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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1127.85737" ID-Pensoft-Pub="1313-2970-1127-1" ID-Pensoft-UUID="D72971CE12A85992AFFD69F186474E85" ID-ZooBank="4759AFC32EFD47A7853F645FB32829BA" ModsDocID="1313-2970-1127-1" checkinTime="1667414686597" checkinUser="pensoft" docAuthor="Ramahandrison, Andriamirado Tahina, Rakouth, Bakolimalala &amp; Manuel, Michael" docDate="2022" docId="DBD337F8D15C574DA2B62586C506AB51" docLanguage="en" docName="ZooKeys 1127: 1-60" docOrigin="ZooKeys 1127" docPubDate="2022-11-02" docSource="http://dx.doi.org/10.3897/zookeys.1127.85737" docTitle="Copelatus zanabato Manuel &amp; Ramahandrison 2022, sp. nov." docType="treatment" docUuid="7D683E52-D50E-485A-B2CA-5E44D3F4CC5C" docUuidSource="ZooBank" docVersion="1" id="D72971CE12A85992AFFD69F186474E85" lastPageNumber="1" masterDocId="D72971CE12A85992AFFD69F186474E85" masterDocTitle="The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae)" masterLastPageNumber="60" masterPageNumber="1" pageNumber="1" updateTime="1667414686597" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ramahandrison, Andriamirado Tahina</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0833-8730</mods:nameIdentifier>
<mods:affiliation>Departement de Biologie et Ecologie Vegetales, Faculte des Sciences, BP 906, Universite d'Antananarivo, Antananarivo, Madagascar &amp; Sorbonne Universite, Institut de Systematique, Evolution, Biodiversite (UMR 7205), MNHN SU CNRS EPHE UA, Case 05, 7 quai St Bernard, Paris, France</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rakouth, Bakolimalala</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5710-2006</mods:nameIdentifier>
<mods:affiliation>Departement de Biologie et Ecologie Vegetales, Faculte des Sciences, BP 906, Universite d'Antananarivo, Antananarivo, Madagascar</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Manuel, Michael</mods:namePart>
<mods:affiliation>Sorbonne Universite, Institut de Systematique, Evolution, Biodiversite (UMR 7205), MNHN SU CNRS EPHE UA, Case 05, 7 quai St Bernard, Paris, France</mods:affiliation>
<mods:nameIdentifier type="email">manuel1570@yahoo.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-11-02</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1127</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>60</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1127.85737</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1127.85737</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1127-1</mods:identifier>
<mods:identifier type="ZooBank">4759AFC32EFD47A7853F645FB32829BA</mods:identifier>
<mods:identifier type="Pensoft-UUID">D72971CE12A85992AFFD69F186474E85</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:7D683E52-D50E-485A-B2CA-5E44D3F4CC5C" httpUri="http://treatment.plazi.org/id/DBD337F8D15C574DA2B62586C506AB51" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="https://zoobank.org/7D683E52-D50E-485A-B2CA-5E44D3F4CC5C" authority="Manuel &amp; Ramahandrison" authorityName="Manuel &amp; Ramahandrison" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus zanabato" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="zanabato" status="sp. nov.">Copelatus zanabato Manuel &amp; Ramahandrison</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 3" captionStartId="F4" captionText="Figure 3. Habitus in dorsal view. Scale bars: 2 mm A Copelatus malavergnorum sp. nov. (holotype) B Copelatus zanabato sp. nov. (holotype)." figureDoi="10.3897/zookeys.1127.85737.figure3" httpUri="https://binary.pensoft.net/fig/764729" pageId="0" pageNumber="1">Figs 3B</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F5" captionText="Figure 4. Male genitalia. Scale bars: 200 µm A-D Copelatus malavergnorum sp. nov. (holotype) E-H Copelatus zanabato sp. nov. (holotype) A, E Median lobe of aedeagus in right lateral view B, F Median lobe of aedeagus in ventral view C, G Median lobe of aedeagus in left lateral view D, H Left paramere." figureDoi="10.3897/zookeys.1127.85737.figure4" httpUri="https://binary.pensoft.net/fig/764730" pageId="0" pageNumber="1">, 4E-H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
Madagascar, Toliara province, Malaimbandy municipality, Makay massif (northern part), ca. 21 km W of Tsimazava, ca.
<geoCoordinate degrees="20" direction="south" minutes="42" orientation="latitude" precision="925" value="-20.7">20°42'S</geoCoordinate>
,
<geoCoordinate degrees="45" direction="east" minutes="30" orientation="longitude" precision="925" value="45.5">45°30'E</geoCoordinate>
, altitude ca. 430 m a.s.l.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
♂: &quot;Madagascar. Ex-prov. Toliara / Makay massif, ca. 21 km / W Tsimazava /
<geoCoordinate degrees="20" direction="south" minutes="42" orientation="latitude" precision="15" seconds="10" value="-20.702778">20°42'10&quot;S</geoCoordinate>
,
<geoCoordinate degrees="45" direction="east" minutes="30" orientation="longitude" precision="15" seconds="18" value="45.505">45°30'18&quot;E</geoCoordinate>
[pr.] // 12.IV.2018. Ramahandrison leg. / Alt. 433 m. Pool in a / very narrow and dark / canyon. [pr.] // Holotype /
<taxonomicName authorityName="Manuel &amp; Ramahandrison" authorityYear="2022" class="Insecta" family="Polygonaceae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus zanabato" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="zanabato">Copelatus zanabato</taxonomicName>
sp. nov. / Manuel &amp; Ramahandrison, 2022 [red, pr.]&quot; (MNHN). Paratypes: 1 ♀: same as holotype. 1 ♂: &quot;Madagascar. Ex-prov. Toliara / Makay massif, ca. 20 km / WSW Tsimazava /
<geoCoordinate degrees="20" direction="south" minutes="43" orientation="latitude" precision="15" seconds="54" value="-20.731667">20°43'54&quot;S</geoCoordinate>
,
<geoCoordinate degrees="45" direction="east" minutes="31" orientation="longitude" precision="15" seconds="10" value="45.519444">45°31'10&quot;E</geoCoordinate>
/ 14.IV.2018. Ramahandrison leg. [pr.] // Alt. 437 m. Small / pond in a canyon at / Ampasimaiky [pr.]&quot; (CMM). Both paratypes with respective red label.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species belongs to the
<taxonomicName authorityName="Guerin-Meneville" authorityYear="1847" class="Insecta" family="Polygonaceae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus erichsonii" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="erichsonii">
<emphasis italics="true" pageId="0" pageNumber="1">Copelatus erichsonii</emphasis>
</taxonomicName>
-group. It is externally rather similar to
<taxonomicName family="Cyperaceae" kingdom="Animalia" lsidName="C. acamas" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="acamas">
<emphasis italics="true" pageId="0" pageNumber="1">C. acamas</emphasis>
</taxonomicName>
, from which it differs by distinctly smaller size; habitus narrower with sides more parallel, dorsally much flatter; discal stria IX on elytron more strongly abbreviated anteriorly; strioles on pronotum surface sparser and much more weakly impressed; shape of median lobe of aedeagus very different. Among
<taxonomicName authorityName="Erichson" authorityYear="1832" class="Insecta" family="Polygonaceae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Copelatus</emphasis>
</taxonomicName>
species known from Madagascar, the aedeagus of
<taxonomicName family="Cyperaceae" kingdom="Animalia" lsidName="C. zanabato" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="zanabato">
<emphasis italics="true" pageId="0" pageNumber="1">C. zanabato</emphasis>
</taxonomicName>
sp. nov. is most similar to that of
<taxonomicName family="Cyperaceae" kingdom="Animalia" lsidName="C. andobonicus" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="andobonicus">
<emphasis italics="true" pageId="0" pageNumber="1">C. andobonicus</emphasis>
</taxonomicName>
. From the latter, the new species differs by: habitus narrower with sides more parallel, dorsally much flatter; pronotum paler and elytra with darker linear colouration following the striae much less contrasted with respect to paler background; strioles on pronotum surface denser, present on whole surface (in
<taxonomicName family="Cyperaceae" kingdom="Animalia" lsidName="C. andobonicus" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="andobonicus">
<emphasis italics="true" pageId="0" pageNumber="1">C. andobonicus</emphasis>
</taxonomicName>
almost without striae in anterior disk region); median lobe of aedeagus in lateral view with broad flat protuberance on ventral side ca. halfway between base and apex (in
<taxonomicName family="Cyperaceae" kingdom="Animalia" lsidName="C. andobonicus" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="andobonicus">
<emphasis italics="true" pageId="0" pageNumber="1">C. andobonicus</emphasis>
</taxonomicName>
with much smaller protuberance at ca. basal third) and apical third much broader, in ventral view with apical region much broader and evenly narrowed, twisted on the left farther from apex.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description of holotype.</paragraph>
<paragraph pageId="0" pageNumber="1">
Body shape elongate oval, with sides subparallel (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F4" captionText="Figure 3. Habitus in dorsal view. Scale bars: 2 mm A Copelatus malavergnorum sp. nov. (holotype) B Copelatus zanabato sp. nov. (holotype)." figureDoi="10.3897/zookeys.1127.85737.figure3" httpUri="https://binary.pensoft.net/fig/764729" pageId="0" pageNumber="1">3B</figureCitation>
), dorsally weakly convex. Pronotum sides evenly curved and converging from posterior angle. Ratio between maximum width of pronotum and maximum body width ~ 0.90. Head uniformly light rufo-testaceous. Ratio between interocular distance and maximum width of head ~ 0.68. Pronotum medially rufous, laterally colour becoming gradually rufo-testaceous. Elytra light chestnut brown, with darker linear colouration along striae (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F4" captionText="Figure 3. Habitus in dorsal view. Scale bars: 2 mm A Copelatus malavergnorum sp. nov. (holotype) B Copelatus zanabato sp. nov. (holotype)." figureDoi="10.3897/zookeys.1127.85737.figure3" httpUri="https://binary.pensoft.net/fig/764729" pageId="0" pageNumber="1">3B</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">Elytra with ten well-impressed discal and one submarginal striae. Stria IX abbreviated anteriorly. Striae I and II diverging anteriorly. Submarginal stria starting slightly before elytron midlength, fragmented anteriorly. Head, pronotum and elytra with fine reticulation and fine punctation. Whole surface of pronotum with rather dense, short and fine strioles, in medial disk region strioles even finer.</paragraph>
<paragraph pageId="0" pageNumber="1">Ventral side uniformly rufo-testaceous. Metacoxal plates with moderately impressed short strioles; visible abdominal ventrites I-III with denser and longer very fine strioles. Prosternal process short and broad, with rounded apex. Metacoxal lines short, ending anteriorly at large distance from posterior margin of metaventrite, moderately diverging anteriorly.</paragraph>
<paragraph pageId="0" pageNumber="1">Appendages: Antennae, palps, forelegs and midlegs testaceous, hindlegs rufo-testaceous. First three pro- and mesotarsomeres widened and ventrally equipped with suction cups; number of suction cups per article (I-III) 7:4:4 on both pro- and mesotarsus. Protibia at base shortly narrow, with shallow protuberance along ventral margin, distally broadened. Pro- and mesotarsal claws unmodified.</paragraph>
<paragraph pageId="0" pageNumber="1">
Median lobe and parameres as in Fig.
<figureCitation captionStart="Figure 4" captionStartId="F5" captionText="Figure 4. Male genitalia. Scale bars: 200 µm A-D Copelatus malavergnorum sp. nov. (holotype) E-H Copelatus zanabato sp. nov. (holotype) A, E Median lobe of aedeagus in right lateral view B, F Median lobe of aedeagus in ventral view C, G Median lobe of aedeagus in left lateral view D, H Left paramere." figureDoi="10.3897/zookeys.1127.85737.figure4" httpUri="https://binary.pensoft.net/fig/764730" pageId="0" pageNumber="1">4E-H</figureCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Female.</emphasis>
Strioles on pronotum surface denser. Pro- and mesotarsomeres and protibia unmodified.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="1">Holotype: TL 6.35 mm, TL without head 5.7 mm, MW 2.9 mm, ratio TL/MW 2.20. Paratypes: TL 6.55-6.8 mm, TL without head 5.9-6.1 mm, MW 3.0-3.1 mm, ratio TL/MW 2.17.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="variation">
<paragraph pageId="0" pageNumber="1">Variation.</paragraph>
<paragraph pageId="0" pageNumber="1">In the male paratype, strioles on pronotum are longer and more deeply impressed than in the holotype, and the metacoxal lines are slightly longer. In the female paratype, the elytral stria V is slightly abbreviated anteriorly.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">The species name literally means &quot;son of the rock&quot; in Malagasy. It is an invariable name standing in apposition.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">So far known only from northern Makay in Madagascar.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="habitat">
<paragraph pageId="0" pageNumber="1">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species was collected at two sites located in two nearby canyons in northern inner Makay. Two specimens were sampled at site MAK-46, an isolated pool (~ 1 m
<normalizedToken originalValue="×">x</normalizedToken>
3 m) on the bottom of a narrow and dark canyon, and one specimen at site MAK-50, a stagnant temporary pond (~ 3 m
<normalizedToken originalValue="×">x</normalizedToken>
7 m) situated in a wider canyon and in a more open environment. Both habitats were characterised by sandy bottom with some plant debris, absence of visible inflow / outflow, somewhat turbid water and no vegetation. Other species of aquatic
<taxonomicName kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" rank="subOrder" subOrder="Adephaga">Adephaga</taxonomicName>
(all
<taxonomicName authorityName="Leach" authorityYear="1815" class="Insecta" family="Dytiscidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Dytiscidae</taxonomicName>
) sampled at the same sites:
<taxonomicName authorityName="Guignot" authorityYear="1955" class="Insecta" family="Polygonaceae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus acamas" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="acamas">
<emphasis italics="true" pageId="0" pageNumber="1">Copelatus acamas</emphasis>
</taxonomicName>
,
<taxonomicName family="Cyperaceae" kingdom="Animalia" lsidName="C. ruficapillus" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="ruficapillus">
<emphasis italics="true" pageId="0" pageNumber="1">C. ruficapillus</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Cybister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cybister operosus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="operosus">
<emphasis italics="true" pageId="0" pageNumber="1">Cybister operosus</emphasis>
</taxonomicName>
Sharp, 1882,
<taxonomicName authorityName="Aube" authorityYear="1838" class="Insecta" family="Dytiscidae" genus="Hydaticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydaticus sobrinus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sobrinus">
<emphasis italics="true" pageId="0" pageNumber="1">Hydaticus sobrinus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Aubé">Aube</normalizedToken>
, 1838,
<taxonomicName authorityName="Regimbart" authorityYear="1895" class="Insecta" family="Dytiscidae" genus="Hyphydrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyphydrus separandus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="separandus">
<emphasis italics="true" pageId="0" pageNumber="1">Hyphydrus separandus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Régimbart">Regimbart</normalizedToken>
, 1895,
<taxonomicName authorityName="Manuel &amp; Ramahandrison" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus makay" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="makay">
<emphasis italics="true" pageId="0" pageNumber="1">Laccophilus makay</emphasis>
</taxonomicName>
,
<taxonomicName class="Insecta" family="Dytiscidae" genus="Pachynectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachynectes" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pachynectes</emphasis>
</taxonomicName>
sp. Ma1, and
<taxonomicName genus="P." lsidName="P." pageId="0" pageNumber="1" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">P.</emphasis>
</taxonomicName>
sp. Ma4.
</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName id="2294E1E56A90487ACD7AACD0C40AA836" authority="Ehrh." authorityName="Ehrh." class="Magnoliopsida" family="Brassicaceae" genus="Erysimum" kingdom="Plantae" order="Brassicales" pageId="null" pageNumber="244" phylum="Tracheophyta" rank="species" species="diffusum">
<pageBreakToken id="C3751621A10B74A6433CEDA08B76C719" pageId="null" pageNumber="244" start="start">Erysimum</pageBreakToken>
diffusum Ehrh.
</taxonomicName>
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</subSubSection>
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<paragraph id="22B86EA1D2EF6489983330B83691DA8F" pageId="null" pageNumber="244">
(
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<emphasis id="A216E016E71132B69F2961ED48BDB7F8" italics="true" pageId="null" pageNumber="244">E. canescens</emphasis>
Roth
</taxonomicName>
)
</paragraph>
</subSubSection>
<subSubSection id="3815B01BA78CBAD212F19444353CE4B6" pageId="null" pageNumber="244" type="vernacular_names">
<paragraph id="59684E87226E5B94B609C9D421416D07" pageId="null" pageNumber="244">
Grauer
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</paragraph>
</subSubSection>
</title>
<subSubSection id="5E84729B03E49089D4E8F1ADD2E3ADDE" pageId="null" pageNumber="244" type="description">
<paragraph id="42F1749B02A1D567E2DDFD58E26BA766" pageId="null" pageNumber="244">
Meist 2
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; meist ohne sterile Blattrosetten; 30-100 cm hoch.
<normalizedToken id="DC1165A477651537407E5B09E5F6BE92" originalValue="Kelchblätter" pageId="null" pageNumber="244">Kelchblaetter</normalizedToken>
5-9 mm lang,
<emphasis id="489BD8C7C0013EF77F3E05D1B857AA51" italics="true" pageId="null" pageNumber="244">
am Grunde nicht ausgebuchtet.
<normalizedToken id="E10E0D1237DE743A428C12733277F76B" originalValue="Kronblätter" pageId="null" pageNumber="244">Kronblaetter</normalizedToken>
8-14 mm lang
</emphasis>
, gelb.
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30-80 mm lang und
<emphasis id="1541227D1A1E50F73FE5DCB3C16036B5" italics="true" pageId="null" pageNumber="244">0,6-1 mm</emphasis>
<emphasis id="11BC268EA1F7AA8870247F49C2FC2C75" italics="true" pageId="null" pageNumber="244">
<pageBreakToken id="31FBA125ABDE5AA26DCBC03F7407E57B" pageId="null" pageNumber="244">dick</pageBreakToken>
</emphasis>
, auf den Kanten zerstreut, auf den
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dicht behaart (Epidermis meist nicht sichtbar); Griffel 0,5-1,5 mm lang. Samen 0,8-1,3 mm lang. -
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:
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Sommer.
</paragraph>
<paragraph id="A44C5A54C65F42B1D063E20BE6D154D9" pageId="null" pageNumber="244">
<emphasis id="DBA6EB64D89211EDC600732CB38E06BB" italics="true" pageId="null" pageNumber="244">Zytologische Angaben. 2n</emphasis>
=
<emphasis id="A001DE77941AB359867316D5D9D69E48" italics="true" pageId="null" pageNumber="244">28:</emphasis>
Material aus
<normalizedToken id="1E45CB4B2E68F2E83556ECD8A8C33649" originalValue="Niederösterreich" pageId="null" pageNumber="244">Niederoesterreich</normalizedToken>
und dem Burgenland (Polatschek 1966).
<emphasis id="50FF8C2294BFA7256E1F8A10CBB13262" italics="true" pageId="null" pageNumber="244">2n ca. 32:</emphasis>
Material aus Ungarn (Baksay in
<normalizedToken id="03E091D1AF2D28ABE91B6D94DBF6BBD0" originalValue="Löve" pageId="null" pageNumber="244">Loeve</normalizedToken>
und
<normalizedToken id="2FAA7AE585F6A8327874360B27E2BF77" originalValue="Löve" pageId="null" pageNumber="244">Loeve</normalizedToken>
1961).
<emphasis id="F9890F66276D828A2CBDF0C2329DD1A8" italics="true" pageId="null" pageNumber="244">2n</emphasis>
=
<emphasis id="53ED953331736E5617D892AB89C39C9F" italics="true" pageId="null" pageNumber="244">70:</emphasis>
Material von 4 Stellen aus
<normalizedToken id="1A5A4B044F12635F743BE9678ED786A1" originalValue="Niederröstereich" pageId="null" pageNumber="244">Niederroestereich</normalizedToken>
(Polatschek 1966).
<emphasis id="84D7BA87CFFC67C2EDE518F1BBDD0D2C" italics="true" pageId="null" pageNumber="244">2n</emphasis>
=
<emphasis id="FAE77C28BCD1035A697E08A6BBC7EAB6" italics="true" pageId="null" pageNumber="244">72:</emphasis>
Material aus botanischem Garten (Manton 1932), aus
<normalizedToken id="A12BF9727080F62EED7FF8A768817CDB" originalValue="Niederösterreich" pageId="null" pageNumber="244">Niederoesterreich</normalizedToken>
(Polatschek 1966). Favarger (1964)
<normalizedToken id="7829B52BCC27F2C24061D6E061BA0900" originalValue="zählte" pageId="null" pageNumber="244">zaehlte</normalizedToken>
an Material aus dem Aostatal (
<normalizedToken id="85D9A286903B89856618D13078C76DC9" originalValue="möglicherweise" pageId="null" pageNumber="244">moeglicherweise</normalizedToken>
Bastard mit
<emphasis id="4ED71F0C42EC26CE667510749F3187EC" italics="true" pageId="null" pageNumber="244">E. helvcticum</emphasis>
) 2n ca. 60.
</paragraph>
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<emphasis id="8032E78BC3801972E376F72423AD08BD" italics="true" pageId="null" pageNumber="244">Standort.</emphasis>
Kollin und montan. Trockene, steinige
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in warmen Lagen. Felsige
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, Trockenwiesen,
<normalizedToken id="8FB5C941B37903F24F14188F7740E7A1" originalValue="Wegränder" pageId="null" pageNumber="244">Wegraender</normalizedToken>
.
</paragraph>
<paragraph id="B1CF20E3A7228626039ECEBDEA97FAF9" pageId="null" pageNumber="244">
<emphasis id="483CC594A0DDB4CAF3334E833A6CD1F1" italics="true" pageId="null" pageNumber="244">
Verbreitung.
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Pflanze:
</emphasis>
West- und
<normalizedToken id="A92E002DD59554FE6EB6A27848E4CA1F" originalValue="nordwärts" pageId="null" pageNumber="244">nordwaerts</normalizedToken>
bis Aostatal (?),
<normalizedToken id="587A4ECB6EB4AEDE177B011ED59DA077" originalValue="Oberösterreich" pageId="null" pageNumber="244">Oberoesterreich</normalizedToken>
,
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, Ukraine;
<normalizedToken id="0B52C31BDBBF97D5611CBD26DFE229F8" originalValue="süd-" pageId="null" pageNumber="244">sued-</normalizedToken>
und
<normalizedToken id="1323EC44125FBBD800C45D4005AB1BA6" originalValue="ostwärts" pageId="null" pageNumber="244">ostwaerts</normalizedToken>
bis Mazedonien, Kaukasus; Sibirien (?). - Im Gebiet: Angaben aus den Bergamasker Alpen (Monte Resegone, Monte Bracca), dem Vintschgau, dem Valsesia und dem Aostatal; die
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der Pflanzen
<normalizedToken id="45BC4A911E0A63EE158F9D82084AAACA" originalValue="muß" pageId="null" pageNumber="244">muss</normalizedToken>
indessen
<normalizedToken id="2BD145D4D427A4472F374F983CBB6E4F" originalValue="abgeklärt" pageId="null" pageNumber="244">abgeklaert</normalizedToken>
werden; sonst gelegentlich adventiv.
</paragraph>
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132
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<mods:role>
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</mods:role>
<mods:namePart>Newton, Alfred F.</mods:namePart>
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<mods:role>
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<mods:role>
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<treatment ID-GBIF-Taxon="159361976" LSID="urn:lsid:plazi:treatment:DBD37F6DF026677EBDD4EEAF17F86FC0" httpUri="http://treatment.plazi.org/id/DBD37F6DF026677EBDD4EEAF17F86FC0" lastPageNumber="143" pageId="142" pageNumber="143">
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<paragraph pageId="142" pageNumber="143">
Subfamily
<taxonomicName authority="Champion, 1897" authorityName="Champion" authorityYear="1897" lsidName="" pageId="142" pageNumber="143" rank="subFamily" subFamily="Anchytarsinae">Anchytarsinae Champion, 1897</taxonomicName>
</paragraph>
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<subSubSection pageId="142" pageNumber="143" type="reference_group">
<paragraph pageId="142" pageNumber="143">
<taxonomicName genus="Coloboderides" lsidName="Coloboderides" pageId="142" pageNumber="143" rank="genus">Coloboderides</taxonomicName>
Erichson, 1847a: 174 [stem: Coloboder-]. Type genus:
<taxonomicName class="Insecta" genus="Colobodera" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Colobodera" order="Coleoptera" pageId="142" pageNumber="143" phylum="Arthropoda" rank="genus">Colobodera</taxonomicName>
Klug, 1838 [syn. of
<taxonomicName class="Insecta" family="Dascillidae" genus="Daemon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Daemon" order="Coleoptera" pageId="142" pageNumber="143" phylum="Arthropoda" rank="genus">Daemon</taxonomicName>
Laporte, 1836].
</paragraph>
<paragraph pageId="142" pageNumber="143">
<taxonomicName lsidName="" pageId="142" pageNumber="143" rank="tribe" tribe="Anchytarsini">Anchytarsini</taxonomicName>
Champion, 1897: 593 [stem: Anchytars-]. Type genus:
<taxonomicName class="Insecta" family="Ptilodactylidae" genus="Anchytarsus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anchytarsus" order="Coleoptera" pageId="142" pageNumber="143" phylum="Arthropoda" rank="genus">Anchytarsus</taxonomicName>
<normalizedToken originalValue="Guérin-Méneville">Guerin-Meneville</normalizedToken>
, 1843. Comment:
<taxonomicName lsidName="" pageId="142" pageNumber="143" rank="subFamily" subFamily="Coloboderinae">Coloboderinae</taxonomicName>
Erichson, 1847 is the oldest name for this tribe, however since
<taxonomicName lsidName="" pageId="142" pageNumber="143" rank="subFamily" subFamily="Coloboderinae">Coloboderinae</taxonomicName>
has not been used as valid after 1899 to our knowledge, we believe that usage of
<taxonomicName lsidName="" pageId="142" pageNumber="143" rank="subFamily" subFamily="Anchytarsinae">Anchytarsinae</taxonomicName>
Champion, 1897 should be conserved and an application be submitted to the Commission to suppress the older name.
</paragraph>
<paragraph pageId="142" pageNumber="143">
<taxonomicName genus="Epilichadinae" lsidName="Epilichadinae" pageId="142" pageNumber="143" rank="genus">*Epilichadinae</taxonomicName>
Lawrence and Stribling, 1992: 19 [stem: Epilichad-]. Type genus:
<taxonomicName class="Insecta" family="Ptilodactylidae" genus="Epilichas" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Epilichas" order="Coleoptera" pageId="142" pageNumber="143" phylum="Arthropoda" rank="genus">Epilichas</taxonomicName>
A. White, 1859. Comment: although Ivie (2002: 136) treated this name as available we do not believe that requirements of Art. 13.1 were met and we therefore treat this name here as unavailable.
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135
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<mods:start id="08749F5B2915DBD4CE784844B5256EC7">111</mods:start>
<mods:end id="0B17391959765AB3DA03645F610AB684">184</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="61D703A3205405A4959579E2B5DA589D">book chapter</mods:classification>
<mods:identifier id="272FEBA8BBFE061040EB9B8BC9C9CF6A" type="ISBN">0-8018-8221-4</mods:identifier>
<mods:identifier id="38A847606E334C68DDBAE99C26F2B29E" type="DOI">10.5281/zenodo.7316519</mods:identifier>
</mods:mods>
<treatment id="DBD3CB1C5A9719F2BF6AAF61A59936EA" ID-DOI="http://doi.org/10.5281/zenodo.11338801" ID-GBIF-Taxon="231542360" ID-Zenodo-Dep="11338801" LSID="urn:lsid:plazi:treatment:DBD3CB1C5A9719F2BF6AAF61A59936EA" httpUri="http://treatment.plazi.org/id/DBD3CB1C5A9719F2BF6AAF61A59936EA" lastPageNumber="164" pageNumber="164" sourceUrl="http://www.departments.bucknell.edu/biology/resources/msw3/browse.asp?id=12100565">
<subSubSection id="BED51B76C0E3F79F8ABAF12939F9591E" pageNumber="164" type="nomenclature">
<paragraph id="4F47CB394C2F2737BA9AEB9B5A24BE84" pageNumber="164">
<taxonomicName id="03A42AD201D53551AEE805A275860653" ID-CoL="3WWP8" authority="(Linnaeus 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Mammalia" family="Cercopithecidae" genus="Macaca" kingdom="Animalia" order="Primates" pageNumber="164" phylum="Chordata" rank="species" species="sylvanus">
<emphasis id="248CD8E200B47C5982EC1226FC53437C" italics="true" pageNumber="164">Macaca sylvanus</emphasis>
(Linnaeus 1758)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="88B15DE2B6CF601D8BCE9FC62BE48DB1" pageNumber="164" type="reference_group">
<paragraph id="A4A92BCF3539BC870723EE2700F013FF" pageNumber="164">
<treatmentCitation id="825DC0B039F8A15C6071161AA8143E09" ID-DOI="http://doi.org/10.5281/zenodo.3913052" ID-Zenodo-Dep="3913052" LSID="urn:lsid:zoobank.org:act:C1FC5985-3C54-41F8-8F13-BA4F9BC1E881" author="Linnaeus" authority="Linnaeus 1758" authorityName="Linnaeus" authorityYear="1758" class="Mammalia" family="Cercopithecidae" genus="Simia" httpUri="http://treatment.plazi.org/id/C2A86FC2FE9E978E58BF6DFBC4C916DE" kingdom="Animalia" order="Primates" page="25" pageNumber="164" phylum="Chordata" rank="species" species="sylvanus" title="Simia sylvanus" volumeTitle="Syst. Nat., 10th ed., Vol. 1" year="1758">
<taxonomicName id="915EB7E2070FE74311A56FDCCA6179FA" ID-CoL="7QQD" authority="Linnaeus 1758" authorityName="Linnaeus" authorityYear="1758" class="Mammalia" family="Cercopithecidae" genus="Simia" kingdom="Animalia" order="Primates" pageNumber="164" phylum="Chordata" rank="species" species="sylvanus">
<emphasis id="5854056E6FB7821E40FFE92BF3896901" italics="true" pageNumber="164">[Simia] sylvanus</emphasis>
<bibRefCitation id="03E9CDED6A5EA78D01E3AF33B4C1442C" author="Linnaeus, C." refId="ref213949" refString="Linnaeus, C. 1758. Systema Naturae per regna tria naturae, secundum classis, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Tenth ed. Vol. 1. Laurentii Salvii, Stockholm, 824 pp." year="1758">Linnaeus 1758</bibRefCitation>
</taxonomicName>
,
<bibCitation id="9CEFA0EEE3F3B7971C7CE27B97E8158F" author="Linnaeus" pageNumber="164" pagination="25" title="Simia sylvanus" volumeTitle="Syst. Nat., 10th ed., Vol. 1" year="1758">Syst. Nat., 10th ed., Vol. 1: 25</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CEBF7460E74F2669965D1BD8A4D780C2" pageNumber="164" type="materials_examined">
<paragraph id="D27728A85C81A01D0350E444EB9B5707" pageNumber="164">
<emphasis id="2DD2462DFE1860FBEE72B4C705D60C21" inLineHeading="true" pageNumber="164">Type Locality:</emphasis>
<materialsCitation id="636DDA9E0CB270D374013C20E85F4893" ID-GBIF-Occurrence="4867331378" pageNumber="164">North Africa, &quot;Barbary coast&quot;.</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="1E63FBA6113A7D32C54C49524A27DCAC" pageNumber="164" type="vernacular_names">
<paragraph id="57C33C53A8A5B3B44F2A1B73889D3184" pageNumber="164">
<emphasis id="103E7602D8CE807A30545C28A153218E" inLineHeading="true" pageNumber="164">Vernacular Names:</emphasis>
<vernacularName id="7085B90A004DD9ED296971B94EEAFB95" language="eng" pageNumber="164">Barbary Macaque</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="2342D48C0D67F05AF47FE4A2558E343F" pageNumber="164" type="synonym_list">
<paragraph id="83B6A80D4A8FAD10FE764C139175F36C" pageNumber="164">
<emphasis id="CC3F5CC9B8F66A0BD08793C768283C11" inLineHeading="true" pageNumber="164">Synonyms:</emphasis>
<taxonomicName id="3F5A6F25AAAD9FCEA277E6DC02EA38CB" authority="(É. Geoffroy 1812)" baseAuthorityName="É. Geoffroy" baseAuthorityYear="1812" class="Mammalia" family="Cercopithecidae" genus="Macaca" kingdom="Animalia" order="Primates" pageNumber="164" phylum="Chordata" rank="species" species="ecaudatus">
<emphasis id="AE94F20CD41A98A90427AC0F5C94058A" italics="true" pageNumber="164">Macaca ecaudatus</emphasis>
(É. Geoffroy 1812)
</taxonomicName>
;
<taxonomicName id="89C1B6F8A5D1678261A3544A79A82D1B" authority="(Linnaeus 1766)" baseAuthorityName="Linnaeus" baseAuthorityYear="1766" class="Mammalia" family="Cercopithecidae" genus="Macaca" kingdom="Animalia" order="Primates" pageNumber="164" phylum="Chordata" rank="species" species="inuus">
<emphasis id="639EE18310CE56FC5D5780CF231EAA96" italics="true" pageNumber="164">Macaca inuus</emphasis>
(
<bibRefCitation id="A54BBB4C2D9421BC2947EC5271BCB1D6" author="Linnaeus, C." refId="ref213994" refString="Linnaeus, C. 1766 - 1768. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis synonymis, locis. Vol. 1. Regnum Animale. pt. 1, pp. 1 - 532 [1766]; vol. 3, Appendix pp. 223 - 235 [1768]. 12 th ed. [Vol. 3, Regnum Lapideum, pp. 1 - 222, rejected for nomenclatural purposes, ICZN, opinion 296]. Laurentii Salvii, Stockholm, 3 vols." year="1766">Linnaeus 1766</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="3D92DFB3D7DAB779871145D9E3586F20" authority="(Schreber 1799)" baseAuthorityName="Schreber" baseAuthorityYear="1799" class="Mammalia" family="Cercopithecidae" genus="Macaca" kingdom="Animalia" order="Primates" pageNumber="164" phylum="Chordata" rank="species" species="pithecus">
<emphasis id="28DB07B8F38409C65495F25C5442E88A" italics="true" pageNumber="164">Macaca pithecus</emphasis>
(Schreber 1799)
</taxonomicName>
;
<taxonomicName id="EE1385A2955578F1BB9A634CCC86D2AD" authority="(Reichenbach 1863)" baseAuthorityName="Reichenbach" baseAuthorityYear="1863" class="Mammalia" family="Cercopithecidae" genus="Macaca" kingdom="Animalia" order="Primates" pageNumber="164" phylum="Chordata" rank="species" species="pygmaeus">
<emphasis id="ADDE94809B987FB901297BE82DA04102" italics="true" pageNumber="164">Macaca pygmaeus</emphasis>
(Reichenbach 1863)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="D522FF25D9C5A75CDD32FEFCC26324CB" pageNumber="164" type="distribution">
<paragraph id="7AFCF267DC31E3EEEE71A80A01A34287" pageNumber="164">
<emphasis id="272C3910346C5AD74EE0B74DCD8F42E4" inLineHeading="true" pageNumber="164">Distribution:</emphasis>
<collectingCountry id="A25068BEB227303A9D5CD465CC3C3FD2" name="Morocco">Morocco</collectingCountry>
,
<collectingCountry id="601466400331BA34945E65A06B2A01EA" name="Algeria">Algeria</collectingCountry>
,
<collectingCountry id="8CBE19ADB88A27438695C889A612F772" name="Gibraltar">Gibraltar</collectingCountry>
(introduced).
</paragraph>
</subSubSection>
<subSubSection id="A9C89687867E90011B3101020ABC9614" pageNumber="164" type="conservation">
<paragraph id="F11CE568F4A8C9D84F5C3F336E90BD4C" pageNumber="164">
<emphasis id="E84ABDDC0660CAB3ABE6D1AAD34BAE6B" inLineHeading="true" pageNumber="164">Conservation:</emphasis>
<collectionCode id="B5FAE40EB2F1C0D116B0AEC799DC2588">CITES</collectionCode>
– Appendix II;
<collectionCode id="719C58D81DE920E32984277B27F7BE2E">IUCN</collectionCode>
– Vulnerable.
</paragraph>
</subSubSection>
<subSubSection id="BD9E4A66BCF4DF6EA2051369484B5324" pageNumber="164" type="discussion">
<paragraph id="00005D02B26FC026191F4C20A329363B" pageNumber="164">
<emphasis id="DB0C69C6519FA551FC4CCB746CD36AB5" inLineHeading="true" pageNumber="164">Discussion:</emphasis>
<taxonomicName id="5F2460B012218FBF9C94DDE7DF92295E" authorityName="Linnaeus" authorityYear="1758" class="Mammalia" family="Cercopithecidae" genus="Simia" kingdom="Animalia" order="Primates" phylum="Chordata" rank="species" species="sylvanus">
<emphasis id="599EC771706A40B1E2D8EB22A3D39A1F" italics="true" pageNumber="164">M. sylvanus</emphasis>
</taxonomicName>
species group. See
<bibRefCitation id="DA83E744083A95053AD3C35BA2BEE68F" author="Fooden, J." refId="ref106390" refString="Fooden, J. 1976. Provisional classification and key to living species of macaques. Folia Primatologia, 25: 225 - 236." year="1976">Fooden (1976:226)</bibRefCitation>
for the use of this name.
</paragraph>
</subSubSection>
</treatment>
</document>