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<mods:title id="4CEDEDF2F1614C6D14E5458D33869AD7">Checklist of British and Irish Hymenoptera - Ichneumonidae</mods:title>
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<mods:part id="3A9F89FE05C1766134224C5C74272E4A">
<mods:date id="16511F6A462BA1D5A9D28F828E10FECD">2016</mods:date>
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Isadelphus
<normalizedToken id="824ECBA3D69B901EBED1B3C4320C56B5" originalValue="Förster">Foerster</normalizedToken>
, 1869
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<taxonomicName id="4857A0C951AB0C146F6EC80B6485FBA8" class="Insecta" family="Ichneumonidae" genus="Perosis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="PEROSIS" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="genus">PEROSIS</taxonomicName>
<normalizedToken id="5CF187E831F46168B6AA10173735FC1C" originalValue="Förster">Foerster</normalizedToken>
, 1869
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<taxonomicName id="7F7F524122900E63233F3936BD176683" class="Insecta" family="Ichneumonidae" genus="Cecidonomus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="CECIDONOMUS" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="genus">CECIDONOMUS</taxonomicName>
Bridgman, 1880
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<paragraph id="C6AE02E004CE9E874F6E87E886C12444" pageId="0" pageNumber="9042">Notes</paragraph>
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See note for
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.
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Type status:
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<taxonomicName ID-CoL="74VBB" authority="Linnaeus" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Lamiaceae" genus="Origanum" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="syriacum">
<emphasis bold="true" pageNumber="708">Origanum syriacum</emphasis>
Linnaeus
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,
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2
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&quot;Habitat - - -
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RCN: 4295.
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<typeStatus>
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</typeStatus>
(Paton in Jarvis &amp; al. in
<emphasis italics="true" pageNumber="708">Taxon</emphasis>
50: 516. 2001): [icon]
<emphasis italics="true" pageNumber="708">&quot;Marum syriacum&quot;</emphasis>
in Plantin, Pl. Stirp. Icon.: 499. 1581.
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<paragraph pageNumber="708">
Current name:
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<emphasis italics="true" pageNumber="708">
<emphasis bold="true" italics="true" pageNumber="708">Origanum syriacum</emphasis>
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L.
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(
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<paragraph pageNumber="708">
<emphasis italics="true" pageNumber="708">Note:</emphasis>
As pointed out by Harley (in
<emphasis italics="true" pageNumber="708">Watsonia</emphasis>
14: 86. 1982), Ietswaart (in
<emphasis italics="true" pageNumber="708">Leiden Bot. Ser.</emphasis>
4: 87. 1980) designated 743.12 (LINN) as the type of both
<taxonomicName authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Lamiaceae" genus="Origanum" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="syriacum">
<emphasis italics="true" pageNumber="708">O. syriacum</emphasis>
</taxonomicName>
and
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<emphasis italics="true" pageNumber="708">O. maru</emphasis>
L. Harley
</taxonomicName>
stated &quot;while the sheet indicated is unquestionably the type of
<taxonomicName authorityName="Linnaeus" authorityYear="1763" class="Magnoliopsida" family="Lamiaceae" genus="Origanum" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="maru">
<emphasis italics="true" pageNumber="708">O. maru</emphasis>
</taxonomicName>
, it can hardly be the type of
<taxonomicName authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Lamiaceae" genus="Origanum" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="syriacum">
<emphasis italics="true" pageNumber="708">O. syriacum</emphasis>
</taxonomicName>
, which apparently does not occur among the sheets in
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herbarium in London, and must be looked for elsewhere&quot;. Meikle (
<emphasis italics="true" pageNumber="708">Fl. Cyprus</emphasis>
2: 1267. 1985) stated &quot;TYPE: Probably the illustration in Lobel...t. 499 (1591)&quot;, i.e. the Plantin illustration, but his use of the word
<normalizedToken originalValue="“probably”">&quot;probably&quot;</normalizedToken>
contradicts Art. 7.11 of the Code.
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<paragraph id="E7310E0185E06902B53CAF1916D026FF" pageNumber="79">3. Avena panicula attenuata, calycibus bifloris seminibus villosis, aristis calyce duplo longioribus.</paragraph>
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<emphasis id="AA233DE4F7F2121D350FF61EDDD5B879" italics="true">Habitat in</emphasis>
Pensylvania
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<mods:title>Lizards (Reptilia: Squamata) from the Caatinga, northeastern Brazil: Detailed and updated overview</mods:title>
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<mods:namePart>Uchoa, Lucas Rafael</mods:namePart>
<mods:affiliation>Centro de Estudos Superiores de Caxias, Universidade Estadual do Maranhao, Programa de Pos-Graduacao em Biodiversidade, Ambiente e Saude, Praca Duque de Caxias, 65604 - 380, Caxias, MA, Brazil</mods:affiliation>
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<mods:namePart>Delfim, Fagner Ribeiro</mods:namePart>
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<mods:namePart>Mesquita, Daniel Oliveira</mods:namePart>
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<mods:namePart>Colli, Guarino Rinaldi</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Garda, Adrian Antonio</mods:namePart>
<mods:affiliation>Departamento de Botanica e Zoologia, Universidade Federal do Rio Grande do Norte, 59078 - 900, Natal, RN, Brazil</mods:affiliation>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Guedes, Thais B.</mods:namePart>
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<mods:affiliation>Departamento de Biologia Animal, Instituto de Biologia, Universidade Estadual de Campinas, 13083 - 862, Campinas, SP, Brazil &amp; University of Gothenburg, Gothenburg Global Biodiversity Center, Department of Biological and Environmental Sciences, Box 461, SE- 405 30, Goeteborg, Sweden</mods:affiliation>
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<taxonomicName LSID="7C13F3D7-3BAA-54EE-B1CC-DF53D65F4A0F" authority="(Manzani &amp; Abe, 1990)" baseAuthorityName="Manzani &amp; Abe" baseAuthorityYear="1990" class="Reptilia" family="Tropiduridae" genus="Tropidurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropidurus helenae" order="Squamata" pageId="0" pageNumber="599" phylum="Chordata" rank="species" species="Helenae">Tropidurus helenae (Manzani &amp; Abe, 1990)</taxonomicName>
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Lizards recorded in the Caatinga region. 12.1 Tropidurus helenae; 12.2 Tropidurus hispidus; 12.3 Tropidurus jaguaribanus; 12.4 Tropidurus montanus; 12.5 Tropidurus mucujensis; 12.6 Tropidurus oreadicus; 12.7 Tropidurus pinima and 12.8 Tropidurus semitaeniatus. Photograph credits: Daniel Mesquita (12.1, 12.6, 12.8), Adrian Garda (12.2), Daniel Passos (12.3, 12.4, 12.7), Marco A. Freitas (12.5)." figureDoi="10.3897/vz.72.e78828.figure12" httpUri="https://binary.pensoft.net/fig/726820" pageId="0" pageNumber="599">Figs 12.1 and 20</figureCitation>
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<paragraph pageId="0" pageNumber="599">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="599">
Toca de Cima dos
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, municipality of
<normalizedToken originalValue="São">Sao</normalizedToken>
Raimundo Nonato, state of
<normalizedToken originalValue="Piauí">Piaui</normalizedToken>
, Brazil.
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<paragraph pageId="0" pageNumber="599">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="599">
Caatinga endemic species. It is recorded only in the state of
<normalizedToken originalValue="Piauí">Piaui</normalizedToken>
. It shows restricted distribution in the Caatinga and with annual mean temperature 20 to 28°C along three ecoregions (Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Lizards from the Caatinga. Summarized data about distribution, ecology, and conservation. Caatinga ecoregions (Silva et al. 2017) - B: Brejos, CCD: Complexo Chapada Diamantina, CIA: Complexo Ibiapaba-Araripe, DSM: Depressao Sertaneja Meridional, DSS: Depressao Sertaneja Setentorial, DSF: Dunas do Sao Francisco, PB: Planalto da Borborema, RC: Raso da Catarina, SFG: Sao Francisco - Gurgueia. Distribution Patterns - WD: widespread (EOO greater than 20,000 km 2, see Material and Methods for details), R: restricted (EOO less than 20,000 km 2). Habitat - O: open areas; F: forested areas; Preferred habitat type: AT: ant hills and termite mounds, B: bromeliads, G: general, LL: leaf litter, P: psammophilous, A: anthropic; S: saxicolous and TB: trees and bushes; Activity - D: diurnal, N: nocturnal and C: crepuscular. Conservation status (IUCN 2021; ICMBio 2018) - Critically Endangered (CR), Data Deficient (DD), Endangered (EN), Least Concern (LC), Near Threatened (NT), Not Evaluated (NE) and Vulnerable (VU). Caatinga endemic species ' * '." httpUri="http://table.plazi.org/id/DCA21B21C4ADC68A0D52968CF9844DE2" pageId="0" pageNumber="599" tableUuid="DCA21B21C4ADC68A0D52968CF9844DE2">1</tableCitation>
; Appendix S3). Distributed in low to medium elevation areas (259-387 m a.s.l.), with annual mean temperature 26 to 27°C, and average annual rainfall between 702 and 849 mm.
</paragraph>
<caption doi="10.3897/vz.72.e78828.figure12" httpUri="https://binary.pensoft.net/fig/726820" pageId="0" pageNumber="599" start="Figure 12" startId="F12">
<paragraph pageId="0" pageNumber="599">
<emphasis bold="true" pageId="0" pageNumber="599">Figure 12.</emphasis>
Lizards recorded in the Caatinga region.
<emphasis bold="true" pageId="0" pageNumber="599">12.1</emphasis>
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<emphasis italics="true" pageId="0" pageNumber="599">Tropidurus helenae</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="599">12.2</emphasis>
<taxonomicName baseAuthorityName="Spix" baseAuthorityYear="1825" class="Reptilia" family="Tropiduridae" genus="Tropidurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropidurus hispidus" order="Squamata" pageId="0" pageNumber="599" phylum="Chordata" rank="species" species="Hispidus">
<emphasis italics="true" pageId="0" pageNumber="599">Tropidurus hispidus</emphasis>
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;
<emphasis bold="true" pageId="0" pageNumber="599">12.3</emphasis>
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<emphasis italics="true" pageId="0" pageNumber="599">Tropidurus jaguaribanus</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="599">12.4</emphasis>
<taxonomicName authorityName="Rodrigues" authorityYear="1987" class="Reptilia" family="Tropiduridae" genus="Tropidurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropidurus montanus" order="Squamata" pageId="0" pageNumber="599" phylum="Chordata" rank="species" species="Montanus">
<emphasis italics="true" pageId="0" pageNumber="599">Tropidurus montanus</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="599">12.5</emphasis>
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<emphasis italics="true" pageId="0" pageNumber="599">Tropidurus mucujensis</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="599">12.6</emphasis>
<taxonomicName authorityName="Rodrigues" authorityYear="1987" class="Reptilia" family="Tropiduridae" genus="Tropidurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropidurus oreadicus" order="Squamata" pageId="0" pageNumber="599" phylum="Chordata" rank="species" species="Oreadicus">
<emphasis italics="true" pageId="0" pageNumber="599">Tropidurus oreadicus</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="599">12.7</emphasis>
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<emphasis italics="true" pageId="0" pageNumber="599">Tropidurus pinima</emphasis>
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and
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<taxonomicName baseAuthorityName="Spix" baseAuthorityYear="1825" class="Reptilia" family="Tropiduridae" genus="Tropidurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropidurus semitaeniatus" order="Squamata" pageId="0" pageNumber="599" phylum="Chordata" rank="species" species="Semitaeniatus">
<emphasis italics="true" pageId="0" pageNumber="599">Tropidurus semitaeniatus</emphasis>
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. Photograph credits: Daniel Mesquita (12.1, 12.6, 12.8), Adrian Garda (12.2), Daniel Passos (12.3, 12.4, 12.7), Marco A. Freitas (12.5).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="599" type="ecological notes">
<paragraph pageId="0" pageNumber="599">Ecological notes.</paragraph>
<paragraph pageId="0" pageNumber="599">
Terrestrial and diurnal. It is a saxico-lous species specializing in rocky areas with crevices (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2930.1.5" author="Passos, D C" journalOrPublisher="Zootaxa" pageId="0" pageNumber="599" pagination="60 - 68" refId="B151" refString="Passos, D C, Lima, D C, Borges-Nojosa, D M, 2011a. A new species of Tropidurus (Squamata, Tropiduridae) of the semitaeniatus group from a semiarid area in Northeastern Brazil. Zootaxa 2930: 60 - 68, DOI: https://doi.org/10.11646/zootaxa.2930.1.5" title="A new species of Tropidurus (Squamata, Tropiduridae) of the semitaeniatus group from a semiarid area in Northeastern Brazil." url="https://doi.org/10.11646/zootaxa.2930.1.5" volume="2930" year="2011 a">Passos et al. 2011a</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1111/aec.12486" author="Pelegrin, N" journalOrPublisher="Austral Ecology" pageId="0" pageNumber="599" pagination="677 - 689" refId="B156" refString="Pelegrin, N, Mesquita, DO, Albinati, P, Caldas, FLS, Cavalcanti, LBQ, Costa, TB, Falico, DA, Galdino, JYA, Tucker, DB, Garda, AA, 2017. Extreme specialization to rocky habitats in Tropidurus lizards from Brazil: Tradeoffs between a fitted ecomorph and autoecology in a harsh environment. Austral Ecology 42: 677 - 689, DOI: https://doi.org/10.1111/aec.12486" title="Extreme specialization to rocky habitats in Tropidurus lizards from Brazil: Tradeoffs between a fitted ecomorph and autoecology in a harsh environment." url="https://doi.org/10.1111/aec.12486" volume="42" year="2017">Pelegrin et al. 2017</bibRefCitation>
). It is found in limestone and sandstone formations and is occasionally found on the ground or in tree trunks (
<bibRefCitation author="Manzani, PR" journalOrPublisher="Herpetologica" pageId="0" pageNumber="599" pagination="462 - 467" refId="B118" refString="Manzani, PR, Abe, AS, 1990. A new species of Tapinurus from the Caatinga of Piaui, northeastern Brazil (Squamata: Tropiduridae). Herpetologica 46: 462 - 467" title="A new species of Tapinurus from the Caatinga of Piaui, northeastern Brazil (Squamata: Tropiduridae)." volume="46" year="1990">Manzani and Abe 1990</bibRefCitation>
). Diet based mainly on arthropods, being
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,
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, and
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the most representative items (
<bibRefCitation DOI="https://doi.org/10.1111/aec.12486" author="Pelegrin, N" journalOrPublisher="Austral Ecology" pageId="0" pageNumber="599" pagination="677 - 689" refId="B156" refString="Pelegrin, N, Mesquita, DO, Albinati, P, Caldas, FLS, Cavalcanti, LBQ, Costa, TB, Falico, DA, Galdino, JYA, Tucker, DB, Garda, AA, 2017. Extreme specialization to rocky habitats in Tropidurus lizards from Brazil: Tradeoffs between a fitted ecomorph and autoecology in a harsh environment. Austral Ecology 42: 677 - 689, DOI: https://doi.org/10.1111/aec.12486" title="Extreme specialization to rocky habitats in Tropidurus lizards from Brazil: Tradeoffs between a fitted ecomorph and autoecology in a harsh environment." url="https://doi.org/10.1111/aec.12486" volume="42" year="2017">Pelegrin et al. 2017</bibRefCitation>
). Oviparous, no detailed data is known about the number of eggs laid by the species, but it could be similar to other
<taxonomicName class="Reptilia" family="Tropiduridae" genus="Tropidurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropidurus" order="Squamata" pageId="0" pageNumber="599" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="599">Tropidurus</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.2307/1564798" author="Vitt, LJ" journalOrPublisher="Journal of Herpetology" pageId="0" pageNumber="599" pagination="79 - 90" refId="B252" refString="Vitt, LJ, 1991. An introduction to the ecology of Cerrado lizards. Journal of Herpetology 25: 79 - 90, DOI: https://doi.org/10.2307/1564798" title="An introduction to the ecology of Cerrado lizards." url="https://doi.org/10.2307/1564798" volume="25" year="1991">Vitt 1991</bibRefCitation>
;
<bibRefCitation author="Avila, RW" journalOrPublisher="Herpetological Review" pageId="0" pageNumber="599" pagination="430 - 433" refId="B16" refString="Avila, RW, Cunha-Avellar, LR, Ferreira, VL, 2008. Diet and reproduction of the lizard Tropidurus etheridgei in rocky areas of central Brazil. Herpetological Review 39: 430 - 433" title="Diet and reproduction of the lizard Tropidurus etheridgei in rocky areas of central Brazil." volume="39" year="2008">
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et al. 2008
</bibRefCitation>
).
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<emphasis id="C6CE9B05FFE2FF8454A05FB7FDCCF836" bold="true" box="[151,571,2008,2034]" pageId="2" pageNumber="51">
Family
<taxonomicName id="33BA3C94FFE2FF8454C15FB7FDCCF836" ID-CoL="7JD" ID-ENA="479187" authority="Zevina, 1978" authorityName="Zevina" authorityYear="1978" box="[246,571,2008,2034]" class="Maxillopoda" family="Calanticidae" kingdom="Animalia" order="Pedunculata" pageId="2" pageNumber="51" phylum="Arthropoda" rank="family">
Calanticidae
<bibRefCitation id="902B3AE6FFE2FF8455AC5FB7FDCCF836" author="Zevina" box="[411,571,2008,2034]" pageId="2" pageNumber="51" refString="Zevina, G. B. (1978) A new classification of the family Scalpellidae Pilsbry (Cirripedia, Thoracica). 1. Subfamilies Lithotryinae, Calanticinae, Pollicipinae, Scalpellinae, Brochiinae and Scalpellopsinae. Zoologichesky Zhurnal, 57 (7), 998 - 1007. (In Russian)." type="journal article" year="1978">Zevina, 1978</bibRefCitation>
</taxonomicName>
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<paragraph id="F4054717FFE3FF8554A058F7FBA1FED8" blockId="3.[151,1436,151,285]" pageId="3" pageNumber="52">
<emphasis id="C6CE9B05FFE3FF8554A058F7FEFCFF74" bold="true" box="[151,267,152,176]" pageId="3" pageNumber="52">Remarks.</emphasis>
The
<taxonomicName id="33BA3C94FFE3FF85557158F8FE25FF6B" box="[326,466,151,175]" class="Maxillopoda" family="Calanticidae" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Calanticidae</taxonomicName>
includes 10 extant and five extinct genera (Newman 1996,
<bibRefCitation id="902B3AE6FFE3FF85504258F7FB00FF6B" author="Jones" box="[1141,1271,151,175]" pageId="3" pageNumber="52" refString="Jones, D. S. (1998) New genus and species of Calanticidae (Cirripedia, Thoracica, Scalpellomorpha) from Australian waters. Zoosystema, 20 (2), 239 - 253." type="journal article" year="1998">Jones 1998</bibRefCitation>
,
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), the most generalized of which,
<taxonomicName id="33BA3C94FFE3FF85563658D3FD7FFF10" box="[513,648,188,212]" class="Maxillopoda" family="Calanticidae" genus="Scillaelepas" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFE3FF85563658D3FD7FFF10" box="[513,648,188,212]" italics="true" pageId="3" pageNumber="52">Scillaelepas</emphasis>
</taxonomicName>
, was known from the fossil record (
<bibRefCitation id="902B3AE6FFE3FF85501A58D3FB15FF10" author="Seguenza" box="[1069,1250,188,212]" pageId="3" pageNumber="52" refString="Seguenza, G. (1876) Ricerche paleontologiche intorno ai Cirripedi terziari della provincia di Messina. ParteII. Terza famiglia Lepadi Darwin. Atti dell'Accademia Pontoniana, Naples, 10, 1 - 112." type="journal article" year="1876">Seguenza, 1876</bibRefCitation>
) before a living representative was discovered in the deep-sea (
<bibRefCitation id="902B3AE6FFE3FF85568558B0FC8EFF33" author="Aurivillius" box="[690,889,223,247]" pageId="3" pageNumber="52" refString="Aurivillius, C. W. S. (1894) Studien uber Cirripeden. Kongliga Svenska Vetenskaps Akademiens Handlingar, Uppsala, 26 (7), 5 - 107 + 9 pls." type="journal article" year="1894">Aurivillius, 1894</bibRefCitation>
).
<collectingCountry id="8CAD0787FFE3FF8557B9588FFC5AFF3C" box="[910,941,224,248]" name="American Samoa" pageId="3" pageNumber="52">As</collectingCountry>
will be seen below, the calanticid reported upon herein belongs to
<emphasis id="C6CE9B05FFE3FF8555AA596BFDE4FED8" box="[413,531,260,284]" italics="true" pageId="3" pageNumber="52">
<taxonomicName id="33BA3C94FFE3FF8555AA596BFDF9FED8" box="[413,526,260,284]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">Calantica</taxonomicName>
,
</emphasis>
a genus that has diverged little from the stem form.
</paragraph>
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<heading id="AF4DF07BFFE3FF8554A05E80FE6CF8CD" bold="true" box="[151,411,1775,1801]" fontSize="11" level="1" pageId="3" pageNumber="52" reason="1">
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<emphasis id="C6CE9B05FFE3FF8554A05E80FE6CF8CD" bold="true" box="[151,411,1775,1801]" pageId="3" pageNumber="52">
<emphasis id="C6CE9B05FFE3FF8554A05E80FEFAF8CD" bold="true" box="[151,269,1775,1801]" italics="true" pageId="3" pageNumber="52">Calantica</emphasis>
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</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="BCA0149CFFE3FF8554A05F5AFE84F85F" pageId="3" pageNumber="52" type="diagnosis">
<paragraph id="F4054717FFE3FF8554A05F5AFE84F85F" blockId="3.[151,1436,1845,2021]" pageId="3" pageNumber="52">
<emphasis id="C6CE9B05FFE3FF8554A05F5AFEE6F88A" bold="true" box="[151,273,1845,1870]" pageId="3" pageNumber="52">Diagnosis.</emphasis>
<taxonomicName id="33BA3C94FFE3FF85552C5F59FE7EF88A" box="[283,393,1846,1870]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFE3FF85552C5F59FE7EF88A" box="[283,393,1846,1870]" italics="true" pageId="3" pageNumber="52">Calantica</emphasis>
</taxonomicName>
(
<typeStatus id="2B01F9B5FFE3FF8555AC5F59FE38F88A" box="[411,463,1846,1870]" pageId="3" pageNumber="52">type</typeStatus>
,
<taxonomicName id="33BA3C94FFE3FF8555EA5F59FCB3F88A" authority="Leach, 1824" authorityName="Leach" authorityYear="1824" box="[477,836,1846,1870]" class="Maxillopoda" family="Pollicipedidae" genus="Pollicipes" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="villosus">
<emphasis id="C6CE9B05FFE3FF8555EA5F59FD5CF88A" box="[477,683,1846,1870]" italics="true" pageId="3" pageNumber="52">Pollicipes villosus</emphasis>
<bibRefCitation id="902B3AE6FFE3FF8556825F59FCB3F88A" author="Leach" box="[693,836,1846,1870]" pageId="3" pageNumber="52" refString="Leach, W. E. (1824) Cirripedes. Supplement to the 4 th, 5 t and 6 th editions of the Encyclopaedia Britannica, 3 (1), 168 - 171 + Pl. 57." type="journal article" year="1824">Leach, 1824</bibRefCitation>
</taxonomicName>
) is limited to species with the same basic 13-plated pattern of
<taxonomicName id="33BA3C94FFE3FF85553E5F32FE67F8B1" box="[265,400,1885,1909]" class="Maxillopoda" family="Calanticidae" genus="Scillaelepas" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFE3FF85553E5F32FE67F8B1" box="[265,400,1885,1909]" italics="true" pageId="3" pageNumber="52">Scillaelepas</emphasis>
</taxonomicName>
, sometimes supplemented by first a subrostrum (SR1), a second latus (L2) or a second carinolatus (CL2) (Fig. 6).
</paragraph>
</subSubSection>
<subSubSection id="BCA0149CFFE3FF8154F25FC7FACEF837" lastPageId="7" lastPageNumber="56" pageId="3" pageNumber="52" type="discussion">
<paragraph id="F4054717FFE3FF8254F25FC7FDE9FF33" blockId="3.[151,1436,1845,2021]" lastBlockId="4.[151,1436,151,576]" lastPageId="4" lastPageNumber="53" pageId="3" pageNumber="52">
<emphasis id="C6CE9B05FFE3FF8554F25FC7FECEF804" bold="true" box="[197,313,1960,1984]" pageId="3" pageNumber="52">Remarks.</emphasis>
<taxonomicName id="33BA3C94FFE3FF8555765FC7FE58F804" box="[321,431,1960,1984]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFE3FF8555765FC7FE58F804" box="[321,431,1960,1984]" italics="true" pageId="3" pageNumber="52">Calantica</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="33BA3C94FFE3FF85569A5FC7FCC3F804" box="[685,820,1960,1984]" class="Maxillopoda" family="Calanticidae" genus="Scillaelepas" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFE3FF85569A5FC7FCC3F804" box="[685,820,1960,1984]" italics="true" pageId="3" pageNumber="52">Scillaelepas</emphasis>
</taxonomicName>
in having small peduncular scales covered with a distinct cuticle, and both genera are distinguished from the next higher grade,
<emphasis id="C6CE9B05FFE3FF8557CD5FA3FBAAF820" box="[1018,1117,1996,2020]" italics="true" pageId="3" pageNumber="52">
<taxonomicName id="33BA3C94FFE3FF8557CD5FA3FBAEF820" box="[1018,1113,1996,2020]" class="Maxillopoda" family="Scalpellidae" genus="Smilium" kingdom="Animalia" order="Pedunculata" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">Smilium</taxonomicName>
,
</emphasis>
in not having the latus (L), reduced or conspicuously elevated above the carinolatus (CL). There are borderline species between
<taxonomicName id="33BA3C94FFE4FF8250C958F7FA9AFF74" box="[1278,1389,152,176]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFE4FF8250C958F7FA9AFF74" box="[1278,1389,152,176]" italics="true" pageId="4" pageNumber="53">Calantica</emphasis>
</taxonomicName>
and
<taxonomicName id="33BA3C94FFE4FF8254A058D3FF04FF10" box="[151,243,188,212]" class="Maxillopoda" family="Scalpellidae" genus="Smilium" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFE4FF8254A058D3FF04FF10" box="[151,243,188,212]" italics="true" pageId="4" pageNumber="53">Smilium</emphasis>
</taxonomicName>
over which there will be disagreements until some revisionary breakthrough has been made, but fortunately they do not concern us here.
</paragraph>
<paragraph id="F4054717FFE4FF8254F2596BFF00FDDE" blockId="4.[151,1436,151,576]" pageId="4" pageNumber="53">
While Zevinas (1981) monograph on the scalpellomorphs includes 13 species of
<taxonomicName id="33BA3C94FFE4FF825065596BFA9FFED8" authority="Young (2003)" authorityName="Young" authorityYear="2003" box="[1106,1384,260,285]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFE4FF825065596BFB37FED8" box="[1106,1216,260,284]" italics="true" pageId="4" pageNumber="53">Calantica</emphasis>
,
<bibRefCitation id="902B3AE6FFE4FF8250F9596AFA9FFED8" author="Young" box="[1230,1384,260,285]" pageId="4" pageNumber="53" refString="Young, P. S. (2003) Redescription of the calanticids (Cirripedia, Scalpellomorpha) described by Wilhelm Weltner. Zoosystematics and Evolution, 79 (1), 181 - 201." type="journal article" year="2003">Young (2003)</bibRefCitation>
</taxonomicName>
lists 15 species after the inadvertent duplication of
<taxonomicName id="33BA3C94FFE4FF8256A95947FCD8FE84" box="[670,815,296,320]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="moskalevi">
<emphasis id="C6CE9B05FFE4FF8256A95947FCD8FE84" box="[670,815,296,320]" italics="true" pageId="4" pageNumber="53">C. moskalevi</emphasis>
</taxonomicName>
is removed. The list can arguably be further reduced to 11 by transfer of
<taxonomicName id="33BA3C94FFE4FF8255615922FDA8FEA0" authority="Ren, 1989" authorityName="Ren" authorityYear="1989" box="[342,607,332,356]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="flagellata">
<emphasis id="C6CE9B05FFE4FF8255615922FE15FEA0" box="[342,482,332,356]" italics="true" pageId="4" pageNumber="53">C. flagellata</emphasis>
<bibRefCitation id="902B3AE6FFE4FF8255DD5922FDA8FEA0" author="Ren" box="[490,607,332,356]" pageId="4" pageNumber="53" refString="Ren, X. (1989) On a collection of Cirripedia Thoracica from Madagascar and adjacent waters. Bulletin du Museum national d'Histoire naturelle, Paris, (4), 11, A, (2), 431 - 468." type="journal article" year="1989">Ren, 1989</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="33BA3C94FFE4FF82565C5922FC13FEA0" authority="Broch, 1931" authorityName="Broch" authorityYear="1931" box="[619,996,332,356]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="pedunculostriata">
<emphasis id="C6CE9B05FFE4FF82565C5922FCA7FEA0" box="[619,848,332,356]" italics="true" pageId="4" pageNumber="53">C. pedunculostriata,</emphasis>
<bibRefCitation id="902B3AE6FFE4FF8257605923FC13FEA0" author="Broch" box="[855,996,332,356]" pageId="4" pageNumber="53" refString="Broch, H. (1931) Papers from Dr Th. Mortensen's Pacific Expedition 1914 - 1916. LVI. Indomalayan Cirripedia. Videnskabelige Meddelelser fra Dansk naturhistorisk Forening i Kobenhaven, 91, 1 - 146." type="journal article" year="1931">Broch, 1931</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="33BA3C94FFE4FF8257D85922FA6BFEA0" authority="Quoy and Gaimard, 1834" authorityName="Quoy and Gaimard" authorityYear="1834" box="[1007,1436,332,356]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="spinosa">
<emphasis id="C6CE9B05FFE4FF8257D85922FB91FEA0" box="[1007,1126,333,356]" italics="true" pageId="4" pageNumber="53">C. spinosa</emphasis>
(
<bibRefCitation id="902B3AE6FFE4FF8250435923FA63FEA0" author="Quoy" box="[1140,1428,332,356]" pageId="4" pageNumber="53" refString="Quoy, J. R. C. &amp; Gaimard, J. P. (1834) Voyage de decouvertes de l'Astrolabe, execut par ordre du Roi, pendant les annees 1826 - 29, sous le commandement de M. J. Dumond d'Urville. Zoologie, Mollusca, 3, Paris. 712 pp." type="book" year="1834">Quoy and Gaimard, 1834</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="33BA3C94FFE4FF8254FF591FFE0EFE43" authority="Foster, 1979" authorityName="Foster" authorityYear="1979" box="[200,505,367,392]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="spinilatera">
<emphasis id="C6CE9B05FFE4FF8254FF591FFE94FE4C" box="[200,355,368,392]" italics="true" pageId="4" pageNumber="53">C. spinilatera</emphasis>
<bibRefCitation id="902B3AE6FFE4FF82555B591FFE0EFE43" author="Foster" box="[364,505,367,391]" pageId="4" pageNumber="53" refString="Foster, B. A. (1979) The Marine Fauna of New Zealand; Barnacles (Cirripedia: Thoracica). New Zealand Oceanographic Institute Memoir, 69, 1 - 159 (1978)." type="journal article" year="1979">Foster, 1979</bibRefCitation>
</taxonomicName>
to
<taxonomicName id="33BA3C94FFE4FF825617591FFD8BFE4C" box="[544,636,368,392]" class="Maxillopoda" family="Scalpellidae" genus="Smilium" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFE4FF825617591FFD8BFE4C" box="[544,636,368,392]" italics="true" pageId="4" pageNumber="53">Smilium</emphasis>
</taxonomicName>
or elsewhere, and then increased to 14 when
<taxonomicName id="33BA3C94FFE4FF8250BF591FFF2FFE68" authority="Annandale, 1909" authorityName="Annandale" authorityYear="1909" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="kampeni">
<emphasis id="C6CE9B05FFE4FF8250BF591FFAFDFE4C" box="[1160,1290,368,392]" italics="true" pageId="4" pageNumber="53">C. kampeni</emphasis>
(
<bibRefCitation id="902B3AE6FFE4FF82512D5900FF27FE68" author="Annandale" pageId="4" pageNumber="53" refString="Annandale, N. (1909) Description of a barnacle of the genus Scalpellum from Malaysia. Records of the Indian Museum, 3 (3), 267 - 270." type="journal article" year="1909">Annandale, 1909</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="33BA3C94FFE4FF82553959FAFDABFE68" authority="Hoek, 1907" authorityName="Hoek" authorityYear="1907" box="[270,604,404,428]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="pollicipoides">
<emphasis id="C6CE9B05FFE4FF82553959FAFE37FE68" box="[270,448,404,428]" italics="true" pageId="4" pageNumber="53">C. pollicipoides</emphasis>
(
<bibRefCitation id="902B3AE6FFE4FF8255F959FBFDA4FE68" author="Hoek" box="[462,595,404,428]" pageId="4" pageNumber="53" refString="Hoek, P. P. C. (1907) Cirripedia of the Siboga-Expedition. A. Cirripedia Pedunculata. Siboga-Expeditie, 31 A, 1 - 127 + pls. 1 - 10. Brill, Leyden." type="journal article" year="1907">Hoek, 1907</bibRefCitation>
)
</taxonomicName>
are returned from
<taxonomicName id="33BA3C94FFE4FF82571A59FBFC7EFE68" box="[813,905,404,428]" class="Maxillopoda" family="Scalpellidae" genus="Smilium" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFE4FF82571A59FBFC7EFE68" box="[813,905,404,428]" italics="true" pageId="4" pageNumber="53">Smilium</emphasis>
</taxonomicName>
and
<taxonomicName id="33BA3C94FFE4FF82578859FAFACEFE68" authority="Jones and Hosie, 2009" authorityName="Jones and Hosie" authorityYear="2009" box="[959,1337,404,428]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="darwini">
<emphasis id="C6CE9B05FFE4FF82578859FAFBC1FE68" box="[959,1078,404,428]" italics="true" pageId="4" pageNumber="53">C. darwini</emphasis>
<bibRefCitation id="902B3AE6FFE4FF82500A59FAFACEFE68" author="Jones" box="[1085,1337,404,428]" pageId="4" pageNumber="53" refString="Jones, D. S. &amp; Hosie, A. M. (2009) A new species of Calantica from Western Australian waters (Thoracica: Scalpellomorpha: Calanticidae). Records of the Western Australian Museum, 25, 239 - 246." type="journal article" year="2009">Jones and Hosie, 2009</bibRefCitation>
</taxonomicName>
is added (see below). Fortunately, “borderline” cases do not concern us here; to the contrary, the Axial form is a true albeit a relatively generalized
<taxonomicName id="33BA3C94FFE4FF8255AE59B0FDF0FE33" box="[409,519,479,503]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFE4FF8255AE59B0FDF0FE33" box="[409,519,479,503]" italics="true" pageId="4" pageNumber="53">Calantica</emphasis>
</taxonomicName>
, especially when, and then apparently more commonly than not, CL2s fail to develop.
</paragraph>
<paragraph id="F4054717FFE4FF8254F05A47FB3DFD84" blockId="4.[151,1436,151,576]" box="[199,1226,551,576]" pageId="4" pageNumber="53">
For convenience, the 14 species of
<taxonomicName id="33BA3C94FFE4FF8256665A48FD48FDFB" box="[593,703,551,575]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFE4FF8256665A48FD48FDFB" box="[593,703,551,575]" italics="true" pageId="4" pageNumber="53">Calantica</emphasis>
</taxonomicName>
can be divided into the following four groups:
</paragraph>
<paragraph id="F4054717FFE4FF8254A05A1FFAA1FD43" blockId="4.[151,1366,623,647]" box="[151,1366,623,647]" pageId="4" pageNumber="53">
I. Species with only the basic 13 capitular plates of which
<taxonomicName id="33BA3C94FFE4FF82570B5A1FFC5CFD43" box="[828,939,623,647]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="studeri">
<emphasis id="C6CE9B05FFE4FF82570B5A1FFC5CFD43" box="[828,939,623,647]" italics="true" pageId="4" pageNumber="53">C. studeri</emphasis>
</taxonomicName>
appears most similar to
<taxonomicName id="33BA3C94FFE4FF82508A5A1FFAB9FD43" box="[1213,1358,623,647]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="moskalevi">
<emphasis id="C6CE9B05FFE4FF82508A5A1FFAB9FD43" box="[1213,1358,623,647]" italics="true" pageId="4" pageNumber="53">C. moskalevi</emphasis>
</taxonomicName>
:
</paragraph>
<paragraph id="F4054717FFE4FF8254A05AD8FC8AFD0B" blockId="4.[151,1436,695,1042]" box="[151,893,695,719]" pageId="4" pageNumber="53">
1)
<taxonomicName id="33BA3C94FFE4FF8254F05AD7FE0CFD0B" authority="Broch, 1922: 232" authorityName="Broch" authorityPageNumber="232" authorityYear="1922" box="[199,507,695,719]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="C6CE9B05FFE4FF8254F05AD7FEDAFD0B" box="[199,301,695,719]" italics="true" pageId="4" pageNumber="53">C. affinis</emphasis>
<bibRefCitation id="902B3AE6FFE4FF8255035AD8FE34FD0B" author="Broch" box="[308,451,695,719]" pageId="4" pageNumber="53" refString="Broch, H. (1922) Studies on Pacific cirripeds. In: Papers from Dr. Th. Mortensen's Pacific Expedition 1914 - 1916. X. Videnskabelige meddelelser fra Dansk Naturhistorisk Forening i Kobenhaven, 73, 215 - 358 + 559 (correction)." type="journal article" year="1922">Broch, 1922</bibRefCitation>
: 232
</taxonomicName>
,
<collectingCountry id="8CAD0787FFE4FF8256315AD8FD73FD0B" box="[518,644,695,719]" name="Philippines" pageId="4" pageNumber="53">Philippines</collectingCountry>
(Zamboanga),
<specimenCount id="E2BC8C9EFFE4FF8257185AD8FC8AFD0B" box="[815,893,695,719]" pageId="4" pageNumber="53" type="generic">
<quantity id="3342EAF2FFE4FF8257185AD8FC8EFD0B" box="[815,889,695,719]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="4" pageNumber="53" unit="m" value="350.0">350 m</quantity>
.
</specimenCount>
</paragraph>
<paragraph id="F4054717FFE4FF8254A05AB5FD76FD36" blockId="4.[151,1436,695,1042]" box="[151,641,730,755]" pageId="4" pageNumber="53">
2)
<taxonomicName id="33BA3C94FFE4FF8254F05AB4FE2FFD36" authority="Pilsbry, 1907: 7" authorityName="Pilsbry" authorityPageNumber="7" authorityYear="1907" box="[199,472,730,755]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="eos">
<emphasis id="C6CE9B05FFE4FF8254F05AB4FEFAFD37" box="[199,269,731,755]" italics="true" pageId="4" pageNumber="53">C. eos</emphasis>
(
<bibRefCitation id="902B3AE6FFE4FF82552C5AB5FE38FD36" author="Pilsbry" box="[283,463,730,754]" pageId="4" pageNumber="53" refString="Pilsbry, H. A. (1907) The barnacles (Cirripedia) contained in the collections of the U. S. National Museum. Bulletin of the U. S. National Museum, 60, 1 - 122 + pls 1 - 11." type="journal article" year="1907">Pilsbry, 1907: 7</bibRefCitation>
)
</taxonomicName>
,
<collectingCountry id="8CAD0787FFE4FF8255D25AB4FDDFFD36" box="[485,552,731,754]" name="Japan" pageId="4" pageNumber="53">Japan</collectingCountry>
,
<specimenCount id="E2BC8C9EFFE4FF8256045AB5FD76FD36" box="[563,641,730,754]" pageId="4" pageNumber="53" type="generic">
<quantity id="3342EAF2FFE4FF8256045AB5FD8BFD36" box="[563,636,730,754]" metricMagnitude="2" metricUnit="m" metricValue="1.33" pageId="4" pageNumber="53" unit="m" value="133.0">133 m</quantity>
.
</specimenCount>
</paragraph>
<paragraph id="F4054717FFE4FF8254A05A90FC53FCD3" blockId="4.[151,1436,695,1042]" box="[151,932,767,791]" pageId="4" pageNumber="53">
3)
<taxonomicName id="33BA3C94FFE4FF8254F05B6FFD8AFCD3" authority="Jones and Hosie, 2009: 240" authorityName="Jones and Hosie" authorityPageNumber="240" authorityYear="2009" box="[199,637,767,791]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="darwini">
<emphasis id="C6CE9B05FFE4FF8254F05B6FFEC9FCD3" box="[199,318,767,791]" italics="true" pageId="4" pageNumber="53">C. darwini</emphasis>
<bibRefCitation id="902B3AE6FFE4FF8255725B6FFDB2FCD3" author="Jones" box="[325,581,767,791]" pageId="4" pageNumber="53" refString="Jones, D. S. &amp; Hosie, A. M. (2009) A new species of Calantica from Western Australian waters (Thoracica: Scalpellomorpha: Calanticidae). Records of the Western Australian Museum, 25, 239 - 246." type="journal article" year="2009">Jones and Hosie, 2009</bibRefCitation>
: 240
</taxonomicName>
, W
<collectingCountry id="8CAD0787FFE4FF82569E5A90FCE4FCD3" box="[681,787,767,791]" name="Australia" pageId="4" pageNumber="53">Australia</collectingCountry>
,
<quantity id="3342EAF2FFE4FF8257295A90FC57FCD3" box="[798,928,767,791]" metricMagnitude="2" metricUnit="m" metricValue="1.51" metricValueMax="1.56" metricValueMin="1.46" pageId="4" pageNumber="53" unit="m" value="151.0" valueMax="156.0" valueMin="146.0">
146
<specimenCount id="E2BC8C9EFFE4FF8257605A90FC57FCD3" box="[855,928,767,791]" pageId="4" pageNumber="53" type="generic">156 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph id="F4054717FFE4FF8254A05B4DFCDBFCFE" blockId="4.[151,1436,695,1042]" box="[151,812,802,827]" pageId="4" pageNumber="53">
4)
<taxonomicName id="33BA3C94FFE4FF8254F05B4CFDF9FCFE" authority="Rosell, 1991: 13" authorityName="Rosell" authorityPageNumber="13" authorityYear="1991" box="[199,526,802,827]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="graphica">
<emphasis id="C6CE9B05FFE4FF8254F05B4CFEBBFCFF" box="[199,332,803,827]" italics="true" pageId="4" pageNumber="53">C. graphica</emphasis>
<bibRefCitation id="902B3AE6FFE4FF8255635B4DFE12FCFE" author="Rosell" box="[340,485,802,826]" pageId="4" pageNumber="53" refString="Rosell, N. C. (1991) Crustacea Cirripedia Thoracica: MUSORSTOM 3 Philippines collection. In: Crosnier, A. (ed.) Resultats des Campagnes MUSORSTOM 9. Memoires du Museum national d'Histoire naturelle, (A), 152, 6 - 61." type="journal article" year="1991">Rosell, 1991</bibRefCitation>
: 13
</taxonomicName>
,
<collectingCountry id="8CAD0787FFE4FF82562D5B4DFD6CFCFE" box="[538,667,802,826]" name="Philippines" pageId="4" pageNumber="53">Philippines</collectingCountry>
,
<quantity id="3342EAF2FFE4FF8256915B4DFCDFFCFE" box="[678,808,802,826]" metricMagnitude="2" metricUnit="m" metricValue="1.9899999999999998" metricValueMax="2.05" metricValueMin="1.93" pageId="4" pageNumber="53" unit="m" value="199.0" valueMax="205.0" valueMin="193.0">
193
<specimenCount id="E2BC8C9EFFE4FF8256EA5B4DFCDFFCFE" box="[733,808,802,826]" pageId="4" pageNumber="53" type="generic">205 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph id="F4054717FFE4FF8254A05B28FC74FC9B" blockId="4.[151,1436,695,1042]" box="[151,899,839,863]" pageId="4" pageNumber="53">
5)
<taxonomicName id="33BA3C94FFE4FF8254F05B27FDF5FC9B" authority="Hiro, 1932: 473" authorityName="Hiro" authorityPageNumber="473" authorityYear="1932" box="[199,514,839,863]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="kruegeri">
<emphasis id="C6CE9B05FFE4FF8254F05B27FEB1FC9B" box="[199,326,839,863]" italics="true" pageId="4" pageNumber="53">C. kruegeri</emphasis>
<bibRefCitation id="902B3AE6FFE4FF82557A5B28FE3CFC9B" author="Hiro" box="[333,459,839,863]" pageId="4" pageNumber="53" refString="Hiro, F. (1932) Report on the Japanese species of the genus Calantica (Cirripedia). Annotationes Zoologicae, Japon, 13 (5), 467 - 482." type="journal article" year="1932">Hiro, 1932</bibRefCitation>
: 473
</taxonomicName>
,
<collectingCountry id="8CAD0787FFE4FF82563A5B27FDBAFC9B" box="[525,589,840,863]" name="Japan" pageId="4" pageNumber="53">Japan</collectingCountry>
(Sagami Bay),
<quantity id="3342EAF2FFE4FF8256CA5B28FC88FC9B" box="[765,895,839,863]" metricMagnitude="2" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="4" pageNumber="53" unit="m" value="165.0" valueMax="180.0" valueMin="150.0">
150
<specimenCount id="E2BC8C9EFFE4FF8257015B28FC88FC9B" box="[822,895,839,863]" pageId="4" pageNumber="53" type="generic">180 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph id="F4054717FFE4FF8254A05B05FC8CFC46" blockId="4.[151,1436,695,1042]" box="[151,891,874,899]" pageId="4" pageNumber="53">
6)
<taxonomicName id="33BA3C94FFE4FF8254F05B04FDEEFC46" authority="Utinomi, 1970: 159" authorityName="Utinomi" authorityPageNumber="159" authorityYear="1970" box="[199,537,874,899]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="pusilla">
<emphasis id="C6CE9B05FFE4FF8254F05B04FEC4FC47" box="[199,307,875,899]" italics="true" pageId="4" pageNumber="53">C. pusilla</emphasis>
<bibRefCitation id="902B3AE6FFE4FF82550C5B05FE15FC46" author="Utinomi" box="[315,482,874,898]" pageId="4" pageNumber="53" refString="Utinomi, H. (1970) New and rare commensal pedunculate cirripeds from Amakusa Islands, Western Kyusyu, Japan. Publications of the Seto Marine Biological Laboratory, 18 (3), 157 - 67." type="journal article" year="1970">Utinomi, 1970</bibRefCitation>
: 159
</taxonomicName>
,
<collectingCountry id="8CAD0787FFE4FF8256125B04FD92FC46" box="[549,613,875,898]" name="Japan" pageId="4" pageNumber="53">Japan</collectingCountry>
(Amakusa Is),
<quantity id="3342EAF2FFE4FF8257265B05FC80FC46" box="[785,887,874,898]" metricMagnitude="1" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="4" pageNumber="53" unit="m" value="27.5" valueMax="30.0" valueMin="25.0">
25
<specimenCount id="E2BC8C9EFFE4FF82570C5B05FC80FC46" box="[827,887,874,898]" pageId="4" pageNumber="53" type="generic">30 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph id="F4054717FFE4FF8254A05BE0FE4EFC0E" blockId="4.[151,1436,695,1042]" pageId="4" pageNumber="53">
7)
<taxonomicName id="33BA3C94FFE4FF8254F05BFFFDD1FC63" authority="Weltner, 1922: 100" authorityName="Weltner" authorityPageNumber="100" authorityYear="1922" box="[199,550,911,935]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="studeri">
<emphasis id="C6CE9B05FFE4FF8254F05BFFFEC1FC63" box="[199,310,911,935]" italics="true" pageId="4" pageNumber="53">C. studeri</emphasis>
(
<bibRefCitation id="902B3AE6FFE4FF8255735BE0FDE9FC63" author="Weltner" box="[324,542,911,935]" pageId="4" pageNumber="53" refString="Weltner, W. (1922) Cirripedia der Deutschen Tiefsee-Expedition, Wissenschafteliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot; 1898 - 1899, 23 (2), 69 - 113 + pls. 2 - 4. Jena." type="journal article" year="1922">Weltner, 1922: 100</bibRefCitation>
)
</taxonomicName>
,
<bibRefCitation id="902B3AE6FFE4FF8256055BFFFD4CFC63" author="Foster" box="[562,699,911,935]" pageId="4" pageNumber="53" refString="Foster, B. A. (1979) The Marine Fauna of New Zealand; Barnacles (Cirripedia: Thoracica). New Zealand Oceanographic Institute Memoir, 69, 1 - 159 (1978)." type="journal article" year="1979">Foster 1979</bibRefCitation>
: 45, NW
<collectingCountry id="8CAD0787FFE4FF8257115BE0FC7AFC63" box="[806,909,911,935]" name="Australia" pageId="4" pageNumber="53">Australia</collectingCountry>
and
<collectingCountry id="8CAD0787FFE4FF8257F35BFFFBAAFC63" box="[964,1117,911,935]" name="New Zealand" pageId="4" pageNumber="53">New Zealand</collectingCountry>
,
<quantity id="3342EAF2FFE4FF82505F5BE0FB2BFC63" box="[1128,1244,911,935]" metricMagnitude="2" metricUnit="m" metricValue="1.54" metricValueMax="2.48" metricValueMin="0.6" pageId="4" pageNumber="53" unit="m" value="154.0" valueMax="248.0" valueMin="60.0">60248 m</quantity>
; cf.
<bibRefCitation id="902B3AE6FFE4FF82513C5BFFFA6FFC63" author="Young" box="[1291,1432,911,936]" pageId="4" pageNumber="53" refString="Young, P. S. (2003) Redescription of the calanticids (Cirripedia, Scalpellomorpha) described by Wilhelm Weltner. Zoosystematics and Evolution, 79 (1), 181 - 201." type="journal article" year="2003">Young 2003</bibRefCitation>
: 192 for redescription.
</paragraph>
<paragraph id="F4054717FFE4FF8254A05BB8FC53FBD6" blockId="4.[151,1436,695,1042]" pageId="4" pageNumber="53">
8)
<taxonomicName id="33BA3C94FFE4FF8254F05BB7FDD6FC2B" authority="Hoek, 1883: 72" authorityName="Hoek" authorityPageNumber="72" authorityYear="1883" box="[199,545,983,1007]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="trispinosa">
<emphasis id="C6CE9B05FFE4FF8254F05BB7FEA0FC2B" box="[199,343,984,1007]" italics="true" pageId="4" pageNumber="53">C. trispinosa</emphasis>
(
<bibRefCitation id="902B3AE6FFE4FF8255515BB8FDEEFC2B" author="Hoek" box="[358,537,983,1007]" pageId="4" pageNumber="53" refString="Hoek, P. P. C. (1883) Report on the Cirripedia collected by H. M. S. Challenger during the years 1873 - 1876. In: Thomson, W. and Murray, J. (eds), Report of the Scientific Results of the Voyage of H. M. S. Challenger, Zoology, 8 (25), 1 - 169 + pls. 1 - 13." type="journal article" year="1883">Hoek, 1883: 72</bibRefCitation>
)
</taxonomicName>
; cf.
<bibRefCitation id="902B3AE6FFE4FF8256675BB8FD2CFC2B" author="Rosell" box="[592,731,983,1007]" pageId="4" pageNumber="53" refString="Rosell, N. C. (1991) Crustacea Cirripedia Thoracica: MUSORSTOM 3 Philippines collection. In: Crosnier, A. (ed.) Resultats des Campagnes MUSORSTOM 9. Memoires du Museum national d'Histoire naturelle, (A), 152, 6 - 61." type="journal article" year="1991">Rosell 1991</bibRefCitation>
: 14,
<collectingCountry id="8CAD0787FFE4FF8257385BB8FC67FC2B" box="[783,912,983,1007]" name="Philippines" pageId="4" pageNumber="53">Philippines</collectingCountry>
,
<quantity id="3342EAF2FFE4FF8257AC5BB8FC08FC2B" box="[923,1023,983,1007]" metricMagnitude="1" metricUnit="m" metricValue="8.35" metricValueMax="8.5" metricValueMin="8.2" pageId="4" pageNumber="53" unit="m" value="83.5" valueMax="85.0" valueMin="82.0">8285 m</quantity>
, Java Sea to Sagami Bay,
<quantity id="3342EAF2FFE4FF8251145BB8FA60FC2B" box="[1315,1431,983,1007]" metricMagnitude="2" metricUnit="m" metricValue="1.28" metricValueMax="2.0" metricValueMin="0.5599999999999999" pageId="4" pageNumber="53" unit="m" value="128.0" valueMax="200.0" valueMin="56.0">56200 m</quantity>
;
<bibRefCitation id="902B3AE6FFE4FF8254F05B94FEBFFBD6" author="Jones" box="[199,328,1018,1042]" pageId="4" pageNumber="53" refString="Jones, D. S. (1998) New genus and species of Calanticidae (Cirripedia, Thoracica, Scalpellomorpha) from Australian waters. Zoosystema, 20 (2), 239 - 253." type="journal article" year="1998">Jones 1998</bibRefCitation>
: 247, Gulf of
<collectingCountry id="8CAD0787FFE4FF8255D45B95FDE8FBD6" box="[483,543,1018,1042]" name="Thailand" pageId="4" pageNumber="53">Siam</collectingCountry>
, S.
<collectingCountry id="8CAD0787FFE4FF82567E5B95FD7BFBD6" box="[585,652,1018,1042]" name="China" pageId="4" pageNumber="53">China</collectingCountry>
Sea to
<collectingCountry id="8CAD0787FFE4FF8256E85B94FCD5FBD6" box="[735,802,1019,1042]" name="Japan" pageId="4" pageNumber="53">Japan</collectingCountry>
,
<quantity id="3342EAF2FFE4FF82571B5B95FC57FBD6" box="[812,928,1018,1042]" metricMagnitude="2" metricUnit="m" metricValue="1.775" metricValueMax="3.2" metricValueMin="0.35" pageId="4" pageNumber="53" unit="m" value="177.5" valueMax="320.0" valueMin="35.0">
35
<specimenCount id="E2BC8C9EFFE4FF8257615B95FC57FBD6" box="[854,928,1018,1042]" pageId="4" pageNumber="53" type="generic">320 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph id="F4054717FFE4FF8254A05C29FC02FB99" blockId="4.[151,1013,1090,1117]" box="[151,1013,1090,1117]" pageId="4" pageNumber="53">II. Species with CL2 sometimes appearing carinal of L on one or both sides.</paragraph>
<paragraph id="F4054717FFE4FF8254A05CE2FC57FB61" blockId="4.[151,928,1165,1189]" box="[151,928,1165,1189]" pageId="4" pageNumber="53">
9)
<taxonomicName id="33BA3C94FFE4FF8254F05CE1FD47FB61" authority="Zevina and Galkin, 1989: 134" authorityName="Zevina and Galkin" authorityPageNumber="134" authorityYear="1989" box="[199,688,1165,1189]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="moskalevi">
<emphasis id="C6CE9B05FFE4FF8254F05CE1FF2DFB61" box="[199,218,1166,1189]" italics="true" pageId="4" pageNumber="53">C</emphasis>
.
<emphasis id="C6CE9B05FFE4FF8254D05CE2FEA0FB61" box="[231,343,1165,1189]" italics="true" pageId="4" pageNumber="53">moskalevi</emphasis>
<bibRefCitation id="902B3AE6FFE4FF8255695CE2FD8EFB61" author="Zevina" box="[350,633,1165,1189]" pageId="4" pageNumber="53" refString="Zevina, G. B. &amp; Galkin, S. V. (1989) New species of cirripeds (Cirripedia, Thoracica) from thermal waters. Zoologicheskii Zhurnal, 68 (3), 134 - 136." type="journal article" year="1989">Zevina and Galkin, 1989</bibRefCitation>
: 134
</taxonomicName>
. NE Pacific,
<quantity id="3342EAF2FFE4FF8257745CE2FC6CFB61" box="[835,923,1165,1189]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="53" unit="m" value="1400.0">1400 m</quantity>
.
</paragraph>
<paragraph id="F4054717FFE4FF8254A05CB9FCE6FB29" blockId="4.[151,785,1237,1261]" box="[151,785,1237,1261]" pageId="4" pageNumber="53">III. Species with the basic 13-plate pattern +SR include:</paragraph>
<paragraph id="F4054717FFE4FF8254A05D72FCBFFAF1" blockId="4.[151,1436,1309,1406]" box="[151,840,1309,1333]" pageId="4" pageNumber="53">
10)
<taxonomicName id="33BA3C94FFE4FF8254F05D71FDB7FAF1" authority="Hiro, 1932: 469" authorityName="Hiro" authorityPageNumber="469" authorityYear="1932" box="[199,576,1309,1333]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="quinquelatera">
<emphasis id="C6CE9B05FFE4FF8254F05D71FE73FAF1" box="[199,388,1309,1333]" italics="true" pageId="4" pageNumber="53">C. quinquelatera</emphasis>
<bibRefCitation id="902B3AE6FFE4FF8255BB5D72FDFEFAF1" author="Hiro" box="[396,521,1309,1333]" pageId="4" pageNumber="53" refString="Hiro, F. (1932) Report on the Japanese species of the genus Calantica (Cirripedia). Annotationes Zoologicae, Japon, 13 (5), 467 - 482." type="journal article" year="1932">Hiro, 1932</bibRefCitation>
: 469
</taxonomicName>
,
<collectingCountry id="8CAD0787FFE4FF82567C5D71FD79FAF1" box="[587,654,1310,1333]" name="Japan" pageId="4" pageNumber="53">Japan</collectingCountry>
,?shallow water.
</paragraph>
<paragraph id="F4054717FFE4FF8254A05D2EFABBFAB9" blockId="4.[151,1436,1309,1406]" pageId="4" pageNumber="53">
11)
<taxonomicName id="33BA3C94FFE4FF8254F05D2DFEC5FA9D" box="[199,306,1345,1369]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="villosa">
<emphasis id="C6CE9B05FFE4FF8254F05D2DFEC5FA9D" box="[199,306,1345,1369]" italics="true" pageId="4" pageNumber="53">C. villosa</emphasis>
</taxonomicName>
(
<bibRefCitation id="902B3AE6FFE4FF8255775D2EFD9BFA9D" author="Leach" box="[320,620,1345,1369]" pageId="4" pageNumber="53" refString="Leach, W. E. (1824) Cirripedes. Supplement to the 4 th, 5 t and 6 th editions of the Encyclopaedia Britannica, 3 (1), 168 - 171 + Pl. 57." type="journal article" year="1824">Leach, 1824: 170, Plate 57</bibRefCitation>
) (on p. 170 as
<taxonomicName id="33BA3C94FFE4FF8257275D2EFBF3FA9D" box="[784,1028,1345,1369]" class="Maxillopoda" family="Pollicipedidae" genus="Pollicipes" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="tomentosus">
<emphasis id="C6CE9B05FFE4FF8257275D2EFBF3FA9D" box="[784,1028,1345,1369]" italics="true" pageId="4" pageNumber="53">Pollicipes tomentosus</emphasis>
</taxonomicName>
according to Darwin 1852: 274, and on Plate 57 dated 1817, as
<taxonomicName id="33BA3C94FFE4FF8255C35D08FD93FABA" box="[500,612,1382,1406]" class="Maxillopoda" family="Pollicipedidae" genus="Pollicipes" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="villosus">
<emphasis id="C6CE9B05FFE4FF8255C35D08FD93FABA" box="[500,612,1382,1406]" italics="true" pageId="4" pageNumber="53">P. villosus</emphasis>
</taxonomicName>
);
<typeStatus id="2B01F9B5FFE4FF82564A5D09FD5AFABA" box="[637,685,1382,1406]" pageId="4" pageNumber="53">type</typeStatus>
species;
<bibRefCitation id="902B3AE6FFE4FF8257235D09FC6BFAB9" author="Foster" box="[788,924,1381,1405]" pageId="4" pageNumber="53" refString="Foster, B. A. (1979) The Marine Fauna of New Zealand; Barnacles (Cirripedia: Thoracica). New Zealand Oceanographic Institute Memoir, 69, 1 - 159 (1978)." type="journal article" year="1979">Foster 1979</bibRefCitation>
: 44,
<collectingCountry id="8CAD0787FFE4FF8257E65D09FB9DFAB9" box="[977,1130,1381,1405]" name="New Zealand" pageId="4" pageNumber="53">New Zealand</collectingCountry>
, intertidal to
<specimenCount id="E2BC8C9EFFE4FF8250C95D0AFABBFAB9" box="[1278,1356,1381,1405]" pageId="4" pageNumber="53" type="generic">
<quantity id="3342EAF2FFE4FF8250C95D0AFABFFAB9" box="[1278,1352,1381,1405]" metricMagnitude="2" metricUnit="m" metricValue="2.76" pageId="4" pageNumber="53" unit="m" value="276.0">276 m</quantity>
.
</specimenCount>
</paragraph>
<paragraph id="F4054717FFE4FF8254AE5DDEFE38FA28" blockId="4.[153,1436,1453,1516]" pageId="4" pageNumber="53">
IV. Species with the basic pattern + L2 appearing somewhat rostral of midline of L, one of which,
<taxonomicName id="33BA3C94FFE4FF8250CF5DDEFA8DFA0C" box="[1272,1402,1457,1480]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="siemensi">
<emphasis id="C6CE9B05FFE4FF8250CF5DDEFA8DFA0C" box="[1272,1402,1457,1480]" italics="true" pageId="4" pageNumber="53">C. siemensi</emphasis>
</taxonomicName>
, is closest to
<taxonomicName id="33BA3C94FFE4FF8255005DBAFE30FA28" box="[311,455,1492,1516]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="moskalevi">
<emphasis id="C6CE9B05FFE4FF8255005DBAFE30FA28" box="[311,455,1492,1516]" italics="true" pageId="4" pageNumber="53">C. moskalevi</emphasis>
</taxonomicName>
:
</paragraph>
<paragraph id="F4054717FFE4FF8254A05E73FA6BF9B9" blockId="4.[151,1436,1564,1696]" pageId="4" pageNumber="53">
12)
<taxonomicName id="33BA3C94FFE4FF8254F05E72FDACF9F0" authority="Annandale, 1909: 267" authorityName="Annandale" authorityPageNumber="267" authorityYear="1909" box="[199,603,1564,1588]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="kampeni">
<emphasis id="C6CE9B05FFE4FF8254F05E72FEB1F9F0" box="[199,326,1564,1588]" italics="true" pageId="4" pageNumber="53">C. kampeni</emphasis>
(
<bibRefCitation id="902B3AE6FFE4FF8255625E73FDA4F9F0" author="Annandale" box="[341,595,1564,1588]" pageId="4" pageNumber="53" refString="Annandale, N. (1909) Description of a barnacle of the genus Scalpellum from Malaysia. Records of the Indian Museum, 3 (3), 267 - 270." type="journal article" year="1909">Annandale, 1909: 267</bibRefCitation>
)
</taxonomicName>
; cf.
<bibRefCitation id="902B3AE6FFE4FF8256BD5E72FCFBF9F0" author="Jones" box="[650,780,1564,1588]" pageId="4" pageNumber="53" refString="Jones, D. S. (1998) New genus and species of Calanticidae (Cirripedia, Thoracica, Scalpellomorpha) from Australian waters. Zoosystema, 20 (2), 239 - 253." type="journal article" year="1998">Jones 1998</bibRefCitation>
: 247, Gulf of Aden to Pacific Ocean,
<quantity id="3342EAF2FFE4FF8250855E73FAE0F9F0" box="[1202,1303,1564,1588]" metricMagnitude="2" metricUnit="m" metricValue="1.155" metricValueMax="2.25" metricValueMin="0.06" pageId="4" pageNumber="53" unit="m" value="115.5" valueMax="225.0" valueMin="6.0">
6
<specimenCount id="E2BC8C9EFFE4FF8250F95E73FAE0F9F0" box="[1230,1303,1564,1588]" pageId="4" pageNumber="53" type="generic">225 m</specimenCount>
</quantity>
. 13)
<taxonomicName id="33BA3C94FFE4FF8254F05E2EFDB6F99C" authority="Hoek, 1907: 60" authorityName="Hoek" authorityPageNumber="60" authorityYear="1907" box="[199,577,1600,1624]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="pollicipoides">
<emphasis id="C6CE9B05FFE4FF8254F05E2EFE8FF99C" box="[199,376,1600,1624]" italics="true" pageId="4" pageNumber="53">C. pollicipoides</emphasis>
(
<bibRefCitation id="902B3AE6FFE4FF8255B05E2FFDCEF99C" author="Hoek" box="[391,569,1600,1624]" pageId="4" pageNumber="53" refString="Hoek, P. P. C. (1907) Cirripedia of the Siboga-Expedition. A. Cirripedia Pedunculata. Siboga-Expeditie, 31 A, 1 - 127 + pls. 1 - 10. Brill, Leyden." type="journal article" year="1907">Hoek, 1907: 60</bibRefCitation>
)
</taxonomicName>
, Celebes,
<quantity id="3342EAF2FFE4FF8256835E2FFD07F99C" box="[692,752,1600,1624]" metricMagnitude="1" metricUnit="m" metricValue="5.7" pageId="4" pageNumber="53" unit="m" value="57.0">57 m</quantity>
; cf.
<bibRefCitation id="902B3AE6FFE4FF8257285E2EFC57F99C" author="Jones" box="[799,928,1600,1624]" pageId="4" pageNumber="53" refString="Jones, D. S. (1998) New genus and species of Calanticidae (Cirripedia, Thoracica, Scalpellomorpha) from Australian waters. Zoosystema, 20 (2), 239 - 253." type="journal article" year="1998">Jones 1998</bibRefCitation>
: 247, S. Africa to
<collectingCountry id="8CAD0787FFE4FF82505B5E2FFB1AF99C" box="[1132,1261,1600,1624]" name="Philippines" pageId="4" pageNumber="53">Philippines</collectingCountry>
,
<quantity id="3342EAF2FFE4FF8250CF5E2FFA9BF99C" box="[1272,1388,1600,1624]" metricMagnitude="2" metricUnit="m" metricValue="1.2349999999999999" metricValueMax="1.9" metricValueMin="0.57" pageId="4" pageNumber="53" unit="m" value="123.5" valueMax="190.0" valueMin="57.0">
57
<specimenCount id="E2BC8C9EFFE4FF8251155E2FFA9BF99C" box="[1314,1388,1600,1624]" pageId="4" pageNumber="53" type="generic">190 m</specimenCount>
</quantity>
. 14)
<taxonomicName id="33BA3C94FFE4FF8254F05E0AFDC4F9B8" authority="Weltner, 1922: 97" authorityName="Weltner" authorityPageNumber="97" authorityYear="1922" box="[199,563,1636,1660]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="siemensi">
<emphasis id="C6CE9B05FFE4FF8254F05E0AFEBEF9B8" box="[199,329,1637,1660]" italics="true" pageId="4" pageNumber="53">C. siemensi</emphasis>
(
<bibRefCitation id="902B3AE6FFE4FF82556D5E0BFDDCF9B8" author="Weltner" box="[346,555,1636,1660]" pageId="4" pageNumber="53" refString="Weltner, W. (1922) Cirripedia der Deutschen Tiefsee-Expedition, Wissenschafteliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot; 1898 - 1899, 23 (2), 69 - 113 + pls. 2 - 4. Jena." type="journal article" year="1922">Weltner, 1922: 97</bibRefCitation>
)
</taxonomicName>
;
<bibRefCitation id="902B3AE6FFE4FF8256765E0BFC8DF9B8" author="Zevina" box="[577,890,1636,1660]" pageId="4" pageNumber="53" refString="Zevina, G. B. &amp; Litvinova, N. M. (1970) Supplement to the barnacle fauna (Cirripedia, Thoracica) of the Red Sea. In: Kovalefsky, A. O. (ed.), Marine Biology, (18), 172 - 181. Institute for Biology of the Southern Oceans, Academy of Science USSR." type="journal article" year="1970">Zevina and Litvinova 1970</bibRefCitation>
: 176, Red Sea (Gulf of Aden),
<quantity id="3342EAF2FFE4FF8250D55E0BFAD7F9B8" box="[1250,1312,1636,1660]" metricMagnitude="1" metricUnit="m" metricValue="4.8" pageId="4" pageNumber="53" unit="m" value="48.0">48 m</quantity>
; cf.
<bibRefCitation id="902B3AE6FFE4FF8251645E0AFA6BF9B9" author="Young" box="[1363,1436,1637,1661]" pageId="4" pageNumber="53" refString="Young, P. S. (2003) Redescription of the calanticids (Cirripedia, Scalpellomorpha) described by Wilhelm Weltner. Zoosystematics and Evolution, 79 (1), 181 - 201." type="journal article" year="2003">Young</bibRefCitation>
</paragraph>
<paragraph id="F4054717FFE4FF8254F05EE7FDF7F964" blockId="4.[151,1436,1564,1696]" box="[199,512,1672,1696]" pageId="4" pageNumber="53">
<bibRef id="2D1AD3E1FFE4FF8254F05EE7FDF7F964" box="[199,512,1672,1696]" pageId="4" pageNumber="53" type="book chapter">
<year id="80120644FFE4FF8254F05EE7FEF4F964" box="[199,259,1672,1696]" pageId="4" pageNumber="53">2003</year>
:
<pagination id="CD73B9E3FFE4FF82553A5EE7FEC0F964" box="[269,311,1672,1696]" pageId="4" pageNumber="53" type="pagination">189</pagination>
<title id="86BEAC21FFE4FF8255095EE7FE0BF964" box="[318,508,1672,1696]" pageId="4" pageNumber="53">for redescription</title>
.
</bibRef>
</paragraph>
<paragraph id="F4054717FFE4FF8254F05EBFFDF5F8BA" blockId="4.[151,1436,1744,1918]" pageId="4" pageNumber="53">
Species number 9 above,
<taxonomicName id="33BA3C94FFE4FF8255C05EBFFD16F92C" box="[503,737,1744,1768]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="moskalevi">
<emphasis id="C6CE9B05FFE4FF8255C05EBFFD16F92C" box="[503,737,1744,1768]" italics="true" pageId="4" pageNumber="53">Calantica moskalevi</emphasis>
</taxonomicName>
from Axial Seamount, is somewhat uncomfortably situated between the generalized and advanced members of the genus. This is due to the variable occurrence of the CL2 versus an otherwise relatively generalized,
<emphasis id="C6CE9B05FFE4FF8256525F71FD01F8F2" box="[613,758,1822,1846]" italics="true" pageId="4" pageNumber="53">Scillaelepas-</emphasis>
like facies. What may have been the primordial CL2s were seen in the juveniles described by
<bibRefCitation id="902B3AE6FFE4FF8256245F2DFC52F89E" author="Zevina" box="[531,933,1858,1882]" pageId="4" pageNumber="53" refString="Zevina, G. B. &amp; Galkin, S. V. (1989) New species of cirripeds (Cirripedia, Thoracica) from thermal waters. Zoologicheskii Zhurnal, 68 (3), 134 - 136." type="journal article" year="1989">
Zevina and Galkin (1989: cf.
<figureCitation id="6C815B92FFE4FF82576B5F2DFC6BF89E" box="[860,924,1858,1882]" captionStart="FIGURE 5" captionStartId="6.[151,250,1830,1853]" captionTargetBox="[295,1289,199,1795]" captionTargetId="figure@6.[284,1301,193,1810]" captionTargetPageId="6" captionText="FIGURE 5. Atetrapachylasma dijonesae gen. et sp. nov. in situ on pillow basalt from 2080 m of depth Vance B Seamount, with nozzle of suction sampler of ROV Tiburon largely to their right. A, the remains of several walls broken away by the nozzle can be seen to the right of “ 1 ” leaving membranous bases exposed. B, the barnacles seen between “ 2 ” and “ 3 ” and to the right of “ 4 ” in A have been collected. N. B.: Inspection of MPEGs taken by ROV Tiburon during collection of the barnacles seen removed in these frames detected no loss of material in the process. Therefore, it can be said with confidence that the shell fragments at the base of the basalt below the barnacles (cf. Fig. 4) represent natural postmortem accumulations." httpUri="https://zenodo.org/record/202506/files/figure.png" pageId="4" pageNumber="53">Fig. 5</figureCitation>
)
</bibRefCitation>
and again in two of the 10 mature specimens reported upon herein (Figs 8, 9).
</paragraph>
<caption id="A0C5179FFFE5FF8354A05C41FC40FADF" httpUri="https://zenodo.org/record/202505/files/figure.png" pageId="5" pageNumber="54" targetBox="[256,1323,193,1021]" targetPageId="5">
<paragraph id="F4054717FFE5FF8354A05C41FC40FADF" blockId="5.[151,1436,1070,1307]" pageId="5" pageNumber="54">
<emphasis id="C6CE9B05FFE5FF8354A05C41FEE4FB80" bold="true" box="[151,275,1070,1093]" pageId="5" pageNumber="54">FIGURE 4.</emphasis>
Numerous females of
<taxonomicName id="33BA3C94FFE5FF8355CB5C40FCEAFB81" box="[508,797,1071,1093]" class="Maxillopoda" family="Pachylasmatidae" genus="Atetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="dijonesae">
<emphasis id="C6CE9B05FFE5FF8355CB5C40FCEAFB81" box="[508,797,1071,1093]" italics="true" pageId="5" pageNumber="54">Atetrapachylasma dijonesae</emphasis>
</taxonomicName>
<emphasis id="C6CE9B05FFE5FF8357135C40FC34FB80" bold="true" box="[804,963,1071,1092]" pageId="5" pageNumber="54">gen. et sp. nov.</emphasis>
<emphasis id="C6CE9B05FFE5FF8357FC5C5FFBFDFB81" box="[971,1034,1072,1093]" italics="true" pageId="5" pageNumber="54">in situ</emphasis>
on pillow basalt from 2080 m of depth on Vance B Seamount, with nozzle of suction sampler on ROV
<emphasis id="C6CE9B05FFE5FF8357135C22FC85FBA7" box="[804,882,1101,1123]" italics="true" pageId="5" pageNumber="54">Tiburon</emphasis>
to their right (distance between red laser dots 30 cm). The pristine white appearance of the dead shells accumulating at the base of the slope, as well as of the living individuals, suggests that there is little if any hydrothermal activity nearby to stain them (rather than the barnacle just being short lived, which they are). Thus, as might be expected, dissection revealed feeding appendages (Fig. 13) of the deep-sea rather than hydrothermal type. N.B.: Inspection of MPEGs taken by ROV
<emphasis id="C6CE9B05FFE5FF8356995CA7FD0BFB1A" box="[686,764,1224,1246]" italics="true" pageId="5" pageNumber="54">Tiburon</emphasis>
during collection of most of the barnacles seen in this frame (also, Fig. 5) detected no loss of material in the process. Therefore, it can be said with confidence that the shell fragments at the base of the basalt below the barnacles represent natural, postmortem accumulations.
</paragraph>
</caption>
<paragraph id="F4054717FFE5FF8354F25D24FE35F96D" blockId="5.[151,1436,1352,1886]" pageId="5" pageNumber="54">
The facts, that 1) the mature specimen with but one CL2 as well as that with two came from the same clump as the otherwise rather similar appearing specimens and 2) there are no other species of
<taxonomicName id="33BA3C94FFE5FF8350795D00FB4BFA43" box="[1102,1212,1391,1415]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="5" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFE5FF8350795D00FB4BFA43" box="[1102,1212,1391,1415]" italics="true" pageId="5" pageNumber="54">Calantica</emphasis>
</taxonomicName>
known in the entire Central as well as Eastern Pacific, leaves little doubt the difference is due to variability rather than to a mistaken identification. While we are unaware of such variability having been previously reported in
<taxonomicName id="33BA3C94FFE5FF8350AF5DD8FAF1FA0B" box="[1176,1286,1463,1487]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="5" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFE5FF8350AF5DD8FAF1FA0B" box="[1176,1286,1463,1487]" italics="true" pageId="5" pageNumber="54">Calantica</emphasis>
</taxonomicName>
, more or less comparable variations involving 1) the SRs were noted in
<taxonomicName id="33BA3C94FFE5FF8357745DB4FB00FA37" authority="Gruvel, 1911" authorityName="Gruvel" authorityYear="1911" box="[835,1271,1499,1523]" class="Maxillopoda" family="Calanticidae" genus="Gruvelialepas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="pilsbryi">
<emphasis id="C6CE9B05FFE5FF8357745DB4FBB5FA37" box="[835,1090,1499,1523]" italics="true" pageId="5" pageNumber="54">Gruvelialepas pilsbryi</emphasis>
(
<bibRefCitation id="902B3AE6FFE5FF8350635DB4FB19FA37" author="Gruvel" box="[1108,1262,1499,1523]" pageId="5" pageNumber="54" refString="Gruvel, A. (1911) Sur deux especes nouvelles de Cirrhipedes appartenant a la collection du Museum. Bulletin Museum d'Histoire nataturelle, Paris 17, 290 - 292." type="journal article" year="1911">Gruvel, 1911</bibRefCitation>
)
</taxonomicName>
, (cf.
<bibRefCitation id="902B3AE6FFE5FF8351005DB3FF24F9D3" author="Newman" pageId="5" pageNumber="54" refString="Newman, W. A. (1980) A review of extant Scillaelepas (Cirripedia: Scalpellidae) including recognition of new species from the North Atlantic, Western Indian Ocean and New Zealand. Tethys, 9 (4), 379 - 398." type="journal article" year="1980">Newman 1980</bibRefCitation>
), 2) an additional CL and SC in
<taxonomicName id="33BA3C94FFE5FF8356725D90FABFF9D3" authority="Zevina and Yakhontova, 1987 (Newman, 1993: 405)" authorityName="Zevina and Yakhontova, 1987 (Newman" authorityPageNumber="405" authorityYear="1993" box="[581,1352,1535,1559]" class="Maxillopoda" family="Calanticidae" genus="Newmanilepas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pedunculata" pageId="5" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFE5FF8356725D90FD1DF9D3" box="[581,746,1535,1559]" italics="true" pageId="5" pageNumber="54">Newmanilepas</emphasis>
<bibRefCitation id="902B3AE6FFE5FF8356C55D90FBB2F9D3" author="Zevina" box="[754,1093,1535,1559]" pageId="5" pageNumber="54" refString="Zevina, G. B. &amp; Yakhontova, I. V. (1987) A new barnacle genus of the family Scalpellidae (Crustacea, Cirripedia) from the North Atlantic. Zoologicheskii Zhurnal, 66 (8), 1261 - 1264." type="journal article" year="1987">Zevina and Yakhontova, 1987</bibRefCitation>
(
<bibRefCitation id="902B3AE6FFE5FF8350635D90FAB7F9D3" author="Newman" box="[1108,1344,1535,1559]" pageId="5" pageNumber="54" refString="Newman, W. A. (1993) Darwin and cirripedology. In: Truesdale, J. (ed.), The History of Carcinology. Crustacean Issues, 8, 349 - 434. Balkema, Rotterdam." type="journal article" year="1993">Newman, 1993: 405</bibRefCitation>
)
</taxonomicName>
, and 3) the number of L in
<taxonomicName id="33BA3C94FFE5FF8355445E4CFBC6F9FF" authority="(Quoy and Gaimard, 1834) (Foster 1979: 40)" authorityName="(Quoy and Gaimard, 1834) (Foster" authorityPageNumber="40" authorityYear="1979" box="[371,1073,1571,1595]" class="Maxillopoda" family="Scalpellidae" genus="Smilium" kingdom="Animalia" order="Pedunculata" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="spinosa">
<emphasis id="C6CE9B05FFE5FF8355445E4CFDDAF9FF" box="[371,557,1571,1595]" italics="true" pageId="5" pageNumber="54">Smilium spinosa</emphasis>
(
<bibRefCitation id="902B3AE6FFE5FF83560B5E4CFCAAF9FF" author="Quoy" box="[572,861,1571,1595]" pageId="5" pageNumber="54" refString="Quoy, J. R. C. &amp; Gaimard, J. P. (1834) Voyage de decouvertes de l'Astrolabe, execut par ordre du Roi, pendant les annees 1826 - 29, sous le commandement de M. J. Dumond d'Urville. Zoologie, Mollusca, 3, Paris. 712 pp." type="book" year="1834">Quoy and Gaimard, 1834</bibRefCitation>
) (
<bibRefCitation id="902B3AE6FFE5FF8357425E4CFBDEF9FF" author="Foster" box="[885,1065,1571,1595]" pageId="5" pageNumber="54" refString="Foster, B. A. (1979) The Marine Fauna of New Zealand; Barnacles (Cirripedia: Thoracica). New Zealand Oceanographic Institute Memoir, 69, 1 - 159 (1978)." type="journal article" year="1979">Foster 1979: 40</bibRefCitation>
)
</taxonomicName>
that have been observed. While the examples of the delayed addition of plates suggest that the CL2s of
<taxonomicName id="33BA3C94FFE5FF8357825E25FBB1F9A6" box="[949,1094,1610,1634]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="moskalevi">
<emphasis id="C6CE9B05FFE5FF8357825E25FC3FF9A5" box="[949,968,1610,1633]" italics="true" pageId="5" pageNumber="54">C</emphasis>
.
<emphasis id="C6CE9B05FFE5FF8357E15E25FBB1F9A6" box="[982,1094,1610,1634]" italics="true" pageId="5" pageNumber="54">moskalevi</emphasis>
</taxonomicName>
might possibly appear later in ontogeny, the fact that some of the specimens without them appear as old if not older than the specimen having them renders that unlikely.
</paragraph>
<paragraph id="F4054717FFE5FF8354F25ED9FBE4F89A" blockId="5.[151,1436,1352,1886]" pageId="5" pageNumber="54">
Zevina and Galkin (1998) assigned the specimens to
<taxonomicName id="33BA3C94FFE5FF8357295ED9FC7AF90A" box="[798,909,1718,1742]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="5" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFE5FF8357295ED9FC7AF90A" box="[798,909,1718,1742]" italics="true" pageId="5" pageNumber="54">Calantica</emphasis>
</taxonomicName>
with a parenthetical question mark of caution, “
<taxonomicName id="33BA3C94FFE5FF8354945EB5FEE6F936" box="[163,273,1754,1778]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="5" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFE5FF8354945EB5FEE6F936" box="[163,273,1754,1778]" italics="true" pageId="5" pageNumber="54">Calantica</emphasis>
</taxonomicName>
(?)
<emphasis id="C6CE9B05FFE5FF8355765EB5FE37F936" box="[321,448,1754,1778]" italics="true" pageId="5" pageNumber="54">
<taxonomicName id="33BA3C94FFE5FF8355765EB5FE4EF936" box="[321,441,1754,1778]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="moskalevi">moskalevi</taxonomicName>
</emphasis>
, and they reported it as within
<quantity id="3342EAF2FFE5FF8357135EB6FC68F935" box="[804,927,1753,1777]" metricMagnitude="1" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="5" pageNumber="54" unit="m" value="20.0" valueMax="30.0" valueMin="10.0">10 to 30 m</quantity>
of vents in the caldera of Axial Seamount at
<geoCoordinate id="918E21D0FFE5FF8354A05E91FEE8F8D2" box="[151,287,1790,1814]" direction="north" orientation="latitude" pageId="5" pageNumber="54" precision="92" value="45.928333">45° 55.7 N</geoCoordinate>
,
<geoCoordinate id="918E21D0FFE5FF83551C5E91FE32F8D2" box="[299,453,1790,1814]" direction="west" orientation="longitude" pageId="5" pageNumber="54" precision="92" value="-130.03833">130° 02.3 W</geoCoordinate>
and
<geoCoordinate id="918E21D0FFE5FF8355CA5E91FD72F8D2" box="[509,645,1790,1814]" direction="north" orientation="latitude" pageId="5" pageNumber="54" precision="92" value="45.925">45° 55.5 N</geoCoordinate>
,
<geoCoordinate id="918E21D0FFE5FF8356A55E91FCD4F8D2" box="[658,803,1790,1814]" direction="west" orientation="longitude" pageId="5" pageNumber="54" precision="92" value="-130.02667">130° 01.6W</geoCoordinate>
at depths of
<quantity id="3342EAF2FFE5FF83578F5E91FBF9F8D2" box="[952,1038,1790,1814]" metricMagnitude="3" metricUnit="m" metricValue="1.41" pageId="5" pageNumber="54" unit="m" value="1410.0">1410 m</quantity>
and
<quantity id="3342EAF2FFE5FF8350725E91FB6AF8D2" box="[1093,1181,1790,1814]" metricMagnitude="3" metricUnit="m" metricValue="1.54" pageId="5" pageNumber="54" unit="m" value="1540.0">1540 m</quantity>
, respectively. The two juveniles, upon which the description was based, were designated the
<typeStatus id="2B01F9B5FFE5FF8357945F4EFBF2F8FD" box="[931,1029,1825,1849]" pageId="5" pageNumber="54" type="holotype">holotype</typeStatus>
and
<typeStatus id="2B01F9B5FFE5FF83500C5F4DFB69F8FE" box="[1083,1182,1826,1850]" pageId="5" pageNumber="54" type="paratype">paratype</typeStatus>
, and measured 4.3 and 2.8 mm in height, respectively (
<figureCitation id="6C815B92FFE5FF8355CD5F29FDC8F89A" box="[506,575,1862,1886]" captionStart="FIGURE 7" captionStartId="7.[151,250,1728,1751]" captionTargetBox="[196,1368,929,1688]" captionTargetId="figure@7.[188,1393,910,1714]" captionTargetPageId="7" captionText="FIGURE 7. The holotype and paratype of Calantica moskalevi Zevina and Galkin, 1989: 134, Fig. 1) consisted of two juveniles (measurements below in mm)." httpUri="https://zenodo.org/record/202507/files/figure.png" pageId="5" pageNumber="54">Fig. 7</figureCitation>
). A description of the adult form follows.
</paragraph>
<caption id="A0C5179FFFE6FF8054A05F49FB20F811" httpUri="https://zenodo.org/record/202506/files/figure.png" pageId="6" pageNumber="55" targetBox="[295,1289,199,1795]" targetPageId="6">
<paragraph id="F4054717FFE6FF8054A05F49FB20F811" blockId="6.[151,1436,1830,2006]" pageId="6" pageNumber="55">
<emphasis id="C6CE9B05FFE6FF8054A05F49FEE3F8F8" bold="true" box="[151,276,1830,1853]" pageId="6" pageNumber="55">FIGURE 5.</emphasis>
<taxonomicName id="33BA3C94FFE6FF80552B5F49FDCAF8F8" box="[284,573,1830,1852]" class="Maxillopoda" family="Pachylasmatidae" genus="Atetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="6" pageNumber="55" phylum="Arthropoda" rank="species" species="dijonesae">
<emphasis id="C6CE9B05FFE6FF80552B5F49FDCAF8F8" box="[284,573,1830,1852]" italics="true" pageId="6" pageNumber="55">Atetrapachylasma dijonesae</emphasis>
</taxonomicName>
<emphasis id="C6CE9B05FFE6FF8056725F48FD10F8F8" bold="true" box="[581,743,1831,1852]" pageId="6" pageNumber="55">gen. et sp. nov.</emphasis>
<emphasis id="C6CE9B05FFE6FF8056C75F48FCC7F8F8" box="[752,816,1831,1852]" italics="true" pageId="6" pageNumber="55">in situ</emphasis>
on pillow basalt from 2080 m of depth Vance B Seamount, with nozzle of suction sampler of ROV
<emphasis id="C6CE9B05FFE6FF8056105F2AFD82F89F" box="[551,629,1861,1883]" italics="true" pageId="6" pageNumber="55">Tiburon</emphasis>
largely to their right. A, the remains of several walls broken away by the nozzle can be seen to the right of “1” leaving membranous bases exposed. B, the barnacles seen between “2” and “3” and to the right of “4” in A have been collected. N.B.: Inspection of MPEGs taken by ROV
<emphasis id="C6CE9B05FFE6FF8057825FEDFBF4F85C" box="[949,1027,1922,1944]" italics="true" pageId="6" pageNumber="55">Tiburon</emphasis>
during collection of the barnacles seen removed in these frames detected no loss of material in the process. Therefore, it can be said with confidence that the shell fragments at the base of the basalt below the barnacles (cf. Fig. 4) represent natural postmortem accumulations.
</paragraph>
</caption>
<paragraph id="F4054717FFE7FF8154A05AB1FBDEFCB6" blockId="7.[151,1437,734,882]" pageId="7" pageNumber="56">
<emphasis id="C6CE9B05FFE7FF8154A05AB1FEE5FD30" bold="true" box="[151,274,734,757]" pageId="7" pageNumber="56">FIGURE 6.</emphasis>
The capitular plates of
<taxonomicName id="33BA3C94FFE7FF8155C85AB0FD94FD31" box="[511,611,735,757]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="7" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFE7FF8155C85AB0FD94FD31" box="[511,611,735,757]" italics="true" pageId="7" pageNumber="56">Calantica</emphasis>
</taxonomicName>
can be divided into three more or less arbitrary whorls; 1, S-T; 2, R-C; and 3, RL- L-CL-SC (scutum, tergum, rostrum, carina, rostrolatus, latus, carinolatus and subcarina, respectively), sometimes supplemented by a subrostrum, or second latus or second carinolatus, (SR or L2 or CL2), respectively. The same three whorls are typical of
<taxonomicName id="33BA3C94FFE7FF8154E15B52FEA7FC97" box="[214,336,829,851]" class="Maxillopoda" family="Calanticidae" genus="Scillaelepas" kingdom="Animalia" order="Pedunculata" pageId="7" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFE7FF8154E15B52FEA7FC97" box="[214,336,829,851]" italics="true" pageId="7" pageNumber="56">Scillaelepas</emphasis>
</taxonomicName>
, which differs in not having supplementary capitular plates other than sometime a subrostrum or two, and in having proportionately larger peduncular plates not covered with a relatively thick cuticle.
</paragraph>
<caption id="A0C5179FFFE7FF8154A05EAFFDF5F931" httpUri="https://zenodo.org/record/202507/files/figure.png" pageId="7" pageNumber="56" targetBox="[196,1368,929,1688]" targetPageId="7">
<paragraph id="F4054717FFE7FF8154A05EAFFDF5F931" blockId="7.[151,1436,1728,1781]" pageId="7" pageNumber="56">
<emphasis id="C6CE9B05FFE7FF8154A05EAFFEE3F912" bold="true" box="[151,276,1728,1751]" pageId="7" pageNumber="56">FIGURE 7.</emphasis>
The holotype and paratype of
<taxonomicName id="33BA3C94FFE7FF8156655EAFFB97F912" authority="Zevina and Galkin, 1989: 134" authorityName="Zevina and Galkin" authorityPageNumber="134" authorityYear="1989" box="[594,1120,1728,1750]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="moskalevi">
<emphasis id="C6CE9B05FFE7FF8156655EAFFCD5F912" box="[594,802,1728,1750]" italics="true" pageId="7" pageNumber="56">Calantica moskalevi</emphasis>
Zevina and Galkin, 1989: 134
</taxonomicName>
, Fig. 1) consisted of two juveniles (measurements below in mm).
</paragraph>
</caption>
<paragraph id="F4054717FFE7FF8155AE5F5AFB5DF85A" pageId="7" pageNumber="56">
<table id="86BAB5B7FFE7007954A85F5AFABDF85A" box="[159,1354,1845,1950]" gridcols="5" gridrows="3" pageId="7" pageNumber="56">
<tr id="4A8A4555FFE7007954A85F5AFABDF88F" box="[159,1354,1845,1867]" gridrow="0" pageId="7" pageNumber="56" rowspan-0="1">
<th id="095B2C29FFE7007955AE5F5AFDCBF88F" box="[409,572,1845,1867]" gridcol="1" gridrow="0" pageId="7" pageNumber="56">Capitular height</th>
<th id="095B2C29FFE7007956A55F5AFCC7F88F" box="[658,816,1845,1867]" gridcol="2" gridrow="0" pageId="7" pageNumber="56">Capitular width</th>
<th id="095B2C29FFE7007957BB5F5AFBB4F88F" box="[908,1091,1845,1867]" gridcol="3" gridrow="0" pageId="7" pageNumber="56">Peduncular height</th>
<th id="095B2C29FFE7007950AE5F5AFABDF88F" box="[1177,1354,1845,1867]" gridcol="4" gridrow="0" pageId="7" pageNumber="56">Peduncular width</th>
</tr>
<tr id="4A8A4555FFE7007954A85F30FABDF8B1" box="[159,1354,1887,1909]" gridrow="1" pageId="7" pageNumber="56">
<th id="095B2C29FFE7007954A85F30FF0AF8B1" box="[159,253,1887,1909]" gridcol="0" gridrow="1" pageId="7" pageNumber="56">Holotype</th>
<td id="095B2C29FFE7007955AE5F30FDCBF8B1" box="[409,572,1887,1909]" gridcol="1" gridrow="1" pageId="7" pageNumber="56">2.8</td>
<td id="095B2C29FFE7007956A55F30FCC7F8B1" box="[658,816,1887,1909]" gridcol="2" gridrow="1" pageId="7" pageNumber="56">3.0</td>
<td id="095B2C29FFE7007957BB5F30FBB4F8B1" box="[908,1091,1887,1909]" gridcol="3" gridrow="1" pageId="7" pageNumber="56">1.5</td>
<td id="095B2C29FFE7007950AE5F30FABDF8B1" box="[1177,1354,1887,1909]" gridcol="4" gridrow="1" pageId="7" pageNumber="56">1.0</td>
</tr>
<tr id="4A8A4555FFE7007954A85FE6FABDF85A" box="[159,1354,1929,1950]" gridrow="2" pageId="7" pageNumber="56">
<th id="095B2C29FFE7007954A85FE6FF0AF85A" box="[159,253,1929,1950]" gridcol="0" gridrow="2" pageId="7" pageNumber="56">Paratype</th>
<td id="095B2C29FFE7007955AE5FE6FDCBF85A" box="[409,572,1929,1950]" gridcol="1" gridrow="2" pageId="7" pageNumber="56">1.8</td>
<td id="095B2C29FFE7007956A55FE6FCC7F85A" box="[658,816,1929,1950]" gridcol="2" gridrow="2" pageId="7" pageNumber="56">1.0</td>
<td id="095B2C29FFE7007957BB5FE6FBB4F85A" box="[908,1091,1929,1950]" gridcol="3" gridrow="2" pageId="7" pageNumber="56">--</td>
<td id="095B2C29FFE7007950AE5FE6FABDF85A" box="[1177,1354,1929,1950]" gridcol="4" gridrow="2" pageId="7" pageNumber="56">--</td>
</tr>
</table>
</paragraph>
<paragraph id="F4054717FFE7FF8154A05FD0FACEF837" blockId="7.[151,1436,1983,2036]" pageId="7" pageNumber="56">
The authors note “One scale on each side …” (more or less at the capitulo-pedunclar junction). These can be seen in the
<typeStatus id="2B01F9B5FFE7FF8151515FD0FF34F837" pageId="7" pageNumber="56" type="holotype">holotype</typeStatus>
(A and B) and on one side of the
<typeStatus id="2B01F9B5FFE7FF81562E5FB1FD87F837" box="[537,624,2014,2035]" pageId="7" pageNumber="56" type="paratype">paratype</typeStatus>
(F). See discussion herein re the apparent significance of these scales.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="1C59A89BAB69B9BDBB7CA3D0E1A67A2B">Two Northeast Pacific deep-water barnacle populations (Cirripedia: Calanticidae and Pachylasmatidae) from seamounts of the Juan de Fuca Ridge; &quot; insular &quot; endemics stemming from Tethys, or by subsequent dispersal from the Western Pacific center of distribution?</mods:title>
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<subSubSection id="BCA0149CFFE8FF8E54A058F8FE95FF16" pageId="8" pageNumber="57" type="nomenclature">
<paragraph id="F4054717FFE8FF8E54A058F8FD30FF75" blockId="8.[151,711,151,210]" box="[151,711,151,177]" pageId="8" pageNumber="57">
<heading id="AF4DF07BFFE8FF8E54A058F8FD30FF75" bold="true" box="[151,711,151,177]" fontSize="11" level="1" pageId="8" pageNumber="57" reason="1">
<taxonomicName id="33BA3C94FFE8FF8E54A058F8FD30FF75" authority="Zevina and Galkin, 1989" authorityName="Zevina and Galkin" authorityYear="1989" box="[151,711,151,177]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="8" pageNumber="57" phylum="Arthropoda" rank="species" species="moskalevi">
<emphasis id="C6CE9B05FFE8FF8E54A058F8FD30FF75" bold="true" box="[151,711,151,177]" pageId="8" pageNumber="57">
<emphasis id="C6CE9B05FFE8FF8E54A058F8FE7BFF75" bold="true" box="[151,396,151,177]" italics="true" pageId="8" pageNumber="57">Calantica moskalevi</emphasis>
<bibRefCitation id="902B3AE6FFE8FF8E55A458F8FD30FF75" author="Zevina" box="[403,711,151,177]" pageId="8" pageNumber="57" refString="Zevina, G. B. &amp; Galkin, S. V. (1989) New species of cirripeds (Cirripedia, Thoracica) from thermal waters. Zoologicheskii Zhurnal, 68 (3), 134 - 136." type="journal article" year="1989">Zevina and Galkin, 1989</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="F4054717FFE8FF8E54A058D5FE95FF16" blockId="8.[151,711,151,210]" box="[151,354,186,210]" pageId="8" pageNumber="57">
<figureCitation id="6C815B92FFE8FF8E54A058D5FEF5FF16" box="[151,258,186,210]" captionStart="FIGURE 2" captionStartId="2.[151,250,915,938]" captionTargetBox="[313,1273,363,894]" captionTargetId="figure@2.[313,1273,363,894]" captionTargetPageId="2" captionText="FIGURE 2. Panorama in the caldera of Axial Seamount, at approximately 1434 m of depth. Several yellowish-brown clumps and a few scatters individuals of the stalked barnacle, Calantica moskalevi Zevina and Galkin, 1989, can be seen at roughly 9, 12 and 3 oclock of the right red laser dot (distance between red laser dots 30 cm)." httpUri="https://zenodo.org/record/202503/files/figure.png" pageId="8" pageNumber="57">Figures 2</figureCitation>
,
<figureCitation id="6C815B92FFE8FF8E553958D5FEE9FF16" box="[270,286,186,210]" captionStart="FIGURE 3" captionStartId="3.[151,250,1525,1548]" captionTargetBox="[313,1273,339,1502]" captionTargetId="figure@3.[313,1273,339,1502]" captionTargetPageId="3" captionText="FIGURE 3. Group of several adult hermaphrodites of Calantica moskalevi Zevina and Galkin, 1989 in situ in the caldera of Axial Seamount (Fig. 1). The upper barnacle has its cirral net deployed for passive feeding in gentle currents. The cuticle of the barnacles, stained yellow-brown to gold apparently by ferromanganese oxides from venting 200 meters of so away, has sloughed off the apexes of some of the capitular plates, exposing the underlying calcitic shell to corrosion. Dissection reveals feeding appendages (Fig. 8) of the deep-sea rather than hydrothermal type." httpUri="https://zenodo.org/record/202504/files/figure.png" pageId="8" pageNumber="57">3</figureCitation>
, 810
</paragraph>
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<subSubSection id="BCA0149CFFE8FF8E54A0596DFCABFEDC" box="[151,860,258,280]" pageId="8" pageNumber="57" type="reference_group">
<paragraph id="F4054717FFE8FF8E54A0596DFCABFEDC" blockId="8.[151,860,258,280]" box="[151,860,258,280]" pageId="8" pageNumber="57">
<treatmentCitationGroup id="D4AA6039FFE8FF8E54A0596DFCABFEDC" box="[151,860,258,280]" pageId="8" pageNumber="57">
<taxonomicName id="33BA3C94FFE8FF8E54A0596DFD6AFEDC" authority="Zevina and Galkin, 1989: 134" authorityName="Zevina and Galkin" authorityPageNumber="134" authorityYear="1989" box="[151,669,258,280]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="8" pageNumber="57" phylum="Arthropoda" rank="species" species="moskalevi">
<emphasis id="C6CE9B05FFE8FF8E54A0596DFE91FEDC" box="[151,358,258,280]" italics="true" pageId="8" pageNumber="57">Calantica moskalevi</emphasis>
<treatmentCitation id="751B6106FFE8FF8E555B596DFD6AFEDC" author="Zevina" box="[364,669,258,280]" page="134" pageId="8" pageNumber="57" year="1989">
<bibRefCitation id="902B3AE6FFE8FF8E555B596DFD9BFEDC" author="Zevina" box="[364,620,258,280]" pageId="8" pageNumber="57" refString="Zevina, G. B. &amp; Galkin, S. V. (1989) New species of cirripeds (Cirripedia, Thoracica) from thermal waters. Zoologicheskii Zhurnal, 68 (3), 134 - 136." type="journal article" year="1989">Zevina and Galkin, 1989</bibRefCitation>
: 134
</treatmentCitation>
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,
<figureCitation id="6C815B92FFE8FF8E569F596DFD11FEDC" box="[680,742,258,280]" captionStart="FIGURE 1" captionStartId="1.[151,250,1711,1734]" captionTargetBox="[315,1272,639,1699]" captionTargetId="figure@1.[313,1273,636,1700]" captionTargetPageId="1" captionText="FIGURE 1. Axial and Vance B Seamounts are located on the Juan de Fuca Ridge, at 45 º 55 ' N 130 º 00 ' W and 45 ° 30 ' N 130 ° 40 ' W off Oregon (insert, large and small dot) respectively. Hydrothermally active Axial Seamount resides on the ridge crest near the point of origin of the chain of seamounts ranging to the NW. Vance B, among the seamounts running nearly parallel off the ridge crest some 96 km to the SSW, is hydrothermally inactive (see Table 1 for some physical parameters)." httpUri="https://zenodo.org/record/202502/files/figure.png" pageId="8" pageNumber="57">Figs 1</figureCitation>
ah, 2 ae.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BCA0149CFFE8FF8E54A0592AFF42FDD5" pageId="8" pageNumber="57" type="materials_examined">
<paragraph id="F4054717FFE8FF8E54A0592AFAFAFE61" blockId="8.[151,1436,325,751]" pageId="8" pageNumber="57">
<emphasis id="C6CE9B05FFE8FF8E54A0592AFE77FE9A" bold="true" box="[151,384,325,350]" pageId="8" pageNumber="57">Material examined.</emphasis>
<collectionCode id="92ABDFD2FFE8FF8E55B15929FE15FE9A" box="[390,482,326,350]" pageId="8" pageNumber="57">MBARI</collectionCode>
ROV
<emphasis id="C6CE9B05FFE8FF8E561D592AFD77FE99" box="[554,640,325,349]" italics="true" pageId="8" pageNumber="57">Tiburon</emphasis>
sample nos T1009 A2 and R3, (a clump of ten mature and one juvenile specimen, respectively) and
<collectionCode id="92ABDFD2FFE8FF8E55E85906FDCCFE45" box="[479,571,361,385]" pageId="8" pageNumber="57">MBARI</collectionCode>
ROV
<emphasis id="C6CE9B05FFE8FF8E56BD5906FD15FE45" box="[650,738,361,385]" italics="true" pageId="8" pageNumber="57">Tiburon</emphasis>
sample no. T1010 R7, fragments of
<specimenCount id="E2BC8C9EFFE8FF8E50A45906FAEFFE45" box="[1171,1304,361,385]" pageId="8" pageNumber="57" type="generic">1 specimen</specimenCount>
; Axial seamount (
<geoCoordinate id="918E21D0FFE8FF8E54DA59E2FEAEFE61" box="[237,345,397,421]" direction="north" orientation="latitude" pageId="8" pageNumber="57" precision="925" value="45.933334">45º 56' N</geoCoordinate>
,
<geoCoordinate id="918E21D0FFE8FF8E555359E2FE13FE61" box="[356,484,397,421]" direction="west" orientation="longitude" pageId="8" pageNumber="57" precision="925" value="-130.03334">130º 02' W</geoCoordinate>
), SW and NE caldera wall. Depths of
<quantity id="3342EAF2FFE8FF8E57B859E2FC13FE61" box="[911,996,397,421]" metricMagnitude="3" metricUnit="m" metricValue="1.4769999999999999" pageId="8" pageNumber="57" unit="m" value="1477.0">1477 m</quantity>
and
<quantity id="3342EAF2FFE8FF8E502D59E2FB86FE61" box="[1050,1137,397,421]" metricMagnitude="3" metricUnit="m" metricValue="1.434" pageId="8" pageNumber="57" unit="m" value="1434.0">1434 m</quantity>
, respectively:
</paragraph>
<paragraph id="F4054717FFE8FF8E54F059DEFF42FDD5" blockId="8.[151,1436,325,751]" pageId="8" pageNumber="57">
Deposition of topotypes from sample no. T1009 A2: First topotype,
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cat. no. 1145900; second topotype,
<collectionCode id="92ABDFD2FFE8FF8E54A059BAFF1FFE29" LSID="urn:lsid:biocol.org:col:34871" box="[151,232,469,493]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="57">USNM</collectionCode>
cat. no. 1145901; third topotype,
<collectionCode id="92ABDFD2FFE8FF8E566859BAFD7BFE29" LSID="urn:lsid:biocol.org:col:14844" box="[607,652,469,493]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14844" name="Scripps Institution of Oceanography" pageId="8" pageNumber="57">SIO</collectionCode>
<collectionCode id="92ABDFD2FFE8FF8E56A459BAFD36FE29" box="[659,705,469,493]" pageId="8" pageNumber="57">BIC</collectionCode>
cat. no. C-11193; topotype lot,
<collectionCode id="92ABDFD2FFE8FF8E501659BAFBB9FE29" LSID="urn:lsid:biocol.org:col:14844" box="[1057,1102,469,493]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14844" name="Scripps Institution of Oceanography" pageId="8" pageNumber="57">SIO</collectionCode>
<collectionCode id="92ABDFD2FFE8FF8E506359BAFB74FE29" box="[1108,1155,469,493]" pageId="8" pageNumber="57">BIC</collectionCode>
cat. no. C-11194 (Figs 8, 9).
</paragraph>
</subSubSection>
<subSubSection id="BCA0149CFFE8FF8E54F05A70FDC4FD98" pageId="8" pageNumber="57" type="diagnosis">
<paragraph id="F4054717FFE8FF8E54F05A70FDC4FD98" blockId="8.[151,1436,325,751]" pageId="8" pageNumber="57">
<emphasis id="C6CE9B05FFE8FF8E54F05A70FEB6FDFC" bold="true" box="[199,321,543,568]" pageId="8" pageNumber="57">Diagnosis.</emphasis>
With basic 13 capitular plates but unique in occasionally having second carinolatus (CL2) on one or both sides, without subrostrum (SR).
</paragraph>
</subSubSection>
<subSubSection id="BCA0149CFFE8FF8F54F05A05FEEBF9AB" lastPageId="9" lastPageNumber="58" pageId="8" pageNumber="57" type="description">
<paragraph id="F4054717FFE8FF8E54F05A05FC69FD2B" blockId="8.[151,1436,325,751]" pageId="8" pageNumber="57">
<emphasis id="C6CE9B05FFE8FF8E54F05A05FEA6FD47" bold="true" box="[199,337,618,643]" pageId="8" pageNumber="57">Description</emphasis>
. Hermaphrodites with capitular armament of S-T surrounded by R-RL-L-CL-±CL2-C-SC, capitular height to
<quantity id="3342EAF2FFE8FF8E551D5AE0FE89FD63" box="[298,382,655,679]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="57" unit="mm" value="30.0">30 mm</quantity>
; peduncular length usually from about same as, to as much as twice, capitular height. Plates white, covered with transparent cuticle, cuticle becoming stained yellowish gold, eventually even becoming thinly encrusted with what appears to be ferromanganese oxides (Figs 8, 9).
</paragraph>
<paragraph id="F4054717FFE8FF8E54A05D7FFE7AFA43" blockId="8.[151,1436,1296,1415]" pageId="8" pageNumber="57">
<emphasis id="C6CE9B05FFE8FF8E54A05D7FFEE4FAE2" bold="true" box="[151,275,1296,1319]" pageId="8" pageNumber="57">FIGURE 8.</emphasis>
<taxonomicName id="33BA3C94FFE8FF8E552E5D7FFD18FAE2" authority="Zevina and Galkin, 1989" authorityName="Zevina and Galkin" authorityYear="1989" box="[281,751,1296,1318]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="8" pageNumber="57" phylum="Arthropoda" rank="species" species="moskalevi">
<emphasis id="C6CE9B05FFE8FF8E552E5D7FFE1EFAE2" box="[281,489,1296,1318]" italics="true" pageId="8" pageNumber="57">Calantica moskalevi</emphasis>
<bibRefCitation id="902B3AE6FFE8FF8E55C75D7FFD18FAE2" author="Zevina" box="[496,751,1296,1318]" pageId="8" pageNumber="57" refString="Zevina, G. B. &amp; Galkin, S. V. (1989) New species of cirripeds (Cirripedia, Thoracica) from thermal waters. Zoologicheskii Zhurnal, 68 (3), 134 - 136." type="journal article" year="1989">Zevina and Galkin, 1989</bibRefCitation>
</taxonomicName>
; first topotype (view from right and left sides respectively): Two of the ten mature individuals in the sample had carinolaterals (CL2s) and the one with a CL2 on both sides was chosen as the first topotype. While the sides are virtually mirror images, it will be observed the left CL2 is smaller than that on the right (see
<figureCitation id="6C815B92FFE8FF8E51565D3DFA6BFAAD" box="[1377,1436,1362,1385]" captionStart="FIGURE 7" captionStartId="7.[151,250,1728,1751]" captionTargetBox="[196,1368,929,1688]" captionTargetId="figure@7.[188,1393,910,1714]" captionTargetPageId="7" captionText="FIGURE 7. The holotype and paratype of Calantica moskalevi Zevina and Galkin, 1989: 134, Fig. 1) consisted of two juveniles (measurements below in mm)." httpUri="https://zenodo.org/record/202507/files/figure.png" pageId="8" pageNumber="57">Fig. 7</figureCitation>
for other combinations).
</paragraph>
<paragraph id="F4054717FFE8FF8E54F05DDBFEFEF944" blockId="8.[151,1436,1460,1808]" pageId="8" pageNumber="57">
Trophi (
<figureCitation id="6C815B92FFE8FF8E55165DDBFE85FA08" box="[289,370,1460,1484]" captionStart="FIGURE 10" captionStartId="10.[151,250,1812,1835]" captionTargetBox="[176,1436,309,1770]" captionTargetId="figure@10.[151,1436,309,1788]" captionTargetPageId="10" captionText="FIGURE 10. Calantica moskalevi Zevina and Galkin, 1989; appendages, first topotype: A, portion of crest of labrum with left palp attached; B, mandible; C, first maxillae; D, second maxilla; E, intermediate articles of cirrus VI; and F, penis still attached to the coxal portion of pedicle of the left sixth cirrus." httpUri="https://zenodo.org/record/202508/files/figure.png" pageId="8" pageNumber="57">Fig. 10</figureCitation>
) with strong palps flanking bullate labrum, labral crest supporting row of numerous fine, sharp, contiguous teeth; mandible tridentoid, few well spaced spines between first and second teeth, comb of contiguous spines between third tooth and inferior angle; first maxilla with protuberant central portion of cutting edge supporting comb of closely-spaced spines separating upper field of short contiguous spines from lower field of long and short spines similar to those of inferior angle; second maxilla with typical notch between setose superior and inferior lobes.
</paragraph>
<paragraph id="F4054717FFE8FF8E54F05EE3FB0FF8D4" blockId="8.[151,1436,1460,1808]" pageId="8" pageNumber="57">
Cirri uniformly deployed along thorax (no hiatus between cirrus I and II), cirrus I with rami subequal in length, setation antenniform, remaining cirri ctenopod; intermediate articles of cirrus VI twice as high as wide (
<figureCitation id="6C815B92FFE8FF8E51145EDFFA84F90C" box="[1315,1395,1712,1736]" captionStart="FIGURE 10" captionStartId="10.[151,250,1812,1835]" captionTargetBox="[176,1436,309,1770]" captionTargetId="figure@10.[151,1436,309,1788]" captionTargetPageId="10" captionText="FIGURE 10. Calantica moskalevi Zevina and Galkin, 1989; appendages, first topotype: A, portion of crest of labrum with left palp attached; B, mandible; C, first maxillae; D, second maxilla; E, intermediate articles of cirrus VI; and F, penis still attached to the coxal portion of pedicle of the left sixth cirrus." httpUri="https://zenodo.org/record/202508/files/figure.png" pageId="8" pageNumber="57">Fig. 10</figureCitation>
E), each article supporting five strong and one weak pair of setae along inner margin, clump of four or five strong setae at distal corner of outer margin. Numbers of articles per anterior and posterior rami indicated below.
</paragraph>
<paragraph id="F4054717FFE8FF8E54A85F43FB76F850" pageId="8" pageNumber="57">
<table id="86BAB5B7FFE8007954A05F43FA6BF851" box="[151,1436,1836,1941]" gridcols="7" gridrows="3" pageId="8" pageNumber="57">
<tr id="4A8A4555FFE8007954A05F43FA6BF886" box="[151,1436,1836,1858]" gridrow="0" pageId="8" pageNumber="57">
<th id="095B2C29FFE8007954A05F43FE9FF886" box="[151,360,1836,1858]" gridcol="0" gridrow="0" pageId="8" pageNumber="57">Cirrus I</th>
<th id="095B2C29FFE8007955DC5F43FDF3F886" box="[491,516,1836,1858]" gridcol="1" gridrow="0" pageId="8" pageNumber="57">II</th>
<th id="095B2C29FFE8007956BF5F43FD55F886" box="[648,674,1836,1858]" gridcol="2" gridrow="0" pageId="8" pageNumber="57">III</th>
<th id="095B2C29FFE8007957105F43FCB5F886" box="[807,834,1836,1858]" gridcol="3" gridrow="0" pageId="8" pageNumber="57">IV</th>
<th id="095B2C29FFE8007957FF5F43FC16F886" box="[968,993,1836,1858]" gridcol="4" gridrow="0" pageId="8" pageNumber="57">V</th>
<th id="095B2C29FFE8007950505F43FB75F886" box="[1127,1154,1836,1858]" gridcol="5" gridrow="0" pageId="8" pageNumber="57">VI</th>
<th id="095B2C29FFE80079513F5F43FA6BF886" box="[1288,1436,1836,1858]" gridcol="6" gridrow="0" pageId="8" pageNumber="57">C.A.</th>
</tr>
<tr id="4A8A4555FFE8007954A05F3AFA6BF8AF" box="[151,1436,1877,1899]" gridrow="1" pageId="8" pageNumber="57">
<th id="095B2C29FFE8007954A05F3AFE9FF8AF" box="[151,360,1877,1899]" gridcol="0" gridrow="1" pageId="8" pageNumber="57">anterior 14</th>
<td id="095B2C29FFE8007955DC5F3AFDF3F8AF" box="[491,516,1877,1899]" gridcol="1" gridrow="1" pageId="8" pageNumber="57">19</td>
<td id="095B2C29FFE8007956BF5F3AFD55F8AF" box="[648,674,1877,1899]" gridcol="2" gridrow="1" pageId="8" pageNumber="57">--</td>
<td id="095B2C29FFE8007957105F3AFCB5F8AF" box="[807,834,1877,1899]" gridcol="3" gridrow="1" pageId="8" pageNumber="57">20</td>
<td id="095B2C29FFE8007957FF5F3AFC16F8AF" box="[968,993,1877,1899]" gridcol="4" gridrow="1" pageId="8" pageNumber="57">21</td>
<td id="095B2C29FFE8007950505F3AFB75F8AF" box="[1127,1154,1877,1899]" gridcol="5" gridrow="1" pageId="8" pageNumber="57">20</td>
<td id="095B2C29FFE80079513F5F3AFA6BF8AF" box="[1288,1436,1877,1899]" gridcol="6" gridrow="1" pageId="8" pageNumber="57">absent</td>
</tr>
<tr id="4A8A4555FFE8007954A05F10FA6BF851" box="[151,1436,1919,1941]" gridrow="2" pageId="8" pageNumber="57" rowspan-6="1">
<th id="095B2C29FFE8007954A05F10FE9FF851" box="[151,360,1919,1941]" gridcol="0" gridrow="2" pageId="8" pageNumber="57">posterior 14</th>
<td id="095B2C29FFE8007955DC5F10FDF3F851" box="[491,516,1919,1941]" gridcol="1" gridrow="2" pageId="8" pageNumber="57">18</td>
<td id="095B2C29FFE8007956BF5F10FD55F851" box="[648,674,1919,1941]" gridcol="2" gridrow="2" pageId="8" pageNumber="57">19</td>
<td id="095B2C29FFE8007957105F10FCB5F851" box="[807,834,1919,1941]" gridcol="3" gridrow="2" pageId="8" pageNumber="57">20</td>
<td id="095B2C29FFE8007957FF5F10FC16F851" box="[968,993,1919,1941]" gridcol="4" gridrow="2" pageId="8" pageNumber="57">20</td>
<td id="095B2C29FFE8007950505F10FB75F851" box="[1127,1154,1919,1941]" gridcol="5" gridrow="2" pageId="8" pageNumber="57">20</td>
</tr>
</table>
</paragraph>
<paragraph id="F4054717FFE9FF8F54A05D49FF1CFA7B" blockId="9.[151,1437,1318,1471]" pageId="9" pageNumber="58">
<emphasis id="C6CE9B05FFE9FF8F54A05D49FEE4FAF8" bold="true" box="[151,275,1318,1341]" pageId="9" pageNumber="58">FIGURE 9.</emphasis>
<taxonomicName id="33BA3C94FFE9FF8F552D5D48FD05FAF8" authority="Zevina and Galkin, 1989" authorityName="Zevina and Galkin" authorityYear="1989" box="[282,754,1318,1341]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="58" phylum="Arthropoda" rank="species" species="moskalevi">
<emphasis id="C6CE9B05FFE9FF8F552D5D48FE1EFAF9" box="[282,489,1319,1341]" italics="true" pageId="9" pageNumber="58">Calantica moskalevi</emphasis>
<bibRefCitation id="902B3AE6FFE9FF8F55C65D49FD05FAF8" author="Zevina" box="[497,754,1318,1340]" pageId="9" pageNumber="58" refString="Zevina, G. B. &amp; Galkin, S. V. (1989) New species of cirripeds (Cirripedia, Thoracica) from thermal waters. Zoologicheskii Zhurnal, 68 (3), 134 - 136." type="journal article" year="1989">Zevina and Galkin, 1989</bibRefCitation>
</taxonomicName>
; second and third topotypes: Of the ten mature specimens sampled two had carinolaterals (CL2s). The one having a CL2 on both sides was designated the first topotype (Fig. 8). The other (upper and lower right), having a CL2 on but one side (upper right), was designated the second topotype. The remaining eight lacked CL2s, and one (lower left, to same scale) was designated the third topotype. The remainder (not illustrated) constitute the topotype lot.
</paragraph>
<paragraph id="F4054717FFE9FF8F54F05D84FEEBF9AB" blockId="9.[151,1437,1515,2008]" pageId="9" pageNumber="58">
Caudal appendages, while figured for a juvenile by
<bibRefCitation id="902B3AE6FFE9FF8F572F5D84FB5EF9C7" author="Zevina" box="[792,1193,1515,1539]" pageId="9" pageNumber="58" refString="Zevina, G. B. &amp; Galkin, S. V. (1989) New species of cirripeds (Cirripedia, Thoracica) from thermal waters. Zoologicheskii Zhurnal, 68 (3), 134 - 136." type="journal article" year="1989" yearSuffix="e">
Zevina and Galkin (1989,
<figureCitation id="6C815B92FFE9FF8F50715D84FB7FF9C7" box="[1094,1160,1515,1539]" captionStart="FIGURE 2" captionStartId="2.[151,250,915,938]" captionTargetBox="[313,1273,363,894]" captionTargetId="figure@2.[313,1273,363,894]" captionTargetPageId="2" captionText="FIGURE 2. Panorama in the caldera of Axial Seamount, at approximately 1434 m of depth. Several yellowish-brown clumps and a few scatters individuals of the stalked barnacle, Calantica moskalevi Zevina and Galkin, 1989, can be seen at roughly 9, 12 and 3 oclock of the right red laser dot (distance between red laser dots 30 cm)." httpUri="https://zenodo.org/record/202503/files/figure.png" pageId="9" pageNumber="58">Fig. 2</figureCitation>
e)
</bibRefCitation>
, apparently absent in adult. Penis robust, short, somewhat longer than pedicle of cirrus VI (
<figureCitation id="6C815B92FFE9FF8F57995E60FC08F9E3" box="[942,1023,1551,1575]" captionStart="FIGURE 10" captionStartId="10.[151,250,1812,1835]" captionTargetBox="[176,1436,309,1770]" captionTargetId="figure@10.[151,1436,309,1788]" captionTargetPageId="10" captionText="FIGURE 10. Calantica moskalevi Zevina and Galkin, 1989; appendages, first topotype: A, portion of crest of labrum with left palp attached; B, mandible; C, first maxillae; D, second maxilla; E, intermediate articles of cirrus VI; and F, penis still attached to the coxal portion of pedicle of the left sixth cirrus." httpUri="https://zenodo.org/record/202508/files/figure.png" pageId="9" pageNumber="58">Fig. 10</figureCitation>
F), fully equipped with annulations, sensor setae and tuft of apical setae (therefore judged functionally hermaphroditic); apertural, complemental males not evident.
</paragraph>
</subSubSection>
<subSubSection id="BCA0149CFFE9FF8C54F05E13FB9AFF39" lastPageId="10" lastPageNumber="59" pageId="9" pageNumber="58" type="discussion">
<paragraph id="F4054717FFE9FF8C54F05E13FB9AFF39" blockId="9.[151,1437,1515,2008]" lastBlockId="10.[151,1436,151,254]" lastPageId="10" lastPageNumber="59" pageId="9" pageNumber="58">
<emphasis id="C6CE9B05FFE9FF8F54F05E13FECCF950" bold="true" box="[199,315,1660,1684]" pageId="9" pageNumber="58">Remarks.</emphasis>
<bibRefCitation id="902B3AE6FFE9FF8F55755E14FE3AF957" author="Gray" box="[322,461,1659,1683]" pageId="9" pageNumber="58" refString="Gray, J. E. (1825) A synopsis of the genera of cirripedes arranged in natural families, with a description of some new species. Annals of Philosophy (new ser.), 10 (2), 97 - 107." type="journal article" year="1825">Gray (1825)</bibRefCitation>
proposed the genus
<taxonomicName id="33BA3C94FFE9FF8F56815E13FCD3F950" box="[694,804,1660,1684]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFE9FF8F56815E13FCD3F950" box="[694,804,1660,1684]" italics="true" pageId="9" pageNumber="58">Calantica</emphasis>
</taxonomicName>
, generalized members of which superficially resembled
<taxonomicName id="33BA3C94FFE9FF8F54A05ECFFE28F973" authority="Seguenza, 1876" authorityName="Seguenza" authorityYear="1876" box="[151,479,1695,1720]" class="Maxillopoda" family="Calanticidae" genus="Scillaelepas" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFE9FF8F54A05ECFFEE9F97C" box="[151,286,1696,1720]" italics="true" pageId="9" pageNumber="58">Scillaelepas</emphasis>
<bibRefCitation id="902B3AE6FFE9FF8F551F5EF0FE28F973" author="Seguenza" box="[296,479,1695,1719]" pageId="9" pageNumber="58" refString="Seguenza, G. (1876) Ricerche paleontologiche intorno ai Cirripedi terziari della provincia di Messina. ParteII. Terza famiglia Lepadi Darwin. Atti dell'Accademia Pontoniana, Naples, 10, 1 - 112." type="journal article" year="1876">Seguenza, 1876</bibRefCitation>
</taxonomicName>
. They thereby fell in and out of synonymy, or were considered subgenera (
<bibRefCitation id="902B3AE6FFE9FF8F517C5EF0FF27F91F" author="Kruger" pageId="9" pageNumber="58" refString="Kruger, P. (1940) Cirripedia und Ascothoracida. In: Bronns, H. G., Klassen und Ordnungen des Tierreichs, 5 (1, 3, 3), 1 - 566. Akademische Verlagsgesellschaft, Leipzig" type="journal article" year="1940">Krüger 1940</bibRefCitation>
) until revised (
<bibRefCitation id="902B3AE6FFE9FF8F55485EACFDDCF91F" author="Newman" box="[383,555,1731,1755]" pageId="9" pageNumber="58" refString="Newman, W. A. (1980) A review of extant Scillaelepas (Cirripedia: Scalpellidae) including recognition of new species from the North Atlantic, Western Indian Ocean and New Zealand. Tethys, 9 (4), 379 - 398." type="journal article" year="1980">Newman 1980</bibRefCitation>
).
<collectingCountry id="8CAD0787FFE9FF8F56085EABFDA9F918" box="[575,606,1732,1756]" name="American Samoa" pageId="9" pageNumber="58">As</collectingCountry>
noted in the above, the name of Zevina and Galkins form, when newly proposed, included a parenthetical question mark. This was probably because the two specimens were early juveniles and, since the juveniles of distantly related genera can be quite similar, the identification was deemed uncertain. The specimens (
<figureCitation id="6C815B92FFE9FF8F55A35F40FE2BF883" box="[404,476,1839,1863]" captionStart="FIGURE 7" captionStartId="7.[151,250,1728,1751]" captionTargetBox="[196,1368,929,1688]" captionTargetId="figure@7.[188,1393,910,1714]" captionTargetPageId="7" captionText="FIGURE 7. The holotype and paratype of Calantica moskalevi Zevina and Galkin, 1989: 134, Fig. 1) consisted of two juveniles (measurements below in mm)." httpUri="https://zenodo.org/record/202507/files/figure.png" pageId="9" pageNumber="58">Fig. 7</figureCitation>
) had the usual 13 capitular plates (3 unpaired, 5 paired) which, taken alone, are diagnostic of both
<taxonomicName id="33BA3C94FFE9FF8F555E5F3BFE07F8A8" box="[361,496,1876,1900]" class="Maxillopoda" family="Calanticidae" genus="Scillaelepas" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFE9FF8F555E5F3BFE07F8A8" box="[361,496,1876,1900]" italics="true" pageId="9" pageNumber="58">Scillaelepas</emphasis>
</taxonomicName>
and
<taxonomicName id="33BA3C94FFE9FF8F56105F3BFD62F8A8" box="[551,661,1876,1900]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFE9FF8F56105F3BFD62F8A8" box="[551,661,1876,1900]" italics="true" pageId="9" pageNumber="58">Calantica</emphasis>
</taxonomicName>
(Fig. 6). However, one of the two juveniles had an additional pair of lateral plates while the other had but one, supplementary lateral plates being known in
<taxonomicName id="33BA3C94FFE9FF8F50475F17FB28F854" box="[1136,1247,1912,1936]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFE9FF8F50475F17FB28F854" box="[1136,1247,1912,1936]" italics="true" pageId="9" pageNumber="58">Calantica</emphasis>
</taxonomicName>
but not
<emphasis id="C6CE9B05FFE9FF8F51085F17FF25F870" italics="true" pageId="9" pageNumber="58">
<taxonomicName id="33BA3C94FFE9FF8F51085F17FF39F870" class="Maxillopoda" family="Calanticidae" genus="Scillaelepas" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="58" phylum="Arthropoda" rank="genus">Scillaelepas</taxonomicName>
.
</emphasis>
Curiously, only one of the
<specimenCount id="E2BC8C9EFFE9FF8F56315FF4FD96F877" box="[518,609,1947,1971]" pageId="9" pageNumber="58" type="adult">10 adult</specimenCount>
specimens in the present sample (first topotype) has an additional pair of plates, while another (second topotype) has but one of the pair, and so situated in both as to be referable to as second carinolatera (CL2; Figs 8, 9). The remainder were similar to
<taxonomicName id="33BA3C94FFEAFF8C575D58F4FC2FFF77" box="[874,984,155,179]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="10" pageNumber="59" phylum="Arthropoda" rank="species" species="studeri">
<emphasis id="C6CE9B05FFEAFF8C575D58F4FC2FFF77" box="[874,984,155,179]" italics="true" pageId="10" pageNumber="59">C. studeri</emphasis>
</taxonomicName>
or, ignoring the smallness of the peduncular plates and the cuticle covering them, like
<taxonomicName id="33BA3C94FFEAFF8C569A58D0FCC3FF13" box="[685,820,191,215]" class="Maxillopoda" family="Calanticidae" genus="Scillaelepas" kingdom="Animalia" order="Pedunculata" pageId="10" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFEAFF8C569A58D0FCC3FF13" box="[685,820,191,215]" italics="true" pageId="10" pageNumber="59">Scillaelepas</emphasis>
</taxonomicName>
(
<figureCitation id="6C815B92FFEAFF8C577258D0FC7DFF13" box="[837,906,191,215]" captionStart="FIGURE 7" captionStartId="7.[151,250,1728,1751]" captionTargetBox="[196,1368,929,1688]" captionTargetId="figure@7.[188,1393,910,1714]" captionTargetPageId="7" captionText="FIGURE 7. The holotype and paratype of Calantica moskalevi Zevina and Galkin, 1989: 134, Fig. 1) consisted of two juveniles (measurements below in mm)." httpUri="https://zenodo.org/record/202507/files/figure.png" pageId="10" pageNumber="59">Fig. 7</figureCitation>
, lower left). Whatever, the adult of Zevina and Galkins form is clearly a
<taxonomicName id="33BA3C94FFEAFF8C558D5889FDDEFF3A" box="[442,553,230,254]" class="Maxillopoda" family="Calanticidae" genus="Calantica" kingdom="Animalia" order="Pedunculata" pageId="10" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFEAFF8C558D5889FDDEFF3A" box="[442,553,230,254]" italics="true" pageId="10" pageNumber="59">Calantica</emphasis>
</taxonomicName>
as presently defined, whether CL2 is present or not.
</paragraph>
</subSubSection>
</treatment>
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<paragraph id="F4054717FFEBFF8D54A059C2FD5DFE03" blockId="11.[151,682,429,455]" box="[151,682,429,455]" pageId="11" pageNumber="60">
<heading id="AF4DF07BFFEBFF8D54A059C2FD5DFE03" bold="true" box="[151,682,429,455]" fontSize="11" level="1" pageId="11" pageNumber="60" reason="1">
<emphasis id="C6CE9B05FFEBFF8D54A059C2FD5DFE03" bold="true" box="[151,682,429,455]" pageId="11" pageNumber="60">
Subfamily
<taxonomicName id="33BA3C94FFEBFF8D552859C2FD5DFE03" authority="Utinomi, 1968" authorityName="Utinomi" authorityYear="1968" box="[287,682,429,455]" class="Maxillopoda" family="Pachylasmatidae" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="60" phylum="Arthropoda" rank="subFamily" subFamily="Pachylasmatinae">
Pachylasmatinae
<bibRefCitation id="902B3AE6FFEBFF8D55CD59C2FD5DFE03" author="Utinomi" box="[506,682,429,455]" pageId="11" pageNumber="60" refString="Utinomi, H. (1968) A revision of the deep-sea barnacles Pachylasma and Hexelasma from Japan, with a proposal of new classification of the Chthamalidae (Cirripedia, Thoracica). Publications of the Seto Marine Biological Laboratory, 16 (1), 21 - 39." type="journal article" year="1968">Utinomi, 1968</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BCA0149CFFEBFF8D54A0599BFF16FCB3" pageId="11" pageNumber="60" type="discussion">
<paragraph id="F4054717FFEBFF8D54A0599BFF16FCB3" blockId="11.[151,1436,500,887]" pageId="11" pageNumber="60">
<emphasis id="C6CE9B05FFEBFF8D54A0599BFEFCFDC8" bold="true" box="[151,267,500,524]" pageId="11" pageNumber="60">Remarks.</emphasis>
The
<taxonomicName id="33BA3C94FFEBFF8D5572599BFDF3FDC8" box="[325,516,500,524]" class="Maxillopoda" family="Pachylasmatidae" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="60" phylum="Arthropoda" rank="subFamily" subFamily="Pachylasmatinae">Pachylasmatinae</taxonomicName>
, as per the landmark revision of
<bibRefCitation id="902B3AE6FFEBFF8D5741599AFBFEFDC8" author="Jones" box="[886,1033,500,524]" pageId="11" pageNumber="60" refString="Jones, D. S. (2000) Crustacea Cirripedia Thoracica: Chionelasmatoidea and Pachylasmatoidea (Balanomorpha) of New Caledonia, Vanuatu and Wallis and Futuna Islands, with a review of all currently assigned taxa. In: Crosnier, A. (ed.), Resultats des Campagne MUSORSTOM 21. Memoires du Museum national d'Historie naturelle, 184, 141 - 283." type="journal article" year="2000">Jones (2000)</bibRefCitation>
, encompasses five genera;
<taxonomicName id="33BA3C94FFEBFF8D510B599AFE51FDEB" authority="Jones, 2000: 166" authorityName="Jones" authorityPageNumber="166" authorityYear="2000" class="Maxillopoda" family="Pachylasmatidae" genus="Eutomolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFEBFF8D510B599AFF2DFDF4" italics="true" pageId="11" pageNumber="60">Eutomolasma</emphasis>
<bibRefCitation id="902B3AE6FFEBFF8D54D35A77FE99FDEB" author="Jones" box="[228,366,535,559]" pageId="11" pageNumber="60" refString="Jones, D. S. (2000) Crustacea Cirripedia Thoracica: Chionelasmatoidea and Pachylasmatoidea (Balanomorpha) of New Caledonia, Vanuatu and Wallis and Futuna Islands, with a review of all currently assigned taxa. In: Crosnier, A. (ed.), Resultats des Campagne MUSORSTOM 21. Memoires du Museum national d'Historie naturelle, 184, 141 - 283." type="journal article" year="2000">Jones, 2000</bibRefCitation>
: 166
</taxonomicName>
(four species),
<taxonomicName id="33BA3C94FFEBFF8D566B5A77FC58FDEB" authority="Jones, 2000: 177" authorityName="Jones" authorityPageNumber="177" authorityYear="2000" box="[604,943,535,560]" class="Maxillopoda" family="Pachylasmatidae" genus="Microlasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFEBFF8D566B5A77FD15FDF4" box="[604,738,536,560]" italics="true" pageId="11" pageNumber="60">Microlasma</emphasis>
<bibRefCitation id="902B3AE6FFEBFF8D56DB5A77FC80FDEB" author="Jones" box="[748,887,535,559]" pageId="11" pageNumber="60" refString="Jones, D. S. (2000) Crustacea Cirripedia Thoracica: Chionelasmatoidea and Pachylasmatoidea (Balanomorpha) of New Caledonia, Vanuatu and Wallis and Futuna Islands, with a review of all currently assigned taxa. In: Crosnier, A. (ed.), Resultats des Campagne MUSORSTOM 21. Memoires du Museum national d'Historie naturelle, 184, 141 - 283." type="journal article" year="2000">Jones, 2000</bibRefCitation>
: 177
</taxonomicName>
(four species),
<taxonomicName id="33BA3C94FFEBFF8D50525A77FF36FD90" authority="Darwin, 1854: 185" authorityName="Darwin" authorityPageNumber="185" authorityYear="1854" class="Maxillopoda" family="Pachylasmatidae" genus="Pachylasma" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFEBFF8D50525A77FB19FDF4" box="[1125,1262,536,560]" italics="true" pageId="11" pageNumber="60">Pachylasma</emphasis>
<bibRefCitation id="902B3AE6FFEBFF8D50C05A78FA6FFDEB" author="Darwin" box="[1271,1432,535,559]" pageId="11" pageNumber="60" refString="Darwin, C. (1854) A monograph on the subclass Cirripedia, with figures of all species, the Balanidae, the Verrucidae etc. Pp 1 - 684 + pls 1 - 30. Ray Society, London." type="book chapter" year="1854">Darwin, 1854</bibRefCitation>
: 185
</taxonomicName>
(eight species),
<taxonomicName id="33BA3C94FFEBFF8D554C5A53FD4EFD90" authority="Jones, 2000: 201" authorityName="Jones" authorityPageNumber="201" authorityYear="2000" box="[379,697,572,596]" class="Maxillopoda" family="Pachylasmatidae" genus="Eurylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFEBFF8D554C5A53FE03FD90" box="[379,500,572,596]" italics="true" pageId="11" pageNumber="60">Eurylasma</emphasis>
<bibRefCitation id="902B3AE6FFEBFF8D55CB5A52FD72FD90" author="Jones" box="[508,645,572,596]" pageId="11" pageNumber="60" refString="Jones, D. S. (2000) Crustacea Cirripedia Thoracica: Chionelasmatoidea and Pachylasmatoidea (Balanomorpha) of New Caledonia, Vanuatu and Wallis and Futuna Islands, with a review of all currently assigned taxa. In: Crosnier, A. (ed.), Resultats des Campagne MUSORSTOM 21. Memoires du Museum national d'Historie naturelle, 184, 141 - 283." type="journal article" year="2000">Jones, 2000</bibRefCitation>
: 201
</taxonomicName>
(three species), and
<taxonomicName id="33BA3C94FFEBFF8D57AB5A53FADEFD90" authority="Foster, 1988: 215" authorityName="Foster" authorityPageNumber="215" authorityYear="1988" box="[924,1321,572,596]" class="Maxillopoda" family="Pachylasmatidae" genus="Tetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFEBFF8D57AB5A53FBABFD90" box="[924,1116,572,596]" italics="true" pageId="11" pageNumber="60">Tetrapachylasma</emphasis>
<bibRefCitation id="902B3AE6FFEBFF8D50535A52FB04FD90" author="Foster" box="[1124,1267,572,596]" pageId="11" pageNumber="60" refString="Foster, B. A. (1988) Tetrapachylasma: A new genus for a shallow water pachylasmatid barnacle from the Cook Islands. Crustaceana, 55 (3), 225 - 233." type="journal article" year="1988">Foster, 1988</bibRefCitation>
: 215
</taxonomicName>
(five species). All, except the
<typeStatus id="2B01F9B5FFEBFF8D55BC5A0EFE4CFDBD" box="[395,443,609,633]" pageId="11" pageNumber="60">type</typeStatus>
of
<taxonomicName id="33BA3C94FFEBFF8D55D45A0FFD54FDBC" box="[483,675,608,632]" class="Maxillopoda" family="Pachylasmatidae" genus="Tetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFEBFF8D55D45A0FFD54FDBC" box="[483,675,608,632]" italics="true" pageId="11" pageNumber="60">Tetrapachylasma</emphasis>
</taxonomicName>
, have an eight-plated stage early in ontogeny, albeit the rostrum is generally tripartite, RL-R-RL, and the carinolaterals, CL1-CL2, are partially concrescent. These tendencies are preludes to the pure four-platedness seen in
<taxonomicName id="33BA3C94FFEBFF8D56495AC4FCC9FD07" box="[638,830,683,707]" class="Maxillopoda" family="Pachylasmatidae" genus="Tetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="60" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="C6CE9B05FFEBFF8D56495AC4FCC9FD07" box="[638,830,683,707]" italics="true" pageId="11" pageNumber="60">Tetrapachylasma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDFD267EFFEBFF8D577E5AC4FC9AFD06" box="[841,877,683,706]" pageId="11" pageNumber="60" sensu="stricto">s.s.</taxonomicNameLabel>
Therefore, acceptance of
<taxonomicName id="33BA3C94FFEBFF8D50955AC4FA95FD07" box="[1186,1378,683,707]" class="Maxillopoda" family="Pachylasmatidae" genus="Tetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFEBFF8D50955AC4FA95FD07" box="[1186,1378,683,707]" italics="true" pageId="11" pageNumber="60">Tetrapachylasma</emphasis>
</taxonomicName>
as a monophyletic taxon must be viewed with caution. The wall of the new genus, to be proposed below, is like that of a
<taxonomicName id="33BA3C94FFEBFF8D549C5A9CFE9CFCCF" box="[171,363,755,779]" class="Maxillopoda" family="Pachylasmatidae" genus="Tetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="60" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="C6CE9B05FFEBFF8D549C5A9CFE9CFCCF" box="[171,363,755,779]" italics="true" pageId="11" pageNumber="60">Tetrapachylasma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDFD267EFFEBFF8D55455A9CFE61FCCE" box="[370,406,755,778]" pageId="11" pageNumber="60" sensu="stricto">s.s.</taxonomicNameLabel>
in that it displays not a hint of having had more than four plates making up its wall early in ontogeny. On the other hand, as we shall see, it displays the most generalized opercular plates known to any balanomorph and, therefore, stands well apart from other pachylasmatines including the other species of
<taxonomicName id="33BA3C94FFEBFF8D51225B54FF2DFCB3" class="Maxillopoda" family="Pachylasmatidae" genus="Tetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFEBFF8D51225B54FF2DFCB3" italics="true" pageId="11" pageNumber="60">Tetrapachylasma</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,92 @@
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<treatment id="7C13F601FFEBFF8D54A05BA5FD14FA88" ID-DOI="http://doi.org/10.5281/zenodo.5667573" ID-GBIF-Taxon="119392594" ID-Zenodo-Dep="5667573" LSID="urn:lsid:zoobank.org:act:14AB0CB6-5D96-418B-A250-2D57FBFDDFC8" httpUri="http://treatment.plazi.org/id/7C13F601FFEBFF8D54A05BA5FD14FA88" lastPageNumber="60" pageId="11" pageNumber="60">
<subSubSection id="BCA0149CFFEBFF8D54A05BA5FE11FC20" box="[151,486,970,996]" pageId="11" pageNumber="60" type="nomenclature">
<paragraph id="F4054717FFEBFF8D54A05BA5FE11FC20" blockId="11.[151,486,970,996]" box="[151,486,970,996]" pageId="11" pageNumber="60">
<heading id="AF4DF07BFFEBFF8D54A05BA5FE11FC20" bold="true" box="[151,486,970,996]" fontSize="11" level="1" pageId="11" pageNumber="60" reason="1">
<emphasis id="C6CE9B05FFEBFF8D54A05BA5FE11FC20" bold="true" box="[151,486,970,996]" pageId="11" pageNumber="60">
<taxonomicName id="33BA3C94FFEBFF8D54A05BA5FE86FC20" box="[151,369,970,996]" class="Maxillopoda" family="Pachylasmatidae" genus="Atetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="60" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="C6CE9B05FFEBFF8D54A05BA5FE86FC20" bold="true" box="[151,369,970,996]" italics="true" pageId="11" pageNumber="60">Atetrapachylasma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDFD267EFFEBFF8D554F5BA5FE11FC20" box="[376,486,970,996]" pageId="11" pageNumber="60" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BCA0149CFFEBFF8D54A05C7EFD7DFB24" pageId="11" pageNumber="60" type="diagnosis">
<paragraph id="F4054717FFEBFF8D54A05C7EFD7DFB24" blockId="11.[151,1437,1041,1356]" pageId="11" pageNumber="60">
<emphasis id="C6CE9B05FFEBFF8D54A05C7EFEE6FBEE" bold="true" box="[151,273,1041,1066]" pageId="11" pageNumber="60">Diagnosis.</emphasis>
Wall of female of four plates (
<figureCitation id="6C815B92FFEBFF8D56505C7EFD4AFBED" box="[615,701,1041,1065]" captionStart="FIGURE 11" captionStartId="12.[151,250,1257,1280]" captionTargetBox="[341,1239,759,1225]" captionTargetId="figure@12.[317,1269,727,1235]" captionTargetPageId="12" captionText="FIGURE 11. Atetrapachylasma dijonesae gen. et sp. nov. holotype: A, exterior view of female with males; note largely exposed alae and the pair of males on each side of the tight fitting occludent ridges of the scuta. B, interior view of the same female, note the tight fit of the bearing surfaces of the alae on the adjacent plates, and the close fit of the sutures between plates, especially between the opercular plates and the interior of the orifice." httpUri="https://zenodo.org/record/202509/files/figure.png" pageId="11" pageNumber="60">Figs 11</figureCitation>
,
<figureCitation id="6C815B92FFEBFF8D56FF5C7EFD10FBED" box="[712,743,1041,1065]" captionStart="FIGURE 12" captionStartId="13.[151,250,1312,1335]" captionTargetBox="[370,1206,224,1261]" captionTargetId="figure@13.[348,1239,193,1291]" captionTargetPageId="13" captionText="FIGURE 12. Atetrapachylasma dijonesae gen. et sp. nov., exploded four-plated wall arranged from loose plates, type lot: A, exterior of a rostrum; B and D, left and right carinolaterals; C, a carina; and E, the interior of a portion of a rostrum. Note the variably pronounced depression in the sheath of rostrum (E), which, together with the edges of the carinolateral alae, receives the rostral ends of the occludent ridges of the scuta (Figs 9 A; 11 C, D)." httpUri="https://zenodo.org/record/202510/files/figure.png" pageId="11" pageNumber="60">12</figureCitation>
), compound rostrum (RLRRL), paired compound carinolatals (CL1CL2s) and carina (C); compound plates apparently without sutural traces in dwarf males as well as females. Wall low conical, wider than high; overall round plan constricted at sutures giving the four parietal parts a somewhat lobed appearance (
<figureCitation id="6C815B92FFEBFF8D55875CEFFE0AFB5C" box="[432,509,1152,1176]" captionStart="FIGURE 5" captionStartId="6.[151,250,1830,1853]" captionTargetBox="[295,1289,199,1795]" captionTargetId="figure@6.[284,1301,193,1810]" captionTargetPageId="6" captionText="FIGURE 5. Atetrapachylasma dijonesae gen. et sp. nov. in situ on pillow basalt from 2080 m of depth Vance B Seamount, with nozzle of suction sampler of ROV Tiburon largely to their right. A, the remains of several walls broken away by the nozzle can be seen to the right of “ 1 ” leaving membranous bases exposed. B, the barnacles seen between “ 2 ” and “ 3 ” and to the right of “ 4 ” in A have been collected. N. B.: Inspection of MPEGs taken by ROV Tiburon during collection of the barnacles seen removed in these frames detected no loss of material in the process. Therefore, it can be said with confidence that the shell fragments at the base of the basalt below the barnacles (cf. Fig. 4) represent natural postmortem accumulations." httpUri="https://zenodo.org/record/202506/files/figure.png" pageId="11" pageNumber="60">Figs 5</figureCitation>
,
<figureCitation id="6C815B92FFEBFF8D563B5CEFFDDDFB5C" box="[524,554,1152,1176]" captionStart="FIGURE 11" captionStartId="12.[151,250,1257,1280]" captionTargetBox="[341,1239,759,1225]" captionTargetId="figure@12.[317,1269,727,1235]" captionTargetPageId="12" captionText="FIGURE 11. Atetrapachylasma dijonesae gen. et sp. nov. holotype: A, exterior view of female with males; note largely exposed alae and the pair of males on each side of the tight fitting occludent ridges of the scuta. B, interior view of the same female, note the tight fit of the bearing surfaces of the alae on the adjacent plates, and the close fit of the sutures between plates, especially between the opercular plates and the interior of the orifice." httpUri="https://zenodo.org/record/202509/files/figure.png" pageId="11" pageNumber="60">11</figureCitation>
). Opercular plates simple, triangular in outline, articular ridges and furrows weakly developed; tergum without depressor muscles crests; scutum with strong occludent ridge visible externally and internally, adductor pit and ridge absent.
</paragraph>
</subSubSection>
<subSubSection id="BCA0149CFFEBFF8D54F05C82FD14FA88" pageId="11" pageNumber="60" type="etymology">
<paragraph id="F4054717FFEBFF8D54F05C82FD14FA88" blockId="11.[151,1437,1041,1356]" pageId="11" pageNumber="60">
<emphasis id="C6CE9B05FFEBFF8D54F05C82FEB9FAC1" bold="true" box="[199,334,1261,1285]" pageId="11" pageNumber="60">Etymology.</emphasis>
“A-”
<emphasis id="C6CE9B05FFEBFF8D55A45C82FE40FAC0" box="[403,439,1261,1284]" italics="true" pageId="11" pageNumber="60">not</emphasis>
, in preference to “Para-“
<emphasis id="C6CE9B05FFEBFF8D56EF5C83FCD7FAC0" box="[728,800,1260,1284]" italics="true" pageId="11" pageNumber="60">beside</emphasis>
or
<emphasis id="C6CE9B05FFEBFF8D57715C82FC8EFAC0" box="[838,889,1261,1284]" italics="true" pageId="11" pageNumber="60">near</emphasis>
“, since despite also being four-plated and without traces of vestigial sutures, it is otherwise evidently not closely related to the
<typeStatus id="2B01F9B5FFEBFF8D501A5D7EFB97FAED" box="[1069,1120,1297,1321]" pageId="11" pageNumber="60">type</typeStatus>
,
<taxonomicName id="33BA3C94FFEBFF8D505A5D7FFEABFA88" authority="Foster, 1988: 225" authorityName="Foster" authorityPageNumber="225" authorityYear="1988" class="Maxillopoda" family="Pachylasmatidae" genus="Tetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="trigonum">
<emphasis id="C6CE9B05FFEBFF8D505A5D7FFA6BFAEC" box="[1133,1436,1296,1320]" italics="true" pageId="11" pageNumber="60">Tetrapachylasma trigonum</emphasis>
<bibRefCitation id="902B3AE6FFEBFF8D54A05D5AFED1FA88" author="Foster" box="[151,294,1332,1356]" pageId="11" pageNumber="60" refString="Foster, B. A. (1988) Tetrapachylasma: A new genus for a shallow water pachylasmatid barnacle from the Cook Islands. Crustaceana, 55 (3), 225 - 233." type="journal article" year="1988">Foster, 1988</bibRefCitation>
: 225
</taxonomicName>
, much less to
<taxonomicName id="33BA3C94FFEBFF8D55CC5D5BFD4CFA88" box="[507,699,1332,1356]" class="Maxillopoda" family="Pachylasmatidae" genus="Tetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="60" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="C6CE9B05FFEBFF8D55CC5D5BFD4CFA88" box="[507,699,1332,1356]" italics="true" pageId="11" pageNumber="60">Tetrapachylasma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDFD267EFFEBFF8D56F55D5BFD14FA88" box="[706,739,1332,1356]" pageId="11" pageNumber="60" sensu="lato">s.l.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,428 @@
<document id="E8028478EC5951C6A1D4B7DB96791A36" ID-DOI="10.5281/zenodo.202501" ID-GBIF-Dataset="86a4e805-beba-4f05-9da2-7030b8addcea" ID-ISSN="1175-5326" ID-Zenodo-Dep="202501" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460256919205" checkinUser="plazi" docAuthor="Newman, William A. &amp; Jones, William J." docDate="2011" docId="7C13F601FFEBFF9654A05DCFFAF0FC98" docLanguage="en" docName="zt02789p068.pdf" docOrigin="Zootaxa 2789" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Atetrapachylasma dijonesae Newman &amp; Jones, 2011, sp. nov." docType="treatment" docVersion="8" lastPageNumber="65" masterDocId="802A8E79FFE0FF865437586FFFF7FFC4" masterDocTitle="Two Northeast Pacific deep-water barnacle populations (Cirripedia: Calanticidae and Pachylasmatidae) from seamounts of the Juan de Fuca Ridge; &quot; insular &quot; endemics stemming from Tethys, or by subsequent dispersal from the Western Pacific center of distribution?" masterLastPageNumber="68" masterPageNumber="49" pageNumber="60" updateTime="1698243655103" updateUser="plazi">
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<mods:title id="08159BAC7CDDCCC01DAEB0AAF851822F">Two Northeast Pacific deep-water barnacle populations (Cirripedia: Calanticidae and Pachylasmatidae) from seamounts of the Juan de Fuca Ridge; &quot; insular &quot; endemics stemming from Tethys, or by subsequent dispersal from the Western Pacific center of distribution?</mods:title>
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<mods:namePart id="6C25739B203271D62F53C2C52F1D891D">Newman, William A.</mods:namePart>
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<mods:namePart id="310368A958DBF70BCC646679C908A3DE">Jones, William J.</mods:namePart>
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<mods:date id="BB5B1BAE467BD1F10B2B73CE62AC1B69">2011</mods:date>
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<treatment id="7C13F601FFEBFF9654A05DCFFAF0FC98" ID-DOI="http://doi.org/10.5281/zenodo.5667575" ID-GBIF-Taxon="119392595" ID-Zenodo-Dep="5667575" LSID="urn:lsid:zoobank.org:act:17A58A13-3DFF-4AC9-94F2-1C2259882647" httpUri="http://treatment.plazi.org/id/7C13F601FFEBFF9654A05DCFFAF0FC98" lastPageId="16" lastPageNumber="65" pageId="11" pageNumber="60">
<subSubSection id="BCA0149CFFEBFF8D54A05DCFFE98FA1F" pageId="11" pageNumber="60" type="nomenclature">
<paragraph id="F4054717FFEBFF8D54A05DCFFDB8FA7E" blockId="11.[151,591,1440,1499]" box="[151,591,1440,1466]" pageId="11" pageNumber="60">
<heading id="AF4DF07BFFEBFF8D54A05DCFFDB8FA7E" bold="true" box="[151,591,1440,1466]" fontSize="11" level="1" pageId="11" pageNumber="60" reason="1">
<emphasis id="C6CE9B05FFEBFF8D54A05DCFFDB8FA7E" bold="true" box="[151,591,1440,1466]" pageId="11" pageNumber="60">
<taxonomicName id="33BA3C94FFEBFF8D54A05DCFFE1DFA7E" box="[151,490,1440,1466]" class="Maxillopoda" family="Pachylasmatidae" genus="Atetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="dijonesae" status="sp. nov.">
<emphasis id="C6CE9B05FFEBFF8D54A05DCFFE1DFA7E" bold="true" box="[151,490,1440,1466]" italics="true" pageId="11" pageNumber="60">Atetrapachylasma dijonesae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDFD267EFFEBFF8D55C55DCFFDB8FA7E" box="[498,591,1440,1466]" pageId="11" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="F4054717FFEBFF8D54A05DACFE98FA1F" blockId="11.[151,591,1440,1499]" box="[151,367,1475,1499]" pageId="11" pageNumber="60">
<figureCitation id="6C815B92FFEBFF8D54A05DACFEF5FA1F" box="[151,258,1475,1499]" captionStart="FIGURE 4" captionStartId="5.[151,250,1070,1093]" captionTargetBox="[256,1323,193,1021]" captionTargetId="figure@5.[256,1323,193,1022]" captionTargetPageId="5" captionText="FIGURE 4. Numerous females of Atetrapachylasma dijonesae gen. et sp. nov. in situ on pillow basalt from 2080 m of depth on Vance B Seamount, with nozzle of suction sampler on ROV Tiburon to their right (distance between red laser dots 30 cm). The pristine white appearance of the dead shells accumulating at the base of the slope, as well as of the living individuals, suggests that there is little if any hydrothermal activity nearby to stain them (rather than the barnacle just being short lived, which they are). Thus, as might be expected, dissection revealed feeding appendages (Fig. 13) of the deep-sea rather than hydrothermal type. N. B.: Inspection of MPEGs taken by ROV Tiburon during collection of most of the barnacles seen in this frame (also, Fig. 5) detected no loss of material in the process. Therefore, it can be said with confidence that the shell fragments at the base of the basalt below the barnacles represent natural, postmortem accumulations." httpUri="https://zenodo.org/record/202505/files/figure.png" pageId="11" pageNumber="60">Figures 4</figureCitation>
,
<figureCitation id="6C815B92FFEBFF8D55395DACFEE9FA1F" box="[270,286,1475,1499]" captionStart="FIGURE 5" captionStartId="6.[151,250,1830,1853]" captionTargetBox="[295,1289,199,1795]" captionTargetId="figure@6.[284,1301,193,1810]" captionTargetPageId="6" captionText="FIGURE 5. Atetrapachylasma dijonesae gen. et sp. nov. in situ on pillow basalt from 2080 m of depth Vance B Seamount, with nozzle of suction sampler of ROV Tiburon largely to their right. A, the remains of several walls broken away by the nozzle can be seen to the right of “ 1 ” leaving membranous bases exposed. B, the barnacles seen between “ 2 ” and “ 3 ” and to the right of “ 4 ” in A have been collected. N. B.: Inspection of MPEGs taken by ROV Tiburon during collection of the barnacles seen removed in these frames detected no loss of material in the process. Therefore, it can be said with confidence that the shell fragments at the base of the basalt below the barnacles (cf. Fig. 4) represent natural postmortem accumulations." httpUri="https://zenodo.org/record/202506/files/figure.png" pageId="11" pageNumber="60">5</figureCitation>
,
<figureCitation id="6C815B92FFEBFF8D551D5DACFE98FA1F" box="[298,367,1475,1499]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStart-2="FIGURE 12" captionStart-3="FIGURE 13" captionStart-4="FIGURE 14" captionStart-5="FIGURE 15" captionStartId-0="10.[151,250,1812,1835]" captionStartId-1="12.[151,250,1257,1280]" captionStartId-2="13.[151,250,1312,1335]" captionStartId-3="14.[151,250,1865,1888]" captionStartId-4="15.[151,250,645,668]" captionStartId-5="15.[151,250,1974,1997]" captionTargetBox-0="[176,1436,309,1770]" captionTargetBox-1="[341,1239,759,1225]" captionTargetBox-2="[370,1206,224,1261]" captionTargetBox-3="[323,1234,441,1824]" captionTargetBox-4="[374,1237,208,614]" captionTargetBox-5="[189,1382,809,1944]" captionTargetId-0="figure@10.[151,1436,309,1788]" captionTargetId-1="figure@12.[317,1269,727,1235]" captionTargetId-2="figure@13.[348,1239,193,1291]" captionTargetId-3="figure@14.[323,1264,409,1845]" captionTargetId-4="figure@15.[339,1247,193,624]" captionTargetId-5="figure@15.[185,1390,791,1951]" captionTargetPageId-0="10" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionTargetPageId-4="15" captionTargetPageId-5="15" captionText-0="FIGURE 10. Calantica moskalevi Zevina and Galkin, 1989; appendages, first topotype: A, portion of crest of labrum with left palp attached; B, mandible; C, first maxillae; D, second maxilla; E, intermediate articles of cirrus VI; and F, penis still attached to the coxal portion of pedicle of the left sixth cirrus." captionText-1="FIGURE 11. Atetrapachylasma dijonesae gen. et sp. nov. holotype: A, exterior view of female with males; note largely exposed alae and the pair of males on each side of the tight fitting occludent ridges of the scuta. B, interior view of the same female, note the tight fit of the bearing surfaces of the alae on the adjacent plates, and the close fit of the sutures between plates, especially between the opercular plates and the interior of the orifice." captionText-2="FIGURE 12. Atetrapachylasma dijonesae gen. et sp. nov., exploded four-plated wall arranged from loose plates, type lot: A, exterior of a rostrum; B and D, left and right carinolaterals; C, a carina; and E, the interior of a portion of a rostrum. Note the variably pronounced depression in the sheath of rostrum (E), which, together with the edges of the carinolateral alae, receives the rostral ends of the occludent ridges of the scuta (Figs 9 A; 11 C, D)." captionText-3="FIGURE 13. Atetrapachylasma dijonesae gen. et sp. nov., opercular plates of female: A and B, interior and exterior of terga, and C and D, interior and exterior of scuta. Both the terga and scuta are isosceles triangles, the bases of which are formed by their occludent margins. Note that the weakly developed articular ridges and furrows, as well as the lack of adductor and depressor muscle crests (apodemes), pits or scars, are remarkable. Comparably simple opercular plates are seen in no other balanomorph." captionText-4="FIGURE 14. Atetrapachylasma dijonesae gen. et sp. nov., dwarf males: A, viewed obliquely (R-C axis running between 10: 30 and 4: 30 oclock); B, another individual, viewed from the carinal end, leaning markedly to its right, toward the aperture of the female (see Fig. 9 for four males in situ). Note that the chitinous primary wall is made up of four plates, with the rostrum overlapping the carinolaterals. There is no indication of sutures in the rostrum or the carinolaterals of the male or the female." captionText-5="FIGURE 15. Atetrapachylasma dijonesae gen. et sp. nov., appendages: A, mandible; B, first maxillae; C, second maxilla; D, intermediate articles of cirrus VI; and E, crest of labrum flanked by left palp." httpUri-0="https://zenodo.org/record/202508/files/figure.png" httpUri-1="https://zenodo.org/record/202509/files/figure.png" httpUri-2="https://zenodo.org/record/202510/files/figure.png" httpUri-3="https://zenodo.org/record/202511/files/figure.png" httpUri-4="https://zenodo.org/record/202512/files/figure.png" httpUri-5="https://zenodo.org/record/202513/files/figure.png" pageId="11" pageNumber="60">1015</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BCA0149CFFEBFF8D54A05E65FBBFF94B" pageId="11" pageNumber="60" type="materials_examined">
<paragraph id="F4054717FFEBFF8D54A05E65FE81F983" blockId="11.[151,1436,1546,2003]" pageId="11" pageNumber="60">
<emphasis id="C6CE9B05FFEBFF8D54A05E65FE75F9E7" bold="true" box="[151,386,1546,1571]" pageId="11" pageNumber="60">Material examined.</emphasis>
Vance B Seamount (summit
<quantity id="3342EAF2FFEBFF8D56F85E64FCDEF9E7" box="[719,809,1547,1571]" metricMagnitude="3" metricUnit="m" metricValue="1.932" pageId="11" pageNumber="60" unit="m" value="1932.0">1932 m</quantity>
),
<collectionCode id="92ABDFD2FFEBFF8D57095E64FC6DF9E7" box="[830,922,1547,1571]" pageId="11" pageNumber="60">MBARI</collectionCode>
ROV dive nos T1012 and T1013,
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° 30'N
<geoCoordinate id="918E21D0FFEBFF8D54A05E40FEF8F983" box="[151,271,1583,1607]" direction="west" orientation="longitude" pageId="11" pageNumber="60" precision="925" value="-130.66667">130° 40'W</geoCoordinate>
,
<quantity id="3342EAF2FFEBFF8D552D5E40FE85F983" box="[282,370,1583,1607]" metricMagnitude="3" metricUnit="m" metricValue="2.081" pageId="11" pageNumber="60" unit="m" value="2081.0">2081 m</quantity>
.
</paragraph>
<paragraph id="F4054717FFEBFF8D54F25E3CFBBFF94B" blockId="11.[151,1436,1546,2003]" pageId="11" pageNumber="60">
Deposition of
<typeStatus id="2B01F9B5FFEBFF8D555F5E3BFE54F9A8" box="[360,419,1620,1644]" pageId="11" pageNumber="60">types</typeStatus>
selected from dive no. T1012:
<typeStatus id="2B01F9B5FFEBFF8D573E5E3CFC83F9AF" box="[777,884,1619,1643]" pageId="11" pageNumber="60" type="holotype">Holotype</typeStatus>
,
<collectionCode id="92ABDFD2FFEBFF8D57B65E3CFC25F9AF" LSID="urn:lsid:biocol.org:col:34871" box="[897,978,1619,1643]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="60">USNM</collectionCode>
cat. no. 1145902; first
<typeStatus id="2B01F9B5FFEBFF8D50EC5E3BFAC9F9A8" box="[1243,1342,1620,1644]" pageId="11" pageNumber="60" type="paratype">paratype</typeStatus>
,
<collectionCode id="92ABDFD2FFEBFF8D517C5E3CFA6BF9AF" LSID="urn:lsid:biocol.org:col:34871" box="[1355,1436,1619,1643]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="60">USNM</collectionCode>
cat. no. 1145903; second
<typeStatus id="2B01F9B5FFEBFF8D55835E17FDE3F954" box="[436,532,1656,1680]" pageId="11" pageNumber="60" type="paratype">paratype</typeStatus>
<collectionCode id="92ABDFD2FFEBFF8D562C5E18FDBFF94B" LSID="urn:lsid:biocol.org:col:14844" box="[539,584,1655,1679]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14844" name="Scripps Institution of Oceanography" pageId="11" pageNumber="60">SIO</collectionCode>
<collectionCode id="92ABDFD2FFEBFF8D56785E18FD89F94B" box="[591,638,1655,1679]" pageId="11" pageNumber="60">BIC</collectionCode>
cat. no. 11195;
<typeStatus id="2B01F9B5FFEBFF8D57075E17FC97F954" box="[816,864,1656,1680]" pageId="11" pageNumber="60">type</typeStatus>
lot,
<collectionCode id="92ABDFD2FFEBFF8D57A45E18FC37F94B" LSID="urn:lsid:biocol.org:col:14844" box="[915,960,1655,1679]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14844" name="Scripps Institution of Oceanography" pageId="11" pageNumber="60">SIO</collectionCode>
<collectionCode id="92ABDFD2FFEBFF8D57F05E18FC01F94B" box="[967,1014,1655,1679]" pageId="11" pageNumber="60">BIC</collectionCode>
11196.
</paragraph>
</subSubSection>
<subSubSection id="BCA0149CFFEBFF8D54F05EF5FDFFF970" box="[199,520,1690,1716]" pageId="11" pageNumber="60" type="diagnosis">
<paragraph id="F4054717FFEBFF8D54F05EF5FDFFF970" blockId="11.[151,1436,1546,2003]" box="[199,520,1690,1716]" pageId="11" pageNumber="60">
<emphasis id="C6CE9B05FFEBFF8D54F05EF5FEB6F977" bold="true" box="[199,321,1690,1715]" pageId="11" pageNumber="60">Diagnosis.</emphasis>
<collectingCountry id="8CAD0787FFEBFF8D557F5EF4FE90F977" box="[328,359,1691,1715]" name="American Samoa" pageId="11" pageNumber="60">As</collectingCountry>
for the genus.
</paragraph>
</subSubSection>
<subSubSection id="BCA0149CFFEBFF8A54F05ED1FD93F9F4" lastPageId="12" lastPageNumber="61" pageId="11" pageNumber="60" type="description">
<paragraph id="F4054717FFEBFF8D54F05ED1FD8AF84F" blockId="11.[151,1436,1546,2003]" pageId="11" pageNumber="60">
<emphasis id="C6CE9B05FFEBFF8D54F05ED1FEAEF913" bold="true" box="[199,345,1726,1751]" pageId="11" pageNumber="60">Description.</emphasis>
Female shells white, wall to
<quantity id="3342EAF2FFEBFF8D56AB5ED0FD1DF913" box="[668,746,1727,1751]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="60" unit="mm" value="20.0">20 mm</quantity>
R-C diameter,
<quantity id="3342EAF2FFEBFF8D57A35ED0FC15F913" box="[916,994,1727,1751]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="60" unit="mm" value="15.0">15 mm</quantity>
height. Narrow overlapping margins of R and CL resting on broad, largely exposed alae of CL and C, respectively (
<figureCitation id="6C815B92FFEBFF8D57CB5E8CFBB8F93F" box="[1020,1103,1763,1787]" captionStart="FIGURE 11" captionStartId="12.[151,250,1257,1280]" captionTargetBox="[341,1239,759,1225]" captionTargetId="figure@12.[317,1269,727,1235]" captionTargetPageId="12" captionText="FIGURE 11. Atetrapachylasma dijonesae gen. et sp. nov. holotype: A, exterior view of female with males; note largely exposed alae and the pair of males on each side of the tight fitting occludent ridges of the scuta. B, interior view of the same female, note the tight fit of the bearing surfaces of the alae on the adjacent plates, and the close fit of the sutures between plates, especially between the opercular plates and the interior of the orifice." httpUri="https://zenodo.org/record/202509/files/figure.png" pageId="11" pageNumber="60">Fig. 11</figureCitation>
), kite-shaped operculum suspended snuggly in orifice with rostral ends of S fitting depression in t rostral sheath (
<figureCitation id="6C815B92FFEBFF8D50675F68FB54F8DB" box="[1104,1187,1799,1823]" captionStart="FIGURE 12" captionStartId="13.[151,250,1312,1335]" captionTargetBox="[370,1206,224,1261]" captionTargetId="figure@13.[348,1239,193,1291]" captionTargetPageId="13" captionText="FIGURE 12. Atetrapachylasma dijonesae gen. et sp. nov., exploded four-plated wall arranged from loose plates, type lot: A, exterior of a rostrum; B and D, left and right carinolaterals; C, a carina; and E, the interior of a portion of a rostrum. Note the variably pronounced depression in the sheath of rostrum (E), which, together with the edges of the carinolateral alae, receives the rostral ends of the occludent ridges of the scuta (Figs 9 A; 11 C, D)." httpUri="https://zenodo.org/record/202510/files/figure.png" pageId="11" pageNumber="60">Fig. 12</figureCitation>
). Occludent margin at carinal end of T with fine teeth formed by alternate growth lines interdigitating when plates closed. Opercular plates individually triangular, simple, without notable articular ridges or furrows, crests (apodemes) for depressor muscles or adductor muscle scars (
<figureCitation id="6C815B92FFEBFF8D562A5F1CFD98F84F" box="[541,623,1907,1931]" captionStart="FIGURE 13" captionStartId="14.[151,250,1865,1888]" captionTargetBox="[323,1234,441,1824]" captionTargetId="figure@14.[323,1264,409,1845]" captionTargetPageId="14" captionText="FIGURE 13. Atetrapachylasma dijonesae gen. et sp. nov., opercular plates of female: A and B, interior and exterior of terga, and C and D, interior and exterior of scuta. Both the terga and scuta are isosceles triangles, the bases of which are formed by their occludent margins. Note that the weakly developed articular ridges and furrows, as well as the lack of adductor and depressor muscle crests (apodemes), pits or scars, are remarkable. Comparably simple opercular plates are seen in no other balanomorph." httpUri="https://zenodo.org/record/202511/files/figure.png" pageId="11" pageNumber="60">Fig. 13</figureCitation>
).
</paragraph>
<paragraph id="F4054717FFEBFF8A54F25FF8FEA5FE43" blockId="11.[151,1436,1546,2003]" lastBlockId="12.[151,1437,151,391]" lastPageId="12" lastPageNumber="61" pageId="11" pageNumber="60">
Caudal appendages and penis absent. Labrum thick, crest nearly straight, supporting single row of fine, slightly separated, blunt teeth, flanked by stout mandibular palps (
<figureCitation id="6C815B92FFEBFF8D57175FD4FC87F817" box="[800,880,1979,2003]" captionStart="FIGURE 15" captionStartId="15.[151,250,1974,1997]" captionTargetBox="[189,1382,809,1944]" captionTargetId="figure@15.[185,1390,791,1951]" captionTargetPageId="15" captionText="FIGURE 15. Atetrapachylasma dijonesae gen. et sp. nov., appendages: A, mandible; B, first maxillae; C, second maxilla; D, intermediate articles of cirrus VI; and E, crest of labrum flanked by left palp." httpUri="https://zenodo.org/record/202513/files/figure.png" pageId="11" pageNumber="60">Fig. 15</figureCitation>
E); mandible tridentoid, clothed with fine spines, inferior angle armed with rounded comb of short strong spines. First maxillae with stepwise cutting edge armed with strong and relatively weak spines; second maxillae with typical notch (
<figureCitation id="6C815B92FFECFF8A57CB58D3FBB9FF10" box="[1020,1102,188,212]" captionStart="FIGURE 15" captionStartId="15.[151,250,1974,1997]" captionTargetBox="[189,1382,809,1944]" captionTargetId="figure@15.[185,1390,791,1951]" captionTargetPageId="15" captionText="FIGURE 15. Atetrapachylasma dijonesae gen. et sp. nov., appendages: A, mandible; B, first maxillae; C, second maxilla; D, intermediate articles of cirrus VI; and E, crest of labrum flanked by left palp." httpUri="https://zenodo.org/record/202513/files/figure.png" pageId="12" pageNumber="61">Fig. 15</figureCitation>
B, C). Cirri evenly deployed along thorax; first pair modified as maxillipeds, anterior ramus somewhat antenniform, posterior ramus nearly normal in resembling those of posterior pairs; second pair more closely resembling the following than the proceeding pair, rami generally subequal in length; intermediate articles of cirrus VI twice as high as wide, setation ctenopod, each article supporting one pair of strong and one pair of weak setae (
<figureCitation id="6C815B92FFECFF8A57995923FC08FEA0" box="[942,1023,332,356]" captionStart="FIGURE 15" captionStartId="15.[151,250,1974,1997]" captionTargetBox="[189,1382,809,1944]" captionTargetId="figure@15.[185,1390,791,1951]" captionTargetPageId="15" captionText="FIGURE 15. Atetrapachylasma dijonesae gen. et sp. nov., appendages: A, mandible; B, first maxillae; C, second maxilla; D, intermediate articles of cirrus VI; and E, crest of labrum flanked by left palp." httpUri="https://zenodo.org/record/202513/files/figure.png" pageId="12" pageNumber="61">Fig. 15</figureCitation>
D). Cirral counts for two specimens are given below:
</paragraph>
<paragraph id="F4054717FFECFF8A54A859CCFB89FD4C" pageId="12" pageNumber="61">
<table id="86BAB5B7FFEC007954A059CCFA6AFD4C" box="[151,1437,419,648]" gridcols="7" gridrows="5" pageId="12" pageNumber="61">
<tr id="4A8A4555FFEC007954A059CCFA6AFE7E" box="[151,1437,419,442]" gridrow="0" pageId="12" pageNumber="61">
<th id="095B2C29FFEC007954A059CCFE99FE7E" box="[151,366,419,442]" gridcol="0" gridrow="0" pageId="12" pageNumber="61">Cirrus I</th>
<th id="095B2C29FFEC007955DA59CCFDF0FE7E" box="[493,519,419,442]" gridcol="1" gridrow="0" pageId="12" pageNumber="61">II</th>
<th id="095B2C29FFEC007956BC59CCFD53FE7E" box="[651,676,419,442]" gridcol="2" gridrow="0" pageId="12" pageNumber="61">III</th>
<th id="095B2C29FFEC0079571E59CCFCB4FE7E" box="[809,835,419,442]" gridcol="3" gridrow="0" pageId="12" pageNumber="61">IV</th>
<th id="095B2C29FFEC007957F059CCFC16FE7E" box="[967,993,419,442]" gridcol="4" gridrow="0" pageId="12" pageNumber="61">V</th>
<th id="095B2C29FFEC0079505259CCFB89FE7E" box="[1125,1150,419,442]" gridcol="5" gridrow="0" pageId="12" pageNumber="61">V</th>
<th id="095B2C29FFEC0079513359CCFA6AFE7E" box="[1284,1437,419,442]" gridcol="6" gridrow="0" pageId="12" pageNumber="61">C.A.</th>
</tr>
<tr id="4A8A4555FFEC007954A059A2FA6AFE27" box="[151,1437,461,483]" gridrow="1" pageId="12" pageNumber="61">
<th id="095B2C29FFEC007954A059A2FE99FE27" box="[151,366,461,483]" gridcol="0" gridrow="1" pageId="12" pageNumber="61">anterior 10</th>
<td id="095B2C29FFEC007955DA59A2FDF0FE27" box="[493,519,461,483]" gridcol="1" gridrow="1" pageId="12" pageNumber="61">18</td>
<td id="095B2C29FFEC007956BC59A2FD53FE27" box="[651,676,461,483]" gridcol="2" gridrow="1" pageId="12" pageNumber="61">31</td>
<td id="095B2C29FFEC0079571E59A2FCB4FE27" box="[809,835,461,483]" gridcol="3" gridrow="1" pageId="12" pageNumber="61">36</td>
<td id="095B2C29FFEC007957F059A2FC16FE27" box="[967,993,461,483]" gridcol="4" gridrow="1" pageId="12" pageNumber="61">---</td>
<td id="095B2C29FFEC0079505259A2FB89FE27" box="[1125,1150,461,483]" gridcol="5" gridrow="1" pageId="12" pageNumber="61">44</td>
<td id="095B2C29FFEC0079513359A2FA6AFE27" box="[1284,1437,461,483]" gridcol="6" gridrow="1" pageId="12" pageNumber="61">absent</td>
</tr>
<tr id="4A8A4555FFEC007954A05999FA6AFDC8" box="[151,1437,502,524]" gridrow="2" pageId="12" pageNumber="61" rowspan-6="1">
<th id="095B2C29FFEC007954A05999FE99FDC8" box="[151,366,502,524]" gridcol="0" gridrow="2" pageId="12" pageNumber="61">posterior 9</th>
<td id="095B2C29FFEC007955DA5999FDF0FDC8" box="[493,519,502,524]" gridcol="1" gridrow="2" pageId="12" pageNumber="61">25</td>
<td id="095B2C29FFEC007956BC5999FD53FDC8" box="[651,676,502,524]" gridcol="2" gridrow="2" pageId="12" pageNumber="61">33</td>
<td id="095B2C29FFEC0079571E5999FCB4FDC8" box="[809,835,502,524]" gridcol="3" gridrow="2" pageId="12" pageNumber="61">45</td>
<td id="095B2C29FFEC007957F05999FC16FDC8" box="[967,993,502,524]" gridcol="4" gridrow="2" pageId="12" pageNumber="61">49</td>
<td id="095B2C29FFEC007950525999FB89FDC8" box="[1125,1150,502,524]" gridcol="5" gridrow="2" pageId="12" pageNumber="61">46</td>
</tr>
<tr id="4A8A4555FFEC007954A05A26FA6AFD9B" box="[151,1437,585,607]" gridrow="3" pageId="12" pageNumber="61">
<th id="095B2C29FFEC007954A05A26FE99FD9B" box="[151,366,585,607]" gridcol="0" gridrow="3" pageId="12" pageNumber="61">anterior 10</th>
<td id="095B2C29FFEC007955DA5A26FDF0FD9B" box="[493,519,585,607]" gridcol="1" gridrow="3" pageId="12" pageNumber="61">22</td>
<td id="095B2C29FFEC007956BC5A26FD53FD9B" box="[651,676,585,607]" gridcol="2" gridrow="3" pageId="12" pageNumber="61">30</td>
<td id="095B2C29FFEC0079571E5A26FCB4FD9B" box="[809,835,585,607]" gridcol="3" gridrow="3" pageId="12" pageNumber="61">36</td>
<td id="095B2C29FFEC007957F05A26FC16FD9B" box="[967,993,585,607]" gridcol="4" gridrow="3" pageId="12" pageNumber="61">45</td>
<td id="095B2C29FFEC007950525A26FB89FD9B" box="[1125,1150,585,607]" gridcol="5" gridrow="3" pageId="12" pageNumber="61">---</td>
<td id="095B2C29FFEC007951335A26FA6AFD9B" box="[1284,1437,585,607]" gridcol="6" gridrow="3" pageId="12" pageNumber="61">absent</td>
</tr>
<tr id="4A8A4555FFEC007954A05A1DFA6AFD4C" box="[151,1437,626,648]" gridrow="4" pageId="12" pageNumber="61" rowspan-6="1">
<th id="095B2C29FFEC007954A05A1DFE99FD4C" box="[151,366,626,648]" gridcol="0" gridrow="4" pageId="12" pageNumber="61">posterior 8</th>
<td id="095B2C29FFEC007955DA5A1DFDF0FD4C" box="[493,519,626,648]" gridcol="1" gridrow="4" pageId="12" pageNumber="61">16</td>
<td id="095B2C29FFEC007956BC5A1DFD53FD4C" box="[651,676,626,648]" gridcol="2" gridrow="4" pageId="12" pageNumber="61">33</td>
<td id="095B2C29FFEC0079571E5A1DFCB4FD4C" box="[809,835,626,648]" gridcol="3" gridrow="4" pageId="12" pageNumber="61">41</td>
<td id="095B2C29FFEC007957F05A1DFC16FD4C" box="[967,993,626,648]" gridcol="4" gridrow="4" pageId="12" pageNumber="61">45</td>
<td id="095B2C29FFEC007950525A1DFB89FD4C" box="[1125,1150,626,648]" gridcol="5" gridrow="4" pageId="12" pageNumber="61">---</td>
</tr>
</table>
</paragraph>
<caption id="A0C5179FFFECFF8A54A05C86FCA2FA9F" httpUri="https://zenodo.org/record/202509/files/figure.png" pageId="12" pageNumber="61" targetBox="[341,1239,759,1225]" targetPageId="12">
<paragraph id="F4054717FFECFF8A54A05C86FCA2FA9F" blockId="12.[151,1436,1257,1371]" pageId="12" pageNumber="61">
<emphasis id="C6CE9B05FFECFF8A54A05C86FED5FB3B" bold="true" box="[151,290,1257,1280]" pageId="12" pageNumber="61">FIGURE 11.</emphasis>
<taxonomicName id="33BA3C94FFECFF8A551B5C86FDB8FB3B" box="[300,591,1257,1279]" class="Maxillopoda" family="Pachylasmatidae" genus="Atetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="dijonesae">
<emphasis id="C6CE9B05FFECFF8A551B5C86FDB8FB3B" box="[300,591,1257,1279]" italics="true" pageId="12" pageNumber="61">Atetrapachylasma dijonesae</emphasis>
</taxonomicName>
<emphasis id="C6CE9B05FFECFF8A566D5C85FCF5FB3B" bold="true" box="[602,770,1258,1279]" pageId="12" pageNumber="61">gen. et sp. nov.</emphasis>
holotype: A, exterior view of female with males; note largely exposed alae and the pair of males on each side of the tight fitting occludent ridges of the scuta. B, interior view of the same female, note the tight fit of the bearing surfaces of the alae on the adjacent plates, and the close fit of the sutures between plates, especially between the opercular plates and the interior of the orifice.
</paragraph>
</caption>
<paragraph id="F4054717FFECFF8A54F05DE7FD93F9F4" blockId="12.[151,1436,1416,2027]" pageId="12" pageNumber="61">
Dwarf males located on external surface of S of female, as many as three per side (
<figureCitation id="6C815B92FFECFF8A50BF5DE7FB17FA64" box="[1160,1248,1416,1440]" captionStart="FIGURE 11" captionStartId="12.[151,250,1257,1280]" captionTargetBox="[341,1239,759,1225]" captionTargetId="figure@12.[317,1269,727,1235]" captionTargetPageId="12" captionText="FIGURE 11. Atetrapachylasma dijonesae gen. et sp. nov. holotype: A, exterior view of female with males; note largely exposed alae and the pair of males on each side of the tight fitting occludent ridges of the scuta. B, interior view of the same female, note the tight fit of the bearing surfaces of the alae on the adjacent plates, and the close fit of the sutures between plates, especially between the opercular plates and the interior of the orifice." httpUri="https://zenodo.org/record/202509/files/figure.png" pageId="12" pageNumber="61">Figs 11</figureCitation>
,
<figureCitation id="6C815B92FFECFF8A50D95DE7FAFAFA64" box="[1262,1293,1416,1440]" captionStart="FIGURE 14" captionStartId="15.[151,250,645,668]" captionTargetBox="[374,1237,208,614]" captionTargetId="figure@15.[339,1247,193,624]" captionTargetPageId="15" captionText="FIGURE 14. Atetrapachylasma dijonesae gen. et sp. nov., dwarf males: A, viewed obliquely (R-C axis running between 10: 30 and 4: 30 oclock); B, another individual, viewed from the carinal end, leaning markedly to its right, toward the aperture of the female (see Fig. 9 for four males in situ). Note that the chitinous primary wall is made up of four plates, with the rostrum overlapping the carinolaterals. There is no indication of sutures in the rostrum or the carinolaterals of the male or the female." httpUri="https://zenodo.org/record/202512/files/figure.png" pageId="12" pageNumber="61">14</figureCitation>
). Shell, R-C diameter approximately
<quantity id="3342EAF2FFECFF8A559C5DC3FE07FA00" box="[427,496,1452,1476]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="61" unit="mm" value="1.0">1 mm</quantity>
, consisting of three parts; 1) opercular plates (S and T) composed of chitinous primordial valves supplemented 2) by largely underlying calcareous plates, and 3) two concentric rings, the juvenile chitinous wall of four plates above concrescent, calcareous, columnar, four-plated wall, leaning to varying degrees toward aperture of female (
<figureCitation id="6C815B92FFECFF8A55FF5E77FDEFF9F4" box="[456,536,1560,1584]" captionStart="FIGURE 14" captionStartId="15.[151,250,645,668]" captionTargetBox="[374,1237,208,614]" captionTargetId="figure@15.[339,1247,193,624]" captionTargetPageId="15" captionText="FIGURE 14. Atetrapachylasma dijonesae gen. et sp. nov., dwarf males: A, viewed obliquely (R-C axis running between 10: 30 and 4: 30 oclock); B, another individual, viewed from the carinal end, leaning markedly to its right, toward the aperture of the female (see Fig. 9 for four males in situ). Note that the chitinous primary wall is made up of four plates, with the rostrum overlapping the carinolaterals. There is no indication of sutures in the rostrum or the carinolaterals of the male or the female." httpUri="https://zenodo.org/record/202512/files/figure.png" pageId="12" pageNumber="61">Fig. 14</figureCitation>
A, B).
</paragraph>
</subSubSection>
<subSubSection id="BCA0149CFFECFF9654F25E53FAF0FC98" lastPageId="16" lastPageNumber="65" pageId="12" pageNumber="61" type="etymology">
<paragraph id="F4054717FFECFF8A54F25E53FED9F9BC" blockId="12.[151,1436,1416,2027]" pageId="12" pageNumber="61">
<emphasis id="C6CE9B05FFECFF8A54F25E53FEBBF990" bold="true" box="[197,332,1596,1620]" pageId="12" pageNumber="61">Etymology.</emphasis>
Named for Diana S. Jones, in appreciation of her many works on thoracic
<taxonomicName id="33BA3C94FFECFF8A509C5E53FAD7F990" box="[1195,1312,1596,1620]" class="Maxillopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cirripedia" pageId="12" pageNumber="61" phylum="Arthropoda" rank="order">Cirripedia</taxonomicName>
, especially
<bibRefCitation id="902B3AE6FFECFF8A54A05E0EFEDDF9BC" author="Jones" box="[151,298,1632,1656]" pageId="12" pageNumber="61" refString="Jones, D. S. (2000) Crustacea Cirripedia Thoracica: Chionelasmatoidea and Pachylasmatoidea (Balanomorpha) of New Caledonia, Vanuatu and Wallis and Futuna Islands, with a review of all currently assigned taxa. In: Crosnier, A. (ed.), Resultats des Campagne MUSORSTOM 21. Memoires du Museum national d'Historie naturelle, 184, 141 - 283." type="journal article" year="2000">Jones (2000)</bibRefCitation>
.
</paragraph>
<paragraph id="F4054717FFECFF8A54F25EECFC19F82F" blockId="12.[151,1436,1416,2027]" pageId="12" pageNumber="61">
<emphasis id="C6CE9B05FFECFF8A54F25EECFECCF958" bold="true" box="[197,315,1667,1692]" pageId="12" pageNumber="61">Affinities.</emphasis>
The pachylasmatines constitute a diverse subfamily all members of which, excepting
<taxonomicName id="33BA3C94FFECFF8A513D5EEBFF2DF907" class="Maxillopoda" family="Pachylasmatidae" genus="Atetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="12" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFECFF8A513D5EEBFF2DF907" italics="true" pageId="12" pageNumber="61">Atetrapachylasma</emphasis>
</taxonomicName>
and arguably
<taxonomicName id="33BA3C94FFECFF8A554E5EC4FDCEF907" box="[377,569,1707,1731]" class="Maxillopoda" family="Pachylasmatidae" genus="Tetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="12" pageNumber="61" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="C6CE9B05FFECFF8A554E5EC4FDCEF907" box="[377,569,1707,1731]" italics="true" pageId="12" pageNumber="61">Tetrapachylasma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDFD267EFFECFF8A56775EC3FD93F907" box="[576,612,1708,1731]" pageId="12" pageNumber="61" sensu="stricto">s.s.</taxonomicNameLabel>
, display vestiges of an eight-plated ancestor; R-RL-CL1-CL2-C (cf.
<bibRefCitation id="902B3AE6FFECFF8A51695EC3FF23F923" author="Jones" pageId="12" pageNumber="61" refString="Jones, D. S. (2000) Crustacea Cirripedia Thoracica: Chionelasmatoidea and Pachylasmatoidea (Balanomorpha) of New Caledonia, Vanuatu and Wallis and Futuna Islands, with a review of all currently assigned taxa. In: Crosnier, A. (ed.), Resultats des Campagne MUSORSTOM 21. Memoires du Museum national d'Historie naturelle, 184, 141 - 283." type="journal article" year="2000">Jones 2000</bibRefCitation>
). Ignoring the exception, the number of wall plates of the 23 species representing five genera are variously reduced; reduction being ontogenetic as well as phylogenetic. Reduction is accomplished by partial to near total concrescence of the rostrolaterals with the rostrum, resulting in six-plated forms (RRL-CL1-CL2-C). Concrescences of the carinolaterals, in some cases simply by elimination of CL2 as suggested by
<bibRefCitation id="902B3AE6FFECFF8A50075F2EFB3DF89C" author="Foster" box="[1072,1226,1856,1880]" pageId="12" pageNumber="61" refString="Foster, B. A. (1988) Tetrapachylasma: A new genus for a shallow water pachylasmatid barnacle from the Cook Islands. Crustaceana, 55 (3), 225 - 233." type="journal article" year="1988">Foster (1988)</bibRefCitation>
, follows, reducing the number of plates to four. To put it another way, these configurations generally progress from visible vestigial sutures of a tripartite rostrum (RRL) to a compound rostrum (RRL), followed in close order by the carinolaterals (CL1/CL2 to CL1CL2). In such an ontogenetic series, last vestiges of the sutures are best seen on the interior, and to a lesser extent exteriorly, on the older, apical portions of the plates involved.
</paragraph>
<caption id="A0C5179FFFEDFF8B54A05D4FFCA8FA56" httpUri="https://zenodo.org/record/202510/files/figure.png" pageId="13" pageNumber="62" targetBox="[370,1206,224,1261]" targetPageId="13">
<paragraph id="F4054717FFEDFF8B54A05D4FFCA8FA56" blockId="13.[151,1436,1312,1426]" pageId="13" pageNumber="62">
<emphasis id="C6CE9B05FFEDFF8B54A05D4FFED7FAF2" bold="true" box="[151,288,1312,1335]" pageId="13" pageNumber="62">FIGURE 12.</emphasis>
<taxonomicName id="33BA3C94FFEDFF8B55105D4EFDBFFAF3" box="[295,584,1313,1335]" class="Maxillopoda" family="Pachylasmatidae" genus="Atetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="dijonesae">
<emphasis id="C6CE9B05FFEDFF8B55105D4EFDBFFAF3" box="[295,584,1313,1335]" italics="true" pageId="13" pageNumber="62">Atetrapachylasma dijonesae</emphasis>
</taxonomicName>
<emphasis id="C6CE9B05FFEDFF8B56785D4EFD19FAF2" bold="true" box="[591,750,1313,1334]" pageId="13" pageNumber="62">gen. et sp. nov.</emphasis>
, exploded four-plated wall arranged from loose plates, type lot: A, exterior of a rostrum; B and D, left and right carinolaterals; C, a carina; and E, the interior of a portion of a rostrum. Note the variably pronounced depression in the sheath of rostrum (E), which, together with the edges of the carinolateral alae, receives the rostral ends of the occludent ridges of the scuta (Figs 9A; 11C, D).
</paragraph>
</caption>
<paragraph id="F4054717FFEDFF8B54F25DD5FD29F915" blockId="13.[151,1436,1466,2033]" pageId="13" pageNumber="62">
Five four-plated species were assigned to
<taxonomicName id="33BA3C94FFEDFF8B56A15DD5FC10FA16" authority="Foster, 1988" authorityName="Foster" authorityYear="1988" box="[662,999,1466,1490]" class="Maxillopoda" family="Pachylasmatidae" genus="Tetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="13" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFEDFF8B56A15DD5FCA1FA16" box="[662,854,1466,1490]" italics="true" pageId="13" pageNumber="62">Tetrapachylasma</emphasis>
<bibRefCitation id="902B3AE6FFEDFF8B576B5DD4FC10FA16" author="Foster" box="[860,999,1466,1490]" pageId="13" pageNumber="62" refString="Foster, B. A. (1988) Tetrapachylasma: A new genus for a shallow water pachylasmatid barnacle from the Cook Islands. Crustaceana, 55 (3), 225 - 233." type="journal article" year="1988">Foster, 1988</bibRefCitation>
</taxonomicName>
by
<bibRefCitation id="902B3AE6FFEDFF8B50275DD4FB56FA16" author="Jones" box="[1040,1185,1466,1490]" pageId="13" pageNumber="62" refString="Jones, D. S. (2000) Crustacea Cirripedia Thoracica: Chionelasmatoidea and Pachylasmatoidea (Balanomorpha) of New Caledonia, Vanuatu and Wallis and Futuna Islands, with a review of all currently assigned taxa. In: Crosnier, A. (ed.), Resultats des Campagne MUSORSTOM 21. Memoires du Museum national d'Historie naturelle, 184, 141 - 283." type="journal article" year="2000">Jones (2000)</bibRefCitation>
, but of them only in
<taxonomicName id="33BA3C94FFEDFF8B51B05DD4FE79FA3D" authority="Foster, 1988" authorityName="Foster" authorityYear="1988" class="Maxillopoda" family="Pachylasmatidae" genus="Tetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="trigonum">
<emphasis id="C6CE9B05FFEDFF8B51B05DD4FF09FA3D" italics="true" pageId="13" pageNumber="62">T. trigonum</emphasis>
<bibRefCitation id="902B3AE6FFEDFF8B55335D8DFE79FA3D" author="Foster" box="[260,398,1505,1529]" pageId="13" pageNumber="62" refString="Foster, B. A. (1988) Tetrapachylasma: A new genus for a shallow water pachylasmatid barnacle from the Cook Islands. Crustaceana, 55 (3), 225 - 233." type="journal article" year="1988">Foster, 1988</bibRefCitation>
</taxonomicName>
are the compound plates, RLR and CL1CL2, believed, at least by Foster, to be passed through during ontogeny without traces of the fused sutures. However,
<taxonomicName id="33BA3C94FFEDFF8B57685E69FC14F9D9" box="[863,995,1542,1565]" class="Maxillopoda" family="Pachylasmatidae" genus="Tetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="trigonum">
<emphasis id="C6CE9B05FFEDFF8B57685E69FC14F9D9" box="[863,995,1542,1565]" italics="true" pageId="13" pageNumber="62">T. trigonum</emphasis>
</taxonomicName>
is known from but three mature specimens, measuring between 8 and
<quantity id="3342EAF2FFEDFF8B563E5E46FDAEF985" box="[521,601,1577,1601]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="62" unit="mm" value="10.0">10 mm</quantity>
in R-C, diameter.
<bibRefCitation id="902B3AE6FFEDFF8B571C5E45FC4AF985" author="Jones" box="[811,957,1577,1601]" pageId="13" pageNumber="62" refString="Jones, D. S. (2000) Crustacea Cirripedia Thoracica: Chionelasmatoidea and Pachylasmatoidea (Balanomorpha) of New Caledonia, Vanuatu and Wallis and Futuna Islands, with a review of all currently assigned taxa. In: Crosnier, A. (ed.), Resultats des Campagne MUSORSTOM 21. Memoires du Museum national d'Historie naturelle, 184, 141 - 283." type="journal article" year="2000">Jones (2000)</bibRefCitation>
argues that, since traces of the sutures are seen in the ontogeny of the other four species and Foster did not have an ontogenetic series to judge by, one cannot be certain they are not also passed through in the ontogeny of
<taxonomicName id="33BA3C94FFEDFF8B57565E1DFC13F94D" box="[865,996,1650,1673]" class="Maxillopoda" family="Pachylasmatidae" genus="Tetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="trigonum">
<emphasis id="C6CE9B05FFEDFF8B57565E1DFC13F94D" box="[865,996,1650,1673]" italics="true" pageId="13" pageNumber="62">T. trigonum</emphasis>
</taxonomicName>
. The very small males of
<taxonomicName id="33BA3C94FFEDFF8B51265E1DFA62F94D" box="[1297,1429,1650,1673]" class="Maxillopoda" family="Pachylasmatidae" genus="Tetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="trigonum">
<emphasis id="C6CE9B05FFEDFF8B51265E1DFA62F94D" box="[1297,1429,1650,1673]" italics="true" pageId="13" pageNumber="62">T. trigonum</emphasis>
</taxonomicName>
, measuring half a millimeter or so in R-C diameter, while perhaps four-plated, do not calcify in the carinolateral region, and so one cannot be sure about them either.
</paragraph>
<paragraph id="F4054717FFEDFF8854F25EB2FD39FEA0" blockId="13.[151,1436,1466,2033]" lastBlockId="14.[151,1436,151,356]" lastPageId="14" lastPageNumber="63" pageId="13" pageNumber="62">
On the other hand, the sample of the new species consists of numerous loose shells and a half dozen or so complete specimens having R-C diameters ranging from ~
<quantity id="3342EAF2FFEDFF8B56C65F6EFC84F8DD" box="[753,883,1793,1817]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="15.0" metricValueMin="4.0" pageId="13" pageNumber="62" unit="mm" value="9.5" valueMax="15.0" valueMin="4.0">4 to 15 mm</quantity>
, and no traces of vestigial sutures are seen among them. Furthermore, the males, a millimeter or so in R-C diameter, are clearly without vestigial sutures in the early uncalcified as well as the later calcified parts of the wall and, therefore, they too appear to be but four-plated. However, while this indicates that
<taxonomicName id="33BA3C94FFEDFF8B55D15F01FD9EF841" box="[486,617,1902,1925]" class="Maxillopoda" family="Pachylasmatidae" genus="Tetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="trigonum">
<emphasis id="C6CE9B05FFEDFF8B55D15F01FD9EF841" box="[486,617,1902,1925]" italics="true" pageId="13" pageNumber="62">T. trigonum</emphasis>
</taxonomicName>
could also have been four-plated at the onset, so different are its opercular plates that it does not seem likely it is allied to the new species. Therefore, the genus may not be a monophyletic taxon. This is heightened by the fact that the same four-plated configuration arose independently in the 1) bathylasmatines (
<taxonomicName id="33BA3C94FFEDFF8B54C85FB6FE46F835" box="[255,433,2009,2033]" class="Maxillopoda" family="Bathylasmatidae" genus="Tetrachaelasma" kingdom="Animalia" order="Sessilia" pageId="13" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFEDFF8B54C85FB6FE46F835" box="[255,433,2009,2033]" italics="true" pageId="13" pageNumber="62">Tetrachaelasma</emphasis>
</taxonomicName>
), 2) chthamalines (
<taxonomicName id="33BA3C94FFEDFF8B56B95FB6FCBEF835" box="[654,841,2009,2033]" class="Maxillopoda" family="Chthamalidae" genus="Tetrachthamalus" kingdom="Animalia" order="Sessilia" pageId="13" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFEDFF8B56B95FB6FCBEF835" box="[654,841,2009,2033]" italics="true" pageId="13" pageNumber="62">Tetrachthamalus</emphasis>
</taxonomicName>
), 3) elminiines (
<taxonomicName id="33BA3C94FFEDFF8B503F5FB6FAF3F835" authority="Austrominiu" authorityName="Austrominiu" box="[1032,1284,2009,2033]" class="Maxillopoda" family="Balanidae" genus="Elminius" kingdom="Animalia" order="Sessilia" pageId="13" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFEDFF8B503F5FB6FB9BF835" box="[1032,1132,2009,2033]" italics="true" pageId="13" pageNumber="62">Elminius</emphasis>
,
<emphasis id="C6CE9B05FFEDFF8B504C5FB6FAF3F835" box="[1147,1284,2009,2033]" italics="true" pageId="13" pageNumber="62">Austrominiu</emphasis>
</taxonomicName>
s, etc. (
<bibRefCitation id="902B3AE6FFEDFF8851615FB6FE38FF6B" author="Buckeridge" lastPageId="14" lastPageNumber="63" pageId="13" pageNumber="62" refString="Buckeridge, J. S. &amp; Newman, W. A. (2010) A review of the subfamily Elminiinae (Cirripedia: Thoracica: Austrobalanidae), including a new genus, Protelminius nov., from the Oligocene of New Zealand. Zootaxa, 2349, 39 - 54." type="journal article" year="2010">Buckeridge and Newman 2010</bibRefCitation>
), 4) most
<taxonomicName id="33BA3C94FFEEFF88566558F8FD2AFF6B" box="[594,733,151,175]" class="Maxillopoda" family="Tetraclitidae" kingdom="Animalia" order="Sessilia" pageId="14" pageNumber="63" phylum="Arthropoda" rank="family">Tetraclitidae</taxonomicName>
(
<emphasis id="C6CE9B05FFEEFF8856C358F7FED4FF10" italics="true" pageId="14" pageNumber="63">
<taxonomicName id="33BA3C94FFEEFF8856C358F7FCABFF74" box="[756,860,152,176]" class="Maxillopoda" family="Tetraclitidae" genus="Epopella" kingdom="Animalia" order="Sessilia" pageId="14" pageNumber="63" phylum="Arthropoda" rank="genus">Epopella</taxonomicName>
,
<taxonomicName id="33BA3C94FFEEFF88575958F6FC0EFF74" box="[878,1017,153,176]" class="Maxillopoda" family="Tetraclitidae" genus="Tesseropora" kingdom="Animalia" order="Sessilia" pageId="14" pageNumber="63" phylum="Arthropoda" rank="genus">Tesseropora</taxonomicName>
,
<taxonomicName id="33BA3C94FFEEFF88503D58F7FB62FF74" box="[1034,1173,152,176]" class="Maxillopoda" family="Tetraclitidae" genus="Tetraclitella" kingdom="Animalia" order="Sessilia" pageId="14" pageNumber="63" phylum="Arthropoda" rank="genus">Tetraclitella</taxonomicName>
,
<taxonomicName id="33BA3C94FFEEFF88509058F7FAE1FF74" box="[1191,1302,152,176]" class="Maxillopoda" family="Tetraclitidae" genus="Tetraclita" kingdom="Animalia" order="Sessilia" pageId="14" pageNumber="63" phylum="Arthropoda" rank="genus">Tetraclita</taxonomicName>
,
<taxonomicName id="33BA3C94FFEEFF88511058F7FA60FF74" box="[1319,1431,152,176]" class="Maxillopoda" family="Tetraclitidae" genus="Astroclita" kingdom="Animalia" order="Sessilia" pageId="14" pageNumber="63" phylum="Arthropoda" rank="genus">Astroclita</taxonomicName>
,
<taxonomicName id="33BA3C94FFEEFF8854A058D3FED4FF10" box="[151,291,188,212]" class="Maxillopoda" family="Tetraclitidae" genus="Newmanella" kingdom="Animalia" order="Sessilia" pageId="14" pageNumber="63" phylum="Arthropoda" rank="genus">Newmanella</taxonomicName>
</emphasis>
and
<taxonomicName id="33BA3C94FFEEFF88556B58D3FDF6FF10" box="[348,513,188,212]" class="Maxillopoda" family="Tetraclitidae" genus="Yamaguchiella" kingdom="Animalia" order="Sessilia" pageId="14" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFEEFF88556B58D3FDF6FF10" box="[348,513,188,212]" italics="true" pageId="14" pageNumber="63">Yamaguchiella</emphasis>
</taxonomicName>
), balanoids, including independently the 5) balanids
<taxonomicName id="33BA3C94FFEEFF88506858D3FB05FF10" box="[1119,1266,188,212]" class="Maxillopoda" family="Balanidae" genus="Tetrabalanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="14" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFEEFF88506858D3FB05FF10" box="[1119,1266,188,212]" italics="true" pageId="14" pageNumber="63">Tetrabalanus</emphasis>
</taxonomicName>
and
<emphasis id="C6CE9B05FFEEFF88511B58D3FA8BFF10" box="[1324,1404,188,212]" italics="true" pageId="14" pageNumber="63">
<taxonomicName id="33BA3C94FFEEFF88511B58D3FA8FFF10" box="[1324,1400,188,212]" class="Maxillopoda" family="Balanidae" genus="Zulloa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="14" pageNumber="63" phylum="Arthropoda" rank="genus">Zulloa</taxonomicName>
,
</emphasis>
6) pyrgomatids or coral barnacles (cf.
<bibRefCitation id="902B3AE6FFEEFF88561B58B0FD34FF33" author="Darwin" box="[556,707,223,247]" pageId="14" pageNumber="63" refString="Darwin, C. (1854) A monograph on the subclass Cirripedia, with figures of all species, the Balanidae, the Verrucidae etc. Pp 1 - 684 + pls 1 - 30. Ray Society, London." type="book chapter" year="1854">Darwin 1854</bibRefCitation>
,
<bibRefCitation id="902B3AE6FFEEFF8856E7588FFC8EFF33" author="Newman" box="[720,889,223,247]" pageId="14" pageNumber="63" refString="Newman, W. A. (1967) A new genus of Chthamalidae (Cirripedia, Balanomorpha) from the Red Sea and Indian Ocean. Journal of Zoology, London, 153, 423 - 435." type="journal article" year="1967">Newman 1967</bibRefCitation>
,
<bibRefCitation id="902B3AE6FFEEFF8857B2588FFB68FF33" author="Ross" box="[901,1183,223,247]" pageId="14" pageNumber="63" refString="Ross, A. &amp; Newman, W. A. (1996) A unique experiment in four-platedness by a Miocene barnacle (Cirripedia: Balanidae) that Darwin considered improbable. Journal of Crustacean Biology, 16 (4), 663 - 668." type="journal article" year="1996">Ross and Newman 1996</bibRefCitation>
), and the 7) bryozobiines (
<bibRefCitation id="902B3AE6FFEEFF8854E4596BFDD3FED8" author="Van" box="[211,548,260,284]" pageId="14" pageNumber="63" refString="Van Syoc, R. J. &amp; Newman, W. A. (2010) Morphology and evolutionary ecology of a sponge-barnacle symbiosis: Four new genera of barnacles (Archaeobalanidae, Bryozobiinae). Journal of Experimental Marine Biology and Ecology, 392, 65 - 88." type="journal article" year="2010">Van Syoc and Newman 2010</bibRefCitation>
). There are other four-plated configurations, such as R-RLCL-C in
<taxonomicName id="33BA3C94FFEEFF885114596BFF36FE84" class="Maxillopoda" family="Chthamalidae" genus="Chamaesipho" kingdom="Animalia" order="Sessilia" pageId="14" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFEEFF885114596BFF36FE84" italics="true" pageId="14" pageNumber="63">Chamaesipho</emphasis>
</taxonomicName>
; a configuration arrived at by yet another route. Thus, monophyly of at least
<taxonomicName id="33BA3C94FFEEFF8850555947FAD5FE84" box="[1122,1314,296,320]" class="Maxillopoda" family="Pachylasmatidae" genus="Tetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="14" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFEEFF8850555947FAD5FE84" box="[1122,1314,296,320]" italics="true" pageId="14" pageNumber="63">Tetrapachylasma</emphasis>
</taxonomicName>
if not the pachylasmatines in general is anything but certain.
</paragraph>
<caption id="A0C5179FFFEEFF8854A05F26FEF2F81E" httpUri="https://zenodo.org/record/202511/files/figure.png" pageId="14" pageNumber="63" targetBox="[323,1234,441,1824]" targetPageId="14">
<paragraph id="F4054717FFEEFF8854A05F26FEF2F81E" blockId="14.[151,1436,1865,2010]" pageId="14" pageNumber="63">
<emphasis id="C6CE9B05FFEEFF8854A05F26FED7F89B" bold="true" box="[151,288,1865,1888]" pageId="14" pageNumber="63">FIGURE 13.</emphasis>
<taxonomicName id="33BA3C94FFEEFF8855105F25FDB0F8A4" box="[295,583,1866,1888]" class="Maxillopoda" family="Pachylasmatidae" genus="Atetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="14" pageNumber="63" phylum="Arthropoda" rank="species" species="dijonesae">
<emphasis id="C6CE9B05FFEEFF8855105F25FDB0F8A4" box="[295,583,1866,1888]" italics="true" pageId="14" pageNumber="63">Atetrapachylasma dijonesae</emphasis>
</taxonomicName>
<emphasis id="C6CE9B05FFEEFF88567A5F25FD04F89B" bold="true" box="[589,755,1866,1887]" pageId="14" pageNumber="63">gen. et sp. nov.,</emphasis>
opercular plates of female: A and B, interior and exterior of terga, and C and D, interior and exterior of scuta. Both the terga and scuta are isosceles triangles, the bases of which are formed by their occludent margins. Note that the weakly developed articular ridges and furrows, as well as the lack of adductor and depressor muscle crests (apodemes), pits or scars, are remarkable. Comparably simple opercular plates are seen in no other balanomorph.
</paragraph>
</caption>
<caption id="A0C5179FFFEFFF8954A05AEAFAADFD3C" httpUri="https://zenodo.org/record/202512/files/figure.png" pageId="15" pageNumber="64" targetBox="[374,1237,208,614]" targetPageId="15">
<paragraph id="F4054717FFEFFF8954A05AEAFAADFD3C" blockId="15.[151,1436,645,760]" pageId="15" pageNumber="64">
<emphasis id="C6CE9B05FFEFFF8954A05AEAFEE8FD5F" bold="true" box="[151,287,645,668]" pageId="15" pageNumber="64">FIGURE 14.</emphasis>
<taxonomicName id="33BA3C94FFEFFF8955135AE9FDB3FD58" box="[292,580,646,668]" class="Maxillopoda" family="Pachylasmatidae" genus="Atetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="15" pageNumber="64" phylum="Arthropoda" rank="species" species="dijonesae">
<emphasis id="C6CE9B05FFEFFF8955135AE9FDB3FD58" box="[292,580,646,668]" italics="true" pageId="15" pageNumber="64">Atetrapachylasma dijonesae</emphasis>
</taxonomicName>
<emphasis id="C6CE9B05FFEFFF89567E5AE9FD12FD5F" bold="true" box="[585,741,646,667]" pageId="15" pageNumber="64">gen. et sp. nov.</emphasis>
, dwarf males: A, viewed obliquely (R-C axis running between 10:30 and 4:30 oclock); B, another individual, viewed from the carinal end, leaning markedly to its right, toward the aperture of the female (see Fig. 9 for four males
<emphasis id="C6CE9B05FFEFFF8955DE5AABFDD0FD1D" box="[489,551,708,729]" italics="true" pageId="15" pageNumber="64">in situ</emphasis>
). Note that the chitinous primary wall is made up of four plates, with the rostrum overlapping the carinolaterals. There is no indication of sutures in the rostrum or the carinolaterals of the male or the female.
</paragraph>
</caption>
<caption id="A0C5179FFFEFFF8954A05FD9FC55F82F" httpUri="https://zenodo.org/record/202513/files/figure.png" pageId="15" pageNumber="64" targetBox="[189,1382,809,1944]" targetPageId="15">
<paragraph id="F4054717FFEFFF8954A05FD9FC55F82F" blockId="15.[151,1436,1974,2027]" pageId="15" pageNumber="64">
<emphasis id="C6CE9B05FFEFFF8954A05FD9FED7F808" bold="true" box="[151,288,1974,1997]" pageId="15" pageNumber="64">FIGURE 15.</emphasis>
<taxonomicName id="33BA3C94FFEFFF8955105FD8FDB0F809" box="[295,583,1975,1997]" class="Maxillopoda" family="Pachylasmatidae" genus="Atetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="15" pageNumber="64" phylum="Arthropoda" rank="species" species="dijonesae">
<emphasis id="C6CE9B05FFEFFF8955105FD8FDB0F809" box="[295,583,1975,1997]" italics="true" pageId="15" pageNumber="64">Atetrapachylasma dijonesae</emphasis>
</taxonomicName>
<emphasis id="C6CE9B05FFEFFF8956795FD8FD1AF808" bold="true" box="[590,749,1975,1996]" pageId="15" pageNumber="64">gen. et sp. nov.</emphasis>
, appendages: A, mandible; B, first maxillae; C, second maxilla; D, intermediate articles of cirrus VI; and E, crest of labrum flanked by left palp.
</paragraph>
</caption>
<paragraph id="F4054717FFF0FF9654F258F7FAF0FC98" blockId="16.[151,1437,151,860]" pageId="16" pageNumber="65">
Four of the five species of
<taxonomicName id="33BA3C94FFF0FF9655CC58F7FD4CFF74" box="[507,699,152,176]" class="Maxillopoda" family="Pachylasmatidae" genus="Tetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="16" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFF0FF9655CC58F7FD4CFF74" box="[507,699,152,176]" italics="true" pageId="16" pageNumber="65">Tetrapachylasma</emphasis>
</taxonomicName>
are known to have a calcareous basis, albeit it very thin, as it is centrally in at least two, and
<bibRefCitation id="902B3AE6FFF0FF9655E158D2FD91FF10" author="Jones" box="[470,614,188,212]" pageId="16" pageNumber="65" refString="Jones, D. S. (2000) Crustacea Cirripedia Thoracica: Chionelasmatoidea and Pachylasmatoidea (Balanomorpha) of New Caledonia, Vanuatu and Wallis and Futuna Islands, with a review of all currently assigned taxa. In: Crosnier, A. (ed.), Resultats des Campagne MUSORSTOM 21. Memoires du Museum national d'Historie naturelle, 184, 141 - 283." type="journal article" year="2000">Jones (2000)</bibRefCitation>
infers that the fifth has one too. Unfortunately, its nature (membranous or calcareous) in
<taxonomicName id="33BA3C94FFF0FF96550B588FFD8CFF3C" box="[316,635,224,248]" class="Maxillopoda" family="Pachylasmatidae" genus="Atetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="16" pageNumber="65" phylum="Arthropoda" rank="species" species="dijonesae" status="sp. nov.">
<emphasis id="C6CE9B05FFF0FF96550B588FFD8CFF3C" box="[316,635,224,248]" italics="true" pageId="16" pageNumber="65">Atetrapachylasma dijonesae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDFD267EFFF0FF9656B3588FFD29FF3C" box="[644,734,224,248]" pageId="16" pageNumber="65" rank="species">
<emphasis id="C6CE9B05FFF0FF9656B3588FFD29FF3C" bold="true" box="[644,734,224,248]" pageId="16" pageNumber="65">sp. nov.</emphasis>
</taxonomicNameLabel>
could not be directly ascertained, but the videos taken during capture (cf.
<figureCitation id="6C815B92FFF0FF965528596BFE96FED8" box="[287,353,260,284]" captionStart="FIGURE 5" captionStartId="6.[151,250,1830,1853]" captionTargetBox="[295,1289,199,1795]" captionTargetId="figure@6.[284,1301,193,1810]" captionTargetPageId="6" captionText="FIGURE 5. Atetrapachylasma dijonesae gen. et sp. nov. in situ on pillow basalt from 2080 m of depth Vance B Seamount, with nozzle of suction sampler of ROV Tiburon largely to their right. A, the remains of several walls broken away by the nozzle can be seen to the right of “ 1 ” leaving membranous bases exposed. B, the barnacles seen between “ 2 ” and “ 3 ” and to the right of “ 4 ” in A have been collected. N. B.: Inspection of MPEGs taken by ROV Tiburon during collection of the barnacles seen removed in these frames detected no loss of material in the process. Therefore, it can be said with confidence that the shell fragments at the base of the basalt below the barnacles (cf. Fig. 4) represent natural postmortem accumulations." httpUri="https://zenodo.org/record/202506/files/figure.png" pageId="16" pageNumber="65">Fig. 5</figureCitation>
) show that, in the unlikely event one is present, it is very thin. Caudal appendages in the genus range from as many as 18 segments in
<taxonomicName id="33BA3C94FFF0FF96567E5946FD39FE84" box="[585,718,297,320]" class="Maxillopoda" family="Pachylasmatidae" genus="Tetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="16" pageNumber="65" phylum="Arthropoda" rank="species" species="arcuatum">
<emphasis id="C6CE9B05FFF0FF96567E5946FD39FE84" box="[585,718,297,320]" italics="true" pageId="16" pageNumber="65">T. arcuatum</emphasis>
</taxonomicName>
to as few as one in
<taxonomicName id="33BA3C94FFF0FF96579C5946FBDBFE84" box="[939,1068,297,320]" class="Maxillopoda" family="Pachylasmatidae" genus="Tetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="16" pageNumber="65" phylum="Arthropoda" rank="species" species="trigonum">
<emphasis id="C6CE9B05FFF0FF96579C5946FBDBFE84" box="[939,1068,297,320]" italics="true" pageId="16" pageNumber="65">T. trigonum</emphasis>
</taxonomicName>
, but none were found in the new form. The opercular plates of the previously described species of
<taxonomicName id="33BA3C94FFF0FF9657405923FBC0FEA0" box="[887,1079,332,356]" class="Maxillopoda" family="Pachylasmatidae" genus="Tetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="16" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFF0FF9657405923FBC0FEA0" box="[887,1079,332,356]" italics="true" pageId="16" pageNumber="65">Tetrapachylasma</emphasis>
</taxonomicName>
, while relatively generalized as balanomorphs go, have longitudinally striate scuta (a diagnostic feature of the genus), and terga with an incipient spur with an open furrow and strong depressor muscle crests. The operculum of the new form is macroscopically undistinguished, other than for the strong external and internal occludent ridge of the scuta. Furthermore, the tergum resembles that of the brachylepadomorph,
<taxonomicName id="33BA3C94FFF0FF96568459B3FAE9FE30" authority="Newman" authorityName="Newman" authorityYear="1995" box="[691,1310,476,500]" class="Maxillopoda" family="Brachylepadidae" genus="Neobrachylepas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="16" pageNumber="65" phylum="Arthropoda" rank="species" species="relica">
<emphasis id="C6CE9B05FFF0FF96568459B3FC47FE30" box="[691,944,476,500]" italics="true" pageId="16" pageNumber="65">Neobrachylepas relica</emphasis>
<bibRefCitation id="902B3AE6FFF0FF96578E59B2FAE9FE30" author="Newman" box="[953,1310,476,500]" pageId="16" pageNumber="65" refString="Newman, W. A. &amp; Yamaguchi, T. (1995) A new sessile barnacle (Cirripedia; Brachylepadomorpha) from the Lau Back-Arc Basin, Tonga; first record of a living representative since the Miocene. Bulletin du Museum national d'Histoire naturelle, Paris (4), 17 A (3 - 4), 211 - 243." type="journal article" year="1995">Newman and Yamaguchi, 1995</bibRefCitation>
</taxonomicName>
, in its simplicity; that is, in having a straight basal margin (no indication of a spur, incipient or otherwise) and no tergal depressor muscle crests (apodemes), which is also the case in the otherwise more generalized balanomorphs,
<taxonomicName id="33BA3C94FFF0FF96516B5A4BFF08FDA4" class="Maxillopoda" family="Chionelasmatidae" genus="Chionelasmus" kingdom="Animalia" order="Sessilia" pageId="16" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFF0FF96516B5A4BFF08FDA4" italics="true" pageId="16" pageNumber="65">Chionelasmus</emphasis>
</taxonomicName>
and
<taxonomicName id="33BA3C94FFF0FF96550F5A27FE4AFDA4" box="[312,445,584,608]" class="Maxillopoda" family="Waikalasmatidae" genus="Waikalasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="16" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFF0FF96550F5A27FE4AFDA4" box="[312,445,584,608]" italics="true" pageId="16" pageNumber="65">Waikalasma</emphasis>
</taxonomicName>
. While the scuta are not crossed by the external longitudinal striae seen in
<taxonomicName id="33BA3C94FFF0FF9651225A27FF2DFD40" class="Maxillopoda" family="Pachylasmatidae" genus="Tetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="16" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFF0FF9651225A27FF2DFD40" italics="true" pageId="16" pageNumber="65">Tetrapachylasma</emphasis>
</taxonomicName>
in general, the growth lines alternate to form fine interdigitating teeth along their occludent margins and the external occludent ridge appears to be a unique apomorphy. The simple growth lines of the terga also interdigitate along their occludent margins and this may be a unique apomorphy. Otherwise, the simplicity of the opercular parts of the new form, especially the terga, suggest they have either retrogressed from those of the other species of
<taxonomicName id="33BA3C94FFF0FF96516F5AB6FEEAFCD0" class="Maxillopoda" family="Pachylasmatidae" genus="Tetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="16" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFF0FF96516F5AB6FEEAFCD0" italics="true" pageId="16" pageNumber="65">Tetrapachylasma</emphasis>
</taxonomicName>
, its extant allies, or from some extinct pachylasmatine ancestor, or they are simply the least modified of any known pachylasmatoid, or of any balanomorphs for that matter. Whichever, taken a face value,
<taxonomicName id="33BA3C94FFF0FF96513D5B4FFF2DFC98" class="Maxillopoda" family="Pachylasmatidae" genus="Atetrapachylasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="16" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="C6CE9B05FFF0FF96513D5B4FFF2DFC98" italics="true" pageId="16" pageNumber="65">Atetrapachylasma</emphasis>
</taxonomicName>
is a unique barnacle, and it is recommended it be placed
<emphasis id="C6CE9B05FFF0FF96576C5B2AFC01FC98" box="[859,1014,836,860]" italics="true" pageId="16" pageNumber="65">incertae sedis</emphasis>
in the
<taxonomicName id="33BA3C94FFF0FF9650745B2BFAF4FC98" box="[1091,1283,836,860]" class="Maxillopoda" family="Pachylasmatidae" kingdom="Animalia" order="Sessilia" pageId="16" pageNumber="65" phylum="Arthropoda" rank="subFamily" subFamily="Pachylasmatinae">Pachylasmatinae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>