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<document id="D0D9C1EBABEE66C4B90598E504737A1B" ID-DOI="10.5281/zenodo.209071" ID-GBIF-Dataset="5d6f7106-bc7f-4487-bdfa-0cedaff94c70" ID-ISSN="1175-5326" ID-Zenodo-Dep="209071" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460495772678" checkinUser="plazi" docAuthor="Ota, Yuzo" docDate="2012" docId="DB1B87CDF361FFA5FF38FAE60FE5F84E" docLanguage="en" docName="zt03367p094.pdf" docOrigin="Zootaxa 3367" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Gnathia kumejimensis Ota, 2012, n. sp." docType="treatment" docVersion="9" lastPageNumber="89" masterDocId="2722FFB5F369FFAFFFAFFFB90C02FFDA" masterDocTitle="Gnathiidae from Kumejima Island in the Ryukyu Archipelago, southwestern Japan, with description of three new species (Crustacea: Isopoda) *" masterLastPageNumber="94" masterPageNumber="79" pageNumber="87" updateTime="1698310894548" updateUser="plazi">
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<mods:title id="0465080BD1DFD50C53EFA64F54A84E4F">Gnathiidae from Kumejima Island in the Ryukyu Archipelago, southwestern Japan, with description of three new species (Crustacea: Isopoda) *</mods:title>
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<mods:namePart id="6663CF8989225F89177F1DAEACF063AC">Ota, Yuzo</mods:namePart>
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<mods:date id="0C80530A60FC5225980269C0F9C0307C">2012</mods:date>
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<mods:classification id="E724463D89A188651525C6DFBC7CDAF8">journal article</mods:classification>
<mods:identifier id="7F404B01E9A94740AACF2AB5349F2FE9" type="DOI">10.5281/zenodo.209071</mods:identifier>
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<treatment id="DB1B87CDF361FFA5FF38FAE60FE5F84E" ID-DOI="http://doi.org/10.5281/zenodo.6173346" ID-GBIF-Taxon="119520625" ID-Zenodo-Dep="6173346" LSID="urn:lsid:plazi:treatment:DB1B87CDF361FFA5FF38FAE60FE5F84E" httpUri="http://treatment.plazi.org/id/DB1B87CDF361FFA5FF38FAE60FE5F84E" lastPageId="10" lastPageNumber="89" pageId="8" pageNumber="87">
<subSubSection id="1BA86550F361FFA7FF38FAE60D10FA64" pageId="8" pageNumber="87" type="nomenclature">
<paragraph id="530D36DBF361FFA7FF38FAE60DF3FAA3" blockId="8.[151,775,1375,1470]" box="[151,497,1375,1401]" pageId="8" pageNumber="87">
<heading id="084581B7F361FFA7FF38FAE60DF3FAA3" bold="true" box="[151,497,1375,1401]" fontSize="11" level="1" pageId="8" pageNumber="87" reason="1">
<emphasis id="61C6EAC9F361FFA7FF38FAE60DF3FAA3" bold="true" box="[151,497,1375,1401]" pageId="8" pageNumber="87">
<taxonomicName id="94B24D58F361FFA7FF38FAE60DA5FAA3" ID-CoL="6KNBV" box="[151,423,1375,1401]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="87" phylum="Arthropoda" rank="species" species="kumejimensis" status="sp. nov.">
<emphasis id="61C6EAC9F361FFA7FF38FAE60DA5FAA3" bold="true" box="[151,423,1375,1401]" italics="true" pageId="8" pageNumber="87">Gnathia kumejimensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AF557B2F361FFA7FE00FAE60DF3FAA3" box="[431,497,1375,1401]" pageId="8" pageNumber="87" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="530D36DBF361FFA7FF38FA3A0D10FA64" blockId="8.[151,775,1375,1470]" pageId="8" pageNumber="87">
New Japanese name: Kumejima-nokogiri-umikuwagata (
<figureCitation id="CB892A5EF361FFA7FF30FA1F0CEDFA64" box="[159,239,1446,1470]" captionStart="FIGURE 6" captionStartId="9.[151,250,1946,1969]" captionTargetBox="[212,1380,193,1934]" captionTargetId="figure@9.[212,1380,193,1935]" captionTargetPageId="9" captionText="FIGURE 6. Gnathia kumejimensis n. sp., holotype, adult male (RUMF-ZC- 1430, total length, 4.85 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view (IL, internal lobe); D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209077/files/figure.png" pageId="8" pageNumber="87">Figs. 6</figureCitation>
,
<figureCitation id="CB892A5EF361FFA7FF55FA1F0D0BFA64" box="[250,265,1446,1470]" captionStart="FIGURE 7" captionStartId="11.[151,250,1899,1922]" captionTargetBox="[210,1391,196,1878]" captionTargetId="figure@11.[205,1391,195,1878]" captionTargetPageId="11" captionText="FIGURE 7. Gnathia kumejimensis n. sp., holotype, adult male (RUMF-ZC- 1430, total length, 4.85 mm): A, right maxilliped, ventral view; B, right pylopod, ventral view; C, right pereopod II, medial view; D, left pleopod II, ventral view; E, penes, ventral view." httpUri="https://zenodo.org/record/209078/files/figure.png" pageId="8" pageNumber="87">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1BA86550F361FFA7FF38FA57092AF8D8" pageId="8" pageNumber="87" type="materials_examined">
<paragraph id="530D36DBF361FFA7FF38FA570F54F994" blockId="8.[151,1436,1518,2010]" pageId="8" pageNumber="87">
<emphasis id="61C6EAC9F361FFA7FF38FA570D83F9DD" bold="true" box="[151,385,1518,1543]" pageId="8" pageNumber="87">Material examined.</emphasis>
<typeStatus id="8C098879F361FFA7FE26FA570DF7F9DC" box="[393,501,1518,1542]" pageId="8" pageNumber="87" type="holotype">Holotype</typeStatus>
. Male
<quantity id="944A9B3EF361FFA7FDEDFA570EAAF9DC" box="[578,680,1518,1542]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" pageId="8" pageNumber="87" unit="mm" value="4.85">4.85 mm</quantity>
in total length (RUMF-ZC-1430), from Stn. Dredge 11 (triangular dredge), 26°22.806ʹN, 126°48.035ʹE – 26°22.670ʹN, 126°48.327ʹE,
<quantity id="944A9B3EF361FFA7FC6CF9AB0807F9F0" box="[963,1029,1554,1578]" metricMagnitude="1" metricUnit="m" metricValue="6.8" pageId="8" pageNumber="87" unit="m" value="68.0">68 m</quantity>
depth, off Kumejima Island, the Ryukyu Archipelago, southwestern
<collectingCountry id="2BA5764BF361FFA7FD87F98E0E69F994" box="[552,619,1591,1614]" name="Japan" pageId="8" pageNumber="87">Japan</collectingCountry>
,
<date id="270C101BF361FFA7FDD9F98F0F50F994" box="[630,850,1590,1614]" pageId="8" pageNumber="87" value="2009-11-10">10 November 2009</date>
.
</paragraph>
<paragraph id="530D36DBF361FFA7FF68F9E2092AF8D8" blockId="8.[151,1436,1518,2010]" pageId="8" pageNumber="87">
<typeStatus id="8C098879F361FFA7FF68F9E20D3AF9A9" box="[199,312,1627,1651]" pageId="8" pageNumber="87" type="paratype">Paratypes</typeStatus>
. One male from the same site as the
<typeStatus id="8C098879F361FFA7FD61F9E30F32F9A8" box="[718,816,1626,1650]" pageId="8" pageNumber="87" type="holotype">holotype</typeStatus>
(RUMF-ZC-1431). One male (RUMF-ZC-1432), from dead coral rubble or rock rubble, Stn. Trawl 45, 26°19.907ʹN, 126°43.191ʹE –26°20.056ʹN, 126°42.622ʹE,
<quantity id="944A9B3EF361FFA7FAF2F9C70CD7F960" metricMagnitude="1" metricUnit="m" metricValue="7.175" metricValueMax="7.6" metricValueMin="6.75" pageId="8" pageNumber="87" unit="m" value="71.75" valueMax="76.0" valueMin="67.5">67.5– 76 m</quantity>
depth, off Kumejima Island,
<date id="270C101BF361FFA7FD95F91B0F1DF960" box="[570,799,1698,1722]" pageId="8" pageNumber="87" value="2009-11-16">16 November 2009</date>
, beam trawl, towed by “Daisan-Emimaru”. One male (RUMF-ZC-1433), from rock rubble, Stn. Dredge 76, 26°19.672ʹN, 126°43.328ʹE – 26°19.698ʹN, 126°42.957ʹE,
<quantity id="944A9B3EF361FFA7FF38F9530D1FF8D8" box="[151,285,1770,1794]" metricMagnitude="1" metricUnit="m" metricValue="9.709999999999999" metricValueMax="10.1" metricValueMin="9.32" pageId="8" pageNumber="87" unit="m" value="97.1" valueMax="101.0" valueMin="93.2">93.2–101 m</quantity>
depth, off Kumejima Island,
<date id="270C101BF361FFA7FDC9F9530F03F8D8" box="[614,769,1770,1794]" pageId="8" pageNumber="87" value="2009-11-19">19 Nov. 2009</date>
, triangular dredge, towed by “Daisan-Emimaru”.
</paragraph>
</subSubSection>
<subSubSection id="1BA86550F361FFA5FF68F8B70829F9F6" lastPageId="10" lastPageNumber="89" pageId="8" pageNumber="87" type="description">
<paragraph id="530D36DBF361FFA7FF68F8B70D3EF800" blockId="8.[151,1436,1518,2010]" pageId="8" pageNumber="87">
<emphasis id="61C6EAC9F361FFA7FF68F8B70D5BF8FD" bold="true" box="[199,345,1806,1831]" pageId="8" pageNumber="87">Description.</emphasis>
Male (
<figureCitation id="CB892A5EF361FFA7FE1EF8B70E07F8FC" box="[433,517,1806,1830]" captionStart="FIGURE 6" captionStartId="9.[151,250,1946,1969]" captionTargetBox="[212,1380,193,1934]" captionTargetId="figure@9.[212,1380,193,1935]" captionTargetPageId="9" captionText="FIGURE 6. Gnathia kumejimensis n. sp., holotype, adult male (RUMF-ZC- 1430, total length, 4.85 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view (IL, internal lobe); D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209077/files/figure.png" pageId="8" pageNumber="87">Figs. 6</figureCitation>
,
<figureCitation id="CB892A5EF361FFA7FDBAF8B70E27F8FC" box="[533,549,1806,1830]" captionStart="FIGURE 7" captionStartId="11.[151,250,1899,1922]" captionTargetBox="[210,1391,196,1878]" captionTargetId="figure@11.[205,1391,195,1878]" captionTargetPageId="11" captionText="FIGURE 7. Gnathia kumejimensis n. sp., holotype, adult male (RUMF-ZC- 1430, total length, 4.85 mm): A, right maxilliped, ventral view; B, right pylopod, ventral view; C, right pereopod II, medial view; D, left pleopod II, ventral view; E, penes, ventral view." httpUri="https://zenodo.org/record/209078/files/figure.png" pageId="8" pageNumber="87">7</figureCitation>
). Body
<quantity id="944A9B3EF361FFA7FD28F8B70F10F8FC" box="[647,786,1806,1830]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="8" pageNumber="87" unit="mm" value="5.0" valueMax="5.5" valueMin="4.5">4.5–5.5 mm</quantity>
(4.78 ±
<quantity id="944A9B3EF361FFA7FCD1F8B70FEAF8FC" box="[894,1000,1806,1830]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="8" pageNumber="87" unit="mm" value="0.23">0.23 mm</quantity>
, n = 4). Cephalothorax (
<figureCitation id="CB892A5EF361FFA7FABFF8B70954F8FC" box="[1296,1366,1806,1830]" captionStart="FIGURE 6" captionStartId="9.[151,250,1946,1969]" captionTargetBox="[212,1380,193,1934]" captionTargetId="figure@9.[212,1380,193,1935]" captionTargetPageId="9" captionText="FIGURE 6. Gnathia kumejimensis n. sp., holotype, adult male (RUMF-ZC- 1430, total length, 4.85 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view (IL, internal lobe); D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209077/files/figure.png" pageId="8" pageNumber="87">Fig. 6</figureCitation>
A–C). Cephalothorax almost rectangular, sparsely covered with setae. Dorsal sulcus narrow and shallow. Posterior margin concave. Frontal border concave with 3 setae on 2 superior frontolateral processes; frontal margin with 19 small tubercles. Mediofrontal process rounded with small round process on distal margin. Eyes with 55 ocelli in 8 horizontal rows. Paraocular ornamentation prominent, composed of several tubercles and long setae. Supraocular lobe not acute.
</paragraph>
<caption id="07CD6653F360FFA6FF38F8230FD1F82F" httpUri="https://zenodo.org/record/209077/files/figure.png" pageId="9" pageNumber="88" targetBox="[212,1380,193,1934]" targetPageId="9">
<paragraph id="530D36DBF360FFA6FF38F8230FD1F82F" blockId="9.[151,1436,1946,2037]" pageId="9" pageNumber="88">
<emphasis id="61C6EAC9F360FFA6FF38F8230D10F86A" bold="true" box="[151,274,1946,1969]" pageId="9" pageNumber="88">FIGURE 6.</emphasis>
<taxonomicName id="94B24D58F360FFA6FEB7F8230DFBF86A" box="[280,505,1946,1968]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="88" phylum="Arthropoda" rank="species" species="kumejimensis" status="sp. nov.">
<emphasis id="61C6EAC9F360FFA6FEB7F8230DFBF86A" box="[280,505,1946,1968]" italics="true" pageId="9" pageNumber="88">Gnathia kumejimensis</emphasis>
</taxonomicName>
<emphasis id="61C6EAC9F360FFA6FE50F8220E35F86A" bold="true" box="[511,567,1947,1968]" pageId="9" pageNumber="88">
<taxonomicNameLabel id="7AF557B2F360FFA6FE50F8220E35F86A" box="[511,567,1947,1968]" pageId="9" pageNumber="88" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, adult male (RUMF-ZC-1430, total length, 4.85 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view (IL, internal lobe); D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view.
</paragraph>
</caption>
<paragraph id="530D36DBF363FFA5FF68FF210E4CFEE5" blockId="10.[151,1437,151,1940]" pageId="10" pageNumber="89">
Pereon (
<figureCitation id="CB892A5EF363FFA5FE89FF2E0D69FF75" box="[294,363,151,175]" captionStart="FIGURE 6" captionStartId="9.[151,250,1946,1969]" captionTargetBox="[212,1380,193,1934]" captionTargetId="figure@9.[212,1380,193,1935]" captionTargetPageId="9" captionText="FIGURE 6. Gnathia kumejimensis n. sp., holotype, adult male (RUMF-ZC- 1430, total length, 4.85 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view (IL, internal lobe); D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209077/files/figure.png" pageId="10" pageNumber="89">Fig. 6</figureCitation>
A). Pereonite I slightly shorter than pereonite II, not fused, separated into 3 parts by posterior margin of cephalothorax. Pereonite II and III subequal in length and width, posterior margins almost straight. Pereonite IV with anterior constriction; median groove closed. Pereonite IV with areae laterales; semicircular parts visible. Pereonite VI as long as combined kength of pereonite IV and V. Pereonite VII not extending postero-lateral of pereonite VI, overlapping pleonite I.
</paragraph>
<paragraph id="530D36DBF363FFA5FF68FEF50D71FE5D" blockId="10.[151,1437,151,1940]" pageId="10" pageNumber="89">
Pleon (
<figureCitation id="CB892A5EF363FFA5FEB4FEF50D61FEBE" box="[283,355,332,356]" captionStart="FIGURE 6" captionStartId="9.[151,250,1946,1969]" captionTargetBox="[212,1380,193,1934]" captionTargetId="figure@9.[212,1380,193,1935]" captionTargetPageId="9" captionText="FIGURE 6. Gnathia kumejimensis n. sp., holotype, adult male (RUMF-ZC- 1430, total length, 4.85 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view (IL, internal lobe); D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209077/files/figure.png" pageId="10" pageNumber="89">Fig. 6</figureCitation>
A). All pleonites sparsely covered with setae. Pleonites II–V subequal in length and width; epimera prominent.
</paragraph>
<paragraph id="530D36DBF363FFA5FF68FE2D08D3FE15" blockId="10.[151,1437,151,1940]" pageId="10" pageNumber="89">
Pleotelson (
<figureCitation id="CB892A5EF363FFA5FEE1FE2D0D96FE76" box="[334,404,404,428]" captionStart="FIGURE 6" captionStartId="9.[151,250,1946,1969]" captionTargetBox="[212,1380,193,1934]" captionTargetId="figure@9.[212,1380,193,1935]" captionTargetPageId="9" captionText="FIGURE 6. Gnathia kumejimensis n. sp., holotype, adult male (RUMF-ZC- 1430, total length, 4.85 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view (IL, internal lobe); D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209077/files/figure.png" pageId="10" pageNumber="89">Fig. 6</figureCitation>
D). Width approximately 0.88 length. 3 pairs of setae bearing near lateral margin, central part, and apex. Lateral margins dentate, slightly concave at proximal third and convex at middle.
</paragraph>
<paragraph id="530D36DBF363FFA5FF68FE650D5CFD85" blockId="10.[151,1437,151,1940]" pageId="10" pageNumber="89">
Mandible (
<figureCitation id="CB892A5EF363FFA5FEE8FE650D92FE2E" box="[327,400,476,500]" captionStart="FIGURE 6" captionStartId="9.[151,250,1946,1969]" captionTargetBox="[212,1380,193,1934]" captionTargetId="figure@9.[212,1380,193,1935]" captionTargetPageId="9" captionText="FIGURE 6. Gnathia kumejimensis n. sp., holotype, adult male (RUMF-ZC- 1430, total length, 4.85 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view (IL, internal lobe); D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209077/files/figure.png" pageId="10" pageNumber="89">Fig. 6</figureCitation>
B, C). Mandible approximately 0.8 times as long as cephalothorax; apex curved dorsomesially. Mandibular seta present on mid-dorsal surface near incisor. Dentate blade occupying approximately 0.4 times as long as mandible. Erisma prominent. Internal lobe visible in lateral view, remarkably elongated dorsally, extending incisor.
</paragraph>
<paragraph id="530D36DBF363FFA5FF68FDD50D39FD2A" blockId="10.[151,1437,151,1940]" pageId="10" pageNumber="89">
Antennula (
<figureCitation id="CB892A5EF363FFA5FEE2FDD50D90FD5E" box="[333,402,620,644]" captionStart="FIGURE 6" captionStartId="9.[151,250,1946,1969]" captionTargetBox="[212,1380,193,1934]" captionTargetId="figure@9.[212,1380,193,1935]" captionTargetPageId="9" captionText="FIGURE 6. Gnathia kumejimensis n. sp., holotype, adult male (RUMF-ZC- 1430, total length, 4.85 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view (IL, internal lobe); D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209077/files/figure.png" pageId="10" pageNumber="89">Fig. 6</figureCitation>
E). 3 basal podomeres and 5 flagellar articles. Distal margins of basal podomeres I, II, III, and flagellar articles I and V bearing 3, 3, 1, 2, and 1 penicillate setae, respectively. Basal podomere III bearing 6 long setae on external margin. Flagellar articles III and IV each with 1 aesthetasc. Flagellar article V bearing 3 terminal setae.
</paragraph>
<paragraph id="530D36DBF363FFA5FF68FD450DA8FC5A" blockId="10.[151,1437,151,1940]" pageId="10" pageNumber="89">
Antenna (
<figureCitation id="CB892A5EF363FFA5FE97FD450D7CFCCE" box="[312,382,764,788]" captionStart="FIGURE 6" captionStartId="9.[151,250,1946,1969]" captionTargetBox="[212,1380,193,1934]" captionTargetId="figure@9.[212,1380,193,1935]" captionTargetPageId="9" captionText="FIGURE 6. Gnathia kumejimensis n. sp., holotype, adult male (RUMF-ZC- 1430, total length, 4.85 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view (IL, internal lobe); D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209077/files/figure.png" pageId="10" pageNumber="89">Fig. 6</figureCitation>
F). 4 basal podomeres and 7 flagellar articles. Basal podomeres III and IV bearing 1 and 5 penicillate setae on distal margins, respectively. Basal podomeres III and IV densely covered with long setae on distal margin and lateral margin, respectively. Flagellar articles I–VII with few setae on distal margins; article VII bearing 4 terminal setae.
</paragraph>
<paragraph id="530D36DBF363FFA5FF68FC350898FC12" blockId="10.[151,1437,151,1940]" pageId="10" pageNumber="89">
Maxilliped (
<figureCitation id="CB892A5EF363FFA5FEFDFC350D95FC7E" box="[338,407,908,932]" captionStart="FIGURE 7" captionStartId="11.[151,250,1899,1922]" captionTargetBox="[210,1391,196,1878]" captionTargetId="figure@11.[205,1391,195,1878]" captionTargetPageId="11" captionText="FIGURE 7. Gnathia kumejimensis n. sp., holotype, adult male (RUMF-ZC- 1430, total length, 4.85 mm): A, right maxilliped, ventral view; B, right pylopod, ventral view; C, right pereopod II, medial view; D, left pleopod II, ventral view; E, penes, ventral view." httpUri="https://zenodo.org/record/209078/files/figure.png" pageId="10" pageNumber="89">Fig. 7</figureCitation>
A). Endite reaching distal margin of palp article I. Palp articles I–IV bearing 4, 7, 5, and 7 plumose setae, respectively, on external margins; article IV bearing 4 simple terminal setae.
</paragraph>
<paragraph id="530D36DBF363FFA5FF68FC6D0898FBEE" blockId="10.[151,1437,151,1940]" pageId="10" pageNumber="89">
Pylopod (
<figureCitation id="CB892A5EF363FFA5FE98FC6D0D7FFC36" box="[311,381,980,1004]" captionStart="FIGURE 7" captionStartId="11.[151,250,1899,1922]" captionTargetBox="[210,1391,196,1878]" captionTargetId="figure@11.[205,1391,195,1878]" captionTargetPageId="11" captionText="FIGURE 7. Gnathia kumejimensis n. sp., holotype, adult male (RUMF-ZC- 1430, total length, 4.85 mm): A, right maxilliped, ventral view; B, right pylopod, ventral view; C, right pereopod II, medial view; D, left pleopod II, ventral view; E, penes, ventral view." httpUri="https://zenodo.org/record/209078/files/figure.png" pageId="10" pageNumber="89">Fig. 7</figureCitation>
B). 3-articulate; 13 and 14 setae bearing near distal margin of article I and II, respectively. Article I elliptical with 3 areolae bearing 30 plumose setae on internal margin; 1 penicillate seta bearing near basis. Article II elliptical, fringed with fine setae; 12 setae near distal margin. Article III elliptical.
</paragraph>
<paragraph id="530D36DBF363FFA5FF68FBF90D8CFAEA" blockId="10.[151,1437,151,1940]" pageId="10" pageNumber="89">
Pereopod II (
<figureCitation id="CB892A5EF363FFA5FEF6FBF90D9EFB82" box="[345,412,1088,1112]" captionStart="FIGURE 7" captionStartId="11.[151,250,1899,1922]" captionTargetBox="[210,1391,196,1878]" captionTargetId="figure@11.[205,1391,195,1878]" captionTargetPageId="11" captionText="FIGURE 7. Gnathia kumejimensis n. sp., holotype, adult male (RUMF-ZC- 1430, total length, 4.85 mm): A, right maxilliped, ventral view; B, right pylopod, ventral view; C, right pereopod II, medial view; D, left pleopod II, ventral view; E, penes, ventral view." httpUri="https://zenodo.org/record/209078/files/figure.png" pageId="10" pageNumber="89">Fig. 7</figureCitation>
C). Merus, carpus, and propodus inner margins covered with pectinate scales. Basis oblong, with 3 penicillate setae and 5 processes on outer margin. Ischium approximately 0.8 times as long as basis, becoming wider distally; setae bearing sparsely and 3 long setae near outer distal margin. Merus approximately 0.6 times as long as ischium; 4 setae on distal corner and 5 process on inner margin. Carpus three-fourth length of merus, bearing 1 spine, 3 setae, and 6 processes on inner margin. Propodus rectangular and 1.5 times as long as carpus; 2 spines on inner-middle and inner-distal margins and 1 penicillate seta on distal corner. Dactylus terminating in unguis.
</paragraph>
<paragraph id="530D36DBF363FFA5FF68FA850905FA46" blockId="10.[151,1437,151,1940]" pageId="10" pageNumber="89">
Pleopod II (
<figureCitation id="CB892A5EF363FFA5FEFEFA850D95FA8E" box="[337,407,1340,1364]" captionStart="FIGURE 7" captionStartId="11.[151,250,1899,1922]" captionTargetBox="[210,1391,196,1878]" captionTargetId="figure@11.[205,1391,195,1878]" captionTargetPageId="11" captionText="FIGURE 7. Gnathia kumejimensis n. sp., holotype, adult male (RUMF-ZC- 1430, total length, 4.85 mm): A, right maxilliped, ventral view; B, right pylopod, ventral view; C, right pereopod II, medial view; D, left pleopod II, ventral view; E, penes, ventral view." httpUri="https://zenodo.org/record/209078/files/figure.png" pageId="10" pageNumber="89">Fig. 7</figureCitation>
D). Protopod bearing coupling hook and 1 seta on distal corner; pectinate scales on inner margin. Both rami elliptic and same length; 8 and 9 plumose setae on endopod and exopod, respectively. Appendix masculina absent. Both rami of all pleopods same shapes, length, and same number of plumose setae.
</paragraph>
<paragraph id="530D36DBF363FFA5FF68FA11095AF9D2" blockId="10.[151,1437,151,1940]" pageId="10" pageNumber="89">
Uropod (
<figureCitation id="CB892A5EF363FFA5FE9EFA110D79FA1A" box="[305,379,1448,1472]" captionStart="FIGURE 6" captionStartId="9.[151,250,1946,1969]" captionTargetBox="[212,1380,193,1934]" captionTargetId="figure@9.[212,1380,193,1935]" captionTargetPageId="9" captionText="FIGURE 6. Gnathia kumejimensis n. sp., holotype, adult male (RUMF-ZC- 1430, total length, 4.85 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view (IL, internal lobe); D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209077/files/figure.png" pageId="10" pageNumber="89">Fig. 6</figureCitation>
D). Both rami subequal in length, extending beyond apex of pleotelson. Longest seta on uropodal rami much longer than pleotelson. Exopod bearing 16 or 19 setae and 3 plumose setae laterally. Endopod bearing 3 or 4 setae and 5 or 6 plumose setae laterally. Dorsal surface of endopod bearing 3 penicillate setae.
</paragraph>
<paragraph id="530D36DBF363FFA5FF68F9AC0829F9F6" blockId="10.[151,1437,151,1940]" box="[199,1067,1556,1580]" pageId="10" pageNumber="89">
Penes (
<figureCitation id="CB892A5EF363FFA5FEB8F9AD0D59F9F6" box="[279,347,1556,1580]" captionStart="FIGURE 7" captionStartId="11.[151,250,1899,1922]" captionTargetBox="[210,1391,196,1878]" captionTargetId="figure@11.[205,1391,195,1878]" captionTargetPageId="11" captionText="FIGURE 7. Gnathia kumejimensis n. sp., holotype, adult male (RUMF-ZC- 1430, total length, 4.85 mm): A, right maxilliped, ventral view; B, right pylopod, ventral view; C, right pereopod II, medial view; D, left pleopod II, ventral view; E, penes, ventral view." httpUri="https://zenodo.org/record/209078/files/figure.png" pageId="10" pageNumber="89">Fig. 7</figureCitation>
F). Penes composed of 2 contiguous papillae and not prominent.
</paragraph>
</subSubSection>
<subSubSection id="1BA86550F363FFA5FF68F9810DA0F9AE" pageId="10" pageNumber="89" type="etymology">
<paragraph id="530D36DBF363FFA5FF68F9810DA0F9AE" blockId="10.[151,1437,151,1940]" pageId="10" pageNumber="89">
<emphasis id="61C6EAC9F363FFA5FF68F9810D4CF98A" bold="true" box="[199,334,1592,1616]" pageId="10" pageNumber="89">Etymology.</emphasis>
The scientific name
<taxonomicName id="94B24D58F363FFA5FDEDF9810EDBF98A" box="[578,729,1592,1616]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="89" phylum="Arthropoda" rank="species" species="kumejimensis">
<emphasis id="61C6EAC9F363FFA5FDEDF9810EDBF98A" box="[578,729,1592,1616]" italics="true" pageId="10" pageNumber="89">kumejimensis</emphasis>
</taxonomicName>
is derived from the island, Kumejima where the present new species was discovered.
</paragraph>
</subSubSection>
<subSubSection id="1BA86550F363FFA5FF68F9390F84F8F2" pageId="10" pageNumber="89" type="discussion">
<paragraph id="530D36DBF363FFA5FF68F9390F84F8F2" blockId="10.[151,1437,151,1940]" pageId="10" pageNumber="89">
<emphasis id="61C6EAC9F363FFA5FF68F9390D39F942" bold="true" box="[199,315,1664,1688]" pageId="10" pageNumber="89">Remarks.</emphasis>
The present new species is most similar to
<taxonomicName id="94B24D58F363FFA5FCFCF939099AF942" authority="Kensley, Schotte &amp; Poore, 2009" authorityName="Kensley, Schotte &amp; Poore" authorityYear="2009" box="[851,1432,1664,1688]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="89" phylum="Arthropoda" rank="species" species="serrula">
<emphasis id="61C6EAC9F363FFA5FCFCF939080EF942" box="[851,1036,1664,1688]" italics="true" pageId="10" pageNumber="89">Gnathia serrula</emphasis>
<bibRefCitation id="37234B2AF363FFA5FBB7F939099AF942" author="Kensley" box="[1048,1432,1664,1688]" pageId="10" pageNumber="89" refString="Kensley, B., Schotte, M. &amp; Poore, G. C. B. (2009) Gnathiid isopods (Crustacea: Isopoda: Gnathiidae), mostly new, from the Indian Ocean. Proceedings of the Biological Society of Washington, 122 (1), 32 - 51." type="journal article" year="2009">Kensley, Schotte &amp; Poore, 2009</bibRefCitation>
</taxonomicName>
, because of the following characters; frontal border is concave with mediofrontal process, pleotelson bears two pairs of long setae near the lateral margin and on the apex (
<bibRefCitation id="37234B2AF363FFA5FD5DF9710FD6F93A" author="Kensley" box="[754,980,1736,1760]" pageId="10" pageNumber="89" refString="Kensley, B., Schotte, M. &amp; Poore, G. C. B. (2009) Gnathiid isopods (Crustacea: Isopoda: Gnathiidae), mostly new, from the Indian Ocean. Proceedings of the Biological Society of Washington, 122 (1), 32 - 51." type="journal article" year="2009">
Kensley
<emphasis id="61C6EAC9F363FFA5FCF8F9700F8BF93A" box="[855,905,1736,1760]" italics="true" pageId="10" pageNumber="89">et al</emphasis>
. 2009
</bibRefCitation>
). This species, however, is distinguished from
<taxonomicName id="94B24D58F363FFA5FF78F9540D4BF8DE" box="[215,329,1772,1796]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="89" phylum="Arthropoda" rank="species" species="serrula">
<emphasis id="61C6EAC9F363FFA5FF78F9540D4BF8DE" box="[215,329,1772,1796]" italics="true" pageId="10" pageNumber="89">G. serrula</emphasis>
</taxonomicName>
by dorsally elongated internal lobe on the mandibles and a much larger number of setae on the ventral surface of pylopod (27 setae versus 8;
<bibRefCitation id="37234B2AF363FFA5FD34F8A90F78F8F2" author="Kensley" box="[667,890,1808,1832]" pageId="10" pageNumber="89" refString="Kensley, B., Schotte, M. &amp; Poore, G. C. B. (2009) Gnathiid isopods (Crustacea: Isopoda: Gnathiidae), mostly new, from the Indian Ocean. Proceedings of the Biological Society of Washington, 122 (1), 32 - 51." type="journal article" year="2009">
Kensley
<emphasis id="61C6EAC9F363FFA5FD51F8A80F2DF8F2" box="[766,815,1808,1832]" italics="true" pageId="10" pageNumber="89">et al</emphasis>
. 2009
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="1BA86550F363FFA5FF68F88D0FE5F84E" pageId="10" pageNumber="89" type="reference_group">
<paragraph id="530D36DBF363FFA5FF68F88D0FE5F84E" blockId="10.[151,1437,151,1940]" pageId="10" pageNumber="89">
<taxonomicName id="94B24D58F363FFA5FF68F88D0E51F896" ID-CoL="6KNCC" authority="Tanaka, 2004" authorityName="Tanaka" authorityYear="2004" box="[199,595,1844,1868]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="89" phylum="Arthropoda" rank="species" species="camuripenis">
<emphasis id="61C6EAC9F363FFA5FF68F88D0DB6F896" box="[199,436,1844,1868]" italics="true" pageId="10" pageNumber="89">Gnathia camuripenis</emphasis>
<bibRefCitation id="37234B2AF363FFA5FE15F88D0E51F896" author="Tanaka" box="[442,595,1844,1868]" pageId="10" pageNumber="89" refString="Tanaka, K. (2004) A new species of Gnathia (Isopoda: Cymothoida: Gnathiidae) from Ishigaki Island, the Ryukyus, southwestern Japan. Crustacean Research, 33, 51 - 60." type="journal article" year="2004">Tanaka, 2004</bibRefCitation>
</taxonomicName>
, was also collected at the same locality of this expedition and similar in the whole shape to the present new species.
<taxonomicName id="94B24D58F363FFA5FDC9F8E10F57F8AA" ID-CoL="6KNCC" box="[614,853,1880,1904]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="89" phylum="Arthropoda" rank="species" species="camuripenis">
<emphasis id="61C6EAC9F363FFA5FDC9F8E10F57F8AA" box="[614,853,1880,1904]" italics="true" pageId="10" pageNumber="89">Gnathia camuripenis</emphasis>
</taxonomicName>
, however, has prominent penes bent posteriorly on distal one-fourth length (
<bibRefCitation id="37234B2AF363FFA5FE03F8C50E43F84E" author="Tanaka" box="[428,577,1916,1940]" pageId="10" pageNumber="89" refString="Tanaka, K. (2004) A new species of Gnathia (Isopoda: Cymothoida: Gnathiidae) from Ishigaki Island, the Ryukyus, southwestern Japan. Crustacean Research, 33, 51 - 60." type="journal article" year="2004">Tanaka 2004</bibRefCitation>
), rather than two contiguous papillae.
</paragraph>
</subSubSection>
</treatment>
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<emphasis id="61C6EAC9F365FFA3FF38FDDF0D88FD5A" bold="true" box="[151,394,614,640]" italics="true" pageId="12" pageNumber="91">Tenerognathia visus</emphasis>
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, 26°26ʹN, 127°46ʹE]. —
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Kensley
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One male
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in total length (RUMF-ZC-1435), from dead coral rubble or rock rubble, Stn. Trawl 45 (triangular dredge), 26°19.907ʹN, 126°43.191ʹE – 26°20.056ʹN, 126°42.622ʹE, 67.5–76.0 m depth, off Kumejima Island, the Ryukyu Archipelago, southwestern
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, triangular dredge.
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<emphasis id="61C6EAC9F365FFA3FF68FCD40D39FC5F" bold="true" box="[199,315,877,901]" pageId="12" pageNumber="91">Remarks.</emphasis>
This species, belonging to a monospecific genus, was originally recorded from coral rubble in
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depth at Okinawajima Island, and from reef edge in
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depth at Ishigakijima Island.
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is easily identified from the other genera and species by some remarkable characters; very large eyes occupying whole of the side of head, elliptical cephalosome without dorsal sulcus and paraocular ornamentation, short simple mandible, and narrow and weakly chitinized pereonites (
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), which resembles whole shapes of other gnathiid larvae. The specimens collected in this study were identified as
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, 24°27ʹN, 124°13ʹE]. —
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Ota
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One male
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in total length (RUMF-ZC-1434), from sand and mud sediment, Stn. Dredge 68 (silver dredge), 26°19.095ʹN, 126°49.008ʹE – 26°18.898ʹN, 126°48.969ʹE,
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<emphasis id="61C6EAC9F365FFA3FF68FE260D39FE6D" bold="true" box="[199,315,415,439]" pageId="12" pageNumber="91">Remarks.</emphasis>
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was originally recorded from a coral rubble trap that was placed in 0.5–1.0 m depth at Ishigakijima Island, south Ryukyus. Both adults and larvae were described in detail. The adult male of this species is easily distinguished from the other
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species by remarkably large penes fused into a single crooked blade and mandible with 2 inner lobes (
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<mods:identifier id="66164335948AD2927E47300F730AD3A0" type="GBIF-Dataset">5d6f7106-bc7f-4487-bdfa-0cedaff94c70</mods:identifier>
<mods:identifier id="C49B7A12AF608FFDA26531D006196905" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="9D8C8DB3E6FFFEC2EACC6ADF56AF027A" type="Zenodo-Dep">209071</mods:identifier>
</mods:mods>
<treatment id="DB1B87CDF36BFFAAFF38FF2E0970FBEE" ID-DOI="http://doi.org/10.5281/zenodo.6173342" ID-GBIF-Taxon="119520623" ID-Zenodo-Dep="6173342" LSID="urn:lsid:plazi:treatment:DB1B87CDF36BFFAAFF38FF2E0970FBEE" httpUri="http://treatment.plazi.org/id/DB1B87CDF36BFFAAFF38FF2E0970FBEE" lastPageId="5" lastPageNumber="84" pageId="2" pageNumber="81">
<subSubSection id="1BA86550F36BFFADFF38FF2E0D13FF2C" pageId="2" pageNumber="81" type="nomenclature">
<paragraph id="530D36DBF36BFFADFF38FF2E0DB7FF6B" blockId="2.[151,615,151,246]" box="[151,437,151,177]" pageId="2" pageNumber="81">
<heading id="084581B7F36BFFADFF38FF2E0DB7FF6B" bold="true" box="[151,437,151,177]" fontSize="11" level="1" pageId="2" pageNumber="81" reason="1">
<emphasis id="61C6EAC9F36BFFADFF38FF2E0DB7FF6B" bold="true" box="[151,437,151,177]" pageId="2" pageNumber="81">
<taxonomicName id="94B24D58F36BFFADFF38FF2E0D69FF6B" ID-CoL="3GLBM" box="[151,363,151,177]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="excavata" status="sp. nov.">
<emphasis id="61C6EAC9F36BFFADFF38FF2E0D69FF6B" bold="true" box="[151,363,151,177]" italics="true" pageId="2" pageNumber="81">Gnathia excavata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AF557B2F36BFFADFEDDFF2E0DB7FF6B" box="[370,437,151,177]" pageId="2" pageNumber="81" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="530D36DBF36BFFADFF38FF020D13FF2C" blockId="2.[151,615,151,246]" pageId="2" pageNumber="81">
New Japanese name: Eguri-umikuwagata (
<figureCitation id="CB892A5EF36BFFADFF30FF670CEDFF2C" box="[159,239,222,246]" captionStart="FIGURE 2" captionStartId="3.[151,250,1944,1967]" captionTargetBox="[203,1380,179,1933]" captionTargetId="figure@3.[203,1380,179,1934]" captionTargetPageId="3" captionText="FIGURE 2. Gnathia excavata n. sp., holotype, adult male (RUMF-ZC- 1428, total length, 2.45 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view; D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209073/files/figure.png" pageId="2" pageNumber="81">Figs. 2</figureCitation>
,
<figureCitation id="CB892A5EF36BFFADFF55FF670D0AFF2C" box="[250,264,222,246]" captionStart="FIGURE 3" captionStartId="4.[151,250,1925,1948]" captionTargetBox="[177,1415,193,1909]" captionTargetId="figure@4.[172,1416,193,1909]" captionTargetPageId="4" captionText="FIGURE 3. Gnathia excavata n. sp., holotype, adult male (RUMF-ZC- 1428, total length, 2.45 mm): A, left maxilliped, ventral view; B, left pylopod, ventral view; C, left pereopod II, medial view; D, left pleopod II, ventral view; E, left pleopod IV, ventral view, plumose setae were omitted; F, penes, ventral view." httpUri="https://zenodo.org/record/209074/files/figure.png" pageId="2" pageNumber="81">3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1BA86550F36BFFADFF38FE9F0F54FE5C" pageId="2" pageNumber="81" type="materials_examined">
<paragraph id="530D36DBF36BFFADFF38FE9F0F54FE5C" blockId="2.[151,1437,294,2010]" pageId="2" pageNumber="81">
<emphasis id="61C6EAC9F36BFFADFF38FE9F0DFEFEE5" bold="true" box="[151,508,294,319]" pageId="2" pageNumber="81">
Material examined.
<typeStatus id="8C098879F36BFFADFE27FE9E0DF5FEE5" box="[392,503,295,319]" pageId="2" pageNumber="81" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male,
<quantity id="944A9B3EF36BFFADFDE5FE9F0EADFEE4" box="[586,687,294,318]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" pageId="2" pageNumber="81" unit="mm" value="2.45">2.45 mm</quantity>
in total length (RUMF-ZC-1428), from Stn. Dredge 23 (triangular dredge), 26°16.380ʹN, 126°51.502ʹE – 26°15.982ʹN, 126°51.304ʹE, 147–
<quantity id="944A9B3EF36BFFADFC76FEF30820FEB8" box="[985,1058,330,354]" metricMagnitude="2" metricUnit="m" metricValue="1.25" pageId="2" pageNumber="81" unit="m" value="125.0">125 m</quantity>
depth, off Kumejima Island, the Ryukyu Archipelago, southwestern
<collectingCountry id="2BA5764BF36BFFADFD87FED60E69FE5C" box="[552,619,367,390]" name="Japan" pageId="2" pageNumber="81">Japan</collectingCountry>
,
<date id="270C101BF36BFFADFDD9FED70F50FE5C" box="[630,850,366,390]" pageId="2" pageNumber="81" value="2009-11-12">12 November 2009</date>
.
</paragraph>
</subSubSection>
<subSubSection id="1BA86550F36BFFADFF68FE2B0850F848" pageId="2" pageNumber="81" type="description">
<paragraph id="530D36DBF36BFFADFF68FE2B0F28FE70" blockId="2.[151,1437,294,2010]" box="[199,810,402,427]" pageId="2" pageNumber="81">
<emphasis id="61C6EAC9F36BFFADFF68FE2B0D5BFE71" bold="true" box="[199,345,402,427]" pageId="2" pageNumber="81">Description.</emphasis>
Male (
<figureCitation id="CB892A5EF36BFFADFE07FE2B0DF5FE70" box="[424,503,402,426]" captionStart="FIGURE 2" captionStartId="3.[151,250,1944,1967]" captionTargetBox="[203,1380,179,1933]" captionTargetId="figure@3.[203,1380,179,1934]" captionTargetPageId="3" captionText="FIGURE 2. Gnathia excavata n. sp., holotype, adult male (RUMF-ZC- 1428, total length, 2.45 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view; D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209073/files/figure.png" pageId="2" pageNumber="81">Figs. 2</figureCitation>
,
<figureCitation id="CB892A5EF36BFFADFDACFE2B0E11FE70" box="[515,531,402,426]" captionStart="FIGURE 3" captionStartId="4.[151,250,1925,1948]" captionTargetBox="[177,1415,193,1909]" captionTargetId="figure@4.[172,1416,193,1909]" captionTargetPageId="4" captionText="FIGURE 3. Gnathia excavata n. sp., holotype, adult male (RUMF-ZC- 1428, total length, 2.45 mm): A, left maxilliped, ventral view; B, left pylopod, ventral view; C, left pereopod II, medial view; D, left pleopod II, ventral view; E, left pleopod IV, ventral view, plumose setae were omitted; F, penes, ventral view." httpUri="https://zenodo.org/record/209074/files/figure.png" pageId="2" pageNumber="81">3</figureCitation>
). Body
<quantity id="944A9B3EF36BFFADFDC3FE2B0ED2FE70" box="[620,720,402,426]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" pageId="2" pageNumber="81" unit="mm" value="2.45">2.45 mm</quantity>
(n = 1).
</paragraph>
<paragraph id="530D36DBF36BFFADFF68FE0F0E3FFD58" blockId="2.[151,1437,294,2010]" pageId="2" pageNumber="81">
Cephalothorax (
<figureCitation id="CB892A5EF36BFFADFED3FE0F0DC2FE14" box="[380,448,438,462]" captionStart="FIGURE 2" captionStartId="3.[151,250,1944,1967]" captionTargetBox="[203,1380,179,1933]" captionTargetId="figure@3.[203,1380,179,1934]" captionTargetPageId="3" captionText="FIGURE 2. Gnathia excavata n. sp., holotype, adult male (RUMF-ZC- 1428, total length, 2.45 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view; D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209073/files/figure.png" pageId="2" pageNumber="81">Fig. 2</figureCitation>
A–C). Cephalothorax almost square, dorsal surface sparsely covered with setae. Dorsal sulcus deep; superior frontolateral processes acute, with 1, 2 setae on left, right process, respectively. Posterior margin slightly but widely concave. Mediofrontal process broad, rounded with slightly bifid apex, dorso-ventrally thinner than region surrounded by dorsal sulcus. Anterior parts of marginal carina visible below mediofrontal process in dorsal view. Eyes with 49 ocelli in 8 horizontal rows. Paraocular ornamentation composed of 3 indistinct tubercles. Supraocular lobe not acute.
</paragraph>
<paragraph id="530D36DBF36BFFADFF68FD3608CDFCEC" blockId="2.[151,1437,294,2010]" pageId="2" pageNumber="81">
Pereon (
<figureCitation id="CB892A5EF36BFFADFE8BFD370D65FD7C" box="[292,359,654,678]" captionStart="FIGURE 2" captionStartId="3.[151,250,1944,1967]" captionTargetBox="[203,1380,179,1933]" captionTargetId="figure@3.[203,1380,179,1934]" captionTargetPageId="3" captionText="FIGURE 2. Gnathia excavata n. sp., holotype, adult male (RUMF-ZC- 1428, total length, 2.45 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view; D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209073/files/figure.png" pageId="2" pageNumber="81">Fig. 2</figureCitation>
A). Pereonite 1 slightly shorter than pereonite II, not fused, separated into 2 lateral and 1 central parts by posterior margin of cephalothorax and distal margin of pereonite II (
<figureCitation id="CB892A5EF36BFFADFC51FD0B083EFD10" box="[1022,1084,690,714]" captionStart="FIGURE 2" captionStartId="3.[151,250,1944,1967]" captionTargetBox="[203,1380,179,1933]" captionTargetId="figure@3.[203,1380,179,1934]" captionTargetPageId="3" captionText="FIGURE 2. Gnathia excavata n. sp., holotype, adult male (RUMF-ZC- 1428, total length, 2.45 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view; D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209073/files/figure.png" pageId="2" pageNumber="81">Fig 2</figureCitation>
C). Pereonite II slightly shorter than pereonite III; few setae on lateral margins. Pereonite IV with anterior constriction; anterolateral lobe absent. Pereonite V with areae laterales but not distinct. Pereonite VI slightly longer than combined length of pereonite IV and V. Pereonite VII not extending posterolateral margin of pereonite VI, overlapping pleonite I.
</paragraph>
<paragraph id="530D36DBF36BFFADFF68FCFB085EFC80" blockId="2.[151,1437,294,2010]" box="[199,1116,834,858]" pageId="2" pageNumber="81">
Pleon (
<figureCitation id="CB892A5EF36BFFADFEB9FCFB0D5BFC80" box="[278,345,834,858]" captionStart="FIGURE 2" captionStartId="3.[151,250,1944,1967]" captionTargetBox="[203,1380,179,1933]" captionTargetId="figure@3.[203,1380,179,1934]" captionTargetPageId="3" captionText="FIGURE 2. Gnathia excavata n. sp., holotype, adult male (RUMF-ZC- 1428, total length, 2.45 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view; D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209073/files/figure.png" pageId="2" pageNumber="81">Fig. 2</figureCitation>
A). Pleonites II–V subequal in length and width; epimera prominent.
</paragraph>
<paragraph id="530D36DBF36BFFADFF68FCDF0CEDFC78" blockId="2.[151,1437,294,2010]" pageId="2" pageNumber="81">
Pleotelson (
<figureCitation id="CB892A5EF36BFFADFEE4FCDF0D8CFCA4" box="[331,398,870,894]" captionStart="FIGURE 2" captionStartId="3.[151,250,1944,1967]" captionTargetBox="[203,1380,179,1933]" captionTargetId="figure@3.[203,1380,179,1934]" captionTargetPageId="3" captionText="FIGURE 2. Gnathia excavata n. sp., holotype, adult male (RUMF-ZC- 1428, total length, 2.45 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view; D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209073/files/figure.png" pageId="2" pageNumber="81">Fig. 2</figureCitation>
D). Width 0.90 length. Two pairs of setae on lateral margin and apex; lateral margins slightly convex.
</paragraph>
<paragraph id="530D36DBF36BFFADFF68FC170E87FBD4" blockId="2.[151,1437,294,2010]" pageId="2" pageNumber="81">
Mandible (
<figureCitation id="CB892A5EF36BFFADFEE7FC170D91FC1C" box="[328,403,942,966]" captionStart="FIGURE 2" captionStartId="3.[151,250,1944,1967]" captionTargetBox="[203,1380,179,1933]" captionTargetId="figure@3.[203,1380,179,1934]" captionTargetPageId="3" captionText="FIGURE 2. Gnathia excavata n. sp., holotype, adult male (RUMF-ZC- 1428, total length, 2.45 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view; D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209073/files/figure.png" pageId="2" pageNumber="81">Fig. 2</figureCitation>
B, C). Mandible approximately half length of cephalothorax; apex curved inward and dorsally. Mandibular seta present on mid-dorsal surface near incisor. Dentate blade occupying approximately onethird of mandible length. Erisma prominent.
</paragraph>
<paragraph id="530D36DBF36BFFADFF68FBA30E91FBA0" blockId="2.[151,1437,294,2010]" pageId="2" pageNumber="81">
Antennula (
<figureCitation id="CB892A5EF36BFFADFEE2FBA30D90FBE8" box="[333,402,1050,1074]" captionStart="FIGURE 2" captionStartId="3.[151,250,1944,1967]" captionTargetBox="[203,1380,179,1933]" captionTargetId="figure@3.[203,1380,179,1934]" captionTargetPageId="3" captionText="FIGURE 2. Gnathia excavata n. sp., holotype, adult male (RUMF-ZC- 1428, total length, 2.45 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view; D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209073/files/figure.png" pageId="2" pageNumber="81">Fig. 2</figureCitation>
E). 3 basal podomeres and 5 flagellar articles. Distal margins of basal podomeres I, II, III, and flagellar article V bearing 2, 2, 1, and 1 penicillate setae, respectively. Flagellar articles III–V each with 1 aesthetasc; article V bearing 3 terminal setae.
</paragraph>
<paragraph id="530D36DBF36BFFADFF68FB3E0DA8FB3C" blockId="2.[151,1437,294,2010]" pageId="2" pageNumber="81">
Antenna (
<figureCitation id="CB892A5EF36BFFADFE98FB3F0D7EFB44" box="[311,380,1158,1182]" captionStart="FIGURE 2" captionStartId="3.[151,250,1944,1967]" captionTargetBox="[203,1380,179,1933]" captionTargetId="figure@3.[203,1380,179,1934]" captionTargetPageId="3" captionText="FIGURE 2. Gnathia excavata n. sp., holotype, adult male (RUMF-ZC- 1428, total length, 2.45 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view; D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209073/files/figure.png" pageId="2" pageNumber="81">Fig. 2</figureCitation>
F). 4 basal podomeres and 7 flagellar articles. Distal margins of basal podomeres III and IV bearing 2 and 6 penicillate setae, respectively. Flagellar articles I–VII with few setae on distal margins; article VII bearing 5 terminal setae.
</paragraph>
<paragraph id="530D36DBF36BFFADFF68FB4B08A1FAF4" blockId="2.[151,1437,294,2010]" pageId="2" pageNumber="81">
Maxilliped (
<figureCitation id="CB892A5EF36BFFADFEFCFB4B0D95FAD0" box="[339,407,1266,1290]" captionStart="FIGURE 3" captionStartId="4.[151,250,1925,1948]" captionTargetBox="[177,1415,193,1909]" captionTargetId="figure@4.[172,1416,193,1909]" captionTargetPageId="4" captionText="FIGURE 3. Gnathia excavata n. sp., holotype, adult male (RUMF-ZC- 1428, total length, 2.45 mm): A, left maxilliped, ventral view; B, left pylopod, ventral view; C, left pereopod II, medial view; D, left pleopod II, ventral view; E, left pleopod IV, ventral view, plumose setae were omitted; F, penes, ventral view." httpUri="https://zenodo.org/record/209074/files/figure.png" pageId="2" pageNumber="81">Fig. 3</figureCitation>
A). Endite extending to distal margin of palp article I. Palp articles I–IV external margins bearing 3, 6, 4, and 7 plumose setae, respectively; article IV bearing 5 simple terminal setae.
</paragraph>
<paragraph id="530D36DBF36BFFADFF68FA830814FA40" blockId="2.[151,1437,294,2010]" pageId="2" pageNumber="81">
Pylopod (
<figureCitation id="CB892A5EF36BFFADFE98FA830D7FFA88" box="[311,381,1338,1362]" captionStart="FIGURE 3" captionStartId="4.[151,250,1925,1948]" captionTargetBox="[177,1415,193,1909]" captionTargetId="figure@4.[172,1416,193,1909]" captionTargetPageId="4" captionText="FIGURE 3. Gnathia excavata n. sp., holotype, adult male (RUMF-ZC- 1428, total length, 2.45 mm): A, left maxilliped, ventral view; B, left pylopod, ventral view; C, left pereopod II, medial view; D, left pleopod II, ventral view; E, left pleopod IV, ventral view, plumose setae were omitted; F, penes, ventral view." httpUri="https://zenodo.org/record/209074/files/figure.png" pageId="2" pageNumber="81">Fig. 3</figureCitation>
B). 3 articles expressed; distal margins of article I and II with 3 and 2 setae, respectively. Article I elliptical with 3 areolae bearing 27 plumose setae on internal margin and 3 setae near external margin. Article II elliptical, fringed with fine setae. Article III reduced and semicircular.
</paragraph>
<paragraph id="530D36DBF36BFFADFF68FA1F08CDF994" blockId="2.[151,1437,294,2010]" pageId="2" pageNumber="81">
Pereopod II (
<figureCitation id="CB892A5EF36BFFADFEF6FA1F0D99FA64" box="[345,411,1446,1470]" captionStart="FIGURE 3" captionStartId="4.[151,250,1925,1948]" captionTargetBox="[177,1415,193,1909]" captionTargetId="figure@4.[172,1416,193,1909]" captionTargetPageId="4" captionText="FIGURE 3. Gnathia excavata n. sp., holotype, adult male (RUMF-ZC- 1428, total length, 2.45 mm): A, left maxilliped, ventral view; B, left pylopod, ventral view; C, left pereopod II, medial view; D, left pleopod II, ventral view; E, left pleopod IV, ventral view, plumose setae were omitted; F, penes, ventral view." httpUri="https://zenodo.org/record/209074/files/figure.png" pageId="2" pageNumber="81">Fig. 3</figureCitation>
C). Inner margins of merus, carpus, and propodus with pectinate scales. Basis oblong, outer margin with 3 penicillate setae and 2 processes. Ischium approximately two-thirds length of basis, becoming wider distally; distal margin bearing 1 long seta and 3 shorter setae. Merus approximately half of ischium length, bearing 3 setae on distal margin. Carpus subequal in length to merus, bearing 1 spine on inner-middle margin. Propodus rectangular and 1.3 times as long as carpus; inner-middle and inner-distal margins with 2 spines.
</paragraph>
<paragraph id="530D36DBF36BFFADFF68F9E30E3CF8D8" blockId="2.[151,1437,294,2010]" pageId="2" pageNumber="81">
Pleopod II (
<figureCitation id="CB892A5EF36BFFADFEFCF9E30D9BF9A8" box="[339,409,1626,1650]" captionStart="FIGURE 3" captionStartId="4.[151,250,1925,1948]" captionTargetBox="[177,1415,193,1909]" captionTargetId="figure@4.[172,1416,193,1909]" captionTargetPageId="4" captionText="FIGURE 3. Gnathia excavata n. sp., holotype, adult male (RUMF-ZC- 1428, total length, 2.45 mm): A, left maxilliped, ventral view; B, left pylopod, ventral view; C, left pereopod II, medial view; D, left pleopod II, ventral view; E, left pleopod IV, ventral view, plumose setae were omitted; F, penes, ventral view." httpUri="https://zenodo.org/record/209074/files/figure.png" pageId="2" pageNumber="81">Fig. 3</figureCitation>
D, E). Protopod distomedial corner with 1 coupling hook and 1 seta. Both rami lengths subequal, shape elliptical; endopod and exopod with 7 and 9 plumose setae, respectively. Top of appendix masculina bearing 1 plumose seta; another appendix masculina of left pleopod 2 lacking plumose seta. Pleopods IV and V rami shorter than pleopods I–III; exopods shorter than endopods. Exopods each with 6–11 plumose setae, endopods each with 7–9 simple setae.
</paragraph>
<paragraph id="530D36DBF36BFFADFF68F8B70CD4F8B4" blockId="2.[151,1437,294,2010]" pageId="2" pageNumber="81">
Uropod (
<figureCitation id="CB892A5EF36BFFADFE83F8B70D6DF8FC" box="[300,367,1806,1830]" captionStart="FIGURE 2" captionStartId="3.[151,250,1944,1967]" captionTargetBox="[203,1380,179,1933]" captionTargetId="figure@3.[203,1380,179,1934]" captionTargetPageId="3" captionText="FIGURE 2. Gnathia excavata n. sp., holotype, adult male (RUMF-ZC- 1428, total length, 2.45 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view; D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209073/files/figure.png" pageId="2" pageNumber="81">Fig. 2</figureCitation>
D). Both rami subequal in length, slightly extending to pleotelson apex. Exopod bearing 6 setae and 3 plumose setae laterally. Endopod bearing 8 setae laterally. Dorsal surface of endopod bearing 5 penicillate setae.
</paragraph>
<paragraph id="530D36DBF36BFFADFF68F8C20850F848" blockId="2.[151,1437,294,2010]" box="[199,1106,1914,1938]" pageId="2" pageNumber="81">
Penes (
<figureCitation id="CB892A5EF36BFFADFEB8F8C30D58F848" box="[279,346,1914,1938]" captionStart="FIGURE 3" captionStartId="4.[151,250,1925,1948]" captionTargetBox="[177,1415,193,1909]" captionTargetId="figure@4.[172,1416,193,1909]" captionTargetPageId="4" captionText="FIGURE 3. Gnathia excavata n. sp., holotype, adult male (RUMF-ZC- 1428, total length, 2.45 mm): A, left maxilliped, ventral view; B, left pylopod, ventral view; C, left pereopod II, medial view; D, left pleopod II, ventral view; E, left pleopod IV, ventral view, plumose setae were omitted; F, penes, ventral view." httpUri="https://zenodo.org/record/209074/files/figure.png" pageId="2" pageNumber="81">Fig. 3</figureCitation>
F). Fused but not prominent; tip of penes composed with 3 papillae.
</paragraph>
</subSubSection>
<subSubSection id="1BA86550F36BFFADFF68F8260DC6F800" pageId="2" pageNumber="81" type="etymology">
<paragraph id="530D36DBF36BFFADFF68F8260DC6F800" blockId="2.[151,1437,294,2010]" pageId="2" pageNumber="81">
<emphasis id="61C6EAC9F36BFFADFF68F8260D4CF86D" bold="true" box="[199,334,1951,1975]" pageId="2" pageNumber="81">Etymology.</emphasis>
The scientific name
<taxonomicName id="94B24D58F36BFFADFD93F8190E9DF86D" box="[572,671,1952,1975]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="excavata">
<emphasis id="61C6EAC9F36BFFADFD93F8190E9DF86D" box="[572,671,1952,1975]" italics="true" pageId="2" pageNumber="81">excavata</emphasis>
</taxonomicName>
is derived from the Latin meaning “hollow”, referring to be deeply concaved on dorsal sulcus.
</paragraph>
</subSubSection>
<caption id="07CD6653F36AFFACFF38F8210ED4F831" httpUri="https://zenodo.org/record/209073/files/figure.png" pageId="3" pageNumber="82" targetBox="[203,1380,179,1933]" targetPageId="3">
<paragraph id="530D36DBF36AFFACFF38F8210ED4F831" blockId="3.[151,1436,1944,2027]" pageId="3" pageNumber="82">
<emphasis id="61C6EAC9F36AFFACFF38F8210D11F874" bold="true" box="[151,275,1944,1967]" pageId="3" pageNumber="82">FIGURE 2.</emphasis>
<taxonomicName id="94B24D58F36AFFACFEB5F8210DCCF875" box="[282,462,1944,1967]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="82" phylum="Arthropoda" rank="species" species="excavata" status="sp. nov.">
<emphasis id="61C6EAC9F36AFFACFEB5F8210DCCF875" box="[282,462,1944,1967]" italics="true" pageId="3" pageNumber="82">Gnathia excavata</emphasis>
</taxonomicName>
<emphasis id="61C6EAC9F36AFFACFE7BF8200E0CF874" bold="true" box="[468,526,1945,1966]" pageId="3" pageNumber="82">
<taxonomicNameLabel id="7AF557B2F36AFFACFE7BF8200E0CF874" box="[468,526,1945,1966]" pageId="3" pageNumber="82" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, adult male (RUMF-ZC-1428, total length, 2.45 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view; D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view.
</paragraph>
</caption>
<caption id="07CD6653F36DFFABFF38F83C0EDFF83A" httpUri="https://zenodo.org/record/209074/files/figure.png" pageId="4" pageNumber="83" targetBox="[177,1415,193,1909]" targetPageId="4">
<paragraph id="530D36DBF36DFFABFF38F83C0EDFF83A" blockId="4.[151,1436,1925,2016]" pageId="4" pageNumber="83">
<emphasis id="61C6EAC9F36DFFABFF38F83C0D10F841" bold="true" box="[151,274,1925,1948]" pageId="4" pageNumber="83">FIGURE 3.</emphasis>
<taxonomicName id="94B24D58F36DFFABFEB7F83F0DC9F846" box="[280,459,1926,1948]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="83" phylum="Arthropoda" rank="species" species="excavata" status="sp. nov.">
<emphasis id="61C6EAC9F36DFFABFEB7F83F0DC9F846" box="[280,459,1926,1948]" italics="true" pageId="4" pageNumber="83">Gnathia excavata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AF557B2F36DFFABFE7EF83F0E0BF841" box="[465,521,1926,1947]" pageId="4" pageNumber="83" rank="species">
<emphasis id="61C6EAC9F36DFFABFE7EF83F0E0BF841" bold="true" box="[465,521,1926,1947]" pageId="4" pageNumber="83">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, adult male (RUMF-ZC-1428, total length, 2.45 mm): A, left maxilliped, ventral view; B, left pylopod, ventral view; C, left pereopod II, medial view; D, left pleopod II, ventral view; E, left pleopod IV, ventral view, plumose setae were omitted; F, penes, ventral view.
</paragraph>
</caption>
<subSubSection id="1BA86550F36CFFAAFF68FF210D90FC7F" pageId="5" pageNumber="84" type="discussion">
<paragraph id="530D36DBF36CFFAAFF68FF210E45FEE5" blockId="5.[151,1436,151,1076]" pageId="5" pageNumber="84">
<emphasis id="61C6EAC9F36CFFAAFF68FF210D39FF6A" bold="true" box="[199,315,152,176]" pageId="5" pageNumber="84">Remarks.</emphasis>
Although gnathiid morphology quite differs between male adult, female adult, and larva, the taxonomy has traditionally been based on the male morphology only. Congeners of the adult males are distinguished by the frontal border, pleotelson, pylopod, maxilliped, and their combinations. In addition to these, number of setae on the mouthparts, the presence of tubercles and setae on cephalothorax and appendix masculina on pleopod II are also frequently used.
</paragraph>
<paragraph id="530D36DBF36CFFAAFF68FEF50D90FC7F" blockId="5.[151,1436,151,1076]" pageId="5" pageNumber="84">
Among the 190 species of the family
<taxonomicName id="94B24D58F36CFFAAFD3DFEF50F12FEBE" box="[658,784,332,356]" class="Malacostraca" family="Gnathiidae" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="84" phylum="Arthropoda" rank="family">Gnathiidae</taxonomicName>
,
<taxonomicName id="94B24D58F36CFFAAFC8FFEF50F7EFEBE" box="[800,892,332,356]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="61C6EAC9F36CFFAAFC8FFEF50F7EFEBE" box="[800,892,332,356]" italics="true" pageId="5" pageNumber="84">Gnathia</emphasis>
</taxonomicName>
and
<taxonomicName id="94B24D58F36CFFAAFC12FEF50858FEBE" box="[957,1114,332,356]" class="Malacostraca" family="Gnathiidae" genus="Caecognathia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="61C6EAC9F36CFFAAFC12FEF50858FEBE" box="[957,1114,332,356]" italics="true" pageId="5" pageNumber="84">Caecognathia</emphasis>
</taxonomicName>
species comprise the most number of species of this family (containing approximately 110 and 50 species, respectively). These two genera are closely related and have been even synonymised with each other until
<bibRefCitation id="37234B2AF36CFFAAFC46FE2D08F8FE76" author="Cohen" box="[1001,1274,404,428]" pageId="5" pageNumber="84" refString="Cohen, B. F. &amp; Poore, G. C. B. (1994) Phylogeny and biogeography of the Gnathiidae (Crustacea: Isopoda) with descriptions of new genera and species, most from south-eastern Australia. Memoirs of Museum of Victoria, 54 (2), 271 - 397." type="journal article" year="1994">Cohen &amp; Poore (1994)</bibRefCitation>
reviewed the classification of the
<taxonomicName id="94B24D58F36CFFAAFE32FE0E0E1EFE15" box="[413,540,439,463]" class="Malacostraca" family="Gnathiidae" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="84" phylum="Arthropoda" rank="family">Gnathiidae</taxonomicName>
.
<bibRefCitation id="37234B2AF36CFFAAFD9DFE0E0F57FE15" author="Cohen" box="[562,853,439,464]" pageId="5" pageNumber="84" refString="Cohen, B. F. &amp; Poore, G. C. B. (1994) Phylogeny and biogeography of the Gnathiidae (Crustacea: Isopoda) with descriptions of new genera and species, most from south-eastern Australia. Memoirs of Museum of Victoria, 54 (2), 271 - 397." type="journal article" year="1994">Cohen &amp; Poore (1994)</bibRefCitation>
recognised 10 genera, including
<taxonomicName id="94B24D58F36CFFAAFAAAFE010963FE0A" box="[1285,1377,440,464]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="61C6EAC9F36CFFAAFAAAFE010963FE0A" box="[1285,1377,440,464]" italics="true" pageId="5" pageNumber="84">Gnathia</emphasis>
</taxonomicName>
and
<taxonomicName id="94B24D58F36CFFAAFF38FE650D36FE2E" box="[151,308,476,500]" class="Malacostraca" family="Gnathiidae" genus="Caecognathia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="61C6EAC9F36CFFAAFF38FE650D36FE2E" box="[151,308,476,500]" italics="true" pageId="5" pageNumber="84">Caecognathia</emphasis>
</taxonomicName>
, in the family. According to their key,
<taxonomicName id="94B24D58F36CFFAAFD5FFE650F4EFE2E" box="[752,844,476,500]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="61C6EAC9F36CFFAAFD5FFE650F4EFE2E" box="[752,844,476,500]" italics="true" pageId="5" pageNumber="84">Gnathia</emphasis>
</taxonomicName>
species can be distinguished from the congeners by the presence of a frontal process on the frontal border of the cephalothorax (not deeply excavated frontal border), the presence of paraocular ornamentation, and depth and/or width of dorsal sulcus. On the other hand,
<taxonomicName id="94B24D58F36CFFAAFF38FDF10D36FDBA" box="[151,308,584,608]" class="Malacostraca" family="Gnathiidae" genus="Caecognathia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="61C6EAC9F36CFFAAFF38FDF10D36FDBA" box="[151,308,584,608]" italics="true" pageId="5" pageNumber="84">Caecognathia</emphasis>
</taxonomicName>
species can be distinguished from the congeners by the absence of a frontal process on the frontal border (frontal border often rounded) (
<bibRefCitation id="37234B2AF36CFFAAFDE4FDD50F43FD5E" author="Cohen" box="[587,833,620,644]" pageId="5" pageNumber="84" refString="Cohen, B. F. &amp; Poore, G. C. B. (1994) Phylogeny and biogeography of the Gnathiidae (Crustacea: Isopoda) with descriptions of new genera and species, most from south-eastern Australia. Memoirs of Museum of Victoria, 54 (2), 271 - 397." type="journal article" year="1994">Cohen &amp; Poore 1994</bibRefCitation>
). However, the characters of the frontal “border” and the frontal “process” seem to be ambiguously interpreted by the authors. Some
<taxonomicName id="94B24D58F36CFFAAFBFAFD2908B3FD72" box="[1109,1201,656,680]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="61C6EAC9F36CFFAAFBFAFD2908B3FD72" box="[1109,1201,656,680]" italics="true" pageId="5" pageNumber="84">Gnathia</emphasis>
</taxonomicName>
species have large, rounded frontal “process” (or anteriorly pronounced frontal borders); e.g.
<taxonomicName id="94B24D58F36CFFAAFC4BFD0C0905FD16" authority="Schultz, 1966" authorityName="Schultz" authorityYear="1966" box="[996,1287,692,716]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="84" phylum="Arthropoda" rank="species" species="triobata">
<emphasis id="61C6EAC9F36CFFAAFC4BFD0C085CFD16" box="[996,1118,692,716]" italics="true" pageId="5" pageNumber="84">G. triobata</emphasis>
<bibRefCitation id="37234B2AF36CFFAAFBC7FD0D0905FD16" author="Schultz" box="[1128,1287,692,716]" pageId="5" pageNumber="84" refString="Schultz, G. A. (1966) Submarine canyons of southern Calfornia. Part IV. Systematics: Isopoda. Marine isopods of the submarine canyons of the southern Californian continental shelf. Allan Hancock Pacific Expeditions, 27 (4), 1 - 56." type="journal article" year="1966">Schultz, 1966</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="94B24D58F36CFFAAFABBFD0C0D8DFD2A" authority="Cohen &amp; Poore, 1994" authorityName="Cohen &amp; Poore" authorityYear="1994" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="84" phylum="Arthropoda" rank="species" species="prolasius">
<emphasis id="61C6EAC9F36CFFAAFABBFD0C099EFD16" box="[1300,1436,692,716]" italics="true" pageId="5" pageNumber="84">G. prolasius</emphasis>
<bibRefCitation id="37234B2AF36CFFAAFF38FD610D8DFD2A" author="Cohen" box="[151,399,728,752]" pageId="5" pageNumber="84" refString="Cohen, B. F. &amp; Poore, G. C. B. (1994) Phylogeny and biogeography of the Gnathiidae (Crustacea: Isopoda) with descriptions of new genera and species, most from south-eastern Australia. Memoirs of Museum of Victoria, 54 (2), 271 - 397." type="journal article" year="1994">Cohen &amp; Poore, 1994</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="94B24D58F36CFFAAFE34FD610E9CFD2A" authority="Pires, 1996" authorityName="Pires" authorityYear="1996" box="[411,670,728,752]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="84" phylum="Arthropoda" rank="species" species="ubatuba">
<emphasis id="61C6EAC9F36CFFAAFE34FD610E14FD2A" box="[411,534,728,752]" italics="true" pageId="5" pageNumber="84">G. ubatuba</emphasis>
<bibRefCitation id="37234B2AF36CFFAAFDB2FD610E9CFD2A" author="Pires" box="[541,670,728,752]" pageId="5" pageNumber="84" refString="Pires, A. M. S. (1996) The gnathiids from the Brazilian southeastern continental shelf and slope: distribution, ecological notes and description of three new species. Spixiana, 19 (1), 1 - 16." type="journal article" year="1996">Pires, 1996</bibRefCitation>
</taxonomicName>
;
<emphasis id="61C6EAC9F36CFFAAFD05FD610F22FD2A" box="[682,800,728,752]" italics="true" pageId="5" pageNumber="84">G. r i c a rd o i</emphasis>
<bibRefCitation id="37234B2AF36CFFAAFC84FD610FAEFD2A" author="Pires" box="[811,940,728,752]" pageId="5" pageNumber="84" refString="Pires, A. M. S. (1996) The gnathiids from the Brazilian southeastern continental shelf and slope: distribution, ecological notes and description of three new species. Spixiana, 19 (1), 1 - 16." type="journal article" year="1996">Pires, 1996</bibRefCitation>
. However, these characters has been termed differently by different workers, e.g. “large frontal process” (
<bibRefCitation id="37234B2AF36CFFAAFCE2FD450846FCCE" author="Cohen" box="[845,1092,764,788]" pageId="5" pageNumber="84" refString="Cohen, B. F. &amp; Poore, G. C. B. (1994) Phylogeny and biogeography of the Gnathiidae (Crustacea: Isopoda) with descriptions of new genera and species, most from south-eastern Australia. Memoirs of Museum of Victoria, 54 (2), 271 - 397." type="journal article" year="1994">Cohen &amp; Poore 1994</bibRefCitation>
), “frontal projection” (
<bibRefCitation id="37234B2AF36CFFAAFAE9FD450CD2FCE2" author="Schultz" pageId="5" pageNumber="84" refString="Schultz, G. A. (1966) Submarine canyons of southern Calfornia. Part IV. Systematics: Isopoda. Marine isopods of the submarine canyons of the southern Californian continental shelf. Allan Hancock Pacific Expeditions, 27 (4), 1 - 56." type="journal article" year="1966">Schultz 1966</bibRefCitation>
) or not even noted (
<bibRefCitation id="37234B2AF36CFFAAFE7FFC990E51FCE2" author="Pires" box="[464,595,800,824]" pageId="5" pageNumber="84" refString="Pires, A. M. S. (1996) The gnathiids from the Brazilian southeastern continental shelf and slope: distribution, ecological notes and description of three new species. Spixiana, 19 (1), 1 - 16." type="journal article" year="1996">Pires 1996</bibRefCitation>
). Furthermore, the frontal borders of
<taxonomicName id="94B24D58F36CFFAAFBB6FC9908B4FCE2" box="[1049,1206,800,824]" class="Malacostraca" family="Gnathiidae" genus="Caecognathia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="61C6EAC9F36CFFAAFBB6FC9908B4FCE2" box="[1049,1206,800,824]" italics="true" pageId="5" pageNumber="84">Caecognathia</emphasis>
</taxonomicName>
species are indeed pronounced anteriorly; e.g.
<taxonomicName id="94B24D58F36CFFAAFE74FCFC0F12FC86" authority="Schultz, 1966" authorityName="Schultz" authorityYear="1966" box="[475,784,836,860]" class="Malacostraca" family="Gnathiidae" genus="Caecognathia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="84" phylum="Arthropoda" rank="species" species="hirusta">
<emphasis id="61C6EAC9F36CFFAAFE74FCFC0DECFC86" box="[475,494,837,860]" italics="true" pageId="5" pageNumber="84">C</emphasis>
.
<emphasis id="61C6EAC9F36CFFAAFDAFFCFD0E4DFC86" box="[512,591,836,860]" italics="true" pageId="5" pageNumber="84">hirusta</emphasis>
(
<bibRefCitation id="37234B2AF36CFFAAFDCCFCFD0F05FC86" author="Schultz" box="[611,775,836,860]" pageId="5" pageNumber="84" refString="Schultz, G. A. (1966) Submarine canyons of southern Calfornia. Part IV. Systematics: Isopoda. Marine isopods of the submarine canyons of the southern Californian continental shelf. Allan Hancock Pacific Expeditions, 27 (4), 1 - 56." type="journal article" year="1966">Schultz, 1966</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="94B24D58F36CFFAAFC8FFCFC0851FC86" authority="Hansen, 1916" authorityName="Hansen" authorityYear="1916" box="[800,1107,836,860]" class="Malacostraca" family="Gnathiidae" genus="Caecognathia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="84" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="61C6EAC9F36CFFAAFC8FFCFC0F94FC86" box="[800,918,836,860]" italics="true" pageId="5" pageNumber="84">C. bicolor</emphasis>
(
<bibRefCitation id="37234B2AF36CFFAAFC05FCFD084EFC86" author="Hansen" box="[938,1100,836,860]" pageId="5" pageNumber="84" refString="Hansen, H. J. (1916) Crustacea Malacostraca III. Isopoda. The Danish Ingolf Expedition, 3 (5), 1 - 262." type="journal article" year="1916">Hansen, 1916</bibRefCitation>
)
</taxonomicName>
(redescribed by
<bibRefCitation id="37234B2AF36CFFAAFAB3FCFD0CD1FC5A" author="Svavarsson" pageId="5" pageNumber="84" refString="Svavarsson, J. (1999) The deep water gnathiid Caecognathia bicolor (Hansen, 1916) (Crustacea, Isopoda, Gnathiidae) - redescription and new data on its distribution -. Rit Fiskideildar, 16, 171 - 185." type="journal article" year="1999">Svavarsson 1999</bibRefCitation>
);
<taxonomicName id="94B24D58F36CFFAAFF47FCD10E9DFC5A" authority="Cohen &amp; Poore, 1994" authorityName="Cohen &amp; Poore" authorityYear="1994" box="[232,671,872,896]" class="Malacostraca" family="Gnathiidae" genus="Caecognathia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="84" phylum="Arthropoda" rank="species" species="dolichoderus">
<emphasis id="61C6EAC9F36CFFAAFF47FCD10D9EFC5A" box="[232,412,872,896]" italics="true" pageId="5" pageNumber="84">C. dolichoderus</emphasis>
<bibRefCitation id="37234B2AF36CFFAAFE0BFCD10E9DFC5A" author="Cohen" box="[420,671,872,896]" pageId="5" pageNumber="84" refString="Cohen, B. F. &amp; Poore, G. C. B. (1994) Phylogeny and biogeography of the Gnathiidae (Crustacea: Isopoda) with descriptions of new genera and species, most from south-eastern Australia. Memoirs of Museum of Victoria, 54 (2), 271 - 397." type="journal article" year="1994">Cohen &amp; Poore, 1994</bibRefCitation>
</taxonomicName>
. Future study may provide a better understanding on the systematic position of the genera.
</paragraph>
</subSubSection>
<subSubSection id="1BA86550F36CFFAAFF68FC090970FBEE" pageId="5" pageNumber="84" type="reference_group">
<paragraph id="530D36DBF36CFFAAFF68FC090970FBEE" blockId="5.[151,1436,151,1076]" pageId="5" pageNumber="84">
<taxonomicName id="94B24D58F36CFFAAFF68FC090D93FC12" ID-CoL="3GLBM" box="[199,401,944,968]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="84" phylum="Arthropoda" rank="species" species="excavata" status="sp. nov.">
<emphasis id="61C6EAC9F36CFFAAFF68FC090D93FC12" box="[199,401,944,968]" italics="true" pageId="5" pageNumber="84">Gnathia excavata</emphasis>
</taxonomicName>
<emphasis id="61C6EAC9F36CFFAAFE33FC090DDCFC12" bold="true" box="[412,478,944,968]" pageId="5" pageNumber="84">
<taxonomicNameLabel id="7AF557B2F36CFFAAFE33FC090DDCFC12" box="[412,478,944,968]" pageId="5" pageNumber="84" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is most similar to
<taxonomicName id="94B24D58F36CFFAAFD6EFC09086FFC12" ID-CoL="3GLD7" authority="Cohen &amp; Poore, 1994" authorityName="Cohen &amp; Poore" authorityYear="1994" box="[705,1133,944,968]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="84" phylum="Arthropoda" rank="species" species="notostigma">
<emphasis id="61C6EAC9F36CFFAAFD6EFC090F62FC12" box="[705,864,944,968]" italics="true" pageId="5" pageNumber="84">G. notostigma</emphasis>
<bibRefCitation id="37234B2AF36CFFAAFCC4FC09086FFC12" author="Cohen" box="[875,1133,944,968]" pageId="5" pageNumber="84" refString="Cohen, B. F. &amp; Poore, G. C. B. (1994) Phylogeny and biogeography of the Gnathiidae (Crustacea: Isopoda) with descriptions of new genera and species, most from south-eastern Australia. Memoirs of Museum of Victoria, 54 (2), 271 - 397." type="journal article" year="1994">Cohen &amp; Poore, 1994</bibRefCitation>
</taxonomicName>
, because of the following characters; the mediofrontal process is broad with bifid apex and the whole body is not covered with tubercles (
<bibRefCitation id="37234B2AF36CFFAAFF31FC410D99FBCA" author="Cohen" box="[158,411,1016,1040]" pageId="5" pageNumber="84" refString="Cohen, B. F. &amp; Poore, G. C. B. (1994) Phylogeny and biogeography of the Gnathiidae (Crustacea: Isopoda) with descriptions of new genera and species, most from south-eastern Australia. Memoirs of Museum of Victoria, 54 (2), 271 - 397." type="journal article" year="1994">Cohen &amp; Poore 1994</bibRefCitation>
). However,
<emphasis id="61C6EAC9F36CFFAAFD86FC410EC6FBCA" box="[553,708,1016,1040]" italics="true" pageId="5" pageNumber="84">G. n o t o s t i g m a</emphasis>
has very pronounced paraocular ornamentation forming slight mesolateral ridge (
<bibRefCitation id="37234B2AF36CFFAAFEC8FBA50E5AFBEE" author="Cohen" box="[359,600,1052,1076]" pageId="5" pageNumber="84" refString="Cohen, B. F. &amp; Poore, G. C. B. (1994) Phylogeny and biogeography of the Gnathiidae (Crustacea: Isopoda) with descriptions of new genera and species, most from south-eastern Australia. Memoirs of Museum of Victoria, 54 (2), 271 - 397." type="journal article" year="1994">Cohen &amp; Poore 1994</bibRefCitation>
) rather than three small indistinct tubercles of the present new species.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="4B96CA905A3ED1C0D3FC6930A2DC8353">Gnathiidae from Kumejima Island in the Ryukyu Archipelago, southwestern Japan, with description of three new species (Crustacea: Isopoda) *</mods:title>
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<treatment id="DB1B87CDF36CFFA7FF38FB3D0E1BFAD6" ID-DOI="http://doi.org/10.5281/zenodo.6173344" ID-GBIF-Taxon="119520622" ID-Zenodo-Dep="6173344" LSID="urn:lsid:plazi:treatment:DB1B87CDF36CFFA7FF38FB3D0E1BFAD6" httpUri="http://treatment.plazi.org/id/DB1B87CDF36CFFA7FF38FB3D0E1BFAD6" lastPageId="8" lastPageNumber="87" pageId="5" pageNumber="84">
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<paragraph id="530D36DBF36CFFAAFF38FB3D0D99FB47" blockId="5.[151,657,1155,1250]" box="[151,411,1155,1182]" pageId="5" pageNumber="84">
<heading id="084581B7F36CFFAAFF38FB3D0D99FB47" bold="true" box="[151,411,1155,1182]" fontSize="11" level="1" pageId="5" pageNumber="84" reason="1">
<emphasis id="61C6EAC9F36CFFAAFF38FB3D0D99FB47" bold="true" box="[151,411,1155,1182]" pageId="5" pageNumber="84">
<taxonomicName id="94B24D58F36CFFAAFF38FB3D0D53FB44" ID-CoL="6KNC3" box="[151,337,1156,1182]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="84" phylum="Arthropoda" rank="species" species="scabra" status="sp. nov.">
<emphasis id="61C6EAC9F36CFFAAFF38FB3D0D53FB44" bold="true" box="[151,337,1156,1182]" italics="true" pageId="5" pageNumber="84">Gnathia scabra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AF557B2F36CFFAAFEF6FB3A0D99FB47" box="[345,411,1155,1181]" pageId="5" pageNumber="84" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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<paragraph id="530D36DBF36CFFAAFF38FB1E0D10FB38" blockId="5.[151,657,1155,1250]" pageId="5" pageNumber="84">
New Japanese name: Ibo-eguri-umikuwagata (
<figureCitation id="CB892A5EF36CFFAAFF30FB730CEDFB38" box="[159,239,1226,1250]" captionStart="FIGURE 4" captionStartId="6.[151,250,1940,1963]" captionTargetBox="[186,1404,181,1928]" captionTargetId="figure@6.[183,1407,181,1928]" captionTargetPageId="6" captionText="FIGURE 4. Gnathia scabra n. sp., holotype, adult male (RUMF-ZC- 1429, total length, 2.40 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view; D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209075/files/figure.png" pageId="5" pageNumber="84">Figs. 4</figureCitation>
,
<figureCitation id="CB892A5EF36CFFAAFF55FB730D0AFB38" box="[250,264,1226,1250]" captionStart="FIGURE 5" captionStartId="7.[151,250,1945,1968]" captionTargetBox="[208,1389,187,1932]" captionTargetId="figure@7.[203,1390,187,1932]" captionTargetPageId="7" captionText="FIGURE 5. Gnathia scabra n. sp., holotype, adult male (RUMF-ZC- 1429, total length, 2.40 mm): A, left maxilliped, ventral view; B, left pylopod, ventral view; C, left pereopod II, medial view; D, right pleopod II, ventral view; E, right pleopod III, ventral view, plumose setae were omitted; F, penes, ventral view." httpUri="https://zenodo.org/record/209076/files/figure.png" pageId="5" pageNumber="84">5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1BA86550F36CFFAAFF38FAB2084DFAB6" pageId="5" pageNumber="84" type="materials_examined">
<paragraph id="530D36DBF36CFFAAFF38FAB2084DFAB6" blockId="5.[151,1437,1291,2036]" pageId="5" pageNumber="84">
<emphasis id="61C6EAC9F36CFFAAFF38FAB20D82FAFE" bold="true" box="[151,384,1291,1316]" pageId="5" pageNumber="84">Material examined.</emphasis>
<typeStatus id="8C098879F36CFFAAFE28FAB50DF1FAFE" box="[391,499,1292,1316]" pageId="5" pageNumber="84" type="holotype">Holotype</typeStatus>
. Male,
<quantity id="944A9B3EF36CFFAAFDEBFAB50EAAFAFE" box="[580,680,1292,1316]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="84" unit="mm" value="2.4">2.40 mm</quantity>
in total length (RUMF-ZC-1429), dead coral rubble or rock rubble, Stn. Trawl 45 (beam trawl), 26°19.907ʹN, 126°43.191ʹE – 26°20.056ʹN, 126°42.622ʹE,
<quantity id="944A9B3EF36CFFAAFB0FFA96091CFA9D" box="[1184,1310,1327,1351]" metricMagnitude="1" metricUnit="m" metricValue="7.175" metricValueMax="7.6" metricValueMin="6.75" pageId="5" pageNumber="84" unit="m" value="71.75" valueMax="76.0" valueMin="67.5">67.5–76 m</quantity>
depth, off Kumejima Island, the Ryukyu Archipelago, southwestern
<collectingCountry id="2BA5764BF36CFFAAFC8EFAEC0F66FAB6" box="[801,868,1365,1388]" name="Japan" pageId="5" pageNumber="84">Japan</collectingCountry>
,
<date id="270C101BF36CFFAAFCC0FAED0849FAB6" box="[879,1099,1364,1388]" pageId="5" pageNumber="84" value="2009-11-16">16 November 2009</date>
.
</paragraph>
</subSubSection>
<subSubSection id="1BA86550F36CFFA7FF68FACE0829FBCA" lastPageId="8" lastPageNumber="87" pageId="5" pageNumber="84" type="description">
<paragraph id="530D36DBF36CFFAAFF68FACE0F28FA55" blockId="5.[151,1437,1291,2036]" box="[199,810,1399,1424]" pageId="5" pageNumber="84">
<emphasis id="61C6EAC9F36CFFAAFF68FACE0D5BFA4A" bold="true" box="[199,345,1399,1424]" pageId="5" pageNumber="84">Description.</emphasis>
Male (
<figureCitation id="CB892A5EF36CFFAAFE07FACE0DF5FA55" box="[424,503,1399,1423]" captionStart="FIGURE 4" captionStartId="6.[151,250,1940,1963]" captionTargetBox="[186,1404,181,1928]" captionTargetId="figure@6.[183,1407,181,1928]" captionTargetPageId="6" captionText="FIGURE 4. Gnathia scabra n. sp., holotype, adult male (RUMF-ZC- 1429, total length, 2.40 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view; D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209075/files/figure.png" pageId="5" pageNumber="84">Figs. 4</figureCitation>
,
<figureCitation id="CB892A5EF36CFFAAFDACFACE0E11FA55" box="[515,531,1399,1423]" captionStart="FIGURE 5" captionStartId="7.[151,250,1945,1968]" captionTargetBox="[208,1389,187,1932]" captionTargetId="figure@7.[203,1390,187,1932]" captionTargetPageId="7" captionText="FIGURE 5. Gnathia scabra n. sp., holotype, adult male (RUMF-ZC- 1429, total length, 2.40 mm): A, left maxilliped, ventral view; B, left pylopod, ventral view; C, left pereopod II, medial view; D, right pleopod II, ventral view; E, right pleopod III, ventral view, plumose setae were omitted; F, penes, ventral view." httpUri="https://zenodo.org/record/209076/files/figure.png" pageId="5" pageNumber="84">5</figureCitation>
). Body
<quantity id="944A9B3EF36CFFAAFDC3FACE0ED2FA55" box="[620,720,1399,1423]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="84" unit="mm" value="2.4">2.40 mm</quantity>
(n = 1).
</paragraph>
<paragraph id="530D36DBF36CFFAAFF68FA250DC8F99E" blockId="5.[151,1437,1291,2036]" pageId="5" pageNumber="84">
Cephalothorax (
<figureCitation id="CB892A5EF36CFFAAFED2FA250DC0FA6E" box="[381,450,1436,1460]" captionStart="FIGURE 4" captionStartId="6.[151,250,1940,1963]" captionTargetBox="[186,1404,181,1928]" captionTargetId="figure@6.[183,1407,181,1928]" captionTargetPageId="6" captionText="FIGURE 4. Gnathia scabra n. sp., holotype, adult male (RUMF-ZC- 1429, total length, 2.40 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view; D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209075/files/figure.png" pageId="5" pageNumber="84">Fig. 4</figureCitation>
A–C). Cephaolothorax, covered with setae and tubercles, almost square. Dorsal sulcus deeply concave. Posterior margin slightly concave. Mediofrontal process broad with remarkably concave apex. Superior frontolateral process with 3 setae. Marginal carina slightly visible below antennae in dorsal view. Eyes with 42 ocelli in 7 horizontal rows. Paraocular ornamentation indistinct, composed of several tiny tubercles. Supraocular lobe not acute.
</paragraph>
<paragraph id="530D36DBF36CFFAAFF68F9E80D97F922" blockId="5.[151,1437,1291,2036]" pageId="5" pageNumber="84">
Pereon (
<figureCitation id="CB892A5EF36CFFAAFE87F9F60D6DF9BD" box="[296,367,1615,1639]" captionStart="FIGURE 4" captionStartId="6.[151,250,1940,1963]" captionTargetBox="[186,1404,181,1928]" captionTargetId="figure@6.[183,1407,181,1928]" captionTargetPageId="6" captionText="FIGURE 4. Gnathia scabra n. sp., holotype, adult male (RUMF-ZC- 1429, total length, 2.40 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view; D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209075/files/figure.png" pageId="5" pageNumber="84">Fig. 4</figureCitation>
A). Pereonite I approximately half length of pereonite II, not fused; lateral parts not visible dorsally. Pereonite II and III subequal in length and width; few setae and tubercles on lateral margins. Pereonite IV with anterior constriction; anterolateral lobe absent. Pereonite V with areae laterales. Pereonite VI as long as combined length of pereonite IV and V. Pereonite VII not extending posterolateral margin of pereonite VI, overlapping pleonite I.
</paragraph>
<paragraph id="530D36DBF36CFFAAFF68F8BD0D71F8E5" blockId="5.[151,1437,1291,2036]" pageId="5" pageNumber="84">
Pleon (
<figureCitation id="CB892A5EF36CFFAAFEB4F8BD0D61F8C6" box="[283,355,1796,1820]" captionStart="FIGURE 4" captionStartId="6.[151,250,1940,1963]" captionTargetBox="[186,1404,181,1928]" captionTargetId="figure@6.[183,1407,181,1928]" captionTargetPageId="6" captionText="FIGURE 4. Gnathia scabra n. sp., holotype, adult male (RUMF-ZC- 1429, total length, 2.40 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view; D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209075/files/figure.png" pageId="5" pageNumber="84">Fig. 4</figureCitation>
A). All pleonites sparsely covered with setae. Pleonites II–V subequal in length and width; epimera prominent.
</paragraph>
<paragraph id="530D36DBF36CFFAAFF68F8F50880F85D" blockId="5.[151,1437,1291,2036]" pageId="5" pageNumber="84">
Pleotelson (
<figureCitation id="CB892A5EF36CFFAAFEE3F8F50D93F8BE" box="[332,401,1868,1892]" captionStart="FIGURE 4" captionStartId="6.[151,250,1940,1963]" captionTargetBox="[186,1404,181,1928]" captionTargetId="figure@6.[183,1407,181,1928]" captionTargetPageId="6" captionText="FIGURE 4. Gnathia scabra n. sp., holotype, adult male (RUMF-ZC- 1429, total length, 2.40 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view; D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209075/files/figure.png" pageId="5" pageNumber="84">Fig. 4</figureCitation>
D). Width approximately 1.09 of length. 4 pairs of setae on lateral margin, central part, and apex. Lateral margins slightly concave over proximal third and slightly convex at middle.
</paragraph>
<paragraph id="530D36DBF36CFFAAFF68F82D0E87F82E" blockId="5.[151,1437,1291,2036]" pageId="5" pageNumber="84">
Mandible (
<figureCitation id="CB892A5EF36CFFAAFEE8F82D0D93F876" box="[327,401,1940,1964]" captionStart="FIGURE 4" captionStartId="6.[151,250,1940,1963]" captionTargetBox="[186,1404,181,1928]" captionTargetId="figure@6.[183,1407,181,1928]" captionTargetPageId="6" captionText="FIGURE 4. Gnathia scabra n. sp., holotype, adult male (RUMF-ZC- 1429, total length, 2.40 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view; D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209075/files/figure.png" pageId="5" pageNumber="84">Fig. 4</figureCitation>
B, C). Mandible approximately 0.6 times as long as cephalothorax; apex curved dorsomesially. Mandibular seta present on mid-dorsal surface near incisor. Dentate blade occupying approximately onethird of mandible length. Erisma prominent.
</paragraph>
<caption id="07CD6653F36FFFA9FF38F82D0ED4F835" httpUri="https://zenodo.org/record/209075/files/figure.png" pageId="6" pageNumber="85" targetBox="[186,1404,181,1928]" targetPageId="6">
<paragraph id="530D36DBF36FFFA9FF38F82D0ED4F835" blockId="6.[151,1436,1940,2031]" pageId="6" pageNumber="85">
<emphasis id="61C6EAC9F36FFFA9FF38F82D0D16F870" bold="true" box="[151,276,1940,1963]" pageId="6" pageNumber="85">FIGURE 4.</emphasis>
<taxonomicName id="94B24D58F36FFFA9FEB3F82C0DBEF871" box="[284,444,1941,1963]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="85" phylum="Arthropoda" rank="species" species="scabra" status="sp. nov.">
<emphasis id="61C6EAC9F36FFFA9FEB3F82C0DBEF871" box="[284,444,1941,1963]" italics="true" pageId="6" pageNumber="85">Gnathia scabra</emphasis>
</taxonomicName>
<emphasis id="61C6EAC9F36FFFA9FE6BF82C0DFDF870" bold="true" box="[452,511,1941,1962]" pageId="6" pageNumber="85">
<taxonomicNameLabel id="7AF557B2F36FFFA9FE6BF82C0DFDF870" box="[452,511,1941,1962]" pageId="6" pageNumber="85" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, adult male (RUMF-ZC-1429, total length, 2.40 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view; D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view.
</paragraph>
</caption>
<caption id="07CD6653F36EFFA8FF38F8200F2BF82E" httpUri="https://zenodo.org/record/209076/files/figure.png" pageId="7" pageNumber="86" targetBox="[208,1389,187,1932]" targetPageId="7">
<paragraph id="530D36DBF36EFFA8FF38F8200F2BF82E" blockId="7.[151,1437,1945,2036]" pageId="7" pageNumber="86">
<emphasis id="61C6EAC9F36EFFA8FF38F8200D11F875" bold="true" box="[151,275,1945,1968]" pageId="7" pageNumber="86">FIGURE 5.</emphasis>
<taxonomicName id="94B24D58F36EFFA8FEB5F8230DBBF86A" box="[282,441,1946,1968]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="scabra" status="sp. nov.">
<emphasis id="61C6EAC9F36EFFA8FEB5F8230DBBF86A" box="[282,441,1946,1968]" italics="true" pageId="7" pageNumber="86">Gnathia scabra</emphasis>
</taxonomicName>
<emphasis id="61C6EAC9F36EFFA8FE6FF8230DF8F875" bold="true" box="[448,506,1946,1967]" pageId="7" pageNumber="86">
<taxonomicNameLabel id="7AF557B2F36EFFA8FE6FF8230DF8F875" box="[448,506,1946,1967]" pageId="7" pageNumber="86" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, adult male (RUMF-ZC-1429, total length, 2.40 mm): A, left maxilliped, ventral view; B, left pylopod, ventral view; C, left pereopod II, medial view; D, right pleopod II, ventral view; E, right pleopod III, ventral view, plumose setae were omitted; F, penes, ventral view.
</paragraph>
</caption>
<paragraph id="530D36DBF361FFA7FF68FF2E0DA8FEBE" blockId="8.[151,1437,151,1292]" pageId="8" pageNumber="87">
Antennula (
<figureCitation id="CB892A5EF361FFA7FEE4FF2E0D8CFF75" box="[331,398,151,175]" captionStart="FIGURE 4" captionStartId="6.[151,250,1940,1963]" captionTargetBox="[186,1404,181,1928]" captionTargetId="figure@6.[183,1407,181,1928]" captionTargetPageId="6" captionText="FIGURE 4. Gnathia scabra n. sp., holotype, adult male (RUMF-ZC- 1429, total length, 2.40 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view; D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209075/files/figure.png" pageId="8" pageNumber="87">Fig. 4</figureCitation>
E). 3 basal podomeres and 5 flagellar articles. Basal podomeres I, II, and III bearing 2, 3, and 1 flagellar articles on distal margins, respectively. Flagellar articles I and V bearing 2, 3, 1, 1, and 1 penicillate setae on distal margins, respectively. Articles VI–VIII each with 1 aesthetasc; article VIII bearing 2 terminal setae. Antenna (
<figureCitation id="CB892A5EF361FFA7FEA4FEBD0D56FEC6" box="[267,340,260,284]" captionStart="FIGURE 4" captionStartId="6.[151,250,1940,1963]" captionTargetBox="[186,1404,181,1928]" captionTargetId="figure@6.[183,1407,181,1928]" captionTargetPageId="6" captionText="FIGURE 4. Gnathia scabra n. sp., holotype, adult male (RUMF-ZC- 1429, total length, 2.40 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view; D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209075/files/figure.png" pageId="8" pageNumber="87">Fig. 4</figureCitation>
F). 4 basal podomeres and 7 flagellar articles. Basal podomeres III and IV bearing 2 and 3 penicillate setae on distal margins, respectively. Flagellar articles I–VII with few setae on distal margins; article VII bearing 3 terminal setae.
</paragraph>
<paragraph id="530D36DBF361FFA7FF68FED60896FE76" blockId="8.[151,1437,151,1292]" pageId="8" pageNumber="87">
Maxilliped (
<figureCitation id="CB892A5EF361FFA7FEFDFED60D97FE5D" box="[338,405,367,391]" captionStart="FIGURE 5" captionStartId="7.[151,250,1945,1968]" captionTargetBox="[208,1389,187,1932]" captionTargetId="figure@7.[203,1390,187,1932]" captionTargetPageId="7" captionText="FIGURE 5. Gnathia scabra n. sp., holotype, adult male (RUMF-ZC- 1429, total length, 2.40 mm): A, left maxilliped, ventral view; B, left pylopod, ventral view; C, left pereopod II, medial view; D, right pleopod II, ventral view; E, right pleopod III, ventral view, plumose setae were omitted; F, penes, ventral view." httpUri="https://zenodo.org/record/209076/files/figure.png" pageId="8" pageNumber="87">Fig. 5</figureCitation>
A). Endite reaching distal margin of palp article 1. Palp articles I–IV bearing 3, 7, 4, and 1 plumose setae on external margins, respectively; article IV bearing 7 simple terminal setae.
</paragraph>
<paragraph id="530D36DBF361FFA7FF68FE0E0818FDCD" blockId="8.[151,1437,151,1292]" pageId="8" pageNumber="87">
Pylopod (
<figureCitation id="CB892A5EF361FFA7FE99FE0E0D79FE15" box="[310,379,439,463]" captionStart="FIGURE 5" captionStartId="7.[151,250,1945,1968]" captionTargetBox="[208,1389,187,1932]" captionTargetId="figure@7.[203,1390,187,1932]" captionTargetPageId="7" captionText="FIGURE 5. Gnathia scabra n. sp., holotype, adult male (RUMF-ZC- 1429, total length, 2.40 mm): A, left maxilliped, ventral view; B, left pylopod, ventral view; C, left pereopod II, medial view; D, right pleopod II, ventral view; E, right pleopod III, ventral view, plumose setae were omitted; F, penes, ventral view." httpUri="https://zenodo.org/record/209076/files/figure.png" pageId="8" pageNumber="87">Fig. 5</figureCitation>
B). 3-articulate; 3 setae near distal margin of article I and II, respectively. Article I elliptical with 3 areolae bearing 25 plumose setae on internal margin; 1 seta near external margin and distal margin, respectively. Article II elliptical, fringed with fine setae. Article III semicircular.
</paragraph>
<paragraph id="530D36DBF361FFA7FF68FD9D0FD4FD2A" blockId="8.[151,1437,151,1292]" pageId="8" pageNumber="87">
Pereopod II (
<figureCitation id="CB892A5EF361FFA7FEC5FD9D0DB7FDE6" box="[362,437,548,572]" captionStart="FIGURE 5" captionStartId="7.[151,250,1945,1968]" captionTargetBox="[208,1389,187,1932]" captionTargetId="figure@7.[203,1390,187,1932]" captionTargetPageId="7" captionText="FIGURE 5. Gnathia scabra n. sp., holotype, adult male (RUMF-ZC- 1429, total length, 2.40 mm): A, left maxilliped, ventral view; B, left pylopod, ventral view; C, left pereopod II, medial view; D, right pleopod II, ventral view; E, right pleopod III, ventral view, plumose setae were omitted; F, penes, ventral view." httpUri="https://zenodo.org/record/209076/files/figure.png" pageId="8" pageNumber="87">Fig. 5</figureCitation>
C). Basis oblong, outer margin with 3 penicillate setae and 4 processes. Ischium approximately two-third length of basis, becoming wider distally; fine setae bearing inner and outer margins and 2 long setae near distal margin. Merus approximately two-third length of ischium, bearing 4 setae on distal corner. Inner margins of carpus and propodus covered with pectinate scales. Carpus slightly shorter than merus, bearing 1 spine and 2 processes on inner margin. Propodus rectangular and 1.3 times as long as carpus; inner-middle and inner-distal margins with 2 spines and distal corner with 1 penicillate seta.
</paragraph>
<paragraph id="530D36DBF361FFA7FF68FD450811FC5A" blockId="8.[151,1437,151,1292]" pageId="8" pageNumber="87">
Pleopod II (
<figureCitation id="CB892A5EF361FFA7FEFEFD450D94FCCE" box="[337,406,764,788]" captionStart="FIGURE 5" captionStartId="7.[151,250,1945,1968]" captionTargetBox="[208,1389,187,1932]" captionTargetId="figure@7.[203,1390,187,1932]" captionTargetPageId="7" captionText="FIGURE 5. Gnathia scabra n. sp., holotype, adult male (RUMF-ZC- 1429, total length, 2.40 mm): A, left maxilliped, ventral view; B, left pylopod, ventral view; C, left pereopod II, medial view; D, right pleopod II, ventral view; E, right pleopod III, ventral view, plumose setae were omitted; F, penes, ventral view." httpUri="https://zenodo.org/record/209076/files/figure.png" pageId="8" pageNumber="87">Fig. 5</figureCitation>
D, E). Protopod distomedial corner with coupling hook and 1 seta. Both rami elliptic but exopod slightly shorter than endopod; 9 and 7 plumose setae on endopod and exopod, respectively. Appendix masculina reaching half-length of endopod. Both rami of pleopods IV and V shorter than those of pleopods I–III. Endopods each with 7 or 8 simple setae. Exopods each with 8–9 plumose setae.
</paragraph>
<paragraph id="530D36DBF361FFA7FF68FC350E4AFC36" blockId="8.[151,1437,151,1292]" pageId="8" pageNumber="87">
Uropod (
<figureCitation id="CB892A5EF361FFA7FE9EFC350D78FC7E" box="[305,378,908,932]" captionStart="FIGURE 4" captionStartId="6.[151,250,1940,1963]" captionTargetBox="[186,1404,181,1928]" captionTargetId="figure@6.[183,1407,181,1928]" captionTargetPageId="6" captionText="FIGURE 4. Gnathia scabra n. sp., holotype, adult male (RUMF-ZC- 1429, total length, 2.40 mm): A, body, dorsal view; B, eye and frontal border, dorsal view; C, cephalothorax and pereonite I, lateral view; D, pleotelson and uropods, dorsal view; E, left antennula, medial view; F, left antenna, medial view." httpUri="https://zenodo.org/record/209075/files/figure.png" pageId="8" pageNumber="87">Fig. 4</figureCitation>
D). Both rami subequal in length; endopod extending beyond apex of pleotelson. Exopod bearing 7 setae and 3 plumose setae laterally. Endopod bearing 6 setae and 3 plumose setae laterally. Dorsal surface of endopod bearing 5 penicillate setae.
</paragraph>
<paragraph id="530D36DBF361FFA7FF68FC400829FBCA" blockId="8.[151,1437,151,1292]" box="[199,1067,1016,1040]" pageId="8" pageNumber="87">
Penes (
<figureCitation id="CB892A5EF361FFA7FEB8FC410D58FBCA" box="[279,346,1016,1040]" captionStart="FIGURE 5" captionStartId="7.[151,250,1945,1968]" captionTargetBox="[208,1389,187,1932]" captionTargetId="figure@7.[203,1390,187,1932]" captionTargetPageId="7" captionText="FIGURE 5. Gnathia scabra n. sp., holotype, adult male (RUMF-ZC- 1429, total length, 2.40 mm): A, left maxilliped, ventral view; B, left pylopod, ventral view; C, left pereopod II, medial view; D, right pleopod II, ventral view; E, right pleopod III, ventral view, plumose setae were omitted; F, penes, ventral view." httpUri="https://zenodo.org/record/209076/files/figure.png" pageId="8" pageNumber="87">Fig. 5</figureCitation>
F). Penes composed of 2 contiguous papillae and not prominent.
</paragraph>
</subSubSection>
<subSubSection id="1BA86550F361FFA7FF68FBA50D8EFB82" pageId="8" pageNumber="87" type="etymology">
<paragraph id="530D36DBF361FFA7FF68FBA50D8EFB82" blockId="8.[151,1437,151,1292]" pageId="8" pageNumber="87">
<emphasis id="61C6EAC9F361FFA7FF68FBA50D4CFBEE" bold="true" box="[199,334,1052,1076]" pageId="8" pageNumber="87">Etymology.</emphasis>
The scientific name
<taxonomicName id="94B24D58F361FFA7FDEAFBA50E93FBEE" box="[581,657,1052,1076]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="87" phylum="Arthropoda" rank="species" species="scabra">
<emphasis id="61C6EAC9F361FFA7FDEAFBA50E93FBEE" box="[581,657,1052,1076]" italics="true" pageId="8" pageNumber="87">scabra</emphasis>
</taxonomicName>
is derived from the Latin meaning “rugged”, referring to its many tubercles on the head.
</paragraph>
</subSubSection>
<subSubSection id="1BA86550F361FFA7FF68FBDD0E1BFAD6" pageId="8" pageNumber="87" type="discussion">
<paragraph id="530D36DBF361FFA7FF68FBDD0E1BFAD6" blockId="8.[151,1437,151,1292]" pageId="8" pageNumber="87">
<emphasis id="61C6EAC9F361FFA7FF68FBDD0D39FBA6" bold="true" box="[199,315,1124,1148]" pageId="8" pageNumber="87">Remarks.</emphasis>
<taxonomicName id="94B24D58F361FFA7FEECFBDD0DF1FBA6" box="[323,499,1124,1148]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="87" phylum="Arthropoda" rank="species" species="scabra" status="sp. nov.">
<emphasis id="61C6EAC9F361FFA7FEECFBDD0DF1FBA6" box="[323,499,1124,1148]" italics="true" pageId="8" pageNumber="87">Gnathia scabra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7AF557B2F361FFA7FE55FBDD0E38FBA6" box="[506,570,1124,1148]" pageId="8" pageNumber="87" rank="species">
<emphasis id="61C6EAC9F361FFA7FE55FBDD0E38FBA6" bold="true" box="[506,570,1124,1148]" pageId="8" pageNumber="87">n. sp.</emphasis>
</taxonomicNameLabel>
is most similar to
<taxonomicName id="94B24D58F361FFA7FCA1FBDC083CFBA6" authority="Muller, 1993" authorityName="Muller" authorityYear="1993" box="[782,1086,1124,1148]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="87" phylum="Arthropoda" rank="species" species="lignophila">
<emphasis id="61C6EAC9F361FFA7FCA1FBDC0FA3FBA6" box="[782,929,1124,1148]" italics="true" pageId="8" pageNumber="87">G. lignophila</emphasis>
<bibRefCitation id="37234B2AF361FFA7FC06FBDD083CFBA6" author="Muller" box="[937,1086,1124,1148]" pageId="8" pageNumber="87" refString="Muller, H. - G. (1993) Gnathiidae from coral reefs in the Tioman Archipelago, Malaysia, with description of two new species (Crustacea: Isopoda: Cymothoidea). Mitteilungen aus dem Zoologishen Museum in Berlin, 69, 3 - 17." type="journal article" year="1993">Müller, 1993</bibRefCitation>
</taxonomicName>
, in following characters: many tubercles present on chephalothorax and pereonite I–III, and the apex of mediofrontal process is remarkably concave (
<bibRefCitation id="37234B2AF361FFA7FEA6FB150DA3FB1E" author="Muller" box="[265,417,1196,1220]" pageId="8" pageNumber="87" refString="Muller, H. - G. (1993) Gnathiidae from coral reefs in the Tioman Archipelago, Malaysia, with description of two new species (Crustacea: Isopoda: Cymothoidea). Mitteilungen aus dem Zoologishen Museum in Berlin, 69, 3 - 17." type="journal article" year="1993">Müller 1993</bibRefCitation>
). This new species, however, has appendix masculina on pleopod II (it lacks in
<taxonomicName id="94B24D58F361FFA7FA2BFB140D08FB32" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="87" phylum="Arthropoda" rank="species" species="lignophila">
<emphasis id="61C6EAC9F361FFA7FA2BFB140D08FB32" italics="true" pageId="8" pageNumber="87">G. lignophila</emphasis>
</taxonomicName>
), longer mediofrontal process than frontolateral process (mediofrontal border of
<taxonomicName id="94B24D58F361FFA7FB35FB68092EFB32" box="[1178,1324,1232,1256]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="87" phylum="Arthropoda" rank="species" species="lignophila">
<emphasis id="61C6EAC9F361FFA7FB35FB68092EFB32" box="[1178,1324,1232,1256]" italics="true" pageId="8" pageNumber="87">G. lignophila</emphasis>
</taxonomicName>
is always shorter than frontolateral process).
</paragraph>
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<taxonomicName authority="H. E. Ahles" class="Magnoliopsida" family="Ranunculaceae" genus="Thalictrum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thalictrum cooleyi" order="Ranunculales" pageId="0" pageNumber="1099" phylum="Tracheophyta" rank="species" species="cooleyi">Thalictrum cooleyi H.E. Ahles</taxonomicName>
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<paragraph pageId="0" pageNumber="1099">Ecological interactions</paragraph>
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<paragraph pageId="0" pageNumber="1099">Conservation status</paragraph>
<paragraph pageId="0" pageNumber="1099">State E, Fed E; S2, G2.</paragraph>
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<paragraph pageId="0" pageNumber="1099">Distribution</paragraph>
<paragraph pageId="0" pageNumber="1099">Wet pine savannas (VWLPS).</paragraph>
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Infrequent. Late
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Jul;
<normalizedToken originalValue="Aug–Oct">Aug-Oct</normalizedToken>
. LeBlond 474 (stored in personal collection of the collector), Sorrie 9502 (NCU!). Specimens seen in the vicinity: Sandy Run: Ahles 58369 (NCU!), Gardner s.n. (NCU!), Taggart SARU 193 (WNC!). [= RAB, FNA, Weakley]
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<treatment id="DB1BD106DF5E919FEA2F1053E54C42BB" ID-GBIF-Taxon="231544869" LSID="urn:lsid:plazi:treatment:DB1BD106DF5E919FEA2F1053E54C42BB" httpUri="http://treatment.plazi.org/id/DB1BD106DF5E919FEA2F1053E54C42BB" lastPageNumber="235" pageNumber="235" sourceUrl="http://www.departments.bucknell.edu/biology/resources/msw3/browse.asp?id=13700095">
<subSubSection id="E915C73B2BF8C1EE55DEF7101332999F" pageNumber="235" type="nomenclature">
<paragraph id="22C075C5BE875A69BC66917CAE29FB6C" pageNumber="235">
<taxonomicName id="1B892D6C9307E3BDB88A36E18D6692DF" ID-CoL="ZK56" authority="Robinson and Kloss 1922" authorityName="Robinson and Kloss" authorityYear="1922" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageNumber="235" phylum="Chordata" rank="species" species="indochinensis">
<emphasis id="F34B08C2FC03C98089142AA72F4D65A8" italics="true" pageNumber="235">Crocidura indochinensis</emphasis>
Robinson and Kloss 1922
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="21849443F99D989DE223799A3ABF327F" pageNumber="235" type="reference_group">
<paragraph id="9435366202AC2D3F4376EAEEA631EA7D" pageNumber="235">
<treatmentCitation id="A3E014218226A7516C38E09E4790B7A9" author="Robinson and Kloss" authority="Robinson and Kloss 1922" authorityName="Robinson and Kloss" authorityYear="1922" class="Mammalia" family="Soricidae" genus="Crocidura" journal="Ann. Mag. Nat. Hist., ser. 9" kingdom="Animalia" order="Soricomorpha" page="88" pageNumber="235" phylum="Chordata" rank="species" species="indochinensis" title="Crocidura indochinensis" volume="9" year="1922">
<taxonomicName id="01CA896A6121C4E920B90D0283542CE4" ID-CoL="ZK56" authority="Robinson and Kloss 1922" authorityName="Robinson and Kloss" authorityYear="1922" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageNumber="235" phylum="Chordata" rank="species" species="indochinensis">
<emphasis id="89074435664075B76C94F60A3390D4C7" italics="true" pageNumber="235">Crocidura indochinensis</emphasis>
<bibRefCitation id="D04750F614F3F31329FF235CF957A388" author="Robinson, H. C. &amp; C. B. Kloss" refId="ref307485" refString="Robinson, H. C., and C. B. Kloss. 1922. New mammals from French Indo-China and Siam. Annals and Magazine of Natural History, ser. 9, 9: 87 - 99." year="1922">Robinson and Kloss 1922</bibRefCitation>
</taxonomicName>
,
<bibCitation id="6CBC7E24E57DA4EB6D54842972302758" author="Robinson and Kloss" journal="Ann. Mag. Nat. Hist., ser. 9" pageNumber="235" pagination="88" title="Crocidura indochinensis" volume="9" year="1922">Ann. Mag. Nat. Hist., ser. 9, 9: 88</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2FB5D3DF38C6D559A293969CDDDBBB6C" pageNumber="235" type="materials_examined">
<paragraph id="3606FFF56A84914B6DBF25AAB7C09B4E" pageNumber="235">
<emphasis id="D17EE47BC9D329246F3204AB5CD70454" inLineHeading="true" pageNumber="235">Type Locality:</emphasis>
<materialsCitation id="E9FACCE2E83FAB075AE34FC8104832E3" ID-GBIF-Occurrence="4867338777" country="Vietnam" pageNumber="235">
<collectingCountry id="55754F0F289D660FCAAA7B5DED474591" name="Vietnam">Vietnam</collectingCountry>
, Langbian Plateau, Dalat.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="62EAABB944F1E13FCE884B71FBE23B2B" pageNumber="235" type="vernacular_names">
<paragraph id="29A8CD94F813ADA5FBE85730C5A62E78" pageNumber="235">
<emphasis id="F859B57E6BA6E581DE40450637992E2C" inLineHeading="true" pageNumber="235">Vernacular Names:</emphasis>
<vernacularName id="C1853AA7F11C0698DDFABD7D1D7156C8" language="eng" pageNumber="235">Indochinese Shrew</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="68A98A28CD60BB6E3A1C5811A9D09E8F" pageNumber="235" type="distribution">
<paragraph id="E4526317272D29EA4B2BB6A76B1CDCA7" pageNumber="235">
<emphasis id="26A2BE511DCF74D7B773422D16272ED2" inLineHeading="true" pageNumber="235">Distribution:</emphasis>
<collectingCountry id="8810BAF4B80AFFE5369F7BEFB6E498A4" name="Myanmar">Burma</collectingCountry>
, N
<collectingCountry id="537846278F8038C2D2C5B402175FB255" name="Thailand">Thailand</collectingCountry>
to
<collectingCountry id="E7094E7BDFA927FC3487DEA8265B1680" name="Vietnam">Vietnam</collectingCountry>
;
<collectingRegion id="1E01A9CD71848D9507E81F616878E28C" country="China" name="Yunnan">Yunnan</collectingRegion>
and
<collectingRegion id="1FD5B6FFAE1D0C25F3832867F4D97F1D" country="China" name="Fujian">Fujian</collectingRegion>
(
<collectingCountry id="28A019B79858A41D5B4AC30AA77DF9A6" name="China">China</collectingCountry>
); exact limits unknown.
</paragraph>
</subSubSection>
<subSubSection id="3144169630BCAE05FCD91D9E333DDD0E" pageNumber="235" type="discussion">
<paragraph id="8A23B1010A9CB3FD48BE83FF0843ABCD" pageNumber="235">
<emphasis id="DE7B98DAB4A91B8272950D1BF0FC467E" inLineHeading="true" pageNumber="235">Discussion:</emphasis>
This species is usually included in
<taxonomicName id="1B76E269A74C8A93160CEA8BA93D426D" authorityName="Tomes" authorityYear="1856" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" phylum="Chordata" rank="species" species="horsfieldii">
<emphasis id="02147D573187525812224405B4AB0DA3" italics="true" pageNumber="235">horsfieldii</emphasis>
</taxonomicName>
(
<bibRefCitation id="91FB40C456944C800DD040DB94FC24E9" author="Corbet, G. B. &amp; J. E. Hill" refId="ref67955" refString="Corbet, G. B., and J. E. Hill. 1992. Mammals of the Indomalayan region. A systematic review. Oxford University Press, Oxford, 488 pp." year="1992">Corbet and Hill, 1992</bibRefCitation>
;
<bibRefCitation id="78B2C56C400AD57376D8C7570E10C54D" author="Jameson, E. W. &amp; G. S. Jones" refId="ref171786" refString="Jameson, E. W., and G. S. Jones. 1977. The Soricidae of Taiwan. Proceedings of the Biological Society of Washington, 90: 459 - 482." year="1977">Jameson and Jones, 1977</bibRefCitation>
;
<bibRefCitation id="631F1080A69EB4092B41F690E959C361" author="Jenkins, P. D." refId="ref172696" refString="Jenkins, P. D. 1976. Variation in Eurasian shrews of the genus Crocidura (Insectivora: Soricidae). Bulletin of the British Museum (Natural History), Zoology Series, 30: 271 - 309." year="1976">Jenkins, 1976</bibRefCitation>
;
<bibRefCitation id="283949FC5B2E2E119BC7DB7959842D73" author="Jiang, X. L. &amp; R. S. Hoffmann" refId="ref174024" refString="Jiang, X. L., and R. S. Hoffmann. 2001. A revision of the white-toothed shrews (Crocidura) of southern China. Journal of Mammalogy, 82: 1059 - 1079." year="2001">Jiang and Hoffmann, 2001</bibRefCitation>
), but the restriction of
<taxonomicName id="7B90121E80E7CB8025C1CA88D1CFB199" authorityName="Tomes" authorityYear="1856" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" phylum="Chordata" rank="species" species="horsfieldii">
<emphasis id="A0B442DB80F19E16169591AC4E9C1CB6" italics="true" pageNumber="235">horsfieldii</emphasis>
</taxonomicName>
to
<collectingCountry id="64DCCEF8FE992B2148DBFE2EAA499100" name="Sri Lanka">Sri Lanka</collectingCountry>
and
<collectingCountry id="DF5D5179DA9475FFA65E87130CB94917" name="India">India</collectingCountry>
and the removal of
<emphasis id="E820E4A317E776E8F7FCBD140D4CCD80" italics="true" pageNumber="235">wuchinensis</emphasis>
leaves
<taxonomicName id="439E8E0B99B85C253F3D8678A3CEA0CE" authorityName="Robinson and Kloss" authorityYear="1922" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" phylum="Chordata" rank="species" species="indochinensis">
<emphasis id="29403710EB3A039198A329E9E23A57C0" italics="true" pageNumber="235">indochinensis</emphasis>
</taxonomicName>
as the available name for the populations from
<collectingCountry id="43063071F8938C4924E3716FAFFF094A" name="Myanmar">Burma</collectingCountry>
to
<collectingCountry id="22A5EAF802D049DC3D1AF29BDD3C5BD2" name="China">China</collectingCountry>
. Heaney and Timm (1983) reported this species from
<collectingCountry id="B283FA0C0BFCCDD22DDAD3758E34EB23" name="Vietnam">Vietnam</collectingCountry>
under
<taxonomicName id="81C2F936527DF89C060C9EFCE062208B" authorityName="Tomes" authorityYear="1856" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" phylum="Chordata" rank="species" species="horsfieldii">
<emphasis id="EF2C35F54490291B307A8D8E3AA06D31" italics="true" pageNumber="235">C. horsfieldii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

101
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1134.93999" ID-Pensoft-Pub="1313-2970-1134-171" ID-Pensoft-UUID="EDDE25C0A46950BABBBF4C2E6C4EB9C6" ID-ZooBank="0DCD69C86FF846618AB4E31BAF26F6D7" ModsDocID="1313-2970-1134-171" checkinTime="1670676970680" checkinUser="pensoft" docAuthor="Wichard, Wilfried &amp; Xu, Chunpeng" docDate="2022" docId="DB1BD3E50E2658BF8961ADD0330BECE3" docLanguage="en" docName="ZooKeys 1134: 171-183" docOrigin="ZooKeys 1134" docPubDate="2022-12-09" docSource="http://dx.doi.org/10.3897/zookeys.1134.93999" docTitle="Neureclipsis triangula Wichard &amp; Xu 2022, sp. nov." docType="treatment" docUuid="8F917EDA-F2C2-4193-9AA2-C185281408EE" docUuidSource="ZooBank" docVersion="1" id="EDDE25C0A46950BABBBF4C2E6C4EB9C6" lastPageNumber="171" masterDocId="EDDE25C0A46950BABBBF4C2E6C4EB9C6" masterDocTitle="The family Polycentropodidae (Insecta, Trichoptera) in mid-Cretaceous Burmese Amber" masterLastPageNumber="183" masterPageNumber="171" pageNumber="171" updateTime="1670676970680" updateUser="pensoft">
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<mods:title>The family Polycentropodidae (Insecta, Trichoptera) in mid-Cretaceous Burmese Amber</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wichard, Wilfried</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5933-6292</mods:nameIdentifier>
<mods:affiliation>Universitaet zu Koeln, Institute of Biology and its Didactics, Herbert Lewinstrasse 2, 50931 Cologne, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Xu, Chunpeng</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4991-4279</mods:nameIdentifier>
<mods:affiliation>State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China &amp; University of Chinese Academy of Sciences, Beijing 100049, China</mods:affiliation>
<mods:nameIdentifier type="email">cpxu@nigpas.ac.cn</mods:nameIdentifier>
</mods:name>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-12-09</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1134</mods:number>
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<mods:start>171</mods:start>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1134.93999</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1134.93999</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1134-171</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:8F917EDA-F2C2-4193-9AA2-C185281408EE" httpUri="http://treatment.plazi.org/id/DB1BD3E50E2658BF8961ADD0330BECE3" lastPageNumber="171" pageId="0" pageNumber="171">
<subSubSection pageId="0" pageNumber="171" type="nomenclature">
<paragraph pageId="0" pageNumber="171">
<taxonomicName LSID="https://zoobank.org/8F917EDA-F2C2-4193-9AA2-C185281408EE" authority="Wichard &amp; Xu, 2022" authorityName="Wichard &amp; Xu" authorityYear="2022" class="Insecta" family="Polycentropodidae" genus="Neureclipsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neureclipsis triangula" order="Trichoptera" pageId="0" pageNumber="171" phylum="Arthropoda" rank="species" species="triangula" status="sp. nov.">Neureclipsis triangula</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="171">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="171" type="description">
<paragraph pageId="0" pageNumber="171">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Neureclipsis triangula sp. nov. A male holotype (NIGP 200021) habitus, ventral view B inferior appendages, ventral view C drawing of inferior appendages, ventral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1134.93999.figure1" httpUri="https://binary.pensoft.net/fig/780067" pageId="0" pageNumber="171">Fig. 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="171" type="diagnosis">
<paragraph pageId="0" pageNumber="171">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="171">
The extinct species
<taxonomicName authorityName="Wichard &amp; Xu" authorityYear="2022" class="Insecta" family="Polycentropodidae" genus="Neureclipsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neureclipsis triangula" order="Trichoptera" pageId="0" pageNumber="171" phylum="Arthropoda" rank="species" species="triangula">
<emphasis italics="true" pageId="0" pageNumber="171">Neureclipsis triangula</emphasis>
</taxonomicName>
sp. nov. is characterized by a pair of slightly cupped inferior appendages running parallel in ventral view. Each appendage is tapered at the base, wider near the middle and apically forming a sub-triangular shape. Its sloping apical edge is clearly subapically toothed and highlighted in black.
</paragraph>
<caption doi="10.3897/zookeys.1134.93999.figure1" httpUri="https://binary.pensoft.net/fig/780067" pageId="0" pageNumber="171" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="171">
<emphasis bold="true" pageId="0" pageNumber="171">Figure 1.</emphasis>
<taxonomicName authorityName="Wichard &amp; Xu" authorityYear="2022" class="Insecta" family="Polycentropodidae" genus="Neureclipsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neureclipsis triangula" order="Trichoptera" pageId="0" pageNumber="171" phylum="Arthropoda" rank="species" species="triangula">
<emphasis italics="true" pageId="0" pageNumber="171">Neureclipsis triangula</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="171">A</emphasis>
male holotype (NIGP200021) habitus, ventral view
<emphasis bold="true" pageId="0" pageNumber="171">B</emphasis>
inferior appendages, ventral view
<emphasis bold="true" pageId="0" pageNumber="171">C</emphasis>
drawing of inferior appendages, ventral view. Scale bar: 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="171" type="etymology">
<paragraph pageId="0" pageNumber="171">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="171">Species named after the inferior appendages, subtriangular (Latin adjective = triangulus, -a, -um).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="171" type="holotype">
<paragraph pageId="0" pageNumber="171">Holotype.</paragraph>
<paragraph pageId="0" pageNumber="171">♂; Myanmar, Kachin State, Hukawng Valley; exact locality unknown; Mid-Cretaceous Burmese amber inclusion; deposited in the amber collection of the NIGP; NIGP200021.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="171" type="description">
<paragraph pageId="0" pageNumber="171">Description.</paragraph>
<paragraph pageId="0" pageNumber="171">Genus as described above. Body well preserved and visible in ventral and dorsal views. Forewing length about 4.2 mm, oblong, rounded, light brown. Antennae about two-thirds as long as forewings, with about 36 flagellomeres plus scapus and short pedicellus. Inferior appendages having subtriangular shape, with oblique, subapically toothed, and black terminal margin.</paragraph>
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