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<mods:title id="3B2877474BC3FF7D97E779C4C17AA40C">Macropsychanthus nitidus: a new rank and combination for dioclea rostrata var. nitida (leguminosae, papilionoideae)</mods:title>
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<mods:namePart id="96C1A6572805CA80D4CD73754BD6E805">Fonseca-Cortés, Andrés</mods:namePart>
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<mods:namePart id="56FAF574C29778982309B4ACDE4FC77B">Snak, Cristiane</mods:namePart>
<mods:affiliation id="E38AEFB6C5CDF90CE3446509A944AC9C">Departamento de Engenharia de Pesca e Ciências Biológicas, Universidade do Estado de Santa Catarina, Rua Cel. Fernandes Martins 270, Progresso, 88790 - 000, Laguna, Santa Catarina, Brazil</mods:affiliation>
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<emphasis id="B978EAADFFFCFFF3FF3FD3E98A06FAB2" box="[136,348,1268,1294]" italics="true" pageId="0" pageNumber="293">Macropsychanthus</emphasis>
Harms ex Schumann &amp; Lauterbach (1900: 366)
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<paragraph id="8BB336BFFFFCFFF3FC2AD3E9885BF8BB" blockId="0.[136,1452,1268,2051]" pageId="0" pageNumber="293">
originally comprised four species confined to Southeastern Asia to New
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(Adema, 1998). Recently,
<bibRefCitation id="EF9D4B4EFFFCFFF3FC8AD2058F10FA8E" author="Queiroz, L. P. &amp; Snak, C." box="[829,1098,1304,1330]" pageId="0" pageNumber="293" pagination="67 - 114" refId="ref3578" refString="Queiroz, L. P. &amp; Snak, C. (2020) Revisiting the taxonomy of Dioclea and related genera (Leguminosae, Papilionoideae), with new generic circumscriptions. PhytoKeys 164 (1): 67 - 114. https: // doi. org / 10.3897 / phytokeys. 164.55441" type="journal article" year="2020">Queiroz &amp; Snak (2020)</bibRefCitation>
has demonstrated that
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<emphasis id="B978EAADFFFCFFF3FAE4D2058EF6FA8E" box="[1363,1452,1304,1330]" italics="true" pageId="0" pageNumber="293">Dioclea</emphasis>
Kunth (1824: 437)
</taxonomicName>
is a non-monophyletic genus, with its species falling into three different positions: (1) the puzzling species,
<taxonomicName id="4C0C4D3CFFFCFFF3FF5FD27D883FFAC6" authority="Killip ex R. H. Maxwell (1978: 243)" authorityName="R. H. Maxwell" authorityPageNumber="243" authorityYear="1978" box="[232,869,1376,1403]" class="Magnoliopsida" family="Fabaceae" genus="Dioclea" kingdom="Plantae" order="Fabales" pageId="0" pageNumber="293" phylum="Tracheophyta" rank="species" species="paniculata">
<emphasis id="B978EAADFFFCFFF3FF5FD27D8A9BFAC6" box="[232,449,1376,1402]" italics="true" pageId="0" pageNumber="293">Dioclea paniculata</emphasis>
Killip ex R.H.
<bibRefCitation id="EF9D4B4EFFFCFFF3FDC5D27D883FFAC6" author="Maxwell, R. H." box="[626,869,1376,1402]" pageId="0" pageNumber="293" pagination="243 - 252" refId="ref3534" refString="Maxwell, R. H. (1978) Problems in the placement of Dioclea paniculata Killip MS. (Leguminosae). Phytologia 40: 243 - 252. https: // doi. org / 10.5962 / bhl. part. 28268" type="journal article" year="1978">Maxwell (1978: 243)</bibRefCitation>
</taxonomicName>
, was highly supported as a member of the Galactia clade; (2)
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<emphasis id="B978EAADFFFCFFF3FEB4D2998A06FA22" box="[259,348,1412,1438]" italics="true" pageId="0" pageNumber="293">Dioclea</emphasis>
subg.
<emphasis id="B978EAADFFFCFFF3FE07D2998953FA22" box="[432,521,1412,1438]" italics="true" pageId="0" pageNumber="293">Dioclea</emphasis>
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appeared as sister to a clade composed of
<taxonomicName id="4C0C4D3CFFFCFFF3FBA6D2998E2DFA22" authority="Bentham (1837: 67)" authorityName="Bentham" authorityPageNumber="67" authorityYear="1837" box="[1041,1399,1412,1438]" class="Magnoliopsida" family="Fabaceae" genus="Cleobulia" kingdom="Plantae" order="Fabales" pageId="0" pageNumber="293" phylum="Tracheophyta" rank="genus">
<emphasis id="B978EAADFFFCFFF3FBA6D2998F25FA22" box="[1041,1151,1412,1438]" italics="true" pageId="0" pageNumber="293">Cleobulia</emphasis>
<bibRefCitation id="EF9D4B4EFFFCFFF3FB3BD2998E2DFA22" author="Bentham, G." box="[1164,1399,1412,1438]" pageId="0" pageNumber="293" refId="ref3219" refString="Bentham, G. (1837) Commentationes de Leguminosarum Generibus. Zollinger, Vienna, 488 pp." type="book" year="1837">Bentham (1837: 67)</bibRefCitation>
</taxonomicName>
and
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<emphasis id="B978EAADFFFCFFF3FF3FD2B58A4BFA7E" box="[136,273,1448,1474]" italics="true" pageId="0" pageNumber="293">Cymbosema</emphasis>
<bibRefCitation id="EF9D4B4EFFFCFFF3FEAFD2B58AA1FA7E" author="Bentham, G." box="[280,507,1448,1474]" pageId="0" pageNumber="293" pagination="38 - 103" refId="ref3238" refString="Bentham, G. (1840) Contributions towards a Flora of South America. - Enumeration of Plants collected by Mr. Schomburgk in British Guiana. Le Journal de Botanique 2: 38 - 103. https: // doi. org / 10.1080 / 00222933909512423" type="journal article" year="1840">Bentham (1840: 61)</bibRefCitation>
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; and (3) the species of
<taxonomicName id="4C0C4D3CFFFCFFF3FCB2D2B58804FA7E" authorityName="Kunth" authorityYear="1824" box="[773,862,1448,1474]" class="Magnoliopsida" family="Fabaceae" genus="Dioclea" kingdom="Plantae" order="Fabales" pageId="0" pageNumber="293" phylum="Tracheophyta" rank="genus">
<emphasis id="B978EAADFFFCFFF3FCB2D2B58804FA7E" box="[773,862,1448,1474]" italics="true" pageId="0" pageNumber="293">Dioclea</emphasis>
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subgenera
<emphasis id="B978EAADFFFCFFF3FC69D2B58F28FA7E" box="[990,1138,1448,1474]" italics="true" pageId="0" pageNumber="293">Pachylobium</emphasis>
and
<taxonomicName id="4C0C4D3CFFFCFFF3FB1ED2B58E68FA7E" box="[1193,1330,1448,1474]" class="Magnoliopsida" family="Fabaceae" genus="Platylobium" kingdom="Plantae" order="Fabales" pageId="0" pageNumber="293" phylum="Tracheophyta" rank="genus">
<emphasis id="B978EAADFFFCFFF3FB1ED2B58E68FA7E" box="[1193,1330,1448,1474]" italics="true" pageId="0" pageNumber="293">Platylobium</emphasis>
</taxonomicName>
composed a paraphyletic grade, nesting the genera
<taxonomicName id="4C0C4D3CFFFCFFF3FDFDD2D08820FA5A" authority="Elmer (1907: 220)" authorityName="Elmer" authorityPageNumber="220" authorityYear="1907" box="[586,890,1484,1510]" class="Magnoliopsida" family="Fabaceae" genus="Luzonia" kingdom="Plantae" order="Fabales" pageId="0" pageNumber="293" phylum="Tracheophyta" rank="genus">
<emphasis id="B978EAADFFFCFFF3FDFDD2D089FEFA5A" box="[586,676,1485,1510]" italics="true" pageId="0" pageNumber="293">Luzonia</emphasis>
<bibRefCitation id="EF9D4B4EFFFCFFF3FD1CD2D18820FA5A" author="Elmer, A. D. E." box="[683,890,1484,1510]" pageId="0" pageNumber="293" pagination="220 - 232" refId="ref3338" refString="Elmer, A. D. E. (1907) Some new Leguminosae. Leaflets of Philippine Botany 1: 220 - 232." type="journal article" year="1907">Elmer (1907: 220)</bibRefCitation>
</taxonomicName>
and
<emphasis id="B978EAADFFFCFFF3FC07D2D18FD6FA5A" box="[944,1164,1484,1510]" italics="true" pageId="0" pageNumber="293">
<taxonomicName id="4C0C4D3CFFFCFFF3FC07D2D18FD2FA5A" authorityName="Harms ex K. Schumann &amp; Lauterbach" authorityYear="1900" box="[944,1160,1484,1510]" class="Magnoliopsida" family="Fabaceae" genus="Macropsychanthus" kingdom="Plantae" order="Fabales" pageId="0" pageNumber="293" phylum="Tracheophyta" rank="genus">Macropsychanthus</taxonomicName>
.
</emphasis>
Therefore, they proposed that the circumscription of
<taxonomicName id="4C0C4D3CFFFCFFF3FE03D2ED8957F9B6" authorityName="Kunth" authorityYear="1824" box="[436,525,1520,1546]" class="Magnoliopsida" family="Fabaceae" genus="Dioclea" kingdom="Plantae" order="Fabales" pageId="0" pageNumber="293" phylum="Tracheophyta" rank="genus">
<emphasis id="B978EAADFFFCFFF3FE03D2ED8957F9B6" box="[436,525,1520,1546]" italics="true" pageId="0" pageNumber="293">Dioclea</emphasis>
</taxonomicName>
should be restricted to
<taxonomicName id="4C0C4D3CFFFCFFF3FCB8D2ED8F51F9B6" box="[783,1035,1520,1546]" class="Magnoliopsida" family="Fabaceae" genus="Dioclea" kingdom="Plantae" order="Fabales" pageId="0" pageNumber="293" phylum="Tracheophyta" rank="subGenus" subGenus="Dioclea">
<emphasis id="B978EAADFFFCFFF3FCB8D2ED8832F9B6" box="[783,872,1520,1546]" italics="true" pageId="0" pageNumber="293">Dioclea</emphasis>
subg.
<emphasis id="B978EAADFFFCFFF3FC07D2ED8F51F9B6" box="[944,1035,1520,1546]" italics="true" pageId="0" pageNumber="293">Dioclea</emphasis>
</taxonomicName>
<emphasis id="B978EAADFFFCFFF3FBBCD2ED8F55F9B6" box="[1035,1039,1520,1546]" italics="true" pageId="0" pageNumber="293">,</emphasis>
with 13 species and that the limits of
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<emphasis id="B978EAADFFFCFFF3FF3FD1098A06F992" box="[136,348,1556,1582]" italics="true" pageId="0" pageNumber="293">Macropsychanthus</emphasis>
</taxonomicName>
should be widened to include the genus
<emphasis id="B978EAADFFFCFFF3FCA0D1088822F992" box="[791,888,1557,1582]" italics="true" pageId="0" pageNumber="293">
<taxonomicName id="4C0C4D3CFFFCFFF3FCA0D108882EF992" authorityName="Elmer" authorityYear="1907" box="[791,884,1557,1582]" class="Magnoliopsida" family="Fabaceae" genus="Luzonia" kingdom="Plantae" order="Fabales" pageId="0" pageNumber="293" phylum="Tracheophyta" rank="genus">Luzonia</taxonomicName>
,
</emphasis>
as well as the
<taxonomicName id="4C0C4D3CFFFCFFF3FBA2D1098F34F992" authorityName="Kunth" authorityYear="1824" box="[1045,1134,1556,1582]" class="Magnoliopsida" family="Fabaceae" genus="Dioclea" kingdom="Plantae" order="Fabales" pageId="0" pageNumber="293" phylum="Tracheophyta" rank="genus">
<emphasis id="B978EAADFFFCFFF3FBA2D1098F34F992" box="[1045,1134,1556,1582]" italics="true" pageId="0" pageNumber="293">Dioclea</emphasis>
</taxonomicName>
subgenera
<emphasis id="B978EAADFFFCFFF3FB5DD1098E24F992" box="[1258,1406,1556,1582]" italics="true" pageId="0" pageNumber="293">Pachylobium</emphasis>
and
<emphasis id="B978EAADFFFCFFF3FF3FD1258A43F9EE" box="[136,281,1592,1618]" italics="true" pageId="0" pageNumber="293">
<taxonomicName id="4C0C4D3CFFFCFFF3FF3FD1258A4FF9EE" box="[136,277,1592,1618]" class="Magnoliopsida" family="Fabaceae" genus="Platylobium" kingdom="Plantae" order="Fabales" pageId="0" pageNumber="293" phylum="Tracheophyta" rank="genus">Platylobium</taxonomicName>
.
</emphasis>
Under the current circumscription,
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<emphasis id="B978EAADFFFCFFF3FD14D125882DF9EE" box="[675,887,1592,1618]" italics="true" pageId="0" pageNumber="293">Macropsychanthus</emphasis>
Harms ex K. Schumann &amp; Lauterbach (1900: 366)
</taxonomicName>
comprises ca. 50 species, with a pantropical distribution, mainly confined to the Neotropics (
<bibRefCitation id="EF9D4B4EFFFCFFF3FB15D1418EFAF9CA" author="Queiroz, L. P. &amp; Snak, C." box="[1186,1440,1628,1654]" pageId="0" pageNumber="293" pagination="67 - 114" refId="ref3578" refString="Queiroz, L. P. &amp; Snak, C. (2020) Revisiting the taxonomy of Dioclea and related genera (Leguminosae, Papilionoideae), with new generic circumscriptions. PhytoKeys 164 (1): 67 - 114. https: // doi. org / 10.3897 / phytokeys. 164.55441" type="journal article" year="2020">Queiroz &amp; Snak 2020</bibRefCitation>
). The genus is characterized by its lianescent habit, erect pseudoracemose inflorescences composed of secundiflorous brachyblasts, bicallosal standard petals and turgid or laterally compressed fruits (
<bibRefCitation id="EF9D4B4EFFFCFFF3FBBAD1B98E5DF902" author="Queiroz, L. P. &amp; Snak, C." box="[1037,1287,1700,1726]" pageId="0" pageNumber="293" pagination="67 - 114" refId="ref3578" refString="Queiroz, L. P. &amp; Snak, C. (2020) Revisiting the taxonomy of Dioclea and related genera (Leguminosae, Papilionoideae), with new generic circumscriptions. PhytoKeys 164 (1): 67 - 114. https: // doi. org / 10.3897 / phytokeys. 164.55441" type="journal article" year="2020">Queiroz &amp; Snak 2020</bibRefCitation>
). Phylogenetic studies on the genus supported the presence of two clades, recognized as subgenera by
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: subgenus
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<emphasis id="B978EAADFFFCFFF3FF40D1F18A91F8BA" box="[247,459,1772,1798]" italics="true" pageId="0" pageNumber="293">Macropsychanthus</emphasis>
</taxonomicName>
and subgenus
<taxonomicName id="4C0C4D3CFFFCFFF3FDC6D1F189A0F8BA" box="[625,762,1772,1798]" class="Magnoliopsida" family="Fabaceae" genus="Platylobium" kingdom="Plantae" order="Fabales" pageId="0" pageNumber="293" phylum="Tracheophyta" rank="genus">
<emphasis id="B978EAADFFFCFFF3FDC6D1F189A0F8BA" box="[625,762,1772,1798]" italics="true" pageId="0" pageNumber="293">Platylobium</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB336BFFFFCFFF3FF0AD00D8ECEF7BE" blockId="0.[136,1452,1268,2051]" pageId="0" pageNumber="293">
The subgenus
<taxonomicName id="4C0C4D3CFFFCFFF3FED3D00D8AB7F896" box="[356,493,1808,1834]" class="Magnoliopsida" family="Fabaceae" genus="Platylobium" kingdom="Plantae" order="Fabales" pageId="0" pageNumber="293" phylum="Tracheophyta" rank="genus">
<emphasis id="B978EAADFFFCFFF3FED3D00D8AB7F896" box="[356,493,1808,1834]" italics="true" pageId="0" pageNumber="293">Platylobium</emphasis>
</taxonomicName>
is restricted to South America and is characterized by basifixed triangular stipules, leaves lacking stipels, laterally compressed fruits and elastically dehiscent, and seeds with circular hilum.This subgenus includes nine species:
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<emphasis id="B978EAADFFFCFFF3FE37D04589F1F8CE" box="[384,683,1880,1906]" italics="true" pageId="0" pageNumber="293">Macropsychanthus bicolor</emphasis>
(Benth.) L.P. Queiroz &amp; Snak, (2020:105)
</taxonomicName>
<taxonomicName id="4C0C4D3CFFFCFFF3FB30D0448A92F82A" authority="(Benth.) L. P. Queiroz &amp; Snak (2020: 106)" authorityName="L. P. Queiroz &amp; Snak" authorityPageNumber="106" authorityYear="2020" baseAuthorityName="Benth." class="Magnoliopsida" family="Fabaceae" genus="Macropsychanthus" kingdom="Plantae" order="Fabales" pageId="0" pageNumber="293" phylum="Tracheophyta" rank="species" species="coriaceus">
<emphasis id="B978EAADFFFCFFF3FB30D0448E42F8CE" box="[1159,1304,1881,1906]" italics="true" pageId="0" pageNumber="293">M. coriaceus</emphasis>
(Benth.) L.P.
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</taxonomicName>
,
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<emphasis id="B978EAADFFFCFFF3FE62D060891FF82A" box="[469,581,1916,1942]" italics="true" pageId="0" pageNumber="293">M. duckei</emphasis>
L.P.
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</taxonomicName>
,
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<emphasis id="B978EAADFFFCFFF3FC79D0608F1EF82A" box="[974,1092,1917,1942]" italics="true" pageId="0" pageNumber="293">M. erectus</emphasis>
(Hoehne) L.P.
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</taxonomicName>
,
<taxonomicName id="4C0C4D3CFFFCFFF3FEA2D0BC8F10F806" authority="(R. H. Maxwell) L. P. Queiroz &amp; Snak (2020: 106)" authorityName="L. P. Queiroz &amp; Snak" authorityPageNumber="106" authorityYear="2020" baseAuthorityName="R. H. Maxwell" box="[277,1098,1952,1979]" class="Magnoliopsida" family="Fabaceae" genus="Macropsychanthus" kingdom="Plantae" order="Fabales" pageId="0" pageNumber="293" phylum="Tracheophyta" rank="species" species="hispidimarginatus">
<emphasis id="B978EAADFFFCFFF3FEA2D0BC8952F806" box="[277,520,1952,1978]" italics="true" pageId="0" pageNumber="293">M. hispidimarginatus</emphasis>
(R.H. Maxwell) L.P.
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</taxonomicName>
,
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<emphasis id="B978EAADFFFCFFF3FBEED0BC8E4EF806" box="[1113,1300,1953,1978]" italics="true" pageId="0" pageNumber="293">M. macrocarpus</emphasis>
(Huber) L.P.
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</taxonomicName>
,
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<emphasis id="B978EAADFFFCFFF3FE63D0D88913F862" box="[468,585,1988,2014]" italics="true" pageId="0" pageNumber="293">M. rigidus</emphasis>
(R.S. Cowan) L.P.
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</taxonomicName>
,
<taxonomicName id="4C0C4D3CFFFCFFF3FBC4D0D88AACF7BE" authority="(R. H. Maxwell) L. P. Queiroz &amp; Snak (2020: 107)" authorityName="L. P. Queiroz &amp; Snak" authorityPageNumber="107" authorityYear="2020" baseAuthorityName="R. H. Maxwell" class="Magnoliopsida" family="Fabaceae" genus="Macropsychanthus" kingdom="Plantae" order="Fabales" pageId="0" pageNumber="293" phylum="Tracheophyta" rank="species" species="ruddiae">
<emphasis id="B978EAADFFFCFFF3FBC4D0D88FAAF862" box="[1139,1264,1988,2014]" italics="true" pageId="0" pageNumber="293">M. ruddiae</emphasis>
(R.H. Maxwell) L.P.
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</taxonomicName>
, and
<taxonomicName id="4C0C4D3CFFFCFFF3FD86D0F48F2CF7BE" authority="(Rich.) L. P. Queiroz &amp; Snak (2020: 107)" authorityName="L. P. Queiroz &amp; Snak" authorityPageNumber="107" authorityYear="2020" baseAuthorityName="Rich." box="[561,1142,2024,2051]" class="Magnoliopsida" family="Fabaceae" genus="Macropsychanthus" kingdom="Plantae" order="Fabales" pageId="0" pageNumber="293" phylum="Tracheophyta" rank="species" species="scabrus">
<emphasis id="B978EAADFFFCFFF3FD86D0F489F4F7BE" box="[561,686,2024,2050]" italics="true" pageId="0" pageNumber="293">M. scabrus</emphasis>
(Rich.) L.P.
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</taxonomicName>
(
<bibRefCitation id="EF9D4B4EFFFCFFF3FB32D0F58ED2F7BE" author="Queiroz, L. P. &amp; Snak, C." box="[1157,1416,2024,2050]" pageId="0" pageNumber="293" pagination="67 - 114" refId="ref3578" refString="Queiroz, L. P. &amp; Snak, C. (2020) Revisiting the taxonomy of Dioclea and related genera (Leguminosae, Papilionoideae), with new generic circumscriptions. PhytoKeys 164 (1): 67 - 114. https: // doi. org / 10.3897 / phytokeys. 164.55441" type="journal article" year="2020">Queiroz &amp; Snak, 2020</bibRefCitation>
).
</paragraph>
<paragraph id="8BB336BFFFFDFFF2FF0AD7838E6FFEF4" blockId="1.[136,1452,158,328]" pageId="1" pageNumber="294">
Bentham described
<taxonomicName id="4C0C4D3CFFFDFFF2FE2ED783884DFF04" authority="Benth. (1837: 69)" authorityName="Benth." authorityPageNumber="69" authorityYear="1837" box="[409,791,158,184]" class="Magnoliopsida" family="Fabaceae" genus="Dioclea" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="294" phylum="Tracheophyta" rank="species" species="rostrata">
<emphasis id="B978EAADFFFDFFF2FE2ED783890AFF04" box="[409,592,158,184]" italics="true" pageId="1" pageNumber="294">Dioclea rostrata</emphasis>
Benth. (1837:69)
</taxonomicName>
. Later, in the Flora Brasiliensis,
<bibRefCitation id="EF9D4B4EFFFDFFF2FBCED7838E74FF05" author="Bentham, G." box="[1145,1326,158,185]" pageId="1" pageNumber="294" pagination="15" refId="ref3286" refString="Bentham, G. (1859) Leguminosae 1, Papilionaceae. In: Martius, C. F. P., Endlicher, S. &amp; Urban, I. (Eds.) Flora Brasiliensis, vol. 15, part 1. Frid. Fleisher, Munich." type="book chapter" year="1859">Bentham (1859)</bibRefCitation>
introduced a new variety:
<taxonomicName id="4C0C4D3CFFFDFFF2FE87D7DE8AF6FF60" box="[304,428,195,221]" class="Magnoliopsida" family="Fabaceae" genus="Dioclea" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="294" phylum="Tracheophyta" rank="species" species="rostrata">
<emphasis id="B978EAADFFFDFFF2FE87D7DE8A1EFF60" box="[304,324,195,220]" italics="true" pageId="1" pageNumber="294">D</emphasis>
.
<emphasis id="B978EAADFFFDFFF2FEE4D7DE8AF6FF60" box="[339,428,195,220]" italics="true" pageId="1" pageNumber="294">rostrata</emphasis>
</taxonomicName>
var. “β”
<emphasis id="B978EAADFFFDFFF2FDA4D7DF8901FF60" box="[531,603,194,220]" italics="true" pageId="1" pageNumber="294">
<taxonomicName id="4C0C4D3CFFFDFFF2FDA4D7DF890DFF60" authorityName="R. H. Maxwell. Finally, Queiroz and Snak" authorityYear="2020" baseAuthorityName="Benth." box="[531,599,194,220]" class="Magnoliopsida" family="Fabaceae" genus="Dioclea" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="294" phylum="Tracheophyta" rank="species" species="nitida">nitida</taxonomicName>
,
</emphasis>
only known from the
<typeStatus id="54B7881DFFFDFFF2FCEED7DE88D3FF61" box="[857,905,195,221]" pageId="1" pageNumber="294">type</typeStatus>
collection. A detailed morphological analysis of
<typeStatus id="54B7881DFFFDFFF2FF3FD7FA8BE2FEBD" box="[136,184,231,257]" pageId="1" pageNumber="294">type</typeStatus>
collections and the specimens of this subgenus revealed the presence of more collections of this taxon, mainly identified as
<taxonomicName id="4C0C4D3CFFFDFFF2FEA0D6178A9DFE98" box="[279,455,266,292]" class="Magnoliopsida" family="Fabaceae" genus="Dioclea" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="294" phylum="Tracheophyta" rank="species" species="bicolor">
<emphasis id="B978EAADFFFDFFF2FEA0D6178A9DFE98" box="[279,455,266,292]" italics="true" pageId="1" pageNumber="294">Dioclea bicolor</emphasis>
</taxonomicName>
. Here we propose a new combination and a new rank for
<taxonomicName id="4C0C4D3CFFFDFFF2FBF4D6168E6EFE98" box="[1091,1332,266,293]" class="Magnoliopsida" family="Fabaceae" genus="Dioclea" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="294" phylum="Tracheophyta" rank="variety" species="rostrata" variety="nitida">
<emphasis id="B978EAADFFFDFFF2FBF4D6168FE4FE98" box="[1091,1214,267,292]" italics="true" pageId="1" pageNumber="294">D. rostrata</emphasis>
var.
<emphasis id="B978EAADFFFDFFF2FB44D6178E6EFE98" box="[1267,1332,266,292]" italics="true" pageId="1" pageNumber="294">nitida</emphasis>
</taxonomicName>
, including complete description, illustration, distribution, and comparison within the morphologically similar species.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="2BF4B2A1E518331256BA543505D7FA6A">Evolutionary Systematics</mods:title>
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<mods:part id="E5C3085D67F58FAAA46C8F3F5092D1EB">
<mods:date id="561798B4DB8E438AAEA3C8040482A16B">2024</mods:date>
<mods:detail id="5ADC1F18AA07C53E129BE8208DDE8132" type="pubDate">
<mods:number id="55A324CC40B8730ED49DD990DEDFFC0C">2024-11-20</mods:number>
</mods:detail>
<mods:detail id="AD27F0280AC443441B27D317551C675D" type="volume">
<mods:number id="6654AA4B4E4A5A74F9E75B9B84B5B67D">8</mods:number>
</mods:detail>
<mods:detail id="598D271A4D2EA1AA2A5E22D7187066DC" type="issue">
<mods:number id="AD16AA7968D11798B8457119F83E97AC">2</mods:number>
</mods:detail>
<mods:extent id="A9E09D1EEE1B211997F4BAEFF1AA3D04" unit="page">
<mods:start id="B41962839DB68EA1D575D8E73037A3BC">261</mods:start>
<mods:end id="2D4B68BAFA6965848AFD7EDFEDAD2877">271</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="49DE1A33E5E1BD1215EED174E4D81999">journal article</mods:classification>
<mods:identifier id="704A90BE1A5D290411AE63973B393921" type="DOI">10.3897/evolsyst.8.141847</mods:identifier>
<mods:identifier id="8C55C8539E0A3BB277DF795BC8AE38A0" type="ZooBank">8E0DCC28-3448-4001-90B7-E675BC107381</mods:identifier>
</mods:mods>
<treatment id="DE0471CA9CD752339E0AD4EB8DC33A83" ID-DOI="http://doi.org/10.5281/zenodo.14193091" ID-Zenodo-Dep="14193091" ID-arpha="DE0471CA-9CD7-5233-9E0A-D4EB8DC33A83" ID-zoobank="https://zoobank.org/8C021216-678A-4C8F-B3EA-B885945C8915" LSID="urn:lsid:plazi:treatment:DE0471CA9CD752339E0AD4EB8DC33A83" LSID-ZBK="urn:lsid:zoobank.org:act:8C021216-678A-4C8F-B3EA-B885945C8915" httpUri="http://treatment.plazi.org/id/DE0471CA9CD752339E0AD4EB8DC33A83">
<subSubSection id="9191FEF4BAD36B63C2D48B9918917139" type="nomenclature">
<paragraph id="17C5F2AA1623A7185DB8394A8C7F488C">
<taxonomicName id="C961A5A23C466AD559C1E884AC83D9B0" authorityName="Wermelinger-Moreira &amp; de Souza Castanheira &amp; Baptista" authorityYear="2024" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="candango" status="sp. nov.">
<emphasis id="66344EC07E3F7629A581CEC19CA4A3BA" italics="true">Harmonicon candango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2F116FDB3C9192A4D5BED3040AAA5BDC" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="E723C0609BDFD0F9AA5AA26E7088EE8E" type="description">
<paragraph id="3C0BE045AB6BBD19392CBCAC2DA464A8">
<figureCitation id="8CA3ABF155933E3214099417E96B8232" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">Figs 2 A J</figureCitation>
,
<figureCitation id="49E3AA07FFF2961D6CEBBC4CCFA3D29C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Harmonicon candango sp. nov., female paratype (MNRJ 7626). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view; D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive female specimen in captivity. Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178837">3 A G</figureCitation>
,
<figureCitation id="F22013A35E2AD231437B78418EE9386D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution records of H. rufescens (holotype = red and black circle, specimen here described = red circle) and H. candango sp. nov. (yellow square)." httpUri="https://binary.pensoft.net/fig/1178838">4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZQAG" type="material">
<paragraph id="3DDD56EE62EE34ABD41E174CC1309713">
<heading id="A7AE2DA7924F8A507C383607FBBEB2F7" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="E0F5E1D81DEA9E042330A3692FB60C83">
<materialsCitation id="B27EB2D6F18B2DB40B4B73991E745097" collectingDate="1973-10-18" collectionCode="MNRJ" collectorName="W. R. Lourenco" country="Brazil" latitude="-15.738917" location="Brasilia" longLatPrecision="1" longitude="-47.926918" specimenCode="MNRJ 7625" specimenCount="1" specimenCount-male="1" stateProvince="Distrito Federal" typeStatus="holotype">
<emphasis id="14DB9F00CAD12B90B8E4FF80A8E53D43" bold="true" italics="true">
<typeStatus id="7E958C73CAADEB0C45EDF59DF2BD9970" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="51DCB41176EF965D1BCAAACD008C0235" name="Brazil">Brazil</collectingCountry>
<specimenCount id="E28D0FB125862F10B84B140E84EE4C56" count="1" type="male"></specimenCount>
;
<location id="CA97BEBD74301469D5D54C2D11AFBE02" LSID="urn:lsid:plazi:treatment:DE0471CA9CD752339E0AD4EB8DC33A83:CA97BEBD74301469D5D54C2D11AFBE02" country="Brazil" latitude="-15.738917" longLatPrecision="1" longitude="-47.926918" name="Brasilia" stateProvince="Distrito Federal">Brasília</location>
,
<collectingRegion id="8E50EF45E6E6625FE621DA5503491FF7" country="Brazil" name="Distrito Federal">Distrito Federal</collectingRegion>
[
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EHRAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-47.926917,-15.738917]}">
<geoCoordinate id="524E547F82BEE46AAC3B236BF291B2EC" degrees="15" direction="south" minutes="44" orientation="latitude" precision="1" seconds="20.1" value="-15.738917">15 ° 44 ' 20.1 &quot; S</geoCoordinate>
,
<geoCoordinate id="8B6656A2AEE2E2761EE53AF00CF4637F" degrees="47" direction="west" minutes="55" orientation="longitude" precision="1" seconds="36.9" value="-47.926918">47 ° 55 ' 36.9 &quot; W</geoCoordinate>
</named-content>
];
<date id="6B7EFFA08FD1AF4A43BB3E2FE35DD906" value="1973-10-18">
<collectingDate id="BDDD6C4BFECD6773A22D52129791EDAF" value="1973-10-18">18 October 1973</collectingDate>
</date>
;
<collectorName id="928A8C942E0A64717FC5D652A70A4FB9">W. R. Lourenço</collectorName>
leg.;
<specimenCode id="3090849F963F28C08F6DC187FADC9D1C" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" type="">
<named-content id="580C064A47FB6D36642C4A169CC0918B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-nacional-universidade-federal-do-rio-de-janeiro" xlink_title="Laboratório de Aracnologia, Museu Nacional, Universidade Federal do Rio de Janeiro">MNRJ</named-content>
7625
</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="E6AB511EA75743A755A2B041619D1B28">
<materialsCitation id="4117C08CBF1E671301C9EB204B060274" collectionCode="MNRJ" country="Brazil" specimenCode="MNRJ 7626, MNRJ 13709" specimenCount="5" specimenCount-female="1" specimenCount-male="4" typeStatus="paratype">
<emphasis id="E7068EDB39633B3B24612916D82377FC" bold="true" italics="true">
<typeStatus id="1DBF713C29827605E796819F739331D6" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="D8A153029DC576704C2A7195AC20AD8D" name="Brazil">Brazil</collectingCountry>
<specimenCount id="7D1C226FD4B77CD92B5A4D887BD3D9E7" count="1" type="female"></specimenCount>
; same data as for preceding;
<specimenCode id="79668C536D5F61EDEF772F9AD3F74DB6" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" type="">
<named-content id="F9285BA9FE25867589206ED90E3F1E21" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-nacional-universidade-federal-do-rio-de-janeiro" xlink_title="Laboratório de Aracnologia, Museu Nacional, Universidade Federal do Rio de Janeiro">MNRJ</named-content>
7626
</specimenCode>
;
<specimenCount id="953D2018594FD9D06D56CD289459A54C" count="4" type="male">4 ♂♂</specimenCount>
, 19 j; same data as for preceding;
<specimenCode id="0111BA107B64CA0761908D7A6D8843E8" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" type="">
<named-content id="94BFFCFA3F8BDE6EBF3939ED06DA872E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-nacional-universidade-federal-do-rio-de-janeiro" xlink_title="Laboratório de Aracnologia, Museu Nacional, Universidade Federal do Rio de Janeiro">MNRJ</named-content>
13709
</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6RAG" type="etymology">
<paragraph id="348C6B5F9250E0F6A6D637A4BDAB0426">
<heading id="ED6E9F2E2DB4F761E815E3F44F423EA0" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="D2327EA0722D076022A64434079C5EAD">
The specific epithet
<taxonomicName id="CFC70996E4515FF935890FF00DDBBD02" authorityName="Wermelinger-Moreira &amp; de Souza Castanheira &amp; Baptista" authorityYear="2024" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="candango">
<emphasis id="2CB6895678729262550DA0631AD36EED" italics="true">candango</emphasis>
</taxonomicName>
” is a masculine noun in apposition used to designate the inhabitants of the city of Brasília, especially to the immigrants from Northeast
<collectingCountry id="BE7796EF24648668B794D0DE0F02944F" name="Brazil">Brazil</collectingCountry>
who built the city.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOSAG" type="diagnosis">
<paragraph id="79733421BDE3883E7B386FE8C4866E97">
<heading id="0211D5FF0E9E8E783E8679C744D95F0D" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="4B56C679CD60C0447C018C2891BF1C4C">
<taxonomicName id="4E5DF8936F6BD59BF9518B4B902A9B5C" authorityName="Wermelinger-Moreira &amp; de Souza Castanheira &amp; Baptista" authorityYear="2024" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="candango" status="sp. nov.">
<emphasis id="533FE18507EB2D8ECE845C08D3D34FA8" italics="true">Harmonicon candango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="62BD7CDCB9C6B5A3FE49A221C03A21E6" rank="species">sp. nov.</taxonomicNameLabel>
can be separated from all other
<taxonomicName id="BF11E2BB69FF71291358058EF3CBADAB" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="AF0DD7702879198207524A57644B70A6" italics="true">Harmonicon</emphasis>
</taxonomicName>
by its lyra composed of setae curved from its base and bearing spatulate tips (Figs
<figureCitation id="60EB0735BEB0ED53F4356CB7188B1B27" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 H</figureCitation>
,
<figureCitation id="FCBF2ADEFBF138327FDFF7828E359CA9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Harmonicon candango sp. nov., female paratype (MNRJ 7626). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view; D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive female specimen in captivity. Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178837">3 C</figureCitation>
) (vs. setae with strongly curved, thin, and acute tips) (e. g., Fig.
<figureCitation id="701D9C6142EBDBDBF5BA101EBBF7F83A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 C</figureCitation>
;
<bibRefCitation id="151992EEDEBDF0E73FAD77C6D3CDDE7D" DOI="10.5962/p.268900" author="Marechal and Marty" firstAuthor="Marechal" journalOrPublisher="Zoosystema" pagination="499-504" refId="B9" refString="Maréchal P, Marty C (1998) Réhabilitation du genre Harmonicon (Pickard-Cambridge, 1896) et description d'une nouvelle espèce de Guyane française (Araneae, Mygalomorphae, Dipluridae). Zoosystema 20: 499504. https://doi.org/10.5962/p.268900" title="Réhabilitation du genre Harmonicon (Pickard-Cambridge, 1896) et description d'une nouvelle espèce de Guyane française (Araneae, Mygalomorphae, Dipluridae)." volume="20" year="1998">Maréchal and Marty 1998</bibRefCitation>
, fig. 2 A, B;
<bibRefCitation id="D2BD853A98D52953CAC7ECC962BB801A" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 8996. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
, fig. 6). Males of
<taxonomicName id="3E5397080E4567F2FBB37EC614D37743" authorityName="Wermelinger-Moreira &amp; de Souza Castanheira &amp; Baptista" authorityYear="2024" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="candango" status="sp. nov.">
<emphasis id="43E545B57D1737B9CBC3247FAB12079F" italics="true">H. candango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0DA1692C94C238ABE6B7A9F19446E65E" rank="species">sp. nov.</taxonomicNameLabel>
resemble
<taxonomicName id="A140F65D22795F06BD8F9E2C1CB6B372" authorityName=", Drolshagen and Backstam" authorityYear="2011" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="oiapoqueae">
<emphasis id="95855E82C27AE1AB2DAA2CDA69A24C2E" italics="true">H. oiapoqueae</emphasis>
</taxonomicName>
by the dark colour and the similar pedipalp bulb profile, with a low hump at the bulb near the basis of embolus, which is not crooked and almost straight (Fig.
<figureCitation id="A75DEE3B253F68FB15FE30ECE48B803E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 B, D</figureCitation>
). They may be recognised from those of
<taxonomicName id="CBE4B076D21A55F1F1587B206DE2196C" authorityName=", Drolshagen and Backstam" authorityYear="2011" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="oiapoqueae">
<emphasis id="3E69BF681AE547BE7BE0D63EAE890C47" italics="true">H. oiapoqueae</emphasis>
</taxonomicName>
by the presence of the ventral setae near the retrolateral tubercle at metatarsus I (Fig.
<figureCitation id="C61BBB625EF8B35E1A771EEDDCDE90B2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 J</figureCitation>
), tibia I with retrolateral spur bearing thorn thinner and more curved (Fig.
<figureCitation id="E763E9D84160E381BE3F73A2BCCC6671" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 I, J</figureCitation>
) (vs. spur thick and almost straight;
<bibRefCitation id="23915B686CB24DE8D3C2630DE8842D7C" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 8996. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
, figs 4, 7), and prolateral macroseta opposing the distal spur long and thin (Fig.
<figureCitation id="6960F7C939DE04C8C451962514EE889C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 J</figureCitation>
, just insertion mark) (vs. absent in
<taxonomicName id="00DC55BD70C5A4325D59595EFFE88A41" authorityName=", Drolshagen and Backstam" authorityYear="2011" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="oiapoqueae">
<emphasis id="85B4BD2ACA28DD4C3665214032A09E56" italics="true">H. oiapoqueae</emphasis>
</taxonomicName>
;
<bibRefCitation id="2436E74383D0A875CA74C24594A7E6F3" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 8996. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
, fig. 7). Furthermore, the pedipalp bulb of
<taxonomicName id="30750B32423D2F9646895907B699C186" authorityName="Wermelinger-Moreira &amp; de Souza Castanheira &amp; Baptista" authorityYear="2024" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="candango" status="sp. nov.">
<emphasis id="99C274F5C2809C9A4ED4EE30E47CCA8B" italics="true">H. candango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="847D075BAE09BD0D84127BE3953D6100" rank="species">sp. nov.</taxonomicNameLabel>
has a longer embolus (ca. 1, 5 × longer than the bulb proper, Fig.
<figureCitation id="ACC5E54B8690772069058A733C5E0014" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 B, D</figureCitation>
) than
<taxonomicName id="C4FCE833B793CB1E2498EFB70125A104" authorityName=", Drolshagen and Backstam" authorityYear="2011" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="oiapoqueae">
<emphasis id="01E8121A75360AB3DD624A06DB11D1E6" italics="true">H. oiapoqueae</emphasis>
</taxonomicName>
(ca. 1, 2 ×;
<bibRefCitation id="C801D18FB0E680F01309498EB0846F77" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 8996. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
, fig. 2) in retrolateral view. Females of
<taxonomicName id="CDC542CCAFE4CDF6A8A3F51A8AC07D57" authorityName="Wermelinger-Moreira &amp; de Souza Castanheira &amp; Baptista" authorityYear="2024" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="candango" status="sp. nov.">
<emphasis id="2B5CAA1F0030E7D24623A1148FC9CA74" italics="true">H. candango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="78A9C3FAC536B7B994F47D2E5AC5DD93" rank="species">sp. nov.</taxonomicNameLabel>
are differentiated from
<taxonomicName id="910C6B7324EE6A3CD8A13689545C069A" authorityName=", Drolshagen and Backstam" authorityYear="2011" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="oiapoqueae">
<emphasis id="ACE3B72FCCBC4164111B18BC2E3EC8FD" italics="true">H. oiapoqueae</emphasis>
</taxonomicName>
and
<taxonomicName id="48F9E7723DBB21F7974D5AED1EE511CE" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="76595EFA3A2F6A3148636299FA6743EF" italics="true">H. rufescens</emphasis>
</taxonomicName>
by the spermathecae with median branch bearing 34 apical lobules (Fig.
<figureCitation id="C268DE79C5A0D68A436A983128AD4769" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Harmonicon candango sp. nov., female paratype (MNRJ 7626). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view; D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive female specimen in captivity. Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178837">3 D F</figureCitation>
) (vs. tip round, blunt, and without lobules) (Fig.
<figureCitation id="8666F0C4BA46F1E11C864B633B0B909D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 D F</figureCitation>
;
<bibRefCitation id="DE0DB319490E54EEBC99EDEB6589DAC6" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 8996. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
, fig. 5), and lateral branch with a very wide and multilobulate tip (Fig.
<figureCitation id="2ADD11528CD7A02E73D2103CBE4AA1F8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Harmonicon candango sp. nov., female paratype (MNRJ 7626). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view; D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive female specimen in captivity. Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178837">3 D F</figureCitation>
) (vs. tip with few lobules or folded over the stem) (Fig.
<figureCitation id="0997C58951DA92988D5225211029376F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Harmonicon rufescens, female (UFRJ 2575). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view (arrow points to thick erect and regularly curved setae); D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive immature female specimen in nature (MNRJ 4362). Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178835">1 D F</figureCitation>
;
<bibRefCitation id="18F426ABF76DF7AE7CDBF52EB15EBA5B" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 8996. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
, fig. 5).
</paragraph>
<caption id="08BBEFFA7241ACF3A10035EF2081F774" ID-arpha="4BC8001A-6733-59B1-B46C-9ED064E12408" httpUri="https://binary.pensoft.net/fig/1178836" startId="F2">
<paragraph id="C02791E0B6441891B30239D5ECB904A1">
<label id="FC3C15FD761C92066AF1325B331FA73B">Figure 2.</label>
</paragraph>
<paragraph id="DCD13535B6908F08A5D6A19B1EA4F9A6">
<taxonomicName id="5F190785989E02AC4CE011EA428440C3" authorityName="Wermelinger-Moreira &amp; de Souza Castanheira &amp; Baptista" authorityYear="2024" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="candango" status="sp. nov.">
<emphasis id="F2BD8C438079842A18B9ED952B8D4289" italics="true">Harmonicon candango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02E5A517C6B04D54286D4F006D7CE08B" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype (
<named-content id="E2A6613BD4AC3A1821BE197E3DC4298F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-nacional-universidade-federal-do-rio-de-janeiro" xlink_title="Laboratório de Aracnologia, Museu Nacional, Universidade Federal do Rio de Janeiro">MNRJ</named-content>
7625).
<emphasis id="70F3F01D8C90D8925AAB5BA651A5DF86" bold="true">A.</emphasis>
Dorsal habitus;
<emphasis id="4213CED82EAD1CE4C5EC767A188E1B08" bold="true">B F.</emphasis>
Male left pedipalp:
<emphasis id="4F9D7FA211D45A820C503FCA12C32A42" bold="true">B.</emphasis>
Tibia, retrolateral view;
<emphasis id="BB9382D77A634D165273AB46F15AB72E" bold="true">C F.</emphasis>
Palpal bulb;
<emphasis id="E366FFD068B1649B18A19F92A10AF078" bold="true">C.</emphasis>
Prolateral view;
<emphasis id="D295CA20B28522E085F2E8E78C0BAB6D" bold="true">D.</emphasis>
Retrolateral view;
<emphasis id="60F86FA9A19DC54D0647F5D3FAAED8A0" bold="true">E.</emphasis>
Dorsal view;
<emphasis id="18CA1F82BB43533B6DBF0ADDB7C3CCDF" bold="true">F.</emphasis>
Ventral view;
<emphasis id="49C6A9602A2B06BD1F9F150AD6D339EE" bold="true">G.</emphasis>
Sternum;
<emphasis id="1518D9723E5F4E561C2D8E8F57C1601A" bold="true">H.</emphasis>
Lyra of left maxilla, ventral view;
<emphasis id="69B7D321A8432E484826833F7FEA29B0" bold="true">I J.</emphasis>
Tibia I:
<emphasis id="2E4C111C6C68F60F33B5A43DDC0246E1" bold="true">I.</emphasis>
Retrolateral view;
<emphasis id="37F7A067FEA58DA294B6DB81B7314E7B" bold="true">J.</emphasis>
Ventral view. Scale bars: 5 mm (
<emphasis id="08A4E9CCA8CC5AAD0FE1C5D720C36B66" bold="true">A</emphasis>
); 2 mm (
<emphasis id="36A800FDB4E5C8D12D8E072295FB0BCE" bold="true">B, I J</emphasis>
); 1 mm (
<emphasis id="5D0F5A75D405ECB4B77C75DB3796EC7E" bold="true">C H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ER1AG" type="description">
<paragraph id="B8488416CB526CD7EF238DE8D27287EB">
<heading id="90DC88D546F50FFDC4D5964A4049F8B2" reason="title">Description.</heading>
</paragraph>
<paragraph id="0A1E27A179BF7C76AE69A04CF30148A3">
<emphasis id="7B683A91E7B2CB68CFA52FE01BC78EDC" bold="true">
Male (
<emphasis id="59A2D09797F16C00AFBE2D7B3C6C0F79" bold="true" italics="true">
<typeStatus id="F14C0B1F4B698F7114028B9B0D42D187" type="holotype">holotype</typeStatus>
</emphasis>
,
<named-content id="2E00BD852527617ACEEDDFBE12FDCA57" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-nacional-universidade-federal-do-rio-de-janeiro" xlink_title="Laboratório de Aracnologia, Museu Nacional, Universidade Federal do Rio de Janeiro">
<collectionCode id="C4265A859E095A87CC5B226E210EB2AE" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" type="">MNRJ</collectionCode>
</named-content>
7625)
</emphasis>
. Carapace oblong, wider at middle portion, light orange-brown, caput slightly elevated and darker; cephalic area with darker furrows, covered with dark setae; and margins of thoracic area with a row of long, thick, rigid, erect, and curved dark setae pointing ectally (Fig.
<figureCitation id="50E896977C20B8CCD88A5DE8107EF478" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 A</figureCitation>
). Clypeus long, with frontal margin carrying 15 median-sized to long, thick setae facing forward (Fig.
<figureCitation id="CC8B3FC3116057DC2D2132FAD8709221" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 A</figureCitation>
). Eye tubercle with 10 thick setae on its anterior margin; area between eyes with 10 shorter, thicker setae and others abundant dark but not thicker setae (Fig.
<figureCitation id="1653BAF6E02060A2BCB2AF633E42E3C2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 A</figureCitation>
). Eye formula:
<abbrev id="ABBRID0EM2AG" xlink_title="anterior lateral eyes">ALE</abbrev>
&gt;
<abbrev id="ABBRID0EQ2AG" xlink_title="posterior lateral eyes">PLE</abbrev>
&gt;
<abbrev id="ABBRID0EU2AG" xlink_title="anterior median eyes">AME</abbrev>
&gt;
<abbrev id="ABBRID0EY2AG" xlink_title="posterior median eyes">PME</abbrev>
. Ocular area much wider than long. Anterior eye row slightly recurved. Posterior eye row slightly recurved. Median ocular quadrangle wider than long, wider at its posterior margin.
<abbrev id="ABBRID0E32AG" xlink_title="anterior median eyes">AME</abbrev>
rounded,
<abbrev id="ABBRID0EA3AG" xlink_title="anterior lateral eyes">ALE</abbrev>
and
<abbrev id="ABBRID0EE3AG" xlink_title="posterior lateral eyes">PLE</abbrev>
elliptical, and
<abbrev id="ABBRID0EI3AG" xlink_title="posterior median eyes">PME</abbrev>
reduced, oval.
<abbrev id="ABBRID0EM3AG" xlink_title="anterior median eyes">AME</abbrev>
-
<abbrev id="ABBRID0EQ3AG" xlink_title="anterior median eyes">AME</abbrev>
separated by less than its diameter.
<abbrev id="ABBRID0EU3AG" xlink_title="anterior median eyes">AME</abbrev>
contiguous to
<abbrev id="ABBRID0EY3AG" xlink_title="posterior median eyes">PME</abbrev>
.
<abbrev id="ABBRID0E33AG" xlink_title="anterior median eyes">AME</abbrev>
separated from
<abbrev id="ABBRID0EA4AG" xlink_title="posterior median eyes">PME</abbrev>
by ca. half its diameter.
<abbrev id="ABBRID0EE4AG" xlink_title="anterior lateral eyes">ALE</abbrev>
and
<abbrev id="ABBRID0EI4AG" xlink_title="posterior median eyes">PME</abbrev>
contiguous to
<abbrev id="ABBRID0EM4AG" xlink_title="posterior lateral eyes">PLE</abbrev>
. Fovea transverse and slightly recurved (Fig.
<figureCitation id="CA47F17A8ECA4AE9ED0345AA5E95FB7B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 A</figureCitation>
). Labium wider than long with no cuspules (Fig.
<figureCitation id="DB398BE8419F24FE13C5DC8283306C0A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 G</figureCitation>
). Sternum longer than wide, covered by black erect setae and brownish thin recumbent setae, one pair of labial sigilla comma-shaped and almost touching each other along the median line of the body, three sigilla on each side near margin, increasing in size from first to last, the third pair larger and elliptical (Fig.
<figureCitation id="A1D8CDB9294707FA9E2F14BFDF17EEE1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 G</figureCitation>
). Chelicerae dark red with 11 promarginal teeth at right and 12 promarginal teeth at left, fang furrow with denticles. Plectrum with five separated thick setae on both sides. Maxillae elongated with 22 and 25 cuspules at left and right pedipalps, respectively (Fig.
<figureCitation id="FC2C7E1AC1F7D26771F26B12A659625B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 G</figureCitation>
). Lyra asymmetrical at ventral side of maxilla, formed by 67 modified thick and long setae, increasing in size basally to distally; apical portion with 12 less curved setae and subsequent ones strongly curved. Left lyra with six (Fig.
<figureCitation id="D4FF8E86C866B801171D0234D472A1FA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 H</figureCitation>
) and right one with seven setae. Leg formula IV&gt; I&gt; II&gt; III; length of segments (femur + patella + tibia + metatarsus + tarsus = total length): I 8.0 + 2.7 + 8.6 + 8.2 + 5.3 = 32.8, II 6.4 + 2.0 + 7.2 + 6.7 + 5.1 = 27.4, III 6.2 + 1.5 + 6.1 + 7.7 + 4.9 = 26.4, IV 7.3 + 2.1 + 8.1 + 10.1 + 6.4 = 34.0. Leg I with modified tibia and metatarsus, forming a clasping mechanism (Fig.
<figureCitation id="0248CC6F0DC56BE567D75879417F2B9D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 I, J</figureCitation>
). Tibia I with retrolateral spur (or apophysis) relatively long, elevated and pronounced in both lateral and retrolateral view, thicker than its thorn, curved and blunt (Fig.
<figureCitation id="796E8FC88EF5D59AF1574EBC3AA71CD9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 I, J</figureCitation>
); thorn longer than the spur, relatively thin, curved at retrolateral view (Fig.
<figureCitation id="88473F014273F6B233713C680237E2BE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 I</figureCitation>
), straight at ventral view (Fig.
<figureCitation id="6278499BFF5531495235D91FF331125D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 J</figureCitation>
), pointed at tip, and with prolateral distal spine (fallen off, but indicated by its insertion). Metatarsus I with retrolateral tubercle obtuse, with wide basis, and situated about the same level as the basis of the first ventral macroseta at the retrolateral side, tubercle region slightly thickened in ventral view (Fig.
<figureCitation id="17E758027F4E1290BBB307DB4F3F3FF3" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 I, J</figureCitation>
). Tarsi with few small cracks covering almost all dorsal and lateral faces, except by basis and tip of article; all tarsi with dense scopulae; metatarsus I II with scopulae; metatarsus I with more than metatarsus II; metatarsus III IV with thin scopulae;
<abbrev id="ABBRID0EY5AG" xlink_title="inferior tarsal claw">ITC</abbrev>
without teeth;
<abbrev id="ABBRID0E35AG" xlink_title="superior tarsal claw">STC</abbrev>
teeth: I: 115, II: 135, III: 124, IV: 104. Macrosetae: Leg I: femur pld 00 2, d 32 0 right, d 12 0 left, rld 11 1, patella 0, tibia r 11 0, p 11 1 right, p 01 1 left, v 11 1 ap (apophysis), metatarsus v 11 1; Leg II: femur d 12 0 right, d 22 0 left, pld 02 1 right, rld 01 2 left, patella 0, tibia p 01 1, v 11 2, metatarsus v 12 1 right, v 11 2 left, p 01 0; Leg III: femur d 21 0, rld 11 1, pld 12 1, patella 0, tibia d 01 1 right, d 01 0 left, p 01 1, v 10 2 right, v 11 2 left, metatarsus d 11 1, rld 11 0, pld 02 1 left, p 11 1 right, v 22 3; Leg IV: femur rld 22 1, d 42 0, pld 11 0 right, pld 11 1 left, patella 0, tibia d 01 0, p 11 0, r 01 1, rld 10 0, v 12 2 left, v 22 2 right, metatarsus d 22 2, rld 12 0 right, p 01 1 right, v 21 4, p 11 1 left, plv 00 1 left, r 11 1 left. Abdomen covered with dark setae, dorsum brown without stripes, with a slightly paler area before the cardiac mark, which is outlined by a somewhat darker brown line (Fig.
<figureCitation id="0454683FB27A73C0CDE2F4961378D60F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 A</figureCitation>
), and venter uniform brown.
<abbrev id="ABBRID0EE6AG" xlink_title="posterior lateral spinnerets">PLS</abbrev>
longer than half of abdomen, with articulate fusules in all articles, and last article finger-like and with light patches of ventral cracks (Fig.
<figureCitation id="36CAF91AA0391B858F06D148E2BB5D3A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 A</figureCitation>
). Pedipalp elongated, bearing three macrosetae, one retrolateral placed near the end of its median third (Fig.
<figureCitation id="944EF815D6DF5A0AB9126DC0E48E71BE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 B</figureCitation>
, fallen off, just the insertion), one prolateral at its distal third, and one prolateroventral at its median third. Pedipalp bulb 0.25 × longer than wide, oblong in prolateral (Fig.
<figureCitation id="F5F8DB2D78F3B3CA4FB68FBC3EA9D383" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 D</figureCitation>
) and retrolateral views (Fig.
<figureCitation id="2E20A1887D9B23EF39119938799DDEB9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 C</figureCitation>
), but more globose in dorsal (Fig.
<figureCitation id="36B7EE283AF4FB5EEA69A31DD47330BC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 E</figureCitation>
) and ventral views (Fig.
<figureCitation id="A51BCA87BE668B07D07778BF16D1B877" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 F</figureCitation>
), tapering to merge with embolus basis well demarcated (Fig.
<figureCitation id="50A499FCC572A9DE86CAC30A1007CFD1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 C F</figureCitation>
). Embolus around 1.5 × longer than bulb itself, almost straight, tapering towards its acute, slightly curved tip (Fig.
<figureCitation id="08BF89C55CB232511547FBE209C59DE1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 C F</figureCitation>
).
</paragraph>
<paragraph id="431DA8D3ACD7663179110134A5C65AF1">
<abbrev id="ABBRID0EKABG" xlink_title="total length">TL</abbrev>
20.1. Carapace 9.0 long, 7.8 wide, carapace length / width 1.15.
<abbrev id="ABBRID0EOABG" xlink_title="anterior median eyes">AME</abbrev>
<quantity id="0732F4E0D08239DFE54283C6E7B9DAB0" metricMagnitude="-4" metricUnit="m" metricValue="2.7" unit="mm" value="0.27">0.27 mm</quantity>
,
<abbrev id="ABBRID0ESABG" xlink_title="anterior median eyes">AME</abbrev>
-
<abbrev id="ABBRID0EWABG" xlink_title="anterior median eyes">AME</abbrev>
<quantity id="7FADF84ECED496B6871C94B36DCE937C" metricMagnitude="-4" metricUnit="m" metricValue="2.3" unit="mm" value="0.23">0.23 mm</quantity>
. Abdomen 11.1 long, 5.4 wide. Spinnerets:
<abbrev id="ABBRID0E1ABG" xlink_title="posterior median spinnerets">PMS</abbrev>
1.8 long;
<abbrev id="ABBRID0E5ABG" xlink_title="posterior lateral spinnerets">PLS</abbrev>
13.4 long, basal article 3.5, middle 4.1, distal 5.8.
</paragraph>
<paragraph id="05F523D5E87F29361543A1B777E916FC">
<emphasis id="83A8364934AD8C53EB4AD4CE65F1DE28" bold="true">
Female (
<emphasis id="F0A8A7DCF429FB57C56D4D67ACCD48A7" bold="true" italics="true">
<typeStatus id="9E708C4B9A4A695732C1BD4767687629" type="paratype">paratype</typeStatus>
</emphasis>
,
<named-content id="B27701EDEFA9699893E33B1F99390A2E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-nacional-universidade-federal-do-rio-de-janeiro" xlink_title="Laboratório de Aracnologia, Museu Nacional, Universidade Federal do Rio de Janeiro">
<collectionCode id="65596CC1E9CA7DDF3C11E9AF9876FAC1" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" type="">MNRJ</collectionCode>
</named-content>
7626)
</emphasis>
. Carapace and clypeus as in male (Fig.
<figureCitation id="F74C8471F09EC6D41B79D60AF7BEB9DF" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Harmonicon candango sp. nov., female paratype (MNRJ 7626). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view; D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive female specimen in captivity. Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178837">3 A</figureCitation>
). Eye tubercle with 13 thick setae on its anterior margin; area between eyes with 13 shorter, thicker setae and others abundant grey and thinner setae (Fig.
<figureCitation id="E26881A0A3EE6ADDA043680829DDB160" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Harmonicon candango sp. nov., female paratype (MNRJ 7626). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view; D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive female specimen in captivity. Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178837">3 A</figureCitation>
). Eyes as in male (Fig.
<figureCitation id="6BE6E303F6B18BDFC5B47F202B877639" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Harmonicon candango sp. nov., female paratype (MNRJ 7626). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view; D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive female specimen in captivity. Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178837">3 A</figureCitation>
). Labium and sternum as in male (Fig.
<figureCitation id="85806E38CC9E99E85ED3F3CE090AE1B3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Harmonicon candango sp. nov., female paratype (MNRJ 7626). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view; D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive female specimen in captivity. Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178837">3 B</figureCitation>
). Chelicerae, as in male with 11 promarginal teeth at both sides. Plectrum as in male but with 89 separated thick setae on left side and right side, respectively. Maxillae with shape as in male but bearing 29 and 27 cuspules at the left and right pedipalp, respectively (Fig.
<figureCitation id="5C3EAB49A27D801E992305B88DB72334" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Harmonicon candango sp. nov., female paratype (MNRJ 7626). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view; D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive female specimen in captivity. Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178837">3 B</figureCitation>
). Lyra at ventral side of maxilla similar to male. Left lyra with seven regular setae and an additional distal one much thinner than the others (Fig.
<figureCitation id="AD05886C32FE5C3ADB0A76D68B2AF320" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Harmonicon candango sp. nov., female paratype (MNRJ 7626). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view; D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive female specimen in captivity. Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178837">3 C</figureCitation>
), right lyra with seven setae. Leg formula IV&gt; I&gt; II&gt; III; length of segments (femur + patella + tibia + metatarsus + tarsus = total length): I 6.1 + 2.4 + 6.5 + 5.2 + 3.6 = 23.8, II 5.0 + 2.2 + 5.7 + 4.8 + 3.4 = 21.1, III 5.1 + 1.2 + 5.0 + 5.7 + 3.3 = 20.3, IV 5.8 + 1.6 + 6.7 + 7.9 + 4.3 = 26.3. Tarsi and metatarsi as in male, palpal tarsi with middle to apical scopulae; pedipalp claw with eight teeth;
<abbrev id="ABBRID0EFCBG" xlink_title="inferior tarsal claw">ITC</abbrev>
without teeth;
<abbrev id="ABBRID0EJCBG" xlink_title="superior tarsal claw">STC</abbrev>
teeth: I: 123, II: 84, III: 123, IV: 83. Macrosetae: Leg I (right absent): femur pld 00 1 left, d 11 0 left, rld 00 1 left, patella 0 left, tibia p 01 1 left, v 11 2 left, metatarsus v 12 1 left; Leg II: femur d 11 0, pld 01 1, patella 0, tibia p 01 1, v 01 1 right, v 01 2 left, metatarsus p 01 0 left, v 12 3 left, v 12 2 right; Leg III: femur d 30 0 right, d 20 0 left, rld 11 1, pld 01 1 right, pld 01 0 left, patella 0, tibia rld 10 0, r 11 0 right, r 01 0 left, p 11 0, v 11 2, metatarsus d 00 2, p 11 1, pld 01 1 left, r 11 0, v 22 1 left, v 22 3 right; Leg IV: femur d 21 0 right, d 41 0 left, rld 13 1 left, rld 12 1 right, patella 0, tibia rld 10 0, r 11 0, p 11 0, v 10 2, v 11 2 right, metatarsus p 11 1 left, p 11 0 right pld 01 1, rld 12 1, v 14 3 left, v 23 3 right. Abdomen similar to male, slightly damaged, dorsally with a paler anterior area near the cardiac mark (Fig.
<figureCitation id="DED5AC0EA3D118BBBEED496A365B2BC4" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Harmonicon candango sp. nov., female paratype (MNRJ 7626). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view; D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive female specimen in captivity. Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178837">3 A</figureCitation>
). Spermathecae separated by ca. 1.2 × its length, with two thick stems (or branches) at each side, basal part not sclerotised, translucent, remaining portions sclerotised, opaque, and yellowish brown, each branch with almost same thickness; median branch basally straight with a curvature at the middle portion outwards and then straight up to its tip with 34 lobules; lateral branch straight with tip much larger than its stem; and tip of the median branch multilobulated (Fig.
<figureCitation id="44AF31B62EFAC40E588D6D5037089B84" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Harmonicon candango sp. nov., female paratype (MNRJ 7626). A. Dorsal habitus; B. Sternum; C. Lyra of left maxilla, ventral view; D F. Spermathecae: D. Ventral view; E. Dorsal view; F. Ventral view, partially cleared; G. Alive female specimen in captivity. Scale bars: 5 mm (A); 2 mm (B); 1 mm (C); 0.5 mm (D E). Living specimen photo by Denis Pedroso." httpUri="https://binary.pensoft.net/fig/1178837">3 D F</figureCitation>
).
</paragraph>
<caption id="3EC760956FE394DCA123AD671AC72435" ID-arpha="454BCF7C-73D4-589A-873C-6F9DC33D5785" httpUri="https://binary.pensoft.net/fig/1178837" startId="F3">
<paragraph id="AD99A98D32C022A972A2BB9D495A6ED0">
<label id="1E22546B241A12829C728C8E8B8B7CE5">Figure 3.</label>
</paragraph>
<paragraph id="61716E3CE13AB82FA479116D4EF07D21">
<taxonomicName id="F1E19EE488C51D624F86706672832321" authorityName="Wermelinger-Moreira &amp; de Souza Castanheira &amp; Baptista" authorityYear="2024" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="candango" status="sp. nov.">
<emphasis id="825587B7073FF20AA99628B7CD722B9E" italics="true">Harmonicon candango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2E0882F3C4A163781C63E757572FF852" rank="species">sp. nov.</taxonomicNameLabel>
, female paratype (
<named-content id="64B2C0E5ACBEA678DB3AE869F9D1B974" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museu-nacional-universidade-federal-do-rio-de-janeiro" xlink_title="Laboratório de Aracnologia, Museu Nacional, Universidade Federal do Rio de Janeiro">MNRJ</named-content>
7626).
<emphasis id="C42D923D42E3F7A8171F3A9ADC400D80" bold="true">A.</emphasis>
Dorsal habitus;
<emphasis id="D47E7F4BE9DDE6BFC7BF44F37F4A783D" bold="true">B.</emphasis>
Sternum;
<emphasis id="A868DDD8B311DE1894ED16B950B821E8" bold="true">C.</emphasis>
Lyra of left maxilla, ventral view;
<emphasis id="CAED05FCBFB98F94274A67ACB3587DA3" bold="true">D F.</emphasis>
Spermathecae:
<emphasis id="2032B4C0DC1B7C6F7A442F3BE6D58C87" bold="true">D.</emphasis>
Ventral view;
<emphasis id="5A529F11C62DD58A00182FDA2CCC71D6" bold="true">E.</emphasis>
Dorsal view;
<emphasis id="79711998FBBEA8C02353A93731530994" bold="true">F.</emphasis>
Ventral view, partially cleared;
<emphasis id="85C942A06260E3F0FEEB0BA1C2E94748" bold="true">G.</emphasis>
Alive female specimen in captivity. Scale bars: 5 mm (
<emphasis id="974FADA220F00549C39C5448218D648E" bold="true">A</emphasis>
); 2 mm (
<emphasis id="B1346F727CB59B7E3F51AEE3488D8CB9" bold="true">B</emphasis>
); 1 mm (
<emphasis id="5B1E18D4A10E969D0F1A32D7CF64A723" bold="true">C</emphasis>
); 0.5 mm (
<emphasis id="ACBCACDAF2F3B360826ACC7096A65FD7" bold="true">D E</emphasis>
). Living specimen photo by Denis Pedroso.
</paragraph>
</caption>
<paragraph id="CFE439CD16B3620FEF18DF88914CCFD2">
<abbrev id="ABBRID0ELEBG" xlink_title="total length">TL</abbrev>
24.9. Carapace 9.4 long, 7.6 wide, carapace length / width 1.24.
<abbrev id="ABBRID0EPEBG" xlink_title="anterior median eyes">AME</abbrev>
<quantity id="FEA4F9347EA77745CE0763BFBA442A65" metricMagnitude="-4" metricUnit="m" metricValue="2.9" unit="mm" value="0.29">0.29 mm</quantity>
,
<abbrev id="ABBRID0ETEBG" xlink_title="anterior median eyes">AME</abbrev>
-
<abbrev id="ABBRID0EXEBG" xlink_title="anterior median eyes">AME</abbrev>
<quantity id="CE6DBBF5EE2F65A0BB62477B0E869438" metricMagnitude="-4" metricUnit="m" metricValue="2.6" unit="mm" value="0.26">0.26 mm</quantity>
. Abdomen 15.5 long, 8.8 wide. Spinnerets: absent.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2EBG" type="Taxonomic notes">
<paragraph id="D5ADF3EEBDDF1CF86D5106CEE7437456">
<heading id="A52E32D665F3C89D4471AE63C764567C" reason="title">Taxonomic notes.</heading>
</paragraph>
<paragraph id="7644B5DD734C6DB36929B1C6297DE4F5">
Our interpretation of the published drawings and photos of some
<taxonomicName id="D2C89479EDE3377D64A4F7858E76F2AD">
<emphasis id="CC4EDF308B38457E5518FDD2DE86C012" italics="true">Harmonicon</emphasis>
species
</taxonomicName>
differs from those stated in their respective original publications. For example, the
<taxonomicName id="45985C89EE7AE4988BC348EF3F593BA7" authorityName="Pedroso &amp; Baptista" authorityYear="2014" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="cerberus">
<emphasis id="D5255A03E5720ADA55CA33005263B485" italics="true">H. cerberus</emphasis>
</taxonomicName>
<typeStatus id="726FD41971FF33B0A08D6F14B3DABA7D" type="holotype">holotype</typeStatus>
seems to have suffered a torsion during its collecting and conservation process.
<bibRefCitation id="F7931837C9721D16563C201B625C51D6" DOI="10.3897/zookeys.389.6693" author="Pedroso DR &amp; Baptista RLC" firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="77-88" refId="B12" refString="Pedroso DR, Baptista RLC (2014) A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus. ZooKeys 389: 7788. https://doi.org/10.3897/zookeys.389.6693" title="A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus." volume="389" year="2014">Pedroso and Baptista (2014)</bibRefCitation>
stated that the figures 7, 8, and 9 of the palpal bulb of
<taxonomicName id="0667976789FFD376C8E30FFC11625DC6" authorityName="Pedroso &amp; Baptista" authorityYear="2014" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="cerberus">
<emphasis id="029033CE0408EBBB1C275561659AD0B7" italics="true">H. cerberus</emphasis>
</taxonomicName>
were retrolateral, prolateral, and dorsal views, respectively. However, we now consider their fig. 9 as better interpreted as retrolateral, matching the profile of the palpal bulbs of other
<taxonomicName id="8373EC3A85E0A84C257232B078563C3F">
<emphasis id="B6B9C6C425BADF3AC9A1277C1BBF1F77" italics="true">Harmonicon</emphasis>
species
</taxonomicName>
(i. e., Fig.
<figureCitation id="E37AB534253399FB0DC3219D6A62B7B8" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 D, F</figureCitation>
). Furthermore,
<bibRefCitation id="9D004C4B510C9F1682041404C39654C9" DOI="10.3897/zookeys.389.6693" author="Pedroso and Baptista" firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="77-88" refId="B12" refString="Pedroso DR, Baptista RLC (2014) A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus. ZooKeys 389: 7788. https://doi.org/10.3897/zookeys.389.6693" title="A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus." volume="389" year="2014">Pedroso and Baptista (2014</bibRefCitation>
, fig. 8) seems to be a dorsal view, while
<bibRefCitation id="DEBB8786B1009B9B2CF706098E9D6D31" DOI="10.3897/zookeys.389.6693" author="Pedroso and Baptista" firstAuthor="Pedroso" journalOrPublisher="ZooKeys" pagination="77-88" refId="B12" refString="Pedroso DR, Baptista RLC (2014) A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus. ZooKeys 389: 7788. https://doi.org/10.3897/zookeys.389.6693" title="A new troglomorphic species of Harmonicon (Araneae, Mygalomorphae, Dipluridae) from Pará, Brazil, with notes on the genus." volume="389" year="2014">Pedroso and Baptista (2014</bibRefCitation>
, fig. 7) is then a ventral view (vs. Fig.
<figureCitation id="602EA11D6547C52CCAD70515BC1EFCAE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Harmonicon candango sp. nov., male holotype (MNRJ 7625). A. Dorsal habitus; B F. Male left pedipalp: B. Tibia, retrolateral view; C F. Palpal bulb; C. Prolateral view; D. Retrolateral view; E. Dorsal view; F. Ventral view; G. Sternum; H. Lyra of left maxilla, ventral view; I J. Tibia I: I. Retrolateral view; J. Ventral view. Scale bars: 5 mm (A); 2 mm (B, I J); 1 mm (C H)." httpUri="https://binary.pensoft.net/fig/1178836">2 E, F</figureCitation>
). A similar torsion has probably happened in the male
<typeStatus id="BB2071F7CCF37A713ED97A019614F517" type="holotype">holotype</typeStatus>
of
<taxonomicName id="1CC4DA2CDB21AC97A2952D034EADB0B5" authorityName=", Drolshagen and Backstam" authorityYear="2011" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="oiapoqueae">
<emphasis id="530496346BC4615F95803E972F66B34C" italics="true">H. oiapoqueae</emphasis>
</taxonomicName>
.
<bibRefCitation id="55A7011A017654DA4DBFBFD57504EB72" DOI="10.3897/zookeys.112.1205" author="Drolshagen B &amp; Bäckstam CM" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 8996. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam (2011)</bibRefCitation>
stated that they included drawings of the retrolateral and prolateral views (
<bibRefCitation id="78C5114B57EBE451331B103D606E38E7" DOI="10.3897/zookeys.112.1205" author="Drolshagen and Backstam" firstAuthor="Drolshagen" journalOrPublisher="ZooKeys" pagination="89-96" refId="B4" refString="Drolshagen B, Bäckstam CM (2011) Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana. ZooKeys 112: 8996. https://doi.org/10.3897/zookeys.112.1205" title="Notes on the genus Harmonicon F. O. P. - Cambridge, 1896 (Araneae, Dipluridae) with description of a new species from French Guyana." volume="112" year="2011">Drolshagen and Bäckstam 2011</bibRefCitation>
, figs 1, 2) of the right pedipalp, but careful comparison indicates that they are respectively dorsal and ventral views, both slightly inclined sideways.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMHBG" type="variation">
<paragraph id="E9041D5E87CE07691A99F96CBB5B3121">
<heading id="CB4C4F1CC114755784C170244A1653B2" reason="title">Variation.</heading>
</paragraph>
<paragraph id="EA72AC4810D339A1B07657B0E7C0E20A">
Males (n = 5).
<abbrev id="ABBRID0ESHBG" xlink_title="total length">TL</abbrev>
14.420.1. Chelicerae with 1012 promarginal teeth, plectrum with 58 setae, maxillary cuspules from 2238. Tibia I with prolateral distal macrosetae thinner and longer than the retrolateral thorn (whenever present). Pedipalp tibia with a variable set of macrosetae, sometimes with the prolateral or retrolateral lacking, often with the prolateroventral lacking.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWHBG" type="distribution">
<paragraph id="C79CD3FBAB50052D732D7D7537899803">
<heading id="2D43773A8F0B9B1B2AA76C0C7B134439" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="F9359C4FA1B7CDEF431A35772BBEFAAE">
Only known from Brasília,
<collectingRegion id="9062A472819224501A71EA6260DAD711" country="Brazil" name="Distrito Federal">Distrito Federal</collectingRegion>
,
<collectingCountry id="3D31F6F4A4D049C99C7637A8721362C5" name="Brazil">Brazil</collectingCountry>
, a locality in the Cerrado biome (Fig.
<figureCitation id="9F170368EC031242B7969F4C0674CD8A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution records of H. rufescens (holotype = red and black circle, specimen here described = red circle) and H. candango sp. nov. (yellow square)." httpUri="https://binary.pensoft.net/fig/1178838">4</figureCitation>
).
</paragraph>
<caption id="07A992162C30D7E153C8E7AD7035DFAB" ID-arpha="79E0CF11-C678-518C-B579-DA36B4294E37" httpUri="https://binary.pensoft.net/fig/1178838" startId="F4">
<paragraph id="30993F8B43E94E07DB1EA201C454393E">
<label id="9A16DE4FD3A3826E5B6AE8EAA846E563">Figure 4.</label>
</paragraph>
<paragraph id="E4103E9FC028421983E15E03EB8CA1B3">
Distribution records of
<taxonomicName id="03437E1A86CC97759FF506C8C783C12E" authorityName="F. O. Pickard-Cambridge" authorityYear="1896" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="2CA381268251F119626B0A6031DAB4F6" italics="true">H. rufescens</emphasis>
</taxonomicName>
(holotype = red and black circle, specimen here described = red circle) and
<taxonomicName id="FE65D2578E0E625EED6BAF98A60F5974" authorityName="Wermelinger-Moreira &amp; de Souza Castanheira &amp; Baptista" authorityYear="2024" class="Arachnida" family="Dipluridae" genus="Harmonicon" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="candango" status="sp. nov.">
<emphasis id="1CD9981F66A49B2C0506C4E6A354CAA3" italics="true">H. candango</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B6C498A1B0F65EDA4A6684F6FD79A5D2" rank="species">sp. nov.</taxonomicNameLabel>
(yellow square).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>