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<document ID-DOI="http://dx.doi.org/10.3897/zitteliana.96.73892" ID-GBIF-Dataset="250ca9ac-a54b-4433-a689-ed5779b1dfd3" ID-GBIF-Taxon="193856279" ID-Pensoft-Pub="2747-8106-96-1" ID-Pensoft-UUID="D5F97D7240455065AD6C43481AAD2D6D" ModsDocID="2747-8106-96-1" checkinTime="1646282116023" checkinUser="pensoft" docAuthor="Pandey, Dhirendra Kumar, Fuersich, Franz T., Alberti, Matthias, Das, Ranajit &amp; Saez, Federico Oloriz" docDate="2022" docId="DB2B51B818815CC0AB7C783411FD8E3F" docLanguage="en" docName="Zitteliana 96: 1-49" docOrigin="Zitteliana 96" docPubDate="2022-03-02" docSource="http://dx.doi.org/10.3897/zitteliana.96.73892" docTitle="Ammonites Zittel 1884" docType="treatment" docVersion="3" id="D5F97D7240455065AD6C43481AAD2D6D" lastPageNumber="1" masterDocId="D5F97D7240455065AD6C43481AAD2D6D" masterDocTitle="First population-level study of the ammonite genus Hildoglochiceras Spath, and the Lower Tithonian record of the Hildoglochiceras Horizon in the Kachchh Basin, India" masterLastPageNumber="49" masterPageNumber="1" pageId="0" pageNumber="1" updateTime="1646286631855" updateUser="ExternalLinkService">
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<mods:title>First population-level study of the ammonite genus Hildoglochiceras Spath, and the Lower Tithonian record of the Hildoglochiceras Horizon in the Kachchh Basin, India</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pandey, Dhirendra Kumar</mods:namePart>
<mods:affiliation>Department of Geology, School of Earth, Biological and Environmental Science, Central University of South Bihar SH 7, Gaya-Panchanpur Road, Village Karhara, Post Fatehpur, Gaya 824236 (Bihar), India</mods:affiliation>
<mods:nameIdentifier type="email">dhirendrap@cusb.ac.in</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fuersich, Franz T.</mods:namePart>
<mods:affiliation>FG Palaeoumwelt, GeoZentrum Nordbayern der Friedrich-Alexander-Universitaet Erlangen-Nuernberg, Loewenichstrasse 28, 91054 Erlangen, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Alberti, Matthias</mods:namePart>
<mods:affiliation>State Key Laboratory for Mineral Deposits Research, School of Earth Sciences and Engineering, Centre for Research and Education on Biological Evolution and Environment and Frontiers Science Center for Critical Earth Material Cycling, Nanjing University, Nanjing 210023, Jiangsu Province, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Das, Ranajit</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5308-3477</mods:nameIdentifier>
<mods:affiliation>Yenepoya Research Centre, Yenepoya (Deemed to be University), Mangalore- 575018, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Saez, Federico Oloriz</mods:namePart>
<mods:affiliation>Departamento de Estratigrafia y Paleontologia, Fac. Ciencias, Univ. Granada, Severo Ochoa s / n, Spain</mods:affiliation>
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<mods:titleInfo>
<mods:title>Zitteliana</mods:title>
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<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-03-02</mods:number>
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<mods:detail type="volume">
<mods:number>96</mods:number>
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<mods:start>1</mods:start>
<mods:end>49</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zitteliana.96.73892</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zitteliana.96.73892</mods:identifier>
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<treatment ID-GBIF-Taxon="193856279" LSID="urn:lsid:plazi:treatment:DB2B51B818815CC0AB7C783411FD8E3F" httpUri="http://treatment.plazi.org/id/DB2B51B818815CC0AB7C783411FD8E3F" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
Family
<taxonomicName LSID="DB2B51B8-1881-5CC0-AB7C-783411FD8E3F" authority="Zittel, 1884" authorityName="Zittel" authorityYear="1884" class="Cephalopoda" family="Haploceratidae" genus="Ammonites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="family">Haploceratidae Zittel, 1884</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
The taxa described under this family are
<taxonomicName authorityName="Zittel" authorityYear="1870" class="Cephalopoda" family="Haploceratidae" genus="Haploceras" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploceras" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Haploceras</emphasis>
</taxonomicName>
Zittel, 1870 and
<taxonomicName authority="Spath, 1924" authorityName="Spath" authorityYear="1924" class="Cephalopoda" family="Oppeliidae" genus="Hildoglochiceras" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hildoglochiceras" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Hildoglochiceras</emphasis>
</taxonomicName>
Spath, 1924. The shell of
<taxonomicName authorityName="Zittel" authorityYear="1870" class="Cephalopoda" family="Haploceratidae" genus="Haploceras" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploceras" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Haploceras</emphasis>
</taxonomicName>
shows fine growth lines when epigenized shell is preserved, while inner casts are smooth, with a small but variable umbilicus that initially opens gradually until the last ontogenic stage when it slightly unfolds. Macroconchiate
<taxonomicName authorityName="Zittel" authorityYear="1870" class="Cephalopoda" family="Haploceratidae" genus="Haploceras" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploceras" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Haploceras</emphasis>
</taxonomicName>
show a slightly flexuous peristome, while assumed microconchs are variable in size, incorporate wide and short lappets rather than narrow and pedunculated ones, but can resemble glochiceratins in the absence of peristomal structures. Incomplete specimens of
<taxonomicName authorityName="Zittel" authorityYear="1870" class="Cephalopoda" family="Haploceratidae" genus="Haploceras" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploceras" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Haploceras</emphasis>
</taxonomicName>
make species-level interpretations difficult.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Reports of the genus
<taxonomicName authorityName="Zittel" authorityYear="1870" class="Cephalopoda" family="Haploceratidae" genus="Haploceras" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploceras" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Haploceras</emphasis>
</taxonomicName>
have been variably interpreted before the mid-twentieth century when this genus was commonly applied to Kimmeridgian and Lower Tithonian glochiceratins elsewhere in the world (e.g.,
<bibRefCitation author="Ziegler, B" journalOrPublisher="Palaeontographica A" pageId="0" pageNumber="1" pagination="93 - 164" refId="B157" refString="Ziegler, B, 1958. Monographie der Ammonitengattung Glochiceras im Epikontinentalen Weissjura Mitteleuropas. Palaeontographica A 110: 93 - 164" title="Monographie der Ammonitengattung Glochiceras im Epikontinentalen Weissjura Mitteleuropas." volume="110" year="1958">Ziegler 1958</bibRefCitation>
), while the present knowledge about microconchiates is inconclusive. Across the Trans-Erythraean Trough,
<taxonomicName authorityName="Zittel" authorityYear="1870" class="Cephalopoda" family="Haploceratidae" genus="Haploceras" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploceras" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Haploceras</emphasis>
</taxonomicName>
has been more frequently reported from the northern and eastern margins than from western ones, although it is a rare genus across the Himalayas (from
<bibRefCitation author="Uhlig, V" journalOrPublisher="Colloque sur la limite Jurassique-Cretacee, Lyon, Neuchatel" pageId="0" pageNumber="1" refId="B142" refString="Uhlig, V, 1903. The Fauna of the Spiti Shales. Memoir Geological Survey lndia, Palaeontologica Indica, Calcutta ser. 15, 4(1): 1-132." title="The Fauna of the Spiti Shales. Memoir Geological Survey lndia, Palaeontologica Indica, Calcutta ser. 15, 4 (1): 1 - 132." year="1903">Uhlig 1903</bibRefCitation>
to
<bibRefCitation author="Enay, R" journalOrPublisher="Documents des Laboratoires de Geologie Lyon" pageId="0" pageNumber="1" pagination="1 - 246" refId="B47" refString="Enay, R, 2009. Les faunes d'ammonites de l'Oxfordien au Tithonien et la biostratigraphie des Spiti-Shales (Callovien superieur-Tithonien) de Thakkhola, Nepal Central. Documents des Laboratoires de Geologie Lyon 166: 1 - 246" title="Les faunes d'ammonites de l'Oxfordien au Tithonien et la biostratigraphie des Spiti-Shales (Callovien superieur-Tithonien) de Thakkhola, Nepal Central." volume="166" year="2009">
<normalizedToken originalValue="Énay">Enay</normalizedToken>
2009
</bibRefCitation>
). Southwards across eastern margins,
<taxonomicName authorityName="Zittel" authorityYear="1870" class="Cephalopoda" family="Haploceratidae" genus="Haploceras" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploceras" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Haploceras</emphasis>
</taxonomicName>
has been reported from southern Pakistan to Madagascar throughout the twentieth century (e.g.,
<bibRefCitation author="Lemoine, P" journalOrPublisher="Zitteliana" pageId="0" pageNumber="1" refId="B98" refString="Lemoine, P, 1910-1911. Ammonites du Jurassique superieur du cercle d'Analalava (Madagascar). Annales de Paleontologie, Paris 5: 137-168 (1910); 6: 45-64 (1911)." title="Ammonites du Jurassique superieur du cercle d'Analalava (Madagascar). Annales de Paleontologie, Paris 5: 137 - 168 (1910); 6: 45 - 64 (1911)." year="1910 - 1911">Lemoine 1910</bibRefCitation>
;
<bibRefCitation author="Collignon, M" journalOrPublisher="Rivista Italiana di Paleontologia e Stratigrafia" pageId="0" pageNumber="1" refId="B30" refString="Collignon, M, 1960. Atlas des fossiles caracteristiques de Madagascar. 6 (Tithonique). Republique malgache, Service geologique, Tananarive, pl. 134-175." title="Atlas des fossiles caracteristiques de Madagascar. 6 (Tithonique). Republique malgache, Service geologique, Tananarive, pl. 134 - 175." year="1960">Collignon 1960</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1016/S0016-6995(94)80137-1" author="Fatmi, AN" journalOrPublisher="Symposium volume III, Geological Survey of Denmark, Copenhagen" pageId="0" pageNumber="1" refId="B56" refString="Fatmi, AN, Zeiss, A, 1994. https://doi.org/10.1016/S0016-6995(94)80137-1" url="https://doi.org/10.1016/S0016-6995(94)80137-1" year="1994">Fatmi and Zeiss 1994</bibRefCitation>
). In contrast, across western margins,
<taxonomicName authorityName="Zittel" authorityYear="1870" class="Cephalopoda" family="Haploceratidae" genus="Haploceras" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploceras" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Haploceras</emphasis>
</taxonomicName>
was only registered from southern Yemen (
<bibRefCitation author="Howarth, MK" journalOrPublisher="Bulletin of the Natural History Museum, Geology Series" pageId="0" pageNumber="1" pagination="1 - 32" refId="B71" refString="Howarth, MK, Morris, NJ, 1998. The Jurassic and Lower Cretaceous of Wadi Hajar, southern Yemen. Bulletin of the Natural History Museum, Geology Series 54 (1): 1 - 32" title="The Jurassic and Lower Cretaceous of Wadi Hajar, southern Yemen." volume="54" year="1998">Howarth and Morris 1998</bibRefCitation>
) and Tanzania (from
<bibRefCitation author="Zwierzycki, J" journalOrPublisher="Palaeontographica Supplement" pageId="0" pageNumber="1" refId="B161" refString="Zwierzycki, J, 1914. Die Cephalopodenfauna der Tendaguruschichten in Deutsch-Ostafrika. Archiv fuer Biontologie Band 3, Heft 4: 8-96." title="Die Cephalopodenfauna der Tendaguruschichten in Deutsch-Ostafrika. Archiv fuer Biontologie Band 3, Heft 4: 8 - 96." year="1914">Zwierczyki 1914</bibRefCitation>
to
<bibRefCitation DOI="https://doi.org/10.1002/mmng.200900004" author="Bussert, R" journalOrPublisher="Fossil Record" pageId="0" pageNumber="1" pagination="141 - 174" refId="B23" refString="Bussert, R, Heinrich, WD, Aberhan, M, 2009. The Tendaguru Formation (Late Jurassic to Early Cretaceous, southern Tanzania): definition, palaeoenvironments, and sequence stratigraphy. Fossil Record 12 (2): 141 - 174, DOI: https://doi.org/10.1002/mmng.200900004" title="The Tendaguru Formation (Late Jurassic to Early Cretaceous, southern Tanzania): definition, palaeoenvironments, and sequence stratigraphy." url="https://doi.org/10.1002/mmng.200900004" volume="12" year="2009">Bussert et al. 2009</bibRefCitation>
), being absent or very rare in between.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Oppel" baseAuthorityYear="1870" class="Cephalopoda" family="Haploceratidae" genus="Haploceras" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploceras elimatum" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="elimatum">
<emphasis italics="true" pageId="0" pageNumber="1">Haploceras elimatum</emphasis>
</taxonomicName>
(Oppel, 1965) and allies represent the morphological group more widespread and have been commonly reported throughout epicontinental areas in the Trans-Erythraean Trough.
<taxonomicName baseAuthorityName="Oppel" baseAuthorityYear="1870" class="Cephalopoda" family="Haploceratidae" genus="Haploceras" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploceras elimatum" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="elimatum">
<emphasis italics="true" pageId="0" pageNumber="1">Haploceras elimatum</emphasis>
</taxonomicName>
(Oppel, 1965) and
<taxonomicName baseAuthorityName="Zeuschner" baseAuthorityYear="1846" class="Cephalopoda" family="Haploceratidae" genus="Haploceras" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploceras staszycii" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="staszycii">
<emphasis italics="true" pageId="0" pageNumber="1">Haploceras staszycii</emphasis>
</taxonomicName>
(Zeuschner, 1846) have commonly been recognized as close species, difficult to separate, for instance by
<bibRefCitation author="Rod, E" journalOrPublisher="The Journal of Geology" pageId="0" pageNumber="1" refId="B123" refString="Rod, E, 1937. Stratigraphie des Malm der Graustock-Hutstock-Gruppe (Melchtal, Kanton Obwalden), Dissertation, Bern (Staempfli)." title="Stratigraphie des Malm der Graustock-Hutstock-Gruppe (Melchtal, Kanton Obwalden), Dissertation, Bern (Staempfli)." year="1937">Rod (1937)</bibRefCitation>
and by (
<bibRefCitation author="Hoelder, H" journalOrPublisher="Neues Jahrbuch fuer Geologie und Palaeontologie, Abhandlungen" pageId="0" pageNumber="1" pagination="150 - 214" refId="B69" refString="Hoelder, H, Ziegler, B, 1959. Stratigraphische und faunistische Beziehungen im Weissen Jura (Kimmeridgium) zwischen Sueddeutschland und Ardeche. Neues Jahrbuch fuer Geologie und Palaeontologie, Abhandlungen 108 (2): 150 - 214" title="Stratigraphische und faunistische Beziehungen im Weissen Jura (Kimmeridgium) zwischen Sueddeutschland und Ardeche." volume="108" year="1959">
<normalizedToken originalValue="Hölder">Hoelder</normalizedToken>
and Ziegler 1959
</bibRefCitation>
), even as representing a single species (
<bibRefCitation author="Rod, E" journalOrPublisher="Eclogae geologicae Helvetiae" pageId="0" pageNumber="1" pagination="177 - 198" refId="B124" refString="Rod, E, 1946. Ueber ein Fossillager im oberen Malm der Melchtaleralpen. Eclogae geologicae Helvetiae 39: 177 - 198" title="Ueber ein Fossillager im oberen Malm der Melchtaleralpen." volume="39" year="1946">Rod 1946</bibRefCitation>
). They represent a dimorphic pair (
<bibRefCitation author="Barthel, KW" journalOrPublisher="Abhandlungen der Bayerischen Akademie der Wissenschaften Mathematisch-Naturwissenschaftliche Klasse, Neue Folge, Heft" pageId="0" pageNumber="1" pagination="5 - 30" refId="B8" refString="Barthel, KW, 1962. Zur Ammonitenfauna und Stratigraphie der Neuburger Bankkalke. Abhandlungen der Bayerischen Akademie der Wissenschaften Mathematisch-Naturwissenschaftliche Klasse, Neue Folge, Heft 105: 5 - 30" title="Zur Ammonitenfauna und Stratigraphie der Neuburger Bankkalke." volume="105" year="1962">Barthel 1962</bibRefCitation>
). The macroconch embraces all but one Tithonian haploceratin species with polymorphic microconchs (
<bibRefCitation author="Enay, R" journalOrPublisher="Journal of Asian Sciences" pageId="0" pageNumber="1" refId="B50" refString="Enay, R, Cecca, F, 1986. Structure et evolution des populations tithoniques du genre d'ammonites tethysien Haploceras Zittel, 1868. In: Comitato Centenario Raffaele Piccinini, Pallini G (Eds) Fossili, Evolucione, Ambiente, Atti I Convegno Pergola, 33-53." title="Structure et evolution des populations tithoniques du genre d'ammonites tethysien Haploceras Zittel, 1868. In: Comitato Centenario Raffaele Piccinini, Pallini G (Eds) Fossili, Evolucione, Ambiente, Atti I Convegno Pergola, 33 - 53." year="1986">
<normalizedToken originalValue="Énay">Enay</normalizedToken>
and Cecca 1986
</bibRefCitation>
). According to the information available, the latter is a rather speculative proposal.
<bibRefCitation author="Zeiss, A" journalOrPublisher="Neue Denkschriften des Naturhistorischen Museums in Wien, 6: Band, Verlag Ferdinand Berger &amp; Soehne, Wien-Horn" pageId="0" pageNumber="1" refId="B154" refString="Zeiss, A, 2001. Die Ammonitenfauna der Tithonklippen von Ernstbrunn, Niederoesterreich. Neue Denkschriften des Naturhistorischen Museums in Wien, 6: Band, Verlag Ferdinand Berger &amp; Soehne, Wien-Horn" title="Die Ammonitenfauna der Tithonklippen von Ernstbrunn, Niederoesterreich." year="2001">Zeiss (2001)</bibRefCitation>
highlighted the existing options for interpreting dimorphism in haploceratins, and treated the morphologically close species
<taxonomicName baseAuthorityName="Oppel" baseAuthorityYear="1870" class="Cephalopoda" family="Haploceratidae" genus="Haploceras" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploceras elimatum" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="elimatum">
<emphasis italics="true" pageId="0" pageNumber="1">Haploceras elimatum</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Zeuschner" baseAuthorityYear="1846" class="Cephalopoda" family="Haploceratidae" genus="Haploceras" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploceras staszycii" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="staszycii">
<emphasis italics="true" pageId="0" pageNumber="1">Haploceras staszycii</emphasis>
</taxonomicName>
as separate units only for convenience. Based on all the above mentioned, these two morphospecies are here interpreted as conspecific and referred to as
<taxonomicName baseAuthorityName="Zeuschner" baseAuthorityYear="1846" class="Cephalopoda" family="Haploceratidae" genus="Haploceras" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploceras staszycii" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="staszycii">
<emphasis italics="true" pageId="0" pageNumber="1">Haploceras staszycii</emphasis>
</taxonomicName>
by priority, with reference to the closest nominal morphological species reported from the Trans-Erythraean Trough.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authority="Spath, 1924" authorityName="Spath" authorityYear="1924" class="Cephalopoda" family="Oppeliidae" genus="Hildoglochiceras" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hildoglochiceras" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Hildoglochiceras</emphasis>
</taxonomicName>
is commonly more evolute, characterized by a median lateral groove and, consequently, with an acute-oval to oval whorl-section, venter of variable width and height, a lateral sulcus above the lower one-third of the flanks, a variable shell-size for the beginning of ribbing, and more or less ornamented outer whorls. In the present collection, we recognise two groups within
<taxonomicName authority="Spath, 1924" authorityName="Spath" authorityYear="1924" class="Cephalopoda" family="Oppeliidae" genus="Hildoglochiceras" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hildoglochiceras" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Hildoglochiceras</emphasis>
</taxonomicName>
; one interpreted as microconch and other as the corresponding macroconch. These two morphs have been separated on the basis of shell size and the diameter of the umbilicus. The macroconch shows a subrectangular to oval whorl section. See previous chapters for a revision of reports of
<taxonomicName authority="Spath, 1924" authorityName="Spath" authorityYear="1924" class="Cephalopoda" family="Oppeliidae" genus="Hildoglochiceras" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hildoglochiceras" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Hildoglochiceras</emphasis>
</taxonomicName>
across the Trans-Erythraean Trough.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Another comparable genus to inner whorls of
<taxonomicName authorityName="Zittel" authorityYear="1870" class="Cephalopoda" family="Haploceratidae" genus="Haploceras" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploceras" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Haploceras</emphasis>
</taxonomicName>
and
<taxonomicName authority="Spath, 1924" authorityName="Spath" authorityYear="1924" class="Cephalopoda" family="Oppeliidae" genus="Hildoglochiceras" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hildoglochiceras" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Hildoglochiceras</emphasis>
</taxonomicName>
is
<taxonomicName authorityName="Hyatt" authorityYear="1900" class="Cephalopoda" family="Haploceratidae" genus="Glochiceras" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glochiceras" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Glochiceras</emphasis>
</taxonomicName>
Hyatt, 1900, the shell of which is smaller and shows a variable whorl section, sculpture and peristomal structure. The umbilicus of
<taxonomicName authorityName="Hyatt" authorityYear="1900" class="Cephalopoda" family="Haploceratidae" genus="Glochiceras" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glochiceras" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Glochiceras</emphasis>
</taxonomicName>
opens suddenly at the end of the growth. In addition, some species of
<taxonomicName authorityName="Hyatt" authorityYear="1900" class="Cephalopoda" family="Haploceratidae" genus="Glochiceras" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glochiceras" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Glochiceras</emphasis>
</taxonomicName>
are also characterised by a median lateral groove like in
<taxonomicName authority="Spath, 1924" authorityName="Spath" authorityYear="1924" class="Cephalopoda" family="Oppeliidae" genus="Hildoglochiceras" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hildoglochiceras" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Hildoglochiceras</emphasis>
</taxonomicName>
. But small size, a rather discoid shell with narrow venter, the type of peristome, and the biostratigraphic range of typical
<taxonomicName authorityName="Hyatt" authorityYear="1900" class="Cephalopoda" family="Haploceratidae" genus="Glochiceras" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glochiceras" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Glochiceras</emphasis>
</taxonomicName>
, i.e. from Oxfordian to Kimmeridgian, with scarce records from the Lower Tithonian (e.g.,
<bibRefCitation author="Ziegler, B" journalOrPublisher="Palaeontographica A" pageId="0" pageNumber="1" pagination="93 - 164" refId="B157" refString="Ziegler, B, 1958. Monographie der Ammonitengattung Glochiceras im Epikontinentalen Weissjura Mitteleuropas. Palaeontographica A 110: 93 - 164" title="Monographie der Ammonitengattung Glochiceras im Epikontinentalen Weissjura Mitteleuropas." volume="110" year="1958">Ziegler 1958</bibRefCitation>
;
<bibRefCitation author="Collignon, M" journalOrPublisher="Rivista Italiana di Paleontologia e Stratigrafia" pageId="0" pageNumber="1" refId="B30" refString="Collignon, M, 1960. Atlas des fossiles caracteristiques de Madagascar. 6 (Tithonique). Republique malgache, Service geologique, Tananarive, pl. 134-175." title="Atlas des fossiles caracteristiques de Madagascar. 6 (Tithonique). Republique malgache, Service geologique, Tananarive, pl. 134 - 175." year="1960">Collignon 1960</bibRefCitation>
;
<bibRefCitation author="Barthel, KW" journalOrPublisher="Abhandlungen der Bayerischen Akademie der Wissenschaften Mathematisch-Naturwissenschaftliche Klasse, Neue Folge, Heft" pageId="0" pageNumber="1" pagination="5 - 30" refId="B8" refString="Barthel, KW, 1962. Zur Ammonitenfauna und Stratigraphie der Neuburger Bankkalke. Abhandlungen der Bayerischen Akademie der Wissenschaften Mathematisch-Naturwissenschaftliche Klasse, Neue Folge, Heft 105: 5 - 30" title="Zur Ammonitenfauna und Stratigraphie der Neuburger Bankkalke." volume="105" year="1962">Barthel 1962</bibRefCitation>
;
<bibRefCitation author="Grigore, D" journalOrPublisher="Stiintele Naturii" pageId="0" pageNumber="1" pagination="29 - 36" refId="B67" refString="Grigore, D, 2019. Glochiceras Hyatt (Oppelidae, Ammonoidea) from the Upper Jurassic deposits of the Hăghimas Mts. (the eastern Carpathians - Romania). Oltenia. Studiisicomunicări. Stiintele Naturii 35 (2): 29 - 36" title="Glochiceras Hyatt (Oppelidae, Ammonoidea) from the Upper Jurassic deposits of the Hăghimas Mts. (the eastern Carpathians - Romania). Oltenia. Studiisicomunicări." volume="35" year="2019">Grigore 2019</bibRefCitation>
), may be diagnostic. These morphological features persist in the scarce
<taxonomicName authorityName="Hyatt" authorityYear="1900" class="Cephalopoda" family="Haploceratidae" genus="Glochiceras" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glochiceras" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Glochiceras</emphasis>
</taxonomicName>
reported from the lowermost Tithonian of Europe. Separation of glochiceratins and microconchite haploceratins can be difficult with incomplete specimens lacking the peristome.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Among evolute haploceratins with a comparatively wide ventral region,
<taxonomicName authorityName="Ziegler" authorityYear="1958" genus="Lingulaticeras" lsidName="Lingulaticeras" pageId="0" pageNumber="1" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Lingulaticeras</emphasis>
</taxonomicName>
Ziegler, 1958 and
<taxonomicName authorityName="Ziegler" authorityYear="1958" genus="Paralingulaticeras" lsidName="Paralingulaticeras" pageId="0" pageNumber="1" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Paralingulaticeras</emphasis>
</taxonomicName>
Ziegler, 1958 are relatively close in shell-type to
<taxonomicName authority="Spath, 1924" authorityName="Spath" authorityYear="1924" class="Cephalopoda" family="Oppeliidae" genus="Hildoglochiceras" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hildoglochiceras" order="Ammonoidea" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Hildoglochiceras</emphasis>
</taxonomicName>
. Supposed lowermost Tithonian forms of the former are more involute and show a sculptured venter of variable width, while those of the latter develop a latero-ventral tuberculation.
</paragraph>
</subSubSection>
</treatment>
</document>