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<document id="0312E97F05B74FEC1DA27893319E89D0" ID-CLB-Dataset="29418" ID-DOI="10.11646/zootaxa.4482.3.2" ID-GBIF-Dataset="fd24ef18-76ac-46f1-aec2-2faf5b0ff325" ID-ISSN="1175-5326" ID-Zenodo-Dep="1440707" ID-ZooBank="3FBE11E2-9F97-4A29-92A9-67780C782E0D" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538375301724" checkinUser="plazi" docAuthor="Li, Jheng-Jhang, Rahayu, Dwi Listyo &amp; Ng, Peter K. L." docDate="2018" docId="376A1B4DFFB0FF91D1CDF8F24053FD72" docLanguage="en" docName="zootaxa.4482.3.2.pdf" docOrigin="Zootaxa 4482 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parasesarma purpureum Li &amp; Rahayu &amp; Ng 2018, n. sp." docType="treatment" docVersion="7" lastPageNumber="474" masterDocId="CB536335FFA4FF86D15AFFF74432FFDA" masterDocTitle="Identity of the tree-spider crab, Parasesarma leptosoma (Hilgendorf, 1869) (Decapoda: Brachyura: Sesarmidae), with descriptions of seven new species from the Western Pacific" masterLastPageNumber="490" masterPageNumber="451" pageNumber="471" updateTime="1698559507815" updateUser="ExternalLinkService">
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<mods:title id="13D99535E6C21E9645712D4658E4A7ED">Identity of the tree-spider crab, Parasesarma leptosoma (Hilgendorf, 1869) (Decapoda: Brachyura: Sesarmidae), with descriptions of seven new species from the Western Pacific</mods:title>
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<mods:namePart id="B80697B2CAB8F607EF2AE5C70F410994">Li, Jheng-Jhang</mods:namePart>
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<mods:namePart id="B76055DDE7715E845C829FDCEA625A0A">Rahayu, Dwi Listyo</mods:namePart>
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<mods:namePart id="039FD637AE9756FFFF39F6771BC22534">Ng, Peter K. L.</mods:namePart>
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<mods:title id="B47C2BFB8FEC009AE4B6ECFC9D09D56C">Zootaxa</mods:title>
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<mods:date id="28278FDD918C10952D33EC8CF2DF44D7">2018</mods:date>
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<mods:number id="7C957AC4114370194F8E4B078DA9EA0A">2018-09-19</mods:number>
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<mods:identifier id="E829662E8D15D363A9B2FAB89FEDFDDD" type="DOI">10.11646/zootaxa.4482.3.2</mods:identifier>
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<treatment id="376A1B4DFFB0FF91D1CDF8F24053FD72" ID-DOI="http://doi.org/10.5281/zenodo.5986795" ID-GBIF-Taxon="148421724" ID-Zenodo-Dep="5986795" LSID="urn:lsid:plazi:treatment:376A1B4DFFB0FF91D1CDF8F24053FD72" httpUri="http://treatment.plazi.org/id/376A1B4DFFB0FF91D1CDF8F24053FD72" lastPageId="23" lastPageNumber="474" pageId="20" pageNumber="471">
<subSubSection id="F7D9F9D0FFB0FF92D1CDF8F246BDF89A" pageId="20" pageNumber="471" type="nomenclature">
<paragraph id="BF7CAA5BFFB0FF92D1CDF8F2463DF8C5" blockId="20.[151,655,1797,1857]" box="[151,527,1797,1823]" pageId="20" pageNumber="471">
<heading id="E4341D37FFB0FF92D1CDF8F2463DF8C5" bold="true" box="[151,527,1797,1823]" fontSize="11" level="1" pageId="20" pageNumber="471" reason="1">
<emphasis id="8DB77649FFB0FF92D1CDF8F2463DF8C5" bold="true" box="[151,527,1797,1823]" pageId="20" pageNumber="471">
<taxonomicName id="78C3D1D8FFB0FF92D1CDF8F245F7F8C5" ID-CoL="84G4G" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[151,453,1797,1823]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="purpureum" status="sp. nov.">
<emphasis id="8DB77649FFB0FF92D1CDF8F245F7F8C5" bold="true" box="[151,453,1797,1823]" italics="true" pageId="20" pageNumber="471">Parasesarma purpureum</emphasis>
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<taxonomicNameLabel id="9684CB32FFB0FF92D096F8F2463DF8C5" box="[460,527,1797,1823]" pageId="20" pageNumber="471" rank="species">n. sp.</taxonomicNameLabel>
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</paragraph>
<paragraph id="BF7CAA5BFFB0FF92D1CDF8DF46BDF89A" blockId="20.[151,655,1797,1857]" box="[151,655,1832,1857]" pageId="20" pageNumber="471">
(
<figureCitation id="27F8B6DEFFB0FF92D1C5F8DF44CEF89A" box="[159,252,1832,1857]" captionStart="FIGURE 13" captionStartId="21.[151,250,1778,1800]" captionTargetBox="[152,1435,608,1724]" captionTargetId="figure@21.[152,1435,608,1724]" captionTargetPageId="21" captionText="FIGURE 13. Parasesarma purpureum n. sp.. A–D, holotype male (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; E, F, paratype female (ZRC 2012.0755b), same locality. A, E, dorsal views; B, F, ventral views; C, D, frontal views." httpUri="https://zenodo.org/record/1440735/files/figure.png" pageId="20" pageNumber="471">Figs. 13</figureCitation>
,
<figureCitation id="27F8B6DEFFB0FF92D052F8DF4514F89A" box="[264,294,1832,1856]" captionStart="FIGURE 14" captionStartId="22.[151,250,1911,1933]" captionTargetBox="[198,1357,638,1836]" captionTargetId="figure@22.[152,1435,568,1875]" captionTargetPageId="22" captionText="FIGURE 14. Parasesarma purpureum n. sp., holotype male (15.9×13.7 mm) (ZRC 2012.0755a), Tioman, Peninsular Malaysia. A, dorsal view of carapace; B–D, tip of left cheliped; E–H, left G1; B, lateral view of pectinate cristae on palm; C, dorsal view of palm and dactylar finger; D, lateral view of dactylar finger; E, G, dorsal (sternal) views; F, H, ventral (pleonal) views; E, F, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440737/files/figure.png" pageId="20" pageNumber="471">14</figureCitation>
,
<figureCitation id="27F8B6DEFFB0FF92D068F8DF456CF89A" box="[306,350,1832,1856]" captionStart="FIGURE 19" captionStartId="28.[151,250,1811,1833]" captionTargetBox="[152,1435,199,1754]" captionTargetId="figure@28.[152,1435,199,1754]" captionTargetPageId="28" captionText="FIGURE 19. Outer views of chelae, holotype males. A, Parasesarma leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; C, P. macaco n. sp. (13.8×12.3 mm) (NMNS-7779- 005), Pingtung, Taiwan; D, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440747/files/figure.png" pageId="20" pageNumber="471">19F</figureCitation>
,
<figureCitation id="27F8B6DEFFB0FF92D030F8DF45ABF89A" box="[362,409,1832,1856]" captionStart="FIGURE 20" captionStartId="30.[151,250,1268,1290]" captionTargetBox="[152,1435,393,1246]" captionTargetId="figure@30.[152,1435,393,1246]" captionTargetPageId="30" captionText="FIGURE 20. First to fourth ambulatory legs (P2–P5) (from top to bottom), holotype males. A, Parasesarma gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; B, P. macaco n. sp. male (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; E, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; F, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; G, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440749/files/figure.png" pageId="20" pageNumber="471">20E</figureCitation>
,
<figureCitation id="27F8B6DEFFB0FF92D0FFF8DF45E3F89A" box="[421,465,1832,1856]" captionStart="FIGURE 21" captionStartId="31.[151,250,1815,1837]" captionTargetBox="[152,1435,260,1731]" captionTargetId="figure@31.[152,1435,260,1731]" captionTargetPageId="31" captionText="FIGURE 21. Gonopods of Parasesarma leptosoma species-complex. A, P. leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; C, P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; D, P. kui n. sp., holotype (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp., holotype (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp., holotype (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp., holotype (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp., holotype (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440751/files/figure.png" pageId="20" pageNumber="471">21F</figureCitation>
,
<figureCitation id="27F8B6DEFFB0FF92D086F8DF463DF89A" box="[476,527,1832,1856]" captionStart="FIGURE 22" captionStartId="32.[151,250,1490,1512]" captionTargetBox="[152,1435,193,1469]" captionTargetId="figure@32.[152,1435,193,1469]" captionTargetPageId="32" captionText="FIGURE 22. Colour in life, males, dorsal views. A. Parasesarma gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; B. P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C. P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. purpureum n. sp. (13.4×11.8 mm) (ZRC 2018.0799), Borneo, Malaysia; E, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440753/files/figure.png" pageId="20" pageNumber="471">22D</figureCitation>
,
<figureCitation id="27F8B6DEFFB0FF92D340F8DF467CF89A" box="[538,590,1832,1856]" captionStart="FIGURE 23" captionStartId="33.[151,250,1605,1627]" captionTargetBox="[152,1435,245,1548]" captionTargetId="figure@33.[152,1435,245,1548]" captionTargetPageId="33" captionText="FIGURE 23. Colour in life, males, ventral views. A. Parasesarma gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; B. P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C. P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. purpureum n. sp. (13.4×11.8 mm) (ZRC 2018.0799), Borneo, Malaysia; E, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440755/files/figure.png" pageId="20" pageNumber="471">23D</figureCitation>
,
<figureCitation id="27F8B6DEFFB0FF92D303F8DF46B5F89A" box="[601,647,1832,1856]" captionStart="FIGURE 24" captionStartId="34.[151,250,1531,1553]" captionTargetBox="[151,1436,218,1490]" captionTargetId="figure@34.[151,1436,218,1490]" captionTargetPageId="34" captionText="FIGURE 24. In situ photographs of Parasesarma leptosoma species-complex, specimens not collected. A, P. leptosoma (Hilgendorf, 1869), Kenya (Photograph: S. Cannicci); B, P. gecko n. sp., Okinawa, Japan; C, P. macaco n. sp., Pingtung, Taiwan; D, P. kui n. sp., Pingtung, Taiwan; E, P. purpureum n. sp., Borneo, Malaysia; F, P. tarantula n. sp., Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440757/files/figure.png" pageId="20" pageNumber="471">24E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFB0FF93D1CDF88740AEFE2E" lastPageId="21" lastPageNumber="472" pageId="20" pageNumber="471" type="materials_examined">
<paragraph id="BF7CAA5BFFB0FF93D1CDF88740AEFE2E" blockId="20.[151,1436,1904,2037]" lastBlockId="21.[151,1436,151,501]" lastPageId="21" lastPageNumber="472" pageId="20" pageNumber="471">
<materialsCitation id="0FABA006FFB0FF92D1CDF887459FF876" ID-GBIF-Occurrence="1913889783" collectingDate="2012-07-07" collectionCode="ZRC" collectorName="B. Y. Lee" country="Malaysia" location="Tioman Island" municipality="Material" pageId="20" pageNumber="471" specimenCode="ZRC 2012.0755" specimenCount="1" typeStatus="holotype">
<emphasis id="8DB77649FFB0FF92D1CDF88745B2F853" bold="true" box="[151,384,1904,1929]" pageId="20" pageNumber="471">
<collectingMunicipality id="5F183021FFB0FF92D1CDF88744CDF853" box="[151,255,1904,1929]" pageId="20" pageNumber="471">Material</collectingMunicipality>
examined.
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<typeStatus id="607814F9FFB0FF92D0DCF887462CF852" box="[390,542,1904,1928]" pageId="20" pageNumber="471" type="holotype">HOLOTYPE</typeStatus>
: male (15.9×
<quantity id="783B07BEFFB0FF92D3F4F8874721F852" box="[686,787,1904,1929]" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" pageId="20" pageNumber="471" unit="mm" value="13.7">13.7 mm</quantity>
) (
<specimenCode id="EF650220FFB0FF92D273F88747D6F853" box="[809,996,1904,1929]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="471">ZRC 2012.0755</specimenCode>
a), Peninsular
<collectingCountry id="C7D4EACBFFB0FF92D525F88740D8F853" box="[1151,1258,1904,1929]" name="Malaysia" pageId="20" pageNumber="471">Malaysia</collectingCountry>
:
<location id="BA1CFC80FFB0FF92D5AEF88741AAF853" LSID="urn:lsid:plazi:treatment:376A1B4DFFB0FF91D1CDF8F24053FD72:BA1CFC80FFB0FF92D5AEF88741AAF853" box="[1268,1432,1904,1929]" country="Malaysia" municipality="Material" name="Tioman Island" pageId="20" pageNumber="471">Tioman Island</location>
,
<collectingDate id="DB397573FFB0FF92D1CDF862451EF877" box="[151,300,1940,1965]" pageId="20" pageNumber="471" value="2012-07-07">
<date id="CB7D8C9BFFB0FF92D1CDF862451EF877" box="[151,300,1940,1965]" pageId="20" pageNumber="471" value="2012-07-07">7 July 2012</date>
</collectingDate>
,
<collectorName id="1236CF8DFFB0FF92D065F862459FF876" box="[319,429,1941,1964]" pageId="20" pageNumber="471">B.Y. Lee</collectorName>
</materialsCitation>
. PARATYPES: 3 males (10.7×
<quantity id="783B07BEFFB0FF92D27FF86247B5F876" box="[805,903,1941,1965]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="20" pageNumber="471" unit="mm" value="9.2">9.2 mm</quantity>
, 13.9×12.0 mm, 15.5×
<quantity id="783B07BEFFB0FF92D5C7F863413FF877" box="[1181,1293,1940,1965]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="20" pageNumber="471" unit="mm" value="13.3">13.3 mm</quantity>
),
<materialsCitation id="0FABA006FFB0FF92D472F8624141F80B" ID-GBIF-Occurrence="1913889804" collectingDate="2012-07-07" collectionCode="ZRC" collectorName="B. Y. Lee" country="Malaysia" location="Tioman Island" municipality="Material" pageId="20" pageNumber="471" specimenCode="ZRC 2018.0794" specimenCount="4" specimenCount-female="4" typeStatus="holotype">
<specimenCount id="A9C561D2FFB0FF92D472F86241AEF877" box="[1320,1436,1940,1965]" pageId="20" pageNumber="471" type="female">4 females</specimenCount>
(12.6×
<quantity id="783B07BEFFB0FF92D1BAF84F4579F80A" box="[224,331,1976,2001]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="20" pageNumber="471" unit="mm" value="10.7">10.7 mm</quantity>
, 13.5×12.0 mm, 14.9×
<quantity id="783B07BEFFB0FF92D30FF84F468DF80A" box="[597,703,1976,2000]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="20" pageNumber="471" unit="mm" value="12.7">12.7 mm</quantity>
, 15.6×13.0 mm) (
<specimenCode id="EF650220FFB0FF92D2C9F84F4060F80A" box="[915,1106,1976,2000]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="471">ZRC 2018.0794</specimenCode>
), same data as holotype.
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—3 males (11.9×10.0 mm, 12.7×
<quantity id="783B07BEFFB0FF92D085F82A4675F82E" box="[479,583,2013,2037]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="20" pageNumber="471" unit="mm" value="10.8">10.8 mm</quantity>
, 14.2×
<quantity id="783B07BEFFB0FF92D3CFF82A46CCF82F" box="[661,766,2013,2037]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="20" pageNumber="471" unit="mm" value="12.8">12.8 mm</quantity>
),
<materialsCitation id="0FABA006FFB0FF93D249F82A4113FF6A" ID-GBIF-Occurrence="1913889798" collectingDate="2012-07-04" collectionCode="ZRC" collectorName="B. Y. Lee." country="Malaysia" lastPageId="21" lastPageNumber="472" location="Tioman Island" pageId="20" pageNumber="471" specimenCode="ZRC 2018.0795" specimenCount="4" specimenCount-female="4" typeStatus="holotype">
<specimenCount id="A9C561D2FFB0FF92D249F82A47B2F82F" box="[787,896,2012,2037]" pageId="20" pageNumber="471" type="female">4 females</specimenCount>
(10.9×9.0 mm, 12.1×
<quantity id="783B07BEFFB0FF92D523F82B40D0F82E" box="[1145,1250,2012,2037]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="20" pageNumber="471" unit="mm" value="10.6">10.6 mm</quantity>
, 12.6×
<quantity id="783B07BEFFB0FF92D46AF82B41AAF82E" box="[1328,1432,2012,2037]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="20" pageNumber="471" unit="mm" value="10.5">10.5 mm</quantity>
, 14.0×
<quantity id="783B07BEFFB1FF93D18DFF6F4577FF6A" box="[215,325,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="21" pageNumber="472" unit="mm" value="12.2">12.2 mm</quantity>
) (
<specimenCode id="EF650220FFB1FF93D03EFF6F4615FF6A" box="[356,551,151,176]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="21" pageNumber="472">ZRC 2018.0795</specimenCode>
), Peninsular
<collectingCountry id="C7D4EACBFFB1FF93D392FF604701FF6A" box="[712,819,151,176]" name="Malaysia" pageId="21" pageNumber="472">Malaysia</collectingCountry>
:
<location id="BA1CFC80FFB1FF93D21CFF6047C0FF6A" LSID="urn:lsid:plazi:treatment:376A1B4DFFB0FF91D1CDF8F24053FD72:BA1CFC80FFB1FF93D21CFF6047C0FF6A" box="[838,1010,151,176]" country="Malaysia" name="Tioman Island" pageId="21" pageNumber="472">Tioman Island</location>
,
<collectingDate id="DB397573FFB1FF93D55CFF6F40AEFF6A" box="[1030,1180,151,176]" pageId="21" pageNumber="472" value="2012-07-04">
<date id="CB7D8C9BFFB1FF93D55CFF6F40AEFF6A" box="[1030,1180,151,176]" pageId="21" pageNumber="472" value="2012-07-04">4 July 2012</date>
</collectingDate>
,
<collectorName id="1236CF8DFFB1FF93D5EAFF6F4113FF6A" box="[1200,1313,152,176]" pageId="21" pageNumber="472">B.Y. Lee.</collectorName>
</materialsCitation>
—2 males (11.4×10.0 mm, 11.7×
<quantity id="783B07BEFFB1FF93D0CDFF4B45C6FF0E" box="[407,500,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="21" pageNumber="472" unit="mm" value="9.8">9.8 mm</quantity>
),
<materialsCitation id="0FABA006FFB1FF93D350FF4B45BCFF22" ID-GBIF-Occurrence="1913889790" collectingDate="2012-07-05" collectionCode="ZRC" collectorName="B. Y. Lee." country="Malaysia" location="Tioman Island" pageId="21" pageNumber="472" specimenCode="ZRC 2018.0796" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<specimenCount id="A9C561D2FFB1FF93D350FF4B465CFF0F" box="[522,622,188,213]" pageId="21" pageNumber="472" type="female">1 female</specimenCount>
(13.5×11.2) (
<specimenCode id="EF650220FFB1FF93D257FF4B47F9FF0F" box="[781,971,188,213]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="21" pageNumber="472">ZRC 2018.0796</specimenCode>
), Peninsular
<collectingCountry id="C7D4EACBFFB1FF93D53AFF4B40F9FF0F" box="[1120,1227,188,213]" name="Malaysia" pageId="21" pageNumber="472">Malaysia</collectingCountry>
:
<location id="BA1CFC80FFB1FF93D583FF4B41B2FF0F" LSID="urn:lsid:plazi:treatment:376A1B4DFFB0FF91D1CDF8F24053FD72:BA1CFC80FFB1FF93D583FF4B41B2FF0F" box="[1241,1408,188,213]" country="Malaysia" name="Tioman Island" pageId="21" pageNumber="472">Tioman Island</location>
,
<collectingDate id="DB397573FFB1FF93D4D4FF4B453CFF22" pageId="21" pageNumber="472" value="2012-07-05">
<date id="CB7D8C9BFFB1FF93D4D4FF4B453CFF22" pageId="21" pageNumber="472" value="2012-07-05">5 July 2012</date>
</collectingDate>
,
<collectorName id="1236CF8DFFB1FF93D045FF1745BCFF22" box="[287,398,224,248]" pageId="21" pageNumber="472">B.Y. Lee.</collectorName>
</materialsCitation>
—3 males (12.7×
<quantity id="783B07BEFFB1FF93D307FF1746F8FF2D" box="[605,714,224,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="21" pageNumber="472" unit="mm" value="10.8">10.8 mm</quantity>
, 13.1×
<quantity id="783B07BEFFB1FF93D241FF1747BAFF2D" box="[795,904,224,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="21" pageNumber="472" unit="mm" value="10.8">10.8 mm</quantity>
, 13.1×
<quantity id="783B07BEFFB1FF93D280FF17407BFF22" box="[986,1097,224,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="21" pageNumber="472" unit="mm" value="10.9">10.9 mm</quantity>
),
<materialsCitation id="0FABA006FFB1FF93D538FF174146FEC6" ID-GBIF-Occurrence="1913889801" collectingDate="2012-07-06" collectionCode="ZRC" collectorName="B. Y. Lee." country="Malaysia" location="Tioman Island" pageId="21" pageNumber="472" specimenCode="ZRC 2018.0797" specimenCount="3" specimenCount-female="3" typeStatus="holotype">
<specimenCount id="A9C561D2FFB1FF93D538FF1740E6FF22" box="[1122,1236,223,248]" pageId="21" pageNumber="472" type="female">3 females</specimenCount>
(12.3×
<quantity id="783B07BEFFB1FF93D470FF1741AAFF2D" box="[1322,1432,224,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="21" pageNumber="472" unit="mm" value="10.4">10.4 mm</quantity>
, 13.6×
<quantity id="783B07BEFFB1FF93D18DFEF3450FFEC6" box="[215,317,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="21" pageNumber="472" unit="mm" value="11.5">11.5 mm</quantity>
, 16.1×14.0 mm) (
<specimenCode id="EF650220FFB1FF93D35FFEF346F3FEC6" box="[517,705,260,284]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="21" pageNumber="472">ZRC 2018.0797</specimenCode>
), Peninsular
<collectingCountry id="C7D4EACBFFB1FF93D20BFEF3478EFEC7" box="[849,956,260,285]" name="Malaysia" pageId="21" pageNumber="472">Malaysia</collectingCountry>
:
<location id="BA1CFC80FFB1FF93D29DFEF34059FEC7" LSID="urn:lsid:plazi:treatment:376A1B4DFFB0FF91D1CDF8F24053FD72:BA1CFC80FFB1FF93D29DFEF34059FEC7" box="[967,1131,260,285]" country="Malaysia" name="Tioman Island" pageId="21" pageNumber="472">Tioman Island</location>
,
<collectingDate id="DB397573FFB1FF93D52DFEF340CFFEC6" box="[1143,1277,260,285]" pageId="21" pageNumber="472" value="2012-07-06">
<date id="CB7D8C9BFFB1FF93D52DFEF340CFFEC6" box="[1143,1277,260,285]" pageId="21" pageNumber="472" value="2012-07-06">6 July 2012</date>
</collectingDate>
,
<collectorName id="1236CF8DFFB1FF93D453FEF24146FEC6" box="[1289,1396,260,284]" pageId="21" pageNumber="472">B.Y. Lee.</collectorName>
</materialsCitation>
—1 male (12.2×
<quantity id="783B07BEFFB1FF93D045FED045BBFE9A" box="[287,393,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="21" pageNumber="472" unit="mm" value="10.5">10.5 mm</quantity>
),
<materialsCitation id="0FABA006FFB1FF93D0C7FEDF4575FEBE" ID-GBIF-Occurrence="1913889789" collectingDate="2012-07-07" collectionCode="ZRC" collectorName="B. Y. Lee." country="Malaysia" location="Tioman Island" pageId="21" pageNumber="472" specimenCode="ZRC 2012.0755" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<specimenCount id="A9C561D2FFB1FF93D0C7FEDF4632FE9A" box="[413,512,295,320]" pageId="21" pageNumber="472" type="female">1 female</specimenCount>
(14.5×13.0 mm) (
<specimenCode id="EF650220FFB1FF93D38BFEDF47BFFE9A" box="[721,909,295,320]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="21" pageNumber="472">ZRC 2012.0755</specimenCode>
b), Peninsular
<collectingCountry id="C7D4EACBFFB1FF93D574FED040ABFE9A" box="[1070,1177,295,320]" name="Malaysia" pageId="21" pageNumber="472">Malaysia</collectingCountry>
:
<location id="BA1CFC80FFB1FF93D5FFFED04178FE9A" LSID="urn:lsid:plazi:treatment:376A1B4DFFB0FF91D1CDF8F24053FD72:BA1CFC80FFB1FF93D5FFFED04178FE9A" box="[1189,1354,295,320]" country="Malaysia" name="Tioman Island" pageId="21" pageNumber="472">Tioman Island</location>
,
<collectingDate id="DB397573FFB1FF93D40DFEDF44E0FEBE" pageId="21" pageNumber="472" value="2012-07-07">
<date id="CB7D8C9BFFB1FF93D40DFEDF44E0FEBE" pageId="21" pageNumber="472" value="2012-07-07">7 July 2012</date>
</collectingDate>
,
<collectorName id="1236CF8DFFB1FF93D187FEBA4575FEBE" box="[221,327,332,356]" pageId="21" pageNumber="472">B.Y. Lee.</collectorName>
</materialsCitation>
—3 males (15.0×
<quantity id="783B07BEFFB1FF93D351FEBB4643FEBE" box="[523,625,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="21" pageNumber="472" unit="mm" value="13.5">13.5 mm</quantity>
, 13.4×
<quantity id="783B07BEFFB1FF93D3E7FEBB4711FEBE" box="[701,803,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="21" pageNumber="472" unit="mm" value="11.8">11.8 mm</quantity>
, 10.9×
<quantity id="783B07BEFFB1FF93D235FEBB47F8FEBE" box="[879,970,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="21" pageNumber="472" unit="mm" value="9.8">9.8 mm</quantity>
),
<materialsCitation id="0FABA006FFB1FF93D284FEBB47D8FE52" ID-GBIF-Occurrence="1913889799" collectingDate="2018-02-27" collectionCode="ZRC" collectorName="J. Li" country="Malaysia" location="Borneo" pageId="21" pageNumber="472" specimenCode="ZRC 2018.0799" specimenCount="2" specimenCount-female="2" stateProvince="Sabah" typeStatus="holotype">
<specimenCount id="A9C561D2FFB1FF93D284FEBB4078FEBF" box="[990,1098,332,357]" pageId="21" pageNumber="472" type="female">2 females</specimenCount>
(13.9×13.6, 12.3×11.1) (
<specimenCode id="EF650220FFB1FF93D43EFEBB4521FE52" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="21" pageNumber="472">ZRC 2018.0799</specimenCode>
),
<collectingCountry id="C7D4EACBFFB1FF93D071FE9845A4FE52" box="[299,406,367,392]" name="Malaysia" pageId="21" pageNumber="472">Malaysia</collectingCountry>
:
<location id="BA1CFC80FFB1FF93D0FCFE8745CFFE5D" LSID="urn:lsid:plazi:treatment:376A1B4DFFB0FF91D1CDF8F24053FD72:BA1CFC80FFB1FF93D0FCFE8745CFFE5D" box="[422,509,368,391]" country="Malaysia" name="Borneo" pageId="21" pageNumber="472" stateProvince="Sabah">Borneo</location>
:
<collectingRegion id="7D0764B9FFB1FF93D356FE984661FE52" box="[524,595,367,392]" country="Malaysia" name="Sabah" pageId="21" pageNumber="472">Sabah</collectingRegion>
,
<collectingDate id="DB397573FFB1FF93D33EFE87470BFE52" box="[612,825,367,392]" pageId="21" pageNumber="472" value="2018-02-27">
<date id="CB7D8C9BFFB1FF93D33EFE87470BFE52" box="[612,825,367,392]" pageId="21" pageNumber="472" value="2018-02-27">27 February 2018</date>
</collectingDate>
, J.-
<collectorName id="1236CF8DFFB1FF93D23FFE8747AEFE52" box="[869,924,367,392]" pageId="21" pageNumber="472">J. Li</collectorName>
<emphasis id="8DB77649FFB1FF93D2F2FE8647D4FE52" box="[936,998,367,392]" italics="true" pageId="21" pageNumber="472">et al.</emphasis>
.
</materialsCitation>
—1 male (12.6×
<quantity id="783B07BEFFB1FF93D5F5FE98412FFE52" box="[1199,1309,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="21" pageNumber="472" unit="mm" value="11.5">11.5 mm</quantity>
),
<materialsCitation id="0FABA006FFB1FF93D46FFE874082FE76" ID-GBIF-Occurrence="1913889794" collectingDate="2018-02-27" collectionCode="MZB" collectorName="J. Li" country="Malaysia" location="Borneo" municipality="Cru" pageId="21" pageNumber="472" specimenCount="1" specimenCount-female="1" stateProvince="Sabah" typeStatus="holotype">
<specimenCount id="A9C561D2FFB1FF93D46FFE8741AEFE52" box="[1333,1436,367,392]" pageId="21" pageNumber="472" type="female">1 female</specimenCount>
(15.4×
<quantity id="783B07BEFFB1FF93D1BAFE634574FE76" box="[224,326,404,429]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="21" pageNumber="472" unit="mm" value="13.8">13.8 mm</quantity>
) (
<collectionCode id="D9D2329EFFB1FF93D007FE6345A9FE76" box="[349,411,404,428]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="21" pageNumber="472">MZB</collectionCode>
<collectingMunicipality id="5F183021FFB1FF93D0F8FE6345FEFE76" box="[418,460,404,428]" pageId="21" pageNumber="472">Cru</collectingMunicipality>
4800),
<collectingCountry id="C7D4EACBFFB1FF93D378FE6346BFFE77" box="[546,653,404,429]" name="Malaysia" pageId="21" pageNumber="472">Malaysia</collectingCountry>
:
<location id="BA1CFC80FFB1FF93D3CDFE6246DCFE76" LSID="urn:lsid:plazi:treatment:376A1B4DFFB0FF91D1CDF8F24053FD72:BA1CFC80FFB1FF93D3CDFE6246DCFE76" box="[663,750,405,428]" country="Malaysia" municipality="Cru" name="Borneo" pageId="21" pageNumber="472" stateProvince="Sabah">Borneo</location>
:
<collectingRegion id="7D0764B9FFB1FF93D3A2FE63470DFE77" box="[760,831,404,429]" country="Malaysia" name="Sabah" pageId="21" pageNumber="472">Sabah</collectingRegion>
,
<collectingDate id="DB397573FFB1FF93D211FE634027FE76" box="[843,1045,404,429]" pageId="21" pageNumber="472" value="2018-02-27">
<date id="CB7D8C9BFFB1FF93D211FE634027FE76" box="[843,1045,404,429]" pageId="21" pageNumber="472" value="2018-02-27">27 February 2018</date>
</collectingDate>
, J.-
<collectorName id="1236CF8DFFB1FF93D561FE62405FFE77" box="[1083,1133,404,429]" pageId="21" pageNumber="472">J. Li</collectorName>
<emphasis id="8DB77649FFB1FF93D52EFE62409EFE77" box="[1140,1196,404,429]" italics="true" pageId="21" pageNumber="472">et al.</emphasis>
.
</materialsCitation>
—
<materialsCitation id="0FABA006FFB1FF93D589FE634051FE0A" ID-GBIF-Occurrence="1913889808" collectingDate="2018-02-27" collectorName="J. Li" country="Malaysia" location="Borneo" pageId="21" pageNumber="472" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
<specimenCount id="A9C561D2FFB1FF93D589FE634128FE77" box="[1235,1306,404,429]" pageId="21" pageNumber="472" type="male">1 male</specimenCount>
(13.7×
<quantity id="783B07BEFFB1FF93D430FE6344F6FE0A" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="21" pageNumber="472" unit="mm" value="12.4">12.4 mm</quantity>
) (NMNS-7779-027),
<collectingCountry id="C7D4EACBFFB1FF93D0E4FE40461BFE0A" box="[446,553,439,464]" name="Malaysia" pageId="21" pageNumber="472">Malaysia</collectingCountry>
:
<location id="BA1CFC80FFB1FF93D36CFE4F46BFFE15" LSID="urn:lsid:plazi:treatment:376A1B4DFFB0FF91D1CDF8F24053FD72:BA1CFC80FFB1FF93D36CFE4F46BFFE15" box="[566,653,440,463]" country="Malaysia" name="Borneo" pageId="21" pageNumber="472" stateProvince="Sabah">Borneo</location>
:
<collectingRegion id="7D0764B9FFB1FF93D3C0FE4046D3FE0A" box="[666,737,439,464]" country="Malaysia" name="Sabah" pageId="21" pageNumber="472">Sabah</collectingRegion>
,
<collectingDate id="DB397573FFB1FF93D3B5FE4F478CFE0A" box="[751,958,439,464]" pageId="21" pageNumber="472" value="2018-02-27">
<date id="CB7D8C9BFFB1FF93D3B5FE4F478CFE0A" box="[751,958,439,464]" pageId="21" pageNumber="472" value="2018-02-27">27 February 2018</date>
</collectingDate>
, J.-
<collectorName id="1236CF8DFFB1FF93D2BDFE4F4029FE0A" box="[999,1051,439,464]" pageId="21" pageNumber="472">J. Li</collectorName>
<emphasis id="8DB77649FFB1FF93D57EFE4E406DFE0A" box="[1060,1119,439,464]" italics="true" pageId="21" pageNumber="472">et al.</emphasis>
.
</materialsCitation>
—
<materialsCitation id="0FABA006FFB1FF93D5D8FE4F40AEFE2E" ID-GBIF-Occurrence="1913889796" collectingDate="2018-02-27" collectionCode="NMCR" collectorName="J. Li" country="Malaysia" location="Borneo" pageId="21" pageNumber="472" specimenCode="NMCR 50704" specimenCount="2" specimenCount-male="2" stateProvince="Sabah" typeStatus="holotype">
<specimenCount id="A9C561D2FFB1FF93D5D8FE4F40EEFE0A" box="[1154,1244,439,464]" pageId="21" pageNumber="472" type="male">2 males</specimenCount>
(14.5×13.0 mm, 14.3×
<quantity id="783B07BEFFB1FF93D18DFE2B450FFE2E" box="[215,317,476,500]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="21" pageNumber="472" unit="mm" value="13.4">13.4 mm</quantity>
) (
<specimenCode id="EF650220FFB1FF93D009FE2B45CAFE2F" box="[339,504,476,501]" collectionCode="NMCR" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14936" name="New Mexico State University" pageId="21" pageNumber="472">NMCR 50704</specimenCode>
),
<collectingCountry id="C7D4EACBFFB1FF93D356FE2B4645FE2F" box="[524,631,476,501]" name="Malaysia" pageId="21" pageNumber="472">Malaysia</collectingCountry>
:
<location id="BA1CFC80FFB1FF93D3DBFE2A46EAFE2E" LSID="urn:lsid:plazi:treatment:376A1B4DFFB0FF91D1CDF8F24053FD72:BA1CFC80FFB1FF93D3DBFE2A46EAFE2E" box="[641,728,477,500]" country="Malaysia" name="Borneo" pageId="21" pageNumber="472" stateProvince="Sabah">Borneo</location>
:
<collectingRegion id="7D0764B9FFB1FF93D3B8FE2B471BFE2F" box="[738,809,476,501]" country="Malaysia" name="Sabah" pageId="21" pageNumber="472">Sabah</collectingRegion>
,
<collectingDate id="DB397573FFB1FF93D26FFE2B47CDFE2E" box="[821,1023,476,501]" pageId="21" pageNumber="472" value="2018-02-27">
<date id="CB7D8C9BFFB1FF93D26FFE2B47CDFE2E" box="[821,1023,476,501]" pageId="21" pageNumber="472" value="2018-02-27">27 February 2018</date>
</collectingDate>
, J.-
<collectorName id="1236CF8DFFB1FF93D57FFE2A4065FE2F" box="[1061,1111,476,501]" pageId="21" pageNumber="472">J. Li</collectorName>
<emphasis id="8DB77649FFB1FF93D507FE2A40A4FE2F" box="[1117,1174,476,501]" italics="true" pageId="21" pageNumber="472">et al.</emphasis>
.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="EBBCFAD3FFB1FF93D1CDF9054511F89F" httpUri="https://zenodo.org/record/1440735/files/figure.png" pageId="21" pageNumber="472" startId="21.[151,250,1778,1800]" targetBox="[152,1435,608,1724]" targetPageId="21">
<paragraph id="BF7CAA5BFFB1FF93D1CDF9054511F89F" blockId="21.[151,1436,1778,1861]" pageId="21" pageNumber="472">
<emphasis id="8DB77649FFB1FF93D1CDF905452CF8DD" bold="true" box="[151,286,1778,1800]" pageId="21" pageNumber="472">FIGURE 13</emphasis>
.
<taxonomicName id="78C3D1D8FFB1FF93D06AF9044600F8D2" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[304,562,1779,1800]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="purpureum" status="sp. nov.">
<emphasis id="8DB77649FFB1FF93D06AF9044600F8D2" box="[304,562,1779,1800]" italics="true" pageId="21" pageNumber="472">Parasesarma purpureum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFB1FF93D367F904464EF8D2" box="[573,636,1779,1800]" pageId="21" pageNumber="472" rank="species">
<emphasis id="8DB77649FFB1FF93D367F904464EF8D2" bold="true" box="[573,636,1779,1800]" pageId="21" pageNumber="472">n. sp.</emphasis>
</taxonomicNameLabel>
. A–D, holotype male (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; E, F, paratype female (ZRC 2012.0755b), same locality. A, E, dorsal views; B, F, ventral views; C, D, frontal views.
</paragraph>
</caption>
<subSubSection id="F7D9F9D0FFB1FF91D19DF8854669FF0E" lastPageId="23" lastPageNumber="474" pageId="21" pageNumber="472" type="diagnosis">
<paragraph id="BF7CAA5BFFB1FF90D19DF885416EFE2E" blockId="21.[151,1436,1906,2003]" lastBlockId="22.[151,1437,151,501]" lastPageId="22" lastPageNumber="473" pageId="21" pageNumber="472">
<emphasis id="8DB77649FFB1FF93D19DF8854573F851" bold="true" box="[199,321,1906,1931]" pageId="21" pageNumber="472">Diagnosis.</emphasis>
Carapace (
<figureCitation id="27F8B6DEFFB1FF93D0E4F885467CF850" box="[446,590,1906,1931]" captionStart="FIGURE 13" captionStartId="21.[151,250,1778,1800]" captionTargetBox="[152,1435,608,1724]" captionTargetId="figure@21.[152,1435,608,1724]" captionTargetPageId="21" captionText="FIGURE 13. Parasesarma purpureum n. sp.. A–D, holotype male (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; E, F, paratype female (ZRC 2012.0755b), same locality. A, E, dorsal views; B, F, ventral views; C, D, frontal views." httpUri="https://zenodo.org/record/1440735/files/figure.png" pageId="21" pageNumber="472">Figs. 13A, E</figureCitation>
,
<figureCitation id="27F8B6DEFFB1FF93D303F88446BEF851" box="[601,652,1907,1931]" captionStart="FIGURE 14" captionStartId="22.[151,250,1911,1933]" captionTargetBox="[198,1357,638,1836]" captionTargetId="figure@22.[152,1435,568,1875]" captionTargetPageId="22" captionText="FIGURE 14. Parasesarma purpureum n. sp., holotype male (15.9×13.7 mm) (ZRC 2012.0755a), Tioman, Peninsular Malaysia. A, dorsal view of carapace; B–D, tip of left cheliped; E–H, left G1; B, lateral view of pectinate cristae on palm; C, dorsal view of palm and dactylar finger; D, lateral view of dactylar finger; E, G, dorsal (sternal) views; F, H, ventral (pleonal) views; E, F, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440737/files/figure.png" pageId="21" pageNumber="472">14A</figureCitation>
,
<figureCitation id="27F8B6DEFFB1FF93D3CDF88446FAF851" box="[663,712,1907,1931]" captionStart="FIGURE 22" captionStartId="32.[151,250,1490,1512]" captionTargetBox="[152,1435,193,1469]" captionTargetId="figure@32.[152,1435,193,1469]" captionTargetPageId="32" captionText="FIGURE 22. Colour in life, males, dorsal views. A. Parasesarma gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; B. P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C. P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. purpureum n. sp. (13.4×11.8 mm) (ZRC 2018.0799), Borneo, Malaysia; E, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440753/files/figure.png" pageId="21" pageNumber="472">22D</figureCitation>
) squarish in general outline, 1.2 times broader than long; regions well defined, separated by shallow grooves; postfrontal region distinct, separated into 4 lobes by deep grooves; front deflexed downwards (
<figureCitation id="27F8B6DEFFB1FF93D0BEF84D46B4F809" box="[484,646,1978,2003]" captionStart="FIGURE 13" captionStartId="21.[151,250,1778,1800]" captionTargetBox="[152,1435,608,1724]" captionTargetId="figure@21.[152,1435,608,1724]" captionTargetPageId="21" captionText="FIGURE 13. Parasesarma purpureum n. sp.. A–D, holotype male (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; E, F, paratype female (ZRC 2012.0755b), same locality. A, E, dorsal views; B, F, ventral views; C, D, frontal views." httpUri="https://zenodo.org/record/1440735/files/figure.png" pageId="21" pageNumber="472">Figs. 13C, D</figureCitation>
) margin slightly concave in dorsal view; lateral margin straight, subparallel along most of length before curving to join almost straight posterior carapace margin; cornea extending or just reaching tip of external orbital tooth (
<figureCitation id="27F8B6DEFFB2FF90D3C0FF4B471DFF0E" box="[666,815,188,213]" captionStart="FIGURE 13" captionStartId="21.[151,250,1778,1800]" captionTargetBox="[152,1435,608,1724]" captionTargetId="figure@21.[152,1435,608,1724]" captionTargetPageId="21" captionText="FIGURE 13. Parasesarma purpureum n. sp.. A–D, holotype male (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; E, F, paratype female (ZRC 2012.0755b), same locality. A, E, dorsal views; B, F, ventral views; C, D, frontal views." httpUri="https://zenodo.org/record/1440735/files/figure.png" pageId="22" pageNumber="473">Figs. 13A, E</figureCitation>
,
<figureCitation id="27F8B6DEFFB2FF90D267FF4B475DFF0E" box="[829,879,188,212]" captionStart="FIGURE 14" captionStartId="22.[151,250,1911,1933]" captionTargetBox="[198,1357,638,1836]" captionTargetId="figure@22.[152,1435,568,1875]" captionTargetPageId="22" captionText="FIGURE 14. Parasesarma purpureum n. sp., holotype male (15.9×13.7 mm) (ZRC 2012.0755a), Tioman, Peninsular Malaysia. A, dorsal view of carapace; B–D, tip of left cheliped; E–H, left G1; B, lateral view of pectinate cristae on palm; C, dorsal view of palm and dactylar finger; D, lateral view of dactylar finger; E, G, dorsal (sternal) views; F, H, ventral (pleonal) views; E, F, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440737/files/figure.png" pageId="22" pageNumber="473">14A</figureCitation>
,
<figureCitation id="27F8B6DEFFB2FF90D224FF4B4780FF0E" box="[894,946,188,212]" captionStart="FIGURE 22" captionStartId="32.[151,250,1490,1512]" captionTargetBox="[152,1435,193,1469]" captionTargetId="figure@32.[152,1435,193,1469]" captionTargetPageId="32" captionText="FIGURE 22. Colour in life, males, dorsal views. A. Parasesarma gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; B. P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C. P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. purpureum n. sp. (13.4×11.8 mm) (ZRC 2018.0799), Borneo, Malaysia; E, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440753/files/figure.png" pageId="22" pageNumber="473">22D</figureCitation>
). Ischium of third maxilliped with shallow median sulcus, merus with distinct submedian ridge; exopod slender, tip reaching half-length of outer margin of merus, flagellum long. Male cheliped palm with 2 transverse pectinate crests (12 and 7 corneous teeth, respectively) on upper surface; upper surface of dactylus with 9 or 10 symmetrical, obliquely elongate dactylar tubercles, proximal tubercles steep, sharp, the others tubercles large, distalmost tubercle indistinct (
<figureCitation id="27F8B6DEFFB2FF90D458FEBB41AAFEBE" box="[1282,1432,332,357]" captionStart="FIGURE 14" captionStartId="22.[151,250,1911,1933]" captionTargetBox="[198,1357,638,1836]" captionTargetId="figure@22.[152,1435,568,1875]" captionTargetPageId="22" captionText="FIGURE 14. Parasesarma purpureum n. sp., holotype male (15.9×13.7 mm) (ZRC 2012.0755a), Tioman, Peninsular Malaysia. A, dorsal view of carapace; B–D, tip of left cheliped; E–H, left G1; B, lateral view of pectinate cristae on palm; C, dorsal view of palm and dactylar finger; D, lateral view of dactylar finger; E, G, dorsal (sternal) views; F, H, ventral (pleonal) views; E, F, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440737/files/figure.png" pageId="22" pageNumber="473">Figs. 14C, D</figureCitation>
,
<figureCitation id="27F8B6DEFFB2FF90D1CDFE8744F4FE52" box="[151,198,368,392]" captionStart="FIGURE 19" captionStartId="28.[151,250,1811,1833]" captionTargetBox="[152,1435,199,1754]" captionTargetId="figure@28.[152,1435,199,1754]" captionTargetPageId="28" captionText="FIGURE 19. Outer views of chelae, holotype males. A, Parasesarma leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; C, P. macaco n. sp. (13.8×12.3 mm) (NMNS-7779- 005), Pingtung, Taiwan; D, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440747/files/figure.png" pageId="22" pageNumber="473">19F</figureCitation>
). Ambulatory legs relatively slender for this species-complex, P3 and P4 about 1.6 times carapace width; P3 and P4 coxae without dense setae; P3 merus 2.6 times as long as broad; P3 propodus 4.4 times as long as broad; P3 dactylus 0.4 times length of propodus (
<figureCitation id="27F8B6DEFFB2FF90D316FE404680FE0A" box="[588,690,439,464]" captionStart="FIGURE 20" captionStartId="30.[151,250,1268,1290]" captionTargetBox="[152,1435,393,1246]" captionTargetId="figure@30.[152,1435,393,1246]" captionTargetPageId="30" captionText="FIGURE 20. First to fourth ambulatory legs (P2–P5) (from top to bottom), holotype males. A, Parasesarma gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; B, P. macaco n. sp. male (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; E, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; F, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; G, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440749/files/figure.png" pageId="22" pageNumber="473">Fig. 20E</figureCitation>
). G1 relatively stout (
<figureCitation id="27F8B6DEFFB2FF90D2FCFE404005FE0A" box="[934,1079,439,464]" captionStart="FIGURE 14" captionStartId="22.[151,250,1911,1933]" captionTargetBox="[198,1357,638,1836]" captionTargetId="figure@22.[152,1435,568,1875]" captionTargetPageId="22" captionText="FIGURE 14. Parasesarma purpureum n. sp., holotype male (15.9×13.7 mm) (ZRC 2012.0755a), Tioman, Peninsular Malaysia. A, dorsal view of carapace; B–D, tip of left cheliped; E–H, left G1; B, lateral view of pectinate cristae on palm; C, dorsal view of palm and dactylar finger; D, lateral view of dactylar finger; E, G, dorsal (sternal) views; F, H, ventral (pleonal) views; E, F, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440737/files/figure.png" pageId="22" pageNumber="473">Figs. 14E–H</figureCitation>
,
<figureCitation id="27F8B6DEFFB2FF90D518FE4F4040FE0A" box="[1090,1138,440,464]" captionStart="FIGURE 21" captionStartId="31.[151,250,1815,1837]" captionTargetBox="[152,1435,260,1731]" captionTargetId="figure@31.[152,1435,260,1731]" captionTargetPageId="31" captionText="FIGURE 21. Gonopods of Parasesarma leptosoma species-complex. A, P. leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; C, P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; D, P. kui n. sp., holotype (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp., holotype (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp., holotype (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp., holotype (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp., holotype (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440751/files/figure.png" pageId="22" pageNumber="473">21F</figureCitation>
,); apical process corneous, short, bent at angle of 45°, long, stout, ending in rounded tip. G2 longer than quarter length of G1 (
<figureCitation id="27F8B6DEFFB2FF90D5B1FE2B417DFE2E" box="[1259,1359,476,501]" captionStart="FIGURE 21" captionStartId="31.[151,250,1815,1837]" captionTargetBox="[152,1435,260,1731]" captionTargetId="figure@31.[152,1435,260,1731]" captionTargetPageId="31" captionText="FIGURE 21. Gonopods of Parasesarma leptosoma species-complex. A, P. leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; C, P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; D, P. kui n. sp., holotype (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp., holotype (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp., holotype (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp., holotype (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp., holotype (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440751/files/figure.png" pageId="22" pageNumber="473">Fig. 21F</figureCitation>
).
</paragraph>
<caption id="EBBCFAD3FFB2FF90D1CDF88046BDF833" httpUri="https://zenodo.org/record/1440737/files/figure.png" pageId="22" pageNumber="473" startId="22.[151,250,1911,1933]" targetBox="[198,1357,638,1836]" targetPageId="22">
<paragraph id="BF7CAA5BFFB2FF90D1CDF88046BDF833" blockId="22.[151,1436,1911,2025]" pageId="22" pageNumber="473">
<emphasis id="8DB77649FFB2FF90D1CDF880452DF856" bold="true" box="[151,287,1911,1933]" pageId="22" pageNumber="473">FIGURE 14</emphasis>
.
<taxonomicName id="78C3D1D8FFB2FF90D069F88F4604F857" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[307,566,1912,1933]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="473" phylum="Arthropoda" rank="species" species="purpureum" status="sp. nov.">
<emphasis id="8DB77649FFB2FF90D069F88F4604F857" box="[307,566,1912,1933]" italics="true" pageId="22" pageNumber="473">Parasesarma purpureum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFB2FF90D31EF88046B6F856" box="[580,644,1911,1932]" pageId="22" pageNumber="473" rank="species">
<emphasis id="8DB77649FFB2FF90D31EF88046B6F856" bold="true" box="[580,644,1911,1932]" pageId="22" pageNumber="473">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype male (15.9×13.7 mm) (ZRC 2012.0755a), Tioman, Peninsular Malaysia. A, dorsal view of carapace; B–D, tip of left cheliped; E–H, left G1; B, lateral view of pectinate cristae on palm; C, dorsal view of palm and dactylar finger; D, lateral view of dactylar finger; E, G, dorsal (sternal) views; F, H, ventral (pleonal) views; E, F, distal part of G1. Scale bars: 1.0 mm.
</paragraph>
</caption>
<paragraph id="BF7CAA5BFFB3FF91D19DFF604669FF0E" blockId="23.[151,1437,151,680]" pageId="23" pageNumber="474">
<emphasis id="8DB77649FFB3FF91D19DFF60454DFF6A" bold="true" box="[199,383,151,176]" pageId="23" pageNumber="474">Colour in life.</emphasis>
Carapace dark brown, fingers of chelipeds are pale white, ambulatory legs violet or purplish brown. (
<figureCitation id="27F8B6DEFFB3FF91D03BFF4B45FFFF0E" box="[353,461,188,213]" captionStart="FIGURE 22" captionStartId="32.[151,250,1490,1512]" captionTargetBox="[152,1435,193,1469]" captionTargetId="figure@32.[152,1435,193,1469]" captionTargetPageId="32" captionText="FIGURE 22. Colour in life, males, dorsal views. A. Parasesarma gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; B. P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C. P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. purpureum n. sp. (13.4×11.8 mm) (ZRC 2018.0799), Borneo, Malaysia; E, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440753/files/figure.png" pageId="23" pageNumber="474">Fig. 22D</figureCitation>
,
<figureCitation id="27F8B6DEFFB3FF91D083FF4B463CFF0E" box="[473,526,188,212]" captionStart="FIGURE 23" captionStartId="33.[151,250,1605,1627]" captionTargetBox="[152,1435,245,1548]" captionTargetId="figure@33.[152,1435,245,1548]" captionTargetPageId="33" captionText="FIGURE 23. Colour in life, males, ventral views. A. Parasesarma gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; B. P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C. P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. purpureum n. sp. (13.4×11.8 mm) (ZRC 2018.0799), Borneo, Malaysia; E, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440755/files/figure.png" pageId="23" pageNumber="474">23D</figureCitation>
,
<figureCitation id="27F8B6DEFFB3FF91D340FF4B467CFF0E" box="[538,590,188,212]" captionStart="FIGURE 24" captionStartId="34.[151,250,1531,1553]" captionTargetBox="[151,1436,218,1490]" captionTargetId="figure@34.[151,1436,218,1490]" captionTargetPageId="34" captionText="FIGURE 24. In situ photographs of Parasesarma leptosoma species-complex, specimens not collected. A, P. leptosoma (Hilgendorf, 1869), Kenya (Photograph: S. Cannicci); B, P. gecko n. sp., Okinawa, Japan; C, P. macaco n. sp., Pingtung, Taiwan; D, P. kui n. sp., Pingtung, Taiwan; E, P. purpureum n. sp., Borneo, Malaysia; F, P. tarantula n. sp., Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440757/files/figure.png" pageId="23" pageNumber="474">24E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFB3FF91D19DFF284062FE2E" pageId="23" pageNumber="474" type="discussion">
<paragraph id="BF7CAA5BFFB3FF91D19DFF284062FE2E" blockId="23.[151,1437,151,680]" pageId="23" pageNumber="474">
<emphasis id="8DB77649FFB3FF91D19DFF28450DFF22" bold="true" box="[199,319,223,248]" pageId="23" pageNumber="474">Remarks.</emphasis>
<taxonomicName id="78C3D1D8FFB3FF91D012FF16465EFF22" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[328,620,225,248]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="purpureum" status="sp. nov.">
<emphasis id="8DB77649FFB3FF91D012FF16465EFF22" box="[328,620,225,248]" italics="true" pageId="23" pageNumber="474">Parasesarma purpureum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFB3FF91D32FFF28468AFF22" box="[629,696,223,248]" pageId="23" pageNumber="474" rank="species">
<emphasis id="8DB77649FFB3FF91D32FFF28468AFF22" bold="true" box="[629,696,223,248]" pageId="23" pageNumber="474">n. sp.</emphasis>
</taxonomicNameLabel>
, is most similar to
<taxonomicName id="78C3D1D8FFB3FF91D2FAFF1647ECFF22" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[928,990,223,248]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="kui" status="sp. nov.">
<emphasis id="8DB77649FFB3FF91D2FAFF1647ECFF22" box="[928,990,223,248]" italics="true" pageId="23" pageNumber="474">P. kui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFB3FF91D2B1FF28401CFF22" box="[1003,1070,223,248]" pageId="23" pageNumber="474" rank="species">
<emphasis id="8DB77649FFB3FF91D2B1FF28401CFF22" bold="true" box="[1003,1070,223,248]" pageId="23" pageNumber="474">n. sp.</emphasis>
</taxonomicNameLabel>
(from
<collectingCountry id="C7D4EACBFFB3FF91D5D8FF2840E8FF22" box="[1154,1242,223,248]" name="Taiwan" pageId="23" pageNumber="474">Taiwan</collectingCountry>
) with regards to its relatively broader carapace. However, the following differences are observed; the carapace surface of
<taxonomicName id="78C3D1D8FFB3FF91D4DCFEF2452AFE9A" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="purpureum">
<emphasis id="8DB77649FFB3FF91D4DCFEF2452AFE9A" italics="true" pageId="23" pageNumber="474">P. purpureum</emphasis>
</taxonomicName>
has scattered tufts of short setae, the dactylar tubercles of the male chela are consistently nine in number (
<figureCitation id="27F8B6DEFFB3FF91D05FFEBB45B3FEBE" box="[261,385,332,357]" captionStart="FIGURE 14" captionStartId="22.[151,250,1911,1933]" captionTargetBox="[198,1357,638,1836]" captionTargetId="figure@22.[152,1435,568,1875]" captionTargetPageId="22" captionText="FIGURE 14. Parasesarma purpureum n. sp., holotype male (15.9×13.7 mm) (ZRC 2012.0755a), Tioman, Peninsular Malaysia. A, dorsal view of carapace; B–D, tip of left cheliped; E–H, left G1; B, lateral view of pectinate cristae on palm; C, dorsal view of palm and dactylar finger; D, lateral view of dactylar finger; E, G, dorsal (sternal) views; F, H, ventral (pleonal) views; E, F, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440737/files/figure.png" pageId="23" pageNumber="474">Figs. 14D</figureCitation>
,
<figureCitation id="27F8B6DEFFB3FF91D0C8FEBB45F1FEBE" box="[402,451,332,356]" captionStart="FIGURE 19" captionStartId="28.[151,250,1811,1833]" captionTargetBox="[152,1435,199,1754]" captionTargetId="figure@28.[152,1435,199,1754]" captionTargetPageId="28" captionText="FIGURE 19. Outer views of chelae, holotype males. A, Parasesarma leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; C, P. macaco n. sp. (13.8×12.3 mm) (NMNS-7779- 005), Pingtung, Taiwan; D, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440747/files/figure.png" pageId="23" pageNumber="474">19F</figureCitation>
,); the corneous tip of the G1 is proportionately broader (
<figureCitation id="27F8B6DEFFB3FF91D5D0FEBB411BFEBE" box="[1162,1321,332,357]" captionStart="FIGURE 14" captionStartId="22.[151,250,1911,1933]" captionTargetBox="[198,1357,638,1836]" captionTargetId="figure@22.[152,1435,568,1875]" captionTargetPageId="22" captionText="FIGURE 14. Parasesarma purpureum n. sp., holotype male (15.9×13.7 mm) (ZRC 2012.0755a), Tioman, Peninsular Malaysia. A, dorsal view of carapace; B–D, tip of left cheliped; E–H, left G1; B, lateral view of pectinate cristae on palm; C, dorsal view of palm and dactylar finger; D, lateral view of dactylar finger; E, G, dorsal (sternal) views; F, H, ventral (pleonal) views; E, F, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440737/files/figure.png" pageId="23" pageNumber="474">Figs. 14E–H</figureCitation>
); and the general colour in life is dark purple, or brownish-purplish with white chelae (
<figureCitation id="27F8B6DEFFB3FF91D578FE9840ABFE52" box="[1058,1177,367,392]" captionStart="FIGURE 22" captionStartId="32.[151,250,1490,1512]" captionTargetBox="[152,1435,193,1469]" captionTargetId="figure@32.[152,1435,193,1469]" captionTargetPageId="32" captionText="FIGURE 22. Colour in life, males, dorsal views. A. Parasesarma gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; B. P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C. P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. purpureum n. sp. (13.4×11.8 mm) (ZRC 2018.0799), Borneo, Malaysia; E, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440753/files/figure.png" pageId="23" pageNumber="474">Figs. 22D</figureCitation>
,
<figureCitation id="27F8B6DEFFB3FF91D5FFFE8740EEFE52" box="[1189,1244,368,392]" captionStart="FIGURE 23" captionStartId="33.[151,250,1605,1627]" captionTargetBox="[152,1435,245,1548]" captionTargetId="figure@33.[152,1435,245,1548]" captionTargetPageId="33" captionText="FIGURE 23. Colour in life, males, ventral views. A. Parasesarma gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; B. P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C. P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. purpureum n. sp. (13.4×11.8 mm) (ZRC 2018.0799), Borneo, Malaysia; E, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440755/files/figure.png" pageId="23" pageNumber="474">23D</figureCitation>
). By contrast, in
<taxonomicName id="78C3D1D8FFB3FF91D1CDFE6244EBFE77" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[151,217,404,429]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="kui" status="sp. nov.">
<emphasis id="8DB77649FFB3FF91D1CDFE6244EBFE77" box="[151,217,404,429]" italics="true" pageId="23" pageNumber="474">P. kui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFB3FF91D1BBFE634517FE77" box="[225,293,404,429]" pageId="23" pageNumber="474" rank="species">
<emphasis id="8DB77649FFB3FF91D1BBFE634517FE77" bold="true" box="[225,293,404,429]" pageId="23" pageNumber="474">n. sp.</emphasis>
</taxonomicNameLabel>
, the carapace surface is glabrous; the dactylar tubercles are nine or ten (
<figureCitation id="27F8B6DEFFB3FF91D5D9FE63412CFE76" box="[1155,1310,404,429]" captionStart="FIGURE 10" captionStartId="17.[151,250,1675,1697]" captionTargetBox="[175,1387,477,1591]" captionTargetId="figure@17.[151,1435,445,1654]" captionTargetPageId="17" captionText="FIGURE 10. Parasesarma kui n. sp., holotype male (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan. A, dorsal view of carapace; B–D, tip of left cheliped; E–H, left G1. B, lateral view of pectinated cristae on palm; C, dorsal view of palm and dactylar finger; D, lateral view of dactylar finger; E, G, dorsal (sternal) views; F, H, ventral (pleonal) views; E, F, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440729/files/figure.png" pageId="23" pageNumber="474">Figs. 10C, D</figureCitation>
,
<figureCitation id="27F8B6DEFFB3FF91D471FE634150FE76" box="[1323,1378,404,428]" captionStart="FIGURE 19" captionStartId="28.[151,250,1811,1833]" captionTargetBox="[152,1435,199,1754]" captionTargetId="figure@28.[152,1435,199,1754]" captionTargetPageId="28" captionText="FIGURE 19. Outer views of chelae, holotype males. A, Parasesarma leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; C, P. macaco n. sp. (13.8×12.3 mm) (NMNS-7779- 005), Pingtung, Taiwan; D, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440747/files/figure.png" pageId="23" pageNumber="474">19D</figureCitation>
); the corneous tip of the G1 is proportionately narrower (
<figureCitation id="27F8B6DEFFB3FF91D25DFE404790FE0A" box="[775,930,439,464]" captionStart="FIGURE 10" captionStartId="17.[151,250,1675,1697]" captionTargetBox="[175,1387,477,1591]" captionTargetId="figure@17.[151,1435,445,1654]" captionTargetPageId="17" captionText="FIGURE 10. Parasesarma kui n. sp., holotype male (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan. A, dorsal view of carapace; B–D, tip of left cheliped; E–H, left G1. B, lateral view of pectinated cristae on palm; C, dorsal view of palm and dactylar finger; D, lateral view of dactylar finger; E, G, dorsal (sternal) views; F, H, ventral (pleonal) views; E, F, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440729/files/figure.png" pageId="23" pageNumber="474">Figs. 10E–H</figureCitation>
), and in life, the colour of the carapace is black or light brown, with the chelae possessing orange fingers (
<figureCitation id="27F8B6DEFFB3FF91D2D6FE2B4030FE2E" box="[908,1026,476,501]" captionStart="FIGURE 22" captionStartId="32.[151,250,1490,1512]" captionTargetBox="[152,1435,193,1469]" captionTargetId="figure@32.[152,1435,193,1469]" captionTargetPageId="32" captionText="FIGURE 22. Colour in life, males, dorsal views. A. Parasesarma gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; B. P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C. P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. purpureum n. sp. (13.4×11.8 mm) (ZRC 2018.0799), Borneo, Malaysia; E, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440753/files/figure.png" pageId="23" pageNumber="474">Figs. 22C</figureCitation>
,
<figureCitation id="27F8B6DEFFB3FF91D554FE2B4071FE2E" box="[1038,1091,476,500]" captionStart="FIGURE 23" captionStartId="33.[151,250,1605,1627]" captionTargetBox="[152,1435,245,1548]" captionTargetId="figure@33.[152,1435,245,1548]" captionTargetPageId="33" captionText="FIGURE 23. Colour in life, males, ventral views. A. Parasesarma gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; B. P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C. P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. purpureum n. sp. (13.4×11.8 mm) (ZRC 2018.0799), Borneo, Malaysia; E, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440755/files/figure.png" pageId="23" pageNumber="474">23C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFB3FF91D19DFE0844C6FDE7" pageId="23" pageNumber="474" type="etymology">
<paragraph id="BF7CAA5BFFB3FF91D19DFE0844C6FDE7" blockId="23.[151,1437,151,680]" pageId="23" pageNumber="474">
<emphasis id="8DB77649FFB3FF91D19DFE084561FDC2" bold="true" box="[199,339,511,536]" pageId="23" pageNumber="474">Etymology.</emphasis>
From the Latin
<taxonomicName id="78C3D1D8FFB3FF91D37EFDF64696FDC2" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[548,676,513,536]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="purpureum">
<emphasis id="8DB77649FFB3FF91D37EFDF64696FDC2" box="[548,676,513,536]" italics="true" pageId="23" pageNumber="474">purpureum</emphasis>
</taxonomicName>
for “purple” with reference to the general colour of the new species.
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFB3FF91D19DFDB041AEFD85" box="[199,1436,583,608]" pageId="23" pageNumber="474" type="distribution">
<paragraph id="BF7CAA5BFFB3FF91D19DFDB041AEFD85" blockId="23.[151,1437,151,680]" box="[199,1436,583,608]" pageId="23" pageNumber="474">
<emphasis id="8DB77649FFB3FF91D19DFDB04555FDBA" bold="true" box="[199,359,583,608]" pageId="23" pageNumber="474">Distribution.</emphasis>
So far known from Tioman Island, Peninsular
<collectingCountry id="C7D4EACBFFB3FF91D2D0FDB047CBFDBA" box="[906,1017,583,608]" name="Malaysia" pageId="23" pageNumber="474">Malaysia</collectingCountry>
; and
<collectingRegion id="7D0764B9FFB3FF91D56FFDB0404DFDBA" box="[1077,1151,583,608]" country="Malaysia" name="Sabah" pageId="23" pageNumber="474">Sabah</collectingRegion>
, East
<collectingCountry id="C7D4EACBFFB3FF91D59EFDB04101FDBA" box="[1220,1331,583,608]" name="Malaysia" pageId="23" pageNumber="474">Malaysia</collectingCountry>
, Borneo.
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFB3FF91D19DFD9B4053FD72" pageId="23" pageNumber="474" type="biology_ecology">
<paragraph id="BF7CAA5BFFB3FF91D19DFD9B4053FD72" blockId="23.[151,1437,151,680]" pageId="23" pageNumber="474">
<emphasis id="8DB77649FFB3FF91D19DFD9B4502FD5F" bold="true" box="[199,304,620,645]" pageId="23" pageNumber="474">Ecology.</emphasis>
<taxonomicName id="78C3D1D8FFB3FF91D061FD9A4651FD5E" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[315,611,621,644]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="purpureum" status="sp. nov.">
<emphasis id="8DB77649FFB3FF91D061FD9A4651FD5E" box="[315,611,621,644]" italics="true" pageId="23" pageNumber="474">Parasesarma purpureum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFB3FF91D337FD9B4686FD5F" box="[621,692,620,645]" pageId="23" pageNumber="474" rank="species">
<emphasis id="8DB77649FFB3FF91D337FD9B4686FD5F" bold="true" box="[621,692,620,645]" pageId="23" pageNumber="474">n. sp.</emphasis>
</taxonomicNameLabel>
in Malaysia (Tioman Island and Sabah) is often observed at night and climbing on mangrove tree trunks and aerial roots above the water level.
</paragraph>
</subSubSection>
</treatment>
</document>
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