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<document id="7C1413464C32D4188BF02A0B982ADACC" ID-DOI="10.5852/ejt.2025.981.2811" ID-ISSN="2118-9773" ID-Zenodo-Dep="15120438" ID-ZooBank="75250E3D-A78F-44F3-87F2-4823E5BD59E1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1743518275313" checkinUser="plazi" docAuthor="Hlaváč, Peter" docDate="2025" docId="038C879DFFC5EC29FD96FB89FE99FB05" docLanguage="en" docName="EJT.2025.981.76-95.pdf" docOrigin="European Journal of Taxonomy 981" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2811/12831" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Apoderiger spinosus" docType="treatment" docVersion="3" lastPageNumber="89" masterDocId="FFB5FFE5FFCEEC24FFDEFF9BFFA0FF9C" masterDocTitle="The Pselaphinae (Coleoptera: Staphylinidae) of Madagascar. VIII. Revision of the myrmecophilous clavigerine genus Apoderiger Wasmann, 1897" masterLastPageNumber="95" masterPageNumber="76" pageNumber="87" updateTime="1743681075659" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="07B64502D63AEA37C48C84FF01B90B28">The Pselaphinae (Coleoptera: Staphylinidae) of Madagascar. VIII. Revision of the myrmecophilous clavigerine genus Apoderiger Wasmann, 1897</mods:title>
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<mods:namePart id="E591501BAD3F9608879CC3D6F70F0B79">Hlaváč, Peter</mods:namePart>
<mods:affiliation id="5AC801C76EB7B98DC482AC9636158563">Department of Zoology, Fisheries, Hydrobiology and Apiculture, Faculty of AgriSciences, Mendel University, Zemědělská 1, Brno, CZ 613 00, Czech Republic.</mods:affiliation>
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<mods:date id="0732D8105DACAF4AE9BB74B715FA211F">2025</mods:date>
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<treatment id="038C879DFFC5EC29FD96FB89FE99FB05" LSID="urn:lsid:plazi:treatment:038C879DFFC5EC29FD96FB89FE99FB05" httpUri="http://treatment.plazi.org/id/038C879DFFC5EC29FD96FB89FE99FB05" lastPageId="13" lastPageNumber="89" pageId="11" pageNumber="87">
<subSubSection id="C33F6500FFC5EC2FFD96FB89FC4CFBB1" box="[584,1004,1042,1069]" pageId="11" pageNumber="87" type="nomenclature">
<paragraph id="8B9A368BFFC5EC2FFD96FB89FC4CFBB1" blockId="11.[584,1004,1042,1104]" box="[584,1004,1042,1069]" pageId="11" pageNumber="87">
<heading id="D0D281E7FFC5EC2FFD96FB89FC4CFBB1" box="[584,1004,1042,1069]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="87" reason="2">
<taxonomicName id="4C254D08FFC5EC2FFD96FB89FC4CFBB1" ID-CoL="FRJW" authority="(Dajoz, 1982)" baseAuthorityName="Dajoz" baseAuthorityYear="1982" box="[584,1004,1042,1069]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="87" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B951EA99FFC5EC2FFD96FB89FCE1FBB0" bold="true" box="[584,833,1042,1069]" italics="true" pageId="11" pageNumber="87">Apoderiger spinosus</emphasis>
(
<bibRefCitation id="EFB44B7AFFC5EC2FFC8FFB89FC43FBB1" author="Dajoz R." box="[849,995,1042,1069]" pageId="11" pageNumber="87" pagination="481 - 522" refId="ref8039" refString="Dajoz R. 1982. Contribution a l'etude des Coleopteres Pselaphidae de Madagascar. Bulletin du Museum national d'Histoire naturelle. Section A, Zoologie, Biologie et Ecologie animales (4): 481-522. https://doi.org/10.5962/p.286051" type="journal article" year="1982">Dajoz, 1982</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F6500FFC5EC2FFD11FBADFCC4FBCC" box="[719,868,1077,1104]" pageId="11" pageNumber="87" type="description">
<paragraph id="8B9A368BFFC5EC2FFD11FBADFCC4FBCC" blockId="11.[584,1004,1042,1104]" box="[719,868,1077,1104]" pageId="11" pageNumber="87">
<figureCitation id="131E2A0EFFC5EC2FFD11FBADFCE9FBCC" box="[719,841,1078,1104]" captionStart="Fig" captionStartId="12.[189,232,1892,1918]" captionTargetBox="[210,1368,284,1850]" captionTargetId="figure-12@12.[199,1369,269,1854]" captionTargetPageId="12" captionText="Fig. 4. Dorsal and lateral view of aedeagus. AB. Apoderiger torticornis Jeannel, 1960, lectotype (MNHN). CD. Apoderiger sikorai sp. nov., paratype (NMW). EF. Apoderiger spinosus (Dajoz, 1982), holotype (MNHN).GH. Apoderiger banari sp. nov., holotype (PCPH).IJ. Apoderiger grandis sp. nov., holotype (MHNG)." figureDoi="http://doi.org/10.5281/zenodo.15120446" httpUri="https://zenodo.org/record/15120446/files/figure.png" pageId="11" pageNumber="87">Figs 4EF</figureCitation>
,
<figureCitation id="131E2A0EFFC5EC2FFC8BFBAEFCC4FBCC" box="[853,868,1077,1104]" captionStart="Fig" captionStartId="14.[185,228,1998,2024]" captionTargetBox="[297,1291,509,1959]" captionTargetId="figure-67@14.[297,1291,509,1960]" captionTargetPageId="14" captionText="Fig. 5. Distribution map of Apoderiger Wasmann, 1897." figureDoi="http://doi.org/10.5281/zenodo.15120450" httpUri="https://zenodo.org/record/15120450/files/figure.png" pageId="11" pageNumber="87">5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33F6500FFC5EC2FFF63FBECFAFDFB0E" box="[189,1373,1143,1170]" pageId="11" pageNumber="87" type="reference_group">
<paragraph id="8B9A368BFFC5EC2FFF63FBECFAFDFB0E" blockId="11.[189,1373,1143,1170]" box="[189,1373,1143,1170]" pageId="11" pageNumber="87">
<treatmentCitationGroup id="AB3511A5FFC5EC2FFF63FBECFAFDFB0E" box="[189,1373,1143,1170]" pageId="11" pageNumber="87">
<treatmentCitation id="0A84109AFFC5EC2FFF63FBECFDFCFB0E" author="Dajoz R." box="[189,604,1143,1170]" page="513" pageId="11" pageNumber="87" year="1982">
<taxonomicName id="4C254D08FFC5EC2FFF63FBECFDFCFB0E" authority="Dajoz, 1982: 513" authorityName="Dajoz" authorityPageNumber="513" authorityYear="1982" box="[189,604,1143,1170]" class="Insecta" family="Staphylinidae" genus="Soalala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="87" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B951EA99FFC5EC2FFF63FBECFE25FB0D" box="[189,389,1143,1169]" italics="true" pageId="11" pageNumber="87">Soalala spinosus</emphasis>
<bibRefCitation id="EFB44B7AFFC5EC2FFE53FBECFDFCFB0E" author="Dajoz R." box="[397,604,1143,1170]" pageId="11" pageNumber="87" pagination="481 - 522" refId="ref8039" refString="Dajoz R. 1982. Contribution a l'etude des Coleopteres Pselaphidae de Madagascar. Bulletin du Museum national d'Histoire naturelle. Section A, Zoologie, Biologie et Ecologie animales (4): 481-522. https://doi.org/10.5962/p.286051" type="journal article" year="1982">Dajoz, 1982: 513</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="549E8829FFC5EC2FFDB8FBE3FD00FB0E" box="[614,672,1144,1170]" pageId="11" pageNumber="87">Type</typeStatus>
locality:
<collectingCountry id="F332761BFFC5EC2FFCD1FBECFC3DFB0D" box="[783,925,1143,1169]" name="Madagascar" pageId="11" pageNumber="87">Madagascar</collectingCountry>
Ouest, Soalala.
<typeStatus id="549E8829FFC5EC2FFB82FBE3FB06FB0E" box="[1116,1190,1144,1170]" pageId="11" pageNumber="87">Types</typeStatus>
: HT
<specimenCount id="9D23FD02FFC5EC2FFB01FBE3FB55FB0E" box="[1247,1269,1144,1170]" count="1" pageId="11" pageNumber="87" type="male"></specimenCount>
MNHN.
</treatmentCitationGroup>
</paragraph>
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<subSubSection id="C33F6500FFC5EC2FFF63FB23FBE8FAF7" pageId="11" pageNumber="87" type="diagnosis">
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<emphasis id="B951EA99FFC5EC2FFF63FB23FE97FB4E" bold="true" box="[189,311,1208,1234]" pageId="11" pageNumber="87">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B9A368BFFC5EC2FFF63FB73FBE8FAF7" blockId="11.[189,1400,1255,1387]" pageId="11" pageNumber="87">Last antennomere strongly curved in distal portion; posterior metaventral process long, slender, subtriangular with bunch of about 6 long setae; lacking sutural and discal striae; composite tergite in posterior portion with pair of small, bump-like, widely separated projections; mesotibia with modified distal portion; aedeagus 2.60 × as long as wide; endophallus lacking sclerites.</paragraph>
</subSubSection>
<subSubSection id="C33F6500FFC5EC2FFF63FA09FC6BF9F4" pageId="11" pageNumber="87" type="materials_examined">
<paragraph id="8B9A368BFFC5EC2FFF63FA09FECDFA30" blockId="11.[189,365,1426,1452]" box="[189,365,1426,1452]" pageId="11" pageNumber="87">
<heading id="D0D281E7FFC5EC2FFF63FA09FECDFA30" bold="true" box="[189,365,1426,1452]" fontSize="11" level="3" pageId="11" pageNumber="87" reason="3">
<emphasis id="B951EA99FFC5EC2FFF63FA09FECDFA30" bold="true" box="[189,365,1426,1452]" pageId="11" pageNumber="87">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B4D3CD6FFC5EC2FFF34FA5AFC6BF9F4" collectingDate="1973-02" collectingDateMax="1988-03" collectingDateMin="1973-02" collectionCode="MNHN" collectorName="A. Peyrieras" country="Madagascar" determinerName="Cl. Besuchet" location="SOALALA" pageId="11" pageNumber="87" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<paragraph id="8B9A368BFFC5EC2FFF34FA5AFEFDFA47" blockId="11.[185,1399,1473,1640]" box="[234,349,1473,1499]" pageId="11" pageNumber="87">
<emphasis id="B951EA99FFC5EC2FFF34FA5AFEFDFA47" bold="true" box="[234,349,1473,1499]" pageId="11" pageNumber="87">
<typeStatus id="549E8829FFC5EC2FFF34FA5AFEFDFA47" box="[234,349,1473,1499]" pageId="11" pageNumber="87" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B9A368BFFC5EC2FFF63FA7FFC6BF9F4" blockId="11.[185,1399,1473,1640]" pageId="11" pageNumber="87">
<collectingCountry id="F332761BFFC5EC2FFF63FA7FFE2EFA62" box="[189,398,1508,1534]" name="Madagascar" pageId="11" pageNumber="87">MADAGASCAR</collectingCountry>
<specimenCount id="9D23FD02FFC5EC2FFE74FA7FFE63FA62" box="[426,451,1508,1534]" count="1" pageId="11" pageNumber="87" type="male"></specimenCount>
; six labels “Madagascar:
<taxonomicName id="4C254D08FFC5EC2FFD28FA7FFCDFFA62" authorityName="Dajoz" authorityYear="1982" box="[758,895,1508,1534]" class="Insecta" family="Staphylinidae" genus="Soalala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="87" phylum="Arthropoda" rank="genus">
<location id="8EFA6050FFC5EC2FFD28FA7FFCDFFA62" LSID="urn:lsid:plazi:treatment:038C879DFFC5EC29FD96FB89FE99FB05:8EFA6050FFC5EC2FFD28FA7FFCDFFA62" box="[758,895,1508,1534]" country="Madagascar" name="SOALALA" pageId="11" pageNumber="87">SOALALA</location>
</taxonomicName>
/ forêt cotière
<date id="FF9B104BFFC5EC2FFBF8FA7FFBDBFA62" box="[1062,1147,1508,1534]" pageId="11" pageNumber="87" value="1973-02">
<collectingDate id="EFDFE9A3FFC5EC2FFBF8FA7FFBDBFA62" box="[1062,1147,1508,1534]" pageId="11" pageNumber="87" value="1973-02">II.1973</collectingDate>
</date>
/ litière A. Peyrieras” [blue, handwritten] // “HOLOTYPE” [red, printed] // “
<taxonomicName id="4C254D08FFC5EC2FFCEAF99CFC52F9BD" authorityName="Dajoz" authorityYear="1982" box="[820,1010,1543,1569]" class="Insecta" family="Staphylinidae" genus="Soalala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="87" phylum="Arthropoda" rank="species" species="spinosa">
<emphasis id="B951EA99FFC5EC2FFCEAF99CFC52F9BD" box="[820,1010,1543,1569]" italics="true" pageId="11" pageNumber="87">Soalala spinosa</emphasis>
</taxonomicName>
/ n. g., n. sp. /Dajoz 1981” [blue, handwritten] //“MUSÉUM PARIS” [blue, printed] // “
<taxonomicName id="4C254D08FFC5EC2FFCBAF9B1FBF3F9D9" authority="Wasm." authorityName="Wasmann" authorityYear="1897" box="[868,1107,1578,1605]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="B951EA99FFC5EC2FFCBAF9B1FC4BF9D8" box="[868,1003,1578,1604]" italics="true" pageId="11" pageNumber="87">Apoderiger</emphasis>
Wasm.
</taxonomicName>
/ =
<taxonomicName id="4C254D08FFC5EC2FFB59F9B1FAE4F9D8" authority="Dajoz" authorityName="Dajoz" authorityYear="1982" box="[1159,1348,1578,1604]" class="Insecta" family="Staphylinidae" genus="Soalala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="B951EA99FFC5EC2FFB59F9B1FB42F9D8" box="[1159,1250,1578,1604]" italics="true" pageId="11" pageNumber="87">Soalala</emphasis>
Dajoz
</taxonomicName>
/
<determinerName id="7E46DA4FFFC5EC2FFA8DF9B1FE85F9FB" pageId="11" pageNumber="87">Cl. Besuchet</determinerName>
/ dét.
<date id="FF9B104BFFC5EC2FFEB5F9D5FE6CF9F4" box="[363,460,1614,1640]" pageId="11" pageNumber="87" value="1988-03">
<collectingDate id="EFDFE9A3FFC5EC2FFEB5F9D5FE6CF9F4" box="[363,460,1614,1640]" pageId="11" pageNumber="87" value="1988-03">III 1988</collectingDate>
</date>
” [white, printed and handwritten];
<collectionCode id="ED34AE4EFFC5EC2FFCB7F9D5FC67F9F4" box="[873,967,1614,1640]" pageId="11" pageNumber="87">MNHN</collectionCode>
.
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="C33F6500FFC5EC29FF63F914FE26FCD5" lastPageId="13" lastPageNumber="89" pageId="11" pageNumber="87" type="description">
<paragraph id="8B9A368BFFC5EC2FFF63F914FEEFF935" blockId="11.[189,335,1679,1705]" box="[189,335,1679,1705]" pageId="11" pageNumber="87">
<heading id="D0D281E7FFC5EC2FFF63F914FEEFF935" bold="true" box="[189,335,1679,1705]" fontSize="11" level="3" pageId="11" pageNumber="87" reason="3">
<emphasis id="B951EA99FFC5EC2FFF63F914FEEFF935" bold="true" box="[189,335,1679,1705]" pageId="11" pageNumber="87">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B9A368BFFC5EC2FFF63F925FAD7F94B" blockId="11.[189,1399,1725,1752]" box="[189,1399,1725,1752]" pageId="11" pageNumber="87">
<smallCapsWord id="8D7CA057FFC5EC2FFF63F925FF5EF94A" baselines="1745,1745" box="[189,254,1726,1752]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Body" pageId="11" pageNumber="87">BODY</smallCapsWord>
. Yellowish-brown, less elongated, 2.30 × as long as wide; body
<quantity id="4CDD9B6EFFC5EC2FFC35F926FBF3F944" box="[1003,1107,1725,1752]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="11" pageNumber="87" unit="mm" value="1.4">1.40 mm</quantity>
long and
<quantity id="4CDD9B6EFFC5EC2FFB19F926FA8FF944" box="[1223,1327,1725,1752]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="87" unit="mm" value="0.6">0.60 mm</quantity>
wide.
</paragraph>
<paragraph id="8B9A368BFFC5EC2FFF63F964FBB2F885" blockId="11.[189,1042,1791,1818]" box="[189,1042,1791,1818]" pageId="11" pageNumber="87">
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. Elongate, 2.25 × as long as wide, about 2.1× as long as pronotum.
</paragraph>
<paragraph id="8B9A368BFFC5EC2FFF63F8DBFAA5F8C6" blockId="11.[189,1285,1856,1883]" box="[189,1285,1856,1883]" pageId="11" pageNumber="87">
<smallCapsWord id="8D7CA057FFC5EC2FFF63F8DBFE94F8C5" baselines="1875,1876" box="[189,308,1856,1882]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Antennae" pageId="11" pageNumber="87">ANTENNAE</smallCapsWord>
. Trimerous,
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long, terminal antennomere strongly curved in distal portion.
</paragraph>
<paragraph id="8B9A368BFFC5EC2FFF63F819FD7DF800" blockId="11.[189,733,1921,1948]" box="[189,733,1921,1948]" pageId="11" pageNumber="87">
<smallCapsWord id="8D7CA057FFC5EC2FFF63F819FE96F806" baselines="1941,1941" box="[189,310,1922,1948]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Pronotum" pageId="11" pageNumber="87">PRONOTUM</smallCapsWord>
. Transverse, 1.54 × as wide as long.
</paragraph>
<caption id="DF5A6603FFC2EC28FF63F8FFFDA7F874" ID-DOI="http://doi.org/10.5281/zenodo.15120446" ID-Zenodo-Dep="15120446" httpUri="https://zenodo.org/record/15120446/files/figure.png" pageId="12" pageNumber="88" startId="12.[189,232,1892,1918]" targetBox="[210,1368,284,1850]" targetPageId="12">
<paragraph id="8B9A368BFFC2EC28FF63F8FFFDA7F874" blockId="12.[187,1399,1892,2025]" pageId="12" pageNumber="88">
<emphasis id="B951EA99FFC2EC28FF63F8FFFEB0F8E3" bold="true" box="[189,272,1892,1919]" pageId="12" pageNumber="88">Fig. 4.</emphasis>
Dorsal and lateral view of aedeagus.
<emphasis id="B951EA99FFC2EC28FD34F8FFFC82F8E2" bold="true" box="[746,802,1892,1918]" pageId="12" pageNumber="88">AB</emphasis>
.
<taxonomicName id="4C254D08FFC2EC28FCEBF8FEFB5AF8E3" authority="Jeannel, 1960" authorityName="Jeannel" authorityYear="1960" box="[821,1274,1893,1919]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="88" phylum="Arthropoda" rank="species" species="torticornis">
<emphasis id="B951EA99FFC2EC28FCEBF8FEFBE5F8E3" box="[821,1093,1893,1919]" italics="true" pageId="12" pageNumber="88">Apoderiger torticornis</emphasis>
<bibRefCitation id="EFB44B7AFFC2EC28FB8CF8FEFB5AF8E3" author="Jeannel R." box="[1106,1274,1893,1919]" pageId="12" pageNumber="88" pagination="31 - 216" refId="ref8172" refString="Jeannel R. 1960. Revision des Pselaphides Malgaches. Bulletin de l'Academie malgache 36: 31-216." type="journal article" year="1960">Jeannel, 1960</bibRefCitation>
</taxonomicName>
, lectotype (MNHN).
<emphasis id="B951EA99FFC2EC28FEE0F813FED6F83E" bold="true" box="[318,374,1928,1954]" pageId="12" pageNumber="88">CD</emphasis>
.
<taxonomicName id="4C254D08FFC2EC28FE54F813FDD0F83E" authorityName="Hlaváč" authorityYear="2025" box="[394,624,1928,1954]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="88" phylum="Arthropoda" rank="species" species="sikorai" status="sp. nov.">
<emphasis id="B951EA99FFC2EC28FE54F813FDD0F83E" box="[394,624,1928,1954]" italics="true" pageId="12" pageNumber="88">Apoderiger sikorai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257E2FFC2EC28FDA2F813FD7EF83E" box="[636,734,1928,1954]" pageId="12" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
, paratype (NMW).
<emphasis id="B951EA99FFC2EC28FC0CF813FBA6F83E" bold="true" box="[978,1030,1928,1954]" pageId="12" pageNumber="88">EF</emphasis>
.
<taxonomicName id="4C254D08FFC2EC28FBC7F813FEA4F859" authority="(Dajoz, 1982)" baseAuthorityName="Dajoz" baseAuthorityYear="1982" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="88" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B951EA99FFC2EC28FBC7F813FAB3F83E" box="[1049,1299,1928,1954]" italics="true" pageId="12" pageNumber="88">Apoderiger spinosus</emphasis>
(
<bibRefCitation id="EFB44B7AFFC2EC28FAF9F813FF5CF859" author="Dajoz R." pageId="12" pageNumber="88" pagination="481 - 522" refId="ref8039" refString="Dajoz R. 1982. Contribution a l'etude des Coleopteres Pselaphidae de Madagascar. Bulletin du Museum national d'Histoire naturelle. Section A, Zoologie, Biologie et Ecologie animales (4): 481-522. https://doi.org/10.5962/p.286051" type="journal article" year="1982">Dajoz, 1982</bibRefCitation>
)
</taxonomicName>
, holotype (MNHN).
<emphasis id="B951EA99FFC2EC28FE2CF830FD8FF859" bold="true" box="[498,559,1963,1989]" pageId="12" pageNumber="88">GH</emphasis>
.
<taxonomicName id="4C254D08FFC2EC28FDE3F830FCB9F859" authorityName="Hlaváč" authorityYear="2025" box="[573,793,1963,1989]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="88" phylum="Arthropoda" rank="species" species="banari" status="sp. nov.">
<emphasis id="B951EA99FFC2EC28FDE3F830FCB9F859" box="[573,793,1963,1989]" italics="true" pageId="12" pageNumber="88">Apoderiger banari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257E2FFC2EC28FCC3F837FCD6F85A" box="[797,886,1964,1990]" pageId="12" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (PCPH).
<emphasis id="B951EA99FFC2EC28FB86F830FB21F859" bold="true" box="[1112,1153,1963,1989]" pageId="12" pageNumber="88">IJ</emphasis>
.
<taxonomicName id="4C254D08FFC2EC28FB51F830FAD6F859" authorityName="Hlaváč" authorityYear="2025" box="[1167,1398,1963,1989]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="88" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B951EA99FFC2EC28FB51F830FAD6F859" box="[1167,1398,1963,1989]" italics="true" pageId="12" pageNumber="88">Apoderiger grandis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257E2FFC2EC28FF65F854FEB8F875" box="[187,280,1999,2025]" pageId="12" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (MHNG).
</paragraph>
</caption>
<paragraph id="8B9A368BFFC3EC29FF63FE92FDE3FEDB" blockId="13.[188,1399,265,327]" pageId="13" pageNumber="89">
<smallCapsWord id="8D7CA057FFC3EC29FF63FE92FEC0FEBE" baselines="284,285" box="[189,352,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Metaventrite" pageId="13" pageNumber="89">METAVENTRITE</smallCapsWord>
. With large median impression, posterior metaventral process long, slender, subtriangular with bunch of about 6 long setae.
</paragraph>
<paragraph id="8B9A368BFFC3EC29FF63FEF5FD0DFE53" blockId="13.[188,1398,365,463]" pageId="13" pageNumber="89">
<smallCapsWord id="8D7CA057FFC3EC29FF63FEF5FEB2FE1A" baselines="385,385" box="[189,274,366,392]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Elytra" pageId="13" pageNumber="89">ELYTRA</smallCapsWord>
. 1.70 × as wide as long, 1.55 × as long as pronotum; lacking sutural and discal striae; humeri strongly projecting; lateral margin with some stout setae; posterior elytral margin in lateral third sinuate, with sharp spine on posterior outer corner.
</paragraph>
<paragraph id="8B9A368BFFC3EC29FF63FE6DFDB5FDAF" blockId="13.[189,1398,501,563]" pageId="13" pageNumber="89">
<smallCapsWord id="8D7CA057FFC3EC29FF63FE6DFE8CFD92" baselines="521,521" box="[189,300,502,528]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Abdomen" pageId="13" pageNumber="89">ABDOMEN</smallCapsWord>
. 1.35 × as long as elytra; composite tergite in posterior portion with pair of small, bump-like, widely separated projections.
</paragraph>
<paragraph id="8B9A368BFFC3EC29FF63FDC0FB21FDE9" blockId="13.[189,1153,602,629]" box="[189,1153,602,629]" pageId="13" pageNumber="89">
<smallCapsWord id="8D7CA057FFC3EC29FF63FDC0FE96FDEF" baselines="622,622" box="[189,310,603,629]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Mesotibia" pageId="13" pageNumber="89">MESOTIBIA</smallCapsWord>
. With large, curved apophysis almost as long as simple mesotarsomere.
</paragraph>
<paragraph id="8B9A368BFFC3EC29FF63FD07FAD7FD45" blockId="13.[189,1399,667,729]" pageId="13" pageNumber="89">
<smallCapsWord id="8D7CA057FFC3EC29FF63FD07FE93FD28" baselines="687,687" box="[189,307,668,694]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Aedeagus" pageId="13" pageNumber="89">AEDEAGUS</smallCapsWord>
(
<figureCitation id="131E2A0EFFC3EC29FE9AFD00FE1FFD2A" box="[324,447,667,694]" captionStart="Fig" captionStartId="12.[189,232,1892,1918]" captionTargetBox="[210,1368,284,1850]" captionTargetId="figure-12@12.[199,1369,269,1854]" captionTargetPageId="12" captionText="Fig. 4. Dorsal and lateral view of aedeagus. AB. Apoderiger torticornis Jeannel, 1960, lectotype (MNHN). CD. Apoderiger sikorai sp. nov., paratype (NMW). EF. Apoderiger spinosus (Dajoz, 1982), holotype (MNHN).GH. Apoderiger banari sp. nov., holotype (PCPH).IJ. Apoderiger grandis sp. nov., holotype (MHNG)." figureDoi="http://doi.org/10.5281/zenodo.15120446" httpUri="https://zenodo.org/record/15120446/files/figure.png" pageId="13" pageNumber="89">Fig. 4EF</figureCitation>
). Elongate, 2.60 × as long as wide; basal bulb and distal lobe well-separated, ratio length of basal bulb / length of distal lobe 1.70; endophallus lacking sclerites; dorsal diaphragm elliptical.
</paragraph>
<paragraph id="8B9A368BFFC3EC29FF63FC9BFE13FC86" blockId="13.[189,435,768,794]" box="[189,435,768,794]" pageId="13" pageNumber="89">
<heading id="D0D281E7FFC3EC29FF63FC9BFE13FC86" bold="true" box="[189,435,768,794]" fontSize="11" level="3" pageId="13" pageNumber="89" reason="3">
<emphasis id="B951EA99FFC3EC29FF63FC9BFE13FC86" bold="true" box="[189,435,768,794]" pageId="13" pageNumber="89">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="8B9A368BFFC3EC29FF63FCB4FE26FCD5" blockId="13.[189,390,815,841]" box="[189,390,815,841]" pageId="13" pageNumber="89">As for the genus.</paragraph>
</subSubSection>
<subSubSection id="C33F6500FFC3EC29FF63FCEBFDBDFC25" box="[189,541,880,953]" pageId="13" pageNumber="89" type="distribution">
<paragraph id="8B9A368BFFC3EC29FF63FCEBFEF8FC16" blockId="13.[189,344,880,906]" box="[189,344,880,906]" pageId="13" pageNumber="89">
<heading id="D0D281E7FFC3EC29FF63FCEBFEF8FC16" bold="true" box="[189,344,880,906]" fontSize="11" level="3" pageId="13" pageNumber="89" reason="3">
<emphasis id="B951EA99FFC3EC29FF63FCEBFEF8FC16" bold="true" box="[189,344,880,906]" pageId="13" pageNumber="89">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B9A368BFFC3EC29FF63FC04FDBDFC25" blockId="13.[189,541,927,953]" box="[189,541,927,953]" pageId="13" pageNumber="89">
<collectingCountry id="F332761BFFC3EC29FF63FC04FEEBFC25" box="[189,331,927,953]" name="Madagascar" pageId="13" pageNumber="89">Madagascar</collectingCountry>
(
<taxonomicName id="4C254D08FFC3EC29FE84FC04FE12FC25" authorityName="Dajoz" authorityYear="1982" box="[346,434,927,953]" class="Insecta" family="Staphylinidae" genus="Soalala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="89" phylum="Arthropoda" rank="genus">Soalala</taxonomicName>
district).
</paragraph>
</subSubSection>
<subSubSection id="C33F6500FFC3EC29FF63FC7BFE59FBB5" box="[189,505,992,1065]" pageId="13" pageNumber="89" type="discussion">
<paragraph id="8B9A368BFFC3EC29FF63FC7BFE90FC66" blockId="13.[189,304,992,1018]" box="[189,304,992,1018]" pageId="13" pageNumber="89">
<heading id="D0D281E7FFC3EC29FF63FC7BFE90FC66" bold="true" box="[189,304,992,1018]" fontSize="11" level="3" pageId="13" pageNumber="89" reason="3">
<emphasis id="B951EA99FFC3EC29FF63FC7BFE90FC66" bold="true" box="[189,304,992,1018]" pageId="13" pageNumber="89">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B9A368BFFC3EC29FF63FB94FE59FBB5" blockId="13.[189,505,1039,1065]" box="[189,505,1039,1065]" pageId="13" pageNumber="89">
See remarks for
<taxonomicName id="4C254D08FFC3EC29FEA0FB94FE51FBB5" authorityName="Hlaváč" authorityYear="2025" box="[382,497,1039,1065]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="89" phylum="Arthropoda" rank="species" species="sikorai">
<emphasis id="B951EA99FFC3EC29FEA0FB94FE51FBB5" box="[382,497,1039,1065]" italics="true" pageId="13" pageNumber="89">A. sikorai</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F6500FFC3EC29FF63FBCBFE99FB05" pageId="13" pageNumber="89" type="biology_ecology">
<paragraph id="8B9A368BFFC3EC29FF63FBCBFE89FBF6" blockId="13.[189,297,1104,1130]" box="[189,297,1104,1130]" pageId="13" pageNumber="89">
<heading id="D0D281E7FFC3EC29FF63FBCBFE89FBF6" bold="true" box="[189,297,1104,1130]" fontSize="11" level="3" pageId="13" pageNumber="89" reason="3">
<emphasis id="B951EA99FFC3EC29FF63FBCBFE89FBF6" bold="true" box="[189,297,1104,1130]" pageId="13" pageNumber="89">Host ant</emphasis>
</heading>
</paragraph>
<paragraph id="8B9A368BFFC3EC29FF63FBE4FE99FB05" blockId="13.[189,313,1151,1177]" box="[189,313,1151,1177]" pageId="13" pageNumber="89">Unknown.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,115 @@
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<mods:title id="48B3EC6C3CA8748D2DAB50509505E995">The Pselaphinae (Coleoptera: Staphylinidae) of Madagascar. VIII. Revision of the myrmecophilous clavigerine genus Apoderiger Wasmann, 1897</mods:title>
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<subSubSection id="C33F6500FFCCEC26FDBAFAF1FC70FA19" box="[612,976,1386,1413]" pageId="2" pageNumber="78" type="nomenclature">
<paragraph id="8B9A368BFFCCEC26FDBAFAF1FC70FA19" blockId="2.[612,976,1386,1413]" box="[612,976,1386,1413]" pageId="2" pageNumber="78">
Tribe
<taxonomicName id="4C254D08FFCCEC26FD74FAF1FC70FA19" ID-CoL="8VR7P" authority="Leach, 1815" authorityName="Leach" authorityYear="1815" box="[682,976,1386,1413]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="78" phylum="Arthropoda" rank="tribe" tribe="Clavigerini">
<emphasis id="B951EA99FFCCEC26FD74FAF1FC98FA18" bold="true" box="[682,824,1386,1412]" pageId="2" pageNumber="78">Clavigerini</emphasis>
Leach, 1815
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C33F6500FFCCEC26FF63FA30FDE5F920" pageId="2" pageNumber="78" type="description">
<paragraph id="8B9A368BFFCCEC26FF63FA30FDE5F920" blockId="2.[187,1399,1451,1724]" pageId="2" pageNumber="78">
All
<taxonomicName id="4C254D08FFCCEC26FF36FA30FED7FA59" authorityName="Leach" authorityYear="1815" box="[232,375,1451,1477]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="78" phylum="Arthropoda" rank="superTribe" superTribe="Clavigeritae">Clavigeritae</taxonomicName>
of
<collectingCountry id="F332761BFFCCEC26FE44FA30FD88FA59" box="[410,552,1451,1477]" name="Madagascar" pageId="2" pageNumber="78">Madagascar</collectingCountry>
are currently assigned to the tribe
<taxonomicName id="4C254D08FFCCEC26FC6FFA30FB95FA59" box="[945,1077,1451,1477]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="78" phylum="Arthropoda" rank="tribe" tribe="Clavigerini">Clavigerini</taxonomicName>
as newly defined by
<bibRefCitation id="EFB44B7AFFCCEC26FAFDFA30FEF1FA75" author="Hlavac P. &amp; Parker J. &amp; Maruyama M. &amp; Fikacek M." pageId="2" pageNumber="78" pagination="422 - 452" refId="ref8089" refString="Hlavac P., Parker J., Maruyama M. &amp; Fikacek M. 2021. Diversification of myrmecophilous Clavigeritae beetles (Coleoptera: Staphylinidae: Pselaphinae) and their radiation in New Caledonia. Systematic Entomology 46: 422-452. https://doi.org/10.1111/syen.12469" type="journal article" year="2021">
Hlaváč
<emphasis id="B951EA99FFCCEC26FF63FA4BFF5AFA75" box="[189,250,1487,1513]" italics="true" pageId="2" pageNumber="78">et al.</emphasis>
(2021)
</bibRefCitation>
. There are apparently four genera in this tribe which resemble each other and are easily separated from others.
<taxonomicName id="4C254D08FFCCEC26FE11FA69FCAEF990" authority="Silvestri, 1926" authorityName="Silvestri" authorityYear="1926" box="[463,782,1522,1548]" class="Insecta" family="Staphylinidae" genus="Ambrosiger" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis id="B951EA99FFCCEC26FE11FA69FDFAF990" box="[463,602,1522,1548]" italics="true" pageId="2" pageNumber="78">Ambrosiger</emphasis>
Silvestri, 1926
</taxonomicName>
(from
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,
<collectingCountry id="F332761BFFCCEC26FC1CFA69FBADF990" box="[962,1037,1522,1548]" name="China" pageId="2" pageNumber="78">China</collectingCountry>
),
<taxonomicName id="4C254D08FFCCEC26FBFDFA69FB09F990" authorityName="Wasmann" authorityYear="1897" box="[1059,1193,1522,1548]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis id="B951EA99FFCCEC26FBFDFA69FB09F990" box="[1059,1193,1522,1548]" italics="true" pageId="2" pageNumber="78">Apoderiger</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D08FFCCEC26FB67FA69FAD6F990" authorityName="Jeannel" authorityYear="1960" box="[1209,1398,1522,1548]" class="Insecta" family="Staphylinidae" genus="Micrapoderiger" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis id="B951EA99FFCCEC26FB67FA69FAD6F990" box="[1209,1398,1522,1548]" italics="true" pageId="2" pageNumber="78">Micrapoderiger</emphasis>
</taxonomicName>
and
<taxonomicName id="4C254D08FFCCEC26FF33F98EFEC4F9B3" authorityName="Fairmaire" authorityYear="1898" box="[237,356,1557,1583]" class="Insecta" family="Staphylinidae" genus="Trymalius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis id="B951EA99FFCCEC26FF33F98EFEC4F9B3" box="[237,356,1557,1583]" italics="true" pageId="2" pageNumber="78">Trymalius</emphasis>
</taxonomicName>
(all three from
<collectingCountry id="F332761BFFCCEC26FDCBF98EFD05F9B3" box="[533,677,1557,1583]" name="Madagascar" pageId="2" pageNumber="78">Madagascar</collectingCountry>
) are all originally placed in the tribe
<taxonomicName id="4C254D08FFCCEC26FB97F98EFE21F9CE" authority="(Newton &amp; Chandler, 1989)" baseAuthorityName="Newton &amp; Chandler" baseAuthorityYear="1989" class="Insecta" family="Staphylinidae" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="78" phylum="Arthropoda" rank="tribe" tribe="Apoderigerini">
Apoderigerini (
<bibRefCitation id="EFB44B7AFFCCEC26FB25F98EFED7F9CE" author="Newton A. F. &amp; Chandler D. S." pageId="2" pageNumber="78" pagination="1 - 93" refId="ref8192" refString="Newton A. F. jr. &amp; Chandler D. S. 1989. World catalog of the genera of Pselaphidae (Coleoptera). Fieldiana: Zoology (N. S.) 53: 1-93. https://doi.org/10.5962/bhl.title.3209" type="journal article" year="1989">Newton &amp; Chandler, 1989</bibRefCitation>
)
</taxonomicName>
. All these genera share a smooth surface of the forebody, trimerous antennae with the terminal antennomere very long and dilated distad to form a distal club, and the composite tergite lacking trichomes.
<taxonomicName id="4C254D08FFCCEC26FE7EF9E4FD8AF905" authorityName="Silvestri" authorityYear="1926" box="[416,554,1663,1689]" class="Insecta" family="Staphylinidae" genus="Ambrosiger" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis id="B951EA99FFCCEC26FE7EF9E4FD8AF905" box="[416,554,1663,1689]" italics="true" pageId="2" pageNumber="78">Ambrosiger</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D08FFCCEC26FDE4F9E4FD57F905" authorityName="Jeannel" authorityYear="1960" box="[570,759,1663,1689]" class="Insecta" family="Staphylinidae" genus="Micrapoderiger" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis id="B951EA99FFCCEC26FDE4F9E4FD57F905" box="[570,759,1663,1689]" italics="true" pageId="2" pageNumber="78">Micrapoderiger</emphasis>
</taxonomicName>
and
<taxonomicName id="4C254D08FFCCEC26FCEDF9E4FC0AF905" authorityName="Fairmaire" authorityYear="1898" box="[819,938,1663,1689]" class="Insecta" family="Staphylinidae" genus="Trymalius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis id="B951EA99FFCCEC26FCEDF9E4FC0AF905" box="[819,938,1663,1689]" italics="true" pageId="2" pageNumber="78">Trymalius</emphasis>
</taxonomicName>
are monogeneric and
<taxonomicName id="4C254D08FFCCEC26FB6CF9E4FA99F905" authorityName="Wasmann" authorityYear="1897" box="[1202,1337,1663,1689]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis id="B951EA99FFCCEC26FB6CF9E4FA99F905" box="[1202,1337,1663,1689]" italics="true" pageId="2" pageNumber="78">Apoderiger</emphasis>
</taxonomicName>
after this revision contains six species.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,105 @@
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Key to the genera of the
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genus group
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1. Pronotum with well-defined, transversal antebasal sulcus ......................
<taxonomicName id="4C254D08FFCCEC26FBEBF883FAD7F8AF" authority="Silvestri, 1926" authorityName="Silvestri" authorityYear="1926" box="[1077,1399,1816,1843]" class="Insecta" family="Staphylinidae" genus="Ambrosiger" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis id="B951EA99FFCCEC26FBEBF883FB64F8AF" bold="true" box="[1077,1220,1816,1843]" italics="true" pageId="2" pageNumber="78">Ambrosiger</emphasis>
Silvestri, 1926
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<keyLead id="30D495BEFFCCEC26FF1FF8A6FAD7F8CA" box="[193,1399,1852,1879]" pageId="2" pageNumber="78"> Pronotum lacking transversal antebasal sulcus ................................................................................. 2</keyLead>
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<keyLead id="30D495BEFFCCEC26FF63F8E5FAD7F804" box="[189,1399,1917,1944]" pageId="2" pageNumber="78">2. Elytra with lateral fold, which is a prolongation of the humeral carina ........................................... 3</keyLead>
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Elytra lacking lateral fold and humeral carina ...................................
<taxonomicName id="4C254D08FFCCEC26FBD1F83BFAD7F827" authority="Jeannel, 1960" authorityName="Jeannel" authorityYear="1960" box="[1039,1399,1952,1979]" class="Insecta" family="Staphylinidae" genus="Micrapoderiger" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis id="B951EA99FFCCEC26FBD1F83BFB6EF827" bold="true" box="[1039,1230,1952,1979]" italics="true" pageId="2" pageNumber="78">Micrapoderiger</emphasis>
<bibRefCitation id="EFB44B7AFFCCEC26FB0BF83AFAD7F827" author="Jeannel R." box="[1237,1399,1953,1979]" pageId="2" pageNumber="78" pagination="31 - 216" refId="ref8172" refString="Jeannel R. 1960. Revision des Pselaphides Malgaches. Bulletin de l'Academie malgache 36: 31-216." type="journal article" year="1960">Jeannel, 1960</bibRefCitation>
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3. Head long, clearly longer than wide; pronotum lacking median antebasal fovea .............................. ................................................................................................................
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Wasmann, 1897
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Head short, at most slightly longer than wide; pronotum with large median antebasal fovea ............ .................................................................................................................
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<emphasis id="B951EA99FFCDEC27FBE9FEE9FB13FE11" bold="true" box="[1079,1203,370,397]" italics="true" pageId="3" pageNumber="79">Trymalius</emphasis>
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<mods:number id="2D15E6CE3EFB5BA054BB1A7C1CFE603E">2025-03-11</mods:number>
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<mods:number id="7BB63E124B5CC74509225BB80D2949AF">981</mods:number>
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<mods:start id="1EF730399AA65978433D156C3A100F75">76</mods:start>
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<mods:identifier id="6BA9BF48C5361214B4DE8D7F24DE168A" type="DOI">10.5852/ejt.2025.981.2811</mods:identifier>
<mods:identifier id="2EBDEA5CF2BCF456C7F3DFEA4450CF89" type="ISSN">2118-9773</mods:identifier>
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<treatment id="038C879DFFCDEC23FD94FE2FFAD7FB35" LSID="urn:lsid:plazi:treatment:038C879DFFCDEC23FD94FE2FFAD7FB35" httpUri="http://treatment.plazi.org/id/038C879DFFCDEC23FD94FE2FFAD7FB35" lastPageId="7" lastPageNumber="83" pageId="3" pageNumber="79">
<subSubSection id="C33F6500FFCDEC27FD94FE2FFC49FE52" box="[586,1001,435,463]" pageId="3" pageNumber="79" type="nomenclature">
<paragraph id="8B9A368BFFCDEC27FD94FE2FFC49FE52" blockId="3.[586,1001,435,463]" box="[586,1001,435,463]" pageId="3" pageNumber="79">
<heading id="D0D281E7FFCDEC27FD94FE2FFC49FE52" box="[586,1001,435,463]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="79" reason="2">
Genus
<taxonomicName id="4C254D08FFCDEC27FD43FE28FC49FE52" ID-CoL="8X73T" authority="Wasmann, 1897" authorityName="Wasmann" authorityYear="1897" box="[669,1001,435,463]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="B951EA99FFCDEC27FD43FE28FC86FE52" bold="true" box="[669,806,435,462]" italics="true" pageId="3" pageNumber="79">Apoderiger</emphasis>
Wasmann, 1897
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F6500FFCDEC27FF63FE6EFDEBFD93" box="[189,587,501,528]" pageId="3" pageNumber="79" type="reference_group">
<paragraph id="8B9A368BFFCDEC27FF63FE6EFDEBFD93" blockId="3.[189,1399,501,599]" box="[189,587,501,528]" pageId="3" pageNumber="79">
<taxonomicName id="4C254D08FFCDEC27FF63FE6EFDE7FD93" authority="Wasmann, 1897: 263" authorityName="Wasmann" authorityPageNumber="263" authorityYear="1897" box="[189,583,501,528]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="B951EA99FFCDEC27FF63FE6EFEE4FD93" box="[189,324,501,527]" italics="true" pageId="3" pageNumber="79">Apoderiger</emphasis>
Wasmann, 1897: 263
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F6500FFCDEC27FD8FFE6DFAB5FD8C" box="[593,1301,501,528]" pageId="3" pageNumber="79" type="type_taxon">
<paragraph id="8B9A368BFFCDEC27FD8FFE6DFAB5FD8C" blockId="3.[189,1399,501,599]" box="[593,1301,501,528]" pageId="3" pageNumber="79">
<typeStatus id="549E8829FFCDEC27FD8FFE6DFD2BFD8C" box="[593,651,502,528]" pageId="3" pageNumber="79">Type</typeStatus>
species:
<taxonomicName id="4C254D08FFCDEC27FD29FE6EFBC8FD8C" authority="Wasmann" authorityName="Wasmann" authorityYear="1897" box="[759,1128,501,528]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="79" phylum="Arthropoda" rank="species" species="cervinus">
<emphasis id="B951EA99FFCDEC27FD29FE6EFC4AFD93" box="[759,1002,501,527]" italics="true" pageId="3" pageNumber="79">Apoderiger cervinus</emphasis>
Wasmann
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="C33F6500FFCDEC27FF63FD82FBECFD27" pageId="3" pageNumber="79" type="reference_group">
<paragraph id="8B9A368BFFCDEC27FF63FD82FE74FDCB" blockId="3.[189,1399,501,599]" pageId="3" pageNumber="79">
<treatmentCitationGroup id="AB3511A5FFCDEC27FF63FD82FE54FDAF" box="[189,500,536,563]" pageId="3" pageNumber="79">
<treatmentCitation id="0A84109AFFCDEC27FF63FD82FE50FDAF" author="Dajoz R." box="[189,496,536,563]" page="512" pageId="3" pageNumber="79" year="1982">
<taxonomicName id="4C254D08FFCDEC27FF63FD82FE50FDAF" ID-CoL="8XPWY" authority="Dajoz, 1982: 512" authorityName="Dajoz" authorityPageNumber="512" authorityYear="1982" box="[189,496,536,563]" class="Insecta" family="Staphylinidae" genus="Soalala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="B951EA99FFCDEC27FF63FD82FEB8FDAF" box="[189,280,537,563]" italics="true" pageId="3" pageNumber="79">Soalala</emphasis>
<bibRefCitation id="EFB44B7AFFCDEC27FEFEFD82FE50FDAF" author="Dajoz R." box="[288,496,536,563]" pageId="3" pageNumber="79" pagination="481 - 522" refId="ref8039" refString="Dajoz R. 1982. Contribution a l'etude des Coleopteres Pselaphidae de Madagascar. Bulletin du Museum national d'Histoire naturelle. Section A, Zoologie, Biologie et Ecologie animales (4): 481-522. https://doi.org/10.5962/p.286051" type="journal article" year="1982">Dajoz, 1982: 512</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
<treatmentCitationGroup id="AB3511A5FFCDEC27FE25FD82FE74FDCB" pageId="3" pageNumber="79">
<typeStatus id="549E8829FFCDEC27FE25FD82FD95FDAF" box="[507,565,537,563]" pageId="3" pageNumber="79">Type</typeStatus>
species:
<taxonomicName id="4C254D08FFCDEC27FD7DFD82FC12FDAF" ID-CoL="6YTR3" authority="Dajoz" authorityName="Dajoz" authorityYear="1982" box="[675,946,537,563]" class="Insecta" family="Staphylinidae" genus="Soalala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="79" phylum="Arthropoda" rank="species" species="spinosa">
<emphasis id="B951EA99FFCDEC27FD7DFD82FCC1FDAE" box="[675,865,537,563]" italics="true" pageId="3" pageNumber="79">Soalala spinosa</emphasis>
Dajoz
</taxonomicName>
, by original designation; synonymy in
<treatmentCitation id="0A84109AFFCDEC27FF35FDA7FE70FDCB" author="Besuchet C." box="[235,464,572,599]" page="65" pageId="3" pageNumber="79" year="2008">
<bibRefCitation id="EFB44B7AFFCDEC27FF35FDA7FE70FDCB" author="Besuchet C." box="[235,464,572,599]" pageId="3" pageNumber="79" pagination="1 - 2" refId="ref7940" refString="Besuchet C. 2008. Synonymies et combinasions nouvelles, revalidations et description de taxa nouveaux de Pselaphinae (Coleoptera: Staphylinidae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft 81 (1-2): 61 - 82." type="journal article" year="2008">Besuchet, 2008: 65</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A368BFFCDEC27FF63FDE6FBECFD27" blockId="3.[189,1398,637,699]" pageId="3" pageNumber="79">
<treatmentCitationGroup id="AB3511A5FFCDEC27FF63FDE6FD88FD26" pageId="3" pageNumber="79">
<taxonomicName id="4C254D08FFCDEC27FF63FDE6FEE4FD0B" ID-CoL="8X73T" authorityName="Wasmann" authorityYear="1897" box="[189,324,637,663]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="B951EA99FFCDEC27FF63FDE6FEE4FD0B" box="[189,324,637,663]" italics="true" pageId="3" pageNumber="79">Apoderiger</emphasis>
</taxonomicName>
<treatmentCitation id="0A84109AFFCDEC27FEB6FDE6FDE4FD0B" author="Jeannel R." box="[360,580,637,664]" page="312" pageId="3" pageNumber="79" year="1954">
<bibRefCitation id="EFB44B7AFFCDEC27FEB6FDE6FDE4FD0B" author="Jeannel R." box="[360,580,637,664]" pageId="3" pageNumber="79" pagination="139 - 344" refId="ref8145" refString="Jeannel R. 1954. Les Pselaphides de Madagascar. Memoires de l'Institut scientifique de Madagascar (E: Entomologie) 4: 139-344." type="journal article" year="1954">Jeannel 1954: 312</bibRefCitation>
</treatmentCitation>
(redescription); 1960: 202 (in key); 1960: 203. —
<treatmentCitation id="0A84109AFFCDEC27FB68FDE6FAD6FD0B" author="Celis M. J." box="[1206,1398,637,663]" page="248" pageId="3" pageNumber="79" year="1970">
<bibRefCitation id="EFB44B7AFFCDEC27FB68FDE6FAD6FD0B" author="Celis M. J." box="[1206,1398,637,663]" pageId="3" pageNumber="79" pagination="237 - 269" refId="ref7980" refString="Celis M. J. 1970. Contribution a l'etude des Clavigerinae de Madagascar (Coleoptera Pselaphidae). Clavigerinae recueillis par le Professeur Dr. H. Franz et par M. J. Vadon. Remaniements apportes a la systematique des Clavigerinae malgaches. Revue de Zoologie et de Botanique africaines 82: 237-269." type="journal article" year="1970">Célis 1970: 248</bibRefCitation>
</treatmentCitation>
(diagnosis, key to species).
</treatmentCitationGroup>
<treatmentCitationGroup id="AB3511A5FFCDEC27FDF1FD3AFBECFD27" box="[559,1100,672,699]" pageId="3" pageNumber="79">
<treatmentCitation id="0A84109AFFCDEC27FD8DFD3BFC8DFD27" author="Besuchet C." box="[595,813,672,699]" page="65" pageId="3" pageNumber="79" year="2008">
<bibRefCitation id="EFB44B7AFFCDEC27FD8DFD3BFC8DFD27" author="Besuchet C." box="[595,813,672,699]" pageId="3" pageNumber="79" pagination="1 - 2" refId="ref7940" refString="Besuchet C. 2008. Synonymies et combinasions nouvelles, revalidations et description de taxa nouveaux de Pselaphinae (Coleoptera: Staphylinidae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft 81 (1-2): 61 - 82." type="journal article" year="2008">Besuchet 2008: 65</bibRefCitation>
</treatmentCitation>
(synonymy of
<taxonomicName id="4C254D08FFCDEC27FC3EFD3BFB9BFD26" ID-CoL="8XPWY" authorityName="Dajoz" authorityYear="1982" box="[992,1083,672,698]" class="Insecta" family="Staphylinidae" genus="Soalala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="B951EA99FFCDEC27FC3EFD3BFB9BFD26" box="[992,1083,672,698]" italics="true" pageId="3" pageNumber="79">Soalala</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33F6500FFCDEC27FF63FD79FAC8FB4D" pageId="3" pageNumber="79" type="diagnosis">
<paragraph id="8B9A368BFFCDEC27FF63FD79FE97FD60" blockId="3.[189,311,738,764]" box="[189,311,738,764]" pageId="3" pageNumber="79">
<heading id="D0D281E7FFCDEC27FF63FD79FE97FD60" bold="true" box="[189,311,738,764]" fontSize="11" level="3" pageId="3" pageNumber="79" reason="3">
<emphasis id="B951EA99FFCDEC27FF63FD79FE97FD60" bold="true" box="[189,311,738,764]" pageId="3" pageNumber="79">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B9A368BFFCDEC27FF63FC8BFAC8FB4D" blockId="3.[188,1399,784,1233]" pageId="3" pageNumber="79">Head elongate, at least twice as long as wide; temples shorter than frontal rostrum; lacking frontal foveae, with pair of minute dorsal tentorial pits, situated behind eyes. Neck region subcylindrical, long, about as long as or slightly shorter than posterior part of head capsule, separated from head capsule by deep occipital constriction, partly retracted into prothorax. Antenna trimerous; terminal antennomere very long, dilated and curved distally, its apex truncate, with dense short and long setae. Pronotum transverse, clearly wider than long, lacking antebasal and lateral foveae. Elytra more than 1.50 × as wide as long; lacking basal foveae; humeri strongly prominent; posterior margin of elytron lacking trichomes. Abdomen with composite tergite convex, lacking impressions, keels and lacking trichomes; three paratergites of composite tergite present, first paratergite in anterior portion with trichome; posterior margin of composite tergite with pair of small, bump-like, wide-distant or with pair of long, triangular, short-distant projections. Legs long and slender; femur lacking interlocking ridge; mesotibia differently modified in pre or apical portion. Aedeagus elongate, slender, 2.53.0× as long as wide; basal bulb well-separated from distal lobe; endophallus with or lacking sclerites; dorsal diaphragm round or elliptical.</paragraph>
</subSubSection>
<subSubSection id="C33F6500FFCDEC22FF63FB63FE16FB76" lastPageId="6" lastPageNumber="82" pageId="3" pageNumber="79" type="description">
<paragraph id="8B9A368BFFCDEC27FF63FB63FEEFFA8E" blockId="3.[189,335,1272,1298]" box="[189,335,1272,1298]" pageId="3" pageNumber="79">
<heading id="D0D281E7FFCDEC27FF63FB63FEEFFA8E" bold="true" box="[189,335,1272,1298]" fontSize="11" level="3" pageId="3" pageNumber="79" reason="3">
<emphasis id="B951EA99FFCDEC27FF63FB63FEEFFA8E" bold="true" box="[189,335,1272,1298]" pageId="3" pageNumber="79">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B9A368BFFCDEC27FF63FABCFA9CFADD" blockId="3.[189,1340,1319,1346]" box="[189,1340,1319,1346]" pageId="3" pageNumber="79">
<smallCapsWord id="8D7CA057FFCDEC27FF63FABCFF5EFAA3" baselines="1338,1338" box="[189,254,1319,1345]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Body" pageId="3" pageNumber="79">BODY</smallCapsWord>
(
<figureCitation id="131E2A0EFFCDEC27FED1FABCFEEBFADD" box="[271,331,1319,1345]" captionStart="Fig" captionStartId="4.[191,234,1961,1987]" captionTargetBox="[291,1382,1113,1903]" captionTargetId="figure-263@4.[189,1398,1090,1923]" captionTargetPageId="4" captionText="Fig. 1.Dorsal habitus of male.A. Apoderiger torticornisJeannel, 1960,holotype (MNHN).B. Apoderiger banari sp. nov., holotype (PCPH)." figureDoi="http://doi.org/10.5281/zenodo.15120440" httpUri="https://zenodo.org/record/15120440/files/figure.png" pageId="3" pageNumber="79">Fig.1</figureCitation>
). Yellowish-brown, elongated, more about 2.5 × as long as wide. Length
<quantity id="4CDD9B6EFFCDEC27FB76FABCFA98FADD" box="[1192,1336,1319,1345]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" metricValueMax="2.2" metricValueMin="1.4" pageId="3" pageNumber="79" unit="mm" value="1.8" valueMax="2.2" valueMin="1.4">1.42.2 mm</quantity>
.
</paragraph>
<paragraph id="8B9A368BFFCDEC27FF63FAF2FB85F900" blockId="3.[187,1401,1384,1692]" pageId="3" pageNumber="79">
<smallCapsWord id="8D7CA057FFCDEC27FF63FAF2FF5CFA1D" baselines="1404,1404" box="[189,252,1385,1411]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="3" pageNumber="79">HEAD</smallCapsWord>
. Elongate, at least twice as long as wide, widest at level of eyes; rostrum narrow, rounded at anterior margin; eyes oval, slightly prominent, situated slightly behind midpoint of head capsule; temples shorter than frontal rostrum, behind eyes slightly convergent posteriad; lacking frontal foveae, with pair of minute dorsal tentorial pits, situated behind eyes; clypeus partly visible on sides. Neck region subcylindrical, long, about as long as or slightly shorter than posterior part of head capsule, separated from head capsule by deep occipital constriction, partly retracted into prothorax, with well-defined gular plate (
<figureCitation id="131E2A0EFFCDEC27FE8FF9A0FE13F9CA" box="[337,435,1595,1622]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[200,1387,265,1853]" captionTargetId="figure-13@5.[200,1388,265,1854]" captionTargetPageId="5" captionText="Fig. 2. Apoderiger sikorai sp. nov., paratype, ♂ (PCPH). A. Head in ventral view. B. Mouth part. CD. Antenna.E. Apoderiger torticornis Jeannel, 1960, holotype, ♂ (MNHN), antenna. Abbrevations: bst = basistipes; cd = cardo; cl = clypeus; gp = gular plate; lb = labrum; md = mandible; mn = mentum; mxp = maxillary palpi; ps = peg-like sensilla; ptp = posterior tentorisl pits." figureDoi="http://doi.org/10.5281/zenodo.15120442" httpUri="https://zenodo.org/record/15120442/files/figure.png" pageId="3" pageNumber="79">Fig. 2A</figureCitation>
); posterior tentorial pits (
<figureCitation id="131E2A0EFFCDEC27FD37F9A0FCEBF9CA" box="[745,843,1595,1622]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[200,1387,265,1853]" captionTargetId="figure-13@5.[200,1388,265,1854]" captionTargetPageId="5" captionText="Fig. 2. Apoderiger sikorai sp. nov., paratype, ♂ (PCPH). A. Head in ventral view. B. Mouth part. CD. Antenna.E. Apoderiger torticornis Jeannel, 1960, holotype, ♂ (MNHN), antenna. Abbrevations: bst = basistipes; cd = cardo; cl = clypeus; gp = gular plate; lb = labrum; md = mandible; mn = mentum; mxp = maxillary palpi; ps = peg-like sensilla; ptp = posterior tentorisl pits." figureDoi="http://doi.org/10.5281/zenodo.15120442" httpUri="https://zenodo.org/record/15120442/files/figure.png" pageId="3" pageNumber="79">Fig. 2A</figureCitation>
;
<emphasis id="B951EA99FFCDEC27FC84F9A7FC20F9C9" box="[858,896,1596,1621]" italics="true" pageId="3" pageNumber="79">ptp</emphasis>
= gular foveae of Chandler 2001) small, circular, well-separated, situated in front of transverse impression demarcating neck region ventrally; pre-tentorial gular-submental region elongate, subrectangular and smooth.
</paragraph>
<paragraph id="8B9A368BFFCDEC27FF63F958FCC6F8DB" blockId="3.[189,1399,1731,1863]" pageId="3" pageNumber="79">
<smallCapsWord id="8D7CA057FFCDEC27FF63F958FE87F940" baselines="1750,1751" box="[189,295,1731,1757]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Antenna" pageId="3" pageNumber="79">ANTENNA</smallCapsWord>
. Trimerous (
<figureCitation id="131E2A0EFFCDEC27FE61F958FD9CF941" box="[447,572,1731,1757]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[200,1387,265,1853]" captionTargetId="figure-13@5.[200,1388,265,1854]" captionTargetPageId="5" captionText="Fig. 2. Apoderiger sikorai sp. nov., paratype, ♂ (PCPH). A. Head in ventral view. B. Mouth part. CD. Antenna.E. Apoderiger torticornis Jeannel, 1960, holotype, ♂ (MNHN), antenna. Abbrevations: bst = basistipes; cd = cardo; cl = clypeus; gp = gular plate; lb = labrum; md = mandible; mn = mentum; mxp = maxillary palpi; ps = peg-like sensilla; ptp = posterior tentorisl pits." figureDoi="http://doi.org/10.5281/zenodo.15120442" httpUri="https://zenodo.org/record/15120442/files/figure.png" pageId="3" pageNumber="79">Fig. 2CE</figureCitation>
), semi-circular in cross-section, very long, exceeding posterior margin of pronotum; scape slightly longer than pedicel, both small, subrectangular; terminal antennomere (flagellomere) very long, more than 10 × as long as wide, dilated distad and variably shaped, curved distally, its apex truncate, with dense short and long setae.
</paragraph>
<paragraph id="8B9A368BFFCDEC20FF63F8F4FDD3FE2C" blockId="3.[189,1399,1902,1999]" lastBlockId="4.[187,1401,265,432]" lastPageId="4" lastPageNumber="80" pageId="3" pageNumber="79">
<smallCapsWord id="8D7CA057FFCDEC27FF63F8F4FEEEF81B" baselines="1922,1922" box="[189,334,1903,1929]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Mouthparts" pageId="3" pageNumber="79">MOUTHPARTS</smallCapsWord>
. Rudimentary but well-developed, completely hidden in transversally elliptical bucal cavity (
<figureCitation id="131E2A0EFFCDEC27FF18F80AFEE7F830" box="[198,327,1937,1964]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[200,1387,265,1853]" captionTargetId="figure-13@5.[200,1388,265,1854]" captionTargetPageId="5" captionText="Fig. 2. Apoderiger sikorai sp. nov., paratype, ♂ (PCPH). A. Head in ventral view. B. Mouth part. CD. Antenna.E. Apoderiger torticornis Jeannel, 1960, holotype, ♂ (MNHN), antenna. Abbrevations: bst = basistipes; cd = cardo; cl = clypeus; gp = gular plate; lb = labrum; md = mandible; mn = mentum; mxp = maxillary palpi; ps = peg-like sensilla; ptp = posterior tentorisl pits." figureDoi="http://doi.org/10.5281/zenodo.15120442" httpUri="https://zenodo.org/record/15120442/files/figure.png" pageId="3" pageNumber="79">Fig. 2AB</figureCitation>
); labium with transverse, sub-rectangular mentum (
<figureCitation id="131E2A0EFFCDEC27FC7AF80AFB85F830" box="[932,1061,1937,1964]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[200,1387,265,1853]" captionTargetId="figure-13@5.[200,1388,265,1854]" captionTargetPageId="5" captionText="Fig. 2. Apoderiger sikorai sp. nov., paratype, ♂ (PCPH). A. Head in ventral view. B. Mouth part. CD. Antenna.E. Apoderiger torticornis Jeannel, 1960, holotype, ♂ (MNHN), antenna. Abbrevations: bst = basistipes; cd = cardo; cl = clypeus; gp = gular plate; lb = labrum; md = mandible; mn = mentum; mxp = maxillary palpi; ps = peg-like sensilla; ptp = posterior tentorisl pits." figureDoi="http://doi.org/10.5281/zenodo.15120442" httpUri="https://zenodo.org/record/15120442/files/figure.png" pageId="3" pageNumber="79">Fig. 2AB</figureCitation>
), anterior margin of mentum slightly rounded, its anterior half with long setae placed in two rows, lacking ligula. Maxilla reduced, with large, hemispherical cardo (
<figureCitation id="131E2A0EFFCAEC20FDE9FE92FD31FEBF" box="[567,657,265,291]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[200,1387,265,1853]" captionTargetId="figure-13@5.[200,1388,265,1854]" captionTargetPageId="5" captionText="Fig. 2. Apoderiger sikorai sp. nov., paratype, ♂ (PCPH). A. Head in ventral view. B. Mouth part. CD. Antenna.E. Apoderiger torticornis Jeannel, 1960, holotype, ♂ (MNHN), antenna. Abbrevations: bst = basistipes; cd = cardo; cl = clypeus; gp = gular plate; lb = labrum; md = mandible; mn = mentum; mxp = maxillary palpi; ps = peg-like sensilla; ptp = posterior tentorisl pits." figureDoi="http://doi.org/10.5281/zenodo.15120442" httpUri="https://zenodo.org/record/15120442/files/figure.png" pageId="4" pageNumber="80">Fig. 2B</figureCitation>
); basistipes (
<figureCitation id="131E2A0EFFCAEC20FCFBFE92FCDDFEBF" box="[805,893,265,291]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[200,1387,265,1853]" captionTargetId="figure-13@5.[200,1388,265,1854]" captionTargetPageId="5" captionText="Fig. 2. Apoderiger sikorai sp. nov., paratype, ♂ (PCPH). A. Head in ventral view. B. Mouth part. CD. Antenna.E. Apoderiger torticornis Jeannel, 1960, holotype, ♂ (MNHN), antenna. Abbrevations: bst = basistipes; cd = cardo; cl = clypeus; gp = gular plate; lb = labrum; md = mandible; mn = mentum; mxp = maxillary palpi; ps = peg-like sensilla; ptp = posterior tentorisl pits." figureDoi="http://doi.org/10.5281/zenodo.15120442" httpUri="https://zenodo.org/record/15120442/files/figure.png" pageId="4" pageNumber="80">Fig. 2B</figureCitation>
) minuscule, with one seta; galea and lacinia atrophied. Maxillary palpi (
<figureCitation id="131E2A0EFFCAEC20FE22FEB7FDF7FEDA" box="[508,599,300,326]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[200,1387,265,1853]" captionTargetId="figure-13@5.[200,1388,265,1854]" captionTargetPageId="5" captionText="Fig. 2. Apoderiger sikorai sp. nov., paratype, ♂ (PCPH). A. Head in ventral view. B. Mouth part. CD. Antenna.E. Apoderiger torticornis Jeannel, 1960, holotype, ♂ (MNHN), antenna. Abbrevations: bst = basistipes; cd = cardo; cl = clypeus; gp = gular plate; lb = labrum; md = mandible; mn = mentum; mxp = maxillary palpi; ps = peg-like sensilla; ptp = posterior tentorisl pits." figureDoi="http://doi.org/10.5281/zenodo.15120442" httpUri="https://zenodo.org/record/15120442/files/figure.png" pageId="4" pageNumber="80">Fig. 2B</figureCitation>
) reduced to one palpomere. Mandibles (
<figureCitation id="131E2A0EFFCAEC20FBF2FEB7FB25FEDA" box="[1068,1157,300,326]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[200,1387,265,1853]" captionTargetId="figure-13@5.[200,1388,265,1854]" captionTargetPageId="5" captionText="Fig. 2. Apoderiger sikorai sp. nov., paratype, ♂ (PCPH). A. Head in ventral view. B. Mouth part. CD. Antenna.E. Apoderiger torticornis Jeannel, 1960, holotype, ♂ (MNHN), antenna. Abbrevations: bst = basistipes; cd = cardo; cl = clypeus; gp = gular plate; lb = labrum; md = mandible; mn = mentum; mxp = maxillary palpi; ps = peg-like sensilla; ptp = posterior tentorisl pits." figureDoi="http://doi.org/10.5281/zenodo.15120442" httpUri="https://zenodo.org/record/15120442/files/figure.png" pageId="4" pageNumber="80">Fig. 2B</figureCitation>
) ovaly projecting on outer portion. Labrum (
<figureCitation id="131E2A0EFFCAEC20FE0BFED4FD92FEF5" box="[469,562,335,361]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[200,1387,265,1853]" captionTargetId="figure-13@5.[200,1388,265,1854]" captionTargetPageId="5" captionText="Fig. 2. Apoderiger sikorai sp. nov., paratype, ♂ (PCPH). A. Head in ventral view. B. Mouth part. CD. Antenna.E. Apoderiger torticornis Jeannel, 1960, holotype, ♂ (MNHN), antenna. Abbrevations: bst = basistipes; cd = cardo; cl = clypeus; gp = gular plate; lb = labrum; md = mandible; mn = mentum; mxp = maxillary palpi; ps = peg-like sensilla; ptp = posterior tentorisl pits." figureDoi="http://doi.org/10.5281/zenodo.15120442" httpUri="https://zenodo.org/record/15120442/files/figure.png" pageId="4" pageNumber="80">Fig. 2B</figureCitation>
) transverse with almost straight anterior margin; anteroventral margin with some peg-like sensilla (
<figureCitation id="131E2A0EFFCAEC20FDD0FEE9FDCAFE11" box="[526,618,370,397]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[200,1387,265,1853]" captionTargetId="figure-13@5.[200,1388,265,1854]" captionTargetPageId="5" captionText="Fig. 2. Apoderiger sikorai sp. nov., paratype, ♂ (PCPH). A. Head in ventral view. B. Mouth part. CD. Antenna.E. Apoderiger torticornis Jeannel, 1960, holotype, ♂ (MNHN), antenna. Abbrevations: bst = basistipes; cd = cardo; cl = clypeus; gp = gular plate; lb = labrum; md = mandible; mn = mentum; mxp = maxillary palpi; ps = peg-like sensilla; ptp = posterior tentorisl pits." figureDoi="http://doi.org/10.5281/zenodo.15120442" httpUri="https://zenodo.org/record/15120442/files/figure.png" pageId="4" pageNumber="80">Fig. 2B</figureCitation>
); clypeus (
<figureCitation id="131E2A0EFFCAEC20FD37FEE9FCE3FE11" box="[745,835,370,397]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[200,1387,265,1853]" captionTargetId="figure-13@5.[200,1388,265,1854]" captionTargetPageId="5" captionText="Fig. 2. Apoderiger sikorai sp. nov., paratype, ♂ (PCPH). A. Head in ventral view. B. Mouth part. CD. Antenna.E. Apoderiger torticornis Jeannel, 1960, holotype, ♂ (MNHN), antenna. Abbrevations: bst = basistipes; cd = cardo; cl = clypeus; gp = gular plate; lb = labrum; md = mandible; mn = mentum; mxp = maxillary palpi; ps = peg-like sensilla; ptp = posterior tentorisl pits." figureDoi="http://doi.org/10.5281/zenodo.15120442" httpUri="https://zenodo.org/record/15120442/files/figure.png" pageId="4" pageNumber="80">Fig. 2B</figureCitation>
) large, transverse, with rounded anterior margin which is bearing bunch of long setae.
</paragraph>
<paragraph id="8B9A368BFFCAEC20FF63FE43FE77FDA4" blockId="4.[189,1399,471,568]" pageId="4" pageNumber="80">
<smallCapsWord id="8D7CA057FFCAEC20FF63FE43FE96FE6C" baselines="491,491" box="[189,310,472,498]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Pronotum" pageId="4" pageNumber="80">PRONOTUM</smallCapsWord>
. Transverse, clearly wider than long, about half of length of head; lacking squamouse setae; lateral margin rounded, in anterior and posterior half convergent; lacking antebasal and lateral foveae; lacking sulci or carinae.
</paragraph>
<paragraph id="8B9A368BFFCAEC20FF63FDFBFA97FCB5" blockId="4.[187,1400,607,810]" pageId="4" pageNumber="80">
<smallCapsWord id="8D7CA057FFCAEC20FF63FDFBFEEDFDE4" baselines="627,627" box="[189,333,608,634]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Prosternum" pageId="4" pageNumber="80">PROSTERNUM</smallCapsWord>
(
<figureCitation id="131E2A0EFFCAEC20FE80FDC4FE1BFDE5" box="[350,443,607,634]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[189,1398,373,1784]" captionTargetId="figure-11@10.[189,1398,373,1784]" captionTargetPageId="10" captionText="Fig. 3. Apoderiger sikoraisp. nov., paratype,♂ (PCPH).A. Prosternum and metaventrite.B. Metaventrite and abdomen. C. Distal part of tibia and tarsus. Abbrevations: amtp = anterior metaventral process; ihy = inner hypomeral region; hyr = hypomeral ridges; hyg = hypomeral groove; lpcf = lateral procoxal fovea; msvp = mesoventral process; mtvp = posterior metaventral process; ohy = outer hypomeral region; psp = prosternal process; pst = prosternum; sbl = tarsal subapical tooth; tpat = tarsal preapical tooth; IIIIX = visible tergites." figureDoi="http://doi.org/10.5281/zenodo.15120444" httpUri="https://zenodo.org/record/15120444/files/figure.png" pageId="4" pageNumber="80">Fig. 3A</figureCitation>
). Laterally fused with hypomera; basisternal region longer than coxal region, with sparse setae, with pair of small, widely separated asetose lateral procoxal foveae (
<figureCitation id="131E2A0EFFCAEC20FB58FD19FB42FD00" box="[1158,1250,642,668]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[189,1398,373,1784]" captionTargetId="figure-11@10.[189,1398,373,1784]" captionTargetPageId="10" captionText="Fig. 3. Apoderiger sikoraisp. nov., paratype,♂ (PCPH).A. Prosternum and metaventrite.B. Metaventrite and abdomen. C. Distal part of tibia and tarsus. Abbrevations: amtp = anterior metaventral process; ihy = inner hypomeral region; hyr = hypomeral ridges; hyg = hypomeral groove; lpcf = lateral procoxal fovea; msvp = mesoventral process; mtvp = posterior metaventral process; ohy = outer hypomeral region; psp = prosternal process; pst = prosternum; sbl = tarsal subapical tooth; tpat = tarsal preapical tooth; IIIIX = visible tergites." figureDoi="http://doi.org/10.5281/zenodo.15120444" httpUri="https://zenodo.org/record/15120444/files/figure.png" pageId="4" pageNumber="80">Fig. 3A</figureCitation>
;
<emphasis id="B951EA99FFCAEC20FB2DFD19FABFFD00" box="[1267,1311,642,668]" italics="true" pageId="4" pageNumber="80">lpcf</emphasis>
= term of Chandler 2001; in fact, these foveae are situated on prosternum, anterad procoxal cavities), median procoxal fovea absent; prosternal process (
<figureCitation id="131E2A0EFFCAEC20FD77FD52FCA3FD7F" box="[681,771,713,739]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[189,1398,373,1784]" captionTargetId="figure-11@10.[189,1398,373,1784]" captionTargetPageId="10" captionText="Fig. 3. Apoderiger sikoraisp. nov., paratype,♂ (PCPH).A. Prosternum and metaventrite.B. Metaventrite and abdomen. C. Distal part of tibia and tarsus. Abbrevations: amtp = anterior metaventral process; ihy = inner hypomeral region; hyr = hypomeral ridges; hyg = hypomeral groove; lpcf = lateral procoxal fovea; msvp = mesoventral process; mtvp = posterior metaventral process; ohy = outer hypomeral region; psp = prosternal process; pst = prosternum; sbl = tarsal subapical tooth; tpat = tarsal preapical tooth; IIIIX = visible tergites." figureDoi="http://doi.org/10.5281/zenodo.15120444" httpUri="https://zenodo.org/record/15120444/files/figure.png" pageId="4" pageNumber="80">Fig. 3A</figureCitation>
) rounded, not separating procoxae. Hypomera divided by incomplete hypomeral ridges (
<figureCitation id="131E2A0EFFCAEC20FD97FD77FD05FC9A" box="[585,677,748,774]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[189,1398,373,1784]" captionTargetId="figure-11@10.[189,1398,373,1784]" captionTargetPageId="10" captionText="Fig. 3. Apoderiger sikoraisp. nov., paratype,♂ (PCPH).A. Prosternum and metaventrite.B. Metaventrite and abdomen. C. Distal part of tibia and tarsus. Abbrevations: amtp = anterior metaventral process; ihy = inner hypomeral region; hyr = hypomeral ridges; hyg = hypomeral groove; lpcf = lateral procoxal fovea; msvp = mesoventral process; mtvp = posterior metaventral process; ohy = outer hypomeral region; psp = prosternal process; pst = prosternum; sbl = tarsal subapical tooth; tpat = tarsal preapical tooth; IIIIX = visible tergites." figureDoi="http://doi.org/10.5281/zenodo.15120444" httpUri="https://zenodo.org/record/15120444/files/figure.png" pageId="4" pageNumber="80">Fig. 3A</figureCitation>
) into narrow, elongate inner hypomeral region (
<figureCitation id="131E2A0EFFCAEC20FB03FD77FA9AFC9A" box="[1245,1338,748,774]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[189,1398,373,1784]" captionTargetId="figure-11@10.[189,1398,373,1784]" captionTargetPageId="10" captionText="Fig. 3. Apoderiger sikoraisp. nov., paratype,♂ (PCPH).A. Prosternum and metaventrite.B. Metaventrite and abdomen. C. Distal part of tibia and tarsus. Abbrevations: amtp = anterior metaventral process; ihy = inner hypomeral region; hyr = hypomeral ridges; hyg = hypomeral groove; lpcf = lateral procoxal fovea; msvp = mesoventral process; mtvp = posterior metaventral process; ohy = outer hypomeral region; psp = prosternal process; pst = prosternum; sbl = tarsal subapical tooth; tpat = tarsal preapical tooth; IIIIX = visible tergites." figureDoi="http://doi.org/10.5281/zenodo.15120444" httpUri="https://zenodo.org/record/15120444/files/figure.png" pageId="4" pageNumber="80">Fig. 3A</figureCitation>
) and broad, smooth outer hypomeral region (
<figureCitation id="131E2A0EFFCAEC20FD4EFC94FD4EFCB5" box="[656,750,783,809]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[189,1398,373,1784]" captionTargetId="figure-11@10.[189,1398,373,1784]" captionTargetPageId="10" captionText="Fig. 3. Apoderiger sikoraisp. nov., paratype,♂ (PCPH).A. Prosternum and metaventrite.B. Metaventrite and abdomen. C. Distal part of tibia and tarsus. Abbrevations: amtp = anterior metaventral process; ihy = inner hypomeral region; hyr = hypomeral ridges; hyg = hypomeral groove; lpcf = lateral procoxal fovea; msvp = mesoventral process; mtvp = posterior metaventral process; ohy = outer hypomeral region; psp = prosternal process; pst = prosternum; sbl = tarsal subapical tooth; tpat = tarsal preapical tooth; IIIIX = visible tergites." figureDoi="http://doi.org/10.5281/zenodo.15120444" httpUri="https://zenodo.org/record/15120444/files/figure.png" pageId="4" pageNumber="80">Fig. 3A</figureCitation>
), latter with shallow hypomeral groove (
<figureCitation id="131E2A0EFFCAEC20FB15FC94FA89FCB5" box="[1227,1321,783,809]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[189,1398,373,1784]" captionTargetId="figure-11@10.[189,1398,373,1784]" captionTargetPageId="10" captionText="Fig. 3. Apoderiger sikoraisp. nov., paratype,♂ (PCPH).A. Prosternum and metaventrite.B. Metaventrite and abdomen. C. Distal part of tibia and tarsus. Abbrevations: amtp = anterior metaventral process; ihy = inner hypomeral region; hyr = hypomeral ridges; hyg = hypomeral groove; lpcf = lateral procoxal fovea; msvp = mesoventral process; mtvp = posterior metaventral process; ohy = outer hypomeral region; psp = prosternal process; pst = prosternum; sbl = tarsal subapical tooth; tpat = tarsal preapical tooth; IIIIX = visible tergites." figureDoi="http://doi.org/10.5281/zenodo.15120444" httpUri="https://zenodo.org/record/15120444/files/figure.png" pageId="4" pageNumber="80">Fig. 3A</figureCitation>
).
</paragraph>
<paragraph id="8B9A368BFFCAEC20FF63FCCAFD0AFCF6" blockId="4.[189,682,848,875]" box="[189,682,848,875]" pageId="4" pageNumber="80">
<smallCapsWord id="8D7CA057FFCAEC20FF63FCCAFEEAFCF5" baselines="868,868" box="[189,330,849,875]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Mesonotum" pageId="4" pageNumber="80">MESONOTUM</smallCapsWord>
<smallCapsWord id="8D7CA057FFCAEC20FE8CFCCDFEDFFCF5" baselines="868" box="[338,383,854,873]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="4" pageNumber="80">AND</smallCapsWord>
<smallCapsWord id="8D7CA057FFCAEC20FE58FCCDFDABFCF5" baselines="868" box="[390,523,854,873]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="metanotum" pageId="4" pageNumber="80">METANOTUM</smallCapsWord>
. Not studied.
</paragraph>
<paragraph id="8B9A368BFFCAEC22FF63FC09FE0AFE10" blockId="4.[188,1398,914,1011]" lastBlockId="6.[189,1400,265,396]" lastPageId="6" lastPageNumber="82" pageId="4" pageNumber="80">
<smallCapsWord id="8D7CA057FFCAEC20FF63FC09FEC4FC36" baselines="933,933" box="[189,356,914,940]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Mesoventrite" pageId="4" pageNumber="80">MESOVENTRITE</smallCapsWord>
. Fused with metaventrite, with sparse setae being denser on metaventral disc, lacking foveae; mesocoxae separated, width of isthmus about half of diameter of mesocoxal cavity; mesoventral and anterior metaventral process subequal in length, truncate; disc of metaventrite slightly convex in anterior part, declining to posterior margin, lacking projection, patch of trichome-like setae or strong median longitudinal keel; metaventral posterior process (
<figureCitation id="131E2A0EFFC8EC22FCB4FEB7FC67FEDA" box="[874,967,300,326]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[189,1398,373,1784]" captionTargetId="figure-11@10.[189,1398,373,1784]" captionTargetPageId="10" captionText="Fig. 3. Apoderiger sikoraisp. nov., paratype,♂ (PCPH).A. Prosternum and metaventrite.B. Metaventrite and abdomen. C. Distal part of tibia and tarsus. Abbrevations: amtp = anterior metaventral process; ihy = inner hypomeral region; hyr = hypomeral ridges; hyg = hypomeral groove; lpcf = lateral procoxal fovea; msvp = mesoventral process; mtvp = posterior metaventral process; ohy = outer hypomeral region; psp = prosternal process; pst = prosternum; sbl = tarsal subapical tooth; tpat = tarsal preapical tooth; IIIIX = visible tergites." figureDoi="http://doi.org/10.5281/zenodo.15120444" httpUri="https://zenodo.org/record/15120444/files/figure.png" pageId="6" pageNumber="82">Fig. 3A</figureCitation>
) wide, short, with straight posterior margin or long, semitriangular, with pointed and setose projecting posterior part exceeding part of abdominal sternites.
</paragraph>
<caption id="DF5A6603FFCAEC20FF61F832FDEFF87B" ID-DOI="http://doi.org/10.5281/zenodo.15120440" ID-Zenodo-Dep="15120440" httpUri="https://zenodo.org/record/15120440/files/figure.png" pageId="4" pageNumber="80" startId="4.[191,234,1961,1987]" targetBox="[291,1382,1113,1903]" targetPageId="4">
<paragraph id="8B9A368BFFCAEC20FF61F832FDEFF87B" blockId="4.[189,1399,1961,2023]" pageId="4" pageNumber="80">
<emphasis id="B951EA99FFCAEC20FF61F832FEAAF85F" bold="true" box="[191,266,1961,1987]" pageId="4" pageNumber="80">Fig. 1.</emphasis>
Dorsal habitus of male.
<emphasis id="B951EA99FFCAEC20FDCAF832FD89F85F" bold="true" box="[532,553,1961,1987]" pageId="4" pageNumber="80">A</emphasis>
.
<taxonomicName id="4C254D08FFCAEC20FDE6F832FC41F85F" authority="Jeannel, 1960" authorityName="Jeannel" authorityYear="1960" box="[568,993,1961,1987]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="80" phylum="Arthropoda" rank="species" species="torticornis">
<emphasis id="B951EA99FFCAEC20FDE6F832FCE0F85F" box="[568,832,1961,1987]" italics="true" pageId="4" pageNumber="80">Apoderiger torticornis</emphasis>
<bibRefCitation id="EFB44B7AFFCAEC20FC9FF832FC41F85F" author="Jeannel R." box="[833,993,1961,1987]" pageId="4" pageNumber="80" pagination="31 - 216" refId="ref8172" refString="Jeannel R. 1960. Revision des Pselaphides Malgaches. Bulletin de l'Academie malgache 36: 31-216." type="journal article" year="1960">Jeannel, 1960</bibRefCitation>
</taxonomicName>
,holotype (MNHN).
<emphasis id="B951EA99FFCAEC20FB13F832FB41F85F" bold="true" box="[1229,1249,1961,1987]" pageId="4" pageNumber="80">B</emphasis>
.
<taxonomicName id="4C254D08FFCAEC20FB2EF832FEACF87A" authorityName="Hlaváč" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="80" phylum="Arthropoda" rank="species" species="banari" status="sp. nov.">
<emphasis id="B951EA99FFCAEC20FB2EF832FEACF87A" italics="true" pageId="4" pageNumber="80">Apoderiger banari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257E2FFCAEC20FECAF856FED1F87B" box="[276,369,1997,2023]" pageId="4" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (PCPH).
</paragraph>
</caption>
<caption id="DF5A6603FFCBEC21FF63F8FFFB86F874" ID-DOI="http://doi.org/10.5281/zenodo.15120442" ID-Zenodo-Dep="15120442" httpUri="https://zenodo.org/record/15120442/files/figure.png" pageId="5" pageNumber="81" startId="5.[189,232,1892,1918]" targetBox="[200,1387,265,1853]" targetPageId="5">
<paragraph id="8B9A368BFFCBEC21FF63F8FFFB86F874" blockId="5.[188,1399,1892,2025]" pageId="5" pageNumber="81">
<emphasis id="B951EA99FFCBEC21FF63F8FFFEB0F8E3" bold="true" box="[189,272,1892,1919]" pageId="5" pageNumber="81">Fig. 2.</emphasis>
<taxonomicName id="4C254D08FFCBEC21FEC5F8FEFDA0F8E3" authorityName="Hlaváč" authorityYear="2025" box="[283,512,1893,1919]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="81" phylum="Arthropoda" rank="species" species="sikorai" status="sp. nov.">
<emphasis id="B951EA99FFCBEC21FEC5F8FEFDA0F8E3" box="[283,512,1893,1919]" italics="true" pageId="5" pageNumber="81">Apoderiger sikorai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257E2FFCBEC21FDD2F8FEFDCDF8E3" box="[524,621,1893,1919]" pageId="5" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♂ (PCPH).
<emphasis id="B951EA99FFCBEC21FC58F8FFFC3BF8E2" bold="true" box="[902,923,1892,1918]" pageId="5" pageNumber="81">A</emphasis>
. Head in ventral view.
<emphasis id="B951EA99FFCBEC21FB1FF8FEFB75F8E3" bold="true" box="[1217,1237,1893,1919]" pageId="5" pageNumber="81">B</emphasis>
. Mouth part.
<emphasis id="B951EA99FFCBEC21FF63F813FF56F83E" bold="true" box="[189,246,1928,1954]" pageId="5" pageNumber="81">CD</emphasis>
. Antenna.
<emphasis id="B951EA99FFCBEC21FEAFF813FE25F83E" bold="true" box="[369,389,1928,1954]" pageId="5" pageNumber="81">E</emphasis>
.
<taxonomicName id="4C254D08FFCBEC21FE4BF813FCEAF83E" authority="Jeannel, 1960" authorityName="Jeannel" authorityYear="1960" box="[405,842,1928,1954]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="81" phylum="Arthropoda" rank="species" species="torticornis">
<emphasis id="B951EA99FFCBEC21FE4BF813FD00F83E" box="[405,672,1928,1954]" italics="true" pageId="5" pageNumber="81">Apoderiger torticornis</emphasis>
<bibRefCitation id="EFB44B7AFFCBEC21FD79F813FCEAF83E" author="Jeannel R." box="[679,842,1928,1954]" pageId="5" pageNumber="81" pagination="31 - 216" refId="ref8172" refString="Jeannel R. 1960. Revision des Pselaphides Malgaches. Bulletin de l'Academie malgache 36: 31-216." type="journal article" year="1960">Jeannel, 1960</bibRefCitation>
</taxonomicName>
, holotype, ♂ (MNHN), antenna. Abbrevations:
<emphasis id="B951EA99FFCBEC21FF62F830FF7FF859" box="[188,223,1963,1989]" italics="true" pageId="5" pageNumber="81">bst</emphasis>
= basistipes;
<emphasis id="B951EA99FFCBEC21FE5EF830FE3CF859" box="[384,412,1963,1989]" italics="true" pageId="5" pageNumber="81">cd</emphasis>
= cardo;
<emphasis id="B951EA99FFCBEC21FDD2F830FD81F859" box="[524,545,1963,1989]" italics="true" pageId="5" pageNumber="81">cl</emphasis>
= clypeus;
<emphasis id="B951EA99FFCBEC21FD77F837FD66F859" box="[681,710,1964,1989]" italics="true" pageId="5" pageNumber="81">gp</emphasis>
= gular plate;
<emphasis id="B951EA99FFCBEC21FCACF830FC29F859" box="[882,905,1963,1989]" italics="true" pageId="5" pageNumber="81">lb</emphasis>
= labrum;
<emphasis id="B951EA99FFCBEC21FBD5F830FB8FF859" box="[1035,1071,1963,1989]" italics="true" pageId="5" pageNumber="81">md</emphasis>
= mandible;
<emphasis id="B951EA99FFCBEC21FB15F837FB4FF859" box="[1227,1263,1964,1989]" italics="true" pageId="5" pageNumber="81">mn</emphasis>
= mentum;
<emphasis id="B951EA99FFCBEC21FF63F854FF4EF874" box="[189,238,1999,2024]" italics="true" pageId="5" pageNumber="81">mxp</emphasis>
= maxillary palpi;
<emphasis id="B951EA99FFCBEC21FE11F854FE49F874" box="[463,489,1999,2024]" italics="true" pageId="5" pageNumber="81">ps</emphasis>
= peg-like sensilla;
<emphasis id="B951EA99FFCBEC21FD06F854FD5EF874" box="[728,766,1999,2024]" italics="true" pageId="5" pageNumber="81">ptp</emphasis>
= posterior tentorisl pits.
</paragraph>
</caption>
<paragraph id="8B9A368BFFC8EC22FF63FE2EFE38FDA4" blockId="6.[189,1400,436,568]" pageId="6" pageNumber="82">
<smallCapsWord id="8D7CA057FFC8EC22FF63FE2EFEB2FE51" baselines="456,456" box="[189,274,437,463]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Elytra" pageId="6" pageNumber="82">ELYTRA</smallCapsWord>
. 1.601.70× as wide as long, 1.501.70 × as long as pronotum, with unevenly distributed short setae; lacking basal foveae; with or without discal and sutural striae; postero-lateral corner of elytron obliquely bevelled mesad, lacking projection; humeri strongly prominent; posterior margin of elytron lacking trichomes.
</paragraph>
<paragraph id="8B9A368BFFC8EC22FF63FDC4FE42FD7F" blockId="6.[189,1399,607,739]" pageId="6" pageNumber="82">
<smallCapsWord id="8D7CA057FFC8EC22FF63FDC4FE8CFDE4" baselines="626,627" box="[189,300,607,633]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Abdomen" pageId="6" pageNumber="82">ABDOMEN</smallCapsWord>
. With composite tergite convex, lacking impressions or keels, lacking trichomes; three paratergites of composite tergite present, first paratergite in anterior portion with trichome; posterior margin of composite tergite with pair of small, bump-like, wide-distant or with pair of long, triangular, short-distant projections.
</paragraph>
<paragraph id="8B9A368BFFC8EC22FF63FC91FAC3FCDB" blockId="6.[189,1398,778,840]" pageId="6" pageNumber="82">
<smallCapsWord id="8D7CA057FFC8EC22FF63FC91FE8CFCBF" baselines="797,798" box="[189,300,778,804]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Abdomen" pageId="6" pageNumber="82">ABDOMEN</smallCapsWord>
(
<figureCitation id="131E2A0EFFC8EC22FE9FFC91FE00FCB8" box="[321,416,778,805]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[189,1398,373,1784]" captionTargetId="figure-11@10.[189,1398,373,1784]" captionTargetPageId="10" captionText="Fig. 3. Apoderiger sikoraisp. nov., paratype,♂ (PCPH).A. Prosternum and metaventrite.B. Metaventrite and abdomen. C. Distal part of tibia and tarsus. Abbrevations: amtp = anterior metaventral process; ihy = inner hypomeral region; hyr = hypomeral ridges; hyg = hypomeral groove; lpcf = lateral procoxal fovea; msvp = mesoventral process; mtvp = posterior metaventral process; ohy = outer hypomeral region; psp = prosternal process; pst = prosternum; sbl = tarsal subapical tooth; tpat = tarsal preapical tooth; IIIIX = visible tergites." figureDoi="http://doi.org/10.5281/zenodo.15120444" httpUri="https://zenodo.org/record/15120444/files/figure.png" pageId="6" pageNumber="82">Fig. 3B</figureCitation>
). With first visible sternite (III) narrow, more than 6 × as short as second visible sternite (IV) which is almost twice as long as third (V); all sternites simple, lacking any modification.
</paragraph>
<paragraph id="8B9A368BFFC8EC22FF63FCF4FAE2FC53" blockId="6.[189,1399,879,975]" pageId="6" pageNumber="82">
<smallCapsWord id="8D7CA057FFC8EC22FF63FCF4FF56FC1B" baselines="898,898" box="[189,246,879,905]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="6" pageNumber="82">LEGS</smallCapsWord>
. Moderately long and slender; femur lacking interlocking ridge; mesotibia differently modified in pre or apical portion, sometimes with mesotibial preapical tooth (
<figureCitation id="131E2A0EFFC8EC22FC2FFC09FBF0FC30" box="[1009,1104,914,940]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[189,1398,373,1784]" captionTargetId="figure-11@10.[189,1398,373,1784]" captionTargetPageId="10" captionText="Fig. 3. Apoderiger sikoraisp. nov., paratype,♂ (PCPH).A. Prosternum and metaventrite.B. Metaventrite and abdomen. C. Distal part of tibia and tarsus. Abbrevations: amtp = anterior metaventral process; ihy = inner hypomeral region; hyr = hypomeral ridges; hyg = hypomeral groove; lpcf = lateral procoxal fovea; msvp = mesoventral process; mtvp = posterior metaventral process; ohy = outer hypomeral region; psp = prosternal process; pst = prosternum; sbl = tarsal subapical tooth; tpat = tarsal preapical tooth; IIIIX = visible tergites." figureDoi="http://doi.org/10.5281/zenodo.15120444" httpUri="https://zenodo.org/record/15120444/files/figure.png" pageId="6" pageNumber="82">Fig. 3C</figureCitation>
); tarsomere 1 minuscule (
<figureCitation id="131E2A0EFFC8EC22FF16FC2EFE83FC53" box="[200,291,949,975]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[189,1398,373,1784]" captionTargetId="figure-11@10.[189,1398,373,1784]" captionTargetPageId="10" captionText="Fig. 3. Apoderiger sikoraisp. nov., paratype,♂ (PCPH).A. Prosternum and metaventrite.B. Metaventrite and abdomen. C. Distal part of tibia and tarsus. Abbrevations: amtp = anterior metaventral process; ihy = inner hypomeral region; hyr = hypomeral ridges; hyg = hypomeral groove; lpcf = lateral procoxal fovea; msvp = mesoventral process; mtvp = posterior metaventral process; ohy = outer hypomeral region; psp = prosternal process; pst = prosternum; sbl = tarsal subapical tooth; tpat = tarsal preapical tooth; IIIIX = visible tergites." figureDoi="http://doi.org/10.5281/zenodo.15120444" httpUri="https://zenodo.org/record/15120444/files/figure.png" pageId="6" pageNumber="82">Fig. 3C</figureCitation>
), 2 much longer, 3 considerably longer than 2, with subtriangular distal lamella (
<figureCitation id="131E2A0EFFC8EC22FB06FC2EFA95FC53" box="[1240,1333,949,975]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[189,1398,373,1784]" captionTargetId="figure-11@10.[189,1398,373,1784]" captionTargetPageId="10" captionText="Fig. 3. Apoderiger sikoraisp. nov., paratype,♂ (PCPH).A. Prosternum and metaventrite.B. Metaventrite and abdomen. C. Distal part of tibia and tarsus. Abbrevations: amtp = anterior metaventral process; ihy = inner hypomeral region; hyr = hypomeral ridges; hyg = hypomeral groove; lpcf = lateral procoxal fovea; msvp = mesoventral process; mtvp = posterior metaventral process; ohy = outer hypomeral region; psp = prosternal process; pst = prosternum; sbl = tarsal subapical tooth; tpat = tarsal preapical tooth; IIIIX = visible tergites." figureDoi="http://doi.org/10.5281/zenodo.15120444" httpUri="https://zenodo.org/record/15120444/files/figure.png" pageId="6" pageNumber="82">Fig. 3C</figureCitation>
).
</paragraph>
<paragraph id="8B9A368BFFC8EC22FF63FC6CFD7AFBCB" blockId="6.[188,1398,1014,1111]" pageId="6" pageNumber="82">
<smallCapsWord id="8D7CA057FFC8EC22FF63FC6CFE93FB93" baselines="1034,1034" box="[189,307,1015,1041]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Aedeagus" pageId="6" pageNumber="82">AEDEAGUS</smallCapsWord>
(
<figureCitation id="131E2A0EFFC8EC22FE96FC6DFE64FB8D" box="[328,452,1014,1041]" captionStart="Fig" captionStartId="12.[189,232,1892,1918]" captionTargetBox="[210,1368,284,1850]" captionTargetId="figure-12@12.[199,1369,269,1854]" captionTargetPageId="12" captionText="Fig. 4. Dorsal and lateral view of aedeagus. AB. Apoderiger torticornis Jeannel, 1960, lectotype (MNHN). CD. Apoderiger sikorai sp. nov., paratype (NMW). EF. Apoderiger spinosus (Dajoz, 1982), holotype (MNHN).GH. Apoderiger banari sp. nov., holotype (PCPH).IJ. Apoderiger grandis sp. nov., holotype (MHNG)." figureDoi="http://doi.org/10.5281/zenodo.15120446" httpUri="https://zenodo.org/record/15120446/files/figure.png" pageId="6" pageNumber="82">Fig. 4AJ</figureCitation>
). Elongate, slender, 2.53.0× as long as wide; basal bulb well-separated from distal lobe, ratio length of basal bulb /length of distal lobe variable 0.92.4; endophallus with or lacking sclerites; dorsal diaphragm round or elliptical.
</paragraph>
<paragraph id="8B9A368BFFC8EC22FF63FBE5FE13FB04" blockId="6.[189,435,1150,1176]" box="[189,435,1150,1176]" pageId="6" pageNumber="82">
<heading id="D0D281E7FFC8EC22FF63FBE5FE13FB04" bold="true" box="[189,435,1150,1176]" fontSize="11" level="3" pageId="6" pageNumber="82" reason="3">
<emphasis id="B951EA99FFC8EC22FF63FBE5FE13FB04" bold="true" box="[189,435,1150,1176]" pageId="6" pageNumber="82">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="8B9A368BFFC8EC22FF63FB36FE16FB76" blockId="6.[189,1398,1197,1258]" pageId="6" pageNumber="82">Females always with short, wide posterior metaventral process, mesotibia always simple, lacking any kind of modification.</paragraph>
</subSubSection>
<subSubSection id="C33F6500FFC8EC22FF63FA89FA81FAC7" pageId="6" pageNumber="82" type="biology_ecology">
<paragraph id="8B9A368BFFC8EC22FF63FA89FE89FAB0" blockId="6.[189,297,1298,1324]" box="[189,297,1298,1324]" pageId="6" pageNumber="82">
<heading id="D0D281E7FFC8EC22FF63FA89FE89FAB0" bold="true" box="[189,297,1298,1324]" fontSize="11" level="3" pageId="6" pageNumber="82" reason="3">
<emphasis id="B951EA99FFC8EC22FF63FA89FE89FAB0" bold="true" box="[189,297,1298,1324]" pageId="6" pageNumber="82">Host ant</emphasis>
</heading>
</paragraph>
<paragraph id="8B9A368BFFC8EC22FF63FADBFA81FAC7" blockId="6.[189,1313,1344,1371]" box="[189,1313,1344,1371]" pageId="6" pageNumber="82">
<taxonomicName id="4C254D08FFC8EC22FF63FADBFD53FAC6" authority="(Forel, 1892)" baseAuthorityName="Forel" baseAuthorityYear="1892" box="[189,755,1344,1370]" class="Insecta" family="Formicidae" genus="Paratrechina" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="82" phylum="Arthropoda" rank="subSpecies" species="amblyops" subSpecies="rubescens">
<emphasis id="B951EA99FFC8EC22FF63FADBFDF1FAC6" box="[189,593,1344,1370]" italics="true" pageId="6" pageNumber="82">Paratrechina amblyops rubescens</emphasis>
(Forel, 1892)
</taxonomicName>
is the only host ant known so far for the genus.
</paragraph>
</subSubSection>
<subSubSection id="C33F6500FFC8EC22FF63FA1AFC33F9E7" pageId="6" pageNumber="82" type="distribution">
<paragraph id="8B9A368BFFC8EC22FF63FA1AFEF8FA07" blockId="6.[189,344,1409,1435]" box="[189,344,1409,1435]" pageId="6" pageNumber="82">
<heading id="D0D281E7FFC8EC22FF63FA1AFEF8FA07" bold="true" box="[189,344,1409,1435]" fontSize="11" level="3" pageId="6" pageNumber="82" reason="3">
<emphasis id="B951EA99FFC8EC22FF63FA1AFEF8FA07" bold="true" box="[189,344,1409,1435]" pageId="6" pageNumber="82">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B9A368BFFC8EC22FF63FA2BFC33F9E7" blockId="6.[188,1400,1456,1659]" pageId="6" pageNumber="82">
The centre of the distribution of the genus
<taxonomicName id="4C254D08FFC8EC22FD6BFA2BFC9CFA56" authorityName="Wasmann" authorityYear="1897" box="[693,828,1456,1482]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="82" phylum="Arthropoda" rank="genus">
<emphasis id="B951EA99FFC8EC22FD6BFA2BFC9CFA56" box="[693,828,1456,1482]" italics="true" pageId="6" pageNumber="82">Apoderiger</emphasis>
</taxonomicName>
is the eastern part of central
<collectingCountry id="F332761BFFC8EC22FB4DFA2BFA83FA56" box="[1171,1315,1456,1482]" name="Madagascar" pageId="6" pageNumber="82">Madagascar</collectingCountry>
, where four species are present,
<taxonomicName id="4C254D08FFC8EC22FE37FA4FFDFBFA71" authorityName="Hlaváč" authorityYear="2025" box="[489,603,1491,1518]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="82" phylum="Arthropoda" rank="species" species="banari" status="sp. nov.">
<emphasis id="B951EA99FFC8EC22FE37FA4FFDFBFA71" box="[489,603,1491,1518]" italics="true" pageId="6" pageNumber="82">A. banari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257E2FFC8EC22FDBAFA4FFD60FA72" box="[612,704,1492,1518]" pageId="6" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C254D08FFC8EC22FD0EFA4FFCE5FA71" authorityName="Hlaváč" authorityYear="2025" box="[720,837,1491,1518]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="82" phylum="Arthropoda" rank="species" species="sikorai" status="sp. nov.">
<emphasis id="B951EA99FFC8EC22FD0EFA4FFCE5FA71" box="[720,837,1491,1518]" italics="true" pageId="6" pageNumber="82">A. sikorai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257E2FFC8EC22FC8FFA4FFC0EFA72" box="[849,942,1492,1518]" pageId="6" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
in the massif Anjozorobe Andasibe,
<taxonomicName id="4C254D08FFC8EC22FF61FA6CFEE4F98C" authorityName="Wasmann" authorityYear="1897" box="[191,324,1527,1553]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="82" phylum="Arthropoda" rank="species" species="cervinus">
<emphasis id="B951EA99FFC8EC22FF61FA6CFEE4F98C" box="[191,324,1527,1553]" italics="true" pageId="6" pageNumber="82">A. cervinus</emphasis>
</taxonomicName>
on the
<taxonomicName id="4C254D08FFC8EC22FE42FA6DFDF5F98C" authority="Plateau" authorityName="Plateau" box="[412,597,1526,1552]" class="Insecta" family="Pyralidae" genus="Imerina" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="82" phylum="Arthropoda" rank="genus">Imerina Plateau</taxonomicName>
and one, the most southern record of an undescribed species, in the Ranomafana National Park. There are three isolated species located more to the north,
<taxonomicName id="4C254D08FFC8EC22FB7AF981FA89F9AF" baseAuthorityName="Dajoz" baseAuthorityYear="1982" box="[1188,1321,1562,1588]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="82" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B951EA99FFC8EC22FB7AF981FA89F9AF" box="[1188,1321,1562,1588]" italics="true" pageId="6" pageNumber="82">A. spinosus</emphasis>
</taxonomicName>
on the western coast,
<taxonomicName id="4C254D08FFC8EC22FEB5F9A6FE61F9CB" authorityName="Dajoz" authorityYear="1982" box="[363,449,1597,1623]" class="Insecta" family="Staphylinidae" genus="Soalala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="82" phylum="Arthropoda" rank="genus">Soalala</taxonomicName>
district,
<taxonomicName id="4C254D08FFC8EC22FDF3F9A6FD0AF9CB" authorityName="Hlaváč" authorityYear="2025" box="[557,682,1597,1623]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="82" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B951EA99FFC8EC22FDF3F9A6FD0AF9CB" box="[557,682,1597,1623]" italics="true" pageId="6" pageNumber="82">A. grandis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257E2FFC8EC22FD6DF9A6FCAFF9CB" box="[691,783,1597,1623]" pageId="6" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
on northeastern part, Manongarivo Special Reserve and
<taxonomicName id="4C254D08FFC8EC22FF31F9FBFE2DF9E6" authorityName="Jeannel" authorityYear="1960" box="[239,397,1632,1658]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="82" phylum="Arthropoda" rank="species" species="torticornis">
<emphasis id="B951EA99FFC8EC22FF31F9FBFE2DF9E6" box="[239,397,1632,1658]" italics="true" pageId="6" pageNumber="82">A. torticornis</emphasis>
</taxonomicName>
in Antongil Bay, north-eastern part (
<figureCitation id="131E2A0EFFC8EC22FCE2F9FBFC24F9E7" box="[828,900,1632,1659]" captionStart="Fig" captionStartId="14.[185,228,1998,2024]" captionTargetBox="[297,1291,509,1959]" captionTargetId="figure-67@14.[297,1291,509,1960]" captionTargetPageId="14" captionText="Fig. 5. Distribution map of Apoderiger Wasmann, 1897." figureDoi="http://doi.org/10.5281/zenodo.15120450" httpUri="https://zenodo.org/record/15120450/files/figure.png" pageId="6" pageNumber="82">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33F6500FFC8EC23FF63F939FDEFFEDA" lastPageId="7" lastPageNumber="83" pageId="6" pageNumber="82" type="discussion">
<paragraph id="8B9A368BFFC8EC22FF63F939FE90F920" blockId="6.[189,304,1698,1724]" box="[189,304,1698,1724]" pageId="6" pageNumber="82">
<heading id="D0D281E7FFC8EC22FF63F939FE90F920" bold="true" box="[189,304,1698,1724]" fontSize="11" level="3" pageId="6" pageNumber="82" reason="3">
<emphasis id="B951EA99FFC8EC22FF63F939FE90F920" bold="true" box="[189,304,1698,1724]" pageId="6" pageNumber="82">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B9A368BFFC8EC23FF63F94AFDEFFEDA" blockId="6.[188,1399,1744,1982]" lastBlockId="7.[189,1399,265,326]" lastPageId="7" lastPageNumber="83" pageId="6" pageNumber="82">
Two well-defined groups of species can be recognized within the genus which are readily separated by the different structure of a pair of projections on the posterior margin of the composite tergite. The first group, containing
<taxonomicName id="4C254D08FFC8EC22FE4AF88CFD66F8AD" authority="(Dajoz, 1982)" baseAuthorityName="Dajoz" baseAuthorityYear="1982" box="[404,710,1814,1841]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="82" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B951EA99FFC8EC22FE4AF88CFDBCF8AC" box="[404,540,1815,1841]" italics="true" pageId="6" pageNumber="82">A. spinosus</emphasis>
(
<bibRefCitation id="EFB44B7AFFC8EC22FDF5F88DFD1EF8AD" author="Dajoz R." box="[555,702,1814,1841]" pageId="6" pageNumber="82" pagination="481 - 522" refId="ref8039" refString="Dajoz R. 1982. Contribution a l'etude des Coleopteres Pselaphidae de Madagascar. Bulletin du Museum national d'Histoire naturelle. Section A, Zoologie, Biologie et Ecologie animales (4): 481-522. https://doi.org/10.5962/p.286051" type="journal article" year="1982">Dajoz, 1982</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C254D08FFC8EC22FCDEF88CFCD4F8AC" authorityName="Hlaváč" authorityYear="2025" box="[768,884,1814,1841]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="82" phylum="Arthropoda" rank="species" species="sikorai" status="sp. nov.">
<emphasis id="B951EA99FFC8EC22FCDEF88CFCD4F8AC" box="[768,884,1814,1841]" italics="true" pageId="6" pageNumber="82">A. sikorai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257E2FFC8EC22FCA5F88CFC79F8AD" box="[891,985,1815,1841]" pageId="6" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
, has posterior margin of composite tergite with pair of small, bump-like, wide-distant projections and strongly curved distal portion of terminal antennomere. The second group, with
<taxonomicName id="4C254D08FFC8EC22FCFAF8C6FC66F8EB" authorityName="Jeannel" authorityYear="1960" box="[804,966,1885,1911]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="82" phylum="Arthropoda" rank="species" species="torticornis">
<emphasis id="B951EA99FFC8EC22FCFAF8C6FC66F8EB" box="[804,966,1885,1911]" italics="true" pageId="6" pageNumber="82">A. torticornis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D08FFC8EC22FC07F8C6FBF9F8EB" authorityName="Hlaváč" authorityYear="2025" box="[985,1113,1885,1911]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="82" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B951EA99FFC8EC22FC07F8C6FBF9F8EB" box="[985,1113,1885,1911]" italics="true" pageId="6" pageNumber="82">A. grandis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257E2FFC8EC22FBBAF8C6FB62F8EB" box="[1124,1218,1885,1911]" pageId="6" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C254D08FFC8EC22FADDF8C6FAD6F8EB" authorityName="Hlaváč" authorityYear="2025" box="[1283,1398,1885,1911]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="82" phylum="Arthropoda" rank="species" species="banari" status="sp. nov.">
<emphasis id="B951EA99FFC8EC22FADDF8C6FAD6F8EB" box="[1283,1398,1885,1911]" italics="true" pageId="6" pageNumber="82">A. banari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257E2FFC8EC22FF63F81AFEB7F807" box="[189,279,1921,1947]" pageId="6" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
, has posterior margin of composite tergite with pair of long, triangular, short-distant projections which are or not exceeding posterior margin of abdomen and slightly curved distal portion of terminal antennomere. Except
<taxonomicName id="4C254D08FFC9EC23FE62FE92FDE0FEBF" authorityName="Wasmann" authorityYear="1897" box="[444,576,265,291]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="83" phylum="Arthropoda" rank="species" species="cervinus">
<emphasis id="B951EA99FFC9EC23FE62FE92FDE0FEBF" box="[444,576,265,291]" italics="true" pageId="7" pageNumber="83">A. cervinus</emphasis>
</taxonomicName>
, because of unknown male, the remaining species of the genus can be separated using the following key.
</paragraph>
</subSubSection>
<subSubSection id="C33F6500FFC9EC23FF63FEF5FEC0FE14" box="[189,352,366,392]" pageId="7" pageNumber="83" type="multiple">
<paragraph id="8B9A368BFFC9EC23FF63FEF5FEC0FE14" blockId="7.[189,352,366,392]" box="[189,352,366,392]" pageId="7" pageNumber="83">
<heading id="D0D281E7FFC9EC23FF63FEF5FEC0FE14" bold="true" box="[189,352,366,392]" fontSize="11" level="3" pageId="7" pageNumber="83" reason="3">
<emphasis id="B951EA99FFC9EC23FF63FEF5FEC0FE14" bold="true" box="[189,352,366,392]" pageId="7" pageNumber="83">Key to males</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F6500FFC9EC23FF63FE07FB71FE2A" box="[189,1233,412,438]" pageId="7" pageNumber="83" type="description">
<paragraph id="8B9A368BFFC9EC23FF63FE07FB71FE2A" blockId="7.[189,1233,412,438]" box="[189,1233,412,438]" pageId="7" pageNumber="83">
(
<taxonomicName id="4C254D08FFC9EC23FF19FE07FE19FE2A" ID-CoL="FRJV" authorityName="Wasmann" authorityYear="1897" box="[199,441,412,438]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="83" phylum="Arthropoda" rank="species" species="cervinus">
<emphasis id="B951EA99FFC9EC23FF19FE07FE19FE2A" box="[199,441,412,438]" italics="true" pageId="7" pageNumber="83">Apoderiger cervinus</emphasis>
</taxonomicName>
is missing in the key because the male of this species is unknown).
</paragraph>
</subSubSection>
<subSubSection id="C33F6500FFC9EC23FF63FE45FAD7FB35" pageId="7" pageNumber="83" type="key">
<key id="FFB461BAFFC9EC23FF63FE45FAD7FB35" pageId="7" pageNumber="83">
<keyStep id="30D12E2EFFC9EC23FF63FE45FAD7FDFD" pageId="7" pageNumber="83">
<paragraph id="8B9A368BFFC9EC23FF63FE45FAD7FD87" blockId="7.[189,1399,478,610]" pageId="7" pageNumber="83">
<keyLead id="30D495BEFFC9EC23FF63FE45FAD7FD87" pageId="7" pageNumber="83">
1. Posterior margin of composite tergite with pair of small, bump-like, wide-distant projections (
<taxonomicName id="4C254D08FFC9EC23FF2AFD9AFEDBFD87" ID-CoL="FRJW" baseAuthorityName="Dajoz" baseAuthorityYear="1982" box="[244,379,513,539]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="83" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B951EA99FFC9EC23FF2AFD9AFEDBFD87" box="[244,379,513,539]" italics="true" pageId="7" pageNumber="83">A. spinosus</emphasis>
</taxonomicName>
species group) .............................................................................................................. 2
</keyLead>
</paragraph>
<paragraph id="8B9A368BFFC9EC23FF1FFDBEFAD7FDFD" blockId="7.[189,1399,478,610]" pageId="7" pageNumber="83">
<keyLead id="30D495BEFFC9EC23FF1FFDBEFAD7FDFD" pageId="7" pageNumber="83">
Posterior margin of composite tergite with pair of long, triangular, short-distant projections (
<taxonomicName id="4C254D08FFC9EC23FF2AFDD3FE32FDFD" ID-CoL="FRJX" authorityName="Jeannel" authorityYear="1960" box="[244,402,584,610]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="83" phylum="Arthropoda" rank="species" species="torticornis">
<emphasis id="B951EA99FFC9EC23FF2AFDD3FE32FDFD" box="[244,402,584,610]" italics="true" pageId="7" pageNumber="83">A. torticornis</emphasis>
</taxonomicName>
species group) ........................................................................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D12E2EFFC9EC23FF63FD12FAD7FC91" pageId="7" pageNumber="83">
<paragraph id="8B9A368BFFC9EC23FF63FD12FAD6FD5A" blockId="7.[189,1399,649,781]" pageId="7" pageNumber="83">
<keyLead id="30D495BEFFC9EC23FF63FD12FAD6FD5A" pageId="7" pageNumber="83">
2. Small species, about
<quantity id="4CDD9B6EFFC9EC23FE3CFD12FDEBFD3F" box="[482,587,649,675]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="7" pageNumber="83" unit="mm" value="1.4">1.40 mm</quantity>
long; apex of mesotibia with inner projection ................................... ...................................................................................................................
<taxonomicName id="4C254D08FFC9EC23FB9EFD37FAD6FD5A" ID-CoL="FRJW" authority="(Dajoz, 1982)" baseAuthorityName="Dajoz" baseAuthorityYear="1982" box="[1088,1398,684,710]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="83" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B951EA99FFC9EC23FB9EFD37FB6CFD5A" bold="true" box="[1088,1228,684,710]" italics="true" pageId="7" pageNumber="83">A. spinosus</emphasis>
(
<bibRefCitation id="EFB44B7AFFC9EC23FB02FD37FACEFD5A" author="Dajoz R." box="[1244,1390,684,710]" pageId="7" pageNumber="83" pagination="481 - 522" refId="ref8039" refString="Dajoz R. 1982. Contribution a l'etude des Coleopteres Pselaphidae de Madagascar. Bulletin du Museum national d'Histoire naturelle. Section A, Zoologie, Biologie et Ecologie animales (4): 481-522. https://doi.org/10.5962/p.286051" type="journal article" year="1982">Dajoz, 1982</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9A368BFFC9EC23FF1FFD4BFAD7FC91" blockId="7.[189,1399,649,781]" pageId="7" pageNumber="83">
<keyLead id="30D495BEFFC9EC23FF1FFD4BFAD7FC91" pageId="7" pageNumber="83">
Large species, about
<quantity id="4CDD9B6EFFC9EC23FE33FD54FD3BFD76" box="[493,667,719,746]" metricMagnitude="-3" metricUnit="m" metricValue="1.775" metricValueMax="1.85" metricValueMin="1.7" pageId="7" pageNumber="83" unit="mm" value="1.775" valueMax="1.85" valueMin="1.7">1.701.85 mm</quantity>
long; apex of mesotibia lacking inner projection, only with small preapical tooth (
<figureCitation id="131E2A0EFFC9EC23FE2EFD69FDEAFC90" box="[496,586,754,780]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[189,1398,373,1784]" captionTargetId="figure-11@10.[189,1398,373,1784]" captionTargetPageId="10" captionText="Fig. 3. Apoderiger sikoraisp. nov., paratype,♂ (PCPH).A. Prosternum and metaventrite.B. Metaventrite and abdomen. C. Distal part of tibia and tarsus. Abbrevations: amtp = anterior metaventral process; ihy = inner hypomeral region; hyr = hypomeral ridges; hyg = hypomeral groove; lpcf = lateral procoxal fovea; msvp = mesoventral process; mtvp = posterior metaventral process; ohy = outer hypomeral region; psp = prosternal process; pst = prosternum; sbl = tarsal subapical tooth; tpat = tarsal preapical tooth; IIIIX = visible tergites." figureDoi="http://doi.org/10.5281/zenodo.15120444" httpUri="https://zenodo.org/record/15120444/files/figure.png" pageId="7" pageNumber="83">Fig. 3C</figureCitation>
) ..............................................................................
<emphasis id="B951EA99FFC9EC23FB40FD69FA87FC90" bold="true" box="[1182,1319,754,781]" pageId="7" pageNumber="83">
<taxonomicName id="4C254D08FFC9EC23FB40FD69FAB4FC91" authorityName="Hlaváč" authorityYear="2025" box="[1182,1300,754,781]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="83" phylum="Arthropoda" rank="species" species="sikorai">
<emphasis id="B951EA99FFC9EC23FB40FD69FAB4FC91" bold="true" box="[1182,1300,754,781]" italics="true" pageId="7" pageNumber="83">A. sikorai</emphasis>
</taxonomicName>
s
</emphasis>
p. nov.
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D12E2EFFC9EC23FF63FCAFFAD7FC62" pageId="7" pageNumber="83">
<paragraph id="8B9A368BFFC9EC23FF63FCAFFAD7FC09" blockId="7.[189,1399,820,1022]" pageId="7" pageNumber="83">
<keyLead id="30D495BEFFC9EC23FF63FCAFFAD7FC09" pageId="7" pageNumber="83">
3. Triangular paired projections on posterior portion of composite tergite shorter, not exceeding posterior margin of abdomen; posterior subtriangular metaventral process exceedingly over first visible sternite (III) ................................................................................................
<taxonomicName id="4C254D08FFC9EC23FB7FFCE1FAB5FC09" authorityName="Hlaváč" authorityYear="2025" box="[1185,1301,890,917]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="83" phylum="Arthropoda" rank="species" species="banari" status="sp. nov.">
<emphasis id="B951EA99FFC9EC23FB7FFCE1FAB5FC09" bold="true" box="[1185,1301,890,917]" italics="true" pageId="7" pageNumber="83">A. banari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257E2FFC9EC23FAC2FCE0FAD7FC09" box="[1308,1399,891,917]" pageId="7" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B9A368BFFC9EC23FF1FFC05FAD7FC62" blockId="7.[189,1399,820,1022]" pageId="7" pageNumber="83">
<keyLead id="30D495BEFFC9EC23FF1FFC05FAD7FC62" pageId="7" pageNumber="83"> Triangular paired projections on posterior portion of composite tergite longer, clearly exceeding posterior margin of abdomen; posterior metaventral process wide, short, with almost straight posterior margin, not exceedingly over first visible sternite (III) ..................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D12E2EFFC9EC23FF63FBBEFAD7FB35" pageId="7" pageNumber="83">
<paragraph id="8B9A368BFFC9EC23FF63FBBEFAD7FBFF" blockId="7.[189,1399,1061,1193]" pageId="7" pageNumber="83">
<keyLead id="30D495BEFFC9EC23FF63FBBEFAD7FBFF" pageId="7" pageNumber="83">
4. Elytra with discal and sutural striae; large species, about
<quantity id="4CDD9B6EFFC9EC23FC5AFBBEFC7EFBDC" box="[900,990,1061,1088]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="83" unit="mm" value="2.2">2.2 mm</quantity>
long; head less than 1.60 × as long as pronotum ..........................................................................................................
<taxonomicName id="4C254D08FFC9EC23FB48FBD3FAB4FBFF" authorityName="Hlaváč" authorityYear="2025" box="[1174,1300,1096,1123]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="83" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="B951EA99FFC9EC23FB48FBD3FAB4FBFF" bold="true" box="[1174,1300,1096,1123]" italics="true" pageId="7" pageNumber="83">A. grandis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26257E2FFC9EC23FAC2FBD2FAD7FBFF" box="[1308,1399,1097,1123]" pageId="7" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B9A368BFFC9EC23FF1FFBF7FAD7FB35" blockId="7.[189,1399,1061,1193]" pageId="7" pageNumber="83">
<keyLead id="30D495BEFFC9EC23FF1FFBF7FAD7FB35" pageId="7" pageNumber="83">
Elytra lacking discal and sutural striae; small species, about
<quantity id="4CDD9B6EFFC9EC23FC7CFBF7FB8EFB1A" box="[930,1070,1132,1158]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="7" pageNumber="83" unit="mm" value="1.85" valueMax="1.9" valueMin="1.8">1.81.9 mm</quantity>
long; head more than 1.75 × as long as pronotum ...............................................................................
<taxonomicName id="4C254D08FFC9EC23FBF2FB14FAD7FB35" ID-CoL="FRJX" authority="Jeannel, 1960" authorityName="Jeannel" authorityYear="1960" box="[1068,1399,1167,1193]" class="Insecta" family="Staphylinidae" genus="Apoderiger" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="83" phylum="Arthropoda" rank="species" species="torticornis">
<emphasis id="B951EA99FFC9EC23FBF2FB14FB6EFB35" bold="true" box="[1068,1230,1167,1193]" italics="true" pageId="7" pageNumber="83">A. torticornis</emphasis>
<bibRefCitation id="EFB44B7AFFC9EC23FB0BFB14FAD7FB35" author="Jeannel R." box="[1237,1399,1167,1193]" pageId="7" pageNumber="83" pagination="31 - 216" refId="ref8172" refString="Jeannel R. 1960. Revision des Pselaphides Malgaches. Bulletin de l'Academie malgache 36: 31-216." type="journal article" year="1960">Jeannel, 1960</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>