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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.905.39592" ID-GBIF-Dataset="ee5a0817-7cfc-4c21-ba7d-3beabb06cccd" ID-PMC="PMC7004242" ID-Pensoft-Pub="1313-2970-905-1" ID-Pensoft-UUID="4C7E0CB428DF56BDB15B7AC8D6707961" ID-PubMed="32063726" ID-ZooBank="F4C766E4633A41039FFDE952718F41FB" ModsDocID="1313-2970-905-1" checkinTime="1579539902851" checkinUser="pensoft" docAuthor="Salata, Sebastian &amp; Fisher, Brian L." docDate="2020" docId="DBE9B8793CDD200FA2CC7A5A8A3EAAD1" docLanguage="en" docName="ZooKeys 905: 1-235" docOrigin="ZooKeys 905" docSource="http://dx.doi.org/10.3897/zookeys.905.39592" docTitle="Pheidole bemarahaensis Salata &amp; Fisher 2020, sp. nov." docType="treatment" docUuid="C491FD00-0251-4F41-957D-8CDAEA87616E" docUuidSource="ZooBank" docVersion="5" id="4C7E0CB428DF56BDB15B7AC8D6707961" lastPageNumber="1" masterDocId="4C7E0CB428DF56BDB15B7AC8D6707961" masterDocTitle="Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups" masterLastPageNumber="235" masterPageNumber="1" pageNumber="1" updateTime="1668168337901" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Salata, Sebastian</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fisher, Brian L.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>905</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>235</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.905.39592</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.905.39592</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-905-1</mods:identifier>
<mods:identifier type="ZooBank">F4C766E4633A41039FFDE952718F41FB</mods:identifier>
<mods:identifier type="Pensoft-UUID">4C7E0CB428DF56BDB15B7AC8D6707961</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="161777033" LSID="urn:lsid:zoobank.org:act:C491FD00-0251-4F41-957D-8CDAEA87616E" httpUri="http://treatment.plazi.org/id/DBE9B8793CDD200FA2CC7A5A8A3EAAD1" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/C491FD00-0251-4F41-957D-8CDAEA87616E" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole bemarahaensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bemarahaensis" status="sp. nov.">Pheidole bemarahaensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole bemarahaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0425239) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0425236)." figureDoi="10.3897/zookeys.905.39592.figure61" httpUri="https://binary.pensoft.net/fig/373108" pageId="0" pageNumber="1">Figs 61A-F</figureCitation>
<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">, 84G</figureCitation>
<figureCitation captionStart="Figure 86" captionStartId="F86" captionText="Figure 86. Distribution. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure86" httpUri="https://binary.pensoft.net/fig/373133" pageId="0" pageNumber="1">, 86G</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
.
</emphasis>
Madagascar. •1 major worker; Mahajanga; Parc National Tsingy de Bemaraha, 2.5 km 62°ENE Bekopaka, Ankidrodroa River;
<geoCoordinate degrees="19.13222" direction="south" orientation="latitude" precision="1" value="-19.13222">-19.13222</geoCoordinate>
,
<geoCoordinate degrees="44.81467" direction="east" orientation="longitude" precision="1" value="44.81467">44.81467</geoCoordinate>
; alt. 100 m; 15 Nov 2001; B.L. Fisher et al. leg.; BLF04388, CASENT0425236, bottom specimen (CASC).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
.
</emphasis>
Madagascar. •7w., 1s.; same data as for holotype; CASENT0425237-CASENT0425239, CASENT0872078, CASENT0872169-CASENT0872172 (CASC).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
.
</emphasis>
Small species: HL: 1.05-1.04, HW: 1.01-0.98, WL: 0.76-0.69; head in full-face view square, anterior and posterior sides of eyes slightly convex; in lateral view sub-oval; ventral and dorsal faces convex; dorsal face not depressed posteriorly; sides of the head with sparse, relatively long, erect pilosity; frons with dense, thick, and longitudinal rugae, interspaces smooth, only posterior part of frons finely foveolate; inner hypostomal teeth distinct, closely spaced, triangular with relatively narrow base, apex rounded; outer hypostomal teeth slightly higher than inner hypostomal teeth, and with wider base, triangular; gaster indistinctly shagreened; body brown to dark brown.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
.
</emphasis>
Head and mesosoma foveolate; promesonotum low, long, slightly convex, with relatively steep posterior declivity; propodeal spines moderately long.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
Measurements (
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
= 2): HL: 1.05-1.04; HW: 1.01-0.98; SL: 0.49-0.51; EL: 0.14-0.14; WL: 0.76-0.69; PSL: 0.15-0.15; MTL: 0.49-0.46; PNW: 0.45-0.45; PTW: 0.11-0.12; PPW: 0.32-0.33; CI: 95.5-94.2; SI: 49.0-51.9; PSLI: 14.3-14.6; PPI: 34.5-34.5; PNI: 44.9-46.3; MTI: 48.6-47.4.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
In full-face view square, anterior and posterior sides of eyes slightly convex (Fig.
<figureCitation captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole bemarahaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0425239) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0425236)." figureDoi="10.3897/zookeys.905.39592.figure61" httpUri="https://binary.pensoft.net/fig/373108" pageId="0" pageNumber="1">61B</figureCitation>
). In lateral view sub-oval; ventral and dorsal faces convex; dorsal face not depressed posteriorly; inner hypostomal teeth visible. Sides of the head with sparse, relatively long, erect pilosity; whole head with dense, short, suberect to erect pilosity. Antennal scrobes very indistinct and not delimited by carinulae; scrobe surface shiny, finely foveolate with sparse, thick, longitudinal to irregular in posterior part rugulae. Occipital lobes shiny, with fine foveolae and distinct, sparse, irregular rugae, sculpture not weakening posteriorly; frons and malar area with dense, thick, and longitudinal rugae, interspaces smooth, and only posterior part of frons finely foveolate; genae shiny, with fine rugo-foveolae. Centre of clypeus smooth and shiny, lateral sides with longitudinal rugae; median notch present, narrow, and shallow; median longitudinal carina present; lateral longitudinal carinae present. Scape, when laid back, slightly exceeding the midlength of head; pilosity suberect to erect (Fig.
<figureCitation captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole bemarahaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0425239) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0425236)." figureDoi="10.3897/zookeys.905.39592.figure61" httpUri="https://binary.pensoft.net/fig/373108" pageId="0" pageNumber="1">61B, D</figureCitation>
). Inner hypostomal teeth distinct, closely spaced, triangular with relatively narrow base, apex rounded; outer hypostomal teeth slightly higher than outer hypostomal teeth, and with wider base, triangular (Fig.
<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">84G</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
.
</emphasis>
In lateral view, promesonotum moderately short, relatively low and convex, posterior mesonotum steep, with small tubercle-like projections; promesonotal groove absent; metanotal groove absent; propodeal spines small, triangular, with rounded apex; humeral area laterally well produced (Fig.
<figureCitation captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole bemarahaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0425239) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0425236)." figureDoi="10.3897/zookeys.905.39592.figure61" httpUri="https://binary.pensoft.net/fig/373108" pageId="0" pageNumber="1">61D</figureCitation>
). Surface shiny, with fine but distinct foveolae, additional sparse transverse to irregular rugae on promesonotal dorsum, sculpture slightly weakening on dorsal surface. Pilosity sparse, long, and erect (Fig.
<figureCitation captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole bemarahaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0425239) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0425236)." figureDoi="10.3897/zookeys.905.39592.figure61" httpUri="https://binary.pensoft.net/fig/373108" pageId="0" pageNumber="1">61D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
Shiny and with fine foveolae; peduncle short, with indistinct horizontal lobes on its basal part; node relatively high, triangular, with rounded apex, in rear view node slightly convex; pilosity moderately sparse and erect (Fig.
<figureCitation captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole bemarahaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0425239) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0425236)." figureDoi="10.3897/zookeys.905.39592.figure61" httpUri="https://binary.pensoft.net/fig/373108" pageId="0" pageNumber="1">61D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
.
</emphasis>
Shiny, with fine and sparse foveolae, smooth on dorsum; in dorsal view sides with short, acute, and triangular projections; pilosity long, moderately long, and erect (Fig.
<figureCitation captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole bemarahaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0425239) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0425236)." figureDoi="10.3897/zookeys.905.39592.figure61" httpUri="https://binary.pensoft.net/fig/373108" pageId="0" pageNumber="1">61D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
Shiny and indistinctly shagreened; pilosity sparse, short, and erect (Fig.
<figureCitation captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole bemarahaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0425239) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0425236)." figureDoi="10.3897/zookeys.905.39592.figure61" httpUri="https://binary.pensoft.net/fig/373108" pageId="0" pageNumber="1">61D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
.
</emphasis>
Unicolourous, brown to dark brown; malar area and lower frons bright brown; legs yellow (Fig.
<figureCitation captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole bemarahaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0425239) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0425236)." figureDoi="10.3897/zookeys.905.39592.figure61" httpUri="https://binary.pensoft.net/fig/373108" pageId="0" pageNumber="1">61D, F</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.905.39592.figure61" httpUri="https://binary.pensoft.net/fig/373108" pageId="0" pageNumber="1" start="Figure 61" startId="F61">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 61.</emphasis>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole bemarahaensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bemarahaensis">Pheidole bemarahaensis</taxonomicName>
sp. nov., full-face view (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
), profile (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
), and dorsal view (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
) of paratype minor worker (CASENT0425239) and full-face view (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
), profile (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
), and dorsal view (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
) of holotype major worker (CASENT0425236).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
Measurements (
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
= 7): HL: 0.48-0.51 (0.49); HW: 0.43-0.47 (0.45); SL: 0.42-0.46 (0.45); EL: 0.1-0.11 (0.1); WL: 0.56-0.59 (0.58); PSL: 0.09-0.11 (0.1); MTL: 0.32-0.37 (0.35); PNW: 0.28-0.31 (0.29); PTW: 0.06-0.08 (0.07); PPW: 0.12-0.13 (0.12); CI: 88.8-93.6 (91.1); SI: 98.1-105.1 (100.2); PSLI: 17.9-21.0 (19.8); PPI: 53.4-61.4 (56.3); PNI: 64.1-65.9 (65.2); MTI: 72.5-80.4 (77.1).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
Occipital margin straight or indistinctly concave; occipital carina absent (Fig.
<figureCitation captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole bemarahaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0425239) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0425236)." figureDoi="10.3897/zookeys.905.39592.figure61" httpUri="https://binary.pensoft.net/fig/373108" pageId="0" pageNumber="1">61A</figureCitation>
). Pilosity sparse, short, suberect to erect. Head foveolate. Clypeus with fine and sometimes reduced foveolae and rugae; median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by one-fifth of its length; pilosity suberect to erect (Fig.
<figureCitation captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole bemarahaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0425239) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0425236)." figureDoi="10.3897/zookeys.905.39592.figure61" httpUri="https://binary.pensoft.net/fig/373108" pageId="0" pageNumber="1">61A, C</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
.
</emphasis>
In lateral view, promesonotum low, long, slightly convex, with relatively steep posterior declivity; promesonotal groove absent; metanotal groove present; propodeal spines moderately long, triangular, apex acute (Fig.
<figureCitation captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole bemarahaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0425239) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0425236)." figureDoi="10.3897/zookeys.905.39592.figure61" httpUri="https://binary.pensoft.net/fig/373108" pageId="0" pageNumber="1">61C</figureCitation>
). Sculpture foveolate. Pilosity sparse, short, and erect (Fig.
<figureCitation captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole bemarahaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0425239) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0425236)." figureDoi="10.3897/zookeys.905.39592.figure61" httpUri="https://binary.pensoft.net/fig/373108" pageId="0" pageNumber="1">61C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
Shiny and with fine foveolae; peduncle short and thin with ventral face slightly convex; node moderately high, triangular, and small; with few short, erect setae (Fig.
<figureCitation captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole bemarahaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0425239) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0425236)." figureDoi="10.3897/zookeys.905.39592.figure61" httpUri="https://binary.pensoft.net/fig/373108" pageId="0" pageNumber="1">61C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
.
</emphasis>
Short, low, and slightly convex; with few short, erect setae (Fig.
<figureCitation captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole bemarahaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0425239) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0425236)." figureDoi="10.3897/zookeys.905.39592.figure61" httpUri="https://binary.pensoft.net/fig/373108" pageId="0" pageNumber="1">61C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
With moderately sparse, erect pilosity (Fig.
<figureCitation captionStart="Figure 61" captionStartId="F61" captionText="Figure 61. Pheidole bemarahaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0425239) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0425236)." figureDoi="10.3897/zookeys.905.39592.figure61" httpUri="https://binary.pensoft.net/fig/373108" pageId="0" pageNumber="1">61C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
.
</emphasis>
Unicolourous, yellow (Fig.
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</paragraph>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">From the type locality.</paragraph>
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<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
<paragraph pageId="0" pageNumber="1">The species was collected at 100 m in elevation, in tropical dry forest. Nest was located in a rotten log.</paragraph>
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<subSubSection pageId="0" pageNumber="1" type="comments">
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Pheidole bemarahaensis</emphasis>
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sp. nov. is most similar to
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<emphasis italics="true" pageId="0" pageNumber="1">P. bemarivoensis</emphasis>
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sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
.
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<emphasis italics="true" pageId="0" pageNumber="1">Pheidole bemarahaensis</emphasis>
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sp. nov. differs from
<taxonomicName lsidName="P. bemarivoensis" pageId="0" pageNumber="1" rank="species" species="bemarivoensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. bemarivoensis</emphasis>
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sp. nov. in at least lower part of frons smooth between rugae, more distinct rugo-foveolate sculpture on genae, and thicker dorsal surface of promesonotum.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
.
</emphasis>
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<emphasis italics="true" pageId="0" pageNumber="1">P. bemarahaensis</emphasis>
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sp. nov. differs from
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<emphasis italics="true" pageId="0" pageNumber="1">P. bemarivoensis</emphasis>
</taxonomicName>
sp. nov. in moderately long propodeal spines.
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Tribe
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Coiffait, 1963: 381 [stem: Metrotyphl-]. Type genus:
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</mods:role>
<mods:namePart>Hu, Wenjia</mods:namePart>
</mods:name>
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</mods:role>
<mods:namePart>Zheng, Xinqing</mods:namePart>
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<mods:namePart>Affendi, Yang Amri</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ooi, Jillian Lean Sim</mods:namePart>
</mods:name>
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ma, Zhiyuan</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rizman-Idid, Mohammed</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chan, Albert Apollo</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Biodiversity Data Journal</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>7</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>47537</mods:start>
<mods:end>47537</mods:end>
</mods:extent>
</mods:part>
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<mods:url>http://dx.doi.org/10.3897/BDJ.7.e47537</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/BDJ.7.e47537</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2828-7-e47537</mods:identifier>
<mods:identifier type="ZooBank">F940F7FD0A3541E98BDD33F83C2369D5</mods:identifier>
<mods:identifier type="Pensoft-UUID">AE1BE74780565E8D9B3522053F3B0983</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:DBE9CFF350B55509B4DEA3BD98718E9D" httpUri="http://treatment.plazi.org/id/DBE9CFF350B55509B4DEA3BD98718E9D" lastPageNumber="47537" pageId="0" pageNumber="47537">
<subSubSection pageId="0" pageNumber="47537" type="nomenclature">
<paragraph pageId="0" pageNumber="47537">
<taxonomicName LSID="DBE9CFF3-50B5-5509-B4DE-A3BD98718E9D" authority="Quoy &amp; Gaimard, 1834" authorityName="Quoy &amp; Gaimard" authorityYear="1834" class="Actinopterygii" family="Blenniidae" genus="Aspidontus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aspidontus taeniatus" order="Perciformes" pageId="0" pageNumber="47537" phylum="Chordata" rank="species" species="taeniatus">Aspidontus taeniatus Quoy &amp; Gaimard, 1834</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47537" type="materials_examined">
<paragraph pageId="0" pageNumber="47537">Materials</paragraph>
<paragraph pageId="0" pageNumber="47537">
<materialsCitation country="Malaysia" location="Redang islands" pageId="0" pageNumber="47537" specimenCount="1" typeStatus="Other material">
<emphasis bold="true" pageId="0" pageNumber="47537">Type status:</emphasis>
<typeStatus pageId="0" pageNumber="47537">Other material</typeStatus>
.
<emphasis bold="true" pageId="0" pageNumber="47537">Occurrence:</emphasis>
occurrenceID: BDJ_12482_27;
<emphasis bold="true" pageId="0" pageNumber="47537">Location:</emphasis>
country:
<collectingCountry pageId="0" pageNumber="47537">Malaysia</collectingCountry>
; locality:
<location LSID="urn:lsid:plazi:treatment:DBE9CFF350B55509B4DEA3BD98718E9D:23B92FDC5F557CC890A510C0147F89D4" country="Malaysia" name="Redang islands" pageId="0" pageNumber="47537">Redang islands</location>
;
<emphasis bold="true" pageId="0" pageNumber="47537">Identification:</emphasis>
identifiedBy:
<determinerName pageId="0" pageNumber="47537">Loh KH and Du Jianguo</determinerName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47537" type="notes">
<paragraph pageId="0" pageNumber="47537">Notes</paragraph>
<paragraph pageId="0" pageNumber="47537">Newly recorded in Redang islands + This study.</paragraph>
</subSubSection>
</treatment>
</document>