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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1024.56927" ID-GBIF-Dataset="7a0ded71-ddb1-49e5-9c2c-10c01fd03a1f" ID-PMC="PMC7990858" ID-Pensoft-Pub="1313-2970-1024-157" ID-Pensoft-UUID="086BAB631B8D5B888571137F0CEFF686" ID-PubMed="33786006" ID-ZooBank="FDD37EA991214385B67D51AD313CB49E" ModsDocID="1313-2970-1024-157" checkinTime="1615979483707" checkinUser="pensoft" docAuthor="Nestor-Arriola, Jorge Ismael, Toledo-Hernandez, Victor Hugo, Solis, Angel, Gonzalez, Guillermo &amp; Vetrovec, Jaroslav" docDate="2021" docId="81015BD6C5715B37AFA30DBAEF4ED794" docLanguage="en" docName="ZooKeys 1024: 157-196" docOrigin="ZooKeys 1024" docPubDate="2021-03-16" docSource="http://dx.doi.org/10.3897/zookeys.1024.56927" docTitle="Brachiacantha bistripustulata" docType="treatment" docVersion="5" id="086BAB631B8D5B888571137F0CEFF686" lastPageNumber="157" masterDocId="086BAB631B8D5B888571137F0CEFF686" masterDocTitle="The Brachiacantha Dejean, 1837 (Coleoptera, Coccinellidae) of Central America" masterLastPageNumber="196" masterPageNumber="157" pageNumber="157" updateTime="1668149905347" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The Brachiacantha Dejean, 1837 (Coleoptera, Coccinellidae) of Central America</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nestor-Arriola, Jorge Ismael</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2394-1586</mods:nameIdentifier>
<mods:affiliation>Centro de Investigacion en Biodiversidad y Conservacion (CIByC), Universidad Autonoma del Estado de Morelos. Av. Universidad # 1001, Col. Chamilpa, Cuernavaca, Morelos, C. P. 62209, Mexico</mods:affiliation>
<mods:nameIdentifier type="email">jorge.nestorarr@uaem.edu.mx</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Toledo-Hernandez, Victor Hugo</mods:namePart>
<mods:affiliation>Centro de Investigacion en Biodiversidad y Conservacion (CIByC), Universidad Autonoma del Estado de Morelos. Av. Universidad # 1001, Col. Chamilpa, Cuernavaca, Morelos, C. P. 62209, Mexico</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Solis, Angel</mods:namePart>
<mods:affiliation>Departamento de Historia Natural, Museo Nacional de Costa Rica. Santo Domingo de Heredia, Costa Rica</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gonzalez, Guillermo</mods:namePart>
<mods:affiliation>La Reina, Santiago, Chile</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vetrovec, Jaroslav</mods:namePart>
<mods:affiliation>Buzulucka, Hradec Kralove, Ceska Republika</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-03-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1024</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>157</mods:start>
<mods:end>196</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1024.56927</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1024.56927</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1024-157</mods:identifier>
<mods:identifier type="ZooBank">FDD37EA991214385B67D51AD313CB49E</mods:identifier>
<mods:identifier type="Pensoft-UUID">086BAB631B8D5B888571137F0CEFF686</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="180585853" LSID="urn:lsid:plazi:treatment:81015BD6C5715B37AFA30DBAEF4ED794" httpUri="http://treatment.plazi.org/id/81015BD6C5715B37AFA30DBAEF4ED794" lastPageNumber="157" pageId="0" pageNumber="157">
<subSubSection pageId="0" pageNumber="157" type="nomenclature">
<paragraph pageId="0" pageNumber="157">
<taxonomicName LSID="81015BD6-C571-5B37-AFA3-0DBAEF4ED794" authority="(Fabricius)" baseAuthorityName="Fabricius" class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachiacantha bistripustulata" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="bistripustulata">Brachiacantha bistripustulata (Fabricius)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="157" type="reference_group">
<paragraph pageId="0" pageNumber="157">
<taxonomicName authorityName="Thunberg" authorityYear="1781" class="Insecta" family="Coccinellidae" genus="Coccinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccinella laevis" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="laevis">Coccinella laevis</taxonomicName>
Thunberg, 1781: 20.
</paragraph>
<paragraph pageId="0" pageNumber="157">
<taxonomicName class="Insecta" family="Coccinellidae" genus="Coccinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccinella bistripustulata" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="bistripustulata">Coccinella bistripustulata</taxonomicName>
Fabricius, 1801: 383.
</paragraph>
<paragraph pageId="0" pageNumber="157">
<taxonomicName class="Insecta" family="Coccinellidae" genus="Brachyacantha" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Brachyacantha bis-tripustulata" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="bis-tripustulata">Brachyacantha bis-tripustulata</taxonomicName>
:
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.8953" author="Mulsant, ME" journalOrPublisher="Annales des Sciences Physiques et Naturelles, d'agriculture et d'industrie Lyon" pageId="0" pageNumber="157" pagination="1 - 1104" refId="B22" refString="Mulsant, ME, 1850. Species de Coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles, d'agriculture et d'industrie Lyon 2: 1 - 1104, DOI: https://doi.org/10.5962/bhl.title.8953" title="Species de Coleopteres trimeres securipalpes." url="https://doi.org/10.5962/bhl.title.8953" volume="2" year="1850">Mulsant 1850</bibRefCitation>
: 528.
</paragraph>
<paragraph pageId="0" pageNumber="157">
<taxonomicName class="Insecta" family="Coccinellidae" genus="Brachyacantha" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Brachyacantha bistripustulata" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="bistripustulata">Brachyacantha bistripustulata</taxonomicName>
:
<bibRefCitation author="Gorham, HS" journalOrPublisher="Biologia Centrali-Americana, Insecta, Coleoptera" pageId="0" pageNumber="157" pagination="177 - 208" refId="B17" refString="Gorham, HS, 1894. Coccinellidae. Biologia Centrali-Americana, Insecta, Coleoptera 7: 177 - 208" title="Coccinellidae." volume="7" year="1894">Gorham 1894</bibRefCitation>
: 188.
<bibRefCitation author="Casey, TL" journalOrPublisher="Journal of the New York Entomological Society" pageId="0" pageNumber="157" pagination="7 - 169" refId="B5" refString="Casey, TL, 1899. A revision of the American Coccinellidae. Journal of the New York Entomological Society 7: 7 - 169" title="A revision of the American Coccinellidae." volume="7" year="1899">Casey 1899</bibRefCitation>
: 119.
<bibRefCitation author="Leng, CW" journalOrPublisher="Bulletin American Museum of Natural History" pageId="0" pageNumber="157" pagination="279 - 333" refId="B19" refString="Leng, CW, 1911. The species of Brachyacantha of North and South America. Bulletin American Museum of Natural History 30: 279 - 333" title="The species of Brachyacantha of North and South America." volume="30" year="1911">Leng 1911</bibRefCitation>
: 296.
<bibRefCitation author="Korschefsky, R" journalOrPublisher="Junk, Berlin" pageId="0" pageNumber="157" refId="B18" refString="Korschefsky, R, 1931. Coccinellidae I. Coleopterorum Catalogus. Part 118. W. Junk, Berlin" title="Coccinellidae I. Coleopterorum Catalogus. Part 118. W." year="1931">Korschefsky 1931</bibRefCitation>
: 203.
<bibRefCitation DOI="https://doi.org/10.5479/si.03629236.185.3" author="Blackwelder, RE" journalOrPublisher="Bulletin of the United States National Museum" pageId="0" pageNumber="157" pagination="343 - 550" refId="B1" refString="Blackwelder, RE, 1945. Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies and South America. Part 3. Bulletin of the United States National Museum 185: 343 - 550, DOI: https://doi.org/10.5479/si.03629236.185.3" title="Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies and South America. Part 3." url="https://doi.org/10.5479/si.03629236.185.3" volume="185" year="1945">Blackwelder 1945</bibRefCitation>
: 449.
</paragraph>
<paragraph pageId="0" pageNumber="157">
<taxonomicName class="Insecta" family="Coccinellidae" genus="Brachyacantha" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Brachyacantha erythrocephala" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="erythrocephala">Brachyacantha erythrocephala</taxonomicName>
Gorham, 1894: 188 (non
<taxonomicName class="Insecta" family="Coccinellidae" genus="Coccinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccinella erythrocephala" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="erythrocephala">Coccinella erythrocephala</taxonomicName>
Fabricius).
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.8975" author="Crotch, GR" journalOrPublisher="Janson EW, London" pageId="0" pageNumber="157" refId="B9" refString="Crotch, GR, 1874. A Revision of the Coleopterous Family Coccinellidae. Janson EW, London, DOI: https://doi.org/10.5962/bhl.title.8975" title="A Revision of the Coleopterous Family Coccinellidae." url="https://doi.org/10.5962/bhl.title.8975" year="1874">Crotch 1874</bibRefCitation>
: 211.
</paragraph>
<paragraph pageId="0" pageNumber="157">
<taxonomicName class="Insecta" family="Coccinellidae" genus="Brachyacantha" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Brachyacantha decora" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="decora">Brachyacantha decora</taxonomicName>
Casey, 1899: 119.
<bibRefCitation author="Bowditch, FC" journalOrPublisher="Entomological News" pageId="0" pageNumber="157" pagination="205 - 207" refId="B3" refString="Bowditch, FC, 1902. Notes on Casey's revision of the American Coccinellidae. Entomological News 13: 205 - 207" title="Notes on Casey's revision of the American Coccinellidae." volume="13" year="1902">Bowditch 1902</bibRefCitation>
: 206.
</paragraph>
<paragraph pageId="0" pageNumber="157">
<taxonomicName class="Insecta" family="Coccinellidae" genus="Brachyacantha" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Brachyacantha bistripustulata subsp. var. var. guttata" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="variety" species="bistripustulata" subSpecies="var." variety="guttata">Brachyacantha bistripustulata var. guttata</taxonomicName>
Weise, 1885: 231.
<bibRefCitation author="Leng, CW" journalOrPublisher="Bulletin American Museum of Natural History" pageId="0" pageNumber="157" pagination="279 - 333" refId="B19" refString="Leng, CW, 1911. The species of Brachyacantha of North and South America. Bulletin American Museum of Natural History 30: 279 - 333" title="The species of Brachyacantha of North and South America." volume="30" year="1911">Leng 1911</bibRefCitation>
: 297.
<bibRefCitation author="Korschefsky, R" journalOrPublisher="Junk, Berlin" pageId="0" pageNumber="157" refId="B18" refString="Korschefsky, R, 1931. Coccinellidae I. Coleopterorum Catalogus. Part 118. W. Junk, Berlin" title="Coccinellidae I. Coleopterorum Catalogus. Part 118. W." year="1931">Korschefsky 1931</bibRefCitation>
: 203.
</paragraph>
<paragraph pageId="0" pageNumber="157">
<taxonomicName class="Insecta" family="Coccinellidae" genus="Brachyacantha" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Brachyacantha bistripustulata subsp. var. var. decora" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="variety" species="bistripustulata" subSpecies="var." variety="decora">Brachyacantha bistripustulata var. decora</taxonomicName>
: Leng, 1911: 298.
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.8777" author="Leng, CW" journalOrPublisher="Mount Vernon, New York" pageId="0" pageNumber="157" refId="B20" refString="Leng, CW, 1920. Catalogue of the Coleoptera of America, north of Mexico. Mount Vernon, New York, DOI: https://doi.org/10.5962/bhl.title.8777" title="Catalogue of the Coleoptera of America, north of Mexico." url="https://doi.org/10.5962/bhl.title.8777" year="1920">Leng 1920</bibRefCitation>
: 213.
</paragraph>
<paragraph pageId="0" pageNumber="157">
<taxonomicName class="Insecta" family="Coccinellidae" genus="Brachyacantha" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Brachyacantha bistripustulata subsp. var. var. minor" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="variety" species="bistripustulata" subSpecies="var." variety="minor">Brachyacantha bistripustulata var. minor</taxonomicName>
Leng, 1911: 298.
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.8777" author="Leng, CW" journalOrPublisher="Mount Vernon, New York" pageId="0" pageNumber="157" refId="B20" refString="Leng, CW, 1920. Catalogue of the Coleoptera of America, north of Mexico. Mount Vernon, New York, DOI: https://doi.org/10.5962/bhl.title.8777" title="Catalogue of the Coleoptera of America, north of Mexico." url="https://doi.org/10.5962/bhl.title.8777" year="1920">Leng 1920</bibRefCitation>
: 213.
</paragraph>
<paragraph pageId="0" pageNumber="157">
<taxonomicName class="Insecta" family="Coccinellidae" genus="Brachyacantha" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Brachyacantha bistripustulata subsp. var. var. obscura" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="variety" species="bistripustulata" subSpecies="var." variety="obscura">Brachyacantha bistripustulata var. obscura</taxonomicName>
Leng, 1911: 299.
<bibRefCitation author="Korschefsky, R" journalOrPublisher="Junk, Berlin" pageId="0" pageNumber="157" refId="B18" refString="Korschefsky, R, 1931. Coccinellidae I. Coleopterorum Catalogus. Part 118. W. Junk, Berlin" title="Coccinellidae I. Coleopterorum Catalogus. Part 118. W." year="1931">Korschefsky 1931</bibRefCitation>
: 204.
</paragraph>
<paragraph pageId="0" pageNumber="157">
<taxonomicName class="Insecta" family="Coccinellidae" genus="Brachyacantha" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Brachyacantha bistripustulata subsp. var. var. quichiana" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="variety" species="bistripustulata" subSpecies="var." variety="quichiana">Brachyacantha bistripustulata var. quichiana</taxonomicName>
Leng, 1911: 298.
</paragraph>
<paragraph pageId="0" pageNumber="157">
<taxonomicName authority="decora" class="Insecta" family="Coccinellidae" genus="Brachyacantha" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Brachyacantha bistripustulata subsp. ab." order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="subSpecies" species="bistripustulata" subSpecies="ab.">Brachyacantha bistripustulata ab. decora</taxonomicName>
:
<bibRefCitation author="Korschefsky, R" journalOrPublisher="Junk, Berlin" pageId="0" pageNumber="157" refId="B18" refString="Korschefsky, R, 1931. Coccinellidae I. Coleopterorum Catalogus. Part 118. W. Junk, Berlin" title="Coccinellidae I. Coleopterorum Catalogus. Part 118. W." year="1931">Korschefsky 1931</bibRefCitation>
: 203.
</paragraph>
<paragraph pageId="0" pageNumber="157">
<taxonomicName authority="minor" class="Insecta" family="Coccinellidae" genus="Brachyacantha" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Brachyacantha bistripustulata subsp. ab." order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="subSpecies" species="bistripustulata" subSpecies="ab.">Brachyacantha bistripustulata ab. minor</taxonomicName>
:
<bibRefCitation author="Korschefsky, R" journalOrPublisher="Junk, Berlin" pageId="0" pageNumber="157" refId="B18" refString="Korschefsky, R, 1931. Coccinellidae I. Coleopterorum Catalogus. Part 118. W. Junk, Berlin" title="Coccinellidae I. Coleopterorum Catalogus. Part 118. W." year="1931">Korschefsky 1931</bibRefCitation>
: 203.
</paragraph>
<paragraph pageId="0" pageNumber="157">
<taxonomicName authorityName="Casey" authorityYear="1899" class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachiacantha decora" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="decora">Brachiacantha decora</taxonomicName>
:
<bibRefCitation author="Gordon, RD" journalOrPublisher="Journal of the New York Entomological Society" pageId="0" pageNumber="157" pagination="1 - 912" refId="B14" refString="Gordon, RD, 1985. The Coccinellidae (Coleoptera) of America North of Mexico. Journal of the New York Entomological Society 93 (1): 1 - 912" title="The Coccinellidae (Coleoptera) of America North of Mexico." volume="93" year="1985">Gordon 1985</bibRefCitation>
: 561.
</paragraph>
<paragraph pageId="0" pageNumber="157">
<taxonomicName class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachiacantha laevis" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="laevis">Brachiacantha laevis</taxonomicName>
:
<bibRefCitation DOI="https://doi.org/10.1163/187631287X00025" author="Pope, RD" journalOrPublisher="Baudouin, Paris" pageId="0" pageNumber="157" refId="B28" refString="Pope, RD, 1987. The Coccinellidae (Coleoptera) described by C. P. Thunberg. Insect Systematics and Evolution 18(1): 51-66. https://doi.org/10.1163/187631287X00025" title="The Coccinellidae (Coleoptera) described by C. P. Thunberg. Insect Systematics and Evolution 18 (1): 51 - 66." url="https://doi.org/10.1163/187631287X00025" year="1987">Pope 1987</bibRefCitation>
: 58.
</paragraph>
<paragraph pageId="0" pageNumber="157">
<taxonomicName class="Insecta" family="Coccinellidae" genus="Brachiacantha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachiacantha bistripustulata" order="Coleoptera" pageId="0" pageNumber="157" phylum="Arthropoda" rank="species" species="bistripustulata">Brachiacantha bistripustulata</taxonomicName>
:
<bibRefCitation DOI="https://doi.org/10.1590/S0073-47212007000400010" author="Milleo, J" journalOrPublisher="Iheringia, Serie Zoologia, Porto Alegre" pageId="0" pageNumber="157" pagination="418 - 424" refId="B21" refString="Milleo, J, de Almeida, LM, 2007. Notas taxonomicas sobre os Brachiacanthini neotropicais (Coleoptera, Coccinellidae, Hyperaspinae). Iheringia, Serie Zoologia, Porto Alegre 97 (4): 418 - 424, DOI: https://doi.org/10.1590/S0073-47212007000400010" title="Notas taxonomicas sobre os Brachiacanthini neotropicais (Coleoptera, Coccinellidae, Hyperaspinae)." url="https://doi.org/10.1590/S0073-47212007000400010" volume="97" year="2007">
<normalizedToken originalValue="Milléo">Milleo</normalizedToken>
and Almeida 2007
</bibRefCitation>
: 420.
<bibRefCitation author="Gordon, RD" journalOrPublisher="Insecta Mundi" pageId="0" pageNumber="157" pagination="1 - 76" refId="B16" refString="Gordon, RD, Canepari, C, Hanley, G, 2014. South American Coccinellidae (Coleoptera) Part XVI: systematic revision of Brachiacantha Dejean (Coccinellinae: Hyperaspidini). Insecta Mundi 390: 1 - 76" title="South American Coccinellidae (Coleoptera) Part XVI: systematic revision of Brachiacantha Dejean (Coccinellinae: Hyperaspidini)." volume="390" year="2014">Gordon et al. 2014</bibRefCitation>
: 10.
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4701.6.3" author="Nestor-Arriola, JI" journalOrPublisher="Zootaxa" pageId="0" pageNumber="157" pagination="537 - 552" refId="B25" refString="Nestor-Arriola, JI, Toledo-Hernandez, VH, 2019. Contribution to the knowledge of the dentipes group of the genus Brachiacantha (Coleoptera: Coccinellidae). Zootaxa 4701 (6): 537 - 552, DOI: https://doi.org/10.11646/zootaxa.4701.6.3" title="Contribution to the knowledge of the dentipes group of the genus Brachiacantha (Coleoptera: Coccinellidae)." url="https://doi.org/10.11646/zootaxa.4701.6.3" volume="4701" year="2019">
Nestor-Arriola and
<normalizedToken originalValue="Toledo-Hernández">Toledo-Hernandez</normalizedToken>
2019
</bibRefCitation>
: 539.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="157" type="materials_examined">
<paragraph pageId="0" pageNumber="157">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="157">
<materialsCitation ID-GBIF-Occurrence="3063616354" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="CEAM, CNIN, FSCA, USNMNH, OUNMH, MNCR, MUCR, MZCR" country="Costa Rica" county="Frontera" location="Oeste" municipality="Tegucigalpa" specimenCount="496" stateProvince="Cocle">
<specimenCount type="generic">496 specimens</specimenCount>
from
<collectingCountry name="Costa Rica">Costa Rica</collectingCountry>
: Alajuela, Cartago, Heredia and Puntarenas;
<collectingCountry name="El Salvador">El Salvador</collectingCountry>
:
<normalizedToken originalValue="Ahuachapán">Ahuachapan</normalizedToken>
, La Libertad and San Salvador;
<collectingCountry name="Guatemala">Guatemala</collectingCountry>
: El
<normalizedToken originalValue="Quiché">Quiche</normalizedToken>
, El Progreso,
<collectingCounty>Frontera</collectingCounty>
, Jutiapa, Chimaltenango, Baja Verapaz, Santa Rosa and Zacapa;
<collectingCountry name="Honduras">Honduras</collectingCountry>
: El
<normalizedToken originalValue="Paraíso">Paraiso</normalizedToken>
,
<collectingMunicipality>Tegucigalpa</collectingMunicipality>
,
<normalizedToken originalValue="Atlántida">Atlantida</normalizedToken>
,
<normalizedToken originalValue="Colón">Colon</normalizedToken>
,
<normalizedToken originalValue="Cortés">Cortes</normalizedToken>
, Comayagua, Francisco
<normalizedToken originalValue="Morazán">Morazan</normalizedToken>
, La Paz, Olancho and Yoro;
<normalizedToken originalValue="Panamá">
<collectingCountry name="Panama">Panama</collectingCountry>
</normalizedToken>
:
<location LSID="urn:lsid:plazi:treatment:81015BD6C5715B37AFA30DBAEF4ED794:4DA5AE2A89C3E51B27EDC7C440E2A3F4" country="Costa Rica" county="Frontera" municipality="Tegucigalpa" name="Oeste" stateProvince="Cocle">Oeste</location>
,
<normalizedToken originalValue="Coclé">
<collectingRegion country="Panama" name="Cocle">Cocle</collectingRegion>
</normalizedToken>
,
<normalizedToken originalValue="Chiquirí">
<location LSID="urn:lsid:plazi:treatment:81015BD6C5715B37AFA30DBAEF4ED794:6892346A9083F24D4C0E1BD481C02F27" country="Costa Rica" county="Frontera" municipality="Tegucigalpa" name="Chiquiri" stateProvince="Cocle">Chiquiri</location>
</normalizedToken>
and
<collectingRegion country="Panama" name="Darien">Darien</collectingRegion>
(
<collectionCode country="Mexico" httpUri="http://grbio.org/cool/ff28-ftqi" name="Centro de Entomologica y Acarologia" type="">CEAM</collectionCode>
,
<collectionCode country="Mexico" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" type="">CNIN</collectionCode>
,
<collectionCode country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" type="Museum">FSCA</collectionCode>
,
<collectionCode>USNMNH</collectionCode>
,
<collectionCode>OUNMH</collectionCode>
,
<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" type="">MNCR</collectionCode>
,
<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/ww9x-1v7b" name="Museo de Insectos" type="">MUCR</collectionCode>
,
<collectionCode country="Costa Rica" httpUri="http://grbio.org/cool/qfj6-taa9" name="Museo de Zoologia" type="">MZCR</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="157" type="diagnosis">
<paragraph pageId="0" pageNumber="157">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="157">
Length
<quantity metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="5.4" metricValueMin="3.2" unit="mm" value="4.3" valueMax="5.4" valueMin="3.2">3.2-5.4 mm</quantity>
. Width
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.2" metricValueMax="4.0" metricValueMin="2.4" unit="mm" value="3.2" valueMax="4.0" valueMin="2.4">2.4-4 mm</quantity>
. Dorsally black with three to four orange, yellow or red spots on each elytron, spots sometimes fused. Males with pale-yellow head and pronotal anterior border. Third ventrite bicuspid in males, cusps short, wide and with divergent apex. Male genitalia asymmetrical, penis guide with apical hook.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="157" type="distribution">
<paragraph pageId="0" pageNumber="157">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="157">
From the South of the
<collectingCountry name="United States of America">USA</collectingCountry>
to
<collectingCountry name="Brazil">Brazil</collectingCountry>
and
<collectingCountry name="Peru">Peru</collectingCountry>
, including
<collectingCountry name="Jamaica">Jamaica</collectingCountry>
,
<collectingCountry name="Cuba">Cuba</collectingCountry>
,
<collectingCountry name="Puerto Rico">Puerto Rico</collectingCountry>
and
<collectingCountry name="Dominican Republic">Dominican Republic</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="157" type="discussion">
<paragraph pageId="0" pageNumber="157">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="157">This species is very similar to others in the group, but the shape of the male ventral cusps is different in being wider than long, apically divergent and not strongly pronounced, and is usually enough to differentiate it.</paragraph>
</subSubSection>
</treatment>
</document>
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