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data/FE/09/81/FE09815160B3B3B3AA634675D471D1BE.xml Normal file
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<mods:title id="35449C80031C03943C13A66F48BF814F">Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico</mods:title>
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<subSubSection id="1DD75ED19C3F9CABA25E2CD04E3AE9A6" pageId="742" pageNumber="743" type="nomenclature">
<paragraph id="15FBB7B9477F8D11C26598E805ED1FF6" pageId="742" pageNumber="743">
<taxonomicName id="BFB9BF73502BFD9296EE5C5E4B9AA821" ID-CoL="42KVN" LSID="FE098151-60B3-B3B3-AA63-4675D471D1BE" authority="Barr, 1982" authorityName="Barr" authorityYear="1982" class="Insecta" family="Carabidae" genus="Mexisphodrus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Mexisphodrus valverdensis" order="Coleoptera" pageId="742" pageNumber="743" phylum="Arthropoda" rank="species" species="valverdensis">Mexisphodrus valverdensis Barr, 1982</taxonomicName>
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<taxonomicName id="8F833E6BB568D8F588FBD40E8166FF79" authorityName="Barr" authorityYear="1982" class="Insecta" family="Carabidae" genus="Mexisphodrus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="742" pageNumber="743" phylum="Arthropoda" rank="species" species="valverdensis">Mexisphodrus valverdensis</taxonomicName>
Barr, 1982a: 177. Type locality: &quot;Ladder Cave, 25 km N[orth]W[est] Del Rio, Val Verde Co[unty], Texas&quot; (original citation). Holotype (♂) in AMNH.
</paragraph>
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<subSubSection id="F3314015040C8BA92F6763D14628F7EF" pageId="742" pageNumber="743" type="distribution">
<paragraph id="31F28C7169D768074D61D95D226CFD4D" pageId="742" pageNumber="743">Distribution.</paragraph>
<paragraph id="3855CA328E0EC6DE723D573B35682E18" pageId="742" pageNumber="743">This species is known from two nearby caves in southwestern Texas.</paragraph>
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<paragraph id="AC9D664181DC0EA351C1920127C1D90D" pageId="742" pageNumber="743">Records.</paragraph>
<paragraph id="8C7A5D596D482D12EA3DCB9969D421D6" pageId="742" pageNumber="743">
<emphasis id="E0C74143C6A656AF9A9917B8495D7DA6" bold="true" pageId="742" pageNumber="743">USA</emphasis>
: TX
</paragraph>
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<subSubSection id="7F2C5075926EFE37DD308B27D65113E1" pageId="742" pageNumber="743" type="note">
<paragraph id="763881D14C62848DF41901B167D12BE0" pageId="742" pageNumber="743">Note.</paragraph>
<paragraph id="7CB41D5A5BA9D2F7A17660033D72A566" pageId="742" pageNumber="743">
This species belongs to the
<emphasis id="089042718828A5CC276062A925A2BAEB" italics="true" pageId="742" pageNumber="743">boneti</emphasis>
species group which includes only the polymorphic
<taxonomicName id="DEC0172BEDA6467CA234B4F0592B56C1" baseAuthorityName="Bolivar y Pieltain &amp; Hendrichs" baseAuthorityYear="1964" class="Insecta" family="Carabidae" genus="Mexisphodrus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="742" pageNumber="743" phylum="Arthropoda" rank="species" species="boneti">
<emphasis id="BB66D79B3D6FCBA9EA6EECD7378C396B" italics="true" pageId="742" pageNumber="743">Mexisphodrus boneti</emphasis>
</taxonomicName>
(
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y Pieltain and Hendrichs) from Nuevo
<normalizedToken id="D1E31081CCD91D356729E3CEDB73F1B8" originalValue="León">Leon</normalizedToken>
and
<taxonomicName id="4CA15346F6BAA502AEA857FA37EFEC59" authorityName="Barr" authorityYear="1982" class="Insecta" family="Carabidae" genus="Mexisphodrus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="742" pageNumber="743" phylum="Arthropoda" rank="species" species="valverdensis">
<emphasis id="FE924B4FF549CD165B4A3FE8FC54B4B7" italics="true" pageId="742" pageNumber="743">Mexisphodrus valverdensis</emphasis>
</taxonomicName>
Barr.
</paragraph>
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</treatment>
</document>

136
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, Ban Ko Muang &amp; H. Malicky leg. (
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, Jiří Kolibáč leg. (
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],
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a.s.l., small pools in secondary forest,
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, Balke &amp; Hendrich leg. (
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, further specimens in coll. Hendrich, Berlin).
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This taxon was described from eastern Sumatra. Although
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considered it as a subspecies and presented specimens from
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with median lobe intermediate to
<emphasis id="44D4EACBFFF9E93BFBE1E82DFF0A10EA" italics="true" pageId="6" pageNumber="57">L. t. lituratus</emphasis>
, its status remains unresolved. The currently known distribution of
<emphasis id="44D4EACBFFF9E93BFC1DE8CDFB9C10EA" box="[961,1126,917,941]" italics="true" pageId="6" pageNumber="57">L. t. leptogonus</emphasis>
and
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does not support the hypothesis of geographical subspecies (see also under
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). We believe that
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represents a separate species, but more material is desirable to answer the question. First records from
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,
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and
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.
</paragraph>
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[
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], Rawa Aopa Nat. Park, Aopa vill.,
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, M. Štrba &amp; I. Jeniš leg. (
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).
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Widely distributed in the Oriental and Australian Region. The nominotypical subspecies was recorded from
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, New
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and northern
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(
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, BALKE et al. 1997). First record from Sulawesi.
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Sharp, 1882
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1
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, Dolok Merangir,
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99°11′
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,
<date id="021E1019FFF9E93BFC2BEF37FB9817C0" box="[1015,1122,1135,1159]" pageId="6" pageNumber="57" value="1991-02-21">21.ii.1991</date>
, Malicky leg. (
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).
</paragraph>
<paragraph id="761F36D9FFF9E93BFF6AEFF7FD281640" blockId="6.[151,1127,1132,1287]" pageId="6" pageNumber="57">
This subspecies was recorded from continental Southeast Asia –
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,
<collectingCountry id="0EB77649FFF9E93BFC7FEFF7FBFE1780" box="[931,1028,1199,1223]" name="Thailand" pageId="6" pageNumber="57">Thailand</collectingCountry>
,
<collectingCountry id="0EB77649FFF9E93BFBD1EFF7FB9D1780" box="[1037,1127,1199,1223]" name="Vietnam" pageId="6" pageNumber="57">Vietnam</collectingCountry>
and
<collectingCountry id="0EB77649FFF9E93BFF18EF97FECB17A0" box="[196,305,1231,1255]" name="Cambodia" pageId="6" pageNumber="57">Cambodia</collectingCountry>
(
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,
<bibRefCitation id="12314B28FFF9E93BFE27EF88FD8017A0" author="HAJEK J." box="[507,634,1231,1255]" pageId="6" pageNumber="57" pagination="115 - 123" refId="ref3833" refString="HAJEK J. 2003: Dytiscidae: III. The genus Laccophilus Leach in China (Coleoptera). Pp. 115 - 123. In: JACH M. A. &amp; JI L. (eds.): Water Beetles of China, Vol. III. Zoologisch-Botanische Gesellschaft in Osterreich and Wiener Coleopterologenverein, Wien, vi + 572 pp." type="book chapter" year="2003">HÁJEK 2003</bibRefCitation>
). The present finding from
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represents the first record from
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and the Sunda Islands.
</paragraph>
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104
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Aubé, 1838
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2 JJ
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, Nakai env.,
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a.s.l.,
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).
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Widely distributed Oriental species, known from
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,
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,
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and
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(
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134
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, Ban Lok,
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,
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, J. Mlíkovský leg. (
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). 1 J,
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, Taleban [= Thale Ban,
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] (See),
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<geoCoordinate id="1394501EFFFAE938FFADEACBFF2F12EC" box="[113,213,402,427]" degrees="100" direction="east" minutes="10" orientation="longitude" pageId="5" pageNumber="56" precision="925" value="100.166664">100°10′E</geoCoordinate>
,
<quantity id="B1589B3CFFFAE938FF02EACBFEE812EC" box="[222,274,403,427]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="56" unit="m" value="80.0">80 m</quantity>
a.s.l.,
<date id="021E1019FFFAE938FE8CEACBFE3D12EC" box="[336,455,403,427]" pageId="5" pageNumber="56" value="1993-05-07" valueMax="1993-05-08" valueMin="1993-05-07">7.-8.v.1993</date>
, Malicky leg. (
<collectionCode id="10B1AE1CFFFAE938FD81EACBFD5712EC" box="[605,685,403,427]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="5" pageNumber="56">NMPC</collectionCode>
). 1J,
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</emphasis>
,
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, Langkawi, Pantai Kok,
<date id="021E1019FFFAE938FF2AEAEBFEA6128C" box="[246,348,435,459]" pageId="5" pageNumber="56" value="1992-01-30">30.i.1992</date>
, M. A. Jäch leg. (14) (
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).
</paragraph>
<paragraph id="761F36D9FFFAE938FF4CEA8BFDB8112C" blockId="5.[113,1090,336,619]" pageId="5" pageNumber="56">
Previously recorded from border area between the Palaearctic and Oriental Regions –
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,
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, northern
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,
<collectingCountry id="0EB77649FFFAE938FE6EEAABFDCA114C" box="[434,560,499,523]" name="Bangladesh" pageId="5" pageNumber="56">Bangladesh</collectingCountry>
,
<collectingCountry id="0EB77649FFFAE938FDE5EAACFD5B114B" box="[569,673,500,524]" name="Myanmar" pageId="5" pageNumber="56">Myanmar</collectingCountry>
, southern
<collectingCountry id="0EB77649FFFAE938FCD5EAABFCB1114C" box="[777,843,499,523]" name="China" pageId="5" pageNumber="56">China</collectingCountry>
, Ryukyu Islands, northern
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, northern
<collectingCountry id="0EB77649FFFAE938FE82E94CFE6B116C" box="[350,401,532,555]" name="Laos" pageId="5" pageNumber="56">Laos</collectingCountry>
and northern
<collectingCountry id="0EB77649FFFAE938FDC2E94BFD8D116C" box="[542,631,531,555]" name="Vietnam" pageId="5" pageNumber="56">Vietnam</collectingCountry>
(
<bibRefCitation id="12314B28FFFAE938FD58E94BFCCB116C" author="BRANCUCCI M." box="[644,817,531,555]" pageId="5" pageNumber="56" pagination="241 - 426" refId="ref3760" refString="BRANCUCCI M. 1983: Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey 31 / 32: 241 - 426." type="journal article" year="1983">BRANCUCCI 1983</bibRefCitation>
,
<bibRefCitation id="12314B28FFFAE938FCE6E94CFC42116C" author="HAJEK J." box="[826,952,531,555]" pageId="5" pageNumber="56" pagination="115 - 123" refId="ref3833" refString="HAJEK J. 2003: Dytiscidae: III. The genus Laccophilus Leach in China (Coleoptera). Pp. 115 - 123. In: JACH M. A. &amp; JI L. (eds.): Water Beetles of China, Vol. III. Zoologisch-Botanische Gesellschaft in Osterreich and Wiener Coleopterologenverein, Wien, vi + 572 pp." type="book chapter" year="2003">HÁJEK 2003</bibRefCitation>
). The present records extend the distribution area of
<taxonomicName id="B1A04D5AFFFAE938FDDAE96CFD79110C" authorityName="Boheman" authorityYear="1858" box="[518,643,563,587]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="44D4EACBFFFAE938FDDAE96CFD79110C" box="[518,643,563,587]" italics="true" pageId="5" pageNumber="56">L. chinensis</emphasis>
</taxonomicName>
southwards in continental Southeast Asia. First records from
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and
<collectingCountry id="0EB77649FFFAE938FE05E90BFDC4112C" box="[473,574,595,619]" name="Malaysia" pageId="5" pageNumber="56">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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98
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<emphasis id="44D4EACBFFFAE938FEBEEFF0FD681786" bold="true" box="[354,658,1192,1217]" italics="true" pageId="5" pageNumber="56">Laccophilus punctatissimus</emphasis>
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2 JJ
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,
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Division, Chatthin Wildlife Sanctuary, Kinsan Camp,
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,
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a.s.l.,
<date id="021E1019FFFAE938FD3AEE5BFCA8165C" box="[742,850,1283,1307]" pageId="5" pageNumber="56" value="1998-10-13">13.x.1998</date>
, H. Schillhammer leg. (14) (
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,
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).
</paragraph>
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Described from
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,
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,
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and
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(
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). First record from
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119
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<emphasis id="44D4EACBFFFAE938FEF2E9CBFE0911EC" bold="true" box="[302,499,659,683]" italics="true" pageId="5" pageNumber="56">Laccophilus chini</emphasis>
<bibRefCitation id="12314B28FFFAE938FE26E9CBFC7E11EC" author="BALKE M. &amp; MAZZOLDI P. &amp; HENDRICH L." box="[506,900,659,683]" pageId="5" pageNumber="56" pagination="71 - 77" refId="ref3715" refString="BALKE M., MAZZOLDI P. &amp; HENDRICH L. 1998: Two new Laccophilus species (Coleoptera: Dytiscidae) from Southeast Asia, and notes on other species of the genus. Raffles Bulletin of Zoology 46: 71 - 77." type="journal article" year="1998">Balke, Mazzoldi &amp; Hendrich, 1998</bibRefCitation>
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<emphasis id="44D4EACBFFFAE938FFADE996FECC11A1" bold="true" box="[113,310,718,742]" pageId="5" pageNumber="56">Material studied.</emphasis>
3 JJ,
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<collectingCountry id="0EB77649FFFAE938FE57E996FE2911A1" box="[395,467,718,742]" name="Laos" pageId="5" pageNumber="56">LAOS</collectingCountry>
</emphasis>
,
<collectingRegion id="B464F83BFFFAE938FE3FE996FD5011A2" box="[483,682,718,741]" country="Laos" name="Khammouan" pageId="5" pageNumber="56">KHAMMOUAN Prov.</collectingRegion>
, Nakai env.,
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, 500- -
<quantity id="B1589B3CFFFAE938FFA5E9B6FF451042" box="[121,191,750,774]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="56" unit="m" value="600.0">600 m</quantity>
a.s.l.,
<date id="021E1019FFFAE938FEDBE9B6FE501041" box="[263,426,750,774]" pageId="5" pageNumber="56" value="2001-05-22" valueMax="2001-06-08" valueMin="2001-05-22">22.v.-8.vi.2001</date>
, E. Jendek &amp; O. Šauša leg. (
<collectionCode id="10B1AE1CFFFAE938FD3FE9B6FCCE1041" box="[739,820,750,774]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="5" pageNumber="56">NMPC</collectionCode>
). 1 J
<specimenCount id="60A6FD50FFFAE938FCA4E9B6FC641040" box="[888,926,747,775]" pageId="5" pageNumber="56" type="female">1 ♀</specimenCount>
,
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<collectingCountry id="0EB77649FFFAE938FC73E9B6FC0D1041" box="[943,1015,750,774]" name="Laos" pageId="5" pageNumber="56">LAOS</collectingCountry>
</emphasis>
, VIEN-
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Prov.,
<quantity id="B1589B3CFFFAE938FEDEE856FEBF1061" box="[258,325,782,806]" metricMagnitude="4" metricUnit="m" metricValue="5.5" pageId="5" pageNumber="56" unit="km" value="55.0">55 km</quantity>
NE Vientianne, Lao Pako env.,
<quantity id="B1589B3CFFFAE938FD40E856FD1A1062" box="[668,736,782,806]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="56" unit="m" value="200.0">200 m</quantity>
a.s.l.,
<date id="021E1019FFFAE938FCFAE856FC5A1061" box="[806,928,782,806]" pageId="5" pageNumber="56" value="2004-05-01" valueMax="2004-05-04" valueMin="2004-05-01">1.-4.v.2004</date>
, J. Bezděk leg. (
<collectionCode id="10B1AE1CFFFAE938FFA5E876FF311001" box="[121,203,814,838]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="5" pageNumber="56">NMPC</collectionCode>
).
</paragraph>
<paragraph id="761F36D9FFFAE938FF4CE816FCDD10E2" blockId="5.[113,1089,715,934]" pageId="5" pageNumber="56">
The species was described from southern
<collectingCountry id="0EB77649FFFAE938FD95E816FD591021" box="[585,675,846,870]" name="Vietnam" pageId="5" pageNumber="56">Vietnam</collectingCountry>
and
<collectingCountry id="0EB77649FFFAE938FD0AE816FCCD1021" box="[726,823,846,870]" name="Malaysia" pageId="5" pageNumber="56">Malaysia</collectingCountry>
(
<collectingRegion id="B464F83BFFFAE938FC99E816FC6C1021" box="[837,918,846,870]" country="Malaysia" name="Pahang" pageId="5" pageNumber="56">Pahang</collectingRegion>
) by
<bibRefCitation id="12314B28FFFAE938FC18E816FF4110C1" author="BALKE M. &amp; MAZZOLDI P. &amp; HENDRICH L." pageId="5" pageNumber="56" pagination="71 - 77" refId="ref3715" refString="BALKE M., MAZZOLDI P. &amp; HENDRICH L. 1998: Two new Laccophilus species (Coleoptera: Dytiscidae) from Southeast Asia, and notes on other species of the genus. Raffles Bulletin of Zoology 46: 71 - 77." type="journal article" year="1998">BALKE et al. (1998)</bibRefCitation>
. We could not study the specimen they recorded from
<collectingRegion id="B464F83BFFFAE938FCCDE836FCAE10C1" box="[785,852,878,902]" country="Malaysia" name="Sabah" pageId="5" pageNumber="56">Sabah</collectingRegion>
, but we expect that it represents
<taxonomicName id="B1A04D5AFFFAE938FF38E8D7FEA110E1" authority="Hájek &amp; Šťastný, 2005" authorityName="Hájek &amp; Šťastný" authorityYear="2005" box="[228,347,910,934]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="uncletan" status="sp. nov.">
<emphasis id="44D4EACBFFFAE938FF38E8D7FEA110E1" box="[228,347,910,934]" italics="true" pageId="5" pageNumber="56">L. uncletan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5FE757B0FFFAE938FEBEE8D6FE4F10E2" box="[354,437,910,933]" pageId="5" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
First record of
<taxonomicName id="B1A04D5AFFFAE938FD86E8D7FD5110E1" authorityName="Balke, Mazzoldi &amp; Hendrich" authorityYear="1998" box="[602,683,910,934]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="chini">
<emphasis id="44D4EACBFFFAE938FD86E8D7FD5110E1" box="[602,683,910,934]" italics="true" pageId="5" pageNumber="56">L. chini</emphasis>
</taxonomicName>
from
<collectingCountry id="0EB77649FFFAE938FD31E8D6FCDE10E2" box="[749,804,910,933]" name="Laos" pageId="5" pageNumber="56">Laos</collectingCountry>
.
</paragraph>
</subSubSection>
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88
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<emphasis id="44D4EACBFFFAE938FE5EEEFBFD9616FC" bold="true" box="[386,620,1443,1467]" italics="true" pageId="5" pageNumber="56">Laccophilus ritsemae</emphasis>
Régimbart, 1880
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1 J,
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</emphasis>
, LOMBOK, Ampenan [
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], at light,
<date id="021E1019FFFAE938FFADEEA6FF1F1551" box="[113,229,1534,1558]" pageId="5" pageNumber="56" value="1984-06-15">15.vi.1984</date>
, G. De Rougemont leg. (
<collectionCode id="10B1AE1CFFFAE938FE33EEA6FDC31551" box="[495,569,1534,1558]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="56">NMW</collectionCode>
).
</paragraph>
<paragraph id="761F36D9FFF9E93BFF6AEB87FEC81270" blockId="6.[151,1127,223,311]" pageId="6" pageNumber="57">
Distributed in the Malayan Peninsula and the western part of Indonesian Archipelago –
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,
<collectingRegion id="B464F83BFFF9E93BFEDCEA58FEC81250" box="[256,306,256,279]" country="Indonesia" name="Jawa Timur" pageId="6" pageNumber="57">Java</collectingRegion>
, Borneo and
<collectingRegion id="B464F83BFFF9E93BFE19EBA7FE0B1250" box="[453,497,255,279]" country="Indonesia" name="Bali" pageId="6" pageNumber="57">Bali</collectingRegion>
(
<bibRefCitation id="12314B28FFF9E93BFDDFEBA7FD4F1250" author="BRANCUCCI M." box="[515,693,255,279]" pageId="6" pageNumber="57" pagination="241 - 426" refId="ref3760" refString="BRANCUCCI M. 1983: Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey 31 / 32: 241 - 426." type="journal article" year="1983">BRANCUCCI 1983</bibRefCitation>
,
<bibRefCitation id="12314B28FFF9E93BFD1FEA58FC2E1250" author="HENDRICH L. &amp; BALKE M." box="[707,980,255,279]" pageId="6" pageNumber="57" pagination="29 - 56" refId="ref3898" refString="HENDRICH L. &amp; BALKE M. 1995: Die Schwimmkafer der Sunda-Insel Bali: Faunistik, Taxonomie, Okologie, Besiedlungsgeschichte und Beschreibung von vier neuen Arten (Insecta: Coleoptera: Dytiscidae). Faunistische Abhandlungen des Museums fur Tierkunde in Dresden 20: 29 - 56." type="journal article" year="1995">HENDRICH &amp; BALKE 1995</bibRefCitation>
). First record from Lombok.
</paragraph>
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234
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<document id="6C9067EA46EB93A443AA1B2EFC3B88C8" ID-DOI="10.5281/zenodo.5176618" ID-ISSN="0374-1036" ID-Zenodo-Dep="5176618" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628601973277" checkinUser="carolina" docAuthor="Hájek, Jiří &amp; Šťastný, Jaroslav" docDate="2005" docId="FE0987CFFFFDE93EFE06EF87FEC11550" docLanguage="en" docName="ActaEntMusNatPra.45.51-58.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 45" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Laccophilus uncletan Hájek &amp; Šťastný 2005, sp. nov." docType="treatment" docVersion="3" lastPageNumber="54" masterDocId="0230FFB7FFFFE93DFFDCEB58FFFA1347" masterDocTitle="Two new species of Laccophilus Leach, 1815 (Coleoptera: Dytiscidae) from India and Borneo with notes on other species of the genus" masterLastPageNumber="58" masterPageNumber="51" pageNumber="53" updateTime="1699047269640" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="3C98FD8625F68B9EB68D348E83A769B9">Two new species of Laccophilus Leach, 1815 (Coleoptera: Dytiscidae) from India and Borneo with notes on other species of the genus</mods:title>
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<mods:name id="74439810FFC6312312B7D37CC19A3E4B" type="personal">
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<mods:namePart id="3041B689C3FC015328834F9957C6F209">Hájek, Jiří</mods:namePart>
<mods:affiliation id="A1913C77CE2CD28463A29CB7EBC967E1">Department of Entomology, National Museum, Kunratice 1, CZ- 148 00 Praha 4, Czech Republic; e-mail: jiri _ hajek @ nm. cz</mods:affiliation>
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<mods:namePart id="AB6AE19C87E66D01A42F7A4989843357">Šťastný, Jaroslav</mods:namePart>
<mods:affiliation id="4D7C2856BD6FD740E5C25720F16D51A4">Kosmonautů 359, CZ- 460 05 Liberec, Czech Republic; e-mail: stastnyj @ jergym. hiedu. cz</mods:affiliation>
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<mods:title id="088E85DA07765F59375D5873093618A1">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="8FD8EE6ED293C7EEF8585FD5BA8F70E0">
<mods:date id="87FF21EECC15CD10EB25848919184514">2005</mods:date>
<mods:detail id="C32AFB8A2EDE411F6C7E887F30AE2300" type="pubDate">
<mods:number id="4D50D90A06A6BF8F3EE516998C29BEAE">2005-12-31</mods:number>
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<mods:number id="1FB880499DDA60B48DFA8856E27839A1">45</mods:number>
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<mods:start id="60CC192A406A5F9726F528FFB73328F1">51</mods:start>
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<paragraph id="761F36D9FFFDE93FFE06EF87FCDE17B0" blockId="2.[474,804,1247,1271]" box="[474,804,1247,1271]" pageId="2" pageNumber="53">
<heading id="2D5781B5FFFDE93FFE06EF87FCDE17B0" bold="true" box="[474,804,1247,1271]" centered="true" fontSize="8" level="2" pageId="2" pageNumber="53" reason="3">
<emphasis id="44D4EACBFFFDE93FFE06EF87FCDE17B0" bold="true" box="[474,804,1247,1271]" pageId="2" pageNumber="53">
<taxonomicName id="B1A04D5AFFFDE93FFE06EF87FD3C17B0" ID-CoL="3RMCH" authority="Hájek &amp; Šťastný, 2005" authorityName="Hájek &amp; Šťastný" authorityYear="2005" box="[474,710,1247,1271]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="uncletan" status="sp. nov.">
<emphasis id="44D4EACBFFFDE93FFE06EF87FD3C17B0" bold="true" box="[474,710,1247,1271]" italics="true" pageId="2" pageNumber="53">Laccophilus uncletan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5FE757B0FFFDE93FFD12EF87FCDE17B0" box="[718,804,1247,1271]" pageId="2" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="3EBA6552FFFDE93FFD8CEFA7FD541655" box="[592,686,1279,1298]" pageId="2" pageNumber="53" type="description">
<paragraph id="761F36D9FFFDE93FFD8CEFA7FD541655" blockId="2.[151,742,1279,1333]" box="[592,686,1279,1298]" pageId="2" pageNumber="53">
(
<figureCitation id="EE9B2A5CFFFDE93FFD8AEFA7FD5D1655" box="[598,679,1279,1298]" captionStart="Figs 1-4. 1, 3" captionStartId="4.[151,188,1486,1505]" captionTargetBox="[167,1112,215,1451]" captionTargetId="figure-9@4.[167,1112,215,1451]" captionTargetPageId="4" captionText="Figs 1-4. 1, 3 – Laccophilus boukali sp. nov. 1 – habitus; 3 – median lobe in lateral view. 2, 4 – L. uncletan sp. nov. 2 – habitus; 4 – aedeagus (a – median lobe in lateral view; b – apex of median lobe in dorsal view; c – left paramere; d – right paramere). Scale bars: 2 mm (Figs 1-2); 0.5 mm (Figs 3-4)." figureDoi="http://doi.org/10.5281/zenodo.5176620" httpUri="https://zenodo.org/record/5176620/files/figure.png" pageId="2" pageNumber="53">Figs. 2, 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3EBA6552FFFDE93FFF4BEE7AFD1C1672" box="[151,742,1314,1333]" pageId="2" pageNumber="53" type="reference_group">
<paragraph id="761F36D9FFFDE93FFF4BEE7AFD1C1672" blockId="2.[151,742,1279,1333]" box="[151,742,1314,1333]" pageId="2" pageNumber="53">
<treatmentCitationGroup id="56B011F7FFFDE93FFF4BEE7AFD1C1672" box="[151,742,1314,1333]" pageId="2" pageNumber="53">
?
<treatmentCitation id="F70110C8FFFDE93FFF7DEE7AFD691672" author="BALKE M. &amp; MAZZOLDI P. &amp; HENDRICH L." box="[161,659,1314,1333]" page="72" pageId="2" pageNumber="53" year="1998">
<taxonomicName id="B1A04D5AFFFDE93FFF7DEE7AFD691672" ID-CoL="3RM63" authority="Balke, Mazzoldi &amp; Hendrich, 1998: 72" authorityName="Balke, Mazzoldi &amp; Hendrich" authorityPageNumber="72" authorityYear="1998" box="[161,659,1314,1333]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="chini">
<emphasis id="44D4EACBFFFDE93FFF7DEE7AFEC31672" box="[161,313,1314,1333]" italics="true" pageId="2" pageNumber="53">Laccophilus chini</emphasis>
<bibRefCitation id="12314B28FFFDE93FFE9CEE7AFD691672" author="BALKE M. &amp; MAZZOLDI P. &amp; HENDRICH L." box="[320,659,1314,1333]" pageId="2" pageNumber="53" pagination="71 - 77" refId="ref3715" refString="BALKE M., MAZZOLDI P. &amp; HENDRICH L. 1998: Two new Laccophilus species (Coleoptera: Dytiscidae) from Southeast Asia, and notes on other species of the genus. Raffles Bulletin of Zoology 46: 71 - 77." type="journal article" year="1998">Balke, Mazzoldi &amp; Hendrich, 1998: 72</bibRefCitation>
</taxonomicName>
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(partim).
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</paragraph>
</subSubSection>
<subSubSection id="3EBA6552FFFDE93FFF4BEE0FFD9C1568" pageId="2" pageNumber="53" type="materials_examined">
<paragraph id="761F36D9FFFDE93FFF4BEE0FFC6A1629" blockId="2.[151,1127,1367,1583]" box="[151,912,1367,1391]" pageId="2" pageNumber="53">
<emphasis id="44D4EACBFFFDE93FFF4BEE0FFED41628" bold="true" box="[151,302,1367,1391]" pageId="2" pageNumber="53">
<typeStatus id="A91B887BFFFDE93FFF4BEE0FFF341628" box="[151,206,1367,1391]" pageId="2" pageNumber="53">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="0EB77649FFFDE93FFEE8EE0FFE621628" box="[308,408,1367,1391]" name="Malaysia" pageId="2" pageNumber="53">Malaysia</collectingCountry>
,
<collectingRegion id="B464F83BFFFDE93FFE7FEE0FFE1C1628" box="[419,486,1367,1391]" country="Malaysia" name="Sabah" pageId="2" pageNumber="53">Sabah</collectingRegion>
, Kinabatangan river, Uncle Tan’s camp.
</paragraph>
<paragraph id="761F36D9FFFDE93FFF4BEE2FFD9C1568" blockId="2.[151,1127,1367,1583]" pageId="2" pageNumber="53">
<emphasis id="44D4EACBFFFDE93FFF4BEE2FFEC616C8" bold="true" box="[151,316,1399,1423]" pageId="2" pageNumber="53">Type material.</emphasis>
<materialsCitation id="C6C83C84FFFDE93FFE9EEE2FFD9116E8" collectorName="J. Stastny" country="Malaysia" location="Sabah" pageId="2" pageNumber="53" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus id="A91B887BFFFDE93FFE9EEE2FFE5316CA" box="[322,425,1399,1422]" pageId="2" pageNumber="53" type="holotype">HOLOTYPE</typeStatus>
: J, ‘
<collectingCountry id="0EB77649FFFDE93FFE06EE2FFDB816C8" box="[474,578,1399,1423]" name="Malaysia" pageId="2" pageNumber="53">Malaysia</collectingCountry>
,
<collectingRegion id="B464F83BFFFDE93FFD91EE2FFD7416C8" box="[589,654,1399,1423]" country="Malaysia" name="Sabah" pageId="2" pageNumber="53">Sabah</collectingRegion>
/ Kinabatangan riv.[er] / 8.-15.6.[20]03, Uncle Tan’s / camp,
<collectorName id="DB55530FFFFDE93FFE84EECFFE3916E8" box="[344,451,1425,1455]" pageId="2" pageNumber="53">J. Šťastný</collectorName>
lgt. [printed] //
</materialsCitation>
<materialsCitation id="C6C83C84FFFDE93FFDAFEECFFC961688" collectionCode="NMPC" country="Malaysia" determinerName="J. Hajek &amp; J. Stastny" pageId="2" pageNumber="53" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus id="A91B887BFFFDE93FFDAFEECFFD0516E8" box="[627,767,1431,1455]" pageId="2" pageNumber="53" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="B1A04D5AFFFDE93FFCCBEECFFC3416E8" authorityName="Leach" authorityYear="1815" box="[791,974,1431,1455]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="genus">LACCOPHILUS</taxonomicName>
/ uncletan
<taxonomicNameLabel id="5FE757B0FFFDE93FFB94EECFFF3F1689" pageId="2" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
/
<determinerName id="83C3DA1DFFFDE93FFF07EEEFFEC91688" box="[219,307,1463,1487]" pageId="2" pageNumber="53">J. Hájek</determinerName>
&amp;
<determinerName id="83C3DA1DFFFDE93FFE84EEEFFE381688" box="[344,450,1457,1487]" pageId="2" pageNumber="53">J. Šťastný</determinerName>
det. 2005’ [printed, red label] (
<collectionCode id="10B1AE1CFFFDE93FFCC8EEEFFC9F1688" box="[788,869,1463,1487]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="53">NMPC</collectionCode>
)
</materialsCitation>
.
<typeStatus id="A91B887BFFFDE93FFCA4EEEFFC1E168A" box="[888,996,1463,1486]" pageId="2" pageNumber="53" type="paratype">PARATYPES</typeStatus>
: 2JJ
<specimenCount id="60A6FD50FFFDE93FFBF1EEEFFBA21697" box="[1069,1112,1460,1488]" pageId="2" pageNumber="53" type="female">5♀♀</specimenCount>
, same label data as
<typeStatus id="A91B887BFFFDE93FFEBAEE8FFE3916A8" box="[358,451,1495,1519]" pageId="2" pageNumber="53" type="holotype">holotype</typeStatus>
(
<collectionCode id="10B1AE1CFFFDE93FFE0FEE8FFDE816A8" box="[467,530,1495,1519]" pageId="2" pageNumber="53">JSCL</collectionCode>
,
<collectionCode id="10B1AE1CFFFDE93FFDC3EE8FFD9716A8" box="[543,621,1495,1519]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="53">NMPC</collectionCode>
,
<collectionCode id="10B1AE1CFFFDE93FFDA5EE8FFD3816A8" box="[633,706,1495,1519]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="53">NMW</collectionCode>
). Each
<typeStatus id="A91B887BFFFDE93FFCCFEE8FFC9416A8" box="[787,878,1495,1519]" pageId="2" pageNumber="53" type="paratype">paratype</typeStatus>
is provided with a red label similar to that of the
<typeStatus id="A91B887BFFFDE93FFE68EEAFFDE91548" box="[436,531,1527,1551]" pageId="2" pageNumber="53" type="holotype">holotype</typeStatus>
, except ‘PARATYPE’ instead of ‘HOLOTYPE’, the respective sex symbol and collection number.
</paragraph>
</subSubSection>
<subSubSection id="3EBA6552FFFCE93EFFADEB87FE951710" pageId="3" pageNumber="54" type="description">
<paragraph id="761F36D9FFFCE93EFFADEB87FDDE1270" blockId="3.[113,1090,223,1559]" pageId="3" pageNumber="54">
<emphasis id="44D4EACBFFFCE93EFFADEB87FF0113B0" bold="true" box="[113,251,223,247]" pageId="3" pageNumber="54">Description.</emphasis>
Oval and oblong; head testaceous, pronotum testaceous with blackish basal band and two subbasal spots, elytra blackish with testaceous pattern as depicted in
<figureCitation id="EE9B2A5CFFFCE93EFC26EBA7FBC71250" box="[1018,1085,255,279]" captionStart="Figs 1-4. 1, 3" captionStartId="4.[151,188,1486,1505]" captionTargetBox="[167,1112,215,1451]" captionTargetId="figure-9@4.[167,1112,215,1451]" captionTargetPageId="4" captionText="Figs 1-4. 1, 3 – Laccophilus boukali sp. nov. 1 – habitus; 3 – median lobe in lateral view. 2, 4 – L. uncletan sp. nov. 2 – habitus; 4 – aedeagus (a – median lobe in lateral view; b – apex of median lobe in dorsal view; c – left paramere; d – right paramere). Scale bars: 2 mm (Figs 1-2); 0.5 mm (Figs 3-4)." figureDoi="http://doi.org/10.5281/zenodo.5176620" httpUri="https://zenodo.org/record/5176620/files/figure.png" pageId="3" pageNumber="54">Fig. 2</figureCitation>
. Ventral parts and appendages testaceous.
</paragraph>
<paragraph id="761F36D9FFFCE93EFF4CEA18FCD61230" blockId="3.[113,1090,223,1559]" pageId="3" pageNumber="54">
Measurements. Body length
<quantity id="B1589B3CFFFCE93EFE1FEA67FDA21210" box="[451,600,319,343]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="3.8" metricValueMin="3.5" pageId="3" pageNumber="54" unit="mm" value="3.65" valueMax="3.8" valueMin="3.5">3.50-3.80 mm</quantity>
(mean value =
<quantity id="B1589B3CFFFCE93EFCDDEA67FC991210" box="[769,867,319,343]" metricMagnitude="-3" metricUnit="m" metricValue="3.62" pageId="3" pageNumber="54" unit="mm" value="3.62">3.62 mm</quantity>
,
<typeStatus id="A91B887BFFFCE93EFCB2EA67FC311210" box="[878,971,319,343]" pageId="3" pageNumber="54" type="holotype">holotype</typeStatus>
<quantity id="B1589B3CFFFCE93EFC0EEA67FBCF1210" box="[978,1077,319,343]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" pageId="3" pageNumber="54" unit="mm" value="3.65">3.65 mm</quantity>
), width
<quantity id="B1589B3CFFFCE93EFF68EA07FEB31230" box="[180,329,351,375]" metricMagnitude="-3" metricUnit="m" metricValue="1.89" metricValueMax="1.95" metricValueMin="1.83" pageId="3" pageNumber="54" unit="mm" value="1.89" valueMax="1.95" valueMin="1.83">1.83-1.95 mm</quantity>
(mean value =
<quantity id="B1589B3CFFFCE93EFE33EA07FDAA1230" box="[495,592,351,375]" metricMagnitude="-3" metricUnit="m" metricValue="1.88" pageId="3" pageNumber="54" unit="mm" value="1.88">1.88 mm</quantity>
,
<typeStatus id="A91B887BFFFCE93EFD86EA07FD4D1230" box="[602,695,351,375]" pageId="3" pageNumber="54" type="holotype">holotype</typeStatus>
<quantity id="B1589B3CFFFCE93EFD61EA07FCDA1230" box="[701,800,351,375]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="3" pageNumber="54" unit="mm" value="1.85">1.85 mm</quantity>
).
</paragraph>
<paragraph id="761F36D9FFFCE93EFF4CEA27FBE71290" blockId="3.[113,1090,223,1559]" pageId="3" pageNumber="54">Head. Coloration testaceous with indistinct darker spots. Reticulation double, composed of almost imperceptible polygonal meshes, and microreticulation of fine isodiametric meshes. Frons between eyes and clypeus with several large punctures. Mouthparts testaceous.</paragraph>
<paragraph id="761F36D9FFFCE93EFF4CEAB8FCD91110" blockId="3.[113,1090,223,1559]" pageId="3" pageNumber="54">Pronotum. Coloration testaceous with thin basal blackish band and two subbasal blackish spots. Reticulation double, composed of slightly impressed large polygonal meshes and microreticulation of fine isodiametric meshes similar to that of head. Anterior margin, basal margin and posterior angles with numerous setigerous punctures.</paragraph>
<paragraph id="761F36D9FFFCE93EFF4CE907FCD011B0" blockId="3.[113,1090,223,1559]" pageId="3" pageNumber="54">Elytra. Coloration blackish with testaceous markings comprising subbasal transverse sinuous band, mediolateral spot, subapical transverse sinuous band and apical spot. Sutural part of elytra blackish; in some specimens all testaceous markings connected in epipleural part. Reticulation on whole elytra distinctly double, similar to that of pronotum. Each elytron with sutural, discal and lateral row of large serial setigerous punctures.</paragraph>
<paragraph id="761F36D9FFFCE93EFF4CE9A7FF341030" blockId="3.[113,1090,223,1559]" pageId="3" pageNumber="54">Ventral side. Testaceous and microreticulated. Prosternal process long and acute. Abdominal sternites laterally with oblique grooves and several large punctures. Posterior margin of anal sternite prominent medially with cluster of setigerous punctures, and moderately incised laterally.</paragraph>
<paragraph id="761F36D9FFFCE93EFF4CE8D8FF481090" blockId="3.[113,1090,223,1559]" pageId="3" pageNumber="54">Legs. Completely testaceous. Male tarsomeres 1-3 of fore and middle legs only weakly dilated with microtrichia on ventral side. Claws of fore as well as middle tarsi simple and equal.</paragraph>
<paragraph id="761F36D9FFFCE93EFF4CE887FCBB10B0" blockId="3.[113,1090,223,1559]" box="[144,833,991,1015]" pageId="3" pageNumber="54">
Male genitalia. Shape of median lobe and parameres as in
<figureCitation id="EE9B2A5CFFFCE93EFD20E887FCC710B0" box="[764,829,991,1015]" captionStart="Figs 1-4. 1, 3" captionStartId="4.[151,188,1486,1505]" captionTargetBox="[167,1112,215,1451]" captionTargetId="figure-9@4.[167,1112,215,1451]" captionTargetPageId="4" captionText="Figs 1-4. 1, 3 – Laccophilus boukali sp. nov. 1 – habitus; 3 – median lobe in lateral view. 2, 4 – L. uncletan sp. nov. 2 – habitus; 4 – aedeagus (a – median lobe in lateral view; b – apex of median lobe in dorsal view; c – left paramere; d – right paramere). Scale bars: 2 mm (Figs 1-2); 0.5 mm (Figs 3-4)." figureDoi="http://doi.org/10.5281/zenodo.5176620" httpUri="https://zenodo.org/record/5176620/files/figure.png" pageId="3" pageNumber="54">Fig. 4</figureCitation>
.
</paragraph>
<paragraph id="761F36D9FFFCE93EFF4CE8A7FE951710" blockId="3.[113,1090,223,1559]" pageId="3" pageNumber="54">Female. Generally similar to male. Posterior margin of anal sternite laterally only slightly incised, medially more prominent than in male, with truncate apex. Tarsomeres of fore and middle tarsi not dilated.</paragraph>
</subSubSection>
<subSubSection id="3EBA6552FFFCE93EFFADEF07FE8C1650" pageId="3" pageNumber="54" type="diagnosis">
<paragraph id="761F36D9FFFCE93EFFADEF07FE8C1650" blockId="3.[113,1090,223,1559]" pageId="3" pageNumber="54">
<emphasis id="44D4EACBFFFCE93EFFADEF07FE931730" bold="true" box="[113,361,1119,1143]" pageId="3" pageNumber="54">Differential diagnosis.</emphasis>
Based on the double elytral reticulation and the long and acute prosternal process, the new species is classified into the
<taxonomicName id="B1A04D5AFFFCE93EFD54EF27FC8F17D0" authorityName="Sharp" authorityYear="1882" box="[648,885,1151,1175]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="44D4EACBFFFCE93EFD54EF27FC8F17D0" box="[648,885,1151,1175]" italics="true" pageId="3" pageNumber="54">Laccophilus siamensis</emphasis>
</taxonomicName>
-group sensu BRAN- CUCCI (1983). Its sister species is
<taxonomicName id="B1A04D5AFFFCE93EFE13EFF8FC6517F0" authority="Balke, Mazzoldi &amp; Hendrich, 1998" authorityName="Balke, Mazzoldi &amp; Hendrich" authorityYear="1998" box="[463,927,1183,1207]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="chini">
<emphasis id="44D4EACBFFFCE93EFE13EFF8FDDA17F0" box="[463,544,1183,1207]" italics="true" pageId="3" pageNumber="54">L. chini</emphasis>
<bibRefCitation id="12314B28FFFCE93EFDFBEFC7FC6517F0" author="BALKE M. &amp; MAZZOLDI P. &amp; HENDRICH L." box="[551,927,1183,1207]" pageId="3" pageNumber="54" pagination="71 - 77" refId="ref3715" refString="BALKE M., MAZZOLDI P. &amp; HENDRICH L. 1998: Two new Laccophilus species (Coleoptera: Dytiscidae) from Southeast Asia, and notes on other species of the genus. Raffles Bulletin of Zoology 46: 71 - 77." type="journal article" year="1998">Balke, Mazzoldi &amp; Hendrich, 1998</bibRefCitation>
</taxonomicName>
, from which
<taxonomicName id="B1A04D5AFFFCE93EFBF7EFF8FF361790" authority="Hájek &amp; Šťastný, 2005" authorityName="Hájek &amp; Šťastný" authorityYear="2005" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="uncletan" status="sp. nov.">
<emphasis id="44D4EACBFFFCE93EFBF7EFF8FF361790" italics="true" pageId="3" pageNumber="54">L. uncletan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5FE757B0FFFCE93EFF0FEF98FEDC1790" box="[211,294,1216,1239]" pageId="3" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
can be reliably distinguished only by the male genitalia. In
<taxonomicName id="B1A04D5AFFFCE93EFC78EF98FBE11790" authority="Hájek &amp; Šťastný, 2005" authorityName="Hájek &amp; Šťastný" authorityYear="2005" box="[932,1051,1215,1239]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="uncletan" status="sp. nov.">
<emphasis id="44D4EACBFFFCE93EFC78EF98FBE11790" box="[932,1051,1215,1239]" italics="true" pageId="3" pageNumber="54">L. uncletan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5FE757B0FFFCE93EFBFEEF98FF6517B0" pageId="3" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
the median lobe is more slender, ventrally without a subbasal rectangular spur, and has an almost rounded apex.
</paragraph>
</subSubSection>
<subSubSection id="3EBA6552FFFCE93EFFADEE47FC3316F0" pageId="3" pageNumber="54" type="distribution">
<paragraph id="761F36D9FFFCE93EFFADEE47FCDC1670" blockId="3.[113,1090,223,1559]" box="[113,806,1311,1335]" pageId="3" pageNumber="54">
<emphasis id="44D4EACBFFFCE93EFFADEE47FEF81670" bold="true" box="[113,258,1311,1335]" pageId="3" pageNumber="54">Distribution.</emphasis>
Known only from northern
<collectingRegion id="B464F83BFFFCE93EFDF2EE47FD5C1670" box="[558,678,1311,1335]" country="Malaysia" name="Sabah" pageId="3" pageNumber="54">Sabah state</collectingRegion>
(
<collectingCountry id="0EB77649FFFCE93EFD68EE47FCE11670" box="[692,795,1311,1335]" name="Malaysia" pageId="3" pageNumber="54">Malaysia</collectingCountry>
).
</paragraph>
<paragraph id="761F36D9FFFCE93EFFADEE67FC3316F0" blockId="3.[113,1090,223,1559]" pageId="3" pageNumber="54">
<emphasis id="44D4EACBFFFCE93EFFADEE67FF261610" bold="true" box="[113,220,1343,1367]" pageId="3" pageNumber="54">Bionomy.</emphasis>
The specimens were collected in primary lowland forest in a shallow oxbow of the river Kinabatangan together with
<taxonomicName id="B1A04D5AFFFCE93EFE0AEE38FCFC1630" authority="Brancucci, 1983" authorityName="Brancucci" authorityYear="1983" box="[470,774,1375,1399]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="bacchusi">
<emphasis id="44D4EACBFFFCE93EFE0AEE38FDAA1630" box="[470,592,1375,1399]" italics="true" pageId="3" pageNumber="54">L. bacchusi</emphasis>
<bibRefCitation id="12314B28FFFCE93EFD8BEE07FCFC1630" author="BRANCUCCI M." box="[599,774,1375,1399]" pageId="3" pageNumber="54" pagination="241 - 426" refId="ref3760" refString="BRANCUCCI M. 1983: Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey 31 / 32: 241 - 426." type="journal article" year="1983">Brancucci, 1983</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B1A04D5AFFFCE93EFCCDEE38FBBB1630" authority="Regimbart, 1880" authorityName="Regimbart" authorityYear="1880" box="[785,1089,1375,1399]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="ritsemae">
<emphasis id="44D4EACBFFFCE93EFCCDEE38FC721630" box="[785,904,1376,1399]" italics="true" pageId="3" pageNumber="54">L. ritsemae</emphasis>
Régimbart, 1880
</taxonomicName>
and
<emphasis id="44D4EACBFFFCE93EFF42EED8FF3116D0" box="[158,203,1408,1431]" italics="true" pageId="3" pageNumber="54">L. s.</emphasis>
<taxonomicName id="B1A04D5AFFFCE93EFF0EEED8FE3816D0" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[210,450,1407,1431]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="44D4EACBFFFCE93EFF0EEED8FEC316D0" box="[210,313,1408,1431]" italics="true" pageId="3" pageNumber="54">siamensis</emphasis>
Sharp, 1882
</taxonomicName>
. They inhabited muddy water near the banks of small drains flowing to the oxbow. The substrate was silty with accumulated decaying leaves.
</paragraph>
</subSubSection>
<subSubSection id="3EBA6552FFFCE93EFFADEEE7FEC11550" pageId="3" pageNumber="54" type="etymology">
<paragraph id="761F36D9FFFCE93EFFADEEE7FEC11550" blockId="3.[113,1090,223,1559]" pageId="3" pageNumber="54">
<emphasis id="44D4EACBFFFCE93EFFADEEE7FF0B1690" bold="true" box="[113,241,1471,1495]" pageId="3" pageNumber="54">Etymology.</emphasis>
This species is dedicated to Mr. Tan Su Lim known as ‘Uncle Tan’, an enthusiastic supporter of ecology popularisation and a leading environmentalist in
<collectingRegion id="B464F83BFFFCE93EFCABEE87FC4116B0" box="[887,955,1503,1527]" country="Malaysia" name="Sabah" pageId="3" pageNumber="54">Sabah</collectingRegion>
. The noun is used in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>

243
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<mods:title id="70E7602EC1D7F4A189AC56C43621562F">Two new species of Laccophilus Leach, 1815 (Coleoptera: Dytiscidae) from India and Borneo with notes on other species of the genus</mods:title>
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<mods:affiliation id="4C1338B5176BC72704065004BF27640D">Department of Entomology, National Museum, Kunratice 1, CZ- 148 00 Praha 4, Czech Republic; e-mail: jiri _ hajek @ nm. cz</mods:affiliation>
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<heading id="2D5781B5FFFEE93CFE67E99AFD0D119D" bold="true" box="[443,759,706,730]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="52" reason="3">
<emphasis id="44D4EACBFFFEE93CFE67E99AFD0D119D" bold="true" box="[443,759,706,730]" pageId="1" pageNumber="52">
<taxonomicName id="B1A04D5AFFFEE93CFE67E99AFD60119D" ID-CoL="3RM5M" authority="Hájek &amp; Šťastný, 2005" authorityName="Hájek &amp; Šťastný" authorityYear="2005" box="[443,666,706,730]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="boukali" status="sp. nov.">
<emphasis id="44D4EACBFFFEE93CFE67E99AFD60119D" bold="true" box="[443,666,706,730]" italics="true" pageId="1" pageNumber="52">Laccophilus boukali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5FE757B0FFFEE93CFD7EE99AFD0D119D" box="[674,759,706,730]" pageId="1" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3EBA6552FFFEE93CFDF6E9BAFD7211B2" box="[554,648,738,758]" pageId="1" pageNumber="52" type="description">
<paragraph id="761F36D9FFFEE93CFDF6E9BAFD7211B2" blockId="1.[554,648,738,758]" box="[554,648,738,758]" pageId="1" pageNumber="52">
(
<figureCitation id="EE9B2A5CFFFEE93CFDECE9BAFD7811B2" box="[560,642,738,758]" captionStart="Figs 1-4. 1, 3" captionStartId="4.[151,188,1486,1505]" captionTargetBox="[167,1112,215,1451]" captionTargetId="figure-9@4.[167,1112,215,1451]" captionTargetPageId="4" captionText="Figs 1-4. 1, 3 – Laccophilus boukali sp. nov. 1 – habitus; 3 – median lobe in lateral view. 2, 4 – L. uncletan sp. nov. 2 – habitus; 4 – aedeagus (a – median lobe in lateral view; b – apex of median lobe in dorsal view; c – left paramere; d – right paramere). Scale bars: 2 mm (Figs 1-2); 0.5 mm (Figs 3-4)." figureDoi="http://doi.org/10.5281/zenodo.5176620" httpUri="https://zenodo.org/record/5176620/files/figure.png" pageId="1" pageNumber="52">Figs. 1, 3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3EBA6552FFFEE93CFFADE840FC6C1757" pageId="1" pageNumber="52" type="materials_examined">
<paragraph id="761F36D9FFFEE93CFFADE840FD571068" blockId="1.[113,1089,791,1040]" box="[113,685,791,816]" pageId="1" pageNumber="52">
<emphasis id="44D4EACBFFFEE93CFFADE840FEF21068" bold="true" box="[113,264,791,816]" pageId="1" pageNumber="52">
<typeStatus id="A91B887BFFFEE93CFFADE840FF521077" box="[113,168,792,816]" pageId="1" pageNumber="52">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="0EB77649FFFEE93CFED2E840FEBD1077" box="[270,327,792,816]" name="India" pageId="1" pageNumber="52">India</collectingCountry>
,
<collectingRegion id="B464F83BFFFEE93CFE8EE840FE381077" box="[338,450,792,816]" country="India" name="Karnataka" pageId="1" pageNumber="52">Karnataka</collectingRegion>
, Coorg, Kakkabe env.
</paragraph>
<paragraph id="761F36D9FFFEE93CFFADE860FC6C1757" blockId="1.[113,1089,791,1040]" pageId="1" pageNumber="52">
<emphasis id="44D4EACBFFFEE93CFFADE860FEE81008" bold="true" box="[113,274,823,848]" pageId="1" pageNumber="52">Type material.</emphasis>
<materialsCitation id="C6C83C84FFFEE93CFECAE860FD361037" collectingDate="1998-12-21" collectorName="D. Boukal" country="India" elevation="1050" latitude="12.25" location="Kakkabe" longLatPrecision="1293" longitude="75.583336" municipality="Coorg" pageId="1" pageNumber="52" specimenCount="1" stateProvince="Karnataka" typeStatus="holotype">
<typeStatus id="A91B887BFFFEE93CFECAE860FE811009" box="[278,379,824,847]" pageId="1" pageNumber="52" type="holotype">HOLOTYPE</typeStatus>
: J, ‘
<collectingCountry id="0EB77649FFFEE93CFE75E860FE061017" box="[425,508,824,848]" name="India" pageId="1" pageNumber="52">INDIA</collectingCountry>
:
<collectingRegion id="B464F83BFFFEE93CFDD8E860FD881017" box="[516,626,824,848]" country="India" name="Karnataka" pageId="1" pageNumber="52">Karnataka</collectingRegion>
,
<collectingMunicipality id="967BACA3FFFEE93CFDA6E860FD411017" box="[634,699,824,848]" pageId="1" pageNumber="52">Coorg</collectingMunicipality>
/
<location id="737F6002FFFEE93CFD17E860FCD21017" LSID="urn:lsid:plazi:treatment:FE0987CFFFFEE93FFE67E99AFEBE17F0:737F6002FFFEE93CFD17E860FCD21017" box="[715,808,824,848]" country="India" latitude="12.25" longLatPrecision="1293" longitude="75.583336" municipality="Coorg" name="Kakkabe" pageId="1" pageNumber="52" stateProvince="Karnataka">Kakkabe</location>
env.,
<date id="021E1019FFFEE93CFCBFE860FC221017" box="[867,984,824,848]" pageId="1" pageNumber="52" value="1998-12-21">
<collectingDate id="125AE9F1FFFEE93CFCBFE860FC221017" box="[867,984,824,848]" pageId="1" pageNumber="52" value="1998-12-21">21.12.1998</collectingDate>
</date>
/
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<geoCoordinate id="1394501EFFFEE93CFFADE800FF331028" box="[113,201,854,880]" degrees="75" direction="east" minutes="35" orientation="longitude" pageId="1" pageNumber="52" precision="925" value="75.583336">75°35′E</geoCoordinate>
,
<quantity id="B1589B3CFFFEE93CFF0BE800FE981037" box="[215,354,856,880]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="1" pageNumber="52" unit="m" value="1050.0" valueMax="1200.0" valueMin="900.0">
<elevation id="FD8DD1EAFFFEE93CFF0BE800FE981037" box="[215,354,856,880]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="1" pageNumber="52" unit="m" value="1050.0" valueMax="1200.0" valueMin="900.0">900 - 1200m</elevation>
</quantity>
/ leg.
<collectorName id="DB55530FFFFEE93CFE77E800FDE11037" box="[427,539,856,880]" pageId="1" pageNumber="52">D. Boukal</collectorName>
(12) [printed] //
</materialsCitation>
<materialsCitation id="C6C83C84FFFEE93CFD09E800FC5510D7" collectionCode="NMW" country="India" determinerName="J. Hajek &amp; J. Stastny" pageId="1" pageNumber="52" specimenCount="1" stateProvince="Karnataka" typeStatus="holotype">
<typeStatus id="A91B887BFFFEE93CFD09E800FC9B1037" box="[725,865,856,880]" pageId="1" pageNumber="52" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="B1A04D5AFFFEE93CFCA6E800FBCB1037" authorityName="Leach" authorityYear="1815" box="[890,1073,856,880]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="genus">LACCOPHILUS</taxonomicName>
/ boukali
<taxonomicNameLabel id="5FE757B0FFFEE93CFF14E820FEE010C8" box="[200,282,888,911]" pageId="1" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
/
<determinerName id="83C3DA1DFFFEE93CFEF3E820FE7F10D7" box="[303,389,888,912]" pageId="1" pageNumber="52">J. Hájek</determinerName>
&amp;
<determinerName id="83C3DA1DFFFEE93CFE74E820FDE810D7" box="[424,530,882,912]" pageId="1" pageNumber="52">J. Šťastný</determinerName>
det. 2005’ [printed, red label] (
<collectionCode id="10B1AE1CFFFEE93CFC82E820FC5210D7" box="[862,936,888,912]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="52">NMW</collectionCode>
)
</materialsCitation>
.
<typeStatus id="A91B887BFFFEE93CFC65E820FBDF10C9" box="[953,1061,888,911]" pageId="1" pageNumber="52" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="60A6FD50FFFEE93CFBEFE820FF7410F6" pageId="1" pageNumber="52" type="female">2 ♀♀</specimenCount>
, same label data as
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(
<collectionCode id="10B1AE1CFFFEE93CFE00E8C0FDDC10F7" box="[476,550,920,944]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="52">NMW</collectionCode>
). 1 J, ‘
<collectingCountry id="0EB77649FFFEE93CFDAFE8C0FD3D10F7" box="[627,711,920,944]" name="India" pageId="1" pageNumber="52">INDIA</collectingCountry>
:
<collectingRegion id="B464F83BFFFEE93CFD0EE8C0FCB810F7" box="[722,834,920,944]" country="India" name="Karnataka" pageId="1" pageNumber="52">Karnataka</collectingRegion>
, Coorg / Madikeri env.,
<date id="021E1019FFFEE93CFFADE8E0FF131097" box="[113,233,952,976]" pageId="1" pageNumber="52" value="1998-12-19">19.12.1998</date>
/
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, Talakaveri /
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<geoCoordinate id="1394501EFFFEE93CFD92E8E0FD5F1088" box="[590,677,950,976]" degrees="75" direction="east" minutes="30" orientation="longitude" pageId="1" pageNumber="52" precision="925" value="75.5">75°30′E</geoCoordinate>
/ leg. D. Boukal (5)’ (
<collectionCode id="10B1AE1CFFFEE93CFC79E8E0FC0D1097" box="[933,1015,952,976]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="52">NMPC</collectionCode>
). Each
<typeStatus id="A91B887BFFFEE93CFFADE880FF3610B7" box="[113,204,984,1008]" pageId="1" pageNumber="52" type="paratype">paratype</typeStatus>
is provided with a red label similar to that of the
<typeStatus id="A91B887BFFFEE93CFD2EE880FCAB10B7" box="[754,849,984,1008]" pageId="1" pageNumber="52" type="holotype">holotype</typeStatus>
, except ‘PARATYPE’ instead of ‘HOLOTYPE’, the respective sex symbol and collection number.
</paragraph>
</subSubSection>
<subSubSection id="3EBA6552FFFEE93FFFADEF7FFC4B1170" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52" type="description">
<paragraph id="761F36D9FFFEE93CFFADEF7FFBFB1719" blockId="1.[113,1090,1063,1599]" pageId="1" pageNumber="52">
<emphasis id="44D4EACBFFFEE93CFFADEF7FFF011778" bold="true" box="[113,251,1063,1087]" pageId="1" pageNumber="52">Description.</emphasis>
Oval and oblong; head and pronotum testaceous, larger part of elytra brown with testaceous pattern as depicted in
<figureCitation id="EE9B2A5CFFFEE93CFDD9EF1FFDB91718" box="[517,579,1095,1119]" captionStart="Figs 1-4. 1, 3" captionStartId="4.[151,188,1486,1505]" captionTargetBox="[167,1112,215,1451]" captionTargetId="figure-9@4.[167,1112,215,1451]" captionTargetPageId="4" captionText="Figs 1-4. 1, 3 – Laccophilus boukali sp. nov. 1 – habitus; 3 – median lobe in lateral view. 2, 4 – L. uncletan sp. nov. 2 – habitus; 4 – aedeagus (a – median lobe in lateral view; b – apex of median lobe in dorsal view; c – left paramere; d – right paramere). Scale bars: 2 mm (Figs 1-2); 0.5 mm (Figs 3-4)." figureDoi="http://doi.org/10.5281/zenodo.5176620" httpUri="https://zenodo.org/record/5176620/files/figure.png" pageId="1" pageNumber="52">Fig. 1</figureCitation>
. Ventral parts and appendages testaceous.
</paragraph>
<paragraph id="761F36D9FFFEE93CFF4CEF3FFCD617D8" blockId="1.[113,1090,1063,1599]" pageId="1" pageNumber="52">
Measurements. Body length
<quantity id="B1589B3CFFFEE93CFE1FEF3FFDA21739" box="[451,600,1127,1151]" metricMagnitude="-3" metricUnit="m" metricValue="3.275" metricValueMax="3.35" metricValueMin="3.2" pageId="1" pageNumber="52" unit="mm" value="3.275" valueMax="3.35" valueMin="3.2">3.20-3.35 mm</quantity>
(mean value =
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,
<typeStatus id="A91B887BFFFEE93CFCB2EF3FFC311738" box="[878,971,1127,1151]" pageId="1" pageNumber="52" type="holotype">holotype</typeStatus>
<quantity id="B1589B3CFFFEE93CFC0EEF3FFBCF1738" box="[978,1077,1127,1151]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="1" pageNumber="52" unit="mm" value="3.2">3.20 mm</quantity>
), width
<quantity id="B1589B3CFFFEE93CFF68EFDFFEB317D9" box="[180,329,1159,1183]" metricMagnitude="-3" metricUnit="m" metricValue="1.825" metricValueMax="1.85" metricValueMin="1.8" pageId="1" pageNumber="52" unit="mm" value="1.825" valueMax="1.85" valueMin="1.8">1.80-1.85 mm</quantity>
(mean value =
<quantity id="B1589B3CFFFEE93CFE33EFDFFDAA17D9" box="[495,592,1159,1183]" metricMagnitude="-3" metricUnit="m" metricValue="1.81" pageId="1" pageNumber="52" unit="mm" value="1.81">1.81 mm</quantity>
,
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<quantity id="B1589B3CFFFEE93CFD61EFDFFCDA17D8" box="[701,800,1159,1183]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="52" unit="mm" value="1.8">1.80 mm</quantity>
).
</paragraph>
<paragraph id="761F36D9FFFEE93CFF4CEFFFFDFD1659" blockId="1.[113,1090,1063,1599]" pageId="1" pageNumber="52">Head. Coloration testaceous, in some specimens with indistinct light brown markings. Reticulation double, composed of moderately deeply impressed transverse polygonal meshes on anterior part and irregular meshes on posterior part, and microreticulation of very fine round meshes. Mouthparts testaceous.</paragraph>
<paragraph id="761F36D9FFFEE93CFF4CEE7FFCA31639" blockId="1.[113,1090,1063,1599]" pageId="1" pageNumber="52">Pronotum. Coloration testaceous, with two small brown spots at basal margin, and with anterior margin slightly darkened. Reticulation double, similar to that of head, but meshes larger. Anterior margin and posterior angles with numerous punctures.</paragraph>
<paragraph id="761F36D9FFFEE93FFF4CEEDFFE421270" blockId="1.[113,1090,1063,1599]" lastBlockId="2.[151,1127,223,1207]" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52">Elytra. Elytral disc brown with testaceous markings, lateral parts testaceous. Subbasal testaceous band with anterior border almost straight, posterior border with distinct elongation near elytral suture and in most specimens also with smaller discal elongation. Subapical band divided into five longitudinal stripes. Remaining elytral surface with sinuous stripes, often indistinct in middle third of elytra, but widening apically, with elytral apex nearly testaceous in some specimens. Reticulation on whole elytra distinctly double, composed of irregular polygonal meshes and microreticulation, which is moderately deeply impressed and more distinct than on head or pronotum. Each elytron with sutural, discal and lateral row of large serial setigerous punctures.</paragraph>
<paragraph id="761F36D9FFFDE93FFF6AEA67FDB812DF" blockId="2.[151,1127,223,1207]" pageId="2" pageNumber="53">Ventral side. Testaceous and microreticulated. Prosternal process long and acute. Abdominal sternites laterally with oblique grooves. Anal sternite tectiform, its surface with large punctures, setae and transverse grooves.</paragraph>
<paragraph id="761F36D9FFFDE93FFF6AEAF8FF3612B0" blockId="2.[151,1127,223,1207]" pageId="2" pageNumber="53">Legs. Completely testaceous. Male tarsomeres 1-3 of fore and middle legs weakly dilated with microtrichia on ventral side. Claws of fore tarsi simple and equal, as well as those of mid tarsi.</paragraph>
<paragraph id="761F36D9FFFDE93FFF6AEAA7FD3F1150" blockId="2.[151,1127,223,1207]" box="[182,709,511,535]" pageId="2" pageNumber="53">
Male genitalia. Shape of median lobe as in
<figureCitation id="EE9B2A5CFFFDE93FFD5DEAA7FD3B1150" box="[641,705,511,535]" captionStart="Figs 1-4. 1, 3" captionStartId="4.[151,188,1486,1505]" captionTargetBox="[167,1112,215,1451]" captionTargetId="figure-9@4.[167,1112,215,1451]" captionTargetPageId="4" captionText="Figs 1-4. 1, 3 – Laccophilus boukali sp. nov. 1 – habitus; 3 – median lobe in lateral view. 2, 4 – L. uncletan sp. nov. 2 – habitus; 4 – aedeagus (a – median lobe in lateral view; b – apex of median lobe in dorsal view; c – left paramere; d – right paramere). Scale bars: 2 mm (Figs 1-2); 0.5 mm (Figs 3-4)." figureDoi="http://doi.org/10.5281/zenodo.5176620" httpUri="https://zenodo.org/record/5176620/files/figure.png" pageId="2" pageNumber="53">Fig. 3</figureCitation>
.
</paragraph>
<paragraph id="761F36D9FFFDE93FFF6AE947FC4B1170" blockId="2.[151,1127,223,1207]" box="[182,945,543,567]" pageId="2" pageNumber="53">Female. Similar to male; tarsomeres of fore and middle legs not dilated.</paragraph>
</subSubSection>
<subSubSection id="3EBA6552FFFDE93FFF4BE967FDF01030" pageId="2" pageNumber="53" type="diagnosis">
<paragraph id="761F36D9FFFDE93FFF4BE967FDF01030" blockId="2.[151,1127,223,1207]" pageId="2" pageNumber="53">
<emphasis id="44D4EACBFFFDE93FFF4BE967FE751110" bold="true" box="[151,399,575,599]" pageId="2" pageNumber="53">Differential diagnosis.</emphasis>
Based on the double reticulation of pronotum and elytra and the elytral pattern (brown sinuous stripes), the new species belongs to the
<taxonomicName id="B1A04D5AFFFDE93FFCAEE907FBA71130" authorityName="Boheman" authorityYear="1858" box="[882,1117,607,631]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="44D4EACBFFFDE93FFCAEE907FBA71130" box="[882,1117,607,631]" italics="true" pageId="2" pageNumber="53">Laccophilus chinensis</emphasis>
</taxonomicName>
- group as defined by
<bibRefCitation id="12314B28FFFDE93FFEABE9D8FDC311D0" author="BRANCUCCI M." box="[375,569,639,663]" pageId="2" pageNumber="53" pagination="241 - 426" refId="ref3760" refString="BRANCUCCI M. 1983: Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey 31 / 32: 241 - 426." type="journal article" year="1983">BRANCUCCI (1983)</bibRefCitation>
. Among the
<collectingRegion id="B464F83BFFFDE93FFD18E927FCE111D0" box="[708,795,639,663]" country="Democratic Republic of the Congo" name="Orientale" pageId="2" pageNumber="53">Oriental</collectingRegion>
species of this group, the new species is similar to
<taxonomicName id="B1A04D5AFFFDE93FFEAAE9F8FD5511F0" authority="Regimbart, 1880" authorityName="Regimbart" authorityYear="1880" box="[374,687,671,695]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="ritsemae">
<emphasis id="44D4EACBFFFDE93FFEAAE9F8FE1511F0" box="[374,495,672,695]" italics="true" pageId="2" pageNumber="53">L. ritsemae</emphasis>
Régimbart, 1880
</taxonomicName>
, which it resembles in the habitus, body length and coloration. Both species can only be distinguished by the male genitalia. The median lobe of
<taxonomicName id="B1A04D5AFFFDE93FFEE4E9B8FE5B11B0" authority="Hájek &amp; Šťastný, 2005" authorityName="Hájek &amp; Šťastný" authorityYear="2005" box="[312,417,735,759]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="boukali" status="sp. nov.">
<emphasis id="44D4EACBFFFDE93FFEE4E9B8FE5B11B0" box="[312,417,735,759]" italics="true" pageId="2" pageNumber="53">L. boukali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5FE757B0FFFDE93FFE7BE9B8FE0211B0" box="[423,504,736,759]" pageId="2" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
is short and broad with a distinct ventral angle in the middle third of its length, while the median lobe of
<taxonomicName id="B1A04D5AFFFDE93FFDAFE858FD111050" authorityName="Regimbart" authorityYear="1880" box="[627,747,768,791]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="ritsemae">
<emphasis id="44D4EACBFFFDE93FFDAFE858FD111050" box="[627,747,768,791]" italics="true" pageId="2" pageNumber="53">L. ritsemae</emphasis>
</taxonomicName>
is long and slender with its ventral edge sinuous. From other sympatrically occurring species,
<taxonomicName id="B1A04D5AFFFDE93FFCD7E878FBC01070" authority="(Walker, 1859)" baseAuthorityName="Walker" baseAuthorityYear="1859" box="[779,1082,799,823]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="inefficiens">
<emphasis id="44D4EACBFFFDE93FFCD7E878FC6E1070" box="[779,916,799,823]" italics="true" pageId="2" pageNumber="53">L. inefficiens</emphasis>
(Walker, 1859)
</taxonomicName>
and
<taxonomicName id="B1A04D5AFFFDE93FFF4BE818FE3B1010" authority="Brancucci, 1983" authorityName="Brancucci" authorityYear="1983" box="[151,449,831,855]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="kaszabi">
<emphasis id="44D4EACBFFFDE93FFF4BE818FEFF1010" box="[151,261,831,855]" italics="true" pageId="2" pageNumber="53">L. kaszabi</emphasis>
<bibRefCitation id="12314B28FFFDE93FFED3E867FE3B1010" author="BRANCUCCI M." box="[271,449,831,855]" pageId="2" pageNumber="53" pagination="241 - 426" refId="ref3760" refString="BRANCUCCI M. 1983: Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey 31 / 32: 241 - 426." type="journal article" year="1983">Brancucci, 1983</bibRefCitation>
</taxonomicName>
, the new species differs by the generally smaller size and the different shape of the median lobe.
</paragraph>
</subSubSection>
<subSubSection id="3EBA6552FFFDE93FFF4BE827FEA11710" pageId="2" pageNumber="53" type="distribution">
<paragraph id="761F36D9FFFDE93FFF4BE827FF2F10F0" blockId="2.[151,1127,223,1207]" pageId="2" pageNumber="53">
<emphasis id="44D4EACBFFFDE93FFF4BE827FEDD10D0" bold="true" box="[151,295,895,919]" pageId="2" pageNumber="53">Distribution.</emphasis>
So far known only from two close localities in
<collectingRegion id="B464F83BFFFDE93FFCCDE827FC4E10D0" box="[785,948,895,919]" country="India" name="Karnataka" pageId="2" pageNumber="53">Karnataka state</collectingRegion>
in south-western
<collectingCountry id="0EB77649FFFDE93FFF4BE8C7FF2B10F0" box="[151,209,927,951]" name="India" pageId="2" pageNumber="53">India</collectingCountry>
.
</paragraph>
<paragraph id="761F36D9FFFDE93FFF4BE8E7FEA11710" blockId="2.[151,1127,223,1207]" pageId="2" pageNumber="53">
<emphasis id="44D4EACBFFFDE93FFF4BE8E7FEF81090" bold="true" box="[151,258,959,983]" pageId="2" pageNumber="53">Bionomy.</emphasis>
The specimens from Kakkabe were collected in a larger stream with the main channel ca
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wide, with large boulders, a rocky bed with some stones and gravel in riffles and silt in pools. The specimen from Madikeri was collected in an artificial reservoir at a hilltop; the examined site had a dense shore vegetation, muddy bed, and was little shaded (D. S. BOUKAL in litt.).
</paragraph>
</subSubSection>
<subSubSection id="3EBA6552FFFDE93FFF4BEF07FEBE17F0" pageId="2" pageNumber="53" type="etymology">
<paragraph id="761F36D9FFFDE93FFF4BEF07FEBE17F0" blockId="2.[151,1127,223,1207]" pageId="2" pageNumber="53">
<emphasis id="44D4EACBFFFDE93FFF4BEF07FEED1730" bold="true" box="[151,279,1119,1143]" pageId="2" pageNumber="53">Etymology.</emphasis>
The new species is dedicated to its collector David S. Boukal (České Budějovice,
<collectingCountry id="0EB77649FFFDE93FFF0CEF27FE8617D0" box="[208,380,1151,1175]" name="Czech Republic" pageId="2" pageNumber="53">Czech Republic</collectingCountry>
), our friend, theoretical mathematician, photographer, and specialist on the Dryopoidea.
</paragraph>
</subSubSection>
</treatment>
</document>

136
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<taxonomicName authority="(Sseliwanoff, 1881)" baseAuthorityName="Sseliwanoff" baseAuthorityYear="1881" box="[227,663,829,851]" class="Chilopoda" family="Lithobiidae" genus="Disphaerobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiomorpha" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="loricatus">
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<emphasis bold="true" box="[227,464,829,851]" italics="true" pageId="2" pageNumber="29">Disphaerobius loricatus</emphasis>
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MATERIAL EXAMINED.
<emphasis bold="true" box="[536,713,872,894]" pageId="2" pageNumber="29">Kostanay Region</emphasis>
: Amangeldy District, Altyn Dala Reserve,
</paragraph>
<paragraph blockId="2.[227,1128,872,1060]" pageId="2" pageNumber="29">
<materialsCitation collectingDate="2013-07-04" county="river Uly-Zhilanshik" latitude="49.266666" location="Ulyzhilanshik" longLatPrecision="1105" longitude="65.23333" municipality="Rakhmet Village" pageId="2" pageNumber="29" specimenCount="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:FE0987D3FFC0FFCA1294D82AFE42FBBB:737F601EFFC0FFCA1294D893FED7FC05" box="[227,362,900,922]" county="river Uly-Zhilanshik" latitude="49.266666" longLatPrecision="1105" longitude="65.23333" municipality="Rakhmet Village" name="Ulyzhilanshik" pageId="2" pageNumber="29">Ulyzhilanshik</location>
site, near destroyed
<collectingMunicipality box="[567,734,900,922]" pageId="2" pageNumber="29">Rakhmet Village</collectingMunicipality>
, bank of the
<collectingCounty box="[865,1067,900,922]" pageId="2" pageNumber="29">river Uly-Zhilanshik</collectingCounty>
, plant association with
<taxonomicName box="[390,534,928,949]" class="Insecta" family="Pyralidae" genus="Anabasis" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="salsa">
<emphasis box="[390,534,928,949]" italics="true" pageId="2" pageNumber="29">Anabasis salsa</emphasis>
</taxonomicName>
,
<collectingMethod box="[547,661,928,950]" pageId="2" pageNumber="29">pitfall traps</collectingMethod>
,
<geoCoordinate box="[671,760,927,950]" degrees="49" direction="north" minutes="16" orientation="latitude" pageId="2" pageNumber="29" precision="925" value="49.266666">49°16′ N</geoCoordinate>
,
<geoCoordinate box="[771,858,927,950]" degrees="65" direction="east" minutes="14" orientation="longitude" pageId="2" pageNumber="29" precision="925" value="65.23333">65°14′ E</geoCoordinate>
,
<date box="[869,989,928,949]" pageId="2" pageNumber="29" value="2013-07-04">
<collectingDate box="[869,989,928,949]" pageId="2" pageNumber="29" value="2013-07-04">04.VII 2013</collectingDate>
</date>
,
<specimenCount box="[1000,1032,928,949]" pageId="2" pageNumber="29" type="male">1♂</specimenCount>
</materialsCitation>
;
<materialsCitation collectingDate="2014-05-07" county="river Uly-Zhilanshik" latitude="49.266666" location="Ulyzhilanshik" longLatPrecision="1105" longitude="65.23333" municipality="Rakhmet Village" pageId="2" pageNumber="29" specimenCount="1" specimenCount-male="1">
same locality, under shelter,
<date box="[424,525,955,976]" pageId="2" pageNumber="29" value="2014-05-07">
<collectingDate box="[424,525,955,976]" pageId="2" pageNumber="29" value="2014-05-07">07.V 2014</collectingDate>
</date>
,
<specimenCount box="[535,565,955,976]" pageId="2" pageNumber="29" type="male">1♂</specimenCount>
(PSU-871)
</materialsCitation>
;
<materialsCitation box="[687,1109,955,977]" collectingDate="2014-05-07" county="river Uly-Zhilanshik" latitude="49.266666" location="Ulyzhilanshik" longLatPrecision="1105" longitude="65.23333" municipality="Rakhmet Village" pageId="2" pageNumber="29" specimenCount="1" specimenCount-male="1">
same locality, under shelter,
<date box="[964,1068,955,976]" pageId="2" pageNumber="29" value="2014-05-07">
<collectingDate box="[964,1068,955,976]" pageId="2" pageNumber="29" value="2014-05-07">07.V 2014</collectingDate>
</date>
,
<specimenCount box="[1077,1109,955,976]" pageId="2" pageNumber="29" type="male">1♂</specimenCount>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="29" type="distribution">
<paragraph blockId="2.[227,1128,872,1060]" box="[265,1128,983,1005]" pageId="2" pageNumber="29">
DISTRIBUTION.
<collectingCountry box="[450,566,983,1005]" name="Kazakhstan" pageId="2" pageNumber="29">Kazakhstan</collectingCountry>
: Aktobe,
<collectingRegion box="[666,830,983,1005]" country="Kazakhstan" name="Shyghys Qazaqstan Oblysy" pageId="2" pageNumber="29">East Kazakhstan</collectingRegion>
and
<collectingRegion box="[885,957,983,1005]" country="Kazakhstan" name="Almaty Oblysy" pageId="2" pageNumber="29">Almaty</collectingRegion>
regions;
<collectingCountry box="[1056,1124,983,1005]" name="Russia" pageId="2" pageNumber="29">Russia</collectingCountry>
:
</paragraph>
<paragraph blockId="2.[227,1128,872,1060]" box="[227,1128,1011,1033]" pageId="2" pageNumber="29">
<collectingRegion box="[227,395,1011,1033]" country="Russia" name="Orenburg" pageId="2" pageNumber="29">Orenburg Region</collectingRegion>
(Farzalieva &amp; Zalesskaja, 2003; Farzalieva
<emphasis box="[828,872,1011,1032]" italics="true" pageId="2" pageNumber="29">et al</emphasis>
., 2017; Dyachkov, 2019).
</paragraph>
<paragraph blockId="2.[227,1128,872,1060]" box="[227,511,1038,1060]" pageId="2" pageNumber="29">New to the Kostanay Region.</paragraph>
</subSubSection>
</treatment>
</document>

132
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<heading id="2D5781A9FFC0FFCA1294DAE3FD5CFD96" box="[227,737,500,522]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="29" reason="2">
<taxonomicName id="B1A04D46FFC0FFCA1294DAE3FD5CFD96" ID-CoL="3VJ92" authority="L. Koch, 1862" authorityName="L. Koch" authorityYear="1862" box="[227,737,500,522]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="crassipes" subGenus="Monotarsobius">
<emphasis id="44D4EAD7FFC0FFCA1294DAE3FD5CFD96" bold="true" box="[227,737,500,522]" pageId="2" pageNumber="29">
<emphasis id="44D4EAD7FFC0FFCA1294DAE3FEFFFD95" bold="true" box="[227,322,500,522]" italics="true" pageId="2" pageNumber="29">Lithobius</emphasis>
(
<emphasis id="44D4EAD7FFC0FFCA1327DAE3FE58FD95" bold="true" box="[336,485,500,522]" italics="true" pageId="2" pageNumber="29">Monotarsobius</emphasis>
)
<emphasis id="44D4EAD7FFC0FFCA1384DAE3FDF1FD96" bold="true" box="[499,588,500,521]" italics="true" pageId="2" pageNumber="29">crassipes</emphasis>
L. Koch, 1862
</emphasis>
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<paragraph id="761F36C5FFC0FFCA137ED937FBDBFDAB" blockId="2.[227,1127,543,731]" box="[265,1126,543,565]" pageId="2" pageNumber="29">
MATERIAL EXAMINED.
<emphasis id="44D4EAD7FFC0FFCA1068D908FD6EFDAA" bold="true" box="[543,723,543,565]" pageId="2" pageNumber="29">Kostanay Region</emphasis>
: Kostanay District, Stoun (Kamennoye)
</paragraph>
<paragraph id="761F36C5FFC0FFCA1294D92CFBD8FDCE" blockId="2.[227,1127,543,731]" box="[227,1125,571,593]" pageId="2" pageNumber="29">
<materialsCitation id="C6C83C98FFC0FFCA1294D92CFDF5FDCE" box="[227,584,571,593]" collectionCode="V" location="Lake" pageId="2" pageNumber="29" specimenCount="1">
<location id="737F601EFFC0FFCA1294D92CFEA8FDCE" LSID="urn:lsid:plazi:treatment:FE0987D3FFC0FFCA1294DAE3FCC0FC9F:737F601EFFC0FFCA1294D92CFEA8FDCE" box="[227,277,571,593]" name="Lake" pageId="2" pageNumber="29">Lake</location>
, birch forest, in the litter,
<date id="021E1005FFC0FFCA106FD92CFDF5FDCE" box="[536,584,571,593]" pageId="2" pageNumber="29" value="2017-05-16">
16.
<collectionCode id="10B1AE00FFC0FFCA1043D92CFDF5FDCE" box="[564,584,571,593]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="29" type="Museum">V</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="C6C83C98FFC0FFCA1027D92CFBD8FDCE" box="[592,1125,571,593]" collectionCode="V" location="Kostanay town" pageId="2" pageNumber="29" specimenCount="1" specimenCount-male="1">
2017,
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(PSU-869);
<location id="737F601EFFC0FFCA115DD92CFC00FDCF" LSID="urn:lsid:plazi:treatment:FE0987D3FFC0FFCA1294DAE3FCC0FC9F:737F601EFFC0FFCA115DD92CFC00FDCF" box="[810,957,571,592]" name="Kostanay town" pageId="2" pageNumber="29">Kostanay town</location>
, in the flat, 15.
<collectionCode id="10B1AE00FFC0FFCA1625D92CFBD8FDCE" box="[1106,1125,571,593]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="29" type="Museum">V</collectionCode>
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</paragraph>
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2017,
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.
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<paragraph id="761F36C5FFC0FFCA137ED965FBDBFD17" blockId="2.[227,1127,543,731]" box="[265,1126,626,648]" pageId="2" pageNumber="29">
DISTRIBUTION. Europe (including
<collectingCountry id="0EB77655FFC0FFCA10F5D965FD7BFD17" box="[642,710,626,648]" name="Russia" pageId="2" pageNumber="29">Russia</collectingCountry>
: European part and Western Siberia), N
</paragraph>
<paragraph id="761F36C5FFC0FFCA1294D999FBDAFD3C" blockId="2.[227,1127,543,731]" box="[227,1127,654,676]" pageId="2" pageNumber="29">
Africa, Western Asia, introduced in
<collectingCountry id="0EB77655FFC0FFCA1023D999FD25FD3B" box="[596,664,654,676]" name="Taiwan" pageId="2" pageNumber="29">Taiwan</collectingCountry>
and
<collectingCountry id="0EB77655FFC0FFCA10A5D999FD43FD3C" box="[722,766,654,675]" name="United States of America" pageId="2" pageNumber="29">USA</collectingCountry>
(Zalesskaja, 1978; Nefediev
<emphasis id="44D4EAD7FFC0FFCA165BD999FBE6FD3C" box="[1068,1115,654,675]" italics="true" pageId="2" pageNumber="29">et al</emphasis>
.,
</paragraph>
<paragraph id="761F36C5FFC0FFCA1294D9BEFBDAFD20" blockId="2.[227,1127,543,731]" box="[227,1127,681,703]" pageId="2" pageNumber="29">
2016).
<collectingCountry id="0EB77655FFC0FFCA135ED9BEFE20FD20" box="[297,413,681,703]" name="Kazakhstan" pageId="2" pageNumber="29">Kazakhstan</collectingCountry>
:
<collectingRegion id="B464F827FFC0FFCA13DFD9BDFD2AFD20" box="[424,663,681,703]" country="Kazakhstan" name="Shyghys Qazaqstan Oblysy" pageId="2" pageNumber="29">East Kazakhstan Region</collectingRegion>
(Dyachkov
<emphasis id="44D4EAD7FFC0FFCA1164D9BDFCFDFD21" box="[787,832,681,703]" italics="true" pageId="2" pageNumber="29">et al</emphasis>
., 2016); new to the Kostanay
</paragraph>
<paragraph id="761F36C5FFC0FFCA1294D9D2FE8DFD44" blockId="2.[227,1127,543,731]" box="[227,304,709,731]" pageId="2" pageNumber="29">Region.</paragraph>
<paragraph id="761F36C5FFC0FFCA13BAD9FCFCC0FC9F" blockId="2.[461,893,747,769]" box="[461,893,747,769]" pageId="2" pageNumber="29">
<emphasis id="44D4EAD7FFC0FFCA13BAD9FCFCC0FC9F" bold="true" box="[461,893,747,769]" pageId="2" pageNumber="29">
Subfamily
<taxonomicName id="B1A04D46FFC0FFCA104BD9FCFCC0FC9F" authority="Verhoeff, 1933" authorityName="Verhoeff" authorityYear="1933" box="[572,893,747,769]" class="Chilopoda" family="Lithobiidae" kingdom="Animalia" order="Lithobiomorpha" pageId="2" pageNumber="29" phylum="Arthropoda" rank="subFamily" subFamily="Pterygoterginae">Pterygoterginae Verhoeff, 1933</taxonomicName>
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140
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<paragraph id="761F36C5FFC0FFCA1294DE20FD17FAD3" blockId="2.[227,682,1334,1357]" box="[227,682,1334,1357]" pageId="2" pageNumber="29">
<heading id="2D5781A9FFC0FFCA1294DE20FD17FAD3" box="[227,682,1334,1357]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="29" reason="2">
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<emphasis id="44D4EAD7FFC0FFCA1294DE20FD17FAD3" bold="true" box="[227,682,1334,1357]" pageId="2" pageNumber="29">
<emphasis id="44D4EAD7FFC0FFCA1294DE20FE55FAD2" bold="true" box="[227,488,1335,1357]" italics="true" pageId="2" pageNumber="29">Hessebius multicalcaratus</emphasis>
Folkmanová, 1958
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MATERIAL EXAMINED.
<emphasis id="44D4EAD7FFC0FFCA1068DE75FD6EFAE7" bold="true" box="[543,723,1378,1400]" pageId="2" pageNumber="29">Kostanay Region</emphasis>
: Karasu District, Karasu Village, under dry dung,
<date id="021E1005FFC0FFCA1330DE69FE10FA0C" box="[327,429,1406,1428]" pageId="2" pageNumber="29" value="2013-05-16">
16.
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<materialsCitation id="C6C83C98FFC0FFCA130BDE69FE10FA0C" box="[380,429,1406,1427]" collectorName="Tersek" location="Aulyekol District" municipality="Naurzum Reserve" pageId="2" pageNumber="29" specimenCount="4" specimenCount-female="2" specimenCount-male="2">2013</materialsCitation>
</date>
,
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,
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;
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,
<collectingMunicipality id="967BACBFFFC0FFCA10C9DE69FCD6FA0C" box="[702,875,1406,1427]" pageId="2" pageNumber="29">Naurzum Reserve</collectingMunicipality>
,
<collectorName id="DB555313FFC0FFCA1102DE69FC0AFA0B" box="[885,951,1406,1428]" pageId="2" pageNumber="29">Tersek</collectorName>
,
<taxonomicName id="B1A04D46FFC0FFCA11B5DE69FBDAFA0C" box="[962,1127,1406,1427]" class="Liliopsida" family="Poaceae" genus="Stipa" kingdom="Plantae" order="Poales" pageId="2" pageNumber="29" phylum="Tracheophyta" rank="species" species="lessingiana">
<emphasis id="44D4EAD7FFC0FFCA11B5DE69FBDAFA0C" box="[962,1127,1406,1427]" italics="true" pageId="2" pageNumber="29">Stipa lessingiana</emphasis>
</taxonomicName>
steppe,
<date id="021E1005FFC0FFCA135BDE8EFE29FA31" box="[300,404,1433,1455]" pageId="2" pageNumber="29" value="1998-05-15">
15.
<collectionCode id="10B1AE00FFC0FFCA133FDE8EFEE1FA30" box="[328,348,1433,1455]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="29" type="Museum">V</collectionCode>
<materialsCitation id="C6C83C98FFC0FFCA1315DE8EFE29FA31" box="[354,404,1433,1454]" collectionCode="T" determinerName="Zalesskaja" pageId="2" pageNumber="29" specimenCount="3">1998</materialsCitation>
</date>
,
<specimenCount id="60A6FD4CFFC0FFCA13E9DE8EFE5CFA30" box="[414,481,1433,1455]" pageId="2" pageNumber="29" type="generic">3 spec.</specimenCount>
(N.
<collectionCode id="10B1AE00FFC0FFCA1071DE8DFDAAFA30" box="[518,535,1434,1455]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="29">T</collectionCode>
.
<determinerName id="83C3DA01FFC0FFCA1057DE8EFD38FA30" box="[544,645,1433,1455]" pageId="2" pageNumber="29">Zalesskaja</determinerName>
det.).
</paragraph>
</subSubSection>
<subSubSection id="3EBA654EFFC0FFCA137EDEA2FCD7F9A7" pageId="2" pageNumber="29" type="distribution">
<paragraph id="761F36C5FFC0FFCA137EDEA2FBDAFA79" blockId="2.[227,1127,1378,1593]" pageId="2" pageNumber="29">
DISTRIBUTION. East-European steppe species.
<collectingRegion id="B464F827FFC0FFCA109BDEA2FCEDFA54" box="[748,848,1461,1483]" country="Russia" name="Volgograd" pageId="2" pageNumber="29">Volgograd</collectingRegion>
and Rostov-on-Don Regions of
<collectingCountry id="0EB77655FFC0FFCA128ADEC7FE83FA79" box="[253,318,1488,1510]" name="Russia" pageId="2" pageNumber="29">Russia</collectingCountry>
, the
<collectingRegion id="B464F827FFC0FFCA131CDEC7FDF1FA79" box="[363,588,1488,1510]" country="Ukraine" name="Dnipropetrovs'k" pageId="2" pageNumber="29">Dnepropetrovsk Region</collectingRegion>
of
<collectingCountry id="0EB77655FFC0FFCA101BDEC7FD05FA79" box="[620,696,1488,1510]" name="Ukraine" pageId="2" pageNumber="29">Ukraine</collectingCountry>
(Zalesskaja, 1978; Zuev &amp; Evsyukov, 2016);
</paragraph>
<paragraph id="761F36C5FFC0FFCA1294DEFBFBDAF99D" blockId="2.[227,1127,1378,1593]" box="[227,1127,1516,1538]" pageId="2" pageNumber="29">
border of the
<collectingRegion id="B464F827FFC0FFCA1310DEFBFDB1F99D" box="[359,524,1516,1538]" country="Russia" name="Orenburg" pageId="2" pageNumber="29">Orenburg Region</collectingRegion>
of
<collectingCountry id="0EB77655FFC0FFCA1058DEFBFDD0F99D" box="[559,621,1516,1538]" name="Russia" pageId="2" pageNumber="29">Russia</collectingCountry>
and the Kostanay Region of
<collectingCountry id="0EB77655FFC0FFCA11FEDEFBFC4BF99D" box="[905,1014,1516,1538]" name="Kazakhstan" pageId="2" pageNumber="29">Kazakhstan</collectingCountry>
(Farzalieva
</paragraph>
<paragraph id="761F36C5FFC0FFCA1294DD1FFBDBF981" blockId="2.[227,1127,1378,1593]" box="[227,1126,1544,1566]" pageId="2" pageNumber="29">
&amp; Esyunin, 2008: as
<taxonomicName id="B1A04D46FFC0FFCA13CBDD1FFDF8F982" box="[444,581,1544,1565]" class="Chilopoda" family="Lithobiidae" genus="Hessebius" kingdom="Animalia" order="Lithobiomorpha" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="44D4EAD7FFC0FFCA13CBDD1FFDA0F982" box="[444,541,1544,1565]" italics="true" pageId="2" pageNumber="29">Hessebius</emphasis>
sp.
</taxonomicName>
).
<collectingCountry id="0EB77655FFC0FFCA102DDD1FFD73F981" box="[602,718,1544,1566]" name="Kazakhstan" pageId="2" pageNumber="29">Kazakhstan</collectingCountry>
:
<collectingRegion id="B464F827FFC0FFCA10ACDD1FFC3BF981" box="[731,902,1544,1566]" country="Kazakhstan" name="Batys Qazaqstan Oblysy" pageId="2" pageNumber="29">West Kazakhstan</collectingRegion>
and Kostanay regions
</paragraph>
<paragraph id="761F36C5FFC0FFCA1294DD34FCD7F9A7" blockId="2.[227,1127,1378,1593]" box="[227,874,1571,1593]" pageId="2" pageNumber="29">(Vsevolodova-Perel, 2009; Bragina, 2012, 2016; Dyachkov, 2019).</paragraph>
</subSubSection>
</treatment>
</document>

277
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C.L. Koch, 1847
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MATERIAL EXAMINED.
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: Auliekol District, Naurzum Reserve,
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Tersek, virgin
<taxonomicName id="B1A04D46FFC1FFCB1306D963FDAAFD16" box="[369,535,628,649]" class="Liliopsida" family="Poaceae" genus="Stipa" kingdom="Plantae" order="Poales" pageId="3" pageNumber="30" phylum="Tracheophyta" rank="species" species="lessingiana">
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steppe, in the soil,
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. Zalesskaya det.); same locality,
</paragraph>
<paragraph id="761F36C5FFC1FFCB1294D998FBDAFD3A" blockId="3.[227,1128,600,978]" box="[227,1127,654,677]" pageId="3" pageNumber="30">
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in soil (
<quantity id="B1589B20FFC1FFCB135AD998FE3AFD3B" box="[301,391,655,676]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="3" pageNumber="30" unit="cm" value="5.0" valueMax="10.0" valueMin="0.0">0–10 cm</quantity>
),
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<collectingDate id="125AE9EDFFC1FFCB13EFD998FDBDFD3B" box="[408,512,655,677]" pageId="3" pageNumber="30" value="2018-05-26">26.V 2018</collectingDate>
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,
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,
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,
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,
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<paragraph id="761F36C5FFC1FFCB1294D9BEFBD5FD5F" blockId="3.[227,1128,600,978]" box="[227,1128,681,705]" pageId="3" pageNumber="30">
<geoCoordinate id="13945002FFC1FFCB1294D9BEFE87FD5F" box="[227,314,681,704]" degrees="63" direction="east" minutes="52" orientation="longitude" pageId="3" pageNumber="30" precision="925" value="63.866665">63°52′ E</geoCoordinate>
,
<date id="021E1005FFC1FFCB1332D9BCFE08FD5F" box="[325,437,683,704]" pageId="3" pageNumber="30" value="2013-04-14">14.IV 2013</date>
,
<specimenCount id="60A6FD4CFFC1FFCB13B6D9BCFE5DFD5F" box="[449,480,683,704]" pageId="3" pageNumber="30" type="female">1♀</specimenCount>
;
<materialsCitation id="C6C83C98FFC1FFCB139AD9BCFC07FD5E" box="[493,954,683,705]" location="Stoun (Kamennoye) Lake" pageId="3" pageNumber="30" specimenCount="1">
<location id="737F601EFFC1FFCB139AD9BCFD54FD5E" LSID="urn:lsid:plazi:treatment:FE0987D3FFC1FFCB1294D93AFCDEFB97:737F601EFFC1FFCB139AD9BCFD54FD5E" box="[493,745,683,705]" name="Stoun (Kamennoye) Lake" pageId="3" pageNumber="30">Stoun (Kamennoye) Lake</location>
, birch forest, in litter
</materialsCitation>
,
<geoCoordinate id="13945002FFC1FFCB11B2D9BDFBA0FD5F" box="[965,1053,682,704]" degrees="53" direction="north" minutes="12" orientation="latitude" pageId="3" pageNumber="30" precision="925" value="53.2">53°12' N</geoCoordinate>
, 63°44'
</paragraph>
<paragraph id="761F36C5FFC1FFCB1294D9D1FBDAFD43" blockId="3.[227,1128,600,978]" box="[227,1127,710,732]" pageId="3" pageNumber="30">
E,
<date id="021E1005FFC1FFCB1377D9D1FED4FD44" box="[256,361,710,732]" pageId="3" pageNumber="30" value="2017-05-16">
16.
<collectionCode id="10B1AE00FFC1FFCB136BD9D1FE8DFD43" box="[284,304,710,732]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="30" type="Museum">V</collectionCode>
2017
</date>
,
<specimenCount id="60A6FD4CFFC1FFCB1301D9D1FE2BFD44" box="[374,406,710,731]" pageId="3" pageNumber="30" type="female">2♀</specimenCount>
;
<materialsCitation id="C6C83C98FFC1FFCB13D5D9D1FCF1FD44" box="[418,844,710,732]" latitude="53.2" longLatPrecision="1078" longitude="63.733334" pageId="3" pageNumber="30" specimenCount="1">
same locality, lakeside,
<geoCoordinate id="13945002FFC1FFCB10F8D9D1FD55FD44" box="[655,744,710,732]" degrees="53" direction="north" minutes="12" orientation="latitude" pageId="3" pageNumber="30" precision="925" value="53.2">53°12' N</geoCoordinate>
,
<geoCoordinate id="13945002FFC1FFCB1082D9D1FCF1FD44" box="[757,844,710,732]" degrees="63" direction="east" minutes="44" orientation="longitude" pageId="3" pageNumber="30" precision="925" value="63.733334">63°44' E</geoCoordinate>
</materialsCitation>
,
<date id="021E1005FFC1FFCB112FD9D1FC7CFD44" box="[856,961,710,732]" pageId="3" pageNumber="30" value="2017-05-16">
16.
<collectionCode id="10B1AE00FFC1FFCB1103D9D1FC35FD43" box="[884,904,710,732]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="30" type="Museum">V</collectionCode>
2017
</date>
,
<specimenCount id="60A6FD4CFFC1FFCB11B9D9D1FC47FD44" box="[974,1018,710,731]" pageId="3" pageNumber="30" type="female">10♀</specimenCount>
; Zatobolsk
</paragraph>
<paragraph id="761F36C5FFC1FFCB1294D9F6FBDBFD68" blockId="3.[227,1128,600,978]" box="[227,1126,737,759]" pageId="3" pageNumber="30">
<materialsCitation id="C6C83C98FFC1FFCB1294D9F6FBDBFD68" box="[227,1126,737,759]" collectingDate="2017-05-14" latitude="53.2" location="Zarechnone Village" longLatPrecision="1078" longitude="63.683334" pageId="3" pageNumber="30" specimenCount="2" specimenCount-female="2">
Village,
<geoCoordinate id="13945002FFC1FFCB134FD9F6FE2FFD68" box="[312,402,737,759]" degrees="53" direction="north" minutes="12" orientation="latitude" pageId="3" pageNumber="30" precision="925" value="53.2">53°12' N</geoCoordinate>
,
<geoCoordinate id="13945002FFC1FFCB13E8D9F6FE4AFD68" box="[415,503,737,759]" degrees="63" direction="east" minutes="41" orientation="longitude" pageId="3" pageNumber="30" precision="925" value="63.683334">63°41' E</geoCoordinate>
,
<date id="021E1005FFC1FFCB1073D9F6FDD3FD69" box="[516,622,737,758]" pageId="3" pageNumber="30" value="2017-05-14">
<collectingDate id="125AE9EDFFC1FFCB1073D9F6FDD3FD69" box="[516,622,737,758]" pageId="3" pageNumber="30" value="2017-05-14">14.V 2017</collectingDate>
</date>
,
<specimenCount id="60A6FD4CFFC1FFCB100CD9F6FD26FD69" box="[635,667,737,758]" pageId="3" pageNumber="30" type="female">2♀</specimenCount>
;
<location id="737F601EFFC1FFCB10DFD9F6FCD6FD68" LSID="urn:lsid:plazi:treatment:FE0987D3FFC1FFCB1294D93AFCDEFB97:737F601EFFC1FFCB10DFD9F6FCD6FD68" box="[680,875,737,759]" latitude="53.2" longLatPrecision="1078" longitude="63.683334" name="Zarechnone Village" pageId="3" pageNumber="30">Zarechnone Village</location>
, garden,
<collectingMethod id="AFE14ED2FFC1FFCB11BFD9F6FBA6FD68" box="[968,1051,737,759]" pageId="3" pageNumber="30">soil trap</collectingMethod>
, 15.VII
</materialsCitation>
</paragraph>
<paragraph id="761F36C5FFC1FFCB1294D9EAFBDAFC8D" blockId="3.[227,1128,600,978]" box="[227,1127,764,787]" pageId="3" pageNumber="30">
2017,
<materialsCitation id="C6C83C98FFC1FFCB1357D9EAFDF4FC8D" box="[288,585,765,787]" collectingDate="2018-04-17" pageId="3" pageNumber="30" specimenCount="1" specimenCount-female="1">
<specimenCount id="60A6FD4CFFC1FFCB1357D9EAFE82FC8D" box="[288,319,765,786]" pageId="3" pageNumber="30" type="female">1♀</specimenCount>
; same locality,
<date id="021E1005FFC1FFCB13AED9EAFDF4FC8D" box="[473,585,765,786]" pageId="3" pageNumber="30" value="2018-04-17">
<collectingDate id="125AE9EDFFC1FFCB13AED9EAFDF4FC8D" box="[473,585,765,786]" pageId="3" pageNumber="30" value="2018-04-17">17.IV 2018</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="C6C83C98FFC1FFCB1023D9EAFBDFFC8D" box="[596,1122,764,787]" latitude="53.216667" location="Kostanay town" pageId="3" pageNumber="30" specimenCount="6" specimenCount-female="6">
<specimenCount id="60A6FD4CFFC1FFCB1023D9EAFDCEFC8C" box="[596,627,765,787]" pageId="3" pageNumber="30" type="female">6♀</specimenCount>
, 2 fragm.;
<location id="737F601EFFC1FFCB10A8D9EAFCCEFC8D" LSID="urn:lsid:plazi:treatment:FE0987D3FFC1FFCB1294D93AFCDEFB97:737F601EFFC1FFCB10A8D9EAFCCEFC8D" box="[735,883,765,786]" latitude="53.216667" name="Kostanay town" pageId="3" pageNumber="30">Kostanay town</location>
, in the garden,
<geoCoordinate id="13945002FFC1FFCB167DD9EBFBDFFC8D" box="[1034,1122,764,786]" degrees="53" direction="north" minutes="13" orientation="latitude" pageId="3" pageNumber="30" precision="925" value="53.216667">53°13' N</geoCoordinate>
</materialsCitation>
,
</paragraph>
<paragraph id="761F36C5FFC1FFCB1294D80FFBDAFCB1" blockId="3.[227,1128,600,978]" box="[227,1127,791,814]" pageId="3" pageNumber="30">
<geoCoordinate id="13945002FFC1FFCB1294D80FFE84FCB2" box="[227,313,792,814]" degrees="63" direction="east" minutes="37" orientation="longitude" pageId="3" pageNumber="30" precision="925" value="63.616665">63°37' E</geoCoordinate>
,
<date id="021E1005FFC1FFCB1334D80FFE11FCB2" box="[323,428,792,813]" pageId="3" pageNumber="30" value="2017-05-07">
07.
<collectionCode id="10B1AE00FFC1FFCB1317D80FFECEFCB2" box="[352,371,792,813]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="30" type="Museum">V</collectionCode>
2017
</date>
,
<specimenCount id="60A6FD4CFFC1FFCB13C0D80FFE66FCB2" box="[439,475,792,813]" pageId="3" pageNumber="30" type="female">3 ♀</specimenCount>
,
<specimenCount id="60A6FD4CFFC1FFCB139FD80FFD99FCB1" box="[488,548,792,814]" pageId="3" pageNumber="30" type="generic">3 juv.</specimenCount>
;
<materialsCitation id="C6C83C98FFC1FFCB105AD80FFC04FCB1" box="[557,953,792,814]" location="Amangeldy District" municipality="Amangeldy Village" pageId="3" pageNumber="30" specimenCount="1">
<location id="737F601EFFC1FFCB105AD80FFD4DFCB1" LSID="urn:lsid:plazi:treatment:FE0987D3FFC1FFCB1294D93AFCDEFB97:737F601EFFC1FFCB105AD80FFD4DFCB1" box="[557,752,792,814]" municipality="Amangeldy Village" name="Amangeldy District" pageId="3" pageNumber="30">Amangeldy District</location>
,
<collectingMunicipality id="967BACBFFFC1FFCB108DD80FFC04FCB1" box="[762,953,792,814]" pageId="3" pageNumber="30">Amangeldy Village</collectingMunicipality>
</materialsCitation>
,
<geoCoordinate id="13945002FFC1FFCB11B3D800FBA0FCB2" box="[964,1053,791,814]" degrees="50" direction="north" minutes="11" orientation="latitude" pageId="3" pageNumber="30" precision="925" value="50.183334">50°11′ N</geoCoordinate>
, 65°11′
</paragraph>
<paragraph id="761F36C5FFC1FFCB1294D823FBDAFCD6" blockId="3.[227,1128,600,978]" box="[227,1127,819,841]" pageId="3" pageNumber="30">
E,
<date id="021E1005FFC1FFCB1376D824FED0FCD7" box="[257,365,819,841]" pageId="3" pageNumber="30" value="2017-05-16">
16.
<collectionCode id="10B1AE00FFC1FFCB136AD824FE8CFCD6" box="[285,305,819,841]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="30" type="Museum">V</collectionCode>
<materialsCitation id="C6C83C98FFC1FFCB134CD824FED0FCD7" box="[315,365,819,840]" latitude="53.45" location="Karabalyk District" longLatPrecision="1077" longitude="63.316666" municipality="Ozernoye Village" pageId="3" pageNumber="30" specimenCount="1" specimenCount-female="1">2017</materialsCitation>
</date>
,
<specimenCount id="60A6FD4CFFC1FFCB130DD824FE24FCD7" box="[378,409,819,840]" pageId="3" pageNumber="30" type="female">1♀</specimenCount>
;
<location id="737F601EFFC1FFCB13DED824FDDDFCD6" LSID="urn:lsid:plazi:treatment:FE0987D3FFC1FFCB1294D93AFCDEFB97:737F601EFFC1FFCB13DED824FDDDFCD6" box="[425,608,819,841]" latitude="53.45" longLatPrecision="1077" longitude="63.316666" municipality="Ozernoye Village" name="Karabalyk District" pageId="3" pageNumber="30">Karabalyk District</location>
,
<collectingMunicipality id="967BACBFFFC1FFCB1019D824FCA3FCD6" box="[622,798,819,841]" pageId="3" pageNumber="30">Ozernoye Village</collectingMunicipality>
, in the soil,
<geoCoordinate id="13945002FFC1FFCB11D6D824FC41FCD6" box="[929,1020,819,841]" degrees="53" direction="north" minutes="27" orientation="latitude" pageId="3" pageNumber="30" precision="925" value="53.45">53°27' N</geoCoordinate>
,
<geoCoordinate id="13945002FFC1FFCB167DD824FBDEFCD6" box="[1034,1123,819,841]" degrees="63" direction="east" minutes="19" orientation="longitude" pageId="3" pageNumber="30" precision="925" value="63.316666">63°19' E</geoCoordinate>
,
</paragraph>
<paragraph id="761F36C5FFC1FFCB1294D858FCA8FCE0" blockId="3.[227,1128,600,978]" pageId="3" pageNumber="30">
<date id="021E1005FFC1FFCB1294D858FEF6FCFB" box="[227,331,847,868]" pageId="3" pageNumber="30" value="2017-05-12">
12.
<collectionCode id="10B1AE00FFC1FFCB1288D858FEAFFCFB" box="[255,274,847,868]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="30" type="Museum">V</collectionCode>
2017
</date>
,
<specimenCount id="60A6FD4CFFC1FFCB1321D858FEC8FCFA" box="[342,373,847,869]" pageId="3" pageNumber="30" type="female">5♀</specimenCount>
; Auliekol District, Naurzum Reserve, Tersek,
<taxonomicName id="B1A04D46FFC1FFCB114ED858FC31FCFB" box="[825,908,847,868]" class="Liliopsida" family="Poaceae" genus="Triticum" kingdom="Plantae" order="Poales" pageId="3" pageNumber="30" phylum="Tracheophyta" rank="genus">
<emphasis id="44D4EAD7FFC1FFCB114ED858FC31FCFB" box="[825,908,847,868]" italics="true" pageId="3" pageNumber="30">Tríticum</emphasis>
</taxonomicName>
field near virgin
<taxonomicName id="B1A04D46FFC1FFCB1642D858FEEDFCE0" class="Liliopsida" family="Poaceae" genus="Stipa" kingdom="Plantae" order="Poales" pageId="3" pageNumber="30" phylum="Tracheophyta" rank="species" species="lessingiana">
<emphasis id="44D4EAD7FFC1FFCB1642D858FEEDFCE0" italics="true" pageId="3" pageNumber="30">Stipa lessingiana</emphasis>
</taxonomicName>
steppe, in the soil (
<quantity id="B1589B20FFC1FFCB107CD87DFDD2FCE0" box="[523,623,874,895]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="3" pageNumber="30" unit="cm" value="25.0" valueMax="30.0" valueMin="20.0">20–30 cm</quantity>
),
<date id="021E1005FFC1FFCB10F7D87DFD5AFCE0" box="[640,743,874,895]" pageId="3" pageNumber="30" value="2018-05-18">
18.
<collectionCode id="10B1AE00FFC1FFCB10ECD87DFD12FCE0" box="[667,687,874,895]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="30" type="Museum">V</collectionCode>
2018
</date>
,
<specimenCount id="60A6FD4CFFC1FFCB1086D87DFCACFCE0" box="[753,785,874,895]" pageId="3" pageNumber="30" type="female">3♀</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="3EBA654EFFC1FFCB137ED891FCDEFB97" pageId="3" pageNumber="30" type="distribution">
<paragraph id="761F36C5FFC1FFCB137ED891FBDAFC04" blockId="3.[227,1128,600,978]" box="[265,1127,901,923]" pageId="3" pageNumber="30">DISTRIBUTION. West-Palearctic polyzonal species (Barber &amp; Jones, 1999; Barber,</paragraph>
<paragraph id="761F36C5FFC1FFCB1294D8B6FD0DFC4E" blockId="3.[227,1128,600,978]" pageId="3" pageNumber="30">
<materialsCitation id="C6C83C98FFC1FFCB1294D8B6FD11FC4E" collectorName="Dyachkov &amp; Tuf" country="Kazakhstan" location="East Kazakhstan" pageId="3" pageNumber="30" specimenCount="1" stateProvince="Shyghys Qazaqstan Oblysy">
2009; Nefediev
<emphasis id="44D4EAD7FFC1FFCB13F2D8B6FE0EFC29" box="[389,435,929,950]" italics="true" pageId="3" pageNumber="30">et al</emphasis>
., 2017).
<collectingCountry id="0EB77655FFC1FFCB1066D8B6FD38FC28" box="[529,645,929,951]" name="Kazakhstan" pageId="3" pageNumber="30">Kazakhstan</collectingCountry>
:
<collectingRegion id="B464F827FFC1FFCB10E6D8B6FC8AFC28" box="[657,823,929,951]" country="Kazakhstan" name="Shyghys Qazaqstan Oblysy" pageId="3" pageNumber="30">East Kazakhstan</collectingRegion>
, Karaganda,
<collectingRegion id="B464F827FFC1FFCB11CBD8B6FBDAFC28" box="[956,1127,929,951]" country="Kazakhstan" name="Batys Qazaqstan Oblysy" pageId="3" pageNumber="30">West Kazakhstan</collectingRegion>
and Kostanay regions (
<collectorName id="DB555313FFC1FFCB13C8D8ABFD9EFC4D" box="[447,547,956,978]" pageId="3" pageNumber="30">Dyachkov</collectorName>
&amp;
<collectorName id="DB555313FFC1FFCB1035D8ABFDDAFC4D" box="[578,615,956,978]" pageId="3" pageNumber="30">Tuf</collectorName>
, 2019)
</materialsCitation>
.
</paragraph>
<paragraph id="761F36C5FFC1FFCB1391D8E4FCDEFB97" blockId="3.[486,867,1011,1033]" box="[486,867,1011,1033]" pageId="3" pageNumber="30">
<emphasis id="44D4EAD7FFC1FFCB1391D8E4FCDEFB97" bold="true" box="[486,867,1011,1033]" pageId="3" pageNumber="30">
Family
<taxonomicName id="B1A04D46FFC1FFCB1043D8E4FCDEFB97" authority="Bollman, 1893" authorityName="Bollman" authorityYear="1893" box="[564,867,1011,1033]" class="Chilopoda" family="Himantariidae" kingdom="Animalia" order="Geophilomorpha" pageId="3" pageNumber="30" phylum="Arthropoda" rank="family">Himantariidae Bollman, 1893</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

129
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Zalesskaja, 1978
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MATERIAL EXAMINED.
<emphasis id="44D4EAD7FFC1FFCB1057DA25FD6AFED7" bold="true" box="[544,727,306,328]" pageId="3" pageNumber="30">Kostanay Region</emphasis>
: Aulyekol District, Naurzum Reserve,
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<paragraph id="761F36C5FFC1FFCB1294DA5AFC76FEFC" blockId="3.[227,1127,306,410]" box="[227,971,333,355]" pageId="3" pageNumber="30">
<materialsCitation id="C6C83C98FFC1FFCB1294DA5AFC7BFEFC" box="[227,966,333,355]" collectingDate="1998-07-01" collectionCode="T" determinerName="Zalesskaja" pageId="3" pageNumber="30" specimenCount="2">
Tersek, the
<taxonomicName id="B1A04D46FFC1FFCB1321DA59FE46FEFD" box="[342,507,333,355]" class="Liliopsida" family="Poaceae" genus="Stipa" kingdom="Plantae" order="Poales" pageId="3" pageNumber="30" phylum="Tracheophyta" rank="species" species="lessingiana">
<emphasis id="44D4EAD7FFC1FFCB1321DA59FE46FEFD" box="[342,507,333,355]" italics="true" pageId="3" pageNumber="30">Stipa lessingiana</emphasis>
</taxonomicName>
steppe,
<date id="021E1005FFC1FFCB103DDA59FD08FEFC" box="[586,693,334,355]" pageId="3" pageNumber="30" value="1998-07-01">
<collectingDate id="125AE9EDFFC1FFCB103DDA59FD08FEFC" box="[586,693,334,355]" pageId="3" pageNumber="30" value="1998-07-01">1.VII 1998</collectingDate>
</date>
,
<specimenCount id="60A6FD4CFFC1FFCB10C8DA59FD53FEFC" box="[703,750,334,355]" pageId="3" pageNumber="30" type="generic">2 ex.</specimenCount>
(N.
<collectionCode id="10B1AE00FFC1FFCB1164DA59FC99FEFC" box="[787,804,334,355]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="30">T</collectionCode>
.
<determinerName id="83C3DA01FFC1FFCB115ADA5AFC2FFEFC" box="[813,914,333,355]" pageId="3" pageNumber="30">Zalesskaja</determinerName>
det.)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3EBA654EFFC1FFCB137EDA7EFCD8FE4F" pageId="3" pageNumber="30" type="distribution">
<paragraph id="761F36C5FFC1FFCB137EDA7EFBDAFEE1" blockId="3.[227,1127,306,410]" box="[265,1127,361,383]" pageId="3" pageNumber="30">
DISTRIBUTION.
<collectingCountry id="0EB77655FFC1FFCB13B7DA7EFD8CFEE1" box="[448,561,361,382]" name="Kyrgyzstan" pageId="3" pageNumber="30">Kyrgyzstan</collectingCountry>
,
<collectingCountry id="0EB77655FFC1FFCB104ADA7EFD20FEE0" box="[573,669,361,383]" name="Tajikistan" pageId="3" pageNumber="30">Tajikistan</collectingCountry>
(Zalesskaja, 1978) and
<collectingCountry id="0EB77655FFC1FFCB11FADA7EFBBCFEE0" box="[909,1025,361,383]" name="Kazakhstan" pageId="3" pageNumber="30">Kazakhstan</collectingCountry>
: Kostanay
</paragraph>
<paragraph id="761F36C5FFC1FFCB1294DA93FC54FE06" blockId="3.[227,1127,306,410]" box="[227,1001,388,410]" pageId="3" pageNumber="30">
(Bragina, 2012, 2016) and
<collectingRegion id="B464F827FFC1FFCB1392DA93FD90FE05" box="[485,557,388,410]" country="Kazakhstan" name="Almaty Oblysy" pageId="3" pageNumber="30">Almaty</collectingRegion>
regions (Dyachkov, 2019: as
<taxonomicName id="B1A04D46FFC1FFCB113ADA92FC61FE06" authorityName="Zalesskaja" authorityYear="1978" box="[845,988,388,410]" class="Chilopoda" family="Lithobiidae" genus="Hessebius" isUncertain="true" kingdom="Animalia" order="Lithobiomorpha" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="plumatus">
<emphasis id="44D4EAD7FFC1FFCB113ADA92FCE3FE05" box="[845,862,389,410]" italics="true" pageId="3" pageNumber="30">H</emphasis>
. cf
<emphasis id="44D4EAD7FFC1FFCB11F3DA93FC61FE06" box="[900,988,388,409]" italics="true" pageId="3" pageNumber="30">plumatus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="761F36C5FFC1FFCB1392DAACFCD8FE4F" blockId="3.[485,869,443,465]" box="[485,869,443,465]" pageId="3" pageNumber="30">
<emphasis id="44D4EAD7FFC1FFCB1392DAACFCD8FE4F" bold="true" box="[485,869,443,465]" pageId="3" pageNumber="30">
Order
<taxonomicName id="B1A04D46FFC1FFCB105CDAACFCD8FE4F" authority="Pocock, 1896" authorityName="Pocock" authorityYear="1896" box="[555,869,443,465]" class="Chilopoda" kingdom="Animalia" order="Geophilomorpha" pageId="3" pageNumber="30" phylum="Arthropoda" rank="order">Geophilomorpha Pocock, 1896</taxonomicName>
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<paragraph id="761F36C5FFC1FFCB1388DAF6FCF7FE69" blockId="3.[511,842,481,503]" box="[511,842,481,503]" pageId="3" pageNumber="30">
<emphasis id="44D4EAD7FFC1FFCB1388DAF6FCF7FE69" bold="true" box="[511,842,481,503]" pageId="3" pageNumber="30">
Family
<taxonomicName id="B1A04D46FFC1FFCB103ADAF6FCF7FE69" authority="Leach, 1816" authorityName="Leach" authorityYear="1816" box="[589,842,481,503]" class="Chilopoda" family="Geophilidae" kingdom="Animalia" order="Geophilomorpha" pageId="3" pageNumber="30" phylum="Arthropoda" rank="family">Geophilidae Leach, 1816</taxonomicName>
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<emphasis id="44D4EAD7FFC1FFCB1294DF28FE7CFBCA" bold="true" box="[227,449,1087,1109]" italics="true" pageId="3" pageNumber="30">Polyporogaster porosa</emphasis>
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MATERIAL EXAMINED.
<emphasis id="44D4EAD7FFC1FFCB106FDF7CFD74FB1F" bold="true" box="[536,713,1130,1152]" pageId="3" pageNumber="30">Kostanay Region</emphasis>
: Amangeldy District, Altyn Dala Reserve,
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<materialsCitation id="C6C83C98FFC1FFCB1294DF91FBF9FB04" box="[227,1092,1158,1180]" collectingDate="2013-07-04" location="Ulyzhilanshik" municipality="Rakhmet Village" pageId="3" pageNumber="30" specimenCount="1" specimenCount-female="1">
<location id="737F601EFFC1FFCB1294DF91FED7FB03" LSID="urn:lsid:plazi:treatment:FE0987D3FFC1FFCB1294DF28FE6FFA96:737F601EFFC1FFCB1294DF91FED7FB03" box="[227,362,1158,1180]" municipality="Rakhmet Village" name="Ulyzhilanshik" pageId="3" pageNumber="30">Ulyzhilanshik</location>
site, near destroyed
<collectingMunicipality id="967BACBFFFC1FFCB1047DF91FD68FB03" box="[560,725,1158,1180]" pageId="3" pageNumber="30">Rakhmet Village</collectingMunicipality>
, steppe, grazing site,
<date id="021E1005FFC1FFCB11D4DF91FBA6FB04" box="[931,1051,1158,1179]" pageId="3" pageNumber="30" value="2013-07-04">
<collectingDate id="125AE9EDFFC1FFCB11D4DF91FBA6FB04" box="[931,1051,1158,1179]" pageId="3" pageNumber="30" value="2013-07-04">04.VII 2013</collectingDate>
</date>
,
<specimenCount id="60A6FD4CFFC1FFCB1652DF91FBF9FB04" box="[1061,1092,1158,1179]" pageId="3" pageNumber="30" type="female">1♀</specimenCount>
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.
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<paragraph id="761F36C5FFC1FFCB137EDFB6FBDAFB28" blockId="3.[227,1127,1130,1289]" box="[265,1127,1185,1207]" pageId="3" pageNumber="30">
REMARKS. Central Asian species.
<collectingCountry id="0EB77655FFC1FFCB1006DFB6FD60FB28" box="[625,733,1185,1207]" name="Uzbekistan" pageId="3" pageNumber="30">Uzbekistan</collectingCountry>
(Sseliwanoff, 1881; Lignau, 1929) and
</paragraph>
<paragraph id="761F36C5FFC1FFCB1294DFABFBDAFB72" blockId="3.[227,1127,1130,1289]" pageId="3" pageNumber="30">
<collectingCountry id="0EB77655FFC1FFCB1294DFABFEEAFB4D" box="[227,343,1212,1234]" name="Kazakhstan" pageId="3" pageNumber="30">Kazakhstan</collectingCountry>
: Mangystau, Kyzylorda, Turkistan and
<collectingRegion id="B464F827FFC1FFCB10AADFABFC98FB4D" box="[733,805,1212,1234]" country="Kazakhstan" name="Almaty Oblysy" pageId="3" pageNumber="30">Almaty</collectingRegion>
regions (Dyachkov, 2020). This species, the genus
<taxonomicName id="B1A04D46FFC1FFCB13EFDFCFFD90FB72" authorityName="Verhoeff" authorityYear="1899" box="[408,557,1240,1261]" class="Chilopoda" family="Himantariidae" genus="Polyporogaster" kingdom="Animalia" order="Geophilomorpha" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="44D4EAD7FFC1FFCB13EFDFCFFD90FB72" box="[408,557,1240,1261]" italics="true" pageId="3" pageNumber="30">Polyporogaster</emphasis>
</taxonomicName>
and family
<taxonomicName id="B1A04D46FFC1FFCB10D4DFCFFC91FB71" authorityName="Bollman" authorityYear="1893" box="[675,812,1240,1262]" class="Chilopoda" family="Himantariidae" kingdom="Animalia" order="Geophilomorpha" pageId="3" pageNumber="30" phylum="Arthropoda" rank="family">Himantariidae</taxonomicName>
are recorded from the Kostanay
</paragraph>
<paragraph id="761F36C5FFC1FFCB1294DFE4FE6FFA96" blockId="3.[227,1127,1130,1289]" box="[227,466,1267,1289]" pageId="3" pageNumber="30">Region for the first time.</paragraph>
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232
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<mods:title id="091301ED79F79624D6C1F6DE542A7B84">NEW DATA ON THE CENTIPEDE FAUNA (MYRIAPODA: CHILOPODA) OF KOSTANAY REGION, KAZAKHSTAN</mods:title>
</mods:titleInfo>
<mods:name id="A15C0EB5D38BB7C9F4A8DD0236523820" type="personal">
<mods:role id="242638142AB3FC2C690E6BB07B48B3B2">
<mods:roleTerm id="0A7B9EBDE876EFCE302CEBFFF8478645">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C25FE83A59303CF8ED677B8780188E6E">Bragina, T. M.</mods:namePart>
</mods:name>
<mods:name id="042C51B9B6C699DFB3F2CBA169FF50D9" type="personal">
<mods:role id="CBF471E03E3C1D246E73EF5D1B3F6AEE">
<mods:roleTerm id="E5969E4DD23F4FF0BEAF9C625C037856">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="509A229C8C815002A844D039E4873A68">Dyachkov, Yu. V.</mods:namePart>
</mods:name>
<mods:name id="9F05980447E4776BDBDAC0BB7BC8E7C3" type="personal">
<mods:role id="D8B6B5F19DDE4AD5027B5B9B351349B3">
<mods:roleTerm id="5B05E52D0991E285FADC5E44D644B409">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A2ACB876E37F9D8294CE3C89348BF39A">Farzalieva, G. Sh.</mods:namePart>
</mods:name>
<mods:typeOfResource id="EC2F2E96B0D7B187F92BA1E361FF0683">text</mods:typeOfResource>
<mods:relatedItem id="1D5D4B5BD1BD15B196EC5629DA5BD0E6" type="host">
<mods:titleInfo id="788F260EF219590A2040B41579B007C2">
<mods:title id="30C10535C0BD3166DCCEACF1EBE83B19">Far Eastern Entomologist</mods:title>
</mods:titleInfo>
<mods:part id="13B380C858F7F22D587D8A4570593F83">
<mods:date id="25A4ED297ADDA60E406A3B6940D536E0">2020</mods:date>
<mods:detail id="385C828526F139B511EA023DA7EA1ED6" type="pubDate">
<mods:number id="3D8FCD6571DA00F919C1BF87D6441B34">2020-05-01</mods:number>
</mods:detail>
<mods:detail id="52D8F6A06D58E32723DFF4A5D364A980" type="volume">
<mods:number id="4BCD59F1D3BDA652265FCD977A1C8A37">406</mods:number>
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<mods:start id="E098E8CFF893B3848136B5BC175890BC">27</mods:start>
<mods:end id="4CAA0830FA19A1EFF9756A52ACF36816">32</mods:end>
</mods:extent>
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<mods:location id="D769B873E1E18A198BDD0905EC550189">
<mods:url id="164CA90C4D261755A1A9F2EBFEF20E38">http://dx.doi.org/10.25221/fee.406.4</mods:url>
</mods:location>
<mods:classification id="596A018EC4DB0802EA36CB123425614E">journal article</mods:classification>
<mods:identifier id="C37C05EB14B970A47B18372851944147" type="DOI">10.25221/fee.406.4</mods:identifier>
<mods:identifier id="A513512FD869B262F17A9D540BE6F846" type="ISSN">2713-2196</mods:identifier>
<mods:identifier id="BFB2195C66522A00839DA1637679912E" type="Zenodo-Dep">7165314</mods:identifier>
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</mods:mods>
<treatment id="FE0987D3FFC3FFC91294D9CBFCD8FB09" LSID="urn:lsid:plazi:treatment:FE0987D3FFC3FFC91294D9CBFCD8FB09" httpUri="http://treatment.plazi.org/id/FE0987D3FFC3FFC91294D9CBFCD8FB09" lastPageNumber="28" pageId="1" pageNumber="28">
<subSubSection id="3EBA654EFFC3FFC91294D9CBFDDAFD6D" box="[227,615,732,754]" pageId="1" pageNumber="28" type="nomenclature">
<paragraph id="761F36C5FFC3FFC91294D9CBFDDAFD6D" blockId="1.[227,615,732,754]" box="[227,615,732,754]" pageId="1" pageNumber="28">
<heading id="2D5781A9FFC3FFC91294D9CBFDDAFD6D" box="[227,615,732,754]" centered="true" fontSize="9" level="2" pageId="1" pageNumber="28" reason="2">
<taxonomicName id="B1A04D46FFC3FFC91294D9CBFDDAFD6D" ID-CoL="4W38V" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[227,615,732,754]" class="Chilopoda" family="Scutigeridae" genus="Scutigera" kingdom="Animalia" order="Scutigeromorpha" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="coleoptrata">
<emphasis id="44D4EAD7FFC3FFC91294D9CBFDDAFD6D" bold="true" box="[227,615,732,754]" pageId="1" pageNumber="28">
<emphasis id="44D4EAD7FFC3FFC91294D9CBFE0BFD6D" bold="true" box="[227,438,732,754]" italics="true" pageId="1" pageNumber="28">Scutigera coleoptrata</emphasis>
(Linnaeus, 1758)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3EBA654EFFC3FFC9137ED81FFDFCFC43" pageId="1" pageNumber="28" type="materials_examined">
<paragraph id="761F36C5FFC3FFC9137ED81FFBDAFC82" blockId="1.[227,1127,775,1044]" box="[265,1127,775,797]" pageId="1" pageNumber="28">
MATERIAL EXAMINED.
<emphasis id="44D4EAD7FFC3FFC91053D810FD67FC82" bold="true" box="[548,730,775,797]" pageId="1" pageNumber="28">Kostanay Region</emphasis>
: Naurzum District, Naurzum Reserve,
</paragraph>
<paragraph id="761F36C5FFC3FFC91294D834FBDAFCA7" blockId="1.[227,1127,775,1044]" box="[227,1127,803,825]" pageId="1" pageNumber="28">
<materialsCitation id="C6C83C98FFC3FFC91294D834FD70FCA7" box="[227,717,803,825]" collectionCode="V" location="Naurzum" pageId="1" pageNumber="28" specimenCount="1">
<location id="737F601EFFC3FFC91294D834FE81FCA7" LSID="urn:lsid:plazi:treatment:FE0987D3FFC3FFC91294D9CBFCD8FB09:737F601EFFC3FFC91294D834FE81FCA7" box="[227,316,803,824]" name="Naurzum" pageId="1" pageNumber="28">Naurzum</location>
site, inspector’s house “Garden”,
<date id="021E1005FFC3FFC910EAD834FD70FCA7" box="[669,717,803,824]" pageId="1" pageNumber="28" value="2014-05-04">
04.
<collectionCode id="10B1AE00FFC3FFC910CDD834FD70FCA7" box="[698,717,803,824]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="28" type="Museum">V</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="C6C83C98FFC3FFC910A0D834FBDAFCA7" box="[727,1127,803,825]" collectionCode="VI" location="Karamendy Village" pageId="1" pageNumber="28" specimenCount="1">
2014,
<specimenCount id="60A6FD4CFFC3FFC9116FD834FCF2FCA7" box="[792,847,803,824]" pageId="1" pageNumber="28" type="generic">1 ex.</specimenCount>
;
<location id="737F601EFFC3FFC9112AD834FB9CFCA6" LSID="urn:lsid:plazi:treatment:FE0987D3FFC3FFC91294D9CBFCD8FB09:737F601EFFC3FFC9112AD834FB9CFCA6" box="[861,1057,803,825]" name="Karamendy Village" pageId="1" pageNumber="28">Karamendy Village</location>
, 20.
<collectionCode id="10B1AE00FFC3FFC9163BD834FBDAFCA7" box="[1100,1127,803,824]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="1" pageNumber="28">VI</collectionCode>
</materialsCitation>
</paragraph>
<paragraph id="761F36C5FFC3FFC91294D829FBD9FCF0" blockId="1.[227,1127,775,1044]" pageId="1" pageNumber="28">
2017,
<specimenCount id="60A6FD4CFFC3FFC91353D829FED9FCCB" box="[292,356,830,852]" pageId="1" pageNumber="28" type="generic">1 juv.</specimenCount>
; same place,
<date id="021E1005FFC3FFC91383D829FDCCFCCC" box="[500,625,830,851]" pageId="1" pageNumber="28" value="2017-07-30">30.VII 2017</date>
,
<materialsCitation id="C6C83C98FFC3FFC910F7D829FCC8FCCC" box="[640,885,830,852]" location="Kostanay town" pageId="1" pageNumber="28" specimenCount="1" specimenCount-female="1">
<specimenCount id="60A6FD4CFFC3FFC910F7D829FD1DFCCC" box="[640,672,830,851]" pageId="1" pageNumber="28" type="female">1♀</specimenCount>
;
<location id="737F601EFFC3FFC910C6D828FCF4FCCB" LSID="urn:lsid:plazi:treatment:FE0987D3FFC3FFC91294D9CBFCD8FB09:737F601EFFC3FFC910C6D828FCF4FCCB" box="[689,841,831,852]" name="Kostanay town" pageId="1" pageNumber="28">Kostanay town</location>
,
<date id="021E1005FFC3FFC9112FD829FCC8FCCC" box="[856,885,830,851]" pageId="1" pageNumber="28" value="2014-05-17">17.</date>
</materialsCitation>
<collectionCode id="10B1AE00FFC3FFC91103D829FC35FCCC" box="[884,904,830,851]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="28" type="Museum">V</collectionCode>
2014,
<materialsCitation id="C6C83C98FFC3FFC911A2D829FDAEFCF0" collectingDate="2017-02-20" location="Kostanay town" pageId="1" pageNumber="28" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="60A6FD4CFFC3FFC911A2D829FC48FCCC" box="[981,1013,830,851]" pageId="1" pageNumber="28" type="male">1♂</specimenCount>
,
<specimenCount id="60A6FD4CFFC3FFC91672D829FB99FCCC" box="[1029,1060,830,851]" pageId="1" pageNumber="28" type="female">1♀</specimenCount>
; same locality, in the flat,
<date id="021E1005FFC3FFC913DDD84DFDAEFCF0" box="[426,531,858,879]" pageId="1" pageNumber="28" value="2017-02-20">
<collectingDate id="125AE9EDFFC3FFC913DDD84DFDAEFCF0" box="[426,531,858,879]" pageId="1" pageNumber="28" value="2017-02-20">20.II 2017</collectingDate>
</date>
</materialsCitation>
,
<specimenCount id="60A6FD4CFFC3FFC91057D84DFD82FCF0" box="[544,575,858,879]" pageId="1" pageNumber="28" type="female">1♀</specimenCount>
; in the house,
<date id="021E1005FFC3FFC910A2D84EFC82FCF0" box="[725,831,857,879]" pageId="1" pageNumber="28" value="2017-05-15">
15.
<collectionCode id="10B1AE00FFC3FFC91086D84EFCB8FCF0" box="[753,773,857,879]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="28" type="Museum">V</collectionCode>
2017
</date>
,
<materialsCitation id="C6C83C98FFC3FFC9113BD84DFBD9FCF0" box="[844,1124,857,879]" collectingDate="2017-02-20" collectionCode="V" location="Kostanay town" pageId="1" pageNumber="28" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<specimenCount id="60A6FD4CFFC3FFC9113BD84DFCD4FCF0" box="[844,873,858,879]" pageId="1" pageNumber="28" type="male">1♂</specimenCount>
,
<specimenCount id="60A6FD4CFFC3FFC9110ED84DFC2BFCF0" box="[889,918,858,879]" pageId="1" pageNumber="28" type="female">2♀</specimenCount>
; same locality, 15.
<collectionCode id="10B1AE00FFC3FFC91626D84EFBD9FCF0" box="[1105,1124,857,879]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="28" type="Museum">V</collectionCode>
</materialsCitation>
</paragraph>
<paragraph id="761F36C5FFC3FFC91294D862FBDAFC39" blockId="1.[227,1127,775,1044]" pageId="1" pageNumber="28">
2017,
<materialsCitation id="C6C83C98FFC3FFC91357D862FD49FC15" box="[288,756,885,907]" collectingDate="2017-03-31" pageId="1" pageNumber="28" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="60A6FD4CFFC3FFC91357D862FE82FC15" box="[288,319,885,906]" pageId="1" pageNumber="28" type="male">1♂</specimenCount>
,
<specimenCount id="60A6FD4CFFC3FFC9133CD862FED7FC15" box="[331,362,885,906]" pageId="1" pageNumber="28" type="female">1♀</specimenCount>
; same locality, in the house,
<date id="021E1005FFC3FFC910F2D862FD49FC15" box="[645,756,885,906]" pageId="1" pageNumber="28" value="2017-03-31">
<collectingDate id="125AE9EDFFC3FFC910F2D862FD49FC15" box="[645,756,885,906]" pageId="1" pageNumber="28" value="2017-03-31">31.III 2017</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="C6C83C98FFC3FFC91088D862FE31FC3A" collectingDate="2018-07-26" location="Zarechnone Village" pageId="1" pageNumber="28" specimenCount="1" specimenCount-male="1">
<specimenCount id="60A6FD4CFFC3FFC91088D862FCA2FC15" box="[767,799,885,906]" pageId="1" pageNumber="28" type="male">1♂</specimenCount>
;
<location id="737F601EFFC3FFC9115DD862FC56FC14" LSID="urn:lsid:plazi:treatment:FE0987D3FFC3FFC91294D9CBFCD8FB09:737F601EFFC3FFC9115DD862FC56FC14" box="[810,1003,885,907]" name="Zarechnone Village" pageId="1" pageNumber="28">Zarechnone Village</location>
, garden,
<collectingMethod id="AFE14ED2FFC3FFC91633D862FEB6FC39" pageId="1" pageNumber="28">soil trap</collectingMethod>
,
<date id="021E1005FFC3FFC91363D887FE31FC3A" box="[276,396,912,934]" pageId="1" pageNumber="28" value="2018-07-26">
<collectingDate id="125AE9EDFFC3FFC91363D887FE31FC3A" box="[276,396,912,934]" pageId="1" pageNumber="28" value="2018-07-26">26.VII 2018</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="C6C83C98FFC3FFC913E1D887FC2DFC3A" box="[406,912,912,934]" collectingDate="2017-04-18" location="Kamysty District" municipality="Kamysty Village" pageId="1" pageNumber="28" specimenCount="1" specimenCount-female="1">
<specimenCount id="60A6FD4CFFC3FFC913E1D887FE08FC3A" box="[406,437,912,933]" pageId="1" pageNumber="28" type="female">1♀</specimenCount>
;
<location id="737F601EFFC3FFC913B7D886FDD5FC39" LSID="urn:lsid:plazi:treatment:FE0987D3FFC3FFC91294D9CBFCD8FB09:737F601EFFC3FFC913B7D886FDD5FC39" box="[448,616,912,934]" municipality="Kamysty Village" name="Kamysty District" pageId="1" pageNumber="28">Kamysty District</location>
,
<collectingMunicipality id="967BACBFFFC3FFC91006D886FCAAFC39" box="[625,791,912,934]" pageId="1" pageNumber="28">Kamysty Village</collectingMunicipality>
,
<date id="021E1005FFC3FFC91157D887FC2DFC3A" box="[800,912,912,933]" pageId="1" pageNumber="28" value="2017-04-18">
<collectingDate id="125AE9EDFFC3FFC91157D887FC2DFC3A" box="[800,912,912,933]" pageId="1" pageNumber="28" value="2017-04-18">18.IV 2017</collectingDate>
</date>
</materialsCitation>
,
<specimenCount id="60A6FD4CFFC3FFC911EDD887FC07FC3A" box="[922,954,912,933]" pageId="1" pageNumber="28" type="male">2♂</specimenCount>
,
<specimenCount id="60A6FD4CFFC3FFC911B3D887FC5FFC3A" box="[964,994,912,933]" pageId="1" pageNumber="28" type="female">1♀</specimenCount>
; Denisovskyi
</paragraph>
<paragraph id="761F36C5FFC3FFC91294D8BCFBDAFC5E" blockId="1.[227,1127,775,1044]" box="[227,1127,939,961]" pageId="1" pageNumber="28">
<materialsCitation id="C6C83C98FFC3FFC91294D8BCFBDEFC5E" box="[227,1123,939,961]" collectingDate="2014-05-04" county="Amangeldy District" location="District" municipality="Ayat River" pageId="1" pageNumber="28" specimenCount="1">
<location id="737F601EFFC3FFC91294D8BCFE93FC5E" LSID="urn:lsid:plazi:treatment:FE0987D3FFC3FFC91294D9CBFCD8FB09:737F601EFFC3FFC91294D8BCFE93FC5E" box="[227,302,939,961]" county="Amangeldy District" municipality="Ayat River" name="District" pageId="1" pageNumber="28">District</location>
,
<location id="737F601EFFC3FFC91336D8BCFE54FC5E" LSID="urn:lsid:plazi:treatment:FE0987D3FFC3FFC91294D9CBFCD8FB09:737F601EFFC3FFC91336D8BCFE54FC5E" box="[321,489,939,961]" county="Amangeldy District" municipality="Ayat River" name="Vyatsky Village" pageId="1" pageNumber="28">Vyatsky Village</location>
, near
<collectingMunicipality id="967BACBFFFC3FFC91042D8BBFD11FC5E" box="[565,684,939,961]" pageId="1" pageNumber="28">Ayat River</collectingMunicipality>
,
<date id="021E1005FFC3FFC910C9D8BBFC92FC5E" box="[702,815,940,961]" pageId="1" pageNumber="28" value="2014-05-04">
<collectingDate id="125AE9EDFFC3FFC910C9D8BBFC92FC5E" box="[702,815,940,961]" pageId="1" pageNumber="28" value="2014-05-04">04.V 2014</collectingDate>
</date>
,
<specimenCount id="60A6FD4CFFC3FFC91134D8BBFC3AFC5E" box="[835,903,939,961]" pageId="1" pageNumber="28" type="generic">1 juv.</specimenCount>
;
<collectingCounty id="9F7E4E49FFC3FFC911EED8BCFBDEFC5E" box="[921,1123,939,961]" pageId="1" pageNumber="28">Amangeldy District</collectingCounty>
</materialsCitation>
,
</paragraph>
<paragraph id="761F36C5FFC3FFC91294D8D0FDFCFC43" blockId="1.[227,1127,775,1044]" box="[227,577,967,989]" pageId="1" pageNumber="28">
<materialsCitation id="C6C83C98FFC3FFC91294D8D0FD80FC43" box="[227,573,967,989]" collectingDate="2017-05-16" location="Amangeldy Village" pageId="1" pageNumber="28" specimenCount="1" specimenCount-male="1">
<location id="737F601EFFC3FFC91294D8D0FE1FFC42" LSID="urn:lsid:plazi:treatment:FE0987D3FFC3FFC91294D9CBFCD8FB09:737F601EFFC3FFC91294D8D0FE1FFC42" box="[227,418,967,989]" name="Amangeldy Village" pageId="1" pageNumber="28">Amangeldy Village</location>
,
<date id="021E1005FFC3FFC913DBD8D0FDAEFC43" box="[428,531,967,989]" pageId="1" pageNumber="28" value="2017-05-16">
<collectingDate id="125AE9EDFFC3FFC913DBD8D0FDAEFC43" box="[428,531,967,989]" pageId="1" pageNumber="28" value="2017-05-16">16.V 2017</collectingDate>
</date>
,
<specimenCount id="60A6FD4CFFC3FFC9106AD8D0FD80FC43" box="[541,573,967,988]" pageId="1" pageNumber="28" type="male">1♂</specimenCount>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3EBA654EFFC3FFC9137ED8F5FCDFFBD6" pageId="1" pageNumber="28" type="distribution">
<paragraph id="761F36C5FFC3FFC9137ED8F5FC32FB8B" blockId="1.[227,1127,775,1044]" pageId="1" pageNumber="28">
DISTRIBUTION. Cosmopolitan species (Edgecombe, 2011). Recorded from the
<collectingRegion id="B464F827FFC3FFC91663D8F4FE3DFB8B" country="Kazakhstan" name="Shyghys Qazaqstan Oblysy" pageId="1" pageNumber="28">East Kazakhstan Region</collectingRegion>
(Dyachkov
<emphasis id="44D4EAD7FFC3FFC91380D8E9FD9CFB8C" box="[503,545,1022,1043]" italics="true" pageId="1" pageNumber="28">et al</emphasis>
., 2016); new to the Kostanay Region.
</paragraph>
<paragraph id="761F36C5FFC3FFC91390DF23FCDFFBD6" blockId="1.[487,866,1076,1098]" box="[487,866,1076,1098]" pageId="1" pageNumber="28">
<emphasis id="44D4EAD7FFC3FFC91390DF23FCDFFBD6" bold="true" box="[487,866,1076,1098]" pageId="1" pageNumber="28">
Order
<taxonomicName id="B1A04D46FFC3FFC9105ADF23FCDFFBD6" authority="Pocock, 1895" authorityName="Pocock" authorityYear="1895" box="[557,866,1076,1098]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lithobiomorpha" pageId="1" pageNumber="28" phylum="Tracheophyta" rank="order">Lithobiomorpha Pocock, 1895</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="3EBA654EFFC3FFC91382DF4DFCD8FB09" pageId="1" pageNumber="28" type="description">
<paragraph id="761F36C5FFC3FFC91382DF4DFCE8FBEF" blockId="1.[501,853,1114,1136]" box="[501,853,1114,1136]" pageId="1" pageNumber="28">
<emphasis id="44D4EAD7FFC3FFC91382DF4DFCE8FBEF" bold="true" box="[501,853,1114,1136]" pageId="1" pageNumber="28">
Family
<taxonomicName id="B1A04D46FFC3FFC91034DF4DFCE8FBEF" authority="Newport, 1844" authorityName="Newport" authorityYear="1844" box="[579,853,1114,1136]" class="Chilopoda" family="Lithobiidae" kingdom="Animalia" order="Lithobiomorpha" pageId="1" pageNumber="28" phylum="Arthropoda" rank="family">Lithobiidae Newport, 1844</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="761F36C5FFC3FFC91393DF97FCD8FB09" blockId="1.[484,869,1152,1175]" box="[484,869,1152,1175]" pageId="1" pageNumber="28">
<emphasis id="44D4EAD7FFC3FFC91393DF97FCD8FB09" bold="true" box="[484,869,1152,1175]" pageId="1" pageNumber="28">
Subfamily
<taxonomicName id="B1A04D46FFC3FFC91024DF96FCD8FB09" authority="Newport, 1844" authorityName="Newport" authorityYear="1844" box="[595,869,1153,1175]" class="Chilopoda" family="Lithobiidae" kingdom="Animalia" order="Lithobiomorpha" pageId="1" pageNumber="28" phylum="Arthropoda" rank="subFamily" subFamily="Lithobiinae">Lithobiinae Newport, 1844</taxonomicName>
</emphasis>
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<mods:title id="6F94C148EE67C2BBDA56575C5173D846">NEW DATA ON THE CENTIPEDE FAUNA (MYRIAPODA: CHILOPODA) OF KOSTANAY REGION, KAZAKHSTAN</mods:title>
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<taxonomicName id="B1A04D46FFC3FFC91294DFDBFD53FB7D" ID-CoL="3VJ95" authority="C. L. Koch, 1847" authorityName="C. L. Koch" authorityYear="1847" box="[227,750,1228,1251]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="curtipes" subGenus="Monotarsobius">
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<emphasis id="44D4EAD7FFC3FFC91294DFDBFEFFFB7D" bold="true" box="[227,322,1228,1250]" italics="true" pageId="1" pageNumber="28">Lithobius</emphasis>
(
<emphasis id="44D4EAD7FFC3FFC91327DFDBFE58FB7D" bold="true" box="[336,485,1228,1250]" italics="true" pageId="1" pageNumber="28">Monotarsobius</emphasis>
)
<emphasis id="44D4EAD7FFC3FFC91384DFDAFDFFFB7D" bold="true" box="[499,578,1229,1250]" italics="true" pageId="1" pageNumber="28">curtipes</emphasis>
C.L. Koch, 1847
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MATERIAL EXAMINED.
<emphasis id="44D4EAD7FFC3FFC91068DFEFFD6EFA91" bold="true" box="[543,723,1272,1294]" pageId="1" pageNumber="28">Kostanay Region</emphasis>
: Kostanay District, Stoun (Kamennoye)
</paragraph>
<paragraph id="761F36C5FFC3FFC91294DE04FCD5FAB6" blockId="1.[227,1126,1272,1403]" box="[227,872,1299,1321]" pageId="1" pageNumber="28">
Lake, birch forest, in litter,
<date id="021E1005FFC3FFC9139CDE04FDEFFAB7" box="[491,594,1299,1321]" pageId="1" pageNumber="28" value="2017-05-16">16.V 2017</date>
,
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,
<specimenCount id="60A6FD4CFFC3FFC910F1DE04FD18FAB6" box="[646,677,1299,1321]" pageId="1" pageNumber="28" type="female">5♀</specimenCount>
,
<specimenCount id="60A6FD4CFFC3FFC910C7DE04FD72FAB7" box="[688,719,1299,1320]" pageId="1" pageNumber="28" type="male">1♂</specimenCount>
/
<specimenCount id="60A6FD4CFFC3FFC910AEDE04FD55FAB7" box="[729,744,1299,1320]" pageId="1" pageNumber="28" type="female">♀</specimenCount>
, 3 agenitalis.
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DISTRIBUTION. Trans-Palaearctic polyzonal species: from Europe to Western Asia, Siberia and
<collectingCountry id="0EB77655FFC3FFC91348DE5DFE26FAFF" box="[319,411,1354,1376]" name="Mongolia" pageId="1" pageNumber="28">Mongolia</collectingCountry>
(Zalesskaja, 1978; Farzalieva &amp; Esyunin, 2008).
<collectingCountry id="0EB77655FFC3FFC9111DDE5DFC66FAFF" box="[874,987,1354,1376]" name="Kazakhstan" pageId="1" pageNumber="28">Kazakhstan</collectingCountry>
: Kostanay and
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<collectingRegion id="B464F827FFC3FFC91294DE71FE37FAE4" box="[227,394,1381,1403]" country="Kazakhstan" name="Batys Qazaqstan Oblysy" pageId="1" pageNumber="28">West Kazakhstan</collectingRegion>
regions (Bragina, 2012; Vsevolodova-Perel, 2009; Dyachkov, 2019).
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