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211 KiB
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695 lines
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<document id="4AA65F8949A10318646791795BBEEEB0" ID-CLB-Dataset="56571" ID-DOI="10.1016/j.jcz.2022.11.010" ID-GBIF-Dataset="5b10a3d9-1d42-4874-abfe-ef46cd21e183" ID-ISSN="1873-2674" ID-Zenodo-Dep="8164163" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1689773170588" checkinUser="felipe" docAuthor="Herranz, María, Moreleon, Alexis D., Rho, Hyun Soo & Sørensen, Martin V." docDate="2023" docId="039F6C5ADA3FFFEBFF31A443FBD4FA8E" docLanguage="en" docName="ZoolAnz.302.67-74.pdf" docOrigin="Zoologischer Anzeiger 302" docSource="http://dx.doi.org/10.1016/j.jcz.2022.11.010" docStyle="DocumentStyle:32483D6D135DE2FF927BA3F1400E0FFD.1:ZoolAnz.2022-.journal_article" docStyleId="32483D6D135DE2FF927BA3F1400E0FFD" docStyleName="ZoolAnz.2022-.journal_article" docStyleVersion="1" docTitle="Echinoderes inaequalis Herranz & Moreleon & Rho & Sørensen 2023, sp. nov." docType="treatment" docVersion="5" lastPageNumber="72" masterDocId="FFA61422DA3EFFEEFFB5A169FFB8FF90" masterDocTitle="Expanding the Echinoderes coulli group (Kinorhyncha: Cyclorhagida) with a new species from the Chuuk Islands, Micronesia" masterLastPageNumber="74" masterPageNumber="67" pageNumber="68" updateTime="1699544063373" updateUser="plazi">
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<mods:title id="57BD302857E788F2F11FEBEBE374DDEF">Expanding the Echinoderes coulli group (Kinorhyncha: Cyclorhagida) with a new species from the Chuuk Islands, Micronesia</mods:title>
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<mods:namePart id="5D342697BB973BCD2AEDEFEA82730738">Herranz, María</mods:namePart>
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<mods:namePart id="CA9C2A6B1CBFCB824D4885BDEA90D410">Moreleon, Alexis D.</mods:namePart>
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<mods:name id="F89A263BB02F1FC9F92DA604EBBAE8AA" type="personal">
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<mods:namePart id="39AD3B408B84E12CEAEB4EF0A164306D">Rho, Hyun Soo</mods:namePart>
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<mods:namePart id="1F5BC998D01A251C30171914D78AEAFA">Sørensen, Martin V.</mods:namePart>
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<mods:title id="17140DFEDCD5FC7D5FF24C7FA9D7635E">Zoologischer Anzeiger</mods:title>
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<mods:date id="701CC4CEBE01C8A1F981067232B84B54">2023</mods:date>
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<mods:number id="F709F519B13F6401AAEF4000E3832585">2023-01-31</mods:number>
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<mods:url id="D7FD1D99B24FCE836F97CC904E0541D7">http://dx.doi.org/10.1016/j.jcz.2022.11.010</mods:url>
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<mods:classification id="E6A47E4B5AADB6E66BCCA4533615B2EF">journal article</mods:classification>
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<treatment id="039F6C5ADA3FFFEBFF31A443FBD4FA8E" ID-DOI="http://doi.org/10.5281/zenodo.8171330" ID-GBIF-Taxon="212417501" ID-Zenodo-Dep="8171330" LSID="urn:lsid:plazi:treatment:039F6C5ADA3FFFEBFF31A443FBD4FA8E" httpUri="http://treatment.plazi.org/id/039F6C5ADA3FFFEBFF31A443FBD4FA8E" lastPageId="5" lastPageNumber="72" pageId="1" pageNumber="68">
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<subSubSection id="C32C8EC7DA3FFFEFFF31A443FE2AFAAD" box="[132,402,1321,1341]" pageId="1" pageNumber="68" type="nomenclature">
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<paragraph id="8B89DD4CDA3FFFEFFF31A443FE2AFAAD" blockId="1.[100,470,1266,1397]" box="[132,402,1321,1341]" pageId="1" pageNumber="68">
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Species
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<taxonomicName id="4C36A6CFDA3FFFEFFF7AA440FEF9FAAD" ID-CoL="7a56aa36-6a66-4b33-9767-fb9bc2e692f4" authority="Herranz & Moreleon & Rho & Sørensen, 2023" authorityName="Herranz & Moreleon & Rho & Sørensen" authorityYear="2023" box="[207,321,1321,1341]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="1" pageNumber="68" phylum="Kinorhyncha" rank="species" species="inaequalis" status="sp. nov.">
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<emphasis id="B942015EDA3FFFEFFF7AA440FEF9FAAD" bold="true" box="[207,321,1321,1341]" italics="true" pageId="1" pageNumber="68">E. inaequalis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A271BC25DA3FFFEFFEF2A443FE2AFAAD" box="[327,402,1322,1341]" pageId="1" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C8EC7DA3FFFEFFF31A42FFA3BFEA3" pageId="1" pageNumber="68" type="description">
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<paragraph id="8B89DD4CDA3FFFEFFF31A42FFE6EFAC9" blockId="1.[100,470,1266,1397]" box="[132,470,1350,1369]" pageId="1" pageNumber="68">urn:lsid:zoobank.org:act:41BC4BF8-</paragraph>
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<paragraph id="8B89DD4CDA3FFFEFFFD1A408FE00FAE4" blockId="1.[100,470,1266,1397]" box="[100,440,1377,1397]" pageId="1" pageNumber="68">E3B4–445C–87F1–9C6D2CC97AA2.</paragraph>
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<paragraph id="8B89DD4CDA3FFFEFFCE4A1FDFB95FF37" blockId="1.[849,1069,148,167]" box="[849,1069,148,167]" pageId="1" pageNumber="68">
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(
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<figureCitation id="130DC1C9DA3FFFEFFCECA1FDFC16FF37" box="[857,942,148,167]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="2.[100,130,1313,1330]" captionStartId-1="3.[100,130,1675,1692]" captionStartId-2="4.[100,130,1675,1692]" captionTargetBox-0="[188,1399,149,1284]" captionTargetBox-1="[147,1439,149,1646]" captionTargetBox-2="[147,1440,149,1646]" captionTargetId-0="figure-421@2.[187,1400,148,1285]" captionTargetId-1="figure-8@3.[146,1441,148,1647]" captionTargetId-2="figure-8@4.[146,1441,148,1647]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." captionText-1="Fig. 3. Light micrographs (DIC) showing traits in Echinoderes inaequalis sp. nov. Male paratype NHMD-1176545 (A, B, D, E, H), female holotype NHMD-1176540 (C, K, I) female paratype NHMD-1176543 (F, G), and male paratype NHMD-1176542 (J). (A) Male overview, ventral view. (B) Segments 1–2, ventral view. (C) Segments 1–2, dorsal view. (D) Segments 5–7, ventral view. (E) Segments 4–5, dorsal view. (F) Segments 7–8, lateroventral view. (G) Segments 6–7, dorsolateral view. (H) Male, segments 10–11 ventral view. (I) Female, segment 11, ventral view. (J) Segments 9–10, ventral view. (J’) Detail of sieve plate. (K) Segment 8, focused on laterodorsal area. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mdgco1, middorsal glandular cell outlet type 1; mlgco2, midlateral glandular cell outlet type 2; mlt, midlateral tube; mvp, midventral placid; pdgco1, paradorsal glandular cell outlet type 1; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; te, tergal extension; tp, trichoscalid plate; vlgco2, ventrolateral glandular cell outlet type 2; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." captionText-2="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral gla
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,
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<tableCitation id="C6B4E8F7DA3FFFEFFC0FA1FDFB98FF37" box="[954,1056,148,167]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="5.[100,150,150,166]" captionStartId-1="5.[100,150,1552,1568]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionText-0="Table 1 Measurements (in μm) from light microscopy (except for the middorsal spine of segment 4) of adult Echinoderes inaequalis sp. nov. Abbreviations: LTAS, lateral terminal accessory spine; LTS, lateral terminal spine; LVT, lateroventral tube; MD, middorsal spine; MLT, midlateral tube; MSW, maximum sternal width; n, number of specimens; SD, standard deviation; SDT, subdorsal tube; SI, sieve plate length; SW, standard width; S, segment lengths; TL, trunk length.Numbers, where inserted, indicate segment number." captionText-1="Table 2 Summary of the nature and location of sensory spots, glandular cell outlets, spines and tubes in E. inaequalis sp. nov. Abbreviations: LA: lateral accessory; LD: laterodorsal; LV: lateroventral; MD: middorsal; ML: midlateral; PD: paradorsal; SD: subdorsal; SL: sublateral; VL: ventrolateral; VM: ventromedial; ac, acicular spine; gco1/2, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; pe, penile spines; si, sieve plate; ss, sensory spot; tu, tube; tu*, unpaired tube; (Ƌ), male condition of sexually dimorphic character; (♀), female condition of sexually dimorphic character." httpUri-0="http://table.plazi.org/id/DF498DC4DA3BFFEBFFD1A1FFFE73FECA" httpUri-1="http://table.plazi.org/id/DF498DC4DA3BFFEBFFD1A779FBCEF916" pageId="1" pageNumber="68" tableUuid-0="DF498DC4DA3BFFEBFFD1A1FFFE73FECA" tableUuid-1="DF498DC4DA3BFFEBFFD1A779FBCEF916">Tables 1–2</tableCitation>
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).
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</paragraph>
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<paragraph id="8B89DD4CDA3FFFEFFC87A1A3FC68FF4E" blockId="1.[818,1487,202,307]" box="[818,976,202,222]" pageId="1" pageNumber="68">
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<emphasis id="B942015EDA3FFFEFFC87A1A3FC68FF4E" bold="true" box="[818,976,202,222]" italics="true" pageId="1" pageNumber="68">3.1.1. Etymology</emphasis>
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</paragraph>
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<paragraph id="8B89DD4CDA3FFFEFFCE4A18EFA3BFEA3" blockId="1.[818,1487,202,307]" pageId="1" pageNumber="68">The species name comes from the Latin in- (un or not) and -aequ¯alis (equal or even), meaning, the uneven one, referring to the distinctive asymmetric distribution of subdorsal tubes on segments 5 to 7.</paragraph>
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</subSubSection>
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<subSubSection id="C32C8EC7DA3FFFEFFC87A03FFB7BFD61" pageId="1" pageNumber="68" type="diagnosis">
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<paragraph id="8B89DD4CDA3FFFEFFC87A03FFC70FEFA" blockId="1.[818,1488,342,753]" box="[818,968,342,362]" pageId="1" pageNumber="68">
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<emphasis id="B942015EDA3FFFEFFC87A03FFC70FEFA" bold="true" box="[818,968,342,362]" italics="true" pageId="1" pageNumber="68">3.1.2. Diagnosis</emphasis>
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</paragraph>
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<paragraph id="8B89DD4CDA3FFFEFFCE4A01BFB7BFD61" blockId="1.[818,1488,342,753]" pageId="1" pageNumber="68">
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<taxonomicName id="4C36A6CFDA3FFFEFFCE4A01BFC02FE16" baseAuthorityName="Yamasaki" baseAuthorityYear="2022" box="[849,954,370,390]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="1" pageNumber="68" phylum="Kinorhyncha" rank="genus">
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<emphasis id="B942015EDA3FFFEFFCE4A01BFC02FE16" bold="true" box="[849,954,370,390]" italics="true" pageId="1" pageNumber="68">Echinoderes</emphasis>
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</taxonomicName>
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with a single, minute middorsal spine on segment 4. Lateroventral tubes on segments 5, 7–9, midlateral tubes on segment 8, and sexually dimorphic laterodorsal tubes on segment 10. Single subdorsal tubes usually in alternating positions on segments 5–7 with intraspecific variation. Small glandular cell outlets
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<typeStatus id="548D63EEDA3FFFEFFAA0A08AFA86FE66" box="[1301,1342,483,502]" pageId="1" pageNumber="68">type</typeStatus>
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<quantity id="4CCE70A9DA3FFFEFFAF1A08AFAD0FE65" box="[1348,1384,482,502]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="1" pageNumber="68" unit="in" value="2.0">2 in</quantity>
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subdorsal, laterodorsal and ventrolateral positions on segment 2, subdorsal position on segment 4, and midlateral positions on segments 5–7; large, conspicuous, laterodorsal glandular cell outlets
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<typeStatus id="548D63EEDA3FFFEFFB4AA35EFA90FDDA" box="[1279,1320,567,586]" pageId="1" pageNumber="68">type</typeStatus>
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2 are present on segment 8. Large nephridial areas composed of elongated sieve plates and a posterior pore in sublateral position on segment 9. Short and stout lateral terminal spines with fringed edges. Males with three pairs of slender penile spines, and well-developed laterodorsal tubes on segment 10. Females with short and thin lateral terminal accessory spines, and reduced laterodorsal tubes on segment 10.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C8EC7DA3FFFEFFC87A27DFB6CFB77" pageId="1" pageNumber="68" type="materials_examined">
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<paragraph id="8B89DD4CDA3FFFEFFC87A27DFBA4FCB8" blockId="1.[818,1488,788,1255]" box="[818,1052,788,808]" pageId="1" pageNumber="68">
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<emphasis id="B942015EDA3FFFEFFC87A27DFBA4FCB8" bold="true" box="[818,1052,788,808]" italics="true" pageId="1" pageNumber="68">3.1.3. Material examined</emphasis>
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</paragraph>
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<paragraph id="8B89DD4CDA3FFFEFFCE4A258FB79FC24" blockId="1.[818,1488,788,1255]" pageId="1" pageNumber="68">
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<materialsCitation id="3B5ED711DA3FFFEFFCE4A258FB05FC24" ID-GBIF-Occurrence="4163027301" collectingDate="2007-03-21" country="Micronesia" elevation="2" latitude="7.4530554" location="Islands" longLatPrecision="21" longitude="151.90416" pageId="1" pageNumber="68" specimenCount="1" stateProvince="Chuuk" typeStatus="holotype">
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Holotypic female collected by HSR on
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<date id="FF88FB8CDA3FFFEFFB0AA258FADEFCD4" box="[1215,1382,817,836]" pageId="1" pageNumber="68" value="2007-03-21">
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<collectingDate id="EFCC0264DA3FFFEFFB0AA258FADEFCD4" box="[1215,1382,817,836]" pageId="1" pageNumber="68" value="2007-03-21">March 21st, 2007</collectingDate>
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</date>
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from coral sand at
|
|||
|
|
<elevation id="001B3A7FDA3FFFEFFC34A224FC45FCF0" box="[897,1021,845,864]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="1" pageNumber="68" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">
|
|||
|
|
<quantity id="4CCE70A9DA3FFFEFFC34A224FC06FCF0" box="[897,958,845,864]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="1" pageNumber="68" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1–2 m</quantity>
|
|||
|
|
depth
|
|||
|
|
</elevation>
|
|||
|
|
at
|
|||
|
|
<collectingRegion id="49F213AEDA3FFFEFFB94A224FBE6FCF0" box="[1057,1118,845,864]" country="Micronesia" name="Chuuk" pageId="1" pageNumber="68">Chuuk</collectingRegion>
|
|||
|
|
<location id="8EE98B97DA3FFFEFFBD2A224FB13FCF0" LSID="urn:lsid:plazi:treatment:039F6C5ADA3FFFEBFF31A443FBD4FA8E:8EE98B97DA3FFFEFFBD2A224FB13FCF0" box="[1127,1195,845,864]" country="Micronesia" latitude="7.4530554" longLatPrecision="21" longitude="151.90416" name="Islands" pageId="1" pageNumber="68" stateProvince="Chuuk">Islands</location>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="F3219DDCDA3FFFEFFB02A224FAA6FCF0" box="[1207,1310,845,864]" name="Micronesia" pageId="1" pageNumber="68">Micronesia</collectingCountry>
|
|||
|
|
(locality MAP-19:
|
|||
|
|
<geoCoordinate id="EE02BB8BDA3FFFEFFC87A200FC0DFCEC" box="[818,949,870,892]" degrees="007" direction="north" minutes="27" orientation="latitude" pageId="1" pageNumber="68" precision="15" seconds="11" value="7.4530554">
|
|||
|
|
007
|
|||
|
|
<superScript id="7C437004DA3FFFEFFCE3A20FFCE5FCE3" attach="none" box="[854,861,870,883]" fontSize="6" pageId="1" pageNumber="68">◦</superScript>
|
|||
|
|
27’11’’N
|
|||
|
|
</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE02BB8BDA3FFFEFFC0AA200FB87FCEC" box="[959,1087,870,892]" degrees="151" direction="east" minutes="54" orientation="longitude" pageId="1" pageNumber="68" precision="15" seconds="15" value="151.90416">
|
|||
|
|
151
|
|||
|
|
<superScript id="7C437004DA3FFFEFFC56A20FFC52FCE3" attach="none" box="[995,1002,870,883]" fontSize="6" pageId="1" pageNumber="68">◦</superScript>
|
|||
|
|
54’15’’E
|
|||
|
|
</geoCoordinate>
|
|||
|
|
) (
|
|||
|
|
<figureCitation id="130DC1C9DA3FFFEFFBE1A200FB31FCEC" box="[1108,1161,873,892]" captionStart="Fig" captionStartId="1.[100,130,1939,1956]" captionTargetBox="[108,766,1439,1909]" captionTargetId="figure-1009@1.[106,767,1437,1911]" captionTargetPageId="1" captionText="Fig. 1. Map showing (A) the position of the Chuuk Islands, and (B) zoom of the sampling localities." figureDoi="http://doi.org/10.5281/zenodo.8164165" httpUri="https://zenodo.org/record/8164165/files/figure.png" pageId="1" pageNumber="68">Fig. 1</figureCitation>
|
|||
|
|
); mounted in
|
|||
|
|
<collectorName id="26C3B89ADA3FFFEFFAA5A200FA18FCEC" box="[1296,1440,873,892]" pageId="1" pageNumber="68">Fluoromount G</collectorName>
|
|||
|
|
on a glass slide and deposited at the
|
|||
|
|
<collectorName id="26C3B89ADA3FFFEFFBD3A2ECFAEAFC08" box="[1126,1362,901,920]" pageId="1" pageNumber="68">Natural History Museum</collectorName>
|
|||
|
|
of Denmark, under catalogue number NHMD-1176540
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B89DD4CDA3FFFEFFCE4A2D4FBC8FB3F" blockId="1.[818,1488,788,1255]" pageId="1" pageNumber="68">
|
|||
|
|
<materialsCitation id="3B5ED711DA3FFFEFFCE4A2D4FBD4FB3F" ID-GBIF-Occurrence="4163027303" collectingDate="2007-03-09" collectorName="Fluoromount G" country="Denmark" elevation="30" latitude="7.4575" longLatPrecision="21" longitude="151.89778" pageId="1" pageNumber="68" specimenCount="3" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
|
|||
|
|
<typeStatus id="548D63EEDA3FFFEFFCE4A2D4FC14FC40" box="[849,940,957,976]" pageId="1" pageNumber="68" type="paratype">Paratypes</typeStatus>
|
|||
|
|
series include
|
|||
|
|
<specimenCount id="9D3016C5DA3FFFEFFB80A2D4FB78FC40" box="[1077,1216,957,976]" pageId="1" pageNumber="68" type="generic">five specimens</specimenCount>
|
|||
|
|
,
|
|||
|
|
<specimenCount id="9D3016C5DA3FFFEFFB7DA2D4FAFCFC40" box="[1224,1348,957,976]" pageId="1" pageNumber="68" type="female">three females</specimenCount>
|
|||
|
|
and
|
|||
|
|
<specimenCount id="9D3016C5DA3FFFEFFAC4A2D4FA77FC40" box="[1393,1487,957,976]" pageId="1" pageNumber="68" type="male">two males</specimenCount>
|
|||
|
|
all of them mounted in
|
|||
|
|
<collectorName id="26C3B89ADA3FFFEFFBA4A2B0FB19FC7C" box="[1041,1185,985,1004]" pageId="1" pageNumber="68">Fluoromount G</collectorName>
|
|||
|
|
on glass slides and deposited at the Natural History Museum of
|
|||
|
|
<collectingCountry id="F3219DDCDA3FFFEFFBC7A29DFB72FB97" box="[1138,1226,1012,1031]" name="Denmark" pageId="1" pageNumber="68">Denmark</collectingCountry>
|
|||
|
|
, under catalogue numbers NHMD-1176541 to 45. Of those, male paratypes NHMD-1176542, NHMD-1176545 and female paratype NHMD-1176541 were collected at the same locality as the holotype; whereas the two remaining female paratypes, NHMD-1176543 and NHMD-1176544, were collected on
|
|||
|
|
<date id="FF88FB8CDA3FFFEFFC87A5E9FC69FB03" box="[818,977,1152,1171]" pageId="1" pageNumber="68" value="2007-03-09">
|
|||
|
|
<collectingDate id="EFCC0264DA3FFFEFFC87A5E9FC69FB03" box="[818,977,1152,1171]" pageId="1" pageNumber="68" value="2007-03-09">March 9th, 2007</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
from subtidal sediment at
|
|||
|
|
<elevation id="001B3A7FDA3FFFEFFB67A5E9FAF8FB03" box="[1234,1344,1152,1171]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="68" unit="m" value="30.0">
|
|||
|
|
<quantity id="4CCE70A9DA3FFFEFFB67A5E9FABBFB03" box="[1234,1283,1152,1171]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="68" unit="m" value="30.0">30 m</quantity>
|
|||
|
|
depth
|
|||
|
|
</elevation>
|
|||
|
|
(locality MAP-01:
|
|||
|
|
<geoCoordinate id="EE02BB8BDA3FFFEFFCE3A5F5FC62FB3F" box="[854,986,1177,1199]" degrees="007" direction="north" minutes="27" orientation="latitude" pageId="1" pageNumber="68" precision="15" seconds="27" value="7.4575">
|
|||
|
|
007
|
|||
|
|
<superScript id="7C437004DA3FFFEFFCCFA5F0FC39FB36" attach="none" box="[890,897,1177,1190]" fontSize="6" pageId="1" pageNumber="68">◦</superScript>
|
|||
|
|
27’27’’N
|
|||
|
|
</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE02BB8BDA3FFFEFFC56A5F5FBDDFB3F" box="[995,1125,1177,1199]" degrees="151" direction="east" minutes="53" orientation="longitude" pageId="1" pageNumber="68" precision="15" seconds="52" value="151.89778">
|
|||
|
|
151
|
|||
|
|
<superScript id="7C437004DA3FFFEFFBB2A5F0FBB6FB36" attach="none" box="[1031,1038,1177,1190]" fontSize="6" pageId="1" pageNumber="68">◦</superScript>
|
|||
|
|
53’52’’E
|
|||
|
|
</geoCoordinate>
|
|||
|
|
)
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B89DD4CDA3FFFEFFCE4A5D1FB6CFB77" blockId="1.[818,1488,788,1255]" pageId="1" pageNumber="68">
|
|||
|
|
<materialsCitation id="3B5ED711DA3FFFEFFCE4A5D1FB68FB77" ID-GBIF-Occurrence="4163027302" collectionCode="SEM" pageId="1" pageNumber="68" specimenCount="17">
|
|||
|
|
Additional material includes
|
|||
|
|
<specimenCount id="9D3016C5DA3FFFEFFBD8A5D1FB56FB5B" box="[1133,1262,1208,1227]" pageId="1" pageNumber="68" type="generic">17 specimens</specimenCount>
|
|||
|
|
mounted for
|
|||
|
|
<collectionCode id="ED274589DA3FFFEFFACCA5D1FA1BFB5B" box="[1401,1443,1208,1227]" pageId="1" pageNumber="68">SEM</collectionCode>
|
|||
|
|
and stored in MVS personal reference collection
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C32C8EC7DA3FFFEBFC87A463FBD4FA8E" lastPageId="5" lastPageNumber="72" pageId="1" pageNumber="68" type="description">
|
|||
|
|
<paragraph id="8B89DD4CDA3FFFEFFC87A463FC6FFA8E" blockId="1.[818,1488,1290,1980]" box="[818,983,1290,1310]" pageId="1" pageNumber="68">
|
|||
|
|
<emphasis id="B942015EDA3FFFEFFC87A463FC6FFA8E" bold="true" box="[818,983,1290,1310]" italics="true" pageId="1" pageNumber="68">3.1.4. Description</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B89DD4CDA3FFFEFFCE4A44EFA68FA3A" blockId="1.[818,1488,1290,1980]" pageId="1" pageNumber="68">
|
|||
|
|
Adults with head, neck and eleven trunk segments (
|
|||
|
|
<figureCitation id="130DC1C9DA3FFFEFFAF5A44EFA38FAAA" box="[1344,1408,1319,1338]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="1" pageNumber="68">Figs. 2</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="130DC1C9DA3FFFEFFA3EA44EFA1DFAAA" box="[1419,1445,1319,1338]" captionStart="Fig" captionStartId="3.[100,130,1675,1692]" captionTargetBox="[147,1439,149,1646]" captionTargetId="figure-8@3.[146,1441,148,1647]" captionTargetPageId="3" captionText="Fig. 3. Light micrographs (DIC) showing traits in Echinoderes inaequalis sp. nov. Male paratype NHMD-1176545 (A, B, D, E, H), female holotype NHMD-1176540 (C, K, I) female paratype NHMD-1176543 (F, G), and male paratype NHMD-1176542 (J). (A) Male overview, ventral view. (B) Segments 1–2, ventral view. (C) Segments 1–2, dorsal view. (D) Segments 5–7, ventral view. (E) Segments 4–5, dorsal view. (F) Segments 7–8, lateroventral view. (G) Segments 6–7, dorsolateral view. (H) Male, segments 10–11 ventral view. (I) Female, segment 11, ventral view. (J) Segments 9–10, ventral view. (J’) Detail of sieve plate. (K) Segment 8, focused on laterodorsal area. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mdgco1, middorsal glandular cell outlet type 1; mlgco2, midlateral glandular cell outlet type 2; mlt, midlateral tube; mvp, midventral placid; pdgco1, paradorsal glandular cell outlet type 1; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; te, tergal extension; tp, trichoscalid plate; vlgco2, ventrolateral glandular cell outlet type 2; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164174" httpUri="https://zenodo.org/record/8164174/files/figure.png" pageId="1" pageNumber="68">3A</figureCitation>
|
|||
|
|
and
|
|||
|
|
<figureCitation id="130DC1C9DA3FFFEFFC87A42AFCE8FAC6" box="[818,848,1347,1366]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="1" pageNumber="68">4A</figureCitation>
|
|||
|
|
). Measurements and dimensions are given in
|
|||
|
|
<tableCitation id="C6B4E8F7DA3FFFEFFB4FA42AFAE2FAC6" box="[1274,1370,1347,1367]" captionStart="Table 1" captionStartId="5.[100,150,150,166]" captionTargetPageId="5" captionText="Table 1 Measurements (in μm) from light microscopy (except for the middorsal spine of segment 4) of adult Echinoderes inaequalis sp. nov. Abbreviations: LTAS, lateral terminal accessory spine; LTS, lateral terminal spine; LVT, lateroventral tube; MD, middorsal spine; MLT, midlateral tube; MSW, maximum sternal width; n, number of specimens; SD, standard deviation; SDT, subdorsal tube; SI, sieve plate length; SW, standard width; S, segment lengths; TL, trunk length.Numbers, where inserted, indicate segment number." httpUri="http://table.plazi.org/id/DF498DC4DA3BFFEBFFD1A1FFFE73FECA" pageId="1" pageNumber="68" tableUuid="DF498DC4DA3BFFEBFFD1A1FFFE73FECA">Table 1. A</tableCitation>
|
|||
|
|
summary of sensory spots, spines, tubes and glandular cell outlet positions is provided in
|
|||
|
|
<tableCitation id="C6B4E8F7DA3FFFEFFC34A412FC7EFA1E" box="[897,966,1403,1422]" captionStart="Table 2" captionStartId="5.[100,150,1552,1568]" captionTargetPageId="5" captionText="Table 2 Summary of the nature and location of sensory spots, glandular cell outlets, spines and tubes in E. inaequalis sp. nov. Abbreviations: LA: lateral accessory; LD: laterodorsal; LV: lateroventral; MD: middorsal; ML: midlateral; PD: paradorsal; SD: subdorsal; SL: sublateral; VL: ventrolateral; VM: ventromedial; ac, acicular spine; gco1/2, glandular cell outlet type 1/2; ltas, lateral terminal accessory spine; lts, lateral terminal spine; pe, penile spines; si, sieve plate; ss, sensory spot; tu, tube; tu*, unpaired tube; (Ƌ), male condition of sexually dimorphic character; (♀), female condition of sexually dimorphic character." httpUri="http://table.plazi.org/id/DF498DC4DA3BFFEBFFD1A779FBCEF916" pageId="1" pageNumber="68" tableUuid="DF498DC4DA3BFFEBFFD1A779FBCEF916">Table 2</tableCitation>
|
|||
|
|
. Mouth cone and introvert could not be studied in detail due to the poor condition of the SEM specimens prepared 14 years ago.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B89DD4CDA3FFFEFFCE4A4DAFAF4F919" blockId="1.[818,1488,1290,1980]" pageId="1" pageNumber="68">
|
|||
|
|
The neck consists of 16 placids numbered clockwise from the midventral 1 (
|
|||
|
|
<figureCitation id="130DC1C9DA3FFFEFFC26A4A6FBACFA72" box="[915,1044,1487,1506]" captionStart="Fig" captionStartId="3.[100,130,1675,1692]" captionTargetBox="[147,1439,149,1646]" captionTargetId="figure-8@3.[146,1441,148,1647]" captionTargetPageId="3" captionText="Fig. 3. Light micrographs (DIC) showing traits in Echinoderes inaequalis sp. nov. Male paratype NHMD-1176545 (A, B, D, E, H), female holotype NHMD-1176540 (C, K, I) female paratype NHMD-1176543 (F, G), and male paratype NHMD-1176542 (J). (A) Male overview, ventral view. (B) Segments 1–2, ventral view. (C) Segments 1–2, dorsal view. (D) Segments 5–7, ventral view. (E) Segments 4–5, dorsal view. (F) Segments 7–8, lateroventral view. (G) Segments 6–7, dorsolateral view. (H) Male, segments 10–11 ventral view. (I) Female, segment 11, ventral view. (J) Segments 9–10, ventral view. (J’) Detail of sieve plate. (K) Segment 8, focused on laterodorsal area. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mdgco1, middorsal glandular cell outlet type 1; mlgco2, midlateral glandular cell outlet type 2; mlt, midlateral tube; mvp, midventral placid; pdgco1, paradorsal glandular cell outlet type 1; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; te, tergal extension; tp, trichoscalid plate; vlgco2, ventrolateral glandular cell outlet type 2; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164174" httpUri="https://zenodo.org/record/8164174/files/figure.png" pageId="1" pageNumber="68">Fig. 3B and C</figureCitation>
|
|||
|
|
). Placids 2–16 are trapezoid measuring 9 μm at the base while the midventral placid is more rectangular and wider measuring 15 μm (Fig.. 2B, 3B–C). All placids articulate with the first trunk segment. Trichoscalid plates bearing trichoscalids appear dorsally on placids 6, 8, 10, 12 and ventrally on placids 2 and 16. Ventral trichoscalid plates are triangular with rounded edges, while dorsal trichoscalid plates are rounded and smaller (
|
|||
|
|
<figureCitation id="130DC1C9DA3FFFEFFB77A71FFAB8F919" box="[1218,1280,1654,1673]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="1" pageNumber="68">Figs. 2</figureCitation>
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,
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<figureCitation id="130DC1C9DA3FFFEFFAB9A71FFAF9F919" box="[1292,1345,1654,1673]" captionStart="Fig" captionStartId="3.[100,130,1675,1692]" captionTargetBox="[147,1439,149,1646]" captionTargetId="figure-8@3.[146,1441,148,1647]" captionTargetPageId="3" captionText="Fig. 3. Light micrographs (DIC) showing traits in Echinoderes inaequalis sp. nov. Male paratype NHMD-1176545 (A, B, D, E, H), female holotype NHMD-1176540 (C, K, I) female paratype NHMD-1176543 (F, G), and male paratype NHMD-1176542 (J). (A) Male overview, ventral view. (B) Segments 1–2, ventral view. (C) Segments 1–2, dorsal view. (D) Segments 5–7, ventral view. (E) Segments 4–5, dorsal view. (F) Segments 7–8, lateroventral view. (G) Segments 6–7, dorsolateral view. (H) Male, segments 10–11 ventral view. (I) Female, segment 11, ventral view. (J) Segments 9–10, ventral view. (J’) Detail of sieve plate. (K) Segment 8, focused on laterodorsal area. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mdgco1, middorsal glandular cell outlet type 1; mlgco2, midlateral glandular cell outlet type 2; mlt, midlateral tube; mvp, midventral placid; pdgco1, paradorsal glandular cell outlet type 1; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; te, tergal extension; tp, trichoscalid plate; vlgco2, ventrolateral glandular cell outlet type 2; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164174" httpUri="https://zenodo.org/record/8164174/files/figure.png" pageId="1" pageNumber="68">3B–C</figureCitation>
|
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).
|
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</paragraph>
|
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|
|
<paragraph id="8B89DD4CDA3FFFECFCE4A7FBFDA4F9B8" blockId="1.[818,1488,1290,1980]" lastBlockId="2.[100,771,1473,1967]" lastPageId="2" lastPageNumber="69" pageId="1" pageNumber="68">
|
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|
|
Trunk segments 1 and 2 consist of one closed cuticular ring (
|
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|
|
<figureCitation id="130DC1C9DA3FFFEFFA39A7FBFA71F935" box="[1420,1481,1682,1701]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="1" pageNumber="68">Figs. 2</figureCitation>
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,
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<figureCitation id="130DC1C9DA3FFFEFFC87A7C7FC83F951" box="[818,827,1710,1729]" captionStart="Fig" captionStartId="3.[100,130,1675,1692]" captionTargetBox="[147,1439,149,1646]" captionTargetId="figure-8@3.[146,1441,148,1647]" captionTargetPageId="3" captionText="Fig. 3. Light micrographs (DIC) showing traits in Echinoderes inaequalis sp. nov. Male paratype NHMD-1176545 (A, B, D, E, H), female holotype NHMD-1176540 (C, K, I) female paratype NHMD-1176543 (F, G), and male paratype NHMD-1176542 (J). (A) Male overview, ventral view. (B) Segments 1–2, ventral view. (C) Segments 1–2, dorsal view. (D) Segments 5–7, ventral view. (E) Segments 4–5, dorsal view. (F) Segments 7–8, lateroventral view. (G) Segments 6–7, dorsolateral view. (H) Male, segments 10–11 ventral view. (I) Female, segment 11, ventral view. (J) Segments 9–10, ventral view. (J’) Detail of sieve plate. (K) Segment 8, focused on laterodorsal area. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mdgco1, middorsal glandular cell outlet type 1; mlgco2, midlateral glandular cell outlet type 2; mlt, midlateral tube; mvp, midventral placid; pdgco1, paradorsal glandular cell outlet type 1; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; te, tergal extension; tp, trichoscalid plate; vlgco2, ventrolateral glandular cell outlet type 2; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164174" httpUri="https://zenodo.org/record/8164174/files/figure.png" pageId="1" pageNumber="68">3</figureCitation>
|
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A-C, 4A-B) while segments 3–11 are composed of one tergal and two sternal plates (
|
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|
|
<figureCitation id="130DC1C9DA3FFFEFFC08A7A3FC44F94D" box="[957,1020,1738,1757]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="1" pageNumber="68">Figs. 2</figureCitation>
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,
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<figureCitation id="130DC1C9DA3FFFEFFBB3A7A3FB99F94D" box="[1030,1057,1738,1757]" captionStart="Fig" captionStartId="3.[100,130,1675,1692]" captionTargetBox="[147,1439,149,1646]" captionTargetId="figure-8@3.[146,1441,148,1647]" captionTargetPageId="3" captionText="Fig. 3. Light micrographs (DIC) showing traits in Echinoderes inaequalis sp. nov. Male paratype NHMD-1176545 (A, B, D, E, H), female holotype NHMD-1176540 (C, K, I) female paratype NHMD-1176543 (F, G), and male paratype NHMD-1176542 (J). (A) Male overview, ventral view. (B) Segments 1–2, ventral view. (C) Segments 1–2, dorsal view. (D) Segments 5–7, ventral view. (E) Segments 4–5, dorsal view. (F) Segments 7–8, lateroventral view. (G) Segments 6–7, dorsolateral view. (H) Male, segments 10–11 ventral view. (I) Female, segment 11, ventral view. (J) Segments 9–10, ventral view. (J’) Detail of sieve plate. (K) Segment 8, focused on laterodorsal area. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mdgco1, middorsal glandular cell outlet type 1; mlgco2, midlateral glandular cell outlet type 2; mlt, midlateral tube; mvp, midventral placid; pdgco1, paradorsal glandular cell outlet type 1; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; te, tergal extension; tp, trichoscalid plate; vlgco2, ventrolateral glandular cell outlet type 2; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164174" httpUri="https://zenodo.org/record/8164174/files/figure.png" pageId="1" pageNumber="68">3A</figureCitation>
|
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and
|
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<figureCitation id="130DC1C9DA3FFFEFFBE5A7A3FBD6F94D" box="[1104,1134,1738,1757]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="1" pageNumber="68">4A</figureCitation>
|
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). Glandular cell outlets
|
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<typeStatus id="548D63EEDA3FFFEFFAFBA7A3FACFF94D" box="[1358,1399,1738,1757]" pageId="1" pageNumber="68">type</typeStatus>
|
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|
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1 consist of numerous minute pores arranged in a circle, situated in the anterior part of the segments, usually hidden under the posterior part of the previous segment and quite inconspicuous. Dorsal glandular cell outlets
|
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<typeStatus id="548D63EEDA3FFFEFFC87A653FCE3F8DD" box="[818,859,1850,1869]" pageId="1" pageNumber="68">type</typeStatus>
|
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1 are located middorsally on segments 1–3, 10–11 and paradorsally on segments 4–9 (
|
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<figureCitation id="130DC1C9DA3FFFEFFC55A63CFBAEF8F8" box="[992,1046,1877,1896]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="1" pageNumber="68">Fig. 2</figureCitation>
|
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). Ventral glandular cell outlets
|
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<typeStatus id="548D63EEDA3FFFEFFAFDA63FFAC9F8F9" box="[1352,1393,1878,1897]" pageId="1" pageNumber="68">type</typeStatus>
|
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1 are lateroventral on segment 1 and ventromedial on segments 2–10 (
|
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|
|
<figureCitation id="130DC1C9DA3FFFEFFA3EA618FA79F814" box="[1419,1473,1905,1924]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="1" pageNumber="68">Fig. 2</figureCitation>
|
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). Primary pectinate fringe well developed in all segments, composed of very short fringe tips on segments 1 and 2 (
|
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|
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<figureCitation id="130DC1C9DA3FFFEFFB70A6C0FABAF82C" box="[1221,1282,1961,1980]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="1" pageNumber="68">Figs. 2</figureCitation>
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and
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<figureCitation id="130DC1C9DA3FFFEFFA9AA6C0FAF1F82C" box="[1327,1353,1961,1980]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="1" pageNumber="68">4B</figureCitation>
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) and long and flexible tips in remaining segments (
|
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|
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<figureCitation id="130DC1C9DA3CFFECFE06A4A8FE57FA44" box="[435,495,1473,1492]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="2" pageNumber="69">Figs. 2</figureCitation>
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and
|
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<figureCitation id="130DC1C9DA3CFFECFDAFA4A8FD8EFA44" box="[538,566,1473,1492]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="2" pageNumber="69">4B</figureCitation>
|
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). Secondary pectinate fringe absent on segment 1, and on segments 2–11 consisting of a single line of minute and regular teeth usually hidden under the primary pectinate fringe of previous segment (
|
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<figureCitation id="130DC1C9DA3CFFECFE7EA77CFDA8F9B8" box="[459,528,1557,1576]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="2" pageNumber="69">Fig. 4B</figureCitation>
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).
|
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</paragraph>
|
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<caption id="DF498DC4DA3FFFEFFFD1A6FAFEB2F82D" ID-DOI="http://doi.org/10.5281/zenodo.8164165" ID-Zenodo-Dep="8164165" httpUri="https://zenodo.org/record/8164165/files/figure.png" pageId="1" pageNumber="68" startId="1.[100,130,1939,1956]" targetBox="[108,766,1439,1909]" targetPageId="1" targetType="figure">
|
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<paragraph id="8B89DD4CDA3FFFEFFFD1A6FAFEB2F82D" blockId="1.[100,770,1938,1981]" pageId="1" pageNumber="68">
|
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|
|
<emphasis id="B942015EDA3FFFEFFFD1A6FAFF24F834" bold="true" box="[100,156,1939,1956]" pageId="1" pageNumber="68">Fig. 1.</emphasis>
|
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|
Map showing (A) the position of the Chuuk Islands, and (B) zoom of the sampling localities.
|
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|
</paragraph>
|
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|
|
</caption>
|
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|
|
<caption id="DF498DC4DA3CFFECFFD1A448FA68FA07" ID-DOI="http://doi.org/10.5281/zenodo.8164169" ID-Zenodo-Dep="8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="2" pageNumber="69" startId="2.[100,130,1313,1330]" targetBox="[188,1399,149,1284]" targetPageId="2" targetType="figure">
|
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|
|
<paragraph id="8B89DD4CDA3CFFECFFD1A448FA68FA07" blockId="2.[100,1488,1312,1433]" pageId="2" pageNumber="69">
|
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|
|
<emphasis id="B942015EDA3CFFECFFD1A448FF25FAA2" bold="true" box="[100,157,1313,1330]" pageId="2" pageNumber="69">Fig. 2.</emphasis>
|
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Line art illustrations of
|
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|
<taxonomicName id="4C36A6CFDA3CFFECFEDEA449FD98FAA2" authority="Herranz & Moreleon & Rho & Sørensen, 2023" authorityName="Herranz & Moreleon & Rho & Sørensen" authorityYear="2023" box="[363,544,1312,1330]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="2" pageNumber="69" phylum="Kinorhyncha" rank="species" species="inaequalis" status="sp. nov.">
|
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|
|
<emphasis id="B942015EDA3CFFECFEDEA449FD98FAA2" bold="true" box="[363,544,1312,1330]" italics="true" pageId="2" pageNumber="69">Echinoderes inaequalis</emphasis>
|
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</taxonomicName>
|
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<taxonomicNameLabel id="A271BC25DA3CFFECFD90A448FDD0FAA1" box="[549,616,1313,1329]" pageId="2" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
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(A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm.
|
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|
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</paragraph>
|
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</caption>
|
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<paragraph id="8B89DD4CDA3CFFECFF31A758FD13F8CB" blockId="2.[100,771,1473,1967]" pageId="2" pageNumber="69">
|
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|
Segment 1 with rounded sensory spots formed by a collar of short papillae surrounding at least one pore (
|
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|
<figureCitation id="130DC1C9DA3CFFECFE42A725FDF9F9F0" box="[503,577,1612,1632]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="2" pageNumber="69">Fig. 4B</figureCitation>
|
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|
); sensory spots are located near the anterior segment margin in subdorsal and laterodorsal positions, and centrally in ventromedial position (
|
|||
|
|
<figureCitation id="130DC1C9DA3CFFECFDF9A7EDFD35F907" box="[588,653,1668,1687]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="2" pageNumber="69">Figs. 2</figureCitation>
|
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,
|
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|
<figureCitation id="130DC1C9DA3CFFECFD2EA7EDFD77F907" box="[667,719,1668,1688]" captionStart="Fig" captionStartId="3.[100,130,1675,1692]" captionTargetBox="[147,1439,149,1646]" captionTargetId="figure-8@3.[146,1441,148,1647]" captionTargetPageId="3" captionText="Fig. 3. Light micrographs (DIC) showing traits in Echinoderes inaequalis sp. nov. Male paratype NHMD-1176545 (A, B, D, E, H), female holotype NHMD-1176540 (C, K, I) female paratype NHMD-1176543 (F, G), and male paratype NHMD-1176542 (J). (A) Male overview, ventral view. (B) Segments 1–2, ventral view. (C) Segments 1–2, dorsal view. (D) Segments 5–7, ventral view. (E) Segments 4–5, dorsal view. (F) Segments 7–8, lateroventral view. (G) Segments 6–7, dorsolateral view. (H) Male, segments 10–11 ventral view. (I) Female, segment 11, ventral view. (J) Segments 9–10, ventral view. (J’) Detail of sieve plate. (K) Segment 8, focused on laterodorsal area. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mdgco1, middorsal glandular cell outlet type 1; mlgco2, midlateral glandular cell outlet type 2; mlt, midlateral tube; mvp, midventral placid; pdgco1, paradorsal glandular cell outlet type 1; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; te, tergal extension; tp, trichoscalid plate; vlgco2, ventrolateral glandular cell outlet type 2; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164174" httpUri="https://zenodo.org/record/8164174/files/figure.png" pageId="2" pageNumber="69">3B–C</figureCitation>
|
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,
|
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|
|
<figureCitation id="130DC1C9DA3CFFECFD6EA7EDFD40F907" box="[731,760,1668,1687]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="2" pageNumber="69">4B</figureCitation>
|
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|
). Cuticular hairs are abundant and distributed forming a wide belt covering most of the dorsal and ventral surface of the segment, except for the anterior margin of the segment, ventral area with a less dense hair covering (
|
|||
|
|
<figureCitation id="130DC1C9DA3CFFECFF42A79DFE80F897" box="[247,312,1780,1799]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="2" pageNumber="69">Figs. 2</figureCitation>
|
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,
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<figureCitation id="130DC1C9DA3CFFECFEFDA79DFEC4F897" box="[328,380,1780,1799]" captionStart="Fig" captionStartId="3.[100,130,1675,1692]" captionTargetBox="[147,1439,149,1646]" captionTargetId="figure-8@3.[146,1441,148,1647]" captionTargetPageId="3" captionText="Fig. 3. Light micrographs (DIC) showing traits in Echinoderes inaequalis sp. nov. Male paratype NHMD-1176545 (A, B, D, E, H), female holotype NHMD-1176540 (C, K, I) female paratype NHMD-1176543 (F, G), and male paratype NHMD-1176542 (J). (A) Male overview, ventral view. (B) Segments 1–2, ventral view. (C) Segments 1–2, dorsal view. (D) Segments 5–7, ventral view. (E) Segments 4–5, dorsal view. (F) Segments 7–8, lateroventral view. (G) Segments 6–7, dorsolateral view. (H) Male, segments 10–11 ventral view. (I) Female, segment 11, ventral view. (J) Segments 9–10, ventral view. (J’) Detail of sieve plate. (K) Segment 8, focused on laterodorsal area. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mdgco1, middorsal glandular cell outlet type 1; mlgco2, midlateral glandular cell outlet type 2; mlt, midlateral tube; mvp, midventral placid; pdgco1, paradorsal glandular cell outlet type 1; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; te, tergal extension; tp, trichoscalid plate; vlgco2, ventrolateral glandular cell outlet type 2; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164174" httpUri="https://zenodo.org/record/8164174/files/figure.png" pageId="2" pageNumber="69">3B–C</figureCitation>
|
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,
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|
|
<figureCitation id="130DC1C9DA3CFFECFE3CA79DFE1DF897" box="[393,421,1780,1799]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="2" pageNumber="69">4B</figureCitation>
|
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). All cuticular hairs emerging from round perforation sites in this and the following segments. The posterior segment margin is straight along the dorsal and lateral side but extends slightly posteriorly in the ventromedial and midventral areas.
|
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</paragraph>
|
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<paragraph id="8B89DD4CDA3CFFECFF31A60DFB20F9D3" blockId="2.[100,771,1473,1967]" lastBlockId="2.[818,1488,1473,1967]" pageId="2" pageNumber="69">
|
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Segment 2 with glandular cell outlets
|
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<typeStatus id="548D63EEDA3CFFECFE47A60DFDA3F8E7" box="[498,539,1892,1911]" pageId="2" pageNumber="69">type</typeStatus>
|
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2, located in subdorsal, laterodorsal and ventrolateral positions (
|
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|
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<figureCitation id="130DC1C9DA3CFFECFE58A616FDCDF803" box="[493,629,1919,1939]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="2" pageNumber="69">Fig. 2A and B</figureCitation>
|
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,
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<figureCitation id="130DC1C9DA3CFFECFD34A616FD0DF802" box="[641,693,1919,1939]" captionStart="Fig" captionStartId="3.[100,130,1675,1692]" captionTargetBox="[147,1439,149,1646]" captionTargetId="figure-8@3.[146,1441,148,1647]" captionTargetPageId="3" captionText="Fig. 3. Light micrographs (DIC) showing traits in Echinoderes inaequalis sp. nov. Male paratype NHMD-1176545 (A, B, D, E, H), female holotype NHMD-1176540 (C, K, I) female paratype NHMD-1176543 (F, G), and male paratype NHMD-1176542 (J). (A) Male overview, ventral view. (B) Segments 1–2, ventral view. (C) Segments 1–2, dorsal view. (D) Segments 5–7, ventral view. (E) Segments 4–5, dorsal view. (F) Segments 7–8, lateroventral view. (G) Segments 6–7, dorsolateral view. (H) Male, segments 10–11 ventral view. (I) Female, segment 11, ventral view. (J) Segments 9–10, ventral view. (J’) Detail of sieve plate. (K) Segment 8, focused on laterodorsal area. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mdgco1, middorsal glandular cell outlet type 1; mlgco2, midlateral glandular cell outlet type 2; mlt, midlateral tube; mvp, midventral placid; pdgco1, paradorsal glandular cell outlet type 1; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; te, tergal extension; tp, trichoscalid plate; vlgco2, ventrolateral glandular cell outlet type 2; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164174" httpUri="https://zenodo.org/record/8164174/files/figure.png" pageId="2" pageNumber="69">3B–C</figureCitation>
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,
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|
<figureCitation id="130DC1C9DA3CFFECFD75A616FD40F802" box="[704,760,1919,1938]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="2" pageNumber="69">4B, E</figureCitation>
|
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|
). These are small with delicate fringed margins. Sensory spots present in middorsal, laterodorsal, midlateral and ventromedial positions; sensory spots on this and the following segments are oval. Hairs densely distributed in a belt covering the segment showing hairless areas in the anterior margin of the segment in ventromedial position (
|
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|
|
<figureCitation id="130DC1C9DA3CFFECFAFAA77CFA7DF9B8" box="[1359,1477,1557,1576]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="2" pageNumber="69">Fig. 4B–C, E</figureCitation>
|
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|
). Hairs increasing in length posteriorly.
|
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|
|
</paragraph>
|
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|
|
<paragraph id="8B89DD4CDA3CFFECFCE4A724FB60F95F" blockId="2.[818,1488,1473,1967]" pageId="2" pageNumber="69">
|
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|
|
Segment 3 with sensory spots in subdorsal and sublateral positions (
|
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|
|
<figureCitation id="130DC1C9DA3CFFECFC8FA701FC02F9EB" box="[826,954,1640,1659]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="2" pageNumber="69">Fig. 2A and B</figureCitation>
|
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). Dorsal hair pattern dense, cuticular hairs on the ventral side are covering the sternal plates except for hairless areas on the anterior margin of the sternal plates and narrow patches in ventromedial positions in this and the following segments.
|
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|
|
</paragraph>
|
|||
|
|
<paragraph id="8B89DD4CDA3CFFECFCE4A7B1FC3FF8CB" blockId="2.[818,1488,1473,1967]" pageId="2" pageNumber="69">
|
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|
|
Segment 4 with a very short acicular middorsal spine (ca. 11 μm measured from SEM) (
|
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|
|
<figureCitation id="130DC1C9DA3CFFECFBB7A79DFB39F897" box="[1026,1153,1780,1799]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="2" pageNumber="69">Fig. 2A and B</figureCitation>
|
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,
|
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<figureCitation id="130DC1C9DA3CFFECFB3FA79DFB1CF897" box="[1162,1188,1780,1799]" captionStart="Fig" captionStartId="3.[100,130,1675,1692]" captionTargetBox="[147,1439,149,1646]" captionTargetId="figure-8@3.[146,1441,148,1647]" captionTargetPageId="3" captionText="Fig. 3. Light micrographs (DIC) showing traits in Echinoderes inaequalis sp. nov. Male paratype NHMD-1176545 (A, B, D, E, H), female holotype NHMD-1176540 (C, K, I) female paratype NHMD-1176543 (F, G), and male paratype NHMD-1176542 (J). (A) Male overview, ventral view. (B) Segments 1–2, ventral view. (C) Segments 1–2, dorsal view. (D) Segments 5–7, ventral view. (E) Segments 4–5, dorsal view. (F) Segments 7–8, lateroventral view. (G) Segments 6–7, dorsolateral view. (H) Male, segments 10–11 ventral view. (I) Female, segment 11, ventral view. (J) Segments 9–10, ventral view. (J’) Detail of sieve plate. (K) Segment 8, focused on laterodorsal area. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mdgco1, middorsal glandular cell outlet type 1; mlgco2, midlateral glandular cell outlet type 2; mlt, midlateral tube; mvp, midventral placid; pdgco1, paradorsal glandular cell outlet type 1; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; te, tergal extension; tp, trichoscalid plate; vlgco2, ventrolateral glandular cell outlet type 2; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164174" httpUri="https://zenodo.org/record/8164174/files/figure.png" pageId="2" pageNumber="69">3E</figureCitation>
|
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,
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<figureCitation id="130DC1C9DA3CFFECFB18A79DFB72F897" box="[1197,1226,1780,1799]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="2" pageNumber="69">4C</figureCitation>
|
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|
|
). Glandular cell outlets
|
|||
|
|
<typeStatus id="548D63EEDA3CFFECFA13A79DFA77F897" box="[1446,1487,1780,1799]" pageId="2" pageNumber="69">type</typeStatus>
|
|||
|
|
2 present in subdorsal position (
|
|||
|
|
<figureCitation id="130DC1C9DA3CFFECFBC4A679FB7AF8B3" box="[1137,1218,1808,1827]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="2" pageNumber="69">Figs. 2A</figureCitation>
|
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|
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,
|
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|
|
<figureCitation id="130DC1C9DA3CFFECFB7AA679FB50F8B3" box="[1231,1256,1808,1827]" captionStart="Fig" captionStartId="3.[100,130,1675,1692]" captionTargetBox="[147,1439,149,1646]" captionTargetId="figure-8@3.[146,1441,148,1647]" captionTargetPageId="3" captionText="Fig. 3. Light micrographs (DIC) showing traits in Echinoderes inaequalis sp. nov. Male paratype NHMD-1176545 (A, B, D, E, H), female holotype NHMD-1176540 (C, K, I) female paratype NHMD-1176543 (F, G), and male paratype NHMD-1176542 (J). (A) Male overview, ventral view. (B) Segments 1–2, ventral view. (C) Segments 1–2, dorsal view. (D) Segments 5–7, ventral view. (E) Segments 4–5, dorsal view. (F) Segments 7–8, lateroventral view. (G) Segments 6–7, dorsolateral view. (H) Male, segments 10–11 ventral view. (I) Female, segment 11, ventral view. (J) Segments 9–10, ventral view. (J’) Detail of sieve plate. (K) Segment 8, focused on laterodorsal area. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mdgco1, middorsal glandular cell outlet type 1; mlgco2, midlateral glandular cell outlet type 2; mlt, midlateral tube; mvp, midventral placid; pdgco1, paradorsal glandular cell outlet type 1; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; te, tergal extension; tp, trichoscalid plate; vlgco2, ventrolateral glandular cell outlet type 2; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164174" httpUri="https://zenodo.org/record/8164174/files/figure.png" pageId="2" pageNumber="69">3E</figureCitation>
|
|||
|
|
and
|
|||
|
|
<figureCitation id="130DC1C9DA3CFFECFAA8A679FA82F8B3" box="[1309,1338,1808,1827]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="2" pageNumber="69">4C</figureCitation>
|
|||
|
|
). Sensory spots present in laterodorsal position. Other characters similar to previous segment.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B89DD4CDA3CFFEBFCE4A60DFE6FFB43" blockId="2.[818,1488,1473,1967]" lastBlockId="5.[100,771,1132,1486]" lastPageId="5" lastPageNumber="72" pageId="2" pageNumber="69">
|
|||
|
|
Segment 5 with lateroventral tubes and a single additional tube in subdorsal position (
|
|||
|
|
<figureCitation id="130DC1C9DA3CFFECFC5EA616FBD5F803" box="[1003,1133,1919,1939]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="2" pageNumber="69">Fig. 2A and B</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="130DC1C9DA3CFFECFBC3A616FBC7F802" box="[1142,1151,1919,1938]" captionStart="Fig" captionStartId="3.[100,130,1675,1692]" captionTargetBox="[147,1439,149,1646]" captionTargetId="figure-8@3.[146,1441,148,1647]" captionTargetPageId="3" captionText="Fig. 3. Light micrographs (DIC) showing traits in Echinoderes inaequalis sp. nov. Male paratype NHMD-1176545 (A, B, D, E, H), female holotype NHMD-1176540 (C, K, I) female paratype NHMD-1176543 (F, G), and male paratype NHMD-1176542 (J). (A) Male overview, ventral view. (B) Segments 1–2, ventral view. (C) Segments 1–2, dorsal view. (D) Segments 5–7, ventral view. (E) Segments 4–5, dorsal view. (F) Segments 7–8, lateroventral view. (G) Segments 6–7, dorsolateral view. (H) Male, segments 10–11 ventral view. (I) Female, segment 11, ventral view. (J) Segments 9–10, ventral view. (J’) Detail of sieve plate. (K) Segment 8, focused on laterodorsal area. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mdgco1, middorsal glandular cell outlet type 1; mlgco2, midlateral glandular cell outlet type 2; mlt, midlateral tube; mvp, midventral placid; pdgco1, paradorsal glandular cell outlet type 1; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; te, tergal extension; tp, trichoscalid plate; vlgco2, ventrolateral glandular cell outlet type 2; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164174" httpUri="https://zenodo.org/record/8164174/files/figure.png" pageId="2" pageNumber="69">3</figureCitation>
|
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|
D-E, 4D, G). Each tube consists of a short and smooth basal part, and a longer distal part in this and the following segments (
|
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|
|
<figureCitation id="130DC1C9DA3BFFEBFE99A505FECEFBEF" box="[300,374,1132,1151]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="5" pageNumber="72">Fig. 4G</figureCitation>
|
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). Glandular cell outlets
|
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|
|
<typeStatus id="548D63EEDA3BFFEBFDE8A505FD3EFBEF" box="[605,646,1132,1151]" pageId="5" pageNumber="72">type</typeStatus>
|
|||
|
|
2 present in midlateral position (
|
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|
|
<figureCitation id="130DC1C9DA3BFFEBFE99A5E1FEC3FB0B" box="[300,379,1160,1179]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="5" pageNumber="72">Figs. 2A</figureCitation>
|
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|
and
|
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|
|
<figureCitation id="130DC1C9DA3BFFEBFE07A5E1FE69FB0B" box="[434,465,1160,1179]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="5" pageNumber="72">4G</figureCitation>
|
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|
|
). Sensory spots present in subdorsal, laterodorsal and ventromedial positions (
|
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|
|
<figureCitation id="130DC1C9DA3BFFEBFD81A5CAFD02FB27" box="[564,698,1187,1207]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="5" pageNumber="72">Fig. 2A and B</figureCitation>
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). Other characters similar to previous segment.
|
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|
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</paragraph>
|
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|
|
<caption id="DF498DC4DA3DFFEDFFD1A7E2FCDEF811" ID-DOI="http://doi.org/10.5281/zenodo.8164174" ID-Zenodo-Dep="8164174" httpUri="https://zenodo.org/record/8164174/files/figure.png" pageId="3" pageNumber="70" startId="3.[100,130,1675,1692]" targetBox="[147,1439,149,1646]" targetPageId="3" targetType="figure">
|
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|
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<paragraph id="8B89DD4CDA3DFFEDFFD1A7E2FCDEF811" blockId="3.[100,1488,1674,1921]" pageId="3" pageNumber="70">
|
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|
|
<emphasis id="B942015EDA3DFFEDFFD1A7E2FF25F90C" bold="true" box="[100,157,1675,1692]" pageId="3" pageNumber="70">Fig. 3.</emphasis>
|
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|
|
Light micrographs (DIC) showing traits in
|
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|
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<taxonomicName id="4C36A6CFDA3DFFEDFDBFA7E3FD06F90C" authority="Herranz & Moreleon & Rho & Sørensen, 2023" authorityName="Herranz & Moreleon & Rho & Sørensen" authorityYear="2023" box="[522,702,1674,1692]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="3" pageNumber="70" phylum="Kinorhyncha" rank="species" species="inaequalis" status="sp. nov.">
|
|||
|
|
<emphasis id="B942015EDA3DFFEDFDBFA7E3FD06F90C" bold="true" box="[522,702,1674,1692]" italics="true" pageId="3" pageNumber="70">Echinoderes inaequalis</emphasis>
|
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|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A271BC25DA3DFFEDFD76A7E2FCBEF90B" box="[707,774,1675,1691]" pageId="3" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
|
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|
|
Male paratype NHMD-1176545 (A, B, D, E, H), female holotype NHMD-1176540 (C, K, I) female paratype NHMD-1176543 (F, G), and male paratype NHMD-1176542 (J). (A) Male overview, ventral view. (B) Segments 1–2, ventral view. (C) Segments 1–2, dorsal view. (D) Segments 5–7, ventral view. (E) Segments 4–5, dorsal view. (F) Segments 7–8, lateroventral view. (G) Segments 6–7, dorsolateral view. (H) Male, segments 10–11 ventral view. (I) Female, segment 11, ventral view. (J) Segments 9–10, ventral view. (J’) Detail of sieve plate. (K) Segment 8, focused on laterodorsal area. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mdgco1, middorsal glandular cell outlet type 1; mlgco2, midlateral glandular cell outlet type 2; mlt, midlateral tube; mvp, midventral placid; pdgco1, paradorsal glandular cell outlet type 1; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; te, tergal extension; tp, trichoscalid plate; vlgco2, ventrolateral glandular cell outlet type 2; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="DF498DC4DA3AFFEAFFD1A7E2FD21F811" ID-DOI="http://doi.org/10.5281/zenodo.8164177" ID-Zenodo-Dep="8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="4" pageNumber="71" startId="4.[100,130,1675,1692]" targetBox="[147,1440,149,1646]" targetPageId="4" targetType="figure">
|
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|
|
<paragraph id="8B89DD4CDA3AFFEAFFD1A7E2FD21F811" blockId="4.[100,1488,1674,1921]" pageId="4" pageNumber="71">
|
|||
|
|
<emphasis id="B942015EDA3AFFEAFFD1A7E2FF26F90C" bold="true" box="[100,158,1675,1692]" pageId="4" pageNumber="71">Fig. 4.</emphasis>
|
|||
|
|
Scanning electron micrographs (SEM) showing overviews and details of
|
|||
|
|
<taxonomicName id="4C36A6CFDA3AFFEAFCA7A7E3FC70F90C" authority="Herranz & Moreleon & Rho & Sørensen, 2023" authorityName="Herranz & Moreleon & Rho & Sørensen" authorityYear="2023" box="[786,968,1674,1692]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="4" pageNumber="71" phylum="Kinorhyncha" rank="species" species="inaequalis" status="sp. nov.">
|
|||
|
|
<emphasis id="B942015EDA3AFFEAFCA7A7E3FC70F90C" bold="true" box="[786,968,1674,1692]" italics="true" pageId="4" pageNumber="71">Echinoderes inaequalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A271BC25DA3AFFEAFC7BA7E2FBAAF90B" box="[974,1042,1675,1691]" pageId="4" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
(A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="DF498DC4DA3BFFEBFFD1A1FFFE73FECA" ID-Table-UUID="DF498DC4DA3BFFEBFFD1A1FFFE73FECA" httpUri="http://table.plazi.org/id/DF498DC4DA3BFFEBFFD1A1FFFE73FECA" pageId="5" pageNumber="72" startId="5.[100,150,150,166]" targetBox="[116,753,366,1078]" targetIsTable="true" targetPageId="5" targetType="table">
|
|||
|
|
<paragraph id="8B89DD4CDA3BFFEBFFD1A1FFFF1FFF37" blockId="5.[100,771,150,346]" box="[100,167,150,167]" pageId="5" pageNumber="72">
|
|||
|
|
<emphasis id="B942015EDA3BFFEBFFD1A1FFFF1FFF37" bold="true" box="[100,167,150,167]" pageId="5" pageNumber="72">Table 1</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B89DD4CDA3BFFEBFFD1A1D9FE73FECA" blockId="5.[100,771,150,346]" pageId="5" pageNumber="72">
|
|||
|
|
Measurements (in μm) from light microscopy (except for the middorsal spine of segment 4) of adult
|
|||
|
|
<taxonomicName id="4C36A6CFDA3BFFEBFEA5A1A1FE7DFF4A" authority="Herranz & Moreleon & Rho & Sørensen, 2023" authorityName="Herranz & Moreleon & Rho & Sørensen" authorityYear="2023" box="[272,453,200,218]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="5" pageNumber="72" phylum="Kinorhyncha" rank="species" species="inaequalis" status="sp. nov.">
|
|||
|
|
<emphasis id="B942015EDA3BFFEBFEA5A1A1FE7DFF4A" bold="true" box="[272,453,200,218]" italics="true" pageId="5" pageNumber="72">Echinoderes inaequalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A271BC25DA3BFFEBFE7EA1A3FDB7FF4A" box="[459,527,202,218]" pageId="5" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
Abbreviations: LTAS, lateral terminal accessory spine; LTS, lateral terminal spine; LVT, lateroventral tube; MD, middorsal spine; MLT, midlateral tube; MSW, maximum sternal width; n, number of specimens; SD, standard deviation; SDT, subdorsal tube; SI, sieve plate length; SW, standard width; S, segment lengths; TL, trunk length.Numbers, where inserted, indicate segment number.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8B89DD4CDA3BFFEBFFC1A007FD50FBA5" pageId="5" pageNumber="72">
|
|||
|
|
<table id="F9362FECDA3B0011FFC1A007FD49FBA6" box="[116,753,366,1078]" gridcols="5" gridrows="30" pageId="5" pageNumber="72">
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A007FD49FEEC" box="[116,753,366,380]" gridrow="0" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A007FF60FEEC" box="[116,216,366,380]" gridcol="0" gridrow="0" pageId="5" pageNumber="72">Character</th>
|
|||
|
|
<th id="76D7B672DA3B0011FE8CA007FEFBFEEC" box="[313,323,366,380]" gridcol="1" gridrow="0" pageId="5" pageNumber="72">n</th>
|
|||
|
|
<th id="76D7B672DA3B0011FE16A007FE5DFEEC" box="[419,485,366,380]" gridcol="2" gridrow="0" pageId="5" pageNumber="72">Range</th>
|
|||
|
|
<th id="76D7B672DA3B0011FDF3A007FDC8FEEC" box="[582,624,366,380]" gridcol="3" gridrow="0" pageId="5" pageNumber="72">Mean</th>
|
|||
|
|
<th id="76D7B672DA3B0011FD65A007FD49FEEC" box="[720,753,366,380]" gridcol="4" gridrow="0" pageId="5" pageNumber="72">SD</th>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A0F8FD49FE0F" box="[116,753,401,415]" gridrow="1" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A0F8FF60FE0F" box="[116,216,401,415]" gridcol="0" gridrow="1" pageId="5" pageNumber="72">TL</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA0F8FEFBFE0F" box="[313,323,401,415]" gridcol="1" gridrow="1" pageId="5" pageNumber="72">6</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A0F8FE5DFE0F" box="[419,485,401,415]" gridcol="2" gridrow="1" pageId="5" pageNumber="72">359–386</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A0F8FDC8FE0F" box="[582,624,401,415]" gridcol="3" gridrow="1" pageId="5" pageNumber="72">372</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A0F8FD49FE0F" box="[720,753,401,415]" gridcol="4" gridrow="1" pageId="5" pageNumber="72">10.6</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A0C1FD49FE26" box="[116,753,424,438]" gridrow="2" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A0C1FF60FE26" box="[116,216,424,438]" gridcol="0" gridrow="2" pageId="5" pageNumber="72">MSW (8)</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA0C1FEFBFE26" box="[313,323,424,438]" gridcol="1" gridrow="2" pageId="5" pageNumber="72">6</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A0C1FE5DFE26" box="[419,485,424,438]" gridcol="2" gridrow="2" pageId="5" pageNumber="72">64–72</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A0C1FDC8FE26" box="[582,624,424,438]" gridcol="3" gridrow="2" pageId="5" pageNumber="72">69</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A0C1FD49FE26" box="[720,753,424,438]" gridcol="4" gridrow="2" pageId="5" pageNumber="72">3.3</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A0D7FD49FE5D" box="[116,753,446,461]" gridrow="3" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A0D7FF60FE5D" box="[116,216,446,461]" gridcol="0" gridrow="3" pageId="5" pageNumber="72">MSW/TL (%)</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA0D7FEFBFE5D" box="[313,323,446,461]" gridcol="1" gridrow="3" pageId="5" pageNumber="72">6</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A0D7FE5DFE5D" box="[419,485,446,461]" gridcol="2" gridrow="3" pageId="5" pageNumber="72">18–19</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A0D7FDC8FE5D" box="[582,624,446,461]" gridcol="3" gridrow="3" pageId="5" pageNumber="72">18</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A0D7FD49FE5D" box="[720,753,446,461]" gridcol="4" gridrow="3" pageId="5" pageNumber="72">0.5</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A0BCFD49FE74" box="[116,753,469,484]" gridrow="4" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A0BCFF60FE74" box="[116,216,469,484]" gridcol="0" gridrow="4" pageId="5" pageNumber="72">SW</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA0BCFEFBFE74" box="[313,323,469,484]" gridcol="1" gridrow="4" pageId="5" pageNumber="72">6</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A0BCFE5DFE74" box="[419,485,469,484]" gridcol="2" gridrow="4" pageId="5" pageNumber="72">57–65</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A0BCFDC8FE74" box="[582,624,469,484]" gridcol="3" gridrow="4" pageId="5" pageNumber="72">61</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A0BCFD49FE74" box="[720,753,469,484]" gridcol="4" gridrow="4" pageId="5" pageNumber="72">2.8</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A085FD49FE6A" box="[116,753,492,506]" gridrow="5" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A085FF60FE6A" box="[116,216,492,506]" gridcol="0" gridrow="5" pageId="5" pageNumber="72">SW/TL (%)</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA085FEFBFE6A" box="[313,323,492,506]" gridcol="1" gridrow="5" pageId="5" pageNumber="72">6</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A085FE5DFE6A" box="[419,485,492,506]" gridcol="2" gridrow="5" pageId="5" pageNumber="72">16–17</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A085FDC8FE6A" box="[582,624,492,506]" gridcol="3" gridrow="5" pageId="5" pageNumber="72">16</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A085FD49FE6A" box="[720,753,492,506]" gridcol="4" gridrow="5" pageId="5" pageNumber="72">0.6</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A36AFD49FD81" box="[116,753,515,529]" gridrow="6" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A36AFF60FD81" box="[116,216,515,529]" gridcol="0" gridrow="6" pageId="5" pageNumber="72">S1</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA36AFEFBFD81" box="[313,323,515,529]" gridcol="1" gridrow="6" pageId="5" pageNumber="72">6</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A36AFE5DFD81" box="[419,485,515,529]" gridcol="2" gridrow="6" pageId="5" pageNumber="72">29–31</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A36AFDC8FD81" box="[582,624,515,529]" gridcol="3" gridrow="6" pageId="5" pageNumber="72">30</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A36AFD49FD81" box="[720,753,515,529]" gridcol="4" gridrow="6" pageId="5" pageNumber="72">0.9</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A373FD49FDB8" box="[116,753,538,552]" gridrow="7" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A373FF60FDB8" box="[116,216,538,552]" gridcol="0" gridrow="7" pageId="5" pageNumber="72">S2</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA373FEFBFDB8" box="[313,323,538,552]" gridcol="1" gridrow="7" pageId="5" pageNumber="72">6</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A373FE5DFDB8" box="[419,485,538,552]" gridcol="2" gridrow="7" pageId="5" pageNumber="72">26–31</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A373FDC8FDB8" box="[582,624,538,552]" gridcol="3" gridrow="7" pageId="5" pageNumber="72">29</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A373FD49FDB8" box="[720,753,538,552]" gridcol="4" gridrow="7" pageId="5" pageNumber="72">2.0</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A358FD49FDAF" box="[116,753,561,575]" gridrow="8" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A358FF60FDAF" box="[116,216,561,575]" gridcol="0" gridrow="8" pageId="5" pageNumber="72">S3</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA358FEFBFDAF" box="[313,323,561,575]" gridcol="1" gridrow="8" pageId="5" pageNumber="72">6</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A358FE5DFDAF" box="[419,485,561,575]" gridcol="2" gridrow="8" pageId="5" pageNumber="72">30–34</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A358FDC8FDAF" box="[582,624,561,575]" gridcol="3" gridrow="8" pageId="5" pageNumber="72">32</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A358FD49FDAF" box="[720,753,561,575]" gridcol="4" gridrow="8" pageId="5" pageNumber="72">1.4</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A32EFD49FDC6" box="[116,753,583,598]" gridrow="9" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A32EFF60FDC6" box="[116,216,583,598]" gridcol="0" gridrow="9" pageId="5" pageNumber="72">S4</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA32EFEFBFDC6" box="[313,323,583,598]" gridcol="1" gridrow="9" pageId="5" pageNumber="72">6</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A32EFE5DFDC6" box="[419,485,583,598]" gridcol="2" gridrow="9" pageId="5" pageNumber="72">33–36</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A32EFDC8FDC6" box="[582,624,583,598]" gridcol="3" gridrow="9" pageId="5" pageNumber="72">34</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A32EFD49FDC6" box="[720,753,583,598]" gridcol="4" gridrow="9" pageId="5" pageNumber="72">1.2</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A337FD49FDFD" box="[116,753,606,621]" gridrow="10" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A337FF60FDFD" box="[116,216,606,621]" gridcol="0" gridrow="10" pageId="5" pageNumber="72">S5</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA337FEFBFDFD" box="[313,323,606,621]" gridcol="1" gridrow="10" pageId="5" pageNumber="72">6</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A337FE5DFDFD" box="[419,485,606,621]" gridcol="2" gridrow="10" pageId="5" pageNumber="72">36–40</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A337FDC8FDFD" box="[582,624,606,621]" gridcol="3" gridrow="10" pageId="5" pageNumber="72">37</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A337FD49FDFD" box="[720,753,606,621]" gridcol="4" gridrow="10" pageId="5" pageNumber="72">1.5</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A31CFD49FD14" box="[116,753,629,644]" gridrow="11" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A31CFF60FD14" box="[116,216,629,644]" gridcol="0" gridrow="11" pageId="5" pageNumber="72">S6</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA31CFEFBFD14" box="[313,323,629,644]" gridcol="1" gridrow="11" pageId="5" pageNumber="72">6</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A31CFE5DFD14" box="[419,485,629,644]" gridcol="2" gridrow="11" pageId="5" pageNumber="72">41–44</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A31CFDC8FD14" box="[582,624,629,644]" gridcol="3" gridrow="11" pageId="5" pageNumber="72">43</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A31CFD49FD14" box="[720,753,629,644]" gridcol="4" gridrow="11" pageId="5" pageNumber="72">1.4</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A3E5FD49FD0A" box="[116,753,652,666]" gridrow="12" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A3E5FF60FD0A" box="[116,216,652,666]" gridcol="0" gridrow="12" pageId="5" pageNumber="72">S7</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA3E5FEFBFD0A" box="[313,323,652,666]" gridcol="1" gridrow="12" pageId="5" pageNumber="72">6</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A3E5FE5DFD0A" box="[419,485,652,666]" gridcol="2" gridrow="12" pageId="5" pageNumber="72">44–48</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A3E5FDC8FD0A" box="[582,624,652,666]" gridcol="3" gridrow="12" pageId="5" pageNumber="72">46</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A3E5FD49FD0A" box="[720,753,652,666]" gridcol="4" gridrow="12" pageId="5" pageNumber="72">1.5</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A3CAFD49FD21" box="[116,753,675,689]" gridrow="13" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A3CAFF60FD21" box="[116,216,675,689]" gridcol="0" gridrow="13" pageId="5" pageNumber="72">S8</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA3CAFEFBFD21" box="[313,323,675,689]" gridcol="1" gridrow="13" pageId="5" pageNumber="72">6</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A3CAFE5DFD21" box="[419,485,675,689]" gridcol="2" gridrow="13" pageId="5" pageNumber="72">49–55</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A3CAFDC8FD21" box="[582,624,675,689]" gridcol="3" gridrow="13" pageId="5" pageNumber="72">52</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A3CAFD49FD21" box="[720,753,675,689]" gridcol="4" gridrow="13" pageId="5" pageNumber="72">2.2</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A3D3FD49FD58" box="[116,753,698,712]" gridrow="14" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A3D3FF60FD58" box="[116,216,698,712]" gridcol="0" gridrow="14" pageId="5" pageNumber="72">S9</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA3D3FEFBFD58" box="[313,323,698,712]" gridcol="1" gridrow="14" pageId="5" pageNumber="72">6</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A3D3FE5DFD58" box="[419,485,698,712]" gridcol="2" gridrow="14" pageId="5" pageNumber="72">46–55</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A3D3FDC8FD58" box="[582,624,698,712]" gridcol="3" gridrow="14" pageId="5" pageNumber="72">51</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A3D3FD49FD58" box="[720,753,698,712]" gridcol="4" gridrow="14" pageId="5" pageNumber="72">3.1</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A3B8FD49FD4F" box="[116,753,721,735]" gridrow="15" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A3B8FF60FD4F" box="[116,216,721,735]" gridcol="0" gridrow="15" pageId="5" pageNumber="72">S10</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA3B8FEFBFD4F" box="[313,323,721,735]" gridcol="1" gridrow="15" pageId="5" pageNumber="72">6</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A3B8FE5DFD4F" box="[419,485,721,735]" gridcol="2" gridrow="15" pageId="5" pageNumber="72">39–50</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A3B8FDC8FD4F" box="[582,624,721,735]" gridcol="3" gridrow="15" pageId="5" pageNumber="72">43</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A3B8FD49FD4F" box="[720,753,721,735]" gridcol="4" gridrow="15" pageId="5" pageNumber="72">4.0</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A38EFD49FD66" box="[116,753,743,758]" gridrow="16" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A38EFF60FD66" box="[116,216,743,758]" gridcol="0" gridrow="16" pageId="5" pageNumber="72">S11</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA38EFEFBFD66" box="[313,323,743,758]" gridcol="1" gridrow="16" pageId="5" pageNumber="72">6</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A38EFE5DFD66" box="[419,485,743,758]" gridcol="2" gridrow="16" pageId="5" pageNumber="72">33–41</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A38EFDC8FD66" box="[582,624,743,758]" gridcol="3" gridrow="16" pageId="5" pageNumber="72">38</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A38EFD49FD66" box="[720,753,743,758]" gridcol="4" gridrow="16" pageId="5" pageNumber="72">3.6</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A27CFD49FCB3" box="[116,753,789,803]" gridrow="17" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A27CFF60FCB3" box="[116,216,789,803]" gridcol="0" gridrow="17" pageId="5" pageNumber="72">MD4</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA27CFEFBFCB3" box="[313,323,789,803]" gridcol="1" gridrow="17" pageId="5" pageNumber="72">1</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A27CFE5DFCB3" box="[419,485,789,803]" gridcol="2" gridrow="17" pageId="5" pageNumber="72">11</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A27CFDC8FCB3" box="[582,624,789,803]" gridcol="3" gridrow="17" pageId="5" pageNumber="72">–</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A27CFD49FCB3" box="[720,753,789,803]" gridcol="4" gridrow="17" pageId="5" pageNumber="72">–</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A245FD49FCAA" box="[116,753,812,826]" gridrow="18" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A245FF60FCAA" box="[116,216,812,826]" gridcol="0" gridrow="18" pageId="5" pageNumber="72">SDT5</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA245FEFBFCAA" box="[313,323,812,826]" gridcol="1" gridrow="18" pageId="5" pageNumber="72">1</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A245FE5DFCAA" box="[419,485,812,826]" gridcol="2" gridrow="18" pageId="5" pageNumber="72">18</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A245FDC8FCAA" box="[582,624,812,826]" gridcol="3" gridrow="18" pageId="5" pageNumber="72">–</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A245FD49FCAA" box="[720,753,812,826]" gridcol="4" gridrow="18" pageId="5" pageNumber="72">–</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A22AFD49FCC1" box="[116,753,835,849]" gridrow="19" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A22AFF60FCC1" box="[116,216,835,849]" gridcol="0" gridrow="19" pageId="5" pageNumber="72">SDT6</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA22AFEFBFCC1" box="[313,323,835,849]" gridcol="1" gridrow="19" pageId="5" pageNumber="72">2</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A22AFE5DFCC1" box="[419,485,835,849]" gridcol="2" gridrow="19" pageId="5" pageNumber="72">12–17</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A22AFDC8FCC1" box="[582,624,835,849]" gridcol="3" gridrow="19" pageId="5" pageNumber="72">15</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A22AFD49FCC1" box="[720,753,835,849]" gridcol="4" gridrow="19" pageId="5" pageNumber="72">2.8</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A233FD49FCF8" box="[116,753,858,872]" gridrow="20" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A233FF60FCF8" box="[116,216,858,872]" gridcol="0" gridrow="20" pageId="5" pageNumber="72">SDT7</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA233FEFBFCF8" box="[313,323,858,872]" gridcol="1" gridrow="20" pageId="5" pageNumber="72">1</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A233FE5DFCF8" box="[419,485,858,872]" gridcol="2" gridrow="20" pageId="5" pageNumber="72">15</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A233FDC8FCF8" box="[582,624,858,872]" gridcol="3" gridrow="20" pageId="5" pageNumber="72">–</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A233FD49FCF8" box="[720,753,858,872]" gridcol="4" gridrow="20" pageId="5" pageNumber="72">–</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A218FD49FCEF" box="[116,753,881,895]" gridrow="21" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A218FF60FCEF" box="[116,216,881,895]" gridcol="0" gridrow="21" pageId="5" pageNumber="72">MLT8</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA218FEFBFCEF" box="[313,323,881,895]" gridcol="1" gridrow="21" pageId="5" pageNumber="72">3</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A218FE5DFCEF" box="[419,485,881,895]" gridcol="2" gridrow="21" pageId="5" pageNumber="72">10–20</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A218FDC8FCEF" box="[582,624,881,895]" gridcol="3" gridrow="21" pageId="5" pageNumber="72">16</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A218FD49FCEF" box="[720,753,881,895]" gridcol="4" gridrow="21" pageId="5" pageNumber="72">5.1</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A2EEFD49FC06" box="[116,753,903,918]" gridrow="22" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A2EEFF60FC06" box="[116,216,903,918]" gridcol="0" gridrow="22" pageId="5" pageNumber="72">LVT5</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA2EEFEFBFC06" box="[313,323,903,918]" gridcol="1" gridrow="22" pageId="5" pageNumber="72">4</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A2EEFE5DFC06" box="[419,485,903,918]" gridcol="2" gridrow="22" pageId="5" pageNumber="72">10–22</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A2EEFDC8FC06" box="[582,624,903,918]" gridcol="3" gridrow="22" pageId="5" pageNumber="72">14</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A2EEFD49FC06" box="[720,753,903,918]" gridcol="4" gridrow="22" pageId="5" pageNumber="72">5.5</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A2F7FD49FC3D" box="[116,753,926,941]" gridrow="23" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A2F7FF60FC3D" box="[116,216,926,941]" gridcol="0" gridrow="23" pageId="5" pageNumber="72">LVT7</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA2F7FEFBFC3D" box="[313,323,926,941]" gridcol="1" gridrow="23" pageId="5" pageNumber="72">3</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A2F7FE5DFC3D" box="[419,485,926,941]" gridcol="2" gridrow="23" pageId="5" pageNumber="72">10–15</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A2F7FDC8FC3D" box="[582,624,926,941]" gridcol="3" gridrow="23" pageId="5" pageNumber="72">13</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A2F7FD49FC3D" box="[720,753,926,941]" gridcol="4" gridrow="23" pageId="5" pageNumber="72">2.5</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A2DCFD49FC53" box="[116,753,949,963]" gridrow="24" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A2DCFF60FC53" box="[116,216,949,963]" gridcol="0" gridrow="24" pageId="5" pageNumber="72">LVT8</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA2DCFEFBFC53" box="[313,323,949,963]" gridcol="1" gridrow="24" pageId="5" pageNumber="72">2</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A2DCFE5DFC53" box="[419,485,949,963]" gridcol="2" gridrow="24" pageId="5" pageNumber="72">15–16</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A2DCFDC8FC53" box="[582,624,949,963]" gridcol="3" gridrow="24" pageId="5" pageNumber="72">16</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A2DCFD49FC53" box="[720,753,949,963]" gridcol="4" gridrow="24" pageId="5" pageNumber="72">0.7</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A2A5FD49FC4A" box="[116,753,972,986]" gridrow="25" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A2A5FF60FC4A" box="[116,216,972,986]" gridcol="0" gridrow="25" pageId="5" pageNumber="72">LVT9</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA2A5FEFBFC4A" box="[313,323,972,986]" gridcol="1" gridrow="25" pageId="5" pageNumber="72">2</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A2A5FE5DFC4A" box="[419,485,972,986]" gridcol="2" gridrow="25" pageId="5" pageNumber="72">20</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A2A5FDC8FC4A" box="[582,624,972,986]" gridcol="3" gridrow="25" pageId="5" pageNumber="72">20</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A2A5FD49FC4A" box="[720,753,972,986]" gridcol="4" gridrow="25" pageId="5" pageNumber="72">0</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A28AFD49FC61" box="[116,753,995,1009]" gridrow="26" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A28AFF60FC61" box="[116,216,995,1009]" gridcol="0" gridrow="26" pageId="5" pageNumber="72">SI</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA28AFEFBFC61" box="[313,323,995,1009]" gridcol="1" gridrow="26" pageId="5" pageNumber="72">2</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A28AFE5DFC61" box="[419,485,995,1009]" gridcol="2" gridrow="26" pageId="5" pageNumber="72">13–15</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A28AFDC8FC61" box="[582,624,995,1009]" gridcol="3" gridrow="26" pageId="5" pageNumber="72">14</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A28AFD49FC61" box="[720,753,995,1009]" gridcol="4" gridrow="26" pageId="5" pageNumber="72">1.4</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A293FD49FB98" box="[116,753,1018,1032]" gridrow="27" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A293FF60FB98" box="[116,216,1018,1032]" gridcol="0" gridrow="27" pageId="5" pageNumber="72">LTS</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA293FEFBFB98" box="[313,323,1018,1032]" gridcol="1" gridrow="27" pageId="5" pageNumber="72">6</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A293FE5DFB98" box="[419,485,1018,1032]" gridcol="2" gridrow="27" pageId="5" pageNumber="72">41–47</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A293FDC8FB98" box="[582,624,1018,1032]" gridcol="3" gridrow="27" pageId="5" pageNumber="72">43</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A293FD49FB98" box="[720,753,1018,1032]" gridcol="4" gridrow="27" pageId="5" pageNumber="72">2.4</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A579FD49FB8F" box="[116,753,1040,1055]" gridrow="28" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A579FF60FB8F" box="[116,216,1040,1055]" gridcol="0" gridrow="28" pageId="5" pageNumber="72">LTS/TL (%)</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA579FEFBFB8F" box="[313,323,1040,1055]" gridcol="1" gridrow="28" pageId="5" pageNumber="72">6</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A579FE5DFB8F" box="[419,485,1040,1055]" gridcol="2" gridrow="28" pageId="5" pageNumber="72">11–13</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A579FDC8FB8F" box="[582,624,1040,1055]" gridcol="3" gridrow="28" pageId="5" pageNumber="72">12</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A579FD49FB8F" box="[720,753,1040,1055]" gridcol="4" gridrow="28" pageId="5" pageNumber="72">0.7</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="3506DF0EDA3B0011FFC1A54EFD49FBA6" box="[116,753,1063,1078]" gridrow="29" pageId="5" pageNumber="72">
|
|||
|
|
<th id="76D7B672DA3B0011FFC1A54EFF60FBA6" box="[116,216,1063,1078]" gridcol="0" gridrow="29" pageId="5" pageNumber="72">LTAS</th>
|
|||
|
|
<td id="76D7B672DA3B0011FE8CA54EFEFBFBA6" box="[313,323,1063,1078]" gridcol="1" gridrow="29" pageId="5" pageNumber="72">2</td>
|
|||
|
|
<td id="76D7B672DA3B0011FE16A54EFE5DFBA6" box="[419,485,1063,1078]" gridcol="2" gridrow="29" pageId="5" pageNumber="72">20–22</td>
|
|||
|
|
<td id="76D7B672DA3B0011FDF3A54EFDC8FBA6" box="[582,624,1063,1078]" gridcol="3" gridrow="29" pageId="5" pageNumber="72">21</td>
|
|||
|
|
<td id="76D7B672DA3B0011FD65A54EFD49FBA6" box="[720,753,1063,1078]" gridcol="4" gridrow="29" pageId="5" pageNumber="72">1.4</td>
|
|||
|
|
</tr>
|
|||
|
|
</table>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B89DD4CDA3BFFEBFF31A5B2FEA0FAEA" blockId="5.[100,771,1132,1486]" pageId="5" pageNumber="72">
|
|||
|
|
Segment 6 with a single subdorsal tube usually, but not consistently, in alternating position in respect to the single tube on segment 5 (
|
|||
|
|
<figureCitation id="130DC1C9DA3BFFEBFFD9A47AFF01FAB6" box="[108,185,1299,1318]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="5" pageNumber="72">Figs. 2A</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="130DC1C9DA3BFFEBFF74A47AFF64FAB6" box="[193,220,1299,1318]" captionStart="Fig" captionStartId="3.[100,130,1675,1692]" captionTargetBox="[147,1439,149,1646]" captionTargetId="figure-8@3.[146,1441,148,1647]" captionTargetPageId="3" captionText="Fig. 3. Light micrographs (DIC) showing traits in Echinoderes inaequalis sp. nov. Male paratype NHMD-1176545 (A, B, D, E, H), female holotype NHMD-1176540 (C, K, I) female paratype NHMD-1176543 (F, G), and male paratype NHMD-1176542 (J). (A) Male overview, ventral view. (B) Segments 1–2, ventral view. (C) Segments 1–2, dorsal view. (D) Segments 5–7, ventral view. (E) Segments 4–5, dorsal view. (F) Segments 7–8, lateroventral view. (G) Segments 6–7, dorsolateral view. (H) Male, segments 10–11 ventral view. (I) Female, segment 11, ventral view. (J) Segments 9–10, ventral view. (J’) Detail of sieve plate. (K) Segment 8, focused on laterodorsal area. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mdgco1, middorsal glandular cell outlet type 1; mlgco2, midlateral glandular cell outlet type 2; mlt, midlateral tube; mvp, midventral placid; pdgco1, paradorsal glandular cell outlet type 1; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; te, tergal extension; tp, trichoscalid plate; vlgco2, ventrolateral glandular cell outlet type 2; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164174" httpUri="https://zenodo.org/record/8164174/files/figure.png" pageId="5" pageNumber="72">3G</figureCitation>
|
|||
|
|
and
|
|||
|
|
<figureCitation id="130DC1C9DA3BFFEBFEBDA47AFE9FFAB6" box="[264,295,1299,1318]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="5" pageNumber="72">4H</figureCitation>
|
|||
|
|
). Glandular cell outlets
|
|||
|
|
<typeStatus id="548D63EEDA3BFFEBFDB7A47AFD93FAB6" box="[514,555,1299,1318]" pageId="5" pageNumber="72">type</typeStatus>
|
|||
|
|
2 present in midlateral position (
|
|||
|
|
<figureCitation id="130DC1C9DA3BFFEBFF09A446FEBFFAD2" box="[188,263,1327,1346]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="5" pageNumber="72">Figs. 2A</figureCitation>
|
|||
|
|
and
|
|||
|
|
<figureCitation id="130DC1C9DA3BFFEBFE87A446FEE9FAD2" box="[306,337,1327,1346]" captionStart="Fig" captionStartId="3.[100,130,1675,1692]" captionTargetBox="[147,1439,149,1646]" captionTargetId="figure-8@3.[146,1441,148,1647]" captionTargetPageId="3" captionText="Fig. 3. Light micrographs (DIC) showing traits in Echinoderes inaequalis sp. nov. Male paratype NHMD-1176545 (A, B, D, E, H), female holotype NHMD-1176540 (C, K, I) female paratype NHMD-1176543 (F, G), and male paratype NHMD-1176542 (J). (A) Male overview, ventral view. (B) Segments 1–2, ventral view. (C) Segments 1–2, dorsal view. (D) Segments 5–7, ventral view. (E) Segments 4–5, dorsal view. (F) Segments 7–8, lateroventral view. (G) Segments 6–7, dorsolateral view. (H) Male, segments 10–11 ventral view. (I) Female, segment 11, ventral view. (J) Segments 9–10, ventral view. (J’) Detail of sieve plate. (K) Segment 8, focused on laterodorsal area. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mdgco1, middorsal glandular cell outlet type 1; mlgco2, midlateral glandular cell outlet type 2; mlt, midlateral tube; mvp, midventral placid; pdgco1, paradorsal glandular cell outlet type 1; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; te, tergal extension; tp, trichoscalid plate; vlgco2, ventrolateral glandular cell outlet type 2; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164174" httpUri="https://zenodo.org/record/8164174/files/figure.png" pageId="5" pageNumber="72">3G</figureCitation>
|
|||
|
|
). Sensory spots present in subdorsal, sublateral and ventromedial positions (
|
|||
|
|
<figureCitation id="130DC1C9DA3BFFEBFEC5A422FE49FACE" box="[368,497,1355,1374]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="5" pageNumber="72">Fig. 2A and B</figureCitation>
|
|||
|
|
). Remaining characters as on previous segments.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B89DD4CDA3BFFEBFF31A4EAFC7FFF6B" blockId="5.[100,771,1132,1486]" lastBlockId="5.[818,1488,148,1311]" pageId="5" pageNumber="72">
|
|||
|
|
Segment 7 with lateroventral tubes, and an additional single tube located in subdorsal position, the latter usually alternating in position with the unpaired tube of the preceding segment (
|
|||
|
|
<figureCitation id="130DC1C9DA3BFFEBFDF4A4D3FD7CFA5E" box="[577,708,1466,1486]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="5" pageNumber="72">Fig. 2A and B</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="130DC1C9DA3BFFEBFD7AA4D2FD60FA5E" box="[719,728,1467,1486]" captionStart="Fig" captionStartId="3.[100,130,1675,1692]" captionTargetBox="[147,1439,149,1646]" captionTargetId="figure-8@3.[146,1441,148,1647]" captionTargetPageId="3" captionText="Fig. 3. Light micrographs (DIC) showing traits in Echinoderes inaequalis sp. nov. Male paratype NHMD-1176545 (A, B, D, E, H), female holotype NHMD-1176540 (C, K, I) female paratype NHMD-1176543 (F, G), and male paratype NHMD-1176542 (J). (A) Male overview, ventral view. (B) Segments 1–2, ventral view. (C) Segments 1–2, dorsal view. (D) Segments 5–7, ventral view. (E) Segments 4–5, dorsal view. (F) Segments 7–8, lateroventral view. (G) Segments 6–7, dorsolateral view. (H) Male, segments 10–11 ventral view. (I) Female, segment 11, ventral view. (J) Segments 9–10, ventral view. (J’) Detail of sieve plate. (K) Segment 8, focused on laterodorsal area. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mdgco1, middorsal glandular cell outlet type 1; mlgco2, midlateral glandular cell outlet type 2; mlt, midlateral tube; mvp, midventral placid; pdgco1, paradorsal glandular cell outlet type 1; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; te, tergal extension; tp, trichoscalid plate; vlgco2, ventrolateral glandular cell outlet type 2; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164174" httpUri="https://zenodo.org/record/8164174/files/figure.png" pageId="5" pageNumber="72">3</figureCitation>
|
|||
|
|
F-G, 4H). Glandular cell outlets
|
|||
|
|
<typeStatus id="548D63EEDA3BFFEBFBE4A1FDFBC2FF37" box="[1105,1146,148,167]" pageId="5" pageNumber="72">type</typeStatus>
|
|||
|
|
2 present in midlateral position (
|
|||
|
|
<figureCitation id="130DC1C9DA3BFFEBFC8FA1D9FC3BFF53" box="[826,899,176,195]" captionStart="Fig" captionStartId="3.[100,130,1675,1692]" captionTargetBox="[147,1439,149,1646]" captionTargetId="figure-8@3.[146,1441,148,1647]" captionTargetPageId="3" captionText="Fig. 3. Light micrographs (DIC) showing traits in Echinoderes inaequalis sp. nov. Male paratype NHMD-1176545 (A, B, D, E, H), female holotype NHMD-1176540 (C, K, I) female paratype NHMD-1176543 (F, G), and male paratype NHMD-1176542 (J). (A) Male overview, ventral view. (B) Segments 1–2, ventral view. (C) Segments 1–2, dorsal view. (D) Segments 5–7, ventral view. (E) Segments 4–5, dorsal view. (F) Segments 7–8, lateroventral view. (G) Segments 6–7, dorsolateral view. (H) Male, segments 10–11 ventral view. (I) Female, segment 11, ventral view. (J) Segments 9–10, ventral view. (J’) Detail of sieve plate. (K) Segment 8, focused on laterodorsal area. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mdgco1, middorsal glandular cell outlet type 1; mlgco2, midlateral glandular cell outlet type 2; mlt, midlateral tube; mvp, midventral placid; pdgco1, paradorsal glandular cell outlet type 1; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; te, tergal extension; tp, trichoscalid plate; vlgco2, ventrolateral glandular cell outlet type 2; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164174" httpUri="https://zenodo.org/record/8164174/files/figure.png" pageId="5" pageNumber="72">Fig. 3G</figureCitation>
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). Sensory spots present in subdorsal, laterodorsal and ventrolateral positions (
|
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|
|
<figureCitation id="130DC1C9DA3BFFEBFC6CA1A2FBE7FF4F" box="[985,1119,203,223]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="5" pageNumber="72">Fig. 2A and B</figureCitation>
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,
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<figureCitation id="130DC1C9DA3BFFEBFBDFA1A5FB31FF4F" box="[1130,1161,204,223]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="5" pageNumber="72">4H</figureCitation>
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). Remaining characters as on previous segments.
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</paragraph>
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<paragraph id="8B89DD4CDA3BFFEBFCE4A06AFC37FE16" blockId="5.[818,1488,148,1311]" pageId="5" pageNumber="72">
|
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Segment 8 with tubes in lateroventral and midlateral positions (
|
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<figureCitation id="130DC1C9DA3BFFEBFC8FA076FC0EFEA3" box="[826,950,287,307]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="5" pageNumber="72">Fig. 2A and B</figureCitation>
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,
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<figureCitation id="130DC1C9DA3BFFEBFC0BA076FC61FEA2" box="[958,985,287,306]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="5" pageNumber="72">4F</figureCitation>
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). Large glandular cell outlets
|
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<typeStatus id="548D63EEDA3BFFEBFB52A049FAA8FEA3" box="[1255,1296,288,307]" pageId="5" pageNumber="72">type</typeStatus>
|
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2 with fringed edges present in laterodorsal position (
|
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<figureCitation id="130DC1C9DA3BFFEBFBDFA052FB48FEDE" box="[1130,1264,315,334]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="5" pageNumber="72">Fig. 2A and B</figureCitation>
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,
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<figureCitation id="130DC1C9DA3BFFEBFB4EA052FAAFFEDE" box="[1275,1303,315,334]" captionStart="Fig" captionStartId="3.[100,130,1675,1692]" captionTargetBox="[147,1439,149,1646]" captionTargetId="figure-8@3.[146,1441,148,1647]" captionTargetPageId="3" captionText="Fig. 3. Light micrographs (DIC) showing traits in Echinoderes inaequalis sp. nov. Male paratype NHMD-1176545 (A, B, D, E, H), female holotype NHMD-1176540 (C, K, I) female paratype NHMD-1176543 (F, G), and male paratype NHMD-1176542 (J). (A) Male overview, ventral view. (B) Segments 1–2, ventral view. (C) Segments 1–2, dorsal view. (D) Segments 5–7, ventral view. (E) Segments 4–5, dorsal view. (F) Segments 7–8, lateroventral view. (G) Segments 6–7, dorsolateral view. (H) Male, segments 10–11 ventral view. (I) Female, segment 11, ventral view. (J) Segments 9–10, ventral view. (J’) Detail of sieve plate. (K) Segment 8, focused on laterodorsal area. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mdgco1, middorsal glandular cell outlet type 1; mlgco2, midlateral glandular cell outlet type 2; mlt, midlateral tube; mvp, midventral placid; pdgco1, paradorsal glandular cell outlet type 1; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; te, tergal extension; tp, trichoscalid plate; vlgco2, ventrolateral glandular cell outlet type 2; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164174" httpUri="https://zenodo.org/record/8164174/files/figure.png" pageId="5" pageNumber="72">3K</figureCitation>
|
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,
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<figureCitation id="130DC1C9DA3BFFEBFA97A052FA85FEDE" box="[1314,1341,315,334]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="5" pageNumber="72">4F</figureCitation>
|
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|
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). Sensory spots present in subdorsal position. Other characters similar to previous segments.
|
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|
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</paragraph>
|
|||
|
|
<paragraph id="8B89DD4CDA3BFFEBFCE4A0E6FC5EFDBE" blockId="5.[818,1488,148,1311]" pageId="5" pageNumber="72">
|
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|
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Segment 9 with tubes in lateroventral position. Sensory spots present in subdorsal, laterodorsal, midlateral and ventrolateral positions (
|
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|
|
<figureCitation id="130DC1C9DA3BFFEBFC8FA0AEFC7CFE4A" box="[826,964,455,474]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="5" pageNumber="72">Fig. 2A and B</figureCitation>
|
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,
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<figureCitation id="130DC1C9DA3BFFEBFC64A0AEFC51FE4A" box="[977,1001,455,474]" captionStart="Fig" captionStartId="3.[100,130,1675,1692]" captionTargetBox="[147,1439,149,1646]" captionTargetId="figure-8@3.[146,1441,148,1647]" captionTargetPageId="3" captionText="Fig. 3. Light micrographs (DIC) showing traits in Echinoderes inaequalis sp. nov. Male paratype NHMD-1176545 (A, B, D, E, H), female holotype NHMD-1176540 (C, K, I) female paratype NHMD-1176543 (F, G), and male paratype NHMD-1176542 (J). (A) Male overview, ventral view. (B) Segments 1–2, ventral view. (C) Segments 1–2, dorsal view. (D) Segments 5–7, ventral view. (E) Segments 4–5, dorsal view. (F) Segments 7–8, lateroventral view. (G) Segments 6–7, dorsolateral view. (H) Male, segments 10–11 ventral view. (I) Female, segment 11, ventral view. (J) Segments 9–10, ventral view. (J’) Detail of sieve plate. (K) Segment 8, focused on laterodorsal area. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mdgco1, middorsal glandular cell outlet type 1; mlgco2, midlateral glandular cell outlet type 2; mlt, midlateral tube; mvp, midventral placid; pdgco1, paradorsal glandular cell outlet type 1; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; te, tergal extension; tp, trichoscalid plate; vlgco2, ventrolateral glandular cell outlet type 2; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164174" httpUri="https://zenodo.org/record/8164174/files/figure.png" pageId="5" pageNumber="72">3J</figureCitation>
|
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). Elongated nephridial areas present in sublateral position, and consisting of an oval perforated sieve plate (ca. 12 μm) with a posterior round pore (
|
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|
<figureCitation id="130DC1C9DA3BFFEBFBF6A097FBC0FD81" box="[1091,1144,510,529]" captionStart="Fig" captionStartId="3.[100,130,1675,1692]" captionTargetBox="[147,1439,149,1646]" captionTargetId="figure-8@3.[146,1441,148,1647]" captionTargetPageId="3" captionText="Fig. 3. Light micrographs (DIC) showing traits in Echinoderes inaequalis sp. nov. Male paratype NHMD-1176545 (A, B, D, E, H), female holotype NHMD-1176540 (C, K, I) female paratype NHMD-1176543 (F, G), and male paratype NHMD-1176542 (J). (A) Male overview, ventral view. (B) Segments 1–2, ventral view. (C) Segments 1–2, dorsal view. (D) Segments 5–7, ventral view. (E) Segments 4–5, dorsal view. (F) Segments 7–8, lateroventral view. (G) Segments 6–7, dorsolateral view. (H) Male, segments 10–11 ventral view. (I) Female, segment 11, ventral view. (J) Segments 9–10, ventral view. (J’) Detail of sieve plate. (K) Segment 8, focused on laterodorsal area. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mdgco1, middorsal glandular cell outlet type 1; mlgco2, midlateral glandular cell outlet type 2; mlt, midlateral tube; mvp, midventral placid; pdgco1, paradorsal glandular cell outlet type 1; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; te, tergal extension; tp, trichoscalid plate; vlgco2, ventrolateral glandular cell outlet type 2; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164174" httpUri="https://zenodo.org/record/8164174/files/figure.png" pageId="5" pageNumber="72">Fig. 3</figureCitation>
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J- J’, 4I). Other characters similar to previous segments.
|
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</paragraph>
|
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|
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<paragraph id="8B89DD4CDA3BFFEBFCE4A35FFB21FCF0" blockId="5.[818,1488,148,1311]" pageId="5" pageNumber="72">
|
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Segment 10 with tubes in laterodorsal positions, at or near the posterior segment margin. In males the tubes are long and similar of those described on segments 5–9 (Fig.. 2C and 4M), whereas in females the tubes lack the basal part, showing just a flexible and short tubular structure (
|
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|
|
<figureCitation id="130DC1C9DA3BFFEBFC20A3CFFC59FD29" box="[917,993,678,697]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="5" pageNumber="72">Figs. 2A</figureCitation>
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and
|
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<figureCitation id="130DC1C9DA3BFFEBFBA7A3CFFB92FD29" box="[1042,1066,678,697]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="5" pageNumber="72">4J</figureCitation>
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). Very elongated sensory spots composed of two or three rows of short parallel papillae present in subdorsal and ventrolateral positions (
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<figureCitation id="130DC1C9DA3BFFEBFBA6A3B4FBE9FD61" box="[1043,1105,733,753]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="5" pageNumber="72">Figs. 2</figureCitation>
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<figureCitation id="130DC1C9DA3BFFEBFBE8A3B7FBD1FD61" box="[1117,1129,734,753]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="5" pageNumber="72">4</figureCitation>
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J-K). The posterior segment margin of the tergal plate is straight and with a small, but well-developed, pectinate fringe, whereas the margins of the sternal plates are concave, extending posteriorly near the midventral junction, and with well-developed fringe tips (
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<figureCitation id="130DC1C9DA3BFFEBFBB3A224FBDEFCF0" box="[1030,1126,845,864]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="5" pageNumber="72">Fig. 2B, D</figureCitation>
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<figureCitation id="130DC1C9DA3BFFEBFBC5A224FB36FCF0" box="[1136,1166,845,864]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="5" pageNumber="72">4K</figureCitation>
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).
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</paragraph>
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<paragraph id="8B89DD4CDA3BFFEBFCE4A200FBD4FA8E" blockId="5.[818,1488,148,1311]" pageId="5" pageNumber="72">
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Segment 11 with short and stout lateral terminal spines with lateral fringes, and a short fringe wrapping the spine medially (
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<figureCitation id="130DC1C9DA3BFFEBFAF6A2ECFA38FC08" box="[1347,1408,901,920]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="5" pageNumber="72">Figs. 2</figureCitation>
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and
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<figureCitation id="130DC1C9DA3BFFEBFA1AA2ECFCEDFC24" captionStart="Fig" captionStartId="3.[100,130,1675,1692]" captionTargetBox="[147,1439,149,1646]" captionTargetId="figure-8@3.[146,1441,148,1647]" captionTargetPageId="3" captionText="Fig. 3. Light micrographs (DIC) showing traits in Echinoderes inaequalis sp. nov. Male paratype NHMD-1176545 (A, B, D, E, H), female holotype NHMD-1176540 (C, K, I) female paratype NHMD-1176543 (F, G), and male paratype NHMD-1176542 (J). (A) Male overview, ventral view. (B) Segments 1–2, ventral view. (C) Segments 1–2, dorsal view. (D) Segments 5–7, ventral view. (E) Segments 4–5, dorsal view. (F) Segments 7–8, lateroventral view. (G) Segments 6–7, dorsolateral view. (H) Male, segments 10–11 ventral view. (I) Female, segment 11, ventral view. (J) Segments 9–10, ventral view. (J’) Detail of sieve plate. (K) Segment 8, focused on laterodorsal area. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mdgco1, middorsal glandular cell outlet type 1; mlgco2, midlateral glandular cell outlet type 2; mlt, midlateral tube; mvp, midventral placid; pdgco1, paradorsal glandular cell outlet type 1; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; te, tergal extension; tp, trichoscalid plate; vlgco2, ventrolateral glandular cell outlet type 2; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164174" httpUri="https://zenodo.org/record/8164174/files/figure.png" pageId="5" pageNumber="72">3A, H–I</figureCitation>
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<figureCitation id="130DC1C9DA3BFFEBFCD4A2C8FCC2FC24" box="[865,890,929,948]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="5" pageNumber="72">4J</figureCitation>
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). Males with three pairs of tubular, and flexible penile spines, two of them long, and one slightly shorter (
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<figureCitation id="130DC1C9DA3BFFEBFB62A2D4FAE5FC40" box="[1239,1373,957,976]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="5" pageNumber="72">Fig. 2B and C</figureCitation>
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<figureCitation id="130DC1C9DA3BFFEBFAD2A2D4FA3DFC40" box="[1383,1413,957,976]" captionStart="Fig" captionStartId="3.[100,130,1675,1692]" captionTargetBox="[147,1439,149,1646]" captionTargetId="figure-8@3.[146,1441,148,1647]" captionTargetPageId="3" captionText="Fig. 3. Light micrographs (DIC) showing traits in Echinoderes inaequalis sp. nov. Male paratype NHMD-1176545 (A, B, D, E, H), female holotype NHMD-1176540 (C, K, I) female paratype NHMD-1176543 (F, G), and male paratype NHMD-1176542 (J). (A) Male overview, ventral view. (B) Segments 1–2, ventral view. (C) Segments 1–2, dorsal view. (D) Segments 5–7, ventral view. (E) Segments 4–5, dorsal view. (F) Segments 7–8, lateroventral view. (G) Segments 6–7, dorsolateral view. (H) Male, segments 10–11 ventral view. (I) Female, segment 11, ventral view. (J) Segments 9–10, ventral view. (J’) Detail of sieve plate. (K) Segment 8, focused on laterodorsal area. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mdgco1, middorsal glandular cell outlet type 1; mlgco2, midlateral glandular cell outlet type 2; mlt, midlateral tube; mvp, midventral placid; pdgco1, paradorsal glandular cell outlet type 1; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; te, tergal extension; tp, trichoscalid plate; vlgco2, ventrolateral glandular cell outlet type 2; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164174" httpUri="https://zenodo.org/record/8164174/files/figure.png" pageId="5" pageNumber="72">3H</figureCitation>
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<figureCitation id="130DC1C9DA3BFFEBFA25A2D4FA23FC40" box="[1424,1435,957,976]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="5" pageNumber="72">4</figureCitation>
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L-M). Females with a pair of short, thin and hairy lateral terminal accessory spines (
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<figureCitation id="130DC1C9DA3BFFEBFCCFA29DFC45FB98" box="[890,1021,1012,1032]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="5" pageNumber="72">Fig. 2A and B</figureCitation>
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<figureCitation id="130DC1C9DA3BFFEBFBBDA29CFBA2FB98" box="[1032,1050,1013,1032]" captionStart="Fig" captionStartId="3.[100,130,1675,1692]" captionTargetBox="[147,1439,149,1646]" captionTargetId="figure-8@3.[146,1441,148,1647]" captionTargetPageId="3" captionText="Fig. 3. Light micrographs (DIC) showing traits in Echinoderes inaequalis sp. nov. Male paratype NHMD-1176545 (A, B, D, E, H), female holotype NHMD-1176540 (C, K, I) female paratype NHMD-1176543 (F, G), and male paratype NHMD-1176542 (J). (A) Male overview, ventral view. (B) Segments 1–2, ventral view. (C) Segments 1–2, dorsal view. (D) Segments 5–7, ventral view. (E) Segments 4–5, dorsal view. (F) Segments 7–8, lateroventral view. (G) Segments 6–7, dorsolateral view. (H) Male, segments 10–11 ventral view. (I) Female, segment 11, ventral view. (J) Segments 9–10, ventral view. (J’) Detail of sieve plate. (K) Segment 8, focused on laterodorsal area. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mdgco1, middorsal glandular cell outlet type 1; mlgco2, midlateral glandular cell outlet type 2; mlt, midlateral tube; mvp, midventral placid; pdgco1, paradorsal glandular cell outlet type 1; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; te, tergal extension; tp, trichoscalid plate; vlgco2, ventrolateral glandular cell outlet type 2; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164174" httpUri="https://zenodo.org/record/8164174/files/figure.png" pageId="5" pageNumber="72">3I</figureCitation>
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<figureCitation id="130DC1C9DA3BFFEBFB92A29CFB8BFB98" box="[1063,1075,1013,1032]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="5" pageNumber="72">4</figureCitation>
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J-K). Sensory spots are present in subdorsal position (two pairs) and in ventromedial position. The ventromedial sensory spots are very small, and located at the posterior margin of the sternal plate (
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<figureCitation id="130DC1C9DA3BFFEBFC07A521FC40FBCB" box="[946,1016,1096,1115]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="5" pageNumber="72">Fig. 4K</figureCitation>
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). One of the subdorsal pairs of sensory spots is also small and located on the margin of the tergal extensions (
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<figureCitation id="130DC1C9DA3BFFEBFAD5A50DFA2EFBE7" box="[1376,1430,1124,1143]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="5" pageNumber="72">Fig. 2</figureCitation>
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). The tergal plate is covered in hair-like extensions without perforation sites; sternal plates with a ventromedial fringed area (
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<figureCitation id="130DC1C9DA3BFFEBFABEA5F5FAD7FB3F" box="[1291,1391,1180,1199]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="5" pageNumber="72">Fig. 2B, D</figureCitation>
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,
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<figureCitation id="130DC1C9DA3BFFEBFAC9A5F5FA22FB3F" box="[1404,1434,1180,1199]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="5" pageNumber="72">4K</figureCitation>
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). The margins of the sternal plates are curved and covered by long hairs that get shorter towards the lateroventral area (
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<figureCitation id="130DC1C9DA3BFFEBFB65A5BDFAA1FB77" box="[1232,1305,1236,1255]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="5" pageNumber="72">Fig. 4K</figureCitation>
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). Tergal extensions are pointed extending further than the sternal plates and with a small distal notch (
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<figureCitation id="130DC1C9DA3BFFEBFC1AA462FC54FA8E" box="[943,1004,1291,1310]" captionStart="Fig" captionStartId="2.[100,130,1313,1330]" captionTargetBox="[188,1399,149,1284]" captionTargetId="figure-421@2.[187,1400,148,1285]" captionTargetPageId="2" captionText="Fig. 2. Line art illustrations of Echinoderes inaequalis sp. nov. (A) Female, dorsal view. (B) Female, ventral view. (C) Male, detail of segments 10–11, dorsal view. (D) Male, detail of segments 10–11, ventral view. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdt, subdorsal tube; si, sieve plate; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmss, ventromedial sensory spot. Scale bar, 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8164169" httpUri="https://zenodo.org/record/8164169/files/figure.png" pageId="5" pageNumber="72">Figs. 2</figureCitation>
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,
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<figureCitation id="130DC1C9DA3BFFEBFC4CA462FB90FA8F" box="[1017,1064,1291,1311]" captionStart="Fig" captionStartId="3.[100,130,1675,1692]" captionTargetBox="[147,1439,149,1646]" captionTargetId="figure-8@3.[146,1441,148,1647]" captionTargetPageId="3" captionText="Fig. 3. Light micrographs (DIC) showing traits in Echinoderes inaequalis sp. nov. Male paratype NHMD-1176545 (A, B, D, E, H), female holotype NHMD-1176540 (C, K, I) female paratype NHMD-1176543 (F, G), and male paratype NHMD-1176542 (J). (A) Male overview, ventral view. (B) Segments 1–2, ventral view. (C) Segments 1–2, dorsal view. (D) Segments 5–7, ventral view. (E) Segments 4–5, dorsal view. (F) Segments 7–8, lateroventral view. (G) Segments 6–7, dorsolateral view. (H) Male, segments 10–11 ventral view. (I) Female, segment 11, ventral view. (J) Segments 9–10, ventral view. (J’) Detail of sieve plate. (K) Segment 8, focused on laterodorsal area. Abbreviations: ldgco2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mdgco1, middorsal glandular cell outlet type 1; mlgco2, midlateral glandular cell outlet type 2; mlt, midlateral tube; mvp, midventral placid; pdgco1, paradorsal glandular cell outlet type 1; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; te, tergal extension; tp, trichoscalid plate; vlgco2, ventrolateral glandular cell outlet type 2; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164174" httpUri="https://zenodo.org/record/8164174/files/figure.png" pageId="5" pageNumber="72">3H–I</figureCitation>
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,
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<figureCitation id="130DC1C9DA3BFFEBFB87A462FB86FA8E" box="[1074,1086,1291,1310]" captionStart="Fig" captionStartId="4.[100,130,1675,1692]" captionTargetBox="[147,1440,149,1646]" captionTargetId="figure-8@4.[146,1441,148,1647]" captionTargetPageId="4" captionText="Fig. 4. Scanning electron micrographs (SEM) showing overviews and details of Echinoderes inaequalis sp. nov. (A) Male, lateroventral overview. (B) Segments 1–2 ventral view. (C) Segments 2–4, dorsal view. (D) Segments 5–6, lateroventral view. (E) Segment 2, laterodorsal view. (F) Detail of segment 8, lateral view. (G) Detail of segment 5, lateroventral view. (H) Segments 6–7, middorsal view. (I) Detail of sieve plate. (J) Female, segments 10–11, dorsal view. (K) Segments 9–11, ventral view. (L) Male, detail of penile spines, lateral view. (M) Male, detail of penile spines and laterodorsal tube, laterodorsal view. Abbreviations: ldgo2, laterodorsal glandular cell outlet type 2; ldss, laterodorsal sensory spot; ldt, laterodorsal tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvgco1, lateroventral glandular cell outlet type 1; lvt, lateroventral tube; mds, middorsal spine; mdss, middorsal sensory spot; mlgco2, midlateral glandular cell outlet type 2; mlss, midlateral sensory spot; mlt, midlateral tube; pe, penile spine; sdgco2, subdorsal glandular cell outlet type 2; sdss, subdorsal sensory spot; sdt, subdorsal tube; si, sieve plate; spf, secondary pectinate fringe; te, tergal extension; vlgco2, ventrolateral glandular cell outlet type 2; vlss, ventrolateral sensory spot; vmgco1, ventromedial glandular cell outlet type 1; vmss, ventromedial sensory spot. Circles indicate the position of glandular cell outlets type 1. Dashed circles indicate the position of sensory spots. Digits after abbreviations refer to segment number." figureDoi="http://doi.org/10.5281/zenodo.8164177" httpUri="https://zenodo.org/record/8164177/files/figure.png" pageId="5" pageNumber="72">4</figureCitation>
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J-K).
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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