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<document ENCODING="UTF-8" ID-DOI="https://doi.org/10.5962/bhl.title.542" ID-GBIF-Dataset="57ea6fdb-2533-44e5-b60b-736758a88d19" ID-Zenodo-Dep="3922206" ID-ZooBank="2C6327E1-5560-4DB4-B9CA-76A0FA03D975" checkinTime="1593218702860" checkinUser="admin" docAuthor="Linnaeus, Carolus" docDate="1758" docId="FE5B087A19DAB4171E26506A1F3600D0" docLanguage="de" docName="Linnaeus1758Treatments.xml" docOrigin="Stockholm: Laurentius Salvius" docSource="https://archive.org/download/mobot31753000798865/mobot31753000798865.pdf" docTitle="Tubipora Linnaeus, 1758, gen. nov." docType="treatment" docUuid="EC2CF37B-B4C0-4879-8876-F0BBA00483BF" docUuidSource="ZooBank" docVersion="13" lastPageNumber="789" masterDocId="3F896AFC21C459639D0D71D9EDB8875C" masterDocTitle="Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis" pageNumber="789" updateTime="1643600433100" updateUser="ExternalLinkService">
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<mods:title>Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Linnaeus, Carolus</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:placeTerm>Stockholm</mods:placeTerm>
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<mods:location>
<mods:url>https://archive.org/download/mobot31753000798865/mobot31753000798865.pdf</mods:url>
</mods:location>
<mods:classification>book</mods:classification>
<mods:identifier type="ZooBank">2C6327E1-5560-4DB4-B9CA-76A0FA03D975</mods:identifier>
<mods:identifier type="DOI">https://doi.org/10.5962/bhl.title.542</mods:identifier>
<mods:identifier type="Zenodo-Dep">3922206</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3915602" ID-GBIF-Taxon="164807630" ID-Zenodo-Dep="3915602" LSID="urn:lsid:zoobank.org:act:EC2CF37B-B4C0-4879-8876-F0BBA00483BF" httpUri="http://treatment.plazi.org/id/FE5B087A19DAB4171E26506A1F3600D0" lastPageNumber="789" pageNumber="789" sourcePageUrl="https://www.biodiversitylibrary.org/page/727700">
<subSubSection type="nomenclature">
<paragraph pageNumber="789">
<taxonomicName LSID="urn:lsid:zoobank.org:act:EC2CF37B-B4C0-4879-8876-F0BBA00483BF" authorityName="Linnaeus" authorityYear="1758" class="Anthozoa" family="Tubiporidae" genus="Tubipora" kingdom="Animalia" order="Alcyonacea" phylum="Cnidaria" rank="genus" status="gen. nov." zbkClass="Vermes" zbkKingdom="Animalia" zbkOrder="Lithophyta">Tubipora</taxonomicName>
[
<taxonomicNameLabel rank="genus">gen. nov.</taxonomicNameLabel>
]
</paragraph>
</subSubSection>
<subSubSection type="description">
<paragraph pageNumber="789">
<emphasis italics="true">Animalia</emphasis>
Nereides.
</paragraph>
<paragraph pageNumber="789">
<emphasis italics="true">Corallium</emphasis>
tubis subcylindricis, laevibus, basin usque cavis.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.384.6197" ID-GBIF-Dataset="f8902ab2-4591-440e-93f1-fb2d2f850112" ID-PMC="PMC3950428" ID-Pensoft-Pub="1313-2970-384-1" ID-PubMed="24624022" ID-ZBK="832EF8274642416895252C2AD202EB9B" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-384-1" ModsDocOrigin="ZooKeys 384" ModsDocTitle="A revision of the genus Arenivaga (Rehn) (Blattodea, Corydiidae), with descriptions of new species and key to the males of the genus" checkinTime="1451246226561" checkinUser="pensoft" docAuthor="Hopkins, Heidi" docDate="2014" docId="FE5B22D9FD7C11B327F457D381252514" docLanguage="en" docName="ZooKeys 384: 1-256" docOrigin="ZooKeys 384" docSource="http://dx.doi.org/10.3897/zookeys.384.6197" docTitle="Arenivaga mortisvallisensis Hopkins, 2014, sp. n." docType="treatment" docUuid="3086A1CB-8F24-43DE-89B7-E9EF84621413" docUuidSource="ZooBank" docVersion="4" lastPageNumber="85" masterDocId="FF86641EFFB8B30DFFC9F429FFD4FFA1" masterDocTitle="A revision of the genus Arenivaga (Rehn) (Blattodea, Corydiidae), with descriptions of new species and key to the males of the genus" masterLastPageNumber="256" masterPageNumber="1" pageNumber="84" updateTime="1668157791821" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A revision of the genus Arenivaga (Rehn) (Blattodea, Corydiidae), with descriptions of new species and key to the males of the genus</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hopkins, Heidi</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>384</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>256</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.384.6197</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.384.6197</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-384-1</mods:identifier>
<mods:identifier type="ZBK">832EF8274642416895252C2AD202EB9B</mods:identifier>
<mods:identifier type="ZooBank">832EF8274642416895252C2AD202EB9B</mods:identifier>
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<treatment ID-GBIF-Taxon="152051378" LSID="urn:lsid:zoobank.org:act:3086A1CB-8F24-43DE-89B7-E9EF84621413" httpUri="http://treatment.plazi.org/id/FE5B22D9FD7C11B327F457D381252514" lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84">
<subSubSection pageId="83" pageNumber="84" type="nomenclature">
<paragraph pageId="83" pageNumber="84">
<taxonomicName LSID="http://zoobank.org/3086A1CB-8F24-43DE-89B7-E9EF84621413" class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga mortisvallisensis" order="Blattodea" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="mortisvallisensis">
<pageBreakToken pageId="83" pageNumber="84" start="start">Arenivaga</pageBreakToken>
mortisvallisensis
</taxonomicName>
<taxonomicNameLabel pageId="83" pageNumber="84">sp. n.</taxonomicNameLabel>
Figures 108-110
</paragraph>
</subSubSection>
<subSubSection pageId="83" pageNumber="84" type="type locality">
<paragraph pageId="83" pageNumber="84">Type locality.</paragraph>
<paragraph pageId="83" pageNumber="84">USA, California, Inyo Co., Death Valley, Sand Spring</paragraph>
</subSubSection>
<subSubSection pageId="83" pageNumber="84" type="material examined">
<paragraph pageId="83" pageNumber="84">Material examined.</paragraph>
<paragraph pageId="83" pageNumber="84">
Holotype: ♂ in CSCA labeled &quot;CALIF: Inyo Co., Sand Spring 15.5 mi NW Scottys Castle, 3100', II-87 to IX-1987, Antifreeze pit trap&quot; &quot;HOLOTYPE
<taxonomicName class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga mortisvallisensis" order="Blattodea" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="mortisvallisensis">Arenivaga mortisvallisensis</taxonomicName>
Hopkins, 2012&quot; [red label with black border].
</paragraph>
<paragraph pageId="83" pageNumber="84">Paratypes: None at this time.</paragraph>
</subSubSection>
<subSubSection pageId="83" pageNumber="84" type="etymology">
<paragraph pageId="83" pageNumber="84">Etymology.</paragraph>
<paragraph pageId="83" pageNumber="84">This species is named for its locality, Death Valley, CA.</paragraph>
</subSubSection>
<subSubSection pageId="83" pageNumber="84" type="distribution">
<paragraph pageId="83" pageNumber="84">Distribution.</paragraph>
<paragraph pageId="83" pageNumber="84">This species is only known from one specimen from Death Valley NM, CA. See Fig. 110.</paragraph>
</subSubSection>
<subSubSection pageId="83" pageNumber="84" type="diagnosis">
<paragraph pageId="83" pageNumber="84">Diagnosis.</paragraph>
<paragraph pageId="83" pageNumber="84">
<taxonomicName class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga mortisvallisensis" order="Blattodea" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="mortisvallisensis">Arenivaga mortisvallisensis</taxonomicName>
is smaller than average for
<taxonomicName class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga" order="Blattodea" pageId="83" pageNumber="84" phylum="Arthropoda" rank="genus">Arenivaga</taxonomicName>
but may be confused with
<taxonomicName class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga delicata" order="Blattodea" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="delicata">Arenivaga delicata</taxonomicName>
which has sympatric distribution.
<taxonomicName class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga mortisvallisensis" order="Blattodea" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="mortisvallisensis">Arenivaga mortisvallisensis</taxonomicName>
may be distinguished by the medial margin of the right dorsal phallomere that projects anteriorly in pronounced manner into rounded, shagreened lobe with toothed margin. See Figs 109 and 40.
</paragraph>
</subSubSection>
<subSubSection lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84" type="description">
<paragraph pageId="83" pageNumber="84">Description.</paragraph>
<paragraph pageId="83" pageNumber="84">Male.Measurements. Approximate due to poor condition of specimen. Holotype TL = 15.3 mm, GW = 6.2 mm, PW = 4.3 mm, PL = 3.24 mm, TL/GW = 2.47, PL/PW = 0.75. EW = 0.60 mm; OW = 0.40 mm.</paragraph>
<paragraph pageId="83" pageNumber="84">
Head. Two ocelli large, ovoid and less protruding than on most species (0.3
<normalizedToken originalValue="×">x</normalizedToken>
0.2 mm); vertex unusually broad, dark brown with small ridges in rays around upper apices of eyes and extending onto ocellar tubercles; interocellar space concave, smooth, dark brown; two teardrop shaped indentations medial to ocelli. Frons very dark brown, posterior half flat, tectiform just below indentations; anterior portion of frons bulbous, very dark brown; very dark brown smooth anteclypeus. See Fig. 108d.
</paragraph>
<paragraph pageId="83" pageNumber="84">Pronotum. Pronotum translucent, waxy beige; dorsal surface of pronotum with short fine golden setae centrally and posteriorly grading to longer, thicker setae laterally and anteriorly; pronotal pattern dark brown &quot;panther face&quot;, impressed, with little discernible detail; no aura. See Fig. 108c.</paragraph>
<paragraph pageId="83" pageNumber="84">Body. Wing brace present. Two tarsal claws present. Legs and body orange-brown; abdomen missing from specimen; species, or perhaps just this specimen, unique in its almost complete absence of deposits of uric acid, and those only very minimally in forewings. Subgenital plate cleared in KOH with genitalia therefore color uncertain; strongly asymmetrical with rounded apices. See Fig. 108b.</paragraph>
<paragraph pageId="83" pageNumber="84">Forewings. Wings extended beyond abdominal apex though distance cannot be estimated as abdomen is missing; transparent pale brown with veins darker brown; surface hyaline. See Fig. 108a.</paragraph>
<paragraph pageId="84" pageNumber="85">
<pageBreakToken pageId="84" pageNumber="85" start="start">Genitalia</pageBreakToken>
. Right dorsal phallomere composed of bulbous lightly sclerotized lobe, articulated with right ventral phallomere on lateral side; central field lightly sclerotized; medial margin strongly projected anteriorly into rounded, shagreened lobe with toothed margin. Small central sclerite of nondescript shape, finely punctate, mostly flat with ventrally bent rim slightly more sclerotize and densely punctate; right ventral phallomere extends from articulation to form rounded somewhat elongate smooth structure that becomes shorter and punctate anteriorly; attached anteriorly is mildly dorsally projecting flanged arm, shagreened with lightly toothed edge. Folded anterior portion of left phallomere setose, otherwise unmodified. Genital hook with short distance to pointed head with short hook; arm very delicate. See Fig. 109.
</paragraph>
<caption pageId="84" pageNumber="85">
<paragraph pageId="84" pageNumber="85">
Figure 108.
<taxonomicName class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga mortisvallisensis" order="Blattodea" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="mortisvallisensis">Arenivaga mortisvallisensis</taxonomicName>
a dorsal habitus b ventral habitus c pronotum d head.
</paragraph>
</caption>
<caption pageId="84" pageNumber="85">
<paragraph pageId="84" pageNumber="85">
Figure 109.
<taxonomicName class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga mortisvallisensis" order="Blattodea" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="mortisvallisensis">Arenivaga mortisvallisensis</taxonomicName>
, genitalia: a) right dorsal phallomere b right ventral phallomere c small central sclerite d genital hook. Arrow(s) indicate diagnostic characters (see text).
</paragraph>
</caption>
<caption pageId="84" pageNumber="85">
<paragraph pageId="84" pageNumber="85">
Figure 110.
<taxonomicName class="Insecta" family="Corydiidae" genus="Arenivaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arenivaga mortisvallisensis" order="Blattodea" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="mortisvallisensis">Arenivaga mortisvallisensis</taxonomicName>
, distribution.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="84" pageNumber="85" type="habitat and natural history">
<paragraph pageId="84" pageNumber="85">Habitat and natural history.</paragraph>
<paragraph pageId="84" pageNumber="85">All life history elements remain unobserved.</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="5852BB5BEB029BABB74A8544AA2883E7" ID-DOI="10.5281/zenodo.169958" ID-GBIF-Dataset="c3f67774-ce96-432d-a2ab-9c176d7e3219" ID-ISSN="1175­5326" ID-Zenodo-Dep="169958" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459749106037" checkinUser="plazi" docAuthor="Pitts, James P." docDate="2005" docId="FE5B5F28FFF0B722FEEDF6C0FB8DFC4B" docLanguage="en" docName="zt01050p043.pdf" docOrigin="Zootaxa 1050" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Sphaeropthalma nocticaro Pitts 2005, New Species" docType="treatment" docVersion="11" lastPageNumber="43" masterDocId="02622750FFF1B726FFE5F241FF80FFC5" masterDocTitle="Description of a new species in the Sphaeropthalma pensylvanica species­group (Hymenoptera: Mutillidae)" masterLastPageNumber="43" masterPageNumber="39" pageNumber="40" updateTime="1698211302435" updateUser="plazi">
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<mods:title id="A4BAEBB9D281518032A4F24FA5601302">Description of a new species in the Sphaeropthalma pensylvanica species­group (Hymenoptera: Mutillidae)</mods:title>
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<mods:namePart id="50CB48F80B3F664DDADD992472C38404">Pitts, James P.</mods:namePart>
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<mods:date id="8A80DAA58E46EE1D9A7C2B02B215165E">2005</mods:date>
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<taxonomicName id="B1F295BDFFF0B727FEEDF6C0FD0CFB5E" authority="Pitts" authorityName="Pitts" authorityYear="2005" box="[264,652,1153,1179]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="nocticaro" status="sp. nov.">
<emphasis id="4486322CFFF0B727FEEDF6C0FDCAFB5E" bold="true" box="[264,586,1153,1179]" italics="true" pageId="1" pageNumber="40">Sphaeropthalma nocticaro</emphasis>
Pitts
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,
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(
<figureCitation id="EEC9F2BBFFF0B727FCA7F6C0FC34FB5E" box="[834,948,1153,1179]" captionStart-0="FIGURE 1 – 3" captionStart-1="FIGURE 4 – 5" captionStartId-0="2.[264,367,1260,1284]" captionStartId-1="3.[264,367,954,978]" captionTargetBox-0="[264,1322,473,1246]" captionTargetBox-1="[537,1080,443,917]" captionTargetId-0="figure@2.[264,1323,472,1248]" captionTargetId-1="figure@3.[486,1101,441,928]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." captionText-1="FIGURE 4 – 5. Holotype of Sphaeropthalma nocticaro, sp. nov., genitalia. 4. Dorsal view, right, ventral view, left; 5. Lateral view." httpUri-0="https://zenodo.org/record/169959/files/figure.png" httpUri-1="https://zenodo.org/record/169960/files/figure.png" pageId="1" pageNumber="40">Figs. 1–5</figureCitation>
)
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<subSubSection id="3EE8BDB5FFF0B727FEEDF68DFB3EFAF3" pageId="1" pageNumber="40" type="diagnosis">
<paragraph id="764DEE3EFFF0B727FEEDF68DFB3EFAF3" blockId="1.[264,1323,1228,1894]" pageId="1" pageNumber="40">
<emphasis id="4486322CFFF0B727FEEDF68DFE07FB20" box="[264,391,1228,1253]" italics="true" pageId="1" pageNumber="40">Diagnosis.</emphasis>
This species is easily differentiated from other members of the
<taxonomicName id="B1F295BDFFF0B727FB91F68DFAAAFB20" box="[1140,1322,1228,1253]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="pensylvanica">
<emphasis id="4486322CFFF0B727FB91F68DFAAAFB20" box="[1140,1322,1228,1253]" italics="true" pageId="1" pageNumber="40">S. pensylvanica</emphasis>
</taxonomicName>
species­group by the piceous integumental coloration (
<figureCitation id="EEC9F2BBFFF0B727FC7CF6B5FC60FACB" box="[921,992,1268,1294]" captionStart="FIGURE 1 – 3" captionStartId="2.[264,367,1260,1284]" captionTargetBox="[264,1322,473,1246]" captionTargetId="figure@2.[264,1323,472,1248]" captionTargetPageId="2" captionText="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="1" pageNumber="40">Fig. 1</figureCitation>
), the white brachyplumose setae, and T3 being completely covered in distinct, white, plumose setae (
<figureCitation id="EEC9F2BBFFF0B727FB8DF75DFB2DFAF3" box="[1128,1197,1308,1334]" captionStart="FIGURE 1 – 3" captionStartId="2.[264,367,1260,1284]" captionTargetBox="[264,1322,473,1246]" captionTargetId="figure@2.[264,1323,472,1248]" captionTargetPageId="2" captionText="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="1" pageNumber="40">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3EE8BDB5FFF0B725FEDDF705FD99FA12" lastPageId="3" lastPageNumber="42" pageId="1" pageNumber="40" type="description">
<paragraph id="764DEE3EFFF0B727FEDDF705FB6DF98B" blockId="1.[264,1323,1228,1894]" pageId="1" pageNumber="40">
<emphasis id="4486322CFFF0B727FEDDF705FBA9FA98" box="[312,1065,1348,1373]" italics="true" pageId="1" pageNumber="40">
Description of Male
<typeStatus id="A949509CFFF0B727FDD9F705FD2BFA98" box="[572,683,1348,1373]" pageId="1" pageNumber="40" type="holotype">Holotype</typeStatus>
. Coloration and Setal Pattern.
</emphasis>
Integument piceous, flagellum black (
<figureCitation id="EEC9F2BBFFF0B727FE35F72DFD96FA43" box="[464,534,1388,1414]" captionStart="FIGURE 1 – 3" captionStartId="2.[264,367,1260,1284]" captionTargetBox="[264,1322,473,1246]" captionTargetId="figure@2.[264,1323,472,1248]" captionTargetPageId="2" captionText="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="1" pageNumber="40">Fig. 1</figureCitation>
). Body covered in sparse erect white brachyplumose setae, except front also with indistinct piceous brachyplumose setae present, and mesonotum, tibiae, tarsi, disk of T2, T4–T7 and S6–S7 also with piceous brachyplumose setae (
<figureCitation id="EEC9F2BBFFF0B727FB65F7FDFB45FA13" box="[1152,1221,1468,1494]" captionStart="FIGURE 1 – 3" captionStartId="2.[264,367,1260,1284]" captionTargetBox="[264,1322,473,1246]" captionTargetId="figure@2.[264,1323,472,1248]" captionTargetPageId="2" captionText="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="1" pageNumber="40">Fig. 1</figureCitation>
). Fringe of T1–T3 with dense distinct white plumose setae (
<figureCitation id="EEC9F2BBFFF0B727FC62F7A5FC51FA3B" box="[903,977,1508,1534]" captionStart="FIGURE 1 – 3" captionStartId="2.[264,367,1260,1284]" captionTargetBox="[264,1322,473,1246]" captionTargetId="figure@2.[264,1323,472,1248]" captionTargetPageId="2" captionText="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="1" pageNumber="40">Fig. 1</figureCitation>
). T3 disk covered in dense plumose white setae obscuring sculpture; some plumose setae yellow tinged (
<figureCitation id="EEC9F2BBFFF0B727FB28F44DFA97F9E3" box="[1229,1303,1548,1574]" captionStart="FIGURE 1 – 3" captionStartId="2.[264,367,1260,1284]" captionTargetBox="[264,1322,473,1246]" captionTargetId="figure@2.[264,1323,472,1248]" captionTargetPageId="2" captionText="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="1" pageNumber="40">Fig. 1</figureCitation>
). Wings infuscate with simple brown setae (
<figureCitation id="EEC9F2BBFFF0B727FD1DF475FCBDF98B" box="[760,829,1588,1614]" captionStart="FIGURE 1 – 3" captionStartId="2.[264,367,1260,1284]" captionTargetBox="[264,1322,473,1246]" captionTargetId="figure@2.[264,1323,472,1248]" captionTargetPageId="2" captionText="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="1" pageNumber="40">Fig. 1</figureCitation>
). Pterostigma and veins dark brown.
</paragraph>
<paragraph id="764DEE3EFFF0B724FEDDF41DFC08FE43" blockId="1.[264,1323,1228,1894]" lastBlockId="2.[264,1323,284,390]" lastPageId="2" lastPageNumber="41" pageId="1" pageNumber="40">
<emphasis id="4486322CFFF0B727FEDDF41DFEFFF9B0" box="[312,383,1628,1653]" italics="true" pageId="1" pageNumber="40">Head.</emphasis>
Subquadrate posteriorly; parallel behind eyes; moderately punctate throughout (
<figureCitation id="EEC9F2BBFFF0B727FEF4F4C5FE03F95B" box="[273,387,1668,1694]" captionStart="FIGURE 1 – 3" captionStartId="2.[264,367,1260,1284]" captionTargetBox="[264,1322,473,1246]" captionTargetId="figure@2.[264,1323,472,1248]" captionTargetPageId="2" captionText="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="1" pageNumber="40">Figs. 2, 3</figureCitation>
). Eye small, barely protruding (
<figureCitation id="EEC9F2BBFFF0B727FD1AF4C5FCF1F95B" box="[767,881,1668,1694]" captionStart="FIGURE 1 – 3" captionStartId="2.[264,367,1260,1284]" captionTargetBox="[264,1322,473,1246]" captionTargetId="figure@2.[264,1323,472,1248]" captionTargetPageId="2" captionText="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="1" pageNumber="40">Figs. 2, 3</figureCitation>
). Ocellus small, interocellar distance ~2.75X lateral ocellus diameter, ocellocular distance ~4.5X lateral ocellus diameter (
<figureCitation id="EEC9F2BBFFF0B727FB1EF4EDFE99F92B" captionStart="FIGURE 1 – 3" captionStartId="2.[264,367,1260,1284]" captionTargetBox="[264,1322,473,1246]" captionTargetId="figure@2.[264,1323,472,1248]" captionTargetPageId="2" captionText="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="1" pageNumber="40">Fig. 2</figureCitation>
). Glabrous triangular region present anterior to median ocellus. Antennal scrobe carinate dorsally; tubercle present. Mandible tridentate (well­worn in
<typeStatus id="A949509CFFF0B727FC32F4BDFBC2F8D3" box="[983,1090,1788,1814]" pageId="1" pageNumber="40" type="holotype">holotype</typeStatus>
), broad and vertical throughout (
<figureCitation id="EEC9F2BBFFF0B727FE79F565FE66F8FB" box="[412,486,1828,1854]" captionStart="FIGURE 1 – 3" captionStartId="2.[264,367,1260,1284]" captionTargetBox="[264,1322,473,1246]" captionTargetId="figure@2.[264,1323,472,1248]" captionTargetPageId="2" captionText="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="1" pageNumber="40">Fig. 3</figureCitation>
). Mandible with slight ventral excision resulting in basal angulation. Clypeus truncate apically, distinctly curved dorsally, with 2 small tubercles set back from corners of truncation. Clypeus greatly depressed below dorsal margin of mandibles (
<figureCitation id="EEC9F2BBFFF3B724FB3AF35DFE98FE9B" captionStart="FIGURE 1 – 3" captionStartId="2.[264,367,1260,1284]" captionTargetBox="[264,1322,473,1246]" captionTargetId="figure@2.[264,1323,472,1248]" captionTargetPageId="2" captionText="FIGURE 1 – 3. Holotype of Sphaeropthalma nocticaro, sp. nov., 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view." httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="2" pageNumber="41">Fig. 2, 3</figureCitation>
). Malar space ~
<date id="024CC8FEFFF3B724FE00F305FD89FE9B" box="[485,521,324,350]" pageId="2" pageNumber="41">2X</date>
observed length of pedicel. First flagellomere ~2.25X length of pedicle. Second flagellomere slightly longer than first.
</paragraph>
<caption id="228DBEB6FFF3B724FEEDF6ADFDF9FAE3" httpUri="https://zenodo.org/record/169959/files/figure.png" pageId="2" pageNumber="41" targetBox="[264,1322,473,1246]" targetPageId="2">
<paragraph id="764DEE3EFFF3B724FEEDF6ADFDF9FAE3" blockId="2.[264,1322,1260,1318]" pageId="2" pageNumber="41">
<emphasis id="4486322CFFF3B724FEEDF6ADFE26FAC1" bold="true" box="[264,422,1260,1284]" pageId="2" pageNumber="41">FIGURE 1–3.</emphasis>
Holotype of
<taxonomicName id="B1F295BDFFF3B724FDDDF6ACFCD4FAC1" authority="Pitts, 2005" authorityName="Pitts" authorityYear="2005" box="[568,852,1261,1284]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="41" phylum="Arthropoda" rank="species" species="nocticaro" status="sp. nov.">
<emphasis id="4486322CFFF3B724FDDDF6ACFCD4FAC1" box="[568,852,1261,1284]" italics="true" pageId="2" pageNumber="41">Sphaeropthalma nocticaro</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5FB58F57FFF3B724FC87F6ADFC38FAC6" box="[866,952,1260,1283]" pageId="2" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
, 1. Habitus, lateral view; 2. Head, dorsal view; 3. Head, frontal view.
</paragraph>
</caption>
<paragraph id="764DEE3EFFF3B724FEDDF7C3FC0CF949" blockId="2.[264,1325,1410,1876]" pageId="2" pageNumber="41">
<emphasis id="4486322CFFF3B724FEDDF7C3FE3DFA5E" box="[312,445,1410,1435]" italics="true" pageId="2" pageNumber="41">
<taxonomicName id="B1F295BDFFF3B724FEDDF7C3FE39FA5E" box="[312,441,1410,1435]" class="Crustacea" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="41" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
.
</emphasis>
Moderately punctate. Notali complete. Parapsidal furrows present on posterior ~0.66 of mesonotum. Pronotum nitid anterior to epaulets, area almost reaching anterior margin. Propodeum coarsely punctate dorsally, vertical and lateral faces reticulate, becoming coarsely punctate ventrally. Dorsal face with median area longitudinally concave. Mesosternum unarmed. Metasternum truncate with small median tooth present. Coxae and trochanters unarmed. Marginal cell ~
<date id="024CC8FEFFF3B724FBE9F40BFBB1F9A1" box="[1036,1073,1610,1636]" pageId="2" pageNumber="41">2X</date>
length of stigma as measured along costa. Wings with 3 submarginal cells.
</paragraph>
<paragraph id="764DEE3EFFF3B725FEDDF4DBFBE5FE9B" blockId="2.[264,1325,1410,1876]" lastBlockId="3.[264,1322,284,350]" lastPageId="3" lastPageNumber="42" pageId="2" pageNumber="41">
<emphasis id="4486322CFFF3B724FEDDF4DBFE3AF976" box="[312,442,1690,1715]" italics="true" pageId="2" pageNumber="41">Metasoma.</emphasis>
T1 nodose with T2. T1 moderately punctate laterally and apically, otherwise sparsely punctate with interstitial area nitid. T2 with small dense punctures; interstitial width less than diameter of punctures, nitid. T3–T6 micropunctate, appearing granulate. T7 wider than long; nitid, except for apical margin with setigerous punctures bearing fringe of short hooked setae. S2 felt line absent, although micropunctures present in felt line area. Punctation of sternites similar to respective tergites, except S7 with small punctations. S7 wider than long. Apical margin of S7 broadly triangulate.
</paragraph>
<caption id="228DBEB6FFF2B725FEEDF1FBFDF0FC32" httpUri="https://zenodo.org/record/169960/files/figure.png" pageId="3" pageNumber="42" targetBox="[537,1080,443,917]" targetPageId="3">
<paragraph id="764DEE3EFFF2B725FEEDF1FBFDF0FC32" blockId="3.[264,1322,952,1015]" pageId="3" pageNumber="42">
<emphasis id="4486322CFFF2B725FEEDF1FBFE2AFC17" bold="true" box="[264,426,952,978]" pageId="3" pageNumber="42">FIGURE 4–5.</emphasis>
Holotype of
<taxonomicName id="B1F295BDFFF2B725FDA5F1FBFCDEFC14" authority="Pitts, 2005" authorityName="Pitts" authorityYear="2005" box="[576,862,954,977]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="nocticaro" status="sp. nov.">
<emphasis id="4486322CFFF2B725FDA5F1FBFCDEFC14" box="[576,862,954,977]" italics="true" pageId="3" pageNumber="42">Sphaeropthalma nocticaro</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5FB58F57FFF2B725FC8AF1FBFC47FC14" box="[879,967,954,977]" pageId="3" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
, genitalia. 4. Dorsal view, right, ventral view, left; 5. Lateral view.
</paragraph>
</caption>
<paragraph id="764DEE3EFFF2B725FEDDF617FB82FA42" blockId="3.[264,1323,1109,1895]" pageId="3" pageNumber="42">
<emphasis id="4486322CFFF2B725FEDDF617FE2EFBAA" box="[312,430,1110,1135]" italics="true" pageId="3" pageNumber="42">Genitalia.</emphasis>
Paramere arcuate, stout at base, tapering toward apex, curved dorsally (
<figureCitation id="EEC9F2BBFFF2B725FEF4F63CFE0EFB52" box="[273,398,1149,1175]" captionStart="FIGURE 4 – 5" captionStartId="3.[264,367,954,978]" captionTargetBox="[537,1080,443,917]" captionTargetId="figure@3.[486,1101,441,928]" captionTargetPageId="3" captionText="FIGURE 4 – 5. Holotype of Sphaeropthalma nocticaro, sp. nov., genitalia. 4. Dorsal view, right, ventral view, left; 5. Lateral view." httpUri="https://zenodo.org/record/169960/files/figure.png" pageId="3" pageNumber="42">Figs. 4, 5</figureCitation>
). Dense setae present on ventral margin of paramere (
<figureCitation id="EEC9F2BBFFF2B725FBA4F63CFB3EFB52" box="[1089,1214,1149,1175]" captionStart="FIGURE 4 – 5" captionStartId="3.[264,367,954,978]" captionTargetBox="[537,1080,443,917]" captionTargetId="figure@3.[486,1101,441,928]" captionTargetPageId="3" captionText="FIGURE 4 – 5. Holotype of Sphaeropthalma nocticaro, sp. nov., genitalia. 4. Dorsal view, right, ventral view, left; 5. Lateral view." httpUri="https://zenodo.org/record/169960/files/figure.png" pageId="3" pageNumber="42">Figs. 4, 5</figureCitation>
). Cuspis elongate, 0.8X free length of paramere, basal portion cylindrical, dilated and flatted towards apex, inner and outer margin with long dense setae, tips of setae plumose with many branches, basal portion with plumose tipped setae ventrally, small basal pit present with long simple to brachyplumose setae, dorsum asetose (
<figureCitation id="EEC9F2BBFFF2B725FC23F75CFBBAFAF2" box="[966,1082,1309,1335]" captionStart="FIGURE 4 – 5" captionStartId="3.[264,367,954,978]" captionTargetBox="[537,1080,443,917]" captionTargetId="figure@3.[486,1101,441,928]" captionTargetPageId="3" captionText="FIGURE 4 – 5. Holotype of Sphaeropthalma nocticaro, sp. nov., genitalia. 4. Dorsal view, right, ventral view, left; 5. Lateral view." httpUri="https://zenodo.org/record/169960/files/figure.png" pageId="3" pageNumber="42">Figs. 4, 5</figureCitation>
). Basal lobe present on cuspis, setose (
<figureCitation id="EEC9F2BBFFF2B725FE01F704FDD9FA9A" box="[484,601,1349,1375]" captionStart="FIGURE 4 – 5" captionStartId="3.[264,367,954,978]" captionTargetBox="[537,1080,443,917]" captionTargetId="figure@3.[486,1101,441,928]" captionTargetPageId="3" captionText="FIGURE 4 – 5. Holotype of Sphaeropthalma nocticaro, sp. nov., genitalia. 4. Dorsal view, right, ventral view, left; 5. Lateral view." httpUri="https://zenodo.org/record/169960/files/figure.png" pageId="3" pageNumber="42">Figs. 4, 5</figureCitation>
). Digitus cylindrical, short, minutely pubescent (
<figureCitation id="EEC9F2BBFFF2B725FB43F704FA9CFA9A" box="[1190,1308,1349,1375]" captionStart="FIGURE 4 – 5" captionStartId="3.[264,367,954,978]" captionTargetBox="[537,1080,443,917]" captionTargetId="figure@3.[486,1101,441,928]" captionTargetPageId="3" captionText="FIGURE 4 – 5. Holotype of Sphaeropthalma nocticaro, sp. nov., genitalia. 4. Dorsal view, right, ventral view, left; 5. Lateral view." httpUri="https://zenodo.org/record/169960/files/figure.png" pageId="3" pageNumber="42">Figs. 4, 5</figureCitation>
). Penis valve bidentate ventrally, not angulate dorsally (
<figureCitation id="EEC9F2BBFFF2B725FC66F72CFC73FA42" box="[899,1011,1389,1415]" captionStart="FIGURE 4 – 5" captionStartId="3.[264,367,954,978]" captionTargetBox="[537,1080,443,917]" captionTargetId="figure@3.[486,1101,441,928]" captionTargetPageId="3" captionText="FIGURE 4 – 5. Holotype of Sphaeropthalma nocticaro, sp. nov., genitalia. 4. Dorsal view, right, ventral view, left; 5. Lateral view." httpUri="https://zenodo.org/record/169960/files/figure.png" pageId="3" pageNumber="42">Figs. 4, 5</figureCitation>
).
</paragraph>
<paragraph id="764DEE3EFFF2B725FEDDF7D7FD81FA6A" blockId="3.[264,1323,1109,1895]" box="[312,513,1429,1455]" pageId="3" pageNumber="42">
<emphasis id="4486322CFFF2B725FEDDF7D7FE11FA6A" box="[312,401,1430,1455]" italics="true" pageId="3" pageNumber="42">Length.</emphasis>
~
<quantity id="B10A43DBFFF2B725FE42F7D4FE7DFA6A" box="[423,509,1429,1455]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="42" unit="mm" value="14.0">14 mm</quantity>
.
</paragraph>
<paragraph id="764DEE3EFFF2B725FEDDF7FFFD99FA12" blockId="3.[264,1323,1109,1895]" box="[312,537,1469,1495]" pageId="3" pageNumber="42">
<emphasis id="4486322CFFF2B725FEDDF7FFFE17FA12" box="[312,407,1470,1495]" italics="true" pageId="3" pageNumber="42">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="3EE8BDB5FFF2B725FEDDF7A7FC2DF98A" pageId="3" pageNumber="42" type="materials_examined">
<paragraph id="764DEE3EFFF2B725FEDDF7A7FC2DF98A" blockId="3.[264,1323,1109,1895]" pageId="3" pageNumber="42">
<emphasis id="4486322CFFF2B725FEDDF7A7FE6AFA3A" box="[312,490,1510,1535]" italics="true" pageId="3" pageNumber="42">
<typeStatus id="A949509CFFF2B725FEDDF7A7FEEFFA3A" box="[312,367,1510,1535]" pageId="3" pageNumber="42">Type</typeStatus>
Material.
</emphasis>
<emphasis id="4486322CFFF2B725FE10F7A4FDF1FA3A" bold="true" box="[501,625,1509,1535]" pageId="3" pageNumber="42">
<typeStatus id="A949509CFFF2B725FE10F7A4FDEBFA3A" box="[501,619,1509,1535]" pageId="3" pageNumber="42" type="holotype">Holotype</typeStatus>
:
</emphasis>
“Temascal, Oax[aca], Mex[ico],
<date id="024CC8FEFFF2B725FBE8F7A4FBFCFA3A" box="[1037,1148,1509,1535]" pageId="3" pageNumber="42" value="1964-06-22">VI­22­64</date>
, D.H. Janzen, Collector”. Deposited into
<collectionCode id="10E376FBFFF2B725FDA8F44CFD12F9E2" box="[589,658,1549,1575]" pageId="3" pageNumber="42">CISC</collectionCode>
.
<typeStatus id="A949509CFFF2B725FD44F44CFC93F9E2" box="[673,787,1549,1575]" pageId="3" pageNumber="42" type="paratype">
<emphasis id="4486322CFFF2B725FD44F44CFC93F9E2" bold="true" box="[673,787,1549,1575]" pageId="3" pageNumber="42">Paratype</emphasis>
</typeStatus>
: “
<collectingCountry id="0EE5AEAEFFF2B725FCD5F44CFC35F9E2" box="[816,949,1549,1575]" name="Guatemala" pageId="3" pageNumber="42">Guatemala</collectingCountry>
, Petén, Ruinas de Tikal,
<date id="024CC8FEFFF2B725FB0DF44CFEDFF98A" pageId="3" pageNumber="42" value="1927-08-29">27/29 August</date>
, 1974, E.M. Fisher, collr.” Deposited into
<collectionCode id="10E376FBFFF2B725FCB5F474FC29F98A" LSID="urn:lsid:biocol.org:col:34802" box="[848,937,1589,1615]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="3" pageNumber="42">LACM</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="3EE8BDB5FFF2B725FEDDF41FFD24F95A" pageId="3" pageNumber="42" type="etymology">
<paragraph id="764DEE3EFFF2B725FEDDF41FFD24F95A" blockId="3.[264,1323,1109,1895]" pageId="3" pageNumber="42">
<emphasis id="4486322CFFF2B725FEDDF41FFE3BF9B2" box="[312,443,1630,1655]" italics="true" pageId="3" pageNumber="42">Etymology.</emphasis>
From the Latin
<emphasis id="4486322CFFF2B725FD9DF41FFD39F9B2" box="[632,697,1630,1655]" italics="true" pageId="3" pageNumber="42">noctu</emphasis>
“by night” and Latin
<emphasis id="4486322CFFF2B725FC5CF41FFC6EF9B2" box="[953,1006,1630,1655]" italics="true" pageId="3" pageNumber="42">caro</emphasis>
“flesh”, in reference to the black coloration of the integument.
</paragraph>
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<emphasis id="4486322CFFF2B725FEDDF4EFFE26F902" box="[312,422,1710,1735]" italics="true" pageId="3" pageNumber="42">Remarks.</emphasis>
This species easily keys out to the subgenus
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<emphasis id="4486322CFFF2B725FC31F4EFFB15F902" box="[980,1173,1710,1735]" italics="true" pageId="3" pageNumber="42">Sphaeropthalma</emphasis>
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(now the
<taxonomicName id="B1F295BDFFF2B725FAF1F4EFFE21F92A" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="pensylvanica">
<emphasis id="4486322CFFF2B725FAF1F4EFFE21F92A" italics="true" pageId="3" pageNumber="42">S. pensylvanica</emphasis>
</taxonomicName>
species­group; see
<bibRefCitation id="126393CFFFF2B725FD6EF494FCCAF92A" author="Pitts" box="[651,842,1749,1775]" pageId="3" pageNumber="43" refString="Pitts, J. P., Parker, F. D. &amp; Pitts-Singer, T. L. (2004) A review of the Sphaeropthalma uro speciesgroup (Hymenoptera: Mutillidae), with taxonomic changes. Journal of the Kansas Entomological Society, 77, 223 - 234." type="journal article" year="2004">
Pitts
<emphasis id="4486322CFFF2B725FD2CF497FC85F92A" box="[713,773,1750,1775]" italics="true" pageId="3" pageNumber="42">et al.</emphasis>
2004
</bibRefCitation>
) in
<bibRefCitation id="126393CFFFF2B725FC9EF494FB1FF92A" author="Manley" box="[891,1183,1749,1775]" pageId="3" pageNumber="43" refString="Manley, D. G. &amp; Pitts, J. P. (2002) Key to the genera and subgenera of Mutillidae of America North of Mexico. Journal of Hymenoptera Research, 11, 72 - 101." type="journal article" year="2002">Manley and Pitts (2002)</bibRefCitation>
. In general, this species is quite similar morphologically to other species in this species­group, except for the coloration and white plumose setal mentioned previously. This species is also probably diurnal like the other members of this group as deduced from the very small ocelli and dark integumental coloration. The genitalia also differ from the other species. For
<taxonomicName id="B1F295BDFFF5B722FEDDF305FE44FE98" authorityName="Pitts" authorityYear="2005" box="[312,452,324,349]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="nocticaro">
<emphasis id="4486322CFFF5B722FEDDF305FE44FE98" box="[312,452,324,349]" italics="true" pageId="4" pageNumber="43">S. nocticaro</emphasis>
</taxonomicName>
, the paramere has ventral setae present much further towards the apex, the basal portion of cuspis is not as densely setose ventrally, and the digitus is much larger in comparison to the cuspis. There is also a slight difference in the shape of the apex of the cuspis between
<taxonomicName id="B1F295BDFFF5B722FE27F3FDFDCEFE10" authorityName="Pitts" authorityYear="2005" box="[450,590,444,469]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="nocticaro">
<emphasis id="4486322CFFF5B722FE27F3FDFDCEFE10" box="[450,590,444,469]" italics="true" pageId="4" pageNumber="43">S. nocticaro</emphasis>
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and the other species.
</paragraph>
<paragraph id="764DEE3EFFF5B722FEDDF3A5FB8DFC4B" blockId="4.[264,1323,284,910]" pageId="4" pageNumber="43">
As with most organisms, morphological variation has been documented for many of the
<taxonomicName id="B1F295BDFFF5B722FED0F04DFE76FDE0" box="[309,502,524,549]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="4486322CFFF5B722FED0F04DFE76FDE0" box="[309,502,524,549]" italics="true" pageId="4" pageNumber="43">Sphaeropthalma</emphasis>
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species. Because only two specimens are available, variation cannot be adequately addressed for
<taxonomicName id="B1F295BDFFF5B722FD8EF075FD7CFD88" authorityName="Pitts" authorityYear="2005" box="[619,764,564,589]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="nocticaro">
<emphasis id="4486322CFFF5B722FD8EF075FD7CFD88" box="[619,764,564,589]" italics="true" pageId="4" pageNumber="43">S. nocticaro</emphasis>
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at this time. Furthermore, the
<typeStatus id="A949509CFFF5B722FB68F075FB74FD8B" box="[1165,1268,564,590]" pageId="4" pageNumber="43" type="holotype">holotype</typeStatus>
and
<typeStatus id="A949509CFFF5B722FEEDF01DFEEDFDB3" box="[264,365,604,630]" pageId="4" pageNumber="43" type="paratype">paratype</typeStatus>
are quite similar morphologically, differing only in wear of the mandibular teeth. Normal variation with male
<taxonomicName id="B1F295BDFFF5B722FDB9F0C5FC9DFD58" box="[604,797,644,669]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="4486322CFFF5B722FDB9F0C5FC9DFD58" box="[604,797,644,669]" italics="true" pageId="4" pageNumber="43">Sphaeropthalma</emphasis>
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usually includes a large range in body size. For smaller individuals, the diameter and coarseness of punctations are normally reduced, and the integumental coloration is lighter (
<bibRefCitation id="126393CFFFF5B722FCFEF095FC5DFD2B" author="Ferguson" box="[795,989,724,750]" pageId="4" pageNumber="43" refString="Ferguson, W. E. (1962) Biological characteristics of the mutillid subgenus Photopsis Blake and their systematic values (Hymenoptera). University of California Publications in Entomology, 27, 1 - 92." type="journal article" year="1962">Ferguson, 1962</bibRefCitation>
). Variation of this
<typeStatus id="A949509CFFF5B722FB3DF095FA8BFD2B" box="[1240,1291,724,750]" pageId="4" pageNumber="43">type</typeStatus>
is expected for
<taxonomicName id="B1F295BDFFF5B722FE4EF0BDFDBBFCD0" authorityName="Pitts" authorityYear="2005" box="[427,571,764,789]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="nocticaro">
<emphasis id="4486322CFFF5B722FE4EF0BDFDBBFCD0" box="[427,571,764,789]" italics="true" pageId="4" pageNumber="43">S. nocticaro</emphasis>
</taxonomicName>
once more specimens are found. The diagnostic characters, however, used to differentiate this species from other members of the
<taxonomicName id="B1F295BDFFF5B722FB8FF165FE21FCA0" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="pensylvanica">
<emphasis id="4486322CFFF5B722FB8FF165FE21FCA0" italics="true" pageId="4" pageNumber="43">Sphaeropthalma pensylvanica</emphasis>
</taxonomicName>
species­group are unlikely to vary given the normal variation seen in other members of this group, as well as for
<taxonomicName id="B1F295BDFFF5B722FD27F135FC03FC48" box="[706,899,884,909]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="4486322CFFF5B722FD27F135FC03FC48" box="[706,899,884,909]" italics="true" pageId="4" pageNumber="43">Sphaeropthalma</emphasis>
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as a whole.
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116
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Westwood, 1842
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<paragraph pageId="0" pageNumber="8051">
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Type status:
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. Taxon: scientificName: Megapomponiaimperatoria (Westwood, 1842); Location: continent: Asia; locality:
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<paragraph pageId="0" pageNumber="8051">Distribution</paragraph>
<paragraph pageId="0" pageNumber="8051">[Metcalf, 1963] East Indies; Indian Islands; Nepal; India; Assam; Malaya; Borneo; Sumatra; Penang; Cambodia; Laos; Siam; Malay Peninsula; Java; Malay Islands; Malay States; Sarawak; Malacca; Perak; Singapore Island; Johore. [Duffels and van der Laan, 1985] Nepal. [Sanborn, 2014] Thailand, India, Nepal, Malaysia, Borneo, Cambodia, Laos, Sarawak, Indonesia, Indian Islands, Peninsular Malaysia.</paragraph>
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<paragraph pageId="0" pageNumber="8051">
Authority:
<bibRefCitation author="Westwood, J. O." journalOrPublisher="The Annals and Magazine of Natural History" pageId="0" pageNumber="8051" pagination="118 - 119" title="Insectorm novorum centuria, auctore" volume="9" year="1842">Westwood 1842</bibRefCitation>
; Multiple localities listed in species description. Not from India:
<bibRefCitation author="Metcalf, Z. P." journalOrPublisher="North Carolina State College Contribution" pageId="0" pageNumber="8051" pagination="587 - 919" title="General catalogue of the Homoptera. Fascicle VIII. Cicadoidea. Part 1. Cicadidae. Section II. Gaeaninae and Cicadinae." volume="1502" year="1963">Metcalf (1963)</bibRefCitation>
, in error, listed &quot;India, Assam, Nepal&quot; in reference to
<bibRefCitation author="Walker, Francis" journalOrPublisher="Edward Newman, London" pageId="0" pageNumber="8051" title="List of the specimens of Homopterous insects in the collection of the British Museum. Part I" year="1850">Walker (1850)</bibRefCitation>
. The type description does not give a specific locality (East Indies and the Indian Islands), while recent literature indicates that this species is confined to the Malaysian Peninsula (
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). Subsequent references to India (and Nepal) in
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,
<bibRefCitation author="Duffels, J. P" journalOrPublisher="Springer Netherlands" pageId="0" pageNumber="8051" title="Catalogue of the Cicadoidea (Homoptera, Auchenorhyncha) 1956 - 1980" year="1985">Duffels and van der Laan (1985)</bibRefCitation>
and
<bibRefCitation author="Sanborn, A" journalOrPublisher="Elsevier" pageId="0" pageNumber="8051" title="Catalogue of the Cicadoidea (Hemiptera: Auchenorrhyncha)" year="2014">Sanborn (2014)</bibRefCitation>
are likely to be based on this previous error.
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<heading id="2D0581F899408350089EE9FBFDEAFC7D" bold="true" box="[151,595,909,936]" fontSize="11" level="1" pageId="1" pageNumber="277" reason="1">
<taxonomicName id="B1F24D1799408350089EE9FBFDEAFC7D" ID-CoL="9X3LR" authority="Kosterin, 2016" authorityName="Kosterin" authorityYear="2016" box="[151,595,909,936]" class="Insecta" family="Gomphidae" genus="Microgomphus" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="277" phylum="Arthropoda" rank="species" species="alani">
<emphasis id="4486EA8699408350089EE9FBFDEAFC7D" bold="true" box="[151,595,909,936]" pageId="1" pageNumber="277">
<emphasis id="4486EA8699408350089EE9FBFE2CFC72" bold="true" box="[151,405,909,936]" italics="true" pageId="1" pageNumber="277">Microgomphus alani</emphasis>
Kosterin, 2016
</emphasis>
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<figureCitation id="EEC92A1199408350089EE9C7FF40FC11" box="[151,249,945,971]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="2.[151,250,1450,1475]" captionStartId-1="3.[151,250,1924,1949]" captionStartId-2="4.[151,250,1902,1927]" captionStartId-3="5.[151,250,1906,1931]" captionStartId-4="6.[151,250,1669,1694]" captionStartId-5="8.[151,250,1901,1926]" captionStartId-6="9.[151,250,1973,1998]" captionTargetBox-0="[151,1436,553,1431]" captionTargetBox-1="[151,1431,805,1905]" captionTargetBox-2="[266,1321,166,1874]" captionTargetBox-3="[151,1424,792,1868]" captionTargetBox-4="[170,1436,174,1602]" captionTargetBox-5="[161,1426,166,1866]" captionTargetBox-6="[345,1242,166,1896]" captionTargetId-0="figure@2.[151,1436,553,1431]" captionTargetId-1="figure@3.[151,1436,805,1905]" captionTargetId-2="figure@4.[266,1321,166,1883]" captionTargetId-3="figure@5.[151,1436,769,1886]" captionTargetId-4="figure@6.[151,1436,174,1650]" captionTargetId-5="figure@8.[161,1426,166,1881]" captionTargetId-6="figure@9.[345,1242,166,1953]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="8" captionTargetPageId-6="9" captionText-0="FIGURE 1. Male (a, d–e) and female (b–c) specimens of Microgomphus alani in live colours from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–e) of Cambodia, 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e). Not to scale" captionText-1="FIGURE 2. Dry male (a, d–e) and female (b–c, f) specimens of Microgomphus alani from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–f) of Cambodia, collected on 3 viii 2016 (f), 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e)." captionText-2="FIGURE 3. Details of male specimens of Microgomphus alani collected on 26 vi 2018 in Mondulkiri Province (a–d; the same specimen as in Fig. 2d) and on 17 vi 2018 in the Phnom Kulen Mts (a–h, the same specimen as in Fig. 1a and Fig 2a): a,e—head and thorax, lateral view; b,f—pterothorax, dorsal view; c,g—head, frontal (c) and dorsofrontal (g) view; d,h—secondary genitalia, lateral view. Scale bars. from top to bottom, 4 mm, 2 mm, 2 mm and 0.5 mm." captionText-3="FIGURE 4. Anal appendages, in dorsal (left) and lateral (right) view, of mature males of Microgomphus alani, collected on 26 vi 2018 in Mondulkiri Province (a–b, respectively corresponding to the specimens in Fig. 2d and Fig. 2e) and on 17 vi 2018 in the Phnom Kulen Mts (c, correspond to Fig. 1a and Fig 2a). Scale bar 1 mm." captionText-4="FIGURE 5. Frontal view of head (left) and occiput (right) of female specimens of Microgomphus alani collected on 3 viii 2016 in Mondulkiri Province (a; the same specimen as in Fig. 2f) and in the Phnom Kulen Mts on 11 vi 2018 (b, the same specimen as in Figs 1c, 2c) and 30 vi 2018 (c, the same specimen as in Figs 1b, 2b). Scale bar 1 mm." captionText-5="FIGURE 6. Details of female specimens of Microgomphus alani collected in Mondulkiri Province on 3 viii 2016 (c,e; the same specimen as in Fig. 2f) and in the Phnom Kulen Mts on 11 vi 2018 (a,f; the same specimen as in Fig. 1c and Fig 2c) and on 30 vi 2018 (b,d,g; the same specimen as in Figs 1b, 2b): a–b—head and thorax, lateral view; c–d—pterothorax, dorsal view; e–g—vulvar lamina, ventral view. Scale bars, from top to bottom, 4 mm, 2 mm and 0.5 mm." captionText-6="FIGURE 7. Hind legs of Microgomphus alani, a—male from Mondulkiri Province, 13 vi 2014; b—female from Modulkiri Province, 3 viii 2016; c—female from Phnom Kulen Mts, 30 vi 2018. Scale bar 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3558041" figureDoi-1="http://doi.org/10.5281/zenodo.3558043" figureDoi-2="http://doi.org/10.5281/zenodo.3558045" figureDoi-3="http://doi.org/10.5281/zenodo.3558047" figureDoi-4="http://doi.org/10.5281/zenodo.3558049" figureDoi-5="http://doi.org/10.5281/zenodo.3558051" figureDoi-6="http://doi.org/10.5281/zenodo.3558053" httpUri-0="https://zenodo.org/record/3558041/files/figure.png" httpUri-1="https://zenodo.org/record/3558043/files/figure.png" httpUri-2="https://zenodo.org/record/3558045/files/figure.png" httpUri-3="https://zenodo.org/record/3558047/files/figure.png" httpUri-4="https://zenodo.org/record/3558049/files/figure.png" httpUri-5="https://zenodo.org/record/3558051/files/figure.png" httpUri-6="https://zenodo.org/record/3558053/files/figure.png" pageId="1" pageNumber="277">Figs 1–7</figureCitation>
</paragraph>
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<subSubSection id="3EE8651F99408350089EE983FEEDFAFB" pageId="1" pageNumber="277" type="reference_group">
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<treatmentCitation id="F753108599408350089EE983FDFBFBD7" author="Asahina, S." box="[151,578,1012,1037]" page="15" pageId="1" pageNumber="277" year="1986">
<taxonomicName id="B1F24D1799408350089EE983FD98FBD7" ID-CoL="3KB8R" authority="- Asahina (1986)" authorityName="Asahina" authorityYear="1986" box="[151,545,1012,1037]" class="Insecta" family="Gomphidae" genus="Heliogomphus" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="277" phylum="Arthropoda" rank="species" species="selysi">
<emphasis id="4486EA8699408350089EE983FED1FBD7" box="[151,360,1013,1037]" italics="true" pageId="1" pageNumber="277">Heliogomphus selysi</emphasis>
—
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: 15
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–16, figs 30–32, 37–38: (most probably a misidentified female from
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, Chieng Dao—description, drawings of the head, thorax, hind femora, vulvar lamina).
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</paragraph>
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,
</emphasis>
<taxonomicNameLabel id="5FB557FD99408350097FEE45FE10FB91" box="[374,425,1074,1099]" pageId="1" pageNumber="277" rank="species">sp. n.</taxonomicNameLabel>
—
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<bibRefCitation id="12634B659940835009CAEE44FDCCFB91" author="Kosterin, O. E." box="[451,629,1074,1099]" pageId="1" pageNumber="277" pagination="341 - 350" refId="ref9406" refString="Kosterin, O. E. (2016 a) Microgomphus alani (Odonata, Gomphidae) sp. nov. from Cambodia. Zootaxa, 4114 (3), 341 - 350. https: // doi. org / 10.11646 / zootaxa. 4114.3.9" type="journal article" year="2016">Kosterin (2016a)</bibRefCitation>
: 341
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–450, figs 1–4 (original description by two males from
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of
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, collected in teneral state and hardened in captivity, differential diagnosis, habitat, distribution; illustrations of general habitus, wings, head, thorax, secondary genitalia including vesica spermalis and anal appendages).
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</paragraph>
<paragraph id="764D369499408350089EEEF8FDD3FB1F" blockId="1.[151,1436,1012,1313]" pageId="1" pageNumber="277">
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<taxonomicName id="B1F24D1799408350089EEEF8FE33FB7C" ID-CoL="42X2T" authorityName="Karube" authorityYear="2000" box="[151,394,1166,1190]" class="Insecta" family="Gomphidae" genus="Microgomphus" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="277" phylum="Arthropoda" rank="species" species="jurzitzai">
<emphasis id="4486EA8699408350089EEEF8FE33FB7C" box="[151,394,1166,1190]" italics="true" pageId="1" pageNumber="277">Microgomphus jurzitzai</emphasis>
</taxonomicName>
(misidentification)—
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<bibRefCitation id="12634B65994083500A6FEEF8FCA1FB7D" author="Kosterin, O. E." box="[614,792,1166,1191]" pageId="1" pageNumber="277" pagination="1 - 86" refId="ref9453" refString="Kosterin, O. E. (2016 b) A survey of Odonata of Mondulkiri, the elevated eastern province of Cambodia, for ten days in June 2014. International Dragonfly Fund Report, 98, 1 - 86." type="journal article" year="2016">Kosterin (2016b)</bibRefCitation>
: 60
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–61 (1 misidentified
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reported for
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of
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, the Monorom Waterfall area).
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</paragraph>
<paragraph id="764D369499408350089EEEBAFEEDFAFB" blockId="1.[151,1436,1012,1313]" pageId="1" pageNumber="277">
<taxonomicName id="B1F24D1799408350089EEEBAFE92FB3E" ID-CoL="62TNM" authorityName="Laidlaw" authorityYear="1922" box="[151,299,1228,1252]" class="Insecta" family="Gomphidae" genus="Heliogomphus" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="4486EA8699408350089EEEBAFE92FB3E" box="[151,299,1228,1252]" italics="true" pageId="1" pageNumber="277">Heliogomphus</emphasis>
</taxonomicName>
sp. (misidentification)—
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: 73–75, fig. 58 (1 misidentified
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reported for
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of
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(the Buu Sraa Waterfal area); general habitus, head, thorax, fore wing base, genital lamina illustrated by photographs).
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<emphasis id="4486EA8699408350089EEF39FD2EFAB0" bold="true" box="[151,663,1359,1386]" pageId="1" pageNumber="277">
Specimens examined.
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:
</emphasis>
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, the
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(in alcohol)—
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,
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,
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ENE of
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, ‘
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Rivulet’,
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,
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[
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,
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a.s.l.,
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(
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: figs 1a,c, 2a–f,h, 3a–b,e–f, 4).
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, the paratype—Cambodia,
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,
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SW Thma Bang village, the ‘
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River’, the slow reach upstream of the bridge,
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,
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[
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,
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],
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a.s.l.,
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(
<bibRefCitation id="12634B65994083500B65EFA9FBA7FA23" author="Kosterin, O. E." box="[876,1054,1503,1529]" pageId="1" pageNumber="277" pagination="341 - 350" refId="ref9406" refString="Kosterin, O. E. (2016 a) Microgomphus alani (Odonata, Gomphidae) sp. nov. from Cambodia. Zootaxa, 4114 (3), 341 - 350. https: // doi. org / 10.11646 / zootaxa. 4114.3.9" type="journal article" year="2016">Kosterin 2016a</bibRefCitation>
: figs 1b;
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; 3c–d).
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Phnom Kulen Mts,
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:
</emphasis>
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,
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,
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,
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)—
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,
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, Phnom Kulen Mts, the
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River just upstream of the bridge,
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W of Preah Ang Thom,
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,
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,
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a.s.l.,
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(
<figureCitation id="EEC92A1199408350095CEC3DFE13F9BC" box="[341,426,1611,1638]" captionStart="FIGURE 1" captionStartId="2.[151,250,1450,1475]" captionTargetBox="[151,1436,553,1431]" captionTargetId="figure@2.[151,1436,553,1431]" captionTargetPageId="2" captionText="FIGURE 1. Male (a, d–e) and female (b–c) specimens of Microgomphus alani in live colours from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–e) of Cambodia, 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e). Not to scale" figureDoi="http://doi.org/10.5281/zenodo.3558041" httpUri="https://zenodo.org/record/3558041/files/figure.png" pageId="1" pageNumber="277">Figs 1a</figureCitation>
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e–h, 4c)—
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, Phnom Kulen Mts, the
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River downstream of the waterfall,
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NNW of Preah Ang Thom,
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,
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,
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a.s.l.,
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.
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(
<figureCitation id="EEC92A11994083500896ECE5FF4DF977" box="[159,244,1683,1709]" captionStart="FIGURE 1" captionStartId="2.[151,250,1450,1475]" captionTargetBox="[151,1436,553,1431]" captionTargetId="figure@2.[151,1436,553,1431]" captionTargetPageId="2" captionText="FIGURE 1. Male (a, d–e) and female (b–c) specimens of Microgomphus alani in live colours from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–e) of Cambodia, 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e). Not to scale" figureDoi="http://doi.org/10.5281/zenodo.3558041" httpUri="https://zenodo.org/record/3558041/files/figure.png" pageId="1" pageNumber="277">Figs 1b</figureCitation>
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, Phnom Kulen Mts, a brook, the left
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River tributary,
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SE of Preah Ang Thom, 13.551–554 N, 104.117–120 E,
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a.s.l.,
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<collectingRegion id="B436F876994083500D52ECCEFEE6F92C" country="Cambodia" name="Mondolkiri" pageId="1" pageNumber="277">Mondulkiri Province</collectingRegion>
:
</emphasis>
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, Monorom Waterfall area, the left grassy slope of the valley of the ‘Ochracea rivulet’, the left tributary of the main river,
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WSW of Sen Monorom,
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,
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a.s.l.,
<date id="024C10549940835009DBED52FDFEF8E4" box="[466,583,1827,1854]" pageId="1" pageNumber="277" value="2014-06-13">13 vi 2014</date>
(
<bibRefCitation id="12634B65994083500A5DED55FCBCF8E7" author="Kosterin, O. E." box="[596,773,1827,1853]" pageId="1" pageNumber="277" pagination="1 - 86" refId="ref9453" refString="Kosterin, O. E. (2016 b) A survey of Odonata of Mondulkiri, the elevated eastern province of Cambodia, for ten days in June 2014. International Dragonfly Fund Report, 98, 1 - 86." type="journal article" year="2016">Kosterin 2016b</bibRefCitation>
, misidentified as
<taxonomicName id="B1F24D17994083500BCFED52FBFFF8E7" authorityName="Karube" authorityYear="2000" box="[966,1094,1828,1853]" class="Insecta" family="Gomphidae" genus="Microgomphus" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="277" phylum="Arthropoda" rank="species" species="jurzitzai">
<emphasis id="4486EA86994083500BCFED52FBFFF8E7" box="[966,1094,1828,1853]" italics="true" pageId="1" pageNumber="277">M. jurzitzai</emphasis>
</taxonomicName>
; coordinates mentioned in that source for the relevant locality 4a were slightly incorrect).
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(
<bibRefCitation id="12634B65994083500B5BED31FBBDF8BB" author="Kosterin, O. E." box="[850,1028,1863,1889]" pageId="1" pageNumber="277" pagination="1 - 86" refId="ref9453" refString="Kosterin, O. E. (2016 b) A survey of Odonata of Mondulkiri, the elevated eastern province of Cambodia, for ten days in June 2014. International Dragonfly Fund Report, 98, 1 - 86." type="journal article" year="2016">Kosterin 2016b</bibRefCitation>
: fig. 58;
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,
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,
<figureCitation id="EEC92A11994083500CEDED31FAADF8BB" box="[1252,1300,1863,1889]" captionStart="FIGURE 6" captionStartId="8.[151,250,1901,1926]" captionTargetBox="[161,1426,166,1866]" captionTargetId="figure@8.[161,1426,166,1881]" captionTargetPageId="8" captionText="FIGURE 6. Details of female specimens of Microgomphus alani collected in Mondulkiri Province on 3 viii 2016 (c,e; the same specimen as in Fig. 2f) and in the Phnom Kulen Mts on 11 vi 2018 (a,f; the same specimen as in Fig. 1c and Fig 2c) and on 30 vi 2018 (b,d,g; the same specimen as in Figs 1b, 2b): a–b—head and thorax, lateral view; c–d—pterothorax, dorsal view; e–g—vulvar lamina, ventral view. Scale bars, from top to bottom, 4 mm, 2 mm and 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3558051" httpUri="https://zenodo.org/record/3558051/files/figure.png" pageId="1" pageNumber="277">6c,e</figureCitation>
,
<figureCitation id="EEC92A11994083500D17ED3EFA8EF8B8" box="[1310,1335,1864,1890]" captionStart="FIGURE 7" captionStartId="9.[151,250,1973,1998]" captionTargetBox="[345,1242,166,1896]" captionTargetId="figure@9.[345,1242,166,1953]" captionTargetPageId="9" captionText="FIGURE 7. Hind legs of Microgomphus alani, a—male from Mondulkiri Province, 13 vi 2014; b—female from Modulkiri Province, 3 viii 2016; c—female from Phnom Kulen Mts, 30 vi 2018. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3558053" httpUri="https://zenodo.org/record/3558053/files/figure.png" pageId="1" pageNumber="277">7c</figureCitation>
)—Cam- bodia,
<collectingRegion id="B436F8769940835008ECED1DFE6AF85F" box="[229,467,1899,1925]" country="Cambodia" name="Mondolkiri" pageId="1" pageNumber="277">Mondulkiri Province</collectingRegion>
, just downstream of Buu Sraa Waterfall, the
<emphasis id="4486EA86994083500BDEED1DFC59F85C" bold="true" box="[983,992,1899,1926]" pageId="1" pageNumber="277">‘</emphasis>
Loringae brook’ lowermost reaches, an open grassy area at the main river right bank,
<geoCoordinate id="13C65053994083500AA1EDF9FC9AF870" box="[680,803,1935,1962]" degrees="12.5717" direction="north" orientation="latitude" pageId="1" pageNumber="277" precision="5" value="12.5717">12.5717 N</geoCoordinate>
,
<geoCoordinate id="13C65053994083500B39EDE6FC0DF870" box="[816,948,1936,1962]" degrees="107.4173" direction="east" orientation="longitude" pageId="1" pageNumber="277" precision="5" value="107.4173">107.4173 E</geoCoordinate>
,
<quantity id="B10A9B71994083500BCBEDE6FBB2F870" box="[962,1035,1936,1962]" metricMagnitude="2" metricUnit="m" metricValue="4.32" pageId="1" pageNumber="277" unit="m" value="432.0">432 m</quantity>
a.s.l.,
<date id="024C1054994083500C5EEDE6FB6DF873" box="[1111,1236,1935,1962]" pageId="1" pageNumber="277" value="2016-08-03">3 viii 2016</date>
(
<bibRefCitation id="12634B65994083500CEDEDF9FA21F873" author="Kosterin, O. E." box="[1252,1432,1935,1961]" pageId="1" pageNumber="277" pagination="1 - 86" refId="ref9453" refString="Kosterin, O. E. (2016 b) A survey of Odonata of Mondulkiri, the elevated eastern province of Cambodia, for ten days in June 2014. International Dragonfly Fund Report, 98, 1 - 86." type="journal article" year="2016">Kosterin 2016b</bibRefCitation>
, misidentified as
<taxonomicName id="B1F24D1799408350095BEDC2FE4CF814" authorityName="Laidlaw" authorityYear="1922" box="[338,501,1972,1998]" class="Insecta" family="Gomphidae" genus="Heliogomphus" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="4486EA8699408350095BEDC2FE4CF814" box="[338,501,1972,1998]" italics="true" pageId="1" pageNumber="277">Heliogomphus</emphasis>
</taxonomicName>
sp.).
<specimenCount id="60F4FD1D994083500A3FEDC2FDE7F817" box="[566,606,1971,1998]" pageId="1" pageNumber="277" type="male">1 ♂</specimenCount>
—
<collectingCountry id="0EE57604994083500A75EDC5FD4AF817" box="[636,755,1971,1997]" name="Cambodia" pageId="1" pageNumber="277">Cambodia</collectingCountry>
,
<collectingRegion id="B436F876994083500AF6EDC5FC54F817" box="[767,1005,1971,1997]" country="Cambodia" name="Mondolkiri" pageId="1" pageNumber="277">Mondulkiri Province</collectingRegion>
,
<quantity id="B10A9B71994083500BF3EDC2FBEEF817" box="[1018,1111,1971,1998]" metricMagnitude="4" metricUnit="m" metricValue="1.28" pageId="1" pageNumber="277" unit="km" value="12.8">12.8 km</quantity>
NE of Sen Monorom, lower reaches of a brook—right tributary of Pulung (?) River, 12.5186–5189 N, 107.2931–2954 E,
<quantity id="B10A9B71994083500C96EDA1FAA4F828" box="[1183,1309,2007,2034]" metricMagnitude="2" metricUnit="m" metricValue="5.365" metricValueMax="5.4399999999999995" metricValueMin="5.29" pageId="1" pageNumber="277" unit="m" value="536.5" valueMax="544.0" valueMin="529.0">529–544 m</quantity>
a.s.l.,
<date id="024C1054994083530D6DEDA1FF6AFF69" lastPageId="2" lastPageNumber="278" pageId="1" pageNumber="277" value="2018-06-25">25 vi 2018</date>
.
<specimenCount id="60F4FD1D9943835308D5EAEFFEA3FF69" box="[220,282,153,179]" pageId="2" pageNumber="278" type="male">2 ♂♂</specimenCount>
(
<figureCitation id="EEC92A1199438353092EEAEFFED0FF69" box="[295,361,153,179]" captionStart="FIGURE 1" captionStartId="2.[151,250,1450,1475]" captionTargetBox="[151,1436,553,1431]" captionTargetId="figure@2.[151,1436,553,1431]" captionTargetPageId="2" captionText="FIGURE 1. Male (a, d–e) and female (b–c) specimens of Microgomphus alani in live colours from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–e) of Cambodia, 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e). Not to scale" figureDoi="http://doi.org/10.5281/zenodo.3558041" httpUri="https://zenodo.org/record/3558041/files/figure.png" pageId="2" pageNumber="278">Figs 1</figureCitation>
d–e, 2d–e, 3a–d, 4a–b)—
<collectingCountry id="0EE57604994383530A8AEAEFFD43FF6E" box="[643,762,153,180]" name="Cambodia" pageId="2" pageNumber="278">Cambodia</collectingCountry>
,
<collectingRegion id="B436F876994383530B0DEAEFFC56FF69" box="[772,1007,153,179]" country="Cambodia" name="Mondolkiri" pageId="2" pageNumber="278">Mondulkiri Province</collectingRegion>
,
<quantity id="B10A9B71994383530BF0EAEFFB86FF69" box="[1017,1087,153,179]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="278" unit="km" value="16.0">16 km</quantity>
NNE of Sen Monorom, a small river, 12.584–587 N, 107.253–256 E,
<quantity id="B10A9B71994383530A49EACBFD06FF0D" box="[576,703,189,215]" metricMagnitude="2" metricUnit="m" metricValue="4.1" metricValueMax="4.2" metricValueMin="4.0" pageId="2" pageNumber="278" unit="m" value="410.0" valueMax="420.0" valueMin="400.0">400–420 m</quantity>
a.s.l.,
<date id="024C1054994383530B01EACBFC3DFF0D" box="[776,900,189,215]" pageId="2" pageNumber="278" value="2018-06-26">26 vi 2018</date>
.
<specimenCount id="60F4FD1D994383530B87EACBFC0CFF0D" box="[910,949,189,215]" pageId="2" pageNumber="278" type="male">1 ♂</specimenCount>
—the same place,
<date id="024C1054994383530C8EEACBFABAFF0D" box="[1159,1283,189,215]" pageId="2" pageNumber="278" value="2018-06-27">27 vi 2018</date>
.
</paragraph>
<paragraph id="764D36949943835308CEEA97FD98FE99" blockId="2.[151,1437,153,540]" pageId="2" pageNumber="278">
All specimens are collected by the author. The
<typeStatus id="A9498836994383530AE7EA97FCEAFF21" box="[750,851,225,251]" pageId="2" pageNumber="278" type="holotype">holotype</typeStatus>
, one male collected on
<date id="024C1054994383530C60EA97FB5EFF21" box="[1129,1255,225,251]" pageId="2" pageNumber="278" value="2018-06-26">26 vi 2018</date>
and the female collected on
<date id="024C1054994383530923EB73FE1DFEC5" box="[298,420,261,287]" pageId="2" pageNumber="278" value="2018-06-11">11 vi 2018</date>
are kept in Naturalis Biodiversity Centre, Leiden,
<collectingCountry id="0EE57604994383530BE0EB73FB19FEFA" box="[1001,1184,261,288]" name="Netherlands" pageId="2" pageNumber="278">the Netherlands</collectingCountry>
), the remaining specimens are in the author’s collection.
</paragraph>
<paragraph id="764D36949943835308CEEB3AFC30FDC1" blockId="2.[151,1437,153,540]" pageId="2" pageNumber="278">
<emphasis id="4486EA869943835308CEEB3AFE17FEBD" bold="true" box="[199,430,332,359]" pageId="2" pageNumber="278">Variation in males.</emphasis>
The
<typeStatus id="A94988369943835309E7EB3BFDE9FEBD" box="[494,592,333,359]" pageId="2" pageNumber="278" type="holotype">holotype</typeStatus>
and
<typeStatus id="A9498836994383530A85EB3BFD55FEBD" box="[652,748,333,359]" pageId="2" pageNumber="278" type="paratype">paratype</typeStatus>
<materialsCitation id="C69A3CC9994383530AFFEB3BFC3CFDC1" ID-GBIF-Occurrence="2460219569" collectingDate="2014-12-13" collectorName="FW Ax &amp; The &amp; Phnom Kulen &amp; Ab &amp; However" location="Ab" municipality="Cardamom Mts" pageId="2" pageNumber="278" specimenCount="1" stateProvince="Mondolkiri" typeStatus="holotype">
of
<taxonomicName id="B1F24D17994383530B11EB38FCC3FEBD" authorityName="Kosterin" authorityYear="2019" box="[792,890,333,359]" class="Insecta" family="Gomphidae" genus="Microgomphus" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="278" phylum="Arthropoda" rank="species" species="alani">
<emphasis id="4486EA86994383530B11EB38FCC3FEBD" box="[792,890,333,359]" italics="true" pageId="2" pageNumber="278">M. alani</emphasis>
</taxonomicName>
, originating from the
<collectingMunicipality id="9629ACEE994383530C75EB3BFA89FEBD" box="[1148,1328,333,359]" pageId="2" pageNumber="278">Cardamom Mts</collectingMunicipality>
, are very small: HW 23 and
<quantity id="B10A9B71994383530965EB06FE6AFE51" box="[364,467,368,395]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" pageId="2" pageNumber="278" unit="mm" value="21.5">21.5 mm</quantity>
,
<location id="732D604F9943835309D7EB07FDB9FE51" LSID="urn:lsid:plazi:treatment:FE5B87829940835B089EE9FBFAEEFC35:732D604F9943835309D7EB07FDB9FE51" box="[478,512,369,395]" municipality="Cardamom Mts" name="Ab" pageId="2" pageNumber="278" stateProvince="Mondolkiri">Ab</location>
25 and
<quantity id="B10A9B71994383530A52EB07FD15FE51" box="[603,684,369,395]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="278" unit="mm" value="23.0">23 mm</quantity>
,
<collectorName id="DB075342994383530AB0EB07FCB3FE51" box="[697,778,369,395]" pageId="2" pageNumber="278">FW Ax</collectorName>
<date id="024C1054994383530B1BEB07FC61FE51" box="[786,984,369,395]" pageId="2" pageNumber="278" value="2014-12-13" valueMax="2014-12-13" valueMin="2014-12-13">
<collectingDate id="1208E9BC994383530B1BEB07FC61FE51" box="[786,984,369,395]" pageId="2" pageNumber="278" value="2014-12-13" valueMax="2014-12-13" valueMin="2014-12-13">12–13 and 13–14</collectingDate>
</date>
, of which the second primary one is 5th and 4th/5
<superScript id="81879BDC9943835308F6EBE2FEB5FE78" attach="none" box="[255,268,404,418]" fontSize="6" pageId="2" pageNumber="278">th</superScript>
, respectively (
<bibRefCitation id="12634B659943835309A7EBE3FDD9FE6A" author="Kosterin, O. E." box="[430,608,405,432]" pageId="2" pageNumber="278" pagination="341 - 350" refId="ref9406" refString="Kosterin, O. E. (2016 a) Microgomphus alani (Odonata, Gomphidae) sp. nov. from Cambodia. Zootaxa, 4114 (3), 341 - 350. https: // doi. org / 10.11646 / zootaxa. 4114.3.9" type="journal article" year="2016">Kosterin 2016a</bibRefCitation>
).
<collectorName id="DB075342994383530A78EBE3FD24FE75" box="[625,669,405,431]" pageId="2" pageNumber="278">The</collectorName>
male from
<collectorName id="DB075342994383530B12EBE3FC0CFE75" box="[795,949,405,431]" pageId="2" pageNumber="278">Phnom Kulen</collectorName>
is only very slightly larger: HW
<quantity id="B10A9B71994383530D28EBE3FAC8FE75" box="[1313,1393,405,431]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="278" unit="mm" value="24.0">24 mm</quantity>
,
<collectorName id="DB075342994383530D73EBE3FA25FE75" box="[1402,1436,405,431]" pageId="2" pageNumber="278">Ab</collectorName>
<quantity id="B10A9B7199438353089EEBCFFF51FE09" box="[151,232,441,467]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="278" unit="mm" value="26.0">26 mm</quantity>
,
<collectorName id="DB0753429943835308FDEBCFFEFCFE09" box="[244,325,441,467]" pageId="2" pageNumber="278">FW Ax</collectorName>
15, of which the second primary is 5
<superScript id="81879BDC994383530AEEEBCEFD4DFE1C" attach="none" box="[743,756,440,454]" fontSize="6" pageId="2" pageNumber="278">th</superScript>
/6th.
<collectorName id="DB075342994383530B27EBCFFC2FFE09" box="[814,918,441,467]" pageId="2" pageNumber="278">However</collectorName>
, the five males from
<collectingRegion id="B436F876994383530C8DEBCFFABDFE09" box="[1156,1284,441,467]" country="Cambodia" name="Mondolkiri" pageId="2" pageNumber="278">Mondulkiri</collectingRegion>
are distinctly larger: HW 26–27 (mean 26.6) mm,
<collectorName id="DB075342994383530A3BEBABFDEDFE2D" box="[562,596,477,503]" pageId="2" pageNumber="278">Ab</collectorName>
29.0–30.5 (mean 29.8) mm,
<collectorName id="DB075342994383530B95EBABFC57FE2D" box="[924,1006,477,503]" pageId="2" pageNumber="278">FW Ax</collectorName>
13–15 (of which 5
<superScript id="81879BDC994383530CCCEBAAFB6BFE30" attach="left" box="[1221,1234,476,490]" fontSize="6" pageId="2" pageNumber="278">th</superScript>
(7 cases) or 6
<superScript id="81879BDC994383530D78EBAAFAC7FE30" attach="left" box="[1393,1406,476,490]" fontSize="6" pageId="2" pageNumber="278">th</superScript>
(3 cases) are primary); they are thus 15–20% larger than the
<typeStatus id="A9498836994383530B16E877FC3CFDC1" box="[799,901,513,539]" pageId="2" pageNumber="278" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
<caption id="228D661C99438353089EEFDCFE7CF9D1" ID-DOI="http://doi.org/10.5281/zenodo.3558041" ID-Zenodo-Dep="3558041" httpUri="https://zenodo.org/record/3558041/files/figure.png" pageId="2" pageNumber="278" startId="2.[151,250,1450,1475]" targetBox="[151,1436,553,1431]" targetPageId="2">
<paragraph id="764D369499438353089EEFDCFE7CF9D1" blockId="2.[151,1437,1450,1547]" pageId="2" pageNumber="278">
<emphasis id="4486EA8699438353089EEFDCFEADFA19" bold="true" box="[151,276,1450,1475]" pageId="2" pageNumber="278">FIGURE 1.</emphasis>
Male (a, d
<emphasis id="4486EA8699438353098CEFDAFE2BFA19" box="[389,402,1452,1475]" italics="true" pageId="2" pageNumber="278">–</emphasis>
e) and female (b
<emphasis id="4486EA86994383530A33EFDAFDFEFA19" box="[570,583,1452,1475]" italics="true" pageId="2" pageNumber="278">–</emphasis>
c) specimens of
<taxonomicName id="B1F24D17994383530AE6EFDDFC7BFA19" authorityName="Kosterin" authorityYear="2019" box="[751,962,1451,1475]" class="Insecta" family="Gomphidae" genus="Microgomphus" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="278" phylum="Arthropoda" rank="species" species="alani">
<emphasis id="4486EA86994383530AE6EFDDFC7BFA19" box="[751,962,1451,1475]" italics="true" pageId="2" pageNumber="278">Microgomphus alani</emphasis>
</taxonomicName>
in live colours from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d
<emphasis id="4486EA86994383530AF3EFA6FCBEFA3D" box="[762,775,1488,1511]" italics="true" pageId="2" pageNumber="278">–</emphasis>
e) of Cambodia, 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d
<emphasis id="4486EA86994383530916EF82FE95F9D1" box="[287,300,1524,1547]" italics="true" pageId="2" pageNumber="278">–</emphasis>
e). Not to scale
</paragraph>
</caption>
<paragraph id="764D36949943835308CEEC42FEE1F86E" blockId="2.[151,1437,1588,2044]" pageId="2" pageNumber="278">
A degree of inflation of the pale pattern correlates with size. The
<typeStatus id="A9498836994383530BA9EC43FBBAF995" box="[928,1027,1589,1615]" pageId="2" pageNumber="278" type="paratype">paratype</typeStatus>
, which is the smallest specimen, has no antealar spots on the mesepisterna (
<bibRefCitation id="12634B65994383530A4AEC2EFD4CF9A8" author="Kosterin, O. E." box="[579,757,1624,1650]" pageId="2" pageNumber="278" pagination="341 - 350" refId="ref9406" refString="Kosterin, O. E. (2016 a) Microgomphus alani (Odonata, Gomphidae) sp. nov. from Cambodia. Zootaxa, 4114 (3), 341 - 350. https: // doi. org / 10.11646 / zootaxa. 4114.3.9" type="journal article" year="2016">Kosterin 2016a</bibRefCitation>
: fig.1b). The slightly larger
<typeStatus id="A9498836994383530C23EC2EFB35F9A8" box="[1066,1164,1624,1650]" pageId="2" pageNumber="278" type="holotype">holotype</typeStatus>
has their traces (
<bibRefCitation id="12634B65994383530D4EEC2EFF47F94F" author="Kosterin, O. E." pageId="2" pageNumber="278" pagination="341 - 350" refId="ref9406" refString="Kosterin, O. E. (2016 a) Microgomphus alani (Odonata, Gomphidae) sp. nov. from Cambodia. Zootaxa, 4114 (3), 341 - 350. https: // doi. org / 10.11646 / zootaxa. 4114.3.9" type="journal article" year="2016">Kosterin 2016a</bibRefCitation>
: fig1a, 2a–b), the further, slightly larger male from Phnom Kulen, has well developed but small antealar spots (
<figureCitation id="EEC92A119943835308E8ECE9FE8FF963" box="[225,310,1695,1721]" captionStart="FIGURE 1" captionStartId="2.[151,250,1450,1475]" captionTargetBox="[151,1436,553,1431]" captionTargetId="figure@2.[151,1436,553,1431]" captionTargetPageId="2" captionText="FIGURE 1. Male (a, d–e) and female (b–c) specimens of Microgomphus alani in live colours from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–e) of Cambodia, 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e). Not to scale" figureDoi="http://doi.org/10.5281/zenodo.3558041" httpUri="https://zenodo.org/record/3558041/files/figure.png" pageId="2" pageNumber="278">Figs 1a</figureCitation>
,
<figureCitation id="EEC92A1199438353094CECE9FEDBF963" box="[325,354,1695,1721]" captionStart="FIGURE 2" captionStartId="3.[151,250,1924,1949]" captionTargetBox="[151,1431,805,1905]" captionTargetId="figure@3.[151,1436,805,1905]" captionTargetPageId="3" captionText="FIGURE 2. Dry male (a, d–e) and female (b–c, f) specimens of Microgomphus alani from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–f) of Cambodia, collected on 3 viii 2016 (f), 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e)." figureDoi="http://doi.org/10.5281/zenodo.3558043" httpUri="https://zenodo.org/record/3558043/files/figure.png" pageId="2" pageNumber="278">2a</figureCitation>
,
<figureCitation id="EEC92A11994383530967ECE9FE25F960" box="[366,412,1695,1722]" captionStart="FIGURE 3" captionStartId="4.[151,250,1902,1927]" captionTargetBox="[266,1321,166,1874]" captionTargetId="figure@4.[266,1321,166,1883]" captionTargetPageId="4" captionText="FIGURE 3. Details of male specimens of Microgomphus alani collected on 26 vi 2018 in Mondulkiri Province (a–d; the same specimen as in Fig. 2d) and on 17 vi 2018 in the Phnom Kulen Mts (a–h, the same specimen as in Fig. 1a and Fig 2a): a,e—head and thorax, lateral view; b,f—pterothorax, dorsal view; c,g—head, frontal (c) and dorsofrontal (g) view; d,h—secondary genitalia, lateral view. Scale bars. from top to bottom, 4 mm, 2 mm, 2 mm and 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3558045" httpUri="https://zenodo.org/record/3558045/files/figure.png" pageId="2" pageNumber="278">3e,f</figureCitation>
), while these spots are relatively large in the males from
<collectingRegion id="B436F876994383530C20ECE9FAACF963" box="[1065,1301,1695,1721]" country="Cambodia" name="Mondolkiri" pageId="2" pageNumber="278">Mondulkiri Province</collectingRegion>
(
<figureCitation id="EEC92A11994383530D2CECE9FAD0F963" box="[1317,1385,1695,1721]" captionStart="FIGURE 1" captionStartId="2.[151,250,1450,1475]" captionTargetBox="[151,1436,553,1431]" captionTargetId="figure@2.[151,1436,553,1431]" captionTargetPageId="2" captionText="FIGURE 1. Male (a, d–e) and female (b–c) specimens of Microgomphus alani in live colours from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–e) of Cambodia, 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e). Not to scale" figureDoi="http://doi.org/10.5281/zenodo.3558041" httpUri="https://zenodo.org/record/3558041/files/figure.png" pageId="2" pageNumber="278">Figs 1</figureCitation>
d–e, 2d–e, 3a–b). In the type series males from the Cardamoms and the Phnom Kulen male all anteriolateral pale spots on S3–S6 are isolated from each other and dorsal pale streaks (
<figureCitation id="EEC92A11994383530B5AEC91FC1BF8DB" box="[851,930,1767,1793]" captionStart="FIGURE 2" captionStartId="3.[151,250,1924,1949]" captionTargetBox="[151,1431,805,1905]" captionTargetId="figure@3.[151,1436,805,1905]" captionTargetPageId="3" captionText="FIGURE 2. Dry male (a, d–e) and female (b–c, f) specimens of Microgomphus alani from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–f) of Cambodia, collected on 3 viii 2016 (f), 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e)." figureDoi="http://doi.org/10.5281/zenodo.3558043" httpUri="https://zenodo.org/record/3558043/files/figure.png" pageId="2" pageNumber="278">Fig. 2a</figureCitation>
) but in the
<collectingRegion id="B436F876994383530C28EC91FB18F8DB" box="[1057,1185,1767,1793]" country="Cambodia" name="Mondolkiri" pageId="2" pageNumber="278">Mondulkiri</collectingRegion>
males those on S5–S6 (in one male only on S6) are fused to the anterior element of the dorsal streaks to form tripartite anterior spots (
<figureCitation id="EEC92A11994383530D66ED7CFF0CF892" captionStart="FIGURE 2" captionStartId="3.[151,250,1924,1949]" captionTargetBox="[151,1431,805,1905]" captionTargetId="figure@3.[151,1436,805,1905]" captionTargetPageId="3" captionText="FIGURE 2. Dry male (a, d–e) and female (b–c, f) specimens of Microgomphus alani from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–f) of Cambodia, collected on 3 viii 2016 (f), 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e)." figureDoi="http://doi.org/10.5281/zenodo.3558043" httpUri="https://zenodo.org/record/3558043/files/figure.png" pageId="2" pageNumber="278">Fig. 2e</figureCitation>
). The two males from the Cardamoms (the type series) (
<bibRefCitation id="12634B65994383530B31ED58FC52F892" author="Kosterin, O. E." box="[824,1003,1838,1864]" pageId="2" pageNumber="278" pagination="341 - 350" refId="ref9406" refString="Kosterin, O. E. (2016 a) Microgomphus alani (Odonata, Gomphidae) sp. nov. from Cambodia. Zootaxa, 4114 (3), 341 - 350. https: // doi. org / 10.11646 / zootaxa. 4114.3.9" type="journal article" year="2016">Kosterin 2016a</bibRefCitation>
: fig. 1a), the male from Phnom Kulen (
<figureCitation id="EEC92A119943835308A9ED24FF4BF8B6" box="[160,242,1874,1900]" captionStart="FIGURE 4" captionStartId="5.[151,250,1906,1931]" captionTargetBox="[151,1424,792,1868]" captionTargetId="figure@5.[151,1436,769,1886]" captionTargetPageId="5" captionText="FIGURE 4. Anal appendages, in dorsal (left) and lateral (right) view, of mature males of Microgomphus alani, collected on 26 vi 2018 in Mondulkiri Province (a–b, respectively corresponding to the specimens in Fig. 2d and Fig. 2e) and on 17 vi 2018 in the Phnom Kulen Mts (c, correspond to Fig. 1a and Fig 2a). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3558047" httpUri="https://zenodo.org/record/3558047/files/figure.png" pageId="2" pageNumber="278">Fig. 4c</figureCitation>
) and two males from
<collectingRegion id="B436F8769943835309F9ED24FDC9F8B6" box="[496,624,1874,1900]" country="Cambodia" name="Mondolkiri" pageId="2" pageNumber="278">Mondulkiri</collectingRegion>
have no pale spots on S8 while three males from
<collectingRegion id="B436F876994383530CB9ED24FA24F8B6" box="[1200,1437,1874,1900]" country="Cambodia" name="Mondolkiri" pageId="2" pageNumber="278">Mondulkiri Province</collectingRegion>
have a very small, elongate and slanting, lateral spot at the S8 posterior margin, at the middle of its height in lateral view (
<figureCitation id="EEC92A119943835308D4EDEFFE99F86E" box="[221,288,1945,1972]" captionStart="FIGURE 4" captionStartId="5.[151,250,1906,1931]" captionTargetBox="[151,1424,792,1868]" captionTargetId="figure@5.[151,1436,769,1886]" captionTargetPageId="5" captionText="FIGURE 4. Anal appendages, in dorsal (left) and lateral (right) view, of mature males of Microgomphus alani, collected on 26 vi 2018 in Mondulkiri Province (a–b, respectively corresponding to the specimens in Fig. 2d and Fig. 2e) and on 17 vi 2018 in the Phnom Kulen Mts (c, correspond to Fig. 1a and Fig 2a). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3558047" httpUri="https://zenodo.org/record/3558047/files/figure.png" pageId="2" pageNumber="278">Fig. 4</figureCitation>
a–b).
</paragraph>
<paragraph id="764D36949943835208CEEDCBFAD6FF02" blockId="2.[151,1437,1588,2044]" lastBlockId="3.[151,1437,153,792]" lastPageId="3" lastPageNumber="279" pageId="2" pageNumber="278">
Pterostigmata are dark brownish in all mature males, while pale brownish in the teneral males of the
<typeStatus id="A9498836994383530D16EDC8FAF6F802" box="[1311,1359,1982,2008]" pageId="2" pageNumber="278">type</typeStatus>
series. Unlike these teneral males, the lower part of the pterothorax, the coxae, trochanters, femora and secondary genitalia in all mature males were covered with white pruinescence (
<figureCitation id="EEC92A11994283520B29EAEFFC3BFF6E" box="[800,898,153,180]" captionStart="FIGURE 1" captionStartId="2.[151,250,1450,1475]" captionTargetBox="[151,1436,553,1431]" captionTargetId="figure@2.[151,1436,553,1431]" captionTargetPageId="2" captionText="FIGURE 1. Male (a, d–e) and female (b–c) specimens of Microgomphus alani in live colours from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–e) of Cambodia, 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e). Not to scale" figureDoi="http://doi.org/10.5281/zenodo.3558041" httpUri="https://zenodo.org/record/3558041/files/figure.png" pageId="3" pageNumber="279">Fig. 1a,d</figureCitation>
). This was later washed out by acetone treatment (
<figureCitation id="EEC92A11994283520896EACBFF48FF0D" box="[159,241,189,215]" captionStart="FIGURE 2" captionStartId="3.[151,250,1924,1949]" captionTargetBox="[151,1431,805,1905]" captionTargetId="figure@3.[151,1436,805,1905]" captionTargetPageId="3" captionText="FIGURE 2. Dry male (a, d–e) and female (b–c, f) specimens of Microgomphus alani from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–f) of Cambodia, collected on 3 viii 2016 (f), 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e)." figureDoi="http://doi.org/10.5281/zenodo.3558043" httpUri="https://zenodo.org/record/3558043/files/figure.png" pageId="3" pageNumber="279">Figs 2a</figureCitation>
,d–e, 3a,e) but is well preserved in a male of
<date id="024C1054994283520AD3EACBFCE8FF0D" box="[730,849,189,215]" pageId="3" pageNumber="279" value="2014-06-13">13 vi 2014</date>
, which was not treated with acetone (not shown).
</paragraph>
<paragraph id="764D36949942835208CEEA97FC08FD5D" blockId="3.[151,1437,153,792]" pageId="3" pageNumber="279">
The anal appendages are deformed in the
<typeStatus id="A9498836994283520AAAEA97FCBAFF21" box="[675,771,225,251]" pageId="3" pageNumber="279" type="paratype">paratype</typeStatus>
and, as will be discussed below, not fully expanded in the
<typeStatus id="A949883699428352089EEB73FF40FEC5" box="[151,249,261,287]" pageId="3" pageNumber="279" type="holotype">holotype</typeStatus>
(although the original description was based on them, see
<bibRefCitation id="12634B65994283520B97EB73FBE8FEFA" author="Kosterin, O. E." box="[926,1105,261,288]" pageId="3" pageNumber="279" pagination="341 - 350" refId="ref9406" refString="Kosterin, O. E. (2016 a) Microgomphus alani (Odonata, Gomphidae) sp. nov. from Cambodia. Zootaxa, 4114 (3), 341 - 350. https: // doi. org / 10.11646 / zootaxa. 4114.3.9" type="journal article" year="2016">Kosterin 2016a</bibRefCitation>
); hence the type series is not fully comparable with the presently available and here considered mature specimens. In these specimens, the cerci look identical in dorsal view (
<figureCitation id="EEC92A119942835209E2EB3BFD96FEBD" box="[491,559,333,359]" captionStart="FIGURE 4" captionStartId="5.[151,250,1906,1931]" captionTargetBox="[151,1424,792,1868]" captionTargetId="figure@5.[151,1436,769,1886]" captionTargetPageId="5" captionText="FIGURE 4. Anal appendages, in dorsal (left) and lateral (right) view, of mature males of Microgomphus alani, collected on 26 vi 2018 in Mondulkiri Province (a–b, respectively corresponding to the specimens in Fig. 2d and Fig. 2e) and on 17 vi 2018 in the Phnom Kulen Mts (c, correspond to Fig. 1a and Fig 2a). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3558047" httpUri="https://zenodo.org/record/3558047/files/figure.png" pageId="3" pageNumber="279">Fig. 4</figureCitation>
a–c) and differ from those in the
<typeStatus id="A9498836994283520BAEEB3BFBB0FEBD" box="[935,1033,333,359]" pageId="3" pageNumber="279" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="12634B65994283520C10EB3BFB74FEBD" author="Kosterin, O. E." box="[1049,1229,333,359]" pageId="3" pageNumber="279" pagination="341 - 350" refId="ref9406" refString="Kosterin, O. E. (2016 a) Microgomphus alani (Odonata, Gomphidae) sp. nov. from Cambodia. Zootaxa, 4114 (3), 341 - 350. https: // doi. org / 10.11646 / zootaxa. 4114.3.9" type="journal article" year="2016">Kosterin 2016a</bibRefCitation>
: fig. 2c) in having a divergent rather than parallel alignment, and having their inner branches slightly curved and protruding further behind than the main parts (‘outer branches’) of cerci, rather than strongly curved and not reaching the level of the main parts of cerci. The cercus main part has the same principal shape as in the
<typeStatus id="A9498836994283520C11EBCFFBC4FE09" box="[1048,1149,441,467]" pageId="3" pageNumber="279" type="holotype">holotype</typeStatus>
, with a bluntly bifid apex and a blunt outer tooth, but is longer. Interestingly, the epiproct in the
<typeStatus id="A9498836994283520BBBEBABFBADFE2D" box="[946,1044,477,503]" pageId="3" pageNumber="279" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="12634B65994283520C2DEBABFB6EFE2D" author="Kosterin, O. E." box="[1060,1239,477,503]" pageId="3" pageNumber="279" pagination="341 - 350" refId="ref9406" refString="Kosterin, O. E. (2016 a) Microgomphus alani (Odonata, Gomphidae) sp. nov. from Cambodia. Zootaxa, 4114 (3), 341 - 350. https: // doi. org / 10.11646 / zootaxa. 4114.3.9" type="journal article" year="2016">Kosterin 2016a</bibRefCitation>
: fig. c–d) and the male from Phnom Kulen (
<figureCitation id="EEC92A119942835209B0E877FDB1FDC1" box="[441,520,513,539]" captionStart="FIGURE 4" captionStartId="5.[151,250,1906,1931]" captionTargetBox="[151,1424,792,1868]" captionTargetId="figure@5.[151,1436,769,1886]" captionTargetPageId="5" captionText="FIGURE 4. Anal appendages, in dorsal (left) and lateral (right) view, of mature males of Microgomphus alani, collected on 26 vi 2018 in Mondulkiri Province (a–b, respectively corresponding to the specimens in Fig. 2d and Fig. 2e) and on 17 vi 2018 in the Phnom Kulen Mts (c, correspond to Fig. 1a and Fig 2a). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3558047" httpUri="https://zenodo.org/record/3558047/files/figure.png" pageId="3" pageNumber="279">Fig. 4c</figureCitation>
) is similar in both dorsal and lateral view, while in the
<collectingRegion id="B436F876994283520C6EE877FB5EFDC1" box="[1127,1255,513,539]" country="Cambodia" name="Mondolkiri" pageId="3" pageNumber="279">Mondulkiri</collectingRegion>
males it is more strongly curved up (
<figureCitation id="EEC92A1199428352097CE853FE01FD9A" box="[373,440,549,576]" captionStart="FIGURE 4" captionStartId="5.[151,250,1906,1931]" captionTargetBox="[151,1424,792,1868]" captionTargetId="figure@5.[151,1436,769,1886]" captionTargetPageId="5" captionText="FIGURE 4. Anal appendages, in dorsal (left) and lateral (right) view, of mature males of Microgomphus alani, collected on 26 vi 2018 in Mondulkiri Province (a–b, respectively corresponding to the specimens in Fig. 2d and Fig. 2e) and on 17 vi 2018 in the Phnom Kulen Mts (c, correspond to Fig. 1a and Fig 2a). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3558047" httpUri="https://zenodo.org/record/3558047/files/figure.png" pageId="3" pageNumber="279">Fig. 4</figureCitation>
a–b). This difference is strong enough to suppose that these males are not conspecific with the males from the Cardamom and Phnom Kulen Mts., but the whole complex of evidence is not solid enough and made me to consider this as a case of geographical structural variation.
</paragraph>
<paragraph id="764D36949942835208CEE8E7FABDFCC2" blockId="3.[151,1437,153,792]" pageId="3" pageNumber="279">
The posterior hamulus in lateral view is slightly curved apically in the
<typeStatus id="A9498836994283520BE3E8E7FBF5FD71" box="[1002,1100,657,683]" pageId="3" pageNumber="279" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="12634B65994283520C52E8E7FAB7FD71" author="Kosterin, O. E." box="[1115,1294,657,683]" pageId="3" pageNumber="279" pagination="341 - 350" refId="ref9406" refString="Kosterin, O. E. (2016 a) Microgomphus alani (Odonata, Gomphidae) sp. nov. from Cambodia. Zootaxa, 4114 (3), 341 - 350. https: // doi. org / 10.11646 / zootaxa. 4114.3.9" type="journal article" year="2016">Kosterin 2016a</bibRefCitation>
: figs 2f, 3a), more strongly curved in the Phnom Kulen and
<collectingRegion id="B436F876994283520AAFE8C3FC9FFD15" box="[678,806,693,719]" country="Cambodia" name="Mondolkiri" pageId="3" pageNumber="279">Mondulkiri</collectingRegion>
males (
<figureCitation id="EEC92A11994283520B77E8C3FC5DFD0A" box="[894,996,693,720]" captionStart="FIGURE 3" captionStartId="4.[151,250,1902,1927]" captionTargetBox="[266,1321,166,1874]" captionTargetId="figure@4.[266,1321,166,1883]" captionTargetPageId="4" captionText="FIGURE 3. Details of male specimens of Microgomphus alani collected on 26 vi 2018 in Mondulkiri Province (a–d; the same specimen as in Fig. 2d) and on 17 vi 2018 in the Phnom Kulen Mts (a–h, the same specimen as in Fig. 1a and Fig 2a): a,e—head and thorax, lateral view; b,f—pterothorax, dorsal view; c,g—head, frontal (c) and dorsofrontal (g) view; d,h—secondary genitalia, lateral view. Scale bars. from top to bottom, 4 mm, 2 mm, 2 mm and 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3558045" httpUri="https://zenodo.org/record/3558045/files/figure.png" pageId="3" pageNumber="279">Fig. 3d,h</figureCitation>
) and even more strongly hooked in the
<typeStatus id="A949883699428352089EE8AFFF4EFD29" box="[151,247,729,755]" pageId="3" pageNumber="279" type="paratype">paratype</typeStatus>
(
<bibRefCitation id="12634B6599428352090CE8AFFE0FFD29" author="Kosterin, O. E." box="[261,438,729,755]" pageId="3" pageNumber="279" pagination="341 - 350" refId="ref9406" refString="Kosterin, O. E. (2016 a) Microgomphus alani (Odonata, Gomphidae) sp. nov. from Cambodia. Zootaxa, 4114 (3), 341 - 350. https: // doi. org / 10.11646 / zootaxa. 4114.3.9" type="journal article" year="2016">Kosterin 2016a</bibRefCitation>
: figs
<quantity id="B10A9B719942835209E6E8AFFDB4FD29" box="[495,525,729,755]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="3" pageNumber="279" unit="g" value="2.0">2g</quantity>
, 3c). The anterior hamulus is similarly simply curved in the type series and Phnom Kulen male (
<figureCitation id="EEC92A11994283520921E88BFEC3FCC2" box="[296,378,765,792]" captionStart="FIGURE 3" captionStartId="4.[151,250,1902,1927]" captionTargetBox="[266,1321,166,1874]" captionTargetId="figure@4.[266,1321,166,1883]" captionTargetPageId="4" captionText="FIGURE 3. Details of male specimens of Microgomphus alani collected on 26 vi 2018 in Mondulkiri Province (a–d; the same specimen as in Fig. 2d) and on 17 vi 2018 in the Phnom Kulen Mts (a–h, the same specimen as in Fig. 1a and Fig 2a): a,e—head and thorax, lateral view; b,f—pterothorax, dorsal view; c,g—head, frontal (c) and dorsofrontal (g) view; d,h—secondary genitalia, lateral view. Scale bars. from top to bottom, 4 mm, 2 mm, 2 mm and 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3558045" httpUri="https://zenodo.org/record/3558045/files/figure.png" pageId="3" pageNumber="279">Fig. 3h</figureCitation>
) but somewhat S-like curved and more robust in the
<collectingRegion id="B436F876994283520BC5E88BFBF5FCCD" box="[972,1100,765,791]" country="Cambodia" name="Mondolkiri" pageId="3" pageNumber="279">Mondulkiri</collectingRegion>
males (
<figureCitation id="EEC92A11994283520CADE88BFB4EFCC2" box="[1188,1271,765,792]" captionStart="FIGURE 3" captionStartId="4.[151,250,1902,1927]" captionTargetBox="[266,1321,166,1874]" captionTargetId="figure@4.[266,1321,166,1883]" captionTargetPageId="4" captionText="FIGURE 3. Details of male specimens of Microgomphus alani collected on 26 vi 2018 in Mondulkiri Province (a–d; the same specimen as in Fig. 2d) and on 17 vi 2018 in the Phnom Kulen Mts (a–h, the same specimen as in Fig. 1a and Fig 2a): a,e—head and thorax, lateral view; b,f—pterothorax, dorsal view; c,g—head, frontal (c) and dorsofrontal (g) view; d,h—secondary genitalia, lateral view. Scale bars. from top to bottom, 4 mm, 2 mm, 2 mm and 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3558045" httpUri="https://zenodo.org/record/3558045/files/figure.png" pageId="3" pageNumber="279">Fig. 3h</figureCitation>
).
</paragraph>
<caption id="228D661C99428352089EEDF2FE51F83F" ID-DOI="http://doi.org/10.5281/zenodo.3558043" ID-Zenodo-Dep="3558043" httpUri="https://zenodo.org/record/3558043/files/figure.png" pageId="3" pageNumber="279" startId="3.[151,250,1924,1949]" targetBox="[151,1431,805,1905]" targetPageId="3">
<paragraph id="764D369499428352089EEDF2FE51F83F" blockId="3.[151,1436,1924,2021]" pageId="3" pageNumber="279">
<emphasis id="4486EA8699428352089EEDF2FEAAF847" bold="true" box="[151,275,1924,1949]" pageId="3" pageNumber="279">FIGURE 2.</emphasis>
Dry male (a, d
<emphasis id="4486EA869942835209A5EDF3FE00F846" box="[428,441,1925,1948]" italics="true" pageId="3" pageNumber="279">–</emphasis>
e) and female (b
<emphasis id="4486EA86994283520A57EDF3FDD2F846" box="[606,619,1925,1948]" italics="true" pageId="3" pageNumber="279">–</emphasis>
c, f) specimens of
<taxonomicName id="B1F24D17994283520B2FEDF3FC40F847" authorityName="Kosterin" authorityYear="2019" box="[806,1017,1925,1949]" class="Insecta" family="Gomphidae" genus="Microgomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="279" phylum="Arthropoda" rank="species" species="alani">
<emphasis id="4486EA86994283520B2FEDF3FC40F847" box="[806,1017,1925,1949]" italics="true" pageId="3" pageNumber="279">Microgomphus alani</emphasis>
</taxonomicName>
from the Phnom Kulen Mountains, Siem Reap Province (a
<emphasis id="4486EA8699428352094CEDDFFEEBF81A" box="[325,338,1961,1984]" italics="true" pageId="3" pageNumber="279">–</emphasis>
c) and Mondulkiri Province (d
<emphasis id="4486EA86994283520A8DEDDFFD28F81A" box="[644,657,1961,1984]" italics="true" pageId="3" pageNumber="279">–</emphasis>
f) of Cambodia, collected on 3 viii 2016 (f), 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d
<emphasis id="4486EA869942835209C8EDBBFE77F83E" box="[449,462,1997,2020]" italics="true" pageId="3" pageNumber="279">–</emphasis>
e).
</paragraph>
</caption>
<caption id="228D661C99458355089EED18FC58F828" ID-DOI="http://doi.org/10.5281/zenodo.3558045" ID-Zenodo-Dep="3558045" httpUri="https://zenodo.org/record/3558045/files/figure.png" pageId="4" pageNumber="280" startId="4.[151,250,1902,1927]" targetBox="[266,1321,166,1874]" targetPageId="4">
<paragraph id="764D369499458355089EED18FC58F828" blockId="4.[151,1437,1902,2035]" pageId="4" pageNumber="280">
<emphasis id="4486EA8699458355089EED18FEAAF85D" bold="true" box="[151,275,1902,1927]" pageId="4" pageNumber="280">FIGURE 3.</emphasis>
Details of male specimens of
<taxonomicName id="B1F24D17994583550A4FED19FCA0F85D" authorityName="Kosterin" authorityYear="2019" box="[582,793,1903,1927]" class="Insecta" family="Gomphidae" genus="Microgomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="280" phylum="Arthropoda" rank="species" species="alani">
<emphasis id="4486EA86994583550A4FED19FCA0F85D" box="[582,793,1903,1927]" italics="true" pageId="4" pageNumber="280">Microgomphus alani</emphasis>
</taxonomicName>
collected on 26 vi 2018 in Mondulkiri Province (a
<emphasis id="4486EA86994583550D15ED06FA90F85D" box="[1308,1321,1904,1927]" italics="true" pageId="4" pageNumber="280">–</emphasis>
d; the same specimen as in Fig. 2d) and on 17 vi 2018 in the Phnom Kulen Mts (a
<emphasis id="4486EA86994583550B46EDE2FCE5F871" box="[847,860,1940,1963]" italics="true" pageId="4" pageNumber="280">–</emphasis>
h, the same specimen as in Fig. 1a and Fig 2a): a,e—head and thorax, lateral view; b,f—pterothorax, dorsal view; c,g—head, frontal (c) and dorsofrontal (g) view; d,h—secondary genitalia, lateral view. Scale bars. from top to bottom, 4 mm, 2 mm, 2 mm and 0.5 mm.
</paragraph>
</caption>
<paragraph id="764D36949944835408CEEAEFFDD7FE51" blockId="5.[151,1437,153,755]" pageId="5" pageNumber="281">
<emphasis id="4486EA869944835408CEEAEFFE9CFF69" bold="true" box="[199,293,153,179]" pageId="5" pageNumber="281">Female.</emphasis>
Three females are available for study. The female collected in
<quantity id="B10A9B71994483540BDEEAEFFB92FF69" box="[983,1067,153,179]" metricMagnitude="1" metricUnit="m" metricValue="5.12064" pageId="5" pageNumber="281" unit="in" value="2016.0">2016 in</quantity>
<collectingRegion id="B436F876994483540C39EAEFFAA3FF69" box="[1072,1306,153,179]" country="Cambodia" name="Mondolkiri" pageId="5" pageNumber="281">Mondulkiri Province</collectingRegion>
has already been briefly characterised, illustrated and compared to ‘
<taxonomicName id="B1F24D17994483540B1BEACBFB44FF0D" authority="' in Kosterin (2016 b" authorityName="' in Kosterin" authorityYear="2016" box="[786,1277,189,215]" class="Insecta" family="Gomphidae" genus="Heliogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="svihleri">
<emphasis id="4486EA86994483540B1BEACBFBABFF0D" box="[786,1042,189,215]" italics="true" pageId="5" pageNumber="281">Heliogomphus svihleri</emphasis>
’ in
<bibRefCitation id="12634B65994483540C49EACBFB44FF0D" author="Kosterin, O. E." box="[1088,1277,189,215]" pageId="5" pageNumber="281" pagination="1 - 86" refId="ref9453" refString="Kosterin, O. E. (2016 b) A survey of Odonata of Mondulkiri, the elevated eastern province of Cambodia, for ten days in June 2014. International Dragonfly Fund Report, 98, 1 - 86." type="journal article" year="2016">Kosterin (2016b</bibRefCitation>
</taxonomicName>
: fig. 56), who misidentified it as
<taxonomicName id="B1F24D17994483540961EA97FDB2FF21" authorityName="Laidlaw" authorityYear="1922" box="[360,523,225,251]" class="Insecta" family="Gomphidae" genus="Heliogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="281" phylum="Arthropoda" rank="genus">
<emphasis id="4486EA86994483540961EA97FDB2FF21" box="[360,523,225,251]" italics="true" pageId="5" pageNumber="281">Heliogomphus</emphasis>
</taxonomicName>
sp. Like males from the same province, it is considerably larger than the females from Phnom Kulen Mts. as well as the males of the type series from the Cardamoms. Since the male from Phnom Kulen more closely resembles the
<typeStatus id="A9498836994483540A17EB5FFD39FE99" box="[542,640,297,323]" pageId="5" pageNumber="281" type="holotype">holotype</typeStatus>
in size than those from
<collectingRegion id="B436F876994483540B9AEB5FFB39FE99" box="[915,1152,297,323]" country="Cambodia" name="Mondolkiri" pageId="5" pageNumber="281">Mondulkiri Province</collectingRegion>
, one of the females from Phnom Kulen, that of
<date id="024C105499448354099CEB3BFDA9FEBD" box="[405,528,333,359]" pageId="5" pageNumber="281" value="2016-06-30">30 vi 2016</date>
(
<figureCitation id="EEC92A11994483540A28EB3BFDC1FEBD" box="[545,632,333,359]" captionStart="FIGURE 1" captionStartId="2.[151,250,1450,1475]" captionTargetBox="[151,1436,553,1431]" captionTargetId="figure@2.[151,1436,553,1431]" captionTargetPageId="2" captionText="FIGURE 1. Male (a, d–e) and female (b–c) specimens of Microgomphus alani in live colours from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–e) of Cambodia, 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e). Not to scale" figureDoi="http://doi.org/10.5281/zenodo.3558041" httpUri="https://zenodo.org/record/3558041/files/figure.png" pageId="5" pageNumber="281">Figs 1b</figureCitation>
,
<figureCitation id="EEC92A11994483540A8EEB3BFD1CFEBD" box="[647,677,333,359]" captionStart="FIGURE 2" captionStartId="3.[151,250,1924,1949]" captionTargetBox="[151,1431,805,1905]" captionTargetId="figure@3.[151,1436,805,1905]" captionTargetPageId="3" captionText="FIGURE 2. Dry male (a, d–e) and female (b–c, f) specimens of Microgomphus alani from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–f) of Cambodia, collected on 3 viii 2016 (f), 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e)." figureDoi="http://doi.org/10.5281/zenodo.3558043" httpUri="https://zenodo.org/record/3558043/files/figure.png" pageId="5" pageNumber="281">2b</figureCitation>
,
<figureCitation id="EEC92A11994483540ABBEB3AFD76FEBD" box="[690,719,332,359]" captionStart="FIGURE 5" captionStartId="6.[151,250,1669,1694]" captionTargetBox="[170,1436,174,1602]" captionTargetId="figure@6.[151,1436,174,1650]" captionTargetPageId="6" captionText="FIGURE 5. Frontal view of head (left) and occiput (right) of female specimens of Microgomphus alani collected on 3 viii 2016 in Mondulkiri Province (a; the same specimen as in Fig. 2f) and in the Phnom Kulen Mts on 11 vi 2018 (b, the same specimen as in Figs 1c, 2c) and 30 vi 2018 (c, the same specimen as in Figs 1b, 2b). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3558049" httpUri="https://zenodo.org/record/3558049/files/figure.png" pageId="5" pageNumber="281">5c</figureCitation>
,
<figureCitation id="EEC92A11994483540AD5EB3BFC9DFEBD" box="[732,804,333,359]" captionStart="FIGURE 6" captionStartId="8.[151,250,1901,1926]" captionTargetBox="[161,1426,166,1866]" captionTargetId="figure@8.[161,1426,166,1881]" captionTargetPageId="8" captionText="FIGURE 6. Details of female specimens of Microgomphus alani collected in Mondulkiri Province on 3 viii 2016 (c,e; the same specimen as in Fig. 2f) and in the Phnom Kulen Mts on 11 vi 2018 (a,f; the same specimen as in Fig. 1c and Fig 2c) and on 30 vi 2018 (b,d,g; the same specimen as in Figs 1b, 2b): a–b—head and thorax, lateral view; c–d—pterothorax, dorsal view; e–g—vulvar lamina, ventral view. Scale bars, from top to bottom, 4 mm, 2 mm and 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3558051" httpUri="https://zenodo.org/record/3558051/files/figure.png" pageId="5" pageNumber="281">6b,d,g</figureCitation>
,
<figureCitation id="EEC92A11994483540B3BEB3BFCF6FEBD" box="[818,847,333,359]" captionStart="FIGURE 7" captionStartId="9.[151,250,1973,1998]" captionTargetBox="[345,1242,166,1896]" captionTargetId="figure@9.[345,1242,166,1953]" captionTargetPageId="9" captionText="FIGURE 7. Hind legs of Microgomphus alani, a—male from Mondulkiri Province, 13 vi 2014; b—female from Modulkiri Province, 3 viii 2016; c—female from Phnom Kulen Mts, 30 vi 2018. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3558053" httpUri="https://zenodo.org/record/3558053/files/figure.png" pageId="5" pageNumber="281">7c</figureCitation>
), was chosen for the description provided below, in spite of its aberrant asymmetrical occiput.
</paragraph>
<paragraph id="764D36949944835408CEEBE3FB19FD29" blockId="5.[151,1437,153,755]" pageId="5" pageNumber="281">
<emphasis id="4486EA869944835408CEEBE3FEBDFE75" box="[199,260,405,431]" italics="true" pageId="5" pageNumber="281">Head</emphasis>
(
<figureCitation id="EEC92A1199448354091DEBE3FED2FE75" box="[276,363,404,431]" captionStart="FIGURE 5" captionStartId="6.[151,250,1669,1694]" captionTargetBox="[170,1436,174,1602]" captionTargetId="figure@6.[151,1436,174,1650]" captionTargetPageId="6" captionText="FIGURE 5. Frontal view of head (left) and occiput (right) of female specimens of Microgomphus alani collected on 3 viii 2016 in Mondulkiri Province (a; the same specimen as in Fig. 2f) and in the Phnom Kulen Mts on 11 vi 2018 (b, the same specimen as in Figs 1c, 2c) and 30 vi 2018 (c, the same specimen as in Figs 1b, 2b). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3558049" httpUri="https://zenodo.org/record/3558049/files/figure.png" pageId="5" pageNumber="281">Figs 5c</figureCitation>
). Face black with pale yellow pattern. Labium pale but apical border of mentum and movable hooks black. Mandible bases pale. Labrum with a pair of large ovoid (tapering to external ends) yellow spots on either side. Anteclypeus with a broad rectangular central pale spot adjacent to its upper margin but leaving a black border along lower margin. Lower lateral parts of postclypeus broadly pale (these pale spots crossed by an additional seam separating lateroventral lobes of postclypeus). Frons dorsal surface with a broad pale stripe, broadest at middle. Vertex and occiput black. Vertex with a pair of bluntly conical tubercles behind ocelli, set twice as close to each other as to lateral ocelli and with a U-shaped depression between them. Occiput with a high blunt central hump (raising as high as eye tops), on the right side of which there is a narrow, pointed horn, the corresponding place to the left smooth, without a horn; occipital crest gently rises laterally towards eyes. Vertex and occiput with long sparse hairs; shorter hairs on sides of anteriolateral lobes of postclypeus, on labrum and mandibles.
</paragraph>
<caption id="228D661C99448354089EED04FC19F808" ID-DOI="http://doi.org/10.5281/zenodo.3558047" ID-Zenodo-Dep="3558047" httpUri="https://zenodo.org/record/3558047/files/figure.png" pageId="5" pageNumber="281" startId="5.[151,250,1906,1931]" targetBox="[151,1424,792,1868]" targetPageId="5">
<paragraph id="764D369499448354089EED04FC19F808" blockId="5.[151,1437,1906,2003]" pageId="5" pageNumber="281">
<emphasis id="4486EA8699448354089EED04FEAAF851" bold="true" box="[151,275,1906,1931]" pageId="5" pageNumber="281">FIGURE 4.</emphasis>
Anal appendages, in dorsal (left) and lateral (right) view, of mature males of
<taxonomicName id="B1F24D17994483540C2BED05FB4CF851" authorityName="Kosterin" authorityYear="2019" box="[1058,1269,1907,1931]" class="Insecta" family="Gomphidae" genus="Microgomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="281" phylum="Arthropoda" rank="species" species="alani">
<emphasis id="4486EA86994483540C2BED05FB4CF851" box="[1058,1269,1907,1931]" italics="true" pageId="5" pageNumber="281">Microgomphus alani</emphasis>
</taxonomicName>
, collected on 26 vi 2018 in Mondulkiri Province (a
<emphasis id="4486EA869944835409FAEDE1FDB9F874" box="[499,512,1943,1966]" italics="true" pageId="5" pageNumber="281">–</emphasis>
b, respectively corresponding to the specimens in Fig. 2d and Fig. 2e) and on 17 vi 2018 in the Phnom Kulen Mts (c, correspond to Fig. 1a and Fig 2a). Scale bar 1 mm.
</paragraph>
</caption>
<caption id="228D661C99478357089EECF3FB96F93F" ID-DOI="http://doi.org/10.5281/zenodo.3558049" ID-Zenodo-Dep="3558049" httpUri="https://zenodo.org/record/3558049/files/figure.png" pageId="6" pageNumber="282" startId="6.[151,250,1669,1694]" targetBox="[170,1436,174,1602]" targetPageId="6">
<paragraph id="764D369499478357089EECF3FB96F93F" blockId="6.[151,1437,1669,1766]" pageId="6" pageNumber="282">
<emphasis id="4486EA8699478357089EECF3FEABF944" bold="true" box="[151,274,1669,1694]" pageId="6" pageNumber="282">FIGURE 5.</emphasis>
Frontal view of head (left) and occiput (right) of female specimens of
<taxonomicName id="B1F24D17994783570BDEECF0FB13F944" authorityName="Kosterin" authorityYear="2019" box="[983,1194,1670,1694]" class="Insecta" family="Gomphidae" genus="Microgomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="282" phylum="Arthropoda" rank="species" species="alani">
<emphasis id="4486EA86994783570BDEECF0FB13F944" box="[983,1194,1670,1694]" italics="true" pageId="6" pageNumber="282">Microgomphus alani</emphasis>
</taxonomicName>
collected on 3 viii 2016 in Mondulkiri Province (a; the same specimen as in Fig. 2f) and in the Phnom Kulen Mts on 11 vi 2018 (b, the same specimen as in Figs 1c, 2c) and 30 vi 2018 (c, the same specimen as in Figs 1b, 2b). Scale bar 1 mm.
</paragraph>
</caption>
<paragraph id="764D36949947835608CEED62FCD3FD71" blockId="6.[151,1437,1812,2019]" lastBlockId="7.[151,1437,153,1979]" lastPageId="7" lastPageNumber="283" pageId="6" pageNumber="282">
<emphasis id="4486EA869947835708CEED62FEA1F8F4" box="[199,280,1812,1838]" italics="true" pageId="6" pageNumber="282">Thorax</emphasis>
in life with a strong white pruinescence on coxae and lower parts of mesinfraepisternum and metinfraepisternum (
<figureCitation id="EEC92A1199478357095EED4EFE11F889" box="[343,424,1848,1875]" captionStart="FIGURE 1" captionStartId="2.[151,250,1450,1475]" captionTargetBox="[151,1436,553,1431]" captionTargetId="figure@2.[151,1436,553,1431]" captionTargetPageId="2" captionText="FIGURE 1. Male (a, d–e) and female (b–c) specimens of Microgomphus alani in live colours from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–e) of Cambodia, 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e). Not to scale" figureDoi="http://doi.org/10.5281/zenodo.3558041" httpUri="https://zenodo.org/record/3558041/files/figure.png" pageId="6" pageNumber="282">Fig. 1b</figureCitation>
), completely washed out
<emphasis id="4486EA86994783570ACFED4FFCEBF888" box="[710,850,1849,1874]" italics="true" pageId="6" pageNumber="282">post mortem</emphasis>
by acetone treatment (
<figureCitation id="EEC92A11994783570C46ED4EFB1FF888" box="[1103,1190,1848,1874]" captionStart="FIGURE 2" captionStartId="3.[151,250,1924,1949]" captionTargetBox="[151,1431,805,1905]" captionTargetId="figure@3.[151,1436,805,1905]" captionTargetPageId="3" captionText="FIGURE 2. Dry male (a, d–e) and female (b–c, f) specimens of Microgomphus alani from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–f) of Cambodia, collected on 3 viii 2016 (f), 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e)." figureDoi="http://doi.org/10.5281/zenodo.3558043" httpUri="https://zenodo.org/record/3558043/files/figure.png" pageId="6" pageNumber="282">Figs 2b</figureCitation>
,
<figureCitation id="EEC92A11994783570CB8ED4EFB69F889" box="[1201,1232,1848,1875]" captionStart="FIGURE 6" captionStartId="8.[151,250,1901,1926]" captionTargetBox="[161,1426,166,1866]" captionTargetId="figure@8.[161,1426,166,1881]" captionTargetPageId="8" captionText="FIGURE 6. Details of female specimens of Microgomphus alani collected in Mondulkiri Province on 3 viii 2016 (c,e; the same specimen as in Fig. 2f) and in the Phnom Kulen Mts on 11 vi 2018 (a,f; the same specimen as in Fig. 1c and Fig 2c) and on 30 vi 2018 (b,d,g; the same specimen as in Figs 1b, 2b): a–b—head and thorax, lateral view; c–d—pterothorax, dorsal view; e–g—vulvar lamina, ventral view. Scale bars, from top to bottom, 4 mm, 2 mm and 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3558051" httpUri="https://zenodo.org/record/3558051/files/figure.png" pageId="6" pageNumber="282">6b</figureCitation>
). Prothorax black, anterior lobe and large spots at sides of median lobe yellow. Pterothorax (
<figureCitation id="EEC92A11994783570BC4ED2AFB9AF8AC" box="[973,1059,1884,1910]" captionStart="FIGURE 1" captionStartId="2.[151,250,1450,1475]" captionTargetBox="[151,1436,553,1431]" captionTargetId="figure@2.[151,1436,553,1431]" captionTargetPageId="2" captionText="FIGURE 1. Male (a, d–e) and female (b–c) specimens of Microgomphus alani in live colours from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–e) of Cambodia, 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e). Not to scale" figureDoi="http://doi.org/10.5281/zenodo.3558041" httpUri="https://zenodo.org/record/3558041/files/figure.png" pageId="6" pageNumber="282">Figs 1b</figureCitation>
,
<figureCitation id="EEC92A11994783570C39ED2AFBF6F8AC" box="[1072,1103,1884,1910]" captionStart="FIGURE 2" captionStartId="3.[151,250,1924,1949]" captionTargetBox="[151,1431,805,1905]" captionTargetId="figure@3.[151,1436,805,1905]" captionTargetPageId="3" captionText="FIGURE 2. Dry male (a, d–e) and female (b–c, f) specimens of Microgomphus alani from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–f) of Cambodia, collected on 3 viii 2016 (f), 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e)." figureDoi="http://doi.org/10.5281/zenodo.3558043" httpUri="https://zenodo.org/record/3558043/files/figure.png" pageId="6" pageNumber="282">2b</figureCitation>
,
<figureCitation id="EEC92A11994783570C53ED2AFB2AF8AD" box="[1114,1171,1884,1911]" captionStart="FIGURE 6" captionStartId="8.[151,250,1901,1926]" captionTargetBox="[161,1426,166,1866]" captionTargetId="figure@8.[161,1426,166,1881]" captionTargetPageId="8" captionText="FIGURE 6. Details of female specimens of Microgomphus alani collected in Mondulkiri Province on 3 viii 2016 (c,e; the same specimen as in Fig. 2f) and in the Phnom Kulen Mts on 11 vi 2018 (a,f; the same specimen as in Fig. 1c and Fig 2c) and on 30 vi 2018 (b,d,g; the same specimen as in Figs 1b, 2b): a–b—head and thorax, lateral view; c–d—pterothorax, dorsal view; e–g—vulvar lamina, ventral view. Scale bars, from top to bottom, 4 mm, 2 mm and 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3558051" httpUri="https://zenodo.org/record/3558051/files/figure.png" pageId="6" pageNumber="282">6b, d</figureCitation>
) black and pale yellow. Its dorsum (
<figureCitation id="EEC92A11994783570915EDF6FED5F841" box="[284,364,1920,1947]" captionStart="FIGURE 6" captionStartId="8.[151,250,1901,1926]" captionTargetBox="[161,1426,166,1866]" captionTargetId="figure@8.[161,1426,166,1881]" captionTargetPageId="8" captionText="FIGURE 6. Details of female specimens of Microgomphus alani collected in Mondulkiri Province on 3 viii 2016 (c,e; the same specimen as in Fig. 2f) and in the Phnom Kulen Mts on 11 vi 2018 (a,f; the same specimen as in Fig. 1c and Fig 2c) and on 30 vi 2018 (b,d,g; the same specimen as in Figs 1b, 2b): a–b—head and thorax, lateral view; c–d—pterothorax, dorsal view; e–g—vulvar lamina, ventral view. Scale bars, from top to bottom, 4 mm, 2 mm and 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3558051" httpUri="https://zenodo.org/record/3558051/files/figure.png" pageId="6" pageNumber="282">Fig. 6d</figureCitation>
) black, dorsal yellow stripes broad, posteriorly rounded, not reaching alar sinus, anteriorly fused to broad stripes of mesothoracic collar at their lower ends forming two inverted 7s; collar stripes narrowly interrupted with black at medial carina. Antealar spots well expressed but rather small, elongate. A broad yellow stripe occupies ca 2/3 of mesepimeron width; it is shifted towards its anterior margin so that the anterior of the two dark remaining borders is only half as wide as the posterior, wavy one. Mesepisternum also with a broad yellow stripe occupying ca 2/3 of its width, which is strongly shifted to its posterior margin, with only a narrow black border there (
<figureCitation id="EEC92A119946835608E9EB73FE8CFEFA" box="[224,309,261,288]" captionStart="FIGURE 6" captionStartId="8.[151,250,1901,1926]" captionTargetBox="[161,1426,166,1866]" captionTargetId="figure@8.[161,1426,166,1881]" captionTargetPageId="8" captionText="FIGURE 6. Details of female specimens of Microgomphus alani collected in Mondulkiri Province on 3 viii 2016 (c,e; the same specimen as in Fig. 2f) and in the Phnom Kulen Mts on 11 vi 2018 (a,f; the same specimen as in Fig. 1c and Fig 2c) and on 30 vi 2018 (b,d,g; the same specimen as in Figs 1b, 2b): a–b—head and thorax, lateral view; c–d—pterothorax, dorsal view; e–g—vulvar lamina, ventral view. Scale bars, from top to bottom, 4 mm, 2 mm and 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3558051" httpUri="https://zenodo.org/record/3558051/files/figure.png" pageId="7" pageNumber="283">Fig. 6b</figureCitation>
). Metepimeron yellow but its anterior border and anteroventral corner black; posterior margin with a short and narrow black border in its upper part. Mesinfraepisternum black with a long vertical yellow spot in its lower part; metinfraepisternum pale with anterior and posterior black borders (
<figureCitation id="EEC92A11994683560C06EB3BFBDAFEB2" box="[1039,1123,333,360]" captionStart="FIGURE 6" captionStartId="8.[151,250,1901,1926]" captionTargetBox="[161,1426,166,1866]" captionTargetId="figure@8.[161,1426,166,1881]" captionTargetPageId="8" captionText="FIGURE 6. Details of female specimens of Microgomphus alani collected in Mondulkiri Province on 3 viii 2016 (c,e; the same specimen as in Fig. 2f) and in the Phnom Kulen Mts on 11 vi 2018 (a,f; the same specimen as in Fig. 1c and Fig 2c) and on 30 vi 2018 (b,d,g; the same specimen as in Figs 1b, 2b): a–b—head and thorax, lateral view; c–d—pterothorax, dorsal view; e–g—vulvar lamina, ventral view. Scale bars, from top to bottom, 4 mm, 2 mm and 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3558051" httpUri="https://zenodo.org/record/3558051/files/figure.png" pageId="7" pageNumber="283">Fig. 6b</figureCitation>
). Generally, sides of pterothorax yellow with two parallel black stripes along mesopleural and metapleural sutures, the former thrice as broad, with wavy anterior margin producing a projection narrowly embracing the spiracle with black (
<figureCitation id="EEC92A11994683560CD4EBE3FA8CFE75" box="[1245,1333,405,431]" captionStart="FIGURE 1" captionStartId="2.[151,250,1450,1475]" captionTargetBox="[151,1436,553,1431]" captionTargetId="figure@2.[151,1436,553,1431]" captionTargetPageId="2" captionText="FIGURE 1. Male (a, d–e) and female (b–c) specimens of Microgomphus alani in live colours from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–e) of Cambodia, 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e). Not to scale" figureDoi="http://doi.org/10.5281/zenodo.3558041" httpUri="https://zenodo.org/record/3558041/files/figure.png" pageId="7" pageNumber="283">Figs 1b</figureCitation>
,
<figureCitation id="EEC92A11994683560D4DEBE3FADAFE75" box="[1348,1379,405,431]" captionStart="FIGURE 2" captionStartId="3.[151,250,1924,1949]" captionTargetBox="[151,1431,805,1905]" captionTargetId="figure@3.[151,1436,805,1905]" captionTargetPageId="3" captionText="FIGURE 2. Dry male (a, d–e) and female (b–c, f) specimens of Microgomphus alani from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–f) of Cambodia, collected on 3 viii 2016 (f), 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e)." figureDoi="http://doi.org/10.5281/zenodo.3558043" httpUri="https://zenodo.org/record/3558043/files/figure.png" pageId="7" pageNumber="283">2b</figureCitation>
,
<figureCitation id="EEC92A11994683560D79EBE3FA36FE6A" box="[1392,1423,405,432]" captionStart="FIGURE 6" captionStartId="8.[151,250,1901,1926]" captionTargetBox="[161,1426,166,1866]" captionTargetId="figure@8.[161,1426,166,1881]" captionTargetPageId="8" captionText="FIGURE 6. Details of female specimens of Microgomphus alani collected in Mondulkiri Province on 3 viii 2016 (c,e; the same specimen as in Fig. 2f) and in the Phnom Kulen Mts on 11 vi 2018 (a,f; the same specimen as in Fig. 1c and Fig 2c) and on 30 vi 2018 (b,d,g; the same specimen as in Figs 1b, 2b): a–b—head and thorax, lateral view; c–d—pterothorax, dorsal view; e–g—vulvar lamina, ventral view. Scale bars, from top to bottom, 4 mm, 2 mm and 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3558051" httpUri="https://zenodo.org/record/3558051/files/figure.png" pageId="7" pageNumber="283">6b</figureCitation>
). Subalar ridges black but that of metepisternum mostly yellow. Ventral side yellow. Coxae black anteriorly, yellow posteriorly. Legs black but outer side of profemur with a broad long yellowish spot occupying ca 2/3 of its length. Metafemur reaches ca 0.4 of S2 length, its flexor surface with two rows of robust and sparse spines, becoming shorter but not denser towards its base; between these rows there is a third median row of small, very short and more densely set spinulets. Mesofemur with smaller but similarly sparse long spines, without distinct intermittent row of fine spinulets, profemur with three rows of very small, short spinulets. Trochanters and femora were covered with a slight white pruinescence (
<figureCitation id="EEC92A119946835609CAE8E7FDAEFD76" box="[451,535,657,684]" captionStart="FIGURE 2" captionStartId="3.[151,250,1924,1949]" captionTargetBox="[151,1431,805,1905]" captionTargetId="figure@3.[151,1436,805,1905]" captionTargetPageId="3" captionText="FIGURE 2. Dry male (a, d–e) and female (b–c, f) specimens of Microgomphus alani from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–f) of Cambodia, collected on 3 viii 2016 (f), 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e)." figureDoi="http://doi.org/10.5281/zenodo.3558043" httpUri="https://zenodo.org/record/3558043/files/figure.png" pageId="7" pageNumber="283">Fig. 2b</figureCitation>
), later washed out by acetone.
</paragraph>
<paragraph id="764D36949946835608CEE8C3FAD7FC59" blockId="7.[151,1437,153,1979]" pageId="7" pageNumber="283">
Wings (
<figureCitation id="EEC92A11994683560916E8C3FECBFD0A" box="[287,370,693,720]" captionStart="FIGURE 2" captionStartId="3.[151,250,1924,1949]" captionTargetBox="[151,1431,805,1905]" captionTargetId="figure@3.[151,1436,805,1905]" captionTargetPageId="3" captionText="FIGURE 2. Dry male (a, d–e) and female (b–c, f) specimens of Microgomphus alani from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–f) of Cambodia, collected on 3 viii 2016 (f), 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e)." figureDoi="http://doi.org/10.5281/zenodo.3558043" httpUri="https://zenodo.org/record/3558043/files/figure.png" pageId="7" pageNumber="283">Fig. 2b</figureCitation>
) hyaline with a very slight brownish infumation, basally with a slight yellowish tint gradually disappearing towards triangles. Venation black but outer surface inside nodes whitish. FW Ax (1
<superScript id="81879BDC994683560CDFE8AEFB58FD3C" attach="left" box="[1238,1249,728,742]" fontSize="6" pageId="7" pageNumber="283">st</superScript>
and 5
<superScript id="81879BDC994683560D2FE8AEFA8AFD3C" attach="left" box="[1318,1331,728,742]" fontSize="6" pageId="7" pageNumber="283">th</superScript>
primary) 14 (left)–15 (right), HW Ax 10; FW Px 9 (right)–10 (left); HW Px 8 (right)–9(left). Crossveins between Arc and R1–R4 junction: 6 (left)–7 (right) above Rs and 4 (left)–5 (right) below Rs on FW, 3 (left)–4 (right) above Rs and 3 below Rs on HW. One cubito-anal crossvein in all wings. Tornus smoothly curving. Pterostigmata brown, almost transparent, in FW covering 4–4.5 cells below, in HW 3.5–4 cells, bordering longitudinal veins inflated, black.
</paragraph>
<paragraph id="764D36949946835608CEE9FBFE69FAA1" blockId="7.[151,1437,153,1979]" pageId="7" pageNumber="283">
Abdomen black with yellow maculation (
<figureCitation id="EEC92A11994683560A98E9FBFD5FFC7D" box="[657,742,909,935]" captionStart="FIGURE 1" captionStartId="2.[151,250,1450,1475]" captionTargetBox="[151,1436,553,1431]" captionTargetId="figure@2.[151,1436,553,1431]" captionTargetPageId="2" captionText="FIGURE 1. Male (a, d–e) and female (b–c) specimens of Microgomphus alani in live colours from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–e) of Cambodia, 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e). Not to scale" figureDoi="http://doi.org/10.5281/zenodo.3558041" httpUri="https://zenodo.org/record/3558041/files/figure.png" pageId="7" pageNumber="283">Figs 1b</figureCitation>
,
<figureCitation id="EEC92A11994683560AFBE9FBFCA8FC72" box="[754,785,909,936]" captionStart="FIGURE 2" captionStartId="3.[151,250,1924,1949]" captionTargetBox="[151,1431,805,1905]" captionTargetId="figure@3.[151,1436,805,1905]" captionTargetPageId="3" captionText="FIGURE 2. Dry male (a, d–e) and female (b–c, f) specimens of Microgomphus alani from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–f) of Cambodia, collected on 3 viii 2016 (f), 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e)." figureDoi="http://doi.org/10.5281/zenodo.3558043" httpUri="https://zenodo.org/record/3558043/files/figure.png" pageId="7" pageNumber="283">2b</figureCitation>
). S1–S6 with yellow dorsal stripes throughout their length, with uneven margins, becoming narrower on subsequent segments so that those on S1–S3 rather broad, those on S4–S5 narrow with a broader base and tapering to apex, that on S6 linear, hardly noticeable, with a small roundish spot at anterior segment margin. Laterally, S1 with yellow ventrolateral parts occupying ca 2/3 of segment height; dorsal black colour produces a narrow protrusion into this yellow part at about middle of its upper margin. S2 with broad, yellow lateral stripes with even margins throughout its length. S3–S6 with anteriolateral spots expanding along segment anterior margins (especially on S3), and ending before medial suture, that on S3 here truncated, other rounded. In line with these spots and behind them (and medial suture), there are isolated lateral stripes twice as short as respective segments; that on S3 with a truncated anterior end, other ends rounded. On S3, the anteriolateral spot and further stripe are separated by just a black streak along midsegmental suture, on S4–S6 by a broader gap. S7 with large anteriolateral spots occupying ¼ of segment length and a shorter dorsal spot almost fused to them, separated only with vague and irregular blackish streaks. Rest of abdomen black. Cerci narrowly pointed, yellowish, paraprocts (short structures as seen between cerci) also yellowish. Sternites with a slight white pruinescence (
<figureCitation id="EEC92A11994683560D66EF4BFF0DFAA6" captionStart="FIGURE 1" captionStartId="2.[151,250,1450,1475]" captionTargetBox="[151,1436,553,1431]" captionTargetId="figure@2.[151,1436,553,1431]" captionTargetPageId="2" captionText="FIGURE 1. Male (a, d–e) and female (b–c) specimens of Microgomphus alani in live colours from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–e) of Cambodia, 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e). Not to scale" figureDoi="http://doi.org/10.5281/zenodo.3558041" httpUri="https://zenodo.org/record/3558041/files/figure.png" pageId="7" pageNumber="283">Fig. 1b</figureCitation>
), washed out by acetone.
</paragraph>
<paragraph id="764D36949946835608CEEFF3FC8CFA3D" blockId="7.[151,1437,153,1979]" pageId="7" pageNumber="283">Vulvar lamina outline approaches an equilateral triangle but with incised apex as split into two lobes for 60% of its length, anterior margin somewhat convex and lateral margins slightly concave; inner margins of lobes convex so that lobes almost touch each other for 2/3 of their length.</paragraph>
<paragraph id="764D36949946835608CEEF87FF03F9F5" blockId="7.[151,1437,153,1979]" pageId="7" pageNumber="283">Measurements (mm). Hindwing 23; abdomen without appendages 25; total length (with head and appendages) 36.</paragraph>
<paragraph id="764D36949946835608CEEC4CFB72F8DD" blockId="7.[151,1437,153,1979]" pageId="7" pageNumber="283">
<emphasis id="4486EA869946835608CEEC4CFE13F989" box="[199,426,1593,1619]" italics="true" pageId="7" pageNumber="283">Variation in females</emphasis>
. The second female from Phnom Kulen, of
<date id="024C1054994683560B93EC4FFBACF989" box="[922,1045,1593,1619]" pageId="7" pageNumber="283" value="2018-06-11">11 vi 2018</date>
, has
<quantity id="B10A9B71994683560C44EC4EFB1AF989" box="[1101,1187,1592,1619]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="283" unit="mm" value="0.5">0.5 mm</quantity>
shorter hind wing but
<quantity id="B10A9B7199468356089EEC2BFF60F9AD" box="[151,217,1629,1655]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="283" unit="mm" value="2.0">2 mm</quantity>
longer abdomen (
<figureCitation id="EEC92A119946835609AEEC2BFE44F9AD" box="[423,509,1629,1655]" captionStart="FIGURE 1" captionStartId="2.[151,250,1450,1475]" captionTargetBox="[151,1436,553,1431]" captionTargetId="figure@2.[151,1436,553,1431]" captionTargetPageId="2" captionText="FIGURE 1. Male (a, d–e) and female (b–c) specimens of Microgomphus alani in live colours from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–e) of Cambodia, 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e). Not to scale" figureDoi="http://doi.org/10.5281/zenodo.3558041" httpUri="https://zenodo.org/record/3558041/files/figure.png" pageId="7" pageNumber="283">Figs 1c</figureCitation>
,
<figureCitation id="EEC92A11994683560A02EC2BFD9FF9AD" box="[523,550,1629,1655]" captionStart="FIGURE 2" captionStartId="3.[151,250,1924,1949]" captionTargetBox="[151,1431,805,1905]" captionTargetId="figure@3.[151,1436,805,1905]" captionTargetPageId="3" captionText="FIGURE 2. Dry male (a, d–e) and female (b–c, f) specimens of Microgomphus alani from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–f) of Cambodia, collected on 3 viii 2016 (f), 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e)." figureDoi="http://doi.org/10.5281/zenodo.3558043" httpUri="https://zenodo.org/record/3558043/files/figure.png" pageId="7" pageNumber="283">2c</figureCitation>
) (HW 25.5, Ab 28.5, total length 37; FW Ax 14). The latter may be related to its somewhat immature state, while the female described above was obviously fully mature as collected during oviposition (
<figureCitation id="EEC92A11994683560908ECD3FEEBF965" box="[257,338,1701,1727]" captionStart="FIGURE 1" captionStartId="2.[151,250,1450,1475]" captionTargetBox="[151,1436,553,1431]" captionTargetId="figure@2.[151,1436,553,1431]" captionTargetPageId="2" captionText="FIGURE 1. Male (a, d–e) and female (b–c) specimens of Microgomphus alani in live colours from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–e) of Cambodia, 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e). Not to scale" figureDoi="http://doi.org/10.5281/zenodo.3558041" httpUri="https://zenodo.org/record/3558041/files/figure.png" pageId="7" pageNumber="283">Fig. 1c</figureCitation>
). The second Phnom Kulen female has vestigial yellow maculation also on S8: a small triangular anteriodorsal spot, lateral streaks at anterior segment margin accompanied by a pair of tiny elongate spots just above them at some distance from segment margin. In this female the paraprocts are also dark greyish.
</paragraph>
<paragraph id="764D36949946835608CEED67FD64F84D" blockId="7.[151,1437,153,1979]" pageId="7" pageNumber="283">
The only female from
<collectingRegion id="B436F8769946835609C5ED67FD00F8F1" box="[460,697,1809,1835]" country="Cambodia" name="Mondolkiri" pageId="7" pageNumber="283">Mondulkiri Province</collectingRegion>
(of
<date id="024C1054994683560AE2ED67FCD2F8F6" box="[747,875,1809,1836]" pageId="7" pageNumber="283" value="2016-08-03">3 viii 2016</date>
) (
<figureCitation id="EEC92A11994683560B8DED67FC6AF8F6" box="[900,979,1809,1836]" captionStart="FIGURE 2" captionStartId="3.[151,250,1924,1949]" captionTargetBox="[151,1431,805,1905]" captionTargetId="figure@3.[151,1436,805,1905]" captionTargetPageId="3" captionText="FIGURE 2. Dry male (a, d–e) and female (b–c, f) specimens of Microgomphus alani from the Phnom Kulen Mountains, Siem Reap Province (a–c) and Mondulkiri Province (d–f) of Cambodia, collected on 3 viii 2016 (f), 11 vi 2018 (c), 17 vi 2018 (a), 30 vi 2018 (b) and 26 vi 2018 (d–e)." figureDoi="http://doi.org/10.5281/zenodo.3558043" httpUri="https://zenodo.org/record/3558043/files/figure.png" pageId="7" pageNumber="283">Fig. 2f</figureCitation>
) is ca 10% larger than the one used for description (HW 27.5, Ab 29.5; total length 41; FW Ax 14/15). Its pale stripe across the frons is interrupted at the middle (
<figureCitation id="EEC92A119946835608FDED2FFEFCF8A9" box="[244,325,1880,1907]" captionStart="FIGURE 5" captionStartId="6.[151,250,1669,1694]" captionTargetBox="[170,1436,174,1602]" captionTargetId="figure@6.[151,1436,174,1650]" captionTargetPageId="6" captionText="FIGURE 5. Frontal view of head (left) and occiput (right) of female specimens of Microgomphus alani collected on 3 viii 2016 in Mondulkiri Province (a; the same specimen as in Fig. 2f) and in the Phnom Kulen Mts on 11 vi 2018 (b, the same specimen as in Figs 1c, 2c) and 30 vi 2018 (c, the same specimen as in Figs 1b, 2b). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3558049" httpUri="https://zenodo.org/record/3558049/files/figure.png" pageId="7" pageNumber="283">Fig. 5a</figureCitation>
); cerci and paraprocts whitish. Although treated with acetone, this specimen has a strong while pruinescence of the coxae and prothorax lower surface.
</paragraph>
<paragraph id="764D36949946835608CEEDD7FA31F861" blockId="7.[151,1437,153,1979]" box="[199,1416,1952,1979]" pageId="7" pageNumber="283">
Both these females have a symmetrical occiput with two horns at either side of the central hump (
<figureCitation id="EEC92A11994683560D05EDD7FAF7F861" box="[1292,1358,1952,1979]" captionStart="FIGURE 5" captionStartId="6.[151,250,1669,1694]" captionTargetBox="[170,1436,174,1602]" captionTargetId="figure@6.[151,1436,174,1650]" captionTargetPageId="6" captionText="FIGURE 5. Frontal view of head (left) and occiput (right) of female specimens of Microgomphus alani collected on 3 viii 2016 in Mondulkiri Province (a; the same specimen as in Fig. 2f) and in the Phnom Kulen Mts on 11 vi 2018 (b, the same specimen as in Figs 1c, 2c) and 30 vi 2018 (c, the same specimen as in Figs 1b, 2b). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3558049" httpUri="https://zenodo.org/record/3558049/files/figure.png" pageId="7" pageNumber="283">Fig. 5</figureCitation>
a–b).
</paragraph>
<caption id="228D661C99498359089EED1BFB86F82B" ID-DOI="http://doi.org/10.5281/zenodo.3558051" ID-Zenodo-Dep="3558051" httpUri="https://zenodo.org/record/3558051/files/figure.png" pageId="8" pageNumber="284" startId="8.[151,250,1901,1926]" targetBox="[161,1426,166,1866]" targetPageId="8">
<paragraph id="764D369499498359089EED1BFB86F82B" blockId="8.[151,1437,1901,2034]" pageId="8" pageNumber="284">
<emphasis id="4486EA8699498359089EED1BFEACF85C" bold="true" box="[151,277,1901,1926]" pageId="8" pageNumber="284">FIGURE 6.</emphasis>
Details of female specimens of
<taxonomicName id="B1F24D17994983590A62ED1BFCF8F85F" authorityName="Kosterin" authorityYear="2019" box="[619,833,1901,1925]" class="Insecta" family="Gomphidae" genus="Microgomphus" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="284" phylum="Arthropoda" rank="species" species="alani">
<emphasis id="4486EA86994983590A62ED1BFCF8F85F" box="[619,833,1901,1925]" italics="true" pageId="8" pageNumber="284">Microgomphus alani</emphasis>
</taxonomicName>
collected in Mondulkiri Province on 3 viii 2016 (c,e; the same specimen as in Fig. 2f) and in the Phnom Kulen Mts on 11 vi 2018 (a,f; the same specimen as in Fig. 1c and Fig 2c) and on 30 vi 2018 (b,d,g; the same specimen as in Figs 1b, 2b): a
<emphasis id="4486EA86994983590AF4EDC0FCB3F817" box="[765,778,1974,1997]" italics="true" pageId="8" pageNumber="284">–</emphasis>
b—head and thorax, lateral view; c
<emphasis id="4486EA86994983590C65EDC0FBC0F817" box="[1132,1145,1974,1997]" italics="true" pageId="8" pageNumber="284">–</emphasis>
d—pterothorax, dorsal view; e
<emphasis id="4486EA869949835908ABEDACFF16F82B" box="[162,175,2010,2033]" italics="true" pageId="8" pageNumber="284">–</emphasis>
g—vulvar lamina, ventral view. Scale bars, from top to bottom, 4 mm, 2 mm and 0.5 mm.
</paragraph>
</caption>
<caption id="228D661C99488358089EEDC3FBBFF82B" ID-DOI="http://doi.org/10.5281/zenodo.3558053" ID-Zenodo-Dep="3558053" httpUri="https://zenodo.org/record/3558053/files/figure.png" pageId="9" pageNumber="285" startId="9.[151,250,1973,1998]" targetBox="[345,1242,166,1896]" targetPageId="9">
<paragraph id="764D369499488358089EEDC3FBBFF82B" blockId="9.[151,1436,1973,2034]" pageId="9" pageNumber="285">
<emphasis id="4486EA8699488358089EEDC3FEACF814" bold="true" box="[151,277,1973,1998]" pageId="9" pageNumber="285">FIGURE 7.</emphasis>
Hind legs of
<taxonomicName id="B1F24D179948835809A1EDC3FDC4F817" authorityName="Kosterin" authorityYear="2019" box="[424,637,1973,1997]" class="Insecta" family="Gomphidae" genus="Microgomphus" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="285" phylum="Arthropoda" rank="species" species="alani">
<emphasis id="4486EA869948835809A1EDC3FDC4F817" box="[424,637,1973,1997]" italics="true" pageId="9" pageNumber="285">Microgomphus alani</emphasis>
</taxonomicName>
, a—male from Mondulkiri Province, 13 vi 2014; b—female from Modulkiri Province, 3 viii 2016; c—female from Phnom Kulen Mts, 30 vi 2018. Scale bar 1 mm.
</paragraph>
</caption>
<paragraph id="764D3694994B835B08CEEAEFFD42FE51" blockId="10.[151,1437,152,1007]" pageId="10" pageNumber="286">
<emphasis id="4486EA86994B835B08CEEAEFFD99FF69" bold="true" box="[199,544,152,179]" pageId="10" pageNumber="286">
<emphasis id="4486EA86994B835B08CEEAEFFDA0FF69" bold="true" box="[199,537,152,179]" italics="true" pageId="10" pageNumber="286">Main differences from males</emphasis>
.
</emphasis>
Besides the reproductive organs, the females of
<taxonomicName id="B1F24D17994B835B0C47EAECFB16FF69" authorityName="Kosterin" authorityYear="2019" box="[1102,1199,153,179]" class="Insecta" family="Gomphidae" genus="Microgomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="286" phylum="Arthropoda" rank="species" species="alani">
<emphasis id="4486EA86994B835B0C47EAECFB16FF69" box="[1102,1199,153,179]" italics="true" pageId="10" pageNumber="286">M. alani</emphasis>
</taxonomicName>
differ from males in having a central hump and a pair of horns on the occiput (which is simple and straight in the male;
<figureCitation id="EEC92A11994B835B0CEDEACBFAF0FF0D" box="[1252,1353,189,215]" captionStart="FIGURE 3" captionStartId="4.[151,250,1902,1927]" captionTargetBox="[266,1321,166,1874]" captionTargetId="figure@4.[266,1321,166,1883]" captionTargetPageId="4" captionText="FIGURE 3. Details of male specimens of Microgomphus alani collected on 26 vi 2018 in Mondulkiri Province (a–d; the same specimen as in Fig. 2d) and on 17 vi 2018 in the Phnom Kulen Mts (a–h, the same specimen as in Fig. 1a and Fig 2a): a,e—head and thorax, lateral view; b,f—pterothorax, dorsal view; c,g—head, frontal (c) and dorsofrontal (g) view; d,h—secondary genitalia, lateral view. Scale bars. from top to bottom, 4 mm, 2 mm, 2 mm and 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3558045" httpUri="https://zenodo.org/record/3558045/files/figure.png" pageId="10" pageNumber="286">Fig. 3c,g</figureCitation>
), a pair of strong blunt tubercles behind the ocelli (vs gentle swellings set more apart from each other in males;
<figureCitation id="EEC92A11994B835B0D20EA97FA37FF21" box="[1321,1422,225,251]" captionStart="FIGURE 3" captionStartId="4.[151,250,1902,1927]" captionTargetBox="[266,1321,166,1874]" captionTargetId="figure@4.[266,1321,166,1883]" captionTargetPageId="4" captionText="FIGURE 3. Details of male specimens of Microgomphus alani collected on 26 vi 2018 in Mondulkiri Province (a–d; the same specimen as in Fig. 2d) and on 17 vi 2018 in the Phnom Kulen Mts (a–h, the same specimen as in Fig. 1a and Fig 2a): a,e—head and thorax, lateral view; b,f—pterothorax, dorsal view; c,g—head, frontal (c) and dorsofrontal (g) view; d,h—secondary genitalia, lateral view. Scale bars. from top to bottom, 4 mm, 2 mm, 2 mm and 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3558045" httpUri="https://zenodo.org/record/3558045/files/figure.png" pageId="10" pageNumber="286">Fig. 3c,g</figureCitation>
). The metafemur bears two rows of strong and sparse spines in females (with an intermittent row or fine spinulets) but only fine and dense spinulets in males, as discussed below. In maculation, the main difference of females from males is the presence of long lateral (subventral) pale stripes behind the short anteriolateral spots on S3
<emphasis id="4486EA86994B835B0CC4EB38FB62FEBD" box="[1229,1243,334,359]" italics="true" pageId="10" pageNumber="286">–</emphasis>
S6. Profemur has a large yellowish area in females but is black in males.
</paragraph>
<paragraph id="764D3694994B835B08CEEBE2FAEEFC35" blockId="10.[151,1437,152,1007]" pageId="10" pageNumber="286">
<emphasis id="4486EA86994B835B08CEEBE2FDE3FE75" bold="true" box="[199,602,404,431]" pageId="10" pageNumber="286">
<emphasis id="4486EA86994B835B08CEEBE2FDEAFE75" bold="true" box="[199,595,404,431]" italics="true" pageId="10" pageNumber="286">Differential diagnosis for females</emphasis>
.
</emphasis>
The prominent central hump on the occipital plate is unique among the known females of
<taxonomicName id="B1F24D17994B835B0979EBCFFDA0FE09" authorityName="Selys" authorityYear="1858" box="[368,537,441,467]" class="Insecta" family="Gomphidae" genus="Microgomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="286" phylum="Arthropoda" rank="genus">
<emphasis id="4486EA86994B835B0979EBCFFDA0FE09" box="[368,537,441,467]" italics="true" pageId="10" pageNumber="286">Microgomphus</emphasis>
</taxonomicName>
spp. The female head sculpture of
<taxonomicName id="B1F24D17994B835B0BBEEBCFFA25FE09" authority="Lieftinck, 1929" authorityName="Lieftinck" authorityYear="1929" box="[951,1436,441,467]" class="Insecta" family="Gomphidae" genus="Microgomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="286" phylum="Arthropoda" rank="species" species="telyphonus">
<emphasis id="4486EA86994B835B0BBEEBCFFB5BFE09" box="[951,1250,441,467]" italics="true" pageId="10" pageNumber="286">Microgomphus telyphonus</emphasis>
<bibRefCitation id="12634B65994B835B0CE4EBCFFA25FE09" author="Lieftinck, M. A." box="[1261,1436,441,467]" pageId="10" pageNumber="286" pagination="109 - 147" refId="ref9545" refString="Lieftinck, M. A. (1929) Contributions to the dragonfly fauna of the Sondaic area. Tijdschrift voor Entomologie, 72 (2), 109 - 147." type="journal article" year="1929">Lieftinck, 1929</bibRefCitation>
</taxonomicName>
(presently considered a subspecies of
<taxonomicName id="B1F24D17994B835B0A36EBA8FCFDFE2D" authority="Selys, 1858" authorityName="Selys" authorityYear="1858" box="[575,836,476,503]" class="Insecta" family="Gomphidae" genus="Microgomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="286" phylum="Arthropoda" rank="species" species="chelifer">
<emphasis id="4486EA86994B835B0A36EBA8FD03FE2D" box="[575,698,477,503]" italics="true" pageId="10" pageNumber="286">M. chelifer</emphasis>
Selys, 1858
</taxonomicName>
) was described (and illustrated) as follows: “The free margin of the occipital plate, in frontal view, with a low but sharply crested elevation, directed upwards under right angle; this margin slightly concave in the middle, with a row of 5
<emphasis id="4486EA86994B835B0B8DE850FC2BFDE5" box="[900,914,550,575]" italics="true" pageId="10" pageNumber="286">–</emphasis>
6 robust, irregular black spines on either side; anterior surface with a low, central tubercle, just behind the two lateral ocelli” (
<bibRefCitation id="12634B65994B835B0C2BE83FFAF8FDB9" author="Lieftinck, M. A." box="[1058,1345,585,611]" pageId="10" pageNumber="286" pagination="109 - 147" refId="ref9545" refString="Lieftinck, M. A. (1929) Contributions to the dragonfly fauna of the Sondaic area. Tijdschrift voor Entomologie, 72 (2), 109 - 147." type="journal article" year="1929">
Lieftinck 1929: 126
<emphasis id="4486EA86994B835B0D0CE83CFAAAFDB9" box="[1285,1299,586,611]" italics="true" pageId="10" pageNumber="286">–</emphasis>
127
</bibRefCitation>
).
<bibRefCitation id="12634B65994B835B0D5FE83FFF5BFD5D" author="Fraser, F. C." pageId="10" pageNumber="286" refId="ref9153" refString="Fraser, F. C. (1934) The Fauna of British India, including Ceylon and Burma. Odonata. Vol. II. Taylor and Francis, London, 461 pp." type="book" year="1934">Fraser (1934)</bibRefCitation>
stated that the female occiput in
<taxonomicName id="B1F24D17994B835B0A42E81BFC44FD5D" authority="(Selys, 1854)" authorityName="Selys" authorityYear="1854" baseAuthorityName="Selys" baseAuthorityYear="1854" box="[587,1021,620,647]" class="Insecta" family="Gomphidae" genus="Microgomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="286" phylum="Arthropoda" rank="species" species="troquatus">
<emphasis id="4486EA86994B835B0A42E81BFCDAFD5D" box="[587,867,621,647]" italics="true" pageId="10" pageNumber="286">Microgomphus troquatus</emphasis>
(Selys, 1854)
</taxonomicName>
is “with a low, slightly concave crest, which is bordered with about ten minute spines” (that is like in
<taxonomicName id="B1F24D17994B835B0B7CE8E4FBCAFD71" box="[885,1139,657,683]" class="Insecta" family="Gomphidae" genus="Microgomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="286" phylum="Arthropoda" rank="subSpecies" species="chelifer" subSpecies="telyphonus">
<emphasis id="4486EA86994B835B0B7CE8E4FBCAFD71" box="[885,1139,657,683]" italics="true" pageId="10" pageNumber="286">M. chelifer telyphonus</emphasis>
</taxonomicName>
). He described that of
<taxonomicName id="B1F24D17994B835B0D77E8E4FE1AFD15" authority="Fraser, 1923" authorityName="Fraser" authorityYear="1923" class="Insecta" family="Gomphidae" genus="Microgomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="286" phylum="Arthropoda" rank="species" species="lilliputians">
<emphasis id="4486EA86994B835B0D77E8E4FEA9FD15" italics="true" pageId="10" pageNumber="286">M. lilliputians</emphasis>
Fraser, 1923
</taxonomicName>
as “simple, sinuous, a little notched at its middle”, that or
<emphasis id="4486EA86994B835B0C3CE8C0FBEDFD15" box="[1077,1108,694,719]" italics="true" pageId="10" pageNumber="286">M.</emphasis>
(?)
<emphasis id="4486EA86994B835B0C88E8C3FB46FD15" box="[1153,1279,693,719]" italics="true" pageId="10" pageNumber="286">minusculus</emphasis>
(Selys, 1878) as “simple, straight”, but provided no information on the female head sculpture of
<taxonomicName id="B1F24D17994B835B0C40E8AFFF6BFCCD" authority="Fraser, 1924" authorityName="Fraser" authorityYear="1924" class="Insecta" family="Gomphidae" genus="Microgomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="286" phylum="Arthropoda" rank="species" species="souteri">
<emphasis id="4486EA86994B835B0C40E8AFFAF1FD29" box="[1097,1352,729,755]" italics="true" pageId="10" pageNumber="286">Microgomphus souteri</emphasis>
Fraser, 1924
</taxonomicName>
,
<emphasis id="4486EA86994B835B08D6E888FF47FCCD" box="[223,254,766,791]" italics="true" pageId="10" pageNumber="286">M.</emphasis>
(?)
<emphasis id="4486EA86994B835B0924E88BFE2CFCCD" box="[301,405,765,791]" italics="true" pageId="10" pageNumber="286">verticalis</emphasis>
(Selys, 1873) and
<taxonomicName id="B1F24D17994B835B0A67E888FCC4FCCD" authority="Fraser, 1923" authorityName="Fraser" authorityYear="1923" box="[622,893,765,791]" class="Insecta" family="Gomphidae" genus="Microgomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="286" phylum="Arthropoda" rank="species" species="loogali">
<emphasis id="4486EA86994B835B0A67E888FD5DFCCD" box="[622,740,765,791]" italics="true" pageId="10" pageNumber="286">M. loogali</emphasis>
Fraser, 1923
</taxonomicName>
, that could imply absence of notable structures. The female of
<taxonomicName id="B1F24D17994B835B0934E957FD54FCE1" authority="Lieftinck, 1940" authorityName="Lieftinck" authorityYear="1940" box="[317,749,801,827]" class="Insecta" family="Gomphidae" genus="Microgomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="286" phylum="Arthropoda" rank="species" species="wijaya">
<emphasis id="4486EA86994B835B0934E957FD81FCE1" box="[317,568,801,827]" italics="true" pageId="10" pageNumber="286">Microgomphus wijaya</emphasis>
<bibRefCitation id="12634B65994B835B0A36E957FD54FCE1" author="Lieftinck, M. A." box="[575,749,801,827]" pageId="10" pageNumber="286" pagination="79 - 117" refId="ref9577" refString="Lieftinck, M. A. (1940) On some Odonata collected in Ceylon, with descriptions of new species and larvae. Ceylon Journal of Sciences, Biology, 22 (1), 79 - 117." type="journal article" year="1940">Lieftinck, 1940</bibRefCitation>
</taxonomicName>
is said to have the “occiput as in the opposite sex”, in which “occipital plate straight posteriorly, armed with a row of microscopical, erect, finely pointed denticles along margin” (
<bibRefCitation id="12634B65994B835B0897E91FFEC8FC59" author="Lieftinck, M. A." box="[158,369,873,899]" pageId="10" pageNumber="286" pagination="79 - 117" refId="ref9577" refString="Lieftinck, M. A. (1940) On some Odonata collected in Ceylon, with descriptions of new species and larvae. Ceylon Journal of Sciences, Biology, 22 (1), 79 - 117." type="journal article" year="1940">Lieftinck 1940: 98</bibRefCitation>
, 100). The female of
<taxonomicName id="B1F24D17994B835B0A55E91CFD45FC59" authorityName="Coniff &amp; Limbu" authorityYear="2018" box="[604,764,873,899]" class="Insecta" family="Gomphidae" genus="Microgomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="286" phylum="Arthropoda" rank="species" species="phewataali">
<emphasis id="4486EA86994B835B0A55E91CFD45FC59" box="[604,764,873,899]" italics="true" pageId="10" pageNumber="286">M. phewataali</emphasis>
</taxonomicName>
has “occipital margin with two small close-set tubercles less prominent than in male, upper frons ... peaked in middle with shallow cleft” (Coniff &amp; Singh Limbu 2018: 282). The head of the female of
<taxonomicName id="B1F24D17994B835B0981E9C4FE43FC11" authorityName="Makbun &amp; Fleck" authorityYear="2018" box="[392,506,945,971]" class="Insecta" family="Gomphidae" genus="Microgomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="286" phylum="Arthropoda" rank="species" species="farrelli">
<emphasis id="4486EA86994B835B0981E9C4FE43FC11" box="[392,506,945,971]" italics="true" pageId="10" pageNumber="286">M. farrelli</emphasis>
</taxonomicName>
is not described except for the statement of “absence of occipital horns (presence in
<taxonomicName id="B1F24D17994B835B089EE9A0FEA8FC35" authorityName="' in Kosterin" authorityYear="2016" box="[151,273,981,1007]" class="Insecta" family="Gomphidae" genus="Heliogomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="286" phylum="Arthropoda" rank="species" species="svihleri">
<emphasis id="4486EA86994B835B089EE9A0FEA8FC35" box="[151,273,981,1007]" italics="true" pageId="10" pageNumber="286">M. svihleri</emphasis>
</taxonomicName>
)...”, the illustration shows a simple straight occiput margin (
<bibRefCitation id="12634B65994B835B0BBEE9A3FB4AFC35" author="Makbun, N. &amp; Fleck, G." box="[951,1267,980,1007]" pageId="10" pageNumber="286" pagination="442 - 450" refId="ref9617" refString="Makbun, N. &amp; Fleck, G. (2018) Description of Microgomphus farrelli sp. nov. (Odonata: Anisoptera: Gomphidae) based on adults of both sexes and larvae from Northern Thailand. Zootaxa, 4422 (3), 442 - 450. https: // doi. org / 10.11646 / zootaxa. 4422.3.10" type="journal article" year="2018">Makbun &amp; Fleck 2018: 445</bibRefCitation>
, fig. 10).
</paragraph>
</subSubSection>
</treatment>
</document>

134
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<treatment id="FE5BCDCC6176779F32E337C12861E0FF" ID-GBIF-Taxon="231544920" LSID="urn:lsid:plazi:treatment:FE5BCDCC6176779F32E337C12861E0FF" httpUri="http://treatment.plazi.org/id/FE5BCDCC6176779F32E337C12861E0FF" lastPageNumber="240" pageNumber="240" sourceUrl="http://www.departments.bucknell.edu/biology/resources/msw3/browse.asp?id=13700128">
<subSubSection id="FFAD5DCEFF68B5855FBB2595EA003B29" pageNumber="240" type="nomenclature">
<paragraph id="4343B84ECDD379E519323902ED51A9A1" pageNumber="240">
<taxonomicName id="E707BF687AB5A9072251CC1790A6ABCC" ID-CoL="ZK62" authority="Sody 1936" authorityName="Sody" authorityYear="1936" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageNumber="240" phylum="Chordata" rank="species" species="maxi">
<emphasis id="AEC30160706B4CF26F3C7C055759FDBD" italics="true" pageNumber="240">Crocidura maxi</emphasis>
Sody 1936
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="2680194FCCF65D043CA91AAC38144B45" pageNumber="240" type="reference_group">
<paragraph id="B6830C8AB01C8EE9D3BFB0C4FDF06C21" pageNumber="240">
<treatmentCitation id="56C72299D8BA389A9FE682384704E572" author="Sody" authority="Sody 1936" authorityName="Sody" authorityYear="1936" class="Mammalia" family="Soricidae" genus="Crocidura" journal="Natuurk. Tijdschr. Ned.-Ind." kingdom="Animalia" order="Soricomorpha" page="53" pageNumber="240" phylum="Chordata" rank="species" species="maxi" title="Crocidura maxi" volume="96" year="1936">
<taxonomicName id="6F9B8E77D52DBC4CC042ECEECA53FFBA" ID-CoL="ZK62" authority="Sody 1936" authorityName="Sody" authorityYear="1936" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageNumber="240" phylum="Chordata" rank="species" species="maxi">
<emphasis id="45287DE980B4BB62894CD8F721CAA773" italics="true" pageNumber="240">Crocidura maxi</emphasis>
Sody 1936
</taxonomicName>
,
<bibCitation id="E8E4E1AB06B9BEA537383E7CC98FBC54" author="Sody" journal="Natuurk. Tijdschr. Ned.-Ind." pageNumber="240" pagination="53" title="Crocidura maxi" volume="96" year="1936">Natuurk. Tijdschr. Ned.-Ind., 96: 53</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="1B52B19FC65F1372E8640A30AF450250" pageNumber="240" type="materials_examined">
<paragraph id="2D0D71676E0D81D5F437045444E3EC3E" pageNumber="240">
<emphasis id="0C8D6B759FF428DC6B2518C0BB98C4D1" inLineHeading="true" pageNumber="240">Type Locality:</emphasis>
<materialsCitation id="731CECBBCF59A186EACF9C20E4305A04" ID-GBIF-Occurrence="4867338316" country="Indonesia" pageNumber="240" stateProvince="Jawa Timur">
<collectingCountry id="F5B6717AFFCBE022ECA29B911ABCFCC2" name="Indonesia">Indonesia</collectingCountry>
,
<collectingRegion id="526B9F8BF20201119EDD301CF9F8E0F7" country="Indonesia" name="Jawa Timur">Java</collectingRegion>
,
<collectingRegion id="59CB51D8C7E05F4D76487E00C4784721" country="Indonesia" name="Jawa Timur">East Java</collectingRegion>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="08295BC68E369586DCD91FF00596A8EC" pageNumber="240" type="vernacular_names">
<paragraph id="A8BD59BA4A51F74B196C403D38B9D50B" pageNumber="240">
<emphasis id="2933BED27B277E0F528F6D7CBF513F73" inLineHeading="true" pageNumber="240">Vernacular Names:</emphasis>
<vernacularName id="D7780F4C4C60E5DC61D5F3FE39480361" language="eng" pageNumber="240">Javanese Shrew</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="BC6DF8F5E293B8D08E0769C992CF7721" pageNumber="240" type="distribution">
<paragraph id="1B3484B67290140422C13AE403539155" pageNumber="240">
<emphasis id="F58BEACD858CEFED402F43597BE43BFD" inLineHeading="true" pageNumber="240">Distribution:</emphasis>
<collectingRegion id="8E708E51BDCBDE1919E06A64D0389AE3" country="Indonesia" name="Jawa Timur">Java</collectingRegion>
, Lesser Sunda Isls (
<collectingRegion id="B52EA1EE1D6DD104B1748CE4B5800A4A" country="Indonesia" name="Bali">Bali</collectingRegion>
, Sumbawa, Komodo, Sumba,
<collectingCountry id="CB8773F867B97FC51565753C81E57122" name="Indonesia">Flores</collectingCountry>
, Alor, Roti, Timor), and
<collectingCountry id="2349B1DB975FE2894C2FE2FFF2A0DEA6" name="Indonesia">Ambon</collectingCountry>
(
<collectingCountry id="0FB056D83D8E6504E58D7F09E69A7957" name="Indonesia">Moluccas</collectingCountry>
,
<collectingCountry id="39CE6CF86ED12AF9919EC241C888FDEE" name="Indonesia">Indonesia</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="6C7063A3EECEF1179D6A458471C5AFDB" pageNumber="240" type="conservation">
<paragraph id="ACB4E25B29C3AB7D14149268082AEC07" pageNumber="240">
<emphasis id="B747619811999005302F69243A6B3ECE" inLineHeading="true" pageNumber="240">Conservation:</emphasis>
<collectionCode id="1FA6593E525F3B09635B150878BD7796">IUCN</collectionCode>
– Lower Risk (lc).
</paragraph>
</subSubSection>
<subSubSection id="BD474E1025C770C0AB6669C6CE239E42" pageNumber="240" type="discussion">
<paragraph id="F838A5E2D918A5FFAF2DF70C830A3FFE" pageNumber="240">
<emphasis id="DEE096D5BB2C636EF57889DF2C7E790F" inLineHeading="true" pageNumber="240">Discussion:</emphasis>
Occurs sympatrically with
<taxonomicName id="452B5C36F1C4B0D77591C968182EC0E3" authorityName="Peters" authorityYear="1870" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" phylum="Chordata" rank="species" species="monticola">
<emphasis id="57AEC0F0A686B9AA63E9C8891F117012" italics="true" pageNumber="240">monticola</emphasis>
</taxonomicName>
in
<collectingRegion id="6DE14B5494DB76AC4449407C3832F812" country="Indonesia" name="Jawa Timur">Java</collectingRegion>
; see Jenkins (1982) and
<bibRefCitation id="FDB4486CA434EA8DD59000F90B087B96" author="Kitchener, D. J. &amp; S. Hisheh &amp; L. H. Schmitt &amp; A. Suyanto" refId="ref188137" refString="Kitchener, D. J., S. Hisheh, L. H. Schmitt, and A. Suyanto. 1994 a. Shrews (Soricidae: Crocidura) from the Lesser Sunda Islands and southeast Maluku, eastern Indonesia. Australian Mammalogy, 17: 7 - 17." year="1994">
Kitchener et al. (1994
<emphasis id="11AAC6067F8B161A89C3AAFCA27F94DB" italics="true" pageNumber="240">a</emphasis>
)
</bibRefCitation>
. Ruedi (1995) questioned the validity of
<taxonomicName id="819D639C43EF857E98BC4BD74FC1AF48" authorityName="Sody" authorityYear="1936" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" phylum="Chordata" rank="species" species="maxi">
<emphasis id="7D527A2F7B13CFF17CF214A7E6F52248" italics="true" pageNumber="240">maxi</emphasis>
</taxonomicName>
but stated (p. 243) that &quot;Eastern representatives, which are usually larger, with a flatter skull profile than nominal and central forms, could be treated as a distinct subspecies,
<taxonomicName id="9A01A563CFB4CF8DC25993CE718ED8E1" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" phylum="Chordata" rank="subSpecies" species="monticola" subSpecies="maxi">
<emphasis id="F6ECDC59557376A9EEF513B15288281F" italics="true" pageNumber="240">C. monticola maxi</emphasis>
</taxonomicName>
.&quot;.
</paragraph>
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130
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<taxonomicName id="73830B075E0D5B3035B17D375C863104" ID-CoL="3GJFM" LSID-ZBK="urn:lsid:zoobank.org:act:629CDB4F-2E7B-4F96-A91B-164A78A4171E" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" lsidName-ZBK="Glyptothorax zanaensis Wu, He &amp; Chu 1981:74 (English p. 79)" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="zanaensis">Glyptothorax zanaensis Wu, He &amp; Chu 1981</taxonomicName>
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<paragraph id="2277A0A3ECC5E1A63E0FBA8B28B3EA80" pageNumber="57">
<taxonomicName id="51D857CB18B9B3C531D87F924FFF3FB4" LSID-ZBK="urn:lsid:zoobank.org:act:629CDB4F-2E7B-4F96-A91B-164A78A4171E" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" lsidName-ZBK="Glyptothorax zanaensis Wu, He &amp; Chu 1981:74 (English p. 79)" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="zanaensis">Glyptothorax zanaensis Wu, He &amp; Chu 1981</taxonomicName>
: 74 (English p. 79), fig. 1 (1-b).
<materialsCitation id="2AC0D4FE0418B22FBF6428560E56E3DB" ID-GBIF-Occurrence="923901431" collectionCode="Inst. Hydrobiol. Acad. Sin." country="China" location="Type locality: Qinghai-Xizang plateau region" specimenCode="Inst. Hydrobiol. Acad. Sin. 606164 - 66, 606168, 606170 - 74, 606178, 6006651 - 53" typeStatus="Syntypes">
<location id="E80458284507E57328A6AE604BC8D9A2" LSID="urn:lsid:plazi:treatment:FE5BD2DF623CB3D7D42E2BB02EBFFF2E:E80458284507E57328A6AE604BC8D9A2" country="China" name="Type locality: Qinghai-Xizang plateau region">Type locality: Qinghai-Xizang plateau region</location>
,
<collectingCountry id="1FA939BA7CA64FDD10B70B57585B4E7D">China</collectingCountry>
.
<typeStatus id="A703422A00F68A2D7AFF0583A3DD8588">Syntypes</typeStatus>
:
<specimenCode id="82FABD1DC71114F6594A6A8825136C2E">
<collectionCode id="7DCC63A92598CD77DA432232726BBD74">Inst. Hydrobiol. Acad. Sin.</collectionCode>
606164-66
</specimenCode>
(3),
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(1),
<specimenCode id="5D09DBC703EABCFDE10AABA854B3BD4F">606170-74</specimenCode>
(5),
<specimenCode id="0A1762D2D7E2E046A5CAEB68A0097900">606178</specimenCode>
(1),
<specimenCode id="C6B6B6A5472E6DD5CCE22CBCCBF23980">6006651-53</specimenCode>
(3).
</materialsCitation>
</paragraph>
<paragraph id="6E4AD2D16A1A08BA1C286F2D2B200176" pageNumber="57">
<taxonomicName id="B293F3993CB9293EC6F64859CE407FF0" LSID-ZBK="urn:lsid:zoobank.org:act:78E015B3-28D5-4A1B-B06A-880C3304DBF3" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" lsidName-ZBK="Glyptothorax longinema Li 1984:[English p. 88]" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="longinema">Glyptothorax longinema Li 1984b</taxonomicName>
: [English p. 88], fig. 6.
<materialsCitation id="9E1583C2812A0E42310C0DF3FBB0B4F7" ID-GBIF-Occurrence="923901444" collectionCode="KIZ" country="China" location="Type locality [of Glyptothorax longinema Li]: Lushui Xian [Salween drainage]" specimenCode="KIZ 741097 [and others]" stateProvince="Yunnan" typeStatus="Holotype, Paratypes">
<location id="0A141CAED98A631CEAA30B07D4954C38" LSID="urn:lsid:plazi:treatment:FE5BD2DF623CB3D7D42E2BB02EBFFF2E:0A141CAED98A631CEAA30B07D4954C38" country="China" name="Type locality: Lushui Xian [Salween drainage]" stateProvince="Yunnan">Type locality: Lushui Xian [Salween drainage]</location>
,
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,
<collectingCountry id="6014F99BE6AA4E83B9EDEF068AF72662">China</collectingCountry>
.
<typeStatus id="3A13D106AB05826DD6AA09050BF5E1D9">Holotype</typeStatus>
:
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741097
</specimenCode>
.
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:
<collectionCode id="8F05B2289F8851EFC5250D087B940C85" collectionName="KIZ">KIZ</collectionCode>
(3).
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Synonomized with
<taxonomicName id="0C0E904C5C25EC2C9A29B5C369193741" LSID-ZBK="urn:lsid:zoobank.org:act:629CDB4F-2E7B-4F96-A91B-164A78A4171E" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" lsidName-ZBK="Glyptothorax zanaensis Wu, He &amp; Chu 1981:74 (English p. 79)" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="zanaensis">Glyptothorax zanaensis</taxonomicName>
by Mo &amp; Chu (1986).
</paragraph>
<paragraph id="3157594B01D29A5FD850851E7A4C78E3" pageNumber="57">
<taxonomicName id="F5546C31F1EBCBD4C043A807D1E472D2" LSID-ZBK="urn:lsid:zoobank.org:act:90C71FFE-BB6C-4642-91AA-61357BF95357" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" lsidName-ZBK="Glyptothorax rubermentus Li 1984:[English p. 88]" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="rubermentus">Glyptothorax rubermentus Li 1984b</taxonomicName>
: [English p. 88], fig. 8.
<materialsCitation id="F4A03063FFE6486D79316D067685F8DF" ID-GBIF-Occurrence="923901437" collectionCode="KIZ" country="China" location="Type locality [of Glyptothorax rubermentus Li]: Baoshan Xian [Salween drainage]" specimenCode="KIZ 749356 [and others]" stateProvince="Yunnan" typeStatus="Holotype, Paratypes">
<location id="C5235F90282464E11F9CC1EA9B52D9F6" LSID="urn:lsid:plazi:treatment:FE5BD2DF623CB3D7D42E2BB02EBFFF2E:C5235F90282464E11F9CC1EA9B52D9F6" country="China" name="Type locality: Baoshan Xian [Salween drainage]" stateProvince="Yunnan">Type locality: Baoshan Xian [Salween drainage]</location>
,
<collectingRegion id="EB70DDE403675350EABBEA212BF6787B">Yunnan</collectingRegion>
,
<collectingCountry id="8E90C7DD3CD2178225812D3169E9E7FC">China</collectingCountry>
.
<typeStatus id="A6D944E4018DF58BEBB931FB52DF89B6">Holotype</typeStatus>
:
<specimenCode id="6283B2799581CE29AC39521B52BEA9C0">
<collectionCode id="59E8981B9050B5650DA6E962D9ADEE30">KIZ</collectionCode>
749356
</specimenCode>
.
<typeStatus id="1EB83AF7CFC98BF3DA07B74E9FCBF37F">Paratypes</typeStatus>
:
<collectionCode id="FA0EE4EED3D87E5F7173EFFB36073A7B" collectionName="KIZ">KIZ</collectionCode>
(28).
</materialsCitation>
Synonomized with
<taxonomicName id="B7ABDCCEA7C1171A94A4676073FA8D71" LSID-ZBK="urn:lsid:zoobank.org:act:629CDB4F-2E7B-4F96-A91B-164A78A4171E" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" lsidName-ZBK="Glyptothorax zanaensis Wu, He &amp; Chu 1981:74 (English p. 79)" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="species" species="zanaensis">Glyptothorax zanaensis</taxonomicName>
by Mo &amp; Chu (1986).
</paragraph>
</subSubSection>
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<paragraph id="2E0C6D9D25DA3256DE81B13DE5218549" pageNumber="57">Distribution: Mekong and Salween drainages, Yunnan, China (Chu et al., 1999; Kottelat, 2001a).</paragraph>
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118
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<taxonomicName LSID="FE5BD536-109D-E15E-8FD5-F1D1FD5CEAF5" authority="(Soula, 2006)" baseAuthorityName="Soula" baseAuthorityYear="2006" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota louzadai" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="louzadai">Pelidnota louzadai (Soula, 2006)</taxonomicName>
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<taxonomicName authorityName="Soula" authorityYear="2006" class="Insecta" family="Scarabaeidae" genus="Strigidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strigidia louzadai" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="louzadai">Strigidia louzadai</taxonomicName>
Soula, 2006: 12, 55-56 [original combination].
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Soula" baseAuthorityYear="2006" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota louzadai" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="louzadai">Pelidnota louzadai</taxonomicName>
(Soula) [new combination by
<bibRefCitation author="Soula, M" journalOrPublisher="AECFT, Saintry" pageId="0" pageNumber="1" refId="B360" refString="Soula, M, 2009. Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot;Pelidnotina&quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry" title="Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot; Pelidnotina &quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale." year="2009">Soula 2009</bibRefCitation>
: 115].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
BRAZIL: Mato Grosso (
<bibRefCitation author="Soula, M" journalOrPublisher="AECFT, Saintry" pageId="0" pageNumber="1" refId="B358" refString="Soula, M, 2006. Les Coleopteres du Nouveau Monde. Volume 1: Rutelini 1. Revision des Pelidnotina 1 et des Lasiocalina. Une revision des genres Strigidia, Chalcoplethis, Epichalcoplethis, Sorocha, Lasiocala, Minilasiocala, Pseudochlorota, Homeochlorota, Pachacama (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: Pelidnotina et Lasiocalina). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry" title="Les Coleopteres du Nouveau Monde. Volume 1: Rutelini 1. Revision des Pelidnotina 1 et des Lasiocalina. Une revision des genres Strigidia, Chalcoplethis, Epichalcoplethis, Sorocha, Lasiocala, Minilasiocala, Pseudochlorota, Homeochlorota, Pachacama (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: Pelidnotina et Lasiocalina). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale." year="2006">Soula 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="types">
<paragraph pageId="0" pageNumber="1">Types.</paragraph>
<paragraph pageId="0" pageNumber="1">
The following specimens are deposited at CCECL. 1 ♂ holotype, 1 ♀ allotype: &quot;BRASIL - MT Faz.Sao Tiago 12.35S-56.20W XI-81//Holotype 2006
<taxonomicName authorityName="Soula" authorityYear="2006" class="Insecta" family="Scarabaeidae" genus="Strigidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strigidia louzadai" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="louzadai">
<emphasis italics="true" pageId="0" pageNumber="1">Strigidia louzadai</emphasis>
</taxonomicName>
S. Soula&quot; (47030437); &quot;BRASIL - MT Faz.Sao Tiago 12.35S-56.20W XI-81//Allotype 2006
<taxonomicName authorityName="Soula" authorityYear="2006" class="Insecta" family="Scarabaeidae" genus="Strigidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strigidia louzadai" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="louzadai">
<emphasis italics="true" pageId="0" pageNumber="1">Strigidia louzadai</emphasis>
</taxonomicName>
S. Soula&quot; (47030438). The genitalia are card-mounted underneath the male holotype. Box 4618663 SOULA.
</paragraph>
</subSubSection>
</treatment>
</document>