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<document ID-DOI="http://dx.doi.org/10.3897/BDJ.12.e99555" ID-Pensoft-Pub="1314-2828-12-e99555" ID-Pensoft-UUID="767A4AD8287A5FE99D4806177D4BACF0" ModsDocID="1314-2828-12-e99555" checkinTime="1706571555167" checkinUser="pensoft" docAuthor="Jimoh, Bunmi Omowumi, Gomez, Kiko, Kemabonta, Kehinde Abike, Wakanjuola, Winifred Ayinke, Phiri, Ethel Emmarantia &amp; Mothapo, Palesa Natasha" docDate="2024" docId="DB6B48BB04FF5D1382D644B8B66C78F9" docLanguage="en" docName="BiodivDatJour 12: e99555" docOrigin="Biodiversity Data Journal 12" docPubDate="2024-01-29" docSource="http://dx.doi.org/10.3897/BDJ.12.e99555" docTitle="Crematogaster africana subsp. alligatrix Forel 1911" docType="treatment" docVersion="1" id="767A4AD8287A5FE99D4806177D4BACF0" lastPageNumber="99555" masterDocId="767A4AD8287A5FE99D4806177D4BACF0" masterDocTitle="A checklist of Nigerian ants (Hymenoptera, Formicidae): a review, new records and exotic species" masterLastPageNumber="99555" masterPageNumber="99555" pageNumber="99555" updateTime="1706571858046" updateUser="ExternalLinkService">
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<mods:title id="9A312725E5944AD116ECBBBFF15B576F">A checklist of Nigerian ants (Hymenoptera, Formicidae): a review, new records and exotic species</mods:title>
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<mods:namePart id="BDD7CCF46072587CC6BDCD8057746400">Jimoh, Bunmi Omowumi</mods:namePart>
<mods:affiliation id="FBBABFEE13AC343D9ECEE2D3FFDCA669">University of Lagos, Lagos, Nigeria</mods:affiliation>
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<mods:namePart id="E1B27407C2715E8418DB12D97AECC50B">Gomez, Kiko</mods:namePart>
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<mods:affiliation id="3C566655C7ECFB5D00FE021F4CBDB8E4">Independent Researcher, Barcelona, Spain</mods:affiliation>
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<mods:namePart id="816914D5782D306A2B1B0D422730DD68">Kemabonta, Kehinde Abike</mods:namePart>
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<mods:affiliation id="0F9A9B848860F04AEA03F39EDEAD0929">University of Lagos, Lagos, Nigeria</mods:affiliation>
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<mods:namePart id="E9B2CD524AF845E1A68380D67DBB8C65">Phiri, Ethel Emmarantia</mods:namePart>
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<mods:namePart id="E7A4C7EDE800B93524ED2DDBD304B0D8">Mothapo, Palesa Natasha</mods:namePart>
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<mods:date id="07CE93BE20DBBE67E91D0C68FA9D7D07">2024</mods:date>
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<taxonomicName id="56DEB6BC4ED7F8F48C9892B59247056A" ID-CoL="5GNJ7" LSID="DB6B48BB-04FF-5D13-82D6-44B8B66C78F9" authority="Forel, 1911" authorityName="Forel" authorityYear="1911" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName="Crematogaster africana subsp. alligatrix" order="Hymenoptera" pageId="0" pageNumber="99555" phylum="Arthropoda" rank="subSpecies" species="africana" subSpecies="alligatrix">Crematogaster africana alligatrix Forel, 1911</taxonomicName>
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<paragraph id="0D8D2F7556930C895A4E634CED029112" pageId="0" pageNumber="99555">Notes</paragraph>
<paragraph id="379A19C9CACD200D0D84B5BEFE83DE41" pageId="0" pageNumber="99555">
(
<bibRefCitation id="C3AFD6F0DE255D2D16C74ABCC5E95FFD" author="Forel, A." journalOrPublisher="Revue Zoologique Africaine (Brussels)" pageId="0" pageNumber="99555" pagination="352 - 353" refId="B7735049" refString="Forel, A., 1913. Fourmis de Nigerie. Revue Zoologique Africaine (Brussels) 2: 352 - 353" title="Fourmis de Nigerie" volume="2" year="1913">Forel 1913</bibRefCitation>
,
<bibRefCitation id="177370711C2BDDBE3F41C49418295AE6" author="Wheeler, W. M." journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="0" pageNumber="99555" pagination="711 - 1004" refId="B7741538" refString="Wheeler, W. M., 1922. Ants of the American Museum Congo expedition. A contribution to the myrmecology of Africa. VIII. A synonymic list of the ants of the Ethiopian region. Bulletin of the American Museum of Natural History 45: 711 - 1004" title="Ants of the American Museum Congo expedition. A contribution to the myrmecology of Africa. VIII. A synonymic list of the ants of the Ethiopian region" volume="45" year="1922">Wheeler 1922</bibRefCitation>
,
<bibRefCitation id="F66E576C5A4A45D212F0A78B2B218BD2" author="Soulie, J." journalOrPublisher="Annales de l'Universite d'Abidjan. Serie Sciences." pageId="0" pageNumber="99555" pagination="85 - 106" refId="B7740544" refString="Soulie, J., Dicko, L. D., 1965. La repartition des genres de fourmis de la tribu des &quot; Cremastogastrini &quot; dans la faune ethiopienne et malgache. Hymenoptera - Formicoidea - Myrmicidae. Annales de l'Universite d'Abidjan. Serie Sciences. 1: 85 - 106" title="La repartition des genres de fourmis de la tribu des &quot; Cremastogastrini &quot; dans la faune ethiopienne et malgache. Hymenoptera - Formicoidea - Myrmicidae" volume="1" year="1965">
<normalizedToken id="8785D5F2AEEC3067D57C57791C87EAF1" originalValue="Soulié">Soulie</normalizedToken>
and Dicko 1965
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)
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<heading id="083581E8FF80713DFEFD8FB7FD44EDCB" bold="true" box="[264,714,1444,1470]" fontSize="11" level="1" pageId="37" pageNumber="38" reason="1">
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<taxonomicName id="94C24D07FF80713DFEFD8FB7FDD1EDCB" authority="Lelej" authorityName="Lelej" box="[264,607,1444,1470]" class="Insecta" family="Mutillidae" genus="Vanhartenidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="arabica" status="sp. nov.">
<emphasis id="61B6EA96FF80713DFEFD8FB7FD99EDCB" bold="true" box="[264,535,1444,1470]" italics="true" pageId="37" pageNumber="38">Vanhartenidia arabica</emphasis>
Lelej
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,
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(
<figureCitation id="CBF92A01FF80713DFEE48FD4FEE8ED94" box="[273,358,1479,1505]" captionStart="FIGURES 22 – 29. 22 – 23" captionStartId="40.[264,382,1451,1475]" captionTargetBox="[543,1028,890,1425]" captionTargetId="figure@40.[539,1048,889,1428]" captionTargetPageId="40" captionText="FIGURES 22 – 29. 22 – 23. Spinulomutilla yemenita sp. nov., male, holotype. 22. Sterna 6 – 8, ventral view; 23. Penial valves, ventral view. 24. Vanhartenidia arabica sp. nov., female, holotype. Foretarsus. 25 – 29. V. felix sp. nov., paratype (25 – 28, male; 29, female). 25. Hind coxa, ventral view; 26. Genitalia, dorsal view; 27. Volsella, dorsal view; 28. Penial valves, ventral view; 29. Pygidial area. Scale = 1 mm for 22, 0.5 mm for 23 and 26, and 0.25 mm for 27 and 28." httpUri="https://zenodo.org/record/172675/files/figure.png" pageId="37" pageNumber="38">Fig. 24</figureCitation>
)
</paragraph>
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<subSubSection id="1BD8650FFF80713DFEFD8C04FAAAEE2C" box="[264,1316,1559,1625]" pageId="37" pageNumber="38" type="diagnosis">
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<emphasis id="61B6EA96FF80713DFEFD8C04FEF1EE45" box="[264,383,1559,1584]" italics="true" pageId="37" pageNumber="38">Diagnosis</emphasis>
</paragraph>
<paragraph id="537D3684FF80713DFECD8C2CFAAAEE2C" blockId="37.[264,1316,1559,1625]" box="[312,1316,1599,1625]" pageId="37" pageNumber="38">The differences beyween the new species and related ones are noted in the key below.</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF80713EFEFD8C9CFD9BEBFB" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38" type="description">
<paragraph id="537D3684FF80713DFEFD8C9CFE1CEEDD" blockId="37.[264,1323,1679,1905]" box="[264,402,1679,1704]" pageId="37" pageNumber="38">
<emphasis id="61B6EA96FF80713DFEFD8C9CFE1CEEDD" box="[264,402,1679,1704]" italics="true" pageId="37" pageNumber="38">Description</emphasis>
</paragraph>
<paragraph id="537D3684FF80713DFECD8CA4FAA5EF04" blockId="37.[264,1323,1679,1905]" pageId="37" pageNumber="38">
FEMALE. Body length
<quantity id="943A9B61FF80713DFDA18CA4FD3CEEA4" box="[596,690,1719,1745]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="37" pageNumber="38" unit="mm" value="7.6">7.6 mm</quantity>
. Relation of pedicel and first three flagellomeres 0.4: 1.5: 0.8: 1.2. Ratio eye height / gena height (head in lateral view) 1.7X. Genal carina weak. Mesosoma widened posteriorly, 1.3X longer than its maximal width. Pronotum anteriorly arcuate, with lateral sides divergent posteriorly, slightly wider than mesonotum. Scutellar scale wide, prescutellar, scutellar and postscutellar row of tubercles weak.
</paragraph>
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Propodeum posteriorly reticulate. Foretarsus strongly pectinate. Carina on S1 posteriorly without tubercle. Pygidial area narrower than that of
<taxonomicName id="94C24D07FF83713EFC778B57FC5CE928" box="[898,978,324,349]" class="Insecta" family="Mutillidae" genus="Vanhartenidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="felix">
<emphasis id="61B6EA96FF83713EFC778B57FC5CE928" box="[898,978,324,349]" italics="true" pageId="38" pageNumber="39">V. felix</emphasis>
</taxonomicName>
. Frons and vertex with large deep dense, sometimes confluent punctures. T2 laterally with shallow punctures. Mesosoma dorsally with larger, deeper confluent punctures. Mesopleura, metapleura, and propodeum glabrous.
</paragraph>
<paragraph id="537D3684FF83713EFECD8BF7FDE8EB13" blockId="38.[264,1322,284,910]" pageId="38" pageNumber="39">Head black, reddish beneath. Mesosoma ferruginous­red. Mandible ferruginous­red with black apex. Antenna black. Legs castaneous. Metasoma yellowish­red includes cuticula under black setae. T6 brown. Frons, vertex and mesosomal dorsum with sparse recumbent yellowish setae mixed with erect black setae. Gena, paraocular area, malar space with dense silvery setae. Mesopleura and propleura with spot of silvery setae. T2 with two large basal golden pubescent spots, distance between them less than scutellar scale width. T3–T5 with golden bands. Metasomal terga except golden spots and bands with black setae, felt lines yellowish. T1 and T2 posteriorly with golden fringes. T6 laterally with whitish and black setae. Metasomal sterna with silvery setae, S2–S5 with apical fringes of silvery setae.</paragraph>
<paragraph id="537D3684FF83713EFECD8967FD9BEBFB" blockId="38.[264,1322,284,910]" box="[312,533,884,910]" pageId="38" pageNumber="39">MALE. Unknown.</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF83713EFEFD89D7FC56EC73" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="537D3684FF83713EFEFD89D7FE27EBA8" blockId="38.[264,984,964,1030]" box="[264,425,964,989]" pageId="38" pageNumber="39">
<emphasis id="61B6EA96FF83713EFEFD89D7FE27EBA8" box="[264,425,964,989]" italics="true" pageId="38" pageNumber="39">
<typeStatus id="8C798826FF83713EFEFD89D7FEB1EBA8" box="[264,319,964,989]" pageId="38" pageNumber="39">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="537D3684FF83713EFECD89FFFC56EC73" blockId="38.[264,984,964,1030]" box="[312,984,1003,1030]" pageId="38" pageNumber="39">
<typeStatus id="8C798826FF83713EFECD89FFFE2BEC73" box="[312,421,1004,1030]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
Ψ,
<collectingCountry id="2BD57614FF83713EFE3B89FFFDAAEC73" box="[462,548,1004,1030]" name="Yemen" pageId="38" pageNumber="39">Yemen</collectingCountry>
: Al Kowd,
<date id="277C1044FF83713EFD5F89FFFC93EC73" box="[682,797,1004,1030]" pageId="38" pageNumber="39" value="2001-01-16">16.I.2001</date>
, AvH [
<collectionCode id="35D3AE41FF83713EFC9B89FFFC40EC73" LSID="urn:lsid:biocol.org:col:34975" box="[878,974,1004,1030]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="38" pageNumber="39">ZMAN</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF83713EFEFD8E2FFE03EC0B" box="[264,407,1084,1150]" pageId="38" pageNumber="39" type="distribution">
<paragraph id="537D3684FF83713EFEFD8E2FFE19EC20" blockId="38.[264,407,1084,1150]" box="[264,407,1084,1109]" pageId="38" pageNumber="39">
<emphasis id="61B6EA96FF83713EFEFD8E2FFE19EC20" box="[264,407,1084,1109]" italics="true" pageId="38" pageNumber="39">Distribution</emphasis>
</paragraph>
<paragraph id="537D3684FF83713EFECD8E77FE03EC0B" blockId="38.[264,407,1084,1150]" box="[312,397,1124,1150]" pageId="38" pageNumber="39">
<collectingCountry id="2BD57614FF83713EFECD8E77FE09EC0B" box="[312,391,1124,1150]" name="Yemen" pageId="38" pageNumber="39">Yemen</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF83713EFEFD8EA7FD77ED6B" pageId="38" pageNumber="39" type="etymology">
<paragraph id="537D3684FF83713EFEFD8EA7FE0BECB8" blockId="38.[264,1321,1204,1310]" box="[264,389,1204,1229]" pageId="38" pageNumber="39">
<emphasis id="61B6EA96FF83713EFEFD8EA7FE0BECB8" box="[264,389,1204,1229]" italics="true" pageId="38" pageNumber="39">Etymology</emphasis>
</paragraph>
<paragraph id="537D3684FF83713EFECD8ECFFD77ED6B" blockId="38.[264,1321,1204,1310]" pageId="38" pageNumber="39">
<taxonomicName id="94C24D07FF83713EFECD8ECFFE18EC80" box="[312,406,1244,1269]" class="Insecta" family="Mutillidae" genus="Vanhartenidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="43" phylum="Arthropoda" rank="species" species="arabica" status="sp. nov.">
<emphasis id="61B6EA96FF83713EFECD8ECFFE18EC80" box="[312,406,1244,1269]" italics="true" pageId="38" pageNumber="39">Arabica</emphasis>
</taxonomicName>
is a Latin adjective derived from Arabia, the name of peninsula referring to the area where the species has been found.
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF83713EFEFD8F47FB42EDE3" box="[264,1228,1364,1430]" pageId="38" pageNumber="39" type="discussion">
<paragraph id="537D3684FF83713EFEFD8F47FEE0ED18" blockId="38.[264,1228,1364,1430]" box="[264,366,1364,1389]" pageId="38" pageNumber="39">
<emphasis id="61B6EA96FF83713EFEFD8F47FEE0ED18" box="[264,366,1364,1389]" italics="true" pageId="38" pageNumber="39">Remarks</emphasis>
</paragraph>
<paragraph id="537D3684FF83713EFECD8F6FFB42EDE3" blockId="38.[264,1228,1364,1430]" box="[312,1228,1404,1430]" pageId="38" pageNumber="39">
It is quite possible that
<taxonomicName id="94C24D07FF83713EFDBD8F6FFCDEEDE0" box="[584,848,1404,1429]" class="Insecta" family="Mutillidae" genus="Vanhartenidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="arabica">
<emphasis id="61B6EA96FF83713EFDBD8F6FFCDEEDE0" box="[584,848,1404,1429]" italics="true" pageId="38" pageNumber="39">Vanhartenidia arabica</emphasis>
</taxonomicName>
is the opposite sex of
<taxonomicName id="94C24D07FF83713EFBA28F6FFB4BEDE0" box="[1111,1221,1404,1429]" class="Insecta" family="Mutillidae" genus="Vanhartenidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="tihama">
<emphasis id="61B6EA96FF83713EFBA28F6FFB4BEDE0" box="[1111,1221,1404,1429]" italics="true" pageId="38" pageNumber="39">V. tihama</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

167
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<treatment id="DB6B8792FF837130FEFD8FE7FA91EB4B" ID-DOI="http://doi.org/10.5281/zenodo.6252994" ID-GBIF-Taxon="119344680" ID-Zenodo-Dep="6252994" LSID="urn:lsid:plazi:treatment:DB6B8792FF837130FEFD8FE7FA91EB4B" httpUri="http://treatment.plazi.org/id/DB6B8792FF837130FEFD8FE7FA91EB4B" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39">
<subSubSection id="1BD8650FFF83713EFEFD8FE7FE10EE44" pageId="38" pageNumber="39" type="nomenclature">
<paragraph id="537D3684FF83713EFEFD8FE7FD2AEE7B" blockId="38.[264,676,1524,1585]" box="[264,676,1524,1550]" pageId="38" pageNumber="39">
<heading id="083581E8FF83713EFEFD8FE7FD2AEE7B" bold="true" box="[264,676,1524,1550]" fontSize="11" level="1" pageId="38" pageNumber="39" reason="1">
<emphasis id="61B6EA96FF83713EFEFD8FE7FD2AEE7B" bold="true" box="[264,676,1524,1550]" pageId="38" pageNumber="39">
<taxonomicName id="94C24D07FF83713EFEFD8FE7FDB4EE7B" authority="Lelej" authorityName="Lelej" box="[264,570,1524,1550]" class="Insecta" family="Mutillidae" genus="Vanhartenidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="felix" status="sp. nov.">
<emphasis id="61B6EA96FF83713EFEFD8FE7FE7CEE7B" bold="true" box="[264,498,1524,1550]" italics="true" pageId="38" pageNumber="39">Vanhartenidia felix</emphasis>
Lelej
</taxonomicName>
,
<taxonomicNameLabel id="7A8557EDFF83713EFDB28FE7FD2AEE7B" box="[583,676,1524,1550]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="537D3684FF83713EFEFD8C04FE10EE44" blockId="38.[264,676,1524,1585]" box="[264,414,1559,1585]" pageId="38" pageNumber="39">
(
<figureCitation id="CBF92A01FF83713EFEE58C04FE18EE44" box="[272,406,1559,1585]" captionStart="FIGURES 22 – 29. 22 – 23" captionStartId="40.[264,382,1451,1475]" captionTargetBox="[543,1028,890,1425]" captionTargetId="figure@40.[539,1048,889,1428]" captionTargetPageId="40" captionText="FIGURES 22 – 29. 22 – 23. Spinulomutilla yemenita sp. nov., male, holotype. 22. Sterna 6 – 8, ventral view; 23. Penial valves, ventral view. 24. Vanhartenidia arabica sp. nov., female, holotype. Foretarsus. 25 – 29. V. felix sp. nov., paratype (25 – 28, male; 29, female). 25. Hind coxa, ventral view; 26. Genitalia, dorsal view; 27. Volsella, dorsal view; 28. Penial valves, ventral view; 29. Pygidial area. Scale = 1 mm for 22, 0.5 mm for 23 and 26, and 0.25 mm for 27 and 28." httpUri="https://zenodo.org/record/172675/files/figure.png" pageId="38" pageNumber="39">Figs 25–29</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF83713EFEFD8C74FA90EEDC" box="[264,1310,1639,1705]" pageId="38" pageNumber="39" type="diagnosis">
<paragraph id="537D3684FF83713EFEFD8C74FEF1EEF5" blockId="38.[264,1310,1639,1705]" box="[264,383,1639,1664]" pageId="38" pageNumber="39">
<emphasis id="61B6EA96FF83713EFEFD8C74FEF1EEF5" box="[264,383,1639,1664]" italics="true" pageId="38" pageNumber="39">Diagnosis</emphasis>
</paragraph>
<paragraph id="537D3684FF83713EFECD8C9CFA90EEDC" blockId="38.[264,1310,1639,1705]" box="[312,1310,1679,1705]" pageId="38" pageNumber="39">The differences between the new species and related ones are noted in the key below.</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF83713FFEFD8CCCFC2CEEDB" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39" type="description">
<paragraph id="537D3684FF83713EFEFD8CCCFE1CEE8D" blockId="38.[264,1323,1759,1905]" box="[264,402,1759,1784]" pageId="38" pageNumber="39">
<emphasis id="61B6EA96FF83713EFEFD8CCCFE1CEE8D" box="[264,402,1759,1784]" italics="true" pageId="38" pageNumber="39">Description</emphasis>
</paragraph>
<paragraph id="537D3684FF83713FFECD8D14FE5AE98B" blockId="38.[264,1323,1759,1905]" lastBlockId="39.[264,1324,284,1870]" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
MALE. Body length 10.5–12.0 mm. Eyes projecting over head. POD: OOD 0.9X. Relation of pedicel and first three flagellomeres 0.8: 1.9: 2.0: 2.1. Clypeus with deep fovea on median elevated part delimited laterally by a carina. Mid coxa with lateral longitudinal carina. Parascutal carinae developed.
<collectingCountry id="2BD57614FF82713FFC858B0FFC25E943" box="[880,939,284,310]" name="India" pageId="39" pageNumber="40">Hind</collectingCountry>
coxa inside with carina which ends in weak tubercle. Carina on S1 posteriorly with additional high carina. Genitalia as in
<figureCitation id="CBF92A01FF82713FFEFD8B7FFEECE9F3" box="[264,354,364,390]" captionStart="FIGURES 22 – 29. 22 – 23" captionStartId="40.[264,382,1451,1475]" captionTargetBox="[543,1028,890,1425]" captionTargetId="figure@40.[539,1048,889,1428]" captionTargetPageId="40" captionText="FIGURES 22 – 29. 22 – 23. Spinulomutilla yemenita sp. nov., male, holotype. 22. Sterna 6 – 8, ventral view; 23. Penial valves, ventral view. 24. Vanhartenidia arabica sp. nov., female, holotype. Foretarsus. 25 – 29. V. felix sp. nov., paratype (25 – 28, male; 29, female). 25. Hind coxa, ventral view; 26. Genitalia, dorsal view; 27. Volsella, dorsal view; 28. Penial valves, ventral view; 29. Pygidial area. Scale = 1 mm for 22, 0.5 mm for 23 and 26, and 0.25 mm for 27 and 28." httpUri="https://zenodo.org/record/172675/files/figure.png" pageId="39" pageNumber="40">Fig. 26</figureCitation>
. Head and mesosoma dorsally densely punctured. Propodeum dorsally, laterally, and posteriorly reticulate. T2 and S2 densely punctured, punctures larger on S2 and sparser on disc of T2. T7 with median longitudinal glabrous gibbosity, which ends before abrupt posterior surface.
</paragraph>
<paragraph id="537D3684FF82713FFECD881FFE1DEBFB" blockId="39.[264,1324,284,1870]" pageId="39" pageNumber="40">
Head and mesosoma ferruginous­red, metasoma except last segment yellowish­red. Mandible castaneous, preapically red. Antennae black, palps and legs brown. T7 brown to black. Forewing bright yellow in proximal half and dark fuscous in distal half. Head and mandible with long erect and short recumbent silvery setae denser on paraocular area and lateral part of clypeus. Genae with sparse silvery setae. Dorsum of mesosoma with dense recumbent golden setae hidden the sculpture, sparser on scutellum. Mesopleura and legs with sparse whitish setae. T2 with golden pubescence, sparser on disc, T2 posteriorly and T3–T6 with bands of dense recumbent golden setae. T7 with black setae. Metasomal sterna with sparse silvery setae, S2–S6 with apical fringes of silvery setae. Genitalia as in
<figureCitation id="CBF92A01FF82713FFEFD8967FE01EBFB" box="[264,399,884,910]" captionStart="FIGURES 22 – 29. 22 – 23" captionStartId="40.[264,382,1451,1475]" captionTargetBox="[543,1028,890,1425]" captionTargetId="figure@40.[539,1048,889,1428]" captionTargetPageId="40" captionText="FIGURES 22 – 29. 22 – 23. Spinulomutilla yemenita sp. nov., male, holotype. 22. Sterna 6 – 8, ventral view; 23. Penial valves, ventral view. 24. Vanhartenidia arabica sp. nov., female, holotype. Foretarsus. 25 – 29. V. felix sp. nov., paratype (25 – 28, male; 29, female). 25. Hind coxa, ventral view; 26. Genitalia, dorsal view; 27. Volsella, dorsal view; 28. Penial valves, ventral view; 29. Pygidial area. Scale = 1 mm for 22, 0.5 mm for 23 and 26, and 0.25 mm for 27 and 28." httpUri="https://zenodo.org/record/172675/files/figure.png" pageId="39" pageNumber="40">Figs 26–28</figureCitation>
.
</paragraph>
<paragraph id="537D3684FF82713FFECD898FFC85ED6B" blockId="39.[264,1324,284,1870]" pageId="39" pageNumber="40">
FEMALE. Body length 9.0 mm. Relation of pedicel and first three flagellomeres 0.6: 1.8: 0.8: 1.1. Ratio eye height / gena height (head in lateral view) 1.9. Genal carina weak. Mesosoma 1.3X longer than its maximal width. Pronotum anteriorly arcuate, with parallel lateral sides, slightly wider than mesonotum. Scutellar scale wide, prescutellar, scutellar and postscutellar row of tubercles well developed. Foretarsus strongly pectinate. Propodeum posteriorly longitudinally rugulate. Carina on S1 posteriorly with additional triangular carina. Pygidial area as in
<figureCitation id="CBF92A01FF82713FFD248E9FFCA2ECD3" box="[721,812,1164,1190]" captionStart="FIGURES 22 – 29. 22 – 23" captionStartId="40.[264,382,1451,1475]" captionTargetBox="[543,1028,890,1425]" captionTargetId="figure@40.[539,1048,889,1428]" captionTargetPageId="40" captionText="FIGURES 22 – 29. 22 – 23. Spinulomutilla yemenita sp. nov., male, holotype. 22. Sterna 6 – 8, ventral view; 23. Penial valves, ventral view. 24. Vanhartenidia arabica sp. nov., female, holotype. Foretarsus. 25 – 29. V. felix sp. nov., paratype (25 – 28, male; 29, female). 25. Hind coxa, ventral view; 26. Genitalia, dorsal view; 27. Volsella, dorsal view; 28. Penial valves, ventral view; 29. Pygidial area. Scale = 1 mm for 22, 0.5 mm for 23 and 26, and 0.25 mm for 27 and 28." httpUri="https://zenodo.org/record/172675/files/figure.png" pageId="39" pageNumber="40">Fig. 29</figureCitation>
. Frons and vertex with large deep dense, sometimes confluent punctures. T2 laterally with shallow punctures. Mesosoma dorsally with larger, deeper confluent punctures with tending to approach rugosity. Mesopleura, metapleura, and allied propodeum glabrous.
</paragraph>
<paragraph id="537D3684FF82713FFECD8F3FFC2CEEDB" blockId="39.[264,1324,284,1870]" pageId="39" pageNumber="40">Head and mesosoma ferruginous­red. Mandible ferruginous­red with black apex. Antenna black. Legs castaneous. Metasoma yellowish­red with T1, T5–T6 and T2 black, except for large yellowish­red cuticular spots under golden pubescent spots. Frons, vertex, and mesosomal dorsum with sparse recumbent brown setae mixed with erect black setae. Head below with silvery, legs with yellowish setae. Mesosoma laterally with dense silvery micropubescence. T2 with two large basal golden spots, distance between them slightly more than scutellar scale width. T3–T4 with bands of golden setae. Metasomal terga except golden spots and bands with black setae, felt lines yellowish. T1 and T2 with apical fringes of sparse black setae. T6 laterally with yellowish setae. Metasomal sterna with whitish setae, S2–S5 with apical fringes of whitish setae.</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF827130FEFD8CAFFC07E92B" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="materials_examined">
<paragraph id="537D3684FF82713FFEFD8CAFFE27EEA0" blockId="39.[264,1324,284,1870]" box="[264,425,1724,1749]" pageId="39" pageNumber="40">
<emphasis id="61B6EA96FF82713FFEFD8CAFFE27EEA0" box="[264,425,1724,1749]" italics="true" pageId="39" pageNumber="40">
<typeStatus id="8C798826FF82713FFEFD8CAFFEB1EEA0" box="[264,319,1724,1749]" pageId="39" pageNumber="40">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="537D3684FF827130FECD8CF7FC07E92B" blockId="39.[264,1324,284,1870]" lastBlockId="40.[264,1323,283,350]" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
<typeStatus id="8C798826FF82713FFECD8CF7FE2BEE8B" box="[312,421,1764,1790]" pageId="39" pageNumber="40" type="holotype">Holotype</typeStatus>
ɗ,
<collectingCountry id="2BD57614FF82713FFE258CF7FDA8EE8B" box="[464,550,1764,1790]" name="Yemen" pageId="39" pageNumber="40">Yemen</collectingCountry>
:
<quantity id="943A9B61FF82713FFDC78CF7FDF3EE8B" box="[562,637,1764,1790]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="39" pageNumber="40" unit="km" value="12.0">12 km</quantity>
NW Manakhah,
<date id="277C1044FF82713FFCBF8CF7FBBCEE8B" box="[842,1074,1764,1790]" pageId="39" pageNumber="40" value="2001-07-03" valueMax="2001-08-21" valueMin="2001-07-03">3.VII–21.VIII.2001</date>
, MT, AvH [
<collectionCode id="35D3AE41FF82713FFB4B8CF7FA91EE8B" LSID="urn:lsid:biocol.org:col:34992" box="[1214,1311,1764,1790]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="39" pageNumber="40">RMNH</collectionCode>
].
<typeStatus id="8C798826FF82713FFEFD8D1FFEF0EF53" box="[264,382,1804,1830]" pageId="39" pageNumber="40" type="paratype">Paratypes</typeStatus>
. 5 ɗ, 1 Ψ with the same label as
<typeStatus id="8C798826FF82713FFCDF8D1FFC1EEF53" box="[810,912,1804,1830]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
[
<collectionCode id="35D3AE41FF82713FFC568D1FFB8FEF53" LSID="urn:lsid:biocol.org:col:34992" box="[931,1025,1804,1830]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="39" pageNumber="40">RMNH</collectionCode>
,
<collectionCode id="35D3AE41FF82713FFBE78D1FFBD8EF53" box="[1042,1110,1804,1830]" pageId="39" pageNumber="40">IBSS</collectionCode>
]; the same place,
<date id="277C1044FF82713FFEFD8D27FE56EF3B" box="[264,472,1844,1870]" pageId="39" pageNumber="40" value="2003-05-15" valueMax="2003-06-23" valueMin="2003-05-15">15.V–23.VI.2003</date>
, MT, AvH, 2 ɗ [
<collectionCode id="35D3AE41FF82713FFD498D27FC94EF3B" LSID="urn:lsid:biocol.org:col:34992" box="[700,794,1844,1870]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="39" pageNumber="40">RMNH</collectionCode>
,
<collectionCode id="35D3AE41FF82713FFCD88D27FCFEEF3B" box="[813,880,1844,1870]" pageId="39" pageNumber="40">IBSS</collectionCode>
];
<date id="277C1044FF82713FFC7E8D27FBD2EF3B" box="[907,1116,1844,1870]" pageId="39" pageNumber="40" value="2003-09-15" valueMax="2003-10-22" valueMin="2003-09-15">15.IX–22.X.2003</date>
, MT, AvH, 2 ɗ [
<collectionCode id="35D3AE41FF8D7130FEFA8B0FFEFEE943" LSID="urn:lsid:biocol.org:col:34992" box="[271,368,284,310]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="40" pageNumber="41">RMNH</collectionCode>
]; Al Lahima,
<date id="277C1044FF8D7130FDDF8B0FFC8AE943" box="[554,772,284,310]" pageId="40" pageNumber="41" value="2000-10-16" valueMax="2000-12-31" valueMin="2000-10-16">16.X–31.XII.2000</date>
, MT, AvH &amp; A.M. Hager, 1 ɗ [
<collectionCode id="35D3AE41FF8D7130FB358B0FFA91E943" LSID="urn:lsid:biocol.org:col:34975" box="[1216,1311,284,310]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="40" pageNumber="41">ZMAN</collectionCode>
];
<date id="277C1044FF8D7130FEFD8B57FE22E92B" box="[264,428,324,350]" pageId="40" pageNumber="41" value="2001-01-01" valueMax="2001-04-09" valueMin="2001-01-01">1.I–9.IV.2001</date>
, MT, AvH &amp; A.M. Hager, 1 ɗ [
<collectionCode id="35D3AE41FF8D7130FCEA8B57FCF1E92B" LSID="urn:lsid:biocol.org:col:34975" box="[799,895,324,350]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="40" pageNumber="41">ZMAN</collectionCode>
].
</paragraph>
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<emphasis id="61B6EA96FF8D7130FEFD8B87FE19E9D8" box="[264,407,404,429]" italics="true" pageId="40" pageNumber="41">Distribution</emphasis>
</paragraph>
<paragraph id="537D3684FF8D7130FECD8BAFFE03E9A3" blockId="40.[264,407,404,470]" box="[312,397,444,470]" pageId="40" pageNumber="41">
<collectingCountry id="2BD57614FF8D7130FECD8BAFFE09E9A3" box="[312,391,444,470]" name="Yemen" pageId="40" pageNumber="41">Yemen</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF8D7130FEFD881FFB05EA03" pageId="40" pageNumber="41" type="etymology">
<paragraph id="537D3684FF8D7130FEFD881FFE0BEA50" blockId="40.[264,1323,524,630]" box="[264,389,524,549]" pageId="40" pageNumber="41">
<emphasis id="61B6EA96FF8D7130FEFD881FFE0BEA50" box="[264,389,524,549]" italics="true" pageId="40" pageNumber="41">Etymology</emphasis>
</paragraph>
<paragraph id="537D3684FF8D7130FECD8827FB05EA03" blockId="40.[264,1323,524,630]" pageId="40" pageNumber="41">
The specific name is a Latin adjective derived from &quot;Arabia
<taxonomicName id="94C24D07FF8D7130FBEB8827FBD0EA3B" box="[1054,1118,564,590]" class="Insecta" family="Mutillidae" genus="Vanhartenidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="43" phylum="Arthropoda" rank="species" species="felix" status="sp. nov.">Felix</taxonomicName>
&quot; (Happy Arabia, currently
<collectingCountry id="2BD57614FF8D7130FE8C884FFE5EEA03" box="[377,464,604,630]" name="Yemen" pageId="40" pageNumber="41">Yemen</collectingCountry>
), referring to the country where the species has been found.
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF8D7130FEFD88BFFA91EB4B" pageId="40" pageNumber="41" type="discussion">
<paragraph id="537D3684FF8D7130FEFD88BFFEE0EAB0" blockId="40.[264,1323,684,830]" box="[264,366,684,709]" pageId="40" pageNumber="41">
<emphasis id="61B6EA96FF8D7130FEFD88BFFEE0EAB0" box="[264,366,684,709]" italics="true" pageId="40" pageNumber="41">Remarks</emphasis>
</paragraph>
<paragraph id="537D3684FF8D7130FECD88C7FA91EB4B" blockId="40.[264,1323,684,830]" pageId="40" pageNumber="41">
The collecting of six males and one female by a Malaise trap at the same time permits us to predict that one pair was probably caught
<emphasis id="61B6EA96FF8D7130FC9188EFFC59EB60" box="[868,983,764,789]" italics="true" pageId="40" pageNumber="41">in copula</emphasis>
. Furthermore the male and female have similarly shaped carina on S1, similar silvery spots on gena and mesopleura.
</paragraph>
</subSubSection>
</treatment>
</document>

110
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*
<taxonomicName id="94C24D07FF857138FEE28B0FFD40E940" authority="Bischoff 1920" authorityName="Bischoff" authorityYear="1920" box="[279,718,283,310]" class="Insecta" family="Mutillidae" genus="Dentotilla" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="auricoma">
<emphasis id="61B6EA96FF857138FEE28B0FFD84E943" bold="true" box="[279,522,284,310]" italics="true" pageId="32" pageNumber="33">Dentotilla auricoma</emphasis>
(Bischoff 1920)
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<emphasis id="61B6EA96FF857138FEFD8B74FE7AE90B" box="[264,500,359,382]" italics="true" pageId="32" pageNumber="33">Trogaspidia auricoma</emphasis>
Bischoff 1920: 428
</taxonomicName>
, ♂,
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, &quot;
<collectingCountry id="2BD57614FF857138FC9D8B75FC3BE908" box="[872,949,358,381]" name="Eritrea" pageId="32" pageNumber="33">Eritrea</collectingCountry>
, Ghinda&quot;.
<emphasis id="61B6EA96FF857138FEFD8B94FE6FE9EB" box="[264,481,391,414]" italics="true" pageId="32" pageNumber="33">
Dentotilla
<taxonomicName id="94C24D07FF857138FE8E8B94FE6FE9EB" authority="Nonveiller 1977: 276" authorityName="Nonveiller" authorityPageNumber="276" authorityYear="1977" box="[379,481,391,414]" class="Insecta" family="Mutillidae" genus="Trogaspidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="auricoma">auricoma</taxonomicName>
</emphasis>
:
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<bibRefCitation id="37534B75FF857138FE1A8B95FD2EE9E8" author="Nonveiller" box="[495,672,390,413]" pageId="32" pageNumber="49" refString="Nonveiller, G. (1977) Recherches sur les Mutillides de l'Afrique (Hymenoptera, Mutillidae). VII. Dentotilla gen. nov. (dΨ). Beitrage zur Entomologie, Berlin, 27 (2), 245 - 300." type="journal article" year="1977">Nonveiller 1977</bibRefCitation>
: 276
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, ♂.
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<emphasis id="61B6EA96FF857138FEFD8BDCFE68E99D" box="[264,486,463,488]" italics="true" pageId="32" pageNumber="33">Material examined</emphasis>
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<paragraph id="537D3684FF857138FECD8BE4FCD8EA4C" blockId="32.[264,1323,463,569]" pageId="32" pageNumber="33">
<collectingCountry id="2BD57614FF857138FECD8BE4FE00EA64" box="[312,398,503,529]" name="Yemen" pageId="32" pageNumber="33">Yemen</collectingCountry>
: Al Lahima,
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, MT, AvH &amp; A.H. Hager, 1 ɗ [
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<emphasis id="61B6EA96FF857138FEFD887CFE19EAFD" box="[264,407,623,648]" italics="true" pageId="32" pageNumber="33">Distribution</emphasis>
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<collectingCountry id="2BD57614FF857138FECD8884FE04EAC4" box="[312,394,663,689]" name="Yemen" pageId="32" pageNumber="33">Yemen</collectingCountry>
(new record),
<collectingCountry id="2BD57614FF857138FDC38884FD08EAC4" box="[566,646,663,689]" name="Eritrea" pageId="32" pageNumber="33">Eritrea</collectingCountry>
(
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148
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<heading id="083581E8FF857138FEFD8EA2FD6AECBE" bold="true" box="[264,740,1201,1227]" fontSize="11" level="1" pageId="32" pageNumber="33" reason="1">
<emphasis id="61B6EA96FF857138FEFD8EA2FD6AECBE" bold="true" box="[264,740,1201,1227]" pageId="32" pageNumber="33">
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<emphasis id="61B6EA96FF857138FEFD8EA2FDBFECBE" bold="true" box="[264,561,1201,1227]" italics="true" pageId="32" pageNumber="33">Spinulomutilla yemenita</emphasis>
Lelej
</taxonomicName>
,
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</emphasis>
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(
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)
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF857138FEFD8F37FAAFEE23" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="537D3684FF857138FEFD8F37FEF1ED48" blockId="32.[264,1323,1316,1622]" box="[264,383,1316,1341]" pageId="32" pageNumber="33">
<emphasis id="61B6EA96FF857138FEFD8F37FEF1ED48" box="[264,383,1316,1341]" italics="true" pageId="32" pageNumber="33">Diagnosis</emphasis>
</paragraph>
<paragraph id="537D3684FF857138FECD8F5FFAAFEE23" blockId="32.[264,1323,1316,1622]" pageId="32" pageNumber="33">
The male of this new species has weak carinae on S7 and hypopygium and resembles that of
<taxonomicName id="94C24D07FF857138FE948F67FCE5EDFB" authority="Nonveiller 1994" authorityName="Nonveiller" authorityYear="1994" box="[353,875,1396,1422]" class="Insecta" family="Mutillidae" genus="Spinulomutilla" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="flavogastra">
<emphasis id="61B6EA96FF857138FE948F67FD10EDF8" box="[353,670,1396,1421]" italics="true" pageId="32" pageNumber="33">Spinulomutilla flavogastra</emphasis>
Nonveiller 1994
</taxonomicName>
described from
<collectingCountry id="2BD57614FF857138FBC68F67FB3FEDFB" box="[1075,1201,1396,1422]" name="Cameroon" pageId="32" pageNumber="33">Cameroon</collectingCountry>
. It can be separated from the latter species in having less asymmetrical genitalia, golden pubescence on terga inverted medially (reverted back in
<taxonomicName id="94C24D07FF857138FCC08FD7FC53EDA8" box="[821,989,1476,1501]" class="Insecta" family="Mutillidae" genus="Spinulomutilla" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="flavogastra">
<emphasis id="61B6EA96FF857138FCC08FD7FC53EDA8" box="[821,989,1476,1501]" italics="true" pageId="32" pageNumber="33">S. flavogastra</emphasis>
</taxonomicName>
). The male of this species resembles that of
<taxonomicName id="94C24D07FF857138FE2F8FFFFC82EE73" authority="Magretti 1906" authorityName="Magretti" authorityYear="1906" box="[474,780,1516,1542]" class="Insecta" family="Mutillidae" genus="Spinulomutilla" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="sennae">
<emphasis id="61B6EA96FF857138FE2F8FFFFDC4EE70" box="[474,586,1516,1541]" italics="true" pageId="32" pageNumber="33">S. sennae</emphasis>
(Magretti 1906)
</taxonomicName>
from
<collectingCountry id="2BD57614FF857138FCA08FFFFC2BEE73" box="[853,933,1516,1542]" name="Eritrea" pageId="32" pageNumber="33">Eritrea</collectingCountry>
and
<collectingCountry id="2BD57614FF857138FC158FFFFBCAEE73" box="[992,1092,1516,1542]" name="Ethiopia" pageId="32" pageNumber="33">Ethiopia</collectingCountry>
in having similarly shaped genitalia and similar coloration of metasoma. It can be separated from the latter species by having differently shaped penial valves, and unarmed S6 (armed in
<taxonomicName id="94C24D07FF857138FB578C2FFA9EEE20" box="[1186,1296,1596,1621]" class="Insecta" family="Mutillidae" genus="Spinulomutilla" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="sennae">
<emphasis id="61B6EA96FF857138FB578C2FFA9EEE20" box="[1186,1296,1596,1621]" italics="true" pageId="32" pageNumber="33">S. sennae</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
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<paragraph id="537D3684FF857138FEFD8CA7FE1CEEB8" blockId="32.[264,1325,1716,1902]" box="[264,402,1716,1741]" pageId="32" pageNumber="33">
<emphasis id="61B6EA96FF857138FEFD8CA7FE1CEEB8" box="[264,402,1716,1741]" italics="true" pageId="32" pageNumber="33">Description</emphasis>
</paragraph>
<paragraph id="537D3684FF857139FECD8CCFFDB7EA3B" blockId="32.[264,1325,1716,1902]" lastBlockId="33.[264,1324,284,1030]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
MALE. Body length
<quantity id="943A9B61FF857138FDC78CCFFD1EEE83" box="[562,656,1756,1782]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="32" pageNumber="33" unit="mm" value="6.6">6.6 mm</quantity>
. Head width 1.3X its height. POD: OOD 1.0X. Relation of pedicel and first three flagellomeres 0.6: 1.1: 1.1: 1.3. Clypeus with rounded median deep shiny concavity and narrow preapical setose line bordered laterally by weak tubercles. Mandible bidentate, not excised beneath, without subbasal tooth on inner border. Parascutal carinae not developed.
<collectingCountry id="2BD57614FF847139FCF48B0FFCB2E943" box="[769,828,284,310]" name="India" pageId="33" pageNumber="34">Hind</collectingCountry>
coxa with carina inside which ends in a sharp tubercle. S1 anteriorly with rounded carina. Lateral felt lines located on T2 only. T7 with lateral and apical carinae and median swelled wide glabrous area. S6 not carinate, S7 with weak posterolateral carinae, hypopygium with strongly curved lateral carinae in basal half and weak subapical ones (
<figureCitation id="CBF92A01FF847139FD748BAFFD53E9A3" box="[641,733,444,470]" captionStart="FIGURES 22 – 29. 22 – 23" captionStartId="40.[264,382,1451,1475]" captionTargetBox="[543,1028,890,1425]" captionTargetId="figure@40.[539,1048,889,1428]" captionTargetPageId="40" captionText="FIGURES 22 – 29. 22 – 23. Spinulomutilla yemenita sp. nov., male, holotype. 22. Sterna 6 – 8, ventral view; 23. Penial valves, ventral view. 24. Vanhartenidia arabica sp. nov., female, holotype. Foretarsus. 25 – 29. V. felix sp. nov., paratype (25 – 28, male; 29, female). 25. Hind coxa, ventral view; 26. Genitalia, dorsal view; 27. Volsella, dorsal view; 28. Penial valves, ventral view; 29. Pygidial area. Scale = 1 mm for 22, 0.5 mm for 23 and 26, and 0.25 mm for 27 and 28." httpUri="https://zenodo.org/record/172675/files/figure.png" pageId="33" pageNumber="34">Fig. 22</figureCitation>
). Penial valves short, weakly asymmetrical (
<figureCitation id="CBF92A01FF847139FB0E8BAFFEA7E98B" captionStart="FIGURES 22 – 29. 22 – 23" captionStartId="40.[264,382,1451,1475]" captionTargetBox="[543,1028,890,1425]" captionTargetId="figure@40.[539,1048,889,1428]" captionTargetPageId="40" captionText="FIGURES 22 – 29. 22 – 23. Spinulomutilla yemenita sp. nov., male, holotype. 22. Sterna 6 – 8, ventral view; 23. Penial valves, ventral view. 24. Vanhartenidia arabica sp. nov., female, holotype. Foretarsus. 25 – 29. V. felix sp. nov., paratype (25 – 28, male; 29, female). 25. Hind coxa, ventral view; 26. Genitalia, dorsal view; 27. Volsella, dorsal view; 28. Penial valves, ventral view; 29. Pygidial area. Scale = 1 mm for 22, 0.5 mm for 23 and 26, and 0.25 mm for 27 and 28." httpUri="https://zenodo.org/record/172675/files/figure.png" pageId="33" pageNumber="34">Fig. 23</figureCitation>
). Mesoscutum and scutellum densely punctate, tegulae sparsely punctate. Propodeum dorsally and posteriorly reticulate. T2 and S2 sparsely punctate, punctures larger on S2 and sparser on tergal disc.
</paragraph>
<paragraph id="537D3684FF847139FECD884FFEC4EBAB" blockId="33.[264,1324,284,1030]" pageId="33" pageNumber="34">Head, mesosoma, coxae, tibiae, and femorae black; mandible black, reddish preapically; tegula glabrous, castaneous; tarsi ferruginous­red; antennae black with castaneous pedicel and apical part of scape; forewings fuscous. Metasoma ferruginous except brown S1. Head, except clypeal median part, pronotum dorsally and propodeum dorsally with dense recumbent and erect silvery setae; mesoscutum, scutellum and tegulae with black setae; mesopleurae with silvery setae; legs with sparse white setae; T2–T6 posteriorly with bands of dense golden recumbent setae inverted medially, T2 basally with wide band concave posteriorly of golden setae reverted back. T7 with sparse black setae. Metasomal sterna with sparse yellowish setae, S2–S7 with apical fringes of yellowish setae.</paragraph>
<paragraph id="537D3684FF847139FECD89FFFDB9EC73" blockId="33.[264,1324,284,1030]" box="[312,567,1004,1030]" pageId="33" pageNumber="34">FEMALE. Unknown.</paragraph>
</subSubSection>
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<typeStatus id="8C798826FF847139FEFD8E2FFEB1EC20" box="[264,319,1084,1109]" pageId="33" pageNumber="34">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="537D3684FF847139FECD8E77FA91EC0B" blockId="33.[264,1311,1084,1150]" box="[312,1311,1123,1150]" pageId="33" pageNumber="34">
<typeStatus id="8C798826FF847139FECD8E77FE2BEC0B" box="[312,421,1124,1150]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
ɗ,
<collectingCountry id="2BD57614FF847139FE3B8E77FDAAEC0B" box="[462,548,1124,1150]" name="Yemen" pageId="33" pageNumber="34">Yemen</collectingCountry>
: Al Kadan,
<date id="277C1044FF847139FD5A8E77FCF5EC0B" box="[687,891,1124,1150]" pageId="33" pageNumber="34" value="1998-02-17" valueMax="1998-03-31" valueMin="1998-02-17">17.II–31.III.1998</date>
, MT, AvH &amp; H.M. Naser [
<collectionCode id="35D3AE41FF847139FB418E77FA9AEC0B" LSID="urn:lsid:biocol.org:col:34975" box="[1204,1300,1124,1150]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="33" pageNumber="34">ZMAN</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF847139FEFD8EA7FE03EC83" box="[264,407,1204,1270]" pageId="33" pageNumber="34" type="distribution">
<paragraph id="537D3684FF847139FEFD8EA7FE19ECB8" blockId="33.[264,407,1204,1270]" box="[264,407,1204,1229]" pageId="33" pageNumber="34">
<emphasis id="61B6EA96FF847139FEFD8EA7FE19ECB8" box="[264,407,1204,1229]" italics="true" pageId="33" pageNumber="34">Distribution</emphasis>
</paragraph>
<paragraph id="537D3684FF847139FECD8ECFFE03EC83" blockId="33.[264,407,1204,1270]" box="[312,397,1244,1270]" pageId="33" pageNumber="34">
<collectingCountry id="2BD57614FF847139FECD8ECFFE09EC83" box="[312,391,1244,1270]" name="Yemen" pageId="33" pageNumber="34">Yemen</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF847139FEFD8F3FFE4FEDE3" pageId="33" pageNumber="34" type="etymology">
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<emphasis id="61B6EA96FF847139FEFD8F3FFE0BED30" box="[264,389,1324,1349]" italics="true" pageId="33" pageNumber="34">Etymology</emphasis>
</paragraph>
<paragraph id="537D3684FF847139FECD8F47FE4FEDE3" blockId="33.[264,1324,1324,1430]" pageId="33" pageNumber="34">
The specific name originates from
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, referring to the country where the species has been found.
</paragraph>
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</treatment>
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<emphasis id="61B6EA96FF86713BFEFD88EFFC0BEB63" bold="true" box="[264,901,764,790]" pageId="35" pageNumber="36">
<emphasis id="61B6EA96FF86713BFEFD88EFFE16EB63" bold="true" box="[264,408,764,790]" italics="true" pageId="35" pageNumber="36">Trogaspidia</emphasis>
(
<emphasis id="61B6EA96FF86713BFE5C88EFFDB7EB63" bold="true" box="[425,569,764,790]" italics="true" pageId="35" pageNumber="36">Trogaspidia</emphasis>
)
<emphasis id="61B6EA96FF86713BFDBF88EFFD5AEB63" bold="true" box="[586,724,764,790]" italics="true" pageId="35" pageNumber="36">miniaticeps</emphasis>
Bischoff 1920
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF86713BFEFD8954FC9DEBE8" pageId="35" pageNumber="36" type="reference_group">
<paragraph id="537D3684FF86713BFEFD8954FC9DEBE8" blockId="35.[264,1323,838,925]" pageId="35" pageNumber="36">
<taxonomicName id="94C24D07FF86713BFEFD8954FD6BEB28" authority="Bischoff 1920: 393" authorityName="Bischoff" authorityPageNumber="393" authorityYear="1920" box="[264,741,838,862]" class="Insecta" family="Mutillidae" genus="Trogaspidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="miniaticeps">
<emphasis id="61B6EA96FF86713BFEFD8954FD85EB2B" box="[264,523,839,862]" italics="true" pageId="35" pageNumber="36">Trogaspidia miniaticeps</emphasis>
Bischoff 1920: 393
</taxonomicName>
(Ψ), 449 (♂, as subspecies of
<taxonomicName id="94C24D07FF86713BFBC78954FB02EB2B" box="[1074,1164,839,862]" class="Insecta" family="Mutillidae" genus="Trogaspidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="divisa">
<emphasis id="61B6EA96FF86713BFBC78954FB02EB2B" box="[1074,1164,839,862]" italics="true" pageId="35" pageNumber="36">T. divisa</emphasis>
</taxonomicName>
),
<typeStatus id="8C798826FF86713BFB508955FA8BEB28" box="[1189,1285,838,861]" pageId="35" pageNumber="36" type="holotype">holotype</typeStatus>
, Ψ, North Aden (
<collectingCountry id="2BD57614FF86713BFE318975FD9AEB08" box="[452,532,870,893]" name="Yemen" pageId="35" pageNumber="36">Yemen</collectingCountry>
), Lakey,
<date id="277C1044FF86713BFD818975FD6FEB08" box="[628,737,870,893]" pageId="35" pageNumber="36" value="1895-03-06">6.III.1895</date>
, Bingham [ZMHU]; Hammer 1962: 9, Ψ (Aden);
<bibRefCitation id="37534B75FF86713BFB018975FDB6EBE8" author="Nonveiller" pageId="35" pageNumber="49" refString="Nonveiller, G. (1995) A preliminary approach to a revision of the Afrotropical representatives of the genus Trogaspidia Ashmead, 1899 (sensu Bischoff, 1920) (Hymenoptera, Mutillidae). Studies on African Mutillidae XXI. (Third contribution to the knowledge of the afrotropical Trogaspidia). Annales de la Societe Entomologique de France, (N. S.), 31 (4), 349 - 368." type="journal article" year="1995">Nonveiller &amp; Petersen 1995</bibRefCitation>
: 192, Ψ, ♂ (
<collectingCountry id="2BD57614FF86713BFD428995FC86EBE8" box="[695,776,902,925]" name="Yemen" pageId="35" pageNumber="36">Yemen</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF86713BFEFD89DCFE71ECFC" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="537D3684FF86713BFEFD89DCFE68EB9D" blockId="35.[264,1323,975,1161]" box="[264,486,975,1000]" pageId="35" pageNumber="36">
<emphasis id="61B6EA96FF86713BFEFD89DCFE68EB9D" box="[264,486,975,1000]" italics="true" pageId="35" pageNumber="36">Material examined</emphasis>
</paragraph>
<paragraph id="537D3684FF86713BFECD89E4FE71ECFC" blockId="35.[264,1323,975,1161]" pageId="35" pageNumber="36">
<collectingCountry id="2BD57614FF86713BFECD89E4FE00EC64" box="[312,398,1015,1041]" name="Yemen" pageId="35" pageNumber="36">Yemen</collectingCountry>
: Al Kowd,
<date id="277C1044FF86713BFDE089E4FD33EC64" box="[533,701,1015,1041]" pageId="35" pageNumber="36" value="1993-02-15" valueMax="1993-02-28" valueMin="1993-02-15">15–28.II.1993</date>
, MT, AvH, 1 ɗ [
<collectionCode id="35D3AE41FF86713BFC7489E4FC6FEC64" LSID="urn:lsid:biocol.org:col:34975" box="[897,993,1015,1041]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="35" pageNumber="36">ZMAN</collectionCode>
]; Lahj,
<date id="277C1044FF86713BFBCC89E4FB62EC64" box="[1081,1260,1015,1041]" pageId="35" pageNumber="36" value="2000-06-15" valueMax="2000-06-30" valueMin="2000-06-15">15–30.VI.2000</date>
, MT, AvH &amp; A. Sallam, 1 ɗ [
<collectionCode id="35D3AE41FF86713BFDDC8E0CFDE1EC4C" box="[553,623,1055,1081]" pageId="35" pageNumber="36">IBSS</collectionCode>
];
<date id="277C1044FF86713BFD718E0CFD54EC4C" box="[644,730,1055,1081]" pageId="35" pageNumber="36" value="2000-05">V.2000</date>
, MT, AvH &amp; A. Sallam, 1 ɗ [
<collectionCode id="35D3AE41FF86713BFBB78E0CFB2CEC4C" LSID="urn:lsid:biocol.org:col:34992" box="[1090,1186,1055,1081]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="35" pageNumber="36">RMNH</collectionCode>
];
<date id="277C1044FF86713BFB5B8E0CFAA8EC4C" box="[1198,1318,1055,1081]" pageId="35" pageNumber="36" value="2000-08">VIII.2000</date>
, MT, AvH &amp; A. Sallam, 1 ɗ [
<collectionCode id="35D3AE41FF86713BFD928E54FD46EC14" LSID="urn:lsid:biocol.org:col:34992" box="[615,712,1095,1121]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="35" pageNumber="36">RMNH</collectionCode>
]; Al Lahima,
<date id="277C1044FF86713BFC9B8E54FBC6EC14" box="[878,1096,1095,1121]" pageId="35" pageNumber="36" value="2000-10-16" valueMax="2000-12-31" valueMin="2000-10-16">16.X–31.XII.2000</date>
, MT, AvH &amp; A.H. Hager, 1 ɗ [
<collectionCode id="35D3AE41FF86713BFE608E7CFE7BECFC" LSID="urn:lsid:biocol.org:col:34975" box="[405,501,1135,1161]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="35" pageNumber="36">ZMAN</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF86713BFEFD8EACFB82ED74" box="[264,1036,1215,1281]" pageId="35" pageNumber="36" type="distribution">
<paragraph id="537D3684FF86713BFEFD8EACFE19ECAD" blockId="35.[264,1036,1215,1281]" box="[264,407,1215,1240]" pageId="35" pageNumber="36">
<emphasis id="61B6EA96FF86713BFEFD8EACFE19ECAD" box="[264,407,1215,1240]" italics="true" pageId="35" pageNumber="36">Distribution</emphasis>
</paragraph>
<paragraph id="537D3684FF86713BFECD8EF4FB82ED74" blockId="35.[264,1036,1215,1281]" box="[312,1036,1255,1281]" pageId="35" pageNumber="36">
<collectingCountry id="2BD57614FF86713BFECD8EF4FE03ED74" box="[312,397,1255,1281]" name="Yemen" pageId="35" pageNumber="36">Yemen</collectingCountry>
,
<collectingCountry id="2BD57614FF86713BFE6D8EF4FDBBED74" box="[408,565,1255,1281]" name="Saudi Arabia" pageId="35" pageNumber="36">Saudi Arabia</collectingCountry>
,
<collectingCountry id="2BD57614FF86713BFDB48EF4FD2FED74" box="[577,673,1255,1281]" name="Somalia" pageId="35" pageNumber="36">Somalia</collectingCountry>
(
<bibRefCitation id="37534B75FF86713BFD458EF4FC71ED74" author="Nonveiller" box="[688,1023,1255,1281]" pageId="35" pageNumber="49" refString="Nonveiller, G. (1995) A preliminary approach to a revision of the Afrotropical representatives of the genus Trogaspidia Ashmead, 1899 (sensu Bischoff, 1920) (Hymenoptera, Mutillidae). Studies on African Mutillidae XXI. (Third contribution to the knowledge of the afrotropical Trogaspidia). Annales de la Societe Entomologique de France, (N. S.), 31 (4), 349 - 368." type="journal article" year="1995">Nonveiller &amp; Petersen 1995</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

112
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<subSubSection id="1BD8650FFF86713BFEFD8F4DFCEBED0D" box="[264,869,1374,1400]" pageId="35" pageNumber="36" type="nomenclature">
<paragraph id="537D3684FF86713BFEFD8F4DFCEBED0D" blockId="35.[264,869,1374,1400]" box="[264,869,1374,1400]" pageId="35" pageNumber="36">
<heading id="083581E8FF86713BFEFD8F4DFCEBED0D" bold="true" box="[264,869,1374,1400]" fontSize="11" level="1" pageId="35" pageNumber="36" reason="1">
<taxonomicName id="94C24D07FF86713BFEFD8F4DFCEBED0D" authority="Bischoff 1920" authorityName="Bischoff" authorityYear="1920" box="[264,869,1374,1400]" class="Insecta" family="Mutillidae" genus="Trogaspidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="nilotica" subGenus="Acutitropidia">
<emphasis id="61B6EA96FF86713BFEFD8F4DFCEBED0D" bold="true" box="[264,869,1374,1400]" pageId="35" pageNumber="36">
<emphasis id="61B6EA96FF86713BFEFD8F4DFE16ED0D" bold="true" box="[264,408,1374,1400]" italics="true" pageId="35" pageNumber="36">Trogaspidia</emphasis>
(
<emphasis id="61B6EA96FF86713BFE5C8F4DFDC7ED0D" bold="true" box="[425,585,1374,1400]" italics="true" pageId="35" pageNumber="36">Acutitropidia</emphasis>
)
<emphasis id="61B6EA96FF86713BFDAF8F4DFD3BED0D" bold="true" box="[602,693,1374,1400]" italics="true" pageId="35" pageNumber="36">nilotica</emphasis>
Bischoff 1920
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF86713BFEFD8FB9FB77ED95" pageId="35" pageNumber="36" type="reference_group">
<paragraph id="537D3684FF86713BFEFD8FB9FB77ED95" blockId="35.[264,1322,1449,1504]" pageId="35" pageNumber="36">
<treatmentCitationGroup id="73D211AAFF86713BFEFD8FB9FB77ED95" pageId="35" pageNumber="36">
<taxonomicName id="94C24D07FF86713BFEFD8FB9FD5FEDB5" authority="Bischoff 1920: 416" authorityName="Bischoff" authorityPageNumber="416" authorityYear="1920" box="[264,721,1449,1473]" class="Insecta" family="Mutillidae" genus="Trogaspidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="nilotica">
<emphasis id="61B6EA96FF86713BFEFD8FB9FE6BEDB4" box="[264,485,1450,1473]" italics="true" pageId="35" pageNumber="36">Trogaspidia nilotica</emphasis>
Bischoff 1920: 416
</taxonomicName>
, ♂,
<typeStatus id="8C798826FF86713BFCFD8FBAFCE6EDB5" box="[776,872,1449,1472]" pageId="35" pageNumber="36" type="holotype">holotype</typeStatus>
, &quot;Tura b. Heliopolis,
<date id="277C1044FF86713BFB928FBAFB28EDB5" box="[1127,1190,1449,1472]" pageId="35" pageNumber="36" value="1919-09">IX.19</date>
, Schwabel&quot; (
<collectingCountry id="2BD57614FF86713BFECA8FDAFE0FED95" box="[319,385,1481,1504]" name="Egypt" pageId="35" pageNumber="36">Egypt</collectingCountry>
) [ZMHU];
<treatmentCitation id="D2631095FF86713BFE0B8FDAFCEFED95" author="Nonveiller" box="[510,865,1481,1504]" page="183" pageId="35" pageNumber="36" year="1995">
<bibRefCitation id="37534B75FF86713BFE0B8FDAFCA2ED95" author="Nonveiller" box="[510,812,1481,1504]" pageId="35" pageNumber="49" refString="Nonveiller, G. (1995) A preliminary approach to a revision of the Afrotropical representatives of the genus Trogaspidia Ashmead, 1899 (sensu Bischoff, 1920) (Hymenoptera, Mutillidae). Studies on African Mutillidae XXI. (Third contribution to the knowledge of the afrotropical Trogaspidia). Annales de la Societe Entomologique de France, (N. S.), 31 (4), 349 - 368." type="journal article" year="1995">Nonveiller &amp; Petersen 1995</bibRefCitation>
: 183
</treatmentCitation>
, ♂;
<treatmentCitation id="D2631095FF86713BFC798FDAFBFDED95" author="Nonveiller" box="[908,1139,1481,1504]" page="360" pageId="35" pageNumber="36" year="1995">
<bibRefCitation id="37534B75FF86713BFC798FDAFBB0ED95" author="Nonveiller" box="[908,1086,1481,1504]" pageId="35" pageNumber="49" refString="Nonveiller, G. (1995) A preliminary approach to a revision of the Afrotropical representatives of the genus Trogaspidia Ashmead, 1899 (sensu Bischoff, 1920) (Hymenoptera, Mutillidae). Studies on African Mutillidae XXI. (Third contribution to the knowledge of the afrotropical Trogaspidia). Annales de la Societe Entomologique de France, (N. S.), 31 (4), 349 - 368." type="journal article" year="1995">Nonveiller 1995</bibRefCitation>
: 360
</treatmentCitation>
, ♂ (
<collectingCountry id="2BD57614FF86713BFB688FDAFB60ED95" box="[1181,1262,1481,1504]" name="Yemen" pageId="35" pageNumber="36">Yemen</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF86713BFEFD8C02FDE5EE26" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="537D3684FF86713BFEFD8C02FE68EE5F" blockId="35.[264,1048,1553,1699]" box="[264,486,1553,1578]" pageId="35" pageNumber="36">
<emphasis id="61B6EA96FF86713BFEFD8C02FE68EE5F" box="[264,486,1553,1578]" italics="true" pageId="35" pageNumber="36">Material examined</emphasis>
</paragraph>
<paragraph id="537D3684FF86713BFECD8C2AFDE5EE26" blockId="35.[264,1048,1553,1699]" box="[312,619,1593,1619]" pageId="35" pageNumber="36">Not found in our material.</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF86713BFEFD8C72FB96EED6" box="[264,1048,1633,1699]" pageId="35" pageNumber="36" type="distribution">
<paragraph id="537D3684FF86713BFEFD8C72FE19EE0F" blockId="35.[264,1048,1553,1699]" box="[264,407,1633,1658]" pageId="35" pageNumber="36">
<emphasis id="61B6EA96FF86713BFEFD8C72FE19EE0F" box="[264,407,1633,1658]" italics="true" pageId="35" pageNumber="36">Distribution</emphasis>
</paragraph>
<paragraph id="537D3684FF86713BFECD8C9AFB96EED6" blockId="35.[264,1048,1553,1699]" box="[312,1048,1673,1699]" pageId="35" pageNumber="36">
<collectingCountry id="2BD57614FF86713BFECD8C9AFE03EED6" box="[312,397,1673,1699]" name="Yemen" pageId="35" pageNumber="36">Yemen</collectingCountry>
,
<collectingCountry id="2BD57614FF86713BFE6D8C9AFDBBEED6" box="[408,565,1673,1699]" name="Saudi Arabia" pageId="35" pageNumber="36">Saudi Arabia</collectingCountry>
,
<collectingCountry id="2BD57614FF86713BFDB48C9AFD02EED6" box="[577,652,1673,1699]" name="Oman" pageId="35" pageNumber="36">Oman</collectingCountry>
,
<collectingCountry id="2BD57614FF86713BFD6D8C9AFD6AEED6" box="[664,740,1673,1699]" name="Sudan" pageId="35" pageNumber="36">Sudan</collectingCountry>
,
<collectingCountry id="2BD57614FF86713BFD1A8C9AFCBBEED6" box="[751,821,1673,1699]" name="Egypt" pageId="35" pageNumber="36">Egypt</collectingCountry>
(
<bibRefCitation id="37534B75FF86713BFCB08C9AFB85EED6" author="Nonveiller" box="[837,1035,1673,1699]" pageId="35" pageNumber="49" refString="Nonveiller, G. (1995) A preliminary approach to a revision of the Afrotropical representatives of the genus Trogaspidia Ashmead, 1899 (sensu Bischoff, 1920) (Hymenoptera, Mutillidae). Studies on African Mutillidae XXI. (Third contribution to the knowledge of the afrotropical Trogaspidia). Annales de la Societe Entomologique de France, (N. S.), 31 (4), 349 - 368." type="journal article" year="1995">Nonveiller 1995</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

159
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&quot;. Synonymized by
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, 1 Ψ, A. Mochi [
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<treatment id="DB6B8792FF87713BFEFD8FA2FCFDEAEB" ID-DOI="http://doi.org/10.5281/zenodo.6252986" ID-GBIF-Taxon="119344672" ID-Zenodo-Dep="6252986" LSID="urn:lsid:plazi:treatment:DB6B8792FF87713BFEFD8FA2FCFDEAEB" httpUri="http://treatment.plazi.org/id/DB6B8792FF87713BFEFD8FA2FCFDEAEB" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
<subSubSection id="1BD8650FFF87713AFEFD8FA2FCB4EDBE" box="[264,826,1457,1483]" pageId="34" pageNumber="35" type="nomenclature">
<paragraph id="537D3684FF87713AFEFD8FA2FCB4EDBE" blockId="34.[264,826,1457,1483]" box="[264,826,1457,1483]" pageId="34" pageNumber="35">
<heading id="083581E8FF87713AFEFD8FA2FCB4EDBE" bold="true" box="[264,826,1457,1483]" fontSize="11" level="1" pageId="34" pageNumber="35" reason="1">
<taxonomicName id="94C24D07FF87713AFEFD8FA2FCB4EDBE" authority="Klug 1829" authorityName="Klug" authorityYear="1829" box="[264,826,1457,1483]" class="Insecta" family="Mutillidae" genus="Trogaspidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="floralis" subGenus="Trogaspidia">
<emphasis id="61B6EA96FF87713AFEFD8FA2FCB4EDBE" bold="true" box="[264,826,1457,1483]" pageId="34" pageNumber="35">
<emphasis id="61B6EA96FF87713AFEFD8FA2FE16EDBE" bold="true" box="[264,408,1457,1483]" italics="true" pageId="34" pageNumber="35">Trogaspidia</emphasis>
(
<emphasis id="61B6EA96FF87713AFE5C8FA2FDB7EDBE" bold="true" box="[425,569,1457,1483]" italics="true" pageId="34" pageNumber="35">Trogaspidia</emphasis>
)
<emphasis id="61B6EA96FF87713AFDBF8FA2FD2EEDBE" bold="true" box="[586,672,1457,1483]" italics="true" pageId="34" pageNumber="35">floralis</emphasis>
(
<bibRefCitation id="37534B75FF87713AFD5A8FA2FCBCEDBE" author="Klug" box="[687,818,1457,1483]" pageId="34" pageNumber="48" refString="Klug, J. C. F. (1829) Symbolae physicae seu icones et descriptiones insectorum quae ex itinere per Africam borealem et Asiam occidentalem Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg studio novae aut illustratae redierunt. Hymenoptera. Decas Prima [genera Mutilla et Apterogyna]. Venditur a Mittlero, Berolini. [15 - 21] + tab. 4, 5." type="journal volume" year="1829">Klug 1829</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF87713AFEFD8FE4FEFEEF18" pageId="34" pageNumber="35" type="reference_group">
<paragraph id="537D3684FF87713AFEFD8FE4FA81EE78" blockId="34.[264,1322,1526,1901]" box="[264,1295,1526,1550]" pageId="34" pageNumber="35">
<treatmentCitationGroup id="73D211AAFF87713AFEFD8FE4FA81EE78" box="[264,1295,1526,1550]" pageId="34" pageNumber="35">
<taxonomicName id="94C24D07FF87713AFEFD8FE4FD0CEE78" authority="Olivier 1811: 62" authorityName="Olivier" authorityPageNumber="62" authorityYear="1811" box="[264,642,1526,1550]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="interrupta">
<emphasis id="61B6EA96FF87713AFEFD8FE4FE47EE7B" box="[264,457,1527,1550]" italics="true" pageId="34" pageNumber="35">Mutilla interrupta</emphasis>
<treatmentCitation id="D2631095FF87713AFE258FE5FD0CEE78" author="Olivier" box="[464,642,1526,1549]" page="62" pageId="34" pageNumber="35" year="1811">
<bibRefCitation id="37534B75FF87713AFE258FE5FDD4EE78" author="Olivier" box="[464,602,1526,1549]" pageId="34" pageNumber="49" refString="Olivier, A. G. (1811) Encyclopedie methodique. Histoire naturelle. Insectes. 8. H. Agasse, Paris, 722 pp. [Mutillidae - p. 48 - 66, 128 - 131]." type="book chapter" year="1811">Olivier 1811</bibRefCitation>
: 62
</treatmentCitation>
</taxonomicName>
, Ψ, &quot;se trouve en Arabie&quot;, nom. praeocc., non Latreille 1792.
</treatmentCitationGroup>
</paragraph>
<paragraph id="537D3684FF87713AFEFD8C04FB79EE38" blockId="34.[264,1322,1526,1901]" pageId="34" pageNumber="35">
<taxonomicName id="94C24D07FF87713AFEFD8C04FDA6EE58" authority="Klug 1829" authorityName="Klug" authorityYear="1829" box="[264,552,1558,1582]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="floralis">
<emphasis id="61B6EA96FF87713AFEFD8C04FE25EE5B" box="[264,427,1559,1582]" italics="true" pageId="34" pageNumber="35">Mutilla floralis</emphasis>
<bibRefCitation id="37534B75FF87713AFE478C05FDA6EE58" author="Klug" box="[434,552,1558,1581]" pageId="34" pageNumber="48" refString="Klug, J. C. F. (1829) Symbolae physicae seu icones et descriptiones insectorum quae ex itinere per Africam borealem et Asiam occidentalem Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg studio novae aut illustratae redierunt. Hymenoptera. Decas Prima [genera Mutilla et Apterogyna]. Venditur a Mittlero, Berolini. [15 - 21] + tab. 4, 5." type="journal volume" year="1829">Klug 1829</bibRefCitation>
</taxonomicName>
: [19], tab. 5, fig. 1, 2, ♂, Fajum (
<collectingCountry id="2BD57614FF87713AFC798C05FC5FEE58" box="[908,977,1558,1581]" name="Egypt" pageId="34" pageNumber="35">Egypt</collectingCountry>
),
<typeStatus id="8C798826FF87713AFC118C05FBC8EE58" box="[996,1094,1558,1581]" pageId="34" pageNumber="35" type="lectotype">lectotype</typeStatus>
(designated Bischoff 1920: 449), &quot;Fajum, Ehrenberg&quot; [ZMHU]. Synonymized with
<taxonomicName id="94C24D07FF87713AFC268C24FB9DEE3B" box="[979,1043,1591,1614]" class="Insecta" family="Mutillidae" genus="Trogaspidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="divisa">
<emphasis id="61B6EA96FF87713AFC268C24FB9DEE3B" box="[979,1043,1591,1614]" italics="true" pageId="34" pageNumber="35">divisa</emphasis>
</taxonomicName>
by
<bibRefCitation id="37534B75FF87713AFBCE8C25FB33EE38" author="Andre" box="[1083,1213,1590,1613]" pageId="34" pageNumber="47" refString="Andre, E. (1896) Notes pour servir a la connaissance de mutilles palearctiques et description de quelques especes nouvelles. Deuxieme partie. Memoirs de la Societe Zoologique de France, 9, 261 - 277." type="journal article" year="1896">André 1896</bibRefCitation>
: 271.
</paragraph>
<paragraph id="537D3684FF87713AFEFD8C44FBBCEE18" blockId="34.[264,1322,1526,1901]" box="[264,1074,1622,1646]" pageId="34" pageNumber="35">
<taxonomicName id="94C24D07FF87713AFEFD8C44FDC2EE18" authority="Klug 1829" authorityName="Klug" authorityYear="1829" box="[264,588,1622,1646]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="interrupta">
<emphasis id="61B6EA96FF87713AFEFD8C44FE47EE1B" box="[264,457,1623,1646]" italics="true" pageId="34" pageNumber="35">Mutilla interrupta</emphasis>
:
<bibRefCitation id="37534B75FF87713AFE228C45FDC2EE18" author="Klug" box="[471,588,1622,1645]" pageId="34" pageNumber="48" refString="Klug, J. C. F. (1829) Symbolae physicae seu icones et descriptiones insectorum quae ex itinere per Africam borealem et Asiam occidentalem Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg studio novae aut illustratae redierunt. Hymenoptera. Decas Prima [genera Mutilla et Apterogyna]. Venditur a Mittlero, Berolini. [15 - 21] + tab. 4, 5." type="journal volume" year="1829">Klug 1829</bibRefCitation>
</taxonomicName>
: [18], tab. 4, fig. 11, Ψ; Magretti 1906: 46, Ψ.
</paragraph>
<paragraph id="537D3684FF87713AFEFD8C64FD2EEED8" blockId="34.[264,1322,1526,1901]" pageId="34" pageNumber="35">
<treatmentCitationGroup id="73D211AAFF87713AFEFD8C64FD2EEED8" pageId="34" pageNumber="35">
<taxonomicName id="94C24D07FF87713AFEFD8C64FDC8EEF8" authority="Smith 1855: 11" authorityName="Smith" authorityPageNumber="11" authorityYear="1855" box="[264,582,1654,1678]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="divisa">
<emphasis id="61B6EA96FF87713AFEFD8C64FE15EEFB" box="[264,411,1655,1678]" italics="true" pageId="34" pageNumber="35">Mutilla divisa</emphasis>
<treatmentCitation id="D2631095FF87713AFE568C65FDC8EEF8" author="Smith" box="[419,582,1654,1677]" page="11" pageId="34" pageNumber="35" year="1855">
<bibRefCitation id="37534B75FF87713AFE568C65FDAFEEF8" author="Smith" box="[419,545,1654,1677]" pageId="34" pageNumber="50" refString="Smith, F. (1855) Catalogue of Hymenopterous Insects in the Collection of the British Museum. Part III. Mutillidae and Pompilidae. Taylor and Francis, London. 206 pp. + 5 pls. [Mutillidae - p. l - 66, pl. 1]." type="book" year="1855">Smith 1855</bibRefCitation>
: 11
</treatmentCitation>
</taxonomicName>
, nom. nov. pro
<taxonomicName id="94C24D07FF87713AFD1F8C64FC68EEF8" authority="Olivier 1811" authorityName="Olivier" authorityYear="1811" box="[746,998,1654,1678]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="interrupta">
<emphasis id="61B6EA96FF87713AFD1F8C64FCD8EEFB" box="[746,854,1655,1678]" italics="true" pageId="34" pageNumber="35">interrupta</emphasis>
<bibRefCitation id="37534B75FF87713AFCA98C65FC68EEF8" author="Olivier" box="[860,998,1654,1677]" pageId="34" pageNumber="49" refString="Olivier, A. G. (1811) Encyclopedie methodique. Histoire naturelle. Insectes. 8. H. Agasse, Paris, 722 pp. [Mutillidae - p. 48 - 66, 128 - 131]." type="book chapter" year="1811">Olivier 1811</bibRefCitation>
</taxonomicName>
. Synonymized with
<taxonomicName id="94C24D07FF87713AFB488C64FA84EEFB" box="[1213,1290,1655,1678]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="floralis">
<emphasis id="61B6EA96FF87713AFB488C64FA84EEFB" box="[1213,1290,1655,1678]" italics="true" pageId="34" pageNumber="35">floralis</emphasis>
</taxonomicName>
by
<treatmentCitation id="D2631095FF87713AFECD8C85FD15EED8" author="Nonveiller" box="[312,667,1686,1709]" page="184" pageId="34" pageNumber="35" year="1995">
<bibRefCitation id="37534B75FF87713AFECD8C85FDE8EED8" author="Nonveiller" box="[312,614,1686,1709]" pageId="34" pageNumber="49" refString="Nonveiller, G. (1995) A preliminary approach to a revision of the Afrotropical representatives of the genus Trogaspidia Ashmead, 1899 (sensu Bischoff, 1920) (Hymenoptera, Mutillidae). Studies on African Mutillidae XXI. (Third contribution to the knowledge of the afrotropical Trogaspidia). Annales de la Societe Entomologique de France, (N. S.), 31 (4), 349 - 368." type="journal article" year="1995">Nonveiller &amp; Petersen 1995</bibRefCitation>
: 184
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="537D3684FF87713AFEFD8CA4FD79EEB8" blockId="34.[264,1322,1526,1901]" box="[264,759,1718,1742]" pageId="34" pageNumber="35">
<treatmentCitationGroup id="73D211AAFF87713AFEFD8CA4FD79EEB8" box="[264,759,1718,1742]" pageId="34" pageNumber="35">
<taxonomicName id="94C24D07FF87713AFEFD8CA4FDEFEEB8" authority="Andre 1896: 271" authorityName="Andre" authorityPageNumber="271" authorityYear="1896" box="[264,609,1718,1742]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="divisa">
<emphasis id="61B6EA96FF87713AFEFD8CA4FE13EEBB" box="[264,413,1719,1742]" italics="true" pageId="34" pageNumber="35">Mutilla divisa</emphasis>
:
<treatmentCitation id="D2631095FF87713AFE5E8CA5FDEFEEB8" author="Andre" box="[427,609,1718,1741]" page="271" pageId="34" pageNumber="35" year="1896">
<bibRefCitation id="37534B75FF87713AFE5E8CA5FDA3EEB8" author="Andre" box="[427,557,1718,1741]" pageId="34" pageNumber="47" refString="Andre, E. (1896) Notes pour servir a la connaissance de mutilles palearctiques et description de quelques especes nouvelles. Deuxieme partie. Memoirs de la Societe Zoologique de France, 9, 261 - 277." type="journal article" year="1896">André 1896</bibRefCitation>
: 271
</treatmentCitation>
</taxonomicName>
, Ψ, ♂ (Aden).
</treatmentCitationGroup>
</paragraph>
<paragraph id="537D3684FF87713AFEFD8CC4FC91EE98" blockId="34.[264,1322,1526,1901]" box="[264,799,1750,1774]" pageId="34" pageNumber="35">
<taxonomicName id="94C24D07FF87713AFEFD8CC4FDF5EE98" authority="Magretti 1906: 46" authorityName="Magretti" authorityPageNumber="46" authorityYear="1906" box="[264,635,1750,1774]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="floralis">
<emphasis id="61B6EA96FF87713AFEFD8CC4FE24EE9B" box="[264,426,1751,1774]" italics="true" pageId="34" pageNumber="35">Mutilla floralis</emphasis>
: Magretti 1906: 46
</taxonomicName>
, Ψ, ♂ (
<collectingCountry id="2BD57614FF87713AFD308CC5FC9AEE98" box="[709,788,1750,1773]" name="Eritrea" pageId="34" pageNumber="35">Eritrea</collectingCountry>
).
</paragraph>
<paragraph id="537D3684FF87713AFEFD8CE4FCB5EF78" blockId="34.[264,1322,1526,1901]" box="[264,827,1782,1806]" pageId="34" pageNumber="35">
<taxonomicName id="94C24D07FF87713AFEFD8CE4FD27EF78" authority="Bischoff 1920: 386" authorityName="Bischoff" authorityPageNumber="386" authorityYear="1920" box="[264,681,1782,1806]" class="Insecta" family="Mutillidae" genus="Trogaspidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="divisa">
<emphasis id="61B6EA96FF87713AFEFD8CE4FE40EF7B" box="[264,462,1783,1806]" italics="true" pageId="34" pageNumber="35">Trogaspidia divisa</emphasis>
: Bischoff 1920: 386
</taxonomicName>
(Ψ), 449 (♂).
</paragraph>
<paragraph id="537D3684FF87713AFEFD8D04FCF5EF58" blockId="34.[264,1322,1526,1901]" box="[264,891,1814,1838]" pageId="34" pageNumber="35">
<taxonomicName id="94C24D07FF87713AFEFD8D04FC8BEF58" authority="Hammer 1962: 23" authorityName="Hammer" authorityPageNumber="23" authorityYear="1962" box="[264,773,1814,1838]" class="Insecta" family="Mutillidae" genus="Trogaspidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="subSpecies" species="divisa" subSpecies="klugiana">
<emphasis id="61B6EA96FF87713AFEFD8D04FDBCEF5B" box="[264,562,1815,1838]" italics="true" pageId="34" pageNumber="35">Trogaspidia divisa klugiana</emphasis>
: Hammer 1962: 23
</taxonomicName>
, ♂ (Aden).
</paragraph>
<paragraph id="537D3684FF87713AFEFD8D24FEFEEF18" blockId="34.[264,1322,1526,1901]" pageId="34" pageNumber="35">
<treatmentCitationGroup id="73D211AAFF87713AFEFD8D24FEFEEF18" pageId="34" pageNumber="35">
<taxonomicName id="94C24D07FF87713AFEFD8D24FC60EF38" authority="Nonveiller &amp; Petersen 1995: 183" authorityName="Nonveiller &amp; Petersen" authorityPageNumber="183" authorityYear="1995" box="[264,1006,1846,1870]" class="Insecta" family="Mutillidae" genus="Trogaspidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="floralis" subGenus="Trogaspidia">
<emphasis id="61B6EA96FF87713AFEFD8D24FE06EF3B" box="[264,392,1847,1870]" italics="true" pageId="34" pageNumber="35">Trogaspidia</emphasis>
(
<emphasis id="61B6EA96FF87713AFE6C8D24FD97EF3B" box="[409,537,1847,1870]" italics="true" pageId="34" pageNumber="35">Trogaspidia</emphasis>
)
<emphasis id="61B6EA96FF87713AFDDF8D24FDF9EF3B" box="[554,631,1847,1870]" italics="true" pageId="34" pageNumber="35">floralis</emphasis>
:
<treatmentCitation id="D2631095FF87713AFD738D25FC60EF38" author="Nonveiller" box="[646,1006,1846,1869]" page="183" pageId="34" pageNumber="35" year="1995">
<bibRefCitation id="37534B75FF87713AFD738D25FC36EF38" author="Nonveiller" box="[646,952,1846,1869]" pageId="34" pageNumber="49" refString="Nonveiller, G. (1995) A preliminary approach to a revision of the Afrotropical representatives of the genus Trogaspidia Ashmead, 1899 (sensu Bischoff, 1920) (Hymenoptera, Mutillidae). Studies on African Mutillidae XXI. (Third contribution to the knowledge of the afrotropical Trogaspidia). Annales de la Societe Entomologique de France, (N. S.), 31 (4), 349 - 368." type="journal article" year="1995">Nonveiller &amp; Petersen 1995</bibRefCitation>
: 183
</treatmentCitation>
</taxonomicName>
, Ψ, ♂;
<treatmentCitation id="D2631095FF87713AFBC88D25FAA8EF38" author="Nonveiller" box="[1085,1318,1846,1869]" page="361" pageId="34" pageNumber="35" year="1995">
<bibRefCitation id="37534B75FF87713AFBC88D25FB7EEF38" author="Nonveiller" box="[1085,1264,1846,1869]" pageId="34" pageNumber="49" refString="Nonveiller, G. (1995) A preliminary approach to a revision of the Afrotropical representatives of the genus Trogaspidia Ashmead, 1899 (sensu Bischoff, 1920) (Hymenoptera, Mutillidae). Studies on African Mutillidae XXI. (Third contribution to the knowledge of the afrotropical Trogaspidia). Annales de la Societe Entomologique de France, (N. S.), 31 (4), 349 - 368." type="journal article" year="1995">Nonveiller 1995</bibRefCitation>
: 361
</treatmentCitation>
, Ψ, ♂.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF86713BFEFD8B0FFD1AE98B" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="537D3684FF86713BFEFD8B0FFE68E940" blockId="35.[264,1323,284,510]" box="[264,486,284,309]" pageId="35" pageNumber="36">
<emphasis id="61B6EA96FF86713BFEFD8B0FFE68E940" box="[264,486,284,309]" italics="true" pageId="35" pageNumber="36">Material examined</emphasis>
</paragraph>
<paragraph id="537D3684FF86713BFECD8B57FD1AE98B" blockId="35.[264,1323,284,510]" pageId="35" pageNumber="36">
Form with red head and mesosoma (f.
<emphasis id="61B6EA96FF86713BFCDF8B57FC2CE928" box="[810,930,324,349]" italics="true" pageId="35" pageNumber="36">moslemita</emphasis>
Bischoff).
<collectingCountry id="2BD57614FF86713BFBC68B57FB07E92B" box="[1075,1161,324,350]" name="Yemen" pageId="35" pageNumber="36">Yemen</collectingCountry>
:
<quantity id="943A9B61FF86713BFB6E8B57FB65E92B" box="[1179,1259,324,350]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="35" pageNumber="36" unit="km" value="12.0">12 km</quantity>
NW Manakhah,
<date id="277C1044FF86713BFE618B7FFDF3E9F3" box="[404,637,364,390]" pageId="35" pageNumber="36" value="2001-07-03" valueMax="2001-08-21" valueMin="2001-07-03">3.VII–21.VIII.2001</date>
, MT, AvH, 14 ɗ [
<collectionCode id="35D3AE41FF86713BFCAD8B7FFC38E9F3" LSID="urn:lsid:biocol.org:col:34992" box="[856,950,364,390]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="35" pageNumber="36">RMNH</collectionCode>
,
<collectionCode id="35D3AE41FF86713BFC318B7FFB86E9F3" box="[964,1032,364,390]" pageId="35" pageNumber="36">IBSS</collectionCode>
]; Al Lahima,
<date id="277C1044FF86713BFB5B8B7FFAA8E9F3" box="[1198,1318,364,390]" pageId="35" pageNumber="36" value="2001-01" valueMax="2001-04-31" valueMin="2001-01-00">I–IV.2001</date>
, MT, AvH &amp; A.M. Hajer, 1 ɗ [
<collectionCode id="35D3AE41FF86713BFD858B87FD5EE9DB" LSID="urn:lsid:biocol.org:col:34975" box="[624,720,404,430]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="35" pageNumber="36">ZMAN</collectionCode>
]; Sanaa,
<date id="277C1044FF86713BFCCC8B87FC3CE9DB" box="[825,946,404,430]" pageId="35" pageNumber="36" value="1931-11-05">5.XI.1931</date>
, 1 ɗ, Zhenzhurist (N. Filippov) [
<collectionCode id="35D3AE41FF86713BFEFA8BAFFEFAE9A3" box="[271,372,444,470]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="35" pageNumber="36">ZMMU</collectionCode>
]. Form with black head and mesosoma (f.
<emphasis id="61B6EA96FF86713BFC8E8BAFFC6FE9A0" box="[891,993,444,469]" italics="true" pageId="35" pageNumber="36">klugiana</emphasis>
André).
<collectingCountry id="2BD57614FF86713BFBA48BAFFB28E9A3" box="[1105,1190,444,470]" name="Yemen" pageId="35" pageNumber="36">Yemen</collectingCountry>
, Al Kowd,
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<collectingCountry id="2BD57614FF86713BFECD884FFE03EA03" box="[312,397,604,630]" name="Yemen" pageId="35" pageNumber="36">Yemen</collectingCountry>
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,
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<bibRefCitation id="37534B75FF86713BFDE28897FCE8EAEB" author="Nonveiller" box="[535,870,644,670]" pageId="35" pageNumber="49" refString="Nonveiller, G. (1995) A preliminary approach to a revision of the Afrotropical representatives of the genus Trogaspidia Ashmead, 1899 (sensu Bischoff, 1920) (Hymenoptera, Mutillidae). Studies on African Mutillidae XXI. (Third contribution to the knowledge of the afrotropical Trogaspidia). Annales de la Societe Entomologique de France, (N. S.), 31 (4), 349 - 368." type="journal article" year="1995">Nonveiller &amp; Petersen 1995</bibRefCitation>
).
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135
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*
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<emphasis id="61B6EA96FF887135FEE28995FDD6EBD5" bold="true" box="[279,600,902,928]" italics="true" pageId="45" pageNumber="46">Tricholabiodes sudanensis</emphasis>
Suárez 1967
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<emphasis id="61B6EA96FF887135FEFD89C1FE21EB9C" box="[264,431,978,1001]" italics="true" pageId="45" pageNumber="46">Tricholabioides</emphasis>
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(!)
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<collectingCountry id="2BD57614FF887135FECD8E92FE00ECEE" box="[312,398,1153,1179]" name="Yemen" pageId="45" pageNumber="46">Yemen</collectingCountry>
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<emphasis id="61B6EA96FF887135FEFD8BAFFDBBE9A3" bold="true" box="[264,565,444,470]" italics="true" pageId="45" pageNumber="46">Tricholabiodes scorteccii</emphasis>
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<emphasis id="61B6EA96FF887135FEFD8814FD94EA6B" box="[264,538,519,542]" italics="true" pageId="45" pageNumber="46">Tricholabiodes scorteccii</emphasis>
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, ♂Ψ,
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, ♂ &quot;Ingeramis&quot; [
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, Scortecci [Museo Civico di Storia Naturale di Milano];
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<bibRefCitation id="37534B75FF887135FC798835FB9AEA48" author="Suarez" box="[908,1044,550,573]" pageId="45" pageNumber="50" refString="Suarez, F. J. (1989) Notas sobre algunos Tricholabioides (Hymenoptera, Mutillidae). Eos, Madrid, 65 (1), 125 - 188, 34 figs." type="journal article" year="1989">Suárez 1989</bibRefCitation>
: 185
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, ♂.
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</paragraph>
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<paragraph id="537D3684FF887135FEFD887CFE68EAFD" blockId="45.[264,619,623,689]" box="[264,486,623,648]" pageId="45" pageNumber="46">
<emphasis id="61B6EA96FF887135FEFD887CFE68EAFD" box="[264,486,623,648]" italics="true" pageId="45" pageNumber="46">Material examined</emphasis>
</paragraph>
<paragraph id="537D3684FF887135FECD8884FDE5EAC4" blockId="45.[264,619,623,689]" box="[312,619,663,689]" pageId="45" pageNumber="46">Not found in our material.</paragraph>
</subSubSection>
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<emphasis id="61B6EA96FF887135FEFD88F4FE19EB75" box="[264,407,743,768]" italics="true" pageId="45" pageNumber="46">Distribution</emphasis>
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<collectingCountry id="2BD57614FF887135FECD891CFE03EB5C" box="[312,397,783,809]" name="Yemen" pageId="45" pageNumber="46">Yemen</collectingCountry>
,
<collectingCountry id="2BD57614FF887135FE6D891CFDBAEB5C" box="[408,564,783,809]" name="Yemen" pageId="45" pageNumber="46">South Arabia</collectingCountry>
(
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).
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110
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*
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<emphasis id="61B6EA96FF897134FEE2898FFDB5EBC3" bold="true" box="[279,571,924,950]" italics="true" pageId="44" pageNumber="45">Tricholabiodes niloticus</emphasis>
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<taxonomicName id="94C24D07FF897134FEFD89F4FE21EB8B" box="[264,431,999,1022]" class="Insecta" family="Mutillidae" genus="Tricholabioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="61B6EA96FF897134FEFD89F4FE21EB8B" box="[264,431,999,1022]" italics="true" pageId="44" pageNumber="45">Tricholabioides</emphasis>
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(!)
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<emphasis id="61B6EA96FF897134FE2889F4FDB7EB8B" box="[477,569,999,1022]" italics="true" pageId="44" pageNumber="45">niloticus</emphasis>
Suárez 1967: 569
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, &quot;Helwan, Eg.,
<date id="277C1044FF897134FBB189F5FB14EB88" box="[1092,1178,998,1021]" pageId="44" pageNumber="45" value="1933-07-17">17­7­33</date>
, W. Wittner&quot; (
<collectingCountry id="2BD57614FF897134FECA8E15FE0FEC68" box="[319,385,1030,1053]" name="Egypt" pageId="44" pageNumber="45">Egypt</collectingCountry>
) [MSNG]; 1989: 144, ♂.
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<collectingCountry id="2BD57614FF897134FECD8E64FE00ECE4" box="[312,398,1143,1169]" name="Yemen" pageId="44" pageNumber="45">Yemen</collectingCountry>
: Al Kowd,
<date id="277C1044FF897134FDD18E64FD06ECE4" box="[548,648,1143,1169]" pageId="44" pageNumber="45" value="1999-09">IX.1999</date>
, LT, AvH &amp; S. Al Haruri, 1 ɗ [
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NW Manakhah,
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, MT, AvH, 1 ɗ [
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<emphasis id="61B6EA96FF897134FEFD8EFCFE19ED7D" box="[264,407,1263,1288]" italics="true" pageId="44" pageNumber="45">Distribution</emphasis>
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<collectingCountry id="2BD57614FF897134FECD8F04FE04ED44" box="[312,394,1303,1329]" name="Yemen" pageId="44" pageNumber="45">Yemen</collectingCountry>
(new record),
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(
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132
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*
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<emphasis id="61B6EA96FF897134FEE28F9DFD18EDDD" bold="true" box="[279,662,1422,1448]" italics="true" pageId="44" pageNumber="45">Tricholabiodes pedunculatoides</emphasis>
Bischoff 1920
</taxonomicName>
</emphasis>
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<emphasis id="61B6EA96FF897134FEFD8FC9FD53ED84" box="[264,733,1498,1521]" italics="true" pageId="44" pageNumber="45">Tricholabiodes semistriata pedunculatoides</emphasis>
Bischoff 1920: 105
</taxonomicName>
, ♂,
<typeStatus id="8C798826FF897134FC178FCAFBCCED85" box="[994,1090,1497,1520]" pageId="44" pageNumber="45" type="holotype">holotype</typeStatus>
, ♂ &quot;
<collectingCountry id="2BD57614FF897134FB858FCAFB4FED85" box="[1136,1217,1497,1520]" name="Somalia" pageId="44" pageNumber="45">Somali</collectingCountry>
, Damaso, B.V. Erlanger&quot; [ZMHU].
</paragraph>
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<emphasis id="61B6EA96FF897134FEFD8C09FE21EE44" box="[264,431,1562,1585]" italics="true" pageId="44" pageNumber="45">Tricholabioides</emphasis>
</taxonomicName>
(!)
<taxonomicName id="94C24D07FF897134FE228C09FCDEEE45" authority="Suarez 1989: 161" authorityName="Suarez" authorityPageNumber="161" authorityYear="1989" box="[471,848,1561,1585]" class="Insecta" family="Mutillidae" genus="Tricholabiodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="pedunculatoides">
<emphasis id="61B6EA96FF897134FE228C09FD08EE44" box="[471,646,1562,1585]" italics="true" pageId="44" pageNumber="45">pedunculatoides</emphasis>
:
<treatmentCitation id="D2631095FF897134FD618C0AFCDEEE45" author="Suarez" box="[660,848,1561,1584]" page="161" pageId="44" pageNumber="45" year="1989">
<bibRefCitation id="37534B75FF897134FD618C0AFC92EE45" author="Suarez" box="[660,796,1561,1584]" pageId="44" pageNumber="50" refString="Suarez, F. J. (1989) Notas sobre algunos Tricholabioides (Hymenoptera, Mutillidae). Eos, Madrid, 65 (1), 125 - 188, 34 figs." type="journal article" year="1989">Suárez 1989</bibRefCitation>
: 161
</treatmentCitation>
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, ♂.
</treatmentCitationGroup>
</paragraph>
<paragraph id="537D3684FF897134FEFD8C29FC07EE25" blockId="44.[264,1323,1497,1617]" box="[264,905,1593,1617]" pageId="44" pageNumber="45">
<taxonomicName id="94C24D07FF897134FEFD8C29FC8AEE25" authority="Magretti 1906: 39" authorityName="Magretti" authorityPageNumber="39" authorityYear="1906" box="[264,772,1593,1617]" class="Insecta" family="Mutillidae" genus="Tricholabiodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="pedunculata">
<emphasis id="61B6EA96FF897134FEFD8C29FDBCEE24" box="[264,562,1594,1617]" italics="true" pageId="44" pageNumber="45">Tricholabiodes pedunculata</emphasis>
: Magretti 1906: 39
</taxonomicName>
, ♂ (
<collectingCountry id="2BD57614FF897134FCDB8C2AFCF3EE25" box="[814,893,1593,1616]" name="Eritrea" pageId="44" pageNumber="45">Eritrea</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF897134FEFD8CBAFE18EF4E" pageId="44" pageNumber="45" type="materials_examined">
<paragraph id="537D3684FF897134FEFD8CBAFE68EEB7" blockId="44.[264,1322,1705,1851]" box="[264,486,1705,1730]" pageId="44" pageNumber="45">
<emphasis id="61B6EA96FF897134FEFD8CBAFE68EEB7" box="[264,486,1705,1730]" italics="true" pageId="44" pageNumber="45">Material examined</emphasis>
</paragraph>
<paragraph id="537D3684FF897134FECD8CC2FE18EF4E" blockId="44.[264,1322,1705,1851]" pageId="44" pageNumber="45">
<collectingCountry id="2BD57614FF897134FECD8CC2FE00EE9E" box="[312,398,1745,1771]" name="Yemen" pageId="44" pageNumber="45">Yemen</collectingCountry>
: Ta’izz,
<date id="277C1044FF897134FE1A8CC2FDE8EE9E" box="[495,614,1745,1771]" pageId="44" pageNumber="45" value="2000-08">VIII.2000</date>
, LT, AvH &amp; A.R. Yarimi, 1 ɗ [
<collectionCode id="35D3AE41FF897134FC388CC2FBA3EE9E" LSID="urn:lsid:biocol.org:col:34975" box="[973,1069,1745,1771]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="44" pageNumber="45">ZMAN</collectionCode>
]; Al Kowd,
<date id="277C1044FF897134FB4D8CC2FAA8EE9E" box="[1208,1318,1745,1771]" pageId="44" pageNumber="45" value="2000-12">XII.2000</date>
, LT, AvH &amp; S. Al Haruri, 1 ɗ [
<collectionCode id="35D3AE41FF897134FD908CEAFD24EF66" box="[613,682,1785,1811]" pageId="44" pageNumber="45">IBSS</collectionCode>
]; Suq Bani Mansour,
<date id="277C1044FF897134FC508CEAFB1DEF66" box="[933,1171,1785,1811]" pageId="44" pageNumber="45" value="2001-08-28" valueMax="2001-11-14" valueMin="2001-08-28">28.VIII–14.XI.2001</date>
, MT, AvH, 1 ɗ [
<collectionCode id="35D3AE41FF897134FEDF8D32FE05EF4E" LSID="urn:lsid:biocol.org:col:34992" box="[298,395,1825,1851]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="44" pageNumber="45">RMNH</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF887135FEFD8B0FFB97E92B" box="[264,1049,284,350]" pageId="45" pageNumber="46" type="distribution">
<paragraph id="537D3684FF887135FEFD8B0FFE19E940" blockId="45.[264,1049,284,350]" box="[264,407,284,309]" pageId="45" pageNumber="46">
<emphasis id="61B6EA96FF887135FEFD8B0FFE19E940" box="[264,407,284,309]" italics="true" pageId="45" pageNumber="46">Distribution</emphasis>
</paragraph>
<paragraph id="537D3684FF887135FECD8B57FB97E92B" blockId="45.[264,1049,284,350]" box="[312,1049,324,350]" pageId="45" pageNumber="46">
<collectingCountry id="2BD57614FF887135FECD8B57FE04E92B" box="[312,394,324,350]" name="Yemen" pageId="45" pageNumber="46">Yemen</collectingCountry>
(new record),
<collectingCountry id="2BD57614FF887135FDC38B57FD17E92B" box="[566,665,324,350]" name="Somalia" pageId="45" pageNumber="46">Somalia</collectingCountry>
,
<collectingCountry id="2BD57614FF887135FD508B57FD7BE92B" box="[677,757,324,350]" name="Kenya" pageId="45" pageNumber="46">Kenya</collectingCountry>
,
<collectingCountry id="2BD57614FF887135FCF48B57FCEAE92B" box="[769,868,324,350]" name="Ethiopia" pageId="45" pageNumber="46">Ethiopia</collectingCountry>
(
<bibRefCitation id="37534B75FF887135FC818B57FB85E92B" author="Suarez" box="[884,1035,324,350]" pageId="45" pageNumber="50" refString="Suarez, F. J. (1989) Notas sobre algunos Tricholabioides (Hymenoptera, Mutillidae). Eos, Madrid, 65 (1), 125 - 188, 34 figs." type="journal article" year="1989">Suárez 1989</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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108
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Key to the species of
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1. Head and mesosoma ferruginous­red. Forewing bright yellow in proximal half and dark fuscous in distal half. 10.5–12.0 mm. (
<collectingCountry id="2BD57614FF8C7131FCC38D44FC02EF04" box="[822,908,1879,1905]" name="Yemen" pageId="41" pageNumber="42">Yemen</collectingCountry>
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..............................
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<taxonomicNameLabel id="7A8557EDFF8C7131FB388D45FAA4EF05" box="[1229,1322,1878,1904]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
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<emphasis id="61B6EA96FF8F7132FEFD8B08FE9CE940" bold="true" box="[264,274,283,309]" pageId="42" pageNumber="43">­</emphasis>
Head and mesosoma black. Forewing dark fuscous with basal quarter yellowish or hyaline ........................................................................................................................... 2
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<paragraph id="537D3684FF8F7132FEFD8B7FFAA4E9D8" blockId="42.[264,1324,283,510]" pageId="42" pageNumber="43">
<keyLead id="E83395B1FF8F7132FEFD8B7FFAA4E9D8" pageId="42" pageNumber="43">
2. Basal quarter of forewing hyaline. Metasoma yellowish­red including apical segments. 10.0–12.0 mm. (
<collectingCountry id="2BD57614FF8F7132FDB88B87FD2DE9DB" box="[589,675,404,430]" name="Yemen" pageId="42" pageNumber="43">Yemen</collectingCountry>
)
<emphasis id="61B6EA96FF8F7132FD458B87FB48E9D8" box="[688,1222,404,429]" italics="true" pageId="42" pageNumber="43">
.........................................................
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<emphasis id="61B6EA96FF8F7132FB388B80FAA4E9D8" bold="true" box="[1229,1322,403,429]" pageId="42" pageNumber="43">sp. nov.</emphasis>
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<emphasis id="61B6EA96FF8F7132FEFD8BA8FE9CE9A0" bold="true" box="[264,274,443,469]" pageId="42" pageNumber="43">­</emphasis>
Basal quarter of forewing yellowish. Metasoma black with ferruginous­red segments 1 and 2. 12.0 mm. (
<collectingCountry id="2BD57614FF8F7132FDF38BF7FDC8E98B" box="[518,582,484,510]" name="India" pageId="42" pageNumber="43">India</collectingCountry>
,
<collectingCountry id="2BD57614FF8F7132FDA78BF7FD36E98B" box="[594,696,484,510]" name="Pakistan" pageId="42" pageNumber="43">Pakistan</collectingCountry>
,
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)
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<emphasis id="61B6EA96FF8F7132FB848BF7FB44E988" box="[1137,1226,484,509]" italics="true" pageId="42" pageNumber="43">V. dives</emphasis>
(Smith)
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117
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Lelej
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<emphasis id="61B6EA96FF8D7130FEFD8D0FFEF1EF40" box="[264,383,1820,1845]" italics="true" pageId="40" pageNumber="41">Diagnosis</emphasis>
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MALE. Body length 10.0–12.0 mm. Eyes projecting over head. POD: OOD 0.85X. Relation of pedicel and first three flagellomeres 0.6: 1.5: 1.8: 1.9. Clypeus slightly concave on median elevated part, delimited laterally by an obtuse carina. Mid coxa without lateral longitudinal carina. Parascutal carinae developed.
<collectingCountry id="2BD57614FF8C7131FBEA8BAFFBD4E9A3" box="[1055,1114,444,470]" name="India" pageId="41" pageNumber="42">Hind</collectingCountry>
coxa inside with carina which ends in a well­developed tubercle. Carina on S1 posteriorly with additional small acute carina. Genitalia are similar to those of
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species. Head and mesosoma densely punctured dorsally. Propodeum dorsally, laterally, and posteriorly reticulate. T2 and S2 densely punctate, punctures larger on S2 and sparser on disc of T2. T7 with median longitudinal glabrous gibbosity, which ends before preapical transversal carina.
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ɗ,
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The specific epithet is a noun derived from
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, or Tehama, a narrow, extremely arid lowland region along the West Coast of the Arabian Peninsula, from the Gulf of Aqaba south to the Bab el Mandeb straits, and refers to the area where the species has been collected.
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Suárez 1967
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, ♂,
<typeStatus id="8C798826FF8E7133FCD689EAFC0DEC65" box="[803,899,1017,1040]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
, ♂ &quot;de Uadi el Bark, Hadhramaut (Arabia meridional.),
<date id="277C1044FF8E7133FE068E0AFD13EC45" box="[499,669,1049,1072]" pageId="43" pageNumber="44" value="1962-03-28" valueMax="1962-03-29" valueMin="1962-03-28">28–29.III. 1962</date>
, G. Scortecci&quot; [Museo Civico di Storia Naturale di Milano]; 1989: 126, ♂ (
<collectingCountry id="2BD57614FF8E7133FE258E2AFD91EC25" box="[464,543,1081,1104]" name="Yemen" pageId="43" pageNumber="44">Yemen</collectingCountry>
,
<collectingCountry id="2BD57614FF8E7133FDDF8E2AFDFEEC25" box="[554,624,1081,1104]" name="Oman" pageId="43" pageNumber="44">Oman</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF8E7133FEFD8E49FCFFECC5" pageId="43" pageNumber="44" type="reference_group">
<paragraph id="537D3684FF8E7133FEFD8E49FCCDEC05" blockId="43.[264,1323,1017,1201]" box="[264,835,1113,1137]" pageId="43" pageNumber="44">
<treatmentCitationGroup id="73D211AAFF8E7133FEFD8E49FCCDEC05" box="[264,835,1113,1137]" pageId="43" pageNumber="44">
<taxonomicName id="94C24D07FF8E7133FEFD8E49FD33EC05" authority="Gribodo 1884: 390" authorityName="Gribodo" authorityPageNumber="390" authorityYear="1884" box="[264,701,1113,1137]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="pedunculata">
<emphasis id="61B6EA96FF8E7133FEFD8E49FE6FEC04" box="[264,481,1114,1137]" italics="true" pageId="43" pageNumber="44">Mutilla pedunculata</emphasis>
:
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<bibRefCitation id="37534B75FF8E7133FE1A8E4AFD06EC05" author="Gribodo" box="[495,648,1113,1136]" pageId="43" pageNumber="47" refString="Gribodo, G. (1884) Viaggio ad Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso &quot; Esploratore &quot; dal 16 Novembre, 1879 al 26 Febbraio 1880. III. Imenotteri. Annali del Museo Civico di Storia Naturale di Genova, 20, 381 - 392. [Mutillidae - p. 389 - 392]." type="journal article" year="1884">Gribodo 1884</bibRefCitation>
: 390
</treatmentCitation>
</taxonomicName>
, ♂ (
<collectingCountry id="2BD57614FF8E7133FD128E4AFCB6EC05" box="[743,824,1113,1136]" name="Yemen" pageId="43" pageNumber="44">Yemen</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="537D3684FF8E7133FEFD8E69FCFCECE5" blockId="43.[264,1323,1017,1201]" box="[264,882,1145,1169]" pageId="43" pageNumber="44">
<treatmentCitationGroup id="73D211AAFF8E7133FEFD8E69FCFCECE5" box="[264,882,1145,1169]" pageId="43" pageNumber="44">
<taxonomicName id="94C24D07FF8E7133FEFD8E69FD62ECE5" authority="Invrea 1965b: 61" authorityName="Invrea" authorityPageNumber="61" authorityYear="1965" box="[264,748,1145,1169]" class="Insecta" family="Mutillidae" genus="Tricholabiodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="semistriata">
<emphasis id="61B6EA96FF8E7133FEFD8E69FDABECE4" box="[264,549,1146,1169]" italics="true" pageId="43" pageNumber="44">Tricholabiodes semistriata</emphasis>
:
<treatmentCitation id="D2631095FF8E7133FDC68E6AFD62ECE5" author="Invrea" box="[563,748,1145,1168]" page="61" pageId="43" pageNumber="44" year="1965">
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: 61
</treatmentCitation>
</taxonomicName>
, ♂ (
<collectingCountry id="2BD57614FF8E7133FCE38E6AFCE9ECE5" box="[790,871,1145,1168]" name="Yemen" pageId="43" pageNumber="44">Yemen</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="537D3684FF8E7133FEFD8E89FCFFECC5" blockId="43.[264,1323,1017,1201]" box="[264,881,1177,1201]" pageId="43" pageNumber="44">
<treatmentCitationGroup id="73D211AAFF8E7133FEFD8E89FCFFECC5" box="[264,881,1177,1201]" pageId="43" pageNumber="44">
<taxonomicName id="94C24D07FF8E7133FEFD8E89FD64ECC5" authority="Invrea 1965b: 62" authorityName="Invrea" authorityPageNumber="62" authorityYear="1965" box="[264,746,1177,1201]" class="Insecta" family="Mutillidae" genus="Tricholabiodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="aegyptiaca">
<emphasis id="61B6EA96FF8E7133FEFD8E89FDADECC4" box="[264,547,1178,1201]" italics="true" pageId="43" pageNumber="44">Tricholabiodes aegyptiaca</emphasis>
:
<treatmentCitation id="D2631095FF8E7133FDC78E8AFD64ECC5" author="Invrea" box="[562,746,1177,1200]" page="62" pageId="43" pageNumber="44" year="1965">
<bibRefCitation id="37534B75FF8E7133FDC78E8AFD4DECC5" author="Invrea" box="[562,707,1177,1200]" pageId="43" pageNumber="48" refString="Invrea, F. (1965 b) Missione 1962 del Prof. Giuseppe Scortecci nell'Arabia meridionale (Hymenoptera: Apterogynidae, Mutillidae et Chrysididae). Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano, 104 (1), 59 - 66." type="journal article" year="1965" yearSuffix="b">Invrea 1965b</bibRefCitation>
: 62
</treatmentCitation>
</taxonomicName>
, ♂ (
<collectingCountry id="2BD57614FF8E7133FCE08E8AFCE8ECC5" box="[789,870,1177,1200]" name="Yemen" pageId="43" pageNumber="44">Yemen</collectingCountry>
).
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</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF8E7133FEFD8EF2FEF7EDEE" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="537D3684FF8E7133FEFD8EF2FE68EC8F" blockId="43.[264,1323,1249,1435]" box="[264,486,1249,1274]" pageId="43" pageNumber="44">
<emphasis id="61B6EA96FF8E7133FEFD8EF2FE68EC8F" box="[264,486,1249,1274]" italics="true" pageId="43" pageNumber="44">Material examined</emphasis>
</paragraph>
<paragraph id="537D3684FF8E7133FECD8F1AFEF7EDEE" blockId="43.[264,1323,1249,1435]" pageId="43" pageNumber="44">
<collectingCountry id="2BD57614FF8E7133FECD8F1AFE01ED56" box="[312,399,1289,1315]" name="Yemen" pageId="43" pageNumber="44">Yemen</collectingCountry>
: Al Kowd,
<date id="277C1044FF8E7133FDED8F1AFD01ED56" box="[536,655,1289,1315]" pageId="43" pageNumber="44" value="1999-08">VIII.1999</date>
, LT, AvH &amp; S. Al Haruri, 3 ɗ [
<collectionCode id="35D3AE41FF8E7133FBF08F1AFBEFED56" LSID="urn:lsid:biocol.org:col:34975" box="[1029,1121,1289,1315]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="43" pageNumber="44">ZMAN</collectionCode>
,
<collectionCode id="35D3AE41FF8E7133FB9B8F1AFB3CED56" box="[1134,1202,1289,1315]" pageId="43" pageNumber="44">IBSS</collectionCode>
];
<date id="277C1044FF8E7133FB338F1AFAA8ED56" box="[1222,1318,1289,1315]" pageId="43" pageNumber="44" value="2000-04">IV.2000</date>
, LT, AvH &amp; S. Al Haruri, 1 ɗ [
<collectionCode id="35D3AE41FF8E7133FD988F22FD43ED3E" LSID="urn:lsid:biocol.org:col:34975" box="[621,717,1329,1355]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="43" pageNumber="44">ZMAN</collectionCode>
];
<date id="277C1044FF8E7133FD158F22FCC0ED3E" box="[736,846,1329,1355]" pageId="43" pageNumber="44" value="2000-07">VII.2000</date>
, LT, AvH &amp; S. Al Haruri, 1 ɗ [
<collectionCode id="35D3AE41FF8E7133FB358F22FA91ED3E" LSID="urn:lsid:biocol.org:col:34975" box="[1216,1311,1329,1355]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="43" pageNumber="44">ZMAN</collectionCode>
];
<date id="277C1044FF8E7133FEFD8F4AFEF1ED06" box="[264,383,1369,1395]" pageId="43" pageNumber="44" value="2000-08">VIII.2000</date>
, LT, AvH &amp; S. Al Haruri, 1 ɗ [
<collectionCode id="35D3AE41FF8E7133FD098F4AFCD2ED06" LSID="urn:lsid:biocol.org:col:34975" box="[764,860,1369,1395]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="43" pageNumber="44">ZMAN</collectionCode>
];
<date id="277C1044FF8E7133FC858F4AFC30ED06" box="[880,958,1369,1395]" pageId="43" pageNumber="44" value="2001-01">I.2001</date>
, LT, AvH &amp; S. Al Haruri, 1 ɗ [
<collectionCode id="35D3AE41FF8E7133FEFA8F92FEE0EDEE" LSID="urn:lsid:biocol.org:col:34975" box="[271,366,1409,1435]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="43" pageNumber="44">ZMAN</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF8E7133FEFD8FC2FD1AEE66" box="[264,660,1489,1555]" pageId="43" pageNumber="44" type="distribution">
<paragraph id="537D3684FF8E7133FEFD8FC2FE19ED9F" blockId="43.[264,660,1489,1555]" box="[264,407,1489,1514]" pageId="43" pageNumber="44">
<emphasis id="61B6EA96FF8E7133FEFD8FC2FE19ED9F" box="[264,407,1489,1514]" italics="true" pageId="43" pageNumber="44">Distribution</emphasis>
</paragraph>
<paragraph id="537D3684FF8E7133FECD8FEAFD1AEE66" blockId="43.[264,660,1489,1555]" box="[312,660,1529,1555]" pageId="43" pageNumber="44">
<collectingCountry id="2BD57614FF8E7133FECD8FEAFE03EE66" box="[312,397,1529,1555]" name="Yemen" pageId="43" pageNumber="44">Yemen</collectingCountry>
,
<collectingCountry id="2BD57614FF8E7133FE6D8FEAFE6EEE66" box="[408,480,1529,1555]" name="Oman" pageId="43" pageNumber="44">Oman</collectingCountry>
(
<bibRefCitation id="37534B75FF8E7133FE1A8FEAFD09EE66" author="Suarez" box="[495,647,1529,1555]" pageId="43" pageNumber="50" refString="Suarez, F. J. (1989) Notas sobre algunos Tricholabioides (Hymenoptera, Mutillidae). Eos, Madrid, 65 (1), 125 - 188, 34 figs." type="journal article" year="1989">Suárez 1989</bibRefCitation>
).
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</treatment>
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177
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<emphasis id="61B6EA96FF8E7133FEFD8CD2FC9FEEAE" bold="true" box="[264,785,1729,1755]" pageId="43" pageNumber="44">
<emphasis id="61B6EA96FF8E7133FEFD8CD2FDC4EEAE" bold="true" box="[264,586,1729,1755]" italics="true" pageId="43" pageNumber="44">Tricholabiodes chloroticus</emphasis>
(
<bibRefCitation id="37534B75FF8E7133FDAF8CD2FC87EEAE" author="Gribodo" box="[602,777,1729,1755]" pageId="43" pageNumber="47" refString="Gribodo, G. (1884) Viaggio ad Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso &quot; Esploratore &quot; dal 16 Novembre, 1879 al 26 Febbraio 1880. III. Imenotteri. Annali del Museo Civico di Storia Naturale di Genova, 20, 381 - 392. [Mutillidae - p. 389 - 392]." type="journal article" year="1884">Gribodo 1884</bibRefCitation>
)
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</subSubSection>
<subSubSection id="1BD8650FFF8E7133FEFD8D1FFBDEEF16" pageId="43" pageNumber="44" type="reference_group">
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<treatmentCitationGroup id="73D211AAFF8E7133FEFD8D1FFD3AEF36" pageId="43" pageNumber="44">
<taxonomicName id="94C24D07FF8E7133FEFD8D1FFD15EF56" authority="Gribodo 1884: 391" authorityName="Gribodo" authorityPageNumber="391" authorityYear="1884" box="[264,667,1804,1827]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="chlorotica">
<emphasis id="61B6EA96FF8E7133FEFD8D1FFE47EF56" box="[264,457,1804,1827]" italics="true" pageId="43" pageNumber="44">Mutilla chlorotica</emphasis>
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: 391
</treatmentCitation>
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, ♂,
<typeStatus id="8C798826FF8E7133FD318D1FFCA8EF56" box="[708,806,1804,1827]" pageId="43" pageNumber="44" type="lectotype">lectotype</typeStatus>
♂ &quot;
<collectingCountry id="2BD57614FF8E7133FCB98D1FFC14EF56" box="[844,922,1804,1827]" name="Yemen" pageId="43" pageNumber="44">Yemen</collectingCountry>
merid., Tes,
<date id="277C1044FF8E7133FBD68D1FFBE2EF56" box="[1059,1132,1804,1827]" pageId="43" pageNumber="44" value="1800-01">I­1800</date>
, R. Mazoni&quot;, designated
<treatmentCitation id="D2631095FF8E7133FE788D3FFDC6EF36" author="Suarez" box="[397,584,1836,1859]" page="131" pageId="43" pageNumber="44" year="1989">
<bibRefCitation id="37534B75FF8E7133FE788D3FFD9BEF36" author="Suarez" box="[397,533,1836,1859]" pageId="43" pageNumber="50" refString="Suarez, F. J. (1989) Notas sobre algunos Tricholabioides (Hymenoptera, Mutillidae). Eos, Madrid, 65 (1), 125 - 188, 34 figs." type="journal article" year="1989">Suárez 1989</bibRefCitation>
: 131
</treatmentCitation>
[MSNG].
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</paragraph>
<paragraph id="537D3684FF8E7133FEFD8D5FFBDEEF16" blockId="43.[264,1323,1803,1891]" box="[264,1104,1867,1891]" pageId="43" pageNumber="44">
<treatmentCitationGroup id="73D211AAFF8E7133FEFD8D5FFBDEEF16" box="[264,1104,1867,1891]" pageId="43" pageNumber="44">
<taxonomicName id="94C24D07FF8E7133FEFD8D5FFD6FEF16" authority="Invrea 1965b: 64" authorityName="Invrea" authorityPageNumber="64" authorityYear="1965" box="[264,737,1868,1891]" class="Insecta" family="Mutillidae" genus="Tricholabiodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="chlorotica">
<emphasis id="61B6EA96FF8E7133FEFD8D5FFD95EF16" box="[264,539,1868,1891]" italics="true" pageId="43" pageNumber="44">Tricholabiodes chlorotica</emphasis>
:
<treatmentCitation id="D2631095FF8E7133FDDC8D5FFD6FEF16" author="Invrea" box="[553,737,1868,1891]" page="64" pageId="43" pageNumber="44" year="1965">
<bibRefCitation id="37534B75FF8E7133FDDC8D5FFD34EF16" author="Invrea" box="[553,698,1868,1891]" pageId="43" pageNumber="48" refString="Invrea, F. (1965 b) Missione 1962 del Prof. Giuseppe Scortecci nell'Arabia meridionale (Hymenoptera: Apterogynidae, Mutillidae et Chrysididae). Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano, 104 (1), 59 - 66." type="journal article" year="1965" yearSuffix="b">Invrea 1965b</bibRefCitation>
: 64
</treatmentCitation>
</taxonomicName>
, ♂ (
<collectingCountry id="2BD57614FF8E7133FCF88D5FFCD0EF16" box="[781,862,1868,1891]" name="Yemen" pageId="43" pageNumber="44">Yemen</collectingCountry>
);
<treatmentCitation id="D2631095FF8E7133FC9A8D5FFBA2EF16" author="Suarez" box="[879,1068,1868,1891]" page="131" pageId="43" pageNumber="44" year="1989">
<bibRefCitation id="37534B75FF8E7133FC9A8D5FFC76EF16" author="Suarez" box="[879,1016,1868,1891]" pageId="43" pageNumber="50" refString="Suarez, F. J. (1989) Notas sobre algunos Tricholabioides (Hymenoptera, Mutillidae). Eos, Madrid, 65 (1), 125 - 188, 34 figs." type="journal article" year="1989">Suárez 1989</bibRefCitation>
: 131
</treatmentCitation>
, ♂.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF897134FEFD8B0FFCDCE9A3" pageId="44" pageNumber="45" type="materials_examined">
<paragraph id="537D3684FF897134FEFD8B0FFE68E940" blockId="44.[264,1323,284,470]" box="[264,486,284,309]" pageId="44" pageNumber="45">
<emphasis id="61B6EA96FF897134FEFD8B0FFE68E940" box="[264,486,284,309]" italics="true" pageId="44" pageNumber="45">Material examined</emphasis>
</paragraph>
<paragraph id="537D3684FF897134FECD8B57FCDCE9A3" blockId="44.[264,1323,284,470]" pageId="44" pageNumber="45">
<collectingCountry id="2BD57614FF897134FECD8B57FE00E92B" box="[312,398,324,350]" name="Yemen" pageId="44" pageNumber="45">Yemen</collectingCountry>
: Al Kowd,
<date id="277C1044FF897134FDE18B57FDF9E92B" box="[532,631,324,350]" pageId="44" pageNumber="45" value="1999-09">IX.1999</date>
, LT, AvH &amp; S. Al Haruri, 1 ɗ [
<collectionCode id="35D3AE41FF897134FC118B57FBCDE92B" LSID="urn:lsid:biocol.org:col:34975" box="[996,1091,324,350]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="44" pageNumber="45">ZMAN</collectionCode>
];
<date id="277C1044FF897134FBA38B57FB34E92B" box="[1110,1210,324,350]" pageId="44" pageNumber="45" value="2000-11">XI.2000</date>
, LT, AvH &amp; S. Al Haruri, 1 ɗ [
<collectionCode id="35D3AE41FF897134FDF28B7FFDE9E9F3" LSID="urn:lsid:biocol.org:col:34975" box="[519,615,364,390]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="44" pageNumber="45">ZMAN</collectionCode>
];
<date id="277C1044FF897134FD8E8B7FFD66E9F3" box="[635,744,364,390]" pageId="44" pageNumber="45" value="2000-12">XII.2000</date>
, LT, AvH &amp; S. Al Haruri, 1 ɗ [
<collectionCode id="35D3AE41FF897134FB968B7FFB4DE9F3" LSID="urn:lsid:biocol.org:col:34975" box="[1123,1219,364,390]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="44" pageNumber="45">ZMAN</collectionCode>
];
<date id="277C1044FF897134FB228B7FFAA8E9F3" box="[1239,1318,364,390]" pageId="44" pageNumber="45" value="2001-01">I.2001</date>
, LT, AvH &amp; S. Al Haruri, 1 ɗ [
<collectionCode id="35D3AE41FF897134FD8B8B87FD4AE9DB" box="[638,708,404,430]" pageId="44" pageNumber="45">IBSS</collectionCode>
];
<date id="277C1044FF897134FD2F8B87FCB2E9DB" box="[730,828,404,430]" pageId="44" pageNumber="45" value="2001-02">II. 2001</date>
, LT, AvH &amp; S. Al Haruri, 1 ɗ [
<collectionCode id="35D3AE41FF897134FB358B87FA91E9DB" LSID="urn:lsid:biocol.org:col:34975" box="[1216,1311,404,430]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="44" pageNumber="45">ZMAN</collectionCode>
];
<collectingCountry id="2BD57614FF897134FEFD8BAFFE2BE9A3" box="[264,421,444,470]" name="Saudi Arabia" pageId="44" pageNumber="45">Saudi Arabia</collectingCountry>
, 1 ɗ (coll. Schmid­Egger,
<collectingCountry id="2BD57614FF897134FD298BAFFCC2E9A3" box="[732,844,444,470]" name="Germany" pageId="44" pageNumber="45">Germany</collectingCountry>
]
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF897134FEFD881FFC0AEA3B" box="[264,900,524,590]" pageId="44" pageNumber="45" type="distribution">
<paragraph id="537D3684FF897134FEFD881FFE19EA50" blockId="44.[264,900,524,590]" box="[264,407,524,549]" pageId="44" pageNumber="45">
<emphasis id="61B6EA96FF897134FEFD881FFE19EA50" box="[264,407,524,549]" italics="true" pageId="44" pageNumber="45">Distribution</emphasis>
</paragraph>
<paragraph id="537D3684FF897134FECD8827FC0AEA3B" blockId="44.[264,900,524,590]" box="[312,900,564,590]" pageId="44" pageNumber="45">
<collectingCountry id="2BD57614FF897134FECD8827FE04EA3B" box="[312,394,564,590]" name="Yemen" pageId="44" pageNumber="45">Yemen</collectingCountry>
(
<bibRefCitation id="37534B75FF897134FE6C8827FDBEEA3B" author="Suarez" box="[409,560,564,590]" pageId="44" pageNumber="50" refString="Suarez, F. J. (1989) Notas sobre algunos Tricholabioides (Hymenoptera, Mutillidae). Eos, Madrid, 65 (1), 125 - 188, 34 figs." type="journal article" year="1989">Suárez 1989</bibRefCitation>
),
<collectingCountry id="2BD57614FF897134FDB08827FD51EA3B" box="[581,735,564,590]" name="Saudi Arabia" pageId="44" pageNumber="45">Saudi Arabia</collectingCountry>
(new record).
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFF897134FEFD8897FED0EB4B" pageId="44" pageNumber="45" type="discussion">
<paragraph id="537D3684FF897134FEFD8897FEE0EAE8" blockId="44.[264,1323,644,830]" box="[264,366,644,669]" pageId="44" pageNumber="45">
<emphasis id="61B6EA96FF897134FEFD8897FEE0EAE8" box="[264,366,644,669]" italics="true" pageId="44" pageNumber="45">Remarks</emphasis>
</paragraph>
<paragraph id="537D3684FF897134FECD88BFFED0EB4B" blockId="44.[264,1323,644,830]" pageId="44" pageNumber="45">
Bischoff (1920) erroneously synonymized
<taxonomicName id="94C24D07FF897134FCBE88BFFB66EAB3" authority="Gribodo, 1884" authorityName="Gribodo" authorityYear="1884" box="[843,1256,684,710]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="chlorotica">
<emphasis id="61B6EA96FF897134FCBE88BFFBA9EAB0" box="[843,1063,684,709]" italics="true" pageId="44" pageNumber="45">Mutilla chlorotica</emphasis>
<bibRefCitation id="37534B75FF897134FBC388BFFB66EAB3" author="Gribodo" box="[1078,1256,684,710]" pageId="44" pageNumber="47" refString="Gribodo, G. (1884) Viaggio ad Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso &quot; Esploratore &quot; dal 16 Novembre, 1879 al 26 Febbraio 1880. III. Imenotteri. Annali del Museo Civico di Storia Naturale di Genova, 20, 381 - 392. [Mutillidae - p. 389 - 392]." type="journal article" year="1884">Gribodo, 1884</bibRefCitation>
</taxonomicName>
with
<taxonomicName id="94C24D07FF897134FEFD88C7FDF3EA9B" authority="Klug 1829" authorityName="Klug" authorityYear="1829" box="[264,637,724,750]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="61B6EA96FF897134FEFD88C7FE5BEA98" box="[264,469,724,749]" italics="true" pageId="44" pageNumber="45">Mutilla testacea</emphasis>
<bibRefCitation id="37534B75FF897134FE1888C7FDF3EA9B" author="Klug" box="[493,637,724,750]" pageId="44" pageNumber="48" refString="Klug, J. C. F. (1829) Symbolae physicae seu icones et descriptiones insectorum quae ex itinere per Africam borealem et Asiam occidentalem Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg studio novae aut illustratae redierunt. Hymenoptera. Decas Prima [genera Mutilla et Apterogyna]. Venditur a Mittlero, Berolini. [15 - 21] + tab. 4, 5." type="journal volume" year="1829">Klug 1829</bibRefCitation>
</taxonomicName>
. The former belongs to the genus
<taxonomicName id="94C24D07FF897134FB8E88C7FE75EB63" authority="Radoszkowski 1885" authorityName="Radoszkowski" authorityYear="1885" class="Insecta" family="Mutillidae" genus="Tricholabiodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="61B6EA96FF897134FB8E88C7FAA5EA98" box="[1147,1323,724,749]" italics="true" pageId="44" pageNumber="45">Tricholabiodes</emphasis>
<bibRefCitation id="37534B75FF897134FEFD88EFFE75EB63" author="Radoszkowski" box="[264,507,764,790]" pageId="44" pageNumber="49" refString="Radoszkowski, O. (1885) Revision des armures copulatrices des males de la famille de Mutillides. Horae Societatis Entomologicae Rossicae, 19 (1 - 2), 3 - 49, 9 pl." type="journal article" year="1885">Radoszkowski 1885</bibRefCitation>
</taxonomicName>
, the latter to the genus
<taxonomicName id="94C24D07FF897134FCD688EFFB9AEB63" authority="Lelej 1980" authorityName="Lelej" authorityYear="1980" box="[803,1044,764,790]" class="Insecta" family="Mutillidae" genus="Dentilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="61B6EA96FF897134FCD688EFFC0DEB60" box="[803,899,764,789]" italics="true" pageId="44" pageNumber="45">Dentilla</emphasis>
<bibRefCitation id="37534B75FF897134FC7B88EFFB9AEB63" author="Lelej" box="[910,1044,764,790]" pageId="44" pageNumber="48" refString="Lelej, A. S. &amp; Kabakov, O. N. (1980) On Bradynobaenidae and Mutillidae (Hymenoptera) of Afghanistan. Entomologicheskoe Obozrenie, 59 (1), 181 - 196, 14 figs. (In Russian). (English translation in Entomological Review, 59, 141 - 156)." type="journal article" year="1980">Lelej 1980</bibRefCitation>
</taxonomicName>
. Both species occur in
<collectingCountry id="2BD57614FF897134FEFD8937FED9EB4B" box="[264,343,804,830]" name="Yemen" pageId="44" pageNumber="45">Yemen</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

210
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<paragraph id="537D3684FFA3711EFEFD8CDDFC81EE9D" blockId="6.[264,783,1742,1803]" box="[264,783,1742,1768]" pageId="6" pageNumber="7">
<heading id="083581E8FFA3711EFEFD8CDDFC81EE9D" bold="true" box="[264,783,1742,1768]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="61B6EA96FFA3711EFEFD8CDDFC81EE9D" bold="true" box="[264,783,1742,1768]" pageId="6" pageNumber="7">
<emphasis id="61B6EA96FFA3711EFEFD8CDDFE9AEE9D" bold="true" box="[264,276,1742,1768]" italics="true" pageId="6" pageNumber="7">*</emphasis>
<taxonomicName id="94C24D07FFA3711EFEE18CDDFC81EE9D" authority="Invrea 1962" authorityName="Invrea" authorityYear="1962" box="[276,783,1742,1768]" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="mascatiana">
<emphasis id="61B6EA96FFA3711EFEE18CDDFDFDEE9D" bold="true" box="[276,627,1742,1768]" italics="true" pageId="6" pageNumber="7">Pseudophotopsis mascatiana</emphasis>
Invrea 1962
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="537D3684FFA3711EFEFD8CE2FE0CEF7E" blockId="6.[264,783,1742,1803]" box="[264,386,1777,1803]" pageId="6" pageNumber="7">
(
<figureCitation id="CBF92A01FFA3711EFEE58CE2FEF6EF7E" box="[272,376,1777,1803]" captionStart="FIGURES 2 – 10. 2 – 5" captionStartId="8.[264,382,1060,1084]" captionTargetBox="[541,1044,411,1033]" captionTargetId="figure@8.[535,1051,406,1036]" captionTargetPageId="8" captionText="FIGURES 2 – 10. 2 – 5. Pseudophotopsis mascatiana, male. 2. Forewing; 3. Head; 4. Metasomal tergum 1, dorsal view; 5. Mandible, lateral view. 6 – 8. Arnoldtilla achterbergi sp. nov., female, holotype. 6. Habitus, dorsal view; 7. Mandible, dorsal view; 8. Pedicel and flagellomeres 1 – 3. 9 – 10. Omotilla conjuncta, male. 9. Mandible, dorsal view; 10. Genitalia, ventral view. Scale = 1 mm for 2 – 4, 6 and 0.5 mm for 5, 7 – 10." httpUri="https://zenodo.org/record/172673/files/figure.png" pageId="6" pageNumber="7">Figs 2–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFA3711EFEFD8D24FB6CEF18" pageId="6" pageNumber="7" type="reference_group">
<paragraph id="537D3684FFA3711EFEFD8D24FB6CEF18" blockId="6.[264,1322,1846,1901]" pageId="6" pageNumber="7">
<taxonomicName id="94C24D07FFA3711EFEFD8D24FC9FEF38" authority="Invrea 1962: 324" authorityName="Invrea" authorityPageNumber="324" authorityYear="1962" box="[264,785,1846,1870]" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="mascatiana">
<emphasis id="61B6EA96FFA3711EFEFD8D24FDCDEF3B" box="[264,579,1847,1870]" italics="true" pageId="6" pageNumber="7">Pseudophotopsis mascatiana</emphasis>
Invrea 1962: 324
</taxonomicName>
, ♂,
<typeStatus id="8C798826FFA3711EFCB68D25FC2EEF38" box="[835,928,1846,1869]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
&quot;
<collectingCountry id="2BD57614FFA3711EFC418D25FB80EF38" box="[948,1038,1846,1869]" name="Somalia" pageId="6" pageNumber="7">Somalia</collectingCountry>
sett. (Migiurtinia), Passo Bocc.,
<quantity id="943A9B61FFA3711EFE768D45FE58EF18" box="[387,470,1878,1901]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="7" unit="m" value="1300.0">1300 m</quantity>
,
<date id="277C1044FFA3711EFE158D45FDD2EF18" box="[480,604,1878,1901]" pageId="6" pageNumber="7" value="1957-09-27">27.IX.1957</date>
, leg. Scortecci [Museo Civico di Storia Naturale di Milano].
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFA2711FFEFD8B0FFE24EA00" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="537D3684FFA2711FFEFD8B0FFEF1E940" blockId="7.[264,1323,284,550]" box="[264,383,284,309]" pageId="7" pageNumber="8">
<emphasis id="61B6EA96FFA2711FFEFD8B0FFEF1E940" box="[264,383,284,309]" italics="true" pageId="7" pageNumber="8">Diagnosis</emphasis>
</paragraph>
<paragraph id="537D3684FFA2711FFECD8B57FBE1EA53" blockId="7.[264,1323,284,550]" pageId="7" pageNumber="8">
The male of
<taxonomicName id="94C24D07FFA2711FFE188B57FCC3E928" box="[493,845,324,349]" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="mascatiana">
<emphasis id="61B6EA96FFA2711FFE188B57FCC3E928" box="[493,845,324,349]" italics="true" pageId="7" pageNumber="8">Pseudophotopsis mascatiana</emphasis>
</taxonomicName>
can be distinguished from those of congeneric species by very long metasomal segment 1. The male of
<taxonomicName id="94C24D07FFA2711FFB948B7FFA84E9F0" box="[1121,1290,364,389]" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="mascatiana">
<emphasis id="61B6EA96FFA2711FFB948B7FFA84E9F0" box="[1121,1290,364,389]" italics="true" pageId="7" pageNumber="8">P. mascatiana</emphasis>
</taxonomicName>
is similar to that of East Mediterranean
<taxonomicName id="94C24D07FFA2711FFD258B87FB82E9DB" authority="Smith 1855" authorityName="Smith" authorityYear="1855" box="[720,1036,404,430]" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="obliterata">
<emphasis id="61B6EA96FFA2711FFD258B87FCEAE9D8" box="[720,868,404,429]" italics="true" pageId="7" pageNumber="8">P. obliterata</emphasis>
(
<bibRefCitation id="37534B75FFA2711FFC838B87FB8AE9DB" author="Smith" box="[886,1028,404,430]" pageId="7" pageNumber="50" refString="Smith, F. (1855) Catalogue of Hymenopterous Insects in the Collection of the British Museum. Part III. Mutillidae and Pompilidae. Taylor and Francis, London. 206 pp. + 5 pls. [Mutillidae - p. l - 66, pl. 1]." type="book" year="1855">Smith 1855</bibRefCitation>
)
</taxonomicName>
in having similar sized ocelli and similar body coloration. It can be distinguished from the latter by having a much longer metasomal segment 1, well developed metanotal spines (very weak in
<taxonomicName id="94C24D07FFA2711FFB538BF7FA94E988" box="[1190,1306,484,509]" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="obliterata">
<emphasis id="61B6EA96FFA2711FFB538BF7FA94E988" box="[1190,1306,484,509]" italics="true" pageId="7" pageNumber="8">obliterata</emphasis>
</taxonomicName>
), and by a weak median basal tubercle on S2 (well developed in
<taxonomicName id="94C24D07FFA2711FFC1E881FFBD1EA50" box="[1003,1119,524,549]" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="obliterata">
<emphasis id="61B6EA96FFA2711FFC1E881FFBD1EA50" box="[1003,1119,524,549]" italics="true" pageId="7" pageNumber="8">obliterata</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="537D3684FFA2711FFEFD884FFE24EA00" blockId="7.[264,1323,604,1430]" box="[264,426,604,629]" pageId="7" pageNumber="8">
<emphasis id="61B6EA96FFA2711FFEFD884FFE24EA00" box="[264,426,604,629]" italics="true" pageId="7" pageNumber="8">Redescription</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFA2711FFECD8897FDB9EDE3" pageId="7" pageNumber="8" type="description">
<paragraph id="537D3684FFA2711FFECD8897FD40EC0B" blockId="7.[264,1323,604,1430]" pageId="7" pageNumber="8">
MALE. Body length 8.8–12.0 mm. Head length (from anterior clypeal border to occiput) 0.95–1.0X its maximal width. Ocelli large (
<figureCitation id="CBF92A01FFA2711FFC8488BFFC36EAB3" box="[881,952,684,710]" captionStart="FIGURES 2 – 10. 2 – 5" captionStartId="8.[264,382,1060,1084]" captionTargetBox="[541,1044,411,1033]" captionTargetId="figure@8.[535,1051,406,1036]" captionTargetPageId="8" captionText="FIGURES 2 – 10. 2 – 5. Pseudophotopsis mascatiana, male. 2. Forewing; 3. Head; 4. Metasomal tergum 1, dorsal view; 5. Mandible, lateral view. 6 – 8. Arnoldtilla achterbergi sp. nov., female, holotype. 6. Habitus, dorsal view; 7. Mandible, dorsal view; 8. Pedicel and flagellomeres 1 – 3. 9 – 10. Omotilla conjuncta, male. 9. Mandible, dorsal view; 10. Genitalia, ventral view. Scale = 1 mm for 2 – 4, 6 and 0.5 mm for 5, 7 – 10." httpUri="https://zenodo.org/record/172673/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
), POD: OOD 1.25–1.5X; ratio of distance between posterior ocellus and posterior head border to longitudinal posterior ocellus diameter 1.5–1.6X. Mandibles excised beneath, with tooth near base, distance between its apex and upper mandibular ridge less than mandibular basal height (
<figureCitation id="CBF92A01FFA2711FFB258937FA94EB4B" box="[1232,1306,804,830]" captionStart="FIGURES 2 – 10. 2 – 5" captionStartId="8.[264,382,1060,1084]" captionTargetBox="[541,1044,411,1033]" captionTargetId="figure@8.[535,1051,406,1036]" captionTargetPageId="8" captionText="FIGURES 2 – 10. 2 – 5. Pseudophotopsis mascatiana, male. 2. Forewing; 3. Head; 4. Metasomal tergum 1, dorsal view; 5. Mandible, lateral view. 6 – 8. Arnoldtilla achterbergi sp. nov., female, holotype. 6. Habitus, dorsal view; 7. Mandible, dorsal view; 8. Pedicel and flagellomeres 1 – 3. 9 – 10. Omotilla conjuncta, male. 9. Mandible, dorsal view; 10. Genitalia, ventral view. Scale = 1 mm for 2 – 4, 6 and 0.5 mm for 5, 7 – 10." httpUri="https://zenodo.org/record/172673/files/figure.png" pageId="7" pageNumber="8">Fig. 5</figureCitation>
). First flagellomere 2.0X its width and equal flagellomere 2. Metanotum with a pair of spines. Radial cell of forewing with acuminate apex. Forewing venation as in
<figureCitation id="CBF92A01FFA2711FFB448967FB74EBFB" box="[1201,1274,884,910]" captionStart="FIGURES 2 – 10. 2 – 5" captionStartId="8.[264,382,1060,1084]" captionTargetBox="[541,1044,411,1033]" captionTargetId="figure@8.[535,1051,406,1036]" captionTargetPageId="8" captionText="FIGURES 2 – 10. 2 – 5. Pseudophotopsis mascatiana, male. 2. Forewing; 3. Head; 4. Metasomal tergum 1, dorsal view; 5. Mandible, lateral view. 6 – 8. Arnoldtilla achterbergi sp. nov., female, holotype. 6. Habitus, dorsal view; 7. Mandible, dorsal view; 8. Pedicel and flagellomeres 1 – 3. 9 – 10. Omotilla conjuncta, male. 9. Mandible, dorsal view; 10. Genitalia, ventral view. Scale = 1 mm for 2 – 4, 6 and 0.5 mm for 5, 7 – 10." httpUri="https://zenodo.org/record/172673/files/figure.png" pageId="7" pageNumber="8">Fig. 2</figureCitation>
. T1 length 1.7–1.75X its maximal width (
<figureCitation id="CBF92A01FFA2711FFD4B898FFC88EBC3" box="[702,774,924,950]" captionStart="FIGURES 2 – 10. 2 – 5" captionStartId="8.[264,382,1060,1084]" captionTargetBox="[541,1044,411,1033]" captionTargetId="figure@8.[535,1051,406,1036]" captionTargetPageId="8" captionText="FIGURES 2 – 10. 2 – 5. Pseudophotopsis mascatiana, male. 2. Forewing; 3. Head; 4. Metasomal tergum 1, dorsal view; 5. Mandible, lateral view. 6 – 8. Arnoldtilla achterbergi sp. nov., female, holotype. 6. Habitus, dorsal view; 7. Mandible, dorsal view; 8. Pedicel and flagellomeres 1 – 3. 9 – 10. Omotilla conjuncta, male. 9. Mandible, dorsal view; 10. Genitalia, ventral view. Scale = 1 mm for 2 – 4, 6 and 0.5 mm for 5, 7 – 10." httpUri="https://zenodo.org/record/172673/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
). T2 length 0.95X its maximal width. Segment 2 with lateral felt lines on T2 and S2, their length on T2 less than distance between felt line and posterior tergal border. S2 with weak median basal tubercle. T2 shiny, densely punctate basally and laterally mixed with small punctures, disc sparsely punctate. T7 coarsely punctate. Hypopygium with median apical tubercle. Genitalia are similar to those of
<taxonomicName id="94C24D07FFA2711FFEDD8E77FE21EC08" box="[296,431,1124,1149]" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kabakovi">
<emphasis id="61B6EA96FFA2711FFEDD8E77FE21EC08" box="[296,431,1124,1149]" italics="true" pageId="7" pageNumber="8">P. kabakovi</emphasis>
</taxonomicName>
(
<bibRefCitation id="37534B75FFA2711FFE4B8E77FD32EC08" author="Lelej" box="[446,700,1124,1150]" pageId="7" pageNumber="48" refString="Lelej, A. S. (1985) Osy-nemki (Hymenoptera, Mutillidae) fauny SSSR i sopredelnykh stran [The velvet ants (Hymenoptera, Mutillidae) of the USSR and neighbouring countries]. Nauka, Leningrad. 268 pp., 64 tables. (In Russian)." type="book" year="1985">
Lelej 1985, Fig. 32,
<emphasis id="61B6EA96FFA2711FFD5B8E77FD32EC08" box="[686,700,1124,1149]" italics="true" pageId="7" pageNumber="8">4</emphasis>
</bibRefCitation>
).
</paragraph>
<paragraph id="537D3684FFA2711FFECD8E9FFE27ED1B" blockId="7.[264,1323,604,1430]" pageId="7" pageNumber="8">Head and mesosoma red­testaceous, ocellar spot darkened. Legs castaneous with lighter tarsi. Metasomal segment 1 reddish­brown to piceous, segments 2–7 dark brown. Flagellomeres reddish­brown, lighter beneath, palps pale brown. Wings slightly infuscate in apical half with brown pterostigma and veins. Body and legs clothed with short recumbent and scattered long erect white pubescence. T2–T6 and S2–S6 with weak apical white fringes.</paragraph>
<paragraph id="537D3684FFA2711FFECD8F6FFDB9EDE3" blockId="7.[264,1323,604,1430]" box="[312,567,1404,1430]" pageId="7" pageNumber="8">FEMALE. Unknown.</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFA2711FFEFD8FDFFC78EF3B" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="537D3684FFA2711FFEFD8FDFFE68ED90" blockId="7.[264,1323,1484,1870]" box="[264,486,1484,1509]" pageId="7" pageNumber="8">
<emphasis id="61B6EA96FFA2711FFEFD8FDFFE68ED90" box="[264,486,1484,1509]" italics="true" pageId="7" pageNumber="8">Material examined</emphasis>
</paragraph>
<paragraph id="537D3684FFA2711FFECD8FE7FC78EF3B" blockId="7.[264,1323,1484,1870]" pageId="7" pageNumber="8">
<collectingCountry id="2BD57614FFA2711FFECD8FE7FE00EE7B" box="[312,398,1524,1550]" name="Yemen" pageId="7" pageNumber="8">Yemen</collectingCountry>
: Sanaa,
<date id="277C1044FFA2711FFE0D8FE7FDEBEE7B" box="[504,613,1524,1550]" pageId="7" pageNumber="8" value="1991-07">VII.1991</date>
, LT, AvH, 1 ɗ [
<collectionCode id="35D3AE41FFA2711FFCC38FE7FC18EE7B" LSID="urn:lsid:biocol.org:col:34975" box="[822,918,1524,1550]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="7" pageNumber="8">ZMAN</collectionCode>
]; Ma’bar,
<date id="277C1044FFA2711FFBE28FE7FB48EE7B" box="[1047,1222,1524,1550]" pageId="7" pageNumber="8" value="1992-07" valueMax="1992-08-31" valueMin="1992-07-00">VII–VIII.1992</date>
, LT, M. Mahyoub, 2 ɗ [
<collectionCode id="35D3AE41FFA2711FFE3C8C0FFDA7EE43" LSID="urn:lsid:biocol.org:col:34975" box="[457,553,1564,1590]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="7" pageNumber="8">ZMAN</collectionCode>
]; Qa Al Boun,
<date id="277C1044FFA2711FFD118C0FFC1CEE43" box="[740,914,1564,1590]" pageId="7" pageNumber="8" value="1992-07" valueMax="1992-08-31" valueMin="1992-07-00">VII–VIII.1992</date>
, LT, M. Mahyoub, 6 ɗ [
<collectionCode id="35D3AE41FFA2711FFB4A8C0FFA90EE43" LSID="urn:lsid:biocol.org:col:34975" box="[1215,1310,1564,1590]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="7" pageNumber="8">ZMAN</collectionCode>
]; Ta’izz,
<date id="277C1044FFA2711FFE958C57FE7EEE2B" box="[352,496,1604,1630]" pageId="7" pageNumber="8" value="1999-01-03" valueMax="1999-01-24" valueMin="1999-01-03">3–24.I.1999</date>
, AvH &amp; M. Mahyoub, 1 ɗ [
<collectionCode id="35D3AE41FFA2711FFCCB8C57FC13EE2B" LSID="urn:lsid:biocol.org:col:34975" box="[830,925,1604,1630]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="7" pageNumber="8">ZMAN</collectionCode>
];
<date id="277C1044FFA2711FFC448C57FB84EE2B" box="[945,1034,1604,1630]" pageId="7" pageNumber="8" value="1999-10">X.1999</date>
, LT, AvH &amp; A. Awad, 1 ɗ [
<collectionCode id="35D3AE41FFA2711FFED88C7FFE03EEF3" LSID="urn:lsid:biocol.org:col:34975" box="[301,397,1644,1670]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="7" pageNumber="8">ZMAN</collectionCode>
];
<date id="277C1044FFA2711FFE568C7FFD89EEF3" box="[419,519,1644,1670]" pageId="7" pageNumber="8" value="2000-09">IX.2000</date>
, LT, AvH &amp; A. R. Al Yarimi, 1 ɗ [
<collectionCode id="35D3AE41FFA2711FFC368C7FFBADEEF3" LSID="urn:lsid:biocol.org:col:34975" box="[963,1059,1644,1670]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="7" pageNumber="8">ZMAN</collectionCode>
]; Sanaa,
<date id="277C1044FFA2711FFB648C7FFB7BEEF3" box="[1169,1269,1644,1670]" pageId="7" pageNumber="8" value="1999-06">VI.1999</date>
, LT, AvH, 2 ɗ [
<collectionCode id="35D3AE41FFA2711FFE688C87FE73EEDB" LSID="urn:lsid:biocol.org:col:34975" box="[413,509,1684,1710]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="7" pageNumber="8">ZMAN</collectionCode>
]; Al Lahima,
<date id="277C1044FFA2711FFD418C87FC00EEDB" box="[692,910,1684,1710]" pageId="7" pageNumber="8" value="2000-10-16" valueMax="2000-12-31" valueMin="2000-10-16">16.X–31.XII.2000</date>
, MT, AvH &amp; A.M. Hager, 2 ɗ [
<collectionCode id="35D3AE41FFA2711FFEFA8CAFFEE1EEA3" LSID="urn:lsid:biocol.org:col:34975" box="[271,367,1724,1750]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="7" pageNumber="8">ZMAN</collectionCode>
];
<quantity id="943A9B61FFA2711FFE7B8CAFFE6AEEA3" box="[398,484,1724,1750]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="8" unit="km" value="12.0">12 km</quantity>
NW Manakhah,
<date id="277C1044FFA2711FFD218CAFFC33EEA3" box="[724,957,1724,1750]" pageId="7" pageNumber="8" value="2001-07-03" valueMax="2001-08-21" valueMin="2001-07-03">3.VII–21.VIII.2001</date>
, MT, AvH, 6 ɗ [
<collectionCode id="35D3AE41FFA2711FFB4B8CAFFA91EEA3" LSID="urn:lsid:biocol.org:col:34992" box="[1214,1311,1724,1750]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="7" pageNumber="8">RMNH</collectionCode>
];
<date id="277C1044FFA2711FFEFD8CF7FE68EE8B" box="[264,486,1764,1790]" pageId="7" pageNumber="8" value="2003-06-24" valueMax="2003-08-04" valueMin="2003-06-24">24.VI–4.VIII.2003</date>
, MT, AvH, 1 ɗ [
<collectionCode id="35D3AE41FFA2711FFD458CF7FC9FEE8B" LSID="urn:lsid:biocol.org:col:34992" box="[688,785,1764,1790]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="7" pageNumber="8">RMNH</collectionCode>
];
<date id="277C1044FFA2711FFCD08CF7FC78EE8B" box="[805,1014,1764,1790]" pageId="7" pageNumber="8" value="2003-09-15" valueMax="2003-10-22" valueMin="2003-09-15">15.IX–22.X.2003</date>
, MT, AvH, 1 ɗ [
<collectionCode id="35D3AE41FFA2711FFB4B8CF7FA91EE8B" LSID="urn:lsid:biocol.org:col:34992" box="[1214,1311,1764,1790]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="7" pageNumber="8">RMNH</collectionCode>
]; Jebel Burra, Primärwald (primary forest),
<quantity id="943A9B61FFA2711FFCF28D1FFCDBEF53" box="[775,853,1804,1830]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="8" unit="km" value="25.0">25 km</quantity>
SE Bajil,
<geoCoordinate id="36F65043FFA2711FFC228D1FFBB2EF53" box="[983,1084,1804,1830]" direction="north" orientation="latitude" pageId="7" pageNumber="8" precision="925" value="14.883333">14°53’N</geoCoordinate>
<geoCoordinate id="36F65043FFA2711FFBB28D1FFB22EF53" box="[1095,1196,1804,1830]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="925" value="43.45">43°27’E</geoCoordinate>
, 1000 NN Jem­Bar,
<date id="277C1044FFA2711FFE8C8D27FE73EF3B" box="[377,509,1844,1870]" pageId="7" pageNumber="8" value="1997-04-16">16.IV.1997</date>
, lif Brechtel, Würst, Ehrmann, 1 ɗ [
<collectionCode id="35D3AE41FFA2711FFC508D27FC65EF3B" box="[933,1003,1844,1870]" pageId="7" pageNumber="8">IBSS</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFAD7110FEFD8B0FFCB0E92B" box="[264,830,284,350]" pageId="8" pageNumber="9" type="distribution">
<paragraph id="537D3684FFAD7110FEFD8B0FFE19E940" blockId="8.[264,830,284,350]" box="[264,407,284,309]" pageId="8" pageNumber="9">
<emphasis id="61B6EA96FFAD7110FEFD8B0FFE19E940" box="[264,407,284,309]" italics="true" pageId="8" pageNumber="9">Distribution</emphasis>
</paragraph>
<paragraph id="537D3684FFAD7110FECD8B57FCB0E92B" blockId="8.[264,830,284,350]" box="[312,830,324,350]" pageId="8" pageNumber="9">
<collectingCountry id="2BD57614FFAD7110FECD8B57FE04E92B" box="[312,394,324,350]" name="Yemen" pageId="8" pageNumber="9">Yemen</collectingCountry>
(new record),
<collectingCountry id="2BD57614FFAD7110FDC38B57FD18E92B" box="[566,662,324,350]" name="Somalia" pageId="8" pageNumber="9">Somalia</collectingCountry>
(Migiurtinia).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:identifier id="B22A30C977D5FD59B00D2323D55A2061" type="DOI">10.5281/zenodo.172671</mods:identifier>
<mods:identifier id="C5C8FEDC33CBD9B40AB0CBE7C061A783" type="GBIF-Dataset">b0d82648-af13-4bba-8318-49fa28bd2515</mods:identifier>
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<treatment id="DB6B8792FFA3711EFEFD8925FBFEEE0E" ID-DOI="http://doi.org/10.5281/zenodo.6252931" ID-GBIF-Taxon="119344657" ID-Zenodo-Dep="6252931" LSID="urn:lsid:plazi:treatment:DB6B8792FFA3711EFEFD8925FBFEEE0E" httpUri="http://treatment.plazi.org/id/DB6B8792FFA3711EFEFD8925FBFEEE0E" lastPageNumber="7" pageId="6" pageNumber="7">
<subSubSection id="1BD8650FFFA3711EFEFD8925FD34EB25" box="[264,698,822,848]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="537D3684FFA3711EFEFD8925FD34EB25" blockId="6.[264,698,822,848]" box="[264,698,822,848]" pageId="6" pageNumber="7">
<heading id="083581E8FFA3711EFEFD8925FD34EB25" bold="true" box="[264,698,822,848]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<taxonomicName id="94C24D07FFA3711EFEFD8925FD34EB25" authority="Klug 1829" authorityName="Klug" authorityYear="1829" box="[264,698,822,848]" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="aurea">
<emphasis id="61B6EA96FFA3711EFEFD8925FD34EB25" bold="true" box="[264,698,822,848]" pageId="6" pageNumber="7">
<emphasis id="61B6EA96FFA3711EFEFD8925FD91EB25" bold="true" box="[264,543,822,848]" italics="true" pageId="6" pageNumber="7">Pseudophotopsis aurea</emphasis>
(
<bibRefCitation id="37534B75FFA3711EFDDA8925FD3CEB25" author="Klug" box="[559,690,822,848]" pageId="6" pageNumber="48" refString="Klug, J. C. F. (1829) Symbolae physicae seu icones et descriptiones insectorum quae ex itinere per Africam borealem et Asiam occidentalem Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg studio novae aut illustratae redierunt. Hymenoptera. Decas Prima [genera Mutilla et Apterogyna]. Venditur a Mittlero, Berolini. [15 - 21] + tab. 4, 5." type="journal volume" year="1829">Klug 1829</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFA3711EFEFD896FFE60ECC6" pageId="6" pageNumber="7" type="reference_group">
<paragraph id="537D3684FFA3711EFEFD896FFDE2EBC6" blockId="6.[264,1323,891,1203]" pageId="6" pageNumber="7">
<taxonomicName id="94C24D07FFA3711EFEFD896FFD92EBE6" authority="Klug 1829" authorityName="Klug" authorityYear="1829" box="[264,540,892,915]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="aurea">
<emphasis id="61B6EA96FFA3711EFEFD896FFE12EBE6" box="[264,412,892,915]" italics="true" pageId="6" pageNumber="7">Mutilla aurea</emphasis>
<bibRefCitation id="37534B75FFA3711EFE50896FFD92EBE6" author="Klug" box="[421,540,892,915]" pageId="6" pageNumber="48" refString="Klug, J. C. F. (1829) Symbolae physicae seu icones et descriptiones insectorum quae ex itinere per Africam borealem et Asiam occidentalem Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg studio novae aut illustratae redierunt. Hymenoptera. Decas Prima [genera Mutilla et Apterogyna]. Venditur a Mittlero, Berolini. [15 - 21] + tab. 4, 5." type="journal volume" year="1829">Klug 1829</bibRefCitation>
</taxonomicName>
: [18], tab. 4, fig. 13, Ψ &quot;Ex Habissinia et Arabia deserta&quot; [
<collectingCountry id="2BD57614FFA3711EFB6B896FFB75EBE6" box="[1182,1275,892,915]" name="Ethiopia" pageId="6" pageNumber="7">Ethiopia</collectingCountry>
and Arabia],
<typeStatus id="8C798826FFA3711EFE62898FFE7AEBC6" box="[407,500,924,947]" pageId="6" pageNumber="7" type="syntype">syntypes</typeStatus>
in ZMHU.
</paragraph>
<paragraph id="537D3684FFA3711EFEFD89AFFB2FEBA6" blockId="6.[264,1323,891,1203]" box="[264,1185,955,979]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="73D211AAFFA3711EFEFD89AFFB2FEBA6" box="[264,1185,955,979]" pageId="6" pageNumber="7">
<taxonomicName id="94C24D07FFA3711EFEFD89AFFD05EBA6" authority="Magretti 1898: 42" authorityName="Magretti" authorityPageNumber="42" authorityYear="1898" box="[264,651,956,979]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="kassalina">
<emphasis id="61B6EA96FFA3711EFEFD89AFFE4CEBA6" box="[264,450,956,979]" italics="true" pageId="6" pageNumber="7">Mutilla kassalina</emphasis>
<treatmentCitation id="D2631095FFA3711EFE3C89AFFD05EBA6" author="Magretti" box="[457,651,956,979]" page="42" pageId="6" pageNumber="7" year="1898">
<bibRefCitation id="37534B75FFA3711EFE3C89AFFDEAEBA6" author="Magretti" box="[457,612,956,979]" lastPageId="6" lastPageNumber="49" pageId="6" pageNumber="48" refString="Magretti, P. (1898) Imenotteri della seconda spedizione di Don Eugenio dei principi Ruspoli nei paesi Galla e Somali. Annali del Museo Civico di Storia Naturale di Genova, ser. 2 a, 19, 37 - 42." type="journal article" year="1898">Magretti 1898</bibRefCitation>
: 42
</treatmentCitation>
</taxonomicName>
, ♂,
<typeStatus id="8C798826FFA3711EFD4389AFFC9DEBA6" box="[694,787,956,979]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
&quot;Kassala&quot; [
<collectingCountry id="2BD57614FFA3711EFC6589AFFC54EBA6" box="[912,986,956,979]" name="Sudan" pageId="6" pageNumber="7">Sudan</collectingCountry>
].
<emphasis id="61B6EA96FFA3711EFC1E89A8FB17EBA6" bold="true" box="[1003,1177,955,979]" pageId="6" pageNumber="7">New Synonymy</emphasis>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="537D3684FFA3711EFEFD89CFFA95EB86" blockId="6.[264,1323,891,1203]" box="[264,1307,987,1011]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="73D211AAFFA3711EFEFD89CFFA95EB86" box="[264,1307,987,1011]" pageId="6" pageNumber="7">
<taxonomicName id="94C24D07FFA3711EFEFD89CFFD77EB86" authority="Magretti 1906: 40" authorityName="Magretti" authorityPageNumber="40" authorityYear="1906" box="[264,761,988,1011]" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="kassalina">
<emphasis id="61B6EA96FFA3711EFEFD89CFFDA6EB86" box="[264,552,988,1011]" italics="true" pageId="6" pageNumber="7">Pseudophotopsis kassalina</emphasis>
: Magretti 1906: 40
</taxonomicName>
, ♂; Bischoff 1920: 99, ♂;
<treatmentCitation id="D2631095FFA3711EFBE489CFFB79EB86" author="Nonveiller" box="[1041,1271,988,1011]" page="105" pageId="6" pageNumber="7" year="1974">
<bibRefCitation id="37534B75FFA3711EFBE489CFFB4CEB86" author="Nonveiller" box="[1041,1218,988,1011]" pageId="6" pageNumber="49" refString="Nonveiller, G. (1974) Recherches sur les Mutillides de l'Afrique (Hymenoptera, Mutillidae). V. Description du male encore inconnu de Pseudophotopsis continua F. seul representant du genre en Afrique occidentale. Annales de la Faculte des Sciences du Cameroun, 18, 95 - 108." type="journal article" year="1974">Nonveiller 1974</bibRefCitation>
: 105
</treatmentCitation>
, ♂.
</treatmentCitationGroup>
</paragraph>
<paragraph id="537D3684FFA3711EFEFD89EFFD88EC46" blockId="6.[264,1323,891,1203]" pageId="6" pageNumber="7">
<taxonomicName id="94C24D07FFA3711EFEFD89EFFE61EC66" box="[264,495,1020,1043]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="continua">
<emphasis id="61B6EA96FFA3711EFEFD89EFFE61EC66" box="[264,495,1020,1043]" italics="true" pageId="6" pageNumber="7">Ephutomma continua</emphasis>
</taxonomicName>
<taxonomicName id="94C24D07FFA3711EFE0C89EFFCCEEC66" authority="Magretti 1906: 37" authorityName="Magretti" authorityPageNumber="37" authorityYear="1906" box="[505,832,1020,1043]" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="variety" species="continua" variety="aurea">
var.
<emphasis id="61B6EA96FFA3711EFDDF89EFFDE9EC66" box="[554,615,1020,1043]" italics="true" pageId="6" pageNumber="7">aurea</emphasis>
: Magretti 1906: 37
</taxonomicName>
, Ψ (
<collectingCountry id="2BD57614FFA3711EFC8489EFFC4EEC66" box="[881,960,1020,1043]" name="Eritrea" pageId="6" pageNumber="7">Eritrea</collectingCountry>
); Bischoff 1920: 151, ♂ (
<collectingCountry id="2BD57614FFA3711EFB2F89EFFAA9EC66" box="[1242,1319,1020,1043]" name="Eritrea" pageId="6" pageNumber="7">Eritrea</collectingCountry>
,
<collectingCountry id="2BD57614FFA3711EFECD8E0FFE1AEC46" box="[312,404,1052,1075]" name="Djibouti" pageId="6" pageNumber="7">Djibouti</collectingCountry>
,
<collectingCountry id="2BD57614FFA3711EFE6B8E0FFE74EC46" box="[414,506,1052,1075]" name="Somalia" pageId="6" pageNumber="7">Somalia</collectingCountry>
).
</paragraph>
<paragraph id="537D3684FFA3711EFEFD8E2FFC05EC26" blockId="6.[264,1323,891,1203]" box="[264,907,1083,1107]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="73D211AAFFA3711EFEFD8E2FFC05EC26" box="[264,907,1083,1107]" pageId="6" pageNumber="7">
<taxonomicName id="94C24D07FFA3711EFEFD8E2FFD79EC26" authority="Invrea 1956: 301" authorityName="Invrea" authorityPageNumber="301" authorityYear="1956" box="[264,759,1084,1107]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subSpecies" species="continua" subSpecies="aurea">
<emphasis id="61B6EA96FFA3711EFEFD8E2FFDBEEC26" box="[264,560,1084,1107]" italics="true" pageId="6" pageNumber="7">Ephutomma continua aurea</emphasis>
:
<treatmentCitation id="D2631095FFA3711EFDCA8E2FFD79EC26" author="Invrea" box="[575,759,1084,1107]" page="301" pageId="6" pageNumber="7" year="1956">
<bibRefCitation id="37534B75FFA3711EFDCA8E2FFD4DEC26" author="Invrea" box="[575,707,1084,1107]" pageId="6" pageNumber="48" refString="Invrea, F. (1956) Missione del Prof. Guiseppe Scortecci in Migiurtinia con il contributo del consiglio nazionale delle ricerche. II. Apterogynidae e Mutillidae (Hymenoptera). Annali del Museo Civico di Storia Naturale Giacomo Doria, 68, 297 - 305." type="journal article" year="1956">Invrea 1956</bibRefCitation>
: 301
</treatmentCitation>
</taxonomicName>
, Ψ (
<collectingCountry id="2BD57614FFA3711EFCD78E2FFCF1EC26" box="[802,895,1084,1107]" name="Somalia" pageId="6" pageNumber="7">Somalia</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="537D3684FFA3711EFEFD8E4FFCF2EC06" blockId="6.[264,1323,891,1203]" box="[264,892,1115,1139]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="73D211AAFFA3711EFEFD8E4FFCF2EC06" box="[264,892,1115,1139]" pageId="6" pageNumber="7">
<taxonomicName id="94C24D07FFA3711EFEFD8E4FFD7BEC06" authority="Nonveiller 1974: 105" authorityName="Nonveiller" authorityPageNumber="105" authorityYear="1974" box="[264,757,1116,1139]" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="aurea">
<emphasis id="61B6EA96FFA3711EFEFD8E4FFD8EEC06" box="[264,512,1116,1139]" italics="true" pageId="6" pageNumber="7">Pseudophotopsis aurea</emphasis>
:
<treatmentCitation id="D2631095FFA3711EFDFA8E4FFD7BEC06" author="Nonveiller" box="[527,757,1116,1139]" page="105" pageId="6" pageNumber="7" year="1974">
<bibRefCitation id="37534B75FFA3711EFDFA8E4FFD4FEC06" author="Nonveiller" box="[527,705,1116,1139]" pageId="6" pageNumber="49" refString="Nonveiller, G. (1974) Recherches sur les Mutillides de l'Afrique (Hymenoptera, Mutillidae). V. Description du male encore inconnu de Pseudophotopsis continua F. seul representant du genre en Afrique occidentale. Annales de la Faculte des Sciences du Cameroun, 18, 95 - 108." type="journal article" year="1974">Nonveiller 1974</bibRefCitation>
: 105
</treatmentCitation>
</taxonomicName>
, Ψ (
<collectingCountry id="2BD57614FFA3711EFCD58E4FFCFFEC06" box="[800,881,1116,1139]" name="Yemen" pageId="6" pageNumber="7">Yemen</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="537D3684FFA3711EFEFD8E6FFE60ECC6" blockId="6.[264,1323,891,1203]" pageId="6" pageNumber="7">
<taxonomicName id="94C24D07FFA3711EFEFD8E6FFCB3ECE6" authority="Hammer 1962: 2" authorityName="Hammer" authorityPageNumber="2" authorityYear="1962" box="[264,829,1148,1171]" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subSpecies" species="continua" subSpecies="arabica">
<emphasis id="61B6EA96FFA3711EFEFD8E6FFDF2ECE6" box="[264,636,1148,1171]" italics="true" pageId="6" pageNumber="7">Pseudophotopsis continua arabica</emphasis>
Hammer 1962: 2
</taxonomicName>
, Ψ,
<typeStatus id="8C798826FFA3711EFC9C8E6FFC48ECE6" box="[873,966,1148,1171]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
&quot;Sufean, Lahej, Aden&quot; [
<collectingCountry id="2BD57614FFA3711EFB3B8E6FFAAFECE6" box="[1230,1313,1148,1171]" name="Yemen" pageId="6" pageNumber="7">Yemen</collectingCountry>
].
<emphasis id="61B6EA96FFA3711EFECD8E88FE66ECC6" bold="true" box="[312,488,1179,1203]" pageId="6" pageNumber="7">New Synonymy</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFA3711EFEFD8EF7FEF5ED03" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="537D3684FFA3711EFEFD8EF7FE68EC88" blockId="6.[264,1323,1252,1398]" box="[264,486,1252,1277]" pageId="6" pageNumber="7">
<emphasis id="61B6EA96FFA3711EFEFD8EF7FE68EC88" box="[264,486,1252,1277]" italics="true" pageId="6" pageNumber="7">Material examined</emphasis>
</paragraph>
<paragraph id="537D3684FFA3711EFECD8F1FFEF5ED03" blockId="6.[264,1323,1252,1398]" pageId="6" pageNumber="7">
<collectingCountry id="2BD57614FFA3711EFECD8F1FFE01ED53" box="[312,399,1292,1318]" name="Yemen" pageId="6" pageNumber="7">Yemen</collectingCountry>
: Sanaa,
<date id="277C1044FFA3711EFE1A8F1FFDFDED53" box="[495,627,1292,1318]" pageId="6" pageNumber="7" value="1998-02" valueMax="1998-03-31" valueMin="1998-02-00">II–III.1998</date>
, MT, AvH, 1 Ψ [
<collectionCode id="35D3AE41FFA3711EFCC38F1FFC1BED53" LSID="urn:lsid:biocol.org:col:34975" box="[822,917,1292,1318]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="6" pageNumber="7">ZMAN</collectionCode>
]; Ta’izz,
<date id="277C1044FFA3711EFBF48F1FFB37ED53" box="[1025,1209,1292,1318]" pageId="6" pageNumber="7" value="1998-04-26" valueMax="1998-04-28" valueMin="1998-04-26">26–28.IV. 1998</date>
, LT, AvH &amp; A. Awad, 1 ɗ [
<collectionCode id="35D3AE41FFA3711EFE1C8F27FDC7ED3B" LSID="urn:lsid:biocol.org:col:34975" box="[489,585,1332,1358]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="6" pageNumber="7">ZMAN</collectionCode>
];
<quantity id="943A9B61FFA3711EFD958F27FD21ED3B" box="[608,687,1332,1358]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="7" unit="km" value="12.0">12 km</quantity>
NW Manakhah,
<date id="277C1044FFA3711EFC738F27FBEBED3B" box="[902,1125,1332,1358]" pageId="6" pageNumber="7" value="2003-06-24" valueMax="2003-08-04" valueMin="2003-06-24">24.VI–4.VIII.2003</date>
, MT, AvH, 1 Ψ [
<collectionCode id="35D3AE41FFA3711EFEFA8F4FFEFEED03" LSID="urn:lsid:biocol.org:col:34992" box="[271,368,1372,1398]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="7">RMNH</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFA3711EFEFD8F8CFC27ED94" box="[264,937,1439,1505]" pageId="6" pageNumber="7" type="distribution">
<paragraph id="537D3684FFA3711EFEFD8F8CFE19EDCD" blockId="6.[264,937,1439,1505]" box="[264,407,1439,1464]" pageId="6" pageNumber="7">
<emphasis id="61B6EA96FFA3711EFEFD8F8CFE19EDCD" box="[264,407,1439,1464]" italics="true" pageId="6" pageNumber="7">Distribution</emphasis>
</paragraph>
<paragraph id="537D3684FFA3711EFECD8FD4FC27ED94" blockId="6.[264,937,1439,1505]" box="[312,937,1479,1505]" pageId="6" pageNumber="7">
<collectingCountry id="2BD57614FFA3711EFECD8FD4FE03ED94" box="[312,397,1479,1505]" name="Yemen" pageId="6" pageNumber="7">Yemen</collectingCountry>
,
<collectingCountry id="2BD57614FFA3711EFE6D8FD4FE65ED94" box="[408,491,1479,1505]" name="Eritrea" pageId="6" pageNumber="7">Eritrea</collectingCountry>
,
<collectingCountry id="2BD57614FFA3711EFE028FD4FDD2ED94" box="[503,604,1479,1505]" name="Djibouti" pageId="6" pageNumber="7">Djibouti</collectingCountry>
,
<collectingCountry id="2BD57614FFA3711EFD928FD4FD49ED94" box="[615,711,1479,1505]" name="Somalia" pageId="6" pageNumber="7">Somalia</collectingCountry>
(
<bibRefCitation id="37534B75FFA3711EFD238FD4FC12ED94" author="Nonveiller" box="[726,924,1479,1505]" pageId="6" pageNumber="49" refString="Nonveiller, G. (1974) Recherches sur les Mutillides de l'Afrique (Hymenoptera, Mutillidae). V. Description du male encore inconnu de Pseudophotopsis continua F. seul representant du genre en Afrique occidentale. Annales de la Faculte des Sciences du Cameroun, 18, 95 - 108." type="journal article" year="1974">Nonveiller 1974</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFA3711EFEFD8C02FBFEEE0E" pageId="6" pageNumber="7" type="discussion">
<paragraph id="537D3684FFA3711EFEFD8C02FEE0EE5F" blockId="6.[264,1322,1553,1659]" box="[264,366,1553,1578]" pageId="6" pageNumber="7">
<emphasis id="61B6EA96FFA3711EFEFD8C02FEE0EE5F" box="[264,366,1553,1578]" italics="true" pageId="6" pageNumber="7">Remarks</emphasis>
</paragraph>
<paragraph id="537D3684FFA3711EFECD8C2AFBFEEE0E" blockId="6.[264,1322,1553,1659]" pageId="6" pageNumber="7">
This is a very distinct species. Golden apical fringes on T2–T
<quantity id="943A9B61FFA3711EFBD48C2AFBC0EE26" box="[1057,1102,1593,1619]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="6" pageNumber="7" unit="in" value="5.0">5 in</quantity>
female (T2–T
<quantity id="943A9B61FFA3711EFB088C2AFAA4EE26" box="[1277,1322,1593,1619]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="6" pageNumber="7" unit="in" value="6.0">6 in</quantity>
male) and hypostomal carina shape are common characters for both sexes.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="3E6CB74F0C90ACAC656B8D7694CDE350" ID-DOI="10.5281/zenodo.172671" ID-GBIF-Dataset="b0d82648-af13-4bba-8318-49fa28bd2515" ID-ISSN="1175­5326" ID-Zenodo-Dep="172671" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459760090076" checkinUser="plazi" docAuthor="Lelej, Arkady S. &amp; Harten, Antonius Van" docDate="2006" docId="DB6B8792FFA87116FEFD898FFC1FEC5C" docLanguage="en" docName="zt01226p050.pdf" docOrigin="Zootaxa 1226" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Mutilla conjuncta Klug 1829, comb. nov." docType="treatment" docVersion="8" lastPageNumber="15" masterDocId="2752FFEAFFA57118FFF58A13FF8EE875" masterDocTitle="A review of the Mutillidae (Hymenoptera) of Yemen" masterLastPageNumber="50" masterPageNumber="1" pageNumber="14" updateTime="1698214875622" updateUser="plazi">
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<mods:title id="04E7201A3E3EE3B7129326096C0C7B73">A review of the Mutillidae (Hymenoptera) of Yemen</mods:title>
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<subSubSection id="1BD8650FFFA87115FEFD898FFE1EEBAC" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="537D3684FFA87115FEFD898FFCAFEBC3" blockId="13.[264,801,924,985]" box="[264,801,924,950]" pageId="13" pageNumber="14">
<heading id="083581E8FFA87115FEFD898FFCAFEBC3" bold="true" box="[264,801,924,950]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="61B6EA96FFA87115FEFD898FFCAFEBC3" bold="true" box="[264,801,924,950]" pageId="13" pageNumber="14">
<emphasis id="61B6EA96FFA87115FEFD898FFEFCEBC3" bold="true" box="[264,370,924,950]" italics="true" pageId="13" pageNumber="14">Omotilla</emphasis>
<taxonomicName id="94C24D07FFA87115FE8C898FFD03EBC3" authority="(Klug 1829)" authorityName="Klug" authorityYear="1829" baseAuthorityName="Klug" baseAuthorityYear="1829" box="[377,653,924,950]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="conjuncta" status="comb. nov.">
<emphasis id="61B6EA96FFA87115FE8C898FFE7FEBC3" bold="true" box="[377,497,924,950]" italics="true" pageId="13" pageNumber="14">conjuncta</emphasis>
(
<bibRefCitation id="37534B75FFA87115FDF4898FFD0BEBC3" author="Klug" box="[513,645,924,950]" pageId="13" pageNumber="48" refString="Klug, J. C. F. (1829) Symbolae physicae seu icones et descriptiones insectorum quae ex itinere per Africam borealem et Asiam occidentalem Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg studio novae aut illustratae redierunt. Hymenoptera. Decas Prima [genera Mutilla et Apterogyna]. Venditur a Mittlero, Berolini. [15 - 21] + tab. 4, 5." type="journal volume" year="1829">Klug 1829</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="7A8557EDFFA87115FD6F898FFCAFEBC3" box="[666,801,924,950]" pageId="13" pageNumber="14">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="537D3684FFA87115FEFD89ACFE1EEBAC" blockId="13.[264,801,924,985]" box="[264,400,959,985]" pageId="13" pageNumber="14">
(
<figureCitation id="CBF92A01FFA87115FEE589ACFE09EBAC" box="[272,391,959,985]" captionStart="FIGURES 2 – 10. 2 – 5" captionStartId="8.[264,382,1060,1084]" captionTargetBox="[541,1044,411,1033]" captionTargetId="figure@8.[535,1051,406,1036]" captionTargetPageId="8" captionText="FIGURES 2 – 10. 2 – 5. Pseudophotopsis mascatiana, male. 2. Forewing; 3. Head; 4. Metasomal tergum 1, dorsal view; 5. Mandible, lateral view. 6 – 8. Arnoldtilla achterbergi sp. nov., female, holotype. 6. Habitus, dorsal view; 7. Mandible, dorsal view; 8. Pedicel and flagellomeres 1 – 3. 9 – 10. Omotilla conjuncta, male. 9. Mandible, dorsal view; 10. Genitalia, ventral view. Scale = 1 mm for 2 – 4, 6 and 0.5 mm for 5, 7 – 10." httpUri="https://zenodo.org/record/172673/files/figure.png" pageId="13" pageNumber="14">Figs 9, 10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFA87115FEFD8E1CFB0AECF0" pageId="13" pageNumber="14" type="reference_group">
<paragraph id="537D3684FFA87115FEFD8E1CFD69EC10" blockId="13.[264,1323,1038,1158]" pageId="13" pageNumber="14">
<taxonomicName id="94C24D07FFA87115FEFD8E1CFDC0EC50" authority="Klug 1829" authorityName="Klug" authorityYear="1829" box="[264,590,1038,1062]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="conjuncta">
<emphasis id="61B6EA96FFA87115FEFD8E1CFE44EC53" box="[264,458,1039,1062]" italics="true" pageId="13" pageNumber="14">Mutilla conjuncta</emphasis>
<bibRefCitation id="37534B75FFA87115FE208E1DFDC0EC50" author="Klug" box="[469,590,1038,1061]" pageId="13" pageNumber="48" refString="Klug, J. C. F. (1829) Symbolae physicae seu icones et descriptiones insectorum quae ex itinere per Africam borealem et Asiam occidentalem Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg studio novae aut illustratae redierunt. Hymenoptera. Decas Prima [genera Mutilla et Apterogyna]. Venditur a Mittlero, Berolini. [15 - 21] + tab. 4, 5." type="journal volume" year="1829">Klug 1829</bibRefCitation>
</taxonomicName>
: [16], tab. 5, fig. 5, Ψ, &quot;Arabia
<taxonomicName id="94C24D07FFA87115FC4D8E1DFC67EC50" box="[952,1001,1038,1061]" class="Insecta" family="Mutillidae" genus="Vanhartenidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="felix">felix</taxonomicName>
et Ambukohl&quot;,
<typeStatus id="8C798826FFA87115FB6A8E1DFA8DEC50" box="[1183,1283,1038,1061]" pageId="13" pageNumber="14" type="lectotype">lectotype</typeStatus>
, Ψ, &quot;Ambukohl, Ehrenberg&quot; (
<collectingCountry id="2BD57614FFA87115FDBF8E3DFD1FEC30" box="[586,657,1070,1093]" name="Sudan" pageId="13" pageNumber="14">Sudan</collectingCountry>
), designated Bischoff 1920: 73 [ZMHU].
<taxonomicName id="94C24D07FFA87115FEFD8E5CFD4DEC10" authority="Bischoff 1920: 73" authorityName="Bischoff" authorityPageNumber="73" authorityYear="1920" box="[264,707,1102,1126]" class="Insecta" family="Mutillidae" genus="Odontotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="conjuncta">
<emphasis id="61B6EA96FFA87115FEFD8E5CFE7DEC13" box="[264,499,1103,1126]" italics="true" pageId="13" pageNumber="14">Odontotilla conjuncta</emphasis>
: Bischoff 1920: 73
</taxonomicName>
, Ψ.
</paragraph>
<paragraph id="537D3684FFA87115FEFD8E7CFB0AECF0" blockId="13.[264,1323,1038,1158]" box="[264,1156,1134,1158]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="73D211AAFFA87115FEFD8E7CFB0AECF0" box="[264,1156,1134,1158]" pageId="13" pageNumber="14">
<taxonomicName id="94C24D07FFA87115FEFD8E7CFC2BECF0" authority="Nonveiller 1996a: 331" authorityName="Nonveiller" authorityPageNumber="331" authorityYear="1996" box="[264,933,1134,1158]" class="Insecta" family="Mutillidae" genus="Odontotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="conjuncta" subGenus="Conjunctotilla">
<emphasis id="61B6EA96FFA87115FEFD8E7CFE0DECF3" box="[264,387,1135,1158]" italics="true" pageId="13" pageNumber="14">Odontotilla</emphasis>
(
<emphasis id="61B6EA96FFA87115FE668E7CFDA3ECF3" box="[403,557,1135,1158]" italics="true" pageId="13" pageNumber="14">Conjunctotilla</emphasis>
)
<emphasis id="61B6EA96FFA87115FDC98E7CFD2BECF3" box="[572,677,1135,1158]" italics="true" pageId="13" pageNumber="14">conjuncta</emphasis>
:
<treatmentCitation id="D2631095FFA87115FD468E7DFC2BECF0" author="Nonveiller" box="[691,933,1134,1157]" page="331" pageId="13" pageNumber="14" year="1996">
<bibRefCitation id="37534B75FFA87115FD468E7DFCFFECF0" author="Nonveiller" box="[691,881,1134,1157]" pageId="13" pageNumber="49" refString="Nonveiller, G. (1996 a) Revision du genre Afrotropical Odontotilla Bischoff, 1920 (male et femelle) (Hymenoptera: Mutillidae). Annales de la Societe Entomologique de France, (N. S.), 32 (3), 299 - 357." type="journal article" year="1996" yearSuffix="a">Nonveiller 1996a</bibRefCitation>
: 331
</treatmentCitation>
</taxonomicName>
, Ψ (Aden, Hodeida).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFA87115FEFD8EA4FB61EDEC" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="537D3684FFA87115FEFD8EA4FEF1ECA5" blockId="13.[264,1323,1207,1433]" box="[264,383,1207,1232]" pageId="13" pageNumber="14">
<emphasis id="61B6EA96FFA87115FEFD8EA4FEF1ECA5" box="[264,383,1207,1232]" italics="true" pageId="13" pageNumber="14">Diagnosis</emphasis>
</paragraph>
<paragraph id="537D3684FFA87115FECD8ECCFB61EDEC" blockId="13.[264,1323,1207,1433]" pageId="13" pageNumber="14">
The male of
<emphasis id="61B6EA96FFA87115FE3E8ECCFD23EC8D" box="[459,685,1247,1272]" italics="true" pageId="13" pageNumber="14">
Omotilla
<taxonomicName id="94C24D07FFA87115FDCF8ECCFD23EC8D" box="[570,685,1247,1272]" class="Insecta" family="Mutillidae" genus="Odontotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="conjuncta">conjuncta</taxonomicName>
</emphasis>
is similar to that of the
<typeStatus id="8C798826FFA87115FC348ECCFC7DEC8C" box="[961,1011,1247,1273]" pageId="13" pageNumber="14">type</typeStatus>
species
<taxonomicName id="94C24D07FFA87115FBA08ECCFA83EC8D" box="[1109,1293,1247,1272]" class="Insecta" family="Mutillidae" genus="Odontotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="curvidentata">
<emphasis id="61B6EA96FFA87115FBA08ECCFA83EC8D" box="[1109,1293,1247,1272]" italics="true" pageId="13" pageNumber="14">
<collectingCountry id="2BD57614FFA87115FBA08ECCFBE3EC8D" box="[1109,1133,1247,1272]" name="Iceland" pageId="13" pageNumber="14">O</collectingCountry>
. curvidentata
</emphasis>
</taxonomicName>
in having a dentate propodeal lateral border, similar wing venation, a similarly shaped tooth on S2, a similar carina on hypopygium, and similar apical white fringes on metasoma. It can be distinguished from the latter species in having a red mesosoma (black in
<taxonomicName id="94C24D07FFA87115FAFB8F44FE12EDED" class="Insecta" family="Mutillidae" genus="Odontotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="curvidentata">
<emphasis id="61B6EA96FFA87115FAFB8F44FE12EDED" italics="true" pageId="13" pageNumber="14">
<collectingCountry id="2BD57614FFA87115FAFB8F44FAA8ED05" box="[1294,1318,1367,1392]" name="Iceland" pageId="13" pageNumber="14">O</collectingCountry>
. curvidentata
</emphasis>
</taxonomicName>
) and black metasoma (metasomal segments 1–3 red in
<taxonomicName id="94C24D07FFA87115FBD38F6CFB50EDED" box="[1062,1246,1407,1432]" class="Insecta" family="Mutillidae" genus="Odontotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="curvidentata">
<emphasis id="61B6EA96FFA87115FBD38F6CFB50EDED" box="[1062,1246,1407,1432]" italics="true" pageId="13" pageNumber="14">
<collectingCountry id="2BD57614FFA87115FBD38F6CFBB0EDED" box="[1062,1086,1407,1432]" name="Iceland" pageId="13" pageNumber="14">O</collectingCountry>
. curvidentata
</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFA87116FEFD8FDCFD30E92B" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="description">
<paragraph id="537D3684FFA87115FEFD8FDCFE1CED9D" blockId="13.[264,1326,1487,1873]" box="[264,402,1487,1512]" pageId="13" pageNumber="14">
<emphasis id="61B6EA96FFA87115FEFD8FDCFE1CED9D" box="[264,402,1487,1512]" italics="true" pageId="13" pageNumber="14">Description</emphasis>
</paragraph>
<paragraph id="537D3684FFA87115FECD8FE4FCA0EF74" blockId="13.[264,1326,1487,1873]" pageId="13" pageNumber="14">
MALE (hitherto unknown). Body length
<quantity id="943A9B61FFA87115FCEE8FE4FC22EE64" box="[795,940,1527,1553]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" metricValueMax="8.8" metricValueMin="6.4" pageId="13" pageNumber="14" unit="mm" value="7.6" valueMax="8.8" valueMin="6.4">6.4–8.8 mm</quantity>
. Head width 1.1–1.2X its height. Ocelli small, POD: OOD 0.5–0.6X. Eyes slightly emarginated inside. Relation of pedicel and first three flagellomeres 0.5(0.6): 0.9(1.0): 1.4(1.6): 1.3(1.5). Clypeus with two weak tubercles on anterior border. Mandible widened, not excised beneath (
<figureCitation id="CBF92A01FFA87115FBCE8C7CFB0DEEFC" box="[1083,1155,1647,1673]" captionStart="FIGURES 2 – 10. 2 – 5" captionStartId="8.[264,382,1060,1084]" captionTargetBox="[541,1044,411,1033]" captionTargetId="figure@8.[535,1051,406,1036]" captionTargetPageId="8" captionText="FIGURES 2 – 10. 2 – 5. Pseudophotopsis mascatiana, male. 2. Forewing; 3. Head; 4. Metasomal tergum 1, dorsal view; 5. Mandible, lateral view. 6 – 8. Arnoldtilla achterbergi sp. nov., female, holotype. 6. Habitus, dorsal view; 7. Mandible, dorsal view; 8. Pedicel and flagellomeres 1 – 3. 9 – 10. Omotilla conjuncta, male. 9. Mandible, dorsal view; 10. Genitalia, ventral view. Scale = 1 mm for 2 – 4, 6 and 0.5 mm for 5, 7 – 10." httpUri="https://zenodo.org/record/172673/files/figure.png" pageId="13" pageNumber="14">Fig. 9</figureCitation>
). Notauli well developed. Propodeum abrupt, lateral border dentate. S2 with strong dent oriented backward, hypopygium with median triangular carina. T2 with long lateral felt line; S2 without lateral felt line. Genitalia as in
<figureCitation id="CBF92A01FFA87115FD218CF4FCA4EF74" box="[724,810,1767,1793]" captionStart="FIGURES 2 – 10. 2 – 5" captionStartId="8.[264,382,1060,1084]" captionTargetBox="[541,1044,411,1033]" captionTargetId="figure@8.[535,1051,406,1036]" captionTargetPageId="8" captionText="FIGURES 2 – 10. 2 – 5. Pseudophotopsis mascatiana, male. 2. Forewing; 3. Head; 4. Metasomal tergum 1, dorsal view; 5. Mandible, lateral view. 6 – 8. Arnoldtilla achterbergi sp. nov., female, holotype. 6. Habitus, dorsal view; 7. Mandible, dorsal view; 8. Pedicel and flagellomeres 1 – 3. 9 – 10. Omotilla conjuncta, male. 9. Mandible, dorsal view; 10. Genitalia, ventral view. Scale = 1 mm for 2 – 4, 6 and 0.5 mm for 5, 7 – 10." httpUri="https://zenodo.org/record/172673/files/figure.png" pageId="13" pageNumber="14">Fig. 10</figureCitation>
.
</paragraph>
<paragraph id="537D3684FFA87116FECD8D1CFD30E92B" blockId="13.[264,1326,1487,1873]" lastBlockId="14.[264,1321,284,350]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">Head black to red with black frons. Antenna castaneous beneath, mandible red with dark denticles. Mesosoma red (pronotum anteriorly may be darkened), legs dark brown, wings infuscate. Metasoma black, T1 basally and S1 red. T1–T4 posteriorly with bands of dense white setae narrowed medially.</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFAB7116FEFD8B87FD8CEAEB" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="537D3684FFAB7116FEFD8B87FE68E9D8" blockId="14.[264,1323,404,670]" box="[264,486,404,429]" pageId="14" pageNumber="15">
<emphasis id="61B6EA96FFAB7116FEFD8B87FE68E9D8" box="[264,486,404,429]" italics="true" pageId="14" pageNumber="15">Material examined</emphasis>
</paragraph>
<paragraph id="537D3684FFAB7116FECD8BAFFD8CEAEB" blockId="14.[264,1323,404,670]" pageId="14" pageNumber="15">
<collectingCountry id="2BD57614FFAB7116FECD8BAFFE00E9A3" box="[312,398,444,470]" name="Yemen" pageId="14" pageNumber="15">Yemen</collectingCountry>
: Al Kadan,
<date id="277C1044FFAB7116FDD18BAFFD28E9A3" box="[548,678,444,470]" pageId="14" pageNumber="15" value="1997-12-03">3.XII.1997</date>
–
<date id="277C1044FFAB7116FD428BAFFCBEE9A3" box="[695,816,444,470]" pageId="14" pageNumber="15" value="1998-02-17">17.II.1998</date>
, MT, 1 Ψ, AvH &amp; H.M. Naser [
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];
<date id="277C1044FFAB7116FEFD8BF7FE5BE98B" box="[264,469,484,510]" pageId="14" pageNumber="15" value="1998-02-17" valueMax="1998-03-31" valueMin="1998-02-17">17.II–31.III.1998</date>
, MT, 1 ɗ, AvH &amp; H.M. Naser [
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];
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NW Manakhah,
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(
<bibRefCitation id="37534B75FFAB7116FD838937FCC7EB4B" author="Nonveiller" box="[630,841,804,830]" pageId="14" pageNumber="49" refString="Nonveiller, G. (1996 a) Revision du genre Afrotropical Odontotilla Bischoff, 1920 (male et femelle) (Hymenoptera: Mutillidae). Annales de la Societe Entomologique de France, (N. S.), 32 (3), 299 - 357." type="journal article" year="1996" yearSuffix="a">Nonveiller 1996a</bibRefCitation>
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The described male is associated with the female of
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Omotilla
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</emphasis>
because both sexes have similarly shaped mandibles, dentate propodeal lateral border, armed hypopygium of male (this character corresponds to armed S6 of female). Furthermore only one species of this small group is distributed in
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<emphasis id="61B6EA96FFAA7117FEFD895DFE9AEB1D" bold="true" box="[264,276,846,872]" italics="true" pageId="15" pageNumber="16">*</emphasis>
<taxonomicName id="94C24D07FFAA7117FEE1895DFDF7EB1D" authority="(Andre 1901)" authorityName="Andre" authorityYear="1901" baseAuthorityName="Andre" baseAuthorityYear="1901" box="[276,633,846,872]" class="Insecta" family="Mutillidae" genus="Ronisia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="idonea">
<emphasis id="61B6EA96FFAA7117FEE1895DFE43EB1D" bold="true" box="[276,461,846,872]" italics="true" pageId="15" pageNumber="16">Ronisia idonea</emphasis>
(
<bibRefCitation id="37534B75FFAA7117FE2E895DFDFEEB1D" author="Andre" box="[475,624,846,872]" pageId="15" pageNumber="47" refString="Andre, E. (1901) Materiaux por servir a la connaissance des Mutillides d'Afrique. Zeitschrift fur systematische Hymenopterologie und Dipterologie, 1, 279 - 288, 305 - 352." type="journal article" year="1901">André 1901</bibRefCitation>
)
</taxonomicName>
</emphasis>
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<emphasis id="61B6EA96FFAA7117FEFD8989FE28EBC4" box="[264,422,922,945]" italics="true" pageId="15" pageNumber="16">Mutilla idonea</emphasis>
<treatmentCitation id="D2631095FFAA7117FE59898AFDEDEBC5" author="Andre" box="[428,611,921,944]" page="341" pageId="15" pageNumber="16" year="1901">
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: 341
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, Ψ,
<typeStatus id="8C798826FFAA7117FD7B898AFD32EBC5" box="[654,700,921,944]" pageId="15" pageNumber="16">type</typeStatus>
locality &quot;
<collectingCountry id="2BD57614FFAA7117FCD4898AFCF3EBC5" box="[801,893,921,944]" name="Senegal" pageId="15" pageNumber="16">Senegal</collectingCountry>
&quot;.
<taxonomicName id="94C24D07FFAA7117FEFD89A9FD74EBA5" authority="Bischoff 1920: 201" authorityName="Bischoff" authorityPageNumber="201" authorityYear="1920" box="[264,762,953,977]" class="Insecta" family="Mutillidae" genus="Pycnotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="idoneaeformis">
<emphasis id="61B6EA96FFAA7117FEFD89A9FD97EBA4" box="[264,537,954,977]" italics="true" pageId="15" pageNumber="16">Pycnotilla idoneaeformis</emphasis>
Bischoff 1920: 201
</taxonomicName>
, Ψ,
<typeStatus id="8C798826FFAA7117FCD989AAFC02EBA5" box="[812,908,953,976]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
,
<collectingCountry id="2BD57614FFAA7117FC6F89AAFB9DEBA5" box="[922,1043,953,976]" name="Mauritania" pageId="15" pageNumber="16">Mauritania</collectingCountry>
. Synonymized by
<treatmentCitation id="D2631095FFAA7117FB1789AAFE28EB85" author="Suarez" page="925" pageId="15" pageNumber="16" year="1963">
<bibRefCitation id="37534B75FFAA7117FB1789AAFEFFEB85" author="Suarez" pageId="15" pageNumber="50" refString="Suarez, F. J. (1963) Hymenopteres Tiphiides, Apterogynides et mutillides recoltes par J. Mateu dans l'Ennedi et au Tchad. Bulletin de l'Institut fondamental d'Afrique Noire, Dakar. Serie A: Sciences Naturelles, 25 (3), 912 - 943, 10 figs." type="journal article" year="1963">Suárez 1963</bibRefCitation>
: 925
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="537D3684FFAA7117FEFD89E9FD2CEC65" blockId="15.[264,1323,921,1041]" box="[264,674,1017,1041]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="73D211AAFFAA7117FEFD89E9FD2CEC65" box="[264,674,1017,1041]" pageId="15" pageNumber="16">
<taxonomicName id="94C24D07FFAA7117FEFD89E9FD10EC65" authority="Nonveiller 1996b: 94" authorityName="Nonveiller" authorityPageNumber="94" authorityYear="1996" box="[264,670,1017,1041]" class="Insecta" family="Mutillidae" genus="Ronisia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="idonea">
<emphasis id="61B6EA96FFAA7117FEFD89E9FE27EC64" box="[264,425,1018,1041]" italics="true" pageId="15" pageNumber="16">Ronisia idonea</emphasis>
:
<treatmentCitation id="D2631095FFAA7117FE4289EAFD10EC65" author="Nonveiller" box="[439,670,1017,1040]" page="94" pageId="15" pageNumber="16" year="1996">
<bibRefCitation id="37534B75FFAA7117FE4289EAFDF8EC65" author="Nonveiller" box="[439,630,1017,1040]" pageId="15" pageNumber="49" refString="Nonveiller, G. (1996 b) Types de repartition dans la region Afrotropicale bases sur l'etude des Mutillides (Hymenoptera, Mutillidae). Bulletin de la Societe Entomologique de France, 101 (1), 89 - 112." type="journal article" year="1996" yearSuffix="b">Nonveiller 1996b</bibRefCitation>
: 94
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
</treatment>
</document>

120
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<emphasis id="61B6EA96FFAC7111FEFD8FA2FD93EDBE" bold="true" box="[264,541,1457,1483]" italics="true" pageId="9" pageNumber="10">Arnoldtilla achterbergi</emphasis>
Lelej
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,
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(
<figureCitation id="CBF92A01FFAC7111FEE58FC7FEF6ED9B" box="[272,376,1492,1518]" captionStart="FIGURES 2 – 10. 2 – 5" captionStartId="8.[264,382,1060,1084]" captionTargetBox="[541,1044,411,1033]" captionTargetId="figure@8.[535,1051,406,1036]" captionTargetPageId="8" captionText="FIGURES 2 – 10. 2 – 5. Pseudophotopsis mascatiana, male. 2. Forewing; 3. Head; 4. Metasomal tergum 1, dorsal view; 5. Mandible, lateral view. 6 – 8. Arnoldtilla achterbergi sp. nov., female, holotype. 6. Habitus, dorsal view; 7. Mandible, dorsal view; 8. Pedicel and flagellomeres 1 – 3. 9 – 10. Omotilla conjuncta, male. 9. Mandible, dorsal view; 10. Genitalia, ventral view. Scale = 1 mm for 2 – 4, 6 and 0.5 mm for 5, 7 – 10." httpUri="https://zenodo.org/record/172673/files/figure.png" pageId="9" pageNumber="10">Figs 6–8</figureCitation>
)
</paragraph>
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<subSubSection id="1BD8650FFFAC7112FEFD8C37FD9BEC23" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="description">
<paragraph id="537D3684FFAC7111FEFD8C37FE1CEE48" blockId="9.[264,1325,1572,1878]" box="[264,402,1572,1597]" pageId="9" pageNumber="10">
<emphasis id="61B6EA96FFAC7111FEFD8C37FE1CEE48" box="[264,402,1572,1597]" italics="true" pageId="9" pageNumber="10">Description</emphasis>
</paragraph>
<paragraph id="537D3684FFAC7112FECD8C5FFB77EA3B" blockId="9.[264,1325,1572,1878]" lastBlockId="10.[264,1324,284,1110]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
FEMALE (
<figureCitation id="CBF92A01FFAC7111FE488C5FFD8BEE13" box="[445,517,1612,1638]" captionStart="FIGURES 2 – 10. 2 – 5" captionStartId="8.[264,382,1060,1084]" captionTargetBox="[541,1044,411,1033]" captionTargetId="figure@8.[535,1051,406,1036]" captionTargetPageId="8" captionText="FIGURES 2 – 10. 2 – 5. Pseudophotopsis mascatiana, male. 2. Forewing; 3. Head; 4. Metasomal tergum 1, dorsal view; 5. Mandible, lateral view. 6 – 8. Arnoldtilla achterbergi sp. nov., female, holotype. 6. Habitus, dorsal view; 7. Mandible, dorsal view; 8. Pedicel and flagellomeres 1 – 3. 9 – 10. Omotilla conjuncta, male. 9. Mandible, dorsal view; 10. Genitalia, ventral view. Scale = 1 mm for 2 – 4, 6 and 0.5 mm for 5, 7 – 10." httpUri="https://zenodo.org/record/172673/files/figure.png" pageId="9" pageNumber="10">Fig. 6</figureCitation>
). Body length
<quantity id="943A9B61FFAC7111FD478C5FFC9EEE13" box="[690,784,1612,1638]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="9" pageNumber="10" unit="mm" value="4.4">4.4 mm</quantity>
. Head large, wide, 1.4X wider than pronotum, area behind eyes equal to eye length. Genal carina not developed. Clypeus anteriorly not dentate. Carina between supraantennal tubercle and compound eye well developed. Hypostomal bridge medially not tuberculate. Mandible tridentate, beneath not excised, inner border with two preapical teeth (
<figureCitation id="CBF92A01FFAC7111FD208CFFFC90EF73" box="[725,798,1772,1798]" captionStart="FIGURES 2 – 10. 2 – 5" captionStartId="8.[264,382,1060,1084]" captionTargetBox="[541,1044,411,1033]" captionTargetId="figure@8.[535,1051,406,1036]" captionTargetPageId="8" captionText="FIGURES 2 – 10. 2 – 5. Pseudophotopsis mascatiana, male. 2. Forewing; 3. Head; 4. Metasomal tergum 1, dorsal view; 5. Mandible, lateral view. 6 – 8. Arnoldtilla achterbergi sp. nov., female, holotype. 6. Habitus, dorsal view; 7. Mandible, dorsal view; 8. Pedicel and flagellomeres 1 – 3. 9 – 10. Omotilla conjuncta, male. 9. Mandible, dorsal view; 10. Genitalia, ventral view. Scale = 1 mm for 2 – 4, 6 and 0.5 mm for 5, 7 – 10." httpUri="https://zenodo.org/record/172673/files/figure.png" pageId="9" pageNumber="10">Fig. 7</figureCitation>
). Scape curved, flagellomeres not flattened, pedicel long (
<figureCitation id="CBF92A01FFAC7111FE5E8D07FE7DEF5B" box="[427,499,1812,1838]" captionStart="FIGURES 2 – 10. 2 – 5" captionStartId="8.[264,382,1060,1084]" captionTargetBox="[541,1044,411,1033]" captionTargetId="figure@8.[535,1051,406,1036]" captionTargetPageId="8" captionText="FIGURES 2 – 10. 2 – 5. Pseudophotopsis mascatiana, male. 2. Forewing; 3. Head; 4. Metasomal tergum 1, dorsal view; 5. Mandible, lateral view. 6 – 8. Arnoldtilla achterbergi sp. nov., female, holotype. 6. Habitus, dorsal view; 7. Mandible, dorsal view; 8. Pedicel and flagellomeres 1 – 3. 9 – 10. Omotilla conjuncta, male. 9. Mandible, dorsal view; 10. Genitalia, ventral view. Scale = 1 mm for 2 – 4, 6 and 0.5 mm for 5, 7 – 10." httpUri="https://zenodo.org/record/172673/files/figure.png" pageId="9" pageNumber="10">Fig. 8</figureCitation>
), relation of pedicel and first three flagellomeres 1.0: 1.5: 1.0: 0.9. Frons and vertex with dense punctures. Mesosoma 1.35X longer than minimal width (before propodeal spiracles). Mesosoma with acuminate humeral angle and weak tubercles laterally. Mesopleura without precoxal carina. Propodeum posteriorly with three well developed teeth. Fore coxa with blunt tubercle. Mesosoma dorsally coarsely punctured, laterally glabrous, posterior propodeal face reticulate in dorsal half. Metasomal segment 1 broad, anterior and dorsal surfaces not divided by transverse carina. T1 baso­laterally with lamellate carina. Lateral felt line located on T2 only. S2 without median basal carina, not swollen postero­laterally. S6 without any tubercles. T6 without distinct pygidial area, glabrous shiny rounded area does not touch base, nor carinated laterally in apical part.
</paragraph>
<paragraph id="537D3684FFAF7112FECD884FFCA4EC5B" blockId="10.[264,1324,284,1110]" pageId="10" pageNumber="11">Head yellowish­red, frons somewhat fuscous; antenna yellowish­red, flagellomeres above brownish; mandible yellowish­red with darker denticles; palps pale brown. Mesosoma and legs yellowish­red, tibial spurs whitish. Metasoma brown, T1, T6 yellowish­red, sterna pale brown. Head, mandible, scape with sparse long erect and short recumbent yellowish setae, brownish ones on frons. Mesosoma dorsally with long sparse erect and dense recumbent yellowish setae, laterally with whitish recumbent micropubescence. Legs with long erect yellowish setae. T1 dorsally with band of dense yellowish setae. T2 basally with large median spot of yellowish setae and apically with narrow band widened medially of dense yellowish setae. T3 completely covered by dense yellowish setae. T2 and T6 laterally, T4 and T5 wholly with sparse long erect yellowish setae. Lateral felt line on T2 yellowish. Metasoma ventrally with sparse yellowish setae and apical fringes of yellowsh setae on S2–S5.</paragraph>
<paragraph id="537D3684FFAF7112FECD8E2FFD9BEC23" blockId="10.[264,1324,284,1110]" box="[312,533,1084,1110]" pageId="10" pageNumber="11">MALE. Unknown.</paragraph>
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<emphasis id="61B6EA96FFAF7112FEFD8E9FFE27ECD0" box="[264,425,1164,1189]" italics="true" pageId="10" pageNumber="11">
<typeStatus id="8C798826FFAF7112FEFD8E9FFEB1ECD0" box="[264,319,1164,1189]" pageId="10" pageNumber="11">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="537D3684FFAF7112FECD8EA7FBF6ECBB" blockId="10.[264,1144,1164,1230]" box="[312,1144,1203,1230]" pageId="10" pageNumber="11">
<typeStatus id="8C798826FFAF7112FECD8EA7FE26ECBB" box="[312,424,1204,1230]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
, Ψ,
<collectingCountry id="2BD57614FFAF7112FE208EA7FDA2ECBB" box="[469,556,1204,1230]" name="Yemen" pageId="10" pageNumber="11">Yemen</collectingCountry>
: Wadi Hadramaut,
<date id="277C1044FFAF7112FCF98EA7FC33ECBB" box="[780,957,1204,1230]" pageId="10" pageNumber="11" value="1992-04-25" valueMax="1992-04-27" valueMin="1992-04-25">25–27.IV.1992</date>
, AvH [
<collectionCode id="35D3AE41FFAF7112FBF88EA7FBE3ECBB" LSID="urn:lsid:biocol.org:col:34975" box="[1037,1133,1204,1230]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="10" pageNumber="11">ZMAN</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFAF7112FEFD8F17FE03ED33" box="[264,407,1284,1350]" pageId="10" pageNumber="11" type="distribution">
<paragraph id="537D3684FFAF7112FEFD8F17FE19ED68" blockId="10.[264,407,1284,1350]" box="[264,407,1284,1309]" pageId="10" pageNumber="11">
<emphasis id="61B6EA96FFAF7112FEFD8F17FE19ED68" box="[264,407,1284,1309]" italics="true" pageId="10" pageNumber="11">Distribution</emphasis>
</paragraph>
<paragraph id="537D3684FFAF7112FECD8F3FFE03ED33" blockId="10.[264,407,1284,1350]" box="[312,397,1324,1350]" pageId="10" pageNumber="11">
<collectingCountry id="2BD57614FFAF7112FECD8F3FFE09ED33" box="[312,391,1324,1350]" name="Yemen" pageId="10" pageNumber="11">Yemen</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFAF7112FEFD8F6FFD00EE7B" pageId="10" pageNumber="11" type="etymology">
<paragraph id="537D3684FFAF7112FEFD8F6FFE0BEDE0" blockId="10.[264,1323,1404,1550]" box="[264,389,1404,1429]" pageId="10" pageNumber="11">
<emphasis id="61B6EA96FFAF7112FEFD8F6FFE0BEDE0" box="[264,389,1404,1429]" italics="true" pageId="10" pageNumber="11">Etymology</emphasis>
</paragraph>
<paragraph id="537D3684FFAF7112FECD8FB7FD00EE7B" blockId="10.[264,1323,1404,1550]" pageId="10" pageNumber="11">
This species is dedicated to Kees van Achterberg, world authority of
<taxonomicName id="94C24D07FFAF7112FB868FB7FB77EDCB" box="[1139,1273,1444,1470]" class="Insecta" family="Braconidae" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Braconidae</taxonomicName>
and curator of the
<taxonomicName id="94C24D07FFAF7112FE438FDFFDD8ED93" box="[438,598,1484,1510]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
and Diptera collections in the National Museum of Natural History, Leiden,
<collectingCountry id="2BD57614FFAF7112FE388FE7FD04EE7B" box="[461,650,1524,1550]" name="Netherlands" pageId="10" pageNumber="11">the Netherlands</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

166
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<emphasis id="61B6EA96FFAD7110FEFD8F31FD66ED49" bold="true" box="[264,744,1314,1340]" pageId="8" pageNumber="9">
<emphasis id="61B6EA96FFAD7110FEFD8F31FE9AED49" bold="true" box="[264,276,1314,1340]" italics="true" pageId="8" pageNumber="9">*</emphasis>
<taxonomicName id="94C24D07FFAD7110FEE18F31FD66ED49" authority="Bischoff 1920" authorityName="Bischoff" authorityYear="1920" box="[276,744,1314,1340]" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="maura">
<emphasis id="61B6EA96FFAD7110FEE18F31FDB9ED49" bold="true" box="[276,567,1314,1340]" italics="true" pageId="8" pageNumber="9">Pseudophotopsis maura</emphasis>
Bischoff 1920
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFAD7110FEFD8F7EFB2EEDB1" pageId="8" pageNumber="9" type="reference_group">
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<treatmentCitationGroup id="73D211AAFFAD7110FEFD8F7EFB2EEDB1" pageId="8" pageNumber="9">
<taxonomicName id="94C24D07FFAD7110FEFD8F7EFCD7EDF1" authority="Bischoff 1920: 98" authorityName="Bischoff" authorityPageNumber="98" authorityYear="1920" box="[264,857,1389,1412]" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="subSpecies" species="kokpetica" subSpecies="maura">
<emphasis id="61B6EA96FFAD7110FEFD8F7EFD0FEDF1" box="[264,641,1389,1412]" italics="true" pageId="8" pageNumber="9">Pseudophotopsis kokpetica maura</emphasis>
Bischoff 1920: 98
</taxonomicName>
, ♂,
<typeStatus id="8C798826FFAD7110FC7B8F7EFC60EDF1" box="[910,1006,1389,1412]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
, ♂,
<collectingCountry id="2BD57614FFAD7110FBD68F7EFBFBEDF1" box="[1059,1141,1389,1412]" name="Tunisia" pageId="8" pageNumber="9">Tunisia</collectingCountry>
, &quot;Gabes, Weiss, 1906&quot;;
<treatmentCitation id="D2631095FFAD7110FE738F9EFDC3EDD1" author="Invrea" box="[390,589,1421,1444]" page="462" pageId="8" pageNumber="9" year="1932">
<bibRefCitation id="37534B75FFAD7110FE738F9EFD99EDD1" author="Invrea" box="[390,535,1421,1444]" pageId="8" pageNumber="48" refString="Invrea, F. (1932 b) Spedizione scientifica all'oasi di Cufra (marzo-luglio 1931). Mutillidae e Chrysididae (Hymenoptera). Annali del Museo Civico di Storia Naturale Giacomo Doria, 55, 457 - 465. [Mutillidae - p. 458 - 464, 1 fig.]." type="journal article" year="1932" yearSuffix="b">Invrea 1932b</bibRefCitation>
: 462
</treatmentCitation>
, ♂ (
<collectingCountry id="2BD57614FFAD7110FD828F9EFD35EDD1" box="[631,699,1421,1444]" name="Libya" pageId="8" pageNumber="9">Libya</collectingCountry>
); 1932b: 66, ♂ (
<collectingCountry id="2BD57614FFAD7110FC938F9EFC24EDD1" box="[870,938,1421,1444]" name="Libya" pageId="8" pageNumber="9">Libya</collectingCountry>
);
<treatmentCitation id="D2631095FFAD7110FC488F9EFBF4EDD1" author="Suarez" box="[957,1146,1421,1444]" page="920" pageId="8" pageNumber="9" year="1963">
<bibRefCitation id="37534B75FFAD7110FC488F9EFBCBEDD1" author="Suarez" box="[957,1093,1421,1444]" pageId="8" pageNumber="50" refString="Suarez, F. J. (1963) Hymenopteres Tiphiides, Apterogynides et mutillides recoltes par J. Mateu dans l'Ennedi et au Tchad. Bulletin de l'Institut fondamental d'Afrique Noire, Dakar. Serie A: Sciences Naturelles, 25 (3), 912 - 943, 10 figs." type="journal article" year="1963">Suárez 1963</bibRefCitation>
: 920
</treatmentCitation>
, ♂ (
<collectingCountry id="2BD57614FFAD7110FB5E8F9EFB67EDD1" box="[1195,1257,1421,1444]" name="Chad" pageId="8" pageNumber="9">Chad</collectingCountry>
).
<taxonomicName id="94C24D07FFAD7110FEFD8FBEFD40EDB1" authority="Invrea 1965a: 62" authorityName="Invrea" authorityPageNumber="62" authorityYear="1965" box="[264,718,1453,1476]" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="maura">
<emphasis id="61B6EA96FFAD7110FEFD8FBEFD87EDB1" box="[264,521,1453,1476]" italics="true" pageId="8" pageNumber="9">Pseudophotopsis maura</emphasis>
:
<treatmentCitation id="D2631095FFAD7110FDE28FBEFD40EDB1" author="Invrea" box="[535,718,1453,1476]" page="62" pageId="8" pageNumber="9" year="1965">
<bibRefCitation id="37534B75FFAD7110FDE28FBEFD26EDB1" author="Invrea" box="[535,680,1453,1476]" pageId="8" pageNumber="48" refString="Invrea, F. (1965 a) Studi sugli Apterogynidi e Mutillidi della Palestina (Hymenoptera). Memorie della Societa Entomologica Italiana, 44, 53 - 93." type="journal article" year="1965" yearSuffix="a">Invrea 1965a</bibRefCitation>
: 62
</treatmentCitation>
</taxonomicName>
, ♂ (
<collectingCountry id="2BD57614FFAD7110FD0C8FBEFCB4EDB1" box="[761,826,1453,1476]" name="Israel" pageId="8" pageNumber="9">Israel</collectingCountry>
);
<treatmentCitation id="D2631095FFAD7110FCB98FBEFBBFEDB1" author="Nonveiller" box="[844,1073,1453,1476]" page="103" pageId="8" pageNumber="9" year="1974">
<bibRefCitation id="37534B75FFAD7110FCB98FBEFC70EDB1" author="Nonveiller" box="[844,1022,1453,1476]" pageId="8" pageNumber="49" refString="Nonveiller, G. (1974) Recherches sur les Mutillides de l'Afrique (Hymenoptera, Mutillidae). V. Description du male encore inconnu de Pseudophotopsis continua F. seul representant du genre en Afrique occidentale. Annales de la Faculte des Sciences du Cameroun, 18, 95 - 108." type="journal article" year="1974">Nonveiller 1974</bibRefCitation>
: 103
</treatmentCitation>
(
<collectingCountry id="2BD57614FFAD7110FBCA8FBEFB1BEDB1" box="[1087,1173,1453,1476]" name="Algeria" pageId="8" pageNumber="9">Algeria</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFAD7110FEFD8FE6FD0EEEA2" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="537D3684FFAD7110FEFD8FE6FE68EE7B" blockId="8.[264,1323,1525,1751]" box="[264,486,1525,1550]" pageId="8" pageNumber="9">
<emphasis id="61B6EA96FFAD7110FEFD8FE6FE68EE7B" box="[264,486,1525,1550]" italics="true" pageId="8" pageNumber="9">Material examined</emphasis>
</paragraph>
<paragraph id="537D3684FFAD7110FECD8C0EFD0EEEA2" blockId="8.[264,1323,1525,1751]" pageId="8" pageNumber="9">
<collectingCountry id="2BD57614FFAD7110FECD8C0EFE00EE42" box="[312,398,1565,1591]" name="Yemen" pageId="8" pageNumber="9">Yemen</collectingCountry>
: Al Kowd,
<date id="277C1044FFAD7110FDE18C0EFD05EE42" box="[532,651,1565,1591]" pageId="8" pageNumber="9" value="1999-08">VIII.1999</date>
, LT, AvH &amp; S. Al Haruri, 2 ɗ [
<collectionCode id="35D3AE41FFAD7110FC0C8C0EFBD6EE42" LSID="urn:lsid:biocol.org:col:34975" box="[1017,1112,1565,1591]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="8" pageNumber="9">ZMAN</collectionCode>
];
<date id="277C1044FFAD7110FB9E8C0EFB34EE42" box="[1131,1210,1565,1591]" pageId="8" pageNumber="9" value="2000-01">I.2000</date>
, LT, AvH &amp; S. Al Haruri, 1 ɗ [
<collectionCode id="35D3AE41FFAD7110FDF38C56FDE8EE2A" LSID="urn:lsid:biocol.org:col:34975" box="[518,614,1605,1631]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="8" pageNumber="9">ZMAN</collectionCode>
];
<date id="277C1044FFAD7110FD8F8C56FD5BEE2A" box="[634,725,1605,1631]" pageId="8" pageNumber="9" value="2000-10">X.2000</date>
, LT, AvH &amp; S. Al Haruri, 2 ɗ [
<collectionCode id="35D3AE41FFAD7110FBBB8C56FB20EE2A" LSID="urn:lsid:biocol.org:col:34975" box="[1102,1198,1605,1631]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="8" pageNumber="9">ZMAN</collectionCode>
];
<date id="277C1044FFAD7110FB378C56FAA8EE2A" box="[1218,1318,1605,1631]" pageId="8" pageNumber="9" value="2000-11">XI.2000</date>
, LT, AvH &amp; S. Al Haruri, 1 ɗ [
<collectionCode id="35D3AE41FFAD7110FD908C7EFD4BEEF2" LSID="urn:lsid:biocol.org:col:34975" box="[613,709,1645,1671]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="8" pageNumber="9">ZMAN</collectionCode>
];
<date id="277C1044FFAD7110FD228C7EFCBFEEF2" box="[727,817,1645,1671]" pageId="8" pageNumber="9" value="2003-10">X.2003</date>
, LT, AvH &amp; S. Al Haruri, 1 ɗ [
<collectionCode id="35D3AE41FFAD7110FB6C8C7EFB74EEF2" LSID="urn:lsid:biocol.org:col:34992" box="[1177,1274,1645,1671]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="8" pageNumber="9">RMNH</collectionCode>
];
<quantity id="943A9B61FFAD7110FAF88C7EFEA3EEDA" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="9" unit="km" value="12.0">12 km</quantity>
NW Manakhah,
<date id="277C1044FFAD7110FDF88C86FD78EEDA" box="[525,758,1685,1711]" pageId="8" pageNumber="9" value="2001-07-03" valueMax="2001-08-21" valueMin="2001-07-03">3.VII–21.VIII.2001</date>
, MT, AvH, 1 ɗ [
<collectionCode id="35D3AE41FFAD7110FC288C86FBB0EEDA" LSID="urn:lsid:biocol.org:col:34992" box="[989,1086,1685,1711]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="8" pageNumber="9">RMNH</collectionCode>
];
<collectingCountry id="2BD57614FFAD7110FBAC8C86FB3BEEDA" box="[1113,1205,1685,1711]" name="Yemen" pageId="8" pageNumber="9">Socotra</collectingCountry>
, Nogeed,
<date id="277C1044FFAD7110FEFD8CAEFE03EEA2" box="[264,397,1725,1751]" pageId="8" pageNumber="9" value="1993-04-16">16.IV.1993</date>
, AvH, 1 ɗ [
<collectionCode id="35D3AE41FFAD7110FDE38CAEFDF8EEA2" box="[534,630,1725,1751]" pageId="8" pageNumber="9">ZMAH</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFAD7110FEFD8D1EFB48EF3A" box="[264,1222,1805,1871]" pageId="8" pageNumber="9" type="distribution">
<paragraph id="537D3684FFAD7110FEFD8D1EFE19EF53" blockId="8.[264,1222,1805,1871]" box="[264,407,1805,1830]" pageId="8" pageNumber="9">
<emphasis id="61B6EA96FFAD7110FEFD8D1EFE19EF53" box="[264,407,1805,1830]" italics="true" pageId="8" pageNumber="9">Distribution</emphasis>
</paragraph>
<paragraph id="537D3684FFAD7110FECD8D26FB48EF3A" blockId="8.[264,1222,1805,1871]" box="[312,1222,1845,1871]" pageId="8" pageNumber="9">
<collectingCountry id="2BD57614FFAD7110FECD8D26FE04EF3A" box="[312,394,1845,1871]" name="Yemen" pageId="8" pageNumber="9">Yemen</collectingCountry>
(new record),
<collectingCountry id="2BD57614FFAD7110FDC38D26FDF4EF3A" box="[566,634,1845,1871]" name="Israel" pageId="8" pageNumber="9">Israel</collectingCountry>
,
<collectingCountry id="2BD57614FFAD7110FD738D26FD40EF3A" box="[646,718,1845,1871]" name="Libya" pageId="8" pageNumber="9">Libya</collectingCountry>
,
<collectingCountry id="2BD57614FFAD7110FD2C8D26FC94EF3A" box="[729,794,1845,1871]" name="Chad" pageId="8" pageNumber="9">Chad</collectingCountry>
,
<collectingCountry id="2BD57614FFAD7110FCD38D26FC0EEF3A" box="[806,896,1845,1871]" name="Tunisia" pageId="8" pageNumber="9">Tunisia</collectingCountry>
,
<collectingCountry id="2BD57614FFAD7110FC798D26FC6AEF3A" box="[908,996,1845,1871]" name="Algeria" pageId="8" pageNumber="9">Algeria</collectingCountry>
(
<bibRefCitation id="37534B75FFAD7110FC068D26FB37EF3A" author="Nonveiller" box="[1011,1209,1845,1871]" pageId="8" pageNumber="49" refString="Nonveiller, G. (1974) Recherches sur les Mutillides de l'Afrique (Hymenoptera, Mutillidae). V. Description du male encore inconnu de Pseudophotopsis continua F. seul representant du genre en Afrique occidentale. Annales de la Faculte des Sciences du Cameroun, 18, 95 - 108." type="journal article" year="1974">Nonveiller 1974</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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212
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<document id="E75E1DD6559C5AA23238786A4CEF7D39" ID-DOI="10.5281/zenodo.172671" ID-GBIF-Dataset="b0d82648-af13-4bba-8318-49fa28bd2515" ID-ISSN="1175­5326" ID-Zenodo-Dep="172671" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459760090076" checkinUser="plazi" docAuthor="Lelej, Arkady S. &amp; Harten, Antonius Van" docDate="2006" docId="DB6B8792FFAE7113FEFD8B08FDBAEDD4" docLanguage="en" docName="zt01226p050.pdf" docOrigin="Zootaxa 1226" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Myrmilla parvula Magretti 1906, stat. nov." docType="treatment" docVersion="8" lastPageNumber="12" masterDocId="2752FFEAFFA57118FFF58A13FF8EE875" masterDocTitle="A review of the Mutillidae (Hymenoptera) of Yemen" masterLastPageNumber="50" masterPageNumber="1" pageNumber="12" updateTime="1698214875622" updateUser="plazi">
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<mods:title id="F48CF0FA6A2A18989D7BE38B62E5E0D7">A review of the Mutillidae (Hymenoptera) of Yemen</mods:title>
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<mods:namePart id="9D052133589C965781704DB298812883">Lelej, Arkady S.</mods:namePart>
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<mods:namePart id="7BEE434AEB580B8AADA9B1E99AD1398B">Harten, Antonius Van</mods:namePart>
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<mods:date id="4B3E7687602239FA5192F9FCAAE43515">2006</mods:date>
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<treatment id="DB6B8792FFAE7113FEFD8B08FDBAEDD4" ID-DOI="http://doi.org/10.5281/zenodo.6252939" ID-GBIF-Taxon="119344648" ID-Zenodo-Dep="6252939" LSID="urn:lsid:plazi:treatment:DB6B8792FFAE7113FEFD8B08FDBAEDD4" httpUri="http://treatment.plazi.org/id/DB6B8792FFAE7113FEFD8B08FDBAEDD4" lastPageNumber="12" pageId="11" pageNumber="12">
<subSubSection id="1BD8650FFFAE7113FEFD8B08FE10E9E8" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="537D3684FFAE7113FEFD8B08FCAFE940" blockId="11.[264,801,283,310]" box="[264,801,283,310]" pageId="11" pageNumber="12">
<heading id="083581E8FFAE7113FEFD8B08FCAFE940" bold="true" box="[264,801,283,310]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="61B6EA96FFAE7113FEFD8B08FCAFE940" bold="true" box="[264,801,283,310]" pageId="11" pageNumber="12">
*
<taxonomicName id="94C24D07FFAE7113FEE28B0FFD2AE940" authority="Magretti 1906" authorityName="Magretti" authorityYear="1906" box="[279,676,283,310]" class="Insecta" family="Mutillidae" genus="Myrmilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="parvula" status="stat. nov.">
<emphasis id="61B6EA96FFAE7113FEE28B0FFE65E943" bold="true" box="[279,491,284,310]" italics="true" pageId="11" pageNumber="12">Myrmilla parvula</emphasis>
Magretti 1906
</taxonomicName>
,
<taxonomicNameLabel id="7A8557EDFFAE7113FD448B08FCAFE940" box="[689,801,283,309]" pageId="11" pageNumber="12">stat. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="537D3684FFAE7113FEFD8B74FE10E9E8" blockId="11.[264,1322,358,446]" pageId="11" pageNumber="12">
<taxonomicName id="94C24D07FFAE7113FEFD8B74FE53E90B" box="[264,477,359,382]" class="Insecta" family="Mutillidae" genus="Myrmilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="cephalica">
<emphasis id="61B6EA96FFAE7113FEFD8B74FE53E90B" box="[264,477,359,382]" italics="true" pageId="11" pageNumber="12">Myrmilla cephalica</emphasis>
</taxonomicName>
<taxonomicName id="94C24D07FFAE7113FE1C8B75FCCAE908" authority="Magretti 1906: 41" authorityName="Magretti" authorityPageNumber="41" authorityYear="1906" box="[489,836,358,382]" class="Insecta" family="Mutillidae" genus="Myrmilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="variety" species="rubida" variety="parvula">
var.
<emphasis id="61B6EA96FFAE7113FDEE8B74FDE0E90B" box="[539,622,359,382]" italics="true" pageId="11" pageNumber="12">parvula</emphasis>
Magretti 1906: 41
</taxonomicName>
, Ψ,
<typeStatus id="8C798826FFAE7113FC838B75FC5DE908" box="[886,979,358,381]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
&quot;Adi­Caié, Andreini&quot; (
<collectingCountry id="2BD57614FFAE7113FB228B75FAADE908" box="[1239,1315,358,381]" name="Eritrea" pageId="11" pageNumber="12">Eritrea</collectingCountry>
) [MSNG].
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFAE7113FEFD8BB4FC07E9C8" box="[264,905,422,446]" pageId="11" pageNumber="12" type="reference_group">
<paragraph id="537D3684FFAE7113FEFD8BB4FC07E9C8" blockId="11.[264,1322,358,446]" box="[264,905,422,446]" pageId="11" pageNumber="12">
<taxonomicName id="94C24D07FFAE7113FEFD8BB4FC8AE9C8" authority="Bischoff 1920: 58" authorityName="Bischoff" authorityPageNumber="58" authorityYear="1920" box="[264,772,422,446]" class="Insecta" family="Mutillidae" genus="Myrmilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subSpecies" species="cephalica" subSpecies="parvula">
<emphasis id="61B6EA96FFAE7113FEFD8BB4FDBDE9CB" box="[264,563,423,446]" italics="true" pageId="11" pageNumber="12">Myrmilla cephalica parvula</emphasis>
: Bischoff 1920: 58
</taxonomicName>
, Ψ (
<collectingCountry id="2BD57614FFAE7113FCDB8BB5FCF3E9C8" box="[814,893,422,445]" name="Eritrea" pageId="11" pageNumber="12">Eritrea</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFAE7113FEFD8BFCFD93EB54" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="537D3684FFAE7113FEFD8BFCFEF1EA7D" blockId="11.[264,1323,495,801]" box="[264,383,495,520]" pageId="11" pageNumber="12">
<emphasis id="61B6EA96FFAE7113FEFD8BFCFEF1EA7D" box="[264,383,495,520]" italics="true" pageId="11" pageNumber="12">Diagnosis</emphasis>
</paragraph>
<paragraph id="537D3684FFAE7113FECD8804FD93EB54" blockId="11.[264,1323,495,801]" pageId="11" pageNumber="12">
The female of
<taxonomicName id="94C24D07FFAE7113FE1D8804FDE2EA45" box="[488,620,535,560]" class="Insecta" family="Mutillidae" genus="Myrmilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="parvula">
<emphasis id="61B6EA96FFAE7113FE1D8804FDE2EA45" box="[488,620,535,560]" italics="true" pageId="11" pageNumber="12">M. parvula</emphasis>
</taxonomicName>
can be distinguished from that of
<taxonomicName id="94C24D07FFAE7113FBFC8804FB0EEA45" box="[1033,1152,535,560]" class="Insecta" family="Mutillidae" genus="Myrmilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="rubida">
<emphasis id="61B6EA96FFAE7113FBFC8804FB0EEA45" box="[1033,1152,535,560]" italics="true" pageId="11" pageNumber="12">M. rubida</emphasis>
</taxonomicName>
by its smaller size (4.0 mm against 6.0–7.0 mm in
<taxonomicName id="94C24D07FFAE7113FD25882CFCC7EA2D" box="[720,841,575,600]" class="Insecta" family="Mutillidae" genus="Myrmilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="rubida">
<emphasis id="61B6EA96FFAE7113FD25882CFCC7EA2D" box="[720,841,575,600]" italics="true" pageId="11" pageNumber="12">M. rubida</emphasis>
</taxonomicName>
), in lacking a humeral projection (well developed in
<taxonomicName id="94C24D07FFAE7113FE5E8874FDACEAF5" box="[427,546,615,640]" class="Insecta" family="Mutillidae" genus="Myrmilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="rubida">
<emphasis id="61B6EA96FFAE7113FE5E8874FDACEAF5" box="[427,546,615,640]" italics="true" pageId="11" pageNumber="12">M. rubida</emphasis>
</taxonomicName>
), and in having the frons darkened. The male of
<taxonomicName id="94C24D07FFAE7113FB808874FB74EAF5" box="[1141,1274,615,640]" class="Insecta" family="Mutillidae" genus="Myrmilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="parvula">
<emphasis id="61B6EA96FFAE7113FB808874FB74EAF5" box="[1141,1274,615,640]" italics="true" pageId="11" pageNumber="12">M. parvula</emphasis>
</taxonomicName>
has similarly shaped mandibles and body coloration to the male of
<taxonomicName id="94C24D07FFAE7113FBFE889CFB0DEADD" box="[1035,1155,655,680]" class="Insecta" family="Mutillidae" genus="Myrmilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="rubida">
<emphasis id="61B6EA96FFAE7113FBFE889CFB0DEADD" box="[1035,1155,655,680]" italics="true" pageId="11" pageNumber="12">M. rubida</emphasis>
</taxonomicName>
but differs in having smaller size (4.8–6.0 mm against
<quantity id="943A9B61FFAE7113FD0588A4FCE6EAA4" box="[752,872,695,721]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" metricValueMax="1.9304" metricValueMin="1.6256" pageId="11" pageNumber="12" unit="in" value="7.0" valueMax="7.6" valueMin="6.4">6.4–7.6 in</quantity>
<taxonomicName id="94C24D07FFAE7113FC8588A4FC68EAA5" box="[880,998,695,720]" class="Insecta" family="Mutillidae" genus="Myrmilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="rubida">
<emphasis id="61B6EA96FFAE7113FC8588A4FC68EAA5" box="[880,998,695,720]" italics="true" pageId="11" pageNumber="12">M. rubida</emphasis>
</taxonomicName>
), in having well developed denticles on the anterior clypeal border (weak in
<taxonomicName id="94C24D07FFAE7113FCB288CCFC32EA8D" box="[839,956,735,760]" class="Insecta" family="Mutillidae" genus="Myrmilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="rubida">
<emphasis id="61B6EA96FFAE7113FCB288CCFC32EA8D" box="[839,956,735,760]" italics="true" pageId="11" pageNumber="12">M. rubida</emphasis>
</taxonomicName>
), and in having a shorter, more acuminated gonostylus.
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFAE7113FEFD8944FE18EC14" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="537D3684FFAE7113FEFD8944FE68EB05" blockId="11.[264,1323,855,1121]" box="[264,486,855,880]" pageId="11" pageNumber="12">
<emphasis id="61B6EA96FFAE7113FEFD8944FE68EB05" box="[264,486,855,880]" italics="true" pageId="11" pageNumber="12">Material examined</emphasis>
</paragraph>
<paragraph id="537D3684FFAE7113FECD896CFE18EC14" blockId="11.[264,1323,855,1121]" pageId="11" pageNumber="12">
<collectingCountry id="2BD57614FFAE7113FECD896CFE00EBEC" box="[312,398,895,921]" name="Yemen" pageId="11" pageNumber="12">Yemen</collectingCountry>
: near Sanaa,
<date id="277C1044FFAE7113FDD2896CFD3DEBEC" box="[551,691,895,921]" pageId="11" pageNumber="12" value="1998-03" valueMax="1998-04-31" valueMin="1998-03-00">III–IV.1998</date>
, PT, AvH, 1 Ψ [
<collectionCode id="35D3AE41FFAE7113FC9F896CFC44EBEC" LSID="urn:lsid:biocol.org:col:34975" box="[874,970,895,921]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="11" pageNumber="12">ZMAN</collectionCode>
]; Sanaa,
<date id="277C1044FFAE7113FBC7896CFB1CEBEC" box="[1074,1170,895,921]" pageId="11" pageNumber="12" value="1998-04">IV.1998</date>
, MT, AvH, 2 ɗ [
<collectionCode id="35D3AE41FFAE7113FEDB89B4FE03EBB4" LSID="urn:lsid:biocol.org:col:34975" box="[302,397,935,961]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="11" pageNumber="12">ZMAN</collectionCode>
];
<date id="277C1044FFAE7113FE5089B4FD9CEBB4" box="[421,530,935,961]" pageId="11" pageNumber="12" value="1998-07">VII.1998</date>
, MT, AvH, 1 ɗ [
<collectionCode id="35D3AE41FFAE7113FD1F89B4FCC7EBB4" LSID="urn:lsid:biocol.org:col:34975" box="[746,841,935,961]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="11" pageNumber="12">ZMAN</collectionCode>
];
<date id="277C1044FFAE7113FC9489B4FC66EBB4" box="[865,1000,935,961]" pageId="11" pageNumber="12" value="1998-05" valueMax="1998-06-31" valueMin="1998-05-00">V–VI.1998</date>
, MT, AvH, 1 ɗ [
<collectionCode id="35D3AE41FFAE7113FB3589B4FA91EBB4" LSID="urn:lsid:biocol.org:col:34975" box="[1216,1311,935,961]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="11" pageNumber="12">ZMAN</collectionCode>
];
<date id="277C1044FFAE7113FEFD89DCFE1DEB9C" box="[264,403,975,1001]" pageId="11" pageNumber="12" value="1998-03" valueMax="1998-04-31" valueMin="1998-03-00">III–IV.1998</date>
, PT, AvH, 1 ɗ [
<collectionCode id="35D3AE41FFAE7113FDA489DCFD3FEB9C" LSID="urn:lsid:biocol.org:col:34975" box="[593,689,975,1001]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="11" pageNumber="12">ZMAN</collectionCode>
]; Al Kowd,
<date id="277C1044FFAE7113FCB789DCFC2FEB9C" box="[834,929,975,1001]" pageId="11" pageNumber="12" value="1993-04">IV.1993</date>
, MT, AvH, 1 ɗ [
<collectionCode id="35D3AE41FFAE7113FB9E89DCFB45EB9C" LSID="urn:lsid:biocol.org:col:34975" box="[1131,1227,975,1001]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="11" pageNumber="12">ZMAN</collectionCode>
];
<quantity id="943A9B61FFAE7113FB2A89DCFAA5EB9C" box="[1247,1323,975,1001]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="12" unit="km" value="12.0">12 km</quantity>
NW Manakhah,
<date id="277C1044FFAE7113FE3889E4FD38EC64" box="[461,694,1015,1041]" pageId="11" pageNumber="12" value="2001-07-03" valueMax="2001-08-21" valueMin="2001-07-03">3.VII–21.VIII.2001</date>
, MT, AvH, 2 ɗ [
<collectionCode id="35D3AE41FFAE7113FC8B89E4FC51EC64" LSID="urn:lsid:biocol.org:col:34992" box="[894,991,1015,1041]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="11" pageNumber="12">RMNH</collectionCode>
];
<date id="277C1044FFAE7113FC0189E4FB03EC64" box="[1012,1165,1015,1041]" pageId="11" pageNumber="12" value="2003-01-01" valueMax="2003-05-07" valueMin="2003-01-01">1.I–7.V.2003</date>
, MT, AvH, 1 ɗ [
<collectionCode id="35D3AE41FFAE7113FEDE8E0CFE02EC4C" LSID="urn:lsid:biocol.org:col:34992" box="[299,396,1055,1081]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="11" pageNumber="12">RMNH</collectionCode>
];
<date id="277C1044FFAE7113FE558E0CFDFEEC4C" box="[416,624,1055,1081]" pageId="11" pageNumber="12" value="2003-05-15" valueMax="2003-06-23" valueMin="2003-05-15">15.V–23.VI.2003</date>
, MT, AvH, 3 ɗ [
<collectionCode id="35D3AE41FFAE7113FCCC8E0CFC14EC4C" LSID="urn:lsid:biocol.org:col:34992" box="[825,922,1055,1081]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="11" pageNumber="12">RMNH</collectionCode>
];
<date id="277C1044FFAE7113FC5A8E0CFB03EC4C" box="[943,1165,1055,1081]" pageId="11" pageNumber="12" value="2003-06-24" valueMax="2003-08-04" valueMin="2003-06-24">24.VI–4.VIII.2003</date>
, MT, AvH, 2 ɗ [
<collectionCode id="35D3AE41FFAE7113FEDF8E54FE05EC14" LSID="urn:lsid:biocol.org:col:34992" box="[298,395,1095,1121]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="11" pageNumber="12">RMNH</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFAE7113FEFD8E84FD00ECAC" box="[264,654,1175,1241]" pageId="11" pageNumber="12" type="distribution">
<paragraph id="537D3684FFAE7113FEFD8E84FE19ECC5" blockId="11.[264,654,1175,1241]" box="[264,407,1175,1200]" pageId="11" pageNumber="12">
<emphasis id="61B6EA96FFAE7113FEFD8E84FE19ECC5" box="[264,407,1175,1200]" italics="true" pageId="11" pageNumber="12">Distribution</emphasis>
</paragraph>
<paragraph id="537D3684FFAE7113FECD8EACFD00ECAC" blockId="11.[264,654,1175,1241]" box="[312,654,1215,1241]" pageId="11" pageNumber="12">
<collectingCountry id="2BD57614FFAE7113FECD8EACFE04ECAC" box="[312,394,1215,1241]" name="Yemen" pageId="11" pageNumber="12">Yemen</collectingCountry>
(new record),
<collectingCountry id="2BD57614FFAE7113FDC38EACFD04ECAC" box="[566,650,1215,1241]" name="Eritrea" pageId="11" pageNumber="12">Eritrea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFAE7113FEFD8F1CFDBAEDD4" pageId="11" pageNumber="12" type="discussion">
<paragraph id="537D3684FFAE7113FEFD8F1CFEE0ED5D" blockId="11.[264,1322,1295,1441]" box="[264,366,1295,1320]" pageId="11" pageNumber="12">
<emphasis id="61B6EA96FFAE7113FEFD8F1CFEE0ED5D" box="[264,366,1295,1320]" italics="true" pageId="11" pageNumber="12">Remarks</emphasis>
</paragraph>
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The varieties
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<emphasis id="61B6EA96FFAE7113FE2A8F24FDA3ED25" box="[479,557,1335,1360]" italics="true" pageId="11" pageNumber="12">rubida</emphasis>
</taxonomicName>
and
<taxonomicName id="94C24D07FFAE7113FD9B8F24FD47ED25" box="[622,713,1335,1360]" class="Insecta" family="Mutillidae" genus="Myrmilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="parvula">
<emphasis id="61B6EA96FFAE7113FD9B8F24FD47ED25" box="[622,713,1335,1360]" italics="true" pageId="11" pageNumber="12">parvula</emphasis>
</taxonomicName>
described by Magretti (1906) have nothing to do with Palaearctic
<taxonomicName id="94C24D07FFAE7113FE3B8F4CFC9BED0C" authority="Fabricius 1775" authorityName="Fabricius" authorityYear="1775" box="[462,789,1375,1401]" class="Insecta" family="Mutillidae" genus="Myrmilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="glabrata">
<emphasis id="61B6EA96FFAE7113FE3B8F4CFDD2ED0D" box="[462,604,1375,1400]" italics="true" pageId="11" pageNumber="12">M. glabrata</emphasis>
Fabricius 1775
</taxonomicName>
(=
<taxonomicName id="94C24D07FFAE7113FCB48F4CFA92ED0C" authority="Sichel et Radoszkowski 1869" authorityName="Sichel et Radoszkowski" authorityYear="1869" box="[833,1308,1375,1401]" class="Insecta" family="Mutillidae" genus="Myrmilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="cephalica">
<emphasis id="61B6EA96FFAE7113FCB48F4CFC3DED0D" box="[833,947,1375,1400]" italics="true" pageId="11" pageNumber="12">cephalica</emphasis>
Sichel et Radoszkowski 1869
</taxonomicName>
), so we change their status.
</paragraph>
</subSubSection>
</treatment>
</document>

131
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*
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<emphasis id="61B6EA96FFAE7113FEE28FEDFE50EE6D" bold="true" box="[279,478,1534,1560]" italics="true" pageId="11" pageNumber="12">Myrmilla rubida</emphasis>
Magretti 1906
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,
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<taxonomicName id="94C24D07FFAE7113FEFD8C59FE54EE14" box="[264,474,1610,1633]" class="Insecta" family="Mutillidae" genus="Myrmilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="cephalica">
<emphasis id="61B6EA96FFAE7113FEFD8C59FE54EE14" box="[264,474,1610,1633]" italics="true" pageId="11" pageNumber="12">Myrmilla cephalica</emphasis>
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var.
<emphasis id="61B6EA96FFAE7113FDFB8C59FDDBEE14" box="[526,597,1610,1633]" italics="true" pageId="11" pageNumber="12">rubida</emphasis>
Magretti 1906: 41
</taxonomicName>
, Ψ,
<typeStatus id="8C798826FFAE7113FCBC8C5AFC29EE15" box="[841,935,1609,1632]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
&quot;Saati, Belli&quot; (
<collectingCountry id="2BD57614FFAE7113FBB28C5AFB1DEE15" box="[1095,1171,1609,1632]" name="Eritrea" pageId="11" pageNumber="12">Eritrea</collectingCountry>
) [MSNG].
<taxonomicName id="94C24D07FFAE7113FEFD8C79FD76EEF5" authority="Bischoff 1920: 58" authorityName="Bischoff" authorityPageNumber="58" authorityYear="1920" box="[264,760,1641,1665]" class="Insecta" family="Mutillidae" genus="Myrmilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subSpecies" species="cephalica" subSpecies="rubida">
<emphasis id="61B6EA96FFAE7113FEFD8C79FDA9EEF4" box="[264,551,1642,1665]" italics="true" pageId="11" pageNumber="12">Myrmilla cephalica rubida</emphasis>
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, Ψ ♂ (
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<emphasis id="61B6EA96FFAE7113FEFD8CCAFE68EE87" box="[264,486,1753,1778]" italics="true" pageId="11" pageNumber="12">Material examined</emphasis>
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<collectingCountry id="2BD57614FFAE7113FECD8D12FE01EF6E" box="[312,399,1793,1819]" name="Yemen" pageId="11" pageNumber="12">Yemen</collectingCountry>
: Sanaa,
<date id="277C1044FFAE7113FE1B8D12FD1CEF6E" box="[494,658,1793,1819]" pageId="11" pageNumber="12" value="1930-08-16">16.VIII. 1930</date>
, Zhenzhurist (N. Filippov), 2 Ψ [
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<date id="277C1044FFAE7113FB7F8D12FAA8EF6E" box="[1162,1318,1793,1819]" pageId="11" pageNumber="12" value="1930-08-20">20.VIII.1930</date>
, Zhenzhurist, 1 Ψ, 1 ɗ [
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<date id="277C1044FFAE7113FD788D3AFC94EF36" box="[653,794,1833,1859]" pageId="11" pageNumber="12" value="1931-08-08">8.VIII.1931</date>
, Zhenzhurist, 1 Ψ [
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,
<date id="277C1044FFAE7113FEFD8D42FE1EEF1E" box="[264,400,1873,1899]" pageId="11" pageNumber="12" value="1931-11-15">15.XI.1931</date>
, Zhenzhurist, 6 Ψ, 1 ɗ [
<collectionCode id="35D3AE41FFAE7113FD528D42FC85EF1E" box="[679,779,1873,1899]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="11" pageNumber="12">ZMMU</collectionCode>
];
<date id="277C1044FFAE7113FCEB8D42FC19EF1E" box="[798,919,1873,1899]" pageId="11" pageNumber="12" value="1998-09-08">8.IX.1998</date>
, MT, AvH, 1 ɗ [
<collectionCode id="35D3AE41FFAE7113FBAF8D42FB34EF1E" LSID="urn:lsid:biocol.org:col:34975" box="[1114,1210,1873,1899]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="11" pageNumber="12">ZMAN</collectionCode>
];
<date id="277C1044FFAE7113FB398D42FAA8EF1E" box="[1228,1318,1873,1899]" pageId="11" pageNumber="12" value="1998-10">X.1998</date>
, PT, AvH, 1 Ψ [
<collectionCode id="35D3AE41FFA97114FE4B8B0FFD90E943" LSID="urn:lsid:biocol.org:col:34975" box="[446,542,284,310]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="12" pageNumber="13">ZMAN</collectionCode>
]; Hodeida, 1930, Zhenzhurist, 1 ɗ [
<collectionCode id="35D3AE41FFA97114FC3B8B0FFBBCE943" box="[974,1074,284,310]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="12" pageNumber="13">ZMMU</collectionCode>
]; Ma’bar,
<date id="277C1044FFA97114FB5A8B0FFAA8E943" box="[1199,1318,284,310]" pageId="12" pageNumber="13" value="1992-08">VIII.1992</date>
, swept in alfalfa field, A. Drews, 1 ɗ [
<collectionCode id="35D3AE41FFA97114FD4A8B57FC91E92B" LSID="urn:lsid:biocol.org:col:34975" box="[703,799,324,350]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="12" pageNumber="13">ZMAN</collectionCode>
].
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<emphasis id="61B6EA96FFA97114FEFD8B87FE19E9D8" box="[264,407,404,429]" italics="true" pageId="12" pageNumber="13">Distribution</emphasis>
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<collectingCountry id="2BD57614FFA97114FECD8BAFFE04E9A3" box="[312,394,444,470]" name="Yemen" pageId="12" pageNumber="13">Yemen</collectingCountry>
(new record),
<collectingCountry id="2BD57614FFA97114FDC38BAFFD04E9A3" box="[566,650,444,470]" name="Eritrea" pageId="12" pageNumber="13">Eritrea</collectingCountry>
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<treatment id="DB6B8792FFB0710DFEFD89BDFB4DEDCE" ID-DOI="http://doi.org/10.5281/zenodo.6252963" ID-GBIF-Taxon="119344638" ID-Zenodo-Dep="6252963" LSID="urn:lsid:plazi:treatment:DB6B8792FFB0710DFEFD89BDFB4DEDCE" httpUri="http://treatment.plazi.org/id/DB6B8792FFB0710DFEFD89BDFB4DEDCE" lastPageNumber="22" pageId="21" pageNumber="22">
<subSubSection id="1BD8650FFFB0710DFEFD89BDFCA0EBBD" box="[264,814,942,968]" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="537D3684FFB0710DFEFD89BDFCA0EBBD" blockId="21.[264,814,942,968]" box="[264,814,942,968]" pageId="21" pageNumber="22">
<heading id="083581E8FFB0710DFEFD89BDFCA0EBBD" bold="true" box="[264,814,942,968]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<emphasis id="61B6EA96FFB0710DFEFD89BDFCA0EBBD" bold="true" box="[264,814,942,968]" pageId="21" pageNumber="22">
<taxonomicName id="94C24D07FFB0710DFEFD89BDFD14EBBD" authority="Hammer 1962" authorityName="Hammer" authorityYear="1962" box="[264,666,942,968]" class="Insecta" family="Mutillidae" genus="Dentilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="arabica" status="comb. nov.">
<emphasis id="61B6EA96FFB0710DFEFD89BDFE42EBBD" bold="true" box="[264,460,942,968]" italics="true" pageId="21" pageNumber="22">Dentilla arabica</emphasis>
(Hammer 1962)
</taxonomicName>
,
<taxonomicNameLabel id="7A8557EDFFB0710DFD5D89BDFCA0EBBD" box="[680,814,942,968]" pageId="21" pageNumber="22">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB0710DFEFD89E9FCB1EC45" pageId="21" pageNumber="22" type="reference_group">
<paragraph id="537D3684FFB0710DFEFD89E9FCB1EC45" blockId="21.[264,1322,1017,1072]" pageId="21" pageNumber="22">
<taxonomicName id="94C24D07FFB0710DFEFD89E9FD46EC65" authority="Hammer 1962: 32" authorityName="Hammer" authorityPageNumber="32" authorityYear="1962" box="[264,712,1017,1041]" class="Insecta" family="Mutillidae" genus="Smicromyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="arabica">
<emphasis id="61B6EA96FFB0710DFEFD89E9FE79EC64" box="[264,503,1018,1041]" italics="true" pageId="21" pageNumber="22">Smicromyrme arabica</emphasis>
Hammer 1962: 32
</taxonomicName>
, Ψ,
<typeStatus id="8C798826FFB0710DFD0389EAFCDDEC65" box="[758,851,1017,1040]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
— &quot;Fiyush Dist., Aden, leg. Sewell, 1915, Desert fauna&quot; [Indian Museum, Kolikata,
<collectingCountry id="2BD57614FFB0710DFD0D8E0AFCBAEC45" box="[760,820,1049,1072]" name="India" pageId="21" pageNumber="22">India</collectingCountry>
].
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB0710DFEFD8E72FC17ECD6" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="537D3684FFB0710DFEFD8E72FE68EC0F" blockId="21.[264,921,1121,1187]" box="[264,486,1121,1146]" pageId="21" pageNumber="22">
<emphasis id="61B6EA96FFB0710DFEFD8E72FE68EC0F" box="[264,486,1121,1146]" italics="true" pageId="21" pageNumber="22">Material examined</emphasis>
</paragraph>
<paragraph id="537D3684FFB0710DFECD8E9AFC17ECD6" blockId="21.[264,921,1121,1187]" box="[312,921,1161,1187]" pageId="21" pageNumber="22">
<collectingCountry id="2BD57614FFB0710DFECD8E9AFE03ECD6" box="[312,397,1161,1187]" name="Yemen" pageId="21" pageNumber="22">Yemen</collectingCountry>
, Wadi Bana,
<date id="277C1044FFB0710DFDD28E9AFD2BECD6" box="[551,677,1161,1187]" pageId="21" pageNumber="22" value="1992-10-26">26.X.1992</date>
, AvH, 1 Ψ [
<collectionCode id="35D3AE41FFB0710DFCDA8E9AFC00ECD6" LSID="urn:lsid:biocol.org:col:34975" box="[815,910,1161,1187]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="21" pageNumber="22">ZMAN</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB0710DFEFD8ECAFDDAED6E" box="[264,596,1241,1307]" pageId="21" pageNumber="22" type="distribution">
<paragraph id="537D3684FFB0710DFEFD8ECAFE19EC87" blockId="21.[264,596,1241,1307]" box="[264,407,1241,1266]" pageId="21" pageNumber="22">
<emphasis id="61B6EA96FFB0710DFEFD8ECAFE19EC87" box="[264,407,1241,1266]" italics="true" pageId="21" pageNumber="22">Distribution</emphasis>
</paragraph>
<paragraph id="537D3684FFB0710DFECD8F12FDDAED6E" blockId="21.[264,596,1241,1307]" box="[312,596,1281,1307]" pageId="21" pageNumber="22">
<collectingCountry id="2BD57614FFB0710DFECD8F12FE04ED6E" box="[312,394,1281,1307]" name="Yemen" pageId="21" pageNumber="22">Yemen</collectingCountry>
(Hammer 1962).
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB0710DFEFD8F42FB4DEDCE" pageId="21" pageNumber="22" type="discussion">
<paragraph id="537D3684FFB0710DFEFD8F42FEE0ED1F" blockId="21.[264,1323,1361,1467]" box="[264,366,1361,1386]" pageId="21" pageNumber="22">
<emphasis id="61B6EA96FFB0710DFEFD8F42FEE0ED1F" box="[264,366,1361,1386]" italics="true" pageId="21" pageNumber="22">Remarks</emphasis>
</paragraph>
<paragraph id="537D3684FFB0710DFECD8F6AFB4DEDCE" blockId="21.[264,1323,1361,1467]" pageId="21" pageNumber="22">
It is quite possible that
<taxonomicName id="94C24D07FFB0710DFD918F6AFCA5EDE7" box="[612,811,1401,1426]" class="Insecta" family="Mutillidae" genus="Dentilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="arabica">
<emphasis id="61B6EA96FFB0710DFD918F6AFCA5EDE7" box="[612,811,1401,1426]" italics="true" pageId="21" pageNumber="22">Dentilla arabica</emphasis>
</taxonomicName>
is the opposite sex of
<taxonomicName id="94C24D07FFB0710DFBA68F6AFB52EDE7" box="[1107,1244,1401,1426]" class="Insecta" family="Mutillidae" genus="Dentilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="61B6EA96FFB0710DFBA68F6AFB52EDE7" box="[1107,1244,1401,1426]" italics="true" pageId="21" pageNumber="22">D. testacea</emphasis>
</taxonomicName>
or
<taxonomicName id="94C24D07FFB0710DFAFB8F6AFE1AEDCF" class="Insecta" family="Mutillidae" genus="Dentilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="echrenbergi">
<emphasis id="61B6EA96FFB0710DFAFB8F6AFE1AEDCF" italics="true" pageId="21" pageNumber="22">D. echrenbergi</emphasis>
</taxonomicName>
. Both male species have been collected in Al Kowd near Wadi Bana.
</paragraph>
</subSubSection>
</treatment>
</document>

226
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<treatment id="DB6B8792FFB0710FFEFD8C0AFD81EB63" ID-DOI="http://doi.org/10.5281/zenodo.6252965" ID-GBIF-Taxon="119344639" ID-Zenodo-Dep="6252965" LSID="urn:lsid:plazi:treatment:DB6B8792FFB0710FFEFD8C0AFD81EB63" httpUri="http://treatment.plazi.org/id/DB6B8792FFB0710FFEFD8C0AFD81EB63" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22">
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<paragraph id="537D3684FFB0710DFEFD8C0AFD26EE46" blockId="21.[264,680,1561,1622]" box="[264,680,1561,1587]" pageId="21" pageNumber="22">
<heading id="083581E8FFB0710DFEFD8C0AFD26EE46" bold="true" box="[264,680,1561,1587]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<emphasis id="61B6EA96FFB0710DFEFD8C0AFD26EE46" bold="true" box="[264,680,1561,1587]" pageId="21" pageNumber="22">
<taxonomicName id="94C24D07FFB0710DFEFD8C0AFDB3EE46" authority="Lelej" authorityName="Lelej" box="[264,573,1561,1587]" class="Insecta" family="Mutillidae" genus="Dentilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="ehrenbergi" status="sp. nov.">
<emphasis id="61B6EA96FFB0710DFEFD8C0AFE7BEE46" bold="true" box="[264,501,1561,1587]" italics="true" pageId="21" pageNumber="22">Dentilla ehrenbergi</emphasis>
Lelej
</taxonomicName>
,
<taxonomicNameLabel id="7A8557EDFFB0710DFDBE8C0AFD26EE46" box="[587,680,1561,1587]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="537D3684FFB0710DFEFD8C2FFEE3EE23" blockId="21.[264,680,1561,1622]" box="[264,365,1596,1622]" pageId="21" pageNumber="22">
(
<figureCitation id="CBF92A01FFB0710DFEE48C2FFEEDEE23" box="[273,355,1596,1622]" captionStart="FIGURES 11 – 16. 11" captionStartId="26.[264,383,923,947]" captionTargetBox="[549,1042,566,897]" captionTargetId="figure@26.[538,1048,566,898]" captionTargetPageId="26" captionText="FIGURES 11 – 16. 11. Dentilla ehrenbergi sp. nov., male, holotype. Genitalia, left part, inner view. 12 – 16. Paglianotilla hogenesi sp. nov., male, paratype. 12. Metasoma, dorsal view; 13. Pedicel and flagellomeres 1 – 3; 14. Mandible, dorsal view; 15. Genitalia, ventral view; 16. Penial valves, ventral view. Scale = 1 mm for 12 and 0.5 mm for 11, 13, and 15." httpUri="https://zenodo.org/record/172674/files/figure.png" pageId="21" pageNumber="22">Fig. 11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB0710DFEFD8C9FFBA6EEB6" pageId="21" pageNumber="22" type="reference_group">
<paragraph id="537D3684FFB0710DFEFD8C9FFBA6EEB6" blockId="21.[264,1087,1675,1731]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="73D211AAFFB0710DFEFD8C9FFBA6EEB6" pageId="21" pageNumber="22">
<taxonomicName id="94C24D07FFB0710DFEFD8C9FFC80EED6" authority="Bischoff 1920: 151" authorityName="Bischoff" authorityPageNumber="151" authorityYear="1920" box="[264,782,1676,1699]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subSpecies" species="continua" subSpecies="aurea">
<emphasis id="61B6EA96FFB0710DFEFD8C9FFDBEEED6" box="[264,560,1676,1699]" italics="true" pageId="21" pageNumber="22">Ephutomma continua aurea</emphasis>
: Bischoff 1920: 151
</taxonomicName>
, ♂ non Ψ. Misidentification.
<taxonomicName id="94C24D07FFB0710DFEFD8CBFFD2DEEB6" authority="Invrea 1965b: 65" authorityName="Invrea" authorityPageNumber="65" authorityYear="1965" box="[264,675,1708,1731]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="61B6EA96FFB0710DFEFD8CBFFE52EEB6" box="[264,476,1708,1731]" italics="true" pageId="21" pageNumber="22">Ephutomma bicolor</emphasis>
:
<treatmentCitation id="D2631095FFB0710DFE1E8CBFFD2DEEB6" author="Invrea" box="[491,675,1708,1731]" page="65" pageId="21" pageNumber="22" year="1965">
<bibRefCitation id="37534B75FFB0710DFE1E8CBFFDF2EEB6" author="Invrea" box="[491,636,1708,1731]" pageId="21" pageNumber="48" refString="Invrea, F. (1965 b) Missione 1962 del Prof. Giuseppe Scortecci nell'Arabia meridionale (Hymenoptera: Apterogynidae, Mutillidae et Chrysididae). Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano, 104 (1), 59 - 66." type="journal article" year="1965" yearSuffix="b">Invrea 1965b</bibRefCitation>
: 65
</treatmentCitation>
</taxonomicName>
, ♂ (Hadhramaut). Misidentification.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB0710EFEFD8CE7FDF8E9DB" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="537D3684FFB0710DFEFD8CE7FEF1EF78" blockId="21.[264,1323,1780,1886]" box="[264,383,1780,1805]" pageId="21" pageNumber="22">
<emphasis id="61B6EA96FFB0710DFEFD8CE7FEF1EF78" box="[264,383,1780,1805]" italics="true" pageId="21" pageNumber="22">Diagnosis</emphasis>
</paragraph>
<paragraph id="537D3684FFB0710EFECD8D0FFDF8E9DB" blockId="21.[264,1323,1780,1886]" lastBlockId="22.[264,1323,284,430]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
The male of this new species can be distinguished from that of the Palaearctic
<taxonomicName id="94C24D07FFB0710DFB3F8D0FFD64EF2B" authority="Sichel et Radoszkowski 1870" authorityName="Sichel et Radoszkowski" authorityYear="1870" class="Insecta" family="Mutillidae" genus="Dentilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="61B6EA96FFB0710DFB3F8D0FFEECEF28" italics="true" pageId="21" pageNumber="22">Dentilla dichroa</emphasis>
(
<bibRefCitation id="37534B75FFB0710DFE838D57FD6CEF2B" author="Sichel" box="[374,738,1860,1886]" pageId="21" pageNumber="50" refString="Sichel, J. &amp; Radoszkowski, O. (1869 - 1870) Essai d'une monographie des Mutilles de l'ancien continent. Horae Societatis Entomologicae Rossicae, (1869) 6 (3), 139 - 172; (1870) 6 (4), 173 - 309, tab. 6 - 11." type="journal article" year="1870">Sichel et Radoszkowski 1870</bibRefCitation>
)
</taxonomicName>
in having the apex of T7 rounded (straight or emarginated in
<taxonomicName id="94C24D07FFB3710EFE358B0FFD94E940" box="[448,538,284,309]" class="Insecta" family="Mutillidae" genus="Dentilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="61B6EA96FFB3710EFE358B0FFD94E940" box="[448,538,284,309]" italics="true" pageId="22" pageNumber="23">dichroa</emphasis>
</taxonomicName>
), in having golden felt lines on T2 and S2 (whitish in
<taxonomicName id="94C24D07FFB3710EFB558B0FFB74E940" box="[1184,1274,284,309]" class="Insecta" family="Mutillidae" genus="Dentilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="61B6EA96FFB3710EFB558B0FFB74E940" box="[1184,1274,284,309]" italics="true" pageId="22" pageNumber="23">dichroa</emphasis>
</taxonomicName>
), in having a more infuscated forewing (infuscated spot beyond the marginal cell in
<taxonomicName id="94C24D07FFB3710EFB4A8B57FA97E928" box="[1215,1305,324,349]" class="Insecta" family="Mutillidae" genus="Dentilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="61B6EA96FFB3710EFB4A8B57FA97E928" box="[1215,1305,324,349]" italics="true" pageId="22" pageNumber="23">dichroa</emphasis>
</taxonomicName>
), in having differring shape and setae of volsella (long setae of basivolsella project over apex of gonostylus in
<taxonomicName id="94C24D07FFB3710EFDFE8B87FDEBE9D8" box="[523,613,404,429]" class="Insecta" family="Mutillidae" genus="Dentilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="61B6EA96FFB3710EFDFE8B87FDEBE9D8" box="[523,613,404,429]" italics="true" pageId="22" pageNumber="23">dichroa</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB3710EFEFD8BF7FDB9EE2B" pageId="22" pageNumber="23" type="description">
<paragraph id="537D3684FFB3710EFEFD8BF7FE1CE988" blockId="22.[264,1326,484,1630]" box="[264,402,484,509]" pageId="22" pageNumber="23">
<emphasis id="61B6EA96FFB3710EFEFD8BF7FE1CE988" box="[264,402,484,509]" italics="true" pageId="22" pageNumber="23">Description</emphasis>
</paragraph>
<paragraph id="537D3684FFB3710EFECD881FFC9EED33" blockId="22.[264,1326,484,1630]" pageId="22" pageNumber="23">
MALE. Body length 8.0–
<quantity id="943A9B61FFB3710EFD9E881FFD56EA53" box="[619,728,524,550]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="22" pageNumber="23" unit="mm" value="12.8">12.8 mm</quantity>
. Head width 0.9X thorax width including tegulae. Clypeus deeply concave with narrow median process anteriorly and short acute basal median carina, clypeal surface sparsely finely punctate. Scape distinctly bicarinate beneath, upper ridge visible basally. Ocelli large, POD: OOD 0.7X; POD equal to maximal diameter of posterior ocellus; distance between posterior ocellus and posterior head margin 2.0X POD. Frons with deep longitudinal median furrow. First flagellomere length 1.2–1.3X its width, 2.4–2.7X pedicel, and 0.55–0.6X flagellomere 2, the latter 1.1–1.2X flagellomere 3. Antennal sockets with arcuate carina. Mandible quadridentate with strong curved upper carina, with large tooth beneath near the base, all inner teeth more or less equal, subbasal one with hair tuft ventrally. Vertex sparsely punctate, genae densely punctate. Pronotal dorsum almost straight anteriorly, pronotal width 1.2–1.3X propodeal width (on spiracle line). Scutum with well developed parascutal carinae. Metasternal process with denticles. Posterior coxae carinate inside. Tegulae slightly projecting over mesoscuto­scutellar suture, shining, glabrous, with a few punctures inside. Propodeum abrupt, reticulate with median dorsal longitudinal cell distinctly delimited posteriorly by two denticles. Relative length of
<emphasis id="61B6EA96FFB3710EFCBB8E77FCEEEC08" box="[846,864,1124,1149]" italics="true" pageId="22" pageNumber="23">R</emphasis>
­abscissas is 1.3(1.0): 1.0: 1.4(1.1): 2.0(1.4). Metasomal segment 1 carinate beneath, its length 1.0–1.1X its maximal width; segment 2 with long lateral felt lines on T2 and short ones on S2, length of felt line on S2 approximately equal to distance between felt line and posterior border of S2; T2 with dense large punctures, sparser on disc. T7 weakly rounded apically, with large dense punctures. Volsella of genitalia as in
<figureCitation id="CBF92A01FFB3710EFD428F3FFC82ED33" box="[695,780,1324,1350]" captionStart="FIGURES 11 – 16. 11" captionStartId="26.[264,383,923,947]" captionTargetBox="[549,1042,566,897]" captionTargetId="figure@26.[538,1048,566,898]" captionTargetPageId="26" captionText="FIGURES 11 – 16. 11. Dentilla ehrenbergi sp. nov., male, holotype. Genitalia, left part, inner view. 12 – 16. Paglianotilla hogenesi sp. nov., male, paratype. 12. Metasoma, dorsal view; 13. Pedicel and flagellomeres 1 – 3; 14. Mandible, dorsal view; 15. Genitalia, ventral view; 16. Penial valves, ventral view. Scale = 1 mm for 12 and 0.5 mm for 11, 13, and 15." httpUri="https://zenodo.org/record/172674/files/figure.png" pageId="22" pageNumber="23">Fig. 11</figureCitation>
.
</paragraph>
<paragraph id="537D3684FFB3710EFECD8F47FB40EE43" blockId="22.[264,1326,484,1630]" pageId="22" pageNumber="23">Head, palps, mesosoma, tegulae, legs, T1 basally, and S1 yellowish­red. Antenna yellowish­red with brown apex. Mandible yellowish­red with dark denticles. Metasoma black. Mid­ and hind tibial spurs whitish. Wings hyaline with brown veins, forewing (distal of cells) distinctly infuscated, darker toward anterior margin. Body and legs clothed with recumbent short and scattered long erect whitish pubescence; T2–T6 and S2–S6 with apical fringes of whitish setae which denser on T2; felt lines on T2 and S2 golden.</paragraph>
<paragraph id="537D3684FFB3710EFECD8C57FDB9EE2B" blockId="22.[264,1326,484,1630]" box="[312,567,1604,1630]" pageId="22" pageNumber="23">FEMALE. Unknown.</paragraph>
</subSubSection>
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<typeStatus id="8C798826FFB3710EFEFD8C87FEB1EED8" box="[264,319,1684,1709]" pageId="22" pageNumber="23">Type</typeStatus>
material
</emphasis>
</paragraph>
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<typeStatus id="8C798826FFB3710EFECD8CAFFE2BEEA3" box="[312,421,1724,1750]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
ɗ,
<collectingCountry id="2BD57614FFB3710EFE3B8CAFFDADEEA3" box="[462,547,1724,1750]" name="Yemen" pageId="22" pageNumber="23">Yemen</collectingCountry>
, Al Kowd,
<date id="277C1044FFB3710EFD5C8CAFFC85EEA3" box="[681,779,1724,1750]" pageId="22" pageNumber="23" value="2001-03">III.2001</date>
, LT, AvH &amp; S. Al Haruri [
<collectionCode id="35D3AE41FFB3710EFBB58CAFFB2EEEA3" LSID="urn:lsid:biocol.org:col:34975" box="[1088,1184,1724,1750]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="22" pageNumber="23">ZMAN</collectionCode>
].
<typeStatus id="8C798826FFB3710EFB448CAFFAA9EEA3" box="[1201,1319,1724,1750]" pageId="22" pageNumber="23" type="paratype">Paratypes</typeStatus>
. Al Kowd,
<date id="277C1044FFB3710EFE728CF7FE65EE8B" box="[391,491,1764,1790]" pageId="22" pageNumber="23" value="1999-09">IX.1999</date>
, LT, AvH &amp; S. Al Haruri, 1 ɗ [
<collectionCode id="35D3AE41FFB3710EFC9E8CF7FC3FEE8B" box="[875,945,1764,1790]" pageId="22" pageNumber="23">IBSS</collectionCode>
];
<date id="277C1044FFB3710EFC338CF7FBC3EE8B" box="[966,1101,1764,1790]" pageId="22" pageNumber="23" value="2000-05" valueMax="2000-06-31" valueMin="2000-05-00">V–VI.2000</date>
, LT, AvH &amp; S. Al Haruri, 4 ɗ [
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,
<collectionCode id="35D3AE41FFB3710EFDF48D1FFDCAEF53" box="[513,580,1804,1830]" pageId="22" pageNumber="23">IBSS</collectionCode>
];
<date id="277C1044FFB3710EFDAD8D1FFD4BEF53" box="[600,709,1804,1830]" pageId="22" pageNumber="23" value="2000-07">VII.2000</date>
, LT, AvH &amp; S. Al Haruri, 1 ɗ [
<collectionCode id="35D3AE41FFB3710EFBD88D1FFB02EF53" LSID="urn:lsid:biocol.org:col:34975" box="[1069,1164,1804,1830]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="22" pageNumber="23">ZMAN</collectionCode>
];
<date id="277C1044FFB3710EFB6A8D1FFB77EF53" box="[1183,1273,1804,1830]" pageId="22" pageNumber="23" value="2000-10">X.2000</date>
, LT, AvH &amp; S. Al Haruri, 1 ɗ [
<collectionCode id="35D3AE41FFB3710EFD968D27FD4EEF3B" LSID="urn:lsid:biocol.org:col:34975" box="[611,704,1844,1870]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="22" pageNumber="23">ZMAN</collectionCode>
;
<date id="277C1044FFB3710EFD248D27FC91EF3B" box="[721,799,1844,1870]" pageId="22" pageNumber="23" value="2001-01">I.2001</date>
, LT, AvH &amp; S. Al Haruri, 1 ɗ [
<collectionCode id="35D3AE41FFB3710EFB358D27FA91EF3B" LSID="urn:lsid:biocol.org:col:34975" box="[1216,1311,1844,1870]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="22" pageNumber="23">ZMAN</collectionCode>
];
<date id="277C1044FFB2710FFEFD8B0FFEE5E943" box="[264,363,284,310]" pageId="23" pageNumber="24" value="2003-09">IX.2003</date>
, LT, AvH &amp; S. Al Haruri, 3 ɗ [
<collectionCode id="35D3AE41FFB2710FFC9C8B0FFC44E943" LSID="urn:lsid:biocol.org:col:34992" box="[873,970,284,310]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="24">RMNH</collectionCode>
];
<quantity id="943A9B61FFB2710FFC188B0FFBC6E943" box="[1005,1096,284,310]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="23" pageNumber="24" unit="km" value="12.0">12 km</quantity>
NW Manakhah,
<date id="277C1044FFB2710FFEFD8B57FE7EE92B" box="[264,496,324,350]" pageId="23" pageNumber="24" value="2001-07-03" valueMax="2001-08-21" valueMin="2001-07-03">3.VII–21.VIII.2001</date>
, MT, AvH, 1 ɗ [
<collectionCode id="35D3AE41FFB2710FFD348B57FC89E92B" box="[705,775,324,350]" pageId="23" pageNumber="24">IBSS</collectionCode>
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<date id="277C1044FFB2710FFCE88B57FC60E92B" box="[797,1006,324,350]" pageId="23" pageNumber="24" value="2003-09-15" valueMax="2003-10-22" valueMin="2003-09-15">15.IX–22.X.2003</date>
, MT, AvH, 1 ɗ [
<collectionCode id="35D3AE41FFB2710FFB4B8B57FA91E92B" LSID="urn:lsid:biocol.org:col:34992" box="[1214,1311,324,350]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="24">RMNH</collectionCode>
]; Jabal Bura,
<date id="277C1044FFB2710FFE698B7FFDABE9F3" box="[412,549,364,390]" pageId="23" pageNumber="24" value="1988-09-25">25.IX.1988</date>
, LT, J. Meerman, 1 ɗ [
<collectionCode id="35D3AE41FFB2710FFCB88B7FFC1CE9F3" box="[845,914,364,390]" pageId="23" pageNumber="24">IBSS</collectionCode>
]; Al Hudaydah Tal,
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E
<collectingCountry id="2BD57614FFB2710FFAFF8B7FFAA5E9F3" box="[1290,1323,364,390]" name="American Samoa" pageId="23" pageNumber="24">As</collectingCountry>
Sukhnah,
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, 500 NN Jem­Suk,
<date id="277C1044FFB2710FFCCF8B87FC31E9DB" box="[826,959,404,430]" pageId="23" pageNumber="24" value="1997-04-12">12.IV.1997</date>
, lif Brechtel, Würst, Ehrmann, 1 ɗ [
<collectionCode id="35D3AE41FFB2710FFEB58BAFFE08E9A3" box="[320,390,444,470]" pageId="23" pageNumber="24">IBSS</collectionCode>
].
<collectingCountry id="2BD57614FFB2710FFE6D8BAFFDBBE9A3" box="[408,565,444,470]" name="Saudi Arabia" pageId="23" pageNumber="24">Saudi Arabia</collectingCountry>
, Al Hijaz (Hejaz), Jeddah,
<date id="277C1044FFB2710FFC858BAFFC65E9A3" box="[880,1003,444,470]" pageId="23" pageNumber="24" value="1931-05-28">28.V.1931</date>
, Zhenzhurist (N. Filippov), 1 ɗ [
<collectionCode id="35D3AE41FFB2710FFEB58BF7FE2BE98B" box="[320,421,484,510]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="23" pageNumber="24">ZMMU</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB2710FFEFD8827FDB4EA03" box="[264,570,564,630]" pageId="23" pageNumber="24" type="distribution">
<paragraph id="537D3684FFB2710FFEFD8827FE19EA38" blockId="23.[264,570,564,630]" box="[264,407,564,589]" pageId="23" pageNumber="24">
<emphasis id="61B6EA96FFB2710FFEFD8827FE19EA38" box="[264,407,564,589]" italics="true" pageId="23" pageNumber="24">Distribution</emphasis>
</paragraph>
<paragraph id="537D3684FFB2710FFECD884FFDB4EA03" blockId="23.[264,570,564,630]" box="[312,570,604,630]" pageId="23" pageNumber="24">
<collectingCountry id="2BD57614FFB2710FFECD884FFE03EA03" box="[312,397,604,630]" name="Yemen" pageId="23" pageNumber="24">Yemen</collectingCountry>
,
<collectingCountry id="2BD57614FFB2710FFE6D884FFDB8EA03" box="[408,566,604,630]" name="Saudi Arabia" pageId="23" pageNumber="24">Saudi Arabia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB2710FFEFD88BFFD81EB63" pageId="23" pageNumber="24" type="etymology">
<paragraph id="537D3684FFB2710FFEFD88BFFE0BEAB0" blockId="23.[264,1323,684,790]" box="[264,389,684,709]" pageId="23" pageNumber="24">
<emphasis id="61B6EA96FFB2710FFEFD88BFFE0BEAB0" box="[264,389,684,709]" italics="true" pageId="23" pageNumber="24">Etymology</emphasis>
</paragraph>
<paragraph id="537D3684FFB2710FFECD88C7FD81EB63" blockId="23.[264,1323,684,790]" pageId="23" pageNumber="24">
This species is dedicated to Christian Gottfried Ehrenberg, who was the first collector of mutillids in
<collectingCountry id="2BD57614FFB2710FFE4088EFFD85EB63" box="[437,523,764,790]" name="Yemen" pageId="23" pageNumber="24">Yemen</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

126
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*
<taxonomicName id="94C24D07FFB2710FFEE28967FDF4EBFB" authority="Klug 1829" authorityName="Klug" authorityYear="1829" box="[279,634,884,910]" class="Insecta" family="Mutillidae" genus="Dentilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="61B6EA96FFB2710FFEE28967FE6EEBFB" bold="true" box="[279,480,884,910]" italics="true" pageId="23" pageNumber="24">Dentilla testacea</emphasis>
(
<bibRefCitation id="37534B75FFB2710FFE1A8967FDFCEBFB" author="Klug" box="[495,626,884,910]" pageId="23" pageNumber="48" refString="Klug, J. C. F. (1829) Symbolae physicae seu icones et descriptiones insectorum quae ex itinere per Africam borealem et Asiam occidentalem Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg studio novae aut illustratae redierunt. Hymenoptera. Decas Prima [genera Mutilla et Apterogyna]. Venditur a Mittlero, Berolini. [15 - 21] + tab. 4, 5." type="journal volume" year="1829">Klug 1829</bibRefCitation>
)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<paragraph id="537D3684FFB2710FFEFD89ACFDB8EB80" blockId="23.[264,1323,958,1078]" pageId="23" pageNumber="24">
<taxonomicName id="94C24D07FFB2710FFEFD89ACFDB6EBA0" authority="Klug 1829" authorityName="Klug" authorityYear="1829" box="[264,568,958,982]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="61B6EA96FFB2710FFEFD89ACFE39EBA3" box="[264,439,959,982]" italics="true" pageId="23" pageNumber="24">Mutilla testacea</emphasis>
<bibRefCitation id="37534B75FFB2710FFE3489ADFDB6EBA0" author="Klug" box="[449,568,958,981]" pageId="23" pageNumber="48" refString="Klug, J. C. F. (1829) Symbolae physicae seu icones et descriptiones insectorum quae ex itinere per Africam borealem et Asiam occidentalem Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg studio novae aut illustratae redierunt. Hymenoptera. Decas Prima [genera Mutilla et Apterogyna]. Venditur a Mittlero, Berolini. [15 - 21] + tab. 4, 5." type="journal volume" year="1829">Klug 1829</bibRefCitation>
</taxonomicName>
: [20], tab. 5, fig. 4, ♂,
<typeStatus id="8C798826FFB2710FFCC989ADFC17EBA0" box="[828,921,958,981]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
— &quot;Ex Arabia deserta&quot; (central and north Arabia) [ZMHU].
</paragraph>
<paragraph id="537D3684FFB2710FFEFD89ECFD67EC60" blockId="23.[264,1323,958,1078]" box="[264,745,1022,1046]" pageId="23" pageNumber="24">
<taxonomicName id="94C24D07FFB2710FFEFD89ECFD4AEC60" authority="Bischoff 1920: 154" authorityName="Bischoff" authorityPageNumber="154" authorityYear="1920" box="[264,708,1022,1046]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="61B6EA96FFB2710FFEFD89ECFE69EC63" box="[264,487,1023,1046]" italics="true" pageId="23" pageNumber="24">Ephutomma testacea</emphasis>
: Bischoff 1920: 154
</taxonomicName>
, ♂.
</paragraph>
<paragraph id="537D3684FFB2710FFEFD8E0CFDE0EC40" blockId="23.[264,1323,958,1078]" box="[264,622,1054,1078]" pageId="23" pageNumber="24">
<treatmentCitationGroup id="73D211AAFFB2710FFEFD8E0CFDE0EC40" box="[264,622,1054,1078]" pageId="23" pageNumber="24">
<taxonomicName id="94C24D07FFB2710FFEFD8E0CFDE5EC40" authority="Lelej 2002: 52" authorityName="Lelej" authorityPageNumber="52" authorityYear="2002" box="[264,619,1054,1078]" class="Insecta" family="Mutillidae" genus="Dentilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="61B6EA96FFB2710FFEFD8E0CFE30EC43" box="[264,446,1055,1078]" italics="true" pageId="23" pageNumber="24">Dentilla testacea</emphasis>
:
<treatmentCitation id="D2631095FFB2710FFE3E8E0DFDE5EC40" author="Lelej" box="[459,619,1054,1077]" page="52" pageId="23" pageNumber="24" year="2002">
<bibRefCitation id="37534B75FFB2710FFE3E8E0DFDCDEC40" author="Lelej" box="[459,579,1054,1077]" pageId="23" pageNumber="48" refString="Lelej, A. S. (2002) Catalogue of the Mutillidae (Hymenoptera) of the Palaearctic region. Dalnauka, Vladivostok. 171 pp." type="book" year="2002">Lelej 2002</bibRefCitation>
: 52
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB2710FFEFD8E74FCFDEC8C" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="537D3684FFB2710FFEFD8E74FE68ECF5" blockId="23.[264,1323,1127,1273]" box="[264,486,1127,1152]" pageId="23" pageNumber="24">
<emphasis id="61B6EA96FFB2710FFEFD8E74FE68ECF5" box="[264,486,1127,1152]" italics="true" pageId="23" pageNumber="24">Material examined</emphasis>
</paragraph>
<paragraph id="537D3684FFB2710FFECD8E9CFCFDEC8C" blockId="23.[264,1323,1127,1273]" pageId="23" pageNumber="24">
<collectingCountry id="2BD57614FFB2710FFECD8E9CFE03ECDC" box="[312,397,1167,1193]" name="Yemen" pageId="23" pageNumber="24">Yemen</collectingCountry>
, Al Kowd,
<date id="277C1044FFB2710FFDE68E9CFDE5ECDC" box="[531,619,1167,1193]" pageId="23" pageNumber="24" value="2001-02">II.2001</date>
, LT, AvH &amp; S. Al Haruri, 1 ɗ [
<collectionCode id="35D3AE41FFB2710FFC228E9CFBB9ECDC" LSID="urn:lsid:biocol.org:col:34975" box="[983,1079,1167,1193]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="23" pageNumber="24">ZMAN</collectionCode>
];
<date id="277C1044FFB2710FFBBC8E9CFB76ECDC" box="[1097,1272,1167,1193]" pageId="23" pageNumber="24" value="2001-07-08" valueMax="2001-07-12" valueMin="2001-07-08">8–12.VII.2001</date>
, LT, AvH &amp; S. Al Haruri, 2 ɗ [
<collectionCode id="35D3AE41FFB2710FFD9C8EA4FD49ECA4" LSID="urn:lsid:biocol.org:col:34992" box="[617,711,1207,1233]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="24">RMNH</collectionCode>
,
<collectionCode id="35D3AE41FFB2710FFD2F8EA4FC90ECA4" box="[730,798,1207,1233]" pageId="23" pageNumber="24">IBSS</collectionCode>
].
<collectingCountry id="2BD57614FFB2710FFCC28EA4FC54ECA4" box="[823,986,1207,1233]" name="Saudi Arabia" pageId="23" pageNumber="24">Saudi Arabia</collectingCountry>
, Al Hijaz (Hejaz), Jeddah,
<date id="277C1044FFB2710FFEFD8ECCFE0DEC8C" box="[264,387,1247,1273]" pageId="23" pageNumber="24" value="1931-05-28">28.V.1931</date>
, Zhenzhurist (N. Filippov), 1 ɗ [
<collectionCode id="35D3AE41FFB2710FFCF68ECCFCE6EC8C" box="[771,872,1247,1273]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="23" pageNumber="24">ZMMU</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB2710FFEFD8F3CFEDAEDEC" pageId="23" pageNumber="24" type="distribution">
<paragraph id="537D3684FFB2710FFEFD8F3CFE19ED3D" blockId="23.[264,1323,1327,1433]" box="[264,407,1327,1352]" pageId="23" pageNumber="24">
<emphasis id="61B6EA96FFB2710FFEFD8F3CFE19ED3D" box="[264,407,1327,1352]" italics="true" pageId="23" pageNumber="24">Distribution</emphasis>
</paragraph>
<paragraph id="537D3684FFB2710FFECD8F44FEDAEDEC" blockId="23.[264,1323,1327,1433]" pageId="23" pageNumber="24">
<collectingCountry id="2BD57614FFB2710FFECD8F44FE04ED04" box="[312,394,1367,1393]" name="Yemen" pageId="23" pageNumber="24">Yemen</collectingCountry>
(new record),
<collectingCountry id="2BD57614FFB2710FFDB08F44FD6AED04" box="[581,740,1367,1393]" name="Saudi Arabia" pageId="23" pageNumber="24">Saudi Arabia</collectingCountry>
(new record), North and Central Arabia (
<bibRefCitation id="37534B75FFB2710FFB1A8F44FEC9EDEC" author="Klug" pageId="23" pageNumber="48" refString="Klug, J. C. F. (1829) Symbolae physicae seu icones et descriptiones insectorum quae ex itinere per Africam borealem et Asiam occidentalem Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg studio novae aut illustratae redierunt. Hymenoptera. Decas Prima [genera Mutilla et Apterogyna]. Venditur a Mittlero, Berolini. [15 - 21] + tab. 4, 5." type="journal volume" year="1829">Klug 1829</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

179
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<heading id="083581E8FFB47109FEFD8BF7FD3DE988" bold="true" box="[264,691,483,510]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<emphasis id="61B6EA96FFB47109FEFD8BF7FD3DE988" bold="true" box="[264,691,483,510]" pageId="17" pageNumber="18">
<emphasis id="61B6EA96FFB47109FEFD8BF7FE9AE98B" bold="true" box="[264,276,484,510]" italics="true" pageId="17" pageNumber="18">*</emphasis>
<taxonomicName id="94C24D07FFB47109FEE18BF7FD3DE988" authority="(Magretti 1906)" authorityName="Magretti" authorityYear="1906" baseAuthorityName="Magretti" baseAuthorityYear="1906" box="[276,691,483,510]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bellii">
<emphasis id="61B6EA96FFB47109FEE18BF7FE66E98B" bold="true" box="[276,488,484,510]" italics="true" pageId="17" pageNumber="18">Tropidotilla bellii</emphasis>
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<taxonomicName id="94C24D07FFB47109FEFD883CFDD0EA30" authority="Magretti 1906: 68" authorityName="Magretti" authorityPageNumber="68" authorityYear="1906" box="[264,606,558,582]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bellii">
<emphasis id="61B6EA96FFB47109FEFD883CFE1AEA33" box="[264,404,559,582]" italics="true" pageId="17" pageNumber="18">Mutilla bellii</emphasis>
Magretti 1906: 68
</taxonomicName>
, fig. 10, ♂,
<typeStatus id="8C798826FFB47109FD29883DFCB7EA30" box="[732,825,558,581]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
&quot;Saati&quot; [
<collectingCountry id="2BD57614FFB47109FC6C883DFC69EA30" box="[921,999,558,581]" name="Eritrea" pageId="17" pageNumber="18">Eritrea</collectingCountry>
, South of Massawa],
<date id="277C1044FFB47109FB38883DFAA9EA30" box="[1229,1319,558,581]" pageId="17" pageNumber="18" value="1901-03">III.1901</date>
, leg. V. Belli [MSNG].
</paragraph>
<paragraph id="537D3684FFB47109FEFD887CFD4AEAF0" blockId="17.[264,1322,558,646]" box="[264,708,622,646]" pageId="17" pageNumber="18">
<taxonomicName id="94C24D07FFB47109FEFD887CFD2FEAF0" authority="Bischoff 1920: 157" authorityName="Bischoff" authorityPageNumber="157" authorityYear="1920" box="[264,673,622,646]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bellii">
<emphasis id="61B6EA96FFB47109FEFD887CFE4AEAF3" box="[264,452,623,646]" italics="true" pageId="17" pageNumber="18">Tropidotilla bellii</emphasis>
: Bischoff 1920: 157
</taxonomicName>
, ♂.
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB47109FEFD88CCFBD0ECFC" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="537D3684FFB47109FEFD88CCFE68EA8D" blockId="17.[264,1323,735,1161]" box="[264,486,735,760]" pageId="17" pageNumber="18">
<emphasis id="61B6EA96FFB47109FEFD88CCFE68EA8D" box="[264,486,735,760]" italics="true" pageId="17" pageNumber="18">Material examined</emphasis>
</paragraph>
<paragraph id="537D3684FFB47109FECD8914FBD0ECFC" blockId="17.[264,1323,735,1161]" pageId="17" pageNumber="18">
<collectingCountry id="2BD57614FFB47109FECD8914FE00EB54" box="[312,398,775,801]" name="Yemen" pageId="17" pageNumber="18">Yemen</collectingCountry>
: Al Kowd,
<date id="277C1044FFB47109FDEF8914FD52EB54" box="[538,732,775,801]" pageId="17" pageNumber="18" value="1992-10-27" valueMax="1992-11-06" valueMin="1992-10-27">27.X–6.XI.1992</date>
, MT, AvH 1 ɗ [
<collectionCode id="35D3AE41FFB47109FC578914FB8CEB54" LSID="urn:lsid:biocol.org:col:34975" box="[930,1026,775,801]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="17" pageNumber="18">ZMAN</collectionCode>
]; Al Kadan,
<date id="277C1044FFB47109FB6E8914FECBEB3C" pageId="17" pageNumber="18" value="1998-02-17" valueMax="1998-03-31" valueMin="1998-02-17">17.II–31.III. 1998</date>
, MT, AvH &amp; H.M. Naser, 2 ɗ [
<collectionCode id="35D3AE41FFB47109FD31893CFCADEB3C" LSID="urn:lsid:biocol.org:col:34975" box="[708,803,815,841]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="17" pageNumber="18">ZMAN</collectionCode>
];
<date id="277C1044FFB47109FCCD893CFC34EB3C" box="[824,954,815,841]" pageId="17" pageNumber="18" value="1997-12-03">3.XII.1997</date>
–
<date id="277C1044FFB47109FC3E893CFBCAEB3C" box="[971,1092,815,841]" pageId="17" pageNumber="18" value="1998-02-17">17.II.1998</date>
, MT, AvH &amp; H.M. Naser, 1 ɗ [
<collectionCode id="35D3AE41FFB47109FE6D8944FE79EB04" LSID="urn:lsid:biocol.org:col:34975" box="[408,503,855,881]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="17" pageNumber="18">ZMAN</collectionCode>
]; Lahj,
<date id="277C1044FFB47109FDA78944FD63EB04" box="[594,749,855,881]" pageId="17" pageNumber="18" value="1999-11" valueMax="1999-12-31" valueMin="1999-11-00">XI–XII.1999</date>
, MT, AvH &amp; A. Sallam, 1 ɗ [
<collectionCode id="35D3AE41FFB47109FBAC8944FB37EB04" LSID="urn:lsid:biocol.org:col:34975" box="[1113,1209,855,881]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="17" pageNumber="18">ZMAN</collectionCode>
];
<date id="277C1044FFB47109FB3B8944FAA8EB04" box="[1230,1318,855,881]" pageId="17" pageNumber="18" value="2000-02">II.2000</date>
, MT, AvH &amp; A. Sallam, 1 ɗ [
<collectionCode id="35D3AE41FFB47109FD82896CFD59EBEC" LSID="urn:lsid:biocol.org:col:34975" box="[631,727,895,921]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="17" pageNumber="18">ZMAN</collectionCode>
];
<date id="277C1044FFB47109FD18896CFC30EBEC" box="[749,958,895,921]" pageId="17" pageNumber="18" value="2000-05-17" valueMax="2000-06-15" valueMin="2000-05-17">17.V–15.VI.2000</date>
, MT, AvH &amp; A. Sallam, 4 ɗ [
<collectionCode id="35D3AE41FFB47109FEFA89B4FEFEEBB4" LSID="urn:lsid:biocol.org:col:34992" box="[271,368,935,961]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="17" pageNumber="18">RMNH</collectionCode>
];
<date id="277C1044FFB47109FE7789B4FDB8EBB4" box="[386,566,935,961]" pageId="17" pageNumber="18" value="2000-06-15" valueMax="2000-06-30" valueMin="2000-06-15">15–30.VI.2000</date>
, MT, AvH &amp; A. Sallam, 8 ɗ [
<collectionCode id="35D3AE41FFB47109FC7A89B4FC7EEBB4" LSID="urn:lsid:biocol.org:col:34992" box="[911,1008,935,961]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="17" pageNumber="18">RMNH</collectionCode>
];
<date id="277C1044FFB47109FBF789B4FBFEEBB4" box="[1026,1136,935,961]" pageId="17" pageNumber="18" value="2000-07">VII.2000</date>
, MT, AvH &amp; A. Sallam, 3 ɗ [
<collectionCode id="35D3AE41FFB47109FE6A89DCFD8EEB9C" LSID="urn:lsid:biocol.org:col:34992" box="[415,512,975,1001]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="17" pageNumber="18">RMNH</collectionCode>
];
<date id="277C1044FFB47109FDE689DCFD07EB9C" box="[531,649,975,1001]" pageId="17" pageNumber="18" value="2000-08">VIII.2000</date>
, MT, AvH &amp; A. Sallam, 5 ɗ [
<collectionCode id="35D3AE41FFB47109FC1689DCFBCAEB9C" LSID="urn:lsid:biocol.org:col:34992" box="[995,1092,975,1001]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="17" pageNumber="18">RMNH</collectionCode>
];
<date id="277C1044FFB47109FBA389DCFB3FEB9C" box="[1110,1201,975,1001]" pageId="17" pageNumber="18" value="2000-10">X.2000</date>
, MT, AvH &amp; A. Sallam, 1 ɗ [
<collectionCode id="35D3AE41FFB47109FE1C89E4FDC4EC64" LSID="urn:lsid:biocol.org:col:34992" box="[489,586,1015,1041]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="17" pageNumber="18">RMNH</collectionCode>
];
<date id="277C1044FFB47109FDAB89E4FD4FEC64" box="[606,705,1015,1041]" pageId="17" pageNumber="18" value="2000-11">XI.2000</date>
, MT, AvH &amp; A. Sallam, 2 ɗ [
<collectionCode id="35D3AE41FFB47109FBDD89E4FB07EC64" LSID="urn:lsid:biocol.org:col:34992" box="[1064,1161,1015,1041]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="17" pageNumber="18">RMNH</collectionCode>
];
<date id="277C1044FFB47109FB6889E4FB62EC64" box="[1181,1260,1015,1041]" pageId="17" pageNumber="18" value="2001-01">I.2001</date>
, MT, AvH &amp; A. Sallam, 1 ɗ [
<collectionCode id="35D3AE41FFB47109FDB98E0CFD23EC4C" LSID="urn:lsid:biocol.org:col:34992" box="[588,685,1055,1081]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="17" pageNumber="18">RMNH</collectionCode>
];
<date id="277C1044FFB47109FD328E0CFCA9EC4C" box="[711,807,1055,1081]" pageId="17" pageNumber="18" value="2001-04">IV.2001</date>
, MT, AvH &amp; A. Sallam, 2 ɗ [
<collectionCode id="35D3AE41FFB47109FB4B8E0CFA91EC4C" LSID="urn:lsid:biocol.org:col:34992" box="[1214,1311,1055,1081]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="17" pageNumber="18">RMNH</collectionCode>
];
<date id="277C1044FFB47109FEFD8E54FE2DEC14" box="[264,419,1095,1121]" pageId="17" pageNumber="18" value="2001-07" valueMax="2001-09-31" valueMin="2001-07-00">VII–IX.2001</date>
, MT, AvH &amp; A. Sallam, 2 ɗ [
<collectionCode id="35D3AE41FFB47109FCD78E54FC0DEC14" LSID="urn:lsid:biocol.org:col:34992" box="[802,899,1095,1121]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="17" pageNumber="18">RMNH</collectionCode>
];
<date id="277C1044FFB47109FC6F8E54FBD5EC14" box="[922,1115,1095,1121]" pageId="17" pageNumber="18" value="2001-10-01" valueMax="2001-11-17" valueMin="2001-10-01">1.X–17.XI.2001</date>
, MT, AvH &amp; A. Sallam, 2 ɗ [
<collectionCode id="35D3AE41FFB47109FE558E7CFD8FECFC" LSID="urn:lsid:biocol.org:col:34992" box="[416,513,1135,1161]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="17" pageNumber="18">RMNH</collectionCode>
];
<date id="277C1044FFB47109FDE18E7CFD1BECFC" box="[532,661,1135,1161]" pageId="17" pageNumber="18" value="2002-03" valueMax="2002-05-31" valueMin="2002-03-00">III–V.2002</date>
, MT, AvH &amp; A. Sallam, 2 ɗ [
<collectionCode id="35D3AE41FFB47109FC078E7CFBDDECFC" LSID="urn:lsid:biocol.org:col:34992" box="[1010,1107,1135,1161]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="17" pageNumber="18">RMNH</collectionCode>
].
</paragraph>
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<subSubSection id="1BD8650FFFB47109FEFD8EACFCC1ED74" box="[264,847,1215,1281]" pageId="17" pageNumber="18" type="distribution">
<paragraph id="537D3684FFB47109FEFD8EACFE19ECAD" blockId="17.[264,847,1215,1281]" box="[264,407,1215,1240]" pageId="17" pageNumber="18">
<emphasis id="61B6EA96FFB47109FEFD8EACFE19ECAD" box="[264,407,1215,1240]" italics="true" pageId="17" pageNumber="18">Distribution</emphasis>
</paragraph>
<paragraph id="537D3684FFB47109FECD8EF4FCC1ED74" blockId="17.[264,847,1215,1281]" box="[312,847,1255,1281]" pageId="17" pageNumber="18">
<collectingCountry id="2BD57614FFB47109FECD8EF4FE04ED74" box="[312,394,1255,1281]" name="Yemen" pageId="17" pageNumber="18">Yemen</collectingCountry>
(new record),
<collectingCountry id="2BD57614FFB47109FDC38EF4FD08ED74" box="[566,646,1255,1281]" name="Eritrea" pageId="17" pageNumber="18">Eritrea</collectingCountry>
(Magretti 1906).
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<paragraph id="537D3684FFB47109FEFD8F24FEE0ED25" blockId="17.[264,1235,1335,1401]" box="[264,366,1335,1360]" pageId="17" pageNumber="18">
<emphasis id="61B6EA96FFB47109FEFD8F24FEE0ED25" box="[264,366,1335,1360]" italics="true" pageId="17" pageNumber="18">Remarks</emphasis>
</paragraph>
<paragraph id="537D3684FFB47109FECD8F4CFB5DED0C" blockId="17.[264,1235,1335,1401]" box="[312,1235,1375,1401]" pageId="17" pageNumber="18">
See
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<emphasis id="61B6EA96FFB47109FE9C8F4CFDD3ED0D" box="[361,605,1375,1400]" italics="true" pageId="17" pageNumber="18">Tropidotilla frontalis</emphasis>
</taxonomicName>
for the discussion of the taxonomic status of
<taxonomicName id="94C24D07FFB47109FB868F4CFB42ED0D" box="[1139,1228,1375,1400]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bellii">
<emphasis id="61B6EA96FFB47109FB868F4CFB42ED0D" box="[1139,1228,1375,1400]" italics="true" pageId="17" pageNumber="18">T. bellii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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110
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<heading id="083581E8FFB47109FEFD8FC5FD60ED85" bold="true" box="[264,750,1494,1520]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<emphasis id="61B6EA96FFB47109FEFD8FC5FD60ED85" bold="true" box="[264,750,1494,1520]" pageId="17" pageNumber="18">
<emphasis id="61B6EA96FFB47109FEFD8FC5FE9AED85" bold="true" box="[264,276,1494,1520]" italics="true" pageId="17" pageNumber="18">*</emphasis>
<taxonomicName id="94C24D07FFB47109FEE18FC5FD60ED85" authority="Bischoff 1920" authorityName="Bischoff" authorityYear="1920" box="[276,750,1494,1520]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="carinulifera">
<emphasis id="61B6EA96FFB47109FEE18FC5FDB3ED85" bold="true" box="[276,573,1494,1520]" italics="true" pageId="17" pageNumber="18">Tropidotilla carinulifera</emphasis>
Bischoff 1920
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<taxonomicName id="94C24D07FFB47109FEFD8C31FD6FEE4D" authority="Bischoff 1920: 158" authorityName="Bischoff" authorityPageNumber="158" authorityYear="1920" box="[264,737,1569,1593]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="carinulifera">
<emphasis id="61B6EA96FFB47109FEFD8C31FD82EE4C" box="[264,524,1570,1593]" italics="true" pageId="17" pageNumber="18">Tropidotilla carinulifera</emphasis>
Bischoff 1920: 158
</taxonomicName>
, ♂,
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&quot;Metemma&quot; [North
<collectingCountry id="2BD57614FFB47109FBB68C32FB2DEE4D" box="[1091,1187,1569,1592]" name="Ethiopia" pageId="17" pageNumber="18">Ethiopia</collectingCountry>
],
<date id="277C1044FFB47109FB468C32FAA4EE4D" box="[1203,1322,1569,1592]" pageId="17" pageNumber="18" value="1883-03-24">24.III.1883</date>
[MSNG].
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<paragraph id="537D3684FFB47109FEFD8C9AFE68EED7" blockId="17.[264,1190,1673,1739]" box="[264,486,1673,1698]" pageId="17" pageNumber="18">
<emphasis id="61B6EA96FFB47109FEFD8C9AFE68EED7" box="[264,486,1673,1698]" italics="true" pageId="17" pageNumber="18">Material examined</emphasis>
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<paragraph id="537D3684FFB47109FECD8CA2FB28EEBE" blockId="17.[264,1190,1673,1739]" box="[312,1190,1713,1739]" pageId="17" pageNumber="18">
<collectingCountry id="2BD57614FFB47109FECD8CA2FE00EEBE" box="[312,398,1713,1739]" name="Yemen" pageId="17" pageNumber="18">Yemen</collectingCountry>
: Lahj,
<date id="277C1044FFB47109FE288CA2FD1EEEBE" box="[477,656,1713,1739]" pageId="17" pageNumber="18" value="2000-06-15" valueMax="2000-06-30" valueMin="2000-06-15">15–30.VI.2000</date>
, MT, AvH &amp; A. Sallam, 3 ɗ [
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,
<collectionCode id="35D3AE41FFB47109FBA28CA2FB15EEBE" box="[1111,1179,1713,1739]" pageId="17" pageNumber="18">IBSS</collectionCode>
].
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<emphasis id="61B6EA96FFB47109FEFD8D12FE19EF6F" box="[264,407,1793,1818]" italics="true" pageId="17" pageNumber="18">Distribution</emphasis>
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<paragraph id="537D3684FFB47109FECD8D3AFC20EF36" blockId="17.[264,942,1793,1859]" box="[312,942,1833,1859]" pageId="17" pageNumber="18">
<collectingCountry id="2BD57614FFB47109FECD8D3AFE04EF36" box="[312,394,1833,1859]" name="Yemen" pageId="17" pageNumber="18">Yemen</collectingCountry>
(new record), North
<collectingCountry id="2BD57614FFB47109FD748D3AFD6BEF36" box="[641,741,1833,1859]" name="Ethiopia" pageId="17" pageNumber="18">Ethiopia</collectingCountry>
(Bischoff 1920).
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<emphasis id="61B6EA96FFB7710AFEFD8B0FFEE0E940" box="[264,366,284,309]" italics="true" pageId="18" pageNumber="19">Remarks</emphasis>
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The specimens from
<collectingCountry id="2BD57614FFB7710AFDB28B57FD17E92B" box="[583,665,324,350]" name="Yemen" pageId="18" pageNumber="19">Yemen</collectingCountry>
are totally black whereas the
<typeStatus id="8C798826FFB7710AFBD28B57FBD7E92B" box="[1063,1113,324,350]" pageId="18" pageNumber="19">type</typeStatus>
specimens from
<collectingCountry id="2BD57614FFB7710AFEFD8B7FFEE5E9F3" box="[264,363,364,390]" name="Ethiopia" pageId="18" pageNumber="19">Ethiopia</collectingCountry>
have a ferruginous­red propodeum. Body length 5.2–6.0 mm.
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262
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<mods:identifier id="D1BAFAC069022AC84DA5D1C6539F76BE" type="DOI">10.5281/zenodo.172671</mods:identifier>
<mods:identifier id="9FBB2E04A1CFAEE114CBEC250AEAAC6D" type="GBIF-Dataset">b0d82648-af13-4bba-8318-49fa28bd2515</mods:identifier>
<mods:identifier id="AAED86093019F3D6B165F96F606D3F65" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="B9336FCB523EF02605C3296933A1CFDC" type="Zenodo-Dep">172671</mods:identifier>
</mods:mods>
<treatment id="DB6B8792FFB57108FEFD8B0FFA9DEEF4" ID-DOI="http://doi.org/10.5281/zenodo.6252949" ID-GBIF-Taxon="119344616" ID-Zenodo-Dep="6252949" LSID="urn:lsid:plazi:treatment:DB6B8792FFB57108FEFD8B0FFA9DEEF4" httpUri="http://treatment.plazi.org/id/DB6B8792FFB57108FEFD8B0FFA9DEEF4" lastPageNumber="17" pageId="16" pageNumber="17">
<subSubSection id="1BD8650FFFB57108FEFD8B0FFC81E940" box="[264,783,283,310]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="537D3684FFB57108FEFD8B0FFC81E940" blockId="16.[264,783,283,310]" box="[264,783,283,310]" pageId="16" pageNumber="17">
<heading id="083581E8FFB57108FEFD8B0FFC81E940" bold="true" box="[264,783,283,310]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="61B6EA96FFB57108FEFD8B0FFC81E940" bold="true" box="[264,783,283,310]" pageId="16" pageNumber="17">
<emphasis id="61B6EA96FFB57108FEFD8B0FFE33E943" bold="true" box="[264,445,284,310]" italics="true" pageId="16" pageNumber="17">*Strangulotilla</emphasis>
<taxonomicName id="94C24D07FFB57108FE318B0FFC81E940" authority="(Magretti 1906)" authorityName="Magretti" authorityYear="1906" baseAuthorityName="Magretti" baseAuthorityYear="1906" box="[452,783,283,310]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="samharica">
<emphasis id="61B6EA96FFB57108FE318B0FFDCDE943" bold="true" box="[452,579,284,310]" italics="true" pageId="16" pageNumber="17">samharica</emphasis>
(Magretti 1906)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB57108FEFD8B74FB95E988" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="537D3684FFB57108FEFD8B74FC60E9C8" blockId="16.[264,1323,358,510]" pageId="16" pageNumber="17">
<taxonomicName id="94C24D07FFB57108FEFD8B74FD27E908" authority="Magretti 1906: 55" authorityName="Magretti" authorityPageNumber="55" authorityYear="1906" box="[264,681,358,382]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="samharica">
<emphasis id="61B6EA96FFB57108FEFD8B74FE5FE90B" box="[264,465,359,382]" italics="true" pageId="16" pageNumber="17">Mutilla samharica</emphasis>
Magretti 1906: 55
</taxonomicName>
, fig. 5, Ψ,
<typeStatus id="8C798826FFB57108FCD98B75FC00E908" box="[812,910,358,381]" pageId="16" pageNumber="17" type="lectotype">lectotype</typeStatus>
(designated
<bibRefCitation id="37534B75FFB57108FBEA8B75FB4BE908" author="Bischoff" box="[1055,1221,358,381]" pageId="16" pageNumber="47" refString="Bischoff, H. (1920 - 1921) Monographie der Mutilliden Afrikas. Archiv fur Naturgeschichte, (1920) 86 A (1 / 3), 1 - 480; (1921) 86 A (4), 481 - 830 + plates 1 - 7." type="journal article" year="1921">Bischoff, 1921</bibRefCitation>
: 545) —
<collectingCountry id="2BD57614FFB57108FECD8B95FE0DE9E8" box="[312,387,390,413]" name="Eritrea" pageId="16" pageNumber="17">Eritrea</collectingCountry>
, Allato,
<date id="277C1044FFB57108FE2A8B95FDC9E9E8" box="[479,583,390,413]" pageId="16" pageNumber="17" value="1900-01-24">24.I.1900</date>
, Sabagruma, lost (
<bibRefCitation id="37534B75FFB57108FCE58B95FC4AE9E8" author="Nonveiller" box="[784,964,390,413]" pageId="16" pageNumber="49" refString="Nonveiller, G. (1978) Recherches sur les Mutillides de l'Afrique (Hymenoptera, Mutillidae). VIII. Revision des genres Ctenotilla, Cephalotilla et Pseudocephalotilla sensu Bischoff. Memoires publies par l'Institut pour la Protection des Plantes, Belgrad, 13, 1 - 184." type="journal article" year="1978">Nonveiller 1978</bibRefCitation>
);
<typeStatus id="8C798826FFB57108FC2D8B95FBA3E9E8" box="[984,1069,390,413]" pageId="16" pageNumber="17" type="neotype">neotype</typeStatus>
(designated
<bibRefCitation id="37534B75FFB57108FB4D8B95FEFFE9C8" author="Nonveiller" pageId="16" pageNumber="49" refString="Nonveiller, G. (1978) Recherches sur les Mutillides de l'Afrique (Hymenoptera, Mutillidae). VIII. Revision des genres Ctenotilla, Cephalotilla et Pseudocephalotilla sensu Bischoff. Memoires publies par l'Institut pour la Protection des Plantes, Belgrad, 13, 1 - 184." type="journal article" year="1978">Nonveiller 1978</bibRefCitation>
: 51) —
<collectingCountry id="2BD57614FFB57108FE328BB5FDB7E9C8" box="[455,569,422,445]" name="Cameroon" pageId="16" pageNumber="17">Cameroon</collectingCountry>
, Station de Guétalé,
<date id="277C1044FFB57108FCE08BB5FC0FE9C8" box="[789,897,422,445]" pageId="16" pageNumber="17" value="1967-09-05">5.IX.1967</date>
[MSNG].
</paragraph>
<paragraph id="537D3684FFB57108FEFD8BD4FC74E9A8" blockId="16.[264,1323,358,510]" box="[264,1018,454,478]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="73D211AAFFB57108FEFD8BD4FC74E9A8" box="[264,1018,454,478]" pageId="16" pageNumber="17">
<taxonomicName id="94C24D07FFB57108FEFD8BD4FCE0E9A8" authority="Bischoff 1921: 545" authorityName="Bischoff" authorityPageNumber="545" authorityYear="1921" box="[264,878,454,478]" class="Insecta" family="Mutillidae" genus="Ctenotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="thoracosulcata" subSpecies="samharica">
<emphasis id="61B6EA96FFB57108FEFD8BD4FD1EE9AB" box="[264,656,455,478]" italics="true" pageId="16" pageNumber="17">Ctenotilla thoracosulcata samharica</emphasis>
:
<treatmentCitation id="D2631095FFB57108FD6A8BD5FCE0E9A8" author="Bischoff" box="[671,878,454,477]" page="545" pageId="16" pageNumber="17" year="1921">
<bibRefCitation id="37534B75FFB57108FD6A8BD5FCB4E9A8" author="Bischoff" box="[671,826,454,477]" pageId="16" pageNumber="47" refString="Bischoff, H. (1920 - 1921) Monographie der Mutilliden Afrikas. Archiv fur Naturgeschichte, (1920) 86 A (1 / 3), 1 - 480; (1921) 86 A (4), 481 - 830 + plates 1 - 7." type="journal article" year="1921">Bischoff 1921</bibRefCitation>
: 545
</treatmentCitation>
</taxonomicName>
, Ψ, (
<collectingCountry id="2BD57614FFB57108FC558BD5FC61E9A8" box="[928,1007,454,477]" name="Eritrea" pageId="16" pageNumber="17">Eritrea</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="537D3684FFB57108FEFD8BF4FB95E988" blockId="16.[264,1323,358,510]" box="[264,1051,486,510]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="73D211AAFFB57108FEFD8BF4FB95E988" box="[264,1051,486,510]" pageId="16" pageNumber="17">
<emphasis id="61B6EA96FFB57108FEFD8BF4FD9DE98B" box="[264,531,487,510]" italics="true" pageId="16" pageNumber="17">
Strangulotilla
<taxonomicName id="94C24D07FFB57108FE578BF4FD9DE98B" authority="Nonveiller 1978: 50" authorityName="Nonveiller" authorityPageNumber="50" authorityYear="1978" box="[418,531,487,510]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="samharica">samharica</taxonomicName>
</emphasis>
:
<treatmentCitation id="D2631095FFB57108FDD48BF5FD74E988" author="Nonveiller" box="[545,762,486,509]" page="50" pageId="16" pageNumber="17" year="1978">
<bibRefCitation id="37534B75FFB57108FDD48BF5FD5CE988" author="Nonveiller" box="[545,722,486,509]" pageId="16" pageNumber="49" refString="Nonveiller, G. (1978) Recherches sur les Mutillides de l'Afrique (Hymenoptera, Mutillidae). VIII. Revision des genres Ctenotilla, Cephalotilla et Pseudocephalotilla sensu Bischoff. Memoires publies par l'Institut pour la Protection des Plantes, Belgrad, 13, 1 - 184." type="journal article" year="1978">Nonveiller 1978</bibRefCitation>
: 50
</treatmentCitation>
, Ψ ♂ (
<collectingCountry id="2BD57614FFB57108FCB68BF5FC1EE988" box="[835,912,486,509]" name="Eritrea" pageId="16" pageNumber="17">Eritrea</collectingCountry>
,
<collectingCountry id="2BD57614FFB57108FC6E8BF5FB81E988" box="[923,1039,486,509]" name="Cameroon" pageId="16" pageNumber="17">Cameroon</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB57108FEFD8844FC6FEC84" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="537D3684FFB57108FEFD8844FE68EA05" blockId="16.[264,1323,599,1265]" box="[264,486,599,624]" pageId="16" pageNumber="17">
<emphasis id="61B6EA96FFB57108FEFD8844FE68EA05" box="[264,486,599,624]" italics="true" pageId="16" pageNumber="17">Material examined</emphasis>
</paragraph>
<paragraph id="537D3684FFB57108FECD886CFC6FEC84" blockId="16.[264,1323,599,1265]" pageId="16" pageNumber="17">
Form with black propodeum.
<collectingCountry id="2BD57614FFB57108FD5C886CFC8EEAEC" box="[681,768,639,665]" name="Yemen" pageId="16" pageNumber="17">Yemen</collectingCountry>
: Al Lahima,
<date id="277C1044FFB57108FC52886CFB0FEAEC" box="[935,1153,639,665]" pageId="16" pageNumber="17" value="2000-10-16" valueMax="2000-12-31" valueMin="2000-10-16">16.X–31.XII.2000</date>
, MT, AvH &amp; A.M. Hager, 1 ɗ [
<collectionCode id="35D3AE41FFB57108FE1A88B4FDC1EAB4" LSID="urn:lsid:biocol.org:col:34975" box="[495,591,679,705]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="16" pageNumber="17">ZMAN</collectionCode>
];
<date id="277C1044FFB57108FD9388B4FCC4EAB4" box="[614,842,679,705]" pageId="16" pageNumber="17" value="2001-07-24" valueMax="2001-09-17" valueMin="2001-07-24">24.VII–17.IX.2001</date>
, MT, AvH, 1 ɗ [
<collectionCode id="35D3AE41FFB57108FBD088B4FB08EAB4" LSID="urn:lsid:biocol.org:col:34992" box="[1061,1158,679,705]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">RMNH</collectionCode>
];
<quantity id="943A9B61FFB57108FB6B88B4FB63EAB4" box="[1182,1261,679,705]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="16" pageNumber="17" unit="km" value="12.0">12 km</quantity>
NW Manakhah,
<date id="277C1044FFB57108FE6688DCFDF5EA9C" box="[403,635,719,745]" pageId="16" pageNumber="17" value="2001-07-03" valueMax="2001-08-21" valueMin="2001-07-03">3.VII–21.VIII.2001</date>
, MT, AvH, 14 ɗ [
<collectionCode id="35D3AE41FFB57108FCA488DCFC3CEA9C" LSID="urn:lsid:biocol.org:col:34992" box="[849,946,719,745]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">RMNH</collectionCode>
];
<date id="277C1044FFB57108FC3388DCFB01EA9C" box="[966,1167,719,745]" pageId="16" pageNumber="17" value="2003-05-15" valueMax="2003-06-23" valueMin="2003-05-15">15.V–23VI.2003</date>
, MT, AvH, 2 ɗ [
<collectionCode id="35D3AE41FFB57108FEDF88E4FE05EB64" LSID="urn:lsid:biocol.org:col:34992" box="[298,395,759,785]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">RMNH</collectionCode>
];
<date id="277C1044FFB57108FE6A88E4FDFBEB64" box="[415,629,759,785]" pageId="16" pageNumber="17" value="2003-06-24" valueMax="2003-07-04" valueMin="2003-06-24">24.VI–4.VII.2003</date>
, MT, AvH, 6 ɗ [
<collectionCode id="35D3AE41FFB57108FCCE88E4FC12EB64" LSID="urn:lsid:biocol.org:col:34992" box="[827,924,759,785]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">RMNH</collectionCode>
];
<date id="277C1044FFB57108FC4588E4FB01EB64" box="[944,1167,759,785]" pageId="16" pageNumber="17" value="2003-08-04" valueMax="2003-09-15" valueMin="2003-08-04">4.VIII–15.IX.2003</date>
, MT, AvH, 2 ɗ [
<collectionCode id="35D3AE41FFB57108FEDF890CFE05EB4C" LSID="urn:lsid:biocol.org:col:34992" box="[298,395,799,825]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">RMNH</collectionCode>
]. Form with ferruginous­red propodeum.
<collectingCountry id="2BD57614FFB57108FC80890CFC42EB4C" box="[885,972,799,825]" name="Yemen" pageId="16" pageNumber="17">Yemen</collectingCountry>
: Al Kadan,
<date id="277C1044FFB57108FBAD890CFAABEB4C" box="[1112,1317,799,825]" pageId="16" pageNumber="17" value="1998-02-17" valueMax="1998-03-31" valueMin="1998-02-17">17.II–31.III.1998</date>
, MT, AvH &amp; H.M. Naser, 1 ɗ [
<collectionCode id="35D3AE41FFB57108FD9A8954FD40EB14" LSID="urn:lsid:biocol.org:col:34975" box="[623,718,839,865]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="16" pageNumber="17">ZMAN</collectionCode>
]; Al Lahima,
<date id="277C1044FFB57108FC858954FBC4EB14" box="[880,1098,839,865]" pageId="16" pageNumber="17" value="2000-10-16" valueMax="2000-12-31" valueMin="2000-10-16">16.X–31.XII.2000</date>
, MT, AvH &amp; A.M. Hager, 2 ɗ [
<collectionCode id="35D3AE41FFB57108FE54897CFD8EEBFC" LSID="urn:lsid:biocol.org:col:34975" box="[417,512,879,905]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="16" pageNumber="17">ZMAN</collectionCode>
]; Lahj,
<date id="277C1044FFB57108FD94897CFCBFEBFC" box="[609,817,879,905]" pageId="16" pageNumber="17" value="2000-05-17" valueMax="2000-06-15" valueMin="2000-05-17">17.V–15.VI.2000</date>
, MT, AvH &amp; A. Sallam, 37 ɗ [
<collectionCode id="35D3AE41FFB57108FB4B897CFA91EBFC" LSID="urn:lsid:biocol.org:col:34992" box="[1214,1311,879,905]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">RMNH</collectionCode>
];
<date id="277C1044FFB57108FEFD8984FE35EBC4" box="[264,443,919,945]" pageId="16" pageNumber="17" value="2000-06-15" valueMax="2000-06-30" valueMin="2000-06-15">15–30.VI.2000</date>
, MT, AvH &amp; A. Sallam, 8 ɗ [
<collectionCode id="35D3AE41FFB57108FCE28984FCF6EBC4" LSID="urn:lsid:biocol.org:col:34992" box="[791,888,919,945]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">RMNH</collectionCode>
];
<date id="277C1044FFB57108FC7E8984FC76EBC4" box="[907,1016,919,945]" pageId="16" pageNumber="17" value="2000-07">VII.2000</date>
, MT, AvH &amp; A. Sallam, 4 ɗ [
<collectionCode id="35D3AE41FFB57108FED989ACFE03EBAC" LSID="urn:lsid:biocol.org:col:34992" box="[300,397,959,985]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">RMNH</collectionCode>
];
<date id="277C1044FFB57108FE5689ACFD97EBAC" box="[419,537,959,985]" pageId="16" pageNumber="17" value="2000-08">VIII.2000</date>
, MT, AvH &amp; A. Sallam, 2 ɗ [
<collectionCode id="35D3AE41FFB57108FC7F89ACFC65EBAC" LSID="urn:lsid:biocol.org:col:34992" box="[906,1003,959,985]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">RMNH</collectionCode>
];
<date id="277C1044FFB57108FBF589ACFBEAEBAC" box="[1024,1124,959,985]" pageId="16" pageNumber="17" value="2000-09">IX.2000</date>
, MT, AvH &amp; A. Sallam, 6 ɗ [
<collectionCode id="35D3AE41FFB57108FE5689F4FD8AEC74" LSID="urn:lsid:biocol.org:col:34992" box="[419,516,999,1025]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">RMNH</collectionCode>
];
<date id="277C1044FFB57108FDED89F4FDFCEC74" box="[536,626,999,1025]" pageId="16" pageNumber="17" value="2000-10">X.2000</date>
, MT, AvH &amp; A. Sallam, 2 ɗ [
<collectionCode id="35D3AE41FFB57108FC2089F4FBB8EC74" LSID="urn:lsid:biocol.org:col:34992" box="[981,1078,999,1025]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">RMNH</collectionCode>
];
<date id="277C1044FFB57108FBBF89F4FB23EC74" box="[1098,1197,999,1025]" pageId="16" pageNumber="17" value="2000-11">XI.2000</date>
, MT, AvH &amp; A. Sallam, 13 ɗ [
<collectionCode id="35D3AE41FFB57108FE038E1CFDD9EC5C" LSID="urn:lsid:biocol.org:col:34992" box="[502,599,1039,1065]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">RMNH</collectionCode>
];
<date id="277C1044FFB57108FD9E8E1CFD37EC5C" box="[619,697,1039,1065]" pageId="16" pageNumber="17" value="2001-01">I.2001</date>
, MT, AvH &amp; A. Sallam, 9 ɗ [
<collectionCode id="35D3AE41FFB57108FBEA8E1CFB0EEC5C" LSID="urn:lsid:biocol.org:col:34992" box="[1055,1152,1039,1065]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">RMNH</collectionCode>
];
<date id="277C1044FFB57108FB618E1CFB62EC5C" box="[1172,1260,1039,1065]" pageId="16" pageNumber="17" value="2001-02">II.2001</date>
, MT, AvH &amp; A. Sallam, 8 ɗ [
<collectionCode id="35D3AE41FFB57108FDD58E24FD0FEC24" LSID="urn:lsid:biocol.org:col:34992" box="[544,641,1079,1105]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">RMNH</collectionCode>
];
<date id="277C1044FFB57108FD618E24FD7BEC24" box="[660,757,1079,1105]" pageId="16" pageNumber="17" value="2001-03">III.2001</date>
, MT, AvH &amp; A. Sallam, 2 ɗ [
<collectionCode id="35D3AE41FFB57108FBA78E24FB3DEC24" LSID="urn:lsid:biocol.org:col:34992" box="[1106,1203,1079,1105]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">RMNH</collectionCode>
];
<date id="277C1044FFB57108FB338E24FAA8EC24" box="[1222,1318,1079,1105]" pageId="16" pageNumber="17" value="2001-04">IV.2001</date>
, MT, AvH &amp; A. Sallam, 8 ɗ [
<collectionCode id="35D3AE41FFB57108FD818E4CFD5BEC0C" LSID="urn:lsid:biocol.org:col:34992" box="[628,725,1119,1145]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">RMNH</collectionCode>
];
<date id="277C1044FFB57108FD198E4CFCCCEC0C" box="[748,834,1119,1145]" pageId="16" pageNumber="17" value="2001-05">V.2001</date>
, MT, AvH &amp; A. Sallam, 1 ɗ [
<collectionCode id="35D3AE41FFB57108FB4B8E4CFA91EC0C" LSID="urn:lsid:biocol.org:col:34992" box="[1214,1311,1119,1145]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">RMNH</collectionCode>
];
<date id="277C1044FFB57108FEFD8E94FE2DECD4" box="[264,419,1159,1185]" pageId="16" pageNumber="17" value="2001-07" valueMax="2001-09-31" valueMin="2001-07-00">VII–IX.2001</date>
, MT, AvH &amp; A. Sallam, 5 ɗ [
<collectionCode id="35D3AE41FFB57108FCF88E94FCE0ECD4" LSID="urn:lsid:biocol.org:col:34992" box="[781,878,1159,1185]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">RMNH</collectionCode>
];
<date id="277C1044FFB57108FC768E94FBE8ECD4" box="[899,1126,1159,1185]" pageId="16" pageNumber="17" value="2001-07-24" valueMax="2001-09-17" valueMin="2001-07-24">24.VII–17.IX.2001</date>
, MT, AvH &amp; A. Sallam, 2 ɗ [
<collectionCode id="35D3AE41FFB57108FE498EBCFD93ECBC" LSID="urn:lsid:biocol.org:col:34992" box="[444,541,1199,1225]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">RMNH</collectionCode>
];
<date id="277C1044FFB57108FDCF8EBCFC8BECBC" box="[570,773,1199,1225]" pageId="16" pageNumber="17" value="2001-10-01" valueMax="2001-12-17" valueMin="2001-10-01">1.X–17.XII.2001</date>
, MT, AvH &amp; A. Sallam, 23 ɗ [
<collectionCode id="35D3AE41FFB57108FB4B8EBCFA91ECBC" LSID="urn:lsid:biocol.org:col:34992" box="[1214,1311,1199,1225]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">RMNH</collectionCode>
];
<date id="277C1044FFB57108FEFD8EC4FE16EC84" box="[264,408,1239,1265]" pageId="16" pageNumber="17" value="2001-12-17">17.XII.2001</date>
–
<date id="277C1044FFB57108FE5C8EC4FD97EC84" box="[425,537,1239,1265]" pageId="16" pageNumber="17" value="2002-01-31">31.I.2002</date>
, MT, AvH &amp; A. Sallam, 9 ɗ [
<collectionCode id="35D3AE41FFB57108FC808EC4FC58EC84" LSID="urn:lsid:biocol.org:col:34992" box="[885,982,1239,1265]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">RMNH</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB57108FEFD8F34FC7EED1C" box="[264,1008,1319,1385]" pageId="16" pageNumber="17" type="distribution">
<paragraph id="537D3684FFB57108FEFD8F34FE19ED35" blockId="16.[264,1008,1319,1385]" box="[264,407,1319,1344]" pageId="16" pageNumber="17">
<emphasis id="61B6EA96FFB57108FEFD8F34FE19ED35" box="[264,407,1319,1344]" italics="true" pageId="16" pageNumber="17">Distribution</emphasis>
</paragraph>
<paragraph id="537D3684FFB57108FECD8F5CFC7EED1C" blockId="16.[264,1008,1319,1385]" box="[312,1008,1359,1385]" pageId="16" pageNumber="17">
<collectingCountry id="2BD57614FFB57108FECD8F5CFE04ED1C" box="[312,394,1359,1385]" name="Yemen" pageId="16" pageNumber="17">Yemen</collectingCountry>
(new record),
<collectingCountry id="2BD57614FFB57108FDC38F5CFD07ED1C" box="[566,649,1359,1385]" name="Eritrea" pageId="16" pageNumber="17">Eritrea</collectingCountry>
,
<collectingCountry id="2BD57614FFB57108FD618F5CFC80ED1C" box="[660,782,1359,1385]" name="Cameroon" pageId="16" pageNumber="17">Cameroon</collectingCountry>
(
<bibRefCitation id="37534B75FFB57108FCE88F5CFC6DED1C" author="Nonveiller" box="[797,995,1359,1385]" pageId="16" pageNumber="49" refString="Nonveiller, G. (1978) Recherches sur les Mutillides de l'Afrique (Hymenoptera, Mutillidae). VIII. Revision des genres Ctenotilla, Cephalotilla et Pseudocephalotilla sensu Bischoff. Memoires publies par l'Institut pour la Protection des Plantes, Belgrad, 13, 1 - 184." type="journal article" year="1978">Nonveiller 1978</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB57108FEFD8F8CFA9DEEF4" pageId="16" pageNumber="17" type="discussion">
<paragraph id="537D3684FFB57108FEFD8F8CFEE0EDCD" blockId="16.[264,1324,1439,1665]" box="[264,366,1439,1464]" pageId="16" pageNumber="17">
<emphasis id="61B6EA96FFB57108FEFD8F8CFEE0EDCD" box="[264,366,1439,1464]" italics="true" pageId="16" pageNumber="17">Remarks</emphasis>
</paragraph>
<paragraph id="537D3684FFB57108FECD8FD4FA9DEEF4" blockId="16.[264,1324,1439,1665]" pageId="16" pageNumber="17">
The coloration of the male’s mesosoma is variable from wholly ferruginous­red to ferruginous­red with black propodeum. The latter form has been described by
<bibRefCitation id="37534B75FFB57108FB588FFCFED9EE44" author="Nonveiller" pageId="16" pageNumber="49" refString="Nonveiller, G. (1978) Recherches sur les Mutillides de l'Afrique (Hymenoptera, Mutillidae). VIII. Revision des genres Ctenotilla, Cephalotilla et Pseudocephalotilla sensu Bischoff. Memoires publies par l'Institut pour la Protection des Plantes, Belgrad, 13, 1 - 184." type="journal article" year="1978">Nonveiller (1978)</bibRefCitation>
from
<collectingCountry id="2BD57614FFB57108FE5D8C04FDACEE44" box="[424,546,1559,1585]" name="Cameroon" pageId="16" pageNumber="17">Cameroon</collectingCountry>
and also occurs in
<collectingCountry id="2BD57614FFB57108FCEA8C04FCFAEE44" box="[799,884,1559,1585]" name="Yemen" pageId="16" pageNumber="17">Yemen</collectingCountry>
, but most abundant in
<collectingCountry id="2BD57614FFB57108FB6D8C04FB64EE44" box="[1176,1258,1559,1585]" name="Yemen" pageId="16" pageNumber="17">Yemen</collectingCountry>
(
<specimenCount id="45C4FD0DFFB57108FB0A8C04FE0CEE2C" pageId="16" pageNumber="17" type="generic">147 specimens</specimenCount>
from 170) is the form with ferruginous­red propodeum. There are several specimens with intermediate propodeal castaneous coloration. Body length
<quantity id="943A9B61FFB57108FB8B8C74FA9EEEF4" box="[1150,1296,1639,1665]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" metricValueMax="8.8" metricValueMin="4.8" pageId="16" pageNumber="17" unit="mm" value="6.8" valueMax="8.8" valueMin="4.8">4.8–8.8 mm</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

238
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<treatment id="DB6B8792FFB6710DFEFD8C72FC03E92B" ID-DOI="http://doi.org/10.5281/zenodo.6252961" ID-GBIF-Taxon="119344632" ID-Zenodo-Dep="6252961" LSID="urn:lsid:plazi:treatment:DB6B8792FFB6710DFEFD8C72FC03E92B" httpUri="http://treatment.plazi.org/id/DB6B8792FFB6710DFEFD8C72FC03E92B" lastPageId="21" lastPageNumber="21" pageId="19" pageNumber="20">
<subSubSection id="1BD8650FFFB6710BFEFD8C72FD40EE0E" box="[264,718,1633,1659]" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="537D3684FFB6710BFEFD8C72FD40EE0E" blockId="19.[264,718,1633,1659]" box="[264,718,1633,1659]" pageId="19" pageNumber="20">
<heading id="083581E8FFB6710BFEFD8C72FD40EE0E" bold="true" box="[264,718,1633,1659]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
<emphasis id="61B6EA96FFB6710BFEFD8C72FD40EE0E" bold="true" box="[264,718,1633,1659]" pageId="19" pageNumber="20">
<emphasis id="61B6EA96FFB6710BFEFD8C72FE9AEE0E" bold="true" box="[264,276,1633,1659]" italics="true" pageId="19" pageNumber="20">*</emphasis>
<taxonomicName id="94C24D07FFB6710BFEE18C72FD40EE0E" authority="(Magretti 1898)" authorityName="Magretti" authorityYear="1898" baseAuthorityName="Magretti" baseAuthorityYear="1898" box="[276,718,1633,1659]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="milmili">
<emphasis id="61B6EA96FFB6710BFEE18C72FD8CEE0E" bold="true" box="[276,514,1633,1659]" italics="true" pageId="19" pageNumber="20">Tropidotilla milmili</emphasis>
(
<bibRefCitation id="37534B75FFB6710BFDE78C72FD4BEE0E" author="Magretti" box="[530,709,1633,1659]" lastPageId="19" lastPageNumber="49" pageId="19" pageNumber="48" refString="Magretti, P. (1898) Imenotteri della seconda spedizione di Don Eugenio dei principi Ruspoli nei paesi Galla e Somali. Annali del Museo Civico di Storia Naturale di Genova, ser. 2 a, 19, 37 - 42." type="journal article" year="1898">Magretti 1898</bibRefCitation>
)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<paragraph id="537D3684FFB6710BFEFD8CBFFD54EF76" pageId="19" pageNumber="20">
<table id="21C2C424FFB68EE7FEFD8CB8FAA5EF76" box="[264,1323,1707,1795]" gridcols="7" gridrows="3" pageId="19" pageNumber="20">
<tr id="EDF234C6FFB68EE7FEFD8CB8FAA5EEB6" box="[264,1323,1707,1731]" gridrow="0" pageId="19" pageNumber="20">
<th id="AE235DBAFFB68EE7FEFD8CB8FC9AEEB6" box="[264,788,1707,1731]" gridcol="0" gridrow="0" pageId="19" pageNumber="20">
<taxonomicName id="94C24D07FFB6710BFEFD8CBFFD0AEEB6" authority="Magretti 1898: 40" authorityName="Magretti" authorityPageNumber="40" authorityYear="1898" box="[264,644,1708,1731]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="milmili">
<emphasis id="61B6EA96FFB6710BFEFD8CBFFE23EEB6" box="[264,429,1708,1731]" italics="true" pageId="19" pageNumber="20">Mutilla milmili</emphasis>
Magretti 1898: 40
</taxonomicName>
, ♂, holotype
</th>
<th id="AE235DBAFFB68EE7FCEA8CB8FCF1EEB6" box="[799,895,1707,1731]" gridcol="1" gridrow="0" pageId="19" pageNumber="20">&quot;Milmil&quot;</th>
<th id="AE235DBAFFB68EE7FC7E8CB8FC5CEEB6" box="[907,978,1707,1731]" gridcol="2" gridrow="0" pageId="19" pageNumber="20">[North</th>
<th id="AE235DBAFFB68EE7FC288CB8FBCAEEB6" box="[989,1092,1707,1731]" gridcol="3" gridrow="0" pageId="19" pageNumber="20">Somalia],</th>
<th id="AE235DBAFFB68EE7FBBA8CB8FB14EEB6" box="[1103,1178,1707,1731]" gridcol="4" gridrow="0" pageId="19" pageNumber="20">I.1893,</th>
<th id="AE235DBAFFB68EE7FB508CB8FB43EEB6" box="[1189,1229,1707,1731]" gridcol="5" gridrow="0" pageId="19" pageNumber="20">leg.</th>
<th id="AE235DBAFFB68EE7FB2D8CB8FAA5EEB6" box="[1240,1323,1707,1731]" gridcol="6" gridrow="0" pageId="19" pageNumber="20">Ruspoli</th>
</tr>
<tr id="EDF234C6FFB68EE7FEFD8CDFFAA5EE96" box="[264,1323,1740,1763]" gridrow="1" pageId="19" pageNumber="20" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="AE235DBAFFB68EE7FEFD8CDFFC9AEE96" box="[264,788,1740,1763]" gridcol="0" gridrow="1" pageId="19" pageNumber="20">[MCSG].</th>
</tr>
<tr id="EDF234C6FFB68EE7FEFD8CF8FAA5EF76" box="[264,1323,1771,1795]" gridrow="2" pageId="19" pageNumber="20" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="AE235DBAFFB68EE7FEFD8CF8FC9AEF76" box="[264,788,1771,1795]" gridcol="0" gridrow="2" pageId="19" pageNumber="20">
<taxonomicName id="94C24D07FFB6710BFEFD8CFFFD38EF76" authority="Bischoff 1920: 159" authorityName="Bischoff" authorityPageNumber="159" authorityYear="1920" box="[264,694,1772,1795]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="milmili">
<emphasis id="61B6EA96FFB6710BFEFD8CFFFE56EF76" box="[264,472,1772,1795]" italics="true" pageId="19" pageNumber="20">Tropidotilla milmili</emphasis>
: Bischoff 1920: 159
</taxonomicName>
, ♂.
</th>
</tr>
</table>
</paragraph>
<subSubSection id="1BD8650FFFB1710CFEFD8B0FFE08EA3B" pageId="20" pageNumber="21" type="diagnosis">
<paragraph id="537D3684FFB1710CFEFD8B0FFEF1E940" blockId="20.[264,1323,284,590]" box="[264,383,284,309]" pageId="20" pageNumber="21">
<emphasis id="61B6EA96FFB1710CFEFD8B0FFEF1E940" box="[264,383,284,309]" italics="true" pageId="20" pageNumber="21">Diagnosis</emphasis>
</paragraph>
<paragraph id="537D3684FFB1710CFECD8B57FE08EA3B" blockId="20.[264,1323,284,590]" pageId="20" pageNumber="21">
The female of
<taxonomicName id="94C24D07FFB1710CFE1F8B57FD5FE928" box="[490,721,324,349]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="milmili">
<emphasis id="61B6EA96FFB1710CFE1F8B57FD5FE928" box="[490,721,324,349]" italics="true" pageId="20" pageNumber="21">Tropidotilla milmili</emphasis>
</taxonomicName>
is similar to that of
<taxonomicName id="94C24D07FFB1710CFC3F8B57FBDAE928" box="[970,1108,324,349]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="fimbriata">
<emphasis id="61B6EA96FFB1710CFC3F8B57FBDAE928" box="[970,1108,324,349]" italics="true" pageId="20" pageNumber="21">T. fimbriata</emphasis>
</taxonomicName>
in having similar color patterns and size. It can be distinguished from the latter species in having a differently shaped mesosoma (slightly widened posteriorly and length 1.15X its maximal width in
<taxonomicName id="94C24D07FFB1710CFE818BAFFE73E9A0" box="[372,509,444,469]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="fimbriata">
<emphasis id="61B6EA96FFB1710CFE818BAFFE73E9A0" box="[372,509,444,469]" italics="true" pageId="20" pageNumber="21">T. fimbriata</emphasis>
</taxonomicName>
), in having a well developed genal carina (weak in
<taxonomicName id="94C24D07FFB1710CFB9B8BAFFB76E9A0" box="[1134,1272,444,469]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="fimbriata">
<emphasis id="61B6EA96FFB1710CFB9B8BAFFB76E9A0" box="[1134,1272,444,469]" italics="true" pageId="20" pageNumber="21">T. fimbriata</emphasis>
</taxonomicName>
), in having narrow apical fringe on T2 (narrow band widened medially in
<taxonomicName id="94C24D07FFB1710CFBCA8BF7FB47E988" box="[1087,1225,484,509]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="fimbriata">
<emphasis id="61B6EA96FFB1710CFBCA8BF7FB47E988" box="[1087,1225,484,509]" italics="true" pageId="20" pageNumber="21">T. fimbriata</emphasis>
</taxonomicName>
), and in having a different shape and position of carina on S1 (weak carina located anteriorly in
<taxonomicName id="94C24D07FFB1710CFAE0881FFEFBEA38" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="fimbriata">
<emphasis id="61B6EA96FFB1710CFAE0881FFEFBEA38" italics="true" pageId="20" pageNumber="21">T. fimbriata</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB1710CFEFD8897FD07EC73" pageId="20" pageNumber="21" type="description">
<paragraph id="537D3684FFB1710CFEFD8897FE1CEAE8" blockId="20.[264,1325,644,1030]" box="[264,402,644,669]" pageId="20" pageNumber="21">
<emphasis id="61B6EA96FFB1710CFEFD8897FE1CEAE8" box="[264,402,644,669]" italics="true" pageId="20" pageNumber="21">Description</emphasis>
</paragraph>
<paragraph id="537D3684FFB1710CFECD88BFFD07EC73" blockId="20.[264,1325,644,1030]" pageId="20" pageNumber="21">FEMALE (hitherto unknown). Body length 5.2–6.0 mm. Head rounded behind eyes. Distance between posterior eye line and posterior headline 0.7X eye length. First flagellomere 1.6X flagellomere 2. Genal carina well developed and connected to hypostomal carina, latter widened at this point. Mesosoma slightly narrow posteriorly, length 1.3X its maximal width. Scutellar scale well developed. Metasoma elongated. T1 transversal with distinct dorsal surface delimited anteriorly by serrate carina and posteriorly by narrow band of yellowish pubescence. S1 with triangular carina in median part. Head, antennae, mesosoma and legs ferruginous­red, metasoma black. T2–T5 with apical fringes of yellowish setae.</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB1710CFEFD8E2FFD48EC83" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="537D3684FFB1710CFEFD8E2FFE68EC20" blockId="20.[264,1323,1084,1270]" box="[264,486,1084,1109]" pageId="20" pageNumber="21">
<emphasis id="61B6EA96FFB1710CFEFD8E2FFE68EC20" box="[264,486,1084,1109]" italics="true" pageId="20" pageNumber="21">Material examined</emphasis>
</paragraph>
<paragraph id="537D3684FFB1710CFECD8E77FD48EC83" blockId="20.[264,1323,1084,1270]" pageId="20" pageNumber="21">
<collectingCountry id="2BD57614FFB1710CFECD8E77FE00EC0B" box="[312,398,1124,1150]" name="Yemen" pageId="20" pageNumber="21">Yemen</collectingCountry>
:
<quantity id="943A9B61FFB1710CFE6B8E77FE63EC0B" box="[414,493,1124,1150]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="20" pageNumber="21" unit="km" value="12.0">12 km</quantity>
NW Manakhah,
<date id="277C1044FFB1710CFD328E77FC3EEC0B" box="[711,944,1124,1150]" pageId="20" pageNumber="21" value="2001-07-03" valueMax="2001-08-21" valueMin="2001-07-03">3.VII–21.VIII.2001</date>
, MT, AvH, 2 Ψ 8 ɗ [
<collectionCode id="35D3AE41FFB1710CFB3D8E77FAA8EC0B" LSID="urn:lsid:biocol.org:col:34992" box="[1224,1318,1124,1150]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="21">RMNH</collectionCode>
,
<collectionCode id="35D3AE41FFB1710CFEFD8E9FFEC5ECD3" box="[264,331,1164,1190]" pageId="20" pageNumber="21">IBSS</collectionCode>
];
<date id="277C1044FFB1710CFE968E9FFDCCECD3" box="[355,578,1164,1190]" pageId="20" pageNumber="21" value="2003-06-24" valueMax="2003-08-04" valueMin="2003-06-24">24.VI–4.VIII.2003</date>
, MT, AvH, 1 Ψ [
<collectionCode id="35D3AE41FFB1710CFCE98E9FFCF3ECD3" LSID="urn:lsid:biocol.org:col:34992" box="[796,893,1164,1190]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="21">RMNH</collectionCode>
];
<date id="277C1044FFB1710CFC618E9FFBEBECD3" box="[916,1125,1164,1190]" pageId="20" pageNumber="21" value="2003-09-15" valueMax="2003-10-22" valueMin="2003-09-15">15.IX–22.X.2003</date>
, MT, AvH, 2 ɗ [
<collectionCode id="35D3AE41FFB1710CFEFA8EA7FEE3ECBB" LSID="urn:lsid:biocol.org:col:34992" box="[271,365,1204,1230]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="21">RMNH</collectionCode>
,
<collectionCode id="35D3AE41FFB1710CFE8F8EA7FE33ECBB" box="[378,445,1204,1230]" pageId="20" pageNumber="21">IBSS</collectionCode>
]; Lahj,
<date id="277C1044FFB1710CFDED8EA7FD66ECBB" box="[536,744,1204,1230]" pageId="20" pageNumber="21" value="2000-05-17" valueMax="2000-06-15" valueMin="2000-05-17">17.V–15.VI.2000</date>
, MT, AvH &amp; A. Sallam, 1 ɗ [
<collectionCode id="35D3AE41FFB1710CFBA48EA7FB3CECBB" LSID="urn:lsid:biocol.org:col:34992" box="[1105,1202,1204,1230]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="21">RMNH</collectionCode>
];
<date id="277C1044FFB1710CFB338EA7FAA8ECBB" box="[1222,1318,1204,1230]" pageId="20" pageNumber="21" value="2001-04">IV.2001</date>
, MT, AvH &amp; A. Sallam, 1 ɗ [
<collectionCode id="35D3AE41FFB1710CFDAC8ECFFD34EC83" LSID="urn:lsid:biocol.org:col:34992" box="[601,698,1244,1270]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="21">RMNH</collectionCode>
];
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB1710CFEFD8F34FBA0ED1C" box="[264,1070,1319,1385]" pageId="20" pageNumber="21" type="distribution">
<paragraph id="537D3684FFB1710CFEFD8F34FE19ED35" blockId="20.[264,1070,1319,1385]" box="[264,407,1319,1344]" pageId="20" pageNumber="21">
<emphasis id="61B6EA96FFB1710CFEFD8F34FE19ED35" box="[264,407,1319,1344]" italics="true" pageId="20" pageNumber="21">Distribution</emphasis>
</paragraph>
<paragraph id="537D3684FFB1710CFECD8F5CFBA0ED1C" blockId="20.[264,1070,1319,1385]" box="[312,1070,1359,1385]" pageId="20" pageNumber="21">
<collectingCountry id="2BD57614FFB1710CFECD8F5CFE04ED1C" box="[312,394,1359,1385]" name="Yemen" pageId="20" pageNumber="21">Yemen</collectingCountry>
(new record),
<collectingCountry id="2BD57614FFB1710CFDC38F5CFD17ED1C" box="[566,665,1359,1385]" name="Somalia" pageId="20" pageNumber="21">Somalia</collectingCountry>
,
<collectingCountry id="2BD57614FFB1710CFD518F5CFD7EED1C" box="[676,752,1359,1385]" name="Sudan" pageId="20" pageNumber="21">Sudan</collectingCountry>
,
<collectingCountry id="2BD57614FFB1710CFD088F5CFCE8ED1C" box="[765,870,1359,1385]" name="Tanzania" pageId="20" pageNumber="21">Tanzania</collectingCountry>
(Bischoff 1920).
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB1710DFEFD8F8AFC03E92B" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="discussion">
<paragraph id="537D3684FFB1710CFEFD8F8AFEE0EDC7" blockId="20.[264,1323,1433,1579]" box="[264,366,1433,1458]" pageId="20" pageNumber="21">
<emphasis id="61B6EA96FFB1710CFEFD8F8AFEE0EDC7" box="[264,366,1433,1458]" italics="true" pageId="20" pageNumber="21">Remarks</emphasis>
</paragraph>
<paragraph id="537D3684FFB1710CFECD8FD2FD44EE5E" blockId="20.[264,1323,1433,1579]" pageId="20" pageNumber="21">
The described female is associated with the male of
<taxonomicName id="94C24D07FFB1710CFC628FD2FB8AEDAF" box="[919,1028,1473,1498]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="milmili">
<emphasis id="61B6EA96FFB1710CFC628FD2FB8AEDAF" box="[919,1028,1473,1498]" italics="true" pageId="20" pageNumber="21">T. milmili</emphasis>
</taxonomicName>
because both sexes were collected at the same locality at the same time and have a similar carina on S1.
<emphasis id="61B6EA96FFB1710CFEFD8C02FD44EE5E" bold="true" box="[264,714,1553,1579]" pageId="20" pageNumber="21">Tribe Smicromyrmini Bischoff 1920</emphasis>
</paragraph>
<paragraph id="537D3684FFB1710DFEFD8C4FFC03E92B" blockId="20.[264,1323,1628,1894]" lastBlockId="21.[264,1323,284,350]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
This tribe is the taxonomically most difficult of the Afrotropical fauna. Bischoff (
<date id="277C1044FFB1710CFEE58C97FE19EEEB" box="[272,407,1668,1694]" pageId="20" pageNumber="21" value="1920" valueMax="1921">1920–1921</date>
) placed 90 Afrotropical species in the genus
<taxonomicName id="94C24D07FFB1710CFC268C97FBFBEEE8" box="[979,1141,1668,1693]" class="Insecta" family="Mutillidae" genus="Smicromyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="61B6EA96FFB1710CFC268C97FBFBEEE8" box="[979,1141,1668,1693]" italics="true" pageId="20" pageNumber="21">Smicromyrme</emphasis>
</taxonomicName>
, that currently belong to many genera. In the last two decades ASL described 11 genera and three subgenera in the tribe Smicromyrmini from the Palaearctic and Oriental regions (
<bibRefCitation id="37534B75FFB1710CFB188CC7FECBEF63" author="Lelej" pageId="20" pageNumber="48" refString="Lelej, A. S. (1985) Osy-nemki (Hymenoptera, Mutillidae) fauny SSSR i sopredelnykh stran [The velvet ants (Hymenoptera, Mutillidae) of the USSR and neighbouring countries]. Nauka, Leningrad. 268 pp., 64 tables. (In Russian)." type="book" year="1985">Lelej 1985</bibRefCitation>
,
<bibRefCitation id="37534B75FFB1710CFEA98CEFFE17EF63" author="Lelej" box="[348,409,1788,1814]" pageId="20" pageNumber="48" refString="Lelej, A. S. (1995) To the knowledge of East Asian species of the tribe Smicromyrmini Bischoff, 1920 (Hymenoptera, Mutillidae) with description of four new genera and eight new species. Far Eastern entomologist, 13, 1 - 28." type="journal article" year="1995">1995</bibRefCitation>
,
<bibRefCitation id="37534B75FFB1710CFE448CEFFE7EEF63" author="Lelej" box="[433,496,1788,1814]" pageId="20" pageNumber="48" refString="Lelej, A. S. (2005) Catalogue of the Mutillidae (Hymenoptera) of the Oriental region. Dalnauka, Vladivostok. 252 pp." type="book" year="2005">2005</bibRefCitation>
). We include here the species from the genera
<taxonomicName id="94C24D07FFB1710CFB818CEFFAA8EF63" authority="Lelej" authorityName="Lelej" box="[1140,1318,1788,1814]" class="Insecta" family="Mutillidae" genus="Dentilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="61B6EA96FFB1710CFB818CEFFB5AEF60" box="[1140,1236,1788,1813]" italics="true" pageId="20" pageNumber="21">Dentilla</emphasis>
Lelej
</taxonomicName>
,
<taxonomicName id="94C24D07FFB1710CFEFD8D37FE2DEF48" box="[264,419,1828,1853]" class="Insecta" family="Mutillidae" genus="Paglianotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="61B6EA96FFB1710CFEFD8D37FE2DEF48" box="[264,419,1828,1853]" italics="true" pageId="20" pageNumber="21">Paglianotilla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7A8557EDFFB1710CFE5E8D37FD97EF4B" box="[427,537,1828,1854]" pageId="20" pageNumber="21" rank="genus">gen. nov.</taxonomicNameLabel>
, and
<taxonomicName id="94C24D07FFB1710CFDAC8D37FC8CEF48" box="[601,770,1828,1853]" class="Insecta" family="Mutillidae" genus="Rasnitsynitilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="61B6EA96FFB1710CFDAC8D37FC8CEF48" box="[601,770,1828,1853]" italics="true" pageId="20" pageNumber="21">Rasnitsynitilla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7A8557EDFFB1710CFCFE8D37FCF8EF4B" box="[779,886,1828,1854]" pageId="20" pageNumber="21" rank="genus">gen. nov.</taxonomicNameLabel>
In addition, we recognize more than 10 species from the genera
<taxonomicName id="94C24D07FFB1710CFDB28D5FFC00EF13" authority="Schuster 1949" authorityName="Schuster" authorityYear="1949" box="[583,910,1868,1894]" class="Insecta" family="Mutillidae" genus="Physetopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="61B6EA96FFB1710CFDB28D5FFD50EF10" box="[583,734,1868,1893]" italics="true" pageId="20" pageNumber="21">Physetopoda</emphasis>
Schuster 1949
</taxonomicName>
,
<taxonomicName id="94C24D07FFB1710CFC6F8D5FFB7AEF13" authority="Thomson 1870" authorityName="Thomson" authorityYear="1870" box="[922,1268,1868,1894]" class="Insecta" family="Mutillidae" genus="Smicromyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="61B6EA96FFB1710CFC6F8D5FFBB2EF10" box="[922,1084,1868,1893]" italics="true" pageId="20" pageNumber="21">Smicromyrme</emphasis>
Thomson 1870
</taxonomicName>
, and
<taxonomicName id="94C24D07FFB0710DFEFD8B0FFD81E943" authority="Lelej 1995" authorityName="Lelej" authorityYear="1995" box="[264,527,284,310]" class="Insecta" family="Mutillidae" genus="Ephucilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="61B6EA96FFB0710DFEFD8B0FFEF4E940" box="[264,378,284,309]" italics="true" pageId="21" pageNumber="22">Ephucilla</emphasis>
<bibRefCitation id="37534B75FFB0710DFE728B0FFD81E943" author="Lelej" box="[391,527,284,310]" pageId="21" pageNumber="48" refString="Lelej, A. S. (1995) To the knowledge of East Asian species of the tribe Smicromyrmini Bischoff, 1920 (Hymenoptera, Mutillidae) with description of four new genera and eight new species. Far Eastern entomologist, 13, 1 - 28." type="journal article" year="1995">Lelej 1995</bibRefCitation>
</taxonomicName>
. The identification of these species requires the revision of the Afrotropical species placed in the genus
<taxonomicName id="94C24D07FFB0710DFD108B57FC08E928" box="[741,902,324,349]" class="Insecta" family="Mutillidae" genus="Smicromyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="61B6EA96FFB0710DFD108B57FC08E928" box="[741,902,324,349]" italics="true" pageId="21" pageNumber="22">Smicromyrme</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<subSubSection id="1BD8650FFFB7710AFEFD8BF7FD26E988" box="[264,680,483,510]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="537D3684FFB7710AFEFD8BF7FD26E988" blockId="18.[264,680,483,510]" box="[264,680,483,510]" pageId="18" pageNumber="19">
<heading id="083581E8FFB7710AFEFD8BF7FD26E988" bold="true" box="[264,680,483,510]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<taxonomicName id="94C24D07FFB7710AFEFD8BF7FD26E988" authority="Klug 1829" authorityName="Klug" authorityYear="1829" box="[264,680,483,510]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="fimbriata">
<emphasis id="61B6EA96FFB7710AFEFD8BF7FD26E988" bold="true" box="[264,680,483,510]" pageId="18" pageNumber="19">
<emphasis id="61B6EA96FFB7710AFEFD8BF7FD80E98B" bold="true" box="[264,526,484,510]" italics="true" pageId="18" pageNumber="19">Tropidotilla fimbriata</emphasis>
(
<bibRefCitation id="37534B75FFB7710AFDE98BF0FD2EE988" author="Klug" box="[540,672,483,509]" pageId="18" pageNumber="48" refString="Klug, J. C. F. (1829) Symbolae physicae seu icones et descriptiones insectorum quae ex itinere per Africam borealem et Asiam occidentalem Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg studio novae aut illustratae redierunt. Hymenoptera. Decas Prima [genera Mutilla et Apterogyna]. Venditur a Mittlero, Berolini. [15 - 21] + tab. 4, 5." type="journal volume" year="1829">Klug 1829</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB7710AFEFD883CFE30EAB2" pageId="18" pageNumber="19" type="reference_group">
<paragraph id="537D3684FFB7710AFEFD883CFC82EA10" blockId="18.[264,1323,558,711]" pageId="18" pageNumber="19">
<taxonomicName id="94C24D07FFB7710AFEFD883CFDB1EA30" authority="Klug 1829" authorityName="Klug" authorityYear="1829" box="[264,575,558,582]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="fimbriata">
<emphasis id="61B6EA96FFB7710AFEFD883CFE4FEA33" box="[264,449,559,582]" italics="true" pageId="18" pageNumber="19">Mutilla fimbriata</emphasis>
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</taxonomicName>
: [15], tab. 4, fig. 3, Ψ &quot;ex Aegypto, ... varietas duplo minor in Arabia felici&quot; [
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,
<collectingCountry id="2BD57614FFB7710AFE23885DFDA8EA10" box="[470,550,590,613]" name="Yemen" pageId="18" pageNumber="19">Yemen</collectingCountry>
],
<typeStatus id="8C798826FFB7710AFDC3885DFD1DEA10" box="[566,659,590,613]" pageId="18" pageNumber="19" type="syntype">syntypes</typeStatus>
in ZMHU.
</paragraph>
<paragraph id="537D3684FFB7710AFEFD887CFC96EAF0" blockId="18.[264,1323,558,711]" box="[264,792,622,646]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="73D211AAFFB7710AFEFD887CFC96EAF0" box="[264,792,622,646]" pageId="18" pageNumber="19">
<taxonomicName id="94C24D07FFB7710AFEFD887CFE3CEAF3" box="[264,434,623,646]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="litoralis">
<emphasis id="61B6EA96FFB7710AFEFD887CFE3CEAF3" box="[264,434,623,646]" italics="true" pageId="18" pageNumber="19">Mutilla litoralis</emphasis>
</taxonomicName>
<taxonomicName id="94C24D07FFB7710AFE4D887DFD7DEAF0" authority="Andre 1910: 7" authorityName="Andre" authorityPageNumber="7" authorityYear="1910" box="[440,755,622,646]" pageId="18" pageNumber="19" rank="variety" variety="fimbriata">
var.
<emphasis id="61B6EA96FFB7710AFE13887CFDC7EAF3" box="[486,585,623,646]" italics="true" pageId="18" pageNumber="19">fimbriata</emphasis>
:
<treatmentCitation id="D2631095FFB7710AFDAD887DFD7DEAF0" author="Andre" box="[600,755,622,645]" page="7" pageId="18" pageNumber="19" year="1910">
<bibRefCitation id="37534B75FFB7710AFDAD887DFD54EAF0" author="Andre" box="[600,730,622,645]" pageId="18" pageNumber="47" refString="Andre, E. (1910) Revision monographique des mutillides de l'Egypte. Memoires de la Societe Entomologique d'Egypte, 1 (2), 1 - 94, 3 plates." type="book chapter" year="1910">André 1910</bibRefCitation>
: 7
</treatmentCitation>
</taxonomicName>
, Ψ.
</treatmentCitationGroup>
</paragraph>
<paragraph id="537D3684FFB7710AFEFD889CFE30EAB2" blockId="18.[264,1323,558,711]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="73D211AAFFB7710AFEFD889CFE30EAB2" pageId="18" pageNumber="19">
<taxonomicName id="94C24D07FFB7710AFEFD889CFD52EAD0" authority="Bischoff 1920: 156" authorityName="Bischoff" authorityPageNumber="156" authorityYear="1920" box="[264,732,654,678]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="fimbriata">
<emphasis id="61B6EA96FFB7710AFEFD889CFE7DEAD3" box="[264,499,655,678]" italics="true" pageId="18" pageNumber="19">Tropidotilla fimbriata</emphasis>
: Bischoff 1920: 156
</taxonomicName>
, Ψ;
<treatmentCitation id="D2631095FFB7710AFCF8889DFC42EAD0" author="Invrea" box="[781,972,654,677]" page="206" pageId="18" pageNumber="19" year="1960">
<bibRefCitation id="37534B75FFB7710AFCF8889DFC1AEAD0" author="Invrea" box="[781,916,654,677]" pageId="18" pageNumber="48" refString="Invrea, F. (1960) Descrizione di nuovi apterogynidi e mutillidi dell'Egitto con note su altre species della regione (Hymenoptera: Apterogynidae et Mutillidae). Memorie della Societa Entomologica Italiana, 38, 197 - 214, 1 fig." type="journal article" year="1960">Invrea 1960</bibRefCitation>
: 206
</treatmentCitation>
, Ψ;
<treatmentCitation id="D2631095FFB7710AFC0B889DFB4CEAD0" author="Suarez" box="[1022,1218,654,677]" page="925" pageId="18" pageNumber="19" year="1963">
<bibRefCitation id="37534B75FFB7710AFC0B889DFB04EAD0" author="Suarez" box="[1022,1162,654,677]" pageId="18" pageNumber="50" refString="Suarez, F. J. (1963) Hymenopteres Tiphiides, Apterogynides et mutillides recoltes par J. Mateu dans l'Ennedi et au Tchad. Bulletin de l'Institut fondamental d'Afrique Noire, Dakar. Serie A: Sciences Naturelles, 25 (3), 912 - 943, 10 figs." type="journal article" year="1963">Suárez 1963</bibRefCitation>
: 925
</treatmentCitation>
, Ψ;
<treatmentCitation id="D2631095FFB7710AFB01889DFE16EAB2" author="Lelej" page="49" pageId="18" pageNumber="19" year="2002">
<bibRefCitation id="37534B75FFB7710AFB01889DFEFFEAB2" author="Lelej" pageId="18" pageNumber="48" refString="Lelej, A. S. (2002) Catalogue of the Mutillidae (Hymenoptera) of the Palaearctic region. Dalnauka, Vladivostok. 171 pp." type="book" year="2002">Lelej 2002</bibRefCitation>
: 49
</treatmentCitation>
, Ψ.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB7710AFEFD8932FC6BEBC6" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="537D3684FFB7710AFEFD8932FE68EB4F" blockId="18.[264,1323,801,947]" box="[264,486,801,826]" pageId="18" pageNumber="19">
<emphasis id="61B6EA96FFB7710AFEFD8932FE68EB4F" box="[264,486,801,826]" italics="true" pageId="18" pageNumber="19">Material examined</emphasis>
</paragraph>
<paragraph id="537D3684FFB7710AFECD895AFC6BEBC6" blockId="18.[264,1323,801,947]" pageId="18" pageNumber="19">
<collectingCountry id="2BD57614FFB7710AFECD895AFE01EB16" box="[312,399,841,867]" name="Yemen" pageId="18" pageNumber="19">Yemen</collectingCountry>
: Wadi Surdud,
<date id="277C1044FFB7710AFDB2895AFC8BEB16" box="[583,773,841,867]" pageId="18" pageNumber="19" value="1991-12-28" valueMax="1991-12-29" valueMin="1991-12-28">28–29.XII.1991</date>
, AvH, 1 Ψ [
<collectionCode id="35D3AE41FFB7710AFC63895AFC78EB16" LSID="urn:lsid:biocol.org:col:34975" box="[918,1014,841,867]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="18" pageNumber="19">ZMAN</collectionCode>
]; Lahj,
<date id="277C1044FFB7710AFBA4895AFB65EB16" box="[1105,1259,841,867]" pageId="18" pageNumber="19" value="1999-11" valueMax="1999-12-31" valueMin="1999-11-00">XI–XII.1999</date>
, MT, AvH &amp; A. Sallam, 1 Ψ [
<collectionCode id="35D3AE41FFB7710AFDDF8962FDE1EBFE" box="[554,623,881,907]" pageId="18" pageNumber="19">IBSS</collectionCode>
];
<quantity id="943A9B61FFB7710AFD708962FD5FEBFE" box="[645,721,881,907]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="18" pageNumber="19" unit="km" value="12.0">12 km</quantity>
NW Manakhah,
<date id="277C1044FFB7710AFC558962FBFEEBFE" box="[928,1136,881,907]" pageId="18" pageNumber="19" value="2003-05-15" valueMax="2003-06-23" valueMin="2003-05-15">15.V–23.VI.2003</date>
, MT, AvH, 1 Ψ [
<collectionCode id="35D3AE41FFB7710AFEFA898AFEFEEBC6" LSID="urn:lsid:biocol.org:col:34992" box="[271,368,921,947]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="18" pageNumber="19">RMNH</collectionCode>
]; Aden, Mohur,
<date id="277C1044FFB7710AFDDB898AFD35EBC6" box="[558,699,921,947]" pageId="18" pageNumber="19" value="1983-08-09">9.VIII.1983</date>
, 1 Ψ, A. Mochi [
<collectionCode id="35D3AE41FFB7710AFC8B898AFC54EBC6" box="[894,986,921,947]" pageId="18" pageNumber="19">MRSN</collectionCode>
].
</paragraph>
</subSubSection>
<paragraph id="537D3684FFB7710AFEFD89FAFEDAEC26" pageId="18" pageNumber="19">
<table id="21C2C424FFB78EE7FEFD89FAFAA3EC26" box="[264,1325,1001,1107]" colsContinueFrom="14.[264,1323,1799,1905]" gridcols="10" gridrows="3" pageId="18" pageNumber="19">
<tr id="EDF234C6FFB78EE7FEFD89FAFAA3EC77" box="[264,1325,1001,1026]" gridrow="0" pageId="18" pageNumber="19" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="AE235DBAFFB78EE7FEFD89FAFE5EEC77" box="[264,464,1001,1026]" gridcol="0" gridrow="0" pageId="18" pageNumber="19">
<emphasis id="61B6EA96FFB7710AFEFD89FAFE19EC77" box="[264,407,1001,1026]" italics="true" pageId="18" pageNumber="19">Distribution</emphasis>
</th>
</tr>
<tr id="EDF234C6FFB78EE7FEFD8E02FAA3EC5E" box="[264,1325,1041,1067]" gridrow="1" pageId="18" pageNumber="19">
<th id="AE235DBAFFB78EE7FEFD8E02FE5EEC5E" box="[264,464,1041,1067]" gridcol="0" gridrow="1" pageId="18" pageNumber="19">Yemen (new</th>
<td id="AE235DBAFFB78EE7FE2E8E02FDB9EC5E" box="[475,567,1041,1067]" gridcol="1" gridrow="1" pageId="18" pageNumber="19">record),</td>
<td id="AE235DBAFFB78EE7FDB68E02FD06EC5E" box="[579,648,1041,1067]" gridcol="2" gridrow="1" pageId="18" pageNumber="19">Syria,</td>
<td id="AE235DBAFFB78EE7FD618E02FD65EC5E" box="[660,747,1041,1067]" gridcol="3" gridrow="1" pageId="18" pageNumber="19">Arabia,</td>
<td id="AE235DBAFFB78EE7FD028E02FCCAEC5E" box="[759,836,1041,1067]" gridcol="4" gridrow="1" pageId="18" pageNumber="19">Egypt,</td>
<td id="AE235DBAFFB78EE7FCBA8E02FC1AEC5E" box="[847,916,1041,1067]" gridcol="5" gridrow="1" pageId="18" pageNumber="19">Chad,</td>
<td id="AE235DBAFFB78EE7FC558E02FB85EC5E" box="[928,1035,1041,1067]" gridcol="6" gridrow="1" pageId="18" pageNumber="19">Ethiopia,</td>
<td id="AE235DBAFFB78EE7FBE28E02FBE0EC5E" box="[1047,1134,1041,1067]" gridcol="7" gridrow="1" pageId="18" pageNumber="19">Eritrea,</td>
<td id="AE235DBAFFB78EE7FB8F8E02FB52EC5E" box="[1146,1244,1041,1067]" gridcol="8" gridrow="1" pageId="18" pageNumber="19">Djibouti</td>
<td id="AE235DBAFFB78EE7FB128E02FAA3EC5E" box="[1255,1325,1041,1067]" gridcol="9" gridrow="1" pageId="18" pageNumber="19">(Lelej</td>
</tr>
<tr id="EDF234C6FFB78EE7FEFD8E2AFAA3EC26" box="[264,1325,1081,1107]" gridrow="2" pageId="18" pageNumber="19" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="AE235DBAFFB78EE7FEFD8E2AFE5EEC26" box="[264,464,1081,1107]" gridcol="0" gridrow="2" pageId="18" pageNumber="19">2002).</th>
</tr>
</table>
</paragraph>
<subSubSection id="1BD8650FFFB7710AFEFD8E9AFD46EDE6" pageId="18" pageNumber="19" type="discussion">
<paragraph id="537D3684FFB7710AFEFD8E9AFEE0ECD7" blockId="18.[264,1324,1161,1427]" box="[264,366,1161,1186]" pageId="18" pageNumber="19">
<emphasis id="61B6EA96FFB7710AFEFD8E9AFEE0ECD7" box="[264,366,1161,1186]" italics="true" pageId="18" pageNumber="19">Remarks</emphasis>
</paragraph>
<paragraph id="537D3684FFB7710AFECD8EA2FD46EDE6" blockId="18.[264,1324,1161,1427]" pageId="18" pageNumber="19">
The female of
<taxonomicName id="94C24D07FFB7710AFDF58EA2FC87ECBF" box="[512,777,1201,1226]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="fimbriata">
<emphasis id="61B6EA96FFB7710AFDF58EA2FC87ECBF" box="[512,777,1201,1226]" italics="true" pageId="18" pageNumber="19">Tropidotilla fimbriata</emphasis>
</taxonomicName>
has almost quadrate mesosoma as it was mentioned in the original description (
<bibRefCitation id="37534B75FFB7710AFD3B8ECAFCDCEC86" author="Klug" box="[718,850,1241,1267]" pageId="18" pageNumber="48" refString="Klug, J. C. F. (1829) Symbolae physicae seu icones et descriptiones insectorum quae ex itinere per Africam borealem et Asiam occidentalem Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg studio novae aut illustratae redierunt. Hymenoptera. Decas Prima [genera Mutilla et Apterogyna]. Venditur a Mittlero, Berolini. [15 - 21] + tab. 4, 5." type="journal volume" year="1829">Klug 1829</bibRefCitation>
). The other female species has elongate mesosoma (mesosoma length 1.3X its maximal width) and we associated it with males of
<taxonomicName id="94C24D07FFB7710AFEFD8F3AFEF2ED37" box="[264,380,1321,1346]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="milmili">
<emphasis id="61B6EA96FFB7710AFEFD8F3AFEF2ED37" box="[264,380,1321,1346]" italics="true" pageId="18" pageNumber="19">T. milmili</emphasis>
</taxonomicName>
that have a narrower mesosoma than
<taxonomicName id="94C24D07FFB7710AFCA28F3AFC54ED37" box="[855,986,1321,1346]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="61B6EA96FFB7710AFCA28F3AFC54ED37" box="[855,986,1321,1346]" italics="true" pageId="18" pageNumber="19">T. frontalis</emphasis>
</taxonomicName>
and
<taxonomicName id="94C24D07FFB7710AFBE98F3AFBF4ED37" box="[1052,1146,1321,1346]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bellii">
<emphasis id="61B6EA96FFB7710AFBE98F3AFBF4ED37" box="[1052,1146,1321,1346]" italics="true" pageId="18" pageNumber="19">T. bellii</emphasis>
</taxonomicName>
. Definitely,
<taxonomicName id="94C24D07FFB7710AFAE08F3AFEFBED1F" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="fimbriata">
<emphasis id="61B6EA96FFB7710AFAE08F3AFEFBED1F" italics="true" pageId="18" pageNumber="19">T. fimbriata</emphasis>
</taxonomicName>
is the opposite sex of
<taxonomicName id="94C24D07FFB7710AFD888F42FD72ED1F" box="[637,764,1361,1386]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="61B6EA96FFB7710AFD888F42FD72ED1F" box="[637,764,1361,1386]" italics="true" pageId="18" pageNumber="19">T. frontalis</emphasis>
</taxonomicName>
or
<taxonomicName id="94C24D07FFB7710AFCD18F42FCF3ED1F" box="[804,893,1361,1386]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bellii">
<emphasis id="61B6EA96FFB7710AFCD18F42FCF3ED1F" box="[804,893,1361,1386]" italics="true" pageId="18" pageNumber="19">T. bellii</emphasis>
</taxonomicName>
, but the synonymy will be proposed when the status of
<taxonomicName id="94C24D07FFB7710AFE118F6AFDB3EDE7" box="[484,573,1401,1426]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bellii">
<emphasis id="61B6EA96FFB7710AFE118F6AFDB3EDE7" box="[484,573,1401,1426]" italics="true" pageId="18" pageNumber="19">T. bellii</emphasis>
</taxonomicName>
is clarified.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:identifier id="FB642B00423F425C75437F3D01629F1F" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="46D9C07744CD0D5B7FE8D2213E8C647E" type="Zenodo-Dep">172671</mods:identifier>
</mods:mods>
<treatment id="DB6B8792FFB7710BFEFD8FDAFE47EE7B" ID-DOI="http://doi.org/10.5281/zenodo.6252959" ID-GBIF-Taxon="119344635" ID-Zenodo-Dep="6252959" LSID="urn:lsid:plazi:treatment:DB6B8792FFB7710BFEFD8FDAFE47EE7B" httpUri="http://treatment.plazi.org/id/DB6B8792FFB7710BFEFD8FDAFE47EE7B" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<subSubSection id="1BD8650FFFB7710AFEFD8FDAFD23ED96" box="[264,685,1481,1507]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="537D3684FFB7710AFEFD8FDAFD23ED96" blockId="18.[264,685,1481,1507]" box="[264,685,1481,1507]" pageId="18" pageNumber="19">
<heading id="083581E8FFB7710AFEFD8FDAFD23ED96" bold="true" box="[264,685,1481,1507]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<emphasis id="61B6EA96FFB7710AFEFD8FDAFD23ED96" bold="true" box="[264,685,1481,1507]" pageId="18" pageNumber="19">
<emphasis id="61B6EA96FFB7710AFEFD8FDAFE9AED96" bold="true" box="[264,276,1481,1507]" italics="true" pageId="18" pageNumber="19">*</emphasis>
<taxonomicName id="94C24D07FFB7710AFEE18FDAFD23ED96" authority="(Klug 1829)" authorityName="Klug" authorityYear="1829" baseAuthorityName="Klug" baseAuthorityYear="1829" box="[276,685,1481,1507]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="61B6EA96FFB7710AFEE18FDAFD9CED96" bold="true" box="[276,530,1481,1507]" italics="true" pageId="18" pageNumber="19">Tropidotilla frontalis</emphasis>
(
<bibRefCitation id="37534B75FFB7710AFDD78FDAFD2BED96" author="Klug" box="[546,677,1481,1507]" pageId="18" pageNumber="48" refString="Klug, J. C. F. (1829) Symbolae physicae seu icones et descriptiones insectorum quae ex itinere per Africam borealem et Asiam occidentalem Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg studio novae aut illustratae redierunt. Hymenoptera. Decas Prima [genera Mutilla et Apterogyna]. Venditur a Mittlero, Berolini. [15 - 21] + tab. 4, 5." type="journal volume" year="1829">Klug 1829</bibRefCitation>
)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB7710AFEFD8C07FB1AEE1E" pageId="18" pageNumber="19" type="reference_group">
<paragraph id="537D3684FFB7710AFEFD8C07FD4AEE3E" blockId="18.[264,1207,1555,1643]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="73D211AAFFB7710AFEFD8C07FD4AEE3E" pageId="18" pageNumber="19">
<taxonomicName id="94C24D07FFB7710AFEFD8C07FDBCEE5E" authority="Klug 1829" authorityName="Klug" authorityYear="1829" box="[264,562,1556,1579]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="61B6EA96FFB7710AFEFD8C07FE38EE5E" box="[264,438,1556,1579]" italics="true" pageId="18" pageNumber="19">Mutilla frontalis</emphasis>
<bibRefCitation id="37534B75FFB7710AFE488C07FDBCEE5E" author="Klug" box="[445,562,1556,1579]" pageId="18" pageNumber="48" refString="Klug, J. C. F. (1829) Symbolae physicae seu icones et descriptiones insectorum quae ex itinere per Africam borealem et Asiam occidentalem Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg studio novae aut illustratae redierunt. Hymenoptera. Decas Prima [genera Mutilla et Apterogyna]. Venditur a Mittlero, Berolini. [15 - 21] + tab. 4, 5." type="journal volume" year="1829">Klug 1829</bibRefCitation>
</taxonomicName>
: [15], tab. 4, fig. 2, ♂,
<typeStatus id="8C798826FFB7710AFCD18C07FC0FEE5E" box="[804,897,1556,1579]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
&quot;Angabe&quot; (
<collectingCountry id="2BD57614FFB7710AFC0A8C07FBCFEE5E" box="[1023,1089,1556,1579]" name="Egypt" pageId="18" pageNumber="19">Egypt</collectingCountry>
) [ZMHU].
<taxonomicName id="94C24D07FFB7710AFEFD8C27FD2FEE3E" authority="Andre 1910: 27" authorityName="Andre" authorityPageNumber="27" authorityYear="1910" box="[264,673,1588,1611]" class="Insecta" family="Mutillidae" genus="Ephutomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="61B6EA96FFB7710AFEFD8C27FE66EE3E" box="[264,488,1588,1611]" italics="true" pageId="18" pageNumber="19">Ephutomma frontalis</emphasis>
:
<treatmentCitation id="D2631095FFB7710AFE028C27FD2FEE3E" author="Andre" box="[503,673,1588,1611]" page="27" pageId="18" pageNumber="19" year="1910">
<bibRefCitation id="37534B75FFB7710AFE028C27FDF7EE3E" author="Andre" box="[503,633,1588,1611]" pageId="18" pageNumber="47" refString="Andre, E. (1910) Revision monographique des mutillides de l'Egypte. Memoires de la Societe Entomologique d'Egypte, 1 (2), 1 - 94, 3 plates." type="book chapter" year="1910">André 1910</bibRefCitation>
: 27
</treatmentCitation>
</taxonomicName>
, ♂.
</treatmentCitationGroup>
</paragraph>
<paragraph id="537D3684FFB7710AFEFD8C47FB1AEE1E" blockId="18.[264,1207,1555,1643]" box="[264,1172,1619,1643]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="73D211AAFFB7710AFEFD8C47FB1AEE1E" box="[264,1172,1619,1643]" pageId="18" pageNumber="19">
<taxonomicName id="94C24D07FFB7710AFEFD8C47FD4AEE1E" authority="Bischoff 1920: 160" authorityName="Bischoff" authorityPageNumber="160" authorityYear="1920" box="[264,708,1620,1643]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="61B6EA96FFB7710AFEFD8C47FE68EE1E" box="[264,486,1620,1643]" italics="true" pageId="18" pageNumber="19">Tropidotilla frontalis</emphasis>
: Bischoff 1920: 160
</taxonomicName>
, ♂;
<treatmentCitation id="D2631095FFB7710AFD1A8C47FC29EE1E" author="Invrea" box="[751,935,1620,1643]" page="206" pageId="18" pageNumber="19" year="1960">
<bibRefCitation id="37534B75FFB7710AFD1A8C47FCFDEE1E" author="Invrea" box="[751,883,1620,1643]" pageId="18" pageNumber="48" refString="Invrea, F. (1960) Descrizione di nuovi apterogynidi e mutillidi dell'Egitto con note su altre species della regione (Hymenoptera: Apterogynidae et Mutillidae). Memorie della Societa Entomologica Italiana, 38, 197 - 214, 1 fig." type="journal article" year="1960">Invrea 1960</bibRefCitation>
: 206
</treatmentCitation>
, ♂;
<treatmentCitation id="D2631095FFB7710AFC268C47FBFEEE1E" author="Lelej" box="[979,1136,1620,1643]" page="49" pageId="18" pageNumber="19" year="2002">
<bibRefCitation id="37534B75FFB7710AFC268C47FBC7EE1E" author="Lelej" box="[979,1097,1620,1643]" pageId="18" pageNumber="48" refString="Lelej, A. S. (2002) Catalogue of the Mutillidae (Hymenoptera) of the Palaearctic region. Dalnauka, Vladivostok. 171 pp." type="book" year="2002">Lelej 2002</bibRefCitation>
: 49
</treatmentCitation>
, ♂.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB7710BFEFD8C8FFC3FE98B" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="537D3684FFB7710AFEFD8C8FFE68EEC0" blockId="18.[264,1327,1692,1878]" box="[264,486,1692,1717]" pageId="18" pageNumber="19">
<emphasis id="61B6EA96FFB7710AFEFD8C8FFE68EEC0" box="[264,486,1692,1717]" italics="true" pageId="18" pageNumber="19">Material examined</emphasis>
</paragraph>
<paragraph id="537D3684FFB7710BFECD8CD7FC3FE98B" blockId="18.[264,1327,1692,1878]" lastBlockId="19.[264,1323,283,510]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<collectingCountry id="2BD57614FFB7710AFECD8CD7FE00EEAB" box="[312,398,1732,1758]" name="Yemen" pageId="18" pageNumber="19">Yemen</collectingCountry>
: Al Mahrah, Al Ghayda,
<date id="277C1044FFB7710AFD3D8CD7FCF2EEAB" box="[712,892,1732,1758]" pageId="18" pageNumber="19" value="1997-11-12" valueMax="1997-11-14" valueMin="1997-11-12">12–14.XI.1997</date>
, A. Sallam &amp; S. Ba Angood, 1 ɗ [
<collectionCode id="35D3AE41FFB7710AFEFA8CFFFEE1EF73" LSID="urn:lsid:biocol.org:col:34975" box="[271,367,1772,1798]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="18" pageNumber="19">ZMAN</collectionCode>
]; Al Kadan,
<date id="277C1044FFB7710AFDF78CFFFD0AEF73" box="[514,644,1772,1798]" pageId="18" pageNumber="19" value="1997-12-03">3.XII.1997</date>
–
<date id="277C1044FFB7710AFD608CFFFC80EF73" box="[661,782,1772,1798]" pageId="18" pageNumber="19" value="1998-02-17">17.II.1998</date>
, MT, AvH &amp; H.M. Naser, 1 ɗ[
<collectionCode id="35D3AE41FFB7710AFB8F8CFFFB54EF73" LSID="urn:lsid:biocol.org:col:34975" box="[1146,1242,1772,1798]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="18" pageNumber="19">ZMAN</collectionCode>
]; Lahj,
<date id="277C1044FFB7710AFEFD8D07FE48EF5B" box="[264,454,1812,1838]" pageId="18" pageNumber="19" value="2000-04-01" valueMax="2000-05-17" valueMin="2000-04-01">1.IV–17.V.2000</date>
, MT, AvH &amp; A. Sallam, 1 ɗ [
<collectionCode id="35D3AE41FFB7710AFCDD8D07FC06EF5B" LSID="urn:lsid:biocol.org:col:34975" box="[808,904,1812,1838]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="18" pageNumber="19">ZMAN</collectionCode>
];
<date id="277C1044FFB7710AFC6F8D07FBE5EF5B" box="[922,1131,1812,1838]" pageId="18" pageNumber="19" value="2000-05-17" valueMax="2000-06-15" valueMin="2000-05-17">17.V–15.VI.2000</date>
, MT, AvH &amp; A. Sallam, 6 ɗ [
<collectionCode id="35D3AE41FFB7710AFE518D2FFD8BEF23" LSID="urn:lsid:biocol.org:col:34992" box="[420,517,1852,1878]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="18" pageNumber="19">RMNH</collectionCode>
];
<date id="277C1044FFB7710AFDEC8D2FFD40EF23" box="[537,718,1852,1878]" pageId="18" pageNumber="19" value="2000-06-15" valueMax="2000-06-30" valueMin="2000-06-15">15–30.VI.2000</date>
, MT, AvH &amp; A. Sallam, 10 ɗ [
<collectionCode id="35D3AE41FFB7710AFBB68D2FFB2AEF23" LSID="urn:lsid:biocol.org:col:34992" box="[1091,1188,1852,1878]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="18" pageNumber="19">RMNH</collectionCode>
];
<date id="277C1044FFB7710AFB4C8D2FFAA8EF23" box="[1209,1318,1852,1878]" pageId="18" pageNumber="19" value="2000-07">VII.2000</date>
, MT, AvH &amp; A. Sallam, 7 ɗ [
<collectionCode id="35D3AE41FFB6710BFD938B0FFD49E943" LSID="urn:lsid:biocol.org:col:34992" box="[614,711,284,310]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="19" pageNumber="20">RMNH</collectionCode>
];
<date id="277C1044FFB6710BFD298B0FFCDCE943" box="[732,850,284,310]" pageId="19" pageNumber="20" value="2000-08">VIII.2000</date>
, MT, AvH &amp; A. Sallam, 5 ɗ [
<collectionCode id="35D3AE41FFB6710BFB4B8B0FFA91E943" LSID="urn:lsid:biocol.org:col:34992" box="[1214,1311,284,310]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="19" pageNumber="20">RMNH</collectionCode>
];
<date id="277C1044FFB6710BFEFD8B57FEE5E92B" box="[264,363,324,350]" pageId="19" pageNumber="20" value="2000-11">XI.2000</date>
, MT, AvH &amp; A. Sallam, 2 ɗ [
<collectionCode id="35D3AE41FFB6710BFD048B57FCDCE92B" LSID="urn:lsid:biocol.org:col:34992" box="[753,850,324,350]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="19" pageNumber="20">RMNH</collectionCode>
];
<date id="277C1044FFB6710BFC9F8B57FC36E92B" box="[874,952,324,350]" pageId="19" pageNumber="20" value="2001-01">I.2001</date>
, MT, AvH &amp; A. Sallam, 2 ɗ [
<collectionCode id="35D3AE41FFB6710BFEFA8B7FFEFEE9F3" LSID="urn:lsid:biocol.org:col:34992" box="[271,368,364,390]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="19" pageNumber="20">RMNH</collectionCode>
];
<date id="277C1044FFB6710BFE778B7FFE6AE9F3" box="[386,484,364,390]" pageId="19" pageNumber="20" value="2001-03">III.2001</date>
, MT, AvH &amp; A. Sallam, 1 ɗ [
<collectionCode id="35D3AE41FFB6710BFCC98B7FFC13E9F3" LSID="urn:lsid:biocol.org:col:34992" box="[828,925,364,390]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="19" pageNumber="20">RMNH</collectionCode>
];
<date id="277C1044FFB6710BFC458B7FFB81E9F3" box="[944,1039,364,390]" pageId="19" pageNumber="20" value="2001-04">IV.2001</date>
, MT, AvH &amp; A. Sallam, 5 ɗ [
<collectionCode id="35D3AE41FFB6710BFECA8B87FE2EE9DB" LSID="urn:lsid:biocol.org:col:34992" box="[319,416,404,430]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="19" pageNumber="20">RMNH</collectionCode>
];
<date id="277C1044FFB6710BFE468B87FD86E9DB" box="[435,520,404,430]" pageId="19" pageNumber="20" value="2001-05">V.2001</date>
, MT, AvH &amp; A. Sallam, 2 ɗ [
<collectionCode id="35D3AE41FFB6710BFC948B87FC4CE9DB" LSID="urn:lsid:biocol.org:col:34992" box="[865,962,404,430]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="19" pageNumber="20">RMNH</collectionCode>
];
<date id="277C1044FFB6710BFC218B87FBE1E9DB" box="[980,1135,404,430]" pageId="19" pageNumber="20" value="2001-07" valueMax="2001-09-31" valueMin="2001-07-00">VII–IX.2001</date>
, MT, AvH &amp; A. Sallam, 5 ɗ [
<collectionCode id="35D3AE41FFB6710BFE548BAFFD8CE9A3" LSID="urn:lsid:biocol.org:col:34992" box="[417,514,444,470]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="19" pageNumber="20">RMNH</collectionCode>
];
<date id="277C1044FFB6710BFDE18BAFFD58E9A3" box="[532,726,444,470]" pageId="19" pageNumber="20" value="2001-10-01" valueMax="2001-11-17" valueMin="2001-10-01">1.X–17.XI.2001</date>
, MT, AvH &amp; A. Sallam, 1 ɗ [
<collectionCode id="35D3AE41FFB6710BFBC18BAFFB1BE9A3" LSID="urn:lsid:biocol.org:col:34992" box="[1076,1173,444,470]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="19" pageNumber="20">RMNH</collectionCode>
];
<quantity id="943A9B61FFB6710BFB5D8BAFFB7CE9A3" box="[1192,1266,444,470]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="19" pageNumber="20" unit="km" value="12.0">12 km</quantity>
NW Manakhah,
<date id="277C1044FFB6710BFE678BF7FDF2E98B" box="[402,636,484,510]" pageId="19" pageNumber="20" value="2001-07-03" valueMax="2001-08-21" valueMin="2001-07-03">3.VII–21.VIII.2001</date>
, MT, AvH, 3 ɗ [
<collectionCode id="35D3AE41FFB6710BFCB18BF7FC2BE98B" LSID="urn:lsid:biocol.org:col:34992" box="[836,933,484,510]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="19" pageNumber="20">RMNH</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB6710BFEFD8827FD0AEA03" box="[264,644,564,630]" pageId="19" pageNumber="20" type="distribution">
<paragraph id="537D3684FFB6710BFEFD8827FE19EA38" blockId="19.[264,644,564,630]" box="[264,407,564,589]" pageId="19" pageNumber="20">
<emphasis id="61B6EA96FFB6710BFEFD8827FE19EA38" box="[264,407,564,589]" italics="true" pageId="19" pageNumber="20">Distribution</emphasis>
</paragraph>
<paragraph id="537D3684FFB6710BFECD884FFD0AEA03" blockId="19.[264,644,564,630]" box="[312,644,604,630]" pageId="19" pageNumber="20">
<collectingCountry id="2BD57614FFB6710BFECD884FFE04EA03" box="[312,394,604,630]" name="Yemen" pageId="19" pageNumber="20">Yemen</collectingCountry>
(new record),
<collectingCountry id="2BD57614FFB6710BFDC3884FFD0EEA03" box="[566,640,604,630]" name="Egypt" pageId="19" pageNumber="20">Egypt</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB6710BFEFD88BFFE47EE7B" pageId="19" pageNumber="20" type="discussion">
<paragraph id="537D3684FFB6710BFEFD88BFFEE0EAB0" blockId="19.[264,1324,684,1550]" box="[264,366,684,709]" pageId="19" pageNumber="20">
<emphasis id="61B6EA96FFB6710BFEFD88BFFEE0EAB0" box="[264,366,684,709]" italics="true" pageId="19" pageNumber="20">Remarks</emphasis>
</paragraph>
<paragraph id="537D3684FFB6710BFECD88C7FD67EC73" blockId="19.[264,1324,684,1550]" pageId="19" pageNumber="20">
This species is known only from the male. The head color in the specimens from
<collectingCountry id="2BD57614FFB6710BFEFD88EFFED4EB63" box="[264,346,764,790]" name="Yemen" pageId="19" pageNumber="20">Yemen</collectingCountry>
varies from wholly ferruginous­red with a weak darkened spot on frons to almost black with ferruginous­red spots between eye and antennal socket. Mesosoma also varies in color from ferruginous­red darkened ventrally to ferruginous­red with black mesoscutum and rarely darkened mesoscutellum and pronotum. In spite of the head and mesosoma coloration, the legs are always brown to piceous with lighter tarsi. Antennae castaneous, flagellomeres beneath ferruginous­red; flagellomere 1 beneath always lighter (yellowish­red) than other flagellomeres.
</paragraph>
<paragraph id="537D3684FFB6710BFECD8E07FAAEEC83" blockId="19.[264,1324,684,1550]" pageId="19" pageNumber="20">
It is quite possible that
<taxonomicName id="94C24D07FFB6710BFDAF8E07FC76EC5B" authority="Magretti 1906" authorityName="Magretti" authorityYear="1906" box="[602,1016,1044,1070]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bellii">
<emphasis id="61B6EA96FFB6710BFDAF8E07FCA2EC58" box="[602,812,1044,1069]" italics="true" pageId="19" pageNumber="20">Tropidotilla bellii</emphasis>
(Magretti 1906)
</taxonomicName>
, which is known only as male and has variable coloration of mesosoma, overlaps with
<taxonomicName id="94C24D07FFB6710BFBF68E2FFB09EC20" box="[1027,1159,1084,1109]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="61B6EA96FFB6710BFBF68E2FFB09EC20" box="[1027,1159,1084,1109]" italics="true" pageId="19" pageNumber="20">T. frontalis</emphasis>
</taxonomicName>
and that this black headed species is a color form of
<taxonomicName id="94C24D07FFB6710BFCD18E77FC23EC08" box="[804,941,1124,1149]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="61B6EA96FFB6710BFCD18E77FC23EC08" box="[804,941,1124,1149]" italics="true" pageId="19" pageNumber="20">T. frontalis</emphasis>
</taxonomicName>
, because other morphological characters like the posterolateral denticle of S2 and the absence of a carina on S1 are similar. In
<collectingCountry id="2BD57614FFB6710BFE798EA7FE50ECBB" box="[396,478,1204,1230]" name="Yemen" pageId="19" pageNumber="20">Yemen</collectingCountry>
both species are collected practically at the same localities and at the same time of the year. Body length of
<taxonomicName id="94C24D07FFB6710BFD3C8ECFFCC6EC80" box="[713,840,1244,1269]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="61B6EA96FFB6710BFD3C8ECFFCC6EC80" box="[713,840,1244,1269]" italics="true" pageId="19" pageNumber="20">T. frontalis</emphasis>
</taxonomicName>
<quantity id="943A9B61FFB6710BFCBA8ECFFC63EC83" box="[847,1005,1244,1270]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" metricValueMax="10.4" metricValueMin="5.2" pageId="19" pageNumber="20" unit="mm" value="7.8" valueMax="10.4" valueMin="5.2">5.2–10.4 mm</quantity>
,
<taxonomicName id="94C24D07FFB6710BFC0C8ECFFBDCEC80" box="[1017,1106,1244,1269]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bellii">
<emphasis id="61B6EA96FFB6710BFC0C8ECFFBDCEC80" box="[1017,1106,1244,1269]" italics="true" pageId="19" pageNumber="20">T. bellii</emphasis>
</taxonomicName>
— 6.0–11.0 mm.
</paragraph>
<paragraph id="537D3684FFB6710BFECD8F17FE47EE7B" blockId="19.[264,1324,684,1550]" pageId="19" pageNumber="20">
<bibRefCitation id="37534B75FFB6710BFECD8F17FE53ED6B" author="Invrea" box="[312,477,1284,1310]" pageId="19" pageNumber="48" refString="Invrea, F. (1960) Descrizione di nuovi apterogynidi e mutillidi dell'Egitto con note su altre species della regione (Hymenoptera: Apterogynidae et Mutillidae). Memorie della Societa Entomologica Italiana, 38, 197 - 214, 1 fig." type="journal article" year="1960">Invrea (1960)</bibRefCitation>
and
<bibRefCitation id="37534B75FFB6710BFDD48F17FD6EED6B" author="Bischoff" box="[545,736,1284,1310]" pageId="19" pageNumber="47" refString="Bischoff, H. (1920 - 1921) Monographie der Mutilliden Afrikas. Archiv fur Naturgeschichte, (1920) 86 A (1 / 3), 1 - 480; (1921) 86 A (4), 481 - 830 + plates 1 - 7." type="journal article" year="1921">Bischoff (1921)</bibRefCitation>
regarded
<taxonomicName id="94C24D07FFB6710BFCAA8F17FB57ED6B" authority="Andre 1902" authorityName="Andre" authorityYear="1902" box="[863,1241,1284,1310]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="pharaonis">
<emphasis id="61B6EA96FFB6710BFCAA8F17FBB7ED68" box="[863,1081,1284,1309]" italics="true" pageId="19" pageNumber="20">Mutilla pharaonis</emphasis>
<bibRefCitation id="37534B75FFB6710BFBB38F17FB57ED6B" author="Andre" box="[1094,1241,1284,1310]" pageId="19" pageNumber="47" refString="Andre, E. (1899 - 1903) Species de Hymenopteres d'Europe &amp; d'Algerie. 8. Les mutillides. V ve Dubosclard, Editeur, Paris. 478 pp. + pl. 1 - 15. (1899) 1 - 64, (1900) 65 - 144, (1901) 145 - 304, (1902) 305 - 384, (1903) 385 - 478 + pl. 1 - 15." type="book chapter" year="1902">André 1902</bibRefCitation>
</taxonomicName>
, male, described from
<collectingCountry id="2BD57614FFB6710BFE378F3FFD86ED33" box="[450,520,1324,1350]" name="Egypt" pageId="19" pageNumber="20">Egypt</collectingCountry>
(
<bibRefCitation id="37534B75FFB6710BFDED8F3FFD29ED33" author="Andre" box="[536,679,1324,1350]" pageId="19" pageNumber="47" refString="Andre, E. (1899 - 1903) Species de Hymenopteres d'Europe &amp; d'Algerie. 8. Les mutillides. V ve Dubosclard, Editeur, Paris. 478 pp. + pl. 1 - 15. (1899) 1 - 64, (1900) 65 - 144, (1901) 145 - 304, (1902) 305 - 384, (1903) 385 - 478 + pl. 1 - 15." type="book chapter" year="1902">André 1902</bibRefCitation>
) as a probable synonym of
<taxonomicName id="94C24D07FFB6710BFC078F3FFBFFED30" box="[1010,1137,1324,1349]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="61B6EA96FFB6710BFC078F3FFBFFED30" box="[1010,1137,1324,1349]" italics="true" pageId="19" pageNumber="20">T. frontalis</emphasis>
</taxonomicName>
or
<taxonomicName id="94C24D07FFB6710BFB6E8F3FFB7AED30" box="[1179,1268,1324,1349]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bellii">
<emphasis id="61B6EA96FFB6710BFB6E8F3FFB7AED30" box="[1179,1268,1324,1349]" italics="true" pageId="19" pageNumber="20">T. bellii</emphasis>
</taxonomicName>
. We have not possibility to study the
<typeStatus id="8C798826FFB6710BFD638F47FD72ED1B" box="[662,764,1364,1390]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
of
<taxonomicName id="94C24D07FFB6710BFCDC8F47FC44ED18" box="[809,970,1364,1389]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="pharaonis">
<emphasis id="61B6EA96FFB6710BFCDC8F47FC44ED18" box="[809,970,1364,1389]" italics="true" pageId="19" pageNumber="20">M. pharaonis</emphasis>
</taxonomicName>
but according to the original description this species has antennae and legs ferruginous­red that never observed in
<taxonomicName id="94C24D07FFB6710BFAE08F6FFEE4EDC8" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="61B6EA96FFB6710BFAE08F6FFEE4EDC8" italics="true" pageId="19" pageNumber="20">T. frontalis</emphasis>
</taxonomicName>
and
<taxonomicName id="94C24D07FFB6710BFE598FB7FD87EDC8" box="[428,521,1444,1469]" class="Insecta" family="Mutillidae" genus="Tropidotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bellii">
<emphasis id="61B6EA96FFB6710BFE598FB7FD87EDC8" box="[428,521,1444,1469]" italics="true" pageId="19" pageNumber="20">T. bellii</emphasis>
</taxonomicName>
. We support here the opinion of
<bibRefCitation id="37534B75FFB6710BFC5C8FB7FBC2EDCB" author="Andre" box="[937,1100,1444,1470]" pageId="19" pageNumber="47" refString="Andre, E. (1899 - 1903) Species de Hymenopteres d'Europe &amp; d'Algerie. 8. Les mutillides. V ve Dubosclard, Editeur, Paris. 478 pp. + pl. 1 - 15. (1899) 1 - 64, (1900) 65 - 144, (1901) 145 - 304, (1902) 305 - 384, (1903) 385 - 478 + pl. 1 - 15." type="book chapter" year="1902">André (1902)</bibRefCitation>
who regarded his species related to
<taxonomicName id="94C24D07FFB6710BFE138FDFFDC8ED90" box="[486,582,1484,1509]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="barbara">
<emphasis id="61B6EA96FFB6710BFE138FDFFDC8ED90" box="[486,582,1484,1509]" italics="true" pageId="19" pageNumber="20">barbara</emphasis>
</taxonomicName>
­group [currently the genus
<taxonomicName id="94C24D07FFB6710BFC6B8FDFFB03ED93" authority="Costa 1858" authorityName="Costa" authorityYear="1858" box="[926,1165,1484,1510]" class="Insecta" family="Mutillidae" genus="Ronisia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="61B6EA96FFB6710BFC6B8FDFFC76ED90" box="[926,1016,1484,1509]" italics="true" pageId="19" pageNumber="20">Ronisia</emphasis>
Costa 1858
</taxonomicName>
(=
<taxonomicName id="94C24D07FFB6710BFB478FDFFE3BEE7B" authority="Bischoff 1920" authorityName="Bischoff" authorityYear="1920" class="Insecta" family="Mutillidae" genus="Pycnotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="61B6EA96FFB6710BFB478FDFFAA4ED90" box="[1202,1322,1484,1509]" italics="true" pageId="19" pageNumber="20">Pycnotilla</emphasis>
Bischoff 1920
</taxonomicName>
)].
</paragraph>
</subSubSection>
</treatment>
</document>

140
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<subSubSection id="1BD8650FFFB87105FEFD898FFEE0EBAC" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="537D3684FFB87105FEFD898FFC9FEBC3" blockId="29.[264,785,924,985]" box="[264,785,924,950]" pageId="29" pageNumber="30">
<heading id="083581E8FFB87105FEFD898FFC9FEBC3" bold="true" box="[264,785,924,950]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
<emphasis id="61B6EA96FFB87105FEFD898FFC9FEBC3" bold="true" box="[264,785,924,950]" pageId="29" pageNumber="30">
<taxonomicName id="94C24D07FFB87105FEFD898FFD29EBC3" authority="Lelej" authorityName="Lelej" box="[264,679,924,950]" class="Insecta" family="Mutillidae" genus="Rasnitsynitilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="schmideggeri" status="sp. nov.">
<emphasis id="61B6EA96FFB87105FEFD898FFDD0EBC3" bold="true" box="[264,606,924,950]" italics="true" pageId="29" pageNumber="30">Rasnitsynitilla schmideggeri</emphasis>
Lelej
</taxonomicName>
,
<taxonomicNameLabel id="7A8557EDFFB87105FD41898FFC9FEBC3" box="[692,785,924,950]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="537D3684FFB87105FEFD89ACFEE0EBAC" blockId="29.[264,785,924,985]" box="[264,366,959,985]" pageId="29" pageNumber="30">(Fig. 21)</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB87105FEFD8E1CFC3FEC24" box="[264,945,1039,1105]" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="537D3684FFB87105FEFD8E1CFEF1EC5D" blockId="29.[264,945,1039,1105]" box="[264,383,1039,1064]" pageId="29" pageNumber="30">
<emphasis id="61B6EA96FFB87105FEFD8E1CFEF1EC5D" box="[264,383,1039,1064]" italics="true" pageId="29" pageNumber="30">Diagnosis</emphasis>
</paragraph>
<paragraph id="537D3684FFB87105FECD8E24FC3FEC24" blockId="29.[264,945,1039,1105]" box="[312,945,1079,1105]" pageId="29" pageNumber="30">Diagnostic characters are mentioned in the key below.</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB87105FEFD8E94FDB9EF54" pageId="29" pageNumber="30" type="description">
<paragraph id="537D3684FFB87105FEFD8E94FE1CECD5" blockId="29.[264,1325,1159,1825]" box="[264,402,1159,1184]" pageId="29" pageNumber="30">
<emphasis id="61B6EA96FFB87105FEFD8E94FE1CECD5" box="[264,402,1159,1184]" italics="true" pageId="29" pageNumber="30">Description</emphasis>
</paragraph>
<paragraph id="537D3684FFB87105FECD8EBCFDC3EDCC" blockId="29.[264,1325,1159,1825]" pageId="29" pageNumber="30">MALE. Body length 8.0 mm. Relative width of head and thorax including tegulae 4.5: 5.3. POD: OOD 0.5X. Relation of pedicel and first three flagellomeres 0.5: 1.1: 1.5: 1.5. Pronotum anteriorly with rounded humeri. Forewing with small pterostigma, venation as in Fig. 21. Tegula densely punctate. Propodeum reticulate, dorsally with median triangular cell. Pronotum dorsally, mesoscutum, and scutellum densely punctate. T1 width 2.0X T1 length. S1 carinate. T2 and S2 densely punctate, punctures larger on S2. T7 with glabrous area in apical half.</paragraph>
<paragraph id="537D3684FFB87105FECD8FD4FA97EE8C" blockId="29.[264,1325,1159,1825]" pageId="29" pageNumber="30">Head, mesosoma, and segment 1 of metasoma, and legs ferruginous­red. Mandible ferruginous­red with dark teeth. Palps ferruginous­red. Scape and pedicel ferruginous­red, flagellomeres black, castaneous beneath. Tegulae concolorous with mesosoma. Forewing hyaline, slightly infuscate beyond of cells. Head, mesosoma, and legs with sparse erect and recumbent yellowish setae. T1 posteriorly with fringe of whitish setae. T2 posteriorly and T3–T4 with bands of dense white setae. T5 and T7 with sparse whitish setae mixed with black ones. T6 with black setae. Lateral felt line of T2 and S2 yellowish. Metasoma ventrally with sparse yellowish setae, S2–S6 with weak apical fringes of yellowish setae.</paragraph>
<paragraph id="537D3684FFB87105FECD8D14FDB9EF54" blockId="29.[264,1325,1159,1825]" box="[312,567,1799,1825]" pageId="29" pageNumber="30">FEMALE. Unknown.</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFBB7106FEFD8B0FFC73E92B" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="537D3684FFBB7106FEFD8B0FFE27E940" blockId="30.[264,1021,284,350]" box="[264,425,284,309]" pageId="30" pageNumber="31">
<emphasis id="61B6EA96FFBB7106FEFD8B0FFE27E940" box="[264,425,284,309]" italics="true" pageId="30" pageNumber="31">
<typeStatus id="8C798826FFBB7106FEFD8B0FFEB1E940" box="[264,319,284,309]" pageId="30" pageNumber="31">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="537D3684FFBB7106FECD8B57FC73E92B" blockId="30.[264,1021,284,350]" box="[312,1021,323,350]" pageId="30" pageNumber="31">
<typeStatus id="8C798826FFBB7106FECD8B57FE2BE92B" box="[312,421,324,350]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
ɗ,
<collectingCountry id="2BD57614FFBB7106FE3B8B57FDADE92B" box="[462,547,324,350]" name="Yemen" pageId="30" pageNumber="31">Yemen</collectingCountry>
, Al Kowd,
<date id="277C1044FFBB7106FD5C8B57FC86E92B" box="[681,776,324,350]" pageId="30" pageNumber="31" value="1993-04">IV.1993</date>
, MT, AvH [
<collectionCode id="35D3AE41FFBB7106FC678B57FC7CE92B" LSID="urn:lsid:biocol.org:col:34975" box="[914,1010,324,350]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="30" pageNumber="31">ZMAN</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFBB7106FEFD8B87FE03E9A3" box="[264,407,404,470]" pageId="30" pageNumber="31" type="distribution">
<paragraph id="537D3684FFBB7106FEFD8B87FE19E9D8" blockId="30.[264,407,404,470]" box="[264,407,404,429]" pageId="30" pageNumber="31">
<emphasis id="61B6EA96FFBB7106FEFD8B87FE19E9D8" box="[264,407,404,429]" italics="true" pageId="30" pageNumber="31">Distribution</emphasis>
</paragraph>
<paragraph id="537D3684FFBB7106FECD8BAFFE03E9A3" blockId="30.[264,407,404,470]" box="[312,397,444,470]" pageId="30" pageNumber="31">
<collectingCountry id="2BD57614FFBB7106FECD8BAFFE09E9A3" box="[312,391,444,470]" name="Yemen" pageId="30" pageNumber="31">Yemen</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFBB7106FEFD881FFD4CECBA" pageId="30" pageNumber="31" type="etymology">
<paragraph id="537D3684FFBB7106FEFD881FFE0BEA50" blockId="30.[264,1323,524,630]" box="[264,389,524,549]" pageId="30" pageNumber="31">
<emphasis id="61B6EA96FFBB7106FEFD881FFE0BEA50" box="[264,389,524,549]" italics="true" pageId="30" pageNumber="31">Etymology</emphasis>
</paragraph>
<paragraph id="537D3684FFBB7106FECD8827FED9EA03" blockId="30.[264,1323,524,630]" pageId="30" pageNumber="31">
This species is dedicated to Christian Schmid­Egger (
<collectingCountry id="2BD57614FFBB7106FC478827FBABEA3B" box="[946,1061,564,590]" name="Germany" pageId="30" pageNumber="31">Germany</collectingCountry>
), expert in Palaearctic wasps.
</paragraph>
<paragraph id="537D3684FFBB7106FEFD8E43FD4CECBA" blockId="30.[264,1323,1104,1231]" pageId="30" pageNumber="31">
<emphasis id="61B6EA96FFBB7106FEFD8E43FE3CEC1D" bold="true" box="[264,434,1104,1128]" pageId="30" pageNumber="31">Figures 17–21.</emphasis>
17–18.
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<emphasis id="61B6EA96FFBB7106FDE78E42FCB7EC1D" box="[530,825,1105,1128]" italics="true" pageId="30" pageNumber="31">Rasnitsynitilla brachyptera</emphasis>
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, male,
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. 17. Forewing; 18. Pedicel and flagellomeres
<date id="277C1044FFBB7106FDDD8E60FD27ECFF" box="[552,681,1139,1162]" pageId="30" pageNumber="31" value="1919-01-20" valueMax="1919-03-20" valueMin="1919-01-20">1–3. 19–20.</date>
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<emphasis id="61B6EA96FFBB7106FD448E60FC92ECFF" box="[689,796,1139,1162]" italics="true" pageId="30" pageNumber="31">R. invreai</emphasis>
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,
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. 19. Forewing; 20. Mandible, lateral view. 21.
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<emphasis id="61B6EA96FFBB7106FE498E85FDE4ECD8" box="[444,618,1174,1197]" italics="true" pageId="30" pageNumber="31">R. schmideggeri</emphasis>
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<taxonomicNameLabel id="7A8557EDFFBB7106FD878E85FD46ECD8" box="[626,712,1174,1197]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
, male,
<typeStatus id="8C798826FFBB7106FCE38E85FCF9ECD8" box="[790,887,1174,1197]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
. Forewing. Scale
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for 17 and 19,
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for 18 and 20, and
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for 21.
</paragraph>
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112
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<emphasis id="61B6EA96FFB97104FEFD8EA7FD9CECBB" bold="true" box="[264,530,1204,1230]" italics="true" pageId="28" pageNumber="29">Rasnitsynitilla invreai</emphasis>
Lelej
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<paragraph id="537D3684FFB97104FEFD8EC4FE10EC84" blockId="28.[264,709,1204,1265]" box="[264,414,1239,1265]" pageId="28" pageNumber="29">(Figs 19, 20)</paragraph>
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<emphasis id="61B6EA96FFB97104FEFD8F34FEF1ED35" box="[264,383,1319,1344]" italics="true" pageId="28" pageNumber="29">Diagnosis</emphasis>
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<emphasis id="61B6EA96FFB97104FEFD8F8CFE1CEDCD" box="[264,402,1439,1464]" italics="true" pageId="28" pageNumber="29">Description</emphasis>
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<paragraph id="537D3684FFB97104FECD8FD4FB44EEA4" blockId="28.[264,1325,1439,1905]" pageId="28" pageNumber="29">MALE. Body length 4.0 mm. Relative width of head and thorax including tegulae 4.5: 5.3. POD: OOD 1.5X. Flagellomeres short, relation of pedicel and first three flagellomeres 0.3: 0.5: 0.9: 0.9. Pronotum anteriorly with rounded humeri. Forewing with small chitinized pterostigma, and very short radial cell, venation as in Fig. 19. Tegula glabrous. Propodeum reticulate, dorsally with median triangular cell. Pronotum dorsally, mesoscutum, and scutellum with dense shallow punctures. T1 width 1.1X T1 length. S1 carinate. T2 and S2 densely punctate, punctures larger on S2. T7 mostly glabrous.</paragraph>
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ɗ,
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, Mayfa’ah,
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.
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<emphasis id="61B6EA96FFB87105FEFD88C7FE0BEA98" box="[264,389,724,749]" italics="true" pageId="29" pageNumber="30">Etymology</emphasis>
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This species is dedicated to Fabio Invrea (
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and
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<paragraph id="537D3684FFBD7100FEFD8C7DFD44EEFD" blockId="24.[264,714,1646,1707]" box="[264,714,1646,1672]" pageId="24" pageNumber="25">
<heading id="083581E8FFBD7100FEFD8C7DFD44EEFD" bold="true" box="[264,714,1646,1672]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
<emphasis id="61B6EA96FFBD7100FEFD8C7DFD44EEFD" bold="true" box="[264,714,1646,1672]" pageId="24" pageNumber="25">
<taxonomicName id="94C24D07FFBD7100FEFD8C7DFDD1EEFD" authority="Lelej" authorityName="Lelej" box="[264,607,1646,1672]" class="Insecta" family="Mutillidae" genus="Paglianotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="hogenesi" status="sp. nov.">
<emphasis id="61B6EA96FFBD7100FEFD8C7DFD99EEFD" bold="true" box="[264,535,1646,1672]" italics="true" pageId="24" pageNumber="25">Paglianotilla hogenesi</emphasis>
Lelej
</taxonomicName>
,
<taxonomicNameLabel id="7A8557EDFFBD7100FD998C7DFD44EEFD" box="[620,714,1646,1672]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="537D3684FFBD7100FEFD8C82FE11EEDE" blockId="24.[264,714,1646,1707]" box="[264,415,1681,1707]" pageId="24" pageNumber="25">
(
<figureCitation id="CBF92A01FFBD7100FEE58C82FE18EEDE" box="[272,406,1681,1707]" captionStart="FIGURES 11 – 16. 11" captionStartId="26.[264,383,923,947]" captionTargetBox="[549,1042,566,897]" captionTargetId="figure@26.[538,1048,566,898]" captionTargetPageId="26" captionText="FIGURES 11 – 16. 11. Dentilla ehrenbergi sp. nov., male, holotype. Genitalia, left part, inner view. 12 – 16. Paglianotilla hogenesi sp. nov., male, paratype. 12. Metasoma, dorsal view; 13. Pedicel and flagellomeres 1 – 3; 14. Mandible, dorsal view; 15. Genitalia, ventral view; 16. Penial valves, ventral view. Scale = 1 mm for 12 and 0.5 mm for 11, 13, and 15." httpUri="https://zenodo.org/record/172674/files/figure.png" pageId="24" pageNumber="25">Figs 12–16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFBD7100FEFD8CCFFDB7EF1B" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="537D3684FFBD7100FEFD8CCFFEF1EE80" blockId="24.[264,1323,1756,1902]" box="[264,383,1756,1781]" pageId="24" pageNumber="25">
<emphasis id="61B6EA96FFBD7100FEFD8CCFFEF1EE80" box="[264,383,1756,1781]" italics="true" pageId="24" pageNumber="25">Diagnosis</emphasis>
</paragraph>
<paragraph id="537D3684FFBD7100FECD8D17FDB7EF1B" blockId="24.[264,1323,1756,1902]" pageId="24" pageNumber="25">
The male of the new species can be easily separated from that of the undescribed species from
<collectingCountry id="2BD57614FFBD7100FE508D3FFDB2EF33" box="[421,572,1836,1862]" name="South Africa" pageId="24" pageNumber="25">South Africa</collectingCountry>
in having a central silvery setal spot on T2 and in lacking large propleural tooth ventrally.
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFBC7101FEFD8B0FFDB9EC23" pageId="25" pageNumber="26" type="description">
<paragraph id="537D3684FFBC7101FEFD8B0FFE1CE940" blockId="25.[264,1323,284,1110]" box="[264,402,284,309]" pageId="25" pageNumber="26">
<emphasis id="61B6EA96FFBC7101FEFD8B0FFE1CE940" box="[264,402,284,309]" italics="true" pageId="25" pageNumber="26">Description</emphasis>
</paragraph>
<paragraph id="537D3684FFBC7101FECD8B57FC9BEAEB" blockId="25.[264,1323,284,1110]" pageId="25" pageNumber="26">
MALE. Body length
<quantity id="943A9B61FFBC7101FDC58B57FD4EE92B" box="[560,704,324,350]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" metricValueMax="8.8" metricValueMin="4.8" pageId="25" pageNumber="26" unit="mm" value="6.8" valueMax="8.8" valueMin="4.8">4.8–8.8 mm</quantity>
. Relative width of head and thorax including tegulae 5.3: 6.0. POD: OOD 0.75–1.0X. Frons with strong antennal scrobes. Prementum not tuberculate. Hypostomal bridge without projection. Relation of pedicel and first three flagellomeres in lateral view 0.6: 0.7: 1.7: 1.8, in dorsal view 0.6: 1.2: 1.6: 1.7. Pronotum anteriorly straight. Tegula strongly sculptured. Propodeum reticulate, dorsally with large median triangular cell. Pronotum dorsally, mesoscutum and scutellum densely punctate. Carina on S1 with sharp tooth posteriorly. T2 and S2 densely punctate, punctures larger on S2. T7 densely punctate throughout. Hypopygium weakly bi­emarginated. Genitalia and penial valves as in
<figureCitation id="CBF92A01FFBC7101FD7E8897FC9FEAEB" box="[651,785,644,670]" captionStart="FIGURES 11 – 16. 11" captionStartId="26.[264,383,923,947]" captionTargetBox="[549,1042,566,897]" captionTargetId="figure@26.[538,1048,566,898]" captionTargetPageId="26" captionText="FIGURES 11 – 16. 11. Dentilla ehrenbergi sp. nov., male, holotype. Genitalia, left part, inner view. 12 – 16. Paglianotilla hogenesi sp. nov., male, paratype. 12. Metasoma, dorsal view; 13. Pedicel and flagellomeres 1 – 3; 14. Mandible, dorsal view; 15. Genitalia, ventral view; 16. Penial valves, ventral view. Scale = 1 mm for 12 and 0.5 mm for 11, 13, and 15." httpUri="https://zenodo.org/record/172674/files/figure.png" pageId="25" pageNumber="26">Figs 15, 16</figureCitation>
.
</paragraph>
<paragraph id="537D3684FFBC7101FECD88BFFE49EC5B" blockId="25.[264,1323,284,1110]" pageId="25" pageNumber="26">
Head, mesosoma, and metasoma black. Mandible black, reddish preapically. Antenna black, brown beneath. Tegulae brownish­red. Legs brown to reddish­brown. Forewing weakly infuscated. Frons, antennal scrobes, clypeus, and genae with dense silvery setae, pronotum with sparser silvery setae. Mesopleura with dense silvery setae. Mesoscutum with black setae. Tegulae with whitish setae. Scutellum with sparse erect whitish setae. Tegulae posteriorly, axillae, metanotum and allied narrow part of propodeum with dense short recumbent silvery setae. T1–T2 posteriorly and whole of T3 with bands of silvery setae narrowed on T3. T2 with central large spot of silvery setae (
<figureCitation id="CBF92A01FFBC7101FBFC89D7FBEFEBAB" box="[1033,1121,964,990]" captionStart="FIGURES 11 – 16. 11" captionStartId="26.[264,383,923,947]" captionTargetBox="[549,1042,566,897]" captionTargetId="figure@26.[538,1048,566,898]" captionTargetPageId="26" captionText="FIGURES 11 – 16. 11. Dentilla ehrenbergi sp. nov., male, holotype. Genitalia, left part, inner view. 12 – 16. Paglianotilla hogenesi sp. nov., male, paratype. 12. Metasoma, dorsal view; 13. Pedicel and flagellomeres 1 – 3; 14. Mandible, dorsal view; 15. Genitalia, ventral view; 16. Penial valves, ventral view. Scale = 1 mm for 12 and 0.5 mm for 11, 13, and 15." httpUri="https://zenodo.org/record/172674/files/figure.png" pageId="25" pageNumber="26">Fig. 12</figureCitation>
). Lateral felt line of T2 yellowish. Metasoma beneath with sparse whitish setae, S2–S6 with apical fringes of whitish setae.
</paragraph>
<paragraph id="537D3684FFBC7101FECD8E2FFDB9EC23" blockId="25.[264,1323,284,1110]" box="[312,567,1084,1110]" pageId="25" pageNumber="26">FEMALE. Unknown.</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFBC7101FEFD8E9FFDA6EE8B" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="537D3684FFBC7101FEFD8E9FFE27ECD0" blockId="25.[264,1323,1164,1790]" box="[264,425,1164,1189]" pageId="25" pageNumber="26">
<emphasis id="61B6EA96FFBC7101FEFD8E9FFE27ECD0" box="[264,425,1164,1189]" italics="true" pageId="25" pageNumber="26">
<typeStatus id="8C798826FFBC7101FEFD8E9FFEB1ECD0" box="[264,319,1164,1189]" pageId="25" pageNumber="26">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="537D3684FFBC7101FECD8EA7FDA6EE8B" blockId="25.[264,1323,1164,1790]" pageId="25" pageNumber="26">
<typeStatus id="8C798826FFBC7101FECD8EA7FE2BECBB" box="[312,421,1204,1230]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
ɗ,
<collectingCountry id="2BD57614FFBC7101FE208EA7FDA4ECBB" box="[469,554,1204,1230]" name="Yemen" pageId="25" pageNumber="26">Yemen</collectingCountry>
,
<quantity id="943A9B61FFBC7101FDCC8EA7FD08ECBB" box="[569,646,1204,1230]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="25" pageNumber="26" unit="km" value="12.0">12 km</quantity>
NW Manakhah,
<date id="277C1044FFBC7101FCAF8EA7FBA5ECBB" box="[858,1067,1204,1230]" pageId="25" pageNumber="26" value="2003-09-15" valueMax="2003-10-22" valueMin="2003-09-15">15.IX–22.X.2003</date>
, MT, AvH [
<collectionCode id="35D3AE41FFBC7101FB4A8EA7FAAEECBB" LSID="urn:lsid:biocol.org:col:34992" box="[1215,1312,1204,1230]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="25" pageNumber="26">RMNH</collectionCode>
].
<typeStatus id="8C798826FFBC7101FEFD8ECFFEF0EC83" box="[264,382,1244,1270]" pageId="25" pageNumber="26" type="paratype">Paratypes</typeStatus>
. 1 ɗ with the same label as
<typeStatus id="8C798826FFBC7101FD388ECFFCBAEC83" box="[717,820,1244,1270]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
[
<collectionCode id="35D3AE41FFBC7101FCB68ECFFC2AEC83" LSID="urn:lsid:biocol.org:col:34992" box="[835,932,1244,1270]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="25" pageNumber="26">RMNH</collectionCode>
];
<collectingCountry id="2BD57614FFBC7101FC4C8ECFFB80EC83" box="[953,1038,1244,1270]" name="Yemen" pageId="25" pageNumber="26">Yemen</collectingCountry>
,
<quantity id="943A9B61FFBC7101FBEF8ECFFBE8EC83" box="[1050,1126,1244,1270]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="25" pageNumber="26" unit="km" value="12.0">12 km</quantity>
NW Manakhah,
<date id="277C1044FFBC7101FEFD8F17FE7FED6B" box="[264,497,1284,1310]" pageId="25" pageNumber="26" value="2001-07-03" valueMax="2001-08-21" valueMin="2001-07-03">3.VII–21.VIII.2001</date>
, MT, AvH, 10 ɗ [
<collectionCode id="35D3AE41FFBC7101FD398F17FCA4ED6B" LSID="urn:lsid:biocol.org:col:34992" box="[716,810,1284,1310]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="25" pageNumber="26">RMNH</collectionCode>
,
<collectionCode id="35D3AE41FFBC7101FCCD8F17FCF5ED6B" box="[824,891,1284,1310]" pageId="25" pageNumber="26">IBSS</collectionCode>
];
<date id="277C1044FFBC7101FC648F17FBEEED6B" box="[913,1120,1284,1310]" pageId="25" pageNumber="26" value="2003-05-15" valueMax="2003-06-23" valueMin="2003-05-15">15.V–23.VI.2003</date>
, MT, AvH, 11 ɗ [
<collectionCode id="35D3AE41FFBC7101FEFA8F3FFEE3ED33" LSID="urn:lsid:biocol.org:col:34992" box="[271,365,1324,1350]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="25" pageNumber="26">RMNH</collectionCode>
,
<collectionCode id="35D3AE41FFBC7101FE8E8F3FFE30ED33" box="[379,446,1324,1350]" pageId="25" pageNumber="26">IBSS</collectionCode>
];
<date id="277C1044FFBC7101FE218F3FFD3DED33" box="[468,691,1324,1350]" pageId="25" pageNumber="26" value="2003-06-24" valueMax="2003-08-04" valueMin="2003-06-24">24.VI–4.VIII.2003</date>
, MT, AvH, 3 ɗ [
<collectionCode id="35D3AE41FFBC7101FC748F3FFC51ED33" LSID="urn:lsid:biocol.org:col:34992" box="[897,991,1324,1350]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="25" pageNumber="26">RMNH</collectionCode>
,
<collectionCode id="35D3AE41FFBC7101FC188F3FFBBEED33" box="[1005,1072,1324,1350]" pageId="25" pageNumber="26">IBSS</collectionCode>
];
<date id="277C1044FFBC7101FBB28F3FFAA8ED33" box="[1095,1318,1324,1350]" pageId="25" pageNumber="26" value="2003-08-04" valueMax="2003-09-15" valueMin="2003-08-04">4.VIII–15.IX.2003</date>
, MT, AvH, 2 ɗ [
<collectionCode id="35D3AE41FFBC7101FE3E8F47FDA2ED1B" LSID="urn:lsid:biocol.org:col:34992" box="[459,556,1364,1390]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="25" pageNumber="26">RMNH</collectionCode>
]; Lahj,
<date id="277C1044FFBC7101FD7F8F47FCC6ED1B" box="[650,840,1364,1390]" pageId="25" pageNumber="26" value="2000-04-01" valueMax="2000-05-17" valueMin="2000-04-01">1.IV–17.V.2000</date>
, MT, AvH &amp; A. Sallam, 2 ɗ [
<collectionCode id="35D3AE41FFBC7101FB358F47FA91ED1B" LSID="urn:lsid:biocol.org:col:34975" box="[1216,1311,1364,1390]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="25" pageNumber="26">ZMAN</collectionCode>
];
<date id="277C1044FFBC7101FEFD8F6FFE52EDE3" box="[264,476,1404,1430]" pageId="25" pageNumber="26" value="2000-05-17" valueMax="2000-06-15" valueMin="2000-05-17">17.V.–15.VI.2000</date>
, MT, AvH &amp; A. Sallam, 8 ɗ [
<collectionCode id="35D3AE41FFBC7101FCB48F6FFC2CEDE3" LSID="urn:lsid:biocol.org:col:34992" box="[833,930,1404,1430]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="25" pageNumber="26">RMNH</collectionCode>
];
<date id="277C1044FFBC7101FC408F6FFBE7EDE3" box="[949,1129,1404,1430]" pageId="25" pageNumber="26" value="2000-06-15" valueMax="2000-06-30" valueMin="2000-06-15">15–30.VI.2000</date>
, MT, AvH &amp; A. Sallam, 5 ɗ [
<collectionCode id="35D3AE41FFBC7101FE518FB7FD8BEDCB" LSID="urn:lsid:biocol.org:col:34992" box="[420,517,1444,1470]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="25" pageNumber="26">RMNH</collectionCode>
];
<date id="277C1044FFBC7101FDEC8FB7FD08EDCB" box="[537,646,1444,1470]" pageId="25" pageNumber="26" value="2000-07">VII.2000</date>
, MT, AvH &amp; A. Sallam, 4 ɗ [
<collectionCode id="35D3AE41FFBC7101FC198FB7FBC4EDCB" LSID="urn:lsid:biocol.org:col:34992" box="[1004,1098,1444,1470]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="25" pageNumber="26">RMNH</collectionCode>
,
<collectionCode id="35D3AE41FFBC7101FBA28FB7FB15EDCB" box="[1111,1179,1444,1470]" pageId="25" pageNumber="26">IBSS</collectionCode>
];
<date id="277C1044FFBC7101FB5A8FB7FAA9EDCB" box="[1199,1319,1444,1470]" pageId="25" pageNumber="26" value="2000-08">VIII.2000</date>
, MT, AvH &amp; A. Sallam, 3 ɗ [
<collectionCode id="35D3AE41FFBC7101FD778FDFFD6EED93" LSID="urn:lsid:biocol.org:col:34992" box="[642,736,1484,1510]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="25" pageNumber="26">RMNH</collectionCode>
,
<collectionCode id="35D3AE41FFBC7101FD078FDFFCBBED93" box="[754,821,1484,1510]" pageId="25" pageNumber="26">IBSS</collectionCode>
];
<date id="277C1044FFBC7101FCBA8FDFFC3DED93" box="[847,947,1484,1510]" pageId="25" pageNumber="26" value="2000-09">IX.2000</date>
, MT, AvH &amp; A. Sallam, 2 ɗ [
<collectionCode id="35D3AE41FFBC7101FEFA8FE7FEFEEE7B" LSID="urn:lsid:biocol.org:col:34992" box="[271,368,1524,1550]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="25" pageNumber="26">RMNH</collectionCode>
];
<date id="277C1044FFBC7101FE768FE7FE68EE7B" box="[387,486,1524,1550]" pageId="25" pageNumber="26" value="2000-11">XI.2000</date>
, MT, AvH &amp; A. Sallam, 8 ɗ [
<collectionCode id="35D3AE41FFBC7101FCB68FE7FC2AEE7B" LSID="urn:lsid:biocol.org:col:34992" box="[835,932,1524,1550]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="25" pageNumber="26">RMNH</collectionCode>
];
<date id="277C1044FFBC7101FC428FE7FB1FEE7B" box="[951,1169,1524,1550]" pageId="25" pageNumber="26" value="2000-10-16" valueMax="2000-12-17" valueMin="2000-10-16">16.X–17.XII.2000</date>
, MT, AvH &amp; A.M. Hager, 1 ɗ [
<collectionCode id="35D3AE41FFBC7101FE158C0FFDB1EE43" LSID="urn:lsid:biocol.org:col:34975" box="[480,575,1564,1590]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="25" pageNumber="26">ZMAN</collectionCode>
];
<date id="277C1044FFBC7101FDA18C0FFD22EE43" box="[596,684,1564,1590]" pageId="25" pageNumber="26" value="2001-02">II.2001</date>
, MT, AvH &amp; A. Sallam, 1 ɗ [
<collectionCode id="35D3AE41FFBC7101FBE08C0FFBF8EE43" LSID="urn:lsid:biocol.org:col:34992" box="[1045,1142,1564,1590]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="25" pageNumber="26">RMNH</collectionCode>
];
<date id="277C1044FFBC7101FB7F8C0FFB62EE43" box="[1162,1260,1564,1590]" pageId="25" pageNumber="26" value="2001-03">III.2001</date>
, MT, AvH &amp; A. Sallam, 2 ɗ [
<collectionCode id="35D3AE41FFBC7101FDB68C57FD2FEE2B" LSID="urn:lsid:biocol.org:col:34992" box="[579,673,1604,1630]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="25" pageNumber="26">RMNH</collectionCode>
,
<collectionCode id="35D3AE41FFBC7101FD478C57FD7BEE2B" box="[690,757,1604,1630]" pageId="25" pageNumber="26">IBSS</collectionCode>
]; I–
<date id="277C1044FFBC7101FCDC8C57FC13EE2B" box="[809,925,1604,1630]" pageId="25" pageNumber="26" value="2001-04-09">9.IV.2001</date>
, MT, AvH &amp; A.M. Hager, 2 ɗ [
<collectionCode id="35D3AE41FFBC7101FEFA8C7FFEE1EEF3" LSID="urn:lsid:biocol.org:col:34975" box="[271,367,1644,1670]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="25" pageNumber="26">ZMAN</collectionCode>
];
<date id="277C1044FFBC7101FE768C7FFE6DEEF3" box="[387,483,1644,1670]" pageId="25" pageNumber="26" value="2001-04">IV.2001</date>
, MT, AvH &amp; A. Sallam, 8 ɗ [
<collectionCode id="35D3AE41FFBC7101FCB98C7FFC24EEF3" LSID="urn:lsid:biocol.org:col:34992" box="[844,938,1644,1670]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="25" pageNumber="26">RMNH</collectionCode>
,
<collectionCode id="35D3AE41FFBC7101FC4D8C7FFC72EEF3" box="[952,1020,1644,1670]" pageId="25" pageNumber="26">IBSS</collectionCode>
];
<date id="277C1044FFBC7101FBE48C7FFBE8EEF3" box="[1041,1126,1644,1670]" pageId="25" pageNumber="26" value="2001-05">V.2001</date>
, MT, AvH &amp; A. Sallam, 3 ɗ [
<collectionCode id="35D3AE41FFBC7101FE4E8C87FD97EEDB" LSID="urn:lsid:biocol.org:col:34992" box="[443,537,1684,1710]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="25" pageNumber="26">RMNH</collectionCode>
,
<collectionCode id="35D3AE41FFBC7101FDD88C87FDFEEEDB" box="[557,624,1684,1710]" pageId="25" pageNumber="26">IBSS</collectionCode>
];
<date id="277C1044FFBC7101FD788C87FC93EEDB" box="[653,797,1684,1710]" pageId="25" pageNumber="26" value="2001-12-17">17.XII.2001</date>
–
<date id="277C1044FFBC7101FCDB8C87FC10EEDB" box="[814,926,1684,1710]" pageId="25" pageNumber="26" value="2002-01-31">31.I.2002</date>
, MT, AvH &amp; A. Sallam, 4 ɗ [
<collectionCode id="35D3AE41FFBC7101FEFA8CAFFEFEEEA3" LSID="urn:lsid:biocol.org:col:34992" box="[271,368,1724,1750]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="25" pageNumber="26">RMNH</collectionCode>
];
<date id="277C1044FFBC7101FE738CAFFD86EEA3" box="[390,520,1724,1750]" pageId="25" pageNumber="26" value="2002-03" valueMax="2002-05-31" valueMin="2002-03-00">III–V.2002</date>
, MT, AvH &amp; A. Sallam, 7 ɗ [
<collectionCode id="35D3AE41FFBC7101FC768CAFFC6AEEA3" LSID="urn:lsid:biocol.org:col:34992" box="[899,996,1724,1750]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="25" pageNumber="26">RMNH</collectionCode>
]; Al Kowd,
<date id="277C1044FFBC7101FB888CAFFAA8EEA3" box="[1149,1318,1724,1750]" pageId="25" pageNumber="26" value="1993-02-15" valueMax="1993-02-28" valueMin="1993-02-15">15–28.II.1993</date>
, AvH, MT, 1 ɗ [
<collectionCode id="35D3AE41FFBC7101FE4B8CF7FD93EE8B" LSID="urn:lsid:biocol.org:col:34975" box="[446,541,1764,1790]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="25" pageNumber="26">ZMAN</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFBF7102FEFD8B0FFE03E92B" box="[264,407,284,350]" pageId="26" pageNumber="27" type="distribution">
<paragraph id="537D3684FFBF7102FEFD8B0FFE19E940" blockId="26.[264,407,284,350]" box="[264,407,284,309]" pageId="26" pageNumber="27">
<emphasis id="61B6EA96FFBF7102FEFD8B0FFE19E940" box="[264,407,284,309]" italics="true" pageId="26" pageNumber="27">Distribution</emphasis>
</paragraph>
<paragraph id="537D3684FFBF7102FECD8B57FE03E92B" blockId="26.[264,407,284,350]" box="[312,397,324,350]" pageId="26" pageNumber="27">
<collectingCountry id="2BD57614FFBF7102FECD8B57FE09E92B" box="[312,391,324,350]" name="Yemen" pageId="26" pageNumber="27">Yemen</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFBF7102FEFD8B87FC06E98B" pageId="26" pageNumber="27" type="etymology">
<paragraph id="537D3684FFBF7102FEFD8B87FE0BE9D8" blockId="26.[264,1323,404,510]" box="[264,389,404,429]" pageId="26" pageNumber="27">
<emphasis id="61B6EA96FFBF7102FEFD8B87FE0BE9D8" box="[264,389,404,429]" italics="true" pageId="26" pageNumber="27">Etymology</emphasis>
</paragraph>
<paragraph id="537D3684FFBF7102FECD8BAFFC06E98B" blockId="26.[264,1323,404,510]" pageId="26" pageNumber="27">
This species is dedicated to Willem Hogenes, curator of the
<taxonomicName id="94C24D07FFBF7102FC068BAFFB1DE9A3" box="[1011,1171,444,470]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
collection in the Zoological Museum, Amsterdam,
<collectingCountry id="2BD57614FFBF7102FD328BF7FC0AE98B" box="[711,900,484,510]" name="Netherlands" pageId="26" pageNumber="27">the Netherlands</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

141
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<mods:title id="596D66989F310A6A7929F74FF9472A99">A review of the Mutillidae (Hymenoptera) of Yemen</mods:title>
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<mods:namePart id="27F8545859C86BC7CE4E8F5B8D4C4741">Lelej, Arkady S.</mods:namePart>
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<treatment id="DB6B8792FFBE7104FEFD8F47FB73EC23" ID-DOI="http://doi.org/10.5281/zenodo.6252973" ID-GBIF-Taxon="119344628" ID-Zenodo-Dep="6252973" LSID="urn:lsid:plazi:treatment:DB6B8792FFBE7104FEFD8F47FB73EC23" httpUri="http://treatment.plazi.org/id/DB6B8792FFBE7104FEFD8F47FB73EC23" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
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<taxonomicName id="94C24D07FFBE7103FEFD8F47FD18ED1B" authority="Lelej" authorityName="Lelej" box="[264,662,1364,1390]" class="Insecta" family="Mutillidae" genus="Rasnitsynitilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="brachyptera" status="sp. nov.">
<emphasis id="61B6EA96FFBE7103FEFD8F47FDC0ED1B" bold="true" box="[264,590,1364,1390]" italics="true" pageId="27" pageNumber="28">Rasnitsynitilla brachyptera</emphasis>
Lelej
</taxonomicName>
,
<taxonomicNameLabel id="7A8557EDFFBE7103FD518F47FC8CED1B" box="[676,770,1364,1390]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="537D3684FFBE7103FEFD8F64FE10EDE4" blockId="27.[264,770,1364,1425]" box="[264,414,1399,1425]" pageId="27" pageNumber="28">(Figs 17, 18)</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFBE7103FEFD8FD4FC3FEE7C" box="[264,945,1479,1545]" pageId="27" pageNumber="28" type="diagnosis">
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<emphasis id="61B6EA96FFBE7103FEFD8FD4FEF1ED95" box="[264,383,1479,1504]" italics="true" pageId="27" pageNumber="28">Diagnosis</emphasis>
</paragraph>
<paragraph id="537D3684FFBE7103FECD8FFCFC3FEE7C" blockId="27.[264,1323,1479,1905]" box="[312,945,1519,1545]" pageId="27" pageNumber="28">Diagnostic characters are mentioned in the key below.</paragraph>
</subSubSection>
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<emphasis id="61B6EA96FFBE7103FEFD8C04FE1CEE45" box="[264,402,1559,1584]" italics="true" pageId="27" pageNumber="28">Description</emphasis>
</paragraph>
<paragraph id="537D3684FFBE7103FECD8C2CFE05EF04" blockId="27.[264,1323,1479,1905]" pageId="27" pageNumber="28">MALE. Body length 3.6–4.0 mm. Relative width of head and thorax including tegulae 4.7: 5.3. POD: OOD 0.67–0.72X. Flagellomeres short (Fig. 18), relation of pedicel and first three flagellomeres 0.3: 0.5: 0.9: 0.9. Pronotum anteriorly with rounded humeri. Forewing with small chitinized pterostigma, venation as in Fig. 17. Tegula with shallow punctures. Propodeum reticulate, dorsally with elongate median triangular cell. Pronotum dorsally, mesoscutum, and scutellum with shallow punctures. T1 width 1.2X T1 length. S1 carinate. T2 and S2 densely punctate, punctures larger on S2. T7 with glabrous area in apical half.</paragraph>
<paragraph id="537D3684FFB97104FECD8B0FFE57E98B" blockId="28.[264,1322,284,550]" pageId="28" pageNumber="29">Head, mesosoma, segment 1 of metasoma, and legs yellowish­red. Mandible yellowish­red with dark teeth. Palps whitish. Scape and pedicel yellowish­red; flagellomeres brownish, reddish beneath. Tegulae concolorous with mesosoma. Forewing hyaline. Head, mesosoma, metasoma, and legs with sparse erect and recumbent yellowish setae. T1–T6 and S2–S6 with weak apical fringes of yellowish setae. Lateral felt line of T2 and S2 yellowish.</paragraph>
<paragraph id="537D3684FFB97104FECD881FFDB9EA53" blockId="28.[264,1322,284,550]" box="[312,567,524,550]" pageId="28" pageNumber="29">FEMALE. Unknown.</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB97104FEFD884FFD82EB4B" pageId="28" pageNumber="29" type="materials_examined">
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<emphasis id="61B6EA96FFB97104FEFD884FFE27EA00" box="[264,425,604,629]" italics="true" pageId="28" pageNumber="29">
<typeStatus id="8C798826FFB97104FEFD884FFEB1EA00" box="[264,319,604,629]" pageId="28" pageNumber="29">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="537D3684FFB97104FECD8897FD82EB4B" blockId="28.[264,1323,604,830]" pageId="28" pageNumber="29">
<typeStatus id="8C798826FFB97104FECD8897FE2BEAEB" box="[312,421,644,670]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
ɗ,
<collectingCountry id="2BD57614FFB97104FE2A8897FDBAEAEB" box="[479,564,644,670]" name="Yemen" pageId="28" pageNumber="29">Yemen</collectingCountry>
, Lahj,
<date id="277C1044FFB97104FD618897FCE6EAEB" box="[660,872,644,670]" pageId="28" pageNumber="29" value="2000-05-17" valueMax="2000-06-15" valueMin="2000-05-17">17.V.–15.VI.2000</date>
, MT, AvH &amp; A. Sallam [
<collectionCode id="35D3AE41FFB97104FB4B8897FA91EAEB" LSID="urn:lsid:biocol.org:col:34992" box="[1214,1311,644,670]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="28" pageNumber="29">RMNH</collectionCode>
].
<typeStatus id="8C798826FFB97104FEFD88BFFEF0EAB3" box="[264,382,684,710]" pageId="28" pageNumber="29" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="2BD57614FFB97104FE6188BFFE65EAB3" box="[404,491,684,710]" name="Yemen" pageId="28" pageNumber="29">Yemen</collectingCountry>
: 10 ɗ with the same label as
<typeStatus id="8C798826FFB97104FC6288BFFC73EAB3" box="[919,1021,684,710]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
[
<collectionCode id="35D3AE41FFB97104FBE288BFFBFBEAB3" LSID="urn:lsid:biocol.org:col:34992" box="[1047,1141,684,710]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="28" pageNumber="29">RMNH</collectionCode>
,
<collectionCode id="35D3AE41FFB97104FB7E88BFFB41EAB3" box="[1163,1231,684,710]" pageId="28" pageNumber="29">IBSS</collectionCode>
]; Lahj,
<date id="277C1044FFB97104FEFD88C7FE35EA9B" box="[264,443,724,750]" pageId="28" pageNumber="29" value="2000-06-15" valueMax="2000-06-30" valueMin="2000-06-15">15–30.VI.2000</date>
, MT, AvH &amp; A. Sallam, 5 ɗ [
<collectionCode id="35D3AE41FFB97104FCE288C7FCF6EA9B" LSID="urn:lsid:biocol.org:col:34992" box="[791,888,724,750]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="28" pageNumber="29">RMNH</collectionCode>
];
<date id="277C1044FFB97104FC7E88C7FC76EA9B" box="[907,1016,724,750]" pageId="28" pageNumber="29" value="2000-07">VII.2000</date>
, MT, AvH &amp; A. Sallam, 3 ɗ [
<collectionCode id="35D3AE41FFB97104FED888EFFE00EB63" LSID="urn:lsid:biocol.org:col:34992" box="[301,398,764,790]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="28" pageNumber="29">RMNH</collectionCode>
];
<date id="277C1044FFB97104FE5088EFFD87EB63" box="[421,521,764,790]" pageId="28" pageNumber="29" value="2000-09">IX.2000</date>
, MT, AvH &amp; A. Sallam, 1 ɗ [
<collectionCode id="35D3AE41FFB97104FC7688EFFC6AEB63" LSID="urn:lsid:biocol.org:col:34992" box="[899,996,764,790]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="28" pageNumber="29">RMNH</collectionCode>
];
<date id="277C1044FFB97104FC0E88EFFBD0EB63" box="[1019,1118,764,790]" pageId="28" pageNumber="29" value="2000-11">XI.2000</date>
, MT, AvH &amp; A. Sallam, 1 ɗ [
<collectionCode id="35D3AE41FFB97104FE558937FD8FEB4B" LSID="urn:lsid:biocol.org:col:34992" box="[416,513,804,830]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="28" pageNumber="29">RMNH</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB97104FEFD8967FE03EBC3" box="[264,407,884,950]" pageId="28" pageNumber="29" type="distribution">
<paragraph id="537D3684FFB97104FEFD8967FE19EBF8" blockId="28.[264,407,884,950]" box="[264,407,884,909]" pageId="28" pageNumber="29">
<emphasis id="61B6EA96FFB97104FEFD8967FE19EBF8" box="[264,407,884,909]" italics="true" pageId="28" pageNumber="29">Distribution</emphasis>
</paragraph>
<paragraph id="537D3684FFB97104FECD898FFE03EBC3" blockId="28.[264,407,884,950]" box="[312,397,924,950]" pageId="28" pageNumber="29">
<collectingCountry id="2BD57614FFB97104FECD898FFE09EBC3" box="[312,391,924,950]" name="Yemen" pageId="28" pageNumber="29">Yemen</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="1BD8650FFFB97104FEFD89FFFB73EC23" pageId="28" pageNumber="29" type="etymology">
<paragraph id="537D3684FFB97104FEFD89FFFE0BEC70" blockId="28.[264,1322,1004,1110]" box="[264,389,1004,1029]" pageId="28" pageNumber="29">
<emphasis id="61B6EA96FFB97104FEFD89FFFE0BEC70" box="[264,389,1004,1029]" italics="true" pageId="28" pageNumber="29">Etymology</emphasis>
</paragraph>
<paragraph id="537D3684FFB97104FECD8E07FB73EC23" blockId="28.[264,1322,1004,1110]" pageId="28" pageNumber="29">
<taxonomicName id="94C24D07FFB97104FECD8E07FE45EC58" box="[312,459,1044,1069]" class="Insecta" family="Mutillidae" genus="Rasnitsynitilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="31" phylum="Arthropoda" rank="species" species="brachyptera" status="sp. nov.">
<emphasis id="61B6EA96FFB97104FECD8E07FE45EC58" box="[312,459,1044,1069]" italics="true" pageId="28" pageNumber="29">Brachyptera</emphasis>
</taxonomicName>
is derived from the Greek adjective
<emphasis id="61B6EA96FFB97104FC768E07FC6EEC58" box="[899,992,1044,1069]" italics="true" pageId="28" pageNumber="29">brachys</emphasis>
, pertaining to short and the noun
<emphasis id="61B6EA96FFB97104FEBF8E2FFE1BEC20" box="[330,405,1084,1109]" italics="true" pageId="28" pageNumber="29">pteron</emphasis>
(plural
<emphasis id="61B6EA96FFB97104FE078E2FFDBEEC20" box="[498,560,1084,1109]" italics="true" pageId="28" pageNumber="29">ptera</emphasis>
), pertaining to wing (wings), referring to its short forewings.
</paragraph>
</subSubSection>
</treatment>
</document>

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[TL:
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, Elman Dagh], following
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, note 14) this name is a
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. It was described by Gaede as follows: ‘a pale
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North
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similar to
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Fruhst.
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’ (Translation). Although
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attributed the name to Rebel (undated), no description by Rebel or anyone else has been found.
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Hesselbarth
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. (1995: 1031)
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suggested that it may have been a misspelling of
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Rebel, but Gaede treated the names separately. As far as the authors are aware there has not been any illustration of “almana” and it was not mentioned by
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.
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[TL: Askhabad,
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. The Type Locality is just across the border from north-eastern
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Tshikolovets
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. 2014: 319–320
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). The type material, present in the Natural History Museum, London, consists of a
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male (
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), an
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female and
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and
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<typeStatus id="8C798804FFF0FFEEE5EF0181FAE0FE36" box="[1237,1347,475,501]" pageId="8" pageNumber="33" type="paratype">paratypes</typeStatus>
, All the specimens display the wing characters typical of
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, of which it was considered previously to be a subspecies (
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Tshikolovets
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. 2014: 319
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and
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, note 33).
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[TL:
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, Sarvistan, SE of Shiraz salt lake] has been considered a distinct species,
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<emphasis id="61B6EAB4FFF0FFEEE01702D5FEFDFD6B" box="[301,350,654,680]" italics="true" pageId="8" pageNumber="33">et al</emphasis>
. (2014: 321–322
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&amp; plate LX: figs 13-18).
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<bibRefCitation id="37534B57FFF0FFEEE08A02E9FD95FD0F" author="Varga, Z. S." box="[432,566,690,717]" pageId="8" pageNumber="33" pagination="119 - 137" refId="ref11743" refString="Varga, Z. S. (1967) A Melitaea phoebe Schiff. Delkelet-europai populacioinak taxonomiai elemzese, ket uj alfaj leirasaval [A taxonomic analysis of south-eastern populations of Melitaea phoebe Schiff. With descriptions of two new subspecies]. Acta Biologica Debrecina, 5, 119 - 137." type="journal article" year="1967">Varga, 1967</bibRefCitation>
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[TL:
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, Kiverce, Podalia],
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Varga
<emphasis id="61B6EAB4FFF0FFEEE2CE02E9FB86FD0F" box="[1012,1061,690,716]" italics="true" pageId="8" pageNumber="33">et al</emphasis>
. (2005)
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placed this taxon with
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<emphasis id="61B6EAB4FFF0FFEEE44402E9FF40FD33" italics="true" pageId="8" pageNumber="33">M. ogygia</emphasis>
<bibRefCitation id="37534B57FFF0FFEEE1D0028CFE0FFD33" author="Fruhstorfer, H." box="[234,428,726,752]" pageId="8" pageNumber="33" pagination="310" refId="ref9058" refString="Fruhstorfer, H. (1908) Neue Argynnis und Melitaea. Internationale Entomologische Zeitschrift, 1, 310." type="journal article" year="1908">Fruhstorfer, 1908</bibRefCitation>
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following which
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Tóth
<emphasis id="61B6EAB4FFF0FFEEE391028DFD7FFD33" box="[683,732,726,752]" italics="true" pageId="8" pageNumber="33">et al</emphasis>
. (2013)
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placed it with
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<emphasis id="61B6EAB4FFF0FFEEE2E2028DFBEBFD33" box="[984,1096,727,752]" italics="true" pageId="8" pageNumber="33">M. ornata</emphasis>
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<bibRefCitation id="37534B57FFF0FFEEE56F028CFA34FD32" author="Russell, PJ &amp; Tennent, W. J." box="[1109,1431,726,753]" pageId="8" pageNumber="33" pagination="27 - 56" refId="ref10922" refString="Russell, PJ. C. &amp; Tennent, W. J. (2016) A synonymic list of names associated with western Palaearctic Melitaea phoebe (Denis &amp; Schiffermuller, 1775) species group taxa M. phoebe; M. punica Oberthur, 1876; M. ornata Christoph, 1893) (Lepidoptera, Nymphalidae). Nota Lepidopterologica, 39 (1), 27 - 56. https: // doi. org / 10.3897 / nl. 39.5929" type="journal article" year="2016">Russell &amp; Tennent (2016: 45</bibRefCitation>
, note 21) were in agreement with the latter association, as are the present authors.
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[TL:
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, Budakeszi],
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Varga
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(2005)
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<emphasis id="61B6EAB4FFF0FFEEE4160345FA3FFCFB" box="[1324,1436,799,824]" italics="true" pageId="8" pageNumber="33">M. ogygia</emphasis>
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and later
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placed it with
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,
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, note 45) agreed with the latter association and the current authors also agree.
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<treatment id="DB6B87B0FFF0FFEEE1FD03D1FCA1F88F" ID-DOI="http://doi.org/10.5281/zenodo.6580855" ID-GBIF-Taxon="195626728" ID-Zenodo-Dep="6580855" LSID="urn:lsid:plazi:treatment:DB6B87B0FFF0FFEEE1FD03D1FCA1F88F" httpUri="http://treatment.plazi.org/id/DB6B87B0FFF0FFEEE1FD03D1FCA1F88F" lastPageNumber="33" pageId="8" pageNumber="33">
<subSubSection id="1BD8652DFFF0FFEEE1FD03D1FDD7FC66" box="[199,628,906,933]" pageId="8" pageNumber="33" type="nomenclature">
<paragraph id="537D36A6FFF0FFEEE1FD03D1FDD7FC66" blockId="8.[151,1437,150,2013]" box="[199,628,906,933]" pageId="8" pageNumber="33">
<taxonomicName id="94C24D25FFF0FFEEE1FD03D1FDD7FC66" authority="Gross &amp; Ebert, 1975" authorityName=", Gross &amp; Ebert" authorityYear="1975" box="[199,628,906,933]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="abbas">
<emphasis id="61B6EAB4FFF0FFEEE1FD03D1FE2AFC66" bold="true" box="[199,393,906,933]" italics="true" pageId="8" pageNumber="33">M. phoebe abbas</emphasis>
<bibRefCitation id="37534B57FFF0FFEEE0B503D0FDD7FC66" author="Gross, F. J. &amp; Ebert, G." box="[399,628,906,933]" pageId="8" pageNumber="33" pagination="8 - 45" refId="ref9178" refString="Gross, F. J. &amp; Ebert, G. (1975) Beitrage zur Kenntis der Rhopaloceren Irans. 2. Beitrag. Neue Taxa der Pieridae, Satyridae und Nymphalidae. Journal of the Entomological Society of Iran Supplement, 1, 8 - 45." type="journal article" year="1975">Gross &amp; Ebert, 1975</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="1BD8652DFFF0FFEEE34003D1FCA1F88F" pageId="8" pageNumber="33" type="discussion">
<paragraph id="537D36A6FFF0FFEEE34003D1FDCDFBF7" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="33">
[TL:
<quantity id="943A9B43FFF0FFEEE38903D0FD5AFC67" box="[691,761,906,933]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="33" unit="km" value="50.0">50 km</quantity>
NW Ardekan, Tange-Surkh,
<collectingCountry id="2BD57636FFF0FFEEE50603D1FBCFFC66" box="[1084,1132,907,933]" name="Iran" pageId="8" pageNumber="33">Iran</collectingCountry>
,
<quantity id="943A9B43FFF0FFEEE54D03D0FB73FC66" box="[1143,1232,906,933]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="8" pageNumber="33" unit="m" value="2250.0">2250 m</quantity>
]: junior subjective synonym of
<taxonomicName id="94C24D25FFF0FFEEE01903F4FDD6FC0B" authority="Higgins, 1941" authorityName="Higgins" authorityYear="1941" box="[291,629,942,968]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="33" phylum="Arthropoda" rank="species" species="enoch">
<emphasis id="61B6EAB4FFF0FFEEE01903F4FE73FC0B" box="[291,464,942,968]" italics="true" pageId="8" pageNumber="33">Melitaea enoch</emphasis>
<bibRefCitation id="37534B57FFF0FFEEE0EC03F4FDD6FC0B" author="Higgins, L. G." box="[470,629,942,968]" pageId="8" pageNumber="33" pagination="175 - 365" refId="ref9372" refString="Higgins, L. G. (1941) An illustrated catalogue of the Palearctic Melitaea (Lep. Rhopalocera). Transactions of the Royal Entomological Society of London, 91, 175 - 365. https: // doi. org / 10.1111 / j. 1365 - 2311.1941. tb 01045. x" type="journal article" year="1941">Higgins, 1941</bibRefCitation>
</taxonomicName>
(
<emphasis id="61B6EAB4FFF0FFEEE3BF03F4FD4FFC0B" bold="true" box="[645,748,942,968]" pageId="8" pageNumber="33">syn. nov.</emphasis>
). The synonymy of
<taxonomicName id="94C24D25FFF0FFEEE2F403F4FBB2FC0B" authorityName="Gross &amp; Ebert" authorityYear="1975" box="[974,1041,942,968]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="33" phylum="Arthropoda" rank="species" species="abbas">
<emphasis id="61B6EAB4FFF0FFEEE2F403F4FBB2FC0B" box="[974,1041,942,968]" italics="true" pageId="8" pageNumber="33">abbas</emphasis>
</taxonomicName>
with
<taxonomicName id="94C24D25FFF0FFEEE56A03F5FB1BFC0B" authorityName="Higgins" authorityYear="1941" box="[1104,1208,942,968]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="33" phylum="Arthropoda" rank="species" species="enoch">
<emphasis id="61B6EAB4FFF0FFEEE56A03F5FB1BFC0B" box="[1104,1208,942,968]" italics="true" pageId="8" pageNumber="33">M. enoch</emphasis>
</taxonomicName>
is based on the very close similarity of the wing morphology of the
<specimenCount id="45C4FD2FFFF0FFEEE3990389FCE3FC2F" box="[675,832,978,1005]" pageId="8" pageNumber="33" type="generic" typeStatus="holotypes">two holotypes</specimenCount>
(see figs 17A &amp; B and 16A &amp; B, respectively) and the fact that
<bibRefCitation id="37534B57FFF0FFEEE1C303ACFE1CFBD2" author="Toth, J. P. &amp; Varga, Z. &amp; Bereczki, J." box="[249,447,1014,1041]" pageId="8" pageNumber="33" pagination="106 - 115" refId="ref11405" refString="Toth, J. P., Varga, Z. &amp; Bereczki, J. (2016) Long-term survival and diversification of an endemic Melitaea species in mountains of Iran and adjacent areas. Journal of Zoological Systematics and Evolutionary Research, 54 (2), 106 - 115. https: // doi. org / 10.1111 / jzs. 12122" type="journal article" year="2016">
Tóth
<emphasis id="61B6EAB4FFF0FFEEE00F03ADFEC4FBD3" box="[309,359,1014,1040]" italics="true" pageId="8" pageNumber="33">et al</emphasis>
. (2016)
</bibRefCitation>
could find no genetic differences between the taxon
<taxonomicName id="94C24D25FFF0FFEEE52803ACFBF6FBD3" authorityName="Gross &amp; Ebert" authorityYear="1975" box="[1042,1109,1014,1040]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="33" phylum="Arthropoda" rank="species" species="abbas">
<emphasis id="61B6EAB4FFF0FFEEE52803ACFBF6FBD3" box="[1042,1109,1014,1040]" italics="true" pageId="8" pageNumber="33">abbas</emphasis>
</taxonomicName>
from the northern (where
<taxonomicName id="94C24D25FFF0FFEEE44403ADFF79FBF7" authorityName="Higgins" authorityYear="1941" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="33" phylum="Arthropoda" rank="species" species="enoch">
<emphasis id="61B6EAB4FFF0FFEEE44403ADFF79FBF7" italics="true" pageId="8" pageNumber="33">M. enoch</emphasis>
</taxonomicName>
is found) and southern populations.
</paragraph>
<paragraph id="537D36A6FFF0FFEEE1FD0464FCA2FA5F" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="33">
The original description (
<bibRefCitation id="37534B57FFF0FFEEE0DA0464FD49FB9B" author="Gross, F. J. &amp; Ebert, G." box="[480,746,1086,1113]" pageId="8" pageNumber="33" pagination="8 - 45" refId="ref9178" refString="Gross, F. J. &amp; Ebert, G. (1975) Beitrage zur Kenntis der Rhopaloceren Irans. 2. Beitrag. Neue Taxa der Pieridae, Satyridae und Nymphalidae. Journal of the Entomological Society of Iran Supplement, 1, 8 - 45." type="journal article" year="1975">Gross &amp; Ebert 1975: 44</bibRefCitation>
) indicated the presence of a
<typeStatus id="8C798804FFF0FFEEE51E0464FB25FB9B" box="[1060,1158,1086,1112]" pageId="8" pageNumber="33" type="holotype">holotype</typeStatus>
male, an
<typeStatus id="8C798804FFF0FFEEE5CA0464FAE9FB9B" box="[1264,1354,1086,1112]" pageId="8" pageNumber="33" type="allotype">allotype</typeStatus>
female and
<specimenCount id="45C4FD2FFFF0FFEEE1FC0438FEB2FBBF" box="[198,273,1122,1148]" pageId="8" pageNumber="33" type="male">9 male</specimenCount>
<typeStatus id="8C798804FFF0FFEEE0220439FE24FBBE" box="[280,391,1123,1149]" pageId="8" pageNumber="33" type="paratype">paratypes</typeStatus>
. The
<typeStatus id="8C798804FFF0FFEEE0F90438FD86FBBF" box="[451,549,1122,1148]" pageId="8" pageNumber="33" type="holotype">holotype</typeStatus>
(
<figureCitation id="CBF92A23FFF0FFEEE30E0438FD7CFBBF" box="[564,735,1122,1149]" captionStart="FIGURES 17–20" captionStartId="9.[151,264,1710,1734]" captionTargetBox="[151,1436,381,1687]" captionTargetId="figure-121@9.[151,1436,381,1687]" captionTargetPageId="9" captionText="FIGURES 17–20. 17, Holotype of Melitaea phoebe abbas, Gross &amp; Ebert, 1975. © R. Trusch. 18, Allotype of Melitaea phoebe abbas, Gross &amp; Ebert, 1975. © Alipanah. 19, Holotype of Melitaea ornata adversaria Korb, Stradomsky &amp; Kuznetsov, 2015. 20, Paratype of Melitaea ornata reliquiae Korb, Stradomsky &amp; Kuznetsov, 2015. Figs 19, 20 © S.K. Korb, B.V. Stradomsky &amp; G.V. Kuznetsov.A = upperside, B = underside, C = labels on specimen pin." figureDoi="http://doi.org/10.5281/zenodo.6577646" httpUri="https://zenodo.org/record/6577646/files/figure.png" pageId="8" pageNumber="33">Figs 17A, B, C</figureCitation>
) and
<specimenCount id="45C4FD2FFFF0FFEEE2270438FCD8FBBF" box="[797,891,1122,1148]" pageId="8" pageNumber="33" type="male">six male</specimenCount>
<typeStatus id="8C798804FFF0FFEEE2B80439FC4EFBBE" box="[898,1005,1123,1149]" pageId="8" pageNumber="33" type="paratype">paratypes</typeStatus>
are present in the Staatliches Museum für Naturkunde Karlsruhe (
<bibRefCitation id="37534B57FFF0FFEEE0F104DCFCB7FB62" author="Hesselbarth, G. &amp; van Oorschot, H. &amp; Wagener, S." box="[459,788,1158,1185]" pageId="8" pageNumber="33" pagination="1 - 847" refId="ref9287" refString="Hesselbarth, G., van Oorschot, H. &amp; Wagener, S. (1995) Die Tagfalter der Turkei unter Berucksichtigung der angrenzenden Lander, Sigbert Wagener, Bocholt, Germany, 1, 1 - 757, 2, 758 - 1354, 3, 1 - 847." type="journal article" year="1995">
Hesselbarth
<emphasis id="61B6EAB4FFF0FFEEE36304DDFD2FFB63" box="[601,652,1158,1184]" italics="true" pageId="8" pageNumber="33">et al</emphasis>
. 1995:1031
</bibRefCitation>
) and an
<typeStatus id="8C798804FFF0FFEEE24204DCFC71FB63" box="[888,978,1158,1184]" pageId="8" pageNumber="33" type="allotype">allotype</typeStatus>
female (
<figureCitation id="CBF92A23FFF0FFEEE50D04DCFB45FB63" box="[1079,1254,1158,1185]" captionStart="FIGURES 17–20" captionStartId="9.[151,264,1710,1734]" captionTargetBox="[151,1436,381,1687]" captionTargetId="figure-121@9.[151,1436,381,1687]" captionTargetPageId="9" captionText="FIGURES 17–20. 17, Holotype of Melitaea phoebe abbas, Gross &amp; Ebert, 1975. © R. Trusch. 18, Allotype of Melitaea phoebe abbas, Gross &amp; Ebert, 1975. © Alipanah. 19, Holotype of Melitaea ornata adversaria Korb, Stradomsky &amp; Kuznetsov, 2015. 20, Paratype of Melitaea ornata reliquiae Korb, Stradomsky &amp; Kuznetsov, 2015. Figs 19, 20 © S.K. Korb, B.V. Stradomsky &amp; G.V. Kuznetsov.A = upperside, B = underside, C = labels on specimen pin." figureDoi="http://doi.org/10.5281/zenodo.6577646" httpUri="https://zenodo.org/record/6577646/files/figure.png" pageId="8" pageNumber="33">Figs 18A, B, C</figureCitation>
) and a
<typeStatus id="8C798804FFF0FFEEE40604DDFA3FFB62" box="[1340,1436,1159,1185]" pageId="8" pageNumber="33" type="paratype">paratype</typeStatus>
male are present in the Hayk Mirzayans Insect Museum, Iranian Research Institute of Plant Protection,
<collectingRegion id="9106F844FFF0FFEEE40604F0FA3AFB07" box="[1340,1433,1194,1220]" country="Iran" name="Tehran" pageId="8" pageNumber="33">Teheran</collectingRegion>
,
<collectingCountry id="2BD57636FFF0FFEEE1AD0495FF6BFB2A" box="[151,200,1231,1257]" name="Iran" pageId="8" pageNumber="33">Iran</collectingCountry>
. This was confirmed by
<bibRefCitation id="37534B57FFF0FFEEE0E40494FD77FB2B" author="Alipanah, H." box="[478,724,1230,1256]" pageId="8" pageNumber="33" pagination="191 - 210" refId="ref8644" refString="Alipanah, H. (2016) List of the Lepidoptera type material deposited in the Hayk Mirzayans Insect Museum (HMIM), Iranian Research Institute of Plant Protection (IRIPP) (Insecta: Lepidoptera). SHILAP Revista de Lepidopterologia, 44 (174), 191 - 210." type="journal article" year="2016">Alipanah (2016: 200)</bibRefCitation>
, who stated that Tange-Surkh is a misspelling of Tang-e Sorkh and Ardekan is currently known as Sepidân. Of the remaining
<specimenCount id="45C4FD2FFFF0FFEEE24A04A9FC4BFACF" box="[880,1000,1266,1293]" pageId="8" pageNumber="33" type="male">two males</specimenCount>
, one is deposited in the Zoologisches Forschungsmuseum Alexander Koenig in Bonn,
<collectingCountry id="2BD57636FFF0FFEEE3F3054CFC92FAF3" box="[713,817,1302,1328]" name="Germany" pageId="8" pageNumber="33">Germany</collectingCountry>
and the other in Staatliches Museum für Naturkunde Karlsruhe (SMNK),
<collectingCountry id="2BD57636FFF0FFEEE0460560FE45FA97" box="[380,486,1338,1364]" name="Germany" pageId="8" pageNumber="33">Germany</collectingCountry>
. Recent research by
<bibRefCitation id="37534B57FFF0FFEEE3F30560FC26FA97" author="Toth, J. P. &amp; Bereczki, J. &amp; Varga, Z. &amp; Rota, J. &amp; Sramko, G. &amp; Wahlberg, N." box="[713,901,1338,1365]" pageId="8" pageNumber="33" pagination="749 - 757" refId="ref11329" refString="Toth, J. P., Bereczki, J., Varga, Z., Rota, J., Sramko, G. &amp; Wahlberg, N. (2014) Relationships within the Melitaea phoebe species group (Lepidoptera: Nymphalidae): new insights from molecular and morphometric information. Systematic Entomology, 39, 749 - 757. https: // doi. org / 10.1111 / syen. 12083" type="journal article" year="2014">
Tóth
<emphasis id="61B6EAB4FFF0FFEEE23E0561FC97FA97" box="[772,820,1338,1364]" italics="true" pageId="8" pageNumber="33">et al</emphasis>
. (2014
</bibRefCitation>
,
<bibRefCitation id="37534B57FFF0FFEEE2AA0560FC6FFA96" author="Toth, J. P. &amp; Varga, Z. &amp; Bereczki, J." box="[912,972,1338,1365]" pageId="8" pageNumber="33" pagination="106 - 115" refId="ref11405" refString="Toth, J. P., Varga, Z. &amp; Bereczki, J. (2016) Long-term survival and diversification of an endemic Melitaea species in mountains of Iran and adjacent areas. Journal of Zoological Systematics and Evolutionary Research, 54 (2), 106 - 115. https: // doi. org / 10.1111 / jzs. 12122" type="journal article" year="2016">2016</bibRefCitation>
), indicated that the taxon
<taxonomicName id="94C24D25FFF0FFEEE5D20560FA88FA97" authorityName="Gross &amp; Ebert" authorityYear="1975" box="[1256,1323,1338,1364]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="33" phylum="Arthropoda" rank="species" species="abbas">
<emphasis id="61B6EAB4FFF0FFEEE5D20560FA88FA97" box="[1256,1323,1338,1364]" italics="true" pageId="8" pageNumber="33">abbas</emphasis>
</taxonomicName>
should be elevated to species level. There was no label on the pin of the
<typeStatus id="8C798804FFF0FFEEE26C0504FC13FABB" box="[854,944,1374,1400]" pageId="8" pageNumber="33" type="allotype">allotype</typeStatus>
to indicate its status, this has been rectified and its current specific identification noted (
<figureCitation id="CBF92A23FFF0FFEEE3BC05D8FD57FA5F" box="[646,756,1410,1436]" captionStart="FIGURES 17–20" captionStartId="9.[151,264,1710,1734]" captionTargetBox="[151,1436,381,1687]" captionTargetId="figure-121@9.[151,1436,381,1687]" captionTargetPageId="9" captionText="FIGURES 17–20. 17, Holotype of Melitaea phoebe abbas, Gross &amp; Ebert, 1975. © R. Trusch. 18, Allotype of Melitaea phoebe abbas, Gross &amp; Ebert, 1975. © Alipanah. 19, Holotype of Melitaea ornata adversaria Korb, Stradomsky &amp; Kuznetsov, 2015. 20, Paratype of Melitaea ornata reliquiae Korb, Stradomsky &amp; Kuznetsov, 2015. Figs 19, 20 © S.K. Korb, B.V. Stradomsky &amp; G.V. Kuznetsov.A = upperside, B = underside, C = labels on specimen pin." figureDoi="http://doi.org/10.5281/zenodo.6577646" httpUri="https://zenodo.org/record/6577646/files/figure.png" pageId="8" pageNumber="33">Fig. 18 C</figureCitation>
).
</paragraph>
<paragraph id="537D36A6FFF0FFEEE1FD05FCFCA1F88F" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="33">
There is some muddle in the figures of this species under the name
<taxonomicName id="94C24D25FFF0FFEEE28C05FDFBC8FA03" box="[950,1131,1446,1472]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="telona" subSpecies="abbas">
<emphasis id="61B6EAB4FFF0FFEEE28C05FDFBC8FA03" box="[950,1131,1446,1472]" italics="true" pageId="8" pageNumber="33">M. telona abbas</emphasis>
</taxonomicName>
presented by
<bibRefCitation id="37534B57FFF0FFEEE43C05FDFEC4FA27" author="Van Oorschot, H. &amp; Coutsis, J. G." pageId="8" pageNumber="33" refId="ref11693" refString="Van Oorschot, H. &amp; Coutsis, J. G. (2014) The genus Melitaea Fabricius, 1807 (Lepidoptera, Nymphalidae, Nymphalinae) Taxonomy and systematics with special reference to the male genitalia. Tshikolovets publications, Pardubice, Czech Republic, 360 pp." type="book" year="2014">van Oorschot and Coutsis (2014</bibRefCitation>
: plate 13 figs 11, 12, 13 &amp; 16); their figure 16 correctly shows the
<typeStatus id="8C798804FFF0FFEEE56C0590FB1BFA27" box="[1110,1208,1482,1508]" pageId="8" pageNumber="33" type="holotype">holotype</typeStatus>
in SMNK; however the captions for the specimens (numbers 1463–1465) in figures 11–13 indicate firstly that they are all female and secondly that they are housed in the SMNK (
<bibRefCitation id="37534B57FFF0FFEEE3A60648FBE1F9EE" author="Van Oorschot, H. &amp; Coutsis, J. G." box="[668,1090,1554,1581]" pageId="8" pageNumber="33" refId="ref11693" refString="Van Oorschot, H. &amp; Coutsis, J. G. (2014) The genus Melitaea Fabricius, 1807 (Lepidoptera, Nymphalidae, Nymphalinae) Taxonomy and systematics with special reference to the male genitalia. Tshikolovets publications, Pardubice, Czech Republic, 360 pp." type="book" year="2014">van Oorschot &amp; Coutsis (2014: 167)</bibRefCitation>
; figure 13 does appear to be a female but figures 11 and 12 appear to be males. There are no female
<typeStatus id="8C798804FFF0FFEEE2A3066DFBA7F992" box="[921,1028,1591,1617]" pageId="8" pageNumber="33" type="paratype">paratypes</typeStatus>
among the specimens in SMNK, the
<typeStatus id="8C798804FFF0FFEEE1AD0600FF52F9B7" box="[151,241,1626,1652]" pageId="8" pageNumber="33" type="allotype">allotype</typeStatus>
and only female is in the
<collectingRegion id="9106F844FFF0FFEEE32A0600FDC9F9B7" box="[528,618,1626,1652]" country="Iran" name="Tehran" pageId="8" pageNumber="33">Teheran</collectingRegion>
museum (see above) and was taken at the same locality but on
<date id="277C1066FFF0FFEEE4160600FA3FF9B7" box="[1324,1436,1626,1652]" pageId="8" pageNumber="33" value="1972-06-16">16.6.1972</date>
(see our fig. 19C). The
<specimenCount id="45C4FD2FFFF0FFEEE0A30625FE5CF95B" box="[409,511,1662,1689]" pageId="8" pageNumber="33" type="male">two male</specimenCount>
specimens illustrated were taken apparently on
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and
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(
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. These dates exactly match those on two of the labels on the pins of
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specimens in SMNK, but those specimens do not appear to match the specimens figured by
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: plate 13 figs 11 &amp; 12), the
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illustrated have differently orientated antennae and wing sections missing which do not match any specimens in the collection of
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in the SMNK. Neither John Coutsis nor the present authors are able to explain this discrepancy.
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111
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TL: W.
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NNW from
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Mountains. Types figured by the authorities: Plates V &amp; VI (upperside &amp; underside respectively), figs 10 &amp; 11 (male
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, 54, note 85), who mistakenly indicated at first that the specimens were taken in August, 2003 (p. 33) rather than correctly in July (p. 54). The
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<treatment id="DB6B87B0FFF1FFECE1FD073AFDA8FD46" ID-DOI="http://doi.org/10.5281/zenodo.6580861" ID-GBIF-Taxon="195626726" ID-Zenodo-Dep="6580861" LSID="urn:lsid:plazi:treatment:DB6B87B0FFF1FFECE1FD073AFDA8FD46" httpUri="http://treatment.plazi.org/id/DB6B87B0FFF1FFECE1FD073AFDA8FD46" lastPageId="10" lastPageNumber="34" pageId="9" pageNumber="34">
<subSubSection id="1BD8652DFFF1FFEFE1FD073AFDCCF8B8" box="[199,623,1888,1915]" pageId="9" pageNumber="34" type="nomenclature">
<paragraph id="537D36A6FFF1FFEFE1FD073AFDCCF8B8" blockId="9.[151,1437,1888,2023]" box="[199,623,1888,1915]" pageId="9" pageNumber="34">
<taxonomicName id="94C24D25FFF1FFEFE1FD073AFDCCF8B8" authority="Toth &amp; Varga, 2011" authorityName="Toth &amp; Varga" authorityYear="2011" box="[199,623,1888,1915]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="34" phylum="Arthropoda" rank="species" species="zagrosi">
<emphasis id="61B6EAB4FFF1FFEFE1FD073AFE25F8B8" bold="true" box="[199,390,1888,1915]" italics="true" pageId="9" pageNumber="34">Melitaea zagrosi</emphasis>
<bibRefCitation id="37534B57FFF1FFEFE0B4073BFDCCF8B8" author="Toth, J. P. &amp; Varga, Z." box="[398,623,1889,1915]" pageId="9" pageNumber="34" pagination="258 - 268" refId="ref11248" refString="Toth, J. P. &amp; Varga, Z. (2011) Inter- and intraspecific variation in the genitalia of the ' Melitaea phoebe group' (Lepidoptera. Nymphalidae). Zoologischer Anzeiger, 250, 258 - 268. [6 pp. of Appendix with key and figures of genitalia only present in the on-line version, www. sciencedirect. com] https: // doi. org / 10.1016 / j. jcz. 2011.05.002" type="book chapter" year="2011">Tóth &amp; Varga, 2011</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="1BD8652DFFF1FFECE34D073BFDA8FD46" lastPageId="10" lastPageNumber="35" pageId="9" pageNumber="34" type="discussion">
<paragraph id="537D36A6FFF1FFECE34D073BFDA8FD46" blockId="9.[151,1437,1888,2023]" lastBlockId="10.[151,1437,151,933]" lastPageId="10" lastPageNumber="35" pageId="9" pageNumber="34">
[TL:
<collectingCountry id="2BD57636FFF1FFEFE389073BFD40F8B8" box="[691,739,1889,1915]" name="Iran" pageId="9" pageNumber="34">Iran</collectingCountry>
, between Kazeroun and Buschir (Konar Takhteh)], subjective junior synonym of
<taxonomicName id="94C24D25FFF1FFEFE05507DFFD67F85C" authority="Higgins, 1941" authorityName="Higgins" authorityYear="1941" box="[367,708,1925,1951]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="34" phylum="Arthropoda" rank="species" species="enoch">
<emphasis id="61B6EAB4FFF1FFEFE05507DFFDBEF85C" box="[367,541,1925,1951]" italics="true" pageId="9" pageNumber="34">Melitaea enoch</emphasis>
<bibRefCitation id="37534B57FFF1FFEFE31E07DFFD67F85C" author="Higgins, L. G." box="[548,708,1925,1951]" pageId="9" pageNumber="34" pagination="175 - 365" refId="ref9372" refString="Higgins, L. G. (1941) An illustrated catalogue of the Palearctic Melitaea (Lep. Rhopalocera). Transactions of the Royal Entomological Society of London, 91, 175 - 365. https: // doi. org / 10.1111 / j. 1365 - 2311.1941. tb 01045. x" type="journal article" year="1941">Higgins, 1941</bibRefCitation>
</taxonomicName>
(
<emphasis id="61B6EAB4FFF1FFEFE3EE07DFFC9EF85C" bold="true" box="[724,829,1925,1951]" pageId="9" pageNumber="34">syn. nov.</emphasis>
). This taxon was distinguished from
<taxonomicName id="94C24D25FFF1FFEFE5DB07DCFA3FF85C" baseAuthorityName="Toth &amp; Varga" baseAuthorityYear="2011" box="[1249,1436,1925,1951]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="ornata" subSpecies="abbas">
<emphasis id="61B6EAB4FFF1FFEFE5DB07DCFA3FF85C" box="[1249,1436,1925,1951]" italics="true" pageId="9" pageNumber="34">M. ornata abbas</emphasis>
</taxonomicName>
on the basis of genitalia and wing patterns by
<bibRefCitation id="37534B57FFF1FFEFE39A07F3FC2CF800" author="Toth, J. P. &amp; Varga, Z." box="[672,911,1961,1987]" pageId="9" pageNumber="34" pagination="258 - 268" refId="ref11248" refString="Toth, J. P. &amp; Varga, Z. (2011) Inter- and intraspecific variation in the genitalia of the ' Melitaea phoebe group' (Lepidoptera. Nymphalidae). Zoologischer Anzeiger, 250, 258 - 268. [6 pp. of Appendix with key and figures of genitalia only present in the on-line version, www. sciencedirect. com] https: // doi. org / 10.1016 / j. jcz. 2011.05.002" type="book chapter" year="2011">Tóth &amp; Varga (2011)</bibRefCitation>
.
<bibRefCitation id="37534B57FFF1FFEFE2A107F3FA8AF807" author="Van Oorschot, H. &amp; Coutsis, J. G." box="[923,1321,1961,1988]" pageId="9" pageNumber="34" refId="ref11693" refString="Van Oorschot, H. &amp; Coutsis, J. G. (2014) The genus Melitaea Fabricius, 1807 (Lepidoptera, Nymphalidae, Nymphalinae) Taxonomy and systematics with special reference to the male genitalia. Tshikolovets publications, Pardubice, Czech Republic, 360 pp." type="book" year="2014">van Oorschot &amp; Coutsis (2014: 64)</bibRefCitation>
discussed at great length the characters used to distinguish this taxon and considered that they were not sufficiently reliable as diagnostic characters, particularly in view of the fact that the
<typeStatus id="8C798804FFF2FFECE24F00CDFC0EFF72" box="[885,941,151,177]" pageId="10" pageNumber="35">Type</typeStatus>
Locality of
<taxonomicName id="94C24D25FFF2FFECE50600CDFB2EFF73" authorityName="Toth &amp; Varga" authorityYear="2011" box="[1084,1165,151,176]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="zagrosi">
<emphasis id="61B6EAB4FFF2FFECE50600CDFB2EFF73" box="[1084,1165,151,176]" italics="true" pageId="10" pageNumber="35">zagrosi</emphasis>
</taxonomicName>
was in the midst of the distribution of
<taxonomicName id="94C24D25FFF2FFECE00300E1FE4DFF16" baseAuthorityName="Toth &amp; Varga" baseAuthorityYear="2011" box="[313,494,187,213]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="subSpecies" species="ornata" subSpecies="abbas">
<emphasis id="61B6EAB4FFF2FFECE00300E1FE4DFF16" box="[313,494,187,213]" italics="true" pageId="10" pageNumber="35">M. ornata abbas</emphasis>
</taxonomicName>
in the Zagrosi Mountains (
<bibRefCitation id="37534B57FFF2FFECE22800E1FBF0FF16" author="Tshikolovets, V. V. &amp; Naderi A. &amp; Eckweiler, W." box="[786,1107,187,213]" pageId="10" pageNumber="35" refId="ref11652" refString="Tshikolovets, V. V., Naderi A. &amp; Eckweiler, W. (2014) The butterflies of Iran and Iraq. Tshikolovets Publications, Pardubice, Czech Republic, 366 pp. + 65 colour plates." type="book" year="2014">
Tshikolovets
<emphasis id="61B6EAB4FFF2FFECE29D00E1FC75FF16" box="[935,982,187,213]" italics="true" pageId="10" pageNumber="35">et al</emphasis>
. 2014: 320
</bibRefCitation>
, map).Although
<bibRefCitation id="37534B57FFF2FFECE43400E1FEBEFF3A" author="Tshikolovets, V. V. &amp; Naderi A. &amp; Eckweiler, W." pageId="10" pageNumber="35" refId="ref11652" refString="Tshikolovets, V. V., Naderi A. &amp; Eckweiler, W. (2014) The butterflies of Iran and Iraq. Tshikolovets Publications, Pardubice, Czech Republic, 366 pp. + 65 colour plates." type="book" year="2014">
Tshikolovets
<emphasis id="61B6EAB4FFF2FFECE1AD0085FF65FF3A" box="[151,198,223,249]" italics="true" pageId="10" pageNumber="35">et al</emphasis>
. (2014)
</bibRefCitation>
sank
<taxonomicName id="94C24D25FFF2FFECE0630085FE6EFF3B" authorityName="Toth &amp; Varga" authorityYear="2011" box="[345,461,223,248]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="zagrosi">
<emphasis id="61B6EAB4FFF2FFECE0630085FE6EFF3B" box="[345,461,223,248]" italics="true" pageId="10" pageNumber="35">M. zagrosi</emphasis>
</taxonomicName>
in synonymy with
<taxonomicName id="94C24D25FFF2FFECE3A70085FCF0FF3A" baseAuthorityName="Toth &amp; Varga" baseAuthorityYear="2011" box="[669,851,223,249]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="subSpecies" species="ornata" subSpecies="abbas">
<emphasis id="61B6EAB4FFF2FFECE3A70085FCF0FF3A" box="[669,851,223,249]" italics="true" pageId="10" pageNumber="35">M. ornata abbas</emphasis>
</taxonomicName>
, they acknowledged that there is insufficient material to reach a conclusion regarding the status of the two taxa. The present authors’ only additional comment is that the elevation of the
<typeStatus id="8C798804FFF2FFECE068017DFE29FE82" box="[338,394,295,321]" pageId="10" pageNumber="35">Type</typeStatus>
Localities of
<taxonomicName id="94C24D25FFF2FFECE310017DFCFBFE82" baseAuthorityName="Toth &amp; Varga" baseAuthorityYear="2011" box="[554,856,294,321]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="subSpecies" species="ornata" subSpecies="abbas">
<emphasis id="61B6EAB4FFF2FFECE310017DFD4BFE82" box="[554,744,295,321]" italics="true" pageId="10" pageNumber="35">M. ornata abbas</emphasis>
(
<quantity id="943A9B43FFF2FFECE3C2017CFCF3FE82" box="[760,848,294,321]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="10" pageNumber="35" unit="m" value="2250.0">2250 m</quantity>
)
</taxonomicName>
and the taxon
<taxonomicName id="94C24D25FFF2FFECE53F017DFBF5FE83" authorityName="Toth &amp; Varga" authorityYear="2011" box="[1029,1110,295,320]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="zagrosi">
<emphasis id="61B6EAB4FFF2FFECE53F017DFBF5FE83" box="[1029,1110,295,320]" italics="true" pageId="10" pageNumber="35">zagrosi</emphasis>
</taxonomicName>
(
<quantity id="943A9B43FFF2FFECE55D017CFB13FE82" box="[1127,1200,294,321]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="35" unit="m" value="500.0">500 m</quantity>
– Konartakhteh) are very different and environmental differences might account for the differences in wing morphology.
<bibRefCitation id="37534B57FFF2FFECE4390111FEC1FE4A" author="Van Oorschot, H. &amp; Coutsis, J. G." pageId="10" pageNumber="35" refId="ref11693" refString="Van Oorschot, H. &amp; Coutsis, J. G. (2014) The genus Melitaea Fabricius, 1807 (Lepidoptera, Nymphalidae, Nymphalinae) Taxonomy and systematics with special reference to the male genitalia. Tshikolovets publications, Pardubice, Czech Republic, 360 pp." type="book" year="2014">Van Oorschot &amp; Coutsis (2014)</bibRefCitation>
have already suggested that small differences in genitalia measurements could be accounted for by the coverslips compressing the genitalia on the slide. Recently
<bibRefCitation id="37534B57FFF2FFECE28301C9FB24FE6D" author="Toth, J. P. &amp; Varga, Z. &amp; Bereczki, J." box="[953,1159,403,430]" pageId="10" pageNumber="35" pagination="106 - 115" refId="ref11405" refString="Toth, J. P., Varga, Z. &amp; Bereczki, J. (2016) Long-term survival and diversification of an endemic Melitaea species in mountains of Iran and adjacent areas. Journal of Zoological Systematics and Evolutionary Research, 54 (2), 106 - 115. https: // doi. org / 10.1111 / jzs. 12122" type="journal article" year="2016">
Tóth
<emphasis id="61B6EAB4FFF2FFECE2CD01C9FB89FE6E" box="[1015,1066,403,429]" italics="true" pageId="10" pageNumber="35">et al</emphasis>
. (2016)
</bibRefCitation>
, using a combination of morphometric measurements on the genitalia and DNA, elevated the taxon
<taxonomicName id="94C24D25FFF2FFECE2DA01EDFB80FE12" authorityName="Gross &amp; Ebert" authorityYear="1975" box="[992,1059,439,465]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="abbas">
<emphasis id="61B6EAB4FFF2FFECE2DA01EDFB80FE12" box="[992,1059,439,465]" italics="true" pageId="10" pageNumber="35">abbas</emphasis>
</taxonomicName>
to species level and synonymised the taxon
<taxonomicName id="94C24D25FFF2FFECE0390181FEF7FE37" authorityName="Toth &amp; Varga" authorityYear="2011" box="[259,340,475,500]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="zagrosi">
<emphasis id="61B6EAB4FFF2FFECE0390181FEF7FE37" box="[259,340,475,500]" italics="true" pageId="10" pageNumber="35">zagrosi</emphasis>
</taxonomicName>
with it, in contrast to their previous conclusions, which elevated
<taxonomicName id="94C24D25FFF2FFECE5190181FBD7FE37" authorityName="Toth &amp; Varga" authorityYear="2011" box="[1059,1140,475,500]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="zagrosi">
<emphasis id="61B6EAB4FFF2FFECE5190181FBD7FE37" box="[1059,1140,475,500]" italics="true" pageId="10" pageNumber="35">zagrosi</emphasis>
</taxonomicName>
to species level separating it from
<taxonomicName id="94C24D25FFF2FFECE1D201A5FD2EFDDA" authority="(Toth &amp; Varga 2011)" baseAuthorityName="Toth &amp; Varga" baseAuthorityYear="2011" box="[232,653,511,537]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="subSpecies" species="ornata" subSpecies="abbas">
<emphasis id="61B6EAB4FFF2FFECE1D201A5FE03FDDA" box="[232,416,511,537]" italics="true" pageId="10" pageNumber="35">M. ornata abbas</emphasis>
(
<bibRefCitation id="37534B57FFF2FFECE09701A5FD26FDDA" author="Toth, J. P. &amp; Varga, Z." box="[429,645,511,537]" pageId="10" pageNumber="35" pagination="258 - 268" refId="ref11248" refString="Toth, J. P. &amp; Varga, Z. (2011) Inter- and intraspecific variation in the genitalia of the ' Melitaea phoebe group' (Lepidoptera. Nymphalidae). Zoologischer Anzeiger, 250, 258 - 268. [6 pp. of Appendix with key and figures of genitalia only present in the on-line version, www. sciencedirect. com] https: // doi. org / 10.1016 / j. jcz. 2011.05.002" type="book chapter" year="2011">Tóth &amp; Varga 2011</bibRefCitation>
)
</taxonomicName>
, with further research by
<bibRefCitation id="37534B57FFF2FFECE29F01A5FBC5FDDA" author="Toth, J. P. &amp; Bereczki, J. &amp; Varga, Z. &amp; Rota, J. &amp; Sramko, G. &amp; Wahlberg, N." box="[933,1126,511,537]" pageId="10" pageNumber="35" pagination="749 - 757" refId="ref11329" refString="Toth, J. P., Bereczki, J., Varga, Z., Rota, J., Sramko, G. &amp; Wahlberg, N. (2014) Relationships within the Melitaea phoebe species group (Lepidoptera: Nymphalidae): new insights from molecular and morphometric information. Systematic Entomology, 39, 749 - 757. https: // doi. org / 10.1111 / syen. 12083" type="journal article" year="2014">
Tóth
<emphasis id="61B6EAB4FFF2FFECE2E501A5FBACFDDA" box="[991,1039,511,537]" italics="true" pageId="10" pageNumber="35">et al</emphasis>
. (2014)
</bibRefCitation>
confirming this conclusion. However, their current thinking (
<bibRefCitation id="37534B57FFF2FFECE32C0279FD7AFDFD" author="Toth, J. P. &amp; Varga, Z. &amp; Bereczki, J." box="[534,729,547,574]" pageId="10" pageNumber="35" pagination="106 - 115" refId="ref11405" refString="Toth, J. P., Varga, Z. &amp; Bereczki, J. (2016) Long-term survival and diversification of an endemic Melitaea species in mountains of Iran and adjacent areas. Journal of Zoological Systematics and Evolutionary Research, 54 (2), 106 - 115. https: // doi. org / 10.1111 / jzs. 12122" type="journal article" year="2016">
Tóth
<emphasis id="61B6EAB4FFF2FFECE36D0279FD2FFDFE" box="[599,652,547,573]" italics="true" pageId="10" pageNumber="35">et al</emphasis>
. 2016
</bibRefCitation>
), appeared to be that
<taxonomicName id="94C24D25FFF2FFECE2E00279FBE5FDFE" authorityName="Gross &amp; Ebert" authorityYear="1975" box="[986,1094,547,573]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="abbas">
<emphasis id="61B6EAB4FFF2FFECE2E00279FBE5FDFE" box="[986,1094,547,573]" italics="true" pageId="10" pageNumber="35">M. abbas</emphasis>
</taxonomicName>
is a species distinct from
<taxonomicName id="94C24D25FFF2FFECE4440279FF4BFDA2" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFF2FFECE4440279FF4BFDA2" italics="true" pageId="10" pageNumber="35">M. phoebe</emphasis>
</taxonomicName>
,
<taxonomicName id="94C24D25FFF2FFECE1CF021DFEC6FDA3" authorityName=", Russell &amp; Tennent" authorityYear="2016" box="[245,357,583,608]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="61B6EAB4FFF2FFECE1CF021DFEC6FDA3" box="[245,357,583,608]" italics="true" pageId="10" pageNumber="35">M. ornata</emphasis>
</taxonomicName>
and
<taxonomicName id="94C24D25FFF2FFECE0A3021DFDA0FDA2" authorityName="Fruhstorfer" authorityYear="1908" box="[409,515,583,609]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="telona">
<emphasis id="61B6EAB4FFF2FFECE0A3021DFDA0FDA2" box="[409,515,583,609]" italics="true" pageId="10" pageNumber="35">M. telona</emphasis>
</taxonomicName>
and that the taxon
<taxonomicName id="94C24D25FFF2FFECE3EE021DFC86FDA3" authorityName="Toth &amp; Varga" authorityYear="2011" box="[724,805,583,608]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="zagrosi">
<emphasis id="61B6EAB4FFF2FFECE3EE021DFC86FDA3" box="[724,805,583,608]" italics="true" pageId="10" pageNumber="35">zagrosi</emphasis>
</taxonomicName>
is a subspecies of
<taxonomicName id="94C24D25FFF2FFECE2C8021DFBF9FDA2" authorityName="Gross &amp; Ebert" authorityYear="1975" box="[1010,1114,583,609]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="abbas">
<emphasis id="61B6EAB4FFF2FFECE2C8021DFBF9FDA2" box="[1010,1114,583,609]" italics="true" pageId="10" pageNumber="35">M. abbas</emphasis>
</taxonomicName>
. We place
<taxonomicName id="94C24D25FFF2FFECE5E8021DFA80FDA3" authorityName="Toth &amp; Varga" authorityYear="2011" box="[1234,1315,583,608]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="zagrosi">
<emphasis id="61B6EAB4FFF2FFECE5E8021DFA80FDA3" box="[1234,1315,583,608]" italics="true" pageId="10" pageNumber="35">zagrosi</emphasis>
</taxonomicName>
as a junior subjective synonym of
<taxonomicName id="94C24D25FFF2FFECE0A60231FDA7FD46" authorityName="Higgins" authorityYear="1941" box="[412,516,619,645]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="enoch">
<emphasis id="61B6EAB4FFF2FFECE0A60231FDA7FD46" box="[412,516,619,645]" italics="true" pageId="10" pageNumber="35">M. enoch</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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: plate 1 figs 3 &amp; 4). From the photographs of the
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Korb
<emphasis id="61B6EAB4FFF2FFECE5E50319FAB3FC9E" box="[1247,1296,835,861]" italics="true" pageId="10" pageNumber="35">et al</emphasis>
. 2015
</bibRefCitation>
: plate 1: figs 5 &amp; 6). The specimens photographed (
<figureCitation id="CBF92A23FFF2FFECE39E033DFCFDFC42" box="[676,862,871,897]" captionStart="FIGURES 17–20" captionStartId="9.[151,264,1710,1734]" captionTargetBox="[151,1436,381,1687]" captionTargetId="figure-121@9.[151,1436,381,1687]" captionTargetPageId="9" captionText="FIGURES 17–20. 17, Holotype of Melitaea phoebe abbas, Gross &amp; Ebert, 1975. © R. Trusch. 18, Allotype of Melitaea phoebe abbas, Gross &amp; Ebert, 1975. © Alipanah. 19, Holotype of Melitaea ornata adversaria Korb, Stradomsky &amp; Kuznetsov, 2015. 20, Paratype of Melitaea ornata reliquiae Korb, Stradomsky &amp; Kuznetsov, 2015. Figs 19, 20 © S.K. Korb, B.V. Stradomsky &amp; G.V. Kuznetsov.A = upperside, B = underside, C = labels on specimen pin." figureDoi="http://doi.org/10.5281/zenodo.6577646" httpUri="https://zenodo.org/record/6577646/files/figure.png" pageId="10" pageNumber="35">Figs 20 A, B, C</figureCitation>
) clearly show the wing morphology and spatulate antennae, characteristics of
<taxonomicName id="94C24D25FFF2FFECE0F403D1FD9DFC67" authorityName=", Russell &amp; Tennent" authorityYear="2016" box="[462,574,907,932]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="61B6EAB4FFF2FFECE0F403D1FD9DFC67" box="[462,574,907,932]" italics="true" pageId="10" pageNumber="35">M. ornata</emphasis>
</taxonomicName>
rather than
<taxonomicName id="94C24D25FFF2FFECE3FF03D1FC98FC66" box="[709,827,907,933]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFF2FFECE3FF03D1FC98FC66" box="[709,827,907,933]" italics="true" pageId="10" pageNumber="35">M. phoebe</emphasis>
</taxonomicName>
.
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<treatment id="DB6B87B0FFF9FFE5E1FD0625FDBCFE6E" ID-DOI="http://doi.org/10.5281/zenodo.6580816" ID-GBIF-Taxon="195626723" ID-Zenodo-Dep="6580816" LSID="urn:lsid:plazi:treatment:DB6B87B0FFF9FFE5E1FD0625FDBCFE6E" httpUri="http://treatment.plazi.org/id/DB6B87B0FFF9FFE5E1FD0625FDBCFE6E" lastPageId="3" lastPageNumber="28" pageId="1" pageNumber="26">
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<paragraph id="537D36A6FFF9FFE7E1FD0625FDEBF95A" blockId="1.[151,1437,870,2013]" box="[199,584,1662,1689]" pageId="1" pageNumber="26">
<taxonomicName id="94C24D25FFF9FFE7E1FD0625FDEBF95A" ID-CoL="6QXTY" authority="Alpheraky, 1881" authorityName="Alpheraky" authorityYear="1881" box="[199,584,1662,1689]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="sibina">
<emphasis id="61B6EAB4FFF9FFE7E1FD0625FE28F95A" bold="true" box="[199,395,1662,1689]" italics="true" pageId="1" pageNumber="26">M. phoebe sibina</emphasis>
Alphéraky, 1881
</taxonomicName>
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<subSubSection id="1BD8652DFFF9FFE5E3740625FDBCFE6E" lastPageId="3" lastPageNumber="28" pageId="1" pageNumber="26" type="discussion">
<paragraph id="537D36A6FFF9FFE7E3740625FADCF81E" blockId="1.[151,1437,870,2013]" pageId="1" pageNumber="26">
[TL: Peoples
<collectingCountry id="2BD57636FFF9FFE7E3DE0625FC11F95A" box="[740,946,1663,1689]" name="Taiwan" pageId="1" pageNumber="26">Republic of China</collectingCountry>
, NW Xinjiang, Yining, Sibo, Ili Valley]. The locality data label (see
<figureCitation id="CBF92A23FFF9FFE7E0A106F9FE52F97E" box="[411,497,1699,1725]" captionStart="FIGURES 7–11" captionStartId="4.[151,264,1857,1881]" captionTargetBox="[152,1436,560,1833]" captionTargetId="figure-219@4.[151,1436,560,1833]" captionTargetPageId="4" captionText="FIGURES 7–11. 7, Lectotype of Melitaea phoebe sibina Alphéraky, 1881. 8, Melitaea phoebe ab. tatara Krulikovsky, 1891. 9, Lectotype of Melitaea phoebe ornata Christoph, 1893. 10, Lectotype of Melitaea phoebe uralensis Krulikovsky, 1897 (= ufensis Krulikovsky 1902). 11, Lectotype of Melitaea phoebe var. tungusa Herz, 1899. Figs 7, 9, 11 © V.A. Lukhtanov. Figs 8, 10 © expired. A = upperside, B = underside, C = labels on specimen pin." figureDoi="http://doi.org/10.5281/zenodo.6577635" httpUri="https://zenodo.org/record/6577635/files/figure.png" pageId="1" pageNumber="26">Fig. 7C</figureCitation>
) has the location in Russian ‘Kuldscha, Kainak’, since this was Russian territory in 1879)]; for some considerable time this taxon has been given specific status, as
<taxonomicName id="94C24D25FFF9FFE7E53E069DFB12F922" authorityName="Alpheraky" authorityYear="1881" box="[1028,1201,1735,1761]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="sibina">
<emphasis id="61B6EAB4FFF9FFE7E53E069DFB12F922" box="[1028,1201,1735,1761]" italics="true" pageId="1" pageNumber="26">Melitaea sibina</emphasis>
</taxonomicName>
, for example: Grum- Grshimailo, 1895: 291,
<bibRefCitation id="37534B57FFF9FFE7E09C06B1FDC4F8C6" author="Schultz, O." box="[422,615,1771,1797]" pageId="1" pageNumber="26" pagination="90 - 92" refId="ref11069" refString="Schultz, O. (1908) Ueber einige Farbungs- und Zeichnungs- anomalien aus dem Genus Melitaea F. Entomologische Zeitschrift, 22, 90 - 92." type="journal article" year="1908">Schultz 1908: 91</bibRefCitation>
and
<bibRefCitation id="37534B57FFF9FFE7E39A06B1FCDEF8C6" author="Higgins, L. G." box="[672,893,1771,1797]" pageId="1" pageNumber="26" pagination="175 - 365" refId="ref9372" refString="Higgins, L. G. (1941) An illustrated catalogue of the Palearctic Melitaea (Lep. Rhopalocera). Transactions of the Royal Entomological Society of London, 91, 175 - 365. https: // doi. org / 10.1111 / j. 1365 - 2311.1941. tb 01045. x" type="journal article" year="1941">Higgins, 1941: 347</bibRefCitation>
. However,
<bibRefCitation id="37534B57FFF9FFE7E2C406B1FB4FF8C6" author="Toth, J. P. &amp; Varga, Z." box="[1022,1260,1771,1797]" pageId="1" pageNumber="26" pagination="258 - 268" refId="ref11248" refString="Toth, J. P. &amp; Varga, Z. (2011) Inter- and intraspecific variation in the genitalia of the ' Melitaea phoebe group' (Lepidoptera. Nymphalidae). Zoologischer Anzeiger, 250, 258 - 268. [6 pp. of Appendix with key and figures of genitalia only present in the on-line version, www. sciencedirect. com] https: // doi. org / 10.1016 / j. jcz. 2011.05.002" type="book chapter" year="2011">Tóth &amp; Varga (2011)</bibRefCitation>
were unable to separate
<taxonomicName id="94C24D25FFF9FFE7E1C30755FEC1F8EA" authorityName="Alpheraky" authorityYear="1881" box="[249,354,1807,1833]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="sibina">
<emphasis id="61B6EAB4FFF9FFE7E1C30755FEC1F8EA" box="[249,354,1807,1833]" italics="true" pageId="1" pageNumber="26">M. sibina</emphasis>
</taxonomicName>
from
<taxonomicName id="94C24D25FFF9FFE7E09C0755FDBFF8EA" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[422,540,1807,1833]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFF9FFE7E09C0755FDBFF8EA" box="[422,540,1807,1833]" italics="true" pageId="1" pageNumber="26">M. phoebe</emphasis>
</taxonomicName>
using morphometric measurements of male and female genitalia and
<bibRefCitation id="37534B57FFF9FFE7E4190755FE82F88E" author="Lukhtanov, V. A. &amp; Sourakov, A. &amp; Zakharov, E. V. &amp; Hebert, P. D. N." pageId="1" pageNumber="26" pagination="1302 - 1310" refId="ref10220" refString="Lukhtanov, V. A., Sourakov, A., Zakharov, E. V. &amp; Hebert, P. D. N. (2009) DNA barcoding Central Asian butterflies: increasing geographical dimension does not significantly reduce the success of species identification. Molecular Ecology Resources, 9, 1302 - 1310. https: // doi. org / 10.1111 / j. 1755 - 0998.2009.02577. x" type="journal article" year="2009">
Lukhtanov
<emphasis id="61B6EAB4FFF9FFE7E1AD0769FF64F88E" box="[151,199,1843,1869]" italics="true" pageId="1" pageNumber="26">et al</emphasis>
. (2009)
</bibRefCitation>
,
<bibRefCitation id="37534B57FFF9FFE7E0160769FDB9F88E" author="Leneveu J. &amp; Chichvarkhin, A. &amp; Wahlberg, N." box="[300,538,1843,1869]" pageId="1" pageNumber="26" pagination="346 - 361" refId="ref10155" refString="Leneveu J., Chichvarkhin, A. &amp; Wahlberg, N. (2009) Varying rates of diversification in the genus Melitaea (Lepidoptera: Nymphalidae) during the past 20 million years. Biological Journal of the Linnean Society, 97, 346 - 361. https: // doi. org / 10.1111 / j. 1095 - 8312.2009.01208. x" type="journal article" year="2009">
Leneveu
<emphasis id="61B6EAB4FFF9FFE7E0A80769FE6AF88E" box="[402,457,1843,1869]" italics="true" pageId="1" pageNumber="26">et al.</emphasis>
(2009)
</bibRefCitation>
and
<bibRefCitation id="37534B57FFF9FFE7E3770769FCB3F88E" author="Toth, J. P. &amp; Bereczki, J. &amp; Varga, Z. &amp; Rota, J. &amp; Sramko, G. &amp; Wahlberg, N." box="[589,784,1843,1869]" pageId="1" pageNumber="26" pagination="749 - 757" refId="ref11329" refString="Toth, J. P., Bereczki, J., Varga, Z., Rota, J., Sramko, G. &amp; Wahlberg, N. (2014) Relationships within the Melitaea phoebe species group (Lepidoptera: Nymphalidae): new insights from molecular and morphometric information. Systematic Entomology, 39, 749 - 757. https: // doi. org / 10.1111 / syen. 12083" type="journal article" year="2014">
Tóth
<emphasis id="61B6EAB4FFF9FFE7E3B20769FD1AF88E" box="[648,697,1843,1869]" italics="true" pageId="1" pageNumber="26">et al</emphasis>
. (2014)
</bibRefCitation>
were also unable to separate these species using molecular data. The present authors consider that
<taxonomicName id="94C24D25FFF9FFE7E36D070DFCA4F8B2" authorityName="Alpheraky" authorityYear="1881" box="[599,775,1879,1905]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="sibina">
<emphasis id="61B6EAB4FFF9FFE7E36D070DFCA4F8B2" box="[599,775,1879,1905]" italics="true" pageId="1" pageNumber="26">Melitaea sibina</emphasis>
</taxonomicName>
(
<figureCitation id="CBF92A23FFF9FFE7E222070DFC18F8B2" box="[792,955,1879,1905]" captionStart="FIGURES 7–11" captionStartId="4.[151,264,1857,1881]" captionTargetBox="[152,1436,560,1833]" captionTargetId="figure-219@4.[151,1436,560,1833]" captionTargetPageId="4" captionText="FIGURES 7–11. 7, Lectotype of Melitaea phoebe sibina Alphéraky, 1881. 8, Melitaea phoebe ab. tatara Krulikovsky, 1891. 9, Lectotype of Melitaea phoebe ornata Christoph, 1893. 10, Lectotype of Melitaea phoebe uralensis Krulikovsky, 1897 (= ufensis Krulikovsky 1902). 11, Lectotype of Melitaea phoebe var. tungusa Herz, 1899. Figs 7, 9, 11 © V.A. Lukhtanov. Figs 8, 10 © expired. A = upperside, B = underside, C = labels on specimen pin." figureDoi="http://doi.org/10.5281/zenodo.6577635" httpUri="https://zenodo.org/record/6577635/files/figure.png" pageId="1" pageNumber="26">Figs 7A, B, C</figureCitation>
) should be considered as a closely related species separate from
<taxonomicName id="94C24D25FFF9FFE7E0AC0721FDACF856" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[406,527,1915,1941]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFF9FFE7E0AC0721FDACF856" box="[406,527,1915,1941]" italics="true" pageId="1" pageNumber="26">M. phoebe</emphasis>
</taxonomicName>
due to its significantly different wing pattern, in agreement with
<bibRefCitation id="37534B57FFF9FFE7E43E0721FE2FF87A" author="Van Oorschot, H. &amp; Coutsis, J. G." pageId="1" pageNumber="26" refId="ref11693" refString="Van Oorschot, H. &amp; Coutsis, J. G. (2014) The genus Melitaea Fabricius, 1807 (Lepidoptera, Nymphalidae, Nymphalinae) Taxonomy and systematics with special reference to the male genitalia. Tshikolovets publications, Pardubice, Czech Republic, 360 pp." type="book" year="2014">van Oorschot &amp; Coutsis (2014: 65)</bibRefCitation>
and
<bibRefCitation id="37534B57FFF9FFE7E0F207C5FD24F87A" author="Korb, S. K." box="[456,647,1950,1977]" pageId="1" pageNumber="26" pagination="53 - 60" refId="ref9650" refString="Korb, S. K. (2018) Butterfly fauna of Dzhumagal-Too mountain ridge in Kyrgyzstan (Lepidoptera: Papilionidea and Hesperioidea), Acta Biologica Sibirica, 4 (2), 53 - 60. https: // doi. org / 10.14258 / abs. v 4 i 2.4124" type="journal article" year="2018">Korb (2018: 58)</bibRefCitation>
. As a point of interest the label in Russian (
<figureCitation id="CBF92A23FFF9FFE7E5BD07C5FB40F87A" box="[1159,1251,1951,1977]" captionStart="FIGURES 7–11" captionStartId="4.[151,264,1857,1881]" captionTargetBox="[152,1436,560,1833]" captionTargetId="figure-219@4.[151,1436,560,1833]" captionTargetPageId="4" captionText="FIGURES 7–11. 7, Lectotype of Melitaea phoebe sibina Alphéraky, 1881. 8, Melitaea phoebe ab. tatara Krulikovsky, 1891. 9, Lectotype of Melitaea phoebe ornata Christoph, 1893. 10, Lectotype of Melitaea phoebe uralensis Krulikovsky, 1897 (= ufensis Krulikovsky 1902). 11, Lectotype of Melitaea phoebe var. tungusa Herz, 1899. Figs 7, 9, 11 © V.A. Lukhtanov. Figs 8, 10 © expired. A = upperside, B = underside, C = labels on specimen pin." figureDoi="http://doi.org/10.5281/zenodo.6577635" httpUri="https://zenodo.org/record/6577635/files/figure.png" pageId="1" pageNumber="26">Fig. 7C</figureCitation>
): ‘Кол.[лекциЯ] Вел.[икого] Кн.[ЯЗЯ]/ НиколаЯ/ Михайловича’ translates to ‘Collection of Grand Duke Nikolai Mikhailovich’.
</paragraph>
<caption id="07BD662EFFFAFFE4E1AD06EFFB05F842" ID-DOI="http://doi.org/10.5281/zenodo.6577631" ID-Zenodo-Dep="6577631" httpUri="https://zenodo.org/record/6577631/files/figure.png" pageId="2" pageNumber="27" startId="2.[151,264,1717,1741]" targetBox="[153,1436,200,1692]" targetPageId="2">
<paragraph id="537D36A6FFFAFFE4E1AD06EFFB05F842" blockId="2.[151,1437,1717,1924]" pageId="2" pageNumber="27">
<emphasis id="61B6EAB4FFFAFFE4E1AD06EFFE9AF90E" bold="true" box="[151,313,1717,1741]" pageId="2" pageNumber="27">FIGURES 1–6.</emphasis>
1,
<taxonomicName id="94C24D25FFFAFFE4E06C06EFFE59F90E" authorityName="Christoph" authorityYear="1893" box="[342,506,1717,1741]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="27" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="61B6EAB4FFFAFFE4E06C06EFFE59F90E" box="[342,506,1717,1741]" italics="true" pageId="2" pageNumber="27">Melitaea ornata</emphasis>
</taxonomicName>
male underside, Russian Federation, southern Urals, Orenburg Province, Guberli,
<emphasis id="61B6EAB4FFFAFFE4E40806ECFAE9F90E" box="[1330,1354,1718,1741]" italics="true" pageId="2" pageNumber="27">ca</emphasis>
. 190 m, 14 May 2014. 2,
<taxonomicName id="94C24D25FFFAFFE4E0720683FE51F932" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[328,498,1753,1777]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="27" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFAFFE4E0720683FE51F932" box="[328,498,1753,1777]" italics="true" pageId="2" pageNumber="27">Melitaea phoebe</emphasis>
</taxonomicName>
male underside, Russian Federation, southern Urals, Orenburg Province, Krasnoznamenka,
<emphasis id="61B6EAB4FFFAFFE4E1AD06A4FF0CF8D6" box="[151,175,1790,1813]" italics="true" pageId="2" pageNumber="27">ca</emphasis>
. 210 m, 16 May 2014. 3,
<taxonomicName id="94C24D25FFFAFFE4E08C06A7FDF8F8D6" authorityName="Christoph" authorityYear="1893" box="[438,603,1789,1813]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="27" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="61B6EAB4FFFAFFE4E08C06A7FDF8F8D6" box="[438,603,1789,1813]" italics="true" pageId="2" pageNumber="27">Melitaea ornata</emphasis>
</taxonomicName>
female underside, Slovenia, Koper,
<emphasis id="61B6EAB4FFFAFFE4E2F606A4FC49F8D6" box="[972,1002,1790,1813]" italics="true" pageId="2" pageNumber="27">ca.</emphasis>
2km NW of Rakitovec
<emphasis id="61B6EAB4FFFAFFE4E5E706A4FB58F8D6" box="[1245,1275,1790,1813]" italics="true" pageId="2" pageNumber="27">ca.</emphasis>
525 m, 17 May 2011. 4,
<taxonomicName id="94C24D25FFFAFFE4E1CB077BFE3FF8FA" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[241,412,1825,1849]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="27" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFAFFE4E1CB077BFE3FF8FA" box="[241,412,1825,1849]" italics="true" pageId="2" pageNumber="27">Melitaea phoebe</emphasis>
</taxonomicName>
female underside, Slovenia, Koper,
<emphasis id="61B6EAB4FFFAFFE4E2280778FC92F8FA" box="[786,817,1826,1849]" italics="true" pageId="2" pageNumber="27">ca.</emphasis>
2km S. of Rakitovec
<emphasis id="61B6EAB4FFFAFFE4E5290778FB91F8FA" box="[1043,1074,1826,1849]" italics="true" pageId="2" pageNumber="27">ca.</emphasis>
525 m, 17 May 2011. 5, Lectotype female of
<taxonomicName id="94C24D25FFFAFFE4E1C4071FFD6AF89E" authority="Staudinger, 1870" authorityName="Staudinger" authorityYear="1870" box="[254,713,1861,1885]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="27" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="caucasica">
<emphasis id="61B6EAB4FFFAFFE4E1C4071FFDB0F89E" box="[254,531,1861,1885]" italics="true" pageId="2" pageNumber="27">Melitaea phoebe caucasica</emphasis>
<bibRefCitation id="37534B57FFFAFFE4E323071FFD6AF89E" author="Staudinger, O." box="[537,713,1861,1885]" pageId="2" pageNumber="27" pagination="3" refId="ref11169" refString="Staudinger, O. (1870) Beitrag zur Lepidopterenfauna Griechenlands. Horae Societatis Entomologicae Rossicae, 7, [3] - 304." type="journal article" year="1870">Staudinger, 1870</bibRefCitation>
</taxonomicName>
. 6, Paralectotype male of
<taxonomicName id="94C24D25FFFAFFE4E2F4071FFA3AF89E" authority="Staudinger, 1870" authorityName="Staudinger" authorityYear="1870" box="[974,1433,1861,1885]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="27" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="caucasica">
<emphasis id="61B6EAB4FFFAFFE4E2F4071FFB40F89E" box="[974,1251,1861,1885]" italics="true" pageId="2" pageNumber="27">Melitaea phoebe caucasica</emphasis>
<bibRefCitation id="37534B57FFFAFFE4E5D0071FFA3AF89E" author="Staudinger, O." box="[1258,1433,1861,1885]" pageId="2" pageNumber="27" pagination="3" refId="ref11169" refString="Staudinger, O. (1870) Beitrag zur Lepidopterenfauna Griechenlands. Horae Societatis Entomologicae Rossicae, 7, [3] - 304." type="journal article" year="1870">Staudinger, 1870</bibRefCitation>
</taxonomicName>
. Figs 1–4 © P. Russell. Figs 5–6 © T. Léger. A = upperside, B = underside, C = labels on specimen pin.
</paragraph>
</caption>
<paragraph id="537D36A6FFFBFFE5E1FD00CDFDBCFE6E" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="28">
It is noted that
<bibRefCitation id="37534B57FFFBFFE5E04F00CDFD8DFF71" author="Korb, S. K." box="[373,558,151,178]" pageId="3" pageNumber="28" pagination="61 - 62" refId="ref9613" refString="Korb, S. K. (1998) Some corrections on the diurnal butterflies (Lepidoptera: Rhopalocera) from Xinjiang Province, China. Entomotaxonomia, 20 (1), 61 - 62." type="journal article" year="1998">Korb (1998: 62)</bibRefCitation>
synonymised
<taxonomicName id="94C24D25FFFBFFE5E3E900CDFAADFF72" authority="Huang &amp; Murayama, 1992" authorityName="Huang &amp; Murayama" authorityYear="1992" box="[723,1294,151,177]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="subSpecies" species="scotosia" subSpecies="weiwueria">
<emphasis id="61B6EAB4FFFBFFE5E3E900CDFC70FF73" box="[723,979,151,176]" italics="true" pageId="3" pageNumber="28">M. scotosia weiwueria</emphasis>
<bibRefCitation id="37534B57FFFBFFE5E2E600CDFAADFF72" author="Huang, R. - X. &amp; Murayama, S." box="[988,1294,151,177]" pageId="3" pageNumber="28" pagination="1 - 22" refId="ref9428" refString="Huang, R. - X. &amp; Murayama, S. (1992) Butterflies of Xinjiang Province, China. Tyo to Ga, 43 (1), 1 - 22." type="journal article" year="1992">Huang &amp; Murayama, 1992</bibRefCitation>
</taxonomicName>
[TL:
<collectingCountry id="2BD57636FFFBFFE5E46800CDFA3BFF72" box="[1362,1432,151,177]" name="China" pageId="3" pageNumber="28">China</collectingCountry>
,
<collectingRegion id="9106F844FFFBFFE5E1AD00E1FF5FFF16" box="[151,252,187,213]" country="China" name="Xinjiang Uygur" pageId="3" pageNumber="28">Xinjiang</collectingRegion>
, Altai, Jiadengu], with nominate
<taxonomicName id="94C24D25FFFBFFE5E34E00E1FC29FF16" authority="Butler, 1878" authorityName="Butler" authorityYear="1878" box="[628,906,187,213]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="scotosia">
<emphasis id="61B6EAB4FFFBFFE5E34E00E1FD55FF17" box="[628,758,187,212]" italics="true" pageId="3" pageNumber="28">M. scotosia</emphasis>
<bibRefCitation id="37534B57FFFBFFE5E3C400E1FC29FF16" author="Butler, A. G." box="[766,906,187,213]" pageId="3" pageNumber="28" pagination="281 - 286" refId="ref8862" refString="Butler, A. G. (1878) On some butterflies recently sent home from Japan by Mr. Montague Fenton. Cistula Entomologica, 2 (19), 281 - 286." type="journal article" year="1878">Butler, 1878</bibRefCitation>
</taxonomicName>
and
<bibRefCitation id="37534B57FFFBFFE5E2F800E1FB2BFF15" author="Lang, S. - Y." box="[962,1160,187,214]" pageId="3" pageNumber="28" refId="ref10103" refString="Lang, S. - Y. (2012) The Nymphalidae of China (Lepidoptera: Rhopalocera) Part 1: Lebytheinae, Calinaginae, Morphinae, Heliconiinae, Nymplalinae, Charaxinae, Apaturinae, Cyrestinae, Biblidinae, Limenitidae. Tshikolovets Publications Pardubice, Czech Republic, 443 pp." type="book" year="2012">Lang (2012: 160)</bibRefCitation>
synonymised it with
<taxonomicName id="94C24D25FFFBFFE5E44400E1FD80FF3A" authority="Higgins (1940: 53)" authorityName="Higgins" authorityPageNumber="53" authorityYear="1940" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="subSpecies" species="scotosia" subSpecies="butleri">
<emphasis id="61B6EAB4FFFBFFE5E44400E1FEE6FF3A" italics="true" pageId="3" pageNumber="28">M. scotosia butleri</emphasis>
<bibRefCitation id="37534B57FFFBFFE5E0710085FD80FF3A" author="Higgins, L. G." box="[331,547,222,249]" pageId="3" pageNumber="28" pagination="51 - 53" refId="ref9340" refString="Higgins, L. G. (1940) A new species and two new subspecies of Melitaea (Lep. Nymphalidae). Entomologist, 73, 51 - 53." type="journal article" year="1940">Higgins (1940: 53)</bibRefCitation>
</taxonomicName>
. From Huang &amp; Murayama’s monochrome figure of a worn specimen (1992: 8, fig. 17 no. 19), it is almost impossible to provide specific identification. However,
<bibRefCitation id="37534B57FFFBFFE5E57C0159FA3FFEDE" author="Korb, S. K. &amp; Bolshakov, L. V." box="[1094,1436,259,285]" pageId="3" pageNumber="28" pagination="1 - 123" refId="ref9703" refString="Korb, S. K. &amp; Bolshakov, L. V. (2011) A catalogue of butterflies (Lepidoptera: Papilioformes) of the former USSR. Second edition, reformatted and updated. Eversmannia Supplement 2, 1 - 123. [In Russian]" type="journal article" year="2011">Korb &amp; Bolshakov (2011: 42)</bibRefCitation>
and
<bibRefCitation id="37534B57FFFBFFE5E1F2017DFDF4FE82" author="Van Oorschot, H. &amp; Coutsis, J. G." box="[200,599,294,321]" pageId="3" pageNumber="28" refId="ref11693" refString="Van Oorschot, H. &amp; Coutsis, J. G. (2014) The genus Melitaea Fabricius, 1807 (Lepidoptera, Nymphalidae, Nymphalinae) Taxonomy and systematics with special reference to the male genitalia. Tshikolovets publications, Pardubice, Czech Republic, 360 pp." type="book" year="2014">van Oorschot &amp; Coutsis (2014: 65)</bibRefCitation>
placed this taxon in synonymy with
<taxonomicName id="94C24D25FFFBFFE5E2C1017DFBC5FE82" authorityName="Alpheraky" authorityYear="1881" box="[1019,1126,295,321]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="sibina">
<emphasis id="61B6EAB4FFFBFFE5E2C1017DFBC5FE82" box="[1019,1126,295,321]" italics="true" pageId="3" pageNumber="28">M. sibina</emphasis>
</taxonomicName>
, which seems to be a more plausible interpretation (although not indisputable, possibly requiring further investigation), since the distance to Altai in
<collectingRegion id="9106F844FFFBFFE5E1C90135FE1DFE4A" box="[243,446,367,393]" country="China" name="Xinjiang Uygur" pageId="3" pageNumber="28">Xinjiang province</collectingRegion>
is more than
<quantity id="943A9B43FFFBFFE5E36C0135FD1FFE4A" box="[598,700,367,393]" metricMagnitude="6" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="28" unit="km" value="1000.0">1000 km</quantity>
. northwest of the widely recognized distribution of
<taxonomicName id="94C24D25FFFBFFE5E5C40135FA23FE4B" authorityName="Butler" authorityYear="1878" box="[1278,1408,367,392]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="scotosia">
<emphasis id="61B6EAB4FFFBFFE5E5C40135FA23FE4B" box="[1278,1408,367,392]" italics="true" pageId="3" pageNumber="28">M. scotosia</emphasis>
</taxonomicName>
in northeast
<collectingCountry id="2BD57636FFFBFFE5E03F01C9FEE8FE6E" box="[261,331,403,429]" name="China" pageId="3" pageNumber="28">China</collectingCountry>
,
<collectingCountry id="2BD57636FFFBFFE5E06C01C9FE3EFE6E" box="[342,413,403,429]" name="South Korea" pageId="3" pageNumber="28">Korea</collectingCountry>
, and
<collectingCountry id="2BD57636FFFBFFE5E0E201C9FDB8FE6E" box="[472,539,403,429]" name="Japan" pageId="3" pageNumber="28">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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170
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Identification of
<taxonomicName id="94C24D25FFF9FFE7E06400CDFE72FF72" ID-CoL="3ZKDK" authorityName="Christoph" authorityYear="1893" box="[350,465,151,177]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="ornata">
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and
<taxonomicName id="94C24D25FFF9FFE7E33700CDFD26FF72" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[525,645,150,177]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFF9FFE7E33700CDFD26FF72" bold="true" box="[525,645,150,177]" italics="true" pageId="1" pageNumber="26">M. phoebe</emphasis>
</taxonomicName>
based on underside wing morphology and antennae
</emphasis>
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Identification of individual museum specimens of
<taxonomicName id="94C24D25FFF9FFE7E3D000E1FC96FF17" authorityName="Christoph" authorityYear="1893" box="[746,821,187,212]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="61B6EAB4FFF9FFE7E3D000E1FC96FF17" box="[746,821,187,212]" italics="true" pageId="1" pageNumber="26">ornata</emphasis>
</taxonomicName>
and
<taxonomicName id="94C24D25FFF9FFE7E25F00E1FC15FF16" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[869,950,187,213]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFF9FFE7E25F00E1FC15FF16" box="[869,950,187,213]" italics="true" pageId="1" pageNumber="26">phoebe</emphasis>
</taxonomicName>
may never be achieved with 100% accuracy; although there are morphological features to accommodate identification in many cases, accurate identification requires an examination of their respective late instar larvae. However, the shape of the submarginal markings on the wing undersides and the tips of the antenna usually provides a very good indication of species, particularly when there is a series of
<typeStatus id="8C798804FFF9FFE7E05A0111FE61FEA6" box="[352,450,331,357]" pageId="1" pageNumber="26" type="syntype">syntypes</typeStatus>
from the same population, taken at the same time (cf.
<bibRefCitation id="37534B57FFF9FFE7E52B0111FB4DFEA6" author="Toth, J. P. &amp; Varga, Z." box="[1041,1262,331,357]" pageId="1" pageNumber="26" pagination="258 - 268" refId="ref11248" refString="Toth, J. P. &amp; Varga, Z. (2011) Inter- and intraspecific variation in the genitalia of the ' Melitaea phoebe group' (Lepidoptera. Nymphalidae). Zoologischer Anzeiger, 250, 258 - 268. [6 pp. of Appendix with key and figures of genitalia only present in the on-line version, www. sciencedirect. com] https: // doi. org / 10.1016 / j. jcz. 2011.05.002" type="book chapter" year="2011">Tóth &amp; Varga, 2011</bibRefCitation>
: appendix only;
<bibRefCitation id="37534B57FFF9FFE7E1AD0135FED3FE4A" author="Russell, P. J. C. &amp; Bartolozzi, L. &amp; Hawkins, R. L. &amp; Tennent, W. J. &amp; Leger, T." box="[151,368,367,393]" pageId="1" pageNumber="26" pagination="449 - 472" refId="ref10659" refString="Russell, P. J. C., Bartolozzi, L. Hawkins, R. L., Tennent, W. J. &amp; Leger, T. (2020) Designation of lectotypes for some Spanish and other western European Melitaea taxa, some with mixed syntypic series of M. phoebe ([Denis &amp; Schiffermuller], 1775) and M. ornata Christoph, 1893 (Lepidoptera: Nymphalidae). SHILAP Revista de lepidopterologia, 48, 449 - 472." type="journal article" year="2020">
Russell
<emphasis id="61B6EAB4FFF9FFE7E1C80135FE87FE4A" box="[242,292,367,393]" italics="true" pageId="1" pageNumber="26">et al</emphasis>
. 2020
</bibRefCitation>
). These differences can be seen on figures 1–4, which show the undersides of male specimens from two sympatric and partially synchronic populations of
<taxonomicName id="94C24D25FFF9FFE7E27701C9FC63FE6F" authorityName="Christoph" authorityYear="1893" box="[845,960,403,428]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="61B6EAB4FFF9FFE7E27701C9FC63FE6F" box="[845,960,403,428]" italics="true" pageId="1" pageNumber="26">M. ornata</emphasis>
</taxonomicName>
and
<taxonomicName id="94C24D25FFF9FFE7E2C601C9FBD6FE6E" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[1020,1141,403,429]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFF9FFE7E2C601C9FBD6FE6E" box="[1020,1141,403,429]" italics="true" pageId="1" pageNumber="26">M. phoebe</emphasis>
</taxonomicName>
from the Southern Urals,
<collectingCountry id="2BD57636FFF9FFE7E1AD01EDFED1FE12" box="[151,370,439,465]" name="Russia" pageId="1" pageNumber="26">Russian Federation</collectingCountry>
and females from similar populations near Rakitovec,
<collectingCountry id="2BD57636FFF9FFE7E2D501EDFBF6FE12" box="[1007,1109,439,465]" name="Slovenia" pageId="1" pageNumber="26">Slovenia</collectingCountry>
. Identifications of
<taxonomicName id="94C24D25FFF9FFE7E41401EDFA3FFE12" authorityName="Fruhstorfer" authorityYear="1908" box="[1326,1436,439,465]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="telona">
<emphasis id="61B6EAB4FFF9FFE7E41401EDFA3FFE12" box="[1326,1436,439,465]" italics="true" pageId="1" pageNumber="26">M. telona</emphasis>
</taxonomicName>
and
<taxonomicName id="94C24D25FFF9FFE7E1FC0181FE25FE36" authorityName=", Gross &amp; Ebert" authorityYear="1975" box="[198,390,475,501]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="abbas">
<emphasis id="61B6EAB4FFF9FFE7E1FC0181FE25FE36" box="[198,390,475,501]" italics="true" pageId="1" pageNumber="26">M. phoebe abbas</emphasis>
</taxonomicName>
were based solely on morphometric measurements of genitalia and the biochemical results of
<bibRefCitation id="37534B57FFF9FFE7E1AD01A5FEF0FDDA" author="Toth, J. P. &amp; Bereczki, J. &amp; Varga, Z. &amp; Rota, J. &amp; Sramko, G. &amp; Wahlberg, N." box="[151,339,511,537]" pageId="1" pageNumber="26" pagination="749 - 757" refId="ref11329" refString="Toth, J. P., Bereczki, J., Varga, Z., Rota, J., Sramko, G. &amp; Wahlberg, N. (2014) Relationships within the Melitaea phoebe species group (Lepidoptera: Nymphalidae): new insights from molecular and morphometric information. Systematic Entomology, 39, 749 - 757. https: // doi. org / 10.1111 / syen. 12083" type="journal article" year="2014">
Tóth
<emphasis id="61B6EAB4FFF9FFE7E1E801A5FEA1FDDA" box="[210,258,511,537]" italics="true" pageId="1" pageNumber="26">et al</emphasis>
. (2014
</bibRefCitation>
,
<bibRefCitation id="37534B57FFF9FFE7E06701A5FE36FDDA" author="Toth, J. P. &amp; Varga, Z. &amp; Bereczki, J." box="[349,405,511,537]" pageId="1" pageNumber="26" pagination="106 - 115" refId="ref11405" refString="Toth, J. P., Varga, Z. &amp; Bereczki, J. (2016) Long-term survival and diversification of an endemic Melitaea species in mountains of Iran and adjacent areas. Journal of Zoological Systematics and Evolutionary Research, 54 (2), 106 - 115. https: // doi. org / 10.1111 / jzs. 12122" type="journal article" year="2016">2016</bibRefCitation>
&amp;
<bibRefCitation id="37534B57FFF9FFE7E08C01A5FE50FDDA" author="Toth, J. P. &amp; Varga, Z. &amp; Verovnik, R. &amp; Wahlberg, N. &amp; Varadi, A. &amp; Bereczki, J." box="[438,499,511,537]" pageId="1" pageNumber="26" pagination="267 - 281" refId="ref11547" refString="Toth, J. P., Varga, Z., Verovnik, R., Wahlberg, N., Varadi, A. &amp; Bereczki, J. (2017) Mito-nuclear discordance helps to reveal the phylogeographic patterns of Melitaea ornata (Lepidoptera: Nymphalidae). Biological Journal of the Linnean Society, 121, 267 - 281. https: // doi. org / 10.1093 / biolinnean / blw 037" type="journal article" year="2017">2017</bibRefCitation>
). DNA analysis has confirmed the specific status of
<taxonomicName id="94C24D25FFF9FFE7E50B01A5FB3FFDDA" authorityName="Fruhstorfer" authorityYear="1908" box="[1073,1180,511,537]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="telona">
<emphasis id="61B6EAB4FFF9FFE7E50B01A5FB3FFDDA" box="[1073,1180,511,537]" italics="true" pageId="1" pageNumber="26">M. telona</emphasis>
</taxonomicName>
and is widely regarded as a tool for the reliable separation of closely related species. However, it is not definitive, as exemplified by the similarity of the results of the analysis of the
<emphasis id="61B6EAB4FFF9FFE7E3B4021DFD1DFDA2" box="[654,702,583,609]" italics="true" pageId="1" pageNumber="26">COI</emphasis>
gene in
<taxonomicName id="94C24D25FFF9FFE7E226021DFC2FFDA3" authorityName="Christoph" authorityYear="1893" box="[796,908,583,608]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="61B6EAB4FFF9FFE7E226021DFC2FFDA3" box="[796,908,583,608]" italics="true" pageId="1" pageNumber="26">M. ornata</emphasis>
</taxonomicName>
and
<taxonomicName id="94C24D25FFF9FFE7E2F8021DFB94FDA2" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[962,1079,583,609]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFF9FFE7E2F8021DFB94FDA2" box="[962,1079,583,609]" italics="true" pageId="1" pageNumber="26">M. phoebe</emphasis>
</taxonomicName>
from western populations. Also DNA analysis has so far been unable to separate
<taxonomicName id="94C24D25FFF9FFE7E3940231FC80FD46" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[686,803,619,645]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFF9FFE7E3940231FC80FD46" box="[686,803,619,645]" italics="true" pageId="1" pageNumber="26">M. phoebe</emphasis>
</taxonomicName>
from
<taxonomicName id="94C24D25FFF9FFE7E25F0231FB28FD46" authority="Alpheraky, 1881" authorityName="Alpheraky" authorityYear="1881" box="[869,1163,619,645]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="sibina">
<emphasis id="61B6EAB4FFF9FFE7E25F0231FC6EFD46" box="[869,973,619,645]" italics="true" pageId="1" pageNumber="26">M. sibina</emphasis>
Alphéraky, 1881
</taxonomicName>
(
<bibRefCitation id="37534B57FFF9FFE7E5A20231FA34FD46" author="Lukhtanov, V. A. &amp; Sourakov, A. &amp; Zakharov, E. V. &amp; Hebert, P. D. N." box="[1176,1431,619,645]" pageId="1" pageNumber="26" pagination="1302 - 1310" refId="ref10220" refString="Lukhtanov, V. A., Sourakov, A., Zakharov, E. V. &amp; Hebert, P. D. N. (2009) DNA barcoding Central Asian butterflies: increasing geographical dimension does not significantly reduce the success of species identification. Molecular Ecology Resources, 9, 1302 - 1310. https: // doi. org / 10.1111 / j. 1755 - 0998.2009.02577. x" type="journal article" year="2009">
Lukhtanov
<emphasis id="61B6EAB4FFF9FFE7E4230231FAEAFD46" box="[1305,1353,619,645]" italics="true" pageId="1" pageNumber="26">et al</emphasis>
., 2009
</bibRefCitation>
;
<bibRefCitation id="37534B57FFF9FFE7E1AD02D5FE23FD6A" author="Leneveu J. &amp; Chichvarkhin, A. &amp; Wahlberg, N." box="[151,384,655,681]" pageId="1" pageNumber="26" pagination="346 - 361" refId="ref10155" refString="Leneveu J., Chichvarkhin, A. &amp; Wahlberg, N. (2009) Varying rates of diversification in the genus Melitaea (Lepidoptera: Nymphalidae) during the past 20 million years. Biological Journal of the Linnean Society, 97, 346 - 361. https: // doi. org / 10.1111 / j. 1095 - 8312.2009.01208. x" type="journal article" year="2009">
Leneveu
<emphasis id="61B6EAB4FFF9FFE7E1C502D5FE94FD6A" box="[255,311,655,681]" italics="true" pageId="1" pageNumber="26">et al.</emphasis>
, 2009
</bibRefCitation>
;
<bibRefCitation id="37534B57FFF9FFE7E0B102D5FDE1FD6A" author="Toth, J. P. &amp; Bereczki, J. &amp; Varga, Z. &amp; Rota, J. &amp; Sramko, G. &amp; Wahlberg, N." box="[395,578,655,681]" pageId="1" pageNumber="26" pagination="749 - 757" refId="ref11329" refString="Toth, J. P., Bereczki, J., Varga, Z., Rota, J., Sramko, G. &amp; Wahlberg, N. (2014) Relationships within the Melitaea phoebe species group (Lepidoptera: Nymphalidae): new insights from molecular and morphometric information. Systematic Entomology, 39, 749 - 757. https: // doi. org / 10.1111 / syen. 12083" type="journal article" year="2014">
Tóth
<emphasis id="61B6EAB4FFF9FFE7E0FD02D5FE5CFD6A" box="[455,511,655,681]" italics="true" pageId="1" pageNumber="26">et al.</emphasis>
2014
</bibRefCitation>
), whilst differences in their wing morphology are distinctive.
</paragraph>
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90
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<emphasis id="61B6EAB4FFF9FFE7E1AD033DFEB3FC42" bold="true" box="[151,272,870,897]" italics="true" pageId="1" pageNumber="26">M. phoebe</emphasis>
var.
<emphasis id="61B6EAB4FFF9FFE7E074033CFE11FC42" bold="true" box="[334,434,870,897]" italics="true" pageId="1" pageNumber="26">aetherea</emphasis>
</emphasis>
Eversmann, 1851
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="1BD8652DFFF9FFE7E3BE033DFADAFC2E" pageId="1" pageNumber="26" type="discussion">
<paragraph id="537D36A6FFF9FFE7E3BE033DFADAFC2E" blockId="1.[151,1437,870,2013]" pageId="1" pageNumber="26">
[TL: Southern
<collectingCountry id="2BD57636FFF9FFE7E210033DFBA6FC42" box="[810,1029,870,897]" name="Russia" pageId="1" pageNumber="26">Russian Federation</collectingCountry>
] is considered by the authors to be a lightly marked but brightly coloured variety
<taxonomicName id="94C24D25FFF9FFE7E3A803D1FCAAFC66" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[658,777,907,933]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFF9FFE7E3A803D1FCAAFC66" box="[658,777,907,933]" italics="true" pageId="1" pageNumber="26">M. phoebe</emphasis>
</taxonomicName>
with reduced black markings on both wing surfaces. This nomenclatural association agrees with those of
<bibRefCitation id="37534B57FFF9FFE7E39103F5FC24FC0A" author="Higgins, L. G." box="[683,903,943,969]" pageId="1" pageNumber="26" pagination="175 - 365" refId="ref9372" refString="Higgins, L. G. (1941) An illustrated catalogue of the Palearctic Melitaea (Lep. Rhopalocera). Transactions of the Royal Entomological Society of London, 91, 175 - 365. https: // doi. org / 10.1111 / j. 1365 - 2311.1941. tb 01045. x" type="journal article" year="1941">Higgins (1941: 331</bibRefCitation>
, plate 14, fig. 9),
<bibRefCitation id="37534B57FFF9FFE7E57703F5FACBFC0A" author="Tshikolovets, V. V." box="[1101,1384,943,969]" pageId="1" pageNumber="26" refId="ref11624" refString="Tshikolovets, V. V. (2011) Butterflies of Europe &amp; the Mediterranean area. Tshikolovets Publications, Pardubice, Czech Republic, 544 pp." type="book" year="2011">Tshikolovets (2011: 497)</bibRefCitation>
,
<bibRefCitation id="37534B57FFF9FFE7E44E03F5FE51FC2D" author="Van Oorschot, H. &amp; Coutsis, J. G." pageId="1" pageNumber="26" refId="ref11693" refString="Van Oorschot, H. &amp; Coutsis, J. G. (2014) The genus Melitaea Fabricius, 1807 (Lepidoptera, Nymphalidae, Nymphalinae) Taxonomy and systematics with special reference to the male genitalia. Tshikolovets publications, Pardubice, Czech Republic, 360 pp." type="book" year="2014">van Oorschot &amp; Coutsis (2014: 60)</bibRefCitation>
and
<bibRefCitation id="37534B57FFF9FFE7E3130389FCCEFC2E" author="Russell, PJ &amp; Tennent, W. J." box="[553,877,979,1005]" pageId="1" pageNumber="26" pagination="27 - 56" refId="ref10922" refString="Russell, PJ. C. &amp; Tennent, W. J. (2016) A synonymic list of names associated with western Palaearctic Melitaea phoebe (Denis &amp; Schiffermuller, 1775) species group taxa M. phoebe; M. punica Oberthur, 1876; M. ornata Christoph, 1893) (Lepidoptera, Nymphalidae). Nota Lepidopterologica, 39 (1), 27 - 56. https: // doi. org / 10.3897 / nl. 39.5929" type="journal article" year="2016">Russell &amp; Tennent (2016: 43</bibRefCitation>
, note 9), with which the present authors agree.
</paragraph>
</subSubSection>
</treatment>
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<subSubSection id="1BD8652DFFF9FFE7E1FD03ADFD3FFBD2" box="[199,668,1014,1041]" pageId="1" pageNumber="26" type="nomenclature">
<paragraph id="537D36A6FFF9FFE7E1FD03ADFD3FFBD2" blockId="1.[151,1437,870,2013]" box="[199,668,1014,1041]" pageId="1" pageNumber="26">
<taxonomicName id="94C24D25FFF9FFE7E1FD03ADFD3FFBD2" ID-CoL="3ZK6S" authority="Staudinger, 1870" authorityName="Staudinger" authorityYear="1870" box="[199,668,1014,1041]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="caucasica">
<emphasis id="61B6EAB4FFF9FFE7E1FD03ADFE6BFBD2" bold="true" box="[199,456,1014,1041]" italics="true" pageId="1" pageNumber="26">M. phoebe caucasica</emphasis>
<bibRefCitation id="37534B57FFF9FFE7E0EC03ADFD3FFBD2" author="Staudinger, O." box="[470,668,1015,1041]" pageId="1" pageNumber="26" pagination="3" refId="ref11169" refString="Staudinger, O. (1870) Beitrag zur Lepidopterenfauna Griechenlands. Horae Societatis Entomologicae Rossicae, 7, [3] - 304." type="journal article" year="1870">Staudinger, 1870</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="1BD8652DFFF9FFE7E39003ADFB6BF9B6" pageId="1" pageNumber="26" type="discussion">
<paragraph id="537D36A6FFF9FFE7E39003ADFB6BF9B6" blockId="1.[151,1437,870,2013]" pageId="1" pageNumber="26">
[TL: “Kindermann ganz ähnliche Stücke im Caucasus fing (?- Helenendorf; Kindermann leg.)”]. The name
<emphasis id="61B6EAB4FFF9FFE7E3B50441FD5CFBF7" box="[655,767,1051,1076]" italics="true" pageId="1" pageNumber="26">caucasica</emphasis>
was preoccupied by
<taxonomicName id="94C24D25FFF9FFE7E2D00441FA3FFBF6" authority="Staudinger, 1861" authorityName="Staudinger" authorityYear="1861" box="[1002,1436,1051,1077]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="subSpecies" species="didyma" subSpecies="caucasica">
<emphasis id="61B6EAB4FFF9FFE7E2D00441FB7BFBF7" box="[1002,1240,1051,1077]" italics="true" pageId="1" pageNumber="26">M. didyma caucasica</emphasis>
<bibRefCitation id="37534B57FFF9FFE7E5E40441FA3FFBF6" author="Staudinger, O." box="[1246,1436,1051,1077]" pageId="1" pageNumber="26" refId="ref11128" refString="Staudinger, O. (1861) In Staudinger, O. &amp; Wocke, M. F. Catalog der Lepidopteren Europa's und der angrenzenden Lander, Staudinger &amp; Burdach, Dresden, Germany, XVI + 192 pp." type="book" year="1861">Staudinger, 1861</bibRefCitation>
</taxonomicName>
and the name was replaced first by
<taxonomicName id="94C24D25FFF9FFE7E30F0465FC6CFB9A" authority="Fruhstorfer 1917" authorityName="Fruhstorfer" authorityYear="1917" box="[565,975,1087,1113]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="ottonis">
<emphasis id="61B6EAB4FFF9FFE7E30F0465FCA0FB9B" box="[565,771,1087,1113]" italics="true" pageId="1" pageNumber="26">M. phoebe ottonis</emphasis>
<bibRefCitation id="37534B57FFF9FFE7E2370465FC6CFB9A" author="Fruhstorfer, H." box="[781,975,1087,1113]" pageId="1" pageNumber="26" pagination="19" refId="ref9078" refString="Fruhstorfer, H. (1917) Neue Rhopaloceren aus der Sammlung Leonhard. Societas Entomologica, 32 (5), 19." type="journal article" year="1917">Fruhstorfer 1917</bibRefCitation>
</taxonomicName>
. A
<typeStatus id="8C798804FFF9FFE7E2C20465FBFCFB9A" box="[1016,1119,1087,1113]" pageId="1" pageNumber="26" type="lectotype">lectotype</typeStatus>
female and a
<typeStatus id="8C798804FFF9FFE7E43F0465FA3FFB9A" box="[1285,1436,1087,1113]" pageId="1" pageNumber="26" type="paralectotype">paralectotype</typeStatus>
male were designated by Nekrutenko (
<bibRefCitation id="37534B57FFF9FFE7E36F0439FC27FBBE" author="Hesselbarth, G. &amp; van Oorschot, H. &amp; Wagener, S." box="[597,900,1122,1149]" pageId="1" pageNumber="26" pagination="1 - 847" refId="ref9287" refString="Hesselbarth, G., van Oorschot, H. &amp; Wagener, S. (1995) Die Tagfalter der Turkei unter Berucksichtigung der angrenzenden Lander, Sigbert Wagener, Bocholt, Germany, 1, 1 - 757, 2, 758 - 1354, 3, 1 - 847." type="journal article" year="1995">
Hesselbarth
<emphasis id="61B6EAB4FFF9FFE7E3DF0439FCBAFBBE" box="[741,793,1123,1149]" italics="true" pageId="1" pageNumber="26">et al</emphasis>
. 1995: 2
</bibRefCitation>
: 1028) from the Staudinger collection, housed at Zoologisches Museum der Humboldt Universität,
<collectingRegion id="9106F844FFF9FFE7E23904DDFCEAFB62" box="[771,841,1159,1185]" country="Germany" name="Berlin" pageId="1" pageNumber="26">Berlin</collectingRegion>
(figs 5A, B, C &amp; 6A, B, C). Verity subsequently also proposed a replacement name,
<emphasis id="61B6EAB4FFF9FFE7E30F04F1FD18FB06" box="[565,699,1195,1221]" italics="true" pageId="1" pageNumber="26">caucasicola</emphasis>
<bibRefCitation id="37534B57FFF9FFE7E3FE04F1FCF1FB06" author="Verity, R." box="[708,850,1195,1221]" pageId="1" pageNumber="26" pagination="26 - 31" refId="ref11898" refString="Verity, R. (1919) Seasonal Polymorphism and Races of some European Grypocera and Rhopalocera. Entomologist's Record and Journal of Variation 31 (2), 26 - 31, (3), 43 - 48, (5), 87 - 89, (7), 121 - 129, (10), 178 - 184, (11), 193 - 201." type="journal article" year="1919">Verity, 1919</bibRefCitation>
, this being a synonym of
<emphasis id="61B6EAB4FFF9FFE7E5B804F1FB6DFB07" box="[1154,1230,1195,1220]" italics="true" pageId="1" pageNumber="26">ottonis</emphasis>
.
<bibRefCitation id="37534B57FFF9FFE7E5E504F1FEAFFB2A" author="Kemal, M. &amp; Kocak, A. O." pageId="1" pageNumber="26" refId="ref9502" refString="Kemal, M. &amp; Kocak, A. O. (2011) A synonymical and distributional checklist of the Papilionoidea and Hesperioidea of East Mediterranean countries, including Turkey (Lepidoptera). Priamus, 25 (Supplement), 1 - 162, Plates 1 - 42." type="book" year="2011">Kemal &amp; Koçak (2011: 44)</bibRefCitation>
used the name ‘
<taxonomicName id="94C24D25FFF9FFE7E0FF0495FD09FB2A" box="[453,682,1231,1257]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="subGenus" subGenus="Cinclidia">
<emphasis id="61B6EAB4FFF9FFE7E0FF0495FD8AFB2A" box="[453,553,1231,1257]" italics="true" pageId="1" pageNumber="26">Melitaea</emphasis>
(
<emphasis id="61B6EAB4FFF9FFE7E3030495FD02FB2A" box="[569,673,1231,1257]" italics="true" pageId="1" pageNumber="26">Cinclidia</emphasis>
)
</taxonomicName>
(
<taxonomicName id="94C24D25FFF9FFE7E3860495FCAEFB2A" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[700,781,1231,1257]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFF9FFE7E3860495FCAEFB2A" box="[700,781,1231,1257]" italics="true" pageId="1" pageNumber="26">phoebe</emphasis>
</taxonomicName>
)
<emphasis id="61B6EAB4FFF9FFE7E2270495FCC8FB2A" box="[797,875,1231,1257]" italics="true" pageId="1" pageNumber="26">sextilis</emphasis>
<bibRefCitation id="37534B57FFF9FFE7E2480495FB88FB2A" author="Jachontov, A. A." box="[882,1067,1231,1257]" pageId="1" pageNumber="26" pagination="282 - 292" refId="ref9463" refString="Jachontov, A. A. (1909) Notice sur les Lepidopteres Rhopaloceres du Caucase. Russkoe entomologicheskoe obozreniie (Revue Russe d'Entomologie), 8 (1908), 282 - 292. [In Russian]" type="journal article" year="1909">Jachontov, 1909</bibRefCitation>
’ as a replacement name giving it subspecific(?) status; however,
<bibRefCitation id="37534B57FFF9FFE7E0CD04A9FD50FACE" author="Jachontov, A. A." box="[503,755,1266,1293]" pageId="1" pageNumber="26" pagination="282 - 292" refId="ref9463" refString="Jachontov, A. A. (1909) Notice sur les Lepidopteres Rhopaloceres du Caucase. Russkoe entomologicheskoe obozreniie (Revue Russe d'Entomologie), 8 (1908), 282 - 292. [In Russian]" type="journal article" year="1909">Jachontov (1909: 285)</bibRefCitation>
used this name for a variety of second generation
<taxonomicName id="94C24D25FFF9FFE7E41C04A9FA3FFACE" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[1318,1436,1267,1293]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFF9FFE7E41C04A9FA3FFACE" box="[1318,1436,1267,1293]" italics="true" pageId="1" pageNumber="26">M. phoebe</emphasis>
</taxonomicName>
and, so far as the authors are aware, no author since has used the name
<emphasis id="61B6EAB4FFF9FFE7E2E3054DFB84FAF2" box="[985,1063,1303,1329]" italics="true" pageId="1" pageNumber="26">sextilis</emphasis>
in favour of
<emphasis id="61B6EAB4FFF9FFE7E5F9054DFAACFAF3" box="[1219,1295,1303,1328]" italics="true" pageId="1" pageNumber="26">ottonis</emphasis>
<bibRefCitation id="37534B57FFF9FFE7E422054DFF71FA96" author="Fruhstorfer, H." pageId="1" pageNumber="26" pagination="19" refId="ref9078" refString="Fruhstorfer, H. (1917) Neue Rhopaloceren aus der Sammlung Leonhard. Societas Entomologica, 32 (5), 19." type="journal article" year="1917">Fruhstorfer, 1917</bibRefCitation>
. In fact the
<taxonomicName id="94C24D25FFF9FFE7E0580561FE7FFA96" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[354,476,1339,1365]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFF9FFE7E0580561FE7FFA96" box="[354,476,1339,1365]" italics="true" pageId="1" pageNumber="26">M. phoebe</emphasis>
</taxonomicName>
species group portrayed by
<bibRefCitation id="37534B57FFF9FFE7E2120561FBC9FA96" author="Kemal, M. &amp; Kocak, A. O." box="[808,1130,1339,1365]" pageId="1" pageNumber="26" refId="ref9502" refString="Kemal, M. &amp; Kocak, A. O. (2011) A synonymical and distributional checklist of the Papilionoidea and Hesperioidea of East Mediterranean countries, including Turkey (Lepidoptera). Priamus, 25 (Supplement), 1 - 162, Plates 1 - 42." type="book" year="2011">Kemal &amp; Koçak (2011: 44)</bibRefCitation>
, in their article on eastern Mediterranean butterflies, included
<taxonomicName id="94C24D25FFF9FFE7E3170505FD03FABB" authorityName="Oberthur" authorityYear="1876" box="[557,672,1375,1400]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="punica">
<emphasis id="61B6EAB4FFF9FFE7E3170505FD03FABB" box="[557,672,1375,1400]" italics="true" pageId="1" pageNumber="26">M. punica</emphasis>
</taxonomicName>
, a species absent from the eastern Mediterranean. This perpetuates confusion, which the first author with others has been trying to resolve.
<bibRefCitation id="37534B57FFF9FFE7E28905D9FB7AFA5E" author="Hesselbarth, G. &amp; van Oorschot, H. &amp; Wagener, S." box="[947,1241,1410,1437]" pageId="1" pageNumber="26" pagination="1 - 847" refId="ref9287" refString="Hesselbarth, G., van Oorschot, H. &amp; Wagener, S. (1995) Die Tagfalter der Turkei unter Berucksichtigung der angrenzenden Lander, Sigbert Wagener, Bocholt, Germany, 1, 1 - 757, 2, 758 - 1354, 3, 1 - 847." type="journal article" year="1995">
Hesselbarth
<emphasis id="61B6EAB4FFF9FFE7E50405D9FBCDFA5E" box="[1086,1134,1411,1437]" italics="true" pageId="1" pageNumber="26">et al</emphasis>
. (1995: 3
</bibRefCitation>
, Tafel 80/81: figs 30–
<specimenCount id="45C4FD2FFFF9FFE7E1F805FDFF5EFA02" box="[194,253,1447,1473]" pageId="1" pageNumber="26" type="male">33 ♂</specimenCount>
; Tafel 82/83: figs 1–
<specimenCount id="45C4FD2FFFF9FFE7E0DF05FDFDACFA02" box="[485,527,1447,1473]" pageId="1" pageNumber="26" type="female">4 ♀</specimenCount>
) placed
<emphasis id="61B6EAB4FFF9FFE7E34905FDFD1CFA03" box="[627,703,1447,1472]" italics="true" pageId="1" pageNumber="26">ottonis</emphasis>
as a synonym of
<taxonomicName id="94C24D25FFF9FFE7E2BD05FDFC5DFA02" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[903,1022,1447,1473]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFF9FFE7E2BD05FDFC5DFA02" box="[903,1022,1447,1473]" italics="true" pageId="1" pageNumber="26">M. phoebe</emphasis>
</taxonomicName>
. Although the
<typeStatus id="8C798804FFF9FFE7E59D05FDFAADFA02" box="[1191,1294,1447,1473]" pageId="1" pageNumber="26" type="lectotype">lectotype</typeStatus>
female does not show all the characters typical of
<taxonomicName id="94C24D25FFF9FFE7E3070591FD17FA26" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[573,692,1483,1509]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFF9FFE7E3070591FD17FA26" box="[573,692,1483,1509]" italics="true" pageId="1" pageNumber="26">M. phoebe</emphasis>
</taxonomicName>
, for instance the underside submarginal black arches do not touch the intervening veins (see
<figureCitation id="CBF92A23FFF9FFE7E08405B5FDB6F9CA" box="[446,533,1518,1545]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1717,1741]" captionTargetBox="[153,1436,200,1692]" captionTargetId="figure-26@2.[151,1436,181,1693]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, Melitaea ornata male underside, Russian Federation, southern Urals, Orenburg Province, Guberli, ca. 190 m, 14 May 2014. 2, Melitaea phoebe male underside, Russian Federation, southern Urals, Orenburg Province, Krasnoznamenka, ca. 210 m, 16 May 2014. 3, Melitaea ornata female underside, Slovenia, Koper, ca. 2km NW of Rakitovec ca. 525 m, 17 May 2011. 4, Melitaea phoebe female underside, Slovenia, Koper, ca. 2km S. of Rakitovec ca. 525 m, 17 May 2011. 5, Lectotype female of Melitaea phoebe caucasica Staudinger, 1870. 6, Paralectotype male of Melitaea phoebe caucasica Staudinger, 1870. Figs 1–4 © P. Russell. Figs 5–6 © T. Léger. A = upperside, B = underside, C = labels on specimen pin." figureDoi="http://doi.org/10.5281/zenodo.6577631" httpUri="https://zenodo.org/record/6577631/files/figure.png" pageId="1" pageNumber="26">Fig. 5B</figureCitation>
), the
<typeStatus id="8C798804FFF9FFE7E36E05B5FD48F9CA" box="[596,747,1519,1545]" pageId="1" pageNumber="26" type="paralectotype">paralectotype</typeStatus>
underside (
<figureCitation id="CBF92A23FFF9FFE7E25405B5FC67F9C9" box="[878,964,1519,1546]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1717,1741]" captionTargetBox="[153,1436,200,1692]" captionTargetId="figure-26@2.[151,1436,181,1693]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, Melitaea ornata male underside, Russian Federation, southern Urals, Orenburg Province, Guberli, ca. 190 m, 14 May 2014. 2, Melitaea phoebe male underside, Russian Federation, southern Urals, Orenburg Province, Krasnoznamenka, ca. 210 m, 16 May 2014. 3, Melitaea ornata female underside, Slovenia, Koper, ca. 2km NW of Rakitovec ca. 525 m, 17 May 2011. 4, Melitaea phoebe female underside, Slovenia, Koper, ca. 2km S. of Rakitovec ca. 525 m, 17 May 2011. 5, Lectotype female of Melitaea phoebe caucasica Staudinger, 1870. 6, Paralectotype male of Melitaea phoebe caucasica Staudinger, 1870. Figs 1–4 © P. Russell. Figs 5–6 © T. Léger. A = upperside, B = underside, C = labels on specimen pin." figureDoi="http://doi.org/10.5281/zenodo.6577631" httpUri="https://zenodo.org/record/6577631/files/figure.png" pageId="1" pageNumber="26">Fig. 6B</figureCitation>
) certainly shows all the characters typical of
<taxonomicName id="94C24D25FFF9FFE7E18C0649FE8EF9EE" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[182,301,1555,1581]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFF9FFE7E18C0649FE8EF9EE" box="[182,301,1555,1581]" italics="true" pageId="1" pageNumber="26">M. phoebe</emphasis>
</taxonomicName>
. Recent authors, such
<bibRefCitation id="37534B57FFF9FFE7E3160649FC9CF9EE" author="Tshikolovets, V. V." box="[556,831,1555,1581]" pageId="1" pageNumber="26" refId="ref11624" refString="Tshikolovets, V. V. (2011) Butterflies of Europe &amp; the Mediterranean area. Tshikolovets Publications, Pardubice, Czech Republic, 544 pp." type="book" year="2011">Tshikolovets (2011: 497</bibRefCitation>
; 2003: plate 24: figs
<specimenCount id="45C4FD2FFFF9FFE7E5140649FBCCF9EE" box="[1070,1135,1555,1581]" pageId="1" pageNumber="26" type="generic">
<quantity id="943A9B43FFF9FFE7E5140649FBC8F9EE" box="[1070,1131,1555,1581]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="26" unit="m" value="16.0">16 m</quantity>
.
</specimenCount>
and
<specimenCount id="45C4FD2FFFF9FFE7E59C0649FB7FF9ED" box="[1190,1244,1555,1582]" pageId="1" pageNumber="26" type="generic">17 f.</specimenCount>
),
<bibRefCitation id="37534B57FFF9FFE7E5C80649FEDAF992" author="Tshikolovets, V. V. &amp; Naderi A. &amp; Eckweiler, W." pageId="1" pageNumber="26" refId="ref11652" refString="Tshikolovets, V. V., Naderi A. &amp; Eckweiler, W. (2014) The butterflies of Iran and Iraq. Tshikolovets Publications, Pardubice, Czech Republic, 366 pp. + 65 colour plates." type="book" year="2014">
Tshikolovets
<emphasis id="61B6EAB4FFF9FFE7E4B20649FF17F992" italics="true" pageId="1" pageNumber="26">et al.</emphasis>
(2014: 318–319)
</bibRefCitation>
,
<bibRefCitation id="37534B57FFF9FFE7E0B9066DFCAFF991" author="Van Oorschot, H. &amp; Coutsis, J. G." box="[387,780,1591,1618]" pageId="1" pageNumber="26" refId="ref11693" refString="Van Oorschot, H. &amp; Coutsis, J. G. (2014) The genus Melitaea Fabricius, 1807 (Lepidoptera, Nymphalidae, Nymphalinae) Taxonomy and systematics with special reference to the male genitalia. Tshikolovets publications, Pardubice, Czech Republic, 360 pp." type="book" year="2014">van Oorschot &amp; Coutsis (2014: 60)</bibRefCitation>
and
<bibRefCitation id="37534B57FFF9FFE7E278066DFB26F992" author="Russell, PJ &amp; Tennent, W. J." box="[834,1157,1590,1617]" pageId="1" pageNumber="26" pagination="27 - 56" refId="ref10922" refString="Russell, PJ. C. &amp; Tennent, W. J. (2016) A synonymic list of names associated with western Palaearctic Melitaea phoebe (Denis &amp; Schiffermuller, 1775) species group taxa M. phoebe; M. punica Oberthur, 1876; M. ornata Christoph, 1893) (Lepidoptera, Nymphalidae). Nota Lepidopterologica, 39 (1), 27 - 56. https: // doi. org / 10.3897 / nl. 39.5929" type="journal article" year="2016">Russell &amp; Tennent (2016: 45</bibRefCitation>
, note 22) have all agreed that this is a subspecies of
<taxonomicName id="94C24D25FFF9FFE7E0F80601FD9BF9B6" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[450,568,1627,1653]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFF9FFE7E0F80601FD9BF9B6" box="[450,568,1627,1653]" italics="true" pageId="1" pageNumber="26">M. phoebe</emphasis>
</taxonomicName>
and not
<taxonomicName id="94C24D25FFF9FFE7E3A30601FCA9F9B7" authorityName="Christoph" authorityYear="1893" box="[665,778,1627,1652]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="61B6EAB4FFF9FFE7E3A30601FCA9F9B7" box="[665,778,1627,1652]" italics="true" pageId="1" pageNumber="26">M. ornata</emphasis>
</taxonomicName>
, with which the present authors concur.
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="537D36A6FFFBFFE5E1FD01EDFD2BFE12" blockId="3.[151,1437,151,2013]" box="[199,648,438,465]" pageId="3" pageNumber="28">
<emphasis id="61B6EAB4FFFBFFE5E1FD01EDFE10FE12" bold="true" box="[199,435,438,465]" pageId="3" pageNumber="28">
<taxonomicName id="94C24D25FFFBFFE5E1FD01EDFE9CFE12" ID-CoL="6QY6N" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[199,319,438,465]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFBFFE5E1FD01EDFE9CFE12" bold="true" box="[199,319,438,465]" italics="true" pageId="3" pageNumber="28">M. phoebe</emphasis>
</taxonomicName>
ab.
<emphasis id="61B6EAB4FFFBFFE5E05501EDFE10FE12" bold="true" box="[367,435,439,465]" italics="true" pageId="3" pageNumber="28">tatara</emphasis>
</emphasis>
<bibRefCitation id="37534B57FFFBFFE5E08301EDFD2BFE12" author="Krulikovsky, A." box="[441,648,439,465]" pageId="3" pageNumber="28" pagination="200 - 251" refId="ref9900" refString="Krulikovsky, A. (1891) Experimental catalogue of Lepidoptera of Kazakh Province. Bullitin de la Societe Imperiale des Naturalistes de Moscou (New Series) 4, 200 - 251. [In Russian, colour pls. I - VIII]" type="journal article" year="1891">Krulikovsky, 1891</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="1BD8652DFFFBFFE5E3B401EDFB99FCFA" pageId="3" pageNumber="28" type="discussion">
<paragraph id="537D36A6FFFBFFE5E3B401EDFB99FCFA" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="28">
[TL:
<emphasis id="61B6EAB4FFFBFFE5E3FD01EDFCA3FE13" box="[711,768,439,464]" italics="true" pageId="3" pageNumber="28">circa</emphasis>
Kazan,
<collectingCountry id="2BD57636FFFBFFE5E26301EDFB90FE12" box="[857,1075,438,465]" name="Russia" pageId="3" pageNumber="28">Russian Federation</collectingCountry>
] was considered by Krulikovsky to be a hybrid. In 1897 Krulikovsky said: ‛Earlier I mentioned that I caught
<taxonomicName id="94C24D25FFFBFFE5E53A0181FBDBFE36" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[1024,1144,475,501]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFBFFE5E53A0181FBDBFE36" box="[1024,1144,475,501]" italics="true" pageId="3" pageNumber="28">M. phoebe</emphasis>
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(male) in copula with
<taxonomicName id="94C24D25FFFBFFE5E4440181FF5EFDDA" baseAuthorityName="Esper" baseAuthorityYear="1783" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="arduinna">
<emphasis id="61B6EAB4FFFBFFE5E4440181FF5EFDDA" italics="true" pageId="3" pageNumber="28">M. arduinna</emphasis>
</taxonomicName>
(female). I suspect that ab.
<emphasis id="61B6EAB4FFFBFFE5E31401A5FDD1FDDB" box="[558,626,511,536]" italics="true" pageId="3" pageNumber="28">tatara</emphasis>
, which was shown by me in Bull. de
<collectingRegion id="9106F844FFFBFFE5E53501A5FBCFFDDA" box="[1039,1132,511,537]" country="Russia" name="Moscow" pageId="3" pageNumber="28">Moscou</collectingRegion>
, 1890, pl VIII, f. g, perhaps is a hybrid between
<taxonomicName id="94C24D25FFFBFFE5E0410279FE51FDFE" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[379,498,547,573]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFBFFE5E0410279FE51FDFE" box="[379,498,547,573]" italics="true" pageId="3" pageNumber="28">M. phoebe</emphasis>
</taxonomicName>
and
<taxonomicName id="94C24D25FFFBFFE5E3100279FD44FDFE" authority=", Rott." authorityName="Rott." box="[554,743,547,573]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="athalia">
<emphasis id="61B6EAB4FFFBFFE5E3100279FD3CFDFE" box="[554,671,547,573]" italics="true" pageId="3" pageNumber="28">M. athalia</emphasis>
, Rott.
</taxonomicName>
’.[Translation by G. Kuznetsov] [The species mentioned refer to
<taxonomicName id="94C24D25FFFBFFE5E189021DFE7AFDA2" authority="(Esper, 1783)" baseAuthorityName="Esper" baseAuthorityYear="1783" box="[179,473,583,609]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="arduinna">
<emphasis id="61B6EAB4FFFBFFE5E189021DFE9EFDA2" box="[179,317,583,609]" italics="true" pageId="3" pageNumber="28">M. arduinna</emphasis>
(
<bibRefCitation id="37534B57FFFBFFE5E070021DFE73FDA2" author="Esper, E. J. C." box="[330,464,583,609]" pageId="3" pageNumber="28" pagination="157 - 172" refId="ref8959" refString="Esper, E. J. C. (1783) Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I: Die Tagschmetterlinge. Fortsetzung. Band 2 (7), 141 - 156, plates 81, 82, (8), 157 - 172, plates 83 - 88." type="journal article" year="1783">Esper, 1783</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="94C24D25FFFBFFE5E337021DFCCAFDA2" authority="(Rottemburg, 1775)" baseAuthorityName="Rottemburg" baseAuthorityYear="1775" box="[525,873,582,609]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="athalia">
<emphasis id="61B6EAB4FFFBFFE5E337021DFD23FDA2" box="[525,640,583,609]" italics="true" pageId="3" pageNumber="28">M. athalia</emphasis>
(
<bibRefCitation id="37534B57FFFBFFE5E3B4021DFCC3FDA2" author="Rottemburg, S. A." box="[654,864,582,609]" pageId="3" pageNumber="28" pagination="1 - 34" refId="ref10490" refString="Rottemburg, S. A. (1775) Unmertungen zu den Hufnagelifchen Tabellen der Schmitterlinge Der Naturforscher, 6, 1 - 34." type="journal article" year="1775">Rottemburg, 1775</bibRefCitation>
)
</taxonomicName>
.] Unfortunately the search by
<bibRefCitation id="37534B57FFFBFFE5E58E021DFEF0FD46" author="Kostjuk I. Y. &amp; Zolotuhin V. V." pageId="3" pageNumber="28" pagination="586 - 593" refId="ref9806" refString="Kostjuk I. Y. &amp; Zolotuhin V. V. (2017) The types of moths and butterflies described from the Volga-Ural Region by Leonid Krulikowsky (Krulikovsky). Part 2. Macrolepidoptera. In: Anikin V. V., Sachkov S. A., Zolotuhin, V. V. (Eds.), Fauna lepidopterologica Volgo-Uralensis: from P. Pallas to present days. Munich-Vilnius. Proceedings of the Museum Witt. Munich, 7, 443 - 450, 586 - 593." type="journal article" year="2017">Kostjuk &amp; Zolotuhin (2017: 445–446)</bibRefCitation>
in the collections of the Zoologisches Museum of Shevchenko University, Kyiv failed to find the
<typeStatus id="8C798804FFFBFFE5E1AD02D5FF5EFD6A" box="[151,253,655,681]" pageId="3" pageNumber="28" type="holotype">holotype</typeStatus>
. The habitat was ‘swampy meadows’ unsuitable for
<taxonomicName id="94C24D25FFFBFFE5E26B02D5FC60FD6B" authorityName="Christoph" authorityYear="1893" box="[849,963,655,680]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="61B6EAB4FFFBFFE5E26B02D5FC60FD6B" box="[849,963,655,680]" italics="true" pageId="3" pageNumber="28">M. ornata</emphasis>
</taxonomicName>
but acceptable to both
<taxonomicName id="94C24D25FFFBFFE5E5F702D5FAE6FD6A" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[1229,1349,655,681]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFBFFE5E5F702D5FAE6FD6A" box="[1229,1349,655,681]" italics="true" pageId="3" pageNumber="28">M. phoebe</emphasis>
</taxonomicName>
and
<taxonomicName id="94C24D25FFFBFFE5E44402D5FF45FD0E" baseAuthorityName="Rottemburg" baseAuthorityYear="1775" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="athalia">
<emphasis id="61B6EAB4FFFBFFE5E44402D5FF45FD0E" italics="true" pageId="3" pageNumber="28">M. athalia</emphasis>
</taxonomicName>
; also Kazan is outside the known distribution of
<taxonomicName id="94C24D25FFFBFFE5E22202E9FC29FD0F" authorityName="Christoph" authorityYear="1893" box="[792,906,691,716]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="61B6EAB4FFFBFFE5E22202E9FC29FD0F" box="[792,906,691,716]" italics="true" pageId="3" pageNumber="28">M. ornata</emphasis>
</taxonomicName>
. Hybrids between
<taxonomicName id="94C24D25FFFBFFE5E55B02E9FABEFD0E" box="[1121,1309,691,717]" pageId="3" pageNumber="28">
<emphasis id="61B6EAB4FFFBFFE5E55B02E9FB66FD0E" box="[1121,1221,691,717]" italics="true" pageId="3" pageNumber="28">Melitaea</emphasis>
species
</taxonomicName>
are known to occur (
<bibRefCitation id="37534B57FFFBFFE5E038028DFE78FD32" author="Russell, P. J. C. &amp; Pateman, J. &amp; Verovnik, R." box="[258,475,727,753]" pageId="3" pageNumber="28" pagination="135 - 153" refId="ref10857" refString="Russell, P. J. C., Pateman, J. &amp; Verovnik, R. (2014) First record of M. ornata Christoph, 1893, from Slovenia, with notes on its confirmed distribution and hybridisation with M. phoebe ([Denis &amp; Schiffermuller], 1775). Entomologist's Gazette, 65, 135 - 153." type="journal article" year="2014">
Russell
<emphasis id="61B6EAB4FFFBFFE5E064028DFE32FD32" box="[350,401,727,753]" italics="true" pageId="3" pageNumber="28">et al</emphasis>
. 2014
</bibRefCitation>
;
<bibRefCitation id="37534B57FFFBFFE5E0DD028DFCF2FD32" author="Pazhenkova, E. A. &amp; Lukhtanov, V. A." box="[487,849,727,753]" pageId="3" pageNumber="28" pagination="4819 - 4832" refId="ref10403" refString="Pazhenkova, E. A. &amp; Lukhtanov, V. A. (2021) Genomic introgression from a distant congener in the Levant fritillary butterfly, Melitaea acentria. Molecular Ecology, 30 (19), 4819 - 4832. https: // doi. org / 10.1111 / mec. 16085" type="journal article" year="2021">Pazhenkova &amp; Lukhtanov 2021</bibRefCitation>
); also numerous interspecific, intergeneric and even intersubfamily copulations have been observed (
<bibRefCitation id="37534B57FFFBFFE5E38302A1FCF8FCD6" author="Russell, P. J. C." box="[697,859,763,789]" pageId="3" pageNumber="28" pagination="11 - 17" refId="ref10515" refString="Russell, P. J. C. (2012 a) Intergeneric pairings involving Maniola jurtina L. (Lepidoptera: Nymphalidae: Satyrinae): are sight and smell overridden by this butterfly's sex drive? Entomologists' Gazette, 64, 11 - 17." type="journal article" year="2012">Russell 2012a</bibRefCitation>
, b). In the absence of the specimen the authors can only provide a copy of Krulikovsky’s figure of this aberration or hybrid(?) (
<figureCitation id="CBF92A23FFFBFFE5E2DD0345FB8FFCFA" box="[999,1068,799,825]" captionStart="FIGURES 7–11" captionStartId="4.[151,264,1857,1881]" captionTargetBox="[152,1436,560,1833]" captionTargetId="figure-219@4.[151,1436,560,1833]" captionTargetPageId="4" captionText="FIGURES 7–11. 7, Lectotype of Melitaea phoebe sibina Alphéraky, 1881. 8, Melitaea phoebe ab. tatara Krulikovsky, 1891. 9, Lectotype of Melitaea phoebe ornata Christoph, 1893. 10, Lectotype of Melitaea phoebe uralensis Krulikovsky, 1897 (= ufensis Krulikovsky 1902). 11, Lectotype of Melitaea phoebe var. tungusa Herz, 1899. Figs 7, 9, 11 © V.A. Lukhtanov. Figs 8, 10 © expired. A = upperside, B = underside, C = labels on specimen pin." figureDoi="http://doi.org/10.5281/zenodo.6577635" httpUri="https://zenodo.org/record/6577635/files/figure.png" pageId="3" pageNumber="28">Fig. 8</figureCitation>
).
</paragraph>
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</treatment>
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111
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<taxonomicName id="94C24D25FFFBFFE5E1FD0319FD0AFC9E" ID-CoL="7KC4R" authority="Staudinger, 1892" authorityName="Staudinger" authorityYear="1892" box="[199,681,834,861]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="variety" species="phoebe" variety="saturata">
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<bibRefCitation id="37534B57FFFBFFE5E0D30319FD0AFC9E" author="Staudinger, O." box="[489,681,835,861]" pageId="3" pageNumber="28" pagination="300 - 393" refId="ref11225" refString="Staudinger, O. (1892) Lepidopteren des Kentei-Gebirges. Deutsche Entomologische Zeitschrift, Iris, 5, 300 - 393." type="journal article" year="1892">Staudinger, 1892</bibRefCitation>
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[TL: Kentai Mountains,
<collectingCountry id="2BD57636FFFBFFE5E2F10318FB98FC9E" box="[971,1083,834,861]" name="Mongolia" pageId="3" pageNumber="28">Mongolia</collectingCountry>
] A large and brightly coloured form typical of ‘mountain forms’ (
<bibRefCitation id="37534B57FFFBFFE5E326033DFD50FC42" author="Higgins, L. G." box="[540,755,871,897]" pageId="3" pageNumber="28" pagination="175 - 365" refId="ref9372" refString="Higgins, L. G. (1941) An illustrated catalogue of the Palearctic Melitaea (Lep. Rhopalocera). Transactions of the Royal Entomological Society of London, 91, 175 - 365. https: // doi. org / 10.1111 / j. 1365 - 2311.1941. tb 01045. x" type="journal article" year="1941">Higgins 1941: 342</bibRefCitation>
).
<bibRefCitation id="37534B57FFFBFFE5E23D033DFBC4FC42" author="Van Oorschot, H. &amp; Coutsis, J. G." box="[775,1127,871,897]" pageId="3" pageNumber="28" refId="ref11693" refString="Van Oorschot, H. &amp; Coutsis, J. G. (2014) The genus Melitaea Fabricius, 1807 (Lepidoptera, Nymphalidae, Nymphalinae) Taxonomy and systematics with special reference to the male genitalia. Tshikolovets publications, Pardubice, Czech Republic, 360 pp." type="book" year="2014">Van Oorschot &amp; Coutsis (2014</bibRefCitation>
: plate 12, fig. 25) figured a brightly coloured male
<typeStatus id="8C798804FFFBFFE5E0A603D1FE50FC66" box="[412,499,907,933]" pageId="3" pageNumber="28" type="syntype">syntype</typeStatus>
from the Staudinger collection in Zoologisches Museum der Humboldt Universität, Berlin. Both authors synonymised this
<taxonomicName id="94C24D25FFFBFFE5E37203F5FD6DFC0A" box="[584,718,943,969]" pageId="3" pageNumber="28" rank="variety" variety="with">variety with</taxonomicName>
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<emphasis id="61B6EAB4FFFBFFE5E3E903F5FCEAFC0A" box="[723,841,943,969]" italics="true" pageId="3" pageNumber="28">M. phoebe</emphasis>
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, as did
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, note 80). On the internet www.funet.fi (accessed
<date id="277C1066FFFBFFE5E0C60389FDDFFC2D" box="[508,636,979,1006]" pageId="3" pageNumber="28" value="2020-10-26">26.10.2020</date>
) has a photograph (© Kosterin) of the underside of a female(?), from the Iskitim District, Novosibirst Province, Western Siberia,
<collectingCountry id="2BD57636FFFBFFE5E23903ADFC79FBD2" box="[771,986,1015,1041]" name="Russia" pageId="3" pageNumber="28">Russian Federation</collectingCountry>
on
<date id="277C1066FFFBFFE5E53803ADFB00FBD2" box="[1026,1187,1015,1041]" pageId="3" pageNumber="28" value="1997-06-21">
21
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June 1997
</date>
; this shows clearly the wing markings and antennae distinctive of
<taxonomicName id="94C24D25FFFBFFE5E3410441FD52FBF6" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[635,753,1051,1077]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFBFFE5E3410441FD52FBF6" box="[635,753,1051,1077]" italics="true" pageId="3" pageNumber="28">M. phoebe</emphasis>
</taxonomicName>
rather than of
<taxonomicName id="94C24D25FFFBFFE5E2AD0441FBA4FBF7" authorityName="Christoph" authorityYear="1893" box="[919,1031,1051,1076]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="61B6EAB4FFFBFFE5E2AD0441FBA4FBF7" box="[919,1031,1051,1076]" italics="true" pageId="3" pageNumber="28">M. ornata</emphasis>
</taxonomicName>
. We confirm that the name
<emphasis id="61B6EAB4FFFBFFE5E4050441FA3FFBF7" box="[1343,1436,1051,1076]" italics="true" pageId="3" pageNumber="28">saturata</emphasis>
is referable to
<taxonomicName id="94C24D25FFFBFFE5E0020465FE0DFB9A" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[312,430,1087,1113]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFBFFE5E0020465FE0DFB9A" box="[312,430,1087,1113]" italics="true" pageId="3" pageNumber="28">M. phoebe</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="0EF68CFBED678F083B5AA9D498332953">Reassessment of the status of some European and Asian Melitaea taxa described as subspecies of Melitaea phoebe ([Denis &amp; Schiffermüller], 1775), with designations of lectotypes where appropriate (Lepidoptera: Nymphalidae)</mods:title>
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<treatment id="DB6B87B0FFFBFFE5E1FD0439FA32F9CA" ID-DOI="http://doi.org/10.5281/zenodo.6580824" ID-GBIF-Taxon="195626717" ID-Zenodo-Dep="6580824" LSID="urn:lsid:plazi:treatment:DB6B87B0FFFBFFE5E1FD0439FA32F9CA" httpUri="http://treatment.plazi.org/id/DB6B87B0FFFBFFE5E1FD0439FA32F9CA" lastPageNumber="28" pageId="3" pageNumber="28">
<subSubSection id="1BD8652DFFFBFFE5E1FD0439FD27FBBE" box="[199,644,1122,1149]" pageId="3" pageNumber="28" type="nomenclature">
<paragraph id="537D36A6FFFBFFE5E1FD0439FD27FBBE" blockId="3.[151,1437,151,2013]" box="[199,644,1122,1149]" pageId="3" pageNumber="28">
<taxonomicName id="94C24D25FFFBFFE5E1FD0439FD27FBBE" ID-CoL="7KC4T" authority="Christoph, 1893" authorityName="Christoph" authorityYear="1893" box="[199,644,1122,1149]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="variety" species="phoebe" variety="ornata">
<emphasis id="61B6EAB4FFFBFFE5E1FD0439FE6AFBBE" bold="true" box="[199,457,1122,1149]" pageId="3" pageNumber="28">
<emphasis id="61B6EAB4FFFBFFE5E1FD0439FEE3FBBE" bold="true" box="[199,320,1122,1149]" italics="true" pageId="3" pageNumber="28">M. phoebe</emphasis>
var.
<emphasis id="61B6EAB4FFFBFFE5E0440439FE6AFBBE" bold="true" box="[382,457,1123,1149]" italics="true" pageId="3" pageNumber="28">ornata</emphasis>
</emphasis>
<bibRefCitation id="37534B57FFFBFFE5E0EB0439FD27FBBE" author="Christoph, H." box="[465,644,1123,1149]" pageId="3" pageNumber="28" pagination="86 - 89" refId="ref8897" refString="Christoph, H. (1893) Lepidoptera nova faunae Palaearcticae. Deutsche Entomologische Zeitschrift ' Iris ', 6, 86 - 89." type="journal article" year="1893">Christoph, 1893</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="1BD8652DFFFBFFE5E3B60439FA32F9CA" pageId="3" pageNumber="28" type="discussion">
<paragraph id="537D36A6FFFBFFE5E3B60439FA32F9CA" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="28">
[TL: c
<emphasis id="61B6EAB4FFFBFFE5E3E90439FD5DFBBF" box="[723,766,1123,1148]" italics="true" pageId="3" pageNumber="28">irca</emphasis>
‘Guberli’,
<emphasis id="61B6EAB4FFFBFFE5E2470439FABDFBBE" box="[893,1310,1123,1149]" italics="true" pageId="3" pageNumber="28">promontorium uralensium australium</emphasis>
(Guberlya, Southern Urals,
<collectingRegion id="9106F844FFFBFFE5E07104DDFDBCFB62" box="[331,543,1159,1185]" country="Russia" name="Orenburg" pageId="3" pageNumber="28">Orenburg Province</collectingRegion>
,
<collectingCountry id="2BD57636FFFBFFE5E31004DDFCA1FB62" box="[554,770,1158,1185]" name="Russia" pageId="3" pageNumber="28">Russian Federation</collectingCountry>
)] separated from
<taxonomicName id="94C24D25FFFBFFE5E2F804DDFB95FB62" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[962,1078,1159,1185]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFBFFE5E2F804DDFB95FB62" box="[962,1078,1159,1185]" italics="true" pageId="3" pageNumber="28">M. phoebe</emphasis>
</taxonomicName>
simultaneously by
<bibRefCitation id="37534B57FFFBFFE5E43704DDFF40FB06" author="Russell, P. J. C. &amp; Gascoigne-Pees, M. &amp; Pateman, J. E. &amp; Tennent, W. J." pageId="3" pageNumber="28" pagination="263 - 268" refId="ref10748" refString="Russell, P. J. C., Gascoigne-Pees, M., Pateman, J. E. &amp; Tennent, W. J. (2005) Melitaea emipunica (Verity, 1919) stat. nov.: a hitherto unrecognised butterfly species from Europe (Lepidoptera: Nymphalidae). Entomologist's Gazette, 56, 263 - 268." type="journal article" year="2005">
Russell
<emphasis id="61B6EAB4FFFBFFE5E45F04DDFA3EFB62" box="[1381,1437,1159,1185]" italics="true" pageId="3" pageNumber="28">et al.</emphasis>
(2005)
</bibRefCitation>
, under the name
<taxonomicName id="94C24D25FFFBFFE5E09804F1FD3CFB06" authority="Verity 1919" authorityName="Verity" authorityYear="1919" box="[418,671,1195,1221]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="emipunica">
<emphasis id="61B6EAB4FFFBFFE5E09804F1FDB7FB07" box="[418,532,1195,1220]" italics="true" pageId="3" pageNumber="28">emipunica</emphasis>
<bibRefCitation id="37534B57FFFBFFE5E32104F1FD3CFB06" author="Verity, R." box="[539,671,1195,1221]" pageId="3" pageNumber="28" pagination="26 - 31" refId="ref11898" refString="Verity, R. (1919) Seasonal Polymorphism and Races of some European Grypocera and Rhopalocera. Entomologist's Record and Journal of Variation 31 (2), 26 - 31, (3), 43 - 48, (5), 87 - 89, (7), 121 - 129, (10), 178 - 184, (11), 193 - 201." type="journal article" year="1919">Verity 1919</bibRefCitation>
</taxonomicName>
, and
<bibRefCitation id="37534B57FFFBFFE5E3E004F1FC0FFB06" author="Varga, Z. &amp; Szabo, S. &amp; Kozma, P." box="[730,940,1194,1221]" pageId="3" pageNumber="28" pagination="65 - 68" refId="ref11796" refString="Varga, Z., Szabo, S. &amp; Kozma, P. (2005) Melitaea ogygia kovacsi Varga 1967 (Lepidoptera: Nymphalidae) in the Pannonian region: taxonomy, bionomy, conservation biology. In: Kuhn, E., Feldmann, R., Thomas, J. &amp; Settele, J. (Eds.), Studies on the Ecology and Conservation of Butterflies in Europe, 1: general concepts and case studies, pp. 65 - 68. [Conference Proceedings, UFZ Leipzig - Halle, December 2005]" type="proceedings paper" year="2005">
Varga
<emphasis id="61B6EAB4FFFBFFE5E21B04F1FCF1FB06" box="[801,850,1195,1221]" italics="true" pageId="3" pageNumber="28">et al</emphasis>
. (2005)
</bibRefCitation>
, under the name
<taxonomicName id="94C24D25FFFBFFE5E55104F1FA23FB06" authority="Fruhstorfer, 1908" authorityName="Fruhstorfer" authorityYear="1908" box="[1131,1408,1195,1221]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="ogygia">
<emphasis id="61B6EAB4FFFBFFE5E55104F1FB15FB07" box="[1131,1206,1195,1220]" italics="true" pageId="3" pageNumber="28">ogygia</emphasis>
<bibRefCitation id="37534B57FFFBFFE5E58704F1FA23FB06" author="Fruhstorfer, H." box="[1213,1408,1195,1221]" pageId="3" pageNumber="28" pagination="310" refId="ref9058" refString="Fruhstorfer, H. (1908) Neue Argynnis und Melitaea. Internationale Entomologische Zeitschrift, 1, 310." type="journal article" year="1908">Fruhstorfer, 1908</bibRefCitation>
</taxonomicName>
. A
<typeStatus id="8C798804FFFBFFE5E1AD0495FF58FB2A" box="[151,251,1231,1257]" pageId="3" pageNumber="28" type="lectotype">lectotype</typeStatus>
(
<figureCitation id="CBF92A23FFFBFFE5E0310495FE04FB2A" box="[267,423,1231,1257]" captionStart="FIGURES 7–11" captionStartId="4.[151,264,1857,1881]" captionTargetBox="[152,1436,560,1833]" captionTargetId="figure-219@4.[151,1436,560,1833]" captionTargetPageId="4" captionText="FIGURES 7–11. 7, Lectotype of Melitaea phoebe sibina Alphéraky, 1881. 8, Melitaea phoebe ab. tatara Krulikovsky, 1891. 9, Lectotype of Melitaea phoebe ornata Christoph, 1893. 10, Lectotype of Melitaea phoebe uralensis Krulikovsky, 1897 (= ufensis Krulikovsky 1902). 11, Lectotype of Melitaea phoebe var. tungusa Herz, 1899. Figs 7, 9, 11 © V.A. Lukhtanov. Figs 8, 10 © expired. A = upperside, B = underside, C = labels on specimen pin." figureDoi="http://doi.org/10.5281/zenodo.6577635" httpUri="https://zenodo.org/record/6577635/files/figure.png" pageId="3" pageNumber="28">Figs 9A, B, C</figureCitation>
) selected by Lvovsky (Anikin
<emphasis id="61B6EAB4FFFBFFE5E3C50495FC8CFB2A" box="[767,815,1231,1257]" italics="true" pageId="3" pageNumber="28">et al</emphasis>
. 2017: 521) demonstrates clearly the wing and antennal characteristics of
<taxonomicName id="94C24D25FFFBFFE5E06104A9FE68FACF" authorityName="Christoph" authorityYear="1893" box="[347,459,1267,1292]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="61B6EAB4FFFBFFE5E06104A9FE68FACF" box="[347,459,1267,1292]" italics="true" pageId="3" pageNumber="28">M. ornata</emphasis>
</taxonomicName>
. Unfortunately
<bibRefCitation id="37534B57FFFBFFE5E34604A9FBA4FACD" author="Van Oorschot, H. &amp; Coutsis, J. G." box="[636,1031,1267,1294]" pageId="3" pageNumber="28" refId="ref11693" refString="Van Oorschot, H. &amp; Coutsis, J. G. (2014) The genus Melitaea Fabricius, 1807 (Lepidoptera, Nymphalidae, Nymphalinae) Taxonomy and systematics with special reference to the male genitalia. Tshikolovets publications, Pardubice, Czech Republic, 360 pp." type="book" year="2014">van Oorschot &amp; Coutsis (2014: 60)</bibRefCitation>
referred to ‘
<specimenCount id="45C4FD2FFFFBFFE5E5AB04A9FB5FFACE" box="[1169,1276,1267,1293]" pageId="3" pageNumber="28" type="male">two male</specimenCount>
syntypes’ and placed the taxon
<taxonomicName id="94C24D25FFFBFFE5E074054DFE34FAF3" authorityName="Christoph" authorityYear="1893" box="[334,407,1303,1328]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="61B6EAB4FFFBFFE5E074054DFE34FAF3" box="[334,407,1303,1328]" italics="true" pageId="3" pageNumber="28">ornata</emphasis>
</taxonomicName>
as a subspecies of
<taxonomicName id="94C24D25FFFBFFE5E35E054DFD75FAF2" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[612,726,1303,1329]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFBFFE5E35E054DFD75FAF2" box="[612,726,1303,1329]" italics="true" pageId="3" pageNumber="28">M. phoebe</emphasis>
</taxonomicName>
. Their figures (Plate 13, figs 2 &amp; 3), purporting to be ‘
<specimenCount id="45C4FD2FFFFBFFE5E424054DFA95FAF2" box="[1310,1334,1303,1329]" pageId="3" pageNumber="28" type="male">♂</specimenCount>
<typeStatus id="8C798804FFFBFFE5E406054DFA3FFAF2" box="[1340,1436,1303,1329]" pageId="3" pageNumber="28" type="syntype">syntypes</typeStatus>
from Zoologisches Museum der Humboldt Universität, Berlin’ are clearly females; all the figured specimens show the characters typical of
<taxonomicName id="94C24D25FFFBFFE5E0880505FD80FABB" authorityName="Christoph" authorityYear="1893" box="[434,547,1375,1400]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="61B6EAB4FFFBFFE5E0880505FD80FABB" box="[434,547,1375,1400]" italics="true" pageId="3" pageNumber="28">M. ornata</emphasis>
</taxonomicName>
. Most authors now accept this taxon to be a species distinct from
<taxonomicName id="94C24D25FFFBFFE5E41F0505FA3FFABA" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[1317,1436,1375,1401]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFBFFE5E41F0505FA3FFABA" box="[1317,1436,1375,1401]" italics="true" pageId="3" pageNumber="28">M. phoebe</emphasis>
</taxonomicName>
(
<bibRefCitation id="37534B57FFFBFFE5E1A405D9FE05FA5E" author="Tshikolovets, V. V." box="[158,422,1411,1437]" pageId="3" pageNumber="28" refId="ref11624" refString="Tshikolovets, V. V. (2011) Butterflies of Europe &amp; the Mediterranean area. Tshikolovets Publications, Pardubice, Czech Republic, 544 pp." type="book" year="2011">Tshikolovets 2011: 498</bibRefCitation>
;
<bibRefCitation id="37534B57FFFBFFE5E08905D9FD45FA5D" author="Lafranchis, T. &amp; Jutzeler, D. &amp; Guillosson, J-Y. &amp; Kan, P. &amp; Kan, B." box="[435,742,1410,1438]" pageId="3" pageNumber="28" refId="ref10054" refString="Lafranchis, T., Jutzeler, D., Guillosson, J-Y., Kan, P. &amp; Kan, B. (2015) La Vie des Papillons, Ecologie, Biologie et Comportement des Rhopaloceres de France. Diatheo, Paris, 751 pp." type="book" year="2015">
Lafranchis
<emphasis id="61B6EAB4FFFBFFE5E30B05D9FDC1FA5E" box="[561,610,1411,1437]" italics="true" pageId="3" pageNumber="28">et al</emphasis>
. 2015: 468
</bibRefCitation>
). Some authors have used different names for the species (see
<bibRefCitation id="37534B57FFFBFFE5E1AD05FDFE00FA01" author="Russell, PJ &amp; Tennent, W. J." box="[151,419,1447,1474]" pageId="3" pageNumber="28" pagination="27 - 56" refId="ref10922" refString="Russell, PJ. C. &amp; Tennent, W. J. (2016) A synonymic list of names associated with western Palaearctic Melitaea phoebe (Denis &amp; Schiffermuller, 1775) species group taxa M. phoebe; M. punica Oberthur, 1876; M. ornata Christoph, 1893) (Lepidoptera, Nymphalidae). Nota Lepidopterologica, 39 (1), 27 - 56. https: // doi. org / 10.3897 / nl. 39.5929" type="journal article" year="2016">Russell &amp; Tennent 2016</bibRefCitation>
), whilst others continue to consider all
<taxonomicName id="94C24D25FFFBFFE5E27105FDFC1DFA02" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[843,958,1447,1473]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFBFFE5E27105FDFC1DFA02" box="[843,958,1447,1473]" italics="true" pageId="3" pageNumber="28">M. phoebe</emphasis>
</taxonomicName>
group taxa to be conspecific (
<bibRefCitation id="37534B57FFFBFFE5E43F05FDFEA9FA26" author="Kudrna, O. &amp; Pennstorfer, J &amp; Lux, K." pageId="3" pageNumber="28" refId="ref10020" refString="Kudrna, O., Pennstorfer, J &amp; Lux, K. (2015) Distribution Atlas of European Butterflies and Skippers. Wissenschsftlicher PEKS, Schwanfeld, Germany, 632 pp." type="book" year="2015">
Kudrna
<emphasis id="61B6EAB4FFFBFFE5E46505FDFA2DFA02" box="[1375,1422,1447,1473]" italics="true" pageId="3" pageNumber="28">et al</emphasis>
., 2015: 379
</bibRefCitation>
). However, a large set of molecular and morphological data unequivocally indicate that
<taxonomicName id="94C24D25FFFBFFE5E5E30591FAE4FA27" authorityName="Christoph" authorityYear="1893" box="[1241,1351,1483,1508]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="61B6EAB4FFFBFFE5E5E30591FAE4FA27" box="[1241,1351,1483,1508]" italics="true" pageId="3" pageNumber="28">M. ornata</emphasis>
</taxonomicName>
and
<taxonomicName id="94C24D25FFFBFFE5E4440591FF45F9CA" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFBFFE5E4440591FF45F9CA" italics="true" pageId="3" pageNumber="28">M. phoebe</emphasis>
</taxonomicName>
are different species, which are found sympatrically in a number of localities (
<bibRefCitation id="37534B57FFFBFFE5E57B05B5FB59F9CA" author="Toth, J. P. &amp; Bereczki, J. &amp; Varga, Z. &amp; Rota, J. &amp; Sramko, G. &amp; Wahlberg, N." box="[1089,1274,1519,1545]" pageId="3" pageNumber="28" pagination="749 - 757" refId="ref11329" refString="Toth, J. P., Bereczki, J., Varga, Z., Rota, J., Sramko, G. &amp; Wahlberg, N. (2014) Relationships within the Melitaea phoebe species group (Lepidoptera: Nymphalidae): new insights from molecular and morphometric information. Systematic Entomology, 39, 749 - 757. https: // doi. org / 10.1111 / syen. 12083" type="journal article" year="2014">
Tóth
<emphasis id="61B6EAB4FFFBFFE5E54705B5FB17F9CA" box="[1149,1204,1519,1545]" italics="true" pageId="3" pageNumber="28">et al.</emphasis>
, 2014
</bibRefCitation>
,
<bibRefCitation id="37534B57FFFBFFE5E43F05B5FA9DF9CA" author="Toth, J. P. &amp; Varga, Z. &amp; Bereczki, J." box="[1285,1342,1519,1545]" pageId="3" pageNumber="28" pagination="106 - 115" refId="ref11405" refString="Toth, J. P., Varga, Z. &amp; Bereczki, J. (2016) Long-term survival and diversification of an endemic Melitaea species in mountains of Iran and adjacent areas. Journal of Zoological Systematics and Evolutionary Research, 54 (2), 106 - 115. https: // doi. org / 10.1111 / jzs. 12122" type="journal article" year="2016">2016</bibRefCitation>
,
<bibRefCitation id="37534B57FFFBFFE5E47305B5FA26F9CA" author="Toth, J. P. &amp; Varga, Z. &amp; Verovnik, R. &amp; Wahlberg, N. &amp; Varadi, A. &amp; Bereczki, J." box="[1353,1413,1519,1545]" pageId="3" pageNumber="28" pagination="267 - 281" refId="ref11547" refString="Toth, J. P., Varga, Z., Verovnik, R., Wahlberg, N., Varadi, A. &amp; Bereczki, J. (2017) Mito-nuclear discordance helps to reveal the phylogeographic patterns of Melitaea ornata (Lepidoptera: Nymphalidae). Biological Journal of the Linnean Society, 121, 267 - 281. https: // doi. org / 10.1093 / biolinnean / blw 037" type="journal article" year="2017">2017</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

159
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<mods:title id="664FECB9A5EAE9FC07495D719FA73082">Reassessment of the status of some European and Asian Melitaea taxa described as subspecies of Melitaea phoebe ([Denis &amp; Schiffermüller], 1775), with designations of lectotypes where appropriate (Lepidoptera: Nymphalidae)</mods:title>
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<paragraph id="537D36A6FFFBFFE5E1FD0649FD65F9EE" blockId="3.[151,1437,151,2013]" box="[199,710,1554,1581]" pageId="3" pageNumber="28">
<taxonomicName id="94C24D25FFFBFFE5E1FD0649FD65F9EE" ID-CoL="3ZKHS" authority="Krulikovsky, 1897" authorityName="Krulikovsky" authorityYear="1897" box="[199,710,1554,1581]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="variety" species="phoebe" variety="uralensis">
<emphasis id="61B6EAB4FFFBFFE5E1FD0649FE48F9EE" bold="true" box="[199,491,1554,1581]" pageId="3" pageNumber="28">
<emphasis id="61B6EAB4FFFBFFE5E1FD0649FEE1F9EE" bold="true" box="[199,322,1554,1581]" italics="true" pageId="3" pageNumber="28">M. phoebe</emphasis>
var.
<emphasis id="61B6EAB4FFFBFFE5E0BB0648FE48F9EE" bold="true" box="[385,491,1554,1581]" italics="true" pageId="3" pageNumber="28">uralensis</emphasis>
</emphasis>
<bibRefCitation id="37534B57FFFBFFE5E0CE0649FD65F9EE" author="Krulikovsky, A." box="[500,710,1555,1581]" pageId="3" pageNumber="28" pagination="315 - 328" refId="ref9945" refString="Krulikovsky, A. (1897) Notes on the fauna Macrolepidoptera of the area of the city of Ufa. Metrialy k Pooznaniyu Fauny i Flory Rossiiskoi Imperialis, Otdel Zoologicheskij, 3, 315 - 328. [In Russian]" type="journal article" year="1897">Krulikovsky, 1897</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="1BD8652DFFFBFFE5E3F50649FDF8F81E" pageId="3" pageNumber="28" type="discussion">
<paragraph id="537D36A6FFFBFFE5E3F50649FDF8F81E" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="28">
[TL: District of Ufa,
<collectingCountry id="2BD57636FFFBFFE5E2F90649FB03F9EE" box="[963,1184,1554,1581]" name="Russia" pageId="3" pageNumber="28">Russian Federation</collectingCountry>
] a name subsequently replaced by
<taxonomicName id="94C24D25FFFBFFE5E01C066DFD70F992" authority="Krulikovsky, 1902" authorityName="Krulikovsky" authorityYear="1902" box="[294,723,1591,1617]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="ufensis">
<emphasis id="61B6EAB4FFFBFFE5E01C066DFE55F992" box="[294,502,1591,1617]" italics="true" pageId="3" pageNumber="28">M. phoebe ufensis</emphasis>
<bibRefCitation id="37534B57FFFBFFE5E33A066DFD70F992" author="Krulikovsky, A." box="[512,723,1591,1617]" pageId="3" pageNumber="28" pagination="535 - 560" refId="ref9988" refString="Krulikovsky, A. (1902) On the fauna of Lepidoptera of the Yaroslavl Province. Horae Societatis Entomologicae Rossicae, 35, 535 - 560. [In Russian]" type="journal article" year="1902">Krulikovsky, 1902</bibRefCitation>
</taxonomicName>
(
<emphasis id="61B6EAB4FFFBFFE5E3DC066DFCEEF992" box="[742,845,1591,1617]" italics="true" pageId="3" pageNumber="28">uralensis</emphasis>
preoccupied by
<taxonomicName id="94C24D25FFFBFFE5E535066DFF70F9B6" authority="Eversmann, 1844" authorityName="Eversmann" authorityYear="1844" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="subSpecies" species="arduinna" subSpecies="uralensis">
<emphasis id="61B6EAB4FFFBFFE5E535066DFAADF992" box="[1039,1294,1591,1617]" italics="true" pageId="3" pageNumber="28">M. arduinna uralensis</emphasis>
<bibRefCitation id="37534B57FFFBFFE5E42D066DFF70F9B6" author="Eversmann, E. F." pageId="3" pageNumber="28" refId="ref9016" refString="Eversmann, E. F. (1844) Fauna lepidopterologica Volgo-Uralensis. Exhibens. Lepidopterorum species quar per quinque annos in provinciis Volgam fluvium inter et montes Uralenses sitis observavit et descripsit. Typis Universitatis, Casani, 633 рp." type="book" year="1844">Eversmann, 1844</bibRefCitation>
</taxonomicName>
).
<bibRefCitation id="37534B57FFFBFFE5E1DC0601FE6AF9B6" author="Higgins, L. G." box="[230,457,1627,1653]" pageId="3" pageNumber="28" pagination="175 - 365" refId="ref9372" refString="Higgins, L. G. (1941) An illustrated catalogue of the Palearctic Melitaea (Lep. Rhopalocera). Transactions of the Royal Entomological Society of London, 91, 175 - 365. https: // doi. org / 10.1111 / j. 1365 - 2311.1941. tb 01045. x" type="journal article" year="1941">Higgins (1941: 343)</bibRefCitation>
equated this name to
<taxonomicName id="94C24D25FFFBFFE5E3870601FC2BF9B6" authority="Staudinger" authorityName="Staudinger" box="[701,904,1627,1653]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="61B6EAB4FFFBFFE5E3870601FCABF9B7" box="[701,776,1627,1652]" italics="true" pageId="3" pageNumber="28">ornata</emphasis>
Staudinger
</taxonomicName>
(an error of authority). This synonymy with
<taxonomicName id="94C24D25FFFBFFE5E4440601FF41F95B" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="61B6EAB4FFFBFFE5E4440601FF41F95B" italics="true" pageId="3" pageNumber="28">M. ornata</emphasis>
</taxonomicName>
was accepted by
<bibRefCitation id="37534B57FFFBFFE5E09E0625FD46F95A" author="Russell, PJ &amp; Tennent, W. J." box="[420,741,1662,1689]" pageId="3" pageNumber="28" pagination="27 - 56" refId="ref10922" refString="Russell, PJ. C. &amp; Tennent, W. J. (2016) A synonymic list of names associated with western Palaearctic Melitaea phoebe (Denis &amp; Schiffermuller, 1775) species group taxa M. phoebe; M. punica Oberthur, 1876; M. ornata Christoph, 1893) (Lepidoptera, Nymphalidae). Nota Lepidopterologica, 39 (1), 27 - 56. https: // doi. org / 10.3897 / nl. 39.5929" type="journal article" year="2016">Russell &amp; Tennent (2016: 56</bibRefCitation>
, note 98); however,
<bibRefCitation id="37534B57FFFBFFE5E2FD0625FAEEF959" author="Van Oorschot, H. &amp; Coutsis, J. G." box="[967,1357,1663,1690]" pageId="3" pageNumber="28" refId="ref11693" refString="Van Oorschot, H. &amp; Coutsis, J. G. (2014) The genus Melitaea Fabricius, 1807 (Lepidoptera, Nymphalidae, Nymphalinae) Taxonomy and systematics with special reference to the male genitalia. Tshikolovets publications, Pardubice, Czech Republic, 360 pp." type="book" year="2014">van Oorschot &amp; Coutsis (2014: 60)</bibRefCitation>
placed it as a synonym of
<taxonomicName id="94C24D25FFFBFFE5E05B06F9FE75F97E" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[353,470,1699,1725]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFBFFE5E05B06F9FE75F97E" box="[353,470,1699,1725]" italics="true" pageId="3" pageNumber="28">M. phoebe</emphasis>
</taxonomicName>
, possibly because they incorrectly synonymised
<taxonomicName id="94C24D25FFFBFFE5E2D706F9FBFFF97F" box="[1005,1116,1699,1724]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="61B6EAB4FFFBFFE5E2D706F9FBFFF97F" box="[1005,1116,1699,1724]" italics="true" pageId="3" pageNumber="28">M. ornata</emphasis>
</taxonomicName>
with
<taxonomicName id="94C24D25FFFBFFE5E5AD06F9FAA8F97E" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[1175,1291,1699,1725]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFBFFE5E5AD06F9FAA8F97E" box="[1175,1291,1699,1725]" italics="true" pageId="3" pageNumber="28">M. phoebe</emphasis>
</taxonomicName>
;
<bibRefCitation id="37534B57FFFBFFE5E42206F9FEBCF922" author="Hesselbarth, G. &amp; van Oorschot, H. &amp; Wagener, S." pageId="3" pageNumber="28" pagination="1 - 847" refId="ref9287" refString="Hesselbarth, G., van Oorschot, H. &amp; Wagener, S. (1995) Die Tagfalter der Turkei unter Berucksichtigung der angrenzenden Lander, Sigbert Wagener, Bocholt, Germany, 1, 1 - 757, 2, 758 - 1354, 3, 1 - 847." type="journal article" year="1995">
Hesselbarth
<emphasis id="61B6EAB4FFFBFFE5E1AD069DFF6CF922" box="[151,207,1735,1761]" italics="true" pageId="3" pageNumber="28">et al.</emphasis>
(1995)
</bibRefCitation>
overlooked this name. It is known that both
<taxonomicName id="94C24D25FFFBFFE5E237069DFC20F922" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[781,899,1735,1761]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFBFFE5E237069DFC20F922" box="[781,899,1735,1761]" italics="true" pageId="3" pageNumber="28">M. phoebe</emphasis>
</taxonomicName>
and
<taxonomicName id="94C24D25FFFBFFE5E282069DFB84F923" box="[952,1063,1735,1760]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="61B6EAB4FFFBFFE5E282069DFB84F923" box="[952,1063,1735,1760]" italics="true" pageId="3" pageNumber="28">M. ornata</emphasis>
</taxonomicName>
fly in the southern Urals (
<bibRefCitation id="37534B57FFFBFFE5E472069DFEC2F8C6" author="Russell, P. J. C. &amp; Tennent, W. J." pageId="3" pageNumber="28" pagination="199 - 212" refId="ref11007" refString="Russell, P. J. C. &amp; Tennent, W. J. (2022) Butterflies observed on the western (European) slopes of the southern Ural Mountains during three spring visits in 2011, 2012 and 2014 (Lepidoptera Papilionoidea). Neue Entomologische Nachrichten 80: 199 - 212. [including 8 colour plates]" type="journal article" year="2022">Russell &amp; Tennent (2022)</bibRefCitation>
, see
<figureCitation id="CBF92A23FFFBFFE5E0AC06B1FDADF8C6" box="[406,526,1771,1797]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1717,1741]" captionTargetBox="[153,1436,200,1692]" captionTargetId="figure-26@2.[151,1436,181,1693]" captionTargetPageId="2" captionText="FIGURES 1–6. 1, Melitaea ornata male underside, Russian Federation, southern Urals, Orenburg Province, Guberli, ca. 190 m, 14 May 2014. 2, Melitaea phoebe male underside, Russian Federation, southern Urals, Orenburg Province, Krasnoznamenka, ca. 210 m, 16 May 2014. 3, Melitaea ornata female underside, Slovenia, Koper, ca. 2km NW of Rakitovec ca. 525 m, 17 May 2011. 4, Melitaea phoebe female underside, Slovenia, Koper, ca. 2km S. of Rakitovec ca. 525 m, 17 May 2011. 5, Lectotype female of Melitaea phoebe caucasica Staudinger, 1870. 6, Paralectotype male of Melitaea phoebe caucasica Staudinger, 1870. Figs 1–4 © P. Russell. Figs 5–6 © T. Léger. A = upperside, B = underside, C = labels on specimen pin." figureDoi="http://doi.org/10.5281/zenodo.6577631" httpUri="https://zenodo.org/record/6577631/files/figure.png" pageId="3" pageNumber="28">Figs 1 &amp; 2</figureCitation>
). Anikin
<emphasis id="61B6EAB4FFFBFFE5E34F06B1FD0EF8C6" box="[629,685,1771,1797]" italics="true" pageId="3" pageNumber="28">et al.</emphasis>
(2017: 522) and
<bibRefCitation id="37534B57FFFBFFE5E25606B1FB75F8C6" author="Kostjuk I. Y. &amp; Zolotuhin V. V." box="[876,1238,1771,1797]" pageId="3" pageNumber="28" pagination="586 - 593" refId="ref9806" refString="Kostjuk I. Y. &amp; Zolotuhin V. V. (2017) The types of moths and butterflies described from the Volga-Ural Region by Leonid Krulikowsky (Krulikovsky). Part 2. Macrolepidoptera. In: Anikin V. V., Sachkov S. A., Zolotuhin, V. V. (Eds.), Fauna lepidopterologica Volgo-Uralensis: from P. Pallas to present days. Munich-Vilnius. Proceedings of the Museum Witt. Munich, 7, 443 - 450, 586 - 593." type="journal article" year="2017">Kostjuk &amp; Zolotuhin (2017: 446</bibRefCitation>
&amp; plate 24, fig 4) figured the ‘Typus’, a specimen taken at Ufa on
<date id="277C1066FFFBFFE5E3940755FCE1F8EA" box="[686,834,1807,1833]" pageId="3" pageNumber="28" value="1894-06-10">10 June 1894</date>
(= 24 June in today’s Gregorian calendar, not adopted in
<collectingCountry id="2BD57636FFFBFFE5E1890769FF5DF88E" box="[179,254,1843,1869]" name="Russia" pageId="3" pageNumber="28">Russia</collectingCountry>
until
<date id="277C1066FFFBFFE5E0040769FE47F88E" box="[318,484,1843,1869]" pageId="3" pageNumber="28" value="1918-02">February 1918</date>
), which they designated as the
<typeStatus id="8C798804FFFBFFE5E2040769FC0BF88E" box="[830,936,1843,1869]" pageId="3" pageNumber="28" type="lectotype">lectotype</typeStatus>
. The specimen is figured here with the labels (
<figureCitation id="CBF92A23FFFBFFE5E1A5070DFEFCF8B2" box="[159,351,1879,1905]" captionStart="FIGURES 7–11" captionStartId="4.[151,264,1857,1881]" captionTargetBox="[152,1436,560,1833]" captionTargetId="figure-219@4.[151,1436,560,1833]" captionTargetPageId="4" captionText="FIGURES 7–11. 7, Lectotype of Melitaea phoebe sibina Alphéraky, 1881. 8, Melitaea phoebe ab. tatara Krulikovsky, 1891. 9, Lectotype of Melitaea phoebe ornata Christoph, 1893. 10, Lectotype of Melitaea phoebe uralensis Krulikovsky, 1897 (= ufensis Krulikovsky 1902). 11, Lectotype of Melitaea phoebe var. tungusa Herz, 1899. Figs 7, 9, 11 © V.A. Lukhtanov. Figs 8, 10 © expired. A = upperside, B = underside, C = labels on specimen pin." figureDoi="http://doi.org/10.5281/zenodo.6577635" httpUri="https://zenodo.org/record/6577635/files/figure.png" pageId="3" pageNumber="28">Figures 10A &amp; B</figureCitation>
). From the underside of the specimen it can be seen that the submarginal black arches are slim and touch the intervening veins and the antenna are club-shaped rather than spatulate; thus the present authors consider that this specimen is
<taxonomicName id="94C24D25FFFBFFE5E0B907C5FE59F87A" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[387,506,1951,1977]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFBFFE5E0B907C5FE59F87A" box="[387,506,1951,1977]" italics="true" pageId="3" pageNumber="28">M. phoebe</emphasis>
</taxonomicName>
rather than
<taxonomicName id="94C24D25FFFBFFE5E3B807C5FD51F87B" box="[642,754,1951,1976]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="61B6EAB4FFFBFFE5E3B807C5FD51F87B" box="[642,754,1951,1976]" italics="true" pageId="3" pageNumber="28">M. ornata</emphasis>
</taxonomicName>
. Krulikovsky probably considered this to be a
<taxonomicName id="94C24D25FFFBFFE5E43807C5FADCF87A" box="[1282,1407,1951,1977]" pageId="3" pageNumber="28" rank="variety" variety="due">variety due</taxonomicName>
to its heavily marked and dark appearance.
</paragraph>
</subSubSection>
</treatment>
</document>

109
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<mods:title id="0A652CC1F73598E2320FC3FCAC3D33F8">Reassessment of the status of some European and Asian Melitaea taxa described as subspecies of Melitaea phoebe ([Denis &amp; Schiffermüller], 1775), with designations of lectotypes where appropriate (Lepidoptera: Nymphalidae)</mods:title>
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<taxonomicName id="94C24D25FFFCFFE2E1FD00CDFDBBFF73" ID-CoL="7KC4K" authority="Herz, 1899" authorityName="Herz" authorityYear="1899" box="[199,536,150,177]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="tungusa">
<emphasis id="61B6EAB4FFFCFFE2E1FD00CDFE3FFF72" bold="true" box="[199,412,150,177]" italics="true" pageId="4" pageNumber="29">M.phoebe tungusa</emphasis>
<bibRefCitation id="37534B57FFFCFFE2E09A00CDFDBBFF73" author="Herz, O." box="[416,536,150,177]" pageId="4" pageNumber="29" pagination="209 - 265" refId="ref9249" refString="Herz, O. (1899) Reise nach Nordost-Sibirien in das Lenagebiet in den Jahren 1888 und 1889 nebst einem Verzeichnisse der dort erbeuteten Macrolepidopteren. Deutsche Entomologische Zeitschrift Iris, 11, 209 - 265." type="journal article" year="1899">Herz,1899</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
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<paragraph id="537D36A6FFFCFFE2E32100CDFCA5FDDA" blockId="4.[151,1437,150,537]" pageId="4" pageNumber="29">
[TL:
<collectingCountry id="2BD57636FFFCFFE2E36800CCFC8BFF73" box="[594,808,150,176]" name="Russia" pageId="4" pageNumber="29">Russian Federation</collectingCountry>
,Yakutia,River Witim];wing morphology and particularly the club shaped antenna indicate that this subspecies is referable to
<taxonomicName id="94C24D25FFFCFFE2E2AF00E1FBAFFF17" baseAuthorityName="Denis &amp; Schiffermuller" baseAuthorityYear="1775" box="[917,1036,186,212]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFCFFE2E2AF00E1FBAFFF17" box="[917,1036,186,212]" italics="true" pageId="4" pageNumber="29">M. phoebe</emphasis>
</taxonomicName>
. Lukhtanov here designates a male as
<typeStatus id="8C798804FFFCFFE2E18C0084FEBEFF3B" box="[182,285,222,248]" pageId="4" pageNumber="29" type="lectotype">lectotype</typeStatus>
for the name
<emphasis id="61B6EAB4FFFCFFE2E0820085FDB2FF3B" box="[440,529,223,248]" italics="true" pageId="4" pageNumber="29">tungusa</emphasis>
<bibRefCitation id="37534B57FFFCFFE2E3220085FD34FF3B" author="Herz, O." box="[536,663,222,249]" pageId="4" pageNumber="29" pagination="209 - 265" refId="ref9249" refString="Herz, O. (1899) Reise nach Nordost-Sibirien in das Lenagebiet in den Jahren 1888 und 1889 nebst einem Verzeichnisse der dort erbeuteten Macrolepidopteren. Deutsche Entomologische Zeitschrift Iris, 11, 209 - 265." type="journal article" year="1899">Herz, 1899</bibRefCitation>
. The
<typeStatus id="8C798804FFFCFFE2E3EE0084FC98FF3B" box="[724,827,222,248]" pageId="4" pageNumber="29" type="lectotype">lectotype</typeStatus>
is shown in
<figureCitation id="CBF92A23FFFCFFE2E2F30084FB29FF3A" box="[969,1162,222,249]" captionStart="FIGURES 7–11" captionStartId="4.[151,264,1857,1881]" captionTargetBox="[152,1436,560,1833]" captionTargetId="figure-219@4.[151,1436,560,1833]" captionTargetPageId="4" captionText="FIGURES 7–11. 7, Lectotype of Melitaea phoebe sibina Alphéraky, 1881. 8, Melitaea phoebe ab. tatara Krulikovsky, 1891. 9, Lectotype of Melitaea phoebe ornata Christoph, 1893. 10, Lectotype of Melitaea phoebe uralensis Krulikovsky, 1897 (= ufensis Krulikovsky 1902). 11, Lectotype of Melitaea phoebe var. tungusa Herz, 1899. Figs 7, 9, 11 © V.A. Lukhtanov. Figs 8, 10 © expired. A = upperside, B = underside, C = labels on specimen pin." figureDoi="http://doi.org/10.5281/zenodo.6577635" httpUri="https://zenodo.org/record/6577635/files/figure.png" pageId="4" pageNumber="29">Figures 11A &amp; B</figureCitation>
, together with the labels (
<figureCitation id="CBF92A23FFFCFFE2E19A0158FEA7FEDF" box="[160,260,258,284]" captionStart="FIGURES 7–11" captionStartId="4.[151,264,1857,1881]" captionTargetBox="[152,1436,560,1833]" captionTargetId="figure-219@4.[151,1436,560,1833]" captionTargetPageId="4" captionText="FIGURES 7–11. 7, Lectotype of Melitaea phoebe sibina Alphéraky, 1881. 8, Melitaea phoebe ab. tatara Krulikovsky, 1891. 9, Lectotype of Melitaea phoebe ornata Christoph, 1893. 10, Lectotype of Melitaea phoebe uralensis Krulikovsky, 1897 (= ufensis Krulikovsky 1902). 11, Lectotype of Melitaea phoebe var. tungusa Herz, 1899. Figs 7, 9, 11 © V.A. Lukhtanov. Figs 8, 10 © expired. A = upperside, B = underside, C = labels on specimen pin." figureDoi="http://doi.org/10.5281/zenodo.6577635" httpUri="https://zenodo.org/record/6577635/files/figure.png" pageId="4" pageNumber="29">Fig. 11C</figureCitation>
). The labels on the pin are as follows: (1) on red paper handwritten in black “
<typeStatus id="8C798804FFFCFFE2E55A0158FB71FEDF" box="[1120,1234,258,284]" pageId="4" pageNumber="29" type="lectotype">Lectotype</typeStatus>
/
<taxonomicName id="94C24D25FFFCFFE2E5DB0158FF42FE83" authority="Knock" authorityName="Knock" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="phoebe">Melitaea Phoebe Knock</taxonomicName>
/
<taxonomicName id="94C24D25FFFCFFE2E1C8017DFE62FE83" authority="Herz" authorityName="Herz" authorityYear="1899" box="[242,449,294,321]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="variety" species="phoebe" variety="tungusa">var. Tungusa Herz</taxonomicName>
/ V. Lukhtanov design; (2) on black-edged white paper, printed in black “Кол.[лекциЯ] Вел.[икого] Кн.[ЯЗЯ]/ НиколаЯ/ Михайловича” (translates to: Collection of Grand Duke Nikolai Mikhailovich); (3) on white paper with a broad black surround printed in black on the upperside “Witim” and on the underside “O. Herz”; (4) on white paper (discolouring to brown) handwritten in black “1/12/7” (This means: 01 July [1888] (old style)/12 July [1888] (new style). No year is given on the data label but it can be deduced from
<bibRefCitation id="37534B57FFFCFFE2E5E901EDFA3BFE13" author="Herz, O." box="[1235,1432,438,465]" pageId="4" pageNumber="29" pagination="209 - 265" refId="ref9249" refString="Herz, O. (1899) Reise nach Nordost-Sibirien in das Lenagebiet in den Jahren 1888 und 1889 nebst einem Verzeichnisse der dort erbeuteten Macrolepidopteren. Deutsche Entomologische Zeitschrift Iris, 11, 209 - 265." type="journal article" year="1899">Herz (1899: 240)</bibRefCitation>
. This diagnosis agrees with those of
<bibRefCitation id="37534B57FFFCFFE2E3180180FCA5FE37" author="Higgins, L. G." box="[546,774,474,500]" pageId="4" pageNumber="29" pagination="175 - 365" refId="ref9372" refString="Higgins, L. G. (1941) An illustrated catalogue of the Palearctic Melitaea (Lep. Rhopalocera). Transactions of the Royal Entomological Society of London, 91, 175 - 365. https: // doi. org / 10.1111 / j. 1365 - 2311.1941. tb 01045. x" type="journal article" year="1941">Higgins (1941: 342)</bibRefCitation>
,
<bibRefCitation id="37534B57FFFCFFE2E22B0180FBFEFE37" author="Korb, S. K. &amp; Bolshakov, L. V." box="[785,1117,474,500]" pageId="4" pageNumber="29" pagination="1 - 123" refId="ref9703" refString="Korb, S. K. &amp; Bolshakov, L. V. (2011) A catalogue of butterflies (Lepidoptera: Papilioformes) of the former USSR. Second edition, reformatted and updated. Eversmannia Supplement 2, 1 - 123. [In Russian]" type="journal article" year="2011">Korb &amp; Bolshakov (2011: 41)</bibRefCitation>
and
<bibRefCitation id="37534B57FFFCFFE2E5AB0181FEB3FDDA" author="Van Oorschot, H. &amp; Coutsis, J. G." pageId="4" pageNumber="29" refId="ref11693" refString="Van Oorschot, H. &amp; Coutsis, J. G. (2014) The genus Melitaea Fabricius, 1807 (Lepidoptera, Nymphalidae, Nymphalinae) Taxonomy and systematics with special reference to the male genitalia. Tshikolovets publications, Pardubice, Czech Republic, 360 pp." type="book" year="2014">van Oorschot &amp; Coutsis (2014: 60)</bibRefCitation>
; not considered by
<bibRefCitation id="37534B57FFFCFFE2E0D301A4FCA1FDDA" author="Hesselbarth, G. &amp; van Oorschot, H. &amp; Wagener, S." box="[489,770,510,537]" pageId="4" pageNumber="29" pagination="1 - 847" refId="ref9287" refString="Hesselbarth, G., van Oorschot, H. &amp; Wagener, S. (1995) Die Tagfalter der Turkei unter Berucksichtigung der angrenzenden Lander, Sigbert Wagener, Bocholt, Germany, 1, 1 - 757, 2, 758 - 1354, 3, 1 - 847." type="journal article" year="1995">
Hesselbarth
<emphasis id="61B6EAB4FFFCFFE2E34F01A5FD05FDDB" box="[629,678,510,536]" italics="true" pageId="4" pageNumber="29">et al</emphasis>
. (1995)
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.
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[TL Mongolia] a specimen of unspecified sex figured by Seitz (plate 65, fig. h, 1) is large and heavily marked, particularly the forewing post-discal black square spots.
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(see above) and his illustrations (
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: plate 14, figs 7 &amp; 8), show a similarity in spite of the large disparity in size.
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(D), they did not illustrate either but placed this form under
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<emphasis id="61B6EAB4FFFDFFE3E1FD00CDFE2DFF72" bold="true" box="[199,398,150,177]" italics="true" pageId="5" pageNumber="30">M. phoebe telona</emphasis>
<bibRefCitation id="37534B57FFFDFFE3E0AE00CDFDF4FF72" author="Fruhstorfer, H." box="[404,599,151,177]" pageId="5" pageNumber="30" pagination="310" refId="ref9058" refString="Fruhstorfer, H. (1908) Neue Argynnis und Melitaea. Internationale Entomologische Zeitschrift, 1, 310." type="journal article" year="1908">Fruhstorfer, 1908</bibRefCitation>
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[TL: ‘Palastina’ – generally accepted as environs of
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,
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]; this taxon has been elevated fairly recently to specific status by
<bibRefCitation id="37534B57FFFDFFE3E20C00E1FC5CFF16" author="Toth, J. P. &amp; Bereczki, J. &amp; Varga, Z. &amp; Rota, J. &amp; Sramko, G. &amp; Wahlberg, N." box="[822,1023,187,213]" pageId="5" pageNumber="30" pagination="749 - 757" refId="ref11329" refString="Toth, J. P., Bereczki, J., Varga, Z., Rota, J., Sramko, G. &amp; Wahlberg, N. (2014) Relationships within the Melitaea phoebe species group (Lepidoptera: Nymphalidae): new insights from molecular and morphometric information. Systematic Entomology, 39, 749 - 757. https: // doi. org / 10.1111 / syen. 12083" type="journal article" year="2014">
Tóth
<emphasis id="61B6EAB4FFFDFFE3E24800E1FC07FF16" box="[882,932,187,213]" italics="true" pageId="5" pageNumber="30">et al</emphasis>
. (2014)
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,
<bibRefCitation id="37534B57FFFDFFE3E53100E1FACEFF16" author="Van Oorschot, H. &amp; Coutsis, J. G." box="[1035,1389,187,213]" pageId="5" pageNumber="30" refId="ref11693" refString="Van Oorschot, H. &amp; Coutsis, J. G. (2014) The genus Melitaea Fabricius, 1807 (Lepidoptera, Nymphalidae, Nymphalinae) Taxonomy and systematics with special reference to the male genitalia. Tshikolovets publications, Pardubice, Czech Republic, 360 pp." type="book" year="2014">van Oorschot &amp; Coutsis (2014)</bibRefCitation>
and confirmed recently by
<bibRefCitation id="37534B57FFFDFFE3E0AC0085FDC2FF3A" author="Toth, J. P. &amp; Varga, Z. &amp; Verovnik, R. &amp; Wahlberg, N. &amp; Varadi, A. &amp; Bereczki, J." box="[406,609,223,249]" pageId="5" pageNumber="30" pagination="267 - 281" refId="ref11547" refString="Toth, J. P., Varga, Z., Verovnik, R., Wahlberg, N., Varadi, A. &amp; Bereczki, J. (2017) Mito-nuclear discordance helps to reveal the phylogeographic patterns of Melitaea ornata (Lepidoptera: Nymphalidae). Biological Journal of the Linnean Society, 121, 267 - 281. https: // doi. org / 10.1093 / biolinnean / blw 037" type="journal article" year="2017">
Tóth
<emphasis id="61B6EAB4FFFDFFE3E0E90085FDA7FF3A" box="[467,516,223,249]" italics="true" pageId="5" pageNumber="30">et al</emphasis>
. (2017)
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. There is considerable confusion between authors regarding Fruhstorfer’s
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material; this is being actively investigated and the results will be published separately in due course.
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[TL:
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,
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, Sajan (or Sayan) Mountains] uppersides of a male and female were figured by Seitz (plate 65, figs h 6 &amp; 7). www.funet.fi (accessed
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), however, gives a photograph (© Smirnov) of both upper and undersides of a female of this subspecies from
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, West of the Ujukskiy Mountains (
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, from the underside of which the typical characters of
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<taxonomicName id="94C24D25FFFDFFE3E1FD01A5FDE8FDDA" ID-CoL="3ZK6V" authority="Seitz, 1909" authorityName="Seitz" authorityYear="1909" box="[199,587,510,537]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="30" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="changaica">
<emphasis id="61B6EAB4FFFDFFE3E1FD01A5FE60FDDA" bold="true" box="[199,451,510,537]" italics="true" pageId="5" pageNumber="30">M. phoebe changaica</emphasis>
Seitz, 1909
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="1BD8652DFFFDFFE3E36E01A5FBE6FC9E" pageId="5" pageNumber="30" type="discussion">
<paragraph id="537D36A6FFFDFFE3E36E01A5FBE6FC9E" blockId="5.[151,1437,150,862]" pageId="5" pageNumber="30">
[TL:
<collectingCountry id="2BD57636FFFDFFE3E3AA01A5FCA3FDDA" box="[656,768,511,537]" name="Mongolia" pageId="5" pageNumber="30">Mongolia</collectingCountry>
, Changai (Khangai or Hangayn) Mountains] uppersides of a male and female were figured by Seitz (plate 65, figs h 4 &amp; 5).However, www.funet.fi (accessed
<date id="277C1066FFFDFFE3E42F0279FA36FDFD" box="[1301,1429,547,574]" pageId="5" pageNumber="30" value="2020-10-26">26.10.2020</date>
) provides a photograph (© Kosterin) of undersides of a male and female of this this subspecies in
<emphasis id="61B6EAB4FFFDFFE3E435021DFAF8FDA2" box="[1295,1371,583,609]" italics="true" pageId="5" pageNumber="30">copula</emphasis>
from Chikhalan, Chita Province, S. Transbaikalia, Siberia,
<collectingCountry id="2BD57636FFFDFFE3E3CE0231FC6FFD46" box="[756,972,619,645]" name="Russia" pageId="5" pageNumber="30">Russian Federation</collectingCountry>
(
<date id="277C1066FFFDFFE3E2E70231FBD7FD45" box="[989,1140,619,646]" pageId="5" pageNumber="30" value="1996-07-12">12 July 1996</date>
); from the photograph the characteristics typical of
<taxonomicName id="94C24D25FFFDFFE3E08C02D5FD8CFD6A" box="[438,559,655,681]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="30" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFDFFE3E08C02D5FD8CFD6A" box="[438,559,655,681]" italics="true" pageId="5" pageNumber="30">M. phoebe</emphasis>
</taxonomicName>
can be observed, in particular the black submarginal markings clearly touch the intervening veins on both wings of the
<specimenCount id="45C4FD2FFFFDFFE3E34F02E9FCBEFD0E" box="[629,797,691,717]" pageId="5" pageNumber="30" type="generic">two specimens</specimenCount>
. Also figured (© Kosterin) is a post-diapause larvae, also from the Chita region in Siberia (
<date id="277C1066FFFDFFE3E330028DFD01FD32" box="[522,674,726,753]" pageId="5" pageNumber="30" value="1995-06-19">19 June 1995</date>
), with a black head carapace typical of
<taxonomicName id="94C24D25FFFDFFE3E560028DFB73FD32" box="[1114,1232,727,753]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="30" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFDFFE3E560028DFB73FD32" box="[1114,1232,727,753]" italics="true" pageId="5" pageNumber="30">M. phoebe</emphasis>
</taxonomicName>
. A specimen from Changai in the Staudinger collection (
<figureCitation id="CBF92A23FFFDFFE3E37C02A1FD3BFCD6" box="[582,664,763,789]" captionStart="FIGURE 12" captionStartId="5.[151,250,1868,1892]" captionTargetBox="[151,1436,885,1845]" captionTargetId="figure-371@5.[151,1436,885,1845]" captionTargetPageId="5" captionText="FIGURE 12. Melitaea phoebe changaica Seitz, 1909, male underside. © T. Léger." figureDoi="http://doi.org/10.5281/zenodo.6577642" httpUri="https://zenodo.org/record/6577642/files/figure.png" pageId="5" pageNumber="30">Fig. 12</figureCitation>
) clearly shows the club-shaped antennae and submarginal markings on the underside typical of
<taxonomicName id="94C24D25FFFDFFE3E0FD0345FD9EFCFA" box="[455,573,799,825]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="30" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFDFFE3E0FD0345FD9EFCFA" box="[455,573,799,825]" italics="true" pageId="5" pageNumber="30">M. phoebe</emphasis>
</taxonomicName>
. This taxon is therefore considered to be a subspecies of
<taxonomicName id="94C24D25FFFDFFE3E5820345FA8DFCFA" box="[1208,1326,799,825]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="30" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFDFFE3E5820345FA8DFCFA" box="[1208,1326,799,825]" italics="true" pageId="5" pageNumber="30">M. phoebe</emphasis>
</taxonomicName>
, agreeing with
<bibRefCitation id="37534B57FFFDFFE3E1EA0319FDF9FC9D" author="Van Oorschot, H. &amp; Coutsis, J. G." box="[208,602,835,862]" pageId="5" pageNumber="30" refId="ref11693" refString="Van Oorschot, H. &amp; Coutsis, J. G. (2014) The genus Melitaea Fabricius, 1807 (Lepidoptera, Nymphalidae, Nymphalinae) Taxonomy and systematics with special reference to the male genitalia. Tshikolovets publications, Pardubice, Czech Republic, 360 pp." type="book" year="2014">van Oorschot &amp; Coutsis (2014: 60)</bibRefCitation>
and
<bibRefCitation id="37534B57FFFDFFE3E3AB0319FC76FC9E" author="Russell, PJ &amp; Tennent, W. J." box="[657,981,834,861]" pageId="5" pageNumber="30" pagination="27 - 56" refId="ref10922" refString="Russell, PJ. C. &amp; Tennent, W. J. (2016) A synonymic list of names associated with western Palaearctic Melitaea phoebe (Denis &amp; Schiffermuller, 1775) species group taxa M. phoebe; M. punica Oberthur, 1876; M. ornata Christoph, 1893) (Lepidoptera, Nymphalidae). Nota Lepidopterologica, 39 (1), 27 - 56. https: // doi. org / 10.3897 / nl. 39.5929" type="journal article" year="2016">Russell &amp; Tennent (2016: 45</bibRefCitation>
, note 24).
</paragraph>
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<taxonomicName id="94C24D25FFFEFFE0E1FD0111FD33FEA6" ID-CoL="5JM9D" authority="Wagner, 1913" authorityName="Wagner" authorityYear="1913" box="[199,656,330,357]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="variety" species="phoebe" variety="alatauica">
<emphasis id="61B6EAB4FFFEFFE0E1FD0111FE49FEA6" bold="true" box="[199,490,330,357]" italics="true" pageId="6" pageNumber="31">M. phoebe var. alatauica</emphasis>
<bibRefCitation id="37534B57FFFEFFE0E0C90111FD33FEA6" author="Wagner, F." box="[499,656,331,357]" pageId="6" pageNumber="31" pagination="22 - 288" refId="ref11971" refString="Wagner, F. (1913) Beitrag zur Lepidopterenfauna des Iligebietes sowie des Sary-Dschas (Asia centr.) Entomologische Mitteilungen Berlin, 2 (5), 22 - 288." type="journal article" year="1913">Wagner, 1913</bibRefCitation>
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</subSubSection>
<subSubSection id="1BD8652DFFFEFFE0E3A30111FE7AFDFE" pageId="6" pageNumber="31" type="discussion">
<paragraph id="537D36A6FFFEFFE0E3A30111FE7AFDFE" blockId="6.[151,1436,151,573]" pageId="6" pageNumber="31">
[TL:
<collectingCountry id="2BD57636FFFEFFE0E3EF0111FCF8FEA6" box="[725,859,331,357]" name="Kazakhstan" pageId="6" pageNumber="31">Kazakhstan</collectingCountry>
, Taldi Kurgan and mountains near Issyk-Kul Lake and Ili River], name preoccupied by
<taxonomicName id="94C24D25FFFEFFE0E3160135FC52FE4A" authority="Staudinger, 1881" authorityName="Staudinger" authorityYear="1881" box="[556,1009,367,393]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="parthenie" subSpecies="alatauica">
<emphasis id="61B6EAB4FFFEFFE0E3160135FC8FFE4A" box="[556,812,367,393]" italics="true" pageId="6" pageNumber="31">M. parthenie alatauica</emphasis>
<bibRefCitation id="37534B57FFFEFFE0E20B0135FC52FE4A" author="Staudinger, O." box="[817,1009,367,393]" pageId="6" pageNumber="31" pagination="193 - 215" refId="ref11194" refString="Staudinger, O. (1881) Beitrag zur Lepidopterenfauna Central-Asiens. Stettin Entomologischen Zeitung, 42 (7 - 9), 193 - 215 + 253 - 300." type="journal article" year="1881">Staudinger, 1881</bibRefCitation>
</taxonomicName>
. The name was replaced by
<taxonomicName id="94C24D25FFFEFFE0E41D0135FE68FE6E" authority="Wnukowsky, 1929" authorityName="Wnukowsky" authorityYear="1929" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="wagneri">
<emphasis id="61B6EAB4FFFEFFE0E41D0135FF50FE6F" italics="true" pageId="6" pageNumber="31">M. phoebe wagneri</emphasis>
<bibRefCitation id="37534B57FFFEFFE0E1C201C9FE68FE6E" author="Wnukowsky, W." box="[248,459,403,429]" pageId="6" pageNumber="31" pagination="221 - 224" refId="ref12039" refString="Wnukowsky, W. (1929) Einige faunistische Angaben uber die Insekten Siberiens und des Ussuri-Gebietes. Zoologischer Anzeiger, 83 (9 / 10), 221 - 224." type="journal article" year="1929">Wnukowsky, 1929</bibRefCitation>
</taxonomicName>
. Male specimens of this
<taxonomicName id="94C24D25FFFEFFE0E3E001C9FC2AFE6E" box="[730,905,403,429]" pageId="6" pageNumber="31" rank="variety" variety="together">variety together</taxonomicName>
with typical
<taxonomicName id="94C24D25FFFEFFE0E52201C9FB2EFE6E" box="[1048,1165,403,429]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFEFFE0E52201C9FB2EFE6E" box="[1048,1165,403,429]" italics="true" pageId="6" pageNumber="31">M. phoebe</emphasis>
</taxonomicName>
were figured by
<bibRefCitation id="37534B57FFFEFFE0E47C01C9FEA6FE12" author="Wagner, F." pageId="6" pageNumber="31" pagination="22 - 288" refId="ref11971" refString="Wagner, F. (1913) Beitrag zur Lepidopterenfauna des Iligebietes sowie des Sary-Dschas (Asia centr.) Entomologische Mitteilungen Berlin, 2 (5), 22 - 288." type="journal article" year="1913">Wagner (1913: 89</bibRefCitation>
, figs 5 &amp; 6), exemplifying the heavy post discal black spotting, differentiating it from typical
<taxonomicName id="94C24D25FFFEFFE0E42501EDFA36FE12" box="[1311,1429,439,465]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFEFFE0E42501EDFA36FE12" box="[1311,1429,439,465]" italics="true" pageId="6" pageNumber="31">M. phoebe</emphasis>
</taxonomicName>
. This variety is included here because
<bibRefCitation id="37534B57FFFEFFE0E3000181FCBFFE36" author="Higgins, L. G." box="[570,796,475,501]" pageId="6" pageNumber="31" pagination="175 - 365" refId="ref9372" refString="Higgins, L. G. (1941) An illustrated catalogue of the Palearctic Melitaea (Lep. Rhopalocera). Transactions of the Royal Entomological Society of London, 91, 175 - 365. https: // doi. org / 10.1111 / j. 1365 - 2311.1941. tb 01045. x" type="journal article" year="1941">Higgins (1941: 338)</bibRefCitation>
suggested it might be referable to
<taxonomicName id="94C24D25FFFEFFE0E5A70181FAA5FE36" authorityName="Alpheraky" authorityYear="1881" box="[1181,1286,475,501]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="sibina">
<emphasis id="61B6EAB4FFFEFFE0E5A70181FAA5FE36" box="[1181,1286,475,501]" italics="true" pageId="6" pageNumber="31">M. sibina</emphasis>
</taxonomicName>
. It was noted above that
<bibRefCitation id="37534B57FFFEFFE0E02801A5FE5FFDDA" author="Toth, J. P. &amp; Varga, Z." box="[274,508,511,537]" pageId="6" pageNumber="31" pagination="258 - 268" refId="ref11248" refString="Toth, J. P. &amp; Varga, Z. (2011) Inter- and intraspecific variation in the genitalia of the ' Melitaea phoebe group' (Lepidoptera. Nymphalidae). Zoologischer Anzeiger, 250, 258 - 268. [6 pp. of Appendix with key and figures of genitalia only present in the on-line version, www. sciencedirect. com] https: // doi. org / 10.1016 / j. jcz. 2011.05.002" type="book chapter" year="2011">Tóth &amp; Varga (2011)</bibRefCitation>
were unable to separate these two species; however, the present authors accept the taxon
<taxonomicName id="94C24D25FFFEFFE0E1E60279FE83FDFE" authorityName="Alpheraky" authorityYear="1881" box="[220,288,547,573]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="31" phylum="Arthropoda" rank="species" species="sibina">
<emphasis id="61B6EAB4FFFEFFE0E1E60279FE83FDFE" box="[220,288,547,573]" italics="true" pageId="6" pageNumber="31">sibina</emphasis>
</taxonomicName>
at species level.
</paragraph>
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</treatment>
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<treatment id="DB6B87B0FFFFFFE1E1FD00CDFAB7FD0E" ID-DOI="http://doi.org/10.5281/zenodo.6580840" ID-GBIF-Taxon="195626709" ID-Zenodo-Dep="6580840" LSID="urn:lsid:plazi:treatment:DB6B87B0FFFFFFE1E1FD00CDFAB7FD0E" httpUri="http://treatment.plazi.org/id/DB6B87B0FFFFFFE1E1FD00CDFAB7FD0E" lastPageNumber="32" pageId="7" pageNumber="32">
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<paragraph id="537D36A6FFFFFFE1E1FD00CDFDE2FF72" blockId="7.[151,1437,150,2013]" box="[199,577,150,177]" pageId="7" pageNumber="32">
<taxonomicName id="94C24D25FFFFFFE1E1FD00CDFDE2FF72" ID-CoL="93YD4" authority="Rebel, 1917" authorityName=", Rebel" authorityYear="1917" box="[199,577,150,177]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="amanica">
<emphasis id="61B6EAB4FFFFFFE1E1FD00CDFE0CFF72" bold="true" box="[199,431,150,177]" italics="true" pageId="7" pageNumber="32">M. phoebe amanica</emphasis>
<bibRefCitation id="37534B57FFFFFFE1E08200CDFDE2FF72" author="Rebel, H." box="[440,577,151,177]" pageId="7" pageNumber="32" pagination="243 - 282" refId="ref10458" refString="Rebel, H. (1917) Eine Lepidopterenausbeute aus dem Amanusgebirge (Alman Dagh). Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften Mathematisch-Naturwissenschaftliche Wien, 126, 243 - 282." type="journal article" year="1917">Rebel, 1917</bibRefCitation>
</taxonomicName>
</paragraph>
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<subSubSection id="1BD8652DFFFFFFE1E37000CDFAB7FD0E" pageId="7" pageNumber="32" type="discussion">
<paragraph id="537D36A6FFFFFFE1E37000CDFAB7FD0E" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="32">
[TL: Kushdjula, Taurus Mountains; Das Dagh, Amanus Mountains,
<collectingCountry id="2BD57636FFFFFFE1E46E00CDFA3BFF72" box="[1364,1432,151,177]" name="Turkey" pageId="7" pageNumber="32">Hatay</collectingCountry>
,
<collectingCountry id="2BD57636FFFFFFE1E1AD00E0FF4EFF16" box="[151,237,186,213]" name="Turkey" pageId="7" pageNumber="32">Turkey</collectingCountry>
]; syntypic material consists of a single male and
<specimenCount id="45C4FD2FFFFFFFE1E22300E1FC01FF16" box="[793,930,187,213]" pageId="7" pageNumber="32" type="female">two females</specimenCount>
in the Naturhistorisches Museum in Vienna, each having a red ‘type’ label.
<bibRefCitation id="37534B57FFFFFFE1E0D90085FCA6FF3A" author="Hesselbarth, G. &amp; van Oorschot, H. &amp; Wagener, S." box="[483,773,222,249]" pageId="7" pageNumber="32" pagination="1 - 847" refId="ref9287" refString="Hesselbarth, G., van Oorschot, H. &amp; Wagener, S. (1995) Die Tagfalter der Turkei unter Berucksichtigung der angrenzenden Lander, Sigbert Wagener, Bocholt, Germany, 1, 1 - 757, 2, 758 - 1354, 3, 1 - 847." type="journal article" year="1995">
Hesselbarth
<emphasis id="61B6EAB4FFFFFFE1E3560085FD00FF3A" box="[620,675,223,249]" italics="true" pageId="7" pageNumber="32">et al.</emphasis>
(1995: 2
</bibRefCitation>
, 1031) and
<bibRefCitation id="37534B57FFFFFFE1E2B90085FAA0FF39" author="Van Oorschot, H. &amp; Coutsis, J. G." box="[899,1283,223,250]" pageId="7" pageNumber="32" refId="ref11693" refString="Van Oorschot, H. &amp; Coutsis, J. G. (2014) The genus Melitaea Fabricius, 1807 (Lepidoptera, Nymphalidae, Nymphalinae) Taxonomy and systematics with special reference to the male genitalia. Tshikolovets publications, Pardubice, Czech Republic, 360 pp." type="book" year="2014">van Oorschot &amp; Coutsis (2014: 63)</bibRefCitation>
synonymised this subspecies with
<emphasis id="61B6EAB4FFFFFFE1E0410159FD2DFEDE" box="[379,654,259,285]" italics="true" pageId="7" pageNumber="32">
<taxonomicName id="94C24D25FFFFFFE1E0410159FDE1FEDF" authorityName="Oberthur" authorityYear="1876" box="[379,578,259,285]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="punica">M. phoebe punica</taxonomicName>
<taxonomicName id="94C24D25FFFFFFE1E3720159FD2DFEDE" authorityName="Fruhstorfer" authorityYear="1908" box="[584,654,259,285]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="telona">telona</taxonomicName>
</emphasis>
and
<taxonomicName id="94C24D25FFFFFFE1E3F80159FC8EFEDE" authorityName="Fruhstorfer" authorityYear="1908" box="[706,813,259,285]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="telona">
<emphasis id="61B6EAB4FFFFFFE1E3F80159FC8EFEDE" box="[706,813,259,285]" italics="true" pageId="7" pageNumber="32">M. telona</emphasis>
</taxonomicName>
, respectively; the present authors certainly consider that this subspecies belongs to either
<taxonomicName id="94C24D25FFFFFFE1E33E017DFDD7FE83" box="[516,628,295,320]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="61B6EAB4FFFFFFE1E33E017DFDD7FE83" box="[516,628,295,320]" italics="true" pageId="7" pageNumber="32">M. ornata</emphasis>
</taxonomicName>
or
<taxonomicName id="94C24D25FFFFFFE1E3A2017DFCA0FE82" authorityName="Fruhstorfer" authorityYear="1908" box="[664,771,295,321]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="telona">
<emphasis id="61B6EAB4FFFFFFE1E3A2017DFCA0FE82" box="[664,771,295,321]" italics="true" pageId="7" pageNumber="32">M. telona</emphasis>
</taxonomicName>
rather than
<taxonomicName id="94C24D25FFFFFFE1E2B2017DFC5DFE82" box="[904,1022,295,321]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFFFFE1E2B2017DFC5DFE82" box="[904,1022,295,321]" italics="true" pageId="7" pageNumber="32">M. phoebe</emphasis>
</taxonomicName>
.
<bibRefCitation id="37534B57FFFFFFE1E530017DFB6CFE82" author="Toth, J. P. &amp; Bereczki, J. &amp; Varga, Z. &amp; Rota, J. &amp; Sramko, G. &amp; Wahlberg, N." box="[1034,1231,295,321]" pageId="7" pageNumber="32" pagination="749 - 757" refId="ref11329" refString="Toth, J. P., Bereczki, J., Varga, Z., Rota, J., Sramko, G. &amp; Wahlberg, N. (2014) Relationships within the Melitaea phoebe species group (Lepidoptera: Nymphalidae): new insights from molecular and morphometric information. Systematic Entomology, 39, 749 - 757. https: // doi. org / 10.1111 / syen. 12083" type="journal article" year="2014">
Tóth
<emphasis id="61B6EAB4FFFFFFE1E57C017DFBDDFE82" box="[1094,1150,295,321]" italics="true" pageId="7" pageNumber="32">et al.</emphasis>
(2014)
</bibRefCitation>
demonstrated that
<taxonomicName id="94C24D25FFFFFFE1E1AD0111FEA7FEA6" authorityName="Fruhstorfer" authorityYear="1908" box="[151,260,331,357]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="telona">
<emphasis id="61B6EAB4FFFFFFE1E1AD0111FEA7FEA6" box="[151,260,331,357]" italics="true" pageId="7" pageNumber="32">M. telona</emphasis>
</taxonomicName>
was present in
<collectingCountry id="2BD57636FFFFFFE1E08D0111FDBFFEA6" box="[439,540,331,357]" name="Lebanon" pageId="7" pageNumber="32">Lebanon</collectingCountry>
, to the South of the Nur mountain range and
<bibRefCitation id="37534B57FFFFFFE1E5180111FAEDFEA6" author="Russell, P. J. C. &amp; Pateman, J. E." box="[1058,1358,331,357]" pageId="7" pageNumber="32" pagination="85 - 94" refId="ref10812" refString="Russell, P. J. C. &amp; Pateman, J. E. (2012) Some observations on populations of Melitaea telona Fruhstorfer, 1908 (Lepidoptera: Nymphalidae) in Turkey. Entomologist's Gazette, 63, 85 - 94." type="journal article" year="2012">Russell &amp; Pateman (2012)</bibRefCitation>
reared an egg batch from a female captured at Tuzlabeli geçidi, Muðla, western
<collectingCountry id="2BD57636FFFFFFE1E2E10135FB88FE4A" box="[987,1067,367,393]" name="Turkey" pageId="7" pageNumber="32">Turkey</collectingCountry>
and demonstrated from the L4+ larval head colour that the population there was not
<taxonomicName id="94C24D25FFFFFFE1E3D701C9FCC6FE6E" box="[749,869,403,429]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFFFFE1E3D701C9FCC6FE6E" box="[749,869,403,429]" italics="true" pageId="7" pageNumber="32">M. phoebe</emphasis>
</taxonomicName>
but
<taxonomicName id="94C24D25FFFFFFE1E2A001C9FBAFFE6F" box="[922,1036,403,428]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="61B6EAB4FFFFFFE1E2A001C9FBAFFE6F" box="[922,1036,403,428]" italics="true" pageId="7" pageNumber="32">M. ornata</emphasis>
</taxonomicName>
. The first author considers that the specimens should probably be associated with
<taxonomicName id="94C24D25FFFFFFE1E39D01EDFCB0FE12" authorityName="Fruhstorfer" authorityYear="1908" box="[679,787,439,465]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="telona">
<emphasis id="61B6EAB4FFFFFFE1E39D01EDFCB0FE12" box="[679,787,439,465]" italics="true" pageId="7" pageNumber="32">M. telona</emphasis>
</taxonomicName>
but further DNA analysis is required to confirm to which species they belong. In order that this subspecific name can be in future firmly associated with a species we hereby formally designate the male specimen as
<typeStatus id="8C798804FFFFFFE1E35901A5FD69FDDA" box="[611,714,511,537]" pageId="7" pageNumber="32" type="lectotype">lectotype</typeStatus>
for
<taxonomicName id="94C24D25FFFFFFE1E3C201A5FBC6FDDA" authority="Rebel, 1917" authorityName=", Rebel" authorityYear="1917" box="[760,1125,511,537]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="amanica">
<emphasis id="61B6EAB4FFFFFFE1E3C201A5FC77FDDB" box="[760,980,511,537]" italics="true" pageId="7" pageNumber="32">M. phoebe amanica</emphasis>
<bibRefCitation id="37534B57FFFFFFE1E2E101A5FBC6FDDA" author="Rebel, H." box="[987,1125,511,537]" pageId="7" pageNumber="32" pagination="243 - 282" refId="ref10458" refString="Rebel, H. (1917) Eine Lepidopterenausbeute aus dem Amanusgebirge (Alman Dagh). Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften Mathematisch-Naturwissenschaftliche Wien, 126, 243 - 282." type="journal article" year="1917">Rebel, 1917</bibRefCitation>
</taxonomicName>
. From figures 13A, B it can be seen that this specimen has the wing morphological characters typical of both
<taxonomicName id="94C24D25FFFFFFE1E5000279FB0FFDFF" box="[1082,1196,547,572]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="61B6EAB4FFFFFFE1E5000279FB0FFDFF" box="[1082,1196,547,572]" italics="true" pageId="7" pageNumber="32">M. ornata</emphasis>
</taxonomicName>
and
<taxonomicName id="94C24D25FFFFFFE1E5DF0279FAF1FDFE" authorityName="Fruhstorfer" authorityYear="1908" box="[1253,1362,547,573]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="telona">
<emphasis id="61B6EAB4FFFFFFE1E5DF0279FAF1FDFE" box="[1253,1362,547,573]" italics="true" pageId="7" pageNumber="32">M. telona</emphasis>
</taxonomicName>
rather than those of
<taxonomicName id="94C24D25FFFFFFE1E00A021DFE05FDA2" box="[304,422,583,609]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFFFFE1E00A021DFE05FDA2" box="[304,422,583,609]" italics="true" pageId="7" pageNumber="32">M. phoebe</emphasis>
</taxonomicName>
. The labels on the specimen pin (
<figureCitation id="CBF92A23FFFFFFE1E21B021DFC26FDA2" box="[801,901,583,609]" captionStart="FIGURES 13–16" captionStartId="6.[151,264,1911,1935]" captionTargetBox="[151,1436,623,1883]" captionTargetId="figure-232@6.[151,1436,597,1888]" captionTargetPageId="6" captionText="FIGURES 13–16. 13, Lectotype of Melitaea phoebe amanica, Rebel, 1917. 14, Lectotype male of Melitaea phoebe dorae Graves, 1925. 15, Paralectotype male of Melitaea phoebe dorae Graves, 1925. 16, Holotype of Melitaea phoebe enoch Higgins, 1941. Fig. 13 © H. Bruckner. Figs 14–16 © J. Tennent.A = upperside, B = underside, C = labels on specimen pin." figureDoi="http://doi.org/10.5281/zenodo.6577644" httpUri="https://zenodo.org/record/6577644/files/figure.png" pageId="7" pageNumber="32">Fig. 13C</figureCitation>
) are as follows: (1) on white paper handwritten in black “Taurus/ Kushdjula/ 22.V.14”; (2) on white paper handwritten in black “
<emphasis id="61B6EAB4FFFFFFE1E50F0231FB54FD47" box="[1077,1271,619,645]" italics="true" pageId="7" pageNumber="32">
<taxonomicName id="94C24D25FFFFFFE1E50F0231FB22FD46" box="[1077,1153,619,645]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="phoebe">phoebe</taxonomicName>
/ amanica
</emphasis>
/ Type [in red] Rbl”; (3) on white card typed in black “Misident:/
<taxonomicName id="94C24D25FFFFFFE1E3E902D5FB8AFD6A" authority=", Russell 2021" authorityName=", Russell" authorityYear="2021" box="[723,1065,655,681]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="telona">
<emphasis id="61B6EAB4FFFFFFE1E3E902D5FC27FD6A" box="[723,900,655,681]" italics="true" pageId="7" pageNumber="32">Melitaea telona</emphasis>
, Russell 2021
</taxonomicName>
”; (4) on red paper typed in black “
<typeStatus id="8C798804FFFFFFE1E19A02E9FEE1FD0E" box="[160,322,691,717]" pageId="7" pageNumber="32" type="lectotype">LECTOTYPE</typeStatus>
/
<taxonomicName id="94C24D25FFFFFFE1E06802E9FDD7FD0F" authorityName=", Rebel" authorityYear="1917" box="[338,628,691,717]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="amanica">
<emphasis id="61B6EAB4FFFFFFE1E06802E9FDD7FD0F" box="[338,628,691,717]" italics="true" pageId="7" pageNumber="32">Melitaea phoebe amanica</emphasis>
</taxonomicName>
/
<bibRefCitation id="37534B57FFFFFFE1E3BE02E9FC82FD0E" author="Rebel, H." box="[644,801,691,717]" pageId="7" pageNumber="32" pagination="243 - 282" refId="ref10458" refString="Rebel, H. (1917) Eine Lepidopterenausbeute aus dem Amanusgebirge (Alman Dagh). Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften Mathematisch-Naturwissenschaftliche Wien, 126, 243 - 282." type="journal article" year="1917">REBEL, 1917</bibRefCitation>
/ designated Russell &amp;/ Gaal-Haszler 2021”.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="192E5213BCF63D4974A493415FEF313C">Reassessment of the status of some European and Asian Melitaea taxa described as subspecies of Melitaea phoebe ([Denis &amp; Schiffermüller], 1775), with designations of lectotypes where appropriate (Lepidoptera: Nymphalidae)</mods:title>
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<mods:namePart id="CB7F189D6592CF6B264C5652A5589E9B">Lukhtanov, Vladimir A.</mods:namePart>
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<subSubSection id="1BD8652DFFFFFFE1E1FD028DFD8FFD32" box="[199,556,726,753]" pageId="7" pageNumber="32" type="nomenclature">
<paragraph id="537D36A6FFFFFFE1E1FD028DFD8FFD32" blockId="7.[151,1437,150,2013]" box="[199,556,726,753]" pageId="7" pageNumber="32">
<taxonomicName id="94C24D25FFFFFFE1E1FD028DFD8FFD32" ID-CoL="5JM9P" authority="Graves, 1925" authorityName="Graves" authorityYear="1925" box="[199,556,726,753]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="dorae">
<emphasis id="61B6EAB4FFFFFFE1E1FD028DFE2FFD32" bold="true" box="[199,396,726,753]" italics="true" pageId="7" pageNumber="32">M. phoebe dorae</emphasis>
<bibRefCitation id="37534B57FFFFFFE1E0AF028DFD8FFD32" author="Graves, P." box="[405,556,726,753]" pageId="7" pageNumber="32" pagination="17 - 126" refId="ref9143" refString="Graves, P. (1925) The Rhopalocera and Grypocera of Palestine and Transjordania. Transactions of the entomological Society of London, 73, 17 - 126, plates 4, 5." type="journal article" year="1925">Graves, 1925</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="1BD8652DFFFFFFE1E30E028DFCBAFB2A" pageId="7" pageNumber="32" type="discussion">
<paragraph id="537D36A6FFFFFFE1E30E028DFB50FBD2" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="32">
[TL: Petra, Nabatea,
<collectingCountry id="2BD57636FFFFFFE1E21F028CFCD5FD32" box="[805,886,726,753]" name="Jordan" pageId="7" pageNumber="32">Jordan</collectingCountry>
]. Graves gave a long and detailed description of
<emphasis id="61B6EAB4FFFFFFE1E1AD02A1FF7BFCD6" box="[151,216,763,789]" italics="true" pageId="7" pageNumber="32">dorae</emphasis>
and a related discussion, including details of localities of where the type series was collected. This might have been rather confusing— and appears to have been so to whoever labelled the specimens subsequently since a series of
<specimenCount id="45C4FD2FFFFFFFE1E1C60319FE90FC9E" box="[252,307,835,861]" pageId="7" pageNumber="32" type="male">13 ♂</specimenCount>
and
<specimenCount id="45C4FD2FFFFFFFE1E0530319FE67FC9E" box="[361,452,835,861]" pageId="7" pageNumber="32" type="female">seven ♀</specimenCount>
‘
<emphasis id="61B6EAB4FFFFFFE1E0EE0319FDB6FC9E" box="[468,533,835,861]" italics="true" pageId="7" pageNumber="32">dorae</emphasis>
’ in the Natural History Museum London are labelled as ‘cotypes’ (
<emphasis id="61B6EAB4FFFFFFE1E43B0319FABEFC9F" box="[1281,1309,835,860]" italics="true" pageId="7" pageNumber="32">i.e</emphasis>
.
<typeStatus id="8C798804FFFFFFE1E4100319FA32FC9E" box="[1322,1425,835,861]" pageId="7" pageNumber="32" type="syntype">syntypes</typeStatus>
). However, Graves very clearly said (
<bibRefCitation id="37534B57FFFFFFE1E30F033DFD5CFC42" author="Graves, P." box="[565,767,870,897]" pageId="7" pageNumber="32" pagination="17 - 126" refId="ref9143" refString="Graves, P. (1925) The Rhopalocera and Grypocera of Palestine and Transjordania. Transactions of the entomological Society of London, 73, 17 - 126, plates 4, 5." type="journal article" year="1925">Graves 1925:104</bibRefCitation>
): “Described from
<specimenCount id="45C4FD2FFFFFFFE1E2E1033DFBB5FC42" box="[987,1046,871,897]" pageId="7" pageNumber="32" type="male">13 ♂</specimenCount>
and
<specimenCount id="45C4FD2FFFFFFFE1E56B033DFBDDFC42" box="[1105,1150,871,897]" pageId="7" pageNumber="32" type="female">3 ♀</specimenCount>
, taken in the last half of
<date id="277C1066FFFFFFE1E1AD03D1FEBCFC66" box="[151,287,907,933]" pageId="7" pageNumber="32" value="1922-03">March 1922</date>
and 1923 at Petra in Nabataea (
<quantity id="943A9B43FFFFFFE1E3BF03D1FD46FC66" box="[645,741,907,933]" metricMagnitude="-1" metricUnit="m" metricValue="9.4488" pageId="7" pageNumber="32" unit="ft" value="3.1">3,100 ft.</quantity>
approx. [
<emphasis id="61B6EAB4FFFFFFE1E26B03D1FCC8FC67" box="[849,875,907,932]" italics="true" pageId="7" pageNumber="32">ca</emphasis>
<quantity id="943A9B43FFFFFFE1E24803D0FC1FFC66" box="[882,956,906,933]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="7" pageNumber="32" unit="m" value="950.0">950 m</quantity>
]) by Mr H. St. J. Philby, and now in the B. M.”. This is emphasised in an “index of nigrescence of
<taxonomicName id="94C24D25FFFFFFE1E23603F5FC20FC0A" box="[780,899,943,969]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFFFFE1E23603F5FC20FC0A" box="[780,899,943,969]" italics="true" pageId="7" pageNumber="32">M. phoebe</emphasis>
</taxonomicName>
races” (
<bibRefCitation id="37534B57FFFFFFE1E2DA03F5FB08FC0A" author="Graves, P." box="[992,1195,942,969]" pageId="7" pageNumber="32" pagination="17 - 126" refId="ref9143" refString="Graves, P. (1925) The Rhopalocera and Grypocera of Palestine and Transjordania. Transactions of the entomological Society of London, 73, 17 - 126, plates 4, 5." type="journal article" year="1925">Graves 1925: 105</bibRefCitation>
) where 16 individual
<emphasis id="61B6EAB4FFFFFFE1E1AD0389FF7BFC2E" box="[151,216,979,1005]" italics="true" pageId="7" pageNumber="32">dorae</emphasis>
specimens are listed numerically, all from Petra. Some of the NHM specimens labelled as cotypes are from, for example, the Bir Arub Road in
<collectingCountry id="2BD57636FFFFFFE1E32703ADFD18FBD2" box="[541,699,1015,1041]" name="Jordan" pageId="7" pageNumber="32">Transjordania</collectingCountry>
, and are specifically excluded (
<bibRefCitation id="37534B57FFFFFFE1E52003ADFB44FBD2" author="Graves, P." box="[1050,1255,1014,1041]" pageId="7" pageNumber="32" pagination="17 - 126" refId="ref9143" refString="Graves, P. (1925) The Rhopalocera and Grypocera of Palestine and Transjordania. Transactions of the entomological Society of London, 73, 17 - 126, plates 4, 5." type="journal article" year="1925">Graves 1925: 104</bibRefCitation>
).
</paragraph>
<paragraph id="537D36A6FFFFFFE1E1FD0441FCBAFB2A" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="32">
Described as a race of
<taxonomicName id="94C24D25FFFFFFE1E08D0441FD88FBF6" box="[439,555,1051,1077]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="phoebe">
<emphasis id="61B6EAB4FFFFFFE1E08D0441FD88FBF6" box="[439,555,1051,1077]" italics="true" pageId="7" pageNumber="32">M. phoebe</emphasis>
</taxonomicName>
, the name
<emphasis id="61B6EAB4FFFFFFE1E3A70441FD7DFBF6" box="[669,734,1051,1077]" italics="true" pageId="7" pageNumber="32">dorae</emphasis>
clearly refers to
<taxonomicName id="94C24D25FFFFFFE1E2AA0441FC5AFBF6" authorityName=", Russell" authorityYear="2021" box="[912,1017,1051,1077]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="telona">
<emphasis id="61B6EAB4FFFFFFE1E2AA0441FC5AFBF6" box="[912,1017,1051,1077]" italics="true" pageId="7" pageNumber="32">M. telona</emphasis>
</taxonomicName>
.We hereby designate a male
<typeStatus id="8C798804FFFFFFE1E40C0441FA3EFBF6" box="[1334,1437,1051,1077]" pageId="7" pageNumber="32" type="lectotype">lectotype</typeStatus>
(
<figureCitation id="CBF92A23FFFFFFE1E1A50465FEF1FB9A" box="[159,338,1087,1113]" captionStart="FIGURES 13–16" captionStartId="6.[151,264,1911,1935]" captionTargetBox="[151,1436,623,1883]" captionTargetId="figure-232@6.[151,1436,597,1888]" captionTargetPageId="6" captionText="FIGURES 13–16. 13, Lectotype of Melitaea phoebe amanica, Rebel, 1917. 14, Lectotype male of Melitaea phoebe dorae Graves, 1925. 15, Paralectotype male of Melitaea phoebe dorae Graves, 1925. 16, Holotype of Melitaea phoebe enoch Higgins, 1941. Fig. 13 © H. Bruckner. Figs 14–16 © J. Tennent.A = upperside, B = underside, C = labels on specimen pin." figureDoi="http://doi.org/10.5281/zenodo.6577644" httpUri="https://zenodo.org/record/6577644/files/figure.png" pageId="7" pageNumber="32">Figs 14A, B, C</figureCitation>
) of
<taxonomicName id="94C24D25FFFFFFE1E0BF0465FD2EFB9A" authorityName="Graves" authorityYear="1925" box="[389,653,1087,1113]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="dorae">
<emphasis id="61B6EAB4FFFFFFE1E0BF0465FD2EFB9A" box="[389,653,1087,1113]" italics="true" pageId="7" pageNumber="32">Melitaea phoebe dorae</emphasis>
</taxonomicName>
in the NHM, London, labelled as follows: (1) typed or handwritten “Trans
<collectingCountry id="2BD57636FFFFFFE1E1D60439FE98FBBE" box="[236,315,1123,1149]" name="Jordan" pageId="7" pageNumber="32">Jordan</collectingCountry>
: / Petra./
<date id="277C1066FFFFFFE1E0900439FD89FBBE" box="[426,554,1123,1149]" pageId="7" pageNumber="32" value="1923-03-19">19-iii-1923</date>
./ H.St.J.Philby./ Brit.Mus.1923-457”; (2) typed or handwritten “B.M. TYPE/ No. Rh 8269./
<taxonomicName id="94C24D25FFFFFFE1E00604DDFE10FB62" box="[316,435,1159,1185]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="phoebe">M. phoebe</taxonomicName>
/ dorae/
<specimenCount id="45C4FD2FFFFFFFE1E33504DDFD87FB62" box="[527,548,1159,1185]" pageId="7" pageNumber="32" type="male">♂</specimenCount>
Graves”; (3) printed, red-bordered, circular “Type”; (4) typed “
<typeStatus id="8C798804FFFFFFE1E5D004DDFA33FB62" box="[1258,1424,1159,1185]" pageId="7" pageNumber="32" type="lectotype">LECTOTYPE</typeStatus>
:/
<taxonomicName id="94C24D25FFFFFFE1E1AD04F1FE03FB06" authorityName="Graves" authorityYear="1925" box="[151,416,1195,1221]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="dorae">
<emphasis id="61B6EAB4FFFFFFE1E1AD04F1FE03FB06" box="[151,416,1195,1221]" italics="true" pageId="7" pageNumber="32">Melitaea phoebe dorae</emphasis>
</taxonomicName>
/
<bibRefCitation id="37534B57FFFFFFE1E08804F1FDEEFB06" author="Graves, P." box="[434,589,1194,1221]" pageId="7" pageNumber="32" pagination="17 - 126" refId="ref9143" refString="Graves, P. (1925) The Rhopalocera and Grypocera of Palestine and Transjordania. Transactions of the entomological Society of London, 73, 17 - 126, plates 4, 5." type="journal article" year="1925">Graves, 1925</bibRefCitation>
; designated/ by
<bibRefCitation id="37534B57FFFFFFE1E23104F1FB8BFB06" author="Russell, P. J. C. &amp; Tennent, W. J." box="[779,1064,1195,1221]" pageId="7" pageNumber="32" pagination="199 - 212" refId="ref11007" refString="Russell, P. J. C. &amp; Tennent, W. J. (2022) Butterflies observed on the western (European) slopes of the southern Ural Mountains during three spring visits in 2011, 2012 and 2014 (Lepidoptera Papilionoidea). Neue Entomologische Nachrichten 80: 199 - 212. [including 8 colour plates]" type="journal article" year="2022">Russell &amp; Tennent, 2022</bibRefCitation>
/ (=
<taxonomicName id="94C24D25FFFFFFE1E56904F1FAA4FB06" authorityName=", Russell" authorityYear="2021" box="[1107,1287,1195,1221]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="telona">
<emphasis id="61B6EAB4FFFFFFE1E56904F1FAA4FB06" box="[1107,1287,1195,1221]" italics="true" pageId="7" pageNumber="32">Melitaea telona</emphasis>
</taxonomicName>
/
<bibRefCitation id="37534B57FFFFFFE1E42204F1FF71FB2A" author="Fruhstorfer, H." pageId="7" pageNumber="32" pagination="310" refId="ref9058" refString="Fruhstorfer, H. (1908) Neue Argynnis und Melitaea. Internationale Entomologische Zeitschrift, 1, 310." type="journal article" year="1908">Fruhstorfer, 1908</bibRefCitation>
)” (5) printed purple-bordered, circular “
<typeStatus id="8C798804FFFFFFE1E3A90495FCAFFB2A" box="[659,780,1231,1257]" pageId="7" pageNumber="32" type="lectotype">Lectotype</typeStatus>
”.
</paragraph>
</subSubSection>
<subSubSection id="1BD8652DFFFFFFE1E1FD04A9FE05F81E" pageId="7" pageNumber="32" type="materials_examined">
<paragraph id="537D36A6FFFFFFE1E1FD04A9FB4DF9B6" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="32">
We recognise
<specimenCount id="45C4FD2FFFFFFFE1E05004A9FE07FACE" box="[362,420,1267,1293]" pageId="7" pageNumber="32" type="male">11 ♂</specimenCount>
(we have not been able to identify the remaining
<specimenCount id="45C4FD2FFFFFFFE1E2D004A9FC5CFACE" box="[1002,1023,1267,1293]" pageId="7" pageNumber="32" type="male">♂</specimenCount>
<typeStatus id="8C798804FFFFFFE1E53304A9FBC1FACE" box="[1033,1122,1267,1293]" pageId="7" pageNumber="32" type="syntype">syntype</typeStatus>
, presumably with the B.M. TYPE No. Rh 8279) and
<specimenCount id="45C4FD2FFFFFFFE1E087054DFDB0FAF2" box="[445,531,1303,1329]" pageId="7" pageNumber="32" type="female">three ♀</specimenCount>
<typeStatus id="8C798804FFFFFFE1E326054DFD1CFAF2" box="[540,703,1303,1329]" pageId="7" pageNumber="32" type="paralectotype">paralectotypes</typeStatus>
, all in the NHM, London, labelled as follows:
<specimenCount id="45C4FD2FFFFFFFE1E5E2054DFAA5FAF2" box="[1240,1286,1303,1329]" pageId="7" pageNumber="32" type="male">1 ♂</specimenCount>
, (1) typed or handwritten “Trans
<collectingCountry id="2BD57636FFFFFFE1E04B0561FE1DFA96" box="[369,446,1339,1365]" name="Jordan" pageId="7" pageNumber="32">Jordan</collectingCountry>
:/ Petra./
<date id="277C1066FFFFFFE1E3270561FD35FA96" box="[541,662,1339,1365]" pageId="7" pageNumber="32" value="1923-03-24">24.iii.1923</date>
./ H.St.J.Philby./ Brit. Mus. 1923-457”; (2) typed or handwritten “B.M. TYPE/ No. Rh 8271/
<taxonomicName id="94C24D25FFFFFFE1E0AD0505FDB3FABA" box="[407,528,1375,1401]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="phoebe">M. phoebe</taxonomicName>
/ dorae,/
<specimenCount id="45C4FD2FFFFFFFE1E3400505FD2CFABA" box="[634,655,1375,1401]" pageId="7" pageNumber="32" type="male">♂</specimenCount>
Graves”; (3) printed, yellow-bordered, circular “Cotype”; (4) typed “
<typeStatus id="8C798804FFFFFFE1E19A05D9FE2EFA5E" box="[160,397,1411,1437]" pageId="7" pageNumber="32" type="paralectotype">PARALECTOTYPE</typeStatus>
:/
<taxonomicName id="94C24D25FFFFFFE1E09A05D9FD00FA5E" authorityName="Graves" authorityYear="1925" box="[416,675,1411,1437]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="dorae">
<emphasis id="61B6EAB4FFFFFFE1E09A05D9FD00FA5E" box="[416,675,1411,1437]" italics="true" pageId="7" pageNumber="32">Melitaea phoebe dorae</emphasis>
</taxonomicName>
/
<bibRefCitation id="37534B57FFFFFFE1E38B05D9FCEAFA5E" author="Graves, P." box="[689,841,1410,1437]" pageId="7" pageNumber="32" pagination="17 - 126" refId="ref9143" refString="Graves, P. (1925) The Rhopalocera and Grypocera of Palestine and Transjordania. Transactions of the entomological Society of London, 73, 17 - 126, plates 4, 5." type="journal article" year="1925">Graves, 1925</bibRefCitation>
; designated/ by
<bibRefCitation id="37534B57FFFFFFE1E2C405D9FAB2FA5E" author="Russell, P. J. C. &amp; Tennent, W. J." box="[1022,1297,1411,1437]" pageId="7" pageNumber="32" pagination="199 - 212" refId="ref11007" refString="Russell, P. J. C. &amp; Tennent, W. J. (2022) Butterflies observed on the western (European) slopes of the southern Ural Mountains during three spring visits in 2011, 2012 and 2014 (Lepidoptera Papilionoidea). Neue Entomologische Nachrichten 80: 199 - 212. [including 8 colour plates]" type="journal article" year="2022">Russell &amp; Tennent, 2022</bibRefCitation>
/ (=
<taxonomicName id="94C24D25FFFFFFE1E40005D9FF7FFA02" authorityName=", Russell" authorityYear="2021" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="telona">
<emphasis id="61B6EAB4FFFFFFE1E40005D9FF7FFA02" italics="true" pageId="7" pageNumber="32">Melitaea telona</emphasis>
</taxonomicName>
/
<bibRefCitation id="37534B57FFFFFFE1E1D005FDFE0EFA02" author="Fruhstorfer, H." box="[234,429,1447,1473]" pageId="7" pageNumber="32" pagination="310" refId="ref9058" refString="Fruhstorfer, H. (1908) Neue Argynnis und Melitaea. Internationale Entomologische Zeitschrift, 1, 310." type="journal article" year="1908">Fruhstorfer, 1908</bibRefCitation>
” (5) printed blue-bordered, circular “PARA-LECTO-TYPE”;
<specimenCount id="45C4FD2FFFFFFFE1E56105FDFB0AFA02" box="[1115,1193,1447,1473]" pageId="7" pageNumber="32" type="male">1 male</specimenCount>
, ditto, but
<date id="277C1066FFFFFFE1E42705FDFA34FA02" box="[1309,1431,1447,1473]" pageId="7" pageNumber="32" value="1923-03-26">26.iii.1923</date>
; B.M. TYPE/ No. Rh 8272;
<specimenCount id="45C4FD2FFFFFFFE1E0FE0591FDB2FA26" box="[452,529,1483,1509]" pageId="7" pageNumber="32" type="male">1 male</specimenCount>
, ditto, B.M. TYPE/ No. Rh 8273;
<specimenCount id="45C4FD2FFFFFFFE1E2B20591FC76FA26" box="[904,981,1483,1509]" pageId="7" pageNumber="32" type="male">1 male</specimenCount>
, ditto, B.M. TYPE/ No. Rh 8274;
<specimenCount id="45C4FD2FFFFFFFE1E4760591FA3AFA26" box="[1356,1433,1483,1509]" pageId="7" pageNumber="32" type="male">1 male</specimenCount>
, ditto, B.M. TYPE/ No. Rh 8275, and label printed “Brit. Mus. 1922-466”;
<specimenCount id="45C4FD2FFFFFFFE1E2F605B5FBBAF9CA" box="[972,1049,1519,1545]" pageId="7" pageNumber="32" type="male">1 male</specimenCount>
, ditto, but
<date id="277C1066FFFFFFE1E5B705B5FA8DF9CA" box="[1165,1326,1519,1545]" pageId="7" pageNumber="32" value="1922-03-30" valueMax="1922-03-31" valueMin="1922-03-30">30-31-iii-1922</date>
and B.M. TYPE/ No. Rh 8276;
<specimenCount id="45C4FD2FFFFFFFE1E0B10649FE7AF9EE" box="[395,473,1555,1581]" pageId="7" pageNumber="32" type="male">1 male</specimenCount>
, ditto, B.M. TYPE/ No. Rh 8277”;
<specimenCount id="45C4FD2FFFFFFFE1E25C0649FC17F9EE" box="[870,948,1555,1581]" pageId="7" pageNumber="32" type="male">1 male</specimenCount>
, ditto, but
<date id="277C1066FFFFFFE1E5160649FB08F9EE" box="[1068,1195,1555,1581]" pageId="7" pageNumber="32" value="1923-03-28">28-iii-1923</date>
, B.M. TYPE/ No. Rh 8278;
<specimenCount id="45C4FD2FFFFFFFE1E1E5066DFE8DF992" box="[223,302,1591,1617]" pageId="7" pageNumber="32" type="male">1 male</specimenCount>
, ditto, B.M. TYPE/ No. Rh 8280;
<specimenCount id="45C4FD2FFFFFFFE1E38C066DFCA5F992" box="[694,774,1591,1617]" pageId="7" pageNumber="32" type="male">1 male</specimenCount>
, ditto, B.M. TYPE/No. Rh 8281, and label handwritten, in pencil “P35”;
<specimenCount id="45C4FD2FFFFFFFE1E00E0601FE21F9B6" box="[308,386,1627,1653]" pageId="7" pageNumber="32" type="male">1 male</specimenCount>
, ditto, but “B.M. TYPE/ No. Rh 8282”, and handwritten label, in pencil “P37”.
</paragraph>
<paragraph id="537D36A6FFFFFFE1E1FD0625FE05F81E" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="32">
<specimenCount id="45C4FD2FFFFFFFE1E1FD0625FE89F95A" box="[199,298,1663,1689]" pageId="7" pageNumber="32" type="female">1 female</specimenCount>
(
<figureCitation id="CBF92A23FFFFFFE1E0010625FE4FF95A" box="[315,492,1662,1689]" captionStart="FIGURES 13–16" captionStartId="6.[151,264,1911,1935]" captionTargetBox="[151,1436,623,1883]" captionTargetId="figure-232@6.[151,1436,597,1888]" captionTargetPageId="6" captionText="FIGURES 13–16. 13, Lectotype of Melitaea phoebe amanica, Rebel, 1917. 14, Lectotype male of Melitaea phoebe dorae Graves, 1925. 15, Paralectotype male of Melitaea phoebe dorae Graves, 1925. 16, Holotype of Melitaea phoebe enoch Higgins, 1941. Fig. 13 © H. Bruckner. Figs 14–16 © J. Tennent.A = upperside, B = underside, C = labels on specimen pin." figureDoi="http://doi.org/10.5281/zenodo.6577644" httpUri="https://zenodo.org/record/6577644/files/figure.png" pageId="7" pageNumber="32">Figs 15A, B, C</figureCitation>
), (1) typed or handwritten “Trans
<collectingCountry id="2BD57636FFFFFFE1E24C0625FC67F95A" box="[886,964,1663,1689]" name="Jordan" pageId="7" pageNumber="32">Jordan</collectingCountry>
:/ Petra./
<date id="277C1066FFFFFFE1E5100625FB09F95A" box="[1066,1194,1663,1689]" pageId="7" pageNumber="32" value="1923-03-14">14-iii-1923</date>
./ H.St.J.Philby./ Brit. Mus. 1923-457”; (2) typed or handwritten “B.M. TYPE/ No. Rh 8270/
<taxonomicName id="94C24D25FFFFFFE1E28506F9FB94F97E" box="[959,1079,1699,1725]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="phoebe">M. phoebe</taxonomicName>
/ dorae/
<specimenCount id="45C4FD2FFFFFFFE1E5AF06F9FB09F97E" box="[1173,1194,1699,1725]" pageId="7" pageNumber="32" type="female">♀</specimenCount>
Graves”; (3) printed, red-bordered, circular “Type”; (4) typed “
<typeStatus id="8C798804FFFFFFE1E35C069DFCF6F922" box="[614,853,1735,1761]" pageId="7" pageNumber="32" type="paralectotype">PARALECTOTYPE</typeStatus>
:/
<taxonomicName id="94C24D25FFFFFFE1E252069DFBC9F922" authorityName="Graves" authorityYear="1925" box="[872,1130,1735,1761]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="dorae">
<emphasis id="61B6EAB4FFFFFFE1E252069DFBC9F922" box="[872,1130,1735,1761]" italics="true" pageId="7" pageNumber="32">Melitaea phoebe dorae</emphasis>
</taxonomicName>
/
<bibRefCitation id="37534B57FFFFFFE1E543069DFAB2F922" author="Graves, P." box="[1145,1297,1734,1761]" pageId="7" pageNumber="32" pagination="17 - 126" refId="ref9143" refString="Graves, P. (1925) The Rhopalocera and Grypocera of Palestine and Transjordania. Transactions of the entomological Society of London, 73, 17 - 126, plates 4, 5." type="journal article" year="1925">Graves, 1925</bibRefCitation>
; designated/ by
<bibRefCitation id="37534B57FFFFFFE1E18706B1FE7AF8C6" author="Russell, P. J. C. &amp; Tennent, W. J." box="[189,473,1771,1797]" pageId="7" pageNumber="32" pagination="199 - 212" refId="ref11007" refString="Russell, P. J. C. &amp; Tennent, W. J. (2022) Butterflies observed on the western (European) slopes of the southern Ural Mountains during three spring visits in 2011, 2012 and 2014 (Lepidoptera Papilionoidea). Neue Entomologische Nachrichten 80: 199 - 212. [including 8 colour plates]" type="journal article" year="2022">Russell &amp; Tennent, 2022</bibRefCitation>
/ (=
<taxonomicName id="94C24D25FFFFFFE1E33F06B1FD1BF8C6" authorityName=", Russell" authorityYear="2021" box="[517,696,1771,1797]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="telona">
<emphasis id="61B6EAB4FFFFFFE1E33F06B1FD1BF8C6" box="[517,696,1771,1797]" italics="true" pageId="7" pageNumber="32">Melitaea telona</emphasis>
</taxonomicName>
/
<bibRefCitation id="37534B57FFFFFFE1E3F306B1FC31F8C6" author="Fruhstorfer, H." box="[713,914,1771,1797]" pageId="7" pageNumber="32" pagination="310" refId="ref9058" refString="Fruhstorfer, H. (1908) Neue Argynnis und Melitaea. Internationale Entomologische Zeitschrift, 1, 310." type="journal article" year="1908">Fruhstorfer, 1908</bibRefCitation>
)” (5) printed blue-bordered, circular “PARALECTO-TYPE”;
<specimenCount id="45C4FD2FFFFFFFE1E0670755FE61F8EA" box="[349,450,1807,1833]" pageId="7" pageNumber="32" type="female">1 female</specimenCount>
, (1) typed or handwritten “Trans
<collectingCountry id="2BD57636FFFFFFE1E2040755FC2FF8EA" box="[830,908,1807,1833]" name="Jordan" pageId="7" pageNumber="32">Jordan</collectingCountry>
:/ Petra./
<date id="277C1066FFFFFFE1E2CB0755FBCEF8EA" box="[1009,1133,1807,1833]" pageId="7" pageNumber="32" value="1923-03-26">26-iii-1923</date>
/ H.St.J.Philby./ Brit. Mus. 1923-457”; (2) typed or handwritten “B.M. TYPE/ No. Rh 8284/
<taxonomicName id="94C24D25FFFFFFE1E2570769FC47F88E" box="[877,996,1843,1869]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="phoebe">M. phoebe</taxonomicName>
/ dorae/
<specimenCount id="45C4FD2FFFFFFFE1E5040769FBF0F88E" box="[1086,1107,1843,1869]" pageId="7" pageNumber="32" type="female">♀</specimenCount>
Graves”; (3) printed, yellowbordered, circular “Cotype”; (4) typed “
<typeStatus id="8C798804FFFFFFE1E361070DFCE9F8B2" box="[603,842,1879,1905]" pageId="7" pageNumber="32" type="paralectotype">PARALECTOTYPE</typeStatus>
:/
<taxonomicName id="94C24D25FFFFFFE1E265070DFBC6F8B2" authorityName="Graves" authorityYear="1925" box="[863,1125,1879,1905]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="dorae">
<emphasis id="61B6EAB4FFFFFFE1E265070DFBC6F8B2" box="[863,1125,1879,1905]" italics="true" pageId="7" pageNumber="32">Melitaea phoebe dorae</emphasis>
</taxonomicName>
/
<bibRefCitation id="37534B57FFFFFFE1E54C070DFAACF8B2" author="Graves, P." box="[1142,1295,1878,1905]" pageId="7" pageNumber="32" pagination="17 - 126" refId="ref9143" refString="Graves, P. (1925) The Rhopalocera and Grypocera of Palestine and Transjordania. Transactions of the entomological Society of London, 73, 17 - 126, plates 4, 5." type="journal article" year="1925">Graves, 1925</bibRefCitation>
; designated/ by
<bibRefCitation id="37534B57FFFFFFE1E1860721FE74F856" author="Russell, P. J. C. &amp; Tennent, W. J." box="[188,471,1915,1941]" pageId="7" pageNumber="32" pagination="199 - 212" refId="ref11007" refString="Russell, P. J. C. &amp; Tennent, W. J. (2022) Butterflies observed on the western (European) slopes of the southern Ural Mountains during three spring visits in 2011, 2012 and 2014 (Lepidoptera Papilionoidea). Neue Entomologische Nachrichten 80: 199 - 212. [including 8 colour plates]" type="journal article" year="2022">Russell &amp; Tennent, 2022</bibRefCitation>
/ (=
<taxonomicName id="94C24D25FFFFFFE1E3380721FD17F856" authorityName=", Russell" authorityYear="2021" box="[514,692,1915,1941]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="32" phylum="Arthropoda" rank="species" species="telona">
<emphasis id="61B6EAB4FFFFFFE1E3380721FD17F856" box="[514,692,1915,1941]" italics="true" pageId="7" pageNumber="32">Melitaea telona</emphasis>
</taxonomicName>
/
<bibRefCitation id="37534B57FFFFFFE1E3FF0721FC2EF856" author="Fruhstorfer, H." box="[709,909,1915,1941]" pageId="7" pageNumber="32" pagination="310" refId="ref9058" refString="Fruhstorfer, H. (1908) Neue Argynnis und Melitaea. Internationale Entomologische Zeitschrift, 1, 310." type="journal article" year="1908">Fruhstorfer, 1908</bibRefCitation>
)” (5) printed blue-bordered, circular “PARA-/ LECTO-/ TYPE; (6) printed “NHMUK 014172682”;
<specimenCount id="45C4FD2FFFFFFFE1E3D107C5FCEDF87A" box="[747,846,1951,1977]" pageId="7" pageNumber="32" type="female">1 female</specimenCount>
, ditto, but
<date id="277C1066FFFFFFE1E2F807C4FBE0F87A" box="[962,1091,1950,1977]" pageId="7" pageNumber="32" value="1923-03-25">25-iii-1923</date>
, B.M. TYPE/ No. Rh 8283 and “NHMUK 014172683”.
</paragraph>
</subSubSection>
</treatment>
</document>