treatments-xml/data/CF/46/87/CF46879EFFB5FFB0FF55719DFB012835.xml

<document id="15058F566A817B27B96C7FBF8E606074" ID-CLB-Dataset="82886" ID-DOI="10.1080/00222933.2021.2017499" ID-GBIF-Dataset="5cd01cfe-52bb-4092-97b2-804c3c7d145d" ID-ISSN="1464-5262" ID-Zenodo-Dep="6758313" ID-ZooBank="D766FC5E-CEC8-4184-BE90-C25350B90612" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1656311158681" checkinUser="plazi" docAuthor="Costa, Leidiane O., Duarte, Paulo R. M., Iannuzzi, Luciana &amp; Grossi, Paschoal C." docDate="2022" docId="CF46879EFFB5FFB0FF55719DFB012835" docLanguage="en" docName="JNATHIST.56.1-4.191-225.pdf" docOrigin="Journal of Natural History 56 (1 - 4)" docSource="http://dx.doi.org/10.1080/00222933.2021.2017499" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Oryctini Mulsant 1842" docType="key" docVersion="9" lastPageNumber="222" masterDocId="337FFFE6FFAAFF90FFF57271FF852A10" masterDocTitle="Taxonomic revision and notes on natural history of the enigmatic beetle genus Gibboryctes Endrödi (Coleoptera: Melolonthidae: Dynastinae)" masterLastPageNumber="225" masterPageNumber="191" pageNumber="221" updateTime="1732481095859" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9402DFDE206B2FFE8BB3A1C6BE42808D">Taxonomic revision and notes on natural history of the enigmatic beetle genus Gibboryctes Endrödi (Coleoptera: Melolonthidae: Dynastinae)</mods:title>
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<mods:namePart id="37CC84AAE5A118987A9E86BA63C105D6">Costa, Leidiane O.</mods:namePart>
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<mods:namePart id="99928B8086C02E503BFF9913A0882FEA">Duarte, Paulo R. M.</mods:namePart>
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<mods:namePart id="53FFF798FB7A6CC9856CDFC55AB2ADAB">Iannuzzi, Luciana</mods:namePart>
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<mods:namePart id="DA0E203184DA04363CB03376B8D8C7F2">Grossi, Paschoal C.</mods:namePart>
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<mods:title id="F49C53BBE229E999FC7CAA4F830C7B64">Journal of Natural History</mods:title>
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<mods:date id="2DB602666A2DFDCE9623A8A4F649343E">2022</mods:date>
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<treatment id="CF46879EFFB5FFB0FF55719DFB012835" ID-DOI="http://doi.org/10.5281/zenodo.6761938" ID-GBIF-Taxon="196400119" ID-Zenodo-Dep="6761938" LSID="urn:lsid:plazi:treatment:CF46879EFFB5FFB0FF55719DFB012835" httpUri="http://treatment.plazi.org/id/CF46879EFFB5FFB0FF55719DFB012835" lastPageId="32" lastPageNumber="222" pageId="31" pageNumber="221" scope_genus="Gibboryctes" scope_order="Coleoptera" scope_subFamily="Dynastinae">
<subSubSection id="0FF56503FFB5FF8FFF55719DFC442E15" box="[160,961,1004,1029]" pageId="31" pageNumber="221" type="nomenclature">
<paragraph id="47503688FFB5FF8FFF55719DFC442E15" blockId="31.[160,961,1004,1029]" box="[160,961,1004,1029]" pageId="31" pageNumber="221">
<emphasis id="759BEA9AFFB5FF8FFF55719DFC442E15" bold="true" box="[160,961,1004,1029]" pageId="31" pageNumber="221">
Identification key to the genera of adult 
<taxonomicName id="80EF4D0BFFB5FF8FFD7F719DFD6C2E15" ID-CoL="8RYTF" authorityName="Mulsant" authorityYear="1842" box="[650,745,1004,1029]" class="Insecta" family="Scarabaeidae" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="221" phylum="Arthropoda" rank="tribe" tribe="Oryctini">Oryctini</taxonomicName>
of South America
</emphasis>
</paragraph>
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<subSubSection id="0FF56503FFB5FFB0FF4A765EFB012835" lastPageId="32" lastPageNumber="222" pageId="31" pageNumber="221" type="key">
<key id="337E61B9FFB5FFB0FF4A765EFB012835" lastPageId="32" lastPageNumber="222" pageId="31" pageNumber="221">
<keyStep id="FC1B2E2DFFB5FF8FFF4A765EFB062EA0" pageId="31" pageNumber="221">
<paragraph id="47503688FFB5FF8FFF4A765EFB062E7B" blockId="31.[191,1156,1071,1720]" pageId="31" pageNumber="221">
<keyLead id="FC1E95BDFFB5FF8FFF4A765EFB062E7B" pageId="31" pageNumber="221">1. Body convex dorsally in lateral view. Ventral area of metatarsomeres with a longitudinal row of bristles ......................................................................................................... 2</keyLead>
</paragraph>
<paragraph id="47503688FFB5FF8FFF4A7605FB062EA0" blockId="31.[191,1156,1071,1720]" pageId="31" pageNumber="221">
<keyLead id="FC1E95BDFFB5FF8FFF4A7605FB062EA0" pageId="31" pageNumber="221">
- Body flattened dorsally in lateral view. Ventral area of metatarsomeres bearing a brush of bristles........................................................................................ 
<taxonomicName id="80EF4D0BFFB5FF8FFC6976E6FB062EA0" authority="Ohaus" authorityName="Ohaus" authorityYear="1910" box="[924,1155,1175,1200]" class="Insecta" family="Dynastidae" genus="Platyphileurus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="221" phylum="Arthropoda" rank="genus">
<emphasis id="759BEA9AFFB5FF8FFC6976E6FBB02EA0" box="[924,1077,1175,1200]" italics="true" pageId="31" pageNumber="221">Platyphileurus</emphasis>
Ohaus
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</keyLead>
</paragraph>
</keyStep>
<keyStep id="FC1B2E2DFFB5FF8FFF4A76CBFB062EE5" pageId="31" pageNumber="221">
<paragraph id="47503688FFB5FF8FFF4A76CBFB062EC3" blockId="31.[191,1156,1071,1720]" box="[191,1155,1210,1235]" pageId="31" pageNumber="221">
<keyLead id="FC1E95BDFFB5FF8FFF4A76CBFB062EC3" box="[191,1155,1210,1235]" pageId="31" pageNumber="221">2. Protibia tridentate ............................................................................................................................... 3</keyLead>
</paragraph>
<paragraph id="47503688FFB5FF8FFF4A76ADFB062EE5" blockId="31.[191,1156,1071,1720]" box="[191,1155,1244,1269]" pageId="31" pageNumber="221">
<keyLead id="FC1E95BDFFB5FF8FFF4A76ADFB062EE5" box="[191,1155,1244,1269]" pageId="31" pageNumber="221">- Protibia quadridentate....................................................................................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="FC1B2E2DFFB5FF8FFF4A768EFB012F2B" pageId="31" pageNumber="221">
<paragraph id="47503688FFB5FF8FFF4A768EFB062F08" blockId="31.[191,1156,1071,1720]" box="[191,1155,1279,1304]" pageId="31" pageNumber="221">
<keyLead id="FC1E95BDFFB5FF8FFF4A768EFB062F08" box="[191,1155,1279,1304]" pageId="31" pageNumber="221">3. Colour black. Elytra nearly smooth, except for a deep sutural stria ............................... 4</keyLead>
</paragraph>
<paragraph id="47503688FFB5FF8FFF4A7753FB012F2B" blockId="31.[191,1156,1071,1720]" box="[191,1156,1314,1339]" pageId="31" pageNumber="221">
<keyLead id="FC1E95BDFFB5FF8FFF4A7753FB012F2B" box="[191,1156,1314,1339]" pageId="31" pageNumber="221">
- Colour reddish brown. Elytra with distinct rows of punctures................ 
<taxonomicName id="80EF4D0BFFB5FF8FFC1D7753FB012F2B" authority="Hope" authorityName="Hope" authorityYear="1837" box="[1000,1156,1314,1339]" class="Insecta" family="Dynastidae" genus="Coelosis" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="221" phylum="Arthropoda" rank="genus">
<emphasis id="759BEA9AFFB5FF8FFC1D7753FBBA2F2B" box="[1000,1087,1314,1339]" italics="true" pageId="31" pageNumber="221">Coelosis</emphasis>
Hope
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</paragraph>
</keyStep>
<keyStep id="FC1B2E2DFFB5FF8FFF4A7735FB062F90" pageId="31" pageNumber="221">
<paragraph id="47503688FFB5FF8FFF4A7735FB012F4D" blockId="31.[191,1156,1071,1720]" box="[191,1156,1348,1373]" pageId="31" pageNumber="221">
<keyLead id="FC1E95BDFFB5FF8FFF4A7735FB012F4D" box="[191,1156,1348,1373]" pageId="31" pageNumber="221">
4. Mandibles bidentate............................................................................................ 
<taxonomicName id="80EF4D0BFFB5FF8FFC3E7735FB012F4D" authority="Hope" authorityName="Hope" authorityYear="1837" box="[971,1156,1348,1373]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="221" phylum="Arthropoda" rank="genus">
<emphasis id="759BEA9AFFB5FF8FFC3E7735FBBA2F4D" box="[971,1087,1348,1373]" italics="true" pageId="31" pageNumber="221">Megaceras</emphasis>
Hope
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</keyLead>
</paragraph>
<paragraph id="47503688FFB5FF8FFF4A7716FB062F90" blockId="31.[191,1156,1071,1720]" box="[191,1155,1383,1408]" pageId="31" pageNumber="221">
<keyLead id="FC1E95BDFFB5FF8FFF4A7716FB062F90" box="[191,1155,1383,1408]" pageId="31" pageNumber="221">
- Mandibles tridentate.......................................................................... 
<taxonomicName id="80EF4D0BFFB5FF8FFC927716FB062F90" authority="Dechambre" authorityName="Dechambre" authorityYear="1976" box="[871,1155,1383,1408]" class="Insecta" family="Dynastidae" genus="Megaceropsis" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="221" phylum="Arthropoda" rank="genus">
<emphasis id="759BEA9AFFB5FF8FFC927716FC7F2F90" box="[871,1018,1383,1408]" italics="true" pageId="31" pageNumber="221">Megaceropsis</emphasis>
Dechambre
</taxonomicName>
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<paragraph id="47503688FFB5FF8FFF4A77FBFB062FD5" blockId="31.[191,1156,1071,1720]" pageId="31" pageNumber="221">
<keyLead id="FC1E95BDFFB5FF8FFF4A77FBFB062FD5" pageId="31" pageNumber="221">5. Male head bearing a tubercle, never with a horn. Female pronotum with an anterior tubercle or concavity, or both combined .................................................................................. 6</keyLead>
</paragraph>
<paragraph id="47503688FFB5FF8FFF4A77BEFB062C1B" blockId="31.[191,1156,1071,1720]" pageId="31" pageNumber="221">
<keyLead id="FC1E95BDFFB5FF8FFF4A77BEFB062C1B" pageId="31" pageNumber="221">- Male head bearing a prominent horn. Female pronotum simple, without anterior tubercle or concavity.......................................................................................................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="FC1B2E2DFFB5FF8FFF4A7465FB062C40" pageId="31" pageNumber="221">
<paragraph id="47503688FFB5FF8FFF4A7465FB012C3D" blockId="31.[191,1156,1071,1720]" box="[191,1156,1556,1581]" pageId="31" pageNumber="221">
<keyLead id="FC1E95BDFFB5FF8FFF4A7465FB012C3D" box="[191,1156,1556,1581]" pageId="31" pageNumber="221">
6. Tergite VII bearing stridulatory apparatus.......................... .......................... 
<taxonomicName id="80EF4D0BFFB5FF8FFC227465FB012C3D" authority="Kirby." authorityName="Kirby." box="[983,1156,1556,1581]" class="Insecta" family="Dynastidae" genus="Strategus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="221" phylum="Arthropoda" rank="genus">
<emphasis id="759BEA9AFFB5FF8FFC227465FBB82C3D" box="[983,1085,1556,1581]" italics="true" pageId="31" pageNumber="221">Strategus</emphasis>
Kirby.
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="47503688FFB5FF8FFF4A7446FB062C40" blockId="31.[191,1156,1071,1720]" box="[191,1155,1591,1616]" pageId="31" pageNumber="221">
<keyLead id="FC1E95BDFFB5FF8FFF4A7446FB062C40" box="[191,1155,1591,1616]" pageId="31" pageNumber="221">- Tergite VII without stridulatory apparatus................................................................................. 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="FC1B2E2DFFB5FFB0FF4A742BFB012AFD" lastPageId="32" lastPageNumber="222" pageId="31" pageNumber="221">
<paragraph id="47503688FFB5FF8FFF4A742BFB012CA8" blockId="31.[191,1156,1071,1720]" pageId="31" pageNumber="221">
<keyLead id="FC1E95BDFFB5FF8FFF4A742BFB012CA8" pageId="31" pageNumber="221">
7. Teeth small and triangular at galea. Pronotum similar in both sexes, without horn or deep concavity. Female abdomen bearing the ventrite V shorter than ventrite VI...... .............................................................................................................................. 
<taxonomicName id="80EF4D0BFFB5FF8FFC5574EEFB012CA8" authority="Endrodi." authorityName="Endrodi." box="[928,1156,1695,1720]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="221" phylum="Arthropoda" rank="genus">
<emphasis id="759BEA9AFFB5FF8FFC5574EEFB9A2CA8" box="[928,1055,1695,1720]" italics="true" pageId="31" pageNumber="221">Gibboryctes</emphasis>
Endrödi.
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="47503688FF8AFFB0FF4A72FEFB012AFD" blockId="32.[191,1156,143,549]" pageId="32" pageNumber="222">
<keyLead id="FC1E95BDFF8AFFB0FF4A72FEFB012AFD" pageId="32" pageNumber="222">
- Teeth strong and scythe-shaped at galea. Male pronotum with an anterior horn. Pronotal concavity deep in both sexes. Female abdomen bearing the ventrite V longer than ventrite VI.................... .................... 
<taxonomicName id="80EF4D0BFF8AFFB0FD1572A5FB942AFD" ID-CoL="999KW" authority="Duarte and Grossi" authorityName="Duarte and Grossi" authorityYear="2022" box="[736,1041,212,237]" class="Insecta" family="Scarabaeidae" genus="Tetragus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="222" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="759BEA9AFF8AFFB0FD1572A5FCB82AFD" box="[736,829,212,237]" italics="true" pageId="32" pageNumber="222">Tetragus</emphasis>
Duarte and Grossi
</taxonomicName>
, 
<taxonomicNameLabel id="6EA857E1FF8AFFB0FBEE72A5FB012AFD" box="[1051,1156,212,237]" pageId="32" pageNumber="222" rank="genus">gen. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
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<keyStep id="FC1B2E2DFF8AFFB0FF4A7286FB012B68" pageId="32" pageNumber="222">
<paragraph id="47503688FF8AFFB0FF4A7286FB062B23" blockId="32.[191,1156,143,549]" pageId="32" pageNumber="222">
<keyLead id="FC1E95BDFF8AFFB0FF4A7286FB062B23" pageId="32" pageNumber="222">8. Prosternal process conspicuous, produced between coxae. Female head with 1–2 tubercles, never with a prominent horn ................................................................................... 9</keyLead>
</paragraph>
<paragraph id="47503688FF8AFFB0FF4A734DFB012B68" blockId="32.[191,1156,143,549]" pageId="32" pageNumber="222">
<keyLead id="FC1E95BDFF8AFFB0FF4A734DFB012B68" pageId="32" pageNumber="222">
- Prosternal process quite reduced, nearly obsolete, confined below coxae. Female head with a prominent horn......................................... ......................................... 
<taxonomicName id="80EF4D0BFF8AFFB0FC1A732EFB012B68" authority="Hope." authorityName="Hope." box="[1007,1156,351,376]" class="Insecta" family="Dynastidae" genus="Enema" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="222" phylum="Arthropoda" rank="genus">
<emphasis id="759BEA9AFF8AFFB0FC1A732EFBBC2B68" box="[1007,1081,351,376]" italics="true" pageId="32" pageNumber="222">Enema</emphasis>
Hope.
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="FC1B2E2DFF8AFFB0FF4A73F3FB012835" pageId="32" pageNumber="222">
<paragraph id="47503688FF8AFFB0FF4A73F3FB012BAD" blockId="32.[191,1156,143,549]" pageId="32" pageNumber="222">
<keyLead id="FC1E95BDFF8AFFB0FF4A73F3FB012BAD" pageId="32" pageNumber="222">
9. Body elongated in overall aspect, subparallel laterally. Anterior margin of clypeus with 2 teeth separated by a wide emargination............ ............ 
<taxonomicName id="80EF4D0BFF8AFFB0FC7173D5FB012BAD" authority="Burmeister." authorityName="Burmeister." box="[900,1156,420,445]" class="Insecta" family="Dynastidae" genus="Podischnus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="222" phylum="Arthropoda" rank="genus">
<emphasis id="759BEA9AFF8AFFB0FC7173D5FC782BAD" box="[900,1021,420,445]" italics="true" pageId="32" pageNumber="222">Podischnus</emphasis>
Burmeister.
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="47503688FF8AFFB0FF4A73B6FB012835" blockId="32.[191,1156,143,549]" pageId="32" pageNumber="222">
<keyLead id="FC1E95BDFF8AFFB0FF4A73B6FB012835" pageId="32" pageNumber="222">
- Body broader in overall aspect, rounded laterally. Anterior margin of clypeus convergent, rounded, or with 2 teeth separated by a small notch............................................. ................................................................................................................ 
<taxonomicName id="80EF4D0BFF8AFFB0FCA5707DFB012835" authority="Burmeister." authorityName="Burmeister." box="[848,1156,524,549]" class="Insecta" family="Dynastidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="222" phylum="Arthropoda" rank="genus">
<emphasis id="759BEA9AFF8AFFB0FCA5707DFC782835" box="[848,1021,524,549]" italics="true" pageId="32" pageNumber="222">Heterogomphus</emphasis>
Burmeister.
</taxonomicName>
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</paragraph>
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