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<mods:title id="9691CA7E909C5813218B568FD8C2E332">evdsdon of the darkldng beetle genus Eurynotus (Blaptdnae: Platynotdnd % and new records of ovovdvdpary dn ºenebrdonddae</mods:title>
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<mods:namePart id="9B09DBAB68029D228D5B343C4FF1274A">Kamdńskd, Marcdn J.</mods:namePart>
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<mods:date id="98CD54FFD751C416ADF3EEE5731557E7">2024</mods:date>
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<subSubSection id="02C0653AFFE0FFCF9A050201FD767058" box="[119,757,1852,1879]" pageId="18" pageNumber="376" type="nomenclature">
<paragraph id="4A6536B1FFE0FFCF9A050201FD767058" blockId="18.[113,762,1852,2009]" box="[119,757,1852,1879]" pageId="18" pageNumber="376">
<heading id="112D81DDFFE0FFCF9A050201FD767058" box="[119,757,1852,1879]" centered="true" fontSize="9" level="2" pageId="18" pageNumber="376" reason="4">
<emphasis id="78AEEAA3FFE0FFCF9A050201FD767058" bold="true" box="[119,757,1852,1879]" pageId="18" pageNumber="376">
2.
<taxonomicName id="8DDA4D32FFE0FFCF9AFC0201FDAB7059" authority="(Mulsant and Rey)" baseAuthorityName="Mulsant and Rey" box="[142,552,1852,1879]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="asperipennis">
<emphasis id="78AEEAA3FFE0FFCF9AFC0201FEE17059" bold="true" box="[142,354,1852,1879]" italics="true" pageId="18" pageNumber="376">Biolus asperipennis</emphasis>
(Mulsant and Rey)
</taxonomicName>
stat. res.,comb. nov.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="02C0653AFFE0FFCF9B290246FB0A7162" pageId="18" pageNumber="376" type="description">
<paragraph id="4A6536B1FFE0FFCF9B290246FD92709A" blockId="18.[113,762,1852,2009]" box="[347,529,1915,1941]" pageId="18" pageNumber="376">
<heading id="112D81DDFFE0FFCF9B290246FD92709A" box="[347,529,1915,1941]" centered="true" fontSize="9" level="2" pageId="18" pageNumber="376" reason="3">
<emphasis id="78AEEAA3FFE0FFCF9B290246FD92709A" bold="true" box="[347,529,1915,1941]" pageId="18" pageNumber="376">
(
<figureCitation id="D2E12A34FFE0FFCF9B140246FE30709A" box="[358,435,1915,1941]" captionStart="Figure 2" captionStartId="3.[129,194,1510,1534]" captionTargetBox="[137,1469,147,1474]" captionTargetId="figure-202@3.[129,1473,144,1483]" captionTargetPageId="3" captionText="Figure 2. Biolus stat.nov.specdes.A, Biolus analisetosus (Koch% comb. nov. Holotype; B, B.asperipennis (Mulsant and Rey% stat.res., comb.nov. Lectotype; C, B. brunoi sp. nov. Holotype; D, B. denticosta (Mulsant and Rey% comb. nov. Lectotype; E, B. graafi (Koch% comb. nov. Holotype; F,B. granulatus (Fabrdcdus% comb. nov. Lectotype; G, B. omeri (Koch% comb.nov.Holotype; H, B. sauroides (Koch% comb. nov. Holotype; I, Eurynotus norrisi Mulsant and Rey syn. nov. of B. granulatus, Lectotype." pageId="18" pageNumber="376">Figs 2B</figureCitation>
,
<figureCitation id="D2E12A34FFE0FFCF9BCF0246FE5C709A" box="[445,479,1915,1941]" captionStart="Figure 6" captionStartId="7.[129,194,1700,1724]" captionTargetBox="[131,1472,145,1670]" captionTargetId="figure-63@7.[129,1473,144,1673]" captionTargetPageId="7" captionText="Figure 6. Characters used dn the claddstdc analysds.Arrows dnddcate partdcular states of gdven characters from the morphologdcal matrdx [e.g." pageId="18" pageNumber="376">6G</figureCitation>
,
<figureCitation id="D2E12A34FFE0FFCF9B9B0246FD85709A" box="[489,518,1915,1941]" captionStart="Figure 8" captionStartId="13.[129,194,1898,1922]" captionTargetBox="[134,1473,837,1871]" captionTargetId="figure-393@13.[129,1473,834,1871]" captionTargetPageId="13" captionText="Figure 8. Aedeagal morphology of Eurynotdna specdes.A, Biolus analisetosus; B, B.asperipennis; C, B. brunoi sp. nov.; D, B. denticosta; E, B. graafi; F, B. granulatus; G, B. omeri; H, B. sauroides; I, Eurynotus asperatus; J, E.barbosai; K, E. capensis; L, E. graniderma; M, Capidium tuberculatum; N, Oncotus cedrimontis; O, Heteropsectropus amaroides; P, Schyzoschelus africanus. Parameres hdghldghted red. Scale = 1mm." pageId="18" pageNumber="376">8B</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="4A6536B1FFE0FFCF9A03029FFD7970D6" blockId="18.[113,762,1852,2009]" pageId="18" pageNumber="376">
<pageTitle id="0A45EED6FFE0FFCF9A03029FFD7970D6" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF9A03029FFED670B6" box="[113,341,1954,1977]" italics="true" pageId="18" pageNumber="376">Eurynotus asperipennis</emphasis>
Mulsant and Rey, 1854a: 23. Mulsant and Rey 1854b: 169; Lacordadre 1859: 239; Gebden 1938: 414;
</pageTitle>
</paragraph>
<paragraph id="4A6536B1FFE0FFCF995805ADFC1B77C8" blockId="18.[810,1459,144,199]" pageId="18" pageNumber="376">
Koch 1954a: 280; 1954b: 2;
<bibRefCitation id="2E4B4B40FFE0FFCF9E2205ADFAC477A7" author="Banaszkdewdcz M" box="[1104,1351,144,168]" pageId="18" pageNumber="376" pagination="59 - 77" refId="ref25294" refString="Banaszkdewdcz M. Comparatdve study of female gendtalda dn Peddndnd sens Iwan 2004 (Coleoptera: ºenebrdonddae: Peddndnd %, wdth notes on the classdficatdon. Annales Zoologici 2006; 56: 59 - 77." type="journal article" year="2006">Banaszkdewdcz 2006: 63</bibRefCitation>
; Kamdńskd 2016: 235.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF995805D4FB0E7577" blockId="18.[810,1460,233,632]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF995805D4FC13760F" box="[810,912,233,256]" italics="true" pageId="18" pageNumber="376">Diagnosis:</emphasis>
Lacks sclerdte dn bursa copulatrdx (present dn
<taxonomicName id="8DDA4D32FFE0FFCF9FED05D4FC1E762F" authorityName="Koch" authorityYear="1954" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="analisetosus">
<emphasis id="78AEEAA3FFE0FFCF9FED05D4FC1E762F" italics="true" pageId="18" pageNumber="376">B. analisetosus</emphasis>
</taxonomicName>
and
<taxonomicName id="8DDA4D32FFE0FFCF99920434FBE9762F" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[992,1130,264,288]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="78AEEAA3FFE0FFCF99920434FBE9762F" box="[992,1130,264,288]" italics="true" pageId="18" pageNumber="376">B. granulatus</emphasis>
</taxonomicName>
%.
<taxonomicName id="8DDA4D32FFE0FFCF9EF80435FACF762F" box="[1162,1356,264,288]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="asperipennis">
<emphasis id="78AEEAA3FFE0FFCF9EF80435FACF762F" box="[1162,1356,264,288]" italics="true" pageId="18" pageNumber="376">Biolus asperipennis</emphasis>
</taxonomicName>
has two puncture
<typeStatus id="95618813FFE0FFCF99E30415FC4B764F" box="[913,968,296,320]" pageId="18" pageNumber="376">types</typeStatus>
on ventrdte V (sdngle
<typeStatus id="95618813FFE0FFCF9EC80415FB6B764F" box="[1210,1256,296,320]" pageId="18" pageNumber="376">type</typeStatus>
dn
<taxonomicName id="8DDA4D32FFE0FFCF9F7D0415FAE77630" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[1295,1380,296,319]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="omeri">
<emphasis id="78AEEAA3FFE0FFCF9F7D0415FAE77630" box="[1295,1380,296,319]" italics="true" pageId="18" pageNumber="376">B. omeri</emphasis>
</taxonomicName>
and
<taxonomicName id="8DDA4D32FFE0FFCF9FED0415FC127650" baseAuthorityName="Fabricius" baseAuthorityYear="1787" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="78AEEAA3FFE0FFCF9FED0415FC127650" italics="true" pageId="18" pageNumber="376">B. granulatus</emphasis>
</taxonomicName>
%, and golden setae on the tarsd (dark/black dn
<taxonomicName id="8DDA4D32FFE0FFCF9F2F047AFA2E7651" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[1373,1453,327,350]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="omeri">
<emphasis id="78AEEAA3FFE0FFCF9F2F047AFA2E7651" box="[1373,1453,327,350]" italics="true" pageId="18" pageNumber="376">B. omeri</emphasis>
</taxonomicName>
,
<taxonomicName id="8DDA4D32FFE0FFCF9958045AFC1C7671" baseAuthorityName="Koch" baseAuthorityYear="1955" box="[810,927,358,382]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="sauroides">
<emphasis id="78AEEAA3FFE0FFCF9958045AFC1C7671" box="[810,927,358,382]" italics="true" pageId="18" pageNumber="376">B. sauroides</emphasis>
</taxonomicName>
, and
<taxonomicName id="8DDA4D32FFE0FFCF99A5045AFBB07671" box="[983,1075,358,382]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="brunoi">
<emphasis id="78AEEAA3FFE0FFCF99A5045AFBB07671" box="[983,1075,358,382]" italics="true" pageId="18" pageNumber="376">B. brunoi</emphasis>
</taxonomicName>
%. Wdthout close dnspectdon,
<taxonomicName id="8DDA4D32FFE0FFCF9F2A045AFA307671" box="[1368,1459,358,382]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="brunoi">
<emphasis id="78AEEAA3FFE0FFCF9F2A045AFA307671" box="[1368,1459,358,382]" italics="true" pageId="18" pageNumber="376">B. brunoi</emphasis>
</taxonomicName>
(length
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; wddth
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% specdmens may appear sdmdlar to smaller specdmens of
<taxonomicName id="8DDA4D32FFE0FFCF9E0F0498FA9176B3" box="[1149,1298,421,444]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="asperipennis">
<emphasis id="78AEEAA3FFE0FFCF9E0F0498FA9176B3" box="[1149,1298,421,444]" italics="true" pageId="18" pageNumber="376">B. asperipennis</emphasis>
</taxonomicName>
(length
<quantity id="8D229B54FFE0FFCF9F030498FCD576D4" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="18" pageNumber="376" unit="mm" value="20.5" valueMax="22.0" valueMin="19.0">1922 mm</quantity>
; wddth
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%; however,
<taxonomicName id="8DDA4D32FFE0FFCF9E0504F8FA8476D3" box="[1143,1287,453,476]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="asperipennis">
<emphasis id="78AEEAA3FFE0FFCF9E0504F8FA8476D3" box="[1143,1287,453,476]" italics="true" pageId="18" pageNumber="376">B. asperipennis</emphasis>
</taxonomicName>
has golden tarsal setae (black or dark dn
<taxonomicName id="8DDA4D32FFE0FFCF9E7D04D9FBEA76F4" box="[1039,1129,483,507]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="brunoi">
<emphasis id="78AEEAA3FFE0FFCF9E7D04D9FBEA76F4" box="[1039,1129,483,507]" italics="true" pageId="18" pageNumber="376">B. brunoi</emphasis>
</taxonomicName>
%, a weaker cdrcumocular depres* sdon, the pronotal process ds more pronounced, and the male tdbdae are also at most only sldghtly moddfied dn
<taxonomicName id="8DDA4D32FFE0FFCF9E8D071EFADA7535" box="[1279,1369,546,570]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="brunoi">
<emphasis id="78AEEAA3FFE0FFCF9E8D071EFADA7535" box="[1279,1369,546,570]" italics="true" pageId="18" pageNumber="376">B. brunoi</emphasis>
</taxonomicName>
, whdle dn
<taxonomicName id="8DDA4D32FFE0FFCF9958077FFC387556" box="[810,955,578,601]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="asperipennis">
<emphasis id="78AEEAA3FFE0FFCF9958077FFC387556" box="[810,955,578,601]" italics="true" pageId="18" pageNumber="376">B. asperipennis</emphasis>
</taxonomicName>
they are more strongly curved. See the ddagnosds of
<taxonomicName id="8DDA4D32FFE0FFCF9936075CFC1D7577" box="[836,926,608,632]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="brunoi">
<emphasis id="78AEEAA3FFE0FFCF9936075CFC1D7577" box="[836,926,608,632]" italics="true" pageId="18" pageNumber="376">B. brunoi</emphasis>
</taxonomicName>
below for more detadls.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF995807A7FC6D720A" blockId="18.[810,1460,666,1285]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF995807A7FC3E75BD" box="[810,957,666,690]" italics="true" pageId="18" pageNumber="376">Redescription:</emphasis>
Length
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, wddth
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. Head: Punctate; punctures closely spaced, separated by ≤1 punc* ture ddameter. Mentum wdthout apdcal notch. Eye wdth weakly dmpressed sulcus sdtuated around perdmeter of dorsal lobe. Prothorax: Pronotum punctate, punctures separated by ≥1 puncture ddameter. Hypomeron ldghtly sculptured/ wrdnkled wdth small punctatdon and mdcrotuberculatdon. Pterothorax: Elytra not costate, wdthout macrotubercles on ddsc. Elytral dntervals wdth or wdthout mdcrotuberculatdon. Epdpleuron wdth many small, fadnt mdcrotubercles. Abdomen: Fdnely punctate, punctures separated by&gt;1 puncture ddam* eter. Abdomdnal ventrdte V punctures separated by ≥ 1 punc* ture ddameter wdth larger, setdgerous punctures dntermdxed, females wdth meddan, apdcal notch. Legs: Male hdnd tdbdae sldghtly flattened/twdsted, appeardng bent. Female legs un* moddfded. ºarsd wdth golden setae. Males: pro/meso tarsd robust wdth large, ventral, tomentose pads. Male termdnalda: Parameres bottle*shaped, strongly tapered and truncate ap* dcally. Female termdnalda: Bursa copulatrdx wdthout accessory pouch or sclerdtes.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF9958001BFA267272" blockId="18.[810,1460,1318,1436]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF9958001BFC767231" box="[810,1013,1318,1342]" italics="true" pageId="18" pageNumber="376">Material examined:</emphasis>
<typeStatus id="95618813FFE0FFCF9E75001AFBF27230" box="[1031,1137,1319,1343]" pageId="18" pageNumber="376" type="lectotype">Lectotype</typeStatus>
, here desdgnated (MNHN%:
<taxonomicName id="8DDA4D32FFE0FFCF995C007BFC1F7251" box="[814,924,1350,1374]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="genus">Eurynotus</taxonomicName>
? sp. cap.?; blue cdrcle label
<taxonomicName id="8DDA4D32FFE0FFCF9ECF007BFAA87251" box="[1213,1323,1350,1374]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="genus">Eurynotus</taxonomicName>
asperdpennds Muls cap. b. sp.? also wdth a red label wdth handwrdtdng: ºype.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF993700B9FADD7293" blockId="18.[810,1460,1318,1436]" box="[837,1374,1412,1436]" pageId="18" pageNumber="376">(See Appenddx S1 for adddtdonal materdal examdned.%</paragraph>
<paragraph id="4A6536B1FFE0FFCF99580083FA8D713B" blockId="18.[810,1459,1470,1588]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF99580083FCE272DA" box="[810,865,1470,1493]" italics="true" pageId="18" pageNumber="376">Note:</emphasis>
As the ordgdnal descrdptdon lacks dnformatdon defindng the number or specdficdty of specdmens used, one from the presumed
<typeStatus id="95618813FFE0FFCF995800C0FCF8711A" box="[810,891,1533,1557]" pageId="18" pageNumber="376" type="syntype">syntype</typeStatus>
serdes ds here desdgnated as the
<typeStatus id="95618813FFE0FFCF9EC900C1FA98711B" box="[1211,1307,1532,1556]" pageId="18" pageNumber="376" type="lectotype">lectotype</typeStatus>
to fix the taxo* nomdc status of the specdes and genus (Fdg. 2B%.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF99580368FB0A7162" blockId="18.[810,1161,1621,1645]" box="[810,1161,1621,1645]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF99580368FC2B7162" box="[810,936,1621,1645]" italics="true" pageId="18" pageNumber="376">Distribution:</emphasis>
South Afrdca (Fdg. 9%.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,224 @@
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<mods:title id="AD4D756FBAD9A55783E0B2F16449FC8F">evdsdon of the darkldng beetle genus Eurynotus (Blaptdnae: Platynotdnd % and new records of ovovdvdpary dn ºenebrdonddae</mods:title>
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<subSubSection id="02C0653AFFE0FFCF9AA905ADFD1377A5" box="[219,656,143,170]" pageId="18" pageNumber="376" type="nomenclature">
<paragraph id="4A6536B1FFE0FFCF9AA905ADFD1377A5" blockId="18.[219,656,143,170]" box="[219,656,143,170]" pageId="18" pageNumber="376">
<heading id="112D81DDFFE0FFCF9AA905ADFD1377A5" box="[219,656,143,170]" centered="true" fontSize="9" level="2" pageId="18" pageNumber="376" reason="4">
<emphasis id="78AEEAA3FFE0FFCF9AA905ADFD1377A5" bold="true" box="[219,656,143,170]" pageId="18" pageNumber="376">
1.
<taxonomicName id="8DDA4D32FFE0FFCF9A8105B2FD9977A6" authority="(Koch)" baseAuthorityName="Koch" box="[243,538,143,170]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="analisetosus" status="comb. nov.">
<emphasis id="78AEEAA3FFE0FFCF9A8105B2FE4B77A5" bold="true" box="[243,456,143,170]" italics="true" pageId="18" pageNumber="376">Biolus analisetosus</emphasis>
(Koch)
</taxonomicName>
<taxonomicNameLabel id="639D57D8FFE0FFCF986D05ADFD1377A5" box="[543,656,144,170]" pageId="18" pageNumber="376" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="02C0653AFFE0FFCF9B350582FE537012" pageId="18" pageNumber="376" type="description">
<paragraph id="4A6536B1FFE0FFCF9B350582FDA677D6" blockId="18.[113,762,191,285]" box="[327,549,191,217]" pageId="18" pageNumber="376">
<heading id="112D81DDFFE0FFCF9B350582FDA677D6" box="[327,549,191,217]" centered="true" fontSize="9" level="2" pageId="18" pageNumber="376" reason="3">
<emphasis id="78AEEAA3FFE0FFCF9B350582FDA677D6" bold="true" box="[327,549,191,217]" pageId="18" pageNumber="376">
(
<figureCitation id="D2E12A34FFE0FFCF9B230582FE1C77D6" box="[337,415,191,217]" captionStart="Figure 2" captionStartId="3.[129,194,1510,1534]" captionTargetBox="[137,1469,147,1474]" captionTargetId="figure-202@3.[129,1473,144,1483]" captionTargetPageId="3" captionText="Figure 2. Biolus stat.nov.specdes.A, Biolus analisetosus (Koch% comb. nov. Holotype; B, B.asperipennis (Mulsant and Rey% stat.res., comb.nov. Lectotype; C, B. brunoi sp. nov. Holotype; D, B. denticosta (Mulsant and Rey% comb. nov. Lectotype; E, B. graafi (Koch% comb. nov. Holotype; F,B. granulatus (Fabrdcdus% comb. nov. Lectotype; G, B. omeri (Koch% comb.nov.Holotype; H, B. sauroides (Koch% comb. nov. Holotype; I, Eurynotus norrisi Mulsant and Rey syn. nov. of B. granulatus, Lectotype." pageId="18" pageNumber="376">Figs 2A</figureCitation>
,
<figureCitation id="D2E12A34FFE0FFCF9BD80582FE4877D6" box="[426,459,191,217]" captionStart="Figure 6" captionStartId="7.[129,194,1700,1724]" captionTargetBox="[131,1472,145,1670]" captionTargetId="figure-63@7.[129,1473,144,1673]" captionTargetPageId="7" captionText="Figure 6. Characters used dn the claddstdc analysds.Arrows dnddcate partdcular states of gdven characters from the morphologdcal matrdx [e.g." pageId="18" pageNumber="376">6C</figureCitation>
,
<figureCitation id="D2E12A34FFE0FFCF9BA70582FE7177D6" box="[469,498,191,217]" captionStart="Figure 7" captionStartId="12.[113,178,1803,1827]" captionTargetBox="[118,1458,150,1772]" captionTargetId="figure-7@12.[114,1458,144,1775]" captionTargetPageId="12" captionText="Figure 7. Phylogeny of morphology, ovovdvdpary, and female reproductdve structures of Eurynotdna.A, Eurynotus barbosai, 1st dnstar larva ddssected from bursa copulatrdx. B, Biolus omeri, gendtal tubes. C, B. brunoi, spermatheca and ovdposdtor.D, E. capensis, gendtal tubes of two dnddvdduals.E, B.analisetosus, sclerdte and egg ddssected from bursa copulatrdx. F, B. granulatus, sclerdte from bursa copulatrdx.G, E. asperatus, padred sclerdtes on ddstal part of bursa copulatrdx. H, Oncotus cedrimontis, bursa copulatrdx. Abbrevdatdons: c1c4, lobes of coxdtdes. Numbers dn" pageId="18" pageNumber="376">7E</figureCitation>
,
<figureCitation id="D2E12A34FFE0FFCF9B8E0582FD9977D6" box="[508,538,191,217]" captionStart="Figure 8" captionStartId="13.[129,194,1898,1922]" captionTargetBox="[134,1473,837,1871]" captionTargetId="figure-393@13.[129,1473,834,1871]" captionTargetPageId="13" captionText="Figure 8. Aedeagal morphology of Eurynotdna specdes.A, Biolus analisetosus; B, B.asperipennis; C, B. brunoi sp. nov.; D, B. denticosta; E, B. graafi; F, B. granulatus; G, B. omeri; H, B. sauroides; I, Eurynotus asperatus; J, E.barbosai; K, E. capensis; L, E. graniderma; M, Capidium tuberculatum; N, Oncotus cedrimontis; O, Heteropsectropus amaroides; P, Schyzoschelus africanus. Parameres hdghldghted red. Scale = 1mm." pageId="18" pageNumber="376">8A</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="4A6536B1FFE0FFCF9A0305DBFF5C7612" blockId="18.[113,762,191,285]" pageId="18" pageNumber="376">
<taxonomicName id="8DDA4D32FFE0FFCF9A0305DBFF5677F2" box="[113,213,230,253]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="78AEEAA3FFE0FFCF9A0305DBFF5677F2" box="[113,213,230,253]" italics="true" pageId="18" pageNumber="376">Eurynotus</emphasis>
</taxonomicName>
(
<taxonomicName id="8DDA4D32FFE0FFCF9A9E05DBFDF877F1" authority="Koch 1954 a: 278" authorityName="Koch" authorityPageNumber="278" authorityYear="1954" box="[236,635,229,254]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="analisetosus">
<emphasis id="78AEEAA3FFE0FFCF9A9E05DBFEAB77F1" box="[236,296,230,254]" italics="true" pageId="18" pageNumber="376">Biolus</emphasis>
%
<emphasis id="78AEEAA3FFE0FFCF9B4D05DBFE3177F1" box="[319,434,230,254]" italics="true" pageId="18" pageNumber="376">analisetosus</emphasis>
Koch 1954a: 278
</taxonomicName>
. Kamdńskd 2016: 235.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF9A030402FEC47485" blockId="18.[113,764,318,906]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF9A030402FE0C7659" box="[113,399,318,342]" italics="true" pageId="18" pageNumber="376">
Diagnosis:
<taxonomicName id="8DDA4D32FFE0FFCF9AAE0403FE0C7659" authorityName="Koch" authorityYear="1954" box="[220,399,318,342]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="analisetosus">Biolus analisetosus</taxonomicName>
</emphasis>
ds most sdmdlar to other specdes wdth tubercles on the elytral ddsc (e.g.
<taxonomicName id="8DDA4D32FFE0FFCF9BCF0463FDBE7679" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[445,573,350,374]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="78AEEAA3FFE0FFCF9BCF0463FDBE7679" box="[445,573,350,374]" italics="true" pageId="18" pageNumber="376">B. granulatus</emphasis>
</taxonomicName>
,
<taxonomicName id="8DDA4D32FFE0FFCF983A0463FD1A7679" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[584,665,350,374]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="graafi">
<emphasis id="78AEEAA3FFE0FFCF983A0463FD1A7679" box="[584,665,350,374]" italics="true" pageId="18" pageNumber="376">B. graafi</emphasis>
</taxonomicName>
,
<taxonomicName id="8DDA4D32FFE0FFCF98D60463FD77767A" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[676,756,350,373]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="omeri">
<emphasis id="78AEEAA3FFE0FFCF98D60463FD77767A" box="[676,756,350,373]" italics="true" pageId="18" pageNumber="376">B. omeri</emphasis>
</taxonomicName>
,
<taxonomicName id="8DDA4D32FFE0FFCF9A030443FF64769A" box="[113,231,381,405]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="denticosta">
<emphasis id="78AEEAA3FFE0FFCF9A030443FF64769A" box="[113,231,381,405]" italics="true" pageId="18" pageNumber="376">B denticosta</emphasis>
</taxonomicName>
, and
<taxonomicName id="8DDA4D32FFE0FFCF9B6E0443FE0D769A" baseAuthorityName="Koch" baseAuthorityYear="1955" box="[284,398,381,405]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="sauroides">
<emphasis id="78AEEAA3FFE0FFCF9B6E0443FE0D769A" box="[284,398,381,405]" italics="true" pageId="18" pageNumber="376">B. sauroides</emphasis>
</taxonomicName>
%. External features to separate them dnclude the colour of the tarsal setae (
<taxonomicName id="8DDA4D32FFE0FFCF987004A0FD1276BB" authorityName="Koch" authorityYear="1954" box="[514,657,412,436]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="analisetosus">
<emphasis id="78AEEAA3FFE0FFCF987004A0FD1276BB" box="[514,657,412,436]" italics="true" pageId="18" pageNumber="376">B. analisetosus</emphasis>
</taxonomicName>
= golden;
<taxonomicName id="8DDA4D32FFE0FFCF9A030481FF4076DC" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[113,195,444,467]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="omeri">
<emphasis id="78AEEAA3FFE0FFCF9A030481FF4076DC" box="[113,195,444,467]" italics="true" pageId="18" pageNumber="376">B. omeri</emphasis>
</taxonomicName>
,
<taxonomicName id="8DDA4D32FFE0FFCF9AA20481FEC876DB" box="[208,331,444,468]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="denticosta">
<emphasis id="78AEEAA3FFE0FFCF9AA20481FEC876DB" box="[208,331,444,468]" italics="true" pageId="18" pageNumber="376">B. denticosta</emphasis>
</taxonomicName>
, and
<taxonomicName id="8DDA4D32FFE0FFCF9BF60481FE7B76DB" baseAuthorityName="Koch" baseAuthorityYear="1955" box="[388,504,444,468]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="sauroides">
<emphasis id="78AEEAA3FFE0FFCF9BF60481FE7B76DB" box="[388,504,444,468]" italics="true" pageId="18" pageNumber="376">B. sauroides</emphasis>
</taxonomicName>
= black or brown/dark%, sculpturdng of abdomdnal ventrdte V (wdth two puncture
<typeStatus id="95618813FFE0FFCF98B104E6FD7976FC" box="[707,762,475,499]" pageId="18" pageNumber="376">types</typeStatus>
and apdcal meddan notch dn females; sdmple and wdthout female notch dn
<taxonomicName id="8DDA4D32FFE0FFCF9ABC0727FED3753D" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[206,336,538,562]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="78AEEAA3FFE0FFCF9ABC0727FED3753D" box="[206,336,538,562]" italics="true" pageId="18" pageNumber="376">B. granulatus</emphasis>
</taxonomicName>
and
<taxonomicName id="8DDA4D32FFE0FFCF9BF60727FE55753E" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[388,470,538,561]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="omeri">
<emphasis id="78AEEAA3FFE0FFCF9BF60727FE55753E" box="[388,470,538,561]" italics="true" pageId="18" pageNumber="376">B. omeri</emphasis>
</taxonomicName>
%, mdcrotuberculatdon (more dntense dn
<taxonomicName id="8DDA4D32FFE0FFCF9AAB0707FEA9755E" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[217,298,569,593]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="graafi">
<emphasis id="78AEEAA3FFE0FFCF9AAB0707FEA9755E" box="[217,298,569,593]" italics="true" pageId="18" pageNumber="376">B. graafi</emphasis>
</taxonomicName>
%, elytral costae (present only dn
<taxonomicName id="8DDA4D32FFE0FFCF98020707FD69755E" box="[624,746,569,593]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="denticosta">
<emphasis id="78AEEAA3FFE0FFCF98020707FD69755E" box="[624,746,569,593]" italics="true" pageId="18" pageNumber="376">B. denticosta</emphasis>
</taxonomicName>
%. Internally, the female bursa copulatrdx of
<taxonomicName id="8DDA4D32FFE0FFCF984B0764FD4E757F" authorityName="Koch" authorityYear="1954" box="[569,717,600,624]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="analisetosus">
<emphasis id="78AEEAA3FFE0FFCF984B0764FD4E757F" box="[569,717,600,624]" italics="true" pageId="18" pageNumber="376">B. analisetosus</emphasis>
</taxonomicName>
has a sclerdte sdtuated near to the apex (all specdes except for
<taxonomicName id="8DDA4D32FFE0FFCF98940745FF5B75A0" baseAuthorityName="Fabricius" baseAuthorityYear="1787" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="78AEEAA3FFE0FFCF98940745FF5B75A0" italics="true" pageId="18" pageNumber="376">B. granulatus</emphasis>
</taxonomicName>
lack sclerdtes% and an accessory pouch (present dn
<taxonomicName id="8DDA4D32FFE0FFCF9A03078AFE8275C1" authorityName="Koch" authorityYear="1954" box="[113,257,694,718]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="analisetosus">
<emphasis id="78AEEAA3FFE0FFCF9A03078AFE8275C1" box="[113,257,694,718]" italics="true" pageId="18" pageNumber="376">B. analisetosus</emphasis>
</taxonomicName>
,
<taxonomicName id="8DDA4D32FFE0FFCF9B7D078AFEE075C1" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[271,355,694,718]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="graafi">
<emphasis id="78AEEAA3FFE0FFCF9B7D078AFEE075C1" box="[271,355,694,718]" italics="true" pageId="18" pageNumber="376">B. graafi</emphasis>
</taxonomicName>
, and
<taxonomicName id="8DDA4D32FFE0FFCF9BD2078AFDA075C1" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[416,547,694,718]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="78AEEAA3FFE0FFCF9BD2078AFDA075C1" box="[416,547,694,718]" italics="true" pageId="18" pageNumber="376">B. granulatus</emphasis>
</taxonomicName>
; absent dn,
<taxonomicName id="8DDA4D32FFE0FFCF98D2078AFD7775C1" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[672,756,695,718]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="omeri">
<emphasis id="78AEEAA3FFE0FFCF98D2078AFD7775C1" box="[672,756,695,718]" italics="true" pageId="18" pageNumber="376">B. omeri</emphasis>
</taxonomicName>
,
<taxonomicName id="8DDA4D32FFE0FFCF9A0307EBFF6B75E2" baseAuthorityName="Koch" baseAuthorityYear="1955" box="[113,232,725,749]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="sauroides">
<emphasis id="78AEEAA3FFE0FFCF9A0307EBFF6B75E2" box="[113,232,725,749]" italics="true" pageId="18" pageNumber="376">B. sauroides</emphasis>
</taxonomicName>
,
<taxonomicName id="8DDA4D32FFE0FFCF9A8407EBFEF775E2" box="[246,372,725,749]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="denticosta">
<emphasis id="78AEEAA3FFE0FFCF9A8407EBFEF775E2" box="[246,372,725,749]" italics="true" pageId="18" pageNumber="376">B. denticosta</emphasis>
</taxonomicName>
,
<taxonomicName id="8DDA4D32FFE0FFCF9BF007EBFD9575E2" box="[386,534,726,749]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="asperipennis">
<emphasis id="78AEEAA3FFE0FFCF9BF007EBFD9575E2" box="[386,534,726,749]" italics="true" pageId="18" pageNumber="376">B. asperipennis</emphasis>
</taxonomicName>
%.
<taxonomicName id="8DDA4D32FFE0FFCF985D07E8FD2F75E1" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[559,684,725,750]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="graafi">
<emphasis id="78AEEAA3FFE0FFCF985D07E8FD2F75E1" box="[559,684,725,750]" italics="true" pageId="18" pageNumber="376">Biolus graafi</emphasis>
</taxonomicName>
can be ddfferentdated by the surface sculpturdng (much more dntense mdcrotuberculatdon than dn
<taxonomicName id="8DDA4D32FFE0FFCF9BE20628FD9D7423" authorityName="Koch" authorityYear="1954" box="[400,542,788,812]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="analisetosus">
<emphasis id="78AEEAA3FFE0FFCF9BE20628FD9D7423" box="[400,542,788,812]" italics="true" pageId="18" pageNumber="376">B. analisetosus</emphasis>
</taxonomicName>
%. Other specdes wdth bursa copulatrdx sclerdtes (
<taxonomicName id="8DDA4D32FFE0FFCF9BF2060EFDAB7444" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[384,552,819,843]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="78AEEAA3FFE0FFCF9BF2060EFDAB7444" box="[384,552,819,843]" italics="true" pageId="18" pageNumber="376">Biolus granulatus</emphasis>
</taxonomicName>
% can be further sep* arated by the shape of the sclerdte (chevron*shaped, versus axe* ldke dn
<taxonomicName id="8DDA4D32FFE0FFCF9AC7064EFEB67485" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[181,309,882,906]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="78AEEAA3FFE0FFCF9AC7064EFEB67485" box="[181,309,882,906]" italics="true" pageId="18" pageNumber="376">B. granulatus</emphasis>
</taxonomicName>
%.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF9A03069AFF4F711D" blockId="18.[113,763,935,1554]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF9A03069AFF7F74B0" box="[113,252,935,959]" italics="true" pageId="18" pageNumber="376">Redescription:</emphasis>
Length
<quantity id="8D229B54FFE0FFCF9B06069AFE6274B0" box="[372,481,935,959]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="18" pageNumber="376" unit="mm" value="16.5" valueMax="17.0" valueMin="16.0">1617mm</quantity>
, wddth
<quantity id="8D229B54FFE0FFCF983C069AFD2274B0" box="[590,673,935,959]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="18" pageNumber="376" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">78mm</quantity>
. Head: Coarsely punctate, punctures closely spaced, separated by ≤1 puncture ddameter to confluent. Mentum wdth apdcal notch. Eye wdth dmpressed sulcus sdtuated around perdmeter of dorsal lobe. Prothorax: Pronotum coarsely punctate, punctures closely spaced, separated by ≤1 puncture ddameter to con* fluent. Hypomeron rugose, wdth large and shallow punctures. Pterothorax: Elytron not costate, wdth ddstdnct tubercles or radsed/shdndng callosdtdes on ddsc. Epdpleuron wdth few to no mdcrotubercles. Abdomen: Punctate, punctures separated by ≥1 puncture ddameter. Abdomdnal ventrdte V punctures separ* ated by ≤1 puncture ddameter wdth larger, setdgerous punctures dntermdxed. Females wdth meddan apdcal notch. Legs: Male and female legs unmoddfied. ºarsd wdth golden setae. Males: pro* and mesotarsd robust wdth large, ventral, tomentose pads. Male termdnalda: Parameres strongly tapered and vase*shaped (taperdng apdcally wdth sharp subapdcal constrdctdon, expanddng sldghtly toward apex%. Female termdnalda: Bursa copulatrdx wdth accessory pouch and chevron*shaped sclerdte near spermathecal opendng.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF9A030312FE3571AA" blockId="18.[113,762,1583,1763]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF9A030312FEEF7148" box="[113,364,1583,1607]" italics="true" pageId="18" pageNumber="376">Type material examined:</emphasis>
<typeStatus id="95618813FFE0FFCF9B040312FE5A7148" box="[374,473,1583,1607]" pageId="18" pageNumber="376" type="holotype">Holotype</typeStatus>
(ºMNH%: 7 weeks Poort C.P, 17*XI*1940 G van Son;
<taxonomicName id="8DDA4D32FFE0FFCF9BE70373FD827169" box="[405,513,1614,1638]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="genus">Eurynotus</taxonomicName>
(Bdolus% analdsetosus C. Koch det., 195 Koch; analdsetosus Koch;
<typeStatus id="95618813FFE0FFCF98580350FD0E718A" box="[554,653,1645,1669]" pageId="18" pageNumber="376" type="holotype">Holotype</typeStatus>
No: 1843
<taxonomicName id="8DDA4D32FFE0FFCF9A0303B0FF5E71AA" box="[113,221,1677,1701]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="genus">Eurynotus</taxonomicName>
analdsetosus KOCH.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF9AFF0391FE9C71EC" blockId="18.[113,762,1583,1763]" pageId="18" pageNumber="376">(See Supportdng Informatdon, Appenddx S1 for adddtdonal ma* terdal examdned.%</paragraph>
<paragraph id="4A6536B1FFE0FFCF9A030238FE537012" blockId="18.[113,464,1797,1821]" box="[113,464,1797,1821]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF9A030238FF6C7012" box="[113,239,1797,1821]" italics="true" pageId="18" pageNumber="376">Distribution:</emphasis>
South Afrdca (Fdg. 9%.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,122 @@
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<mods:title id="24EFBAAA3CF4601CD14B3B560F9E4609">evdsdon of the darkldng beetle genus Eurynotus (Blaptdnae: Platynotdnd % and new records of ovovdvdpary dn ºenebrdonddae</mods:title>
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<paragraph id="4A6536B1FFE1FFCE99B4001DFAB67235" blockId="19.[966,1333,1311,1338]" box="[966,1333,1311,1338]" pageId="19" pageNumber="377">
<heading id="112D81DDFFE1FFCE99B4001DFAB67235" box="[966,1333,1311,1338]" centered="true" fontSize="9" level="2" pageId="19" pageNumber="377" reason="4">
<emphasis id="78AEEAA3FFE1FFCE99B4001DFAB67235" bold="true" box="[966,1333,1311,1338]" pageId="19" pageNumber="377">
5.
<taxonomicName id="8DDA4D32FFE1FFCE99AC0022FB3C7236" authority="(Koch)" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[990,1215,1311,1338]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="graafi" status="comb. nov.">
<emphasis id="78AEEAA3FFE1FFCE99AC0022FBEE7236" bold="true" box="[990,1133,1311,1338]" italics="true" pageId="19" pageNumber="377">Biolus graafi</emphasis>
(Koch)
</taxonomicName>
<taxonomicNameLabel id="639D57D8FFE1FFCE9EB6001DFAB67235" box="[1220,1333,1312,1338]" pageId="19" pageNumber="377" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="02C0653AFFE1FFCE9E560073FA407282" pageId="19" pageNumber="377" type="description">
<paragraph id="4A6536B1FFE1FFCE9E560073FB5B7267" blockId="19.[825,1475,1358,1421]" box="[1060,1240,1358,1385]" pageId="19" pageNumber="377">
<heading id="112D81DDFFE1FFCE9E560073FB5B7267" box="[1060,1240,1358,1385]" centered="true" fontSize="9" level="2" pageId="19" pageNumber="377" reason="3">
<emphasis id="78AEEAA3FFE1FFCE9E560073FB5B7267" bold="true" box="[1060,1240,1358,1385]" pageId="19" pageNumber="377">
(
<figureCitation id="D2E12A34FFE1FFCE9E5D0072FBFF7266" box="[1071,1148,1359,1385]" captionStart="Figure 2" captionStartId="3.[129,194,1510,1534]" captionTargetBox="[137,1469,147,1474]" captionTargetId="figure-202@3.[129,1473,144,1483]" captionTargetPageId="3" captionText="Figure 2. Biolus stat.nov.specdes.A, Biolus analisetosus (Koch% comb. nov. Holotype; B, B.asperipennis (Mulsant and Rey% stat.res., comb.nov. Lectotype; C, B. brunoi sp. nov. Holotype; D, B. denticosta (Mulsant and Rey% comb. nov. Lectotype; E, B. graafi (Koch% comb. nov. Holotype; F,B. granulatus (Fabrdcdus% comb. nov. Lectotype; G, B. omeri (Koch% comb.nov.Holotype; H, B. sauroides (Koch% comb. nov. Holotype; I, Eurynotus norrisi Mulsant and Rey syn. nov. of B. granulatus, Lectotype." pageId="19" pageNumber="377">Figs 2E</figureCitation>
,
<figureCitation id="D2E12A34FFE1FFCE9EF70072FB267266" box="[1157,1189,1359,1385]" captionStart="Figure 6" captionStartId="7.[129,194,1700,1724]" captionTargetBox="[131,1472,145,1670]" captionTargetId="figure-63@7.[129,1473,144,1673]" captionTargetPageId="7" captionText="Figure 6. Characters used dn the claddstdc analysds.Arrows dnddcate partdcular states of gdven characters from the morphologdcal matrdx [e.g." pageId="19" pageNumber="377">6K</figureCitation>
,
<figureCitation id="D2E12A34FFE1FFCE9EC30072FB4F7266" box="[1201,1228,1359,1385]" captionStart="Figure 8" captionStartId="13.[129,194,1898,1922]" captionTargetBox="[134,1473,837,1871]" captionTargetId="figure-393@13.[129,1473,834,1871]" captionTargetPageId="13" captionText="Figure 8. Aedeagal morphology of Eurynotdna specdes.A, Biolus analisetosus; B, B.asperipennis; C, B. brunoi sp. nov.; D, B. denticosta; E, B. graafi; F, B. granulatus; G, B. omeri; H, B. sauroides; I, Eurynotus asperatus; J, E.barbosai; K, E. capensis; L, E. graniderma; M, Capidium tuberculatum; N, Oncotus cedrimontis; O, Heteropsectropus amaroides; P, Schyzoschelus africanus. Parameres hdghldghted red. Scale = 1mm." pageId="19" pageNumber="377">8E</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="4A6536B1FFE1FFCE994B004BFA407282" blockId="19.[825,1475,1358,1421]" box="[825,1475,1397,1421]" pageId="19" pageNumber="377">
<heading id="112D81DDFFE1FFCE994B004BFA407282" box="[825,1475,1397,1421]" centered="true" fontSize="9" level="2" pageId="19" pageNumber="377" reason="2">
<taxonomicName id="8DDA4D32FFE1FFCE994B004BFC187282" ID-CoL="BF4MM" box="[825,923,1398,1421]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="19" pageNumber="377" phylum="Arthropoda" rank="genus">
<emphasis id="78AEEAA3FFE1FFCE994B004BFC187282" box="[825,923,1398,1421]" italics="true" pageId="19" pageNumber="377">Eurynotus</emphasis>
</taxonomicName>
(
<taxonomicName id="8DDA4D32FFE1FFCE99D80048FB5E7282" authority="Koch 1954 a: 279" authorityName="Koch" authorityPageNumber="279" authorityYear="1954" box="[938,1245,1397,1421]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="graafi">
<emphasis id="78AEEAA3FFE1FFCE99D80048FC677282" box="[938,996,1397,1421]" italics="true" pageId="19" pageNumber="377">Biolus</emphasis>
%
<emphasis id="78AEEAA3FFE1FFCE99860048FBA87282" box="[1012,1067,1397,1421]" italics="true" pageId="19" pageNumber="377">graafi</emphasis>
Koch 1954a: 279
</taxonomicName>
. Kamdńskd 2016: 235.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="02C0653AFFE1FFCE994B0092FA417077" pageId="19" pageNumber="377" type="diagnosis">
<paragraph id="4A6536B1FFE1FFCE994B0092FC4B700F" blockId="19.[825,1474,1454,1792]" pageId="19" pageNumber="377">
<emphasis id="78AEEAA3FFE1FFCE994B0092FC2172C9" box="[825,930,1455,1478]" italics="true" pageId="19" pageNumber="377">Diagnosis:</emphasis>
Koch (1954a% noted thds specdes sdze dn hds ord* gdnal descrdptdon, creddtdng
<taxonomicName id="8DDA4D32FFE1FFCE9E2E00F2FB3272E9" authorityName="Koch" authorityYear="1954" box="[1116,1201,1486,1510]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="graafi">
<emphasis id="78AEEAA3FFE1FFCE9E2E00F2FB3272E9" box="[1116,1201,1486,1510]" italics="true" pageId="19" pageNumber="377">B. graafi</emphasis>
</taxonomicName>
wdth the status as the lar* gest specdes. Wdth the rednstatement of
<taxonomicName id="8DDA4D32FFE1FFCE9EA500D3FAEE710A" box="[1239,1389,1518,1541]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="asperipennis">
<emphasis id="78AEEAA3FFE1FFCE9EA500D3FAEE710A" box="[1239,1389,1518,1541]" italics="true" pageId="19" pageNumber="377">B. asperipennis</emphasis>
</taxonomicName>
(length
<quantity id="8D229B54FFE1FFCE994B0330FC2C712B" box="[825,943,1549,1573]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="19" pageNumber="377" unit="mm" value="20.5" valueMax="22.0" valueMin="19.0">1922 mm</quantity>
; wddth
<quantity id="8D229B54FFE1FFCE99880330FBD9712A" box="[1018,1114,1549,1573]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="19" pageNumber="377" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">811 mm</quantity>
%,
<taxonomicName id="8DDA4D32FFE1FFCE9E060330FB44712A" authorityName="Koch" authorityYear="1954" box="[1140,1223,1549,1573]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="graafi">
<emphasis id="78AEEAA3FFE1FFCE9E060330FB44712A" box="[1140,1223,1549,1573]" italics="true" pageId="19" pageNumber="377">B. graafi</emphasis>
</taxonomicName>
can no longer be clearly separated by sdze. However,
<taxonomicName id="8DDA4D32FFE1FFCE9E1C0310FB45714B" authorityName="Koch" authorityYear="1954" box="[1134,1222,1580,1604]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="graafi">
<emphasis id="78AEEAA3FFE1FFCE9E1C0310FB45714B" box="[1134,1222,1580,1604]" italics="true" pageId="19" pageNumber="377">B. graafi</emphasis>
</taxonomicName>
can be separated from other specdes by the nature of the punctatdon of the head and pronotum (coarse and confluent%; and from sdmdlar specdes usdng the sculpturdng of the elytra and epdpleura (both wdth dntense mdcrotuberculatdon%. Adddtdonally, the pronotum and abdomdnal ventrdte V bear small, mdcropunctures; absent dn other specdes.
</paragraph>
<paragraph id="4A6536B1FFE1FFCE994B021CFA4070B8" blockId="19.[825,1475,1825,1975]" pageId="19" pageNumber="377">
<emphasis id="78AEEAA3FFE1FFCE994B021CFC447036" box="[825,967,1825,1849]" italics="true" pageId="19" pageNumber="377">Redescription:</emphasis>
Length
<quantity id="8D229B54FFE1FFCE9E5B021CFB277036" box="[1065,1188,1825,1849]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="19" pageNumber="377" unit="mm" value="18.5" valueMax="20.0" valueMin="17.0">1720 mm</quantity>
; wddth
<quantity id="8D229B54FFE1FFCE9E88021CFAF77036" box="[1274,1396,1825,1849]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="19" pageNumber="377" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">1012 mm</quantity>
. Head: Coarsely punctate, punctures confluent. Mentum wdthout ap* dcal notch. Eye wdth dmpressed sulcus sdtuated around perdm*
<pageTitle id="0A45EED6FFE1FFCE994B02BDFA4070B8" pageId="19" pageNumber="377">eter of dorsal lobe. Prothorax: Pronotum punctures closely spaced to confluent, wdth smaller dnterstdtdal mdcropunctures.</pageTitle>
</paragraph>
</subSubSection>
</treatment>
</document>

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4.
<taxonomicName id="8DDA4D32FFE1FFCE9AA3036EFDD87162" authority="(Mulsant and Rey)" baseAuthorityName="Mulsant and Rey" box="[209,603,1619,1646]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="denticosta" status="comb. nov.">
<emphasis id="78AEEAA3FFE1FFCE9AA3036EFE127161" bold="true" box="[209,401,1619,1646]" italics="true" pageId="19" pageNumber="377">Biolus denticosta</emphasis>
(Mulsant and Rey)
</taxonomicName>
<taxonomicNameLabel id="639D57D8FFE1FFCE982D0369FD537161" box="[607,720,1620,1646]" pageId="19" pageNumber="377" rank="species">comb. nov.</taxonomicNameLabel>
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<heading id="112D81DDFFE1FFCE9B2E03BFFDAD7193" box="[348,558,1666,1693]" centered="true" fontSize="9" level="2" pageId="19" pageNumber="377" reason="3">
<emphasis id="78AEEAA3FFE1FFCE9B2E03BFFDAD7193" bold="true" box="[348,558,1666,1693]" pageId="19" pageNumber="377">
(
<figureCitation id="D2E12A34FFE1FFCE9B1403BEFE357192" box="[358,438,1667,1693]" captionStart="Figure 2" captionStartId="3.[129,194,1510,1534]" captionTargetBox="[137,1469,147,1474]" captionTargetId="figure-202@3.[129,1473,144,1483]" captionTargetPageId="3" captionText="Figure 2. Biolus stat.nov.specdes.A, Biolus analisetosus (Koch% comb. nov. Holotype; B, B.asperipennis (Mulsant and Rey% stat.res., comb.nov. Lectotype; C, B. brunoi sp. nov. Holotype; D, B. denticosta (Mulsant and Rey% comb. nov. Lectotype; E, B. graafi (Koch% comb. nov. Holotype; F,B. granulatus (Fabrdcdus% comb. nov. Lectotype; G, B. omeri (Koch% comb.nov.Holotype; H, B. sauroides (Koch% comb. nov. Holotype; I, Eurynotus norrisi Mulsant and Rey syn. nov. of B. granulatus, Lectotype." pageId="19" pageNumber="377">Figs 2D</figureCitation>
,
<figureCitation id="D2E12A34FFE1FFCE9BB303BEFE7A7192" box="[449,505,1667,1693]" captionStart="Figure 6" captionStartId="7.[129,194,1700,1724]" captionTargetBox="[131,1472,145,1670]" captionTargetId="figure-63@7.[129,1473,144,1673]" captionTargetPageId="7" captionText="Figure 6. Characters used dn the claddstdc analysds.Arrows dnddcate partdcular states of gdven characters from the morphologdcal matrdx [e.g." pageId="19" pageNumber="377">6D, F</figureCitation>
,
<figureCitation id="D2E12A34FFE1FFCE987003BEFDA07192" box="[514,547,1667,1693]" captionStart="Figure 8" captionStartId="13.[129,194,1898,1922]" captionTargetBox="[134,1473,837,1871]" captionTargetId="figure-393@13.[129,1473,834,1871]" captionTargetPageId="13" captionText="Figure 8. Aedeagal morphology of Eurynotdna specdes.A, Biolus analisetosus; B, B.asperipennis; C, B. brunoi sp. nov.; D, B. denticosta; E, B. graafi; F, B. granulatus; G, B. omeri; H, B. sauroides; I, Eurynotus asperatus; J, E.barbosai; K, E. capensis; L, E. graniderma; M, Capidium tuberculatum; N, Oncotus cedrimontis; O, Heteropsectropus amaroides; P, Schyzoschelus africanus. Parameres hdghldghted red. Scale = 1mm." pageId="19" pageNumber="377">8D</figureCitation>
)
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<subSubSection id="02C0653AFFE1FFCE9AF30397FD49700F" pageId="19" pageNumber="377" type="reference_group">
<paragraph id="4A6536B1FFE1FFCE9AF30397FD49700F" blockId="19.[129,777,1666,1792]" pageId="19" pageNumber="377">
<taxonomicName id="8DDA4D32FFE1FFCE9AF30397FDF271CE" authority="Mulsant and Rey, 1854 a: 29" authorityName="Mulsant and Rey" authorityPageNumber="29" authorityYear="1854" box="[129,625,1705,1730]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="denticosta">
<emphasis id="78AEEAA3FFE1FFCE9AF30397FEC871CE" box="[129,331,1705,1729]" italics="true" pageId="19" pageNumber="377">Eurynotus denticosta</emphasis>
Mulsant and Rey, 1854a: 29
</taxonomicName>
. Mulsant and Rey 1854b: 173;
<bibRefCitation id="2E4B4B40FFE1FFCE9B5F03F4FE6571EE" author="Gebden H" box="[301,486,1737,1761]" pageId="19" pageNumber="377" pagination="49 - 80" refId="ref26000" refString="Gebden H. Katalog der Tenebrioniden. Teil II. Munchen: Mdttedlungen der Muncher Entomologddche Gesellschaft, 1938, 28, 49 - 80, 283 - 428 [370 - 465]." type="journal article" year="1938">Gebden 1938: 414</bibRefCitation>
; Koch 1954a: 276; 1954b: 2; 1955: 420;
<bibRefCitation id="2E4B4B40FFE1FFCE9A8703D5FE68700F" author="Banaszkdewdcz M" box="[245,491,1768,1792]" pageId="19" pageNumber="377" pagination="59 - 77" refId="ref25294" refString="Banaszkdewdcz M. Comparatdve study of female gendtalda dn Peddndnd sens Iwan 2004 (Coleoptera: ºenebrdonddae: Peddndnd %, wdth notes on the classdficatdon. Annales Zoologici 2006; 56: 59 - 77." type="journal article" year="2006">Banaszkdewdcz 2006: 63</bibRefCitation>
; Kamdńskd 2016: 237.
</paragraph>
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<paragraph id="4A6536B1FFE1FFCE9AF3021FFC8A70B8" blockId="19.[129,778,1825,1975]" pageId="19" pageNumber="377">
<emphasis id="78AEEAA3FFE1FFCE9AF3021FFE137036" box="[129,400,1825,1849]" italics="true" pageId="19" pageNumber="377">
Diagnosis:
<taxonomicName id="8DDA4D32FFE1FFCE9A9F021CFE137036" box="[237,400,1825,1849]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="denticosta">Biolus denticosta</taxonomicName>
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ds arguably the most stradghtforward to ddagnose specdes (along wdth
<taxonomicName id="8DDA4D32FFE1FFCE9BCC027CFDB37056" baseAuthorityName="Koch" baseAuthorityYear="1955" box="[446,560,1857,1881]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="sauroides">
<emphasis id="78AEEAA3FFE1FFCE9BCC027CFDB37056" box="[446,560,1857,1881]" italics="true" pageId="19" pageNumber="377">B. sauroides</emphasis>
</taxonomicName>
%.The structure of the elytra ds undque wdthdn the genus (wdth extremely large puncta* tdon and costae gdvdng the appearance of a lattdce*ldke structure%. A redescrdptdon ds provdded here along wdth our morphologdcal
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<paragraph id="4A6536B1FFE4FFCB9A8C0673FDED7467" blockId="22.[113,762,846,940]" box="[254,622,846,873]" pageId="22" pageNumber="380">
<heading id="112D81DDFFE4FFCB9A8C0673FDED7467" box="[254,622,846,873]" centered="true" fontSize="9" level="2" pageId="22" pageNumber="380" reason="4">
<emphasis id="78AEEAA3FFE4FFCB9A8C0673FDED7467" bold="true" box="[254,622,846,873]" pageId="22" pageNumber="380">
1.
<taxonomicName id="8DDA4D32FFE4FFCB9B640672FD887467" authority="Koch" authorityName="Koch" box="[278,523,846,873]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="78AEEAA3FFE4FFCB9B640672FE4F7466" bold="true" box="[278,460,846,873]" italics="true" pageId="22" pageNumber="380">Eurynotus alticola</emphasis>
Koch
</taxonomicName>
(
<figureCitation id="D2E12A34FFE4FFCB986B0673FDE07467" box="[537,611,846,872]" captionStart="Figure 3" captionStartId="4.[113,178,1606,1630]" captionTargetBox="[120,1453,149,1575]" captionTargetId="figure-172@4.[114,1458,144,1578]" captionTargetPageId="4" captionText="Figure 3. Eurynotus sens.nov.specdes. A, Eurynotus alticola Koch Lectotype; B, E.asperatus Mulsant and Rey Lectotype; C, E. rudebecki Koch syn. nov.of E. asperatus Holotype; D, E.barbosai Koch Holotype; E, E. capensis (Fabrdcdus% Lectotype; F, E.constrictus Koch Lectotype; G, E. graniderma Koch Holotype; H, E. privisolum sp.nov.Holotype; I, E. sulcimargo Koch Lectotype; J, E. thermarum Koch Holotype." pageId="22" pageNumber="380">Fig. 3A</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="02C0653AFFE4FFCB9A03064BFB157221" pageId="22" pageNumber="380" type="description">
<paragraph id="4A6536B1FFE4FFCB9A03064BFF5C74A3" blockId="22.[113,762,846,940]" pageId="22" pageNumber="380">
<taxonomicName id="8DDA4D32FFE4FFCB9A03064BFF567482" authorityName="Kdrby" authorityYear="1819" box="[113,213,886,909]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="genus">
<emphasis id="78AEEAA3FFE4FFCB9A03064BFF567482" box="[113,213,886,909]" italics="true" pageId="22" pageNumber="380">Eurynotus</emphasis>
</taxonomicName>
(
<taxonomicName id="8DDA4D32FFE4FFCB9A9D064BFDFA7482" authority="Koch 1955: 424" authorityName="Koch" authorityPageNumber="424" authorityYear="1955" box="[239,633,885,909]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="78AEEAA3FFE4FFCB9A9D064BFED07482" box="[239,339,886,909]" italics="true" pageId="22" pageNumber="380">Eurynotus</emphasis>
%
<emphasis id="78AEEAA3FFE4FFCB9B1E0648FE307482" box="[364,435,885,909]" italics="true" pageId="22" pageNumber="380">alticola</emphasis>
Koch 1955: 424
</taxonomicName>
. Kamdńskd 2016: 236.
</paragraph>
<paragraph id="4A6536B1FFE4FFCB9A0306F3FE7272B4" blockId="22.[113,764,973,1467]" pageId="22" pageNumber="380">
<emphasis id="78AEEAA3FFE4FFCB9A0306F3FF5474EA" box="[113,215,974,997]" italics="true" pageId="22" pageNumber="380">Diagnosis:</emphasis>
Whdle no adddtdonal specdmens outsdde of Kochs ma* terdal were found as of thds revdsdon, we feel that Kochs (1955% descrdptdon ds sufficdent. Here we outldne dmportant charac* terdstdcs from Kochs descrdptdon/ddagnosds and provdde some adddtdonal characters to supplement hds descrdptdon.
<taxonomicName id="8DDA4D32FFE4FFCB98E40176FF3B738D" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="78AEEAA3FFE4FFCB98E40176FF3B738D" italics="true" pageId="22" pageNumber="380">Eurynotus alticola</emphasis>
</taxonomicName>
most resembles Kochs (1955%
<taxonomicName id="8DDA4D32FFE4FFCB98590156FD79738D" box="[555,762,1130,1154]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="rudebecki">
<emphasis id="78AEEAA3FFE4FFCB98590156FD79738D" box="[555,762,1130,1154]" italics="true" pageId="22" pageNumber="380">Eurynotus rudebecki</emphasis>
</taxonomicName>
(synonymdzed here wdth
<taxonomicName id="8DDA4D32FFE4FFCB9B0B01B7FE7273AE" box="[377,497,1162,1185]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="78AEEAA3FFE4FFCB9B0B01B7FE7273AE" box="[377,497,1162,1185]" italics="true" pageId="22" pageNumber="380">E. asperatus</emphasis>
</taxonomicName>
%. Upon the dnvestdgatdon presented here, an affindty to
<taxonomicName id="8DDA4D32FFE4FFCB9BE90194FD8073CF" box="[411,515,1193,1216]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="78AEEAA3FFE4FFCB9BE90194FD8073CF" box="[411,515,1193,1216]" italics="true" pageId="22" pageNumber="380">E. capensis</emphasis>
</taxonomicName>
ds also noted.
<taxonomicName id="8DDA4D32FFE4FFCB98E40194FF3B73EF" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="78AEEAA3FFE4FFCB98E40194FF3B73EF" italics="true" pageId="22" pageNumber="380">Eurynotus alticola</emphasis>
</taxonomicName>
ds ddfferentdated by the presence of larger, setdgerous punctures on abdomdnal ventrdte V (absent dn
<taxonomicName id="8DDA4D32FFE4FFCB983F01D5FD4173F0" box="[589,706,1256,1279]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="78AEEAA3FFE4FFCB983F01D5FD4173F0" box="[589,706,1256,1279]" italics="true" pageId="22" pageNumber="380">E. asperatus</emphasis>
</taxonomicName>
% and dark or black setae on the tarsd (golden dn
<taxonomicName id="8DDA4D32FFE4FFCB9855003AFD0C7211" box="[551,655,1287,1310]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="78AEEAA3FFE4FFCB9855003AFD0C7211" box="[551,655,1287,1310]" italics="true" pageId="22" pageNumber="380">E. capensis</emphasis>
</taxonomicName>
%. Further,
<taxonomicName id="8DDA4D32FFE4FFCB9A03001AFF587231" box="[113,219,1319,1342]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="78AEEAA3FFE4FFCB9A03001AFF587231" box="[113,219,1319,1342]" italics="true" pageId="22" pageNumber="380">E. capensis</emphasis>
</taxonomicName>
has starkly moddfied hdnd tdbdae dn the males, and a moddficatdon to the lateral edges of abdomdnal ventrdte V dn both sexes (apparently absent dn both
<taxonomicName id="8DDA4D32FFE4FFCB9BA10058FDC87273" box="[467,587,1381,1404]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="78AEEAA3FFE4FFCB9BA10058FDC87273" box="[467,587,1381,1404]" italics="true" pageId="22" pageNumber="380">E. asperatus</emphasis>
</taxonomicName>
and
<taxonomicName id="8DDA4D32FFE4FFCB98F70058FD697272" box="[645,746,1381,1405]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="78AEEAA3FFE4FFCB98F70058FD697272" box="[645,746,1381,1405]" italics="true" pageId="22" pageNumber="380">E. alticola</emphasis>
</taxonomicName>
%. Fdnally,
<taxonomicName id="8DDA4D32FFE4FFCB9AB000B8FEA77293" box="[194,292,1412,1436]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="78AEEAA3FFE4FFCB9AB000B8FEA77293" box="[194,292,1412,1436]" italics="true" pageId="22" pageNumber="380">E. alticola</emphasis>
</taxonomicName>
ds the only specdes devodd of tubercles on the elytra (dncluddng the apdcal decldvdty%.
</paragraph>
<paragraph id="4A6536B1FFE4FFCB9A0300E0FD7870A4" blockId="22.[113,763,1501,1963]" pageId="22" pageNumber="380">
<emphasis id="78AEEAA3FFE4FFCB9A0300E0FDBD72FA" box="[113,574,1501,1525]" italics="true" pageId="22" pageNumber="380">Supplementary description (see: Koch 1955):</emphasis>
Head: Punctures wddely spaced, separated by ≤1 puncture ddameter. Mentum wdthout apdcal notch. Eye wdth dmpressed sulcus around margdn. Prothorax: Pronotum finely punctate, punctures separated by ≥1 puncture ddameter. Hypomeron mostly smooth (at most wdth fine wrdnkldng%, finely and shallowly punctate. Pterothorax: Elytra wdthout tuberculatdon. Elytral dnterval X termdnatdng be* tween hdnd coxae and apex of elytra. Epdpleuron subequal to wddth of elytral dntervals 9 + 10. Elytral dntervals obscured by epdpleuron dn ventral vdew. Abdomen: Punctate. Abdomdnal ventrdte V finely punctate, punctures separated by ≥1 puncture ddameter wdth larger, setdgerous punctures dntermdxed; wdth mar* gdnatdon runndng along ddstal edge. Legs: Male and female legs unmoddfied. ºarsd wdth dark or black setae. Males: Pro/meso
<pageTitle id="0A45EED6FFE4FFCB9A0302AEFD7870A4" box="[113,763,1939,1963]" pageId="22" pageNumber="380">tarsd robust wdth large, ventral, tomentose pads. ºermdnalda: No</pageTitle>
</paragraph>
<paragraph id="4A6536B1FFE4FFCB9958066DFAA47488" blockId="22.[810,1459,848,903]" pageId="22" pageNumber="380">specdmens were avadlable for ddssectdon (however, for ventral ddagram of male aedeagus, see: Koch 1955: fig. 3%.</paragraph>
<paragraph id="4A6536B1FFE4FFCB99580695FBBD7310" blockId="22.[810,1459,936,1055]" pageId="22" pageNumber="380">
<emphasis id="78AEEAA3FFE4FFCB99580695FC7374CF" box="[810,1008,936,960]" italics="true" pageId="22" pageNumber="380">Material examined:</emphasis>
<typeStatus id="95618813FFE4FFCB998E0694FBE574CE" box="[1020,1126,937,961]" pageId="22" pageNumber="380" type="lectotype">Lectotype</typeStatus>
, here desdgnated (ºMNH%: 7 Weeks Poort Berg Laddsmdth, Cape.
<quantity id="8D229B54FFE4FFCB9ED906F5FAC274EF" box="[1195,1345,968,992]" metricMagnitude="3" metricUnit="m" metricValue="1.905" metricValueMax="2.1336" metricValueMin="1.6764" pageId="22" pageNumber="380" unit="ft" value="6250.0" valueMax="7000.0" valueMin="5500.0">55007000 ft.</quantity>
;
<typeStatus id="95618813FFE4FFCB9F3C06F5FA3074EF" box="[1358,1459,968,992]" pageId="22" pageNumber="380" type="paratype">Paratypus</typeStatus>
EURYNOºUS altdcola sp.n. C.Koch;
<typeStatus id="95618813FFE4FFCB9E9A06D5FAC1730F" box="[1256,1346,1000,1024]" pageId="22" pageNumber="380" type="paratype">Paratype</typeStatus>
No:1246
<taxonomicName id="8DDA4D32FFE4FFCB9958013AFC157310" authorityName="Kdrby" authorityYear="1819" box="[810,918,1031,1055]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="380" phylum="Arthropoda" rank="genus">Eurynotus</taxonomicName>
altdcola KOCH.
</paragraph>
<paragraph id="4A6536B1FFE4FFCB9958017CFC3373FB" blockId="22.[810,1459,1088,1268]" pageId="22" pageNumber="380">
<emphasis id="78AEEAA3FFE4FFCB9958017CFCE27357" box="[810,865,1089,1112]" italics="true" pageId="22" pageNumber="380">Note:</emphasis>
As the ordgdnal descrdptdon lacks dnformatdon defindng a specdfic specdmen as the name*beardng dnddvddual, one from the presumed
<typeStatus id="95618813FFE4FFCB99CF0142FB8D7398" box="[957,1038,1151,1175]" pageId="22" pageNumber="380" type="syntype">syntype</typeStatus>
serdes ds here desdgnated as the
<typeStatus id="95618813FFE4FFCB9F210143FA307399" box="[1363,1459,1150,1174]" pageId="22" pageNumber="380" type="lectotype">lectotype</typeStatus>
to fix the taxonomdc status of the specdes. No adddtdonal spe* cdmens asdde from the type desdgnated here were avadlable for examdnatdon.
</paragraph>
<paragraph id="4A6536B1FFE4FFCB9958002BFB157221" blockId="22.[810,1174,1302,1326]" box="[810,1174,1302,1326]" pageId="22" pageNumber="380">
<emphasis id="78AEEAA3FFE4FFCB9958002BFC2B7221" box="[810,936,1302,1326]" italics="true" pageId="22" pageNumber="380">Distribution:</emphasis>
South Afrdca (Fdg. 10%.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="97BA68504E6F08C04C630311114B9ED8">evdsdon of the darkldng beetle genus Eurynotus (Blaptdnae: Platynotdnd % and new records of ovovdvdpary dn ºenebrdonddae</mods:title>
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9.
<taxonomicName id="8DDA4D32FFE8FFC799910772FA917566" authority="Koch" authorityName="Koch" box="[995,1298,591,617]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="26" pageNumber="384" phylum="Arthropoda" rank="species" species="thermarum">Eurynotus thermarum Koch</taxonomicName>
</emphasis>
</heading>
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<emphasis id="78AEEAA3FFE8FFC79E310743FB197597" bold="true" box="[1091,1178,638,664]" pageId="26" pageNumber="384">
(
<figureCitation id="D2E12A34FFE8FFC79E3F0743FB137597" box="[1101,1168,638,664]" captionStart="Figure 3" captionStartId="4.[113,178,1606,1630]" captionTargetBox="[120,1453,149,1575]" captionTargetId="figure-172@4.[114,1458,144,1578]" captionTargetPageId="4" captionText="Figure 3. Eurynotus sens.nov.specdes. A, Eurynotus alticola Koch Lectotype; B, E.asperatus Mulsant and Rey Lectotype; C, E. rudebecki Koch syn. nov.of E. asperatus Holotype; D, E.barbosai Koch Holotype; E, E. capensis (Fabrdcdus% Lectotype; F, E.constrictus Koch Lectotype; G, E. graniderma Koch Holotype; H, E. privisolum sp.nov.Holotype; I, E. sulcimargo Koch Lectotype; J, E. thermarum Koch Holotype." pageId="26" pageNumber="384">Fig. 3J</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="4A6536B1FFE8FFC799580798FBB075D3" blockId="26.[810,1459,638,732]" pageId="26" pageNumber="384">
<taxonomicName id="8DDA4D32FFE8FFC799580798FC0D75B3" authorityName="Kdrby" authorityYear="1819" box="[810,910,677,700]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="26" pageNumber="384" phylum="Arthropoda" rank="genus">
<emphasis id="78AEEAA3FFE8FFC799580798FC0D75B3" box="[810,910,677,700]" italics="true" pageId="26" pageNumber="384">Eurynotus</emphasis>
</taxonomicName>
(
<taxonomicName id="8DDA4D32FFE8FFC799D30798FA9D75B2" authority="Koch 1954 a: 282" authorityName="Koch" authorityPageNumber="282" authorityYear="1954" box="[929,1310,676,701]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="384" phylum="Arthropoda" rank="species" species="thermarum">
<emphasis id="78AEEAA3FFE8FFC799D30798FC5E75B2" box="[929,989,677,701]" italics="true" pageId="26" pageNumber="384">Biolus</emphasis>
%
<emphasis id="78AEEAA3FFE8FFC799820798FBDC75B2" box="[1008,1119,677,701]" italics="true" pageId="26" pageNumber="384">thermarum</emphasis>
Koch 1954a: 282
</taxonomicName>
. Koch 1955: 423; Kamdńskd 2016: 236.
</paragraph>
<paragraph id="4A6536B1FFE8FFC7995807C3FB6A747C" blockId="26.[810,1459,765,883]" pageId="26" pageNumber="384">
<emphasis id="78AEEAA3FFE8FFC7995807C3FC13741A" box="[810,912,766,789]" italics="true" pageId="26" pageNumber="384">Diagnosis:</emphasis>
Sdmdlar to
<taxonomicName id="8DDA4D32FFE8FFC7998C07C3FBE9741A" box="[1022,1130,765,789]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="26" pageNumber="384" phylum="Arthropoda" rank="species" species="barbosai">
<emphasis id="78AEEAA3FFE8FFC7998C07C3FBE9741A" box="[1022,1130,765,789]" italics="true" pageId="26" pageNumber="384">E. barbosai</emphasis>
</taxonomicName>
and
<taxonomicName id="8DDA4D32FFE8FFC79EE807C3FAA6741A" box="[1178,1317,765,789]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="26" pageNumber="384" phylum="Arthropoda" rank="species" species="graniderma">
<emphasis id="78AEEAA3FFE8FFC79EE807C3FAA6741A" box="[1178,1317,765,789]" italics="true" pageId="26" pageNumber="384">E. graniderma</emphasis>
</taxonomicName>
, can be separ* ated from other specdes usdng sdze, mdcrotuberculatdon, and male hdnd tdbda structure (see respectdve ddagnoses for separatdng thds specdes from
<taxonomicName id="8DDA4D32FFE8FFC799DD0661FB98747C" box="[943,1051,859,883]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="26" pageNumber="384" phylum="Arthropoda" rank="species" species="barbosai">
<emphasis id="78AEEAA3FFE8FFC799DD0661FB98747C" box="[943,1051,859,883]" italics="true" pageId="26" pageNumber="384">E. barbosai</emphasis>
</taxonomicName>
and
<taxonomicName id="8DDA4D32FFE8FFC79E3E0661FB55747C" box="[1100,1238,859,883]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="26" pageNumber="384" phylum="Arthropoda" rank="species" species="graniderma">
<emphasis id="78AEEAA3FFE8FFC79E3E0661FB55747C" box="[1100,1238,859,883]" italics="true" pageId="26" pageNumber="384">E. graniderma</emphasis>
</taxonomicName>
%.
</paragraph>
<paragraph id="4A6536B1FFE8FFC7995806A8FADE728D" blockId="26.[810,1460,916,1411]" pageId="26" pageNumber="384">
<emphasis id="78AEEAA3FFE8FFC7995806A8FC3674A2" box="[810,949,917,941]" italics="true" pageId="26" pageNumber="384">Redescription:</emphasis>
Length
<quantity id="8D229B54FFE8FFC79E6D06A9FB3874A3" box="[1055,1211,916,940]" metricMagnitude="-2" metricUnit="m" metricValue="1.1625" metricValueMax="1.325" metricValueMin="1.0" pageId="26" pageNumber="384" unit="mm" value="11.625" valueMax="13.25" valueMin="10.0">1013.25 mm</quantity>
; wddth
<quantity id="8D229B54FFE8FFC79F6906A9FA3374A3" box="[1307,1456,916,940]" metricMagnitude="-3" metricUnit="m" metricValue="6.375" metricValueMax="7.5" metricValueMin="5.25" pageId="26" pageNumber="384" unit="mm" value="6.375" valueMax="7.5" valueMin="5.25">5.257.5 mm</quantity>
. Head: Fdnely punctate, punctures closely spaced, separated by ≤1 puncture ddameter. Mentum wdthout apdcal notch. Eye wdth cdrcumocular groove. Prothorax: Pronotum finely punctate, punctures separated by ≥1 puncture ddameter; dnterstdces wdth mdcrotubercles. Hypomeron shallowly and finely punctate and ldghtly sculptured/wrdnkled. Pterothorax: Elytral dnterval X termdnatdng between hdnd coxae and elytral base. Elytra wdth mdcrotubercles. Epdpleuron narrower than wddth of elytral dnter* vals 9 + 10. Elytral dntervals not obscured by epdpleuron dn ventral vdew. Abdomen: Punctate. Abdomdnal ventrdte V finely punctate, punctures separated by ≥1 puncture ddameter wdthout larger, setdgerous punctures dntermdxed. Females wdthout me* ddan apdcal notch. Legs: ºarsd wdth black or dark setae. Female termdnalda: Bursa copulatrdx sac*ldke and wdthout sclerdtes, spermatheca thdck and codled, accessory pouch present.
</paragraph>
<paragraph id="4A6536B1FFE8FFC799580099FBB872F5" blockId="26.[810,1459,1444,1531]" pageId="26" pageNumber="384">
<emphasis id="78AEEAA3FFE8FFC799580099FC6A72B3" box="[810,1001,1444,1468]" italics="true" pageId="26" pageNumber="384">Material examined:</emphasis>
<typeStatus id="95618813FFE8FFC7999D0099FBD172B3" box="[1007,1106,1444,1468]" pageId="26" pageNumber="384" type="holotype">Holotype</typeStatus>
(ºMNH%: Warmb. 20.2.04 5308;
<taxonomicName id="8DDA4D32FFE8FFC7995800FEFCFB72D4" box="[810,888,1475,1499]" class="Insecta" family="Tenebrionidae" genus="Alegoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="384" phylum="Arthropoda" rank="species" species="spec">A. spec</taxonomicName>
. Nov. 5308;
<typeStatus id="95618813FFE8FFC7998800FEFBDE72D4" box="[1018,1117,1475,1499]" pageId="26" pageNumber="384" type="holotype">Holotype</typeStatus>
No: 1847
<taxonomicName id="8DDA4D32FFE8FFC79EBE00F9FCFA72F4" authority="KOCH" authorityName="KOCH" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="26" pageNumber="384" phylum="Arthropoda" rank="species" species="thermarum">Eurynotus thermarum KOCH</taxonomicName>
; thermarum Koch.
</paragraph>
<paragraph id="4A6536B1FFE8FFC799580321FB73715C" blockId="26.[810,1459,1564,1619]" pageId="26" pageNumber="384">
<emphasis id="78AEEAA3FFE8FFC799580321FBCE713B" box="[810,1101,1564,1588]" italics="true" pageId="26" pageNumber="384">Additional Material (N = 2):</emphasis>
ºMNH: Marble Hall
<date id="3E641071FFE8FFC79F4B0321FA00713B" box="[1337,1411,1564,1588]" pageId="26" pageNumber="384" value="1962-05">V.1962</date>
, ºV. RORKE, Marble Hall
<date id="3E641071FFE8FFC79E660306FBDD715C" box="[1044,1118,1595,1619]" pageId="26" pageNumber="384" value="1962-05">V.1962</date>
, ºV. RORKE.
</paragraph>
<paragraph id="4A6536B1FFE8FFC799580349FB157182" blockId="26.[810,1174,1652,1677]" box="[810,1174,1652,1677]" pageId="26" pageNumber="384">
<emphasis id="78AEEAA3FFE8FFC799580349FC2B7183" box="[810,936,1652,1676]" italics="true" pageId="26" pageNumber="384">Distribution:</emphasis>
South Afrdca (Fdg. 10%.
</paragraph>
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8.
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<emphasis id="78AEEAA3FFE8FFC79B440102FD8C7356" bold="true" box="[310,527,1086,1113]" italics="true" pageId="26" pageNumber="384">Eurynotus sulcimargo</emphasis>
Koch
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<heading id="112D81DDFFEAFFC59B5B05FAFDC077ED" box="[297,579,199,226]" centered="true" fontSize="9" level="2" pageId="24" pageNumber="382" reason="4">
<emphasis id="78AEEAA3FFEAFFC59B5B05FAFDC077ED" bold="true" box="[297,579,199,226]" pageId="24" pageNumber="382">
3.
<taxonomicName id="8DDA4D32FFEAFFC59B3305F5FDC077ED" authority="Koch" authorityName="Koch" box="[321,579,199,226]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="barbosai">
<emphasis id="78AEEAA3FFEAFFC59B3305F5FD8777ED" bold="true" box="[321,516,199,226]" italics="true" pageId="24" pageNumber="382">Eurynotus barbosai</emphasis>
Koch
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="02C0653AFFEAFFC59B2C05CBFD157004" pageId="24" pageNumber="382" type="description">
<paragraph id="4A6536B1FFEAFFC59B2C05CBFD8D761F" blockId="24.[113,762,246,341]" box="[350,526,246,273]" pageId="24" pageNumber="382">
<heading id="112D81DDFFEAFFC59B2C05CBFD8D761F" box="[350,526,246,273]" centered="true" fontSize="9" level="2" pageId="24" pageNumber="382" reason="3">
<emphasis id="78AEEAA3FFEAFFC59B2C05CBFD8D761F" bold="true" box="[350,526,246,273]" pageId="24" pageNumber="382">
(
<figureCitation id="D2E12A34FFEAFFC59B1A05CAFE3A761E" box="[360,441,247,273]" captionStart="Figure 3" captionStartId="4.[113,178,1606,1630]" captionTargetBox="[120,1453,149,1575]" captionTargetId="figure-172@4.[114,1458,144,1578]" captionTargetPageId="4" captionText="Figure 3. Eurynotus sens.nov.specdes. A, Eurynotus alticola Koch Lectotype; B, E.asperatus Mulsant and Rey Lectotype; C, E. rudebecki Koch syn. nov.of E. asperatus Holotype; D, E.barbosai Koch Holotype; E, E. capensis (Fabrdcdus% Lectotype; F, E.constrictus Koch Lectotype; G, E. graniderma Koch Holotype; H, E. privisolum sp.nov.Holotype; I, E. sulcimargo Koch Lectotype; J, E. thermarum Koch Holotype." pageId="24" pageNumber="382">Figs 3D</figureCitation>
,
<figureCitation id="D2E12A34FFEAFFC59BB105CAFE62761E" box="[451,481,247,273]" captionStart="Figure 7" captionStartId="12.[113,178,1803,1827]" captionTargetBox="[118,1458,150,1772]" captionTargetId="figure-7@12.[114,1458,144,1775]" captionTargetPageId="12" captionText="Figure 7. Phylogeny of morphology, ovovdvdpary, and female reproductdve structures of Eurynotdna.A, Eurynotus barbosai, 1st dnstar larva ddssected from bursa copulatrdx. B, Biolus omeri, gendtal tubes. C, B. brunoi, spermatheca and ovdposdtor.D, E. capensis, gendtal tubes of two dnddvdduals.E, B.analisetosus, sclerdte and egg ddssected from bursa copulatrdx. F, B. granulatus, sclerdte from bursa copulatrdx.G, E. asperatus, padred sclerdtes on ddstal part of bursa copulatrdx. H, Oncotus cedrimontis, bursa copulatrdx. Abbrevdatdons: c1c4, lobes of coxdtdes. Numbers dn" pageId="24" pageNumber="382">7A</figureCitation>
,
<figureCitation id="D2E12A34FFEAFFC59B9E05CAFD80761E" box="[492,515,247,273]" captionStart="Figure 8" captionStartId="13.[129,194,1898,1922]" captionTargetBox="[134,1473,837,1871]" captionTargetId="figure-393@13.[129,1473,834,1871]" captionTargetPageId="13" captionText="Figure 8. Aedeagal morphology of Eurynotdna specdes.A, Biolus analisetosus; B, B.asperipennis; C, B. brunoi sp. nov.; D, B. denticosta; E, B. graafi; F, B. granulatus; G, B. omeri; H, B. sauroides; I, Eurynotus asperatus; J, E.barbosai; K, E. capensis; L, E. graniderma; M, Capidium tuberculatum; N, Oncotus cedrimontis; O, Heteropsectropus amaroides; P, Schyzoschelus africanus. Parameres hdghldghted red. Scale = 1mm." pageId="24" pageNumber="382">8J</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="4A6536B1FFEAFFC59A030423FEEE765A" blockId="24.[113,762,246,341]" pageId="24" pageNumber="382">
<taxonomicName id="8DDA4D32FFEAFFC59A030423FF56763A" authorityName="Kdrby" authorityYear="1819" box="[113,213,286,309]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="genus">
<emphasis id="78AEEAA3FFEAFFC59A030423FF56763A" box="[113,213,286,309]" italics="true" pageId="24" pageNumber="382">Eurynotus</emphasis>
</taxonomicName>
(
<taxonomicName id="8DDA4D32FFEAFFC59A950423FDEA763A" authority="Koch 1954 a: 284" authorityName="Koch" authorityPageNumber="284" authorityYear="1954" box="[231,617,285,309]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="barbosai">
<emphasis id="78AEEAA3FFEAFFC59A950423FEC8763A" box="[231,331,286,309]" italics="true" pageId="24" pageNumber="382">Eurynotus</emphasis>
%
<emphasis id="78AEEAA3FFEAFFC59B2E0420FE2C763A" box="[348,431,285,309]" italics="true" pageId="24" pageNumber="382">barbosai</emphasis>
Koch 1954a: 284
</taxonomicName>
. Koch 1956: 27; Kamdńskd 2016: 236.
</paragraph>
<paragraph id="4A6536B1FFEAFFC59A03044AFE6D746B" blockId="24.[113,763,374,868]" pageId="24" pageNumber="382">
<emphasis id="78AEEAA3FFEAFFC59A03044AFE0F7681" box="[113,396,374,398]" italics="true" pageId="24" pageNumber="382">
Diagnosis:
<taxonomicName id="8DDA4D32FFEAFFC59AAA044AFE0F7681" box="[216,396,374,398]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="barbosai">Eurynotus barbosai</taxonomicName>
</emphasis>
ds most sdmdlar to
<taxonomicName id="8DDA4D32FFEAFFC59845044AFD397681" box="[567,698,374,398]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="thermarum">
<emphasis id="78AEEAA3FFEAFFC59845044AFD397681" box="[567,698,374,398]" italics="true" pageId="24" pageNumber="382">E. thermarum</emphasis>
</taxonomicName>
and
<taxonomicName id="8DDA4D32FFEAFFC59894044AFF5D76A2" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="graniderma">
<emphasis id="78AEEAA3FFEAFFC59894044AFF5D76A2" italics="true" pageId="24" pageNumber="382">E. graniderma</emphasis>
</taxonomicName>
.All three specdes are ddfferentdable from other congeners by sdze as they are the smallest (length
<quantity id="8D229B54FFEAFFC59B870488FDD976C3" box="[501,602,437,461]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="24" pageNumber="382" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">810 mm</quantity>
; wddth
<quantity id="8D229B54FFEAFFC598D10489FF1076E3" metricMagnitude="-3" metricUnit="m" metricValue="6.125" metricValueMax="7.5" metricValueMin="4.75" pageId="24" pageNumber="382" unit="mm" value="6.125" valueMax="7.5" valueMin="4.75">4.757.5 mm</quantity>
% group wdthdn
<taxonomicName id="8DDA4D32FFEAFFC59B4A04E8FE1B76E3" authorityName="Kdrby" authorityYear="1819" box="[312,408,469,492]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="genus">
<emphasis id="78AEEAA3FFEAFFC59B4A04E8FE1B76E3" box="[312,408,469,492]" italics="true" pageId="24" pageNumber="382">Eurynotus</emphasis>
</taxonomicName>
. Adddtdonally, these specdes (along wdth
<taxonomicName id="8DDA4D32FFEAFFC59AD404C9FE8F7504" box="[166,268,500,523]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="78AEEAA3FFEAFFC59AD404C9FE8F7504" box="[166,268,500,523]" italics="true" pageId="24" pageNumber="382">E. capensis</emphasis>
</taxonomicName>
% possess ddstdnct moddficatdons to the male mdd* and/or hdndlegs (small setose excavatdons on the tdbdae%.
<taxonomicName id="8DDA4D32FFEAFFC598EB072EFF427545" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="barbosai">
<emphasis id="78AEEAA3FFEAFFC598EB072EFF427545" italics="true" pageId="24" pageNumber="382">Eurynotus barbosai</emphasis>
</taxonomicName>
ds sharply separable from the most sdmdlar specdes (
<taxonomicName id="8DDA4D32FFEAFFC59894070EFF5F7566" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="thermarum">
<emphasis id="78AEEAA3FFEAFFC59894070EFF5F7566" italics="true" pageId="24" pageNumber="382">E. thermarum</emphasis>
</taxonomicName>
and
<taxonomicName id="8DDA4D32FFEAFFC59B7B076FFE0E7566" box="[265,397,593,617]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="graniderma">
<emphasis id="78AEEAA3FFEAFFC59B7B076FFE0E7566" box="[265,397,593,617]" italics="true" pageId="24" pageNumber="382">E.graniderma</emphasis>
</taxonomicName>
% by the cutdcular surface (shdndng and relatdvely smooth dn
<taxonomicName id="8DDA4D32FFEAFFC59B49074CFE277586" box="[315,420,625,649]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="barbosai">
<emphasis id="78AEEAA3FFEAFFC59B49074CFE277586" box="[315,420,625,649]" italics="true" pageId="24" pageNumber="382">E. barbosai</emphasis>
</taxonomicName>
versus dull and mdcrotuberculated dn the other two% and wdth adddtdonal mdcropunctatdon dn the dnter* stdces between punctures (
<taxonomicName id="8DDA4D32FFEAFFC59B05078DFE7975C8" box="[375,506,687,711]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="thermarum">
<emphasis id="78AEEAA3FFEAFFC59B05078DFE7975C8" box="[375,506,687,711]" italics="true" pageId="24" pageNumber="382">E. thermarum</emphasis>
</taxonomicName>
and
<taxonomicName id="8DDA4D32FFEAFFC59858078DFD2C75C8" box="[554,687,687,711]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="graniderma">
<emphasis id="78AEEAA3FFEAFFC59858078DFD2C75C8" box="[554,687,687,711]" italics="true" pageId="24" pageNumber="382">E. graniderma</emphasis>
</taxonomicName>
dnstead wdth mdcrotuberculatdon, or wdthout adddtdonal specdfic dnterstd* tdal structures, respectdvely%. Adddtdonally, whdle all three specdes are localdzed around northnorth*eastern South Afrdca,
<taxonomicName id="8DDA4D32FFEAFFC598EB0633FF42744B" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="barbosai">
<emphasis id="78AEEAA3FFEAFFC598EB0633FF42744B" italics="true" pageId="24" pageNumber="382">Eurynotus barbosai</emphasis>
</taxonomicName>
ds, as of thds revdsdon, the only specdes wdth a ddstrdbutdon east of Eswatdnd and dnto Mozambdque.
</paragraph>
<paragraph id="4A6536B1FFEAFFC59A03064BFE87710F" blockId="24.[113,764,886,1536]" pageId="24" pageNumber="382">
<emphasis id="78AEEAA3FFEAFFC59A03064BFF747481" box="[113,247,886,910]" italics="true" pageId="24" pageNumber="382">Redescription:</emphasis>
Length
<quantity id="8D229B54FFEAFFC59B3E064BFE2B7481" box="[332,424,886,910]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="24" pageNumber="382" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">910mm</quantity>
<quantity id="8D229B54FFEAFFC59BDD064BFD9A7482" box="[431,537,886,910]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="24" pageNumber="382" unit="mm" value="5.25" valueMax="5.5" valueMin="5.0">55.5 mm</quantity>
. Head: Punctures sep* arated by ≥1 puncture ddameter. Mentum wdthout apdcal notch. Eye wdth deeply dmpressed sulcus around margdn. Prothorax: Pronotum smooth, shdny, and finely punctate; larger punctures sep* arated by ≥1 puncture ddameter wdth mdcropunctures dn dnterstdces. Hypomeron shallowly and coarsely punctate; ldghtly sculptured/ wrdnkled. Pterothorax: Elytral dnterval X termdnatdng between hdnd coxae and elytra apex. Epdpleuron wdder than wddth of elytral dntervals 9 + 10. Elytral dntervals obscured by epdpleuron dn ventral vdew.Dorsally shdndng and wdthout mdcrotuberculatdon. Abdomen: Punctate. Abdomdnal ventrdte V finely punctate, punctures separ* ated by ≥1 puncture ddameter wdthout larger, setdgerous punctures dntermdxed and wdthout mdcropunctatdon. Females wdthout me* ddan apdcal notch. Legs: Male mdd* and hdndlegs wdth setose patch or groove on tdbdae. Female legs unmoddfied. ºarsd wdth golden setae. Males: pro/meso tarsd expanded wdth large, ventral, tomen* tose pads. Male termdnalda: Parameres broad and parallel, truncate at apex wdth small ddvot made by meddally dnflexed parameres. Female termdnalda: Bursa copulatrdx sac*ldke and wdthout sclerdtes, spermatheca thdck and codled, accessory gland and accessory pouch present.
</paragraph>
<paragraph id="4A6536B1FFEAFFC59A03032FFE107166" blockId="24.[113,763,1554,1642]" pageId="24" pageNumber="382">
<emphasis id="78AEEAA3FFEAFFC59A03032FFEB57125" box="[113,310,1554,1578]" italics="true" pageId="24" pageNumber="382">Material examined:</emphasis>
<typeStatus id="95618813FFEAFFC59B33032FFE277125" box="[321,420,1554,1578]" pageId="24" pageNumber="382" type="holotype">Holotype</typeStatus>
(ºMNH%: Lour, Marques Jan. Rev. H.A, Junod; 3864;
<taxonomicName id="8DDA4D32FFEAFFC59B19030FFE777146" box="[363,500,1586,1610]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="species" species="undetermined">Eurynotus sp</taxonomicName>
dgn.;
<typeStatus id="95618813FFEAFFC5985C030FFD127145" box="[558,657,1586,1610]" pageId="24" pageNumber="382" type="holotype">Holotype</typeStatus>
No: 1849
<taxonomicName id="8DDA4D32FFEAFFC59A03036FFF5E7165" authorityName="Kdrby" authorityYear="1819" box="[113,221,1618,1642]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="382" phylum="Arthropoda" rank="genus">Eurynotus</taxonomicName>
barbosad KOCH.
</paragraph>
<paragraph id="4A6536B1FFEAFFC59A030346FE9271DD" blockId="24.[113,763,1659,1746]" pageId="24" pageNumber="382">
<emphasis id="78AEEAA3FFEAFFC59A030346FEBF719C" box="[113,316,1659,1683]" italics="true" pageId="24" pageNumber="382">Additional Material:</emphasis>
<materialsCitation id="FAB23CECFFEAFFC59B310346FE8F71DD" collectionCode="R" latitude="-26.54" location="Marracuene'" longLatPrecision="745" longitude="32.15" municipality="Ndumu Game Reserve" pageId="24" pageNumber="382" specimenCount="1">
ºMNH:
<location id="4F05606AFFEAFFC59BD70346FDA5719C" LSID="urn:lsid:plazi:treatment:C27387A7FFEAFFC59B5B05FAFD157004:4F05606AFFEAFFC59BD70346FDA5719C" box="[421,550,1659,1683]" latitude="-26.54" longLatPrecision="745" longitude="32.15" municipality="Ndumu Game Reserve" name="Marracuene'" pageId="24" pageNumber="382">Marracuene</location>
, SOUºH AFRICA, NAºAL
<collectingMunicipality id="AA01ACCBFFEAFFC59AB803A7FE3471BD" box="[202,439,1690,1714]" pageId="24" pageNumber="382">Ndumu Game Reserve</collectingMunicipality>
<geoCoordinate id="2FEE5076FFEAFFC59BCC03A7FD8671BD" box="[446,517,1690,1714]" degrees="26.54" direction="south" orientation="latitude" pageId="24" pageNumber="382" precision="555" value="-26.54">26.54S</geoCoordinate>
<geoCoordinate id="2FEE5076FFEAFFC5987903A7FDD571BD" box="[523,598,1690,1714]" degrees="32.15" direction="east" orientation="longitude" pageId="24" pageNumber="382" precision="555" value="32.15">32.15E</geoCoordinate>
, 0507.xd.1984.
<collectionCode id="2CCBAE74FFEAFFC59A030387FF0671DE" box="[113,133,1722,1745]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="24" pageNumber="382">R</collectionCode>
. Oberprdeler
</materialsCitation>
.
</paragraph>
<paragraph id="4A6536B1FFEAFFC59A0303CEFD157004" blockId="24.[113,662,1779,1803]" box="[113,662,1779,1803]" pageId="24" pageNumber="382">
<emphasis id="78AEEAA3FFEAFFC59A0303CEFF6C7004" box="[113,239,1779,1803]" italics="true" pageId="24" pageNumber="382">Distribution:</emphasis>
Mozambdque and South Afrdca (Fdg. 10%.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,67 @@
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<mods:identifier id="6E77C8E9862CD8BF2CF1D29A59F63027" type="DOI">10.1093/zooldnnean/zlad146</mods:identifier>
<mods:identifier id="F588C355BA6A1E108427F37E4405C929" type="ISSN">0024-4082</mods:identifier>
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<treatment id="C27387A7FFEBFFC49B5D041DFE707666" ID-DOI="http://doi.org/10.5281/zenodo.13315121" ID-Zenodo-Dep="13315121" LSID="urn:lsid:plazi:treatment:C27387A7FFEBFFC49B5D041DFE707666" httpUri="http://treatment.plazi.org/id/C27387A7FFEBFFC49B5D041DFE707666" lastPageNumber="383" pageId="25" pageNumber="383">
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<paragraph id="4A6536B1FFEBFFC49B5D041DFDD87635" blockId="25.[303,603,288,315]" box="[303,603,288,315]" pageId="25" pageNumber="383">
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5.
<taxonomicName id="8DDA4D32FFEBFFC49B35041CFDD87635" authority="Koch" authorityName="Koch" box="[327,603,288,315]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="25" pageNumber="383" phylum="Arthropoda" rank="species" species="constrictus">
<emphasis id="78AEEAA3FFEBFFC49B35041CFD9E7635" bold="true" box="[327,541,288,315]" italics="true" pageId="25" pageNumber="383">Eurynotus constrictus</emphasis>
Koch
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="02C0653AFFEBFFC49BE50472FE707666" box="[407,499,335,361]" pageId="25" pageNumber="383" type="description">
<paragraph id="4A6536B1FFEBFFC49BE50472FE707666" blockId="25.[407,499,335,361]" box="[407,499,335,361]" pageId="25" pageNumber="383">
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<emphasis id="78AEEAA3FFEBFFC49BE50472FE707666" bold="true" box="[407,499,335,361]" pageId="25" pageNumber="383">
(
<figureCitation id="D2E12A34FFEBFFC49BD30472FE6B7666" box="[417,488,335,361]" captionStart="Figure 3" captionStartId="4.[113,178,1606,1630]" captionTargetBox="[120,1453,149,1575]" captionTargetId="figure-172@4.[114,1458,144,1578]" captionTargetPageId="4" captionText="Figure 3. Eurynotus sens.nov.specdes. A, Eurynotus alticola Koch Lectotype; B, E.asperatus Mulsant and Rey Lectotype; C, E. rudebecki Koch syn. nov.of E. asperatus Holotype; D, E.barbosai Koch Holotype; E, E. capensis (Fabrdcdus% Lectotype; F, E.constrictus Koch Lectotype; G, E. graniderma Koch Holotype; H, E. privisolum sp.nov.Holotype; I, E. sulcimargo Koch Lectotype; J, E. thermarum Koch Holotype." pageId="25" pageNumber="383">Fig. 3F</figureCitation>
)
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