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<paragraph pageId="1" pageNumber="88">Taxon classification Animalia Coleoptera Carabidae</paragraph>
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Taxon classification Animalia
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<taxonomicName id="B37921225111F7D571031D67F89284A4" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
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<taxonomicName authority="LaFerte-Senectere, 1851" authorityName="LaFerte-Senectere" authorityYear="1851" class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">
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<paragraph id="D9D8E9C33D8C204E2FFD8953D4C606E9" pageId="1" pageNumber="88">
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<taxonomicName id="C85C0C65E03B1DD00F285615E367730F" ID-CoL="8KX8N" authority="LaFerte-Senectere, 1851" authorityName="LaFerte-Senectere" authorityYear="1851" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">
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Hololeius
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, 1851
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</paragraph>
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<subSubSection pageId="1" pageNumber="88" type="type species">
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<paragraph pageId="1" pageNumber="88">Type species.</paragraph>
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<paragraph pageId="1" pageNumber="88">
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<taxonomicName class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius nitidulus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="nitidulus">Chlaenius nitidulus</taxonomicName>
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Dejean, 1826
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<subSubSection id="5A487C7C67F0B988F74C25A582CC2AC6" pageId="1" pageNumber="88" type="type species">
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<paragraph id="5D866AEED83FFA02FFFCE91BE82FFDE9" pageId="1" pageNumber="88">Type species.</paragraph>
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<paragraph id="00A745662B2276A2AD987B371100FD1A" pageId="1" pageNumber="88">
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<taxonomicName id="B709B1E0256672F77A420E635B702999" authority="Dejean, 1826" authorityName="Dejean" authorityYear="1826" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" lsidName="Chlaenius nitidulus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="nitidulus">Chlaenius nitidulus Dejean, 1826</taxonomicName>
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</paragraph>
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<subSubSection pageId="1" pageNumber="88" type="diagnosis">
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<paragraph pageId="1" pageNumber="88">Diagnosis.</paragraph>
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<paragraph pageId="1" pageNumber="88">Small to medium-sized specimens (11-13 mm) for Oriental chlaeniines, with the following characters: habitus semi-elongate; color mostly olivaceous; integument punctate and pubescent, pubescence denser on sides of elytra and on sides of abdomen, less developed, scattered or lacking on rest of body; head with conspicuously large eyes and minute tempora; penultimate segment of labial palpomere without setae; antennomeres 1-3 lighter than other antennomeres, scape and pedicel glabrous, antennomere 3 with a few fine and scattered setae; pronotum subquadrate, without protruded angles, with basal margin laterally oblique towards posterior angle; elytra more densely pubescent and punctate laterally and apically, with only a few hairs basally and on disc; striae 1-7 of elytra punctiform; striae 5-7 obliterated anteriorly; stria 8 linear, somewhat more impressed than other striae; mesocoxa with two long setae, one lateral seta and one posteromedial seta; sterna 4-5 with pair of long ambulatory setae, sternum 3 without such setae; last abdominal sternum with two pores in male, four pores in female; male protarsomeres 1-3 longer than wide and with protarsomere 1 longer than each of following two protarsomeres separately; tarsomere 5 of all legs setose ventrally.</paragraph>
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<subSubSection id="1D51AC8E96C4F388135A60E83C23C543" pageId="1" pageNumber="88" type="diagnosis">
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<paragraph id="48906EDEA69F7F54A3E0BEEA90C240A8" pageId="1" pageNumber="88">Diagnosis.</paragraph>
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||||
<paragraph id="8966415A3C2FF9919EC1A9CA3A676686" pageId="1" pageNumber="88">Small to medium-sized specimens (11-13 mm) for Oriental chlaeniines, with the following characters: habitus semi-elongate; color mostly olivaceous; integument punctate and pubescent, pubescence denser on sides of elytra and on sides of abdomen, less developed, scattered or lacking on rest of body; head with conspicuously large eyes and minute tempora; penultimate segment of labial palpomere without setae; antennomeres 1-3 lighter than other antennomeres, scape and pedicel glabrous, antennomere 3 with a few fine and scattered setae; pronotum subquadrate, without protruded angles, with basal margin laterally oblique towards posterior angle; elytra more densely pubescent and punctate laterally and apically, with only a few hairs basally and on disc; striae 1-7 of elytra punctiform; striae 5-7 obliterated anteriorly; stria 8 linear, somewhat more impressed than other striae; mesocoxa with two long setae, one lateral seta and one posteromedial seta; sterna 4-5 with pair of long ambulatory setae, sternum 3 without such setae; last abdominal sternum with two pores in male, four pores in female; male protarsomeres 1-3 longer than wide and with protarsomere 1 longer than each of following two protarsomeres separately; tarsomere 5 of all legs setose ventrally.</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="88" type="taxonomic position">
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<paragraph pageId="1" pageNumber="88">Taxonomic position.</paragraph>
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<paragraph pageId="1" pageNumber="88">
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||||
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<paragraph id="4651C0BEEB68BE734959BE4B554D2466" pageId="1" pageNumber="88">Taxonomic position.</paragraph>
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(1851
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: 274) proposed
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for
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<taxonomicName class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius nitidulus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="nitidulus">Chlaenius nitidulus</taxonomicName>
|
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Dejean, 1826, due to: 1, pronotum at the base narrower than the base of elytra; 2, lack of pubescence; 3, antennomere 3 not longer than subsequent antennomeres. He placed the genus in tribe
|
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<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
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, 1851, concluding:
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||||
, due to: 1, pronotum at the base narrower than the base of elytra; 2, lack of pubescence; 3, antennomere 3 not longer than subsequent antennomeres. He placed the genus in tribe
|
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, 1851
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, concluding:
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de la largeur du corselet, cet insecte
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tout les
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desquels nous avons
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les Oodites des
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. In terms of the present knowledge, I can state that the first and third characters are true, but not the second one. When examined it was found that
|
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<taxonomicName class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius ceylanicus" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="ceylanicus">Hololeius ceylanicus</taxonomicName>
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|
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has the integument partially and diffusely punctate and pubescent, both dorsally and ventrally. This state is opposed to one of the basic features of the
|
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<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
|
||||
<taxonomicName id="9C2E9439B5454EF1CA781537FAF040A9" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
|
||||
, namely the lack of pubescence and lack of extensive punctation of the integument (
|
||||
<bibRefCitation author="Bousquet, Y" journalOrPublisher="The Canadian Entomologist" pageId="10" pageNumber="97" pagination="443 - 537" title="Taxonomic revision of Nearctic, Mexican, and West Indian Oodini (Coleoptera: Carabidae)." url="10.4039/Ent128443-3" volume="128" year="1996">Bousquet 1996</bibRefCitation>
|
||||
<bibRefCitation id="0167231B1C4122A98FCF8827C329F8E4" author="Bousquet, Y" journalOrPublisher="The Canadian Entomologist" pageId="10" pageNumber="97" pagination="443 - 537" title="Taxonomic revision of Nearctic, Mexican, and West Indian Oodini (Coleoptera: Carabidae)." url="10.4039/Ent128443-3" volume="128" year="1996">Bousquet 1996</bibRefCitation>
|
||||
: 448). Hence, the three characters
|
||||
<normalizedToken originalValue="LaFerté-Sénectére">LaFerte-Senectere</normalizedToken>
|
||||
<normalizedToken id="2417E4CDCE414310CA582CFFA3896580" originalValue="LaFerté-Sénectére">LaFerte-Senectere</normalizedToken>
|
||||
mentioned are typical of
|
||||
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
|
||||
<taxonomicName id="96991110C4FF72CC1E5C6498093DE269" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
|
||||
. The length and ratio of antennomere 3 to the following antennomeres is not of tribal significance.
|
||||
</paragraph>
|
||||
<paragraph pageId="1" pageNumber="88">
|
||||
<taxonomicName class="Insecta" family="Carabidae" genus="Hololeius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hololeius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Hololeius</taxonomicName>
|
||||
<paragraph id="0547C711A3C700F1A45D7A1AD7DBE938" pageId="1" pageNumber="88">
|
||||
<taxonomicName id="97773DCCC047C182CE54E929464BB637" class="Insecta" family="Carabidae" genus="Hololeius" kingdom="Animalia" lsidName="Hololeius" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Hololeius</taxonomicName>
|
||||
is usually placed in the beginning of the genera of
|
||||
<taxonomicName lsidName="" pageId="1" pageNumber="88" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
|
||||
<taxonomicName id="34D27C5BE87138CADD428574E185B9A0" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
|
||||
(
|
||||
<bibRefCitation author="Lorenz, W" journalOrPublisher="Tutzing" pageId="11" pageNumber="98" title="Systematic list of extant ground beetles of the world (Insecta Coleoptera " Geadephaga ": Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae)" year="1998">Lorenz 1998</bibRefCitation>
|
||||
<bibRefCitation id="3C0797117AE4A0047CD584BAFF41016F" author="Lorenz, W" journalOrPublisher="Tutzing" pageId="11" pageNumber="98" title="Systematic list of extant ground beetles of the world (Insecta Coleoptera " Geadephaga ": Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae)" year="1998">Lorenz 1998</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation author="Lorenz, W" journalOrPublisher="Tutzing" pageId="11" pageNumber="98" title="Systematic list of extant ground beetles of the World. (Insecta, Coleoptera, Adephaga: Trachypachidae & Carabidae incl. Paussinae, Cicindelinae, Rhysodinae)" year="2005">2005</bibRefCitation>
|
||||
<bibRefCitation id="8BA1C2D4019BDB88EB25E9E0ED9295BF" author="Lorenz, W" journalOrPublisher="Tutzing" pageId="11" pageNumber="98" title="Systematic list of extant ground beetles of the World. (Insecta, Coleoptera, Adephaga: Trachypachidae & Carabidae incl. Paussinae, Cicindelinae, Rhysodinae)" year="2005">2005</bibRefCitation>
|
||||
), presumably because of some odd features that it possesses. Actually, the genus combines characters distinctive for either one or the other tribe and the importance of each is discussed below (see
|
||||
<normalizedToken originalValue="‘Affinities’">'Affinities'</normalizedToken>
|
||||
<normalizedToken id="D2CB4519D2FC62E1B737A19FBCCE0906" originalValue="‘Affinities’">'Affinities'</normalizedToken>
|
||||
under
|
||||
<taxonomicName class="Insecta" family="Carabidae" genus="Bamaroodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bamaroodes" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">Bamaroodes</taxonomicName>
|
||||
gen. n.,
|
||||
<normalizedToken originalValue="‘Discussion’">'Discussion'</normalizedToken>
|
||||
<taxonomicName id="6DB58F4FE315024FB9B87FD4BD902F27" class="Insecta" family="Carabidae" genus="Bamaroodes" kingdom="Animalia" lsidName="Bamaroodes" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus" status="gen. nov.">Bamaroodes</taxonomicName>
|
||||
<taxonomicNameLabel id="25DEB8619171D7EB8B290F79962E7643" rank="genus">gen. n.</taxonomicNameLabel>
|
||||
,
|
||||
<normalizedToken id="C262250E3A71C0AF2A80DF49E45E67C8" originalValue="‘Discussion’">'Discussion'</normalizedToken>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
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