulletin of the American Museum of Natural History, 457, 1 - 350. https: // doi. org / 10.5531 / sd. sp. 54" type="journal article" year="2022">
Beck
<emphasis id="82B8EAD0E036FFA29892F976FBEEF96B" box="[1018,1075,1683,1709]" italics="true" pageId="10" pageNumber="545">et al.</emphasis>
(2022)
</bibRefCitation>
list the following cranial and dental features as diagnosing
<taxonomicName id="77CC4D41E036FFA29966F952FDA2F917" ID-CoL="NB3" authorityName="Desmarest" authorityYear="1818" box="[526,639,1719,1745]" class="Mammalia" family="Acrobatidae" genus="Acrobates" kingdom="Animalia" order="Diprotodontia" pageId="10" pageNumber="545" phylum="Chordata" rank="genus">
<emphasis id="82B8EAD0E036FFA29966F952FDA2F917" box="[526,639,1719,1745]" italics="true" pageId="10" pageNumber="545">Acrobates</emphasis>
</taxonomicName>
from
<taxonomicName id="77CC4D41E036FFA299A9F952FC8BF917" ID-CoL="472M" box="[705,854,1719,1745]" class="Mammalia" family="Acrobatidae" genus="Distoechurus" kingdom="Animalia" order="Diprotodontia" pageId="10" pageNumber="545" phylum="Chordata" rank="genus">
<emphasis id="82B8EAD0E036FFA299A9F952FC8BF917" box="[705,854,1719,1745]" italics="true" pageId="10" pageNumber="545">Distoechurus</emphasis>
</taxonomicName>
: differing cranially in having a proportionally larger and more inflated neurocranium, a proportionally shorter and deeper rostrum, a vertically expanded sphenorbital fissure; a deep sulcus that houses the transverse canal vein extends laterally from the carotid canal and is floored ventrally by a strut of the alisphenoid vs by the posterior part of the pterygoid in
<taxonomicName id="77CC4D41E036FFA29F68F8C6FB48F8FB" ID-CoL="472M" box="[1024,1173,1827,1853]" class="Mammalia" family="Acrobatidae" genus="Distoechurus" kingdom="Animalia" order="Diprotodontia" pageId="10" pageNumber="545" phylum="Chordata" rank="genus">
<emphasis id="82B8EAD0E036FFA29F68F8C6FB48F8FB" box="[1024,1173,1827,1853]" italics="true" pageId="10" pageNumber="545">Distoechurus</emphasis>
</taxonomicName>
, a narrower mandibular angular process; differing dentally in having P
<superScript id="47B99B8AE036FFA299F0F8A3FD7DF892" attach="left" box="[664,672,1862,1876]" fontSize="6" pageId="10" pageNumber="545">3</superScript>
relatively large and subequal in size to P
<superScript id="47B99B8AE036FFA29F0DF8A3FBB0F892" attach="left" box="[1125,1133,1862,1876]" fontSize="6" pageId="10" pageNumber="545">1</superScript>
(reduced in
<taxonomicName id="77CC4D41E036FFA29F9FF8A2FA51F8A7" ID-CoL="472M" box="[1271,1420,1863,1889]" class="Mammalia" family="Acrobatidae" genus="Distoechurus" kingdom="Animalia" order="Diprotodontia" pageId="10" pageNumber="545" phylum="Chordata" rank="genus">
<emphasis id="82B8EAD0E036FFA29F9FF8A2FA51F8A7" box="[1271,1420,1863,1889]" italics="true" pageId="10" pageNumber="545">Distoechurus</emphasis>
</taxonomicName>
), M
<superScript id="47B99B8AE036FFA29BD8F88FFF1BF8BE" attach="left" box="[176,198,1898,1912]" fontSize="6" pageId="10" pageNumber="545">1-2</superScript>
square rather than triangular, with the protocone positioned lingual to the paracone, and with more prominent buccal cingula, M
<superScript id="47B99B8AE036FFA29A09F86BFEB4F85A" attach="left" box="[353,361,1934,1948]" fontSize="6" pageId="10" pageNumber="545">3</superScript>
relatively large, more than one half the occlusal area of M
<superScript id="47B99B8AE036FFA29890F86BFBDDF85A" attach="left" box="[1016,1024,1934,1948]" fontSize="6" pageId="10" pageNumber="545">2</superScript>
(
<figureCitation id="28F72A47E036FFA29F78F86AFB89F86F" box="[1040,1108,1935,1961]" captionStart="FIGURE 6" captionStartId="11.[151,250,1165,1189]" captionTargetBox="[239,1348,181,1142]" captionTargetId="figure-18@11.[239,1348,181,1142]" captionTargetPageId="11" captionText="FIGURE 6. Images of cranium and mandible of Acrobates via 3D reconstructions from Micro-CT scans. AE) A. frontalis ANWC CM29876 Julatten Qld, FJ) A. pygmaeus ANWC CM3745 Bondo NSW, KO) Distoechurus pennatus AMS M14752 Mt Karimui, South Simbu Province, PNG, in dorsal, ventral, lateral cranial views and buccal and occlusal mandible views. The crania, mandibles and teeth of the two Acrobates species are very similar (also see Supplementary Fig. S2 for illustration of variation within each species). Acrobates differs from Distoechurus cranially in having a proportionally larger and more inflated neurocranium, a proportionally shorter and deeper rostrum, a narrower mandibular angular process. See the text for a more extensive description." figureDoi="http://doi.org/10.5281/zenodo.14703426" httpUri="https://zenodo.org/record/14703426/files/figure.png" pageId="10" pageNumber="545">Fig. 6</figureCitation>
).
</paragraph>
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<paragraph id="B07336C2E037FFA39BFFFA2AFDEAFA2F" blockId="11.[151,896,1487,1550]" box="[151,567,1487,1514]" pageId="11" pageNumber="546">
<heading id="EB3B81AEE037FFA39BFFFA2AFDEAFA2F" bold="true" box="[151,567,1487,1514]" fontSize="11" level="1" pageId="11" pageNumber="546" reason="1">
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<emphasis id="82B8EAD0E037FFA39BFFFA2AFDEAFA2F" bold="true" box="[151,567,1487,1514]" pageId="11" pageNumber="546">
<emphasis id="82B8EAD0E037FFA39BFFFA2AFE4DFA2C" bold="true" box="[151,400,1487,1514]" italics="true" pageId="11" pageNumber="546">Acrobates pygmaeus</emphasis>
(
<bibRefCitation id="D45D4B33E037FFA39AF7FA2AFDF3FA2F" author="Shaw, G." box="[415,558,1487,1513]" pageId="11" pageNumber="546" refId="ref20439" refString="Shaw, G. (1794) Zoology of New Holland. James Sowerby, London, 33 pp." type="book" year="1794">Shaw, 1794</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F8D66549E037FFA39BFFFA11FC5DF9C8" box="[151,896,1524,1550]" pageId="11" pageNumber="546" type="distribution">
<paragraph id="B07336C2E037FFA39BFFFA11FC5DF9C8" blockId="11.[151,896,1487,1550]" box="[151,896,1524,1550]" pageId="11" pageNumber="546">
<emphasis id="82B8EAD0E037FFA39BFFFA11FDD8F9C8" bold="true" box="[151,517,1524,1550]" pageId="11" pageNumber="546">Recommended common name.</emphasis>
Narrowtoed feathertailed glider
</paragraph>
</subSubSection>
</treatment>
</document>